°msm :m ULTURU PUBLICATION NO. 556 ISSUED APRIL 1937 TECHNICAL BULLETIN NO. 7 FIRST PRINTING DOMINION OF CANADA, DEPARTMENT OF AGRICULTURE PEACE-ATHABASKA WEEDS A RECONNAISSANCE APPRAISAL HERBERT GROH Botanist l + l Agriculture Canada Canadian Agriculture Library Bibliotheque canadienne de I'agriculture Ottawa K1 A 0C5 DIVISION OF BOTANY EXPERIMENTAL FARMS BRANCH 630.4 C212 P556 1937 c3 Published by authority of the Hon. JAMES G. GARDINER. Minister of Agriculture Ottawa, Canada PEACE-ATHABASKA WEEDS A RECONNAISSANCE APPRAISAL A brief visit in October, 1929, to Beaverlodge in the heart of the Peace River country,* and three weeks more in the late summer of 1934 have afforded opportunity to become somewhat acquainted with the region and with its weed flora. A few days in 1935 were also given to examination of the Athabaska sector so far only traversed by rail. The 1934 trip, the second in a projected quinquennial survey designed to preserve for at least one such area in Canada a more or less adequate weed history, was fortunate in the facilities provided for penetrating to almost all settled parts. This was due largely to the aid of Messrs. W. D. Albright, J. Travis and W. J. Thomson, of the Dominion, the British Columbia and the Alberta departments of agriculture respectively, and their interest and effort are deeply appreciated. The limited time allowed was conserved for study of the vegetation of settlement. Only incidentally in travel between prairies, was other plant cover much noticed. The native flora has always received major attention, first of the earlier explorers who would find little else, and later of botanists more intent upon phytogeographical and ecological problems than upon the influx of alien species. Recent botanists have been chiefly attracted to the still scarcely disturbed frontiers and beyond; whereas the primary con- cern of the agriculturist is to know the adventive plants, the manner of their advent and spread, and the prospect of retarding their increase in these new farming sections. In the lists so far published it is amazing what little men- tion there has been of weeds already becoming well naturalized. The fruits of past botanical exploration are not, however, to be despised, even though they leave so much need for further weed survey. That same virgin flora is temporarily, and many elements of it long continue to be, the cover to be ousted by plants of cultivation and their attendant weeds; thus any survey is a contribution to weed survey. It is particularly opportune that a recent work on the botany of the Peace and Liard River regions,f and other published contributions from these hinterlands of Canada, by Dr. Hugh M. Raup, have brought together the valuable results of his own and earlier explorations in and beyond this area. This augments considerably the all too meagre results of the present trips; and what is of equal importance, it provides the support of authenticated records for many observations, which in a weed reconnaissance too hurried for much collecting of specimens, remain doubtful or with incomplete identification. DEFINITION AND SCOPE Some definition of terms employed will help to delimit and clarify the subject in hand preparatory to as brief a treatment of the materials under review as will suffice. Peace-Athabaska. — Geographically this term indicates the extensive drainage basins of the Peace and Athabaska rivers; agriculturally it is restricted to a part of the upper or southern reaches of these rivers and their tributaries. Here the name is made to apply to the fairly homogeneous territory stretching *Groh, Herbert. Grande Prairie weeds. Dorn. Can. Dept. of Agr. Pamph. No. 117 New Series. 1930. fRaup, Hugh M. Phytogeograpkic studies in the Peace and Upper Liard River regions, Canada. Contr. from the Arnold Arboretum of Harvard Univ. VI. 1934. 32987—2 from the town of A bh aba ska on the east, to just beyond Montney, north of Fort St. John; and from the town of Peace River on the north, to somewhat short of Edmonton. There remain beyond these bounds settlements where plants can behave as weeds, as at Fort Vermilion; 'and there are wooded tracts alternating with the prairies within where the flora is but little weedy in character. Phytogeographically the area is reasonably common ground for study, and it comprises -approximately what has been embraced in these weed surveys. Weeds. — A weed has been defined as " a plant out of place," meaning of course, that in some' person's estimation it is out of place. From the stand- point of the plant it is always in place when it can gain a foothold and meet the competition of other plants, adapt itself to the environment, and enjoy sufficient respite from human interference. As man sees it, some plants are always and only weeds, others are occasionally or in some circumstances weedy, and some, even of the most useful crop plants are weedy when they escape from cultivation or persist when their place is to be devoted to other uses. Some plants, moreover, are weeds not so much in an agricultural sense, as by reason of being directly harmful or poisonous to man or beast, or as being unsightly or in any way an occasion of trouble or expense. For present purposes a weed may be considered to be any native plant persisting, or any adventive plant encroaching in man's domain, in ways inimical to his interests. Reconnaissance. — A reconnaissance survey is not a census in any degree. The enumeration of what is present on a quadrat or other limited area may be feasible, and has its uses for ecological and other special purposes, but is out of the question here. Only methods can apply which will gather a broad sweep of the information sought in a modicum of time. It is not botanical survey in the stricter srnso. Weed survey, which is all this can claim to be, relies as far as possible on previous botanica'l surveys, dispensing with much of the collecting of common and obvious species in favour of simple recording. The weed surveyor, be it understood, does not undervalue accuracy; but in much of his particular field the risks of inaccuracy are neither great nor serious, since weeds in general are the commonest and most readily distinguished of plants. Collecting is done chiefly in support of records of doubtfully identified, scarce, or otherwise notable plants of weedy character. The surveys here presented are, as already intimated, the first of a series intended to follow developments as they unfold in a new field such as this. It is true that pioneer history has already been left behind for much of even this new north; but the present frontier settlements still present about the same setting, and beyond these may still be found the indigenous flora of the country, unmodified except by those ubiquitous vagrants that seem to dog the steps of even the trail-blazer — wild buckwheat, lamb's quarters, and other freely dis- persed annuals. The principal reconnaissance in 1934 extended through the three weeks commencing August 28. There was opportunity for fairly extensive, though never exhaustive, field work in each of the several prairies which go to make up the bulk of the developed lands, i.e. at High Prairie, in the McLennan-Falher district, in the extensive Peace River-Fairview-Dunvegan belt north of the river, at Beaverlodge and Hythe, at Pouce Coupe and Dawson Creek, and in the Fort St. John district, at that town and about Baldonnel, Rose Prairie, and for some distance north of Montney, as well as at Taylor Flat overlooking the river. Briefer forays were possible at many intermediate points. Moreover, useful observations of the more conspicuous and familiar weeds were made from the moving train or automobile; especially during stops, which allowed a certain amount of scrutiny of the environs of stations along the way or of highways and towns, excellent short lists could be secured. Frequently instead of a general survey, some special environment as a crop or a weedy summer-fallow, was the basis for a list. Altogether 110 lists of varying length and character, and embracing among them practically all plant species distinguished in the course of that year's trip, entered into the study. The Athabaska observations to and from the Peace, were reinforced in 1935 by trips to Barrhead and Athabaska, on which an excellent idea of inter- vening country was secured. These, together with the 1929 surveys, bring the total of lists to 174. Appraisal. — While determining occurrence, surely a useful end in itself, weed survey goes a step farther, and availing itself of the constant repetition of lists, seeks to establish some measure of abundance, if not actual at least relative. In place of dependence on the indefinite " common/' " scarce," " rare/' etc., of popular parlance, there is possible a sort of index of frequency. Any particular weed will be found to have appeared in one, ten, or perhaps ninety per cent of the lists, indicating with some precision whether it is rare, occasional, or quite general in occurrence. Position, high or low in the lists, in a rough way recording prompt or merely chance observation of a plant's presence, can be some guide as well, but is a refinement of method probably unwarranted in view of other refinements lacking. Such appraisal as is secured must always be accepted with due reservations. To illustrate: common dandelion, at the height of early bloom, would be more heavily recorded than at another season when chiefly the prostrate foliage is in evidence. A little later when both fruit and flowers are to be seen, it might also be found that what appeared to be common dandelion is that and red-seeded dandelion together, neither one of which separately would rank so high. A further consideration would be that dandelion, while an undoubted nuisance in its own way, is not often the occasion of great losses in crops. The prevalence found, even supposing that all weeds could be equally conspicuous, in season, and uniformly distributed, is not, either, always the prime factor in weediness. Fireweed and yarrow, native plants unsurpassed for prevalence, are after all less dangerous than many new-comers, still sparingly recorded but ready to forge to a position of real menace. The index of prevalence, while not an absolute measure of weediness or even of abundance, is at least something tangible and an approximation to the truth. Other attributes of plants which, together with abundance, may con- stitute them weedy, are difficult to assess in other than general terms based upon practical judgment. A prevalence percentage, however derived, is highly desir- able information which the botanical manuals, when indicating range, can give scant light upon; and what they give can be valid for the time only, since weeds are ever on the march, and subject also to changes affecting agriculture itself. General Considerations. — Those plants observed which are in no way to be regarded as weeds, are excluded from notice. Weedy species are listed in two tables; first: those indigenous which tend to persist as weeds, or for other reasons belong in that category; second: those adventive or escaped from cultivation, and becoming naturalized and more or less weedy. In each case they are arranged in a descending order of prevalence; and for each of eight meridians from west to east a frequency percentage figure is given, or locality indicated for those infrequent. Discussion of weeds follows in the sequence of the current botanical treat- ments. It is for the greater part an evaluation of the plant as a Peace-Athabaska weed, actual or potential. It is not to any great extent descriptive of the weed, nor of its control, since these are functions of the existing weed manuals. Nomenclature has quite closely followed that adopted by Raup in his works on the vegetation of this part of Canada. As a second report, though on a broader field, this treatment enlarges upon Pamph. No. 117, New Series, Grande Prairie Weeds, and in many cases repeats 6 the language of that report. Being essentially but another interim report, based on all too slight observation, it still anticipates the more detailed and critical investigation which should be possible at the third step in the project. The several meridians may be designated as: 113 — Athabaska district, east. 114 — Athabaska district, west. 115 — Lesser Slave Lake district. 116 — High Prairie — McLennan district. 117 — Father and Peace River districts. 118 — Grande Prairie, east — Spirit River and Fairview districts. 119 — Grande Prairie district, west. 120 — Pouce Coupe and Fort St. John districts. lO »o OS o CO "* <* CO CO OJ CO 00 os 10 OS CM 00 os os 00 OS 0 00 00 CO CO CO »o CO t^. t-» "*< CO 0 0 •* > eo U) c3 -»j c CO o u co Ph >s o c co S3 cr CO o 01 10 00 OS eo _l t^ 00 TJH CO rH "* 00 CO CO "* CO 0 CO «5 "* CO lO CO CO CO r-H 00 00 ;# b- OS 00 00 t~ co t>. CO »o CO CO CO OS o CO CO CO Tt< CO »o OS OS 00 7— 1 CO OS . OS 00 00 t^ co 00 CO !>. us »o t-~ CO CO -* !>. •* CO J— 1 CO CO OS t^ 1— < >o 00 00 O -* 00 00 CO CO "5 t^ *o 00 "* CO Ph co c CO O O (H CO o .5 c3 dl O w* CO »hK *-• S3 co T3 O J5» CO as* O Ph o ■«* co • « .3-3 i^s ^ IS 53 s ° rScQ^i-H t3 eo O 00 13 CO1-1 0) ^ C3 > C3 cj S3 co-2 fl rj 00 T3S 2^ ^? » 5-53 •si !^ ^ g to Sag? o ^^ IS1 o s T3hH* |2 : *« p. so«2 si co s^ co 1 .3 o s S'S«' Ph io a x +S CO a d c3 w Ph 2 2§>;3^ ill* 1^ CO CO fc. C3 co k'S'd Ph M O O.L, 3 'S'a go ,~; P4 JJ3 32987-3 8 l— 1 CO cnj 00 co CO 0 £2 CD 00 00 00 - CO o CO CO CO a; Ph B cw a CD s s .2 J . h sM s § e s Si « u g~. e3 S feo CO cy 6 o 00 3 a GO 8'E O o m ** 'tu ?? e. -^ el V 3 S bo s- o 03 2 ^ ©,W <=> r- «J o e GO H i o e £ i bC-S !» O S ^ o~ o-£ S 03 %£ °^ §U « S? o > £ 03 oCQ c3 la gal O Ph 00 > C 2 § flJlJ -SOS 5^ in R O -2, 2 K _ C3 03 -43 43 bD oo W5 CO CO s oo CO 00 Id 00 CO a CO s o 00 • CO 00 O 1— 1 CO »o > CO 1—1 l> 00 3 42 'C -4J 02 •1-4 0) •l-H .§ e3 3 42 — +3 03 X) i— I -4-3 3 -4-> •1-4 00 o3 _ -u 03 +3 • M "o a 3 "■+3 o3 3 +3 *s 43 -4-3 3 O 43 bO 3 O 43 T3 O -* h s o S 3 >> o bfl a> •+» -'3 3. >> 03 - 0> O "3 M _ O 3 a 3 t— 1 -4-3 -4^> cS o 0Q o #3 N CO N M H "* CO o 00 t-~ > 3 :*!! 03 •*» '^ ;? S§ g ° S > •— > to 3 •§88 S e K 8.-2 8 .MM ►*5 ?3 Ji,42 «0 p-* Fh 2^ S+3 s O o3 COpH eoPn c3 > S3 h 0 e 3 g -» tn-5 Osi_J Si 03 ^-3^13 H3 o3 s M 3 •2 CJO T* o3, Si 03 e o „ 5 o o> e o •i.sss ^a > e 03 >i cd c 3 cy W) 5 CD -2 U •*H Q. CI o 00 J? 03 0 a CD B h 3 J -♦-> 03 O GO II CO HH S CD s T3 CD ^H ^C *» 13 Ph tc 03 OO CO a> •p-i > '3 a ^ bD £ o PQ CD Cd o " o 2W 03 CJ PH a, 3 O U CD CD O Ph bo -3 o TQ CD ss CD PQ 00 .2 ft ■3 § 2oP4 ill 3 Q <5 CD U PH CD « CD o CD 03 cD-r 1°* P, cj;g H ^ o X CD 03 o *o s m ^ CD J & • <■"-< o -^ •S ft M ££§ 1°8 • ^) o *- _! S^^ 2 if III 0 3 U«),g oPh O % K i- *~ ^ B W CO £ CD _j Ph^ ■§^§ S bC g S O o •So S p^ - 1/J •« « S co CD ^13^ 03 s CD ■» si CD u < 03 03 C- 03 O O CD^? ■43 !> S 8 fl S a Si o Ph CD v Ph^ «5»-SS §|W.g CD ^"^ <» ^^ CD^ CD § Co • ^j O c332 fe fn^ cdO CO Ph a ■T3 « CD S CD | IS s 11 a d M j£ to 03 o3 +3 OS -*3 "ts-3 ^ 2* O i-i 03 ,±4 8 £> o3 XJ CQ o3 O c3 © Xi pq p © . p © .i £.2 8 §.2-q^o >"^ O o3 . © Eg 3* 5.9 s a © .44--V o8.8"S * p © o3 bc-ft c — e3 ©, Ph © £ p p © hh* o a o3 5f © © o3 © Ph 03 u si © Sis O © a> £ °> s^ 2 PP W © .£«° 'p* ©^d^Q Ph © M .fc © Ph" o3 © P*< o o3 © Ph Pm -sift — ."S ft »H . g M*,-,,1 2 X ODJ 03 © © bfi o © > o3 © © © bO _o © © PQ _ +» © p P O Is pq Ph © > cPh o, P - o, >>Ph o3 m ° * o3U b =3 88' © Ph pq 03 H dp-J8^ o p^' o °3 PmH © •5 a ^ *-" 03 © 3 _ Ph £ © o3° 2 p 03 t— , P tH P 03 -u 03 O ' P4 gpq ^ .a o3 u Pm o ©_ a •31 § Ph§6 o «« 2 Ph < Ph pin Ph O a © eo O S ~= p s © 3 03 rp © ?5 O q C o 03 i — > 8 • XJ o p &5^ o § « 5 s O a © o -03^ Ph © o p o © 'pT3 «<-c t— ■ • i-H -CO ^<{ © J~ pq O co c3 © 03 ti S » is ©^ P^ Ph §5 C •S ^© Jb4 o gg£ -d s^ O o 03^ 03 P •3 2 p-S +3"43 o3{/5 &H 12 a M 09 oi •*j Si m a M -c CO o3 JO a < m s o a O GQ © o 03 eS 0) ^ r*( ^ w . 00 . CO ei -u o3 +a 03 ,n CG^ oon -u S3 ^ ■5 «.s << • 3 o tf.gSg«:gS§-3 c3 o3 g c3.j §"■*■ Ah CD • i-H « CD o o3 O Ah CD > c3 CD fa CD > M O •"■* ci • 3 Ah £ 03 CD Ah 0 > ^p "o. CO CD •FH 03 Ah 3 C5 to CD > 3 3 Q 03 Ah cd bfl 13 O 11 CD > o3 CD M CD ■IS -2 bfl CD bfl -d "p CD > o3 o A5 CD bfl o > 03 CD A3 CD bfl CD O ■e » >> > JU 03 A3 CD bfl -a CD O £^ >> > W « •^ CD A3 o bfl 13 JO CD CD CD bfl 13 O TJ CD CD -2 03 03 — / H Ah^ CO 03 H _0 +j hj CD 2/£c>£oAh£3ocdg3 Jh^ O 03 OT »ipH J2 "t^ r3 >»o 03^ CO o fa a 3 o O CD O 3 O Ah Ah Ah Ph Ah Ah Ah CD Ah "el CD C CD o CD 15 Pi O #3 co CD a 03 S^H e<3 bfl s o3 jj, 030 M O <» Pi Cs CD •«■» w 13 O O s Pi o 13 o > I s o .s • 03 : > '.* : o ■Ah I o I -a ^5 .2 CD I > So is co 3 ® . < 2 CD 3 aAn Ah a _ p. 13 o CD«f^ B"5 3 ^ bfl CD *» o^ «S Ah g ^ § CD «S O g ' co 03 p S^ CO o o CD^ o 2 o is CD. 2 o 03 %> CD CD~ CD •—> O P CO 3 0 fa S K a •«» ?> 11 J p CO a ^ CD a co '5 2 C3-5 ^1^- 3 P H M o gggl- >>.2.S CD 3 P1^ bO^ > o < w • CD-p 00 ^ « ^ o3 § e \o o3 R o :0 :«8 . cs . -o CD'** S "5 3 S> 31 13 oj .44 m c3 . -°-s 03.14 ,d73 S8 m n a -u X> oa 73 o © 03 | 73 © ^4 J .111 £ d goS to-3 o3 © © © 2:M © ^" Ph 03 © > © © o3 © © © Eh S-4 © © o 03 © Ph © « « 03 © Ph cj © Ph © © eg c3 >-> u OPh a? o3 . - bfl^ 5 . o P £ © © > ■^ © © bfl 73 © © pq _© "3 73 en a 5 © © bO 73 O © > 03 © PQ © bO jo © > 03 © pq © bfl 73 O i— < © 03 © pq © bfl 73 O 13 © > o3 © pq © bfl 73 © > 03 © pq .a 03 Ph o Ph" © S3 ©J2 © Ph J3 o *3 03 H J2 -03 -2 3 o &Ph >> ft 03 rd 03 ^>1»ft co^ o Ph Ph Ph Ph pq Ph < < < < < < Ph < Ph Oh © c M © ll Ph-2 03 « £ <1 to .3 o3 »s» bfl o o> O a I I pq S OPh © bO ^ ^ gPn S 03 3 ® § bfl"S^ 2 H 02 O 3 c 13 St3 3 5» S 05 bflg © ?73^2 E^ ^ a 8 •^ Q, © «2^ ^ §^73 ._ e © ?3pH -d © P! - © © OT3 . ^ 03-^ «o w 3 ■s-sp ^§^g73g ©'t^ O ^ 3 G © © 8 a Q Ph a § «o S2 ^5 3 ^ g'bc -^"S'8 J3 Shh S,g © a^ a TZ.v^o s ©^o^73 O H pq H a ft S w a to 8 53 ^ o 3 © ca 2Ph S Ph 8.S Ph J > S.2 o3 *■» © © <» to T ft o ►> co O ^1 ."S_ c3^ © ^1 h4 "a 2 3 . . a ^ 4-i a 0 © =s bfl^s -a .3 3 0 0 h,0 n 03 © a i-i aj2 3 u 0 on O -1 O •rH pq a 05 o3 •O 373 o a © e ? % -a • © © 14 c8 M n bS *a ^3 tf) & ^5 © I co 0 o o co O Ph CO o a H Q «} co Q to < O & h- I H co t— I CO H Ph co H Ph CO P o o Q ^ ■ © M 03 +^ d CD © © 3 © © >> k Ph1 Ph1 © © © o eft eft © © Ph Ph (-1 © © c3 ^s w Ph J5 o3 © be o © © PQ -d a eft © •FH H Hi , hj ■^ , © J*J ,~^ © .44 ^^ © .J*.-* >+i d e >~£ C d > "£ a c eft w *£J 3 . t>- t» t^ »o ■* o> 1—1 ■«* •* r* CO CO H Z W > o CD bC S3 CD o Sh Ph e3 3 o Ph OS ■* CO lO OS lO "■*! CO o us o CO w 00 CO (M lO OS f^. 00 t^ CO CO »c ■* OS C5 o CO ^H . cc . CO . CD > : . - — ' CD - Q • ■*a • » '•2 hi : . e . CJ C3 . • k C3 - 1 S 3 • ^ .-is ?^o *h co _, t^ ^ to S PQ & 73 CD T3 CD Hi 3 CO Sh • CO ^W S oo ^a _S +3'*-! •s» « <» i-j r£ O C» ^ ^CG0 CD «0 C/J > a—, ^ ^ .^^ S^ !§ CO PJ log 8 S3 o +3 CD ^3 W) . S3 11 h4^ 1.2 CD S3 3 O O) Pi CQ S3 ft O ft T3 Q o Q CO a. o CX2 S a "S ^"^ co CD CO CO Ph P4 cu Fh o ^ « CD R 15 ^ « £.33 u (1> f 32987—4 16 M © 3 Q H W £ H PQ O H H Ph I— I > P H ►q pq iS O — . © 03 >> o3 ^3 .2 © U O © © J3 03 >-. Ph A 5 © a o Q © •l-H "3 © to -3 jo © > 03 © PQ © © ©'-' 03 © Ph 4J fljCJ © ©OS ST © pq © "3,° pqZ <"S 3 O Ph a § © © > PS oS © Ph 03 © T3 JO © > 03 © pq 5 »3 oPh t, * oH Ph 9 Q << Ph pq Ph << Ph o +3 £h3 a3-« 5r*^ co © SPh.H >' W O P4 © -; Si S5 03 rrt o3 £ 3* £ «! <1 2 Eh CQPh 03 *> JH 'o3 .T5 03 ccPm Teepee Creek Grande Prairie Hythe Beaverlodge Hythe 03 bfi J3 Th 03 > 03 PQ 03 M H3 O Th 03 > 03 03 pq 03 bfl O »-M (H 03 > 03 03 PQ Dawson Taylor Flat Pouce Coupe Dawson Creek Fort St. John Baldonnel 03 >> w o l-S m Eh O PH Fort St. John Beaverlodge Montney Baldonnel Dawson Creek Pouce Coupe 03 •H d Ph 03^3 03 U o o Ch' PC < < Cm < < < Cm Cm o <3 Sh o3 O 03 S «B e8* O O O Q3 '«■» ^ < u a 03 03 o3 . m Ho o c = b 2 o3 Cs c3 S 03 8 03 S osO 5 >>§ 03 w 03^ «o 03 a> w hi 2 co « 5 s 9 h • HOD 03(V w. m O S>^H O x o 03< g 03 © X fe ^x «W" ° 2 S 4 ^T3 43^ § .Hi 03 ^S H © rs .2 03 ^-^ «^-2 H ^ S5 o »Ph k - ^1 3& 03 "12 03 CJ 03 « o . <^ , e ^ t-< « N S r* §CQ 03 -JS 03 S hQ a© 03^ -i Q o3 o P g ■a <>a O H Ch Ph PQ W 32987— 4i 18 d OQ d •s X\ 3 o O I* Q W H £ H PQ O H H Ph H > Q H tUD d Ph a CD Eh f3 K c3 m Ph a> cd cd hi) h() hil Td T3 T3 o O O M h h o CD cd > > > a 03 d CD a) a> pq « W o a> hi) hi) T3 T3 O O * (-4 cd a > > a a CD CD Ph « > a F c pq M d Ph cd > o o d a Ph o o cd hn hi) hi) Ti T3 T3 0 o O (-. g — o a> cd ► > > d d a 0) 0) 8 m « « CD Q Ph Ph t-i o • • •«* SO spills a ■s§3 a d o OD j> a> CO. e ° S § c3 « |rt.« g ^ to o o ^ e S ^ "S ■ ^^ d o Tl C3 - PC H 73 CD O O 8 C ^ •, <» a © CO. f-< ta S 2 ©> — 1 co oj s^ *3 Ph £ gc02 Si a c3 Cs >1 d .'o o o d d a> o d c3,C Q cj ~» o P pq IK a 5.9. a c T5 o a c3 o w CO H tO (D O 3"^ •^» CD 2 » d 3 s d co CO Q co £t3 d ££ II 8.8 v. a Ph O O PQ o3 oS 03 M M M 03 03 w a a a Xi Xi Xi a o3 a xi xi xi © >> o 19 03 M 03 . o3 +=> .a* "o3 0> bO T3 O © 03 bfl _o > o3 © PQ o3r^ Pm o t-i PQ Pm Pm Ph C £ £ * bC oe=i C+s so O o3 g o'C e o o3 ' en I 01 eT3 ca © I 03 Si «H 03 S 2 « CO 03 S» CO © O CO o go I frhilni . (=1 g 03 ■»■» Q W