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UNIVERSITY OF LONDON

FRANCIS GALTON LABORATORY FOR NATIONAL EUGENICS

EUGENICS LABORATORY MEMOIRS. IV.

ON THE MEASURE OF THE RESEMBLANCE OF FIRST COUSINS

BY ETHEL M. ELDERTON,

GALTON RESEARCH SCHOLAR IN NATIONAL EUGENICS, UNIVERSITY OF LONDON

ASSISTED BY

KARL PEARSON, F.R.S.

LONDON :

PUBLISHED BY DULAU AND CO., 37, SOHO SQUARE, W. 1907

Price Tliree Shillings and Sixpence

THE FRANCIS GALTON EUGENICS LABORATORY.

University of London, University College, Gower Street, W.C. The Laboratory is under the supervision of Professor Karl Pearson, F.R.S., in consultation with Mr Francis Galton, F.R.S.

Francis Galton Fellow in National Eugenics : David Heron, M.A.

Francis Galton Scholar : Miss E. M. Elderton.

Computer : Miss Amy Barrington.

Advisory Committee. The following have kindly consented to aid the Staff of the Laboratory in special forms of enquiry :

W. Palin Elderton. E. Nettleship.

J. Macpherson, M.D. Edgar Schuster, M.A.

F. Mott, M.D., F.R.S. Lieut. -Colonel R. J. Simpson, R.A.M.C.

J. F. Tocher.

National Eugenics is the study of agencies under social control that may

improve or impair the racial qualities of future generations,

either physically or mentally.

It is the intention of the Founder, Mr Francis Galton, that the Laboratory shall act (i) as a storehouse for statistical material bearing on the mental and physical conditions in man and the relation of these conditions to inheritance and environment, (ii) as a centre for the publication or other form of distribution of information concerning National Eugenics. Provision is made in association with the Biometric Laboratory at University College for training in Statistical Method and for assisting research workers in special Eugenics Problems.

Short courses' of instruction will be provided for those engaged in social, anthropometric; or medical work and desirous of applying- modern methods of analysis to the reduction of their observations.

UNIVERSITY OF LONDON FRANCIS GALTON LABORATORY FOR NATIONAL EUGENICS

EUGENICS LABORATORY MEMOIRS. IV.

ON THE MEASURE OF THE RESEMBLANCE OF FIRST COUSINS

BY ETHEL M. ELDERTON,

GALTON RESEARCH SCHOLAR IN NATIONAL EUGENICS, UNIVERSITY OF LONDON

ASSISTED BY

KARL PEARSON, F.R.S.

LONDON :

PUBLISHED BY DULAU AND CO., 37, SOHO SQUARE, W.

1907

^v*V^

70709 Some reconstruction of the Francis Galton Laboratory having taken place, it

seemed desirable to provide the workers associated with it with a direct channel

of publication of their own, in which their more extended memoirs should appear.

It is hoped that the present sei'ies may be issued at short intervals. Subscribers

should notify their intention of taking in the memoirs as they are published

to Messrs Dulau & Co. Requests to exchange with similar publications, with

archives and journals dealing with demographic and sociological problems, or

with census reports, should be directed to The Editor, Eugenics Laboratory,

University College, Gower Street, London, W.C.

On a Measure of the Resemblance of First Cousins in Man.

J

By Ethel M. Eldertox, Galton Research Scholar in National Eugenics in the University of London ; assisted by Karl Pearson, F.R.S.

1. Introductory. While a very large amount of data has been collected, reduced and published relating to the degree of resemblance in physical and psychical characters of a considerable number of pairs of relatives in man especially in the direct line and between collaterals of the first degree but little has yet been done with regard to collaterals in higher degrees. As far as we are aware tbe only quanti- tative measures yet determined are those for eye colour in man between uncle or aunt and nephew or niece*. No measure of resemblance has yet been determined for cousins. Yet it is precisely among collaterals of the second degree that the question of consanguineous marriages becomes in practical life of great importance, inequality of age being bere less marked, and thus the degree of resemblance between such collaterals has not only scientific but eugenic value. According to local law and religious custom cousin marriages are permitted or forbidden ; thus it would appear that we are here concerned with divergent human experiences, unconsciously formu- lated, as to the relative value of endogamy and exogamy. If we take a character whicb is detrimental to tbe individual, it will, at least in primitive communities, be in the bulk of cases a hindrance to mating. Hence, as a rule, we must classify such a detrimental character as recessive in the Mendelian senset, otherwise selection would have weeded it out. Now consider for a moment a population of dominants with notation DD, and suppose one of these to mate with an individual of detrimental attribute and constitution RR. The result will be the hybrid sibship marked by DR, in which the recessive character R will be latent. If brother-sister mating is for- bidden the next generation will be obtained (assuming the recessive individuals RR to be extremely rare) by mating with the population of dominant character, and the result will be equal numbers of DD and DR. Thus the generation of cousins would consist of 50 p. c. of dominants and 50 p. c. apparent dominants with the detrimental character recessive. ' It therefore follows that it would be as detrimental for some cousins to marry as for all brothers and sisters of the first hybrid sibship. That is to

* Pearson and Lee : Phil. Trans. Vol. 195, A, p. 114 et seq.

t Thus albinism and the tuberculous diathesis are, if not true and complete recessives in the Mendelian sense, still more nearly recessive than dominant characteristics. But there are other abnormalities, e.g. certain digital deformations, which are nearer to, if perhaps not true, dominants.

1—2

4 ETHEL M. ELDERTON

say while a brother and sister marriage would lead to 25 p. c. of the offspring having the harmful character patent and another 50 p. c. having it latent, the intermarriage of the cousins of the DR class among themselves would lead to the same baneful results as this brother-sister marriage, while the intermarriage of the DR class of cousin with the DD would also lead to 50 p. c. with the latent detrimental character. In other words endogamy as far as brothers and sisters are concerned would lead to :

25 p. c. hale. 50 p. c. latent evil. 25 p. c. patent evil,

while endogamy in the cousinship would give us:

56"25 p. c. hale. 37"5 p. c. latent evil. 6-25 p. c. patent evil.

The explanation therefore of the wide-spread social feeling against endogamy in the first degree, even between apparently hale individuals, is on the surface of it explicable on the Mendelian theory ; also we see that, whether we look upon cousin marriage as producing on the average more than six per cent, of patent evil, or in the other aspect, that some cousin marriages are as detrimental as brother-sister marriages, reasons can be found for their all being forbidden by tribal custom or religious ordinance. But this is after all only to look on one side of the picture, because the RR characteristic might be a patent good quality suddenly introduced from outside into a population ; in such a case cousin marriage is distinctly to be commended, and brother-sister marriage would be more effectual still. In this way the endogamy of many early communities receives its due sanction. As long as a species is likely to vary advantageously, endogamy between collaterals of the first degree will produce 75 p. c. with patent or latent good quality, and between collaterals of the second degree 62 5 p. c* ; even endogamy of ascendants and descendants may be advantageous. It is probable that whenever selection is extremely stringent the relative advantages of endogamy become apparent and are emphasised by tribal custom. But the Mendelian theory cannot be considered as demonstrated, and if it were, we could hardly at present apply it to man. We have no means of separating the DR's from the DD's, short of that experimental breeding which the Mendelians tell us is the only reliable guide to the gametic constitution. We cannot, however, afford to bring defective children into the world to test where the endogamous union will be an advantage, where a failure. The somatic characters of the individual and of his or her ancestry are at present our sole possible guide to his or her gametic constitution. From this standpoint we may ask what is the quantitative value of the cousin in the problem of inheritance ? In predicting the probable offspring of an individual is the cousinship of more or less importance than the parents' brothers and sisters ? Is a knowledge of the grandparents' characters of greater value than that of the cousin ? It will be clear that the cousinship while generally less accessible than the sibship, is often far more accessible than the grandparentage, or in the case of orphans than even the parentage ; and for the special purpose of medical diagnosis

* This supposes that endogamy in the first degree is forbidden.

MEASURE OF RESEMBLANCE OF FIRST COUSINS 5

may be of great relevance. The existing state of doubt as to the quantitative value of cousinship may be illustrated from such a vital problem as that of the hereditary predisposition to mental disease where some medical authorities would exclude entirely evidence drawn from the cousinship*, while they retain the inquiry as to the direct line and as to collaterals in the first degree. As the cousinship often combines many lines, it requires of course careful handling, but its size and relative accessibility may be factors which give it equal importance with the grandparentage or the parental sibships.

2. With the object of throwing light on the value of the record of cousinship, an inquiry as to the physical and psychological resemblance of first cousins was set on foot by Karl Pearson some five years ago and a grant obtained from the Government Grant Committee to assist the investigation. The assistance derived from this source is here gratefully acknowledged. The plan followed was twofold. Two independent collections were started. The first part of the investigation was based upon very general inquiries as to the physical and psychical characteristics of families. At present about 300 families have supplied very full particulars of ancestors and collaterals as far as the personal knowledge of the recorders extend. These Family Records supply the material upon which the bulk of the present paper is based, and provide sufficient pairs of cousins to give a fair idea of the general intensity of re- semblance in cousins. The family schedules asked for the following information:

(1) Present Age or Age at Death of each individual.

(2) Ailments in Life.

(3) Cause of Death, if dead.

(4) General Health under the Categories : Very Robust, Robust, Normally Healthy, Delicate, and Very Delicate.

(5) Ability under the categories :

A. Mentally Defective. Capable of holding in the mind only the simplest

facts, and incapable of perceiving or reasoning about the relationship

between facts. B. Slow Dull. Capable of perceiving relationship between facts in some

few fields with long and continuous effort ; but not generally or without

much assistance. C. Slow. Very slow in thought generally, but with time understanding is

reached. D. Slow Intelligent. Slow generally, although possibly more rapid in

certain fields ; quite sure of knowledge when once acquired. Ex. Fairly Intelligent. Ready to grasp, and capable of perceiving facts in

most fields ; capable of understanding without much effort.

* Bucknill and Tuke : Psychological Medicine, 2nd edn. p. 266.

6 ETHEL M. ELDERTON

E... Distinctly Capable. A mind quick in perception and in reasoning

rightly about the perceived. F. Very Able. Quite exceptionally able intellectually, as evidenced either

by the person's career or by consensus of opinion of acquaintances. During a part of the investigation E, and E2 were classed together as E, but

a large number of D, E and E, F entries (i.e. 'Betwixt' entries) occurring,

this category of E was divided as above into Ej and E.,.

(6) Temper under the categories : Sullen Temper, Quick Temper, Even Temper, Weak Temper (not ' even,' but weak good nature).

(7) Temperament under three divisions (a) Reserved, Expressive or Betwixt ; (b) Sympathetic, Callous or Betwixt ; (c) Excitable, Calm or Betwixt.

(8) Success in Life under the categories : Marked Success : An individual who is not only marked above his family, but above his fellow citizens for achievement in life. One who has made a name which would find a place in the Dictionary of National Biography. Prosperous Career: An individual who has advanced beyond his family level but not necessarily marked among his fellow men. An active successful life or career. Average Career: An individual who has not fallen below the family standard of life, whether in profession, trade or craft. Difficult Career : An individual who has found it difficult to maintain the previous family standard. One who has had a struggling and unprosperous career. Failure : An individual who has more or less failed in life ; a bankrupt, or ne'er-do-well ; this letter (F.) may be used to cover the black sheep of a family.

Considerable care was taken in distributing the schedules* among those likely to be interested in the investigation and having a sense of responsibility for the frank- ness and fullness of the information provided. A considerable number of schedules were returned to the recorders for corrections or additions which were at once supplied. In less than two per cent, of cases was it needful to reject a schedule as untrustworthy, or so incomplete as to be useless. Each Family Record contains on the average particulars of about 40 individuals and, as it is hoped to raise the total number of records from the present 300 to 1000, we shall then possess an account of a fairly random sample of the general population of about 40,000 persons.

3. The various types of cousinship distinguished in the schedules are : (^4) Cousins are sons of two brothers. (B) Cousins are sons of two sisters. (C) Cousins are sons of a brother and a sister. (D) Cousins are daughters of two brothers. (E) Cousins are daughters of two sisters. (F) Cousins are daughters of a brother and sister. (G) Cousins are son and daughter of two brothers. (H) Cousins are son and daughter of two sisters. (/) One cousin is daughter of a brother, the other is son of a sister. (K) One cousin is daughter of a sister, the other is son of a brother.

* They provided for information with regard to four generations in the direct line, and three generations of collaterals.

MEASURE OF RESEMBLANCE OF FIRST CO.USINS 7

A, B and C are types of male, D, E and F are types of female, G, H, I, A' of male and female cousins. There are thus ten types of simple first cousins. It was con- sidered desirable to keep these ten types distinct in order to ascertain how far resemblance was modified by change of sex in descent from a common ancestor. Special cases of abnormal cousinship in the first degree were not included. All individuals dealt with were adults. The four characteristics : General Health, Ability, Temper and Success in Life, providing 5, 7, 4 and 5 categories, admitted at once of tables of contingency being formed of at least 4x4 groups, and these were at once reduced by the method of mean square contingency. This was done for all the ten types of cousinship. In the case of Temperament there were only the alternatives and the ' Betwixt ' groups. We were thus compelled to use either con- tingency on a 3 x 3-fold grouping or else assume the material to have a Gaussian distribution and apply fourfold table divisions. In the latter case the results will vary somewhat according to the alternative with which the ' Betwixt ' group is associated.

The Temperament results are, however, in our opinion the least reliable of the series. We believe this to be due to the fact that the ability, success, health and to some extent the temper of an individual are matters of common knowledge or repute ; but that temperament as we have classified it is less generally realised. There is little doubt that some of the recorders had not previously formed a general estimate of temperament, and have taken that of a particular individual as a standard to classify other members of the family by. We should not be inclined accordingly to place much stress on the Temperament results as proving anything beyond the basal principle that temperament is an inherited charactei\

In the second series of investigations, it was considered that it would not be without interest to deal with an entirely novel physical character, i.e. novel from the standpoint of inheritance, and accordingly the hand was selected as easily accessible and, at any rate for some characters*, capable of fairly accurate measurement. Other physical characters readily ascertainable were eye and hair colours and general health. Accordingly a cousin schedule was issued with the directions for measurement noted below. See Appendix A, p. 21. Much time and energy had already been spent over an endeavour to reproduce in a cheap manner the eye and hair scales used as standards in the Biometric Laboratory. Ultimately we had to content ourselves with an admittedly imperfect chromolithograph of hair coloursf. For the eye scale we used a hand-painted scale. Miss Mary Beeton kindly painted on a printed blank the irides of 24 eyes, painting one eye at a time in about 100 copies from a standard glass eye. To these scales was added a cheap but quite efficient hand-spanner prepared by the Cambridge Scientific Instrument Company. The scales, spanners, directions and schedules were circulated in the same manner as the similar material

* R. S. Proc. Vol. 65, pp. 126—151 : "Data for the Problem of Evolution in Man. A First Study of the Human Hand." By M. A. Whiteley and Karl Pearson, t Reproduced Biometrika, Vol. v. p. 474.

8 ETHEL M. ELDEKTON

for the Family Measurements of six years ago* : they were loaned to College students, personal friends of members of the Biometric Laboratory and others. It was, however, soon obvious that we had miscalculated the ease with which pairs of cousins could be found and measured. The work went forward extremely slowly, most investigators sent in only two or three pairs ; and when the question of repeating or verifying a measurement arose, the delay or even the impossibility of supplementing the data was much more common than in the case of the Family Records. In fact cousins are not like brothers and sisters, or pai'ents and offspring, in daily touch with each other; and at the end of three or four years, we are far from having reached a sufficient supply of cousin pairs. Hardly indeed have 300 pairs been yet measured. Accord- ingly this side of the enquiry is incomplete and will only be used as a control series. We need scarcely say that we shall be very glad indeed to loan spanner and scales to any reader of this memoir who will undertake to measure pairs of adult cousins.

4. We now turn to the analytical methods by which the material was reduced. We have already pointed out that contingency was used throughout the whole of the first series, but that in the case of Temperament the 3 x 3-fold tables were not finely enough grouped to make contingency thus obtained really comparable with that found from higher-fold tables. Accordingly the temperament tables were only worked out by 3 x 3-fold contingency in the case of the three groups : male cousin pairs, female cousin pairs, and male and female cousin pairs. The fourfold table methodt was used on the same material in thirty cases, namely the three classes of temperament in the ten classes of cousins.

The mean value of the degree of resemblance between cousins as found from 40 contingency tables was "271 + '009 with a standard deviation of "083 + "006. The mean value of the degree of resemblance in temperament as found from 30 tables by fourfold process was -258 + -014, but the variability in this case was 35 p. c. greater, the standard deviation being •115 + 'OrO.

The temperament tables worked by contingency with three types of cousins gave the result -238 ±"010, with the reduced variability indicated by -045 + -007. These results are collected in Table I. They suffice to show that mean square contingency methods give more uniform results than the fourfold tables i. But the mean found from contingency for Health, Ability, Temper and Success does not sensibly differ from that found for the three divisions of temperament by fourfold tables leading to the coefficient of correlation. If we combine the results of both methods so as to

* Directions and form of schedule for this case are reproduced in Biometrika, Vol. n. pp. 359-60.

t The fourfold tables were worked for two groupings for the alternatives Excitable or Calm ; the Betwixts being thrown first into one group and then into the other, the average value of the correlation coefficient is that given in Table III. In the case of the alternatives Reserved and Exjjressive the Betwixts were thrown into the Reserved, and in that of the alternatives Sympathetic and Callous into the latter group. This was done after some consideration and enquiry as to the popular weight of terming an individual ' reserved ' or ' callous.'

\ The corresponding nine fourfold tables were somewhat erratic and gave a mean of only -19.

MEASURE OF RESEMBLANCE OF FIRST COUSINS

obtain a general average degree of resemblance between the ten types of cousins for seven characters, we find for the whole seventy tables :

Mean value '267 ± '008. Standard Deviation = -093 + -006. Accordingly we may conclude that the average degree of resemblance of cousins lies between -25 and "30, say at '27.

Table I. Mean Results by Different Methods.

Characters

Method

No. of Cases

Mean

Standard Deviation

Three Phases of Temperament

Ditto

Health Ability

Temper Success

Whole Series

Fourfold Division 1 Mean Square Contingency \ 3 x 3-fold Table

Mean Square Contingency 4 x 4-fold and higher- fold Tables

' Contingency Fourfold Division

: 1-

40 30

•258±-014

-238+ -010

•271 ±-009 | -267* ± -008

•115±-010

•045 ± -070

•083 + -006 •093 t -005

Diagram of Frequency of Coefficients of Resemblance in Cousins and Brethren.

325" 375 425 475 525 575 625 675

Cousinships Sibships

Mean from grouped values ; ungrouped value = -265.

2

LO

ETHEL M. ELDERTON

The fluctuation is no doubt considerable in our results. But we think that it lies far more in the difficulty of estimating psychical characters, than in any real variation in the degree of resemblance. The fluctuation is greatest precisely in those characters where personal bias and sex bias make the judgment more difficult.

We are now in a position to compare the intensity of resemblance between cousins with that between brethren. Diagram I, p. 9, shows graphically the distribution of the degree of resemblance of the 70 cases of cousins in this first series and of 65 cases of brethren, physical and psychical. The cases were numerically distributed as follows :

Table II.

o

i

o

3

I

I

1

IN

I 1

1

1

I

!

1

1

1

Totals

Cousinsbip Sibship

2

i

9

10

18

11

10 2

6 •5

1

14-5

2 22

15

8

3

70 65

Mean Cousinship : '27 Mean Sibship : '51

■008. Standard Deviation : -093 ± -005. •006. Standard Deviation : -068 + -004.

An examination of the graph shows that the cousinship group clusters at 25 and the sibship group at '5. These may, we think, safely be taken as working values for cousinship and sibship resemblance for either sex, and we may safely assert that brethren are on the average twice as closely related as cousins. This halving of the degree of resemblance corresponding to the fact that normal first cousins have two common grandparents, whole sibs have four.

It must be noted : (a) that our data for cousins are not drawn from the same records as those for brothers and sisters. While the Family Records here used for cousins enable us also to deal with brothers, these have not yet been tabled and reduced, except in the one instance of Intelligence. Here the adult brothers gave "54 as against the average of three cases of adult male cousins giving '34. Schuster found -56 from adult Oxford graduates for ability. Pearson found '52 for brothers at school and Schuster for schoolboys "56. The half of these fraternal values would be -27, which agrees well with the general cousin average, but not so well with the "34 which is a definitely higher value.

(b) that our data for cousins and sibs are neither from the same records, nor for the same range of characters. In the case of the sibships 21 values were for definitely measurable characters, a much more reliable class of material ; while the consulships of the first series do not present a single measurable character, and only one definitely physical estimate, that of Health. The characters which are common to

MEASURE OF RESEMBLANCE OF FIRST COUSINS

I I

both series are : Health, Ability and Temper in adult cousins, and in sibships of school children ; the following table gives the results for three classes :

Table II a.

Type

Intelligence

Health

Temper

Cousinship

Sibship

Cousinship

Sibship

Cousinship

Sibship

Male and Male Female and Female Male and Female

■34

•34

•34

•40 •47 •H

•31 •33

•30

■52

•51 •57

-18 •19 •25

•51 •49 ■51

Mean

•34

•46

■31

•53

•20

•50

Mean Cousinship "28. Half Mean Sibship -25.

Treated alone these cases would show a definitely larger degree of l-eseinblance for the cousinship, than for half the sibship, but this is not borne out for the whole material. The differences also of the ages of the subjects, adults and school children, and the methods of recording, by relatives and by school teachers, must also be borne in mind.

We consider on the basis of this first series that "25 is a good round working number for the cousinship. This denotes on the assumptions of linear regression and the equal variability of the cousins, that two cousins of an individual selected from unrelated stocks (i.e. maternal and paternal cousins) will give the same probable value for the character of an individual, as a brother of that individual with the same character as the mean of the cousins*. On the other hand the accuracy of the estimate will not be so great. In the first case it is o^ v 1 ('5)2 and in the second case o^ v 1 (-25)2 (-25)2, which measures the variability of the array; these are as 8'6G to 9*35. Thus the prediction from the brother would be somewhat better than from two mutually unrelated cousins. It is clear, however, that a knowledge of two such cousins may be very useful indeed, especially if facts as to the sibship are not forthcoming.

If we turn to other collateral relationships, the avuncular worked out for the eight possible cases in eye colourt, is, as far as we know, the only one yet published. The mean value of the eight cases is *265. We should accordingly conclude from this that for purposes of inheritance a knowledge of the cousin is equally important with a knowledge of the parental sibships. For example, there is no justification in medical histories of lunacy for including the facts as to the parents' brothers and sisters and

* Regression equation for 1 on 2 and 3, the latter being independent, is

h, = 'Ji^i 1H + 'J^ h3 = rnh2 + ruh3

<T., 0-3

t Phil. Trans. Vol. 195, A, p. 114.

if o-,

•50 x J {lh + A,), if

12 ETHEL M. ELDERTON

omitting the cousins from the record. On the same ground the marriage of niece or nephew with uncle or aunt seems to be a marriage of exactly the same degree of kinship as a marriage between first cousins.

The only grandparental data at present reduced for man* are those for eye-colour and the eight cases give a mean value of "32 1. This is somewhat higher than the value ("27) for cousins, and pigmentation data in horses have given an almost equal value. Still other species show rather smaller intensity, and until further data are reduced for the case of man, especially for psychical characters, we are not convinced that the grandparental relationship is definitely more important than the cousinship. At any rate, even with our present values (-27 as against "32) it will be seen that it is not reasonable for the purposes of medical or actuarial diagnosis to neglect the cousins, and make a considerable point as to the grandparental constitution. The grandparent, the uncle or aunt and the cousin are practically on the same footing with regard to relationship or intensity of kinship as measured by degree of likeness of character ; and it seems probable that any scientific marriage enactments would equally allow or equally forbid marriage between grandparent and grandchild, uncle and niece, aunt and nephew and between first cousins. This conclusion is reached on the assumption that the undesirability of marriage depends on the closeness of likeness in the gametic constitution, and that on the average the i-esemblance of the somatic characters may be taken as a measure of the average gametic resemblance between any two classes.

5. We now turn to the details of Table III.

We first ask whether there is any sensible difference between the intensity of inheritance in males and females. We note that the probable error of any individual result runs from about -02 for cousins of same sex to "03 for cousins of different sexes. Our table shows us that the average for pairs of male cousins is the same as that for pairs of female cousins, i.e. "26. If we could lay any stress on the difference "02, we should assert that cousins of different sexes were more alike than cousins of the same sex. But we certainly cannot, and thus, as far as our data go, we can only conclude that difference of sex makes no difference in degree of likeness.

In the next place we may consider whether type of cousinship makes any differ- ence in the intensity of resemblance. Our mean values for all the characters range from '22 to -31 according to the type, and it might be thought that this offered sufficient range to answer the question. As defining the types there are two considera- tions to be noted, (i) a difference of sex in either generation, parental or cousinal, and (ii) a change of sex in descent. Neglecting the first we have :

No change of sex in descent in : A (SO), E (-27) or K (-29) ;

One change of sex in descent in : C (-23), F (-28), G (-24) or H ('31) ;

Two changes of sex in descent in : B ('24), D ('22), or / ('29).

* The Family .Records of the present series provide unreduced material for seven characters, and this will shortly be dealt with.

t Phil. Trans. Vol. 195, A, p. 115.

MEASURE OF RESEMBLANCE OF FIRST COUSINS

13

The means for the three groups are '287, -265 and "250 respectively. It may possibly be therefore that change of sex slightly weakens the intensity of inheritance in the stock. If we turn to the first consideration the change of sex in the same generation, the connected parents are of the same sex and the cousins of the same sex in A and A. but in K the connected parents are of different sexes and the cousins o\' different sexes. A" is not. however, the least of the three. In C and /''there is a

Table III. General Results of First Scries. Characters of Cousins.

Type of Cousins

Health

Intelligence

Success

Temper

Temperament

Reserved or Expressive

Sympathetic or Callous

Excitable or Calm

Method -►

Male. Type A

B

c

M.S.C.

M. S. C.

M.S.C.

M.S.C.

F. T.

M.S.C.

F. T.

M.S.C.

F. T.

M.S.C.

•34 ■32 •26

•-tl •30 •32

•24 •15 •19

•23 •16 •15

■38

•06 •23

1 -20

•31 •36 •30

h

■21 •36

•17

•30

•24 ■23

Mean

•31

•34

•19

•18

•22

(■20)

■32

(•24)

•25

(•34)

•26

Female. Type D F

•51 •24

•23

•34 ■38 •31

•16 •27 ■35

•14

•24 •18

•21 ■30 •26

},

•03 •42 •37

1 -20

■15 •04

•23

|,

•22 •27

•28

Mean

•33

•34

•26

■19

•26

(•19)

•27

(■20)

■14

(•22)

•26

Male & Female. Tvpe G H I

•23 •32 •29

•37

•36 •27 ■38 ■34

•19 •33 •27 •26

■20 ■30

•24

•25

•23

•28

•42

1,4 I

•52 ■44 •19

■24

■24

14 ■16

■IS

-•28

•24 •31 •29 ■29

Mean

•30

•34

•26

•25

•29

(•24)

•34

(•24)

•18

(•28)

•28

General Means

•31

•34

•24

•21

•26

(•21)

•32

(■2.".)

•19

(•28)

■265

M. S. C. = Mean Square Contingency. F. T. = Fourfold Table, difference of sex in the connecting parents, but not in cousins ; in G and H, there is no difference in the connecting parents but one in the cousins. We might therefore expect no difference in the four values. But we find C and G contrasted in magnitude with F and H. In cither words two males related by a male and female go with a male and female related by two males ; and again two females related by a male and female go with a male and female related by two females. We can throw no light on this point, and it may only be a strange result of random sampling. In the third group there are no sex differences in the types B and D for either parental or cousinal generations.

14 ETHEL M. ELDERTON

For / there are such changes for both generations ; and yet / is larger than B and D. We must thus consider that a difference of sex in the same generation makes no difference in the intensity of resemblance so far as our present data go. Accordingly, if change of sex in descent does to some extent weaken inheritance*, it does not appear connected with sex differences in the same generation. The differences noted are, however, too slender and the whole system of values too fluctuating to build up any hypothesis as to sex influence in heredity.

If we now turn to the separate characters, and compare irrespective of cousin type the general means of each, we find an even wider range of results ('19 to '34). We attribute this only in part to real differences in the intensity of resemblance ; we consider it more due to (a) difficulties of estimating some of the characters dealt with, especially as in the case of cousins they are usually not in daily contact with each other; (b) differences of method employed, and the assumption that temperament follows a normal distribution of frequency.

Accordingly we shall draw no conclusions as to divergences in resemblance, believing our data may be relied upon to give a " general average resemblance " of cousins, but cannot be pressed beyond such a result to discriminate between individual of character.

6. We now turn to the results of the second series of quantitative measure- ments. These measurements as we have already noted are far from complete. They give for the four measurements 107 pairs of female cousins, 34 pairs of male cousins, and 111 pairs of male and female cousins, the two first sets giving 214 and 68 pairs in the symmetrical table.

The following table gives the statistical constants of the series of measurements. It will be observed that we have two series for each sex, but not all the individuals in each series are different.

This table shows at once considerable irregularities, which may be due to the paucity of data, or to the defective handling of the spanners. While the finger joint measurements give a sex-ratio for the absolute lengths = '91, very nearly the usual 11/12 of stature and of bone measurements in man, the widths of hand and wrist (involving a good deal more care in determination and allowing of more personal equation) give ratios of about H'5/12. For these also the man is both absolutely and relatively more variable than the woman. For the joint measurements the woman is equally variable absolutely with the man, and relatively more variable. It seems improbable that this equal absolute variability is correct. It is not true for the majority of bone measurements in man and woman. It is further to be noted that in the Male and Female Cousins series, where there was a much larger return of measure-

* The influence of change of sex has been very elusive ; it would appear to have some bearing on the inheritance of eye-colour in man (Biometrika, Vol. II. pp. 237-40), but we have failed to find it in coat-colour in horses (Ibid. Vol. n. pp. 229-34). It is doubtfully significant in the cases of coat-colour of Greyhounds (Ibid. Vol. in. pp. 257-8), and of Shorthorns (Ibid. Vol. iv. pp. 449-51).

MEASURE OF RESEMBLANCE OF FIRST COUSINS

15

Hunts made by male students than in the case of the Female and Female Cousins series, the absolute variabilities of the women are in every case less than in the latter series. In a certain number of cases it was actually found that the user of the hand-spanner had read from the sliding edge and not from the index point, but the difference amounting to about 20 mm. was obvious on the face of the measurements and at once allowed for by measuring the particular hand-spanner which had been used. It is believed that no residual error has crept in in this manner, but the point will be again dealt with below.

Table IV. Statistical Constants of Measurements of Hand in Man and Woman,

Sez

Series

Width of Hand

Width of Wrist

Joint,

Index Finger

Joint,

Little Finger

M.

S. D.

C. of V.

M.

S. D.

C.ofV.

M.

S. D.

C.ofV.

M.

S. D.

C.ofV.

Male

Male with Male Male with Female

83-2 82-1

6-20 5-77

745

7-03

58-2 57-9

3 90 4-17

6-70 7-20

62-5 62 0

2-87 3-23

4-59 5-21

51-0 50-9

2-72 301

5-33 591

Mean

82-6

5-98

7-24

58-0

4-04

7-00

62-2

3-05

4-90

51-0

2-87

5-63

Female

Female with Female Female with Male

72-5 71-4

4-79 3-64

6-61 5-10

51-8 51 0

3-38

2-78

6-53 5-45

57'2 56-3

3-47 3-06

6-07 5-44

46-4 46-1

2-94 2-70

6-34 5-86

"

Mean

72-0

4-21

5-84

51-4

3-08

5-99

56-8

3-26

5-74

463

2-82

6 09

Sex Ratio, Male and Female

•87

•70

•81

•89

•76

•86

•91

1-07

1-17

•91

■98

1 -08

The measurements are in mm.

The correlations as found by the product moment method without grouping are given in Table V. Now if we looked simply at the general mean 336 of all 12 results, we might conclude that the intelligence and health characters of our first series had given us the more reliable results, the temperament and temper being more difficult of estimate, and thence conclude that the average resemblance of cousinship was I .",.

Table V. Correlation of Measured Characters in Pairs of Cousins.

Character

Male and Male

Male and Female

Female and Female

Means

Width of Hand Width of Wrist Joint, Index Finger Joint, Little Finger

■3.3

•17 •19 •29

•21 •26 ■34 •37

•40 •43 •49 •56

•314 ■286 •340

•404

Means -245

•295

•470

•336

16 ETHEL M. ELDERTON

This view might be confirmed possibly by noting that the less easy measurements, those on hand and wrist, gave lower results than the joint measurements. But on further inspection of the table we notice that it is the female-female series which diverges so much from our previous results. The eight cases in which a male was one of the pair, and presumably worked the spanner, give a mean of -270, agreeing excellently with the -267 of our much larger first series. It is the pairs of female cousins, with their excessive variabilities, which give an intensity of resemblance equal to that of a sibship, and raising the average from -270 to "336.

To test the matter further the following steps were taken. A formula has been given by Pearson* which is based on normal distribution of frequency and gives the correlation coefficient in terms of the sum, S(x y), of the positive differences of correlated variates which have the same mean and s.D. Now this is precisely the case of the 106 cousin pairs if we treat them as a symmetrical distribution of 212 pairs. The formula is :

,.-1 y{s(x-y)Y

Applied to the data for the joint of the little finger in pairs of female cousins, we find r = '5578, while found by the product moment method the answer is "5579, a very close agreement. But it will be clear that if the measurer had a personal equation of the nature of a constant error for each pair, it would drop out in the difference x y for that pair. Hence the formula above is convenient to use when such an error for the individual pair is suspected to exist, and the variability cr can be found from other considerations. If in this particular case we adopt : (a) the standard deviation of the women in the series of male and female cousins, (b) the standard deviation found for the women on the assumption that the coefficient of variation for the women ought to be (what it usually is) practically the same as for the men, i.e. if we take the two values 2701 and 2-474, we find that the above formula gives r=-48 and "38 respectively. Thus indicating some considerable reduction from the value -56. It is therefore possible that an adding or subtracting of a constant difference in some of the measurements is the source of the exaggerated values of the female cousins resemblance. Such an error would not only have exaggerated the standard deviation, but it would have resulted at once, if a wrong correction had been applied to the measurements of those helpers who read at the edge, and not at the index point, of the spanner. It is believed that no spanners were removed from the numbered boxes until the measurements had been corrected ; but the doubt, however slight, to those who had the control of the instruments, is sufficient to. make it needful to repeat as soon as possible the whole series of measurements on female-female cousins.

We are able to use this second series as a control series also for the characters, hair colour, eye colour and general health. The method used was that of contingency

* "On further methods of determining Correlation," Drapers' Research Memoirs, Biometric Series IV. p. 4 et seq. (Dulau it Co., Soho Square).

MEASURE OF RESEMBLANf 'E OK FIRST COUSINS

17

but it must be remembered that the series were short, i.e. treated as symmetrical tables we had only 68, and 218 entries, and for male and female cousins 113. The results given in the following- table were reached.

Table VI. Non-quantitative Characters, Second Scries.

Character Male and Male

Male and Female

Female and Female

Means

Health Eye Colour Hair Colour

•38 ■44 •34

■29 •48 •26

•18 •38

•2G

•282

•434

•286

Means

■3S6

•343

■273

■33 1

It will be seen at once that (i) the preponderating intensity of pairs of female cousins no longer exists, (ii) the eye colour values are, however, very high, in one case at least approaching the intensity of the resemblance of siblings, and (iii) the generally higher value obtained in the case of the measurable characters of the same series is maintained. We have already noted that eye and hair scales were used in these observations, 24 eye and 24 hair tints being given. There were 6 categories in the Health graduation, but the " Very Delicate " and the " Very Robust " categories were only very slightly represented, so that for Health merely 3 x 3-fold contingency tables, " Robust," " Normally Healthy," " Delicate " seemed possible. For eye colour the 24 eve tints were first classed as " Pure Blue," " Blue with some orange," "Pure Grey," " Grey with some orange," " Hazel-Green," " Hazel-Brown," " Brown " ; the two greys were then clubbed together, as also the two hazels to form a 5 x 5-fold table for contingency. The 7 x 7-fold table seemed far too fine for the numbers, 68, involved in the male and male cousin tables, and it was desirable to treat all three tables alike. The 24 tints of the hair scale were first grouped into : " Very Dark," " Dark-Brown," " Brown," " Light-Brown," " Fair," " Red." But for the male data only a single "fair" and a single "red" occurred and only three "browns." Accordingly the 2nd and 3rd categories and also the 5th and 6th were grouped together and a 4 x 4-fold table used for the contingency of hair-colour. As samples, the Eye and Hair colour tables for pairs of female cousins are given in Appendix B as Tables LXXX and LXXXI. Now while we frankly admit that this Second Series, whether of measurable or pigmentation characters, has a much too inadequate frequency to be conclusive, still its drift is undoubtedly to confirm the view, that the average resemblance of cousins is higher than that given by the Family Record results. It approaches nearer the value indicated by the more precise of the " Record " characters, and the more accurate of the hand measurements. The numbers in the first series are large as compared with those of the second, and the second series also involves several

18 ETHEL M. ELDERTON

points of doubt and difficulty ; for this reason we have not yet modified the general average of the First Series by including these higher results. But it is conceivable that we may have to raise the general measure of resemblance of cousins from "28 to •33, when other large series already observed have been tabulated and reduced.

7. One further point may be finally touched upon, namely the inheritance of disease. We cannot in the least hope here for accurate numerical estimates, but the data of our first series may suffice to show that cousins are of value even from the standpoint of medical diagnosis. The difficulties of accurate determination are as follows :

((() While on the schedules the record of brothers and sisters, of children, of parents and of grandparents is fairly complete, that of cousins must necessarily be defective. It is quite possible- nay not infrequent— to have more than 50 first cousins. And while one or two recorders actually were patient enough to enter details of a cousinship as large or even larger than this, the bulk of recorders contented themselves with entering a much more limited number, 10 to 12, and thus we have the first limitation; our cousins, as the recorders themselves state, are a selection. It is probable, also, that the selection has been made more frequently of living than of dead cousins, and more frequently of accessible than possibly inaccessible cousins ; thus the individuals suffering from phthisis or insanity, or having died from these diseases, may without direct intention to deceive have been more frequently omitted than in the case of relatives all of whom were included.

(b) The cousins in our family record schedules are those of the subject. In order to get full ancestral information a young adult has been very often taken as the subject and the cousins belong accordingly to the third generation, and are themselves often young adults. It follows accordingly that their medical history is in many cases incomplete. They have not passed wholly through the danger zone in the case of either tuberculosis or insanity.

In the case of tuberculosis, we have for instance among males only 206 tuberculous out of 2990 individuals, and among females only 205 out of 3242, whereas 10 p.c. would probably be affected if we had the full record.

In our records for example there are in the case of women 130 cases of individuals classed as cousins with some form of brain disease or mental defect*. These 130 individuals have 6 insane and 124 sane cousins. If in the remainder of their lives 4 persons out of those 124 sane cousins were to suffer from some form of brain attack, then our table would be as follows :

* "Insanity" for the purpose of this investigation has been taken to include the neuroses: confirmed alcoholism and marked hysteria. These were not included by Heron in using Pearson's Family Records (Eugenics Laboratory Pali/iratiiut.i, n. p. 33). Its use here approaches "want of mental balance."

MEASURE OF RESEMBLANCE OF FIRST COUSINS

First Female Cousin

I

L9

Insane Sane

Totals

Insane Sane

14 120 120 2996

134 3116

Totals

134

3116

3250

instead of the actual

First Female Cousin

Insane Sane

Totals

Insane Sane

i 6 124 124 2996

130 3120

Totals

130 3120

3250

The fourfold table method gives the correlation of the first table about -33 and that of the second "03. Now it is not suggested that four additional cases of insanity are what we have to expect in the case of 124 persons chiefly young adults of insane stock. What we wish to point out is that with a disease so relatively rare as this, the transference when the record is completed of comparatively few individuals from the sane to the insane category is sufficient to raise the intensity of resemblance to a value quite equal to that which we have found for other characters in cousins.

The following are the results reached for insanity and tuberculosis :

Table VII. Inheritance of Pathological Condition in Cousins with

incomplete Record.

Male Cousins Female Cousins

Male and Female Cousins

Means

Insanity -18

Tuberculosis ! -07

•03 ■12

■08 •19

•10 •13

Means -12

•08

•13

•11

In all six series and they number in each case about 3000 pairs we have a positive relationship, and the value is definitely significant in all cases but possibly that of insanity in female cousins. Yet this, owing to the fact that a considerable amount of insanity in the case of women is connected with change of life, is precisely what we might have anticipated considering the ages of our cousins. The fact also that insanity has a later average incidence than tuberculosis may explain why the average value for tuberculosis is higlier than for insanity. We should conclude that so far as our data go they show that the tendency to both insanity and tuberculosis runs in stocks, and that with the incompleteness of the record there is no reason to

20 ETHEL M. ELDERTON

suggest that disease tendencies are not inherited at the same rate as physical and psychological characters in cousins.

8. General Conclusions. Our memoir has dealt with two series of cousin records. The quantitative measure of the resemblance of cousins is of great importance not only on account of its bearing on eugenic marriages, but because cousins form often the principal living record to assist medical diagnosis. Its determination, however, presents considerable difficulties. It is not hard to collect data as to the characters of cousins, when these characters can be judged without the actual presence of the cousins. This was done in our first series. But when we come to the quantitative measurement of cousins our experience has been unfavourable to the rapid accumulation of extensive material. The passing from brethren to cousins - although the latter are a far wider group has more than trebled the difficulty of obtaining measurements. Further our choice of the hand as the organ to be dealt with has possibly led to difficulty, as the treatment and use of the spanner needed more care than a simple measuring tape. It was possible to explain and illustrate the use of the spanner to all the male students to whom it was loaned, but in the case of women helpers we had often to trust to written directions. This may be the source of the high values found for the resemblance of women cousins, but we confess frankly that we are not satisfied that it is so, and we must await the reduction of further material before settling this point. If we turn to the 70 cases dealt with on the basis of our first series, we find an average resemblance of about -27, which tallies with the average found from the eight quantitative series involving male cousins in our second investigation. If this value be confirmed we should say that cousins have as much significance as the parental brothers and sisters. On the other hand an examination of our table shows that what may be treated as the more easily judged and reliable results, show a rather higher value than '27, approximating rather to the '33 of the grand parental resemblance. The pigmentation results of the second series tend to confirm this view.

We should conclude accordingly from the present results that for the purposes of eugenics cousins must be classed as equally important with uncles and aunts, and that they may eventually turn out to be as important as grandparents. For practical purposes it would hardly seem possible in the matter of marriage restrictions based solely on the gametic resemblance judged by somatic characters, to differentiate between the three classes. This equality of resemblance which may appear at first sight paradoxical will be confirmed for uncles and aunts in a forthcoming memoir. Its physiological bearing appears to us of fundamental importance as indicating that a determinantal theory of heredity, emphasising alternate inheritance, must take pre- cedence of any theory of simple blending for the bulk of the characters here dealt with.

We do not consider that our data show any difference between the inheritance of physical, psychical and pathological characters, which could not be accounted for by (a) the difficulty of appreciating temperament, and (b) the incompleteness of the cousin record.

APPENDIX A.

HEREDITARY RESEMBLANCE OF FIRST COUSINS*.

I. OBJECT OF MEASUREMENTS AND GENERAL INSTRUCTIONS.

I.— The present state of our knowledge of the laws of inheritance in man may be summed up as follows :

We know well for a variety of organs direct inheritance from parent to offspring, and the collateral relationship between brothers and sisters. We have less complete, but still valuable data for the direct line in the case of grandparent and great-grand- parents, and for the collateral line in the case of uncles and aunts. To supplement our knowledge, one of the most urgent problems is the determination of the degree of resemblance between cousins. It is with a view of solving this problem of cousin relationship that I appeal for cooperative observations and issue the present paper and schedules.

II. For the purposes of the present investigation we are to understand by the word cousin ;

(i) Full blood First Cousins, that is children of two whole (not half) brothers, of two whole (not half) sisters, or of a whole (not half) sister and brother. Such cousins are to have one and only one grandparental pair in common, and we term them normal cousins.

(ii) " Abnormal " first cousins are to be excluded.

It may happen that two brothers of one family have married to two sisters of another family, or that a sister and brother of one family have married a brother and sister of a second. The issue of such marriages are "doubly" first cousins having all their grandparents in common. Again, a brother and sister in one family might marry an aunt and a nephew in a second family, or again, might marry a woman and a man who are cousins in a second family, or, two brothers may marry two half sisters in a second family. Indeed cases of abnormal cousinship occur in which the abnormal cousins have 1, 2, 3 or 4 common grandparents. All such cases are excluded from the present investigation, which is concerned only with normal cousinship as defined under (i).

(iii) Normal cousins for the purpose of this investigation must be between 18 and 45 years of age.

* Issued by Professor Pearson, 1902 and onwards.

22 ETHEL M. ELDERTON

We cannot for a longer period consider the eye and hair colour to remain even approximately constant. With a shorter period we might fail to obtain sufficient material for statistical purposes.

III. There are ten kinds of normal first cousins. Let A and B stand for the two cousins, thus :

Tiro (A and B may be sons of two brothers. Male - A and B may be sons of two sisters. Cousins\A and B may be sons of a sister and brother.

Two (A and B may be daughters of two brothers. Female-, A and B may be daughters of two sisters. ( husins\A and B may be daughters of a brother and sister. Male I A and B may be son and daughter of two brothers. and ) A and B may be son and daughter of two sisters. Female \A may be the son of a sister and B the daughter of a brother. Cousins [A may be the son of a brother and B the daughter of a sister. In this classification in the last group of " male and female cousins," A is taken as the male and B as the female cousin. But in the actual schedule provision is made for the case where the observer has taken A for the female and B for the male cousin.

The observer, after entering his or her own name, should fill in the names of the cousins A and B and their sex by putting a cross under male or female. Next, under type of cousinship, a cross should be put in the last column against the special type of the two cousins observed. This is very important, because we have reason to believe from the grandparental and avuncular relationships that the degree of resemblance varies a good deal with the type.

IV. Any individual cousin A may be dealt with in any number of cases, but it is not desirable to compare one cousin A with more than four other cousins who are brothers and sisters to each other, and of these, not more than two should be of one sex. Subject to this limitation A may appear, or A's brothers and sisters, in any number of cousinships. A fresh schedule should be used for each such cousin- ship. It is not, however, necessary to fill in on these additional schedules all the measurements and characters. The name and sex only of the repeated cousin, and the type of cousinship, need to be inserted. A cross reference to the number of the observer's series in which the cousin is fully recorded will then suffice. A blank is left for this reference under the name and sex of cousin. For example, if P, Q, R, S be children of four different brothers and sisters, we first fill up a schedule of P and Q, then one of R referring to the schedule containing P ; and another schedule referring to the schedule containing R and to the one containing Q, but giving the type of Q and R cousinship. Then we measure S and refer to P, and

MEASURE OF RESEMBLANCE OF FIRST COUSINS 23

finally two more schedules give merely the names of S and Q, and <S and R with their types of cousinship, and refer to the proper schedules for the observations on S, Q and R. Or again, the observer may till in one schedule for himself or herself, and then with simple reference to the number of that schedule and the type of cousinship, till in separate papers for twenty or thirty of his or her cousins. Then another series of schedule papers may be filled with simple references to the indi- viduals among these twenty or thirty persons (without repeating their measurements) who are cousins among themselves apart from their relationship to the observer. In each case the type of cousinship must he marked on the new schedule paper.

V. Directions for recording Observations.

(1) Hair Colour.

On the hair-colour scale in the box, pick out the number of the hair corresponding most nearly to the colour under observation. If the hair considered falls between two tints, so exactly that you cannot say that it is nearer to the one than the other, give both tints, thus 5 6. If the hair has turned grey before 45, say so ; and if the hair is of tint distinctly not on the scale, fasten a very small sample, sufficient to show colour, on the data sheet with the border of a sheet of postage stamp, or other strip of gummed paper.

(2) Eye Colour.

In judging eye colour, first fix the attention on the amount of orange-brown pigment in the iris. If there be no orange-brown pigment, the eye is (l) Dark Blue, (2) Blue, (3) Light Blue, (4) Light Grey. With hardly visible amounts of the orange- brown pigment we have next (5) Blue-green, (G) Dark Grey, (7) Hazel. Lastly, with clearly marked orange-brown pigment, we have (8) Light Brown, (9) Brown, (10) Dark Brown, (11) Very Dark Brown, (12) "Black." Samples of these eye types are given on the eye-colour scale. Look at the eye with the light upon it from a distance of about 1 8 inches and compare it with the scale. If the eye falls between two types on the scale, give the numbers of both types ; if it agrees fairly well with any type, give the number of that single type only. Thus G 7 would mean that the eye in question fell between 6 and 7 of the scale, but 7 would signify that it was closer to the 7 than to the 6 of the scale.

(3) Health.

Place a cross against the category under which the general health falls.

(4) Measurements of Hand.

These are to be made with the hand-spanner which will be found in the box. All the readings are to be taken to the nearest mark on the scale, and the observer need not give fractions of the units on the scale, if the length falls between two marks. If in any case the observer finds it quite impossible to determine which is the nearer mark, then give both units, e.g., 34 35.

24

ETHEL M. ELDERTON

Fig. (i).

Self-measurement of the left hand by means of the hand-spanner.

(i) Width of Wrist. See Figure (i).

Feel for and satisfy yourself as to the positions of the bony protuberances on either side of the main joint of the wrist. They are the sides of the ends of the two bones of the forearm. The space between the outer sides of these has to be measured with tbe spanner. Hold the spanner in the right hand, resting its fixed jaw against tbe breast, and manipulate the movable jaw with the spare fingers of the right hand. Lay the left wrist back upwards between the jaws of the spanner, so that the bony protuberances come against the jaws. Close the jaws with gentle pressure and clamp the movable jaw with the clamping nut under- neath. Repeat this at least once, and if time will allow twice, taking the reading each time and entering it on the schedule. Do not be surprised if your measurements are not exactly the same. Only suspect something is wrong, difference in your results. If this be so, test carefully again. Do not fill in column marked "mean," but leave this to those who have to reduce the observations.

(ii) Width of Hand. (Left hand, as before.) See Figure (ii).

Feel for and satisfy yourself as to the positions of the outer sides of the knuckles, the one side being formed at the joint at the base of the little finger, the other at that of the forefinger. The hand is to be placed with the fingers close together, with the palm upwards, and all the knuckles t muLiug the spanner. Measure the width between the outer sides with the spanner held with the fixed jaw against the breast and the scale horizontally upwards. Bring the movable jaw without pressure against the knuckle at the base of the forefinger. Clamp and read the scale. N.B. Take care to make two or three trials.

you

find

two units

MEASUIiF OF ItFSFMRLAXOE OF FIRST COUSINS

25

(iii) Length of First Joint of Index and Little Fingei

See Figure (iii).

Close the fist (thumb outside) and thumb uppermost and spanner hori- zontal— the lengths from knuckle to first joint of (a) the Index Finger ; (b) the Little Finger. The outside of first joint is put against the tixed jaw of the spanner, and the movable jaw is brought against the outside of the knuckle with gentle pressure. Clamp and read as before, making two or three trials.

All the measurements should be made with care. The above instruc- tions are intended for self-measurement, but it is easy for one observer to measure both of a pair of cousins, or him or herself and then the cousin

Fig. (iii).

VI. In case of any difficulty, please apply at once to Professor Karl Pearson, University College, London, W.C. The Box and papers should not be kept longer than a month, unless the observer finds it possible to undertake a large series of cousins. About a thousand pairs of cousins of each type, 10,000 in all, will be required. Hence every co-operator will appreciate the necessity for rapid circulation of the boxes, of which only a limited number can be provided.

The name of the observer and address should always be given, in case it is necessary to ask questions as to any special measurement or observation. The cousins, if it be preferred, may be simply denoted by the initials of their christian and surnames, as these will suffice for the observer to identify them*.

* Spanners and schedules are still (November, 1907) being issued, measurements will be gratefully accepted.

id help in further cousin

26

ETHEL M. ELDERTON

II. SCHEDULE.

Kindly make no attempt to fill this Schedule in until the General Instructions have been carefully read through.

Hereditary Resemblance of First Cousins.

Ages between 18 and 45.

Observer Name

Number in Observer's Series

Number in whole Series*

* Leave this space blank.

against Sex of Cousin.)

1 Male

Female

Male Female

Cousin B.

Type of Cousinship. (Place cross against type in right-hand column.

Male Cousins

(i) A and B are sons of two brothers (ii) A and B are sons of two sisters

r A is son of brother, B is son of sister 1 A is son of sister, B is son of brother

Female Cousins

(iv) A and B are daughters of two brothers (v) A and B are daughters of two sisters .. .

f A is daughter of brother, B is daughter of sister ^Tl' \ A is daughter of sister, B is daughter of brother

Male and Female Cousins

f A is daughter, B is son of two brothers ^V11' \ A is son, B is daughter of two brothers

. J A is daughter, B is sou of two sisters \VU1' [ A is son, B is daughter of two sisters

f A is daughter of a brother, B is son of a sister

^'X' t A is son of a sister, B is daughter of a brother

1 A is daughter of a sister, B is son of a brother

^X' 1 A is son of a brother, B is daughter of a sister

A's measurements are already given on Schedule No.

B's measurements are already given on Schedule No.

only to be used if A and B have already been scheduled for other pairs of cousinships,

MEASURE OF RESEMBLANCE OF FIRST COUSINS

27

(1) Hair-Colour.

Insert number of nearest tint on hair-colour scale

(2) Eye-Colour.

Insert number of nearest tint on eye-colour scale

A

B

A

B

(3) Health.

Place a cross against the category which seems best to describe A's general health and a second in the last column for B's health.

(4) Measurements of Left Hand.

To be made with the hand-spanner as described in the General Instructions.

A

B

Very Robust

Robust

Normally Healthy

Rather Delicate

Delicate ...

Very Delicate ...

Measurement

A

B

1st Trial

•2nd Trial

3rd

Trial

Mean

1st Trial

•2nd Trial

3rd Trial

Mean*

(i) Width of Wrist

(ii) Width of Hand

(iii) Length of First Joint, Index Finger

(iv) Length of First Joint, Little Finger

* This column is to be left blank.

To ensure accuracy it is desirable that two or three trials should be made of these measurements, if they are not taken by an independent observer who has measured already a considerable number of pairs.

Kindly return this Schedule when tilled in to Professor Karl Pearson, University College, London, W.C.

APPENDIX B.

TABLES OF DATA.

Health. Male Cousins.

Table I. Type A.

First Male Cousin

Very Robust

Robust

Normally

Healthy

Rather Delicate

Delicate

Totals

Very Robust

"

21

5

2

3

31

a

3

Robust

21

64

77

5

22

189

1

Normally Healthy

5

77

206

17

79

384

1

Rather Delicate

2

5

17

-

2

26

Delicate

3

22

79

2

26

132

Totals

31

189

384

26

132

762

Table II. Type B.

First Male Cousin

Table III. Type C.

First Male Cousin

V. R.

R.

N. H.

R. D.

D.

Totals

o U

1 -a o 1

V. R.

R.

N. H.

R. D.

D.

Totals

V. R.

30

31

28

1

5

95

V. R.

18

52

45

2

18

135

o

R.

31

98

77

5

2.3

236

R.

52

156

136

13

28

385

1

N. H.

28

77

196

13

35

349

N. H.

45

136

314

15

70

580

o 02

R. D.

1

5

13

-

2

21

R. D.

2

13

15

4

2

36

D.

5

25

35

2

14

81

D.

18

28

70

2

28

146

Totals

95

236

349

21

81

782

Totals

135

385

580

36

146

1282

MEASURE OF RESEMBLANCE OF FIRST COUSINS

29

Health. Female ( Iousins

Table IV. Type />.

First Female Cousin

Table V. Type E.

First Female Cousin

V. B.

B.

N.H.

B. D.

D.

Totals

15

Second Female Cousin

V. B.

B.

N.H.

B. D.

D.

Totals

V. E.

24

7

10

-

4

V. B.

-

(i 46

15

-

6

27

■5

0

c

R.

7

44

99

5

is

203

B.

6

60

1

14-

127

Is

a

X. H.

10

99

2 7 1'

9

127

517

N.H.

15

CO

200

4

78

357

I

B. D.

-

5

9

8

3

25

B. D.

-

1

4

-

-

5

D.

4

48

127

3

74

256

D.

6

14

78

-

22

120

Totals

45

203

517

25

256

1046

Totals

27

127

357

5

120

636

Table VI. Type F.

First F

-male

Cousii

V. E.

E.

N.H.

B. D.

D.

Totals

V. E.

8

31

22

-

14

75

1

E.

31

66

123

1

66

287

1

N.H.

22

123

354

17

136

652

a

z

I

X

B. D.

-

1

17

-

2

20

D.

14

66

136

2

58

276

Totals

75

287

652

20

276

1310

30

ETHEL M. ELDERTON

Health. Male and Female Cousins.

Table VII. Type G.

Male Cousin

Table VIII. Type H.

Male Cousin

V. R.

R.

N. H.

R. D.

D.

Totals

3

O

a

V. R.

R.

N. H. 22

R. D.

D.

Totals

V. R.

4

20

8

12 22

5

27

82

2 29 145

37

V. R.

18

6

-

2

48

.s

R.

10

66

85

200

R.

19

52

45

2

21

130

o

o

"3

N. H.

15

86

231

436

N.H.

27

63

163

14

58

325

b

R. D.

-

7

10

21 190 884

R. D.

1

3

4

-

1

9

D.

10

57

92

2

D.

16

31

84

1

33

165

Totals

39

236

126

38

Totals

SI

155

318

17

106

677

Table IX. Type I.

Male Cousin

Table X. Type K.

Male Cousin

V. R.

R.

N.H.

R. D.

D.

Totals

a o O

c

V. R.

R.

N.H.

R. D.

D.

Totals

V. R.

8

8

13

-

6

35

V. R.

14

13

9

-

5

41

p

R.

18

55

67

0

26

168

R.

25

46

41

2

16

130

o O

N.H.

17

82

217

13

38

367

N.H.

11

95

219

18

47

390

i

R. D.

-

4

9

4

1

18

R. D.

-

-

5

2

-

'

D.

7

33

55

2

18

115

D.

17

34

81

1

24

157

Totals

50

182

361

21

89

703

Totals

67

188

355

23

92

725

MEASURE OF RESEMBLANCE OF FIRST COUSINS

31

Intelligence. Male Cousins.

Table XL Type A.

First Male Cousin

F&E2

E

E, & D

C,B*A

Totals

r & E,

14

42

36

5

97

E

42

208

:,:,

14

319

E, AD

36

55

L08

20

219

C.B&A

5

14

20

18

57

Totals

97

319

219

57

692

Table XII. Type B.

First Male Cousin

F&ES

E

Ej &D

C,B&A

Totals

F &E„

26

47

31

3

107

E

17 246

57

23

373

E, & D

31

57

54

19

161

C.B&A

3

23

19

6

51

Totals

107

373

161

51

692

Table XIII. Type C.

First Male Cousin

F&E2

E

E, &D

C.B&A

Totals

-.

F & E,

26

87

40

8

161

3

E

87

448

147

26

708

E, &D

40

147

118

30

335

X

C.BcfcA

8

26

30

30

94

Totals

16]

708

335

94

1298

32

ETHEL M. ELDERTON

Intelligence. Female Cousins.

Table XIV. Type D.

First Female Cousin

F&E2

E

Ej&D

C.B&A

Totals

•1

F&E,

6

23

29

1

59

o

E

23

336

133

14

506

"c«

£

e,*d

29

133

198

37

397

T3

C.B&A

1

14

37

8

60

Totals

59

506

397

60

1022

Table XV. Type E.

First Female Cousin

F&E2

E

Ej & D

C.B&A

Totals

F & E.:

8

9

17

1

35

E

9

262

97

22

390

E, &D

17

97

56

10

180

C,B&A

1

22

10

18

51

Totals

35

390

180

51

656

Table XVI. Type F.

First Female Cousin

F & E.,

E

Ej&D

C.B4A

Totals

F&E,

8

37

15

4

64

E

37

606

155

21

819

El(fcD

15

155

160

24

354

C.B&A

4

21

24

8

57

Totals

64

819

354

57

1294

MEASURE OF RESEMBLANCE OF FIIiST COUSINS

33

Intelligence. Male and Female Cousins.

Table XVII. Type G.

Male Cousin

F&E.

E

E, & D

C,B<S \

Totals

F&E,

11

30

15

56

B

3 0

E

09

266

60

31

429

"5

=

fa

E, ct D

63

93

124

50

330

C.B&A

•2

12

21

13

48

Totals

1 15

401

220

97

s.;:;

Table XVIII. Type II.

Male Cousin

1 A Es

E

Bj .v- D

CB&A

Totals

P & )•:,

13

46

12

2

73

c

3 O

O

E

29

279

74

20

102

S

E, & D

25

107

50

5

187

C.B&A

4

6

10

7

27

Totals

71

438

146

34

689

Table XIX. Type I.

Male Cousin

F & Ea

E

E, & DC, BAA

Totals

F & E,

9

23

13 ' 5

50

c

3

s

E

55

285

67 17

424

1 fa

E1 &T>

22

60

77 9

168

C.B&A

4

14

12 13

43

Totals

90

382

169 14

685

Table XX. Type A".

Male Cousin

FAE.

E

E, & D

CB&A

Totals

F & E,

17

23

9

1

50

1

E

34

334

93

19

480

"3 fa

E, & D

12

67

70

13

162

c.b&a

6

11

8

4

29

Totals

69

435

180

37

721

34

ETHEL M. ELDERTON

Success. Male CousrNS. Table XXI. Type A.

First Male Cousiu

Marked &

Prosperous

Average

Difficult

Failure

Totals

Marked & Prosperous

39

102-25

27-75

15

184

Average

102-25

196-5

40-25

16

355

Difficult

27-75

40-25

28

6

102

Failure

15

16

6

10

47

Totals

184

355

102

47

688

Table XXII. Type B.

First Male Cousin

M. & P.

A.

D.

F.

Totals

■1

M. & P.

90

96

35

14-5

235-5

0

8

0>

A.

96

139

47-5

14-5

297

1 £

D.

35

47-5

22

7

111-5

F.

14-5

14-5

7

,

44

Totals

235-5

297

111-5

44

688

Table XXIII. Type C.

First Male Cousin

M. & P.

A.

D.

F.

Totals

a

M. &P.

79

147-75

47-25

26

300

o O

A.

147-75

272

80-75

24

524-5

D.

47-25

Si 1-75

51-5

6

185-5

1

F.

26

24

6

10

66

Totals

300

524-5

185-5

66

1076

MEASURE OF RESEMBLANCE OF FIIIKT COUSINS

35

SrccKss. Female ( 'oi sins.

Table XXIV. Typi />. First Female I lonsin

M.A P.

A.

D.

F.

roi ii

M.A P.

21

5975

6-25

-

87

A.

59-75

450-5

30-25

8

548-5

D.

6-25

30-25

-

-

36-5

F.

-

8

-

-

8

Totals

87

548-5

36-5

8

680

I \i;i.k \\\ Type I: Firsi Female Cousin

Table XXVI. Type F.

First Female Cousin

M.AP.

A.

D.

F.

Totals

M. & V.

64

94

6-5

12

176-5

A.

94

604-5

33-5

4

736

D.

6-5

33-5

5-5

-

45-5

F.

12

4

-

16

Totals

176-5

736

45-5

16

974

36

ETHEL M. ELDERTON

Success. Male and Female Cousins.

Table XXVII. Type G.

Male Cousin

M. & P.

A.

D.

F.

Totals

M. & P.

20

32-25

23-75

17

93

a

A.

104-5

224-75

83-75

43

456

a

D.

6-5

21

8-5

4

40

F.

4

1

-

-

5

Totals

135

279

116

64

594

Table XXVIII. Type H.

Male Cousin

M. & P.

A.

D.

F.

Totals

M. d- P.

23-5

24-5

5

11

64

A.

78-75

1 18-75

44

10

281-5

D.

6-25

21-25

24

6

57-5

F.

5

6

1

2

14

Totals

113-5

200-5

74

29

417

Table XXIX. Type I.

Male Cousin

M.&P.

A.

D.

F.

Totals

M.&P.

20-5

35-5

31-5

9

96-5

A.

91-75

228-25

54-5

9

383-5

D.

8-75

9-25

5

-

23

F.

1

-

-

-

1

Totals

122

273

91

18

504

Table XXX. Type K.

Male Cousin

M. & P.

A.

D.

F.

Totals

M. A P.

38

24-75

12-75

9-5

85

1

A.

80-75

191-75

75

17

364-5

1

D.

5-75

7-5

5-75

1-5

20-5

F.

3

2

4

•2

11

Totals

127-5

•226

97-5

30

481

MEASURE OF KFSEMBLANCF OF FIRST COUSINS

37

Temper. Male ( Iousinj

Table XXXF Type A.

First Male Cousin

Even

Quick

Sullen

Weak

Totals

a

Even

239

68

28-5

20-5

356

o O

Quick

68

65

1625

12-75

162

a -

Sullen

28-5

16-25

c,.-.

375

55

oa

Weak

20-5

1H-7--.

3-75

4

41

Totals

356

162

55

41

614

Table XXXI 1. Type 11.

First Male Cousin

Even

Quick

Sullen

Weak

Totals

_

Even

202

84-5

17-5

17-5

321-5

0

O

Quick

84-5

■_".i

9-5

7-5

130-5

C

8

V.

Sullen

17-5

9-5

-

5

32

Weak

17-5

7-5

5

2

32

Totals

321-5

130-5

32

32

516

Table XXXIII. Type C.

First Male Cousin

Even

Quick

Sullen

Weak

Totals

Even

419-5

179

43-5

24

666

Quick

179

96-5

22

17

314-5

Sullen

13-5

22

11

8-5

85

Weak

24

17

8-5

5

54-5

Totals

666

314-5

85

54-5

1120

38

ETHEL M. ELDERTON

Temper. Female Cousins.

Table XXXIV. Type D.

First Female Cousin

Even

Quick

Sullen

Weak

Totals

•i

Even

361

118-25

61-25

18-5

559

o Q

Quick

118-25

77-5

27-75

8-5

232

Sullen

61-25

27-75

10

5-5

104-5

as

Weak

18-5

S-5

5-5

-

32-5

Totals

559

232

104-5

32-5

928

Table XXXV. Type E.

First Female Cousin

Even

Quick

Sullen

Weak

Totals

Even

269-5

75-5

16-5

6

367-5

Quick

75-5

64-5

7-5

4

151-5

Sullen

16-5

7-5

3

27

Weak

6

4

-

10

Totals

367-5

151-5

27

10

556

Table XXXVI. Type F.

First Female Cousin

Even

Quick

Sullen

Weak

Totals

Even

545-0

194-5

49-5

11-5

801

6

Quick

194-5

116-5

22

6-5

339-5

5

Sullen

49-5

22

4

6

81-5

o

1

Weak

11-5

6-5

6

-

24

Totals

801

339-5

81-5

24

1246

MEASURE OF RESEMBLANCE OF FIRST COUSINS

39

Temper. Male and Female Cousins. Table XXXVII. Type G. Table XXXVIII. Type II.

Male (

ousin

Even

Quick

Sullen

Weak

Totals

Even

265-5

98-5

500

23

137

s

6

Quick

104 23

72-25

•29-0

19

•224-5

"3 =

Sullen

33

8-5

9-5

2

53

Weak

12-75

6-75

3

7

29-5

Totals

415-5

186

91-5

«

744

Male C

ousin

Even

Quick

•Sullen

Weak

Totals

Even

206-5

74

25-5

25-5

33 1 -5

o O

Quick

89

49

26

7-5

171-5

ta

Sullen

13-5

6-5

6

1

27

Weak

■2

1

-

5

8

Totals

311

130-5

57-5

39

538

Table XXXIX. Type I.

Male Cousin

Even

Quick

Sullen

Weak

Totals

Even

230

78

21

11-5

340-5

o O

Quick

108-25

64-75

21

9-5

203-5

X

Sullen

1 1-25

10-75

4

2

31

Weak

6

5

-

-

11

Totals

358-5

158-5

46

23

586

Table XI, Type K.

Male Cousin

Even

Quick

Sullen

Weak

Totals

Even

2735

65-25

20-75

12

371-5

a

o O

Quick

100-25

61-25

11

14-5

187

S

Sullen

14-25

14:5

5-25

3-5

37-5

Weak

10

3

1

2

16

Totals

398

1 11

38

32

612

40

ETHEL M. ELDERTON

Temperament Reserved or Expressive. Male Cousins. Table XLI. Type A. Table XLII. Type B.

First Male Cousin

Reserved

Betwixt

Expressive

Totals

» o O

1

Reserved

100

47

46

193

Betwixt

47

58

31

136

1

Expressive

46

31

68

145

Totals

193

136

145

474

First Male Cousin

Reserved Betwixt

Expressive Totals

Reserved

32

44

62

138

_0>

1

Betwixt

44

64 27

135

Expressive

62

27 50

139

Totals

138

135 139

412

Table XLII1. Type C.

First Male Cousin

Reserved

Betwixt

Expressive

Totals

Reserved

202

105

107

414

Betwixt

105

108

68

281

Expressive

107

68

112

287

Totals

414

281

287

982

MEASURE OF RESEMBLANCE OF FIRST CoUSlXS

41

Temperament Reserved or Expressive. Female Cousins.

Table XLIV. Type D.

First Female Cousin

Table XLV. Type E.

First Female Cousin

Reserved

Betwixt

Expressive

Totals

1

<D

-c

1

Reserved

Betwixt

Expressive

Totals

s •S3

1

Reserved

94

83

100

277

Reserved

32

34

53

119

1

Betwixt

83

110

62

255

Betwixt

34

74

50

158

a o a

Expressive

100

62

122

284

Expressive

53

50

134

237

Totals

277

255

284

816

Totals

119

158

237

514

Table XLVI. Type F.

First Female Cousin

Reserved

Betwixt

Expressive

Totals

o O

Reserved

86

111

117

314

1=5

Betwixt

111

120

110

341

o a; 02

Expressive

117

110

240

467

Totals

314

341

467

1122

42

ETHEL M. ELDERTON

Temperament Reserved or Expressive. Male and Female Cousins.

Table XL VII. Type G

Male Cousin

Reserved

Betwixt

Expressive

Totals

a

Reserved

87

50

53

190

3

Eetwixt

57

87

41

185

&

Expn ssive

106

47

96

249

Totals

250

184

190

624

Table XLVIII. Type H.

Male Cousin

Reserved

Betwixt

Expressive

Totals

.2

Ki-rni'd

26

26

27

79

o

a

_2

Betwixt

72

69

33

174

£

Expressive

105

31

89

225

Totals

203

126

149

478

Table XLIX. Type I.

Male Cousin

Table L. Type K.

Male Cousin

Reserved

Betwixt

Expressive

Totals

5 O

1

Reserved

Betwixt

Expressive

Totals

Reserved

77

35

35

147

Reserved

63

59

25

147

Betwixt

64

36

61

161

Betwixt

70

72

45

187

Expressive

100

29

105

234

Expressive

71

27

83

181

Totals

241

100

201

542

Totals

204

158

153

515

MKASrKK OF liFSFMIUAXOF OF FIliST COUSINS

l::

Temperament Reserved or Expressive. Cousins. All Types. Table LI. Male Cousins.

First Male Cousin

Reserved

Betwixt

Expressive

Totals 71-")

Reserved

334

196

2 1 5

Betwixt

196

■230

126

552

Expressive

215

126

230

571

Totals

745

552

571

L868

Table LII. Female Cousins.

First Female Cousin

Keserved

Betwixt

Expressive

Totals

Reserved

212

228

270

710

Betwixt

228

304

222

754

Expressive

270

222

496

988

Totals

710

754

988

2452

Table LII I. Male and Female Cousins.

Male Cousin

1 Reserved

Betwixt Expressive

Totals

a

Reserved

253

170

140

563

o O

<E

Betwixt

263

264

180

707

id

Expressive

382

134

373

889

Totals

898

568

693

2159

41

ETHEL M. ELDERTON

Temperament Sympathetic or Callous. Male Cousins.

Table LIV. Type A.

First Male Cousin

Table LV. Type B.

First Male Cousin

Symp.

Betwixt

Callous

Totals

o O

Symp.

Betwixt

Callous

Totals

Syrnp.

91

28

■2

121

Symp.

67

33

9

109

Betwixt

35

36

5

76

"3

3

Betwixt

24

49

5

78

Callous

31

15

2

48

o 0Q

Callous

6

7

10

23

Totals

157

79

9

245

Totals

97

89

24

210

Table LVI. Type C.

First Male Cousin

Symp.

Betwixt

Callous

Totals

Symp.

170

67

18

255

Betwixt

74

84

19

177

Callous

39

20

2

61

Totals

283

171

39

493

MEASURE OF RESEMBLANCE < >F MUST COUSINS

15

Temperament Sympathetic or Callous. Female Cousins.

Table LVII. Type D.

First Female Cousin

Symp.

Betwixt

Callous

Totals

Syrup.

133

59

11

203

Betwixt

102

61

8

171

Callous

30

12

2

44

Totals

265

132

21

418

Table LVIII. Typt K.

First Female Cousin

Symp.

Betwixt

Callous

Totals

1

Symp.

151

40

7

198

o

~

Betwixt

26

27

1

54

r*

5

s

Callous

15

7

7

29

xn

Totals

192

74

15

281

Table LIX. Type F.

First Female Cousin

Symp.

Betwixt

Callous

Totals

o O

Symp. ! 321

82

14

417

Betwixt

53

46

6

105

a

o

1

Callous

25

11

4

40

Totals

399

139

24

562

46

ETHEL M. ELDERTON

Temperament Sympathetic or Callous. Male and Female Cousins.

Table LX. Type G.

Male Cousin

Symp.

Betwixt

Callous

Totals

Symp.

257

113

-

412

Betwixt

85

87

25

197

Callous

28

9

2

39

Totals

370

209

69

648

Table LXI. Type H.

Male Cousin

Symp.

Betwixt

Callous

Totals

Symp.

205

91

15

311

"3

c

Betwixt

41

92

12

145

3

b

Ph

Callous

13

4

12

29

Totals

259

187

39

485

Table LXII. Type I.

Male Cousin

Table LXIII. Type K.

Male Cousin

Symp.

Betwixt

Callous

Totals

o O

f2

Symp.

Betwixt

Callous

Totals

.3

Symp.

293

66

43

402

Symp.

225

100

33

358

o O

Betwixt

50

53

16

119

Betwixt

70

67

9

146

Callous

11

8

3

22

Callous

14

5

1

20

Totals

354

127

62

543

Totals

309

172

43

524

MEASURE OF RESEMBLANCE OF FIRST COUSINS

Temperament Sympathetic oe Callous. Cousins. All Types. Table LXIV. Male Cousins.

First Male Cousin

17

Symp.

Betwixt

Callous

Tola],

Symp.

656

261

105

1022

Betwixt

261

338

71

670

Callous

105

71

28

204

Totals

1022 670

204

1896

Table LXV. Female Cousins.

First Female Cousin

Symp.

Betwixt

Callous

Totals

Symp.

1210

362

102

1674

Betwixt

362

268

45

675

Callous

102

45

26

173

Totals

1674

r,::,

173

2522

Table LXVI. Male and Female Cousins.

Male Cousin

Symp.

Betwixt

Callous

Totals

a

Symp.

980

370

133

1 183

o O

Betwixt

24G

299

02

607

3

Callous

66

26

18

110

Totals

1292

695

213

2200

48

ETHEL M. ELDERTON

Temperament Excitable or Calm. Male Cousins.

Table LXVII. Type A.

First Male Cousin

Table LXVIII. Type B.

First Male Cousin

Excit.

Betwixt

Calm

Totals

o O

"3 1

Excit.

Betwixt

Calm

Totals

o 1

Excit.

48

21

60

129

Excit.

40

18

42

100

Betwixt

21

68

41

130

Betwixt

18

90

34

142

o

1

Calm

60

41

126

227

Calm

42

34

94

170

Totals

129

130

227

486

Totals

100

142

170

412

Table LXIX. Type C.

First Male Cousin

Excit.

Betwixt

Calm

Totals

o O

1

Excit.

78

56

122

256

Betwixt

56

166

93

315

CO

Calm

122

93

224

439

Totals

256

315

439

1010

MEASURE OF RESEMBLANCE OF FIRST COUSINS

i:»

Temperament -Excitable ob Calm. Female Cousins.

Table LXX. Type D.

First Female Cousin

Excit.

Betwixt

Calm

Totals

=

6

Excit.

58

30

110

198

Betwixt

30

70

99

199

8

Calm

110

99

226

435

Totals

198

199

435

832

Table LXXI. Type E.

First Female Cousin

Excit.

Betwixt

Calm Totals

Excit.

70

37

73 180

Betwixt

37

36

47 120

Calm

73

47

82

202

Totals

180

120

202

502

Table LXXII. Type F.

First Female Cousin

Excit.

Betwixt

Calm

Totals

I

= fa

a I

Excit.

136

54

163

353

Betwixt

54

110

76

240

Calm

163

76

322

561

Totals

353

240

561

1154

50

ETHEL M. ELDERTON

Temperament Excitable or Calm. Male and Female Cousins.

Table LXXIII. Type G.

Male Cousin

Excit.

Betwixt

Calm

Totals

Excit.

45

39

81

165

•-

b

Betwixt

12

70

54

136

-

H

£

Calm

47

82

1G3

292

Totals

104

191

298

593

Table LXXIV. Type If.

Male Cousin

Excit.

Betwixt

Calm

Totals

Excit.

48

26

76

150

"53

o

Betwixt

31

87

66

184

~3

£

Calm

42

31

70

143

Totals

121

144

212

477

Table LXXV. Type I.

Excit.

Betwixt

Calm

Totals

Excit.

66

27

115

208

Betwixt

23

42

49

114

Calm

51

41

135

227

Totals

140

110

299

549

Table LXXVI. Type K.

Male Cousin

Exeit.

Betwixt

Calm

Totals

Excit.

44

21

87

152

Betwixt

18

65

32

115

Calm

60

50

152

262

Totals

122

136

271

529

MKASIIJK OF IIESKMBLANCE OK FIRST COI'SINS

51

Temperament Excitable oe Calm. Cousins. All Types. Table LXXVII. Male Cousins.

First Male Cousin

Excit.

Betwixt

Calm

Totals

a 1

Exoit.

166

95

221

485

Betwixt

95

324

168

587

c q I

Calm

•J24

168

444

836

Totals

is:,

587

836

1908

Table LX XVIII. Female Cousins.

First Female Cousin

Excit.

Betwixt

Calm

Totals

Excit.

264

121

346

731

Betwixt

121

216

222

559

Calm

346

222

630

1198

Totals

731

559

1198

2488

Table LXXIX. Male and Female Cousins.

Male Cousin

Excit.

Betwixt

Calm

Totals

_g

Excit.

203

113

359

675

3

o

Betwixt

84

264

201

549

£

Calm

200

204

520

924

Totals

487

581

1080

2148

52

ETHEL M. ELDERTON

PlGMENT-ATK >X CHARACTERS.

Table LXXX. Eye Colour, Female Cousins.

First Female Cousin

Tints

Pure Blue

Blue Orange

Pure Grey

Grey Orange

Hazel Green

Hazel Brown

Brown

Totals

c

Pure Blue

9

1-5

8

2-5

4-5

2-5

4-5

32-5

o

Blue Orange

1-5

6

7

9

3-5

3-5

8-5

39

(J

Pure Grey

8

7

4

5

1

2

1-5

28-5

2

Grey Orange

2-5

9

5

6

1

6

2

31-5

Ph

Hazel Green

4-5

3-5

1

1

1-5

4-5

3

19

"3

Hazel Brown

•2-5

3-5

2

6

4-5

8-5

5-5

325

tc

Brown

4-5

8-5

1-5

2

3

5-5

10

35

Totals

32-5

39

28-5

31-5

19

32-5

35

218

Table LXXXI. Hair Colour, Female Cousins.

First Female Cousin

Tints

Very Dark

Park Brown

Brown

Light Brown

Fair

Red

Totals

-3

Very Dark

9

11

3-5

11

1-5

36

o O

Dark Brown

11

13

12

10

9

1

56

•%

Brown

3-5

12

8

9-75

3-25

3

39-5

ti

Light Brown

11

10

9-75

16-5

9-25

1

57-5

a

O

Fair

1-5

9

3-25

9-25

1

24

Red

1

3

1

_

_

5

Totals

36

56

39-5

57-5

24

5

218

M.KASU11K OF RESKMBLANOF OF FIRST COUSINS

53

Pathological Characters.

Table LXXXII. Tuberculosis.

First Male Cousin

Tuberculous

Non-tuberoulous

Totals

Tuberculous Non-tuberculous

18 188

L88 2596

206 2784

Totals

206

2784

2990

Table LXXXII 1. Insanity*.

First Mala Cousin

Insane

Sane

Totals

Insane Sane

38 221

221 2510

259

2731

Totals

259

2731

2990

Table LXXXIV. Tuberculosis.

First Female Cousin

Tuberculous

Non-tuberculous

Totals

Tuberculous Non-tuberculous

20

185

185 2S52

205 3037

Totals

205

3037

3242

Table LXXXV. Insanity*.

First Female Cousin

.5

5 O

Insane

Sane

Totals

Insane Sane

6 124

124 2996

130 3120

0

Totals

130

3120

3250

Table LXXXVI. Tuberculosis.

Male Cousin

Tuberculous

Non-tuberculous

Totals

Tuberculous Non-tuberculous

20 237

148 2698

168

2935

Totals

257

2846

3103

Table LXXXVII. Insanity-

Male Cousin

Insane

Sane

Totals

Insane Sane

12 187

128 2791

140

2978

Totals 199 2919

3118

Includes marked neuroses, alcoholism and hysteria

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On tht

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resemblance of first cousins