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ON THE
LYSIAXASSA MAGELLANICA H. Milne Edwards,
AND
ON THE CRUSTACEA OF THE SUBORDER
AMPHIPODA
AND SUBFAMILY
LYSIANASSINA
FOUND AN THE COAST OF SWEDEN AND NORWAY.
BY
\* il I S % *B II 11,1 I Oti>[((.
PBOFESBOB "l /. .. in ii. v.
WITH 5 PLATES.
-#>•«&
UPSALA
THE ROY. ACAD. PRESS.
MDCCCLXV.
J- lie Lysianassa magdlanica is one of the most remarkable Amphipods on account of its unusual size. In this respect indeed it surpasses all other animals of the same suborder, and is sometimes cited as an example of the size to which this suborder of Crustacea can attain. These in fact seldom reach the length of 1 inch, whereas the Lysianassa magellanica is more than 3 inches long and is moreover of stout form and strongly built. We know of a species of the family Hyperidce, the Cystosoma Neptwni Guerin-Men'EVII-LE, from the Indian Ocean, the individuals of which are more than 3 inches long, but their form is much slighter; and a species of the family Oxyce- phalidcB, Rhabdosoma. armatum White, found between Aniboina & Van Die- mens land, which attains a length of 4 y, inches, but the form of the body is almost threadlike. The Lysianassa magellanica may therefore be justly considered as the largest of all yet known Amphipoda.
Another circumstance renders it still more remarkable and that is its extensive geographical distribution. It has been briefly described by H. Milne Edwards *), who says, that it was found by D'Orbigny in the belly of a fish near Cape Horn, and was by that naturalist presented to the Museum of the Jardin des Plantes at Paris. It was accordingly a matter of considerable surprise when last autumn Mr Tn. M. Fries 2). Junior Pro- fessor of Botany, brought hither from the Norwegian Finmarken and pre- sented to the Zoological Museum of this University three specimens of that rare Amphipod, with certain testimony, that he had received tliem from a shipper in Hammcrfest, who had found them in the belly of an "Haaskier-
') Annates des Sciences naturelles 3rd Series, Zoology. Tome 9. 1848 p. 398, without drawings. — C. Spence Hate (Catalogue of the specimens of Amphipodous Crustacea in the Collection of the British Museum, 1862, p. 66. Tab. X. fig. 5) has since described and figured it after a drawing communicated by M. Lucas. But neither the description nor the drawing is good, and the author cited seems not to have been aware of Milne Edwards' previous description.
2) We desire hereby iu the name of our Science to express our thankfulness to Mr. Tn. Fries both for this and for several other similar valuable presents.
4 W. LlLLJEBORG,
ding" (Scymnus borealis (Scoresby) Nilsson, on the banks by Beeren Is- land. The discovery of so highly developed an animal common to botli the Arctic and Antarctic Oceans, was something so uncommon that it caused us to doubt the acuracy of Milne Edwards' statement as regards the lo- cality, Cape Horn, and to suspect that the specimen described by that author might perhaps have been obtained in the French Scientific Expedition to Spitzbergen, under P. Gaimahd 1838 — 1840. In order fully to ascer- tain this we have in a letter to Prof. H. Milne Edwards communicated the fact that the Lysianassa magellanica has been found in the neighbourhood of Spitzbergen, and we appended a drawing of the same to compare with the specimen in the Paris Museum, and asked whether the alleged locality, Cape Horn, were perfectly reliable. Mr. Milne Edwards replied through his Son Mr. Alphonse Milne Edwards that our drawing was on comparison found to correspond exactly with the specimen referred to, ("ce me parait etre bien la meme espece et jc n'ai pu y trouver aucune difference appre- ciable") moreover that it was certainly true, that that specimen had been brought by D'OllBlGNY from the Strait of Magellan, and is entered in his spe- cial catalogue of his natural-historical collections from that region.
This animal is then widely distributed both in the Arctic and Ant- arctic Oceans, and as it has only been met with in the stomachs of fishes it seems probable that it is only to be found at a considerable depth. Its tolerably large size and rapid motion render it difficult to catch with the so-called dredge ("bottenskrapa" Swed.), and it has not, as we have been kindly informed by Professor S. Loven, been met with in the Swedish scientific expeditions to Spitzbergen.
Its being found in both the Polar seas, and not in the intermediate waters, — which however is not a unique phoenomenon — is without doubt a matter of deep scientific interest. It shows either that the same species may have several centres of origination and geographical distribution, or else that there have been periods in the developement of the earth, when certain species of animals & vegetables were, in consequence of uniform temperature and similar climatical relations, spread over the whole earth, which, on a subsequent variation of these circumstances, have retired to tracts and regions where tin; original and to them appropriate climate & temperature continued to prevail It is thus that the appearance of the La- gomu tdpina Nilsson on the fells of Lappland, on the Alps & on the Py- renees, but not in the interjacent lowlands has been explained. There has
On the Lysianassa magellanica etc. 5
been a glacial-period which has connected the faunas of these now widely separated tracts.
Between the faunas and floras of the Arctic and Antarctic regions, it is generally known that there prevails, a certain correspondence, so that one not nnfrequently meets in both with representatives of the same family & genus, but it is extremely rare to find in both representatives of the same species, and the instances hitherto recorded appear most generally the result of confounding different species. As regards animalia vertebrata we with certainty know of only one species ') common to both zones, and among land- animals not one common distinguishing genus occurs. Among these the ge- nera which are represented in both zones are in general of a eosmopolitical nature. It is among the inhabitants of the seas that we find examples of an agreement between the zones. Within the class of Mammalia such an example occurs among the Phocidae in the genus Ci/tfophora NlLSSON, al- though that genus includes a species from the West-Indian seas. From the Arctic Ocean we have the species Cystophora cristata (Erxl.) and from the Antarctic Cystophora leonina (Lin.) or proboscidea (Desm.) Nils., which however has by J. E. Gray, though apparently without sufficient reason, been considered a separate genus, Morunga. Within the same class we have among the Cetacea the genus Delphinaptervs or Beluga with two spe- cies: D. leucas (Pallas) from the Arctic and JJ. Kingii (J. E. Gray) from the Antarctic Ocean.
The feathered vertebrata or birds which on their swift wings move to distant tracts with great rapidity, not unfrequcntlv afford, as might be expected, examples of a very extensive geographical dispersion, and we find in the Arctic zone many species common to Europe, Asia and North America, which have been termed circumpolars, and others that are extended from the polar circles to the Equatorial regions. Nevertheless perhaps not
') Otus braehyotus has been mentioned by r'Orbigny as found at the Strait of Magellan, but Gould considers it to be a different species and calls it Otus Gala- pagotmsis as also occurring in the neighbourhood of the Galapagos Isles (Voyage of Beaglel, but Sciilegel has since (Museum des Pays-Bas) quoted that form, marking it however with a note of interrogation, under the name of Otus brachyotus. D'Ok- b.igny has also taken up the ProceUaria glaciates as found in the Strait of Magellan, but it has since been found to be of a distinct species, and has even been referred to another genus, lludassoica Reichenbach, Thalass. glacialoides Reich. Bomip. D'Or- bigny has further stated that Hirundo rustica, urlica and riparia as well as Totanus fuscus are found in Patagonia, but in this also he appears to have confounded dif- ferent species.
6 W. LlLLJEBORG,
more than one species can be named, that is common both to the Arctic and Antarctic zones. This is the Falco peregrinus Lath., which has in- deed received several different specitic names, as the slight varieties of co- lour which it displays in widely separated localities, have by some ornitho- logists been looked upon as indicating different species, though for our own part we cannot but agree with Schlegel ') in considering these as me- rely local varieties. The variety that occurs in North & South America has been called Falco anatum Bonap., and the Australian form has been named Falco mebinogenys GOULD.
The fishes found in the fresh waters of Patagonia consist, it ap- pears, only of two or three species of the Salmonoid family, as is also the case with the fresh water fish belonging to the most northern fauna.
Among the section of Mollusca are some instances which present the phoenomenon of an extensive geographical distribution, though their capabi- lities for locomotion are very limited. Thus for example the Tercbratnla caput serpentis is found from Spitsbergen to the Mediterranean and on the eastern coast of North America, and the Rhynchonella Psittacea from Spitz- bergen and Greenland to England. Massachussets and Sitcha on the western coast of N. America. Some species c. g. Secxicava arctica, 1 emts pullastra and Pecten purio are found both on our northern coasts and at the Cape of Good Hope though not in the intermediate tropical regions. We may per- haps be able to show with certainty any species distinctly belonging to the Arctic Zone, which also occurs in the Antarctic, though one or two peculiar genera have been found that have their representatives in both Zones. Hronn *) states that the species of Litnaeina which belongs to the south Polar Ocean can not be distinguished from the Limacitta arctiea be- longing to the northern, but it has by WOODWARD 8) been classed as a se- parate species with the, name Limaeina antarctica. That genus has no re- presentatives in the intermediate seas. The same is the ease with the ge- nus of PunctureUa, which embraces two Species, of which the one belongs to the arctic the other to the antarctic seas (in the neighbourhood of Tierra del Fuego). Of the genus Clio we have in the northern Polar Seas the CHo borealis, which is there found in such plenty as to constitute a considerable
portion of the Greenland Whale's food. Passing over the intermediate oceans
') Museum des Pay-Bas. 1 Livraisim. Falconcs. p. 1.
') Klassen unci Orel nun yen des Thierreiche. 3. Bd. p. 648.
*) Manual of the Mollusca p. L'uT.
ON THE LYSIANASSA MAGELLANICA ETC. 7
that genus is according to Woodward represented in the Antarctic Ocean by some few species, but according to H. & A. Adams1) by only one, the Clio anstralis Brug. The genus of Buccinum, including about 20 typical species, also belongs only to the arctic and antarctic seas. The same is the case with the genus Trophon, containing about 14 species, the chief part of which are from the Northern Seas (Woodward).
The Bryozoa often present examples of more widely spread geo- graphical distribution than other animals of the lower section, and of these the same species have been occasionally met with in both the arctic and antarctic seas. Thus for example the Retepora celhdosa, which is not un- common in our arctic region, has by Ross been found at a depth of 1620 feet in the South Polar-Sea 2), Lepralia Malusi Busk, at Cape Horn, in the Mediterranean and in northern Europe, and Flustra foliacea Esper in the Southern Ocean and in Northern Europe 3).
The difference between the Crustacea, which strictly belong to the Arctic & Antarctic Zones, is not so great as that between them and those which are found in the warmer seas, but we know as yet of only 2 species common to both, both of the lowest group, and only a few peculiars genera common to both Zones. Our knowledge af the Crustacea of the Antarctic Re- gions is however as yet so imperfect as not at present to justify the utte- ring of a decided opinion on the relation, that may exist between the ant- arctic & the arctic Zones in this matter. The following genera are common to both zones. Order Podophthalmia: LithoJes Latr., Munida Leach, Euphausia Dana4). Suborder Amphipoda: Orchestia Leach, Anonyx KroYER, Ijihimedia H. RATHKE, Atylus LEACH, Amphithoe LEACH, 1 ' fyperia Latreille, Themisto Guerin-Meneville , Cyamus Lamarck. Suborder Is o- poda: Idothea EABRIC. , Ghiptonolm EIGHTS, 1'orcellio LATR., Oniscus LlN., Jaera Leach, Sphaeroma LATR. 5); Order Copepoda: Cetochilus ROUS-
') The genera of recent Molluscn. T. 1. pag. 62. 2) Bronn: Klassen und Ordnungen etc. 3 Bd. p. 90. s) Ibid. p. 92.
4) Mr Th. Fries has kindly presented to our University's Zoological Museum some specimens of a Schizopod that appears to constitute a link uniting the Eapktauia Dana with the Thysanopoda M. Edwards, found by him after a storm thrown upon the shore of Warangerfjord in the norwegian Fiunmarken. The last pair but one of the truncal feet has only the outer branch or palp, very small, and an almost im- perceptible rudiment of the inner branch or stem. We shall call it Euphausia glacialis n. sp. , as it appears to approach somewhat nearer to the genus Euphausia.
5) We have found at Oresund (the harbour of Landskrona) a species of the
8 W. LlLLJEBORG,
SEL DE VAUZEME; Order Cirrhipedia: Lepas LlN., Balanus LlN., Coro- nula Lam., Verruca Schumacher. Of these genera only the Lithodes, An o- nyx, Themisto and Glyptonotus are peculiar to the Zones in question. The others are of a cosmopolitical nature, and are met with also in the temperate and torrid Zones. This is partly the case with the genera Anomix and Themisto, but the greatest number of the species in those genera as well as , with respect to the first named genus, the greater plenteousness and larger dimensions of the individuals, sufficiently indicate preference for the polar seas and more especially the northern. The genus Atylus according to Spence Bate l) numbers 16 species. Of these 4 are from Greenland, Sweden and Norway, 3 from England, 2 from Southern Europe, 1 from N. America, 1 from Valparaiso, 3 from Tierra del Fuego, 1 from the Cape of Good Hope and one from New Holland. It belongs then more especially to the cold & temperate waters, but is hoAvever in a very considerable degree of a cosmopolitical nature 2).
Of the 4 above-named distinguishing genera, Lithodes comprises 10 species, of which 6 are from the northern seas — North-Sea, N. Atlantic and Icy Ocean and from the NE-coast of Asia — 1 from the eastern coast of America, Lithodes australis Bell 3), and 3 from Tierra del Fuego and the Antarctic Ocean; Anonyx about 28 species, of which 20 are from Green- land, Sweden and Norway and England, 4 from the Eastern coast of N. America, 3 from the seas of Nort-eastern Asia, and one from Tierra del Fuego ; Themisto 5 species, of which 2 are from the Antarctic Ocean, 1 from the southern part of the Atlantic & 2 from Greenland (S. Bate) ; and lastly Glyptonotus 3 species, of which 2 are from the northern Icy Ocean and Baltic and 1 from the southern Icy Ocean. The genus Lysianassa M. Edwards, of which Spence Bate reckons 15 species, comes so near the genus Anonyx Kroyer, that we as yet have no certain characteristics, by which these two genera can be distinguished, and it is probable that Kroyer would not have set up Anonyx as an separate genus, had he been aware that also the 2:d
genus Sphaeroma, S. rugicauda Leach, which lists as yet never been discovered on the western coast of Norway, but probably exists there.
Whether the species of the genus Ipfrimedia found at Tierra del Fuego belong reaally to that genus is not quite certain.
M Catalogue of the specimens of Anipbipodous Crustacea etc. p 133.
*) Atyhu axrimahu (FaBB.), which had previously only been found in Green- land, we have met with at Molde and Cbristiansund in Norway :it a depth of 40 — 50 fathoms.
5) British Stalk-Hyed Crustacea, p. Iti4.
On the Lysianassa magellanica etc. 9
pair of the truncal feets have a claw. If we unite the two genera, the geographical distribution will require some little modification, as becoming somewhat less arctic or antarctic. Of the 13 species of the genus Lysia- nassa 2 are from Greenland and Norway, 7 from the Mediterranean, the Atlantic, England and Norway, 1 from Van Diemens Land, 1 from the Cape of Good Hope, and 2 from Rio Janeiro. None of these species is common to both the arctic and antarctic seas. We have not included the Lysia- nassa magellanica in this calculation, because, as we in the following pages shall show, it constitutes the type of a separate genus. In fact the Cirri- peds are the only other order, in which we have examples of species com- mon to both these zones, for the Lepas Jlillii and fascicularis, which are spread over the whole earth (Darwin), are also found in both the above mentioned zones.
The genus Glyptonotus has been formed by ElGTHS ') to define a gi- gantic species of the Idothehbe, found in South Shetland, which approaches the Idothea entomon and is accordingly genetically united with it and with the Idothea Sabini. It has been called Glyptonotus antarctica ElGTHS (according
to DANA). This genus, the geographical distribution of which is, as we have before stated, confined to the polar tracts, although the one species, Ghiptonotus entomon (LlN.), as a relic of an extinct glacialfauna (8. Lovkn), is still occasionally found in more southern parts, as for instance in the Baltic, accordingly presents a most striking and remarkable example of the coincidence of disposition that exists between the two marine polar fauna}. The consideration of this lessens in some degree OUT astonishment, at the discovery of so higly developed a species as the Lysianassa magellanica common to both the arctic and antarctic zones.
Professor Til. Fries has been kind enough to furnish us with some information relative the Flora of the arctic and antarctic Zones, which we here communicate, as being a valuable assistance in judgeing of the relation between the Fauna; of these Zones. "Among the vegetable productions of antarctic America there are not a few found that also belong to the Flora of Europe. The greater part of these however consist of such easily ac- climatized species as Senecio vulgaris, Taraxacum officinale, Sonchvs olera- cens, Galium aparine, Brassica campestris, Capsella. bursa pastoris, Stellaria media, Urtica urens etc., which are now found, one may say, spread over the whole world. Among the vegetables which cannot be referred to this
') Transact. Albany Institute. II. pag. 331. (according to Dana).
Nova Aila Reg. Sue. Sc U]s., Ill' Sme. 2
10 W. LlLLJEBORG,
class, we especially remark certain hill plants common to the mountains of Europe and those of the antarctic Zone, but not met whith in the inter- mediate tracts. Of these Hookek enumerates Erigeron alpinns, Carta /estiva, Phleum alpinum and Trisetum subspicatum, but it is probable that on closer examination these will be found to be nearly related but different species. A remarkable example of a species common to both the Arctic and Ant- arctic regions and not met with elsewhere, is afforded by the beautiful and easily distinguised moss-species Usnea melaxantha, which is met with in Greenland and Spitsbergen as well as in New-Zealand and the most southerly portions of America. The only difference between the northern and southern forms is that the latter seems more thriving and fructifies richly, whereas the former is a more delicate plant and has never yet been met with in a fructificating state. It is also curious that a so remar- kably distinct form as the Nephroma arcticnm, which is so generally met with in the northern alpine and subalpine regions, should nowhere else be represented by any analogous or similar form excepting at Magellan's strait, where the very similar and nearly related Nephroma antarcticum is met with. Among phanerogamous plants the genus Empetnm presents the same phcenomenon, being in the North principally represented by the E. nigrum, whereas in antarctic America the E. rubrum is the prevailing species, unless (as I have lately seen asserted) this latter be also found in Northern America."
We now proceed to describe the remarkable Amphipod Lgsianassa magellanica.
It differs in many important features very considerably from the other species included in the subfamily Lysianastina, and we arc accordingly in- duced to consider it as the type of a separate genus. The first basal joint of the lower antennae is large and swelling, and uncovered at the side of the head, and is limited above by a projecting point of the slicll of the head, which gives the head, when seen in profil ■ peculiar appea- rance. The first pair of maxillae is furnished with a thin and long palp, at the end of which arc two or three coarse bristles or small prickles. The 7'1' caudal segment or caudal appendage, which is deeply forked, is not provided with any moveable spine at the extremity of the lobes. At least one such spine is found in all other Lvsianassina which have the caudal appendage forked. On account of its extensive geographical distri-
ON THE LYSIANASSA MAGELLANICA ETC. 11
bution, we give to this genus the name Eurytenes 1), and characterize it in the following manner:
EURYTENES, nov. gen.
Corporis forma crassa et robnsta, epimeris magnis et pedibus brevibus. Antennoe superior es fiagello appendicitlari proeditce, pedunculo crasso et ejus segmentis 2:do et 3:tio brevibus, et jiagelli segmento l:mo longo. Antenna' in- feriores segmento pedunculi hmo magno et inflato et extus visibili. Mandibular palpAgerce acie Icevi et tuberculo molari magno instructae. Maxillae l:mi parts palpo biartictdato angusto, ajdce duas vel tres setas vel acideos minores mobiles gerente, et earam ramus interior latus et brevis et setts multis ciliatis in- structus. Maxillipedum lamina trunci segmenti 2:di, sive lamina exterior margine interiore tenuissime noduloso, et eorum palpus ijuadriarticulatus et unguiferus. Pedes trunci sive thoracici l:mi et 2:di parts subcheliformes , illi validi et breves, ungue bene evoluto , hi longiores et graciliores, ungue minu- tissimo. lleliqui pedes trunci forma solita, robusti. Laminae branchiales simplices minimeque pectinatini pdicatae. Pedes caudales ultimi paris ramis lamellosis. Segmentum 7:mum sive ultimum caudae profunde bi/idum, laciniis accuminatis ad apicem vero non spiniferis. — Tantummodo una species:
EUKYTEINES M VGELLANICUS (II. Milne Edwaiids)
Lysianassa magellanica, 11. Milne EDWAKDS: Annales des Sciences natu-
rclles, 3:me serie, Zoologie, Tome 9:me; 1848; pag. 398. „ „ C. Spence Bate: Catalogue of the specimens of
Amphipodous Crustacea in the Collection of the British Museum, pag. 66, tab. X, tig. 5. — 1862.
Description: Length of body from end of caudal feet 215/16 inches or 73 millim. The three specimens we possess, which are all females, are all of the same size. Form of body (PI. I. Fig. 1) stout and strongly built, with the 2"1 to 4 ''pairs of epimera (coxae S. Bate) large, with the truncal feet, with the exception of the 2"*1 pair, short and strongly formed. The l:st truncal segment's epimera less than usual with that group, which causes the base of the lower anteniue and the appendages of the mouth to appear exposed. The last segment of the trunc and the first 5 of the tail have above a low longitudinal ridge, and the 6:th tailsegment has above on both sides a ridge, extending backwards, and terminating in a compressed obtuse
') From the Greek tvQitizrtjc, which signifies widely stretched.
12 W. LlLLJEBORG,
process. The 4th truncal segment's epimera, which are the largest, are almost rhomhoidal with the corners rounded off, and witli a wide hollow at the upper and back corner. The 5th pair of epimera are almost rect- angular with the corners rounded off, and a little hollow in the middle of the lower edge, into which hollow the upper edge of the 5"1 pair of feet's second joint is inserted. The epimera of the l:st tail-segment have the lower and back corner rounded off, and those of the 2"li segment have the same corner extented into a short point, from which on the outer side of the epimera a tolerably sharp edge or ridge (Fig. 21) stretches itself obliquely upwards and forwards. The 4th tail-segment's epimera have at their lower extremity a large spine turned backwards.
The eyes are not visible and have accordingly not been introduced in the figure of the animal, but seem, judging from the inner portions, to have been large and kidney- or bottle-shaped, and red.
The head (fig. 2) is somewhat convex, the forehead almost trun- cated, and has in the middle only an obtuse angle, as a slight indication of a rostrum. On the sides of the head is a projecting obtuse-angled lobe between the upper and lower antenna. The hollow under that lobe is ter- minated at its lower extremity by a projecting process.
The shaft of the upper-antennaj is short, and its first joint longer than the two following together, and the 2ml a little longer than the third. The flagellum consists of about 27 joints, the first of which is a little shorter than the shaft, and has on the inner side numerous long thickly set bristles of a brownish colour. The flagellum when laid back reaches about to the middle of the 2"'1 thoracal segment. The flagellum appcndieulare or ap- pendage, which is but litttle shorter than half the flagellum, consists of 9 joints of which the first is the longest.
The lower-antennae are more than double as long as tin' upper, and their flagellum consists of about 56 joints furnished with long bristles on the under-side. The first joint in their shafts (Kig. 1 & 2, b') is espe- cially large and distended, and for the greatest part of its length unpro- tected by the head-shield. The 2nd segment of the said shaft has on its underside evidently a sharp process. ("Olfactory denticle," >S. BATE).
The upper and underlips [Fig. .">, e & 1i^'. I & 5), which are about 5 millim. long, are of a very complicated construction. The former consti- tutes the central piece (a) and the latter the two side pieces (l>, b) which are united at the base so as to be in some degree moveable, and can ap-
On the Lysianassa magellanica etc. 13
proacli to or recede from one another like jaws, to which they have some resemblance. The external build is composed partly of thinner chi- tinous laminae and partly of more solid chitinous ribs which form the solid support of the former. The upper lip is somewhat distended and convex and has near its apex a sharp indentation, from which that apex bends itself inwards almost like a bird of prey's beak. The middle portion of the inner or palate side has two bristley-ribs united both in front and back and diverging in the middle, separated at the back by a notch from the curved apex of the beak. The side-pieces of the under lip are more solid but thin and flattened though uneven. (Fig. 5. b, b from the exterior, fig. 5 from the interior). At the projecting, free, indented end (a) they are yellow- ish, and their inner edge from the point to the base is armed with a thick row of short bristles, so that they are evidently employed in dividing the animal's food and conveying it to the swallow. The mandibular are inserted between the upper and lower lips and are thus for the greatest part of their length separated from the maxillae.
The niandibulac (Fig. 3. d. J and figg. li & 7, the left) are about 5 '/j millim. long and peculiarly strongly formed. The masticating extre- mity (Fig. (i (i from the interior) which is bent inwards as we see it in fig. 7. a, has a sharp, cutting and even or toothless edge. This cd^c has but a slight groove on each side, from which proceeds on the inside a curved and somewhat raised line. At the middle of that line is a very small and simple accessory process ("processus acccssorius", BliiZKLirs ') ), and from that a raised edge thickly armed with at first somewhat coarser and longer but afterwards shorter and finer bristles, which afterwards passes over to the "tuberculum molare" BruzkuuS, (b) of which it forms the outer edge, and when there is thickly set with short bristles. Between these outer edges the tuberculum molare is hollowed out. On the fore side of the man- dibula about halfway between the masticating end or edge and the insertion of the palp is a strong almost ball-like notch (Fig. (5. c). The palp (</) is large, and consists as usual of 3 joints of which the middle one is the largest and longest, and is broadest in the middle, and it, as well as the 3! joint, carry a number of sharp bristles. The palp reaches to about the end of the last joint but one of the lover antennae's shaft. Fig. 7 shows the left mandibula seen from the outer side. As the masticating extremity
') Bruzelius (Skancbs Amphipoda Gammaridea) denies the existence of such a process in the genus Anoriyx; we have however found it in the most species.
14 W. LlLLJEBORG,
is sharp and cutting, it is probable that it serves to divide the food, which is afterwards ground by the tuberculosa ruolare J).
The first pair of maxillae (fig. 8) are also of a strong and solid con- struction, and their length about 7 millimetres. The outer branch 2) (a) is, at the upper end (a'), spread out somewhat in the manner of a hand, hollowed on the inside, and armed at the edges with 11 stout moveable spines, which with the exception of some few of the longest are furnished on the one edge with side-prickles. The same branch's inner edge is also bristled. The inner branch (//) is thin, soft, short, and broad, and has at its point about 11 ciliated bristles. The palp (c) is two-jointed and rea- ches beyond the point of the outer branch. The second joint is in the form of a sabre and has a pair of short bristles at the point, as also a pair of very small moveable prickles. These maxillai are in immediate contact with the mandibulae.
The 21"1 pair of maxillae (fig. 9) are smaller thinner and of less solid construction. Their length is 5 millim. They consist of two bran- ches, and are of about the same obtusangular lancet-form, and the outer (a) l/, longer than the inner (b). Both have at the poiut and along the inner edge numerous filiated bristles.
The maxillipeds (Fig. 3 g g & fig. 10) are 10 millim. long, and consist as usual of a stem of 2 joints (fig. 10. a, b) and of a palp (c). On the inner side of the basal joint of each maxilliped is an oblong almost troncatly terminated lamina (lamina interior) which at the point and the inner border bears a number of longer bristles. These laminae (Fig. 11) come into contact with eachother, and form by their union a raised ridge on the inner side of the maxillipeds, and reach to the end of the first third of the second joint's lamina. The second joint of the stem (Fig. 10. b) is on the inner fore-part expanded into a large almost elliptic lamina, the so called lamina exterior (fig. 3. g'. g' & fig. 10. b'), rounded off in front and there famished with bristles. Its Interior edge (fig. 12), is made uneven by small knobs. The Palp (fig. 3. g" g and fig. 10. r, c) consists of 4 joints of which the 2"d is the largest, and the fourth has the form of a claw, and all. with the exception of the last, are furnished with numerous bristles.
') Spenee Bate considers thai this form of the mandibole indicates that the animal consumes vegetable food.
5) The outer branch may he considered as the stem of these maxillae.
On the Lysianassa magellanica etc. 15
The truncal feet1) of the first pair (fig. 13) are short, particularly strongly built and subcheliform. Their length is about 15 millini. The 2nd joint2) is the longest and the hand or 6"' is almost rectangular, rather smaller at the lower end, with the fore edge somewhat bent and the back concave and bearing about 9 clusters of bristles. Its lower end is obliquely hollowed out, and in front of the notch is fastened a strong, sharp, crooked and flexible claw. The 5th joint which is triangular is shorter than the 6'1'. Its feet are without gill-sack and lamina to cover the eggs.
The truncal feet of the 2hl pair (fig. 14), which are also subcheli- form, are finer and longer than the preceeding. Their length is about 30 millim., and the breadth of their 2"! joint is 2 millim. The 3nl joint is scarcely half so long as the 2 "' , which is almost of uniform breadth and somewhat bent back at the lower end. The hand or 6:!' joint is shorter than the 5'1', oblong and of almost uniform breadth, and obliquely rounded off at the lower end. It lias numerous long bristles, as have also the joints already described. The claw is very small and hardly perceptible. The lamina for covering the eggs (b) and the gill-sack (c) are fastened to the 1st joint or epimerum (a). The former is narrow and of uniform breadth, and bordered with long bristles; the latter simple, of considerable size, and almost kidney-shaped. One observes on it only a few small irregular wrinkles which have probably arisen after death.
The 3"1 and 4lh pairs of truncal-feet (fig. 15, right foot of 3"' pair) are similar to each other and about 24 millim. long. The 4th joint is longer than the 5th, terminated obliquely at the lower end, where, at the obtuse projecting point, it is provided with long bristles. The 6"' joint is about the same length as the 4', almost uniformly broad, somewhat curved, and at the back border provided with bristles. The claw is strong.
The truncal-feet of the 5lb— 7th pairs (fig. 16, the 6fh pair, right foot) bear a close resemblance to eachofher, and differ from all the preceeding
') We here adopt the denominations proposed by T. THOBELL, (Ofvers. af Kongl. Vetensk. Akachs Fiirh 1864. pag. 9) according to which that part of the body of the Crustacea, which is commonly called thorax, is denominated trunc (truncus), and that part, which by Other writers is called t,l>th>m<>n, is denominated tail (eauda).
2) The joint which we call the 2"' has in general been considered as the 1". We consider that that part, to which the gill-sack and lamina for covering the eggs are attached, is the first, although it may be joined to the epimerum, or perhaps more correctly speaking (according to Spencb Bate) constitutes what is called the epime- rum. The gill has no doubt an insertion similar to that of the Podophthalmia.
16 W. LlLLJEBoRG,
in having the 2'"1 joint strongly expanded, and in being directed forwards. The foot of the 61' pair here represented is 24 millim. long. The 5th pair, as may be seen in fig. 1, is distinguised by having the Ist joint larger, and the back part somewhat higher than the forepart, which is almost semi- circular. The second joint is shorter, and has the posterior spreading portion almost semicircular. In those of the 6* pair this joint is rounded orally, and the border rough with indentations or notches. These inden- tations are only slightly apparent in the 5 ll pair. At the base of the la- mina for covering the eggs, attached to these last mentioned feet the fe- male genital aperture is very clearly apparent. The 7th pair of truncal-feet differ from both the proceeding in having the 2nd joint longer and oval, with small but clearly visible indentations in the back border.
Each pair from the 2 ld to the 5"' inclusive is provided with laminae for covering the eggs, all of the same form as that, which is represented in our plate, and which belong to the 2nd pair: the G,h and. 7 * pairs are destitute of this appendage. In the 2nd, 3"1 and 4th pairs the gill-sack is of the same form (fig. 14 c). In the 5tlr and 6th pairs it presents the same form (fig. 17), but differs from the proceeding in being of a firmer structure, and having a caecum-like appendage {b) containing granulated matter, con- sisting of fat globuli and other formative particles. In the 7''1 pair this appendage is wanting, but in other respects its gill-sack is similar to that of the foregoing pair.
The tail-feet of the first 3 pair's, or so-called swimming-feel, arc of the usual form (fig. 18, 1 ' pair, right foot) and arc similar to eachother excepting that they diminish in size, so that the 3,a pair are shorter than the foregoing. They consist of a strong, oblong stem, (a) tapering towards the lower end, and two terminal branches (i,b) tapering gradually to a point, longer than the stem, composed of a great Dumber of joints, and on both sides provided with long ciliated bristles. The 1- pair are 19 millim. long.
The tail-feet of the last 3 pairs (i. e. the 4th, 5* & 6*) are as usual formed for leaping, and consist of a stem and two simple terminal branches without swimming-bristles. The 2 first pairs are as nearly as may lie similar to one another, hut are longer and have their terminal branches more pointed than those of the 6* pair. Those of the 4 h pair (Fig. 19, right leg, outer side: are somewhat longer and slenderer than those of the 5*J their length is about 14 millim. Along the outside of the Stem there is a raised ridge, and a similar elevation runs along the middle of both terminal branches both on the outer and inner side, though higher
ON THE LYSLANASSA MAGELLANICA ETC. 17
on the former, and highest on the outer branch (a): This branch has at its point a spine, separated by a suture, and may by this be distinguished from the inner (b), which is moreover somewhat shorter and broader.
The tail-feet of the last or 6 th pair (Fig. 20, right foot) are shor- ter, broader and more moveable. Their length is 8 millim. The stem is almost rhomboidal, with the lower external angle projecting acutely. The external terminal branch (a) is also somewhat longer than the internal (b) and has also a detached spine at the point. Both branches have about the same shape, are tolerably thin, though thicker at the outer edge, of a broad lancet form, and are provided with numerous bristles on the inner border. Also the stem has similar bristles on its inner border and at the outer and lower acute angle. These feet stretch somewhat behind the ends of the other tail-feet and even somewhat further back than the 7,h tail-segment.
The 7h tail-segment or, as it is called, caudal appendage (fig. 21) is of considerable size, extending almost to the ends of the last tail-feet. Its length is 6 millim. It is forked a little below the middle, and the ope- ning between the two halt's somewhat wider towards their termination. It is tolerably thick with a blunt ridge which on both sides goes parallel with and close beside the external edges of the lobes, with a hollow along the middle between the base and the opening. The lobes are brought up smoothly to points, and are destitute of the moveable spines at the end, and no spines are visible on their sides.
The Group of the family Gammaridae, which the subfamily Lysia- nastina constitutes, seems to be tolerably natural and well defined, but it happens with it, as with many other natural groups, e. g. Felidae among mammalia, Ct/prinidae among fishes, etc., that the forms belonging to it, especially the different species within the same genus, not unfrequently pre- sent such slight differences, that it requires a very minute examination to distinguish them. This group is distinguished by the form of the tipper antennae and mandibular, the former having a peculiarly thick shaft of which the two last joints are very short, and the latter a more or less sharp edge with few or no teeth (fig. 52), and the inner process (a) — processus ac- cessorius, Bruzelius — little or sometimes not at all developed. To this may be added that the 2nd pair of truncal feet are very long and slender, having in general a very small clawr, which however is in one instance absent.
Nova Acta Reg. Sue. So. Ops., I lie SSrie. 3
18 W. LlLLJEBORG,
Dana *) and Spence Bate '), who give to this group a wider range, lay stress on the size of the epimera, (coxae, S. Bate), but this is no distin- guishing feature, for there are others e. g. Stegocephalus , Pleustes, Odins (Otus S. Bate) with equally large or larger epimera. Dana reckons up as belonging to this under-family the genera: Lysianassa M. Edwards, Phlias Gueiun-Mexeville, Opis Kroyer, Uristes Dana, Anonyx Kroyer and Urolhoe Dana. From these we reject Phlias, Uristes and Urotlwr, the two first on the ground that they have no flagellum appendiculare on the upper antennae, and the last on that of its not having a thick shaft to the upper antennae, and moreover that it nearly approaches the genus Phoxits Kroyer. SPENCE Bate includes in the sub-family of Lysianassidae the genera Lysia- nassa, Anony.r, Pontoporeia KROYER, Opis, Ichnopus C'OsTA, Ca&uoma COSTA, Alibrotus M. Edwards, Jlyale H. Rathke, PUias and Uristes. Of these we reject Pontoporeia on acconnt of the different form of the mandibular and the genera AUbrotus and Hyale on account of the entirely different con- struction of the upper antennae and second pair of feet. The genus Ich- nopus appears to us identical with that of Lysianassa, and as regards Phlias and Uristes we have already stated our opinion. We therefore include in this sub-family only the genera Lysianassa, Anonyx, Callisoma and Opis, to which we add the two new genera Eurytenes and Acidostoma. In order to illustrate our view of the family Gammaridae and the underfamily Ly- sianassina, we adduce here first a tableau of the families included in the suborder Amphipoda, and next a similar tableau of the sub-families com- prised in the family Gammaridae, and, as therewith connected, of the ge- nera within the same family belonging to our-own fauna. We do this the rather since no new synopsis of the kind has appeared since the year 1859, when Doctor Bruzelius in Kongl. Wctenskaps-Akadeniiens lland- lingar. (new series) Vol. 3, p. 1. published his excellent paper "Bidrag till kiinnedom om Skandinavicns Amphipoda Gammaridea" (Contributions to the knowledge of the Amphipoda Gammaridea of Scandinavia). Our knowledge of these Crustacea has since that time received considerable additions from the labours of Candidate A. BOEGK, Professor M. SAKS, and the observations we have ourselves had the opportunity of making on the Western Coast of Norway, and we can accordingly now include 36 genera as belonging to our fauna, while BBDZELIUS lias only 19. Of these genera 4 are new viz. Eurytenes, Acidostoma, '/'iron and Oediceropsis , of which genus Eurytenes
') Bxplor. F.xpe.l. Crustacea. T. II. p. '.MIS. *) Catalogue etc. p. 64.
Pag. 18.
Gammaridce. Antenna luperio-
flagello appendicular!
u radius,
; lapul antica ■
nou produc-
tuni.
Pcdunculus
nutemiariiiii snp
cra*sus, segmeutis
llttobUI nil, mi- l.rc-
vlbuj. Miiiiiiibulio
solito rnodo .dentnta.
. Lysinnnssina, Dana. MuxilltO l:mi pnris . - .
2, StiUfamilia ;
Fontnporciiiu, Dana. Pedes trunci (thorneici) 2:dl pnris
palpo pficditic, Pedes trunci fihovocici,) l:mi, pnris iiadom 2:df pans . . .
'manu nibchelfform
11 Biibcholiformi. Seg- m on turn l:mntp psudunculi union nar am inferiorom ml Intern capitis OXttlfl . . .
Genera:
1- LysisnaJsa, M, Edwards.
isibilo, magnum et inrlutmu 2. Eurytcncs, nov. gen.
3. Anonyx, KrGycr.
i crussiores , sed longiorcs ct nngue. indistim
palpo cardites inguc cardites
unguiferi
mnnu BiibchjUformi vel cheliform manu BubchuSformi priedlti . .
mediucris et tegmenta -jus ultima duo
■longnta. —
> producrum .ii basin antauni
3. yuLAi mil
Gammarinn Pedes maxillures
Dai
lamioii intcrioribirs pnediti, liimin&qnc haj et extcrioroa . .
bene cvolutte ct basin palpi cxccdentOfl, Segmentum 5:tom pedum trunci l:mi ct 2;di pane lixuui ad
Spicooa BUpcriorcm segroenti 6:ti sivcfuninrticulatus g
manus. Pedum cuuduliuni ultimo- I rum ramus exterior I
lliiarticulatu?, longissiiuus ]o. ronrginem anieriorcm segmenti 6:ti aive manus u.
el interiores vix bnsin palpi astequcnlcs. Mandibular inter k . .
j
tdis!
(tutu tuliiformcs. — I Pedes I
(Pboxidcs, S. Bate; unci (ilioracici) l:mi et S:di pnris
mumi Bubohcliformi prffldlti. t clongttta ct solito modo formntn Seginentii Sidum ct 3:tium i [
palpi pedttn moxillnriui non prckausili a . 1 1
Ua et laminaria ]gi
cbelifotmi carcntcs jy
. Callisoma, Costa. . Acidostoraa, nov. gen. . Balliyporein, Lindstroni. Stcgoccphalus, KrOyer. . Pontoporcia, KrOyer.
, Gammarus, Fubricins.
Eriopia, Brozelios. . Eusirus, KrOyer.
. Microplax, nov. nom. (Idnaa, A. Boeck)
. Sicippe, Bruzelins.
Pardalisca, KrOyer.
Phoxus, KrGycr.
DrothoC, Dana.
Tiro
n , nov. gen.
5. Bobfomilia: Triscbisostomatina . . (Pi'OKtDjnatai , A. Boruk).
Bagcllo sppendiculai
carentcB,
pculi
'<llll|»lSltl.
Pedes trunci (thoi r'iei) T;mi puris UT In I'lli'uiihus . . .
tnulto longiorcs, segmento ultimo [lingua) longo, recto ct slilifornii.
C. Siibffiiuiliii : Oedicerina. Pedes trunci 7:i
niitecfdeiitilius non duplo tQngiores
parum product duplo vel magii longiorcs.
Siiruicuiiiii] 5:tum
Five cnrpUB (*cdi
trunci S:di pnris longc productum
post ice
. Trischinostotua.Esmark. . Oediceropsis, nov. gen. , Oedicerus, KrOyer.
parum (ongiovei segmento ultiiiK guiformi. Laramie pedum
\illariuni . . .
. Suljfaniilin : Lcucotho Mandibulra .
palpigerm. Manus pedum
mis . I
Jihcliformis
chcliformis, carpo digitum irumobilem eflicicntc . .
8. Subiitniili
lie lie
olutrc. - Atylinn.
Pedes trunci
l:mi paris manu
Pulptis pedum maxilljriuui .
non chcliformis. framis duuliu I Pedes eaudnles I Itiltimi pnris . . [rauio uno ,
pnlpo carcntcs
(quadriurticulntus tr
21. Mnnoeulodes, Stimpson.
22. KrGyera, Spenco Bate.
23. Leueothoe, Leacb. •a. Pleuites, S. Bate.
25. Montagun, S. Bate.
26. StenothoG, Dana.
| IN. 11 In III. ,111 ■■
lion elielifui-mi. piei'UQiquC fuli- el ie I ifunui.
Palpus pedum :' — j artieula- muxillurium .
Pedes trunci
Corpus (
lubchelifonn
Mnndibtilio
pal pi genu. " "anus poduri
Ii |,„n, . .
palpo i
iulto ninjor oadci :mi pnris . .
uidem l;ml pnris mugnitu- dine citaitar mqualiB. Epimera seg- mciUorum trund ante no-
acuminata .
•diusa ct .otttndata .
2". Odins, nov. nom. (Otus, S. Bate)
28. Ipbimedia, H. Rathke.
29. Laphystius, KrOyer.
30. Calliopius, nov. nom. (Calliope Lcaeh, mser. sc-
enndum S. Bate.)
31. Par&n]phithoe,B''UKclius.
. Atylus, Leach.
. D examine, Lcaeb.
non preliensili : tiequo cheliforra
ni'ijiio stibclicliformi ....
Bubfamilia: Ampeliseim
(Amiieliscides, S. Bate)
Oculi . . . .
. Acnnihonotus, Owen.
, Huploops, Lilljeburg.
Aiupclisca, Krfiycr.
g. 18.
Amphipoda.
Pedum cnuda- lium
paria adsunt. Normalia, S. Bate. Pedes maxi Hares lt« maxillipcdcs
porfeeti ct pnlpigcri. Segmcntn enn- daliii posteriori
non coaliia. Segmentuni eaudic 7:mnrn
Familiie:
Inminnre, integrum vel divisum I. Gammurid k, Daim.
2. Orchcstidje, Dann.
erussum ct tu- /magna. Mandibular berculifonno . Ipajpo plcrunv|uc enrontes , Epimera . . .!
Ipnrvti vcl rnodiocna.
(Mandibular pnlpigcnt; . ,
|
unnm vel plur paria ahtunt - |
- Aberrantia, S. Bate. Cauda |
lobsolcta. ^Corpus |
3. Cor a phi die, Dunn.
conlita 4. Chelurida,-, Allinim.
iniperfecti ct palpo enrontes 5. Hyperidre, Dana.
obsoleln, segmentis 6 composite G. Dulichidie, Dann.
clungutiim ct angustum 7. Cuprcllida\ Dann.
depressurn et latiusctuiim ....".. 8. Cyamidn:, Dann.
Pag. 22.
Anonyx. Lamina exterior pedum moxilln- rium margins interiore ...
Inloso.
Segm
I it
plus vel mi-
Anguli po- iteriorei Beg*
menti 3:tii eau-
atcrrimi, fere lageniformcs .
Species:
ampnlla(FhippB); KrOyei
cocci nei vet rubesccntc*. Pedes trunci Itml pans . .
plus vcl minus breves ct ro- busii. Unguii pedum l:mi pari* dente ar-
Laluum . .
longe porrectum 2. longipes, Spenec Bate.
3. gulosus, KrOyer.
scg-
ante medii
Manna pedum I:mi paris . .
roetangulnris , earpo lon-rdimidio segmenti 9edi longius.
gior. Scgmcniiini 3:tiura lOcns unguis validus
pcdUQl I:nii puns . . J ,
ldimiiiio pegmenti 2:di brevius.
IDona unguis minimus 4.
■ unguis minimus 5. p
oides, n.
sp.
sp.
Seguientum 3:tiuni CJjadlC ulrifiqua
postice ...
tnngissimi
hevc.
Fiwurn segmenti 7
non nnte medium segmenti ineipiens, et line segmentum breve et laic
■ dente?
G braehycercus, n. sp.
et gracillimi, segraento 3:tio diinidio 2:di longiorc. Unguis ped, l:mi j
nlc medium segmenti, Dens unguis ped. l:mi paris minimus 8.
ione medium (Ogmontj, Manus pedum l:uii paris curpo loiigior 9,
at urn 10.
mdic ineipiens
-I
l.-htirul,(|
|cmnrgimitum. Unguis pedum lani paris i 'sub-in
, denticulhn
Idiscrctis et majoriboB. Unguis pedum l:r
dentieuhitus ct serratus 11
uatuin. Unguis pedum l:mi paris intus dentieulatus et serratus 12,
[ennfertis et longis. Unguis pedum l:mi paris longissimus ct Levis . 13,
paris brcri6 !*■
heviusculo, et tantummodo aculeis parvis, paucis et diserctis intra murgincm positis, horrido. Oculi nigresccntcs 15
Bruzelii. A. Boeck. nanus, KrOyer. pinguis, A- Boeck. serratus, A. Boeck.
Edwardsli, KrOyer. litoralis, KrOyer. HolbOllii, KrOyer. obtusifrons, A. Boeck. tumidus, KrOyer.
On the Lysianassa magellanica etc. 19
lias been already described, and of the genus Acidostoma full details vill be given in the sequel.
The genus Tiron x) is typified in a species, found by us at Christian- sund in Norway at a depth of 30 to 40 fathoms, which we in our public lectures have called Tiron acatithnrus. n. sp.
Forma capitis ex parte cum eadem gen. Oediceri congruit, antennae superiores vero flagello appendicular! longo sunt praeditae, et pedes trunci 7:mi paris longitudine pedes anteriores aequant, et breves, crassi et unguiferi sunt. Pedes trunci l:mi et 2:di paris graciles, ungue tamen non flexibili instruct! Segraenta caudalia superne in medion lungitudinalitcr carinata, carina ad marginem posteriorem segmcntorum in aculeuin, qui in segnicntis 4:to et 5:to niagnus est, et adliuc in segmento 6:to observatur, excurrente. Antennae superiores longitudine pedunculo antennarum inferiorum ajquales. Frons aliquanto producta, basin antennarum superiorum obtegens, rostro brevi sed acuto. Oculi rubri. Longitudo circ. 10 milliin.
The genus Oediceropsis is also typified by a single species, found by us in the sea off Moldc in Norway at a depth of 40 or 50 fathoms, which we in our public lectures have called Oediceropsis brevicornis n. sp., the upper antennae being particularly short.
Forma corporis eidem gen. Oediceri valde similis, caput tamen rostro caret, et pedes trunci 7:mi pans, qui longuin et rectum ungucm habent, et longi et graciles suut, tamen pedibus anterioribus (j:ti paris non duplo — circ. sesqui — longiores sunt. Antennae superiores non finem articuli pen- ultimi pedunculi antennarum inferiorum assequuntur, et flagello appendicu- lar! carent. Antennae inferiores magnae, fere pediformes, articulo penultimo pedunculi ceteris majore et ad apicem infra setam magnam gerente. Oris partes appendiculares et hujus et anterioris speciei structura solita. Pedes trunci l:mi et 2:di paris forma inter se similes, manu subeheliformi, ovali, carpo postice aliquantum producto. Pedes trunci 3:tii et 4:ti paris parvi et graciles. Segmentum candale 7:mum integrum et parvum sed lammare. Pedes caudales ultimi ramis duobus angustis, fere aequalibus. Color flave- scens; oculi rubescentes, sed parum visibiles. Longitudo circ. 8 millim.
To the genera Microplax , Odius and Calliopius we have given new names instead of the names Iduna, Otus and Calliope, which had already before been given to other animals.
') Tu'owv Proper name.
20 W. LlLLJEBORG,
This being premised we proceed to give an account of such spe- cies of the sub-family Lysianassina as belong to our Fauna, describing somewhat more in detail those which are new to science or even additions to our own Fauna.
Gen. LYSIANASSA H. M. Edwards.
Pedes trunci s. thoracici l:mi pans manu subrheliformi carentes, vngue non fleocibili, seamento 6:to sive manu apicem versus attenuato ibidemque basi unguis vix crassiore. Mandibulae tuberculo molari minimo. Laminae eccte- riores maxillipedum tnaryine interiore noduloso vel laeviusculo.
We have in Sweden and Norway only 3 species of that genus di- stinguishable in the following manner.
Lysianassa. ( tissum. ( pectinatim plicatSB ... 1. gpinioorms (A. Boeck).
Segmentum ! J*"*"*
° nranclnalcs . . . .'
7:mum Cauda; ... ' I ... 0 ■- ,;• nr ■■ x
non plicatse 2. I ulih (Kroyer).
non fissum, margine posteriore con- l vexo 3. Costa:, M. Edwards.
1. L. Spinicornis (A. Boeck).
Jchnopus spinicornis, A. Boeck: Forhandl. ved de Skand. Naturfs 8:de
Mode, 1860, pag. 645.
Antenna; longte, inferiores superioribus longiorea. Scgnienta 2:dum et 3:tium pedunculi antennarura superiorum brevissima, et segnientum l:mum ejusdem pedunculi ad apicem infra aculeo armatum. Flagellum harinu an- tennarum maris circit. 85 — 100, et feminae circit. 60 — 66 aiticulis brevibus compositum, articulo l:mo ultimis duobus aiticulis pedunculi conjunctis lon- giore, et intus circit. 23 paribus fasciculorum pilorum transversis pra'dito. Flagellum appendiculare longum, aiticulis arc. 10. Flagclluni antennarom inferiorum articulis circ. 80 — 120.
.Mandihuhe acie obliqua, Qtrinqne dciiticulo obsoleto munita, processu accessorio carentes. Tubciciiluin niolarc parvum , lvuYxum, acuminatum et pilosum. Palpus magnua, segmento 2:do latiusculo, et 3:tio arcuate et ad latus alteram pectinatim aculeate. Maxilhe l:mi paria ramo exteriore valido, ad apicem aculcis magma, ex parte pectinatis, et pilia annate; ramo inte- riore minore apice pilos duos gerente; et palpo biarticolato, apice tnmcato, striate et aculeate. Maxilhe 2:di paria ramie anguatia et ad apicem acu- leatia. Pedes niaxillarcs sive maxillipedea laminia ioterioribna brevibus ad
On the Lysianassa magellanica etc. 21
apicem oblique truncatis et nodulis 2, et setis circ. 5; et laminis exterioribus niagnis margine interiore noduloso.
Pedes trunci l:mi paris segmcnto 6:to, sive manu, elongato et paullulum arcuato, ungue sat magno et intus fasciculo aculeorum arraato. Pedes trunci 2:di paris manu subcheliformi , obovali et setis lougis dense obsita.
Vesiculse branchiales pectinatim plicatse, plicis a racbi media exe- untibus.
Epimera segmenti 3:tii caudal angulo inferiore et posteriore aculeato. Pedes caudalea ultimi ramie elongato-lanceolatis, marginibus exteriore et in- teriore aculeatis et pilosis. Segmentum 7:mum caudas ultra medium fissum, laciniis contiguis, obtusis et aculeo ad apicem munitis. Oculi magni, reni- formes, fusci. — Longitudo 30 — 40 millim.
Ad Bergen et Trondlijem in Norvegia accepta, baud frequens.
2. L. Yahli (KroYER).
Anonyx Vahli, H. KroYER: Gronlands Ampliipodcr, pag. 5.
„ „ Idem: Naturhist. Tidskr. 2 Rffikke, \U\. 1, pag. 599.
„ „ R. BRUZELroS: Skandinavicns Ampbipoda Gammaridea;
Kongl. Wetensk. Akad:s Handliiigar, ny mljd, Bd. 3,
1 liaftet.
Segmenta 2:dum et 3:tium pcdunculi antennarum superiorum brevis- sima, diniidio segmcnti l:mi brevioia. Flagellom appendicnlare harom an- tennarum articulis 5 — G. Pedes caudalea ultimi ramis latiusculis, et eorum exterior marginibus piliferis. Segmentum ultimum caiuUc sive appendix cau- dalis fere usque ad dimidiam partem fissum, laciniis apice rotundatis. Oculi aterrimi, reniformes. Longitudo U — 20 millim.
Habitat ad oras nostras occidcntales a Bergen in Norvegia ad Fin- markiam, rara.
3. L. Costae, II. Milne Edwards.
Lysianassa Costae, H. MlLNE EDWARDS: Histoire naturelle des Crustaces,
T. IH, pag. 21. „ „ C. Spence Bate: Catalogue of the specimens of Am-
pbipodous Crustacea in the Collection of the British
Museum, pag. G9, tab. 10, fig. 11. „ „ Idem & J. O. Westwood: History of British Sessile-Eyed
Crustacea, Ampbipoda, T. I, pag. 74.
20
W. LlLLJEBORG,
This being premised we proceed to give an account of such spe- cies of the sab-family Lysianasdna as belong to our Fauna, describing somewhat more in detail those which are new to science or even additions to our own Fauna.
r
ft
Gen. LYSIANASSA H. M. Edwards.
Pedes trunci s. thoracici l:mi paris manu subcheliformi carentes, vngue non flecribili, seprnento 6:to sive manu apicem versus attenuato ibidemque basi unpuis vix rrnssiore. Mandibrdae tuberculo molari minimo. Laminae exte- riores ma.rillipedum margins iuteriore noduloso vel laeviusculo.
We have in Sweden and Norway only 3 species of that genus di- stinguishable in the following manner.
Tji/siaiiassa. Segmeutum 7:mum Cauda; .
tissum. Laminae
branchiales
( pectinatim plicate ... 1. xjnnicvrnU (A. Boeck).
I
I non plicata; 2. J 'uhli (Kroyer).
non fissum, niargine posteriore con- 1 vexo 3. Costa', M. Edwards.
1. L. Spinicornis (A. BOECK).
Irhnopus spinicornis, A. BOECK: Forhandl. ved de Skand. Naturfs 8:de
M6de, I860, pag. 645.
Autennse longse, inferiores superioribus longiores. Segmenta 2:dom et 3:tium peduneuli antennarum superiorum brevissima, et segmentnm l.-mum ejusdem peduneuli ad apicem infra aculeo armatum. Flagellum harinu an- tennarum maris circit. 85 — 100, et femime circit 60 — 66 articulis brevibos compositam, artieulo l:mo ultimis duobus articulis peduneuli conjunctis lon- giore, et intus circit. 23 paribus fasciculorum pilorum transversis prsedito. Flagellum appendiculare longum. articulis circ. 10. Flagellum antennarum inferiorum articulis circ. 80 — 120.
Mandibuhe acie obliqua, Qtrinque dcnticulo obsoleto munita, processu accessorio carentes. Tuberculosa molare parvum, reflexum, acuminatum et pilosum. Palpus magnus, segmento 2:do latiasculo, et 3:tio arcoato et ad latus alteram pectinatim aculeato. Maxilla l:mi paris ramo exteriore valido, ad apicem acoleia magnis, ex parte pectinatis, et pilis annate; ramo iute- riore minore apice pfloa duoa gerente; et palpo biarticolato, apice tnmcato, Striate et aculeato. Maxillse 2:di paris rainis angustis et ad apicem acu- lcatis. l'cdcs lnaxillarcs sive maxillipedes laniinis interioribns brevibus ad
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On the Lysianassa magellanica etc.
21
apicem oblique truncatis et nodulis 2, et setis circ. 5; et laminis exterioribus inagnis marline interiore noduloso.
Pedes trunci l:mi paris segmento 6:to, sive manu, elongato et paullulum arcuato, ungue sat maguo et intus fasciculo aculeorum armato. Pedes trunci 2:di paris maim subcheliformi, obovali et setis longis dense obsita.
Vesiculae branchiales pectinatim plicatse, plicis a racbi media exe- untibus.
Epimera scgmenti 3:tii caadee angnlo iuferiore et posteriore aculeato. Pedes caudalcs ultimi ramie elongato-lanceolatis, marginibus exteriore et in- teriore aculeatis et pilosis. Segmentum 7:nmm cauda? ultra medium fissum, laciniis contiguis, obtusis et aculeo ad apicem niunitis. Oculi magni, reni- formes, fusei. — Longitudo 30 — 40 millim.
Ad Bergen et Trondbjem in Norvegia aecepta, baud frequens.
2. L. Yahli (Kroyer).
Anonyx Vahli, H. KroyeR: Gninlands Anq)bipoder, pag. 5.
„ „ Idem: Naturhist. Tidskr. 2 Ka-kke, Bd. 1, pag. 599.
„ „ R. Bruzelius: Skandinaviens Ampbipoda Gammaridea;
Kongl. Wetensk. Akad:s Handlingar, ny fi'djd, Bd. 3,
1 baftet.
Segmenta 2:dum et 3:tium peduncoli antennaram saperioram brevis- sima, dimidio scgmcnti l:mi breviora. Flagellom appendiculare barum an- tennarum articulis 5 — G. Pedes caudales ultimi ramis latiuseulis, et eorum exterior marginibus piliferis. Segmentum ultimum cauda; sive appendix eau- dalis fere usque ad dimidiam partem fissum, laciniis apice rotundatis. Oculi atcrrimi, rcnit'ormcs. Longitudo 9 — -20 millim.
Habitat ad oras nostras occidentales a Bergen in Norvegia ad Fin- markiam, rara.
3. L. Costae, H. Milne Edwards.
Lysianassa Costae, H. Milne EDWABD8: Histoire naturelle des Crustaces,
T. IB, pag. 21. „ „ C. SPENCE Bate: Catalogue of tbe specimens of Am-
pbipodous Crustacea in tbe Collection of the British
Museum, pag. G9, tab. 10, fig. 11. „ „ Idem & J. 0. Westwood: History of British Sessile-Eyed
Crustacea, Ampbipoda, T. I, pag. 74.
I
22 W. LlLLJEBORG,
Lobuli laterales testae capitis anteriores longe producti. Epimcra magna. Epiraera segraenti 3:tii caudae angulo posteriore et inferiore aculeo brevi et superne vergente. Oculi niagni, supra approxiiuati , reniformes , fusci. Segmenta 2:dum et 3:tium pedunculi antennarum superiorum longi- uscula, una segmento l:mo longitudine circ. aequalia. Antennae superiores et inferiores apud fcminam longitudine circ. a-quales, flagello superiorum pilis longis, et articulis 9 — 11 composite, articulo l:mo brevi. Elagellum ap- pendicular biarticulatum articulo ultimo minimo. Mandibular acie utrinque denticido obsoleto, processu accessorio longo et tenui, tuberculo vel pro- cessu molari minuto, fere evanescente, tantummodo lobulum parvum seti- feruni praebente. Palpus mand. longus. Maxillae l:mi parte palpo biarticu- lato, ad apicem aculeis 5 brcvibus, quorum uno mobili; ramo intoriore sat magno. Maxillae 2:di paris ramo interiore latiore, oblongo-ovali. Maxilli- pedes laminis interioribus magnis, et exterioribus vix apiceiu articuli 2:di palpi attingentibus, ovatis, apice rotundato et non sctifcro. margineque into- riore laeviusculo et vix noduloso. Pedes trunci, l:mi paris segmento 6:to, sive manu, conico et ungue parvo. Pedes trunci 2:di paris gracilcs nianu fere oblongo-ovali carpoque breviore et minore. Pedes trunci 3:tii et 4:ti paris etiam graciles. Pedescaudales longi, et eorum ultimum par ramis sub- ulatia minimcque setiferis. Scgmcnttini 7:inum caudae, sive appendix cau- dalis, integrum, ovale, margine postico leviter convexo et utrinque seta una minima. — Longit. circ. 10 millim. — Ad Christianssund in Norvegia earn hand frequenteni invenimus.
2. Gen. EURYTENES oov. gen.
Vide supra!
3. Gen. ANONYX, KrSyer.
Pedes trunci (thoracici) hmi /"iris manu subrheliformi armati, ungue ticribili, margine inferiore manus jdas vel minus definite Mandibular tuber- culo molari mediocri vel magna. Laminae exteriores pedum maziUarium mar- aim' interiors /derumuue noduloso, raro di'uta/o vel aculeato.
We have on our coasts at least Often species of that genus, and of these two are as yet undescribed, one confounded with another species, and one as yet found only on the English coast and in the neighbourhood of the Shetland Isles. In order to facilitate the often difficult work of discri- minating the different species, we subjoin a synoptical table of these spe- cies, and call attention to such distinguishing marks as seem to 08 impor
On the Lysianassa magellanica etc. 23
tant. We further add the accompanying list with such descriptions and re- marks as the circumstances give rise to.
1. A. AMPULLA (Phipps); Kroyer. Anonyx ampulla, H. KROYER: Naturhist. Tidsskr. 2 Rsekke, 1 Bd. pag. 578.
„ „ R. Bruzelius: Skandinaviens Amphipoda Gammaridea, pag.
39; Wetensk. Akad:s Handl., ny foljd, Bd. 3 1859.
This species does not appear in Spence Bate's and J. 0. West- wood's "British Sessile-eyed Crustacea." The species there introduced under this name is a totally different one, as will presently be shown.
2. A. L0NG1PES, Spence Bate.
Anonyx longipes, C. Spence Bate : Catalogue of the specimens of Amphi-
podous Crustacea in the Collection of the British Museum,
pag. 79, pi. XIH, tig. 4. — Femina. „ „ C. Spence Bate & J. 0. Westwood: History of British
Sessile-Eyed Crustacea, T. I, pag. 113. — Femina. „ ampulla, C. Spence Bate: Catalogue etc., pag 79, pi. XIII, fig.
5. — Mas. „ „ C. Spence Bate & J. 0. Westwood: History etc. pag.
116. — Mas.
Descr. Longitudo cor]). 12 — 13 millim. Forma corporis gracilis ct elongata, epimeris mediocribus, capite parvo et ejus lohulis lateralibna Men- tis, segmento 3:tio caudse postice gibho, ej usque angulis lateralilms poste- rioribus (fig. 30, a) in aculemn longum et recurvum productis. Pedes truuci longi et graciles.
Antennae superiores feminae (fig. 23) segmentis 2:do et 3:tio pedun- culi brcvissimis , flagelli articulis circ. 15, et flagelli appendicularis arti- culis 5. Antennae mferiores superioribus fere longitudinc aequales. Antenna? superiores maris iisdem femiiue longiores, et ejus antennae inferiores flagello gracillimo et longissimo, Ut partem posteriorem caudae iuterdum asse- quantur.
Labrum (Fig. 24) parte superiore (a) in longum acumen durum, com- pressum et recurvum porrecta, et infra ad ejus basin lobulis duobus mol- libus (b, b) basin prope infra hispidis. Labium (c, c) ramis duobus ad api- cem setiferis confectum.
24 W. LlLLJEBORG,
Mandibularura (fig. 25) acies tantummodo uno denticulo. Processus aocessorius forma aculei magni et arcuati distinctus, et tubereulum molare magnum excavatum et hispidum aeuleis brevissiruis.
Maxillae l:mi paris (fig. 26) forma solita, ramo interiore et minore ad apicem duas sctas gerente, et ramo exteriore crasso apiceque circ. 7 — 8 dentibus magnis et serratis armato. Palpus ad apicem denticulis brevibua circ. 8. — Maxillae 2:di paris ramis ambo fere latitiuline ajqualibus.
Maxillipedes laminis interiuribus medium laminarum exteriorum asse- quentibus, et hse lamina} (enter.) ad marginem interiorem nodulis confertia prseditae, ad apicem vero aeuleis carentcs.
Pedes trunci l:mi paris (fig. 27) minus robusti, segmento 3:tio <li- midio segmenti 2:di multo breviore. Maims rectangularis, earpo fere ae- qualis, ungue parvo dente uno prope apicem iutus armato. — Pedes 2:di paris (fig. 28) gracillimi, segmento 3:tio dimidio segmenti 2:di fere a*quali. Manus oblongus, fere rectangularis, carpo parum brevior sed latior, et mar- gine posteriore convexo, ungue miniito, intus aeuleato.
Pedes trunci trium parium posteriorum longi et graciles ungue longo et vix arcuato. — Pedes caudales ultimi (fig. 29) ramis fere aequalibus, lan- ceolatis, et acideatis, et ad marginem interiorem setiferis.
Segmentum 7:mum caudai sive appendix eaudalis (fig. 31) supra utrinque aeuleis tribus, et fissura basin propius ineipiente marginibusque di- vergentibus. Laeinise ad apicem 1 — 3 aculeos gcrentcs.
Color flavido-albus. Oculi rubri.
Ad Haugesund, Molde et Cliristianssund in Norvegia banc sjieciem non infrequentem invenimus, et Doctor G. LiNDSiKii.M earn ad Farsund in eadem terra accepit; in profunditate 15 — 60 orgyiarum, et plemmque in fundo arenoso. Antea tantammodo in Anglia accepta.
3. A. GULOSUS, Kroyi:r.
Anonyx fjulosus, H. KuiiYER: Naturbist. Tiilskr. 2 K;ekkc, 1 lid, pag. (ill. „ norvegicus, LlLLJEBOfiO: Ofvcis. at' Kongl. Wetensk. Akatleiniens
F5rh. 1851, pag. 22. n „ K. BRDZELIUS: Skandinaviens Amphijioda (iaiinnaridea
pag. 44. „ HolboUi, Spencb Bate and Wbstwood: British Sessile-Eyed Cru- stacea, T. I, pag. 104
On the Lysianassa magellanica etc. 25
This species is easily recognized by the following' characteristics. The 1st pair of feet have the 3rd joint as long as or longer than half of the 2nd; the claw is armed at the inner edge with a strong tooth; the edge of the hand, to which the claw is attached is furnished with a row of ex- tremely fine spines arranged like the teeth of a comb, within which are scantily set bristles ; and last the 3 rd tail-segment has the lower back angle but little drawn out, but acute. The processus accessorius of the mandibula is particularly prominent, and on one side provided with teeth.
In very young individuals the 3rd joint of the 1st pair of feet is shor- ter than half of the second, and they closely approach the A. nanoides, from which however they may be distinguished by the abovementioned pair of feet beeing longer and slenderer, and by the fore and back borders of the hands being somewhat curved but not convergent, as in the latter spe- cies. The carpus of the 2 nd pair of feet is not quite double as long as the hand.
This species occurs on our west coast from Bohusliin to Finniar- ken, and is extended northward to Greenland and southward to England.
4. A. NANOIDKS, n. sp.
It is probably this species that R. Bruzelius (1. c.) has described un- der the name of Anonyx nanus Krdyer with a note of interrogation. In his description he has omitted the minute denticle on the inner side of the claw of the Is pah' of feet.
Descr. Minor; longitudo circ. 5 — 6 millim. Corporis forma sat obesa, tameii compressa, epimeris magnis, et pedibus brevibus, et unguibus pedum trunci posteriorum arcuatis. Segmentum o-.tium eaudae angulis posterioribus et inferioribus (fig. 34 a) recurvis sed minus acutis.
Antennae superiores inferioribus paullulum breviores, segmentis 2:do et 3:o pedunculi brevissimis. Flagellum articulis 10, quorum primus maximus, fere 3:tiae parti flagelli sequalis. Flagellum appendiculare articulo l:mo fla- gelli longius, circ. finem articuli 5:ti assequens, articulis 6. Antennae infe- riores articulis flagelli circ. 20. Maxillipedes laminis exterioribus ad medium circiter articuli 3:tii palpi porrectis, oblongo-ovatis, apice obtuse angulato, margine exteriore apicem propius eroso ibique seta minore, mar- ginoque interiore noduloso, nodulis minimis et paucis. Laminae interiores ad medium exterioruni extensae.
Nova Acta Reg. Sue. Sc. Ops., Ill* Sine. 4
26 W. LlLLJEBORG,
Pedes trunci l:mi paris (fig. 32) sat robusti, segmento 3:tio dimidio segmenti 2:di breviore, maim rectangulari, apicem versus parum angustiore. longitudine carpo circ. aequali. Unguis crassus et brevis, intus aculeo et setis.
Pedes trunci 2:di paris gracillimi, manu (fig. 33) ovali. longitudine dimidia carpi, et angulo inferiore et posteriore producto et aeuto.
Rami pedum caudalium ultimorum aculeati non vero setiferi, exterior interiore longior.
Oculi rubri.
Tantummodo specimen unum Junius ad Molde in Norvegia accepi.
5. A. PUMILUS, b. sp.
Descr. Minor; longitudo circ. 5 millim. Forma corporis minus obesa quam apud prsecedentem, epimeris et pedibus mediocribus, dorso vero la- tiusculo. Ungues pedum trunci posteriorum longi et parum arcuati. Seg- mentum 3:tium caudai angulis postcrioribus et inferioribus (fig. 41) productis. acuminatis et recurvis.
Antennae superiores (fig. 35) inferioribus parum breviores, segmentis 2:do et 3:tio pedunculi brevissimis. Flagellum articulis 7, quorum l:mus ceteris major est. Flagellum appendiculare articulis 3, ad medium flagelli porre- ctum. — Antennse inferiores articulis flagelli 8.
Mandibular (fig. 36) acie utrinque dente munita, proccssu aeeesaorio et ruberculo molari distinctis. — Pedes maxillarcs laminis extcrioribus ob- longo-ovatis non apicem articuli 2:di palpi attingentibus, ad apicem aculeos setiformes 5 gerentibus, margine interiore noduloso, nodulis minimis paucis (6 — 7) et discretis. Laminse interiorcs vix dimidiam lain, exteriorum attin- gentes, ad apicem dentibus 3 — 4, et setis 3.
Pedes trunci l:mi paris robusti, segmento 3:tio dimidio 2;di multo breviore, et manu (fig. 37) pyramidata vel triangnlari, et margine palms vix definite. Unguis magnns, intus denticnlo minimo et setis duabns vel trilius. — Pedes trunei 2:di paris parum graeiles, manu (fig. 38 ovali et carpo breviore, angulo posteriore et interiore fere recto, ungneque sat magno.
Pedes caudales ultimi (fig. 3'.)) ramis subulatis et setis carentibus, cxtcriore longiorc.
Segmentiini eamhe 7:mum (fig. 40) sat elongatnm, prot'nndc — ultra medium — fissuni, laciniis contiguis, ad apicem emarginatis. ibiqne acu- leuni gerentibus.
On the Lysianassa magellanica etc. 27
Color dilute ruber. — Oeuli obsoleti.
Tantummodo specimen unum ad Molde in Norvegia in profundo 40 — 50 orgyiarum et fundo argillaceo inveui.
6. A. BRACHYCERCUS, n. sp.
Descr. Minimus; longitudo 4 millim. Forma corporis valde crassa et brevis, epimeris magnis, et pedibus brevibus et robustis, unguibus arcuatis. Segmentum 3:rium caudae angulis posterioribus et inferioribus (fig. 49) pro- ductis, acutis et recurvis.
Antennae breves et crassa, supcriores et inferiores longitudine circ. aequales; supcriores (fig. 42) segmentis 2:do et 3:o pedunculi brevissi- mis, flagelli articulis 4, quorum l:mo maximo. Flagellum appendiculare biarticulatum, articulo l:mo magno, et 2:do miuimo; — inferiores flagelli ar- ticulis 4, quorum lrmus non multum ab articulo ultimo pedunculi diversus est.
Mandibular (fig. 43) acie utrinque dente obsoleto, processu accessorio distincto, et tuberculo molari elongato et compresso, et subtilissime hispido, nulli vcro aculci inter boc et processum accessorium adsunt.
Maxillae l:mi paris forma solita, ramo intcriore bisetoso. — Maxillae 2:di pans ramo interiore fere latiore sed breviore.
Maxillipcdes (fig. 44) laminis interioribus (a) elongatis, ad medium laminarum extcriorum porrcctis, ad apicem nodulis 3 et setis 3. Laminae ex- teriorcs magnae, ultra medium articuli 3:tii palpi extensaj, ad marginem in- teriorem tantummodo apicem propius nodulosae, nodulis discretis 4, et ad marginem exteriorem setam unam et pilos minimos gerentes.
Pedes trunci l:mi paris breves et robusti, segmento 3:tio dimidiae segmenti 2:di longitudini circ. aequali, et manu (fig. 45) carpo longiore, versus apicem inferiorem angustiore, marginibusque anteriore et posteriore arcuatis. Unguis magnus, intus dente uuo et setis tribus instructus. — Pedes trunci 2:di paris (fig. 4G) sat robusti, segmento 2:do sinuato, et 3:tio dimidia longitudine segmenti 2:di fere longiore. Maims carpo brevior, ob- longo-ovalis, angulo inferiore et posteriore acuto, ungue crasso et intus den- tato. — Pedes trunci parium trium posteriorum segmento 2:do sequentibus segmentis conjunctis multo longiore. — Pedes caudales ultimi (fig. 47) breves et crassi, ramo cxtcriore longiore et interiore apice minutissime bifido, et ramis ambo aculeis et setis carentibus.
28 W. LlLLJEBORG,
Segmentum ultimum caudse (fig. 48) late ovaturn, ad medium circ. fissum, laeiniis ad apicem aeuleum gerentibus.
Oeuli rubri.
Specimen unum ad Grip juxta Christianssund in Norvegia e profundo 20 — 30 orgyiarum cepi.
7. A. BRUZELII, A. Boeck.
Anonyx Biiizelii, Axel Boeck: Forliandlinger ved de Skandinaviske Natur- forskeres 8:de Mode, 1860 (tr. 1861) pag. 643.
Of this species we have no other knowledge than what is derived from Boeck's above-cited description, having never met with a specimen. It was found by the author referred to on the west coast of Norway. As regards the construction of the first pair of feet it somewhat approaches A. gulosus; nevertheless these feet are a little longer and slenderer than in that species, and it is not stated that the claw is furnished with any tooth on the inner side. The external lamina of the niaxillipeds lias only a few knobs on its inner border, whereas the A. gulosus has many.
8. A. NANUS, Kroyer.
Anonyx nanus, H. Kroyer: Naturhistorisk Tidsskr. 2 Bsekke, 2 Bd. pag. 30.
Distinguitvr: Anguli posteriores et inferiores segment! 3:tii caudse obtusi. Flagellum appendiculare antenn. super, triarticulatus, ad finem ar- ticuli 3:tii flagelli porrectum, articulo l:mo ceteris pintles longiore. Arti- culus loans flagelli reliquis eonjunctis longitudine requalis. Maxillipodum laminae intcriores ad marginem apicalem dentibus obtusis tribus et setis dua- bus. Laminai eorum exteriorcs ad marginem interiorem uodulis, sive den- tibus obtusis confertis. Pedes trunci l:mi paris breves et robusti, segmento 3:tio diniidia longitudine segm. 2:di breviore, manu carpo longitudine mquali, rectangulari, marginibns anteriore et posteriore arcuatis, et margine infe- riore truncate; uugue crasso et intus dente Sat valido ariuato.
Peduiii caudaliuni ultimorum ramus exterior ad marginem interiorem setis ciliatis longis circ. 10 instructus, et ramus interior apice bicuspide. Segmentum 7:nuim caiuhe ultra medium fissum, fissura aperta. Longit. circ. 5 m. m.
Oeuli rubri.
A Kroyer in Kattegat ad Bornbcek (parte australi), a Doctore G. Lindstkom ad Farsund in Norvegia aeceptus.
ON THE LYSIANASSA MAGELLANTCA ETC. 29
9. A. PINGUIS, A. Boeck.
Anonyx pinguis, Axel Boeck: Forhandl: ved de Skand. Naturfrs 8:de Mode, 1860 (tr. 1861), pag. 642.
This animal is also known to ns only by the description cited. It is found on the west coast of Norway. In the form of the 7* tail-seg- ment, which is divided only half-way, it approaches the above described A. brachycercus , as also probably in a thick and stout form of body, but is easily distinguished from it by having the lower back angles of the 3 rd cau- dal segment rounded off. The flagelhim of the upper antennae has 20 joints and the flagellum appendiculare 4, of which the first is long, with teeth on the under side. The first pair of feet have the hand longer than the car- pus, cut off at right angles at the lower end.
10. A. SERRATUS, A. Boeck.
Anonyx serratus, Axel BOECK: Forhandl. ved de Skand. Naturfis 8:de Mode, 1860, pag. 641. „ Edwardsii, C. SPENCE Bate : Catalogue etc. pag. 73, PI. XI, fig. 5. „ „ Spence Bate and J. O.Westwood: British Sessile-K v., 1
Crustacea, T. I, pag. 94.
Easily distinguished from all our other species of the genus Anonyx by the back border of the 31' caudalsegment's epimera being saw-toothed, (fig. 50) and by the lower angles of these epimera being rounded off. The back borders of the \>x and 2'"1 caudalsegments are also saw-toothed.
Forma corporis obesa, cpimeris niagnis, pedibus gracilibus non vero longis, unguibus arcuatis. Longitudo circ. 7 millim.
Antcnnse superiores inferioribus breviores, flagello 9, et flagello ap- pendicular! 5 articulis.
Mandibularum acies altero dente laterali plane obsoleto, et processu accessorio distincto et ad latus alteram dentato. — Maxillarum l:mi pans ramus interior elongatus, ultra medium rami exterioris porrectus, apiceque bisetoso, palpus ad apicem truncatum dentibus 6, et aculeo uno. — Maxilla 2:di paris ramo exteriore interiore longiore et latiore. — Maxillipedes s. pedes maxillares laminis inferioribus ad medium laminarum exteriorum por- rectis, et ad apicem subsinuatum tuberculis tribus et setis ciliatis tribus; laminis exterioribus ad finem articuli 2:di palpi extensis, et ad marginem interiorem confertim nodulosis.
30 W. LlLLJEBORG,
Pedes trimci l:mi paris breves et robusti, segmento 3:tio dimidia lon- gitudine segmenti 2:di multo breviore. Manus carpo longior, rectangularis , marginibus anteriorc et posteriore arcuatis, et margine inferiore sive palmaj erenulato. Unguis crassus et brevis intus denticulo minimo. — Pedes trunci 2:di paris forma solita, manu carpo breviore et angustiore, subsemilunari , hirsutissima, angulo posteriore et inferiore producto et acuto, ungue minimo intus dentato. — Pedes caudales ultimi ramis lanceolatis et aeuleatis non vero setiferis, exteriore longiore.
Segmentum caudje 7:mum usque ad sed non ultra medium fissum, fissura aperta. Latinise ad apicem aculeum unum, et in latere superiore aculeos tres gerentes.
Ad Grip juxta Cbristianssund in Norvegia specimiua duo e 20 — 30 orgyarura profuudo et fundo arenoso accepi.
11. A. EDWARDSII, Kroyer.
Anonyx Edwardsii, H. Kroyer: Naturhist. Tidsskr. 2 Rsekke, 2 Bd. pag. 1. „ „ R. BruzeliuS: Skandinaviens Ampliipoda Gammaridea,
pag. 46.
This species also we have never had an opportunity of examining, and accordingly know it only by the descriptions given by Kroyer and Bruzelius. According to the last named writer it is seldom met with on our western coasts from Bohuslan to Finmarken. It is distinguished by the form of the 7"' caudal-segment, which is short and broad, and has at the back border a very shallow but clearly visible notch ; it has also a pro- perty in common only with the following species, viz. that the claw of the first pair of feet has on the inner side a tooth, within which is a row of very fine saw-teeth.
12. A. L1TOKALIS, Kroyer.
Anonyx UtoraHs, H. KROYER: Naturhist. Tidsskr. 2 Rsekke, 1 B. pag. 621. R. Bruzelius : Skandinaviens Ampliipoda Gammaridea,
pag. 46.
This species, like the proceeding, is distinguished by the form of the last caudal-segment, which is onforked, and in the back border has only a slight indication of inward curvature. It also has a tooth on the inner side of the claw of Is pair of feet, and inside that a row of fine saw-teeth, hut both these last and the tooth itself are larger than in the proceeding species.
On the Lysianassa magellanica etc. 31
On our coasts it is confined to the extreme north. M:r Doct. Th. Fries has brought home & presented to the University Zoological Museum a specimen taken by him on the coast of Warangarfjord.
Longit. 14 — 15 millim. Segmentum 3:tium caudae angulis inferiori- bus et posterioribus acutis, non vero productis. Mandibular acie subtilissime crenulata, utrinque dente sat valido, processu accessorio distincto, solito raodo stiliformi, tuberculo molari medioeri denticulato et hispido. — Maxil- lae l:mi paris ramo interiore parvo, rotundato. setis duabus, et palpo ad apicem aculeis quinque. — Maxillae 2:di paris ramo exteriore interiore fere duplo majore. — Maxillipedes laminis interioribus brevibus, ad apicem tu- bcrculis tribus et setis duabus vel tribus; laminis cxterioribus etiam brevibus, non apicem articuli 2:di palpi assequentibus, margine interiore noduloso, nodulis discrctis (circ. 10), et ad margineni extcriorem prope apicem aculeo mobili. — Pedes trunci l:mi paris brcvissimi et robustissinii, maim fere quadrangulari, infra oblique truncata. Oculi rubri, o\ ato-reniformes.
13. A. HOLBOLLII, Kroyer.
Anonyx Holbollii, H. Kroyer: Naturhist. Tidsskr. 2 Radcke, 2 Bd. pag. 8. „ „ K. Bruzelius: Skandinaviens Ampliipuda Oaminaridea,
pag. 43. ,, denticulatus , C. Spence Bate: Catalogue etc. pag. 74; pi. 12, fig.
2. - Mas. „ „ C. Spence Bate and Westwood: British Sessile-Eyed
Crustacea, T. I, pag. 101. — Mas.
This species is distinguished from all our other species by having the maxillipeds' outer laminae, which are crescent-shaped, furnished on the inner margin with thickly set, comb-like, long, sharp teeth, 17 to 20 in number, and by the baud of the 1st pah' of feet, which is oblong-oval, and has the lower back angle obliquely rounded off, and is furnished with a smooth and peculiarly long claw, which when pressed in reaches with its point back to the middle of the hand. It might therefore be veury well considered as the type of a separate genus. It is also remarkable for the 3 rd caudal- segment's lower and back angles, which are carried out into a very, long sharp point bent upwards, and for the mandibulae which have the cutting or eggbearing end very small and the tuberculum molare very large hard and prominent. The 1 st pair of maxillae have no side-teeth on the large spines at the end of the outer brauch, and their palp has a great number of den- ticles at the point. The second pair of maxillae have the branches very
32 W. LlLLJEBORG,
broad. Tlie flagellum appendiculare of the upper antennae has 3 joints, with the Is joint longer than the other two pat together, and the point al- ways turned upward. The lower antennae are in both sexes, but especially in the male, considerably longer than the upper. The claw of the back truncal feet is long and but slightly curved. We have found the body's length 14 millini.
It is met with on our western coasts form Bohuslan to Finmarken, an in one or two places, as for example Haugesund and Molde in Norway we have found it not uncommon, where the bottom is sandy, and at a depth of 12 to 50 fathoms.
14. A. OBTUSIFRONS, A. Boeck.
Anonyx Obtusifrons, Axel BOECK: Forhandl. ved de Skand. Naturf:s 8:de
Mode, 1860, pag. 643.
This species is known to me only by Boeck's description. The flagellum of the upper autennae has 8 joints, of which the first is as long as all the rest together. The first joint of the flagellum appendiculare is particularly long and provided with several spines on the lower side. The exterior laminae of the maxillipcds have some few scattered coarse teeth on the inner margin. The 3nl caudal segment has its lower and back an- gles drawn out into a long hook or point bent upwards. &c. It is taken on the western coast of Norway.
15. A. TUMIDUS, Kroyer.
Anonyx Uimidus, H. Kroyer: Naturhist. Tidskr. 2 Raikke, 2 Bd. pag. 16. „ „ R. Bruzelius: Skandinaviens Amphipoda Gammaridea,
pag. 41.
This species is by the nature of the mouth's appendages, by the form of the hand belonging to the first pair of feet, and by the peculiarity of its habits, so distinguished from every other species of the same genus, that it might very well be considered as the type of a distinct genus. The mamlibulae are remarkably small, have the egg-bearing end very small and both side teeth exceedingly small, and are without processus aceessorius. The tuberculum molare is very large but thin, brought up to a point and destitute of bristles. The 1 pair of maxillae are short and have both branches, especially the outer, very broad. The inner branch has at its termination 5 coarse ciliated bristles, and the outer has a great many spi- nes. The palp has the usual form. The 2nd pair of maxillae, which are
ON THE Ly.sIAXASSA MAGELLANICA ETC. 33
also short, have the inner branch very broad, and the outer much narrower. The maxillipeds (fig. 51) are also short, but have the laminae exteriores very large, and reaching about to the end of the 2 nd joint of the palp. The inner lamina? are wery short, armed with coarse ciliated bristles at the end and inner side. The exterior laminae have the internal margin almost smooth , but immediately within the edge is a row of about 7 short spines , directed for- wards, the points of which reach beyond the edge of the laminae, by which this latter shows itself to be thinly set with spines. At the point they have a long spine and beyond that three bristles. The hands of the first pair of feet approach in form those of the genus Lysianassa, tapering towards the point, almost conical, and the palm-edge not clearly defined. But the claw is moveable and in some measure doubled back upon the posterior edge of the hand. The whole back edge of the hand is furnished with small spi- nes , and in front of these are a few (5) of larger dimensions. The claw is not, as Kroyer states, divided at the point, but has however on the inner side an almost invisible prickle. The branches of the last pair of caudal-feet are distinguished by having, like those of the genus Hyperia, fine sawteeth, the outer branch only on the inner margin, but the inner branch on both margins.
It is found in the branchial-sack of Ascidice on our western coast from the southern part of the Kattegat at least up to Christianssund in Norway.
\. Gen. CALLISOMA, A. Costa.
Cum genere Anonycis maxima ex parte congruens, distinguitar tamen: Pedes trunci l:mi pari* iisdem 2:di parvs nen crassiores, iiiterdnm grariliores, sed longiorcs, ungue obsoleto vel absente. Ramus interior ma.rillarum l:mi paris setis ciliatis sat multis praeditxs. — Laminae exteriores ma«iUipedum margine interiore aculeato. Aculei ad apicem palpi maxittarwm l:mi paris bifurcati.
Of this genus 4 species arc known, two froni Naples (Callis. Hopei and pnnctatum A. Costa), 1 from England (Callis. crenata. S. Bate), and one from Sweden and Norway.
1. C. KROYERI (Bruzelius).
Anonym Krdyeri, R. Bruzelius: Skandinaviens Amphipoda Gammaridea, pag. 45, tab. II, fig. 7.
Callisoma „ C. Spence Bate : Catalogue etc. pag. 371.
Nova Acta Reg. Sue. Sc. Dps. . IIIe Sc;rk'. 5
34 W. LlLLJEBORG,
The 1st pair of truncal-feet are long and slender, longer than the 2" , and their hands have a very small and rudimentary (daw, concealed by long bristles, on the baek border above the lower and back angle. The hand of the 2ml pair of truneal feet has the lower and baek corner drawn out so as to be almost cheliform. The upper antennae are shorter than the lower; the 3,a caudal-segment's lower and back corners are rounded off; and the 7 th caudal-segment deeply cloven, &c.
According to Bruzelius this species is to be met with on our we- stern coasts from Bohuslan to Finmarken.
5. Gen. AC1DOSTOMA. n. gen. *)
Forma corporis et antennarnm rum genere Anonycis congruit, oiis partes appendiculares tamen plane diversae. Labii rami laterales angusti. Mandibulae processu accessorio, maxillae (rmi paris palpo, et palpus maanlli- pedum ungue carentes, et hae partes oris conjunction acumen production prae- bent. Pedes trunci l:mi parts rolatsti, manu prehensUL Pedes 2:di parts gra- ciles, ungue carentes.
Of this genus we know of only one species, namely:
1. ACIDOSTOMA OBEgXJM (Spence Bate).
Anonyx obesus, C. Spence Bate: Catalogue etc. pag. 74.
„ „ C. Spence Bate & J. 0. Westwood: British Sissilc-Kycd
Crustacea, T. I, p. lJ8.
Descr. Corporis forma (fig. 53) obesa, dorso latiusculo et rotundato. epimeris magnis, et pedibus brevibus et robustis, unguibus parum arcuatis, angulis posterioribus et inferioribus scgmeiiti 3:tii eaudai obtusis. Longitudo circ. 5 millim. — Color ruber; oculi rotundati, fusci
Caput parvuni, lobi laterales parum producti. Antenna siiperiores (tig. 54) peduucoli segmentis 2:do et 3:tio brevibus. flagello articulis circ. 7 quorum primo brevissimo; flagelloqne appeudiculari longitudine flagello fere sequali, articulis 5. — Antenna inferiores longitudine superioribua circ. eeqnales, flagelli articulis 7.
Labium (fig. f>5, a) vonicrit'ornu', et labii partes laterales she rami angusti (b,b), ad apiceni aeuininati et intus hispidi. .Mandibuhe (fig. 56) elon-
') From ilxi; a point and (fiti/tr month, because the mouth and its appendages form a long projecting point.
On the Lysianassa magellanica etc. 25
gatse, processu accessorio carentes, acie utrinque denticulo niinutissimo. Tu- berculum molare (a) non po.stice definitum. Palpus longus et gracilis, segraento 2:do ceteris longiore, et segmento 3:tio ad apicem setifero. — Maxillae l:mi pane (fig. 57) palpo cardites, angustse, ramo exteriore (a) et ma- jore ad apicem dentibus curvatis et partim serratis circ. 8 armato, ramo in- teriore minore ad apicem setas paucas gerente. — Maxilla? 2:di paris (fig. 58) minores et adhuc angustiores, ramo exteriore interiore longiore, et anibo ad apicem setiferis. — Maxillipedes (fig. 59) laminis interioribus mcdiocribus, ad apicem setis 3 — 4, et laminis exterioribus maximis, fere semilunaribus, marginibusque lsevibus, interioribus tantuni apicem versus nodulis minutis- simis et obsoletis paucis instructis. Palpus ungue carens. laminae exteriori longitudine circ. ajqualis, articulis elongatis, sed corum 4:to brevissimo.
Pedes trunci l:mi paris (fig. 60) breves et robusti, segmento 3:tio dimidia longitudine segmenti 2:di breviore. Manus caipo circ. a-qualis, co- nica, margine anteriore areuato et posteriore recto et minutissiine hupido, prajtereaque setis majoribus. Unguis flexibilis, arcuatus et hvvis. — Pedes trunci 2:di paris (fig. Gl) iisdem l:mi paris graeiliores et longiores, manu valdc bispida, oblonga, fere sequilata, parum arcnata et carpo breviore et angustiore, apice obtuso et ungue carente. — Pedes 3:tii et 4:ti paris (fig. 62, pes 3:tii paris) ungue artieulo unguit'ero paullubini breviore. — Pe- des trunci trium pariuin posteriorum inter se fere eadem forma, breves et lati (fig. 63, pes 7:mi paris), segments 2:d<> maximo et se(|uentibus eon- junctis longitudine fere aiquali, postice infra serrato et antiee aeiileato; unguis validus et modicc arcuatus. Pedes 5:ti et 6:ti paris segmento 2:do eodem pedum 7:mi paris paullulum breviore. — Vesicula; branchiales aimplices.
Pedes caudales ultimi (fig. Ii4) ramis lanceolatis, exteriore angustiore et longiore, apiceque tuberculo minimo.
Segmentum 7:mum candse (fig. G5) latum, fere rotundatum, ultra me- dium fissum, laciniis apice obtuso et aculeo brevissimo instructo.
Ad Molde in Norvegia e profundo 30 — 40 orgyiarum et fundo argil- laceo banc speciem hand fi-equentem invenimus. Doctor Lixdstkom earn ad Farsund in Norvegia et in Bahusia accepit.
36 W. LlLLJEBORG,
» »
EXPLICATIO FIGURARUM.
Tabula I.
Eurytenes magellanicus.
Fig. 1. Animal, magnitudine naturali. b', segmentum l:mum pedunculi antennae inferioris.
„ 2. Caput cum antennis, a latere dextro visum, b, segmentum l:mum pedun- culi antennaa inferioris.
„ 3. Caput, visum a latere inferiore. «{, «, antennae superiores. b, b, antenna' iuferiores. e, c, labrum. d, </, mandibular d', d' , palpi mandibularum. g, g, segmenta l:ma truncorum pedum maxillarium sive maxillipedum. ;/,</', laminae extcriores maxillipedum, et g", g" eorum palpi.
„ 4. Labrum, «, et labium, b, a latere sinistro visa.
5. a, labrum. b, b, rami ainbo labii. be. Labium (rami ambo) a latere superiors visum.
6. Mandibula sinistra, a latere interiore visa, a, acies. b, tubereulum mo- lare. c, processus lateralis, d, palpus.
7. Mandibula eadem, a latere exteriore visa, eodem mode- signata.
8. Maxilla l:mi paris. a & a' ramus exterior sive truueus. b, ramus interior. <.-, palpus.
„ '.). Maxilla 2:di paris. a, ramus exterior, b, ramus interior.
Tabula II.
Eutytenes magellanicus.
Fig. 10. Pedes maxillarcs sive maxillipedes. a, a, segmenta l.-ma truneorum. b, b, segmenta 2:da eorum. //, lamina exteriores. c, c, palpi. Lamina? intoriores maxillipedum.
Pan Laminae exterioris eorum cum margine interiore.
Pes dexter l:mi paris pad. trunci sive thoracicorom. a & 1, segmentum lntuim. 2 — G Begmenta -alum — 6:tom. 14. res dexter trum-i 2:di pans. a& 1, segmentum Lnium, b, lamina matri- calis sive capsularis. c, vesicula brancliialis.
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On the Lysianassa magellanica etc. 37
Fig. 15. Pes dexter trunci 3:tii paris.
„ 16. Pes dexter trunci 6:ti paris. 1 &2, segmenta l:mum & 2:dum.
„ 17. Vesicula branchialis pedis trunci dextri 6:ti paris. a, truncus.
„ 18. Pes caudalis dexter l:mi paris. b, b, rami.
Tabula III.
Fig. 19 — 22. Eurytenes magellanicus.
Pes caudalis dexter 4:ti paris. a, ramus exterior, b, ramus interior. Pes caudalis dexter 6:te paris. a, ramus exterior, b, ramus interior. Segmentum 7:mum caudae sive appendix caudalis. Epimerum dextrum segmenti 3:tii caudae.
Fig. 23 — 31. Anonyx longipes.
Antenna superior.
a&b, l>, labrum; c, c, labium, a latere dextro visa.
Mandibula.
Maxilla l:mi paris.
Segmenta ultima pedis trunci l:mi paris.
Segmenta ultima pedis trunci 2:di paris.
Pes caudalis 6:ti paris sinister, a & b, rami exterior et interior.
Epimerum segmenti 3:tii caudae.
Segmentum 7:mum cauche.
Fig. 32 — 34. Anonyx nanoides.
Pes trunci l:mi paris. Pes trunci 2:di paris. Epimerum segmenti 3:tii caudae.
Tabula IV.
Fig. 35 — 41. Anonyx pumilus
Fig. 35. Antenna superior.
36. Mandibula.
37. Segmenta ultima pedis trunci l:mi paris.
38. Segmenta idtima pedis trunci 2:di paris.
39. Pes caudalis G:ti paris. a & b, rami exterior & interior.
40. Segmentum 7:mum caudae.
41. Epimerum segmenti 3:ti caudae.
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Fig. 42 — 49. Anonyx brachycercus.
Fig. 42. Antenna superior.
„ 43. Mandibula.
„ 44. Pes maxillaris. a, lamina interior, b, lamina exterior.
„ 45. Segmenta ultima pedis trunci lani paris.
Nova Acta Reg. Soc. Sc. Ups., III? Serie.
38
Fig. 4G. Pes trunci 2:di paris.
„ 47. Pes caudalis 6:ti paris. a, ramus exterior.
„ 48. Segmentum 7:mum caudse.
„ 49. Epiinerum segmenti 3:tii caudse.
Fig. 50. Anonyx serratus. Fig. 50. Epimerum segmenti 3:tii caudae.
Fig. 51. Anonyx tumidus. Fig. 51. Pes maxillaris. a, lamina interior. b, lamina exterior.
Fig. 52. Anonyx ampulla. Fig. 52. Mandibula. a, processus accessorius. b, tuberculum molare. c, palpus.
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Tabula V.
Acidostoma obesnm.
Animal a latere dextro visum.
Antenna superior.
a, labrum. b, b, rami labii.
Mandibula. «, tuberculum molare.
Maxilla l:mi paris. a & b, rami exterior & interior.
Maxilla 2:di paris. a & b, rami exterior & interior.
Maxillipedes.
Pes trunci l:mi paris.
Pes trunci 2:di paris. a, lamina matricalis.
Pes trunci 3:tii paris.
Pes trunci 7:mi paris.
Pes caudalis 6:ti paris. a, ramus exterior.
Segmentum 7:mum Cauda;.
A't'i'it Acta. Sodei Sen/*/ tfjtxal Si r .*> "-
ri i
Eurvtenes ma^ellanicus
\'<itff -irftt Sociri Sciewtt UpsaX Ser 3 ~
/'/.//
■ ■
Kim inics maeellanicus
Wova ./(in SodttSdcmt UptaL Ser3-
I 'I III
■
Wt. 1 9 -22.Eurvten.es mafcellamcus. Fi*>2.V")l. Anonvx lonOipcs. Y\'i .VJ .Vi.Auonvx nanoxd'e s.
Nova .Uia Societ Sclent Upetd Sar 3i
PI II'
Fife.55-41 .\iion\.\ pumilus. Elg.42-49.An.onyx brachyceTeus, Yvi 50. Anonvx serratus. Fi?. 51. Anonvx tumidus. Fifc.52 Anonvx ampulla.
NbvaActa SocictSeicni UpsalSa .'!'
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Acidostoma obesum
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