eur er ., 4 * , Pe od] * rl 7 - i b i “ae = | ry he sat sat Mn eC Ait ef ‘ i ‘a ' af | ii i : ' i wa - Sais = | eh ah) f cat Ee | ial ake Wit. a ALD S : euy ¢ iD «=> By,” o* > an 7 AS =) Digitized ey the Internet Archive — kar in 2010 with funding from | University of Toronto (bo a Osi Rhy Tipe | Peet Ot Bot Flory H SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM CONTRIBUTIONS FROM THE UNITED STATES NATIONAL HERBARIUM VOLUME 15 THE NORTH AMERICAN SPECIES OF PANICUM - 4 oO Nae By A. SSHITCHCOCK and AGNES CHASE WASHINGTON GOVERNMENT PRINTING OFFICE 1910 BULLETIN OF THE UNITED STATES NATIONA | -Issvrp Ocroper 22, 1910. PREFACE. The accompanying paper, entitled The North American species of - Panicum, by A. S. Hitchcock, Systematic Agrostologist of the United States Department of Agriculture, and Agnes Chase, Assistant in Systematic Agrostology, is the result of an exhaustive study of the material of this genus in the United States National Herbarium and in the other large herbaria of the United States. All the more impor- tant herbaria of Europe were visited by Mr. Hitchcock for the purpose of examining the type specimens of American species described by European authors. This opportunity is taken to acknowledge the many courtesies extended by the curators of the herbaria visited. In addition to the knowledge of the genus gained through an exami- nation of many thousand herbarium specimens, the authors have had opportunity to collect material throughout the United States and have observed nearly all the species of that region in their native habitats. Because of the lack of material and of the necessary field studies, it has been impossible to present the species of tropical America with the same degree of detail as the species of the United States, but it has seemed advisable to include these tropical species in order to bring together in one paper our present knowledge of the genus in North America. The authors describe 197 species and 8 subspecies, each of which is accompanied by a text figure illustrating the spikelet and fruit. The descriptions of species occurring within the limits of the United States are also accompanied by an outline map graphically represent- ing the geographical distribution. FREDERICK V. COVILLE, Curator of the United States National Herbarium. Il a : ’ — a CONTENTS. Page eo on Ss cic a aa a edie Sancta apes emda sus ws wa 1 I MeNenGH Of ENG WOIK....222- 220. 0- 02 0c aes dascnwcncctvcnccees 1 DPemeriGris GCEAMNNON.-.-2-.. - Fos. Jo se~ ste de cue sacemssece ns 1 ES ES CU) ee eee 2 MHI G2 525 cool cais (a eaW Us cue oumhe See won ae asm ne 4 NONI Sc a8 Sao te apis oa Se Raion oF ee se Secon weno 5 MC RMIEION 205 oso hans om Eo = Seid 2 die win oe eee oe 6 Deen ERDEDECICS, ANd TOTMS.. 00. Sasa os = oe ween ener ene enone 7 Geographical distribution...... Ma yn eS SS. Soc oumon Summa cate 8 0 SGT 1: Sa os a eI ere 8 RMR RIMROIOUS «© open sys cans Sees oan wc ea eens anes 10 I AE Serta hl hath 5 8 woe Naan ana cow eawar cera e 10 mastary and limitation of the genus.........-....---.----- cess rceseeeee il Pre-Linnzean use of the name.........-.-- FETE E DDR, ohare EOP 11 CMM CME OICUT 6 aso oon osetia se Denes cases ena ee tees ia 13 SEL ERMINE GCE DS |. Jai ee wins» owas = oes baie Sie eS eine tie 15 Genera here excluded.........-.--- ete ene a pia Beedle Soe erat 16 EIR IE DERE 0S ood pe aes ben oe wee a eae 17 nn tits PORUAANG SPCCICS. -.- <2... co eww fe ten we etree neue ence on 18 te se De 2g Sad Ste eee Aarts an wn Bawah owas 329 List of new subgenera and species and new names.........-...-------++---++- 332 IRE GUGOTNGNS 22-5 a5 cesta teks dhe aS ae ewe a> soup es oe 333 a is dine xn Ca a a oA e aah dae wn eee eae hun tenes = 389 CONonh WN HE ILLUSTRATIONS. TEXT FIGURES. EEE MMTIUIVOTINL «oo — Sposa oe no wawene chro nen eign wane sds aes IE TTINENTILPRLT IOC 5 ea ese crete oa a eh I Io ewe Ve quinn ase ’an 4 aoa INR 2 horn ak os ks Keo Onda Kaataeptinag SSS Smeg emaesadehice ses IERIE he ose oen cae Oe Sete en oak 3 aia EPISTIDULIGH, OFF. TEVEMCRONLE =< 2. = enn 64 47, Distribution of P. hirtieawlé. 2c -ccistaete es tee --- >. 2+ - <= 65 45. oP. POMPINOSUM «w00e op oo enone eae sioaeh bree sr +s. >>. --— on 66 49, Distribution of P.. pampinoswm...-.--- 2. -..<---+2-<---5--eeeee ae 66 BOD P; atraminewninss =~ inseas se oo oe oes eee ese eeescecscsiecs nn 67 51. Distribution-of P.. stramineum. ==. 3-222... --.222->-.---.-aeee 67 52. JP SONOTUM a 2ojic cose ec oc os cease oe eae ene oe oe sien =. a rr 68 5S. PD. POTOUMis doce eee cea ne ween Maes ce ee aioe ee oo een ei te a 68 54. (P: mMilacewin 203.2322 focee cash ae see seo ece ee rr 69 55. Distribution of P. miliaceum. . 62.2 -2.26.222c J.<--15+-- ee 70 56: P. cmyennenger. sc) oe oe keel ees toe pen See {mee ae oe eer 70 57. P. capilarioides.s. 2. 25 )o 2 eee oe eee ee ee 72 68. Distribution of P. capillariides: 2.00.2 s2cc+-s ++ -- se seen 72 69; SP: Gif USUI... Bhs 6 obs ac ooas os canis Se og see Sale eae a= a re 73 CO. PLP pS... a6 eons hoes eben weds ceeeenae aes cane eee 73 61. Distribution of P. filipes.. 2.2. -22....2..-./4--- 2202 see 74 G22 PY halls. ccc. ood ie ns Plea so occ eee eee ee ee 74 63; Distribution of P. halle... 22.2. 22-2 Sn oeee desc cee en er 75 64. P: lepidulumte. coi tele fos c ce te le vhs eon oe sen calles or 76 65; P. ghiesbreghttt 0.52 22.. 52 cnn sos coe tees face 2 ee 76 66. Pi hirsutim,. 0202-23 00 ooo. ek ee eee s ose = eer 77 67. Po MAMIMUN ac oda ee ees ooh isncte see ee eee er 78 68. Distribution of P. maximum: . ..iv...--- 2-222 -.-1 223) eee 79 69: P. plenum. . 0.6 5.20.0s Sage oe. Seee s seas Oe ee 80 70; Distribution of P. plenum. -....2.55-..-+--2- 2-2-2 «== == 81 TALOP. bulbosum. -.i sc. Ss oe Se Joes ee se seer ee ee ere 82 72. Distribution of P. bulbosum...............- 2k s. 2.2. a 83 73. P. bulbosum sciaphilum..:.2....2222cc0. +.) = ==. 2 + 83 74. Distribution of P. bulbosum sciaphilum.........-.---2 sees 84 1D. PePePens. oie we occa cicilcencccess cases deess een es 6a en 85 76. Distribution of P. repens. .....-......-J 22.2. <= «2-65 86 ihe Ps QOUiNE, See ce Se cs eee sche eee ee ee hs oles 86 78. Distribution of P. gowini........ 02.0. =2> seen 25-2 87 19. Po OUMGGUIN oe avin van cea ese acne seceeneneena ene ee =e 89 80.. Distribution of P. virgatum...........2...- ..eebe. 0) «eee 90 Sl. P. virgatum cubense. .. ... 000. cancccccus sss eee ase = eee 92 82. Distribution of P. virgatum cubense............0..--- 2-24 92 S85 Ps NOVARA c wnic'cin tin b's bs «mn avn wee Se ei ee a 93 84, Distributionol P. havardit. 02... cc cccccawed sone + a= eee 93 85> POM OTUMN Sc ows Saw eae ola Oo 5.5 wha otic Se Ee Oe el ee ee 94 86. Distribution of-P. amarwi. ... ...csscescneus0=< ===> a> = =k een 95 ST) BL AMOATUUIN, «we 5c wc e lea = os cos ae ee an oe pee en 96 88. Distribution of P. amarulum..... 2. c0s000 1b ices oss 5a 96 C9 e Pe MENU ade Sow dna a we SSS OK Sean ie ee ce Ses Ss asc en 97 90: Distribution of P. tenerwim... 20. cecenede ens os been.) oan «on 98 OU Pe GUANO ER. 5 desea a sy 0s secs behind Siete wn n= nb «nis pk ma 98 OZ) Pagrostotdes. 2.05 oo. 25 Se ey 100 98.. Distribution of P. agrostotdes..... 2... 6. o ne ocd d ne nes cce ses een 101 Oe PS CONMETIS TN eas dave a < cracerire cee ee eae Doebicts sk ce «ce 02 ILLUSTRATIONS. Ix Page ees ANetrenOn OF 2. CONMENSUM 086 2 oc. -- we wee ee eee 103 PL 20 wee cores cc ete serena b neae crn edsecedeay senna 104 NE 0 a ea 104 ET RS ee that ede ae ee nS 105 eRe en GE t > /LOTMPIONUME <6 5 = we a wie oe seen een once 106 RC Be EO A gS re hae, en 106 arti Het OFF. COMMA -.52- FINS Fe 2s SEU AL Dc eee net ce te eles 119 IS Rn. SE Re ee ee on ye eh aoe et yn Soe 120 IE NNLSEN (7 80S tI A enn Seat oe a ok eae oon 121 NINES 6 3050, * «oo toss = OE as ws Soe sc soso ne Se 122 IM AE 92-52 ECE AT Sh os Se. Se oc nom wou te ye 123 EE ae AS Air arity TRE a, PE EE ho eS 124 TN SR tate ad Sal pa agile) =e EE ee er Se 125 EE See S57 Ra 1S a cp a cm ee eeea= oe 125 EE SES aS aA ae Pe pe ag ee Sp Re i a” Pt Poe ce 126 EINE. 20.90 Gree h a rot oa cise So hae an wna eee ono ns 127 nen Gh F. VETUCOSUM.. 525... 2-2 ~ cece ese cane cane tee een es 127 IG 2 2d St ee et re eee bp em eae 128 i riDution of fF. brachyanthum......-..----..+=--2<----s+----++--- 129 ESTE 2 SR eat ee ee, oe leoe5, 5 i pene sei A a 130 ETT NENT | Rags BT RDN 2 ol RII et 2, oo Dye Si merce eed Se 131 EI st song Bre ott ne a aaa < aoe ae 133 nretOn OF F. UPDIULEONUM-..- Sol css cen ene ns wae ee ees wie 133 NT ERAT Se ee POP Sas Di ap Tk: eh rae le a 134 NNN say Sckntas sno taut ket ove mennee ue an «we ohne sake ae 135 EES 1 SNR eps Bet ha eatin 28, Ee A a a 137 IN ee Secs cn ok Sate ae ee Se ee ae ns Sa cae =e rg a 138 toe oe he yt oie on eee whee ue cease awekes,s 139 SEE A RE Rn ey EO en ere oa 140 Ee RE OR IES Se eA A A eee eee Ane ee 141 I nn oa Wt, wee aka ied wenn dee shaeas 152 Eee arettipution of P; depauperattim.....-.-..-..---00+---- 20-20-2052 153 cn a oa es nas oe an ae wba hee a oe 154 NREL GAP. SPIONGUND so So oan wos ne oe eee ee ose nen 154 NE EEO OE CE EE te 155 nee eeeeripution of P. linearvfoltum..........--- 2260-02 0--- nn en ene nenne 156 IIE ods ee emilee, See ae abn oe 157 I PG yy ee ne ee ee 157 ne Ge oe ae snc wa eed aenfanewsy 158 EES 9 eS | a ee ee 159 Te Ml en a im nm cmv ne as a teme nee 160 EE a a a eT 160 Ke ILLUSTRATIONS. Page. Hue.148. P. salapense strichiramevm..- 5-2» = caesarean ne 33) - -o0 2< 5 161 149. Distribution of P. calapense strictirameum..........-«..-----n-seeeme 161 ES ae ge 2.) ie ae ey eee 162 15; Distribution of =P iciliatuimn= 2 seees. co ec 2 <3 22 = oe = ee 163 152.) PP POlgCGwlont ania wnn enna sine n anne = wegen sean snlln 64 mee ern 163 153. ‘Distribution of P. polycaulon.....2..-..-.+--------+----5-e eer 164 54 GP SIPIQ OS WIN Sisjnim sine Selatan = wale Sia2 sin ot in, oo bie hsmeiel = Sh> S:2 5 164 155. Distribution of 2. sirigosum.........-.-:.2-..-------- 20298 165 VAG ee. ACICWLATE sees ote a. acne saa oie oe alee apne see De aos ee 167 157; Distributioniol P. aciculares....-2.<<-'s sc 2 .0t2 seconde eee 168 | 158. P. chrysopsidifolwm... «22s s2ce dose = gab esc O12 = <-5 een 168 159. Distribution of P. chrysopsidifolaum._.......---...-+.-+4-:08een 169 160... Ps CONSONGUINGUML«..-~ -0sdaes snakes dee eed oe cee ona 2a 169 161.. Distribution of P: consanguinewm.---. 2... =~. 22 2-22 .2-ee 170 162. PGngustifolnwm .. oo: sie ac 2 = cece bo ccm cere ese an == enn 171 163. Distributionor P. angustifoluun.-- 2-2-2 22> 5-2. 71 164. PB: fUst[Orme-aiscioeisecacleto ete stee sae cess oases. ee 172 165. Distribution’ of P. fusiformes.-..- <= -cs---- ans 2 ee oe 173 166. 4P: arentcoloides. cc 3. n.d occ Sacc = odes e+ hs shee eee 173 167. Distribution of 2. arentcoloides. ..<......-_ 252-0 eee 174 168: 4P. OVINWIN 2 ie oe 8s 5 be ok Ae eet ee ee ee ee eee 175 169. Distribution of Ps ovinuni......5..<2cs---- = .2ces seer 175 170. Po newranthwm. ssc. ecb secs koe oo bas tee cnet. Se ee 176 171. Distribution: of P. neuranthum.........-.---+..+--46s- eee 176 W2. Py brveknelliticcecccencew se oeeteee <4 loo oss s Hots. eee 177 173. Distribution ot -P. biceknellit-s.....-.- 22-82. 5e eee 177 14, 2. calliphyllim.. 05s. sanseew s+ - she gee +. = ote + 05 ge een 178 175. Distribution of P. calliphyllum. ........-.--..5--0- 7---24-e eee 178 WIGS BS MUGICOUE 358s Sia Penn oe eee he ewe See bee 179 77 Distribution of P. nudicaule...<5-s2.4c5-22-56+-6> 25455 179 178. P-MAUCTOCORPON 2s oa io «.si0 35510 om a0 wisi ciaee es oes 5. 182 179. Distribution of P. microcarpon..........-.<2s-.+-+<.-425ne 182 IO PR ie WA aac oi Re RT oa oe jis cates cn acicn oa 184 181). Distribution of P. nitidum.....--.-. QIDEMATIENSEL << ford eter nie vist sce eek week eee ee eee er ee oe, See ee ee . Distribution of P. implicatum........... SUMEEDILIIGH. OF FP . RUOCKUCHE sn ~ uch eaeee eee ttt Seek ee eee eek se Be SIEACHUCRC SULURCOU: « oeeraee ck. Hee De eee ee nee Oa ee pe lustzipution of P. huachucae silvicqla......02+-22.0- 2225.2 ee se sees OS LORS SDT OES a eee Oe ene, pie OT Tee: a Dey a eke Boe gS a MINIM 6 sade Win a ner CUE RIOR EL ot eee eke Oka Ue SRE ITLION OF PF. LANUGUNOS UM 2. <0 aera swine ene Se a De oe oes mermiaon. OF J. auburne.....osstuee Ce eee Sk ee eee ET INSEE ek one nig oe CRE RRR OE Ee ote . Distribution of P. thurowii...............-...-- fe ester. se ITTY TD re on ce OEE Ele LOE RES eee MEIN in nisin cee gia see mn aC EME Ith. ace LLL LU. ote MIO CLF. STUCCIIUS. S502 5 oon ceccew becnmes ys SEKe Le oe MEN C0 FUTUR... aad nip en edo a OLE Sce sou ae NNN i rete a sy ol np EARS daw KUL Lode eeebk ont eemrenuenon Of P. Pangiidum... 2 nn oe nnn n nee wee eee tls seteos XII Fig. 254. 255. 256. 257. 258. 259. 260. 261. 262. 263. 264. 265. 266. 267. 268. 269. 270. Pele 272k Biles 274. 275. 276. marti 278. 279. 280. 281. 282. 283. 284. 285. 286. 287. 288. 289. 290. 291. 292. 293. 294. 295. 296. 297. 298. 299. 300. 301. 302. 303. 304. 305. 306. ILLUSTRATIONS. Page. Distribution of P. scopartoides.....-.-.cccccscccescccsesccconcscaee 239 (Pe SHASTENISC® osc, « <0.2 ae w= c= = sicsinie ste wlainisie sialaie,es ae >ie= =| == 239 Distribution of P. shastense... 02. 0 cccccnosncss.s-tescess ocean 240 IPI OLACON << sccenergntreoe = <5. = 20 rioee een es, 7s ein ie enn 241 Distribution of P. malacon ..... 2. - 22 0<0000025 000 700s oe nn 241 BP COMMONSIONIUIMN.« «2 w= 2's 2.5 oo ore amie mes ils e oie oe > ae oe te 244 Distribution of 2. wilmingionense: « «./<-.--.- «<= ects 10-5 ee 245 PPG ELOTUAN os o.o mya > ey ce = nieie eine se oe 52 > 2 ie So el 245 Distribution of P. tSugetorum. - . «<< <20.02,060:0 0008886 3s se 246 IP NCOULOTANIUT s «3 2a = tie = ote inia ies lee eioee emia seo oe oe aoa ene een 247 Distribution of P. columbianwm.. - «200-0005 60=5 55 <0 onsen 248 P. columbianune thinvum.--~. 2-2 = eheistee.e 2 el ole ae ee 248 Distribution of P. columbianum thintwm....-...-.<2.-.-.sseeeeeeee 249 BP. OTCOMG aa sac a noe da 2 oo = bo = cidin,o Siskel PO slates Sle =,5 eee s ee eee 249 Distribution of P: orieola -- - 2... 22 ais bee eel ele ee 250 P 8pNQErOCOPPON «6. 2.0 = 22 < on onin = ola Se cls wskee = e els o 251 Distribution of P. sphacrocarpon......-- -.--....00-0-=0s 05 = 252 P. sphaerocarpon inflatum. ......- 2200s 6ose een + ne 254 Distribution of P. sphaerocarpon inflatum.....-.------------++++++-- 254 P. POWarthes.... ssc 2ocs ne paees bres case nceS = s 46s eee 255 Distribution of P. polyanthes ......--- 20-005 -.s00-26 =e en 256 P erectifoliitins «2 2 ons cnine a no 2sic wa onielaeieinidiclotn ee ee 257 Distribution of P. erectifolium . ...-......+.--i02 2.058) aoe 257 PONE Se voc on cease Sass cieceae + Seelseee eee eee pee en 259 DistnibUtt ONOmeatenUuceseearees see eee eer ee tere °c 260 P. albomargimatumn. .-.-- 2-0-0 «2 oe Senn aise 5 6 = si< ieee 260 Distribution of P. albomarginatum......-...<+..----2-2ss5e= ee 261 Ps PU OUMM wna Bie c on a ancm eo 2 wis iwc mca ela wale am age 261 Distribution of P. irifolium..... 2.2... sss 32sec ==. ~ nee 262 PUT ONG oa cpenin wea ae so oe eine se ee nee ge 262 Distribution of P. flavovirens... 2... 22522 see on ee 263 P\CONCINTAUS. 2.52 <2 0aceaes Son pn ea cee ge eae See ee umearinitiion Of F’. oligosanihes .... ocd. coc ccke nce sdneseneee ss . P. ravenelii EAMRRME SLT. TOTCHOING 2 200 ooo Se rcncis os man eieamen gees kiseuenah 2s REINO 5s ce oa ES ie oe bei os ios Rad aes ooo 2 RILEY EE? PADRE oh he eden ee osm ane b5~ IN SINOIINON sk re ia ao ols IS re oe ns Hee See ae aids 4a emaripimaon of P. zanthophysum. ! 2-222. --- 2. =... 2.22 - econ - es ENE AN oe esc 2 Le A. RRS 2 Sos coo ee owe bce cesce macunninen OF P. pedeellatum: ... . 2.2. Sods~ fas cela s de con cesses es . P. nodatum nee GEE MEIN sos 2 5 eas cin onan oo eka p awe sk I SES A Ree Oe ee ee tr ne eee . Distribution of P. scoparium..........-.----- Bes er the en er EEN oi ae ast ee one ee a 8 cok Oh cise oe tog oe aon ence- MINS EY feo 87. Unban ee BA oe Sa CnMILAIN OLE, CEMICACUM 2. ne 2 Se tea soe eiee sea ass Sewstense NT SE ee eee ae eae Se ee nS ay Smeaton Of. SeGUriisenlUeMn «22 eae we a awe oe oo asc sa nen NNN Frc sk pole ae ee el ARSE AL Oona was nsnOon Of Fs cryplanihem.2. 552 -. on ae we oe a nnn e enced ee EASES Ta SY age (72, gee, Die cl OE MILIEU Se oe een te eee pe one cc Sits en De oes ake PEABO GION Of -P. CONUNULATUM. —. .. . 22 Soochow ec ecco ccclhcewcavn BPEEIITLION OFF. Wulapiess = oles. so Sales Scien Soe eon ecescs IN Woe oa Ws en OE hl ge Mains wit wk inl dom ae eo a ee eee: eee re EN oi Ss re hag Oth ERR SUES ool xed shes cad oD ie ke eRMMOR 2, BOMEIOIONE, ia hae ts. sis bass ken fens cu eaiene ee ee a AT RS ee a = EN Oe rd RG att at 6 ak Peis 22 alan mabe wn Pee x dab eta de eee SNEIPIRIAL ONE, COPMOUMNE 5 cin noel owe bee rab noecs tue nens PIGMENT E WOOSCI, «ode nns 2s cee oh oo ocd nnd Se Sedends LET TIE NES Re ES tn eS ee ee XIV ILLUSTRATIONS. Page. Rie; 360. Disizibution Of /P. (b08eit Molle. o.ccccnc. oc nonacnass kee e eee 320 SGU 5, Ee OOVUBUAN breve ists Swing Al onsid Svs e Owice eke Oe san ee Ves kil eee 321 362; Disitibution of P: ObtUsUihs. «noon oss conlwncosbenbie te See 322 963. eR HeMMOMON. b6 bcoz oso own o IE ee ae eee be). DAS ee 323 $64.- Distribution Of.) Remitomonsanccrcc-cnss< ss cceeice ees ls eee 324 365). Po CUAEESUNU Mac Britton, Man. 89. 1901. f Nov. Gen. & Sp. 1:103. pl. 38.1815. ¢ Proc. Biol. Soc. Washington 21: 6.1908. 9 Allg. Bot. Zeitschr. 12: 60. 1906. dF]. Brit. W. Ind. 551. 1864. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 17 GROUPING OF THE SPECIES. The genus Panicum, as here limited, contains two groups sufficiently well marked to warrant their segregation as subgenera, namely, Paurochaetium and Dichanthelium. Nearly all the remaining species group themselves around the central idea of the genus as typified by P.miliaceum. In order to avoid assigning to the main portion of the genus a name of formal nomenclatorial significance, such as Eupan- icum, this group is called true Panicum. There are five outlying species which show no close relationships with the others, but are not sufficiently distinct to be assigned generic rank. These are placed in a final group under “ Miscellaneous Species.’’ It is probable that a further study of the species in other parts of the world will indicate that at least some of these species may, together with extra North American allies, be placed in definite subgenera. The group true Panicum and the subgenus Dichanthelium are susceptible of further division into minor groups, the names of which are the plurals of the characteristic species of each group. These names are not intended to be formal and should have no nomencla- torial standing. The term Angustifolia is used as if we were to say, P. angustifolium and its allies. A few tropical species of true Pani- cum remain ungrouped, as they have no is allies in North America, and there is no advantage in making a group for each species. The groups of these two main divisions are arranged to represent our judgment as to their relationship, so far as this can be done in a lineal sequence. In true Panicum the first group, the Geminata, is furthest removed from the typical species, the inflorescence resem- bling that of Paspalum. The Capillaria, Diffusa, and Virgata, typical groups, are near the center of the series. In the same way, the Depauperata are an outlying group of Dichanthelium, the typical groups being the Dichotoma and Lanuginosa. The species of each group are also arranged to represent their affinities, but it is impossible to indicate the difference in the degree of relationship. Some of the groups are manifestly more homoge- neous than others. The group Geminata, for example, includes two closely allied species, P. geminatum and P. paludivagum, and one, P. barbinode, in which the affinity is less evident. The latter species is placed in the same group as the other two partly as a matter of convenience. If there were several other species more closely allied to this than to P. geminatum it would have been more convenient to segregate these as a separate group. One more example will suffice to illustrate the unequal value of the groups. Panicum reptans is placed in the group Fasciculata because it is more closely allied to the other species of that group than to those of any other group, yet the 41616°—vo1 15—10——2 18 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. degree of difference between this and any other species of the group is much greater than that between such species as P. fasciculatum, P. arizonicum, and P. adspersum. But other groups, such as the Lanuginosa, are made up of closely allied species, connected with one another by intergrades, to form a composite taxonomic network whose component parts cannot be definitely distinguished by clean-cut lines of demarcation. DESCRIPTION OF THE GENUS AND SPECIES. PANICUM L. Panicum L. Sp. Pl. 55. 1753. : Steinchisma Raf. Bull. Bot. Seringe 220. 1830.4 Phanopyrum (Raf.) Nash in Small, Fl. Southeast. U. 8. 104. 1903.5 Spikelets articulated below the glumes, falling entire, more or less compressed dorsoventrally, arranged in panicles, rarely in racemes; glumes two, herbaceous, nerved, usually very unequal, the first often minute, the second typically equaling the sterile lemma, the latter of the same texture and simulating a third glume, bearing in its axila membranaceous or hyaline palea and sometimes a staminate flower, the palea rarely wanting; fertile lemma chartaceous-indurated, typically obtuse, the nerves obsolete, the margins inrolled over an inclosed palea of the same texture, a lunate line of thinner texture at the back just above the base, the radicle protruding through this at germination; stamens threé, styles two, stigmas plumose; grain dorsoventrally compressed, with a punctiform hilum, free within the firmly closed lemma and palea. Annual or perennial herbaceous grasses, of various habit, confined to the warmer regions of both hemispheres. A number of species here included in the genus Panicum depart in some measure from these generic characters. Thesubgenus Paurochaetium approaches Chaetochloa Scribn. and the group hitherto referred to Panicum section Ptychophylium A. Br., in that the uppermost spikelet of each branchlet is subtended by a bristle-like prolonga- tion of the axis. Panicum geminatum Forsk. and P. paludivagum Hitche. & Chase have a racemose inflorescence as in Brachiaria Griseb., but the spikelets are placed with the back of the fruit turned toward the rachis as in true Panicum, not in the reverse position asin Brachiaria. In P. barbinode Trin., P. arizonicum Scribn. & Merr., and P. tevanum Buckl. spikelets toward the ends of the branches are often placed in the reverse position characteristic of Brachiaria, while others on the same branch are placed with the back of the fruit toward the axis, showing that in an inflorescence not strictly racemose this character of the position of the spikelets in relation to the axis is not of taxonomic significance, since it depends on whether one or the other of a pair of spikelets on a one-flowered branchlet has been developed. Hence this charac- ter, while distinguishing between Paspalum L. on the one hand and Axonopus Beauy. and Brachiaria Griseb. on the other, does not alone clearly separate the latter from Panicum, but must be taken in connection with the strictly racemose inflorescence. In Panicum elephantipes Nees the thin but not hyaline margins of the acuminate lemma are not inrolled above the middle, the fruit thus suggesting an approach to Valota Adans., but in texture it is not cartilaginous and papillose as in that genus nor does P. elephantipes approach Valota in habit or inflorescence. In the Verrucosa and the related Trichoidia the firm-margined lemmas are not inrolled except at the base. a See discussion under P. hians, p. 118. b See discussion under P. gymnocarpon, p. 327. Zl The first glume in this genus is typically not more than half the length of the spikelet, and is commonly much shorter than this, but in a few species, as in P. glutinosum Swartz, P. urvilleanum Kunth, and in most of those included under ‘‘ Miscellaneous Species ’’ the first glume is nearly as long as the second. All the outlying species at the end, nos. 192 to 196, depart in some particular from typical Panicum, such departure being discussed under each species. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 19 KEY TO SPECIES AND GROUPS. Axis of branchlets extending beyond the base of the upper- most spikelet as a point or bristle 1 to 6 mm. I xterra s Sele cate n re eee we saben ate See subgenus PAUROCHAETIUM, SB ae. Axis of branchlets not extending into a bristle. (In P. gemi- natum and P. paludivagum the somewhat flattened axis pointed but not bristle-form.) Plants annual. Inflorescence consisting of several more or less secund spike-like racemes. Fruit smooth and shining; spikelets about 1.5 mm. long. Rachis bearing slender bristles (these wanting in exceptional specimens); nodes usually villous.. 59. P. pilosum. Rachis without bristles; nodes glabrous. ........-.-- 60. P. larum. Fruit transversely rugose; spikelets 2 mm. or more IR la aca = tee Ma niin eee is a ea See FascicuLatTa, ; p. 30. Inflorescence a more or less diffuse panicle. ° Peers, SRPTOMISLe..o25 U5) ksh ose lleced Joyous See VERRUCOSA, p. 126. Spikelets not tuberculate. Spikelets 1.2 to 1.4 mm. long; tropical species ...... See TrICHOIDIA, p. 129: Spikelets 1.7 mm. or more long. Glumes and sterile lemma hispid along the mar- ES er ee te ae gens dee aS 76. P. costaricense. Glumes and sterile lemma glabrous. First glume not over one-fourth the length of the spikelet, truncate or triangular-tipped.....See DicHoTomIFLoRA, p. 47. ~ First glume usually as much as one-half the length of the spikelet, acute or acuminate..See CAPILLARIA, p. 54. Plants perennial. Spikelets short-pediceled along one side of the rachises, forming spike-like racemes. First glume nearly equaling the sterile lemma. Racemes spreading; fruit not over one-third the RORtas On WO GDIMAIOE. oo. 2 occ eco e en beka viene 196. P. gymnocarpon. 20 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Racemes appressed; fruit nearly as long as the spikelet. Spikelets silky-villous, pointed...........-....--- 194. Spikelets glabrous,,obtuse....s,..%-Jiesp os. sees 192. First glume much shorter than the sterile lemma. Spikelets hispid and with two crateriform glands on the sterile lemma. Spikelets not over 2 mm. long; blades not over AW CIN HOM Et. cts sie 22 ois cine HME wipe Swi Se Se 66. Spikelets 3.6 mm. long; blades 4 to 10 cm. long.. 67. Spikelets glabrous. Blades lanceolate to ovate-lanceolate; glumes strongly carinate; spikelets 2.3 to 2.8 mm. long; tropical species. Blades not over 5 cm., usually 2 to 3 cm. long; second glume rather blunt and shorter than fhe Brenie lemma, :..' =<. o20 7 eeheo es ween 64. Blades 5 to 11 cm. long; second glume acute, but slightly shorter than the sterile lemma. - . - - 65. Blades linear, often elongated; glumes not carinate or but slightly so. Spikelets not over 1.5 mm. long. Spikelets pointed, not expanded at maturity by an enlarged sterile palea.....-.-....- 58. Spikelets blunt, expanded at maturity by the enlarged sterile palea. Rachis bearing slender bristles (these want- ing in exceptional specimens); nodes usually villous) ==... ee aetaee toe 59. Rachis without bristles; nodes glabrous. ... 60. Spikelets 2.5 mm. or more long. PARIS IPULORON SL ScialS oo. ond slew ee 57. Axis not pilose. Fruit transversely rugose..-..-.....-..... See Bruit not rigose. .. /s22er20 Se Mee Lost vend te 193. Spikelets in open or sometimes in contracted or congested panicles, but not in spike-like racemes. Basal leaves usually distinctly different from those of the culm, forming a winter rosette; culms at first simple, the spikelets of the primary panicle not per- fecting seed, later usually becoming much branched, the small secondary panicles with cleis- togamous, fruitful spikelets..........-.. See subgenus Basal leaves similar to culm leaves, not forming a winter rosette; spikelets all fertile. Prat transversely rugose. *.... . 2... eee een See Fruit not transversely rugose (minutely papillose- roughened in P. millegrana). P. ciliatissimum. P. obtusum. P. pulchellum. P. biglandulare. P. stoloniferum. P.. frondescens. P. polygonatum. P. pilosum. P. laxum. P. longum. GEMINATA, p. 30. P. hemitomon. DICHANTHE- LIuM, p. 142. <———S ee Maxima, p. 78. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Gl First glume very small, not over one-fourth the length of the small, obovate, blunt spike- let; tropical species. (See also P. repens with permtenispreeleis) S020 sds 2d Sie. eto. See First glume as much as one-third the length of the spikelet (shorter in P. repens). Spikelets pubescent. Fruit silky-villous; spikelets 6 to 7 mm. long, Yo ge | Lh ee 75. Fruit smooth and shining; spikelets not over 3.5 mm. long. r Culms and sheaths glabrous or softly pubes- cent; blades ovate-lanceolate.......... 78. Culms and sheaths densely harshly villous; blades menrss>..30))2 2o0 Sees. 80. Spikelets glabrous. Sterile palea enlarged and indurated at ma- turity, expanding the spikelet; blades scarcely wider than their sheaths. Spikelets 3 mm. long, congested; panicles ark puree. 205.2 32) eee 3 S22 2 63. Spikelets not over 2.4 mm. long; panicles green or pale. Panicle branches spikelet or branchlet- bearing along the upper half or toward jie -pnies Gabyels $s soe fast 62. Panicle branches branchlet - bearing throughout their length or nearly so. 61. Sterile palea, if present, not enlarged and indurated. Plants forming conspicuous creeping, scaly rootstocks. ; Spikelets long - pediceled, not secund, arranged in open or contracted panicles .n25 soctee steed kS 2. See Spikelets short-pediceled, more or less se- cund along the nearly simple panicle branches. Panicles open; spikelets 3.4 to 3.8 mm. long (shorter in exceptional SPOCHREDS ise. sawee See ohn od 55. Panicles more or less contracted; spike- lets not over 2.8 mm. long......... 56. Plants not forming creeping, scaly rootstocks. Fruit crested at the apex; spikelets 5.5 to 6 mm. long; tropical species..........195. Fruit not crested. PARVIGLUMIA, p. 124. P. urvilleanum. P. millegrana. P. rudgei. P. cupreum. P. hians. P. exiguiflorum. VirGATa, p. 84. P. anceps. P. rhizomatum. P. zizanioides. 22 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicles narrow and few-flowered; culms erect and wiry; blades drying in- WOLUMeS o> sha seo teelepece Beee See TENERA, p. 97. Panicles open or contracted, many- flowered. Panicles 40 to 60 cm. long, the numer- ous elongated branches in verticils; iYoplcal, species... 22 jsiniecaee = 81. P. megiston. Panicles mostly much less than 40 cm. long; branches not verticillate. Spikelets short-pediceled along the nearly simple panicle branches.See AGROSTOIDIA, p. 99. Spikelets long -pediceled; panicle open (dense in P. hirsutum). Spikelets viscid; first glume about as long as the second; tropical BPOCIESs a5isb5 vies eet oes = 79. P. glutinosum. Spikelets not viscid; first glume much shorter than the second. Culms erect or rarely spreading; blades linear, usually elon- gated; spikelets pointed..See Drrrusa, p. 71. Culms decumbent at base; blades ovate to oblong-lan- ceolate; spikelets blunt; tropical species. Panicles mostly less than 5 cm. long; plantsglaucous. 77. P. parvifolium. Panicles 10 to 20 cm. long; plants not glaucous...... 78. P. millegrana, Subgenus PAUROCHAETIUM subgen. nov. Perennials; culms tufted, erect, blades not over 7 mm. wide; inflorescence narrow, more or less interrupted, the branches short and appressed, the ultimate branchlets bearing 1 to several spikelets, produced beyond the uppermost spikelet as a bristle 1 to 6 mm. long; spikelets 1.5 to 3.5 mm. long, much swollen on the face, glabrous; fruit transversely rugose, apiculate. The type species is P. distantiflorum Michx. This group approaches Chaetochloa in having branchlets produced into bristles, and in the shape of the spikelets and rugose fruits. Blades elongated, usually more than 15 cm. long, narrowed toward the base. Spikelets about 3.5mm. lone. ...:. co seeeen eae eee 5. P. reverchoni. Spikelets about 2 mm. long, or less. First glume rounded or truncate; second glume about Ss long Sac irUit. 5. teenies. cies eae ces eee 3. P. chapmani. » HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 23 First glume acute, second glume about two-thirds as long as fruit. Spikelets 1.5 mm. long; blades involute........ 1. P. distantiflorum. Spikelets 2 mm. long; blades scarcely involute.. 2. P. utowanaeum. Blades usually less than 10 cm. long, not narrowed toward the base; spikelets 2.5 to 3 mm. long. Blades of mid-culm long-acuminate, usually 2 to 3 mm. MER ee ree Es Satis FAS hE eo aN EE ed US 4. P.ramisetum. Blades of mid-culm abruptly acute, usually 4 to 6 mm. SIG opts eke ake ere ne ee 6. P. firmulum. 1. Panicum distantiflorum Rich. Panicum distantiflorum Rich. in Sagra, Hist. Cuba 11: 304. 1850. ‘“‘Crescit in graminosis montosis insulae Cubae.’? The type in the Paris Herbarium is labeled *‘in montosis ins. Cubae,”’ and was received from Sagra. In the same herbarium is a specimen of Panicum megiston Schult., from Cayenne, which bears a slip with the name ‘‘Panicum distantiflorum,’’ accompanied by a diagnosis and drawings of spike- lets. The diagnosis and drawings apply to the Cuban specimen and not to the very different Cayenne specimen. It would appear that the drawings had been attached to the wrong sheet. DESCRIPTION. Plants cespitose, glabrous; culms 60 to 80 cm. high, slender, wiry, compressed, pro- ducing slender, sometimes fascicled branches from all the nodes; sheaths longer than the internodes, but narrow and sheathing the joints only at the base, flattened, a minute tuft of hairs on the auricles; ligule a ring of very short hairs; blades erect, firm, narrower than the summit of the sheath, linear to almost capillary, as much as 30 cm. long, 1 to 2 mm. wide, mostly strongly invo- lute, at least the lower commonly more or less curled, usually with a few hairs at the base; panicles numerous, 2 to 7 cm. long, very narrow, the branches appressed, scarcely overlap- Hie Lace" dieeeaaen. ping, the lower 8 to 15 mm. long, the branchlets bearing 1 to From type specimen. 3 subsessile spikelets, the setiform prolongation of the axis rarely equaling the spikelet, usually not more than 1 mm. long; spikelets 1.5 mm. long, 0.7 mm. wide, ellipsoid, acute, glabrous; first glume about half as long as the spikelet, acute, strongly 5-nerved; second glume obtuse, two-thirds to three-fourths as long as the fruit and the strongly 7-nerved, acute, sterile lemma; fruit 1.3 mm. long, 0.6 mm. wide, elliptic, pointed, finely rugose. DISTRIBUTION. Open rocky soil, Bahamas and Cuba; apparently rare. Banamas: Inagua, Hitchcock in 1890, Nash & Taylor 893 (both in Field Mus, Herb.). Cusa: Playa de Cojimar, near Habana, Hitchcock 144; Colombia near Habana, Leén 305b, 567; Santiago de Cuba, Leén 912, 917; Playa de Marianao, Leén in 1909. 24 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 2. Panicum utowanaeum Scribn. Panicum utowanaeum Scribn. in Millsp. Field Columb. Mus. Bot. 2: 25. 1900. “No. 702 [Millspaugh Plant. Utowan.], from a dry hillside near Guanica, Porto Rico, Jan. 22, 1899. Type in Field Col. Mus. Herb. No. 60702.”’ In this specimen the root- stock is scarcely visible, but in the duplicate in the National Herbarium the slender rootstock is shown. Panicum sintenisii Nash, Bull. Torrey Club 80: 382. 1903. ‘‘In woods, Guanica, Sintenis 3463.’’ The type, in the herbarium of the New York Botanical Garden, is labeled ‘“‘Guanica, in sylva ad Cerro de la Ensenada, Porto Rico. Sintenis 3463, Jan, 28, 1886.’’ The specimen shows only a trace of the slender rootstock. DESCRIPTION. Plants tufted from the joints of short, slender rootstocks, glabrous; culms ascend- ing or somewhat spreading, 25 to 60 cm. high, slender, compressed, sparingly branching; sheaths shorter than the internodes, compressed, especially the lower- most, ciliate at the auriculate summit; ligule a minute ring of stiff hairs; blades erect or spreading above, 10 to 20 cm. long, 1 to 4 mm. wide, slightly scabrous on the margin (sometimes sparsely pilose on the upper surface at the base), narrowed and more or less involute at the base and much nar- rower than the sheath; panicles 3 to 10cm. long, very slender, the scattered, erect branches 1 to 3 cm. long, the bristle usu- ally equaling or exceeding the spikelet; spikelets subsessile, 2 to 2.1 mm. long, 0.6 to 0.7 mm. wide, elliptic, somewhat beaked at the summit, glabrous, pale with green nerves; firet glume half as long as the spikelet, acute, 3-nerved; second glume two-thirds to three-fourths as long as the fruit, 3 to 5- nerved, sterile lemma 5-nerved, abruptly pointed; fruit 1.9 Fig. 2.—P. utowanaewm. : seed : ; From type specimen. mm. long, 0.6 mm. wide, elliptic, minutely rugose; slightly beaked at the acute apex. This species is closely allied to P. distantiflorum, but may be distinguished from it by the wider, flat or scarcely involute blades and larger, pale, less strongly nerved spikelets. The rootstock is very slender and so easily broken off in collecting that only about half the specimens cited below show it. DISTRIBUTION. Open rocky soil, mostly near the coast, Cuba, Porto Rico, and Guadeloupe. Cusa: Triscornia, near Habana, Hitchcock 141, Tracy 9089; without locality, Wright 3452 (Gray Herb.). Porto Rico: Guanica, Millspaugh Pl. Utow. 702, Sintenis 3365, 3416, 3463 (N. Y. Bot. Gard. Herb.). LEEWARD IsLANDsS: Guadeloupe, Duss 3177. 3. Panicum chapmani Vasey. Panicum chapmani Vasey, Bull. Torrey Club 11: 61. 1884. No locality nor speci- men is cited, but the author says: ‘‘This is the Panicum tenuiculmum of Chapman’s Flora, but is not the P. tenuiculmum of Meyer.’’ A specimen in the National Herba- rium from the Chapman Herbarium labeled ‘‘ Panicum tenuiculmum §, Fl. 8. Florida” in Chapman’s hand, and ‘“‘ Panicum Chapmani Vasey,’’ in Vasey’s hand, is chosen as the type. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 25 DESCRIPTION. Plants cespitose, glabrous; eulms ascending or spreading, 40 cm. to 1 meter high, slender, compressed, wiry, sparingly branching; sheaths about as long as the inter- “nodes, compressed, pubescent at the scarcely au- riculate summit, sometimes ciliate on the margin; ligule a ring of very short hairs; blades erect, rather firm, linear, 15 to 40 cm. long, 2 to 5 mm. wide, acuminate, narrowed to the base, more or less involute when dry, scabrous on the margin and upper surface, the latter usually sparsely pilose toward the base; panicles elongated, sometimes as much as 30 cm. long, of remote, appressed, raceme-like branches bearing few to several sub- Fig. 3.—P, chapmani. Fromtype sassile, somewhat crowded spikelets, the setiform Sil prolongation of the axis 3 to 6 mm. long; spikelets 2 to 2.2 mm. long, 1 to 1.2 mm. wide, obovate, abruptly pointed, turgid, pale green or yellowish; first glume about one-third the length of the spikelet, obtuse or truncate, 3-nerved; second glume F slightly shorter than the fruit and sterile lemma, strongly 5 to 7-nerved, obscurely reticulated; fruit 1.8 mm. long, 1 to 1.1 mm. wide, elliptic, abruptly acute, minutely rugose, the margins of the lemma inrolled only at base. As observed on Key Largo the blades in this species are flat on plants growing in shaded situations and in- volute on plants in the sun. The flat blades become more or less involute in drying. Fig, 4.—Distribution of P. chapmani. DISTRIBUTION. Coral sand and shell mounds, southern Florida and the Bahamas. Frorma: Marco, Hitchcock Lee Co. Pl. 487; Cape Sable, Simpson 157; Key Largo, Chase 3926, Curtiss 5457; Little Pine Key, Curtiss 3607; Key West, Garber in 1877; ‘‘Shores of Manettee River,’’@ Rugel 394; without locality, Blodgett, Chapman. Banamas: New Providence, Britton & Brace 401; Rose Island, Britton & Mills- paugh 2137; Great Exuma, Britton & @Millspaugh 3076 (all in Field Mus. Herb.). 4. Panicum ramisetum Scribn. Panicum subspicatum Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8: 25. 1889, not Desv. 1831. “Texas (Buckley, Nealley).’? Both specimens cited by Vasey are in the National Herbarium. The second of these has been chosen as the type for the following reasons: The first specimen cited, S. B. Buckley in 1881, does not bear the specific name in Vasey’s hand, and furthermore is a mixture of P. ramisetum and P. reverchoni; the second specimen, collected in Texas by G..C. Nealley in 1887, bears the specific name, “‘subspitatum V.’’ in Vasey’s hand. Another Nealley specimen bears the name in Vasey’s hand, but was collected in 1892, after the publication of the species. Panicum ramisetum Scribn. U. 8. Dept. Agr. Div. Agrost. Circ. 27:9. 1900. Based on Panicum subspicatum Vasey, not Desv.. aThis locality, if meant for Manatee River, is probably an error. 26 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Plants pale green, tufted, from short horizontal rootstocks; culms erect or ascending, 25 to 60 cm. high, commonly branching at the base and lower nodes, scabrous at least below the nodes; sheaths nearly as long as the internodes or the lower overlapping, not com- pressed, sparingly papillose-pilose, especially along the margins and at the summit; ligule about 1 mm. long, with longer hairs at the sides; blades rather firm, erect or ascending, 5 to 12cm. long, the lower shorter and more spreading, 2 to 4 mm. wide, tapering to an involute tip, not narrowed at base, but about as broad as the sheath, sparsely papillose-pilose on both sur- faces, at least toward the base; sometimes sparsely ciliate; panicles very slender, 5 to 20 cm. long, net conspicuously interrupted, their branches erect, the ultimate branchlets of 1 to 4 subsessile spikelets, the setiform prolongation of the axis usually not exceeding the short-pediceled spikelet; spikelets 2.4 to 2.6 mm. long, 1.4 to 1.5 mm. wide, obovate, subacute, turgid, plano-convex; first glume clasping, about half the length of the spikelet, subacute or acute, 5-nerved; second glume and sterile lemma subequal, scarcely covering the fruit at matu- rity, strongly 7 to 9-nerved; fruit 2.2 to 2.3 mm. long, 1.4 mm. wide, ellip- tic, acute. Fig. 5.—P. ramisetum. From type specimen. DISTRIBUTION. Sandy plains and prairies, southern Texas and northern Mexico. Texas: Big Springs, Tracy 7958, 8229; Kingsville, Tracy 8879; Encinal, Griffiths 6380; Laredo, Nealley in 1891, Pringle 2377, Sauvignet in 1891; Eagle Pass, Havard 98; San Diego, Nealley 62, Smith in 1897; without locality, Buckley in 1881, Nealley in 1887, 1888, 1889, and 1892. Mexico: State of Coahuila, near Diaz, Pringle 8323. Fig. 6.—Distribution of P. ramisetum. 5. Panicum reverchoni Vasey. Panicum reverchoni Vasey, U. S$ Dept. Agr. Div. Bot. Bull. 8: 25. 1889. “‘Texas (Reverchon).’’? The type, in the National Herbarium, was collected by J. Rever- chon, near Dallas, Texas, and distributed in ‘‘Curtiss, North American Plants No, I,” as Setaria uniseta Fourn. DESCRIPTION. Plants tufted from short rootstocks, branching at the base; culms stiffly erect, 30 to 70 cm. high, simple or occasionally bearing one or two sterile branches, slender, sub- compressed, glabrous or the lower internodes strigose, the nodes appressed-pubescent; sheaths mostly longer than the internodes, ciliate on the margin toward the summit, otherwise glabrous, often slightly scabrous, or the lowermost sometimes sparsely strigose; ligule about 0.3 mm. long; blades erect, stiff, 5 to 20 cm. long, 2 to3 mm. wide, flat or involute toward the apex and base (the blades of the basal shoots commonly involute-setaceous), scabrous on both surfaces, especially the upper, at the base nar- rower than the sheath, the lower commonly disarticulating at this point; panicles HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 27 long-exserted, very slender, 5 to 20 cm. long, the branches scattered, short, erect, the branchlets bearing 1 or 2 subsessile spikelets, the setiform prolongation of the axis mostly equaling or ex- ceeding the spikelet; spikelets 3.5 to 3.8 mm. long, 1.8 to 2mm. wide, elliptic, turgid, abruptly pointed; first glume about half the length of the spikelet, subacute, strongly 5 to 7-nerved; second glume and sterile lemma _ scarcely equaling the fruit, strongly 5 to 7-nerved; fruit 3.1 to 3.3 mm. long, 1.8 to 1.9 mm. wide, elliptic, minutely pointed, obscurely transversely ru- gose, This species differs from P. rami- setum in having longer, narrower blades, narrower at the base than the summit of the sheath, and larger spikelets, commonly but one to a branchlet, hence most of the spikelets subtended by a bristle. Fic. 7.—P. reverchoni. From type specimen. DISTRIBUTION. Rocky or sandy prairies and lime- stone hills, Texas. _ Texas: Dallas, Bebb 1321, Bush 674, Reverchon 1096, in Cur- tiss N. Amer. Pl. 3618*, L; Abilene, Tracy 7940; Colo- rado, Tracy 7939; Weather- ford, Tracy 7948; Kerrville, Heller 1603; Gillespie County, Jermy 39; Bexar County, Fig. 8.—Distribution of P. reverchoni. Jermy 233, 234; San Antonio, Jermy; southwest Texas, Nealley in 1890; without locality, Wright 792. 6. Panicum firmulum sp. nov. DESCRIPTION. Plants light olive green, rather loosely tufted, ascending or decumbent at base, from creeping knotted rootstocks as much as 5 cm. long; culms 30 to 40 cm. high, simple or with a few appressed branches, glabrous, the nodes glabrous or strigose; sheaths overlapping, striate, papillose-pubes- cent, papillose only, or nearly gla- brous, a tuft of stiff hairs 3 mm. long on the sides at the summit; ligule dense, about 1.5 mm. long; blades ascending or spreading, firm,4 to 10 cm. long, the lower shorter and more spreading, 4 to 7 mm. wide, ab- ruptly acuminate, rounded at the base Fic. 9.—P. firmulum. From type specimen. and wider than the sheath, sparsely papillose-ciliate, at least toward the base, scabrous on the upper surface; panicles slender, interrupted, their brancheserect, 28 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. the branchlets bearing 1 to 3 short-pediceled spikelets, the setiform prolongation of the axis usually about as long as the spikelets, sometimes twice as long; spikelets 3 to 3.2 mm. long, 1.7 to 1.8 mm. wide, obo- vate, subacute, turgid, strongly nerved ; first glume clasping, half the length of the spikelet, pointed, 5 to 7- nerved; second glume and _ sterile lemma subequal, scarcely covering the fruit, 5 to 7-nerved, the glume ob- scurely reticulated toward the summit; fruit 2.7 to 2.8 mm. long, 1.6 to 1.7mm. wide, obovate-elliptic, abruptly acute, very turgid. Type U.S. National Herbarium no. 592755, collected May 27, 1904, Elsordo, Zapata County, Texas, by David Griffiths (no. 6446). This species resembles P. ramisetum, from which it differs in the larger spikelets, usually longer setz, broader, more or less ciliate blades, and markedly knotty rootstock. Fig. 10.—Distribution of P. frmulum. DISTRIBUTION. Sandy prairies, southern Texas. Texas: Elsordo, Griffiths 6446; Sarita, Hitchcock 3866; without locality, Nealley. TRUE PANICUM. SYNOPSIS OF GROUPS. Inflorescence consisting of several spike-like racemes along a main axis; fruit transversely rugose. Perennials; culms spreading or creeping; spikelets gla- TOUR LSeO: ace asecet en = Oe eee rere eeeeeee GEMINATA (p. 30). Annuals; spikelets pubescent or glabrous.............--- FAScICcULATA (p. 35). Inflorescence an open or contracted panicle, or if with race- mose branches fruit not transversely rugose. Annuals; panicles open, usually diffuse. (See also Laxa and P. costaricense no. 76.) * Spikelets glabrous, not warty nor rugulose; fruit pol- ished. First glume less than one-fourth the length of the spikelet, obtuse or truncate; sheaths glab- rous except in P. bartowense.............- DICHOTOMIFLORA~ . 47). First glume nearly half the length of the spikelet cen or more; sheaths hispid (sometimes glab- rous in P. decolorans and P. stramineum)..CAPILLARIA (p. 54). Spikelets warty, rugulose, or hispid; fruit not polished, margins of lemma not inrolled. Spikelets 2mm. or more long; not tropical species. VERRUCOSA (p. 126). Spikelets not over 1.4mm. long; tropical species. TRicHorprIA (p. 129). Perennials (two species in Laxa annuals). Fruit transversely rugose (very faintly so in P. ple- num); spikelets ellipsoid, glabrous; plants MODE. eetene ete casc.. Greate eins see smbiee eeeer Maxma (p. 78). Fruit not transversely rugose. Spikelets densely silky-villous, 6 to 7 mm. long; lemma silky on the margins.............. URVILLEANA (p. 182). —— HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 29 Spikelets not silky-villous. Panicles more or less diffuse (somewhat con- tracted in P. hirsutum and P. gowini); the spikelets not short-pediceled along raceme-like branches. Spikelets pointed, glabrous; culms te- rete. Rootstocks wanting; sheaths usually Ree os oo eo ae aes t Drerusa (p. 71). ~ Rootstocks present; sheaths gla- CES penn aencaver eaters pane ae Vireata (p. 84). ~ Spikelets obtuse and glabrous or pointed and sparsely hispid........----.-.See ungrouped tropical species, nos. 76 to 81. Panicles more or less contracted, or the spikelets short-pediceled along the main branches. First glume usually about one-fifth the length of the rounded-obtuse spikelets; tropical species...-.--. PARVIGLUMIA (p. 124). First glume usually more than one-third the length of the acute spikelets (subobtuse in P. stenodes and cer- tain species of Laxa). Culms erect or stiffly ascending, not geniculate; sterile palea not enlarged at maturity; panicle branches not conspicuously raceme-like; blades linear. Panicles few-flowered; con- tracted; sterile palea not . enlarged at maturity ....TENERA (p. 97). Panicles many-flowered, open or contracted; the short- pediceled, pointed spike- lets often secund...-.-..-- AGROSTOIDIA (p. 99). Culms often decumbent or more or less geniculate, if stiff and erect the sterile palea enlarged at maturity; panicle branches raceme-like (except in P. hians and P. exiguiflorum). Second glume and sterile lemma boat-shaped or the latter - bearing two crateriform glands; spikelets glabrous or pubescent...........- SroLoNnIFERA (p. 120). Second glume and sterile lemma not boat-shaped (or the glume butslightly so) nor gland-bearing; spikelets glabrous or scabrous at the apex only.....-..... Laxa (p. 110). 30 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Geminata.—Perennials; culms tall, spreading or creeping; inflorescence consisting of several erect, spike-like racemes distributed along an elongated axis; spikelets secund, glabrous, fruit more or less transversely rugose or roughened. Growing in water or wet places. Au forotaicp] s1es 1x0 (so) Ae baie nAaes eae Smee Fa 7S: ORR A Donates Byte aoa pep 9. P. barbinode. Nodes glabrous. Spikelets 3 mm. long; glumes and sterile lemma papery... -- 8. P. paludivagum. Spikelets not over 2.4mm. long; glumes and sterile lemma MOU DADO: sce oe aaa rit eRe ie cow ete hee eee 7. P. geminatum. 7. Panicum geminatum Forsk. Panicum geminatum Forsk. Fl. Aegypt. Arab. 18. 1775. ‘‘Rosettae in pratis ad littora Nili.’”’? We have not seen the type of this, but the description applies to the American plant which appears to be the same as the Old World species. We are informed by Mr. A. B. Rendle that the type is not in the herbarium of the British Museum. Paspalum appressum Lam. Tabl. Encycl. 1: 176. 1791. ‘‘Ex America merid. Com. D. Richard.’’ We have not seen the type specimen. Digitaria appressa Pers. Syn. 1: 85. 1805. Based on Paspalum appressum Lam. ? Digitaria affinis Roem. & Schult. Syst. Veg. 2: 470.1817. ‘“‘In Santa Fé de Bogota. Ab amiciss. Zea nobiscum communicata.’’ This specimen has not been examined, but the form to which Nees@ and Trinius refer Roemer and Schultes’s name, as shown by a specimen from Bahia in the Trinius Herbarium sent by Nees, is the one here described. Roemer and Schultes’s description leaves the species in doubt, however, since it would only apply to an unusually small specimen and may possibly refer to a different species from that to which Nees and Trinius applied the : name. Panicum beckmanniaeforme Mikan; Trin. in Spreng. Neu. Entd. 2:83. 1821. ‘‘ Hab. — : in Brasilia.’? A portion of the type is in the Trinius Herbarium. It is from Brazil sent by Mikan. Panicum truncatum Trin. Gram. Pan. 130. 1826. The author states he has seen specimens from “Ind. or., Egypt (LinpLey. Steper in hb. Maur. I. I. no. 28.) e Brasil.—s. nom. Panic. affine Schult.—N. aB Esens.”’ The specimen figured in the Icones ® is from Egypt. The specimen has not been examined, but the plate shows the species to be the same as the above-mentioned specimen sent by Nees under the name of P. affine Schult. ? Panicum affine Nees, Agrost. Bras. 113. 1829. Among several other names “Digitaria afinis R. & Sch.” is cited as a synonym. Since this specific name is retained, this is considered as based on Digitaria affinis Roem. & Schult., though Nees’s description is evidently based on the specimen first cited, one collected by Martius in Bahia, Brazil, and preserved in the Munich Herbarium, and belonging to the species here described. ' Panicum brizaeforme Presl, Rel. Haenk. 1: 302. 1830. ‘Hab. in Luzonia.” ,The type, in the herbarium of the German University at Prague, is labeled ‘‘Acapulco. : Haenke.’’ The specimen in the Bernhardi Herbarium at the Missouri Botanical Garden, which is the same species, is from Luzon.¢ Panicum carnosum Salzm.; Steud. Syn. Pl. Glum. 1: 60. 1854. This is mentioned as a synonym under P. paspaloides, with the citation ‘‘Hrbr. Bahia.’’ There is a specimen of this collection in the Trinius Herbarium and another in the United States National Herbarium. a Panicum affine Nees, Agrost. Bras. 113. 1829. 6 Trin. Gram. Icon. 2: pl. 168. 1829. ¢ Scribner, Rep. Mo. Bot. Gard. 10: 46. 1899. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 31 Panicum glomeratum Buckl. Prel. Rep. Geol. Agr. Surv. Tex. App. 3. 1866, not Moench, 1794. ‘‘Western Texas.’’ The type, in the herbarium of the Philadelphia Academy, is a single culm; the turgid spikelets are 2.3 mm. long. Panicum appressum Lam.; Doell in Mart. Fl. Bras. 27: 184. 1877, not Forsk. 1775. Based on Paspalum appressum Lam. This species has usually been called P. paspalodes Pers. The latter, however, is based on P. brizoides Lam.,> not L.¢ The published locality for Panicum brizoides Lam. is “‘India.’’ The type, in the Paris Herbarium, is labeled “‘herb. certo i. de France [Mauritius] Commerson.’’ It belongs to the species described by Hooker @ as P. punctatum Burm.,¢ to which, however, Burmann’s description does not well apply. DESCRIPTION. Plants glabrous throughout; culms cespitose, usually numerous, 25 to 80 cm. high, spreading from a more or less decumbent base, scarcely succulent; sheaths usually overlapping, rather close; ligule ciliate, 1 mm. long; blades 10 to 20 cm. long, 3 to 6 mm. wide, rather stiffly spreading or erect, flat, or involute toward apex, somewhat scabrous on the upper surface; panicle short-exserted or included at the base, 12 to 30 cm. long; axis angled, smooth except toward the summit; racemes 12 to 18, erect or narrowly ascend- ing, the lower rarely distant more than their own length, gradually approximate, the lower 2.5 to 3 cm. long, gradually shorter upward, the axis usually ending in a more or less well-marked, pointed prolongation; spikelets subsessile, 2.2 to 2.4 mm. long, 1.4 mm. wide, turgid, straptly and minutely pointed; first glume about one-third the length of the spikelet, truncate or obtuse; second glume nearly as long as fruit (exceeded only by the point of latter) 4-nerved; sterile lemma 5-nerved, abruptly pointed, equaling the fruit and like the second glume very faintly reticulate toward the summit, inclosing a hyaline palea and usually an abor- tive staminate flower; fruit 2.2 mm. long, 1.2 mm. wide, elliptic, abruptly pointed, strongly transversely rugose. In many of the specimens cited below the base is lacking. Other specimens show a cespitose base with fibrous roots, and a single specimen from Cuba (J//itchcock 142) shows in addition to the cespitose base long slender stolons. The specimen grew in moist soil and the stolons extended over the mud, rooting at the nodes and sending up vertical shoots. These stolons appear very different from the succulent submerged bases of P. paludivagum. Fic. 11.—P.geminatum. From typespecimen of P. brizaeforme Presl. DISTRIBUTION. Moist ground, mostly near the coast, southern Florida and Texas, south through Mexico and the West Indies to Brazil and Peru; also in warmer parts of the Old World. Fioria: Manatee, Tracy 7381; Key Largo, Curtiss 3601*; Key West, Blodgett, Hitchcock 613; Rugel 123. a Syn. Pl. 1: 81. 1805. bTabl. Encycl. 1: 170. 1791. Persoon quotes Lamarck’s description. ¢Mant. Pl. 2: 184.1771. Thisis Echinochloa colona (L.) Link. dF), Brit. Ind. 7: 29. 1896. € Fl. Ind. 26. 1768. 32 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Texas: Dallas, Reverchon 1078; Giddings, Hall 823; Houston, Thurow in 1898; Pierce, Tracy 7392; Spofford, Griffiths 6306; without locality Nealley in 1884. Mexico: Lower California, El Taste, Brandegee in 1893; San José del Cabo, Bran- degee 14, 36; Guaymas, Palmer BPs in 1887; ean River, Palmer 15 in 1869; Acapulco, Palmer 289 in 1894; Yucatan, Izamal, Gawmer 1027 (itcheook Herb.) GUATEMALA: San José, Kellerman 6250. Banamas: Nassau, Curtiss 175. Cusa: Santiago de las Vegas, Hitchcock 143; Habana, Leon 918; Batabano, Hitchcock 142; Hanabana, Wright 761; Guana- bacoa, Leén 920; Guantanamo, Britton 2281. Jamaica: Gordon Town, Hart 806. Porto Rico: Guanica, Sintenis 3367. DanisH West Inpviz&s: St. Croix, Ricksecker 212; St. Thomas, Eggers in 1882. LEEWARD IsLANnDs: Guadeloupe, Duss 2690, 3584, L’ Herminier. WINDWARD IsLaNnps: Martinique, Duss 1293. CoLoMBIA: Santa Marta, Smith 151. British Guiana: Jenman 3969, 4488, 6022. DutcH Guiana: ‘‘Surinam” Hering (Acad. Phil. Herb.). Brazi: Bahia, Salzmann. Peru: Lima, Wilkes Expl. Exped. GaALAPaGcos Istanps: Snodgrass & Heller 557, 746 (Gray Herb.). Fig. 12.—Distribution of P. geminatum. 5 8. Panicum paludivagum sp. nov. DESCRIPTION. Plants apparently perennial, glabrous throughout; culms elongated, from a long, creeping base, rooting at the nodes, rather soft and succulent, as much as 2 meters long, the lower submerged portion loosely branching, the internodes, except the uppermost, some- what swollen, the nodes constricted and often dark colored; sheaths papery and more or less inflated, especially the lower, mostly deciduous from the submerged portion; ligule ciliate, about 1 mm. long; blades 15 to 40 cm. long, conduplicate at base, flat above, long-acumi- nate, rather lax, very scabrous on the upper surface, smooth below, the lower much reduced or rudimentary; panicle usually overtopped or Fig. 13.—P. paludivagum. Fromtype ©dualed by the leaves, 25 to 35 cm. long; axis specimen. angled, smooth; racemes 12 to 15, erect, the lower distant, 3 cm. long, the upper gradually approximate and shorter, the axis ending in a rudimentary spikelet or sometimes in a short, slender-pointed prolongation; spikelets appressed to the angled rachis, 2.8 to 3 mm. long, 1.4 to 1.6 mm. wide, narrowly ovate, not turgid, subacute; first glume about one-fifth the length of the spikelet, nerveless, erose-truncate; second glume half to two-thirds the length of the spikelet, faintly 3-nerved; sterile lemma as long as the fruit, very faintly nerved toward the summit, inclosing a palea of equal length and a staminate flower; fruit 2.8 to 3 mm. long, 1.2 to 1.3 mm. wide, narrowly ovate, acute, very obscurely rugose, the margins scarcely inrolled. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 33 Type U.S. National Herbarium no. 207685, collected May 16 to 31, 1894, ‘‘in water,” in the vicinity of Eustis, Lake County, Florida, by George V. Nash (no. 746). This species differs from P. geminatum in the succulent stems, the lower part submerged, branching and rooting at the nodes, the loose papery sheaths, the lower nearly bladeless, the elongated aerial blades, and the longer, not turgid spikelets, scarcely nerved glumes, shorter second glume, and nearly smooth fruit. This species has usually been distributed as Panicum paspalodes Pers. It is closely related to the species described by Hooker as P. punctatum Burm.¢ but differs espe- cially in the papery, nearly nerveless glumes and sterile lemma and the nearly smooth fruit. The preceding species, P. geminatum, grows along the seacoast, while this species appears to be an inhabitant of fresh-water lakes and rivers. Lake Amatitlan, the cited locality for the Guatemala specimens, lies at an altitude of 1,200 meters. Pringle’s no. 9556 is labeled ‘‘In water, Valley of Zamora, 5000 ft.,’’ and his no. 3336, ‘‘Shallows of Lake Patzcuaro.’’ DISTRIBUTION. Growing more or less submerged in fresh-water rivers and lakes of the interior at least up to 1,600 meters altitude, in Florida, Texas, Mexico, and Central America; also in Uruguay. Fioripa: Grasmere, Combs 760, 1052; Eustis, Nash 746; Man- atee, Tracy 7412; Braiden- town, Combs 1253; Little River, Garber in 1877. Texas: Without locality, Nealley in 1888. Mexico: Guadalajara, Palmer 429 in 1886; State of Michoacan, Valley of Zamora, Pringle 9556; Lake Patzcuaro, Prin- Fig. 14.—Distribution of P. paludivagum. gle 3336. GuaTeMALA: Amatitlan, Tuerckheim 8790; Lake Amatitlan, Kellerman 6253, 6254, Pittier 101. UruGuay: Montevideo, Arechavaleta. 9. Panicum barbinode Trin. _ Panicum purpurascens Raddi, Agrost. Bras. 47. 1823, not H. B. K. 1815. Raddi states that this species is cultivated ‘“‘in Provincia Rio janeiro,’’ and also grows spon- taneously. We have not seen the type, but Raddi’s description applies well to P. barbinode. Panicum barbinode Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 256. 1834. Trinius cites ‘‘ Panicum barbinode Trin. ic. gr. XX VII. tab. 318,’’ then unpublished, and states that his specimen is from Brazil. In the Icones ® the habitat isgiven as Bahia. This specimen, in the Trinius Herbarium, which is the type, is labeled “Bahia, Riedel 1831.”’ Panicum guadaloupense Steud. Syn. Pl. Glum. 1: 61. 1854. ‘Ins. Guadaloupe.”’ We have not seen the type, but the description applies well to P. barbinode Trin. Panicum equinum Salzm.; Steud. Syn. Pl. Glum. 1: 67. 1854. ‘“‘Salzmann in Bahia” is the first specimen cited. The specimen in the De Candolle Herbarium labeled ‘‘P. equinum Salz., Bahia, Salzm.”’ is probably the type. ' @ See footnote d, page 31, [P. geminatum] and the paragraph to which itis appended, b Gram. Icon. 3: pl. 318. 1836. 41616°—vo1 15—10——3 34 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum pictigluma Steud. Syn. Pl. Glum. 1: 73. 1854. “Brasil.’? We have not seen the type of this, but Steudel cites P. purpurascens Raddi as a synonym and his description applies well to P. barbinode. Panicum paraguayense Steud.; Doell in Mart. Fl. Bras. 2?: 189. 1877. This is given as a synonym under P. numidianum Lam., and credited to ‘‘Steudel in planta- rum Renggeri schedulis.’’? We have not seen the type. The name P. numidianum Lam. was taken up as the earliest one for this species by Nees,@ Doell,® Hitchcock, ¢ and others, but the type specimen of P. numidianum, ‘“Ex numidia,’’? in the Lamarck Herbarium, does not agree in all respects with the type of P. barbi- node. The lower glume is longer and is 3-nerved instead of 1-nerved, the pedicels of the stalked spikelets are longer, and the rachis lacks the long hairs of P. barbinode. Panicum muticum Forsk.¢ is accepted for this species by Hooker ¢ and others, but the identity of Forskal’s species is uncertain, as we have not seen the type and the description is insufficient to identify it. Forsk&l’s plant was collected at Rosetta and is said to be allied to Panicum colonum. We are informed by Mr. A. B. Rendle that the type is not in the herbarium of the British Museum. Recent American authors / have applied the name P. molle Swartz to this species, but an examination of Swartz’s type g shows it to belong to a very different species. Fia. 15.—P. barbinode. From type specimen. . DESCRIPTION. Plants perennial, sending out widely creeping stolons; culms decumbent at base, rooting at the lower nodes, 2 to 5 or 6 meters high, or higher in cultivation, robust, simple, or producing leafy shoots only, glabrous, the nodes densely villous; sheaths softly or harshly villous to merely papillose or even glabrous toward the summit, densely pubescent at the juncture with the blades; ligules membranaceous, densely ciliate, about 1 mm. long; blades ascending or spreading, 10 to 30 cm. long, 10 to 15 mm. wide, rounded at the base, glabrous on both surfaces, the margin scabrous; panicle 12 to 20 cm. long, about half as wide, the rather distant, subracemose, densely flowered branches ascending or spreading, the main axis and the somewhat flattened branches scabrous on the edges, densely pubescent in the axils, a few stiff hairs on the very short pedicels; spikelets 3 mm. long, 1.3 mm. wide, elliptic; first glume about one-fourth the length of the spikelet, 1-nerved, acute; second glume and sterile lemma subequal, both exceeded by the sterile palea; fruit about 2.5 mm. long, 1.1 mm, wide, obtuse, minutely transversely rugose. This species, commercially known as “‘ Para grass,’’ is cultivated in South America, the West Indies, and Mexico, and has been introduced into the Gulf States. @ Agrost. Bras. 122. 1829. b Mart. Fl. Bras. 27: 188. 1877. ¢Contr. Nat. Herb. 12: 224. 1909. dF. Aegypt. Arab. 20. 1775. éFl. Brit. Ind. 7: 34. 1896. f/Scribner, U. 8S. Dept. Agr. Div. Agrost. Bull. 14: 54. 1900; Nash in Small, Fl. Southeast. U. 8. 90. 1903. 9See P. molle Swartz, page 42; and for a full discussion of Swartz’s types, see Hitchcock, Contr. Nat. Herb. 12: 135-143. 1908. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 35 DISTRIBUTION. Cultivated and waste ground, escaped from cultivation, Florida to Texas, and throughout subtropical and tropical America; native of Brazil. Frioria: Merrimack, Baker 48; Braidentown, Combs 1265, 1311, Tracy 7763. AtaBAMA: Mobile, Mohr. in 1880. Texas: Without locality, Nealley. Mexico: Cuantla, Holway 3045; Manzanillo, Palmer 1078 in 1890; Colima, Em- , rick 8; Lower California, Brandegee 46. GuaTeMALA: Alta Vera Paz, Tuerckheim 7799, 8617; Mazatenango, Mazondé Hay 3476. _Sarvapor: Without locality, Renson 214. NicaraGcua: Chinandega, Baker 2053. Costa Rica: Térraba, Pittier 412; border of Rio Tufs, Tonduz 11393. Bawamas: Nassau, Curtiss W. Ind. Pl. 115. Cusa: Puentes Grandes, Leén 283; near Cienfuegos, Pringle 26, Habana, Leén 568; Romelie, Eggers 4870. Porto Rico: Bayamon, Heller 100, Millspaugh 324; Guanica, Millspaugh 727; Yauco, Heller 6293; Los Mameges, Eggers 1328. DanisH West Inp1Es: St. Croix, Ricksecker 300. Leewarp Istanps: Guadeloupe, Duss 2689, L’ Herminier in part. Wrypwarp Istanps: Martinique, Duss 539; Granada, Broadway in 1904 and 1905. CotomBiA: Santa Marta, Smith 211. British Guiana: Jenman 5998. Braziu: Bahia, Salzmann; Campinas, Novaes 1242; Sao Sebastiao, Léfgren 3142; without locality, Riedel. Paraauay: Morong 779a. Ecuapor: Lehmann 5744. This species occurs in the warmer parts of the Old World, where it was probably introduced from South America. Hooker @ states that it is ‘cultivated or naturalized” in Bengaland Ceylon. Trimen? makes the following statement concerning it: “A well-known fodder-grass in Ceylon, but there is no record of its introduc- tion into the island. According to Roxburgh seeds were received at the Calcutta Botanical Garden fromSumatra in 1804, through Dr. Charles Campbell. As it is a native of tropical America, the Dutch, who then held ports in Sumatra, may have imported it from | Surinam.” Durand and Schinz ¢ state Fig. 16.—Distribution of P. barbinode. concerning P. barbinode, which is re- ferred to P. molle Swartz, ‘Maurice, Seychelles.—Distrib.: Originaire des Indes occid. et abondamment répandu maintenant dans l’ancien monde (Baker).”’ Fasciculata.—Annuals with flat, usually rather wide blades; ligules ciliate or mem- branaceous-ciliate, not over 1 mm. long; inflorescence of several narrow or spike-like racemes along a main axis; second glume and sterile lemma usually more or less reticulate-veined, at least toward the apex, the lemma, excepting in occasional specimens of P. molle, inclosing a palea of nearly equal length . and often a staminate flower; fruit transversely rugose. @ FI. Brit. Ind. 7: 35.1896. Hooker gives here several synonyms based upon Asiatic specimens, which we have not examined. 5 Fl. Ceylon 5: 140. 1900. ¢Consp. Fl. Afr. 5: 755. 1895. 36 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Spikelets 5}. to 6mm. long. :.: 4.050 ee seb eet - Cees pe ce 15. P. texanum, Spikelets 2 to 4 mm. long. Spikelets strongly reticulate-veined, 2 to 3 mm. long; glabrous. Panicle branches long and spreading; blades pubes- Gent ar SbanbOlee see eee eg eer ate = og ll. P. fasciculatum. Panicle branches short, appressed; blades narrow, WEWONCONG SS Serer te are apes ae ary one ate lla. P. fasciculatum chartaginense. Spikelets scarcely reticulate-veined or only near apex. Spikelets not over 2 mm. long, glabrous. ..........- 10. P. reptans. Spikelets over 3 mm. long, pubescent. Rachis scabrous but not bristly.....-......--.-.- 13. P. adspersum. Rachis pilose with bristly hairs. Plant more or less velvety, sheaths not PR Se eae Peer oe oe ine 12. P. molle. Plant not velvety; sheaths papillose........ 14. P. arizonicum. 10. Panicum reptans L. Panicum reptans L. Syst. Nat. ed. 10. 2: 870. 1759. No locality is here given, but the same specimen is described more fully by Linneeus, under the name Panicum grossarium, later in his list of Jamaica plants.¢ The type specimen, in the Linnzan Herbarium, is marked ‘‘Br” [for Browne who sent the plant] and on the sheet the word ‘‘reptans” was written and then crossed out. A full discussion of the type of this and P. grossarium is given in another place. Panicum grossarium L. Syst. Nat. ed. 10. 2: 871. 1759. The type specimen is the same as that of P. reptans. Panicum prostratum Lam. Tabl. Encycl. 1: 171. 1791. ‘‘ Ex Insulis Caribeeis.’’ The type, in the Lamarck Herbarium, is from Santo Domingo. Panicum caespitosum Swartz, Fl. Ind. Occ. 1: 146.1797. ‘Habitat in pascuis siccioribus Jamaicae.’’ The type,¢ in the Swartz Herbarium, is from ‘Jamaica, Swartz.”’ Panicum insularum Steud. Syn. Pl. Glum. 1: 61. 1854. ‘‘Ins. Antillae minores.’’ The type, in the Steudel Herbarium, is labeled ‘‘Panicum insularum Steud. An- tillae minores. Hohenacker.”’ Bracharia prostrata Griseb. Abh. Ges. Wiss. Géttingen 7: 263.1857. Based on Panicum prostratum Lam. Panicum aurelianum Hale in Wood, Class-book ed. 3. 787.1861. ‘‘Damp soils, about N. Orleans (Hale).’’ We have not been able to locate the type of this, but in the herbarium of the Missouri Botanical Garden there is a specimen of P. reptans bearing a ticket reading ‘‘ Panicum aurelianum New Orleans Dr. Hale.’’ This agrees perfectly with Hale’s description. Panicum prostratum pilosatum] Eggers, Fl. St. Croix & Virgin Isl. 104. 1879. “‘St. Croix (La Grange).’’ We have not seen the type. The description applies to the common form of P. reptans with pilose rachises. a@ Amoen. Acad. 5: 392. 1759. 6 Hitchcock, Contr. Nat. Herb. 12: 119. 1908. ¢ For an account of Swartz’s American grasses, which are preserved in the Natural History Museum at Stockholm, see Hitchcock, Contr. Nat. Herb. 12: 138. 1908. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 37 * DESCRIPTION, Plants spreading, usually prostrate, or with a decumbent base, rooting at the lower nodes; culms slender, usually freely branching, ascending 10 to 30 cm. above the decumbent or creeping base, glabrous, the nodes usually puberulent; sheaths loose, glabrous, densely ciliate, shorter than the inter- nodes; ligule a dense ring of hairs 1 mm. long or less; blades lanceolate or ovate-lanceolate, 1.5 to 6 cm. long, 4 to 12 mm. wide, cordate, glabrous or puberulent on both surfaces, the white, undulate margin hispid- scabrous, ciliate at base with long, stiff hairs; inflores- cence finally long-exserted, 2 to 6 cm. long, consist- Ce tatenn Haws type ing of 3 to 12 spike-like, ascending or spreading racemes specimen of P. prostratum Lam. ®franged along a main axis; racemes solitary or some- times somewhat fascicled, the upper approximate and shorter, the lower rather distant and 2 or 3 cm. long; rachises and pedicels scabrous and usually sparsely pilose with long, weak hairs; spikelets borne on one side of the rachis, irregularly and rather densely clustered, on pubescent pedicels 1 mm. or less in length, elliptic, 1.9 to 2 mm. long, 0.9 to 1 mm. wide, acute, glabrous; first glume about one-sixth the length of the spikelet, rounded or truncate; second glume and t sterile lemma slightly exceeding the fruit, strongly 5 to 7-nerved; fruit 1.7 mm. long, ; 0.8 mm. wide, elliptic, apiculate. hq Asa whole this species is very uniform, but the long hairs on the secondary rachises + and pedicels are sometimes wanting. DISTRIBUTION. Moist open ground and a frequent weed in waste places and cultivated soil, Florida to Texas, Mexico, and the West Indies, south to northern South America; also in the . tropical regions of the Eastern Hemi- sphere. Fiormwa: Apalachicola, Biltmore Distr. Chapman Herb. 4276 % (Gray Herb.). ALABAMA: Mobile, Mohr in 1884. LovuisiANA: Pointe a la Hache, Langlois 45 in 1882, 154 in 1883; Burnside, Combs 1431; New Orleans, Ridell in 1840; 4 Algiers, Tracy 1837. $ Texas: Columbia, Bush 266, 1296; Fic. 18.—Distribution of P. reptans. Industry, Wurzlow 6; Houston, Thurow 21 in 1903; Pierce, Tracy 7387; Lynchburg, Joor 39 in 1884; without } locality, Nealley in 1884 and 1888. . Mexico: Cuicatlan, Nelson 1622. x Cusa: Habana, Curtiss 691, Hitchcock 146, Leén 276, 292, 297, 566, 576, 906, 910c; , Madruga, Curtiss 536; San Antonio, Hitchcock 145; Herradura, Tracy 9103; a Cienfuegos, Pringle 73; Baracoa, Pollard, Palmer & Palmer 19; La Magdalena, ‘ Earle & Baker 2455; Santiago, Leén 910; Guayabal, Leén 910b; without local- : ity, Wright 3857. ' Jamaica: Hope, Harris 6845; Gordon Town, Hart 838. Porto Rico: Guanica, Millspaugh Pl. Utow. 726, Sintenis 3368; Salinas de Cabo, Sintenis 847; Coamo Springs, Goll 662; Ponce, Heller 497. Danisu West Inptes: St. Croix, Ricksecker 77; St. Thomas, Eggers 293 (Hitch- cock Herb.). 38 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Lezwarp Istanps: Guadeloupe, Duss 3529. ‘Winpwarp Istanps: Martinique, Duss 1290. Cotomsta: Santa Marta, Smith 173. VENEZUELA: Island of Margarita, Miller & Johnston 171. British GuIANA: Jenman 6024. 11. Panicum fasciculatum Swartz. Panicum fasciculatum Swartz, Prodr. Veg. Ind. Occ. 22. 1788. ‘‘ Jamaica.” The type, in the Swartz Herbarium, has sparsely papillose-hispid sheaths and spikelets 2.1 to 2.2 mm. long. Panicum fuscum Swartz, Prodr. Veg. Ind. Occ. 23. 1788. ‘‘ Jamaica.” The type, in the Swartz Herbarium, consists of two smaller, more branching plants, with some- what more hispid sheaths and more contracted panicles than in the type of P. fascicu- latum; the spikelets are 2.1 to 2.3 mm. long. Panicum flavescens Swartz, Prodr. Veg. Ind. Occ. 23.1788. ‘‘ Jamaica.” The type, in the Swartz Herbarium, consists of the upper portion of two culms with panicles somewhat more open than those of the type of P. fasciculatum, but otherwise very like that; the spikelets are 2.2 to 2.3mm. long. This is not the species described under this name by Grisebach® and by Hooker, ® which is a species of the section Ptychophyllum. Panicum fusco-rubens Lam. Tabl. Encycl. 1: 171. 1791. ‘‘Ex Ins. Carbiaeis.” The type, in the Paris Herbarium, is a portion of a large plant with a rather open panicle and spikelets 2.5 mm. long. Panicum fastigiatum Poir. in Lam. imnayel: Suppl. 4: 277. 1816. Based on Panicum fascic- ulatum Swartz, the name changed because of P. fasciculatum Lam. 1798. Panicum nigricans Willd.; Spreng. Syst. Veg. : vin 1: 310.1825. This is given asa synonym under sai er aha Bree) \ ae Jasciculatum. The type, in the Willdenow Herbarium, is from ‘‘Amer. Merid.” collected by Humboldt. The spikelets are 2.1 to 2.2 mm. long. Panicum fuscatum Presl; Nees, Agrost. Bras. 152. 1829. This is given as asynonym> under P. fasciculatum ;. We have not seen Presl’s specimen. Panicum spithamaeum Willd.; Nees, Agrost. Bras. 152. 1829. This name is men- tioned in a note under P. fasciculatum. The type, in the Willdenow Herbarium, from Humboldt, is labeled, ‘‘Amer. Merid.” This name is mispelled ‘‘spithamineum ” by Steudel.¢ Panicum illinoniense Desv. Opusc. 91.1831. ‘‘ Habitat in America boreali.” The type, in the Desvaux Herbarium, hears a slip with the name ‘‘ Panicum illinoniense DesV. Op. p. 91,’’ and ‘‘Hab. Carol.’’ The locality, if meant for Carolina, is clearly an error, but there are many errors in the data on the labels of Desvaux’s plants. The specimen is much like the type of P. fuscum. ' Panicum reticulatum Griseb. Abh. Ges. Wiss. Géttingen 7: 264. 1857, not Torrey, 1852. Grisebach states that his specimen was collected by Duchaissing either in the Caribbees or in the Isthmus of Panama. We have not seen the type, but Grisebach later 4 refers this species to P. fuscum Swartz. Panicum fuscum fasciculatum Griseb. Fl. Brit. W. Ind. 547. 1864. Based on P. Jasciculatum Swartz. a Fl. Brit. W. Ind. 547. 1864. ¢ Syn. Pl. Glum. 1: 80. 1854. b Fl. Brit. Ind. 7: 56. 1896. ] } . a + 4 ‘ re ae HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 39 Panicum fasciculatum genuinum Doell in Mart. Fl. Bras. 2?: 204. 1877. Based on P. fasciculatum Swartz. Panicum fasciculatum flavescens Doell in Mart. FT. Bras. 2?: 205.1877. Based on P. flavescens Swartz. Panicum fasciculatum fuscum Doell in Mart. Fl. Bras. 2?: 205. 1877. Based on P. Suscum Swartz. Panicum fasciculatum was described under a phrase name and figured by Sloane, whose type is at the British Museum of Natural History. Kuntze misapplies the name Panicum paniculatum (L.) Kuntze,¢ based on Paspalum paniculatum L., to this species, owing to the fact that Linnzeus erroneously cites Sloane’s plate of Panicum fasciculatum after his description of Paspalum paniculatum, the type of which is in the Linnzan Herbarium.? Nashé also later made the combination Panicum paniculatum on the same grounds. DESCRIPTION. Plants erect or spreading from a decumbent base, the more robust becoming much branched from the lower nodes; culms 30 to 100 cm. or more high, glabrous or scabrous, or sometimes pubescent below the panicle or hispid below the appressed-pubescent nodes; sheaths sometimes shorter, sometimes longer than the internodes, glabrous or more or less papillose-hispid, densely ciliate, pubescent at the juncture with the blades; ligule a dense ring of hairs about 1 mm. long; blades flat, 4 to 30 em. long, 6 to 20 mm. wide, glabrous, usually scabrous above, sometimes sparsely hispid on one or both surfaces, the nerves in the larger blades conspicuous, sometimes appearing somewhat plicate; inflorescence short-exserted or included at base until maturity, consisting of a series of spike-like racemes arranged along a scabrous, sometimes pilose, main axis, 5 to 15 cm. long, the racemes 5 to 10 cm. long, solitary or fascicled, narrowly ascending to somewhat spreading, spikelet-bearing from the base, or naked below, the short-pediceled spikelets approximate or somewhat crowded, borne singly, or two or three together on short branchlets, along the under side of the axis; spikelets bronze to mahogany colored, 2.1 to 2.5 mm. long, in occasional specimens as much as 3 mm. long, obovate, turgid, abruptly short-pointed, glabrous; first glume clasping, about one-third the length of the spikelet, subacute, 5 to 7-nerved; second glume and sterile lemma slightly exceeding the fruit, 9-nerved, faintly to strongly transversely wrinkled between the nerves; fruit 1.9 to 2.3 mm. long, obovate, obscurely apiculate. This species is variable in the amount of pubescence and in the size of the spikelets. Almost all the West Indian specimens cited below have spikelets not over 2.3 mm. long. The greater number of specimens from Mexico and the United States have spikelets 2.5 to 2.8 mm. long, while about half the Central American specimens have the larger spikelets. This difference in size can not be correlated with any other character. f DISTRIBUTION. Moist open ground, often a weed in fields and along roadsides, southern Florida and Texas, southward through Mexico and the West Indies to Brazil and Ecuador. Fiorma: Cape Canaveral, Curtiss 3589; Lastero Bay, Garber 36; Sneeds Island, Tracy 6455; Caxambas Island, Simpson 275; Marco, Hitchcock Lee Co. Pl. 484; Key West, Rugel; without locality, Blodgett. @Voy. Jam. 1: 115. pl. 72. f. 2. 1707. 6b See Hitchcock, Contr. Nat. Herb. 12: 131. 1908, for an account of Sloane’s Ja- maica grasses. * ¢ Rey. Gen. Pl. 3: 363. 1898. 4 See Hitchcock, Contr. Nat. Herb. 12: 116. 1908. € Bull. Torrey Club 30: 381. 1903. 40 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Texas: Robbstown, Griffiths 6508. Mexico: Santa Ana, Griffiths 6857; Guaymas, Palmer 158 and 207 in 1887, Alamos, Palmer 694 in 1890; Hermosillo, Hitehcock 3598; Chihuahua, Palmer la in 1885; Colima, Palmer 19 in 1897; Topolobampo, Palmer 241 in 1897; Culiacan, Palmer 1557 in 1891; Teotaleingo, Liebmann 277; Santa Maria Tlatella, Tiebmann 279; Cordoba, Finck in 1893; State of Chiapas, Nelson 2874, 2958; Merida, Schott 384; Rosario, Rose 1834, 1884. GuaTteMALA: Alta Vera Paz, Goll | 81; Chicacao, Heyde & Lux Fic .20.—Distribution of P. fasciculatum. 6404; Esquintla, J. D. Smith 2233; Dept. Huehuetenango, Seler 2704; Gualan, Deam 6267. Honpuras: San Pedro Sul4, Thieme 195, 5584. Nicaraaua: Flint in 1868. Costa Rica: Puerto Viejo, Biolley 7471; Matina, Pittier 9727; Nicoya, Tonduz 13749; near the Rio Grande, Pittier 2035. Panama: Bocas del Toro, Hart 78. Banamas: Turks Island, Madiana (Gray Herb.). Cusa: San Antonio, Eggers 4875; Cienfuegos, Pringle 74, 124; Santiago de las Vegas, Wilson 593; Habana, Leén 573; Herradura, Tracy 9091; Sancti Spiritus, Leén 916; Santiago, Leén & Boillot 813. Jamaica: Bath, Mazon 2361; Hope Gardens, Mazon 1659; Gordon Town, Hart 785, 840. Harti: Jacquemont (Gray Herb.). Porto Rico: Rio Piedras, Barrett 63, Heller & Heller 135; Culebra, Britton & Wheeler 137; Caguas, Goll 385, 588; Ponce, Heller 6226, 6302; Guinaca, Sintenis 3647; ‘‘Monte Goyo,’’ Sintenis 1901; Aguadilla, Heller 4528; Aibonito, Underwood & Griggs 462; without locality, Underwood & Griggs 824. Danisu West Inpres: St. Croix, Ricksecker 317; St. Thomas, Eggers in 1882. LEEWARD IsLANps: Guadeloupe, Duss 2691, L’ Herminier. Winpwarp Istanpbs: Martinique, Duss 537, 538; Hahn in 1867-1870; Granada, Broadway in 1905. CotomsiA: Santa Marta, H. H. Smith 208; Santa Ana, Pittier 1610. VENEZUELA: Island of Margarita, Miller & Johnston 180. TrrniwaD: Botanic Gardens Herb. 2283, 3192. FrencuH Guiana: Without data (Gray Herb.). Braziu: Piauhy, Gardner 2357. Ecuapor: El Recreo, Eggers 15418, 15834. GALAPAGOS ISLANDS: Agassiz in 1891. lla. Panicum fasciculatum chartaginense (Swartz) Doell. Panicum chartaginense Swartz, Prodr. Veg. Ind. Occ. 22. 1788. ‘‘ America meridio- nalis, Chartagena.’’ The type, in the Swartz Herbarium, is a more or less prostrate- spreading plant, with short, crowded leaves and narrow, compact panicles somewhat included at base. The blades, and especially the sheaths, are hispid; the spikelets are 2.6 to 2.8 mm. long. Panicum reticulatum Torr. in Marey, Expl. Red Riv. 299. 1852. ‘‘Main fork of Red River, [Texas] July.’’ The type, in the Torrey Herbarium, consists of three slender HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 41 plants with papillose-hispid sheaths and blades and contracted panicles; the spikelets are 2.8 mm. long. Panicum fasciculatum c{hlarthaginense Doell in Mart. Fl. Bras. 27: 205. 1877. Based on P. chartaginense Swartz. Panicum fasciculatum reticulatum Beal, Grasses N. Amer. 2: 117. 1896. Based on _ . P. reticulatum Torr. Panicum fuscum reticulatum Scribn. & Merr- U.S. Dept. Agr. Div. Agrost. Cire. 82: 4.1901. Based on P. reticulatum Torr. The specimen here referred to P. chartaginense is P. ramo- sum L.¢ of Asia. DESCRIPTION. Differing from P. fasciculatum in having smaller, more compact panicles, the branches ascending or appressed, narrower blades, usually pubescent on both surfaces, and spikelets 2.6 to 3 mm. long. There are many intermediate specimens. The well-marked form is usually smaller, with appressed branches and blades. It occurs in the drier regions of the Mexican Plateau. Some specimens, such as Griffiths 1545 and 1616, cited under P. fuscum reticulatum by Scribner and Merrill, are nearly glabrous. The following specimens from Texas seem to be intermediate between the species and subspecies. They are mostly large plants with rather open panicles and spikelets about 3 mm. long: Fort Worth, Tracy 8171; Dona, Tracy 8890; College Station, Price in 1895; Abilene, Bentley in 1899; Victoria, Plank 74; San Antonio, Havard, Heller 1698, Jermy 202; Dallas, Reverchon 94; Houston, Thurow 16. A closely allied species, P. multiculmum Anders.,¢ from the Galapagos Islands, has been referred to P. chartaginense Swartz by Grisebach.¢@ Fic. 21.—P, fasciculatum chartaginense. From type specimen. DISTRIBUTION. Prairies, fields, and waste ground, Texas and Arizona to Mexico; also in Venezuela (Swartz). Texas: Uvalde, Reverchon 1086; Dallas, Bush 1157; Taylor, Ball in 1901; Abi- lene, Bentley in 1899; Waco, Plank 11; Big Springs, Tracy 8289; Columbia, Bush 270; Manor, Hall 825; San Antonio, Havard in 1882, Hitchcock 162, Jermy 203, Plank 47; Bexar County, Jermy 22; Laredo, Sauvignet in 1892; without locality, Nealley in 1887, Wright 797. New Mexico: Socorro, Plank 38. Arizona: Tucson, Griffiths 1545, 1616,3362, Hitchcock 3495; Prin- gle in 1881; Papayo Reserva- tion, Griffiths 1654; Santa Rita Fig. 22.—Distribution of P. fasciculatum chartaginense. Mountains, Griffiths 7297, 7299. Mexico: Chihuahua, Pringle 379, 380; San Dieguito, Palmer 152 in 1904; Vic- toria, Palmer 412 in 1907; Guaymas, Hitchcock 3561, Palmer 159 in 1887 in part. @ See page 44. ¢ Vet. Akad. Handl. Stockh. 1853: 133. 1855. » Loc. cit. @ Fl. Brit. W. Ind. 546. 1864. 42 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, 12. Panicum molle Swartz. Panicum molle Swartz, Prodr. Veg. Ind. Occ. 22.1788. ‘‘ India occidentalis.”? The type, in the Swartz Herbarium, has already been discussed. The sterile lemma bears in its axis a well-developed palea. There is some uncertainty as to the original locality of the type specimen of P. molle. It is said by Swartz to come from the West Indies, but we haye seen no other specimens of this species from that region. Panicum velutinosum Nees; Trin. Gram. Pan. 144. 1826. This is given as a syn- onym of Panicum petiverit Trin. 8. Trinius’s specimen was from “‘ Brasil,’’ commu- nicated by ‘‘N. as Esens.’’ This is in the Trinius Herbarium and is the original of the plate? of P. velutinosum in the Icones. The spikelets differ from those of the type of P. molle in that the palea of the sterile lemma is wanting. Nees¢ later described P. velutinosum, giving the locality as follows: ‘‘ Habitat in sylvaticis prope Villa da Cachoeira, provinciae Bahiensis.’’ [Brazil]. His specimen, of which Trinius’s is evi- dently a duplicate, is in the Munich Herbarium. There are six plants on the sheet, all much smaller than Swartz’s plant, and having shorter, broader blades. The Argentine specimens, which have been referred to P. velutinosum, have a well- developed palea in the sterile floret. Doell@ recognized the two species as distinct chiefly because of this character. More material is needed definitely. to determine whether or not these two forms should be segregated. In Kunth’s Enumeratioé the name is misprinted P. velutinum Nees. DESCRIPTION. Plants ascending or spreading from a decumbent base, usually branching; culms 30 to 70 cm. high, softly pubescent, at least below the pubescent nodes; sheaths usu- ally shorter than the internodes, loose, softly pubescent between the nerves, sometimes obscurely so, densely ciliate; ligule a dense ring of hairs about 1 mm. long; blades ascending or spreading, 4 to 15 cm. long, rarely longer, 7 to 15 mm. wide, rounded at the base, finely and softly pubescent on both surfaces or nearly glabrous on the upper; panicles short-exserted or, especially those of the branches, included at base, 6 to 15 cm. long, the several to many subracemose branches ascending, rarely widely spread- ing at maturity, the main axis and those of the branches densely softly pubescent and also beset with stiff, spreading hairs about 1 mm. long, the short pedicels of the somewhat crowded spikelets similarly hirsute; spikelets 3.4 to 3.8 mm. long, 1.5 to 1.7 mm. wide, obovate, turgid, abruptly pointed, short-attenuate at base, a distinct internode of the rachilla between the first and second glumes; first glume clasping, half as long as the spikelet or more, acute, 5-nerved, the nerves usually anastomosing toward the apex, pilose; second glume and sterile lemma pointed beyond the fruit, 5-nerved, obscurely reticulated between the nerves, pilose, often densely so; fruit 2.6 to 3 mm. long, 1.5 to 1.6 mm. wide, elliptic, apiculate. The Argentine specimens are less velvety and approach in appearance the large forms of P. arizonicum, but these as well as the Mexican specimens lack the papillae commonly present on the sheaths of P. arizonicum. Fig. 23.—P. molle. From type specimen. @ See footnote ¢, p. 36. ¢ Agrost. Bras. 121. 1829. b Trin. Gram. Icon. 2: pl. 180. 1829. d Mart. Fl. Bras. 2?: 187. 1877. é¢ Enum. Pl. 1: 92. 1833. a ; J HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 43 DISTRIBUTION. River banks and moist places, Mexico to Argentina. Mexico: Colima, Palmer 149 in 1897; Lodiego near Culiacdn, Palmer 1660 in 1891; Saltillo, Brandegee 17 in 1893 (Univ. Cal. Herb.); Yucatan, Schott 592 (Field Mus. Herb.). GUATEMALA: Agua Caliente, Deam 6143. Braziu: Piauhy, Gardner 2353, 2361; Prov. Cear4, Gardner 1876 (all in Gray Herb.). ARGENTINA: (6rdoba, Stuckert 11719, 56 in Kneucker Gram. Exs. 366. 13. Panicum adspersum Trin. Panicum adspersum Trin. Gram. Pan. 146. 1826. Trinius states as to the origin of his specimen, ‘‘V.sp. Doming. (SPRENGEL, sub nomine Pan. caespitosi.)’’ The type, in the Trinius Herbarium, is labeled, ‘‘ Pani- cum adspersum m. St. Domingense s.[ub] n.[omine] P. caespitosum Lam. (!) mis. cl. Sprengel.’? This specimen was afterwards figured by Trinius.¢ The spikelets are 3.2 mm. long. Panicum thomasianum Steud.; Doell in Mart. Fl. Bras. 27: 188. 1877. This is men- tioned as a synonym under P. adspersum Trin. The type, collected by Duchaissing in St. Thomas, is in the Steudel Herbarium. This species has been referred by many authors to P. grossarium L., but that name isa synonym of P. reptans. Fia. 24.—P. adspersum. From type specimen. DESCRIPTION. Plants light green, glabrous except as noted, ascending or spreading from a decum- bent base, rooting at the lower nodes, commonly rather freely branching; culms 30 to 100 cm. high, compressed; sheaths shorter than the internodes, rather loose, densely ciliate at least toward the summit; ligule a ciliate-membranaceous ring scarcely 1 mm. long; blades ascending or spreading, 5 to 15 cm., rarely as much as 20 cm. long, 8 to 20 mm. wide, abruptly acuminate, sometimes ciliate at the rounded base, scabrous on the margin; panicles rather short-exserted, 6 to 15 cm. long, composed of few to many ascending spike-like racemes, 3 to 10 cm. long, the slender axes angled, scabrous, usually pubescent in the axils, bearing approximate, short-pediceled spike- lets singly or two or three together on short branchlets along the under side; spikelets 3.2 to 4 mm. long, 1.5 to 1.8 mm. wide, fusiform, turgid, abruptly acuminate; first glume clasping, about one-third the length of the spikelet, subacute, 5-nerved, glabrous; second glume and sterile lemma exceeding the fruit and pointed beyond it, 5 to 7-nerved, hispid at least toward the summit, or sometimes hispidulous only, rarely glabrous, sometimes obscurely reticulate; fruit 2.2 to 3 mm. long, obovate, obtuse. This species varies much in size and habit. The Florida specimens are more robust than many of those from the West Indies, including the type specimen. There ap- pears, however, to be no characters by which these can be separated. Some of the Cuban specimens, such as Curtiss 748, are equally robust. In a specimen from St. Croix, Eggers in 1876, the spikelets are strongly papillose-hispid. @4Gram. Icon. (2: pl. 169. 1829. 44 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Moist open ground, Florida and the West Indies, often a weed in pastures and culti- vated fields. It has been collected as a ballast plant by Mohr at Mobile, Alabama, by Scribner at Philadelphia, Pennsylvania, and by Martindale at Camden, New Jersey. Fiorina: St. Augustine, Curtiss 6705, Kearney 176, Ricker 952; Sanibal Island, Simpson 292; Marco, Hitchcock Lee Co. Pl. 485; Miami, Chase 3851, Hitchcock 650; Sand Key, Curtiss 3606** ; Key West, Curtiss 5431, Hitch- cock 611, 618, 620; without lo- cality Chapman. ALABAMA: Mobile, on ballast, Mohr in 1891. Bawamas: Nassau, Curtiss 113. Cusa: Habana, Curtiss 748, Ledn 291, 570; Santiago de las Vegas, Baker & Wilson 512, Hitchcock 147, 148, Tracy 9109; Triscor- nia, Hitchcock 159; Cabanas, Palmer & Riley 746,771; Her- radura, T’racy 9102; Sancti Spiritus, Leén 925; Guines, Ledn 924; without locality, Wright 3869. Jamaica: Without locality, March (Gray Herb.). Porto Rico: Between Coamo and Aibonito, Sintenis 1957. DanisH West Inpiks: St. Croix, Eggers in 1876, Ricksecker 66, 384. LEEWARD IsLANDs: Guadeloupe, Duss 3180. Fig. 25.—Distribution of P. adspersum. Panicum RAMosuM L. Mant. Pl. 1: 29. 1767, an Asiatic species of this group and somewhat resembling P. adspersum, but with smaller spikelets, having a finely trans- versely rugose sterile lemma, in appearance much like the fertile lemma, was collected on ballast, at Mobile, Ala., Sept. 16, 1891, by Dr. Charles Mohr. This is the specimen referred by Scribner @ to P. chartaginense. 14. Panicum arizonicum Scribn. & Merr. Panicum dissitiflorum Vasey in 8. Wats. Proc. Amer. Acad. 24: 80.1889. This is listed without description, Palmer’s numbers 159 and 190, Guaymas, Mexico, being cited. Two species were distributed under Palmer 159, P. arizonicwm and P. fasciculatum chartaginense, a speci- men of each of which is on the sheet of no. 159 which was in the National Herbarium in the time of Doctor Vasey. The plant of P. arizonicum is taken as the type, since other specimens of this species are named P. dissitiflorum in Vasey’s writing. Panicum fuscum major[us] Vasey, U. S. Dept. Agr. Div. Bot. Bull. 8: 26. 1889. ‘Mexico (Dr. E. Palmer).’’ The type, in the Na- tional Herbarium, is from south- western Chihuahua, collected Au- Fig. 26.—P. arizonicum. From type specimen of P. dissitiflorwm Vaeey, . gust to November, 1885, no. 1 b. It is a robust specimen, 60 cm. high, lacking the base, the blades as much as 15 cm. long and 15 mm. wide, the large panicle 15 cm. long, the sheaths and under surface of the blades papillose-hispid. aU.S8. Dept. Agr. Div. Agrost. Circ. 32: 4. 1901. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 45 Panicum arizonicum Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Cire. 32: 2. 1901. Based on ‘(Panicum (sine nomine) Scribn. Bul. Torr. Bot. Club, 9: 76. 1882; P. fasciculatum dissitiflorum Vasey, in herb. Not P. dissitiflorwm Steud. 1841).” The authors also cite, ‘Type specimen collected on mesas near Camp Lowell, Santa Cruz Valley, Arizona, 465 C. G. Pringle, 1881.’’ As indicated above this species was first mentioned as ‘‘ Panicum (Virgaria) sp.’’ where the specimen referred to is Pringle 465. The same specimen, which is in the National Herbarium, was marked by Doctor Vasey, “‘ Panicum fasciculatum var. dissitiflorum,’’and later by Scribnerand Merrill as the type of P. arizonicum. It is about 60 cm. high, but more slender than Palmer’s specimen mentioned above; the sheaths and blades bear only a few scattered papillz, mostly without hairs. Panicum fasciculatum dissitiflorum Vasey; Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Cire. 32: 2.1901. This herbarium name is given as a synonym of P. arizonicum of which it is a typonym. Panicum arizonicum tenue Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Circ. 32: 3.1901. “Type specimen collected at Fort Huachuca, Arizona, by T. E. Wilcox in 1894.’’ The type, in the National Herbarium, is the small form common in sterile soil. The largest specimen is 17cm. high. Some of the sheaths are sparsely papillose- hispid, some glabrous. Panicum arizonicum laeviglume Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Circ. 32:3.1901. ‘‘Type specimen collected at Mescal, Arizona, 1810 David Griffiths, October, 1900.’’ The type, in the National Herbarium, is a plant about 20 cm. high, with glabrous spikelets and glabrous to sparsely papillose-hispid sheaths. Panicum arizonicum major[us] Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Circ. 32:3.1901. Based on P. fuscum majus Vasey. DESCRIPTION. Plants erect or ascending, sometimes decumbent at base and rooting at the lower nodes, branching at the base and lower nodes; culms 20 to 60 cm. high, glabrous except below the panicle, the nodes sometimes slightly pubescent; sheaths shorter than the internodes or the upper often overlapping, rather loose, glabrous to strongly papillose- hispid; ligule a ring of hairs about 1 mm. long; blades rather thin, ascending or spreading, 5 to 15 cm. long, 6 to 12 mm. wide, rounded at base, glabrous on both surfaces, or scabrous to papillose-hispid beneath, the scabrous, thin, cartilaginous margin usually papillose-ciliate at base; panicles usually long-exserted, 7 to 20 cm. long, the solitary, ascending, slender branches loosely flowered, the spikelets borne on very short, appressed branchlets, the pedicels and axes of branchlets, branches, and the entire panicle finely pubescent and also copiously papillose-hirsute; spikelets 3.5 to 3.8 mm. long, obovate-elliptic, abruptly pointed, attenuate at base as in P. molle, densely hirsute to glabrous; first glume clasping, half the length of the spikelet, acute, 5-nerved ; second glume and sterile lemma pointed beyond the fruit, 5-nerved, the nerves sometimes anastomosing as in P. molle; fruit 2.9 to 3 mm. long, 1.5 to 1.6 mm. wide, obovate-elliptic, apiculate. This species is variable in size and in the amount of pubescence. In cultivated or moist soil it is robust as in the type of the species or of Scribner and Merrill’s subspecies majus. 'Thecommoner form is smaller, more like the type of Scribner and Merrill’s sub- _ species tenue. The form separated by Scribner and Merrill as subspecies laeviglume, because of the glabrous spikelets, appears to have no other distinguishing characters. The following specimens are this form, though in some cases the spikelets are sparsely pubescent or some of the spikelets are glabrous and some are pubescent: Canby 8, Griffiths 1913, 6152, 6168, 6929, 6938, 6939, 6990, Griffiths & Thornber 75, 230, 239, Merton 1694, Metcalfe 768, Pringle 487, Smith in 1896, Wilcox in 1894. 46 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Open sandy or stony ground, western Texas to southern California and northern Mexico. Texas: El Paso, Jones in 1884; Presidio County, Nealley in 1892. New Mexico: Mangas, Smith in 1896, Metcalfe in 1897; Las Cruces, Griffiths 7399 in part; Mogollon Mountains, Metcalfe 768; Sierra County, Metcalfe 1294. Arizona: Tucson Mountains, Grif- jiths 6152, 6938, 6939; Tucson, Griffiths 1596, 3356, 6168, 6737, 7017, Mitchcock 3482; Santa Rita Mountains, Griffiths 5981, 6894, 6990, Grifiths & Thornber Fia. 27.—Distribution of P. arizonicum. 75, 230, 239; Santa Catalina Mountains, Griffiths 7143, 7148, Lemmon 3062; Sasabe, Griffiths 6929;¢ Bowie, Toumey in 1896; Lowell, Pringle 465; Mescal, Griffiths 1810; Fort Huachuca, Wilcox in 1894; Patagonia, Hitchcock 3695; Bisbee, Mearns 1072; San Pedro River, Merton 1694; Cochise, Griffiths 1913; without locality, Lemmon 353. CaLIForNIA: Jamacha, Canby 8 in 1894. Mexico: San José del Cabo, Brandegee 18 in 1890; south of Nogales, Hitchcock 3637; Arroyo San Lazaro, Brandegee in 1902; Guaymas, Hitchcock 3562, Palmer 159 in 1887 in part; Hermosillo, Hitchcock 3542; Nogales, Griffiths 6747, 6759; Loquka, [Lacuca?] Sonora, Griffiths 6891; Topolobampo, Palmer 250 in 1897; State of Chihuahua, Palmer 1 b in 1885, Pringle 487; State of Durango, Rose 2280. 15. Panicum texanum Buckl. Panicum texanum Buckl. Prel. Rep. Geol. Agr. Surv. Tex. App. 3. 1866. ‘‘Austin, Texas.” The type specimen, in the herbarium of the Philadelphia Academy, consists of two overmature plants lacking the base. DESCRIPTION. Plants erect or ascending, often decumbent and rooting at the lower nodes, branching from the base and com- monly from the lower and middle nodes; culms stout, 50 to 150 cm. high, or in robust specimens as much as 3 meters high, softly pubescent at least below the nodes and be- low the panicles; sheaths softly pubescent, often papillose; densely ciliate, the lower shorter than the internodes, the upper usu- ally overlapping; ligules about 1 mm. long; blades ascending or spreading, 8 to 20 cm. long, 7 to 15mm. Fia. 28.—P, texanum. From type specimen. wide, rounded at the base, softly pubescent on both surfaces, often finely papillose, panicles finally exserted, 8 to 20 cm. long, 1 to 3 cm. a HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 47 wide, the main axis much exceeding the erect branches, the axes densely clothed with short pubescence having long, stiff hairs intermixed, the short-pediceled spikelets somewhat crowded; spikelets 5 to 6 mm. long, about 2 mm. wide, fusiform, pointed, short-attenuate at base, pilose; first glume clasping, more than half the length of the spikelet, acute, 3 to 5- nerved; second glume and sterile lemma exceeding the fruit, 5-nerved, often obscurely reticulate; fruit 3.7 to 3.8 mm. long, about 2 mm. wide, elliptic, apiculate. DISTRIBUTION. Prairies and open ground, especially on low land along streams, often a weed in waste ground and cultivated fields, Fig. 29.—Distribution of P. tezanum. Texas and northern Mexico. This is sparingly cultivated under the name of Colorado grass. Texas: Dallas, Reverchon 1226 in Curtiss N. Amer. Pl. 3607A; Corsicana, Rever- chon 2228; Pierce, Tracy 7748; Victoria, Plank 73; Austin, Plank 31; Har- vester, Thurow in 1898; Wallisville, Wallis in 1880; Goliad County, Lea in 1874; San Antonio, Bush 1198, Havard in 1882; Corpus Christi, Hitchcock 163. Mexico: Monterey, Hitchcock 5540. Dichotomiflora.—Annual plants with smooth culms, mostly large, spreading pani- cles, the branchlets short and appressed along the ascending or rarely spread- ing main branches; ligule membranaceous below, densely ciliate above, 1 to 3 mm. long; spikelets glabrous, narrow, acute or acuminate, 2 to 5 mm. long, the first glume one-fifth to one-fourth as long, truncate or with a broadly trian- gular tip; fruit smooth and shining. Panicles narrow, less than 1 cm. wide.............---------- 16..P. vaseyanum. Panicles open, the branches usually ascending. Fruit acuminate; culms with a long, rooting base... .-.- - 19. P. elephantipes. Fruit not acuminate. ‘ Sheaths papillose-hispid.........,..-.-------------- 18. P. bartowense. j ee ee a Ee ee hn i ae 17. P. dichotomiflorum. 16. Panicum vaseyanum Scribn. Panicum vaseyanum Scribn.; Beal, Grasses N. Amer. 2:140. 1896. The only speci- men cited is ‘‘Mexico, Pringle 1415.” The type specimen, in the National Herba- : rium, was collected in the State of Chihuahua, in ‘“‘Wet places, pine plains, base of Sierra Madre,’’ Sep- tember 30, 1887, by C. G. Pringle. DESCRIPTION. Plants spreading, branching at base and at the lower and middle nodes, glabrous throughout; culms 50 to 70 cm. long, somewhat compressed; sheaths shorter than the elongated internodes; ligules 1 to 2mm. long; blades 5 to 20 cm. long, 3 to7 mm. wide, linear, scarcely narrowed at the folded or enveloping base; panicles terminal and from the axils of the upper leaves of the main culms and large branches, narrow, 4 to 7 cm. long, less than 1 cm. wide, partially included, equaled or exceeded by the erect uppermost blade; spikelets short-pediceled, narrowly ovate, 2.5 mm. long, 1.1 to 1.2 mm. wide, subacute; first glume about one- Fig. 30.—P. vaseyanum. From type specimen. 48 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. fifth the length of the spikelet, truncate or obtuse; second glume slightly shorter than the sterile lemma, both 7-nerved, palea of the sterile floret obsolete; fruit 2.1 mm. long, 1 mm. wide, elliptic, apiculate. This very distinct species is known from a single collection only, that distributed by Pringle, mentioned above. 17. Panicum dichotomifiorum Michx. Panicum miliaceum Walt. Fl. Carol. 72. 1788, not L. 1753. Since Walter does not give Linnzeus as authority nor use his diagnosis, this is evidently intended as a new species. No specimen of this is found in Walter’s herbarium,? but the description indicates P. dichotomiflorum, which, together with Walter’s name, Elliott® refers to P. geniculatum Muhl. Panicum dichotomiflorum Michx. Fl. Bor. Amer. 1: 48. 1803. ‘‘ Has. in occi- dentalibus montium Alleghanis.’’ The type is in the herbarium of Drake de Castillo. It was sent by Richard, having been collected by Michaux ‘‘ad occidentum montium Alleghanis.’’ The specimen of this in the Michaux Herbarium is labeled ‘‘in regione Illinoensium.’’ Both of these specimens are the common glabrous form of the United States as represented by Chase in Kneucker, Gram. Exs. no. 546. Panicum geniculatum Muhl. Cat. Pl. 9. 1813. Based on P. dichotomiflorum Michx. The specimen in the Muhlenberg Herbarium is in folio aie marked ‘‘ Panicum geniculatum (dichotomiflorum) M. 114.” Panicum aquaticum Poir. in Lam. Encycl. Suppl. 4: 281. 1816. ‘‘Cette plante croit & Porto-Ricco; elle m’a été communiquée par M. Ledru.’’ The type, in the Cosson Herbarium, is from Porto Rico. In the description the species is compared to P. melicarium Michx., and the label of the type bears the abbreviated statement» ‘aff. P. melicario Mich.’’ In the Desvaux Herbarium there is a similar specimen from the Antilles, labeled ‘“‘P. aquaticum Desv. in Poir. Enc. Suppl.’”’ In both specimens the spikelets are nearly 3 mm. long as in Wright 3861. In the original publication there is no indication that Desvaux is the author of the species. This name was erroneously referred by Hitchcock¢ to P. elephantipes. The fruit is not acuminate as in that species Panicum multiflorum Poir. in Lam. Encycl. Suppl. 4: 282. 1816. ‘“‘Cette plante croit & la Caroline; elle m’a été communiquée par M. Bosc.’’ We take the specimen labeled ‘‘bosc. caroline,’’ in the Cosson Herbarium, to be the type. Another specimen of the same collection is in the Desfontaines Herbarium. It is labeled, ‘“‘Am. Sept. Bosc,’’ and also ‘‘ Panicum brachiatum Bosc.’’ These are the typical form. Panicum brachiatum Bosc; Spreng. Syst. Veg. 1: 321. 1825, not Poir. 1816. The locality given by Sprengel is “‘Jns. Bermud.’’ As indicated above, the specimen of P. multiflorum from Bose is labeled P. brachiatum Bosc. There is also in the Delessert Herbarium a specimen so labeled, collected in South Carolina by Bose. We have seen no specimen of Bosc’s from the Bermudas, the published locality, and we find no record that Bose visited the Bermudas. The meager description applies to P. dichotomiflorum. Panicum chloroticum Nees; Trin. Gram. Pan. 236. 1826. Trinius describes a variety ‘‘e (agreste N. ab Es.)”’ which is the equivalent of the species, and ‘‘ (sylvestre N. ab Es.),”’ both from Brazil, ‘‘V. utriusque spp. Brasil (N. AaB Esens. LANGs- porFF).”’ The latter differs in having a more open panicle, larger spikelets, and narrower leaves. Nees@ described the same species later, with three varieties, @ For an account of Walter’s grasses see Hitchcock, Rep. Mo. Bot. Gard. 16: 31-56. 1905. b Bot. 8S. C. & Ga. 1: 117. 1816. ¢Contr. Nat. Herb. 12: 218. 1909. @ Agrost. Bras. 164. 1829. —_ = = . : TS li Oe ee ¢ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 49 aagreste, the equivalent of the species, ‘‘ Habitat in graminosis cultis prope Soteropo- lin et Oeiras provinciae Bahiensis et Piauhiensis;’’ 8 sylvestre, ‘‘ Habitat in sylvis ad Almada, Ferradas et in via Felisbertia districtus Insulanorum proy. Bahiensis, (Martius et Maximil. Princ. Neovid.);’’ y pingue, ‘‘ Habitat in cultis ad Soteropolin, provinciae Bahiensis.’’ Specimens of none of these could be found in the Trinius Herbarium, and therefore the specimens described by Nees, which are in the Munich Herbarium, may be considered the types. These are all labeled with the published data as given above. The types of agreste and pingue are similar, having broad blades and rather dense panicles of small spikelets 2.2 to 2.3 mm. long as in Chase 4234 from Florida and Morong 543 from Paraguay. The type of variety sylvestre differs in having narrow and shorter blades, smaller, more open panicle, and larger spikelets about 3 mm. long as in Riedel 959 from Brazil. This form may prove to be a distinct species. None of the specimens shows the base of the plant. Kunth@ erroneously refers variety sylvestre to Panicum brachiatum Poir., which is a species of Chaetochloa. Panicum elliottii Trin.; Nees, Agrost. Bras. 170. 1829. This is mentioned as a syn- onym under P. proliferum Lam. which latter name Nees applies to P. dichotomiflorum Michx. The type was not found in the Trinius Herbarium nor at Munich. Panicum retrofractum Delile; Desv. Opusc. 96. 1831. Desvaux gives no locality other than ‘‘America borealis.’’ The type, in the Jussieu Herbarium, is from “Oaroline,’’ and is the typical form. Panicum hygrophilum Salzm.; Steud. Syn. Pl. Glum. 1: 71. 1854. ‘‘Bahia.’’ In the National Herbarium is a specimen labeled P. hygrophilum Salzm. from Bahia, which agrees with Nees’s variety sylvestre. There is a specimen of the same in Van Heurck’s herbarium, where is located the original set of Salzmann, and duplicates in Hackel’s and other European herbaria, but we do not know which specimen was seen by Steudel. Panicum proliferum pilosum Griseb. Cat. Pl. Cub. 232. 1866. ‘‘Wr[ight] a. 1865, ad lagunas.’’ The type, in the Grisebach Herbarium, is labeled, ‘‘ Around lagunas in wet or damp ground, Hanabana,’’ no. 186. This is a small plant with spreading or decumbent culms, papillose-hispid sheaths and blades villous above. Nash’s no. 567 from Eustis, Florida, is similar to this but has somewhat larger spikelets. Panicum proliferum strictum Griseb. Cat. Pl. Cub. 232. 1866. ‘‘Wr[ight] 3456.” The type, Wright’s no. 3456 from Cuba, is in the Grisebach Herbarium. The spikelets are about 3 mm. long, the sheaths smooth, the blades villous above. Panicum proliferum geniculatum Wood, Bot. & Flor. 392. 1874. This is probably based on P. geniculatum Ell., though that name is not mentioned; no locality nor specimen is cited. Vasey makes the same combination, basing it upon P. genicu- latum Ell. Panicum amplectans Chapm. Bot. Gaz. 3:20. 1878. ‘‘South Florida.’’ The type, in the Chapman Herbarium at Biltmore, was collected by Blodgett. Panicum francavillanum Fourn. Mex. Pl. 2: 25. 1886.¢ ‘‘Tacabaya (ScHarrn[ER] @Enum. Pl. 1: 155. 1833. b Grasses U. 8. 12. 1883. ¢ The date given on the title-page of this work is 1886. A set of proof sheets was supplied to Bentham in 1880 and is referred to by the latter author in his paper, Notes on Gramineae, read November 3, 1881, and published in the Journal of the Linnaean Society (Botany 19: 14-134. 1881). Fournier’s names are also cited by Hemsley (Biol. Centr. Amer. 3: 1885), to which work they are referred by the Index Kewensis, but the names are there usually nomina nuda. The proof sheets mentioned above are in the library at Kew, marked, ‘‘Proof sheets of Mr. Fournier Gramineae, 1881. From Mr. Bentham.’’ They are stamped, ‘‘lre Epreuve 18 Mai 1880." Ben- tham says of these (Notes on Gramineae, p. 20), ‘‘Eugéne Fournier’s ‘Enumeration of 41616°—vort 15—10——4 t 50 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. n.301).’’ The type is in the herbarium of Drake de Castillo. The name was earlier mentioned by Hemsley.¢ Panicum proliferum chloroticum Hack. in Repert. Nov. Sp. Fedde 7: 348. 1909. Based on P. chloroticum Nees. This species was referred by Pursh,) as it has been by most later authors, to P. proliferum Lam. The latter is, however, the same as P. miliare Lam., an Old World species. DESCRIPTION. Plants usually freely branching, ascending or spreading from a geniculate base, or sometimes erect, usually smooth throughout, or, in tropical forms, more or less pubes- cent; culms somewhat compressed, often thick and succulent, drying furrowed, usually 50 to 100 cm. long, in robust specimens as much as 2 meters long, the nodes . smooth, at least the lower swollen; sheaths often com- pressed, usually longer than the internodes, ciliate on the margin toward the summit; ligules 1 to 2mm. long; blades flat or in small specimens sometimes folded, glabrous or - sparsely pilose above, 10 to 50 cm. long, 3 to 20 mm. wide, at base about as wide as sheath, the white midnerve usu- ally prominent; panicles terminal and axillary, included at base or tardily short-exserted, many-flowered, 10 to 40 cm. long or more, the main branches ascending, or finally spreading or even reflexed, the short branchlets appressed, Fig. 31.—P. dichotomiflorum. }earing short-pediceled, often rather crowded spikelets, From specimen of 2. genict- +6 axes angled and scabrous; spikelets narrowly oblong- latum Muhl. in Elliott Her- 5 , bart } ovate, 2 to 3.2 mm., usually about 2.5 mm. long, about 0.9 mm. wide, acute, often greenish purple; first glume one- fifth to one-fourth the length of the spikelet, truncate or broadly triangular; second glume and sterile lemma more or less pointed beyond the fruit, rather faintly 7-nerved, the palea of the sterile floret present or wanting; fruit 1.8 to 2 mm. long, about 0.8 mm. wide, elliptic. This species as it occurs in the United States is usually glabrous throughout but varies much in the size of the blades and of the spikelets, the latter varying from 2 to 3.2 mm. in length. Not uncommonly specimens occur with the upper surface of some or all of the blades sparsely or even densely pilose, such as: CoNNECTICUT, Wilson 1248; New Yorx, Young in 1872; PENNSYLVANIA, Heller in 1900; DELAWARE, Commons 230; Kansas, Carleton in 1892; Forma, Chase 4294, Combs 94, 1251. One series of specimens from Florida, Nash 567,¢ is low, 20 to 30 cm. high, with narrow blades pubescent above, and papillose-hispid sheaths. Nash’s no. 372 from the same locality is glabrous throughout, except the ciliate margin of the sheaths, but otherwise is the same as his no. 567. Two Cuban specimens, Hitchcock 149 and Wright 3860, are like Nash’s no. 567. Many of the West Indian specimens have blades pilose above, some of which have spikelets about 2 mm. long and others about 3 mm. long. Such are: Brace 3742, Britton & Cowell 432, Curtiss 177, Duss 3178, Eggers 4405, 4512, Geogr. Soc. Baltimore 489, Hitchcock 150, Wright 3861. The South American specimens cited are glabrous. Those from Arechavaleta and Morong 543 have small spikelets as in the Mexican Gramineae’ is not yet published; but being already printed off and M. Fournier having obligingly supplied me with a copy, I feel bound in so far as I am con- cerned, to tre&t it as having already taken date.’’? The Kew copy ends with page 150 and lacks index, title-page, and plates. @ Biol. Centr. Amer. Bot. 3: 489. 1885. ob Fl. Amer. Sept. 1: 68. 1814. ¢ This number was distributed under an unpublished varietal name. _ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 5} type of P. chloroticum, while Morong 1002 and Riedel 959 have spikelets 3 mm. long, as in the type of P. chloroticum 8 sylvestre. These different variations in pubescence, size of spikelet, and habit can not be in any way correlated and each is connected by intergrading specimens with the typical form. Nor has any variation a separate geographical range, though specimens with pilose blades are commoner in the West Indies than elsewhere. The specimens cited below from Brazil have larger spikelets. They belong to the form mentioned above under P. chloroticum as variety sylvestre. DISTRIBUTION. Moist ground, along streams, and a weed in waste places and cultivated soil, Maine to Nebraska, south to Florida and Texas;,also in California, Mexico, the West In- dies, and South America to Uruguay. Marne: North Berwick, Parlin in 1891 (Gray Herb.). Massacuusetts: Cambridge, Morong in 1876; Newburyport, Leavitt & Eaton in 1902; Plymouth, Oakes. Connecticut: Stamford, Driggs 8; South Glastonbury, Wilson 1248; Bridgeport, Eames in 1895. _ New York: Northville, Young in 1872 (Hitchcock Herb.). New Jersey: Clifton, Nash in 1891; Weehauken, Van Sickle in 1895; Freehold, Pearce in 1884. PENNSYLVANIA: Easton, Porter in 1895; Chambersburg, Porter in 1898; West- moreland County, Pierron in 1877; Lancaster County, Heller in 1900. Onto: Niles, Ingraham in 1891; Sheffield, Ricksecker in 1894. Inprana: Wells County, Deam in 1903; Lafayette, Dorner 23, 86. Inurvotrs: Chicago, Moffatt 374, Umbach in 1896; Pine Rock, Waite in 1885; Peoria, Brendel, McDonald 71; St. Clair County, Eggert 110. Iowa: Mount Pleasant, Mills in 1894; Jefferson, Wilcox 27; Manchester, Ball 39; Murray, Morris A 287; Fayette County, Fink 409. Nesraska: Talmage, Elmore 71; without locality, Holmes in 1889. Missourt: Courtney, Bush 10 in 1892. Kansas: Topeka, Smyth 331; Riley County, Hitchcock 3838, Norton 567, Carleton in 1892. DELAWARE: Wilmington, Commons 229, 230, Canby in 1896. MARYLAND: Garrett County, Smith in 1879. District or CoLtumpBia: Chase in Kneucker Gram. Exs. 546, McCarthy in 1886, Pollard 682, Steele in 1896, Vasey in 1887, Ward in 1876, Williams in 1896. Viremia: Virginia Beach, Hitch- cock 217; Gravelly Run, Ward FiG. 32.—Distribution of P. dichotomiflorum. in 1886. West Virernia: Aurora, Steele in 1898; Tygarts Valley, Smith in 1879. Norra Caroma: Chapel Hill, Ashe; West Raleigh, Stanton 1280; Biltmore, Biltmore Herb. 702a; Magnetic City, Wetherby 19. Soutn Caro.tina: Lexington, Corley in 1879; St. Helena Island, Cuthbert in 1904. Groraia: Dalton, Harper 382; Augusta, Kearney 208. Frorma: Duval County, Fredholm 395; Lake City, Chase 4234, Combs 94; Eustis, Nash 372, 567, 874; Crystal, Combs 990; Manatee, Chapman, C a 1251. Kentucky: Bell County, Kearney 374. TENNESSEE: Knoxville, Ruth 71; Cocke County, Kearney 966; Nashville, Gat- tinger in 1879. 52 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, AtABAMA: Mobile, Mohr in 1878; Deatsville, Pollard & Mazon 307. Mississippi: Starkville, Kearney 9, 18; Nicholson, Kearney 372 in part; Biloxi, Tracy 4618, 6507; Mississippi City, Tracy 77. Lourstana: Calhoun, Ball 67; McCall, Combs 1436; Burnside, Combs 1419; Coushatta, Ball 135; Natchitoches, Ball 139, 164; Rayville, Ball 31, 24; Oberlin, Ball 215, 229; South Pass, Tracy & Lloyd 471; Baton Rouge, Joor 25; Lake Charles, Chase 4392; without locality, Langlois 29. Texas: Houston, Hall817; Waller County, Thurow in 1898 and 1903; Texarkana, Heller 4210, 4246; Clarksville, Plank 8; Llano, Plank 18 in part; Kerrville, Heller 1883 in part; Santa Maria, Nealley in 1889; without locality, Joor, Nealley in 1886. OKLAHOMA: False Washita, Palmer 37p in 1868. New Mexico: Las Cruces, Plank 29. CairorniA: Fresno, Biolett: 140. Mexico: Head of Mazatlan River, Wright 1317 (Gray Herb.). Bermupas: Hamilton, Millspaugh Pl. Utow. 126. Bawamas: Hog Island, Eggers 4405, 4512; Nassau, Curtiss 177; Cat Cay, Brace 3742; Watlings, Geogr. Soc. Baltimore 489. Cusa: Herradura, Tracy 9055, 9342; Santiago de las Vegas, Hitchcock 151; Bata- bano, Hitchcock 150; Guanabacoa, Leén 919; without locality, Wright 3456, 3860 in part, 3861. Porto Rico: Utuado, Britton & Cowell 432. LEEWARD IsLANpDs: Guadeloupe, Duss 3178. Braziu: Bahia, Salzmann; without locality, Riedel 959. Paraauay: Morong 543, 1002 in part. Uruguay: Montevideo, Arechavaleta. 18. Panicum bartowense Scribn. & Merr. Panicum bartowense Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Cire. 35: 3. 1901. ‘‘Type specimen collected in wet, reclaimed swamps at Bartow, Polk County, Fla. No. 1220, Robert Combs, September 29, 1898.”’ The type, in the National Herbarium, is an erect, sim- ple plant about 1.75 meters high, with conspicuously hispid sheaths and nearly glabrous blades, the spikelets about 2.5 mm. long. DESCRIPTION. Plants simple or sparingly branching, as much as 2 meters high, erect; culms glabrous, the larger as much as 7 mm. thick; sheaths mostly longer than the inter- nodes, papillose-hispid; ligules 2 to 3 mm. long, the ciliz more or less segregated in tufts; blades 15 to 40 cm. long, 5 to 13 mm. wide, glabrous or more or less pilose above, rarely sparsely hispid beneath, rather prominently papillose on the margin near the round but scarcely cordate base; panicles large and finally loosely spreading, 15 to 60 em. long, the branches at first ascending, finally spreading, the short branchlets and short-pediceled spikelets appressed as in P. dichotomiflorum; spikelets 2.2 to 2.7 mm. long, the glumes and fruit as in P. dichotomiflorum. This species is closely allied to P. dichotomiflorum, and may be only an extreme form of that species. As limited here, it differs in having tall, erect, simple, or nearly simple culms and papillose-hispid sheaths. The blades are usually pilose above, though the type specimen has nearly glabrous blades, but this is the case Fig. 33.—P. bartowense. From type specimen. SE eE—————————————ee eee ee — elle eee Oe”) CO HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 53 with some of the specimens referred to P. dichotomiflorum. A few specimens men- tioned under the latter species have papillose-hispid sheaths, but are low branching plants with the habit of that species rather than of P.bartowense. Although most of the specimens cited below are erect and simple, one, Chase 3850, is much branched and spreading at the base like P. dichotomiflorum, and it is possible that the erect, simple habit has no special significance as a specific character. DISTRIBUTION. Low ground, often growing in shallow Fig. 34.—Distribution of P. bartowense. water, Florida and the Bahamas. Fioripa: Homosassa, Combs 971; Titusville, Chase 4007; Manatee, Tracy 6691; Braidentown, Tracy 7738; Palma Sola, Tracy 7740; Bartow, Combs 1220; Myers, Hitchcock Lee Co. Pl. 483; Palm Beach, Curtiss 5386; Little River, Eaton 467; Miami, Chase 3850, Eaton 164 in part, Hitchcock 648, 658, 697; without locality, Simpson in 1889. Banamas: Great Bahama, Britton & Millspaugh 2706; North Bimini, Brace 3467 (all in Field Mus. Herb.). 19. Panicum elephantipes Nees. Panicum elephantipes Nees, Agrost. Bras. 165. 1829. ‘‘ Habitat in sylvis udis archipelagi Paraénsis.’’ The type, in the Munich Herbarium, labeled as above, consists of a large detached panicle, a leaf, and a few inches of a culm. Panicum fistulosum Hochst.; Steud. Syn. Pl. Glum. 1: 71. 1854. The locality mentioned is, ‘‘Surinam”’ and the specimen cited is ‘‘Hrbr. Kappler nr. 1434.’’ A specimen of Kappler 1434 was examined at the Florence Herbarium and another at Stockholm. As no specimen of this num- ber was found among the Steudel plants at Paris, we are unable to locate the type. In India is found a similar species, described in Hooker’s Flora of India@ as P. proliferum (P. palu- dosum Roxb.) which, judging from the specimens in the U. S. National Herbarium, is a smaller plant, with small, tardily exserted panicles 10 to 15 cm. long. DESCRIPTION. Culms ascending from a decumbent, often widely creeping base, rooting at the nodes, succulent, as much as 2 cm. thick, apparently a meter or more me ys isabaamagg high, glabrous, the nodes glabrous, usually conspicu- ously dark colored; sheaths glabrous, longer than the internodes, loose, the lower often tesselated by cross partitions between the nerves; ligules about 3 mm. long; blades 15 to 50 cm. long, 7 to 20 mm. wide, glabrous beneath, pilose above, at least near the base; panicles large and open, as much as 40 cm. long, aF\. Brit. Ind. 7: 50. 1896. ae 54 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. the short branchlets appressed along the ascending branches; spikelets 4 to 5 mm. long, lanceolate, acuminate; first glame one-fifth to one-fourth the length of the spikelet, subacute or obtuse, rarely pointed and one-third the length of the spikelet; second glume and sterile lemma subequal, 7 to 9-nerved, the palea of the sterile floret wanting; fruit 3 to 4 mm. long, about 0.8 mm. wide, lanceolate, acuminate, the mar- gins of the lemma above the middle thin and not inrolled. This large, succulent, semi-aquatic, apparently annual species, often producing dense masses of roots at the submerged nodes, is readily distinguished from P. dichoto- miflorum by the acuminate fruit. The Mexican specimens cited below have narrower blades than the others and none show the basal portion, but the floral characters agree with the type, except that the fruit is more turgid and less acuminate. DISTRIBUTION. In ponds and shallow water, West Indies and southern Mexico, south to Argentina. An introduced specimen was collected in North Carolina, ‘‘ Eastern part of state along seacoast,’’? by McCarthy in 1898. Mexico: Near Mexico City, Bourgeau 529, Pringle 6322,4 9577.4 GuaTemaLa: Alta Vera Paz, Goll 35A. Cupa: San Antonio, Hitchcock 152; Habana, Ledén 335. Porto Rico: Fajardo, Sintenis 938. ParaGcuay: Morong 1002 in part. . Urueuay: Montevideo, Arechavaleta in 1876; without locality, Arechavaleta in 1892. ARGENTINA: Lagos de la Darsesso, Buenos Aires in 1892, name of collector not given. Capillaria.—Annuals; papillose-hispid at least on the sheaths, or rarely glabrous, ligules membranaceous, ciliate, 1 to 3 mm. long; panicles many-flowered; more or less diffuse, often breaking away at maturity and rolling before the wind; spikelets pointed, glabrous, the first glume large and clasping, the fruit often falling from the spikelet before the disarticulation of the latter, smooth and shining, usually olive brown at maturity, the nerves showing as faint pale lines. Panicles more or less drooping. Spikelets not over 3.5 mm. long; Mexican species... -- - 28. P. sonorum. Spikelets 4.5 to5 mm. long; introduced from Old Worid . 30. P. miliaceum. Panicles erect. Inflorescence elongated, composed of several approximate, aie plichie wnmeles. ... 625.005 ots 2 eee 31. P. cayennense. Inflorescence not composed of approximate nor implicate panicles. Panicles more than half the length of the entire plant. Panicles narrow, usually less than half as broad SE ADOEEE «04 nz = os oes oe he eee eee 20. P. flexile. Panicle as broad as long. Spikelets 2 to 2.2, rarely 2.5 mm. long; blades not crowded toward the base. 23. P. capillare. Spikelets 3 to 3.3, rarely only 2.5 mm. long; blades usually crowded toward the MRE. . 6.2.60 wim ke Peas pee eee 24. P. barbipulvinatum. a These numbers were distributed as a variety of P. proliferum Lam., the varietal name being unpublished. eS e—E———EEeeeeee ee a - HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 55 Panicles not more than one-third the entire height of the plant. | , Spikelets not over 2mm. long, acute but not long- acuminate. Culms stout; blades about 1 cm. wide; Bomelets CPI... 5 S522 Os ae as - 2S 21. P. gattingeri. Culms slender; blades not over 6 mm. wide; spikelets not turgid.................. 22. P. philadelphicum. Spikelets 2.7 to 6 mm. long, acuminate. First glume about one-third the length of the spikelet, subacute or blunt..........-. 27. P. stramineum. First glume usually more than half the length of the spikelet, acuminate. Spikelets 4.5 to 6 mm. long. Spikelets 6 mm. long, scattered.. 29. P. parcum. Spikelets scarcely over 5 mm. long, approximate........- 26A. P. decolorans. Spikelets not over 4 mm. long. First glume more than three-fourths the length of the spikelet; spikelets 4 mm. long. ....-.-- 26. P. pampinosum. First glume half to two-thirds the length of the spikelet; spike- | lets not over 3.3 mm. long... 25. P. hirticaule. 20. Panicum flexile (Gattinger) Scribn. Panicum capillare flerile Gattinger, Tenn. Fl. 94. 1887. ‘Characteristic of the cedar glades.’’ In the Gattinger Herbarium are two specimens labeled ‘‘Panicum capillare L. var. flerile Gattinger” in Gattinger’s hand. The larger specimen is chosen as the type. This is labeled ‘‘Cedar glade near Nashville, Sept. ’88. A. Gattinger.’’ Panicum flexile Scribn. in Kearney, Bull. Torrey Club 20: 476. 1893. Based on P. capillare flerile Gattinger. Panicum minus Nash, Bull. Torrey Club 22: 421. 1895. Based on ‘‘Panicum capillare var. minus Muhl.,’’ though Nash’s description applies to P. philadelphicum. Scribner and Merrill @ applied to this species the name P. philadelphicum Bernh., but this name belongs to a different species. Muhlenberg ® describes P. flerile as a variety of P. capillare, but without giving a varietal name, saying, ‘‘ Varietas minor oceurrit ubique in cultis magis aridis,’’ and following this by a description. This specimen in the Muhlenberg Herbarium is labeled, ‘‘183 Panicum capillare var. minor.”’ Pursh ¢ describes P. flevile under the name of P. diffuswum Swartz. It was also described by Trinius @ as P. capillare 8 Panicula depauperata. DESCRIPTION. Plants erect, much branched from the base, 20 to 70 cm. high; culms slender, glabrous, or somewhat hispid below, nodes pubescent; sheaths papillose-hispid, the hairs shorter than in P. capillare; blades erect but not stiff, glabrous or sparsely hispid, @U.S8. Dept. Agr. Div. Agrost. Cire. 27: 3. 1900. A discussion of ‘‘The grasses in the Herbarium of Dr. H. Muhlenberg.” - bDescr. Gram. 124. 1817. eFl. Amer. Sept. 1: 68. 1814. @Gram. Pan. 215. 1826. See note under P. philadelphicum, p. 58. 56 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, as much as 30 cm. long, 2 to 6 mm. wide, rarely narrower; panicles rather few-flowered, oblong, narrow, 10 to 20 cm., rarely 30 cm. long, about one-third as wide, the branches at first narrowly ascend- ing, somewhat spreading at maturity, the peduncle of the panicle not brittle and readily breaking as in P. capillare; spikelets long-pediceled, 3.1 to 3.5 mm. long, 0.9 to 1 mm. wide, lanceolate, acuminate; first glume about one-third the length of the spikelet; second glume slightly longer than the sterile lemma, both 7 to 9-nerved, much exceeding the fruit, the palea of the sterile floret wanting; fruit 2 mm. long, 0.9 mm. wide, elliptic. This species is distinguished from P. capillare by the FiG. 36.—P. flerile. From type ™more slender culms, less dense pubescence, narrower specimen. blades, the narrow, less diffuse panicles and the longer, acuminate spikelets; and from P. philadelphicum by the narrow panicle and larger, acuminate spikelets. DISTRIBUTION. Sandy, mostly damp soil, meadows and open woods, Ontario to South Dakota, south to Florida and Texas. ' Ontario: Sarnia, Dodge 128, 130, Macoun 26332; St. Clair River, Dodge 124; Birch Island, Macoun 26331; Point Edward, Macoun 26330. PENNSYLVANIA: Lancaster County, Heller 701, 4777, 4786, Porter in 1862, Small in 1890 and 1892. Ox10: Erie County, Moseley in 1895; Columbus, Werner 6777. InprAnaA: Clarke, Umbach in 1898. Iuunots: Chicago, Bebb 2928, Chase 1474, 1479, 1729, 2006, Hill in 1901; Ingraham in 1893; Beach, Umbach 2543; St. Clair County, Eggert 241. Micuiaan: Port Huron, Dodge in 1909; Orion, Farwell 893; Jack- son County, Wheeler in 1892. Souta Daxora: Stearns, Wallace 46. Towa: Cedar Rapids, Pammel in FiG. 37.—Distribution of P. flezile. 1889 (Mo. Bot. Gard. Herb.). Missouri: Glenwood, Bush 3295; St. Louis, Lggert in 1875; Dodson, Bush 848; Eagle Rock, Bush 384; Noel, Bush 5259. District or CotumBIA: Steele in 1896 and 1898. VireintaA: Four-Mile Run, Chase 5444. West VirainiA: Peters Mountain, Steele 281. Fioripa: Without locality, Curtiss in 1886. Kentucky: Bell County, Kearney 312. TENNESSEE: Knoxville, Kearney in 1893, Ruth 66; Johnson City, Canby 221; Cocke County, Kearney 960; Nashville, Gattinger in Curtiss N. Amer. Pl. 3581 J, 3582a, and J. AxaBAMA: Monte Sano, Baker in 1897; Jackson County, Chase 4492. Mississippi: Starkville, Chase 4453, Kearney 72, 88 in part. Arkansas: Benton County, Plank 12, 107; Fulton, Bush 906. Texas: Clarksville, Plank 6, 10 in part. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 57 21. Panicum gattingeri Nash. Panicum capillare campestre Gattinger, Tenn. Fl. 94. 1887, not P. campestre Nees, 1826. No definite locality in Tennessee is given. In the Gattinger Herbarium are four specimens very much alike labeled ‘Panicum capillare L. var. campestre Gattinger’’ in Gattinger’s hand. The specimen with the following data is chosen as the type: ‘‘Cedar glades near Nashville, Sept. A Gattinger.”’ Panicum capillare geniculatum Scribn. in Kearney, Bull. Torrey Club 20: 447. 1893, not P. geniculatum Lam. 1798. ‘‘In the neighborhood of Wasiota,’’ [Bell Sates P. cattingeri. From County, Kentucky]. No type is indicated. Kearney type specimen. (on page 479) lists numbers 317, 335, 378, 497 as P. capillare geniculatum. A sheet of Kearney’s no. 378, col- lected near Wasiota, Bell County, Kentucky, in 1893, in the National Herbarium, and evidently the only one of the cited series examined by Scribner, is taken as the type. Panicum capillare gattingeri Nash in Britt. & Brown, Illust. Fl. 1: 123. 1896. Based on P. capillare campestre Gattinger. Panicum gattingeri Nash in Small, Fl. Southeast U. S. 92 and 1327. 1903. Based on P. capillare campestre Gattinger. DESCRIPTION. Plants at first erect, but soon decumbent-spreading and rooting at the lower nodes, freely branching from the lower and middle nodes; culms papillose-hispid, in robust specimens as much as | meter in length; sheaths hispid like the culms; blades 10 to 20 cm. long, 6 to 10 mm. wide, narrow to a rounded base, more or less hispid on both surfaces or nearly glabrous; panicles numerous, terminating the culms and main branches and auxillary at most of the nodes, short-exserted or, especially the axillary, included at base, oval or elliptic in outline, the terminal 10 to 15 cm. long, two-thirds as wide, the lateral smaller, rather densely flowered, the branches ascending or tardily spreading; spikelets shorter-pediceled than in P. capillare and more turgid, 2 mm. long, 0.9 to 1 mm. wide, elliptic; first glume about two-fifths as long as the spikelet, acute or blunt; second glume and sterile lemma equal, 5-nerved, but slightly exceed- ing the fruit, the palea of the sterile floret wanting; fruit 1.6 mm. long, 0.8 mm. wide, elliptic. This species differs from P. capillare in the branching, spreading habit, and the numerous panicles, oval in outline and less diffuse, produced from all the nodes. The spikelets in P. gattingeri are not so variable in length as in the other species in this group. DISTRIBUTION. : Nene ; 9 inet Sood ad | i aay : Open ground and waste places, often a weed in cultivated soil, Pennsyl- vania to Iowa and Missouri, south to North Carolina and Tennessee. This is the form introduced into Fig. 39.—Distribution of P. gattingeri. South Africa and described by Stapf as P. capillare.4 ‘Ontario: Kingston, Fowler in 1897 (Field Mus. Herb.). PENNSYLVANIA: Lancaster County, Heller in 1901. @ Dyer, Fl. Cap. 7: 407. 1898. A 58 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Ouro: Sheffield, Ricksecker in 1894; Cincinnati, Lloyd in 1883. Inp1ana: Lafayette, Dorner 87. Inurots: Peoria, Brendel; Wady Petra, V. H. Chase 124 in 1897, 783; Canton, Wolf in 1882. Minnesota: Hennepin County, Sandberg in 1890 (Univ. Cal. Herb.). Iowa: Emmett County, Pammel & Cratty 850. Missourt: Sugar Creek, Bush 4824; Monteer, Bush 5116. Maryann: Bethesda, Steeie in 1899; Plummers Island, Steele in 1897; Cabin John, Chase 2628. Districr or CotumsBia: Hitchcock 167, Williams in 1900. Vrraimnia: Arlington, Chase 5443. West Vireinia: Aurora, Séeele in 1898. Norts Carouina: Magnetic City, Wetherby 21. Kentucky: Bell County, Kearney 378. TENNESSEE: Cocke County, Kearney 962, 963; Knoxville, Ruth 59 in part; Nash- ville, Gattinger. 22. Panicum philadelphicum Bernh. Panicum capillare sylvaticum Torr. Fl. North. & Mid. U. 8. 149. 1824, not P. sylvaticum Lam. 1798. ‘‘In dry woods near New York.’”’ The type is in Columbia University Herbarium. Panicum philadelphicum Bernh.; Trin. Gram. Pan. 216. 1826. This is mentioned by Trinius as a variety of P. capillare, similar to his 8 [P. flexile], but with spikelets only acute [not acuminate as in the others], “‘misit s. n Pan. philadelphici sibi cl. BerNHaRDI.’’ Enough description is given to technically constitute publication. Nees later@ describes the species more fully. The type, in*the Trinius Herbarium, is from Philadelphia, sent by Bernhardi. On the same sheet is a specimen of P. flevile from the same source, but a drawing by Trinius with the name of P. phila- delphicum indicates which specimen is the type. Fia.40.—P.. philadelphicum. Panicum porphyrium Trin.; Nees, Agrost. Bras. 198. 1829. From duplicatetypespec- This is given as a synonym andere philadelphicum Bernh. imen in Stockholm Her- We have not seen the type. barium. Panicum torreyi Fourn. in Hemsl. Biol. Centr. Amer. Bot. 8: 497. 1885. Based on P. capillare sylvaticwum Torr. Fournier’s description > does not apply to this species and a specimen in the Fournier Herbarium, cited by him under this name, is P. bulboswm. Panicum capillare minimum Engelm.; Gattinger, Tenn. Fl. 94. 1887. No definite locality in Tennessee is mentioned. The type in the Gattinger Herbarium bears the name in Gattinger’s writing and the data “Greenbriar, Sept. ’78. A. Gattinger.” This name is initialed ‘‘F. L. S.’”’ by Scribner, hence the same name published¢ later by him is also based on this specimen. Panicum minimum Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Cire. 27: 4. 1900. Based on P. capillare minimum Engelm. Panicum soboliferum Tuckerm.; Scribn. & Merr. Rhodora 3: 106. 1901. This is given as a synonym under P. minimum. Tuckerman’s specimen, in the Gray Her- barium, is from ‘‘Head of Lake Memphremagog, Vt., Sept. 1859.” @ Agrost. Bras. 198. 1829. b Mex. Pl. 2: 28. 1886. ¢Tenn. Agr. Exp. Bull. 7: 44. pl. 10. f. 89. 1894. Ee j HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 59 DESCRIPTION. Plants light yellowish green, in small tufts, freely branching, erect or rarely decum- bent at base, 15 to 50 cm. high, depauperate, northern specimens sometimes forming small mats; culms slender, papillose-hispid to nearly glabrous, more or less zigzig at base, the lower internodes much shortened; sheaths mostly longer than the internodes, papillose-hispid; blades erect or ascending, 4 to 15 cm. long, 2 to 6 mm. wide, rather sparsely hirsute, rarely nearly glabrous; panicles exserted, diffuse, ovoid, 10 to 20 cm. long, forming one-third the entire height of the plant or more, few-flowered, the capillary, scabrous branchlets solitary, bearing rather short-pediceled spikelets, usually in twos, at the ends; spikelets 1.7 to 2 mm. long, 0.7 mm. wide, elliptic; -first glume about two-fifths the length of the spikelet, 5-nerved, acute;second glume and sterile lemma equal, only slightly exceeding the fruit, the palea of the sterile floret wanting; fruit 1.5 mm. long, 0.6 mm. wide, elliptic. This species differs from P. capillare in its narrow, erect blades, more slender culms, and smaller, fewer-flowered panicles, with more divergent branches and spikelets mostly in twos. The spikelets are usually slightly smaller, but the spikelets of the type of P. philadelphicum and of several other specimens are 2 mm. long. Two specimens from Stone Mountain, Georgia, Harper 184 and Hitchcock 439, have spikelets 2.2 mm. long, and are erect, narrow-leaved plants with somewhat the aspect of P. flerile. DISTRIBUTION. Dry open or sandy ground, Maine to Wisconsin and Oklahoma, south to Georgia and Mississippi. QueEBEc: Chandiere River, Macoun 7444 (Herb. Geol. Survey Can.). Marve: Holden, Knight in 1893; Mattawamkeag, Fernald 2802; Henderson, Parlin 1776. New Hampsuire: Alstead, Fernald 361. Vermont: Lake Memphremagog, Tuckerman in 1859 (Gray Herb.). Connecticut: Hadlyme Ferry, Graves 167. RuopeE Isianp: Lincoln, Fernald in 1906 (Gray Herb.). New York: Kinderhook Lake, Peck; Verona, Haberer in 1900. New Jersey: Lakehurst, Macken- zie 2366. PENNSYLVANIA: LancasterCounty, Heller 4789. Onto: Ottawa, Kellerman in 1900 (Univ. Ohio Herb.). Wisconsin: Sauk City, Zuders in 1885; Granite Heights, Cheney 2911. Missouri: Monteer, Bush 5119, 5120; Readings Mill, Bush 5203. DELAWARE: Collins Beach, Com- mons in 1879. MaryLanp: Glen Echo, Kearney in 1897; Chevy Chase, Chase 2599. District or CotumsBia: Chase 5441, Kearney in 1897, Steele in 1899, Vasey in 1882, Williams 10 in part, 12. Viren: Four-Mile Run, Hitchcock 166; Alexandria, House 413; Portsmouth, Chase 3687; Craigs, Steele 16. ‘West Virernia: Aurora, Steele in 1898; Baileysville, Morris 1283. Norra Carouna: Chapel Hill, Ashe. Sour Caro.ina: Orangeburg, Hitchcock 7. Grorais: Thomson, Bartlett 1024; Stone Mountain, Harper 184, Hitchcock 439. Fic. 41.—Distribution of P. philadelphicum. 60 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Kentucky: Harlan County, Kearney 151. TENNESSEE: Bluff City, Hitchcock 165; Cocke County, Kearney 961; Chester County, Bain in 1892. AvaBama: Cullman County, Eggert 58. OxiaHomMA: Sapulpa, Bush 722 in 1894. Texas: Dallas County, Reverchon 1842 (Mo. Bot. Gard. Herb.). 23. Panicum capillare L. Panicum capillare L. Sp. Pl. 58. 1753. Linneeus gives no description of his own but bases his name upon a phrase name of Gronovius@ which he quotes. Hence the type of P. capillare is the same as the type of Gronovius’s species, namely, Clayton no. 454, cited by Gronovius. This specimen, in the herbarium of the British Museum, is the common form of this species with broad blades and ample panicle; the spikelets are 2 mm. long. Linneus also cites a phrase name and a figure from Sloane? as a synonym, the Sloane plant, also in the British Museum, being Panicum trichoides Swartz. On the strength of these two citations, Linnzus gives the habitat as “‘ Vir- ginia, Jamaica.’’ In the Linnean Herbarium there is a specimen of P. capillare from ‘‘H. U.’’ [Hortus Upsalensis] upon which Linnzeus has written the name. Milium capillare Moench, Meth. Pl. 203. 1794. Based on Panicum capillare L. Panicum bobarti Lam. Encycl. 4: 748. 1798. Lamarck cites the following: “Gramen paniculatum virginianum, locustis minimis Bobarti. Moris. hist. 3. p. 202. no. 33. Ex herb. Vaill.’”’ It would seem that Lamarck is quoting the data on a specimen and not direct from Morison’s History, since the name Bobart nowhere occurs in Morison’s description or plate,¢ which applies to some species of Panicularia. In the Lamarck Herbarium is a fragmentary specimen of P. capillare bearing in Lamarck’s writing the data he quotes and in addition, also in his writing, the name ‘‘panicum Bobarti, Lam. dict.”” Since Fig. 42.—P. capillare. From ],amarck’s description applies to this it is taken as the type. eee eae eae Panicum capillare agreste Gattinger, Tenn. Fl. 94, 1887. es No definite locality in Tennessee is mentioned. The type specimen, in the Gattinger Herbarium, is labeled in Gattinger’s hand ‘‘ Panicum capillare L. var. agreste. Fields, Ridgetop, Sumner Co., 14. IX. ’82.”’ Collected by Dr. A. Gattinger. It is a medium-sized specimen of P. capillare. Panicum capillare vulgaris{e] Scribn. Tenn. Agr. Exp. Sta. Bull. 7: 44. 1894. No specimen is cited and no definite locality in Tennessee is given, but Scribner states that this variety is the same as ‘‘the variety agreste of Dr. Gattinger.”’ DESCRIPTION. Plants erect or ascending, simple or sparingly branched at the base or sometimes above, 20 to 80 cm. high; culms papillose-hispid to nearly glabrous, the pubescence dense at the nodes; sheaths usually longer than the internodes, densely papillose-hispid; blades 10 to 25 em. long, 5 to 15 mm. wide, scarcely narrowed toward the rounded base, hispid on both surfaces, the midrib prominent ; panicle densely flowered, large and very diffuse, often half the length of the entire plant, included at the base until maturity, the solitary or fascicled branches at first ascending, at maturity divaricately spreading, the whole panicles breaking away and rolling before the wind, the main a Fl, Virg. 1:13. 1739. See Hitchcock, Contr. Nat. Herb. 12: 118. 1908. b Voy. Jam. 1: 115. pl. 72. f. 8. 1707. ¢ Moris. Pl. Hist. 3: 202. sect. 8. pl. 6. f. 33. 1715. ———————————— HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 61 axis and branches sparsely pilose, the numerous capillary, scabrous branchlets bearing long-pediceled spikelets toward their ends; spikelets 2 to 2.5 mm. long, 0.8 to 0.9 mm. wide, elliptic; first glume about half the length of the spikelet, acute, 5 to 7-nerved; second glume and sterile lemma equal, more or less acuminate beyond the fruit, the palea of the sterile floret wanting; fruit about 1.5 mm. long, 0.7 to 0.8 mm. wide, elliptic. This common and widely distributed species is variable, occasional specimens approaching or intergrading with each of its closely allied species. Thus Bush 3318, Chase 2008,and V. H. Chase 183 are in- termediate between this and P. phila- delphicum; Bush 4138, Gattinger, Nash- ville, Tenn., in 1882, and Steele 243, between this and P. gattingeri; Bebb 2917 and Chase 1480 approach P. /flerile, and Blankinship, Huntsville, Okla- homa in 1896, and Griffiths 15 and 120 approach P. barbipulvinatum. Besides these there is a group of rather low, freely branching specimens with fewer- flowered, divaricately branched pani- cles forming as much as two-thirds the entire height of the plant, and acuminate spikelets 2.4 to 2.6 mm. long, tending to be in twos as in P. philadelphicum. These are the following and are not cited in the general distribution. Marne: Westbrook, Ricker 975; Massacuuserts: South Hadley, Cook in 1887; Pennsytvanta: Williamsport, Small & Heller in 1890; New York: Verona, Haberer in 1900; Int1nors: Chicago, Nelson 3000. A Florida specimen, Combs 665, has exceptionally turgid spikelets 2.5 mm. long. Fic. 43. Distribution of P. capillare. DISTRIBUTION. Open ground, fields and waste places, Maine to Dakota and Colorado, south to Florida and Texas; also introduced in the Bermudas, MAINE: Bangor, Knight 65. New Hampsuire: Jaffrey, Robinson 396. Vermont: Manchester, Day in 1898. Massacuusetts: Boston, Hitchcock in 1903 (Hitchcock Herb.). Connecticut: South Glastonbury, Wilson 1249. New York: New Hannibal, Pearce in 1883; Oxford, Coville in 1884. Ontario: Galt, Herriot 69; Kingston, Fowler in 1897; Wilmot Center, Umbach in 1899; Windsor, Macoun 26317; Sarnia, Morris A171; Belleville, Macoun in 1864. New Jersey: Stockholm, Van Sickle.in 1895. PENNSYLVANIA: Philadelphia, Scribner 48; Smith 110; Easton, Porter in 1894. Onto: Berea, Watson in 1897; St. Marys, Wetzstein in Kneucker Gram. Exs. 189; Columbus, Kellerman 6765. Intros: Chicago, Bebb 2917, Chase 1480; Naperville, Umbach in 1898; Roberts, Wilcox 109; Mokena, Chase 2008; Wady Petra, V. H. Chase 125 in 1897, 183, 774. Micuican: Port Austin, Morris A 226. Wisconsin: Sauk City, Duders in 1885; Ashland, Hitchcock 5099, 5111. Manirozpa: Killarney, Macoun 13225. Minnesota: Spicer, Frost in 1892; Fort Snelling, Mearns 756; Montevideo, _ Moyer 43. ba LG 62 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. SoutH Daxota: Yankton, Bruce 10; Aurora, Wilcox in 1892; Redfield, Grif- jiths 207; Huron, Griffiths 15; Aberdeen, Griffiths 120. Iowa: Thayer, Morris A 252; Decatur County, Fitzpatrick 31; Ames, Ball 120; Dudgeon in Pammel Amer. Weeds 20; Manchester, Ball 720; Ledyard, Pammel 767. Missourt: Creve Coeur Lake, Kellogg 20; Courtney, Bush 3318. Kansas: Manhattan, Hitchcock 2380, 3842, 3851, Norton 568. DELAWARE: Greenbank, Commons 30; Townsend, Canby in 1896. MARYLAND: Garrett County, Smith in 1879. = District or CoLumMBIA: Steele in 1896; Blanchard in 1891. Virernia: Arlington, Chase 5442. West Vrreinia: Sweetsprings, Steele 243. Nort Carouina: Wilmington, McCarthy in 1885. Froriwwa: Bay Head, Combs 665. TENNESSEE: Knoxville, Ruth 59 in part; Nashville, Gattinger in 1882. ALABAMA: Scottsboro, Chase 4495. Misstssipri: Starkville, Chase 4461, Kearney 80. ARKANSAS: Benton County, Plank 84, 159. Louistana: Natchitoches, Ball 149; McCall, Combs 1435. Texas: Seguin, Plank 99; Ennis, Smith in 1897; Texarkana, Plank 82; Dallas, Bebb 1299; Waller, Thurow in 1898; Galveston Island, Tracy 7409; Chilli- cothe, Ball 966. OxtaHomA: Huntsville, Blankinship in 1896. Montana: Columbia Falls, Hitchcock 4932. Cotorapo: Above Manitou, Williams 2184. CaLirorNia: Pinegrove, Hansen 599. Bermupas: Brown & Britton 21. 24. Panicum barbipulvinatum Nash. Panicum capillare brevifolium Vasey; Rydb. & Shear, U.S. Dept. Agr. Diy. Agrost. Bull. 5: 21. 1897, not P. brevifolium L. 1753. ‘‘Montana: Manhattan, on a shaded sandbar in the Gallatin River; July 19, [Shear] 436.’’ The type, collected by C. L. Shear, is in the National Herbarium. Panicum barbipulvinatum Nash in Rydb. Mem. N.Y. Bot. Gard. 1: 21. 1900. ‘‘Pani- cum capillare brevifolium Vasey * * * not Panicum brevifolium L.”’ is cited, but as a description follows and a new type is indicated, ‘*“YELLOWSTONE Parxk: Lower Geyser Basin, August 4, 1897, Rydberg & Bessey, 3544 (type),” this should not be considered as primarily a change of name. The specimen, in the herbarium of the New York Botanical Garden, agrees with Shear’s no. 436, men- tioned above. DESCRIPTION. Plants erect, 15 to 50 cm. high, freely branching : Sets J at the base, the branches often much shorter than Fig. 44.—P. barbipulvinatum. From ‘ : type specimen. the main culm and spreading; culms rather slender, glabrous or hispid below the nodes, the lower inter- nodes much shortened, the nodes often somewhat geniculate; sheaths usually longer than the internodes, papillose-hispid; blades erect or erect-recurving, 3 to 15 cm. long, 3 to 12 mm. wide, longer or wider in exceptional specimens, hispid on both surfaces or sometimes nearly glabrous; panicles soon exserted, about half the length of the -entire plant, rather few-flowered, the branches early divaricate, the pulvini often —- ——— HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 63 prominent, hispid, the axis and branches scabrous only or sometimes sparsely pilose; spikelets 2.5 to 3.3 mm. long (usually about 3 mm. long), 1 mm. wide, turgid, lanceolate-elliptic, acuminate; first glume about two-fifths the length of the spikelet, pointed, the midnerve scabrous toward the apex; second glume slightly longer than the sterile lemma, both much ex- ceeding the fruit, 9-nerved, the nerves scabrous toward the apex, the palea of the stenile lemma wanting; fruit 1.7 to 1.8 mm. long, 0.9 mm. wide, elliptic. This species differs from P. capillare in being on the average lower, in hav- ing shorter, less pubescent blades crowded toward the base of the plant, shorter, exserted panicles with divari- Fig. 45.—Distribution of P. barbipulvinatum. cate branches, and larger spikelets. The plants are often depauperate and tufted and the blades sometimes white- margined. ; Occasional specimens, as Mearns 743, Parish 1081 and Rydberg 1538, scarcely branch- ing at the base and with longer blades, approach P. capillare. DISTRIBUTION. Open ground, waste piaces and cultivated fields, Wisconsin, Minnesota to British Columbia, south to Texas and southern California. Inuinois: Oquawka, Patterson. Wisconsin: Madison, Agr. Exp. Sta. (probably transient adventive). Mrxynesora: Fort Snelling, Mearns 755; Hennepin County, Sandberg in 1889; Duluth, Hitchcock 5083. Norrs Dakota: Medora, Brannon 134, Bismarck, Field 1867; Devils Lake, Hitch- cock 5050. Souts Daxora: Hot Springs, Rydberg 1096; White Horse Camp, Griffiths 278; Cheyenne River, Wallace 2; Bellefourche, Griffiths 399; Huron, Griffiths 7. Nesraska: Simeon, Bates 1114; North Platte, Plank 38; Central City, Rydberg 2011, Shear 264; Niobrara, Clements 2705; Hooker County, Rydberg 1538; Grant County, Rydberg 1788. Kansas: Bucklin, Hitchcock in 1892; Ulysses, Thompson 56; Syracuse, Thomp- son 134; Tribune, Reed in 1892; Osborne City, Shear 152. Texas: Abilene, Tracy 8295; Paloduro, Gardner 24; without locality, Nealley in 1889. Montana: Great Falls, R. S. Williams 843; Silesia, Griffiths & Lange 15; Upper Big Horn River, Blankinship 179; Missoula River, Scribner 328; Glendive, Ward in 1883; Billings, Williams & Griffiths 246; Manhattan, Shear 436. Wyomine: No Wood Creek, Williams 2847; Sand Creek, E. Nelson 4984; Sybille Creek, E. Nelson 481; Badger, A. Nelson 8346; Cheyenne River, Geyer in 1839; Buffalo Fork, Tweedy 93, 95; Newcastle, Griffiths 684; Yellowstone National Park, Chase 5251, 5253, Hitchcock 2114, Rydberg & Bessey 3544. Ipano: St. Anthony, Merrill & Wilcor 152; Merrill 43; Shoshone, Hvermann in 1893; Rathdrum, Sandberg, Heller & MacDougal 713; Caldwell, Chase 4746. Wasuinaton: Kittitas County, Sandberg & Leiberg 431, Waitsburg, Horner 526; Streptoe, Vasey 3065; Prosser, Cotton 625, 891; Spokane, Chase 4986; Lake Chelan, Elmer 484, 848. British Cotumsia: Sicamous, Macoun 6 in 1889, 64 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. OREGON: Union, Cusick 3326¢; Malheur County, Griffiths & Morris 6764; Ontario, Griffiths & Morris 935; Harney County, Griffiths & Morris 891, 900; Malheur Lake, Griffiths & Morris 738; Manns Lake, Griffiths & Morris 696; Beulah, Griffiths & Morris 864; Princeville, Leiberg 834; Milton, Brown 34; Upper Klamath Lake, Leiberg 714; Sauvies Island, Howell in 1882; Corvallis, Hitchcock in 1901; Roseburg, Hitchcock 2789, 2798. Cotoravo: Denver, Eastwood 17a, Letterman in 1884; Manitou, Chase 5292, 5307; Rocky Ford, Griffiths 3313; Fort Collins, Agr. College 3368, Cowen 549; Golden, Rydberg 2505, Shear 755; Canyon City, Shear 965, Piedra, Baker ing 1899; Durango, Tweedy 386; Black Canyon, Baker 676; Steamboat Springs, Hast- wood 14; Grand Junction, Hitchcock 2204; Montrose, Hitchcock 2205; above Manitou, Hitchcock 2374; Ouray, Hitchcock 2275; Trinidad, Chase 5345. Urau: Cottonwood Canyon, Watson 1349; Logan, Rydberg 2351; Gunnison, Ward 679; Elk Ranch, Jones 6035; Ephraim, Tidestrom 2482. Nevapa: Leonard Creek, Griffiths & Morris 270; Big Creek, Griffiths & Morris 188; Ruby Valley, Watson 1349; Reno, Tracy 192; Virginia City, Bloomer 2265. New Mexico: Tierra Amarilla, Wooton 2948; Cliff, Smith in 1897; White Moun- tains, Wooton 303; Albuquerque, Harward 21a; Las Cruces, Plank 2; Mogol- lon Mountains, Metcalfe 434; Pecos, Standley 4948; Cimarron Canyon, Griffiths 5551; Organ Mountains, Wooton 1071; Deming, Hitchcock 3760; Las Vegas Hot Springs, Grant 5536; Mesilla Park, Hitchcock 3819; Dona Ana County, Wooton & Standley 3184. Arizona: San Francisco Mountains, Leiberg 5783; Canyon de Chelly, Griffiths 5852; Big Valley Mountains, Baker & Nutting in 1894; San Bernardino Ranch, Mearns 743, 788. CALIFORNIA: Castle Crag, Hitchcock 3069; Mt. Shasta, Palmer 2649 in 1892; Yreka, Butler 871; on the Sacramento, Wilkes Expl. Exp.; Tulare County, Michener & Bioletti 115, Palmer 2709 in 1892; Modoc County, Baker & Nutting in 1894; Riverside, Hall in 1901, Reed 1137; Colorado River, Cooper 2228; San Ber- nardino, Parish 1081. 25. Panicum hirticaule Presl. Panicum hirticaulum{e] Presl, Rel. Haenk. 1: 308. 1830. The locality as given by Presl is ‘‘ad Acapulco, Mexico.’”? The type, in the herbarium of the National Museum at Prague, is labeled ‘‘Mexico.”’ It rep- resents the medium form of the species, with nearly simple culms, narrowly ascending lower panicle branches and reddish brown spikelets, 3 mm. long. Panicum flabellatum Fourn. Bull. Soe. Bot. France II. 27: 293. 1880, not Steud. 1854. This is published in a list of plants collected in Nicaragua by Paul Lévy, ‘‘Omotepe (n. 1166)” being cited. The type, Lévy no. 1166 in the Paris Herbarium, was collected October, 1869, on ‘‘Prairies, Ile d’Omotepe.”’ Panicum polygamum hirticaulum[e] Fourn, Mex. Pl. 2: 28.1886. Based on ‘‘P. hirticaulum Presl,”’ but the specimens here cited are referable to P. maximum. This species was listed by Brandegee® as Panicum capillare var. glabrum Vasey, without description, having been so named by Vasey. The specimen in the National Herbarium consists of several small plants of P. hirticaule. Fic. 46.—P. hirticaule. From type Fspecimen. a Proc. Calif. Acad. II. 2: 211. 1889. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 65 DESCRIPTION. Plants erect, simple or nearly so, or sometimes branching and decumbent at base, 15 to 70 cm. high; culms papillose-hispid to glabrous, nodes spreading-hispid; sheaths papillose-hispid, but sometimes sparsely so; blades 5 to 15 cm. long, 4 to 13 mm. wide, often cordate at base, sparsely hispid or nearly glabrous, ciliate toward the base; panicles exserted, 5 to 15 cm. long, scarcely one-third the entire height of the plant, rather many-flowered, the branches ascending, the lower usually narrowly so, scabrous but not pilose, bearing rather short and appressed-pediceled spikelets along half to two-thirds their length, the glabrous pulvini inconspicuous; spikelets 2.7 to 3.3 mm. long, 1 to 1.1 mm. wide (smaller in occasional specimens), lanceolate-fusiform, acuminate, typi- cally reddish brown; first glume half to three-fourths the length of the spikelet, acuminate, the midnerve scabrous toward the apex; second glumeslightly longer than the sterile lemma, both much exceeding the fruit, strongly many-nerved, the midnerves scabrous toward the summit, the palea of the sterile floret small, nerveless; fruit 2 mm. long, 1 mm. wide, elliptic, a scar sometimes showing on either side at base. This species is variable; the Mexican specimens are mostly fairly typical, but the more northern ones are often rather freely branched or the panicles are less strict or the spikelets not reddish. In the following Mexican and Central American specimens the spikelets, though reddish brown and borne on strict branches, are only 2.2 to 2.5 mm. long. Mexico: Colima, Palmer 14, 143, and 145 in 1897; Alamos, Palmer 690 in 1890; Territorio de Tepic, Rose 3351. Nicaracua: Without locality, Flint in 1868. Fig. 47.—Distribution of P. hirticaule. DISTRIBUTION, Rocky or sandy soil, Texas to southern California and south through Mexico; also in the Galapagos Islands. Texas: El Paso, Jones 4212; without locality, Nealley in 1887. WaAsHINGTON: Bottomland near Bingen, Suksdorf 2330. This is probably intro- duced, as it is far out of its range. New Mexico: Organ Mountains, Wooton in 1907, Standley in 1906; Florida Moun- tains, Mulford 1012, 1078; Mangas, Metcalfe in 1897, Smith in 1898; Rio Gila, Greene 258; Hillsboro, Metcalfe 1442; San Luis Mountains, Mearns 2093. ‘Arizona: Tucson, Griffiths 1520, 3358, Hitchcock 3481, 3494, 3509, Pringle in 1881; Santa Rita Mountains, Griffiths 7005, 7194, Griffiths & Thornber 4, 28, 256; Salero Mountains, Griffiths 6123; Santa Catalina Mountains, Griffiths 7146; Huachuca Mountains, Holzner 1659; Fort Huachuca, Wilcox in 1894; Pearce, Griffiths 1938; Cochise, Griffiths 1918; Phoenix, Griffiths 7317; Patagonia, Hitchcock 3658, 3675; Benson, Hitcheock 3730; near monument 82, Mearns 1905; near Mexican boundary, Mearns 738, 767. CauirorNiA: Sierra Nevada Mountains, Lemmon in 1875; Jamacha, Canby 6 in 1894, 41616°—vo1 15—10——_5 66 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Mexico: Near International Boundary, Mearns 2294, Merton 1705; northern - Sonora, Griffiths 6758, 6799, 6892; Nogales, Hitchcock 3631; Llano, Hitch- cock 3526; Hermosillo, Hitchcock 3541, 3578, 3604; Guaymas, Hitchcock 3547, 3553, Palmer 208 and 346 in 1887; Alamos, Palmer 695 and 750 in 1890; Purisima, Lower California, Brandegee 8 in 1889, 42 in 1899; State of Chihua- hua, Nelson 6297, 6355, Palmer 1b in 1885 in part; Topolobampo, Palmer 249 and 251 in 1897; Culiacdn, Palmer 1544, 1545, and 1554 in pen in 1891. Gaudpacos IsLaANps: Agassiz in 1891. 26. Panicum pampinosum sp. nov. DESCRIPTION. Plants freely branching from the base; culms ascending from a decumbent base, branching at the lower and middle nodes, compressed, glabrous, or sparsely pilose below the panicle, the nodes pubescent; sheaths loose, papillose-hispid, ciliate; ligules of very stiff hairs 2 to 3 mm. long; blades 5 to 12 cm. long, 5 to 10 mm. wide, flat, scarcely narrowed at the rounded or subcordate base, sparsely pilose above, sparsely hispid beneath, the margin pilose and more or less undulate; pan- icles long-exserted, 6 to 15 cm. long, half to two-thirds as wide, the branches solitary, the lower as much as 5 to 8 cm. long, often bear- ing a branchlet at base, stiffly ascending, bearing several, mostly short, appressed branchlets with rather crowded, sbort-pedi- celed spikelets; spikelets about 4 mm. long, 1.4mm. wide, very turgid, pointed; first glume more than three-fourths the length of the spikelet, pointed; second glume and sterile lemma subequal, exceeding the fruit _and pointed beyond it, the sterile palea short; fruit 2.2 mm. long, 1.3 mm. wide, oval, an indistinct scar at the base. Type U. 8S. National Herbarium no. 592754, collected August 25, 1903, on range reserve, altitude 2,600 feet, Wilmot, Arizona, by Prof. J.J. Thornber (no. 193). Fig. 48.—P. pampinosum. From type specimen. DISTRIBUTION. On mesas, New Mexico and Arizona, New Mexico: Organ Mountains, Wooton 2014; Grant County, Rusby 444; without locality, Wright 2084. Arizona: Tucson Mountains, Griffiths 69393; Wilmot, Thornber 193 and Septem- ber 8, 1903 (the latter in Univ. Ariz. Herb.). Fia. 49.—Distribution of P. pampinosum. 26A. Panicum decolorans H. B. K.@ aWhile this revision was in press this species was collected by A. 8. Hitchcock at its type locality, Querétaro, Mexico. It has therefore been removed from the list of doubtful species. The description and illustration will be found on page 328. . HITCHCOCK AND CHASE—NORTH AMERICAN PAYICUM. 67 27. Panicum stramineum sp. nov. * DESCRIPTION. Plants ascending or widely spreading, sparingly branching at the base and lower nodes, or simple, 20 to 50 em. high; culms glabrous, the nodes appressed-villous; sheaths glabrous, sparsely papillose or papillose-hispid; blades erect orascending, 10 to 30 cm. long, 4 to 15 mm. wide, rounded or somewhat cordate at base, glabrous, scabrous on the margin and sometimes on the upper surface, some- times ciliate at base; panicles finally ex- serted, one-fourth to one-third the entire height of the plant, ovoid in outline, rather many-flowered, the pedicelsscarcely as appressed as in P. hirticaule; spikelets 3.2 to 3.7 mm. long, 1.5 mm. wide, elliptic, abruptly acuminate, turgid, pale stramine- ous; first glume one-third the length of the spikelet, blunt or subacute, the nerves usually anastomosing; second glume and sterile lemma equal, or the glume slightly shorter, not much exceeding the fruit; the palea of the sterile floret as long as the fruit, 2-nerved; fruit 2.2 mm. long, 1.3 mm. wide, obovate-elliptic, turgid, a rather prominent scar on either side at base. Type U.S. National Herbarium no. 592753, collected 1887, Guaymas, Sonora, Mex- ico, by Dr. Edward Palmer (no. 206). This species differs from P. hirti- caule in being nearly glabrous through- out, in the longer blades, more turgid, less long-pointed spikelets with shorter, scarcely acute first glume. Fic. 50.—P. stramineum. From type specimen. DISTRIBUTION. Rich bottom lands and damp soil, southern Arizona and northwestern Mexico. Arizona: Near the Mexican boun- dary, Pringle in 1884; Tucson, Thornber in 1901, and 219 (the latter in N. M. Agr. Col. Herb.). Mexico: Guaymas, Palmer 168a and 206 in 1887; Culiacan, Palmer 1538 in 1891; State of Sinaloa, Rose 1878, 1883; Acaponeta, Rose 1889, 3281. Fia. 51.—Distribution of P. stramineum. 28. Panicum sonorum Beal. Panicum capillare miliaceum Vasey, Contr. Nat. Herb. 1: 28 1890, not P. miliaceum L. 1753. Collected at Lerdo, Mexico, at the head of the Gulf of California, in 1889, Palmer 947. This is not based on P. miliaceum L. The type is in the National Her- barium. Panicum sonorum Beal, Grasses N. Amer. 2: 130.1896. Based on P.capillare milia- ceum Vasey. 68 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. . Plants robust, erect or spreading, 60 cm. to over 1 meter high; culms glabrous or sparsely papillose-hispid, the nodes pubescent; sheaths papillose-hispid or nearly glabrous; blades 15 to 40 cm. long, 15 to 30 mm. wide, cordate-clasping at base, rather prominently nerved, glabrous or sparsely papillose- hispid; panicles large and more or less drooping, 20 to 30 cm. long, densely flowered, the numerous branches narrowly ascending; spikelets 3 to 3.3 mm. long, 1.1 mm. wide, lanceolate, strongly nerved, brownish; first glume half to two-thirds the length of the spikelet, acuminate; second glume slightly exceeding the sterile lemma, the palea of the sterile floret wanting; fruit 2.1mm. long, 1 mm. wide, oblong-obovate, subacute. This species may be a cultivated form of P. hirticaule. It is large in all its vegetative parts. Fig. 52.—P. sonorum. From type Palmer states that it is used as food by the Cocopa Beta Indians, the seed being sown in spring on wet ground. A specimen from the State of Chiapas in southern Mexico, Nelson 2959, is intermediate between this and P. hirticaule but is more robust than the latter species, the blades being 15 cm. long and 18 mm. wide. DISTRIBUTION. Rich bottom land, northwestern Mexico. Mexico: Lerdo, Palmer 947 in 1889; southwestern Chihuahua, Palmer 1c in 1885; Culiacdn, Palmer 1539 and 1554 in part in 1891. 29. Panicum parcum sp. nov. DESCRIPTION. Plants sparingly branching from the middle or upper nodes; culms 30 to 50 cm. high, slender, erect or somewhat geniculate at base, glabrous; sheaths rather sparingly papillose-hispid, glabrate toward the base; ligules 1 to 2 mm. long; blades ascending, rather thin, linear, elongated, 10 to 30 cm. long, 2 to 6 mm. wide, slightly narrowed to the base, acuminate, sparsely pilose on both surfaces or glabrate, more or less ciliate; panicles short-exserted, the termi- nal 10 to 20 cm. long, half to two-thirds as wide (the axillary smaller), few-flowered, the few, slender, but not capillary, flex- uous branches solitary, remote, ascending, bearing ascending or appressed branchlets with scattered, rather long-pediceled spikelets; spikelets about 6 mm. long, 1.8 mm. wide, turgid, acuminate-pointed; first glume about half the length of the spikelet, pointed; second glume longer than the sterile lemma, both exceeding the fruit and pointed beyond it, the sterile palea about half as long as its lemma; fruit 3.3 mm. long, 1.4 mm. wide. Type U. S. National Herbarium’ no. 471378, collected October 9 to 15, 1891,.on “mountain side, not very common,’’ Lodiego, on the Culiacan River, Sinaloa, Mexico, by Edward Palmer (no, 1657). Fig. 53.—P. parcum. From type specimen. . ‘4 . : HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 69 The only other specimen examined is also from the vicinity of Culiacdn, Sinaloa; Copradia, Brandegee 4 in 1904. 30. Panicum miliaceum L. Panicum miliaceum L. Sp. Pl. 58. 1753. ‘Habitat in India” is the only citation given. We have not seen the type, which may not be in existence. Milium panicum Mill. Gard. Dict. no. 1. 1759. Based on Paniewm miliaceum L. Milium esculentum Moench, Meth. Pl. 203. 1794. Based on Panicum miliaceum L. Panicum milium Pers. Syn. Pl. 1:83. 1805. Based on P. miliaceum L. the original description of which is copied but very slightly augmented. Panicum asperrimum Fisch.; Jacq. Eclog. Gram. 46. pl. 31. 1815-1820. The description is based on a specimen grown in the garden of the University at Vienna from seed received from Count Razoumovsky of Gorenki [near Moscow]. This name was earlier listed by Fischer¢ without description. We have not seen the type, but Jacquin’s description and plate identify the species. DESCRIPTION. Plants erect or decumbent at base, usually branching from the basal nodes, 20 cm. to as much as 1 meter high; culms stout, hispid below the pubescent nodes or glab- rous; sheaths loose, sometimes - longer than the internodes, papillose-hispid; blades dry- ing yellowish green, more or less pilose on both surfaces, or glabrate, as much as 30 cm. long and 2cm. wide, rounded at base, gradually narrowed to the apex; panicles usually more or less included at base, 10 to 30 cm. long, more or less nodding, usually rather com- pact, the numerous branches narrowly ascending, very scabrous, spikelet-bearing to- ward thesummit; spikelets4.5 to5 mm. long, ovate, acumi- nate, strongly many-nerved; first glume half the length of the spikelet or more, acuminate; second glume and sterile lemma subequal, a small palea in the sterile floret; fruit 3 mm. long, 2 mm. wide, elliptic, stramineous to reddish brown. Fig. 54.—P. miliacewm. From Griffith’s no. 6490, East Himalaya. DISTRIBUTION. Waste places, introduced from the Old World. Cultivated under the name of broomcorn millet or hog millet. Scattered specimens, introduced or escaped from cultivation, are found in all the cooler parts of the United States, especially eastward. Cultivated specimens may be larger than indicated in the above description, while in dry soil depauperate specimens occur. _ Marne: Orono, Harvey in 1897. Vermont: Burlington, Brainerd in 1895 (Gray Herb.). Massacuusetts: Essex County, Conant in 1880. @ Cat. Hort.Gorenk. ed. 2. 3. 1812. 70 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Connecticut: Bridgeport, Hames in 1895. New Yor«: Ontario Beach, Britton in 1892. New Jersey: Califon, Fisher in 1899; Landisville, Goss in 1883. PENNSYLVANIA: Luzerne County, Small & Heller in 1890. Ouro: Cleveland, Claasen in 1891 (Gray Herb.). InprAna: Miller, Umbach 2363. Inurvors: Chicago, Chase 1633 (Hitchcock Herb.). MicuiGgANn: Without locality, Far- well 1414. DELAWARE: Wilmington, Com- mons 57. District oF CotumBiA: Steele in 1899. Fya. 55.—Distribution of P. miliacewm. Froriwa: Pensacola, Curtiss 6867. New Mexico: Lincoln County, Wooton & Standley 3528. CaLirorniA: Kenwood, Smith in 1898; Sacramento, Williams in 1906. 31. Panicum cayennense Lam. Panicum cayennense Lam. Tabl. Encycl. 1: 173. 1791. “‘Cayenna. D. Stoupy.” The type, in the Lamarck Herbarium, is labeled ‘‘ Panicum cayennense Lam. ill. gen. aff. Panico capillare.’? Lamarck states in the original description that this species is near P. capillare. In the general herbarium at Paris is a specimen of P. rudgeit from Cayenne collected by Le Blond, which is labeled in Lamarck’s hand ‘‘Panicum cayennense lam. illustr. et dict.” This specimen does not correspond to Lamarck’s m descriptions, especially the later one,@ so well as Te IGNRAIe Be cauenionse e eeuaeastye does the preceding specimen, which is therefore specimen. taken as the type. Panicum floribundum Rich.; Lam. Encycl. 4:742. 1798. This is given as a synonym under P. cayennense, and credited to ‘‘ Rich. herb.” The type, in the Richard Herbarium, is from Cayenne. This specimen agrees with the first of those mentioned above. Panicum pedunculare Willd.; Steud. Syn. Pl. Glum. 1:77. 1854. ‘‘Brasil.’’ This is credited to ‘‘ Willd. hrbr.’’ The name occursearlier®> asanomennudum. The type, in the Willdenow Herbarium, was collected by Humboldt in “‘ Amer. meridion.”’ Panicum cayennense patulum Doell in Mart. Fl. Bras. 2*: 220. 1877. Based on P. cayennense Lam. DESCRIPTION. Plants, erect, or, when much-branched at the base, spreading; culms 20 to 50 cm. high, often zigzag, glabrous or more or less papillose-pilose below the pilose nodes; sheaths papillose-pilose, ciliate; blades linear, 10 to 20 cm. long, 4 to 10 mm. wide, rather stiffly ascending, rounded at the scarcely narrowed base, rather sparsely papil- lose-pilose on both surfaces or sometimes nearly glabrous; panicles terminal and @ Lam. Encycl. 4: 742. 1798. b Steud. Nom. ed. 2. 2: 261. 1841. axillary from the upper 2 or 3 sheaths, rarely fascicled, approximate and forming an elongated inflorescence from two-thirds to almost the entire height of the plant, the individual panicles included at the base, 8 to 20 cm. long, half to two-thirds as wide, the slender, scabrous branches and branchlets and the long, flexuous pedicels divar- icate; spikelets 2.2 to 2.3 mm. long, 1.2 mm. wide, obovoid, turgid; first glume about half the length of the spikelet, acute; second glume and sterile lemma equal, slightly exceeding the fruit, abruptly short-pointed, strongly 5 to 7-nerved, the sterile palea about half the length of its lemma; fruit 1.7 mm. long, 1.1 mm. wide, oval, turgid. In its elongated inflorescence composed of several approximate panicles this species resembles P. rudgei Roem. & Schult., with which it has commonly been confused. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 71 DISTRIBUTION, Pine woods, Costa Rica, Cuba, and South America. Costa Rica: Buenos Aires, Tonduz 3685. Cusa: Pinar del Rio, Wright 3865; Herradura, Tracy 9073; Isle of Pines, Curtiss 267, Palmer & Riley 1086, Taylor 34. ; Frencu Guiana: Cayenne (Paris Herb.). Braz: Burchell 8350. Diffusa.—Perennials; culms stiff, somewhat tufted, sheaths mostly hirsute, ligules membranaceous, ciliate, 1 to 3 mm. long; blades long and narrow; spikelets mostly narrowly ovate, acuminate, glabrous; first glume clasping, the equal second glume and sterile lemma exceeding the fruit and pointed beyond it, the palea of the sterile floret about half as long as its lemma; fruit smooth and shining. The species of this group often resemble those of Capillaria, espe- cially in their spikelets, but the latter are all annual. Second glume and sterile lemma elongated, at least three DESO MELUS INUIT ecw e hac be seers era's 32. P. capillarioides. Second glume and sterile lemma not elongated. Panicles narrow, compact; blades 2 cm. or more wide... 38. P. hirsutum. Panicles diffuse, at least at maturity; blades not over e cm. wide. Blades 1 to3 mm. wide, plants spreading or ascending 33. P. diffusum. Blades mostly over 5 mm. wide, plants erect. Spikelets 4 to 4.2 mm. long, the midnerves of glumes and sterile lemma scabrous toward tre apeE. Us. SPE eee. 36. P. lepidulum. Spikelets usually less than 3.5 mm. long. Blades hirsute on both surfaces (sometimes glabrescent), not at all glaucous...... 37. P. ghiesbreghtii. Blades glabrous on both surfaces or with a few hairs on either surface, glaucous above. Panicles much exceeding the leaves; spikelets 3 to 3.5 mm. (rarely 3.7 RE) Se, See ge ose de 35. P. halivi. Panicles usually equaled or exceeded by the uppermost blades; spikelets 2 See I NS as eB wh 5a 34. P. filipes. "2 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 32. Panicum capillarioides Vasey. Panicum capillarioides Vasey in Coulter, Contr. Nat. Herb. 1: 54.1890. ‘Point Isabel,’’ collected by ‘‘G. C. Nealley, in the region of the Rio Grande, in Texas.’’ The type specimen, in the National Herbarium, bears the name in Vasey’s writing and the serial number 634 as in the list in which the species is published. DESCRIPTION. Plants in tufts of few to several culms from a knotted crown, erect or ascending, 30 to 55 cm. high; culms stiff, simple or sparingly branching, ap- pressed-pubescent or sometimes glabrate, the nodes densely ascending-pubescent; sheaths mostly equal- ing or exceeding the rather short internodes, papil- lose-pubescent; ligules about 1.5 mm. long; blades rather stiff, ascending, 10 to 30 cm. long, 2 to 10 mm. wide, scarcely narrowed to the rounded base, flat or drying somewhat involute, harshly papillose-pubes- \ cent on both surfaces, usually sparsely so beneath; Eig. 57.—P. capillarioides. From anicles short-exserted, usually nearly equaled by Bed lae aa the upper blades, diffuse, few-flowered, 10 to 20 cm. long, as wide or wider, the capillary branches stiffly spreading at maturity, bearing rather short-pediceled spikelets toward the ends, the axis and branches scabrous, the rather conspicuous pulvini pubescent; spikelets 5 to 6 mm. long,-1 to 1.2 mm. wide, lanceolate, long-acuminate; first glume one-third to half the length of the spikelet, acute, 7-nerved; second glume and sterile lemma, subequal, many-nerved, at least three times as long as the fruit, usually more or less inflated above it; fruit 1.6 to 1.8 mm. long, 0.8 to 0.9 mm. wide, elliptic. This species is readily distinguished from all others by the elongated second glume and sterile lemma, greatly exceeding the small fruit. Fig. 58.—Distribution of P. capillarioides. DISTRIBUTION. Prairie, southern Texas and northern Mexico. Texas: San Diego, Croft 240, Nealley 69 in 1892; Pena, Nealley 30 in 1891; Kings- ville, Piper in 1906; Corpus Christi to Brownsville, Hitchcock 218; without locality or date, Buckley (Hitchcock Herb.). Mexico: Monterey, Hitchcock 5547. 833. Panicum diffusum Swartz. Panicum diffusum Swartz, Prodr. Veg. Ind. Occ. 23.1788. ‘‘ Jamaica, Hispaniola.” The type,@ in the Swartz Herbarium, is labeled ‘‘diffusum fl. ind. occ.” Panicum caespititium Lam. Tabl. Encycl. 1: 173.1791. ‘Ex Amer. merid. com- munic. D. Richard.’ The type specimen, in the Lamarck Herbarium, is marked ‘‘ Ex D. Richard.”’ “See footnote ¢, page 36. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 18 DESCRIPTION. Plants in small, dense tufts, spreading, or ascending from a decumbent base, simple or sparingly branching, rarely repeatedly branching, 25 to 50 cm. high; culms slender, wiry, glabrous, the nodes appressed-pubescent; sheath, shorter than the internodes, striate, glabrous, or pubes cent along the margin toward the summit or the lower sparsely so throughout; ligules about 1 mm. long; blades erect from the sheath but often spreading at the ends, 5 to 20 cm. long, 1 to 3 mm. wide, flat or drying subinvo- lute, sparsely pilose on the upper surface, glabrous or sparingly pubescent beneath; panicles exserted, 5 to 10 em. long, nearly as wide, the few capillary branches at first ascending, stiffly spreading at maturity, bearing F¥6-59.—P.diffusum. From type a few short-pediceled spikelets toward the ends; spike- RF Ge: lets 2.1 to 2.5 mm. long, about 1 mm. wide; first glume about half the length of the spikelet, acute, 5-nerved; second glume and sterile lemma 7 to 9-nerved; fruit 1.5 to 1.6 mm. long, 0.9 mm. wide, elliptic. A collection from Santo Domingo, Wright, Parry & Brummel 627, is doubtiully referred here. It consists of tall plants, 60 to 70 cm. high, with spikelets 2.8 to 3 mm. long. The blades are narrow as in P. diffuswm, but hirsute on both surfaces as in P. ghiesbreghtii. DISTRIBUTION. Savannas and borders of open woods, West Indies. Banamas: New Providence, Britton & Brace 424. Cusa: Habana, Ledén 190, 305 in part, 923, 923b; Santiago de las Vegas, Baker & Wilson 511, Tracy 9111, Van Hermann 1444, Wilson 1405; Triscornia, Tracy 9082; Guanajay, Palmer & Riley 802; Prov. of Santa Clara, Leén 923c; with- out locality, Wright 3852, 3860 in part, 3877; Isle of Pines, Nueva Gerona, Curtiss 384, 494. ; DanisH West Inpres: St. Thomas, Eggers in 1882. Wriypwarp Istanps: Martinique, Duss 536. 34. Panicum filipes Scribn. Panicum filipes Scribn. in Heller, Contr. Herb. Frankl. Marsh. Coll.1: 13, 1895. “Growing in rich shaded ground in the upper part of the ‘ Arroyo,’ at Corpus Christi,”’ Texas. The type, in Hitchcock’s herbarium, was col- lected May 31, 1894, by A. A. Heller, no. 1809. The panicle is rather more lax and has longer, more delicate branchlets than usual in the species, and spikelets 2.5 to 2.6 mm. long. DESCRIPTION. Plants pale green, in small dense tufts, erect or ascending, 30 to 80 cm. high; culms simple or spar- ingly branching, stiff, glabrous except the appressed- BLOF) flipes. From type pubescent nodes; sheaths shorter than the internodes, specimen. glabrous or sparsely appressed-hispid toward the sum- mit; ligules about 1.5 mm. long; blades thin, ascend- ing or laxly spreading, 10 to 25 cm. long, 3 to 8 mm. wide, flat, glaucous and glabrous on the upper surface, glabrous beneath or very sparsely papillose-hirsute; panicles exserted, usually equaled or exceeded by the uppermost blades, 7 to 25 cm. long, about as wide, the distant branches spreading, the branchlets rather more numerous and the spikelets usually longer-pediceled than in P. diffusum; spikelets 2 to 2.6 mm. 74. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. long, about 1 mm. wide; first glume about two-thirds the length of the spikelet, acuminate, 3 to 5-nerved, second glume and sterile lemma 5 to 7-nerved; fruit 1.5 to 1.7 mm. long, about 0.8 mm. wide, elliptic. This species has been confused with both P. diffuswm and P. hallii, to both of which it is closely related. From the first 1t is distinguished by the taller, erect or nearly erect culms, and wider blades glabrous and glaucous on the upper surface. The typical form has a much more loosely-flowered panicle than has P. diffusum, but most of the specimens are less well marked. From P. hallii it is distinguished by the looser panicles of usually longer pediceled, smaller spikelets, by the usually taller culms and longer blades, often equaling or exceeding the panicle. The following specimens have spike- Fic. 61.—Distribution of P. filipes. lets 2.8 to 3 mm. long and appear to be intermediate between P. filipes and P. hallii. Texas: Abilene, Bentley in 1899; Daffan, Bodin 310; Dallas, Bush 1156; Spofford, Griffiths 6323; Olmito, Tracy 8908; Corpus Christi, Heller 1490. Mexico: Guerrero, Lagunillas, Langlassé 263. A specimen from ‘‘Overflow land along Colorado River,’’ southern California, Schellenger 3, is doubtfully referred here. It appears to be an annual but is probably P. filipes fruiting the first year from seed. DISTRIBUTION. Low open ground or among chaparral, southern Texas. Texas: Arlington, Reverchon 3526; Terrell, Warburton in 1904; Burnet, Plank 38; Llano, Plank 2; Kingsville, Piper in 1906; Houston, Nealley in 1886; Hemp- stead, Thurow in 1906; Kerrville, Heller 1883 in part; Seguin, Plank 97; Abi- lene, Bentley in 1899; Del Rio, Plank 85; San Diego, Nealley in 1894; Encinal, Griffiths 6387; Corpus Christi, Nealley 28 in 1891, in 1893 and 1894; Santa Maria, Nealley in 1889; Brownsville, Hitchcock 220, without locality, Buckley in 1881 and 1883, Drummond 286, 384, 394, Nealley in 1888. 35. Panicum hallii Vasey. Panicum hallii Vasey, Bull. Torrey Club 11: 64. 1884. ‘‘This is number 816 of KE. Hall’s Texas collection, distributed as P. gigantewm, Scheele.’’ The type, in the National Herbarium, was collected on ‘‘Dry hills, Austin, Eastern Texas, May 18, 1872, by Elihu Hall.”? Two species were distributed by Hall un- der 816, the other being P. jilipes. DESCRIPTION. Plants rather glaucous green, in small tufts, erect, 15 to 60 cm. high; culms simple or sparingly branch- ing from the lower nodes, glabrous except the ap- pressed-pubescent nodes; leaves commonly more or less crowded toward the base, the blades becoming curled or twisted, the lower sheaths overlapping on theshortinternodes; sheathssparsely papillose-hispid to glabrous; ligules about 1.5 mm. long; blades erect or nearly so, 4 to 15 cm. long, 2 to 6 mm. wide, flat, usually sparsely papillose-ciliate toward the base, otherwise glabrous or with a few long, delicate hairs on the upper | 1 Fic. 62.—P. hallii. From type specimen. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 75 surface or sparingly papillose-hispid beneath, often with a thin, cartilaginous, white margin; panicles usually long-exserted and much exceeding the leaves, 6 to 20 cm. long, rather narrowly flabellate in outline, the few branches stiffly ascending, bearing short, appressed branchlets with approximate spikelets on short, appressed pedicels; spikelets 3 to 3.7 mm. long, 1.1 to 1.5 mm. wide, turgid; first glume half to two-thirds the length of the spikelet, acuminate, 3 to 5-netved; second glume and sterile lemma strongly 5 to 7-nerved; fruit 1.7 to 2 mm. long, 1 to 1.3 mm. wide, oval, obtuse, dark olive brown at maturity. The following specimens have looser panicles than typical and spikelets only 2.8 to 3.2 mm. long, the two Plank specimens having also laxer blades. These appear to be intermediate between P. filipes and P. hallii but rather nearer the latter. Texas: Abilene, Tracy 7941, Del Rio, Plank 44, 57. Leiberg’s no. 5916, collected on cinder cones, is a depauperate form, scarcely 10 cm. high. DISTRIBUTION. Dry prairie, rocky and gravelly hills and canyons and in bottomlands, and irri- gated fields, Texas to Arizona and south to central Mexico. Texas: Texline, Griffiths 5600; Baird, Letterman in 1882; Austin, Hall 816, Stiles in 1884; Abilene, Bentley in 1899, Tracy 7950; Colorado, Tracy 7945; Big Springs, Tracy 7953; Kimble County, Reverchon 1682; Kerrville, Smith in 1897; San Antonio, Hitchcock 219, Plank 46, 53; Corpus Christi, Hitchcock 221; Olmito, Tracy 9338; Spofford, Griffiths 6288; Del Rio, Plank 72 in part; Midland, Tracy 7952, 7954; Guadalupe Mountains, Bailey 719; Fort Davis, Nealley in 1893, Marfa, Havard 23; Sierra Blanca, Nealley in 1893; El Paso, Vasey in 1881; with- out locality, Nealley in 1887. Fic. 63.—Distribution of P. hallii. New Mexico: Cimarron Canyon, Griffiths 5504; Roswell, Earle 302; Carlsbad, Tracy 8200; Organ Mountains, Hitchcock 3783; Las Cruces, Griffiths 7408; Dona Ana County, Wooton & Standley 3983; Deming, Hitchcock 3762; Grant County, Metcalfe 807, Smith in 1896 and 1897. J Arizona: San Francisco Peaks, Leiberg 5916; Ash Fork, Griffiths 7357; Prescott, Toumey in 1894; Clifton, Davidson 3la, 414a; Mescal, Griffiths 1813; Santa Rita Mountains, Griffiths 3388, Griffiths & Thornber 238, 309; Patagonia, Hitchcock 3706; Paradise, Blumer 1683; Huachuca Mountains, Holzner Inter- nat. Bound. Comm. 1566; without locality, Lemmon in 1883. Mexico: Tamaulipas, Palmer 554 in 1907; Coahuila, Palmer 1338 in 1880; Chi- huahua, Pringle 376. 36. Panicum lepidulum sp. nov. DESCRIPTION. Plants solitary or in small tufts, erect, 25 to 70 cm. high; culms usually producing one or two erect branches from the lower nodes, sparsely papillose-pilose to merely scabrous toward the summit; sheaths longer than the short lower internodes, shorter than the middle and upper ones, papillose-hispid, the hairs ascending; ligules about 2mm. long; blades erect, or spreading at the apex, 7 to 30 cm. long, 5 to 10 mm. wide, scarcely narrowed to the more or less infolding base, flat or folded, sparsely papillose- pilose to nearly glabrous on both surfaces, glaucous on the upper surface; terminal 76 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. panicles rather long-exserted, those of the branches short-exserted or slightly included at base, 7 to 20 cm. long, usually scarcely half as wide, the flexuous branches ascending, bearing short, rather spreading branchlets with 1 to 3 spikelets toward their ends, the whole forming a more evenly flowered panicle than in P. hallii; spikelets 4 to 4.2 mm. long, about 1.5 mm. wide, narrowly ovate, turgid, acuminate; first glume about half the length of the spikelet, acuminate, 5- nerved, the midnerve scabrous toward the apex; second glume and sterile lemma strongly 7 to 9-nerved, the midnerves scab- rous toward the apex; fruit 2.3 to 2.6mm. long, 1.38 to 1.5 mm. wide, oval, turgid, obtuse. Type U. 8S. National Herbarium no. 155163, collected September 22, 1885, by streams, rocky hills near Chihuahua, State of Chihuahua, Mexico, by C. G. Pringle (no. 497), and distributed as P. diffusum Swartz. This species is distinguished from P. hallii by the more evenly flowered, narrower panicle of larger spikelets, and by the greater amount of pubescence. The plants average taller than P. hallii, though Pal- mer 533 is a depauperate specimen only 15 cm. high. A specimen from Santa Catalina Mountains, Arizona, Griffiths 7063, with spikelets only 3.8 mm. long, is doubtfully referred here. Fie.64.—P.lepidulum. From type specimen. DISTRIBUTION. Moist places in the mountains, Chihuahua to the City of Mexico. Mexico: Chihuahua, Pringle 497; Durango, Palmer 525 in 1896, 533 in 1906; City of Mexico, Hitchcock 5958. 37. Panicum ghiesbreghtii Fourn. Panicum ghiesbreghtii Fourn. Mex. Pl. 2: 29. 1886. This name was earlier listed by Hemsley@ without description. Fournier cites three specimens, the first being ‘“Absque loco (GuiEsBREGHT),”’ which, since the species is named for this collector, is taken as the type. This isin the Paris Herbarium. It was collected in Mexico in 1845. Aspecimen collected by Ghiesbreght in Mexico and labeled P. ghiesbreghtii in the herbarium of the Botanical Garden in St. Petersburg, is P. filipes. Panicum hirtivaginum Hitche. Contr. Nat. Herb. 12: 223.1909. ‘‘Type specimen Wright 758, Cuba, U. 8. National Herbarium no. 559958.’’ The label on this specimen gives no locality in Cuba. DESCRIPTION. Plants in small tufts, rather robust; culms erect, pa- pillose, ascending-hirsute, 60 to 80 cm. high, the nodes densely hirsute; sheaths mostly longer than the inter- ; . i Fiq. 65.—P. ghiesbreghtii. From nodes, hirsute like the culms; ligules about 2 mm. type specimen. long; blades erect or ascending, as much as 60 cm. long and 12 mm. wide, flat, not narrowed to the rounded base, papillose-hirsute on both surfaces or glabrescent; panicle short-exserted, nearly equaled by the « Biol. Centr. Amer. Bot. 3 : 489. 1885. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 77 upper blades, 20 to 30 cm. long, usually less than half as wide, the branches ascending, naked at the base, the branchlets more or less appressed, bearing short-pediceled, approximate, but not crowded spikelets 3 mm. long, 1 mm. wide; first glume half to two-thirds the length of the spikelet, acute, 3 to 5-nerved; second glume and sterile lemma strongly 7 to 9-nerved; fruit 1.9 to 2 mm. long, 0.9 mm. wide, elliptic. This species differs from P. diffusum in its robust habit, wider, flat blades, hirsute on both surfaces, and in its larger spikelets. DISTRIBUTION. Low moist ground, Mexico, Central America, and the West Indies; also in Ecuador. Mexico: Borrego prés Orizaba, Bourgeau 2751; Yucatan, Izamal, Gawmer 2477. Costa Rica: Salinas Bay, Pittier & Durand 2633. Cusa: Santiago de las Vegas, Tracy 9116; Herradura, Tracy 9068; La Soledad, Eggers 5406; Calvario, Leén 922; Guayabal, Ledn 922b; without locality, Wright 758. Porto Rico: Tabucoa, Sintenis 4983. LEEWARD IsLANDs: Guadeloupe, Duss 3184 and a specimen without number or date. Ecuapor: Manabi, Zggers 15419. 38. Panicum hirsutum Swartz. Panicum hirsutum Swartz, Fl. Ind. Oce. 1: 173. 1797. ‘‘Habitat in Jamaicae et Hispaniolae graminosis.’’ The type, in the Swartz Herbarium, is from ‘‘Jamaica, Swartz.”’ DESCRIPTION. Plants robust, culms as much as 1 cm. thick and 1.5 meters or more high, simple or sparingly branching, glabrous or with a few scattered hairs, the nodes appressed- pubescent; sheaths mostly overlapping, appressed papil-. lose hirsute, the hairs sometimes dense at the summit; ligules dense, about 3 mm. long; blades flat, 20 to 50 cm. long, 20 to 35 mm. wide, scarcely narrowed to the rounded base, glabrous or with a few hairs toward the base, the margin serrulate; panicles short-exserted or included at base, 20 to 35 cm. long, 3 to 12 mm. wide, compact, densely flowered, the long lower branches erect, naked at base; spikelets 2 to 2.2 mm. long, about 0.9 mm. wide, rather turgid, abruptly pointed; first glume scarcely half =‘ Fi. 66.—P-. hirsutum. From the length of the spikelet, acute, 3 to 5-nerved; second Oi ae! ange glume and sterile lemma 5 to 7-nerved; fruit 1.5 mm. long, 0.7 mm. wide, elliptic. This species is much more robust than any other in this group, to which, because of its compact panicle, it is not very closely allied. DISTRIBUTION. Gravelly river banks and wet places, Mexico, Central America, and the West Indies. Leewarp Istanps: Guadeloupe, Duss 3917. ; Mexico: Oaxaca, Pringle 5573. Costa Rica: Confluence of Puerto Viejo and Sarapiquf, Biolley 7467; Port Limon, Pittier in 1904. Trinwwavb: Botanic Gardens Herb. 2295, Broadway 2629. 78 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum Bera Arechay.2 a South American species with numerous leaves clustered at the base, hispid sheaths, involute blades, and very diffuse panicles, a third or more the entire height of the plant, with verticillate lower branches, conspicuously pilose in the axils, and short-pointed spikelets 2.2 to 2.6 mm. long, was collected on ballast at Mobile, Alabama, in July and August, 1891, by Mohr and at Galveston, Texas, in 1903, by Hitchcock (the latter in Hitchcock Herb.). Maxima.—Perennials; culms mostly robust and more than 1 meter high, simple or branching at the base only, or with small sterile shoots from the lower nodes; ligules membranaceous, ciliate; blades linear, flat; panicles large, many- flowered; spikelets ellipsoid, glabrous, mostly faintly nerved, the sterile floret with a well-developed palea and in P. maximum a staminate flower; fruit strongly to very obscurely transversely rugose, puberulent at the apex. Culms with a corm-like base. Blades mostly over 5 mm. wide; culms more than 1 MNIGLCRUMIM ee isc Gis /e oS pa lelas'e SERS Aita. Spteas tee 41. P. bulbosum. Blades less than 5 mm. wide; culms rarely as much as 1 mptom ini ast sider. fous ioei.% Bue be ee 4la. P. bulbosum _ sci- aphilum. Culms from a creeping rootstock, not corm-like at base. Nodes hirstte; ligules 4 to 6 mm, long; fruit strongly TUPORO SpA dyd -NENMER shale Sans Se peewee rere 39. P. maximum. Nodes glabrous; ligules 2 mm. long; fruit very obscurely FHUPOSOL Seer bic cee e ck ae ick alate o edaees seen 40. P. plenum. 39. Panicum maximum Jacq. Panicum maximum Jacq. Coll. Bot. 1: 76. 1786. ‘‘In insula Guadeloupe sponte crescit.’’ In the Vienna Herbarium is a specimen from ‘‘Hb. Jacq. fil.’’® This is probably the most authentic specimen to be obtained. The plate in Jacquin’s Icones¢ is an excellent illus- tration of the species as commonly understood. Panicum polygamum Swartz, Prodr. Veg. Ind. Occ. 24.1788. ‘‘ India occidentalis” is the only local- ity cited. The type specimen, marked ‘Jamaica. Swartz. P. polygamum prodr.,” is in the Swartz Herbarium. Panicum laeve Lam. Tabl. Encycl. 1: 172. 1791. ‘‘E Domingo, ins. Franc.’’ is the only citation given. The type specimen, in the Lamarck Herbarium, is marked ‘‘Panicum laeven, Lam.ill.gen. * * * Ste. Dominique.”’ Panicum jumentorum Pers. Syn. Pl. 1: 83. 1805. Based on P. polygamum Swartz, the description of which Persoon copies. Panicum scaberrimum Lag. Gen. & Sp. Nov. 2. 1816. ‘‘Habitat in Nova Hispania. Introd. ann. 1804 ex seminibus per. D. Sessé missis.”” The type specimen, in the Madrid Herbarium, bears a label reading ‘‘Panicum scaberrimum sp. n. Ex h, r. m.¢ an 1804. Habitat in N. Hispania.” Fic. 67.—P. maximum. From type specimen of P. polygamum Swartz. a Anal. Mus. Nac. Montevideo 1: 147. 1894. “‘Campos del Departmento de Monte- video, San José, Florida, Mercedes, etc.”’ > Nicolas Joseph Jacquin the author of the Collectiones. ¢ Icon. Pl. Rar. 1: 2. pl. 18. 1781-1786. This work is dated 1781-1786, but “Jacq. coll. vol. 1” is cited, which would indicate that the Collectiones appeared the earlier. @ Hortus regius Matritensis. oo ie a HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 79 Panicum trichocondylum Steud. Syn. Pl. Glum. 2: 74. 1854. ‘‘Duchaising legit in Ins. Guadaloup.’’ The type specimen, in the Steudel Herbarium, bears a label with the data as published. Panicum praticola Salzm.; Doell in Mart. Fl. Bras. 2?: 203.1877. This is given asa synonym of P. maximum Faca. Doell cites ‘“‘Salzmann Herb. Bahiense N. 683,” which specimen we have not seen. DESCRIPTION. Plants light green, 1 to 2.5 meters high, or taller in cultivation, in tufts of few to many culms, from creeping rootstocks; culms robust, erect or sometimes genicu- late and rooting at the lower nodes, glabrous, the nodes usually densely hirsute; sheaths shorter than the internodes, papillose-hirsute to glabrous, ciliate, usually a dense ring of pubescence at the juncture with the blade; ligules 4 to 6 mm. long, stiffly and densely ciliate from a membranaceous base; blades erect or ascending, flat, 30 to 75 cm. long, 1 to 3.5 cm. wide, very scabrous on the margin, otherwise glabrous, or hirsute on the uppersurface at the base; pani- cles finally long-exserted, 20 to 50 cm. long, usually about one-third as wide, densely flowered, the long, rather stiff branches ascending, naked at the base, the lower in whorls, the axils pilose, the branchlets short, appressed, bear- ing more or less clustered, short-pedi- celed spikelets; spikelets 3 to 3.3mm. long, 1 to 1.1 mm. wide, and about as thick, oblong-ellipsoid, glabrous, somewhat shining, faintly nerved; first glume about one-third the length of thespikelet, obtuse; second glume and sterile lemma subequal, slightly exceeding the fruit, thin in texture, the lemma inclosing a 8taminate flower; fruit 2.3 to 2.5 mm. long, about 1 mm. wide, elliptic, transversely rugose, minutely puberulent at the apex. Fia. 68.—Distribution of P. maximum. DISTRIBUTION. Cultivated for forage under the name of Guinea grass in the Gulf States, especially in Florida, and southward through tropical South America, whence it has escaped into fields and waste places;4 also in the tropical parts of the Old World. Froripa: Duval County, Fredholm 373; Indian River, Curtiss 3597**; Eustis, Nash 1730; Grasmere, Combs 1170; Orange County, Baker in 1897; Braiden- town, Combs 1310; Caximbas Island, Simpson 580; Key West, Blodgett; with- out locality, Simpson. Mexico: State of Vera Cruz, Finck 8 and in 1893, Smith 1409; Cérdoba, Kerber 48; State of Colima, Pmrick 3; Huitamalco, Liebmann 425 in part; Zacualpan, Purpus 3774. @ Trimen (Hand. Fl. Ceyl. 5: 154. 1900) states the following concerning this grass: “The well-known Guinea Grass was introduced from W. Trop. Africa into Jamaica about 1774, by Mr. John Ellis, as food for some birds which he had imported. The birds died, and the seed, being thrown away as useless, yielded a magnificent grass greedily eaten by cattle and horses. It was introduced into India in 1802 by Sir John Sinclair, and must have been rapidly disseminated, for I find a specimen in Rottler’s Herbarium (named P. meneri, miliacea, var. (?) P. nodosum, nob.) received from Heyne, with the date, June 3, 1808. There is no record of its introduction into Ceylon, but it is included in Moon’s Catalogue, published in 1824.”’ 80 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Guatremata: Escuintla, J. D. Smith 2705, 2706; Morales, Kellerman 6267; Alta Vera Paz, Cook & Griggs 579; Gualin, Deam 6268. Honpuras: Cortez, Kellerman 4725. Satvapor: Izalco, Pittier 1960; San Salvador, Velasco in 1906; without locality, Renson 293. Costa Rica: San José, Pittier 9050; Alajuelita, Pittier 2995, Tonduz 2995;4 Boca Banana, Tonduz 9114; Zent Farm, Pittier in 1904. Bermupas: Brown & Britton 20. Bananas: Nassau, Curtiss W. I. Pl. 124; Eleuthra, Geogr. Soc. Baltimore 338. Cusa‘ Pinar del Rio, Palmer & Riley 377; El Guama, Palmer & Riley 178; San Diego de los Bafios Palmer & Riley 542, 545; Guanajay, Palmer & Riley 816; Santiago de las Vegas, Hitchcock 157, Wilson 438; Santiago, Leén & Voisard 915, Pollard, Palmer & Palmer 283; Guines, Leén 427, Liebmann 445 in part; Herradura, Hitchcock 156; Guayabal, Leén 921; Marianao, Leén 957. Jamaica: Gordon Town, Hart 797; Port Antonio, Fredholm 3319. ° Porto Rico: Cayey, Sintenis 2468; Mayaguez, Sintenis 51; El Sobrante, Eggers 1226; Guanica, Sintensis 3366; Martin Pena, Heller & Heller 377. Danisu West Inp1Es: St. Croix, Ricksecker 200, 413; St. Thomas, Eggers in 1887, Millspaugh 454. LEEWARD IsLANDs: Guadeloupe, Duss 3186. Winpwarp IsLanps: Martinique, Duss 1288; Barbados, Dash 259; Granada, Broadway in 1905. Cotomsra: Santa Marta, Smith 2153; Magdalena, Pittier 1617. VENEZUELA: Island of Margarita, Miller & Johnston 177. Braz: Province of Ceard, Gardner in 1838; Rio, Glaziow 18627; without locality, Glaziou 16612, Riedel 53. 40. Panicum plenum sp. nov. DESCRIPTION. Plants mostly in large clumps, 1 to 2 meters high, erect, from a stout rootstock, mostly glaucous; culms robust, compressed, glabrous, usually decumbent at base, sometimes branching at the lower nodes; sheaths overlapping on the short lower internodes, shorter than the upper, glabrous, or the lower sometimes pubescent toward the summit, more or less keeled; ligules densely ciliate, about 2 mm. long; blades erect or ascending, or the lower spreading, flat, 20 to 35 cm. long, 7 to 17 mm. wide, glabrous on both surfaces or rarely sparsely pilose on either surface toward the base, the upper surface scarcely scab- rous; panicles 20 to 50 cm. long, about two-thirds as wide, the slender branches somewhat spreading, the general appearance much like that of P. bul- Fic. 69.—P. plenum. From type bosum but proportionately wider, the main axis specimen. nearly smooth; spikelets 3 to 3.4 mm. long, 1.2 mm. wide, oblong-elliptic, glabrous, rather strongly nerved; first glume scarcely half the length of the spikelet or less, subacute, 3-nerved; second glume and sterile lemma subequal, scarcely exceeding the fruit, the palea aSome of the Pittier collections were distributed with the name of Tonduz as col- lector. The numbers are in one series, some of the plants having been collected by Pittier, some by Tonduz, and a few by Biolley. The labels may be marked, ‘‘Herb. Instit. physico-geogr. nat. costaricensis,’’ or, “‘Plantae costaricenses exsiccatae.”’ —— HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 81 of the sterile floret about as long as its lemma; fruit 2.9 to 3 mm. long, 1 mm. wide, elliptic, acute, only very obscurely rugose, minutely puberulent at the apex. Type U. S. National Herbarium no. 495701, collected September 18, 1903, ‘‘at Mangas Springs, 18 miles northwest of Silver City, Grant County, New Mexico, by O. B. Metcalfe (no. 739), altitude 4,770 feet.”’ This species is related to P. bulbosum from which it is distinguished chiefly by the creeping rootstock and decum- bent, not corm-like, base of the culms. Specimens lacking the base may be recognized by the compressed culm, scarcely scabrous blades, shorter first glume, and only very obscurely rugose fruit, appearing smooth except under a high power lens. Many specimens of this species have been distributed as P. avenaceum but an examination of the type specimen of the latter, together with the statement in the original description that the base is bulbous, shows it to be the same as P. bulbosum. Fic. 70.—Distribution of P. plenum. DISTRIBUTION. Moist places in rocky hills and canyons, Texas to Arizona, south to southern Mexico. Texas: Kerrville, Heller 1898; Colorado, Tracy 8224; without locality, Nealley in 1887. New Mexico: Organ Mountains, Vasey in 1881; Wooton 2017; Mangas Canyon, Smith in 1896; Mangas Springs, Metcalfe 738, 739; Mangas, Metcalfe 6, 80 in part, J. K. Metcalfe in 1897, Smith in 1896; Greenwood Canyon, Smith in 1896; Las Cruces, Griffiths 7400, 7401; ‘‘from Western Texas to El Paso,’’ N. M., Wright 786. 3 Arizona: Santa Rita Forest Reserve, Griffiths 3427; Fort Huachuca, Wilcox in 1894; Patagonia, Hitchcock 3649; Dos Cabezas, MacDougal 789; Mustang Mountains, Pringle 7 in 1884 (Hitchcock Herb.); without locality, Emersley in 1890. Mexico: Chihuahua, Wilkinson in 1885; Durango, Palmer 741 in 1896; Faral, Schumann 1733; Orizaba, Botteri 160; Las Cuevas, Hartman 170. 41. Panicum bulbosum H. B. K. Panicum bulbosum H. B. K. Nov. Gen. & Sp. 1: 99.1815. ‘‘Crescit in Nove His- paniz scopulosis et frigidis juxta Santa Rosa, Los Joares et Guanaxuato, inter 1070 et 1360 hexap. altitudinem.’’ The type specimen, from the Bonpland Herbarium in the Paris Herbarium, has a well developed corm. The accompanying label reads, ‘‘ Pani- eum bulbosum Kunth Synops. 175. in scopulosis & frigidis. Nova Hisp. alt. 1070- 1360 hex. No. 4250.’ The spikelets are 3.7 mm. long. Panicum avenaceum H. B. K. Nov. Gen. & Sp. 1: 99. 1815. ‘‘ Crescit in regno Qui- tensi, in valle amoena Chilloensi et planitie Cachapamba, regione subtemperata, alt. 1340 hexap.”’ The type specimen, from the Bonpland Herbarium in the Paris Herbarium, bears a label with the following data: ‘‘Panicum avenaceum Kunth, Synops. 175. (P. bulbosum proximum) in valle amoena Chilla. alt. 1340 h. regn. Quitensis. No. 3016.”’ The base of the specimen is wanting, though the description states that it is bulbous. The spikelets are 4mm.long. These slightly larger spikelets and the few hairs on the 41616°—voL 15—10—_6 82 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. sheaths are the only characters not agreeing with those of the type of P. bulbosum. The author states it is closely allied to that species and scarcely different from it. Panicum gongylodes Jacq. Eclog. Gram. 30. pl. 21. 1815-1820. The description is based on a plant grown in the University garden at Vienna in 1807 from seed received from the Botanical Garden at Montpellier under the name Panicum altissimum Brouss.¢ of P. kunthii and the specimens cited refer to P. elephantipes. A synonym cited, P. arenarium [Brot. misapplied by] Schlecht., is P. gowini. Panicum ichnanthoides Fourn. Mex. Pl. 2: 30. 1886. This name was earlier listed without description by Hemsley.¢ Fournier cites ‘‘ Orizaba (F. MULL[ER] n. 2082 in herb. Perror.)” The specimen in the herbarium of the Botanical Garden in St. Petersburg labeled as above is Miiller 2002, the number as printed being an error. The panicle is narrow, the spikelets 3.5 to 3.8 mm. long. Panicum virgatum confertum Vasey, Bull. Torrey Club 18: 26. 1886. Vasey gives no definite locality, merely stating that the variety grows, ‘‘particularly on the sea coast.”? The type, in the National Herbarium, is labeled ‘‘Seashore at Atlantic City, N. J., Geo. Vasey, 1884.” The panicles are narrow and compact, about 20 cm. long and 5cm. wide. The spikelets are about 3.5 mm. long. Panicum virgatum elongatum Vasey, Bull. Torrey Club 18: 26. 1886. No defi- nite locality is given. The type, in the National Herbarium, was collected at White River, South Dakota, August, 1892, by E. N. Wilcox, no. 13. This is an immature specimen, the long narrow panicle and slender spikelets, as described, being due to immaturity. This name is not based on P. elongatum Pursh, since Vasey says, “‘per- haps this is the Panicwm elongatum of Pursh.”’ Panicum virgatum diffusum Vasey, Bull. Torrey Club 13: 26.1886. ‘“‘Sandy prai- ries, Kansas, Colorado, etc.’’ No specimen bearing this name can be found in the National Herbarium nor any from Kansas or Colorado collected before 1886. Palmer’s no. 376 from Indian Territory in 1868, with a very large and diffuse panicle and marked ‘‘P. virgatum?’’ by Vasey well answers his brief description. DESCRIPTION. Plants erect, usually 1 to 2 meters high, producing numerous scaly, creeping root- stocks, glabrous throughout except as noted, commonly purple tinged, often glaucous, especially on the internodes and upper surface of the blades; culms in large to small clumps or even solitary, simple, robust, tough and hard; sheaths longer than the rather short lower internodes, usually shorter than the upper ones, often ciliate, sometimes @ Descr. Gram. 120. 1817. ¢ Biol. Centr. Amer. Bot. 3: 490. 1885. 6 Mex. Pl. Gram. 2: 29. 1886. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 89 villous at the throat; ligules dense, 2 to 4 mm. long; blades ascending, 10 to 60 cm. long, 3 to 15 mm. wide, slightly narrowed toward the base, and gradually long-acu- minate, flat, sometimes pilose on the upper surface toward the base, rarely to the apex, margins scabrous; panicles long-exserted, 15 to 50 cm. long, mostly one-third to half as wide, but sometimes contracted, or very loose and nearly as wide as long, usually many-flowered, the slender, scabrous, usually fascicled branches ascending or spreading, naked at base, repeat- edly branching along the upper half or two- thirds; spikelets rather short-pediceled, 3.5 to 5 mm., rarely but 3 or as much as 6 mm. long, 1.2 to 1.5 mm. wide, elliptic-ovate, acuminate, strongly nerved; first glume clasping, two-thirds to three-fourths the length of the spikelet, rarely equaling the sterile lemma, acuminate to cus- pidate, 5-nerved; second glume longer than the sterile lemma, both much exceeding the fruit, 5 to 7-nerved; fruit narrowly ovate, the margins of the lemma inrolled only at base. This species is well marked but variable. The blades are usually glabrousor pilose above near the base only. Sometimes, however, the entire upper surface or even both surfaces are pilose. Examples of this are: Minnesota: Mearns 743; Soutrn Daxora: Thornber; NeprasKa: Rydberg 1561; Kansas: Smyth92; Grorata: Tracy 3604, Harper 631; Fiormpa: Combs 597; ALaBama: Carver 72; Mississippi: Chase 4459. The form named by Vasey P. virgatum confertum, with more or less compact panicles, is represented by: New Jersey: Scribner in 1895, Vasey in 1884, Ward in 1884; Vir- GIntA: Knowlton in 1897; NortH Carouina: McCarthy in 1885; Forma: Kearney 158. The size of the panicle is variable, in northern specimens being often much dwarfed. The branches may be stiffly ascending or laxly spreading or drooping, these characters not being coédrdinate with others. The glaucous character also appears to be without Significance in separating forms, glaucous and green individuals growing under the same conditions. All these variations are connected by all shades of intergradation with the typical form. Throughout the western portion of the range of the species there is found, chiefly on sandy soil, a form with mostly single or loosely cespitose culms, often decumbent at base, pale green or glaucous foliage, and small panicles with ascending branches. We have been unable to separate this form as a subspecies because of the numerous intergrading specimens. ‘The following, which are not cited under the distribution of the species, are representative of this form: Sourn Daxora: Huron, Williams in 1892; Bellefourche, Griffiths 395; White River, Wallace 3, 4, 5; Iowa: Cherokee County, Crozier in 1888; Nepraska: Sidney, Plank 13; Mullen, Rydberg 1597; Kan- sas: Morton County, Hitchcock Pl. Kans. 570a; Texas: Tascora, Reverchon 2844; Channing, Williams 3061; CoLtorapo: Raton Mountains, Griffiths 5463; Arizona: Flagstaff, MacDougal 265; without locality, Lemmon 3154. The spikelets are frequently affected by a smut, this sometimes resulting in abnor- mal forms with spikelets in glomerules or with two to several staminate or abortive florets to a spikelet, as in Sandberg from Minnesota in 1891 and Brandegee from Colorado in 1878. Fria. 79.—P. virgatum. From type specimen in Gronovius Herbarium. DISTRIBUTION. Prairies, moist open ground, open woods and salt marshes, Maine to Wyoming and south to Florida and Arizona, southwest through Mexico to Costa Rica; also in the Bermudas and Trinidad. Marine: Scarboro, Chamberlain 552. New Hampsurre: Walpole, Fernald 271 (N. E. Bot. Club Herb.). CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Vermont: Brattleboro, Grout in 1895. Massacuusetts: South Hadley, Cook in 1887. RuoveE Istanp: Newport, Hitchcock 170. Connecticut: Fairfield, Hames in 1896; South Glastonbury, Wilson 1251. New York: Cayuga Lake, Dudley in 1884. Ontario: Toronto, Macoun 26324. New Jersey: Stockton, Fisher in 1897; Camden, Scribner 104; Atlantic City, Scribner in 1895, Vasey in 1884, Ward in 1884. PENNSYLVANIA: Easton, Porter in 1895; Sayre, Barbour in Kneucker Gram. Exs. 302; Safe Harbor, Heller in1893; Harrisburg, Small in 1888. Ouro: Sandusky, Morris 53, 124; Columbus, Fischer 6801; Cedar Point, Wilkinson 6805. , Inp1ana: Dune Park, Umbach1922; Indiana Harbor, Deam 2638; Pine, Umbach in 1898; Clarke, Umbach in 1896; Lafayette, Dorner 12, 85. Iniinors: Lake County, Gleason & Shobe326; Chicago, Chase 1178, Fia. 80.—Distribution of P. virgatum. 1636; Joliet, Skeels 470; Oregon, Waitein 1885; Thawville, Wilcor100; Wady Petra, V. H. Chase 73in 1897; Prince- ville, V. H. Chase 818; Peoria, Brendel, McDonald 59; Athens, Hall in 1870. Micuican: Lansing, Bratt in 1881 (Hitchcock Herb.). Wisconsin: Witches Gulch, Cheney 3868; without locality, Wood in 1889. Minnesota: Pipestone City, Sheldon in 1891; Chippewa County, Moyer 42; St. Paul, Blanchard in 1890; Fort Snelling, Mearns 743 in part, 758, 769, 793. Manirosa: Brandon, Macown 13231; Assiniboine River, Macoun 29303; Red Deer River, Macoun 29304. SASKATCHEWAN: Saskatoon, Macouwn 73003. Norta Daxora: Minot, Waldron 1813; Davenport, Wright 1866; Dunseith, Brannon 99. Souts Dakota: Aberdeen, Griffiths 82, 132; Black Hills, Rydberg 1097; Frankfort, Griffiths 53; Huron, Griffiths 22; Aurora, Wilcox 54; Brookings, Wilcox 15; Jamesville, Bruce 16; Grindstone, Girifiths 751; Bellefourche, Griffiths 371; Redfield, Griffiths 206; White Horse Camp, Griffiths 295; Zell, Griffiths 242; Dell Rapids, Thornber in 1892; White River, Wilcox 13. Iowa: Armstrong, Shimek 63; Spirit Lake, Geyer in 1839; Fayette County, Fink 459; Mount Pleasant, Mills 770; Iowa City, Somes 189, 219; Murray, Morris A 294. Nesraska: North Platte, Rydberg 2516, Shear 767; Talmage, Elmore 6, 126; Pishelville, Clements 2742; Broken Bow, Plank 57; Mullen, Rydberg 1561. Missouri: Courtney, Bush 705 in part, 4038; Allenton, Letterman in 1892; Jefferson County, Eggert in 1896; Monteer, Bush 5105. Kansas: Manhattan, Hitchcock 3840; Riley County, Norion 570; Hutchinson, Smyth 9; Osborne City, Shear 223; Wichita, Smyth 230, 257; Florence, Gri/- Jiths 5047; Morton County, Hitchcock 6274; Platt, Smyth 92. DELAWARE: Kimensi, Commons in 1897. MaryLanp: Cumberland, Shriver 610; River View, Blanchard in 1891; Bay Ridge, Scribner in 1897; Chesapeake Beach, Chase 2528, Hitchcock 442, 2391, House 358; Anne Arundel County, Smith in 1879; High Island, Dewey 134, Steele in 1896, Scribner in 1894; Great Falls, Steele. District or CoLumBia: Kearney in 1897, Pollard 595, Steele in 1896, Ward in 1871. Virainia: Chain Bridge, Chase 3623; Rosslyn, Topping in 1895; Hampton, McCarthy in 1883; Virginia Beach, Hitchcock 171, Kearney 2018; Dismal Swamp, Chase 3676; Walnut Point, Knowlion in 1897. a HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 91 West Vireria: Tucker County, Greenman 47 (Gray Herb.). Norto Carouina: Wilmington, Kearney 267a; Tarboro, McCarthy in 1883 and 1885; Swanquarter, Ashe in 1898; West Raleigh, Stanton 1283; Biltmore, Bilt- more Herb. 700c. Sours Caro.ra: Charleston, Donaldson in 1888; Isle of Palms, Hitchcock 226; St. Helena Island, Cuthbert in 1905. Georetia: Early County, Harper 1220; Sumter County, Harper 429, 631, 1037; Albany, Tracy 3604. Fiorina: Jacksonville, Curtiss 3609, 5084, Combs 8, 14, Kearney 158; Baldwin, Combs 63, 66; Lake City, Combs 119; Madison, Combs 235, 267; Monticello, Combs 348; Apalachicola, Biltmore Herb. 700b; De Funiak Springs, Combs 437, 458, 473, 475; Bay Head, Combs 645, 646, 647, 654; Chipley, Combs 537, 579, 597; Old Town, Combs 887; Gainesville, Combs 722; Waldo, Combs 695. 703; Eustis, Nash 1761; Manatee, Combs 1297; Arcadia, Combs 1249, 1280; Myers, Hitchcock Lee Co. Pl. 491; Miami, Chase 3900, Hitchcock 728; without locality, Rugel 595. Kentucky: Clays Ferry, Peter in 1834 (Ky. State Univ. Herb.). TENNESSEE: Hollow Rock, Biltmore Herb. 700a. AtaBaMA: Valley Head, Ruth 76; Auburn, Earle & Baker in 1897, Tracy in 1897; Birmingham, McCarthy in 1888; Tuskegee, Carver 72; Mobile, Kearney 20 in part. Mississiprr: Starkville, Chase 4459, 4460, Kearney 82; Waynesboro, Kearney 168; Jackson, Smith in 1885; Batesville, Eggert 126; Biloxi, Tracy 3660, 3762 in part, 3763; Ocean Springs, Kearney 298, Tracy in 1889; Point St. Martin, Tracy 4565; Bay St. Louis, Langlois 34. ArxKansas: Batesville, Coville in 1887; Benton County, Plank 68; without locality, Harvey in 1882. Loursiana: Oberlin, Ball 192. Texas: Dallas, Reverchon 1622; Gill County, Jermy 219; Jacksonville, Plank 18; Clarksville, Plank 4; Trinity Bay, Joor in 1884; Wallisville, Wallis in 1880; Gillespie County, Jermy 787; Luling, Plank 21; Lampasas, Joor 23; Harris County, Joor 34; Paloduro, Gardner 18; Texline, Griffiths 5664; Waller County, Thurow in 1898; Eagle Pass, Havard 84; Fort Concho, Havard in 1881; Moore County, Carleton 422; without locality, Lindheimer 733, Reverchon 22, 32. OxtaHoma: Verdigris River, Blankinship in 1895; False Washita, Palmer 376 in 1868; without locality, Sheldon in 1891. Montana: Glendive, Ward in 1883. Wyomine: Laramie County, A. Nelson 3626, E. Nelson 330, 476; Glen Rock, A. Nelson 8386. Cotorapo: Fort Collins, Crandall & Cowen 550; Denver, Tracy 912; Millers Ranch, Fry 3371; Golden, Rydberg 2508, Shear 756; Meadow Park, Shear 606; La Veta, Shear 819; Canyon City, Shear 980; Trimble Springs, Baker, Earle & Tracy 962, Selig 1264; La Salle, Rydberg 2512. Nevapa: Ash Meadows, Purpus in 1898. New Mexico: Tesuque, Wooton 2936; Grant County, Rusby 445; Pecos, Stand- ley 5289. Arizona: San Francisco Mountains, Leiberg 5732; Tuscon, Towmey 7814; Oak Creek, MacDougal 474; Yavapai County, Rusby in 1883; Beaver Creek, Rusby in 1883; Flagstaff, Toumey in 1894; Turkey Tanks, Wooton 2001. Mexico: State of Jalisco, Palmer 207 and 510 in 1886; Chiapas, Nelson 2975; Orizaba, Botteri 648. Honpuras: San Pedro Sula, Thieme 532. Costa Rica: Buenos Aires, Tonduz 3619. Bermupas: Brown & Britton 358, Moore 2850 (both in Field Mus. Herb.). Triniwap: Without data (Gray Herb.). 92 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 44a, Panicum virgatum cubense Griseb. Panicum virgatum cubense Griseb. Cat. Pl. Cub. 233. 1866. The only specimen mentioned by Grisebach is, ‘‘Wr. a. 1865,” that is, collected in Cuba by Wright in 1865. The type, in the Grisebach Herbarium, bears the number 183 and is labeled “‘Low savannas, Hanabana, May 19.”’ Panicum virgatum obtusum Wood, Bot. & Flor. 392. 1874. ‘“‘N.J.’’ [New Jersey]. The whereabouts of the type, if it be in existence, is not known. The diagnosis ‘‘ Panicle contracted ; spikeletssmaller, not pointed, obtusish,’’ seems sufficiently to indi- cate this subspecies. Panicum virgatum breviramosum Nash, Bull. Tor- rey Club 23: 150. 1896. ‘‘Collected by Dr. Small in clay soil in the pine lands about Augusta, Georgia, where it was common, June 27-July 1, 1895.’’ The type, in Columbia University Herba- rium, is a slender plant with narrow panicles about 12 cm. long and 3 to 4 cm. wide, rather compactly flowered, and as a whole very like Wright’s no. 183 mentioned above. Fig. 81.—P. virgatum cubense. From type specimen. DESCRIPTION. Differs from P. virgatum in having culms more slender than usual in the species, solitary or few in a clump, usually narrow panicles with ascending branches, and. smaller spikelets, 2.8 to 3.2 mm. long, the first glume usually about half the length of the spikelet, acute but usually not acuminate-pointed, the second glume and sterile lemma about equal and but slightly exceeding the fruit, the latter about 2 mm. long. This combination of characters fails to hold throughout. A few of the specimens cited below have open pan- icles, but the small, obtuse spikelets with shorter first glume; others have Fig. 82.—Distribution of P. virgatum cubense. the panicle characteristic of the sub- species but an acuminate-pointed first glume to the small spikelets. The following represent these intermediate specimens: Connecticut: Graves 244; New Jersey: Pearce in 1884; Nort Carouna: McCarthy in 1885; Fiortmpa: Chase 3859, 3860, Hitchcock 743, Hume 37. DISTRIBUTION. Pine woods, the Atlantic Coastal Plain from Connecticut to Florida; also in the Bermudas and Cuba. Connecticut: Groton, Graves 244. New Yorx: Aquebogue, Scribner in 1872 (Hitchcock Herb.). New Jersey: Atsion, Chase 3573; New Durham, Van Sickle in 1895; without locality, Pearce in 1884. PENNSYLVANIA: Philadelphia, Smith 109. MaryYLAND: College Park, Novik in 1907. Vireinta: Ashland, DeChalmot; Portsmouth, Noyes 87. Norra Carouina: Edenton, Kearney 1899; Tarboro, McCarthy in 1885; Wil- mington, Chase 3144, Coville 104, Kearney 267b; Hendersonville, Biltmore Herb. 700e. rou ——__—_ —— ‘ - HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 93 Sours Carona: Elgin, House 2575; Camden, House 2668. GeorciA: Camp Cornelia, Ricker 936; Augusta, Cuthbert 1015, Small in 1895. Frorma: Homosassa, Combs 930; Cedar Key, Combs 774; Jensen, Hitchcock 743; Sanibel Island, Hume 37; Miami, Chase 3859, 3860; Big Pine Key, Simpson 335. Misstsstrrr: Scranton, Pollard 1201; Biloxi, Tracy 3762 in part. Bermupas: Stone in 1888 (Acad. Phil. Herb.). Cusa: Batdbano, Hitchcock 153; Palmer & Riley 1134; without locality, Wright 3873. 45. Panicum havardii Vasey. Panicum virgatum macranthum Vasey, Bull. Torrey Club 13: 26. 1886, not Panicum macranthum Trin. 1826. ‘‘Col- lected by Dr. Havard, in the Guadalupe Mountains, Tex- as.’’ The type, in the Na- tional Herbarium, collected in 1881, is labeled in Vasey’s hand with the data as pub- lished. It consists of the up- per portion of a robust culm, the large and open panicle measuring 40 cm. long and 30 cm. wide, the spikelets nearly 7 mm. long. Panicum havardii Vasey, Bull. Torrey Club 14: 95. 1887. Based upon the type of Panicum virgatum macranthum Vasey. DESCRIPTION. Plants 1 meter or more tall, pale green, glaucous, glabrous throughout; culms robust, sol- itary, erect from creeping rootstocks, simple; sheaths longer than the internodes; ligules dense, about 3 mm. long; blades erect or ascending, 25 to 40 cm. long, 5 to 10 mm. wide, broadest at the base, tapering into long, involute-setaceous tips, some- times pilose on the upper surface at the base; panicles short-exserted, as much as 40 cm. long, half to three-fourths as wide, loosely flowered, the mostly ver- ticillate branches ascending or finally spreading; spikelets 6 to 8 mm. long, about 2 mm. wide, ovate, acuminate, strongly nerved; first glume clasping, half to two-thirds the length of the spikelet, acuminate; second glume slightly shorter than the sterile lemma, both exceeding the fruit, 7 to 9-nerved; fruit 4.5 to 5 mm. long, about 1.8 mm. wide, narrowly ovate, the margin of the lemma inrolled only at base. This apparently rare species resembles P. virgatwm from which it differs in the decumbent base of the solitary culms and in the larger spikelets. Fia. 83.—P. havardii. From type specimen. Fia. 84.—Distribution of P. havardii. 94 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Arroyos and sandhills, western Texas, New Mexico, and northern Mexico. Texas: Guadalupe Mountains, Havard in 1881; without locality, Nealley in 1887. New Mexico: Las Vegas, Vasey in 1880; Roswell, Griffiths 5735. Mexico: Paso del Norte, Pringle 1124 (Hitchcock Herb.). 46. Panicum amarum Ell. Panicum amarum Ell. Bot. 8. C. & Ga. 1: 121.1816. ‘‘ Grows among the sand- hills on the seashore,’’ presumably of South Carolina and Georgia. No specimen of this could be found in the Elliott Herbarium.¢ The description is as follows: ‘‘ Plant very glabrous; leaves thick, glaucous; panicle appressed; glumes acuminate. Root perennial? Stem 2-3 feet high, columnar, thick, nearly an half inch in diameter. Leaves nearly flat, almost coriaceous, the margins very entire; sheaths shorter than the joints, tinged with yellow; the throat contracted, purple; stipules villous. Panicle large, branches all appressed. Flowers very large. Peduncles, which in every other species are very scabrous, and generally hairy, are glabrous and nearly smooth. Calyx 2-flowered, hermaphrodite and male; valves glabrous and tinged with purple. Corolla, valve of the male floret as large as those of the hermaphrodite. * * * Grows among the sandhills on the seashore. Leaves excess- ively bitter.’’ The greater part of this description will be seen to apply equally well to the cespitose species to which the name P. amarum has been applied and to P. amaroides Scribn. & Merr. Scribner and Mer- rill® accepted the cespitose form as the true P. amarum, but the fact that P. amaroides and not the cespi- tose species grows on the coast of North and South Carolina, and espe- cially that it is abundant on the Isle of Palms in Charleston Harbor, Elli- ott’s own locality, casts doubt on the correctness of this identification of Elliott’s species. In the description quoted above ‘‘Panicle appressed” seems to indicate P. amaroides, as does the query after ‘‘root peren- nial.’’ There could be no doubt about the cespitose species (unless one had only a specimen without the base), while in P. amaroides the horizontal rootstock is deep in the sand and the solitary culms are readily detached from it. ‘‘Stem 2-3 feet high” applies much better to P. amaroides, since the allied species is rarely as low as 3 feet. ‘“‘ Leaves excessively bitter” is true of P. amaroides while those of the cespitose species are but slightly or not at all bitter. On the whole the evidence is so strongly in favor of P. amaroides as . Fig. 85.—P.amarum. From type specimen of P. amarum minus Vasey. @ Scribner and Merrill (U. 8. Dept. Agr. Div. Agrost. Cire. 29: 5. 1901) state that “The specimen in the Herbarium of Elliott under this name is a robust form of Panicum virgatum Linn.’’ This name, however, was added later, since it is initialed “H.W. R.”’ [Ravenel.] The original label bears an unpublished name, bU,S. Dept. Agr. Div. Agrost. Circ. 29: 5. 1901, —_— . i HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 95 the plant described by Elliott as P. amarum that we are forced to apply his name to the species named P. amaroides. In the herbarium of the Philadelphia Academy is a specimen of this species bearing two tickets reading respectively “Panicum amarum” and “Elliott S. C.,’’ the former being in the handwriting found in the Elliott Herbarium. While this may not be the type, it would seem to be an authentic specimen. A second specimen of this species in the same herbarium bears a ticket reading ‘‘Georg. Baldw.”’ Panicum amarum minor[us] Vasey & Scribn. U.S. Dept. Agr. Div. Bot. Bull. 8: 38. 1889. ‘Fortress Monroe, Va., and northward, near the coast.’’ The type specimen, in the National Herbarium, was collected by Dr. Vasey at Fort Monroe, Va., in 1879. Panicum amaroides Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Cire. 29:5. f. 1. 1901. Based on P. amarum minus Vasey & Scribn. DESCRIPTION. Plants glaucous and glabrous throughout; culms solitary from the nodes of exten- sively creeping, horizontal rootstocks, simple or occasionally branching from the lower nodes, 30 cm. to 1 meter high; sheaths overlapping but commonly narrow and partially exposing the short, very glaucous internodes; ligules dense and silky, about 3 mm. long; blades ascending or spreading, thick, 10 to 30 cm. long, 5 to 12 mm. wide, broadest at the base, flat below, involute toward the tip, margins smooth; panicles short-exserted or included at base, one-fourth to one-third the height of the plant, or sometimes more, not over 3 cm. wide, mostly few-flowered, the distant, appressed branches bearing scattered, short, appressed branchlets with approximate, short- pediceled spikelets; spikelets 5 to 6.5 mm. long, about 2 mm. wide and as much as 3 mm. thick, narrowly ovate, acuminate, strongly nerved; first glume clasping, two- thirds to three-fourths the length of the spikelet, acuminate, 7 to 9-nerved, the midnerve usually scabrous toward the apex; second glume slightly longer than the sterile lemma, both much exceeding the fruit, 9-nerved, the midnerves scabrous toward the apex; fruit 3.5 mm. long, about 1.5 mm. wide, narrowly ovate. DISTRIBUTION. Sandy seashores and coast dunes, Connecticut to Georgia; also on islands off the coast of Mississippi. An important sand binder. Connecticut: New Haven, Winton in 1887. New York: Long Island, Miller, Young in 1872; Staten Island, Tyler in 1895. New Jersey: Cape May, Burk in 1881 (Hitchcock Herb.). Detaware: Cedar Neck, Com- mons 228 in 1875; Lewes, Hitchcock 227 ; without locality, Canby. Maryann: Bay Ridge, Scribner in 1897. . Vireinia: Fort Monroe, McCarthy in 1883, Vasey in 1879; Ports- mouth, Noyes 66 in 1895; Ocean View, Vasey in 1890; Fig. 86.—Distribution of P. amarum. Virginia Beach, Hitchcock 169, Kearney 2064, Mackenzie 1736, Williams 3089. Norru Carourina: Ocracoke Island, Kearney 2317; Brunswick County, McCarthy; Wilmington, Hitchcock 441; without locality, McCarthy in, 1885. Sourn Carourna: Isle of Palms, Chase 4555. Georata: Tybee Island, Harper 742. Mississirri: Horn Island, Tracy 2854 and in 1897, 96 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 47. Panicum amarulum sp. nov. DESCRIPTION. Plants glaucous but less so than in P.amarum, glabrous throughout, cespitose in large bunches sometimes as much as a meter across, 1 to 2 meters high; culms simple above the base, stout, some- times 1 cm. thick; sheaths mostly overlapping; ligules dense, about 3 mm. long; blades erect or ascending, 20 to 50 cm. long, 5 to 12 mm. wide, broadest at the base, more or less involute, pilose at the base on the upper surface; panicles finally ex- serted, one-third the height of the plant or more, 5 to 10 cm. wide, slightly nodding, compact, densely flowered, the long, fascicled branches erect but more or less spreading at their tips, spikelet-bearing from the base; spikelets short-pediceled, 4.3 to 5.56 mm. long, 1.7 to 2 mm. wide, narrowly ovate, acuminate, strongly nerved; first glume half to two-thirds as long as the spikelet, acuminate, 5 to 7-nerved; second glume slightly longer than the sterile lemma, both 7 to 9-nerved, pointed beyond the fruit; fruit 3 to3.5 mm. long, about 1.4 mm. wide, narrowly ovate, bluntly pointed. Type U.S. National Herbarium no. 592748, collected September 24, 1900, Virginia Beach, Va., by T. A. Williams (no. 3090). This is the species to which the name P. amarum was restricted by Scribner and Merrill, @ Nash, and Hitchcock,¢ when this species and the preceding@ came to be regarded as distinct. The name here proposed is an attempt to associate this species with the name it has so long borne, and also refers to the slightly bitter taste of the foliage. Fic. 87.—P. amarulum. From type specimen. DISTRIBUTION. Sandy seashores and coast dunes, Virginia to Florida and west to Louisi- ana; also in the Bahamas. Vrramnia: Norfolk, Kearney 292; Ocean View, Kearney 1775, Vasey in 1890; Lynn Ha- ven, Iitchcock 440; Virginia Fig. 88.—Distribution of P. amarulum. Beach, Hitchcock 168, Kear- ney 2021, 2063, Mackenzie 1725, Williams 3090. Friorma: Indian River, Curtiss 3578; Cape Malabar, Curtiss in 1879; Lake Worth Inlet, Curtiss 5527; Sea Breeze, Webber 465; Miami, Chase 3899; Cape Florida, aU.8. Dept. Agr. Div. Agrost. Circe. 29: 5. 1901. b Britton, Man. ed. 2. 1048. 1905. ¢ Gray, Man. ed. 7. 104. 1908. @ See note under P. amarum Ell. p. 94. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 97 Chase 3953; Key Largo, Chase 3935, Curtiss in 1884; Upper Metacumbe Key, Chase 3915; Santa Rosa Island, Tracy 6508; without locality, Chapman, Rugel 444. LovursiANa: Battledore Island, Tracy & Lloyd 464. Bawamas: Great Bahama, Britton & Millspaugh 2739; New Providence, Britton & Brace 307, 493 (all in Field Mus. Herb.). Tenera.—Perennials; culms slightly compressed, wiry, the internodes much elon- gated; ligules membranaceous, about 0.5 mm. long; blades linear, at the base narrower than their sheaths; panicles small, narrow, nearly simple; spikelets short-pediceled, glabrous or nearly so; palea of sterile floret a small nerveless scale; fruit elliptic, smooth and shining. Panicles 3 to 8 cm. long; spikelets 2.2 to 2.8 mm. long, pointed....... 48. P. tenerum. Panicles not over 2 cm. long; spikelets not over 1.6 mm. long, blunt. 49. P. stenodes. 48. Panicum tenerum Beyr. Panicum tenerum Beyr. in Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 341. 1834. The citation given is ‘‘Georg. Amer. (in paludibus nemorum cresc.) BEYRICH ms.’’ The label accompanying the type specimen in the Trinius Herbarium reads ‘*Panicum tenerum n. sp. Georgia, in paludibus nemorum,”’ and ‘‘mis. Beyrich 1834” added in Trinius’s writing. A second ticket bears the name ‘‘ Pan. tenerum Beyr.”’ Panicum anceps strictum Chapm. Fl. South. U. 8. 573. 1860. ‘‘Damp sterile soil, Florida.’’ In the Chapman Herbarium at Biltmore is a specimen labeled ‘‘P. anceps strictum from the original locality Apa{lachicola] 1887” and in the Chapman Herbarium in that of Columbia Uni- versity is a similar specimen labeled ‘‘ Panicum anceps L. var. strictum, Southern Flora. Florida,’’ with nothing to indicate whether or not it was collected before the date of publication. In the National Herbarium are two more specimens from Chapman also without date. All the specimens belong to P. tenerum, hence there is no doubt as to the identity of Chapman’s variety, though whether or not the type be in exist- ence can not be determined. This species was described under the name of P. stenodes Griseb. by Vasey, 4 Chap- man, Scribner,¢ and Nash.@ Fic. 89.—P. tenerum. From type specimen. DESCRIPTION. Plants in small tufts from a knotted crown, 40 to 90 cm. high, olivaceous; culms erect, stiff and wiry, producing small, solitary panicles from the upper nodes or remaining simple, glabrous; sheaths much shorter than the internodes, the upper glabrous, the lower sparsely to copiously papillose-pubescent toward the summit with soft, spreading or reflexed hairs; blades 4 to 15 cm. long, 2 to 4 mm. wide, (the uppermost much reduced), erect, firm, drying involute at least toward the summit, pilose on the upper surface toward the base, or the lower sometimes on both surfaces; terminal panicles rather short-exserted, 3 to 8 cm. long, rarely over 5 mm. wide, the short, appressed, subracemose branches bearing rather crowded spikelets throughout their length, the aU.§8. Dept. Agr. Div. Bot. Bull. 8: 25. 1889. b Fl. South. U. S. ed. 3. 583. 1897. eU. 8. Dept. Agr. Div. Agrost. Bull. 7: 64. /. 46. 1897; op. cit. (ed. 2) 52. /. 46. 1900. @Small, Fl. Southeast. U. S. 92. 1903. 41616°—voL 15—10——7 98 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. pedicels usually with a few long hairs at the summit; spikelets 2.2 to 2.8 mm. long, 0.8 to 1 mm. wide, narrowly ovate, pointed; first glume clasping, half as long as the spikelet or more, 1-nerved, glabrous or obscurely strigose toward the summit; second glume and sterile lemma equal, exceeding the fruit, 5 to 7-nerved, glabrous; fruit 1.7 to 1.8 mm. long, about 0.8 mm. wide. DISTRIBUTION. Margins of swamps and wet places in flatwoods and pine barrens near the coast, Georgia to Florida and west fe Py also in the Bahamas and Fig. 90.—Distribution of P. tenerum. uba. Georaia: Cobb, Harper 1045; Ruskin, Ricker 908. Fioripa: Jacksonville, Curtiss 3579 in part, 4035, 5083; Kearney 139; Baldwin, Combs 61, Nash 2249; De Funiak Springs, Combs 476; Apalachicola, Chapman; Pensacola, Combs 530; Avondale, Comvus 487; Jupiter, Curtiss 5576C; Chipley, Combs 574, 619; Grasmere, Combs 1112; Tampa, Garber in 1877; Braidentown, Combs 1266; Myers, Chase 4187, Hitchcock Lee Co. Pl. 492; without locality, Chapman, Garber in 1878, Simpson in 1889. AuaBAMA: Mobile, Kearney 50 in part, Mohr in 1884, 1893, and 1894. Mississipri: Ocean Springs, Kearney 293, Tracy 26; Horn Island, Tracy in 1898. Louistana: Without locality, Hale (Gray Herb.). Texas: Nona, Nealley in 1892; without locality, Nealley in 1884. Banamas: Andros, Brace 7019, 7132 (both in Field Mus. Herb.). Cusa: Pinar del Rio, Wright 3870 in part; Herradura, Hitchcock 154, Tracy 9080. 49. Panicum stenodes Griseb. Panicum stenodes Griseb. Fl. Brit. W. Ind. 547.1864. ‘‘Har. Jamaica!, Pd. [Purdie], in savannahs, Manchester.’’ The type specimen, bearing data as published, is in the Kew Herbarium. Panicum hians Spruce; Griseb. Fl. Brit. W. Ind. 548. 1864, not Ell. 1816. This name is given as a synonym of P. stenodes and credited to ‘‘Pl. Spruce.’’ We have not seen Spruce’s specimen and the name is here referred to P- stenodes on the authority of Grisebach. DESCRIPTION. Plants in small tufts, 25 to 50cm. high, glabrous throughout; culms erect or reclining, very slender and wiry, producing Fig. 91.—P. stenodes. from the middle nodes slender branches about equaling the From Wright’s no. 192 main culm, both this and the branches bearing small, solitary ale Herba- or fascicled panicles from the upper nodes; sheaths very short, ; about 1 to 2cm. long; blades 1 to 4 cm. long, 1 to 2 mm. wide, the upper often reduced to mere points, erect, firm, involute; panicles short-exserted, the lateral often partly included, 1 to 2 em. long, 1 to 2 mm. wide, subracemose; spikelets 1.4 to 1.6 mm. long, about 0.7 mm. wide, elliptic, rather turgid; first glume about half the length of the spikelet, blunt, nerveless or l-nerved; second glume and sterile lemma equal, scarcely exceeding the fruit, 5-nerved; fruit 1.3 mm. long, 0.6 mm. wide. This species is distinguished from P. tenerum by its more slender culms, numerous branches, and smaller panicles of smaller spikelets. ude HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 99 Panicum caricoides Nees,* the type specimen of which, in the Munich Herbarium, was collected by Martius in the province of Para, Brazil, is an allied species. The Martius specimen differs from the West Indian plants in having larger spikelets, 1.8 to 1.9 mm. long, and numerous long, stiff, erect hairs on the pedicels. DISTRIBUTION. Borders of ponds and wet savannas, Costa Rica and the West Indies; also in Brazil. Costa Rica: Buenos Aires, Pittier 10589. Cuspa: Herradura, Baker & Dimmock 4837 (Hitchcock Herb.), Hitchcock 155; without locality, Wright 3871; Hanabana, Wright 192 (Grisebach Herb.). Santo DominGco: Wright, Parry, & Brummel 624. Brazi: Prov. of St. Paul, St. Hilaire 758 (Paris Herb.); Barra, Spruce 1289 (Gray Herb.). Agrostoidia.—Tufted perennials; culms erect, compressed, sheaths more or less keeled; ligules membranaceous, short, sometimes ciliate; blades long and narrow; spikelets lanceolate, glabrous; first glume keeled, scabrous on the keel toward the apex, the second glume and sterile lemma pointed beyond the fruit, more or less keeled, the spikelet thus often appearing laterally compressed, the palea of the sterile floret about half as long as its lemma; fruit elliptic, smooth and shining, a minute tuft of thickish hairs at the apex. The first three species of this group have much the appearance of certain species of Agrostis. Rootstocks present; culms but little compressed; spikelets set obliquely on their appressed pedicels. Panicles open; spikelets 3.4 to 3.8 mm. long (shorter in exceptional specimens)... .. EE 8 ee eee 55. P. anceps. Panicles more or less contracted; spikelets not over 2.8 EE ee ea ee oe ee eee 56. P. rhizomatum. Rootstocks absent; culms strongly compressed with keeled sheaths; spikelets not obliquely disposed. Ligules ciliate; basal leaves half as long as the culm or more; panicle much exceeding the upper leaves. Spikelets not over 2.7 mm., usually 2.5 mm. long, the ' first glume less than half that length; ligules 2 OS ie ee ier 5 | eee 53. P. longifolium. Spikelets 3 to 3.5 mm. long; first glume two-thirds to three-fourths that length; ligule less than 1 hen ote 5 ns dp ee on. a eg 9 54. P. combsit. Ligules erose or lacerate, not ciliate; basal leaves in short tufts, the upper usually nearly equaling the terminal panicle. Fruit stipitate; spikelets 2.5 to 2.8 mm. long, con- MOH CMON Sia ows oo a weg ea OP oe ees a 52. P. stipitatum. Fruit not stipitate; spikelets not conspicuously se- cund. Spikelets 1.8 to 2 mm., in occasional specimens 2.2 mm. long; panicle branches ascending PP I irsictals ata teas cine nc ac dene ns Dlbe, L.. COOLS, Spikelets about 2.5 mm. long; panicle branches GfOCh OF NEARLY. 80. 2 <2 2.2. 4-saeae-c---- 51. P. condensum. a Nees in Trin. Gram. Pan. 149. 1826; Nees, Agrost. Bras, 108. 1829. 100 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 50. Panicum agrostoides Spreng. Panicum agrostoides Spreng. Pl. Pugill. 2: 4.1815. Sprengel first gives an original diagnosis, then cites ‘‘P. agrostidiforme Lam. ill. n. 895. encycl. 4. 738. Habitare videtur in Cayenna. Misit etiam Miithlenbergius e Pensylvania.’’ Then follows an ample description which applies to the plant bearing the name Panicum agrostoides and marked as sent by Muhlenberg which is found in the Willdenow Herbarium. Spren- gel’s herbarium is not segregated from the general herbarium at Berlin, as is Willde- now’s, and no specimen marked Panicum agrostoides from Muhlenberg was found in the general herbarium. Sprengel’s description was doubtless based on the specimen in the Willdenow Herbarium, which is therefore taken as the type, the citation of La- marck’s name @ as a synonym being erroneous. It would seem that the name P. agros- toides was given by Muhlenberg on the specimen sent to Willdenow, since Muhlenberg shortly after ® published this as a new species of hisown.¢ In the Muhlenberg Herbar- ium specimens of both this species and of P. stipitatum are found in the cover marked ‘“Panicum agrostoides M.”’ Panicum rigidulum Bosc; Spreng. Syst. Veg. 1: 320.1825. This is given as a synonym of P. anceps Michx. and is based on P. “‘rigidulum Bosc. (W. herb.).’’ The * specimen in the Willdenow Herbarium is P. agrostoides. Pani- cum rigidulum was described by Nees@ as a new species with Bosc as the author, and based on the specimen in the Willdenow Herbarium. Agrostis polystachya Bosc; Steud. Nom. Bot. ed. 2. 1: 40. 1840. This is given as a synonym of A. composita Poir. No locality is cited. A specimen in the De Candolle Herbarium, collected in Carolina by Bosc, is referable to P. agrostoides. In Fic. 92.—P. agrostoides. the Delessert Herbarium are two specimens labeled by Bose as From type specimen 4 grostis polystachya; one is Panicum anceps and the other is P. in Willdenow Herba- : eit virgatum. Panicum elongatum ramosior|ius] Mohr, Contr. Nat. Herb. 6: 357. 1901. ‘‘AnaBAMA: Damp cultivated ground. Mobile County, Pierce’s Land- ing.’ The type specimen, in the Mohr Herbarium, collected by ‘‘C. Mohr, Oct. 1885,’ is a portion of a plant with an unusually large and open panicle. This species was described by Nashé under the name of P. agrostidiforme Lam. DESCRIPTION. Plants in dense clumps, from a short caudex, with numerous shoots of short leaves at the base, erect, glabrous throughout except as noted, 50cm. to 1 meter or more high; culms rather stout, compressed; sheaths longer than the internodes, keeled, occasion- ally pilose on the sides at the juncture with the blade; ligules erose, about 1 mm. long; blades erect, conduplicate at the base, but flat above or sometimes drying involute, 20 to 50 cm. long, rarely longer, 5 to 12 mm. wide; panicles terminal and axillary, finally long-exserted, 10 to 30 cm. long, rarely longer, usually half to two-thirds as wide but occasionally diffuse and nearly as wide as long, the stiff branches ascending or sometimes spreading at maturity, with more or less divergent, densely flowered branch- lets, commonly from the lower side, the ultimate branchlets and short pedicles a Panicum agrostidiforme Lam., the type of which is in the Lamarck Herbarium, is P. laxum Swartz. b Deser. Gram. 119. 1817. ¢Sprengel does not include his P. agrostoides in the Systema [Syst. Veg. 1: 319. 1825] but gives P. agrostoides Muhl. asasynonym of P. proliferum. @ Acrost. Bras. 163. 1829. e Britt. & Brown, [lust. Fl. 1: 115. /. 249. 1896. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 101 appressed, scabrous, the latter usually bearing at the summit one to several delicate white hairs, these often one-fourth to one-third as long as the spikelet; spikelets 1.8 to 2 mm. long, in occasional specimens 2.2 mm. long, 0.7 to 0.8 mm. wide; first glume hardly half the length of the spikelet; second glume and sterile lemma sub- equal, 5-nerved, the midnerves scabrous at the summit; fruit 1.3 mm. long, 0.6 mm. wide. This widely distributed species is variable in the form of the panicle, occasional specimens approaching P. condensum and others P. stipitatum. The Muhlenberg specimens in the Willdenow and Muhlenberg herbaria are the characteristic form described above. The following specimens have rather turgid spikelets 2 to 2.2 mm. long, more or less crowded on the ascending but not appressed branches and appear to be intermediate between P. agrostoides and P. condensum. These are not cited in the distribution given below. Marne: North Berwick, Parlin 751; Massacnusetrs: Newburyport, Conant; New York: Erastina, Pollard in 1894; Virernta: Boettcher 462; GEorGIA: Leslie, Harper 1730; Fiorta: Jacksonville, Curtiss 5302; Bartow, Combs 1207; Loutstana: Lake Charles, Chase 4410; Texas: Dallas, Reverchon 1083A; Hempstead, Hall 819; without locality, Reverchon 103 in 1879. In the following specimens the spikelets are more or less secund on the branchlets, giving the panicles much the aspect of those of P. stipitatum, but the spikelets are 2 mm. long or scarcely more: Bartlett 1066, Chase 4397, Curtiss 6890, Harper 1239, Plank 24, Riggs, Marshall, Texas. Unusually loosely flowered, open-panicled specimens, such as that named P. elon- gatum var. ramosius, are the following: Chapman, Georgia, Curtiss 6888, Mohr in 1885, Plank 75, Winchell, Alabama. DISTRIBUTION. Wet meadows and shores, Maine to Illinois and Kansas, south to Florida and Texas; also on Vancouver Island and in California. Marne: Orono, Harvey 1240; North Berwick, Parlin in 1896 (Gray Herb.). Massacuusetts: Without local- ity, Conant; Middlesex Fells, Knowlton 910 (Hitchcock Herb.). Connecticut: South Manchester, Hitchcock 129; Huntington, Eames in 1895; Southington, Bissell 5529. Ruope Istranp: Cumberland, Greenman 1804 (Gray Herb.). New York: Cleveland, Rowlee in 1906. New Jersey: Mount Arlington, Mackenzie 2355; Stockholm, Van Sickle in 1895; Atsion, Chase 3529; Mon- mouth Junction, Pearce in 1884. PENNSYLVANIA: Chambersburg, Porter in 1897; Tinicum, Smith 158; Westches- ter, Darlington; without locality, Mc Minn. Onto: Sandusky, Kellerman in 1902 (Univ. Ohio Herb.). Inp1Ana: Miller, Umbach in 1896 (Field Mus. Herb.). InuiNots: Madison, Eggert 227; St. Clair County, Brendel in 1850; southern IIli- nois, Vasey in 1860. Missouri: Lake City, Mackenzie 301; Sibley, Bush 302; Monteer, Bush 5115 in part; Lees Summit, Bush 5234; Dunklin County, Bush 9. Fic. 93.—Distribution of P. agrostoides. 102 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Kansas: Cherokee County, Hitchcock Pl. Kan. 877; Kingman County, Carle- ton 548. DELAWARE: Townsend, Canby in 1891; Talleyville, Commons in 1897. MaryLAND: Lanham, Chase 3469, Hitchcock 2398; Glen Echo, Kearney in 1895; Great Falls, Ball 83. Nortu Carouna: Chapel Hill, Coker in 1908. Sourn Carouina: Aiken, Ravenel in 1869; Orangeburg, Hitchcock 445. GeoraiA: Thomson, Bartlett 1066; Leesburg, Curtiss 6888, 6890; Arlington, Har- per 1239; Augusta, Cuthbert 1128; without locality, Chapman. Fiorina: Old Town, Combs 879; Dunnellon, Combs 915; Lake City, Chase 4286, 4287; Eustis, Nash 1694; Grasmere, Combs 1120; Clarcona, Meislahn 100, 159a; Orange County, Baker 131; Crystal, Combs 1012; Homosassa, Combs 961; Bartow, Combs 1198, 1204; Ellzey, Combs 804; Jenkins, Haton 242; Miami, Eaton 164 in part, 165; without locality, Rugel in 1842. Kentucky: Lexington, Short in 1835 (Gray Herb.). TENNESSEE: Knoxville, Ruth 65. ALABAMA: Eufaula, McCarthy in 1888; Cullman County, Eggert in 1897; Mobile County, Mohr in 1885; without locality, Chapman, Winchell. Mississippi: Starkville, Chase 4445, Kearney 74; Nicholson, Kearney 372 in part; Panola County, Eggert in 1896; Horn Island, Tracy 2036. ARKANSAS: Fulton, Bush 1023; Texarkana, Heller 4235. Louisiana: Coushatta County, Ball 124; Alexandria, Ball 169; Rayville, Ball 20 in part; Natchitoches, Ball 156; Oberlin, Ball 225; Chalmette, Tracy 7651; Shreveport, Ball 111; Pointe a la Hache, Langlois in 1880; Lake Charles, Chase 4397. Texas: Texarkana, Plank 24; Beaumont, Plank 24; Jacksonville, Joor in 1884; Rusk, Plank 75; Marshall County, Riggs 18; Columbia, Bush 1398; Harvester, Thurow in 1898; Hockley, Thurow in 1889; Tyler, Reverchon 2222; Luling, Plank 22; “60 miles south of Fort Scott,’’ Palmer in 1863. OKLAHOMA: Poteau, Hitchcock in 1903 (Hitchcock Herb.); without locality, Shel- don in 1891. BritisH CotumstiaA: Sproat Lake, Vancouver Island, Macown 135, 29348. CaLtrorniA: On the Sacramento, Wilkes Expl. Exped. 51. Panicum condensum Nash. Agrostis purpurascens Bert.; Steud. Nom. Bot. ed. 2. 1: 42. 1840, not Swartz, 1788. “Domingo” is cited, but no description isgiven. Bertero’s specimen, labeled “‘ Agros- tis purpurascens loc. aquosis. §. Domingue Bertero. M. Balbis 1821,’”’ in the Berlin Herbarium, is P. condensum. Panicum contractum Trin.; Steud. Nom. Bot. ed. 2. 2: 254. 1841. ‘‘Trin. mpt. Guadal. Domingo” is cited, but no description isgiven. The specimen in the Berlin Herbarium, bearing this name in Trinius’s writing and labeled “S. Do- mingo, locis aquosis. Balbis misit 1824,’’ is P. condensum. Panicum condensum Nash in Small, Fl. Southeast. U.S. 93. 1903. On page 1327 in the list of new genera and species the following citation is given: “‘Type, Curtiss,’ Second Distr. Pl. So. U. 8., no. 5576, in Herb. C. U.”? The type, in Columbia University Herbarium, is a single plant lacking the base. Curtiss’s no. 5576 was collected October 16, 1895, near Jacksonville, Fla. Fig. 94.—P. condensum. From type specimen. a These were collected on the same date, but have been distributed under two num- bers. 6 Thurber [Torr. Bot. Wilkes Expl. Exped. 1: 480. 1874] calls attention to the fact that this collection was far out of the known range of P. agrostoides. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 103 DESCRIPTION. Plants like P. agrostoides in habit, culms on the average rather taller, sometimes geniculate at base; sheaths appressed-pubescent along the sides toward the summit or glabrous; blades often sparsely pilose on the upper surface at the conduplicate base; panicles terminal and axillary, 10 to 25 cm. long, rarely over 5 cm. wide, the long, solitary or fascicled branches erect, naked at the base, with ap- pressed branchlets bearing crowded spikelets on short, scabrous, but not pilose pedicels, or these occasionally with one or two erect hairs; spikelets 2.2 to 2.5 mm. long, about 0.8 mm. wide, turgid; first glume slightly more than half the length of the spikelet, acuminate; second glume slightly Fic. 95.—Distribution of P. condensum. longer than the sterile lemma, both acuminate, the tips slightly spreading, scabrous on the midnerves toward the apex; fruit 1.4 to 1.5 mm. long, about 0.7 mm. wide. This species is closely related to P. agrostoides, from which it is distinguished by the condensed panicle of slightly larger spikelets. In occasional specimens, as Combs 9644, 973, Hitchcock 174, and Meislahn 159, the panicle branches are ascending, the panicle not contracted, thus approaching P. agrostoides. DISTRIBUTION. Borders of streams and ponds and in wet places, Pennsylvania to Florida and west to Texas, mostly near the coast; also in the West Indies. New Jersey: Cape May County, Van Pelt in 1909 (Acad. Phil. Herb.). PENNSYLVANIA: Chambersburg, Porter in 1897. DELAWARE: Smyrna, Long in 1908 (Acad. Phil. Herb.). District or CotumBia: Steele in 1896. Vrrerta: Brick Haven, Steele in 1896; Four-Mile Run, Chase 2673; Virginia Beach, Hitchcock 174. North Carouina: Wilmington, Hitchcock 444. Soutn Caro.urna: Isle of Palms, Chase 4527. Fioriwa: Jacksonville, Curtiss 5298, 5576, 5576A; Titusville, Chase 4014; Orange County, Baker 18, Combs 1144; Clarcona, Meislahn 159; Homosassa, Combs 957, 9644, 973; Crystal, Combs 1011; Jensen, Hitchcock 747; Miami, Hitchcock 172, 695, 699; Braidentown, Simpson 7093; St. Vincents Island, Chapman, Tracy 6444. Mississrepi: Horn Island, Tracy in 1893 (Hitchcock Herb.). Texas: Dallas County, Reverchon 2368; Waller County, Thurow in 1898; without locality, Lindheimer 728. Banamas: Nassau, Curtiss W. Ind. Pl. 174. Cusa: Without locality, Wright 3862 in part. Granv CayMan: Hitchcock in 1891 (Mo. Bot. Gard. Herb.). LeEwarD IsLtanps: Guadeloupe, Duss 3919. 104 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 52. Panicum stipitatum Nash. Panicum elongatum Pursh, Fl. Amer. Sept. 1: 69. 1814, not Salish. 1796. ‘‘New ” Jersey to Virginia.’’ Among the Pursh plants in Kew Herbarium are two sheets labeled ‘‘Panicum elongatum Pursh. Fl: Amer.,’’ on one of which is a robust specimen and a ticket bearing an unpub- lished herbarium name, and also the name “‘elongatum’”’ and the word ‘‘Delaware.’’ This is taken as the type. On the second sheet are two slender specimens, one of this species and the other of P. agrostoides. Panicum agrostoides elongatum Scribn. Tenn. Agr. Exp. Sta. Bull. 7: 42. pl. 9. f. 34. 1894. Based on P. elongatum Pursh. Panicum stipitatum Nash in Scribn. U.8. Dept. Agr. Div. , sie Agrost. Bull. 1'7 (ed. 2): 56. f. 352. May 22,1901. Based on Bee i Re el P. elongatum Pursh, 1814, not Salisb. 1796. ‘‘Nash, in Britt. Manual, 83, 1901,’’ is here cited as the place of publication, but this must have been taken from proof sheets, since the Manual was not published until after August 24, 1901, that being the date given aiter the preiace. DESCRIPTION. Plants like P. agrostoides in habit, often purple-tinged throughout; culms on the average stouter, strongly compressed; sheaths much overlapping, blades usually equaling or exceeding the terminal panicles, often scabrous on the lower surface; panicles usually several to a culm, sometimes as many as five axillary panicles, com- monly dark purple, short-exserted, 10 to 20 cm. long, one-third to half as wide, densely flowered, the numerous stiff branches ascending, with numerous divaricate branchlets mostly from the lower side and beginning at the base, bearing crowded, subsecund spikelets, the short, scabrous pedicels only rarely with one or two erect hairs; spikelets 2.5 to 2.8 mm. long, about 0.7mm. wide, often curved at the point; first glume about half the length of the spikelet; second glume and sterile lemma subequal, scabrous on the midnerve at the acuminate apex; fruit about 1.5 mm. long, about 0.6 mm. wide, short-stipitate. Typical specimens of this species are characteristic and readily distinguished from P. agrostoides, but less densely panicled forms, with smaller spikelets approach that species. Such are the following: Bush 3658, Chase 4497, Cocks 3008. In these specimens the fruit is stipitate, for which reason they are referred to this species, DISTRIBUTION. Moist soil, Connecticut to South Carolina, west to Kentucky, Missouri, and Texas. Connecticut: Lyme, Graves 236. New Jersey: Camden, Scribner 39 in part; Oradell, Mackenzie Fig. 97.—Distribution of P. stipitatum. 1893. PENNSYLVANIA: Chambersburg, Porter in 1897; Westchester, Darlington in 1827. Outro: Lancaster, Kellerman 6800. Missourt: Williamsville, Bush 3658. DELAWARE: Greenbank, Commons 25 and 305 in 1884. MaryLAnpb: Hyattsville, House 1443. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 105 District or CotumBiA: Greene in 1908, Steele, Vasey. VirGiniA: Four-Mile Run, Chase 2679; Goshen, Steele in 1904. West Vrrernta: ‘Grafton, Guttenberg in 1879, Smith in 1879. Nortx Carouna: Newbern, Kearney 2249; eastern North Carolina, McCarthy in 1885; Henderson County, Smith in 1881. Soutn Carouia: Aiken, Ravenel in 1869; Isle of Palms, Hitchcock 225. Kentucky: Bell County, Kearney 380;2 Harlan County, Kearney 380.4 TENNESSEE: Hollow Rock, Biltmore Herb. 808a, Eggert in 1897; Cocke County, Kearney 969; Madison County, Bain in 1892. ALABAMA: Scottsboro, Chase 4497. Lourstana: Calhoun, Ball 70; Calcasieu, Cocks 3008. Texas: Texarkana, Plank 77. 53. Panicum longifolium Torr. Panicum longifolium Torr. Fl. North. & Mid. U. S. 149. 1824. ‘In the pine barrens of New Jersey. * * * For specimens * * * I am indebted to Mr. James Goldy.” The type, in the Columbia University Herba- rium, is a small specimen 35 cm. high, with an open, few- flowered panicle. Panicum anceps pubescens Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8:37. 1889. ‘‘Mobile, Ala. (Dr. Mohr).” There is no specimen in the National Herbarium marked with this name in Vasey’s writing, but there are two duplicate specimens of P. longifolium with pubescent sheaths and blades, collected by Dr. Charles Mohr, Mobile, Alabama, both labeled Panicum anceps and bearing an unpublished varietal name in Vasey’s writing. These agree with Vasey’s brief description and are doubtless the basis of this name. Vasey applied this unpub- lished varietal name also to specimens of P. anceps and to one of P. rhizomatum, but Dr. Mohr’s, being the only specimen cited, is taken as the type. ° Panicum pseudanceps Nash, Bull. Torrey Club 25: 85. 1898. ‘‘Collected by Mr. J. H. Simpson in Florida in 1889.”” The type, in Nash’s herbarium, consists of two plants 58 and 60 cm. high, the panicles less open than those of the Goldy specimen mentioned above. Fic. 98.—P. longifolium. From type specimen. DESCRIPTION. Plants in dense tufts, 35 to 80 cm. high, usually surrounded by basal leaves nearly half as long as the culm; culms rather slender, much compressed, stiff, glabrous; sheaths mostly shorter than the internodes except at the base, keeled, usually hairy on the sides at the juncture with the blade, otherwise glabrous or villous toward the summit, sometimes densely so; ligules fimbriate-ciliate, 2 to 3 mm. long, the cilize usually at the sides only, not meeting at the back; blades erect or sometimes recurved or tortuous, conduplicate at base, flat above or somewhat involute, 8 to 40 cm. long, 2 to 5 mm. wide, pilose on the upper surface toward the base, sometimes also on the lower surface; lateral panicles few or none; terminal panicles finally long-exserted, much exceeding the leaves, 10 to 25 cm. long, usually half to two-thirds as wide, but sometimes rather contracted, the distant, slender branches solitary or fascicled, ascending, usually nak'd at base, bearing short, appressed, rather closely flowered branchlets, these and the pedicels scabrous, the latter sometimes with a few hairs at the summit; spikelets 2.4 to2.7 mm. long, about 0.7 mm. wide; first glume two-fifths ta scarcely half the length of the spikelet, acute; second glume slightly longer than the sterile lemma, both keeled, usually spreading at the tip, scabrous on the mid- nerve at the apex; fruit 1.6 mm. long, 0.6 mm. wide. a These two collections were distributed under the same number. 106 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. A specimen from Nicholson, Mississippi, Kearney 380, with spreading panicle branches and spikelets only 2 mm. long is doubtfully referred here. DISTRIBUTION. Moist sandy ground, Rhode Island and Connecticut to Florida and west to Texas, mostly near the coast. Connecticut: Groton, Bissell 11596, Graves 248, 256. RuopeE Is~tanp: Shannock, Moore in 1896. New York: New York, Bicknell in 1896; Long Island, Miller in 1873 (both in Hitchcock Herb.). New Jersey: Atsion, Chase 3530; without locality, Holmes in 1890. PENNSYLVANIA: Philadelphia, Saunders in 1898, Smith. DELAWARE: Ellendale, Commons 224, 343, 344. MaryLanp: Berlin, Novik 421. District or CoLuMBIA: Kearney in 1897, Steele in 1897 and 1898. Virernia: Cape Henry, Hitchcock 446; Dismal Swamp, Chase 3662; Virginia Beach, Kearney 2025. Norra Carona: Newbern, Kearney 2242; Wilmington, Biltmore Herb. 3627, Chase 4564, IMitchcock 447; eastern North Carolina, Ashe; West ; Raleigh, Stanton 1270; middle Fig. 99.—Distribution of P. longifolium. North Carolina, Ashe. Fiorina: Jacksonville, Curtiss 5576B; Milton, Chase 4318; Bayhead, Combs 644; Chipley, Combs 571; Gainesville, Chase 4225; Crystal, Combs 1010; Cuetee Combs 1154, 1168; Bartow, Combs 11804, 1235. ALABAMA: Tuskegee, Carver 100, Cullman, Eggert in 1897, Mohr in 1886: Mobile, Mohr. Mississippi: Biloxi, Chase 4346, Kearney 220, Tracy 3627, 3860, 6507a; Fontain- bleau, Tracy 3858, 3859; Ship Island, Tracy 4563; Waynesboro, Kearney 116. Lours1ana: Without locality, Hale (Gray Herb.). Texas: Jefferson, Plank 31; without locality, Nealley in 1885, Wright. 54. Panicum combsii Scribn. & Ball. Panicum combsii Scribn. & Ball, U.S. Dept. Agr. Div. Agrost. Bull. 24: 42. f. 76. 1901. ‘‘Type collected by Robert Combs, No. 583, for whom the species is named, in damp, fertile flat woods at Chipley, Washington County, Florida, August 20, 1898.” The type specimen, in the National Horbarium, is a tuft of several culms with mature panicles. DESCRIPTION. Plants like P. longifolium in habit; leaves somewhat more clustered at base; sheaths glabrous or pubescent along the margin toward the summit; ligules less than 1 mm. long, Fic. 100.—P. combsii, From S°metimes a few longer hairs at the margin; blades aver- type specimen. aging shorter, rarely 25 cm. long, glabrous or pilose on the upper surface at the base, usually sparsely so; lateral panicles wanting or but one or two, terminal panicles 12 to 20 cm. long, two-thirds to three-fourths as wide, few-flowered, the slender, scattered branches ascending; HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 107 spikelets 3 to 3.5 mm. long, 0.7 to 0.8 mm. wide, acuminate; first glume two-thirds to three-fourths the length of the spikelet; second glume and sterile lemma subequal, usually scabrous on the midnerve, much exceeding the fruit; fruit 1.8 to 2 mm. long, 0.6 to 0.8 mm. wide. This species is closely related to P. longifolium from which it may be dis- tinguished by its shorter blades, longer spikelets, and usually by the lack of pubescence. DISTRIBUTION. Margins of ponds and wet woods, Geor- gia to Florida, and west to Louisiana. Fig. 101.—Distribution of P. combsii. Gzorai: Tifton, Harper 1679; Douglas, Harper 2014, Huntington, Harper 1081. Fioripa: Pensacola, Curtiss 6919; Argyle, Curtiss 6925A; Chipley, Combs 583; without locality, Chapman. ALABAMA: Gateswood, Tracy 8408. Misstssrepr: Biloxi, Tracy 4568 in part. Lourstana: Lake Charles, Chase 4434. 55. Panicum anceps Michx. Panicum anceps Michx. Fl. Bor. Amer. 1: 48. 1803. ‘‘Has. in Carolinae herbosis humidis sylvaticis.”. The type specimen, labeled ‘‘Panicum anceps, Hab. in her- bosis humidis Carolinae, Virginiae, Georgiae,”’ is in the Michaux Herbarium. Panicum rostratum Muhl.; Willd. Enum. Pl. 1032. 1809. ‘‘ Habitat in sylvaticis humidis Pensylvaniae, Carolinae.” The type specimen, in the Willdenow Herbarium, is marked ‘‘P. rostratum Am. Boreal. Muhlenberg.” Muh- lenberg later@ published P. rostratum asa new species. The specimen in the Muhlenberg Herbarium is the same species as the one sent to Willdenow. Agrostis nutans Poir. in Lam. Encycl. Suppl. 1: 255. 1810. ‘‘Cette plante a été recueillie dans la Caroline, par M. Bosc.” We have not seen this specimen, but the de- scription applies well to P. anceps, to which species Poiret later > refers his A. nutans. Panicum nutans Desv. Opusc. 93.1831. Based on ‘‘ Agrostis nutans Poir. Enc. suppl.” Fig. 102.—P. anceps. From Panicum anceps angusta[um] Vasey, U. 8. Dept. Agr. Div. Sf alo am Bot. Bull. 8: 37. 1889. ‘‘Texas (G. C. Nealley).” The type specimen, in the National Herbarium, is a tall plant with an immature panicle, the long blades involute. Panicum anceps densiflorum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8:37. 1889. “Texas (J. F. Riggs).”” The type specimen, in the National Herbarium, was collected November, 1884, Marshall, Texas, by J. F. Riggs, no. 91. It consists of the upper por- tions of two robust plants, the panicle branches densely flowered. DESCRIPTION. Plants in tufts of few to many culms, 50 cm. to 1 meter or more high, with numerous stout, scaly rootstocks; culms usually robust, not strongly compressed, glabrous; sheaths usually shorter than the internodes, glabrous to densely papillose-pilose, a Descr. Gram. 121. 1817. b In Lam. Encycl. Suppl. 5: 539. 1817. 108 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. especially at the summit; ligules membranaceous, less than 1 mm. long; blades erect, flat or but slightly conduplicate at base, 20 to 50 cm. long, 4 to 12 mm. wide, pilose and usually ciliate on the upper surface toward the base, otherwise glabrous, scabrous or more or less pubescent on the upper and occasionally on the under surface; panicles terminal only, or narrow, long-peduncled panicles produced from the upper sheaths, the terminal ones finally long-exserted but often nearly equaled by the long blades, 15 to 40 cm. long, usually half to two- thirds as wide (or occasionally narrow), the long, slender, remote, branches somewhat spreading, bearing short, mostly appressed, distant or approxi- mate branchlets with rather crowded, somewhat curved, subsecund spikelets set. obliquely on their short, appressed, scabrous pedicels, the first glume to- ward the main axis, the axes and branches scabrous, usually a few hairs in the axils; spikelets 3.4 to 3.8 mm. long, 1 to 1.2 mm. wide (occasional specimens with smaller spikelets); first glume one-third to half the length of the spike- let, acute; second glume and sterile lemma subequal, forming a beak beyond the fruit, the tips open at maturity; fruit 2 to 2.2 mm. long, about 1 mm. wide. This species is variable in the pubescefice on sheaths and blades and somewhat so in the panicle. The short secondary branchlets may be distant, giving the long branches the appearance of interrupted racemes, or approximate, producing densely flowered branches as in Vasey’s variety densiflorum. A comparatively few “specimens occur with spikelets only 3 to 3.2 mm. long. Such are the following, which, being nearly glabrous plants with open panicles, are referred here, though in the smaller spikelets they approach the next species: Ball 228, Chase 4201, 4393, 4554, Cocks 3001, Combs 717, 878, 1398, Kearney 376, Mohr in 1885, Tracy 4620, 8414. Fig. 103.—Distribution of P. anceps. DISTRIBUTION. Moist sandy soil, New Jersey to Florida and west to Kansas and Texas. New Jersry: Stockton, Fisher in 1897. PENNSYLVANIA: Philadelphia, Smith; Lancaster, Porter in 1861. InumNors: Oquawka, Patterson; Mount Carmel, Waite in 1887. Missourt: Monteer, Bush 5114; St. Louis, Eggert in 1886. Kansas: Cherokee County, Hitchcock Pl. Kan. 876. DELAWARE: Carpenter Station, Commons 225; Greenbank, Commons 303. Marytanp: West Chevy Chase, Chase 2585; Cabin John, Steele in 1896. District or CoLtumsBiA: Blanchard in 1891, Chase 2996, House 257, Kearney 16, Vasey in 1874, Ward in 1878. Viremia: Fairfax County, Chase 3631; Arlington, Dewey 53; Norfolk, Kearney 1748; Virginia Beach, Britton in 1895, Hitchcock 228. Norta Carortna: Madison County, Biltmore Herb. 5839b; West Raleigh, Coit 1301, Stanton 1282; Dunns Mountain, Small in 1894; eastern North Caro- lina, McCarthy in 1885; Wilmington, Kearney 274. Sours Carortna: Aiken, Ravenel in 1869; Orangeburg, Hitchcock 448, 449; Isle of Palms, Chase 4554. Groraia: Thomson, Bartlett 903; Whitfield County, Harper 369; Stone Moun- tain, Hitchcock 230; Leesburg, Curtiss 6887; Augusta, Cuthbert in 1904. Fioriwa: Madison, Combs 282; Quincy, Combs 415; Tallahassee, Combs 381, Kearney 87, Nash 2529; De Funiak Springs, Combs 470; Chattahoochee, Cur- 7 HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 109 tiss 5936; Apalachicola, Biltmore Herb. Dist. Dupl. Chapman 696b; Monti- cello, Combs 320; Homosassa, Combs 935, 972; Lake City, Chase 4284; Crystal, Combs 1021; Gainesville, Chase 4201, Combs 717; Old Town, Combs 878; Chipley, Combs 560, 586; Grasmere, Comds 1073; Orange Bend, Chase 4095; Bartow, Combs 1206; Tampa, Combs 1398; Fort Lauderdale, Eaton 341; without locality, Chapman. Kentucky: Bell County, Kearney 372 in part. TENNESSEE: Bristol, Hitchcock 229; Lavergne, Biltmore Herb. 696a; Knoxville, Ruth in 1895 and 1898; Cocke County, Kearney 965; Grainger County, Smith in 1880. AtaBAMaA: Auburn, 7racy in 1897; Birmingham, McCarthy in 1888; Gadsden, McCarthy in 1888; Nesheka, Carver 7; Cullman, Mohr in 1885. Misstssrerr: Nicholson, Kearney 376; Taylorville, Tracy 8414; Waynesboro, Pollard 1228; Tupelo, Tracy 1535; Starkville, Chase 4464, Kearney 20 in part; Biloxi, Tracy 4620 and in 1893; Bay St. Louis, Langlois in 1883; Pass Christian, Langlois in 1882. ArKaNnsas: Benton County, Plank 65; Miller County, Eggert in 1896; Harvey in 1880, Loutstana: Natchitoches, Ball 166; Rayville, Ball 8; Coushatta County, Ball 125; Calhoun, Ball 46; Oberlin, Ball 228; West Carroll Parish, Moseley in 1903; Lake Charles, Chase 4393; Calcasieu, Cocks 3001. Texas: Texarkana, Heller 4082, Plank 80; Columbia, Bush 1298; Rusk, Plank 76; Beaumont, Plank 25; Marshall, Riggs 91; Waller, Thurow in 1898; Ennis, Smith in 1897; Industry, Wurzlow in 1894; without locality, Nealley in 1887; Reverchon 106. OxtaHoma: On the False Washita, Palmer 380 in 1868; Kingfisher County, Carleton in 1891; Choctaw Agency, Bigelow in 1853-4. 56. Panicum rhizomatum sp. nov. DESCRIPTION. Plants like P. anceps in habit; the culms less robust, the scaly rhizomes slender and more numerous; leaves more or less clustered toward the base, the sheaths, except the lowermost, shorter than the internodes, densely to sparsely villous along the margin and toward the summit, a dense ring of pubescence at the juncture with the blade; ligules nearly obsolete; blades erect or the lower commonly spread- ing, 10 to 40 cm. long (usually not over 30 cm.), 5 to 10 mm. wide, pubescent on both surfaces or sometimes glabrous except on the upper surface toward the base; terminal panicles long-exserted, the usually numerous smaller ax- illary ones short-peduncled or partially included, 10 to 25 Fic. 104,.—P. rhizomatum. ©™- long, usually less than one-third as wide, more or less From type specimen. contracted and densely flowered, rather more compound than in P. anceps, the distant primary branches ascend- ing, bearing numerous branchlets 1 to 3 cm. long, these with appressed, short, approximate branchlets, with crowded spikelets set obliquely on their short, appressed pedicles as in P. anceps, but hardly at all secund; spikelets 2.4 to 2.8 mm. long, about 1 mm. wide; first glume one-third to scarcely half as long as the spikelet, acute; second glume and sterile lemma subequal, beaked as in P. anceps but less strongly so, but little exceeding the fruit; fruit 1.9 mm. long, 0.9 mm. wide. Type U.S. National Herbarium no. 592752, collected August 18, 1905, Orangeburg, S. C., by A. S. Hitchcock (no. 450). This species was referred by Scribner in the herbarium to P. anceps pubescens Vasey and was distributed under this name by Nash and others. It is distinguished from P. anceps by the somewhat contracted, more densely flowered panicles of smaller 7 spikelets and by the shorter leaves more or less crowded at the base. In occasional specimens the panicle is rather open but less so than in P. anceps; but the following three specimens, having all the other characters of P. rhizomatum, have panicles as open as those of P. anceps, the small spikelets secund as in that species, and appear to be intermediate. FLortmpaA: Bay Head, Combs 648; Orange County, Baker 40; Mississreri: Starkville, Kearney 61; Waynesboro, Kearney 108. 110 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Moist sandy woods and savannas, Virginia to Forida and west to Texas. Virernta: Lynn Haven, Hitchcock 175; Portsmouth, Noyes 67 in part. Fig. 105.—Distribution of P. rhizomatum. NortH Carona: Wilmington, Chase 4560, Hitchcock 310. SoutH CaRoLina: Orangeburg, Hitchcock 450; Isle of Palms, Hitchcock 104; with- out locality, Curtiss in 1875. Georaia: Savannah, Kearney 181; Dock Junction, Ricker 962; Camp Cornelia, Ricker 933. Froriwa: Jacksonville, Curtiss 5289, 5747, Kearney 138; Duval County, Curtiss 3579 in part; Madison, Combs 259; Old Town, Combs 884; Lake City, Combs & Rolfs 120, 192; Waldo, Combs 700; Gainesville, Chase 42014, 4220, 4271; Citrus, Bystra in 1906; Titusville, Chase 3984; Sanford, Nash 2258; Grasmere, Combs 1051, 1125; Eustis, Nash 1713; Ellzey, Combs 820; Dunnellon, Combs 919; Bay Head, Combs 633, 648; Tampa, Combs 1395; Avondale, Combs 496; Bartow, Combs 1192, 1217a; Braidentown, Combs 1252, 1264, 1303, Tracy 7093, 7105; Myers, Chase 4191, Hitchcock Lee Co. Pl. 490. AtaBaMa: Without locality, Mohr in 1878. Mississippi: Centerville, Tracy 4564; Nicholson, Kearney 358; Biloxi, Kearney 235, Tracy 3603, 3626, 4619, 4621, and in 1889; Ocean Springs, Forkert 33, Pol- lard 1106; Bay St. Louis, Langlois in 1883; Pass Christian, Langlois in 1880 and 1882. Lourstana: Natchitoches, Ball 137. Texas: Pierce, Tracy 7405; without locality, Nealley. Laxa.—Plants mostly perennial, apparently annual in P. lavum and P. pilosum, more or less decumbent at base and rooting at the lower nodes, creeping in P. polygonatum, commonly not decumbent in P. hians and P.cupreum; culms compressed, glabrous; ligules membranaceous, very short or wanting; spikelets short-pediceled, more or less clustered, not over 3 mm. long, glabrous, the pale& of the sterile floret, except in P. polygonatum and P. longum, becoming more or less enlarged and indurated, expanding the spike- let at maturity; fruit mostly elliptic, minutely papillose-roughened, rather thin in texture. Spikelets pointed, not expanded at maturity by an enlarged sterile palea. Nodes densely pubescent; spikelets not over 1.5 mm. Ifovrh S PRS Ay bE aS emer ete a> ey ee 58. P. polygonatum. Nodes glabrous; spikelets 2.5 mm. long.................. 57. P. longum. Spikelets blunt, expanded at maturity by the enlarged sterile palea. Panicle branches subracemose, the spikelets secund; blades at least 5 mm. wide; the enlarged sterile palea not conspicuous. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 111 Rachises bearing slender bristles (these wanting in exceptional specimens); blades broadest near the cordate or truncate base; nodes usually SIN a hie tes ms ha 9 Bn Ve i ao 59. P. pilosum. Rachises without bristles; blades narrowed toward Gio bane; nodes plabrows:.2.05..'......-.25..- 60. P. laxum. Panicle branches not racemose; blades scarcely wider than their sheaths; the sterile palea conspicuously enlarged. Spikelets 3 mm. long, congested; panicles dark pur- ro RET Eee ee a oP ee 63. P. cupreum. Spikelets not over 2.4 mm. long; panicles green or pale. Panicle branches spikelet or branchlet-bearing along the upper half or toward the ends Gy areueoae sch. SU. pee APA 62. P. hians. Panicle branches branchlet-bearing throughout their length or nearly so...............-- 61. P. exiguiflorum. 57. Panicum longum nom. noy. Panicum munitum Trin.; Steud. Nom. Bot. ed. 2. 2: 260. 1841. This nomen nudum is credited to ‘‘Trin. mpt. Mexico.’? A specimen of P. longum collected near Jalapa by Schiede and Deppe, no. 674, and listed by Schlechtendahl¢ as num- ber 898 ‘‘Panicum proximum piloso Sw.,’’ was examined at Halle. The sheet is marked ‘“‘Panicum munitum”’ by Trinius, and may be the type of this name, but since the species was not described by Trinius P. longum is here based on Scribner’s type, Pringle 8195. Panicum pilosum macranthum Scribn. U. 8. Dept. Agr. Div. Agrost. Circ. 19: 1. 1900, not P. macranthum Trin. 1826. ‘‘Swamps near Jalapa, State of Vera Cruz, altitude 1,230 m., May 21. C. G. Pringle, No. 8195. 1899.’’ The type, in the National Herbarium, is a slender plant 150 cm. high. DESCRIPTION. Plants perennial, ascending or spreading from a more or less geniculate base; culms rather robust, 1. 2 to 2 meters long, simple or sparingly branching from the lower nodes, glabrous, the nodes glabrous; sheaths nearly as long as the internodes or overlapping, papillose or papillose-hirsute (the greater number of papille without hairs); ligule about 0.5mm. long; blades ascending or spreading, 10 to 25 cm. long, 6 to 12 mm. wide, linear, scarcely narrowed to the rounded base, very sparsely papillose-pilose on the upper surface, glabrous beneath or with a few papille; panicles short-exserted or included at the base, 20 to 25 cm. long, 2 to 5 cm. wide, the branches raceme-like, the lower soli- Fie. 106.—P. longum. From tary, distant, the middle and upper opposite or verticillate, type specimen of P. pilosum the rachisesand upper part of the main axis with stiff hairs macranthum Scribn. - J * ° slightly exceeding the short-pediceled, secund, approxi- mate, but scarcely crowded spikelets; spikelets mostly in pairs, 2.5 mm. long, 0.9 mm. wide, pointed, strongly nerved, the nerves minutely scabrous toward the summit; first glume about half the length of the spikelet, pointed; second glume shorter than the sterile lemma, both slightly exceeding the fruit, the sterile palea a7 a? @ Linnaea 6: 35. 1831. bb CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. obsolete; fruit 2.1 mm. long, 0.8 mm. wide, acute, thin in texture, scabrous toward the summit. This species seems to be more nearly related to P. polygonatum than to any other, for which reason it is placed, though somewhat doubtfully, in this group. DISTRIBUTION. Swamps, Mexico. Mexico: Jalapa, Pringle 8195, Schiede & Deppe 674 (Halle Herb.). 58. Panicum polygonatum Schrad. Panicum polygonatum Schrad. in Schult. Mant. 2: 256. 1824. “In Brasilia, ad ripas fluvit Ithéos. Princeps Sereniss. Maxim. Neowidens.’’ The type, in the herba- rium of the Botanical Garden in St. Petersburg, has a large pan- icle, pubescent nodes, and a few bristles on the rachis. Panicum potamium Trin. Gram. Pan. 239. 1826. ‘Brasil. (Lanesporrr).’’ The type, in the Trinius Herbarium, is from near Mandioca, Brazil, collected by Langsdorff ‘‘in udis aquosis puris.”’ This name is spelled ‘‘P. potamicum” by Steudel.¢ Fig. 107.—P. polygo- Panicum hydrophilum Trin.; Nees, Agrost. Bras. 208. 1829. natum. Fromtype This is given asa synonym under P. dubium Lam., the specimen ye AS referred to as ‘‘(Langsdorff.— V. in Herb. Trin.),’’ doubtless being the basis of thisname. We have not seen this specimen, but Trinius ® refers to this name in a note under P. potamium as ‘‘olim P. hydrophilum mihi dictam.”’ Setaria polygonata Kunth, Rév. Gram, 1: 47. 1829. Based on Panicum polygo- natum Schrad. Panicum trichogonum Willd.; Steud. Nom. Bot. ed. 2. 2: 261. 1841. This is given as a synonym under P. polygonatum Schrad. with the following citation: ‘‘ Willd. hrb. (Sec. Trin. mpt.).’’ The type, in the Willdenow Herbarium, is labeled ‘‘Amer. merid. Humboldt.”’ Panicum pilosum polygonatum Doell in Mart. Fl. Bras. 27: 211. 1877. Based on P. polygonatum Schrad. Panicum bourgaet Fourn. Mex. Pl. 2: 25. 1886. This name was earlier listed by Hemsley ¢ without description. Fournier cites a single specimen, ‘‘In valle Cordo- vensi, januario (BouRG[EAU] n. 1662 part.).’’ The specimen of this number seen at Halle is P. polygonatum, while the one in the Gray Herbarium is P. larum. The original description calls for a plant with pubescent nodes, which would indicate P. polygonatum. DESCRIPTION. Plants rather freely branching from the lower nodes, widely spreading from a decum- bent or creeping base, rooting at the nodes; culms 20 to 100 cm. long, the nodes densely pubescent; sheaths shorter than the internodes or sometimes nearly equal, densely ciliate, otherwise glabrous or hirsute toward the summit; ligules less than 0.5 mm. long; blades ascending or spreading, oblong-lanceolate, 3 to 13 cm. long, 8 to 15 mm. wide, usually ciliate at the cordate base, otherwise glabrous or occasionally sparsely hirsute; panicles 7 to 20 cm. long, about half as wide, the lower branches solitary, distant, spreading, the upper sometimes in pairs, the numerous raceme-like branchlets secund from the lower side of the branches, the somewhat clustered, short-pedicled spikelets also secund on the branchlets, the rachises sparsely pilose with long, weak hairs, or a Syn. Pl. Glum. 1:71. 1854. b Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 266. 1834. ¢ Biol. Centr. Amer. Bot. 3: 486. 1885. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 113 sometimes glabrous; spikelets 1.4 to 1.5 mm. long, 0.5 mm. wide, not thickened, pointed; first glume about half the length of the spikelet, 3-nerved, acute; second glume and sterile lemma equal, 3 to 5-nerved, exceeding the fruit, the sterile palea only about half the length of its lemma; fruit 1 mm. long, 0.5 mm. wide, obovate. This species differs from P. larum and P. pilosum in the pointed spikelets which are not expanded by an enlarged sterile palea, and in the compound lower branches of the panicle. DISTRIBUTION. Swamps and moist soil along roads and in open woods, Mexico to Braziland Paraguay. Mexico: Cérdoba, Bourgeau 1662 in part. GuaTeMaLa: Dept. Alta Vera Paz, Collins & Goll 08, Cook & Griggs 530, Tuerck- heim 7797, 8795; Puerto Barrios, Kellerman 5114. Honpuras: San Pedro Suld, Thieme 781, 5578, 5587 in part. Costa Rica: Buenos Aires, Tonduz 4042; Turrialba, Tonduz 4092; Tufs, Tonduz 11396; Talamanca, Tonduz 8557; Giri Biolley 3107; Echeverria, Pittier & Tonduz 2479; San José, Pittier 1183. Panama: Hart 73. Cotomsia: Santa Marta, Smith 206, 2190. Trinipap: Fendler 946 (Kew Herb.). Brazit: Madeira, Rusby 199 in part. ParaGcuay: Morong 441. 59. Panicum pilosum Swartz. Panicum pilosum Swartz, Prodr. Veg. Ind. Occ. 22.1788. ‘‘ Jamaica.’ The type, in the Swartz Herbarium, labeled ‘‘Jamaica, Swartz. P. pilosum fl. ind. occ.,’’ consists of two branching plants. Panicum distichum Lam. Encycl. 4: 731. 1798. ‘‘Cette plante crédit 4 la Jamaique.”’ The type, in the Lamarck Herbarium, consists of a panicle only. No locality is given upon the label. Panicum pilisparsum Meyer, Prim. Fl. Esseq. 57.1818. ‘‘In graminosis humidis plantitionis Hamburg,’’ Essequibo [British Guiana]. The type isin the Géttingen Herbarium, but a portion of this is in the Trinius Herbarium and was examined there. Panicum trichophorum Schrad. in Schult. Mant. 2: 247.1824. “In Brasilia. Princeps Sereniss. Maximil. Neowidensis.’’ The type is in the Schrader Herbarium at the Botanical Garden in St. Petersburg. Setaria disticha Humb.; ; Spreng. Syst. Veg. 1: 305. 1825. Based on ‘‘ Panicum distichon Lam.’’ Panicum distans Willd.; Spreng. Syst. Veg. 1: 305, 1825. This herbarium name of Willdenow is given as a synonym under Setaria disticha Humb. The type, in the Willdenow Herbarium, was collected by Humboldt in ‘‘America meridionale.”’ Panicum densiflorum Willd.; Spreng. Syst. Veg. 1:320.1825. The name is credited to ‘‘W. herb.,’’ and the locality is given as ‘‘ Ad Orinoc.’’ The type dollected by Humboldt, in the Willdenow Herbarium, is labeled ‘‘ Amer. Merid.’’ Setaria pilosa Kunth, Rév. Gram. 1: 47.1829. Based on Panicum pilosum Swartz. Setaria meyeri Kunth, Réy. Gram. 1:47. 1829. Based on Panicum pilisparsum Meyer. Setaria schraderi Kunth, Rév. Gram. 1: 47. 1829. Based on Panicum trichophorum Schrad. Panicum apiculatum Salzm.; Steud. Syn. Pl. Glum. 1: 65. 1854. This is given asasynonym under P. pilosum Swartz. In the Trinius Herbarium and in the United States National Herbarium are specimens bearing this name collected by Salzmann in Bahia, Brazil, which belong to P. pilosum. 41616°—vo1t 15—10——8 114 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum distichum lancifolium Griseb. Fl. Brit. W. Ind. 548. 1864. Grisebach bases this upon a specimen from Trinidad collected by Crueger. The type, Crueger 84, in the Grisebach Herbarium, is an exceptionally robust plant, with numerous racemes and villous nodes. Grisebach states that the rachis of the branches is glabrous, but the type has slender bristles thinly interspersed. It resembles Hart’s no. 732 from Jamaica. DESCRIPTION. Plants annual, usually decumbent or creeping at base, rooting at the nodes and send- ing up erect branches, smaller plants sometimes erect; culms usually branching, 25 to 100 cm. high, the nodes villous or sometimes glabrous or nearly so; sheaths elongated, but usually less so than the internodes, keeled, separating more or less from the culm, exposing the long prophyllum, and inrolled at the summit, somewhat simulating a petiole to the blade, glabrous or sometimes ciliate or sparsely hirsute; ligules wanting; blades ascending or spreading, narrowly fanecolate. 4 to 20 cm. long, 7 to 15 mm. wide, broadest near the cordate or truncate base, puberulent at the very base, otherwise glabrous or sometimes sparsely pubescent; panicles consisting of 10 to 40 epike- like, densely-flowered, somewhat spreading racemes _ alonganaxis 5 to 15 cm. long, the lower distant; racemes 1 to 3 cm. ck gages Z ie long, straight or curved, bearing numerous spikelets in clusters of paoermiedt eatharhs aie secund on the lower side, the rachises copiously to sparsely papillose-hispid, the hairs 1 to 3 mm. long, wanting in occasional specimens; spikelets about 1.5 mm. long, 0.6 mm. wide, and nearly 1 mm. thick; first glume about half the length of the spikelet; second glume and sterile lemma equal, the former 5, the latter 3-nerved, both scabrous on the midnerve at the apex, the sterile palea as long as its lemma, becoming subrigid and forcing open the spikelet; fruit 1.3 mm. long, 0.6 mm. wide. This species differs from P.laxum in the more freely branching habit, compara- tively shorter and uniformly cordate or truncate blades, and smaller panicles of shorter, denser racemes, usually conspicuously bristly. In the following specimens the bristles are wanting: Curtiss 305, Rovirosa 599, Rusby & Squires 79. The latter is also exceptional in having pubescent spikelets. In this species an occasional internode is much shortened, thus giving a few leaves the appearance of being nearly in pairs. Most of the specimens from North America have villous nodes, but those from South America are commonly glabrous on the nodes. DISTRIBUTION. Fields and open woods, Mexico, West Indies, and south to Brazil. Mexico: Mirador, Liebmann 411; Chiapas, Nelson 3056; San Juan Bautista, Rovirosa 599. GuATEMALA: Los Andes, Kellerman 5119; Dept. Alta Vera Paz, Goll 11, Tuerck- heim 8797. Honpuras: San Pedro Sula, Thieme 5587 in part; Bonacco Island, Gaumer. Nicaraaua: Wright. Costa Rica: Rio Hondo, Cook & Doyle 499; Talamanca, Tonduz 9495; Puerto Viejo, Biolley 7463. Panama: Fendler 368. Cusa: Herradura, Tracy 9063, Van Hermann 763; Dayaniguas, Wright 3451 in part; Sancti Spiritus, Ledn 908; Isle of Pines, Curtiss 305, Taylor 36. Jamaica: Gordon Town, Hart 732; Navy Island, Millspaugh 1859 ee Herb.), Port Merant, Hitchcock in 1890 (Hitchcock Herb.). Lrewarp IsLtanps: Guadeloupe, Duss 4154. WINDWARD IsLANDSs: Granada, Broadway in 1904. CotomBiA: Santa Marta, Smith 203. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 115 VENEZUELA: Sacupana, Rusby & Squires 79, 347. Trinipap: Hart 3293. Topaco: Eggers 5534. British Guiana: Jenman 5969. Dutcu Gurana: Hostmann 641 (Gray Herb.). Frencu Guiana: Rotherey 14. Brazi: Falls of Madeira, Rusby 210 in part; Campinas, Novaes 1288; Bahia, Salzmann; Rio Janeiro, Widgren, Wilkes Expl. Exped. in part; without locality, Riedel; Piauhy, Gardner. Bourvia: Mapiri, Rusby 212. Paraauay: Hassler 8192 (Gray Herb.). 60. Panicum laxum Swartz. Panicum larum Swartz, Prod. Veg. Ind. Occ. 23.1788. ‘‘ Jamaica.’’ The type, in the Swartz Herbarium, is the upper part of aculm. The spikelets are 2.2 to2.3 mm. long. A specimen in the Munich Herbarium sent by Swartz as P. lazum is a species of Leptochloa. Panicum agrostidiforme Lam. Tabl. Encycl. 1: 172. 1791. Lamarck cites ‘‘Ex Amer. merid. Communic. A. D. Richard.’’ The type, in the Lamarck Herbarium, is the upper part of a culm. Panicum tenuiculmum{e] Meyer, Prim. Fl. Esseq. 58. 1818. Meyer gives no par- ticular locality. The type is in the Géttingen Herbarium, but the portion examined is in the Trinius Herbarium, having been sent by Meyer to Trinius. Panicum diandrum Kunth, Rév. Gram. 2: 393. pl. 110. 1829. ‘‘Crescit in insula Guadelupz inque Brasilia.’’ The type, in the Berlin Herbarium, being the plant figured, was collected in Guadeloupe by Balbis in 1844. ‘ Panicum leptomerum Presl, Rel. Haenk. 1: 311. 1830. The locality was unknown to Presl, who gives ‘‘Hab......... ’? The type, collected by Haenke, in the Bohemian Museum, bears the name but no locality. ‘ Panicum ramuliflorum Hochst.; Steud. Syn. Pl. Glum. 1: 65.1854. This is based on ‘‘Pl. Kappler surin. nr. 1523.’’ Specimens of this number have been examined at Munich and Leipzig, but none was found in the Steudel Herbarium. Agrostis nigrescens Salzm.; Steud. Syn. Pl. Glum. 1: 65. 1854. ‘‘Bahia.’’ This name is mentioned under Panicum ramuliflorum Hochst. as applying to a variety “ramulis paniculae densiflorus.’’ Steudel gives the name earlier,? but as a nomen nudum. Salzmann specimens from Bahia, bearing this name, in the Trinius Herba- rium and in the United States National Herbarium, are densely flowered forms of P. larum. . Panicum nigrescens Salzm.; Steud. Syn. Pl. Glum. 1: 66. 1854. This is given asa synonym under P. /arum Swartz. Doell® also gives this name in the same way. It is doubtless the same as Agrostis nigrescens, but we have seen no Salzmann specimen labeled Panicum nigrescens. Panicum agrostis Nees; Doell in Mart. Fl. Bras. 2?: 213. 1877. This is given asa synonym under P. larwm Swartz, and is credited to ‘‘ Nees in herb. schedulis.’’ The type, in the Berlin Herbarium, was collected in Brazil by Sello. Panicum larum pubescens Doell in Mart. Fl. Bras. 2?: 213.1877. The first specimen cited by Doell is ‘Prope Santarem (Spruce ‘Panicum n. 26’).’? The type, in the Munich Herbarium, has pubescent spikelets. Panicum pilosum epilosum Fourn. Mex. P|. 2: 24.1886. This appears to be founded on P. agrostidiforme Lam., as this is the first synonym given. The specimens cited - by Fournier belong to P. larum Swartz. aNom. Bot. ed. 2. 1: 41. 1841. b Mart. Fl. Bras. 2?: 213. 1877. 116 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Plants more or less spreading, often rooting at the nodes of the decumbent base; culms simple or sparingly branching, 40 to 100 cm. or more high; sheaths shorter than the elongated internodes, ciliate and hirsute at the juncture with the blade, otherwise glabrous or papillose-hirsute toward the summit; ligules fimbriate, about 0.5 mm. long; blades erect or ascending, conduplicate or flat, 10 to 25 cm. long, 5 to 15 mm. wide, rarely wider, narrowed to the rounded or subcordate base, glabrous or with a few scattered hairs on the upper surface; panicles oblong in outline, 5 to 30 cm. long, composed of many slender, raceme-like branches, the lower distant, spreading, sometimes as much as 10 cm. long, the upper ascending; branchlets very short, mostly secund on the lower'side of the branches, bearing 2 or 3 spikelets, or a few toward the base of the lower branches 5 to 10 mm. long; spikelets 1.1 to 1.5 mm. long, about 0.7 mm. wide and as thick or thicker; first glume one-third to half the length of the spikelet, sub- acute, 1 to 3-nerved; second glume slightly shorter than the sterile lemma, the latter subtending a palea of nearly equal length, this becoming subrigid at maturity forcing open the spikelet; fruit 1 to 1.1 mm. long, about 0.5 mm. wide. This widely distributed species is variable in appearance. The following excep- tionally robust specimens from Mexico and Guatemala have comparatively short, cordate blades 1.5 to 1.8 em. wide, and very turgid spikelets 1.5 mm. long, a few stiff hairs on the branchlets: Finck 3, Liebmann 419, Purpus 2159, 2160,¢ Tuwerckheim 1254. Tonduz’s no. 3071 and his no. 4868, in the Costa Rica Herbarium, have pubescent spikelets as in the Spruce specimen upon which Doell bases his variety pubescens. These appear to be merély exceptional specimens. Fig. 109.—P. lax- um. Fromtype specimen. DISTRIBUTION. Savannas and open woods, Mexico, West Indies, and south to Paraguay. Mexico: Culiac4n, Palmer 1558 in 1891; Mirador, Liebmann 412, 419; Cérdoba, Finck 3; Mayito, Rovirosa 427; Zacualpan, Purpus 2159, 2160 in part; Oriz- aba, Botteri 688. GuATEMALA: Dept. Alta Vera Paz, Tuerckheim 1254, 8803; Santa Rosa, Heyde & Lux 3900; Puerto Barrios, Pittier 361. Honpuras: San Pedro Sula, Thieme 5587 in part. Costa Rica: Buenos Aires, Tonduz 4864, 4871; Rio Tilirf, Tonduz 3071; Turri- alba, Pittier 16123; Las Mesas, Pittier 3117. Banamas: Turk Island, Madiana (Gray Herb.). Cusa: Herradura, Caldwell & Baker 7136, Hitchcock 177, Tracy 9054, 9062, 9072, 9099; Wajay, Earle & Wilson 343; Santiago de las Vegas, Hitchcock 178, Tracy 9114; Retiro, Wright 759; Guanabacoa, Leén 914; Sancti Spiritus, Ledn 907, 909; without locality, Wright 3751 in part, 3863 in part; Isle of Pines, Curtiss 464, Palmer & Riley 1069, Taylor 37. Porto Rico: Utuado, Britton & Cowell 394; Cayey, Heller & Heller 522; Catafio, Heller & Heller 1378; Ponce, Heller in 1902; Mayaguez, Sintenis 360; Fajardo, Sintenis 1254; Mount Morales, Britton & Marble 1068; without locality, Eggers 1329. Danisu West Inptes: St. Thomas, Hggers 165 (Gray Herb.). LEEWARD IsLANpDs: Guadeloupe, Duss 3179. Winpwarp Istanps: St. Lucia, U.S. Fish Com. in 1887; Barbadoes, Dash 450; Granada, Broadway in 1905. a Panicum viscidellum Scribn. was also distributed under this number. ~ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 117 CotomBra: Santa Marta, Smith 202, 204; Cérdoba, Pittier 521, 553; Cali, Pit- tier 665. TrinipapD: Hart 2177, 2289. Britis Guiana: Jenman 6008, 6009. Frencu Guiana: Sagot 692 (Gray Herb.). Brazit: Bahia, Salzmann; Madeira, Rusby 210; Prov. Goyaz, Gardner 3517; Campinas, Novaes 1246; Rio Janeiro, Wilkes Expl. Exped. 10; without locality, Gardner 1183, Riedel 52, Sello. ParaGuay: Hassler 8465, Morong 534, 537, 977, 1574. Urueuay: Arechavaleta 31 in 1888, 2 in 1893. Ecuapor: Balao, Eggers 14585. Bourvia: Mapiri, Rusby 228, 236; Unduavi, Rusby 22; Yungas, Bang 266, in part, 308a; Chaco, Fries 1537. 61. Panicum exiguiflorum Griseb. Panicum minutiflorum Rich. in Sagra, Hist. Cuba 11: 305. 1850, not Rasp. 1825. “Crescit in pratis et locis herbosis insulae Cubae.’’ The type, in the Paris Her- barium, is labeled ‘‘Insula Cuba. Legit Ramon de la Sagra.”’ Panicum laxum variegatum Griseb. Cat. Pl. Cub. 233. 1866. The only specimen cited is Wright 3450. The type, in the Grisebach Herbarium, is from western Cuba, 1863, and is numbered ‘‘909=3450.”’ Panicum exiguiflorum Griseb. Cat. Pl. Cub. 234. 1866. Based on “P. minutiflorum Rich. ex descr., non Hochst.”’ Panicum tricolor Hack. Oesterr. Bot. Zeitschr. 51: 370.1901. ‘‘In insulis Bahama; Fortune Island, inter frutices, Eggers nr. 3978.’’ The type is in Hackel’s herbarium. DESCRIPTION. Plants perennial, cespitose, sometimes sending out prostrate, stolon-like shoots; culms simple or sparingly branching from the upper nodes, erect from a decumbent or ascending base, slender, wiry, 15 to 50 cm. high, the lower nodes geniculate; sheaths much shorter than the elongated internodes, ciliate, otherwise glabrous; ligules membranaceous, delicate, fim- briate; blades appressed or ascending, 2 to 6 cm. long, 0.5 to 1.5 mm. wide, glabrous; panicles 3 to 10 cm. long, one-third to half as wide, the slender branches few, spreading or reflexed at maturity, Fic. 110.—P. erig- bearing short, divergent branchlets with clustered, short-pediceled uiflorum. From spikelets; spikelets 1.4 to 1.5 mm. long, about 0.5 mm. wide, and ‘er twice as thick; first glume about one-third the length of the spikelet, epatum Griseb. 3-nerved; second glume about two-thirds the length of the subequal sterile and fertile florets, the sterile palea very large and firm at maturity, much expanding the spikelet; fruit 1.3 mm. long, 0.5 mm. wide, acute. The immature spikelets are dorsally compressed, as characteristic in this genus, but as they mature the sterile palea becomes greatly enlarged, with broad firm wings, and forces the spikelet open. In this character and in habit this species is most closely related to P. hians Ell. DISTRIBUTION. Low savannas and moist sandy woods, Bahamas and Cuba. Banamas: Fortune Island, Eggers 3978 (Hackel Herb.), Hitchcock in 1890; Inagua, Nash & Taylor 1450; Long Cay, Brace 4158, 4164; Acklins Island, Brace 4380 (all in Field Mus. Herb.). Cusa: Chirigote, Wright 3450; Herradura, Hitchcock 179, Tracy 9075; Prov. of Santa Clara, Leén 902; Guanabacoa, Leén 911, 913; Isle of Pines, Taylor 35. 118 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 62. Panicum hians Eli. Panicum hians Ell. Bot. 8. C. & Ga. 1: 118. 1816. Elliott gives no particular locality, but he states that the species is ‘‘ Very abundant near Michaux’s old farm, 10 miles from Charieston.’’ The type, in the Elliott Herbarium, is labeled ‘‘ Panicum Hians mihi. Hab: in pinetis humidis,’’ but without particular locality. Panicum oblongiflorum Desv. Opusc. 89. 1831. ‘‘Habitat in Carolina * * * & Bosc.”’ The type is in the Desvaux Herbarium. Panicum jejunum Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 2': 103. 1836. “‘Touisiana.’’ The type specimen, in the Trinius Herbarium, is marked ‘‘ Louisiana. mis. Hooker. 1835.” Aira incompleta Bosc; Steud. Nom. Bot. ed. 2. 1: 45. 1840. This isa nomen nudum mentioned as a synonym of Panicum debile [no author cited], but there is no cross reference under Panicum debile. We have seen specimens of this collected by Bosc, in the Trinius Herbarium, in the Padua Herbarium, and in the Delessert Herbarium. All are Panicum hians. In the absence of any evidence as to which is the type specimen we take as the type the one in the Padua Herbarium, which appears to con- tain Bose’s own herbarium. The locality of this is given as ‘‘Caroline.”’ Steinchisma hians Raf.; Ind. Kew. 2: 982. 1895. Based on Panicum hians Ell. Nash @ segregated the genus Steinchisma and was followed by Hitchcock. Steinchisma was first mentioned by Rafinesque ¢ in a letter to De Candolle in which he pro- poses several new genera, this genus appearing as follows: ‘‘Stein- chisma= Panicum divaricatum, hians.’’ This probably refers to Panicum hians Ell. and to P. divaricatum Michx. which is given by Elliott as a queried synonym under P.hians. Panicum divaricatum Michx. is, however, Festuca obtusa Spreng.¢ Rafin- esque’s name is given by Steudelé as ‘‘Steinschisma Rafin. Panicum debile.” Panicum debile Ell. is P. verrucosum Muhl. Nash separated the genus Steinchisma on the character of the enlarged palea of the sterile floret. This character is shared by Panicum eaiquiflorum, P. cupreum, the South American P. decipiens Nees, and, in less pronounced form, by P. larwm and P pilosum, while P. polygonatum, which is evidently allied to P. lJarwm, has a very small palea. This character, since it proves not to be correlated with any other, does not seem to us suflicient for the segregation as a genus of those species showing it, especially since such segregation would place closely allied species in separate genera. Bealf and Scribnerg misapply the name Panicum melicarium Michx. to P. hians Ell. Panicum melicarium Michx. is Panicularia elongata (Torr.) Kuntze, P. melicaria (Michx.) Hitche.’ Fra. 111.—P. hians. From typespecimen. DESCRIPTION. . Plants perennial, cespitose; culms simple or sparingly branching, 20 to 60 cm. high, erect or a few of the outer ones geniculate and rooting at the lower nodes, some- times prostrate and sending up erect branches; sheaths usually much shorter than the aSmall, Fl. Southeast. U. 8S. 105. 1908. b A. Gray, Man. ed. 7. 117. 1908. ¢ Bull. Bot. Seringe 220. 1830. dSee Hitchcock, Contr. Nat. Herb. 12: 149. 1908. ¢Nom. Bot. ed. 2. 2: 635. 1841. f Grasses N. Amer. 2: 127. 1896. g9U.S. Dept. Agr. Div. Agrost. Bull. 7: 66. /. 48. 1897. hSee Hitchcock, Contr. Nat. Herb. 12: 149. 1908. i Oi _ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 119 to 5 mm. wide, usually scarcely wider than the sheaths, erect, flat or folded, pilose on the upper surface near the base, otherwise glabrous; panicles 5 to 20 cm. long, usually loose and open, the primary branches few, slender, distant, spreading or drooping, sometimes rather narrow and compact, with ascending branches, the branchlets borne along the upper half or toward the ends only, the short-pediceled spikelets in more or less secund clusters; spikelets 2.2 to 2.4 mm. long, about 0.8 mm. wide and twice as thick at maturity, rather strongly nerved; first glume about half the length of the spikelet, acute; second glume and sterile lemma subequal, slightly exceeded by the enlarged, indurated sterile palea; fruit 1.8 to 1.9 mm. long, 0.7 mm. wide, the margins of the lemma scarcely inrolled. The immature spikelets are dorsally compressed; it is only as they mature that the characteristic induration of the sterile palea takes place. DISTRIBUTION. Damp soil along ponds and streams, North Carolina to Florida, west to New Mexico and north through Arkansas to southern Missouri and Oklahoma. Missourt: Campbell, Bush 120. NortH Carourmna: Rowan County, Small & Heller 205; Wilsons Mills, Chase 3098. Sours Carona: Columbia, Curtiss 3594; Orangeburg, Hitchcock 451. Georeia: Savannah, Kearney 187; Bulloch County, Harper 838; Oliver, Curtiss 6836. Fioriwa: Jacksonville, Combs 26, Kearney 173; Lake City, Combs 117, 125, 172, 187; Madison, Combs 262; River Junction, Combs 434; Pensacola, Combs 526; Quincy, Combs 427; Chipley, Combs 587; Waldo, Combs 712; Ellzey, Combs 831; Arcadia, Combs 1246; Eustis, Nash 213; Jupiter, Curtiss 5534; Manatee, Rugel 231; Braidentown, Combs 1261, 1325, Tracy 7038; Pease Creek, Curtiss 3594; Myers, Hitchcock 872, Lee Co. Pl. 488, 489; Miami, Chase 3889. . , Atapama: Tuskegee, Carver 47; : Mobile, Kearney 30, Mohr in 1883. Mississippi: Nicholson, Kearney 363; Starkville, Tracy in Pol- lard Distr. 1418; Waynesboro, Kearney 121; Ocean Springs, Seymour 25. ARKANSAS: Central Arkansas, Har- vey; Malvern, Eggert 117. Louisiana: Rayville, Ball 26; Cal- Fig. 112.—Distribution of P. hians. houn, Ball 60, Hitchcock 1282; Alexandria, Ball 535; Oberlin, Ball 190; Red River Parish, Ball 123; Abbe- ville, Langlois in 1884; Lake Charles, Hitchcock 1133, Tracy in 1897. Texas: Jacksonville, Plank 63; Luling, Plank 18; Texarkana, Heller & Heller 4103, 4145; Huntsville, Plank 65; Orange, Plank 46; Pierce, Tracy 7745; Terrell, Warburton in 1904; Hempstead, Hall 815; Indianola, Ravenel in 1869; Ennis, Smith in 1898; Harvester, Hitchcock 1184, Thurow in 1898; Galveston, Plank 35, 83; Kingsville, Piper in 1906; Cold Creek, Bigelow in 1853; without locality, Reverchon 99 in 1879. OxtaHoMA: Sapulpa, Bush 707, 708, 709; on the Washita, Palmer 381 in 1868. New Mexico: Las Cruces, Plank 6. 120 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 63. Panicum cupreum nom. nov. Panicum hians purpurascens Scribn. Proc. Acad. Phila. 1891: 296. 1891, not P. purpurascens H. B. K. 1815. ‘‘(3449) [Pringle] * * * Wet hollows in prairies of Flor de Maria, State of Mexico. September 4.’’ The type is in Hitchcock’s herba- rium. It was collected in 1890. DESCRIPTION. Plants perennial, in small tufts; culms simple, erect, 40 to 60 cm. high, slender, wiry, glabrous; leaves somewhat clustered at the base, the sheaths keeled, glabrous, the lower overlapping, the upper shorter than the inter- nodes; ligules fimbriate, scarcely 0.5 mm. long; blades 5 to 15 cm. long, 2 to 4 mm. wide, at the base scarcely as wide as the sheaths, erect or ascending, folded and more or less twisted, glabrous or with a few long hairs on the upper surface at the base; panicles very long-exserted, 3 to 11 cm. long, dark pur- ple, composed of a few distant, slender, appressed or ascending branches, naked about half their length, bearing short, crowded branchlets with densely clustered spikelets along the upper half or toward the ends; spikelets 3 mm. long, about 1.2 mm. wide, and at maturity nearly twice as thick, rather Fria. 113.—P. cupreum. strongly nerved; first glume one-third the length of the spike- From type specimen of Jo¢ or less, obtuse, concave along the midnerve; second glume P. hians purpurascens : . Scribn. and sterile lemma equal, exceeded by the enlarged sterile palea, the lemma strongly concave along the midnerve below; fruit 2.4 mm. long, 0.8 mm. wide, the margins of the lemma more inrolled than in P. hians, the apex tipped with a minute bit of hyaline membrane. This species is allied to P. hians, from which it is chiefly distinguished by the larger, more congested spikelets, with shorter, concave first glume and concave sterile lemma. The type collection, Pringle’s no. 3449, two specimens of which we have seen, is the only one known of this species. Stolonifera.—Plants perennial, decumbent at base or widely creeping and rooting at the nodes; culms branching, pubescent in lines or glabrate; sheaths (except in P. biglandulare) with a dense line of pubescence at the summit; ligules mem- branaceous, nearly obsolete; blades lanceolate or ovate-lanceolate, acuminate and with a pubescent, petiole-like base; panicles composed of few to several short, densely-flowered racemes along a main axis, a tuft of pubescence at the bases of the rachises; spikelets short-pediceled, mostly in 2’s, secund along the lower side of the rachis, strongly nerved, the second glume and sterile lemma scabrous on the midnerves toward the summit, exceeding the smooth and shining fruit. Spikelets hispid and with 2 crateriform glands on the sterile lemma; second glume and sterile lemma not boat-shaped. Spikelets Bot over 2mm. long; blades not over 4cm.long. 66. P. pulchellum. Spikelets 3.6 mm. long; blades 4 to 10 cm. long.......-.. 67. P. biglandulare. Spikelets glabrous, glandless; second glume and sterile lemma boat-shaped. Blades not over 5 cm., usually 2 or 3 cm. long; second elume rather blunt and shorter than the sterile Wes oat aoY pene 3 os ot Mima eget, ots VSR Ue See ee 64. P. stoloniferum. Blades 5 to 11 cm. long; second glume acute, nearly equal- ing the stemlelomms:. ...iancbeniwasuscswecmene 65. P. frondescens. Cl lis Ee tg bar oS) HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 121 64. Panicum stoloniferum Poir. Panicum stoloniferum Poir. in Lam. Encycl. Suppl. 4: 274. 1816. ‘‘Cette plante créit 4 l’ile de Cayenne.’’ The type is in the Cosson Herbarium. Panicum ctenodes Trin. Gram. Icon. 2: pl. 171. 1829. Trinius cites no specimen nor particular locality, but states that the figure is ‘‘ad specimen Brasiliense.’? There is apparently no specimen in the Trinius Herbarium bearing this name, but in the Berlin Herbarium is a specimen from Trinius which is probably a part of the type, and which bears the name in Trinius’s handwriting. It was collected in Brazil by Beyrich. Panicum leprieurii Steud. Syn. Pl. Glum. 1: 65. 1854. Two specimens are cited, *P. stoloniferum Hochst. Hrbr. Kappler nr. 1500. Surinam et Leprieur in Cayenne.”’ On account of the specific name Leprieur’s specimen is taken as the type. This is in the Steudel Herbarium at Paris. It is a some- what undersized specimen labeled, ‘“‘Cayenne, Leprieur 1835.”’ DESCRIPTION. Plants creeping, freely branching; culms 10 to 50 cm. long, with two opposite lines of minute crisp pubescence, rarely pubescent all over or glabrate, the nodes pubescent or glabrous; sheaths shorter than the internodes or overlapping on the branches, ciliate, other- Fie. 114.—P.stoloni- wise glabrate, or crisp-pubescent; ligules fimbriate, about 0.2 mm. ferum. From jong: blades 1 to 5cm. long, 3 to 15 mm. wide, the margins undu- type specimen. late, glabrous or sparsely pilose on the upper surface, minutely soft- pubescent beneath; panicles 1 to5cm. long, about one-third as wide, racemes 5 to 10 mm. long, Sebending or spreading, the main axis more or less pubescent; spikelets 2.3 to 2.5 mm. long, about 0.6 mm. wide and 1 mm. thick, glabrous; first glume about one-third the length of the spikelet, 3-nerved, blunt; second glume blunt, shorter than the acute sterile lemma, both somewhat boat-shaped, 5-nerved, the sterile palea about half as long as its lemma; fruit 1.3 mm. long, 0.5 mm. wide, acute. ° DISTRIBUTION. Woods and low grounds, Guatemala to Brazil and Ecuador. GuATEMALA: Puerto Barrios, Pittier 364. Trintpap: Broadway 2370; Crueger 79, Bot. Gard. Herb. 2293. British Guiana: Jenman 4081. Durex Guiana: “Surinam” Hering (Acad. Phil. Herb.). Frencw Guiana: Sagot 667. Braziz: Beyrich (Trinius Herb.). Ecuapor: Balao, Eggers 14149. 65. Panicum frondescens Meyer. Panicum frondescens Meyer, Prim. Fl. Esseq. 56. 1818. ‘‘In graminosis umbrosis insulae Arouabisch”’ [British Guiana]. We have not examined the type, which is at Géttingen, but we have seen in the Trinius Herbarium a portion of it which was sent by Meyer. Panicum olyraefolium Raddi, Agrost. Bras. 43. pl. 1./.6. 1823. Raddi states@ that this occurs ‘‘in sepibus prope fossas udas in viciniis Rio-Janeiro.’’ We have not seen the type of this, but the description and figure agree with P. frondescens. a Op. cit. 44 Sindee P. ‘done Yolium. 122 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum ctenodes majus Trin. Gram. Icon. 2: pl. 171. f. A. 1829. In the descrip- tion of P. ctenodes a larger variety is referred to and figure A of the plate represents this form. It does not appear certain that Trinius intended to name this in this way, but in the explanation of the figure ‘‘ad specimen Brasiliense’’ he uses the term “‘ Var. major,’’ and Kunth@ cites the name as ‘‘ Panicum ctenodes var. major.”’ The type is in the Trinius Herbarium. It is a shoot without a label, lying on a sheet of speci- mens of P. stoloniferum from Brazil and appears to be the basis of figure A. One of the loose labels may belong to this specimen. Panicum stoloniferum major[us] Kunth, Rév. Gram. 2: 389. 1831. Based on Pan- icum ctenodes majus Trin. Panicum brachyclados Reichenb.; Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 251. 1834. This is given as a synonym under P. frondescens Meyer, the authorship being given as ‘‘ Rchbch. in Weig. hb. Surin.’’ The type, in the Trinius Herbarium, is labeled ‘‘ Panicum (Echinochloa) brachyclados. Surinam. ex herb. Reichenb.’’ Panicum kegelii Steud. Syn. Pl. Glum. 1: 65.1854. ‘‘Guatemala.’’ There is a specimen in the Berlin Herbarium bearing this name, collected in Guatemala by Kegel (no. 12716), which may be the type. No specimen of this was seen in the Steudel Herbarium. Panicum umbrosum Salzm.; Steud. Syn. Pl. Glum. 1: 65, 1854, not Retz.1786. This is given as a synonym under P. stoloniferum Poir. The type, in the Paris Herbarium, is from Bahia, Brazil. DESCRIPTION. Culms ascending from a decumbent or creeping base, less freely branching than in P. stoloniferum, 30 to 50 cm. high, compressed, glabrous except below the panicle or sometimes with two lines of pubescence toward the summit of the internodes; nodes black; sheaths shorter than the internodes, densely ciliate, otherwise glab- rous; ligules nearly obsolete; blades 5 to 11 em. long, 12 to 20mm. wide, acuminate, narrowed toward the base, glabrous; panicles 5 to 11 em. long, usually less than one-third as wide, rather compact, the numerous, approximate racemes ascending or somewhat spreading, the lower 10 to 25 mm. long; spikelets 2.6 to 2.8 mm. long, about 0.6 mm. wide and 1 mm. thick, glabrous; first glume scarcely one-third the length of the spikelet, acute, scabrous on the midnerve; second glume and sterile lemma subequal, somewhat boat-shaped, acute, the sterile palea about two-thirds as long as its lemma; fruit 1.6 mm. long, 0.5 mm. wide, pointed, short-stipitate. This species resembles P. stoloniferum, but is larger in all its parts, while the numer- ous racemes are usually aggregated into a rather compact panicle. The spikelets of this species and of P. stoloniferwm, with their somewhat boat-shaped second glume and sterile lemma, suggest species of Sacciolepis. In this species the stipitate fruit also shows an approach to that genus, but the habit is wholly different. Fig. 115.—P. fronde- scens. From type specimen. DISTRIBUTION. Moist woods, Mexico to Brazil and Peru. Mexico: Papantla, Liebmann 405; St. Sebastian, Rovirosa 497. GuATEMALA: Dept. Peten, Walker 1138. Trinipap: Broadway 2371. British Guiana: Meyer (Trinius Herb.). Dutcn Guiana: Weigelt (Trinius Herb.); Hering (Acad. Phil. Herb.). FrencH Guana: Sagot 689 (Gray Herb.). a Rév. Gram. 2; 389. 1831. thank 45 HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 123 Braziu: Bahia, Salzmann; Rio Janeiro, Wilkes Expl. Exped. 11; Sao Paulo, Léf- gren & Edwall 2803; Porto Alegre, Reineck & Czermak 241. Perv: Péppig (Trinius Herb.). Bourvia: Mapiri, Rusby 229. 66. Panicum pulchellum Raddi. Panicum pulchellum Raddi, Agrost. Bras. 42. 1823. “‘In sylvaticis prope Catumby, non procul ab Urbe Rio de janeiro.’ There is a specimen in the herbarium of the British Museum marked ‘‘Panicum pulchellum Rad. Rio janeiro. Raddi” which appears to be authentic, but probably is not the type. Eriochloa? pulchella Kunth, Rév. Gram. 1: 30. 1829. Based on Panicum pulchellum Raddi. Panicum leptostachyum Presl, Rel. Haenk. 1: 311. 1830. ‘‘ Hab. in Mexico.” The type, in the herbarium of the Bohemian Museum, is labeled ‘‘Mexico.”’ DESCRIPTION. Plants apparently perennial, decumbent or creeping at base; culms slender, rather freely branching, ascending, pubescent in lines or glabrate, the nodes bearded; sheaths short, softly pubescent or glabrate; ligules ciliate, about 0.3 mm. long; blades thin, spreading, 1.5 to 4 cm. long, 4 to 15 mm. wide, rather abruptly acuminate, un- symmetrically subcordate; panicles oblong, 2 to 12 cm. long, rarely over 2 cm. wide, the racemes 5 to 15 mm. long, all about the same length, ascending or finally spreading, distant or approximate toward the summit; spikelets 1.8 to Be. pulckclum. 2 mm. long, about 0.8 mm. wide, turgid but not thicker From Raddi’s specimen in than wide, hispid, the hairs longer toward the margin; British Museum. first glume one-third to half the length of the spikelet, acute, 3-nerved; second glume and sterile lemma sub- equal, acute, the latter bearing at either side of the midnerve a crateriform gland, the sterile palea about three-fourths as long as its lemma; fruit 1.3 mm. long, 0.6 mm. wide, rather blunt. DISTRIBUTION. Woods and savannas, Mexico to Brazil and Bolivia. Mexico: Cérdoba, Bourgeau 1455; Minatitlan, Smith 589 (Hitchcock Herb.). GuATEMALA: Dept. Alta Vera Paz, Cook & Griggs 284, Maron & Hay 3153,* Tuerckheim 7702, 8794; El Palmar, Kellerman 6246. Costa Rica: Boruca, Pittier 4459, Tonduz 4460 in part; Cafias Gordas, Pittier 7360; Buenos Aires, Tonduz 4881. Braziu: Rio Janeiro, Raddi (British Mus. Herb.). Botrvia: Guanai, Rusby 217. 67. Panicum biglandulare Scribn. & Smith. Panicum biglandulare Scribn. & Smith, U. 8. Dept. Agr. Div. Agrost. Bull. 4: 13. pl. 4. 1897. ‘“‘Near Pinabete, Chiapas, February 8, 1896, at an altitude of 6,500 to 8,000 feet; No.3781,’’ collected by E. W. Nelson. The type, in the National Herba- rium, is a culm 120 cm. long, bearing two ascending branches about the middle, the lower portion being naked. 7 124 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Culms ascending from a decumbent base, becoming spreading and much-branched, somewhat compressed, the line of pubescence sparse or wanting; sheaths densely papillose-ciliate, otherwise glabrous; blades 4 to 10 cm. long, 10 to 18 mm. wide, lanceolate, acu- minate, narrowed to the rounded base, more or less pilose on both surfaces; panicles 5 to 12 cm. long, the few, distant, racemes 0.8 to 2 cm. long, ascending or finally spreading; spikelets 3.6 mm. long, about 1 mm. wide, pointed; first glume scarcely one-third the length of the spikelet, his- pid along the midnerve and margin; second glume shorter than the sterile lemma, 7-nerved, hispid, the hairs longer toward the summit and margin, the sterile lemma 5-nerved, hispidulous and along Fig. 117.—P. biglandulare. Fromtype the margins hispid, bearing at either side of the specimen. midnerve a crateriform gland, these more promi- nent than in P. pulchellum, the sterile palea nearly as long as its lemma, hispidulous; fruit 2.4 mm. long, 0.7 mm. wide, elliptic, minutely stipitate. In the original description of P. biglandulare the margins of the sheaths are de- scribed as ‘‘clothed with glands bearing branching hairs.’’ The hairs are found to be simple and arising from papille. DISTRIBUTION. Among bushes, mountains of Mexico and Guatemala. Mexico: Near Pinabete, Nelson 3781. GUATEMALA: Coban, Twerckheim II 1956. Parviglumia.—Plants erect or ascending, usually from a decumbent base; culms slender; sheaths densely ciliate and with a dense ring of pubescence at the summit; ligules less than 0.5 mm. long; blades firm, lanceolate, constricted into a very short petiole-like base, and having a thin, white, cartilaginous margin; panicles light green, with few, compactly flowered branches; spike- lets not over 2 mm. long, obovate, obtuse, glabrous, the first glume usually about one-fifth the length of the spikelet; fruit, except in P. parviglume, with scattered, appressed, silky hairs. Besides the three here given two Brazilian species belong in this group: P. trichidiachne Doell@ and P. schiffneri Hack.,® and also P. conchatum Fourn.¢ described from a Mexican specimen.4@ Blades 12 to 16 cm. long, 2 to 3 cm. wide; fruit glabrous... . 70. P. parviglume. Blades not over 10 cm. long nor 1.8 cm. wide; fruit with scattered silky hairs. Blades scabrous on the upper surface, not faleate.....-.-. 68. P. virgultorum. Blades sparsely hispid on the upper surface, falcate...-..- 69. P. schmitzii. aIn Mart. Fl. Bras. 2?: 339. pl. 49. 1877. b Denkschr. Math.-Naturw. Akad. Wiss. Wien 79: 11. 1906. ¢Mex. Pl. 2: 25. 1886. dSee P. conchatum Fourn. page 329. a ip ag er al es HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 25 68. Panicum virgultorum Hack. Panicum virgultorum Hack. Oesterr. Bot. Zeitschr. 51: 369.1901. ‘‘Costarica: in virgultis ad La Verbena prope Alajuelito leg. Tonduz (Pittier et Dur. Pl. costar. exs. nr. 8829) et ad rivulos prope tres Rios (Pittier ibid. nr. 4326).’’ The type, Tonduz 8829, is in Hackel’s herbarium. Tonduz®@ listed this species under the name ‘‘ Panicum oblongum Hack.’ This was a herbarium name at first applied by Hackel to Tonduz’s no. 8829, as shown by specimens in Hackel’s herbarium and others distributed by Tonduz. DESCRIPTION. Plants perennial, ascending from a decumbent or sometimes a widely creeping base, branching and rooting at the lower, geniculate nodes; culms slender, wiry, 0.4 to 1 meter or more long, compressed, glabrous or pubes- cent below the nodes; nodes pubescent or glabrous; sheaths often as long as the internodes, usually short- ciliate, otherwise glabrous or the lowermost pubescent; ligules membranaceous; blades 5 to 10 cm. long, 5 to 10 mm. wide, narrowly lanceolate, acuminate, narrowed to the base, scabrous on the upper surface, smooth and glossy beneath; panicles mostly long-exserted, 2.5 to 7 em. long, half to two-thirds as wide, the 2 to 4 branches ascending, compactly flowered except at the base, or the lower sometimes naked one-third its length, the short branchlets and pedicels pubescent; spikelets 1.6 mm. long, 0.9 mm. wide; first glume less than one-fourth the length of the spikelet; second glume and sterile lemma equal, 5-nerved; fruit 1.5 mm. long, 0.8 mm. wide, oval, smooth and shining but with sparse, long, appressed, silky hairs. Fig. 118.—P. virgultorum. From type specimen. DISTRIBUTION. Hedgerows and cultivated fields, Guatemala and Costa Rica. GuaTEMALA: Dept. Huehuetenango, Seler 2708. Costa Rica: Alajuelita, Tonduz 8818, 8829. 69. Panicum schmitzii Hack. Panicum schmitzii Hack. Ann. Naturhist. Hofm. Wien 17: 254.1902. No specimen nor locality is cited. The author states that the specimen was from Mexico and was sent him by Dr. Zahibruckner for identification, having been communicated to the Hofmuseum by Schmitz, but that neither the name of the collector nor the date of collection was given. The type is in Hackel’s herbarium. DESCRIPTION. Plants apparently perennial ascending, or erect from a long-jointed, creeping base, rooting and branching from Fic. 119.—P. schmitzii. From the nodes of the decumbent portion, the suberect branches type specimen. 20 to 45 cm. high, simple or nearly so; culms puberulent at least below the puberulent nodes; sheaths short, puberu- lent, at least toward the summit; ligules ciliate; blades 6 to 10 cm. long, 10 to 18 mm. wide, lanceolate, more or less falcate, narrowed to the rounded base, sparsely papillose-hispid on the upper surface, glabrous or with a few scattered papille or 126 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. branches finally somewhat spreading, pubescent at the base, bearing short, appressed, approximate branchlets of crowded spikelets along the upper two-thirds to three- fourths of their length; spikelets 1.8 mm. long, 0.9 mm. wide; first glume scarcely ° one-fifth the length of the spikelet; second glume slightly longer than the sterile lemma, both 5-nerved; fruit 1.5 mm. long, 0.8 mm. wide, oval, the lemma sparsely clothed with appressed, silky hairs, the palea glabrous. DISTRIBUTION. Shaded rocky slopes, southern Mexico. Mexico: Las Canoas, San Luis Potosi, Pringle 3817. 70. Panicum parviglume Hack. Panicum parviglume Hack. Oesterr. Bot. Zeitschr. 51: 429.1901. ‘“‘Costarica: in ripis fl. Rio Torres prope S. José (Pittier 9080), prope S. Francisco de Guadalupe (Tonduz, 2448).’’ The type, Pittier 9080, is in Hackel’s herbarium. DESCRIPTION. Culms slender, erect, from an ascending base, 100 cm. high, striate, sparsely papil- lose-pilose; sheaths exceeding the internodes, ciliate, sparsely papillose-pilose, striate, more densely pubescent at the juncture with the blade; ligules very short, ciliate; blades lanceolate or linear-lanceolate, 8 to 16 cm. long, 12 to 25 mm, wide, flat, firm, rounded at base, acuminate, sparsely hispidulous, especially above, the margin very sca- brous; panicles ovate, 15 to 25 cm. long, the branches spreading or ascending, the lower distant, solitary, 10 to 12 cm. long, the lower fourth naked, bearded at base; branchlets appressed, the longer 1 to2cm. long, Fig. 120.—P. parviglume. From ; ; i type specimen. bearing 3 to 8 subcontiguous spikelets on slender, flexuous, scabrous pedicels 0.5 to 2 mm. long; spike- lets oval, 2 mm. long, 1.2 mm. wide; first glume about one-fifth the length of the spikelet; second glume and sterile lemma equal, slightly exceeding the fruit, faintly 5-nerved, minutely apiculate; fruit smooth. DISTRIBUTION. Thickets along streams, southern Mexico and Costa Rica. The only complete speci- men in the National Herbarium is Tonduz’s no. 8448 from which the above descrip- tion is drawn. This was collected by Tonduz and Pittier at San Francisco de Guada- lupe, Costa Rica, ‘‘Buissons sur les bords du Rio Torres.”’ In the National Herbarium there is a panicle, with upper leaf, of a plant of this species, with the specimen of Botteri’s no. 150, collected in Mexico. A similar frag- mentary specimen is mixed with the same collection in the Gray Herbarium. Verrucosa.—Glabrous annuals with weak, divaricately branching culms, decumbent at base and usually provided with aerial brace-roots at the lower nodes, the lower internodes much shorter than the middle and upper; ligules ciliate, not over 0.5 mm. long; panicles with divaricate, capillary branches, spikelet- bearing toward the ends; spikelets tuberculate; fruit minutely papillose, the margin of the lemma flat, inrolled only at base. Spikelets about 2 mm. long, glabrous........................ 71. P. verrucosum. Spikelets over'S mimi, lone, Wispid le" Je s Po. How els oor testers 72. P. brachyanthum. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 127 ‘ 71. Panicum verrucosum Muhl. Panicum debile Ell. Bot. S. C. & Ga. 1: 129. 1816, not Desf. 1798. Nospecimen nor locality is cited. The type was not found in the Elliott Herbarium but the descrip- tion clearly identifies the species. Panicum verrucosum Muhl. Descr. Gram. 113.1817. ‘‘Habitat in N. Cesarea, Dela- ware, et Georgia.’’ The typeisin the Muhlenberg Herbarium. On the sheet is written “‘verrucosum” but there is nothing to indicate from which of the three States cited the specimen came. Panicum umbraculum Bosc; Spreng. Syst. Veg. 1: 314. 1825. This and the follow- ing are names found in the Willdenow Herbarium, on specimens collected by Bosc, and published as synonyms of P. verrucosum Muhl. Both belong to this species. Panicum rugosum Bosc; Spreng. loc. cit. DESCRIPTION. Plants bright green, solitary or few together, lax, at first erect but soon decumbent at base, and ascending or widely spreading; culms slender, 20 cm. to 1.5 meters high; sheaths shorter than the internodes, ciliate; blades thin, lax, flat, 5 to 20 cm. long, 4 to 10 mm. wide, somewhat narrowed toward the base, gradually nar- rowed to the acuminate apex, glabrous; panicles finally exserted, 5 to 30 cm. long, about as wide, diffuse, small panicles often produced at the lower nodes, at least the ultimate branchlets scabrous, the branches mostly solitary, the branchlets bearing a Fic. 121.—P. verrucosum. Fromtype {¢W short-pediceled spikelets, mostly in twos, toward specimen. the ends; spikelets 1.8 to 2.1 mm. long, about 1 mm. wide, elliptic-obovate, subacute; first glume one- fourth the length of the spikelet or less; second glume and sterile lemma warty, — glabrous, the glume shorter than the fruit at maturity; fruit 1.8 to.2 mm. long, 1 mm. wide, elliptic, acute. DISTRIBUTION. Wet, mostly shady soil, Atlantic Coastal Plain, Massachusetts to Florida and Texas; also in Indiana and Tennessee. Massacuusetts: Springfield, Andrews 23; Plymouth, Oakes. New York: Staten Island, Tyler in 1898. New Jersey: Atsion, Chase 3546; Egg Harbor, Scribner in 1886, Vasey in 1884; Englishtown, Pearce in 1884. PENNSYLVANIA: Tinicum, Diffen- baugh in 1868, Smith 99. Inprana: Dune Park, Chase 918, Hill 177 in 1898. DELAWARE: Ellendale, Commons 231. Fic. 122.,—Distribution of P. verrucosum. MaryLAnp: Eastern Shore, Canby. District or CotumpBia: Chase 5440, Dewey 408, Kearney in 1895; Steele in 1896, Ward in 1878 and 1879. ~Virertnta: Parksley, Warburton in 1903; Munden, Mackenzie 1671; Portsmouth, Noyes 84; Virginia Beach, Britton in 1895, Hitchcock 164, Kearney 2053; Suffolk, Boettcher 471; Dismal Swamp, Chase 3660. 128 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, NortH Carouina: West Raleigh, Stanton 1271; Swain County, Beardslee & Kofoid in 1891; Wilmington, Ashe in 1897. Souts Carona: Santee Canal, Ravenel (Gray Herb.). GeoraiA: Sumter County, Harper 638; Thomson, Bartlett 1103; Stone Moun- tain, Hitchcock 216; Augusta, Cuthbert in 1903; without locality, Latimer in 1885. Froripa: Jacksonville, Curtiss 3608, 4036, 5252, 5808; Milton, Chase 4314; Orange County, Baker 31, Combs 1049, 1116, Meislahn 22a; Titusville, Chase 3986; Eustis, Chase 4066, Nash 780; Gainesville, Chase 4202; Braidentown, Combs 1269, 1294; Bartow, Combs 1234; without locality, Rugel 598. TENNESSEE: Nashville, Gattinger in 1882. AtaBAMA: Cullman County, Eggert 59; Auburn, Earle & Baker in 1897. Mississippi: Nicholson, Kearney 379; Waynesboro, Kearney 120; Biloxi, Kearney 336 in part, Tracy 4562; Pass Christian, Langlois 35 in 1882.” Louisiana: Oberlin, Ball 204; Lake Charles, Chase 4424. Texas: Jefferson, Plank 30 (Hitchcock Herb.). 72. Panicum brachyanthum Steud. Panicum brachyanthum Steud. Syn. Pl. Glum. 1: 67. 1854. The only specimen cited is, ‘‘ Vinzent Coll. nr. 124. Texas.’’ The type, in the Paris Herbarium, is labeled ‘In sabulosis sylvaticis, Texas, prope coloniam Rusk County, Vincent coll. 124.” Panicum sparsiflorum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 36. 1889, not Doell, 1877. Vasey cites ‘‘(P. angustifolium, Chap. non Ell.),” gives a description, and follows with the range ‘‘South Carolina to Texas.’’ As this is not primarily a change of name, the type is one of the specimens which Vasey had before him when he wrote the description... From among those in the National Herbarium upon which Dr. Vasey has written the name, we have chosen as the type one collected in dry soil at San Bernardino, Texas, October, 1839, by Dr. Ridell, no. 20. This was first named by Dr. Vas2y, P. angustifolium Ell. The authority, ‘‘Ell.,’’ was changed to ‘‘Chap. non Ell.’’ No specimens from South Carolina can be found named P. sparsiflorum by Vasey nor is the species known from that State. DESCRIPTION. Plants weakly ascending or spreading from a decumbent base, freely branching from the lower nodes; culms slender, 30 cm. to 1 meter high; sheaths shorter than the internodes, minutely ciliate; blades 5 to 15 cm. long, 2 to 3 mm. wide, narrowed toward the base, often involute and scabrous toward the apex, the uppermost usually reduced; panicles finally ex- serted, 5 to 15 cm. long, about as wide, the branches few, scabrous, the lower == sometimes as much as 10cm. long, bear- , ing a few short-pediceled spikelets, mostly in twos, toward the ends; spike- lets 3.2 to 3.6 mm. long, 1.5 mm. wide, elliptic-obovate, abruptly pointed; first glume minute; second glume and sterile lemma subequal, the tubercles bearing stiff, spreading hairs; fruit 2.9 to 3 mm. long, 1.4 mm. wide, obovate-elliptic, subacute. Fiq@. 123.—P. brachyanthum. From type specimen. ® HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 129 DISTRIBUTION. Sandy soil, Louisiana, Texas, and Oklahoma. Loutstana: Oberlin, Ball 220; Lake Charles, Chase 4389; without locality, Hale (Hitch- cock Herb.). Texas: Jacksonville, Joor in 1884, Plank 22, 60; Grand Saline, Reverchon 2223; Paris, Heller 4221; College Station, Nealley in 1882; Galveston, Joor 3648; Industry, Wurzlow 9; Mar- shall, Bush 979; Tyler, Rever- chon 2224; Harvester, Thurow in 1898; without locality, Fia. 124.—Distribution of P. brachyanthum. Nealley in 1885. Ox.anHoma: Sapulpa, Bush 738 (Gray Herb.). Trichoidia.—Annuals, decumbent at base and rooting at the lower nodes, rather freely branching; blades oblong-lanceolate to ovate; panicles short-exserted or included at base until maturity, very diffuse, the numerous branches, branchlets and long pedicels capillary; spikelets minute, not over 1.4 mm. long; fruit minutely papillose, the margin of the lemma flat. Blades more than one-fourth as wide as long; spikelets i ee eto pt a.... 73. P. trichoides. Blades less than one-eighth as wide as long; spikelets mi- EE 0 SE i heli go Mite ramet 74. P. trichanthum. 73. Panicum trichoides Swartz. Panicum trichoides Swartz, Prodr. Veg. Ind. Occ. 24. 1788. ‘* Jamaica, Hispaniola.”’ Swartz cites Sloane’s plate 72, figure 3, but does not quote his diagnosis, hence the type @ is not the Sloane plant, which belongs to the same species, but the plant in the Swartz Herbarium labeled ‘‘trichoides fl. ind. occ.’’ from ‘‘Jamaica, Swartz.’ Panicum capillaceum Lam. Tabl. Encycl. 1: 173. 1791. Lamarck gives for the locality “‘Amer. merid.’’ and cites Sloane’s plate 72, figure 3. The type, in the Lamarck Herbarium, was collected in Porto Rico by Le Dru. Persoon,» while quoting Lamarck’s diagnosis, gives the name as ‘‘ Capillaceum (filamentosum).’’ The second word does not seem to be meant either as a change of name or as a variety. What the author intended is not apparent. Panicum acutifolium Willd.; Spreng. Syst. Veg. 1: 320. 1825. This is given asa synonym ynder P. capillare and is credited to ‘‘W. herb.’’ The type, in the Willde- now Herbarium, was collected by Humboldt at Cumanacoa. Panicum capillaceum strictius Doell in Mart. Fl. Bras. 27: 249. 1877. ‘‘In Prov. Piauhyensi (Gardner n. 3509).’’ This specimen we have not seen. This species has usually been referred by authors of American floras to P. brevifolium L., which is from India, and is the same as P. ovalifolium Poir. as described by Hooker, ¢ and a very different species. : @See Hitchcock, Contr. Nat. Herb. 12: 140. 1908. bSyn. Pl. 1: 83. 1805. cFl. Brit. Ind. 7: 44, 1896. 41616°—voLt 15—10-——-9 130 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Plants often widely spreading; culms ascending from a decumbent base, the ascend- ing portion 20 to 40 cm. high, rather slender, leafy, pubescent; sheaths short, but sometimes overlapping toward the summit and on the branches, ciliate and papillose- hirsute, at least tow ard the summit; ligules membranaceous-ciliate, scarcely 0.5 mm. long; blades preadine. thin, 2 to 6 cm. ae 1 to 2 cm. wide, ovate or ovate-lanceolate, somewhat unsymmetrical, acuminate, cordate and somewhat clasping at the ciliate base, glabrous or sparsely hirsute on both surfaces; panicles oval or ovate in out- line, 5 to 20 cm. long, nearly as wide, the axis sparsely pilose, the numerous branches and branchlets and the long pedicels Fic. 125.—P. trichoides. Capillary; spikelets 1.2 to 1.3 mm. long, about 0.5 mm. wide, Fromtype specimen. obovate-elliptic, sparsely hirsute; first glume about half the length of the spikelet, acute, l-nerved; second glume shorter than the sterile lemma, both 3-nerved, and at maturity sometimes ruptured by the expanding fruit, often only the lower part of each remaining; fruit at maturity 1.3 mm. long, 0.6 mm. wide, elliptic, minutely papillose while immature, becoming smooth but not polished. DISTRIBUTION. Woods and open ground, often a weed in cultivated soil, Mexico and the West Indies, south to Ecuador and Brazil. Mexico: Imala, Palmer 1758 in 1891; Manzanillo, Palmer 1083 in 1890; Acapulco, Palmer 287 in 1895; Guadalajara, Pringle 3828; Guatulco, Liebmann 317; Zacualpan, Purpus 2902; Cérdoba, Finck in 1893; La Correa, Langlassé 380; Coahuayutla, Emrick 53; Tabasco, Rovirosa 434; San Juan Bautista, Rovirosa 598; Yucatan, Gawmer 522; Tres Marias Islands, Nelson 4257. GUATEMALA: Santo Thomas, Deam 6041; Dept. Santa Rosa, Heyde & Lux 4299; El Palmar, Kellerman 6263; Alta Vera Paz, Tuerckheim 7801. Honpuras: Wilson 188 (Field Mus. Herb.). SaLvApDoR: San Salvador, Velasco in 1906. Costa Rica: Nicoya, Cooper 10379; Matina, Pittier 9754; Buenos Aires, Pittier 3651, Tonduz 4865; San José, Tonduz 3123; without locality, Pittier 4458, 16081. Cusa: Habana, Curtiss 714. Jamaica: Port Antonio, Fredholm 3282. Porto Rico: Mayaguez, Cowell 522, 583, Sintenis 160; Ponce, Heller 6094; Luquillo Mountains, Wilson 283; Martin Pefia, Heller & Heller 387; Cayey, Heller & Heller 531; Mount Morales, Britton & Cowell 444; Rio Piedras, Barrett 61; Santurce, Heller & Heller 157. Lrewarp Isuanps: St. Christopher, Britton & Cowell 295; Dominica, Eggers in 1881; Guadeloupe, Duss 2681. 2 Winpwarp IstaNnps: Martinique, Duss 1321, Hahn 1047; Granada, Broadway in 1905, Eggers 5987. CotomsiA: State of Magdelena, Pittier 1621; Cali, Pittier 971; Santa Marta, Smith 167. VENEZUELA: Tovar, Fendler 2499. Trinipap: ‘Ex herb. W. Mitten,’’ collector not given. British Guiana: Demerara, Jenman 4403. Braziu: Para, Spruce 466 (Panicum 13); without locality, Burchell 8706. Ecuapor: Balao, Eggers 14481; El Recreo, Eggers 15417. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 131 74. Panicum trichanthum Nees. ? Milium microspermum Lag. Gen. & Sp. Nov. 2. 1816. ‘‘Habitat in Nova His- pania. Semina communicavit perill. D. Sessé.’” We have not seen the type specimen and the short description applies equally well to P. trichoides. Fournier@ gives this as equivalent to P. trichanthum Nees. Panicum trichanthum Nees, Agrost. Bras. 210. 1829. ‘‘ Habitat in Brasilia aequa- toriali (Siber—Vidi in Herb. Reg. Berol.) In regno Mexicano (ab Humboldt, Haenke.) (Vide in Herb. Willd. et Haenk.)’”’ Nees’s first citation, however, following the diag- nosis and preceding his discussion of this and P. trichoides is as follows: ‘‘ Panicum tricho[iJdes, Humb. et K. * * * (fide Herb. Willd.)’’ While Nees’s description distinguishes carefully between this species and P. trichoides, there is some confusion as to the specimens mentioned, since some are referable to P. trichanthuwm and some to P. trichoides. The specimen in the Willdenow Herbarium marked “‘P. trichanthum” in Nees’s writing, which appears to be the specimen referred to as that of ‘‘Humb. et K.,’’ is accepted as the type. Panicum guayaquilense Steud. Syn. Pl. Glum. 1: 85. 1854. ‘‘ Jameson Hrbr. nr. 560, Guayaquil.’’ The type, in the Steudel Herbarium, is labeled ‘“‘Savannis Guaya- quilensibus.”’ Panicum microspermum Fourn.; Hemsley, Biol. Centr. Amer. Bot. 3: 492. 1885. “Panicum trichoides [Swartz, misapplied by] Ch. et Schl. in Linnea, VI. p. 33” and ‘‘ Panicum trichanthum Nees Agrostol. Bras. p. 210,’’ are cited as synonyms, no description being given. Schlechtendal and Chamisso® give without description P. trichoides Swartz as the name of Schiede & Deppe’s no. 894, the specimen of which, in the Berlin Herbarium, is referable to P. trichanthum. Panicum microspermum as used by Hemsley, then, must be considered as based on P. trichanthum Nees. Four- nier@ later describes the species under the name P. microspermum Fourn., citing as synonyms the names given by Hemsley and also ‘‘ Miliwm microspermum Lag.,’’ which, since Fournier takes up Lagasca’s specific name, would be the basis of Fournier’s name as published by himself. While the identity of M. microspermum is doubtful all the specimens cited by Fournier are referable to P. trichanthum. DESCRIPTION. Plants often in large bunches; culms ascending from a decumbent base, as much as 1 to 2 meters long, rather stout, glabrous; sheaths shorter than the internodes, ciliate, and usually with a villous ring at the juncture with the blade, otherwise glabrous or rarely pubescent toward the summit; ligules membranaceous, less than 0.5 mm. long; blades oblong- lanceolate, usually 10 to 15 em. long, 10 to 15 or rarely 25 mm. wide, cordate, rather strongly nerved, glabrous or puberulent, often pilose above the ligule; panicles 10 to 30 cm. long, the ; axis glabrous, the numerous branches and branchlets and the i Reames long pedicels capillary, flexuous; spikelets 1.2 to 1.4 mm. long, ate no. 1643. 0.6 mm. wide, obovate-fusiform, acute; first glume less than one- fifth the length of the spikelet; second glume shorter than the sterile lemma, both 3-nerved and under alens minutely bullate-rugose, often ruptured and breaking off as the fruit matures, as in P. trichoides; fruit 1.2 mm. long, 0.6 mm. wide. a Mex. Pl. 2: 22. 1886. b Linnaea 6: 33. 1831. 132 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Thickets, river banks and rocky slopes, Mexico, the West Indies, and south to Paraguay. ; Mexico: Colima, Palmer 1257 in 1891; Colipa, Liebmann. 432; Jicaltepec, Lieb- mann 320; Vera Cruz, Miiller 2172 in part; San Luis Potosi to Tampico, Pal- mer 1151 in 1879. GuatematLa: Alta Vera Paz, Lewton 377, Tuerckheim 7798; Gualan, Deam 424; Morales, Kellerman 6272. Honpuras: San Pedro Sula, Thieme 5587 in part. Costa Rica: Talamanca, Tonduz 8600, 8670. Panama: Bocas del Toro, Hart 87. Cusa: Trinidad, Wright 753; Romelie, Eggers 5350; Vento, Curtiss 598, Len 557. JamAIcA: Purdie (Gray Herb.). Porto Rico: Cayey, Sintenis 2471. CotomBiA: Santa Marta, Smith 2151. VENEZUELA: Tovar, Fendler 1648. Brazit: Without locality, Burchell 7062, 8791, Riedel 1360. Paraacuay: Morong 317, 1571. Urvilleana.—Perennials with large, densely villous spikelets, the fertile lemma clothed with long hairs on the margin. A South American group of two or three species, of which one extends into the desert region of the southwestern United States. 75. Panicum urvilleanum Kunth. Panicum megastachyum Presl, Rel. Haenk. 1: 305. 1830, not Nees 1826. ‘‘ Hab. in montanis Peruviae huanoccensibus.’’ The type specimen, labeled ‘‘Panicum mega- stachyum Presl. Peruana montano guanoccensis. Haenke,’’ is in the herbarium of the Bohemian Museum. Panicum urvilleanum Kunth, Rév. Gram. 2: 403. pl. 115. 1830. Kunth gives no definite locality other than ‘‘Crescit in regno Chilensi.’? The name is listed with the citation ‘‘Chili: Legit amiciss. Dumont D’Urville,’’ but without description, in an earlier part of the same work.¢ The type specimen, in the Berlin Herbarium, is labeled ‘‘Conception de Chili, D’Urville ded 1815.” Panicum preslet Kunth, Enum. Pl. 1: 121. 1833. Based. on ‘‘P. megastachyum Presl,’’? the name is presumably changed because of P. megastachyum Nees. Presl’s description is copied, Kunth evidently not having seen the plant. Panicum urvilleanum longiglumis[e| Scribn. U.S. Dept. Agr. Div. Agrost. Bull. 17: (ed. 2). 49.1901. ‘‘San Jacinto, Southern Calif. No. 887. 8S. B. & W. F. Parish, June, 1882.’’ The type specimen, in the National Herbarium, has spikelets about 7 mm. long. DESCRIPTION. Plants robust, erect from a creeping rootstock, 0.5 to 1 meter high, culms solitary or few in a tuft, simple or branching at the base only, the nodes densely bearded, rarely visible; sheaths overlapping, loose, densely, retrorsely, harshly villous; ligules densely ciliate, about 2 mm. long; blades 30 to 60 cm. long, 4 to 7 mm. wide, tapering from a flat base to a long involute-setaceous point, retrorsely strigose to nearly glabrous on both surfaces; panicles short-exserted, equaled or exceeded by the upper blades, 25 to 30cm. long, about half as wide, rather many-flowered, the glabrous to pilose, slender, flexuous branches ascending, producing spikelet-bearing branchlets along the upper half to two-thirds of their length; spikelets short-pediceled, 6 to 7 mm. long, about 2 @Op. cit. 1: 35. 1829. - HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 133 mm. wide, and as much as 4 mm. thick, ovate, densely silvery to tawny villous, strongly nerved but the nerves obscured by the pubescence; first glume clasping, two- thirds to nearly as long as the spikelet, sparsely villous or glabrescent toward the —— ~ SASS —S — WS SSS ss \ Fic. 127.—P. urvilleanum. From type specimen. acuminate apex; second glume slightly longer and more pointed than the sterile lemma, both exceeding the fruit, the lemma inclosing a villous palea of equal length and a staminate flower; fruit 4.2 to 4.5 mm. long, about 1.6 mm. wide, the margins of the lemma clothed with long white hairs, otherwise smooth and shining. DISTRIBUTION. Sandy deserts, Arizona and southern California south to Argentina. Arizona: Without locality, Lemmon in 1884. CaAuirorniA: San Jacinto, S. B. & W. F. Parish 887 in 1882; Agua Caliente, S. B. & W. F. Parish 887 in 1881; Barstow, Chase 5766, Tracy 434; Hesperia, Abrams 2164; Colorado Desert, Chase 5519, Parry & Lemmon 400 (Hitchcock Herb.), Wilder 1082. ARGENTINA: Rio Negro, Wilkes S. Pac. Expl. Exped. in 1838- 1842. Cute: Gillies (Gray Herb.). Fig. 128.—Distribution of P. urvilleanum. UNGROUPED SPECIES OF TRUE PANICUM., The following tropical species do not fall into any of the foregoing natural groups nor, with the exception of Panicum rudgei and P. rotundum, which are allied species, do they form such groups among themselves. The Mexican and Central American species are as yet not so well understood as those of the United States, and further study is necessary before our knowledge of the species of the former regions shall be as detailed as that of the species occurring north of the Mexican boundary. This study awaits additional material and extended field work in Mexico and Central America. 134 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 76. Panicum costaricense Hack. Panicum costaricense Hack. Oesterr. Bot. Zeitschr. 51: 428. 1901. ‘‘Costarica: Inter Buenos Aires et Térraba (Pittier 3636); in silva prope Térraba (3673), in vir- gultis ad Rio Ceibo (4860).’’ The first specimen cited, which is taken as the type, is in the Brussels Herbarium, and was examined by Hackel, but is not in his own herba- rium. The two other specimens are in Hackel’s herbarium. DESCRIPTION. Plants apparently annual; culms slender, erect, 0.5 to 1 meter high, sparingly branched, striate, glabrous or minutely pubescent; sheaths much shorter than the elongated internodes, papillose-pilose to glabrate, ciliate at least toward the summit; ligules membranaceous, ciliate, about 0.5 mm. long; blades 5 to 10 cm. long, 6 to 12 mm. wide, narrowed toward the base, long-acuminate, very sparsely pilose on both surfaces or glabrate; panicles ovoid in outline, 10 to 15 cm. long, about two-thirds as wide, loosely flowered, the slender, flexuous branches ascending or spreading, bear- ing delicate branchlets throughout, the scattered spikelets on capillary, flexuous pedicels; spikelets 2.4 mm. long, 0.7 Fi. 129.—P. costaricense. ™m. wide, elliptic, acute, rather strongly nerved; first glume From type specimen. nearly half the length of the spikelet, acute, hirsute toward the margin; second glume and sterile lemmasub qual, pointed beyond the eh hirsute toward the margin and with very minute papillz bordering the nerves; fruit 1.7 mm. long, 0.6 mm. wide, elliptic, smooth and shining, the margins of the lemma near ee base each bowed out into a little angle, giving a somewhat auricled appearance. The somewhat auricled base of the fruit in this species suggests an approach to species of Ichnanthus. DISTRIBUTION. Forests, Costa Rica. Costa Rica: Buenos Aires, Pittier 3661, Tonduz 4860; Cordoncillal, Pittier 3640; Boruca, Pittier 4626; Térraba, Tonduz 3673. Panicum expansum Fourn.,¢ the type specimen of which, Liebmann’s no. 426 from Huitamalco, Mexico, in the Copenhagen Herbarium, was examined at Halle, is appar- ently closely related to P. costaricense. The fruit has the same auricled or angled base, but the spikelets are slightly larger and glabrous, and the blades are glabrous. Lieb- mann’s no. 427 in the same herbarium is the only other specimen of this species we have seen. For the satisfactory placing of this species more material is needed. 77. Panicum parvifolium Lam. Panicum parvifolium Lam. Tabl. Encycl. 1: 173. 1791. The only specimen men- tioned is ‘‘Ex Amer. merid. Communic. D. Richard.’’ The type, in the Lamarck Herbarium, labeled, ‘‘ill. gen. ex D. Richard,’’ is a single slender culm. Panicum brasiliense Spreng. Syst. Veg. 1: 321. 1825. ‘‘Brasil. (P.ascendens W. herb.)’’ In the Willdenow Herbarium is a specimen of P. parvifoliwm labeled, ‘‘Panicum adscendens. Brasil. Hoffmansegg,’? which is evidently the specimen referred to by Sprengel, and is the type of P. brasiliense. Panicum ascendens Willd.; Spreng. Syst. Veg. 1: 321. 1825. This is given asa synonym of P. brasiliense of which it is a typonym. Panicum adscendens Hoffmgg.; Ind. Kew. 2: 410.1894. This is listed with the reference ‘‘ex Schult. Mant. 2: 592,’’ but this is Meh o an error, since the Mantissa a Hie Pl. 2: 26. 1886. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 135 volume 2 contains but 522 pages in all the copies we have seen. The type is undoubt- edly the Hoffmansegg specimen mentioned above. Panicum oplismenoides Nash, Bull. Torrey Club 30: 381. 1903, not Hack. 1888. “Collected on the edge of a ditch at Vega Baja, May 9, 1899, by Heller, no. 1316.” The type is in the herbarium of the New York Botanical Garden. DESCRIPTION. Plants perennial, cespitose, decumbent or creeping, rooting at the lower nodes, glaucous and glabrous throughout, except as noted; culms slender, branching, 20 to 80 cm. long, leafy, with numerous short internodes, the nodes sometimes sparsely pilose; sheaths rarely over 1 cm. long, ciliate, and some- times, especially on young shoots, sparsely pilose; ligules { \\__ nearly obsolete; blades 1 to 3 cm. long, 2 to 6 mm. wide, | oblong-lanceolate, rounded or subcordate at base, spreading j or reflexed, or the upper and often those of young shoots appressed, sometimes sparsely pilose at the base; panicles Fic. 130.—P. parvifolium. Short-exserted, 2 to 4 cm., rarely 6 cm. long, about as wide, From type specimen. loosely flowered, the slender, flexuous branches spreading, the branchlets and pedicels divergent; spikelets about 1.5 mm. long, 0.8 mm. wide, turgid, blunt, glabrous; first glume slightly more than half the length of the spikelet, subacute, 3-nerved; second glume and sterile lemma sub- equal, 5-nerved, the sterile palea nearly as long as its lemma; fruit 1.4 mm. long, 0.8 mm. wide, ovate, smooth and shining. DISTRIBUTION. Damp shady places, Costa Rica and, the West Indies, south to Brazil and Paraguay. Costa Rica: Buenos Aires, Pittier 10594, Tonduz 3631. Cusa: Los Almacigos, Wright 3458; Herradura, Baker 2078, Hitchcock 181, Tracy 9060, 9079. Porto Rico: Sintenis 5719, 1216 (Krug & Urban Herb.); Vega Baja, Heller & Heller 1316. Trinipav: Broadway 2372. British Guiana: Schomburgk 407. Dutcn Gutana: Surinam, no collector given (Gray Herb.). Frencu Guiana: No data (Gray Herb.). Braz: Falls of St. Gabriel, Spruce 2207; Santarem, Spruce 632; Sio Paulo, Léf- gren 1124; Campinas, Novaes 1245; without locality, Riedel 958. ParaGcuay: Morong 519. 78. Panicum millegrana Poir. Panicum hirsutum Lam. Encycl. 4: 741. 1798, not Swartz, 1797. ‘‘Cette plante croit 4 Cayenne, d’ou elle a ét4 envoyée par le citoyen Leblond.’”’ The type, in the Paris Herbarium, labeled ‘‘Cayenne, Le Blond,’’ has glabrous spikelets. Panicum millegrana Poir. in Lam. Encycl. Suppl. 4: 278. 1816. ‘‘Cette plante croit dans l’Amérique méridionale (V. s. in herb. Desfont).’”’ This specimen was not found in the Desfontaine Herbarium at Florence. In the Paris Herbarium is a specimen from ‘‘Cayenne, Martin,’’ labeled ‘‘Panicum millegrana Poir.,’’ which appears to be authentic and may be the type. Poiret’s description applies well to this plant and to the species known as P. rugulosum Trin. The spikelets are glabrous as in the type of that species. ; Panicum rugulosum Trin. Gram. Pan. 195. 1826. ‘Brasil (LANGsporrr.)’’ The type, in the Trinius Herbarium, has glabrous spikelets. 136 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum sellowii Nees, Agrost. Bras. 153. 1829. ‘‘ Habitat in Brasilia meridionali. (Sellow).’’ The type, in the Berlin Herbarium, has spikelets densely papillose- pubescent with short, stiff hairs. Panicum beyrichii Kunth, Rév. i dal P20, ipl, 27. 1830. ‘‘Crescit in nemoribus prope novum Triburgum Brasiliz.’’ Kunth states that he received the plant under the name P. sellowit Nees from Beyrich who collected it. The type, in the Berlin Herbarium, has glabrous spikelets as in the type of P. millegrana. Panicum lasianthum Trin. Gram. Icon. 3: pl. 245.1830. Trinius states that the figure is ‘‘ad specimen Brasilianum.’’ The type, in the Trinius Herbarium, collected in Brazil by Langsdorff, has papillose-hispid spikelets. Panicum puberulum Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 277. 1834, not Kunth, 1829. ‘Trinius states that his specimen is from Brazil. The type, in the Trinius Eierbannan) collected in Brazil by Sello, has papillose-hispid spikelets as in the type of P. sellowtt. Panicum dispersum Trin. Mém. Acad. St. Pétersb. VI. Sci. Nat. 1: 282. 1834. Trinius gives the locality as ‘‘Bahiens.’’ The type, in the Trinius Herbarium, col- lected in Bahia, Brazil, by Riedel in 1831, is a prostrate or decumbent plant with glabrous spikelets. Panicum expansum Trin.; Steid. Nom. Bot. ed. 2. 2: 256. 1841. This is a nomen nudum credited to ‘‘Trin. mpt. Mexico.’’ The type, in the Trinius Herbarium, labeled ‘‘Mexico, Hacienda J.aguna, Schiede,’’ has glabrous spikelets. This is not the P. expansum of Fourn‘sr@ who cites Trinius’s name as a synonym under P.rugulosum.® Panicum pilosum leiogonum Rupr. Bull. Acad. Roy. Belg. 9°: 239. 1842. This name is listed without description. The only specimen mentioned is Galeotti 5728 from Xalapa, Mexico. The type is in the Brussels Herbarium. This name is given by Fournier@ as a synonym under P. rugulosum, but Galeotti’s no. 5728 he cites¢ also under P. pilosum genuinum. Panicum selloww longevaginatum. Rupr. Bull. Acad. Roy. Belg. 9?: 239. 1842. Under this name, which is listed without description, Ruprecht cites two speci- mens, Galeotti 5726 and 5699. The first has glabrous, the second hispid spikelets. This name is given by Hemsley@ and Fournieré as a synonym under P. rugulosum. Panicum valenzuelanum Rich. in Sagra, Hist. Cuba 11: 304. 1850. ‘‘@rescit in locis montosis partis occidentalis insulae Cubae Vuelta de abajo dictae * * * (Don José Maria Valenzuela).’’ The type, in the Richard Herbarium, labeled ‘‘ Vuelta de Abajo (Valenzuelana),’’ has papillose-hispid spikelets. Panicum probandum Steud. Syn. Pl. Glum. 1: 76. 1854. Based on ‘‘P. puberulum Trin. non Kunth.”’ Panicum rugulosum hirtiglume Griseb. Cat. Pl. Cub. 233. 1866. The only speci- men cited is Wright 3455. The type, in the Grisebach Herbarium, has hiSpie wars spikelets. Panicum rugulosum glabrescens Doell in Mart. Fl. Bras. 27: 259. 1877. As no specimen is mentioned, P. beyrichii Kunth, the first synonym cited, is taken as deter- mining the type. Panicum rugulosum pubescens Doell in Mart. Fl. Bras. 2?: 259. 1877. No speci- men is mentioned. The type of the first synonym cited, P. rugulosum Trin., has glabrous spikelets, while Doell describes his variety as having puberulent glumes, hence the type of the seeond synonym cited, P. sellowii Nees, which has hispid spikelets, is taken as the type. a Mex. Pl. 2: 26. 1886. d@ Biol. Centr. Amer. Bot. 3: 495. 1885. b Op. eit. 21. é¢ Mex. Pl. 2: 21. 1886. cOp. eit. 24. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 137 Panicum rugulosum subvelutinum Doell in Mart. Fl. Bras. 27: 259. 1877. ‘‘A el. Wullschlaegel (n. 1612) in Surinamiae districtu Paraénsi lecta.’’ We have not seen this specimen but the description, ‘‘foliorum lamina utrinque subvelutina,’’ would indicate the form with velvety blades. DESCRIPTION. Plants perennial, spreading; culms sparingly branching, 0.5 to 1 meter high, ascending from a decumbent base, softly pubescent to glabrous; sheaths ciliate and with a dense ring of pubescence at the summit, otherwise papillose-pilose to glabrous; ligules membranaceous, scarcely 0.3 mm. long; blades ascending or spreading, thin, ovate-lanceolate, 4 to 15 cm. long, 10 to 30 mm. wide, somewhat unsymmetrical at the rounded or slightly cordate, sometimes ciliate, base, softly pubescent, or sometimes velvety, on both surfaces to glabrate except near the margin and at the base; panicles short-exserted, finally loose and rather few-flowered, 10 to 20 cm. long, about two-thirds as wide when expanded, the rather few, slender, branches stiffly ascending or spreading, bearing toward the ends short, ap- pressed branchlets with 1 to 3 rather short-pedi- celed spikelets; spikelets 2 to 2.3 mm. long, 1 to 1.2mm. wide, obovate, obtuse, turgid, at maturity olivaceous or brown, glabrous or more commonly papillose-hispidulous; first glume about two-thirds as long as the spikelet, acute; second glume slightly shorter than the sterile lemma, exposing the summit of the fruit at maturity, both 5-nerved, in glabrous spikelets the nerves bordered by interrupted rows of minute papille; fruit 1.9 to 2.1 mm. long, about 1 mm. wide, elliptic, obscurely pointed, papillose-rough- ened, becoming dark brown at maturity. This species as here defined is very variable. The examination of a greater num- ber of specimens and field study may show P. sellowii to be distinct from P. millegrana. From the material at hand they can not be satisfactorily separated, for while most of the specimens have either glabrous or papillose-hispidulous spikelets a few have both sorts in the same panicle, and the pubescence of the sheaths and blades can not be correlated with that of the spikelets. The following specimens have glabrous spikelets: Fendler 1641, Heyde & Lux 3927, Holway 3083, Liebmann Pl. Mex. 275, Léfgren 1228, Regnell III 1359**, Riedel, Rusby 233, Smith 2146, Widgren in 1844. In Spruce 603 and Tuerckheim 657 most of the spikelets are glabrous but some in the same panicle are hispidulous, while in Wright 3455 the greater number of the spikelets are hispidulous but glabrous ones are found in the same panicle. Fig. 131.—P. millegrana. From type specimen of P. rugulosum Trin. DISTRIBUTION. Damp woods, Mexico and Cuba, south to Brazil. Mexico: Mirador, Liebmann 275; Jalapa, Holway 3083; State of Chiapas, Heyde & Lux 3927. GuatTeMALA: Dept. Alta Vera Paz, Tuerckheim 657, 8783, 8784. Costa Rica: El General, Pittier 10615. Cusa: Habana, Wright 3462 in part; La Catalina, Wright 3455; Pinar del Rio, Wright 3855; Herradura, Hitchcock 180, Tracy 9098. Cotomsia: Santa Marta, Smith 2146. VENEZUELA: Tovar, Fendler 1641. Brazit: Campinas, Novaes 1249; Sio Paulo, Léfgren 1228; Rio Janeiro, Widgren in 1844; Prov. Minas Geraes, Regnell III 1359**; Santarem, Spruce 603; Madeira, Rusby 233; without locality, Burchell 4146, 4315-2, 4653, Riedel. Paracuay: Laguna Ipacarary, Fiebrig 561 (Field Mus. Herb.). 138 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 79. Panicum glutinosum Swartz. Panicum glutinosum Swartz, Prodr. Veg. Ind. Occ. 24. 1788. ‘‘ Jamaica.” The type is in the Swartz Herbarium. Panicum obtusiflorum Rich. in Sagra, Hist. Cuba 11: 305. 1850. ‘‘Crescit ad marginem rivuli Cauta in provincia Santiago de Cuba. (Linden,n.2143.)” The type, in the Richard Herbarium, is labeled ‘‘St. Yago de Cuba. Linden 2143.” Panicum lindenii Griseb. Cat. Pl. Cub. 233. 1866. Based on ‘‘P. obtusiflorum Rich. Cub. non Hochst. [1851]; Lind[en]. 2143.” DESCRIPTION. Plants perennial, somewhat glaucous; culms erect from a geniculate or decumbent base, often rooting at the lower nodes, 1 to 2 meters high, robust, compressed, glabrous; ° sheaths often longer than the internodes, somewhat keeled, especially the lower, glabrous or sometimes pilose, densely bearded at the juncture with the blade; ligules obso- lete; blades elongated-lanceolate, acuminate, 15 to 50 cm. long, 15 to 25 mm. wide (the up- permost reduced), abruptly or gradually nar- rowed at the base, more or less ciliate along the lower portion, glabrous or sometimes very sparsely pilose; panicles rhomboid in outline, 15 to 30 cm. long, about as wide, the lower branches verticillate, nearly as long as the main axis, stiffly ascending, the axis and Fig. 132.—P. glutinosum. From type branches glabrous, sometimes viscid, bearded specimen. in the axils, bearing slender, flexuous, sca- brous branchlets, with rather long-pediceled spikelets, mostly along the upper half; spikelets 3 mm. long, 1.5 to 2 mm. wide, turgid, obovoid, obtuse, olivaceous to brown, the faintly nerved, very viscid glumes whitish on the margin; first and second glumes about equal, slightly shorter than the fruit, the sterile lemma thinner in texture, mostly entirely concealed beneath the first glume, sterile palea wanting; fruit 2.6 mm. long, 1.2 to 1.5 mm. wide, obovoid- elliptic, olive-brown, densely minutely puberulent at the summit, otherwise smooth and shining, the margins of the lemma toward the summit scarcely inrolled. DISTRIBUTION. Mountain woods, Mexico and the West Indies, south to Paraguay and Bolivia. Mexico: Zacualpan, Purpus 2156, 2903; Mirador, Liebmann 428; San Cristobal, Bourgeau 3192; Orizaba, Botteri; State of Chiapas, Nelson 3357. Costa Rica: Cafias Gordas, Pittier 11017; Diquis Valley, Pittver 12002. Cuspa: Loma Pelada, Wright 757. Jamaica: Gordon Town, Hart 792; Troy, Maxon 2816. Porro Rico: Mayaguez, Sintenis 357; Sierra de Yabucos, Sintenis 2609. Brazit: Prov. Minas Geraes, Widgren in 1845, Regnell III 1370; Sio Paulo, Lofgren 2291, Lofgren & Edwall 2383; Campinas, Novaes 1247. Paracuay: Central Paraguay, Morong 405 A. Boutvia: Mapiri, Rusby 244. f HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 139 80. Panicum rudgei Roem. & Schult. Panicum scoparium Rudge, Pl. Guian. 1:21. pl. 29. 1805, not Lam. 1798. No par- ticular locality in Guiana is mentioned by Rudge. In the Berlin Herbarium is a specimen from Rudge which is authentic though it probably is not the type. The original description and the plate leave no doubt as to the identity of the species. Panicum rudgei Roem. & Schult. Syst. Veg. 2: 444. 1817. Based on ‘Pan. scoparium Rudge.” Panicum rudgei brasiliense Raddi, Agrost. Bras. 48. 1823. ‘‘Species rarissima observata tantum in viciniis fluminis Inhu- mirim.’’ We have not seen the type of this, but the description applies to the type of P. rudget. Panicum dasytrichum Spreng. Syst. Veg. 1: 317. 1825. ‘‘Brasil.”” The type, in the Sprengel Herbarium, was collected by Hoff- mansegg. Panicum hirsutum Willd.; Spreng. Syst. Veg. 1: 317. 1825, not Swartz, 1797. This is given as a synonym under P. dasytrichum and is credited to ‘‘W. herb.” The type, in the Willdenow Herbarium, was collected 3 i Fig. 133.—P. rudgei. From Salzmann’s by Hoffmansegg in Brazil. specimen of P. rigens. Panicum rhigiophyllum Steud. Syn. Pl. Glum. 1: 76.1854. ‘‘P.rigens. Salzm. Hrbr. Bahia.’”’ This specimen was not found in the Steudel Herbarium, but a Salzmann specimen bearing this name was examined at Halle. @ The following species, though not North American, may here be described on account of its relationship to Panicum rudgei: Panicum rotundum sp. noy. Plants perennial, in small tufts; culms 30 to 50 em. high, rather stout, stiff, erect or somewhat geniculate at base, densely ascending-hirsute, the nodes densely bearded; sheaths, except the upper, mostly shorter than the internodes, hirsute like the culms; ligules membranaceous, ciliate, the hairs mingling with those of the blade; blades thick, erect or appressed, linear, 5 to 20 cm. long, 2 to 5 mm. wide, more or less involute, at least toward the long-acuminate apex, only as wide as the sheath at base, the juncture obscure, densely hirsute on the upper surface, harshly velvety beneath; panicles terminal and in the axils of the upper 1 to3 leaves, forming an oblong inflorescence as in P. rudgei, about one-third the height of the plant, the main axis pilose, the slender, angled, scabrous, stiff but flexuous branchlets as- cending or spreading, pilose in the axils, the long pedicels divergent; spikelets 2.3 to 2.5 mm. long, 1 to 1.2 mm. wide, very turgid, abruptly pointed, strongly nerved, a few stiff, appressed hairs here and there between the nerves; first glume over half the length of the spikelet, abruptly pointed; sec- ond glume and sterile lemma subequal, exceeding the fruit, the lemma subtending a palea and staminate flower; fruit 1.8 mm. long, 1 mm. wide, ellipsoid, smooth and shining, a broad scar at the base. Fia. 134.—P. rotundum. From type specimen. e 140 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum rigens Salzm.; Steud. Syn. Pl. Glum. 1:76. 1854, not Swartz, 1788. This is given asa synonym under P. rhigiophyllum Steud. Salzmann specimens from Bahia, bearing this name have been examined in the herbaria at Munich and Halle and in the United States National Herbarium. Panicum cayennense divaricatum Doell in Mart. Fl. Bras. 2?: 220. 1877. Based on “Panicum scoparium Rudge * * * non Lam. * * * nec Michauz.”’ DESCRIPTION. Plants perennial, yellow-green or tawny; culms robust, 30 to 100 cm. high, erect or somewhat geniculate at base, often zigzag, especially above, densely and harshly villous; sheaths nearly equaling the internodes or overlapping, densely papillose- villous or hirsute; ligules membranaceous, ciliate, about 1 mm. long, the hairs of the ligule blending with the hairs of the upper surface of the blade: blades thick, linear, 15 to 40 cm. long, 5 to 10 mm. wide, rather rigidly ascending, flat or folded, slightly narrowed toward the base, gradually long-acuminate, densely short-hirsute on both surfaces or glabrate; panicles terminal and in the axils of the approximate upper 2 to 6 leaves, forming an oblong inflorescence one-third the height of the plant, or more, each more or less included at base in the subtending sheath; branches pilose in the axils, branching freely from the base, the branches and branchlets angled, scabrous, the lower branchlets stiffly ascending, the upper and the long pedicels divaricate, somewhat flexuous; spikelets about 3.5 mm. long, 1.5 mm. wide, turgid, somewhat attenuate at base, strongly nerved, sparsely hirsute, the stiff hairs irregularly dis- tributed; first glume about two-thirds the length of the spikelets, acuminate; second glume and sterile lemma subequal, exceeding the fruit, abruptly pointed, the latter subtending a palea and staminate flower; fruit 2.1 mm. long, 1.1 mm. wide, elliptic, smooth and shining, a cartilaginous flap-like appendage at the base. All the cited specimens from Costa Rica have pubescent blades, while several of those from South America have glabrate blades, e. g., Rusby & Squires 362, Spruce 93. These specimens with glabrate blades, though apparently less common than those with pubescent blades, are the typical form described by Meyer. DISTRIBUTION. Sayannas, Costa Rica to Brazil. Costa Rica: Buenos Aires, Pittier 10576, Tonduz 3679, 4875; Los Palmaies, Pit- tier 10588; Helechales del General, Pittier 12064. VENEZUELA: Santa Catalina, Rusby & Squires 362. British Guiana: Jenman 5978. DutcH Guiana: Surinam, Hostmann 642. Braziu: Para, Spruce 93; Bahia, Salzmann; Organ Mountains, Wilkes Expl. Exped. 9; without locality, Gardner 1178. Type U.S. National Herbarium no. 824039, collected in 1845, in the Province of Minas Geraes, Brazil, by Widgren, ‘‘Ex herb. Brasil. Regnellian. Musei bot. Stock- holm,” and distributed as ‘‘ Panicum cayennense Lam.” This species is related to P. rudgei with which and P. cayennense it has been confused. DISTRIBUTION. Brazil. The data on the labels of the specimens examined do not include habitat. Braziu: Prov. Minas Geraes, Henschen & Regnall 111 1367, Widgren in 1845; with- out locality, Burchell A 101-2 (Gray Herb.), Riedel. —-" HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 141 81. Panicum megiston Schult. Panicum altissimum Meyer, Prim. Fl. Esseq. 63. 1818, not DC. 1817.2 ‘‘In sylvis humidis plantationis Hof van Holland,’’ Essequebo or British Guiana. We have seena portion of the type in the Trinius Herbarium. The type is in the Géttingen Herbarium. Panicum megiston Schult. Mant. 2: 248. 1824. Based on P. altissimwm Meyer. Panicum: tuberculatum Presl, Rel. Haenk. 1: 307. 1830. The locality given by Presl is, “‘ Hab. in Luzonia.’? The type, in the herbarium of the German University at Prague, is labeled ‘‘Luzonia,’’ but it probably came from Mexico. Panicum elatior{us] Kunth, Rév. Gram. 1: 38.1829. Based on P. altissimum Meyer. . Panicum equisetum Nees; Doell in Mart. Fl. Bras. 2?: 206.1877. This is given as a synonym under P. megiston Schult., and is credited to ‘‘ Nees ab Esenbeck in herb. Reg. Berolinensis schedula.’’ The type, in the Berlin Herbarium, was collected by Sello in Bahia, Brazil. DESCRIPTION. Plants perennial; culms tall and robust, glabrous; sheaths papillose-hispid or papillose only; ligules fimbriate, about 1.5 mm. long; blades firm, ascending, 15 to 40 cm. or more long, 1.5 to 3:cm. wide, linear-lanceolate, slightly narrowed to the rounded base, glabrous; panicles finally exserted, 40 to 60 cm. long, the stiff main axis striate-angled, smooth or scabrous, the branches in distant verticils, often as many as 20 to 30 in a verticil, 10 to 20 cm. long, slender, stiffly or sinuously ascending, very scabrous, nearly simple, bearing the scattered, short-pediceled spike- lets along the upper half or third; spikelets usually purplish at maturity, about 3.4 mm. long, 1.5 mm. wide, globular-obovoid, glabrous; first glume scarcely one-third the length of the spikelet, pointed; second glume slightly shorter than the sterile lemma, both abruptly apiculate, 7 to 9-nerved, the glume about equaling the fruit, the sterile palea rather firm, about as long as the fruit; fruit 2.8 mm. long, 1.4 mm. wide, abruptly pointed, smooth and shining. Fic. 135.—P. megiston. From type Meyer states that the culms ascend to a height of 20 specimen of P.altissimum Meyer. or 30 feet and that they are much branched. Our specimens are all, with the exception of Wright 3872, the simple upper part of the culm only. Eggers, on the label accompanying his no. 14345, gives the height as 6 to 8 feet. The Wright specimen, except for the underground portion, is entire. This is simple and measures but 1.2 meters in height. This species somewhat resembles P. oaxacense and P. procerrimum of the genus Lasiacis, but the fruit has not the form and texture characteristic of that genus. DISTRIBUTION. In moist woods, Mexico and Cuba to Paraguay. Mexico: San Juan Bautista, Rovirosa 532. Cusa: St. Cruz de los Pinos, Wright 3872. VENEZUELA: Santa Catalina, Rusby & Squires 355. Brazit: Without locality, Riedel 1239, Gardner 1179. ParaGcuay: Morong 813, 1072. Ecuapvor: Balao, Eggers 14345. ‘a Roem. & Schult. Syst. Veg. 2: 457. 1817. The diagnosis is here referred to De Candolle ‘‘ Hornem. Hort. Hafn.1.p. 84.’’ The latter work we have not seen, nor that cited by the Index Kewensis, ‘‘Elench. Hort. Monsp. 42. 1805,” for P. ‘“‘altissi- mum Brouss., * * * nomen.’ In any case the name P. altissimum is preoc- cupied. 142 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Subgenus DICHANTHELIUM subgen. nov. Perennial, from a crown, rarely from short, matted rootstocks, surrounded by a more or less well-marked rosette of usually short winter leaves, in spring producing simple culms with mostly narrowly lanceolate blades and terminal panicles with numerous’ spikelets, these rarely perfecting seed; the early culms branching at some or all of the nodes (in a few species from the base only) after the maturity of the primary pani- cles or sometimes before; the branches often repeatedly branching, the short branchlets more or less fascicled and bearing usually much reduced leaves; the terminal one or two joints of the primary culms often finally falling, the whole producing an autumnal form usually strikingly different from the vernal form; the secondary panicles re- duced, the latest more or less included in the sheaths, cleistogamous and perfecting their grains. The type species is P. dichotomum L. In this group there is an intermediate stage of branching, in which the plants do not show the characteristic vernal nor autumnal habit. Vernal culms are sometimes produced on plants during the branched condition, because of renewal of activity, due to increased moisture, excess of nutriment, injury, or other causes. SYNOPSIS OF*GROUPS. Blades elongated, not over 5 mm. wide, 20 times as long as wide; autumnal form branching from the base only (from sy the lower nodes ta > WerIlEr) .2.03 2555 fede oat on bas eae DepauPERATA (p. 151). Blades not elongated, (or if so, more than 5 mm. wide and autumnal form not branching from base). Plants branching from the base, finally forming rosettes or cushions, foliage soft and lax; blades prominently ciliate CXCEPL ih 2 LAr ORME: ooaek's See eee acu ee Soe Rea LAXIFLORA (p. 158). Plants branching from the culm nodes or rarely remaining simple. Blades long, stiff; autumnal form bushy-branched above. Spikelets turgid, attenuate at base; mostly pustulose- pubescent; blades conspicuously striate, tapering rom. Wase Wo mpem. ).'... 2. 2. ae eee eee. see ANGUSTIFOLIA (p. 165). Spikelets scarcely turgid, not attenuate at base; blades tapering tovpotmr ends... 20... 52 yee ea Te BICKNELLIANA (p. 176). Blades not long and stiff (somewhat so in P. oligo- santhes, P. malacon, P. commonsianum, and P. equilaterale); not bushy-branched. Plants not forming a distinct winter rosette; spikelets attenuate at base, papillose... o..cs.82.0055.-F.-. PEDICELLATA (p. 292). Plants forming a distinct winter rosette; spikelets not attenuate at base. Spikelets turgid, blunt, strongly nerved (not strongly turgid in P. oligosanthes); blades rarely as much ~ as 1.5 cm. wide (sometimes 2 cm. in P. rav- enellii and P. xanthophysum). Sheaths, or some of them, papillose-hispid (some- ; times all glabrous in P. helleri); spikelets 3 to 4 ) mm. long (2.7 to 3 mm. in P. wilcorianum)....OLIGOSANTHIA (p. 278). Sheaths glabrous or minutely puberulent; spikelets 1.5 to 2.5 mm. long, unsymmetrically pyri- fori; ‘ CUlMawity..c25e. Cees abv. ea LANCEARIA (p. 271). HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 143 Spikelets not turgid, blunt, nor strongly nerved (see, however, P. roanokense and P. caerulescens.) Ligule of conspicuous hairs, usually 3 to 5 mm. long. Sheaths glabrous or only the lowermost some- i oe oe eae Spreta (p. 200). Sheaths strongly pubescent...............-.-.- LANUGINOSA (p. 208). 6 Ligule obsolete or nearly so (manifest in P. vis- cidellum, P. oricola, P. tsugetorum and P. cur- tifolium.) Spikelets spherical at maturity; blades glabrous, firm, cordate; plants sparingly branching. ..SpPHAEROCARPA (p.250). Spikelets usually obovate or elliptic. Blades of mid-culm elongated, less than 1.5 cm. wide; culms usually tall; spikelets pointed, abruptly so in the velvety P. scoparium week PS oiseigellny 2 ee Lyne Spikelets not over 3 mm. long; blades rather thin; sheaths glabrous or sparsely hispid...........----- 170. Spikelets less than 3 mm. long. Second glume and sterile lemma exceeding the fruit and pointed beyond it; spikelets 2.2 to 2.9 mm. long. Blades clustered toward the base...-...-.......------ 101. Blades not clustered toward the base. Sheaths, at least the secondary, hispid..........-.- 181. Sheaths glabrous. Bladestinmnamatthlo mim lOngse.seeetes eee oe 182. * Blades thin; fruit nearly 2 mm. long.............-111. Second glume and sterile lemma not pointed beyond the fruit. Ligule manifest, | to 3 mm. long. Culms rather stout; ligule 2 to 3 mm. long; sheaths PLANOUSE A eae: nis 5. /t sak Wyote Gun ue SPM ere 116 Culms slender; ligule 1 mm. long; sheaths sparsely PIULOSE Scene ee tin co ais bieeibla seis Woes aie Sete 159 Ligule obsolete. Spikelets 1.5 mm. or less long. INOdesibEardedmene:: oc 5 cen ee ek S eeeen ers gee 102. Nodes not bearded. Gulmsrandsbladespilaseas ov seins ee eee eee 90. Culms glabrous. Blades conspicuously ciliate; plants branching at baseronmliys O52 2a kee Ree ae ae 89. Blades not ciliate; plants branching from mid- dle or upper nodes. Vernal culm 50 cm. or more high; spikelets turgid, strongly nerved; autumnal form erect, with fascicled branches shorter than the primary internodes.........-- 113. Vernal culms usually much less than 50 cm. high; autumnal form spreading or re- clining. Spikelets 1.1 to 1.2 mm. long; blades rarely as much as 5 cm. long.....-.--- 160. Spikelets 1.2 to 1.4 mm. long. Blades elongated, at least some of them 8 fomMOrcm along .<2. a le Aeneas 161. Blades not over 3 cm. long......------- 157. Spikelets 2 mm. or more long. Blades elongated, some of them 20 times as long as __ wide; spikelets 2.2 to 2.8 mm. long. Blades erect; branches, when present, from the IGWhr MOU GH ODLY 2. cactauh tia Ate = oc eee 85. Blades spreading; branches from upper nodes... 99. ty ‘vy . depauperatum. . scribnerianum.* . helleri. . nudicaule. . scabriusculum. . eryptanthum. . yadkinense. . Spretum. . curtifolium. . microcar pon. . sirigosum. . polycaulon.. . caerulescens. . chamaelonche. . glabrifolium. ensifolium. . wernert. . bicknellir. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Blades not elongated, about 10 times as long as wide. Culms soon prostrate, vine-like; branches divari- cate. Plants bright green; culms lax; spikelets not over 2.1 mm. long Plants grayish green; culms stiff; spikelets 2.5 Culms not vine-like; Blades not over 6 mm. wide; fruit ainantl Culms wiry, crisp-puberulent; blades ciliate branches not divaricate. Spikelets 2.3 to 2.6 mm. long. Blades, or some of them, at least 8 mm. wide; fruit papillose-roughened......-...-...-- anMmmingls SoC es Se? ate Spikelets 2 mm. long. Culms glabrous; blades not ciliate. Blades Spikelets pubescent. Spikelets 3 mm. or more long. erect, firm; spikelets turgid, strongly nerved; plants grayish olive. Blades spreading; spikelets not turgid. Nodes glabrous; autumnal form erect, branched like a little tree.......... Nodes, at least the lowest, 167. 164. 112. 109. usually bearded; autumnal form topheavy- reclining. ... long as wide. Secondary panicles from basal sheaths only. Spikelets pointed, about 3.5 mm. long Spikelets blunt, 3 mm. or less long i Secondary panicles from upper branches. Spikelets attenuate at base, pustulose-pubescent; lowermost sheaths softly villous 110. Blades elongated, those of the mid-culm at least 15 timesas 95. Spikelets not attenuate at base, not pustulose; lower- most sheaths glabrous or hispid. Upper leaves approximate; sheaths glabrous. ..... Upper leaves distant; at least lower sheaths hispid . wide. Blades velvety-pubescent beneath. Spikelets 3 mm. long; plants velvety-villous through- Spikelets 4 mm. or more long. Sheaths ascending-hirsute; ligule 3 to 4 mm. long. . Sheaths downy-pubescent; ligule obsolete......... Blades not velvety-pubescent beneath. Sheaths glabrous or minutely puberulent only. Nodes bearded; spikelets 4 mm. or more long...... Nodes not bearded; spikelets not over 3.8 mm. long. Spikelets 3.5 to 3.8 mm. long; blades 2 cm. or more wide 41616°—voLt 15—10——-10 187. Blades not elongated, usually less than 10 times as long as 145 . lucidum. . sphagnicola. . webbertanum. . patentifolium. . lancearium. . roanokense. . dichotomum. . barbulatum. . depauperatum. . perlongum. P. fusiforme. . equilaterale. . aculeatum. . maleophyllum. . ravenelii. . boscit molle. . boscii. . latifolium. . 146 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. : Spikelets scarcely more than 3 mm. long. Spikelets turgid, blunt; blades mostly less CHS ACM WIE! eo Eee Sree ae ti eee 170. P. helleri. Spikelets not turgid; blades more than 1 cm. wide. Panicle narrow, branches ascending; spike- lets on long stiff pedicels...........--- 100. P. calliphyllum. Panicle as broad as long, branches spreading. Plants glaucous; basal blades conspicu- OUSIW MoMGO Joe ohh. soc Jb anise oes’ be 185. P. mutabile. Plants not glaucous; basal blades not cili- ate, orat the base only. Culms erect, or autumnal form leaning; blades symmetrical, broadly cor- Gate watss 4 cakes eee eee Pe oe 184. P. commutatum. Culms decumbent; blades usually un- symmetrical and falcate; narrowed to the scarcely cordate base.....-.. 186. P. joorit. Sheaths pubescent. Pubescence ascending or appressed. Spikelets 3 to 3.2 mm. long; first glume conspicu- OUnly TEMNOLC oss 2k ios wn Bvicg Dae =< 142. P. malacon. Spikelets 3.5 to 4 mm. long; first glume not TEIMOUO Meee an sac c ewelte dno ee sme sae eee eee 172. P. oligosanthes. Pubescence spreading, sometimes sparse. Plants robust, about 1 meter high; blades usually Figm, Gkanore Wide; 22. 2c. sce oss eee 189. P. clandestinum. Plants rarely more than 50 cm. high; blades rarely over 1.5 cm. wide. Panicles about as wide as long; blades ascend- ing or spreading. Spikelets attenuate at base, 3.5 to 4 mm. long soos. 2. sos RES eas See PEDICELLATA Spikelets not attenuate at base, not over 3.3 (p. 292). mm. long. Spikelets 3.2 to 3.3 mm. long, blades firm; sheaths, or some of them, more or less Rispid...2 «i232 hod Se eee 171. P. scribnerianum. Spikelets not over 3 mm. long; blades rather thin; sheaths, or some of them, glabrous or sparsely hispid.....----.. 170. P. helleri. Panicles narrow, branches erect (sometimes ascending in P. wilcoxianum), or spreading at anthesis only; blades erect. Spikelets not over 3 mm. long; blades not OveIouim, wide... Cede skieneeeine 168. P. wilcoxianum. Spikelets 3.7 to 4 mm. long; blades 8 to 20 mm. wide. Blades papillose-hispid.............------ 174. P. leibergii. Blades glabrous on both surfaces.......-- 175. P. xanthophysum. Spikelets less than 3 mm. long. Blades elongated, not over 5 mm. wide; secondary pani- cles at the base only or wanting. - oS 4 t 7) Culms single or few in a tuft; spikelets turgid, 2.7 to 3 HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 147 . Linearifolium. BARD g MO ce MEER Co nin on a bine eek es 83. P. perlongum. Culms in large tufts; spikelets not turgid, not over 2.7 mm. long. ORGS DOH be teee oo oO sc Se ait biota ein ww ew ov 84. Sheaths glabrous.............-. nt) BOs Blades usually not elongated; hecondary Spantieles, not at the base. Spikelets attenuate at base, mostly prominently pustu- lose; blades narrow, stiff, strongly nerved, tapering from base to apex. Nodes bearded; plants grayish-villous; autumnal blades flat. TIPPER EEIENY NOUN Coon id otis ca gerne gd Ate Sls s 92. Spikelets 2.5 to 2.8 mm. long.............-......- 93. Nodes not bearded; plants villous only at the base, or nearly glabrous. Autumnal blades flat; lower panicle branches spreading or deflexed..............--..-...-- 94, Autumnal blades involute; lower panicle branches more or less ascending. Plants glabrous or nearly so; autumnal culms erect. Spikelets subsecund along the suberect panicle Dranchoss (St chew coches a he 8 es 5 ete 98. Spikelets not subsecund; panicle loose and Plants pubescent, at least on the lower half. Spikelets about 2.4 mm. long; vernal blades 7 to 12cm. long, autumnal blades not falcate. 96. Spikelets not over 2 mm. long; vernal blades 4 to 6 cm. long, autumnal blades falcate.. 91. Spikelets not attenuate at base. Sheaths retrorsely pilose; blades soft and lax. Panicle branches ascending, forming a compact panicle; spikelets 1.6 mm. long...............87a. Panicle branches loosely spreading. Blades ciliate and more or less pilose on the sur- face; spikelets 2 mm. long.. bake o. Ot Blades glabrous or nearly so on the surface and margin; spikelets 2.2 mm. long. ........... 86. Sheaths not retrorsely pilose. Ligule manifest, mostly 2 to 5 mm. long. Sheaths, or all but the lowest glabrous; spike- lets not over 1.6 mm. long. Panicle narrow, one-fourth to one-third as wide Panicle open, nearly as wide as long. Spikelew 10 mm, long......2.552265....-..117. Spikelets 1.1 mm. long...............------119. Sheaths pubescent. P £. - £ i of ag! wernert. . chrysopsidifo- lium. . consanguineum. . angustifolium. . neuranthum. . ovinum. . arenicoloides. . aciculare. . xalapense stric- tirameum. . calapense. . laxiflorum. . spretum. lindheimeri. longiligulatum. 148 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Spikelets abruptly pointed; blades cordate; MexICODE os Lares sed sateen Leo 3s sees 179. P. viscidellum. Spikelets not pointed at maturity; blades not cordate. Ligule 1 mm. long; sheaths sparsely pilose; spikelets.14 mm-lonp...:-=.-.---.-.0% 159. Ligule usually more than 1 mm. long. Ligule 1 to 1.5 mm. long; culms and sheaths appressed-pubescent; spike- lets 1.5 to 1.9 mm. long. Spikelets 1.8 to 1.9 mm. long; plants UNE NYOTeeH! se ete et ee eee 146. Spikelets 1.5mm. long, nearly globular; plants olivacequs:. -: 2.25. 5ee2.% 148. Ligule 2 to 5 mm. long. Spikelets 1 to 1.3 mm. long; culms and sheaths softly appressed-pubescent. Spikelets 1.2 to 1.8 mm. long......... 118. Spikelets not over 1 mm. long........120. Spikelets mostly more than 1.5 mm. P P . curtifolium. . tsugetorum. ° . oricola. . leucothriz. . wrightianum. long, if less, pubescence spreading. See LANUGINOSA Ligule obsolete or less than 1 mm. long. Nodes bearded (P. scoparitum may appear to be bearded). Spikelets nearly 3 mm. long; plants velvety- Villousihnouchoutdssi0c fos ceeeoes SO. 169. Spikelets rarely as much as 2.5 mm. long; plants not pubescent throughout. Spikelets 1.5 to 1.6 mm. long............... 102. Spikelets 2 mm. or more long. Blades all velvety; autumnal form spar- anply branched: ...<.es222 23 Maes. kee 105. Blades glabrous, or only the lower pubes- cent or velvety. Spikelets 2 mm. long; autumnal form profusely branching. Fruits slightly exposed at maturity; upper sheaths viscid-spotted. -.... 103. Fruits covered at maturity; sheaths not viscid-spotted; Mexican. ....104. Spikelets 2.2 mm. or more long; autum- nal form less profusely branching. Sheaths and upper nodes glabrous. . . . 107. Lower sheaths and all nodes pubes- COntesaioe ts Jee eee ee ee 106. Nodes not bearded. Plants densely gray-velvety throughout, a viscid, glabrous ring below the nodes..... 178. Plants not gray-velvety. Sheaths or some of them pilose or hispid. Pubescence papillose-hispid. Spikelets ovate, pointed, 2.3 to 2.6 mm. (p. 208). . malacophyllum. . microcar pon. . annulum. . nitidum. . multirameum. . clutet. mattamuskee- tense. . scoparium. . scabriusculum. =), ee HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 149 Spikelets obovate, obtuse, nearly 3 mm. long. Blades about 2 cm. wide. .......-.-- 189. P. clandestinum. Blades not over 6 mm. wide......... 168. P. wilcovianum. Pubescence ascending-pilose. Spikelets 2 to 2.5 mm. long. Winter blades elongated, 5 to 10 cm. long; plants bluish green; spike- Reis ean, Wn ss oe a yh 2 ted’ 145. P. wilmingtonense. Winter blades 1 to 3 cm. long; plants olivaceous. Spikelets about 2.4 mm. long; pan- icle open, branches _ stiffly BS SNE a ee at baa i tee he 143. P.commonsianum. Spikelets 2 to 2.1 mm. long; panicle rather dense, branches ascend- tee ROE rege ee aS 3 ee 144. P. addisonii. Spikelets not over 1.7 mm. long. Blades white-margined; spikelets 1.6 to 1.7 mm. long, elliptic......... 152. P. tenue. Blades not white-margined; spikelets 1.3 to 14 mm. long, nearly filo] US eae e)” . ° Sana 147a.P. columbianum thinium. Sheaths glabrous or puberulent only. Spikelets spherieal, not over 1.8 mm. long; blades cordate, ciliate at base........ See SPHAEROCARPA (p. 250). Spikelets not spherical. Culms soon prostrate, vine-like; branches divaricate. Plants bright green; culms lax; spike- lets not over 2.1 mm. long........ 114. P. lucidum. Plants grayish green; culms. stiff; spikelets 2.5 mm. long.......... 115. P. sphagnicola. Culms not vine-like; branches not divari- cate. Spikelets unsymmetrically pyriform, strongly nerved; culms wiry..... See Lancearia (p. 271). Spikelets not pyriform. Blades elongated, especially the upper, about 20 times as long as wide; spikelets about 2.5 mm. long, on long pedicels... .. ee re 99. P. bicknellii. Blades not elongated. (See con- ; tinuation.) (Continuation. ) Spikelets 2 mm. or more long. Spikelets 2.5 to 3 mm. long; blades cordate, usually 1 cm. or more wide. Plants glaucous; basal blades conspicuously ciliate.. 185. P. mutabile. Plants not glaucous; basal blades ciliate at base only. Blades nearly linear, that is; with parallel mar- gins; first glume about half as long as the spikelet; Mekican..............- beaece 188. P.albomaculatum. 150 Blades lanceolate; first glume not more than one- third as long as the spikelet. Culms crisp-puberulent; blades rarely over 1 cm. wide; spikelets about 2.5 mm. long Culms glabrous or obscurely puberulent; blades usually 1.5 cm. or more wide; spikelets 2.7 to'S mm. longs: .--'... - 184. Spikelets not over 2.3 mm. long; blades not cordate, usu- ally less than 1 cm. wide. Blades conspicuously ciliate, soft and lax, crowded at the sbasGetns scree teeter eect ees eres Blades not ciliate or so only at base, not crowded at the base of the culm. Blades not over 6 mm. wide; plants not branch- ing or rarely branching from near the base Blades 7 mm. or more wide; plants branching from middle and upper nodes. Primary blades spreading; Leese eee fruit exposed at summit. Primary blades erect; nmieles eat ae COVELEO\c. 2 Socket ae ee ee eas 108. Spikelets not over 1.7 mm. long. Culms crisp-puberulent; spikelets turgid............-.- 147. Culms glabrous. Blades white-margined, firm. Blades puberulent beneath, often above......-- 152. Blades glabrous. Uppermost blades much reduced; culms branching from lower nodes only, the branches repeatedly branching... . . - 153. Uppermost blades about as long as the others; culms bearing short branches from middle and upper nodes.......--.---- 154. Blades not white-margined or very obscurely so (or if white margin is evident, spikelets only 1.1 mm. long). Culms branching only at ate plants soft, light green. oe. Culms penis at ue Modes” Spikelets 1.1 mm. long; winter blades bluish Preen; not glossyii 2. t so Seen ee 156. Spikelets 1.3 to 1.5 mm. long. Blades involute, falcate, with long stiff hairs on margin near base; plants BLT AN Wiry <2). di eocck eet eee 162. Blades not involute or at tip only, not falcate. Plants bright green; winter blades conspicuous, glossy green. . Plants olive; winter blades not con- spicuous nor glossy........- 157. oLOye . -158. - 155. Pe ‘os CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. . ashei. . commutatum. . ciliatum. . werneri. . clutei. . boreale. . columbianum. . tenue. . albomargina- tum. . trifolium. . vernale. . concinnius. . breve. flavovirens. ensifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 151 Depauperata.—Culms simple, mostly 10 to 40 cm. high; ligules less than 1 mm. long; blades much elongated, 5 to 35 cm. long, 2 to 5 mm. wide, narrowed at the base, long-acuminate at apex, basal blades shorter, but not forming a distinct. rosette in the autumn; spikelets 2.2 to 3.8 mm. long, strongly 7 to 9-nerved. Autumnal form bearing simple branches from the basal or lower nodes, the reduced panicles more or less concealed in the foliage at the base of the plants. Spikelets about 3.5 mm. long, beaked...........-....-.---- 82. P. depauperatum. Spikelets 3 mm. long or less, (sometimes 3.2 mm. long in P. per- longum) not beaked. Culms single or few in a tuft; spikelets turgid, blunt, 2.7 to 3.2.mm. lone; prairie plants..-.-...-------..-- 83. P. perlongum. Culms in large tufts; spikelets not turgid, 2.2 to 2.7 mm. long; plants of dry woods. Sheaths pilose; spikelets 2.2 to 2.7 mm. long, pilose.. 84. P. linearifolium. Sheaths glabrous; spikelets 2.2 to 2.3 mm. long; glab- maid ar sparingly pilose... .— 2 -2---------- 85. P. werneri. 82. Panicum depauperatum Muhl. Panicum strictum Pursh, Fl. Amer. Sept. 1: 69. 1814, not R. Br. 1810. ‘‘On the banks of the Delaware, Pennsylvania.’’ The type, in Kew Herbarium, has pilose sheaths. Panicum depauperatum Muhl. Descr. Gram. 112.1817. ‘‘Habitat in glareosis, floret Maio, Junio, Penns. Carolina.” The type is in the Muhlenberg Herba- rium. Muhlenberg described the species as having glabrous or pubescent leaves, pilose sheaths, glabrous spikelets, and fertile floret a little shorter than the second glume and sterile lemma. This description applies to the plant that has generally been referred to P. depauperatum rather than to the one with smaller spikelets later dis- tinguished as P. linearifolium, although in Muhlenberg’s herbarium specimens of both species are included in the same cover. Furthermore, some specimens with large spikelets have pilose and others glabrous sheaths. Of these, a specimen with pilose sheaths and spikelets 3.5 mm. long has been chosen as the type and has been so indi- cated by attaching a note to the specimen. Panicum rectum Roem. & Schult. Syst. Veg. 2: 457. 1817. Based on P. strictum Pursh, the original description of which is copied. Panicum involutum Torr. Fl. North. & Mid. U. S. 144. 1823. ‘Near Deerfield, Massachusetts. Cooley.’’ The type, in the Torrey Herbarium, is a small clump with culms 20 to 30 cm. high, sparsely pilose sheaths, involute blades with a few hairs on under surface, overmature primary panicles 4 to 5 cm. long, and spikelets 3.8 mm. long, the second glume and sterile lemma with a few hairs; secondary panicles with nearly mature spikelets are present at base. The sheet bears two labels, one, ‘‘From Dr. Cooley, Mass.,’’ the other in Torrey’s handwriting bears the name ‘‘ Panicum involu- tum,*’’ [Torrey used an asterisk to indicate his own species] followed by a diagnosis. Panicum muhlenbergii Spreng. Syst. Veg. 1: 314. 1825. Sprengel states nothing as to the source of his specimen other than ‘‘Amer. bor. (P. acwminatum Muhl.).”’ Since this name immediately follows P. acuminatum Swartz it seems evident that Sprengel meant to name the species which Muhlenberg described@ as Panicum acuminatum Swartz. There is no specimen bearing this name in the Muhlenberg Herbarium. In the Sprengel Herbarium the specimen labeled ‘‘Panicum Muhlen- bergii,’’ and which must be taken as the type, is P. depauperatum Muhl. This is from “Pine barrens, N. Jers., from Dr. Torrey.”’ Panicum junceum Trin. Gram. Pan. 220. 1826. ‘Trinius states that his specimen is from North America and called P. acuminatum by Sprengel. Such a specimen @ Descr. Gram. 125. 1817. 152 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. could not be found in the Trinius Herbarium. Since Sprengel cites ‘‘P. acwminatum Muhl.”’ under his P. muhlenbergii his specimen or a part of it is doubtless the specimen referred to by Trinius. The description well applies to an involute-leaved plant of P. depauperatum. Panicum sprengel Kunth, Rév. Gram. 1: 39. 1829. Based on P. muhlenbergii Spreng., without explanation as to change of name. Panicum depauperatum involutum Wood, Class-book 786. 1861. Based on P. involutum Torr. ? Panicum depauperatum laxalum] Vasey, U. 8S. Dept. Agr. Div. Bot. Bull. 8: 29. 1889. ‘‘Virginia, Florida, Texas, Arkansas, Missouri.’’ Described as ‘‘weaker stemmed, panicle with longer and more spreading branches, * * * spikelets smaller.’’ No type is indicated and there is no specimen in the National Herbarium so marked by Doctor Vasey. The description would seem to apply to P. linearifo- lium, but the range given is south of that in which that species is common, and no specimens of it from any of the States mentioned, except one each from Missouri, Arkansas, and Texas which are labeled ‘‘ Panicum depauperatum Muhl.”’ in Vasey’s writing, were in the National Herbarium in the time of Doctor Vasey. Hitchcock’s no. 1354, Stone Mountain, Georgia, with spikelets 3.1 to 3.2 mm. long, and rather loose panicles may represent Vasey’s variety. DESCRIPTION. Vernal form with culms several to many in a tuft, 20 to 40 cm. high, slender but rather stiff, erect or spreading at the summit, glabrous, puberulent or sometimes pilose; nodes ascending-pubeseent; sheaths, except the lowest, shorter than the internodes, glabrous to papillose-pilose; blades linear, 6 to 15 cm. long, 2 to 5 mm. wide (the lower shorter), often involute in drying, scabrous on both surfaces, sometimes pubescent beneath; panicles exserted, usually not much exceeding the leaves, 4 to 8 cm. long, rarely longer, few-flowered, the rather strict, remote branches narrowly ascending at maturity; spikelets 3.2 to 3.8 mm. long, rarely only 3 mm. oras much as 4 mm. long, 1.5 to 1.7 mm. wide, elliptical, pointed, glabrous or sparsely pubescent; first glume one-third to half the length of the spikelet, subacute; second glume and sterile lemma equal, extending beyond the fruit, forming a beak, strongly 7 to 9-nerved; fruit 2.1 to 2.3 mm. long, 1.4 to 1.5mm. wide, oval, minutely umbonate at the apex. Autumnal form similar to the ver- nal, the reduced secondary panicles produced on branches from the basal or lower nodes, more or less concealed in the tuft of basal leaves. This species is variable as to pubes- cence and size of spikelets. The Fic. 136.—P, depauperatum. Fromtypespecimen. Spikelets of the type specimen are 3.5 mm. long, those of the glabrous plants on the same sheet in the Muhlenberg Herbarium are 3.9 mm. long. Many New Eng- land specimens and occasional specimens from elsewhere have spikelets only 3 mm. long. This form is represented by Chamberlain 298, Chase 3379, and Parlin 1957 from Maine; Burgess in 1893 from Massachusetts; Perron in 1876 from Pennsylvania, and Lansing 2743 from Indiana. The difference in length is often due to the inrolling of the summit of the second glume and sterile lemma, but the glumes show greater pro- portion of variation than usual in this genus. Asa rule the spikelets do not vary on the same plant, but Chase 2402, has spikelets 3 mm. long on the terminal panicle and 3.5 to 4mm. long on the basal panicles. The fruit shows little variation in size. \; . - HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 158 DISTRIBUTION. Open sterile woods, Maine to Minnesota, south to Georgia and Texas. Marne: Canton, Parlin 1957; Chesterville, Chase 3283, 3316; Fayette, Chase 3379; Cumberland, Chamberlain 298. New Hampsuire: Sanbornton, Carter 100 (Hitchcock Herb.). Vermont: Burlington, Flynn in 1902. Massacuusetts: Ipswich, Boott. Connecticut: Hartford, Wilson 1257; Montville, Graves in 1897; Southington, Andrews 58, Bissell 5533. RuopeE Isianp: Gloucester, Collins in 1908. New York: Bronx, Bicknell in 1896; Woodmere, Bicknell in 1905; Rockville Center, Bicknell in 1906; Norwood, Bicknell in 1903; Long Island, Bicknell in 1905. Ontario: Sarnia, Macoun 26322; Toronto, Biltmore Herb. 797c. New Jersey: Atco, Painter 699; Eagle Rock, Mackenzie 1452. PENNSYLVANIA: Lancaster County, Heller 4775; Easton, Porter in 1897; Penryn, Small in 1889; Broad Moun- tain, Pretz 1959. InpiaANnA: Clark, Bebb 513; Miller, Chase 1540, Lansing 2743, Um- bach 1657. Inurnots: Lansing, Chase 867; Ma- kanda, Gleason 1022. Micuican: Port Huron, Dodge 78. Wisconsin: Clear Lake, Cheney 1216. Mrynesora: Hennepin County, Sandberg in 1890; Nicollet, Fic. 137.—Distribution of P. depawperatum. Ballard in 1892. Missouri: St. Louis, Eggert 233; Monteer, Bush 4654; Lees Summit, Bush 3936. Kansas: Lindsborg, Plank 18. DELAWARE: Mount Cuba, Commons 29. MARYLAND: Chesapeake Junction; Hitchcock 2414; West Chevy Chase, Hitchcock 341; Great Falls, Chase 2864, in Kneucker Gram. Exs. 548. District or CotumBiA: Chase 2284, 2402, Hitchcock 340, House 947, Pollard 338, Ward in 1876. Virermia: Luray, Steele 229. Nortu Carona: Biltmore, Biltmore Herb. 797; Watauga County, Small & Heller 279; Chapel Hill, Chase 3053; Henderson County, Biltmore Herb. 797a. Sours Carona: Clemson College, House 2161. Geroraia: Stone Mountain, Eggert 39, Hitchcock 1354, 1360; Augusta, Cuthbert 388. TENNESSEE: Knoxville, Ruth 63; Ducktown, Chambliss 89. ALABAMA: Auburn, Zarle & Baker in 1897. Mississirri: Jackson, Hitchcock 1303. ARKANSAS: Fulton, Bush 2350 (Gray Herb.). Lovurs1aNna: Shreveport, Hitchcock 1249. Texas: Denison, Bebb 2663. 83. Panicum perlongum Nash. Panicum perlongum Nash, Bull. Torrey Club 26: 575. 1899. ‘On prairies and dry soil, Illinois to North Dakota, south to Indian Territory. Type collected in Indian Territory at Creek Nation, by M. A. Carl[e]ton, April 25, 1891, No. 98.’’ The type, 154 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. in Nash’s herbarium, consists of five clumps of several culms each, 7 to 40 cm. high, with mature primary and immature secondary panicles, and spikelets 3 to 3.2 mm. long. Panicum pammeli Ashe, N. C. Agr. Exp. Sta. Bull. 175: 116. 1900. “Prairies of Iowa, June.’’ The type, in Ashe’s herbarium, collected by R. I. Cratty, June 12, 1881, has spikelets 3.1 to 3.2 mm. long. DESCRIPTION. Vernal form similar to that of P. depawperatum, more strict in habit, and in smaller tufts, more constantly pilose and usually papillose, the blades on the average longer and narrower, sometimes 25 cm. long, pubescent on the lower surface; panicles smallerand narrower, the branches erect, hence appearing more densely flowered; spikelets 2.7 to 3.2 mm. long, 1.6 to 1.7 mm. wide, oval, blunt, sparingly pilose; first glume one-fourth to one-third the length of the spikelet, acute or obtuse; second glume and sterile lemma equal, obtuse, not extended beyond the fruit, strongly 7 to 9-nerved; fruit 2.4 mm. long, 1.5 to 1.6 mm. wide, obovate- oval, rounded and minutely umbonate at the summit. Autumnal form with secondary panicles usually more numerous than in P. depau- peratum and sometimes produced from the second node. Fig. 138.—P. perlongum. From type specimen. DISTRIBUTION. Prairies and dry soil, Michigan and Manitoba to Texas. Inprana: Elkhart County, Deam 6753.4 Inuinois: Hanover, Gleason & Gates 2530; Naperville, Umbach 1670; Wady Petra, V. H. Chase 460, 1158, 1731, in Kneucker Gram. Exs. 547; Peoria, Brendel, McDonald 18. MicuicAN: Keweenaw County, Farwell 755. Wisconsin: Racine, Wadmond in 1901. Minnesota: Lake City, Manning in 1883 (Gray Herb.). Manirosa: Lake Winnipeg Valley, Bourgeau in 1837 (Gray Herb.). Souta Daxota: Custer, Rydberg 1100; Clark, Griffiths 863. Towa: Ames, Ball 30, 145; Clinton, Vasey; Iowa City, Somes 245. Nesraska: Ewing, Bates 1120. Missouri: Lees Summit, Bush 3089. Kansas: Manhattan, Hitchcock 2501, 2509. Texas: Llano County, Nealley 79; Wallisville, Wallis 38. OxtaHoma: Creek Nation, Carleton 98. Fia. 139.— Distribution of P. perlongum. @ This specimen and those of P. xalapense, P. ashei, and P. commutatum from Indiana were received too late for representation in the maps. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 155 84. Panicum linearifolium Scribn. Panicum linearifolium Scribn. in Britt. & Brown, Illust. Fl. 3: 500. f. 268a. June, 1898. ‘‘Dry soil, especially hillsides, New York and New Jersey to Missouri.’’ This was again published @ as ‘‘n. sp.’’ a few days later. ‘‘New England, southward to Vir- ginia and westward to Texas.’’ Both descriptions state that the sheaths are glabrous or pilose, both illustrations indicate pilose sheaths. The type, in the National Her- barium, is marked ‘“‘P. linearifolium Scribn. Type.”’ in Scribner’s handwriting, and is labeled ‘‘ Washington, D. C., Vasey, 1882,’’ but was probably collected in Maryland, along the Potomac northwest of Washington, where this species is frequent in rocky woods. The specimen is a tuft of culms 30 to 40 cm. high, with pilose sheaths, mature primary panicles, and much reduced, nearly hidden secondary ones. The spikelets are 2.2 to 2.4 mm. long. DESCRIPTION. Vernal form light green, in dense tufts, often surrounded by the withered, persistent, more or less curled leaves of the previous year, the culms readily separating, 20 to 45 em. high, very slender, erect, spreading or almost drooping at the summit, glabrous, minutely puberulent or rarely pilose; sheaths usually equaling or exceeding the inter- nodes, sparsely to densely papillose-pilose, the papillse often obscure; blades elongated and erect, usually overtopping the panicles until maturity, 10 to 35 cm. long (the lower shorter), 2 to 4 mm. wide, scabrous on both surfaces or often pubescent on the lower, rarely on the upper surface, usually ciliate near the base with long hairs; pani- cles finally long-exserted, 5 to 10 cm. long, half to two-thirds as wide, rather few- flowered, the scabrous, flexuous branches re- mote, ascending; spikelets 2.2 to 2.7 mm. long, 1.3 to 1.5 mm. wide, oblong-elliptic, obtuse, sparsely pilose with weak, spreading hairs; first glume one-fourth to one-third the length of the spikelet, obtuse, or pointed by the inrolling of the margins; second glume and sterile lem- ma equal and equaling the fruit at maturity; fruit 2 to 2.1 mm. long, 1.2 mm. wide, oval, obecarely umbonate : f \ he suxanut. Fig. 140.—P. linearifolium. From Autumnal form similar, the reduced sec- type specimen. ondary panicles produced on short basal branches mostly concealed in the tuft of basal leaves. While the typical form of this species is quite distinct from P.depauperatum, occa- sional specimens, such as the following, seem to be intermediate between the two: Burnham 24, Bush 1555, 4734; Hitchcock P|. Kans. 880; Plank 40. In these the spike- lets are about 3 mm. long and sometimes obscurely short-pointed. The following specimens have the sheaths glabrous or nearly so and approach the closely allied P. werneri: Bissell 5541, Bush 4411A, Deam, Wells County, Indiana, in 1901, Hitchcock 598, Pollard, Washington, D. C., in 1897, Rose & Painter 8153. DISTRIBUTION. Dry woods, Maine to Kansas, south to Georgia and Texas. Marne: Chesterville, Chase 3326; Fayette, Chase 3393; Canton, Parlin 1971. Vermont: Barnet, Blanchard in 1888; Burlington, Hitchcock 598. Massacuusetrts: Williamstown, Churchill in 1901. Connecticut: Southington, Andrews 49, Bissell 5541, 5542; Fairfield, Hames in | 1895. 4 Scribn. U.S. Dept. Agr. Div. Agrost. Bull. 11: 42. pl. 7. July 20, 1898. 156 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. New York: Thousand Islands, Robinson & Maxon 20; Oxford, Coville in 1884; Washington County, Burnham 24; Hempstead, Bicknell in 1903; Jamaica, Bicknell in 1905. OnTaARIO: Galt, Herriot 34. New Jersey: Wildwood, Pollard in 1897; Morris County, Mackenzie 1339, 1398; Springdale, Pretz 1882. PENNSYLVANIA: Newtown, Smith 156; York County, Rose & Painter 8153. Outo: Barnesville, Laughlin 6906. InprANA: Wells County, Deam in 1901; Kosciusko County, Deam 3218. Inurnors: Wheaton, Moffatt 255 in 1893; Makanda, Gleason in 1903; Cobden, Waite in 1885. MicuicaN: Keweenaw County, Farwell 597; Agricultural College, Lake in 1888. Minnesora: Lake Kilpatrick, Bal- lard in 1893 (Univ. Minn. Herb.). Missouri: Monteer, Bush 731 in part, 742, 288la, 4734; Eagle Rock, Bush 153; Pleasant Grove, Bush 320; Carthage, Bush 1555; Swan, Bush 4533, 4549; Chadwick, Bush 4411, 4411A; Pilot Knob, Ward in 1878. Kansas: Manhattan, Hitchcock 2506, Pl. Kans. 880. MaRYLAND: Cabin John, Chase 5411; Plummers Island, Hitchcock 597; Great Falls, Chase 2303. District or CoLumsia: Pollard in 1897, Vasey in 1882. West Virainia: Harpers Ferry, Hitchcock in 1905. GeroraiA: Silver Creek, Biltmore Herb. 7079a (Biltmore Herb.). Kentucky: Lexington, Short 8 (Gray Herb.). Mississippi: Without locality, Johnson in 1886. ARKANSAS: Benton County, Plank 45, 55, 96, 100, 148, 157; northwest Arkansas, Harvey 7. Louisiana: New Orleans, Ridell (Gray Herb.). Texas: Palestine, Plank 40; Jacksonville, Plank in 1894; without locality, Nealley in 1890, Wright (Gray Herb.). OxiaHoma: Sapulpa, Bush 1107 in 1895 (Mo. Bot. Gard. Herb.). Fig. 141.— Distribution of P. linearifolium. 85. Panicum werneri Scribn. Panicum werneri Scribn. in Britt. & Brown, Illust. Fl. 3: 501. f. 268b. 1898. “Dry knolls in swamps, New York and Ohio.’’ The type, in Hitchcock’s herbarium, is a specimen collected by William C. Werner, near Painesville, Ohio, 1889, no. 60. It consists of eight simple culms, mostly lacking the base, glabrous except for the sparsely bearded nodes, with over-mature panicles; spikelets almost glabrous, the sparse hairs obscure. Panicum delawarense Ashe, N. C. Agr. Exp. Sta. Bull. 175: 116. 1900. ‘‘Dry soil near Centreville, Del. Collected by A. Commons, July 6, 1878.’’ The type, in Ashe’s herbarium, consists of four solitary culms with long-exserted, over-mature panicles, and sparsely pubescent spikelets. DESCRIPTION. Vernal form similar to that of P. linearifolium, typical specimens differing as follows: Culms stiffer, nodes usually sparingly pilose, sheaths glabrous, often shorter than the internodes; blades firmer, shorter and wider, 15 cm. long or less, the lower culm blades HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 157 3.5 to 6 cm. long, 3 to 6 mm. wide, a few long hairs at the rounded base, scabrous on both surfaces, not pubescent; spikelets 2.1 to 2.4 mm. long, 1.2 to 1.3 mm. wide, nearly or quite glabrous. Autumnal form similar to the vernal, remaining simple or late in the season bearing simple branches from the lower, rarely from the basal, nodes. The above-mentioned types are both of this form, but material examined shows a much less clear distinction from P. linearifolium, with which this species seems to intergrade. The division is here based on a com- bination of stiffer habit, glabrous sheaths, shorter, broader, and firmer blades, and less pubescent spikelets. In many specimens, however, having the other characters enumerated the blades are as long as in many specimens of P. linearifolium. The following represent these intergrading specimens: Biltmore Herb. 8342, Chase 3299, 3382, Jones, Bur- lington, Vermont, in 1892, Knight 55, 57, Parlin 1190, Porter, Easton, Pennsylvania, in 1895, 1897, and 1898. In habit, especially as seen in the field, P. werneri often suggests P. depauperatum. The following specimens, with slightly pointed spikelets 2.5 to 2.8 mm. long, approach that species: Briggs 1248, Graves 7, Eggleston 1757, H. W. Merrill 44. Fie. 142.—P. werneri. From type specimen. DISTRIBUTION. Sterile woods and knolls, Maine to Minnesota, south to Ohio and Texas. Marne: Penobscot County, Fernald 506, Knight 55, 57; Chesterville, Chase 3299; Fayette, Chase 3382; North Berwick, Parlin 1190, 1502, 1806; Hiram, H. W. Merrill 44, 47; Orono, Briggs 1248. New Hampsuire: Laconia, Carter 101, 242 (Hitchcock Herb.). Vermont: Burlington, Jones in 1892; Eggleston 1757. Massacuusetts: Sheffield, Hoffmann in 1901. Connecticut: Ledyard, Graves 16; Volunteer, Graves 17; Frank- lin, Woodward in 1906; South- ington, Chamberlain & Bissell in 1903; Waterford, Graves 7. New York: Tripoli, Burnham in 1897; Ithaca, Ashe, Rowlee in 1892; New York, Bicknell in 1895. Ontario: Toronto, Biltmore Herb. 8342; Kingston, Fowler in 1898; Kingston Mills, Klugh in 1907; Algonquin Park, Macoun 21957. New Jersey: Berkeley Heights, Mackenzie 2251. PENNSYLVANIA: Easton, Porter in 1895, 1897, and 1898. Onto: Painesville, Werner 60, 65 (both in Hitchcock Herb.). Micaican: Flint, Clark (Field Mus. Herb.). Wisconsin: Rainbow Rapids, Cheney 1345. Minnesota: Chisago County, Sandberg in 1886. Missouri: Swan, Bush 2913, 2926. DELAWARE: Centerville, Commons 358. Texas: Dallas, Reverchon in 1876 (Gray Herb.). Fia. 143.—Distribution of P. werneri. 158 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Laxiflora.—Plants light green, vernal culms 10 to 40 cm. high, numerous in tufts; blades flat, soft, mostly ciliate, basal blades shorter, but not forming true rosettes in the autumn; ligules nearly obsolete; primary panicles long-exserted ; spikelets 1.3 to 2.8 mm. long, obovate, obtuse, turgid, 5 to 7-nerved. Autumnal form freely branching near the base, forming close, flat, soft tufts, the reduced panicles often exceeded by the leaves. Sheaths retrorsely pilose; spikelets papillose-pilose. Panicle branches ascending, forming a rather compact panicle; spikelets 1.6mm. long.....2..--....1..-- 87a. P.xalapense stric- tirameum. Panicle branches loosely spreading. Blades ciliate and more or less pilose on the surface; spikelets’ 2mm lonee eo eee. Poe Aes 87. P. xalapense. Blades glabrous or nearly so on the surface and mar- eins ppikelete 2:2 mimi lone 20s seek. ac! Seis - 86. P. laxiflorum. Sheaths not retrorsely pilose; spikelets pubescent or glabrous. Spikelets pubescent, about 2 mm. long.....----.-------- 88. P. ciliatum. Spikelets glabrous. Blades glabrous on the surface. .2....2::-.-+---...-. 89. P. polycaulon. Blades pilose‘onsthe surface... .. 2.2). -262- e452. 22 90. P. strigosum. 86. Panicum laxiflorum Lam. Panicum laxiflorum Lam. Encycl. 4: 748.1798. ‘‘Cette plante est dans l’herbier du Muséum. Je la crois d’Amérique Septentrionale.’’ The type, labeled in Lamarck’s hand ‘‘panicum laxiflorum lam. dict.,’’ is in the Lamarck Herbarium. It consists of two culms, each with a loose terminal panicle, one leafless, the other with a single blade glabrous on both surfaces; the spikelets are 2.2 mm. long, papillose-pilose, the fruit covered by the second glume and sterile lemma. These characters indicate not the species that has been called by this name in America, but the one of more restricted southern range. Panicum pyriforme Nash, Bull. Torrey Club 26: 579.1899. ‘‘Type collected by the writer in clay soil, at Orange Bend,¢ Lake Co., Florida, March 12-31, 1894, no. 239.” The type, in Nash’s herbarium, consists of a clump of three culms, 15 to 45 em. high. The description states that the blades are glabrous on the margin, but the type, as well as duplicate types in the National and Columbia University herbaria and in Hitchcock’s herbarium, has sev- eral sparingly ciliate blades; the spikelets are said to be ‘about 2.5 mm. long” but measure 2.2 mm. Panicum aureum Muhl.; Scribn. & Merr. U.S. Dept. Agr. Div. Agrost. Cire. 27: 4.1900. This is mentioned as a synonym of P. laxiflorum Lam. The type specimen is in the Muhlenberg Herba- rium in folio ‘‘187, Panicum strigosum.”’ It is labeled ‘‘117 P. aureum M 115.” NAS We Fic. 144.—P. laxiflorum. From type specimen. DESCRIPTION. Vernal form with slender culms 20 to 60 cm. high, erect, or the lower nodes often geniculate, glabrous; nodes bearded with reflexed hairs; sheaths shorter than the internodes, conspicuously retrorsely pilose; blades mostly 10 to 18 cm. long, 7 to 12 mm. wide, acuminate, narrowed toward the base, glabrous on both surfaces and on aThe locality of Nash 239 as given on the label is ‘‘vicinity of Eustis.” Orange Bend is a few miles northwest of Eustis. “ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 159 the margins, or sparsely ciliate; panicles long-exserted, but sometimes equaled by the long upper blades, 8 to 12 cm. long, nearly as wide, lax, few-flowered, the branches flexuous, spreading, the lower often deflexed; spikelets 2.2 to 2.3 mm. long, 1.2 mm. wide, oblong-obovate, obtuse, first glume one-third to two-fifths as long as the spikelet; second glume and sterile lemma equal and covering the fruit at maturity, papillose-pilose; fruit 1.8 mm. long, 1.2 mm.wide, obovate-elliptic, minutely umbonate. Autumnal form branching at the base, forming soft, spreading tufts, the sheaths overlapping and the blades but little or not at all reduced, much exceeding the sec- ondary panicles; spikelets more turgid and obtuse than the primary ones. A specimen collected by Lester F. and Rosamond Ward at Palatka, Fla., in 1891, and another from Dr. Chapman, collected in Florida, without locality or date, have pilose blades like those of P. xalapense; the spikelets are 2.2 to 2.3 mm. long, and the fruit is covered by the equal second glume and sterile lemma. DISTRIBUTION. Rich or damp woods, Georgia to Florida and Alabama. GeoraiA: Stone Mountain, Egger 42; Ocmulgee River Swamp, below Macon, Small in 1895. Fioripa: Monticello, Combs 327; Fig. 145.—Distribution of P. laxiflorum. Lake City, Nash 2156, Hitch- cock 1009; Gainesville, Chase 4217, Combs 743; Mabel, Curtiss 6635 in part;¢ Grasmere, Combs 1064; Spruce Creek, Curtiss in 1885; Manatee, Tracy 7383; Palmetto, Tracy 6707; Eustis, Nash 239, 2034; Orange Bend, Chase 4101; Dunnellon, Combs 909; Titusville, Chase 4023; Ormond, Hitchcock 111. ALABAMA: Springhill, Mohr in 1895. 87. Panicum xalapense H. B. K. Panicum xalapense H. B. K. Nov. Gen. & Sp. 1: 103. 1816. ‘‘Crescit in regno Mexicano prope Xalapa et montem Macultepec.’’ The type, in the Bonpland Herba- rium, is labeled, ‘‘In regno Mexicano prope xalapa, regione temperata.”’ Panicum pumilum Bosc; Nees, Agrost. Bras. 228. 1829, not Lam. 1798. ‘‘(Herb. Willd.) Habitat in America boreali.’’ This is mentioned as a synonym under P. lazi- florum Lam. The specimen referred to, in the Willdenow Herbarium, is the vernal form. Panicum rariflorum Rupr. Bull. Acad. Roy. Belg. 97: 240. 1842, not Lam. 1798. (Coll. H. Galfeotti] No. 5733). Nous avons trouvé cette nouvelle espéce * * * pres de Xalapa.’”’ Thisisanomen nudum. The type, in the Brussels Herbarium, is a poor specimen, but undoubtedly belongs to this species. Panicum ruprechti Fourn. Mex. Pl. 2: 21. 1886, not Fenzl, 1854. This name is based on the type of P. rariflorum Rupr., ‘‘(GaL[Eorri] n. 5733).’’ It was earlier listed without description by Hemsley,> based on ‘‘Panicum rariflorum Rupr. * * * non Lam.,’’ that is, a nomen nudum based on a nomen nudum. Panicum caricifolium Scribn.; Ashe, Journ. Elisha Mitchell Soc. 15:¢ 57. 1898. This is given as a synonym of P. laziflorum Lam. “As distributed by Kearney ‘a Curtiss included P. xalapense also under this number. U.S. National Herbarium no. 388470 is a sheet of Curtiss 6635, with a tuft of each species. b Biol. Centr. Amer. Bot. 3: 495. 1885. ¢ The title page, vol. 15, pt. 1 (pp. 1-75) is incorrectly numbered 4 (IV). ‘ 160 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. (Washington, D. C., May, 1897).’’? The Kearney specimen referred to could not be found in Ashe’s herbarium, but other specimens there under this name are P. zala- pense, as are specimens in the National Herbarium bearing the name “Panicum caricifolium” in Scribner’s writing. This is the species described as P. laxiflorum by American authors. DESCRIPTION. Vernal form similar to P. laxiflorum in texture and habit; culms and blades on the average shorter, the blades pilose on one or both surfaces or nearly glabrous, usually short-ciliate, the uppermost more or less convolute at base around the culm; panicles hardly so few- flowered; spikelets 1.9 to 2 mm. long, 1.1 mm. wide, oblong-obovate, obtuse, the first glume one-fourth to one-third as long as the spikelet; second glume and sterile lemma pilose, less prominently papillose, the glume shorter than the fruit, the latter 1.5 mm. long and 1 mm. wide, oval, minutely umbonate. Autumnal form as in P. laziflorum, but forming usually denser tufts with shorter leaves. This species is conspicuously retrorsely pilose on the sheaths. Canby’s no. 106, Stone Mountain, Ga., in the Gray Herbarium, is exceptional in having almost glabrous sheaths. Fig. 146.—P. zalapense. From type specimen. DISTRIBUTION. Woods, Maryland, Illinois, and Missouri to Florida, Texas, and Mexico; alsc in Santo Domingo. InpIANA: Clarke County, Deam 6883. InuiNois: Jackson County, French in 1905. Missourr: Campbell, Bush 749 in part; Swan, Bush 4638; Jefferson County, Eggert 127; Pacific, Kellogg 19. MaryLanp: Cabin John, Chase 5412, Chase in Kneucker Gram Exs. 549; west District Line, Hitchcock 343; Great Falls, Chase 2316, 2320. Virainia: Richmond, DeChalmot; Smyth County, Small in 1892; Princess Anne County, Kear- ney 1033, 1104, 1179, 1308, 1467, Pollard & Maxon in 1900. Norts Carourna: Biltmore, Bilt- more Herb. 2993a; Madison County, Biltmore Herb. 2993c; Raleigh, Ashe; Chapel Hill, Ashe, Chase 3057; Wilmington, Chase 3111, 3117, 3118. Fic. 147.—Distribution of P. ralapense. SoutH CarRoLina: Orangeburg, Hitchcock 1391, 1415, 1424; Clemson College, House 2177. Groraia: Thomson, Bartlett 1453; Rome, Curtiss 6791; Warm Springs, Tracy 8867; Augusta, Cuthbert 1119, Kearney 212; Union, Harper 1085; Clarke County, Harper 77; Dublin, Harper 2140; Stone Mountain, Chase 4517, Eggert 86, Harper 183, Hitchcock 1361, Small in 1893, Wilson 14. Fiormwa: Jacksonville, Curtiss 6602; Lake City, Combs & Rolfs 141; Tallahassee, Combs 390, Kearney 75; Sanford, Hitchcock 773; Titusville, Chase 4009; Dade County, Curtiss 5537, Eaton 589, 831, Hitchcock 659, 673, Tracy 8849; Manatee County, Tracy 6694; Sarasota Key, Tracy 7202; Alva, Hitchcock Lee Co. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 161 — Pl. 480; Tampa, Hitchcock 927, 935; Myers, Hitchcock 906; without locality, Rugel 392. Kentucky: Harlan County, Kearney 53. TENNESSEE: Knoxville, Ruth 68; Nashville, Gattinger in 1880; Wolf Creek, Kearney in 1894; Cocke County, Kearney 970. AtaBama: Auburn, Earle & Baker in 1897; Tracy 3759; Tuskegee, Carver 55; Scottsboro, Chase 4506. Mississippi: Meridian, Tracy3267; Dekalb, Tracy 3255, 3256; Fairport, Tracy 3211; Acona, Tracy 2058; Starkville, Tracy 1410, 1753; Enterprise, Tracy 3267, 3287; Jackson, Hitchcock 1310; Biloxi, Hitchcock 1073, Tracy 2032, 4574, 4588, 6358. ArxKansas: Fulton, Bush 1440; Texarkana, Bush 2488. Lovuistana: Calhoun, Ball 62, Hitchcock 1260; Coushatta, Ball 122; Shreveport, Cocks 3511, Hitchcock 1239; Lake Charles, Hitchcock 110, 1124; Opelousas, Langlois 36; New Orleans, Drummond 456, 457. Texas: Waller County, Hitchcock 1182, Thurow in 1898 and 1906; Dallas, Reverchon 93, Bush 651; Houston, Bebb 1236; Denison, Bebb 1457; Columbia, Bush 1273; Heiler 4085, 4209; Galveston, Plank 91; Palestine, Plank 51; Nealley in 1884. Ox.taHoma: Wister, Hitchcock in 1903; Poteau, Hitchcock in 1903 (both in Hitch- cock Herb.). Mexico: Jalapa, Pringle 8083, C. L. Smith 1752; Hidalgo, Pringle 13250; Orizaba, Nelson 201; Chinantla, Liebmann 328; Valley of Cérdoba, Bourgeawu 2162; Galeotti 5733 (Brussels Herb.), Schiede & Deppe ‘‘acuminatum c;” Seler 2160 (both in Berlin Herb.). Santo Domrnco: Near Jarabaco, Eggers 2129. 87a. Panicum xalapense strictirameum subsp. nov. DESCRIPTION. Differing from P. xalapense in having ovoid, more compact panicles with ascending branches, somewhat smaller spikelets, 1.7 mm. long, shorter, narrower blades, than common in the species, and shorter culms. Autumnal form in smaller, shorter tufts. Type U.S. National Herbarium no. 558449, col- lected April 28, 1906, Jackson, Miss., by A.S. Hitch- cock (no.1311). This specimen hasshort, appressed blades, the uppermost 2.5 to 4 cm. long, and long- Fig. 148.—P. rala triclirameum. - ees gecimen. “m™ ~~ exserted panicles 2 to 3 cm. long. This form was abundant on wooded hills in the outskirts of Jack- son, where P. zalapense also grew, but from which it differed conspicuously in the form of the panicle. Most of the other specimens referred to this subspecies are less characteristic, but are smaller, with smaller blades than the species usually has, though occasional speci- mens of P. zalapense have the smaller blades of the subspecies. DISTRIBUTION. Dry woods, South Carolina to Loui- siana. South Carona: Lancaster, House 2551. AvaBaMa: Auburn, Hitchcock 1333. Mississippi: Jackson, Hitchcock 1311; Madison, Tracy 1478. Lourstana: Calhoun, Hitchcock 1290; West Feliciana, Cocks 3510. 41616°—vo1 15—10——11 Fic. 149.—Distribution of P. zalapense strictirameum. 162 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 88. Panicum ciliatum Ell. Panicum ciliatum Ell. Bot. 8. C. & Ga. 1: 126. 1816. No locality is cited. The type, in the Elliott Herbarium, consists of two short culms with short-exserted, imma- ture panicles. Panicum leucoblepharis Trin. Clav. Agrost. 234. 1822. Trinius’s full citation is as follows: ‘‘1177. Gr. miliaceum americanum, majus, panicula minore. Pluk. Phytogr. p. 176. Tab. 92. f. 7. Mant. p. 95. (excl. Syn. Sloan. ut ipse Sloaneus monet). Citatur a Gronowio (Virg. p. 12.) ad Pan. paniculatum floribus muticis; sed quid illud? Figura bene convenit cum Panico quodam herb. notsr. [nostr.] ex Amer. bor. (Pan. leucoblepharis m.) praeter cilia foliorum elegantissima, rigidiuscula.—Synon. Recchii ap. Pluk. admodum dubium.’’ Plukenet cites ‘‘Nov. Hispan. Terent. apud Rece. 373” after the phrase name cited by Trinius. The figure, which is not identifi- able, represents blades with cordate bases and no ciliz. A specimen in the Trinius Herbarium is labeled: ‘‘ab Enslino in Am. bor. 1. dt. cl. Trattinick.”’ This is typ- ical P.ciliatum Ell. Trinius¢@ further describes this species, citing the Enslin speci- men. Since the Plukenet figure can not certainly be identified, the Enslin specimen has been chosen as the type. Panicum ciliatifolium Kunth, Rév. Gram. 1: 36. 1829. Based on P. ciliatum Ell. without description or explanation as to reason for change of name. Panicum ciliatifolium Desv. Opusc. 88. 1831. ‘‘Habitat in America boreali.” The type could not be found in the Desvaux Herbarium, but the description leaves no doubt as to the identity of the species. Desvaux cites P. ciliatum Ell. with a query. He uses the name ciliatifolium apparently without reference to its previous use by Kunth for the same species. DESCRIPTION. Vernal form with culms 5 to 30 cm. high, erect or spreading, sparsely pilose toward the summit, the nodes glabrous; sheaths ciliate on the margin, otherwise glabrous, usually overlapping; blades 3 to 6 cm. long, 3 to 8 mm. wide, the uppermost often much sthaller, lanceolate, ciliate on the margin with stiff hairs 2 to 3 mm. long, arising from papillz, panicles 3 to 4 cm. long, about as wide, with more numerous spikelets than those of P. xalapense, the branches spreading, flexuous, the axis pilose; spikelets 1.8 to2 mm. long, 1 mm. wide, obovate-elliptic; first glume half the length of the spikelet, subacute; second glume and sterilé lemma equal, not exceeding the fruit, villous; fruit 1.7 to 1.8 mm. long, 0.9 mm. wide, elliptic, the apex minutely umbonate. Autumnal form in flat, soft mats, similar to that of P. xalapense. A specimen of Nash 807 from Eustis, Fla., in Hitch- cock’s herbarium, which is an autumnal form of this species, consists of two small tufts, of which one has glabrous spikelets, the other pubescent spikelets; the plants are otherwise identical; the spikelets are of the same size and the fruits of the glabrous ones are umbonate as in the pubescent ones. So far as examined, other specimens of this number have ‘pubescent spikelets. Two specimens, Chase 3131 from Wilmington, N. C., and Hitch- cock 1062 from Biloxi, Miss., with pubescent spikelets only 1.6 mm. long are inter- mediate between P. ciliatum and P. polycaulon and might perhaps be considered as a form of P. polycaulon with pubescent spikelets. Fia. 150.—P. ciliatum. From type specimen. aGram. Pan. 219. 1826. - Low pine lands and hammocks, North Carolina to Florida and Louisiana. Norta Carouna: Onslow County, Chase 3184; Roanoke Island, Chase 3214, 3226; New Hanover County, Chase 4583, Hitchcock 1430, 1451, 1490, Kearney 250. SourH Carona: Orangeburg, Hitchcock 342, 1371. Frormwa: Baldwin, Combs 57, Hitchcock 991, 993, 1000; Lake City, Combs 100, 137, Hitch- cock 1019, 1036; Madison, Combs 288; Eustis, Nash 807. ALABAMA: Flomaton, AHitchcock 1040; Mobile, Kearney 24. Misstssrer1: Jackson County, Kear- ney 283, Tracy 162; Biloxi, Chase 4360, Kearney 326 in part; Avondale, Tracy 4582. Lovuistana: New Orleans, Drummond (Gray Herb.). HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 163 DISTRIBUTION. Fic. 151.—Distribution of P. ciliatum. 89. Panicum polycaulon Nash. Panicum polycaulon Nash, Bull. Torrey Club 24: 200. 1897. ‘‘Type specimen collected by the writer on August 20, 1895, in the flatwoods at Tampa, Florida, No. 2420a.’’ The type, in Nash’s herbarium, consists of a spreading tuft of numerous culms, 10 to 17 cm. high, the overmature panicles nearly devoid of spikelets, and the first glume half the length of the spikelets, which are 1.6 mm. long. DESCRIPTION. : ‘ Vernal form similar to that of P. ciliatum, culms rarely over 20 cm. high, glabrous, but pilose in the long-exserted panicle; sheaths sparingly ciliate; blades on the average narrower than those of P. ciliatum; spikelets 1.5 to 1.6 mm. leng (excep- tionally as much as 2 mm. long), 0.8 mm. wide, obovate, blunt, glabrous; first glume one-third to half the length of the spikelet, subacute; second glume and sterile lemma strongly nerved; fruit 1.4 mm. long, 0.8 mm. wide, elliptic, subacute, not umbonate. Autumnal form in flat, soft mats, similar to those of P. xalapense, but smaller. ; The type specimens of P. ciliatum and P. polycaulon differ in the spikelet char- acters, the former having pubescent spikelets 2 mm. long and the latter having gla- brous, more obovate, turgid spikelets 1.6 mm. long. Our numerous specimens, however, do not show these constant differences. Occasional specimens have glabrous spikelets as much as 2 mm. long. A comparatively few specimens have pubescent spikelets that are of the smaller size. We have not found any distinct differences.in habit that can be coérdinated with the spikelet characters. The range of the two forms is somewhat different, P. ciliatum extending from North Carolina to northern Florida, and P. polycaulon throughout Florida and southward into Cuba. It will be observed that the ranges of the two overlap in northern Florida and it is here that the intermediate specimens are found. A specimen from Tampa, Florida, Hitchcock 933, has blades nearly destitute of cilie. The following specimens have spikelets nearly or quite 2 mm. long: Fior- ma: Kalamazoo, Hitchcock 763; Lakeland, Hitchcock 836; Dunedin, Tracy 6698. Mississippi: Mississippi City, Hitchcock 1101. Fic. 152.—P. polycaulon. From type specimen. 164 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Flatwoods and hammocks, Florida and along the Gulf coast to Mississippi; also in Cuba. Froriwa: Live Oak, Tracy 6727; Washington County, Combs 649 in part; Apa- lachicola, Biltmore Herb. 6022a, Kearney 96; Orange County, Baker 68, Combs & Baker 1086, Curtiss 6627;; Orange Bend, Chase 4104; Titusville, Chase 3967; Dunedin, Tracy 6698, 6723; Kalamazoo, Mitch- cock 762; Sanford, Hitchcock 771, 772, 827; Manatee, Mitch- cock 950,974; Lakeland, Hitch- cock 843; Tampa, Combs 1338, Hitchcock 933, 943; Hog Island, Tracy 6710 in part; Lemon Bay, Tracy 7188 in part; Sneeds Island, Tracy 6692; Santa Rosa Island, Tracy 8411; Myers, Hitchcock 868, 903, 9234, Fig. 153.—Distribution of P. polycaulon. Lee Co. Pl. 482; Miami, Chase 3885, Hitchcock 112, 665, 711, 721; Homestead, Hitchcock 6894. AtaBaMA: Fort Morgan, Tracy 7208. Mississrppr: Horn Island, Tracy 6470; Petit Bois Island, Tracy 4606; Ocean . Springs, Skehan in 1895; Biloxi, Chase 4364. Cusa: Herradura, Hitchcock 115; without locality, Wright 3875 in part; Isle of Pines, Palmer & Riley 990. Porto Rico: Near Piedra Blanca, Sintenis 5724. 90. Panicum strigosum Muhl. Panicum strigosum Muhl. in Ell. Bot. 8. C. & Ga. 1: 126. 1816. Elliott cites no locality. The type, in the Elliott Herbarium, is a single immature plant, the panicle only short-exserted. The accompanying label reads ‘‘Panicum strigosum Muhl. Hab. in humidis. Car: & Georg: Flor: Ma-Jun:.”’ ‘ Panicum laxiflorum pubescens Vasey, Contr. Nat. Herb. 3: 30. 1892. No locality nor specimen is cited. Only two specimens bearing this name in Vasey’s writing can be found in the National Herbarium. One of these, Curtiss North American Plants No. H, Duval County, Florida, agrees well with Vasey’s description; the other, a specimen of P. zalapense, does not agree with the description. The first is therefore chosen as the type. Panicum longipedunculatum Scribn. Tenn. Agr. Exp. Sta. Bull. 7: 53. pl. 16. f. 61. 1894. ‘‘Damp woods, White Cliff Springs [Tennessee], July 1890; Tullohoma, July, 1892. A large form of this species is represented by No. 3597* A. H. Curtiss N. Am. Pl.”’ The first specimen cited, which is evidently the plant figured and which is chosen as the type, is in Hitchcock’s herbarium. It consists of two tufts with slender culms 15 to 40 cm. high, more sparsely pilose than usual. DESCRIPTION. Fig. 154.—P. strigosum. From type specimen. Vernal form similar to that of P. ciliatum, but having sparsely pilose culms and sheaths, bearded nodes, and blades on the average a little wider and more or less pilose on both surfaces; panicles larger, with pilose axis and branches, bearing more numerous, smaller, glabrous spikelets, the latter 1.3 to 1.5 mm. long, 0.7 mm. wide, obovate, less turgid than in other species HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 165 of this group; first glume one-third to half the length of the spikelet; second glume and sterile lemma equal, faintly nerved; fruit 1.3 mm. long, 0.6 to 0.7 mm. wide, elliptic, subacute, not umbonate. Autumnal form a dense mat with panicles scarcely rising above the leaves. This species is variable as to pubescence. Some of the specimens from Cuba and Guatemala show only a few scattered hairs upon the surface of some of the blades, but these are conspicuously ciliate on the margin. DISTRIBUTION, Sandy woods, Virginia and Tennessee to Florida and Louisiana; also in Mexico, Guatemala, and Cuba. » * Vrraria: Norfolk County, Kearney 1761. Norra Carouna: Roanoke Island, Chase 3244; Onslow County, Chase 3171; Wilmington, Hitchcock 1450, Kearney 282. Sours Carona: Aiken, Ravenel. Fiorina: Jacksonville, Curtiss 3597*, 4031; ‘Washington County, Combs 567, 584. TENNESSEE: White Cliff Springs, Scribner in 1890 (Hitchcock Herb.). ALABAMA: Cullman County, £g- gert 21; Gateswood, Tracy 8422; Flomaton, Hitchcock 1043. Fig. 155.—Distribution of P. strigosum. Mississtrr1: Saratoga, Tracy 8402. Lourstana: Lake Charles, Hitchcock 1162. Mexico: Minatitlan, J. G. Smith 555 (Hitchcock Herb.). GuaTEeMALA: Secanquim, Pittier 257; Cuesta de Peixh4, Pittier 1800. Cusa: El Guama, Palmer & Riley 213; without locality, Wright 3875 in part. Angustifolia.—Plants mostly dull grayish-green, cespitose; vernal culms erect or ascending from a spreading base, mostly 30 or 40 cm., rarely as much as 100 cm. high, appressed-villous at base or sometimes above, or rarely smooth even at base; ligules ciliate, less than 1 mm. long; blades narrow, ascending, usually firm and rigid, more or less striate with prominent nerves, and some- times longitudinally wrinkled besides, often ciliate at the base; spikelets attenuate at base, rather strongly 7-nerved, usually pubescent, the hairs aris- ing from bullate papille; first glume narrow and sheathing at base. Autum- nal culms repeatedly branching, forming bushy crowns, these remaining erect or becoming decumbent or widely spreading; blades much reduced, often involute; a distinct rosette of basal leaves formed in the fall. Species of the Atlantic Coastal Plain. Nodes bearded; plants grayish-villous; autumnal blades flat. ENR, MUM oe io oo bean aha denen dave Rene oo 92. P. chrysopsi- difolium. AE RE of a 93. P. consangui- neum. . Nodes not bearded; plants villous only at base, or nearly gla- brous; autumnal blades involute or flat. Autumnal blades flat; lower panicle branches spreading NRCG sa aan OF dads dae <> aides aed bene h ae des 94. P. angustifolium. 166 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Autumnal blades involute; lower panicle branches more or less ascending. Spikelets 3.3 to 3.5 mm. long, pointed.......--....-- 95. P. fusiforme. Spikelets less than 3 mm. long, not pointed, or ob- scurely so. Plants glabrous or nearly so; autumnal culms erect. Spikelets subsecund along the suberect pan- icle branchesayaeesaysoaee A-d 2 eis. 98. P. neuranthum. Spikelets not subsecund; panicle loose and (Oy Oe) Nees ee Sia he Ne Gee sre k= eg 97. P. ovinum. Plants pubescent, at least on the lower half. Spikelets about 2.4 mm., rarely only 2.1 mm. long; vernal blades 7 to 12 cm. long; autumnal blades not falcate........-- 96. P. arenicoloides. Spikelets not over 2 mm. long; vernal blades 4 to6cm. long, autumnal blades much Crow@ed,talcategs. ae ceee fa oeerea 91. P. aciculare. 91. Panicum aciculare Desv. Panicum aciculare Desv.; Poir. in Lam. Encycl. Suppl. 4: 274. 1816. ‘‘Cette plante croit dans les Indes orientales. (V.s. in herb. Desv.)’’ Poiret cites ‘‘P. acicu- lare Desv. Herb.’’ The type is in the Paris Herbarium. It is from the Desvaux Herbarium and is labeled in Desvaux’s handwriting ‘‘ Panicum aciculare Desy. in Enc. Suppl. 4. p. 274. habitat in india orientali.’’ It is the autumnal form and is evi- dently the specimen described. The locality is certainly an error, as no such plant is known from the East Indies, and the specimen is without doubt from the southeastern United States.¢ Panicum setaceum Muhl. Descr. Gram. 99.1817. ‘‘Habitat in Georgia.’’ The type, in the Muhlenberg Herbarium, is in a folio marked: ‘‘151 Panicum pungens M., 97 Elliott, 358.’’ Panicum pungens in the herbarium stands in the same relation to the other species as does P. setaceum in Muhlenberg’s book.» In the herbarium folio 151 is the first of the Panicums. In the book P. setacewm is the first species described under this genus. Likewise the sequence in the two places is essentially the same. The specimen, which is the autumnal form, agrees with Muhlenberg’s description of P. setaceum. Panicum subuniflorum Bosc; Spreng. Syst. Veg. 1: 312. 1825. ‘‘Carolin.’’ In the Delessert Herbarium is a specimen from ‘‘Carolina” collected by Bose. In the Will- denow Herbarium is another fragmentary specimen labeled P. subuniflorum Bosc, but without locality or collector. The latter may be the type. Panicum arenicolum{cola] Ashe, Journ. Elisha Mitchell Soc. 15: 56. 1898. ‘‘Type material collected by the writer at Chapel Hill, N. C. June 1898.’ The type could not be found in Ashe’s herbarium. In Hitchcock’s herbarium is a specimen labeled ‘Panicum arenaecolum Ashe” collected in the vicinity of Chapel Hill, North Carolina, by W. W. Ashe, and sent by him to Professor Scribner. The date of collection is not given. This specimen, which is probably a duplicate type, consists of two vernal culms, somewhat appressed-pubescent below; the stiffly ascending blades are gla- brous except the lowermost, which is sparsely pubescent beneath. Panicum filiram[eJum Ashe, Journ. Elisha Mitchell Soc. 16: 88. 1900. ‘‘Sandy woods, eastern North Carolina. Type material collected in New Hanover County, @ Several cases of erroneous localities occur in Desvaux’s herbarium. See Panicum illinonense Desv. under P. fasciculatum. bCompare U.S. Dept. Agr. Div. Agrost. Circ. 277: 2. 1900. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 167 N. C., in June 1899.” The type, in Ashe’s herbarium, consists of two single vernal plants, with slender, villous culms, sheaths less villous, blades nearly glabrous on one plant, sparsely long-pilose on the other, the panicles overmature. Panicum pungens Muhl.; Scribn. & Merr. U.S. Dept. Agr. Div. Agrost. Circ. 27: 2. 1900, not Poir. 1816. This is mentioned as a herbarium name of P. setacewm Muhl., of which it is a typonym. This is the species described in Britton’s Manual@ and in Small’s Flora> under the name Panicum neuranthum Griseb. DESCRIPTION. Vernal culms numerous in a tuft, ascending from a spreading base, appressed- pubescent below, glabrate above, 20 to 50 cm. or, in shaded situations, 60 cm. or more high, the nodes more or less pubescent but not bearded; lower sheaths villous, the upper glabrous except the ciliate margin; blades stiff, spreading or ascending, nar- rowed to an involute point, glabrous or the lower sparsely pilose, somewhat papillose- hispid on the margin at base, the middle culm blades 4 to 6 cm. long, rarely longer, 2 to 5 mm. wide, the uppermost shorter, usually only 1 to 2 cm. long and 1 to 2 mm. wide; panicles open, 3 to 7 cm. long, the flexuous branches spreading at maturity; spikelets 1.9 to 2 mm. long, 1.1 mm. wide, obovate, blunt, basal attenuation short; first glume about one-fourth the length of the spikelet, obtuse or pointed; second glume and sterile lemma equal, papillose- pubescent; fruit 1.6 mm. long, 1 mm. wide, oval- elliptic, glabrous at the apex. Autumnal form consisting of numerous bushy- branched culms 10 to 30 cm. long, spreading and forming dense cushions, the short blades involute, sharp-pointed and usually arcuate, mostly 1 to 3 cm. long; spikelets more turgid than in the vernal form. This species is abundant in the coast region and can be distinguished from all the other species of this group within its range by the small spikelets, and the awl-like blades of the autumnal state. Chase’s no. 3097} is referred here though it is unusually glabrous. Fic. 156.—P..aciculare. From typespeci- men in Florence Herbarium. DISTRIBUTION. Sandy pine woods of the Coastal Plain from New Jersey to northern Florida and eastern Texas. New Jersey: Cape May, Stone in 1909. VirGInia: Vicinity of Cape Henry, Chase 2357, 2936, 3682, 5413, 5414, Hitchcock 345, 347, Kearney 1375, 1566, 2038, Mackenzie 1664, Williams 3099. Nortu Carona: Lake Mattamuskeet, Chase 3205; Raleigh, Chase 3084; Wil- mington, Ashe in 1899, Chase 3123, 3130, 3157, 4578, 4586, Hitchcock 344, 1447,. 1473, Kearney 247, 284 in part; Roanoke Island, Ashe in Curtiss Dist. 6451, Chase 3215, 3216, 3217; Ocracoke Island, Kearney 2272; Edenton, Kearney 1871; Wilsons Mills, Chase 3094, 30974, 3103, 3105; Jacksonville, Chase 3167; Wards Mill, Chase 3185. Sourn Caro.ina: Orangeburg, Hitchcock 17, 346, 1375, 1404; Aiken, Ravenel in 1882; Fripps Island, Cuthbert 1166; Isle of Palms, Chase 4525. GeoraiA: Augusta, Kearney 215; Thomson, Bartlett 1444, 1455; Albany, Tracy 3640; Thomasville, Tracy 3638, 3641; Jessup, Biltmore Herb. 11866. @ Man. 84. 1901. > Fl. Southeast. U. 8. 95. 1903. 168 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Fuioriwa: Baldwin, Hitchcock 989; Lake City, Combs 183; Madison, Combs 216; Monticello, Combs 301, 329; Tallahassee, Combs 372, 380; Quincy, Combs 408; Washington County, Combs 554, 567a; Leon County, Curtiss E; Chatta- hoochee, Tracy 3639; Mari- anna, Tracy 3637; Milton, Chase 4302; Eustis, Nash 1243; Pasco County, Curtiss 6639. AtaBAMA: Tuskegee, Carver 96; Gateswood, Tracy 8420; Mo- bile, Kearney 21 in part. Mississier1: Starkville, Chase 4444; vicinity of Biloxi, Chase 4359, Hitchcock 1082, Kearney 2844, 306 in part, Tracy 1417, 3634. Fig. 157.—Distribution’of P. aciculare. Lovistana: Covington, Langlois A48b in 1884; Calcasieu, Cocks 2194; Lake Charles, Hitchcock 1127, 11394, 1140, Tracy 3650. Texas: Waller County, Hitchcock 1225, Thurow 1 in 1900. OKLAHOMA: Sapulpa, Bush 1388 in 1895 (Mo. Bot. Gard. Herb.). §2. Panicum chrysopsidifolium Nash. Panicum chrysopsidifolium Nash in Small, Fl. Southeast. U. 8. 100. 1903. On page 1327, in the list of new genera and species, the following citation is given: ‘‘Type, Cur- tiss, N. Am. Pl., no. D,in Herb.N. Y.B.G.’’ The type, in the herbarium of the New York Botanical Garden, is labeled ‘‘Hammock land, Leon Co. Fla., May 12, 1886,” and consists of a clump of four vernal culms 30 to 55 cm. high with mature, short- exserted panicles. DESCRIPTION. Vernal form with ascending or spreading, rather slender culms, 30 to 45 cm. high, purplish, grayish-villous, especially below, the nodes bearded; sheaths much shorter than the internodes, villous like the culm, densely so at the summit; blades 5 to 10 cm. long, 3 to 5 mm. wide, tapering from base to apex, conspicuously pointed, villous on both surfaces; panicles finally long-exserted, 4 to 6 cm. long, about three-fourths as wide, the flexuous branches ascending or spread- ing; spikelets 2 mm. long, 1.2 to 1.3 mm. wide, ob- ovate, blunt and turgid; first glume one-third the length of the spikelet, subacute or obtuse; second glume and sterile lemma subequal, scarcely cover- ing the fruit at maturity, villous, the bullate pa- pill prominent; fruit 1.7 mm. long, 1.2 mm. wide, broadly elliptic, minutely puberulent at the apex. Tig tes P ehyghvaelduoleuih. — rom Autumnal form spreading and forming mats, the type specimen. culms slender, often zigzag toward the tip; blades numerous, flat, becoming papery with age, mostly 1 to3 cm. long, 1.5 mm. wide; spikelets more turgid than usual in the primary panicle. Panicum chrysopsidifolium has been confused with P. newranthum Griseb., because both species were distributed by Wright under the same number (Wright 3453). This number in the Grisebach Herbarium is the type of P. newranthum. The same number in several other herbaria consists of the autumnal form of P. chrysopsidifolium.@ @ For further discussion of Wright’s Cuba grasses, see Hitchcock, Contr. Nat. Herb. 12: 183. 1909. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 169 The specimen referred to by Grisebach,@ under P. neuranthum as, ‘“‘forma ascendens, ramosa, foliis planis, spiculis ut in @” is P. chrysopsidifolium. This species can be distinguished from P. consanguineum by the smaller spikelets, and from P. aciculare by the bearded nodes and by the lax culms and flat blades of the autumnal form. DISTRIBUTION. Sandy pine woods of the Coastal Plain, Florida to Louisiana; also in Cuba and Porto Rico. Fiorma: Leon County, Curtiss D; Orange County, Baker 45; San- ford, Hitchcock 775. Sypris Lake Charles, Chase Fig. 159.—Distribution of P. chrysopsidifolium. 4405. Cusa: Consolacion del Sur, Palmer & Riley 481; Herradura, Hitchcock 116; eastern Cuba, Wright 3453 in part; Isle of Pines, Palmer & Riley 982. Porto Rico: Santurce, Heller 982; Las Marias ad Tabomeo, Sintenis 5985. 93. Panicum consanguineum Kunth. Panicum villosum Ell. Bot. S. C. & Ga. 1: 124. 1816, not Lam. 1791. No locality is cited. The type, in the Elliott Herbarium, consists of a single culm lacking the base, bearing four leaves and an immature, scarcely exserted panicle. The accom- panying label reads: ‘‘ Panicum villosum mihi. Hab. in umbrosis. Flor. Ap. Ma.” Panicum consanguineum Kunth, Rév. Gram. 1: 36. 1829. Based on P. villosum Ell., the name presumably changed because of P. villosum Lam. Panicum commutatum consanguineum Beal, Grasses N. Amer, 2: 141. 1896. Based on P. consanguinewm Kunth. ; Panicum georgianum Ashe, Journ. Elisha Mitchell Soc. 15: 36. 1898. ‘‘Georgia: Small; Darden [Darien] Junction, McIntosh Co., June 27, 1895.’’ The type speci- men, which is in the Biltmore Herbarium and which is marked ‘“‘P. georgianum W. W. Ashe,”’ in Ashe’s writing, is the autumnal form. Panicum cahoonianum Ashe, Journ. Elisha Mitchell Soc. 15:4 113.1899. Based on P. georgianum Ashe, the name changed because of Panicum georgicum Spreng. 1825. DESCRIPTION. Vernal form with culms ascending or spreading, often geniculate at base, 20 to 55 cm. high, rather stout, densely felty- villous below, less so above, nodes SIE chsgaiuinente, ‘Wrori type specimen bearded; sheaths villous, the upper often of P. villosum Ell. sparsely so; blades erect or ascending, 7. to 11 cm. long, 5 to 8 mm. wide, (the lowermost shorter and broader), tapering slightly toward the base, more or less invo- lute-pointed, villous on both surfaces or nearly glabrous above, the longitudinal @ Cat. Pl. Cub. 232. 1866. b The title page, vol. 15, pt. 2. (pp. 76-114) is incorrectly numbered 14. (XIV). 170 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. wrinkling conspicuous in the lower blades; panicles 4 to 8 cm. long, one-half to two- thirds as wide, the lower branches usually narrowly ascending; spikelets 2.6 to 2.8 mm. long, 1.6 to 1.8 mm. wide, obovate, blunt, turgid; first glume one-third the length of the spikelet or less; second glume and sterile lemma equal, scarcely cov- ering the fruit at maturity, densely papillose-villous, the bullate papillae prominent; fruit 2 mm. long, 1.5 to 1.7 mm. wide, minutely puberulent at the apex. Autumnal form spreading or decumbent, the numerous branches somewhat flabel- lately fascicled, the blades mostly 3 to 4 cm. long, 2 to 3 mm. wide, flat, thin, and papery. The vernal form of this species may be distinguished from P. angustifolium by the ereater amount of pubescence, the bearded nodes, and the ascending panicle-branches, and the autumnal form by the widely spreading habit and shorter blades. DISTRIBUTION. Sandy pine woods of the Coastal Plain from North Carolina to northern Florida and west to eastern Texas. VirainiA: Virginia Beach, Hitchcock in 1905. Norra Caroutna: Wilmington, Hitchcock 1449, 1471; Wards Mill, Onslow County, Chase 3174; Wilsons Mills, Chase 3095; Roanoke Island, Chase 3230. Souta Caro.ina: Orangeburg, Hitchcock 1373, 1382. Georata: Macon, Small in 1895; Darien Junction, Small in 1895. Fioripa: Baldwin, Hitchcock 986, 999; Washington County, Combs 570, 651; Marianna, Tracy 3633; Milton, Chase 4299; without locality, Rugel 142. ALABAMA: Gateswood, Tracy 8427. Mississippi: Vicinity of Biloxi, Fig. 161.—Distribution of P. consanguinewm. Chase 4351, Hitchcock 1069, 1070, 1083, Kearney 218 in part, 284 in part, Tracy 1884, 2873, 3661, 4614. ARKANSAS: Texarkana, Heller 4238. Lourstana: Lake Charles, Hitchcock 1139, 1155, Langlois in 1884. Texas: Beaumont, Reverchon 4156; Waller County, Thurow 23 in 1906. 94, Panicum angustifolium Ell. ? Panicum ramulosum Michx. Fl. Bor. Amer. 1: 50. 1803. ‘‘Hab. in sylvis Caro- linae.’’ The type specimen, in the Michaux Herbarium, labeled “‘in pratis caespi- tosis Carolinae,’’ belongs to a species of the Angustifolia, apparently P. angustifolium, but on account of the fragmentary condition of the type, which is devoid of spikelets, and the insufficiency of Michaux’s description the identity of this specimen can not be determined with certainty. Panicum angustifolium Ell. Bot. S. C. & Ga. 1: 129. 1816. No locality is cited. The type, in the Elliott Herbarium, consists of a single culm, lacking the base, bearing three leaves and a scarcely exserted, immature panicle; the culm is glabrous, the sheaths are sparingly pilose, densely ciliate on the margin toward the summit; blades 18 cm. long, involute toward the apex, long ciliate on the margin for half to two- thirds their length, otherwise glabrous. The accompanying label reads: “‘ Panicum angustifolium. Hab: in aridis. Flor. Ma.?” , | HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 171 Panicum curtisii Steud. Syn. Pl. Glum. 1: 66. 1854.. ‘‘ M. A. Curtis sub: P. ner- vosum. Miihlbrg. var. legitin Carolina.’’ The type in the Paris Herbarium, labeled by Steudel “Panicum curtisii Steud. Panicum nervosum Muhlb. var.? M. A. Curtis. Carolina australis, Commun. Lenormand,”’ is a somewhat fragmentary specimen, but appears to be P. angustifolium. DESCRIPTION. Vernal form with erect or nearly erect culms 30 to 55 cm. high, the lowermost inter- nodes gray crisp villous, the middle and upper glabrous; nodes glabrous or the lower villous, not bearded; lower sheaths more or less appressed-villous, the upper glabrous, except the usually ciliate margin; blades 8 to 12, rarely 15, cm. long, 4 to 8 mm. wide (lowermost blades shorter and broader and longitudinally wrinkled), stiffly ascending, the upper more appressed, long-acuminate, scarcely narrowed at base; panicles long- exserted, 4 to 10 cm. long, nearly as wide, loosely flowered, the branches at anthesis widely spreading, the lower 3 to 4 cm. long, often reflexed; spikelets 2.5 to 2.8 mm. long, 1.4 to 1.6 mm. wide, elliptic-obovate, turgid; first glume about one-third the length of the spikelet, pointed or obtuse; second glume and sterile lemma equal, covering the fruit at ma- turity, not beaked beyond it, papillose-villous; fruit 2 mm. long, 1.3 to 1.5 mm. wide, broadly elliptic, minutely puberulent at the obscurely umbonate apex. Bi oP, cugusiiolium. From type Autumnal culms stiffly ascending or some- specimen. what topheavy-reclining, not spreading nor mat-like; blades very numerous, flat, ap- pressed, rather thin and papery, panicles reduced (the later ones often to two or three spikelets), overtopped by the leaves; nen commonly more turgid and blunt than those of the primary panicles. . The flat, papery blades of the autumnal form as seen in the spring still attached to the plants bearing the vernal culm are very characteristic of this species and of the two others of this group with flat autumnal blades (P. consanguineum and P. chrysopsidi- folium). DISTRIBUTION. Sandy pine woods along the Coastal Plain from Pennsylvania to northern Florida and westward to eastern Texas. PENNSYLVANIA: “Bank of Schuylkill below Reading, 1849, Thos. C. Porter” (Acad. Phil. Herb.). DELAWARE: Frankford, Commons in 1875. Viremia: Vicinity of Cape Henry, Chase 5415, Hitchcock 348, Kear- ney 1369, 1416, Williams 3100; Dismal Swamp, Chase 3677. Norra Caro.ina: Roanoke Island, Chase 3249, 3250; vicinity of Wilmington, Chase 3138, 3163, 4585, Hitchcock 14664, 1475; Onslow County, Chase 3169; F1G. 163.—Distribution of P. angustifolium. Chapel Hill, Chase 3063; Ral- eigh, Chase 3082; Caraleigh Junction, Chase 3087. Sout Carona: Fripps Island, Cuthbert 1165; Orangeburg, Hitchcock 349, 1408; Aiken, Ravenel in 1882. 172 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Groraia: Augusta, Kearney 206; Albany, Tracy 3635 in part; Lookout Mountain, Ruth 56; Stone Mountain, Hitchcock 1362; Bullock County, Harper 828; Thomson, Bartlett 1462. Fioripa: Jacksonville, Curtiss 3587 in part, 6803; Baldwin, Hitchcock 984; Talla- hassee, Combs 374, 376; Apalachicola, Biltmore Herb. 4278; Chattahoochee, . Tracy 3636; Gainesville, Chase 4229, 4267; Milton, Chase 4297, 4301; Lake- land, Hitchcock 835. TENNESSEE: Knoxville, Scribner in 1892 (Univ. Tenn. Herb.). Auasama: Etowah County, Eggert 10; Pisgah, Chase 4479; Auburn, Hitchcock 1332, 1340, Tracy 3746, 3750, 3758 in part; Tuskegee, Carver 86; Flomaton, Hitchcock 1039; Mobile, Kearney 28 in part, Langlois 48. MississieP1: Jackson, Hitchcock 1298; vicinity of Biloxi, Hitchcock 1087, 1091, 1105, 1115, Kearney 306 in part, Tracy 1730, 1883, 1888, 4578, 4579, 4615, 4616 in part. Lovuistana: Calhoun, Hitchcock 1261, 1264, 1269; Coushatta, Ball 121; Lake Charles, Hitchcock 1151, Tracy 3651, 3657; Calcasieu River, Langlois in 1884. Texas: Waller, Hitchcock 1193, 1209, 1221; Beaumont, Reverchon 4159; Houston, Bebb 1262, Hall 833; Big Sandy, Reverchon 4193; without locality, Wright (Gray Herb.). 95. Panicum fusiforme Hitchc. Panicum neuranthum ramosum Griseb. Cat. Pl. Cub. 232. 1866, not P. ramosum L. 1767. ‘Cuba occ. (Wr[ight] 3454).’”’ The type, in the Grisebach Herbarium, was col- lected by Charles Wright in western Cuba in 1863, and is numbered ‘900 =3454.” It is the autumnal form. This species was also distributed by Wright under nos. 3453 and 3461 in part.¢ Panicum fusiforme Hitche. Contr. Nat. Herb. 12: 222. 1909. Based on P. neu- ranthum ramosum Griseb., not P. ramosum L. DE SCRIPTION . Vernal form similar to that of P. angustifolium; culms 30 to 70 em. high, the basal and lower blades narrower and at least the lowermost softly pubescent beneath, the longitudinal wrinkles obscure, the leaves more or less clustered toward the base of the culms, the panicles thus long-exserted; spikelets 3.3 to 3.5 mm. long, 1.4 to 1.5 mm. wide, elliptic, long-attenuate at base; first glume two-fifths the length of the spikelet, usually obtuse, second glume and sterile lemma exceeding the fruit and somewhat beaked beyond it at maturity, the pubescence as in P. an- gustifolium; fruit 2.5 mm. long, 1.4 to 1.5 mm.wide, broadly elliptic, obscurely puberulent at the subacute apex. Autumnal culms erect or reclining, under favorable conditions formingdense, 3 bushy clusters 30 to 60 cm. in height; Fig. 164.—P. fusiforme. From type specimen of blades soon involute, 3 to 5 em. long; a i as SS eas spikelets more turgid than those of the primary panicles, 3.5 to 3.8 mm. long, more pointed; fruit more turgid. Panicum fusiforme can be distinguished from P. angustifolium in the vernal state by the larger and more pointed spikelets and the soft pubescence on the lower surface of —S= <.. S ee a See Hitchcock, Contr. Nat. Herb. 12: 222. 1909. al the narrower lower blades; in the autumnal state by the involute blades and dis- tinctly longer spikelets. In the autumnal state this species resembles P. arenicoloides, but has larger spikelets. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 173 DISTRIBUTION. Sandy pine woods, southern Georgia to Florida and Mississippi; also in Cuba. Georat: Albany, Tracy 3614, 3635 in part. Fiorma: Lake City, Combs 136; De Funiak Springs, Combs 456; Monticello, Combs 298; Madison, Combs 231; Pensacola, Combs 516; Gainesville, Chase 4248, Combs 731; Sanford, Hitchcock 786,791; Titusville, Chase 3991, 4021; Eustis, Nash 1226, 1856; Orange Bend, Chase 4105; Lake Harris, Chase 4119; Orange County, Combs 1037; Ormond, Hitchcock 114; Lafay- ette County, Combs 853, 899; Lakeland, Hitchcock 834, 837, 850; Wiwauma, Hitchcock 980; Tampa, Combs 1340, 1343; Seminole, Tracy 7163; Bartow, Fia. 165.—Distribution of P. fusiforme. Combs 1241; Braidentown, Hitchcock 964; Manatee County, Tracy 6708, 6710 in part, 6713 in part, 6713a; Perico Island, Tracy 7371; Myers, Chase 4179, 4194, Hitchcock 877, 899, 912, 923; Miami, Chase 3855, Hitchcock 627. ALABAMA: Flomaton, Hitchcock 1054. MisstssirPi: Ocean Springs, Tracy in 1892. Cupa: Herradura, Caldwell & Baker 7139, Hitchcock 117, Tracy 9074; western Cuba, Wright 3453 in part, 3454 in part; Isle of Pines, Curtiss 406. 96. Panicum arenicoloides Ashe. Panicum arenicoloides Ashe, Journ. Elisha Mitchell Soc. 16: 89.1900. ‘‘Shady sandy woods along the coast of North Carolina. Type material collected by me near Wilmington, N. C., June 6, 1899.’’ The type, in Ashe’s herbarium, consists of three vernal culms with involute blades and mature panicles; the spikelets are 2.4 mm. long. Panicum orthophyllum Ashe, Journ. Elisha Mitchell Soc. 16:90. 1900. ‘‘Shady slopes of sand hills, New Hanover County, N.C., June, 1899.’’ The type, in Ashe’s herbarium, consists of a small tuft of vernal culms beginning to branch, about 60 cm. high, the lower nodes geniculate. The primary panicles are mostly devoid of spikelets, the secondary nearly mature; the spikelets are 2.2 mm. long. DESCRIPTION. Vernal form intermediate in appearance between ide ie 2? drenkeelbides. row that of P. angustifolium and P. aciculare, grayish type specimen. green, slender, mostly 30 to 50 cm. high; lower sheathsand blades softly villous; blades 7 to 12cm, long (the lower shorter), 3 to 4, rarely 5 mm. wide, tapering from ‘the base to a more or less involute apex; panicles 4 to 6 cm. long, two-thirds to three-fourths as wide, the lower branches ascending; spikelets 2.1 to 2.5 mm. long, 1.2 to 1.3 mm. wide, obovate, obtuse; first glume one-third the length of the spikelet, truncate or pointed; second glume and sterile lemma scarcely covering the fruit at maturity, papillose- 174 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. pubescent; fruit 1.8 to 1.9 mm. long, 1.1 to 1.2 wide, obscurely puberulent at the apex. Autumnal form bushy-branching, erect or topheavy, the blades involute; spikelets more turgid, the attenuate base in exceptional specimens elongated, lengthening the spikelet to as much as 2.8 mm. The vernal form of this species can be distinguished from P. aciculare by the larger spikelets and longer blades, from P. angustifolium by the smaller spikelets and the ascending branches of the panicle; the autumnal form is distinguished by the invo- lute blades, longer than those of P. aciculare. The following specimens have spikelets with lengthened bases: Frorrpa: Eustis, Nash 598, 1436; Lake City, Chase 4281; Gainesville, Chase 4211. Musstssrppr: Biloxi, Tracy 3632. An exceptional specimen, with beaked spikelets 2.9 mm. long, Chase 4161, Myers, Florida, is doubtfully referred here. DISTRIBUTION. Sandy pine woods, mostly near the coast, from North Carolina to Florida, Arkansas, and Texas; also in Guatemala. Norra Carona: Near Wilmington, Chase 3120, 3143, 3156, 4581, Hitchcock 350; Raleigh, Chase 30824. SoutH Carouina: St. Helena Island, Cuthbert in 1899; Orangeburg, Hitchcock 352; Isle of Palms, Hitchcock 351. Georei: Millen, Harper 757. Fioria: Duval County, Curtiss 3583* in part, 3587* in part, 4028; Lake City, Chase 4291, Combs 164, Hitchcock 1012; Monticello, Combs 300; Leon County, Curtiss B; Citrus County, Combs 1022; Mary Esther, Tracy 9144; Gaines- ville, Chase 4249; Ormond, Hitchcock 108. Misstssipp1: Biloxi, Chase 4340, Hitchcock 1077, Kearney 215 in part. ArKansas: Fulton, Bush 2522. Lours1ana: Breton Island, Tracy 459, 459a; Lake Charles, Chase 4423; Tangi- pohoa, Cocks 3322. Texas: Houston, Eggert in 1899 (Mo. Bot. Gard. Herb.). GuaTEMALA: Between Guald4n and Copan, Pittier 1805b. Fic. 167.—Distribution of P. arenicoloides. 97. Panicum ovinum Scribn. & Smith. P. redivivum Trin.; Steud. Nom. Bot. ed. 2. 2: 262. 1841. This isa nomen nudum, and appears as P. redivivum ‘Trin. mpt. Mexico.” The type, in the Berlin Herba- rium, was collected by Schiede at Hacienda de la Laguna, Mexico. Panicum ovinum Scribn. & Smith, U. 8. Dept. Agr. Div. Agrost. Cire. 16: 3. 1899. “Type collected by F. W. Thurow, Waller County, Texas, May 29, 1898.’’ The type specimen, in the National Herbarium, the vernal form, is glabrous except the ciliate basal portion of some of the lowermost blades. DESCRIPTION. Vernal form with culms usually few in a cluster, erect or nearly so, glabrous, 30 to 50 cm. high; sheaths glabrous or the lowermost appressed pubescent; blades erect or ascending, stiff, glabrous, the lower somewhat ciliate on the margin at base, the lower- HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 175 most ovate or lanceolate, as much as 1 cm. wide, those of the mid-culm, 10 to 15 cm, long, 3 to 6 mm. wide, the uppermost shorter and narrower; panicles usually short- exserted, 5 to 9 cm. long, three-fourths as wide or less, loosely flowered, the lower branches ascend- ing; spikelets 2.1 to 2.2 mm. long, 1.2 to 1.3 mm. wide, obovate-elliptic, obtuse, basal attenuation short; first glume about one-fourth the length of the spikelet, usually truncate; second glume and sterile lemma scarcely equaling the fruit at maturity, papillose-pubescent, sometimes minutely so; fruit 1.8 mm. long, 1.1 mm. wide, oval, puberulent at the apex. Autumnal form erect or nearly so, the blades becoming loosely involute, not much shorter than the vernal blades; spikelets more turgid, sometimes slightly shorter than those of the primary panicle. Panicum ovinum in its vernal form differs from P. aciculare in being nearly smooth, and in having broader spikelets and larger, less exserted panicles, the upper- most blades being proportionately longer. Fig. 168.—P. ovinum. From type specimen. DISTRIBUTION. Dry or moist open ground, Missis- sippi to Arkansas and eastern Texas; also in Mexico. In Texas this species occurs upon the open prairie, on dry ground, and also in swales. Mississippi: Ocean Springs, Tracy a? part; Biloxi, Hitchcock Fia. 169.—Distribution of P. ovinum. Arkansas: Jefferson County, Zggert in 1898 (Mo. Bot. Gard: Herb.). Lourstana: Shreveport, Hitchcock 1250; Lake Charles, Hitchcock 1131, 1141, 1150. Texas: Dallas, Reverchon 1087; Waller County, Hitchcock 1172, 1192, 1210, 1222, Thurow in 1898 and 1906; Montgomery County, Thurow in 1905; Grand Saline, Reverchon 4137; Hempstead, Hall 834 (Gray Herb.); without locality, Nealley in 1884 and 1887, Reverchon 92 in 1879. Mexico: Hacienda de la Laguna, near Jalapa, Schiede (Berlin Herb.). 98. Panicum neuranthum Griseb. Panicum neuranthum Griseb. Cat. Pl. Cub. 232. 1866. ‘‘Cuba or. (Wr[ight] 3453); oce., in savanis pr. Hanabana (Wr[ight] a. 1865: forma ascendens, ramosa, foliis planis, spiculis ut in @).’’ The type specimen, in the Grisebach Herbarium, was collected by Charles Wright in eastern Cuba in-1860 and is numbered ‘‘103=3453.” This is the autumnal form. Another specimen, the second one cited above, was col- lected in 1865 and is labeled “‘a forma ascendens ramosa.”” This specimen is P. chrysopsidifolium. 4 DESCRIPTION. Vernal form with numerous cespitose, stiff, erect, glabrous culms, 30 to 60 cm. high; sheaths glabrous or ciliate on the margin and usually with a few long hairs at the summit, or the lowermost sparsely ascending-pubescent; blades erect or ascending, @ See Hitchcock, Contr. Nat. Herb. 12:183. 1909, for a discussion of Wright’s Cuba grasses, 176 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. glabrous or with a few ciliz toward the base, the short basal blades few or wanting, those of the middle culm usually 10 to 15 cm. (sometimes only 5 to 6 em.) long, 3 to 5 mm. wide; panicles finally long-exserted, 5 to 9 cm. long, narrow, the flexuous branches narrowly ascending, rarely loosely spreading, the branchlets appressed, the short- pediceled spikelets more or less secund along the branches; spikelets 2 mm. long, 1.2 mm. wide, broadly obovate, blunt, the attenuation at base short; first glume about one-third the length of the spikelet, truncate or pointed; second glume and sterile lemma scarcely equaling the fruit at maturity, finely papillose-pubescent; fruit 1.8 mm. long, 1.1 mm. wide, elliptic, puberulent at the subacute Fic. 170.—P. neuranthum. From apex. : type specimen. Autumnal form with erect, usually slender culms nearly as tall as the vernal form; blades involute but not stiff, not conspicuously shorter than the vernal blades, 4 to 10 em. long; spikelets more turgid, the fruit often slightly exceeding the second glume. Most nearly related to P. ovinwm, from which the vernal form may be distinguished by the narrower, fewer-flowered panicles with subsecund, slightly smaller spikelets, by the usually taller culms, and by the absence of the comparatively broad basal blades which distinguish the latter; the au- tumnal form is much taller, with longer, less crowded blades. From P. aciculare this is distinguished by absence of pubescence, much longer blades, subsecund spikelets, and an erect autumnal form. DISTRIBUTION. Savannas and open ground, southern Florida, and along the coast to Missis- Fig. 171.—Distribution of P. neuranthum. sippi; also in the Bahamas and Cuba. Fiorina: Miami, Hitchcock, 109, 705, 710; Alligator Harbor, Tracy 7176; Sanibel Island, Simpson 298; Clearwater, Tracy 7166; Braidentown, Tracy 6711; without locality, Rugel 290. Mississippi: Petit Bois Island, Tracy 4567; Horn Island, Tracy 2859. Banamas: New Providence, Britton & Brace 599 (Field Mus. Herb.). Cusa: Eastern Cuba, Wright 3453 in part. Bicknelliana.—Culms few to several in a tuft, glabrous or nearly so; sheaths glabrous or with a few hairs; ligules nearly obsolete; blades somewhat elongated, stiffly ascending or spreading, ciliate at base; panicles few-flowered, with 7-nerved, long-peduncled spikelets 2.5 to 3 mm. long. Autumnal form sparingly branching from the upper or middle nodes. This group of two species is intermediate in habit between the Depauperata and Dichotoma; the blades are elongated as in the former but the vernal culms and the mode of branching are more like those of the latter. Spikelets 2.5 to 2.8 mm. long, blades not over 9 mm. wide.... 99. P. bicknellit. Spikelets 3 mm. long, blades as much as 12 mm. wide.....-.-- 100. P. calliphyllum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 177 99. Panicum bicknellii Nash. Panicum bicknellii Nash, Bull. Torrey Club 24: 193. 1897. ‘‘The type specimens were collected by Mr. Eugene P. Bicknell * * * in Bronx Park [N. Y.] on July , 21, 1895.’ The type, in Nash’s herbarium, is the early branching form of the plant. The spikelets are sparsely pubescent. Panicum nemopanthum Ashe, Journ. Elisha Mitchell Soc. 15:42. 1898. ‘‘Type material collected by the writer April, 1895, in the Penitentiary woods, Raleigh, N.C.’’ The type could not be found in Ashe’s herbarium, but a specimen from the type material labeled in Ashe’s handwriting is in the National Herbarium. This isa single vernal culm with an immature, partly included panicle; the spikelets are nearly or quite glabrous. Panicum bushii Nash, Bull. Torrey Club 26: 568.1899. ‘‘Collected by B. F. Bush, in dry ground, in McDonald Co., Missouri, July 24, 1893 no. 413.” The type in the Columbia University Herbarium consists of a small tuft of branching culms, the pri- mary panicles devoid of spikelets; most of the primary nodes sparsely pilose, most of the secondary ones glabrous; the spikelets glabrous. Although the types of P. nemopanthum and of P. bushii have glabrous spikelets, later collections of the species in the Peniten- tiary woods, Raleigh (Ashe & Chase 3092) and from B. F. Bush have pubescent spikelets. These two types are exceptional specimens. DESCRIPTION. Vernal form bluish green; culms erect or as- cending, 30 to 50 cm. high, glabrous, or the low- ermost portion puberulent, nodes sparsely bearded or glabrous; sheaths glabrous or the lower sparsely villous especially above the nodes; blades stiffly ascending, or somewhat spreading, elongated, 8 to 15 cm. long, 3 to 8 mm. wide, the uppermost usually longest, narrowed toward the base, there usually ciliate with a few stiff hairs; panicles ovoid, 5 to8cm. long, about two-thirds as wide, the branches ascending, bearing few long-pediceled spikelets, these 2.3 to 2.8 mm. long, 1.1 to 1.2 mm. wide, oblong-elliptic, sparsely pubescent or rarely glabrous; first glume about one-third the length of the spikelet, subacute; second glume and sterile lemma equal, covering the fruit at maturity; fruit 2 mm. long, 1.1 mm. wide, elliptic, subacute. Autumnal form erect, branching from the middle nodes, forming a loose, bushy crown of stiffly ascending blades not much reduced and overtopping the narrow, few-flowered panicles. The long upper blades in this species are noticeable. Vernal specimens some- times resemble P. werneri. One specimen, Bush 3246, has pilose sheaths and scattered long hairs on the blades. Fic. 172.—P. bicknellii. From type specimen. DISTRIBUTION. Dry, sterile or rocky woods, Connect- icut to Georgia and Missouri. Connecticut: Norwich, Graves 15 in 1899. Fi. 173.—Distribution of P. bicknellii. New York: Bronx Park, Bicknell in 1895; Cedarhurst, Bicknell in 1903; Rockville Center, Bicknell in 1906; Woodmere, Bicknell in 1904; Rockport, Bicknell 1905. 41616°—vo1t 15—10——12 178 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. PENNSYLVANIA: Chambersburg, Porter in 1896, 1897, and 1898; Westchester, Windle in 1904. Onto: Vinton, Kellerman 6887. Missouri: McDonald County, Bush 413; Eagle Rock, Bush 3246. Marytsanp: Plummers Island, Hitchcock 118, Steele in 1897; Great Falls, Chase 3783; West Chevy Chase, Chase 24774, 5416. District or CotumBtiA: Hitchcock 353. Norra Carona: Raleigh, Ashe in 1895, Ashe & Chase 3092. Georaia: Stone Mountain, Hitchcock 201. 100. Panicum calliphyllum Ashe. Panicum calliphyilum Ashe, Journ. Elisha Mitchell Soc. 15: 31. 1898. ‘“‘Type ma- terial collected by the writer at Watkins, Lake Seneca, N. Y., Aug. 1898.” The type could not be found in Ashe’s herbarium. In the National Herbarium is a specimen collected by Ashe ‘‘Near Ithaca, N. Y.,’’ which Mr. Ashe has stated orally is from the published locality and is a duplicate type. This specimen consists of two vernal culms, lack- ing basal leaves. DESCRIPTION. Vernal form yellowish green; culms erect, 35 to 50 cm. high, glabrous, the nodes sparsely vil- lous; sheaths glabrous, or the lowermost sparingly pubescent, ciliate on the margin; blades ascend- ing, flat, 8 to 12 cm. long, 9 to 12 mm. wide, gla- brous, ciliate at the rounded base; panicles few- Fig. 174.—P. calliphyllum. From flowered, 7 to 9 cm. long, half as wide or less, with type specimen. 0 . E a few ascending branches; spikelets mostly long- pediceled, 2.9 to 3 mm. long, 1.2 mm. wide, elliptic, sparsely pubescent; first glume about oné-third the length of the spikelet; second glume and sterile lemma equaling the fruit at maturity; fruit 2.3 mm. long, 1.1 mm. wide, elliptic, subacute. Autumnal form sparingly branching from the middle nodes, the branches about as long as the internodes, erect; blades not greatly reduced; panicles narrow, partly included. This little known species has been referred @ to P. xanthophysum, to small specimens of which the above-mentioned dupli- cate type bears some superficial re- semblance. The species is closely related to P. bicknellii. But three specimens have been seen, on none of which is the habitat given. DISTRIBUTION. Massachusetts, New York, and Ohio. Massacuusetts: Medford, Perkins in 1881 (N. E. Bot. Club Herb.). New York: Near Ithaca, Ashe in 1898. Onto: Painesville, Hacker in 1901. Fia. 175.—Distribution of P. calliphyllum. @ Merrill, Bull. Torrey Club 27: 595. 1900. Nudicaulia.—The following species does not seem to be closely allied with any other, and hence is placed tentatively in a group by itself. So far as the technical characters are concerned it might be placed in the group Dichotoma, but it differs from any of the species of that group in the narrow, enveloping base of the blades, and the nearly naked culms. When the autumnal form is known the affinity of the species may be shown. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 179 101. Panicum nudicaule Vasey. Panicum nudicaule Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8: 31. 1889. ‘‘Swamps, Santa Rosa County, Fla. (A. H. Curtiss.)” The type, in the National Herbarium, is the vernal form. It was collected in ‘‘Swamps, Santa Rosa County, N. W. Florida,’’ in May [1886] by A. H. Curtiss (no. 3583*). DESCRIPTION. Vernal culms erect from a somewhat spreading base, 40 to 60 cm. high, glabrous, the lower internodes short, the two upper much elongated, thus producing the effect of a long, nearly leafless stem; nodes gla- brous, only the upper two visible; sheaths glabrous or the overlapping basal ones sometimes sparsely pubescent; ligules ciliate, less than 0.5 mm. long; blades erect, rather thick, 4 to 10 cm. or some of the lower as much as 13 cm. long, 5 to 8 mm. wide (the uppermost more or less reduced), nearly linear, abruptly pointed, slightly narrowed and somewhat enveloping the culm at base, glabrous; panicles long-exserted, 4 to 7 cm. long, hardly as wide, few- flowered, the branches ascending; spikelets 2.7 to 2.9 mm. long, 1 to 1.2 mm. wide, narrowly ovate, acuminate, glabrous; first glume one-fourth to one-third as long as the spikelet, usually pointed; second glume and sterile lemma 7- nerved, exceeding the fruitand pointed beyond it; fruit 2 mm. long, 1 mm. wide, elliptic, acute. Autumnal form unknown. Fig. 176.—P. nudicaule. From type specimen. DISTRIBUTION. Swamps, western Florida and south- ern Alabama. Fia. 177.—Distribution of P. nudicaule. . Fiorina: Blackwater River Swamp, Santa Rosa County, Curtiss, B, 3583*. ALABAMA: Gateswood, Tracy 8431, 8432. Dichotoma.—Culms few to many in a tuft, glabrous, or the nodes only pubescent; sheaths glabrous, or the lower sometimes pubescent, never conspicuously hirsute; ligules ciliate, 0.7 mm. long or less; blades lanceolate, rarely as much as 1 cm. wide, mostly glabrous; panicles usually open; spikelets elliptical, not turgid (except in P. roanokense and P. caerulescens), 1.5 to 2.5 mm. long; second glume and sterile lemma 5 to 7-nerved. Autumnal form usually freely branching, erect, reclining, or prostrate, secondary leaves and panicles much reduced. 180 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Nodes, at least the lower, bearded. Spikelets 1.5 to 1.6 mm. long, glabrous (occasional in- dividuals with pubescent spikelets)........-...---- 102. Spikelets 2 mm. or more long. Spikelets glabrous, 2 mm. long; autumnal form top- Heavy ysTeCMMMne es Shak t set ER ee 110. Spikelets pubescent. Blades all velvety; autumnal form sparingly branche disease ee oe Race ee 105. Blades glabrous, or only the lower pubescent or velvety; autumnal form freely branching. Spikelets 2 mm. long; autumnal form pro- fusely branching. Fruits slightly exposed at maturity; upper sheaths viscid-spotted; autumnal form erect or reclining. .103. Fruits covered at maturity; sheaths not viscid - spotted; autumnal form decumbent, with flabellate - fascicled branches; Mexican..... 104. Spikelets 2.2 mm. or more long; autumnal form less profusely branching. Sheaths and upper nodes glabrous. -. . . - 107. Lower sheaths and all nodes pubescent. . 106. Nodes not bearded. Spikelets pubescent. Culms erect, never becoming vine-like. Primary blades spreading; Ager oes fruit exposed at summit. . Be Oe Primary blades erect; panicles green; fruit, cov- ered (wood ee with ae blades may be distinguished from P. dichotomum by pubescent spikelets, 2.2 mm. long). - - -108. Culms soon prostrate, vine-like; branches divaricate. Plants bright green, culms lax; spikelets not over 2a tam’, longit: 02th <4 sock eee 114. Plants grayish green, culms stiff; spikelets 2.5 THE ORES 2. wince kisiae ec ciee ta aoe ee eee 115. Spikelets glabrous. Culms soon prostrate. Plants bright green, culms lax; spikelets not Ovens lem. Long Sseeees Ae Plants grayish green, Crime stiff; Meeps: 2.5 vada 0p | V0} 0 6. Agen teaches Pep ee 8 ay tN ad 115. ‘Culms erect, or the autumnal form topheavy, never prostrate. Spikelets not over 1.6 mm. long; panicles nar- row; plants glaucous bluish green......... 113. Spikelets 2 mm. or more long; panicles open. Blades erect, firm; spikelets turgid, strongly nerved; plants grayish olive green....112. Blades spreading; spikelets not turgid. P.microcarpon. P. barbulatum. P.annulum. P. nitidum. P.multirameum. P. clutet. P. mattamuskeet- ense. P. clutei. P. boreale. ” P. lucidum. P. sphagnicola. P. lucidum. P. sphagnicola. P. caerulescens. P. roanokense. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 181] Spikelets 2.2 mm. or more long, pointed; sheaths bearing pale glandular Dp Oe ee eee ee lll. P. yadkinense. Spikelets not over 2 mm. long, not pointed. Autumnal form erect, branched like a little tree; primary blades rarely over 5mm. wide; second glume shorter than fruit and sterile lemma...... 109. P. dichotomum. Autumnal form topheavy-reclining; primary blades 6 to 10 mm. wide; second glume equaling fruit and sterile lemma.....- 110. P. barbulatum. 102. Panicum microcarpon Muhl. Panicum microcarpon Muhl.; Ell. Bot. S. C. & Ga. 1: 127.1816. No locality is cited. The type, in the Elliott Herbarium, consists of a single culm, lacking the base, with five leaves and an immature panicle, slightly included at the base. The accompanying label reads: ‘“‘Panicum microcarpon. barbulat: var.? Hab. Georg: Dr. Baldwin.’’ As shown by his description and the above-mentioned type, Elliott misunderstood Muhlenberg’s application of this name and attached it to a different species, though giving Muhlenberg as author. Muhlenberg’s later publication of the name @ for a different species ® is thus invalidated. Panicum nitidum ramulosum Torr. Fl. North. & Mid. U. 8. 146. 1824. ‘‘Near Quaker-Bridge, New-Jersey.’’ The type, in the Torrey Herbarium, is a sparingly branched, vernal culm, with nearly mature terminal panicle. The label reads: “Panicum nitidum f var. [9 is ramulosum] collected in swamps in the pine barrens of New Jersey, near Quaker Bridge, June, 1818.” ?Panicum nitidum barbatum Torr. Fl. North. & Mid. U. 8S. 146. 1824. “‘In woods and meadows.’’ The type could not be found in the Torrey Herbarium. The description seems to apply to the vernal form of P. microcarpon, or possibly to P. barbulatum. This species is the one described by Muhlenberg ¢ under the name of P. discolor Spreng., as shown by a specimen so labeled in the Muhlenberg Herbarium. Panicum heterophyllum Schreb. is here cited as a synonym, erroneously, as Schreber did not publish this name. LP. heterophyllum Muhl.¢ is a nomen nudum. Panicum microcarpon Muhl. is the species described as P. barbulatum by American authors, which proves to be not P. barbulatum Michx. DESCRIPTION. Vernal form cespitose; culms erect, or sometimes geniculate at base, 30 or usually 60 to 100 cm. high, the nodes densely bearded with reflexed hairs; sheaths glabrous, or the lowermost pubescent, often mottled with white spots between the nerves, ciliate on the margin; blades thin, spreading, the upper often reflexed, 10 to 12 cm. long, 8 to 15 mm. wide, narrowed toward the base, glabrous, more or less papillose- ciliate at base; panicles finally long-exserted, many-flowered, ovate in outline, 8 to 12 cm. long, the branches ascending; spikelets 1.6 mm. long, 0.7 mm. wide, elliptic, a Descr. Gram. 111. 1817. b See P. microcarpon Muhl. under P. polyanthes, page 255. ¢ Descr. Gram. 114. 1817. d Trans. Amer. Phil. Soc. 3:160. 1793. 182 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. glabrous, or rarely minutely pubescent; first glume one-fourth the length of the spikelet or less; second glume a little shorter than the sterile lemma and slightly exposing the fruit at maturity; fruit 1.3 to 1.4 mm. long, 0.7 mm. wide, elliptic, slightly pointed. Autumnal form much branched from all the nodes, reclining from the weight of the dense mass of branches; blades reduced, flat, mostly 2 to 4cm.long; the ciliz of the sheaths and the hairs at the base of the blades relatively more conspicuous; panicles much reduced and loosely flowered; the tufted basal blades often large, sometimes as much as 8 cm. long and 15 mm. wide. The following specimens have pubescent spikelets, but differ in no other respect from the typical form: Massa- Fig. 178.—P. microcarpon. causnrts: Bartlett 844; Connecticut: Bissell 5570; New From type specimen in 2 Biliott Herbarium. JERSEY: Heritage 4; PENNSYLVANIA: Smith 102; Ixurnots: Schneck in 1880; DetawaReE: Canby 15; District or Cotum- BIA: Pollard 408, Vasey in 1884; Norra Carouina: Biltmore Herb. 803a, Chase 3204; GeoreiA: Ruth 57; Fioriwa: Curtiss, P.Q.; ALABAMA: Carver 40, Hitchcock 13438; MississiPPi: Tracy 1733, 3388,¢ 3623; ARKANSAS: Heller 4237. The last specimen is also exceptional in being pubescent throughout. Curtiss’s no. 6648, McClenny, Florida, with glabrous spikelets as much as 1.8 mm. long, seems to be intermediate between P. microcarpon and P. nitidum. DISTRIBUTION. Wet woods and swampy places, Massachusetts to Illinois, and south to northern Florida and eastern Texas. MassacuuseEtts: Milton, Bartlett 844. Connecticut: New Haven, Bissell 5566; Groton, Bissell 5569; Stratford, Bissell 5571; Milford, Bissell 5570. RHODE Istanp: Providence, Collins in 1891 (Gray Herb.). New York: Eastern New York, Austin (Mo. Bot. Gard. Herb.). New Jersey: South Amboy, Watherdie 1484, 2161; Morris Plains, Mackenzie 1605; Haworth, Mackenzie 2476; Wildwood, Chase 3516, Heri- tage 4, 4’; Clifton, Nash in 1892. PENNSYLVANIA: Lancaster County, Heller 4770 in part, 4787, Small in 1889; Philadelphia, Smith 102. Onto: Albion, Ashcroft in 1897; Jackson, Kellerman 6778. InprANA: Brazil, Somes 232. .Inuvors: Peoria, Brendel; Mount Fia. 179.—Distribution of P. microcarpon. Carmel, Schneck in 1880. Missourr: Butler County, Eggert 236; Campbell, Bush 747 in part; Dunklin County, Eggert 287. DetawareE: Milton, Commons 346, 347; Centerville, Commons 285; “Townsend, Canby 15. MARYLAND: Chesapeake Beach, Chase 3253, 3262; Hitchcock 1606, 1611; Belts- ville, Chase 3767, 3831; Riverdale, Chase 2367, 2370. District or CotumsiA: Ball 704, Chase 5419, in Kneucker Gram. Exs. 550, Hitchcock 370, 596, Merrill 197, Pollard 362, 406, 408, Vasey in 1884. a Tracy’s no. 3388 was erroneously cited under Panicum wibbertulaia Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Cire. 29: 9. 1901. HITCHCOCK AND GHASE—NORTH AMERICAN PANICUM. 1883 Virernia: Vicinity of Norfolk, Kearney 1307, Mackenzie 1686, Noyes 93; Dismal ‘Swamp, Chase 3668; Smyth County, Small in 1892. West Virarnia: Baileysville, Morris 1193; Peeryville, Morris 1139. Norrs Carouina: Hyde County, Chase 3204, 32103; Onslow County, Chase 3187, 3190; Chapel Hill, Chase 30613; Rowan County, Small in 1894; Biltmore, Biltmore Herb. 803a. Sourn Carona: Orangeburg, Hitchcock 1393, 1411, 1419. Groraia: Clarke County, Harper 74; Stone Mountain, Eggert 89; Thomson, Bart- lett 1081, 1457; Whitfield County, Harper 281, Wilson 125, 137; Gwinnett County, Small in 1893; Lookout Mountain, Ruth 57, 64; Augusta, Cuthbert 1161; Warm Springs, Tracy 8865; Leslie, Harper 1105; Rabun County, Small in 1893. Froriwa: Tallahassee, Combs 391, Kearney 88, Nash 2522; Lake City, Combs 173; Madison, Combs 255, 256; Orange County, Combs 1140. Kentucky: Harlan County, Kearney 50. TENNESSEE: Polk County, Kearney 326 in part; Cocke County, Kearney 967; Carroll County, Eggert 78. AxaBaAMA: Mobile, Mohr in 1892; Jackson County, Chase 4482; Tuskegee, Carver 16, 40; Cullman County, Eggert 15; Auburn, Hitchcock 1343. Mississippi: Biloxi, Tracy 1733, 3623, 3624, 3761, 4609 in part, 7018; Wahalak, Tracy 3224; Coopolis, Tracy 4595; Fairport, Tracy 3207; Centerville, Tracy 3618; Macon, Tracy 3225; Saratoga, Tracy 8417; Starkville, Phares 3623; Morrisonville, Tracey 3388. ArKAnsAs: Little Rock, Coville in 1887; northwest Arkansas, Harvey 4, Tex- arkana, Heller 4237. Lourstana: Plaquemines Parish, Langlois 40a, 40b; Lake Charles, Chase 4426, Hitchcock 1149; Calhqun, Baill 51. Texas: Big Sandy, Reverchon 4194; Texarkana, Heller 4088; Rockland, Nealley 36; Waller County, Thurow 18. 1038. Panicum nitidum Lam. Panicum nitidum Lam. Tabl. Encycl. 1: 172.1791. ‘‘E Carolina. com. D. fraser.”’ The type,? in the Lamarck Herbarium, consists of a panicle and the uppermost joint of the culm with its leaf, the blade reflexed, the node sparsely clothed with reflexed hairs. This specimen does not belong to any of the species to which the name P. nitidum has been applied by American authors. Panicum nodiflorum Lam. Encycl. 4: 744. 1798. Lamarck states as to the origin of his specimen, ‘‘J’en posséde un exemplaire recueilli par M. Fraser dans la Caroline. Le citoyen Michaux |’a trouvée dans la Basse-Caroline.’’ The type, in the Lamarck Herbarium, is a fragment of an autumnal culm with two nodes, at each of which is a fascicle of branchlets with reduced leaves but devoid of spikelets. It is from ““Caroline.”’ Panicum dichotomum nitidum Wodd, Class-book ed. 3. 786. 1861. This is presum- ably based on P. nitidum Lam., though no synonym nor locality is cited. The short description seems to apply best to P. lindheimeri Nash. Panicum subbarbulatum Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Circ. 29: 9.1901. Based on ‘‘( Panicum barbulatum of Ell. Sk. Bot. 8S. C. and Ga. 1: 127. 1817, not Michx. Fl. Bor. Am. 1: 49. 1803.)’’ ‘‘The type of the species in the herbarium of Elliott” consists of three vernal culms lacking the base. @ For a full discussion see Hitchcock, Contr. Nat. Herb. 12: 148. 1908. 184 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Vernal form cespitose; culms erect or somewhat spreading at base, rather stout, usually 30 to 60 cm. high, or sometimes as much as | meter high, the nodes bearded with reflexed hairs; sheaths glabrous, or the lower pubescent, ciliate on the margin, more or less mottled or glandular, especially the upper at anthesis; blades firm, glabrous, sometimes sparsely ciliate at the base, 5 to 12 cm. long, 5 to 10 mm. wide, the lower ascending, the upper usually reflexed; panicles ovoid, 5 to 8 cm. long, nearly as wide, rather densely flowered, the axis and ascending branches viscid-spotted; spikelets elliptic, 2 mm. long, 1 mm. wide; first glume less than one-third the length of the spikelet, pointed; second glume and sterile lemma subequal, the slightly shorter glume scarcely covering the fruit at maturity; fruit elliptic 1.7 mm. long, 1 mm. wide, subobtuse. ee Autumnal form erect or more or less reclining from a ie ans Brae bili weight of the foliage, the branchlets and foliage forming large clusters from the nodes of the vernal culm; reduced blades numerous, | to 3 cm. long, 1 to 3 mm. wide, flat or soon becoming involute; panicles mostly reduced to a few long-pediceled spikelets. This species has been confused with Panicum microcarpon, from which it is distin- guished by the pubescent spikelets, 2 mm. long, and by the erect autumnal form with involute blades. The viscid spots on the sheaths are often conspicuous. Two specimens, Hitchcock 1420 and 1421, while showing all the other characters of this species have glabrous spikelets, but they are fully 2 mm. long. DISTRIBUTION. Low moist or marshy ground from Virginia to Florida and along the coast to Texas; also in the Bahamas. Missouri: Carter County, Eggert 288. Virainia: Virginia Beach, Hitchcock 119. Nort Carona: Scranton, Chase 3200. Sours Carona: Orangeburg, Hitchcock 4, 15,1376, 1389, 1392, 1420, 1421, 1422, 1423. GeorGia: Thomson, Bartlett 1173, 1456. Froripa: Milton, Chase 4307, 4321; Madison County, Combs 266; Citrus County, Combs 979, 1008; Lake County, Chase 4091, Hitchcock 814, Nash 376; Sanford, Chase 4132, Hitchcock 774; Titusville, Chase 3990, 4005, 4017; Myers, Hitchcock 898, 908, Lee Co. Pl. 478, 479; Miami, Hitchcock 718, Tracy 8850; Homestead, Hitchcock 687; Captiva Island, Tracy 7199; Sumter County, Curtiss 3600A in part@; Duval County, Curtiss 3600A in part; Sanibel Island, Tracy 7170; Braiden- town, Hitchcock 961. Mississippi: Biloxi, Hitchcock 1081, Tracy 2031, 4591, 4609 in part; Macon, Tracy 3228. Fig, 181.—Distribution of P. nitidum. a The specimen of this number in the Gray Herbarium consists of P. nitidum and P. equilaterale. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 185 Loutstana: Lake Charles, Hitchcock 1128, 1143, 1148, 1153, 1154. Texas: Hempstead, Hall 834 in part. Banamas: Great Bahama, Britton & Millspaugh 2518, 2736, Brace 3697 (all in Field Mus. Herb.). 104. Panicum multirameum Scribn. Panicum multirameum Scribn. U. S. Dept. Agr. Div. Agrost. Cire. 19: 2. 1900. ‘Gravelly hills near Jalapa, State of Vera Cruz, altitude 1,250 m. (4000 feet). C.G. Pringle 7882, 1889. Orizaba, State of Vera Cruz, February 17, Jared G. Smith, No. 593, 1892.’’ The type, Pringle 7882, in Hitchcock’s herbarium, is the autumnal form with decumbent culms and numerous ascending branches, with a few vernal culms still attached. DESCRIPTION. Vernal culms erect or soon decumbent, 30 to 60 cm. high, nodes villous-bearded; sheaths strongly ciliate on the margin, especially at the summit, the upper glabrous, the lowermost more or less villous; blades rather thick, 3 to 6 cm. long, 3 to 6 mm. wide, glabrous, or rarely puberulent, somewhat ciliate around the base, obscurely white-margined; panicles ovoid, 3 to 6 cm. long, about three-fourths as wide, the branches ascending; spikelets 2 mm. long, 1 mm. wide, elliptic, finely pubescent; first glume one-third the length of the spikelet; second glume and sterile lemma equal and just covering the fruit at maturity; fruit 1.7 mm. long, 0.9 mm. wide, rather abruptly subacute. Autumnal form decumbent, rooting at the lower nodes, flabellately branching before the maturity of the primary panicle, the branchlets in close, appressed clusters at the ends of the secondary branches; blades reduced, flat or somewhat rolled, appressed; panicles reduced to a few spreading branches or long-pediceled spikelets. None of the specimens examined shows a basal rosette of leaves. In technical characters this species is allied to P. nitidwm but differs in the decumbent, flabel- lately-branched autumnal form. Fic.182.—P.multirameum. From type specimen. DISTRIBUTION. Dry hills and gravelly banks, southern Mexico. Mexico: Jalapa, Pringle 7882, 8339,¢ 9209, 9210; Orizaba, J. G. Smith 593, Bourgeau 2383 in part (all in Hitchcock’s herbarium except Pringle 8339); Botteri 703 in part (Gray Herb.). GUATEMALA: Coban, Tuerckheim 56 in part (Gray Herb.). 105. Panicum annulum Ashe. Panicum annulum Ashe, Journ. Elisha Mitchell Soc. 15: 58.1898. ‘‘Dry rocky woods. Maryland to North Carolina and Georgia. Near Washington, D. C. Ward 1892,’’ is the first specimen cited, and is taken as the type. This is in the National Herbarium and consists of three vernal culms with mature panicles. The year of collection is 1882 instead of 1892. Panicum bogueanum Ashe, Journ. Elisha Mitchell Soc. 16: 85. 1900. Based on “P.annulum Ashe, not P. annulatum A. Rich.” 1851. aThree species were distributed under this number: National Herbarium no. 381990 is P. multirameum; P. sphaerocarpon is mixed with this species in the specimen of this number in Hitchcock’s herbarium; National Herbarium no. 823271 is P. olivaceum. 186 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Vernal form usually purplish, in small clumps or solitary; culms 35 to 60 cm. high, the nodes densely bearded; sheaths shorter than the internodes, velvety-pubescent or the upper nearly glabrous; blades 6 to 12 cm. long, 7 to 13 mm. wide, densely velvety-pubescent on both surfaces, the margin ciliate toward the base; panicles 6 to 8 cm. long, about three-fourths as wide, rather numerously flowered, the flexuous branches ascending or later spreading; spikelets 2 mm. long, 0.9 mm. wide, elliptic, blunt; first glume one-fourth to one-third the length of the spikelet, obtuse; second glume and sterile lemma pubescent, the glume slighty shorter; fruit 1.8 to 1.9 mm. long, 0.9 mm. wide, elliptic. Autumnal form erect, bearing in late summer a few short, erect branches at the upper nodes; soon dying to the ground. Fig. 183.—P. annulum. From type specimen. DISTRIBUTION. Dry woods, New Jersey to Georgia, and west to Mississippi; also in Missouri; rare. New Jersey: Milburn, Mackenzie 2138. PENNSYLVANIA: Chester County, Pennell 999, Windle 7; Delaware County, Pen- nell 621, 642, 727, 1184. Missouri: Hunter, Lggert in 1893 (Mo. Bot. Gard. Herb.). Maryuanp: West Chevy Chase, Chase 2947, 3809, 5420. District or CotumBia: Woodley Park, Ward in 1882. Virainia: Great Falls, Chase 3708. Norra Carona: Chapel Hill, Ashe in 1898. Georeaia: Augusta, Cuthberé 431. AuaBama: Mobile, Mohr in 1897; Auburn, Earle & Baker 1544 (Ala. Polyt. Inst. Herb.). Mississippi: Starkville, Tracy in 1888. Fig. 184.—Distribution of P. annulum. 106. Panicum mattamuskeetense Ashe. Panicum mattamuske[eltense Ashe, Journ. Elisha Mitchell Soc. 15:45. 1898. ‘Roadsides, ditch banks, and wet open woods around Lake Mattamuskeet, N.C. * * * Collected by the writer, and Mr. Gilbert Pearson in June, 1898.’’ The type could not be found in Ashe’s herbarium. In the National Herbarium is a specimen labeled ‘‘Panicum Mattamusketense Ashe, Lake Mattamuskeet” in Ashe’s hand- writing, collected ‘‘June 10—July 6, 1898,’’ by “‘W. W. Ashe,”’ evidently a duplicate type. This is a single vernal culm nearly 80 cm. high, with a mature panicle, and agrees in all respects with the description except that the spikelets are described as glabrous, while those of the specimen are pubescent. The two lower sheaths and lowest blade are velvety pilose; the spikelets are 2.3 mm. long. A second duplicate type in Biltmore Herbarium is a better and more characteristic specimen. Panicum flecuosum Muhl.; Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Cire. 27: 3.1900, not Retz. 1791. This herbarium name of Muhlenberg is listed and _ long, the difference in length being HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 187 referred to P. mattamuskeetense Ashe. The species was described by Muhlenberg @ immediately etter Panicum discolor Spreng., under the heading, ‘‘Co-species vel varietas major.’’ The type, in the Muhlenberg Herbarium, is somewhat fragmentary, consisting of a single culm broken into three pieces, but is evidently the same as P. mattamuskeetense. DESCRIPTION. Vernal form in clumps of few to several culms, olivaceous, usually strongly tinged with deep purple; culms stout, erect or subgeniculate at base, the nodes bearded or the upper puberulent only; sheaths less than half the length of the internodes, pilose on the margin, a puberulent ring at the summit, the auricles pilose, the lower sheaths velvety-pilose, the upper glabrous; ligule dense, 0.7 mm. long; blades horizontally spreading, 8 to 12 cm. long, 8 to 12 mm. wide (basal blades much shorter), acuminate, narrowed toward the base, the lower velvety, the upper glabrous, on both surfaces; panicles long-exserted at maturity, 8 to 10 mm., rarely 12 mm. long, about three- fourths as wide, many-flowered, the flexuous branches spreading, short spikelet-bearing branch- lets in the axils; spikelets 2.3 to2.5 mm. long (rarely 2.7 mm. long), 11 mm. wide, elliptic, pointed be- fore maturity, pubescent; first glume about one- third the length of the spikelet, subacute; second glume and sterile lemma subequal, barely cover- ing the fruit at maturity; fruit 2mm. long, 1 mm. wide, elliptic. =: iri os raamgpnge pron Autumnal form erect or becoming somewhat iehahens. decumbent, branching rather sparingly from the middle nodes after the maturity of the primary panicles, the branches iaitics appressed, the reduced crowded blades ascending. The spikelets at maturity are more turgid, shorter, and more obtuse than when immature, the swelling of the ripened fruit siiatease the spikelet in length. In Chase 3744 the spikelets are 2.7 mm. long, while in Chase 3791 from the same place three weeks later the mature spikelets are but 2.4 mm. long, and in Chase 3793, of the same date as the latter, they are but 2.1 mm. long; the fruits, however, are of the same size, 2 mm. due to a varying length of the second glume and sterile lemma. DISTRIBUTION. Low moist ground, New York to Fia. 186,—Distribution of P. mattamuskeetense. North Carolina. New York: Hempstead, Bicknell in 1903; Woodmere, Bicknell in 1902; Hewletts, Bicknell in 1905; Far Rockaway, Bia in 1902. New Jersey: Anglesea, Brown in 1897 (Phila. Acad. Herb.). MARYLAND: Beltsville, Chase 3744, 37444, 3791, 3793, 3826, 3829; Vienna, Novik 182. Norts Carouina: Lake Mattamuskeet, Ashe in 1898; Wilsons Mills, Chase 3099; Wilmington, Hitchcock 354, 1455; Roanoke Island, Chase 3232. 4 Descr. Gram. 115. 1817. 188 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 107. Panicum clutei Nash. Panicum clutei Nash, Bull. Torrey Club 26: 569. 1899. ‘‘Pine-barrens of southern New Jersey. Collected by Mr. W. N. Clute * * * ona trip from Tuckerton to Atsion, July 3-6, 1899.’’ The type, in Nash’s herbarium, consists of three stout culms with mature panicles. ‘The lowermost nodes are sparsely bearded, the upper glabrous, the lowermost sheaths sparsely soft-pilose as in the type of P. mattamus- keetense. The blades are puberulent or pilose on the auricles and the dense ligule is 0.5 mm. long. DESCRIPTION. Vernal form similar in color, size, and habit to P. mattamuskeetense, but more nearly glabrous, only the lowermost nodes, sheaths, and blades velvety, the puberulent ring at the summit of the sheath less dense or wanting; paniclessimilar, on the average smaller; spikelets 2.2 to 2.3 mm. long, 1 mm. wide, indistinguishable from the smaller spikelets of P. mattamuskeetense; fruit the same size and shape. Autumnal form stiffly erect, branching from the lower and middle nodes before the maturity of the primary panicles, the earlier branches nearly equal- ing the primary culms, the later branches shorter, somewhat crowded, the reduced blades spreading. ete | ie eae This form is but doubtfully distinguished from P. specimen. matiamuskeetense. The division is here made on the nearly glabrous character of P. clutei, and spikelets not over 2.3 mm. long, but a few of the specimens are about as referable to one species as to the other. One specimen, Chase 3590, has the smaller panicles and more numerous branches of P. clutei, but the two or three lower sheaths and blades are velvety, while Chase 3793 has Salclets but 2.1 mn. long and upper nodes puberulent only, but is densely velvety below. The latter Specimen is referred to P. mattamuskeetense because its vernal form (Chase 37444) is more like the type of P. matta- muskeetense than like that of P. clutet. One specimen, Chase 3242, has glabrous spikelets. DISTRIBUTION. Low moist ground and cranberry bogs, Massachusetts to North Carolina. Massacuusetts: Framingham, Smith 732; West Falmouth, Churchill in 1894 (Hitchcock Fig. 188,—Distribution of P. clutei. Herb.). New York: Riverhead, Young in 1874 (Field Mus. Herb.). New Jersey: Burlington County, Clute in 1899; Toms River, Bicknell in 1900; Manchester, Chickering in 1877; Atsion, Chase 3545, 3553; Forked River, Chase 3590; Tuckerton, Chase 3598. DeLawaReE: Ellendale, Commons 345. Disrrict or CotumBia: Kenilworth Swamp, Steele in 1899. Virernta: Bedford County, Curtiss in 1871 (Gray Herb.). Nort Carona: Roanoke Island, Chase 3242. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 189 108. Panicum boreale Nash. Panicum boreale Nash, Bull. Torrey Club 22: 421. 1895. ‘‘Moist soil, New- foundland and Ontario to New York, Vermont and Minnesota. This plant was first noted by the writer in 1893 in the Catskill Mts., near Cairo, N. Y.’’ The type in Nash’s herbarium consists of several vernal culms with nearly mature panicles; the lower sheaths sparsely papillose-pubescent at least toward the summit, the lower and middle nodes bear a few reflexed hairs. The label reads: ‘‘Panicum boreale Nash., n.sp. Moist soil, Cairo, Greene Co., N. Y. Alt. 1400 ft. Coll: Geo. V. Nash, June 28, 1893.”’ DESCRIPTION. Vernal form cespitose, the culms erect, or in shaded places sometimes decumbent at base, usually 30 to 50 cm. high, the nodes glabrous or sometimes with a few hairs; sheaths often overlapping, ciliate on the margin, glabrous, or the lower sparsely pubescent; blades erect, or in laxer forms spreading, sparsely ciliate at the rounded base, otherwise glabrous, 6 to 12 cm. long, 7 to 12 mm. wide; panicles loosely rather few- flowered, 5 to 10 cm. long, hardly as wide, the branches ascending or spreading; spikelets 2 to 2.2 mm. long, 1 mm. wide, elliptic, subacute, pubescent; first glume one-third as long as the spikelet or less; second glume and sterile lemma subequal, the glume scarcely as long as the fruit at maturity; fruit 1.9 mm. long, 1 mm. wide, elliptic, subacute. Autumnal form erect, sparingly branched from all the nodes in late summer, the branches erect, the leaves and panicles not greatly reduced. A weak form with geniculate base and lax spread- ing blades occurs in Maine, and is represented by Fernald 512 and 516, and Chase 3355. One specimen, Chase 3437, is unusual in having blades that are puberulent beneath. Two specimens from northern Indiana, Bebb 2030 and 2831, and a specimen from Wisconsin, Cheney 2100, have stouter culms and more compact and more numerously flowered panicles. A specimen from De- troit, Mich., Farwell 1425, is referred to this species doubtfully because of the pubescence on the back of the joint between the blade and the sheath and because of the sparse papillose-pubescence on the under surface of the blades and on some of the sheaths. It resembles P. mattamuskeetense in habit, but the nodes are glabrous and it is far out of the range of that species. A specimen from Rockville Center, Long Island, Bicknell in 1903, is doubtfully referred here. Fic. 189.—P. boreale. From type specimen. DISTRIBUTION. Moist open ground or woods, Newfoundland to New Jersey and west to northern Indiana and Minnesota. NEWFOUNDLAND: Exploits River, Robinson & Schrenk 222. Nova Scorta: Windsor, Macown 29349; without locality, Burgess 16. New Brunswick: Fredericton, Fowler in 1892; Miramichi, Fowler in 1892. QueEBEc: Montmorenci Falls, Macown 69204 (Gray Herb.). Marne: Dover, Fernald 239, 514; Orono, Fernald 513, 516, 517; Holden, Knight 60, 61; North Berwick, Parlin 1029, 1187; Brownsville, Parlin 1701, 1738, 1744; Canton, Parlin 2013, 2034; St. Francis, Fernald 505, Knight 58; Farm, ington, Fernald 512; Hartford, Parlin 1423; Cumberland, Chamberlain 336- 793; Cape Elizabeth, Chase 3458; Chesterville, Chase 3277, 3355, 3437, 3443; Mount Desert Island, Redfield in 1893. 190 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. New Hawmesuire: Jaffrey, Hitchcock 120, Robinson 338, 338a; White Mountains, Hitchcock 121. Vermont: Brattleboro, Jones 33. Massacuusetts: Framingham, Smith 733; South Hadley, Cook in 1887. Connecticut: Southington, Bis- sell 5582; Griswold, Graves 77; Waterford, Graves 82. New York: Cairo, Nash in 1893; Fulton Chain, Peck 2, 2a, 3. Ontario: Almonte, Fowler in 1898; Algonquin Park, Macoun 22022. New Jersey: Budds Lake, Mac- kenzie 2093; Cranberry Lake, Mackenzie 2109. Onto: Wauseon, Kellerman in 1899 (Ohio State Univ.). Inp1AnA: Gibson, Bebb 2935, Hill 97 in 1908; Griffith, Hill 50 in 1909. Micuiaan: Keweenaw County, Farwell 643; shore of Lake Superior, Wood in 1884. Wisconsin: Tomahawk, Cheney 2100; Newbold, Cheney 1700; Granite Heights, Cheney 3088. Minnesota: Hennepin County, Sandberg in 1890 (Univ. Minn. Herb.). Fia. 190.—Distribution of P. boreale. 109. Panicum dichotomum L. Panicum dichotomum L. Sp. Pl. 58. 1753. ‘‘Habitat in Virginia.’’ Since Linnzeus gives no description of his own, but quotes the diagnosis from Gronovius’s Flora Vir- ginica,? the type of this species is Clayton no. 458 which is the specimen cited by Gronovius,? and upon which his diagnosis is based. This specimen, preserved in the Gronovius Herbarium, is the autumnal form answering well to Gronovius’s character- ization, ‘‘vix pedale, in arbusculae formam excrescens.’’ The specimen in the Lin- nean Herbarium¢ collected by Kalm is P. microcarpon. Panicum angustifolium LeC. in Torr. Cat. Pl. N. Y. 91. 1819, not Ell. 1816. No specimen nor locality is cited. A vernal specimen in the Torrey Herbarium penciled “‘aneustifolius (nitid. var)’’ but without data may be the type. Panicum tremulum Spreng. Neu. Entd. 2: 103. 1821. ‘‘Panicum n. 39. Mihlenb. gram. p. 127.* In Nova Caesarea.’”’ [The * indicates a new species.] No locality is cited by Muhlenberg, and this number does not now exist in the Muhlenberg Her- barium. In the Sprengel Herbarium is a specimen consisting of a vernal culm with mature panicle, which bears a label ‘‘Panicum tremulum Spr. Hb. Spr. Torrey.” A second label reads ‘‘Panicum tremulum* Miihlenb. gram. p. 127.” Though no locality is given this specimen is doubtless the basis of the locality cited by Sprengel, and may be considered his type, since he gives a description and had not seen Muhlen- berg’s plant. Panicum dichotomum viride Vasey, U. 8S. Dept. Agr. Div. Bot. Bull. 8: 30. 1889. No locality nor specimen is cited. The diagnosis reads ‘‘Smooth all over, leaves light green and narrower.”’ In the National Herbarium are four specimens marked “‘var. aFil, Virg. 2: 133. 1743. b See Hitchcock, Contr. Nat. Herb. 12: 127. 1908, for an account of the grasses of Gronovius’s Flora Virginica. : c See Hitchcock, Contr. Nat. Herb. 12:114. 1908, for an account of the American grasses in the Linnean Herbarium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 191 viride Vasey” in Vasey’s handwriting. Of these a vernal specimen collected by L. F. Ward, Woodley Park, Washington, D. C., 1881, is chosen as the type, since it is entirely without pubescence, while the other specimens marked “‘ var. viride’’ by Vasey have pubescent spikelets or nodes, or are fragmentary. Panicum dichotomum divaricatum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 30. 1889. No specimen nor locality is cited. There are in the National Herbarium two specimens of the autumnal form of P. dichotomum marked ‘‘var. divaricatum” by Vasey. Of these, S. M. Tracy 127, Lake, Mississippi, is chosen as the type, since the other specimen is nearly devoid of spikelets. Certain other specimens marked by Vasey do not correspond to the description. Panicum nitidum pauciflorum Britton, Trans. N. Y. Acad. 9:14. 1889. Based on *P. dichotomum var. pauciflorum Vasey in Columbia College Herbarium.’’ The type is labeled ‘‘Panicum, Shady moist grounds, July” and consists of several spar- ingly branched culms of P. dichotomum with small panicles. Vasey’s herbarium name had not been published. Panicum nitidum viride Britton, Trans. N. Y. Acad. 9: 14. 1889. Based on P. dicho- tomum viride Vasey. Panicum dichotomum commune Wats. & Coulter in A. Gray, Man. ed. 6. 633. 1890. No specimen nor locality is cited. The name as published is ‘‘(a@) commune” and was probably meant to designate the common vernal form. No specimen marked “var. commune ’’ could be found in the Gray Herbarium. Panicum ramulosum viride Porter, Bull. Torrey Club 20: 194. 1893. Presumably based on P. dichotomum viride Vasey, since ‘‘(Vasey)’’ is given after the varietal name. DESCRIPTION. Vernal form often purplish; culms 30 to 50 cm. high, erect, from a knotted crown, the nodes naked or the lower with a few spreading hairs; sheaths less than half the length of the internodes, sometimes ciliate on the margin, otherwise glabrous, or the jowermost rarely sparingly pubescent; blades spreading, 5 to 11 em. long, 4 to 8 mm. wide, acuminate, slightly narrowed toward the base, glabrous on both surfaces, some- times with a few long hairs on the margin at the base, the basal blades lanceolate- ovate, long-ciliate on the margin near the base; pani- cles long-exserted, 4 to 9 cm. long, the axis and spread- ing branches flexuous, spikelet-bearing toward the ends; spikelets 2mm. long, 0.9 mm. wide, elliptic, glabrous or rarely pubescent; first glume one-third the length of the spikelet, subacute; second glume and sterile lemma rather faintly nerved, the glume shorter than the fruit at maturity; fruit 1.8 mm. ‘ long, 0.9 mm. wide, elliptic. ee Yromtype Autumnal culms much branched at the middle specimen in Gronovius Herbarium. i nodes, the lower portion usually erect and devoid of blades, thus giving the plants the appearance of diminutive trees as described by Gronovius and Linnzeus; blades much reduced and very numerous, often involute. This common and widely distributed species can be distinguished by its lack of pubescence, its smooth spikelets, 2 mm. long, and its erect autumnal form. A few specimens, such as Hitchcock 1292, Pollard 323, and Ward in 1881, from Wash- ington, D. C., which show all the other characters of P. dichotomum have pubescent spikelets. 192 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Dry or sterile woods, New Brunswick to Michigan and south to northern Florida and eastern Texas. New Brunswick: Bass River, Fowler. Marne: Mount Agamenticus, Parlin 1266, Ricker 1309. New Hampsutire: Walpole, Fernald 406. Vermont: West Rutland, Hggleston 1759. Massacuusetts: Framingham, Smith 736. Connecticut: Greens Farms, Pollard 16; Bridgeport, Hames in 1895; Oxford, Harger in Kneucker Gram. Exs. 245. New York: Long Island, Bicknell in 1905; Oxford, Coville in 1884; Washington County, Burnham 13. OntTARIO: Galt, Herriot 13. New Jersey: Morris County, Mackenzie 1405, 2280; Cranberry Lake, Mackenzie 2106. PENNSYLVANIA: Easton, Porter in 1895, 1897, and 1898; Lancaster County, Heller 4769, 4783; York County, Rose & Painter 8134; Mountainville, Pretz 1936. Ou1o: Berlin Heights, Moseley in 1895; Lancaster, Kellerman 6768; Sugar Grove, Kellerman 6891. Inp1iANA: Dune Park, Chase 1919, Umbach 1789; Miller, Chase 1543; Lafayette, Dorner 83. Intinors: Cobden, Waite in 1885. MicuicgANn: Port Huron, Dodge in 1899 and 1909; Grand Beach Springs, Hill 84 in 1908. Missouri: St. Francois County, Fig. 192.—Distribution of P. dichotomum. Eggert 246. DeLAwaARE: Wilmington, Commons 297; Centerville, Commons 294, 295. MARYLAND: Riverdale, Chase 2379, House 949; High Island, Pollard 324; Great Falls, Chase 2315. District or CoLumBiA: Hitchcock 122, 357, Kearney 5, 25, Williams 6, 8, 9. VirGiniA: Four-Mile Run, Hitchcock 358, Norfolk County, Kearney 300 in part, i 1374, Noyes 73, 75, 97; Dismal Swamp, Chase 3657. West Vireinia: Wyoming County, Morris 1193a; Fayette County, Morse in 1903. Norra Carouina: Chimney Rock Mountain, Biltmore Herb. 800c; Biltmore, Biltmore Herb. 800a in part; Blowing Rock Mountain, Small & Heller 480; Chapel Hill, Chase 3048, 30564. Sourn Carouna: Aiken, Kearney 234; Pickens County, Anderson 1201; Orange- burg, Hitchcock 1400, 1403; Graniteville, Cuthbert 962. GeoraiaA: Clark County, Harper 95; Albany, Tracy 3616, 3649; Rowe, Canby 219; Stone.Mountain, Eggert 444, 46, Hitchcock 1352; Pigeon Mountain, Wilson 179; Alecovey Mountain, Small in 1893. Froria: Duval County, Curtiss 3600A in part, Kearney 147; Waldo, Combs 687 in part; Orange Bend, Chase 4100. TrenNeESsSEE: Knoxville, Scribner in 1890; Polk County, Chambliss 39, 77, Kearney 329. AuaBama: Auburn, Tracy 3757, Hitchcock 1330; Flomaton, Tracy 3625 in part; Cullman County, Eggert 20. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 1983 Mississipri: Starkville, Tracy 1755; Enterprise, Tracy 3273; Meridian, Tracy 3253; Fairport, Tracy 3204, 3209; Nicholson, Kearney 367, 384; Biloxi, Tracy 2028, 2050. Louisiana: Calhoun, Hitchcock 1262, 1292; Shreveport, Hitchcock 1240, 1246, 1251; Calcasieu, Langlois 42 in 1884. Texas: Hardin County, Nealley in 1892; Beaumont, Reverchon 4155, 4158. 110. Panicum barbulatum Michx. Panicum barbulatum Michx. Fl. Bor. Amer. 1: 49. 1803. ‘‘Hab. in Carolina.” In the Michaux Herbarium is a sheet upon which are three specimens and two labels. One label reads ‘‘ Panicum barbulatum Hab. in Canada P. capillari affine. Ad ripas amnis: Rivierre a Jacques Cartier dicti legi,’’ the other “‘Rivierre a Jacques Cartier Route a Queb. P. barbulatum.’’ The two larger plants are the vernal form of the species described below, and do not belong to the species to which the name P. barbu- latum has been heretofore applied in our Manuals.b The third plant upon this sheet isasmall specimen of P. lindheimeri Nash. The only Carolina specimen from Michaux labeled P. barbulatum is one in the Drake de Castillo Herbarium sent out by Richard. This is labeled “‘Caroline,’’ but since the specimen, which is P. ashei, has glabrous nodes, and hence does not agree with Michaux’s description, it is rejected and the above specimens with bearded nodes from Canada are chosen as the type. Panicum dichotomum barbulatum Wood, Class-book ed. 3. 786. 1861. This is pre- sumably based on P. barbulatwm Michx., though no synonymy is cited. The descrip- tion given applies to P. microcarpon. Panicum pubescens barbulatum Britton, Cat. Pl. N. J. 280. 1889. Presumably based on P. barbulatum Michx., no synonymy nor description being given. Panicum nitidum barbulatum Chapm. FI. South. U. S. ed. 3. 586. 1897. Based on P. barbulatum Michx., though the description applies to P. microcarpon. Panicum gravius Hitchc. & Chase, Rhodora 8: 205. 1906. ‘‘Type Chase 3620; forming top-heavy tufts, in sandy, rather dry woods, on the old Commons farm, between Centreville and Mt. Cuba, Delaware, July 30, 1906; collected by Agnes Chase. Deposited in National Herbarium.” The name barbulatum is misspelled “‘barbatum” by Persoon,¢ and ‘“‘barbatulum” by Roemer and Schultes,@ the latter spelling being given by Steudel ¢ under P, boscit (page 253), P. deustum (page 255), and P. sphaerocarpon (page 263). DESCRIPTION. Vernal form in large tufts; culms slender, 50 to 80 cm. high, erect, or spreading at the summit, lower nodes usu- Fic.193.—P. barbulatum. ly bearded ; sheaths glabrous except a puberulent ring From type specimen. at the summit, the lower usually softly pubescent; blades spreading, 6 to 10 cm. long, 6 to 10 mm. wide (the lower shorter), acuminate, rounded at the base, glabrous, the lower rarely puberulent; panicles long-exserted, 6 to 11 cm. long, as wide or wider, the slender, flexuous branches fascicled, the lower spreading or drooping at maturity, spikelet-bearing at @ See Hitchcock, Contr. Nat. Herb. 12: 143. 1908, for an account of the American grasses in the Michaux Herbarium. b See P. microcarpon, page 181. ¢ Syn. Pl. 1: 84. 1805. @ Syst. Veg. 2: 447. 1817. ¢ Nom. Bot. ed. 2. 2: 253, 255, 263. 1841. 41616°—vo1 15—10——13 194 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. the ends; spikelets oval, 2 mm. long, 1 mm. wide, glabrous; first glume one-fourth to one-third as long as the spikelet, acute; second glume and sterile lemma equal, cover- ing the fruit at maturity; fruit elliptic, 1.8 mm. long, 1 mm. wide, obscurely apiculate. Autumnal form diffusely branched, forming very large, topheavy reclining bunches, the slender branchlets recurved, the numerous flat blades horizontally spreading. Closely allied to P. dichotomum L., from which it differs in the vernal form in having usually wider blades and bearded lower nodes and fruit covered by the equal second glume and sterile lemma; the autumnal form is distinguished by the large topheavy reclining tufts. DISTRIBUTION. Sterile or rocky woods, Massachusetts to Michigan and south to Georgia and eastern Texas. Massacuusetts: Malden, Frohock in 1879 (N. E. Bot. Club Herb.). Connecticut: Southington, Andrews 18, Bissell 5577; Groton, Graves 12. New York: St. Albans, Bicknell in 1905. New Jersey: South Amboy, Mackenzie 1548. PENNSYLVANIA: Easton, Porter in 1895; Lancaster County, Heller 4776. Onto: Vinton, Kellerman 6886. Inp1iANA: Dune Park, Hill 129 in 1906; Clarke County, Deam 6577; Brown County, Deam 6467a. MicuicaNn: Port Huron, Dodge in 1899; Park Lake, Wheeler 17 (both in Hitchcock Herb.). Missourtr: Eagle Rock, Bush 148; Monteer, Bush 2877, 3529, 4733; Swan, Bush 3369, 3456A, 4473; Pleasant Grove, Bush 313, 720; Howell County, Bush 51; Chadwick, Bush 4458. DeLawaRrReE: Wilmington, Canby 10 in 1891; Centerville, Chase 3620, Commons 296. MARYLAND: Riverdale, Chase 3643; Lanham, Chase 3484; Chesa- peake Junction, Hitchcock 1641, 2411; Beltsville, Chase 3747, 3758; High Island, Pol- Fic. 194.—Distribution of P. barbulatum. lard 323. District or CotumBriA: [Hitchcock 123, 125, Kearney in 1897. VireiniA: Smyth County, Small in 1892; Arlington, Hitchcock 124; Clifton Forge, Tidestrom 22. e West VirGinisA: Fayette County, Kellerman 6903. Norra Carona: Biltmore, Biltmore Herb. 800a in part; Caraleigh Junction, Chase 3090; Wards Mill, Chase 3188. Souts CAROLINA: Orangeburg, Hitchcock 360, 1416. GeoratiA: Stone Mountain, Chase 4519, Hitchcock 359, 1350. Kentucky: Harlan County, Kearney 35 in part, 57 in part. TENNESSEE: Nashville, Gattinger in 1884 (Univ. Tenn. Herb.). ALABAMA: Pisgah, Chase 4469; Scottsboro, Chase 4502. Mississrerr: Saratoga, Tracy 8403. Arxansas: Benton County, Plank 9, 10a, 41; Fulton, Bush 2532. LoursiAna: Shreveport, Hitchcock 1252. Texas: Houston, Plank 93; Mineola, Reverchon 4147. OKLAHOMA: Poteau, Hitchcock in 1903 (Hitchcock Herb.). HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 195 111. Panicum yadkinense Ashe. ? Panicum dumus Desv. Opusc. 88. 1831. ‘‘Habitat in America calidiori.’? The type, in the Desvaux Herbarium, is a fragment of a branch of some species in this group. The immature, glabrous spikelets, 2.3 mm. long, the second glume and sterile lemma pointed beyond the fertile lemma, suggest P. yadkinense, though it may be P. barbulatum, the pointed spikelets being due to immaturity and withering. Panicum maculatum Ashe, Journ. Elisha Mitchell Soc. 15: 44. 1898, not Aubl. 1775. “Collected by the writer at Raleigh, N. C., May, 1895.’’ The type of this could not be found in Ashe’s herbarium. In the National Herbarium is a specimen labeled in Ashe’s handwriting, “‘Panicum maculatum, Raleigh, May, 1895’’ which is probably a duplicate type. This plant belongs to the species here described though it does not agree in all respects with the original description of Panicum maculatum. The spikelets are there said to be { lines long, and ‘‘about the size of those of P. barbu- latum”’ [P. microcarpon of this monograph], and the species is said to be distinguished from P. dichotomum by the smaller spikelets. The specimen from Raleigh has spike- lets larger than those of P. dichotomum, being about 2.5 mm. long. Since this speci- men belongs to the species as understood by Ashe, it is probable that the description of the spikelets was based upon an admixture of P. microcarpon, as the two species are frequently found growing together. Panicum yadkinense Ashe, Journ. Elisha Mitchell Soc. 16: 85. 1900. Based on “P. maculatum Ashe, not P. maculatum Aubl.”’ DESCRIPTION. Vernal form similar to that of P. dichotomum but culms taller and stouter, some- times 1 meter high; sheaths usually bearing pale, glandular spots; blades 9 to 13 cm. long, 8 to 11 mm. wide, panicle about 10 to 12 cm. long, about three-fourths as wide, the long lower branches ascending; spikelets 2.3 to 2.5 mm. long, 1 mm. wide, elliptic to subfusiform, pointed, glabrous; first glume about one-third the length of the spikelet, usually blunt; second glume and sterile lemma rather faintly nerved, equal, exceeding the fruit and forming a slight point beyond it; fruit 1.9 mm. long, 0.9 mm. wide, elliptic, subobtuse. Autumnal form erect or leaning, loosely branch- ing from the middle nodes, the blades smaller but not conspicuously reduced. This species differs from P. dichotomum in the fia. 195.—P. yadkinense. From dupli- vernal form in its larger size and longer, acute ©#t¢ type specimen of P. maculatum . . Ashe in National Herbarium. spikelets, and in the autumnal form in the com- paratively few branches, which do not form a bushy crown. Occasionally the branches are rather numerous, though not closely fascicled and bushy, for example, Harper 1349, and Porter from Pennsylvania in 1895. The following two specimens, Chase 3072 and Hitchcock 1416, are referred to this species, though the spikelets are scarcely over 2 mm. long; that is, the second glume and sterile lemma do not extend into a point as in typical spikelets. DISTRIBUTION. Moist woods and thickets, Pennsylvania to Georgia, west to southern Illinois and Louisiana. PENNSYLVANIA: Easton, Porter in 1895. Inuinors: Makanda, Gleason in 1903; Johnson County, Schneck in 1902 (Hitch- cock Herb.). 196 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DeLAWwARE: Wilmington, Chase 3616. MaryLaAnpb: Little Falls, Vasey in 1884; Cabin John, Chase 2853, 3772; West Chevy Chase, Chase 2946, Hitchcock 361. District or CotumBtIA: House 911, | Kearney 28 in part. Virainia: Arlington, Chase 2964. Nortu Carouina: Raleigh, Ashe in 1895, Chapel Hill, Chase 3059, 3061, 3072; Jacksonville, Chase 3192. South Carona: Orangeburg, Hitchcock 6, 14164. Gxzorata: Dublin, Harper 1349. Fic. 196.—Distribution of P. yadkinense. TENNESSEE: Sumner County, Gat- tinger in 1883 (Univ. Tenn. Herb.). AxaBaMa: Tensaw, Tracy 8029. Lovuistana: Lake Charles, Hitchcock 1164. 112. Panicum roanokense Ashe. Panicum roanokense Ashe, Journ. Elisha Mitchell Soc. 15: 44. 1898. “‘Type material collected by writer in dry soil, Roanoke Island, N. C. June, 1898. Also collected at Rose Bay and Mackleyville, N. C., the same month.’”’ The type could not be found in Ashe’s herbarium. In the Biltmore Herbarium is a specimen from Manteo, Roanoke Island, N. C., collected by Ashe, June 10, 1898, and labeled by him Panicum roanokense Ashe. This is a duplicate type or possibly the type. It con- sists of two vernal culms with mature primary panicles. Panicum curtivaginum Ashe, Journ. Elisha Mitchell Soc. 16:85. 1900. ‘‘Collected at Petit Bois Island, Mississippi, May 8, 1898 by S. M. Tracy.”’ An unmounted specimen of the collection cited was found in a cover marked “P. curtivaginatum sp. nov.’”’ in Ashe’s herbarium. No name was written on the Tracy label, which bears the number 4584. As this was the only specimen of this collection found in Ashe’s herbarium it is taken as the type. It consists of a tuft of three slender vernal culms with over-mature panicles. The autumnal form is not represented, but in a specimen of Tracy 4584 in the National Herbarium the autumnal culms of the previous year are attached to the tuft. The spikelets are described as “‘quite 1.5 mm.”’ long, but they measure 2 mm. DESCRIPTION. Vernal form cespitose, somewhat glaucous olive green; culms erect or ascending, 50 to 100 cm. high; Fig. 197.—P. roanokense. From sheaths half as long as the internodes or less, gla- ete collected by Ashe at} yous, or the lowermost sometimes sparsely pubescent; : blades at first stiffly erect, later ascending or spread- ing, 6 to 9 cm. long, 3 to 8 mm. wide, tapering to both ends, glabrous or with a few hairs around the base; panicles 4 to 8 cm. long, scarcely as wide, the branches spreading; spikelets 2mm. long, 1 mm. wide, ellipsoid-obovoid, very turgid, glabrous; first glume about one-third the length of the spikelet; second glume and sterile lemma strongly nerved, subequal, the glume rather conspicuously purple-tinged at base, scarcely covering the fruit at maturity; fruit 1.6 mm. long, 0.9 mm, wide, ellipsoid. Autumnal form erect or decumbent, branching at the middle and upper nodes, the branches numerous but not in tufts, the primary internodes elongating and becoming HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 197 arched about the time the branches appear; the reduced blades more or less involute, not exceeding the 1.5 to 4 cm. long panicles; basal blades firm, erect, often as much as 5 or 6 cm. long. The plant is glabrous throughout with exceptions mentioned; the glaucous olive green color and very turgid spikelets, purple-stained at base, are characteristic. Harper’s number 458, from Sumter County, Georgia, is doubtfully referred to this species. The first glume is very short, the panicle narrow with few, appressed branches, and the blades are long and narrow. DISTRIBUTION. Open swampy woods or wet peaty meadows, southeastern Virginia to Florida and west to eastern Texas. Virainta: Near Norfolk, Kearney 1514, 2026. Nortu Carona: Rose Bay, Ashe in 1898; Lake Mattamuskeet, Chase 3203; Roanoke Island, Chase 3240, 3247; Wards Mill, Chase 3178. Fiorina: Baldwin, Combs 60, Hitchcock 987, 998; Mabel, Curtiss 6636; Tampa, Hitchcock 9384, 939. AtaBAMA: Flomaton, Tracy 3625 in part. Mississipri: Petit Bois Island, Tracy 4584, Ocean Springs, Tracy 4592. Lourstana: Lake Charles, Hitchcock 1144. Texas: Waller, Hitchcock 1174. Fie. 198.—Distribution of P. roanokense. 113. Panicum caerulescens Hack. ._ Panicum caerulescens Hack.; Hitche. Contr. Nat. Herb. 12: 219. 1909. ‘‘The type is Hitchcock 706. In glade among Spartina, etc., stretching up through the tall grass, Miami, Florida, April 3, 1906, U. S. National Herbarium no. 558380.”’ This specimen consists of two tufts, some of the culms beginning to branch and with over- mature primary panicles, and some freely branching. DESCRIPTION. Vernal form similar to that of P. roanokense; culms more slender, rarely over 75 cm. high; blades ascending or spreading, commonly purplish beneath, 5 to 8 em. long, 4 to 7 mm. wide, the margins nearly parallel for two-thirds their length; panicles usually short-exserted, 3 to 7 cm. long, half as wide or less, the branches narrowly ascending; spikelets 1.5 to 1.6 mm. long, 0.9 mm. wide, obovoid, blunt, very turgid, glabrous; first glume about one-third the length of the spikelets; second glumeand sterile lemma subequal, the glume scarcely as long as the fruit at ma- turity; fruit 1.4 mm. long, 0.8 mm. wide, ellipsoid. Autumnal form erect or leaning, sometimes decumbent at 5 co redsoone, at base, producing short, densely fascicled branches at From type specimen. the middle and upper nodes, these tufts scarcely as long as the primary internodes, the reduced blades ascending, more or less involute, the reduced panicles with a few long-pediceled spikelets. This species is distinguished from P. roanokense by the narrow panicles and smaller spikelets and by the tufted branches of the autumnal form. 198 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. In marshes and swampy woods, southeastern New Jersey to Florida, west to Missis- sippi; also in the Bahamas and Cuba. New Jersey: Cape May, Stone in 1909. Virainia: Lynn Haven, Chase 5417, Hitchcock 356. Fioripa: Levy County, Combs 803; Titusville, Chase 3992; Miami, Hitchcock 706, 715; Homestead, Hitchcock 690; Braidentown, Hitchcock 965; Myers, Hitchcock 897, 904, 915. AuaBAMA: Fort Morgan, Tracy 8401. Mississippi: Horn Island, Tracy in 1903. Baunamas: New Providence, Brit- ton & Brace 597, 599, Mills- paugh 2182, Northrup 248; Great Bahamas, Brace 3524, Britton & Millspaugh 2506, Fig. 200.—Distribution of P. caerulescens. 2668; Andros, Brace 7015 (all in Field Mus. Herb.); New Providence, Eggers 4305 (Hackel Herb.), Eggers 4312 (Krug & Urban Herb.). Cusa: Without locality, Wright 3463 in part. 114. Panicum lucidum Ashe. Panicum lucidum Ashe, Journ. Elisha Mitchell Soc. 15:47. 1898. “Collected in June 1898 by the writer in deep, shady swamps bordering Lake Mattamuskeet, N. C.”? There is no specimen in Ashe’s herbarium from the type locality, but there is a specimen of the vernal form in the National Herbarium collected by Ashe in 1898 at Lake Mattamuskeet. This specimen is either the type or a duplicate type. The label is in Ashe’s handwriting. : Panicum taxodiorum Ashe, Journ. Elisha Mitchell Soc. 16:91. 1900. ‘Type: K. K. McKenzie’s no. 460. Hummocks in cypress swamps. Lake Charles, La., September 1890.’ The type, in Ashe’s herbarium, is a specimen passing from the vernal to the autumnal form and showing the early branching condition. DESCRIPTION. Vernal form at first erect and resembling that of P. dichotomum, but the weak culms soon becoming decumbent, sometimes rooting at the lower nodes; sheaths glabrous, usually ciliate on the margin; blades thin, bright green, shining, glabrous, at first erect, but soon widely spreading, 4 to 7 cm. long, 4 to 6 mm. wide; panicles resembling those of P. dichotomum but fewer-flowered; spikelets 2 to 2.1 mm. long, 1 mm. wide, elliptic, glabrous (rarely obscurely pubescent); first glume about two-fifths the length of the spikelet, pointed; second glume and sterile Jemma more strongly nerved than in P. dichotomum, both shorter than the fruit at maturity; fruit 1.7 mm. long, 0.9 mm. wide, slightly pointed. Autumnal form repeatedly branching, forming large clumps or mats of slender, weak, vine-like culms, the branches elongated and diverging at a wide angle, not fascicled, the blades 2 to 4 cm. long, waxy, flat and spreading; panicles much reduced, with few long-pediceled spikelets; basal blades linear-oblong, as much as 10 cm. long. Under a lens the oblong epidermal cells are visible between the nerves in the blades, especially on the lower surface, giving a minutely bullate surface characteristic of this species and of no other in this group. Fic. 201.—P. lucidum. From type specimen in National Herbarium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 199 DISTRIBUTION. Wet woods and sphagnum swamps, along the Coastal Plain from New York to Florida and west to eastern Texas. New Yorx: Woodmere, Bicknell in 1902; Hempstead, Bicknell in 1903. New Jersey: Speedwell, Stone 7; Wildwood, Pollard in 1897; Tuckerton, Chase 3599; Forked River, Chase 3593; Atsion, Chase 3550, 3554; South Amboy, Mackenzie 2167. Inprana: Dune Park, Umbach 4962. DELAWARE: Ogletown, Canby 11. Marytanp: Beltsville, Chase 3743; Lanham, Chase 3475. District or Cotumsia: Chase 5418, Greene in 1908, Kearney in 1897, Pollard 403, Scribner in 1894, Steele in 1899. Virermia: Fort Myer, Williams in 1898; Lynn Haven, Hitchcock 364. Nort Carouina: Wilmington, Chase 3112, 3159, [Hitchcock 365, 367, 368, 369, 1442, 1470, Kearney 260; Jack- sonville, Chase 3197; Lake Mat- tamuskeet, Ashe in 1898; Rowan County, Smallin 1894; Biltmore, Biltmore Herb. 5066b, Hitchcock 362. Sourn Carouta: Aiken, Kearney 288 in part, Orangeburg, Hitchcock 363, 366. Groreta: Clarke County, Harper 88; Randolph County, Harper 1760; Thomson, Bartlett 1136; Fia. 202.—Distribution of P. lucidum. Augusta, Cuthbert 529. Fiormwa: Jacksonville, Curtiss 6601; eastern Florida, Palmer 632 in 1874; Lake City, Hitchcock 1026; Argyle, Curtiss 6403; Apalachicola, Biltmore Herb. 800b; Milton, Chase 4320; Washington County, Combs 615; Waldo, Combs 687 in part; Homosassa, Combs 934; Eustis, Chase 4068, Nash 337, 500; Bartow, Combs 1218; Myers, [Hitchcock 919, Lee Co. Pl. 481. AvaBAMA: Auburn, Pollard & Mazon 54, Tracy 3749; Flomaton, Hitchcock 1059; Mobile, Kearney 45. Mississippi: Taylorville, Tracy 8405; Magee, Trccy 8504; Waynesboro, Kearney 167; Ocean Springs, Tracy 95. Lovuistana: Oberlin, Ball 202; Lake Charles, Mackenzie 460. Texas: Colmesneil, Nealley 35 in 1892. In the herbarium of the Philadelphia Academy is a specimen said to be from Brazil which appears to be P. lucidum. ' i] : pwm nnn! § See ae } See f 115. Panicum sphagnicola Nash. Panicum sphagnicolum{cola] Nash, Bull. Torrey Club 22: 422. 1895. ‘The late and much branched state was collected by the writer this summer in a sphagnum bog at Lake City, Florida, and will be distributed as No. 2500.’’ The type, in Nash’s herbarium, consists of several culms 45 to 55 cm. high, with long internodes and divaricate branches; the primary panicle is devoid of spikelets, the secondary panicles are small and few-flowered. There are three sheets of this collection in Nash’s her- barium, none of which is marked type. The foregoing refers to the largest specimen. 200 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Vernal form grayish olive green, cespitose; culms slender, strongly flattened, erect, or reclining, 50 to 100 cm. high; sheaths glabrous or the lowermost sparsely papillose- pilose, soon becoming divaricate and enveloping the internodes only at base; blades at first erect, later widely spreading, glabrous, 5 to 8 cm. long, 3 to 7 mm. wide; panicles narrow, 5 to 6 cm. long, the branches ascending or somewhat spreading, not spikelet-bearing at the base; spikelets 2.5 mm. long, 1.1 mm. wide, elliptic; first glume nearly half the length of the spikelet, subacute; second glume and sterile lemma strongly nerved, minutely pubescent to- ward the summit or glabrous, the glume shorter than the fruit; fruit 2 mm. long, 1 to 1.1 mm. wide, elliptic, subobtuse. Fie. 203-—P. sphagnicola, “From Autumnal form decumbent or finally prostrate- type specimen. spreading, divaricately branching from all the nodes, the branches slender and elongated, some- times rooting at the nodes; sheaths divaricately spreading from the stem, usually nearly as long as the blades; blades flat, reduced in length but not much in width, mostly 1 to 2 cm. long, or on the ultimate branchlets only 5 mm. long and 1 mm. wide; panicles rather few, reduced to a few short-pediceled spikelets; basal blades 4 to 8 cm. long, about 1 cm. wide, sometimes sparsely pilose at base. This species is readily distinguished in the autumnal form by its slender, widely- spreading branches and divaricate sheaths. At this stage the primary sheath may subtend two branches, each with its conspicuous prophyllum, 5 to 15 mm. long, ciliate on the keels and bearing a tuft of hairs at the acuminate tip. The leaf of the second branch is much reduced and inclosed in the base of the primary sheath. Panicum lucidum, the only other spe- cies with a like autumnal habit, is much more slender, more leafy, and bright green and shining, and has smaller long-pediceled spikelets. DISTRIBUTION. Edges of cypress swamps, in sphag- num bogs, and in similar moist, shady places, southern Georgia and Florida. Geroreia: Darien, Biltmore Herb. 5066 e (Biltmore Herb.). Fioripa: Lake City, Bitting 18, Combs 73, Hitchcock 1006, Nash 2500; Sanford, Chase 4039; Levy County, Combs 838; eastern Florida, Palmer 633 in 1874 (Gray Herb.). Fig. 204.—Distribution of P. sphagnicola. Spreta.—Culms tufted, rather stiff, mostly glabrous or nearly so; ligules densely hairy, 3 to 5 mm. long; blades not over 8 mm. wide; spikelets 1 to 1.6 mm. long, pubescent or rarely glabrous, second glume and sterile lemma 5 to 7 nerved. Autumnal form with more or less tufted branchlets and niuch reduced leaves and panicles. Panicle narrow, one-fourth to one-third as wide as long... .. 116. P. spretum. Panicle open, two-thirds as wide as long, or more. Spikelots 1h mum done. ... 4. ena tetedis-aokeacboeeee 117. P. lindheimeri. t HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 201 Spikelets 1.3 mm. long or less. Culms and sheaths glabrous..............-...-.+--- 119. P. longiligula- tum. Culms and sheaths appressed-pubescent. Spikelets 1.2 to 1.3 mm. long. ...............-118. P. leucothriz. Spikelets not over 1 mm. long. .......--..--.-- 120. P. wrightianum. 116. Panicum spretum Schult. Panicum spretum Schult. Mant. 2: 248. 1824. Based on ‘‘ Miihlenb. Descr. ub. p. 125. n. 37. (sine nomine).’’ Muhlenberg’s description is copied, but slightly rear- ranged, and the locality ‘‘N. Anglica” also copied. The type specimen, in the Muhlenberg Herbarium, is a vernal culm labeled “‘No. 2. Panicum an capillare? In moist ground. Mon. 184. M. 116a.’”’ On the folio ‘‘N. Angl.” is written after this number. Panicum nitidum densiflorum Rand & Redfield, Fl. Mt. Desert 174. 1894. ‘‘Shore of Ripples Pond (Rand).’’ The type, in Rand’s herbarium, collected July 28, 1892, - has a narrow many-flowered panicle with pubescent spikelets like the above- mentioned plant in the Muhlenberg Herbarium. Panicum eatoni Nash, Bull. Torrey Club 25: 84. 1898. Collected by ‘‘Alvah A. Eaton * * * at Seabrook, N. H.’’ The type, in Nash’s herbarium, consists of two vernal culms with spikelets measuring 1.5 to 1.6 mm. long. Panicum octonodum Smith, U.S. Dept. Agr. Div. Agrost. Bull. 17:73. 7.369. June 30,1899. ‘“‘Texas.’’ This species was republished@ with more complete description as ‘“‘Panicum octonodum Scribn. & Smith, sp. nov.,’’ with the following citation: “‘Waller County, Texas. Collected by F. W. Thurow, May 5, 1898.’’ The type, in the National Herbarium, is a wholly glabrous vernal plant with glabrous spikelets 1.5 mm. long. Panicum paucipilum Nash, Bull. Torrey Club 26: 573. 1899. “‘Type collected by Mr. E. P. Bicknell, at Wildwood, New Jersey, May 30 and 31, 1897.’’ The type, in Nash’s herbarium, consists of four vernal culms, beginning to branch at the middle nodes, the sheaths sparsely ciliate toward the summit, the spikelets pubescent, 1.4 to _ 15 mm. long. Panicum paucipilum was described as differing from P. eatoni in having ‘‘much smaller spikelets with the first scale glabrous.”’ Panicum nitidum octonodum Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Bull. 24: 34.1901. Based on P. octonodum Smith. Panicum spretum has been referred to Panicum nitidum Lam. and was discussed by Scribner® in an article on that species. Scribner, however, based his identification of the latter upon a tracing made by A. H. Baldwin of a specimen in the Michaux Herbarium. The type of P. nitidum Lam.¢ is in the Lamarck Herbarium and is different from P. spretum. The plant from which Baldwin’s tracing was made is P. angustifolium or a closely allied species.¢ DESCRIPTION. Vernal culms tufted, 30 to 90 cm. high, erect or slightly decumbent at base, some- times sending out rootlets from the lower nodes, glabrous, the nodes swollen; sheaths loose, shorter than the internodes, usually ciliate on the margin toward the summit, otherwise glabrous, or the lower sometimes slightly pubescent; ligules 2 to 3 mm. long; blades firm, ascending or often reflexed, 7 to 10 cm. long, 4 to 8 mm. wide, sparingly aU.S. Dept. Agr. Div. Agrost. Circ. 16:5. July 1, 1899. 6U.S8. Dept. Agr. Div. Agrost. Bull. 24: 31. 1901. ¢ See p. 183. d See Hitchcock, Contr. Nat. Herb. 12: 148. 1908. 202 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. long-ciliate at the rounded base, otherwise glabrous; panicles 8 to 12 cm. long, one- fourth to one-third as wide, rather densely flowered, the branches ascending or appressed, short spikelet-bearing branchlets in the axils; spikelets 1.4 to 1.6 mm. (usually 1.5 mm.) long, 0.7 to 0.9 mm. wide, elliptic, obscurely pointed; first glume one-fourth to one-third the length of the spikelet, obtuse or subacute; second glume and sterile lemma equaling the fruit at maturity, pubescent or rarely glabrous; fruit 5 mm. long, 0.7 to 0.8 mm. wide, elliptic, slightly pointed. Autumnal form more or less reclining, branching after the maturity of the primary panicle, the earlier branches elongated, ascending but not appressed, bearing exserted panicles, the subsequent branchlets in short fascicles, the blades much reduced, sometimes minutely pubes- cent, overtopping the small ultimate panicles; winter rosette appearing rather early, the blades glabrous or nearly so. Specimens with spikelets 1.4 mm., or even 1.3 mm. long, occur. The type of P. paucipilum Nash is such Fic. 205.—P. spretwum. From a specimen, as are Chase 2333 and Hitchcock 553. In the ele cars in Muhlenberg herbarium of the Philadelphia Academy are a number ° of such specimens, some with spikelets only 1.3 mm. long. But since no other character can be correlated with the smaller spikelets, and since specimens with spikelets 1.5 mm. long are much more numerous, we are unable to separate specifically the extremes of this species. It does not seem advisable to recognize as a subspecies the form with glabrous spikelets. Besides the Texas plants, in which the spikelets are glabrous, similar specimens have been collected in Delaware, namely, Canby 4, Commons 340, and Hitchcock 553. The Commons specimen consists of two plants, one with glabrous and one with pubescent spikelets, but otherwise alike. DISTRIBUTION. Wet and usually sandy soil, mostly near the coast, Maine to Texas; also in northern Indiana. Marne: York County, Fernald 510. New Hamesuire: East Kingston, Haton in 1898 (Biltmore Herb.). Massacuusetts: Essex County, Conant in 1881; Dedham, Bartlett 807. Connecticut: Waterford, Graves 80, 87; East Lyme, Graves 157. Ruope Istanp: Kingston, Piper in 1907. New Yorx: Riverhead, Bicknell in 1905, Peck 4. New Jersey: Atsion, Chase 3551, 3569; Bear Swamp, Stone 2; Wildwood, Bicknell in 1897. PENNSYLVANIA: Westchester, Win- dle 2 (Hitchcock Herb.). Inpiana: Dune Park, Hill 128 in Fig. 206.—Distribution of P. spretum. 1906, Umbach 1799; Miller, Pepoon in 1898; Michigan City, Hill 162 in 1906. DELAWARE: Cape Henlopen, Commons 340; Townsend, Canby in 1891; Milton, Commons 348; Lewes, Hitchcock 553. Mary.Lanpb: College Park, Novik in 1907. Virainia: Cape Henry, Chase 2333, 5421; Lynn Haven, Hitchcock 378, 379. Nort Carona: Wilsons Mills, Chase 3093. | ' ‘ i i ' ' ‘ ‘ 1 : HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 203 Froripa: Baldwin, Hitchcock 10053; Apalachicola, Biltmore Herb. 6028 in part, Chapman. AtaBaMA: Gateswood, Tracy 8433; without locality, Buckley (Mo. Bot. Gard. Herb.). : Mississippi: Ocean Springs, Tracy 91, 4585; Beauvoir, Tracy 4594. Texas: Waller County, Hitchcock 1175, Thurow 6. 117. Panicum lindheimeri Nash. Panicum lindheimeri Nash, Bull. Torrey Club 24: 196. 1897. ‘‘The type was col- lected by F. Lindheimer in 1846, no. 565.’’ The type, in Nash’s herbarium, consists of two rather slender vernal culms geniculate at the lower nodes, sparsely papillose- pilose below, beginning to branch at some of the nodes. No locality other than Texas is given on the label of the type nor on that of Lindheimer 565 in the National Tlerbarium, but on that of another specimen of this collection in the herbarium of the Missouri Botanical Garden the following data are given: ‘‘Springs, banks of the Guadeloupe, near New Braunfels.’’ Panicum funstoni Scribn. & Merr. U. 8. Dept. Agr. Div. Agrost. Circ. 35: 4. 1901. “Type specimen collected on the bank of Kaweah River at Three Rivers, Tulare County, Cal., no. 1286, Coville & Funston, July 26, 1891.’ The type, in the National Herbarium, consists of two branching culms, one sending out a rootlet at the second node (indicating that the culm was prostrate); sheaths and lower internodes rather strongly papillose-pubescent. This is the species described under P. nitidum Lam. in Britton’s Manual.¢ DESCRIPTION. Vernal culms stiffly ascending or spreading, 30 to 100 cm. high, glabrous, or lower internodes ascending-pubescent, the nodes swollen; sheaths less than half as long as the elongated internodes, ciliate on the margin, otherwise glabrous, or the lower ascending-pubescent; ligules 4 to 5 mm. long; blades usually firm, 5 to 10 cm. long, 6 to 8 mm. wide, at first ascending, soon spreading, papillose-ciliate at the rounded base, glabrous on both surfaces, or minutely puberulent beneath; panicles 4 to 7 em. long (rarely longer), nearly as wide, branches ascending or spreading, loosely flowered; spikelets 1.4 to 1.6 mm. long, 0.8 to 0.9 mm. wide, obovate, obtuse, turgid, pubescent; first glume one-fourth as long as the spikelet or less, usually obtuse; second glume and sterile lemma scarcely equaling the fruit at maturity; fruit 1.3 to 1.4 mm. long, 0.8 mm. wide, elliptic, obtuse. Autumnal! form usually stiffly spreading or radiate-pros- trate, internodes elongated, with tufts of short, appressed branches at the nodes; blades reduced, involute-pointed and often conspicuously ciliate at base. This common and widely distributed species is variable as to pubescence. Usually the plants are glabrous except the lower internodes and sheaths, but sometimes the pubescence extends nearly to the summit. These more pubescent specimens, such as the type of P. funstoni from California, Macown 26338 from Ontario, Chase 3464 from Maryland, and Tracy 7947 from Texas, in the vernal form resemble less pubescent specimens of P. tennesseense but can be distinguished by the smaller spikelets. In the autumnal form the stiffly radiating culms with the tufts of short branches also distinguish this species. Fic. 207.—P. lindheimeri. From type specimen. 4 Man..85. 1901. 204 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Dry sandy or sterile woods or open ground, Maine to northern Florida, and west to southern California. Marne: North Berwick, Parlin 1607 (Gray Herb.). New Hampsnrre: Summers Falls, Eggleston in 1893 (Biltmore Herb.). Vermont: Willoughby, EL. & C. E. Faxon in 1896. MassacuHusetTts: Framingham, Smith 734. Connecticut: Southington, Andrews 14, Bissell 5583; East Lyme, Graves 158; Franklin, Graves 76. New York: Bergen, Hill 1834 in 1907; Long Point, Bicknell in 1906; Mineola, Bicknell in 1906; Mambasset Neck, Bicknell in 1908; Long Island, Bicknell in 1902 and 1995. Ontario: Niagara Falls, Macown 26338; Ottawa, Macoun 65370; Port Colborne, Macoun 26316; Sarnia, Dodge 49. New Jersey: Clifton, Nash in 1892; Forked River, Chase 3589; Atsion, Chase 3528, 3572, Commons 68, 70; Wildwood, Heritage in 1897; South Amboy, Mackenzie 2156, 2160, 2164, 2168, 2349. PENNSYLVANIA: Ridley, Smith 157. Onto: Ashtabula County, Kellerman in 1888 (Ohio State Univ. Herb.). Inprana: Miller, Umbach 2353. Ixtinors: Chicago, Nelson in 1899; Beach, Umbach 2242; Urbana, Seymour in 1880; St. Clair County, Eggert 237; Jackson County, French in 1871 in part. Micuican: Port Huron, Dodge in 1909. Wisconsin: Witches Gulch, Che- ney 3872. Minnesota: Sandy Beach, Mac- Millan & Sheldon 1703 (Univ. Minn. Herb.). Missovrt: Allenton, Kellogg 1, 4, 5; Cliff Cave, Kellogg 10; St. Fic. 208.—Distribution of P. lindheimeri. Louis, Hitchcock 552. DeLAwaRrE: Wilmington, Canby 18, Commons 55, 64, 67, 71, 291, 363; Ogletown, Commons 66; Millsboro, Commons 42; Newport, Canby 3. MaryiAnp: Chesapeake Beach, Chase 3255, 3259, Hitchcock 1603; Lanham, Chase 3464, 34734; Owings, Hitchcock 1622; Chevy Chase, Chase 2887; Belts- ville, Chase 3729. District or CotumBIA: Chase 2985, 5422, Hitchcock 380, Kearney 18, 26, Ward in 1879. Virernia: Alexandria County, Chase 5423, in Kneucker Gram. Exs. 552; Norfolk, Kearney 309; Portsmouth, Chase 3686, Noyes 92. West Virainia: Summers County, Morris 984. Norra Carona: Jacksonville, Chase 3198; Wilsons Mills, Chase 3108; West Durham, Chase 3047; Magnetic City, Wetherby 18, 51; Biltmore, Boynton 5. Groraia: Americus, 7'racy 3889 in part; Stone Mountain, Hitchcock 381. Fioripa: Chattahoochee, Tracy 3615. TENNESSEE: Ducktown, Chambliss 27; Coffee County, Eggert 34. Atapama: Mobile, Kearney 32. Mississippi: Enterprise, Tracy 3285; Meridian, Tracy 3265; Starkville, Chase 4449; Biloxi, Tracy 6736. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 205 Lovrstana: Calhoun, Ball 53; Hitchcock 1278, 1287; Shreveport, Cocks 3508; Mandeville, Langlois 42; Abbeville, Langlois 38; Lake Charles, Chase 4400, Hitchcock 1136, 1138, 1156, 1165. Texas: Waller County, Hitchcock 1202, 1203, 1215; Thurow 3, 4, 12, 13, 14, 16, 19, 20, 24, 26, 27, 28, 29, 33; Columbia, Bush 178; Weatherford, Tracy 7944, 7947; La Grange, Plank 97; Huntsville, Plank 63; Gladewater, Reverchon 2357; Ennis, Smith in 1897; Kerrville, Heller 1752, 1888; Houston, Bebb 1276; Denison, Bebb 1428; New Braunfels, Lindheimer 565; Fort Smith to the Rio Grande, Bigelow; without locality, Nealley in 1884 and 1888. Ox.anoma: Poteau, Hitchcock 551; without locality, Palmer 384 in 1868. New Mexico: Without locality, Wright 2088, 2085 (the latter in Gray Herb.). Catirorni: Three Rivers, Coville & Funston 1286; Sacramento, Michener 142. 118. Panicum leucothrix Nash. Panicum leucothriz Nash, Bull. Torrey Club 24: 41. 1897. ‘Type collected by the writer in the low pine land at Eustis, Lake County, Florida, in the latter part of July, 1894, no’1338.’’ The type, in Nash’s herbarium, consists of somewhat branch- ing primary culms, decumbent at base. The description reads: ‘Spikelets obovate, about 0.65 mm. long, 0.4 mm. wide.’’ This is evidently an error, as the spikelets of the type measure 1.2 mm. as do also those of Nash 334 and 467 cited with the descrip- tion. Panicum parvispiculum Nash, Bull. Torrey Club 24: 347. 1897. ‘Type collected by Dr. John K. Small at Darien Junction, McIntosh Co., Ga., June 25-27, 1895.” The type, in Nash’s herbarium, consists of a tuft of mature vernal culms, beginning to branch. The culms and sheaths are appressed-pubescent, though less copiously so than is the type of P. leucothriz, and the panicles are larger. In the description the spikelets are given as 1.5 mm. long, but those of the type measure 1.3 mm. DESCRIPTION. Vernal plants light olive green, often purplish tinged; culms tufted, 25 to 45 cm. high, erect or ascending, appressed papillose pilose, the nodes scarcely swollen, pubescent; sheaths shorter than the internodes, papillose-pubescent, the hairs less appressed than those of the culm, rarely nearly glabrous, the margins ciliate, densely so at the summit; ligules 3 mm. long; blades rathgr firm, ascending or spreading, 3 to 7 cm. long, 3 to 7 mm. wide, rounded and papillose-ciliate at the base, glabrous or rarely sparsely villous on the upper surface, velvety puberulent beneath; panicles long-exserted, 3 to 8 cm. long, about three-fourths as wide, rather densely flowered, the axis appressed-pubescent, with tufts of long hairs in the axils, the branches ascending; spikelets 1.2 to 1.3 mm. long, 0.7 mm. wide, obovate-elliptic, densely papillose-pubescent; first glume about one-fourth the length of the spikelet, obtuse; second glume and sterile lemma equaling the fruit but not exceeding it; fruit 1.1 mm. long, 0.6 mm. wide, elliptic, slightly pointed. Autumnal form ascending, usually decumbent at base, at first sending out from the lower and middle nodes long branches similar to the vernal culms, later pro- ducing appressed, more or less fascicled branchlets, the flat or somewhat involute blades not greatly reduced. ’ The less copious pubescence and larger panicles of the type of P. parvispiculum prove not to be correlated. A specimen of Wash 467 cited in the description of P. leucothriz has one panicle 8 cm. long as in the type of P. parvispiculum, and six small * Fie. 209.—P. leucothriz. From type specimen. 206 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. ones like those in the type of P. leucothrix. The New Jersey specimens, Chase 3536, 3556, and 3578, as also Hitchcock 1163 and 1398, though small plants with small panicles, are as little pubescent as is the type of P. parvispiculum or even less so. DISTRIBUTION. Low pine lands, New Jersey to Florida and Mississippi; also in Cuba. New Jersey: Atsion, Chase 3536, 3556; Forked River, Chase 3578. Norta Carouina: Wilmington, Hitchcock 377. SoutH CAROLINA: Orangeburg, Hitchcock 14, 376, 1372, 1398. GeoraiA: Darien Junction, Small in 1895; Alapaha, Curtiss 6817 in part. Fioriwa: Jacksonville, Combs 6, Kearney 146; Washington County, Combs 672, 673; Chip- ley, Combs 551, 572, 617; Eus- tis, Hitchcock 800, 805, Nash Fia. 210.—Distribution of P. lewcothriz. 334, 467, 1338, 2075; Seminole, Tracy 7193 in part;¢ Warrenton, Tracy 8410. Mississippi: Ocean Springs, Tracy 43. Louistana: St. Tammany Parish, Cocks 286; Lake Charles, Hitchcock 1163. Cusa: Herradura, Hitchcock 554; without locality, Wright 3460 in part. 119. Panicum longiligulatum Nash. Panicum longiligulatum Nash, Bull. Torrey Club 26: 574. 1899. ‘‘Collected by Dr. Geo. Vasey, at Apalachicola, Florida, in 1892.’’ The type, in Nash’s herbarium, consists of two vernal culms with three autumnal culms of the preceding year attached. Elliott ® described this species under P. ‘‘nitidum? La Marck” as shown by the specimen so labeled in the Elliott Herbarium. DESCRIPTION. Vernal culms usually stout, 30 to 70 cm. high, erect, or ascending at base, glabrous; sheaths glabrous, usually much shorter than the internodes; ligules 2 to 3 mm. long; blades rather thick and firm, 4 to 8 cm. long, 4 to 8 mm. wide, glabrous on the upper surface, puberulent beneath, gradually narrowed to the sharp point, the lower ascend- ing, the upper spreading or often reflexed; panicles ovoid, 3 to 8 cm. long, two-thirds to three-fourths as wide, rather densely flowered, the slender branches usually stiffly ascending, short spikelet-bearing branchlets in the axils; spikelets 1.1 to 1.2 mm. long, 0.7 mm. wide, elliptic, pubescent; first glume one-fourth as long as the spikelet; second glume slightly shorter than the fruit and sterile lemma; fruit 1 mm. long, 0.7 mm. wide, elliptic. Autumnal culms more or less reclining, the branches spreading, usually somewhat recurved, with crowded branchlets and spreading, subinvolute, reduced blades about equaling the reduced panicles of few long-pediceled spikelets; winter rosette prominent, blades glabrous. Smaller, more slender specimens of this species resemble less pubescent specimens of P. leucothriz; these may be distinguished from that species by the glabrous culms and sheaths and slightly smaller spikelets with fruit exposed at the summit. Fig. 211.—P. longiligulat- um. From type speci- men. a Panicum longiligulatum and P. lindheimeri were also distributed under this number. b Bot. 8. C. & Ga. 1: 128. 1816. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 207 DISTRIBUTION. Low pine barrens and swamps of the Coastal Plain, North Carolina to Florida and Louisiana. Nortu Carourma: Roanoke Island, Chase 3213, 3228, 3233; Wards Mill, Chase 3175, 3179; Wilmington, Chase 3136, 3145, 3150, Hitchcock 372. Sourn Carouina: Orangeburg, Hitchcock 16; Hartsville, Coker in 1908. Georaia: Charlton County, Harper 1575; Sumter County, Harper 459; Bullock, Harper 839; Stone Mountain, Hitchcock 375. Fioripa: Jacksonville, Curtiss 4033; Baldwin, Hitchcock 988, 1005; Chipley, Combs 569; Lake City, Combs 115; Milton, Chase 4308; Apalachicola, Chapman; Bay Head, Combs 652; Santa Rosa, Combs 488, Tracy 8398, 8423; Warrington, Tracy 8413; Braidentown, Hitchcock 957, 958; Seminole, Tracy 7193 in part; Myers, Chase 4141, 4172, 4188, Hitchcock 875, 881, Lee Co. Pl. 473. Mississrerr: Biloxi, Chase 4366, Hitchcock 1068; Ocean Springs, Kearney 288 in part, 300 in part; Petit Bois Island, Tracy in 1898. Louisiana: Abita Springs, Cocks 422. Fig. 212. Distribution of P. longiligulatum. 120. Panicum wrightianum Scribn. Panicum strictum Bosc; Roem. & Schult. Syst. Veg. 2: 447. 1817, not R. Br. 1810. This is described in a note under P. barbulatum Michx., from a: specimen collected by Bose in ‘America sept.’’ and received from him under this name. The type, in the Munich Herbarium, is marked ‘‘Panicum strictum Bose. Carolina, comm. Bosc.’’ Two duplicates of it are in the Willdenow Herbarium, one bearing a new name bestowed by Panzer, but we can not find that this name has been published. Panicum minutulum Desv. Opusc. 87. 1833, not Gaud. 1826. The type, in the Desvaux Herbarium, consists of two plants beginning to branch. Panicum wrightianum Scribn. U. 8. Dept. Agr. Div. Agrost. Bull. 11: 44. /. 4. 1898. “Cuba (No. 3463, C. Wright, 1865).’? The type, in the National Herbarium, is the early autumnal form and is the large plant figured with the original description. Trinius described@ this species under the name ‘‘Panicum nodiflorum La M.,”’ citing P. strictum Bose as a synonym. The specimen in the Trinius Herbarium is marked ‘‘Carolina, Bosc. sub nom. P. ~ () strictum Bosc.’’ DESCRIPTION. es Vernal culms weak and slender, ascending from a decumbent ob ck oar ltd base, or rarely at first erect, 15 to 40 cm., or rarely 60 cm. high, specimen. minutely puberulent; sheaths striate, shorter than the inter- nodes, glabrous, except the summit and ciliate margin, or puber- ulent; ligules 2 to 3 mm. long; blades spreading, 2 to 4 cm. long, 3 to 5 or rarely 6 mm. wide, glabrous or puberulent beneath and minutely pilose above; panicles oblong-ovate, 3 to 6 cm. long, one-third to half as wide, the branches ascending, the minute spikelets long-pediceled; spikelets 0.95 to 1 mm. long, 0.5 mm. wide, @Gram. Pan. 241. 1826. 208 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. ellipsoid, turgid, subacute, pubescent; first glume about one-fourth as long as the spikelet; second glume shorter than the fruit and sterile lemma; fruit 0.8 mm. long, 0.5 mm. wide, subacute. Autumnal form decumbent-spreading, the culms sending out from the lower and middle nodes numerous ascending branches, becoming somewhat bushy branched, the flat or subinvolute blades and secondary panicles not much reduced. DISTRIBUTION. Along the margins of streams and ponds in sandy or mucky soil, southern New Jersey to Florida and west to Texas; also in Cuba. New Jersty: Bennett, Stone in 1909. Norra Carona: Wilsons Mills, Chase 3096; Wilmington, Chase 3135, Hitch- cock 373, 374, Kearney 246. GeoraiA: Hawkinsville, Biltmore Herb. 7080a (Biltmore Herb.). Froripa: Live Oak, Curtiss 6652; Madison County, Combs 290; De Funiak Springs, Combs 441, 477, Curtiss 5912; Monticello, Combs 347, 354; Washington County, Combs 552, 664; Pen- sacola, Combs 524; Grasmere, Combs 1068, 1087; Marianna, es — Tra cy 3644. Fia. 214.—Distribution of P. wrightianum. AtaBaMa: Mobile, Kearney 49 in part. Mississippi: Biloxi, Kearney 307; Horn Island, Tracy 2861; Petit Bois Island, Tracy 4611. Texas: Without locality, Wright (Gray Herb.). Cusa: Without locality, Wright 3463 in part. Lanuginosa.—Plants more or less pubescent throughout, usually conspicuously so; ligules densely hairy, 2 to 5 mm. long; blades not over 1 cm. wide, usually narrower; spikelets 1.3 to 3 mm. long, pubescent, the second glume and sterile lemma 5 to 7 or in the larger spikelets 7 to 9-nerved. Autumnal form usually freely branching, secondary leaves and panicles much reduced. These species were usually referred by the earlier American authors to P. pubescens Lam. or Michx. Spikelets not over 2 mm. long. Plants grayish, velvety-pubescent. Spikelets 1.4 to 1.5 mm. long; autumnal blades involute-pointed (see also P. albemarlense)....128. P. auburne. Spikelets 1.8 to 2 mm. long; autumnal blades flat. Plants dark or olive green when dry; spikelets 1.9 to 2 mm. long. Freely branching from lower nodes, decum- bent; vernal blades puberulent on och eurtaces:.. : .},.foaseeaeeeeeeee 130. P. olivaceum. Sparingly branching from middle nodes, erect; vernal blades sparingly pilose onupper-suxess yi! 5 ss 4 ee 129. P. thurowii. Plants light or yellow green when dry. Autumnal form prostrate, branching from base and lower nodes, forming close mats; blades not ciliate; around hot BPMINPS*! tose as cee ee ce = = een eee 135. P. thermale. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 209. Autumnal form ascending or spreading, branching from middle and upper nodes; the reduced, fascicled blades strongly ciliate. Culms 40 to 70 cm. high, autumnal culms usually 40 to 50 cm. long; southeastern U.8. 25.2225. .5204). 126. Culms 20 to 40 cm. high, autumnal culms usually 20 to 30 cm. long, the early branches zigzag; West Perit Bel yee Mae Gi ek 127. Plants pubescent, often villous, but not velvety. Culms conspicuously pilose with long, horizontally spreading hairs; branching before expansion of BIBNEy PaMICheR. oo. une oped 131. Culms variously pubescent, if pilose the hairs not long and horizontally spreading. Vernal blades glabrous or nearly so on the upper surface, firm in texture. Autumnal culms branching from the lower nodes, forming a spreading bunch 10 to 15 cm. high; Pacific slope......... 133. Autumnal culms branching from the middle nodes, forming widely spreading mats; Atlantic slope (see also form of P. hua- PiUCIE SACOM) S 2-52 o2 =. See oes eeeee 125. Vernal blades pubescent on upper surface, some- times pilose near base and margins only. Spikelets 1.3 to 1.5 mm. long; vernal blades long-pilose on upper surface. Autumnal form widely decumbent- spreading, forming a mat; vernal culms soon geniculate-spreading; plants olivaceous..-.......-.:.++- 122. Autumnal form erect or leaning, never forming a mat; plants yellowish- green. Axis of panicle pilose, panicle branches tangled, the lower GEONIIOE a 6a ates ge ie nr w= gn Axis of panicle puberulent only, panicle branches not tangled, the lower ascending. .-........ 121. Spikelets 1.6 to 2 mm. long; vernal blades pilose or pubescent. Upper surface of blades pilose; spike- lets 1.8 to 2 mm. long; autumnal form decumbent-spreading. Spikelets pointed; culms weak and EGER O Seles « DESCRIPTION. Vernal form cespitose, usually stiffly upright, light olivaceous, often purplish, harsh to the touch from the copious, spreading, papillose pubescence of culms and leaves; culms 20 to 60 cm. high; nodes bearded with spreading hairs; sheaths shorter than the internodes; ligules 3 to 4 mm. long; blades firm, stiffly erect or ascending, 4 to 8 cm. long, 6 to8 mm. wide, the veins inconspicuous, the upper surface copiously short-pilose, especially toward the base, the lower surface densely pubescent; panicle rather short- exserted until maturity, 4 to 6 cm. long, nearly as wide, rather densely flowered, the axis and often the branches pilose, the flexuous, fascicled branches ascending or spreading, short spikelet- bearing branchlets at base of the fascicles; spike- lets 1.6 to 1.8 mm. long, 1 mm. wide, obovate, obtuse, turgid, papillose-pubescent; first glume about one-third the length of the spikelet; second glume and sterile lemma subequal, scarcely covering the fruit at maturity; fruit 1.5 to 1.6 mm. long, 1 mm. wide, elliptic, obscurely apiculate. Autumnal form stiffly erect or ascending, the culms and sheaths sometimes papillose only, the branches fascicled, the reduced, crowded leaves ascending, the blades 2 to 3 cm. long, much exceeding the reduced panicles. This species is variable as to amount of pubescence and as to the stiffness of the leaves, and it intergrades with the following subspecies. A specimen collected by Havard at El Paso, Texas, is referred here, though it is an unusual form with wider blades and spreading habit suggesting P. lindheimeri. Fig. 221.—P.huachucae. From type speci- men in National Herbarium. DISTRIBUTION. Prairies and open ground, Maine to South Dakota and south to North Carolina and southern California. Marne: North Berwick, Parlin 1186, 1189. Vermont: Burlington, Hitchcock 133. Massacuusetts: Wellesley, Smith 737. Connecticut: Southington, Andrews 70; New London, Graves 4. New York: Vaughns, Burnham in 1897; Pavilion, Hill 182 in 1907; Westfield, Hill 171 in 1907; Jamaica, Bicknell in 1905; Hempstead, Bicknell in 1903; Woodmere, Bicknell in 1907. Ontario: Galt, Herriot 14; Niagara, Macoun 26337; Belleville, Macoun 29369; Long Point, Herriot 44. New Jersey: Netcong, Mackenzie 2075. @Scribner and Merrill, Rhodora 8 : 121. 1901; Nash in Britton, Man. 1040. 1901. bSee synonymy under P. tenwe Muhl., page 259. 916 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. PENNSYLVANIA: Easton, Porter in 1893. Ouro: Big Darby, Morris 9; Lancaster, Kellerman 6769; Steubenville, Kellerman 6785; Mount Gilead, Kellerman 6873; Vinton, Kellerman 6893. Inprana: Clark Junction, Bebb 520, Umbach 1816; Gibson, Hill 98 in 1908. Intrvo1s: Waukegan, Gleason & Shobe 324; Chicago, Hill 130 in 1905, Somes 210; Beach, Umbach 2237, 2244; Williamsfield, V. H. Chase 1858; Waucanda, Hill 217 in 1898. MicuicaNn: Detroit, Farwell 643b, 1382 in part; Howard Terrace, Wheeler in 1899. Wisconsin: Northwest Wisconsin, p Wood in 1889; Doherty Lake, Cheney 1107; Drummond, Cheney 4104; Madeline Island, Cheney 5638. Minnesora: Nicollet, Ballard in r 1892; Spring Lake, Ballard 544; Minneapolis, Sandberg 316; Center City, Sandberg 666; Thompson, Sandberg 385. Sout Daxora: Black Hills, Ryd- berg 1099. Iowa: Mount Pleasant, Mills in 1894; Fayette County, Fink 562. NEBRASKA: Ewing, Bates 1003; Thomas County, Rydberg 1368. Missouri: Allenton, Kellogg 16. Kansas: Manhattan, Hitchcock 2528, Kellerman in 1888. DELAWARE: Wilmington, Canby in 1898. District oF CoLtumMBiIA: Sudworth in 1889. Virernia: Ashland, De Chalmot. Kentucky: Harlan County, Kearney 58 in part; Lexington, Short 9 (Gray Herb.). Mississippi: Agency, Tracy 3190. ARKANSAS: Fulton, Bush 2332 (Mo. Bot. Gard. Herb.). Texas: El Paso, Havard in 1881. Montana: Without locality, Williams in 1887. Arizona: Huachuca Mountains, Lemmon 2907. CauirorniA: San Bernardino Mountains, Abrams 2737 (Gray Herb.). The Montana specimen, though from beyond the known range of this species, is fairly typical. Fig. 222.—Distribution of P. hwachucae. 124a. Panicum huachucae silvicola Hitche. & Chase. Panicum dichotomum fasciculatum Torr. Fl. North. & Mid. U.S. 145. 1824, not P. fasciculatum Swartz, 1788. ‘‘In sandy fields, New-Jersey.’’ The type, in Columbia University Herbarium, is a late autumnal specimen with tufts of short branches at the nodes. Panicum nitidum ciliatum Torr. Fl. North. & Mid. U. 8. 146. 1824, not P. ciliatum Ell. 1816. ‘‘In the pine-barrens of New-Jersey.”’? The type, in Columbia University Herbarium, is a single vernal culm. Panicum huachucae silvicola Hitche. & Chase in Robinson, Rhodora 10: 64. 1908. “Type, District of Columbia, Chase, no. 2400, in National Herbarium.’’ The speci- men is a small clump of vernal culms beginning to branch and with mature primary panicles. This is the form described by Scribner and Merrill@ under the name P. lanugi- nosum Ell. aU.S8. Dept. Agr. Div. Agrost. Circ. 29: 7. 1901. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 217 DESCRIPTION. Vernal form taller and more slender, brighter green and less densely pubescent than in P. huachucae; culms 30 to 75 cm. high, suberect or ascending, papillose-pilose with spreading hairs; nodes bearded with reflexed hairs, usually a glabrous ring below; sheaths papillose-pilose; blades thin, lax and spreading, 5 to 10 cm. long, 6 to 12 mm. wide, the veins inconspicuous; upper surface sparsely short-pilose or with copious long hairs toward the base; lower surface pubescent and with a satiny luster; panicles exserted, 5 to 8 cm. rarely 10 cm. long, nearly as wide, rather densely flowered, the axis pilose, the flexuous, fascicled branches spreading, with short spikelet-bearing branchlets at the base of the fascicles; spikelets 1.6 to 1.8 mm. long, 0.8 to 1 mm. wide, elliptic-obovate, at maturity subobtuse, pubescent with spreading hairs; first glume one-fourth to one-third the length of the spikelet, obtuse or subacute; second glume and sterile lemma subequal, slightly shorter than the fruit at maturity; fruit 1.5 mm. long, 0.8 to 0.9 mm. wide, elliptic, subacute. Autumnal form more or less decumbent, the numer- ous fascicled branches shorter than the primary inter- nodes, at least late in the season, the reduced spreading leaves sometimes nearly glabrous above except for a few long hairs near the base. The following specimens represent an extreme form with the upper surface of the blades nearly or quite glabrous, thus approaching P. tennesseense. They differ from that species in the thin, lax blades, with no marked white margin and without conspicuous veins. MaIne: Westbrook, Ricker 666; Orono, Fernald 503; Massacnusetts: Framingham, Smith 739; Connecticut: Franklin, Graves 166; RuopE IsLanp: Providence, Battey in 1886; New York: Ithaca, Ashe in 1898; New Jersey: Bear Swamp, Stone 3; PENNSYLVANIA: Easton, Porter in 1898; Germantown, Stone 5, 9; Iowa: Appanoose County, Fitzpatrick 38; District oF Cotumsia: Kearney 33, Steele in 1900; NortH Caroutna: Chapel Hill, Chase 3049, 3062, 3067; TENNESSEE: Polk County, Kearney 328; Texas: Ennis, Smith in 1897; OxtaHomA: Chelsea, Bush 1210. FiG. 223.—P. huachucae silvicola. From type specimen. DISTRIBUTION. Open woods and clearings, Maine to northern Florida, west to Michigan, Nebraska, and Arizona. Marne: Falmouth, Chamberlain 513; Brewer, Knight 51; South Berwick, Parlin 1181; Southport, Fernald 509. New Hampsarre: Langdon, Fernald in 1899., Vermont: Burlington, Jones in 1898; Brandon, Knowlton in 1882. Massacuusetts: Salem, Sears in 1883; Wellesley, Smith 735. Connecticut: Southington, Andrews 36, 64, 66; Groton, Graves 8; Waterford, Graves 156. RuoveE Isianp: Providence, Battey in 1886. New York: Sylvan Beach, Mazon 550; Oneida, Haberer in 1900, House 1136; Washington County, Burnham 14, 23; Gansevoort, Peck in 1897; Jamaica, Bicknell in 1904; Port Washington, Bicknell in 1908. Ontario: Galt, Herriot 53, 61, 93; Windsor, Macoun 26334. New Jersey: Clifton, Nash in 1892; South Amboy, Mackenzie 2159, 2169; Wild- wood, Chase 3508; Forked River, Chase 3580; Berkeley Heights, Mackenzie 2249; Milburn, Mackenzie 2137; Cranberry Lake, Mackenzie 2197. PENNSYLVANIA: Refton, Heller 4791; Mount Hope, Heller 4785; McCalls Ferry, Rose & Painter 8133; Easton, Porter in 1895. 218 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Onto: Columbus, Morris 48; Painesville, Werner in 1885; Berea, Ashcroft in 1897; Erie County, Moseley in 1902; Big Darby, Kellerman 6758; Vinton, Kellerman 6890, 6895. Inp1ANA: Lafayette, Dorner 84, 91, 93; Steuben County, Deam in 1903; Brazil, Somes 230. Intrnots: Downers Grove, Umbach 1820; Athens, Hall in 1861; Peoria, Brendel; Glasford, Wilcox 42; Princeville, V. H. Chase 81; Williamsfield, V. H. Chase 1851; Oregon, Waite in 1885; Mahomet, Gleason 1033; Makanda, Gleason 1028, 1030; Grand Tower, Gleason 1031. MicuicaNn: Port Huron, Dodge in 1909; Detroit, Farwell 597d, 1382 in part; Grand Beach Springs, Hill 83 and 85 in 1908. Iowa: Ames, Ball 42, 157; Lebanon, Ball & Sample 35; Decatur County, Fitz- patrick 37; Fort Dodge, Somes 207. NesrasKka: Pishelville, Clements 2983. Missourt: Courtney, Bush 734, 1718, 2996, 3968; Monteer, Bush, 746, 759, 760; Sibley, Bush 4002, 4803; Wodsen! Bush 4024; Vale, Bush 3915; Pleasant Grove, Bush 309; St. Louis, Eggert 124, 235, Hitchcock 599; Jefferson County, Lggert 244, 289. Kansas: Cherokee Ganty: Hitch- cock Pl. Kan. 882; Manhattan, EMitchcock 2523. DELAWARE: Centerville, Commons 289, 290, 292, 2938, 360, 364; Wilmington, Commons 362. MaryLaNnp: Chesapeake Beach, Chase 3254; Owings, Hitchcock 1621, 1624, 1630. District or CotumsBtiA: Chase 2400, Kearney 22, Vasey in 1882, Ward in 1881. Vireria: Alexandria County, Chase in Kneucker Gram. Exs. 551, Hitchcock 382, 383, Ward in 1879; Clifton Forge, Tidestrom 48. West Virainia: Fairmont, Hitchcock 136; Morgantown, Hitchcock 137. Nortu Carona: Chapel Hill, Ashe, Chase 3065, 3069. Groreia: Americus, Tracy 3889 in part; Augusta, Kearney 218 in part; Stone Mountain, Hitchcock 13574. Fiorrpa: Gainesville, Combs 751. TENNESSEE: Sherwood, Eggert 30, 245; Ducktown, Chambliss 23; Chester County, Bain 197. AtaBama: Sand Mountain, Biltmore Herb. 14879b (Biltmore Herb.). Mississippi: Jackson, Hitchcock 1306; Fairport, Tracy 3208; Macon, Tracy 3223; Starkville, Tracy 1751; Agency, Tracy 3198. ARKANSAS: Prescott, Bush 251; Benton County, Plank 49. Lourstana: New Orleans, Drummond 454; Shreveport, Hitchcock 1255; West Feliciana, Cocks 3509. Texas: Waller County, Hitchcock 1214, Thurow 22; Gillespie County, Jermy 57; without locality, Reverchon 1075; Texarkana, Heller 4084. Oxtanoma: Chelsea, Bush 1210 (Mo. Bot. Gard. Herb.). Arizona: Tucson, Towmey 781. Fia. 224.—Distribution of P. huachucae silvicola. 125. Panicum tennesseense Ashe. Panicum tennesseense Ashe, Journ. Elisha Mitchell Soc. 15: 52.1898. ‘‘Based on No. 7087 Biltmore Herbarium: Cedar glades, La Vergne Co., Tennessee.’’ The type specimen, in the Biltmore Herbarium, is the autumnal form, collected August 7, 1897. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 219 DESCRIPTION. Vernal form suberect or stiffly spreading, bluish green, often purplish; culms 25 to 60 cm. high, slender, papillose-pilose, or the upper portion glabrous; sheaths spread- ing-pubescent, rarely nearly glabrous; ligules dense, 4 to 5 mm. long; blades firm with a thin white cartilaginous margin, ascending or suberect, 6 to 9 em. long, 5 to 8 mm., rarely 10 mm., wide (the upper smaller), often sparsely ciliate at base, the veins usually conspicuous, the upper surface glabrous or with a few long, scattered hairs toward the base, the lower surface appressed-pubescent or nearly glabrous; panicle . 4 to 7 cm. long, nearly as wide, rather densely flowered, the lower branches ascending; spikelets 1.6 to 1.7 mm. long, 0.8 to 1 mm. wide, obovate-obtuse, turgid, pubes- cent; first glume about one-fourth the length of the spikelet; second glume shorter than the sterile lemma, leaving the summit of the fruit exposed at maturity; fruit 1.4mm. long, 0.8 mm. wide, elliptic, obtuse. Autumnal form widely spreading or decumbent, with numerous fascicled, somewhat flabellate, branches, often forming prostrate mats; leaves much reduced, the blades usually ciliate at base; winter rosette formed early. This species resembles P. lindheimeri and P. huachucae silvicola. From the former it differs in the larger spikelets, pilose sheaths, and more or less white-margined blades, which are often pubescent beneath, from the latter, in the firmer blades, glabrous above, and from both in the prostrate, mat-like autumnal form. Two vernal specimens from Connecticut, Graves 13 and 75 in 1899, are referred here doubtfully because of the looser panicle and rather numerous hairs on the upper surface of the blades. Two specimens with spikelets about 2 mm. long are referred doubtiully to P. tennesseense, one from Jefferson County, Missouri (Eggert 242) and one from Sapulpa, Oklahoma (Bush 711). . Fig. 225.—P. tennesseense. From type specimen. DISTRIBUTION. Open rather moist ground and borders of woods, Maine to Minnesota, and south to Georgia, Mississippi, and Arkansas; also in Colorado and Utah. Marne: Dover, G. B. Fernald 507; St. Francis, Fernald 166a; Fort Fairfield, Fernald 166; Cape Elizabeth, Chase 3457; Chesterville, Chase 3301; Fayette, Chase 3399; Hartford, Parlin 2017. New Hamesurre: Nashua, Robin- son 789 (Gray Herb.). VerRMoNT: Westmore, Eggleston 2181 (Gray Herb.). MAssaAcHUSETTsS: Framingham, Smith 741, 743. Connecticut: Hartford, Driggs 3; Preston, Graves 11; Branford, Bissell 5611. Rope Istanp: Providence, Collins in 1891 (Gray Herb.). New York: Thousand Islands, Ball 816, Robinson & Maxon 86; Ithaca, Coville in 1885; Apalachin, Fenno 13, 17; Ausable Chasm, Jones in 1898; Jamaica, Bicknell in 1905; Valley Stream, Bicknell, in 1905; Rosedale, Bicknell in 1904; Rockville Center, Bicknell in 1902; Edgemere, Bicknell in 1902; Hewlett, Bicknell in 1905. Ontario: Algonquin Park, Macown 72965. Fia. 226.—Distribution of P. tennesseense. 220 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. New Jersey: Wildwood, Chase 3503; South Amboy, Mackenzie 1459; Netcong, Mackenzie 2076. PENNSYLVANIA: Easton, Porter in 1892, 1895, and 1898; Lancaster County, Heller 4774, 4778; Germantown, Stone 8, 13; Safe Harbor, Small in 1889; Sayre, Barbour in Kneucker Gram. Exs. 4854; Rockdale, Pretz 2022. Oun10: Berea, Ashcroft in 1897. Inprana: Clark Junction, Bebb 2881; Indiana Harbor, Chase 1904; Anderson, Deam 2065. Inurnors: Zion City, Hill 141 in 1905. Micuican: Grand Beach Springs, Hill 86 in 1908; Petoskey, Hill 162 in 1878 (Hitchcock Herb.). Wisconsin: Racine, Wadmond 3424b; Webster, Cheney 3409; Stevens Point, Cheney 3471. Minnesota: Milaca, Sheldon 2743. Iowa: Fort Dodge, Somes 153. Nesraska: Minden, Hapeman in 1907. Missourr: Williamsville, Eggert 243; Swan, Bush 4532; Monteer, Bush 4684; Vale, Bush 3914. DELAWARE: Wilmington, Chase 3617, Commons 365. Mary.anp: Chesapeake Beach, Chase 3260; Potomac Valley a few miles above Washington, Chase 2463, 2849, 2874, 3274, 3275, 5424, 5425, Hitchcock 138, Kearney in 1897. District or CotumBiA: Ball in 1902, Kearney 29a, Hitchcock 505, Pollard 523. Viren: Fairfax County, Hitchcock 139; Clifton Forge, Tidestrom 5. West VIRGINIA: Quinnemont, Pollard & Maxon 22. NortH Carona: Asheville, Boynton 2; Hendersonville, Biltmore Herb. 5184b; Biltmore, Biltmore Herb. 698b; Macon County, Boynton 9. GeoraiA: Stone Mountain, Hitchcock 1358. Kentucky: Lexington, Peter in 1833 (Ky. State Univ. Herb.): TENNESSEE: Knox County, Kearney in 1893. AxaBAMA: Pisgah, Chase 4477; Scottsboro, Chase 4499. Mississippi: Panola County, Eggert 296. Arxansas: Texarkana, Heller 4160. OKLAHOMA: Sapulpa, Bush 712. Cotorapo: South Boulder, Jones 619. Urau: Springdale, Jones 6069. Arizona: Santa Catalina Mountains, Thornber 308 (Jones Herb.). 126. Panicum lanuginosum Ell. Panicum lanuginosum Ell. Bot. 8. C. & Ga. 1: 123. 1816. ‘‘Grows in Georgia. Sent to me by Dr. Baldwin.”’ The type, in the Elliott Herbarium, consists of a single culm lacking the base, with four leaves and primary panicle included at base; the spikelets, which are immature, are 1.8 mm. long, and 0.8mm. wide. The accompany- ing label reads: ‘‘Panicum Lanuginosum. Hab. Georg: Dr. Baldwin.” Panicum dichotomum lanuginosum Wood, Class-book ed. 3. 786. 1861. Presum- ably based on P. lanuginosum Ell., no synonymy nor locality being cited. Panicum orangensis[e] Ashe, Journ. Elisha Mitchell Soc. 15: 113. 1899. “I have collected the plant at two stations, both in Orange County, N. C.”’ ‘‘Collected in June, 1898.’’ No specimen bearing this name could be found in Ashe’s herbarium. There is, however, a cover containing specimens collected at Chapel Hill, Orange County, North Carolina, June 29, 1898. On this cover are notes which indicate that Ashe considered the species allied to P. lanuginosum. The description of P. orangense a This number in Mo. Bot. Gard. Herb. is P. hwachucae silvicola. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 221 agrees with these specimens except that the spikelets are said to be glabrous. But this statement is probably an error of observation or of description, since the author adds, ‘‘Related to Panicum lanuginosum Ell., and separated from it by having a longer, softer pubescence and its leaves not being ciliate.’? Since the spikelets of P. lanuginosum as described by Ashe (P. huachucae silvicola) are pubescent, this dif- ference would probably have been noted in the contrast of the two species. A por- tion of the specimen mentioned above has been deposited in the National Herbarium and has been chosen as the type of P. orangense Ashe. It is the early autumnal form. Panicum ciliosum Nash, Bull. Torrey Club 26: 568. 1899. “‘Type collected by S. M. Tracy, at Biloxi, Mississippi, September 1, 1898, no. 4580.’’ The type, in Nash’s herbarium, is the early autumnal form with a simple culm and primary panicle attached, and without the winter rosette. The specimen of Tracy 4580 in the National Herbarium has a winter rosette, the blades 4 to 6 cm. long. In the description the ligule is said to be ‘‘about 0.5 mm. long’’ but in the type it measures 3 mm. long. DESCRIPTION. Vernal form grayish olive green, velvety to the touch; culms tufted, usually in large clumps, 40 to 70 cm. long, slender, lax, spreading, densely villous with fine, soft hairs arising from small papille; nodes villous, often a glabrous ring below; sheaths shorter than the internodes, soft-villous like the culm, or the upper puberulent only, ciliate on the margin; ligules 3 to 4 mm. long; blades thickish but not stiff, ascending or spreading, somewhat incurved or spoon-shaped, 5 to 10 cm. long, 5 to 10 mm. wide (the uppermost much smaller), acuminate, narrowed toward the rounded base, the margins sometimes papillose-ciliate, the upper surface clothed with short, soft hairs with long soft hairs intermixed, especif~ly toward the margins and base, the lower surface densely velvety-pubescent; panicles exserted, 6 to 12 cm. long, about as wide, loosely flowered, the axis pubescent, the slender flexuous branches spreading or ascending, the lower often drooping; spikelets 1.8 to 1.9 mm. long, 1 mm. wide, obovate-elliptic, subobtuse, pubescent; first glume one-third the length of the spike- let, obtuse or obscurely pointed; second glume and sterile lemma equal, slightly shorter than the fruit at maturity; fruit 1.6 mm. long, 0.9 mm. wide, ellip- tic, subacute. Autumnal form widely spreading or decumbent, freely branching from the middle nodes, the branches repeatedly branching and much exceeding the internodes, the ultimate branchlets forming flabel- late fascicles; leaves and panicles much reduced, the flat blades almost always ciliate and exceeding the panicles; winter rosette not appearing until late, the blades 4 to 5 cm. long, usually ciliate, otherwise minutely velvety or nearly glabrous. The plant bears some resemblance in color and pubescence to P. scoparium, but is smaller and much more slender. The vernal form also resembles P. hwachucae silvicola but is larger and more velvety and is gray-green in color rather than bright green. It may be that the form described by Nash as P. ciliosum is a distinct species. It differs in having blades glabrous on the upper surface or with a few long hairs only, but not velvety, and winter rosettes of large blades. The typical form has been found only in Tracy’s garden, at Biloxi, in cultivated soil. Other Biloxi specimens lack the large rosettes, probably because not growing in cultivated soil. The following specimens, because of the lack of velvety pubescence on the upper surface of the blades, may be referred to this form: Mississippi: Biloxi, Chase 4331, Hitchcock 1079, Tracy 1735, 2867, 3620, 3622, 3645, 4580, 4605; Ocean Springs, Tracy 6469. Lovistana: Lake Charles, Hitchcock 1152, Chase 4401. Fic. 227.—P. lanuginosumg From type specimen. 222 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Moist sandy woods, mostly near the coast, New Jersey to Florida and Texas. New Jersey: Wildwood, Chase 3488, 3505, Heritage 6. DeLAWARE: Milton, Commons 342; Lewes, Hitchcock 387. MaryLanpD: Between Chesapeake Beach and Chesapeake Junction, Hitchcock 1613, 1638. Virernia: Dismal Swamp, Chase 3663, Tyler in 1905; Norfolk County, Kearney 1559; Virginia Beach, Kearney 2043; Cape Henry, Chase 5426. Norta Carouna: Chapel Hill, Ashe in 1898, Chase 3068, 3076; Raleigh, Chase 3086; Wilsons Mills, Chase 3107; Lake Mattamuskeet, Chase 3208; Scranton, Chase 32014; Roanoke Island, Chase 3221; Wilmington, Chase 4584, Hitchcock 388, 1468. SourH Carouina: Orangeburg, Mtchcock 389, 390, 1395; Isle of Palms, Hitchcock 386, Chase 4532; St. Helena Island, Cuth- bert in 1899. Groreia: Stone Mountain, J. D. Fig. 228.—Distribution of P. lanuginosum. Smith 48 in 1883; Millen, Cur- tiss 6827; Coney, Harper 1399; Burke County, Harper 765; Thomson, Bartlett 1443, 1460. Fioripa: Lake City, Bitting g 13, Chase 4277, 4292,.Combs 174, 194, Hitchcock 1032; Milton, Chase 4305, Curtiss R; Madison County, Combs 215, 294; Gainesville, Chase 4242, Combs 732; Eustis, Nash 375; Orange Bend, Chase 4113; St. Andrews, Tracy 9138. AtaBAMA: Fort Morgan, Tracy 8399. Mississippi: Biloxi, Chase 4331, Hitchcock 1079, Tracy 1735, 2867, 3620, 3622, 3645, 4580, 4605; Ocean Springs, Tracy 6469; Cat Island, Tracy & Lloyd 441; Horn Island, Tracy 2856; Jackson, Hitchcock 1297; Saratoga, Tracy 8416. Lourstana: Shreveport, Hitchcock 1238, 1258; Cameron, Cocks 2191; Calcasieu, Cocks 2193; Breton Island, Tracy & Lloyd 467; Alexandria, Ball 544; Lake Charles, Chase 4401, Hitchcock 1129, 1135, 1147, 1152. Texas: Silver Lake, Reverchon 1884. 127. Panicum acuminatum Swartz. Panicum acuminatum Swartz, Prodr. Veg. Ind. Occ. 23. 1788. ‘*Jamaica.”? In his Flora® Swartz states concerning this species, ‘‘Incolit campos arenosos Jamaicae montosae.’’ The type specimen, in the Swartz Herbarium, consists of three plants of the prostrate autumnal form. Panicum dichotomum acuminatum Swartz; Griseb. Fl. Brit. W. Ind. 553. 1864. Based on P. acuminatum Swartz. In the Grisebach Herbarium is a plant of this species labeled by Grisebach, collected in Jamaica by March. Panicum comophyllum Nash, Bull. Torrey Club 30: 380. 1903. ‘‘Type collected in rich soil at Santurce [Porto Rico], January 9, 1899, by Heller, no.12.’’ The type, in the herbarium of the New York Botanical Garden, is a specimen in the early branch- ing state. @ Fl. Ind. Occ. 1: 152. 1797. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 223 DESCRIPTION. Vernal culms leafy, ascending from a geniculate base, 20 to 70 em. high, densely villous with soft, spreading hairs, rarely glabrate above, the nodes more or less bearded; sheaths velvety papillose villous or the upper glabrate; ligule 2 to 3 mm. long; blades ascending or spreading, 4 to 8 cm. long, 6 to 13 mm. wide, lanceolate, slightly cordate at . base, sharply acuminate, usually ciliate, the lower sur- face velvety papillose puberulent, the upper surface from appressed papillose pubescent to long-villous, or nearly glabrous except for long hairs near the base or margin; panicles 3 to 10 cm. long, about as wide, the axis usually villous, the branches flexuous, the lower spreading or even reflexed; spikelets 1.8 to 1.9 mm. long, 0.9 mm. wide, obovate, turgid, abruptly subacute, pilose; first glume about one-third the length of the spikelet, subacute; second glume and sterile lemma barely equaling the fruit at maturity; fruit 1.3 to 1.4 mm. long, 0.9 mm. wide, elliptic, abruptly acute. Autumnal form appearing early, the primary culms branching ai all but the upper- most nodes before the maturity of the primary panicles, these branches often exceed- ing the culm, more or less zigzag, repeatedly branching, the ultimate branchlets in dense, short, flabellate fascicles, the reduced blades flat or involute-pointed, the long hairs on the margins and upper surface usually conspicuous. Fig. 229.—P. acuminatum. From type specimen. DISTRIBUTION. Sandy pine woods, the West Indies; also in the United States of Colombia. Cusa: Herradura, Hitchcock 140, Tracy 9078; Pinar del Rio, Palmer & Riley 447; Wright 3874; Isle of Pines, Curtiss 307, 328, Palmer 4d: Riley 989, 1083, A. A. Taylor in 1901. Jamaica: Swartz, Hart 736. Santo Dominco: Poiteau (Paris Herb.). Porro Rico: Santurce, Heller in 1903, Maricao, Sintenis 355; Fajardo, Sintenis 1224 in part; Lares, Sintenis 5908. Cotomsia: Near Jamundi, Pittier 932, 982a. 128. Panicum auburne Ashe. Panicum auburne Ashe, N. C. Agr. Exp. Sta. Bull. 175:115. 1900. ‘‘Auburn, Ala., May 7, 1898. Collected by Professors F. S. Earle and C. Baker, of the Alabama Biological Survey, at Auburn, Ala., May 7, 1898. No. 1527.’’ The type specimen, in Ashe’s herbarium, consists of several immature vernal culms with portions of the dead autumnal culms of the pre- ceding year attached. DESCRIPTION. Vernal form grayish velvety-villous throughout; culms tufted, 20 to 50 cm. high, geniculate at base, widely spread- ing, soon becoming branched and decumbent, rather slender, densely papillose silky villous below, velvety with copious silky hairs intermixed above; sheaths usually about half the length of the internodes, villous like the culms; ligules 3 to 4 mm. long; blades rather thin, ascending, 3 to 7 em. long, 3 to 5 mm. wide, acuminate, slightly narrowed toward the base, the upper Fic. 230.—P. auburne. From type specimen. 294 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. surface velvety with copious long, silky hairs intermixed, especially toward the base, the lower surface silky-villous or velvety, the nerves somewhat conspicuous; paniclesshort- exserted, 3to5 cm. long, about as wide, the axis velvety, with long, silky hairs inter- mixed, the flexuous branches ascending or spreading; spikelets 1.3 to 1.4 mm. long, 0.8 to 0.9 mm. wide, obovate, very turgid, densely papillose-pubescent; first glume one-third to half the length of the spikelet, acute; second glume and sterile lemma equal and covering the fruit at maturity; fruit 1.1 to1.2 mm. long, 0.8 mm. wide, obovate-elliptic, minutely pointed. Autumnal form early becoming diffusely branched at all the nodes, prostrate- spreading, forming large mats, the branches curved upward at the ends; earlier branches longer than the primary internodes, the ultimate branchlets in short fascicles with involute-pointed blades 1 to 2 cm. long, the numerous turgid little spikelets clustered at their bases; winter rosette appearing rather late, the lanceolate blades silky-villous like those of the primary culm. The verflal form resembles that of P. lanuginosum but is smaller, more slender and more silky-villous, with smaller, more turgid spikelets; the prostrate autumnal form with upturned branch tips is characteristic. DISTRIBUTION. Sandy pine and oak woods of the Coastal Plain from Virginia to Florida, and west to Louisiana. Vireinia: Cape Henry, Chase 2341; Virginia Beach, Mackenzie 1733, Walliams 3097, 3105; Dismal Swamp, Chase 3680. NortH Caronina: Wilmington, Chase 3132, 4579, Hitchcock 1482; Cumberland County, Stevens 6425. SoutH Carotina: Orangeburg, Hitchcock 10. GeroretA: Bainbridge, Curtiss 6811; Thomson, Bartlett 1172. Froripa: De Funiak Springs, Combs 440. ALABAMA: Gateswood, Tracy 8430 in part; Auburn, Farle & Baker 1527. Lourstana: Shreveport, Cocks 3506, Fia. 231.—Distribution of P. auburne. 129. Panicum thurowii Scribn. & Smith. Panicum thurowti Scribn. & Smith, U.S. Dept. Agr. Div. Agrost. Cire. 16: 5. 1899. ‘“‘Named for Mr. F. W. Thurow, by whom it was collected in Waller County, Texas, June 5, 1898, No. 9.” The type, in the National Herbarium, consists of one simple culm and one beginning to branch, about 40 cm. high, with short-exserted, nearly mature panicles, DESCRIPTION. Vernal form bluish green, but drying olive; culms tufted, 35 to 70 cm. high, erect or ascend- ing, villous, the nodes bearded with spread- ing hairs, usually a glabrous ring below; sheaths long, the lower often overlapping, the upper shorter than the internodes, sparsely or rather densely villous; ligules 4 mm. long; blades rather stiff, ascending or spreading, 7 to 12 cm. long, or the uppermost only 2 to 3 cm. long, 6 to 10 mm. Fig. 232.—P. thurowti. From type specimen. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 225 wide, acuminate, often somewhat involute toward the apex, narrowed toward the rounded base, the upper surface sparingly pilose toward the base and margins, the lower surface densely velvety-villous; panicles short-exserted, 7 to 11 cm. long, nearly as wide, rather densely flowered, the axis sparingly villous near the base, the branches spreading; spikelets 2 mm. long, 1 mm. wide, elliptic, somewhat obovate at maturity, obtuse, pubescent with soft, spreading hairs; first glume one-fifth the length of the spikelet, obtuse or obscurely pointed; second glume and sterile lemma equal, scarcely equaling the fruit at maturity, obtuse or slightly pointed; fruit 1.7 mm. long, 1 mm. wide, elliptic, subacute. Autumnal form erect, after the maturity of the primary panicle bearing at the mid- dle nodes a few appressed or ascending fascicled branches scarcely longer than the primary internodes, the reduced blades flat or somewhat involute at the tips, ciliate. DISTRIBUTION. Prairies and dry open woods, Ala- bama to Texas. AxaBama: In the vicinity of Mo- bile, Mohr in 1895 and 1897. Lovistana: Without locality, Hale (Gray Herb.). Texas: Waller County, Hitchcock 1171, 1195, 1226, Thurow 9, 11; Montgomery County, Thurow in 1905; Hockley, Thurow in 1893 and 1906; Swan, Reverchon 4163; Houston, Ravenel in 1869; Del Rio, Plank 41; without locality, Nealley in 1884 and 1887. Fia. 233.—Distribution of P. thurowii. 130. Panicum olivaceum gp. noy. DESCRIPTION. Vernal culms olive green, erect or somewhat spreading at base, 20 to 40 cm. high, velvety-villous with short hairs, the nodes bearded; sheaths villous like the culm, mostly shorter than the internodes; ligules 3 to 4 mm. long; blades rather stiffly erect or ascending or some of the lower spreading, 4 to 7 cm. long, 5 to 8 mm. wide (the uppermost erect, 1 to 3 cm. long), puberulent on both surfaces, also more or less short-villous above, and often with longer villous hairs toward the base; panicles 3 to 7 cm. long, ovate, the flexuous branches spreading, short spikelet-bearing branchlets in the axils; spike- lets 1.9 to 2 mm. long, 1 mm. wide, obovate, sub- acute, papillose-pilose; first glume one-fourth to one- third the length of the spikelet, usually pointed; second glume scarcely equaling the fruit and sterile lemma; fruit 1.6 mm. long, 1 mm. wide, subacute. Autumnal form upright or becoming decumbent- Fig. 234.—P. olivaceum. From : é : type specimen. spreading, freely branching from the lowerand middle nodes before the maturity of the primary panicle, the reduced branches appressed, or in the decumbent culms curved upward; blades reduced, flat, 1 to 2 cm. long, 2 to 4 mm. wide, usually conspicuously ciliate. Type U. S. National Herbarium no. 823209, collected February, 1888, at Coban, Department of Alta Vera Paz, Guatemala, altitude 1,400 meters, by H. von Tuerck- 41616°—vo1 15—10——15 226 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. heim (no. 428). It consists of four plants with mature primary panicles and freely branching culms. This species is closely allied to P. acuminatum, differing in the olivaceous color, the less velvety pubescence, the stiff, appressed blades, and the larger spikelets. The autumnal form is bushy, the branches evenly distributed, not gathered into distinct fascicles as in P. acwminatum. DISTRIBUTION. Gravelly banks and cultivated fields, Mexico to Costa Rica; also in Venezuela. Mexico: Jalapa, Pringle 8339 in part? (Nat. Herb. no. 823271); Orizaba, Bour- geau 2383 in part, Bottert 99, 101 (both in Brit. Mus. Herb.), 1987 (Paris Herb.); Minatitlan, J. G. Smith 571 (Hitchcock Herb.). GUATEMALA: Coban, Tuerckheim 428 in 1879, 428 in 1888; Seler 3235 (Berlin Herb.). Costa Rica: San Pedro de la Calabaza, Tonduz 10745 in part (Nat. Herb. no. 385918); Tablazo, Tonduz 7944. VENEZUELA: Tovar, Fendler 16388. Cotoms1a: Popayadn, Lehmann 974 (Gray Herb.). 131. Panicum praecocius Hitchc. & Chase. Panicum praecocius Hitche. & Chase, Rhodora 8: 206. 1906. ‘‘Type V. H. Chase 649; dry bank, near Wady Petra, Stark County, [linois, June 30, 1900, collected by Virginius H. Chase.’’ The type, in the National Herbarium, is a clump of branching culms, with mature secondary panicles, the primary ones being devoid of spikelets. DESCRIPTION. Vernal culms tufted, 15 to 25 cm. high, early branching and elongating, sometimes to 30 or 45 cm., at first erect, soon becoming geniculate and spreading, very slender, wiry, abundantly papillose-pilose with weak spreading hairs 3 to 4 mm. long; sheaths, even the lowest, much shorter than the very long internodes, those of the branches usually but 1 to 2 cm. long, pilose like the culm, more prominently papillose; ligules 3 to 4 mm. long; blades rather firm, erect or ascending, 5 to 9 cm. long, 4 to 6 mm. wide, the margins parallel about two-thirds their length, acuminate, long-pilose on both surfaces, the hairs of the upper surface 4 to 5 mm. long, erect from the plane of the blade, the under surface prominently papillose; panicles at first usually overtopped by the upper leaf, but at or past maturity exserted, 4 to 6 cm, long, about as wide, loosely flowered, the axis pilose, the branches flexuous, spreading or ascending; FG, 235;-—-Peraecocius, From spikelets 1.8 to 1.9 mm, long, 1 mm, wide, obo- type specimen. vate, turgid, obtuse, pilose; first glume one-third to half the length of the spikelet, triangular; sec- ond glume and sterile lemma subequal, the glume slightly shorter than the fruit at maturity; fruit 1.6 mm. long, 1 mm. wide, broad-elliptic. Autumnal form ascending from a geniculate base, or in prairie sod erect, forming close bunches 10 to 20 cm. high, the upper portion of the primary culms early decidu- ous, the branches appressed, the scarcely reduced blades erect or narrowly ascending, much exceeding the reduced panicles; winter rosette appearing late, the blades 2 to 3 cm. long, long-pilose. This species scarcely has a simple state, the branches appearing often before the first panicle is expanded. 4See footnote under P. multirameum, page 185. a HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 227 DISTRIBUTION. Dry prairies and clearings, Michigan and Indiana to Minnesota and Texas. InprAna: Hessville, Hill 49 in 1909. Inurnors: Joliet, Hill 37 in 1907; Wady Petra, V. H. Chase 472, 649, 1212, 1214, 1218, 1492, 1515; Marshall County, V. H. Chase 1791; Williamsfield, V. H. Chase 1850; Havana, Gleason 1034. MicuicaNn: Port Huron, Dodge in 1909. Wisconsry: Lauderdale, Bebb 2057. Mrynesorta: Itaska Lake, Sand- berg 1016. Iowa: Fort Dodge, Somes 25; Armstrong, Cratty in 1890; Iowa City, Somes 246. — Nesraska: Broken Bow, Webber4. Fig. 236.—Distribution of P. praecocius. Missourtr: Monteer, Bush 748, 749 in part; McDonald County, Bush 87; Howell County, Bush 54; Lees Sum- mit, Bush 3090, 3935. Kansas: Manhattan, Carleton in 1892, Hitchcock 2500, 2524, 3853, Kellerman 20; Belleville, Hitchcock 3544. Texas: Waller County, Thurow 5, and in 1906; Weatherford, Tracy 7943 in part. Ox.aHomaA: Stillwater, Hitchcock in 1903 (Hitchcock Herb.). 132. Panicum subvillosum Ashe. Panicum subvillosum Ashe, Elisha Mitchell Soc. 16: 86. 1900. ‘‘Collected by the writer at Carlton, Minnesota, in August, in the simple state. Type material pre- served in my herbarium.’’ The type specimen, in Ashe’s herbarium, consists of three tufts of several culms each, 15 to 30 cm. high, with leaves clustered at the base and long-exserted mature panicles. Panicum unciphyllum forma pilosum Scribn. & Merr. Rhodora 3: 124. 1901, not Panicum pilosum Swartz, 1788. ‘‘Dry woods, Orono, Maine, 501 M. L. Fernald, July 7, 1891.’ The type, in the National Herbarium, consists of a tuft of nine slender culms 15 to 35cm. high, with long-exserted nearly mature panicles. DESCRIPTION. Vernal culms tufted, 10 to 45 cm. high, slender, ascending or spreading, pilose with ascending hairs, usually faintly papillose, the lower internodes short, thus making the plant more leafy below, the nodes short-bearded; sheaths sparsely pilose with ascending hairs, the lower overlapping, the upper much shorter than the long internodes; ligules 3 mm. long; blades rather firm, ascending, 4 to 6 cm. long, 4 to 6 mm. wide, rarely wider, acuminate, slightly narrowed toward the base, both surfaces pilose, the hairs on ee en the upper surface 3 to 5 mm. long, shorter on the type specimen. lower; panicles long-exserted, ovate to oblong in outline, 3 to5 em. long, two-thirds to three-fourths as wide, rather densely flowered, the axis pubescent or toward the base pilose, the lower branches ascending; spikelets 1.8 to 1.9 mm. long, 0.9 mm. wide, elliptic, obtuse, pubescent; first glume nearly or quite half the length of the spikelet, acuminate; 228 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. second glume scarcely equaling the fruit at maturity; fruit 1.5 mm. long, 0.8 mm. wide, elliptic, obtuse. Autumnal form widely spreading, sparingly branching from the lower nodes, the leaves and panicles not greatly reduced, the panicles overtopped by the leaves, these less copiously pilose. This species may be distinguished from P. implicatum and P. nieritienale by the larger spikelets, the long-exserted panicles, the aggregation of the leaves toward the base of the vernal culms, and the sparingly branched, almost prostrate autumnal form. In its most characteristic form the panicle branches are strictly ascending at maturity and spikelet-bearing near the ends only, thus forming a compact panicle with a long naked base. DISTRIBUTION, Dry woods and sandy ground, Nova Scotia to Connecticut, and west to Minnesota and northern Indiana. Nova Scotia: Bedford, Macoun 29368. New Brunswick: Kent County, Fowler in 1875. QueseEc: Montmorenci Fails, Macoun 69205 (Gray Herb.). Marne: Chesterville, Chase 3278, 3320; Fayette, Chase 3391; Cape Elizabeth, Chase 3453; Stacyville, Knight 56; North Yarmouth, Cham- berlain 837; Hartford, Parlin 2016; Cumberland, Chamber- lain 787, Ricker 12774; Orono, Fernald 501; Ogunquit, Parlin 1581; Canton, -Parlin 2001. New HampsuHireE: Wiers, Carter in 1902 (Hitchcock Herb.). Vermont: Rutland, Eggleston 1758. Massacuuserts: Ipswich, Oakes Fig. 238.—Distribution of P. subvillosum. (Gray Herb.). Connecticut: Tolland, Bissell 12001. New York: Verona, Haberer in 1900; Hempstead, Bicknell in 1903; Valley Stream, Bicknell in 1905; Rosedale, Bicknell in 1904. OntaARIO: Galt, Herriot in 1898; Algonquin Park, Macoun 22023. InpiANA: Clark Junction, Bebb 2832, 28334. MicuicaNn: Keweenaw County, Farwell 642. Wisconsin: Conover, Cheney 678; Tomahawk Lake, Cheney 1082. Minnesora: Carlton, Ashe in 1899. 138. Panicum occidentale Scribn. Panicum occidentale Scribn. Rep. Mo. Bot. Gard. 10:48. 1899. Based on “P. pubescens [Lam. misapplied by] Presl, not Lam. nor Michx.’’ While the type must be the specimen in Presl’s herbarium, Scribner’s conclusions were based on a dupli- cate in the Bernhardi Herbarium at the Missouri Botanical Garden, labeled in Presl’s handwriting ‘‘Panicwm pubescens Michx.’”’ The type specimen collected by Haenke, which is the basis of Presl’s @ identification, is in the Bohemian Museum at Prague and consists of three culms with mature primary panicles, and with secondary panicles on short branches from the lower nodes, that is, vernal culms showing the commence- ment of the autumnal form. One label reads ‘‘ Panicum pubescens Michx.;’’ another bears the locality ‘‘Archipel,” which refers to the vicinity of Nootka Sound, Van- couver Island, the locality as published by Presl, ‘‘Hab. in Nootka-Sund.”’ ,, @ Rel. Haenk, 1:306. 1830. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 229 DESCRIPTION. Vernal form tufted, yellowish green; culms slender, 15 to 40 cm. high, rarely higher, spreading, the lower internodes usually short, as in P. subvillosum, producing a leafy base as in that species, sparsely papillose-pubescent, the upper more or less elongated, glabrate, the nodes pubescent; sheaths rather sparsely papillose-pubescent, rarely almost glabrous; ligules 3 to4 mm. long; blades firm, erect or ascending, 4 to 8 cm. long, 5 to 7 mm. wide, acuminate, rounded at the base, the upper surface with a few long hairs toward the base and margin, otherwise glabrous, the under surface ap- pressed-pubescent; panicles long-exserted, 4 to 7 em. long, about two-thirds as wide, rather loosely Ee a flowered, the flexuous branches ascending or spread- specimen at Prague. ing; spikelets 1.8 mm. long, 1 mm. wide, elliptic- obovate, subacute, pubescent; first glume one-fourth the length of the spikelet or less, obtuse or pointed; second glume and sterile lemma as long as the fruit at maturity; fruit 1.6 mm. long, 0.9 mm. wide, elliptic, subacute. Autumnal form branching from the lower nodes, forming a spreading bunchor tussock 10 to 15 cm. high; leaves and panicles reduced; winter rosette appearing late, the blades narrowly lanceolate, glabrous or pilose at base. This species is less pubescent than any other in this group. DISTRIBUTION. Peat bogs and moist sandy ground, British Columbia and Idaho to southern California. Ipano: Lake Coeur d’ Alene, Sand- berg, Heller & MacDougal in 1892; Priest Lake, Piper 3778. WasHINGTON: Montesano, Heller 3978; Chelan County, Whited in 1901; Lake Chelan, Elmer 489, Lake & Hull 118; Bingen, Suksdorf 5162, 5174; Granville, Conard 378, Yakima County, Cotton 736, 792. British CotumsBia: Lake Osoyoos, Macoun 77229; Vancouver Is- Fia. 240.—Distribution of P. occidentale. land, Canby 352 (Gray Herb.). OreEGon: Mount Scott, Sheldon in 1902; Columbia River, Sheldon 8706. CALIFORNIA: Crescent City, Davy 5971; Mendocino, Davy 6092, MeMurphy, 425; New York Falls, Hansen 1723; Yosemite Valley, Brewer 1646; Merced River, Torrey 587; San Diego, Orcutt 540; without locality, Bridges 366, Hartweg 2024. 134. Panicum pacificum sp. noy. DESCRIPTION. Vernal form light green; culms tufted, 25 to 50 cm. high, ascending or spreading, leafy (culm leaves 5 or 6), papillose-pilose with spreading hairs, the nodes shortly spreading-pilose; sheaths papillose-pilose, the papillee prominent; ligules 3 to 4 mm, long; blades erect or ascending 5 to 10 cm. long, 5 to 8 mm. wide, acuminate, nar- rowed toward the rounded base, the upper surface papillose-pilose, typically with short hairs intermixed, but these often wanting and the long hairs sometimes sparse, 230 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, the lower surface appressed papillose pubescent; panicles usually rather short- exserted, 5 to 10 cm. long, about three-fourths as wide, the flexuous branches ascend- ing; spikelets 1.8 to 2 mm. long, 1 to 1.1 mm. wide, obovate, obtuse, turgid, papillose- ‘pubescent; first glume one-fourth to one-third the length of the spikelet, truncate; second glume and sterile lemma equaling the fruit at maturity; fruit 1.6 mm. long, 1 mm. wide, elliptic-obovate, obtuse or obscurely pointed. Autumnal form prostrate-spreading, repeatedly branching from the middle and upper nodes after the maturity of the primary panicle, the reduced blades less pilose than the vernal ones, exceeding the reduced panicles; winter rosette appearing rather early, blades only sparsely pubescent. Type U. 8. National Herbarium no. 592751, collected August 3, 1908, in moist places in woods, one-fourth mile east of hotel, Castle Crag, Shasta County, California, by A. 8S. Hitchcock (no. 3070). The specimen is a tuft of several branching culms. This species is distinguished from P. occidentale by the more copious pubescence throughout, the more leafy culms, and in the autumnal form by the branching habit; from P. thermale by the taller, late-branching culms, longer, narrower. blades, and pilose, not velvety, pubescence, also by the branching habit. It most nearly resem- bles P. huachucae, laxer forms resembling its subspecies silvicola, and like these it is variable in amount of pubescence. The spreading habit and larger spikelets, together with a distinct range, make it impossible to include this western form under P. huachucae. Fic. 241.—P. pacijicum. From type specimen. DISTRIBUTION. Sandy shores and slopes, and moist crevices in rocks, ascending to 1,650 meters, British Columbia and Idaho to southern California. Ipauo: Salmon River, Henderson 3569; Lake Coeur d’ Alene, Hitchcock 2171, 2190, Leiberg 1312; Lochsa River, Piper 4056; Sawtooth National Forest, Tide- strom 2636. Wasuineton: Falcon Valley, Suksdorf 124; Wenatchee, Whited 1249; Kittitas Sounty, Sandberg & Leiberg 425; Spokane, Kreager 160; Lake Calispell, Kreager 325; Lake Chelan, Gorman 635; Klickitat County, Suksdorf 6292. British CoLtumMBIA: Vancouver Island, Canby 252, Rosendahl & Brand 107, Waldron 1921. OreGon: Grants Pass, Piper 6493; Belknap Springs, Gorman 1834; without locality, Hall 671 (Gray Herb.). Nevapa: Ruby Valley, Watson 1350. Arizona: Lowell, W. F. Parish 263. : CatirorniA. Castle Crag, Hitchcock 3070, 3071, 3073, 3077; Red- ding, Heller 7856; Yosemite Valley, Bolander 4840, Hall & Babcock 3317, 3362, Hitchcock 3214, 3219, 3232, 3233, 3234; Crow Point, Hansen 1444; Clinton Bar, Hansen 1381; Pine Grove, Hansen 626; North Fork, Griffiths 4438, 4476, 4617; Madera, Griffiths 6586; San Jacinto Mountains, Hall2244; Pine Ridge, Hall & Chandler 239; Santa Cruz, Jones 2294; Point Reyes, Davy 6745, 6780; Requa, Davy & Blasdale 5894; San Bernardino Mountains, S. B. & W. F. Parish 1663; without locality, Bolander 564, Hartweg 2024 (Gray Herb.). Fia. 242.— Distribution of P. pacificum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 231 135. Panicum thermale Boland. Panicum thermale Boland. Proc. Calif. Acad. 2: 181. 1862. ‘On hot rocks and in hot water flowing from the Geyser springs and Geyser mountains, in the northern part ef Sonoma County,’’ California. The type, in the Gray Herbarium, is the early branching form. It is marked ‘“‘I call this: Panicum thermale till I shall know better. It grows in the Geysers Sonoma Co. and on hot rocks.”’ DESCRIPTION. Vernal culms grayish green, densely tufted, velvety-villous, 10 to 30 cm. high, ascending or spreading, the nodes with a dense ring of short hairs; sheaths often over- lapping, velvety-villous; ligules 3 mm. long; blades thick, ascending oy spreading, 3 to 8 cm. (mostly about 5 cm.) long, 5 to 12 mm. wide, acuminate, rounded or sub- cordate at base, both surfaces densely velvety-villous; panicles exserted or in high alpine specimens partly included, 3 to 6 cm. long, about as wide, densely flowered, the axis villous, the flexuous branches spreading, often drooping; spikelets 1.9 to 2 mm. long, 1 mm. wide, obovate-oblong, obtuse, turgid, papillose-pilose; first glume about one- third the length of the spikelet, obtuse or abruptly pointed; second glume and sterile lemma sub- equal, the glume shorter than the fruit at ma- turity; fruit 1.7 mm. long, 1 mm. wide, elliptic, subobtuse. Autumnal form widely spreading, the branches appearing even before the primary panicles are exserted, repeatediy branching, the whole form- ing a dense cushion, the blades and panicles of the ultimate branchlets reduced; winter rosette appearing early, the blades ovate-lanceolate, usually less pubescent than those of the culms. ' In the original description Dr. Bolander states: ‘‘The whole plant is like velvet to the feel. There are, however, some specimens which are rather smooth.’’ This smoother form is represented by part of Merrill 157, one tuft of which has lower blades nearly glabrous, but sheaths and upper blades nearly as velvety as in the type, while other specimens of this collection are fully as villous. The Bolander type collection represents about the average of the species. Some of the specimens cited below are longer villous than the type and some few are smoother. A specimen from Banff, Alberta, McCalla 2318, ‘‘on tufa and old bogs in warm sulphur stream; alt. 4,500 ft.,’’ has short, early-branching culms, broad leaves and small panicles like P. ther- male, but the pubescence is of sparser long hairs, somewhat harsh and promi- nently papillose as in P. pacificum. Fic. 243.—P. thermale. From type specimen. pases DISTRIBUTION. . Wet saline soil in the immediate vicinity of geysers and hot springs ascending to 2,500 meters, Alberta to Wyoming and California. Avperta: Banff, McCalla 2318. Montana: Lo Lo Hot Springs, Williams & Griffiths 306. : Wyromina: Yellowstone National Park, Chase 5252; Hitchcock 1902, 2061, 2086, Mearns 3061, 4050, 4166, 4203, 4789, 4870, 4983, 5064, 5110, 5134, Merrill 157, Fic. 244.—Distribution of P. thermale. 232 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 164, 165, A. Nelson 6174, A. Nelson & E. Nelson 6037, Rydberg & Bessey 3545, 3547, Tweedy 580; Bighorn County, Tweedy 94. CALirorNiA: Sonoma County, Bolander 3941; Napa County, Brewer 861; Lassen Peak, Bolander 2169. * 186. Panicum languidum nom. nov. Panicum unciphyllum forma prostratum Scribn. & Merr. Rhodora 3: 124. 1901, not P. prostratum Lam. 1791. ‘‘South Berwick, Maine, M. L. Fernald, September 26, 1897.” The type, in the National Herbarium, is a lax, decumbent, autumnal speci- men with geniculate nodes, numerous loose branches with immature panicles, and pointed spikelets 2 mm. long. DESCRIPTION. Vernal form tufted; culms 25 to 40 cm. high, weak, slender, ascending or spreading, pilose; sheaths shorter than the internodes, papillose-pilose; ligules about 3 mm. long; blades thin, lax, ascending or spreading, 4 to 7 cm. long, 4 to 9 mm. wide, acumi- nate, slightly narrowed to the rounded base, sparsely pilose on the upper surface, minutely appressed-pubescent beneath, usually with long hairs intermixed; panicles rather long-exserted, 3 to 6 em. long, two-thirds to three-fourths as wide, loosely flowered, the very flexuous branches finally spreading or drooping, the spikelets on long, mostly divaricate, flexuous pedicels, the axis and branches sparsely long-pilose; spikelets 2 mm. Fie. 245.—P. languidum. From type ono 1 mm. wide, elliptic, acute, pilose; first specimen of P. wnciphyllwm forma pro- A : ; Epi eoritu. a ibere: glume about one-third the length of the spikelet, obtuse or acute; second glume and sterile lemma exceeding the fruit and slightly pointed beyond it; fruit 1.5 mm. long, 1 mm. wide, obtuse. ' Autumnal form decumbent, with geniculate, sometimes rooting nodes, branching from all the nodes, the early branches nearly equaling the primary culm, repeatedly branching, forming a large, loose straggling clump, the ultimate blades and panicles scarcely reduced. Type U. 8S. National Herbarium no. 592750 collected September 26, 1897, South Berwick, Maine, by M. L. Fernald, being the type of P. wnciphyllum forma prostratum. This species somewhat resembles P. villosissimum, though much less copiously pilose. It may be distinguished from that species and from P. huachucae sil- vicola by the pointed spikelets 2 mm. long, the second glume and sterile lemma produced in a minute point beyond the fruit. DISTRIBUTION. Dry or sandy open woods, Maine, Massachusetts and eastern New York; apparently rare. Maine: South Berwick, Fernald in 1897, Parlin 938 (Gray Herb.); Island Falls, Fernald in 1897; Mount Desert Island, Fernald in 1892 in part (the last two in N. E. Bot. Club Herb.). MassacuusetTts: Ashburnham, Harris in 1896. New York: Platte Clove, Catskills, Williamson in 1903 (Phila. Acad. Herb.). Fia. 246.—Distribution of P. languidum. — Pw > HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 233 137. Panicum villosissimum Nash. Panicum tectum Willd.; Spreng. Syst. Veg. 1: 313.1825. This is givenasasynonym under P. dichotomum. The type specimen, in the Willdenow Herbarium, is the autumnal form. It is labeled ‘‘Panicum tectum panicula divaricata. * * * Hab. a America boreali.’’ A second specimen so named in the Willdenow Herbarium was sent by Muhlenberg and is P. xalapense. _ Panicum dichotomum villosum Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8: 31. 1889, not P. villosum Ell. 1816. The author cites “‘ P. villosum Ell.?” but since on the same page he gives this name unquestioned as a synonym of P. consanguineum Kunth, Elliott’s species can not be taken as the basis of Vasey’s variety. No locality nor speci- meniscited. A freely branching early autumnal specimen in the National Herbarium marked ‘‘dichotomum var. villosum” in Vasey’s writing, and agreeing well with his description, is chosen as the type. This was collected ‘‘near Pierce’s Mill, Rock Creek, D. C., July 1, 1883,’’ by Dr. Vasey. Panicum nitidum pubescens Scribn. in Kearney, Bull. Torrey Club 20: 479. 1893. This is listed without description as the name of two numbers, 58 and 141, of Kearney’s collection of plants in Harlan and Bell Counties, Kentucky. ‘‘ Paniewm laxiflorum pubescens Chapm.”’ is cited but as the latter name had not at that time been published by Chapman, P. nitidum pubescens must be considered a nomen nudum. The speci- mens of his 58 and 141 in the National Herbarium, and distributed by Kearney, com- prise P. villosissimum and P. huachucae, but the majority are the former. Panicum laxiflorum pubescens Chapm.; Kearney, Bull. Torrey Club 20: 479. 1893, not Vasey 1892. Thisisgivenasasynonym of P. nitidwm pubescens Scribn., but is later described by Chapman4 and based on P. pubescens Lam. Panicum villosissimum Nash, Bull. Torrey Club 23: 149. 1896. ‘‘Collected by Dr. John K. Small in the Ocmulgee River swamp, below Macon [Georgia], May 18-24, 1895.” The type, in Nash’s herbarium, consists of several vernal culms with branches appearing, but secondary panicles not expanded. The spikelets are 2.3 mm. long. Panicum atlanticum Nash, Bull. Torrey Club 24: 346. 1897. :“‘Type specimens collected by the writer on dry somewhat shaded knolls in the grounds of the New York Botanical Garden.’’ The type, in Nash’s herbarium, consists of a small clump and of two single specimens, the culms beginning to branch, primary panicles mature, sec- ondary panicles immature. The spikelets are 2.2 mm. long. This differs from the type of P. villosissimum only in the somewhat stiffer culms and slightly smaller spikelets. Panicum haemacarpon Ashe, Journ. Elisha Mitchell Soc. 15:55. 1898. ‘‘ District of Columbia: Kearney; 1897. Ashe: North Carolina; Chapel Hill, 1898. Iowa: Carver; Jewell Junction, 1895, No. 258.’’ The first specimen cited is chosen as the type. This is in Ashe’s herbarium and consists of a tuft of three simple culms with nearly mature panicles and two autumnal culms of the previous year. Panicum xanthospermum Scribn. & Mohr, Contr. Nat. Herb. 6: 348.1901. ‘‘Type specimen collected by Dr. Charles Mohrin open sandy soil, Greenville, Butler County, Ala., May 8, 1898.’’ This specimen, which is in the National Herbarium, consists of a tuft with two simple culms 18 and 20 cm. high, and the burned bases of others, evi- dently a second growth after a fire. Except in its smaller size it compares well with the type of P. atlanticum. The spikelets, which are immature, are 2.2 mm. long. By selecting the shorter culms it could be matched from many typical clumps of P. villosissimum. This species was described by Scribner? as Panicum pubescens Lam., as indicated by a note upon a sheet, then in his possession, of a duplicate type of P. villosissimum. 2F\. South. U. S. ed. 3. 586. 1897. b Tenn. Agr. Exp. Sta. Bull. 7:52. pl. 15. f. 58. 1894. 234 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Vernal plants light olive green; culms densely tufted, 25 to 45 cm. high, slender, erect or ascending, papillose-pilose with spreading hairs 3 mm. long; sheaths shorter than the internodes, pilose like the culm; ligules 4 to 5 mm. long; blades rather firm, ascending or sometimes spreading, 6 to 10 cm. long, 5 to 10 mm. wide, often subin- volute toward the acuminate apex, little narrowed toward the base, pilose on both surfaces, the hairs of the upper surface appressed, longer and less copious; panicles short-exserted, 4 to 8 cm. long, usually as wide, loosely flowered, the spikelets long- pediceled, the axis sparsely pilose, the branches rather stiffly ascending or spread- ing; spikelets 2.2 to 2.3 mm. long, 1.1 mm. wide, oblong-elliptic, obtuse or obscurely pointed, papillose-pubescent with spreading hairs; first glume sometimes glabrous, one-third to nearly half the length of the spikelet, acute; second glume and sterile lemma subequal, the glume slightly shorter than the fruit at maturity; fruit 1.9 mm. long, 1 mm. wide, elliptic, subacute. Autumnal form at first decumbent, often with geniculate nodes and arched internodes, the first branches appearing at about the maturity of the primary panicle, late in the season prostrate, the leaves of the fascicled branchlets appressed, giving a combed-out appearance, a character conspicuous in the field but less so in the herba- rium; blades not greatly reduced, often with only a few hairs on the upper surface, overtopping the much reduced panicles; winter rosette appearing rather early, blades long, bluish green, densely pilose. This is fairly uniform as a whole for a species of so wide a range, but exceptional specimens with spikelets only 2 mm. long occur, such as Andrews, Southington, Conn., in 1902; Chase 2378, 3762; Dodge 60, 83; Herriot 86; Hitchcock 1635; Smith, Framing- ham, Mass., in 1898. In these the habit and other characteristics are those of the typical form. Another rarer variation with blades nearly or quite glabrous on the upper surface is found, as Ashe, Manteo, N. C., Chase 3121, Commons 52. Fic. 247.—P. villosissimum. From type specimen. DISTRIBUTION. Dry sandy or sterile soil, open woods and hillsides, Massachusetts to Minnesota, south to Florida and Texas. MAssAcHuUSETTS: Framingham, Smith in 1898. ConnecticuT: Franklin, Graves 14; Southington, Andrews in 1902. New York: Bronx Park, Nash in 1897; Long Island, Bicknell in 1902 and 1904. Ontario: Galt, Herriot 86; Squirrel Island, Dodge 60, 83. New Jersey: South Amboy, Mackenzie 1381; Wildwood Junction, Chase 3522. PENNSYLVANIA: Westchester, Windle 12f. (Hitchcock Herb.) Onto: Sandusky, Morris 135. InpianaA: Clark, Bebb 2833, 2834; Miller, Chase 1545, 1563; Umbach 2646 and in 1897. : Iuuino1s: Madison County, Hggert 293; Starved Rock, Chase 1606. MicnicaNn: Carleton, Wheeler in 1890 (Mich. Agr. Col. Herb.). Wisconsin: Juneau County, Mearns 25. Minnesota: Minneapolis, Sandberg 316 in part (Hitchcock Herb.). Missourt: Monteer, Bush 732, 750, 4788; Chadwick, Bush 14; Eagle Rock, Bush 141; Pleasant Grove, Bush 333; Carter County, Eggert 291; Jefferson County, Eggert 292; Noel, Bush 5023. DeLawarRE: Rehoboth, Commons 56 in part; Greenbank, Commons 38; Frank- ford, Commons 52; Lewes, Hitchcock 161. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 235 MARYLAND: Between Chesapeake Beach and Chesapeake Junction, Hitchcock 1608, 1619, 1623, 1627, 1631, 1635, 1643, 1644, 2415; Riverdale, Chase 2378; Beltsville, Chase 37624; Cabin John, Chase 2905; Hyattsville, Steele in 1907. District or CotumsBra: Hitchcock 393, 2403, Kearney 29, Vasey 125, Ward in 1879, Williams 5. Virerntia: Four-Mile Run, Chase in Kneucker Gram. Exs. 555, Hitchcock 391, Pollard 337; Norfolk, Kearney 302; Dismal Swamp, Chase 3653, 3675, Tyler in 1905; Munden, Mackenzie 1710. Norra Carona: Manteo, Ashe in 1898; Raleigh, Chase 3081; Chapel Hill, Chase 3055, 3056, 3058, 3071, 3075; Wilmington, Chase 3121, 3141, 3155, Hitch- cock 394; Biltmore, Boynton 4; Magnetic City, Wetherby 64; Jacksonville, Chase 3191; Lenoir, Hitchcock 395. Sourn Carona: Orangeburg, Hitchcock 392, 1384; Clemson College, House 2158. Gerorara: Lookout Mountain, Ruth 19; Stone Mountain, Hitchcock 396; Rabun County, House 2269; Lagrange, Tracy 8869; Warm Springs, Tracy 8859; Ma- con, Small in 1895; Thomson, Bartlett 1414, 1497; Augusta, Cuthbert 387, 1120. Froriwa: Chipley, Combs 613; Chattahoochee, Tracy 3653, 3658; Leon County, Curtiss F in 1886 in part. Kentucxry: Harlan County, Kear- ney 58 in part, 141. TENNESSEE: Cocke County, Kearney 971; Knoxville, Ruth 72 in part. AvaBaMA: Auburn, Tracy 3751, 3753, 3754, 3758; Pisgah, Chase 4472; Gateswood, Tracy 8426 in part; Tuskegee, Carver 19; Mobile, Kearney 34 in part; Flomaton, Hitchcock 1046; Greenville, Mohr in 1898. Misstssrept: Starkville, Kearney 24 in part; Taylorville, Tracy 8415; Jackson, Hitchcock 1299; Enterprise, Tracy 3272; Biloxi, Kearney 337; Ocean Springs, Tracy 161; Mississippi City, Hitchcock 1109. Arkansas: Benton County, Plank 42, 100. Loutstana: Calhoun, Hitchcock 1259, 1275, 1289; Shreveport, Hitchcock 1256. Texas: Waller County, Thurow 25; Weatherford, Tracy 7943 in part; Angelina County, Reverchon 4136; Denison, Bebb 2661. OxtaHoma: Sapulpa, Bush 1220. Fig. 248.—Distribution of P. villosissimum. 138. Panicum pseudopubescens Nash. Panicum pseudopubescens Nash, Bull. Torrey Club 26: 577. 1899. “‘Type collected at Auburn, Lee Co., Alabama, May 7, 1898, by Messrs. F. 8. Earle and C. F. Baker, No. 1537.” The type, in Nash’s Herbarium, is a clump of a few vernal, mostly imma- ture culms. Some of the blades are nearly naked along the middle of the upper surface. Other specimens cited by Nash under this species, Karle & Baker 1522, 1524, 1526, 1529, have narrower blades than the type, with the upper surface often nearly glabrous; these represent the more usual form of the species. This species was described by Elliott 4 as P. pubescens Lam., as shown by the speci- men so labeled in his herbarium. The culm of this is appressed-pubescent, not ‘‘ very glabrous, ’’ as stated in the description. The — ies described in Gray s Manual? under P. ovale Ell. is P. ee ee a Bot. 8. ©. 4 Gs. 1:125. 1816. b A. Gray, Man. ed 7. 111. 1908. 236 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DESCRIPTION. Vernal form similar to that of P. villosissimuwm; culms somewhat stiffer, the pubes- cence more silky, appressed on the culms, ascending on the sheaths; ligules 2 to 3 mm. long; blades somewhat firmer, the pubescence on the upper surface short like that on the lower and sparse or wanting down the center or occasionally glabrous on the upper surface; panicles averaging larger, the spikelets not so long-pediceled; spikelets 2.25 to 2.4 mm. long, 1.1 mm. wide, obovate-elliptic, obtuse, or slightly pointed, pubescence as in P. villosissimum, second glume slightly shorter than the fruit at maturity; fruit 1.9 mm. long, 1mm. wide, elliptic, subacute. Autumnal form usually stiffly spreading, some- Wig: 2402) papudopubescens. From times prostrate, culms sparingly branching from type specimen. the lower and middle nodes after the maturity of the primary panicle, less freely branching than P. villosissimum,; the upper surface of the reduced blades usually glabrous except along the margins and at the base; winter leaves as in P. villosissimum. This species is very closely allied to P. villosissimum and occasional specimens are about as close to one type as to the other. In these cases the ascending, more silky pubescence of the culms and less pubescent or glabrous upper surface of the blades, together with the stiffer habit, have been used to distinguish P. pseudopu- bescens. DISTRIBUTION. Sandy, open woods, Connecticut to Illinois, south to Florida and Mississippi; also in Mexico. Connecticut: South Britain, Harger 6031 (Bissell Herb.). New Jersey: Atsion, Chase 3566; Camden, Smith; Wildwood Junction, Chase 3520. Outro: Cedar Point, Claassen (Gray Herb.). InprANna: Miller, Chase 1542, Umbach in 1898; Clark, Umbach 1685; Clark Junc- tion, Bebb 514, 2882; Dune Park, V. H. Chase 291, Hill 100 and 101 in 1905, Umbach 1080. Inurnors: Starved Rock, Chase 1605; Hanover, Gleason & Gates 2535, 2539, 2575. MicuHiGcAN: Saginaw Bay, Morris 240 in part; Twin Lakes, Wheeler in 1900. MaRYLAND: Caroline County, Nor- ton in 1907. District or CoLumBIA: Steele in Fig. 250.—Distribution of P. pseudopubescens. 1899. Norra Carouina: Roanoke Island, Ashe in Curtiss Pl. So. U. 8. 6452, Chase 3218, 3222; Wilmington, Chase 3109, 3115, 3147, 3160, Hitchcock 1437, 1474, 1476, 1484, 1485; Biltmore, Boynton in 1906. Sourn Caronina: Orangeburg, Hitchcock 398, 1380, 1383, 1394, 1401, 1402; Fripps Island, Cuthbert 1164. GeoraiaA: Stone Mountain, Hitchcock 397, 1356, 1357. Fioriwa: Lake City, Hitchcock 10134. TENNESSEE: Ducktown, Chambliss 26. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 237 AzaBama: Auburn, Earle & Baker 1522, 1537, Hitchcock 1331, 1336; Gateswood, Tracy 8426 in part; Flomaton, Hitchcock 1048. . Mississrer1: Biloxi, Hitchcock 1080; Mississippi City, Hitchcock 1090, 1096, 1112, Kearney 301 in part. Mexico: San Luis Potosi, Schaffner 146. 139. Panicum ovale Ell. Panicum ovale Ell. Bot. S. C. & Ga. 1: 123. 1816. ‘‘Grows in Carolina and Georgia. Sent from St. Mary’s, Georgia, by Dr. Baldwin.”’ The type, in the Elliott Herbarium, consists of the upper portion of a vernal culm with two leaves and an immature panicle included at base, the culm and sheaths densely pilose with ascending hairs, the blades long pilose along the margin. The ticket attached to this specimen reads: “‘ Panicum Ovale. Hab. St. Mary’s Georg. Dr. Baldwin.’’ Another vernal culm with immature panicle mounted on the same sheet belongs to P. commutatum. To the culm is at- tached a slip marked ‘‘64” but with no data. Since Dr. Baldwin’s is the only speci- men cited, the one with the Baldwin label must be considered the type, though Elliott’s description seems to show he had the two confused. Panicum ciliiferum Nash, Bull. Torrey Club 24: 195. 1897. ‘‘Type collected by the writer in the ‘high pine land’ at Eustis, Lake Co., Florida, March 12-31, 1894, no. 147.’ The type, in Nash’s herbarium, is the vernal form with an old autumnal culm attached. The spikelets are 2.8 mm. long. In a note following the description Nash states that after having examined a specimen named P. ovale by Elliott he con- siders P. ciliiferum as distinct. The specimen referred to is in the Torrey Herbarium, and is labeled Panicum ovale Elliott, ‘‘From Elliott.’’ This is a puberulent narrow- leaved form of P. commutatum, and is the form described in Small’s Flora as P. ovale. Panicum erythrocarpon Ashe, Journ. Elisha Mitchell Soc. 16: 90. 1900. ‘The type material was collected by the writer on the sand hills of New Hanover county, N. C., May 19, 1899.’’ The type, in Ashe’s herbarium, is the vernal form, labeled, ‘‘Shady slopes on the sand hills one mile north of Wilmington [New Hanover County], N. C.”’ DESCRIPTION. Vernal plants light olive green; culms densely tufted, 20 to 50 cm. high, erect or ascending, rather stout, long-pilose below with ascending or appressed hairs, often nearly glabrous above, usually leafy at the base, the nodes densely bearded with short spreading hairs; sheaths shorter than the internodes or the lower overlapping, ascend- ing-pilose, the upper less densely so, rarely nearly glabrous; ligules composed of a ring of hairs about 1 mm. long with a second sparse ring 2 to 3 mm. long above it; blades firm, ascending, 6 to 10 cm. long, 5 to 10 mm. wide (the uppermost much smaller), sharply acumi- nate, rounded at base, the upper surface usually nearly glabrous except for long hairs on or near the margin and base thus giving the blades the appearance of being strongly ciliate, these hairs occasionally wanting except at the base, the lower surface appressed-pubescent; panicles usually short-exserted, 5 to 9 cm. long, about as wide when fully expanded, the lower branches finally spreading, rarely drooping; spikelets 2.7 to 2.9 mm. long, 1.3 mm. wide, oblong-elliptic, obtuse, pilose, sometimes rather sparsely so; first glume one-third to nearly half the length of.the spikelet, usually pointed; second glume Fic. 251—P, ovale. From type specimen. 4} 1. Southeast. U. S. 102. 1903. 238 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. slightly shorter than the fruit and sterile lemma at maturity; fruit 2.2 mm. long, 1.2 mm. wide, elliptic, obtuse. Autumnal form spreading-decumbent, the stiff culms rather loosely branching from the middle and upper nodes, the ultimate branchlets crowded at the ends of the primary branches, the reduced blades erect; winter leaves very firm, conspicuously ciliate; short culms with tufted branches sometimes formed during the winter, the green bushy crown persistent at the base of the tall vernal culms. Curtiss’s no. 4877 is referred here although the specimens resemble P. common- sianum, and the ligule is only 1 mm. long, as in that species; the spikelets, however, are those of P. ovale. DISTRIBUTION. Dry sandy woods, North Carolina to Florida, also in Texas. NortuH Carouna: Near Wilmington, Ashe in 1899, Chase 4589. SoutrH Carona: Isle of Palms, Hitchcock 107; Aiken, Ravenel. Fioripa: Duval County, Curtiss 3583* in part; Jacksonville, Curtiss 4877, 5866, 5813; Lake City, Combs 138, 145, Hitchcock 550, 1013; Madi- son, Combs 225; Chattahoo- chee, Tracy 3617; Gainesville, Chase 4250, 4261; Old Town, Combs 888; Eustis, Curtiss 6616, Nash 75, 103, 147, 1118, 1518, 1857; Lake Harris, Chase 4118; Tavares, Hitchcock 820; Grasmere, Combs 1080; San- ford, Hitchcock 785, 787; Titus- ville, Hitchcock 7614; Ormond, Fig. 252.—Distribution of P. ovale. Mitchcock 160; Dunedin, Tracy 6725; Braidentown, Hitchcock 968; Lakeland, Hitchcock 833, 846, 847, 851; Myers, Chase 4174, Hitchcock 900, 914, Lee Co. Pl. 474; Miami, Chase 3866, 3947, Hitchcock 634, 661, 668, 677, 719, Pollard & Collins 223; Homestead, Hitchcock 688. ; Texas: Waller County, Thurow 17 in 1906. 140. Panicum scoparioides Ashe. Panicum scoparioides Ashe, Journ. Elisha Mitchell Soc. 15: 53. 1898. ‘‘ Based on No. 283, ex. Herb. A. Commons. Dry soil. Centreville, Del. June 1873. Distributed sub nom. P. Scribnerianum Nash.’’ This specimen could not be found in Ashe’s herbarium, but a specimen bearing the above name and data is in the National Herbarium and is doubtless the type. This consists of four vernal culms with immature panicles partly included in the uppermost sheaths. DESCRIPTION. Vernal plants light green; culms few to several in a tuft, 30 to 50cm. high, slender,erect or ascend- Fig. 253.—P. scoparioides. From type é ; ies ‘ = i specimen in National Herbarium. ing, sparsely papillose-hispid with ascending hairs or nearly glabrous, the upper internodes shortened; nodes sometimes sparsely bearded; sheaths papillose-hispid to nearly glabrous, the lower distant, the upper approximate, sometimes overlapping; ligules 2 to 3 mm. long; blades firm, ascending, 7 to 10 cm. long, 6 to 10 mm. wide, tapering to the a@See note on type of P. huachucae, page 215. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 239 rounded base, acuminate, appressed-pubescent beneath, sparsely hispid on the upper surface, usually a few long hairs at the base; panicles short-exserted, usually included at the base until maturity, rather densely flowered, 4 to 7 cm. long, about two-thirds as wide, the branches ascending; spikelets 2.2 to 2.3 mm. long, 1.2 mm. wide, obovate, obtuse or minutely pointed; first glume about one-fourth as long as the spikelet, subacute; second glume and sterile lemma papillose-pubescent, strongly nerved, subequal, as long as the fruit, the margins atthe summit usually inrolled, the midnerve produced into an apiculus; fruit 1.9 mm. long, 1.1mm. wide, elliptic. Autumnal form erect or spreading, culms sparingly branching from the upper and middle nodes after the maturity of the primary panicle, the stiff, reduced blades involute-pointed, much exceeding the panicles. This species is less pubescent than any other in this group except P. oc- cidentale. The smoother specimens somewhat resemble P. boreale. DISTRIBUTION. Dry sandy or gravelly soil, Vermont to Delaware; also in Minnesota; ap- parently rare. Vermont: Hartland, Jones 30. Connecticut: Southington, Bissell 385, 5581, 8084; East Lyme, Graves in 1903 (Hitchcock Herb.). InpIANA: Gary, Umbach 3686. Minnesota: Hennepin sisal Sandberg in 1890; Ramsey County, Oestlund in 1884 in part. DELAWARE: Centerville, pines 283, 359. Fig. 254.— Distribution of P. scoparioides. 141. Panicum shastense Scrib. & Merr. Panicum shastense Scribn. & Merr. U.S. Dept. Agr. Div. Agrost. Cire. 35: 3. 1901. ““Type specimens collected in a moist meadow at the edge of pine forests at Castle Crag, near Mt. Shasta, California, by Louis A. Greata, June, 1899.’’ The type, in the National Herbarium, consists of three vernal culms, 25 to 30 em. high, with short-exserted, nearly mature panicles. DESCRIPTION. Vernal form pale green; culms tufted, 30 to 50 cm. high, slender, ascending from a more or less geniculate base, papillose-pilose with ascending hairs; nodes short-bearded; sheaths papillose- pilose, the hairs spreading; hairs of the ligule rather sparse, 2to 3mm. long; bladesascending, 6 to 8 cm. long, 6 to 8 mm. wide, acuminate, scarcely narrowed toward the base, papillose-pilose on the under surface and with scattered long hairs on the upper; panicles short-exserted, 6 to 8 cm. long, about two-thirds as wide, the axis pilose, the flexuous branches ascending; spikelets 2.4 to 2.6 mm. long, 1.2 to 14mm. wide, obovate-oblong, obtuse, papillose-pubescent; first glume one-fourth to one-third as long as the spikelet, pointed; second glume scarcely equaling the fruit and sterile lemma; fruit 2.1 mm. long, 1.3 mm. wide, elliptic. Fic. 255.—P. shastense. From type specimen. 240 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Autumnal form spreading, with geniculate nodes and elongated, arched internodes, rather sparingly branching from the middle nodes, the primary branches elongated, the ultimate branchlets shorter than the internodes. DISTRIBUTION. Known only from Castle Crag, Shasta County, California, where it is found in meadows. : CauirornNiA: Castle Crag, Greata in 1899, Hitchcock 3072. Columbiana.—Culms stiff, pubescent with appressed or ascending villous hairs, or crisp-puberulent, the sheaths pubescent like the culms or nearly glabrous; ligules usually less than 1 mm. long, rarely 1.5 mm. long; blades firm, thick, stiffly ascending, not over 7 mm. wide; spikelets pubescent, 1.3 to 3.2 mm. long, 5 to 9-nerved. Autumnal form freely branching, the branches and stiff blades mostlyappressed. With the exception of P. tsugetorum, confined to dunes and dry sands of the Coastal Plain, Massachusetts to Florida. The species of this group form a lineal series from P. malacon with spikelets 3.2 mm. long to P. columbianum thinium with spikelets only 1.3 mm. long. With the excep- tion of P. malacon and P. wil- mingtonense, there are con- necting forms between the successive species in this series, especially between P.commonsianum and P. addisonii, and between P. tsugetorum and P. colum- bianum. These intermediate specimens are, however, comparatively rare. On the other hand, P. oricola and hairy forms of P. tsugetorum make a clear cut division between the Lanuginosa and Columbiana impossible. Fig. 256.—Distribution of P. shastense. Spikelets 2 to 3.2 mm. long, mostly elliptic. Winter blades elongated, 5 to 10 cm. long; spikelets 2 sum; lone; plants blue ereen:.-s2.<<2--4.+:0-26 145. P. wilmingtonense. Winter blades 1 to 3 cm. long. Spikelets 3.2 mm. long; first glume conspicuously GIRANIDS ....2.3.. «28a went Sock ae eecee s eee 142. P. malacon. Spikelets not over 2.5 mm. long; first glume not distant. Spikeletsabout 2.4 mm. (2.2 to 2.4 mm.) long; panicle open, branches stiffly spreading ..143. P.commonsianum. Spikelets 2 to 2.1 mm. long; panicle rather dense, Ipranches ascending rc ocso8scktccincce ot om 144. P. addisonit. Spikelets not over 1.9 mm. long, obovate, turgid. Culms crisp-puberulent or appressed-pubescent with crimped hairs; plants bluish or grayish green; pan- icles about 3 to 7 cm. long. Spikelets 1.8 to.1.9 mm. long... <2. semen snes ce aes 146. P. tsugetorum. Spikelets 1.5 to 1.6 mm. long.........-...25.2..). 147. P. columbianum. Culms appressed or ascending-pilose; spikelets not over 1.5 mm. long, rounded and turgid; plants oliva- ceous; panicles rarely more than 3 cm. long. Spikelets 1.5 mm. long; culms rather stout; autumnal form branching from all the nodes..........- 148. P. oricola. duced blades stiff, erect, and involute- pointed. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 241 Spikelets 1.3 to 1.4 mm. long; culms very slender; autumnal form with branches mostly aggre- gated toward the summit.............--...- 147a. P. columbianum thinium. 142. Panicum malacon Nash. Panicum malacon Nash, Bull. Torrey Club 24: 197. 1897. ‘‘Collected by the writer in the ‘high pine land’ at Eustis, Lake County, Florida, May 1-15, 1894, no. 628.’’ The type, in Nash’s herbarium, is the early branching form, the numerous branches appressed, the blades densely puberulent on both surfaces. Panicum strictifolium Nash, Bull. Torrey Club 26:579. 1899. ‘‘Collected by the writer in the high pine land at Eustis, Lake Co., Florida, May 3, 1894, no. 603. Most nearly related to P. malacon, but distinguished by the less copious pubescence which is much finer and softer, and by the glabrous upper surface of the blades.’’ The type, in Nash’s herbarium, is the early branching form, the primary panicles destitute of spikelets and the secondary panicles immature. This is less densely pubescent than is Nash 628, but the pubescence is not softer. The type of P. malacon is more copiously pubescent than are most specimens of this species. The spikelets of the two types are identical except that those of Nash 603 are immature, while those of no. 628 are mature. DESCRIPTION. Vernal form erect or stiffly spreading, pur- plish olive green; culms 30 to 50 cm. high, pubescent with ascending hairs, the nodes short-pubescent; sheaths pubescent like the culms, sometimes sparsely so; blades stiffly ascending or somewhat spreading, 4 Fig. 257.—P. malacon. From type specimen. to 12 cm. long, 3 to 5 mm. wide, rarely wider, sharply acuminate, scarcely narrowed toward the base, puberulent beneath, puberulent or glabrous above, often villous at or near the margin or base with long hairs; panicles 4 to 7 cm. long, three-fourths to nearly as wide, few-flowered, branches few, stiffly ascending, the spikelets on long, stiff pedicels; spikelets 3 to 3.2 mm. long, 1.4 to 1.5 mm. wide, obovate (oblong before maturity), pubescent; first glume distant, half as long as the spikelet or more, sub- acute, 5 to 7-nerved; second glume scarcely equaling the fruit and sterile lemma; fruit short-stipitate, 2.5 mm. long, 1.4 mm. wide, elliptic, acute. Autumnal form more or less decumbent-spreading, branching from the lower and middle nodes, the branches appressed and later rather sparingly producing appressed fascicled branchlets, the re- DISTRIBUTION. Dry pine woods, the so-called ‘‘high pine land,’’ Florida. Fiormwa: East Pass, Tracy 9140; Lake City, Combs 167; Old Town, Combs 855; Grasmere, Combs 1036, 1161; Gainesville, Fic. 258.—Distribution of P. malacon. Chase 4251; Eustis, Chase 4072, 4077, Hitchcock 801, 813, Nash 36, 63, 132, 603, 628; Clearwater, Tracy 6700; Dunedin, Tracy 6725; Lakeland, Hitchcock 845. 41616°—vo1 15—10 16 242 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 143. Panicum commonsianum Ashe. Panicum commonsianum Ashe, Journ. Elisha Mitchell Soc. 15: 55.1898. ‘‘Based on no. 341, Commons. Collected in drifting sands along the coast, Cape May, N. J., June, 1898.’’ The type, in Ashe’s herbarium, ‘‘Ex. Herb. A. Commons,”’ consists of five tufts of vernal culms with mature primary panicles. DESCRIPTION. Vernal plants grayish olive, drying brownish; culms usually in dense tufts 20 to 50 cm. high, stiffly ascending or spreading, papillose-strigose to appressed-pilose, the hairs at the nodes more spreading; sheaths shorter than the internodes, strigose to appressed- pilose like the culms but less densely so; ligules 1 mm. long or less; blades firm, stiffly ascending, 5 to 8 (rarely 9) cm. long, 4 to 7 mm. wide, broadest near the rounded base, the ser- rulate, cartilaginous margin involute toward the acuminate apex, glabrous on the upper surface or with a few long hairs toward the base or margin, strigose on the lower surface or glabrous; panicles long-exserted, 4 to 8 cm. long, about as wide, loosely flowered, the axis and branches strigose to nearly glabrous, the branches stiffly spreading, spikelet-bearing toward the ends; spikelets 2.2 to 2.4 mm. long, 1.2 mm. wide, elliptic, subacute, pubescent; first glume about half as long as the spikelet, sometimes longer, usually pointed, 3-nerved; second glume slightly shorter than the fruit and sterile lemma at maturity; fruit 2mm. long, 1 mm. wide, elliptic, subacute. Autumnal culms branching from the middle and upper nodes, after the maturity of the primary panicles becoming spreading or prostrate, the larger clumps forming mats in the sand, the reduced secondary subinvolute blades rather crowded, stiffly ascending, overtopping the panicles; winter blades lanceolate, commonly more hairy than those of the culm. This species is variable as to pubes- cence. Fig. 259.—P. commonsianum. From type specimen. DISTRIBUTION. Dunes and sandy woods near the coast, Connecticut to northern Florida. Connecticut: North Haven, An- drews in 1901; South Windsor, Bissell 12000. Fia. 260.—Distribution of P. commonsianum. New York: Lake Roukonkoma, Bicknell; Rockville Center, Bicknell in 1906; Valley Stream, Bicknell in 1905; Hempstead, Bicknell in 1903. New Jersey: South Amboy, Mackenzie 1485, 2155, 2165; East Plains, Stone 4, 6; Lakehurst, Mackenzie 2067; Toms River, Bicknell in 1900, Chase 3575; Forked River, Chase 3584, 3596; Atsion, Chase 3531, 3541, 3544, 3570; Cape May, Canby 4 in 1902, Commons 48, 341; Wildwood, Chase 3517}. DELAWARE: Lewes, Hitchcock 408. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 248 VirertA: Region of Cape Henry, Chase 2349, 5428, Hitchcock 407, Kearney 1393, 1454, 1776, 2114, Pollard & Maxon in 1900. NortH Carona: Wilmington, Chase 3161, 4601, Hitchcock 326, 336. SourH Carona: Isle of Palms, Chase 4544. Georaia: Augusta, Kearney 209. Froripa: East Pass, Tracy 9141. 144. Panicum addisonii Nash. Panicum addisonii Nash, Bull. Torrey Club 25: 83. 1898. ‘‘Collected by Mr. E. P. Bicknell in sandy soil at Wildwood, N. J., May 30 and 31, 1897.’’ The type specimen, in Nash’s herbarium, is the early branching form with culms 15 to 30 cm. high, mature primary panicles, and spikelets 2 to 2.1 mm. long. Panicum owenae Bicknell, Bull. Torrey Club 35: 185. 1908. ‘‘Type collected September 20, 1907, on the sandy commons west of the town [Nantucket], deposited in herb. N. Y. Bot. Garden.’’ The type specimen is the autumnal form, the primary panicles destitute of spikelets, the secondary panicles among the crowded autumnal leaves. The blades are nearly smooth except for scattered long hairs near the margin. The vernal form collected by Bicknell at the type locality June 20, 1908, is more pubescent. DESCRIPTION. Vernal form similar to that of P. commonsianum and often closely resembling that species; culms more slender, rarely as much as 40 cm. high, appressed or ascending- pilose at least below, puberulent above; sheaths sparsely ascending-pilose; blades stiffly ascending, 4 to 7 cm. long, 3 to 6 mm. wide, involute-pointed, glabrous on the upper surface, sometimes with a few long hairs near the margin, pubescent or glabrous beneath; panicles long-exserted, 2 to 6 cm. long, two-thirds to three-fourths as wide, the stiff branches ascending, the panicle thus ap- : pearing more densely flowered; spikelets 2 to 2.1 mm. long, 1.1 mm. wide, obovate, blunt, papillose- pubescent, or the papillee obscure; first glume one- third to half as long as the spikelet, usually pointed; second glume and sterile lemma barely equaling the fruit at maturity; fruit 1.7 mm. long, 1 mm. wide, elliptic, subobtuse. Autumnal form more or less spreading, rather Phas pai =P. Gains) Backs freely branching from all the nodes, the branches type specimen. appressed or narrowly ascending, the later branch- lets somewhat fascicled, the stiff blades not greatly reduced, overtopping the num- erous reduced panicles. This species very closely approaches forms of P. commonsianum, the smaller, broader spikelets with shorter first glume affording about the only constant difference, though it is P.commonsianum rather than P. addisonii which varies much. The type of P. owenae and a specimen collected by Steele, Suitland, Maryland, in 1899, both late autumnal forms, have fascicled primary branches from the lower nodes. Usually only the secondary branchlets are fascicled. Two southern specimens, Chase 4580 and Hitchcock 557, have laxer culms and more spreading branches. Jf {| Wil ‘\ f ¥ ae 244 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Sand barrens, Massachusetts to South Carolina. Massacuuserrs: Andover, Blake in 1882; Nantucket, Bicknell in 1907 and 1908. Connecticut: East Lyme, Graves in 1903 (Bissell Herb.). New York: Hempstead, Bicknell in 1903 and 1906. New Jersey: Wildwood, Bicknell in 1897, Chase 3517; Wildwood Junction, Chase 3523; Toms River, Bicknellin 1900; Forked River, Chase3583,3595; Atsion, Chase 3538; Lakehurst, Chase 3574; Somers Point, Canby 5in 1902; Tuckerton, Chase 3603. MaryLAND: Chesapeake Beach, Hitchcock 1617; Suitland, Steele Fic. 262.—Distribution of P. addisonii. in 1899. VirGIniA: Virginia Beach, Hitchcock 556 (Hitchcock Herb.). Nortu Carona: Wilmington, Chase 3166, 4580; Hitchcock 335, 399. Sourn Carona: Orangeburg, Hitchcock 557. 145. Panicum wilmingtonense Ashe. Panicum wilmingtonense Ashe, Journ. Elisha Mitchell Soc. 16: 86.1900. ‘‘The type material collected in May, 1899, on the sand hills near Wilmington, N. C., is preserved in my herbarium.’’ The type, in Ashe’s herbarium, is labeled, ‘‘Shady slopes on the sand hills one mile to north of Wilmington, May 17, 1899. W. W. Ashe, Collector.’”? The plants are the vernal form with some autumnal culms of the preced- ing season attached. Panicum alabamense Ashe, N. C. Agr. Exp. Sta. Bull. 175: 116.1900, not Trin. 1854. “Auburn, Ala., May 7, 1898. Number 1530, Alabama Biological Survey.’’ The type, in Ashe’s herbarium, is a tuft of young vernal culms, the panicles only partly exserted. Mounted on the sheet with this is a specimen of P.lucidwm. Ashe’s description refers to the latter only in so far as the spikelets are said to be glabrous. DESCRIPTION. ~ Vernal form bluish green; culms solitary in small ee ee tufts; slender, erect from an ascending base 20 to type specimen. 40 cm. high, pilose with soft, ascending hairs, the nodes pubescent with short, reflexed hairs; sheaths pubescent like the culms, densely villous-ciliate at the summit; blades rather stiff, ascending, 4 to 9 cm. long, 3 to 7 mm. wide, glabrous on the upper surface, softly pubescent to nearly glabrous beneath, strongly ciliate on margin near base, the thick cartilaginous margin white at least when dry; panicles 5 to 8 cm. long, the branches ascending; spikelets 2 mm. long, 1 mm. wide, elliptic, subacute, first glume one- fourth to one-third as long as the spikelet; second glume and sterile lemma pubes- cent, the glume slightly shorter than the fruit at maturity; fruit 1.7 mm. long, 1 mm. wide, elliptic, obtuse. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 245 Autumnal form spreading, branching from the middle and upper nodes, the branches rather crowded, the reduced involute-pointed blades exceeding the ultimate panicles; blades of the winter rosette as much as7 cm. (rarely 12 cm.) long. DISTRIBUTION. Sandy woods, North Carolina and Alabama; rare. Norra Carouma: Wilmington, Ashe in 1899, Hitchcock 316; Jacksonville, Chase 3195. ALABAMA: Auburn, Alabama Bio- logical Survey, Earle & Baker 1530 in part, Hitchcock 1325; Gateswood, Tracy 8429. Fig. 264.—Distribution of P. wilmingtonense. 146. Panicum tsugetorum Nash. Panicum tsugetorum Nash, Bull. Torrey Club 25: 86. 1898. ‘‘Type material collected by the writer in the Hemlock Grove, New York Botanical Garden, on dry soil, June 22, 1896, no. 287.’ The type, in Nash’s herbarium, consists of a clump of 8 vernal culms 20 to 37 cm. high, decumbent at base and bearing scarcely mature pan- icles. The culms are less stiff and the blades thinner than usual in this species, as the plants grew in the shade. Panicum lanuginosum siccanum Hitche. & Chase, Rhodora 8: 207. 1906. “‘Type Chase 1602. Dry, hot sand of sandstone cliff. Starved Rock, Ill.’’ This specimen, in the National Herbarium, is the early autumnal form, and represents an extremely hairy form of P.tsugetorum. The culms and sheaths are ascending-pilose and the blades are sparsely long-pilose on the upper surface. DESCRIPTION. Vernal plants usually pale bluish green; culms 30 to 50 cm. high, spreading or ascending, the lower nodes often geniculate, densely appressed-pubescent with short, crisp hairs, long hairs more or less copiously intermixed with these on the lower inter- nodes or sometimes nearly to the summit; sheaths commonly not much shorter than the internodes, pubescent like the culm but less densely so, ascending-ciliate on the margin; ligules 1 to 1.5 mm. Jong; blades thickish, firm, with a thin white cartilaginous margin, ascend- ing, 4 to 7 cm. long, 4 to 7 mm. wide, rounded at the base, acuminate, glabrous or with a few long hairs near the base on the upper surface, appressed- puberulent beneath; panicles 3 to 7 cm. long, nearly as wide, the axis and spreading, flexuous branches appressed crisp puberulent; spikelets 1.8 to 1.9 mm. long, 1 mm. wide, obovate-obtuse, rather turgid, short-pubescent; first glume one-third to two-fifths as long as the spikelet, acute; second glume and sterile lemma barely equaling the fruit at maturity; fruit 1.5 mm. long, 1 mm. wide, broadly elliptic, obtuse. Autumnal form decumbent-spreading, branching from the lower and middle nodes often before the maturity of the primary panicles, the branches ascending, the ulti- mate branchlets appressed, the blades not greatly reduced nor crowded; winter rosette appearing rather early, the blades often conspicuously long-pilose. Fic. 265.—P. tsugetorum. From type specimen. 246 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. In this species the blades are typically glabrous on the upper surface and the culms and sheaths are appressed-pubescent with short, crisp hairs, longer hairs intermixed on the lower internodes only, but numerous specimens have blades sparsely long- pilose on the upper surface and culms and sheaths ascending-pilose almost to the summit. This form is represented by the following specimens: MAssacHUSETTS: Smith 740; Ontario: Macoun 26236; New Jersey: Chase 3579, 3608; InpIANA: Chase 1552; Intrnots: Chase 1602 (type of P. lanuginosum siccanum Hitche. & Chase), 1604, 16054; Hill 124 and 129 in 1905; Micutaan: Morris A240. Hill 124 and Chase 3608 are so strongly pilose as to resemble P. implicatum. Two collections from Chesapeake Beach, Chase 3269 and 3270, seem to be interme- diate between P. tsugetorum and P. columbianum, having the habit and pubescence of the former but spikelets only 1.7 mm. long. DISTRIBUTION. Sandy woods, Maine to Illinois, Virginia and Tennessee. Maine: North Berwick, Parlin 1215; Fayette, Chase 3390, 33994; Chesterville, Chase 3321, 3363; Ogunquit, Parlin 1577. Vermont: Salisbury, Brainerd in 1903 (Gray Herb.). Massacuusetts: Framingham, Smith 740. Connecticut: South Manchester, Hitchcock 134; Southington, Andrews 62, 74, Bissell 5594, 5595, 5616. RuopeE Istanp: Gloucester, Collins in 1908. New York: Thousand Islands, Bicknell in 1905; Sylvan Beach, House 1287; Ausable Point, Eggleston 2843; Washington County, Burnham 19; Albany, Peck 6; Oneida Lake, Coville in 1887; Bronx Park, Nash 287; Woodmere, Bicknell in 1902 and 1906; Jamaica, Bick- nell in 1904; Hempstead, Bick- nell in 1903; Norwood, Bick- nell in 1903; Rosedale, Bick- nell in 1904; Valley Stream, Bicknell in 1904; Cedarhurst, Bicknell in 1902. Ontario: Tilsonburg, Macoun 26236. New Jerszy: South Amboy, Mac- kenzie 1379, 2220; Tuckerton, Chase 3601, 3608, 3612; Atsion, Chase 3532, 3543, 3564; Forked River, Chase 3579; Wildwood Junction, Chase 3524. PENNSYLVANIA: Tannersville, Smith 2. Ouro: Defiance County, Fullmer in 1898 (Ohio State Univ. Herb.). Inprana: Dune Park, Chase 1921; Miller, Chase 1544, 1552; Gibson, Bebb 29394, Steuben County, Deam in 1904. Inunors: Starved Rock, Chase 1602, 1604, 16054, Greenman, Lansing & Dixon 155; Oregon, Hill 124 and 129 in 1905. MicuicaNn: Port Huron, Dodge in 1909; Twin Lakes, Wheeler 22; Port Austin, Morris A240 in part. DELAWARE: Point Lookout, Canby 9. MaryLAND: Beltsville, Chase 3752; Riverdale, Chase 3642. District or CoLtumBia: House 907, Scribner in 1894, Ward in 1878. Virani: Patrick County, Heller 1312; Ocean View, Kearney 1447; Lee County, Small in 1892 (Gray Herb.). West Vrrernia: Harpers Ferry, Hitchcock 135. TENNESSEE: Lookout Mountain, Ruth in 1899 (Hitchcock Herb.). Fiq. 266.—Distribution of P. tsugetorum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 247 147. Panicum columbianum Scribn. Panicum heterophyllum Bosc; Nees, Agrost. Bras. 227. 1829, not Spreng. 1822. Based on ‘‘ Panicum heterophyllum Bosc, Herb. Willd.’ The type specimen, in the Willdenow Herbarium, labeled in Bosc’s hand, is the vernal form. Panicum columbianum Scribn. U.S. Dept. Agr. Div. Agrost. Bull. 7: 78. /.60. 1897. “Dry sandy fields, meadows and open woodlands, New England and southward to the Carolinas, and westward to Tennessee and Alabama, mostly near the coast; also in California.’’ The type, the specimen from which figure 60 is drawn, is in Hitchcock’s herbarium. It is labeled as follows in Scribner’s writing: ‘‘ Panicum columbianum Scribn. (Type) Brookland, D. C., July 14, 1894. Coll. F. L.-S.’”’ The specimen con- sists of three branching culms, 25 to 38 cm. high, the primary panicles destitute of spikelets. A duplicate type is in the National Herbarium. Panicum psammophilum Nash, Bull. Torrey Club 26: 576. November, 1899, not Welw. July, 1899. ‘‘New Jersry: * * * Toms River, July 25-31, 1898, W. N. Clute, no. 175 (type).’’ This specimen, in the herbarium of the New York Botanical Garden, consists of five branching culms, 15 to 40 cm. high, the primary panicles desti- tute of spikelets. The lower internodes are appressed-pilose, but otherwise the speci- men is very similar to Scribner’s type. This species has been erroneously referred to Panicum unciphyllum Trin. The specimen sent as a portion of the type from the Trinius Herbarium is P. columbianum, but a subsequent examination of Trinius’s plants showed that there were two groups of specimens lying loose upon a single sheet, one group being the type of P. unciphyllum (P. tenue Muhl.), the other group being P. columbianum. A plant of the latter had been sent with a copy of the label of the former. DESCRIPTION. Vernal plants light grayish green, often purplish; culms tufted, 15 to 50 cm., rarely 60 em., high, stiffly ascending, densely crisp-puberulent with long, ascending, crimped hairs commonly intermixed toward the base, but much less copiously than in P. tsugetorum,; sheaths shorter than the internodes, less pubescent than the culms, some- times puberulent between the nerves only; ligules less than 1 mm. long; blades firm, ascending or erect, 3 to 6 cm. long (seldom over 5 cm. long), 3 to 5 mm. wide, broadest at the rounded base, the serrulate, often white, cartilaginous margin usually involute toward the acuminate apex, typically glabrous on the upper surface but sometimes sparsely pilose toward the base, densely appressed-puberulent to glabrous beneath; panicles 2.5 to 4 cm. (rarely 5 cm.) long, about three-fourths as wide, the lower branches ascending, the axis and branches puberulent to nearly glabrous; spikelets 1.5 to 1.6 mm. long, 1 mm. wide, obovate, obtuse, turgid, densely short-pubescent; first glume one-third to scarcely half as long as the spikelet, acute or subacute; second glume and sterile lemma subequal, scarcely covering the fruit at maturity; fruit 1.3 mm. long, 0.9 mm. wide, broadly elliptic, obtuse. Autumnal culms branching from the middle and upper nodes at the maturity of the primary panicles, becoming widely spreading or decumbent at base, the early branches sometimes nearly equaling the primary culm, the ultimate branchlets in short, appressed fascicles, the crowded blades usually equaling or exceeding the reduced panicles; winter blades thickish, lanceolate, often sparsely pilose or ciliate. Specimens of this species and the preceding often closely resemble each other, since both vary much in pubescence and somewhat in habit. Panicwm columbianum Fig. 267.—P. columbianum. From type specimen. 4 Hitchcock, Bot. Gaz. 41: 66. 1906. 248 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. is typically smaller, more slender and stiffer than P. tsugetorwm, and the culms are densely crisp-puberulent with little of the pilose character of the latter. Occasional specimens, like Bissell 5596, Chase 3822, and Graves 10, are larger, laxer plants with appressed-pilose culms and are referred to P. columbianum because the spikelets are not over 1.6 mm. long. DISTRIBUTION. Sandy woods or open ground, Maine to Virginia.@ Marne: North Berwick, Parlin 1196; York Harbor, Bicknell in 1896. New Hampsuire: Laconia, Carter in 1902 (Hitchcock Herb.). MassacHusetts: Wellesley, Smith'738; Framingham, Smith 742, 744. Connecticut: Griswold, Graves 10; Waterford, Graves 83, 84; Southington, Andrews in 1902, Bissell 5596. New Yorxr: Van Cortlandt Park, Bicknell in 1891; Rockville Center, Bicknell in 1903 and 1906; Cedarhurst, Bicknell in 1903; Hempstead, Bicknell in 1903; Jamaica, Bicknell in 1905. New Jersey: Tuckerton, Chase Fig. 268.—Distribution of P. columbianum. 3606, 3613; Atsion, Chase 3533, 3539; Wildwood, Chase 3519; Toms River, Clute 175; Mount Arlington, Mackenzie 1399; Lakehurst, Mackenzie 2068; Chadwick, Mackenzie 2404; South Amboy, Mackenzie 2247; Sussex County, Mackenzie 2105. PENNSYLVANIA: Stroudsburg, Porter in 1898. MARYLAND: Patuxent River, Hitchcock 1632; Chesapeake Junction, Hitchcock 2407, 2413; Beltsville, Chase 3794, 3832; West Chevy Chase, Chase 5427, District oF CoLtumBIA: Chase 5429, 5430, Hitchcock 2418, Scribner in 1894, Williams in 1896. Virernia: Norfolk, Vasey in 1884 (Gray Herb.). 147a. Panicum columbianum thinium: Hitche. & Chase. Panicum unciphyllum thinium Hitche. & Chase, Rhodora 8: 209. 1906. ‘“‘Type Chase 3577 in National Herbarium. In mats, sandy, open ground, Tom’s River, N. J., July 28,1906; collected by Agnes Chase”’. The type consists of a tuft of many autum- nal culms 12 to 20 em. high, the spikelets 1.3 mm. long. Panicum columbianum thinium Hitche. & Chase in Robinson, Rhodora 10: 64. 1908. Based on P. unei- phyllum thinium Hitche. & Chase. DESCRIPTION. Vernal culms shorter and more slender than in P. colum- bianum, not over 30 cm. high, usually about 20 cm. high, densely tufted, the appressed pubescence longer, stiffer, and arising from minute papillee; sheaths sparsely ascending pilose; blades rarely over 3cm. long, sparsely pilose with long hairs on the upper surface at least near the margin Fig. 269.—P. columbianum thini- um. From type specimen. a The extended range given with the original description was based on misidentifi- cation of various small specimens of P. lindheimeri, as shown by such specimens labeled by Scribner ‘‘Panicwm columbianum” in the National Herbarium and in Hitchcock’s herbarium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 249 and base, the long hairssometimesmixed with appressed pubescence beneath; panicles 1.5 to 4 cm. long, about as wide; spikelets 1.3 to 1.4 mm. long, rounded obovate, very turgid, pubescent; second glume shorter than the fruit at maturity; fruit 1.1 mm. long, 0.8 mm. wide, obtuse. Autumnal form widely spreading, the branches appearing earlier than in the species, shorter and usually more crowded and somewhat aggregated toward the summit. A few specimens intermediate between the species and subspecies occur, as Kearney 10, District of Columbia, which has the habit and pubescence of the subspecies but spikelets 1.5 mm. long; Chase 3559, Atsion, New Jersey, and Commons 58, Rehoboth, Delaware, which have the stouter culms and crisped pubescence of the species but spikelets 1.4 mm. long. Short speci- mens with much crowded branches resemble P. oricola, from which they may be distinguished by the smaller spikelets and less dense pubescence. DISTRIBUTION. Dry sands, Massachusetts to Virginia. Massacuusetts: Nantucket, Bick- nell in 1899 and 1904. New Jersey: Mantoloking, Lyon in 1902; Atsion, Chase 3534, Fie. 270.—Distribution of P. columbianum thinium. 3560, Saunders & Clute2;Toms River, Chase 3577; Forked River, Chase 3588; Tuckerton, Chase 3605. Mary.tanp: Hyattsville, Chase 3806. Vireria: Lynn Haven, Hitchcock 406. 148. Panicum oricola Hitchc. & Chase. Panicum oricola Hitche. & Chase, Rhodora 8: 208. 1906. ‘‘Type Hitchcock 47 in National Herbarium. Prostrate clumps on bare sand on low mounds between marsh and sand dune. Lewes, Del., June 18, 1905, collected by A. S. Hitchcock.’’ This specimen is the early autumnal form. DESCRIPTION. Vernal form grayish, often purplish; culms densely tufted, 10 to 30 cm. high, spreading, densely appressed or ascending pilose, the hairs on the nodes spreading; sheaths usually more than half the length of the internodes, appressed-pilose; ligules 1 to 1.5 mm. long; blades firm, erect or ascending, 2 to5 cm. long, 2 to4 mm. wide, broadest near the base, acuminate, the upper surface pilose with hairs 3 to 5 mm. long, the lower surface appressed-pubescent with longer hairs intermixed; panicles short-exserted, or rarely long-exserted early in the season, 1.8 to 3 cm. long, rarely longer, about two-thirds as wide, rather densely flowered, the axis appressed-pubescent, the flexuous branches ascending or spreading; spikelets 1.5 mm. long, 1 mm. wide, broadly obovate, turgid, obtuse, pubescent with short spreading hairs; first glume one-third to half the length of the spikelet, abruptly pointed; second glume and sterile lemma barely equaling the fruit at maturity; fruit 1.3 mm. long, 0.9 mm. wide, broadly elliptic, very turgid. Fig. 271.—P. oricola. From type specimen. 250 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Autumnal form prostrate, forming mats, with short, fascicled branchesat all the nodes, the branches appearing before the maturity of the primary panicles; leaves and panicles not greatly reduced, the latter overtopped by the blades, which are less pilose than earlier ones; winter blades sparsely pilose above. This species is more copiously pubescent than any other in this group and in this character resembles species of the Lanuginosa, but its affinity seems to be rather with the Columbiana. DISTRIBUTION. Sand barrens along the coast, Massachusetts to Virginia. MassacuusetTts: Nantucket, Bartlett 1368, 1379, Bicknell in 1906, Hitchcock 558; Plymouth, Oakes; Quanquisset, Bartlett 1327. ConneEcTICUT: Groton, Bissell 9306 (Bissell Herb.). New Yorks: Northville, Young in 1871; West Rockaway, Bicknell in 1903; Lawrence, Bicknell in 1906; Hempstead, Bicknell in 1903; Woodmere, Bicknell in 1902; Rockville Center, Bicknell in 1906; Valley Stream, Bicknell in 1905. New Jersey: Absecum, Commons 45; Tuckerton, Chase 3609; East Plains, Stone 8; Atlantic City, Parker; Hammonton, Canby in 1902; South Amboy, Mackenzie 1355; Atsion, Chase 3562; Toms River, Chase 3576; Forked River, Chase 3581. DELAWARE: Rehoboth, Commons 59, 60; Lewes, Hitchcock 47. Virainia: Ocean View, Coville 13, 14, Kearney 1461; Cape Henry, Chase 5431. Fig. 272.—Distribution of P. oricola. Sphaerocarpa.—Culms usually few in a tuft, rather stout, glabrous; ligules obsolete or nearly so; blades mostly thick and firm, cordate and ciliate at base, mar- gins strongly cartilaginous; spikelets obovoid-spherical at maturity, oval when young, | to 1.8 mm. long, puberulent; second glume and sterile lemma 5 to 7-nerved; panicle branches mostly viscid. Autumnal form remaining simple or but sparingly branching, the thick, white-margined blades of the winter rosette conspicuous. Culms spreading; blades obscurely nerved; panicle nearly as broad as long. Ligules obsolete or wanting; blades lanceolate. ....... -- 149. P.sphaerocarpon. Ligules evident, 0.3 to 1 mm. long; margins of blades parallel for at least two-thirds their length. ....... 149a. P. sphaerocarpon inflatum. Culms erect or ascending; blades rather strongly nerved; panicle never more than two-thirds as broad as long, usually less. Spikelets 1.5 to 1.6 mm. long; blades lanceolate, the WPHeL MOL PEMNCCUn fo o5.. fo. ons ee ee 150. P. polyanthes. Spikelets 1 to 1.2 mm. long; blades tapering from base to apex, the upper much smaller than the lower......151. P. erectifoliwm. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 251 149. Panicum sphaerocarpon Ell. Panicum sphaerocarpon Ell. Bot. S.C. & Ga. 1: 125. 1816. ‘‘GrowsinGeorgia. Dr. Baldwin.’’ The type, in the Elliott Herbarium, consists of a single plant with a mature, long-exserted panicle. The accompanying label reads: ‘‘ Panicum sphaero- carpon Hab. Georg. Dr. Baldwin.”’ Panicum kalmii Swartz, Adnot. Bot. 6.1829. ‘‘Hab. in America boreali (Pennsyl- vania?): Prof. Kalm.’’ The type, labeled ‘‘Panicum Kalmii Swartz in Adnot. Bot. p. 6,’ in the Swartz Herbarium, is a single vernal plant. Panicum heterophyllum Swartz, Adnot. Bot. 6. 1829, not Spreng. 1822. This is mentioned as a synonym of P. kalmii Swartz. Panicum dichotomum sphaerocarpum Wood, Class-book ed. 3. 786. 1861. Presum- ably based on P. sphaerocarpon Ell., though the description hardly applies to this species. Panicum nitidum crassifolium Gray; Doell in Mart. Fl. Bras. 2?: 247. 1877. This is described from a ‘‘specimen in New-Jersey lectum, n. 30.’’ Doell’s plant is evi- dently one of the specimens distributed by Gray in Gramineae and Cyperaceae 1: no. 30. 1834, under the above name. The specimen in the Gray Herbarium bears the data ‘‘Hab.—Pine barrens of New-Jersey.’’ This was not described by Gray. It represents the slender form of P. sphaerocarpon. Panicum microcarpum sphaerocarpon Vasey, Grasses U. 8. 12. 1883. Based on “*P. sphaerocarpon, Ell.’’ Panicum vicarium Fourn. Mex. Pl. 2: 20. 1886. Fournier cites only one specimen, “Cordova (ScHAFFN. n. 285).’’ The type is in the Paris Herbarium. This name was earlier listed by Hemsley@ without description. DESCRIPTION. Vernal plants light green, in tufts of few to several culms, 20 to 55 em. high, radiate- spreading, occasionally nearly erect, the nodes appressed-pubescent; sheaths nearly as long as or longer than the comparatively short internodes, loose toward the summit, ciliate on the margin, otherwise glabrous, sometimes with viscid tubercles between the nerves; ligules nearly or quite obsolete; blades thick and firm with usually incon- spicuous nerves, ascending, 6 to 10 cm. long, 7 to 14 mm. wide (rarely longer or wider), the upper and lower smaller, acuminate, slightly narrowed to the subcordate base, rough on the upper surface, smooth below, the cartilaginous, scabrous margins stiffly ciliate toward the base; panicles long-exserted, 5 to 10 cm. long, nearly as wide, rather loosely flowered, the axis and ascending branches with viscid spots; spikelets 1.6 to 1.8 mm. long, 1 to 1.3 mm. wide (in exceptional specimens only 1.5 mm. long), obovoid-spherical at matu- rity (oval when young), puberulent; first glume about one-fourth the length of the spikelet, obtuse; second glume and sterile lemma equaling the fruit at maturity; fruit 1.4 to 1.5 mm. long, 1 to 1.2 mm. wide, obovoid-spherical. Autumnal form prostrate-spreading, sparingly branching late in the season*from the base or lower and middle nodes, the branches short, mostly simple, the blades and panicles not greatly reduced; winter rosettes of many thick, ovate or ovate- lanceolate, white-margined leaves, appearing early. Numerous specimens occur which are intermediate between this species and the following subspecies. These more slender plants with usually narrower blades and slightly smaller spikelets are the form named Panicum nitidum var. crassifolium by Fic. 273.—P. sphaerocarpon. From type specimen. @ Biol. Centr. Amer. Bot. 3: 498. 1885. 252 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Gray @ as shown by the specimen from ‘‘Pine barrens of New-Jersey,’’ in the Boott set of the Gramineae and Cyperaceae now in the Gray Herbarium. This varietal name was never published by Gray and P. nitidwm was included under P. dichotomum in Gray’s Manual from the first to the fifth editions. In the sixth edition the name P. nitidum was applied to P. sphaerocarpon Ell., (which was cited as synonym) as shown by the description and by specimens in the Gray Herbarium. The interme- diate specimens, of which the following are representative, are included in those cited below: Ball 69, Bebb 1259, Chase 3089, 3256, 3489, 3611, Hitchcock 1216, 1607, Mackenzie 2166, Plank 49, Tracy 13, 4607. DISTRIBUTION. Sandy soil, Vermont to northern Florida, west to Illinois and Texas, and south through Mexico to Venezuela. Vermont: Putney, Blanchard 9 (Gray Herb.). MassacuHuseEtts: Boston, Swan in 1886; Framingham, Smith 748. Connecticut: Southington, Andrews 8, 73; Waterford, Graves 85; Berlin, Bissell 5585; Portland, Wilson 1435. RaopDE Istanp: Providence, Olney in 1868 (Brown Univ. Herb.). New York: Niagara County, Townsend 1; Long Island, Bicknell in 1905. New Jersey: Berkeley Heights, Mackenzie 2250; Tuckerton, Chase 3611; Wild- wood, Chase 3489, 3509; Cape May, Pollard in 1897; South Amboy, Mackenzie 2166. PENNSYLVANIA: Easton, Porter in 1893 and 1897; Stroudsburg, Porter in 1898; Germantown, Stone 6; Newtown, Smith 160. Onto: Portsmouth, Kellerman in 1899 (Ohio State Univ. Herb.). Inpiana: Dune Park, Chase 1850, V. H. Chase 242. Iuu1Nots: Jackson, French in 1906. MicuiaAN: Detroit, Farwell 1388; Grand Beach Springs, Hill 88 and 90 in 1908; Magician Lake, Umbach 2153. Missourt: St. Louis, Hggert 254; Cliff Cave, Kellogg 9, 11; Swan, Bush 675; Eagle Rock, Bush 145. Kansas: Cherokee County, Hitchcock Pl. Kans. 883. DeLawakeE: Centerville, Commons 284, 298; Milton, Commons 350; Rehoboth, Commons 49, 50; Lewes, Hitchcock 561. MaryYLAND: Chesapeake Beach, Chase 3256, Hitchcock 1605, 1607, 1615, 1616, 2408; between Chesapeake Beach and Chesapeake Junction, Hitchcock, 1626, 1639. District oF CotumsBtia: Ball 65, Chase 2401, 2412, Kearney 12, Pollard 398, Vasey 34, 111, Ward in 1878, Williams 1, 2. Virainia: Alexandria, House 1058, Chase in Kneucker Gram. Exs. 553; Four- Mile Run, Chase 5432, 5433; Norfolk County, Chase 2332, Kearney 301 in part, 1560; Munden, Mackenzie 1663; Virginia Beach, Williams 3098. Nortu Carouina: Roanoke Island, Chase 3212, 3237; Lake Mattamuskeet, Chase 3206; Wilmington, Chase 4591, Hitchcock 1426; Chapel Hill, Chase 3074; Caraleigh Junction, Chase 3089; Biltmore, Biltmore Herb. 4292b. Fig. 274.—Distribution of P. sphaerocarpon. aGram. & Cyp. 1: no. 30. 1834. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 253 Sout Carona: Orangeburg, Hitchcock 1381; Pacolet, House 2492; Clemson College, House 2413; Aiken, Kearney 243, Ravenel. Georera: Augusta, Cuthbert 1157, 1158; Kearney 217; Lookout Mountain, Ruth 1, 6, 75; Union, Harper 1087; Millen, Curtiss 6828; Thomson, Bartlett 1459. Fioripa: Monticello, Combs 306; St. Andrews Bay, Tracy 9137. Kentucky: Bell County, Kearney 594. TENNESSEE: Cocke County, Kearney 968. AxaBaMA: Pisgah, Chase 4470; Valley Head, Ruth 21; Tuskegee, Carver 28, 67; Cullman County, Eggert 13, 23; Anniston, Tracy 7402; Mobile, Kearney 25; Springhill, Langlois 43. Mississterr: Starkville, Tracy 47; Jackson, Hitchcock 1307; Enterprise, Tracy 3296 in part; Biloxi, Hitchcock 1078, 1084, Kearney 320, Tracy 4576, 4597; Ocean Springs, Tracy 13; Mississippi City, Hitchcock 1095, 1099, 1102; Moss Point, Tracy 4604; Cat Island, Tracy 433; Petit Bois Island, Tracy 4607. ARKANSAS: Little Rock, Coville in 1887; Miller County, Eggert 115; Benton County, Plank 49, 99. Loutstana: Calhoun, Ball 69; Hitchcock 1284; Shreveport, Hitchcock 1241; Breton Island, Tracy & Lloyd 468; Lake Charles, Hitchcock 1126, 1132, 1134, 1136, 1137. Texas: Waller County, Hitchcock 1181, 1190, 1213, 1216, Thurow 2, 4, 10, 30; Houston, Bebb 1259; Hall 832; Denison, Bebb 1434; Terrell, Warburton 12; Columbia, Bush 177; Weatherford, Tracy 7946; Victoria, Plank 4; Milano, Griffiths 6551; Galveston, Plank 34. Oxtanoma: Sapulpa, Bush 1218; Stillwater, Hitchcock 562.. Mexico: San Luis Potosi, Schaffner 1037; Jalapa, Pringle 7883 (Hitchcock Herb.) 8344; Chinantla, Liebmann 327; Orizaba, Botteri; Schiede & Deppe ‘‘acumi- natum b” (Berlin Herb.), Schaffner 138 (Paris Herb.); Cérdoba, Schaffner 285 (Paris Herb.). GuATEMALA: Coban, Tuerckheim 56 in part; crater of Pacaya Volcano, Keller- man 6236. : Costa Rica: Abejonal, Tonduz 7878; San Pedro de la Calabaza, Tonduz 10745 in part; Copey, Tonduz 11866. VENEZUELA: Fendler 1638. 149a. Panicum sphaerocarpon inflatum (Scribn. & Smith) Hitche. Panicum inflatum Scribn. & Smith, U. 8. Dept. Agr. Div. Agrost. Cire. 16: 5. 1899. “Type No. 4622, 8S. M. Tracy, collected at Biloxi, Miss., October, 1898.’’ The type, in the National Herbarium, is a branching plant, decumbent and rooting at the lower nodes, the loose sheaths prominently tuberculate, the loosely flowered panicles mature. Panicum mississippiense Ashe, Journ. Elisha Mitchell Soc. 16: 91. 1900. ‘‘Col- lected by me on the banks of the Mississippi River below New Orleans in October. I also refer here S. M. Tracy’s No. 6777, collected on Horne Island, Miss., in July, 1899.’ The type specimen could not be found in Ashe’s Herbarium. According to Tracy the number of the second specimen mentioned is a misprint for 6471. This specimen is in Ashe’s herbarium. The culms are in the early branching state, slender and widely spreading, the branches elongated. DESCRIPTION. Vernal form similar to that of the species, more ascending, not radiate-spreading; culms on the average taller, more slender; sheaths rather looser, more commonly and prominently viscid-tuberculate; ligules 0.3 to 1 mm. long; blades narrower, 5 to 10 254 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. mm. wide, the margins nearly parallel for two-thirds their length, with fewer ciliz at the base; panicles more loosely flowered; spikelets slightly smaller, 1.4 to 1.5 mm. long, 1 mm. wide. Autumnal form decumbent, rather freely branching from the middle nodes before the maturity of the primary panicles, these early branches long and again branching more freely than in the species, the ultimate blades and panicles not greatly reduced. This subspecies is distinguished by the ligules, slightly smaller spikelets, and nar- rower, parallel-margined blades, taken in combina- tion, and in autumnal specimens by the more freely branching habit. The specimens cited below all show this combination of characters, but about half as many specimens occur which are intermediate between this and the species. These bear a gen- Fig. ee . sphaerocarponinflatum. era] resemblance to the subspecies, having spikelets rom type specimen. about 1.5 mm. long, and narrower, but not always parallel-margined blades, but with no ligule or the merest trace of one. Because of the large proportion of these intermediate specimens P. inflatum Scribn. & Smith is here reduced to a subspecies of P. sphaerocarpon. DISTRIBUTION. Moist sandy ground, Maryland to Florida, and west along the Gulf to Texas, thence north to Missouri. Missourt: Monteer, Bush 747 in part, 753. MaryLANb: Owings, Hitchcock 1618; Chesapeake Junction, Hitchcock 2412. NortH Carona: Wilmington, Chase 3134, 3158. Soutn CAROLINA: Orangeburg, Hitchcock 26. GeEorGIA: Savannah, Kearney 188, 194; Americus, Tracy 3642; Thomasville, Tracy 3656. Frorwwa: Lake City, Combs 182; Quincy, Combs 403, 406; St. Vincent, Tracy 6458. AzaBaMa: Selma, Kearney 7; Fort Morgan, Tracy 8400. Fic. 276.—Distribution of P. sphaerocarpon inflatum. Mississippi: Biloxi, Tracy 4593, 4622; Centerville, Tracy 3619; Horn Island, Tracy 2862, 6471. Lovurstana: Calhoun, Hitchcock 1285; Alexandria, Ball 441, 536; Calcasieu, Cocks 3007; Lake Charles, Chase 4429. Texas: Without locality, Nealley in 1890. OxtaHoma: Poteau, Hitchcock in 1903 (Hitchcock Herb.). 150. Panicum polyanthes Schult. Panicum multiflorum Ell. Bot. 8. C. & Ga. 1: 122. 1816, not Poir. 1816. Elliott gives no exact locality, but his specimen was presumably from the vicinity of Charles- ton as he merely states, ‘‘Grows in shaded, dry soils.’” The type, in the Elliott Herba- rium, consists of a single culm, lacking base, bearing three leaves and an immature panicle, slightly included at base. The accompanying label reads: ‘Panicum multi- florum mihi. Hab. in umbrosis. Flor. May-Jun.”’ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 255 Panicum microcarpon Muhl. Descr. Gram. 111. 1817, not Muhl.; Ell. 1816.¢ ‘‘Semina e Virg. et Cherokee et Delaware.’’ The type in the Muhlenberg Herbarium consists of the upper portion of a culm with two leaves and a short-exserted panicle. The attached label reads: ‘‘40, c Jul. 12. e Cherokee.”’ Panicum polyanthes Schult. Mant. 2: 257. 1824. Based on P. multiflorum Ell. not Poir. That Poiret’s use of the name was earlier is given on the authority of Sprengel. Panicum microcarpon isophyllum Scribn. Tenn. Agr. Exp. Sta. Bull. 7:51. /. 54. 1894. No specimen is cited but in the Scribner Herbarium is a sheet to which is attached a note from Dr. Chapman suggesting the name ‘‘isophyllum” and upon which are four small specimens of P. polyanthes one of which is recognized as the plant figured with the original description. The specimens were collected by E. E. Gayle, Alleghany Springs, Blount Co., Tennessee, August, 1890. DESCRIPTION. Vernal plants light green, in tufts of few to several culms, 30 to 90 cm. high, stout, erect, the nodes glabrous or nearly so; sheaths long, usually overlapping, finely ciliate on the margin, otherwise glabrous; ligules obsolete or wanting; blades rather thin, prominently nerved, ascending, 12 to 23 cm. long, 15 to 25 mm. wide, the upper seldom reduced, long-acuminate, scarcely narrowed toward the cordate base, rough or smooth on the upper surface, smooth below, the cartilaginous, scabrous margin ciliate toward the base; panicles exserted, 8 to 25 cm. long, one-fourth to half as wide, densely flowered, the lower branches narrowly ascending, often distant, the upper fascicled, spikelet-bearing to the base; spikelets 1.5 to 1.6 mm. long, 1 to 1.1 mm. wide, obovoid-spherical at ma- turity, minutely puberulent; first glume one-third to two-fifths the length of the spikelet, obtuse or obscurely pointed; second glume and sterile lemma equaling the fruit at maturity; fruit obovoid- spherical. Autumnal form remaining erect and simple or producing from the lower or middle nodes simple branches with smaller blades and panicles; winter rosettes like those of P. sphaero- carpon, but the leaves larger. This species is distinguished from P. sphaerocarpon by its erect habit, taller, more leafy culms, wider blades and narrow panicles. Specimens not infrequently occur in which, from a twisting of the internodes, the blades are all or mostly on one side. This is especially true of small, late culms. It was tosuch aspecimen the name P. micro- carpon tsophyllum was given. Fig. 277.—P. polyanthes. From type specimen of P. multiflorum Ell. DISTRIBUTION. Damp ground, woods and openings, New Jersey to Oklahoma, south to Georgia and Texas. New Jersey: South Amboy, Mackenzie 1380. PENNSYLVANIA: Lancaster County, Heller 4772, Porter in 1898. Onto: Lancaster, Kellerman 6767. InprANA: Clarke County, Deam 5392; Batesville, Deam 6815. Inuino1s: Cobden, Earle in 1886; Jackson County, French in 1905. _ Missourt: St. Louis, Lggert 250; Pleasant Grove, Bush 232. @ See discussion under P.. microcarpon Muhl.; Ell., page 181. b Neu. Entd. 2: 190. 1821. | 256 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DELAWARE: Wilmington, Commons 307; Townsend, Canby in 1896; Ogletown, Commons 47; Stanton, Commons 306. MarytAnp: Chesapeake Beach, Hitchcock 1609; Chesapeake Junction, Hitchcock 2399; Riverdale, Chase 2368; Hyattsville, Chase 3803, House 450; West Chevy Chase, Chase 3273. District or CotumBIA: Ball 21, 705, Chase in Kneucker Gram. Exs. 554, Hitch- cock 2404, Kearney 13, 20, Merrill 199, Pollard 401, Vasey 110, Ward in 1880. Virarnia: Four-Mile Run, Steele in 1897; Munden, Mackenzie 1745; Ocean View, Kearney 1476; Suffolk, Heller 968; Norfolk, Hitchcock 410. West VirGiniA: Baileysville,. Morris 1186. Norrs Carona: West Raleigh, Coit 1300; Chapel Hill, Ashe; Caldwell County, Small & Heller 463; Hickory, Small & Heller in 1891. Sout Carona: Abbeville, Maier 264 (Gray Herb.). GeoraiA: Lookout Mountain, Ruth 5, 7, 15, 69; Stone Mountain, Small in 1893; Clarke County, Harper 104; Cobb County, Wil- son 8; Thomson, Bartlett 1071, 1498, 1500. Kentucky: Harlan County, Kear- ney 52. TENNESSEE: Robertson County, Eggert 95; Cocke County, Kearney 974; Chester County, Bain 191; White © Cliff Springs, Scribner in 1890. AtaBAmMA: Cullman County, Eggert 14; Pisgah, Chase 4485; Nesheka, Carver 18. MississipPi: Taylorville, Tracy 8418; Heidelberg, Tracy 3316; Centerville, Tracy 3631; Macon, Tracy 3238; Starkville, Kearney 57 in part; without locality, Tracy 3760. ARKANSAS: Greene County, Eggert 240; Prescott, Bush 255. LouIstaNna: Calhoun, Hitchcock 1270. Texas: Palestine, Plank 89; Burnet, Plank 9; without locality, Nealley in 1884 and 1886. OxKLAHoMA: Poteau, Hitchcock in 1903 (Hitchcock Herb.). Fic. 278.—Distribution of P. polyanthes. 151. Panicum erectifolium Nash. Panicum sphaerocarpon floridanum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 33. 1889, not P. floridanum Trin. 1835. ‘‘Florida.’’ Inthe National Herbarium are three specimens from Florida marked in Dr. Vasey’s writing ‘‘Panicum sphaerocarpon Ell. var. Floridanum.’’ Of these Curtiss 3599, from ‘‘ Moist pine barrens, Mosquito Inlet, Florida, May, 1879,’’ has been chosen as the type, since with the other two no locality within the State is given. This specimen consists of two plants 45 and 50 cm. high, with mature panicles of densely puberulent spikelets. Panicum erectifolium Nash, Bull. Torrey Club 23: 148.1896. Based on P. sphaerocarpon floridanum Vasey, not P. floridanum Trin. Panicum floridanum Chapm. Fl. South. U. 8. ed. 3. 585. 1897, not Trin. 1835. ‘In and around shallow ponds, near the coast of West Florida.’’ The type, in the Chapman Herbarium at Biltmore, is from Apalachicola. Chapman presumably intended to base his name upon P. sphaerocarpon floridanum Vasey, but cites ‘‘(P. microcarpon, var., Vasey. P. sphaerocarpum, Flora.)” HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 257 DESCRIPTION. Vernal plants dull green, sometimes bluish, in tufts of few to several culms, 30 to 70 em. high, erect or ascending, usually stout, glabrous, including the nodes; sheaths, except the uppermost, short, rather loose, usually crowded and overlapping at base, ciliate on the margin, otherwise glabrous; ligules about 0.3 mm. long; blades thick and firm with inconspicuous veins, ascending or erect, 7 to 13 cm. long, 6 to 12 mm. wide, the crowded lower ones usually much larger than the others, these suc- cessively smaller upward, tapering from the cordate base to the acuminate apex, smooth on both surfaces, margins scabrous, stiffly ciliate toward the base; panicles exserted, 6 to 12 cm. long, half to two-thirds as wide, densely flowered, the lower branches usually narrowly ascending; spikelets 1 to 1.2 mm. long, 0.8 to 1mm. wide, broadly ovate or subspherical, densely puberulent; first glume one-fifth to one-fourth the length of the spikelet; obtuse, second glume and sterile lemma equaling the fruit at maturity; fruit oval, very obscurely umbonate. Autumnal form remaining erect and simple or late in the season producing branches from the third or fourth node, the branches nearly as long as the primary culm, rarely again branching; winter rosettes appearing late, the blades 3 to 10 cm. long. This species shows an unusual variation in the size of the blades and also the num- ber of leaves to the culm. There are usually 5 to 7, but sometimes as many as 10 leaves, and in a few specimens no blades are over 5 cm. long. The specimens of Combs 553 are small autumnal plants more freely branching than usual. Fic. 279.—P. erectifolium. From type specimen. DISTRIBUTION. Moist pine barrens, swamps, and borders of ponds, North Carolina to Florida and Louisiana; also in Cuba. Norta Caroura: Wilmington, Chase 3137, Hitchcock 411; without locality, McCarthy in 1885. GeoreiA: Allapaha, Curtiss 6817 in part; Huntington, Harper 1394; Charlton County, Har- per 1485. Fioriwa: Jacksonville, Curtiss 4812 (Hitchcock Herb.); Bald- win, Combs 58; Lake City, Combs 114; De Funiak Springs, Combs 467; Washing- ton County, Combs 553, 662; Ellzey, Combs 814; Mosquito Fia. 280.—Distribution of P. erectifolium. Inlet, Curtiss 3599; Grasmere, Combs 1117; Eustis, Nash 1012; Manatee River, Rugel 229; Orange Glade, Eaton 578; Myers, Chase 4154, Hitchcock 874, Lee Co. Pl. 469. Mississippi: Beauvois, Tracy 4596. LourstaANA: St. Tammany Parish, Cocks 292 (Hitchcock Herb.). Cupa: Vuelta Abajo, Wright 3462. 41616°—voL 15—10——17 258 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Ensifolia.—Plants low and slender, usually glabrous and less than 50 cm. high; sheaths glabrous or puberulent (sparsely pilose in P. curtifolium); ligules nearly obsolete (about 1 mm. long in P. curtifolium); panicles small, rarely more than 5 cm. long; spikelets glabrous or pubescent, 1 to 1.7 mm. long, 5 to 7-nerved. Autumnal form sparingly to freely branching, or in P. vernale a distinct autumnal form wanting. Ligules about 1 mm. long; sheaths or some of them sparsely [SfaTg eran Guba) 676) =) 5 Nye amy peerage tn Me § Ee 159. P. curtifolium. Ligules obsolete or nearly so; pubescence if present not spreading. Blades prominently white-margined, firm; spikelets densely puberulent. Blades puberulent beneath, often above; sheaths and sometimes lower internodes ascending RWUCRCOINE <. oC. - cence noe sok meee eee 152. P. tenue. Blades glabrous; sheaths glabrous or minutely ciliate only. Uppermost culm blades much reduced; culms branching from lower nodes only, the branches repeatedly branching........... 153. P. albomarginatum. Uppermost culm blades about as long as the others; culms bearing short branches from the upper and middle nodes.......-.-..-- 154. P. trifolium. Blades not white-margined or very obscurely so (or if white margin is evident spikelets only 1.1 mm. long); spikelets glabrous or puberulent. Culms branching only at base; plants soft, light green.158. P. vernale. Culms branching at the nodes; plants firm or at least not soft. Spikelets glabrous. Spikelets 1.1 to 1.2 mm. long; blades rarely as much a5, cm, jOnes eee coe eee 160. P. chamaelonche. Spikelets 1.2 to 1.5 mm. long. Blades elongated, at least some of them $ to 10 cm. fonesc 2k ence 161. P. glabrifolium. Blades not over 3 cm. long.....-..- RS 157. P. ensifolium. Spikelets puberulent. Spikelets 1.1 mm. long; winter blades bluish green, not glossy /.450 =. .2.5= see eeaek 156. P. concinnius. Spikelets 1.3 to 1.5 mm. long. Blades involute, faleate, with long stiff hairs on margin near base; plants ethifiand withiunsotoeda. eer eee 162. P. breve. Blades not involute, or at tip only, not falcate. Plants bright green; winter blades conspicuous, glossy green... . . 155. P. flavovirens. Plants olive; winter blades not con- spicuous nor glossy....-.-.-- 157. P. ensifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 259 152. Panicum tenue Muhl. Panicum tenue Muhl. Descr. Gram. 118. 1817. No locality isgiven. ‘‘P. deustum Brickell et Enslin,’’ an unpublished name, is cited as synonym. The type, in the Muhlenberg Herbarium, consists of three plants with attached label bearing the name ““Panicum deustum;’’ these are from 10 to 30 cm. high, the panicles immature, the sheaths sparsely appressed-pilose, the blades puberulent on the lower surface and with conspicuous white margins. Panicum deustum Brickell & Enslin; Muhl. Descr. Gram. 119. 1817, not Thunb. 1794. Thisherbarium name isgiven asasynonym of P. tenue, of which it isatyponym. Panicum liton Schult. Mant. 2: 250.1824. Based on P. tenue Muhl., Muhlenberg’s description, slightly rearranged, being copied, “‘ Nomina mutanda, ob tenue Roxb. et deustum Thunb. antiquiora.’’ Panicwm tenue Roxb. 1813,4 is a nomen nudum, the description not being published until 1820.9 Panicum unciphyllum Trin. Gram. Pan. 242. 1826. Trinius states in regard to his specimen, ‘‘V.spp. Am. Bor. (Tratrinick).’’ The type, in the Trinius Herbarium, is the vernal form, with sparsely appressed-villous culms and sheaths and puberulent blades. Asstated under P. columbianum, Trinius’s species was misunderstood because the specimen sent from St. Petersburg as a part of the type proved on a subsequent examination of Trinius’s herbarium to be not the type but a specimen of P. colum- bianum which was on the same sheet with the type specimen. The label accompany- ing the latter reads, ‘‘Panicum unciphyllum m. Pan. heterophyllum Muhl. (teste Nees) an Pluckn. Tay. 92 f. 8, ex herb. Enslini, spmna Am. bor. Trattinick.”’ Panicum macrum Kunth, Rév. Gram. 1:40. 1829. Based on P. tenue Muhl., the name presumably changed because of P. tenwe Roxb. Panicum parvulum Muhl.; Scribn. & Merr. U. 8. Dept. Agr. Diy. Agrost. Oire. 27: 4. 1900, not Trin. 1834. This name, found in Muhlenberg’s herbarium, is given as a synonym of P. tenue. DESCRIPTION. Vernal form olive green; culms in rather small tufts, 20 to 55 cm. high, slender, erect from a more or less geniculate base, glabrous, or the lower internodes sparsely appressed-pubescent, the nodes glabrous, appressed-pubescent, or appressed-pilose; sheaths usually much shorter than the internodes, puberulent between the nerves to sparsely appressed-pilose, or the upper glabrous; ligules 0.3 to 0.5 mm. long, dense; blades distant, ascending or speading, 2 to 5 cm. long, 3 to 4 mm. wide, rather thick and with a cartilaginous, often white, margin, involute-pointed, usually densely puberulent beneath, glabrous on the upper surface or puberulent toward the base; panicles long-exserted, 3 to 5 cm. long, about as wide, pyramidal, open, rather few- flowered, the flexuous branches spreading; spikelets 1.6 to 1.7 mm. long, elliptic, subobtuse, densely puberulent; first glume one-fifth as long as the spikelet or less, obtuse; second glume shorter than the fruit and sterile lemma; fruit 1.4 to 1.5 mm. long, elliptic, subobtuse. Autumnal culms erect or leaning, sparingly branching from the middle nodes, the branches in small fascicles, shorter than the primary inter- nodes, the blades not much reduced; winter rosette con- spicuous, the thick, cartilaginous-margined, involute-pointed blades 3 to 5 cm. long, 4 to 7 mm. wide, persistent (but usu- ally dead) during the succeeding year. This species seems to be intermediate between P. albo- marginatum and P. ensifolium, differing from the first in being pubescent and in having taller, more slender culms, sparingly branched. From P. ensifolium it may be distinguished by the larger, Fig. 281.—P. tenue. From type specimen. 4 Cat. Fl. Ind. 1813. 6 Roxb. Fl. Ind. 1: 313. 1820. 260 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. more pubescent spikelets, the thicker, involute-pointed blades and the large basal rosette of firm leaves. In Hitchcock 1438, from Wilmington, N. C., referred here, the pubescence is so copious as to sug- gest P. leucothrix, but the nearly obso- lete ligule and the size of the spikelets place it, though somewhat doubtfully, in P. tenue. Hitchcock’sno. 1467 is an unusually robust specimen with pani- cles as much as 9 cm. long. DISTRIBUTION. Moist sandy woods, eastern North Carolina and northern Florida. Nort Carouina: Parmele, Ashe Fig. 282.—Distribution of P. tenue. in 1899; Manteo, Ashe in 1898; Wards Mill, Chase 3170, 3172, 3183; Wilmington, Ashe in 1899, Hitchcock 332, 1467. Fioriwa: Lake City, Bitting 20. 153. Panicum albomarginatum Nash. Panicum albomarginatum Nash, Bull. Torrey Club 24: 40. 1897, ‘‘Collected by the writer in low pine land at Eustis, Lake County, Florida, early in June, 1894, no. 925.’’ The type, in Nash’s herbarium, consists of two large tufts in the early branching state, the culms 15 to 28 cm. high, the primary panicles devoid of spikelets. DESCRIPTION. Vernal plants usually grayish green, often purplish; culms densely tufted, 15 to 40 cm. high (rarely taller), slender but firm, ascending or spreading, glabrous includ- ing the nodes; leaves crowded at the base, distant above, sheaths sometimes pubescent on the margin and at the summit, otherwise glabrous, or the lowermost sometimes obscurely pubescent; ligules 0.3 mm. long, dense; blades firm, those of the midculm 4 to 6 cm. long, 4 to 6 mm. wide, rounded at the base, thick and firm, with a prominent white, finely serrulate, cartilaginous margin, ascending or spreading, glabrous, the crowded basal blades as much as 11 cm. long, and the uppermost blade usually much reduced; panicles finally long-exserted, 3 to 6 cm. long, nearly as wide, rather densely flowered, the flexuous branches ascending or spreading; spikelets 1.4 to 1.5 mm. long, 0.7 mm. wide, obovate-elliptic, subobtuse, turgid at maturity, densely puberulent; first glume one-fifth to one-fourth as long as the spikelet, obtuse or subacute; second glume and tie pase ena sterile lemma scarcely equaling the fruit at maturity; fruit tum. From type speci- 1-25 mm. long, 0.65 mm. wide, elliptic, subacute. men. Autumnal form spreading, the primary culms branching from the base and lower nodes, these early branches much longer than the primary internodes and repeatedly branching, forming bushy tufts, the ultimate branchlets and reduced blades appressed; winter blades stiffly erect or spread- ing, very smooth and firm. This species is distinguished by the long crowded basal and distant upper blades, the uppermost usually less than half as long as those of the midculm; and by the autumnal form in which the primary culms branch from the basal and lower, never from the upper, nodes. The specimens collected by Hitchcock in Cuba (no. 555) are robust plants and differ from typical P. albomarginatum in having a ligule 1 mm. long. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 261 DISTRIBUTION. Low sandy soil of the Coastal Plain, from southeastern Virginia to Florida and west to Louisiana; also in Guatemala and Cuba. Virernia: Dismal Swamp, Chase 3658. Norra Carona: Parmele, Ashe in 1899; Wilmington, Hitchcock 1428, 1429, 1434, 1440, Kearney 268. Sours Carona: Orangeburg, Hitchcock 1369, Aiken, Ravenel. Froriwa: Baldwin, Hitchcock 990; Lake City, Combs 112, Hitchcock 1021, 1022; Bay Head, Combs 650; Old Town, Combs 854; Perdido Bay, Tracy 8409; Titusville, Chase 3966, Hitchcock 761; San- ford, Hitchcock 768, 823, 826; Eustis, Chase 4043, Hitchcock 817, Nash 925; Orange County, Baker 119; Lemon Bay, Tracy 7189; Miami, Hitchcock 639, 666, 667, 670, 679, 714, 720; Homestead, Hitchcock 692; Tampa, Hitchcock 945; Lake- land, Hitchcock 838, 839, 848; Braidentown, Hitchcock 949, 963, Tracy 6733; Manatee, Rugel 184; Myers, Chase 4151, Hitchcock 870, 876, 880, 882, 884, 886. AtaBAMA: Tuskegee, Carver 97. Mississippi: Biloxi, Tracy 4605 in part (Gray Herb.). LoursiANa: Caleasieu River, Langlois 42 in 1884. GuaTeMALA: Between Gualén and Copan, Pittier 1805a. Cusa: Herradura, Hitchcock 555; Pinar del Rio, Wright 3463 in part (Sauvalle Herb.); Isle of Pines, Taylor 32. 154. Panicum trifolium Nash. Panicum trifolium Nash, Bull. Torrey Club 26: 580. 1899. ‘Type collected by Dr. John K. Small, in the Ocmulgee River Swamp, below Macon, Georgia, May 18-24, 1895.’’ The type, in Nash’s herbarium, consists of two tufts of slender vernal culms, 25 to 40 cm. high, with leafy bases and elongated internodes, the rather short-exserted panicles immature. Fig. 284.—Distribution of P. albomarginatum. DESCRIPTION. Vernal form similar to that of P. albomarginatum, but the culms in smaller tufts, taller, 20 to 50 cm. high, more slender, erect; leaves less conspicuously crowded at the base, not so stiff, and proportionately not so much longer than those of the mid- culm; sheaths much shorter than the elongated internodes; blades 3 to 5 em. long, 4 to 5 mm. wide, rather less thick _ and firm than those of P. albomarginatum, the uppermost blade not reduced; panicles usually short-exserted, 3 to 5 cm. long, about as wide, loosely flowered; spikelets as in P. albomarginatum but hardly as wide or as turgid, and the fruit rather less exposed at maturity. Autumnal form erect or leaning, sparingly branching from the middle and upper nodes, the branches usually shorter than the primary internodes. This species is very closely allied to the preceding and some vernal specimens are but doubtfully separated from it. Autumnal specimens may be distinguished by the small fascicles of short branches scattered along the slender primary culm. Fia. 285.—P. trifolium. From type specimen. 262 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Occasional specimens, such as Chase 4112, 4166, and 4304, are brighter green than usual, with less pronounced white margins to the blades and resemble P. flavovirens, but in these the primary culms bear short branches from the middle and upper nodes. DISTRIBUTION. Low, mostly moist sandy woods, North Carolina and Tennessee to Florida and Louisiana. Norra CAROLINA: Scranton, Chase 3199; Roanoke Island, Chase 3225, 3238, 3239, 3248: Chapel Hill, Ashe, Chase 3060; east of Wilmington, Chase 3133, 4576. Sourtn Carouina: Orangeburg, Hitchcock 1387; without local- ity, Ravenel. Grorara: Below Macon, Small in 1895; Warm Springs, Tracy 8864; Augusta, Cuthbert 382, 1159. Fioripa: Baldwin, Hitchcock 997; Milton, Chase 4304; Apalachi- cola, Biltmore Herb. 697a; Lake City, Hitchcock 1023, 1038; Madison, Combs 263; Sanford, Hitchcock 779; Grasmere, Combs 1063; Orange Bend, Chase 4112; Tampa, Hitchcock 938, 940; Braiden- town, Hitchcock 953, 962; Dunedin, Tracy 7029; Myers, Chase 4166, Hitchcock 890, 920, 921; Miami, Chase 3946, Hitchcock 712. TENNESSEE: White Cliff Springs, Scribner in 1890 (Hitchcock Herb.). AtaBaMa: Auburn, Earle & Baker 1535 in part; Cullman, Eggert 24; Flomaton, Hitchcock 1042, 1050, 1053. ; Mississippi: Jackson, Hitchcock 1305; Biloxi, Chase 4358, Hitchcock 1063, 1072, 1088, Tracy 2865, 4612; Mississippi City, Hitchcock 1089, 1100, 1111, Avondale, Tracy 4583, 4603; Saundersville, Tracy 3334; Horn Island, Tracy 4601. LovuistaNa: Calhoun, Hitchcock 1267, 1277; Lake Charles, Hitchcock 1130, 1146. Fig. 286.—Distribution of P. trifoliwm. 155. Panicum flavovirens Nash. Panicum flavovirens Nash, Bull. Torrey Club 26: 572. 1899. ‘Type collected by the writer in Lake Co., Florida, June 16-30, 1895, no. 2061; growing in swampy woods along the edge of road leading to the ford near the J. T. & K. W. R. R. bridge across the Wekiva river.’’ The type, in Nash’s herbarium, is a late vernal form, the primary panicles mostly destitute of spikelets. One of the specimens has a tuft of the long, rather thin, bright green, glossy basal leaves that distinguish this species. The other specimen lacks this prominent tuft of basal leaves and in habit resembles the type of P. albomarginatum Nash, but the blades are not firm and leathery nor white-margined, and the panicles are few-flowered, with flexuous branches. DESCRIPTION. Vernal form bright glossy green; culms densely tufted, very slender, ascending or spreading, 15 to 30 em. high, glabrous, more or less striate-angled, the lower leaves somewhat crowded with overlapping sheaths, the upper distant; sheaths often mi- nutely ciliate on the margin, especially at the summit, otherwise glabrous or the lowermost obscurely pubescent; blades ascending or spreading, 2 to 5 cm. long, 3 to Fig. 287.—P. flavovirens. From type specimen. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 263 4 mm. wide, narrowed toward the rounded base, glabrous, or minutely puberulent beneath, thin, the cartilaginous margin inconspicuous or wanting; panicles open, loosely few-flowered, the flexuous branches spreading or the lower somewhat reflexed ; spikelets 1.3 to 1.4 mm. long, 0.7 mm. wide, elliptic, subacute, pubescent; first glume one-fourth to one-third as long as the spikelet, subacute; second glume hardly equal- ing the fruit and sterile lemma; fruit 1.25 mm. long, 0.6 mm. wide, elliptic. Autumnal form spreading, the slender culms mostly decumbent or prostrate, branching from the lower and middle nodes, these early branches usually as long as the primary culms and loosely branching toward the summit, the short branchlets somewhat fascicled, the flat, reduced blades spreading, the ultimate panicles reduced but exserted; winter rosettes appearing early, usually conspicuous and persisting green during the following season as a dense tuft of sterile shoots with somewhat developed internodes, the blades thin, bright glossy green, as much as 7 cm. long, 3 to 5 mm. wide. This species is allied to P. albomarginatum and P. trifolium, from both of which it is distinguished by the thin bright-green glossy blades, which are scarcely or not at all white-margined. The mode of branch- ing is like that of P. albomarginatum, but looser, the thin blades spreading, the small panicles exserted. DISTRIBUTION. Moist shady or mucky soil, North Carolina to Florida and Mississippi. Norta Carona: Wilmington, Hitchcock 337. SourH Carona: Orangeburg, Hitchcock 25. ; Fioripa: Jacksonville, Combs 34; Lake City, Combs 98 in part; Pensacola, Combs 539; Milton, Chase 4310, 4322; Chipley, Combs 585; Sanford, Hitchcock 7674; Eustis, Chase 4059, Nash 2061; Grasmere, Combs 1088; Tampa, Combs 1394; Lemon Bay, Tracy 7188 in part; Myers, Hitchcock 9014, 905. Mississipi: Biloxi, Tracy 2027. Fig. 288.—Distribution of P. flavovirens. 156. Panicum concinnius nom. noy. * Panicum gracilicaule Nash in Small, Fl. Southeast. U. S. 98. 1903, not Rendle, 1899. On page 1327 in the list of new genera and species, the following citation is given: ‘‘Type, Sand Mt., Jackson Co., Ala., Harbison, no. 2415, 1900, in Herb. N. Y. B. G.”’ This specimen, in the herbarium of the New York Botanical Garden, from the Biltmore Herbarium, is the vernal form, the panicles immature. DESCRIPTION. Vernal form bright green; culms tufted, very slender, erect, gla- Fig. 289.—P.concin- }hrous, 12 to 50 cm. high, nodes minutely puberulent; sheaths, ex- ee ae cept the lower, much shorter than the internodes and less than half Gresiiiconlé Nash, 28 long as the blades, puberulent on the margin, otherwise glabrous; ligules about 0.5mm. long; blades 5 to 7 cm. long, 5 to6 mm. wide, erect or spreading, the margins nearly parallel for most of their length, rounded at base, glabrous or obscurely puberulent beneath, rather strongly nerved, faintly white- margined; panicles finally long-exserted, rather few-flowered, 3 to 6 cm. long, about two-thirds as wide, the branches ascending; spikelets 1.1 mm. long, 0.7 mm. wide, obovate, obtuse, pubescent; first glume about one-fifth the length of the spikelet; sec- 264 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. ond glume and sterile lemma slightly shorter than the fruit at maturity; fruit 1 mm. long, elliptic, acute. Autumnal form radiate-spreading, late in the season bearing a few branches with somewhat reduced blades and small exserted panicles; winter rosette appearing early, the numerous, rather firm blades bluish green, about the size of those of the vernal culms. DISTRIBUTION. Moist sandy ground, northern Georgia and Alabama; apparently rare. GeEorGIA: Thomson, Bartlett 1461. AuaBAmMa: ‘‘Sandy soil along a creek, Sand Mt., June 5, 1900,’’ Harbison 2415; ‘‘Bank along roadside above Bryants Creek, Fic. 290.—Distribution of P. concinnius. south of Pisgah, Oct. 14, 1907,”’ Chase 4475; ‘‘In moist spot in woods, south of Pisgah, Oct. 14, 1907,’’ Chase 4476; ‘‘Culms widely spreading, crevices of mossy rocks, north bank of Bryanis Creek, south of Pisgah, Oct. 14, 1907,’’ Chase 4483. The last-mentioned specimen, Chase 4483, was collected at the type locality of the species, as indicated by Mr. Harbison in a letter. 157. Panicum ensifolium Baldw. Panicum ensifolium Baldw.; Ell. Bot. 8. C. & Ga. 1: 126. 1816. ‘‘Grows in damp soils, * * * Georgia. Dr. Baldwin.’’ The type, in the Elliott Her- barium, isa slender plant 33 cm. high, with a tuft of four acuminate basal leaves, the blades 2.5 to 3.5 cm. long, four culm leaves, the upper minutely puberulent through- out on the under surface, the lower toward the tip only, and a long-exserted panicle, with puberulent spikelets 1.5 mm. long. The accompanying label reads: ‘‘ Panicum ensifolium Bald. Hab: in humidis Georg: Dr. Baldwin.’’ The basal blades of the type specimen and of a second specimen from ‘‘Baldw. Georg.’’ in the herbarium of the Philadelphia Academy are firm and sharp-pointed, though to a much less degree than in P. tenue and P. albomarginatum. Panicum nitidum ensifolium Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 29. 1889. Based on Panicum ensifolium Baldw., though the description applies to P. vernale. Panicum brittoni[i] Nash, Bull. Torrey Club 24: 194. 1897. ‘‘In moist sand in the ‘pine barrens’ at Forked River, N. J. Collected by Dr. Britton during an excur- sion of the Torrey Botanical Club to the region May 29-June 2, 1896.’ The type, in Nash’s herbarium, consists of a tuft of slender, simple, vernal culms 10 to 19 em. high, the blades glabrous or minutely puberulent on the under surface, the minutely pubescent spikelets 1.3 to 1.4 mm. long. Panicum cuthbertii Ashe, Journ. Elisha Mitchell Soc. 15:48. 1898. ‘‘South Caro- lina: Cuthbert; St. Helena Island.’’ This specimen could not be found in Ashe’s her- barium, but a piece of the type bearing the above data, sent by Mr. Ashe, is in the National Herbarium. It consists of a single vernal culm lacking the base, with two nodes, the blades broken off, but the sheaths present, the panicle short-exserted, the immature, pubescent spikelets 1.4 mm. long. Ashe states that ‘‘it is separated from P. ensifolium by the strict habit and large basal leaves of the latter,’ but P. ensifolium as understood by Ashe is P. albomarginatum, as shown by his description @ and by his giving P. albomarginatum Nash as a synonym of P. ensifolium. @ Journ. Elisha Mitchell Soc. 15: 46. 1898. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 265 Panicum glabrissimum Ashe, Journ. Elisha Mitchell Soc. 15: 62. 1898. ‘‘The type material was collected by me June, 1898, at Manteo, Dare Co., N. C.’’ The type could not be found in Ashe’s herbarium. In the Mohr Herbarium is a specimen labeled in Ashe’s writing ‘‘ Panicum glabrissimum Ashe” and bearing the cited data. This is a tuft of three vernal culms and agrees with the description, except that the spikelets are said to be glabrous, while these are pubescent. The specimen in the National Herbarium from the same station and sent by Ashe as part of the type collection is P. tenue, and fails in several particulars to agree with the description. While neither of these specimens is the type itself, the one which most nearly agrees with the description is taken to represent the type. Panicum shallotte Ashe, Journ. Elisha Mitchell Soc. 16: 84. 1900. Based on ‘‘P. glabrissimum Ashe, not P. glaberrimum Steud.”’ Panicum parvipaniculatum Ashe, Journ. Elisha Mitchell Soc. 16: 87. 1900. ‘‘Col- lected May 20, in Onslow county, N. C. Type material is preserved in my herba- rium.’’ No specimen so labeled could be found in Ashe’s herbarium, but a cover marked in Ashe’s hand ‘‘P. parvipaniculatum” was found which contained eight sheets of unmounted material, of which two sheets (one within the fold of the other) were accompanied by a label with the following data in Ashe’s writing: ‘‘Panicum gray spikelets? Peaty-soiled thickets sandy flatwoods and ditch banks, 10-18 miles east of Jacksonville, Onslow county, N. C. May 20, 1899.’’ Since these were the only specimens with locality and date according with those published, the specimens on the sheet with the label were chosen as the type, one tuft being deposited in the National Herbarium. These specimens agree with the description except that the species is said to be ‘‘perfectly glabrous except the ligule” while the blades are puberulent beneath and some of them on the upper surface also; that the ligule is given as ‘‘about 2 mm. long,’’ while it is almost obsolete (0.1 to 0.2 mm. long); and that the spikelets are given as ‘‘barely 1 mm. long,’’ while they measure 1.3 to 1.4 mm. long. This type differs from those of P. ensifolium and P. brittonii in having glabrous spikelets. DESCRIPTION. Vernal plants grayish olive green; culms cespitose, slender, erect or reclining, glabrous, 20 to 40 cm. high; sheaths glabrous, usually much shorter than the inter- nodes; blades distant, often reflexed, 1 to 3 cm. long, 1.5 to 3 mm. wide, glabrous on the upper surface or puberulent toward the base, puberulent beneath, at least toward the tip; panicles finally long-exserted, 1.5 to 4 cm. long, nearly as wide, the flexuous branches spreading or the lower reflexed; spikelets 1.3 to 1.5 mm. long, elliptic, subacute, glabrous or puberulent; first glume one- fourth as long as the spikelet or less, acute or obtuse; second glume slightly shorter than the fruit and sterile lemma; fruit 1.2 mm. long, elliptic, subacute. Autumnal culms spreading or reclining, sparingly branch- ing from the middle nodes, the branches mostly remaining simple; winter blades glabrous, usually short, 1.5 to 3 cm. long, 2 to 4 em. wide. In this species the pubescence of the spikelets seems to be very inconstant. The type of P. brittonii and Chase 3557 have pubescent spike- lets, while Chase 3535 and specimens collected by Clute in 1899 and by Bicknell in 1900, also in New Jersey, have glabrous spikelets; of the North Carolina specimens Hitchcock 1425, Chase 30964, 3176, 3177, 3227, and 3234 have pubescent spikelets. Combs’s no. 74, Lake City, Florida, and Tracy 44, Ocean Springs, Mississippi, two very slender autumnal specimens, the first with puberulent, the latter with glabrous, spikelets, are doubtfully referred here. Fic. 2301.—P. ensifolium. From type specimen. 266 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Wet places, mostly sphagnum bogs or swamps, New Jersey to Georgia; also in Mississippi. New Jersey: Forked River, Britton in 1896; Penn Place, Clute in 1899; Toms River, Bicknell in 1900; At- sion, Chase 3535, 3557. Maryann: Beltsville, Chase 3739. NortH Carortna: Roanoke Is- land, Chase 3227, 3234; West Raleigh, Stanton, 1272; Wil- sons Mills, Chase 30964, 3097; Onslow County, Ashe in 1899, Chase 3176, 3177, 3196; Wil- mington, Hitchcock 1425, 14363, 1439. Fia. 292.—Distribution of P. ensifolium. - SoutH Carourna: St. Helena Is- land, Cuthbert in 1887; Orangeburg, Hitchcock 1370, 1379, 1405. GeorGIA: Bulloch County, Harper 829; Augusta, Cuthbert 1160; without locality, Baldwin. Mississippi: Biloxi, Hitchcock 1067. 158. Panicum vernale sp. nov. DESCRIPTION. Vernal plants light green, soft in texture; culms densely cespitose, 15 to 30 cm., rarely to 40 cm. high, very slender, ascending or spreading, glabrous, the nodes glabrous; leaves clustered at the base, the thin, rather soft blades 2 to 7 cm. long, 3 to 5 mm. wide, those of the culm remote, the glabrous sheaths one-fourth to one-third as long as the elongated internodes; ligules almost obsolete; blades 0.7 to 2.5 cm. long, 2 to 3 mm. wide, glabrous or puberulent on the lower surface, occasionally also on the upper surface, at first erect, becoming spreading or reflexed; panicles finally long-exserted, 1.5 to 3 cm. long, nearly as wide, rather few-flowered, the flexuous branches spreading; spikelets 1.4 to 1.5 mm. long, 0.8 mm. wide, obovate-elliptic, subacute, pubescent; first glume about one- fourth as long as the spikelet, subacute; second glume and sterile lemma scarcely as long as the fruit at maturity; fruit 1.2 mm. long, 0.7 to 0.8 mm. wide. ga ented Let Autumnal form like the vernal form in appearance, branching From type speci- , men. from the base, these culms simple and soon dying to the ground, rarely late in the season producing a few short fascicled branchlets at the nodes, the scarcely reduced flat blades spreading; winter leaves numerous, soft, persistent during the vernal stage, linear, rather abruptly narrowed at the apex, not long-acuminate. Type U.S. National Herbarium no. 558416, collected in a ‘‘sphagnum bog, Lake City, Florida, April 16, 1906,’’ by A. 8. Hitchcock (no. 1020). This species has been confused with P. ensifolium Baldw.,¢ from which it is dis- tinguished by the more densely cespitose habit and light green, soft foliage, the very numerous basal blades as much as 7 cm. long, flat, linear, not long-acuminate. « Panicum nitidum ensifolium as described in Chapman’s Flora (Fl. South. U. 8. ed. 3. 586. 1897) is P. vernale. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 267 All the specimens cited below were collected in the spring. Since localities, like Lake City, Eustis, and Miami, Florida, where this species was found in March and April, were visited in September without its being found, it would seem that the plants usually die to the ground in early summer and that the secondary branches appear only rarely. In Hitchcock 931 and 9584 a few sparingly branched dead culms are attached, being the only branching culms seen. Two collections, Hitchcock 809 and Nash 424, have blades pubescent on the upper surface, while Hitchcock 941 and 1092 have some blades that are pubescent and some that are glabrous on the up- persurface. Hitchcock’s nos. 1066 and 1092 have glabrous spikelets. DISTRIBUTION. Moist places, especially sphagnum bogs, Florida to Mississippi. Fioriwa: Baldwin, Hitchcock 1004; Lake City, Bitting 19, Hitch- cock 1020; Apalachicola, Chap- Fic. 294.—Distribution of P. vernale. man; Eustis, Nash 273 in part, 424, Hitchcock 795, 798, 809; Dunedin, Treacy 6699; Braidentown, Hitchcock 9584, 959, 960; Johns Pass Tracy 7180; Tampa, Hitchcock 936, 941; Miami, Hitchcock 931, 942. ALABAMA: Flomaton, Hitchcock 1041. Mississirr1: Biloxi, Hitchcock 1066; Mississippi City, Hitchcock 1092. 159. Panicum curtifolium Nash. Panieum curtifolium Nash, Bull. Torrey Club 26: 569. 1899. ‘‘Collected by S. M. Tracy at Ocean Springs, Mississippi, May 2, 1898, no. 4598.’’ The type, in Nash’s herbarium, consists of a tuft with two slender vernal culms about 30 cm. long, begin- ning to branch at the middle nodes. The blades are glabrous above except at the base and glabrous or sparsely pubescent beneath. Ina duplicate type in the National Herbarium several blades have a few scattered hairs on the upper surface. Panicum earlei Nash, Bull. Torrey Club 26: 571.1899. ‘‘Type collected at Auburn, Lee Co., Alabama, on May 7, 1898, by Messrs. F'. 8. Earle and C. F. Baker, no. 1532.”’ The type, in Nash’s herbarium, consists of a tuft of early vernal culms 8 to 15 cm. high, with immature panicles. The blades are sparsely pilose on the upper surface. Panicum austro-montanum Ashe, Journ. Elisha Mitchell Soc. 16: 85.1900. ‘‘Along mountain streams of Northern Alabama and the adjacent parts of Tennessee. Type material is preserved in my herbarium.’’ The type specimen could not be found in Ashe’s herbarium. In the National Herbarium is a specimen from Sand Moun- tain, Alabama, June, 1899, sent by the Biltmore Herbarium, which was compared by E. D. Merrill in 1900 and said by him to be identical with the type of P. austro- montanum. It is also the same as a specimen from western North Carolina sent by Ashe as representing P. austro-montanum, and furthermore agrees with the original description except that the spikelets are 1 mm. long, instead of 0.7 mm. long. The Biltmore specimen agrees with the types of P. curtifolium and P. earlei. DESCRIPTION. Vernal form in dense colonies, the culms not crowded in the clump; culms 10 to 30 cm. high, slender, weak, angled, erect or spreading, glabrous or sometimes with a few scattered hairs, the nodes sparsely bearded; sheaths much shorter than the elongated internodes, striate-angled, sparsely spreading-pilose, ciliate, especially at 268 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. the summit; ligules about 1 mm. long, the hairs soft, rather sparse; blades spread- ing or reflexed, 1.5 to 3 cm. long, 2 to 5 mm. wide, thin and soft, sparsely pilose on both surfaces or glabrous above except for long soft hairs near the base; panicles short-exserted at least till after maturity, 2 to 3 cm. long, nearly as wide, the branches ascending; spikelets 1.4 mm. long, 0.7 mm. wide, elliptic-obovate, obtuse, glabrous, or minutely pubescent; first glume about one-fifth as long as the spikelet; second glume and sterile lemma both shorter than the fruit at maturity; fruit 1.25 mm. long, 0.7 mm. aS wide, elliptic. Be Acinic ns hay Autumnal form weakly spreading, the culms branching from the middle nodes after the maturity of the primary panicles, the branches exceeding the internodes; ultimate branchlets in small fasci- cles toward the summit of the branches, the reduced blades spreading and the small panicles mostly exserted ; winter rosette appearing early, the soft blades mostly 2 to 3 cm., but sometimes as much as 5 cm. long. This species is the only one of this group with spreading pilose pubescence and manifest ligules. DISTRIBUTION. Boggy soil and shady, moist places, sometimes forming a rather dense car- pet, South Carolina and Tennessee to Florida and Mississippi. Fic. 296.—Distribution of P. curtifolium. Sours Carouna: Aiken, Ravenel; Hartsville, Coker in 1909. Fioriwa: Pensacola, Combs 525; Eustis, Holm 24, Nash 1507; Myers, Hitchcock 867; without locality, Chapman. TENNESSEE: Lookout Mountain, Biltmore Herb. 10715b (Biltmore Herb.). ALABAMA: Sand Mountain, Biltmore Herb. in 1899; Auburn, Harle & Baker 1532, Hitchcock 1337, Tracy 3748 in part, 3752; Flomaton, Hitchcock 1058; Tuskegee, Carver 27, 41; Gateswood, Tracy 8421. MississipPi: Ocean Springs, Tracy 4598, 4599; Mississippi City, Hitchcock 1094, 160. Panicum chamaelonche Trin. Panicum chamaelonche Trin. Gram. Pan. 242. 1826. Trinius states concerning the origin of his specimen, ‘‘V. spp. Am. bor. (TRaTrTrinick, ex. coll. Enslini).’’ The type, in the Trinius Herbarium, is the early branching form. Panicum nitidum minor[us] Vasey, Contr. Nat. Herb. 3: 30.1892. “Florida.’”’ The type, in the National Herbarium, was collected by William C. Canby at St. Augustine, Florida, April, 1869, and is labeled ‘“‘nitidum var. minor” in Dr. Vasey’s writing. This consists of several small tufts of vernal culms. Panicum baldwinit Nutt.; Kearney, U. 8. Dept. Agr. Div. Agrost. Bull. 1: 21. 1895. The citation is as follows: ‘‘Panicum baldwinii Nutt. in Herb. Phila. Acad. (Panicum nitidum minor Vasey Contr. U.S. Nat. Herb. 3: No. 1, 30, 1892.)’” No description is given. Since Nuttall’s herbarium name is taken up, this name should be considered a nomen nudum, and not based on the synonym cited. Panicum baldwinit Nutt.; Chapm. Fl. South. U. 8. ed. 3. 586. 1897. Based on “Panicum baldwinii Nutt. (in Herb.)’’ and described. The type, in the Nuttall Herbarium, labeled ‘‘Panicum Baldwinii, Florida, Bald.’’ is a tuft of vernal culms beginning to branch, with mature and over-mature panicles, “A HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 269 The species described as P. ramulosum Michx. by Chapman is P. chamaelonche as shown by the description, by a specimen in the Chapman Herbarium at the New York Botanical Garden labeled ‘‘ Panicum ramulosum Michx. (nitidum S. Fl.!)> Southern Florida,’”’ and by the fact that Chapman¢ cites “P. ramuloswm Flora” [Southern U. S.] as a synonym under P. baldwinii Nutt. Scribner @ describes and illustrates this species as P. baldwinii ‘‘ Nutt. in herb.”’ and gives as synonym, “‘P. dichotomum var. nitidum Chapman, Southern Flora, first edition.’’ Chapman does not make the combination as stated by Scribner, but refers P. nitidum Ell. to P. dichotomum as a form. Chapman’s plant as stated above is P. chamaelonche, but P. nitidum of Elliott’s herbarium is P. longiligulatum Nash. DESCRIPTION. Vernal form densely tufted; culms 10 to 20 or even 30 cm. high, ascending, glabrous, the nodes glabrous; sheaths, except the basal ones, half as long as the internodes or less, at least the upper rather loose, glabrous or occasionally with a few ciliz on the margin; ligules 0.2 mm. long; blades firm, ascending or spreading, 1.5 to 4 cm., rarely 5 em. long, 2 to 3 mm. wide, more or less involute-pointed, glabrous on both surfaces, often with a few long, stiff hairs on the margin near the base; panicles finally long- exserted, 2.5 to 5 cm. long, nearly as wide, the flexuous branchlets and pedicels spread- ing at nearly right angles; spikelets 1.1 to 1.2 mm. long, 0.6 mm. wide, obovate, obtuse, turgid, glabrous; first glume one-fourth to one-third as long as the spikelet, obtuse; second glume slightly shorter than the fruit and sterile lemma; fruit 0.9 to 1 mm. long, 0.6 mm. wide, elliptic, subobtuse. Autumnal form freely branching from the base and lower nodes, the early branches often as long as the primary culms, Fic. 297.—P. chamaclonche. Tepeatedly branching, forming dense cushions, as much as 50 From type specimen. cm. across, the longer culms upturned at the ends; ultimate branchlets more or less fascicled, the scarcely reduced blades drying involute, overtopping the small panicles; winter rosettes usually persisting green during the vernal state, the rather firm blades 2 to 5 cm. long. The plants are usually purple throughout in both vernal and autumnal state. Occasional specimens, such as Hitchcock 873, are yellow green, and look strikingly different in the field. Hitchcock’s no. 1436, Chase 4570, and Tracy 6732 have unusually large blades, as much as 6 em. long and 5mm. wide. The speci- mens of Hitchcock 952 are 30 to 40 cm. high, with large panicles and rather long lower blades and appear to be intermediate between this and P. glabrifolium. DISTRIBUTION. FIG. 298.—Distribution of P. chamaelonche. Open sandy soil, mostly in the low _ pine land or “‘flatwoods,’’ North Carolina to Florida and Mississippi. Norts Carouna: Vicinity of Wilmington, Chase 3125, 4570, Hitchcock 338, 339, 1427, 1435, 1436, 1483, 1488; Wards Mill, Chase 3182. Georeta: Savannah, Kearney 177. @ Fl. South. U.S. ed. 2. 667. 1889. > This is included as a form under P. dichotomum L. in Chapm. F1. South. U. 8S. 576. 1860. ¢ Fl. South. U. S. ed. 3. 586. 1897. 4U.S. Dept. Agr. Div. Agrost. Bull. 11: 43. /. 3. 1898. 270 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Fiorina: Jacksonville, Curtiss 21, 3602*, 5588, Kearney 145; Baldwin, Hitch- cock 985; Lake City, Combs 127, Hitchcock 1027, 1037; Carabelle, Kearney 92; Dunedin, Tracy 6726; St. Augustine, Canby in 1869; Indian River, Palmer 634 in 1874 in part; Melbourne, Curtiss 5804; Titusville, Chase 3965, Hitchcock 759; Jensen, Hitchcock 729; Sanford, Chase 4032, 4036, 4037, Hitch- cock 769, 778, 781, 828; Eustis, Chase 4053, Hitchcock 794, 806, 807, 818, Nash 51, 71, 335, 778, 1238; Oakland, Curtiss 6628; Lemon Bay, Tracy 7191, 7200; Tampa, Combs 1344, Hitchcock 928, 934, 944; Braidentown, Hitch- cock 951, 952, 954, 973, Tracy 6732; Manatee, Hitchcock 975, Rugel 377; Myers, Chase 4153, 4156, 4171, Hitchcock 865, 866, 873, 885, 887, 895, 916, 924, Lee Co. Pl. 472; Sneeds Island, Tracy 6451, 6464, 6693; Perdido, Tracy 8407. ALABAMA: Fort Morgan, Tracy 7207. Mississippr: Avondale, Tracy 4610; Mississippi City, Hitchcock 1114. In the herbarium of the Philadelphia Academy is a specimen said to be from Surinam which appears to be P. chamaelonche. 161. Panicum glabrifolium Nash. Panicum glabrifolium Nash, Bull. Torrey Club 24: 196. 1897. ‘‘Collected by the writer in the ‘flatwoods’ at Tampa, Florida, on August 20, 1895, no. 2415a.”’ The type, in Nash’s herbarium, is the early branching state. The spikelets are 1.25 mm. long. The statement in the original description, ‘‘the spike- lets slightly exceeding .5 mm. in length” is doubtless a typo- graphical error. DESCRIPTION. Vernal form similar to that of P. chamaelonche, in smaller Fie. 290° P. Glabevonian: ~ tufts; culms stouter, more or less flattened, 15 to 50 cm. high, From type specimen. erect or sometimes subgeniculate at base; blades firm, erect, or narrowly ascending 4 to 12 cm. long, or the lower occasion- ally as muchas 20cm. long, 2 to 4mm. wide, usually involute at least toward the apex, glabrous; panicles 4 to 9 cm. long, two-thirds to three-fourths as wide, the branches ascending, the ultimate branchlets and spikelets more or less secund along the lower side of the branches; spikelets 1.2 to 1.4mm. long, obovate, obtuse, turgid, glabrous; first glume about one-third as long as the spikelet; second glume shorter than the fruit and sterile lemma; fruit 1.1 to 1.2 mm. long, elliptic. Autumnal culms wiry, elongated, and spreading, with geniculate nodes and long internodes; freely branching from the middle and upper nodes, the blades long and narrow, overtopping the somewhat reduced panicles; winter leaves less numerous than in P. chamaelonche, as much as 10 to 15 cm. long, stiffly ascending. In the vernal form this species is distinguished from P. chamaelonche by the taller, stouter culms, more involute blades, larger panicles, and slightly larger spikelets. The autumnal form is distinguished by the different branching habit. There are, however, intermediate specimens that appear to connect the twospecies, such as Hitchcock 893, from Myers, which has the tall culms, elongated lower blades and large panicles of P. glabrifolium but the smaller spikelets of P. chamaelonche. DISTRIBUTION. Lowsandy woods, peninsular Florida. Fiorimwa: Crystal, Combs 1024; Braidentown, Combs 1313, 1316, Hitchcock 966, Tracy 6715; Manatee, Hitchcock 978, Simpson in 1889; Cedar Key, Combs 780; Bartow, Combs 1187. Fig. 300.—Distribution of P. glabrifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 271 162. Panicum breve sp. nov. DESCRIPTION. Vernal form purplish, culms in dense tufts, 5 to 17 cm. high, erect, stiff and wiry, glabrous or appressed-pubescent below; sheaths crowded at the base as in species of Festuca, those of the culms usually longer than the internodes, ciliate on the margin, otherwise glabrous; ligules dense, about 0.3 mm. long; blades firm, erect or ascending, 3 to 6 cm., sometimes 8 cm. long, about 1.5 mm. wide when flattened out, strongly involute, more or less falcate, sometimes strongly so, a few long, stiff hairs on the margin toward the base, otherwise glabrous; panicle short-exserted, 1.5 to 4 cm. long, nearly as wide, loosely flow- Fic.301.—P. breve. From red, the flexuous branches spreading; spikelets 1.3 to 1.4mm. type specimen. long, obovate, obtuse, turgid, puberulent; first glume one-third to half as long as the spikelet; second glume and sterile lemma barely equaling the fruit at maturity; fruit 1.2 mm. long, elliptic. Autumnal form erect, branching from the middle nodes, the fascicled branches strict, the reduced wiry blades overtopping the panicles. Type U.S. National Herbarium no. 558435, collected April 5, 1906, in “‘low pine woods between scrub hills, among palmetto,’’ Jensen, Florida, by A. 8. Hitchcock (no. 734). This species is most closely related to P. glabrifolium, from which it is distinguished by its short, compact habit, by the strongly involute blades bearing long, stiff hairs near the base, and by the pubescent spikelets. DISTRIBUTION. Low pine woods and hammocks, east coast of southern Florida. Fiorina: Indian River, Palmer 634 in 1874 in part; Jensen, Hitchcock 734; Fort Lauder- dale, Small & Carter in 1903; ‘‘About Boca Ratone Lake, below Delray,’’ Small & Carter in 1903 (Biltmore Herb.). Fig. 302.—Distribution of P. breve. Lancearia.—Plants olive green, often purplish; vernal culms usually wiry, minutely crisp-puberulent or glabrous; sheaths glabrous or puberulent, at least at the summit; ligules nearly obsolete; blades glabrous or puberulent, usually strongly ciliate, at least near the base; spikelets unsymmetrically pyriform, that is, more swollen on the face than on the back; first glume thin and _ shining, broad at the summit, obtuse or truncate; second glume and sterile lemma strongly 7 to 9-nerved, puberulent or glabrous. Species of the At- lantic Coastal Plain. UE a ei 163. P. pauciciliatum. Spikelets 2 mm. or more long. Blades, or some of them, at least 8 mm. wide; glabrous on the upper surface; fruit papillose-roughened....... 166. P. webberianum. ' Blades not over 6 mm. wide (or if wider, puberulent on : the upper surface); fruit smooth and shining. Spikelets 2.4 to 2.6 mm. long; blades narrowed to- oo CE a a 5 167. P. patentifolium. 272 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Spikelets not over 2.1 mm. long. Blades firm, glabrous above; culms stiffly ASCCTICINE. 226 oS Se ee eee eee eee eee 164. P. lancearium. Blades lax, softly puberulent on both surfaces; Gilmmcecun bent... .c..25bs aeacme ee tee 165. P. patulum.— 163. Panicum pauciciliatum Ashe. Panicum pauciciliatum Ashe, Elisha Mitchell Soc. 16: 87.1900. ‘‘Collected by me May 20, 1899, growing in dry sand near Wilmington, N.C.” The type, in Ashe’s herbarium, consists of six single culms, beginning to branch, 25 to 30 em. high, with somewhat geniculate nodes, and short-exserted, hardly mature panicles. DESCRIPTION. Vernal culms cespitose, erect or geniculate at base, slender, stiff and wiry, 15 to 30 cm. high, the internodes commonly reddish purple, crisp-puberulent to nearly gla- brous; sheaths much shorter than the internodes, striate, glabrous or crisp-puberulent, usually ciliate; blades firm, 2 to 5 cm. long, 3 to 6 mm. wide, ascending or spreading, glabrous to puberulent, ciliate near the base; panicles 2 to4 cm., rarely 6 or 7 cm., long, two-thirds as wide, the flexuous branches spreading or the lower reflexed, the pedicels and ultimate branchlets often directed toward the under side; spikelets 1.5 to 1.6 mm. long, 1 mm. wide; first glume one-third to half as long as the spikelet, obtuse or truncate; second glume and sterile lemma puberulent, the glume shorter than the fruit and sterile lemma; fruit 1.4mm. long, 1 mm. wide, elliptic-obovoid, obscurely pointed. ‘ Autumnal culms ascending from a decumbent base, branching from all but the uppermost node before the maturity of the primary panicles, the primary inter- nodes often elongating, the terminal joint with its panicle together with the internode below it often falling early, thus giving the ap- pearance of short culms branching at all the nodes characteristic of this species; early branches about equaling these shortened primary culms, repeatedly branching, the ultimate branchlets in fascicles toward the ends, the reduced blades spreading, invo- lute-pointed; winter rosette appearing late, not conspicuous. This species often closely resembles P.lancearium, but the differences, though small, are fairly constant, though Chase 3126 and Ennis in 1899 have spikelets 1.7 to 1.8 mm. long. Chase 3139, Wilmington, N. C., with ligules 0.3 mm. long and scarcely pyriform spikelets, is doubtfully re- ferred here. Fig. 303.—P. pauciciliatum. From type specimen. DISTRIBUTION. Sandy woods of the Coastal Plain, mostly in moist places, North Carolina to Florida and along the Gulf to Texas; also in Cuba and Porto Rico. Norts Carona: Roanoke Island, Chase 3246; Wilmington, Ashe Fig. 304.—Distribution of P. pauciciliatum. in 1899, Chase 3126, 3127, 3128, 3162, 4567, Hitchcock 414, 416, 1432, 1477, 1479, 1487. Fiorwa: Baldwin, Hitchcock 992; Apalachicola, Chapman; Orange County, Baker 41, 70, 71, 72, Combs 1085, Meislahn 169; Eustis, Chase 4045, Hitchcock 793, HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 273 797, 803, 804, 808, 819, Nash 15, 63 in part, 1337, 2076; Sumter County, Curtiss F, 3600A in part; Jensen, Hitchcock 733, 737, 750; Santa Rosa Island, Tracy 6446, 6447; Perdido, Tracy 8406; Myers, Chase 4173, Hitchcock 889. ALABAMA: Fort Morgan, Tracy 8397. Mississreri: Biloxi, Kearney 3314; Mississippi City, Hitchcock 1113; Horn Island, Tracy 2863, 8412. Texas: Narcoossee, Ennis in 1899. Cusa: Without locality, Wright 3876. Porto Rico: Santurce, Heller 982b, 6442; Vega Baja, Heller 639, Underwood & Griggs 955. 164. Panicum lancearium Trin. Panicum lancearium Trin. Gram. Pan. 223. 1826. Trinius here gives a full descrip- tion and states that his specimen was collected in North America by Enslin and com- municated by Trattinick: ‘‘V. spp. Am. bor. (Trattinick ex hbio Enslini).” Tri- nius had previously mentioned the name@ as a probable synonym of a Plukenet species. The type, in the Trinius Herbarium, is the vernal form, with glabrous spike- lets 2 mm. long. It is labeled ‘‘Plukn. Tb. 92. f.6.? In Am. bor. ab Enslino 1. dt. cl. Trattinick.” Panicum nashianum Scribn. U. 8. Dept. Agr. Div. Agrost. Bull. 7: 79. f. 61. 1897. Two specimens are cited, ‘‘4029 Curtiss (1893), and 466 Nash (1894).—Low pine bar- rens, often in moist ground, near the coast, Virginia to Mississippi.” The type (Nash 466, since the species is named for the collector) is in the National Herbarium. It consists of a clump of numerous culms 15 to 30 cm. high with mature and immature panicles, the spikelets minutely pubescent. The accompanying label gives the fol- lowing data: ‘‘Dry sandy soil. Grows in dense clumps 1 ft. across. Collected in vicin- ity of Eustis, Lake county, Florida, by Geo. V. Nash, April 15-30, 1894.” The Curtiss specimen cited by Scribner has glabrous spikelets. DESCRIPTION. Vernal culms cespitose, usually purplish, wiry, stiffly ascending from a more or less geniculate base, 20 to 50 cm. high, minutely grayish crisp-puberulent; sheaths puber- ulent, at least near the margin, much shorter than the internodes; blades ascending or spreading, firm, 2 to 6 cm. long, 3 to7 mm. wide, puberulent or nearly glabrous beneath, usually glabrous on the upper surface, strongly ciliate toward the base, or sometimes nearly to the apex; panicles 3 to 6 cm. long, two-thirds as wide, rather few-flowered, the flexuous branches spreading, or the lower reflexed; spikelets 2 to 2.1 mm. long, 1 to 1.2 mm. wide; first glume one-third to half as long as the spikelet, obtuse or truncate; second glume and sterile lemma puberu- lent or sometimes glabrous, the glume slightly shorter than the fruit and sterile lemma; fruit 1.6 to 1.7 mm. long, 1 mm. wide, obovate-elliptic, minutely puberu- lent at the apex. Autumnal culms geniculate-spreading, ascending at the ends, the stiff internodes occasionally elon- gated, branching from the middle nodes, the branches much longer than the inter- nodes, late in the season bearing fascicles of short branchlets toward the summit, the reduced flat or involute-pointed blades spreading, the ultimate panicles reduced to a few spikelets, partly inclosed in the sheaths. Fig. 305.—P. lancearium. From type specimen. @ Clav. Agrost. 234. 1822. 41616°—vo. 15—10——18 274 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Occasional unusually large specimens, such as Curtiss 6626, Hitchcock 678, and Tracy 7051, resemble P. webberianum, but may be distinguished by the smaller spikelets, with smooth and shining fruits. The following specimens approach P. patulum in habit or in having papillose, more rounded spikelets, but the blades are not pubescent on the upper surface, or but one or two are pubescent, the others being glabrous: Chase 3211, 4543, 4568, 4569, Har per 1689, Hitchcock 1016, Wright 3460. DISTRIBUTION. Low sandy woods of the Coastal Plain, from southeastern Virginia to Florida and Mississippi; also in Cuba. Virainta: Cape Henry, Chase 2345; Norfolk, Vasey in 1884; Dismal Swamp, Chase 3656. Norra Carona: Roanoke Island, Chase 3211, 3224; Wards Mill, Chase 3181; Jacksonville, Ashe in 1899, Chase 3193, 3194; vicinity of Wilmington, Chase 3113, 3129, 4568, 4569, Hitchcock 1431, 1466, 1486. Sourn Carona: Isle of Palms, Ball 792, Chase 4536, 4542, 4543, 4545. Grorata: Savannah, Kearney 178; Tifton, Harper 1689. Fiorma: Duval County, Curtiss 3600 A in part; Jacksonville, Curtiss 4029 (Hitchcock Herb.); Baldwin, Hitchcock 994, 1003; Lake City, Combs 75, 104; Hitchcock 1016, 1025, 1028, 1033; Titusville, Chase 3970, 4004, 4015, Hitchcock 764, 766; Sanford, Hitchcock 770, 784, 821, 824, 825; Eustis, Nash 301, 466; Pensacola, Tracy 8425; Clearwater, J'’racy 7051 in part; Waldo Combs 686; Indian River, Palmer 631 in 1874; Gainesville, Chase 4239, Combs 744, 752; Lakeland, Hitchcock 844, 849; Tampa, Hitchcock 929, 932, 937; Oakland, Curtiss 6626; Madison, Combs 238; Crystal, Combs 1025; Bartow, Combs 1240; Mana- tee, Hitchcock 956, 977, Rugel 376; Santa Rosa Island, Tracy 6466; Pine Island, Tracy 7205; Sneeds Island, Tracy 6452; Perico Island, Tracy 6730; Palma Sola, Tracy 6714; Perdido, Tracy 8409; Sarasota, Tracy 7203; Semi- nole, Tracy 7194, 7195; Mary Esther, Tracy 7175, 9143; Myers, Chase 4149, 4175, Hitchcock 864, 869, 888, 911, 913; Miami, Hitchcock 664, 678, 713; with- out locality, Rugel 291. AtaBAMA: Flomaton, Hitchcock 10394; Fort Morgan, Tracy 7209. Missrssrpp1: Avondale, Tracy 4581 in part; Biloxi, Kearney 330 in part, Tracy 2869, 6465; Mississippi City, Hitchcock 1093, 1298; Ocean Springs, Skehan in 1895. Cusa: San Juan de Buenavista, Wright 3460 (Gray Herb.). Fic. 306.—Distribution of P. lanceariwm. 165. Panicum patulum (Scribn. & Merr.) Hitche. Panicum nashianum patulum Scribn. & Merr. U. S. Dept. Agr. Div. Agrost. Cire. 27:9. 1900. “Type specimen: 1296 Robert Combs, Braidentown, Manatee County, Fla., September 3, 1898.’’ The type, in the National Herbarium, consists of a clump of numerous prostrate culms, 18 to 32 cm. long, with mature primary panicles and numerous branches witli secondary panicles. Panicum patulum Hitche. Rhodora 8: 209. 1906, Based on P. nashianum patulum Scribn. & Merr. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 275 DESCRIPTION. Vernal form densely cespitose, grayish olive green; culms geniculate-decumbent, ascending at the ends, as much as 50 cm. long, the internodes and sheaths densely vel- vety puberulent, the latter usually ciliate on the margin, at least toward the summit; blades rather lax, spreading, 4 to 8 cm. long, 4 to 8 mm. wide, tapering toward both ends, velvety-puberulent beneath, softly pubescent on the upper surface, sometimes obscurely so, ciliate at least half their length; panicles as in P. lancearium, the spikelets more globular-pyriform than in that species, 1.3 mm. wide and nearly as thick, second glume and sterile lemma densely papillose-pubescent; fruit 1.8 mm. long, 1.1 to 1.2 mm. wide; otherwise as in P. lancearium. Autumnal form more freely branching than in P. lancearium, often forming large mats, the de- cumbent culms producing ascending branches from all the nodes at about the maturity of the primary panicles, these repeatedly branching, the ultimate branchlets crowded, but about evenly so throughout, not in fascicles at the summit only, the spreading blades much reduced; winter rosettes appearing early, the blades often 7 or 8 cm. long, glabrous or nearly so on the upper surface. This species is usually readily distinguished from P. lancearium by the decumbent habit and lax blades pubescent on both surfaces, but the blades are sometimes only obscurely pubescent above. Fic. 307.—P. patulum. From type specimen. DISTRIBUTION. Low moist woods of the Coastal Plain, southeastern Virginia to Florida and Louisiana. Virernta: Cape Henry, Chase 5434; Dismal Swamp, Chase 3674. Nortu Carourwa: Wilmington, Chase 3110, 4577. South Carona: Isle of Palms, Chase 4538, Hitchcock 413. Groraia: Thomasville, Small in 1895 (Biltmore Herb.). Fee ee eae Frorma: Jacksonville, Combs 3, _} e a Kearney 140; Lake City, Chase ( { 4282, Combs 132 in part; Mil- “od ton, Chase 4312; Old Town, hey joik i Combs 858, 859; Gainesville, Chase 4209; Grasmere, Combs 1169; Titusville, Chase 4026, Hitchcock 765; Eustis, Chase 4050, 4062, 4086; Nash 50, 151, 1117; Clearwater, Tracy 6701; Tampa, Hitchcock 946; Miami, Hitchcock 654, Tracy 8853; Levy County, Combs 783, 835; Palma Sola, Tracy 6729; Sneeds Island, Tracy 6703; Myers, Chase 4183, Hitchcock 922; Seminole, Tracy 7198. AtaBaMA: Mobile, Hitchcock in 1904 (Hitchcock Herb.). MississipPi: Biloxi, Chase 4357, 4371, Kearney 331, Tracy 4586, 4587; Horn Island, Tracy 3976. Lovuistana: New Orleans, Drummond 452. Fig. 308.—Distribution of P. patulum. 276 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 166. Panicum webberianum Nash. Panicum webberianum Nash, Bull. Torrey Club 23: 149. 1896. ‘‘Collected by the writer on the edge of a clay pit in the low pine land at Eustis, Lake Co., Florida, May 16-31, 1894, No. 781.’’ The type, in Nash’s herbarium, consists of two small tufts of vernal culms 35 to 45 cm. high, with blades as much as 1 cm. wide, and mature primary panicles, the spikelets 2.5 mm. long. Panicum onslowense Ashe, Journ. Elisha Mitchell Soc. 16: 88. 1900. ‘‘Type material was collected near Ward’s Mill,’’ Onslow County, N.C. The type, in Ashe’s herbarium, is the vernal form with immature panicles, the culms glabrous or minutely puberulent, the lower blades as much as 1 cm. wide, the immature spikelets 2.4 mm. long. Other specimens in Ashe’s herbarium and some distributed as P. onslowense and bearing the same data as the type are P. lancearium. DESCRIPTION. Vernal form commonly purplish; culms few to several in a tuft, rather stout, erect or ascending, 20 to 50 cm. high, minutely puberulent or glabrous; leaves somewhat crowded below, distant above; sheaths minutely puberulent at the summit, often ciliate on the margin, otherwise glabrous or nearly so; blades firm, ascending, espe- cially the lower somewhat incurved or spoon-shaped, 3 to 9 cm. long, 4 to 12 mm. wide, usually ciliate at base and sometimes along the margin, rounded or subcordate at base, acute but not long-acuminate; panicles finally long-exserted, 4 to 10 cm. long, two- thirds as wide, the numerous flexuous branches spreading or the lower even reflexed, the branchlets and pedicels usually rather short, thus giving to the main branches a somewhat racemose ee: spikelets 2.3 to 2.5 mm. long (in exceptional speci- mens only 2.1 to 2.2 mm. long), 1.2 to 1.3 mm. wide, obovoid to pyriform, commonly green, conspicuously purple-stained at the base; first glume one-third to two-fifths as long as the spikelet; second glume and sterile lemma mi- nutely pubescent or glabrous, the glumeslightly shorter than the fruit and sterile lemma; fruit = 1.9 to2 mm. long, 1.2 mm. wide, elliptic, under a lens minutely papillose-roughened, puberu- 1G. 309.—P. webberianum. From type lent at the Lae : specimen. Autumnal form spreading or decumbent, flabellately branching at the middleand upper nodes, the branches appressed and rather evenly distributed, sometimes somewhat fascicled; winter blades 3 to 8 cm. long, 5 to 10 mm. wide, strongly stained with purple, forming a flat rosette. As a whole this species is readily distinguished from P. lancearium by its stouter, taller culms, much larger blades, and in typical specimens by the larger spikelets, but a few specimens occur in which only the lower blades are much larger than in P. lan- ceartum, and rather numerous specimens in which the spikelets are only 2.1 to 2.2 mm. long. The minutely papillose-roughened fruit proves constant for all the specimens here referred to P. webberianum, but this character is evident only under a strong lens. DISTRIBUTION. Low pine land, North Carolina to Florida. Nort Carona: Onslow County, Ashe in 1899; Wilmington, Chase 45694, INtcheock 1433, 1472. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 277 Georaia: Brunswick, Ruth in 1893 (Ohio State Univ. Herb.). Fiorina: Jacksonville, Curtiss 4637 (Hitchcock Herb.), J. D. Smith 570; Lake City, Hitchcock 1024; Apalach- ; icola, Biltmore Herb. 6204a; Seville, Curtiss 6610; Titus- ville, Chase 3964; Hitchcock 7664; Grasmere, Combs 1164; Eustis, Chase 40514, Hitchcock 792, 811, Nash 781; Sanford, Hitchcock 782; Tampa, Hitch- cock 9304; Wimauma, Hitchcock 979; Arcadia, Hitchcock 855; Kalamazoo, Hitchcock 7654; Lakeland, Hitchcock 8384, 840; Braidentown, Hitchcock 955, 972, Tracy 6716; Johns Pass, Tracy 7186; Jensen, Hitchcock 735, 748; Myers, Chase 4163, Hitchcock 910, 918, Lee Co. Pl. 470; Miami, Hitchcock 630; without locality, Rugel 443. Fic. 310.—Distribution of P. webberianum. 167. Panicum patentifolium Nash. Panicum patentifolium Nash, Bull. Torrey Club 26: 574. 1899. ‘‘Type collected by the writer at Eustis, Lake Co., Florida, March 12-31, 1894, no. 72, in dry sand ina scrub hammock.’’ The type, in Nash’s herbarium, is the vernal form with two autumnal culms of the preceding year attached; all the blades are narrow, even the basal ones not over 4 mm. wide. DESCRIPTION. Vernal form often purplish throughout; culms several to many ina tuft, slender and wiry, widely decumbent-ascending, 25 to 55 cm. high, minutely puberulent or nearly glabrous; sheaths much shorter than the elongated internodes, a puberulent ring at the summit, otherwise glabrous or nearly so; blades stiffly spreading, 2.5 to 8 em. long, 2 to5 mm. wide, glabrous, acuminate, narrowed and sometimes ciliate toward the base; panicles commonly rather short-exserted, 3 to 7 cm. long, about half as wide, the branches few, ascending; spikelets 2.4 to 2.6 mm. long, 1.3 mm. wide, obovate, turgid; first glume one-third to half as long as the spikelet, obtuse or subacute; second glume and sterile lemma puberu- lent or nearly glabrous, the glume slightly shorter than the fruit and sterile lemma; fruit 2 mm. long, 1.2 mm. wide, elliptic, smooth and shining, mi- nutely puberulent at the apex. Autumnal form decumbent or spreading, Fra. 311.—P. patentifolium. From branching from the middle and upper nodes, the type specimen. branches appressed and somewhat elongated, the secondary branchlets shorter and more or less fascicled, not greatly reduced; winter rosettes appearing late, inconspicuous, the narrow blades ascending. This species differs from P. webberianum in the more slender culms, narrower, spreading culm blades, absence of the large basal blades, and less turgid spikelets in which the fruit is smooth and shining. 278 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Dry sand, especially in ‘‘scrub,”’ Georgia to Florida and Mississippi. Georaia: Dock Junction, Ricker 963 in part. Fiorina: St. Augustine, Ricker 945; Gainesville, Chase 4245; Cedar Key, Combs 777, 778; Eustis, Chase 4051, 4058, 4088, Hitchcock 796, 802, Nash 52, 72; Titusville, Chase 4028, 4029, Hitchcock 767; Ormond, Hitch- cock 563; Clearwater, Tracy 6702; Jensen, Hitchcock 730; Braidentown, Combs 1288, 1333, Hitchcock 969, 970, 971; East Pass, Tracy 6350; Mary Esther, Tracy 9139; Tampa, Hitchcock 930; Arcadia, Hitch- cock 856; Sebastian, Hitchcock 759; Myers, Chase 41634, 4170, Hitchcock 915%. AuaBAMA: Fort Morgan, Tracy Fig. 312.—Distribution of P. patentifolium. 7174. Misstsstprr: Biloxi, Kearney 330 in part. Oligosanthia.—Culms rather stout, usually erect; sheaths more or less hirsute, vil- lous, or sometimes glabrous; ligules inconspicuous except in P. ravenelii; blades firm, not over 2. cm. wide, usually narrower; spikelets about 3 to 4 mm. long, obovate, turgid, usually papillose-hirsute, strongly 7 to 9-nerved. Au- tumnal form with branches more or less crowded toward the summit. Nodes bearded; blades velvety-pubescent beneath. Plants lax, soft-velvety throughout; spikelets not over B Anitih (MOROre ee We let a8 Mee rae eG eee aie 169. P. — malacophyl- lum. Plants stiff, pubescence harsh; spikelets about 4 mm. lonees seems. 8 SSO ee eee 173. P. ravenelit. Nodes not bearded (or but obscurely so in P. wilcoxianum); blades not velvety. Panicle narrow, branches erect, or spreading only at anthesis; blades erect. Spikelets not over 3 mm. long; blades not over 6 mm. WAC. os oh doe me oR ne eee Se ...-- 168. P. wilcorianum. Spikelets 3.7 to 4 mm. long; blades 8 to 20 mm. wide. Blades papillose-hispid..............-..-..------ 174. P. leibergii. Blades glabrous on both surfaces. ...........--- 175. P. xanthophysum. Panicle about as wide as long. Spikelets narrowly obovate, subacute; plants oliva- ceous, appressed-pubescent.........--------- 172. P. oligosanthes. Spikelets broadly obovate, turgid, blunt; plants green, pubescence, if present, not appressed. Blades erect, not over 6 mm. wide; plants copi- ously hirsute throughout............----- 168. P. wileorianum. Blades ascending or spreading, rarely less than 8 mm. wide, usually wider; plants not hirsute throughout. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 279 Spikelets 3.2 to 3.3 mm. long; blades firm; sheaths or some of them more or less FO ee ae ee 171. P. scribnerianum. Spikelets not over 3 mm. long; blades rather thin; sheaths or some of them glabrous OO 2c ns a ee 170. P. helleri. 168. Panicum wilcoxianum Vasey. Panicum wilcoxianum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 32. 1889. ‘“‘Ne- braska (Dr. T. E. Wilcox).’’ The type, in the National Herbarium, consists of several vernal culms beginning to branch, 13 to 17 cm. high, with scarcely mature primary panicles. On the sheet is written in Vasey’s hand, ‘‘ Panicum Wilcoxianum, Vasey n. sp., Niobrara Ft., Nebraska. Dr. T. E. Wilcox, 1888.’’ DESCRIPTION. Vernal form dull green; culms usually in dense tufts, erect, 10 to 25 cm. high, copi- ously papillose-hirsute, as are the rather loose, usually overlapping sheaths; ligules about 1 mm. long; blades firm, erect or ascending, 5 to 8 cm. long, 3 to 6 mm. wide, broadest toward the base (this scarcely wider than the wide sheath), commonly involute-acuminate, long-hirsute on both surfaces; panicles finally exserted, often equaled or exceeded by the upper blades, 2 to 5 em. long, about half as wide, or some- times more expanded at anthesis, rather densely flowered; spikelets 2.7 to 3 mm. long, 1.5 mm. wide, obovate-elliptic, papil- lose-pubescent; first glume about one-third as long as the spikelet, pointed or obtuse; second glume slightly shorter than the fruit and Girila lemma; fruit 2.4 to 2.5 mm. long, 1.3 to 1.4 mm. wide, elliptic. Autumnal form banking from all the nodes, forming bushy tufts with rigid, erect blades much overtopping the reduced panicles; branches appearing early, usually before the maturity of the primary panicles; secondary spikelets usually more turgid than those of the primary panicles. Fig. 313.—P. wilcorianum. From type specimen. DISTRIBUTION. Prairies, Manitoba to North Dakota and south to Iowa and Kansas. Miynesora: Winona, Holzinger 28 in part (Biltmore Herb. ). Manrrosa: Sewell, Macoun 13227. Norto Daxora: Towner, Lunell in 1908. Souta Daxora: Brookings, Williams in 1891, E. N. Wilcox 14; Roberts County, S. D. Agr. Col. & Exp. Sta. 4167; Rosebud, Wallace in 1896; Jamesville, Bruce 80. Iowa: Missouri Valley, Pammel 3198; Gilbert Station, Carver in 1894. Fic. 314.—Distribution of P. wilcorianum. 280 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. NesrasKka: Niobrara Fort, 7. £. Wilcox in 1888; Johnstown, Bates 1084; Thed- ford, Rydberg 1308; Weigand, Clements 2683. Kansas: Manhattan, Hitchcock 2505, Pl. Kan. 879; Courtland, Hitchcock in 1892. Cotorapo: Without locality, American Plains Flora, Hall 231 in 1863 (Mo. Bot. Gard. Herb.). 169. Panicum malacophyllum Nash. Panicum scoparium minor[us]Scribn. Tenn. Agr. Exp. Sta. Bull.'7: 48.1894. ‘‘Mid- dle Tennessee (Gattinger).’’ The type, in the herbarium of the University of Tennes- see, consists of four branching culms with a primary panicle from which the spikelets have fallen and numerous secondary panicles with pilose branchlets and spikelets. On the accompanying label, which reads ‘‘Panicum scoparium Lam. Cedar Glades, Lavergne, Tennessee. Autumnal form 7 VII. ’81. Legit Dr. A. Gattinger,’”’ is written in Scribner’s hand ‘‘var.‘mninor,” and this is the only Gattinger and the only Tennessee specimen so marked by him. No specimen of P. scribnerianum4 can be found which was referred by Scribner to his P. scoparium var. minor, while a specimen collected by Coville, Mountain Park, Arkansas, in 1887, corresponding to the above Gattinger specimen (that is P. malacophyllum Nash), is marked by him ‘‘Panicum scoparium Lam. var. minor Scribn.’’ The description applies to the species repre- sented by the Gattinger specimen except as to the panicle and spikelets: ‘‘ Panicle branches and spikelets nearly smooth, or (subvar. piloswm) densely pilese hairy.”’ Only the Gattinger specimen is cited and this, marked by Scribner as noted above, has pilose spikelets and panicle branches. No specimen can be found marked with the subvarietal name. Were it not that Scribner wrote ‘‘var. minor” on two specimens of P. malacophyllum Nash and on nothing else, the Gattinger citation might possibly be taken to refer to ‘‘subvar. pilosum.’’ It would appear that the author? confused P. scribnerianum and P. malacophyllum; that his description, drawn up from the material in his herbarium, was made to cover both, but more especially the common form, but that, having no Tennessee specimens of the common form he cited a specimen of the form he did have from Tennessee. The author’s remark that ‘‘this is the most widely distributed and best known form of the species” bears out this conclusion. Panicum malacophyllum Nash, Bull. Torrey Club 24: 198. 1897. ‘‘Type collected by Mr. B. F. Bush on May 19, 1895, at Sapulpa, Indian Territory, No. 1228.”’ The type, in Nash’s herba- rium, consists of two early autumnal culms 28 and 35 cm. high, with ma- ture primary panicles. DESCRIPTION. Vernal form velvety or velvety- pilose throughout; culms slender, ; few to several in tufts, 25 to 70 cm. Fig. 315.—P. malacophyllum. From type specimen. high, more or less geniculate at base with arched internodes, ascend- ing or spreading, papillose-pilose with soft, reflexed hairs, the nodes retrorsely bearded; @ See discussion on the type of P. scribnerianum. bAn examination of the specimens of this group in Scribner’s herbarium shows that at that time his idea of true P. scoparium Lam. was P. ravenelii, on a sheet of which he had written ‘Certainly P. scoparium Ell., a form which suggests close relationship with P. Waltert Poir. (P. latifolium Mx.).”’ Accepting Elliott’s interpretation of Lamarck’s species Scribner names this form var. genwinum; his var. pauciflorum (based on P. pauciflorum Ell.) is P. oligosanthes Schult.; his specimens of P. scribnerianum, ten in all (none of them from Tennessee), are all marked in Scribner’s writing P. scoparium Lam., without particular comment, and none as stated above is marked ‘‘var. minor.’’ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 281 sheaths loose, shorter than the internodes, usually less copiously pilose than the culm; ligules 1 to 1.5 mm. long; blades spreading or ascending, 7 to 10 cm. long, 6 to 12 mm. wide, tapering to the rounded base, acuminate, rather thin, velvety on both surfaces, ciliate at least toward the base; panicles usually short-exserted, 3 to 7 cm. long, at first narrow, the lower branches finally spreading, with short, spikelet-bearing branch- lets in the axils; spikelets 2.9 to 3 mm. long, 1.5 to 1.7 mm. wide, elliptic-obovate, obscurely pointed, turgid at maturity, papillose-pilose; first glume about one-third as long as the spikelet; second glume and sterile lemma equaling the fruit at maturity; fruit 2.2 mm. long, 1.5 mm. wide, elliptic. Autumnal form spreading, freely branching from the middle and upper nodes before the maturity of the primary panicle, at length forming bushy, topheavy clumps with reduced blades and numerous secondary panicles. DISTRIBUTION. Sandy woods, Tennessee and Mis- souri to Oklahoma and Texas. Missourtr: Warrenburg, Stigall 8. TENNESSEE: Nashville, Gattinger in 1880; Lavergne, Gattinger in 1881 and 1882. ARKANSAS: Mountain Park, Coville in 1887; Prescott, Bush 263. Texas: Dallas, Bush 642, Reverchon 1831; Weatherford, Tracy 7942; Denison, Bebb 2670 (Hitchcock Herb.). OxtaHomMa: On the False Washita, Palmer 383 in 1868; Wister, Hitchcock 578. Fic. 316.—Distribution of P. malacophyllum. 170. Panicum helleri Nash. Ponicum helleri Nash, Bull. Torrey Club 26: 572. 1899. ‘‘Collected at Kerrville, Kerr Co., Texas, by A. A. Heller, May 14-21, 1894, No. 1759. Differs from P. perner- vosum in the pubescent culm and sheaths, the broader blades of a different shape and the narrow spikelets which are usually sparsely pubescent.’’ The type,in Nash’s herbarium, is a tuft of five vernal culms 24 to 39 cm. high, the lower nodes subgeni- culate; panicles immature, scarcely exserted, the upper spikelets well developed but not mature, 3 mm. long, 1.5 mm. wide, bearing a few scattered hairs; papille on the sheaths not prominent; lower internodes sparsely ascending-pilose, the upper short-pubescent, as frequently found in P. scribnerianum. The blades are 6 to 12 mm. wide, while those of the type of P. pernervosum are 5 to 7 mm. wide, but both are of the same shape, broadest in the middle instead of near the base, as in P. scribnerianum. Panicum pernervosum Nash, Bull. Torrey Club 26: 576. 1899. ‘‘Type collected by Elihu Hall in woods at Houston, Texas, April 16, 1872, No. 830.’’ The type, in the herbarium of the New York Botanical Garden, consists of two vernal plants with culms 27 and 34 cm. high, and an extra piece of culm with a panicle. The culms are glabrous, the sheaths ciliate on the margin, otherwise glabrous, except one which has a few hairs near the summit. The panicles are mature and the spikelets more turgid than in the type of P. helleri, being 3 mm. long and 1.7 mm. wide. A few of them bear a few scattered hairs. DESCRIPTION. Vernal form in clumps of few to several culms, usually somewhat bluish light green; culms 25 to 60 cm. high, slender, ascending or spreading, the lower internodes appressed -pilose, the upper often glabrous; sheaths sparsely papillose-hispid to glabrous, the papille often without hairs as in P. scribnerianum, ciliate on the margin; ligules 282 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. about 1 mm. long; blades ascending or spreading, measuring about the same as those of P. scribnerianum, but broadest about the middle, rather thin, glabrous on both sur- faces or pubescent beneath, ciliate toward the base; panicles finally rather long- exserted, 6 to 12 cm. long, about three-fourths as wide, more open and loosely flowered than in P. scribnerianum; spikelets 2.9 to 3 mm. long, 1.6 to 1.7 mm. wide, obovate, turgid, blunt, glabrous, or with few scattered hairs; first glume about one-third the length of the spikelet, acute; second glume and sterile lemma subequal, slightly exposing the fruit at maturity, strongly nerved; fruit 2.4 to 2.5 mm. long, 1.5 to 1.6 mm. wide, oval, obscurely apiculate. Autumnal form branching at all but the lowest nodes, forming loose sprawling tufts, the branches somewhat divaricate, with sheaths more commonly pubescent than those of the primary culm, the blades widely spreading, not much reduced, the long- pediceled spikelets rather conspicuous among the foliage. This species is closely ‘feladea to P. scribnerianum, and many specimens are but doubtfully differentiated. As here distinguished, the smaller spikelets, thin- ner blades tapering to both ends, and the j lax habit, taken in combination, have Fic. 317.—P. helleri. From type specimen. been used to separate P. helleri. Asin P. scribnerianum little weight can be given to pubescence or lack of it, hispid and glabrous sheaths being found on the same plant, as in Bush 803 and Hitchcock 1173, though P. helleri is more commonly nearly glabrous than is P. scribnerianum. Reverchon’s no. 2857 is probably referable to this species, but the pubescence of culms, sheaths, and lower surface of blades and the long hairs mixed with the short ligule show affinity to P. oligosanthes. The spikelets are 3 mm. long, too immature to show turgidity. A few specimens are intermediate in the size of the spikelets. In Harvey 17 and Hitchcock 1173 the spikelets are 3.1 to 3.2 mm. long; and Hitchcock 1223, with spikelets 3.3 mm. long, is referred here since the specimen shows the sprawling habit of P. helleri. DISTRIBUTION. Open woods and prairies, Missouri and Oklahoma to Louisiana and New Mexico. Missourt: Sheffield, Bush 803, 3903; Levasy, Bush 1685; Courtney, Bush 1710, 3893. ARKANSAS: Fulton, Bush 2529; northern Arkansas, Harvey 17. Loutstana: Cameron, Tracy 8419. Texas: Dallas, Bush 705, Rever- chon 1074 in part, 2444; Dallas County, Reverchon 2342, 2344, 2345; Denison, Bebb 1430; Weatherford, Tracy 7949; Cor- sicana, Reverchon 2855 in part; Grand Saline, Reverchon 4138; Terrell, Warburton 2; Waller, Hitchcock 1173, 1179, 1186, 1212; Waller County, Thurow in 1898; Houston, Bebb 1278, Hall 830, Nealley in 1887; Gal- veston, Plank 82; : Virginia Point, Bray 33; Kerrville, Heller 1759, Smith in 1897; Burnet, Plank 3; Velasco, Smith in 1897; Victoria, Tracy 8870; Kings- ville, Tracy 8885; without locality, Buckley in 1881. OKLAHOMA: On the False Washita, Palmer 382 in 1868. New Mexico: Mogollon Mountains, Metcalfe 354. Fig. 318.—Distribution of P. helleri. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. 283 171. Panicum scribnerianum Nash. Panicum macrocarpon Torr. Fl. North. & Mid. U. 8. 143. 1823, not LeConte 1819. “Has. On the banks of the Connecticut River, near Deerfield, Massachusetts. Sent to me by Dr. Cooley.’’ The type, in herbarium of Columbia University, is a single culm with terminal panicle. It is labeled in Torrey’s hand “‘ Panicwm macrocarpon*,”’ followed by a brief diagnosis, and ‘‘ Near Deerfield, Mass. Dr. Cooley.’’ On the same sheet was mounted the specimen of P. latifolium, which is taken as the type of P. macrocarpon LeConte.¢ Torrey makes no mention of P. macrocarpon LeConte, published a few years earlier in his Catalogue of the Plants of New York. Panicum scribnerianum Nash, Bull. Torrey Club 22: 421. 1895. This is proposed as a new name without description, the following citations being given: “ Panicum scoparium 8. Wats. in A. Gray, Man. Ed. 6, 632.1890. Not Lam. Panicum scoparium var. minor Scribn. Bull. Univ. Tenn. 7: 48. 1894. Not P. capillare var. minor Muhl. 1817.’ The author does not state upon which of these two the new name is based, but since it ‘‘is proposed in honor of Prof. F. L. Scribner, who was the first to indicate its difference from P. scoparium Lam.,’’ it seems evident that the intention is to raise Scribner’s variety to specific rank, changing the name because of P. capillare var. minor Muhl. But examination of Scribner’s type > shows that it is not the species described as P. scoparium in Gray’s Manual and as P. scribnerianum by Nash in the Illustrated Flora,¢ Britton’s Manual,¢ and Small’s Flora.e Owing to the confusion and uncertainty arising from Scribner’s citing a specimen which disagrees in part with his description, it seems best to take the first citation given by Nash as the basis of P. scribnerianum, excluding the reference to P. scoparium var. minor. Panicum scoparium 8. Wats.; Nash, Bull. Torrey Club 22: 421.1895. As synonym under P. scribnerianum Nash. The name is cited by Nash as ‘‘S. Wats. in A. Gray Man. Ed. 6, 632. 1890. Not Lam.,’’ but Watson did not publish this name, since misapplication of a name does not constitute publication. The description of “ P. scoparium Lam.’’ [misapplied] in Gray’s Manual, ed. 6, is identical with that of “P. pauciflorum Ell.?”’ of previous editions back to the first. In the first edition / the range is given as ‘‘ N. Pennsylvania (Carey) and W. New Yorkto Michigan.’’ The Carey specimen, in the Gray Herbarium, is a single branching plant with hispid sheaths, a primary panicle, destitute of spikelets, and two secondary panicles with scarcely mature spikelets. The accompanying label reads: ‘‘ Panicum n. sp.? pauci- florum Ell.? Wysox. Penna. J. Carey, July 1836.’’ This specimen we take as the type of P. scribnerianum. DESCRIPTION. Vernal form in clumps of few to many culms, 20 to 50 cm. high, erect or ascending, often geniculate at base, glabrous or harshly puberulent or sometimes ascending papillose-pilose; sheaths rather loose, conspicuously striate, ciliate on the margin, ascending-pubescent between the nerves and papillose-hispid with spreading or ascending hairs to nearly glabrous, the papille often without hairs; ligules about 1 mm. long; blades ascending or erect, 5 to 10 cm. long, 6 to 12 mm. wide, usually firm, acuminate, rounded and ciliate at base, glabrous on the upper surface, appressed- pubescent to glabrous beneath; panicles short-exserted, 4 to 8 cm. long, rarely longer, two-thirds to three-fourths as wide, the flexuous branches ascending; spikelets 3.2 to 3.3 mm. long, 2 mm. wide, obovate, turgid, blunt, sparsely pubescent to nearly 4See note under P. latifolium L., page 314. . 6See notes on P. scoparium minus Scribn. under P. malacophyllum Nash, page 280. ¢ Britt. & Brown, Lllust. Fl. 1: 118. 1896. d@ Man. 87. 1901. ¢Fl. Southeast U.S. 103. 1903. 7 A. Gray, Man. 613. 1848. 284 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. glabrous; first glume about one-third the length of the spikelet, acute; second glume and sterile lemma subequal, barely or scarcely equaling the fruit at maturity, strongly nerved; fruit 2.8 to 2.9 mm. long, 1.8 to 1.9 mm. wide, broadly elliptic, minutely apiculate. Autumnal form branching from the middle and upper nodes at about the maturity of the primary panicle; the branches longer than the internodes, and late in the season producing crowded branchlets with ascending, not greatly re- duced, blades and small, partially included, panicles from their up- per nodes. This species is very variable in the matter of pubescence. An un- usually hispid Washington speci- men, Elmer 763, was considered worthy of varietal rank by Scrib- ner and Merrill and bears a her- barium name, but Moffatt 1863, Miller, Indiana, is quite and others nearly as hispid, while other Pacific coast specimens are not more hispid than eastern specimens. Glabrous and hispid sheaths are commonly found on the same specimen in this species. A few Mississippi specimens with blades softly pubescent beneath and spikelets 3.2 to 3.6 mm. long, suggest an approach to P. ravenelit Scribn. & Merr. These are: Jackson, Hitchcock 1295, 1296; Starkville, Tracy 42, 1752. Fia. 319.—P. scribnerianum. From type specimen. DISTRIBUTION. Sandy soil or dry prairies, Maine, Ontario, and westward to the Pacific; south to Maryland and Arizona. Marne: South Berwick, Parlin in Harvey, Maine Weeds 1245. New Hampesurre: Walpole, Fernald 280 (Gray Herb.). Vermont: Westminster, Robinson in 1898. Massacuusetts: Wellesley, Smith 731; Ipswich, Oakes; Essex County, Conant in 1881. Connecticut: Oxford, Harger in Kneucker Gram. Exs. 425; Rocky Hill, Wilson 1256; Farmington, Bissell 5563; Southington, Andrews 51. RuopeE Istanp: Providence, Collins in 1887 (Brown Univ. Herb.). New York: Ithaca, Ashe. Ontario: Sarnia, Macown 26328; Sandwich, Macown 26329. New Jersey: Glenlock, Heritage in 1897; Morris Plains, Mackenzie 1611; Passaic County, Nash in 1893. PENNSYLVANIA: Safe Harbor, Porter in 1864; Germantown, Stone 4 in 1889; Easton, Porter in 1895, 1896, and 1898. Outro: Erie County, Mertz 147; Cedar Point, York 6789; Bowling Green, Keller- man 6894. Inprana: Miller, Moffatt 1863, Umbach in 1897; Elston, Dorner 88. InuiNois: Hanover, Gleason & Gates 2598; Beach, Umbach 2365; Chicago, Chase in 1896; Joliet, Skeels 250; Romeo, Umbach 1704; Starved Rock, Chase 1607, Greenman, Lansing & Dixon 156; Wady Petra, V. H. Chase 922; Marshall County, V. H. Chase 1792; Galva, V. H. Chase 1749; Peoria, McDonald 32. Micnican: Grand Rapids, Crozier; without locality, Wheeler 97. Minnesota: Fort Snelling, Mearns 771; Spring Grove, Rosendahl 260; Minneapolis, Sheldon in 1895. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 285 Nortx Daxora: Norfolk, Griffiths 871. SoutH Daxorta: Lead City, Rydberg 1098; Rosebud, Wallace 12. Iowa: Ames, Ball 178; Iowa City, Somes 167, 229. NEBRASKA: Weeping Water, Williams 3009; Thedford, Rydberg 1279, 1493; Fort Niobrara, T. E. Wilcox in 1890; Mullen, Rydberg 1604. Missouri: Wayne County, Eggert 234; Carter County, Eggert 297; St. Louis, Eggert 249; Washington County, Eggert 295; Monteer, Bush 722, 733 in part, 4653; Independence, Bush 729, 740; Watson, Bush 737; Dodson, Bush 1659; Arlington, Kellogg in 1903; Allenton, Kellogg in 1903; Swan, Bush 4568; Westport, Bush 4021; Vale, Bush 3933. Kansas: Osborne City, Shear 85; Manhattan, Hitchcock 2383, 2502, 2511, 2519, 2525, 3854, Pl. Kan. 571, 571a, 921. DELAWARE: Centerville, Commons 281, 282, Chase 3618. Marytanp: Glen Echo, House 831; High Island, Ward in 1879; Great Falls, Chase 2865. District or CoLuMBIA: Chase 3808, Steele in 1901, Ward in 1881 and 1883. TENNESSEE: Knoxville, Ruth 74 in 1898. ArKANSAS: Benton County, Plank 48a, 93. Texas: Denison, Bebb 2715 (Hitch- cock Herb.); without locality, Johnson in 1886. Ox.aHoMA: Limestone Gap, Butler 19; Sapulpa, Bush 1216, 1222; Flora, Bebb 2167; Walker, Bebb 1459. Wyomine: Cambria Canyon, A. Nelson 2524; Inyankara Creek, Williams 2577; Devils Tower, Griffiths 548; Whalen Canyon, A. Nelson 516. Ipano: Nez Perces County, Sandberg, Heller & MacDougal in 1892. WasuHINcTon: Wawawai, Elmer 763; Klickitat County, Suksdorf in 1882 and 1883; Ophir, Elmer 509; Thurston County, Heller 4058; Roy, Hunter 604a; Walla Walla, Wilkes Expl. Exped. 946. British Cotumstra: Chilliwack Valley, Macoun 26333, 77230; Lake Osoyoos, Macoun 77231; Vancouver Island, Macoun 29297. OreGon: Cascades, Kellogg & Harford 1085; Cache Bar, Sheldon 8331; Coos County, Hitchcock 2836; Sauvies Island, Howell 63; Snake River at mouth of Cache Creek, Sheldon 8194; without locality, Hall. Pl. Oreg. 672. Cotorapo: Fort Collins, Baker 36. Arizona: Willow Spring, Palmer 561 in 1890; White Mountains, Griffiths 5401. CalirorniA: Castle Crag, Hitchcock 3074. Fig. 320.—Distribution of P. scribnerianum, 172. Panicum oligosanthes Schult. Panicum pauciflorum Ell. Bot. 8. C. & Ga. 1: 120. 1816, not R. Br. 1810. ‘‘In Georgia, not very rare.’’ The type, in the Elliott Herbarium, is a single culm begin- ning to branch, with five leaves and an exserted panicle 6 cm. long and 6.5 cm. wide. The accompanying label reads: ‘‘Panicum pauciflorum E. Hab: in humidis umbro- sisque. Flor. Ap: Ma.’’ Panicum oligosanthes 4 Schult. Mant. 2: 256. 1824. Based on P. pauciflorum EIl., the name being changed because of P. pauciflorum R. Br. @ This name is incorrectly formed; the word should be oliganthum, but since the incorrect. form has been sanctioned by usage it seems wiser not now to correct it, especially since to do so would invalidate P. oliganthum Schlecht. 1854. 286 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum scoparium angustifolium4 Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8: 32. 1889. ‘‘South Carolina (Dr. Ravenel)’’ is the first specimen cited. This, which is taken as the type, is in the National Herbarium, and consists of several culms in the early branching state, with mature terminal panicles. Panicum scoparium pauciflorum Scribn. Tenn. Agr. Exp. Sta. Bull. 7: 48. 1894. Based on P. pauciflorum Ell. DESCRIPTION. Vernal form olivaceous, in loose tufts of few to several culms 35 to 80 cm. high, erect, often purplish, appressed-pubescent, especially below; sheaths shorter than the internodes or the lower longer, the papillose pubescence ascending; hairs of the ligule 1 to 2mm. long, with longer ones intermixed; blades stiffly spreading or ascending, 6 to 14 cm. long, 5 to 8 (rarely 10) mm. wide, sharply acuminate, narrowed toward the base, glabrous on the upper surface or rarely with a few long hairs, harshly puberulent beneath, stiffly ciliate near the base; panicles finally long-exserted, 6 to 12 em. long, about as wide, loosely flowered, branches usually stiffly ascending or spreading; spike- lets long-pediceled, 3.5 to 4 mm. long, 1.7 to 1.9 mm. wide (smaller in ex- ceptional specimens), oblong-obovate, subacute, sparsely hirsute; first glume less than half the length of the spike- let, acute; second glume slightly shorter than the fruit and sterile lemma; fruit 2.8 to 3 mm. long, 1.5 to 1.6 mm. wide, elliptic. Autumnal form erect or spreading, sometimes topheavy-prostrate, branch- ing sparingly from the lower, freely from the upper, nodes, late in the season the short branchlets aggregated at the summit of the branches, the crowded, reduced blades widely spreading, the panicles more or less included and reduced toa few spikelets, these commonly more turgid and blunt than those of the primary panicle. In this species the spikelets vary more in size than usual in this group. The follow- ing specimens have spikelets only 3.2 to 3.8 mm. long: Bebb 1426, Bush 1225, Hitch- cock 1194, Kearney 1386, Reverchon 1840, 4142, and Thurow in 1899. The shape of the spikelets as well as the other characters of these specimens are those of P. oligosan- thes. A few specimens, however, appear to be intermediate between this and P. scribnerianum, having the narrow blades, appressed pubescence, and open, few-flowered panicles of P. oligosanthes, but very turgid, blunt spikelets, which, however, measure 3.5 to 3.7mm.long. These are: Tracy 1754; Bebb 1389 and 2703, Denison, Texas; and Hitchcock in 1903, Wister, Oklahoma (the last two in Hitchcock Herb.). Fic. 321.—P. oligosanthes. From type specimen of P. pauciflorum Ell. DISTRIBUTION. Sandy, usually moist woods, New Jersey to Illinois, south to Florida and Texas, mostly near the coast. New Jersny: Atsion, Commons 54. Inp1ANA: Dune Park, Hill 201 in 1898. Inurwors: [Wabash County?] Schneck in 1879. a This is not based on P. angustifolium Ell. Vasey gives the latter species on page 29 of the same paper. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 287 DELAWARE: Seaford, Commons in 1882; Greenbank, Commons in 1883; Lewes, Hitchcock 419, 582. District or CoLtumBiA: Steele in 1899. VirerniA: Fort Monroe, Vasey in 1878 and 1883; Cape Henry, Chase 2912, 5435, Kearney 1400; Virginia Beach, Hitchcock 581, Kearney 1386, Williams 3110. Norts Carouina: Wilmington, Hitchcock 418, 1457, 1459, 1460. Souts Carona: Aiken, Hitchcock 580, Ravenel in 1867, Scribner in 1894; Sumter, Hitchcock 579; Orangeburg, Hitchcock 20, 1410. Georeis: Augusta, Cuthbert 1121, 1167, Kearney 216; Clarke County, Harper 146; Stone Mountain, Hitchcock 417. Fiorina: Jacksonville, Curtiss 5864; Lake City, Chase 4276, 4283, Combs 163, Hitchcock 1014; Tallahassee, Combs 382; Gainesville, Chase 4207, 4260, Combs 742. AuaBAMA: Flomaton, Hitchcock FIG. 322.—Distribution of P. oligosanthes. 1055. Mississippi: Columbus, Tracy in 1896; Starkville, Tracy 1754; Waynesboro, Kearney 191; Biloxi, Tracy 3647; Mississippi City, Hitchcock 1106. ARKANSAS: Benton County, Plank 73. Louistana: Calhoun, Ball 64, Hitchcock 1263, 12744, 1293. Texas: Dallas County, Reverchon 1841; Denison, Bebb 1389, 1426; Grand Saline, Reverchon 4142; Silver Lake, Reverchon 1840; Waller, Hitchcock 1194, Thurow in 1898 and 1899; Houston, Bebb 1245, Nealley in 1887; without locality, Wright (Gray Herb.). OKLAHOMA: Sapulpa, Bush 1225. 173. Panicum ravenelii Scribn. & Merr. Panicum scoparium major[us] Vasey, U. S. Dept. Agr. Div. Bot. Bull. 8: 32. 1889 The author states, ‘‘We have only seen this from South Carolina (Dr. Ravenel).”’ The type, in the National Herbarium, consists of three immature vernal culms. Panicum scoparium genuinum Scribn. Tenn. Agr. Exp. Sta. Bull. 7: 48. 1894. “P. scoparium Lam., Ell.’’ is cited, and, as it is evident that Scribner is designating Elliott’s as the genuine P. scoparium, Elliott's specimen is considered the type. Panicum ravenelii Scribn. & Merr. U.S. Dept. Agr. Div. Agrost. Bull. 24: 36. 1901. Published as a new name for ‘‘ Panicum scoparium of Ell. Sk. Bot. 8. C. and Ga. 1: 119. 1817. Not Lam. Encycl. 4: 744. 1797.’’ The type, in the Elliott Herbarium, is a single vernal culm with an immature panicle. No locality is cited by Elliott, and none is given on the label; the specimen is presumably from South Carolina. This species was referred to Panicum scoparium by Elliott, as stated above, and also by Chapman.¢ DESCRIPTION. Vernal form in loose tufts, grayish olive green; culms 30 to 70 cm. high, erect or ascending, densely papillose-hirsute with ascending hairs, the nodes short-bearded; sheaths shorter than the long lower internodes, about equaling the short upper ones or overlapping, papillose-hirsute like the culm; ligules 3 to 4 mm. long; blades thick, ascending or spreading, 8 to 15 cm. long, 1 to 2 cm. wide, sharply acuminate, rounded a2 FI, South. U.S. 575. 1860. 288 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. at the base, glabrous on the upper surface, densely velvety-hirsute beneath, usually short-ciliate nearly to the apex; panicles short-exserted or included at the base, 7 to 12 cm. long, as wide or wider, loosely flowered, the branches finally spreading; spikelets 4 to 4.3 mm. long, 2 to 2.2mm. wide, cbhovate, turgid and blunt, sparsely papillose-pubescent; first glume one-third to two- fifths the length of the spikelet, acute; second glume and sterile lemma subequal,scarcely equal- ing the fruit at maturity, strongly nerved; fruit 3.2 mm. long, 2 mm. wide, elliptic, mi- nutely apiculate. Autumnal form more or less FiG. 323.—P. ravenelii. From type specimen. spreading, branching from the middle and upper nodes, the short branchlets crowded at the summit late in the season, the reduced blades ascending, overtopping the small panicles. DISTRIBUTION. Sandy or gravelly woods or open ground, Maryland to Missouri, Florida, and Texas. Missouri: Swan, Bush in 1899. DeLaware: New Castle County, Canby (Field Mus. Herb.). MaryLanp: Great Falls, Chase 3780; Riverdale, Chase 3796. District or CoLtumBIA: Steele in 1898 and 1900; Ward in 1881. VirciniA: Danville, Small & Heller in 1891 (Biltmore Herb.). Nortu Carona: Wilmington, Hitchcock 420, 1463, 1489; Raleigh, Chase 3088; Chapel Hill, Ashe. Sout Carouina: Aiken, Ravenel; Orangeburg, Hitchcock 1390; Keowee, House 2200. Geroreia: Cobb County, Wilson 31, 47; Thomson, Bartlett 1502. Fioripwa: Apalachicola, Biltmore Herb. 2994a; Mc aticello, Combs 305; Chipley, Combs 602 in part; Lake City, Chase 4279, Mitchcock 1010. TENNESSEE: Henderson, Bain in 1893 (Univ. Tenn. Herb.). AuaBaAMA: Mobile County, Mohr 153; Flomaton, Hitchcock 1047; Auburn, Hiteh- cock 1334; Tuskegee, Carver 18. Mississippi: Starkville, Tracy in 1888; Jackson, Hitchcock 1294; Mississippi City, Hitchcock 1107. Arxansas: Little Rock, Coville in 1887; Fulton, Bush 2526; Benton County, Plank 56, 143; northwestern Arkansas, Harvey 31. Lovistana: Calhoun, Hitchcock 1288; Shreveport, Hitchcock 1245. Texas: Big Sandy, Reverchon 2390; Harvester, Thurow in 1898; Waller, Hitch- cock 1187; without locality, Nealley in 1885, Wright 280 (Gray Herb.). Fig. 324.—Distribution of P. ravenelii. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 289 174. Panicum leibergii (Vasey) Scribn. Panicum scoparium liebergii Vasey, U. S. Dept. Agr. Div. Bot. Bull. 8: 32. 1889. **P. liebergii Scribn.)” is given as synonym, and ‘‘Plymouth County, Iowa (John Lieberg)” [error for Leiberg] is cited. The type, consisting of two vernal culms, is in Hitchcock’s herbarium. The accompanying label reads: ‘‘Panicum Leibergii, Scribn. 1884. Plymouth Co., Iowa. 1878. Legit John Leiberg,’’ the specific name, author, and date being in Scribner’s writing. On the sheet is written in Vasey’s hand, ‘‘P. scoparium var. Leibergii V.’’ Panicum scribnerianum leibergii Scribn. U. 8. Dept. Agr. Div. Agrost. Bull. 6: 32. 1897. Nosynonym is cited and ‘‘South Dakota: Brookings (Wilcox 16)” is the only specimen mentioned. This name must however be considered a new combina- tion based on P. scoparium leibergii Vasey. The Wilcox specimen agrees with that of Leiberg. Panicum liebergii Scribn. in Britt. & Brown, Illust. Fl. 3: 497. 1898. Based on **Panicum scoparium Lam. var. Liebergii Vasey.’’ This name was first mentioned as a synonym under P. scoparium liebergii Vasey.4 DESCRIPTION. Vernal form dull green, in clumps of few to several slender culms, 25 to 75 cm. high, erect from a more or less geniculate base, pilose to merely scabrous; sheaths shorter than the internodes, papillose-hispid with spreading hairs; ligules almost obsolete; blades ascending or erect, rather thin, 6 to 15 cm. long, 7 to 15 mm. wide, narrowed toward the rounded base, acuminate, papillose-hispid on both. surfaces, often sparsely so above, papillose-ciliate from one-third to half their length; panicles finally long-exserted, 8 to 15 cm. long, less than half as wide, the flexuous branches narrowly ascending or somewhat spreading at anthesis; spikelets 3.7 to 4 Fia. 325.—P. leibergii. From type specimen. mm. long, 1.8 to 2 mm. wide, oblong-obovate, turgid, strongly papillose-hispid with spreading hairs; first glume more than half the length of the spikelet, pointed, second glume and sterile lemma subequal, covering the fruit at maturity or the lemma slightly exceeding it; fruit 3 mm. long, 1.7 to 1.8 mm. wide, obovate-oval. Autumnal form more or less leaning, sparingly branching from the middle and lower nodes late in summer, the branches mostly simple, erect, the blades scarcely reduced, usually exceeding the short-exserted panicles. 2U.8. Dept. Agr. Div. Bot. Bull. 8: 32. 1889. 41616°—vo1 15—10——19 290 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Prairies, New York to Manitoba, south to Ohio and Kansas. New York: Head of Seneca Lake (Gray Herb.). Ontario: Squirrel Island, Lake St. Clair, Dodge 17, 20, 62, 84. Onto: Erie County, Moseley in 1897. InpiaAna: Lafayette, Dorner 35. Inurnois: Naperville, Umbach 1669; Emington, Wilcox 129; Joliet, Skeels 334; Wady Petra, V. H. Chase 461, 1455; Knox County, V. H. Chase 1701; Peoria, McDonald 17; Champaign, Waite in 1888. Micnican: Hansens Island, Lake St. Clair, Dodge in 1899 (Hitchcock Herb.). Wisconsin: Racine, Wadmond 46; Elkhart Lake, Hill 91 in 1906. Minnesota: Acton, Frost in 1892; St. Cloud, Campbell in 1887; Montevideo, Moyer in 1894; Spring Grove, Rosendahl 504, 538. ManitoBa: Macgregor, Macoun 73004; Carberry, Macoun 73003. Norta Daxora: Fargo, Bolley 1865; Merrifield, Brannon 10; Leeds, Lunell in 1902. Souta Daxora: Brookings, EH. N. Wilcox 16, Williams 2228; Lake Hendricks, Williams in 1891; Simpson Park, Griffiths 836. Iowa: Plymouth County, Leiberg in 1878; Armstrong, Cratty in 1900; Ames, Ball & Sample 16; Iowa City, Somes 236; De- Fic. 326.—Distribution of P. leibergii. catur County, Fitzpatrick & Fitzpatrick 36; New Albin, Pammel 932; Johnson County, Shimek 69. NesrASKA: Ponca, Clements 2523. Missourtr: Lees Summit, Mackenzie 297; Monteer, Bush 379, 730, 744, 2760; Dod- son, Bush 1652; Levasy, Bush 1684. Kansas: Manhattan; Hitchcock 2504, 2518, Pl. Kan. 571b. This species is listed in the Botany of Stevens’s Report @ as P. clandestinum, as shown by the specimen, without definite locality, in the National Herbarium. 175. Panicum xanthophysum A. Gray. Panicum xanthophysum A. Gray, Gram. & Cyp. 1: no. 28. 1834. This was pub- lished in the set of exsiccatae, a printed description in Latin accompanying no. 28, the locality being given as ‘‘Pine Plains, near Oneida Lake, New-York.’’ In the Gray Herbarium is a specimen, on the sheet of which is written in Dr. Gray’s hand ‘‘Oneida Lake, Wood Creek barrens, P. xanthophysum Gray!’’ This specimen, which we consider the type, is a single culm with an overmature primary panicle, and asmall secondary panicle. Dr. Gray apparently neglected to retain for himself a numbered set of this distribution of Graminee. Thespecimen of no. 28 in the Boott set in Gray Herbarium, and the other specimens of this number which we have seen agree with the above specimen. The species was later described by Gray® in English in a paper on ‘‘New or rare Plants of the State of New York.”’ Panicum xanthophysum forma amplifolium Scribn. in Brainerd, Jones & Eggleston, Fl. Vt. 104. 1900. ‘‘Dry sandy soil, Burlington, Vt. L. R. Jones, collector, August 31, 1893.’? The type, in the National Herbarium, consists of two specimens with overlapping sheaths and blades 1.5 to 2 cm. wide. aA. Gray Pac. R. Rep. 12: 49. 1860, bAnn, Lyc. N. Y. 3: 234, 1835. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 291 DESCRIPTION. Vernal form yellowish green, in loose tufts of few to many culms, 20 to 55 cm. high, erect or ascending, more or less scabrous; sheaths loose, about as long as the internodes or longer, sparsely papillose-pilose and ciliate at least toward the summit, rarely nearly glabrous; ligules 1 mm. long; blades erect or nearly so, rather thin, prominently nerved, 10 to 15 cm. long, rarely longer, 1 to 2 cm. wide, acuminate, slightly narrowed to the rounded papillose-ciliate base, otherwise glabrous, the uppermost blade not reduced, sometimes the largest; panicles finally long-exserted, 5to 12 cm. long, very narrow, sometimes appearing almost racemose, few-flowered, the stiff branches erect or nearly so; spikelets 3.7 to 4 mm. long, 2 to 2.1 mm. wide, obovate, turgid and blunt at maturity, pubescent; first glume about half as long as the spikelet, pointed; sec- ond glume scarcely equal- ing the fruit and _ sterile lemma at maturity; fruit 2.9 to 3 mm. long, 2 mm. wide, obovate-oval, mi- nutely umbonate. Autumnal form erect or Fic. 327.—P. zanthophysum. From type specimen. ascending, branching from the second and third nodes, the branches erect, mostly simple, the blades not reduced, usually equaling the short- exserted panicles, the large, erect blades making the plant appear leafy in the middle. The spikelets of the primary panicles sometimes perfect their grains. As stamens and stigmas are both frequently exserted, the fruitiul spikelets in this species are not always cleistogamous as is commonly the case in Dichanthelium. Three Minnesota specimens, Campbell 66, Ostland 1 and 2 in 1884, and an Ontario specimen, Macoun in 1865, seem to be intermediate between this speciesand P. leibergii. The first of these has the glabrous blades and strict panicles of P. zanthophysum, but the blades are only 5 to7 mm. wide. The Ostland specimens represent vernal and autumnal forms; both have blades that are ciliate for half their length or more but otherwise glabrous; the panicles are looser as in P. leibergii and the pubescence of the spikelets, which are too immature to indicate amount of turgidity, is papillose- spreading as in that species; the autumnal specimen has broader blades and would be referred to P. zanthophysum unhesitatingly but for the intermediate vernal specimen. DISTRIBUTION. Sandy or gravelly soil, Quebec to Minnesota, south to Pennsylvania. QueEBEc: Sorel, Pringle in 1879. Maine: Buckfield, Allen in 1877; Orono, Fernald 345; Madison, Fernald 521; South Berwick, Fernald 519; York, Fernald 520; Bangor, Knight in 1904; Cumberland, Knowlton & Chamberlain 500; East Auburn, Merrill 11; Ches- terville, Chase 3295, 3300. New Hampsurre: Laconia, Carter 101d. Vermont: Burlington, Jones in 1893; Peacham, Blanchard in 1889; Vernon, Grout in 1895. Massacuusetts: Ashburnham, Harris in 1896 (N. E. Bot. Club Herb.). Connecticut: Stafford, Graves in 1903 (Gray Herb.). 292 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. New Yor«: Oneida Lake, Gray; Lake George, Vasey in 1882; Schenectady, Wibbe in 1885; Danby, Coville in 1885; Cairo, Nash in 1893; Tripoli, Burnham in 1897. Ontario: Belleville, Macoun in 1865; Galt, Herriot in 1898 and 1901; Algonquin Park, Macown 22025. PENNSYLVANIA: Tannersville, Brown in 1901; without local- ity, McMinn. MicnicaAN: Keweenaw, Farwell 764; Alma, Davis in 1893. Wisconsin: Rainbow Rapids, Che- ney 1346; Mason, Cheney 4786; Webster, Cheney 3426. Minnesota: Ramsey County, Ost- land 1 and 2 in 1884; Lake Kilpatrick, Ballard in 1893; St. Cloud, Campbell 66. Manitosa: Lake Winnepeg Valley, Bourgeau in 1857 (Gray Herb.). SASKATCHAWAN: Bourgeau in 1858 (Gray Herb.). Fig. 328.—Distribution of P. zanthophysum. Pedicellata.—Clums slender, more or less hirsute; ligules of short hairs; blades not over 6 mm. wide, ciliate; spikelets 3.5 to 4 mm. long, attenuate at base, papillose, 7 to 9-nerved. Autumnal form freely branching, the branches appearing before the maturity of the primary panicle; no distinct winter rosette formed. This group of two species appears to be intermediate between the subgenus Dichanthelium and true Panicum. The plants bear a general resemblance to Oligosanthia but in the absence of a winter rosette and in the branch- ing habit, especially of P. nodatum, they show a departure from Dichanthelium. Culms erect or leaning; blades thin, 5 to 9 cm. long, narrowed LOWEN ERO bane serie ance Leaere nabs hee ee 176. P. pedicellatum. Culms decumbent; blades thick, not over 5 cm. long, not nar- rowed:toward the base. ....s:ces.s 6 Soe hehe ee 177. P. nodatum. 176. Panicum pedicellatum Vasey. Panicum pedicellatum Vasey, U.S. Dept. Agr. Div. Bot. Bull. 8: 28. 1889. ‘Texas, J. Reverchon.’’ The type specimen, in the National Herbarium, Reverchon Texas Plants 1620, bearing the data ‘‘Rocky woods, Kimble Co. June,”’ consists of two branching plants. DESCRIPTION. Vernal form in tufts of few to several erect or ascending culms from a short, knotted rootstock; culms slender, 20 to 50 ecm. high, usually ascending-hirsute at least below, a few spreading hairs on the nodes; sheaths papillose, sparingly hirsute, ciliate on the margin; ligules dense, about 1 mm. long; blades ascend- ing or spreading, 5 to 9 cm. long, 3 to 6 mm. wide, the margin toward the narrowed base sparsely ciliate with long hairs, both surfaces glabrous or sometimes minutely hispid; panicles 3 to 6 cm. long, Fic. 329.—P. pedicellatum. From type specimen. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 293 about three-fourths as wide, the branches few, spreading or ascending; spikelets 3.5 to 3.7 mm. long, 1.4 mm. wide, elliptic, prominently papillose-hispid; first glume nearly or quite half the length of the spikelet, narrow, acute; second glume shorter than the fruit and sterile lemma at maturity; fruit 3 mm. long, 1.3 mm. wide, elliptic, subacute. Autumnal form erect or leaning, branching from all but the uppermost nodes before the maturity of the primary panicle, the branches slightly divaricate, the blades and panicles not greatly re- duced. DISTRIBUTION. Dry woods and prairie, Texas. Texas: Kimble County, Reverchon 1620; Kerrville, Heller 1726, 1736, 1766, Smith in 1897; Aus- tin, Hall 8344 in part; Coman- che Spring, Lindheimer 1265 Fig. 330.—Distribution of P. pedicellatum. in Mo. Bot. Gard. distr; ‘‘in the Sabines bottom,’’ Lindheimer 158 (last two in Gray Herb.). 177. Panicum nodatum sp. noy. DESCRIPTION. Vernal form in tufts from a knotted crown; culms ascending or spreading, slender but hard and wiry, 25 to 35 cm. high, finely papillose, crisp-puberulent; sheaths shorter than the internodes, papillose-hispid between the strong nerves; ligules dense, scarcely 1 mm. long; blades firm, ascending, 3 to 5 cm. long, 3 to 6 mm. wide, broadest at the rounded base, abruptly acute, puberulent on both surfaces, papillose-ciliate with stiff hairs 2 to 3 mm. long; panicles 4 to 5 cm. long, half to two-thirds as wide, few-flowered, the few branches ascending; spike- lets 4 mm. long, 1.7 mm. wide, pyriform, papillose - pubescent; first glume about one-third the length of the spikelet, acuminate; second glume slightly shorter than the fruit and sterile lemma; fruit 3 mm. long, 1.4 mm. wide, obovate- elliptic, minutely white-puberu- lent at the apex. Autumnal form widely genicu- Fig. 331.—P. nodatum. From type specimen. late-decumbent, early branching from all but the uppermost node, the branches somewhat divaricate, equaling or exceeding the main culm, with numerous swollen nodes, the internodes 2 to 3 cm. long, the whole forming a loose tuft, the blades and panicles not reduced. Type U.S. National Herbarium no. 592749, collected August 7, 1904, Sarita, Texas, by A. 8S. Hitchcock (no. 3865). @ This in the Gray Herbarium is numbered 835. 294 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. This species differs from P. pedicellatum in its stiffer, short-jointed culms, shorter, puberulent, prominently ciliate blades, and pyriform spikelets with a shorter first glume. DISTRIBUTION. Oak woods in sand dunes, southern Texas and northern Mexico; appar- ently rare. Texas: Sarita, Hitchcock 3865. Mexico: Matamoros, Berlandier 988, 2418 (last two in Gray Herb.). F1a. 332.—Distribution of P. nodatum. Seoparia.—Vernal culms tall; blades flat, elongated, not over 1.5 em. wide; ligules short; spikelets pointed, 7 to 9-nerved. Pubescence soft-villous or velvety; spikelets abruptly pointed. Vernal culms erect or ascending; plants velvety through- out; spikelets about 2.5 mm. long.............-.-- 178. P. scoparium. Vernal culms decumbent at base; the upper sheaths more or less glabrate; spikelets less than 2 mm. long....- 179. P. viscidellum. Pubescence when present not velvety. Spikelets: elliptic; truit:2 aim. domigsr S222 cone eee 180. P. aculeatum. Spikelets ovate; that is, broadest below the middle; fruit 2 mm. long or less. Sheaths or some of them hispid, rarely glabrous; au- tumnal form with crowded branchitts. ---..--- 181. P. scabriusculum. Sheaths glabrous; autumnal form sparingly branching.182. P. cryptanthum. 178. Panicum scoparium Lam. Panicum scoparium Lam. Encycl. 4: 744.1798. ‘‘Cette plante a été recueillie dans la basse Caroline par le citoyen Michaux.’’ The type specimen, in the Lamarck Herbarium, is a part of a vernal culm with a single leaf and over-mature panicle. The accompanying label reads, ‘‘ Panicum scoparium Lam. dict. donné par le C. Michaux.” Panicum pubescens Lam. Encycl. 4: 748.1798. The author refers to two specimens as follows: ‘‘J’ai vu de cette espéce un individu nain dans l’herbier de Vaillant; il V’avoit regue de Sherard en 1721. Le citoyen Michaux 1’a trouvée dans la Basse- Caroline.’’ The first mentioned specimen is in the Paris Herbarium. It is some species of Brachiaria allied to Panicwm villoswum Lam. as described by Hooker. The other specimen referred to is in the Michaux Herbarium and consists of two freely branching autumnal culms of P. scopariwum. The accompanying label reads “‘ Pani- cum pubescens Lam. Hab. in pratis sylvestribus Carolinae.’’ Since Lamarck’s description applies to the Michaux? rather than to the Sherard specimen we may consider the Michaux plant the type. Panicum viscidum Ell. Bot. 8. C. & Ga. 1: 123. pl. 7. f. 3.¢ 1816. No specimen nor ‘definite locality is cited. The type in the Elliott Herbarium is labeled: ‘‘ Panicum viscidum mihi. Hab: in humidis.’’ It consists of a single culm lacking the base, bearing several short branches and an over-mature primary panicle. Panicum pauciflorum Bosc; Spreng. Syst. Veg. 1: 313. 1825. This, together with P. scoparium Michx., is placed as a synonym under 2’. pubescens Lam. We have seen no authentic specimen. : aF\. Brit. Ind. 7: 34. 1896. bSee Hitchcock, Contr. Nat. Herb. 12: 147. 1908. ¢ Plates 7 to 10 were issued with vol. 2. 1824. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 295 Panicum laxiflorum pubescens Chapm. Fl. South. U. 8. ed. 3. 586. 1897, not Vasey 1892. Based on Panicum pubescens Lam., though the description applies to P. strigo- sum Muhl., the species referred by Chapman to this variety. DESCRIPTION. Vernal plants grayish olive green, velvety-pubescent throughout except as noted; culms 80 to 130 cm. high, stout, erect or ascending, usually geniculate at base, the nodes villous with reflexed hairs, a glabrous, viscid ring below; sheaths about half as long as the long internodes, the velvety pubescence wanting on the back toward the summit, the surface here viscid when fresh; ligules 1 mm. long; blades rather thick, ascending or spreading, often reflexed late in the season, 12 to 20 cm. long, 10 to 18 mm. wide, long-acuminate, slightly narrowed to the rounded base, the uppermost leaf often much reduced; panicles finally long-exserted, 8 to 15 cm. long, nearly as wide, many- flowered, the axis, branches, and pedicels with viscid blotches, the branches ascend- ing or spreading, spikelet-bearing to the base; spikelets 2.4 to 2.6 mm. long, 1.4 to 1.5 mm. wide, obovate, turgid at maturity, abruptly pointed, papillose-pubescent with spreading hairs; first glume one-fifth to one-fourth the length of the spikelet, acute to truncate; second glume and sterile lemma strongly nerved, the glume obtuse, shorter than the fruit at maturity, the lemma abruptly pointed and equaling it; fruit 2 mm. long, 1.4 mm. wide, obovate-elliptic, apiculate. ' Autumnal form leaning or spreading, Fig. 333.—P. scoparium. From type specimen of | branching from the middle nodes after P. viscidum Ell. the maturity of the primary panicle, the branches usually longer than the primary internodes, repeatedly branching, often more or less scorpioid, the ultimate branchlets in flabellate fascicles, the sheaths often swollen toward the summit, con- tracted at the throat, the blades much reduced, overtopping the small, partially included panicles. A well-marked and constant species, easily recognized by its velvety pubescence, the glabrous, viscid ring below the nodes, and the viscid upper portion of the sheath. The viscidity disappears in drying, but the glandular surface is evident. DISTRIBUTION. Wet or damp soil, Massachusetts to Florida, west to Oklahoma and Texas; also in Cuba. Massacuusetts: Cape Cod, Cheney in 1903 (N. E. Bot. Club Herb.). New Jersey: Avon, Mackenzie 1854; Tuckerton, Chase 3600; Wildwood, Chase 3486. PENNSYLVANIA: Tinicum, Smith 159; without locality, Mc Minn. DetawareE: Millsboro, Commons 28; Ellendale, Commons 32. MARYLAND: Eastern Shore, Canby; Anne Arundel County, J. D. Smith in 1879; Chesapeake Junction, Hitchcock 1637. District or CotumBiA: Kearney in 1897, Merrill 233, Sheldon in 1881, Steele in 1896 and 1897, Topping in 1895, Vasey in 1881, Ward in 1878 and 1879. Virani: In the vicinity of Cape Henry, Chase 5438, Coville 17, Hitchcock 593, Kearney 308, 1477, Mackenzie 1688, Noyes 88, 89; Dismal Swamp, McCarthy in 1883. Nortn Carouina: Wilmington, Biltmore Herb. 4290; Hickory, Small & Heller in 1891; Heiligs Mill, Small & Heller 204; West Raleigh, Coit 1304. 296 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. SoutH Carouina: Orangeburg, Hitchcock 437. Groraia: Savannah, Kearney 183; McGuires Mill, Small in 1893; Clarke County, Harper 110, 150; Cobb County, Harper 210; Americus, Tracy in 1897; Dekalb County, Eggert 82; Stone Mountain, Hitchcock 215; Thomson, Bartlett 1170. Fioriwa: Jacksonville, Kearney 156; Lake City, Nash 2204; Apalachicola, Biltmore Herb. 4290a; without locality, Chap- man. Kentucky: Warren County, Price in 1896 (Mo. Bot. Gard. Herb.). } TennesseE: Chester County, Bain Fig. 334.—Distribution of P. scoparium. in 1892. Y AxtaBama: Auburn, Pollard & Mazon 2, 67, Tracy 3978; Cullman, Eggert 12, 60, Mohr in 1895; Selma, Kearney 3; Tuskegee, Carver 52, 87; Mobile, Hitchcock 594, Kearney 67, Tracy 7048. Mississippi: Starkville, Tracy in 1896; Taylorville, Tracy 8591; Pachuta, Tracy 3306. ARKANSAS: Miller County, Eggert 116, Heller 4236; northwest Arkansas, Harvey30. LoursiAna: Arcadia, Ball 77; Ruston, Cocks 3324. Texas: Waller County, Thurow in 1898; Texarkana, Plank 25; Hempstead, Halt 829; Fort Smith to the Rio Grande, Bigelow; without locality, Nealley in 1885, Drummond 381; Vincent 41b (Mo. Bot. Gard. Herb.). Ox.LAHomA: Choctaw Agency, Bigelow. CusBa: Road to Pinal Mayari, Wright 3467 (Gray Herb.). 179. Panicum viscidellum Scribn. Panicum viscidellum Scribn. U. 8S. Dept. Agr. Div. Agrost. Cire. 19: 2. 1900. ‘‘Gravelly banks near Jalapa, State of Vera Cruz, altitude 1,250 m. (4,000 feet). C.G. Pringle, No. 8089. 1899.’’ A second specimen, Liebmann 323, is cited but the first is taken as the type. The Pringle specimen, which is in the National Herbarium, is in the early branching state. The culm appears to be decumbent or creeping, sending up erect branches. DESCRIPTION. Vernal culms ascending from a decumbent, widely spreading or creeping base, rooting at the lower nodes, softly villous or nearly glabrous, the nodes more or less short-bearded; sheaths shorter than the internodes, villous or, especially the upper- most, glabrate; ligules ciliate, 2 to 4 mm. long; blades spreading, rather thick, more or less velvety-pubescent or villous on both surfaces, the uppermost 5 to 13 cm. long, 9 to 13 mm. wide, linear-lanceolate, subcordate, the lower lanceolate, more cordate, shorter and wider; panicles 4 to 11 cm. long, half to two-thirds as wide, usually densely flowered, the numerous branches Fic. 335.—P. viscidellum. From ; : ‘ type specimen. spikelet-bearing from the base; spikelets 1.8 to 1.9mm. long, 1 mm. wide, elliptic, acute, sparsely pubescent; first glume about one-third as long as the spikelet, pointed; second glume and sterile lemma slightly exceeding the fruit at maturity; fruit 1.5 mm. long, 0.8 mm. wide, elliptic, abruptly pointed. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 297 Autumnal form branching from the middle and upper nodes, the branches erect or ascending from the decumbent primary culm, the densely villous sheaths often over- lapping on the shortened internodes, the blades more or less ovate-lanceolate and strongly cordate-clasping, velvety pubescent, the panicles similar to the primary ones but smaller. The autumnal form is not well developed in any of the specimens examined. Judg- ing from the habit of the plant it appears to belong to the subgenus Dichanthelium but more material is necessary to determine this with certainty. The characters of the vernal form would place it in the group Lanuginosa, but the method of branching is different from that of any of the species there segregated. The specimens often resemble P. scoparium, and the species is therefore placed in this group, though somewhat doubtfully because of the manifest ligules and the branching habit. DISTRIBUTION. Gravelly banks, fields and open forests, Mexico to Colombia; also in the Isle of Pines. Mexico: Jalapa, Pringle 8089, C. L. Smith 1617; Zacualpan, Purpus 2160;4 Mirador, Liebmann 323; San Cristobal, Bowrgeaw 3132 (Paris Herb.); Jical- tepec, Liebmann 324 (Paris Herb.); Orizaba, Botteri 705 (Gray Herb.); Schaffner 284 (Paris Herb.). GUATEMALA: Sierra del Mico, Kellerman 6231, 6249. Nicaracua: U.S. Pacific Expl. Exped. Wright. Costa Rica: Aserri, Tonduz 1244; El General, Pittier 3358 (Instit. Costaric. Herb.). Cusa: Isle of Pines, Palmer & Riley 1065. CotomsiA: Near Jamund{, Pittier 940; Popaydn, Lehmann 985 (Gray Herb.). 180. Panicum aculeatum Hitchc. & Chase. Panicum aculeatum Hitche. & Chase, Rhodora 8: 209. 1906. ‘‘Type Chase 2520 in National Herbarium. In large clumps by small slough, border of woods, Takoma Park, D. C., July 27, 1904; collected by Agnes Chase.’’ The type isa vernal specimen beginning to branch, with a mature primary panicle. DESCRIPTION. Vernal plants in large clumps; culms slender, 70 cm. to 1 meter high, ascending, scabrous, harshly pubescent below; sheaths papillose-hispid with stiff, sharp-pointed hairs, a puberulent ring at the summit, the uppermost usually glabrous; ligules minute, membranaceous, ciliate; blades firm, stiffly ascending or spreading, 12 to 20 cm. long, 9 to 13mm. wide, acuminate, involute-pointed, scarcely narrowed to the rounded base, very scabrous on the upper surface and toward the apex beneath; panicles 8 to 12 cm. long, about as wide, few-flowered, the slender, flexuous, fascicled branches ascending or spreading, naked at the base, scabrous, some- times with a few viscid spots; spikelets 3 mm. long, elliptic, minutely pubescent; first glume one-fourth to one-third as long as the spikelet, acute; second glume and sterile lemma abruptly acute, slightly exceeding the fruit; fruit elliptic, 2.7 mm. long, 1.3 mm. wide, minutely umbonate. Fic. 336.—P.aculeatum. From type specimen. a Panicum larum Swartz was also distributed under this number. 298 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Autumnal form branching from the middle nodes, the branches more or less divari- cate, not much crowded, the blades not greatly reduced, the ultimate panicles wholly or partially included in the sheaths. This species is allied to P. scabriusculum, but the panicles are smaller, more open, fewer-flowered, the axis and branches scarcely viscid, the spikelets larger and the glumes only slightly exceed- ing the fruit. Vernal plants bear a . superficial resemblance to P. clandes- ‘ey uit bm tinum. See Pewina ai DISTRIBUTION. Swampy woods, Long Island, New York, District of Columbia, and North Carolina; rare. New Yorx: Rockville Center, Bicknell in 1903; Hempstead, Bicknell in 1906. District oF CoLuMBIA: Chase 2520, 5439, House 1041 (Hitchcock Herb.). Nort Carona: Lake Mattamuskeet, Chase 3210. Fig. 337.—Distribution of P. aculeatum. 181. Panicum scabriusculum Ell. Panicum scabriusculum Ell. Bot. 8. C. & Ga. 1: 121.1816. “Sent to me from Savannah by Dr. Baldwin.’”’ The type, in the Elliott Herbarium, consists of the upper part of a culm with one leaf and a large over-mature panicle. The accompany- ing label reads: ‘‘Panicum scabriusculum. Hab: Georg: Dr. Baldwin.”’ Panicum lanuginosum Bosc; Spreng. Syst. Veg. 1: 319. 1825, not Ell. 1816. ““Georgia.’’ The type is in the Willdenow Herbarium. Panicum eriophorum Schult.;¢ Kunth, Enum. Pl. 1: 128.1833. Based on P. lanuginosum Bosc, the description of which is copied. Panicum nealleyi Vasey, Bull. Torrey Club 138: 25. 1886. ‘‘Collected in Texas by Mr. G. ©. Nealley.’’ The type, in the National Herbarium, is a vernal specimen. It was collected in Texas, May, 1885, the exact locality not being given. Panicum dichotomum elatum Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 31. 1889. No locality nor specimen is mentioned by Vasey. The only specimen in the National Herbarium bearing this name in Vasey’s writing is one of P. scabriusculum in the early branching state. This specimen agrees with Vasey’s description and is taken as the type. It was collected by Charles Mohr in Mobile County, Alabama, June 18, 1888. DESCRIPTION. Vernal form grayish olive green; culms erect, 1 to 1.5 meters high, scabrous at least below the nodes, sometimes puberulent, the nodes glabrous or puberulent; sheaths shorter than the internodes, glabrous to more or less hispid at least toward the sum- mit, often mottled or white-spotted, commonly swollen at the base and contracted toward the summit; ligules short-membranaceous, usually with a ring of hairs above; blades stiffly ascending or spreading, often reflexed, 15 to 25 cm. long, 9 to 12 or rarely 15 mm. wide, glabrous or scabrous, often more or less pubescent beneath, gradually tapering to an involute point, slightly narrowed toward the base; panicles finally exserted, 10 to 20 cm. long, half to two-thirds as wide, rarely wider, many-flowered, the a Kunth cites ‘“‘ Schult. Mant. 3. 591’’ as place of publication, but neither the second nor third volume of Schultes’s Mantissa contains a page 591, nor is this name in either volume. HITCHCOCK AND CHASE-——-NORTH AMERICAN PANICUM. 299 axis glabrous or pubescent, often viscid, the flexuous branches ascending, spikelet- bearing from near the base; spikelets 2.3 to 2.6 mm. long, 1.1 to 1.3 mm. wide, ovate, pointed, glabrous or obscurely puberulent; first glume less than one-sixth as long as the spikelet; second glume and sterile lemma strongly nerved, exceeding the fruit and form- ing an abrupt point beyond it; fruit 1.8 mm. long, 1 mm. wide, elliptic. Autumnal form erect, branching from the mid- dle and upper nodes, the branches appressed, somewhat longer than the internodes, finally bearing fascicled branchlets and forming dense oblong masses along the upper part of the primary culm, the sheaths, especially the later ones, densely papillose-hirsute, the flat, reduced blades ovate-lanceolate, reduced in length much more than in width, the panicles partly or entirely inclosed in the sheaths. This species is very variable in the amount of pubescence; even on the same plant are often found glabrous and hispid sheaths or glabrous and pubescent blades. Other- wise it is an unusually uniform species. Fig. 338.—P. scabriusculum. From type specimen. DISTRIBUTION. Moist ground, especially along ditches, streams, and swamps, near the coast, south- east Virginia to Florida and eastern Texas. New Jersey: Atlantic City, Long in 1909 (Phila. Acad. Herb.). Vireita: Norfolk County, Kearney 1798; Dismal Swamp, Tyler in 1905. Norra Caroutna: Roanoke Is- land, Chase 3235; Wilsons Mills, Chase 3101; Wilmington, Chase 4600, Hitchcock 595, Kearney 270. South Carortna: Orangeburg, Hitchcock 438, 1378; Aiken, Ravenel. Georeia: Bullock County, Harper 881; Leslie, Harper 410. Frioria: Jacksonville, Curtiss Q, 4878; Duval County, Curtiss 3610; Baldwin, Combs 67; Washington, Combs 616; without locality, Chapman. ALABAMA: Flomaton, Hitchcock 1052, Tracy 3643; Mobile, Kearney 27, 39; Mobile County, Mohr in 1888. Mississippi: Beauvoir, Tracy 4617; Biloxi, Tracy 4569. Loutstana: New Orleans, Drummond (Gray Herb.). Texas: Nona, Nealley 38 in 1892; without locality, Nealley in 1885, Wright (Gray Herb.). Fig. 339.—Distribution of P. scabriusculum. 182. Panicum cryptanthum Ashe. Panicum cryptanthum Ashe, N. C. Agr. Exp. Sta. Bull.175: 115.1900. ‘‘Collected by the writer in swamps at Wilson’s Mill, N. C., in July 1897.’’ The type, in Ashe’s herbarium, is a specimen arbitrarily chosen from among four bearing the label, ‘‘ Wil- son’s Mill, N. C. July 15, 1897. W. W. Ashe collector,’’ and with the additional data, “‘In a small swamp on north side of railroad about one mile west of the station.”’ The name does not appear upon any of the sheets, but these plants agree with the 800 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, description and are from the locality as published. These specimens all are the autumnal form, with the reduced panicles partially inclosed in the sheaths. An earlier Panicum cryptanthum “Nuttall. Gen.’’ isa nomen nudum. It is men- tioned without description under Panicum cynodon Reichardt by Hillebrand 4 as a name in Kew Herbarium. DESCRIPTION. Vernal form cespitose; culms erect, 80 to 100 cm. high, glabrous except the usually bearded nodes; sheaths glabrous or the lowermost sparsely hirsute, the upper somewhat in- flated, all more or less ciliate on the margins and pilose at the summit; ligules membranaceous, erose, scarcely 0.5mm. long; blades stiff, ascending or spreading, acuminate, invo- lute-pointed, glabrous, sparingly ciliate at base, 10 to 15 cm. long, 7 to 9 mm. wide; panicles short-exserted, 6 to 10 cm. long, nearly as wide, the axis,and ascending branches viscid-spotted; spikelets 2.2 to 2.4 mm. long, 1 mm. wide, lanceolate-elliptic, pointed; first glume one-fourth to one-third as long as the spikelet; second glume and sterile lemma equal, longer than the fruit and pointed beyond it, glabrous or sparsely pilose; fruit 1.5 mm. long, 0.9 mm. wide, elliptic. Autumnal form erect, glabrate on the nodes, sparingly branching from the middle and upper nodes, the branches stiffly ascending at an angle of 30 to 45 degrees; blades flat, stiffly ascending, 2 to 5 cm. long, 3 to 5 mm. wide, involute-pointed; panicles reduced to a narrow cluster partially hidden in the sheaths. The habit of this species suggests a small P. scabriusculum. Fic. 340.—P. cryptanthum. From type specimen. DISTRIBUTION. Low swampy ground, North Carolina tonorthern Florida, also in Texas; rare. New Jersey: Atlantic City, Long in 1909 (Phila. Acad. Herb.). Nortu Carouina: Wilsons Mills, Ashe in 1897; Wilmington, Hitchcock 371, 1469. Soutu Carona: Orangeburg, Httchcock 1377. GeroraiA: Belair, Zggert in 1899 (Mo. Bot. Gard. Herb.). Frioripa: De Funiak Springs, Combs 444. Texas: Kountze, Nealley 37 in 1892. Fig. 341.—Distribution of P, cryptanthum. Commutata.—Culms rather stout; glabrous or puberulent; ligules obsolete or nearly so; blades usually 1 cm. or more wide (often narrower in P. ashei and P. equilaterale), cordate and more or less ciliate at base; spikelets 2.4 to 3.2 mm. long, elliptic, not very turgid, pubescent, 7 to 9-nerved. Autumnal form usually not very freely branching. Plants glaucous, glabrous; basal blades conspicuously ciliate; vernal, culms usually solitary..2. 02.5... 0-2-cee=snee 185. P. mutabile. Plants not glaucous. Blades nearly linear, that is with parallel margins; first glume about half as long as the spikelet. Primary panicles long-exserted; sheaths mottled Wan WUILO «. on aden ite anaes bm Mee 188. P. albomaculatum. aFl. Hawaiian Isl. 498. 1888. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 301 Primary panicles short-exserted; sheaths not mot- Oe Ae Ae ee ae 187. P. equilaterale. Blades lanceolate. Culms crisp-puberulent; blades usually rigid, sym- metrical, rarely over 10 mm. wide; spikelets ETE ee 183. P. ashei. Culms glabrous or softly puberulent; blades firm or lax; spikelets 2.7 to 3.2 mm. long. Culms erect, or autumnal form leaning; blades symmetrical, broadly cordate...........-- 184. P. commutatum. Culms decumbent; blades usually unsymmetrical and falcate, narrowed to the scarcely cor- eee oO Bee fe are? ee ae 186. P. joorii. 183. Panicum ashei Pearson. Panicum umbrosum LeConte in Torr. Cat. Pl. N. Y. 91. 1819, not Retz. 1786. On page 19 of this work the locality is given as, ‘“‘In woods, Bloomingdale, N. Y.”’ The type, in the Torrey Herbarium, consists of two vernal culms. On the same sheet is mounted a piece of a culm of P.dichotomum. On the label is written ‘‘ Panicum umbrosum mihi” in LeConte’s hand, to which is added in Torrey’s hand, ‘‘(Le Conte).’’ The description as given by LeConte does not apply to this specimen in all respects. The culm and flowers are said to be glabrous, which is true of the speci- men of P. dichotomum, but the other characters apply better to the two culms of P. ashei. Panicum ashei Pearson; Ashe, Journ. Elisha Mitchell Soc. 15:35. 1898. ‘‘P. com- mutatum Schultes var. minor Vasey, Contrib. from U. 8. Nat. Herb., vol. 3, No. I: 32 (1892). Not P. capillare var. minus Muhl. (1817).’’ It would appear from this cita- tion that P. ashei was intended as a change of name, but “‘sp. nov.”’ follows the author’s name, preceding the above citation, and a description is given. Hence the synonym @ may be regarded as an error and the first specimen cited, ‘‘New York: Ashe; Ithaca, July 1898,’’ may be taken as the type. This specimen could not be found in Ashe’s herbarium, but there is a duplicate in the National Herbarium sent by Ashe, and labeled in his writing ‘“‘ Panicum Ashei, G. Pearson, Dry woods, Ithaca, N. Y. W. W. Ashe, July 1898.’’ This consists of two vernal plants beginning to branch, with mature primary panicles; the culms and blades are less rigid than usual for this species. In the description the culms and sheaths are said to be glabrous, but in this specimen the culms and sheaths are crisp-puberulent. DESCRIPTION. Vernal plants usually conspicuously purplish, in loose clumps of few to several culms from a knotted crown; culms 25 to 50 cm. high, erect, stiff and wiry, densely crisp- puberulent, including the nodes; sheaths shorter than the internodes, less densely puberulent, short-ciliate; ligules obsolete; blades usually thick and firm, spreading or ascending, 4 to 8 cm. long, 5 to 10 mm. wide (the lower gradually smaller), acumi- nate, ciliate at the subcordate base and sometimes along the very scabrous margin, glabrous on both surfaces; panicles finally long-exserted, 5 to 8 cm. long, hardly as wide, loosely flowered, the branches usually in distant fascicles, ascending or spread- ing; spikelets 2.4 to 2.7 mm. long, 1.2 to 1.3 mm. wide, oblong-elliptic, obtuse or obscurely pointed, short-pubescent; first glume about one-third the length of the spikelet, subacute; second glume and sterile lemma subequal, slightly exposing the fruit at maturity, obtuse or withering to a point; fruit 2.1 mm. long, 1.1 mm. wide, elliptic, minutely umbonate. a See synonymy under P. commutatum, page 304. 302 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Autumnal form erect or topheavy-reclining, the culms bearing divergent branches from the middle and upper nodes or from the upper only, the terminal joint of the primary culm commonly falling, the sheaths crowded or overlapping, the blades rigid and widely spreading, little reduced except those of late autumn; winter rosette appearing early, the blades thick and firm, usu- ally prominently ciliate nearly to the apex. This unusually uniform species is distinguished from P. commutatum by the rigid habit, thicker, narrower blades, crisp-puberulent culms and sheaths, and by the more freely branching, often topheavy autumnal form. Occasional speci- mens, such as Bissell 5580 and Andrews 63 have spikelets only 2.1 to 2.3 mm. long. Fic. 342.—P. ashei. From duplicatetype There is a form represented by several speci- specimen in National Herbarium. mens which appears to be intermediate between P. ashei and PF. barbulatum. The plants grow in large clumps of numerous stiff culms, with narrow blades ciliate at the base, and with spikelets about 2 mm. long. These specimens are as follows. New Yorx: Law- ence, Bicknell in 1892; NewJersey: Tuckerton, Chase 36014; District or COLUMBIA: Hitchcock 503; SoutH Caroumna: Orangeburg, Hitchcock 1406, 1407. DISTRIBUTION. Dry, especially rocky woods, Massachusetts to northern Florida, west to Michigan, Missouri, and Mississippi. MassacuvuseEtts: Malden, Fernald in 1891 in part; West Quincy, Churchill in 1894 (both in Hitchcock Herb.). Connecticut: Montville, Graves 88; New London, Andrews 63; North Stoning- ton, Bissell 5580. RuHopE IsLanpD: Providence, Olney. New York: Ithaca, Ashe in 1898; Northville, Bicknell in 1903 and 1904, Young 4; Rockdale Center, Bicknell in 1906; Rosedale, Bicknell in 1904. New Jersey: South Amboy, Mackenzie 2163; Sussex County, Mackenzie 2195; Wildwood, Chase 3502; Wildwood Junction, Chase 3521; Atsion, Chase 3542; Egg Harbor, Martindale in 1876; Oradell, Mackenzie 2480. PENNSYLVANIA: Easton, Porter in 1895, 1897, and 1898; Cham- bersburg, Porter in 1898; Lan- caster County, Heller 4779 in part, 4780. Ouro: Niles, Ingraham in 1892; Painesville, Hacker 6878; Sugar Grove, Kellerman 6892. Inp1ANA: Clarke County, Deam 6467, 6905. Fic. 343,—Distribution of P. ashei. MicuicaAN: Muskegan, Wheeler 19. Missouri: Swan, Bush 17, 2911, 3456, 4487; Pleasant Grove, Bush 289, 312; Monteer, Bush 4714; Chadwick, Bush 4412. DeLtawareE: Wilmington, Commons 61; Milton, Commons 349, 356, 357; Green- bank, Commons 37; Ellendale, Commons 308; Frankford, Commons 53. MaryLaNnp: Patuxent, House 961, Hitchcock 1640; Beltsville, Chase 3787; Hyatts- ville, Steele in 1903; Woodside, Chase 2830; West Chevy Chase, Chase 2477; Plummers Island, Hitchcock 564; Chesapeake Junction, Hitchcock 2410. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 3803 District or CotumBIA: Chase in Kneucker Gram. Exs. 556, Hitchcock 422, 425, 568, Kearney in 1897, Ward 2, 7,9, Williams 3, 4, 10. VireiniA: Great Falls, Chase 3704, Four-Mile Run, Pollard 180; Norfolk, Kearney 299, 1029; Portsmouth, Noyes 103; Virginia Beach, Pollard & Maxon in 1900; Munden, Mackenzie 1708; Dismal Swamp, Chase 3679, Tyler in 1905; Clifton Forge, Tidestrom 4. West VirerniA: Summers County, Morris 977; Tibbs Run, Sheldon 566; Fayette County, Kellerman 6901. Nortu Carona: Roanoke Island, Chase 3220; Wilmington, Hitchcock 424, 1461; Chapel Hill, Ashe, Chuse 3051, 3064; Magnetic City, Wetherby 58; Lenoir, Hitchcock 565. Sout Carona: Orangeburg, Hitchcock 1388; Pelzer, House 2430; Clemson College, House 2105; Aiken, Hitchcock 566. Georcia: Augusta, Cuthbert 427, 1163, Kearney 207; Stone Mountain, Eggert 44, Hitchcock 423, 567, 1353; Thomson, Bartlett 1458, 1504. Fiorina: Chattahoochee, Tracy 3629 (Field Mus. Herb.). Kentucky: Harlan County, Kearney 34 in part, 54 in part. TENNESSEE: Polk County, Chambliss 14, 88, Kearney 324; Knoxville, Smith in 1895, Scribner in 1891; Nashville, Gattinger in 1882. AtaBAMA: Pisgah, Chase 4473; Scottsboro, Chase 4498; Auburn, Hitchcock 1323, 1327, Tracy 3747, 3756; Tuskegee, Ball in 1901. Mississreri: Enterprise, Tracy 3275; Meridian, Tracy 3268. 184. Panicum commutatum Schult. Panicum nitidum majus Pursh, Fl. Amer. Sept. 1: 67. 1814. No specimen nor locality is cited. Among the Pursh plants in Kew Herbarium is a sheet upon which are mounted a specimen of P. commutatum and one of P. yadkinense. Above the first is a label reading “‘ Panicum nitidum Mx.,”’ and above the latter one reading ‘‘Panicum nitidum major.’’ Since Pursh’s description of the variety is “‘omnibus partibus multo majus,” it would appear that the labels have been transposed, and that the specimen of P. commutatum should be taken as the type of variety majus. Panicum nervosum Muhl.; Ell. Bot. 8S. C. & Ga. 1: 122. 1816, not Lam. 1797. Elliott states that the species ‘‘grows in dry shaded soils,’’ but no definite locality is given. The type specimen, in the Elliott Herbarium, consists of portions of two culms, the panicle of one destitute of spikelets. The culms are glabrous, the sheaths glabrous or minutely puberulent between the nerves, densely puberulent on the back at the summit, blades papillose-ciliate at base, otherwise glabrous. The accom- panying label reads: ‘‘Panicum nervosum Muhl. Hab. Car. et Geor:’’ This species was also described as P. nervosum by Muhlenberg,¢ but there is no specimen to rep- resent this in the Muhlenberg Herbarium. In this description the ligule is said to be “‘barbaeform,”’ and the species ‘‘P. aquatico? affine.’’ This throws some doubt on the identity of Muhlenberg’s specimen with the type of P. nervosum Muhl.; Ell. in Elliott’s Herbarium. But the statement that the leaves are lanceolate and ciliate at base, and that the panicle branches are numerous and spreading shows that the species has no close affinity to his P. aguaticum, as suggested by Muhlenberg. Since no specimen can be found by which to interpret Muhlenberg’s description, it is here assumed that his type is the same as Elliott’s and that the statement concerning the ligule is an error. Panicum commutatum Schult. Mant. 2: 242. 1824. Based on ‘“‘P. nervosum Miihlenb. Descr. ub. p. 117”’ [error for 116], the name presumably changed because of @ Descr. Gram. 116. 1817. b This is Sacciolepis striata (L.) Nash. 304 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. P. nervosum Lam. 1797. Muhlenberg’s description, including ‘‘ ligula barbiformis, ”’ iscopied. Itis evident that Schultes had not seen a specimen of this.4 Panicum enslini Trin. Gram. Pan. 230. 1826. ‘‘Am.[erica] bor.[ealis] (TratrTiIn- 1cK, e collect. Enslini).’’ The type, in the Trinius Herbarium, is the vernal form. The accompanying label in Trinius’s hand reads “‘Panicum Enslini m. (An Pan. tenue Muhl. quaerit Nees ab Es.) ab Enslino in Am. bor. 1. dt. sine nom. cl. Trattin- ick Wiennae 1820.’’ There is a duplicate specimen in the Vienna Herbarium. Judg- ing from a small portion of the type sent to the National Herbarium from the Trinius Herbarium? this was thought to be the same as P. equilaterale, but from subsequent study of the somewhat fragmentary entire specimen and of the duplicate it appears to be a narrow-leaved form of P. commutatum. Panicum polyneuron Steud. Syn. Pl. Glum. 1: 91. 1854. Based on ‘‘P. nervosum Miihlbrg. Gram. p. 116, Torr. Fl. N. Am. I. 143.’’ The description, however, is a translation of that given in Torrey’s Flora¢ for P. nervosum and applies to the species described as P. macrocarpon LeConte,@4 which proves to be the true P. latifolium L. On the type sheet of P. macrocarpon LeConte, in Columbia University Herbarium, is written in Torrey’s hand ‘‘(P. nervosum Muhl. T.).’’ [T. probably stands for Torrey, that is, P. nervosum Muhl. according to Torrey.] Panicum commutatum minor[us] Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 34. 1889. ‘‘Southern States.’’ The only specimen marked with this name in Vasey’s writing is found in Nash’s herbarium, having been obtained through exchange of duplicates with Dr. Vasey. This is a small, immature, vernal specimen with glab- rous culms, collected by Ravenel at Aiken, S. C., in 1867, and is labeled in Vasey’s hand ‘“‘P. commutatum var. minor Vasey.”’ Panicum commutatum latifolium Scribn. in Kearney, Bull. Torrey Club 20: 476. 1893. This is described in a single line in a footnote to notes on plants collected on Pine Mountain, southeastern Kentucky, in August, 1893, as follows: ‘‘ Leaves very broad, panicle large, the widely spreading branches few-flowered.’’ On page 479 this name is listed as Kearney no. 299. In the Scribner Herbarium is a specimen of P. commutatum with unusually wide biades and large panicles, which is probably the plant referred to, although there are no data with the specimen.é The sheet is marked. P. latifolium Ell., through which is drawn a line and below in Scribner’s hand is written ‘‘Panicum commutatum Schultes.”’ Panicum commelinaefolium Ashe, Journ. Elisha Mitchell Soc. 15: 29. 1898, not Rudge, 1805. ‘‘Based on material collected by Dr. J. K. Small near Stone Mt., Ga., Aug. 1-6, 1895.’’ The type, in the Biltmore Herbarium, consists of several late ver- nal culms beginning to branch, with short internodes and short-exserted panicles. The culms, sheaths, and both surfaces of the blades are puberulent, the blades 1.7 to 2.5 em. wide, the spikelets 2.8 mm. long. Panicum currani Ashe, Journ. Elisha Mitchell Soc. 15: 113. 1899. Based on P. commelinaefolium Ashe, not Kunth. Panicum subsimplex Ashe, N. C. Agr. Exp. Sta. Bull. 175: 115. 1900. ‘‘Col- lected by Mr. A. Commons in dry rocky woods near Wilmington, Del., in August.”’ The type, in Ashe’s herbarium, consists of autumnal culms, with broadly elliptic, somewhat falcate blades 1.5 to 2.5 cm. wide, and small, few-flowered panicles, over- topped by the upper blades. 4 Schultes seems not to have had Elliott’s Botany of South Carolina and Georgia, since where he refers to this work (Mant. 2: 256-257) he does so on the authority of Nuttall’s Genera Plantarum. b See Hitchcock, Bot. Gaz. 41: 65. 1906. ¢Fl. North. & Mid. U. S. 148. 1823. @LeConte in Torr. Cat. Pl. N. Y. 91. 1819. e Other Kearney specimens in the Scribner Herbarium are often marked with collector’s name and number only, without other data. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 305 DESCRIPTION. Vernal plants commonly purple-tinged; culms in clumps of few to many, 40 to 75 em. high, rather stout, erect, glabrous or sometimes softly (not crisp) puberulent; nodes puberulent; sheaths shorter than the long internodes, ciliate on the margin and with a densely puberulent ring at the summit, otherwise glabrous or puberulent between the nerves; ligules nearly obsolete; blades usually firm, spreading or ascend- ing, 5 to 12 cm. long, 12 to 25 mm. wide, the lower and upper smaller than those of the midculm, rather abruptly tapering to an acuminate apex and slightly narrowed to the cordate-clasping base, glabrous on both surfaces or puberulent beneath or sometimes also above, the margin ciliate at the base; panicles usually long-exsérted, 6 to 12 cm. long, as wide or wider, loosely flowered, the axis glabrous or nearly so, the flexuous branches spreading; spikelets 2.6 to 2.8 mm. long, 1.3 mm. wide, oblong- elliptic, obtuse, softly pubescent; first glume about one-fourth the length of the spike- let, triangular, acute or obtuse; second glume and sterile lemma barely covering the fruit at maturity; fruit 2.2 to 2.3 mm. long, 1.2 mm. wide, elliptic, minutely umbonate. Autumnal form erect or leaning, branching from the middle nodes, the portion of the primary culm above the uppermost branch commonly falling away, leaving the branch, with its shortened internodes, crowded, rather loose sheaths, scarcely or not at all reduced blades, and hardly exserted panicle, as the apparent termination of the primary culm; secondary branchlets crowded toward the summit, the reduced blades exceeding the partly included, much reduced panicles; winter rosette appearing rather early, the blades firm, ovate. This species is typically almost glabrous, with stiff culms and firm blades, but puberu- lence occursrather commonly and is not found to be associated with any other character. The type of P. currani is puberulent through- out and has somewhat broader blades than common in P. commutatum, but these charac- ters are too variable to allow of separating this Fic. 344—P. commutatum. From type speci- form as a species. In some specimens the men of P. nervosum Muhl. in Elliott Her- : barium. culms only are puberulent, in others the sheaths or the lower suriace of the blades only. The puberulence can not be codrdinated with the wide blades. Some puberulent specimens have ordinarily wide blades and other specimens with wide blades are glabrous. : Early autumnal specimens in which the upper branch has replaced the terminal portion of the main culm sometimes appear very different from vernal specimens, owing to a somewhat unsymmetrical broadening of the middle of the crowded upper blades. The typeof P. subsimplex issuchaspecimen. A plant collected by Scribner at White Cliff Springs, Tenn. (in Hitchcock’s herbarium), shows several culms of typical P. commutatum, the terminal portions widely divaricate, but not yet fallen, and the upright branches with the unsymmetrically broadened blades as in the type of P. subsimpler. A few southwestern specimens such as Hitchcock 1104, 1253, Langlois 39, and 41 in part, Nealley in 1887 and Tracy 4577, differ in appearance from P. commutatum, having rather slender culms and narrower blades and seem to approach P. joorii, but the spikelets are not over 2.8 mm. long. Most of the Florida specimens are taller and more robust and have blades sometimes as much as 20 cm. long and spikelets 3 to 3.2 mm. long. This form can not be satis- factorily separated even as a subspecies, though extreme specimens differ sufficiently 41616°—vor 15—10——20 306 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. to be recognizable. The following specimens represent this variation: Chase 4205, Combs 89, 156, 886, 926, 1038, Curtiss 3583* in part, Hitchcock 757, 1008, Nash 1119, Tracy 3629. DISTRIBUTION. Woods and copses, Massachusetts to Illinois, south to Florida and Texas. Massacuusetts: Wellesley, Rich in 1899. PENNSYLVANIA: Lancaster County, Heller 4768; Ohiopyle, Ricker 1153. Onto: Vinton, Kellerman 6881. Inpiana: Clarke County, Deam 6595. Inuinois: Without locality, Schneck in 1879. Micnican: Agricultural College, Wheeler in 1890 (Mich. Agr. Col. Herb.). Missouri: Butler County, Eggert 257 in 1893; Carter County, Eggert 290; Monteer, Bush 755, 4685; Malden, Bush 731; Campbell, Bush 748; Pleasant Grove, Bush 238, 287, 308, 322; Grandin, Bush 2736. DELAWARE: Frankfort, Commons 48; Mount Cuba, Chase 3621, Commons 309. MaryLanpD: Great Falls, Chase 5436, Ward in 1882; Riverdale, Chase 2381. District or CotumBiA: Chase in Kneucker Gram. Exs. 557, Hitchcock 426, 569, Kearney 31, Merrill 204, Scribner in 1894, Steele in 1899, Vasey in 1874, 1881, 1883, and 1884, Ward in 1879. Virainia: Near Norfolk, Kearney 1317, 1414, 1463, Noyes 69, 74, Pollard & Mazon in 1900; Dismal Swamp, Chase 3654, 3678; Smyth County, Small 2, 8, and 14, in 1892. West Vrrainia: Fayette County, Kellerman 6902. NortH Carona: Jacksonville, Chase 3168; Wilmington, Chase 3114, Hitchcock 1458; Chapel Hill, Ashe in 1898, Chase 3052; Biltmore, Biltmore Herb. 802b, Hitchcock 570, 571; Lenoir, Hitchcock 428; Magnetic City, Wetherby 59; Colum- bus, Townsend in 1897; Rowan County, Small & Helier 201. SoutH Carouina: Orangeburg, Hitchcock 1413, 1414; Clemson College, House 2106, 2179, 2387. GeoraiA: Chickamauga, Ruth 11; Stone Mountain, Eggert 43, 45, Hitchcock 427, 1348; Augusta, Cuthbert in 1900; Athens, Harper 60; Rabun County, House 2305; Cobb County, Harper 221; Albany, Tracy 3630; Warm Springs, Tracy 8866; Brooks County, Harper 1623. Fioripa: Duval County, Curtiss 3583* in part, 4027, 4636 (last two in Hitchcock Herb.), Fredholm 455; Lake City, Combs 89, 139, 156, Hitchcock 572, 1008; Chattahoochee, Tracy 3629 in part; Apalachicola, Biltmore Herb. 802a; Madi- son, Combs 251; Monticello, Combs 330; Chipley, Combs 589, 601; Old Town, Combs 886; Homosassa, Combs 926; Titusville, Hitchcock 757; Clarcona, Meislahn 168a, 189; Grasmere, Baker 42, Combs 1038, 1098; Eustis, Nash 45 in part, 78, 240 in part, 302, 1119, 1675; Gainesville, Chase 4205; Manatee, Tracy 7382; Manavista, Tracy 6695. Kentucky: Harlan County, Kearney 34 in part, 172. Fig. 345.—Distribution of P. commutatum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 307 TENNESSEE: Hiwassee Gorge, Kearney 327; Wolf Creek, Kearney 972, 973, and in 1894; Knoxville, Ruth61; Lavergne, Biltmore Herb. 2984a; Kingston Springs, Gattinger in 1882; Jackson, Bain in 1893; Greenhill, Gattinger in 1881. ALABAMA: Scottsboro, Chase 4504; Auburn, Earle & Baker 1531, Hitchcock 1319, 1328, 1339; Tuskegee, Carver 60, 88, 93, 94; Blount County, Eggert 22; Etowah County, Eggert 7; Flomaton, Hitchcock 1049; Mobile, Mohr. Mississippi: Starkville, Tracy in 1888; Enterprise, Tracy 3296 in part; Biloxi, Tracy 4577 in part, Tracy in Pollard Distr. 1416; Ocean Springs, Earle in 1895; Petit Bois Island, Tracy 4577 in part; Bayou Gravelin, Tracy 4573; Mis- sissippi City, Hitchcock 1104, 1108. ARKANSAS: Fulton, Bush 234, 1450. Louisiana: Calhoun, Ball 66, Hitchcock 1268, 1274, 1286, 1291; Shreveport, Hitch- cock 1253; Alexandria, Ball 625; Plaquemines Parish, Langlois 39. Texas: Houston, Bebb 1242, Hall 831, Nealley 18, Ravenel 46; Texarkana, Heller 4083; Columbia, Bush 1224; Grand Saline, Reverchon 4143; Palestine, Plank 55. 185. Panicum mutabile Scribn. & Smith. Panicum mutabile Scribn. & Smith; Nash in Small, Fl. Southeast. U. S. 103. 1903. On page 1327 is the following citation: ‘‘Type, Biloxi, Miss., Tracy no. 3074, 1896 in Herb. Nash.’’ The type, in Nash’s herbarium, is the early branching form, the blades from the basal shoots large, 10 to 15 cm. long, 13 to 24 mm. wide, prominently ciliate almost to the apex, the culm blades ciliate toward the base only. DESCRIPTION. Vernal plants blue green, glaucous; culms solitary or few in a tuft, erect, 30 to 70 cm. high, glabrous or rarely minutely pubescent; sheaths much shorter than the internodes, glabrous except the usually sparsely ciliate margin; ligules very short, membranaceous-fimbriate; blades horizontally spreading, 6 to 15 cm. long, 8 to 20 mmm. wide, tapering to both ends, rather thin, glabrous on both surfaces, ciliate on the margin toward the cordate base or the lower ciliate nearly to the apex; panicles 7 to 15 cm. long, about as wide, very loosely few-flowered; spikelets 2.9 to 3 mm. long, 1.2 mm. wide, elliptic, minutely pointed, pubescent; first glume about one-third as long as the spikelet, subacute; second glume and sterile lemma equaling or slightly exceeding the fruit; fruit 2.6 mm. long, 1.1 mm. wide, elliptic, subacute. Autumnal form erect or reclining, sparingly branching from the middle and upper nodes, the branches longer than the internodes, finally producing toward the ends fascicles of shortened branchlets with somewhat reduced leaves. This species differs from P. commutatum in the glaucous blue green color, the more slender culms, solitary or few in a tuft, the glabrous sheaths, the narrower blades and the conspicuously ciliate basal blades. In herbarium specimens the glaucous color may be seen best on the culm below the nodes. Fie. 346.—P. mutabile. From typespecimen. 808 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. DISTRIBUTION. Sandy pine woods or hammocks, southeast Virginia to northern Florida and west to Mississippi. Virainia: Cape Henry, Chase 5437, Hitchcock 429. Norra Caroutna: Wards Mill, Chase 3189; Wilmington, Chase 3154, Hitchcock 573, 1465. Groraia: Stone Mountain, Hitch- cock 430; Clarke County, Har- per 147; Coffee County, Harper 1435; Thomson, Bartlett 1508. Fioripa: Lake City, Chase 4290, Hitchcock 1015, 1035; Madison, Combs 224; Monticello, Combs 299, 307; Bay Head, Combs . 653; Gainesville, Combs 740. Fic. 347.—Distribution of P. mutabile. ALABAMA: Gateswood, Tracy 8424. MisstssipPi: Biloxi, Kearney 336, Tracy 3646; Mississippi City, Hitchcock 1103; Bayou Gravelin, Tracy 4573 (Field Mus. Herb.). 186. Panicum joorii Vasey. Panicum leiophyllum Fourn. Mex. Pl. 2: 20. 1886, not Nees, 1829. The only speci- men cited is ‘‘In valle Cordovensi, januario (BourG[zAU] absque n.).”” This name was earlier listed by Hemsley without description. The type, in the Paris Herba- rium, consists of several primary culms beginning to branch, the culms puberulent, the largest blades about 8 cm. long and 12 mm. wide. Panicum joorti Vasey, U. S. Dept. Agr. Div. Bot. Bull. 8: 31.1889. ‘“‘Louisiana (Dr. J. F. Joor).’’ The type, in the National Herbarium, is a clump of several autum- nal culms, branching at all the nodes, the primary summits mostly fallen and replaced by branches bearing fascicled branchlets, the primary blades as much as 16 em. long and 18 mm. wide, more or less falcate. The accompanying label reads: ‘‘ Panicum Joorii, Creek bank, in dense tufts! near Baton Rouge, La. Oct. 1, ’85. No. 39 Legit J. F. Joor, M. D.” Panicum manatense Nash, Bull. Torrey Club 24:42. 1897. ‘‘Collected by the writer on August 21, 1895, near a sulphair well in a wet hammock northeast of Pal- metto, Manatee County, Florida, no. 2428a.’’ The type, in Nash’s herbarium, is a clump of early autumnal culms branching from all the nodes, the largest primary blades being scarcely 8 cm. long and 15 mm. wide, somewhat falcate. DESCRIPTION. Vernal form rather pale grayish green; culms in clumps of few to many, 20 to 55 cm. high, slender, spreading or ascending from a decumbent base, glabrous or rarely puber- ulent, at least the lower internodes purplish red; sheaths shorter than the internodes, ciliate, otherwise glabrous or rarely puberulent between the nerves; ligules nearly obso- lete; blades ascending or spreading, 6 to 15 cm., rarely 18cm. long, 7 to 18 mm. wide, thin but firm, often subfalcate, acuminate, narrowed toward the rounded base, usually ciliate at base, otherwise glabrous; panicles short-exserted, 5 to 9 cm. long, about two-thirds as wide, loosely flowered, the branches ascending or spreading; spikelets 3 to 3.1 mm. long, 1.2 to 1.3 mm. wide, elliptic, abruptly short-pointed, pubescent; first glume one-third to two-fifths as long as the spikelet, acute; second glume and a Biol. Centr. Amer. Bot. 3: 490. 1885, HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 309 sterile lemma more or less pustulate-papillose between the nerves, the glume slightly shorter than the usually involute-pointed sterile lemma; fruit 2.4 mm. long, 1.2 mm. wide, elliptic, minutely umbonate. Autumnal form widely spreading, bearing more or less divaricate branches from all the nodes, these primary branches longer than the internodes and branching from all or from their upper nodes, the ultimate branchlets in short, dense fascicles, the reduced blades ascending, exceeding the numer- ous, small, partly included panicles; winter rosette a crown of a few short, leafy, basal shoots, with evident internodes. The type of P. joorii is an exceptional speci- men, having unusually large blades. The only other collection seen which entirely matches the type is Chase 4274 (Lake City, Florida), in which the blades are even a little longer than in Joor’s no.39. Harper’s no. 1106 (Georgia) and Tracy 120 and 3318 (Mississippi) are much like the type. The type of P. manatense represents the usual form in its early autumnal state. As a whole this species has glabrous culms, sheaths, and blades, but occasional specimens more or less puberulent are found, such as Combs 860, Heller 4120, and Hitchcock Lee Co. Pl. 477. Three specimens with spikelets only 2.2 to 2.5 mm. long, and with more or less pubescence on culms and sheaths are doubtfully referred here: Ftormpa: Orange Bend, Chase 4094. Lovistana: Burnside, Combs 1427; Plaquemines Parish, Langlois 39. Fia.348.—P. joorii. From typespecimen. DISTRIBUTION. Low or swampy woods, southeastern Virginia to Florida, west to Texas and Arkansas; also in Mexico. VirerniA: Princess Anne County, Mackenzie 1794. Sours Caro.tna: Isle of Palms, Chase 4550. Geroraia: Burke County, Harper 767; Leslie, Harper 1106; Albany, Tracy 3655; Thomson, Bartlett 1501. Fiormwa: Lake City, Chase 4274, Combs 144, Hitchcock 1011; Madison, Combs 270; Chattahoochee, Curtiss in 1884; Quincy, Combs 412; Tallahassee, Combs 373; Old Town, Combs 860; Titusville, Chase 4019, Hitchcock 756, 760; Eustis, Nash 240 in part; Orange Bend, Chase 4099; El- dorado, Chase 4126; Gainesville Chase 4219, 4256; Dunnellon, Combs 921; Mary Esther, Tracy 9142; Bartow, Combs 1237; Tampa, Combs 1393, 1400, Hitchcock 947; Manatee, Hitchcock 976, Rugel 351, 378; Myers, Chase 4181, Hitchcock 909, Lee Co. Pl. 471; Alva, Hitchcock Lee Co. Pl. 477; Miami, Chase 3907, 3948, Eaton 169, Hitchcock 576, 636, 652, 660, 674. ’ AtaABAMA: Chehaw, Hitchcock 577. Mississreri: Starkville, Chase 4468; Tracy 29 in 1889; Morrisville, Tracy 3387; Bludlebury, Tracy 3318; Lake, Tracy 120 in 1888; Meridian, Tracy 3266. ArKansas: Lafayette County, Heller 4120. Fic. 349.—Distribution of P. joorii. 310 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Louisiana: Rayville, Ball 25; Lake Charles, Chase 4430, 4437, Hitchcock 1142; Plaquemines Parish, Langlois 41 in part. Texas: Waller, Hitchcock 1207, 1208, Thurow in 1898; Houston, Hall 828 (Gray Herb.). Mexico: Cérdoba, Bourgeau in 1866 (Paris Herb.). 187. Panicum equilaterale Scribn. Panicum equilaterale Scribn. U. 8. Dept. Agr. Div. Agrost. Bull. 11: 42. pl. 2. 1898. Two specimens are cited as follows: ‘‘In pine lands [Eustis], Florida (No. 1120, George V. Nash, June, 1894); scrubby hammock lands [Eustis], Florida (No. 1674, George V. Nash, August, 1894).’’? The plate cited above is drawn from Scribner’s specimen of Nash 1674, now in Hitchcock’s herbarium, which specimen is therefore the type. It consists of two culms 53 and 65 cm. high, one simple, the other producing short fas- cicled branches at the upper two nodes. The blades are 10 to 17 em. long. Panicum epilifolium Nash, Bull. Torrey Club 26: 571. 1899. ‘‘The type col- lected by the writer in a scrub hammock at Eustis, Lake Co., Florida, March 12-31, 1894, no. 45.’’ The type, in Nash’s herbarium, is the vernal form, with scarcely mature panicles. The longest blade is but 7.5 cm. long, but a duplicate specimen in the National Herbarium has blades as much as 10 cm. long. Two species were dis- tributed under Nash’s no. 45, the other being P. commutatum. DESCRIPTION. Vernal plants glossy, grayish green, in clumps of several to many culms, these 25 to 70 em. high, stiff and erect, glabrous or nearly so, including the nodes; sheaths much shorter than the elongated internodes, or the upper two approximate, glabrous except for the densely short-ciliate margin; ligules nearly obsolete; blades firm, widely spreading or ascending, 6 to 17 cm. long, 6 to 14 mm. wide, very scabrous (the margins nearly parallel), often ciliate at the rounded orsubcordate base, acuminate, glabrous on both surfaces, often drying conduplicate; panicles usually short-exserted, loosely flowered, 5 to 10 cm. long, two-thirds to three-fourths as wide, the branches ascending; spikelet 3.2 mm. long, 1.3 mm. wide, obovate-elliptic, obscurely pointed, pubescent; first glume half the length of the spikelet or more, rather remote, triangular, acute; second glume and sterile lemma subequal, barely covering the fruit at maturity; fruit 2.6 to 2.7 mm. long, 1.25 mm. wide, elliptic, minutely umbonate. Fig. 350.—P. equilaterale. From typespecimen. Autumnal form erect or leaning, branching from the upper and middle nodes after the maturity of the primary panicle, these primary branches often longer than the inter- nodes and producing short, fascicled, appressed branchlets with reduced spreading blades from their uppermost nodes, the numerous small panicles partly included; winter rosette appearing late, the blades lanceolate, firm, sometimes ciliate. This species is distinguished from P. commutatum by its almost linear blades, which vary much in size but are characteristically parallel-margined, and by its branching from the uppermost nodes of both the primary culm and the branches; and from P. manatense, which branches in the same way, by the erect habit and rather distant first glume half as long as the spikelet. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 311 DISTRIBUTION. In pine lands, hammocks, and sandy woods, South Carolina to southern Florida. Sours Caroutna: Isle of Palms, Ball 791, Chase 4537, 4549, Hitchcock 421. Friorma: Duval County, Curtiss 3600 A in part (Gray Herb.); St. Augustine, Ricker 943; Ce- dar Key, Combs 793; Apa- lachicola, Chapman; Pensa- cola, Tracy 8428; Grasmere, Combs 1053; Gainesville, Chase 4241, 4258, Combs 737; Homos- sassa, Combs 9264; Crystal, Combs 1023; Clarcona, Pieters 116; Orange County, Baker 69; Eustis, Chase 4063, 4122, Nash Fig. 351.—Distribution of P. equilaterale. 45 in part, 1120, 1674; Clear- water, Tracy 7167; Palma Sola, Tracy 6713 in part; Sneeds Island, Tracy 6460, 7050, 7051 in part: Miami, Chase 3908, Hitchcock 574, 633, 653, 655; Ormond, Hitchcock 575. 188. Panicum albomaculatum Scribn. Panicum albomaculatum Scribn. U. 8. Dept. Agr. Div. Agrost. Circ. 19: 2. 1900. “Dry rocky hills, Patzcuaro, State of Michoacan [Mexico], October 10. C.G. Pringle, No. 5203, 1892.’’ The type, in Hitchcock’s herbarium, consists of a tuft of primary culms with mature panicles, the culms beginning to branch from the lower nodes. DESCRIPTION. Vernal culms tufted, 50 to 100 cm. high, arising from a knotty crown, erect, slender but stiff, minutely puberulent between the striz, at least below the glabrous nodes; sheaths long, the lower overlapping, minutely puberulent, more or less mottled with white spots, densely short-ciliate on the margin; ligules about 0.2 mm. long; blades i Fig. 352.—P. albomaculatum. From type specimen. firm, spreading or ascending, reflexed when old, 8 to 17 cm. long, 9 to 15 mm. wide, linear- lanceolate, acuminate, rounded and ciliate at the base, roughish on the upper surface and with a narrow line of appressed pubescence along the very scabrous margins, glabrous or obscurely pubescent beneath; panicles long- exserted, ovate in outline, 10 to 16 cm. long, two-thirds to three-fourths as wide, loosely flow- ered, the axis and ascending branches glabrous; spikelets 2.8 mm. long, 1.2 mm. wide, obovate elliptic, obtuse, sparsely pubescent; first glume nearly or quite half the length of the spikelet, obtuse, or subacute; second glume and sterile lemma equaling the fruit at maturity; fruit 2.2 mm. long, 1.2 mm. wide, elliptic, subacute. Autumnal form not known. * Probably most nearly related to P. equilaterale Scribn. a2 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, DISTRIBUTION. Known only from type locality. Mexico: Patzcuaro, State of Michoacan, Pringle 5203. Latifolia.—Culms rather stout, usually more than 50 cm. high; ligules not over 1 mm. long; blades ample, usually more than 1.5 cm. wide, cordate; spikelets 2.7 to 4.5 mm. long, rather turgid, pubescent, 7 to 9-nerved. Autumnal form not very freely branching. Sheaths strongly papillose-hispid, at least the lower and those GLble PEAMCN eet ao. Pes. of.» ales cee oe toe eee ere 189. P.clandestinum. Sheaths glabrous or softly villous. Nodes glabrous; spikelets 3.4 to 3.7 mm. long .......-.---- 190. P. latifolium. Nodes bearded; spikelets 4 to 4.5 mm. long. Blades glabrous or nearly so on both surfaces.......---- 191. P. boscit. Blades yelvety beneath. .......- 2-20 geet ee pee 19la. P. boscit molle, 189. Panicum clandestinum L. Panicum clandestinum L. Sp. Pl. 58. 1753. ‘‘ Habitat in Jamaica, Pensylvania. Kalm.’”’ In the Linnean Herbarium is a specimen marked ‘‘clandestinum K” [Kalm]in Linnzeus’s writing, which is taken as the type since it is the only specimen so marked by Linnzeus, and since it agrees with his description. The Jamaica locality is evidently based upon the Sloane phrase name and figure cited as synonym. This figure represents Hackelochloa granularis (L.) Kuntze, a species to which Linnzeus’s description does not at all apply, for which reason the Jamaica locality is rejected. The Kalm specimen is the autumnal form, the secondary panicles inclosed in the sheaths, which are crowded at the summit. Milium clandestinum Moench, Meth. Pl. 204. 1794. Based on Panicum clandes- tinum L. Panicum latifolium clandestinum Pursh, Fl. Amer. Sept. 1: 68. 1814. Based on P. clandestinum ‘‘Willd. sp. pl. 1. p. 351.’? In Willdenow’s Species Plantarum the original Linnean diagnosis, citations, and description are given, with reference to “Spord.ooo. & Panicam pedunculatum Torr. Fl. North. & Mid. U. 8. 141. 1823. ‘On the Island of New-York.’? The type, in Columbia University Herbarium, is a vernal specimen 80 cm. high, beginning to branch, with an over-mature, long-exserted, primary panicle, the spikelets 2.8 mm. long. The accompanying label, in Torrey’s hand, reads ‘‘Panicum pedunculatum*” [followed by a brief diagnosis] ‘‘In wet meadows, among thickets. Aug.’ Panicum clandestinum pedunculatum Torr. Fl. N. Y. 2: 426. 1843. Based on P. pedunculatum Torr. Panicum decoloratum Nash, Bull. Torrey Club 26: 570. 1899. ‘‘Collected by Mr. E. P. Bicknell on a sandy railroad bank at Tullytown, Pennsylvania, May 30, 1899.” The type, in Nash’s herbarium, consists of four vernal specimens with partially included, immature panicles. The lower sheaths are papillose-hispid, less densely so than common in P. clandestinum, the upper glabrous. The immature spikelets are 2.7 mm. long. DESCRIPTION. Vernal form in large, dense clumps, sometimes with strong rootstocks, 5 to 10 cm. long; culms stout, 70 cm. to 1.5 meters high, erect, scabrous to papillose-hispid, at least below the nodes; sheaths as long as the internodes or overlapping until after the a§Sp. Pl. ed. 2. 86. 1762. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 38138 branches appear, loose, strongly papillose-hispid to nearly glabrous, a puberulent ring at the summit; ligules 0.5 mm. long; blades spreading or finally reflexed, 10 to 20 cm. long, 1.2 to 3 cm. wide, slightly tapering to the cordate-clasping base, acuminate, scabrous on both surfaces, at least toward the end, usually ciliate at the base; panicles finally rather long-exserted, 8 to 15 cm. long, about three-fourths as wide, many- flowered, the flexuous branches in distant fas- cicles, short spikelet-bearing branchlets in the axils; spikelets 2.7 to 3 mm. long, 1.4 tol.5 mm. wide, obovate-oblong, sparsely pubescent; first glume one-third the length of the spikelet, sub- acute or obtuse; second glume slightly shorter than the fruit and sterile lemma; fruit elliptic 2.1 to 2.3 mm. long, 1.2 to 1.3 mm. wide. Autumnal form erect or leaning, sparingly eae Wrenilceitwwes?. Brom typeapect- branching, often before the maturity of the men of P. pedunculatum Torr. primary panicle, from the middle and upper nodes, the branches leafy, the swollen, bristly sheaths overlapping on the shortened internodes and inclosing wholly or partially the secondary panicles; spikelets more turgid than those of the primary panicles. Occasional specimens, such as the type of P. decoloratum, and Eggert 114 and 253, Merrill 198, and one specimen of Small & Heller, Blowing Rock Mountain, Tenn., in 1891 (a second being the ordinary form) are bristly only on the lower sheaths, the upper glabrous or only scabrous. This lack of pubescence is not found to be correlated with smaller spikelets nor with scarcely-exserted panicles. In Andrews 11 the spikelets are exceptionally small, only 2.5 mm. long, but the sheaths and culms are bristly, and the developed panicle is long-exserted and an immature one scarcely exserted. DISTRIBUTION. Moist, mostly sandy ground, Maine to Kansas, south to Florida and Texas. Marne: Foxcroft, Fernald 292, 518; Farmington, Chamberlain & Knowlton in 1902. New Hampsuire: Gilford, Carter 111 (Hitchcock Herb.). Vermont: Burlington, Flynn in 1900 (Hitchcock Herb.). MassacuuseEtts: South Hadley, Cook in 1887. Connecticut: Pine Meadow, Bissell 5544; Southington, Andrews 11, Bissell 5546; South Manchester, Hitchcock 583; South Glastonbury, Wilson 17; Greens Farms, Pollard 92; Oxford, Harger in Kneucker Gram. Exs. 424. Ruope Istanp: Without locality, Congdon (Hitchcock Herb.). New Yor«: Oneida County, Ha- berer 1257. New Jersey: Clifton, Nashin 1891; Stockholm, Van Sickle in 1895;- Woodbury, Smith 112; South Amboy, Mackenzie 1360; Wild- wood, Chase 3513. Fig. 354,—Distribution of P. clandestinum. PENNSYLVANIA: Easton, Porter in 1895 and 1897; Refton, Heller in 1901; Mount Hope, Heller 4781; Tullytown, Bicknell in 1899. Onto: Berea, Ashcroft in 1897; Cadiz Junction, Kellerman 67993; New Ply- mouth, Kellerman 6883; Vinton, Kellerman 6888. 314 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Inp1ANA: Michigan City, Hill 158 in 1906; Brazil, Somes 231. Inuinots: Peoria, Brendel, McDonald in 1903; without locality, Wolf in 1882. MicuicANn: Port Huron, Dodge in 1904 and 1909. Missourt: St. Louis, Eggert 114, 253, Kellogg 18; Sibley, Bush 4003; Independ- ence, Bush 733 in part; Courtney, Bush 3977; Allenton, Kellogg in 1901; Monteer, Bush 4651, 4865. Kansas: Pottawatomie County, Norton 878. DetawareE: Wilmington, Commons 27, 302; Centerville, Commons 300. MaryLanp: High Island, Ward in 1878; West Chevy Chase, Chase 3272, Hitchcock 432; Riverdale, Chase 2366. District or CotumBiA: Ball 702, Hitchcock 431, Kearney 21, 32, Merrill 198, Pol- lard 365, Scribner in 1894, Sudworth in 1890, Vasey in 1874 and 1887, Ward in 1880, Williams 7. VireiniA: Portsmouth, Noyes 68. West Virernia: Aurora, Steele in 1898; Morgantown, Hitchcock 584; Fayette County, Kellerman 6900. Nortu Carona: Biltmore, Biltmore Herb. 804, 804b; Blowing Rock Mountain, Small & Heller in 1891; Heiligs Mill, Small & Heller 348; Magnetic City, Wetherby 32; Chapel Hill, Ashe, Chase 3078. Sours Caroiina: Clemson College, House 2132. Georaia: Gwinnett County, Small in 1893: Stone Mountain, Hitchcock 585; Clarke County, Harper 70. Fiorina: Without locality, Chapman. Kentucky: Poor Fork, Kearney 229. TENNESSEE: Wolf Creek, Kearney in 1897, Ruth 60; Ducktown, Chambliss 18, 78; Hiwassee Gorge, Kearney 330; Blowing Rock Mountain, Small & Heller in 1891. ALABAMA: Without locality, Buckley (Mo. Bot. Gard. Herb.). Mississippi: Starkville, Chase 4463. ARKANSAS: Northwest Arkansas, Harvey 32. Texas: Dallas, Reverchon in 1875 (Gray Herb.). OKLAHOMA: Supulpa, Bush 718 (Mo. Bot. Gard. Herb.). 190. Panicum latifolium L. Panicum latifolium L. Sp. Pl. 58. 1753. ‘‘ Habitat in America.’’ The type, in the Linnzan Herbarium, is a portion of a vernal culm with two leaves and a short- exserted but well-developed primary panicle.@ Milium latifolium Moench, Meth. Pl. 204. 1794. Based on Panicum latifolium L. Panicum macrocarpon LeConte in Torr. Cat. Pl. N. Y. 91.1819. No locality nor specimen is cited. In the Torrey Herbarium is a specimen with the following label a For a full discussion of the reasons for considering this specimen to be the type, see Hitchcock, Contr. Nat. Herb. 12: 118. 1908. The reasons are briefly: In the Linnzan Herbarium are two sheets upon which Linneus has written the name. One of these, which is Panicum zizanioides H. B. K., was received from Browne after the preparation of the manuscript of the Species Plantarum. The second sheet includes two vernal specimens, Panicum macrocarpon LeConte, and P. clandestinum L. The first of these is taken as the type as it corresponds better to the description. The sheet marked ‘‘17 K latifolium,’’ meaning that the specimens were received from Kalm and the species is no. 17, P. latifolium, in the Species Plantarum. Appended to the original description, Linnzeus cites two synonyms, one from Morison, which is prob- ably P. boscit Poir., and one from Sloane, which is P. sloaneit Griseb. The Sloane specimen is considered by some authors to be the type of P. latifolium, but the Sloane synonym is erroneously cited. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 315 in Torrey’s writing: ‘‘‘ Panicum macrocarpon Elliott’ LeConte.’’ Torrey’s note on this specimen would seem to have been written before the publication of the species as above, as he seems to think LeConte credits the name to Elliott. Nodata are given on the label or sheet. No other specimen could be found in the herbarium that could be connected with LeConte’s name. If LeConte had a herbarium and if it be in exist- ence, its whereabouts is unknown. The above specimen, which is taken as the type of P. macrocarpon, is a single plant with simple culm and immature unexpanded panicle. Panicum schnecki Ashe, N. C. Agr. Exp. Sta. Bull. 175: 116. 1900. ‘‘River bot- toms, southern Indiana and Illinois, June.’ The type, in Ashe’s herbarium, consists of two culms, one sterile and one with an immature, scarcely-exserted panicle. On the accompanying label with the printed heading ‘“‘Herbarium of W. W. Ashe” is written in Ashe’s hand ‘‘ Panicum Schnecki W. W. Ashe,’’ but no data whatever are given. This is the species described by Torrey@ as P. nervosum Muhl. Panicum macro- carpon LeConte, though published in his Catalogue seems to have been unknown to Torrey. DESCRIPTION. Vernal plants in clumps of few to several culms from a knotted crown; culms rather stout, 45 cm. to 1 meter high, erect, glabrous, or the lower internodes sometimes sparsely pubescent, the nodes glabrous, rarely with a few hairs; sheaths shorter than the internodes, ciliate on the margin, a pubescent ring at the juncture with the blade, otherwise glabrous, or the lower sparsely downy; ligules nearly obsolete; blades rather thin, ascending or spreading 8 to 18 cm. long, 1.5 to 4 em. wide, the lower smaller, acuminate, cordate-clasping at the base, glabrous, rarely sparsely pubescent on one or both surfaces, short- ciliate at least toward the base; panicles usually short-exserted or sometimes finally long-exserted, 7 to 15 cm. long, about two-thirds as wide, the long, few- flowered, nearly simple branches stiffly ascending; spikelets rather short-pedi- celed, commonly more or less clustered in twos or threes, 3.4 to 3.7 mm. long, 1.8 to 2 mm. wide, oval-obovate, turgid, sparsely pubescent; first glume one- third to half the length of the spikelet, pointed; second glume and sterile lemma scarcely covering the fruit at maturity; fruit 3 mm. long, 1.6 to 1.8 mm. wide, elliptic, minutely pubescent on the obscurely apiculate apex.” Autumnal form more or less spreading, branching from the middle nodes, the upper leaves of the branches crowded and spreading, not much reduced, the small panicles partly included. In this species the culms, nodes, and leaves are typically glabrous, but occasional specimens show more or less pubescence. The following represent this variation: Dorner 89, Shear in 1899, Ward in 1880, Wetherby 53. a4F\. North. & Mid. U.S. 143. 1823. b See P. macrocarpon Torr, under P. scribnerianum, page 283. Fic, 355.—P. latifolium. From type specimen of P. macrocarpon LeConte. 316 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, DISTRIBUTION. Rocky or sandy woods, Maine to Minnesota, south to North Carolina and Kansas. Maine: Orono, Fernald 346; Basin Mills, Knight 53; North Berwick, Parlin 1191; Fayette, Chase 3367; East Auburn, Merrill 1248. New Hampsuire: Surry, Fernald 276 (Gray Herb.). Vermont: Manchester, Day 216; Barnet, Blanchard in 1883. ‘Massacuusetts: South Hadley, Cook in 1887. Connecticut: Southington, Andrews 44, Bissell 5548, 5549, 5550; Old Lyme, Graves 164; Montville, Graves 163; Waterford, Graves in 1898; Stratford, Eames in 1894; Oxford, Harger in Kneucker Gram. Exs. 423. RuoveE Isianp: Providence, Olney. New York: Ithaca, Ashe, Coville in 1885, Dudley in 1885; Oxford, Coville in 1884; Apalachin, Fenno 3, 5, 10, 11, 12; Cairo, Nash in 1893; Long Island, Bicknell in 1903 and 1905. Ontario: Galt, Herriot in 1898 and 1901; Queenstown Heights, Macown 26326, 26327; Tilsonburg, Macown 26325. New Jersey: Clifton, Nash in 1892; Phillipsburg, Porter in 1892 and 1895; Mount Arlington, Mackenzie 1414. PENNSYLVANIA: Easton, Porter in 1895, 1897, and 1898; Germantown, Stone in 1889; Westchester, Windle in 1904; McCalls Ferry, Rose & Painter 8135. Ouro: Niles, Ingraham in 1891; Lancaster, Kellerman 6766; Cadiz Junction, Kellerman 6799. InpiaNnA: Miller, Chase 1541, Um- bach 1791 and in 1898; Pine, Umbach in 1896; Wells Coun- ty, Deam in 1903; Lafayette, Dorner 82. Iuurnots: Beverly Hills, Bebb 882, Chase 1824; Savanna, Chase 1887; Glasford, Wilcox 22, 41; Mossville, Chase 889; Antioch, Gleason & Shobe 268; Peoria, Brendel, McDonald 33; Urbana, Waite in 1886; St. Clair County, Eggert 239; Wabash County, Schreck in 1879 and 1905. MicuicaNn: Detroit, Farwell 1378; Port Huron, Dodge in 1909. Wisconsin: Newbold, Cheney 1545; Milwaukee, Chase 1947. Minnesota: Lindstrom, Yaylor in 1892; Spring Grove, Rosendahl 487; Houston, Mearns 791. Iowa: Fayette County, Fink 593; Ames, Ball 40, 155, Ball & Sample 6; Iowa City, Shimek in 1894; Mount Pleasant, Mills in 1894. Missouri: McDonald County, Bush 91; Independence, Bush in 1884; Swan, Bush 8; Monteer, Bush 190; Courtney, Bush 1732, 3981; Sibley, Bush 4001, 4807; Jefferson Barracks, Kellogg 17; Midway, 2. J. Palmer 1405. Kansas: Manhattan, Kellerman (Hitchcock Herb.). DeLtawaRreE: Centerville, Commons 301 in part. MaryYLANp: Garrett County, J. D. Smith in 1879. District or CoLtumBiA: Shear in 1899, Ward in 1880. VirainiA: Norfolk, Pollard & Maxon in 1900; Munden, Mackenzie 1709; Fairfax County, Hitchcock 587; Carroll County, Small in 1892; Peaks of Otter, Palmer 10. West Virarnia: Aurora, Steele in 1898; Marlinton, Kellerman 6899. Fig. 356.—Distribution of P. latifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 317 Nort Carouna: Magnetic City, Wetherby 53; Blowing Rock Mountain, Small & Heller 394; Biltmore, Hitchcock 588; Linville, Hitchcock 589; Grandfather Mountain, Hitchcock 433. Kentucky: Lexington, Short 5 (Gray Herb.). TENNESSEE: Roan Mountain, Hitchcock 586. 191. Panicum boscii Poir. Panicum boscii Poir. in Lam. Encycl. Suppl. 4: 278.1816. ‘Cette plante a été recueillie 4 la Caroline par M. Bosc. (V.s. in herb. Desfont.).’”” The type, in the Desfontaines Herbarium, consists of three early autumnal culms, the blades nearly glabrous, the spikelets 4.1 mm. long. The accompanying label reads ‘‘Panicum boscii poir. enc. Suppl. (scr. Poiret.) Amer. Sept. & Bosc.” Panicum waltheri Poir. in Lam. Encycl. Suppl. 4: 282. 1816, not P. walteri Pursh, 1814. Based on ‘‘Panicum latifolium Mich. Flor. bor. Amer. 1. pag. 49.—Non Linn. * * * (V.s. in Herb. Mich.).’’ The specimen in the Michaux Herbarium under the name of P. latifolium, and labeled “in pratis sylvestris Virginiz Caro- line,’’ is a somewhat pubescent autumnal specimen of P. boscii. Scribner @ uses this name (as P. walteri Poir.) for P. boscii. Panicum latifolium australis[e] Vasey, U. 8. Dept. Agr. Div. Bot. Bull. 8: 34. 1889. “Alabama to Texas.’’ The type, in the National Herbarium, consists of two vernal culms, the sheaths and blades glabrous or nearly so, the spikelets 4.3 mm. long. The specimen was collected by Dr. Charles Mohr, at Thomasville, Clark County, Ala., April 16, 1888. Panicum porterianum Nash, Bull. Torrey Club 22: 420. 1895. The author pro- poses a new name for ‘Panicum latifolium Walt. Fl. Car. 73. 1788. Not Linneus, 1753. Panicum Walteri Poir. in Lam. Encycl. Suppl. 4: 282. 1816. Not Pursh, 1814. Panicum latifolium var. molle Vasey, Bull. Bot. Div., U. S. Dept. of Agric. 8: 33. [error for 34] 1889.”” No description is given. In Walter’s Flora Linnzeus’s diagnosis of P. latifolium is copied with no additional description, and there is no specimen in Walter’s herbarium ® labeled P. latifolivm by Walter. A fragmentary panicle and two leaves, doubtfully referable to P. commutatum Schult., labeled ‘‘ Panicum 469,” possibly may be the specimen to which Walter applied the name. In the absence of an authentic specimen, and because P. latifolium Walt. must be considered a misappli- cation of P. latifolium L., P. porterianum is here regarded as based upon the next name cited, P. walteri Poir. Since the name was given in honor of Dr. Porter, doubt- less because of his distinguishing this form from what he understood to be P. latifolium L.,¢ it is more fitting also that the name P. porterianum be based on the name which he applied to this form, that is, P. watteri Poir. This species has been described as P. latifolium LL. by many American authors. DESCRIPTION. Vernal form resembling that of P. latifolium, usually in larger clumps; culms 40 to to 70 cm. high, erect or ascending, glabrous or minutely puberulent, rarely somewhat papillose, the nodes retrorsely bearded; sheaths much shorter than the internodes, ciliate, a pubescent ring at the juncture with the blade, otherwise glabrous or sparsely downy-pilose; ligules about 1 mm. long; blades spreading, 7 to 12 cm. long, 1.5 to 3 em. wide, more or less tapering toward the sparsely ciliate, cordate base, acuminate, glabrous on both surfaces or puberulent on the lower and sparsely pubescent on the ‘@Mem. Torrey Club. 5: 32. 1894. b For an account of the grasses in Walter’s herbarium see Hitchcock, Rep. Mo. Bot. Gard. 16: 36. 1905. ; ¢ Bull. Torrey Club 20: 194. 1893, 318 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. upper surface; panicles 6 to 12 cm. long, as wide or wider, the main axis and the flexuous spreading or ascending branches puberulent; spikelets 4 to 4.5 mm. long, 2 to 2.2 mm. wide, oblong-obovate, less turgid than in P. latifolium, papillose- pubescent; first glume one-third to two-fifths the length of the spikelet, pointed; second glume slightly shorter than the fruit and sterile lemma at maturity; fruit 3.2 to 3.5 mm. long, 1.5 to 1.6 mm. wide, elliptic, minutely pubescent, and usually black at the tip. Autumnal form as in P. latifoliwm, rather more freely branching, sometimes top- heavy reclining; spikelets more turgid than those of the primary panicles. As in P. commutatum, what appear to be simple, very leafy, autumnal culms are commonly formed by the replacing of the fallen primary summit by the uppermost branch. These culms are often recurved, the blades all turned into the plane of the branch, or even all to one side, from a twisting of the internodes, and more or less unsym- metrically expanded in the middle, the whole producing a very different aspect from that of the vernal form. This species is closely related to P. latifolium and is often difficult to distinguish from pubescent speci- mens of that species. The form distinguished by Vasey as P. latifolium australe can not be \ satisfactorily separated. The longer Fi. 357.—P. boscii. From type specimen. spikelets and blades narrowed to the base, or rather expanded in the middle, are not correlated. Specimens with large spikelets are mostly of southern range, but those with blades strongly tapering to the base occur throughout, Andrews 40, Connecticut, being a pronounced case. The following specimens represent this variation: Chase 4491, Combs 334, 337, 399, 683, Commons 361, Hall 828, Harper 1812, Mohr in 1888, Nash 2329. In most of the Alabama and Florida specimens the longer spikelets and tapering blades are correlated and might be given subspecific rank but for the more numerous intermediate specimens. The larger spikelets and tapering blades also occur in P. boscii molle. ; The following specimens are intermediate in character between P. boscii and P. boscii molle, having blades more or less downy, or glabrous and downy blades on the same plant: Bush 303, 310, 3348, Chase 4501, Harper 1366, Tracy 3 in 1888. DISTRIBUTION. Woods, Massachusetts to Oklahoma, south to northern Florida and Texas. MassacuuseErtts: Monson, Herb. Western High School of Washington (Biltmore Herb.). Connecticut: Southington, Andrews 40, Bissell 5552; Ledyard, Graves in 1897. New Jersey: Byram Station, Fisher in 1897; Milburn, Mackenzie 2144. PENNSYLVANIA: Lancaster County, Heller 4771; Upper Darby, Smith 154. Ouro: Painesville, Werner in 1886; Cincinnati, Lloyd 3597. InpIANA: Clarke County, Deam 5386, 6586. Inu1nors: Marshall County, V. H. Chase 1489; Jackson County, French in 1905; Tunnel Hill, Ridgway in 1902; Wabash County, Schneck in 1896. Missourt: Allenton, Kellogg 15; Pleasant Grove, Bush 303, 310, 323; Doniphan, Bush 276; Swan, Bush 26, 4534; Monteer, Bush 4652, 4908. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 319 DetawarE: Wilmington, Commons in 1897; Centerville, Commons 361. MARYLAND: Chesapeake Beach, Hitchcock 1604; Riverdale, Chase 2374; Plum- mers Island, Hitchcock 590. District or CotumBia: Ball 14, Kearney in 1897, Scribner in 1894, Sudworth in 1890, Vasey in 1884, Ward in 1876 and 1878. Virerta: Rosslyn, Dewey 235; Glencarlyn, Dewey 280; Marion, Small in 1892; Ocean View, Kearney 1469; Virginia Beach, Kearney 1411. Nort Caroura: Asheville, Canby 220; Biltmore, Hitchcock 591; Columbus, Townsend in 1897; West Raleigh, Coit 1293; Chapel Hill, Chase 3073; Spray, De Chalmot; without locality, Vasey in 1878. Souta Carona: Aiken, Ravenel; Clemson College, House 2136; Orangeburg, Hitchcock 19, 1412. Geroraia: Stone Mountain, Hitchcock 222, Eggert 40; Clyo, Harper 1812; Dublin, Harper 1366; Thomson, Bart- lett 1505; Tallulah Falls, Cuth- bert in 1899. Fiorma: Chipola River, Curtiss K.; Tallahassee, Nash 2329; Monticello, Combs 334, 337; Quincy, Combs 399; Chipley, Combs 683. Kentucky: Pine Mountain, Kear- ney 258; without locality, Short. TENNESSEE: Franklin County, Fig. 358.—Distribution of P. boscii. Eggert 26 in 1897; Hiwassee Gorge, Kearney 325; White Cliff Springs, Scribner in 1890; Knox County, Kearney in 1894. AvaBAMA: Northern Alabama, Vasey in 1878; Scottsboro, Chase 4501; Pisgah to Tennessee River, Chase 4491; Auburn, Hitchcock 1320; Montgomery, Carver 64; Dadeville, Pollard & Maxon 136; Talladega Springs, Pollard & Maxon 247; Thomasville, Mohr in 1888. Mississippi: Meridian, Tracy 3252; Starkville, Tracy 3 in 1888; Jackson, Hitch- cock 1300. ArKansas: Benton County, Plank 7. LovistAna: Calhoun, Hitchcock 1272; Shreveport, Hitchcock 1244. Texas: Monterey, Reverchon 4144 b; Houston, Hall 828 in part; without locality, Nealley in 1889. OKLAHOMA: Without locality, Butler 43 (Mo. Bot. Gard. Herb.). 191la. Panicum boscii molle (Vasey) Hitchc. & Chase. Panicum latifolium molle Vasey U. 8. Dept. Agr. Div. Bot. Bull. 8: 34. 1889. ‘Virginia to Louisiana.’’ The type, in the National Herbarium, consists of two vernal plants, with softly villousculmsand sheaths, bearded nodes, and nearly mature primary panicles. The accompanying label in Vasey’s hand reads: ‘‘ Panicum latifolium var. molle Vasey, a state downy all over, Washington, D. C.,¢ L. F. Ward.”’ Panicum walteri molle Porter, Bull. Torrey Club 20: 194. 1893. Presumably based on P. latifolium molle Vasey, since ‘‘(Vasey)’’ is given as authority, though Vasey’s combination is not cited. a “Washington, D. C.,’’ was commonly given as locality for the surrounding region, and the specimens may have been collected on the Virginia side of the Potomac as indicated in the range given. 320 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Panicum pubifolium Nash, Bull. Torrey Club 26: 577. 1899. “‘P. latifolium var. molle Vasey, Contr. U.S. Nat. Herb. 3: 33. 1892. Not P. molle Sw. 1788,” is cited and a description follows. As no type is cited this is evidently primarily a change of name, based on P. latifolium molle Vasey. Panicum boscii molle Hitche. & Chase in Robinson, Rhodora 10: 64.1908. Based on ‘‘P. lati- folium var. molle Vasey.”’ DESCRIPTION. Closely resembling P. boscii and often scarcely to be distin- guished from it; culms on the average not so tall, downy-villous; sheaths rather sparsely villous; blades velvety on the under sur- face, sparsely appressed - pubes- 1G. 359.—P, boscii molle. From type specimen. cent on the upper; panicle axis and branches puberulent and somewhat pilose; spikelets rather more strongly pubescent than in the species. The Florida specimens, together with Biltmore Herbarium 5185b, Dewey 73 and Steele in 1896, have spikelets mostly larger than 4.5 mm. long, and correspond with the varia- tion of the species which includes P. latifolium australe Vasey. DISTRIBUTION. Woods, Connecticut to Illinois and Arkansas, south to Florida and Louisiana. Connecticut: Norwalk, Bissell 5551. New York: Long Island, Bicknell in 1903, 1904, and 1905. New Jersey: Great Notch, Nash in 1893. PENNSYLVANIA: Germantown, Stone in 1889; Easton, Porter in 1891, 1895, and 1898; Westchester, Windle in 1904; Haines Station, Heller 4779. Onto: Cincinnati, James. Intrno1s: Union County, French in 1872. Missourt: Monteer, Bush 754; Noel, Bush 5060. DetawaRE: Newark, Hitchcock 592. District or CotumBiA: Ball 703, Hitchcock 435, Merrill 202, 203, Fic. 360.—Distribution of P. boscii molle. Pieters in 1896, Pollard 412, Seaman in 1873, Steele in 1896, Vaseyin 1873 and 1884, Wardin 1881 and 1882. Vireinta: Glencarlyn, Dewey 73, 92; Four-Mile Run, Hitchcock 434, 436; Bel- field, Meynke in 1904; Portsmouth, Noyes 104. Nortu Carourna: Asheville, McCarthy in 1888; Biltmore, Biltmore Herb. 5185a, 5185b; Chapel Hill, Chase 3054. Souta Caro.uina: Newry, House 2211. Grorata: Stone Mountain, Hitchcock 1344; Gwinnett County, Small in 1893; Cobb County, Wilson 27; Athens, Harper 15; Warm Springs, Tracy 8868; Augusta, Cuthbert 392, 1162. Froriwa: Lake City, Hitchcock 10103, Rolfs 817; Chattahoochee, Curtiss 6047 (Hitchcock Herb.); Madison, Combs 295; Gainesville, Chase 4208, Combs 738. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 321 Kentucky: Irvine, Biltmore Herb. 9959f (Biltmore Herb.). TENNESSEE: Knoxville, Ruth 70, Scribner in 1892; Madison County, Bain 189. AuaBAMA: Auburn, Hitchcock 1321. Mississippi: Fairport, Tracy 3205; Jackson, Hitchcock 1301; Starkville, Tracy in Pollard Dist. Miss. Pl. 1409. ARKANSAS: Benton County, Plank 16, 46. Louisiana: Covington, Langlois 41 in part; Calhoun, Hitchcock 1283. MISCELLANEOUS SPECIES. 192. Panicum obtusum H. B. K. Panicum obtusum H. B. K. Nov. Gen. & Sp. 1: 98. 1815. ‘‘Crescit in planitie montana regni Mexicani prope Guanaruato et Burras, in humidis, alt. 1080 hexap.”’ The type specimen, in the Bonpland Herbarium, is labeled: ‘‘Panicum obtusum Kunth, Synops. 174, in planitie montana Regni Mexicani, prope Guanaxuato, 1080 hex. No. 4204.” Panicum polygonoides C. Muell. Bot. Zeit. 19: 323. 1861. ‘‘America septentrio- nalis, ubi forsan in Texas legit T. Drummond (Coll. No. 371).’’ The type specimen, bearing the published data, is in the Berlin Herbarium. Panicum repente Buckl. Prel. Rep. Geol. Agr. Surv. Tex. App. 3. 1866. No speci- men nor locality within Texas is cited. The type specimen could not be found in the herbarium of the Philadelphia Academy, where the Buckley collections are deposited. The description amply identifies the species. Brachiaria obtusa Nash in Britton, Man. 77. 1901. Based on Panicum obtusum H. B. K. In this species the spikelets are placed with the back of the fruit to the axis (that is the first glume turned from the axis) as characteristic of true Panicum, not in the reverse position which characterizes Brachiaria.¢ DESCRIPTION. Plants perennial, usually tufted from a more or less knotted rootstock, and produc- ing widely creeping stolons, sometimes 2 or more meters long, with long internodes, and geniculate, swollen, conspicuously villous nodes, these often with a knob-like cluster of hairy scales at the base of the extra- vaginal, erect branches, these clusters being pro- duced sometimes when the branch is not developed; culms wiry, compressed, 20 to 80 cm. high, simple, usually decumbent at base, glabrous, the nodes glabrous; sheaths shorter than the internodes, glab- rous, or the lower and those of the stolons some- times villous; ligules membranaceous, about 1 mm. long; blades 3 to 20 cm. long, 2 to7 mm. wide, erect, firm, usually involute-setaceous toward the tip, glabrous on both surfaces or sometimes with a few long hairs on the upper surface at the base; pani- cle usually short-exserted, 3 to 12 cm. long, about 1 cm. wide, the few, appressed, raceme-like branches densely flowered; spikelets short-pediceled along one side of a slightly flattened rachis, 3 to 3.8 mm. long, 1.5 to 1.8 mm. wide, and about 2 mm. thick, obovoid, blunt, glabrous, usually brownish; first glume nearly FG. 361.—P. ohtusum. From type specimen. 'aThe genus Brachiaria Ledeb. (Fl. Ross. 4: 469. 1853) is based upon Panicum eruciformis Sibth., in which the spikelets are placed with the back of the fruit turned from the rachis. 41616°—vot 15—10——21 822 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. as long as the spikelet, 5-nerved; second glume and sterile lemma subequal, 7 to 9-nerved, the lemma subtending a rather firm palea and a staminate flower; fruit 3 to 3.5 mm. long, 1.5 to 1.7 mm. wide, subacute, smooth and shining, but very obscurely pubescent at the apex. The Brazilian species, P. repandum Nees, is the only known species related to P. obtusum. DISTRIBUTION. Sandy or gravelly soil, mostly along the banks of rivers, arroyos, and irrigation ditches, western Missouri and Colorado to Texas and Arizona and southward to south- ern Mexico. Missouri: Kansas City, Bush 1832, 3107 (Gray Herb.). Kansas: Stanton County, Hitchcock Pl. Kan. 572. Texas: Dallas, Hall 827, Reverchon 1079 and in 1879; Kerrville, Heller 1741, Smith in 1897; Waller County, Thurow in 1898 and 1906; Llano, Plank in 1892; Amarillo, Ball 1139; Fort Worth, Ward in 1877; Abilene, Tracy 7935; Bexar County, Jermy 6; San Antonio, Plank in 1893; El Paso, Jones 4168; Fort Davis, Havard in 1881; Kingsville, Piper in 1906; Texline, Griffiths 5612; without locality, Buckley in 1881; Nealley in 1887. OxLAHOMA: On the False Washita, Palmer 370 in 1868. Cotorapo: Rocky Ford, Griffiths 3309; Canyon City, Shear 975; Trinidad, Shear in 1900; Las Animas County, Chase 5406. New Mexico: Cabra Spring, Pease in 1878; McCarty, Munson & Hopkins in 1889; Socorro, Plank 76, Vasey in 1881; Gray, Skehan 94, 97; Cimarron Canyon Griffiths 5542; Roswell,. Karle 301; Las Cruces, Wooton 1068; Silver City, Metcalfe 749; Mesilla, Wooton 64; Mesilla Park, Hitchcock 3830; Deming, Hitchcock 3763; Grant County, Blumer 205, Smith in 1897; White Water, Mearns 2308; without locality, Vasey in 1881, Wright 2092. Arizona: Moki Reservation, Hough 80; Beaver Creek, MacDougal 569, Rusby 864, 8921; Toumey in 1891; Santa Rita Mountains, Griffiths 3405, 6959, 7288, Gri/- fiths & Thornber in 1902; St. Johns, Griffiths 5196; Holbrook, Zuck in 1896; Tucson, Griffiths 1514, 1546; Benson, Griffiths 2006, Hitchcock 3737; Pearce, Griffiths 1935; Sulphur Spring Valley, Forbes 1645; Patagonia, Hitchcock 3646, 3661; Fort Huachuca, Wilcox in 1894; San Pedro River, Mearns 1130; Bisbee, Mearns 925; San Bernardino Ranch, Mearns 773; near Monument 88 (Mexi- can Boundary), Mearns 1845. Mexico: La Ventura, Nelson 3908; Saltillo, Palmer 394 in 1898; Chihuahua, Nelson 6352, Pringle 476; Nogales, Griffiths 6800; Durango, Palmer 175 in 1896; Conception del Oro, Palmer 266 in 1904; San Luis Potosf, Palmer 590 in 1898, Parry & Palmer 960, Schaffner 148; Faral, Schumann 1714. Fig. 362.—Distribution of P. obtuswm. 193. Panicum hemitomon Schult. Panicum walterit Ell. Bot. S. C. & Ga. 1: 115. 1816, not Pursh, 1814. ‘‘ Grows in damp soils. On Charleston neck, common. Macleod’s pond, 64 miles from Savan- nah, on the Ogechee road.’’ The type, in the Elliott Herbarium, consists of the upper portions of two culms, one sterile, the other with animmature panicle. Attached to this specimen is a label which reads: ‘‘Panicum Walteri mihi. Hab. in humidis HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 323 circa stagnum, 643 a Sav: versus Oquechee. Flor: Ma. 478.’’ Elliott gives ‘ from Paraguay, which is distinguished from this by the smaller panicles, pubescent spikelets, and a more pronounced crest to the fertile lemma. DISTRIBUTION, Woods and copses, Mexico, West Indies, and south to Paraguay. Mexico: Ocuilopa, Nelson 3023; Trapiche de la Concepcion, Liebmann 394; San Juan Bautista, Rovirosa 624. GuatreMALA: Dept. Alta Vera Paz, Tuerckheim 7699, 7700, 8785, 8796. Costa Rica: La Florida, Pittier 11276; Talamanca, Tonduz 8566; San Rafael, Pittier 2598. a For a further discussion see Hitchcock, Contr. Nat. Herb. 12: 118. 1908. One of the sheets upon which Linnzeus has written the name ‘‘latifolium”’ is P. zizanioides. But this was received from Browne in Jamaica after the publication of the first edition of Linnzeus’s Species Plantarum and hence could not be the type of P. latifoliwm. b Ann. N. Y. Acad. Sci. 7: 262. 1893. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 327 Cusa: Laguna Castillano, Baker 4334; Sancti Spiritus, Leén 903; without locality, Wright 3466. Jamaica: Gordon Town, Hart 726; Port Antonio, Maxon 2109. CoLomBIA: Santa Marta, Smith 169. VENEZUELA: Tovar, Fendler 1634 (Gray Herb.). Toxsaao: Eggers 5810. Trintpap: Broadway 2563, Botanic Gardens Herb. 2286, 3188. British Guiana: Jenman 6001. Dourcu Guiana: Surinam, Hostmann (Gray Herb.). Braziu: Santarem, Spruce 706; San Gabriel da Cachoeira, Spruce 2344; Organ Mountains, Wilkes Expl. Exped. 8; Rio Janeiro, Widgren in 1844; without locality, Riedel 960. Paraguay: Morong 536, 1001. Ecuapor: Recreo, Eggers 15422 (Field Mus. Herb.). 196. Panicum gymnocarpon Ell. Panicum gymnocarpon Ell. Bot. S. C. & Ga. 1: 117. 1816. ‘‘Collected near Savannah, by Dr. Baldwin.’’ The type, in the Elliott Herbarium, consists of the upper part of a culm, being a panicle and the uppermost leaf. Panicum monachnoides Desv. Opusc. 86. 1831. ‘Habitat in Brasilio.’’ The type is in the Desvaux Herbarium. The locality given is doubtless an error as is the case with many of Desvaux’s specimens. Panicum drummondii Nees; Steud. Syn. Pl. Glum. 1:63. 1854. ‘‘ Drum[mJond legit in N. Orleans.’’ In the Berlin Herbarium is a specimen labeled ‘‘ Panicum Drum- mondii N. ab E. in Herb. Lindh. New Orleans n. 574,’’ which is probably the type. Phanopyrum gymnocarpum Nash in Small, Fl. Southeast. U. 8. 104. 1903. Based on Panicum gymnocarpon Ell. Rafinesque¢ proposed Phanopyrum as a section of Panicum, including the single species P. gymnocarpon. This section was raised to generic rank by Nash,? the dis- tinguishing characters being the acuminate equal glumes and the short fertile lemma. This species departs somewhat from the usual characters of the genus Panicum, but the divergence does not seem sufficient to justify segregating the single species as the type of a separate genus. DESCRIPTION. Plants perennial, with a succulent, decumbent or creep- ing base, sometimes as much as 2 meters long, rooting at the nodes, glabrous throughout; culms erect or ascending, 60 to 100 cm. high, rather thick and succulent, nodes often dark colored; sheaths shorter than the internodes, sometimes ciliate near the summit, ligules membranaceous, about 1 mm. long, decurrent down the margin of the sheath; blades linear- lanceolate, 20 to 35 em. long, or the upper and lower shorter, usually 15 to 25 mm. wide, flat, scarcely narrowed at the cordate, sparingly ciliate base, narrowed from about the mid- dle to the acute apex, margins very scabrous; panicles finally exserted, 20 to 40 cm. long, about three-fourths as wide, con- sisting of several to many racemes, solitary or fascicled along a-main axis, the racemes stiffly ascending, or somewhat spreading, the middle 8 to 12 em. or oceasionally as much as 18 cm. long, usually spikelet-bearing from base, the spikelets short-pediceled on short, appressed branchlets, thus yee in Fic. 368.—P. gymnocarpon. From type specimen. a Bull. Bot. Seringe 220. 1830. b Small, FI. fonthonst. U. S. 104. 1903. 328 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. somewhat scattered clusters; spikelets 6 to 7 mm. long, about 1.2 mm. wide, and, by the spreading of the glumes, about twice as thick, strongly nerved; first glume nearly as long as the sterile lemma, acuminate-pointed, the second glume exceeding the sterile lemma, both much exceeding the fruit and at maturity spreading and exposing it, acuminate-pointed, the summit of the lemma arcuate, the sterile palea obsolete; fruit 2 mm. long, 1 mm. wide, ob- ovate, stipitate, smooth and shining. DISTRIBUTION. Ditches and muddy banks of streams and lakes, Georgiaand Florida to Texas. Geroraia: Americus, Harper 522. Frorwa: Burnside, Combs 1426; without locality, Chapman, Rugel 599, Simpson in 1890. AuABAMA: Mobile, Mohr in 1887. Fic. 369.—Distribution of P. gymnocarpon. Mississrepr: Saratoga, Tracy 8396. Lovuistana: Plaquemines Parish, Langlois 47, 151; Chalmette, Tracy 7400; Alexandria, Hale; Natchitoches, Ball 157; Lake Charles, Chase 4407. Texas: Harrisburg, Joor in 1875; Mineola, Reverchon 2235; Columbia, Bush 1498; Hempstead, Plank 9; Waller County, Thurow 17, and in 1889. 926A. Panicum decolorans H. B. K. Panicum decolorans H. B. K. Nov. Gen. & Sp. 1: 100. 1815. “‘Crescit in temperatis, apricis regni Mexicani prope Queretaro, alt. 995 hexap.’’ The type specimen from the Bonpland Herbarium in the Paris Herbarium, bearing the published data, consists of two pieces of a culm with flat blades, one with a narrow terminal panicle about 18 cm. long. The spikelets are 5 to 5.2 mm, long. Plants tinged with purple, branching from the base; culms ascending or erect, usually from a geniculate base, simple or bearing simple, usually sterile branches from the lower one or two nodes, strongly striate or almost grooved, glabrous to sparsely papillose-hispidulous, the nodes appressed-pubescent or glabrous; sheaths short, but sometimes overlapping on the shortened lower internodes, glabrous to sparsely papillose-hispid, ciliate; ligules membranaceous-fimbriate, hardly 1 mm. long; blades 8 to 15 cm. long, 7 to 10 mm. wide, flat, glabrous on both surfaces or with a few scattered papille, these with or without short, stiff hairs; panicles finally exserted, 10 to 18 cm. long, usually not more than one-third, but sometimes as much as two-thirds as wide, the rather long branches usually narrowly ascending, the short-pediceled spikelets somewhat crowded on approximate, short, appressed branchlets; spikelets 4.5 to 5.2 mm. long, 1.6 mm. wide, pointed; first glume less than half the length of the spikelet, acute; second elume and sterile lemma subequal, exceeding the fruit and pointed beyond it; fruit 2.8 to 3.2 mm. long, 1.5 mm. wide. Fic. 370,—P, decolorans. From type specimen. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 329 In the original description P. decolorans is given as perennial, but the type lacks the basal portion, while its general character shows its relationship to those species of Capillaria having large spikelets. The pubescence appears to be extremely vari- able even on the same plant. DISTRIBUTION. Fields and waste ground, plateau of central Mexico. Mexico: Querétaro, Hitchcock 5822, Humboldt (Paris Herb.); Cardenas, Hitchcock 5712. DOUBTFUL SPECIES. In the following list are given the names assigned to species of Panicum and credited to North America which have not been accounted for in the preceding pages and which can not definitely be excluded from Panicum as here limited. The list includes several nomina nuda which are mentioned only because the names are given in the Index Kewensis and consequently have become a part of the literature upon the genus. Panicum ambitiosum Fourn. Mex. Pl. 2: 30. 1886. ‘‘Orizaba (THomaAs in herb. Bucuincer); Vera Cruz (herb. Uzac).’’ Wehave not seen the type nor an authentic specimen of this species. From the description it appears to be a species of Ichnan- thus. The name was given by Hemsley as a nomen nudum.4 Panicum arundinariae Trin.; Fourn. Mex. Pl. 2: 25. 1886. Fournier credits this name to ‘‘Trin. in sched. coll. Schiedeanae” and cites as the first of several speci- mens, ‘‘Absque loco (ScHrEDE).’’ We have not seen the Schiede specimen, which is the type. Schaffner’s no. 279, cited by Fournier, which agrees fairly well with his description, is P. virgultorum. Trinius’s name is mentioned earlier as a nomen nudum by Steudel® and Hemsley.¢ Panicum brevifolium Walt. F1. Carol. 73. 1788. No particular locality is given by Walter but his plants were all collected in the valley of the lower Santee River, South Carolina. The author evidently intended to refer his species to P. brevifolium L., as he quotes the Linnzean diagnosis. We may thus consider that Walter misapplied the name. What species Walter had, which he referred to P. brevifolium, is uncertain. Panicum buchingeri Fourn. Mex. Pl. 2: 30. 1886. ‘‘Orizaba (THomas in herb. BucuinGer).’’ We have not seen the type. From the description it appears to be P. virgatum L. The name is listed earlier by Hemsley ¢ without description. Panicum cartilaginewm Muhl. Descr. Gram. 128. 1817. ‘‘Habitat in Georgia.’’ This isnot in the Muhlenberg Herbarium. The description suggests P. leucothriz Nash. Panicum conchatum Fourn. Mex. Pl. 2: 25.1886. ‘‘Sierra de San Cristobal(ScHaFFn. n. 204, octobri).’’ This name was earlier listed by Hemsley ¢ without description. Specimens of this number of Shaffner’s collection were examined at the herbarium of Drake de Castillo and at the Halle Herbarium, but the notes taken at the time are not full enough to enable us to describe this species, which is not represented in the National Herbarium, nor is Fournier’s description at all adequate. This appears to be a distinct species of the Parviglumia and most nearly related to P. schmitzii Hack., from which it differs in having a longer, more oblong spikelet with a longer first glume. Panicum cordifolium Desv. Opusc. 88 [90]. 1831. ‘‘Habitat in America boreali.’”’ We have not seen the type. The description applies well to P. commutatum Schult. Panicum cordovense Fourn. Mex. Pl. 2: 26. 1886. ‘‘ Cordova (ScHarrn. n. 293 in herb. Franq.).’’ The type is in the herbarium of Drake de Castillo. The notes taken upon this specimen do not enable us to identify it with any other Mexican species, hence it is retained among the doubtful species until more material can be _ — 4 Biol. Centr. Amer. Bot. 3: 485. 1885. ¢ Biol. Centr. Amer. Pot. 3: 486. 1885. b Nom. Bot. ed. 2. 2: 253. 1841. d Op. cit. 487. 830 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. examined. The culms and blades are minutely pubescent, the latter 8 cm. long and 12mm. wide. The spikelets are glabrous and 3 mm. long, the first glume obtuse, 3-nerved, nearly as long as the spikelet, the second glume 5-nerved, the sterile lemma 2-nerved, the central nerve being suppressed. Panicum densum Muhl. Descr. Gram. 122. 1817. No locality is given. The type is not in the Muhlenberg Herbarium. The description suggests one of the Lanuginosa. Panicum dichotomum curvatum Torr. Fl. North. & Mid. U. 8. 145. 1824. No locality is given. There is no specimen thus marked in the Torrey Herbarium. The refer- ence to tall culms and curved branches suggests P. yadkinense Ashe. Panicum dichotomum gracile Torr. Fl. North. & Mid. U.S. 145. 1824. ‘‘Common in swamps, New-York.’’ The type is not in the Torrey Herbarium and the form can not be certainly identified, but the description, ‘‘culm tall, slender; leaves mem- branaceous. Common in swamps,’’ suggests the vernal form of P. lucidum Ashe. Panicum dichotomum spathaceum Wood, Bot. & Flor. 393. 1874. No locality nor specimen is mentioned and the form can not be identified. Panicum disciferum Fourn. Mex. Pl. 2: 19. 1886. ‘“‘San Luis de Potosi (Virt[Eet} n. 1292).’’ We have been unable to find the type of this. The name was mentioned earlier, without description, by Hemsley. The description of the inflorescence as consisting of six appressed spikes suggest that this may not be a true Panicum. Panicum discolor Spreng. Mant. Fl. Hal. 31. 1807. The only specimen mentioned is ‘‘E Pensylvania.’’ We have not been able to locate the type and the species can not be identified from the description, though it belongs to the subgenus Dichanthelium. Panicum elliottii Spreng.; Steud. Nom. Bot. ed. 2. 2: 256. 1841, not Trin. 1829. This is given as a synonym of P. pubescens. As the latter name is given without an author, Sprengel’s name can be fixed upon no definite species and is hence a nomen nudum. No type has been seen. Panicum firmandum Steud. Syn. Pl. Glum. 1: 418. 1854. ‘‘Carolina sptr.’’ This was received from M. A. Curtis under the name of P. microcarpon Muhl. We have not seen the type. The description applies well to P. sphaerocarpon except that the spikelets are said to be glabrous while in the latter species they are finely puberulent. Panicum fleruosum Raf.; Desv. Journ. de Bot. 4: 273. 1814, not Retz. 1791. The description is as follows: ‘‘feuilles lancéolées, élatées, ciliees 4 la base; panicule pubescente; pedoncules flexueux; glume ciliee. Dans le nouveau Jersey.” Panicum gracilescens Desv.; Poir. in Lam. Encycl. Suppl. 4: 279. 1816. ‘“‘Cette plante croft 4 la Caroline (V. s. in herb. Desv.).’’ The type could not be found in the Desvaux Herbarium and the species can not be identified. Desvaux gives a later description,® which disagrees in some respects with that of Poiret. Panicum hirsutum Vahl; Griseb. Fl. Brit. W. Ind. 548. 1864, not Swartz, 1797. This is mentioned as a synonym under P..diffusum Swartz. We have not seen the type but the specific name and the statement by Grisebach that the sheaths of P. diffusum may - be glabrous or pilose, suggest that P. hirsutum Vahl may be P. ghiesbreghtii Fourn. Panicum iowense Ashe, N. C. Agr. Exp. Sta. Bull. 175: 115. 1900. ‘‘Dry prairies, eastern Iowa to Kansas, June and July.’’ There is no specimen in Ashe’s herbarium bearing this name nor that can with any degree of certainty be connected with the description. There is a specimen of P. praecocius collected on dry prairies at Arm- strong, Iowa, July, 1890, by R. I. Cratty, which is marked in pencil by Ashe, *‘ Pani- cum prairie.’’ The description applies fairly well to this specimen except that the height is given as ‘‘1—-2 cm.”’ [error for 1 to 2 dm.?], the panicle as ‘‘small, 1-2 cm. long,’’ and the length of the spikelets as ‘‘1.1-1.4 mm.’ [the spikelets are 1.8 to 1.9 mm. long]. The description is too meagre to distinguish this species from P. hua- chucae which is found in the range given, and to which the description of panicles and spikelets better applies. A specimen of P. praecocius, Carver 258, Jewell Junction, Iowa, in the Iowa Agricultural College Herbarium, is marked in Ashe’s writing ‘‘Panicum haemocarpon Ashe sp. noy. affine P. villosissimum Nash.’”’ This adds to the evidence against taking the Cratty specimen as the type of P. iowense. a Biol. Centr. Amer. Bot. 3: 488. 1885. b Opusc. 95. 1831. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 331 Panicum muhlenbergianum Schult. Mant. 2: 230. 1824. Based on ‘‘ Panicum n. 27 (sine nomine) Muhlenb. Descr. ub. p. 118.’’ The type is not in the Muhlenberg Herbarium, and there is no evidence that Schultes saw the plant, the description of which he copies from Muhlenberg. The latter gives the locality as ‘“‘Habitat in Georgia.’’ The description applies well to P. barbulatum Michx. Panicum nitidum glabrum Torr. Fl. North. & Mid. U. 8. 146. 1824. No locality is mentioned, and the type can not be found in the Torrey Herbarium. The descrip- tion suggests P. commutatum Schult. Panicum nitidum gracile Torr. Fl. North. & Mid. U.S. 146. 1824. The only locality mentioned is ‘‘near New-York.’’ The type can not be found in the Torrey Herba- rium. The description applies fairly well to the vernal form of P. dichotomum L. Panicum nitidum major{us] Vasey, Contr. Nat. Herb. 3: 30. 1892. No specimen nor locality is cited and no type can be found in the National Herbarium. Vasey says, ‘‘Here could be placed several variable forms.”’ Panicum ornatum Desv.; Hamilt. Prodr. Pl. Ind. Occ. 11. 1825. ‘‘Herb. Prof. Desy. Porto Rico.’’ This is further described by Desvaux.¢ We have been unable to find the type of this species. It is doubtful if the type came from Porto Rico, as the description does not appear to apply to any of the West Indian species. Panicum pensylvanicum Spreng. Bot. Gart. Halle, Erst. Nacht. 30. 1801. ‘‘ Pensyl- vanien.’’ No type nor authentic specimen of this could be found. The species can not be recognized from the description given. A later description? differs in several essentials and may apply to P. anceps. Willdenow¢ refers P. pensylvanicum “Spreng. cat. hort. halens.’’ to P. rostratum Muhl. (P. anceps). Panicum portoricense Desv.; Hamilt. Prodr. Pl. Ind. Occ. 11. 1825. ‘Herb. Prof. Desv. Porto Rico.’’ A further description is given later by Desvaux,@ where the locality is given as ‘“‘Antillis.”” We have been unable to find the type of this. The description appears to apply to a species of the subgenus Dichanthelium. Panicum pumilum Raf. Med. Repos. N. Y. 5: 353. 1808. This is a nomen nudum with no mention of locality nor specimens. Panicum rafinesquianum Schult. Mant. 2: 257. 1824. Based on P. flexuosum Raf. Panicum reflecopilum Steud. Syn. Pl. Glum. 1: 84. 1854. ‘‘Oaxaca.’’ We have not seen the type. The description applies well to P. viscidellum Scribn. Panicum sessilicaule Desv.; Hamilt. Prodr. Pl. Ind. Oce. 11. 1825. This is men- tioned in a note under P. cayennense Lam. and later described by Desvaux, ¢ who gives the locality as ‘‘Habitat in Carolina?” and states that the plant is only a variety of P. cayennense. It probably did not come from Carolina. We have not seen the type. Panicum speciosum Walt. F\. Carol. 73.1788. No special locality is given, but the general locality is the valley of the lower Santee River, South Carolina. There is no specimen of this in Walter’s herbarium. It can not be identified. Panicum striatum Muhl.; Hemsl. Biol. Centr. Amer. Bot. 3: 492. 1885. This is given as a synonym of P. neuranthum Griseb. The name appears here and also later in Fournier’s work,/ from which Hemsley quotes as ‘‘Panicum striatum Muhl. not Lam.’’ We know of no P. striatum of Muhlenberg. Panicum vilfiforme Wood, Class-book. 785. 1861. ‘‘Wet meadows, E. Tenn.!’’ We have not been able to locate the type of this, and it can not be identified from the description. It appears to be a species of the group Agrostoidia. Panicum virletii Fourn. Mex. Pl. 2: 29. 1886. ‘‘San Luis de Potosi (Vrru. n. 1305, 1371).”’ This name was earlier listed without description by Hemsley.g Neither of the specimens cited could be found. The only description given is ‘‘Differt a P. diffuso gluma inferiore mediam spiculam superante.’’ The specimens cited by Fournier under P. diffusum are mostly P. ghiesbreghtii Fourn. . @Opusc. 88 [90]. 1831. ¢ Opusc. 95. 1831. b Spreng. Mant. Fl. Hal. 1: 31. 1807. J Mex. Pl. 2: 19. 1886. e Enum. PI. 1032. 1809. 9 Biol. Centr. Amer. Bot. 3: 498. 1885. @ Opusc. 89. 1831. LIST OF NEW SUBGENERA AND SPECIES AND NEW NAMES. Page. Dichanthelium Hitchc. & Chase, subgen. nov. ers P.. dichotomum L.)...... 142 Hasiavis (Grisen.) Hitch. 2....-.----+.--+----- : a2 lo oe 16 Lasiacis sect. Panicum Griseb. asia divaricata (1y.) HMitche..... 2... pe ese sees eee 16 Panicum divaricatum L. Panicum amarulum Hitchc. & Chase, sp. nov.........-.---.---+-----+sse00s 96 Panicum breve Hitchc. & Chase, sp. nov.-.....-<-.--..-.- «ss «42a 271 Panicum bulbosum sciaphilum (Rupr.) Hitchc. & Chase..........-------.-- 83 Panicum sciaphilum Rupr. Panicum concinnius Hitche. & Chase...-.......-..:0¢2--<=2+=20905— =n 263 Panicum gracilicaule Nash, 1903, not Rendle, 1899. Panicuni cuprewm Hitche. & Chase...........-...0 Js -ss+2+2 sc 55 eee 120 Panicum hians purpurascens Scribn., not P. purpurascens H. B. K. Panicum firmulum Hitchc. & Chase, sp. nov..........---.------+--sssseces 27 Panicum languidum Hitche. & Chase........0...)....25 .s--2: eee eee 232 Panicum unciphyllum forma prostratum Scribn. & Merr., not P. pros- tratum Lam. Panicum lepidulum Hitchc. & Chase, sp. nov.......-....---------+--++-+-+-+- 75 Panicum lonpuni@#itche. & Chase:..........00.5....0.¢2. 2.32 111 Panicum pilosum macranthum Scribn., not P. macranthum Trin. Panicum nodatum Hitchc. & Chase, sp. nov.......- 1.2.0!) 2 293 Panicum olivaceum Hitchc. & Chase, sp. nov..............--------+---+++: 225 Panicum pacificum Hitchc. & Chase, sp. nov.......-.-...-----------+--e 229 Panicum paludivagum Hitchc. & Chase, sp. nov........-.----------------+- 32 Panicum pampinosum Hitchc. & Chase, sp. nov.......--------------------- 66 Panicum parcum. Hitche. & Chase, sp. Nov...........-:.---<--+: 4:-se=enees 68 Panicum plenum Hitchc. & Chase, sp. nov.........---.--~------<-2--=ss=s6 80 Panicum rhizomatum Hitchc. & Chase, sp. nov.......-.....-----------+--- 109 Panicum rotundum Hitchce. & Chase, sp. nov.....-....---«-=-----=- sasseeme 139 Panicum sphaerocarpon inflatum (Scribn. & Smith) Hitche.........-...-- 253 Panicum inflatum Scribn. & Smith. Panicum stramineum Hitchc. & Chase, sp. nov.......--.----.----+-+------- 67 Panicum vernale Hitchc. & Chase, sp. nov...-.--..-4..0s:-4-:5sseeeemeene 266 Panicum xalapense strictirameum Hitchc. & Chase, subsp. nov........ -- 161 Paurochaetium Hitchc. & Chase, subgen. nov. (type P. distantiflorum Rich.). 22 332 INDEX TO NUMBERED SPECIMENS. The following list includes the numbered specimens distributed in the more important collections of Panicum. Two or more species when here listed under one number were distributed under this number by the collector. ANDREWS, L. Batu, C. R. 8. sphaerocarpon. 8. anceps. ll. clandestinum. 14. boscii. 14. lindheimeri. 20. agrostoides. 18. barbulatum. 21. dichotomiflorum., 20. implicatum. polyanthes, 23. verrucosum. 25. joorii. 36. huachucae silvicola. 30. perlongum. 40. boscii. 34. dichotomiflorum. 44. latifolium. 39. dichotomiflorum. 49. linearifolium. 40. latifolium. 51. scribnerianum. 42. huachucae silvicola. 58. depauperatum. 46. anceps. 62. tsugetorum. 51. microcarpon. 63. ashei. 53. lindheimeri. 64. huachucae silvicola. 60. hians. 66. huachucae silvicola. 62. xalapense. 70. huachucae. 64. oligosanthes. 73. sphaerocarpon. 65. sphaerocarpon. 74. tsugetorum. 66. commutatum. Baker, C. F. 67. dichotomiflorum, 36. scribnerianum. 69. sphaerocarpon. 48. barbinode. 70. stipitatum. 131. agrostoides. 77. scoparium. 676. barbipulvinatum. 83. agrostoides. 2053. barbinode. 111. agrostoides. 2078. parvifolium. 120. capillare. 4334. zizanioides. 121. angustifolium. 4837. stenodes. 122. xalapense. . BAKER, ©. H. 123. hians. 18. condensum. 124. agrostoides. 31. verrucosum. 125. anceps. 40. rhizomatum. 135. dichotomiflorum. 41. pauciciliatum. 137. rhizomatum. 42. commutatum. 139. dichotomiflorum. 45. chrysopsidifolium. 145. perlongum. 68. polycaulon. 149. capillare. 69. equilaterale. 155. latifolium. 70. pauciciliatum. 156. agrostoides. 71. pauciciliatum. 157. gymmnocarpon. 72. pauciciliatum. huachucae silvicola. 119. albomarginatum. 164. dichotomiflorum. 333 334 CONTRIBUTIONS FROM Batu, C. R.—Continued. 166. anceps. 169. agrostoides. 178. scribnerianum. 190. hians. 192. virgatum. 202. lucidum. 204. verrucosum. 215. dichotomiflorum. 220. brachyanthum. 225. agrostoides. 228. anceps. 229. dichotomiflorum. 441. sphaerocarpon inflatum. 535. hians. 536. sphaerocarpon inflatum. 544. lanuginosum. 625. commutatum. 702. clandestinum. 703. boscii molle. 704. microcarpon. 705. polyanthes. 720. capillare. 791. equilaterale. 792. lancearium. 816. tennesseense. 817. implicatum. 966. capillare. 1139. obtusum. BartueEtTT, H. H. 807. spretum. 844. microcarpon. 903. anceps. 1024. philadelphicum. 1066. agrostoides. 1071. polyanthes. 1081. microcarpon. 1103. verrucosum. 1436. lucidum. 1170. scoparium. 1172. auburne. 1173. nitidum. 1327. oricola. 1368. oricola. 1379. oricola. 1414. villosissimum. 1443. lanuginosum. 1444. aciculare. 1453. xalapense. 1455. aciculare. 1456. nitidum. 1457. microcarpon. 1458. ashei. THE NATIONAL Barter, H. 1459. 1460. 1461. 1462. 1497. 1498. 1500. 1501. 1502. 1504. 1505. 1508. Bess, R. 5138. 514. 520. 882. 1236. 1242. 1245. 1259. 1262. 1276. 1278. 1299. 1321. 1426. 1428. 1430. 1434. 1457. 1459. 2057. 2167. 2661. 2663. 2670. 2703. 2715: 2815. 2832. 2833. 28334. 2834. 2881. 2882. 2917. 2928. 2935. 2936. 29394. 2947. HERBARIUM. H.—Continued. sphaerocarpon. lanuginosum. concinnius. angustifolium. villosissimum. polyanthes. polyanthes. joorii. ravenelii. ashei. boscii. mutabile. depauperatum. pseudopubescens. huachucae. latifolium. xalapense. commutatum. oligosanthes. sphaerocarpon. angustifolium. lindheimeri. helleri. capillare. reverchoni. oligosanthes. lindheimeri. helleri. sphaerocarpon. xalapense. scribnerianum. praecocius. scribnerianum. villosissimum. depauperatum. malacophyllum. oligosanthes. scribnerianum. meridionale. subvillosum. villosissimum. subvillosum. villosissimum. tennesseense. pseudopubescens. capillare. flexile. boreale. meridionale. tsugetorum. meridionale. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. BirTMORE HERBARIUM. 696a. 697a. 698b. 700a. 700b. 700c. 700e. 702a. 797. 797a. 797c. 800a. 800b. 800c. 802a. 802b. 803a. 804. 804b. 808a. 2984a. 2993a. 2993c. 2994a. 3627. 4278. 4290. 4290a. 4292b. 5066b. 5066e. 5184b. 5185a. 5185b. 5839b. 6022a. 6028. 6204a. 7079a. 7080a. 8342. 9953c. 9959f. 10715b. 11866. 14879b. BissE Lt, C. H. 385. 5529. 5533. 5541. anceps. trifolium. tennesseense. virgatum. virgatum. virgatum. virgatum cubense. dichotomiflorum. depauperatum. depauperatum. depauperatum. barbulatum. dichotomum. lucidum. dichotomum. commutatum. commutatum. microcarpon. clandestinum. clandestinum. stipitatum. commutatum. xalapense. xalapense. ravenelii. longifolium. angustifolium. scoparium. scoparium. sphaerocarpon. lucidum. sphagnicola. tennesseense. boscii molle. boscii molle. anceps. polycaulon. spretum in part. webberianum. linearifolium. wrightianum. werneri. albemarlense. boscii molle. curtifolium. aciculare. huachucae silvicola. scoparioides. agrostoides. depauperatum. linearifolium. ‘| Bissett, C. H.—Continued. 5542. 5544. 5546. 5548. 5549. 5550. 5551. D002. 5563. 5566. 5569. 5570. 5571. 5977. 5580. 5581. 5582. 5583. 5585. 5590. 5594. 5595. 5596. 5611. 5616. 5622. 8084. 9306. 11596. 12000. 12001. 12002. Borrert, M. 99. 101. 160. 648. 688. 705. 1987. BourGEAU, E. 235. 529. 1455. 1662. 2162. 2383. 2751. 2794. 3132. 3192. linearifolium. clandestinum. clandestinum. latifolium. latifolium. latifolium. boscii molle. boscii. scribnerianum. microcarpon. microcarpon. microcarpon. microcarpon. barbulatum. ashei. scoparioides. boreale. lindheimeri. sphaerocarpon. implicatum. tsugetorum. tsugetorum. columbianum. tennesseense. tsugetorum. implicatum. scoparioides. oricola. longifolium. commonsianum. subvillosum. implicatum. olivaceum. olivaceum. plenum. virgatum. laxum. viscidellum. olivaceum. bulbosum. elephantipes. pulchellum. polygonatum in part. xalapense. multirameum. olivaceum. ghiesbreghtii. bulbosum. viscidellum. glutinosum, 335 336 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Brace, L. J. K. 3467. bartowense. 3524. caerulescens. 3697. nitidum. 3742. dichotomiflorum. 4158. exiguiflorum. 4164. exiguiflorum. 4380. exiguiflorum. 7015. caerulescens. 7019. tenerum. 7132. tenerum. Broapway, W. E. 2370. stoloniferum. 2371. frondescens. 2372. parvifolium. 2563. zizanioides. 2629. hirsutum. BuRCHELL, W. J. Al101-2. rotundum. 4146. millegrana. 4315-2. millegrana. 4653. millegrana. 7062. trichanthum. 8350. cayennense. 8706. trichoides. 8791. trichanthum. Busu, B. F. 8. latifolium. 9. agrostoides. 10. dichotomiflorum. 14. villosissimum. 17. ashei. 26. boscil. 51. barbulatum. 54. praecocius. 87. praecocius. 91. latifolium. 120. hians. 141. villosissimum. 145. sphaerocarpon. 148. barbulatum. 153. linearifolium. 177. sphaerocarpon. 178. lindheimeri. 190. latifolium. 232. polyanthes. 234. commutatum. 238. commutatum. 251. huachucae silvicola. 255. polyanthes. 263. malacophyllum. 266. reptans. 270. fasciculatum chartagi- hense, Busu, B. F.—Continued. 276. 287. 289. 302. 303. 308. 309. 310. 312. 313. 320. 322. 323. 333. 379. 384. 413. 642. 651. 674. 675. 705. 707. 708. 709. TAL. 712. 718. 720. 722. 729. boscil. commutatum. ashei. agrostoides. boscii. commutatum., huachucae silvicola. boscii. ashei. barbulatum. linearifolium. commutatum. boscii. villosissimum. leibergil. flexile. bicknellii. malacophyllum. xalapense. reverchoni. sphaerocarpon. helleri. virgatum. hians. hians. hians. tennesseense. tennesseense. clandestinum. barbulatum. scribnerianum. scribnerianum. leibergii. commutatum. linearifolium. villosissimum. clandestinum. scribnerianum. huachucae silvicola. scribnerianum. brachyanthum. scribnerianum. linearifolium. leibergil. huachucae silvicola. microcarpon. sphaerocarpon inflatum. commutatum. praecocius. praecocius. xalapense. villosissimum. sphaerocarpon inflatum. HITCHCOCK AND CHASE—-NORTH AMERICAN PANICUM. Buss, B. F.—Continued. 754. 755. 759. 760. 803. 848. 906. 979. 1023. 1107. 1156. 1157. 1198. 1210. 1216. 1218. 1220. 1222. 1224. 1225. 1273. 1296. 1298. 1388. 1398. 1440. 1450. 1498. 1555. 1652. 1659. 1684. 1685. 1710. 1713. 1732. 1832. 2332. 2350. 2488. 2522. 2526. 2529. 2532. 2736. 2760. 2877. 2881a. 2911. 2913. 2926. 41616°—vo1 15—10 boscii molle. commutatum. huachucae silvicola. huachucae silvicola. helleri. flexile. flexile. brachyanthum. agrostoides. linearifolium. filipes. fasciculatum chartagi- nense. texanum. huachucae silvicola. scribnerianum. sphaerocarpon. villosissimum. scribnerianum. commutatum. oligosanthes. xalapense. reptans. anceps. aciculare. agrostoides. xalapense. commutatum. gymnocarpon. linearifolium. leibergii. secribnerianum. leibergii. helleri. helleri. huachucae silvicola. latifolium. obtusum. huachucae. depauperatum. xalapense. arenicoloides. ravenelii. helleri. barbulatum. commutatum. leibergii. barbulatum. linearifolium. ashei. werneri. werneri. 22 337 | Busn, B. F.—Continued. 2996. 3089. 3090. 3107. 3246. 3295. 3318. 3369. 3456. 3456a. 3529. 3658. 3893. 3903. 3914. 3915. 3933. 3935. 3936. 3968. 3977. 3981. 4001. 4002. 4003. 4021. 4024. 4038. 4411. 441la. 4412. 4458. 4473. 4487. 4532. 4533. 4534. 4549. 4568. 4638. 4651. 4652. 4653. 4654. 4684. 4685. 4712. 4714. 4733. 4734. A788. 4803. huachucae silvicola. perlongum. praecocius. obtusum. bicknellii. flexile. capillare. barbulatum. ashei. barbulatum. barbulatum. stipitatum. helleri. helleri. tennesseense. huachucae silvicola. scribnerianum. praecocius. depauperatum. huachucae silvicola. clandestinum. latifolium. latifolium. huachucae silvicola. clandestinum. scribnerianum. huachucae silvicola. virgatum. linearifolium. linearifolium. ashel, barbulatum. barbulatum. ashei. tennesseense. linearifolium. boscii. linearifolium. scribnerianum. xalapense. clandestinum. _ boscii. scribnerianum. depauperatum. tennesseense. commutatum. ashei. ashei. barbulatum. linearifolium. villosissimum. huachucae silvicola, 338 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Busu, B. F.—Continued. 4807. 4824. 4865. 4908. 5023. 5060. 5105. 5114, 5115. 5116. 5119. 5120. 5203. 5234. 5259. CHAMBERLAIN, 298. 336. 51S! 552. 787. 793. 837. CuHasE, A. 867. 889. 918. 1178. 1472. 1474. 1479. 1480. 1540. 1541. 1542. 1548. 1544. 1545. 1546. 1552. 1563. 1602. 1604. 1605. 16053. 1606. 1607. 1633. 1636. 1729. 1824. 1850. 1887. latifolium. gattingeri. clandestinum. boscii. villosissimum., boscii molle. virgatum. anceps. agrostoides. gattingeri. philadelphicum. philadelphicum. philadelphicum. agrostoides. flexile. E. B. depauperatum. boreale. huachucae silvicola. virgatum. subvillosum. boreale. subvillosum. depauperatum. latifolium. verrucosum. virgatum. meridionale. flexile. flexile. capillare. depauperatum. latifolium. pseudopubescens. dichotomum. tsugetorum. villosissimum. implicatum. tsugetorum. villosissimum. tsugetorum. tsugetorum. pseudopubescens. tsugetorum. villosissimum. scribnerianum. miliaceum. virgatum. flexile. latifolium. sphaerocarpon. latifolium. Cuase, A.—Continued. 1904. 1919. 1921. 1947. 2006. 2008. 2284. 2303. 2315. 2316. 2320. 2332. 2333. 2339. 2341. 2345. 2349. 2357. 2366. 2367. 2368. 2370. 2374. 2378. 2379. 2381. 2400. 2401. 2402, 2412. 2428. 2463. 2477. 24774. 2520. 2528. 2585. 2599. 2628. 2673. 2679. 2830. 2849. 2853. 2864. 2865. 2874. 2887. 2905. 2912. 2936. 2946. 2947. tennesseense. dichotomum. tsugetorum. latifolium. flexile. capillare. depauperatum. linearifolium. dichotomum. xalapense. xalapense. sphaerocarpon. spretum. albemarlense. auburne. lancearium. commonsianum. aciculare. clandestinum. microcarpon. polyanthes. microcarpon. boscil. villosissimum. dichotomum. commutatum. huachucae silvicola. sphaerocarpon. depauperatum. sphaerocarpon. meridionale. tennesseense. ashel. bicknellii. aculeatum. virgatum. anceps. philadelphicum. gattingeri. condensum. stipitatum. ashei. tennesseense. yadkinense. depauperatum. scribnerianum. tennesseense. lindheimeri. villosissimum. oligosanthes. aciculare. yadkinense. annulum. Cuase, A.—Continued. 2964. 2985. 2996. 3047. 3048. 3049. 3051. 3052. 3053. 3054. 3055. 3056. 3056}. 3057. 3058. 3059. 3060. 3061. 30614. 3062. 3063. 3064. 3065. 3067. 3068. 3069. 3071. 3072. 3073. 3074. 3075. 3076. 3078. 3081. 3082. 30824. 3084. 3086. 3087. 3088. 3089. 3090. 3092. 3093. 3094. 3095. 3096. 30964. 3097. 30974. 3098. 3099. 3100. yadkinense. lindheimeri. anceps. lindheimeri. dichotomum. huachucae silvicola. ashei. commutatum. depauperatum. boscii molle. villosissimum. villosissimum. dichotomum. xalapense. villosissimum. yadkinense. trifolium. yadkinense. microcarpon. huachucae silvicola. angustifolium. ashei. huachucae silvicola. huachucae silvicola. lanuginosum. huachucae silvicola. villosissimum. yadkinense. boscii. sphaerocarpon. villosissimum. lanuginosum. clandestinum. villosissimum. angustifolium. arenicoloides. aciculare. lanuginosum. angustifolium. ravenelii. sphaerocarpon. barbulatum. bicknellii. spretum. aciculare. consanguineum. wrightianum. ensifolium. ensifolium. aciculare. hians. mattamuskeetense. meridionale. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 339 | Cuase, A.—Continued. 31003. 3101. 3103. 3105. 3106. 3107. 3108. 3109. 3110. 3111. 3112. 3113. 3114. 3115. 3117. 3118. 3120. 3121. 3123. 3125. 3126. 3127. 3128. 3129. 3130. 3131. 3132. 3133. 3134. 3135. 3136. 3137. 3138. 3139. 3141. 3143. 3144. 3145. 3147. 3150. 3154. 3155. 3156. 3157. 3158. 3159. 3160. 3161. 3162. 3163. 3166. 3167. albermarlense. scabriusculum. aciculare. aciculare. albemarlense. lanuginosum. lindheimeri. pseudopubescens. patulum. xalapense. lucidum. lancearium. commutatum. pseudopubescens. xalapense. xalapense. arenicoloides. villosissimum. aciculare. chamaelonche. pauciciliatum. pauciciliatum. pauciciliatum. lancearium. aciculare. ciliatum. auburne. trifolium. sphaerocarpon inflatum. wrightianum. longiligulatum. erectifolium. angustifolium. pauciciliatum. villosissimum. arenicoloides. virgatum cubense. longiligulatum. pseudopubescens. longiligulatum. mutabile. villosissimum. arenicoloides. aciculare. sphaerocarpon inflatum. lucidum. pseudopubescens. commonsianum., pauciciliatum. angustifolium. addisonii. aciculare. 340 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. CuaseE, A.—Continued. 3168. 3169. 3170. 3171. 3172. 3174. 3175. 3176. 3177. 3178. 3179. 3181. 3182. 3183. 3184. 3185. 3187. 3188. 3189. 3190. 3191. 3192. 3193. 3194. 3195. 3196. 3197. 3198. 3199. 3200. 3201. 32014. 3203. 3204. 3205. 3206. 3208. 3210. 3210}. 3211. 3212. 3213. 3214. 3215. 3216. 3217. 3218. 3220. 3221, 3999 3224. 3225. 3226. commutatum. angustifolium. tenue. strigosum. tenue. consanguineum. longiligulatum. ensifolium. ensifolium. roanokense. longiligulatum. lancearium. chamaelonche. tenue. ciliatum. aciculare. microcarpon. barbulatum. mutabile. microcarpon. villosissimum. yadkinense. lancearium. lancearium. wilmingtonense. ensifolium. lucidum. lindheimeri. trifolium. nitidum. albemarlense. lanuginosum. roanokense. microcarpon. aciculare. sphaerocarpon. lanuginosum. aculeatum. microcarpon. lancearium. sphaerocarpon. longiligulatum. ciliatum. aciculare. aciculare. aciculare. pseudopubescens. ashei. lanuginosum. pseudopubescens. lancearium. trifolium. ciliatum, Cuase, A.—Continued. 3227. 3228. 3230. 3232. 3233. 3234. 3235. 3237. 3238. 3239. 3240. 3242. 3244, 3246. 3247. 3248. 3249. 3250. 3253. 3254. 3255. 3256. 3259. 3260. 3262. 3269. 3270. 3272. 3273. 3274. 3275. 3277. 3278. 3283. 3295. 3299. 3300. 3301. $316. 3320. 3321. 3326. 3355. 3363. 3367. 3379. 3382. 3390. 3391. 3392. 3393. 3399. 33994. ensifolium. longifolium. consanguineum. mattamuskeetense. longiligulatum. ensifolium. scabriusculum. sphaerocarpon. trifolium. trifolium. roanokense. clutei. strigosum. pauciciliatum. roanokense. trifolium. angustifolium. angustifolium. microcarpon. huachucae silvicola. lindheimeri. — sphaerocarpon. lindheimeri. tennesseense. microcarpon. tsugetorum. tsugetorum. clandestinum. polyanthes. tennesseense. tennesseense. boreale. subvillosum. depauperatum. xanthophysum. werneri. xanthophysum. tennesseense. depauperatum. subvillosum. tsugetorum. linearifolium. boreale. tsugetorum. latifolium. depauperatum. werneri. tsugetorum. subvillosum. implicatum. linearifolium. tennesseense. tsugetorum., HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Cuase, A.—Continued. 3436. 3437. 3443. 3453. 3454. 3457. 3458. 3459. 3464. 3466. 3469. 34734. 3475. 3484. 3486. 3488. 3489. 3502. 3503. 3505. 3508. 3509. 3513. 3516. 3517. 35174. 3519. 3520. 3521. 3522. 3523. 3524. 3528. 3529. 3530. 3531. 3532. 3533. 3534. 35344. 3535. 3536. 3538. 3539. 3541. 3542. 3543. 3544. 3545. 3546. 3550. 3551. 3553. implicatum. boreale. boreale. subvillosum. implicatum. tennesseense. boreale. implicatum. lindheimeri. meridionale. agrostoides. lindheimeri. lucidum. barbulatum. scoparium. lanuginosum. sphaerocarpon. ashei. tennesseense. lanuginosum. huachucae silvicola. sphaerocarpon. clandestinum. microcarpon. addisonii. commonsianum. columbianum. pseudopubescens. ashei. villosissimum. addisonii. tsugetorum. lindheimeri. agrostoides. longifolium. commonsianum. tsugetorum. columbianum. columbianum thinium. meridionale. ensifolium. leucothrix. addisonii. columbianum. commonsianum. ashei. tsugetorum. commonsianum. clutei. verrucosum. lucidum. spretum. clutei. ! | | | 341 CHaseE, A.—Continued. 3554. 3556. 3557. 3559. 3560. 3562. 3564. 3566. 3569. 3570. 3572. 3573. 3574. 3575. 3576. 3577. 3578. 3579. 3580. 3581. 3583. 3584. 3588. 3589. 3590. 3593. 3595. 3596. 3598. 3599. 3600. 3601. 36014. 3603. 3605. 3606. 3608. 3609. 3611. 3612. 3613. 3616. 3617. 3618. 3620. 3621. 3623. 3631. 3642. 3643. 3653. 3654. 3656. lucidum. leucothrix. ensifolium. columbianum thinium. columbianum thinium. oricola. tsugetorum. pseudopubescens. spretum. commonsianum. lindheimeri. virgatum cubense. addisonii. commonsianum. oricola. columbianum thinium. leucothrix. tsugetorum. huachucae silvicola. oricola. addisonil. commonsianum. columbianum thinium, lindheimeri. clutei. lucidum. addisonii. commonsianum. clutei. lucidum. scoparium. tsugetorum. ashei. addisonii. columbianum thinium, columbianum. tsugetorum. oricola. sphaerocarpon. tsugetorum. columbianum. yadkinense. tennesseense. scribnerianuir barbulatum. commutatum. virgatum. anceps. tsugetorum. barbulatum. villosissimum. commutatum. lancearium. 342 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Cuase, A.—Continued. OCxase, A.—Continued. 3657. dichotomum. 3860. virgatum cubense. 3658. albomarginatum. 3866. ovale. 3660. verrucosum. 3885. polycaulon. 3662. longifolium. 3889. hians. 3663. lanuginosum. 3899. amarulum. 3668. microcarpon. 3900. virgatum. 3674. patulum. 3907. joorii. 3675. villosissimum. 3908. equilaterale. 3676. virgatum. 3915. amarulum. 3677. angustifolium. 3926. chapmani. 3678. commutatum. 3935. amarulum. 3679. ashei. 3946. trifolium. 3680. auburne. 3947. ovale. 3682. aciculare. 3948. joorii. 3683. meridionale. 3953. amarulum. 3686. lindheimeri. 3964. webberianum. 3687. philadelphicum. 3965. chamaelonche. 3704. ashei. 3966. albomarginatum. 3708. annulum. 3967. polycaulon. 3729. lindheimeri. 3970. lancearium. 3739. ensifolium. 3984. rhizomatum. 3743. lucidum. 3986. verrucosum. 3744. mattamuskeetense. 3990. nitidum. 37444. mattamuskeetense. 3991. fusiforme. 3745. albemarlense. 3992. caerulescens. 3747. barbulatum. 4004. lancearium. 3752. tsugetorum. 4005. nitidum. 3757. albemarlense. 4007. bartowense. 3758. barbulatum. 4009. xalapense. 3762. albemarlense. 4014. condensum. 37624. villosissimum. 4015. lancearium. 3767. microcarpon. 4017. nitidum. 3772. yadkinense. 4019. joorii. 3780. ravenelii. 4021. fusiforme. 3783. bicknellii. 4023. laxiflorum. 3787. asheli. 4026. patulum. 3791. mattamuskeetense. 4028. patentifolium. 3793. mattamuskeetense. 4029. patentifolium. 3794. columbianum. 4032. chamaelonche. 3796. ravenelii. 4036. chamaelonche. 3803. polyanthes. 4037. chamaelonche. 3806. columbianum thinium. 4039. sphagnicola. 3808. scribnerianum. 4043. albomarginatum. 3809. annulum. 4045. pauciciliatum. 3825. albemarlense. 4050. patulum. 3826. mattamuskeetense. 4051. patentifolium. 3829. mattamuskeetense. 40514. webberianum. 3831. microcarpon. 4053. chamaelonche. 3832. columbianum. 4058. patentifolium. 3850. bartowense. 4059. flavovirens. 3851. adspersum. 4062. patulum. 3855. fusiforme. 4063. equilaterale. 3859. virgatum cubense. 4066. verrucosum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Cuase, A.—Continued. 4068. 4072. 4077. 4086. 4088. _ 4091. 4094. 4095. 4099. 4100. 4101. 4104. 4105. 4112. 4113. 4118. 4119. 4121. 4122. 4126. 4132. 4141. 4149. 4151. 4153. 4154. 4156. 4161. - 4163. 4163}. 4166. 4170. 4171. 4172. 4173. 4174. 4175. 4179. 4181. 4183. 4187. 4188. 4191. 4194. 4201. 42014. 4202. 4205. 4207. 4208. 4209. 4211. 4217. lucidum. malacon. malacon. patulum. patentifolium. nitidum. joorii. anceps. joorii. dichotomum. laxiflorum. polycaulon. fusiforme. trifolium. lanuginosum. ovale. fusiforme. hemitomon. equilaterale. joorii. nitidum. longiligulatum. lancearium. albomarginatum chamaelonche. erectifolium. chamaelonche. arenicoloides. webberianum. patentifolium. trifolium. patentifolium. chamaelonche. longiligulatum. pauciciliatum. ovale. lancearium. fusiforme. joorii. patulum. tenerum. longiligulatum. rhizomatum. fusiforme. anceps. rhizomatum. verrucosum. commutatum. oligosanthes. boscii molle. patulum. arenicoloides. laxiflorum. | Coase, A.—Continued. 4219. 4220. 4225. 4229. 4234. 4239. 424]. 4242. 4245. 4248. 4249. 4250. 4251. 4256. 4258. 4260. 4261. 4267. 4271. 4274. 4276. 4277. 4279. 4281. 4282. 4283. 4284. 4286. 4287. 4290. 4291. 4292. 4297. 4299. 4301. 4302. 4304. 4305. 4307. 4308. 4310. 4312. 4314. 4318. 4320. 4321. 4322. 4331. 4340. 4346. 4351. 4357. 4358. joorii. rhizomatum. longifolium. angustifolium. dichotomiflorum. lancearium. equilaterale. lanuginosum. patentifolium. fusiforme. arenicoloides. ovale. malacon. joorii. equilaterale. oligosanthes. ovale, angustifolium. rhizomatum. joorii. oligosanthes. lanuginosum. ravenelii. arenicoloides. patulum. oligosant hes. anceps. agrostoides, agrostoides. mutabile. arenicoloides. lanuginosum. angustifolium. consanguineum. angustifolium. aciculare. trifolium. lanuginosum. nitidum. longiligulatum. flavovirens. patulum. verrucosur longifolium lucidum. nitidum. flavovirens. lanuginosum. arenicoloides. longifolium. consanguineum. patulum. trifolium. 343 344 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Cuase, A.—Continued. Cuase, A.—Continued. 4359. aciculare. 4506. xalapense. 4360. ciliatum. 4517. xalapense. 4364. polycaulon. 4519. barbulatum. 4366. longiligulatum. 4525. aciculare. 4371. patulum. 4527. condensum. 4377. repens. 4532. lanuginosum. 4389. brachyanthum. 4536. lancearium. 4392. dichotomiflorum. 4537. equilaterale. 4393. anceps. 4538. patulum. 4397. agrostoides. 4542. lancearium. 4400. lindheimeri. 4543. lancearium. 4401. lanuginosum. 4544. commonsianum. 4405. chrysopsidifolium. 4545. lancearium. 4407. gymmnocarpon. 4549. equilaterale. 4410. agrostoides. 4550. jooril. 4423. arenicoloides. 4554. anceps. 4424. verrucosum. 4555. amarum. 4426. microcarpon. 4560. rhizomatum. 4429. sphaerocarpon inflatum. 4564. longifolium. 4430. oorii. 4567. pauciciliatum. 4434. combsii. 4568. lancearium. 4437. joorii. 4569. lancearium. 4444. aciculare. 45694. webberianum. 4445. agrostoides. 4570. chamaelonche. 4449. lindheimeri. 4576. trifolium. 4453. flexile. 4577. patulum. 4459. virgatum. 4578. aciculare. 4460. virgatum. 4579. auburne. 4461. capillare. 4580. addisonii. 4463. clandestinum. 4581. arenicoloides. 4464. anceps. 4583. ciliatum. 4468. joorii. 4584. lanuginosum. 4469. barbulatum. 4585. angustifolium. 4470. sphaerocarpon. 4586. aciculare. 4472. villosissimum. 4589. ovale. 4473. ashel. 4591. sphaerocarpon. 4475. concinnius. 4600. scabriusculum. 4476. concinnius. 4601. commonsianum. 4477. tennesseense. 4746. barbipulvinatum. 4478. meridionale. 4886. barbipulvinatum. 4479. angustifolium. 5251. barbipulvinatum. 4482. microcarpon. 5252. thermale. 4483. concinnius. 5253. barbipulvinatum. 4485. polyanthes. 5292. barbipulvinatum. 4491. boscii. 5307. barbipulvinatum. 4492. flexile. 5345. barbipulvinatum. 4495. capillare. 5406. obtusum. 4497. stipitatum. 5411. linearifolium. 4498. ashei. 5412. xalapense. 4499. tennesseense. 5413. aciculare. 4501. boscii. 5414. aciculare. 4502. barbulatum. 5415. angustifolium. 4504. commutatum. 5416. bicknelli. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 345 CuaseE, A.—Continued. 5417. 5418. 5419. 5420. 5421. 5422. 5423. 5424. 5425. 5426. 5427. 5428. 5429. 5430. 5431. 5432. 5433. 5434. 5435. 5436. 5437. 5438. 5439. 5440. 5441. 5442. 5443. caerulescens. lucidum. microcarpon. annulum. spretum. lindheimeri. lindheimeri. tennesseense. tennesseense. lanuginosum. columbianum. commonsianum. columbianum. columbianum. oricola. sphaerocarpon. sphaerocarpon. patulum. oligosanthes. commutatum. mutabile. scoparium. aculeatum. verrucosum. philadelphicum. capillare. gattingeri. flexile. anceps. urvilleanum. urvilleanum. virgatum. huachucae silvicola. gattingeri. capillare. capillare. sphaerocarpon. pseudopubescens. perlongum. leibergii. praecocius. praecocius. capillare. gattingeri. virgatum. scribnerianum. perlongum. praecocius. praecocius. praecocius. leibergii. boscii. CuasE, V. H.—Continued. 1492. 1515. 1701. 1731. 1749. 1791. 1792. 1850. 1851. 1858. CHENEY, L. S. 678. 1082. 1107. 1216. 1345. 1346. 1545. 1700. 2100. 2911. 3088. 3409. 3426. 3471. 3868. 3872. 4104. 4786. 5638. Cocks, R. 8. 286. 292. 422. 2186. 2191. 2193. 2194. 3001. 3007. 3008. 3322. 3324. 3506. 3508. 3509. 3510. 3511. Coss, R. wm oe praecocius. praecocius. leibergii. perlongum. scribnerianum. praecocius. scribnerianum. praecocius. huachucae silvicola. huachucae. subvillosum. subvillosum. huachucae. depauperatum. werneri. xanthophysum. latifolium. boreale. boreale. philadelphicum. boreale. tennesseense. xanthophysum. tennesseense. virgatum. lindheimeri. huachucae. xanthophysum. huachucae. leucothrix. erectifolium. longiligulatum. repens. lanuginosum. lanuginosum. aciculare. anceps. sphaerocarpon inflatum. stipitatum. arenicoloides. scoparium. auburne. lindheimeri. huachucae silvicola. xalapense strictirameum. xalapense strictirameum. patulum. leucothrix. virgatum. virgatum. 346 Comss, R.—Continued. 26. 34. 57. 58. 60. 61. 63. 66. 67. 69. 73. 74. 75. 87. 89. 94. 98. 100. 104. 112. 114. 115. Wife IS 120. 125. 127. 132. 136. 137. 138. 139. 141. 144. 145. 156. 164. 167. 172. 178. 174. 182. 183. 187. 192. 194. 206. 215. 216. 224. 225. 231. 235. hians. flavovirens. ciliatum. erectifolium. roanokense. tenerum, virgatum. virgatum. scabriusculum. hemitomon. sphagnicola. ensifolium. lancearium. hemitomon. commutatum. dichotomiflorum. flavovirens in part. ciliatum. lancearium. albomarginatum. erectifolium. longiligulatum. hians. virgatum. rhizomatum. hians. chamaelonche. patulum in part. fusiforme. ciliatum. ovale. commutatum. xalapense. joorii. ovale. commutatum. arenicoloides. malacon. hians. microcarpon. lanuginosum. sphaerocarpon inflatum, aciculare. hians. rhizomatum. lanuginosum. hemitomon. lanuginosum. aciculare. mutabile. ovale. fusiforme. virgatum. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Comss, R.—Continued. 238. 201. 255. 256. 259. 262. 263. 266. 267. 270. 282. 287. 288. 290. 294. 295. 298. 299. 300. 301. 305. 306. 307. 320. 327. 329. 330. 334. 337. 347. 348. 304. 372. 373. 374. 376. 380. 381. 382. 390. 391. 399. 403. 406. 408. 412. 415. 427. 434. 437. 440. 441. 443. lancearium. commutatum. microcarpon. microcarpon. rhizomatum. hians. trifolium. nitidum. virgatum. joorii. anceps. hemitomon. ciliatum. wrightianum. lanuginosum. boscii molle. fusiforme. mutabile. arenicoloides. aciculare. ravenelii. sphaerocarpon. mutabile. anceps. laxiflorum. aciculare. commutatum, boscii. boscil. wrightianum., virgatum. wrightianum. aciculare. joorii. angustifolium. angustifolium. aciculare. anceps. oligosanthes. xalapense. microcarpon. boscil. sphaerocarpon inflatum, sphaerocarpon inflatum, aciculare joorii. anceps. hians. hians. virgatum. auburne. wrightianum. hemitomon. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Comss, R.—Continued. 444. 456. 458. 467. 470. 473. 475. 476. 477. 487. 488. 496. 516. 524. 525. 526. 530. 537. 539. 551. 552. 553. 554. 560. 567. 567a. 569. 570. 571. Bie. 574. cryptanthum. fusiforme. virgatum. erectifolium. anceps. virgatum. virgatum. tenerum. wrightianum. tenerum. longiligulatum. rhizomatum. fusiforme. wrightianum. curtifolium. hians. tenerum. virgatum. flavovirens. leucothrix. wrightianum. erectifolium. aciculare. anceps. strigosum. aciculare. longiligulatum. consanguineum. longifolium. leucothrix. tenerum. longifolium. virgatum. combsii. strigosum. flavovirens. anceps. hians. commutatum. virgatum. commutatum. ravenelii in part. villosissimum. lucidum. scabriusculum. leucothrix. tenerum. rhizomatum. longifolium. virgatum. virgatum. virgatum. rhizomatum. Comss, R.—Continued. 649. 650. polycaulon in part. albomarginatum. consanguineum. longiligulatum. mutabile. virgatum. erectifolium. wrightianum. capillare. leucothrix. leucothrix. hemitomon. boscii. lancearium. dichotomum, lucidum. virgatum. rhizomatum. virgatum. hemitomon. hians. anceps. virgatum. fusiforme. lanuginosum., equilaterale. boscii molle. mutabile. oligosanthes. laxiflorum. lancearium. ~ huachucae silvicola. lancearium. paludivagum. virgatum cubense. patentifolium. patentifolium. glabrifolium. patulum. equilaterale. caerulescens. agrostoides. erectifolium. rhizomatum. hians. hemitomon. patulum. hemitomon. sphagnicola. fusiforme. albomarginatum. malacon. patulum. 347 848 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Comss, R.—Continued. 859. 860. 878. 879. 884. 886. 887. 888. 899. 909. 915. 919. 921. 926. 926}. 930. 934. 935. 957. 960. 961. 9644. 971. 972. 973. 979, 990. 1008. 1010. 1011. 1012. 1021. 1022. 1023. 1024. 1025. 1036. 1037. 1038. 1049. 1051. 1052. 1053. 1063. 1064. 1068. 1073. 1080. 1085. 1086. 1087. 1088. 1098. patulum. joorii. anceps. agrostoides. rhizomatum. commutatum. virgatum. ovale. fusiforme. laxiflorum. agrostoides. rhizomatum. joorii. commutatum. equilaterale. virgatum cubense. lucidum. anceps. condensum. hemitomon. agrostoides. condensum. bartowense. anceps. condensum. nitidum. dichotomiflorum. nitidum. longifolium. condensum. agrostoides. anceps. arenicoloides. equilaterale. glabrifolium. lancearium. malacon. fusiforme. commutatum. verrucosum., rhizomatum. paludivagum. equilaterale. trifolium. laxiflorum. wrightianum. anceps. ovale. pauciciliatum. polycaulon. wrightianum. flavovirens. commutatum. Comss, R.—Continued. 1112. 1116. 1117. 1120. 1125. 1140. 1144. 1154. 1161. 1164. 1168. 1169. 1170. 11803. 1187. 1192. 1195. 1198. 1204. 1206. 1207. 1217a. 1218. 1220. 1234, 1235. 1237. 1240. 1241, 1246. 1249. 1251. 1252. 1253. 1261. 1264. 1265. 1266. 1269. 1270. 1280. 1288. 1294, 1297. 1303. 1310. 1311. 1313. 1316. 1323. 1325. 1333. 1338. tenerum. verrucosum. erectifolium. agrostoides, rhizomatum. microcarpon. condensum. longifolium. malacon. webberianum. longifolium. patulum. maximum, longifolium. glabrifolium. rhizomatum. hemitomon. agrostoides. agrostoides. anceps. agrostoides. rhizomatum. lucidum. bartowense. verrucosum. longifolium. joorii. lancearium. fusiforme. hians. virgatum. dichotomifiorum. rhizomatum. paludivagum. hians. rhizomatum, barbinode. tenerum. verrucosum. hemitomon. virgatum. patentifolium. verrucosum. virgatum. rhizomatum. maximum. barbinode. glabrifolium. glabrifolium. hemitomon. hians. patentifolium. polycaulon. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Comss, R.—Continued. 1340. 1343. 1344. 1393. 1394. 1395. 1398. 1400. 1419. 1426. 1427. 1481. 1435. (1436. Commons, A. 23. 25. 27. 28. 29. 30. 32. 37. 38. 230. @ Herbarium Instituto ffsico-geogr4fico. fusiforme. fusiforme. chamaelonche. joorii. flavovirens. rhizomatum. anceps. joorii. dichotomiflorum. gymmnocarpon. joorii. reptans. capillare. dichotomiflorum. hemitomon. stipitatum. clandestinum. scoparium. depauperatum. capillare. scoparium. ashei. villosissimum. lindheimeri. commonsianum. oricola. polyanthes. commutatum. sphaerocarpon. sphaerocarpon. villosissimum. ashei. oligosanthes. lindheimeri. villosissimum in part. miliaceum. columbianum thinium. oricola. oricola. ashei. lindheimeri. lindheimeri. lindheimeri. lindheimeri. lindheimeri. lindheimeri. longifolium. anceps. amarum. dichotomiflorum. dichotomiflorum. Commons, A.—Continued. 231. 281. 282. 283. 284. 285. 286. 289. 290. 291. 292. 293. 294. 295. 296. 297. 298. 300. 301. 356. 357. 358. 359. 360. 361. 362. 363. 364. 365. Costa Rica.@ 412. 1183. 1244. verrucosum. scribnerianum. scribnerianum. scoparioides. sphaerocarpon. microcarpon. meridionale. huachucae silvicola. huachucae silvicola. lindheimeri. huachucae silvicola. huachucae silvicola. dichotomum. dichotomum. barbulatum. dichotomum. sphaerocarpon. clandestinum. latifolium in part. clandestinum. anceps. longifolium. stipitatum. polyanthes. polyanthes. ashei. commutatum. spretum. commonsianum. lanuginosum. longifolium. longifolium. clutei. microcarpon. microcarpon. spretum. ashei. sphaerocarpon. ashei. ashei. werner. scoparioides. huachucae silvicola. boscii. huachucae silvicola. lindheimeri. huachucae silvicola. tennesseense. barbinode. polygonatum. viscidellum. 349 Collectors Biolley, Cooper, Pittier, Tonduz. 350 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Costa Rica—Continued. 2035. 2479. 2598. 2633. 2995. 3071. 3107. 3117. 3123. 3358. 3583. 3619. 3631. 3640. 3651. 3661. 3673. 3679. 3685. 3687. 4042. 4092. 4458. 4459. 4460. 4626. 4860. 4864. 4865. 4871. 4875. 4881. 7360. 7463. 7467. 7471. 7878. 7944. 8557. 8566. 8600. 8670. 8818. 8829. 9050. 9080. 9114. 9495. 9727. 9754. 10379. 10576. 10588. fasciculatum. polygonatum. zizanioides. ghiesbreghtii. maximum. laxum. polygonatum. laxum. trichoides. viscidellum. stenodes. virgatum. parvifolium. costaricense. trichoides. costaricense. costaricense. rudgei. cayennense. costaricense. polygonatum. polygonatum. trichoides. pulchellum. pulchellum in part. costaricense. costaricense. laxum. trichoides. laxum. rudgei. pulchellum. pulchellum. pilosum. hirsutum. fasciculatum. sphaerocarpon. olivaceum. polygonatum. zizaniodes. trichanthum. trichanthum. virgultorum. virgultorum. maximum. parviglume. maximum. pilosum. fasciculatum. trichoides. trichoides. rudgei. rudgei. Costa Rica—Continued. 10589. 10594. 10615. 10745. 11017. 11276. 11393. 11396. 11866. 12002. 12064. 13749. 16081. 16123. stenodes. parvifolium. millegrana. sphaerocarpon. olivaceum. glutinosum. zizanioides. - barbinode. polygonatum. sphaerocarpon. glutinosum. rudgei. fasciculatum. trichoides. laxum. Curtiss, ASHE 3585. 3587. ASE Cn Ce a GIaskkawoet AOA Bo arenicoloides. nudicaule. chrysopsidifolium. pauciciliatum. villosissimum. boscil. scabriusculum. lanuginosum. chamaelonche. adspersum. barbinode. maximum. condensum. geminatum. dichotomifilorum. cayennense. pilosum. acuminatum. acuminatum. diffusum. fusiforme. laxum. diffusum. reptans. trichanthum. reptans. trichoides. adspersum. amarulum. rhizomatum. tenerum. commutatum., arenicoloides. ovale. nudicaule. hemitomon. angustifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Curtiss, A. H.—Continued. 3587*. 3589. 3594. 3597*. 3597**. 3599. 3600A. 3601. 3602*. 3606**. 3607. 3608. 3609. 3610. 4027. 4028. 4029. 4031. 4033. 4035. 4036. 4636. 4637. 4811. 4812. 4877. 4878. 5083. 5084. 5252. 5289. 5298. 5302. 5386. 5431. 5457. 5527. 5534. 5537. 5576. 5576A. 5576B. 5576C. 5588. 5747. 5804. 5808. 5813. 5864. arenicoloides. fasciculatum. hians. strigosum. maximum. erectifolium. dichotomum. equilaterale. lancearium. nitidum. pauciciliatum. geminatum. chamaelonche. adspersum. chapmani. verrucosum. virgatum. scabriusculum. commutatum. arenicoloides. lancearium. strigosum. longiligulatum. tenerum. verrucosum. commutatum. webberianum. hemitomon. erectifolium. ovale. scabriusculum. tenerum. virgatum. verrucosum. rhizomatum. condensum. agrostoides, bartowense. adspersum. chapmani. amarulum. hians. xalapense. condensum. condensum. longifolium. tenerum. chamaelonche. rhizomatum. chamaelonche. verrucosum. ovale. oligosanthes. Curtiss, A. H.—Continued. 5866. 5912. 5936. 6047. 6403. 6513. 6601. 6602. 6610. 6616. 6626. 6627. 6628. 6635. 6636. 6639. 6652. 6705. 6791. 6803. 6811. 6817. 6827. 6828. 6836. 6867. 6887. 6888. 6890. 6919. 6925A. CUTHBERT, A. 382. 387. 388. 392. 427. 431. 529. 962. 1015. 1119. 1120. 1121. 1128. 1157. 1158. 1159. 1160. 1161. 1162. 1163. ovale. wrightianum. anceps. boscii molle. lucidum. repens. lucidum. xalapense. webberianum. ovale. lancearium. polycaulon. chamaelonche. xalapense. roanokense. aciculare. wrightianum. adspersum. xalapense. angustifolium. auburne. erectifolium. leucothrix. lanuginosum. sphaerocarpon. hians. miliaceum. anceps. agrostoides. agrostoides. combsii. combsii. trifolium. villosissimum. depauperatum. boscii molle. ashei. annulum. lucidum. dichotomum. virgatum cubense. xalapense. villosissimum. oligosanthes. agrostoides. sphaerocarpon. sphaerocarpon. trifolium. ensifolium. microcarpon. boscii molle. ashei. 351 352 CutTHBeErtT, A.—Continued. 1164. 1165. 1166. 1167. DAvyY, J: \B: 5894. 5971. 6092. 6745. 6780. Dream, C. C. 424. 2065. 2638. 3218. 5386. 5392. 6041. 6143. 6267. 6268. 6467. 6595. 6753. 6883. Dewey, L. H. 53. oe 92. 134. 235. 280. 408. Dop@e, C. K. 17. 20. 38. 49. 60. 62. 78. 83. 84. 124. 128. 130. Dorner, H. B. pseudopubescens. angustifolium. aciculare. oligosanthes. pacificum. occidentale. occidentale. pacificum. pacificum. trichanthum. tennesseense. virgatum. linearifolium. boscil. polyanthes. trichoides. molle. fasciculatum. maximum. ashei. commutatum. perlongum. xalapense. anceps. boscii molle. boscii molle. virgatum. boscii. boscil. verrucosum. leibergii. leibergii. implicatum. lindheimeri. villosissimum. leibergii. depauperatum. villosissimum. leibergil. flexile. flexile. flexile. virgatum. dichotomiflorum. leibergil. latifolium. dichotomum. huachucae silvicola. virgatum. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Dorner, H. B.—Continued. 86. dichotomiflorum. 87. gattingeri. 88. scribnerianum. 91. huachucae silvicola. 93. huachucae silvicola. DrumMonp, T. 286. filipes. 367. hemitomon. 381. scoparium. 384. filipes. 394. filipes. 452. patulum. 454. huachucae silvicola. 456. xalapense. 457. xalapense. 461. hemitomon. Duss, PmRE. 536. diffusum. 537. fasciculatum. 538. fasciculatum. 539. barbinode. 1288. maximum. 1290. reptans. 1293. geminatum. 321.. trichoides. 2681. trichoides. 2689. barbinode. 2690. geminatum. 2691. fasciculatum. 3177. utowanaeum. 3178. dichotomiflorum. 3179. laxum. 3180. adspersum. 3184. ghiesbreghtii. 3186. maximum. 3529. reptans. 3584. geminatum. 3917. hirsutum. 3919. condensum. 4154. pilosum. Farts, F.8., AnD BAKER, C.F. 1522. pseudopubescens. 1527. auburne. 1530. wilmingtonense in part. 1531. commutatum. 1532. curtifolium. 1535. trifolium in part. 1537. pseudopubescens. 1544. annulum. 2455. reptans. Baton, A. A. 164. agrostoides. bartowense. 165. agrostoides. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Eaton, A. A.—Continued. 169. 242. 341. 467. 574. 578. 589. 831. Eacers, H. F. 165. 293. 1226. 1328. 1329. 2129. 3978. 4305. 4312. 4405. 4512. 4870. 4875. 5350. 5406. 5534. 5810. 5987. 14149. 14345. 14481. 14585. 15417. 15418. 15419. 15422. 15834. Eacert, H. ie 10. 12. 13. 14. 15. 20. Zt. 22. 23. 24. 26. 30. 34. 39. joorii. agrostoides. anceps. bartowense. hemitomon. erectifolium. xalapense. xalapense. A. laxum. reptans. maximum. barbinode. laxum. xalapense. exiguiflorum. caerulescens. caerulescens. dichotomiflorum. dichotomiflorum. barbinode. fasciculatum. trichanthum. ghiesbreghtii. pilosum. zizanioides. trichoides. stoloniferum. megiston. trichoides. laxum. trichoides. fasciculatum. ghiesbreghtii. zizanioides. fasciculatum. commutatum. angustifolium. scoparium. sphaerocarpon. polyanthes. microcarpon. dichotomum. strigosum. commutatum. sphaerocarpon. trifolium. boscii. huachucae silvicola. lindheimeri. depauperatum. 41616°—vo1t 15—10——23 353 Eacert, H.—Continued. 40. 42. 43. 44, 44}. 45. 46. 58. 59. 60. 78. 82. 86. 89. "95. 110. 114. 115. 116. 117. 124, 125. 126. 127. 233. 234. 235. 236. 237. 239. 240. 241. 242. 243. 244. 245. 246. 249. 250. 253. 254. 257. 287. 288. 289. 290. 291. 292. 293. 295. 296. 297. boscii. laxiflorum. commutatum. ashei. dichotomum. commutatum. dichotomum. philadelphicum. verrucosum. scoparium. microcarpon. scoparium. xalapense. microcarpon. polyanthes. dichotomiflorum. clandestinum. sphaerocarpon. scoparium. hians. huachucae silvicola. agrostoides. virgatum. xalapense. depauperatum. scribnerianum. huachucae silvicola. microcarpon. lindheimeri. latifolium. polyanthes. flexile. tennesseense. tennesseense. huachucae silvicola. huachucae silvicola. dichotomum. scribnerianum. polyanthes. clandestinum. sphaerocarpon. commutatum. microcarpon. nitidum. huachucae silvicola. commutatum. villosissimum. villosissimum. villosissimum. scribnerianum. tennesseense. scribnerianum. 354 Eae.eston, W. W. 1757. 1758. 1759. 2181. 2843. FARWELL, O. 597. 597b. 597d. 642. 643. 6438a. 648b. 750. 764. 893. 1378. 1382. 1388. 1414. FENDLER, A. 368. 946. 1634. 1638. 16388. 1641. 1643. 2499. FERNALD, M. 166. 166a. 239. Paral 276. 280. 292. 345. 346. 361. 406. 500. 501. 502. 503. 504. 505. 506. 508. 509. 510. 512. werneri. subvillosum. dichotomum. tennesseense. tsugetorum. JN. linearifolium. implicatum. huachucae silvicola. subvillosum. boreale. implicatum. huachucae. perlongum. xanthophysum. flexile. latifolium. huachucae. huachucae silvicola. sphaerocarpon. miliaceum., pilosum. polygonatum. zizanioides. sphaerocarpon. olivaceum. millegrana. trichanthum. trichoides. ip tennesseense. tennesseense. boreale virgatum. latifolium. scribnerianum. clandestinum. xanthophysum. latifolium. philadelphicum. dichotomum. implicatum. subvillosum. implicatum. huachucae silvicola. implicatum. boreale. werneri. boreale. huachucae silvicola. spretum. boreale. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. FERNALD, M. L.—Continued. 513. 514. 516. 517. 518. 519. 520. 521. 2802. GARDNER, G. 1178. 1179. 1183. 1876. 2353. 2357. 2361. 3517. GRAVES, C. B. 156. 157. 158. 163. 164. 165. 166. 167. 1B Ae 172. 236. 244. 248. 256. boreale. boreale. boreale. boreale. clandestinum. xanthophysum. xanthophysum. xanthophysum. philadelphicum. rudgei. megiston. laxum. molle. molle. fasciculatum. molle. laxum. huachucae. werneri. huachucae silvicola. columbianum. tennesseense. barbulatum. villosissimum. bicknellii. wernerl. werneri. lindheimeri. boreale. implicatum. spretum. boreale. columbianum. columbianum. sphaerocarpon. spretum. ashei. huachucae silvicola. spretum. lindheimeri. latifolium. latifolium. implicatum. huachucae silvicola. philadelphicum. meridionale. meridionale. stipitatum. virgatum cubense. longifolium. longifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Grirrirus, D. Te 15. 22. 53. barbipulvinatum. capillare. virgatum. virgatum. virgatum. capillare. virgatum. virgatum. capillare. virgatum. barbipulvinatum. virgatum. virgatum. virgatum. barbipulvinatum. scribnerianum. barbipulvinatum. virgatum. leibergii. perlongum. scribnerianum. obtusum. hirticaule. fasciculatum chartagi- nense. obtusum. arizonicum. fasciculatum chartagi- nense. fasciculatum chartagi- nense. arizonicum. hallii. arizonicum. hirticaule. obtusum. hirticaule. obtusum. obtusum. barbipulvinatum. arizonicum. hirticaule. fasciculatum chartagi- nense. hallii. obtusum. plenum. pacificum. pacificum. pacificum. bulbosum sciaphilum. virgatum. obtusum., Grirrirus, D.—Continued. 5401. 5463. 5504. 5542. 5551. 5600. 5612. 5664. 5735. 5852. 5981. 6123. 6151. 6168. 6288. 6306. 6323. 6380. 6381. 6387. 6432. 6441. 6445. 6446. 6508. 6551. 6586. 6737. 6747. 6758. 6759. 67854. 6799. 6800. 6857. 6891. 6892. 6894, 6929. 6938. 6939. 69394. 6959. 6990. 7005. 7017. 7063. 7083. 7143. 7146. 7148. 7194, scribnerianum. virgatum. hallii. obtusum. barbipulvinatum. hallii. obtusum. virgatum. havardii. barbipulvinatum. arizonicum. hirticaule. arizonicum. arizonicum. hallii. geminatum. filipes. ramisetum. ciliatissimum. filipes. ciliatissimum. ciliatissimum. ciliatissimum. firmulum. fasciculatum. sphaerocarpon. pacificum. arizonicum. arizonicum. hirticaule. arizonicum. 355 bulbosum sciaphilum. hirticaule. obtusum. fasciculatum. arizonicum. hirticaule. arizonicum. arizonicum. arizonicum. arizonicum. pampinosum. obtusum. arizonicum. hirticaule. arizonicum, lepidulum. bulbosum. arizonicum. hirticaule. arizonicum. hirticaule, 356 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Grirriras, D.—Continued. 7288. 7297. 7299. 7317. 7357. 7399. 7400. 7401. 7408. Hatt, E. 231. 671. 672. 815. 816. 817. 819. 820. 823. 824. 825. 827. 828. 829. 830. 831. 832. 833. 834. HANSEN, G 599. 626. 1381. 1444. 1723. Harper, R. M. 15. 60. 70. 74. ids 88. 95. 104. 110. 146. 147. obtusum., fasciculatum chartagi- nense. fasciculatum chartagi- nense. hirticaule. hallii. arizonicum in part. plenum. plenum. hallii. wilcoxianum. pacificum. scribnerianum. hians. hailii in part. dichotomiflorum. agrostoides. hemitomon. geminatum. ciliatissimum. fasciculatum chartagi- nense. obtusum. joorii. boscii. scoparium. helleri. commutatum. sphaerocarpon. angustifolium. ovinum. pedicellatum. nitidum. capillare. pacificum. pacificum. pacificum. occidentale. boscii molle. commutatum. clandestinum. microcarpon. xalapense. lucidum. dichotomum. polyanthes. scoparium. oligosanthes. mutabile. Harper, R. M.—Continued. 150. 183. 184. 210. 221. 281. 369. 382. 410. 429. 459. 522. 631. 638. 742. 757. 765. 767. 828. 829. 838. 839. 881. 1007. 1037. 1045. 1081. 1085. 1087. 1105. 1106. 1220. 1239. 1349. 1366. 1394. 1399. 1435. 1485. 1575. 1623. 1679. 1689. 1730. 1760. 1812. 2014. 2140. Hart, J. 73. 78. 87. 726. scoparium, xalapense. philadelphicum. scoparium. commutatum. microcarpon. anceps. dichotomiflorum. scabriusculum. virgatum. longiligulatum. gymnocarpon. virgatum. verrucosum. amarum. arenicoloides. lanuginosum. joorii. angustifolium. ensifolium. hians. longiligulatum. scabriusculum., hemitomon. virgatum. tenerum. combsii. xalapense. sphaerocarpon. microcarpon. joorii. virgatum. agrostoides. yadkinense. boscii. erectifolium. lanuginosum. mutabile. erectifolium. longiligulatum. commutatum. combsii. lancearium, agrostoides. lucidum. boscii. combsii. xalapense. polygonatum. fasciculatum. trichanthum, zizanioides. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Hart, J.—Continued. 732. 736. 785. 792. 797. 806. 838. 840. 2177. 2289. 3293. HELLER, A. A. 100. 135. 157. 377. 387. 497. 522. 531. 639. 701. 968. 982. 982b. 1312. 1316. 1378. 1490. 1603. 1726. 1736. 1741. 1752. 1759. 1766. 1883. 1888. 1898. 3978. 4058. 4082. 4083. 4084. 4085. 4088. 4103. 4120. 4145. 4160. 4209. 4210. pilosum. acuminatum. fasciculatum. glutinosum. maximum. geminatum. reptans. fasciculatum. laxum. laxum. pilosum. barbinode. fasciculatum. trichoides. maximum. trichoides. reptans. laxum. trichoides. pauciciliatum. flexile. polyanthes. chrysopsidifolium. pauciciliatum. tsugetorum. parvifolium. laxum. filipes. reverchoni. pedicellatum. pedicellatum. obtusum. lindheimeri. helleri. pedicellatum. filipes. dichotomiflorum. lindheimeri. plenum. occidentale. scribnerianum. anceps. commutatum. huachucae silvicola. xalapense. microcarpon. hians. joorii. hians. tennesseense. xalapense. dichotomiflorum. Heuer, A. A.—Continued. 357 4221. brachyanthum. 4235. agrostoides. 4236. scoparium. 4237. microcarpon. 4238. consanguineum. 4246. dichotomiflorum. 4528. fasciculatum. 4768. commutatum. 4769. dichotomum. 4770. ashei. microcarpon. 4771. boscii. 4772. polyanthes. 4774. tennesseense. 4775. depauperatum. 4776. barbulatum. 4777. flexile. 4778. tennesseense. 4779. boscii molle. 4780. ashei. 4781. clandestinum. 4783. dichotomum. 4785. huachucae silvicola. 4786. flexile. 4787. microcarpon. 4789. philadelphicum. 4790. meridionale. 4791. huachucae silvicola. 6094. trichoides. 6226. fasciculatum. 6293. barbinode. 6302. fasciculatum. 6442. pauciciliatum. 7856. pacificum. How, B.. J. 37 in 1907. praecocius. 49 in 1909. praecocius. 50 in 1909. boreale. 53 in 1907. albemarlense. 83 in 1908. huachucae silvi- cola. 84 in 1908. dichotomum. 85 in 1908. huachucae _silvi- cola. 86 in 1908. tennesseense. 88 in 1908. sphaerocarpon. 90 in 1908. sphaerocarpon. 91 in 1906. leibergii. 97 in 1908. boreale. 98 in 1905. meridionale. 98 in 1908. huachucae. 99 in 1905. implicatum. 100 in 1905. pseudopubescens. 358 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Hu, E. J.—Continued. 101 in 1905. pseudopubescens. 124 in 1905. tsugetorum. 128 in 1906. spretum. 129 in 1905. tsugetorum. 129 in 1906. barbulatum. 130 in 1905. huachucae. 141 in 1905. tennesseense. 145 in 1906. meridionale. 158 in 1906. clandestinum. 162 in 1878. tennesseense. 162 in 1906. spretum. 163 in 1906. implicatum. 171 in 1907. huachucae. 177 in 1898. verrucosum. 182 in 1907. huachucae. 1834 in 1907. lindheimeri. 184 in 1907. implicatum. 185 in 1907. implicatum. 201 in 1898. oligosanthes. 217 in 1898. huachucae. Hitcucock, A. §. 4. nitidum. 6. yadkinense. 7. philadelphicum. 10. auburne. 14. leucothrix. 15. nitidum. 16. longiligulatum. 17. aciculare. 19. boscii. 20. oligosanthes. 25. flavovirens. 26. sphaerocarpon inflatum. 47. oricola. 102. ashei. 104. rhizomatum. 107. ovale. 108. arenicoloides. 109. neuranthum. 110. xalapense. 111. laxiflorum. 112. polycaulon. 114: fusiforme. 115. polycaulon. 116. chrysopsidifolium. 117. fusiforme. 118. bicknellii. 119. nitidum. 120. boreale. 121. boreale. 122. dichotomum. 123. barbulatum. 124. barbulatum. Hircxcocx, A. S.—Continued. 125. 126. 127. 129. 130. 132. 133. 134. 135. 136. 137. 138. 139. 140. 141. 142. 143. 144. 145. 146. 150. 151. 152. 153. 154. 155. 156. 157. 160. 161. 162. 163. 164. 165. 166. 167. 168. 169. 170. iyi 172. 174. 175. Uz. 178. 179. 180. 181. 2C1. 215. 216. 217. barbulatum. albemarlense. implicatum. agrostoides. implicatum. implicatum. huachucae. tsugetorum. tsugetorum. huachucae silvicola. huachucae silvicola. tennesseense, tennesseense. acuminatum. utowanaeum. geminatum. geminatum. distantiflorum. reptans. reptans. dichotomiflorum. dichotomiflorum. elephantipes. virgatum cubense. tenerum. stenodes, maximum. maximum. ovale. villosissimum. fasciculatum chartagi- nense. texanum. verrucosum. philadelphicum. philadelphicum. gattingeri. amarulum. amarum. virgatum. virgatum. condensum. condensum. rhizomatum laxum. laxum. exiguiflorum. millegrana. parvifolium. bicknellii. scoparium, verrucosum. dichotomiflorum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 359 Hirewcocxr, A. S.—Continued. 218. 219. 220. 221. 222. 225. 226. 227. 228. capillarioides. hallii. filipes. hallii. boscii. stipitatum. virgatum. amarum. anceps. anceps. anceps rhizomatum. wilmingtonense. commonsianum. tenue. addisonii. commonsianum flavovirens. chamaelonche. chamaelonche. depauperatum. depauperatum. ciliatum. xalapense. aciculare. aciculare. aciculare. aciculare. angustifolium. angustifolium. arenicoloides. arenicoloides. arenicoloides. bicknellii. mattamuskeetense. caerulescens. dichotomum. barbulatum. barbulatum. yadkinense. lucidum. lucidum. lucidum. lucidum. lucidum. lucidum. lucidum. lucidum. microcarpon. cryptanthum. longiligulatum. wrightianum. wrightianum. Hrirencock, A. §.—Continued. 375. 376. 377. 378. 379. 380. 381. 384. 385. 386. 387. 389. 390. 391. 392. 393. 394. 395. 396. 397. 398. 399. 406. 407. 408. 410. 411. 413. 414. 416. 417. 418. 419. 420. 421. 422. 423. 424. 425. 426. 427. 428. 429. 430. 431. 432. 433. 434. 435. 436. 437. 438. 439. longiligulatum. leucothrix. leucothrix. spretum. spretum. lindheimeri. lindheimeri. meridionale. meridionale. lanuginosum. lanuginosum. lanuginosum. lanuginosum. villosissimum. villosissimum. villosissimum. villosissimum. villosissimum. villosissimum. pseudopubescens. pseudopubescens. addisonii. columbianum thinium, commonsianum. commonsianum. polyanthes. erectifolium. patulum. paucitiliatum. pauciciliatum. oligosanthes. oligosanthes. oligosanthes. ravenelii. equilaterale. ashei. ashei. ashei. ashei. commutatum. commutatum. commutatum. mutabile. mutabile. clandestinum. clandestinum. latifolium. boscii molle. boscii molle. boscii molle. scoparium. scabriusculum. philadelphicum. 360 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Hircucocxk, A. §.—Continued. 440. 441. 442. 444. 445. 446. 447. 448. 449. 450. 451. 503. 505. 500. 501. 582. 593. 504. 555. 506. 597, 508. 562. 563. 564. 565. 566. 567. 568. 569. 570. 571. 572. 573. 574. 575. 576. 577. 578. 579. 580. 581. 582. 583. 584. 585. 586. 587. 588. 589. 590. 591. 592. amarulum. amarum. virgatum. condensum. agrostoides. longifolium. longifolium. anceps. anceps. rhizomatum. hians. ashei. tennesseense. ovale. lindheimeri. lindheimeri. spretum. leucothrix. albomarginatum. addisonii. addisonii. oricola. sphaerocarpon. patentifolium. ashei. ashei. ashei. ashei. ashel. commutatum., commutatum. commutatum. commutatum. mutabile. equilaterale. equilaterale. joorii. joorii. malacophyllum. oligosanthes. oligosanthes. oligosanthes. oligosanthes. clandestinum. clandestinum. clandestinum. latifolium. latifolium. latifolium. latifolium. boscii. boscii. boscii molle. Hircucocr, A. §8.—Continued. 593. 594, 595. 596. 597. 598. 599. 613. 627. 6274. 630. 633. 634. : 636. 639. 648. 652. 653. 654. 655. 658. 659. 660. 661. 664, 665. 666. 667. 668. 670. 673. 674. 677. 678. 679. 687. 688. 6894. 690. 692. 695. 696. 697. 699. 705. 706. 710. 711. 712. 713. 714. 715. 718. scoparium. scoparium. scabriusculum. microcarpon. linearifolium. linearifolium. huachucae silvicola. geminatum. fusiforme. virgatum. webberianum. equilaterale. ovale. joorii. albomarginatum., bartowense. joorii. equilaterale. patulum. equilaterale. bartowense. xalapense. joorii. ovale. lancearium. polycaulon. albomarginatum. albomarginatum. ovale. albomarginatum. xalapense. joorii. ovale. lancearium. albomarginatum. nitidum. ovale. polycaulon. caerulescens. albomarginatum. condensum. hemitomon. bartowense. condensum. neuranthum caerulescens. neuranthum., polycaulon. trifolium. lancearium. albomarginatum. caerulescens. nitidum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Hitcucocr, A. §.—Continued. 719. 720. 721. 728. 729, 730. 733. 734. 735. 737. 743. 744. 747. 748. 750. 75. 756. 757. 759. 760. 761. 7614. 762. 763. 764. 765. 765}. 766. 766}. 767. 767}. 768. 769. 770. 771. 772. 773. 774. 775. 778. 779. 781. 782. 784, 785. 786. 787. 791. 792. 793. 794, 795. 796. ovale. albomarginatum. polycaulon. virgatum. chamaelonche. patentifolium. pauciciliatum. breve. webberianum. pauciciliatum. virgatum cubense. hemitomon. condensum. webberianum. pauciciliatum. patentifolium. joorii. commutatum. chamaelonche. joorii. albomarginatum. ovale. polycaulon. polycaulon. lancearium. patulum. webberianum. lancearium. webberianum. patentifolium. flavovirens. albomarginatum. chamaelonche. lancearium. polycaulon. polycaulon. xalapense. nitidum. chrysopsidifolium. chamaelonche. trifolium. chamaelonche. webberianum. lancearium. ovale. fusiforme. ovale. fusiforme. webberianum. pauciciliatum. chamaelonche. vernale. patentifolium. Hircucock, A. 8.—Continued. 797. 798. 800. 801. 802. 803. 804. 805. 806. 807. 808. 809. 811. 813. 814. 817. 818. 819. 820. 821. 823. 824. 825. 826. 827. 828. 833. 834. 835. 836. 837. 838. 8384. 839. 840. 843. 844. 845. 846. 847, 848. 849, 850. 851. 855. 856. 863. 864. 865. 866. 867. 868. 869. pauciciliatum. vernale. leucothrix. malacon. patentifolium. pauciciliatum. pauciciliatum. leucothrix. chamaelonche. chamaelonche. pauciciliatum. vernale. webberianum., malacon. nitidum. albomarginatum. chamaelonche. pauciciliatum. ovale. lancearium. albomarginatum. lancearium. lancearium. albomarginatum., polycaulon. chamaelonche. ovale. fusiforme. angustifolium. polycaulon. fusiforme. albomarginatum. webberianum. albomarginatum. webberianum. polycaulon. lancearium. malacon. ovale. ovale. albomarginatum. lancearium. fusiforme. ovale. webberianum, patentifolium. hemitomon. lancearium. chamaelonche. chamaelonche. curtifolium. polycaulon. lancearium. 361 362 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Hircucock, A. §.—Continued. Hircncock, A. §.—Continued. 870. albomarginatum. 933. polycaulon. 872. hians. F 934. chamaelonche. 873. chamaelonche. 935. xalapense. 874. erectifolium. 936. vernale. 875. longiligulatum. 937. lancearium. 876. albomarginatum. 938. trifolium. 877. fusiforme. 9384. roanokense. 880. albomarginatum. 939. roanokense. 881. longiligulatum. 940. trifolium. 882. albomarginatum. 941. vernale. 884. albomarginatum. 942. vernale. 885. chamaelonche. 943. polycaulon. 886. albomarginatum. 944. chamaelonche. 887. chamaelonche. 945. albomarginatum. 888. lancearium. 946. patulum. 889. pauciciliatum. 947. joorii. 890. trifolium. 949. albomarginatum. 893. glabrifolium. 950. polycaulon. 895. chamaelonche. 951. chamaelonche. 897. caerulescens. 952. chamaelonche. 898. nitidum. 953. trifolium. 899. fusiforme. 954. chamaelonche. 900. ovale. 955. webberianum. 9014. flavovirens. 956. lancearium. 903. polycaulon. 957. longiligulatum. 904. caerulescens. 958. longiligulatum. 905. flavovirens. 9584. vernale. 906. xalapense. 959. -vernale. 908. nitidum. 960. vernale. 909. joorii. 961. nitidum. 910. webberianum. 962. trifolium. 911. lancearium. 963. albomarginatum. 912. fusiforme. 964. fusiforme. 913. lancearium. 965. caerulescens. 914. ovale. 966. glabrifolium. 915. caerulescens. 968. ovale. 9154. patentifolium. 969. patentifolium. 916. chamaelonche. 970. patentifolium. 918. webberianum. 971. patentifolium. 919. lucidum. 972. webberianum. 920. trifolium. 973. chamaelonche. 921. trifolium. 974. polycaulon. 922. patulum. 975. chamaelonche. 923. fusiforme. 976. joorii. 9234. polycaulon. 977. lancearium. 924. chamaelonche. 978. glabrifolium. 927. xalapense. 979. webberianum. 928. chamaelonche. 980. fusiforme. 929. lancearium. 984. angustifolium. 930. patentifolium. 985. chamaelonche. 9304. webberianum. 986. consanguineum. 931. vernale. 987. roanokense. 932. lancearium. 988. longiligulatum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Hrrcucock, A. S.—Continued. 989. 990. 991. 992. 993. 994. 997. 998. 999. 1000. 1003. 1004. 1005. 10054. 1006. 1008. 1009. 1010. 10104. 1011. 1012. 1013. 10133. 1014. 1015. 1016. 1019. 1020. 1021. 1022. 1023. 1024. 1025. 1026. 1027. 1028. 1032. 1033. 1035. 1036. 1037. 1038. 1039. 10394. 1040. 1041. 1042. 1043. 1046. 1047. 1048. 1049. 1050. aciculare. albomarginatum. ciliatum. pauciciliatum. ciliatum. lancearium. trifolium. roanokense. consanguineum. ciliatum. laneearium. vernale. longiligulatum. spretum. sphagnicola. commutatum. laxiflorum. ravenelii. boscii molle. joorii. arenicoloides. ovale. pseudopubescens. oligosanthes. mutabile. lancearium. ciliatum. vernale. albomarginatum. albomarginatum. trifolium. webberianum. lancearium. lucidum. chamaelonche. lancearium. lanuginosum. lancearium. mutabile. ciliatum. chamaelonche. trifolium. angustifolium. lancearium. ciliatum. vernale. trifolium. strigosum. villosissimum. ravenelii. pseudopubescens, commutatum. trifolium. Hircucock, A. S.—Continued. 1052. 1053. 1054. 1055. 1058. 1059. 1062. 1063. 1066. 1067. 1068. 1069. 1070. 1072. 1073. 1077. 10774. 1078. 1079. 1080. 1081. 1082. 1083. 1084. 1087. 1088. 1089. 1090. 1091. 1092. 1093. 1094. 1095. 1096. 1099. 1100. 1101. 1102. 1103. 1104. 1105. 1106. 1107. 1108. 1109. 1111. 1112, 1113. 1114. 1115. 1124, 1126. 1127. scabriusculum. trifolium. fusiforme. oligosanthes. curtifolium. lucidum. ciliatum. trifolium. vernale. ensifolium. longiligulatum. consanguineum. consanguineum., trifolium. xalapense. arenicoloides. ovinum. sphaerocarpon. lanuginosum. pseudopubescens. nitidum. aciculare. consanguineum. sphaerocarpon. angustifolium. trifolium. trifolium. pseudopubescens. angustifolium. vernale. lancearium. curtifolium. sphaerocarpon. pseudopubescens. sphaerocarpon. trifolium. polycaulon. sphaerocarpon. mutabile. commutatum. angustifolium. oligosanthes. ravenelii. commutatum. villosissimum. trifolium. pseudopubescens. pauciciliatum. chamaelonche. angustifolium. xalapense. sphaerocarpon. aciculare. 363 364 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Hircucock, A. §.—Continued. Hirencocxk, A. §S.—Continued. 1128. nitidum. 1209. angustifolium. 1129. lanuginosum. 1210. ovinum. 1130. trifolium. 1212. helleri. 1131. ovinum. 1213. sphaerocarpon. 1132. sphaerocarpon. 1214. huachucae silvicola. 1133. hians. 1215. lindheimeri. 1134. sphaerocarpon. 1216. sphaerocarpon. 1135. lanuginosum. 1221. angustifolium. 1136. lindheimeri. 1222. ovinum. 1137. sphaerocarpon. 1225. aciculare. 1138. lindheimeri. 1226. thurowii. 1139. consanguineum. 1238. lanuginosum. 11394. aciculare. 1239. xalapense. 1140. aciculare. 1240. dichotomum. 1141. ovinum. 1241. sphaerocarpon. 1142. joorii. 1244. boscii. 1143. nitidum. 1245. ravenelii. 1144, roanokense. 1246. dichotomum. 1146. trifolium. 1249. depauperatum. 1147. lanuginosum. 1250. ovinum. 1148. nitidum. 1251. dichotomum. 1149. microcarpon. 1252. barbulatum. 1150. ovinum. 1253. commutatum. 1151. angustifolium. 1255. huachucae silvicola. 1152. lanuginosum. 1256. . villosissimum, 1153. nitidum. 1258. lanuginosum. 1154. nitidum. 1259. villosissimum. 1155. consanguineum. 1260. xalapense. 1156. lindheimeri. 1261. angustifolium. 1162. strigosum. 1262. dichotomum. 1163. leucothrix. 1263. oligosanthes. 1164. yadkinense. 1264. angustifolium. 1165. lindheimeri. 1267. trifolium. 1171. thurowii. 1268. commutatum. 1172. ovinum. 1269. angustifolium. 1173. helleri. 1270. polyanthes. 1174. roanokense. 1272. boscii. 1175. spretum. 1274. commutatum. 1179. helleri. 12744. oligosanthes. 1181. sphaerocarpon. 1275. villosissimum. 1182. xalapense. 1277. trifolium. 1184. hians. 1278. lindheimeri. 1186. helleri. 1282. hians. 1187. ravenelii. 1283. boscii molle. 1190. sphaerocarpon. 1284. sphaerocarpon. 1192. ovinum. 1285. sphaerocarpon inflatum. * 1193. angustifolium. 1286. commutatum. 1194. oligosanthes. 1287. lindheimeri. 1195. thurowii. 1288. ravenelii. 1202. lindheimeri. 1289. villosissimum. 1203. lindheimeri. 1290. xalapense strictirameum. 1207. joorii. 1291. commutatum. 1208. joorii. 1292. dichotomum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Hircucock, A. 8.—Continued. 1293. 1294. 1295. 1296. 1297. 1298. 1299. 1300. 1301. 1303. 1305. 1306. 1307. 1310. 1311. 1319. 1320. 1321. 1323. 1325. 1327. 1328. 1330. 1331. 1332. 1333. 1334. 1336. 1337. 1339. 1340. 1343. 1344. 1348. 1350. 1352. 1353. 1354. 1356. 1357. 13574. 1358. 1360. 1361. 1362. 1369. 1370. 1371. 1372. 1373. 1375. 1376. 1377. oligosanthes. ravenelii. scribnerianum. scribnerianum. lanuginosum. angustifolium. villosissimum. boscii. boscii molle. depauperatum. trifolium. huachucae silvicola. sphaerocarpon. xalapense. xalapense strictirameum. commutatum. boscii. boscii molle. ashei. wilmingtonense. ashei. commutatum. dichotomum. pseudopubescens. angustifolium. xalapense strictirameum. ravenelii. pseudopubescens. curtifolium. commutatum. angustifolium. microcarpon. boscii molle. commutatum. barbulatum. dichotomum. ashei. depauperatum. pseudopubescens. pseudopubescens. huachucae silvicola. tennesseense. depauperatum. xalapense. angustifolium. albomarginatum. ensifolium. ciliatum. leucothrix. consanguineum. aciculare. nitidum. cryptanthum. Hircscock, A. §.—Continued. 1378. 1379. 1380. 1381. 1382. 1383. 1384. 1387. 1388. 1389. 1390. 1391. 1392. 1393. 1394. 1395. 1398. 1400. 1401. 1402. 1403. 1404. 1405. 1406. 1407. 1408. 1410. 1411. 1412. 1413. 1414. 1415. 1416. 14164. 1419. 1420. 1421. 1422. 1423. 1424. 1425. 1426. 1427. 1428. 1429. 1430. 1431. 1432. 1433. 1434. 1435. 1436. 14364. scabriusculum. ensifolium. pseudopubescens. sphaerocarpon. consanguineum. pseudopubescens. villosissimum. trifolium. ashel. nitidum. ravenelil. xalapense. nitidum. microcarpon. pseudopubescens. lanuginosum. leucothrix. dichotomum. pseudopubescens. pseudopubescens. dichotomum. aciculare. ensifolium. ashei. ashei. angustifolium. oligosanthes. microcarpon. boscii. commutatum. commutatum. xalapense. barbulatum. yadkinense. microcarpon. nitidum. nitidum. nitidum. nitidum. xalapense. ensifolium. sphaerocarpon. chamaelonche. albomarginatum. albomarginatum. ciliatum. lancearium. pauciciliatum. webberianum. albomarginatum. chamaelonche. chamaelonche. ensifolium, 365 366 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Hitcucocrk, A. 8.—Continued. 1437. 1438. 1439. 1440. 1442. 1447, 1449, 1450. 1451. 1455. 1457. 1458. 1459. 1460. 1461. 1463. 1465. 1466. 14664. 1467. 1468. 1469. 1470. 1471. 1472. 1473. 1474, 1475. 1476. 1477. 1479. 1482. 1483. 1484. 1485. 1486. 1487. 1488. 1489. 1490. 1603. 1604. 1605. 1606. 1607. 1608. 1609. 1611. 1612. 1613. 1615. 1616. 1617. pseudopubescens. tenue. ensifolium. albomarginatum. lucidum. aciculare. consanguineum. strigosum. ciliatum. mattamuskeetense. oligosanthes. commutatum. oligosanthes. oligosanthes. ashel. ravenelii. mutabile. lancearium, angustifolium. tenue. lanuginosum. cryptanthum. lucidum. consanguineum. webberianum. aciculare. pseudopubescens. angustifolium. pseudopubescens. pauciciliatum. pauciciliatum. auburne. chamaelonche. pseudopubescens. pseudopubescens. lancearium. pauciciliatum. chamaelonche. ravenelii. ciliatum. lindheimeri. boscii. sphaerocarpon. microcarpon. sphaerocarpon. villosissimum. polyanthes. microcarpon. albemarlense. lanuginosum. sphaerocarpon. sphaerocarpon. addisonii, Hircscock, A. 8.—Continued. 1618. 1619. 1621. 1622. 1623. 1624. 1626. 1627. 1628. 1629. 1630. 1631. 1632. 1635. 1636. 1637. 1638. 1639. 1640. 1641. 1643. 1644. 1902. 2061. 2086. 2114. 2171. 2190. 2204. 2205. 2275. 2374. 2380. 2383. 2391. 2395. 2398. 2399. 2403. 2404. 2407. 2408. 2409. 2410. 2411. 2412. 2413. 2414. 2415. 2418. 2500. 2501. 2502. sphaerocarpon inflatum. villosissimum. huachucae silvicola. lindheimeri. villosissimum. huachucae silvicola. sphaerocarpon. villosissimum. albemarlense. meridionale. huachucae silvicola. villosissimum. columbianum. villosissimum. meridionale. scoparium. lanuginosum. sphaerocarpon. ashei. barbulatum. villosissimum. villosissimum. thermale. thermale. thermale. barbipulvinatum. pacificum. pacificum. barbipulvinatum. barbipulvinatum. barbipulvinatum. barbipulvinatum. capillare. scribnerianum. virgatum. meridionale. agrostoides. polyanthes. villosissimum. polyanthes. columbianum. sphaerocarpon. albemarlense. ashei. barbulatum. sphaerocarpon inflatum. columbianum. depauperatum. villosissimum. columbianum. praecocius. perlongum. scribnerianum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Hircucocr, A. 8.—Continued. 2504. 2505. 2506. 2509. 2511. 2518. 2519. 2523. 2524. 2525. 2528. 2789. 2798. 2836. 3069. 3070. 3071. 3072. 3073. 3074. 3077. 3214. 3219. 3232. 3233. 3234. 3481. 3482. 3494. 3495. 3509. 3526. 3541. 3542. 3544. 3547. 3553. 3561. 3562. 3573. 3598. 3604. 3631. 3637. 3646. 3658. 3661. 3675. 3694. 3695. 3706. leibergii. wilcoxianum. linearifolium. perlongum. scribnerianum. leibergii. scribnerianum. huachucae silvicola. praecocius. scribnerianum. huachucae. barbipulvinatum. barbipulvinatum. scribnerianum. barbipulvinatum. pacificum. pacificum. shastense. pacificum. scribnerianum. pacificum. pacificum. pacificum. pacificum. pacificum. pacificum. hirticaule. arizonicum. hirticaule. fasciculatum chartagi- nense. hirticaule. hirticaule. hirticaule. arizonicum. praecocius. hirticaule. hirticaule. fasciculatum chartagi- nense. arizonicum. hirticaule. fasciculatum. hirticaule. hirticaule. arizonicum. obtusum. hirticaule. obtusum. hirticaule. plenum. arizonicum. hallii. 367 Hitexcocr, A. §.—Continued. 3716. bulbosum sciaphilum. 3730. hirticaule. 3737. obtusum. 3760. barbipulvinatum. 3762. hallii. 3763. obtusum. 3783. hallii. 3784. bulbosum sciaphilum. 3819. barbipulvinatum. 3830. obtusum. 3838. dichotomiflorum. 3840. virgatum. 3842. capillare. 3851. capillare. 3853. praecocius. 3854. scribnerianum. 3865. nodatum. 3866. firmulum. 4932. capillare. 5050. barbipulvinatum. 5083. barbipulvinatum. 5099. capillare. 5111. capillare. 5712. decolorans. 5822. decolorans. Hrircucocx, A.S., LEE Co. Pt. 147. adspersum. 148. adspersum. 159. adspersum. « 469. erectifolium. 470. webberianum. 471. joorii. 472. chamaelonche. 473. longiligulatum. 474, ovale. 477. joorii. 478. nitidum. 479. nitidum. 480. xalapense. 481. lucidum. 482. polycaulon. 483. bartowense. 484. fasciculatur. 485. adspersum. 487. chapmani. 488. hians. 489. hians. 490. rhizomatum. 491. virgatum. 492. tenerum. 611. adspersum. 618. adspersum. 620. adspersum. 650. adspersum. 368 Hitcucock, A. 570a. 571. d71a. 571b. 572. 876. 877. 879. 880. 882. 883. 921. Houssg, H. D. DAE 358. 413. 450. 831. 907. Oi: 947. 949. 957. 961. 1041. 1058. 1136. Sie 1287. 1443. 2105. 2106. 2132. 2136. 2158. 2161. 2177. 2179. 2200. 2211. 2258. 2269. 2305. 2387. 2413. 2430. 2492. 2551. 2575. 2668. JENMAN, G. 3969. 4081. Sf (PL. Kane virgatum. scribnerianum. scribnerianum. leibergii. obtusum. anceps. agrostoides. wilcoxianum. linearifolium. huachucae silvicola. sphaerocarpon. scribnerianum. anceps. virgatum. philadelphicum polyanthes. scribnerianum. tsugetorum. yadkinense. depauperatum. dichotomum. meridionale. ashei. aculeatum. sphaerocarpon. huachucae silvicola. implicatum. tsugetorum. stipitatum. ashei. commutatum. clandestinum. boscil. villosissimum. depauperatum. xalapense. commutatum. ravenelii. boscii molle. meridionale. villosissimum. commutatum. commutatum. sphaerocarpon. ashei. sphaerocarpon. xalapense strictirameum. virgatum cubense. virgatum cubense. geminatum. stoloniferum. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. JENMAN, G.—Continued. 4403. 4438. 5969. 5978. 5998. 6001. 6008. 6009. 6022. 6024. JeRmy, G. 22. 39. 57. 203. 219. 233. 234. 787. Jongs, M. E. 619. 2294. 4019. 4168. 4212. 6035. 6069. KEARNEY, T. a. 25. 26. trichoides. geminatum. pilosum. rudgei. barbinode. zizanioides., laxum. laxum. geminatum. reptans. obtusum. fasciculatum chartagi- nense. reverchoni. huachucae silvicola. fasciculatum chartagi- nense. virgatum. reverchoni. reverchoni. virgatum. tennesseense. pacificum. bulbosum sciaphilum. obtusum. hirticaule. barbipulvinatum. tennesseense. Hi: scoparium. dichotomum., sphaerocarpon inflatum. dichotomiflorum. columbianum thinium. sphaerocarpon. polyanthes. anceps. repens. dichotomiflorum. lindheimeri. anceps. polyanthes. virgatum. aciculare. clandestinum. huachucae silvicola. ciliatum. villosissimum. sphaerocarpon. lindheimeri. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Kearney, T. H.—Continued. 27. 28. 61. 67. 72. 74, 75. 80. 82. 87. 88. 92. 96. 108. 116. 120. 121. 138. 139. 140. 141. 145. 146. 147. 151. 156. 41616°—vo1 15—10——24 albemarlense. scabriusculum. angustifolium. yadkinense. villosissimum. tennesseense. hians. commutatum. clandestinum. lindheimeri. huachucae silvicola. ashei. commutatum. villosissimum. barbulatum in part. scabriusculum. lucidum. wrightianum in part. microcarpon. tenerum. polyanthes. xalapense. ashei in part. barbulatum. polyanthes. huachucae. villosissimum. rhizomatum. scoparium. flexile. agrostoides,. xalapense. capillare. virgatum. anceps. flexile. microcarpon. chamaelonche. polycaulon. rhizomatum. longifolium. verrucosum. hians. rhizomatum. tenerum. patulum. villosissimum. chamaelonche. leucothrix. dichotomum. philadelphicum. scoparium. 369 Kearney, T. H.—Continued. 157. 158. 167. 168. 172. 173. 176. 177. 178. 181. 183. 187. 188. 191. 194. 206. 207. 208. 209. 212. 215. 216. 217. 218. 220. 229. 234. 235. 243. 246. 247. 250. 258. 260. 2674. 267b. 268. 269. 270. 274. 282. 283. 284. 2844. 288. 292. . 293. 298. hemitomon. virgatum. lucidum. virgatum. commutatum. hians. adspersum. chamelonche. lancearium. rhizomatum. scoparium. hians. sphaerocarpon inflatum. oligosanthes. sphaerocarpon inflatum. angustifolium. ashei. dichotomiflorum. commonsianum. xalapense. aciculare. arenicoloides. oligosanthes. sphaerocarpon. consanguineum. huachucae silvicola. longifolium. clandestinum. dichotomum. rhizomatum. sphaerocarpon. wrightianum. aciculare. ciliatum. boscii. lucidum. virgatum. virgatum cubense. albomarginatum. hemitomon. scabriusculum. anceps. strigosum. ciliatum. aciculare. consanguineum. aciculare. lucidum. longiligulatum. amarulum. tenerum. virgatum. 370 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Kearney, T. H.—Continued. Kearney, T. H.—Continued. 299. ashei. 969. stipitatum. 300. dichotomum. 970. xalapense. longiligulatum. 971. villosissimum. 301. pseudopubescens. 972. commutatum. sphaerocarpon. 973. commutatum. 302. villosissimum. 974. polyanthes. 306. aciculare. 1029. ashei. angustifolium. 1033. xalapense. 307. wrightianum. 1104. xalapense. 308. scoparium. 1179. xalapense. 309. lindheimeri. 1307. microcarpon. 312. flexile. 1308. xalapense. 320. sphaerocarpon. 1317. commutatum. 324. ashei. 1369. angustifolium. 325. boscii. 1374. dichotomum. 326. ciliatum. 1375. aciculare. microcarpon. 1386. oligosanthes. 327. commutatum. 1393. commonsianum. 328. huachucae silvicola. 1400. oligosanthes. 329. dichotomum. 1411. _ boscii. 330. clandestinum. 1414. commutatum. lancearium. 1416. angustifolium. patentifolium. 1447. tsugetorum. 331. patulum. 1454. commonsianum. 3314. pauciciliatum. . 1461. oricola. 336. mutabile. 1463. commutatum. verrucosum. 1467. xalapense. 337. villosissimum. 1469. boscii. 342. repens. 1476. polyanthes. 358. rhizomatum. 1477. scoparium. 363. hians. 1514. roanokense. 367. dichotomum. 1559. lanuginosum. 369. virgatum. 1560. sphaerocarpon. * 372. agrostoides. 1566. aciculare. anceps. 1748. anceps. dichotomiflorum. 1761. strigosum. 374. dichotomiflorum. 1775. amarulum. 376. anceps. 1776. commonsianum. 378. gattingeri. 1798. scabriusculum. 379. verrucosum. 1871. aciculare. 380. longifolium. 1899. virgatum cubense. 380a. stipitatum. 2018. virgatum. 384. dichotomum. 2021. amarulum. 594. sphaerocarpon. 2025. longifolium. 960. flexile. 2026. roanokense. 961. philadelphicum. 2038. aciculare. 962. gattingeri. 2043. lanuginosum. 963. gattingeri. 2053. verrucosum. 965. anceps. 2063. amarulum. 966. dichotomiflorum. 2064. amarum. 967. microcarpon. 2114. commonsianum, 968. sphaerocarpon, 2242. longifolium. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 3871 Kearney, T. H.—Continued. KNEUCKER, A., GRAM. Exs.—Continued. 2249. stipitatum. 423. latifolium. 2272. aciculare. 424. clandestinum. 2317. amarum. 425. scribnerianum. KELLERMAN, W. A. 485. tennesseense. 20. praecocius. 546. dichotomiflorum. 4725. maximum. 547. perlongum. 5114. polygonatum. 548. depauperatum. 5119. pilosum. 549. xalapense. 6231. viscidellum. 550. microcarpon. 6236. sphaerocarpon. 551. huachucae silvicola. 6246. pulchellum. 552. lindheimeri. 6249. viscidellum. 553. sphaerocarpon. 6250. geminatum. 554. polyanthes. 6253. paludivagum. 555. villosissimum. 6254. paludivagum. 556. ashei. 6263. trichoides. 557. commutatum. 6267. maximum. LEIBERG, J. B. 6272. trichanthum. 714. barbipulvinatum. 6758. huachucae silvicola. 834. barbipulvinatum. 6765. capillare. 1312. pacificum. 6766. latifolium. 5732. virgatum. 6767. polyanthes. 5783. barbipulvinatum. 6768. dichotomum. 5816. bulbosum sciaphilum. 6769. huachucae. 5916. hallii. 6778. microcarpon. Lemmon, J. G. 6785. huachucae. 353. arizonicum. 6799. latifolium. 2907. huachucae. 67994. clandestinum. 2908. bulbosurh sciaphilum. 6800. stipitatum. 2912. bulbosum. 6873. huachucae. 2914. bulbosum. 6881. commutatum. 2916. bulbosum. 6883. clandestinum. 2922. bulbosum sciaphilum. 6885. implicatum. 3062. arizonicum. 6886. barbulatum: 3152. bulbosum sciaphilum. 6887. bicknellii. 3154. virgatum. 6888. clandestinum. Leon, BroTHeER. 6890. huachucae silvicola. 190. diffusum. 6891. dichotomum. 276. reptans. 6892. ashei. 283. barbinode. 6893. huachucae. 291. adspersum. 6894. scribnerianum. 292. reptans. 6895. huachucae silvicola. 296. repens. 6899. latifolium. 297. reptans. 6900. clandestinum. 305. diffusum. 6901. ashei. 305b. distantiflorum. 6902. commutatum. 335. elephantipes. 6903. barbulatum. 427. maximum. Knevucker, A., Gram. Exs. 557. trichanthum. 189. capillare. 563. repens. 245. dichotomum. 566. reptans. 302. virgatum. 567. distantiflorum. 366. molle. 568. barbinode. 372 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Lre6n, BrotHer—Continued. LINDHEIMER, F. 570. adspersum. 158. pedicellatum. 573. fasciculatum. 565. lindheimeri. 576. reptans. 728. condensum. 813. fasciculatum. 733. virgatum. 902. exiguiflorum. 1265. pedicellatum. 903. zizanioides. MackeEnzigz, K. K. 906. reptans. 297. leibergii. 907. laxum. 301. agrostoides. 908. pilosum. ~ 460. lucidum. 909. laxum. 1339. linearifolium. 910. reptans. 1355. oricola. 910b. reptans. 1360. clandestinum. 910c. reptans. 1379. tsugetorum. 911. exiguiflorum. 1380. polyanthes. 912. distantiflorum. 1381. villosissimum. 913. exiguiflorum. 1398. linearifolium. 914. laxum. 1399. columbianum. 916. fasciculatum. 1405. dichotomum. 917. distantiflorum. 1414. latifolium. 918. geminatum. 1452. depauperatum. 919. dichotomiflorum. 1459. tennesseense. 920. geminatum. 1484. microcarpon. 921. maximum. 1485. commonsianum. 922. ghiesbreghtii. 1548. barbulatum. 922b. ghiesbreghtii. 1605. microcarpon. 923. diffusum. 1611. scribnerianum. 923b. diffusum. 1663. sphaerocarpon. 923c. diffusum. 1664. aciculare. 924. adspersum. 1671. verrucosum. 925. adspersum. 1686. microcarpon. 957. maximum. 1688. scoparium. LizBMANN, F. M. 1708. ashei. 275. millegrana. 1709. latifolium. 277. fasciculatum. 1710. villosissimum. 279. fasciculatum. 1725. amarulum. 317. trichoides. 1733. auburne. 320. trichanthum. 1736. amarum. 323. viscidellum. 1745. polyanthes. 324. viscidellum. 1794. joorii. 327. sphaerocarpon. 1854. scoparium. 328. xalapense. 1893. stipitatum. 394. zizanioides. 2067. commonsianum. 405. frondescens. 2068. columbianum. 411. pilosum. 2075. huachucae. 412. laxum. 2076. tennesseense. 419. laxum. 2093. boreale. 425. maximum in part. 2105. columbianum. 428. gelutinosum. 2106. dichotomum. 432. trichanthum. 2109. boreale. 441. bulbosum. 2137. huachucae silvicola. 442. bulbosum. 2138. annulum. 450. gouini. 2144. boscii. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Mackenziz, K. K.—Continued. 2155. commonsianum. 2156. lindheimeri. 2159. huachucae silvicola. 2160. lindheimeri. 2161. microcarpon. 2163. ashei. 2164. lindheimeri. 2165. commonsianum. 2166. sphaerocarpon. 2167. lucidum. 2168. lindheimeri. 2169. huachucae silvicola. 2170. meridionale. 2195. ashei. 2197. huachucae silvicola. 2220. tsugetorum. 2247. columbianum. 2249. huachucae silvicola. 2250. sphaerocarpon. 2251. werneri. 2280. dichotomum. 2349. lindheimeri. 2355. agrostoides. 2366. philadelphicum. 2404. columbianum. 2476. microcarpon. 2477. meridionale. 2480. ashei. Macovn, J. anp J. M. 135. agrostoides. 7444. philadelphicum. 13003. virgatum. 13225. capillare. 13227. wilcoxianum. 13231. virgatum. 21957. werneri. 22022. boreale. 22023. subvillosum. 22024. implicatum. 22025. xanthophysum. 26236. tsugetorum. 26316. lindheimeri. 26317. capillare. 26322. depauperatum. 26324. virgatum. 26325. latifolium. 26326. latifolium. 26327. latifolium. 26328. scribnerianum. 26329. scribnerianum. 26330. flexile. 26331. flexile. Macouwn, J. anp J. M.—Continued. 26332. 26333. 26334. 26337. 26338. 29297. 29303. 29304. 29348. 29349. 29368. 29369. 65370. 69204. 69205. 72965. 73003. 73004. 77229. 77230. : Mize. Maxon, W. R. 550. 1659. 2109. 2361. 2816. 3153. 3476. Mearns, E. A. 25. 738. 743. 755. 756. 758. 767. 769. (ise ‘is. 788. 791. 793. 925. 1072. 1130. 1845. 1905. 1932. 2093. 2294. 2308. flexile. scribnerianum. huachucae silvicola. huachucae. lindheimeri. scribnerianum. virgatum. virgatum. agrostoides. boreale. subvillosum. huachucae. lindheimeri. boreale. subvillosum. tennesseense. leibergii. leibergii. occidentale. scribnerianum. scribnerianum. huachucae silvicola. fasciculatum. zizanioides. fasciculatum. glutinosum. pulchellum. barbinode. ’ villosissimum. hirticaule. barbipulvinatum. barbipulvinatum. capillare. virgatum. hirticaule. virgatum. scribnerianum. obtusum. barbipulvinatum. latifolium. virgatum. obtusum. arizonicum. obtusum. obtusum. hirticaule. 373 bulbosum sciaphilum. hirticaule. hirticaule. obtusum. 374 Mearns, E. A.—Continued. 2501. bulbosum. 3061. thermale. 4050. thermale. 4166. thermale. 4203. thermale. 4789. thermale. 4870. thermale. 4983. thermale. 5064. thermale. 5110. thermale. 5134. thermale. Merri, E. D. 2. implicatum. 7. implicatum. 8. implicatum.’ 9. implicatum. 11. xanthophysum. 16. implicatum. 43. barbipulvinatum. 152. barbipulvinatum. 157. thermale. 164. thermale. 165. thermale. 197. microcarpon. 198. clandestinum. 199. polyanthes. 202. boscii molle. 203. boscii molle. 204. commutatum. 233. scoparium. 1243. Jatifolium. Mercatre, O. B. 6. plenum. 80. bulbosum. plenum. 354. helleri. 357. bulbosum sciaphilum. 434. barbipulvinatum. 738. plenum. 739. plenum. 749. obtusum. 768. arizonicum. 807. hallii. 1294. arizonicum. 1422. bulbosum. 1442. hirticaule. MituspaueanH, C. F. 126. dichotomiflorum. 324. barbinode. 454. maximum. 702. utowanaeum. 726. reptans. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. MitispaueH, C. F.—Continued. 12. "1859. 2182. Morona, T. SLic 405a. 441. 519. 534. 536. 537. 543. 779a. 813. 977. 1001. 1002. 1072. 1571. 1574. Morris, E. L. 9. 48. 53. » 55. 124. 135. g/l 226. 240. 252. 287. 294. 977. 984. 1139. 1186. 1193. 1198a. 1283. Nasu, G. V. 15. 36. 45. 50. dl. 52. 63. barbinode. pilosum. caerulescens. trichanthum. glutinosum. polygonatum. parvifolium. laxum. zizanioides. laxum. dichotomiflorum. barbinode. megiston. laxum. zizanioides. dichotomiflorum. elephantipes. megiston. trichanthum. laxum. huachucae. huachucae silvicola. virgatum. implicatum. virgatum. villosissimum. capillare. capillare. pseudopubescens. tsugetorum. capillare. dichotomiflorum. virgatum. ashei. lindheimeri. microcarpon. polyanthes. microcarpon. dichotomum. philadelphicum. pauciciliatum. malacon. equilaterale. commutatum. patulum. chamaelonche. patentifolium. malacon. pauciciliatum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Nasu, G. V.—Continued. 71. 72. 75. 78. 103. 182. 147. 151. 213. 239. 240. 273. 287. 301. 302. 334. 335. 337. 372. 375. 1012. 1117. 1118. 1119. 1120. 1226. 1238. 1243. 1337. 1338. 1436. 1507. 1518. 1674. 1675. 1694. chamaelonche. patentifolium. ovale. commutatum. ovale. malacon. ovale. patulum. hians. laxiflorum. commutatum. joorii. vernale in part. tsugetorum. lancearium. commutatum. leucothrix. chamaelonche. lucidum. dichotomiflorum. lanuginosum. nitidum. vernale. lancearium. leucothrix. lucidum. arenicoloides. malacon. malacon. hemitomon. paludivagum. chamaelonche. verrucosum. webberianum. ciliatum. dichotomiflorum. albomarginatum. erectifolium. patulum. ovale. commutatum. equilaterale. fusiforme. chamaelonche. aciculare. pauciciliatum. leucothrix. arenicoloides. curtifolium. ovale. equilaterale. commutatum. agrostoides. Nasu, G. V.—Continued. 375 1713. rhizomatum. 1730. maximum. 1856. fusiforme. 1857. ovale. 2034. laxiflorum. 2075. leucothrix. 2076. pauciciliatum. 2156. laxiflorum. 2204. scoparium. 2249. tenerum. 2258. rhizomatum. 2329. boscii. 2500. sphagnicola. 2522. microcarpon. 2529. anceps. NeEtson, A. 516. scribnerianum. 2524. scribnerianum. 3626. virgatum. 6037. thermale. 6174. thermale. ’ 8346. barbipulvinatum. 8360. virgatum. Netson, E. 330. virgatum. 476. virgatum. 481. barbipulvinatum. 4984. barbipulvinatum. Netson, E. W. 201. xalapense. 1374. bulbosum. 1622. reptans. 2874. fasciculatum. 2958. fasciculatum. 2975. virgatum. 3023. zizanioides. 3056. pilosum. 3357. glutinosum. 3781. biglandulare. 3908. obtusum. 4257. trichoides. 6187. bulbosum. 6297. hirticaule. 6298. bulbosum sciaphilum. 6301. bulbosum. 6352. obtusum. 6355. hirticaule. PALMER, E. lb in 1885. arizonicum. hirticaule. le in 1885. sonorum. 14 in 1897. hirticaule. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 158 in 1887. 159 in 1887. 159 in 1887. 168a in 1887. 175 in 1896. 206 in 1887. 207 in 1886. 207 in 1887. 207a in 1886. 208 in 1887. 241 in 1897. 249 in 1897. 250 in 1897. 251 in 1897. 266 in 1904. 287 in 1895. - 289 in 1894. 340 in 1906. 346 in 1887. 348 in 1906. 349 in 1906. 370 in 1868. 375 in 1868. 376 in 1868. 380 in 1868. 381 in 1868. 382 in 1868. 383 in 1868. 384 in 1868. 394 in 1898. 412 in 1907. 429 in 1886. 467 in 1896. 469 in 1906. 510 in 1886. 525 in 1896. 525a in 1896. PatmerR, E.—Continued. 15 in 1869. 19 in 1897. 143 in 1897. 145 in 1897. 149 in 1897. 152 in 1904. geminatum. fasciculatum. hirticaule. hirticaule. molle. fasciculatum char- taginense. fasciculatum. arizonicum. fasciculatum char- taginense. stramineum. obtusum. stramineum. bulbosum. virgatum. fasciculatum. bulbosum. hirticaule. fasciculatum. hirticaule. arizonicum. hirticaule. obtusum. trichoides. geminatum. bulbosum. hirticaule. bulbosum scia- philum. bulbosum scia- philum. obtusum. dichotomiflorum. virgatum. anceps. hians. helleri. malacophyllum. lindheimeri. obtusum. fasciculatum char- taginense. paludivagum. bulbosum scia- philum. bulbosum. bulbosum scia- philum. virgatum. lepidulum. bulbosum. 554 in 1907. 561 in 1890. 590 in 1898. 631 in 1874. 632 in 1874. 633 in 1874. 634 in 1874, 690 in 1887. 690 in 1890. 694 in 1890. 695 in 1890. 741 in 1896. 750 in 1890. 947 in 1889. 1078 in 1890. 1083 in 1890. 1151 in 1879. 1257 in 1891. 1338 in 1880. 1538 in 1891. 1539 in 1891. 1544 in 1891. 1545 in 1891. 1554 in 1891. 1557 in 1891. 1558 in 1891. 1659 in 1891. 1660 in 1891. 1758 in 1891. 2649 in 1892. 2709 in 1892. PatmeEr, E.—Continued. 533 in 1906. 554 in 1906. lepidulum. bulbosum scia- philum. hallii. scribnerianum. obtusum. lancearium. lucidum. sphagnicola. breve. chamaelonche. geminatum. hirticaule. fasciculatum. hirticaule. plenum. hirticaule. sonorum. barbinode. trichoides. trichanthum. trichanthum. hallii. stramineum, sonorum. hirticaule. hirticaule. hirticaule. sonorum. fasciculatum, laxum. parcum. molle. trichoides. barbipulvinatum., barbipulvinatum. Patmer, W., AND Riney, J. H. 178. 218. 377. 447. 481. 542. 545. 746. Kile 802. 816. 982. 989. 990. 1065. maximum. strigosum. maximum. acuminatum. chrysopsidifolium. maximum. maximum. adspersum. adspersum. diffusum. maximum, chrysopsidifolium. acuminatum. polycaulon. viscidellum. PatMER, W., AND Riwey, J. H.—Cont’d. 1069. laxum. 1083. acuminatum. 1086. cayennense. 1134. virgatum cubense. ParisH, S. B. 263. pacificum. 887. urvilleanum. 1081. barbipulvinatum. 1663. pacificum. Parun, J. C. 751. agrostoides. 938. languidum. 1029. boreale. 1181. huachucae silvicola. 1186. huachucae. 1187. boreale. 1188. implicatum. 1189. huachucae. 1190. werneri. 1191. latifolium. 1196. columbianum. 1198. implicatum. 1215. tsugetorum. 1266. dichotomum. 1423. boreale. 1502. werneri. 1577. tsugetorum. 1581. subvillosum. 1607. lindheimeri. 1701. boreale. 1738. boreale. 1744. boreale. 1776. philadelphicum. 1806. werneri. 1957. depauperatum. 1971. linearifolium. 2000. implicatum. 2001. subvillosum. 2013. boreale. 2016. subvillosum. 2017. tennesseense. 2034. boreale. Pirrier, H. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 101. paludivagum. 257. strigosum. 361. laxum. 364. stoloniferum. 521. laxum. 553. laxum. 665. laxum. 932. acuminatum. 940. viscidellum. 377 Pirtier, H.—Continued. 971. trichoides. 982a. acuminatum. 1617. maximum. 1621. trichoides. 1800. strigosum. 1805a. albomarginatum. 1805b. arenicoloides. 1960. maximum. Poutarp, C. L. 16. dichotomum. 92. clandestinum. 180. ashei. 323. barbulatum. 324. dichotomum. 337. villosissimum. 338. depauperatum. 353. meridionale. 362. microcarpon. 365. clandestinum. 398. sphaerocarpon. 401. polyanthes. 403. lucidum. 406. microcarpon. 408. microcarpon. 412. boscii molle. 523. tennesseense. 595. virgatum. 682. dichotomiflorum. 1106. rhizomatum. 1152. repens. 1201. virgatum cubense. 1228. anceps. PRINGLE, C. G. 7. plenum. 26. barbinode. 73. reptans. 74. fasciculatum. 124. fasciculatum. 376. hallii. 377. bulbosum. 379. fasciculatum chartagi- nense. 380. fasciculatum chartagi- nense. 465. arizonicum. 476. obtusum. 487. arizonicum. 497. lepidulum. 1124. havardii. 1415. vaseyanum. 1406. bulbosum sciaphilum. 2377. ramisetum. 378 PRINGLE, C. G.—Continued. 3336. 3449. 3817. 3828. 5203. 5207. 5569. 5573. 6322. 6418. 7882. 7883. 8083. 8089. 8195. 8323. 8339. 8344. 9209. 9210. 9556. 9575. 9577. 13250. Purpus, C. A. 2156. 2159. 2160. 2902. 2903. 2908. 3774. REVERCHON, J. 22. paludivagum. cupreum. schmitzii. trichoides. albomaculatum. bulbosum. gouini. hirsutum. elephantipes. bulbosum. multirameum. sphaerocarpon. xalapense. viscidellum. longum. ramisetum. multirameum. olivaceum. sphaerocarpon. multirameum. multirameum. paludivagum. bulbosum. elephantipes. xalapense. glutinosum. laxum. laxum. viscidellum. trichoides. glutinosum. bulbosum. maximum. virgatum. virgatum. geminatum. ovinum. xalapense. hians. agrostoides, anceps. helleri in part. huachucae silvicola. geminatum. obtusum. agrostoides. fasciculatum chartagi- nense. ovinum. reverchoni. texanum. CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, REVERCHON, J.—Continued. 1620. 1622. 1682. 1831. 1840. 1841. 1842. 1884. 2222. 2223. 2224. 2228. 2235. 2341. 2342. 2344. 2345. 2357. 2368. 2390. 2444. 2844. 2855. 3526. 4136. 4137. 4138. 4142. 4148. 4144b. 4147. 4155. 4156. 4158. 4159. 4163. 4193. 4194. Ricker, P. L. 666. 908. 933. 936. 943. 945. 952. 962. 963. 975. 1153. 1277. 12774. 1309. pedicellatum. virgatum. hallii. malacophyllum. oligosanthes. oligosanthes. philadelphicum. lanuginosum. agrostoides. brachyanthum. brachyanthum. texanum. gymnocarpon. hemitomon. helleri. helleri. helleri. lindheimeri. condensum. ravenelii. helleri. virgatum. helleri. filipes. villosissimum. ovinum. helleri. oligosanthes. commutatum. boscii. barbulatum. dichotomum. consanguineum. dichotomum. angustifolium. thurowii. angustifolium. microcarpon. huachucae silvicola, tenerum. rhizomatum. virgatum cubense. equilaterale. patentifolium. adspersum. rhizomatum. patentifolium in part. capillare. commutatum. implicatum. subvillosum. dichotomum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. RIcKsEcKeER, A. E. RugeeEt, F.—Continued. 379 66. adspersum. 444. amarulum. 77. reptans. 595. virgatum. 200. maximum. 598. verrucosum. 212. geminatum. 599. gymnocarpon. 300. barbinode. Russy, H. H. 317. fasciculatum. 22. laxum. 384. adspersum. 199. polygonatum in part. 413. maximum. 210. laxum. Riepet, L. pilosum. 52. laxum. 212. pilosum. 53. maximum. 217. pulchellum. 958. parvifolium. 228. laxum. 959. dichotomiflorum. 229. frondescens. 1239. megiston. 233. millegrana. 1360. trichanthum. 236. laxum. Ross, J. N. 244. glutinosum. 1834. fasciculatum. 444. pampinosum. 1878. stramineum. 445. virgatum. 1883. stramineum. 445b. bulbosum. 1884. fasciculatum. 445c. bulbosum sciaphilum in 1889. stramineum. part. 1999. bulbosum. 864. obtusum. 2053. bulbosum sciaphilum. 866. bulbosum sciaphilum. 2280. arizonicum. 8921. obtusum. 2609. bulbosum. Russy, H. H., anp Squires, R. W. 3281. stramineum. 79. pilosum. 3351. hirticaule. 347. pilosum. 3361. bulbosum. 355. megiston. Rovirosa, J. N. 362. rudgei. 427. laxum. Ruta, A. 434. trichoides. 1. sphaerocarpon. 497. frondescens. 5. polyanthes. 532. megiston. 6. sphaerocarpon. 598. trichoides. 7. polyanthes. 599. pilosum. ll. commutatum. 624. zizanioides. 15. polyanthes. Rueaet, F. 19. villosissimum. 123. geminatum. 21. sphaerocarpon. 142. consanguineum. 56. angustifolium. 184. albomarginatum. 57. microcarpon. 229. erectifolium. 59. capillare. 231. hians. gattingeri. 290. neuranthum. 60. clandestinum. 291. lancearium. 61. commutatum. 347. hemitomon. 63. depauperatum. 351. joorii. 64. microcarpon. 376. lancearium. 65. agrostoides. 377. chamaelonche. 66. flexile. 378. jooril. 68. xalapense. 392. xalapense. 69. polyanthes. 394. chapmani. 70. boscii molle. 443. webberianum. 71. dichotomiflorum. 880 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Ruts, A.—Continued. 72. villosissimum in part. 75. sphaerocarpon. 76. virgatum. Rypsere, P. A. 1096. barbipulvinatum. 1097. virgatum. 1098. scribnerianum. 1099. huachucae. 1100. perlongum. 1279. scribnerianum. 1308. wilcoxianum. 1368. huachucae. 1493. scribnerianum. 1538. barbipulvinatum. 1561. virgatum. 1604. scribnerianum. 1788. barbipulvinatum. 2011. barbipulvinatum. 2351. barbipulvinatum. 2505. barbipulvinatum. 2508. virgatum. 2512. virgatum. 2516. virgatum. ScHAFFNER, S. W. 138. sphaerocarpon. 146. pseudopubescens. 148. obtusum. 284. viscidellum. 285. sphaerocarpon. 1037. sphaerocarpon. SHEAR, C. L. 85. scribnerianum. 152. barbipulvinatum. 223. virgatum. 264. barbipulvinatum. 436. barbipulvinatum. 606. virgatum. 755. barbipulvinatum. 756. virgatum. 767. virgatum. 819. virgatum. 965. barbipulvinatum. 975. obtusum. 980. virgatum. Smntenis, P. 51. maximum. 160. trichoides. 355. acuminatum. 357. glutinosum. 360. laxum. 847. reptans. 938. elephantipes. 1216. parvifolium. Sintenis, P.—Continued. 1224. acuminatum in part. 1254. laxum. 1901. fasciculatum. 1957. adspersum. 2468. maximum. 2471. trichanthum. 2609. glutinosum. 3365. utowanaeum. 3366. maximum. 3367. geminatum. 3368. reptans. 3416. utowanaeum. 3463. utowanaeum. 3647. fasciculatum. 4983. ghiesbreghtii. 5719. parvifolium. 5724. polycaulon. 5908. acuminatum. 5985. chrysopsidifolium. Sma., J. K., AND HELLER, A. A. 201. commutatum. 204. scoparium. 205. hians. 279. depauperatum. 348. clandestinum. 394. latifolium. 463. polyanthes. 480. dichotomum. SmitH, H. H. 151. geminatum. 167. trichoides. 169. zizanioides. 173. reptans. 202. laxum. 203. pilosum. 204. laxum. 206. polygonatum. 211. barbinode. 589. pulchellum. 1409. maximum. 2146. millegrana. 2151. trichanthum. 2153. maximum. 2190. polygonatum. Somss, M. P. 25. praecocius. 153. tennesseense. 167. scribnerianum. 189. virgatum. 207. huachucae silvicola. 210. huachucae. 219. virgatum. 229. scribnerianum. ae HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Somes, M. P.—Continued. 230. 231. 232. 236. 245. 246. Spruce, R. 2344. Suxsporr, W. 124. 2330. 5162. 5174. 6292. THIEME, C. 195. 532. 781. 5578. 5584. 5587. TiwestroM, I. 4. 5. 22. 48. 2482. 161. huachucae silvicola. clandestinum. microcarpon. leibergii. perlongum. praecocius. rudgei. trichoides. millegrana. parvifolium. zizanioides. stenodes. parvifolium. zizanioides. N. pacificum, hirticaule. occidentale. occidentale. pacificum. fasciculatum. virgatum. polygonatum. polygonatum. fasciculatum. laxum. pilosum. polygonatum. trichanthum. ashei. tennesseense. barbipulvinatum. huachucae silvicola. barbipulvinatum. pacificum, boscii. sphaerocarpon. tenerum. joorii. repens. scribnerianum. leucothrix. ensifolium. sphaerocarpon. dichotomiflorum. spretum. lucidum. joorii. villosissimum. 381 Tracy, S. M.—Continued. 162. 192. 433. 434. 459. 459a. 912. 1410. 1416. 1417. 1418. 1478. 1535. 1730. ae. 1735. 1751. 1752. 1753. 1754. 1755. 1837. 1883. 1884. 1888. 2027. 2028. 2031. 2032. 2036. 2050. 2058. 2854. 2856. 2859. 2861. 2862. 2863. 2865. 2867. 2869. 2873. 3190. 3198. 3204. 3205. 3207. 3208. 3209. 3211. 3223. 3224. 3225. ciliatum. barbipulvinatum. sphaerocarpon. urvilleanum. arenicoloides. arenicoloides. virgatum. xalapense. commutatum. aciculare. hians. xalapense strictirameum. anceps. angustifolium. microcarpon. lanuginosum. huachucae silvicola. scribnerianum. xalapense. oligosanthes. dichotomum. reptans. angustifolium. consanguineum., angustifolium. flavovirens. dichotomum. nitidum.,: xalapense. agrostoides. dichotomum. xalapense. amarum. lanuginosum. neuranthum. wrightianum. sphaerocarpon inflatum. pauciciliatum. trifolium. lanuginosum. lancearium. consanguineum. huachucae. huachucae silvicola. dichotomum. boscii molle. microcarpon. huachucae silvicola. dichotomum. xalapense. huachucae silvicola. microcarpon. microcarpon. 382 Tracy, S. M.—Continued. 3228. 3238. 3252. 3253. 3255. 3256. 3265. 3266. 3267. 3268. 3272. 3273. 3275. 3285. 3287. 3296. 3306. 3316, 3318. 3334. 3387. 3388. 3388a. 3603. 3604. 3614. 3615. 3616. 3617. 3618. 3619. 3620. 3622. 3623. 3624. 3625. 3626. 3627. 3629. 3630. 3631. 3633. 3634. 3635. 3636. 3637. 3638. 3639. nitidum. polyanthes. boscii. dichotomum. xalapense. xalapense. lindheimeri. joorii. xalapense. asheil. villosissimum. dichotomum. ashei. lindheimeri. xalapense. commutatum. sphaerocarpon. scoparium. polyanthes. joorii. trifolium. joorii. microcarpon. microcarpon. rhizomatum. virgatum. fusiforme. lindheimeri. dichotomum. ovale. microcarpon. sphaerocarpon inflatum. lanuginosum. lanuginosum. microcarpon. microcarpon. dichotomum. roanokense. rhizomatum. longifolium. ashei. commutatum. commutatum. polyanthes. consanguineum. aciculare. angustifolium. fusiforme. angustifolium. aciculare. aciculare. aciculare, CONTRIBUTIONS FROM THE NATIONAL HERBARIUM, Tracy, S. M.—Continued. 3640. 3641. 3642. 3643. 3644. 3645. 3646. 3647. 3649. 3650. 3651. 3653. 3655. 3656. 3657. 3658. 3660. 3661. 3746. 3747. 3748. 3749. 3750. 8751. 3752. 3753. 3754. 3756. 3757. 3758. 3759. 3760. 3761. 3762. 3763. 3858. 3859. 3860. 3861. 3889. 3976. 3978. 4562. 4563. 4564. 4565. 4566. 4567. 4568. aciculare. aciculare. sphaerocarpon inflatum. scabriusculum. wrightianum. lanuginosum. mutabile. oligosanthes. dichotomum. aciculare. angustifolium. villosissimum. joorii. sphaerocarpon inflatum. angustifolium. villosissimum. virgatum. consanguineum. angustifolium. ashel. curtifolium in part. lucidum. angustifolium. villosissimum. curtifolium. villosissimum. villosissimum. ashei. dichotomum. angustifolium. villosissimum. xalapense. polyanthes, microcarpon. virgatum. virgatum cubense. virgatum. longifolium. longifolium. longifolium. repens. lindheimeri. huachucae silvicola. patulum. scoparium. verrucosum. longifolium. rhizomatum. virgatum. — gouini. neuranthum., combsii in part, HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Tracy, S. M.—Continued. 4569. 4573. 4574. 4576. 4577. 4578. 4579. 4580. 4581. 4582. 4583. 4584. 4585. 4586. 4587. 4588. 4591. 4592. 4593. 4594. 4595. 4596. 4597. 4598. 4599. 4601. 4603. 4604. 4605. 4606. 4607. 4609. 4610. 4611. 4612. 4614. 4615. 4616. 4617. 4618. 4619. 4620. 4621. 4622. 6350. 6358. 6444. 6446. scabriusculum. commutatum. mutabile. xalapense. sphaerocarpon. commutatum in part. angustifolium. angustifolium. lanuginosum. lancearium in part. ciliatum. trifolium. roanokense. spretum. patulum. patulum. xalapense. nitidum. roanokense. sphaerocarpon inflatum. spretum. microcarpon. erectifolium. sphaerocarpon. ~ eurtifolium. curtifolium. trifolium. trifolium. sphaerocarpon. lanuginosum. albomarginatum. polycaulon. sphaerocarpon. nitidum. microcarpon. chamaelonche. wrightianum. trifolium. consanguineum., angustifolium. angustifolium. ovinum. scabriusculum. dichotomiflorum. rhizomatum. anceps. rhizomatum. sphaerocarpon inflatum. patentifolium. xalapense. condensum. pauciciliatum, 383 Tracy, S. M.—Continued. 6447. 6451. 6452. 6455. 6458. 6460. 6464. 6465. 6466. 6469. 6470. 6471. 6507. 6507a. 6508. 6691. 6692. 6693. 6694. 6695. 6698. 6699. 6700. 6701. 6702. 6703. 6707. 6708. 6710. 6711. 6713. 6713a. 6714. 6715. 6716. 6723. 6725. 6726. 6727. 6729. 6730. 6731. 6732. 6733. 6736. 7008. 7018. 7029. 7048. 7050. pauciciliatum. chamaelonche. lancearium. fasciculatum. sphaerocarpon inflatum. equilaterale. chamaelonche. lancearium. lancearium. lanuginosum. polycaulon. sphaerocarpon inflatum. dichotomiflorum. longifolium. amarulum. bartowense. polycaulon. chamaelonche. xalapense. commutatum. polycaulon. vernale. malacon. patulum. patentifolium. patulum. laxiflorum. fusiforme. fusiforme. polycaulon. neuranthum. fusiforme. equilaterale. fusiforme. lancearium. glabrifolium. webberianum. polycaulon. malacon. chamaelonche. polycaulon. patulum. lancearium. hemitomon. chamaelonche. albomarginatum. lindheimeri. hians. microcarpon. trifolium. scoparium. equilaterale. 384 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. Tracy, 8. M.—Continued. 7051. 7093. 7105. 7163. 7166. 7167. 7170. 7174. 7175. 7176. 7180. 7186. 7188. 7189. 7191. 7193. 7194. 7195. 7198. 7199. 7200. 7202. 7203. 7205. 7207. 7208. 7209. 7371. 7381. 7382. 7383. 7387. 7392. 7400. 7402. 7405. 7409. 7412. 7651. 7738. 7740. 7745. 7748. 7753. 7763. 7935. 7939. 7940. 7941. 7942, equilaterale. lancearium. rhizomatum. rhizomatum. fusiforme. neuranthum. equilaterale. nitidum. patentifolium. lancearium. neuranthum. vernale. webberianum. flavovirens. polycaulon. albomarginatum. chamaelonche. leucothrix. longiligulatum. lancearium. lancearium. patulum. nitidum. chamaelonche. xalapense. lancearium. lancearium. chamaelonche. polycaulon. lancearium. fusiforme. geminatum. commutatum. laxiflorum. reptans. geminatum. gymnocarpon. sphaerocarpon. rhizomatum. capillare. paludivagum. agrostoides. bartowense. bartowense. hians. texanum. gouini. barbinode. obtusum. reverchoni. reverchoni. hallii. malacon. Tracy, S. M.—Continued. 7943. 7944. 7945. 7946. 7947. 7948. 7949. 7950. 7952. 7953. 7954. 7955. 7958. 8029. 8200. 8224. 8229. 8289. 8290. 8295. 8396. 8397. 8398. 8399. 8400. 8401. 8402. 8403. 8405. 8406. 8407. 8408. 8409. 8410. 8411. 8412. 8413. 8414. 8415. 8416. 8417. 8418. 8419. 8420. 8421. 8422. 8423. 8424. 8425. 8426. praecocius. villosissimum. lindheimeri. hallii. sphaerocarpon. lindheimeri. reverchoni. helleri. hallii. hallii. hallii. hallii. ciliatissimum. ramisetum. yadkinense. hallii. plenum. ramisetum. fasciculatum chartagi- nense. texanum. barbipulvinatum. gymnocarpon. pauciciliatum. longiligulatum. lanuginosum. sphaerocarpon inflatum. caerulescens. strigosum. barbulatum. lucidum. pauciciliatum. chamaelonche. combsii. lancearium. leucothrix. polycaulon. pauciciliatum. longiligulatum. anceps. villosissimum. lanuginosum microcarpon polyanthes. helleri. aciculare. curtifolium. strigosum. longiligulatum. mutabile. lancearium. pseudopubescens. villosissimum. HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. Tracy, S. M.—Continued. 8427. . 8428. 8429. 8430. 8431. 8432. 8433. 8504. 8591. 41616°—vo.t 15—10——25 consanguineum. equilaterale. wilmingtonense. auburne in part. nudicaule. nudicaule. spretum. lucidum. scoparium. hemitomon. xalapense. nitidum. patulum. villosissimum. trifolium. microcarpon. commutatum. xalapense. boscii molle. villosissimum. helleri. ramisetum. helleri. filipes. laxum. dichotomiflorum. parvifolium. laxum. pilosum. ghiesbreghtii. laxum. cayennense. fusiforme. exiguiflorum. acuminatum. parvifolium, tenerum. diffusum. utowanaeum. fasciculatum. millegrana. laxum. adspersum. reptans. adspersum. diffusum. laxum. ghiesbreghtii. sphaerocarpon. lanuginosum. patentifolium. malacon. Tracy, S. M.—Continued. 9141. commonsianum. 9142. joorii. 9143. lancearium. 9144. arenicoloides. 9338. hallii. 9342. dichotomiflorum. TuERCKHEmM, H. von. 56. multirameum. sphaerocarpon. 428. olivaceum. 657. millegrana. 1254. laxum. II. 1956. biglandulare. 7699. zizanioides. 7700. zizanioides. 7702. pulchellum. 7797. polygonatum. 7798. trichanthum. 7799. barbinode. 7801. trichoides. 8617. barbinode. 8783. millegrana. 8784. millegrana. 8785. zizanioides. 8790. paludivagum. 8794. pulchellum. 8795. polygonatum. 8796. zizanioides. 8797. pilosum. 8803. laxum. Umsaca, L. M. 1080. pseudopubescens 1087. meridionale. 1657. depauperatum. 1669. leibergii. 1670. perlongum. 1685. pseudopubescens. 1704. scribnerianum. 1789. dichotomum. 1791. latifolium. 1799. spretum. 1800. meridionale. 1816. huachucae. 1820. huachucae silvicola. 1922. virgatum. 2153. sphaerocarpon. 2155. meridionale. 2237. huachucae. 2242. lindheimeri. 2244. huachucae. 2353. Jlindheimeri. 2363. miliaceum. or 336 UmpBacu, L. M.—Continued. 2365. scribnerianum. 2543. flexile. 2646. villosissimum. 3686. scoparioides. 4962. lucidum. WeTHERBY, A. G. 18. lindheimert. 19. dichotomiflorum. 21. gattingeri. 32. clandestinum. 51. lindheimeri. 53. latifolium. 58. ashei. 59. commutatum. 64. villosissimum. Witcox, T. E. 13. virgatum. 15. virgatum. 22. latifolium. 27. dichotomiflorum. 41. latifolium. 54. virgatum. 100. virgatum. Wiis, T. A. 1. sphaerocarpon. 2. sphaerocarpon. 3. ashe. 4. ashei. 5. villosissimum. 6. dichotomum. 7. clandestinum. 8. dichotomum. 9. dichotomum. 10. ashei. philadelphicum. 12. philadelphicum. 2184. capillare. 2228. leibergii. 2577. scribnerianum. 2847. barbipulvinatum. 3009. scribnerianum. 3061. virgatum. 3089. amarum. 3090. amarulum. 3097. auburne. 3098. sphaerocarpon. 3099. aciculare. 3100. augustifolium. 3105. auburne. 3110. oligosanthes. CONTRIBUTIONS FROM THE NATIONAL | Witson, P. 8. 14. Bitte ot. 47. 125. a kaye 179. 188. 283. oll. 512. 593. 1248. 1249. 1405. Wooton, E. O. 64. 303. 368. 1068. 1071. 2001. 2014. 2017. 2936. 2948. WRriGuHr, C. 280. 753. 757. 758. 759. 761. 786. 792. SNES 1317. 2084. 2085. 2086. 2088. 2092. 3450. 3451. 5452. 3453. HERBARIUM. polyanthes. xalapense. boscii molle. ravenelii. ravenelii. microcarpon. microcarpon. dichotomum. trichoides. trichoides. diffusum. adspersum. fasciculatum. dichotomiflorum. capillare. diffusum. obtusum. barbipulvinatum. bulbosum. obtusum. barbipulvinatum. virgatum. pampinosum. plenum. virgatum. barbipulvinatum., ravenelii. trichanthum. glutinosum. ghiesbreghtii. laxum. geminatum. plenum. reverchoni. fasciculatum chartag1- nense. dichotomiflorum. pampinosum. lindheimeri. bulbosum sciaphilum. lindheimeri. obtusum. exiguiflorum. pilosum. utowanaeum. chrysopsidifolium. fusiforme. neuranthum.~ HITCHCOCK AND CHASE—NORTH AMERICAN PANICUM. 3887 Wriecat, C.—Continued. | Wricut, C.—Continued. 3454. fusiforme in part. 3860. dichotomiflorum. 3455. millegrana. diffusum. 3456. dichotomiflorum. 3861. dichotomiflorum. 3458. parvifolium. 3862. condensum in part. 3460. lJancearium. 3863. laxum in part. leucothrix. 3865. cayennense. 3462. erectifolium. 3869. adspersum. millegrana. 3870. tenerum in part. 3463. albomarginatum. 3871. stenodes. 3463. caerulescens. 3872. megiston. wrightianum. 3873. virgatum cubense. 3466. zizanioides. 3874. acuminatum. 3467. scoparium. 3875. polycaulon. 3751. laxum. strigosum. 3852. diffusum. 3876. pauciciliatum. 3855. millegrana. 3877. diffusum. 3857. reptans. iT ’ ii GT Nee he Bae Ta. HAE RAR Nata } j aie r 1° Seihiooe ae a Ne emt Ter ee PUR eG) aS bh uxt et. Lane § wy pt Pan Weds Paral Mee bi) ey PA 8 Di ha) ett HORNER RA sy "5 LJ i ats eo \ SRC RMT Oe yt ORES spin) We Dipl teh Sutra t ae 5 Sabb bahay Se Aver any Noel A ; Woe * SHH ait rh: Ni . 4 es, / ’ Fee bait r * f ares. iJ i, t ; j ft 4 ist) ' i : f 4 ‘7 q fp i 20% ABS i * i 2 ‘ 7 / i ) F ' ms / ‘ ie ta \/ were, 4s 3 Ree ar tT INDEX. [ Page number of principal entries in bold-face type. Synonyms in italics.) Page. 0 FE ee See 100 SE ee eres 115 co ea ye ES ATS SRS Sd SS 2) Se a 107 DN SSE tae Se aes ee ee 100 eee eer reer 102 SEMIN noe on == sc ec bce sees othe 99 a Se a ee 118 American Codeof Botanical Nomenclature. 5,14, 15 a ie a 165 eae wy. Nerpariim Of... -.............- 2 La Se Se er 18 Barbey, William, herbarium of.............. 3 Beauvois, limitation of genus Panicum by... 14 EIR ee eee ccc ce suecevn 2 Penman MerDAaritm..-...........5...'s.2.- 2 NE oe oes ww on wn on ov ne atcectances 176 oy eS i ae 1 0 ae 3 SS) ear 4 CO NS 3 RE os own oboe ela ne ce vwns « 18 | eens aaa 323 a a ee 321 ee 36 British Museum, herbaria at................ 3 Dc 2 a Se 69 a: 3 Buckley, collections of......................- 1 a Se ee 54 SRMEMMIMOBCUVION § © 62.5 22.205 s05cc0ss0sneess- 13 Oo ee 3 Cenchrus alopecuroides....................--- 12 ear 12,15 os ee pe 11, 12,13, 15 a orp nkm tye c'sdé anew cease 11 4 11,12, 13 EEE os nc cas se aenesncnsenes 1 Citation of specimens. ......................- 8 ER oon 6 noc os ve nnceo stews 16 Se ee 47 0 240 Columbia University Herbarium............ 1 ee 300 Copenhagen, herbarium at.................. 3 eo ae ss nse sv abmaae asec 4 De Candolle Herbarium..................... 3 Oe 3 Ns Pte Wa oa one wd one secu p 151 Desfontaines Herbarium.................... 3 Desvaux, duplicate types of................. 3 eas an = tum awe nokae a ans ote 4 ee ae ee ee 17, 142 eR eM as ys wacaananskecde ee keoe 179 es cry oo s'ns dexrtcenusendas caine 47 TE Pea Ne sie ae kd nn ennnadnnoceDars > 71 / Page. Digitaria affinis...--...........22 5 pees sae te 30 ph et af EE gee Ne hn a Ee Sar 30 Sanguenades 2. 26s ls ies Pelee Ne 15 Distribution, maps illustrating. ............. 10 geographical, of genus Panicum in North Pues A! RE OR eee 8 Diaelll types Ol 22. ose sete eae cee nee 3 Mogiviher species: = 5 sane Sek nee eee 329 Drake de Castillo, herbarium of.............. 4 Eatonia purpurascens..........1-.2.-2------- 87 Behmochinn Colona... 22 senso eet eee es 31 RCNP Ses sais. once sete es eee 11, 12,13 HMGHINGIBDIA: Sook sone oh Cee een eee 16 MOMS Perpariunn. oo. 225-2 che eee eee 2 ‘Engelmann Herbarium...................... 2 BMD yore ac ls ra a Se oe aes epee 258 EGriOuHane INCHENG =. 5225. seven coe aan 123 PEN UNA ee oc 8 Dn oo sae on Ree ee ote 35 MEStUCL OPMINS. <2 ..2c 0 oh aes can once ee es gee 118 Field Museum Herbarium................... 1 MGreNCe NET DALI At. 2-222 2ns sca e eee 3 MOURHICK. GY Pes Olooca es sancawce mecer stant oe 3,4 Franqueville Herbarium...................- 4 Fries, South American collections of......... 4 Galeotti, Mexican collections of.............. 3 yastridium lendigerum....-- pA tiptoe ro 5a ie li Gatinper Herpurigins::** 2.5.0 2.22255. .5-2- 2 DE CSLTELCE | hs app he a a oe ee Re eae 30 Genera excluded from Panicum............- 16 Gerlova, herbaria at. 2. 2202s s eons sae es <= 3 Geographical distribution of the genus Pan- jeumiin: Worth America: 2... 0... ..- 0586s 8 Gottingen, Herbaria at... ..--...5.-...5..s=.- 3 Maray EGR RAUIEIN: 3 Seale oa etn ee ans ot ome 1 GrIsebacCh MernsMum soos costae nant anes 3 PLONOVINS ELVErDSrTM: of wenn ene oe 3 GLOMPINE OL APPCIS! sen we oe ay ewan eases = 17 RUINER OTM... acne ati nas ncmmsewaccas peas 79 Hackel, Eduard, herbarium of............... 2 Hackelochloa granularis.........-... -.--.- 312 PIAeIKE. COMECHIONS Ole 2.5. + 22can st aces caance 4 IB? SBE DARA oe cae ac ae win oo scam bess 3 Herbaria, American, examined.............. 1 European, examined..........:....-.ce-- 2 EOE annth, GRO ro occ ek oan a neem eee 2 SEG Gr nia ts ate etnias cals a oe ee mea ae 3 Berlin, K6niglicher Botanischer Garten. 2 SRRIERNN cet, on eb a scan. cts Oa So etna 2 BUNES sees apse san sh eee 1 DRY ater Daas noe xe vies Sa en ae Se 3 OUD 5 nana n> ome pueee ce ee ene se 4 Herbarium, British Museum of Natural His- Eg Ce RTS PRE peered 3 3 Brussels, Jardin Botanique.............- 3 Columbia University... o... 2 cnc asec sas 1 890 INDEX. Page. Page. Herbarium, Copenhagen, Botanical Garden Herbarium, Stockholm, Naturhistoriska Rik- of the University ..-...-.--.----------- 3 SINUSCUM = 25. 2 ks 6.2 noone 4 GGSS OT erie oe eee res eine anaes 4 Swartie oo k..s.k.n..2 occ ss 4 DeWandolle?.22 2-255 -es- e-em en == 2-1 3 MOLLY cee. 5-\p52e-2 Ho 54-42 = Se 1 Melesserieccccn. tec sens eee semi eee 3 TriNiUS, <.0324¢240200220-0=. cone ae 4 PesfontaineSsscs ee eae eee ee eee 3 United States National... ...22.-7-e-seses 2 DOS VAS 2 on. ck aoc Nee She eee eee eewee 4 Van Huerek:.). 22. .-2. 5 <--2eeee eee ‘2 Drake de Castillo. ..-.322-----2s.+2----< 4 Vienna, Kaiserliches und K6nigliches Oboes cece eee se ne ei tencrsiors mateia1~ 2 Naturhistorisches Hofmuseum........ 4 Eine eli arin semen eee sa see seer tee = te 2 Walter's... . 2.0:5--2420002see eee 3 Mield Musee: sco. .o2s->eackeeee mes 1 Willdenow ........:----+- 2-5) eeeeeeneeen 2 Florence, Orto Botanico.......---------- 3 | History and limitation ofthe genus Panicum. 11 OUI eR Sees cee ec cae es Paci Scenes 4 of Panicum after 1753..2..-—_ ia aseeeneee 15 Mranquevilles sere se= =e ete =a == 4 | Hitchcock, A. 8., herbarium of...........-.- 2 (Goaitincer epee te eer sear renters ee 2 | Hog millet... .....0-.2.. 00 2-50 69 Geneva, Conservatoire et Jardin Bo- Holeus'sorghum...:........-..0sesee= eee 11 fanless se pene oasis Seo ee 3 SPICAIUS.. 50. 220s 5 5. 15 Institut de Botanique del’ Université. 3 | Hymenachnie: J. 222... 00-2 oss oe 16 Géttingen, Botanischer Garten der Uni- Tchnanthus\glaber.. ...:.-+-.s52 see 88 Versititce cscs cen eae emanate ce 3. | Jussiew Merbaritum(.....+: 222 4.eeeee eee 4 (OOS aa Sonate eb acenarr a amciooac ce 1 | Karwinsky, Mexican collections of........-. 4 Gaise pac hiee os sees ete a eee Cees 3 | Kew, herbarium at... 2.22: s0s-eeee ee 3 Gronovius. ...-.-------+-+++++2+20rr eet 3 | Lagasca, duplicate types of..........--.--.-- 3 1 ee at aes aaa 2 types of........22:3.....2- ar 3 ANC ma sce ci ei ean a 3 | Lamarck, duplicate types of..........-....-. 3 Hitchcock’s....----------+++++++--+--++-- 2 Herbarium.....:....2c-2.225.4 eee 4 Jussieu .....--------+--+220022 222s ere 4) ancearia,......:.-:.<.:-- ee 271 Kew. -. 20-9220 en ene een eee ecee ne 3 | Tanupinosa../.0...-.1.1..-2esee 208 Tamarck.....------+--+-------++-200007- 4.|\ Vasiacis...2...2--.oc-.2) 2-4 16 Linnean. ...- BRIS aso FO e tia aed COCR AG 3 divaricatas.......... 2s. .o-de eee 16 London, British Museum of Natural Latifolia... s....22.c2.. ee 312 History....-. .--- we eetee eee es ec ees 3) Taxa........c.:)..cct2 7:00 110 Kew....---.----- Berga ee ae race 3) Daxiflora, 2... .0...<22.082 2 158 Madrid, Jardin Botanico......----------- 3 | Liebmann, Mexican collections of.....2...-- 3 Michaux Scien Reese oan oe is Lindman, South American collections of... .. 4 Missouri Botanical Garden.......-...--- 2 | Linnwan Herbarium ........2-02...eeee00+ 3 AOC): defep ranma ren 2 London, herbaria at... =. .1222 2 -eeesee eee 3 Muhlenberg..-..------------------+------ 2 || Avacinidh hespadl t 3 Munich, Kénigliches Botanisches Mu- rae Sa ania ae CS ITN os. 1 bei oats 5 lake ee 3 Maps, wee distribution: 22se-e-eeeee 10 , | Martius, collections \0f..22272.30e eee eee 3 INES) aCe eee go ado es Seta se See neeennG 1 Mating a New York Botanical Garden......--.--- u tice Carl, ae k of RES FS ee 3 Padua, Orto Botanico......-.----------- 3 Fes a Co 6 ea Paris shunniide eee ean, 4 Michaux Herbarium . vtec ese eseeeeeeseese ess 4 3 Milium, Linnzeus’s limitation of the genus - . 13 WOSSORe eReee eee teen cise = = 4 ; Tea er et eae 4 pre-Linngan use of the name ..........- ll DrkedeCasilon eee aes ss 4 Tournefort’s limitation of the genus.....- 11 Fe Cra, hats ten oe fale a a 4 Capillare 0.222.255.0222. 60 Franqueville. ...- nak 0 ae a 4 clandestinuit....25. 152222 - eee ee 312 Tenet ee ee 4 confertum::::..:<25.2..-.2 aoe 13 Teainaraics ee eee eee: 4 effustim... < e2i228.c4-<. Bee 13, 14,15 NUCH eee ee eC cece 4 escUlenvwm . «oes do oo c inn > eae 69 RiGhard ee eee ne eer 4 latifolium: 2.22.0... ose 2s ae ee 314 Rrendelecte ee eee ee ese 4 microspermum.... <<. J222. see 131 Redeye eo eee eran cee rem es 2 DANICUM..\< 2600s a 22 eae ee 15, 69 Philadelphia Academy.......--------- , 1.) Millet.j. 2 55.2.2: 22252. = 2 eee ll Prague, Botanischer Garten.......-.---- 4 ProOM-COM .< .:-.. -< <5. vee eee ee 69 Museum des KGnigreichs B6hmen... 4 fox-tail. 22. .::2-.----.:--5-=2-= ae 13 [ENR eed gleediten ig Mien Nita yl en eh ot De 4 hogs sccweees sa. oe oon 69 CHAT eae eee nee cceanees 4 peath. 225 o iis ase 15 St. Petersburg, Académie Impériale des Missouri Botanical Garden Herbarium... . .- 2 GIBNGES Fe ete ca oc one cee Seer 4 | Mohr Herbarium). < (2 = cena te eeeeee 2 Botanical Garden ............------- 4 | Mueller, F., Mexican collections of..........-. 4 Sori Pmels case fae sia telere coe see ne Shien 2 | Muhlenberg Herbarium.........-.-.---.---- 2 GHORG oat eon ca oie eect aeeten cmeien aan 9 | Munich, herbarium at...............--.sseee 3 <4 fay: 10 (We epee Se aT Bes oR Ot 3 | Nash, G. V., herbarium of..........-----.<.: 1 rend eletne saerncmearc as sacce shesteinsw eRe 4 | Nees von Esenbeck, types of............-.-- 3 ® INDEX. 391 Page. Page. New York Botanical Garden Herbarium... . 1 | Panicum balbisianum ...........-..------..- 325 Nomenclature, American code of botanical. 5,14,15 UL) ST las eae ee ee ial Se ee SO ee 268, 269 LLU D As Se SC eee ee eee 179 PALDINOdE Wow. = os tacos we 17, 18,83, 34,35 Hemel, collections.of. .............-...--.--- 1 barbipulvinatum ......-.......2.2...... 61,62 LS SN ee eee ee nee 278 barbulatum ...... 6,7, 181, 183, 198, 195, 207,331 Oplismenus compositus...........-..------- 12 ARO WERSE <5 oo 2o oon ace 52, 53 IRIN on onc nn ea Seow 16 DECRTONTEREONME << 3-5 tee 30 CS 3 TEP et che wrk wee te ae te eR 7 PCEEIRCIONGOUL.... ....... +. --.~2+-->0--- 118 OUMICNEN. 9 Sot oe 2 ese eae 136 D3 ee ee 118 JGESIPN EDN Be oe eee 177,178 Panicum aciculare .... 166, 169, 173, 174, 175, 176, 212 Divlandulares ct a.<2 2. sean 120,123, 124 0. eee eee 297 DONE Seek S56 oo ee on oe eee oa 60 SMEAEEITYS. =. Sok 151, 152, 222 , 226 DOQTEIETIUNTU SS 63 oo a Ie tte 185 See ae ee or 129 ORAS Aen tare te en ee ee 189, 239 a ee 240,243 Ose E sae) Ne a 193,314,317, 318 VE eee eee 134 LL) [eee eae 318,319, 320 0 fi SA ees 18,43, 44 DOU GUCLS x Fs ae! Ds 112 0 SS eae eee 30 OPEL 8 oS ora aioe nh ws a a 48,49 Is oe on ss ie we 100,115 prachyanthumi-o...2. 3-0-2 fo ecese 128 eS eae 115 DTQERYCIAGOS: 335)". 52 2 eee ae 122 a@grostoides .............-.- 100, 101, 102, 103, 104 DEQSIUETIRG 2 ooo <=) 32 See ee 134 TE SR, Oe 104 BIPVE sco So... 3 ana cages eee 271 0 nee: 244 preysioliumn:<- 222225 ee eee 13, 62, 129, 329 EES oo ce on eereey 2b DIONE oe ot 5 essa) See ee 264, 265 0S 311 DT POETONING 507-2 8a oe ee 30 albomarginatum...... 259, 260, 261, 262, 263, 264 RIT R = 292". 3 52 eS Po ee 31 0 ID en arena 12 BERN POT «5 foc)025'5 22 anon ea RRO 329 RNIN a a a a oo win ws 82,141 2000] 2 7 6 a eee 58,81, 82,83 EI eset oe a Sd acme 94,95 TORTICEUM 5 « os8n 2 oo ne 82 a toh ee eee oe 96 TTS 5 oes soa ie See esha yee Se 83 AE ee 94,95, 96 Sup ss 22 2 <2 See 83 ee ee eee 95 8, Var. PiOlaceint. 522 320s se as 82 SEER ote So oo o.2a = onion abet wae 329 UUs. Jt se a 2 eS ee ee 177 Ee eee 12,14,15 GEPTEIERORTIRS (07) oc. (see ee eee ee 197 CO ee eee 49 CULE DURORUMN on cos omic o seman vo Re 72 ue: 2 100, 105, 107, 109, 110, 331 caespitosum........---- 8. coat cuban stints 36,43 a ea ere 107 PIL LOD TRUM ara = og ciate cleo eee DER 169 0 Se eee 107,108 Caliphyllrm: ho os <6 5 ce eeee cee 178 eee aT IE SS I ae ee 105, 109 CANINOR UNO 22 = x 55.03. c ect e a sae eee 57 aR aS ES ee ee 97 POEL Sos os 35. oa eh 129 STS oe ep ie a SR Set as capillaceum strictius........:..---.s0s--6 129 128, 170, 171, 172,173, 174, 190, 201 eapillare. .\... 2 -<. 13, 55, 56, 57,59, 60, 63, 70,129 * I ete osm oo eh cam i cioman 185 MONE a ni ne a ae osha 60 es on oscar asa cen wane 185 TI SOOWUNY So St ae a aoe 62 0 ae ee 113 DUNIRERI Ee onic ates ae 57 appressum..... ee ee ey a a € 31 WIEDER ae ax Ne Sl, Secon Skok Ae eee 55 NN te Bn oo oo a ain or en cmee 48, 303 QOMMIQEN Sooo ss eo hek o aa eae 57 RIE 5 5. se an one cence nnn 13 VETOUIOU 2 Saco ee 57 ee 85,88 TEE Dt OD Te = 64 ES es 166 LLL Tog) 1 ae Si Re: ee Bags f SRS? Ooh 67 + 173 TIM ATINUDR aoe ooo Sa oo neem ae 58 SINS... 2s... 3s 18, 42,44, 45 TANNED 58 St IRS ee eee 55, 283, 301 MIs sone cnc Son tac 45 BUEPUUEEIN So Fa = in 3 iran a ee 58 a ee 45 TRONS 62d a2 onto d Se arda ote nae 60 eS 45 GSPIUEIDINes oS. to eos eters 72 0 a 329 | CECILIA Be os ohn hig ars Dar ea ras 159, 160 oS a 134 PALIQOINGR oF. oo seo ines - SSS Ue eee os 99 oo 2 ee 154, 193, 300, 301, 302 COTTE x Shs id dn Os rt 2 323 SE teeta ese on oi oo ascas amas coc 69 COS ONUNI so Bee od nc 2 See See 30 laa ee aE 233 COTOUTHTAUT os oc oc oo anc sb anctwaenubews 323 EMSIRS i Miss wo. c ony o0 aden nes suse on 16 COPTUAGMADOE i. cn os ssa tees = aE 329 SSMU a arian canoe peacdeharca cde 325 CAVEMNBUNG: Le 55 1.2 au san aeb eka ue 70, 140, 331 MERE es ooo er an aidacdannneiron 223 GOON 5 «oo nisin’ pac soe oaks 140 OU! a a ane ia: 36 12.7 1 || ree ean Cee oe 70 IN a Sa oil oo0 o00ce sense ccunh ace « 158 Chamaeionche «co sus. scan soewee .- 268, 269, 270 GuUstrO-MOMANUM.......- 22.222 eecceuae-s 267 ORS MAN 5c ios kes oe lateds onde eee 24 ENN Pts so px ones valexenaanaer xe 81 CROTTOYINONCE sinc n waren «ae 40,41, 44 392 INDEX. Page. Page. Panicum chloroticum.......-0-+-++++0+-+ 48,50,51 | Panicum dichotomum barbulatum.........-.. 193 NEM abe seat ss Seeas yceckee toe ee eee 48,49 COMTVUNTE . -- 2c = ovo See 191 DUG C inom wales oareic Se eriAe ee See 49 CUTIOIUM...< . WS oc oncenpsn cere eee 330 SUINCSIM Ota sin Se tcsc ccs ewe coemeene 48, 49, 51 Givaricatum. 262.72 ose pee eee 191 chrysopsidifolium............. 168, 169,171,175 Claw... 2.53 6s s2s2 Soceee eee 298 CL OMUIBS oe ok ot Sa Sd 22 Seatac pee cere 162 fasciculatum ....2.25..... += ae eeeeee 216 Ciliatissimumss: se d57eiee eee ee seen 324 OTOCIE 3.3 de eee 330 Ciliahiumis . 22. Sees eee eee e 162, 163, 164, 216 laniiginosum:« 222-5. Loo. nceeeeneeeee 220 CUTE apace oes hat canes aceeeeeS 237 NMI Sense eo ee ee antes kee CHOSE Aas ae) sec eee ae Sheets 220 spathaceum...; > .-+<. 2s -oees tea 330 Cliniginirmts dates Prce ee akc.cge cece es 325 Sphacrocarpon..... -..2260ae- eee 251 Clandestinum........-....' 18, 290, 298, 812, 314 VULOSUM = 255 320 2 ae 233 PECUNCULELUM = x\cieiaisctn = cine oe saan 312 VITIE 2s os ct eee eet oe 190, 191 CMUILCI Ee eee eee ee Ue LL Ck sch bade 188 diffusumies=s see seene 6, 55, 72,73, 74, 77, 380, 331 (2 YLaY LAE a Se a ae eas es 34 digitarioides : 3... bv tecec 2s oe eee eee 323 COLOTARUIN eM ae t ck eke te Keen tee Aci cot ters: GiNdIOtUIM 52... os.< + ime 382s 12, 323 columbianum............. 240, 246,247, 248, 259 disciferum. = .).\.<'.2 3. 2 see = bon sic bebS C000 heeyeg peep Somer Se eet 5 ean 240,248 GiSCOLON 2 noi 05 snd ese 181, 187, 330 combsii....... Joey Pts Sas See 106 USPeTSUM S22 .0...5505 Leet 136 CONVINEMNNAEOUUWIMN= «= 5-252 SAaa eee oe 304 CissectuM 22252222 3 2.2 see eee 44,45 commiutatum: ==. .5..24 5442: 6, 154, 237, 301, 302, GONG shoo. i583 Cote cee 113 303 ,304, 305, 307, 310, 317, 318, 329, 331 distantifiorum:.. .29. 22-2 ieee 22,23, 24 CONSONHUINEUN wis secs s acca. cess esos = 169 distichum 232.2. 052360122 sooeee eee 113 LOO INS Soe eee oS oee eee eae oe eee 304 lanctifoliwm.. . 32s.6,.45.2. 22 eee eee 114 MATS S Aedes eco ewe ee eee 301, 304 G20GTICOWWIMN.... cS. aces Shae Chane 16,118 COMOPRYMUN S22 psc he ease tte lees ae 222 CUM MONG... 52... ee 327 COMPOST Ms eee Re ee 12 dubium. 5 223.0205. eee 112 CONEHSMIM 55.5 Ans0 5s seo eee ee ceneee 124,329 QWMUS 1.2 oS ao se se ee 195 CONGEIMMAUS'. Sites see ae eee cata e 263 COMED x San Osos occ Sones coe 267 Condensum 22-8 ses eas cee seen ek 101, 102 COL0NI. one. S22 Sees. oe eee 201 confertwm: §223252 sates oly mote kwrain dail eee 240,241 purpurascens...................-.... 120 WISICOPRYMONG - 9 «5 v0.00 +c Senn ean tne 280, 283 hirsutum Swartz....................... 77 TRULETIEG oan kn 3a nt meade eae waive aie we ves s'Jie'a'enaaaner= a 48, 118 huachucae....... 7,214,215, 217, 230, 233, 238, 330 MICMICIGUAE. scot ne cet t 210, 211, 212, 214, 228 silvicola ........ 7,216, 219, 220, 221 , 230, 232 microcarpon Muhl. in Elliott............ 6, aia a 112 : 181, 182, 184, 190, 193, 195, 255, 330 OS ee 49 microcarpon Muhl..........-..--.+-+-+-- 255 NES oooh 08s a nvceteceancv le 88 isOPhylWM......-- +. ee eeereneneeeeees 255 oS Se eee ST 38, 166 SDR VOEDOR, 5 xcs annianpehecksmaien 251 implicatum .............. 211,218, 214, 228, 246 TCTODNY ME uo 305 shannon adn deganis 210 SS, a reenter 253, 254 ICT ORD EI INN « xan ds bo nn a bebe ee 131 le wn, hia wn 36 “ miliaceum.......... 11, 12, 13, 14, 15, 17, 48, 67,69 OS eer oP ee en een 151 rT Sf ee Ee wep meee fo sta 50 394 INDEX. i eee ' Page. PPAMICUIN TILT nae c cicla 2.210 on aise Kqwleiiae eee 69 | Panicum oryzoides............ oesece hasan 325 miuillemranmaes fos cis actis\cje's cic. eis.c min 135, 136, 137 (1): (ee eR 235, 237, 238 LITT SEO CE SOOO R SOOO Ogee 58 OVAL OLWMN. Sas - « o~000 + =2 050500 13,129 SUMMA R eiade decinais = sss saeesenaeasteesteee 55 OVEBUMN 5 of eibc00.~0 5025 eee 174, 175, 176 TNGULEONUAT s w's,= os 023-0 =0os iano see ers 117 OWEME » 6 o.o00 55a 5:2 «0.00, 3.~ 0 2 243, minutulum..... SoontEaetbosc sab Mache 207 paeificum....... 22.0.5 229, 231 MSSta st PICNSC aicla sexiness a sea 253 paludivagum “*. 2250-5 17,18, 31,82 NOMS ss atarsiais acs aerciasaiate aielels's cles 34, 35,42, 45, 320 paludosum......<-..-=c..cecese ea 53 MOTMICINOUIES 9 <-> win = an '=le\n , Si 292, 298 polyanthes......--.-.----.--.--.. 181,254, 255 MOUAPLONWN ass ss creniaes aaa cloister a eat Se 183, 207 polycaulon.....------+++++++++++-++-- 162, 163 MODOSW Ms rata icfaseratareie hella S Shek Cee 82 polygdMUM...----- +++ +222 ee eeeeeeeees +e 78 MLOLULIIIV ES oe ne ae eee eats 85 gongylodes........-.~---++--+++------ 82 UCHCATIICS Re en Seen een ee. 179 hirticaule....-.--------+----++-----+- 64 vieveant la Glvatibiiipete mbeey tae 5) hehe hepa shania. 34 polygonatum:...--...2--poeaeeeee 110,112, 118 PTTL ACV eRe ep nsf SE 107 polygonoides .....--~---+-++22022-22222-2- 321 DAXUCCTISC Loe eae hee ee nee cane antiees 141 POLYREUTON a.» += -2 = >» =< ne eae See 5, 6,304 OULON GIRO oc cic ieee a oe asia teeret ci mie sis 118 porphyriwm.....-~-++-+---+2e2ee rene eee 58 OHIO ROU ana este: eee eee et ons 125 porterianum ....------+-+++-+++-2+-222+- 317 BHPUSIOTAUNLES esas teen Pace ae ae 138 portoricense......--------------..---505- 331 HUSUNI. cere l eae ee ee ese $21, 322 POLAMICWIM ..- = ae vse oe se cee 112 Opcidentale sie. 6. owes we ete e 228, 230, 239 POLGMIWIMN . - <= -0 cos. sone anne S eee eee 112 DELO NOON Ak cide Soae, «tee oe Ieee elas 201 PYaecocius... .....-<<<= <5. -=5= seen 226, 330 GUEANUNUIMS s- eeeice cts oe See oeetiee ac « 285 | praticold... ..=....2.s2.5+.s==pee eee 79 oligosanthes.......-....00.. 6, 280, 282,285, 286 pre-Linnzan use of the name ..........- il GUVACCUM rece en aiciciatealnneue a «aeee 185, 225 DVENE «o-oo ass esos ewes stn ne 132 OLYTOef OUT ie feiaienlen a ease eee se eee 121 PLOVANAWM . <2... -< +--+ n= ces see eee 136 DNGLODENEC. «ic wren are econ s sens see eee 276 procerrimum.. ©... .....-.:.--5225seeeaee 141 OPLISMENOITER a= ance e aise niece n= te eee 135 WT OUSERWTY, =). iio cin xlaicte Samet 49,50, 53, 54, 100 ORARTENSES ch ierche ta tade stake ees sae ae 220, 221 chioroticum ... ...-.-><-~.-=-+-=eeeeeeee 50 OFICOlG sss sskhc we etckcwemeewesemecmene 240, 249 Geniculatwm. «0. ..-.e sete eee 49 OMATUM Leica cascode dasasceeeaesaameenars 331 DULOSUMN. <.0n.5 5 co sciens asm nee see 49 OF RODAYR UIs ciicemcisusss cs saree enact 173 SHICRUWIN < Ciocock a Shen's ose eee Dee 49 INDEX, 395 Page. PPRMICHI POSALUM . 2. 20. 0ncesncansnecen- 36, 232 LOL OT Se ee eee aS 36 er eee ee 87 ee ne ener 247 0 OS Se ee ees ee 105 pseudopubescens ................-.-.- 235, 236 DINERO io ao one orice Ce cn's 325 a a eee eee 136 pubescens... ...-.... 208, 228, 233, 235, 294, 295, 330 Ro oo cc wae cee sone 193 0 SE eee 320 EMRE eS aca c o nicenicine cece 128, 124 sé ee 159, 331 eee 31,33 TS eee 166, 167 esanoens tt, 5, K 2... 22... 120 Marascens RAAGGI... 2... 22. enscnnne- 33,34 Se 158 0 331 i ee 5, 25, 27, 28 IRR ee cn nan ane ne 41,44, 172 0 115 a 170, 269 0 a ea 191 a ee ae 159 _ 2S 278, 280, 284, 287 oo ee 151 RS eel noe Sa cow cade awnmen 174 SEMIS Noo cons oS ama ac eee 331 Ele co ow occ o cen aceneee 322 A ee 85, 87 0 pean 86 eee 321 J 17,36, 43 0 38, 40, 41 ee eens 49 LS 25,26 ee 139, 140 0 105, 109,110 iy an ae 140 ee 100 0 ae 196, 197 0 107,331 i 133, 139 oo 2 ee 70, 71, 133, 139, 140 SS 139 Ie Soon ss cones accvan's 127 cas ni aac ecacteccs'e's 135, 136 RN 5 05S oo nc soa wnwesec. oo) Somes 203, 217,218, 219 Ui (cee 211, 215, 247,259, 260, 264, 265, 304 TOTETSELITIIS Sone 2 ote nd cee 24, 115 MAUS eae i 3 no re ns os oe oes ae 18,46 RIETIRIE ree oe Ren eh Shee 8, 230,231 LTNORUDU UR a oo 2 ae cls ove 4 tous 2s cee ne 43 TEs)” i ee ety periery sae weve aay 224 POMC 3 Oaks ani wcwinind ntslectan tae sare a 58 Tournefort’s limitation of the genus...... 13 PFEMVENUNT oo oo as we <2 pc tenable sere bd Ne 190 pig tle l Ne AT TW ITs to paqth ke al ene ae SS 131 IIGMMOINGNNG oes sce n a. ccc ane Sas ee a dee 124 trichocondylum.............- ana to ae 79 OMNI Sos, ina iota pa Seat > 8 cdemeane ne 112 PRION ene ots ee ne 60, 129,131 PACROPROTUIN 5 res ooo sn we pao ona Ogi none = 113 THILO oda wie a warn bu conn laa rca ee adeeran 117 PEUONUMMs wash ees.o cet nance eae tesn yas 261, 263 SIEGES TA eras reas ns baa are wr evento 30 BRUREUORINN gn ou cos cone deere 240, 245, 246, 248 PROLTOUMUMUNG, 5 ncn tae en cg tde bas eenwer a 141 PRIRDOM HORII a oi en o's 4 o ama oan ov ane anes © 16 bY DO BPOCCIES Olen 252 06t oe can ture naa ee 13, 14 WINOTUCUIIE sa 0 io oi vacom sees euaecemepr 127 URDTORW ikea na + hud sae ek eee 122, 301 396 INDEX. Page. PanicimWNCInatiil wesc. conc smenpncmeasme 16 | Pre-Linnzan use of the name Milium....... UTCUD WII et ona = cee ceeceten 215, 247,259 | Pre-Linnzean use of the name Panicum..... TONIUAPUOSUIY «coc easesece ss === 01/5 227 | Presi, C. B. and J. 8., types of..-...-.2.-- yey LOLI DTOSUALUM any asne cence 232' |) Ptychophylltm:s2. 215222532422. INP wer ssasccescnsneese cere =: 213.) Pursh, types Of. 5...2 2222.22.02 ee GHETTO a tastina oan cen wean ee ac 211 | Raddi, specimens from. ..............-2---s+ CRINVUNE. os Se ese oe paneee eee aes nie 248: || Rafinesque, types of... ./:5.222 322252 eeeeeee urvilleanum..... Re Eee. ae eee oceee 19,182 | Richard; typesiof,.22::: 25). 532 SSS LONGIQUEM Cz rem ane coe wecee seem 132. | Rudge, specimens from: .-. 22 252-2seseeeeeee AIEO Wy REAC UTI ese seens aeetee se ees s ae 24 | Sacciolepis: . 22: 2222.21. 520.522 eee OULENZUEIONW Mae a2 testes sce aes -s2- 6 136 striata... 22... 4.523-2.2005s2 ee WASOVANUMI Sie sehecectas settee ste so se 47 | St. Petersburg, herbaria at................-- OEUURNOSUIB Dg canon nck nate aeacsereee 42 | Salzmann, Brazilian collections of.........-. AREY EG DASA SOS OSA Meee aS CA e eee 42,212 | Schaffner, Mexican collections of....-........ VENTA OM ae eee esas so cars ates 258,264,266 | Scoparia:e.. 222.522) 2 eee IVEETUICOSUILT yee ee ke cee mcleeisetaae's 118,127 | Scribner, herbarium of ..........-....-.....- WICOTAUMT a ee cannon ee ee aise sins sek 251 | Setariaidistichac:.. ...--2)5 <2 eeeeeeeeeeeee DUPSOVME so sna tae ane Seen tee ane aes 331 meyertes li yee WillosissimlUltleaee se noce eee eee 232, 233, 236, 330 pilost oSeso. . Sanect anole oe eee Ciitosuge Wl (22 Se casecieerace sees 169, 233 DOW GONA. - 22.52. - oe nese RVillostim ain s)32 5 hoses ooceacinascse cess 294 schradert...5.2<- 125.22 s0n- see eee Vireavuenl ==... = 8, 13,87, 88, 92, 93, 94, 100, 329 WNISOD So2 oo be wedcn bo date nee ONEDINAMOSU Maceo ae ese ee ass 92 | Short Herbariumé:- 2122.25. 2625: 5-=eeeeee COMMETIU oe Celene fe eee ceases 88,89: | Sloane Hierbaritum::. 22. - 5.22.2 5seeeeeeeeeeee (bios) gear ES Se SO ee ee 92 |, Species; grouping of- . 22 .csceeeee eee eee CAP USUMD No sess tse eae nes 88 miscellaneous:.... .-.--s.2-0-5ece=eeeeene LONG CLI TI e ate ay se ete ie ee ates ain = 88 subspecies ‘and’ forms... s2se2e-ee-eeeeee MOCTONINUM oe aioe mentees ase ci = 93°}; Specimens) citation of. 22.222. sees eee OULUSTM a, sniseeen elena essa nels ns 92 | Spelling of names:.-:.°*)..22-eeeeeese eee WATPULLOPUO << ceo oe means tees sect Seael= 125,329 | Sphaerocarpas..-.-.. 22280. Seee eee eee 5 rte 2) FO tae SM SW etal) CS ee i ne 331.| Spikelets, text ficures of. - 0222 see essen RUISCIGe llamas seen emee a cee ere 116,296, 331 | Spreta....2. 2.322% 22222 eee RCH ae oars Bee eee a aisiae cae 294: | Steinchisma...:.2.2224 52. 2 aieeeeeee ee PEIILCT Tee ae eee eae cae ae 280, 317, 322, 323 RiGNS oie sn anes oceene lee eee ELST A Bia tS et a Re Sr Re 319 | Stenotaphrum dimidiatum..............-... DELLE EN aise oles clerans eae eins stole niceia.s fan sete 317 | Steudel, types of..2.:-... 52225. aeeeenee MRED DERIAMIM. =. neal sateen 274,246, 277-| Stotkholm, herbaria.at...2 22522 aeeeeeeeeee HORTGMLS St eee sen ae me sete eek 155,156, 157,177 | Stolonifera....: 22-221... 22232 = eee syncope abbr aoe Gc Soe Remco S Sep Eee 279: | Streptostachys--.- << 52225. 5es-5 eee TMM PHOMeCOSCs eeeenece ee ace <= ce === 240,244 | Subspecies, species, and forms..............- RPA es ati ee =~) cles oom se 207 | Swartz Herbarium. ...:-..52225.46e-=eeeeeee xalapense..... 154,159, 160, 161, 162, 163, 164,233 | Swartz, duplicate types of.................-- BUPICUMMMICUIMs sasmecercecme ose acca 161 | Synonymy....°:.:5.-).-.2cepe eee ANtHOpPMYSUML. 2 = -tssssoee- -c~- = 178,290, 291 | Syntherisma filiformis -..........2..5.22.- forma amplifoliwm:.:...------------- 290 sanguinelis.. . 0. [sac ees eee EIR LOR DEN IVUGT TE eae rare aiate = = as ile e's 233 | Denera-: 2.222). 222s=522t-ceee eee VACKINCNSC ae eee neeemaa amass snr 195, 303,330: | Terminology .-...--.----25..2--s55=- eee WAZATNOIGER ance ses ae coe cn cee seat 314,325,326 | Text figures of spikelets...................-- FRANC PAGS ie eas) Seite ei econ sein aye me 8 34 | Torrey Herbariuim:-<22222°2 2 Seseeeee eee IPATIS “HELDANIS AU. =. cece eso seciawccieae se se= ms 4 | Tournefort, limitation of the genus Milium... arty, EUCR Da NtN S cio. ciate n ereleieje!niminieimcee = == = =i 2 limitation of the genus Panicum......... Heprrronles vim nyee siete a aia eapnicya ints = oreiaiecntais oat 124 | Trichoidia -.:2-s2. ..¢:.. 0.3 45-ee eee PSMA Tse cisieiwm sete sme aieale ga cre isintoniein'na's 17,18) ‘Trius ‘Herbarium: ~. << <2 <-osocseeea eee Paspalum appressum..........-<..-----.-..- 30,31 | Type specimens... 2... 2 ccc. ccseee eee CPISSEC lll hosel oc ela Agere aioe sien 12 | Typonyms... 2. 20.- 66...) sce een eee AREINGS CULEUULI cieie iste eae alniele sas ere =e = 39 | United States National Herbarium.......... PRUROCHAG LIS aac ed ace ec ieee. anes 17,18,22 | Urvilleana:...-.: 252... .2) eee FPGHICCL ate cee cten eet cienicice ca elaesis\aleaismintalere « 292 | Valota..2...--/i2:¢.25 23. eee Pennisetum americanum............-..----- 11,12 | Van Huerck, herbarium of.................-- ERORO DUTUM sop ne ckes Hons etc ee oniaste mieten 18 | Verrucoga)../s222522).222 2. ee GYUMNOCUTPUNE = -\- ware ae ence ona aeinemn 327 | Vienna; herbarium at..../2..-2<22oeee=seeeees Philadelphia Academy Herbarium. ......... 1 | Virgata.. 22... 20252-22222) ae IPDITeb uy DES Olis cen eae aac se se wen einen 3,4 | Walter’s herbarium.<-.. ...... 22.22 saseeeeee Polypogon monspeliensis...........--.------ 11 | Willdenow Herbarium... .---<_ 2.2) Speen acnic eames been aaiee Cee 4 | Wood, Alphonso, types of. <........ oc sceeee Prague, herbaria at c) 129 nD s NW We mR Ot Oh to Ore BINUING Lioi VEL [5 193g University of Toronto Library [adr DO NOT REMOVE Pitre CARD FROM THIS POCKET Oo > zy mm oO Acme Library Card Pocket LOWE-MARTIN Co. 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