af (thus 1985
Pp werotocts

_ | 4 international journal to expedite botanical and phytoecological publication

Vol. 538 August 1985 ~ No. 1
CONTENTS

JONES, A.G., An annotated catalogue of type speci-
mens in the University of Illinois herbarium (ILL)
Bee iperdceac, except. PENeTOMia s... 33.85.05. bs dbo nas eei cd ieect. sith, 1

Published by Harold N. Moldenke and Alma L. Moldenke

590 Hemlock Avenue N.W.
Corvallis, Oregon 97330
U.S.A.

Price of this number $3.00; for this volume $15.00 in advance or $16.00
after close of the volume; $5.00 extra to all foreign addresses and domestic
dealers; 512 pages constitute a complete volume; claims for numbers lost
in the mails must be made immediately after receipt of the next following
number for free replacement; back volume prices apply if pent is
received after a volume is closed.

eee

ae 3

es
+o

fe
bag

iene.

te
~

ar:

oe
a

AN ANNOTATED CATALOGUE OF TYPE SPECIMENS IN THE UNIVERSITY OF
ILLINOIS HERBARIUM (ILL)-- 1. PIPERACEAE, EXCEPT PEPEROMIA

Almut G. Jones

Plant Biology, University of Illinois, 289 Morrill Hall,
505 S. Goodwin, Urbana, IL 61801.

INTRODUCTION

Type specimens in the University of Illinois Herbarium would be
of a comparatively modest number, were it not for a concentration
in those genera that were studied and revised by the prolific namers
of plants, William Trelease and Truman G. Yuncker. By far the
largest holdings of type material are to be found in the Piperaceae.

Trelease, Professor of Botany at the University of Illinois
from 1912 to 1926, had become interested in Piperaceae after the
death of Casimir de Candolle in 1917. Aside from many papers and
contributions describing new species, he authored treatments of this
family for Cuba (1926), Costa Rica (1929), Panama (1927, 1938,
1940), and Peru (1936). Notable among his taxonomic revisions is
that of the genus Ottonia (1935). Trelease continued his studies in
Piperaceae until his death in 1945, leaving a considerable amount of
unpublished material.

Yuncker, who had previously collaborated with Trelease and
published a number of his "new species” [e.g., in contributions to
the Flora of Honduras (Yuncker 1938, 1940)], agreed to complete at
least some of the unfinished work and was given an appointment as a
visiting research professor at the University of Illinois for the
academic year 1946/1947. During this time he completed the two-
volume work The Piperaceae of northern South America (Trelease &
Yuncker 1950). Yuncker continued his work in the Piperaceae,
contributing, among others, treatments of this family for Panama
(1950), Trinidad and Tobago (1953), Argentina, Bolivia and Chile
(1955), Suriname (1957), Jamaica (1960), and Brazil (1972, 1973).

The following annotated catalogue includes all names (basio-
nyms) of members of Piperaceae, for which type material (including
paratypes) is located at ILL, except Peperomia, which will be
published separately. Generic names involved are: Anderssoniopiper
Trel., Artanthe Miq., Enckea Kunth, Ottonia Sprengel, Peltobryon
Klotzsch ex Miq., Piper L., Pothomorphe Miq., Sarcorhachis Trel.,
Schillera Kunth, and Trianaeopiper Trel.

Whenever necessary, as well as feasible, I have designated
lectotypes. The entries concerned are marked with an asterisk in

]

2 PHYTOLOGIA Vol. 58, No.1

front of the name, including lectotypes designated by other authors
but not previously published as such. There are 50 such entries.
The status of each specimen was determined or verified by consulting
the original description. Unless indicated otherwise, the original
spelling and endings have been retained.

The list comprises 935 entries. Thirty-one of the marked
lectotypes are at ILL. Some of them actually may be holotypes. In
addition, the holotypes of 205 of the names listed, plus isotypes or
isolectotypes of 577 names are on deposit at ILL. Fragments of
types are cited for 27, and syntypes for 10 entries. For seven of
the names, for which no types were designated in the original
descriptions, I have cited authentic specimens filed at ILL that may
be, in part, duplicate types, or that could be utilized for
lectotypification. A total of 436 collections in 236 entries are
cited in the category of paratype.

In order to increase the information given in this catalogue
and thereby enhance its usefulness, I have cited the place of
original publication for each name, and if the holotype or lectotype
is not at ILL, I have also, whenever available, given the
conventional abbreviation (cf. Holmgren et al. 1981) of the
herbarium in which the type is to be found. However, locations for
isotypes and paratypes in herbaria other than ILL are not included.

Because my knowledge of Piperaceae is limited and I have no
field experience with the group, I have not made any taxonomic
decisions. Whenever available, however, I have added the
dispositions given in the literature. This information may be
incomplete.

The names are listed alphabetically, yeneric names only once,
on a separate line. Authorities for all names entered are cited,
but in the annotations, they are not repeated after the first
mention of a name. They are abbreviated in accordance with the
Gardens, Kew, in 1980. Literature is cited in the format of
Botanico-Periodico-Huntianun,

Except for LECTOTYPE, which is spelled out for emphasis, the
kind of type is generally abbreviated: HT = holotype; IT = isotype;
ILT = isolectotype; ST = syntype; and PT = paratype. For the latter
category, only names of collectors and the collection numbers are
given; locality information is omitted.

A typical entry has the following format: (1) specific epithet
and authority (2) place and date of publication; (3) type locality,
elevation, date of collection; (4) underlined: name of collector(s)
and collection number; (5) in parentheses: location of HT or LT and
type specimen(s) held at ILL; (7) in brackets: disposition of name
by other authors.

1985 Jones, Annotated catalogue of type specimens 3

ANNOTATED CATALOGUE OF TYPE SPECIMENS

Anderssoniopiper Trel.

panamense Trel., Proc. Amer. Philos. Soc. 73: 329, pl. b. 1934.
TYPE: "Panama". April 1852. Andersson s.n. (HT: S; fragment
of type at ILL).-- [This is the type species of
Anderssoniopiper, loc. cit. = Macropiper latifolium (L.f.)
Miq. (ef. Yuncker, 1955a, who pointed out that the type came
from a plant native in Tahiti, and fide Smith, 1975, 1981) ]

Artanthe Miq.
adenophylla Miq., Linnaea 20: 170. 1847. TYPE: Brazil, "In

sylvis umbrosis ad Coralfalga, m. Febr." Pohl 5026 (HT: W;
IT: ILL).-- (= Piper divaricatum Meyer (fide Yuncker, 1972)]

concinna Miq., Linnaea 20: 151. 1847. TYPE: Brazil, “Padre
Correa in umbrosis sylvis." Pohl 5032 (HT: W; IT:
ILL).-- This is also the type mor Piper concinnum (Miq.) C.DC.
in DC., 1869 non Haworth (1821), and the type of; Ps
concinnatoris Yuncker, Bol. Inst. Bot. (S30 Paulo) 32 33).
1966, a renaming of the above under Piper.-- ae latter name
was adopted by Yuncker, 1972]

fuscescens Miq. forma pilosior Miq. in Martius, Fl. Bras. 4(1):
38. 1852. TYPE: Brazil, Santa Cruz do Terza. Pohl 3465 (HT:
W; IT: ILL).-- This is also a ST of Piper estrellense C.DC.
in DC., Prodromus 16(1): 272. 1869, a renaming of A.
fuscescens Miq. under Piper, and an IT of P. estrellense
(var.] 8 pilosius (Miq.) C.DC. in DC., loc.*cit.-- [= P
tectoniifolium Kunth (fide Yuncker, 1972) ]

martinicae Miq., Syst. Piper., 413. 1843. TYPE: Martinique.
Sieber 254 (HT: B?; IT: ILL).-- [= Sarcorhachis incurva
(Sieber ex Schultes) Trel. Note that the name Piper incurvum
Sieber ex Schultes (1822) and the generic name Sarcorhachis
Trel. (1927) are typified by the same collection (cf. Howard,
1973) ]

opulifolia Miq., Linnaea 20: 160. 1847. TYPE: Brazil, San
Izidro. Pohl 1572, 5038 [Syntypes, probably to be
lectotypified: W; probable ST (Pohl s.n., obtained from W) at
ILL].-- (= P. glabratum Kunth (fide Yuncker, 1972)]

polyneura Miq., Syst. Piper. 504. 1844. TYPE: Peru, Cochero.
Poeppig [HT: B; authentic specimen obtained from W, perhaps
an IT, at ILL: Poeppig 1463 (no number cited in the
protologue)].-- {= P. indecorum Kunth (fide Trel. in
Macbr., 1936) ]

4 Po Hyde HOt, OG A Vol. 58, No. 1

schottii Miq., Linnaea 20: 157. 1847. TYPE: Brazil. Pohl 1406,
2006 = syntypes, probably to be lectotypified (ST, received
from W, at ILL: Pohl 1406).-- [= P. schottii (Miq.) C.DC. in
DC. (fide Yuncker, 1972) ]

Enckea Kunth

amalago (L.) Griseb. var. variifolia Griseb., Fl. Brit. West
Ind., 169. 1859. TYPE: Jamaica. Wullschlaegel 1030 (HT: W?;
fragments of type at ILL).-- [= P. amalago L. var. variifolia
(Griseb.) Fawcett & Rendle (1914); this variety recognized by
Yuncker, 1960 and Adams & Collaborators, 1972]

sieberi Miq., Syst. Piper. 358. 1844. TYPE: Trinidad, leg.
Sieber-- several syntypes, probably to be lectotypified (ST
at ILL: Sieber 224).-- [= P. medium Jacq. (fide Yuncker, 1950
and 1957); = P. amalago (fide Howard, 1973) J

Ottonia Sprengel

* aguadana Trel., Proc. Amer. Philos. Soc. 75: 696. 1935. TYPE:
Brazil, Minas Gerais, Fazenda de Aguada. El. 700 m. Mexia
5178 (specimen marked "Type" by Trelease = LECTOTYPE
designated herein: ILL).-- [= 0. propingqua Kunth (fide
Yuncker, 1973)]

apodostachya Trel., Proc. Amer. Philos. Soc. 75: 697. 1935.
TYPE: Brazil, Sta. Catarina, Jaragua. 11 Oct. 1929. Hoehne
24388 [HT: SP (fide Yuncker); IT (2 sheets): ILL].--

[= 0. martiana Miq. var. martiana (fide Yuncker, 1973) ]

* apodostachya var. vidali Trel., Proc. Amer. Philos. Soc. 75:
697. 1935. TYPE: Brazil, Espfritu Santo. 17 Oct. 1924.
Vidal 138 (HT, or LECTOTYPE designated herein: ILL).-- [= O.
martiana var. martiana (fide Yuncker, 1973) ]

armondi Trel., Proc. Amer. Philos. Soc. 75: 697. 1935. TYPE:
Brazil, Rio de Janeiro, Carmo. Armond 156 (HT: R; IT:
ILL).-- [= 0. anisum Sprengel (fide Yuncker, 1973) ]

colliculorum Trel., Proc. Amer. Philos. Soc. 75: 699. 1935.
TYPE: Brazil, Morro de Cavalao, Schwacke 28315 [HT: R (fide
Yuncker, 1973); IT: ILL; PT at ILL: Hoehne 24733].-- [This
species recognized by Yuncker, 1973]

edwalliiiTrel., Proc. ‘Amer. Philos. “Soc? °75: ,702%. 8935. « STXYPE<
Brazil, Sdo Paulo, Ribeira de Iguapé. 29 Oct. 1894. Loefgren
& Edwall 2814 = 12679 [HT: SP (fide Yuncker); IT (2 sheets):
ILL].-- [= 0. propinqua (fide Yuncker, 1973) ]

1985 Jones, Annotated catalogue of type specimens

elocalis Trel., Proc. Amer. Philos. Soc. 75: 703. 1935. TYPE:
Brazil, Sao Paulo, Ipiranga. March 1912. Luederwaldt 12675
(HT: SP (fide Yunker); IT (2 sheets): ILL].-- [= QO. martiana
var. martiana (fide Yuncker, 1973) ]

hammari Trel., Proc. Amer. Philos. Soc. 75: 705. 1935. TYPE:
Brazil, Sdo Paulo, Cantareira. 4 April 1901. Hammar 4551 =
12669 [HT: SP (fide Yuncker); IT (2 sheets): ILL].--
{= O. martiana var. martiana (fide Yuncker, 1973)]

hoehnei Trel., Proc. Amer. Philos. Soc. 75: 705. 1935. ' TYPE:
Brazil, Sdo Paulo, Butantan. 14 Oct. 1918. Hoehne 2480 [HT:
SP (fide Yuncker); IT (2 sheets): ILL].-- [= QO. martiana var.
martiana (fide Yuncker, 1973)]

ituana Trel., Proc. Amer. Philos. Soc. 75: 706. 1935. TYPE:
Brazil, Sao Paulo, ItG. 25 March 1897. Russell 199 = 18268
(HT: SP (fide Yuncker); IT: ILL].-- [= O. propingua (fide
Yuncker, 1973)]

loefgreni Trel.,. Proc. Amer. Philos. Soc. 752707. 1935.) TYPE:
Brazil, Sao Paulo, Pirituba. 30 Oct. 1893. Loefgren 1552 =
12681 (HT: SP (fide Yuncker); IT: ILL].-- [= 0. martiana var.
martiana (fide Yuncker, 1973) ]

* mexiae Trel., Proc. Amer. Philos. Soc. 75: 709. 1935. TYPE:
Brazil, Minas Gerais, Barbada, Vigosa. El. 680 m. 9 Oct.
1930. Mexia 5160 (specimen marked "Type" by Trelease =
LECTOTYPE designated herein: ILL).-- [= 0. anisum (fide
Yuneker, 1973)]

* mexiae var. fertilior Trel., Proc. Amer. Philos. Soc. 75: 709.
1935. TYPE: Brazil, Minas Gerais, Barbada. El. 710 m.
9 April 1930. Mexia 4587 [specimen marked "Type" by Trelease
= LECTOTYPE designated herein: ILL; PT at ILL (2 sheets):
Mexia 4619].-- [= 0. anisum (fide Yuncker, 1973) ]

* miguelitensis Trel., Proc. Amer. Philos. Soc. 75: 709. 1935.
TYPE: Brazil, Minas Gerais, Sao Miguel road, Vigosa. El.
690 m. 4 April 1930. Mexia 4564 [published as 4964--
specimen cited as HT by Yuncker (1973) = LECTOTYPE: US; ILT:
ILL (2 sheets but only one marked "Type" by Trelease); PT at
ILL: Mexia 4137, 4373-a, 4499, 4564-a (published as 4964-a, a
Syntype), 4676 (in part), and 4702].-- [= 0. propinqua (fide
Yuncker, 1973) ]

* miguelitensis var. longipes Trel., Proc. Amer. Philos. Soc. 75:
710. 1935. TYPE: Brazil, Minas Gerais, Fazenda de Graziuma,
Vigosa. El. 700 m. 15 May 1930. Mexia 4702 [HT, or

6 Poy’ TeGokeOrGr iA Vol. 58, No.

LECTOTYPE designated herein (specimen marked "Type" by
Trelease): ILL; IT or ILT (in part cited for ‘var.'

miguelitensis): ILL].-- [= 0. propinqua (fide Yuncker,
1973) ]

* miguelitensis var. parcepilosa Trel., Proc. Amer. Philos. Soc,
75: 710. 1935. TYPE: Brazil, Minas Gerais, Areponga to Sao
Miguel road. El. 850 m. 5 May 1930. Mexia 4676-b
[LECTOTYPE designated herein: ILL (the first of 2 syntypes
cited but only this specimen marked "Type" by Trelease); ST
at ILL: Mexia 4676-a].-- [= 0. propinqua]

mosenii Trel., Proc. Amer. Philos. Soc. 75: 710. 1935. TYPE:
Brazil, Sao Paulo, Serra de Caracol. Mosen 1689 (HT: S;
fragments of type at ILL).-- [= 0. frutescens (C.DC.) Trel.
(fide Yuncker, 1973) ]

palhosana Trel., Proc. Amer. PRI1GS. -Soeso 75? FiZ5)1935- TN TIPE:
Brazil, Palhoga, Florianopolis. 22 Oct. 1929. Hoehne 24462
(HT: SP (fide Yuncker); IT: ILL].-- [= 0. propinqua (fide
Yuncker, 1973) ]

peruibensis Trel., Proc. Amer. Phéltos. Saes°75ee7 122 719852 * TYPE:
Brazil, Sao Paulo, Peruibe. 30 Oct. 1891. Loefgren 1625 =
12677 [HT: SP (fide Yuncker); IT: ILL].-- [= 0. propinqua
(fide Yuncker, 1973) ]

pohliana Miq., Linnaea 20: 179. 1847eo TYPEs Brazily "In sylvis
ad Coralfaga, m. Febr." Pohl 5033 (HT: W; PT at ILL: Pohl

3031 = Mikan 30).-- [= Piper hoffmannseggianum Roemer &
Sehultes (fide Yuncker, 1973) ]

* riedelii /Trel:, Proc: “Asers (Pniidos«Soc..i75< \7133 19935. “TYPE:
Brazil. Riedel s.n. (HT, or LECTOTYPE designated herein: ILL
(the specimen was obtained from W)].-- [= ?0. frutescens
(fide Yuncker, 1973)]

rio-feioana Trel., Proc. Amer. Philos. Soc. 75: 713. 1935.
TYPE: Brazil, "Rfo Feio Expedition". Edwall 12666 [HT: SP
(fide Yuneker); IT: ILL].-- [= O. martiana var. martiana
(fide Yuncker, 1973) ]

Santa-annae Trel., Proc. Amer. Philos. Soc. 75: 713. 1935.
TYPE: Brazil, Sao Paulo, Santa Anna. Nov. 1912. Brade 5786
= 7183 [HT: SP (fide Yuncker); IT: ILL].-- [= 0. propinqua
(fide Yuncker, 1973) ]

taperana Trel., Proc. Amer. Philos. Soc. 75: 715. 1935. TYPE:
Brazil, Pernambuco, Palmares, Tapera. Oct. 1931. Pickel
2789 (HT: US 1541640; IT: ILL).-- [This species recognized
by Yuncker, 1973]

1985 Jones, Annotated catalogue of type specimens 7

usterii Trel., Proc. Amer. Philos. Soc. 75: 716. 1935. TYPE:
Brazil, Sdo Paulo, Sado Caetano. 16 Nov. 1907. Usteri 12683
(HT: SP; IT: ILL).-- [= ?0. propingua (cf. Yuncker, 1973) ]

Peltobryon Klotzsch ex Miq.

poeppigii Klotzsch ex Miq., Syst. Piper. 370. 1844. TYPE: Peru.
Poeppig 1738 [HT: B; IT: ILL (obtained from W)].-- [= Piper
callosum Ruiz & Pavén (fide Yuncker, 1955, 1972) ]

Piper L.

abditum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 130.
1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m. Aug.-Sep.
1929. Killip & Smith 27659 (HT: US; IT: ILL).

acamptostachys Trel. in Feddes Repert. Spec. Nov. Regni Veg. 24:
361. 1928. TYPE: Haiti, La Selle, Morne des Commissaires.
25 Aug. 1926. Ekman 6742 (HT: ILL).

achoteanum Trel., J. Wash. Acad. Sci. 19: 328. 1929. TYPE:
Honduras, Comayagua, El Achote, near Siguatepeque. El.
1500 m. 18 Feb. 1928. Standley 56125 (HT: F 581879; IT:
ILL).-- [This species recognized by Standley & Steyerm.,
1952]

achromatolepis Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 130. 1936. TYPE: Peru, Loreto, Fortaleza, Yurimaguas.
El. 155-210 m. Oct.-Nov. 1929. Williams 4244 (HT: F 624649;
se DIS * PT rete els Killip & Smith 27254).

actae Trel. in Yuncker, Field Mus. Publ. Bot. 17: 340. 1938.
TYPE: Honduras, in sandy soil along the beach [near] Tela.
20 July 1934. Yuncker 4657 (HT: ILL).

adornatispicum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 340.
1938. TYPE: Honduras, Comayagua, in a ravine at El Achote,
above Siguatepeque. El. 1050 m. 15 July 1936. Yuncker,
Dawson & Youse 5885 (HT: ILL).

aduncum L. var, garcia-barrigae Trel. & Yuncker, Piperaceae no.
S. Amer. 1: 251. 1950. TYPE: Colombia, Meta, R{o Negro.
El. 500 m. 25 May 1940. Garcia Barriga 8472 (HT: US
1851560; PT at ILL: Killip 34419).

aduncum var. sigualense Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 23: 5, 8. 1926. TYPE: Cuba, El Sigual, Eggers
4654 (HT: US 798411; PT at ILL: Ekman 14890).

PHY TO LDS ELA Vol. 58, No. 1

aegrum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 132.
1936. TYPE: Peru, Jun{n, Puerto Bermidez. El. 375 m. 14-17
July 1929. Killip & Smith 26478 (HT: US; IT: ILL).

aequale Vahl forma acutispicum Trel. ex Stenlé, Fl. Deser.
Antill. Fr. 2(1): 82. 1940; Bull. Agric. Martinique 9(3):
159. 1940. TYPE: Martinique. 8 Aug. 1939. Stenlé & Stenlé
3382 (HT and IT: ILL).-- [= P. aequale (fide Howard, 1973) ]

aequale {var.] 8 ovalifolium C.DC. in DC., Prodromus 16{1): 311.
1869. TYPE: Trinidad. Sieber 192 (HT: G-DC; IT: ILL).--
[This variety recognized by Yuncker, 1953]

aequale var. variifolium Trel. ex Stehlé, Bull. Agric.
Martinique 9(3): 158. 1940. Martinique. 12 Aug. 1939.
Stenlé & Stehlé 3394 (HT: ILL).-- [= P. aequale (fide Howard,
1973) ]

aereum Trel., Contr. U.S. Nat. Herb. 26(4): 144. 1929. TYPE:
Costa Rica, Cartago, Orosi. Standley 39842 (HT: US 1229481;
PT at ILL: Standley & Valerio 50891, 50970, 51205).-- [This
species recognized by Burger, 1971]

aeruginosibaccum Trel., J. Wash. Acad. Sef ip 1924928557929;
TYPE: Honduras, Atlantida, near La Ceiba. 11 March 1928.
Standley 56735 (HT: F 581879; IT: ILL).-- [= P.
jacquemontianum Kunth (fide Burger, 1971) ]

affectans Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
287, 1940. TYPE: Panama, Chiriquf, vicinity of Casita Alta,
Voledn de Chiriquf. El. 1500-2000 m. 1938. Woodson, Allen
& Seibert 865 (HT: ILL).-- [= P. imperiale (Miq.) C.DC. (fide
Yunceker, 1950 and Burger, 1971) ]

affictum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
132. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 150-350 m.
28-30 Aug. 1929. Killip & Smith PESBiOGHTE*US;: IF:2LLL; PT
at ILL: Killip & Smith 28563).

afflictum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
132. 1936. TYPE: Peru, Loreto, Mainas. 1830. Poeppig s.n.
(HT: W; PT at ILL: Killip & Smith 27067, 29293; Williams
2699).

aguacalientis Trel. in Standley, Field Mus. Publ. Bot. 18: 330.
1937. TYPE: Costa Rica, SW of Agua Caliente. El. 1560 m.
Stork 1317 (HT: ILL).-- [= P. hispidum Sw. (fide Burger,
1971) ]

1985 Jones, Annotated catalogue of type specimens 9

albert-smithii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 413,
fig. 375. 1950. TYPE: Colombia, Santander, vicinity of El
Roble. El. ca 1500 m. 16 Feb. 1927. Killip & Smith 19360
(HT: US 1354616; IT: ILL).-- [= P. ef. veraguense C.DC. in
DC. (cf. Burger, 1971)]

albopapillatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 133. 1936. TYPE: Peru, Junfn, Schunke Hacienda, San
Ramo6n. El. 1400-1700 m. 8-12 June 1929. Killip & Smith
24616) (AT: USGS ITecELL)A

allatum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 341. 1938.
TYPE: Honduras, Comayagua, in a ravine at El Rincon, 10 mi. W
of Siguatepeque. El. 1400-1500 m. 24 July 1936. Yuncker,
Dawson & Youse 6068 (HT: ILL).

alleni Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 25: 826.
1938. TYPE: Panama, Darién, Pinogana-Yavisa Trail.
El. 150me) 17 Mareh: 1937. “Allen 270.(HTs°ILL).--" [=/P.
lucigaudens C.DC. var. allenii (Trel.) Yuncker in Woodson &
Schery, 1950]

alstoni Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 287.
1940. TYPE: Panama, Coclé, north rim of El Valle de Anton.
4 June 1939 [cited as "Feb. 12"]. Alston & Allen 1842 (HT:
ILL).-- [= P. auritum Kunth (fide Yuncker, 1950 and 1957]

alveatum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 287.
1940. TYPE: Panama, Bocas del Toro, vicinity of Nievecita.
Aug. 1938. Woodson, Allen & Seibert 1837 (HT: ILL).-- [= P.
linearifolium C.DC. (fide Yuncker, 1950); = P.
friedrichsthalii C.DC. in DC. (cf. Burger, 1971)]

alveolatifolium Trel., J. Wash. Acad. Sci. 19: 329. 1929. TYPE:
Honduras, Comayagua, near Siguatepeque. El. 1080-1400 m.
Feb. 1928. Standley 56344 (HT: F 581751; IT: ILL).-- [= P.
bredemeyeri Jacq. (fide Burger, 1971) ]

amazonasense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 392,
fig. 352. 1950. TYPE: Venezuela, Amazonas, Isla de San
Carlos de Rfo Negro. El. 100 m. 27 Feb. 1942. Williams
14504 (HT: US 1859700; IT: ILL).

amnigaudens Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
134. 1936. TYPE: Peru, Loreto, steep bank of Rfo Santiago.
El. 200 m. 23 Dec. 1931. Mexia 6332 (HT: US; IT: ILL).

amoenum Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer. 1:
293, fig. 259. 1950. TYPE: Colombia, Chocd, Rfo Nuqui.
El. 200 m. 23 Jan. 1947. Haught 5474 (HT: ILL).

10 Pale ¥i TO. LeOiee dar Vol. 58, No. 1

amphibium Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
288. 1940. TYPE: Panama, Coclé, between Aguadulce and Ant6n.
El. 15-50 m. 12 July 1938. Woodson, Allen & Seibert 1222

(HT: ILL).-- [= P. taboganum C.DC. (fide Yuncker, 1950 and
annotation); = P. pseudo-fuligineum C.DC. (cf. Burger, 1971) ]

amphitrichum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
134. 1936. TYPE: Peru, Junf{n, Colonia Perené. El. 680 m.
14-22 June 1929. Killip & Smith 24938 (HT: US; IT: ILL).

amplectenticaule Trel. & Yuncker, Piperaceae no. S. Amer. 1:
325, fig. 290. 1950. TYPE: Colombia, Meta, Rfo Negrito, 70
km. E of Villavicencio. 25 Dec. 1938. Haught 2491 (HT: US
1707110; IT: ILL; PT at ILL: Haught 4289).

andresense Trel. in Standley, Field Mus. Publ. Bot. 17: 230.
1937. TYPE: Guatemala, Petén, Lake Petén, San Andrés. 3 May
1933. Lundell 3203 (HT: ILL; PT at ILL: Lundell 2073).--

{= P. martensianum C.DC. (fide Standley & Steyerm., 1952) ]

andresense var. subcalvescens Trel. in Standley, Field Mus.
Publ. Bot. 17: 231. 1937. TYPE: Guatemala, Petén, Yaxha.
23 March 1933. Lundell 2011 (HT: ILL).-- [= P. martensianum
(ef. Standley & Steyerm., 1952)]

angustiae Trel. [?ined.] var. quiotepecanum Trel., J. Wash.
Acad. Sci. 19: 329. 1929. TYPE: Mexico, Oaxaca, Pueblo de
Quiotepec, Cuicatlan. El. 500 m. H. & C. Conzatti & Gomez
2389 CHP scbn 246911 Ee: PEEL

anisophyllum Trel., Contr. U.S. Nat. Herb. 26(4): 157. 1929.
Based on P. dilatatum Rich. var. acutifolium CBC. LaEIPEs
Costa Rica, El General. Tonduz 3385 (Pseudo-paratype at
ILL, i.e., specimen cited by Trel.: Standley & Valerio
44228).-- [= P. terrabanum C.DC. (fide Burger, 1971) ]

annulatispicum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 329,
fig. 297. 1950. TYPE: Colombia, Antioquia, "Belén" [the
sheet at ILL is labelled "Medellin"]. Archer 1032 (HT: US
1542614; probable IT: ILL).

* antioquiense C.DC. var. pitaense Yuncker in Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 319. 1950. TYPE: Colombia,
Antioquia, on Quibrada Pita, E of Turbo. El. 100 m.

10 April 1946. Haught 4787 (LECTOTYPE designated herein:
SLs TILE ELE

apertum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 341. 1938.
TYPE: Honduras, Cortés, Lake Yojoa. El. 630 m. 29 July
1934. Yuncker 4821 (HT: Yuncker's herbarium; fragments of
type at ILL).

1985 Jones, Annotated catalogue of type specimens Hi

apertum var. nodosum Trel. in Yuncker, Field Mus. Publ. Bot. 17:
341. 1938. TYPE: Honduras, Cortés, Lake Yojoa. El. 630 m.
29 July 1934. Yuncker 4821-a (HT: ILL).

apodum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 135.
1936. TYPE: Peru, Hudnuco, Muna. El. 2100 m. May-June
1923. Macbride 3987 (HT: F 535049; IT: ILL).

apurimacanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
135. 1936. TYPE: Peru, Ayacucho, Rio Apurimac near
Kimpitiriki. El. 400 m. 10, 11 May 1929. Killip & Smith
22926: {HP: USs (iT: ELL)x

apus Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 136.
1936. TYPE: Peru, Junfn, Pichis Trail, Santa Rosa. El. 625-
900 m. 6, 7 July 1929. Killip & Smith 26192 (HT: US;

Ths eLis)i

aragonense Trel., Contr. U.S. Nat. Herb. 26(4): 146. 1929.
TYPE: Costa Rica, Aragon. El. 700 m. 20 Oct. 1894. Tonduz
9021 (HT: US 796497; PT at ILL: Tonduz 12790).-- [= P.

multiplinervium C.DC. (cf. Burger, 1971)]

araneatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
136. 1936. TYPE: Peru, Junf{n, Colonia Perené. El. 680 m.
14-25 June 1929. Killip & Smith 25242 (HT: US; IT: ILL).

arbelaezii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 155, fig.
134. 1950. TYPE: Colombia, Santander, Barbosa. El. 1500-
1600 m. 24 Feb. 1940. Arbelaez & Cuatrecasas 8127 (HT: US
1796074 acl F -ELis (PT Atak: Killip & Smith 15265).

* arbuscula Trel., J. Arnold Arb. 9: 146. 1928. TYPE: New Guinea,
Papua, U-Uma River. 18 May 1926. Brass 1449 (LECTOTYPE
designated herein: A; ILT: ILL).-- [= P. wichmannii C.DC.
(fide Chew, 1972)]

archeri Trel. & Yuncker, Piperaceae no. S. Amer. 1: 111, fig.
84. 1950. TYPE: Colombia, La Sierra, 18 kmN of Medellin.
El. ca 200 m. 4 Jan. 1931. Archer 1375 [HT: US 1542650; IT:
ILL; PT at ILL: Archer 1555, 2122; Tomas Alberto 7, 1675,
2122, s.n. (2 sheets); Tomas Alberto & Daniel 2731].

archidonense Trel., Ciencia (Mexico) 2: 206. 1941. TYPE:
Ecuador, Napo-Pastaza, Archidona. 4 May 1865. Isern 1791
(HT: MA; fragment of type at ILL).-- [This species upheld by
Trel. & Yuncker, 1950]

12

PR YY THO OiGelsA Vol. 58, No. 1

arctilimbum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:

288. 1940. TYPE: Panama, Coclé, vicinity of El Valle.

23 Feb. 1937. Miller 1813 (HT: US; IT: ILL).-- far Bs
linearifolium (fide Yuncker, 1950); = P. friedrichsthalii
(fide Burger, 1971)]

arctilimbum var. allenii Trel. in Woodson & Schery, Ann. Mo.
Bot. Gard. 27: 288. 1940. TYPE: Panama, Coclé, El Valle de
Anton. El. 600 m. 17 Sep. 1939. Allen 2003 (HT: ILL).--
{= P. linearifolium (fide Yuncker, 1950; P. friedrichsthalii

(fide Burger, 1971)]

arduum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 136.
1936. TYPE: Peru, Loreto, Puerto Arturo, Yurimaguas.
El. 155-210 m. Oct.-Nov. 1929. Williams 5174 (HT: F 618063;
€T22TEb) 2

arenicola Trel. in Yuncker, Field Mus. Publ. Bot. 17: 341. 1938.

TYPE: Honduras, between Tela and Lancetilla. El. 27 m.
Yuncker 4499 (HT: ILL). .

argentamentum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 256,
fig. 218. 1950. TYPE: Colombia, Choc6, Istmina. El. ca
75 m. 29 April 1939. Killip 35452 (HT: US 1772162; IT: ILL;
PT at ILL: Cuatrecasas 16057; Killip 34758; Killip &
Cuatrecasas 39048).

aripoense Trel., Bull. Torrey Bot. Club 51: 5. 1924. TYPE:
Trinidad, heights of Aripo. 16 March 1921. Britton &
Freeman 2353 (HT: NY; IT: ILL).-- [= P. demeraranum (Miq.)
C.DC in DC. (fide Yuncker, 1953) ]

armatum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 243, fig.
211. 1950. TYPE: Ecuador, Napo-Pastaza, Zatzayacu.
El. 400-500 m. 25 March 1935. Mexia 7096 (HT: US 1619103;
ITs $L4).

arrectispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 137. 1936. TYPE: Peru, San Martin, San Roque.
El. 1350-1500 m. Jan.-Feb. 1930. Williams 6993 (HT: F
629617: (IT: “IGL)%

arreptum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
137. 1936. TYPE: Peru, Junfn, Puerto Bermidez. El. 375 m.
14-17 July 1929. Killip & Smith 26487 (HT: US; IT: ILL).

artanthe C.DC. in DC. var. strigosum Trel. & Yuncker, Piperaceae
no. S. Amer. 1: 359. 1950. TYPE: Colombia, between Yarumal
and the Llanos de Cuiba. El. 2300-2700 m. 20 Feb. 1942.
Metcalf & Cuatrecasas 30143 (HT: US 1833392; IT: ILL).

1985 Jones, Annotated catalogue of type specimens A3

arundinetorum Trel., Contr. U.S. Nat. Herb. 26(4): 159. 1929.
TYPE: Costa Rica, San José, near Laguna de la Escuadra, NE of
El Copey. El. 2000-2200 m. 16 Dec. 1925. Standley 41967
(HT: US 1307268; IT: ILL).-- [= P. subsessilifolium C.DC.
(fide Burger, 1971)] my

asclepiadifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 138. 1936. TYPE: Peru, Loreto, Iquitos. El. 100 m.
3-11 Aug. 1929. Killip & Smith 27016 (HT: US; IT: ILL).

aspericaule Trel., J. Wash. Acad. Sci. 19: 329. 1929. TYPE:
Honduras, Atlantida, near Tela. 27 Jan. 1928. Standley
55125 (HT: F 583270; IT: ILL). R

asperrimicaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 138. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 27955 (CHT: US; 31 5.LL).

asterotrichum C.DC. var. yessupense Trel. in Macbr., Field Mus.
Publ. Bot. 13(No. 357): 139. 1936. TYPE: Peru, Junin, Puerto
Yessup. El. 400 m. 10-12 July 1929. Killip & Smith 26271
(HT: US:.i7s ILL).

atlantidanum Trel., J. Wash. Acad. Sci. 19: 329. 1929. TYPE:
Honduras, Atlantida, near La Ceiba. 11 March 1928. Standley
56739 (HT: F 581620; IT: ILL; PT at ILL: Standley 54938,
55736, 56600).-- [= P. pseudo-fuligineum (fide Burger, 1971) ]

atlantidanum var. sibunense Trel. ex Standley in Standley &
Record, Field Mus. Publ. Bot. 12: 406. 1936. TYPE: British
Honduras (Belize), Craig Point, Sibun River. 15 Nov. 1934.
Gentle 1381 (HT: MICH; IT: ILL).-- [= P. cf.

pseud -fuligineum] a

atlantidanum var. yoroense Trel., J. Wash. Acad. Sci. 19: 330.
1929. TYPE: Honduras, Yoro, near Progreso. El. 30 m.
24 Jan. 1928. Standley 55073 (HT: F 583981; IT: ILL) .--

[= P. ef. pseudo-fuligineum]

atrichopus Trel., J. Wash. Acad. Sci. 19: 330. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 150 m.
16 Jan. 1928. Standley 54594 (HT: F 583633; IT: ILL; PT at
ILL: Standley 52616, 52871, 53290, 56757, S5B757 2)

atrichopus var. arborescens Trel. in Yuncker, Field Mus. Publ.
Bot. 9: 276. 1940. TYPE: Honduras, Atlantida, about 25 km E
of La Ceiba. El. 300 m. 21 July 1938. Yuncker, Koepper &
Wagner 8578 (HT: ILL).

14

PRA POR? Se A Vol. 58, No. 1

atrobaccum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 234, fig.
202. 1950. TYPE: Colombia, El Valle, forest above La Cumbre.
El. 1800-2100 m. 14-19 May 1922. Killip 5596 (HT: US
1141684; IT: ILL; PT at ILL: Killip 11598; Killip & Garcfa
33695; Pennell & Killip 7225).

atrobaccum var. pubinervulum Trel. & Yuncker, Piperaceae no. S,
Amer. 1: 235. 1950. TYPE: Colombia, El Valle, Rfo Calima
Valley, El Cairo, between Darién and Mediacanoa. El. 1650-
1750 m. 7 Jan. 1943. Cuatrecasas 13933 (HT: US 1853102; IT:
ILL). '

atropremnon Trel. in Feddes Repert. Spec. Nov. Regni Veg. 24:
361. 1928. TYPE: Haiti, La Hotte. El. 700 m. 30 March
1927. Ekman 7920 (HT: ILL).

attenuatamentum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 139. 1936. TYPE: Peru, Junfn, Puerto BermGdez.
El. 375 m. 14-17 July 1929. Killip & Smith 26513 (HT: US;
i Gt eine be

auriculiferum Trel., Contr. U.S. Nat. Herb. 26(4): 156. 1929.
TYPE: Costa Rica, Guanacaste, El Silencio, near Tilar4n.
El. 750 m. 13 Jan. 1926. Standley & Valerio 44648 (HT: US
1307316; IT: ILL).-- [= P. terrabanum (fide Burger, 1971) ]

auritilaminum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 277.
1940. TYPE: Honduras, Atlantida, lower slopes of Mt.
Cangrejal. 8 July 1938. Yuncker, Koepper & Wagner 8263 (HT:
TLE).

auritilimbum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 277.
1940. TYPE: Honduras, Atlantida, Rio Danto, slopes of Mt.
Cangrejal. 2 Aug. 1938. Yuncker, Koepper & Wagner 8763 (HT:
ILL),

auritum Kunth forma rinconense Trel. in Yuncker, Field Mus.
Publ. Bot. 17: 342. 1938. TYPE: Honduras, Comayagua, El
Rincon, Siguatepeque. El. 1400-1500 m. 24 July 1936.
Yuneker, Dawson & Youse 6064 (HT: ILL).

austro-mexicanum Trel., J. Wash. Acad. Sci. 16: 204. 1926.
TYPE: Mexico, Chiapas, Monserrate. 1925. Purpus 35 (HT: US
12093705" IT: ° TLE).

azupizuanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
139. 1936. TYPE: Peru, Junfn, Pichis Trail, San Nicol&s to
AzupizG. El. 650-900 m. 6 July 1929. Killip & Smith 26094
rs Vesa TEED.

1985 Jones, Annotated catalogue of type specimens 15

baculiferum Trel., Contr. U.S. Nat. Herb. 26(4): 180. 1929.
TYPE: Costa Rica, Guanacaste, El Arenal. El. 500 m.
Jan. 1926. Standley & Valerio 45052 (HT: US 1307342; PT at
ILL: Standley & Valerio 45154).-- [= P. hispidum (fide
Burger, 1971)]

bajanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 390, fig.
350. 1950. TYPE: Colombia, Santander, vicinity of La Baja.
El. 3000 m. Jan. 1927. Killip & Smith 18119 (HT: US
1353586; IT: ILL).

balaoense C.DC., Annuaires Conserv. Jard. Bot. Geneve 21: 258.
1920. TYPE: Ecuador, Guayas, Balao. Jan. 1892. Eggers
14350 (HT: M; fragment of type at ILL).-- [= P. manabinum
C.DC. (fide Trel. & Yuncker, 1950)]

balsapuertanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 140. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 150-
350 m. 28-30 Aug. 1929. Killip & Smith 28394 (HT: US; IT:
ILL)

banaoanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
4, 7. 1926. TYPE: Cuba, El Plurial, Lomas de Banao, Santa
Clara. El. 850 m. 27 Jan. 1926. Ekman 16230 (HT: ILL).

barbatum Kunth [var.] isernii Trel., Ciencia (Mexico) 2: 207.
1941. TYPE: Ecuador, Quito. Isern "Nos. 1271, 1268"
(Syntypes, perhaps to be lectotypified: MA?; ST at ILL: Isern
1268).-- [= P. barbatum var. barbatum (fide Trel. & Yuncker,
1950) ]

barbirostre Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
288. 1940. TYPE: Panama, Chiriquf, vicinity of "New
Switzerland". Jan. 1939. Allen 1365 (HT: ILL).-- [= P.
hispidum (fide Yuncker, 1950 and 1957)]

begoniicolor Trel. & Yuncker, Piperaceae no. S. Amer. 1: 140,
fig. 117. 1950. TYPE: Colombia, Antioquia, Puerto Valdivia,
between Valdivia and Yarumal. El. 1610 m. 20 Feb. 1942.
Metcalf & Cuatrecasas 30099 (HT: US 1833353; IT: ILL; PT at
ILL: Daniel & Gonzalez 2123).

begoniicolor var. lobatum Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 141. 1950. TYPE: Colombia, El Valle, San Antonio,
Woof Cali. El. 1900-2350 m. Feb.-March 1939. Killip &
Garcfa 33915 (HT: US 1770778; IT: ILL; PT at ILL: Ewan
16158).

bejucalanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 29:
28. 1931. TYPE: Santo Domingo, Bejucal, San José de Ocoa,
Azua. 6 March 1929. Ekman 11805 (HT: ILL).

16

RHYL DOL O48 BA Vol. 58, No. 1

belloi Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 85, fig. 73.
1966. TYPE: Brazil, Espirito Santo, Concordia da Cachoeira.
1889. Bello 567 (HT: R; PT at ILL: Sao Paulo, Loefgren
1433).-- [This species upheld by Yuncker, 1973]

bellum Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer. 1:
57, fig. 48. 1950 (publ. as bella). TYPE: Colombia,
Antioquia, near Vijagual, 30 km S of Turbo. 13 April 1945.
Hought 4559 (HT: US 1709141; IT: ILL).

benianum Trel. in Rusby, Mem. N. Y. Bot. Gard. 7: 222. 1927.
TYPE: Bolivia, Huachi, head of Beni River. El. 3000 ft.
Sep. 1921. Cardenas 466 (HT: NY; IT: ILL).-- [= P. callosum
(fide Yuncker, 1955)]

bermudezense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
141. 1936. TYPE: Peru, Junin, Puerto Bermudez. El. 375 m.
14-17 July 1929. Killip & Smith 26424 (HT: US; IT: ILL).

bisasperatum Trel., Contr. U.S. Nat. Herb. 26(4): 173. 1929.
TYPE: Costa Rica, Cartago, Cerro de la Carpintera.
E1,...1500= ee m. Feb. 1924. Standley 35723 (HT: US 1229396;
52002, 52030, 52053).-- [This ees SS ss Sey
1971)

blepharilepidum Trel., Contr. U.S. Nat. Herb. 26(4): 160. 1929.
TYPE: Costa Rica, Guanacaste, Quebrada Serena, SE of Tilar4n.
El. 700 m. 27 Jan. 1926. Standley & Valerio 46217 (HT: US
1307434; IT: ILL).-- [This number is also cited, with a
question mark, among the paratypes of P. verruculigerum Trel.
(op. cit.); = P. bisasperatum (fide Burger, 1971) ]

bogotense C.DC. var. ovalilimbum Trel. & Yuncker, Piperaceae no.
S. Amer. 1: 95. 1950. TYPE: Colombia, Santander, vicinity of
La Baja. El. 2700-3500 m. Jan. 1927. Killip & Smith 18028
(HT: US 1353512; IT: ILL; PT at ILL, from Colombia: Ariste
Joseph 50, 57, A-372; Cuatrecasas 11561; Cuatrecasas & Garcia
Barriga | 10257; Daniel & Gonzalez 1847; | Killip & Smith 16720,
16953, 17127, 19662, 19689; Silvano Jorge 190, 191, 192; from

Venezuela: Tamayo 2402).

bogotense var. usaquemense Trel., Trab. Mus. Nac. Ci. Nat..Ser.
Bot. 33: 48. 1936 (without a Latin diagnosis). TYPE:
Colombia, Cundinamarca, Usaquén. El. 2700 m. Arbeldez 610
(HT: US; PT at ILL: Cuatrecasas 2416).-- [= P. bogotense
(fide Trel. & Yuncker, 1950 and Yuncker, 1950) ]

1985 Jones, Annotated catalogue of type specimens a

bopianum Trel. in Rusby, Mem. N. Y. Bot. Gard. 7: 223. 1927.
TYPE: Bolivia, Bopi River Valley. El. 3000 ft. Sep. 1921.
Rusby 669 (HT: NY; IT: ILL).-- [= P. longestylosum C.DC.

(fide Yuncker, 1955)]

boguianum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 340, fig.
311. 1950. TYPE: Colombia, Caldas, W of Rfo Boqufa Salento.
El. 1600-1900 m. 27 July 1922. Killip & Hazen 8806 (HT: US
t0H2702 5.1 IT 24 184)3

brachypodon (Bentham) C.DC. var. radicescens Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 213. 1950. TYPE: Colombia, Choco,
Tutunendo, 20 km N of Quibdé. El. ca 80 m. May 1931.
Archer 2152 (HT: US 1519167; IT: ILL).

* brachystylum Trel., Ciencia (Mexico) 2: 207. 1941. TYPE:
Ecuador, Archidona. Isern 1333 [syntype, cited as "Type" by
Trel. & Yuncker (1950, 1: 41), i.e., assumed to be the
LECTOTYPE: MA; 2 ST at ILL: Isern 1331, 1334].-- [This
species recognized by Trel. & Yuncker, 1950]

brachytrichum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 141. 1936. TYPE: Peru, Ayacucho, Estrella. El. 500 m.
8, 14 May 1929. Killip & Smith 22652 (HT: US; IT: ILL):

* brassii Trel., J. Arnold Arb. 9: 147. 1928. TYPE: New Guinea,
Papua, Iawarere. El. 350 m. 24 Nov. 1925. Brass 683
(LECTOTYPE designated herein: A; ILT: ILL).-- [= P,
macropiper Pennant (fide Chew, 1972) ]

breve Trel., Contr. U.S. Nat. Herb. 26(2): 38. 1927. TYPE:
Panama, Canal Zone, between Gamboa and Cruces. Pittier 3787
(HT: US 678858; PT at ILL: Piper 5944; Stevens 981).-- [= P.

taboganum (fide Yuncker, 1950); = P. pseudo-fuligineum (cf.
Burger, 1971 and Croat, 1978)]

breve var. bellae-vistae Trel., Contr. U.S. Nat. Herb. 26(2):
39. 1927. TYPE: Panama, Prov. Panama, Bella Vista. 16 Oct.
1922. Killip 12045 (HT: US 1167181; IT: ILL; PT at ILL:
Killip 12004).-- [= P. tobaganum (fide Yuncker); = P. cf.

pseudo-fuligineum]

breve var. divaricatum Trel., Contr. U.S. Nat. Herb. 26(2): 39.
1927. TYPE: Panama, Canal Zone, Fort Sherman. Maxon &
Valentine 6986 (HT: US 1180701; IT: ILL).-- [= P. taboganum

(fide Yuncker); = P. cf. pseudo-fuligineum])
breve var. pallidineurum Trel., Contr. U.S, Nat. Herb. 26(2):

38. 1927. TYPE: Panama, between France Field, Canal Zone and
Catival, Col6én. Standley 30164 [HT: US 1154564; PT at ILL:

18 PHYTO Loe A Vol. 58, No. 1

Killip 12078 (2 sheets); Maxon & Harvey 6591 (cited as
659"); Standley 25203, 26382; Stevens 349, 752, 993].--

—— a ee

"
[= P. taboganum (fide Yuncker); = P. cf. pseudo-fuligineum)

breve var. pseudo-fatoanum Trel., Contr. U.S. Nat. Herb. 26(2):
39. 1927. TYPE: Panama, Canal Zone, Gamboa. Pittier 3709
(HT: US 678777; PT at ILL: Maxon & Harvey 6504, 6543; Maxon &
Valentine 7055; Standley 25565, 25603).-- [= P. taboganum

(fide Yuncker); = P. cf. pseudo-fuligineum]

brevestrigillosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 141. 1936. TYPE: Peru, Ayacucho, Carrapa. El. 1500 m.
May 1929. Killip & Smith 22375 (HT: US; IT: ILL).

brevipedunculum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 141. 1936. TYPE: Peru, Ayacucho, Estrella. El. 500 m.
8, 14 May 1929. Killip & Smith 22677 (HT: US; IT: ILL).

brittonorum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
4, 8. 1926. TYPE: Cuba, Las Tunas, Isle of Pines. Britton,
Britton & Wilson 14743 (HT: NY; PT at ILL: Ekman 12118,
12182).

cabaianum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23: 307,
311. 1927. TYPE: Haiti, Morne Cabaio, Massif de La Selle.
El. 1700 m. 20 Aug. 1924. Ekman 1544 (HT: ILL).

cabaianum var. obliquifolium Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 24: 361. 1928. TYPE: Haiti, Badeau, La Selle.
El. 1300 m. 13 March 1927. Ekman 7850 (HT: ILL).

caballo-cochanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 142. 1936. TYPE: Peru, Loreto, Caballo-Cocha.
Aug. 1929. Williams 2309 (HT: F 608282; IT: ILL).

caeruleifolium Trel., Contr. U.S. Nat. Herb. 26(4): 145. 1929.
TYPE: Costa Rica, Guanacaste, Quebrada Serena, near Tilardn.
El. 700 m. 27 Jan. 1926. Standley & Valerio 46134 (HT: US
1307425; IT: ILL; PT at ILL: Standley & Valerio 46234) .--

[= P. aequale (fide Burger, 1971) ]

cajambrense Trel. & Yuncker var. caguetanum Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 27. 1950. TYPE: Colombia,
Caqueta, Rfo Orteguaza. 2 Aug. 1926. Woronow & Juzepczuk
6364 (HT: ILL).

Calanyanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 66, fig.
59. 1950. TYPE: Colombia, Caquet4, Quebrada de la Calana.
6 April 1940. Cuatrecasas 9186 (HT: US 1795455; PT at ILL:
Antonio Camilo 10; Cuatrecasas 10589).

1985 Jones, Annotated catalogue of type specimens 19

calciseligens Trel. ex Stehlé, Bull. Soc. Bot. France 84: 411.
1938 (nomen subnudum); Fl. Deser. Antill. Fr. 2(1): 97. 1940;
Bull. Agric. Martinique 9(3): 174. 1940. TYPE: Guadeloupe.
El. 10 m. 5 Feb. 1937. Stehlé & Stehlé 1340 (HT: ?;
fragments of type at ILL).-- [= P. dilatatum Rich. (fide
Howard, 1973)]

caliendriferum Trel., J. Wash. Acad. Sci. 19: 330. 1929. TYPE:
Mexico, Oaxaca, Cerro de la Raya, Cuyamecalco. El. 2800 m.
24 June 1909. H. & C. Conzatti & Gomez 2384 (HT: F 246912;
Wp Fae

calvescens Trel. var. aguanum Trel. in Yuncker, Field Mus. Publ.
Bot. 9: 278. 1940. TYPE: Honduras, Yoro, Rfo Agudn, below
Coyoles. 29 June 1938. Yuncker, Koepper & Wagner 8124 (HT:
ILL).

calvescens var. cangrejalense Trel. in Yuncker, Field Mus. Publ.
Bot. 9: 278. 1940. TYPE: Honduras, Atlantida, lower slopes
of Mt. Cangrejal, 16 July 1938. Yuncker, Koepper & Wagner
8464 (HT: ILL).

calvescens var. pedunculatum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 342. 1938. TYPE: Honduras, Cortés, Potrerillos.
31 July 1934. Yuncker 4882 (HT: ILL).

calvescens var. potrerillosense Trel. in Yuncker, Field Mus.
Publ. Bot. 17: 343. 1938 (as v. portrerillosense). TYPE:
Honduras, Cortés, Potrerillos. 31 July 1934. Yuncker 4894
(BT: ILL).

calvescentinerve Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 144. 1936. TYPE: Peru, Junfn, Puerto Yessup.
El. 400 m. 10-12 July 1929. Killip & Smith 26248 (HT: US;
iT: EE}:

camiloi Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.
1: 64, fig. 57. 1950. TYPE: Colombia, Caqueta, Florencia.
8 Jan. 1946. Antonio Camilo 11 (HT: ILL).

camptostachys Urban var. cuabanum Trel. in Feddes Repert. Spec.
Nov. Regni Veg. 29: 28. 1931. TYPE: Santo Domingo. Ekman
12306 (HT: ILL).

candicans Sodiro, Anales Univ. Centr. Ecuador 19: 201. 1905.
TYPE: Ecuador, Esmeraldas. Aug. 1904. Mille s.n. (IT or ST

s.
at ILL).-- [= P. cinereum C.DC. (fide Trel. & Yuncker, 1950) ]

20

PHYTO L.O.6.10A Vol. 58, No. 1

canelosense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 264,
fig. 228. 1950. TYPE: Ecuador, Napo-Pastaza, near Canelos.
El. 300-400 m. Feb. 1935. Mexia 6857 (HT: US 1619089; IT:
ILL; PT at ILL: Mexia 7276).

canescenticaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 145. 1936. TYPE: Peru, Loreto, Puerto Arturo.
El. 135 m. 24-25 Aug. 1929. Killip & Smith 27726 (HT: US;
ITs ILG).

canescenticaule var. expansum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 145. 1936. TYPE: Peru, Loreto, Puerto
Arturo. El. 135 m. 24-25 Aug. 1929. Killip & Smith 27717
(HTsaUSs ITt ILL):

canyazasense Trel. in Waodson & Seibert, Ann. Mo. Bot. Gard. 25:
826. 1938. TYPE: Panama, Veraguas, trail between Canazas and
the foot of the Cordillera Central. El. 300-600 m. 8 Feb.
1937. Allen 185 (HT: ILL).-- [= P. taboganum (fide Yuncker,
1950); = P. pseudo-fuligineum (cf. Burger, 1971) ]

capacibracteum Trel., Contr. U.S. Nat. Herb. 26(4): 183. 1929.
TYPE: Costa Rica, San José, between Aserri and Tarbaca.
El. 1200-1700 m. 6 Dec. 1925. Standley 41397 (HT: US
1307249; IT: ILL; PT at ILL: Standley & Valerio 41812, 41861,
43121, 43245, 43246).-- [This species recognized by Burger,
1971]

capillistipes Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 145. 1936. TYPE: Peru, Loreto, Soledad, Rfo Itaya.
El, 110m, 20-22 Sep. 1929. Killip & Smith 29607 (HT: US;
ITs :2LB)

cararense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 208, fig.

170. 1950. TYPE: Colombia, Santander, Carare Valley, near
Puerto Berrio. 28 March 1935. Haught 1604 (HT: US
1661768; IT: ILL; PT at ILL: Haught 1987, 2192, 2861).

cararense var. productipes Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 209, fig. 171. 1950. TYPE: Colombia, Santander,
vicinity of Barranca Bermeja, Amarilla Creek, Magdalena
Valley. El. 200 m. 11 July 1936. Haught 1928 (HT: US
1742206; IT: ILL; PT at ILL: Killip 11809).

cardenasii Trel. in Rusby, Mem. N. Y. Bot. Gard. 7: 223. 1927.
TYPE: Bolivia, pampas near Lake Rogagua. El. 300 m. 11 Nov.
1921. Cardenas 1663 [HT: NY; IT: ILL; probable PT (from the
type locality) at ILL: Rusby 1688 (the number cited in the
protologue is 1685)].-- [= P. aduncum (fide Trel. & Yuncker,
1950) J

1985 Jones, Annotated catalogue of type specimens 21

carlosii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 197, fig.
157. 1950. TYPE: Colombia, Cundinamarca, Aguadita. 15 Jan.
1943. Silvano Jorge et al. 181 (HT: ILL).

carpapatanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
145. 1936. TYPE: Peru, Junf{n, Carpapata above Huacapistana.
El. 2700-3200 m. 7 June 1929. Killip & Smith 24403 (HT: US;
IT: ILL).

carrapanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
146. 1936. TYPE: Peru, Ayacucho, Carrapa. El. 1200 m.
May 1929. Killip & Smith 22425 (cited as 22485) (HT: US
1357399; IT: ILL).

carrapanum var. lancifolium Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 146. 1936. TYPE: Peru, Junin, Pichis
Trail, Enenas. El. 1600-1900 m. June-July 1929. Killip &
Smith 25758 (HT: US; IT: ILL).

carrilloanum C.DC., Bull. Soc. Bot. Belg. 30(1): 209. 1891.
Syntypes, probably to be lectotypified: Costa Rica, Route de
Carrillo, versant l'Atlantique (Pittier 2525, 2526); bord du
bois a General, 600 m (Pittier 3600)-- (fragments of Pittier
2526, from LE, at ILL).-- [This species recognized by Burger,
1971]

casitense Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
289. 1940. TYPE: Panama, Chiriquf, vicinity of Casita Alta,
Voledn de Chiriquf. El. 1500-2000 m. Woodson, Allen &
Seibert 978 (HT: ILL).-- [= P. hispidum (fide Yuncker, 1950
and 1957) ]

castroanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 221, fig.
182. 1950. TYPE: Colombia, Choc6, Quibd6, Rfo Atrato.
El. ca 60 m. April-May 1931. Archer 1830 (HT: US 1519003;
ETe Ebsbohs

catalinianum Trel. in Standley, Field Mus. Publ. Bot. 18: 335.
1937. TYPE: Costa Rica, Guanacaste. 28 June 1928. Stork
2779 (HT: ILL).-- [= P. jacquemontianum (fide Burger, 1971) ]

cataractarum Trel. in Rusby, Mem. N. Y. Bot. Gard. 7: 224. 1927.
TYPE: Bolivia, cataracts of the Bopi River. El. 3000 ft.
Sep. 1921. Rusby 672 (HT: NY; IT: ILL).-- [= P. hispidum
(fide Yuncker, 1955 and 1957)]

catripense Yuncker in Trel. & Yuncker, Piperaceae no. Ss. Amer.
1: 108, fig. 81. 1950. TYPE: Colombia, Choco, Ensenada,
Catripe. 13 Jan. 1945. Haught 5432 (HT: ILL).

22 Po ¥oT!Ock OubodeA Vol. 58, No. 1

caudefactum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 124,
fig. 99. 1950. TYPE: Colombia, El Valle, Rio Dfgua Valle,
Chorrera La Elsa. El. 975 m. 2, 3 April 1939. Killip 34811
(HT: US 1771602; IT: ILL).

cauranum Trel. ex L. Williams, Explor. Bot. Guayana Venezolana,
175. 1942 (nomen subnudum). TYPE: Venezuela, Bolfvar, La
Prisién, Media Caura. El. 100 m. 29 March 1939. Williams
11628 (HT: F; IT: ILL).-- [= P. demeraranum (fide Trel. &
Yuncker, 1950; Yuncker, 1953, 1957 and 1973)]

cayoense Trel. ex Standley in Standley & Record, Field Mus.
Publ. Bot. 12: 407. 1936. TYPE: British Honduras (Belize),
San Antonio, El Cayo Distr. 6 May 1931. Bartlett 13070 [HT:
MICH; IT at ILL (2 sheets)].-- [This species recognized. by
Standley & Steyerm., 1952]

cejanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 193, f4¢.
152. 1950. TYPE: Colombia, Antioquia, La Ceja. 6 Dec. 1939.
Daniel & Gonzalez 2120 (HT: US 1778619; IT: ILL).

* celatipetiolum Trel. var. brenesii Trel. in Cuf., Archivio Bot.
Sist. Fitogeog. & Genet. 10: 26. 1934. TYPE: Costa Rica,
Pacifica, Golfito ad Golfo dulce. 12 April 1930. Brenes 80
(HT, or LECTOTYPE designated herein: ILL).-- [= P. terrabanum
(fide Burger, 1971)]

celer Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 147.
1936. TYPE: Peru, Loreto, Rio Itaya. May 1929. Williams
132 (HT: F 604312; IT: ILL; PT at ILL: Killip & Smith
27059).

centroense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 298, fig.
264. 1950. TYPE: Colombia, Santander, 20 kmS of El Centro.
El. 100 m. 25 Sep. 1936. Haught 1988 (HT: US 1742249;
STIL

cernuum Vell. Conc. var. biformipilum Yuncker, Bol. Inst. Bot.
(Sao Paulo) 3: 21. 1966. TYPE: Brazil, Minas Gerais, Vi¢gosa,
road to Sado Miguel, near km 3-1/2. El. 685 m. 18 Sep. 1930.
Mexia 5064a (HT: ILL).-- [This variety recognized by Yuncker,
1972)

cerro-puntoense Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
27: 289. 1940. TYPE: Panama, Chiriquf, trail to Cerro Punta,
Rfo Chiriqu{ Viejo Valley. 29 March 1938. White 52 (HT:
ILL).-- [= P. hispidum (fide Yuncker, 1950 and 1957) ]

1985 Jones, Annotated catalogue of type specimens 23

* chalhuapuguianum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 147. 1936. TYPE: Peru, Junfn, Hacienda Chalhuapuquio.
6 Dec. 1924. Stevens 104 (HT, or LECTOTYPE designated
herein: ILL; PT at ILL: Stevens 106, 144).

chanchamayanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 148. 1936. TYPE: Peru, Jun{n, Chanchamayo Valley.
El. 1200 m. Schunke 1595 (HT: F 627279; PT at ILL: Killip &
Smith 22634).

chanekii Trel. ex Standley in Standley & Record, Field Mus.
Publ. Bot. 12: 407. 1936. TYPE: British Honduras (Belize),
El Cayo. March-June 1933. Chanek 10 (HT: MICH; IT:
ILL).-- [= P. scabrum Sw. (fide Standl. & Steyerm., 1952); =

affin. P. hispidum (cf. Howard, 1973)]

chiadoense Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.
1: 337, fig. 307. 1950. TYPE: Colombia, Antioquia, on Rfo
Chiado, 65 km S of Turbo. El. ca 120 m. 7 May 1945. Haught
4642 (HT: US 1709185; IT: ILL).

chrysolepidum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 24:
361. 1928. TYPE: Santo Domingo, between Banane and Gros
Fignier, Cordillera de Bahoruco, Barahona. El. 500 m.

28 Aug. 1926. Ekman 6760 (HT: ILL).

chrysoneurum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
148. 1936. TYPE: Peru, Loreto, Alto Rfo Itaya. El. 145 m.
Sep.-Oct. 1929. Williams 3207 (HT: F; IT: ILL).

cinereonervosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 149. 1936. TYPE: Peru, Loreto, San Antonio.
El. 110 m. 18 Sep. 1929. Killip & Smith 29380 [HT: US; IT
(2 sheets-- one leaf each): ILL].

cinereoramulum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 149. 1936. TYPE: Peru, Loreto, mouth of Rfo Santiago.
El. 200 m. 17 Nov. 1931. Mexia 6117 (HT: UC; IT: ILL).

cirratum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
150. 1936. TYPE: Peru, Loreto, La Victoria on the Amazon
River. Aug.-Sep. 1929. Williams 2969 (HT: F 608159; IT:
ILL).

cisnerosense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 304,
fig. 269. 1950. TYPE: Colombia, El Valle, Cisneros.
El. 300-500 m. 5 May 1939. Killip 35571 (HT: US 1772274;
ITs TLD

24

PRY TS oo rn Vol. 58, No. 1

cispontinum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
150. 1936. TYPE: Peru, Junfn, near Perené Bridge, Rfo
Paucartambo Valley. El. 700 m. 19 June 1929. Killip &
Smith 25284 (HT: US; IT: ILL).

Claudicans Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
151. 1936. TYPE: Peru, Loreto, Iquitos. El. 100 m.
3-11 Aug. 1929. Killip & Smith 26938 (HT: US; IT: ILL).

coactaepilum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
151. 1936. TYPE: Peru, Junfn, Puerto Yessup. El. 400 m.
10-12 July 1929. Killip & Smith 2631C"(RT s. USS ITS SLL).

cobanense Trel. in Standley, Field Mus. Publ. Bot. 10: 159.
1931. “Honduras, Quebrada Seca, also in Guatemala." No type
cited; to be lectotypified?; probably first collected by
Standley (see No. 53930, one of several authentic specimens
at TEE):

cobanense var. sarculatum Trel., J. Wash. Acad. Sci. 19: 331.
1929. TYPE: Honduras, Yoro, Quebrada Seca. El. 30 m. Dec.
1927. Standley 53912 [HT: F 584243; IT (2 sheets): ILL].

cocornanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 297, fig.
262. 1950. TYPE: Colombia, Antioquia, Cocornad. 30 July
1937. Daniel 1431 CHT: Gi; IT: ILL).

cocquericotense Trel. ex Standley in Standley & Record, Field
Mus. Publ. Bot. 12: 407. 1936. TYPE: British Honduras
(Belize), Little Cocquericot, Belize River. 26 March 1933.
Lundell 3829 (HT: MICH; IT: ILL).

colaphitolerans Trel. in Standley, Field Mus. Publ. Bot. 17:
231. 1937. TYPE: Guatemala, Petén, Lake Peteén, Ixlu.
15 June 1933. Lundell 3827 (HT: ILL).-- [= P. psilorhachis
C.DC. (fide Standley & Steyerm., 1952)]

colemanense Trel. in Standley, Field Mus. Publ. Bot. 18: 337.
1937. TYPE: Costa Rica, Heredia, Potreros near Coleman
Finca, above Santo Domingo del Roble. El. 1600 m. 7 Aug.
1936. Dodge & Goerger 9580 (HT: MO; IT: ILL).-- [= P.
aequale (fide Burger, 1971) ]

comasense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
152. 1936. TYPE: Peru, Junfn, Rfo de Comas. El. 2500-
2600 m. Weberbauer 6615 (HT: F 628196; IT: ILL).

comayaguanum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 343.
1938. TYPE: Honduras, Comayagua, 6 km N of Siguatepeque.
El. 1300 m. 4 Aug. 1936. Yuncker, Dawson & Youse 6310 (HT:
ILL).

1985 Jones, Annotated catalogue of type specimens Pde

compactum Trel., Contr. U.S. Nat. Herb. 26(4): 131. 1929. TYPE:
Costa Rica, San José, near Santa Marfa de Dota. El. 1500-
1800 m. Dec. 1925. Standley 41801 (HT: US 1307255; IT: ILL;
PT at ILL: Standley & Valerio 43184-- said to be
broader-leaved).-- [= P. amalago (cf. Burger, 1971) ]

concinnatoris Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 33.
1966.-- [see Artanthe concinna]

concinnifolium Trel., Contr. U.S. Nat. Herb. 26(4): 167. 1929.
TYPE: Costa Rica, Colinas de Piedades, San Ramé6n.
El. 1100 m. 4 June 1901. Brenes 14194 (HT: US 716434; PT at
ILL: Brenes 17765, 17769).-- [= P. aequale (cf. Burger,
1971) ]

conditum Trel. in Macbr., Field Mus. Publ. Bot. wh 357)5
152. 1936. TYPE: Peru, Loreto, Yurimaguas. - 135 mm
Aug.-Sep. 1929. Killip & Smith 28208 (HT: bn ne ILL; PT at
ILL: Killip & Smith 28090) .-- .-- [The number of the PT is also
the type number r for P P. perpunctulatum Trel. in Macbr., op.
cit. 207]

conditum var. angustum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 152. 1936. TYPE: Peru, Loreto, Yurimaguas.
El. 135 m. Aug.-Sep. 1929. Killip & Smith 29105 (HT: US;
Ee sLLL)i.

condotoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 222,
fig. 184. 1950. TYPE: Colombia, Chocé, near confluence of
the Condoto and San Juan rivers. El. 100-150 m. 30 April
1939. Killip 35086 (HT: US 1771840; IT: ILL).

conejoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 46, fig.
35. 1950. TYPE: Colombia, Colombia-Ecuador boundary,
Putumayo, junction of the San Miguel and Conejo rivers.
El. 300 m. 8 Dec. 1940. Cuatrecasas 10896 (HT: US 1798165;
IT Sai Loh

confusionis Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
152. 1936. TYPE: Peru, Loreto, Yurimaguas to Balsapuerto.
El. 135-150 m. 26-31 Aug. 1929. Killip & Smith 28231 (HT:
US ET <=, Dee).

conispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
153. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m.
1-5 Sep. 1929. Killip & Smith 28767 (HT: US; IT: ILL).

26

Pee T Ob BiG add Vol. 58, No. 1

constipatispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 153. 1936. TYPE: Peru, Loreto, San Antonio.
El. 110 m. 18 Sep. 1929. Killip & Smith 29415 (HT: US; IT:
ILL; PT at ILL: Killip & Smith 29302, 29442).

constipatispicum var. lanosum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 153. 1936. TYPE: Peru, Loreto, Iquitos.
El. 110 m. 3-11 Aug. 1929. Killip & Smith 27467 (HT: US;
IT: TLL).

contamanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):

154. 1936. TYPE: Peru, Loreto, Contamana on Rfo Ucayali.
El. 150 m. 29 July 1929. Killip & Smith 26880 (HT: US; IT:
ILL).

conversum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
290. 1940. TYPE: Panama, Chiriquf, Llanos del Volcan.
El. 1300 m. 23 Jan. 1939. Allen 1550 (HT: ILL).-- [This
species upheld by Yuncker, 1950; = P. amalago (fide Burger,
1971) ]

cooperi Yuncker in Woodson & Schery, Ann. Mo. Bot. Gard.) 37: 21.
1950. TYPE: Panama, Bocas del Toro, vicinity of Almirante,
Daytonia Farm. 1928. Cooper 374 (HT: US; IT: ILL).-- [= P.
fimbriulatum C.DC. (fide Burger, 1971) ]

copacabanense Trel. in Daniel, Una ascension Cerro "La Vieja",
14. 1940 (without a Latin diagnosis); Caldasia 1: 88. 1940.
TYPE: Colombia, Copacabana, near Medellin. 28 April 1934.
Daniel 279 (HT: US; IT: ILL).-- [= P. bredemeyeri (fide Trel.
& Yuncker, 1950) ]

corintoananum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 33, fig.
27. 1966. TYPE: Brazil, Minas Gerais, Fazenda do Diamante,
Corrego Diamante, Corinto. El. 590 m. 10 April 1931. Mexia
5583-a (HT: ILL).

correptum Trel. in Macbr., Field Mus. Publ. Bot. 13(Nos 357):
154. 1936. TYPE: Peru, Junin, Puerto Bermudez. El. 375 m.
14-17 July 1929. Killip & Smith 26524 (HT: US; IT: ILL).

* corylistachyopse Trel., J. Arnold Arb. 9: 147.. 4928.: TTYPE: New

Guinea, Papua, Sogeri. El. 450 m. 17 Nov. 1925. Brass 655
(LECTOTYPE, designated herein: A; ILT: ILL).

coyolesense Trel. in Yuncker, Field Mus. Publ. Bot. 9: 278.
1940. TYPE: Honduras, Yoro, near Las Flores, Rfo Aguan.
El. 270 m. 30 June 1938. Yuncker, Koepper & Wagner 8147
(HT: ILL).-- [= P. atlantidanum (fide Standley & Steyerm.,

1952); = P. pseudo-fuligineum (cf. Burger, 1971)]

1985 Jones, Annotated catalogue of type specimens 27

crassamentum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
155. 1936. TYPE: Peru, Junfn, Puerto Yessup. El. 400 m.
10-12 July 1929. Killip & Smith 26354 (HT: US; IT: ILL).

crassicaule Trel. in Yuncker, Field Mus. Publ. Bot. 9: 278.
1940. TYPE: Honduras, Atlantida, ridge near Roma railroad
siding about 25 km E of La Ceiba. El. 240 m. 21 July 1938.
Yuneker, Koepper & Wagner 8572 (HT: ILL).

cricamolense Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
290. 1940. TYPE: Panama, Bocas del Toro, Rfo Cricamola
between Finca St. Louis and Konkintoé. El. ca 10-50 m.

Aug. 1938. Woodson, Allen & Seibert 1927 (HT: ILL).-- [This
species recognized by ~Yuneker, 1950 ]

crispans Trel. in Standley, Field Mus. Publ. Bot. 18: 339. 1937.
TYPE: Costa Rica, La Palma de San Ramon. El. 1050 m. Brenes
5236 (HT: F; fragments of type (Brenes 390 = 5236) at ILL].--
t= P. aequale (fide Burger, 1971) ]

crispatimargine Trel. in Standley, Field Mus. Publ. Bot. 18:
339. 1937. TYPE: Costa Rica, Palmira del Naranjo. El. 1850-
1900 m. Brenes 3499 [HT: F; fragments of type (Brenes 35 =
3499) at ILLJ.-- [= P. poasanum C.DC. (fide Burger, 1971) ]

cristalinanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 129,
fig. 104. 1950. TYPE: Colombia, El Valle, Rio Digua, La
Elsa, Quebrada La Cristalina. El. 1000-1150 m. 30 Sep.
1943. Cuatrecasas 15231 (HT or LT: US; IT or ILT: ILL).

cristicola Trel. in Yuncker, Field Mus. Publ. Bot. 17: 343.
1938. TYPE: Honduras, Comayagua, in forest at summit of the
range above El Achote, near Siguatepeque. El. 1800 m.
Yuncker, Dawson & Youse 6247 (HT: ILL).

cryptocarpum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 281,
fig. 245. 1950. TYPE: Ecuador, Napo-Pastaza, between Tena
and Archidona. El. 400 m. April 1935. Mexia 7132 (HT: US
1619105; IT: ILL; PT at ILL: Isern 1788).

cuatrecasasi Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 48.
1936 (without a Latin diagnosis). TYPE: Colombia, Tolima,
Ibagué. El. 1350 m. 8 May 1932. Cuatrecasas 2426 (HT:
ILL).-- [= P. aduncum (fide Trel. & Yuncker, 1950; Yuncker
1950 and 1955) ]

* cubataonum C.DC., Notizbl. Bot. Gart. Berlin-Dahlem 6: 440.
1917. TYPE: Brazil, Sado Paulo, Cubatado. Usteri 3, 14
[Syntypes: G.-- Yuncker (1973) cited No. 14 as the holotype,
but it is to be regarded as the LECTOTYPE (designated

28

PH Vr OM OxGuln A Vol. 58, No. 1

herein). An unnumbered Usteri specimen at ILL (obtained from
SP under Access. No. 12631) has the same label data as No. 4
and may be an ILT).-- [This species recognized by Yuncker,
1973]

* cufodontii Trel. in Cuf., Archivio Bot. Sist. Fitogeogr. &

Genet. 10: 25. 1934. TYPE: Costa Rica, "Hamburg-Farm, 9 km
ab ore flum. Reventazon." El. 15 m. 2 May 1930. Cufodontis
684 (HT, or LECTOTYPE designated herein: ILL).-- [= P.
arieianum C.DC. (fide Burger, 1971) ]

cumanense Kunth var. pubinerve Trel. & Yuncker, Piperaceae no.

S. Amer. 1: 237. 1950. TYPE: Colombia, Choco, Bahfa Solano,
near Ciudad Mutis. Feb. 1939. Killip & Garcfa 33472 (HT: US
1770407; IT: ILL).

cumanense var. subvillosum Trel. & Yuncker, Piperaceae no. S.

Amer.- 1: 237. 1950. TYPE: Colombia, Cundinamarca, La Vega,
road to Nocaina. El. 950-1200 m. Jan. 1942. Garcfa Barriga
10641 (DHT: US: 1851617; IT:ILL: PT at ILL: Apolinar Marfa 9;

Felipe 104].

cumbricola Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 48.

1936 (without a Latin diagnosis). TYPE: Colombia, La Cumbre.
El. 1650 m. Cuatrecasas 2012 and 2427 = syntypes [No. 2012
cited as "type" by Trel. & Yuncker, 1950, 1: 251 (i.e.,
assumed to be the LECTOTYPE): ILL].-- [= P. aduncum (fide
Trel. & Yuncker, 1950; Yuncker, 1950 and 1955) ]

cumbricola var. montevagum Trel., Trab. Mus. Nac. Ci. Nat. Ser.

Bot. 33: 48. 1936 (without a Latin diagnosis). TYPE:
Colombia, Ibagué. El. 1350 m. 8 May 1932. Cuatrecasas 2425
(HT: MA; fragment of type at ILL).-- [= P. aduncum (fide
Trel. & Yuncker; 1950) ]

cundinamarcanum Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33:

49, 1936 (without a Latin diagnosis). TYPE: Colombia,
Cundinamarca, La Esperanza. Cuatrecasas 2420-b (HT: ILL).--
[= P. holtonii C.DC. in DC. var. cundinamarcanum (Trel.)

Trel. & Yuncker, 1950)

cuniculorum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 234,

fig. 201. 1950. TYPE: Colombia, Putumayo, Columbia-Ecuador
boundary, near the confluence of the San Miguel and Conejo
rivers. El. 300 m. 8 Dec. 1940. Cuatrecasas 10906 (HT: US
47991012. 1T:..ILL).

Cupreatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
156. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 350-550 m.
29 Aug. 1929. Killip & Smith 26509..¢HT.s. USs1IT2 BLL; PT at
ILL: Killip & Smith 28274, 28764).

1985 Jones, Annotated catalogue of type specimens 29

cupreatum var. longum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 156. 1936. TYPE: Peru, Loreto, Iquitos.
El. 100 m. 3-11 Aug. 1929. Killip & Smith 27486 (HT: US;
Ite: ELlad<

curvatipes Trel. in Standley, Field Mus. Publ. Bot. 17: 231.
1937. TYPE: Guatemala, Petén, Lake Petén, Nictun. 4 May
1933. Lundell 3122 (HT: ILL).-- [This species recognized by
Standley & Steyerm., 1952]

cyclophyllum Trel. in Standley, Field Mus. Publ. Bot. 10: 159.
1931. Type locality: Honduras, "frequent in wet forest on
the high hills above Lancetilla." No type indicated;
probably lectotypification needed (authentic specimen at ILL:

Standley 52703).

cyphophyllopse Trel. & Yuncker, Piperaceae no. S. Amer. 1: 334,
fig. 303. 1950. TYPE: Ecuador, Los Rios, Canté6n Vinces,
between Quevedo and Naranjal. El. 60 m. 7, 8 Nov. 1934.
Mexia 6673 (HT: US 1619077; IT: ILL).

cyphophyllopse var. brevipes Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 335. 1950. TYPE: Ecuador, Los Rios, Cant6én Vinces,

near Quevedo. El. 60 m. Oct.-Nov. 1934. Mexia 6628 [HT: US
1619072; IT: ILL; PT at ILL, from Colombia: Felipe 93; Killip
& Garcfa 33051, 33111; Antonio Miguel 3, 25; from Ecuador:
Eggers 14848 (fragment); Rimbach 233].

cyprium Trel. & Yuncker, Piperaceae no. S. Amer. 1: 115, fig.
89. 1950. TYPE: Colombia, Caldas, above Salento. El. 2000-
2500 m. July 1922. Pennell 8925 (HT: US 1142731; IT: ILL).

daedalum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
156. 1936. TYPE: Peru, Loreto, lower R{fo Huallaga. El. 155-
210 m. Oct.-Nov. 1929. Williams 3861 (HT: F; IT: ILL).

daniel-gonzalezii Trel. in Daniel, Una ascensi6n Cerro "La
Vieja", 13. 1940 (without a Latin diagnosis); Caldasia 1: 87.
1940. TYPE: Colombia, Antioquia, La Ceja. Daniel 2113
(cited as 2133) (HT: US 1778612; IT: ILL).-- [This species
recognized by Trel. & Yuncker, 1950]

daniel-gonzalezii var. misellum Trel. & Yuncker, Piperaceae no.
S. Amer. 1: 201. 1950. TYPE: Colombia, Antioquia, San
Antonio. July 1940. Daniel 2316 [HT: US 1800222; PT at
ILL: Daniel 3010 (2 sheets)].

30

Ps Hel: FOO. LivGrG:sbor Vol. 58, No. 1

dawsoni Trel. in Yuncker, Field Mus. Publ. Bot. 17: 343. 1938.
TYPE: Honduras, Comayagua, El Achote, above Siguatepeque.
El. 1350 m. 3 Aug. 1936. Yuncker, Dawson & Youse 6283 (HT:
ILL).

deamii Trel. var. yojoanum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 344. 1938. TYPE: Honduras, Paral, Lake Yojoa.
El. 630 m. 30 July 1934. Yuncker 4875 (HT: ILL).

debilicaule Trel. & Yuncker, Piperaceae no. S. Amer. 1: 333,

fig. 302. 1950. TYPE: Ecuador, Esmeraldas, Parroquia
Concepcion, Playa Rica. El. 105 m. 7 Dec. 1936. Mexia 8411
(HTs°US01778418; ITs ILL).

decorticans Trel. & Yuncker, Piperaceae no. S. Amer. 1: 125,
fig. 100. 1950. TYPE: Colombia, El Valle, 18 kmE of
Buenaventura. El. ca 50 m. 14 Feb. 1939. Killip & Garcia
33236 (HT: US 1770196; IT: ILL).

dedititium Trel., J. Wash. Acad. Sci. 19: 331. 1929. TYPE:
Honduras, Yoro, Quebrada Seca. El. 30 m. Dec. 1927.

Standley 53937 (HT: F; IT: ILL).-- [= P. jacquemontianum
(fide Burger, 1971)]

deflexispicum Trel., Contr. U.S. Nat. Herb. 26(4): 144. 1929.
TYPE: Costa Rica, Alajuela, Viento Fresco. El. 1600-1900 m.
13 Feb. 1926. Standley & Torres 47764 (HT: US 1307512; PT
at ILL: Standley & Torres 47576, 51349; Standley & Valerio
49899).-- [= P. gibbosum C.DC. (fide Burger, 1971) ]

delatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 157.
1936. TYPE: Peru, Loreto, Puerto Arturo. El. 135 m.
24-25 Aug. 1929. Killip & Smith 27713 (HT: US; IT: ILL).

deltoideocarpum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 344,
1938. TYPE: Honduras, Comayagua, river bank near
Siguatepeque. El. 1050 m. 5 July 1936. Yuncker, Dawson &
Youse 5654 (HT: ILL).

dendropse Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
158. 1936. TYPE: Peru, Junin, La Merced. El. 700 m.
May-June 1929. Killip & Smith 23488 (HT: US; IT: ILL).

depressibaccum Trel. in Feddes Repert. Spec. Nov. Regni Veg.
23: 306, 310. 1927. TYPE: Haiti, Port-de-Paix, Haut-Piton.
El. 1000 m. 20 Sep. 1925. Ekman 4890 (HT: ILL).

desultorium Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
158. 1936. TYPE: Peru, Jun{n, Schunke Hacienda, above San
Ramon, La Merced. El. 1400-1700 m. 8-12 June 1929. Killip
& Smith 24654 (HT: US; IT: ILL).

1985 Jones, Annotated catalogue of type specimens 31

detonsum Trel., Contr. U.S. Nat. Herb. 26(4): 141. 1929. TYPE:
Costa Rica, Guanacaste, near Tilardn. El. 500-650 m.
Jan. 1926. Standley & Valerio 44314 (HT: US 1307305; IT:
ILL; PT at ILL: Standley & Valerio 44451, 44601, 44765,
wee (= P. glabrescens (Miq.) C.DC. in DC. (fide Burger,
1971)

diazanum Trel., Contr. U.S. Nat. Herb. 26(2): 34. 1927. TYPE:
Panama, Prov. Panama, E of Juan Diaz. Killip 3114 [HT: US
836136; IT: ILL; PT at ILL: Maxon 6843; Maxon & & Valentine
7012 (2 sheets); Standley 260 06).-- [=P . leptocladum C.D Cc .DC.
(fide Yuneker, 1950); = P. dilatatum Rich. (cf. Burger,
1971)]

diazanum var. viae-kobeanae Trel. in Woodson & Schery, Ann. Mo.
Bot. Gard. 27: 290. 1940. TYPE: Panama, Canal Zone, Fort
Kobe Road. 22 July 1938. Woodson, Allen & Seibert 1412 (HT:
ILL).-- [= P. leptocladum (fide Yuncker, 1950); = P.
dilatatum (er. Burger, 1971)]

dichroostachyum Trel. & Yuncker, Piperaceae no. S, Amer. 1: 294,
fig. 260. 1950. TYPE: Colombia, El Valle, El Silencia,
Yanaconas. El. 1900-2200 m. 28 Feb. 1939. Killip & Garefa
33824 (HT: US 1770703; IT: ILL; PT at ILL: Fosberg 19626).

dichroostachyum var. mayasqueranum Trel. & Yuncker, Piperaceae
no. S. Amer. 1: 295. 1950. TYPE: Colombia, Narino, trail

between Mayasquer and Tambo. El. 3000 m. 2 Aug. 1935.
Mexia 7577 (HT: GH; IT: ILL).

dilatatifolium Trel. & Yuncker, Piperaceae no. S. Amer. 1: 267,
fig. 232. 1950. TYPE: Venezuela, Fed. Distr., Bosque de
Catuche. Pittier 6136 (HT: US 601867; PT at ILL: Colombia,

Killip 34463).

* dilatatum Rich. var. broadwayi Trel. ex Stehlé, Bull. Soc. Bot.
France 83: 627. 1936 (nomen subnudum-- as "broawayi"). TYPE:
Guadeloupe. Jan. 1929. Trelease 69 (LECTOTYPE design. by
Howard, 1973: ILL; ILT: ILL).-- [= P. dilatatum (fide
Howard, 1973) ]

* dilatatum forma diamantense Trel. ex Stehlé, Fl. Deser. Antill.
Fr. 2(1): 96. 1940; Bull. Agric. Martinique 9(3): 173. 1940.
TYPE: Martinique, Diamant. El. 100 m. 29 March 1939.
Stenlé & Stenlé 3215 (LECTOTYPE, designated by Howard, 1973:
ILL).-- [= P. dilatatum (fide Howard, 1973)]

32 PoneY T00%L70 Gdn Vol. 58, No. 1

* dilatatum forma vauclinii Trel. ex Stenlé, Fl. Deser. Antill.
Fr. 2(1): 94. 1940; Bull. Agric. Martinique 9(3): 171. 1940.
TYPE: Martinique. El. 562 m. 8 March 1939. Stehlé & Stenlé
3208 (LECTOTYPE, designated by Howard, 1973: ILL).-- [= P.
dilatatum (fide Howard, 1973)] 7

dimorphophyllum Trel. ex Standley in Standley & Record, Field
Mus. Publ. Bot. 12: 407. 1936. TYPE: British Honduras
(Belize), Sibun River, Craig Point. 15 Nov. 1934. Gentle
1387 (HT: MICH; IT: ILL).-- [= P. aeruginosibaccum (fide
Standley & Steyerm., 1952); = P. jacquemontianum (cf. Burger,
1971)]

discolor Trel. in Standley, Field Mus. Publ. Bot. 17: 231. 1937,
non Sw. (1797)-- the specimens are labelled "P. discolorum".
TYPE: Guatemala, Petén, La Libertad. 7 April 1933. Lundell
2565 (HT: ILL; PT at ILL: Lundell 4886).-- [= P.
aerug inosibaccum (fide Standley & Steyerm., 1952); = P.

jacquemontianum (cf. Burger, 1971) ]

discriminatum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 341,
fig. 313. 1950. TYPE: Colombia, Santander, vicinity of Las
Vegas. Killip & Smitn 16051 (HT: GH; IT: ILL; PT at ILL:
Killip & Smith 16137).

dispansum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
159. 1936. TYPE: Peru, Junfn, Puerto Bermidez. £1. 375 m.
24-25 Aug. 1929. Killip & Smith 26434 (HT: F 607559; PT at
PEGS Killip & Smith 27612, 27684, 27882).

disparipes Trel., Contr. U.S. Nat. Herb. 26(4): 162. 1929.
TYPE: Costa Rica, Guanacaste, El Silencio, Tilaran.
El. 750 m. 13 Jan. 1926. Standley & Valerio 44766 (HT: US
1307327; IT: ILL).-- [= P. umbricola C.DC. (fide Burger,
1971) ]

dissimulans Trel., Contr. U.S. Nat. Herb. 26(4): 133. 1929.
TYPE: Costa Rica, Las Vueltas, Tucurrique. El. 635 m.
Nov. 1898. Tonduz 12773 (HT: US 798447; PT at ILL: Standley
& Valerio 46718).-- [= P. papantlense C.DC. in DC. (fide
Burger, 1971) ]

distichum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 206, fig.
167. 1950. TYPE: Colombia, Chocd, Bahfa Solano, along
Quebrada Jellita. El. 50-100 m. 22 Feb. 1939. Killip &
Garcfa 33556 (HT: US 1770479; IT: ILL; PT at ILL: Haught
4786).

diversipilum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
159. 1936. TYPE: Peru, Hudnuco, Pozuzo. El. 600 m. 20-22
June 1923. Macbride 4666 (HT: F 535740; IT: ILL).

1985 Jones, Annotated catalogue of type specimens 33

divortans Trel. & Yuncker, Piperaceae no. S. Amer. 1: 189, fig.
148. 1950. TYPE: Colombia, Antioquia, Cocorna. 27 Dec.
1939. Daniel & Gonzalez 2119 (HT: US 1778618; IT: ILL; PT
at ILL: Cuatrecasas 4794; Felipe 37; Silvano Jorge 228;
Silvano Jorge & Alfonso 5).

dotanum Trel., Contr. U.S. Nat. Herb. 26(4): 165. 1929. TYPE:
Costa Rica, San José, Santa Marfa de Dota. El. 1500-1800 m.
Dec. 1925. Standley 41739 (HT: US 1307252; IT: ILL; PT at
ILL: Standley 41849).-- [This species recognized by Burger,
1971]

duidaense Trel. var. bolivianum Yuncker, Lilloa 27: 162, pl. 42
[1953] 1955. TYPE: Bolivia, La Paz, Hacienda Simaco, along
road to Tipuani. El. 1400 m. March 1920. Buchtien 5370
(HT: US 1176612; IT: ILL; PT at ILL: Buchtien 601).

dulce Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 160.
1936. TYPE: Peru, Junfn, La Merced. El. 700 m. May-June
1929. Killip & Smith 23705 (HT: US; IT: ILL).

dussii C.DC. var. dugommierianum Trel. ex Stehlé, Bull. Soc.
Bot. France 84: 411. 1938 (nomen subnudum); Fl. Descr.
Antill. Fr. 2(1): 112. 1940 (as forma); Bull. Agric.
Martinique 9(3): 189, tab. 8. 1940. TYPE: Guadeloupe.
5 Dec. 1936. Stehlé & Stenlé 1292 (HT: NY; IT: ILL).--
[= P. hispidum (fide Howard, 1973) ]

echeverrianum Trel., Contr. U.S. Nat. Herb. 26(4): 172. 1929.
TYPE: Costa Rica, Echeverrfa. May 1890. Pittier 2547 [HT:
US 796295; fragment of type (leaf) at ILLJ.-- [= P.
dilatatum (fide Burger, 1971) ]

ecuadorense Sodiro, Anales Univ. Centr. Ecuador 13: 859. 1898;
Piperaceas ecuator, 145. 1900; Fl. Ecuator. Monogr. 1, Ed. 2,
147, 1901. TYPE: Ecuador, Mt. Pichincha, Sodiro s.n. (HT: B;
IT: ILL).-- [This species recognized by Trel. & Yuncker,
1950)

edurum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 160.
1936. TYPE: Peru, Ayacucho, Aina. El. 750-1000 m.
7, 17 May 1929. Killip & Smith 22618 (HT: US; IT: ILL; PT at
ILL: Killip & Smith 22566, 24325).

edwallii Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 36, fig. 30.
1966. TYPE: Brazil, Sao Paulo, Conceicao de Itanhaém.
5 Nov. 1891. Loefgren & Edwall 1668 (HT: ILL).-- [This
species upheld by Yuncker, 1972]

34

PeY oO: 8 6 IA Vol. 58, No. 1

ejuncidum Trel., Contr. U.S. Nat. Herb. 26(4): 164. 1929. TYPE:
Costa Rica, Heredia, Cerro de las Caricias above San Isidro.
El, 2300 m. 11 March 1926, Standley & Valerio 52220 (HT: US
1307670; PT at ILL: Standley & Valerio 49773, 49784,
50260).-- [= P. carpinteranum C.DC. (fide Burger, 1971)]

elasmophyllum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 278.
1940. TYPE: Honduras, Atlantida, lower slopes of Mt.
Cangrejal. 16 July 1938. Yuncker, Koepper & Wagner 6461
(HT: ILL).-- [= P. atlantidanum (fide Standley & Steyern.,
1952); = P. pseudo-fuligineum (cf. Burger, 1971)]

el-bancoanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 90.
1950. TYPE: Colombia, Norte de Santander, El Banco, at the
confluence of Cubugén and Cobaria rivers. El. 320 m.

15 Nov. 1941. Cuatrecasas 13158 (HT: US 1850052; PT at
ILL: Cuatrecasas 11281, 13129; Nicéforo 202).

el-caranyanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 246,
fig. 214. 1950. TYPE: Colombia, Norte de Santander, Rfo
Cubug6n Valley, between El Carano and El Indio. El. 470-
600 m. 12 Nov. 1941. Cuatrecasas 13015 (HT: US 1850452;
IT? Tb)

elliptico-oblongifolium Trel. in Rusby, Mem. N.Y. Bot. Gard. 7:
224. 1927. TYPE: Bolivia, Bopi River Valley. El. 3000 ft.
15 Sep. 1921. Rusby 671 (HT NY:..1T: - ILL). -- [=P
buchtienii C.DC. var. ~ charopampanum (C.DC.) Yuneker (1955) ]

el-metanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 160, fig.
139. 1950. TYPE: Colombia, Meta, Villavicencio. El. 500 m.
19 Jan. 1939. Haught 2539 (HT: US 1707140; IT: ILL; PT at
ILL: Nicéforo 200).

elongatifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 161. 1936. TYPE: Peru, Junfn, La Merced. El. 600 m.
10-24 Aug. 1923. Macbride 5287 (HT: F 536312; IT: ILL; PT
at ILL: Killip & Smith 25113, 25124).

elongatum Vahl var. pampayacuanum Trel. in Macbr., Field Mus.
Publ. Bot. 13(No. 357): 162. 1936 (as pampayacusum,
apparently a misprinting vs. spelling in Trelease's
handwriting on the sheet at ILL). TYPE: Peru, Hudnuco,
Pampayacu, at mouth of Rfo Chinchao. El. 1100 m. 19-25 July
1923. Macbride 5047 (HT: F; IT: ILL).

emancipationis Trel. in Standley, Field Mus. Publ. Bot. 17: 231.
1937. TYPE: Guatemala, Petén, La Libertad. 7 June 1933.
Lundell 3658 (HT: ILL).-- [= P. aeruginosibaccum (fide
Standley & Steyerm., 1952); = P. jacquemontianum (fide
Burger, 1971) ]

1985 Jones, Annotated catalogue of type specimens 35

emancipationis var. longum Trel. in Standley, Field Mus. Publ.
Bot. 17: 232. 1937. TYPE: Guatemala, Petén, La Libertad.
27 April 1933. Lundell 3052 (HT: ILL).-- [= P.
aeruginosibaccum (fide Standley & Steyernm., 1952); = P,
jacquemontianum (fide Burger, 1971) ] ¥

emollitum Trel., Contr. U.S. Nat. Herb. 26(4): 181. 1929. TYPE:
Costa Rica, Guanacaste, El Silencio, Tilaran. El. 750 m.
13 Jan. 1926. Standley & Valerio 44650 (HT: US wie x PT
at ILL: Standley & Valerio LEI 46205).-- [= P.
bisasperatum (fide Burger, 1971) ]

enenyasense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
163. 1936. TYPE: Peru, Junin, Pichis Trail, Enenas.
El. 1600-1900 m. June-July 1929. Killip & Smith 25646 (HT:
US; IT: ILL).-- [This collection was filed by Trelease with
and is also cited as a PT of P. lanosicaule Trel. in Macbr.,
op. cit. 181.]

enganyanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 292, fig.
258. 1950. TYPE: Colombia, El Valle, Rio Digua Valley along
Rio Engafia. El. ca 675 m. April 1939. Killip 34750 (HT: US
VTTVOAT FIT 2 EEG).

entradense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 40. 1950.
TYPE: Ecuador, Guayas, Entrada. 27 June 1942. Haught 3357
CHT? US. 1708091%"IT+ FEL).

eocenicum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23: 307,
311. 1927. TYPE: Haiti, Morne Basil, l'Artibonite.
El. 1300-1450 m. 15 Nov. 1924. Ekman 2502 (HT: ILL).

eocenicum var. cahosense Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 24: 362. 1928. TYPE: Haiti, Morne Lagonne, Los
Cahos. El. 1200 m. 22 Feb. 1926. Ekman 5604 (HT: ILL).

epilosipes Trel. in Yuncker, Field Mus. Publ. Bot. 17: 344.
1938. TYPE: Honduras, Comayagua, El Achote, above
Siguatepeque. El. 1350 m. 3 Aug. 1936. Yuncker, Dawson &
Youse 6296 (HT: ILL).

epilosipes var. heterophyllum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 345. 1938. TYPE: Honduras, El Achote, above
Siguatepeque. El. 1350 m. 26 July 1936. Yuncker, Dawson &
Youse 6097 (HT: ILL).

erectipilum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 95, fig. 82.
1966. TYPE: Brazil, Para, Gurupa. 3 Nov. 1929. Killip &
Smith 30578 (HT: US 1463602; IT: ILL; PT at ILL: Killip &
Smith 30609).-- [This species upheld by Yuncker, 1973]

36

Pte OybebuGrg fA Vol. 58, No. 1

erubescentispicum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
27: 291. 1940 (as erubescentispica). TYPE: Panama, Bocas
del Toro, vicinity of Nievecita. El. 0-50 m. Aug. 1938.
Woodson, Allen & Seibert 1817 (HT: ILL).-- [This species
recognized by Yuncker, 1950]

estrellense C.DC. in DC., Prodromus 16(1): 272. 1869, including
var. pilosius (Miq.) C.DC. in DC., loc. cit.-- [see Artanthe
fuscescens ]

euryphyllum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
163. 1936. TYPE: Peru, Junf{n, E of Quimirf Bridge, near La
Merced. El. 800-1300 m. 1-3 June 1929. Killip & Smith
23891 (cited as 29891) (HT: US 1358464; IT: ILL).-- [This
species recognized by Burger, 1971]

evasum Trel., Contr. U.S. Nat. Herb. 26(4): 155. 1929. TYPE:
Costa Rica, Guanacaste, El Arenal. El. 500 m. 18 Jan. 1926.
Standley & Valerio 45280 (HT: US 1307366; IT: ILL).-- [= P.
imperiale (fide Burger, 1971) ]

evulsipilosum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 279.
1940. TYPE: Honduras, Atlantida, La Ceiba. 25 June 1938.
Yuncker, Koepper & Wagner 8015 (HT: ILL).

ewanii Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer. 1:
278, fig. 240. 1950. TYPE: Colombia, Putumayo, near San
Diego de Colorado...between Umbria and Puerta Asis. 11 Jan.
1945. Ewan 16752 (HT: ILL).

excultum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
164. 1936. TYPE: Peru, San Martf{n, San Roque. El. 1350-
1500 m. Jan.-Feb. 1930. Williams 7461 (HT: F 623031; IT:
ILdAys

exigens Trel. in Yuncker, Field Mus. Publ. Bot. 17: 345. 1938.
TYPE: Honduras, Comayagua, El Achote, above Siguatepeque.
El. 1350 m. 26 July 1936. Yuncker, Dawson & Youse 6124 (HT:
ELL).

exigens var. subovatum Trel. in Yuncker, Field Mus. Publ. Bot.
17: 345. 1938. TYPE: Honduras, El Achote, above
Siguatepeque. El. 1350 m. 21 July 1936. Yuncker, Dawson &
Youse 5997 (HT: ILL).

expolitum Trel. in Macbr., Field Mus. Publ. Bot. T3(NG. 357):
164. 1936. TYPE: Peru, Loreto, Iquitos. El. 120 m.
Oct. 1929. Williams 3590 (HT: F 618287; IT: ILL; PT at ILL:
Killip & Smith 26897, 27228).

1985 Jones, Annotated catalogue of type specimens 37

factum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 165.
1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m. Aug.-Sep.
1929. Killip & Smith 27588 (HT: US; IT: ILL).

fagifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
165. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 155-210 m.
Oct.-Nov. 1929. Williams 4782 (HT: F 629670; IT: ILL; PT at
ILL: Killip & Smith 28743).

ee Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
28: 426. 1941. TYPE: Panama, Chiriquf{, vicinity of Puerto
Armuelles. El. 0-75 m. July 1940. Woodson & Schery 852
(HT: ILL).-- [= P. darienense C.DC. (fide Yuncker, 1950) ]

falanense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 268, fig.
233. 1950. TYPE: Colombia, Tolima, Faladn, canyon of the Rfo
Gualf, road between Fresno and Falan. El. 1120-1700 m.
5 Dec. 1939. Garcfa Barriga 8333 (HT: US 1774154; PT at ILL:
Ewan 16837; Haught 5573; Killip & Garcfa 33292).

falcigerum Trel., Contr. U.S. Nat. Herb. 26(4): 147. 1929.
TYPE: Costa Rica, Guanacaste, El Silencio, Tilaran.
El. 750 m. 13 Jan. 1926. Standley & Valerio 44747 (HT: US
1307323; IT: ILL; PT at ILL: Standley & Valerio 45817) .--
[= P. terrabanum (fide Burger, 1971) ]

falculispicum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 216,
fig. 179. 1950. TYPE: Ecuador, Napo-Pastaza, trail from Tena
to Archidona. El. 400 m. 2 April 1935. Mexia 7129 (HT: US
1619104; IT: ILL; PT at ILL: Mexia 7230).

falculispicum var. cocanum Trel., Ciencia (Mexico) 2: 207. 1941.
TYPE: Ecuador, La Coca. July 1865. Isern 1545 (HT: MA;
fragments of type at ILL).-- [= P. falculispicum Trel. &
Yuncker, 1950]

fallens Trel., J. Wash. Acad. Sci. 19: 332. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. Standley
53968 (HT: F 583884; IT: ILL).-- [This species recognized by
Standley & Steyerm., 1952; = P. near urostachyum Hemsley (cf.
Burger, 1971: 188) ]

fastiditum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 299, fig.
265. 1950. TYPE: Colombia, Santander, vicinity of Charta.
El. 2000-2600 m. Feb. 1927. Killip & Smith 19102 (HT: US
1354402; IT: ILL; PT at ILL: Killip & Smith 16559, 16671,
17057, 18873, 18920, 19011, 19146).

—— ———

38

PHM EOcke BiG 1A Vol. 58, No. 1

fastiditum var. pubescens Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 299. 1950. TYPE: Colombia, Santander, wooded banks
of Rfo de la Baja, below La Baja. El. 2200-2300 m. 26 Jan.
1927. Killip & Smith 18283 (HT: US 1353720; IT: ILL).

faviculiferum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 345,
pl. 11. 1938. TYPE: Honduras, Comayagua, El Achote, above
Siguatepeque. El. 1350 m. 7 Aug. 1936. Yuncker, Dawson &
Youse 6368 (HT: ILL).-- [= P. alveolatifolium (fide Standley

& Steyerm., 1952); = P. bredemeyeri (cf. Burger, 1971) ]

fenianum Trel., J. Wash. Acad. Sci. 16: 204. 1926. TYPE:
Mexico, Chiapas, Hacienda Fenix. 1925. Purpus 386 (HT: US
1209377; IT: ILL).

figlinum Trel., Contr. U.S. Nat. Herb. 26(4): 142. 1929. TYPE:
Costa Rica, San José, vicinity of Santa Marfa de Dota.
El. 1500-1800 m. Standley & Valerio 43112 (HT: US 1307280;
IT: ILL).-- [= P. glabrescens (fide Burger, 1971) ]

firmifolium Trel. in Yuncker, Field Mus. Publ. Bot. 17: 346.
1938. TYPE: Honduras, Comayagua, near El Rincon, about 10
miles west of Siguatepeque. El. 1400-1500 m. 24 July, 1936.
Yuncker, Dawson & Youse 6048 (LECTOTYPE designated herein:
ILL: SLT: SEL),

flagellicuspe Trel. & Yuncker, Piperaceae no. S. Amer. 1: 58,
fig. 49. 1950. TYPE: Ecuador, Napo-Pastaza, near Archidona.
El. 650 m. April 1935. Mexia 7268 (HT: US 1619112; IT: ILL;
PT at ILL: Mexia 8492).

flagellicuspe var. villosulum Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 59. 1950. TYPE: Colombia, Choc6éd, Quibdd, Rfo
Atrato. El. ca 60 m. April-May 1931. Archer 1915 (HT: US
1519059; IT: ILL; PT at ILL: Killip 35412).

flavifructum Trel., J. Arnold Arb. 9: 147. 1928. TYPE: New
Guinea, Papua, Domara River. 4 June 1926. Brass 1644
(LECTOTYPE designated herein: A; ILT: ILL).-- [= P. caninum
Blume (fide Chew, 1972)]

fluvii-initii Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
291. 1940. TYPE: Panama, Chiriqui, trail from Cerro Punta to
headwaters of Rio Caldera. El. 2250-2500 m. 14 Jan. 1939.
Allen 1445 (HT: ILL).-- [= P. petaresanum C.DC. (fide
Yuncker, 1950) ]

fontium Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23: 307,
311. 1927. TYPE: Haiti, Morne Tourrise, Port-de-Paix.
El. 50 m. 24 March 1925. Ekman 3571 (HT: ILL).

1985 Jones, Annotated catalogue of type specimens 39

fortalezanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
166. 1936. TYPE: Peru, Loreto, lower Rf{o Huallaga,
Fortaleza. El. 155-210 m. Oct.-Nov. 1929. Williams 4351
(HT: F 629586; IT: ILL; PT at ILL: Killip & Smith 26281).

* fragrans Trel., J. Arnold Arb. 9: 147. 1928. TYPE: New Guinea,
Papua, Iawarere. El. 550 m. 25 Nov. 1925. Brass 701
(LECTOTYPE designated herein: A; ILT: ILL).

fraguanum Trel., J. Wash. Acad. Sci. 19: 332. 1929. TYPE:
Honduras, Atlantida, La Fragua. 7 Feb. 1928. Standley 55730
(HT: F; IT: ILL).-- [This species recognized by Standley &
Steyerm., 1952; = P. sancti-felicis Trel. (fide Burger,
1971) ]

fraguanum var. viride Trel. in Yuncker, Field Mus. Publ. Bot.
17: 346. 1938. TYPE: Honduras, Lancetilla Valley. El. 27 m.
5 Aug. 1934. Yuncker 4918 (HT: ILL).-- [= P. sancti-felicis
(fide Burger, 1971) ]

fresnoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 237, fig.
204. 1950. TYPE: Colombia, Tolima, El Fresno. El. 1480 m.
7 May 1940. Cuatrecasas 9384 (HT: US 1795883; PT at ILL:
Silvano Jorge 227).

friedrichsthalii C.DC. in DC., Prodromus 16(1): 327. 1869. TYPE:
"In Guatemala Orosi (Friedrichsthal, n. 1424! inh.
Vindob.)." (HT: ?; IT: ILL (obtained from W)].-- [This
species recognized by Standley & Steyerm., 1952, who pointed
out that the type locality actually is in Costa Rica; see
also Burger, 1971]

frijolesanum Trel. {[ined.?] var. grandifolium Trel. in Woodson &
Schery, Ann. Mo. Bot. Gard. 27: 291. 1940. TYPE: Panama,
Coclé, El Valle. El. 600-1100 m. 8 Dec. 1938. Allen 1193
(HT: ILL).-- [= P. imperiale (fide Yuncker, 1950)

fulvescenticaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 167. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 150-
350 m. 28-30 Aug. 1929. Killip & Smith 28574 (HT: US; IT:
LE.) °

furcatipilosum Yuncker in Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 112, fig. 85. 1950. TYPE: Venezuela, Mérida...along
Quebrada of Cuesta del Barro and Mesa del Trapiche...between
Canagua and El Molino. El. 2530-2715 m. 11 May 1944.
Steyermark 56472 (HT: F 1222765; IT: ILL).

40 Pb T Ob OGaA Vol. 58, No. 1

furiae Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 49. 1936
without a Latin diagnosis). TYPE: Colombia, Bogota,
Quebrada de la Vieja. El. 2750 m. 22 April 1932.
Cuatrecasas 2421 (HT: ILL).-- [= P. artanthe (fide Trel. &
Yuncker, 1950) ]

gamboanum (C.DC.) C.DC. var. yapense Trel. in Woodson & Schery,
Ann. Mo. Bot. Gard. 27: 291. 1940. TYPE: Panama, Darién,
vicinity of Yape. El. ca 30 m. 4 Oct. 1938. Allen 854 (HT:
ILL).-- [= P. villiramlum C.DC. (fide Yuncker, 1950)) _

gaudichaudianum Kunth var. minor Yuncker, Bol. Inst. Bot. (Sao
Paulo) 3: 39, fig. 33. 1966. TYPE: Brazil, Rio de Janeiro,
Guanabara, estrada do Corcovado. 23 Sep. 1958. Pereira,
Liene, Benjamin & Duarte 4337 (HT: HB; PT at ILL: Dusen
8234).-- [This variety upheld by Yuncker, 1972]

gentlei Trel. ex Standley in Standley & Record, Field Mus. Publ.
Bot. 12: 407. 1936. TYPE: British Honduras (Belize),
Corozal, San Andrés. Gentle 1077 (HT: MICH; IT: ILL).--
(= P. aeruginosibaccum (fide Standley & Steyerm., 1952);

= P, jacquemontianum (cf. Burger, 1971) ]

gigas Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 292.
1940. TYPE: Panama, Chiriqu{, vicinity of Casita Alta,
Voledn de Chiriquf. El. 1500-2000 m. 1938. Woodson, Allen
& Seibert 846 (HT: ILL).-- [= P. imperiale (fide Yuncker,
1950) J

gilvescens Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
167. 1936. TYPE: Peru, Junf{n, Cahuapanas, on Rfo Pichis.
El. 340 m. 20-21 July 1929. Killip & Smith 26784 (HT: US;
TT °ELL)i

glaberrimicaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 168. 1936. TYPE: Peru, Ayacucho, Estrella. El. 500 m.
8, 14 May 1929. Killip & Smith 22624 (HT: US; IT: ILL).

glabribaccum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
168. 1936. TYPE: Peru, Junin, La Merced. El. 600 m. 10-24
Aug. 1923. Macbride 5335 (HT: F 536362; IT: ILL).

glandulosissimum Yuncker, Bol. Inst. Bot. (S80 Paulo) 3: 40,
fig. 34. 1966. TYPE: Brazil, Pard, Belém. Oct.-Nov. 1929.
Killip & Smith 30260 (HT: US 1356446; PT at ILL: Killip &
Smith 30218).-- [This species upheld by Yuncker, 1972]

1985 Jones, Annotated catalogue of type specimens 4]

gleasonii Yuncker in Maguire & Collaborators, Bull. Torrey Bot.
Club 75: 287. 1948. TYPE: British Guiana (Guyana),
Rockstone. Gleason 857 (HT: NY; PT at ILL: Suriname, Maguire
24112, 24170).-- [This species recognized by Trel. & Yuncker,
1950 and Yuncker, 1957]

goergeri Trel. in Standley, Field Mus. Publ. Bot. 18: 344. 1937.
TYPE: Costa Rica, Finca Castilla, Rfo Reventazon. El. 30 m.
27 July 1936. Dodge & Goerger 9421 (HT: MO; IT: ILL).--
[= P. friedrichsthalii (fide Burger, 1971) ]

gonagricum Trel., Contr. U.S. Nat. Herb. 26(4): 171. 1929.
TYPE: Costa Rica, Heredia, Yerba Buena, above San Isidro.
El. 2000 m. Feb. 1928. Standley & Valerio 49082 (HT: US
1307566; IT: ILL).-- [= P. hispidum (fide Burger, 1971) ]

gorgonae-insulae Trel. & Yuncker, Piperaceae no. S. Amer, 1:
331, fig. 299. 1950. TYPE: Colombia, Narino, south end of
Gorgona Island. 9 Feb. 1939. Killip & Garcfa 33095 (HT: US
1770088; IT: ILL).

gorgonillense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 406,
fig. 368. 1950. TYPE: Colombia, Narino, Gorgonilla Island.
El. 130-200 m. 8 Feb. 1939. Killip & Garcfa 33117 (HT: US
1770108; IT: ILL; PT at ILL: Killip & Garcia 33088).

gracilipedunculum Trel., Contr. U.S. Nat. Herb. 26(4): 148.
1929. TYPE: Costa Rica, Alajuela, Fraijanes. El. 1500-
1700 m. Feb. 1926. Standley & Torres 47519 (HT: US 1307496;
PT at ILL: Standley 42095; Standley & Torres 47582; Standley
& Valerio 49198).-- [= P. decurrens C.DC. (fide Burger,
1971) ]

gracillimum Trel. in Standley, Field Mus. Publ. Bot. 10: 159.
1931. Type locality: Honduras, Lancetilla Valley. "In wet
forest high on the hills above Lancetilla." No type
indicated; probably lectotypification needed (authentic
specimen from area of type locality at ILL: Standley 54684).

gracillimum var. dantoense Trel. in Yuncker, Field Mus. Publ.
Bot. 9: 279. 1940. TYPE: Honduras, Atlantida, R{o Danto, Mt.
Cangrejal. El. 180 m. 16 July 1938. Yuncker, Koepper &
Wagner 8485 (HT: ILL).

grahami Trel., Bull. Torrey Bot. Club 60:° 4877. 1933.- TYPE:
British Guiana (Guyana), Kyk-over-al, junction of the
Mazaruni and Cuyuni rivers. 9 July 1924. Graham 227 (HT:
CM; IT: ILL).-- [= P. dilatatum (fide Trel. & Yuncker, 1950
and Yuncker, 1957) ]

42 PwuY TF OL. 0. & JA Vol. 58, No. 1

granuligerum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
169. 1936. TYPE: Peru, Loreto, San Antonio, Alto Rfo Itaya.
El. 145 m. Sep.-Oct. 1929. Williams 3459 (HT: F 622875;

IT2, Dh)

gratum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 170.
1936. TYPE: Peru, Loreto, Soledad. El. 110 m. 20-22 Sep.
19293, Kil dip & Smith 29785 (HT: US; IT: ILL).

griseocaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
170. 1936. TYPE: Peru, Jun{in, La Merced. El. 1200 m.
Aug.-Sep. 1923. Macbride 5723 (HT: F 536755; IT: ILL; PT at
ILL: Macbride 5595).

griseo-pubens Trel., Contr. U.S. Nat. Herb. 26(4): 176. 1929.
TYPE: Costa Rica, Guanacaste, L{bano. El. 300 m. 15 Jan.
1926. Standley & Valerio 44879 (HT: US 1307328; IT: ILL).--
[= P. pseudo-fuligineum (fide Burger, 1971) ]

griseo-reticulatum Trel. in Feddes Repert. Spec. Nov. Regni
Veg. 24: 362. 1928. TYPE: Haiti, Croix des Bouquets, La
Selle. El. 1200 m. 14 March 1927. Ekman 7852 (HT: ILL).

griseovenosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 170. 1936. TYPE: Peru, Loreto, Puerto Arturo.
El. 135 m. 24-25 Aug. 1929. Killip & Smith 27803 (HT: US;
ITI LG),

guabinachanum [stet!] Trel. in Feddes Repert. Spec. Nov. Regni
Veg. 23: 5, 9. 1926. TYPE: Cuba, Pinar del Rfo, Guabimacho.
6 March 1924. Ekman 18638 (HT and IT: ILL; PT at ILL: Ekman
17970).

guapense Yuncker in Trel. & Yuncker, Piperaceae no. S.. Amer. 1:
28, fig. 19. 1950. TYPE: Colombia, Antioquia, Guapa.
El. ca 40 m. 23 April 1945. Haught 4594 (HT: US 17091 63;
573 1L1).

gurupanum Yuncker, Bol. Inst. Bot. (S¥o Paulo) 3: 100, fig. 87.
1966. TYPE: Brazil, Parad, Rio Amazonas, Gurupa. 3 Nov.
1929. Killip & Smith 30611 /(HT27US 1463625; -IT.sZILL).=
[This species upheld by Yuncker, 1973]

hanckeli Trel. in Standley, Field Mus. Publ. Bot. 18: 345.
1937. TYPE: Costa Rica, Guanacaste, upper slopes of Cerro
San José de L{bano. El. 500-600 m. 15 Feb. 1930. Dodge,
Hanckel & Thomas 6384 (HT: GH; IT: ILL).-- [= P.
chrysostachyum C.DC. (fide Burger, 1971) ]

1985 Jones, Annotated catalogue of type specimens 43

hastularum Yuncker in Woodson & Schery, Ann. Mo. Bot. Gard. 37:
21, fig. 5. 1950. TYPE: Colombia, Chocd, Bahfa Solano...
along Quebrada Jellita. El. 50-100 m. 22 Feb. 1939. Killip
& Garcfa 33568 (HT: US; IT: ILL).-- [This species recognized
by Trel. & Yuncker, 1950]

hastularum var. longipetiolatum Trel. & Yuncker, Piperaceae no.
S. Amer. 1: 137. 1950. TYPE: Colombia, El Valle, Rfo Digua
Valley, Piedra de Moler. El. 900-1180 m. Aug. 1943.
Cuatrecasas 15106 [HT (or LT): US; IT (or ILT): ILL].

haughtii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 263, fig.
226. 1950. TYPE: Colombia, Santander, Magdalena Valley.
9 July 1935. Haught 1831 (HT: US 1661858; IT: ILL; PT at
ILL: Killip, Cuatrecasas & Dryander 39205).

heraldi Trel. var. amplius Trel. in Woodson & Schery, Ann. Mo.
Bot. Gard. 27: 292. 1940. TYPE: Panama, Chiriqui, Finca
Lérida to Boquete. El. 1300-1700 m. July 1938. Woodson,
Allen & Seibert 1095 (HT: ILL).-- [= P. auritum (fide
Yuncker, 1950 and 1957)]

heraldi var. cocleanum Trel. in Woodson & Schery, Ann. Mo. Bot.
Gard. 27: 292. 1940. TYPE: Panama, Coclé, El Valle.
El. 600-1000 m. 8 Dec. 1938. Allen 1192 (HT and IT: ILL).--
(= P. auritum (fide Yuncker, 1950 and 1957) ]

hermannii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 397, fig.
358. 1950. TYPE: Colombia, Rio Vichada, Vichada. Hermann
10998 (HT: US 1906217; PT at ILL: Colombia, Haught 2759;
Venezuela, Williams 14681).

heterocarpum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
172. 1936. TYPE: Peru, Loreto, Yurimaguas to Balsapuerto.
El. 135-150 m. 26-31 Aug. 1929. Killip & Smith 28327 (HT:
Use 30s TLE).

neterotrichum C.DC. var. porrectifolium Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 290. 1950. TYPE: Colombia, El
Valle, Dagua Valley. 22 Sep. 1922. Killip 11529 (HT: US
1143235; IT: ILL).

hispaniolae Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
305, 309. 1927. TYPE: Santo Domingo. Wright, Parry &
Brummel 445 (HT: US; PT at ILL: Leonard 7643).

hispidiseptum Trel., J. Wash. Acad. Sci. 19: 332. 1929. TYPE:
Honduras, Ablant bdo. Lancetilla Valley near Tela. El. 100 m.
8 Dec. 1927. SNES 52715 (HT: F; IT: ILL; PT at ILL:
Standley 55660).-- [= P . fraguanum (fide Standley & Steyerm.,
195273 = <P P. sancti- felicis (fide Burger, 1971)]

44

POT. PLer LA Vol. 58, No. 1

hispidum Sw. var. lanceolatum Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 274, fig. 237. 1950. TYPE: Colombia, Cauca, Rfo
Micay, Caliche. 26 Feb. 1943. Cuatrecasas 14180 [HT: US
1852590; PT at ILL: Ecuador, Mexia 7330 (2 sheets), 8413).

holguinianum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
4, 7. 1926. TYPE: Cuba, Oriente, Holguin. 29 Oct. 1914.
Ekman 3303 (HT: ?; IT: ILL).

hondonadense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 387,
fig. 347. 1950. TYPE: Colombia, Cundinamarca, Salto de
Tequendania...La Hondonada. 2 Oct. 1938. Cuatrecasas 244
(HT: US 1796828; fragment of type (single leaf) at ILL].

nondonadense var. javieri Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 388. 1950. TYPE: Colombia, Cundinamarca, Aguadita.
28 Nov. 1939. Javier 38 (HT: ILL).

hostmannianum (Miq.) C.DC. in DC. var. glabrirameum Trel. &
Yuncker, Piperaceae no. S. Amer. 1: 307. 1950. TYPE:
Venezuela, Amazonas, Tamatama, upper Orinoco. El. 121 m.
4 May 1942. Williams 15129 (HT: US 1877446; IT: ILL; PT
at ILL: Killip 37311).

hostmannianum [var.] ® ramiflorum (Miq.) C.DC. in DC., Prodromus
16(1): 287. 1869 [based on Artanthe ramiflorum Miq. ex C.DC.
in DC., pro syn.]. TYPE: Suriname, Suriname R., near
Victoria. Dec. 1843. Kappler 1397 (HT: ?; IT: ILL).-- [This
variety recognized by Yuncker, 1957]

huacachianum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
172. 1936. TYPE: Peru, Huanuco, Huacachi near Muna.
El. 1950 m. May-June 1923. Macbride 4156 (erroneously cited
as "4167") (HT: F 535230; IT: ILL).-- [Type number verified
by Burger (pers. comm.) ]

huacapistanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 173. 1936. TYPE: Peru, Junin, Huacapistana. 6 Dec.
1924. Stevens 77 (HT: ILL; PT at ILL: Stevens 79).

huallaganum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
173. 1936. TYPE: Peru, Huanuco, Huallaga Canyon below Rfo
Santo Domingo. El. 1200 m. 3 June 1923. Macbride 4235 (HT:
P° 53531430 Tei).

huantanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
173. 1936. TYPE: Peru, Ayacucho, Estrella, between Huanta
and Rio Apurimac. El. 500 m, Killip & Smith 23074 (HT: US;
IT: ILL; PT at ILL: Killip & Smith 25193).

1985 Jones, Annotated catalogue of type specimens 45

humoense Trel. in Standley, Field Mus. Publ. Bot. 18: 346. 1937.
TYPE: Costa Rica, Cartago, Pejivalle Farm above Rfo Humo.
El. 800 m. 25 Sep. 1929. Dodge & Thomas 4371 (HT: GH; IT:
ILL).-- [= P. hispidum (fide Burger, 1971)]

humorigaudens Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
293. 1940. TYPE: Panama, Coclé, N rim of El Valle de Antén.
21 May 1939. Allen 1833 (HT: ILL).-- [This species
recognized by Yuncker, 1950]

* ibaguense Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 50. 1936
(without a Latin diagnosis). TYPE: Colombia, Tolima, Ibagué.
El. 1000-1500 m. 13 Aug. 1922. Killip 9724 (HT, or
LECTOTYPE designated herein: ILL; PT at ILL: Cuatrecasas
2419).-- [= P. aequale (fide Trel. & Yuncker, 1950; Yuncker,
1950 and 1957)]

ignacioanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 280,
fig. 244. 1950. TYPE: Ecuador, Guayas, Tenguel, San Ignacio.
17 Nov. 1919. Holmgren 5 (HT: US 1481980; IT: ILL).

* illudens Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 50. 1936
(without a Latin diagnosis). TYPE: Colombia, Ibagué.
El. 1350 m. 10 May 1932. Cuatrecasas 2424 (HT, or LECTOTYPE
designated herein: ILL).-- [= P. aduncum (fide Trel. &
Yuneker, 1950; Yuncker, 1950, 1955) ]

immite Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 174.
1936. TYPE: Peru, Junfn, La Merced. El. 700 m. May-June
1929. Killip & Smith 23524 (HT: US; IT: ILL; PT at ILL:
Killip & Smith 23437, 23468, 24074).

immutatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
175. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 350-550 m.
29 Aug. 1929. Killip & Smith 28536 (HT: US; IT: ILL).

imperspicuibracteum Trel., J. Wash. Acad. Sci. 19: 332. 1929.
TYPE: Honduras, Atlantida, near Tela. 19 Jan. 1928.
Standley 54770 (HT: F; IT: ILL; PT at ILL: Standley 56602).--
[= P. fraguanum (fide Standley & Steyerm., 1952); = P.
sancti-felicis (fide Burger, 1971)]

inauspicatum Trel., Ciencia (Mexico) 2: 207. 1941. TYPE: Peru,
Chanchamayo. Isern 2184 (HT: MA?; IT: ILL).

incanum Trel. in Standley, J. Wash. Acad. Sei. 13: 436. 1923.
TYPE: El Salvador, vicinity of San Salvador. El. 650-850 m.
20 Dec. 1921-4 Jan. 1922. Standley 19129 (HT: ILL).

incitum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 175.
1936. TYPE: Peru, Junfin, Huacapistana. El. 1800 m. 6 June
1929. Killip & Smith 24304 (HT: US; IT: ILL).

46 Pier TO Or Gs PA Vol. 58, No. 1

inclemens Trel. & Yuncker, Piperaceae no. S. Amer. 1: 321, fig.
286. 1950. TYPE: Ecuador, Esmeraldas, Parroquia Concepcion,
Playa Rica. El. 105 m. 7 Dec. 1936. Mexia 8418 (HT: US
1778442; IT: ILL).

inconsumptum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 346.
1938. TYPE: Honduras, Cortés, forest near Paral, Lake Yojoa.
El. 630 m. 30 July 1934. Yuncker 4871 (HT: ILL; PT at ILL:
Yuncker 4874).

incurvum Sieber ex Schultes, L. Mant. Syst. Veg. 1: 238. 1822.
TYPE: Martinique. Sieber 254 [HT: B?; IT: (marked "Cotype"):
ILL]).-- [This is the type species of Sarcorhachis Trel.,
Contr. U.S. Nat. Herb. 26: 16. 1927, emend. Steyerm.,
Pittieria 3: 29. 1971 = S. incurva (Sieber ex Schultes)
Trel. (see Howard, 1973) ]

indianonum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
176. 1936. TYPE: Peru, Loreto, Iquitos, Fundo Indiano.
El. 110 m. .21 Jan.°1932. Mexia 6396 (HT: UC; ITs ILL; PT at
ILL: Mexia 6441),

indiciflexum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
176. 1936. TYPE: Peru, Junf{n, Huacapistana. El. 1800-
2400 m. 5-8 June 1929. Killip & Smith 24507 (HT: US; IT:
ite es

indignum Trel., 338 Wash. Acad! ‘Sei. 19.333. 1929.: c TYPE:
Honduras, Comayagua, near Siguatepeque. El. 1100 m.
Feb. 1928. Standley 55990 (HT: F; IT: ILL).-- [This species
recognized by Standley & Steyerm., 1952]

infidele Trel. & Yuncker, Piperaceae no. S. Amer. 1: 159, fig.
138. 1950. TYPE: Colombia, Putumayo, near the Colombia-
Ecuador boundary at the confluence of the San Miguel and
Conejo rivers. El. 300 m. 9 Dec. 1940. Cuatrecasas 10921
(HT: US 1799102; IT: ILL).

* insignilaminum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 17,
fig. 5. 1950. TYPE: Colombia, Antioquia, Yarumal.
Oct. 1941. Tomas Alberto 1628 [LECTOTYPE designated herein
(Fig. Sie Tide ALY ALL 4,

insolens Trel., Contr. U.S. Nat. Herb. 26(4): 156. 1929. TYPE:
Costa Rica, San José, Santa Marfa de Dota. El. 1500-1800 m.
Standley & Valerio 43306 (HT: US 1307288; IT: ILL).-- L= Ps
biauritum C,.DC. (cf. Burger, 1971)]

1985 Jones, Annotated catalogue of type specimens 47

instabilipes Trel. in Standley, Field Mus. Publ. Bot. 17: 232.
1937. TYPE: Guatemala, Petén, Santa Teresa, Subin River.
14 April 1933. Lundell 2776 (HT: ILL).-- [This species
recognized by Standley & Steyerm., 1952]

insulicolum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
293. 1940. TYPE: Panama, Prov. Panam&, Isla Taboga. 24 June
1938. Woodson, Allen & Seibert 1527 (HT: ILL).-- [= P.

taboganum (fide Yuncker, 1950); = P. pseudo-fuligineum (cf.
Burger, 1971)]

interitum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
176. 1936. TYPE: Peru, Junin, Pichis Trail, Santa Rosa.
El. 625-900 m. 6, 7 July 1929. Killip & Smith 26150 (HT:
US; ITs ILL);

intersitum Trel. in Daniel, Una ascensién Cerro "La Vieja", 12.
1940 (without a Latin diagnosis); Caldasia 1: 86. 1940.
TYPE: Colombia, Antioquia, Porcecito. Daniel 890 (HT: US;
fragments of type at ILL).-- [= P. aduncum (fide Trel. &
Yuncker, 1950; Yuncker, 1950, 1955 and 1957) ]

intonsum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
177. 1936. TYPE: Peru, Junin, La Merced. El. 600 m.
Macbride 5445 (HT: F 536479; PT at ILL: Williams 6829).

iquitosense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
177. 1936. TYPE: Peru, Loreto, Iquitos. July 1929.
Williams 1432 (HT: F 617448; IT: ILL; PT at ILL: Killip &
Smith 26275, 26899, 28181; Williams 1332).

irazuanum C.DC. var. suborbiculatum Trel., Contr. U.S. Nat.
Herb. 26(4): 137. 1929. TYPE: Costa Rica, San José, near
Finca La Cima, above Los Lotes, N of El Copey. El. 2100-
2400 m. Dec. 1925. Standley 42681 (HT: US 1307274; IT:
ILL; PT at ILL: Standley 41951).-- [= P. lacunosum Kunth
(fide Burger, 1971) ]

irrasum Trel., Contr. U.S. Nat. Herb. 26(4): 151. 1929. TYPE:
Costa Rica, Cartago, El Muneco, Rio Navarro. El. 1400-
1500 m. 6, 7 March 1926. Standley & Valerio 50961 (HT: US
1307609; IT: ILL).-- [= P. imperiale (fide Burger, 1971)]

* jsernii Trel., Ciencia (Mexico) 2: 207. 1941. TYPE: Ecuador, La
Coca. July 1865. Isern 1392 (HT, or LECTOTYPE designated
herein: ILL; IT or ILT: MA).-- [= P. archidonense (fide Trel.
& Yuncker, 1950) ]

48 PHY¥TOLOG.IA Vol. 58, No. 1

itayanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
177. 1936. TYPE: Peru, Loreto, Parafso, Alto Rfio Itaya.
El. 145 m. Williams 3320 (HT: F 623464; PT at ILL: Killip &
Smith 27245, 28130, 29617; Williams 3509).

itayanum var. antoniense Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 178. 1936. TYPE: Peru, Loreto, San Antonio.
El. 110 m. 18 Sep. 1929. Killip & Smith 29450 (HT: US; IT:
ILL; PT at ILL: Killip & Smith 27324).

itayanum var. pichisense Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 178. 1936. TYPE: Peru, Junin, Cahuapanas, Rio
Pichis. El. 340 m. 20, 21 July 1929. Killip & Smith 26738
(HT: US; IT: ILL; PT at ILL: Killip & Smith 26260, 26750).

itayanum var. san-juanum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 178 1936. TYPE: Peru, Loreto, Iquitos, San
Juan. El. 120 m. Oct. 1929. Williams 3720 (HT: F 618346;
it<,; ILE);

jaboncillanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 233,
fig. 200. 1950. TYPE: Colombia, Santander, Rio Surata
Valley, between Bucaramanga and El Jaboncillo. El. 800-
1500 m. 2 Jan. 1927. Killip & Smith 16388 (HT: US 1352133;
E75: ILL);

jaculifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
178. 1936. TYPE: Peru, Junin, La Merced. El. 700 m.
May-June 1929. Killip & Smith 24060 (HT: US; IT: ILL).

jaliscanum S. Watson, Proc. Amer. Acad. Arts 26: 145. 1891.
TYPE: Mexico, canons near Guadalajara. June 1886. Palmer
122, in flower. Dec. 1888. Pringle 2153, in young fruit--
syntypes, probably to be lectotypified (fragment of Palmer
122 at ILL).-- [This species recognized by Standley, 1920]

jamaoanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 29: 29.
1931. TYPE: Santo Domingo, Colonia Jamao, Moca. El. 900 m.
21 May 1925. Ekman 12576 (HT: ILL).

jericoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: $93 T wn fk
359. 1950. TYPE: Colombia, Antioquia, Jeric6. Dec. 1940.
Alejandro 2730 [HT: US 1830390; IT: ILL (2 sheets) ].

ubimarginatum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 8, fig.
6. 1966. TYPE: Brazil, Rio de Janeiro, Maud. 28 Jan. 1897.
Ule 4366 (HT: R; IT: ILL).-- [This species upheld by Yuncker,
1972]

1985 Jones, Annotated catalogue of type specimens 49

kantetulense Trel. ex Standley in Standley & Record, Field Mus.
Publ. Bot. 12: 408. 1936. TYPE: Guatemala, Petén, Kantetul.
2 May 1933. Lundell 3173 (HT: MICH; IT: ILL).-- [= P.
aeruginosibaccum (fide Standley & Steyerm., 1952); i

jacquemontianum (cf. Burger, 1971) ]

kantetulense var. gentlei Trel. ex Standley in Standley &
Record, Field Mus. Publ. Bot. 12: 408. 1936. TYPE: British
Honduras (Belize), Northern River, Prospecto. Nov. 1933.
Gentle 947 (HT: MICH; IT: ILL).-- [= P. aeruginosibaccum
(fide Standley & Steyerm., 1952; = P. jacquemontianum (cf.
Burger, 1971)]

kerberi C.DC., Annuaires Conserv. Jard. Bot. Genéve 2: 258.
1898. TYPE: Mexico, Veracruz, Cordoba. Kerber 26 [HT: LE?;
fragment of type (single leaf) at ILL].

kimpitirikianum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 179. 1936. TYPE: Peru, Ayacucho, Rfo Apurimac Valley,
near Kimpitiriki. El. 400 m. 10 May 1929. Killip & Smith
22647 CHT: US3 IT:° IEE).

koepperi Trel. in Yuncker, Field Mus. Publ. Bot. 9: 279. 1940.
TYPE: Honduras, Yoro, Aguan Valley near Medina. El. 195 m.
Yuneker, Koepper & Wagner 8631 (HT: ILL).

konkintoense Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
293. 1940. TYPE: Panama, Bocas del Toro, between Finca St.
Louis and Konkintoe. El. 10-50 m. Aug. 1938. Woodson,
Allen & Seibert 1894 (HT: ILL).-- [= P. hispidum (fide
Yuncker, 1950, 1955 and 1957); = P. sancti-felicis (fide
Burger, 1971) ]

krukoffii Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 24, fig 18.
1966. TYPE: Brazil, Amazonas, Mun. Humayta, near Livramento,
on Rfo Livramento. Oct.-Nov. 1934. Krukoff 6892 (HT: NY;
IT: ILL).-- [This species upheld by Yuncker, 1972]

laceratibracteum Trel. in Yuncker, Field Mus. Publ. Bot. 17:
346. 1938. TYPE: Honduras, Comayagua, El Achote, above
Siguatepeque. El. 1350 m. 30 July 1936. Yuncker, Dawson &
Youse 6226 (HT: ILL).

lacunosum Kunth var. angustifolium Trel. & Yuncker, Piperaceae
no. S. Amer. 1: 21. 1950. TYPE: Colombia, Caldas, San Felfx
near Salamina. El. 3000 m. July 1944. Tomas Alberto 2459
(HT: US 1564178; PT at ILL: Garcfa Barriga 11882; Pennell
9193). <7 he

50

PHHATO 40:63, 4 Vol. 58, No. 1

lacunosum var. glabricaule Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 21. 1950. TYPE: Colombia, Huila, Balsillas.
El. 2000-2100 m. 3-6 Aug. 1917. Rusby & Pennell 709 (HT: US
1041445; PT at ILL: Cuatrecasas 19215).

la-doradense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 244,
fig. 212. 1950. TYPE: Colombia, Caldas, Quebrada Pontand, ca
18 km W of La Dorado. El. 400 m. 30 Dec. 1936. Haught 2140
(HT: US 1742379; IT: ILL). ,

laedans Trel. in Yuncker, Field Mus, Publ. Bot. 17: 347. 1938.
TYPE: Honduras, Comayagua, El Achote, above Siguatepeque.
El. 1350 m. 13 July 1936. Yuncker, Dawson & Youse 5821 (HT:
ILL).

laeteviride Ekman ex Trel. in Feddes Repert. Spec. Nov. Regni
Veg. 29: 29. 1931. TYPE: Santo Domingo, San Crist6bal,
Cambita, Arroyo Blanco. 27 Jan. 1929. Ekman 11339 (HT:
ILL).

lagenaebaccum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 179. 1936. TYPE: Peru, Junfn, Puerto Yessup.
El. 400 m. 10-12 July 1929. Killip & Smith 26249 (HT: US;
IT: ILL; PT at ILL: Killip & Smith 26175).

laguna-cochanum Trel. & Yuneker, Piperaceae no. S. Amer. 1: 132,
fig. 108. 1950. TYPE: Colombia, Putumayo, south side of
Laguna de la Cocha, Quebrada de Santa Lucfa. El. 2850 m.

8 Jan. 1941. Cuatrecasas 11844 (HT: US 1798192; PT at ILL:
Ewan 16344, 16601).

lamasense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
180. 1936. TYPE: Peru, San Mart{n, Lamas near Tarapoto. El.
840 m. Dec. 1929. Williams 6401 (HT: F 623619; IT: ILL).

lancetillanum Trel., J. Wash. Acad. Sci. 19: 333. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
5 March 1928. Standley 56552 (HT: F 581987; IT: ILL; PT at
ILL: Standley 53231).

lanosibracteum Trel., Contr. U.S. Nat. Herb. 26(4): 182. 1929.
TYPE: Costa Rica, Cartago, Rfo Reventado, N of Cartago.
El. 1500 m. 26 Feb. 1926. Standley & Valerio 49457 (HT: US
1307575; IT: ILL; PT at ILL: Standley & Valerio 49435,
49482).-- [= P. hispidum (fide Burger, 1971) ]

lanosicaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
181. 1936. TYPE: Peru, Junfn, San Ramon, Schunke Hacienda.
El. 1400-1700 m. 8-12 June 1929. Killip & Smith 24723 (HT:
US; IT: ILL; PT at ILL: Killip & Smith 25646).-- [See note

under P. enenyasense ]

1985 Jones, Annotated catalogue of type specimens 5]

late-ovatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
181. 1936. TYPE: Peru, San Martf{n, San Roque. El. 1350-
1500 m. Jan.-Feb. 1930. Williams 7083 (HT: F 623091; IT:
TLL).

laterifissum Trel., J. Wash. Acad. Sci. 19: 333. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 300 m.
31 Dec. 1927. Standley 53943 (HT: F 584143; IT: ILL).

ledezmae Trel. & Yuncker, Piperaceae no. S. Amer. 1: 82, fig.
67. 1950. TYPE: Colombia, Chocé, Corcovada region, upper R{io
San Juan. El. 200-275 m. 24, 25 April 1939. Killip &
Ledezma 35314 (HT: US 1772051; IT: ILL; PT at ILL: Archer
2007).-- [= affin. P. ottoniaefoliumC.DC. (cf. Trel. &
Yuncker, 1950)]

lenticellosum C.DC. var. scandens Yuncker in Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 89. 1950. TYPE: Colombia, El
Valle, valley of Rio Cali, between Puente Soffa and
Yanaconas. El. 1580-1730 m. 6 Nov. 1944. Cuatrecasas 18704
(HT: US 1854284; IT: ILL).

leucofustum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
182. 1936. TYPE: Peru, Junin, La Merced. El. 700 m.
May-June 1929. Killip & Smith 23736 (HT: US; IT: ILE).

leucophaeocaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 183. 1936. TYPE: Peru, Loreto, Lower Rio Nanay.
May-June 1929. Williams 301 (HT: F 603748; IT: ILL; PT at
ILL: Killip & Smith 24629, 27037, 27469).

leucophaeum Trel. in Macbr., Field Mus, Publ. Bot. 13(No. 357):
183. 1936. TYPE: Peru, San Martin, Tarapoto. El. 750 m.
Dec. 1929. Williams 6086 (HT: F 618084; IT: ILL).

leucophlebium Trel., Contr. U.S. Nat. Herb. 26(4): 176. 1929.
TYPE: Costa Rica, Guanacaste, Tilaran. El. 600 m.
Jan. 1926. Standley & Valerio 44277 [HT: US 1307301; IT:

ILL; probable PT at ILL: Standley & Valerio 45708 (cited as
45908) ].-- [= P. villiramulum C.DC. (fide Burger, 1971) ]

leucostachyum Trel, & Yuncker, Piperaceae no. S. Amer. 1: 184,
fig. 143. 1950. TYPE: Colombia, Santander, Rfo Surata valley
above Surata. El. 2000-2300 m. 5, 6 Jan. 1927. Killip &
Smith 16575 (HT: US 1352301; IT: ILL; PT at ILL: Killip &
Smith 16694).

52

py TOY LONG sd oA Vol. 58, No. 1

levilimbum Trel., J. Wash. Acad. Sci. 19: 333. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
22 Dec. 1927. Standley 53483 (HT: F 583185; IT: ILL; PT at

ILL: Standley 53839, 55263).

ligatispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
183. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 28183 (HT: US; IT: ILL).

lincolnense Trel. in Standley, Field Mus. Publ. Bot. 18: 347.
1937. TYPE: Panama, Bocas del Toro, Changuinola Valley,
Lincoln Creek. 3 March 1924. Dunlap 484 (HT: F; IT: ILL).--
[= P. glabrescens (fide Burger, 1971) ]

lineativillosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 321,

fig. 287. 1950. TYPE: Colombia, El Valle, north shore of
Buenaventura Bay, near sea level. 3 June 1944. Killip &
Cuatrecasas 38734 (HT: US 1856798; IT: ILL).

lineatum Ruiz & PavOn var. hirtipetiolatum Trel. in Macbr.,
Field Mus. Publ. Bot. 13(No. 357): 184. 1936. TYPE: Peru,
Hudnuco, Huacachi near Muna. El. 1950 m. May-June 1923.
Macbride 4182 (HT: F 535259; IT: ILL).

lineatum var. leucolepidum Trel. in Macbr., Field Mus, Publ.
Bot. 13(No. 357): 184. 1936. TYPE: Peru, Junin, Carrapa.
El. 1000 m. May 1929. Killip & Smith 22495 (HT: US; IT:
ILL; PT at ILL: Killip & Smith 22451, 22565).

linguifolium Ham. var. brevifolium Trel. in Feddes Repert. Spec.
Nov. Regni Veg. 24: 362. 1928. TYPE: Haiti, Anses a Pitres,
La Selle. 7 Sep. 1926. Ekman 6912 (HT: ILL).

liratinerve Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 24:
186. 1937. TYPE: Panama, Chiriqu{, valley of the upper Rio
Chiriqu{ Viejo, vicinity of Monte Lirio. El. 1300-1900 m.
June 1935. Seibert 158 (HT: MO; IT: ILL).-- [= P.
pseudo-lanceaefolium Trel. (fide Yuncker, 1950); = P.
lanceaefolium Kunth (fide Burger, 1971) ]

llatanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
165. 1936. TYPE: Peru, Huanuco, Llata, Rio Marandén Canyon.
El. 2100 m. 21 Aug. 1922. Macbride & Featherstone 2302 (HT:
F 518785; IT: ILL).

loefgrenii Yuncker, Bol. Inst. Bot. (Sao Paul o}-s2710-7f4c6.. 8.
1966. TYPE: Brazil, Sado Paulo, Araraquara. 28 Sep. 1888.
Loefgren 447 (HT: ILL) .-- [This species upheld by Yuncker,
1972]

1985 Jones, Annotated catalogue of type specimens 53

longegerminatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 185. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m.
1-5 Sep. 1929. Killip & Smith 28839 (HT: US; IT: ILL; PT
at ILL: Killip & Smith 28439).

longichaetum Yuncker, Lilloa 27: 127, pl. 15. [1953] 1955.
TYPE: Bolivia, La Paz, Challana region. £1. 800 m.
Nov. 1933. Cardenas 1325 (HT: ILL). .

longispicum C.DC. var. villipetiolatum Yuncker in Trel. &
Yuncker, Piperaceae no. S. Amer. 1: 140. 1950. TYPE:

Colombia, Narino, Cerro de Puerto above Puerto. El. 2320 m.
July 1944. Ewan 15947 (HT: USNA; IT: ILL).

longivillosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 35,
fig. 27. 1950. TYPE: Colombia, Choco, Rfo Catripe.
El. 100 m. 17 Jan. 1947. Haught 5445 (HT: US 1709497; PT at
ILL: Killip & Garcfa 33491).

loretoanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
186. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 27555 (HT: US; IT: ILL; PT at
ILL: Williams 4758).

losoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 345, fig.
317. 1950. TYPE: Colombia, Norte de Santander, Loso.
El. 2200-2400 m. 6, 7 March 1927. Killip & Smith 20436 (HT:
US 1355482; IT: ILL; PT at ILL: Killip & Smith 16780, 18844,
18907, 18922).

lucaeanum Kunth var. grandifolium Yuncker, Bol. Inst. Bot. (S30
Paulo) 3: 104, fig. 90. 1966. TYPE: Brazil, Parana, Porto de
Cima. 2 Jan. 1914. Dusen 14304 (HT: S; PT at ILL: Dusen
6905, 7025).-- [This variety upheld by Yuncker, 1973]

lundellianum Trel. in Standley, Field Mus. Publ. Bot. 17: 232.
1937. TYPE: Guatemala, Petén, La Libertad. 31 May 1933.
Lundell 3524 (HT: ILL-- the specimen labelled "P.
lundelliifolium, Type").-- [= P. amalago (fide Standley &
Steyerm., 1952) ]

luteobaccum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 29:
29. 1931. TYPE: Santo Domingo, Loma La Campana. E1. 800 m.
11 Feb. 1929. Ekman 11527 (HT: ILL).

macbrideanum Trel. in Macbr., Field Mus. Publ. Bot. 13(2): 187.
1936. TYPE: Peru, Junfn, La Merced. El. 600 m. 10-24 Aug.
1923. Macbride 5516 (HT: F 536556; IT: ILL).

54

Pw’ t Oko 6 oom Vol. 58, No. 1

macedoi Yuncker forma longispicum Yuncker, Bol. Inst. Bot. (Sao
Paulo) 3: 52. 1966. TYPE: Brazil, Minas Gerais, Diamantina,
Barao. El. 1180 m. 19 May 1931. Mexia 5859 (HT: US
1618265; IT: ILL).-- [This variety upheld by Yuncker, 1972]

macerispicum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 404,
fig. 366. 1950. TYPE: Colombia, Caqueta, Florencia.
El. 400 m. 29 March 1940. Cuatrecasas 8807 (HT: US 1795402;
PT at ILL: Klug 1706).

machupicchuense Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 187. 1936. TYPE: Peru, Cuzco, San Miguel, Valle de
Machupicchu. El. 2300 m. 20 July 1928. Herrera 2006 (HT: F
580236; IT: ILL).

maestranum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
6, 10. 1926. TYPE: Cuba, Oriente, Sierra Maestra. 17 July
1922. Ekman 14347 (HT: ILL; PT at ILL: Ekman 1449, 3050,
7245, 14891).

Maguirei Yuncker in Maguire & Collaborators, Bull. Torrey Bot.
Club 75: 289. 1948. TYPE: Suriname, Tafelberg. 19 Sep.
1944, Maguire 24811 (HT: NY; IT: ILL).-- [This species
recognized by Trel. & Yuncker, 1950 and Yuncker, 1957]

malifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):

188. 1936. TYPE: Peru, Huanuco, Pozuzo. El. 600 m.
20-22 June 1923. Macbride 4620 (HT: F 535697; IT: ILL).

malifolium var. mercedanum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 188. 1936. TYPE: Peru, Junfn, La Merced.
El. 600 m. 10-24 Aug. 1923. Macbride 5288 (HT: F 536313;
$F: TL);

manausense Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 107, fig. 93.
1966. TYPE: Brazil, Amazonas, Manaus. El. 25 m. 17 Oct.
1929. Killip & Smith 30136 (HT: US 1463310; IT: bu S oh Me
[This species upheld by Yuncker, 1973]

mandinganum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 153,
fig. 131. 1950. TYPE: Colombia, Choc6, S of Rfo Condoto
between Quebrada Guarapo and Mandinga. El. 120-180 m.

22, 28 April 1939. Killip 35160 (HT: US 1771915; IT: ILL).

marcescentispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 189. 1936. TYPE: Peru, Junfn, Pichis Trail, Santa
Rosa. El. 625-900 m. 6, 7 July 1929. Killip & Smith 26182
(ars USs-21: ILL).

1985 Jones, Annotated catalogue of type specimens 55

margaretae Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
293. 1940. TYPE: Panama, Chiriquf{, island in upper Rfo
Chiriqui Viejo. 13 July 1938. White 158 (HT: ILL).-- {= P.
hispidum (fide Yuncker, 1950 and 1957)] i

margaritatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
189. 1936. TYPE: Peru, San Martin, San Roque. El. 1350-
1500 m. Jan.-Feb. 1930. Williams 7742 (HT: F 623556; IT:
ELL).

marginatum Jacq. var. clausum Yuncker, Ann. Mo. Bot. Gard. Sis
17. 1950. TYPE: Panama, Canal Zone, Empire. 5 March 1937.
Miller 1861 (HT: US; IT: ILL; PT at ILL: Woodson, Allen &
Seibert , 1608). -- [This variety upheld by Yuncker, 1953 and
1957]

marginatum var. domatiiferum Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 23: 306. 1927. TYPE: Haiti, Nord, Mt. Platanna,
St. Michel de l'Atalaye. 7 Dec. 1925. Leonard 7798 (HT:
ILL; PT at ILL: Leonard 7810).

marmoreum Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 24:
186. 1937. TYPE: Panama, Prov. Panama, vicinity of Arenoso,
lower Rfo Trinidad. El. 26-50 m. Aug. 1935. Seibert 622
(HT: ILL).-- [= P. grande Vahl (fide Yuncker, 1950)

marsupiiferum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 189. 1936. TYPE: Peru, Loreto, Soledad, Rfo Itaya.
El. 110 m. 20-22 Sep. 1929. Killip & Smith 29606 (HT: US;
IT?" ILL);

martensianum C.DC. in DC. var. interior Trel., J. Wash. Acad.
Sci. 16: 205. 1926. TYPE: Mexico, Chiapas, Monserrate.
1925. Purpus 172 [HT: US 1209374; IT: ILL (actually a
fragment of the type); PT at ILL: Purpus 143].

Martinense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
190. 1936. TYPE: Peru, San Martf{n, Lamas, near Tarapoto.
El. 840 m. Dec. 1929. Williams 6407 (HT: F 623618; IT:
TE)

masiseanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
190. 1936. TYPE: Peru, Loreto, Masisea. El. 275 m. 25 July
1929. Killip & Smith 26846 (HT: US; IT: ILL).

maternale Trel. in Standley, Field Mus. Publ. Bot. 18: 349.
1937. TYPE: Panama, Bocas del Toro, Flat Rock, Almirante.
1928. Cooper 210 (HT: F; IT: ILL).-- [= P. subcaudatum Trel.
var. maternale (Trel.) Yuncker, 1950; = P. littorale C.DC.
(fide Burger, 1971) ]

56

*

PHYTOLOGIA Vol. 58, No. 1

medium Jacq. var. leonardi Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 23: 305. 1927. TYPE: Haiti, Nord, Marmelade.
20 Dec. 1925. Leonard 8321 (HT: ILL).-- [= P. amalago (cf.
Yuncker, 1962) ]

melanopremnum Trel. & Ekman in Feddes Repert. Spec. Nov. Regni
Veg. 26: 342. 1929. TYPE: Haiti, La Hotte, Quillaud, Les
Roseaux. El. 1500 m. 27 June 1928. Ekman 10139 (HT: ILL;
PT at ILL: Ekman 10298, 10421).

melendezense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
191. 1936. TYPE: Peru, Loreto, Puerto Meléndez, below Pongo
de Manseriche. El. 200 m. 5 Jan. 1932. Mexia 6382 (HT: UC;
aT 2 ELL}.

mellibracteum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 191. 1936. TYPE: Peru, Huanuco, Rfo Huallaga Canyon,
below Rfo Santo Domingo. El. 1200 m. 3 June 1923. Macbride
4234 (HT: F 535313; IT: ILL).-- [= P. elongatum (fide
annotation in Trelease's handwriting) ]

melula Trel., J. Arnold Arb. 9: 148. 1928. TYPE: New Guinea,

Papua, Vaitata River, Hewa. 13 March 1926. Brass 1130
(LECTOTYPE designated herein: A; ILT: ILL).

meraense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 242, fig.
210. 1950. TYPE: Ecuador, Napo-Pastaza, Cant6én Mera.
El. 900 m. 4 May 1935. Mexia 7019 (HT: US 1692972; IT: ILL;
PT at ILL: Ecuador, Mexia 6848; Colombia, Killip & Garcia
33490).

mercedense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
191, 1936. TYPE: Peru, Junfn, La Merced. El. 700 m.
May-June 1929. Killip & Smith 23731 (HT: US; IT: ILL}.

mercens Yuncker, Lilloa 27: 135, pl. 19(1). [1953] 1955.
TYPE: Bolivia, La Paz, S. Yungas, Colaya. El. 1850 m.
13 Dec. 1935. Mexia 7814 (HT: GH; IT: ILL).

meritum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 279. 1940.
TYPE: Honduras, Atlantida, La Ceiba. 12 July 1938. Yuncker,

Koepper & Wagner 8363 (HT: ILL).

metanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 342, fig.
314. 1950. TYPE: Colombia, El Meta, ca 20 km SE of
Villavicencio. El. ca 500 m. 17 March 1939. Killip 34246
(HT: US 1771064; IT: ILL; PT at ILL: Haught 2462, 2769).

1985 Jones, Annotated catalogue of type specimens 57

mikaniaefolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 192. 1936. TYPE: Peru, Loreto, Yurimaguas to
Balsapuerto. El. 135-150 m. 26-31 Aug. 1929. Killip &
Smith 30691 (HT: US; IT: ILL).

milciadesii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 314,
fig. 280. 1950. TYPE: Colombia, El Valle, R{fo Digua Valley
between La Elsa and Rfo Blanco. El. 825 m. April 1939.
Killip & Milciades Dussan 34732 (HT: US 1771530; IT: ILL;
PT at ILL: Haught 4630).

millepunctulatum Trel. & Yuncker, Piperaceae no. S. Amer. 1:
332, fig. 300. 1950. TYPE: Colombia, Chocé, Andagoya.
' El. 70-100 m. April 1939. Killip 35062 (HT: US 1771816; IT:
ILL).

minute-scabiosum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
27: 294. 1940. TYPE: Panama, Coclé, north rim of El Valle de
Ant6n. 12 Feb. 1939. Allen 1639 (HT: ILL).-- [This species
recognized by Yuncker, 1950]

minute-scabiosum var. arborescens Trel. in Woodson & Schery,
Ann. Mo. Bot. Gard. 27: 294. 1940. TYPE: Panama, Coclé, El
Valle. 8 Dec. 1938. Allen 1185 (HT: ILL).-- [= P.
minute-scabiosum (fide Yuncker, 1950) ]

mirabile Trel., Contr. U.S. Nat. Herb. 26(4): 154. 1929. TYPE:
Costa Rica, Santa Clara de Cartago. El. 1950 m 20 July
1923. Maxon & Harvey 8246 (HT: US 1181894; IT: ILL).--
[= P. euryphyllum C.DC. (fide Burger, 1971) ]

mishuyacuense Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 192. 1936. TYPE: Peru, Loreto, Mishuyacu near Iquitos.
El. 100 m. 24-28 Sep. 1929. Killip & Smith 29903 (HT: US;
i: LEY:

mitifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
192. 1936. TYPE: Peru, Junfn, La Merced. El. 700 m.
10-24 Aug. 1923. Macbride 5325 (HT: F 536350. «iT s..ILL; PT at
ILL: Stevens 204).

* molanoi Trel. & Yuncker, Piperaceae no. S. Amer. 1: 127, fig.
102. 1950. TYPE: Colombia, Narino, Gorgonilla Island.
El. 130-200 m. 8 Feb. 1939. Killip, Garcfa & Molano 33119
(LECTOTYPE designated herein: US 1770110 (fig. 102); ILT:
ILL).

58

Pi YT TOR GP A Vol. 58, No. 1

monteluctans Trel. in Yuncker, Field Mus. Publ. Bot. 17: 347.
1938. TYPE: Honduras, Comayagua, creeping over logs in
forest, cloud zone on range above El Achote, near
Siguatepeque. El. 1800 m. Yuncker, Dawson & Youse 6176 (HT:
ILL).

monurum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 336, fig.

- 306. 1950. TYPE: Ecuador, Esmeraldas, Parroquia Concepcion,
Playa Rica. El. 105 m. 21 Dec. 1936. Mexia 8477-a (HT: GH;
IT: SLL).

morator Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 194.

1936. TYPE: Peru, Jun{n, San Ram6n. El. 900-1300 m.
9, 12 June 1929. — Killip & Smith 24743 (HT: US; IT: ILL).

morianum Trel., J. Arnold Arb. 9: 148. 1928. TYPE: New Guinea,

Papua, Lower Mori River. 28 May 1926. Brass 1562
(LECTOTYPE designated herein: A; ILT: ILL).-- [= P.
macropiper (fide Chew, 1972) ]

multinervium M. Martens & Galeotti var. amplum Trel. in Yuncker,

Field Mus. Publ. Bot. 17: 347. 1938. TYPE: Guatemala, Petén,
La Libertad. 7 April 1933. Lundell 2556 (HT: MICH; IT:
ILL).-- [= P. aduncum (fide Standley & Steyerm., 1952) ]

multinervium var, cayoense Trel. in Yuncker, Field Mus. Publ.

Bot. 17: 347. 1938. TYPE: British Honduras (Belize), El
Cayo. 19 Feb. 1931. Bartlett 11570 [HT: MICH; IT (2
sheets): ILL].

multinervium var. flavicans Trel. in Yuncker, Field Mus. Publ.

Bot. 17: 348. 1938. TYPE: Honduras, Comayagua, Siguatepeque.
El. 1200 m. 14 July 1936. Yuncker, Dawson & Youse 5859 (HT:
1).

multinervium var. hirsuticaule Trel. in Yuncker, Field Mus.

Publ. Bot. 17: 348. 1938. TYPE: Honduras, Comayagua,
Siguatepeque. El. 1050 m. 8 July 1936. Yuncker, Dawson &
Youse 5724 (HT: ILL; PT at ILL: Yuncker, Dawson & Youse
5699).

multinervium var. kantelolense Trel. in Yuncker, Field Mus.

Publ. Bot. 17: 348. 1938. TYPE: Guatemala, Petén, La
Libertad. 25 April 1933. Lundell 3008 (HT: MICH; IT: ILL;
PT at ILL: Lundell 3168).-- [= P. aduncum (fide Standley &
Steyerm., 1952) ]

1985 Jones, Annotated catalogue of type specimens 59

* multinervium var. peracutum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 349. 1938. TYPE: Honduras, Atlantida, Lancetilla
Valley near Tela. El. 20-600 m. 1927-1928. Standley 54921
(HT, y LECTOTYPE designated herein: ILL; PT at ILL: Standley
53083 ).

multinervium var. productipes Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 349. 1938. TYPE: Honduras, Comayagua, near El
Achote above Siguatepeque. El. 1350 m. 13 July 1936.
Yuneker, Dawson & Youse 5819 (HT: ILL).

multinervium var. protractifolium Trel. in Yuncker, Field Mus.
Publ. Bot. 17: 349. 1938. TYPE: Honduras, Comayagua, above
Siguatepeque. El. 1050 m. 5 July 1936. Yuncker, Dawson &
Youse 5668 (HT: ILL).

multinervium var. puberulipedunculum Trel. in Yuncker, Field
Mus. Publ. Bot. 17: 349. 1938. TYPE: Honduras, Comayagua, El
Achote, above Siguatepeque. El. 1350 m. 15 July 1936.
Yuncker, Dawson & Youse 5876 (HT: ILL).

multinervium var. pubescenticaule Trel. in Yuncker, Field Mus.
Publ. Bot. 17: 350. 1938. TYPE: Honduras, Comayagua,
Siguatepeque. El. 1050 m. 8 July 1936. Yuncker, Dawson &
Youse 5731 (HT: ILL).

multinervium var. skutchii Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 350. 1938. TYPE: Guatemala, Quezaltenango,
Colombia. El. 900 m. 24 Sep. 1934. Skutch 1299 (HT:
ILL).-- [= P. aduncum (fide Standley & Steyerm., 1952)]

multinervium var. tamashense Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 350. 1938. TYPE: British Honduras (Belize), Tamash
River. El. 30 m. 10 March 1935. Schipp 1382 (HT: ILL).

Mmultinervium var. telanum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 350. 1938. TYPE: Honduras, Atlantida, Tela.
El. 20-600 m. 1926, 1927. Standley 53264 (HT: F; IT: ILL;
PT at ILL: Yuneker 4498, 4512).

multitudinis Trel., Ciencia (Mexico) 2: 207. 1941. TYPE: Peru,
Chanchamayo. 23 Oct. 1863. Isern 2181-c (HT: MA; probable
T= ILE).

multitudinis [var.] latum Trel., Ciencia (Mexico) 2: 207. 1941.
TYPE: Peru, Chanchamayo. 23 Oct. 1863. Isern 2181-b (HT:
MA; IT: ILL).

60

*

*

Ball Yi HsOrk OvGs IsA Vol. 58, No. 1

munchanum C.DC. in DC. var. latum Trel. in Macbr., Field Mus.
Publ. Bot. 13(No. 357): 195. 1936. TYPE: Peru, San Mart{n,
San Roque. El. 1350-1500 m. Jan.-Feb. 1930. Williams 6954
(HT: F 623002; IT: ILL).-- [This species recognized by Trel.
& Yuncker, 1950)

munyanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. '357):
195. 1936. TYPE: Peru, Huanuco, Muna. El. 2400 m. 5-7 June
1923. Macbride 4272 (HT: F 535354; IT: ILL).

musteum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 196.
1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m. 1-5 Sep.
1929. Killip & Smith 28982 (HT: US; IT: ILL).

mutisii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 414, fig.
377. 1950. TYPE: Colombia. Mutis 3463 (HT: MA; IT: ILL).--
[This species recognized by Burger, 1971]

mystaciferum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 350.
1938. TYPE: Honduras, Comayagua, ravine at El Rincon, 10
miles W of Siguatepeque. El. 1300-1400 m. 24 July 1936.
Yuncker, Dawson & Youse 6054 (HT: ILL).

nanayanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
196. 1936. TYPE: Peru, Loreto, S bank of Rfo Amazonas,
Nanay. May-June 1929. Williams 516 (HT: F 603936; IT: ILL).

nanegalense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 25, ore
15. 1950. TYPE: Ecuador, Pichincha, Nanegal. El. 1600 om.
1908. Mille s.n. [LECTOTYPE designated herein (fig. 15):
BLL ALT Sela

negritosense Trel. in Cuf., Archivio Bot. Sist. Fitogeog. &
Genet. 10: 25. 1934. TYPE: Costa Rica, "La Castilla-Los
Negritos". El. 20 m. 8 May 1930. Cufodontis 528 (HT, or
LECTOTYPE designated herein: ILL).-- [= P. dryadumC.DC.
(fide Burger, 1971)]

niceforoi Trel. & Yuncker, Piperaceae no. S. Amer. 1: 79, fig.
66. 1950. TYPE: Colombia, Norte de Santander, Cicuta.
Feb. 1940. Nicéforo 49 (HT: ILL; PT at ILL: Killip, Dugand &
Jaramillo 38293).

nicoyanum C.DC., Bot. Gaz. 70: 174. 1920. TYPE: Costa Rica,
Bords des chemins & Nicoya. Pittier 13689, 13696 [Syntypes,
perhaps to be lectotypified; fragment of Tonduz (= Pittier?)
13689 (from LE) at ILL].-- [= P. amalago (fide Burger, 1971) ]

1985 Jones, Annotated catalogue of type specimens 61

nigro-granulatum Trel. in Rusby, Mem. N.Y. Bot. Gard. 7: 224.
1927. TYPE: Bolivia, Bopi River Valley. El. 3000 ft.
Sep. 1921. Rusby 573 (HT: NY; IT: ILL).-- [= P. lanceolatum
Ruiz & Pavon (fide Yuncker, 1955)]

nitidulifolium Trel. ex Standley in Standley & Record, Field
Mus. Publ. Bot. 12: 408. 1936. TYPE: British Honduras
(Belize), Tower Hill. 1928. Karling 26 (HT: F; IT: ILL).--
[= P. aeruginosibaccum (fide Standley & Steyerm., 1952);
= P. jacquemontianum (cf. Burger, 1971) ]

nobile C.DC. in DC. var. glabratum Yuncker in Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 150. 1950. TYPE: Colombia,
Antioquia, Rfo Saman&, 2 km above Argelia. El. ca 2560 m.
May-June 1944. Ewan 15771 (HT: ILL).

nobile var. parvulum Trel. & Yuncker, Piperaceae no. S. Amer. 1:
151. 1950. TYPE: Venezuela, Distr. Fed., El Junquito.
El. ca 1925 m. 26 March 1943. Killip & Rohl 37177 (HT: US
1855347; IT: ILL).

nonconformans Trel., J. Wash. Acad. Sci. 19: 334. 1929. TYPE:
Honduras, Comayagua, near Siguatepeque. El. 1100 m.
Feb. 1928. Standley 55906 (HT: F; IT: ILL).

nottirbanum Trel. ex Stehlé, Bull. Soc. Bot. France 85: 576.
1939 (nomen); Fl. Deser. Antill. Fr. 2(1): 103. 1940; Bull.
Agric. Martinique 9(3): 180. 1940. TYPE: Guadeloupe,
Malanga. Stehlé & Stenlé 1748 [HT: ILL (annotated as such by
Howard, but this is only a sterile stem with a single
leaf)].-- [= P. glabrescens (fide Howard, 1973) ]

novae-helvetiae Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
27: 294. 1940. TYPE: Panama, Chiriquf, vicinity of "New
Switzerland", Rfo Chiriquf Viejo Valley. El. 1800-2000 m.
6 Jan. 1939. Allen 1359 (HT: ILL; PT at ILL: Allen 1416).--
(= P. crassinervium Kunth (fide Yuncker, 1950) ]

nubigaudens Trel. in Yuncker, Field Mus. Publ. Bote 17> 35.
1938. TYPE: Honduras, Comayagua, El Achote, near
Siguatepeque. El. 1800 m. 21 July 1936. Yuncker, Dawson &
Youse 5991 (HT: ILL).

nubigenum Kunth var. brevifolium Trel. & Yuncker, Piperaceae no.
S. Amer. 1: 354. 1950. TYPE: Colombia, Boyacad, Puente de
Boyacad. El. 2800 m. 24 Feb. 1940. Arbeldez & Cuatrecasas
8075 (HT: US 1774594; PT at ILL: Ecuador, Isern 1004).

62 Phe FTO) be OG4) A Vol. 58, No. 1

nudibracteatum C.DC. var. micropuberulum Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 152. 1950. TYPE: Colombia, El
Valle, La Cumbre. El. 1800-2100 m. May 1922. Pennell &
Killip 5868 [HT: US 1141960; IT: ILL (2 sheets)].

nudibracteatum var. pubescens Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 152. 1950. TYPE: Colombia, Cauca, Cauca Valley.
E1.1500-1600 m. 27 June 1922. Pennell & Killip 7226 [HT: US
sete PT at ILL: Figueroa 915; Killip, Garcla & Lora
33708).

nuncupatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
197. 1936. TYPE: Peru, Junin, Pichis Trail, Yapas.
El. 1350-1600 m. 28, 29 June 1929. Killip & Smith 25577
CHT 2clS: ETE Eb?.

oblanceolatum Trel., Contr. U.S. Nat. Herb. 26(4): 175. 1929.
TYPE: Costa Rica, Guanacaste, Los Ayotes, Tilaran.
El. 650 m. 21 Jan. 1926. Standley & Valerio 45558 (HT: US
1307380; IT: ILL).-- [This species recognized by Burger,
1971]

oblongatifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 198. 1936. TYPE: Peru, Loreto, valley of Rio Maranén.
El. 150 m. Aug.-Sep. 1929. Leg. Dennis; Killip & Smith
29153(HTs. US; IT ILL).

oblongatilimbum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 198. 1936. TYPE: Peru, Loreto, Yurimaguas to
Balsapuerto. El. 135-150 m. 26-31 Aug. 1929. Killip &
Smith -26237> (HT: USe01T:> ILL).

obovatifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 199. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m.
1-5 Sep: - 1929. Killip & Smith 28980 (HT: US; IT: ILL; PT at
ILL: Killip & Smith 28096).

obrutum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 324, fig.
289. 1950. TYPE: Colombia, Santander, northern slopes of
Mesa de los Santos. El. 1000-1500 m. Dec. 1926. Killip &
Smith 15379 (HT: GH; IT: ILL).

obsessum Trel., J. Wash. Acad. Sci. 19: 334. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
22 Jan. 1928. Standley 54925 (HT: F 584233; IT: ILL).--
[= P. fraguanum (fide Standley & Steyerm., 1952);
= P. sancti-felicis (fide Burger, 1971) ]

1985 Jones, Annotated catalogue of type specimens 63

occultum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
199. 1936 (erroneously printed as Peperomia). TYPE: Peru,
Loreto, near Iquitos. El. 120 m. July 1929. Williams 1406
CHT: £ 617357; dt> ILL),

ocoanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 29: 29.
1931. TYPE: Santo Domingo, Bejucal, S. José de Ocoa, Azua.
El. 1300 m. 25 March 1929. Ekman 12046 (HT: ILL).

oculatispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 200. 1936. TYPE: Peru, Loreto, Iquitos. El. 120 m.
March-April 1930. Williams 7954 (HT: F 629718; IT: ILL).

offensum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
200. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 150-350 m.
28-30 Aug. 1929. Killip & Smith 28426. (HT 2 US, IT esEbL).

omega Trel., Contr. U.S. Nat. Herb. 26(4): 146. 1929. TYPE:
Costa Rica, San José, La Hondura. March 1924. Standley
37800 (HT: US 1229449; PT at ILL: Standley & Valerio
50940). -- [= P. carrilloanum (fide Burger, 1971) ]

onerosum Trel., J. Wash. Acad. Sci. 19: 335. 1929. TYPE:
Honduras, Atlantida, near Tela, at sea level. 27 Dec. 1927.
Standley 53696 (HT: F 582949; IT: ILL; PT at ILL: Standley
52682, 55022, 022, 56621).-- [= P. aeruginosibaccum (fide Standley
& & Steyerm. » 1952); = P. jacquemontianum (ef. Burger, 1971)]

opacibracteum Trel., Contr. U.S. Nat. Herb. 26(4): 148. 1929.
TYPE: Costa Rica, Guanacaste, La Tejona, N of Tilaran.
El. 600-700 m. 25 Jan. 1926. Standley & Valerio 45899 (HT:
US 1307414; IT: ILL).-- [= P. xanthostachyum C.DC. (fide
Burger, 1971)]

ornatispicum Trel. in Yuncker, Field Mus: -Publia Bote <b. 5as
1938. TYPE: Honduras, Comayagua, thicket on river bank near
Siguatepeque. El. 1050 m. 3 July 1936. Yuncker, Dawson &
Youse 5618 (HT: ILL).

ornithorhynchum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 201. 1936. TYPE: Peru, Ayacucho, Carrapa. El. 1500 m.
May 1929. Killip & Smith 22334 (HT: US: ITs IbbguP% iat. TLL:
Killip & Smith 23221).

ostii Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 201.
1936. TYPE: Peru, Loreto, mouth of Rio Santiago.
El. 200 m. 24 Nov. 1931. Mexia 6152 (HT: UC; IT: TLL)

64 Pea YS) 'b. OG Vol. 58, No. 1

ottonoides Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 109, fig. 95.
1966. TYPE: Brazil, Parad, Remansdo do Centro, E Ferro
Tocantins. 1 Oct. 1948, Frées 23592 (HT: IAN; PT at ILL:
Minas Gerais, Mexia 5332).-- [This species upheld by Yuncker,
1973]

pachyphloium C.DC. in Kuntze var. macrophyllum Yuncker, Lilloa
27: 107, pl. 4(1). [1953] 1955. TYPE: Bolivia, Cochabamba,
Tungas of Tana Mayu. El. 1800 m. 24 June 1947. Cardenas
3970 (HT: US 1903317; PT at ILL: Cardenas 2161).

paganicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
202. 1936. TYPE: Peru, Loreto, Iquitos. El. 100 m.
3-11 Aug. 1929. Killip & Smith 21432 (CHT: US;. IT: ILE).

pallididorsum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 202. 1936. TYPE: Peru, Junin, La Merced, Schunke
Hacienda. El. 1300 m. Aug.-Sep. 1923. Macbride 5801 (HT: F
536830; IT: ILL).

paludicola Trel. in Yuncker, Field Mus. Piel .. Bot. TT. sons
1938. TYPE: Honduras, Atlantida, in marsh near Tela. 9 Aug.
1934. Yuncker 4977 (HT: ILL).

pamploanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 1G, Tig.
90. 1950. TYPE: Colombia, Norte de Santander, western slope
of Paramo del Hatico en route from Toledo to Pamplona.
El. 2800-2900. 13 March 1927. Killip & Smith 20720 (HT: US
13557163’ IT] ILL).

papillipetiolatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 203. 1936. TYPE: Peru, Loreto, Iquitos. El. 100 m.
3-11 Aug. 1929. Killip & Smith 27407. CHT «US; ITs Thki)}e

papulaecaule Trel. in Standley, Field Mus. Publ. Bot. 18 352%
1937. TYPE: Costa Rica, Guanacaste, Hacienda Santa Maria.
El. 720-920 m. 23 Jan. 1930. Dodge & Thomas 6286 (HT: GH;
IT: ILL).-- [= P. chrysostachyum (fide Burger, 1971) ]

paranapuranum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 204. 1936. TYPE: Peru, Loreto, Rio Parana-pura near
Yurimaguas. El. 180 m. 24 Oct. 1931. Mexia 6075 (HT: UC;
BY isc ds) 4

para-peltobryon Trel. in Gleason, Bull. Torrey Bot. Club 58:
356. 1931. TYPE: Venezuela, Amazonas, summit of Mt. Duida.
El. 1800 m. Tate 711 (HT: NY; IT: ILL).-- [This species
recognized by Trel. & Yuncker, 1950]

1985 Jones, Annotated catalogue of type specimens 65

parcepilosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
204. 1936. TYPE: Peru, Junfn, Carpapata. El. 2700-3200 m.
7 June 1929. Killip & Smith 24385 (HT: US; IT: ILL).

parcum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 311, fig.
277. 1950. TYPE: Ecuador, Los Rios, near Quevedo, Canton
Vinces. El. 60 m. 2 Nov. 1924. Mexia 6604 (HT: US
1619067; IT: ILL).

partiticuspe Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
294. 1940. TYPE: Panama, Coclé, El Valle de Antén. 8 Dec.
1938. Allen 1195 (HT: ILL).-- [This species recognized by
Yuneker, 1950]

paso-anchoense Trel. in Woodson & Schery, Ann. Mo. Bot. Gard.
27: 295. 1940. TYPE: Panama, Chiriquf, trail from Paso Ancho
to Monte Lirio, upper valley of Rio Chiriqu{ Viejo. 16 Jan.
1939. Allen 1579 (HT: ILL).-- [= P. chiriguinum C.DC. (fide
Yuneker, 1950); = P. aequale (fide Burger, 1971) ]

patentifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 205. 1936. TYPE: Peru, Junin, Hacienda Calhuapuquio.
8 Dec. 1924. Stevens 136 (HT: ILL; PT at ILL: Stevens 148).

* patulum Bertol. var. cordifolium Trel. in Standley, J. Wash.
Acad. Sci. 13: 366. 1923. TYPE: El Salvador, Sonsonate,
vicinity of Nahulingo. 21 March 1922. Standley 22046 (HT,
or LECTOTYPE designated herein: ILL).-- [The species
recognized by Standl. & Steyerm. (1952); = P. marginatum
(fide Burger, 1971)]

paucartamboanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 205. 1936. TYPE: Peru, Junfn, near Perené Bridge, R{io
Paucartambo Valley. El. 700 m. 19 June 1929. Killip &
Smith:25388 (HT :°US; ITs! ILL)

pauciramosum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 60, fig.
50. 1966. TYPE: Brazil, Minas Gerais, District Ilhéus, Faz.
da Tabunha. El. 300 m. 13 Aug. 1930. Mexia 4973 [HT: NY;
IT: ILL; PT at ILL: Hoehne 19524 (2 sheets)].-- [This species
upheld by Yuncker, 1972]

* pavimentifolium Trel., J. Arnold Arb. 9: 148. 1928. TYPE: New
Guinea, Papua, Murua River. El. 100 m. 29 March 1926.
Brass 1338 (LECTOTYPE designated herein: A; ILT: ILL).

pebasense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
206. 1936. TYPE: Peru, Loreto, Pebas on the Amazon River.
July 1929. Williams 1746 (HT: F 608248; IT: ILL).

66 PT Ob OG a A Vol. 58, No. 1

pelliticaule Trel., Contr. U.S. Nat. Herb. 26(4): 157. 1929.
TYPE: Costa Rica, Alajuela, vicinity of Fraijanes.
El. 1500-1700 m. Feb. 1926. Standley & Torres 47539 (HT: US
1307499; IT: ILL).-- [= P. bredemeyeri (fide Burger, 1971) ]

peltilimbum Yuncker, Lilloa 27: 155, pl. 37. [1953] 1955. TYPE:
Bolivia, La Paz, S. Yungas, Colaya. El. 1780 m. 14 Dec.
1935. Mexia 7823 (HT: US 1778383; IT: ILL).

pendentispicum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 157,
fig. 136. 1950. TYPE: Colombia, Norte de Santander, Culaga
Valley above Tapata. El. ca 2300 m. 5, 6 March 1927.
Killip & Smith 20338 (HT: US 1355402; IT: ILL).

pendentispicum var. tejamesmeanum Trel. & Yuncker, Piperaceae
no. S. Amer. 1: 158. 1950. TYPE: Colombia, Norte de

Santander, road from Pamplona to Toledo, etc. El. 2800-
3000 m. Feb. 1927. Killip & Smith 20006 (HT: US 1355155;
Ts) 21154

pennellii Trel. & Yuncker, Piperaceae no. S. Amer. 1: 121, fig.
95. 1950. TYPE: Colombia, Caldas, Salento. El. 1400-1700 m.
31 July 1922. Pennell 9056 (HT: US 1142766; IT: ILL).

penyasense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 207.
1950. TYPE: Colombia, Antioquia, Peras Blancas. 26 April
1926. Woronow & Juzepcezuk 4570 (HT: LE; fragment of type at
ILL).

perbrevispicum Yuncker in Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 226, fig. 190. 1950. TYPE: Colombia, Antioquia,
Turbo. El. 300 m. 23 April 1946. Haught 4809 (HT: ILL).

perciliatum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 326,
fig. 293. 1950. TYPE: Venezuela, Amazonas, San Carlos de Rio
Negro. El. 100 m. 7 March 1942. Williams 14683 (HT: US
1E7ONT TS: TPs TL

percostatum Yuncker, Lilloa 27: 157, pl. 39. [1953] 1955. TYPE:
Bolivia, La Paz, Mapiri. El. 1500 m. May 1886. Rusby 2174
(HT: US 79650; PT at ILL: Buchtien 574 (2 sheets), 581].

perenense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
207. 1936. TYPE: Peru, Junin, Colonia Perené. El. 600 m.
16-18 June 1929. Killip & Smith 25108 (BT: USs “ITs ELb; PT
at ILL: Killip & Smith 26442).

perenense var. cupulifolium Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 207. 1936. TYPE: Peru, Junin, La Merced.
El. 700 m. May-June 1929. Killip & Smith 23698 (HT: US; IT:
ILL).

1985 Jones, Annotated catalogue of type specimens 67

perfugii Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 295.
1940. TYPE: Panama, Chiriqu{, vicinity of Casita Alta,
Volean de Chiriquf. El. 1500-2000 m. June 1938, Woodson,
Allen & Seibert 928 (HT: ILL).-- [= P. hirtellipetiolum C. C.0Cc.
var. var. harveyanum (Trel.) Yuncker, 1950; = P. tenuimucronatum
C.DC. (fide Burger, 1971)] e

pergamentifolium Trel. & Standley in Standley & Steyerm.,
Fieldiana Bot. 24 (3): 314. 1952. TYPE: Guatemala,
Quezaltenango, Volean de Zunil. 31 July 1934. Skutch 876
(HT: F; IT: ILL). ie

perinaequilongum Trel. in Standley, Field Mus. Publ. Bot. 10:
160. 1931. Type locality: Honduras, Lancetilla Valley. No
type cited; probably lectotypification needed (authentic
specimens at ILL: near Tela, Standley 54419, 54897, 56893).

perlasense Yuncker in Woodson & Schery, Ann. Mo. Bot. Gard. 37:
67. 1950. TYPE: Panama, Perlas Archipelago, San José Island.
22 Jan. 1946. Johnston 1215 [HT: ILL; PT at ILL: Johnston
134, 262, 717, 1327; Stevens 7 (2 sheets)].

permucronatum Yuncker, Bol. Inst. Bot. (Sado -Paal6)<32 (Tl >ofie.
9. 1966. TYPE: Brazil, Rio de Janeiro, Tijuca, Bom Retiro.
7 Nov. 1946. Rosa 79 (HT: R; PT at ILL: Lutz 27; Sampaio,
Freire, Peckolt & Costa 490).-- [This species upheld by
Yuncker, 1972]

pernigricans Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
207. 1936. TYPE: Peru, Loreto, Mishuyacu. El. 100 m. 24-28
Sep. 1929. Killip & Smith 29976 (HT: US: IT:.JILb).

pernodosum Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.
1: 206, fig. 168. 1950. TYPE: Colombia, Alban. 28 Jan.
1947, Silvano Jorge 45 (HT: ILL).

pernodulosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 120,
fig. 94. 1950. TYPE: Colombia, Cundinamarca, Aguadita.
28 Nov. 1939. Laureano Javier 34 (HT: ILL).

perobumbratum Trel. in Standley, Field Mus. Publ. Bot. 22(1):
11. 1940. TYPE: Brazil, Rio de Janeiro. Pohl 4772 [HT: W;
IT: ILL (2 sheets) ].-- [=P P. arboreum Aublet (fide Yuncker,
oe

perpallidum Ekman, Urban & Trel. in Feddes Repert. Spec. Nov.
Regni Veg. 23: 305, 309. 1927. TYPE: Haiti, Sud, Riviere
Bras Gauche, Trouin. 11 May 1924. Ekman 2365 (HT: ILL; PT
at ILL: Ekman 5093).

68 Pst Ti Oitougnik Vol. 58, No. 1

perpunctulatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 207. 1936. TYPE: Peru, Loreto, Yurimaguas to
Balsapuerto. El. 135-150 m. 26-31 Aug. 1929. Killip &
Smith 28090 (HT: US; IT: ILL).-- [The specimen at ILL is
marked in Trelease's handwriting "= conditum", and this
number is also cited as a PT of P. conditum]

perpurgatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
208. 1936. TYPE: Peru, Loreto, San Antonio, on Rfo Itaya.
El. 110 m. 18 Sep. 1929. Killip & Smith 29455 (HT: US; IT:
ILL; PT at ILL: Killip & Smith 29461, 29707).

perspicuibracteum Trel., J. Wash. Acad. Sci. 19: 335. 1929.
TYPE: Honduras, Atlantida, Lancetilla Valley near Tela.
El, 100 m. 21 Jan. 1928. Standley 54872 (HT: F 584074; IT:
ILL)2

pertractatum Trel., Contr. U.S. Nat. Herb. 26(4): 130. 1929.
TYPE: Costa Rica, Alajuela, Nuestro Amo. Feb. 1919. Jiminez
s.n. (HT: US 1080638; PT at ILL: Standley & Valerio 45694,

45710).-- [= P. pseudo-lindenii C.DC. (fide Burger, 1971) ]

peruligerum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
295. 1940. TYPE: Panama, Bocas del Toro, Isla de Colon.
El. 25-75 m. Aug. 1938. Woodson, Allen & Seibert 1939 (HT:
MO; fragments of type at ILL).-- [This species recognized by
Yuneker, 1950]

perverrucosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 282,
fig. 247. 1950. TYPE: Colombia, Norte de Santander, Caluga
Valley near Tapata (N of Toledo). El. 1500-2100 m.

3-8 March 1927. Killip & Smith 20350 (HT: US 1355408; IT:
ILL; PT at ILL: Killip & Smith 19827).

pervicax Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 296.
1940. TYPE: Panama, Chiriquf, vicinity of Casita Alta,
Volean de Chiriquf{. El. 1500-2000 m. Woodson, Allen &
Seibert 843 (HT: ILL).-- [= P. hispidum (fide Yuncker, 1950
and 1957) ]

pervillosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
208. 1936. TYPE: Peru, San Martin, San Roque. El. 1350-
1500 m. Jan.-Feb. 1930. Williams 6963 (HT: F 622979; IT:
TEL).

pervulgatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
209. 1936. TYPE: Peru, Loreto, Mishuyacu near Iquitos.
El. 100 m. Klug 468 (HT: F 629836; PT at ILL: Killip & Smith
29312; Williams 1889).

1985 Jones, Annotated catalogue of type specimens 69

pervulsum Trel. & Yuncker var. santosense Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 338. 1950. TYPE: Colombia,
Santander, Mesa de los Santos. El. 1500 m. Dec. 1926.
Killip & Smith 15363 (HT: US 1351246; IT: ILL).

petens Trel.' in Yuncker, Field Mus. Publ. Bot. 17: 352..1938:
TYPE: Honduras, Comayagua, El Achote, above Siguatepeque.
El. 1350 m. 26 July 1936. Yuncker, Dawson & Youse 6107 (HT:
fL-E)

phaneropus Trel. in Standley, Field Mus. Publ. Bot. 18: 354.
1937. TYPE: Costa Rica, 5 miles S of Santa Marfa de Dota.
El. 1980 m. 2 May 1928. Stork 1775 (HT: ILL).-- [= P.
epigynium C.DC., a glabrous form (fide Burger, 1971) ]

phegophyllum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
209. 1936. TYPE: Peru, Junin, Puerto BermGdez. El. 375 m.
14-17 July 1929. Killip & Smith 26479 (HT: US; IT: ILL).

picardae C.DC. in Urban var. pilinervium Trel. in Feddes Repert.
Spec. Nov. Regni Veg. 23: 309. 1927. TYPE: Haiti, Motagne
Noire, La Selle. El. 1700 m. 4 Sep» 1924. Ekman 1769 (HT:
ILL).

pichisense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
210. 1936. TYPE: Peru, Junfn, Pichis Trail, San Nicol4s.
El. 1100 m. 4-5 July 1929. Killip & Smith 26012 (HT: US;
DT SeELE).

pictamentum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 352.
1938. TYPE: Honduras, Comayagua, Siguatepeque. El. 1100 m.
17 July 1936. Yuncker, Dawson & Youse 5902 (HT: ILL).--
[= P. achoteanum (fide Standley & Steyerm., 1952) ]

pictamentum var. fluminis Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 352. 1938. TYPE: Honduras, Comayagua, Siguatepeque.
El. 1100 m. 8 July 1936. Yuncker, Dawson & Youse 5718 (HT:
ILL).

piedecuestanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 227,
fig. 191. 1950. TYPE: Colombia, Santander, vicinity of
Piedecuesta. El. 980 m. Dec. 1926. Killip & Smith 15445
(HT: US 1351320; IT: ILL; PT at. ILL: Cuatrecasas & Garcfa

Barriga 9869.)

piluliferifolium Trel. & Yuncker, Piperaceae no. S. Amer. 1:
315, fig. 281. 1950. TYPE: Colombia, Putumayo, Umbria.
El. 325 m. Oct.-Nov. 1930. Klug 1765 (HT: US 1518013; PT
at ILL: Cuatrecasas 11176).

70 Prk ¥yT @ LoOeGriré Vol. 58, No. 1

pinaresanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 202,
fig. 162. 1950. TYPE: Colombia, Caldas, Pinares, near
Salento. El. 2600-2900 m. Aug. 1922. Pennell 9205 [HT: US
1142797; IT: ILL (2 sheets) ).

piritubanum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 61, fig. 51.
1966. TYPE: Brazil, Sao Paulo, Pirituba. 14 Dec. 1913.
Brade 7181 (cited as Brade s.n.) (HT: ILL).-- [This species
upheld by Yuncker, 1972]

piscatorum Trel. ex L. Williams, Explor. Bot. Guayana
Venezolana, 176. 1942 (nomen subnudum); in Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 395, fig. 356. 1950. TYPE:
Venezuela, Bolfvar, La Prisi6én. El. 120 m 3 April 1939.
Williams 11685 (HT: US 1459788; IT: ILL; PT at ILL: Williams
11258).

pitalitoense Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.
1: 309, fig. 275. 1950. TYPE: Colombia, Huila, Pitalito,
Quinche. El. 1300 m. 30 Dec. 1942. Schultes & Villarreal
5101 (HT: GH; IT: ILL).

planipes Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
210. 1936. TYPE: Peru, Junin, Colonia Perené. El. 680 m.
14-22 June 1929. Killip & Smith 24928 (HT: US; IT: ILL).

platannanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
308, 312. 1927. TYPE: Haiti, Mt. Platanna, S. Michel de
l'Atalaye. El. 350 m. 7 Dec. 1925. Leonard 7818 (HT: ILL;
PT at ILL: Leonard 7809).

pleninervosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 211. 1936. TYPE: Peru, Loreto, Puerto Arturo.
El; 135m. 24-25 Aug: 1929. Killip & Smith 27875 (HT: US;
50S IED

plumbeicolor Trel. in Standley, Field Mus. Publ. Bot. 17: 233.
1937. TYPE: Guatemala, Petén, near El Paso. 7 April 1933.
Lundell 2554 (HT: ILL).-- [= P. aeruginosibaccum Trel. (fide
Standley & Steyerm., 1952); = P. jacquemontianum (cf. Burger,
1971) ]

politaereum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
211. 1936. TYPE: Peru, Junin, San Ramon, Schunke Hacienda.
El. 1400-1700 m. 8-12 June 1929. Killip & Smith 24545 (HT:
US; IT: ILL; PT at ILL: Killip & Smith 23905, 24553).

politaereum var. elongatum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 211. 1936. TYPE: Peru, Loreto, Soledad.
El. 110 m. 20-22 Sep. 1929. Killip & Smith 29587 (HT: US;
The SEL).

1985 Jones, Annotated catalogue of type specimens 71

politaereum var. mucronatum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 211. 1936. TYPE: Peru, Loreto, San
Antonio, Rfo Itaya. El. 110 m. 18 Sep. 1929. Killip &
Smith 29460 (HT: US; IT: ILL).

ponganum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
212. 1936. TYPE: Peru, Loreto, Pongo de Manseriche.
El. 200 m. 19 Nov. 1931. Mexia 6123 (HT: UC; IT: ILL).

pontis Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 212.
1936. TYPE: Peru, Junfn, near Perené Bridge, Rfo Paucartambo
Valley. El. 700 m. 19 June 1929. Killip & Smith 25294 (HT:
US: IF 2 ELL).

portalense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
213. 1936. TYPE: Peru, Loreto, Portal, lower R{o Huallaga.
El. 135 m. 5-11 Sep. 1929. Killip & Smith 29258 (HT: US;
ts Fb).

poscitum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 279, fig.
242. 1950. TYPE: Ecuador, Los Rios, near Quevedo, Cantén
Vinces. Oct.-Nov. 1934. Mexia 6594 (HT: US 1619066; IT:
LLL) «

positum Trel. in Macbr., Field Mus. Publ. Bot. 13(¢N0. 5957) 213.
1936. TYPE: Peru, Loreto, Portal, Rfo Huallaga. El. 135 m.
5=71..cep.. 929. Killip & Smith 29256 (HT: US; IT: ILL).

posusanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
213. 1936. TYPE: Peru, Huanuco, Pozuzo. El. 600 m.
20-22 June 1923. Macbride 4609 (HT: F 535686; IT: ILL).

potamophilum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 231,
fig. 197. 1950. TYPE: Colombia, Narifio, S end of Gorgona
Island, near sea level. 9 Feb. 1939. Killip & Garcia 33108
(HT: US 1770099; IT: ILL).

pothophyllum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
214. 1936. TYPE: Peru, Loreto, La Victoria on the Amazon.
Aug.-Sep. 1929. Williams 2998 (HT: F 608343; IT: IL).

praeclarum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
214. 1936. TYPE: Peru, Junin, La Merced, Hacienda Schunke.
El. 1200 m. Aug.-Sep. 1923. Macbride 5623 (HT: F 536662;
ITe SLL) 2

praeclarum Trel., Ciencia (Mexico) 2: 207. 1941, non Trel. in
Macbr. (1936). TYPE: Peru, Chanchamayo. 22 Oct. 1863.
Isern 2183 (HT: MA; IT: ILL).-- [The specimen at ILL was
filed by Trelease under P. mitifolium]

72 PHY T°O°LeO' 6 IVA Vol. 58, No. 1

praemollitum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 352.
1938. TYPE: Honduras, Cortés, near shore of Lake Yojoa.
El. 630 m. 29 July 1934. Yuncker 4825 (HT: ILL).--
tr P. atlantidanum (fide Standley & Steyerm., 1952);

. pseudo-fuligineum (ef. Burger, 1971) ]

praesagium Trel. & Yuncker, Piperaceae no. S, Amer. 1: 308, fig.
273. 1950. TYPE: Colombia, Narino, E side of Gorgona Island.
El. 50-100 m. 11 Feb. 1939. Killip & Garcia 33158 (HT: US
1770140; IT: ILL; PT at ILL: Killip & Garcia 33348).

praeterlatum Trel., J. Wash. Acad. Sci. 19: 335. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
11 Jan. 1928. Standley 54337 (HT: F 583571; IT: ILL).--
(= P. aeruginosibaccum (fide Standley & Steyerm., 1952); = P.

jacquemontianum (cf. Burger, 1971) ]

prismaticum C.DC. var. villosulum Trel., Contr. U.S. Nat. Herb.
26(4): 135. 1929. TYPE: Costa Rica, Limon. Feb. 1926.
Standley & Valerio 48570 (HT: US 1307545; PT at ILL: Standley
& Valerio 50971, 51234).-- [= P. augustum Rudge (cf. Burger,
1971) ]

procerum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
214. 1936. TYPE: Peru, Junin, La Merced, Schunke Hacienda.
El. 1400-1700 m. 8-12 June 1929. Killip & Smith 24636 (HT:
US *EPs FEL);

prodigum Trel., J. Wash. Acad. Sci. 19: 335. 4929.0 “TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
22 Jan. 1928. Standley 54916 (HT: F 582776; IT: ILL).--
[= P. fraguanum (fide Standley & Steyerm., 1952); = P
sancti-felicis (fide Burger, 1971)]

propingquum C.DC. var. brevistylum Trel. in Macbr., Field Mus.
Publ. Bot. 13(No. 357): 215. 1936. TYPE: Peru, Huanuco,
Cochero. Poeppig 1037 [HT: W; the specimen at ILL is Poeppig
1337, obtained from W (that number cited with a BPR) s)- this
either an IT or a PT}].-- f= affin- P. crassinervium (cf.
Yuncker, 1972) ]

propinquum var. grande Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 215. 1936. TYPE: Peru, Loreto, Yurimaguas to
Balsapuerto. El. 135-150 m. 26-31 Aug. 1929. Killip &
Smith 28109 (HT: US; IT: ILL; PT at ILL: Killip & Smith
27757, 28734).-- [= P. crassinervium (cf. Yuncker, 1972) ]

1985 Jones, Annotated catalogue of type specimens 13

provulgatum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 277,
fig. 239. 1950. TYPE: Colombia, Norte de Santander, Tapata,
Caluga Valley. El. 1500-2100 m. March 1927. Killip & Smith
20478 (HT: US 1355516; IT: ILL).

pseudoarboreum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 113, fig.
99. 1966. TYPE: Brazil, Acre, near mouth of Rf{o Macauhan.
8 Aug. 1933. Krukoff 5349 (HT: NY; IT: ILL; PT at ILL:
Krukoff 5713).-- [This species upheld by Yuncker, 1973]

pseudo-callosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 215. 1936. TYPE: Peru, Hudnuco, Huacachi near Muna.
El. 1950 m. May-June 1923. Macbride 4103 (HT: F 535172; IT:
ILL; PT at ILL: Macbride 4333).

pseudoglabrescens Trel. & Yuncker, Piperaceae no. S. Amer. 1:
228, fig. 193. 1950. TYPE: Venezuela, Bolfvar, Rfo Paragua,
Isla El Casabe. El. 265 m. 9 April 1943. Killip 37312 (HT:
US 1855431; IT: ILL).

pseudo-lanceaefolium Trel., Contr. U.S, Nat. Herb. 26( 4): 170.
1929. TYPE: Costa Rica, La Palma. El. 1450 m. Sep. 1898.
Tonduz 12665 (HT: US 798273; PT at ILL: Maxon & Harvey 8375;

Standley 41822; Standley & Valerio 43481, 48046).-- [= P. |
lanceaefolium (cf. Burger, 1971) ]

pseudo-matico Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 217. 1936. TYPE: Peru, Huadnuco, Mito. El. 2700 m.
8-22 July 1922. Macbride & Featherstone 1488 (HT: F 517987;
ITs bhbes (PE at ALby KELL Killip & Smith 24460; “Macbride 3327).

pseudo-mollicomum C.DC. in DC., Prodromus 16(1): 278. 1869.
TYPE: Peru, Loreto, Mainas. Poeppig 2290 (HT: G-DC ?; IT:
EEL).

pseudopopayanense Trel. & Yuncker, Piperaceae no. S. Amer. 1:
191, fig. 150. 1950. TYPE: Colombia, Cauca, highlands of
Popayan. El. 1600-2000 m. Lehmann K310 (HT: US 1420258;
PT at ILL: Ewan 16567; Haught 5290).

pseudo-variabile Trel., Contr. U.S. Nat. Herb. 26(2): 23. 1927.
TYPE: Panama, Colon, Loma de la Gloria, near Fato. Pittier
3845 (HT: US 678920; PT at ILL: Standley 25179).-- [= P
grande (fide Yuncker, 1950) ]

Seeso eta var. collium Trel., Contr. U.S. Nat. Herb.
26(2): 24. 1927. TYPE: Panama, Prov. Panama, along the Rio
Tapia. evi aaa (HT: US 1153478; PT at ILL: Standley
27191, 27555).-- [= P. grande (fide Yuncker, 1950) ]

74 P HVT LO GLA Vol. 58, No.

pseudo-variabile var. pachypus Trel., Contr. U.S. Nat. Herb.
26(2): 24. 1927. TYPE: Panama, between Frijoles and Monte

Lirio. El. 30 m. 18 Oct. 1922. Killip 12145 (HT: US
1167279; IT: ILL (2 sheets)].-- [= P. grande (fide Yuncker,
1950) ]

pseudo-viridicaule Trel. var. nievecitanum Trel. in Woodson &
Schery, Ann. Mo. Bot. Gard. 27: 296. 1940. TYPE: Panama,
Bocas del Toro, vicinity of Nievecita. Aug. 1938. Woodson,
Allen & Seibert 1829 (HT: ILL).-- [= P. hispidum (fide
Yuneker, 1950 and 1957); = P. sancti- felicis (fide Burger,
1971) ]

psilophyllum C.DC. forma longepedunculatum Yuncker, Lilloa 27:
148, pl. 28. [1953] 1955. TYPE: Bolivia, without definite

locality. Bang 2900 (HT: NY; PT at ILL: Cardenas 1331; Tate
639).

psittacorum Endl., Prodromus Fl. Norf., 37. 1833. TYPE: Norfolk
Island. Bauer (HT: W?; IT: ILL).

puberulescens Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 218. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 29057 (HT: US; IT: ILL).

puberulibracteum C.DC. var. monticola Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 144, fig. 122. 1950. TYPE:
Colombia, Santander, vicinity of Las Vegas. El. 2600-3000 m.
Dec. 1926. Killip & Smith 15943 (HT: US 1351739; IT: ILL).

puente-altoanum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 280.
1940. TYPE: Honduras, Atlantida, Puente Alto, on the
railroad about 50 km E of La Ceiba. El. 240 m. 19 July
1938. Yuncker, Koepper & Wagner 8549 (HT: ILL).--

[= P. fraguanum (fide Standley & Steyerm., 1952); = P.
sancti-felicis (fide Burger, 1971)]

* pullibaccum Trel., J. Arnold Arb. 9: 149. 1928. TYPE: New
Guinea, Papua, Vaitata River. 13 Feb. 1926. Brass 966
(LECTOTYPE designated herein: A; ILT: ILL).-- [This species
recognized by Chew, 1972]

pullispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. a57)::
218. 1936. TYPE: Peru, Loreto, Pongo de Manseriche.
El. 280 m. 26 Dec. 1931. Mexia 6352 (HT: UC; IT: ILL).

punctiunculatum Trel., Contr. U.S, Nat. Herb. 26(4): 180. 1929.
TYPE: Costa Rica, Guanacaste, Tilaran. El. 600 m.
Jan. 1926. Standley & Valerio 45712 (HT: US 1307393; PT at
ILL: Standley & Valerio 44975, 45048).

1985 Jones, Annotated catalogue of type specimens 75

puraceanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 205, fig.
166. 1950. TYPE: Colombia, El Cauca, "Canaan", Mt. Puracé.
El. 3100-3300 m. June 1922. Pennell & Killip 6585 (HT: US
Traetass firs TLL);

purpusianum Trel., J. Wash. Acad. Sci. 16: 205. 1926. TYPE:
Mexico, Chiapas, S. Sierra Madre, Hacienda Fenix. 1925.
Purpus 196 (HT: US 1209376; IT: ILL).

putumayoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 47,
fig. 37. 1950. TYPE: Colombia, Putumayo, Rfo Putumayo,
Puerto Ospina. El. 230 m. 14 Nov. 1940. Cuatrecasas 10561
(HT: US 1799093; IT: ILL).

* quentinii Trel. ex Stehlé, Bull. Soc. Bot. France 84: 410. 1938,
nomen subnudum (as quentini); Candollea 8: 74. 1940. TYPE:
Guadeloupe, Ravine Malanga. El. 750 m. 1 Jan. 1937.

Quentin & Stehlé 1337 [LECTOTYPE designated herein: NY;
probable ILT: ILL (the number on the specimen at ILL is 1737
but it was obtained from NY and is probably a duplicate of
No. 1337. Howard annotated the sheet at ILL as HT and the
Sheet at NY as IT); PT at ILL: Martinique, Stehlé & Stehlé
3228, 3351, 3395).-- [= P. glabrescens (fide Howard, 1973) ]

quillandanum [stet!] Trel. & Ekman in Feddes Repert. Spec. Nov.
Regni Veg. 26: 341. 1929. TYPE: Haiti, La Hotte, Quillaud,
Les Roseaux. El. 100 m. 26 June 1928. Ekman 10115 (HT:
ILL; PT at ILL: Ekman 10679).

quimirianum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
219. 1936. TYPE: Peru, Junfn, La Merced, E of Quimiri
Bridge. El. 800-1300 m. 1-3 June 1929. Killip & Smith
23904 (HT: US; IT: ILL).

* guiriguanum Trel. in Standley, Field Mus. Publ. Bot. 10: 160.
1931. Type locality: Honduras, "in wet thickets near
Progreso, also along the coast of Guatemala." [No type cited
but probable HT (or LECTOTYPE designated herein) at ILL-- the
Sheet marked "type" by Trelease: Honduras, Yoro, wet thicket
near Progreso. El. 30 m. 24 Jan. 1928. Standley 54995].--
{= P. patulum (fide Standley & Steyerm., 1952); = P.
marginatum (cf. Burger, 1971) ]

raizudoanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 291,
fig. 257. 1950. TYPE: Colombia, Santander, headwaters of
Dorado Creek, 12 km S of Raizudo. El. ca 300 m. 6 May 1937.
Haught 2191 (HT: US 1742421; IT: ILL; PT at ILL: Haught
1812).

PHYTOL GEDA Vol. 58, No. 1

rasile Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 220.
1936. TYPE: Peru, Jun{n, Puerto Bermddez. El. 375 m. 14-17
July 1929. Killip & Smith 26404 (HT: US; IT: ILL).

ratisportanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 220. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 150-
350 m. 28-30 Aug. 1929. Killip & Smith 28570 (HT: US; IT:
ILL).

recensitum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
220. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m.
1-5) Sep.. 1929. Killip & Smith 28810 (HT: US; IT: ILL).

reciprocum Trel., Ciencia (Mexico) 2: 207. 1941. TYPE: Peru,
Chanchamayo. 23 Oct. 1863. Isern 2310 (HT: MA; IT: ILL).

recreoense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
220. 1936. TYPE: Peru, Loreto, Recreo, Yurimaguas.
El. 155-210 m. Oct.-Nov. 1929. Williams 3928 (HT: F 622942;
a7 2 3h)

rectispicum Trel. in Rusby, Mem. N.Y. Bot. Gard. 7: 225. 1927.
TYPE: Bolivia, Bopi River Valley. El. 3000 ft. Sep. 1921.
Rusby 574 (HT: NY; IT: ILL).-- (= P. mapirense C.DC. (fide
Yuncker, 1955) ]

reductipes Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
221. 1936. TYPE: Peru, Loreto, La Victoria on the Amazon
River. Aug.-Sep. 1929. Williams 2700 (HT: F 617253; IT:
ILL,

reductipes var. amplum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 221. 1936. TYPE: Peru, Loreto, La Victoria on
the Amazon River. Aug.-Sep. 1929. Williams 2971 (HT: F
606157; ITs ILL).

reductipes var. stenostachys Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 221. 1936. TYPE: Peru, Loreto, La Victoria
on the Amazon River. Aug.-Sep. 1929. Williams 2578 (HT: F
6739350 20s ALL )c

regresivanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 239,
fig. 207. 1950. TYPE: Venezuela, Aragua...between Rancho
Grande and Dos Riitos. El. 900 m. 19, 20 May 1943. Killip
& Lasser 37759 (HT: US 1855733; IT: ILL).

rematense Trel. in Standley, Field Mus. Publ. Bot. 17: 233.
1937. TYPE: Guatemala, Petén, Lake Petén, Remate. 25 March
1933. Lundell 2087 (HT: MICH; IT: ILL).-- [= P. martensianum
(fide Standley & Steyerm., 1952)]

1985 Jones, Annotated catalogue of type specimens 77

resalutatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
222. 1936. TYPE: Peru, Junfn, Pichis Trail, San Nicolas.
El. 1100 m. 4, 5 July 1929. Killip & Smith 26024 (HT: US;
fs) LEE

restio Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 222.
1936. TYPE: Peru, Huadnuco, Pozuzo. El. 600 m. 20-22 June
1923. Macbride 4689 (HT: F 535763; IT: ILL).

restrictum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
223. 1936. TYPE: Peru, Junfn, Cahuapanas. El. 340 m.
20, 21 July 1929. Killip & Smith 26737 (HT: US; IT: ILL).

* rhinostachyum Trel. in Macbr., Fiéld Mus. Publ. Bot. 13(No.
357): 224. 1936. TYPE not cited but almost certainly an
inadvertent omission (specimens marked "Type" at US and ILL):
Peru, Junfn, La Merced. El. 700 m. May-June 1929. Killip &
Smith 23461 (LECTOTYPE designated herein: US 1358149; ILT:
FEL}.

* rhizocaule Trel., J. Arnold Arb. 9: 149, 1928. TYPE: New
Guinea, Papua, Vaitata River, Kira. 9 March 1928. Brass
1115 (LECTOTYPE designated herein: A; ILT: ILL).-- [= P.
versteegii C.DC. (fide Chew, 1972) ]

* rhodocarpum Trel., J. Arnold Arb. 9: 149. 1928. TYPE: New
Guinea, Papua, U-Uma River. El. 150 m. 14 May 1926. Brass
1427 (LECTOTYPE designated herein: A; ILT: ILL).-- [= P.
mestonii Bailey (fide Chew, 1972) ]

riitosense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 396, fig.
357. 1950. TYPE: Venezuela, Aragua, Parque Nacional, Dos
Riitos. El. 600 m. 19, 20 May 1943. Killip & Lasser 37755
(HT:, US 18557315.1T: ILL).

rinconense Trel. in Yuncker, Field Mus. Publ. Bot. 17: 353.
1938. TYPE: Honduras, Comayagua, near El Rincon, ca 10 miles
W of Siguatepeque. El. 1400-1500 m. 24 July 1936. Yuncker,
Dawson & Youse 6052 (HT: ILL).

x rio-chiadoense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 229,
fig. 194. 1950. TYPE: Colombia, Antioquia, Rio Chiado, 65 km
S of Turbo. El. 60 m. 7 May 1945. Haught 4643 {HT, or
LECTOTYPE designated herein: US 1709187 (= fig. 194); IT or
ILT: US 1709186; PT at ILL: Cuatrecasas 17590].

riojanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
224. 1936. TYPE: Peru, Amazonas, Rioja, Moyobamba.
El. 2700-3300 m. Jan. 1930. Williams 7600 (HT: F 629721;
it = JES.

78

raRQYiteLteet B Vol. 58, No. |

rio-marguanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 330,
fig. 298. 1950. TYPE: Colombia, Norte de Santander, in
region of Sarare, Rfo Margua Valley. El. 2000 m. 7 Nov.
1941. Cuatrecasas 12847 (HT: US 1851044; IT: ILL).

rio-nechianum Yuncker in Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 211, fig. 174. 1950. TYPE: Colombia, Antioquia, Dos
Bocas, R{o Nechi, near Pato. June 1944. Ewan 15827 (HT:
ILL).

rio-zulianum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 50,
fig. 40. 1950. TYPE: Colombia, Norte de Santander, Rio
Zulia. 18 May 1941. Nicéforo 143 (HT: ILL).

rivialbi Trel. & Yuncker, Piperaceae no. S. Amer. 1: 265, fig.
229. 1950. TYPE: Colombia, El Valle, Rfo Digua Valley,
between La Elsa and Rfo Blanco. El. 825 m. April 1939.
Killip au72s UAT: US" TTT1521= 11:. 1bb).

robleanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 270, fig.
235. 1950. TYPE: Colombia, Santander, vicinity of El Roble.
El. ca 1500 m. 16 Feb. 1927. Killip & Smith 19372 (HT: US
1354629; IT: ILL).

rogaguanum Trel. in Rusby, Mem. N.Y. Bot. Gard. 7: 2255 Sets
TYPE: Bolivia, pampas near Lake Rogagua. El. 1000 ft.
7 Nov. 1921. Rusby 1689 (HT: NY; IT: ILL).-- [This species
recognized by Yuncker, 1955]

roseovenulosum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 353.
1938. TYPE: Honduras, Cortés, near Paral, shore of Lake
Yojoa. El. 630 m. Yuncker 4873 (HT: ILL).

rotundibaccum Trel., Contr. U.S. Nat. Herb. 26(4): 164. 1929.
TYPE: Costa Rica, San José, Quebradillas, Santa Maria de
Dota. El. 1800 m. 24 Dec. 1925. Standley 42902 [HT: US
1307279; PT at ILL: Standley 41285 (identification
goubtten) 1. <-‘ [=P carpinteranum (fide Burger, 1971) ]

rotundibaccum var. fraijanesanum Trel., Contr. U.S. Nat. Herb.
26(4): 164. 1929. TYPE: Costa Rica, Alajuela, Fraijanes.
El, 1600 m. 12, 13 Feb. 1926. Standley & Torres 47669 (HT:
US 1307511; IT: ILL).-- [= P. carpinteranum (fide Burger,
1971) ]

rotundistipulum Trel. & Yuncker, Piperaceae no, Ss. Amer. 1: 406,
fig. 370. 1950. TYPE: Colombia, Putumayo, Rio Putumayo,
Puerto Porvenir, etc. El. 250 m. 17 Nov. 1940. Cuatrecasas
10596 (HT: US 1798150; PT at ILL: Mexia 6951).

1985 Jones, Annotated catalogue of type specimens 79

rubiginosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
224. 1936. TYPE: Peru, Junfn, Puerto Yessup. El. 400 m.
10-12 July 1929. Killip & Smith 26292 (HT: US; IT: ILL).

rubribaccum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
225. 1936. TYPE: Peru, Hudnuco, Pozuzo. El. 600 m.
20-22 June 1923. Macbride 4565 (HT: F 535642; IT: ILL).

rubripes Trel. in Standley, Field Mus. Publ. Bot. 18: 358. 1937.
TYPE: Costa Rica, Guanacaste, upper slopes of Cerro San José
de L{pano. El. 500-960 m. 15 Feb. 1930. Dodge, Hanckel &
Thomas 6382 (HT: GH; IT: ILL).-- [= P. aequale (cf. Burger,
1971) J

rubrospadix Trel. in Standley, Field Mus. Publ. Bot. 18: 358.
1937. TYPE: Costa Rica, Limon, Bonilla Lakes. El. 300-
430 m. 13 Dec. 1929. Dodge, Catt & Thomas 5718 (HT: GH; IT:
ILL).-- [= P. glabrescens (fide Burger, 1971) ]

rugosifolium Trel., Contr. U.S. Nat. Herb. 26(4): 185. 1929.
TYPE: Costa Rica, Cartago, Finca Las Concavas. El. 1200-
1300 m. 7, 8 Dec. 1925. Standley 41533 (HT: US 1307251;
IT: ILL; PT at ILL: Standley & Valerio 49441).-- [= P.
perhispidum C.DC. (fide Burger, 1971) ]

rurrenbaqueanum Trel. in Rusby, Mem. N.Y. Bot. Gard. 7: 225.
1927. TYPE: Bolivia, Rurrenbaque. El. 1000 ft. 4 Oct.
1921. Rusby 836 (HT: NY; IT: ILL).-- [= P. buchtienii (fide
Yuneker, 1955) ]

rusbyi C.DC. var. hirsutum Yuncker, Lilloa 27: 139. [1953] 1955.
TYPE: Bolivia, La Paz, N. Yungas, outskirts of Coroica.
El. 2000 m. 5 Dec. 1935. Mexia 7792 (HT: US 1778378; IT:
ILA)...

Sabanillanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
6, 9. 1926. TYPE: Cuba, Pinar del Rfo, San Diego de Tapias
to Sabanilla, Sierra de los Organos. 3 April 1920. Ekman
10659 [?HT (or fragments of type): ILL].

sagax Trel. in Yuncker, Field Mus. Publ. Bot. 9: 280. 1940.
TYPE: Honduras, Yoro, Coyoles. 27 June 1938. Yuncker,

Koepper & Wagner 8030 (HT: ILL).

Salamancanum Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27:
296. 1940. TYPE: Panama, Canal Zone, vicinity of Salamanca
Hydrographic Station, Rio Pequeni. El. ca 80 m. July 1938.
Woodson, Allen & Seibert 1590 (HT: ILL).-- [= P. pinoganense
Trel. (fide Yuncker, 1950) ]

80

P MutT O. beQom@s Ter Vol. 58, No.

salentoi Trel. & Yuncker, Piperaceae no. S. Amer. 1: 241, fig.
209. 1950. TYPE: Colombia, Caldas, Cordillera Central, Rio
Santa Rita, Salento. El. 1600-1800 m. 29 July 1922. Killip
& Hazen 8962 (HT: PH; PT at ILL: Pennell 8940).

salentoi var. cajamarcanum Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 241. 1950. TYPE: Colombia, Tolima, along Quindfo
Hwy. between Cajamarca and summit of divide. El. 2400 m.
27-28 March 1939. Killip & Varela 34530 (HT: US 1771364;
ifs 1):

salgaranum Trel. ex Trel. & Yuncker, Piperaceae no, S. Amer. 1:
269, fig. 234. 1950. TYPE: Colombia, Antioquia, Salgar,
vicinity of Medellfn. 5 April 1928. Toro 1066 (HT: NY; IT:
ILL).

salicinum Opiz in Presl, Rel. Haenke. 1: 150. 1825-1830. TYPE:
Mariana Islands (Pacific Isl. Trust Territory, U.S.), leg.
Haenke [said to be from Panama-- fide Index Kewensis] (HT: ?;
fragments of type at ILL).

san-andresense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 240,
fig. 208. 1950. TYPE: Colombia, Huila, Cordillera Oriental,
below Gabinete in the ravine of the Abra de San Andrés.

El. 1900-2100 m. 24 March 1940. Cuatrecasas 8615 (HT: US
1796603; IT: ILL).

san-cristobalanum Trel. in Standley, Field Mus. Publ. Bot. 18:
359. 1937. TYPE: Costa Rica, San Cristobal Road.
El. 8000 ft. 27 May 1928. Stork 2205 (HT: ILL).-- [= P.
euryphyllum (fide Burger, 1971) ]

sandaloense Trel. in Standley, Field Mus. Publ. Bot. 18: 359.
1937. TYPE: Costa Rica, Puntarenas, Rfo Sandalo, Peninsula
de Osa, at sea level. Dodge & Goerger 10076 (HT: MO; IT:
ILL).-- [= P. nudifolium C.DC. (fide Burger, 1971)]

sangabrielanum Trel. in Gleason, Bull. Torrey Bot. Club 58: 354.
1931. TYPE: Brazil, Rfo Negro, San Gabriel. 15 Sep. 1928.
Tate s.n., Tyler-Duida Exped. 1002 (HT: NY; IT: ILL) = =P.
Sancarlosianum C.DC. (fide Yuncker, 1973)]

sanguineispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 226. 1936. TYPE: Peru, Loreto, Iquitos. El. 100 m.
3-11 Aug. 1929. Killip & Smith 27135 (HT: US; IT: ILL).

san-joseanum C.DC. var. chiriquinum Trel. in Woodson & Seibert,
Ann. Mo. Bot. Gard. 25: 626. 1938. TYPE: Panama, Chiriqui,
between Rf{o Tabasara and Rio Tinta. 11 Aug. 1937. Woodson,
Allen & Seibert 416 [HT: MO; fragment of type (one leaf) at
ILL).-- [= P. marginatum (fide Yuncker, 1950 and 1957) ]

]

1985 Jones, Annotated catalogue of type specimens 81

san-joseanum var. kobense Trel. in Woodson & Schery, Ann. Mo.
Bot. Gard. 27: 297. 1940. TYPE: Panama, Canal Zone, Fort
Kobe Road. 22 July 1938. Woodson, Allen & Seibert 1423 (HT:
ILL).-- (= P. marginatum (fide Yuncker, 1950 and 1957) J

San-joseanum var. panamanum Trel. in Woodson and Schery, Ann.
Mo. Bot. Gard. 27: 297. 1940. TYPE: Panama, Gorgona Beach.
7 Aug. 1938. Woodson, Allen & Seibert 1690 (HT: ILL).--
(= P. marginatum (fide Yuncker, 1950 and 1957)]

San-joseanum var. remediosense Trel. in Woodson & Schery, Ann.
Mo. Bot. Gard. 27: 297. 1940. TYPE: Panama, Chiriquf, Rfo
Chiriquf to Remedios. El. 15-50 m. 11 July 1938. Woodson,
Allen & Seibert 1191 (HT: ILL).-- [= P. marginatum (fide
Yunecker, 1950 and 1957) ]

san-joseanum var. tabogense Trel. in Woodson & Schery, Ann. Mo.
Bot. Gard. 27: 297. 1940. TYPE: Panama, Prov. Panama, Isla
Taboga. July 1938. Woodson, Allen & Seibert 1531 (HT:
ILL).-- [= P. marginatum (fide Yuncker, 1950 and 1957)]

san-juananum Trel. in Standley, Field Mus. Publ. Bot. 11: 147.
1936. TYPE: El Salvador, San Juan de Tepezones. 1929.
Calderén 2539 (HT: F 603261; IT: ILL).

san-martinense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 410,
fig. 373. 1950. TYPE: Colombia, Santander, E. Cordillera, S
slope of Mt. San Mart{n, near Charta. El. 2300-2500 m.

10 Feb. 1927. Killip & Smith 19160 (HT: US 1354458; IT: ILL;
PT at ILL: Killip & Smith h 19181).

san-miguelense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 280,
fig. 243. 1950. TYPE: Ecuador, San Miguel. 4 Nov. 1924.
Stevens 204 (HT: ILL).

san-nicolasense Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 227. 1936. TYPE: Peru, Junfn, Pichis Trail, San
Nicolas. El. 1100 m. 4-5 July 1929. Killip & Smith 26009
CHF 20S = 2T2 ILbP.

San-ramonense Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 227. 1936. TYPE: Peru, Junfn, San Ramon. El. 1400-
1700 m. 8-12 June 1929. Killip & Smith 24585 (HT: US; IT:
TEL).

san-roqueanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 227. 1936. TYPE: Peru, San Mart{n, San Roque.
El. 1350-1500 m. Jan.-Feb. 1930. Williams THO] (HT: F
623525; IT: ILL; PT at ILL: Killip & Smith 27306).

82

P Mok TO &OcGal Vol. 58, No. |

* santa-ritanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.

357): 227. 1936. TYPE: Peru, Cuzco, Santa Rita, Santa Ana
Valley. El. 2300 m. Aug. 1927. Herrera 1551 (HT, or
LECTOTYPE designated herein: ILL).

san-vicentense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 19,
fig. 7. 1950. TYPE: Colombia, Santander, E.Cordillera, W
Slope of Mt. San Vicente, near Charta. El. 2500-2700 m.

9 Feb. 1927. Killip & Smith 18984 (HT: US 1354299; IT: ILL;
PT at ILL: Killip & Smith 18314).

sapoto-yacuense Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 228. 1936. TYPE: Peru, Loreto, Sapoto Yacu, Santa
Rosa. El. 155-210 m. Oct.-Nov. 1929. Williams 4923 (HT: F
624945; IT: ILL; PT at ILL: Williams 5027).

saramaccanum Yuncker in Maguire & Collaborators, Bull. Torrey
Bot. Club 75: 288. 1948. TYPE: Suriname, km. 3.5, vicinity
Krappa Camp (2), Saramacca River headwaters. 3 Oct. 1944.
Maguire 24887 (HT: NY; IT: ILL).-- [This species recognized
by Trel. & Yuncker, 1950 and by Yuncker, 1957]

sasaimanum Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.

1: 123, fig. 98. 1950. TYPE: Colombia, Cundinamarca,
Estacidén, Santana, above Sasaima. El. 1600-1700 m.

25-29 July 1945. Dugand & Jaramillo 3860 (HT: US 1854010; PT
at ILL: Garcfa Barriga 11900).

satisfactum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):

228. 1936. TYPE: Peru, Junin, Puerto Yessup. El. 400 m.
10-12 July 1929. Killip & Smith 262495 CHT: US; IT: ILL).

scabiosifolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.

357): 229. 1936. TYPE: Peru, Loreto, Puerta Arturo.
Elz, 435 m. »«25-25,Aug. 1929. , Killip..& Smith 2d1307tnL US;
ats. ILL}.

scabrellum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 67, fig. 56.

1966. TYPE: Brazil, Sao Paulo, Itapira. 16 May 1927.
Hoehne 20334 (cited as Hoehne s.n.) (HT: ILL).-- {This
species upheld by Yuncker, 1972]

scabricaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):

229. 1936. TYPE: Peru, Junfn, near Perené Bridge, Rfo
Paucartambo. El. 700 m. 19 June 1929. Killip & Smith 25299
(Hts: US+0T: ELE).

scabridulicaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 229. 1936. TYPE: Peru, Loreto, mouth of Rio Santiago.
14 Nov. 1931. Mexia 6100 [HT: UC; probable IT (Mexia
6106-- marked "Type" by Trelease): ILL].

1985 Jones, Annotated catalogue of type specimens 83

scabridulilimbum Trel. & Yuncker, Piperaceae no. S. Amer. 1:
279, fig. 241. 1950. TYPE: Colombia, Santander, E.
Cordillera, Rfo Surata Valley, between Jaboncillo and Surata.
El. 1500-1800 m. 3 Jan. 1927. Killip & Smith 16465 (HT: GH;
PT at ILL: Killip & Smith 16775).

scabriseptum Trel., J. Wash. Acad. Sci. 19: 336. 1929. TYPE:
Honduras, Atlantida, Lancetilla Valley near Tela. El. 100 m.
8 Dec. 1927. Standley 52681 (HT: F 582776; IT: ILL).

scabriseptum var. reductum Trel. in Yuncker, Field Mus. Publ.
Bot. 17: 353. 1938. TYPE: Honduras, Atlantida, Lancetilla
Valley. El. 27 m. 27 July 1934. Yuncker 4773 (HT: ILL).

scabrum Lam. var. kalacroixense Trel. in Feddes Repert. Spec.
Nov. Regni Veg. 23: 308. 1927. TYPE: Haiti, l'Artibonite,
Kalacroix. El. 700 m. 11 Dec. 1925. Leonard 7939 (HT:
ILL).

scalariforme Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
230. 1936. TYPE: Peru, Loreto, Caballo-Cocha on the Amazon
River. Williams 2057 (HT: F 613146; IT: ILL; PT at ILL:
Killip & Smith 29882; Williams 2056).

scalarispicum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 353.
1938. TYPE: Honduras, Comayagua, above El Achote, near
Siguatepeque. El. 1800-2000 m. 28 July 1936. Yuncker,
Dawson & Youse 6156 (HT: ILL).

scandenticaule Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 68, fig.
57. 1966. TYPE: Brazil, Mato Grosso, Ponte de Pedra.
April 1918. Kuhlmann 1954-K (HT: ILL).-- [This species
upheld by Yuncker, 1972]

scansum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 32, fig. 24H.
1950. TYPE: Colombia, Antioquia, N of Dabeiba, along road to
Turbo. El. 300-350 m. 25 Feb. 1942. Metcalf & Cuatrecasas
30183 (HT: ILL; PT at ILL: Haught 5453).

schippianum Trel. ex Standley in Standley & Record, Field Mus.
Publ. Bot. 12: 104. 1936 (without a Latin diagnosis); in
Standley & Steyerm., Fieldiana Bot. 24(3): 323. 1952. TYPE:
British Honduras (Belize): 19-mile, Stann Creek Railway.
El. 90 m. Schipp 316 (HT: F; fragments of type at ILL).

* scutaespicum Trel., Ciencia (Mexico) 2: 208. 1941. TYPE: Peru,
Chanchamayo. 23 Oct. 1863. Isern 2181-c (HT, or LECTOTYPE
designated herein: ILL).-- [= P. hispidum, fide Trel. &
Yuncker annotation]

84

Pot. eT? Ok, On ah Vol. 58, No. 1

sebastianum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 69, fig.
58. 1966. TYPE: Brazil, Sdo Paulo, Ilha Sado Sebastio.
5 Aug. 1895. Loefgren 3063 (HT: SP; IT: ILL).-- [This
species upheld by Yuncker, 1972]

seminervosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 53.
1950. TYPE: Colombia, Antioquia, Penares Blancas. 5 May
1926. Juzepczuk 4702 (HT: LE; fragments of type at ILL).

seminitidulum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 232. 1936. TYPE: Peru, Junin, La Merced. El. 600 m.
Macbride 5289 (HT: F 536314; IT: ILL; PT at ILL: Killip &
Smith 24093, 24964).

semiplenum Trel., Ciencia (Mexico) 2: 208. 194i. TYPE: Peru,
Chanchamayo. 23 Oct. 1863. Isern 2181-d (HT: MA; IT: ILL).

semitarium Trel. & Yuncker, Piperaceae no. S. Amer. 1: 187, fig.
146. 1950. TYPE: Colombia, Antioquia, JericO. Dec. 1940.
Daniel & Tomas 2732 (HT: US 1830391; IT: ILL).

sericeonervosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 233. 1936. TYPE: Peru, Loreto, La Victoria on the
Amazon River. Aug.-Sep. 1929. Williams 2928 (HT: F 608548;
ITs ILL

serotinum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):

233. 1936. TYPE: Peru, Junin, Perené Bridge, Rio
Paucartambo. El. 700 m. 19 June 1929. Killip & Smith 25344
CHT? US =* iT" TL).

setosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 38, fig. 29.
1950. TYPE: Colombia, El Valle, Cordillera Occidental,
"Calima", on Rio Calima. 14-15 Sep. 1922. Killip 11219 (HT:
US 1143179; IT: ILL; PT at ILL: Cuatrecasas 15403).

setulosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
234. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 27568 (HT: US; IT: ILL).

sibunense Trel. ex Standley in Standley & Record, Field Mus.
Publ. Bot. 12: 408. 1936. TYPE: British Honduras (Belize),
Gracie Rock, Sibun River. 24 March 1935. Gentle 1562 (HT:
MICH; IT: ILL).-- [= P. pseudoasperifolium C.DC. (fide
Standley & Steyerm., 1952) ]

sidipilum Trel. in Macbr., Field Mus. Publ. Bot. 130000357):
234. 1936. TYPE: Peru, Junf{fn, Rfo Perené. El. 600 m.
16-18 June 1929. Killip & Smith 25160 (HT: US; IT: ILL).

1985 Jones, Annotated catalogue of type specimens 85

Siguatepequense Trel. in Yuncker, Field Mus. Publ. Bot. 17: 354.
1938. TYPE: Honduras, Comayagua, near Siguatepeque.
El. 1050 m. 23 July 1936. Yuncker, Dawson & Youse 6042 (HT:
ILL).

Siguatepequense var. subrhombifolium Trel. in Yuneker, Field
Mus. Publ. Bot. 17: 354. 1938. TYPE: Honduras, Comayagua,
vicinity of Siguatepeque. El. 1050 m. 3 July 1936.
Yuneker, Dawson & Youse 5617 (HT: ILL).

Ssilencioi Trel. in Standley, Field Mus. Publ. Bot. 18: 360.
1937. TYPE: Costa Rica, Guanacaste, El Silencio. El. 750 m.
13 Jan. 1926. Standley & Valerio 44644 (HT: US; IT: ILL; PT
at ILL: Standley & Valerio 44656, 45053).-- [= P. nudifolium
(fide Burger, 1971) ]

silvanorum Trel., Contr. U.S. Nat. Herb. 26(4): 169. 1929.
TYPE: Costa Rica, San José, Las Nubes. El. 1500-1900 m.
21 March 1924. Standley 38500 (HT: US 1229454; PT at ILL:
Standley & Valerio 51562, 52116, 52230).-- [= P. poasanum
(fide Burger, 1971)]

Silvarum C.DC. var. minus Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 185, fig. 144. 1950. TYPE: Ecuador, Pichincha,
Coraz6n. Sodiro 1/22 (HT: GH; PT at ILL: Sodiro s.n.).

silvigaudens Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 15, fig.
12. 1966. TYPE: Brazil, Amazonas, Manaus. El. 25 m.
17 Oct. 1929. Killip & Smith 30174 (HT: US 1463342; IT:
ILL).-- [This species upheld by Yuncker, 1972]

Ssilvioberrans Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 234. 1936. TYPE: Peru, Loreto, Yurimaguas to
Balsapuerto. El. 135-150 m. 26-31 Aug. 1929. Killip &
Smith 28119 (HT: US; IT: ILL).-- [= P. subsilvestre C.DC.
(fide Yuncker, 1972)]

Sinuatifolium Trel., Contr. U.S. Nat. Herb. 26(4): 147. 1929.
TYPE: Costa Rica, Guanacaste, Los Ayotes, near Tilaran.
El, 600-700 m. 21 Jan. 1926. Standley & Valerio 45537 (HT:
US 1307377; IT: ILL).-- [= P. terrabanum (fide Burger, 1971) ]

sinuclausum Trel. in Macbr., Field Mus. Publ. 13(No. 357): 235.
1936. TYPE: Peru, Huanuco, Maria del Valle. El. 2100 m.
30 April 1930. Macbride 3564 (HT: F 534640; IT: ILL; PT at
ILL: Herrera 2007).

86

BP Hhs¥vF OL cOoGrirk Vol. 58, No. 1

sinuosirameum Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33: 50.
1936 (without a Latin diagnosis). TYPE: Colombia,
Cundinamarca, La Esperanza. El. + 1500 m. 16 April 1932.
Cuatrecasas 2420(a) (HT: ILL).-- [= P. holtonii var.
cundinamarcanum (fide Trel. & Yuncker, 1950) ]

sipenanum Trel. & Yuncker, Piperaceae no. >. Bers: 225-5 fig.
188. 1950. TYPE: Colombia, Putumayo, Rfo San Miguel,
Quebrada de Sipenae. El. 400 m. 12 Dec. 1940. Cuatrecasas
10996 (HT: US 1799103; IT: ILL).

siquirresense Trel. in Standley, Field Mus. Publ. Bot. 18: 361.
1937. TYPE: Costa Rica, Siquirres. 28 May 1928. Stork 2251
(HT: ILL; IT: ILL (2 sheets)].-- [= P. jacquemontianum (fide
Burger, 1971) ]

sociorum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 225, fig.
189. 1950. TYPE: Colombia, Choc6, Bahfa Solano, along
Quebrada Jellita. El. 50-100 m. 22 Feb. 1939. Killip &
Garcfa 33518 (HT: US 1770452; IT: ILL).

soledadense Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
235. 1936. TYPE: Peru, Loreto, Soledad, Rfo Itaya. El.
110 m. 20-22 Sep. Killip & Smith 29769 (HT: US; IT: ILL; PT
at ILL: Killip & Smith 27101, 27134, 28107, 29598, 29711).

soledadense var. angustum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 236. 1936. TYPE: Peru, Loreto, San Antonio, Rio
Itaya. El. 110 m. 18 Sep. 1929. Killip & Smith 29350 (HT:
US7 IT: Men).

sonsonatense Trel. ex Standley & Calder6n, Lista prelim. pl. El
Salvador, 61. 1925, nomen subnudum. Type locality: El
Salvador, Sonsonate, Nahulingo. Standley 22028 (specimen
marked as "Type" by Trelease at ILL = LECTOTYPE designated
herein).

soricicaudatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 236. 1936. TYPE: Peru, Junfn, Puerto Bermidez.
E14 375 ma! (AZT Juby 19295 5¢KilLip & Smith 26661 (HT: US;
iT >414.),

speratum Trel., J. Wash. Acad. Sci. 19: 336. 1929.5 (TYPE:
Honduras, Atlantida, Tela, at sea level. 27 Jan. 1928.
Standley 55184 (HT: F 583272; IT: ILL).-- [= P. fraguanum
(fide Standley & Steyerm., 1952)]

spoliatum Trel. & Yuncker, Piperaceae no. S) Amer. Tt: 116, fig.
92. 1950. TYPE: Colombia, Narino, E side of Gorgona Island.
El. 50-100 m. 11 Feb. 1939. Killip & Garcia 33168 (HT: US
LT7OINT se ITs ILL}.

1985 Jones, Annotated catalogue of type specimens 87

Sstandleyi Trel. in Standley, J. Wash. Acad. Sci. 13: 366. 1923.
TYPE: El Salvador, Volc&n de San Salvador. El. 1800 m.
7 April 1922. Standley 22894 (HT: ILL).-- [This species
recognized by Standley & Steyerm., 1952]

stehleorum Trel. ex Stehlé, Fl. Deser. Antill. Fr. 2(1)-: 100.
1940; Bull. Agric. Martinique 9(3): 177, tab. 6. 1940. TYPE:
Martinique, "Gorges Se la Falaise." El. 750 m. 24 May 1939.
Stehlé & Stehlé 3257 (HT and IT: ILL; PT at ILL: Stehlé &
Stehlé 3209, 09, 3240, 3 3244).-- [= P. aduncum (fide Howard,
1973) ]

stevensi Trel., Contr. U.S. Nat. Herb. 26(2): 27. 1927. TYPE:
Panama, Canal Zone, Frijoles. 14 Oct. 1924. Stevens 1247
(HT: ILL).-- [This species recognized by Yuncker, 1950]

stevensi Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
237. 1936, non Trel. (1927). TYPE: Peru, Junin,
Huacapistana. 6 Dec. 1924. Stevens 85 (HT: ILL).--
[Originally labelled P. elongatum- probably the correct
disposition]

Stillans Trel. & Standley in Standley & Steyerm., Fieldiana
Bot. 24(3): 325. 1952. TYPE: Guatemala, Quiché, Cerro Putul,
Zona Reina. El. 1200 m. 3 Dec. 1934. Skuteh 1822 (HT: F;
ITEC TES:

Striatinervosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 237. 1936. TYPE: Peru, Loreto, Puerto Arturo.
Et on 35 rm o24-25. Auge N92 o Killip & Smith 27910 (HT: US;
IT seit}.

Strigillosicaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 237. 1936. TYPE: Peru, Loreto, Yurimaguas. El. 135 m.
Aug.-Sep. 1929. Killip & Smith 27615 (HT: US; IT: ILL).

Sstrigillosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
238. 1936. TYPE: Peru, Loreto, Puerto Arturo. El. 135 m.
24-25 Aug. 1929. Killip & Smith 27813 (HT: US; IT: ILL).

Strigosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
238. 1936. TYPE: Peru, Loreto, Balsapuerto. El. 350-550 m.
29 Aug. 1929. Killip & Smith 28538 (HT: US; IT: ILL).

Stuebelii Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
238. 1936. TYPE: Peru, Loreto(?), Rio Huallaga. Stuebel 68
(HT: B; PT at ILL: Williams 7056).

88

PoY TeOoke@® 6 BPA Vol. 58, No. 1

subaequilaterum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 280.
1940. TYPE: Honduras, Atlantida, Roma siding, about 25 km E
of La Ceiba. 21 July 1938. Yuncker, Koepper & Wagner 8565
(HT: ILL).-- [= P. atlantidanum (fide Standley & Steyerm.,
1952); = P. pseudo-fuligineum (cf. Burger, 1971) ]

subcinereum C.DC. var. perhirsutum Yuncker, Bol. Inst. Bot. (Sao
Paulo) 3: 71, fig. 61. 1966. TYPE: Brazil, Minas Gerais,
Mun. de Belo Horizonte, etc. 26 July 1942. Magalhnaes 3224
(HT: US 2124365; PT at ILL: Loefgren 800).-- [This variety
upheld by Yuncker, 1972]

subcordulatum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 239. 1936. TYPE: Peru, Loreto, Soledad, Rfo Itaya.
El. 110 m. 20-22 Sep. 1929. Killip & Smith 29794 (HT: US;
je eee OS

subdilatatum Trel., Contr. U.S. Nat. Herb. 26(2): 35. 1927.

TYPE: Panama, Prov. Panama, Taboga Island. 31 Oct. 1917.
Killip 3194 (HT: US 863177; IT: ILL).-- [= P. taboganum
(fide Yuncker, 1950 and annotation; = P. pseudo-fuligineum
(ef. Burger, 1971)]

sudflavum C.DC. var. longipedunculatum Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 313. 1950. TYPE: Colombia, Cauca
Valley, Popayan, Rfoblanco. El. 1800 m. 9 July 1939.
Arbelaez & Cuatrecasas 5796 (HT: US 1774257; PT at ILL:

Killip 7188).

subpatens Trel. in Macbr., Field Mus. Publ. Bot. 13(No..- 357):
240. 1936. TYPE: Peru, Hudnuco, Pozuzo. El. 600 m.
20-22 June 1923. Macbride 4633 (HT: F 535708; IT: ILL).

subpedale Trel. & Yuncker, Piperaceae no. S. Amer. 1: S3,oF ig.
43. 1950. TYPE: Colombia, Chocd, Andagoya. El. 70-100 m.
20-30 April 1939. Killip 35058 (HT: US 1771813; IT: ELL).

subpuberulum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
4, 7. 1926. TYPE: Cuba, Pinar del Rfo, Rio San Miguel.
Wilson 9291. [HT: ?; PT at ILL: Ekman 13869 (2 sheets),
17328 (2 sheets), 17486].

Subrepens Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 24:
187. 1937. TYPE: Panama, Coclé, western slope and summit of
Cerro Valle Chiquito. El. 700-800 m. 25 July 1935. Seibert
504 (HT: MO; IT: ILL).-- [This species upheld by Yuncker,
1950)

1985 Jones, Annotated catalogue of type specimens 89

subscabridum C.DC., Notizbl. Bot. Gart. Berlin-Dahlem 6: 455.
1917. TYPE: Brazil, Parana, Serra do Mar, Ypiranga in silva
primaeva. Dusen 3626 (HT: S; IT: ILL).-- [= P. dilatatum
(fide Yuncker, 1972) ]

subsessile Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
240. 1936. TYPE: Peru, Loreto, Santa Rosa. El. 135 m.
1-5 Sep. 1929. Killip & Smith 28747 (HT: US; IT: ILL; PT at
ILL: Killip & Smith 28031, , 28818; Wi Williams 3921, 3971).

subtextum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
241. 1936. TYPE: Peru, Junin, Puerto Yessup. El. 400 m.
10-12 July 1929. Killip & Smith 26265 (HE: USsolTseLELeS PT
at TLL: Killip & Smith 26278). 5

subtomentosum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 154,
fig. 132. 1950. TYPE: Colombia, Santander, Rio Surata
Valley, above Surata. El. 2000-2300 m. Jan. 1927. Killip &
Smith 6502 (BTs,US 13523173 ,102 LkL).

subtrinerve Trel., Contr. U.S. Nat. Herb. 26(2): 29. 1927.
TYPE: Panama, Taboga Island. Pittier 3618 (HT: US 678676;
PT at ILL: Piper 5410; Standley 26563) .-- > [=P
hirtellipetiolum var. subtrinerve e (Trel.) Yuncker (fide
Yuneker annotation, but comb. ined.; = var. harveyanum (fide
Yuncker, 1950)-- see next entry]

subtrinerve Trel. var. harveyanum Trel., Contr. U.S. Nat. Herb.
26(2): 30. 1927. TYPE: Panama, Prov. Panama, along the Tapia
River. June 1923. Maxon & Harvey 6684 (HT: US 1180427; IT:
ILL; PT at ILL: Maxon & Harvey 6734).-- [= P.

hirtellipetiolum var. harveyanum (Trel.) Yuneker, 1950)]

subvariabile Trel., Contr. U.S. Nat. Herb. 26(4): 145. 1929.
TYPE: Costa Rica, Guanacaste, near Tilaran. El. 500-650 m.
Jan. 1926. Standley & Valerio 44970 (HT: US 1307334; IT:
ILL; PT at ILL: Standley & Valerio 44294).-- [= P. grande
(fide Burger, 1971) ]

* sumideranum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
5, 9. 1926. TYPE: Cuba, Pinar del Rio, base of Sierra San
Francisco, between Hoyo Colorado and Francisco, Sumidero.
25 Nov. 1923. Ekman 18211 (HT, or LECTOTYPE designated
herein: ILL).

supernum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 1O2 ef ig.
151. 1950. TYPE: Ecuador, Los Rios, Cant6n Vinces, Hacienda
Santa Lucfa; £2. °50°m,s>;18.0ct.. 1994. Mexia. 6561, (HEs,US
1619065; IT: ILL).

Pon AST OE OG 's Vol. 58, No. 1

suratanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 210, fig.
172. 1950. TYPE: Colombia, Santander, Rio Surata Valley,
above Surata. El. 2000-2300 m. 5-6 Jan. 1927. Killip &
Smith 16660 (HT: US 1352380; IT: ILL).

surubresanum Trel., Contr. U.S. Nat. Herb. 26(4): 148. 1929.
TYPE: Costa Rica, San Mateo, along Rio Surubres. 14 Feb.
1909. Biolley 17353 (HT: US 861103; PT at ILL: Standley &
Valerio 46111).-- [= P. chrysostachyum (fide Burger, 1971) ]

svidaefolium Trel. in Rusby, Mem. N.Y. Bot. Gard. 7: 225. 1927.
TYPE: Bolivia, pampas near Lake Rogagua. El. 1000 ft.
7 Nov. 1921. Rusby 1687 (HT: NY; IT: ILL).-~- [= P.
subsilvulanum C.DC. (fide Yuncker, 1955) ]

tabanicidum Trel., Contr. U.S. Nat. Herb. 26(4): 162. 1929.
TYPE: Costa Rica, Guanacaste, Tilaran. El. 600 m.
Jan. 1926. Standley & Valerio 45676 (HT: US 1307383; PT at
ILL: Standley & Valerio 44241, 44896, 44925, 46593).-- [= P.

jacquemontianum (ef. Burger, Seren OTN

tabasaranum Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 25:
827. 1938. TYPE: Panama, Chiriquf, banks of lower Rfo
Tabasara. 12 Aug. 1937. Woodson, Allen & Seibert 440 (HT:
MO; IT: ILL).-- [= P. pinoganense (fide Yuncker, 1950)

tabernillanum Trel. var. anconense Trel. in Woodson & Schery,
Ann. Mo. Bot. Gard. 27: 298. 1940. TYPE: Panama, Canal Zone,
western slope of Anc6én Hill. 20 July 1938. Woodson, Allen &
Seibert 1327 (HT: ILL).-- [= P. taboganum (fide Yuncker, _
1950); = Tse pseudo-fuligineum (cf. Burger, 1971) ]

tachiranum Yuncker in Trel. & Yuncker, Piperaceae no. S. Amer.
1: 298, fig. 263. 1950. TYPE: Venezuela, Tachira, wooded
quebrada in Montana of Rio Tachira at base of Paramo de Tama,
between Betania and Tama, etc. El. 2430 m. 13 July 1944.
Steyermark 57172 (HT: F 1222766; IT: ILL).

taperanum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 126, fig. 110.
1966. TYPE: Brazil, Pernambuco, Tapera. 5 March 1936.
Pickel 4120 (HT: US 1593612; IT: ILL).-- {This species
upheld by Yuncker, 1973]

tapianum Trel., Contr. U.S. Nat. Herb. 26(2): 30. 1927. TYPE:
Panama, Prov. Panama, along the Tapia River, Juan Diaz
region. June 1923. Maxon & Harvey 6709 (HT: US 1180449; IT:

ss PT at ILL: Standley ley 26122). -- [= P. P. hirtellipetiolum
. tapianum (Trel.) Yuncker, 1950]

1985 Jones, Annotated catalogue of type specimens 9]

tardans Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 298.
1940. TYPE: Panama, Chiriquf, vicinity of Casita Alta,
Volean de Chiriqui. El. 1500-2000 m. 1938. Woodson, Allen
& Seibert 848 (HT: ILL).-- [= P. obliquum Ruiz & Pavon (fide
Burger, 1971)]

tardum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 242.
1936. TYPE: Peru, Junin, Colonia Perené. El. 680 m.
14-25 June 1929. Killip & Smith 25025 (HT: US; IT: ILL).

tembladorense Trel. ex L. Williams, Explor. Bot. Guayana
Venezolana, 177. 1942 (nomen subnudum). TYPE: Venezuela, El
Temblador, Bajo Caura. El. 100 m. 20 March 1939. Williams
11551 (HT: US; IT: ILL).-- [= P. taboganum (fide Trel. &
Yuneker, 1950; Yuncker, 1950) ]

tenebricosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
242. 1936. TYPE: Peru, Loreto, Fortaleza, lower Rfo
Huallaga. El. 155-210 m. Oct.-Nov. 1929. Williams 4201
(HT: F 623585; IT: ILL).

tenuiculispicum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 354.
1938. TYPE: Honduras, Atlantida, bank of Tela River above
Lancetilla Valley. El. 27 m. 10 July 1934. Yuncker 4509
(er: FEL):

tenuispicum C.DC., Bot. Gaz. 70: 170. 1920.-- Several syntypes,
probably to be lectotypified: Costa Rica, Las Vueltas.
Pittier 13187; Juan Vinas, Reventazon Valley. Cook & Doyle
294, 295 [Probable ST (from LE) at ILL: Tonduz 13187
(= Pittier 13187?)].-- [= P. aequale (cf. Burger, 1971) ]

tepezontesense Trel. in Standley, Field Mus. Publ. Bot. 11: 148.
1936. TYPE: El Salvador, San Juan Tepezontes. 1929.
Calder6én 2543 (HT: F 603259; IT: ILL; PT at ILL: Calderon
2541).

terronesense Trel. in Standley, Field Mus. Publ. Bot. 18: 365.
1937. TYPE: Costa Rica, Puntarenas, mouth of Rfo Terrones.
28 April 1930. Dodge 7749 (HT: GH; IT: ILL).-- [= P.
nudifolium (fide Burger, 1971) ]

tessaraespicum Trel. in Yuncker, Field Mus. Publ. Bot. 17: 354.
1938. TYPE: Honduras, Comayagua, near Siguatepeque.
El. 1050 m. 5 July 1936. Yuncker, Dawson & Youse 5662 (HT:
ILL).-- [= P. fraguanum (fide Standley & Steyerm., 1952);
= P. sancti-felicis (fide Burger, 1971) ]

thebaudianum Trel. in Feddes Repert. Spec. Nov. Regni Veg. oe
309, 312. 1927. TYPE: Haiti, Massif de La Selle, Riviere
Thébaud. El. 1000-1100 m. Ekman 1924 (HT: ILL).

92

DH) Val DLA AGT of Vol. 58, No. 1

tikalense Trel. in Standley, Field Mus. Publ. Bot. 17: 234.
1937. TYPE: Guatemala, Petén, in forest at Tikal.
12-15 April 1931. Bartlett 12595 (HT: ILL).-- [= P. cayoense
(fide Standley & Steyerm., 1952) ]

tilaranum Trel., Contr. U.S. Nat. Herb. 26(4): 131. 1929. TYPE:
Costa Rica, Guanacaste, near Tilaran. El. 500-650 m.
Jan. 1926. Standley & Valerio 46596 (HT: US 1307464; PT at
ILL: Standley & Valerio 44182, 44247).-- [= P. amalago (fide
Burger, 1971) ]

timothianum A.C. Smith, Sargentia 1: 10. 1942. TYPE: Fiji, Mba
Prov., Viti Levu, Nandarivatu. 22 Nov. 1940. Degener &
Ordonez 13570 (HT: ?; IT: ILL).-- [= Macropiper timothianum
(A.C. Smith) A.C. Smith (fide Smith, 1981)]

tingens Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357): 244.
1936. TYPE: Peru, Loreto, Yurimaguas. El. 180 m. 24 Oct.
1.931.4>Mexia-6072.(HT s-UCs EPs) FEEL).

tomas-albertoi Trel. & Yuncker, Piperaceae no. S. Amer. 1: 113,
fig. 86. 1950. TYPE: Colombia, Antioquia, El Prado. 7 July
1943. Daniel 3022 [HT: US 1834580; IT: ILL; PT at ILL (2
sheets): Toma&s Alberto 1577]. .

tomasi Trel. ex Tomas Alberto, Colegio de San José, Medellin,
Colombia Publ. No. 53-70, 18. 1942 (nomen nudum); ex Trel. &
Yuneker, Piperaceae no. S. Amer. 1: 252. 1950, pro syn.
TYPE: Colombia, Antioquia, Yarumal. March 1941. Tomas
Alberto 1533 (HT: ILL).-- [= P. eriopodon (Miq.) C.DC. in DC.
(fide Trel. & Yuncker, 1950) ]

tonduzii C.DC. var. semiherbaceum Trel., Contr. U.S. Nat. Herb.
26(4): 143. 1929. TYPE: Costa Rica, Guanacaste, Quebrada
Serena near Tilardn. El. 700 m. 27 Jan. 1926. Standley &
Valerio 46261 (HT: US 1307443; PT at ILL: Standley & Valerio
44751).-- [= P. tonduzii (fide Burger, 1971)]

tonoroanum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 328, fig.
295. 1950. TYPE: Venezuela, Bolfvar, Sitio Tonoro, Rfo
Tonoro, etc. 26 Dec. 1943. Cardona 454 (HT: US 1878526;
Fe TL):

torbecanum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23:
306, 310. 1927. TYPE: Haiti, La Mare Proux, Torbec.
Elz 1350: a. ‘EkmanvS279- (HT: 1h) ;

1985 Jones, Annotated catalogue of type specimens 93

tortuosipilum Trel., Contr. U.S. Nat. Herb. 26(4): 148. 1929.
TYPE: Costa Rica, Cartago, El Muneco. El. 1400-1500 m.
March 1926. Standley & Valerio 51026 (HT: US 1307618; PT
at ILL: Standley & Valerio 50973, 51017).-- [= P. biauritum
(cf. Burger, 1971)]

tortuosivenosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 244. 1936. TYPE: Peru, Loreto, mouth of Rio Santiago.
El, 200im... 2-Dees 1931. sHexia 6199 .0HT: UC: IT: ILL (2
sheets) J.

tractifolium Trel., Contr. U.S. Nat. Herb. 26(4): 166. 1929.
TYPE: Costa Rica, Heredia, above San Isidro, Cerro de las
Caricias. El. 2200 m. 11 March 1926. Standley & Valerio
52052 (HT: US 1307663; PT at ILL: Standley & Valerio 49122,
51945).-- [= P. tenuimucronatum C.DC. (cf. Burger, 1971)]

transpontinum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 244. 1936. TYPE: Peru, Junin, Perené Bridge,
Paucartambo Valley. El. 700 m. 19 June 1929. Killip &
Smith 25352 (HT: US; IT: ILL; PT at ILL: Killip & Smith
P5AT3)s

transpontinum var. ecordulatum Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 245. 1936. TYPE: Peru, Junin, Colonia
Perené. El. 680 m. 14-25 June 1929. Killip & Smith 25055
CHE nUS s ete FLL).

trichostylopse Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 245. 1936. TYPE: Peru, Huanuco, Pozuzo. El. 600 m.
20-22 June 1923. Macbride 4646 (HT: F 535721; IT: ILL; PT
at ILL: Macbride 4664, 5336).

trichostylopse var. latum Trel. in Macbr., Field Mus. Publ. Bot.
13(No. 357): 245. 1936. TYPE: Peru, Hudnuco, Cushi.
El. 1500 m. 19-23 June 1923. Macbride 4849 (HT: F 535889;
5 Cam Be

trigonocarpum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 24:
363. 1928. TYPE: Santo Domingo, Las Lagunas, Azua.
El. 750 m. Ekman 6376 (HT: ILL).

triguetrobaccum Trel., Trab. Mus. Nac. Ci. Nat. Ser. Bot. 33:
51. 1936 (without a Latin diagnosis). TYPE: Colombia,
Tolima, Ibagué. El. 1300 m. 10 May 1932. Cuatrecasas 2418
(HT: ILL).-- [= P. cumanense (fide Trel. & Yuncker, 1950) J

triguetrobaccum var. tenuifolium Trel., Trab. Mus. Nac. Ci. Nat.
Ser. Bot. 33: 51. 1936 (without a Latin diagnosis). TYPE:
Colombia, Tolima, Ibagué. 8 May 1935. Cuatrecasas 2417 (HT:
ILL).-- [= P. cumanense (fide Trel. & Yuncker, 1950) ]

94 Path ¥ TrOvk OiGed 2A Vol. 58, No. 1

triquetrofructum Trel. in Standley, Field Mus. Publ. Bot. 18:
366. 1937. TYPE: Costa Rica, Cartago, Rf{o Pejivalle Gorge.
El. 600-650 m. 27 Sep» 1929. Dodge & Thomas 4431 (HT: GH;
IT: ILL).-- (= P. dilatatum (cf. Burger, 1971)]

tristigmum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
246. 1936. TYPE: Peru, Loreto, Caballo-Cocha on the Amazon
River. Aug. 1929. Williams 2029 (HT: F 613314; IT: ILL).

triumphale Trel. in Standley, Field Mus. Publ. Bot. 10: 161.
1931. Type locality: Honduras, "in thickets along the beach
near Tela, Lancetilla Valley." No type cited; perhaps
lectotypification needed (authentic specimen from type
locality at ILL: Standley 53035).

tuberculatum Jacq. var. cangrejalense Trel. in Yuncker, Field
Mus. Publ. Bot. 9: 280. 1940. TYPE: Honduras, Atlantida, in
thicket on lower slopes of Mt. Cangrejal. 5 July 1938.
Yuncker, Koepper & Wagner 8204 (HT: ILL).

tuberculatum var. scandens Trel. & Yuncker, Piperaceae no. S.
Amer. 1: 367. 1950. TYPE: Colombia, Boyacd, San Pablo.
El. 150 m. 21 Feb. 1939. Haught 2620 (HT: US 1707193; IT:
ILL).

tumidicondyli Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357) 2247. °1936.. TYPE: Peru, Loreto, Iquitos,« Rancho
Indzane, Els 110«m., °25 Jan;..1932. Mexia‘6431 G¢HT: UC; IT:
Thi):

tumidipedunculum Trel. in Yuncker, Field Mus. Publ. Bot. 9: 281.
1940. TYPE: Honduras, Yoro, Las Flores, near Coyoles.
El. 300 m. 30 June 1938. Yuncker, Koepper & Wagner 8151
CHE = #EGL Dy

ulceratum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
247. 1936. TYPE: Peru, San Martin, San Roque. El. 1350-
1500 m. Jan.-Feb. 1930. Williams 6962 (HT: F 622978; IT:
EEF

ulceratum var. angustifolium Trel. in Macbr., Field Mus. Publ.
Bot. 13(No. 357): 248. 1936. TYPE: Peru, San Martin, San
Roque. El. 1350-1500 m. Jan.-Feb. 1930. Williams 6982 (HT:
B.6296125 2%¢ ILL).

uncatum Trel. in Standley, J. Wash. Acad. Sci. 13: 367. 1923.
TYPE: El Salvador, San Salvador, vicinity of Tonacatepeque.
30 Dec. 1921. Standley 19435 (HI: Thi) <

1985 Jones, Annotated catalogue of type specimens 95

unguiculatum Ruiz & Pavon, Fl. Peruv. Chil. 1: 34, pl. 57,
fig. b. 1798. TYPE: Peru, Pozuzo and Chinchao. Ruiz & Pavon
S.n. (HT: MA; fragments of type at ILL).-- [This species
discussed by Howard, 1973]

unguiculiferum Trel., Contr. U.S. Nat. Herb. 26(4): 149. 1929.
TYPE: Costa Rica, Cartago, Pejivalle. El. 900 m. 7-8 Feb.
1926. Standley & Valerio 47003 (HT: US 1307485; IT: ILL).--
[= P. xanthostachyum (fide Burger, 1971) ]

vanderveldeanum Trel. in Feddes Repert. Spec. Nov. Regni Veg.
23: 308, 312. 1927. TYPE: Haiti, La Hotte, Camp Perrin,
Morne Vandervelde. El. 900 m. 2 Dec. 1925. Ekman 5224 (HT:
ILL).

variabile C.DC. var. obtentum Trel. in Yuncker, Field Mus. Publ.
Bot. 9: 281. 1940. TYPE: Honduras, Atlantida, Mt. Cangrejal.
El. 150 m. Yuncker, Koepper & Wagner 8495 (HT: ILL).

varium Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 299.
1940. TYPE: Panama, Chiriquf{, trail from Paso Ancho to Monte
Lirio. El. 1500-2000 m. 16 Jan. 1939. Allen 1491 (HT:
ILL).-- [= P. maxonii C.DC. var. varium (Trel.) Yuncker (fide
Yuneker, 1950 and Burger, 1971) ]

velutininervosum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 249. 1936. TYPE: Peru, Junin, E of Quimiri Bridge, La
Merced. El. 800-1300 m. 1-3 June 1929. Killip & Smith
2UOOG “CAT -OUS > (FTF STLL)

* venturii Trel. in Moldenke, Lilloa 10: 303. 1944. TYPE:
Argentina, Tucuman, Tafi. El. 750 m. 8 Aug. 1920. Venturi
866 [HT, or LECTOTYPE designated herein (specimen marked
"Type" by Trelease): ILL; IT or ILT: ILL)].-- [= P. hieronymi
C.DC. (fide Yuncker, 1955)]

venulosum Trel., Contr. U.S. Nat. Herb. 26(4): 132. 1929. TYPE:
Costa Rica, near San Ram6n. Brenes 14192 (HT: US 716432; PT
at ILL: Standley & Valerio 45702).-- [= P. papantlense (cf.
Burger, 1971) ]

veredianum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
250. 1936. TYPE: Peru, Loreto, Caballo-Cocha on the Amazon
River. Aug. 1929. Williams 2106 (HT: F 608810; IT: ILL).

verruculaecaule Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 250. 1936. TYPE: Peru, Ayacucho, Rio Apurfmac Valley,
near Kimpitiriki. El. 400 m. 10, 11 May 1929. Killip &
Smith 23025 (HT: US; IT: ILL).

96 Pith YoT 00k OnpGct ek Vol. 58, No. 1

verruculaefolium Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 250. 1936. TYPE: Peru, Junfn, La Merced. El. 700 m.
May-June 1929. Killip & Smith 24089 (HT: US; IT: ILL).

verruculaepetiolum Trel. in Standley, Field Mus. Publ. Bot. 18:
368. 1937. TYPE: Panama, Bocas del Toro, Farm Six,
Changuinola Valley. 3 March 1924. Dunlap 494 (HT: F; IT:
ILL).-- [= P. terrabanum (fide Burger, 1971) ]

verruculigerum Trel., Contr. U.S. Nat. Herb. 26(4): 165. 1929.
TYPE: Costa Rica, Guanacaste, Quebrada Serena, Tilaran.
El, 700 m. 27 Jan. 1926. Standley & Valerio 46239 (HT: US
1307440; IT: ILL; PT at ILL: Standley & Valerio 44487, 45043,
45679, 45711, 46188, 46233).-- [= P. chry sostachyum yum (fide
Burger, 1971) ]

vexans Trel., J. Wash. Acad. Sci. 19: 336. 1929. TYPE:
Honduras, Atlantida, near Tela, at sea level. 18 Jan. 1928.
Standley 54742 [HT: F 583715; IT: ILL (2 sheets)].-- [= P.
jacquemontianum (fide Burger, 1971)]

viae-marginis Trel. in Yuncker, Field Mus. Publ. Bot. 17: 355.
1938. TYPE: Honduras, Comayagua, near Siguatepeque.
El. 1050 m. 23 July 1936. Yuncker, Dawson & Youse 6044 (HT:
ILL).-- [= P. atlantidanum (fide Standley & Steyerm., » 1952);

= P. pseudo-fuligineum ligineum (cf. Burger, 1971)]

vigosanum Yuncker, Bol. Inst. Bot. (Sao Paulo) 3: 74, fig. 64.
1966. TYPE: Brazil, Minas Gerais, Vigosa, Fazenda do
Deserto. El. 690 m. 3 Nov. 1930. Mexia 5242 (HT: US
1618044; IT: ILL; PT at ILL: Mexia 5401-a).-- [This species
upheld by Yuncker 1972]

villosispicum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 251. 1936. TYPE: Peru, Junfn, Pichis Trail, Santa
Rosa. El. 625-900 m. 6, 7 July 1929. Killip & Smith 26178
(HT: US; IT: ILL; PT at ILL: Killip & Smith 24533).

virgulticola Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
251. 1936. TYPE: Peru, Loreto, Iquitos, Fundo Indiano.
E4211140 mer .25) Jan: °1932.5 Mexia 6430. (HT:, UC; :IT:) DLL):

* viridibaccum Trel., J. Arnold Arb. 9: 150. 1928. TYPE: New
Guinea, Papua, Movabula. 10 May 1926. Brass 1370 (LECTOTYPE
designated herein: A; ILT: ILL).-- [= P. lessertianum C.DC.
(fide Chew annotation) ]

viridilimbum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
251. 1936. TYPE: Peru, Ayacucho, Rfo Apurimac Valley,
Kimpitiriki. El. 400 m. 10, 11 May 1929. Killip & Smith
22990 (HT: US; IT: ILL).

1985 Jones, Annotated catalogue of type specimens 97

viridispicum Trel., Contr. U.S. Nat. Herb. 26(4): 138. 1929.
TYPE: Costa Rica, Limon, Montechristo below Cairo. El. a5 wi.
Feb. 1926. Standley & Valerio 48536 (HT: US 1307543; PT at
ILL: Standley & Valerio 48599).-- [= P. garagaranum C.DC.
(fide Burger, 1971) ]

viscaianum Trel. & Yuncker, Piperaceae no. S. Amer. s2°t39,. fig.
115. 1950. TYPE: Colombia, Santander, Viscaina Creek, 31 km
S of El Centro. El. ca 100 m. 31 Oct. 1936. Haught 2053
CHES CUS :1F42 3192" IFS RILE)?

whiteae Trel. in Woodson & Schery, Ann. Mo. Bot. Gard. 27: 299.
1940. TYPE: Panama, Chiriqu{, valley of the upper Rfo
Chiriquf Viejo, near El Volcdn. 18 July 1938. White 177
CHT" TLL).-= [= P. maxonii (fide Yuncker, 1950 and Burger,
1971) ]

wilsonii Trel. in Standley, Field Mus. Publ. Bot. 10: 161. 1931.
Type locality: Honduras, Atlantida, vicinity of Tela, at sea
level. No type indicated; perhaps lectotypification needed
(authentic specimen from type locality at ILL: Standley
56874).

yalochanum Trel. in Standley, Field Mus. Publ. Bot. 17: 234.
1937. TYPE: Guatemala, Petén, Yaloch Trail, Dos Arroyos.
2 May 1931. Bartlett 12849 (HT: ILL).-- [= P. cayoense (fide
Standley & Steyerm., 1952) ]

yanaconasense Trel. & Yuncker, Piperaceae no. S. Amer. 1: 41,
fig. 32. 1950. TYPE: Colombia, El Valle, El Silencio,
Yanaconas, etc. El. 1900-2200 m. 28 Feb. 1939. Killip &
Garcfa 33796 (HT: US 1770684; IT: ILL).

yananoanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No. 357):
252. 1936. TYPE: Peru, Huanuco, Yanano. El. 1800 m.
13-16 May 1923. Macbride 3796 (HT: F 534858; IT: ILL).

yapeanum Trel. in Woodson & Seibert, Ann. Mo. Bot. Gard. 25:
827. 1938. TYPE: Panama, Darién, near the mouth of R{fo Yape.
El. 20m. July 1937. Allen 351 (HT: ILL).-- [= P.
villiramulum (fide Yuncker, 1950) ]

yaranum Trel. in Feddes Repert. Spec. Nov. Regni Veg. 23: 4, 8.
1926. TYPE: Cuba, Oriente, Rio Yara to R{fo Palmamocha,
Sierra Maestra. 1922. Ekman 14332 (HT: ?; IT: ILL; PT at
ILL: Ekman 6874).

98 PSY 90. bLioGs Fis Vol. 58, No. 1

yaxnanum Trel. in Standley, Field Mus. Publ. Bot. 17: 234. 1937.
TYPE: Guatemala, Petén, Yaxha-Remate Road. 16 June 1933.
Lundell 3826 (HT: ILL).-- [= P. martensianum (fide Standley &
Steyerm., 1952) ]

yoroanum Trel., J. Wash. Acad. Sci. 19: 337. 1929. TYPE:
Honduras, Yoro, Quebrada Seca. El. 30 m. Dec. 1927.
Standley 53889 (HT: F 583866; IT: ILL).

yousei Trel. in Yuncker, Field Mus. Publ. Bot. 17: 355. 1938.
TYPE: Honduras, Comayagua, near El Achote, above
Siguatepeque. El. 1350 m. 21 July 1936. Yuncker, Dawson &
Youse 5999 (HT: ILL; PT at ILL: Yuncker, Dawson & Youse
6070).

yunckeri Trel. in Yuneker, Field Mus. Publ. Bot. 17: 355. 1938.
TYPE: Honduras, Subalpine bog near El Achote, above
Siguatepeque. El. 1350 m. 5 Aug. 1936. Yuncker, Dawson &
Youse 6331 (HT: ILL).

yunckeri var. latum Trel. in Yuncker, Field Mus. Publ. Bot. 17:
355. 1938. TYPE: Honduras, Comayagua, near El Achote, above
Siguatepeque. El. 1350 m. 5 Aug. 1936. Yuncker, Dawson &
Youse 6333 (HT: ILL).

yurimaguasanum Trel. in Macbr., Field Mus. Publ. Bot. 13(No.
357): 253. 1936. TYPE: Peru, ‘Leretopheuttos. yEbactoo a.
3-11 Aug. 1929. - Killip & Smith 27274 (HT: US; ITs ILL; PT at

mr

yzabalanum C.DC. var. pubinerve Trel. in Standley, Field Mus.
Publ. Bot. 10: 161. 1931. Type locality: Honduras,
Atlantida, in thickets about Tela and Lancetilla. No type
indicated; perhaps lectotypification needed (authentic
Specimens from vicinity of type locality at ILL: Standley
55776, 56872).

zingiberinum Trel. in Standley, Field Mus. Publ. Bot. 18: 370.
1937. ~TYPE: .Costaihica, iSanta-Glara Hilis.:° El. 1500.-m.
16 June 1928. Stork 2594 (HT: ILL).-- [= P. verruculosum
C.DC. (fide Burger, 1971)]

zonulatispicum Trel., Contr. U.S. Nat. Herb. 26(4): 164. 1929.
TYPE: Costa Rica, Cartago, El Muneco, Rio Navarro.
El. 1500 m. 7 March 1926. Standley & Valerio 51116 (HT: US
1307636; PT at ILL: Standley & Valerio 51335).-- [= P.
carpinteranum (fide Burger, 1971) ]

1985 Jones, Annotated catalogue of type specimens 99

Pothomorphe Miq.

baileyorum Trel. in Woodson & Schery, Ann. Mot. Bot. Gard. 27:
306. 1940. TYPE: Panama, Canal Zone, Barro Colorado Island.
27 June 1931. L.H. & E.Z. Bailey 414 (HT: ILL).-- {= P P.
peltata (L.) Miq. (fide Yuncker, 1950 and 1957); = Lepianthes
peltata (L.) Raf. (cf. Howard, 1973) ]

baileyorum var. paucispica Trel. in Woodson & Schery, Ann. Mot.
Bot. Gard. 27: 306. 1940. TYPE: Panama, Canal Zone,
Salamanca Hydrographic Station, Rfo Pequeni. El. 80 m.
28 July 1938. Woodson, Allen & Seibert 1573 (HT: ILL).--
[= P. peltata (fide Yuncker, 1950 and 1957); = L. Peltata
(cf. Howard, 1973)]

dussii Trel. ex Stenlé, Fl. Deser. Antill. Fr. 2(1):61, pl. 1.
1940; Bull. Agric. Martinique 9(2): 137, tab. 1. 1940. TYPE:
Martinique, Parnasse, Marin. Duss 1340 (HT: US; PT at ILL:
Stehlé & Stenlé 2318, 3191, 3200, 00, 3239).-- [= L. peltata (cf.
Howard, 1973)]_

tecumensis Trel. var. grandis Trel. in Woodson & Schery, Ann.
Mo. Bot. Gard. 27: 307. 1940. TYPE: Panama, vicinity of
Arraijan. El. ca 15 m. 21 July 1938. Woodson, Allen &
Seibert 1398 (HT: ILL).-- [= P. peltata (fide Yuncker,
1950 and 1957); = L. peltata (ef. Howard, 1973)]

Sarcorhachis Trel., emend. Steyerm.

* incurva (Sieber ex Schultes) Trel. var. stehlei Trel. ex Stehlé,
Bull. Soc. Bot. France 83: 627. 1936 (nomen nudum); Fl
Deser. Antill. Fr.°2(1): 67, pl. 2.1940; Bull. Agric.
Martinique 9(2): 143, tab. 2. 1940. TYPE: Guadeloupe.
El. 650 m. 16 Feb. 1936. Stenhlé & Stehle 380 (Syntype =
LECTOTYPE designated herein: NY; ILT: ILL).-- [= S. incurva
(fide Steyerm., 1971 and Howard, 1973)]

incurva var. treleasii Stenlé, Fl. Deser. Antill. Fr. 2(1): 67.
1940; Bull. Agric. Martinique 9(2): 143. 1940. TYPE:
Guadeloupe. El. 650 m. 27 June 1937. Stehlé & Stehlé 1677
(HT: NY; fragments of type at ILL).-- [= S. incurva (fide
Steyerm., 1971 and Howard, 1973)]

Schillera Kunth

karwinskiana Kunth, Linnaea 13: 700. 1839 [originally named
Piper karwinskianum by Kunth (loc. cit.), i.e., that name
published pro syn. ]. TYPE: Mexico, near Mexico City.
Karwinski [HT, or ST to be lectotypified(?): "in herb.
Lucaeano"; fragments of the original collection (obtained

100 Poe T Oboes fa Yol. 58, No. 1

from LE) at ILL: Karwinski 827, 827b].-- [= Piper
karwinskianum (Kunth) C.DC. in DC.-- the species is known
only from the type locality (fide Standley, 1920) ]

ulmifolia Kunth, Linnaea 13: 698. 1839. TYPE: Martinique.
Sieber 7 [HT: "... herb. Lucaeani"; IT (obtained from W):
ILL; PT at ILL: Trinidad, Sieber 18].-- [= Piper dilatatum
(fide Howard, 1973)]

Trianaeopiper Trel.

confertinodum Trel. & Yuncker, Piperaceae no. S. Amer. 1: 423,
fig. 381. 1950. TYPE: Colombia, Chocdé, S of R{o Condoto,
between Quebrada Guarapo and Mandinga. El. 120-180 m.

April 1939. Killip 35678 (HT: US 1772363; PT at ILL: Killip
35317).

garciae Trel. & Yuncker var. decurtispicum Trel. & Yuncker,
Piperaceae no. S. Amer. 1: 430. 1950. TYPE: Colombia, El
Valle, Dagua Valley, Santa Rosa. El. 200-300 m. 22 Sep.
1922... Killip, £1531. (NTs: Gis, 13s, ALi),

mexiae Trel. & Yuncker, Piperaceae no. S. Amer. 1: 432, fig.
392. 1950. TYPE: Ecuador, Los Rios, Hacienda "El Oasis",
Cant6n Vinces. El. 60 m. 5 Nov. 1934. Mexia 6651 (HT: US
1619074; IT: ILL).

ACKNOWLEDGMENTS

I have received help, via correspondence, from Wm. Burger (F),
P. F. Stevens (A), P. K. Holmgren and A. F. Cholewa (NY), and
P. L. Morrelli (US). Special thanks and appreciation for
assistance in the preparation of this catalogue goes to the typist,
Joyce Roberts, and to Laurel McKee for checking the references and
proofing the manuscript. Warren Lamboy advised me on the format of
the introduction.

1985 Jones, Annotated catalogue of type specimens 101
REFERENCES

Adams, C. D. and Collaborators. 1972. Flowering plants of
Jamaica. University of the West Indies. Mona, Jamaica.

Burger, Wm. 1971. Family No. 41, Piperaceae. In: Flora
Costaricensis. Wm. Burger, Editor. Fieldiana Bot. 35: 5-227.

Chew, W.-L. 1972. The genus Piper (Piperaceae) in New Guinea,
Solomon Islands, and Australia. 1. J. Arnold Arb. 53: 1-25.

Croat, T. B. 1978. Flora of Barro Colorado Island. Stanford
Univ. Press, Stanford, CA.

Holmgren, P. K., W. Keuken and E. K. Schofield. 1981. Index
Herbariorum. Ed. 7. Bohn, Scheltema & Holkema. Utrecht.

Howard, R. A. 1973. Notes on the Piperaceae of the Lesser
Antilles. J. Arnold Arb. 54: 377-411.

Smith, A. C. 1975. The genus Macropiper (Piperaceae).
J.. Linn. Soc. Bot. 71: 1-38, pl. 1-11.

- 1981. Flora Vitiensis Nova. Vol. 2. Pacific
Tropical Bot. Garden. Lawai, Kauai, Hawaii.

Standley, P. C. 1920, 1926. Piperaceae. In: Trees and shrubs of
Mexico. Contr. U. S. Nat. Herb. 23(1): 145-156; 23(5): 1647.

- and J. A. Steyermark. 1952. Piperaceae. In: Flora of
Guatemala. Fieldiana Bot. 24(3): 228-337.

Steyermark, J. A. 1971. Notes on the genus Sarcorhachis Trel.
(Piperaceae). Pittieria 3: 29-38.

Trelease, Wm. 1926. Piperaceae cubenses. Feddes Repert. Spec.
Nov. Regni Veg. 23: 1-31.

- 1927. The Piperaceae of Panama. Contr. U. S. Nat. Herb.
26(2)> 3—50.

- 1929. The Piperaceae of Costa Rica. Contr. U. S. Nat.
Herb. 26(4): 115-226.

- 1935. The pedicellate peppers of South America. Proc.
Amer. Philos. Soc. 75(8): 691-716.

- 1936. Piperaceae. In: Macbride, J. F. Flora of
Peru. Part 2. Field Mus. Publ. Bot. 13 (No. 357): 1-253.

102 Pee T Oe OG" FR Vol. 58, No. 1

- 1938. Piperaceae. In: Woodson, R. E. and R. J. Seibert.
Contributions toward a Flora of Panama. II. Ann. Mo. Bot.
Gard. 253 825-827.

- 1940. Piperaceae. In: Woodson, R. E. and R. W. Schery.
Contributions toward a Flora of Panama. IV. Ann. Mo. Bot.
Gard. 27: 287-307.

. and T. G. Yuncker. 1950. The Piperaceae of northern South
America. 2 vols. The University of Illinois Press. Urbana.

Yuncker, T. G. 1938. A contribution to the Flora of Honduras.
Field Mus. Publ. Bot. 17 (No. 405): 285-407, pl. 1-18.

. 1940. Flora of the Aguan Valley and the coastal regions
near La Ceiba, Honduras. Field Mus. Publ. Bot. 9 (No. 466):
243-346.

- 1950. Piperaceae. In: Woodson, R. E. and R. W.
Schery. Flora of Panama. Part IV, fasc. 1. Ann. Mo. Bot.
Gard > 37(1): 1-120.

- 1953. The Piperaceae of Trinidad and Tobago. Lilloa
26: 239-278.

- [1953] 1955. The Piperaceae of Argentina, Bolivia and
Chiles’ °*Lillea 272°97=303.

- 1955a. The status of the genus Anderssoniopiper
Trelease. Bull. Torrey Bot. Club 82: 355-356.

- 1957. Piperaceae. Pp. 218-290. In: Lanjouw, J.
Editor, Flora of Suriname. Vol. 1, part 2. Royal Tropical
Institute, Amsterdam. Mededeling No. 30.

- 1960. The Piperaceae of Jamaica. Bull. Inst. Jamaica.
Science Ser. 11: 1-56.

- 1962. Nomenclatural notes on Piperaceae. Brittonia 14:
188-190.

- 1972. The Piperaceae of Brazil— I: Piper -- groups I,
II, III, IV. Hoehnea 2: 19-366, figs. 1-167.

- 1973. The Piperaceae of Brazil— II: Piper -- group V;

Ottonia; Pothomorphe; Sarcorhachis. Hoehnea 3: 29-284, figs.
168-292.

1985

Notice to librarians

BACK ISSUES AVAILABLE

103

As of this writing the following back volumes and separate issues
of PHYTOLOGIA are still available from the Publishers, 590 Hemlock
N.W., Corvallis, Oregon 97330:

Avenue

Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume
Volume

UR ERR OF pre Eatin 205s. aig Pate EA ey $ 3.00
See WN ie wore oie 2 arene eee te tee 3.00
ee SUNN PERS ie eae: See a ee te a eee a eee 3.00
Bek, (OS 2 eh Abc re wie le ate oi cie ain areemarere 6.00
Pe Waa Pe nics ain ie ope oro ea aie ne eee 3.00
are ec dia cic ak eeu ene eeear ste 3.00
TIMERS, (Roadie cielo. a aia aw be, Oats aes 3.00
Diet Bass dine vio eh ok ban Rae oe 3.00
ERIN oR Seren aie oa Sell te rye, ete eer eaten 3.00
Sa eS whe es ane, Ger, et oe aan 9.00
SO Nese. bs Soo. ON OL csc. oo ces woe 12.00
MIPEOMDIPLe. 2). 25.26 so a oue enone oe 16.00
ae OMPPELe ys. sc cuca cp eee oe oe eee ae 16.00
cog ho | ak tae a ae Mas SE Tia i Re a ce 16.00
PEMMRD PEERS. @ o5.a sitisin's w helium owes bon 16.00
Pons «aves Re and BS ke Le 13.00
SLMS a Sh De ANG Pee es ot 5 oe 10.00
i eS hs at SANE on eS ee ce 10.00
DMEM BERENS so tta tis ~ ce in ars wae were 16.00
Pe OED RP ECE. a oe ala enn eke ee ee 16.00
Se ORME VE LEGS s-2ic a os 3 ss Stee on ones onl 16.00
Wh CUM PORES ocak ocak barca a wee ede 16,00
SAC ORC E DG o:5 sown ae-< eiee tein oeeeue 16.00
ay \CPMMD ROLE. 00 wic-in sae eke a oe ee 16.00
BA COMULEEE So a's ooo aie sapere. oe ee etetaa ele 16.00
5 EGU GOs do ss sc aa oon noone oe 16.00
Bir SE OMB OLE so ace ico 0 on ciayarding wa mecenne 16.00
SF PEO OES 6c co o.dcahs coos enh a teen 16.00

Material addressed to any USA bookdealer or other

business establishment, add $2.00; this applies also to any

USA, $21
USA, $21
USA, $21
USA, $2]

USA, $18.
00
.00
.00
.00
.00
.00
.00
00
.00
.00
.00
.00

USA, $15
USA, $15
USA, $21
USA, $21
USA, $21
USA, $21
USA, $21
USA, $21
USA, $21
USA, $2]
USA, $2]
USA, $21

.00
.00
-00
.00

00

foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign
foreign

commercial or

addressed to an individual to whom Library Rate postal rates do not

apply

All items are subject to prior sale

104 PHY 7,01 24.22 ¥ei.. 58, Mout

As of this writing the following issues of PHYTOLOGIA MEMOIRS are
still available:

] -- "A Check List of the Vascular Plants of Nicaragua" by F. C.
Seymour, 314 pp. 1980. Available
from Mrs. Frank C. Seymour, 264 Hixville Road, North
Dartmouth, Mass. 02747

I] -- "A Sixth Summary of the Venrbenaceae, Avicenniaceae, Stilbaceae,
ChLoanthaceae, Symphonemaceae, Nyctanthaceae, and Eniocaul-
aceae of the world as to valid taxa, geographic distribution
and synonymy" by H. N. Moldenke, 628 pp. 1980. Available
from Harold N. Moldenke, 590 Hemlock Avenue N.W., Corvallis,
Oregon 97330. USA non-profit institutions $27.00; other
USA addresses $29.00; foreign $31.50

II] -- "Antillean Studies 1. Flora of Hispaniola: Part 1, Celastnales,
Rhamnales, Malvales, Thymekeales, Violales" by A. H. Liogier,
218 pp., 1981. Available from Dr. Alain Liogier, Botanic
Garden, University of Puerto Rico, Central Administration,
G. P. 0. Box 4984-G, San Juan, Puerto Rico 00936

IV -- "Studies in the Begoniaceae" by A. I. Baranov, 88 pp., 198!.
Available from A. Baranov, P. 0. Box 131, Cambridge, Mass.
02140.

V -- "The Flora of New England", second edition, by F. C. Seymour,

illustrations by Andrea Robbins, first impression, 611 pp.,
1982. Available from Harold N. Moldenke, 590 Hemlock Avenue
N.W., Corvallis, Oregon 97330. USA non-profit institutions
$23.00; other USA addresses $23.50; foreign $24.50; a second
impression (1985) is available from Mrs. Frank C. Seymour,
264 Hixville Road, North Dartmouth, Massachusetts 02747

VI -- "A Preliminary Verified List of Plant Collectors in Mexico" by
I. W. Knobloch, 179 pp., 1983. Available from Harold N. Mol-
denke, 590 Hemlock Avenue N.W., Corvallis, Oregon 97330. USA
nonprofit institutions $17.00; other USA commercial insti-
tutions and businesses $17.50; USA individuals $15.00; foreign
$18.50

VII -- "An International Census of the Coniferae. I" by J. Silba, 79 pp.,
1934. Available from John Silba, 198 West Hoffman Avenue,
Lindenhurst, N. Y. 11757.

Also still available from Harold N. Moldenke, 590 Hemlock Avenue N.W.,
Corvallis, Oregon 97330:

"A Fifth Summary of the Verbenaceae, Avicenniaceae, Stilbaceae,
Symphonreraceae, Nyctanthaceae, and Eniocaulaceae of the
world as to valid taxa, geographic distribution, and
synonymy", 2 volumes, 974 pp., 1971. $25.00 plus postage
and handling. Contains much information, keys, bibliography,
etc. not contained in the Sixth Summary

All items subject to prior sale

35185 00288 276/

Inasmuch as we do no editing, papers accepted for publication must be
submitted in exactly the form that the author wants to have them published. They
will then be photographed and printed by photo-offset in exactly the form as
submitted except that we will add page numbers and running-heads.

Typescripts should be prepared single-spaced on clean white heavy bond
smooth and opaque paper. Elite type is probably the most space-economical.
Typescript text must not exceed a rectangle 5% inches wide (horizontal) by 8%
inches high (vertical), not including the running-head and page number.

The title of the paper should be typed in all uppercase (capital) letters with 2
blank lines above the title and one beneath; then the name of the author in
ordinary upper-and lower-case letters, along with his address (if so desired);
followed by 2 blank lines; then the first line of text. It is usually best to leave a
blank line between paragraphs.

All scientific plant and anima! names and group names should be typed either in
italic type (if available) or underscored. Any corrections in the text made by the
author must be complete and neat as they will be photographed as they are.

The finished typescript as submitted by the author will be reduced from the 8%
x 5% inch size as submitted to 6 2/8 x 4 inches by the printer. It is therefore
advisable to place a centimeter or millimeter scale on all text figures and plates
included.

Use a new heavily inked black typewriter ribbon and be sure to clean the type
on the typewriter after each several pages of typing.

Cost of publication at present is $11.00 US per page, with no subsequent rebates,
but this rate may vary depending on inflation and costs, so it is best to inquire as to
current rates. The page charges are due with the typescript and no paper will be
published before payment is received in full. Each author will receive gratis a
proportionate share of the printed copies remaining after paid subscriptions are
filled, but if separates (reprints or offprints) are desired, these will be charged
extra in accord with the current rate for offprints provided by the printer. The cost
of all such separates ordered must also be paid for in advance at the time the
typescript is sent. No orders for separates will be accepted later, nor can additions
or corrections be accepted.

Authors are asked to indicate in light pencil on the reverse side of each page of
their typescript the page number so that no mistakes in sequence occur.

Each number consists of not less than 32 pages. All manuscript accepted will be
published in the next issue, so that the size of numbers may vary greatly. A
volume will contain 512 pages. This plan insures prompt publication of all
accepted manuscript.

Illustrations will be published according to the desires of the authors. No extra
charge is made for line drawings, such as are ordinarily reproduced in zinc, or for
diagrams, tables, or charts, provided they conform to certain limitations of size
and proportion. An extra charge will be made for halftones, depending on their
size, as fixed by the engraver.

- Articles dealing with research in all lines of botany and plant ecology, in any
reasonable length, biographical sketches, and critical reviews and summanies of
literature will be considered for publication.