“* PHYTOLOGIA An international journal to expedite botanical and phytoecological publication Vol. 63 August 1987 No. 4 CONTENTS BEETLE, A.A., Noteworthy grasses from Mexico XIII .......... 209 ADAMS, C.D., Cyperaceae trinitensis novae .................. 298 SCHAAK, C.G., A new Arizona Purshia (Rosaceae) ........... 301 ZARATE P, S., Taxonomic identity of Leucana leucophylla (Lam.) de Wit, with a new combination .................+.-. 304 TURNER, B.L., A new species of Sabazia (Asteraceae- Heliantheae) from Durango, Mexico ...............0000000+ 307 WOLDE, E.R., Two new forms of Helianthus agrestis .......... 310 MOLDENBE Fee OO FEDIEWS. 2 eS ae bio ce cele ce sine ws 311 LIBRARY — JAUG 31 1987 Published by Harold N. Moldenke and Alma I Moldenke YK 590 Hemlock Avenue : Corvallis, Oregon o73SRMGANICAL GARDEN U.S.A. Price of this number $3.00; for this volume $16.00 in advance or $17.00 after close of the volume; $5.00 extra to all foreign addresses and domestic dealers; 512 pages constitute a complete volume; claims for numbers lost in the mail must be made immediately after receipt of the next following number for free replacement; back volume prices apply if payment is received after a volume is closed. NOTEWORTHY GRASSES FROM MEXICO XIII Alan A. Beetle, Range Management Section, Universitv of Wyoming, University Station, P. 0. Box 3354, Laramie, Wyoming, 82071. In 1977 (cf Phytologia 37: 317-407) an annotated list of the grasses reported for Mexico was published. Since that time there has been a concerted effort by the mexican Comisi6én Téc- nico Consultiva de Coeficientes de Agostadero of the Direcci6n General de Normatividad Pecuaria, Secretaria de Agricultura y Recursos Hidraéulicos, to gather information on the taxonomy, distribution and forage value of the grasses of Mexico. Personnel associated with this work have included: Ing. Victor Jaramillo V. M. S. Victor Jaramillo L. Biol. Elizabeth Manrique F. Biol. Alejandro Miranda S. M.S. Aurora Chimal H. Biol. Patricia Guerrero S. M.S. Irama Nunez T. ‘Ing. Rafael Guzman M. Biols Lilia Rico FR. The list presented here is considerably more accurate thar. the first but much remains to be done. For srevious zoners cee Fartologia 27:1974, 2°:1974; 30:1975; 35:1977, 38:1978, 47:1981, 49:33-34; 52:11 - 17. 1981; 54:1983, 209 210 POH XT On 0-G TA Vol. 63, No. 4 ACROCERAS Stapf Paniceae 5 - 6 species, both hemispheres, subtropical. 1. A. zizanioides (HBK) Dandy Pantropical, probably native. Southern México: Veracruz, Oaxaca, Chiapas, Tabasco, Campeche. AEGILOPS L. Hordeae 20 - 30 species, circum-Mediterranean. 2. A. cylindrica Host. Introduced. Northern México: Reported only for Chihuahua where cultivated. AEGOPOGON Humb. & Bonpl. Chlorideae .3 species, New World. 3. A. cenchroides Humb. & Bonpl. var. cenchroides México to Colombia and Venezuela, native. Common from Chihuahua to Chiapas. 3a.A. cenchroides var breviglumis (Scribn.) Beetle Same distribution as the species but less common, native. 4. A. tenellus (DC) Trin. var. tenellus . Arizona to Guatemala, native. Common from Baja Norte and Chihuahua south to Chiapas. 4a.A. tenellus var. abortivus (Fourn.) Beetle. Northern range of the species but less common, native. Sonora south to México, Tlaxcala and Veracruz. AGROPYRON Gaertn. Hordeae 50 - 60 species, both hemispheres, temperate, 5. A. arizonicum Scribn. & Smith Northern Mexico and adjacent United States, native. Mountains: Sonora, Chihuahua, Coahuila, Nuevo Le6én, Zacatecas, San Luis Potosi, México and Morelos. 6. Agropyron parishii Scribn. & Smith var. laeve Scribn.& Smith Southern California and adjacent México, native. Rare, northern Baja California Norte. 7. A. trachycaulum (Link) Malte Boreal North America south in the mountains to México, native. Mountains: Baja Norte, Chihuahua, Coahuila, Nuevo Le6n, Tamaulipas. 8. Agropyron vaillantianum (Wulf. & Schreb.) Trautv. Europe, North America and South America, native. Northern México and the central highlands. AGROSTIS .. Agrostideae 150 species, New World and Old World, both hemispheres, temperate. 9. A. alba L. Introduced. Mesic sites: Chihuahua, Coahuila & Nuevo Le6n south to Oaxaca. 10.A. borealis Hartm. Circumboreal, south in the mountains to México, native. Mountains: Puebla (near Mirador Restaurant). 1987 Beetle, Mexican grasses 7A aL 11.A. bourgaei Fourn. Either an introduced variation of A. alba or an endemic. Mountains: Hidalgo, México, and Tlaxcala and Guanajuato. 12.A. diegoensis Vasey Western United States south to Baja Norte, México Coast range: Baja Norte. 13.A. elliottiana Schultes Maryland to Illinois south to Georgia and Texas; México, native. Waste Places: reported only for Yucat&n. (Where introduced) 14.A. exarata Trin. Alaska and western North America south to México. Moist places: Baja California; Chihuahua, Coahuila south to México. 15.A. ghiesbreghtii Fourn. México, endemic. Central Mountains: Hidalgo, México, Morelos, Guerrero, Oaxaca and Pico de Orizaba. 16.A. hiemalis (Walt.) B.S.P. var. laxiflora (Gray) Beetle Newfoundland to Alaska and south in the mountains to central México. Mountains : Baja California, Chihuahua, Coahuila, south to Chiapas. 17.A. liebmannii (Fourn.) Hitchc. México, endemic. Type loc: Chimantla, Veracruz; also Durango (type loc. for. A. durangensis Mez); Queretaro, México y Oaxaca. Lea: microphylla Steud. California and México, native. 19.A. palustris Huds. Introduced. Marshes: Baja Norte and Chihuahua,Coahuila. 20.A. perennans (Walt.) Tuckerman Quebec to Minesota, and south to Florida and Guatemala. Mountains: Durango, San Luis Potosi, Jalisco, Hidalgo, México, south to Chiapas. 21.A. rosei Scribn. & Merr. México, endemic. Mountains: Durango, Zacatecas, and México. 22.A. schaffneri Fourn. México, endemic. Mountains: Queretaro, Michoacan, México, Tlaxcala, Puebla, Oaxaca & Chiapas. 23.A. semiverticillata (Forsk.) Christ. Introduced. Wet places, common throughout México except only the Yucatdn Peninsula. 24.A. subrepens (Hitchc.) Hitchc. México and Venezuela, native. Wet places: Chihuahua. 25.A. tacubayensis Fourn. México, endemic. Mountains: Michoacan, State of México and Oaxaca. 26.Agrostis tandilensis (Kunth) Parodi Southern California and adjacent México, native. Rare in northern Baja California Norte. 212 PH YT Op LvorGe Iva Vol. 63, No. 4 27. A. thyrsigera Mez México, endemic. Central Montains: 28. A. tolucensis HBK México: south to Chile. Central Montains: 29. A. vinosa Swallen México and Guatemala, native. Central Montains, 30. A. virescens HBK México, endemic. Central Montains. AIRA L. Aveneae About 10 species 9 Old World annuals 31. Aira caryophyllea L. Introduced. Reported from Oaxaca and Baja California Norte. ALLOLEPIS Soderstrom and Decker Eragrosteae One species, North America. 32. A. texana (Vasey) Soderstrom and Decker Texas and México, native. Salt flats: Chihuahua, Coahuila, Tame ilipas and Durango. ALOPECURUS L. Agrostideae 20 - 30 species, Temperate regions. 33. A. geniculatus L. Northern hemisphere, temperate, native. Wet places: at its southern extreme reported from Chihuahua. 34. Alopecurus howellii Vasey From Oregon to the northwest corner of México, native. Rare in Baja California Norte. 35. Alopecurus saccatus Vasey Western Canada south to northern México, native. Rare in Baja California Norte. ANDROPOGON L. Andropogoneae About 100 species, temperate to subtropical, worldwide. 36. A. (Anatherum) bicornis L. México to Argentina, native. Pine savanna or brush: Nayarit, Jalisco, Tamaulipas, Hidal- go, Puebla, Oaxaca, Veracruz,Chiapas, Tabasco, Campeche and Yucatan. 37. A. (Anatherum) bourgaei Hack. México, endemic. Stream draws; Veracruz (Type loc. Rio Blanco), Oaxaca, and Chiapas. 38. A. (Schizachyrium) brevifolius (Sw.) Nees. Tropical and subtropical regions of world, described from "Jamaica", native. Pine savanna: :Sonora south to Chiapas. 39. A. (Schizachyrium) cirratus Hack. México and adjacent United States, native. Pine savannas: northern border states south to Chiapas. 1987 Beetle, Mexican grasses 213 40. A. (Schizachyrium) condensatus HBK As var. elongatum Roberty: México to Paraguay. As subsp. elongatus subvar. exserens Hack. in Mart. Fl. Bras. 2:297. 1883: Brazil and México. Note: by some authors combined with A. microstachyus Desv. 41. A. (Anatherum) elliottii Chapm. i Eastern United States, Cuba, southern México to British Honduras, native. Pine savannas: Chiapas; San Luis Potosi & Veracruz. 42. A. (Schizachyrium) gaumeri (Nash) Hitchc. México, endemic. Brush: Chiapas, Campeche, Yucatan. 43. A. gerardi Vitman United States to Honduras, native. Pine forests: Coahuila, Durango, Sinaloa south to Chiapas. 44. A. (Anatherum) glomeratus (Walt.) B.S.P. Southeastern United States, México, and the West Indies to Panama, native. Common in open areas throughout México. 45. A. hallii Hack. North Dakota and Montana south to northern México, native. Sandy soils: Chihuahua. 46. A. (Schizachyrium) hirtiflorus (Nees) Kunth var. hirtifloru Southern United States and West Indies to Bolivia and Uruguay, native. Pine forests and brush: common throughout México. 46a.A. (Schizachyrium) hirtiflorus var. feensis (Fourn.)Hack. Same distribution as the species. 47. Andropogon hirtifolius Presl native, endemic. Common in Central México, Jalisco and Guanajuato to Chiapas 48. A. leucostachyus HBK Southern México and the West Indies to Argentina, native. .Pine forests: Guerrero, Veracruz, Chiapas, Tabasco, Oaxaca, and Campeche. 49. A. (Anatherum) liebmannii Hack- México, endemic. Pine.forests: Nayarit, Jalisco, Michoac&n, Hidalgo, Puebla, México, Veracruz and Oaxaca. 50. A. maderensis Swallen México, endemic. Canyons: Coahuila. 51. A. (Schizachyrium) malacostachyus Presl México to Costa Rica, native. Rocky hills: Jalisco, Guerrero, Puebla, Oaxaca, Chiapas & Yucatan. 52. A. mexicanus Hitchc. México, endemic. Open woods: Nayarit, Jalisco (Tipe Loc.) and Hidalgo south to Chiapas. 53. A. (Schizachyrium) microstachyus Desv. México and west Indies to Argentina, native. Pine forests or brush: Sonora, Sinaloa, Jalisco, Tamaulipas San Luis Potosi, Hidalgo, Morelos, Guerrero, Puebla, Verazr Oaxaca, Tabasco and Chianas. 214 F HOY FT OFLFOVGL IVA Vol. 63, No. 4 54. A. (Schizachyrium) muelleri (Nash) Hitchc. (a tetraploid variation of A. scoparius) México, endemic. Coastal: Veracruz. Ds A. (Schizachyrium) myosurus Presl México, endemic. Pine woods: Chihuahua, Durango, Sinaloa, Jalisco, Milchoa- can, Guerrero, México, and Oaxaca. 56. A. (Anatherum) pringlei Scribn. & Merr. México, endemic. Central mountains: Michoacan, México, Puebla, Veracruz, Oaxaca. 57. A. salzmanni (Trin.) Nash México to Paraguay (Type from Brazil) native. Brush: Tamaulipas, Veracruz, Oaxaca. 58. Andropogon sanguineus (Retz.)Merr. Florida, México, West Indies to Argentina, native. Brush, worldwide in the subtropics. 59. Andropogon schlumbergeri Fourn. Native, endemic, Durango & San Luis Potosi south to Oaxaca. 60. Andropogon scoparius Michx. var. frequens Hubb. Eastern U.S.A., common Texas, rare in México. 60a.A. scoparius var. littoralis (Nash) Hitchc. Northeastern coast of E.U.A., Great lakes area and Texas, eastern coastal México, native. 60b.A. scoparius var. neomexicanus (Nash) Hitchc. Native, western Texas to Arizona and northern México. 61. A. selloanus (Hack.) Hack. México and West Indies to Argentina, native. Pine woods: Oaxaca, Veracruz, Campeche and Chiapas. 62. A. (Schizachyrium) semiglabrus (Nash) Beetle México, endemic. Chihuahua (Type Loc.) "near Colonia Garcia", Nuevo Le6én, Jalisco, México and Chiapas. 63. A. semitectus Swallen México and Guatemala, native. Brush: Baja California Sur & Sonora south to Guerrero. 64. A. spadiceus Swallen México Coahuila (Type Loc.) 65. A. (Schizachyrium) tener (Nees) Kunth Southern United States and West Indies south to Argentina, native. Brush: Sonora and Tamaulipas south to Chiapas. 66. A. te.narius Michx. De’ aware to Kentucky and Kansas, south to Florida and México, native. Sandy soil: Coahuila. 67. A. virginicus L. Southern United States and West Indies south to Panama,native. Pine woods: Coahuila, Nuevo Le6én, Puebla, Veracruz, Oaxaca and Chiapas. 1987 Beetle, Mexican grasses 215 68. Andropogon wrightii Hack. Native, rocky hills and mesas, southern New México and northern Mexico south to southern Veracruz. 69. A. yucatanus Swallen. Brush: Yucatan and Campeche. ANTHEPHORA Schreb Chlorideae 4 - 5 species in Africa, one in tropical America. 70. A. hermaphodita (L.) Kuntze Florida pee eanee West Indies and Mexico south to Peru and Brazil, native. Brush: Baja California Norte, south to the Yucatdn Peninsula. ANTHOXANTHUM L. Aveneae 3 - 4 species, European. 71. A. odoratum L. Introduced. Occasional in cultivated areas: Hidalgo and Mexico. ARISTIDA L. Aristideae 200 species throughout the subtropics of the world. 72. A. adscensionis L. var. adscensionis Widespread in subtropics of both New and old World,native? Brush: common throughout northern Mexico. 72a.A. adscensionis var. abortiva Beetle California and northern Mexico. native. Dry washes: Baja Sur, Coahuila, Chihuahua and Durango 72b.A. adscensionis var. coarctata (HBK) Kuntze West Indies and Mexico south to Venezuela, native. Brush: Jalisco, Tamaulipas, Campeche, Yucatan 72c.A. adscensionis var. decolorata (Fourn.) Beetle Mexico, endemic. Brush: Baja Sur, Sonora, Guerrero, Oaxaca, Chiapas and Yucatan . 72d.A. adscensionis var. interrupta (Cav.) Beetle Mexico, endemic. Brush: Coahuila, Nuevo Le6n, Tamaulipas, San Luis Potosi, Veracruz and Oaxaca. 72e.A. adscensionis var. modesta Hack. California and Arizona south to Argentina, native. Dry washes: northern borderstates south to Mexico. 72£.A. adscensionis var. nigrescens (Presl) Beetle Mexico, endemic. Dry washes: Baja Sur and Sonora south to Chiapas. 73. A. arizonica Vasey Colorado and Texas south to Central Mexico (acc.Henrard), native. Dry banks: northern border states (except Tamaulipas) south to the central highlands, and Oaxaca. 74. A. barbata Fourn. Texas to Arizona and central Mexico, native. Dry banks: northern border states south to Districto Federal. 216 Typ 76. We 78. Ve 80. 81. 82. 83. 84. 85... 86. 87. 88. POO Y 170) Lr Orel A Vol. 63, No. 4 A. californica Thurb. Southern California and southwestern Arizona to Northwestern Mexico, native. Dry washes: Baja Norte, Baja Sur and Sonora. A. capillacea Lam. Mexico to Bolivia and Brazil, native. Pine forests: Sinaloa, Nayarit and Mexico. A. curvifolia Fourn. Mexico, endemic. Pine savanna: Coahuila, Nuevo Le6én, Durango, Zacatecas, San Luis Potosi and Oaxaca. A. divaricata H. & B. (including A. palmeri Vasey) Kansas to southern California south to Mexico. Dry banks: northern border states south to Oaxaca. A. fendleriana Steud. North Dakota and Montana, south to Nevada, southern California and Mexico, native. Dry banks: Coahuila, Nuevo Leén and San Luis Potosi. A. floridana (Chapman) Vasey Florida and Yucatan Peninsula, native. Brush: Campeche, Yucatan and Quintana Roo. A. fournieriana Hiceher Mexico, endemic. Brush: Veracruz. A. gentilis Henr. Mexico, endemic. Pine woods: Sonora, Durango, Zacatecas, Aguascalientes, Jalisco, Guanajuato, Mexico, Puebla, Oaxaca and Chiapas. A. glabrata (Vasey) Hitchc. Arizona and Mexico, native. Dry banks: Sonora, Baja Norte and Baja Sur. A. glauca (Nees) Walp. Southern California, Nevada and Utah south to Texas and Mexico native. Dry banks: All northern border states and south to Puebla. Aristida gypsophila Beetle Native, endemic. Gypsum soils of Coahuila. A. hamulosa Henr. Southwestern United States to Guatemala, native. Oak forests: Baja Norte, Sonora, Chihuahua, south to Chiapas. A. hintoni Hitchc.- Mexico, endemic. Pine savanna: Tamaulipas, Nayarit, Jalisco, Guanajuato, Guerrero, Michoacan, Mexico, Guerrero and Oaxaca. A. hitchcockiana Henr. Mexico, endemic. Pine savanna: Nayarit, Zacatecas, Guanajuato, Queretaro, Jalisco, Puebla, Oaxaca (Type loc.). 1987 Beetle, Mexican grasses 217 89. A. jacquiniana Tausch Mexico to Ecuador, native. Pine savanna: Jalisco and Guanajuato. 90. A. jaliscana Guzman & Jaramillo Mexico, native endemic. 91 .A. jorulensis Kunth Mexico to Panama, native. Dry banks: Sonora, Sinaloa, Nayarit, Jalisco, Colima, Guerrero, Mexico, Oaxaca, Veracruz and Chiapas. 92. A. lagascae Henr. Mexican, endemic. State of Mexico (Type loc.). 93. A. laxa Cav. var. laxa México, endemic. Dry banks and dunes: Sonora, Chihuahua, Sinaloa, Nayarit, Jalisco, San Luis Potosi, México, Morelos, Puebla and Oaxaca. 93a.A. laxa var. longiramea (Presl) Henr. Mexico, endemic. Dry banks: Jalisco to Oaxaca. 93b.A. laxa var. Karwinskiana (Trin & Rupr.) Henr. Mexico, endemic. Dry banks: Central Mexico. 94. A. liebmanni Fourn. Mexico, endemic. Dry banks: Veracruz (Type loc.) Oaxaca and Chiapas. 95. A. longespica Por. New Hampshire to Michigan south to Florida, Texas, Mexico, native. Sandy soil: Tamaulipas, Coahuila, Chihuahua. 96. A. longiseta Steud. var. longiseta Western U.S. to northern Mexico, native. Plains: Baja Norte, Sonora, Chihuahua, Coahuila and ; Nuevo Le6én and Tamaulipas. 96a.A. longiseta var. robusta Merr. Western United States and New Mexico to northern Mexico, native. Plains: Same rangeas the species but more common northward - 97. A. mexicana Scribn. Mexico, endemic.- reported by Henrard (1929) for Guatemala but not confirmed by Swallen (1955). Dry banks: Mexico, Distr. Fed. and Puebla. 98. A. orizabensis Fourn. Mexico to Panama, native. Brush: Sinaloa, Durango, Jalisco, Guanajuato, Michoac4&n, Mexico, Guerrero, Tlaxcala, Puebla, Veracruz, Oaxaca, Chiapas. 99. A. pansa Woot. & Standl. var. pansa. Texas and Arizona south to central Mexico, native. Plains: Sonora, Chihuahua, Coahuila, Nuevo Le6én, south to Puebla, Oaxaca. 99a.A. pansa var. dissita (Johnston) Beetle. Same distribution as the species. 100.A. parishii Hitchc. Nevada to California and northern Mexico, native. Dry washes: Baja Norte. 218 PH Y TP OP LO Geta Vol. 63), Now 4 101. A. peninsularis Hitchc. Mexico, endemic. Dunes: Baja California Norte and Sonora. 102. A. purpurascens Poir. Massachusetts to Wisconsin and Kansas south to British Honduras, native. Sandy soil: Coahuila. 103. A. purpurea Nutt. Arkansas and Kansas to central Mexico, native. Plains: Coahuila, Nuevo Le6n, Tamaulipas, Durango, San Luis Potosi, Hidalgo and Puebla. 104. A. purpusiana Hitchc. Mexico, endemic. Dry wash: Baja California Sur. 105. A. roemeriana Scheele Texas and New Mexico to central Mexico, native. Plains: northern border States (except. Baja California). Durango, Zacatecas, San Luis Potosi, Puebla and Yucatan. 106. A. schiedeana Trin. & Rupr. Mexico and Guatemala, native. Pine savannas througout Mexico (excpt. Yucatan Peninsula) 107. A. scribneriana Hitchc. Mexico, endemic. Brush: Durango, Jalisco, Guanajuato, Michoacan, Guerrero. 108. Aristida tenuifolia Hitchc. Mexico, endemic. Rocky point, Sulphur Bay. Isla Clarion, Islas Revillagigedo. 109. A. ternipes Cav. var. ternipes Southwestern United States, West Indies to Colombia,native. Brush: common throughout Mexico. 109a.A. ternipes Cav. var. minor (Vasey) Hitchc. Same distribution as the species. 110. A. vaginata Hitchc. Mexico, endemic. Revillagigedo Islands (Soccoro Isld.) type loc. 111. A. wrightii Nash. Southern California and Colorado, Texas, south to central Mexico, native. Plains: Chihuahua, Ccahuila, Nuevo Leon, south to Oaxaca. 112. Arrhenatherum elatius (L.) Presl var. bulbosum (Willd.) Spenner f. striatum (Hubb.) L.B. Smith Introduced, cultivated as an ornamental. ARTHRAXON Beauv. Andropogoneae About 20 species in the Old World tropics. 113. A. hispidus (Thunb.) Makino Introduced. Brush: Chiapas. 114. Arthropogon villosus Nees Mexico and Brazil. Introduced or native. edge of pine woods, Oaxaca. 115. Arthrostylidium venezuelae (Steud.) McClure. Mexico south to Venezuela. Reported only for Chiapas. 1987 Beetle, Mexican grasses 219 ARUNDINELLA Raddi Arundinelleae 20 species, pantropical. 116. A. berteroniana (Schult.)Hitchc. & Chase Mexico to Brazil, native. Brush: Sinaloa, Nuevo Leon, Tamaulipas, San Luis Potosi, Jalisco, Colima, Michoacan, Hidalgo, Mexico, Morelos, Puebla, Guerrero, Veracruz, Oaxaca, Chiapas and Tabasco. 117. A. confinis (Schult.)Hitchc. & Chase Mexico and West Indies south to Panama, native. Dry banks: Sonora, Jalisco, Michoacan, Mexico, Guerrero, Veracruz, Oaxaca, Chiapas and Tabasco. 118. A. deppeana Nees Mexico and West Indies south to Brazil, native. Pine savanna: Nuevo Leon, Sinaloa, Nayarit, Jalisco, Colima, Michoacan,Guerrero, Hidalgo, Mexico, Puebla, Veracruz, Oaxaca, Chiapas, Tabasco. 119. A. palmeri Vasey Mexico, endemic. Dry banks: Sonora, Nayarit, Jalisco, Colima,Michoacan, Mexico, Guerrero, Veracruz. ARUNDO L. Arundinae 5 - 6 species, Old World, subtropics. 120. “An Gdonax i. Intruduced. Cultivated and adventive throughout Mexico. 120a.Arundo donax L. f. versicolor (Mill.) Beetle Introduced, cultivated as an ornamental. 121. Aulonemia laxa (Maekawa) Native, central Mexico. AVENA L. Aveneae About..i0species, Old World, temperate. 122. A. barbata Brot. Introduced. Coastal hills: Baja Norte. 123. A. fatua L. Introduced. Common weed throughout Mexico. 124. A. sativa L. Introduced. Cultivated and an escape, scattered localities from the northern states south to central Mexico. AXONOPUS Beauv. Paniceae about 80 species, tropical and subtropical, New World. 125. A. affinis Chase Southeastern United States, West Indies to Argentina, Native. Wet places: Jalisco, Queretaro, Hidalgo, Puebla, Veracruz and Chiapas and Campeche. 126. A. arseni Swallen Mexico, endemic. Wet places: Colima, Michoacan and Distr. Federal. 220 127. 128. Pele 130. 131. L333). 134. 135 136. 137. IL ite) 133 9%. Pin Wt Or Lora lA Vol. 63, No. 4 A. ater Chase Mexico, endemic. Wet places: Veracruz. A. centralis Chase Mexico, endemic. Wet places: Nayarit, Jalisco and Colima;also Chiapas. A. chrysites (Steud.) Kuhlm. Mexico and Central America, native. Wet places: "Mex. Galeotti" acc. to Black, 1963,Chiapas. A. compressus (Sw.) Beauv. Pantropical (native). Wet places: often cultivated, common throughout Mexico A. deludens Chase Mexico, endemic. Wet places: Jalisco and Sinaloa A. elongatus Swallen Mexico and Guatemala, native. Wet places: Chiapas. A. mexicanus Black Mexico, endemic. Wet places: Sinaloa. A. multipes Swallen Mexico, endemic. Wet places: Veracruz. A. poiophyllus Chase Mexico to Honduras, native. . Wet places: Tamaulipas, Veracruz and Chiapas. A. purpusii (Mez) Chase Mexico to Argentina, native. Wet places: Veracruz, Oaxaca, Chiapas, Tabasco, Puebla. A. reederi Black Mexico, endemic. Wet places: Chiapas. A. rosei (Scribn. & Merr.) Chase Mexico, endemic. Wet places: Nayarit. A. scoparius (Flugge) Kuhlm. Mexico to Peru (acc. to Black, 1963), native. Wet places, "Mex. Galeotti 227" (acc. to Black, 1963). BAMBUSA Schreb. Bambuseae 140. 141. 142). 100 or more in the Old World tropics and in the New World tropics as Section Guadua. B. (Guadua) aculeata (Rupr.)Hitchc. Mexico south to Panama, native. Tropical forest margin, San Luis Potosi, Veracruz, Campeche, Guerrero, Puebla, and Oaxaca. B. (Guadua) amplexifolia (Presl) R.T.S. Mexico to Colombia and Venezuela. Tropical forest margin, Tamaulipas and San Luis Potosi south to Chiapas and Tabasco. Bambusa latifolia HBK Native, Mexico, Trinidad, and northern South America south to Brazil. Reported from San Luis Potosi and Veracruz. 1987 Beetle, Mexican grasses 221 143. B. (Guadua) longifolia (Fourn.) McClure Mexico, endemic. Thickets, Sonora and Chihuahua south to Chiapas and Campeche. 144. Bambusa paniculata (Munro) Hack. Native, Mexico and Central America south to Brazil. Reported from Jalisco, Nayarit, and Veracruz. 145. B. vulgaris Schrad. Introduced. Cultivated, scattered localities throughout Mexico. 145a.Bambusa vulgaris Schad. f. vittatae (A. & C. Rive.)Beetle B. vulgaris var. vittata A. & C. Riv. Bull. Soc. Acclim. rE 53640). 1878) Introduced, cultivated as an ornamental. BLEPHARIDACHNE Hack Eragrosteae 2 species, North America. 146. B. bigelovii (S.Wats.)Hack. Texas and northern Mexico, native. Chihuahuan desert: Coahuila. BLEPHARONEURON Nash Eragrosteae One species, North America. 147. B. tricholepis (Torr.)Nash Colorado and Utah south to central Mexico, native. Pine woods: northern border states south to Puebla. BOTHRIOCHLOA Andropogoneae About 30 species, tropic and subtropics of the world. 148 B. alta (Hitchc.)Henr. Texas and New Mexico, south to Bolivia and Argentina,native Dry banks: Durango, Zacatecas, San Luis Potosi, Jalisco, Guanajuato, Queretaro, Oaxaca, Chiapas. 149. B. barbinodis (Lag.)Herter var. barbinodis California, Colorado and Texas south to central Mexico; “ Argentina and Uruguay, native. 149a.Bothriochloa barbinodis var. palmeri (Hack.) Gould. Native, endemic. Reported from Duranao, Zacatecas, Jalisco, Guanajuato, and Michoacan. 149b.B. barbinodis var. perforata (Trin.)Gould Same distribution as the sp-cies. 150. Bothriochloa bladhi (Retz.)Blake Introduced. Reported from Yucatan. 151. B. hyrida (Gould) Gould Texas and Mexico, native. Dry banks: Coahuila, Nuevo Leon,& Tamaulipas, San Luis Potosi, and Oaxaca. 152. B. ischaemum (L.) Keng var. songarica (Rupr.) Cel.&Henr. Introduced. Cultivated & escaped: Nuevo Leon, Tamaulipas, Michoacan. 153. B. pertusa (L.) Camus Introduced. Dry banks: Tamaulipas, San Luis Potosi, Campeche and Yucatan. 154. B. reevesii (Gould) Gould Mexico, endemic. Dry banks: Coahuila 222 POH Y (TOF O%Geal vA Vol. 63, No. 4 155. B. saccharoides (Sw.)Rydb. var. saccharoides. Southwestern United States and West Indies to Argentina, native. Dry banks: common throughout Mexico exceptfor the Yucatan Peninsula. 155a.B. saccharoides var. laguroides (DC)Beetle Same distribution as the species, native. 155b.B. saccharoides var. longipaniculata (Gould) Gould Texas to Panama, native. Dry banks: Nuevo Leon. 156. B. springfieldii (Gould) Parodi United States and Mexico, native. Dry banks: Nayarit. BOUTELOUA Lag. Chlorideae LSiWic TSS 59%. 160. HEL. Species about 40 in the Americas. B. alamosana Vasey Mexico, endemic. Rocky banks: Sonora (Type loc.)also Zacatecas, Jalisco Oaxaca & Chiapas. Bouteloua americana (L.)Scribn. Native, Carribean and Mexico south to Brazil. Reported only for the Yucatan Peninsula. B. annua Swallen Mexico, endemic. Dry banks: Baja Sur (Type loc.) B. aristidoides (HB)Griseb. (including B. aristidoides var. arizonica Jones). Texas to southern California, Mexico and South America, native. Dry soils: northern border states south to Oaxaca. B. barbata Lag. var. barbata Colorado and Utah south to Mexico; Argentina, native. Dry soils: northern border states south to Oaxaca. 161a.Bouteloua barbata var. major (Vasey) Beetle (B. rothrockii (Cervant) Swallen) Southern California and Arizona to northern Mexico. Native: Mesas;Sonora, Chihuahua, Coahuila, Nuevo Leon, Sinaloa, Durango and Baja Sur, Jalisco. 161b.Bouteloua barbata var. sonorae (Griffiths) Gould 162. 163)< 164. 165) Mexico,endemic. Dry washes: Sonora, Sinaloa and Baja Sur. B. breviseta Vasey U.S. and Mexico, native. Dry washes: Chihuahua, Coahuila, Nuevo Leon, Durango, Zacatecas, San Luis Potosi, and Veracruz. B. chasei Swallen Mexico, endemic. Gypsum: Coahuila, Nuevo Leon, Zacatecas and San Luis Potosi. B. chondrosioides (HBK) Benth. Texas and Arizona south to Honduras, native. Rocky slopes: Sonora and Chihuahua northern border states south to Chiapas. B. curtipendula (Michx.)Torr. var. curtipendula Canada to central Mexico, native. Grassland: northern border states south to central Mexico. 1987 Beetle, Mexican grasses 223 165a. B. curtipendula var. caespitosa Gould and Kapadia U.S. and Mexico; Venezuela to Argentina, native. Dry banks: northern border states south to Chiapas. 165b. B. curtipendula var. tenuis Gould and Kapadia Mexico, endemic. Dry banks: Chihuahua south to Chiapas. 166. B. distans Swallen Mexico, endemic. Dry banks: Oaxaca (Type loc.); Nuevo Leon 167. B. disticha (Kunth) Benth. Mexico and West Indies to Peru and Argentina, native. Dry banks: Nayarit, Jalisco, Michoacan, and Guerrero, Tamaulipas, Mexico, Oaxaca, Campeche, & Yucatan 168. B. elata J.& C. Reeder Mexico, endemic. Rocky cliffs: Nayarit, Jalisco, Colima and Chiapas. 169. B. eludens Griffiths U.S. and Mexico, native. Dry banks: Sonora, 170. B. eriopoda (Torr.)Torr. U.S. and northern Mexico, native. Dry slopes: Sonora, Chihuahua, Coahuila, Nuevo Leon and Durango. 171. B. eriostachya (Swallen) Reeder Mexico, endemic. Dry banks: Coahuila 172. B. gracilis (HBK)Lag.var. gracilis U.S. and Mexico; native; South America (introduced). Grasslands: northern border states to Oaxaca. 172a. Bouteloua gracilis var. major (Vasey) Beetle Native, endemic. Reported from Aguascalientes, Jalisco, Guanajuato and Hidalgo. 172b. Bouteloua gracilis var. polystachya (Nees) Beetle Native, endemic. Central Mexico. 172c. Bouteloua gracilis var. stricta (Vasey)Hitchc. Native, endemic. Reported from San Luis Potosi, Durango and Chihuahua. 173. B. hirsuta Lag. var. hirsuta U.S. and Mexico, native. Thickets: throughout Mexico except for the Yucatan Peninsula. 173a. Bouteloua hirsuta Lag. var. palmeri Vasey Mexico and Guatemala, native. Limestone balds: Sonora, Chihuahua and Baja Sur south to Chiapas. 174. B.johnstonii Swallen Mexico, endemic. Gypsum: Coahuila. 175. B. Karwinskii (Fourn.)Griffiths Mexico, endemic. Gypsum: Coahuila, Nuevo Leon, Tamaulipas, Zacatecas, San Luis Potosi and Tamaulipas. 224 PH YT OSL10*G; TA Vol. 63, No. 4 176. B. media (Fourn.) Gould and Kapadia Mexico, endemic. Dry slopes: Nayarit, San Luis Potosi, Colima, Michoacan, Veracruz, Mexico, Morelos, Guerrero, Oaxaca and Chiapas. 177. B. parryi (Fourn.)Griffiths var. parryi New Mexico and Arizona to México, native. Rocky slopes: northern border states south to Guanajuato. 177a.Bouteloua parryi var. gentryi (Gould)Gould Mexico, endemic. Dry banks: Durango, Sinaloa (Type loc.). 178. B. pedicellata Swallen Mexico, endemic. Rocky slopes: Tlaxcala and Puebla (Type loc.). 179. B. purpurea Gould and Kapadia Mexico, endemic. Heavy black soils: central mountains. 180. B. radicosa (Fourn.)Griffiths Southern New Mexico and southern California to Mexico, native. Dry slopes: northern border states south to Oaxaca. 181. Bouteloua ramosa Vasey Limestone flats and slopes, Texas and northern Mexico. Reported from Chihuahua, Durango, Coahuila, Zacatecas, and Nuevo Leon. 182. B. reflexa Swallen Mexico, endemic. Dry washes: Sonora, Sinaloa, Baja California Sur and Nayarit. 183. B. repens (HBK)Scribn. & Merr. (including B. filiformis (Fourn.)Griffiths. Texas to Arizona, West Indies to Venezuela and Colombia, native. Dry slopes: common throughout Mexico. 184. B. rigideseta (Steud.)Hitchc. Oklahoma and Texas and northern Mexico, native. Grassland: Sonora, Coahuila, Nuevo Leon and Tamaulipas. 185. B. scorpioides Lag. Mexico, endemic. Dry flats: Chihuahua and Nuevo Leon south to Puebla, Oaxaca. 186. B. simplex Lag. Southwestern United States to Argentina, native. Dry flats: northern border states south to Veracruz and Oaxaca. 187: B. triaena’ (fran.)'serzbn. Mexico and Guatemala, native. Tbhickets: Coahuila and Sinaloa south to Yucatan, Peninsula. 18S Bp Ere fidan Thunb Texas to California and northern Mexico, native. Grasslands: northern border states south to Guanajuato. 189. B. uniflora Vasey var. uniflora. United States and northern Mexico, native. Rocky slopes: Nuevo Leon and Coahuila. 190. B. uniflora var. coahuilensis Gould and Kapadia Mexico, endemic. 1987 Beetle, Mexican grasses 225 Rocky slopes: Coahuila, Nuevo Leon, Tamaulipas, Zacatecas, San Luis Potosi, and Aguascalientes. 191. B. williamsii Swallen Mexico to Honduras, native. Rocky slopes: Zacatecas, Nayarit, Jalisco, Guerrero, Oaxaca and Chiapas. BRACHIARIA Griseb. Paniceae About 15, tropics and subtropics of the world. 192. Brachiaria arizonica (Scribn. & Merr.)Black U.S. and Mexico, native. Sandy soil: Baja California, Sonora, Chihuahua and Coahuila, Sinaloa, Durango, Oaxaca and Chiapas. 192a.Brachiaria arizonica var. major (Vasey)Beetle Nativa. endemic. Reported for Chihuahua and Sinaloa. 193. Brachiaria brizantha (Hochst.)Stapf. Introduced, cultivated Reported from Puebla and Yucatan. 194. B. ciliatissima (Buckl.)Chase Arkansas, Oklahoma, Texas and Mexico, native. Sandy soils: Tamaulipas and Nuevo Leon. 195. Brachiaria distachya (L.)Stapf Introduced, cultivated or escaped. Reported from Nuevo Leon, Jalisco, Veracruz and Yucatan. 196. Brachiaria fasciculata (Sw.) Parodi var. fasciculata U.S. , Wets Indies, Mexico south of Brazil, native. weedy, common throughout Mexico. 196a.Brachiaria fasciculata var. reticulata (Torr.)Beetle Same distribution as var. fasciculata. 197. B. meziana Hitchc. Mexico, endemic. Sandy soils: Chihuahua, to Tamaulipas and south to Oaxaca. 198. Brachiaria mollis (Sw.) Parodi Mexico and West Indies south to Argentina, native. coastal plain: Sinaloa south to Chiapas, Veracruz : to Yucatan, native. 199. Brachiaria mutica (Forsk.) Stapf Pantorpica (introduced in the Americas) Cultivated and escaped, grown throughout Mexico. 200. B. ophyrodes Chase Mexico, endemic. Sandy soils: Tamaulipas and Nuevo Leon. 201. B. plantaginea (Link)Hitchc. United States (where introduced); Mexico to Bolivia and Brazil, native. Sandy soils: Durango to Tamaulipas and south to Chiapas. 202. B. platyphylla (Griseb.)Nash U.S. , Cuba and Mexico, native. Wet places: reported for Mexico by Gould (1975). 226 P HY’ T O*RtONGRICA Vol. 63, No. 4 203. Brachiaria texana (Buckl.) Black U.S. and Mexico, native. Weedy, Nuevo Leon and Tamaulipas. BRACHYPODIUM Beauv. Festuceae About 15 species in the Old World; two in subtropical America. 204. Brachypodium distachyon (L.) Beauv. 205. Brachypodium latifolium Fourn. Native, endemic. Reported from Jalisco, Mexico, Veracruz and Oaxaca. 206. B. mexicanum (R. & S.) Link Mexico to Bolivia, native. Dry banks: common from Baja California Sur to Tamaulipas and south to Chiapas. 206a.Brachypodium mexicanum var. inerme Beetle Native, endemic. Reported from Nuevo Leon and Tamaulipas. 207. B. pringlei Scribn. Mexico, endemic. Dry banks: Coahuila, Nuevo Leon, and Tamaulipas. BRIZA L. Festuceae Three annuals in the Old World; about 15 New World. perennials. 208. B. minor L. Introduced. Roadsides: Jalisco, Mexico, Puebla and Veracruz 209. B. subaristata Lam. Mexico and Guatemala, native. Pine savanna: Nuevo Leon and Tamaulipas south to Chiapas. BROMUS L. Brachypodeae About 100 species, temperate zone world-wide. 210. B. anomalus Rupr. United States and Mexico, native. Pine savanna: northern border states south to Veracruz and Oaxaca. 211. Bromus arenarius Labill. Introduced from Australia. sandy roadside, reported only from Baja California Norte 212. B. arizonicus (Shear) Stebbins United States and Mexico, native. Dry washes: Nuevo Leon and Baja Norte and Sonora. 213. B. attenuatus Swallen Mexico, endemic. Pine savanna: Nuevo Leon and Hidalgo. 214. B. carinatus H. & A. United States to central America, native. Pine savanna: common throughout Mexico. 215% Be sciLiatus: Li. Canada and United States south to northern Mexico,native. Pine savanna: Baja Norte, Sonora, Chihuahua, Nuevo Leon, Zacatecas and Jalisco. 216. B. densus Swallen Mexico, endemic. Pine savanna: Coahuila, Nuevo Leon and Tamaulipas 1987 217. 218. 219. 220. 221. 22250 223%. 224. 225% 226. 2272 228. 229). 230. 231. Beetle, Mexican grasses 227 B. diandrus Roth Introduced. Roadsides: Baja Norte, Puebla. B. dolichocarpus Wagnon Mexico, endemic. Pine savanna: Colima, Jalisco, Michoacan, Hidalgo, Mexico, Morelos and Oaxaca. Bromus exaltatus Bernh. Southern Mexico to Panama, native. Jalisco and San Luis Potosi south to Chiapas. Bromus inermis Leyss. Introduced and cultivated. Experimental plots. e.g. in Coahuila. B. lanatipes (Shear) Rydb. (including B. pinetorum Swallen) U.S. and northern Mexico, native. Pine savanna: Coahuila. Bromus madritensis L. Introduced from the Mediterranean Region. Reported only for Baja California Norte. B. marginatus Nees Canada to northern Mexico, native. Pine savanna: Nuevo Leon, Sonora, and Baja California Norte. B. meyeri Swallen Mexico, endemic. Pine savanna: Nuevo Leon. B. mollis L. Introduced. Dry hills: Baja California Norte. B. mucroglumis Wagnon (including B. thysanoglottis Soderstrom & Beaman). Mexico, endemic. Pine savana: Chiahuahua, Durango and Baja California Sur, Coahuila, Nuevo Leon. B. porteri (Coult.)Nash Canada to Mexico, native. Pine savanna: northern border states south to Chiapas and Campeche. B. rubens L. Introduced. Dry hills: Baja California Norte. Bromus sterilis L. Introduced from the Mediterranean Recion. Reported only for Baja California Norte. B. tectorum L. and var. glabratus Spenner. Introduced. TW Cin as Dry hills: Baja California Norte. and Chihuahua. b. trinii Desv. California and northern Mexico; Chile, native. Coastal hills: Baja California Norte. 228 PH YT OF "O"Gr IVa Vol. 63, No. 4 232. B. unioloides HBK. Introduced. Ditches and cultivated: Chihuahua, Coahuila, Nuevo Leon, San Luis Potosi, Hidalgo and Mexico. 233. B. willdenowii Kunth var. willdenowii Introduced. Ditches and cultivated: northern border states south to Chiapas. 233a.Bromus willdenowii Kunth var. lasiophyllus (Goiran.) Beetle Introduced. BUCHLOE Engelm. Chlorideae One species in North America. 234. B. dactyloides (Nutt.)Engelm. Grassland: northern border states south to central mountains. BUCHLOMINUS R-& R.- Chlorideae One, endemic to Mexico. 235. B. nervatus (Swallen)R.& R. Mexico, endemic. Hidalgo and Mexico. CALAMAGROSTIS Adans. Agrostideae About 150 species, temperate, worldwide. 236. G€. exrectifolia Hitchc. Jalisco, endemica. 236a.C. eriantha (HBK) Steud. Mexico, endemic. Pine savanna: Mexico, Puebla, Veracruz and Oaxaca. 237. C. intermedia (Presl) Steud. Native, Mexico to Peru. Reported only from the State of Mexico. 238. C. junciformis (HB) Steud. Mexico and Guatemala, native. Mountain slopes: "Nevado de Toluca", type loc. 239. C. orizabae (Rupr.)Steud. Mexico, endemic. Mountain slopes: "Mount Orizabae", type loc. 240. C. pringlei Beal Mexico (endemic; closely related to C. gquatemalensis Hitchc.) Mountain slopes: Chihuahua, Nuevo Leon, Jalisco, Hidalgo, & Michoacan. 241. C. rigescens (Presl)Scribn. Native, Mexico and South America. Reported only from Cofre de Perote, Veracruz. 1987 242. 243. 244. Beetle, Mexican grasses 229 C. tolucensis (HB)Trin. Mexico, endemic. Mountain slopes: central mountains. C. valida Sohns Mexico, endemic. Mountain slopes: Jalisco. C. vulcanica Swallen Mexico and Guatemala. Mountain slopes: "Vulcan Tacana" Michoacan is in Chiapas". CATHESTECUM Presl Chlorideae 245. 246. 247. 248. Species six, North and Central America. C. brevifolium Swallen Mexico and Central America, native. Dry slopes: Sonora and Chihuahua south to Chiapas. C. erectum Vasey and Hack. Texas to Arizona and adjacent Mexico, native. Dry slopes: Chihuahua. C. prostratum Presl Mexico, endemic. Dry slopes and flats: Queretaro, Jalisco, Morelos, Puebla, Guerrero and Oaxaca. C. varium Swallen Mexico, endemic. Dry slopes: Puebla, Oaxaca and Jalisco. CENCHRUS L. Paniceae 249. 250. 251. 252... 253s 254. Species about 20, worldwide. C. brownii R. & S. Florida and West Indies to Bolivia and Brazil, native. Sandy soil: Baja Sur and Nayarit; Nuevo Leon and Tamaulipas; south to the Yucatan Peninsula. C. ciliaris L. Introduced? (closely related to C. multiflorus Presl which is said to be native). iz Ditches and cultivated: common throughout Mexico. C. echinatus L. Southern United States, West Indies to Argentina,native. Ditches: common throughout Mexico. C. incertus Curtis Southern United States, West Indies to South America, native. Ditches: common throughout Mexico. C. longispinus (Hack.)Fernald United States and West Indies to Venezuela, native. Ditches: Sonora, Durango, Tamaulipas and Campeche, Guanajuato, Michoacan & Oaxaca. C. multiflorus Presl Native? (closely related to C. ciliaris L. which is said to be introduced); also in Central America. Dry flats: southern Sonora and southern Chihuahua south along the west coast of Mexico to Chiapas. 230 255. 256. 257. P HUY T O8L{OsGe vA Vol. 63, No. 4 Cc. myosuroides HBK Florida and West Indies south to South America, native. Northern border states south to Veracruz and Oaxaca. C. palmeri Vasey Mexico, endemic. Sandy soils: Baja Norte, Baja Sur and Sonora (Tipe loc.) Sinaloa. C. pilosus HBK Mexico suth to Peru, native. Dry slopes: Nayarit, Jalisco, Colima south and east to Yucatan. CHABOISSAEA Fourn. Eragrosteae 258. 259. 260. One species, endemic. C. ligulata Fourn. Mexico, endemic. Dry slopes: Chihuahua south to Mexico C. bromoides (Presl) Benth. Mexico and Central America, native. Dry slopes: Chihuahua south to Campeche. Chasmanthium latifolium (Michx.) Yates U.S.A. and Mexico, native. Nuevo Leon, Veracruz, dunas costeras. CHLORIS Sw. 261. 262. 263. 264. 265. 266. 267. 70 species, subtropics, worldwide. C. andropogonoides Fourn. Texas and northern Mexico, native Ditches: Coahuila,Nuevo Leon, Tamaulipas and San Luis Potosi. C. aristata (Cervantes) Swallen Mexico and Central America, native. Coahuila, Nuevo Leon, south to Chiapas. C. brandegei (Vasey) Swallen Mexico, endemic. Arroyos: Baja California Norte and Baja California Sur. C. chloridea (Presl)Hitchc. U.S. and Mexico to Central America, native. Dry slopes: Baja California and Sonora south to Chiapas; Nuevo Leon, Tamaulipas, San Luis Potosi south to Chiapas. C. ciliata Sw. U.S., West Indies and Mexico to Argentina, native. Grassland: Coahuila, Nuevo Leon, Tamaulipas south to Yucatan Peninsula. C. c.inita Lag. Texas south to Argentina, native. Dry washes: Sonora, Chihuahua, Coahuila, Nuevo Leon, Baja California Sur, Durango and San Luis Potosi. C. cucullata Bisch. Texas and northern Mexico, native. Ditches: Coahuila, Nuevo Leon and Tamaulipas. 1987 Beetle, Mexican grasses 268. Chloris dandyana Adams. Native, subtropics of North and South America. Scattered localities throughout Mexico. 269. C. gayana Kunth Introduced. Cultivated and escaped: scattered locations throughout Mexico. 270. C. inflata Link Pantropical and subtropics, introduced. Dry washes: Sonora, Chihuahua, Nayarit, Jalisco, Tamaulipas, Veracruz and the Yucatan Peninsula. 271. C. pluriflora (Fourn.)Clayton Texas, Mexico, Central and South America, native. Dry washes: Coahuila, Nuevo Leon, Tamaulipas, San Luis Potosi, Jalisco, Michoacan, Oaxaca, Campeche. 272. GC. radiata (L.) Sw. West Indies and Mexico to Paraguay, native. Ditches: Nuevo Leon, Chihuahua, San Luis Potosi, Guanajuato, Colima, Veracruz and Oaxaca. 273. C. subdolichostachya Muller Texas and northern Mexico, native. Sandy soils: Coahuila, Nuevo Leon and Tamaulipas. 274. C. submutica HBK Mexico, endemic. Dry washes: common from the northern border states south to Chiapas. 275. C. verticillata Nutt. United States to northern Mexico, native. Plains: reported only from Coahuila. 276. C. virgata Sw. Pantropical, native. (U.S., West Indies south to Argentina). Ditches: common throughout Mexico. CHUSQUEA Bambuseae About 100 species in the Americas. 277. C. bilimeki Fourn. Mexico, endemic. Thickets: San Luis Potosi, Mexico and Veracruz. 278. C. carinata Fourn. Mexico, endemic. Thickets: Veracruz. 235i 232 279. 280. 281. 282. 283). 284. 285. 286. 287. 288. 289. 290. ZS. PH Y¥ T O8LsONGs WA Vol. 63, No. 4 Cc. circinata Soderstrom & Calderon Reported from Jalisco Michoacan (Type loc.) and Jalisco. C. coronalis Soderstrom & Calderon Subtropics, Mexico. C. galeottiana Rupr. Mexico, endemic. Thickets: Guerrero, Oaxaca (Type loc.) and Chiapas. C. lanceolata Hitchc. Reported for Chiapas. C. liebmannii Fourn. Mexico, endemic. Thickets: Sinaloa, Oaxaca (Type loc.) and Chiapas. C. longifolia Swallen Southern Mexico to Panama, native. Thickets: Sinaloa to Chiapas along the West Coast. C. muelleri Munro Mexico, endemic. Tickets: Veracruz. C. nelsonii Scribn.& Smith Mexico, endemic. Thickets: Guerrero (Type loc.). C. pittieri Hack. Chiapas, Guerrero. C. serrulata Pilger Mexico, endemic. Thickets: Chiapas. C. simpliciflora Munro Mexico and Guatemala, native. Thickets: "Mexico" according Swallen: Grasses of Guatemala. C. spinosa Fourn. Mexico, endemic. Thickets: Puebla. C. sulcata Swallen Mexico, endemic. Thickets: Chiapas. CINNA L. Agrostideae 29 2ic 3 species, one in N. America and Eurasia , one inN.Am. and one in Mexico south to South America. C. poaeformis (HBK) Scribn. & Merr. Mexico, Costa Rica, Panama, Colombia, Venezuela, Peru, native. Pine savanna: Hidalgo, Mexico, Morelos, Puebla, Veracruz, Oaxaca, Chiapas. COELORACHIS Brongn. Androfogoneae 293% Tropics of the World, about 25 species. C. ramosa (Fourn.) Nash Mexico to Panama and Colombia, native. Ditches: Michoacan, Mexico, Veracruz, Chiapas, Tabasco and Campeche. 1987 Beetle, Mexican grasses 233 COIX L. Andropoganeae 4 in the Old World tropics. 294. C. lacryma-jobi L. Introduced. Cultivated and escaped: scattered localities throughout Mexico, Coahuila, Nuevo Leon, Veracruz, Oaxaca, Chiapas, Tabasco and Yucatan. CORTADERIA Stapf Arundinae 15 species in South & America. 295. C. selloana (Schult.) Aschners. & Graebn. Introduced. Cultivated as an ornamental: throughout Mexico. COTTEA Kunth Pappophoreae One species, New World. 296. C. pappophoroides Kunth Bicentric: U.S. and Mexico; Ecuador to Peru and Argentina, native. Dry slopes: northern border states south to Oaxaca. CRYPTOCHLOA Swallen Olyreae 4 species in Mexico, Central and South America. 297. C. gramulifera Swallen Mexico to Ecuador, native. Tropical forest: Veracruz and Chiapas. 298. C. strictiflora (Fourn.) Swallen Mexico, endemic. Tropical forest: Veracruz. CTENIUM Panzer Chlorideae 20 species, world tropics. 299. C. planifolium (Presl) Kunth Mexico, endemic. Brush: Oaxaca and Chiapas. 300. C. plumosum (Hitchc.)Swallen : Mexico, endemic. Brush: Sinaloa and Nayarit, Jalisco. CYCLOSTACHYA Reeder and Reeder Chlorideae ‘One species in Mexico. : 301. C. stolonifera (Scribn.) Reeder and Reeder Mexico, endemic. Dry flats: Zacatecas, San Luis Potosi, Aguascalientes, Hidalgo and Mexico. CYMBOPOGON Spreng. Andropogoneae About 60 species in the Old World tropics. 302. C. citratus (DC)Stapf Introduced. Persistent after cultivation: Nuevo Leon, Jalisco, Veracruz, Oaxaca, Chiapas, Tabasco and Yucatan. 234 PH YT OyLpOsGeTvA Vol. 63, No. 4 303. C. nardus (L.) Rendle Introduced. Persistent after cultivation: Nuevo Leon, Oaxaca and Yucatan. CYNODON L. Rich. Chlorideae 6 species, one pantropical, others African. 304. C. dactylon (L.)Pers. Pantropical, native. Common throughout Mexico. 305. C. dactylon (L.) Pers. f. vivipara Beetle Found in Tamaulipas. 306. C. nlemfuensis Vanderyst Introduced from Africa, cultivated and escaped. Reported from Veracruz and Yucatan. 307. C. plectostachyum (Schum.) Pilger Introduced. Persistent after cultivation: scattered localities throughout Mexico. DACTYLIS L. Festuceae About 5 species, temperate, Old World. 308. D. glomerata L. Introduced. Persistent after or escaped from cultivation: Coahuila, San Luis Potosi, Veracruz, Mexico, Puebla, Oaxaca and Chiapas. DACTYLOCTENIUM Willd. Chlorideae 13 Old World species, subtropical. 309. D. aegyptium (L.)Willd. Introduced. Common weed throughout Mexico. 309a.Dactyloctenium aegyptium f. vivipara Beetle Found in Sonora. 310. Dactyloctenium aristatum Link A seashore grass introduced from Africa. Reported from Yucatan. 311. Dactyloctenium geminatum Hack. Introduced from Africa. Reported from Guerrero, Campeche and Quintana Roo. 312. Dactyloctenium scindicum Boiss. Introduced from Africa. Coastal, Baja California Sur and Yucatan. DANTHONIA Lam. & DC. Danthoneae 100 species, temperate, worldwide. 313. D. filifolia Hubbard Mexico, endemic. Pine savanna: Puebla (Type loc.)and Chiapas. 314. D. spicata (L.) Beauv. Canada to northern Mexico, native. Reported from Coahuila and Veracruz. 1987 Beetle, Mexican grasses 235 DESCHAMPASIA Beauv. Aveneae 315). 316. 327. 318). 319. 50 species, temperate, worldwide. D. danthonioides (Trin.)Munro Alaska to Baja California: Chile, native. Moist places: Baja Norte only. D. elongata (Hook.) Munro Alaska to central Mexico; Chile, native. Mt. slopes: Jalisco, Hidalgo, Mexico, Morelos and Veracruz. D. flexuosa (L.)Trin. Amphiatlantic, south in mountain to Mexico, native. Mountain slopes: Chihuahua, Nuevo Leon and Durango. D. liebmanniana (Fourn.)Hitchc. Mexico, endemic. Central mountains: Jalisco Michoacan, Mexico and Puebla. D. straminea Hitchc. Mexico, endemic. Central mountains: Mexico and Puebla. DICHANTHIUM Willemet Andropogoneae 320. 321. 322. 10 species, Old World subtropics. D. annulatum (Forsk.)Stapf Introduced. Cultivated and escaped: Chihuahua, Chiapas, San Luis Potosi and Quintana Roo. D. aristatum (poir.) C.E. Hubb. Introduced. Cultivated and escaped: Nuevo Leon and Tamaulipas. D. sericeum (R.Br.)Camus Introduced. Cultivated and escaped: Sonora, Tamaulipas. DIECTOMIS HBK Andropogoneae 323 324. 325. all species in the Americas. D. angustata Presl Native, Mexico, Central America and northern South America. Reported from Chiapas. D. fastigiata (Sw.) HBK. Pantropical, native. Thickets: Tamaulipas, Sonora, Sinaloa, Nayarit, Jalisco, Colima, Michoacan, Mexico, Morelos, Guerrero, Oaxaca, Chiapas. D. laxa Nees Cuba and Mexico, south to northern Brazil, native. Pine savanna: Sinaloa south to Chiapas, Guerrero. DIGITARIA Heist. Paniceae 326. 327. 328. 300 species, worldwide, temperate and tropical. D. badia (Scribn. & Merr.)Fernald Reported for Sierra de San Felipe, Oaxaca and Jalisco. D. bicornis (Lam.) R. & S. Pantropical, introduced? Mexico, a common weed at lower elevations. D. biformis Willd. Introduced from Asia. Disturbed land, throughout Mexico. 328a.D. biformis Willd. var. chrysoblepharis (Fig.& D.Not.) Beetle. 236 PH Y T 0%.0°0G a *A Vol. 63, No. 4 329. D. californica (Benth.)Henr. U.S. south to Bolivia and Argentina. Dry slopes: northern border states south to Puebla. 330. D. cayoensis Swallen Mexico and British Honduras. Brush: Yucatan and Chiapas. 331. D. ciliaris Retz. Pantropical, native. Common weed throughout Mexico. 332. D. curtigluma Hitchc. Mexico to Panama. Brush: Jalisco, Michoacan, Mexico, Puebla, Oaxaca and Chiapas. 333. D. distans (Chase) Fernald Mexico, endemic. Pond margin: Jalisco. 333a.D. filiformis (L.)Koel. E.U.A. & Mexico, native. 334. D. hitchcockii (Chase) Stuckert Texas and New Mexico, native. Dry slopes: Coahuila, Nuevo Leon, Tamaulipas and San Luis Potosi, also Oaxaca. 335. D. horizontalis Willd. Pantropical, native? Weedy: Sinaloa, Nayarit, Jalisco, Colima, Michoacan, Nuevo Leon, Tamaulipas, San Luis Potosi, Veracruz, Guerrero, Oaxaca, Chiapas, Yucatan and Quintana Roo. 336. D. insularis (L.)Mez United States, Mexico, West Indies south to Argentina, native. Brush: common throughout Mexico except for Baja California and central mountains. 337. D. leucites (Trin.)Henr. var. Jeucites Mexico and Guatemala, native. Dry banks: Nuevo Leon, San Luis Potosi, Jalisco, Michoacan, Mexico, Morelos, Puebla, Veracruz and Chiapas. 338. D. leucites var. glabella (Chase) Henr. Mexico, endemic. Dry banks: Veracruz and Michoacan. 339. D. leucocoma (Nash) Urban Florida, West Indies and Mexico, native. Dry banks: Mexico and Veracruz. 339a.D. obtusa Swallen Southern Mexico to Guatemala, native. Dry banks: Morelos, Veracruz, Oaxaca and Chiapas. 340. D. panicea (Sw.) Urban Eastern United States south to Mexico, native. Weedy: Chihuahua, Coahuila, Durango, Zacatecas, Jalisco, Oaxaca and Chiapas. 341. D. patens (Swallen)Henr. Texas and northern Mexico, native. Sandy soil: Sonora, Coahuila and Nuevo Leon. 1987 Beetle, Mexican grasses 237 342. D. pentzii Stent. var. minor Stent. Introduced. Persistent after cultivation: Tamaulipas, San Luis Potosi, Queretaro, Nayarit, Veracruz, Tabasco, Chiapas and Campeche. 343. D. sanguinalis (L.)Scop. Pantropical, native ?. Scattered localities in northern Mexico south to Queretaro. 344. D. similis Beetle Native, Mexico and West Indies. Reported from Tamaulipas. 345. D. ternata (Rich.)Stapf 346. D. vigiascens Link Pantropical, native? Weedy: Chiapas. DISSANTHELIUM Trin. Aveneae 347. D. californicum (Nutt.)Benth. Islands off the coast nf California and Baja California, native. Dry slopes: Baja California acc. to Hitchcock. 348. D. mathewsii (Ball)Foster & Smith Bicentric: Mexico and Chile, native. Dry slopes: Mexico and San Luis Potosi. DISTICHLIS Raf. Eragrosteae 6 species in the Americas. 349. D. palmeri (Vasey) Fassett Coastal Salt Marshes, native, endemic. Baja California Norte and Sonora. 350. D. spicata (L.) Greene var. spicata Coastal, Canada and U.S. to Central America, native. Salt marshes: Tamaulipas, Veracruz, Tabasco, Yucatan, Quintana Roo and Chiapas and Oaxaca. 351. D. spicata var. divaricata Beetle California and Mexico, native. Western salt deserts: Coahuila, San Luis Potosi and Jalisco west to Baja California. 352. D. spicata var. mexicana Beetle Mexico, endemic. Central plains: alkaline soils, Durango and Coahuila south to Puebla. 353. D. spicata var. stolonifera Beetle California and Mexico, native. West coast salt flats: Baja Norte and Sonora and Jalisco. 354. D. spicata var. stricta (Torr.) Beetle. U.S. and Mexico. Interior salt flats: northern border states south to Puebla. 238 PHY I OvL OC TVA Vol’. 63° Nona ECHINOCHLOA Beauv. Paniceae 20 species, subtropics and temperate, worldwide. 355. E. colonum (L.)Link Pantropical, introduced? Moist places: common througthout Mexico 355a.E. colonum f. vivipara Beetle 356. E. crusgalli (L.)Beauv. ‘ Europe, U.S. , Mexico. Introduced? Weedy: occasional througthout Mexico. 357. E. cruspavonis (HBK)Schult. Subtropics of the world, introduced? Wet places: occasional througthout Mexico. 358. E. holciformis (HBK) Chase Mexico and Guatemala, native. Wet places: Coahuila & Durango south to Mexico. 358a.E. jaliscana McVaugh endemic to central Mexico. 359. E. muricata (Beauv.)Fern. U.S. and northern Mexico, native. Wet places: Sonora and Chihuahua. 360. E. oplismenoides (Fourn.)Hitchc. Mexico, endemic. Wet places: Chihuahua, Durango, Zacatecas and Mexico. 361. E. polystachya (HBK)Hitchc. Wet places: Sonora and Chihuahua, south to Campeche. 362. E. pyramidalis (Lam.)Hitchc. and Chase Introduced. Wet places: Campeche. 363. E. walteri (Pursh) Heller U.S., Cuba, Mexico and Guatemala, native. Wet places: Chihuahua, Coahuila, San Luis Potosi, Tabasco, Campeche. ECHINOLAENA Desv. 364. E. gracilis Swallen Reported for Tabasco. ELEUSINE Gaertn. Chlorideae Species about six, South America and Old World. 365. E. indica (L.)Gaertn. var. indica Introduced. Weedy: common throughout Mexico. 366. E. indica var. brachystachya Trin. (including the cultivated E. corocana (L.)Gaertn.) Introduced. Weedy: Chihuahua, Jalisco, Michoacan, Mexico, Morelos and Yucatan. 367. E. multiflora Hochst. ex A.Rich. Introduced. Weedy: Scattered localities Chihuahua south to Chiapas. ELYMUS L. Hordeae 75 species in temperate parts of the northern hemisphere. 368. E. canadensis L. var. canadensis United States and northern Mexico, native. Dry banks: Chihuahua, Coahuila and Nuevo Leon. 1987 Beetle, Mexican grasses 239 369. E. canadensis var. interruptus (Buchl.) Church E.U.A. and northern Mexico, native. Dry banks : Nuevo Leon, San Luis Potosi, Queretaro. 370. E. condensatus Presl California and Baja California, native. Coastal bluffs: Baja Norte. 371. E. glaucus Buckl. Western N. America from Alaska to Baja California. Open woods and meadows; reported for Baja Calif. Norte. 372. E. pringlei Scribn. & Merr. Mexico, endemic. Dry banks: Coahuila, Nuevo Leon, San Luis Potosi, Queretaro, Hidalgo, Mexico, Puebla and Veracruz. 373. E. triticoides Buckl. Washington and Montana south to Baja California, Arizona and Texas, native. Coastal bluffs: Baja California Norte and Tamaulipas. ELYONURUS Humb. & Bonpl. Andropogoneae Species about 15, subtropical and tropical, worldwide. 374. E. muticus (Spr.)Ktze.var. parviflorus (Scribn.) Beetle U.S. and northern Mexico, native. Grasslands: Sonora, Chihuahua and Durango, Nayarit, Guanajuato, Queretaro. 375. E. tripsacoides Humb. & Bonpl. var. tripsacoides Mexico to Brazil, native. Grasslands: Sinaloa, Michoacan and Veracruz. 376. E. tripsacoides var. sericeus Hack. Mexico, endemic. Rare: Veracruz common, Chihuahua aouth to Chiapas. ENNEAPOGON Desv. Eragrosteae 35 species in the Old World, one in the Americas. 377. E. desvauxii Beauv. U.S. south to Peru and Argentina, native. Grasslands: northern border states south to Oaxaca. ERAGROSTIS Host. Eragrosteae 200 species, temperate and tropical, worldwide. 378. E. acutiflora (HBK) Nees Mexico to Brazil, native. Pine savanna: Western slope. 379. E. atrovirens (Desf.)Steud. Introduced. Tabasco & Chiapas. 380. E. bahiensis Schrad. Mexico, Guatemala, Brazil, native. Wet places: Chiapas and Tabasco. 381. E. barrelieri Dav. Introduced. Scattered localities throughout Mexico. 382. E. capillaris (L.)Nees. Eastern United States and northern Mexico. Open woods, reported from tamaulipas. 383. E. chloromelas Steud. Introduced. Grown experimentally in Coahuila. 240 384. 385. 386. 388. 389. 390. yah Sh 2) 394. 395i. 396. 397. 398i: 399) 400. 401. PHY TO) LO Gel A Vol. 63, No. 4 E. cilianensis (All.)Link Introduced. Scattered localities throughout Mexico. E. ciliaris (L.) R.Br. var. ciliaris Pantropical, introduced. Weedy: common throughout Mexico. E. ciliaris var. laxa Kuntze West Indies and Mexico, native. Reported for Yucatan and Baja California. E. contrerasii Pohl Planicies lodosas, Mexico and Guatemala. Reported from Campeche. E. curtipedicellata Buckl. U.S. to northern Mexico, native. Sandy soil: Coahuila and Nuevo Leon. E. curvula (Schrad.)Nees Introduced. Cultivated: Scatterel location, Chihuahua, Coahuila, Nuevo Leon, south to Mexico. E. ellioitti S. wats. U.S., West Indies, Mexico and Honduras, native. Pine savanna: Tamaulipas, Michoacan, Veracruz, Oaxaca, Chiapas and Quintana Roo. E. erosa Scribn. U.S. and northern Mexico, native. Dry slopes: Sonora, Chihuahua, Coahuila and Nuevo Leon. E. excelsa Griseb. West Indies and Mexico, native. Dunes, Tamaulipas, Veracruz and Quintana Roo. E. glandulosa Harvey Mexico and Guatemala, native. Weedy: galisco, Guerrero and Morelos. E. glomerata (Walt.)L.H. Dewey U.S., Mexico south to Argentina, native. Wet sandy soil: Colima, Guerrero and Morelos. E. guatemalensis Witherspoon Mexico and Guatemala, native. Oak-pine forest, Mexico and Puebla. E. hirsuta (Michx.)Nees Southeastern U.S., México to Central America, native. Pine savanna, Oaxaca, Chiapas and Tabasco. E. hirta Fourn. var. hirta Mexico and Central America, native. Rocky slopes: Chiapas. E. hirta var. longiramea (Swallen)Witherspoon Mexico, endemic. Mountains, Tamaulipas, Nuevo Leon and San Luis Potosi. E. hypnoides (Lam.)B.S.P. U.S. and West Indies, south to Argentina, native. Wet, sandy soil: Baja California Sur, Sonora, Tamaulipas and San Luis Potosi, Guanajuato, Guerrero, Oaxaca, Chiapas, Veracruz, Tabasco. E. intermedia Hitchc. var. intermedia U.S. , Mexico and Guatemala, native. Rocky slopes: common throughgut Mexico. E. intermedia var. appressa Withrspoon Mexico, endemic. Juniper oak forest, border Zacatecas-Jalisco. 1987 Beetle, Mexican grasses 241 402..E. intermedia var. oreophila (Harvey) Witherspoon Mexico, endemic. Rocky slopes: Baja California Sur, Coahuila, Nuevo Leon, Michoacan, and Hidalgo (Type loc.) 403..E. intermedia var. praetermissa (Harvey) Witherspoon Mexico and Central America. Pine oak forests, Tamaulipas, San Luis Potosi, Jalisco, and Mexico. 404. E. lehmanniana Nees Introduced. Cultivated: Sonora, Chihuahua, Coahuila. 405. E. lugens Nees United States and Mexico south to Argentina, native. Dry slopes: northern border states south to Chiapas. 406. E. maypurensis (HBK) Steud. Mexico to Bolivia and Brazil, native. Weedy: Sinaloa and Durango south to Chiapas. 407. E. mexicana (Hornem.)Link var. mexicana U.S. and Mexico south to Brazil, native. Pine savanna: northern border states south to Chiapas. 408. E. mexicana var. pygmaea Doell. Same distribution as the species. Reported from scattered localities throughout Mexico. 409. E. obtusiflora (Fourn.)Scribn. Mexico, endemic. Salt flats: Chihuahua, Coahuila, Nuevo Leon, Mexico and Veracruz. 410. E. orcuttiana Vasey U.S. and northern Mexico, native. Weedy: Baja California Sur, & Baja California Norte. 411. E. palmeri Wats. Texas and northern Mexico, native. Open slopes: northern border states except for Baja California Norte, Durango, Zacatecas, Oaxaca. 412. E. pectinacea (Michx.)Nees 5 United States and Mexico, native. Weedy: Coahuila and Zacatecas. 413. E. pilosa (L.)Beauv. Introduced. ‘Weedy: common, northern border states south to Chiapas. 414. E. plumbea Scribn. Mexico, endemic. Dry slopes: Jalisco (Type loc.)Colima, Michoacan, San Luis Potosi, Puebla, Veracruz, Guanajuato. 415. E. pringlei Mattei Mexico, endemic. Weedy: Chihuahua, Durango, Jalisco and Michoacan. 4i6. E. prolifera (Sw.)Steud. West Indies, southern Mexico and Central America to Brazil, native. Weedy: Michoacan, Veracruz and Quintana Roo. £17. E. reptans (Michx.)Nees U.S. south to northern Mexico, native. Wet, sandy soil: Baja California Norte and Coahuila. 242 PH YT O°LsONC) TA Vol. 63, No. 4 418. E. secundiflora Presl var. secundiflora Brazil and Mexico, native. Peninsula Reported from Guerrero and Oaxaca. 419 .E. secundiflora Presl var. capitata (Fourn.)Beetle Florida to Texas and Mexico, native. Reported for Veracruz and Tamaulipas. 420. E. sessilispica Buckl. U.S. and northern Mexico, native. Sandy prairies: Chihuahua and Tamaulipas. 421. E. silveana Swallen Texas and Mexico, native. Sandy soil: Coahuila, Tamaulipas and San Luis Potosi. 422. E. simpliciflora (Presl)Steud. Southern Mexico to Panama, native. Weedy: "southern Mexico" acc. to Swallen: Grasses of Guatemala. 423. E. spectabilis (pursh)Steud. U.S., Mexico and British Honduras, native. Weedy: "northeastern Mexico" acc. to Harvey. 424. E. spicata Vasey Texas and Mexico, native. Coastal: Tamaulipas and Baja California Sur. 425. E. superba Peyr. Introduced. Cultivated: Sonora, Chihuahua, Coahuila, Nuevo Leon, & Michoacan. 426. E. swalleni Hitchc. Texas and Mexico, native. Sandy soil: San Luis Potosi, Jalisco, Veracruz and Chiapas. 427. E. tenella (L.)Beauv. Pantropical, introduced? Weedy: Baja California Sur, Nayarit, Jalisco, Guerrero, Puebla, Veracruz, Oaxaca, Tabasco, Yucatan and Quintana Roo. 428. E. tenuifolia (A. Rich.)Hochst. Introduced from Africa. Weedy. Reported from Oaxaca. 429. E. tephrosanthos Schult. U.S., West Indies, south to Brazil, Native. Weedy: common throughout Mexico. 430a.E. trichocolea Hack. and Arech. var. floridana (Hitch.) Witherspoon. U.S. and Mexico to Uruguay. Sandy soil: Puebla, Oaxaca and Chiapas. 430b.E. viscosa (Retz.)Trin. Introduced. Sandy soil: Baja California Norte, Baja California Sur, Sinaloa, Durango, Guerrero and Chiapas. 430c.E. yucatana Harvey Mexico, endemic. Sandy soil: Yucatan. EREMOCHLOA Buese 10 species in subtropical Asia. 430d.E. ophiuroides (Munro) Hack. Introduce cultivated as a lawn grass. 1987 Beetle, Mexican grasses 243 ERIANTHUS Michx. Andropogoneae Twenty-five species, temperate & subtropical,worldwide 430e.E. giganteus (Walt.)Muhl. U.S., Cuba and Mexico, native. Swamps: Mexico, Hidalgo and Veracruz. 430f.E. trinii Hack. Mexico and South America, native. Swamps: Nuevo Leon, San Luis Potosi, Queretaro, Hidalgo, Veracruz and Chiapas. ERIOCHLOA HBK Paniceae Twenty-five species, subtropical, worldwide. 430g.E. aristata Vasey Mexico and Guatemala, native. Wet places: common from Sonora and Chihuahua south to Chiapas. 430h.E. boxiana Hitchc. West Indies and Mexico, native. oan a Yucatan. constracta Hitchc. 430i. S.A. & northern Mexico acc. Gould. 4305 a lemmoni Vasey and Scribn. var. lemmoni U.S. and northern Mexico, native. Weedy: Sonora, Chihuahua, Coahuila, Sinaloa, Jalisco, Colima, and Mexico. 430k.E. lemmoni var. minor (Vasey) Beetle Same distribution as the species, native. Weedy: northern border states south to Oaxaca. 4301.E- nelsoni Scribn. & Smith. Mexico, Guatemala and Nicaragua, native. Open slopes: Sonora, and Chihuahua south to along the west slope. Oaxaca (Type loc.) 430m.E- punctata (L.)Desv. -S., West Indies, Mexico south to Argentina, native. Open slopes: Chihuahua, Coahuila and Nuevo Leon, south to Chiapas and Tabasco. 430n.E- sericea (Scheele) Munro U.S. and northern Mexico, native. Grassland: Coahuila. ERIOCHRYSIS Beauv. Andropogoneae Nine species, 4 in America, four in Africa, one in India. 4300.E. cayennensis Beauv. Southern Mexico south to northern Argentina and Brazil, native. Swamps: Oaxaca, Veracruz, Chiapas, Tabasco. ERIONEURON Nash Eragrosteae 5 species, southwest U.S. and Mexico. 430p.E. avenaceum (HBK)Tateoka var. avenaceum U.S. and Mexico, native. Dry slopes: Coahuila and Nuevo Leon south to Oaxaca. 430q.E- avenaceum var. longiaristatum (Kurtz) Beetle U.S. and Mexico, native. Chihuahua, Coahuila and Nuevo Leon, Durango, Zacatecas and San Luis Potosi, Guanajuato, Queretaro, Guerrero. 430r.E- avenaceum (HBK)Tateoka var. nealleyi (Vasey) Gould. U.S. and Mexico, native. 244 PHY TO Lore LA Vol. 63, No. 4 Open range: Chihuahua, Coahuila, Nuevo Leon, Tamaulipas, Durango, Zacatecas and Queretaro, Aguascalientes, San Luis Potosi and Hidalgo. 430s.E. pilosum (Buckl.)Nash U.S. and Mexico, native. Open range: Coahuila, Nuevo Leon, Tamaulipas, Zacatecas, Aguascalientes, San Luis Potosi, and Oaxaca. 430t E. pulchellum (HBK) Tateoka : U.S. and Mexico, native. Open range: northern border states south to Mexico. EUCHLAENA Schrad. Andropogoneae 2 species in Mexico and Central America. 430u.E. mexicana Schrad. Cultivated, endemic? (native of the Americas). Cultivated: northern border states south to Chiapas. 431. Euclasta condylotricha (Hochst.) Stavnf Mexico and West Indies to Colombia, Venezuela and northern Brazil, native. Subtropical clearings: finaloa, Jalisco, Colima, Michoacan, Veracruz, and Oaxaca. EUSTACHYS Desv. Chlorideae 12 species, trorics and subtronics, worldwide. 432. E. petraea (Sw.) Desf. U. €., West Indies, Mexico anc Central America, native. East coast, Nuevo Leon, Tamaulivas, Veracruz, Tabasco and the Yucatan Peninsula. 1987 Beetle, Mexican grasses 245 Pes cuGcAy ti. Festuceae 433. 438. 439. Toc) 440. 440a. 441. 442. About 150 snecies, temnrerate, worldwide. F. amnlissima Rupr. Mexico and Central Ameri-a, native. Baia California Norte & Sonora south also, the west coast to Jalisco; also hich mountains of the central plateau, Michoacan, Hidalgo, Mexico, Morelos, Guerrero, Puebla, Oaxaca and Chianas. F. arundinacea Schreb. Introduced. Cultivated: northen states south to Puebla. tT. brevioluris Swallen Mexico and Central America, native. Hich forests: Jalisco and Guanajuato south to Chianas. Festuca fratercula Ruor. U. &. and northern “exico, native. Pine savanne, Nuevo Leon, Chihuahua, Conora, Durango & Guanajuato alse Veracruz (Orizaba). F. henhaestophila Nees Mexico, endemic. Central mountains: Nuevo Leon (Cerro Potosi), Puebla, and Veracruz (Orizaba). F., licgulata Swallen Texas and northern Mexico. Mountain slores: Coahila. F. mirabilis Pirer Mexico, endemic. “Mountain slores: Chihuahua; also £an Luis Potosi (tvre *Hidalgo and Mexico. F. ovina L. var. brachyphvlla (Schult.) Piper Circumboreal and scuth in the hich mountains, native. Mountain slones: Chihuahua, Durango, Nuevo Leon, Tamaulipas, Mexico and Pueble. Festuca ovina L. var. clauca Fries Introduce, Cultivated as ornamental. F. rosei Piner Fexicc, endemic. Central mountains: Michoacan and Mexico. me Eubra. Le -Circumboreal and south in the hich mountains, native. Mountain slores: Chihuahua and Mexico. 246 443. 444, PHY TO L'o"G TA Vol. 63, No. 4 F. tolucensis HBK Mexico and Central America, native. Pine savanna: Chihuahua south to Chianas. F. wildenoviana Schult. Mexico, endemic. Central mountains: Guerrero, Mexico and Puebla. GASTRIDIUM Beauv. Acrostideae 445. 4 or 5 species, Mediterranean. Gastridium ventricosum (Gouan) Schinz & Thell. Introduced. Dry hills: Baia California Norte. GLYCERIA R. Br. Festuceae 446. 447, 448. 449, 450. 35 snecies, temnerate, worldwide. Glvceria borealis (Nash) Batchelder Native, Alaska, Canada, United States and northern Mexico. Reported from Chihuahua. Gi fluatans) (ie)) bee ses. Circumboreal and south in the high mountains, native. Central mountains: San Luis Potosi, Guanajuato, Querétaro, Hidalco and Mexico. Glyceria mexicana (Kelso) Beetle Mexico and Guatemala, native. Wet rlaces: Hidalgo, Mexico, Morelos Oaxaca and Chiapas. G. septentrionalis Hitchc. Canada south to Mexico, native. Wet places: reported by Hernandez-X¥ for San Luis Potosi. G. striata (Lam.) Hitchc. Canada south to Mexico, native. Wet places: Sonora and Chihuahua. GOUINIA Fourn. Chlorideae 451. 452. 453. 454, 455. 456. G5. 13 species, subtropical, in the Americas. G. guatemalensis (Eack.) Swallen Mexico and Central America, native. Brush: Chiaras, Camneche and Yucatan. G. longiramea Swallen Mexico, endemic. Brush: Yucatan and Quintana Foo. G. mexicana (Scribn.) Vasev Mexico, endemic. Brush: San Luis Potosi. G. pavillosa Swallen Mexico, endemic. Brush: Yucatan and Ouintana Roo. GS. ramosa Swallen “exico, endemic. Brush: Oaxaca, Camneche, Yucatan and Ouintana Roo. G. virgata (Presl) fcribn. Mexico south to Peru, Bolivia and Brazil, native. Brush: Sonora and Nuevo Leon south to Chiapas, Camneche and Yucatan. Griffithsochloa multifida (Griffiths) Pierce Mexico, endemic. Dry slopes: Mexico, Morelos, Guerrero and Oaxaca. 1987 Beetle, Mexican grasses 247 GYMNOTOCGON Beauv. Chlorideae 15 snecies, subtronical, Americas. 458. G. snicatus (Srrenc.) Kuntze Mexico and “est Indies south to Argentina, native. Pine savanna: Verecruz and Chiavnes. GYNERIU™ Willd. Arundineae Monotvric. 459. G. sagittatum (Aubl.) Beauv. Southern Mexico to Paracuav, native. Thickets: Oaxaca, Veracruz, Chianas and Tabasco. KACKELOCHLOA Kuntze Paniceae Monotvnic. 460. H. aranularis (L.) Kuntze Pantrovical, native. weedy: occasional throughout Mexico. 461. Helleria livida (HEBK) Fourn. Mexico, endemic. Central mountains: Mexico, Puebla and Veracruz. EEMARTHRIA R. Br. Androroconeae 12 species, Cld World, trorics and subtronics. 462. H. Introduced. Cnen slopes: Coahuila, Nuevo Leon, San Luis Potosi, Mexico, south to Chiares and Tabasco. HETFEROPOGON Pers. Andronogoneae 8 species, subtrorics, worldwide. 463. H. contortus (L.) Beauv. U. S., west Indies, and Mexico south to Arcentina (native?), also in the Old World. Weedyv: common throughout Mexico. 464. H. melanocarrus (E1ll.) Benth. Pantropical, native? Weedyv: northern border states south to Chiaras. ETE POCHLOS,; R= Br: Species 15-20, temrerate, worldwide. 465. E. mexicana (Rupnr.) Benth. Mexico and Guatemala, native. Pine savanne: Caxaca (tvre loc.) Morelos and Chiapas. HILZRIA HBK Chlorideae Six species, in the 23mericas. 466. H. belanceri (Steud.) Nash var. belangeri U. S. south to central Mexico, native. Grassland: northern border states south to Oaxaca. 466a. H. belanoeri var. longifolia (Yasev) Hitchc. Reported in both U. S. and Mexico. Rocky hills, Sonora (tvpe loc.), Coahuila. 248 PHY T Oi L0G \1yA Vol. 63, No. 4 467. i. cenchroides HBK Mexico ana Guatemala, native. Oren slopes: common from northern border states south to Chiavdas. 468. H. ciliata (Scribn.) Nash Mexico, endemic. Open slones: Sonora, Baja California Sur, Mavarit, Tamaulipas, San Luis Potosi, Jalisco (tvne loc.), south to Oaxaca. 469. E. hintoni Sohns Mexico, endemic. Queretaro, Mexico and Guerrero. 470. H. mutica (Buckl.) Benth. U. S. and northern Mexico, native. Dry flats: Sonora, Chihuahua, Coahuila, Nuevo Leon, Duranco and Zacatecas. 471. H. ricida (Thurb.) Benth. U. S. and northern Mexico, native. Desert: Baja California Norte, Sonora and Chihuahua. 472. HE. semnlei Sohns Mexico, endemic. Llanos, Michoacan (tyve loc.). 473. H. swalleni Cory Texas ana northern Mexico, native. Grasslands: Chihuahua, Coahuila, Nuevo Leon, Duranco, Zacatecas and fan Luis Potosi. HEECUS) ui Avereae About 8 snecies, Old World, temnerate. 474. Hi lanatus L. Introduced. Weedy: Chihuahua, Puebla, Veracruz and Chianras. HOMOLZSPIE Chase Paniceae Three species in trovical America. 475. H. aturensis (EBK) Chase Southern Mexico to Bolivia and Brazil, native. Marshes: Oaxaca, Chianas, Veracruz and Tabasco. MORDEUIS Li. Lordeae Twenty-five snecies, temnerate, worldwide. 47u. Hordeum adscendens HBK Annarently narive? weedy. Coahuila, San Luis Potcsi, Mexico, Aquascalientes and fFuebla. 477. Hordeum arizonicum Covas Native, #£rizona, soutnern Caiifornia and Mexico. Reported by Wicgiis for Eaia California Norte. 476. Hordeum brachyvantherum Nevski Temnerate Kerth America, native. Re-orted Isy Goulc for Baja California isorte. 479. Bordeum depressum (fcribn. & fiitr Rycb. Native, Western Nerth America. “encrted bv Wicairs: "Moist elkaline Gerressio:.: anc flets’ BavarGolatounas , 30s ter 1987 Beetle, Mexican grasses 249 480 H. glaucum Steud. Introduced. — Mediterranean: Baja California Norte. 481. Hordeum hystrix L. Introduced, widespreadin North America. Reported by Gould and Moran for Baja California Norte. 482. Hordeum jubatum L. Introduced: reported by Gould for Baja California Norte. 483. H. pusillum var. pubens Hitchc. U. S. and Northern Mexico, native. Weedy: Coahuila. 484. H. vuigare L. Introduced. Cultivated: scattered localities in Central and northern Mexico. 485. Hydrochloa caroliniensis Beauv. var. caroliniensis. Acuatic, native. Jalisco, etc. 485a. Hydrochloa caroliniensis var. conneri Guzman Native, acuatic, described from & endemic to Jalisco. HYMENACHNE Beauv. Paniceae 10 species, tropics, worldwide. 486. H. amplexicaulis (Rudge) Nees Mexico and West Indies south to Argentina, native. Marshes: West from Sinaloa to Chianas, also Veracruz, Tabasco and Campeche. HYPARRHENIA Andropogoneae Seventy species, tropical and subtropical Africa, one in Americas. 487. H. bracteata (H.& B.) Stapf Mexico to Brazil and Paraguay, native. Marshes: Veracruz, Chiapas, and Oaxaca. 488. H. hirta (Nees) Stapf Introduced. Cultivated: Nuevo Leon, San Luis Potosi, Oaxaca, Chiapas and Yucatan. 489. H. rufa (Nees) Stapf Introduced. Cultivated and escaped: Sinaloa south to the Yucatan Peninsula. 490. Hyparthelia dissoluta (Nees ex Steud.) Clayton Introduced from Africa. Rocky slopes, Sinaloa, Nayarit, Jalisco, Michoacan, Mexico, Chiapas and Yucatan. 491. Hypogynium virgatum (Desv.) Dandy West Indies, Mexico, Central America south to Argentina. Reported from Oaxaca, Tabasco ICHNANTHUS Beauv. Paniceae About 50 species in the New World tropics, 2 in the 01 World tropics. 492. I. calvescens (Nees) Doell. Mexico and British Honduras, native. Oak forest: Oaxaca. POR YT) 0 LOG ak. Vol. 63, No. 4 493. I. lanceolatus Scribn. & Sm. Mexico and British Honduras, native. Brush: Yucatan Peninsula, Chiapas, Tabasco, and the Yucatan Peninsula. 494, I. nemorosus (Sw.) Doell. West Indies and Mexico south to Panama, native. Brush: San Luis Potosi, Hidalgo, Veracruz, Oaxaca and Chianas. 495. I. pallens (Sw.) Munro Mexico and West Indies south to tronical S. America. Brush: San Luis Potosi, Puebla, Veracruz, Oaxaca, Chiapas, Tabasco and Quintana Roo. 495a. I. pallens f. monstrosum (Fourn.) Beetle 496. TI. standleyi Hitchc. Mexico and Central America, native. Forest margin: Chiapas. 497. I. tenuis (Presl) Hitchc. and Chase Mexico, Central America, south to Brazil, native. Brush: Chiapas, Veracruz. IMPERATA Cyrill Andropogoneae Ten species, subtropical, worldwide. 498. I. brasiliensis Trin. Florida, Mexico, and south to Argentina, native. Brush: Oaxaca, Veracruz, Tabasco, Chiapas, and Campeche. 499. I. brevifolia Vasey Texas to California and Mexico, native. Brush: Baja California Norte, Sonora, C hihuahua, Nayarit and Jalisco. 500. I. contracta (HBK) Hitchc. Mexico and West Indies south to Brazil, native. Grassland: Hidalgo, Veracruz, Tabasco, Chiapas and Campeche. 501. I. cylindrica (L.) Beauv. Introduced. Cultivated or escaped: Chiapas, reported by Gould. ISACHNE R. Br. Paniceae Twenty-five species, tropics, worldwide. 502. I. arundinacea (Sw.) Griseb. Mexico, Central America to Peru, native. Pine savanna: Puebla, Veracruz, Oaxaca, Chiapas. 503. I. pubescens Swallen Mexico and Guatemala, native. Pine savanna, Veracruz and Chiapas. ISCHAEMUM L. Paniceae Fifty species, tropics, worldwide. 504. I. latifolium (Spreng) Kunth Mexico and West Indies south to Brazil, native. Weedy: Veracruz, Oaxaca, Chiapas and Campeche 505. Ischaemum rugosum Salisb. Introduced from Asia. Reported from Nayarit and Guerrero. 1987 Beetle, Mexican grasses 251 IXOPHORUS Schlecht. Panicum 506. Ixophorus palmeri (Vasey) Beetle Native, endemic. Reported from Jalisco and Colima. 507. I. unisetus (Presl) Schlecht. Mexico, Cuba and Central America, Colombia and Venezuela, native. Swamps: Sinaloa and Tamaulipas south to Chiapas. JOUVEA Fourn. Eragrosteae Two species, Mexico and Central America. 508. J. pilosa (Presl) Scribn. Mexico and Central America, native. Sand dunes: western coastal states: Tamaulipas, Tabasco, Campeche and Yucatan. 509. J. straminea Fourn. Mexico and Central America, native. Salt marshes: west coast from Sinaloa to Chiapas. KOELERIA Pers. Aveneae Fifty species, temperate, worldwide. 510. K. californica (Domin) Beetle Calif. and Mexico, Native. Dry slopes: northern border states at low 511. Koeleria cristata (L.) Pers. var. cristata Circumboreal, native. Central states at high elevations. 5lla. Koeleria cristata var. aqeniculata (Fourn.) Beetle Mexico, endemic. Pine forests, central highlands. 512. Koeleria phleoides (Vill.) Pers. Introduced from the Mediterranean Region. Reported only for Baja California Norte. LAMARCKIA Moench. Monotynvic. 513. L. aurea (L.) Moench. Introduced. Weedy: Baja California Norte. LASIACIS (Griseb.) Hitchc. Paniceae Thirtv species in American tropics. 514. L. divaricata (L.) Hitchc. var. divaricata 513. Florida, West Indies, Mexico south to Argentina, native. Tropical forest margin: Baja California Sur, Sinaloa, Nuevo Leon, San Luis Potosi south to Yucatan Peninsula. 514a. Lasiacis divaricata var. leptostachva (Hitchc.) Davidse Mexico and Central America, native. Tropical forest margins, Oaxaca and Chiapas. 515. L. grisebachii (Nash) Hitchc. Mexico, Cuba and Central America, native. Tropical forest margin: Veracruz, Chiapas, Camneche and Quintana Poo, also San Luis Potosi, Guerrero, Oaxaca, Puebla. 252 PHY T O}2sOrG, Wh Vol. 63, No. 4 516. Lasiacis linearis Swallen Mexico and Central America, native. Cloud forests or pine woods, Chiapas. 517. Lasiacis nigra Davidse Mexico to Peru, native. Montane, forest margin, common, Nayarit and Nuevo Leon south to Chiapas. 518. L. oaxacensis (Steud.) Hitchc. West Indies, Mexico, Central America, south to Peru, native. Brush: Michoacan, Oaxaca, Veracruz and Chiapas, also Nayarit and Jalisco. 519. L. vrocerrima (Hack.) Hitchc. Mexico to Peru and Brazil, native. Brush: Sinaloa, Durango, Nayarit south to Chiapas and Tabasco 520. L. rhizophora (Fourn.) Hitchc. Mexico and Central America, native. Thickets: Veracruz and Chiapas, also Oaxaca, Guerrero and Morelos. 521. L. rugelii (Griseb.) Hitchc. West Indies and Mexico, native. Thickets: Yucatan, Guanajuato and San Luis Potosi south to the Yucatan Peninsula. 522. L. ruscifolia (HBK) Hitchc. West Indies, Mexico south to Peru and Argentina, native. Brush: common throughout Mexico. 523. L. scabrior Hitchc. Mexico, Central America, Columbia and Ecuador, native. Brush: Veracruz, Oaxaca, and Chiapas. 524. I.sloanei (Griseb.) Hitchc. West Indies and Mexico, Central America, Colombia and Venezuela, Ecuador, native. Brush: Tamaulipas, San Luis Potosi south to Chiapas and the Yucatan Peninsula. 525. L. sorghoidea (Desv.) Hitchc. and Chase West Indies, Mexico, Central America south to Bolivia and Argentina, native. Brush: Mexico, Veracruz, Oaxaca and Chiavas. 526. Lasiacis standleyi Hitchc. Mexico to Venezuela and Ecuador. Shady woods, Chiapas. LEERSIA Sw. Oryzeae Ten species, worldwide, aquatic. 527. L. hexandra Sw. Pantropical, native. Aquatic: Durango, Jalisco, south to Chiapas and Tabasco. $28. “i. ligttlaris Trin. war. Diguiaris Mexico, endemic. Aguatic: Puebla, Veracruz. 528a. Leersia ligularis Trin var. breviligularis (Prod.) Pyrah Mexico and Guatemala, native. Forests Nuevo Leon and Tamaulipas south to Chiapas and Yucatan. 1987 Beetle, Mexican grasses 253 529. L. monandra Sw. U. S., West Indies and Mexico, nstive. Aquatic: Nuevo Leon and Tamaulipas, San Luis Potosi, Tabasco and Yucatan. 530. L. orvzoides (L.) Sw. Canada, U. S. and northern Mexico, native. Aquatic: Nuevo Leon, Tamaulipas and San Luis Potosi. LEPTOCHLOA Beauv. Chlorideae Seventy species, subtropical, worldwide. 531. Ee aquatica Seribn. Mexico, endemic. Aquatic: Jalisco, Michoacan, Morelos (type loc.) and Guerrero. 532. Leptochloa digitatiformis Beetle Mexico, native, endemic. Reported from Sonora. 533. L. domingensis (Jacq.) trin. Mexico, endemic. Swales: Tamaulipas, San Luis Potosi, south to Yucatan Peninsula. 534. L. dubia (HBK)Nees U. S., Mexico and Argentina, native. Swales: northern border states, south to Chiapas. 535. L. fascicularis (Lam.)Gray U. S. and Mexico, Central and south America, native. Swales: common throughout Mexico. 536. L. filiformis (Lam.) Beauv. U. S. south to Argentina, native. Swales, weedy: common throughout Mexico. 536a. Leptochloa filiformis var. latifolia Beetle 536b. Leptochloa filiformis (Lam.)Beauv.var. pulchella Scribn.) Beetle Mexico, native, endemic, mostlv Coastal. 537. L. neallevi Vasey U. S. and Mexico, native. Swales: Tamaulipas, Veracruz and Yucatan; Sinaloa, Nayarit and Jalisco. 538. L. panicoides (Presl)Hitchc. U. S., Mexico and Brazil, native. - Swales: Sonora, Sinaloa, Nayarit, Colima, Guerrero, Morelos, Oaxaca and Tabasco. 539. L. scabra Nees U. S., Mexico, West Indies, south to Brazil, native. Swales: Sonora, Sinaloa, Michoacan, Veracruz, Chiapas and Tabasco. 540. L. uninervia (Presl)Hitchc. and Chase U.S., Mexico, West Indies, south to Argentina, native. Swales: northerm forder states north to Chiapas. 541. L. virgata (L.) Beauv. U. S., Mexico, West Indies, south to Argentina, native. Swales: south to Nayarit, San Luis Potosi and Tamaulipas. Yucatan Peninsula. 254 POH Y oT 0. L,0 CG 1A Vol. 63, No. 4 542. L. viscida (Scribn.) Beal U. S. and northern Mexico, native. Weedy: Baja California Norte, Baja California Sur, Sonora, Sinaloa and Chihuahua. LEPTOCORYPHIUM Nees Paniceae One or two species in tropical America. 543. L. lanatum (HBK) Nees West Indies, Mexico, Central America to Argentina, native. Pine savanna: Chianas, Veracruz and Tabasco. LEPTOLOMA Chase Paniceae Four species, one North America, three in Australia. 544. L. cognatum (Schultes) Chase Canada, U. S. and northern Mexico, native. Sandy soil: Chihuahua, Coahuila, Nuevo Leon, Tamaulipas, Durango, Zacatecas, San Luis Potosi, Guanajuato, Hidalgo and Veracruz. LIMNODEA Dewey Agrostideae Monotypic. 545. L. arkansana(Nutt.) Dewey U.S. and northern Mexico, native. Grassland: Coahuila. LITHACHNE Beauv. Olyreae Four species, tropical America. 546. L. pauciflora (Sw.)Beauv. West Indies and Mexico south to Argentina, native. Tropical forest: Tamaulipas, San Luis Potosi, Veracruz and Chiapas. LOLIUM L. Festuceae Fifteen species, temperate, Old World. 547. L. multiflorum Lam. Introduced. Weedy: scattered localities throughout Mexico. 548. L. perenne L. Introduced. Cultivated and escaped: scattered localities throughout Mexico. 549. Lolium temulentum L. Introduced from the Mediterranean region. Weedy, Baja California Norte, San Luis Potosi, Puebla, Veracruz and Oaxaca. 1987 Beetle, Mexican grasses LUZIOLA Juss. Zizanieae 550. S505 Eight species in tronical America. L. gracillima Prod. Weedy in wet places, Sonora and Jalisco. L. peruviana Gmel. Mexico, Cuba, south to Argentina, native. Marshes: Queretaro, Mexico, Veracruz, Chiapas and Tabasco. LYCURUS HBK Eragrosteae 552). 552b. Six svecies, temverate and subtropical America. L. phleoides HBK var. phleoides U. S. and Mexico, native, Grassland: northern border states south to Chiapas. Lvycurus ovhleoides var. brevifolius Scribn. U.S.A., Mexico and South America. Grasslands, common throughout Mexico except for the Yucatan Peninsula. L. phleoides var. glaucifolius Beal U. S. and Mexico, native. Grassland: Sonora and Chihuahua. MELICA L. Meliceae 5536 554. 554a. 554b. 554¢. 555i 556. S15) 5 Sixty species, temnerate, worldwide. M. frutescens Scribn. California and Baja California, native. 255 Brush: Baja California Norte and Baja California Sur. M. imnerfecta Trin. var. imperfecta. California and Baja California, native. Brush: Baja California Norte and Baja California Sur. M. imperfecta var. flexuosa Beal. M. imperfecta var. minor Scribn. M. imperfecta var. refracta Thurb. M. montezumae Piver (including M. alba Hitchc.) Texas and northern Mexico, native Rocky slopes: Chihuahua, Coahuila and Nuevo Leon. M. nitens (Scribn.) Nutt. U. S. and northern Mexico, native. Woods and grasslands: Coahuila and Nuevo Leon. M. porteri Scribn. U. S. and northern Mexico, native. Grassland: Chihuahua. MELINIS Beauv. Melinideae 553). Fifteen species in Africa. M. minutiflora Beauv. Introduced. Roadsides: Jalisco, Michoacan, Mexico, Hidalgo, Puebla, Veracruz, Oaxaca, Chiapas end Tabasco. MESOSETUM Steud. Paniceae 559). Thirty species, American tronics, mostly Brazil. M. blakei Swallen —~ ; mexico, endemic and Guatemala. Tropical forest margin: Tabasco, Oaxaca and Chiapas. 256 PHY TF 0'L CLG lA Vol. 63, No. 4 560. M. pittieri Hitchc. Mexico and Central America, native. Pine savanna: Oaxaca and Chianas. METCALFIA Conert Festuceae Monotypic. 561. M. mexicana (Scribn.) Conert Mexico, endemic. Pine savanna: Coahuila, Nuevo Leon, San Luis Potosi, Queretaro, Hidalgo, Mexico, Puebla, Oaxaca and Chiapas. MICROCHLOA R. Br. Chlorideae Five or 6 species, tropics, worldwide. 562. M. kunthii Desv. Mexico, Central America and S. America, native. Rocky ground: Baja California Chihuahua south to Chiapas. 563. Miscanthus sinensis Anderson Introduced from China and Japan. Cultivated as an ornamental; Veracruz. 563a. Miscanthus sinensis L. zebrina (Beal) Beetle MONANTHOCHLOE Engelm. Aeluropodeae Three species in the Americas. 564. M. littoralis Engelm. U. S., Cuba and Mexico, native. Baja California Norte, Baja California Sur, Sonora, Sinaloa, Coahuila, Tamaulipas, Chiapas and Yucatan. 564a. Monerma cylindrica (Willd.) Coss & Dur, ntroduced from Europe. Reported from Baja California Norte. MUHLENBERGIA Schreb. Eragrosteae One hundred sixty species, centered in Mexico, but worldwide. 565. M. alamosae Vasey Mexico, endimic. Open slopes: Bajaca California Sur, Sonora, Chihuahua, Sinaloa, Durango, Jalisco , Guanajuato, Mexico, and Morelos. 567. M. angustifolia Swallen Mexico, endemic. Open slopes: Jalisco. 568. M. annua (Vasey) Swallen Mexico, endemic. Open slopes, Chihuahua. 569. M. appressa Gooding Arizona and Mexico, native. Open slopes: Baja California Sur and Baja California Norte. 1987 570. S71 /A Ne 52. S/he 574. SUS 576. 577s Seis Ey) 530. 581, 582. Beetle, Mexican grasses 257 M. arenacea (Buckl.) Hitchc. u. S. and northern Mexico, native. Sandy soils: Sonora, Chihuahua, Coahuila, Zacatecas, and San Luis Potosi. M. arenicola Buckl. U. S. and northern Mexico, native. Sandy soil: Sonora, Chihuahua, Coahuila, Nuevo Leon, Durango, Zacatecas and San Luis Potosi. M. argentea Vasey Mexico, endemic. Open slopes: Chihuahua. M. arizonica Scribn. U. S. and northern Mexico, native. Open slopes: Sonora, Chihuahua,Coahuila, Baja California Sur, Sinaloa and Durango. M. arseni Hitchc. U. S. and northern Mexico, native. Open slopes: Baja California Norte. M. articulata Scribn. Mexico, endemic. Gypsum soils: San Luis Potosi, Guanajuato, Queretaro, Hidalgo and Guerrero. M. biloba Hitchc. Mexico, endemic. Open slopes: Chihuahua and Durango. M. brandegei Reeder Mexico, endemic. Desert: Baja California Sur. M. brevifolia Scribn. Mexico, endemic. Open slopes: Jalisco an Durango. M. brevis Gooding U. S. and Mexico, native. Swales: Chihuahua, Durango, Zacatecas, San Luis Potosi and Mexico. M. breviseta Griseb. Mexico, endemic. Brush: Jalisco, Michoacan, Mexico and Veracruz (type loc.: Orizaba). Muhlenbergia brevivaginata Swallen Mexico, native endemic. Reported only for Jalisco. M. capillaris (Lam.) Trin. U. S., West Indies and Mexico, native. Woodland: Chihuahua, Coahuila, Tamaulipas, Sonora, to Quintana Roo. M. ciliata (HBK) Kunth Mexico, Central America, Ecuador, and Peru, native. Open slopes: Sonora and Chihuahua south to Chiapas M. confusa (Fourn.) Swallen Mexico and Guatemala, native. Open sloves: northern border states south to Chiapas. M. crispiseta Hitchc. Mexico, endemic. Open slopes: Sonora, Chihuahua, Durango, Zacatecas, Nayarit, Jalisco, and San Luis Potosi. 258 PHY T O;¢,0,G 1A Vol. 63, No. 4 586. ™M. curvula Swallen Mexico, endemic. Oak woods: Guanajuato (type loc.) San Luis Potosi, and Jalisco. 587. M. decumbens Swallen Mexico, endemic. Sandy soil: Chihuahua. 588. M. devauperata Scribn. U. S. and Mexico, native. Open slopes: Chihuahua and Coahuila South to Puebla. 589. M. distans Swallen. Mexico and Guatemala, native. Open slopes: Durango, Zacatecas, San Luis Potosi south to Chiapas. 590. M. distichophylla (Presl) Kunth Mexico, endemic. Jalisco south to Chiapas. 591. M. diversiglumis Trin. Mexico, Central America, Colombia and Venezuela to Peru, native. Oak brush: Sinaloa south to Chiapas. 592. M. dubia Fourn. U. S. and Mexico, native. Pine savanna: Chihuahua and Nuevo Leon south to Veracruz and Puebla. 593. M. dubioides Gooding U. S. and northern Mexico, native. Canyons: Coahuila and Chihuahua. 594. M. dumosa Scribn. U. S. and Mexico, native. Cayons, Baja California Sur,Sonora and Chihuahua; south to Oaxaca. 595. M. elongata Scribn. México, endemic. Open slopes: Sonora and Chihuahua; also Mexico acc. to Matuda. 596. M. eludens C. Reeder U. S. and northern Mexico, native. Rocky woods: Sonora, Chihuahua and Durango. 597. M. emersleyi Vasey Southwestern U. S. to Panama, native. Pine-oak forests: northern border states south to Chiapas. 598. M. eriophylla Swallen Mexico, endemic. Oak woods: Mexico. 599. Muhlenbergia expansa (Dc)Trin. United States and Mexico, native. Reported for Veracruz. 600. M. filiformis (Thurb.) Rydb. var. fortis Kelso. U. Sv and Mexico, native. Swales: Baja California Norte and Durango. 601. M. firma Beal Mexico, endemic. Pine savanna: Coahuila, San Luis Potosi south to Oaxaca (tyve loc.). 1987 Beetle, Mexican grasses 259 602. M. flavida Vasey Mexico, endemic. Open slopes: Sonora, Chihuahua, Durango, Jalisco and Hidalgo. 603. M. flaviseta Scribn. Mexico, endemic. Pine savanna: Durango. 604. M. fragilis Swallen U. S. and Mexico, native. Open grond: Northern border states south to Oaxaca y Veracruz. 605. M. gigantea (Fourn.) Hitchc. Mexico, endimic. Pine savanna: Sonora and Durango south to Chiapas. 606. M. glabrata (HBK) Kunth Mexico, endemic. Pine savanna:Nuevo Leon, San Luis Potosi, Guanajuato (type loc.) south to Puebla. 607. M. glauca (Nees) Mez (M. huahuacana Vasey) U. S. And northern Mexico, mative. Pine savanna: Chihuahua an Coahuila south to Mexico. 698. M. goodingii Soderstom Arizona and Sonora, native. Desert: Sonora. 609. M. grandis Vasey Mexico, endemic. Canyon: Nayarit and Jalisco. 610. M. gypsophila Reeder and Reeder Mexico, endemic. Gypsophorous soils: Coahuila, Nuevo Leon and San Luis Potosi. 611. M. hintoni Swallen Mexico, endemic. Central moutains: Mexico. 612. M. implicata (HBK) Kunth Mexico, Central America to Colombia and Venezuela, native. Wet. banks: Chihuahua south to Chiapas. 613. M. iridifolia Soderstrom Mexico, endemic. Pine savanna: Jalisco. 614. Muhlenbergia jaliscana Swallen Mexico, endemic. Pine woods. Reported only for Jalisco. 615. Muhlenbergia laxa Hitchc. Mexico, endemic, wet places Veracruz, "Orizaba". 616. M. leptoura (Piper) Hitchc. Mexico, endemic. Pine savanna: Sonora and Chihuahua. 617. M. lindheimeri, Hitchc. Texas and Coahuila, native. Limestone: Coahuila. 260 POH ¥ T0'L OG 1 A Vol. 63, No. 4 618. M. longiglumis Vasey Mexico, endemic. Pine savanna: Aguascalientes, Jalisco and Michoacan. 619. M. longiligula Hitchc. U. S. and Mexico, native. Canyons: Sonora, Chihuahua, Coahuila, Nuevo Leon and Durango. ; 620. M. lucida Swallen Mexico, endemic. Pine savanna: Chihuahua, Guanajuato and Mexico. 621. M. macrotis (Pipner) Hitchc. Mexico, endemic. Pine savanna: Chihuahua, Sinaloa and Durango south Chiapas. 222. M. macroura (HBK) Hitchc. Mexico and Guatemala, native. Open slopes: Chihuahua and Nuevo Leon south to Chiapas. 624. M. microsperma (DC) Kunth U. S., Mexico, Colombia and Venezuela to Peru, native. Moist banks: south to Chiapas. 625. M. minutissima (Steud.) Swallen U.S. and Mexico, native. Moist banks: Baja California Norte, Chihuahua, and south to Mexico Puebla and Tlaxcala. 626. M. montana (Nutt.) Hitchc. U. S., Mexico and Guatemala, native. Pine savanna: Sonora, Chihuahua, and Coahuila south to Oaxaca. 62:75 \. Mea mMult Cam (RUp I.) Hatcher México, endemic. Pine savanna: Veracruz and Chiapas. 628. M. nigra Hitchc. Mexico and Cuatemala, native. Pine savanna: Guanajuato, Jalisco, Tlaxcala, Puebla Mexico and Chiapas. 629. Muhlenbergia oreophila Swallen Mexico and Guatemala, native. Mexico (type loc. for M. matudae Sohns) 630. M. valmeri Vasey a Mexico, endemic. Pine savanna: Chihuahua and Michoacan. 631. M. parviglumis Vasey U. S., Cuba and Mexico, native. Rocky slopes: Chihuahua, Coahuila, Nuevo Leon Tamaulipas and San Luis Potosi. 632. M. pauciflora Buckl U. S. and northern Mexico, native. Rocky slopes: Baja California Norte, Sonora, Chihuahua, Coahuila and Nuevo Leon. 1987 Beetle, Mexican grasses 261 633. M. pectinata C. Gooding Arizona and Mexico, native. Rocky slopes: Sonora, Chihuahua, Durango, Jalisco, Michoacan, Mexico. 634. M. plumbea (Trin.) Hitchc. Mexico, endemic. Central mountains: Mexico, Puebla and Tlaxcala. 635. M. polycaulis Scribn U. S. and Mexico, native. Rocky slopes: Sonora to Coahuila and south to Hidalgo. 636. M. porteri Scribn. U. S. and Mexico, native. Brush: Baja California Norte, Sonora, Chihuahua, Coahuila Nuevo Leon, Tamaulipas and Durango. 637. M. presliana Hitchc. Mexico and Guatemala, native. Pine savanna: Chiapas. 638. M. pubescens (HBK) Hitchc. Mexico, endemic. Pine savanna: Chihuahua, Coahuila and Nuevo Leon south to Oaxaca. 639. M. pubigluma Swallen Mexico, endemic. Canyons: Chihuahua, Coahuila and Nuevo Leon. 640. M. pulcherrima Scribn. Arizona and Mexico, native. Pine savanna: Chihuahua, Durango and Morelos. 641. M. purpusii Mez Mexico, endemic. Gypsum soils: San Luis Potosi(type loc.) Chihuahua, Coahuila. 642. M. pusilla Steud. Mexico and Guatemala, native. Mountain meadws: Sonora and Chihuahua south to Chiapas. 643. M. quadridentata (HBK) Kunth Mexico and Guatemala, native. Pine savanna: Sonora and Chihuahua south to Chiapas. 643a. M. racemosa (Michx) BSP 644. M. ramulosa (HBK) Swallen ’" MexIco to Costa Rica, native. Meadws: Jalisco and San Luis Potosi, south to Chiapas. 645. M. reederorum Soderstrom Mexico, endemic. Canyons: Durango and Michoacan. 646. M. repens (Presl) Hitchc. U. S. and Mexico, native. Sandy soil: northern border states south to Oaxaca. 647. Muhlenbergia reverchoni Vasey & Scribn. United States and Mexico. Reported from Puebla. 648. M. richardsonis (Trin.) Rydb. Canada south to Mexico, native. Meadows: northern border states south to Puebla. 262 649. 650. 651. G52. 653. 654. 655. 656. 657. 658. 659. 660. 661. 662. 663. 664. 665. POY (TO VEiOiGeT A Vol. 63, No. 4 M. rigens (Benth.) Hitchc. U. S. and Mexico, native. Canyons: northern border states south to Puebla. M. rigida (HKB) Kunth U. S. and Mexico, native. Canyons: northern border states south to Oaxaca. M. robusta (Fourn.) Hitchc. Mexico, endemic. Oak-pine savanna: Sonora and Chihuahua south to Chiapas. M. schmitzii Hack. Mexico, endemic. Pine savanna: Chihuahua, Hidalgo and Mexico. M. schreberi Gmel. U. S. and Mexico, native. Woods: Queretaro, Hidalgo and Veracruz. M. scoparia Vasey Mexico, endemic. Pine savanna: Sonora and Chihuahua south to Michoacan M. seatoni Scribn. Mexico, endemic. ‘Pine savanna: Puebla and Tlaxcala. M. setariodes Fourn. Mexico and Central America, native. Pine savanna: Veracruz and Chiapas. M. setifolia Vasey Texas, New Mexico and northern Mexico, native. Calcareous soil: Chihuahua, Tamaulipas and San Luis Potosi. M. shepherdi (Vasey) Swallen Mexico, endemic. Ledges: Sonora, Chihuahua and Durango. M. sinuosa Swallen U. S. and Mexico, native. Mois slopes: Sonora and Chihuahua. M. speciosa Vasey Mexico, endemic. Pine savanna: Chihuahua south to Puebla. M. spiciformis Trin. Mexico, endemic. Pine savanna: San Luis Potosi, Michoacan, Hidalgo, Veracruz, Oaxaca and Chiapas. M. stricta (Presl) Kunth Mexico, endemic. Pine savanna: Durango, Nayarit, Jalisco, Colima, Michoacan, Mexico, Molrelos, Veracruz and Oaxaca. M. strictior Scribn. Mexico, endemic, Pine savanna: Sonora, Chihuahua, Durango and Mexico. M. subaristata Swallen Mexico, endemic. Canyon: Durango. M. subbiflora Hitchc. Mexico, endemic. Moist slopes: Durango. 1987 666. 667. 668. 669. 670. 671. 672. 673). 674. G7 dy. 676. Tr 678. 67/9); 680. Beetle, Mexican grasses 263 M. tenella (HBK) Trin Mexico to Panama Moist slopes: Sonora and Chihuahua south to Chiapas. M. tenuifolia (HBK) Trin. Mexico, endemic. M. tenuissima (Presl) Kunth Mexico and Panama, native. Moist slopes: Nayarit, Jalisco, Colima and Michoacan M. torreyi (Kunth) Hitchc U. S. and northern Mexico, native. Grasslands: Sonora, Chihuahua and San Luis Potosi. Me Meelis! (Torres) Hiteche. U. S. and Mexico, native. Grassland: Chihuahua, Coahuila, Durango, San Luis Potosi, Hidalgo and Mexico. M. vaginata Swallen Mexico and Guatemala, native. Meadows: Sinaloa, Durango, Guanajuato, Michoacan, Mexico and Hidalgo. M. versicolor Swallen Mexico, endemic. Pine savanna: Nayarit south to Chiapas. Me ve ploras Hatcher Mexico, endemic. Gypsum soils: Coahuila, Nuevo Leon, Durango, Zacatecas, San Luis Potosi and Hidalgo. M. virescens (HBK) Kunth New Mexico and Arizona south to Mexico, native. Canyons: northern border states south to Michoacan and Mexico. M. virletii (Fourn.) Soderstrom Mexico, endemic. High plains: Durango south to Puebla. M. watsoniana Hitchc. Mexico, endemic. Desrt: Chihuahua, San Luis Potosi, and Jalisco. M. wolfii (Vasey) Rydb. U. S. and Mexico, native. Pine savanna: Baja California Norte, Baja California Sur, Sonora, Chihuahua and Durango. M. wrightii Vasey U. S. and Mexico, native. Grasslands: Baja California Norte, Sonora and Chihuahua, also Tamaulipas. M. xanthodos Soderstrom Mexico, endemic. Pine savanna: Chiapas. Muhlenbergia xerovhila C. O. Gooding Arizona and adjacent Mexico, native. Reported from Sonora. MUNROA Torr. Eragrosteae 681. One species in western N. America and two in Argentina. M. sSquarrosa (Nutt.) Torr. Canada, U. S. and Mexico, native. Grassland: Chihuahua and Coahuila. 264 PH Y T OvEVOuGRiea Vol. 63, No. 4 NEYRAUDIA Hook. f. Arundineae Small, Old World genus. 682. N. reynaudiana (Kunth) Keng. Introduced. Cultivated or escaped: Veracruz. OLMECA Soderstrom Two species in Mexico. 632a. O. recta Soderstrom Native, endemic. Veracruz. 682b. O. reflexa Soderstrom Native, endemic. Veracruz, Oaxaca and Chiapas. OLYRA L. Olyreae 25 species in the American tropics, one in Africa. 683. 0. cordifolia HBK Southern Mexico, Venezuela to Paraguay, native. Tropical forest margin: Chiapas, also San Luis Potosi. 684. O. latifolia L. Florida, Mexico and West Indies south to Brazil and northern Argentina, native. Tropical forest margin: Nayarit, San Luis Potosi south to the Yucatan Peninsula. 685. O. yucatana Chase Mexico, Guatemala and British Honduras, native. Tropical forest margin: Oaxaca, Chiapas and the Yucatan Peninsula. OPIZIA Presl Chlorideae Monotypic 686. O. bracteata McVaugh . and from michoacana 686a. O. stolonifera Presl Cuba and Mexico, native. Dry slopes and flats: Nayarit, Tamaulipas south to Yucatan. OPLISMENUS Beauv. Paniaceae Ten species, tropics, wordwide. 687. O. affinis Schult. Pantropical, introduced? Weedy: Sonora and Baja California Sur south to the Yucatan. 687a. O. affinis var. humboldtianus (Nees) Scholz 688. O. hirtellus (L.) Beauv. Mexico and West Indies south to Argentina, native. Forest margin: northern border states south to Yucatan Peninsula. 689. O. rariflorus Presl Southern Mexico south to Peru, native. Pine savanna: Sinaloa south to Chiapas. 1987 Beetle, Mexican grasses 265 690. O. setarius (Lam.) R. & S. U. S., West Indies and Mexico south to northern Argentina, native. Forests: Nuevo Leon an Nayarit and to Quintana Roo. ORCUTTIA Vasey Festuceae Five species in California and Baja California. 691. QO. californica Vasey California and Baja California, native. Vernal pools: Baja Norte 692. O. fragilis Swallen Mexico, endemic. Vernal pool: Baja Sur ORTHOCLADA Beauv. Eragrosteae Monotyvic. 693. O. laxa (L. Rich.) Beauv. Mexico, south to Peru and Brazil, native. Tropical forest margin: Oaxaca, Veracruz, Chiapas and Tabasco. ORYZA L. Ory zeae Twenty-nine species, pantropical. 694. O. alta Swallen Mexico to Paraguay, native. Aquatic: Tabasco, Chiapas and Campeche 695. O. latifolia Desv. Mexico and West Indies south to Paraguay and Brazil, native. Aquatic: Sinaloa, Nayarit, Jalisco, Colima, Oaxaca, and Campeche 696. O. perennis Moench. West Indies, Mexico, Brazil, native. Aquatic: Oaxaca, Vercruz and Chiapas. 697. O. sativa L. Introduced. Cultivated: ORYZOPSIS Michx. Stipeae ‘Twenty species, temperate, worldwide. 698. O. florulenta-vilger Mexico to Colombia, native. Central mountains: Mexico, Tlaxcala and Puebla. 699. O. hymenoides (R. & S.) Ricker U. S. and Mexico, native. Sandy soil: Baja Norte. OTATEA (McClure & Calderon & Soderstrom to species in Mexico. 699a. O. acuminata (Munro) Calderon & Soderstrom Central Mexico, endemic. 699b, O. fimbriata Calderon & Sorderstrom Central and Mexico, endemic. 266 PHY TOLOG TA Vol. 63, No. 4 PANICUM L. Paniceae Probable 500 species, pantropical and subtropical, ocasionable temperate. 700. P. aciculare Desv. var. aciculare United States, Mexico, Honduras and northern South America. Nativa, grasslands. 700a. P. aciculare var. arenicoloides (Ashe) Beetle 700b. P. aciculare var. ovinum (Scribn. & Smith) Beetle Reported from Veracruz. 701. P. acuminatum Sw. var. acuminatum. 70la. BP. acuminatum var. implicatum (Scribn.) Beetle 701b. P. acuminatum var. lindheimeri (Nash) Fernald 7O0ic. = acuminatum var. pilosum (Torr.) Beetle 702: (Dichanthelium) albomaculatum Scribn. Hoxics and Guatemala, native. Rocky hills: Jalisco, Michoacan and Mexico also Nuevo Leon Tamaulipas and Hidalgo. 703. P. altum Hitchc. & Chase Mexico to Panama, native. Reported from Quintana Roo. 704, P. amarulum Hitchc. (# Virgata) U. 5., Bahama, Cuba and Mexico, native. Coastal dunes: Veracruz, Chianas, Tabasco, Camoeche and Quintana Roo. 705. P, amarum Ell. (# Virgata) U. S. and West Indies and Mexico. Coastal dunes: "eastern coast of Mexico" acc. to Gould: Grasses of Texas. 706. P. antidotale Retz. (# Maxima) Introdute Cultivated and escaved: scattered localities throughout Mexico. 1987 Or 708. 709. 710. Vite aa’ WBE Beetle, Mexican grasses 267 P. arundinariae Trin. (# Parviglumia) Mexico to Panama, native. "Southern Mexico" according to Swallen: Grasses of Gautemala. Also reported for San Luis Potosi by Hernandez-X; Vercruz. Veracruz, Oaxaca, Chiapas. P. bartlettii Swallen (# Trichoidea) Mexico, Guatemala and British Honduras, native. Moist woods: San Luis Potosi, Hidalgo, Veracruz, Oaxaca, Chiapas, Campeche and Quintana Roo. P. biglandulare Scribn. & Smith (# Stolonifera) Mexico and Guatemala, native. Pine-oak savanna: Guerrero, Oaxaca and Chiavas. DP, boliviense Hack. Mexico to Argentina, native. Woods: Tamaulipas, San Luis Potosi, Veracruz, Oaxaca, Chiapas and Tabasco. P. breviramosum Swallen Mexico and Guatemala, native. Pine savanna: Chiapas, acc. to Gould. P. bulbosum HBK (# Maxima) U. S., Mexico, Guatemala, Colombia, Ecuador, native. Brush: throughout Mexico excep for Baja California Norte and the Yucatan Peninsula. P. (Dichanthelium) caerulescens Hack. U. S., Cuba and Mexico, native. Marshes: Quintana Roo. P. capillare L. (# Capillaria) U. S. and Mexico, native. Weedy:northermborder states, Baja California Norte, Sonora and Coahuila. P. capillarioides Vasey (# Difusa) Grassland: Texas and northern Mexico, native. Sandy soil: Nuevo Leon, Tamaulipas and San Luis Potosi. 268 PHYTOLOGIA Vol. 63, No. 4 716. P. cayennense Lam. (# Canillaria) Mexico and West Indies to Panama and northem S. America, Native. Oaxaca, Veracruz and Campeche. 717. Panicum clivum Sohns Mexico, native, endemic. States of Hidalgo and Mexico. 718. BP. coloratum L. Introduced. Cultivated: Sonora, Coahuila. 719. BP. (Dichanthelium) commutatum Schult. U. S. and Mexico. Brush: Tamaulinas, Jalisco south to Chiapas. 719a. = camutatum var. joorii (Vasey) Fernald 720. cordovense Fourn. — endemic. Brush: Veracruz, Hidaloo. 721. Panicum craterifemm Sohns. Mexico, native, endemic. Known only fram Guerrero. 722. P. cupreum Hitchc. & Chase (# Laxa) Mexico, endemic. Brush: Mexico (tyne loc.) and Durango. 123); P. cyanescens Nees Mexico, Central America to Peru and Brazil, native. Stream bottoms: Chiapas and Tabasco. 724. P. decolorans HBK (# Canillaria) Mexico and Central America, native. Brush: Chihuahua, Sinaloa, Coahuila, San Luis Potosi, south to Chiapas. 724a. Panicum decolorans EBK var. _Darcum (Hitchc. & Chase) Beetle Mexico, native, endemic. Central states. 725. Panicum dichotomum L. var. dichotomum United States and Mexico, native & various Island of the Carribean. Woods, wet sandy soils, Hidalgo south to Chiapas. 725a. Panicum dichotamum var. tenue (Muhl.) Beetle Dichanthelium dichotomm var. tenue (Muhl.) Gould and Clark, Ann. Missouri Bot. Gard 65:1119. 1878. pera plain of eastern United States and west to Texas and southern Mexico. Reported from Chiapas. P. dichotamiflorum Michx. (# Dichotomiflora) Canada, U.S., and West Indies, Mexico, native. Weedy: Chihuahua to Colima. 726. Panicum diffusum Sw. (# Diffusa) U.S. and West Indies, Mexico to Brazil, native. Weedy: Coahuila, Nuevo Leon and Tamaulinas. 727. ©. (Dichanthelium) ensifolium Baldw. U. Sv and Mexico, native. Bogs: Chihuahua, Hidalgo and Chianas. 728. Panicum filipes Scribn. Texas and Mexico, native. Grasslands, Coahuila, Nuevo Leon and Tamaulinas south to Oaxaca. 729. ~P. frondescens Meyer (# Stolonifera) Mexico and Central America to northern Araentina, native. Woods: Veracruz, Oaxaca, Chianas and Tabasco. 1987 730. gsi W3Ze 733% 734. 135% 736. 737. 738. 738a 738b 738c 139). 740. 741. 742. 743. Beetle, Mexican grasses P. ghiesbreohtii Fourn. (# Diffusa) U.S., West Indies, Cuba and Mexico to northern South America, native. Brush: Throughout Mexico, excent for Baja California. P. glutinosum Sw. Mexico, West Indies to Argentina, native. Forests: Hidalao south to Chianas. P. gouinii Fourn. (# Viragata) U. S. and Mexico, native. Coastal: Veracruz. P. hallii Vasey (# Diffusa) U. S. and Mexico, native. Clav soils: northern border states (except for Baja Calif.) south to Hidalao. P, havardii Vasey (# Viraata) U. S. and Mexico, native. Sandy soil: Chihuahua. P. hians Ell. (# Laxa) U. S. amd Mexico, south to Panama, native. Coastal plain: Tamaulinas end Veracruz; also Nayarit, Jalisco, Guanajuato and Michoacan. P. hintoni Swallen Mexico endemic. Central mountains: Mexico (tyne loc.). P. hirsutum Sw. (# Diffusa) West Indies and Mexico south to Brazil, native. Sandy soil: Colima, San Iuis Potosi south to Campeche. P. hirticaule Presl (# Capillaria Var. hirticaule U. S. Cuba and Mexico south to Arcentina, native. Weedy: common throuchout Mexicc. Panicum hirticaule var. miliaceum (Vasey) Beetle United States and Mexico Common in Baja California and Sonora. Panicum hirticaule var. nampinosum (Fitchc. & Chase) Beetle Mexico and Guatemala, native. Sonora and Chihuahua south to Oaxaca. Panicum hirticaule var. stramineum (Hitchc. & Chase) Beetle U.S. and Mexico, native. Sonora, Sinaloa, Durango, Nuevo Leon, Nayarit, Michoacan and Guerrero. Panicum hirtum Lam. “Mexico and Belice south to South America. Native, reported from Oaxaca and Chiapas. P, ichnanthoides Fourn. Mexico and Central America, native. Dry slopes: Jalisco, Quer6taro, Puebla, Veracruz amd Chiapes. P. (Dichanthelium) laxiflorum Lam. U. A. and Mexico, native. Nuevo Ieon and Tamaulivas south to Chianas. P. laxum Sw. West Indies, Mexico, Central America to Paracuay, native. Coastal brush Sinaloa south to Chianas; Tamaulipas and San Luis Potosi south to Yucatan. P. lepidulum Hitchc. & Chase (#Diffusa) Mexico amd Guatenala, native. Weedv: Sonora, Chihuahua and Coahuila south to Chiapas. 269 748. 749. 750. 751. 751la. 754. 755. 756. 751. 758. PW Y Tf OG G.I A Vol. 63, No. 4 Panicum leucoblepharis Trin. var. nmubescens (Vasev) Beetle United States and Mexico; Wet denressions, Veracruz, Oaxaca, anc Chianas. P. lonoum Hitchc. and Chase (# Laxa) Mexico, endemic. Brush: Veracruz (tyne loc.). Panicum macrospemmum Gould Mexico, native, ememic. Rerorted from San Luis Potosi and Queretaro. P. maximum Jacg. (# Maxima) Introduced. Cultivated and escaned: cormon throuchout Mexico excent for Baja California. P. mertensii Roth Mexico and Central America south to Arcentina, native. Swamps: Tabasco, Jalisco. P. (Dichanthelium) nodatum Hitchc. and Chase Texas and Tamaulinas, native. Grassland: Tamaulipas. P. obtusum HBK U. S. and Mexioo, native. Grasslands: northern border states south to Puebla. P. oligosanthes Schult. U. S. to northern Mexico, native. Woodlands: Panicum oligosanthes var. scribnerianum (Mash) Fernald U. A. and Mexico, native. Brush: Sonora, Chihuahua and Coahuila. P. olivaceum Hitchc. & Chase Mexico, Central America and Venezuela. Pine savanna: Veracruz, Oaxaca end Chianas. Panicun ovale Elliott var. addisonii (Nash) Eastern United States and Mexico. Cormon in oak or oak-pine forest, Nuevo Leon and Tamaulinas south to Chiares. Dichanthelium ovale (Elliott) Gould and Clark Var. addisonii (Nash) Gould land Clark, Brittonia 32:360.1980. P. parviglume Hack. (#Parviglumia) Mexico, Guatemala, British Honduras and Costa Pica, Native. Pine-oak savanna: Veracruz and Chianas. P. (Dichanthelium) vedicellatum Vasey Texas and Mexico, native. Limestone: Coahuila, Nuevo Leon, Tamaulinas and Hidalao. P. pilcana Hack. ee eee : Sr. Texas and N.E. Mexico). Cultivated: Nuevo Leon. P. pilosum Sw. (# Laxa) Mexico West Indies to Argentina, native. Swamps: Nayarit, San Luis Potosi, Veracruz, Guerrero, Oaxaca. Chiapas, Tabasco and Quintana Roo. P. plenum Hitchc. & Chase (# Maxima) U. S. and Mexico, native. Grassland: Sonora, Chihuahua, Coahuila, Duranco, Guerrero and Veracruz. 1987 159% 760. 161. 762. 763. 764. 765. 766. 767. 768. 769. 770. alee CA VSR 774. WD. Beetle, Mexican grasses P. polygonatum Scharad. (# Laxa). Southern Mexico to Paraguay, native. Stream bottoms: Veracruz, Oaxaca and Chiapas. P. (Dichanthelium) pseudopubescens Nash U. S., Mexico and Guatemala, native. Oak-savanna: San Luis Potosi. P. rulchellum Raddi (# Stolonifera) Mexico, British Onduras, Martinique to Bolivia and Brazil, native. Streambottoms: Veracruz, Oaxaca and Chianas. Peak L. Introduced form Eurore. Reported from Tamaulinas and Veracruz. P. rigidulum Nees (# Agrostoidea) U. S., West Indies and lfexico, native. Streambanks: Coahuila, Veracruz and Chianas. P. rudgei R. & S. Mexico, Jamaica and Trinidad to Bolivia and Brazil, native. Brush: Tabasco. P. rugulosim Trin. Mexico to Brazil, native. Swamps: Veracruz, Oaxaca, Chiarnas and Tabasco. P. sabulorum var. patulum (Scribn. & Merr.). P. nashianum var. natulum Scribn. & Merr. P. schiffneri Hack. (# Parviqlumia) Mexico to Brazil, native. Streambanks: Veracruz, San Luis Potosi, Guanajuato, Mexico and Chiapas. P. schmitzii Hack. (# Parviclumia) Mexico, endemic. Brush: San Luis Potosi, Veracruz and Oaxaca. P. sellowii Nees Mexico and West Indies south to Paracuay and Argentina, native. Brush: Veracruz, Oaxaca, Chiamas and Tabasco. P. (Dichanthelium soha m Ell. U. S. and Mexico south to Venezuela, native. Open slopes: northern border states south to Chianas. P. stagnatile Hitchc. and Chase Mexico and Central America, native. Swamps: Veracruz, Chianas and Tabasco. P. stoloniferum Poir. (# Stolonifera) Southern Mexico and Guatemala south to Araentina, native. Swamps: Chiapas. P. sucocosum Hitchc. and Chase Mexico, endemic. Ponds: Jalisco (tyne loc.) and Mexico. Panicum tenerum Beyr. Southeastern U. S., West Indies, Belize. Reported fram Quintana Roo. P. transiens Swallen Mexico and Guatemala, native. Mountain mesa: Nuevo Leon and Tamaulinas (tyne loc.); also Hidalgo. 271 272 776. 777. 782. PHY st (0 LxOvGii A Vol. 63, No. P. trichanthum Nees (# Trichoidea) Mexico and West Indies to Paraguay, native. Swamps: Colima, San Luis Potosi, Veracruz, Caxaca, Campeche and Quintana Roo. P. trichoides Swartz (# Trichoidea) Mexico and West Indies South to Peru and Brazil. . Coastal rmlain, thickets: Sonora south to Chiaras; San Luis Potosi south to Yucatan. Panicum turckheimii Hack. Mexico and Guatemala Reported from Chiapas (Palenque). P. vaseyanum Scribn. (# Dichotomiflora) Mexico, endemic. Brush: Chihuahua, Aguascalientes, Jalisco and Michoacan. P. venosum Swallen Mexico, endenic. Ditch: Michoacan (tvre loc.) P. virgatum L. (# Viroata) U. S. and Mexico, native. Grassland: Sonora, Bfja California Sur, Chihuahua, Jalisco, Nayarit, Oaxaca and Chianas. P. (Dichanthelium) viscidellum Scribn. Mexico, Central America and Colombia, native. Pine savanna: Veracruz. PAPPOPHORUM Schreb. Papnonhoreae 783. 784. 785. 786. Ten species in the Americas. P. bicolor fourn. U.S. and Mexico, native. Grassland: Chihuahua, Coahuila, Nuevo Leon, Tamaulinas, San Luis Potosi, Mexico and Veracruz. P. paopiferum (Lam.) Kuntze Mexico and South America, native. Brush: Veracruz and Oaxaca. P. subbulbosum Arech. Mexico and South America, native. Brush: Sonora. P. vaginatum Buckl. U. S. and Mexico, native. Brush: Sonora, Chihuahua, Coahuila, Nuevo Leon, Durango and Zacatecas. 4 1987 Beetle, Mexican grasses 273 PARAPHOLIS Hubb. Monerneae Five or 6 species, Old World. 787. P. incurvus (L.) C. E. Hubb. Introduced. Sea marshes: Baja Norte. PASPALIDIUM Stapf Paniceae Five or 6 species, Old World. 788. F. geminatum (Forsk.) Stapf var. geminatum see mmea erry Aquatic: scattered localities throughout Mexico. 788a. P. geminatum var. paludivagum (Hitchc. and Chase) Gou U. S. and Mexico south to Araentina, native. Swamps: Jalisco and Michoacan; also Tamaulinas PASPALUM L. Paniceae Four hundred species in tronics and subtropics, ’ worldwide. 789. BP. acuminatum Raddi U. S. and Mexico south to Argentina, native. Aquatic: Michoacan and Morelos. 790. Paspalum acutum Chase Mexico and Panama, native. Irrigation ditches, reported from Veracruz. 791. P. adoperiens (Fourn.) Chase Mexico and Central America, native. Aquatic: Veracruz (type loc.) Qaxaca and Chianas. 792. P. affine Steud. Mexico and Guatemala, native. Swamps: Veracruz, Chiaras, Hidalgo and Oaxaca. 793. P. alcalinum Mez Mexico to Paraguay and Argentina, native. Swamps: San Luis Potosi (type loc.), Jalisco amd Chianas 794. P. arsenei Chase Mexico, endemic. Mountain slopes: Acuascalientes and San Luis Potosi : south to Oaxaca. 795. P. blodgettii Chapm. -U. S., Mexico, West Indies and Central America, native. Yucatan Peninsula 274 796. 797. 798. 799 800. 800a. 800b. 800c. 801. 802. 803. 804. 805. 806. 807. PHYTOLOGIA Vol. 63, No. P. botterii (Fourn.) Chase Mexico and Central America, native. Swamps: Sonora south to Chiapas, Nuevo Leon south to Yucatan. P. caespitosum Flugge U. S., West indies, Mexico and Central America, native. Pine savanna: Tamaulipas and Nuevo Leon, Veracruz, Chiapas, Tabasco and the Yucatan Peninsula. P. candidum (Humb. & Bonpl.) Kunth Southern Mexico to Chile, native. Weedy: Michoacan, Morelos, Guerrero, Veracruz and Chianas. P, clavulifernm Wright Mexico and Central America to Brazil, native. Brush: Sinaloa, Nayarit, Jalisco, Colima, Oaxaca, Chiapas and Yucatan. P. conjugatum Bergius U. S. and Mexico to Argentina, native. Tropical forest margin: Sinaloa south to Chianas; Nuevo Leon and Tamaulipas south to Quintana Poo. P. conjugatum var. parviflorum Doell. Coastal: same distribution as the species. P. conjugatum var. pubescens Doell. Mexico to Brazil, native. Tropical forest margin: San Luis Potosi, Veracruz, Hidalgo, Chiapas and Cammeche. Paspalum conjugatum forma tristachya (Vandervst) Beetle Scattered tions throughout the range of the species. In Mexico reported from Jalisco, Guerrero, Tabasco and Oaxaca. P. sum Schrad. Southern Mexico to Argentina, native. Marshes: Nayarit, jalisco and Morelos, San Luis Potosi, Veracruz and Oaxaca. P. convexum H. & B. Mexico, Carribean to Brazil, native. Oak brush: Sonora and Chihuahua south to Chianas. FP. crassum Chase Mexico, endemic. Brush: Jalisco, Colima (tyne loc.), Michoacan and Mexico, also Guerrero and Chianas. P. crinitum Chase Mexico, endemic. Brush: Coahuila and Nuevo Leon, San Inmis Potosi (type loc.), Jalisco and Puebla. P. culiacanum Vasey Mexico, endemic. Mountains: Sinaloa. P. Cymbi forme Fourn. Mexico Central America, native. Brush: Mexico (type loc.) and Yeracruz, Oaxaca, Chianas. Paspalum densum Poir. West Indies, Panama, Brazil and Bolivia, Native. Reported for Isla Guadalure. 816a 817. 818. 819. 820. 821. 822. Beetle, Mexican grasses 275 P. dilatatum Poir. Introduced, Ditches Veracruz, Michoacan, Baja California Norte. P. distichum L. U. S. and West Indies south to Argentina, native. Ditches: Throughout Mexico excent for the Yucatan Peninsula. P. erectum Chase Mexico, endemic. Colima (type loc.), and Jalisco. P. fasciculatum Willd. Mexico south to Argentina, native. Swamps: Veracruz, Oaxaca, Chiapas, Tabasco and Campeche. P. fimbriatum HBK Introduced? Weedy: Yucatan and Quintana Roo, Baja California (Isla Guadalupe) P, anarum Beetle Mexico, emic. Ledges: Sinaloa (type loc.) and Nayarit. P. hartwegianum Fourn. Sou’ Texas and Mexico, native. Ditches: scattered localities throughout Mexico except Baja California. P. heterotrichon Trin. Mexico south to Brazil, native. Ditches: Chianas. P. humboldtianum Fluoge var. humboldtianum Mexico to Panama, western South America to Argentina, native. Pine forests: throughout Mexico except for the Yucatan Peninsula and Baja California. Paspalum humboldtianum Fluage var. elegantissima Beetle Mexico, emic. Pine or oak woods, Jalisco, Michoacan, Guerrero, Morelos, and Colima. Paspalum intermedium M Mexico south to Peru, native. Swamps: Nararit, Veracruz (tyne loc. for P. plenum) and Tabasco. i P. jaliscanum Chase Mexico to Guatemala, native. Pine forests: Nayarit, Jalisco (tyne loc.), Michoacan, San Luis Potosi, Puebla, Veracruz and Chianas. P. langei (Fourn.) Nash U. S., West Indies and Central America, native. Brush: lowland areas of Mexico (absent from the central plateau and Baja California). P. lentiginosum Presl Mexico Guatemala, native. Swamps: Baja California Sur, Sonora, south to Chiapas; also Yucatan. P. leptachne Chase Mexico, endemic. Brush: Nayarit (type loc.). Paspalum lineare Trin. Mexico, West Indies, Central America and Argentina, native. Reported fram Chianas and Tabasco. 276 823. 824. 825, 826. 827. 828. 829. 830. 831. 832. 833. 834. 835. 836. 837. 838. P HUY oT Os 0yGeT A Vol. 63, No. P. lividum Trin. U. S. and West Indies south to Argentina, native. Ditches: scattered localities throughout Mexico excent for the Yucatan Penincula and Baja California P. longi Nash Mexico, ic. Swamps: west coast, Nayarit south to Oaxaca (tyoe loc. Jalisco), also Campeche. Native, endemic. Described from Socorro Island, Colima. Paspalum luxurlans Guzman and Rico Mexico, native, endemic. Revorted from Jalisco and Nayarit. P. mala lum Trin. Mexico south to Argentina, native. Swamps: San Luis Potosi, Veracruz, Chiapas and Yucatan. P. mayanum Chase Mexico, endemic. Swamps: Yucatan (tyne loc.) and Chiapas. Paspalum melano: Desv. Mexico to Brazil, native. Collected on Isla Guadalupe. P. millegrana Schrad. Mexico south to Brazil, native. Swamps: Nayarit, Tamaulipas, Veracruz, Chiapas, Tabasco and Yucatan. P. minus Fourn. Texas, West Indies south to Paraguay, native. Swamps: Guanajuato, Michoacan, Oaxaca, Veracruz, Chiapas and Tabasco. P. lum monostachyum Vasev i SSS U.S. and Mexico, native. Reported for Veracruz. P. multicaule Poir. Mexico, West Indies to Brazil, native. Pine woods: Veracruz, Nayarit, Mexico and Oaxaca Brush: Chiapas (type loc.), Nayarit. P. notatum Flugge U. S. (where introduced?), Mexico, West Indies south to Araentina (where native?). Swamps: Lowland, Mexico except for Baja California. Paspalum nutans Lam. Mexico to Brazil, native. Collected on Isla Guadalupe. P. orbiculatum Poir Mexico and West Indies south to Paraguay, native. 4 Swamps: Sinaloa, San Luis Potosi, Guerrero, Veracruz, Chiapas and Tabasco. 1987 839. 840. 841. 842. 843. 844, 844a. 845a. 845b. 846. 847. 848. Cie 850. 851". 85la. 851b. Beetle, Mexican grasses P. palmeri Chase Mexico endemic. Brush: Sonora (type loc.) P, paniculatim L. Mexico and West Indies south to Argentina, native. Swamps: Sonora & Baja California Sur south to Chianas and eastern San Luis Potosi southward through the Yucatan Peninsula. P. paucispicatum Vasey Mexico, endemic. Brush: Sonora, Chihuahua and Nuevo Leon south to Oaxaca; also Campeche. P. vectinatum Nees Mexico to southern Brazil, native. Pine woods: Sinaloa south to Tabasco. P. nlicatulum Michx. U.S West Indies south to Argentina, native. Thickets: Sinaloa south to Chiapas, Nuevo Leon and Tamaulipas south to Tabasco. P. prostratum Scribn. & Merr. Mexico, emic. Southern Durango south to Chiapas. P lum prostratum var. pygmaeum Scribn & Merr. P. iflorum Rupr. var. arsenni We BF and Mexico, native. Swamps: northern border states south to Oaxaca and Veracruz. P. vubiflorum var. tinctum Paspalum pulchellum Kunth Mexico a West Indies south to Brazil, native. reported from Oaxaca. Paspalum pygmaeum Hack. Mexico, Peru, Venezuela, Bolivia and Argentina, native. Reported Fram Jalisco, Michoacan, Morelos and Veracruz. Peane Bergius Sou Mexico to northern Argentina, native. ~ Aguatic: Tabasco, Chiapas and Campeche. Paspalum saccharoides Nees Mexico Central America south to Bolivia, native. Collected on the Isla Guadalupe. Paspalum secans Hitchc. & Chase Mexico and West Indies, native. Collected on Isla Guadalupe. P. setaceum Michx. var. setaceum U. S. and Mexico, native. Tamaulipas, Veracruz and Chianas. P. setaceum Michx. var. ciliatifolium (Michx.) Vasey U. S. and Mexico, native. Chihuahua and Veracruz. Paspalum setaceum Michx. var. dispar Guzman Mexico, native, endemic. Reported from Oaxaca. 277 278 PHY TOLOGILIA Vol. 63, No. 4 852. P. sparsum Chase Mexico, endemic. Brush: Yucatan and Campeche. 853. P. squamulatum Fourn. Mexicd and Central America, native. Qak woods: Baja Sur and Sinaloa south to Chiapas. 854. P. stellatm Humb. & Bonpl. Southern Mexico to Argentina, native. Sandy soils, pine woods: Oaxaca and Chiapas. 855. P. tenellum Willd. Mexico south to Brazil, native. Sonora, Aguascalientes, Jalisco, Michoacan, Mexico, Morelos and Chiapas. 856. P. tinctum Chase Mexico and Guatemala, native. Nayarit, Jalisco, Guanajuato (type loc.), Michoacan and Morelos. 857. P. umbratile Chase Mexico and Central America, native. Shady Banks: Nuevo Leon, Tamaulipas, Veracruz and Yucatan. 858. P. unispicatum (Scribn. & Merr.) Nash Mexico, emic. Jalisco, Nuevo Leon, Tamaulinas south to Chiapas (type loc.: Oaxaca). 859. P. urvillei Steud. Introduced. Nuevo Leon, Aguascalientes, Michoacan & Mexico. 860. P. vaginatum Sw. U. S., West Indies south to Argentina and Chile, native. Coastal: Baja California Sur, Baja California Norte, Sonora, south to Chiapas; Tamaulipas south to Yucatan Peninsula. 861. P. variabile (Fourn.) Nash Mexico, endemic. Wood: Nuevo Leon and Tamaulinas south to Veracruz and Chiapas. 862. P. virgatm L GO: Se a West Indies south to Arcentina, native. Swamps: Nayarit, Jalisco, San Luis Potosi and Tamaulipas south to Chiapas and Tabasco. 863. P. virletii Fourn; Brush: Sonora, San Luis Potosi (tvpe loc.) and Veracruz. PENNISETUM L. Paniceae Eighty species, tronics and subtronics, worldwide. 864. P. abericanum (L.). Schum. Introduced and cultivated. Reported from Nuevo Leon and Mexico. 865. P. bambusiforme (Fourn.) Hemsl. Mexico and Central America south to Peru, native. Dry banks: San Luis Potosi, Michoacan, Veracruz (type loc.), Hidalgo, Oaxaca and Chiapas. 1987 866. 867. 868. 869. 870. 871. 872. 873. 874. 875. 876. Beetle, Mexican grasses 279 P. clandestinum Host. Introduced. Commonly cultivated: Throughout Mexico. P. camlanatum (Nees) Hemsl. Mexico and Central America, native. Pine-oak forests: Sinaloa, south to the Yucatan Peninsula. P. crinitum (HBK) Spreng. Mexico, endemic. Brush: Sinaloa, Durango, San Luis Potosi, Nayarit, Jalisco, Guanajuato, Michoacan, Mexico, Veracruz and Guerrero. P. distachyum (Fourn.) Pupr. Guerrero, Puebla, Mexico and Central America, native. Veracruz and Oaxaca. P. durum Beal including P. pringlei Leeke) Mexico, endemic. Chihuahua. P. nervosum (Nees) Trin. Mexico, Guatemala, Ecuador and Brazil to Argentina, native. San Luis Potosi, Veracruz and Chiapas. P. orientale Rich. Introduced, cultivated, cardens, throughout Mexico. Pennisetum prolificum Chase Mexico, endemic. Tropical forest marain: Veracruz (tyne loc.), Guerrero and Oaxaca. P. purpureum Schum. Introduced. Cultivated and escaped: Nuevo Leon south to Chiamas and Tabasco. P. setosum (Sw.) L. Rich. U.S., West Indies and Mexico south to Bolivia and Brazil, native. Pine woods: Sinaloa south to Chianas; Tamaulipas, Veracruz, Mexico. P. villosum R. Br. Introduced. Cultivated: Sinaloa, Durango, Coahuila, Nuevo Leon, Zacatecas, Aguascalientes, Jalisco, Mexico, Tamaulipas Chiapas. PENTARRHAPHIS HBK Chlorideae 877. 878. 879. Two species in Mexico and Central America. Pantarraphis fournierana Vasey Mexico, native, endemic. Jalisco, Guerrero, Oaxaca, Mexico, Morelos. P. polymorpha (Fourn.) Griffiths Mexico, endemic. Rocky places: Sinaloa and Durango south to Oaxaca. P. scabra HBK Mexico, Central America, Colambia, native. Rochy slopes: Querétaro (tyoe loc.), Oaxaca, Chiapas and Tabasco. 280 P iH Y¥ \TiO4Ly01CyT WA Vol. 63, No. PEREILEMA Presl Eracrosteae Three species, Mexico, Central America and northern South America. 880 . P. ciliatum Fourn. Mexico, endemic. Shady banks: Sinaloa, Jalisco, Guanajuato, Mexico, Morelos, Veracruz (type loc.) and Chiapas. 881. BP. crinitum Presl Mexico and Central America south to Colambia, Ecuador and Brazil, native. Shady banks: Baja California Sur, Sonora, Chihuahua south to Chiapas. PEYRITSCHIA Fourn. Aveneae Monotvpic. 882. BP. koelerioides Fourn. Mexico, endemic. Drv slones: Durango, Guanajuato, Michoac&n, México (type loc.), Morelos, Hidalgo, Puebla and Oaxaca. 883. Peyritschia pringlei (Scribn.) Koch mexico & Central America. Sierra Madre: Sonora and Chihuahua south to Puebla. PHALARIS L. Phalarideae Twenty species, mostly northern hemisnvhere, subtemperate. 884. Phalaris anguste Nees Savanna areas of North and South America. Reported for Baja California Norte. 885. Phalaris acuatica L. Introduced, cultivated. Forage trials at exneriment stations, e.g. Chihuahua. 886. P. arundinacea L. Introduced. Cultivated: reported in Chihuahua; Baja California Norte. 887. P. canariensis L. Introduced. __ Cultivated and escaped: scattered localities throughout Mexico excert for Baja California and Yucatan Peninsula. 888. P. caroliniana Walt. U. S. and northern Mexico, native. Ditches: Baja California Norte, Sonora, Chihuahua and Coahuila. 889. P. minor Retz. Introduced. Weedy: reported for Baja California Norte, Sonora, Hidalgo and Mexico. PHARUS L. Olyreae Eight species in the American trovics. 890. P. glaber HBK Mexico and West Indies south to northern Argentina, native. Tropical woods: Chiaras and Tabasco. 1987 Beetle, Mexican grasses 281 891. P. latifolius L. Mexico to Peru and Brazil, native. Moist woods: San Luis Potosi, Mexico, Oaxaca and Veracruz. 892. P. mezii Prodoche Mexico and West Indies to Brazil, native. Tropical woods: Veracruz, Michoacan, Guerrero. PHLEUM L. Aveneae Four species, temperate regions of the world. 893. P. alpinm L. Sea eall and high mountains of the southern hemisphere, native. Mountain grasslands: Nuevo Leon, Mexico and Puebla, Veracruz (Qrizaba) ‘ PHRAGMITES Adans. Arundineae Three species, worldwide. 894. P. australis (Cv.) Trin. Worldwide. Scattered localities throughout Mexico. PHYLLOSTACHYS Sieb. Bambuseae Old World. 895. P. aurea A. & C. Riviere Introduced. Cammonly cultivated: scattered localities throughout Mexico. PIPTOCHAETIUM Presl Stipeae Ten species, Mexico and Central and South America. 896. P. brevical (Fourn.) Ricker Mexico, emic. Pine woods: San Luis Potosi, Hidalgo, Mexico, Tlaxcala and Veracruz. 897. P. fimbriatum (HBK) Hitchc. ; U. 5., Mexico and Guatemala, native. Pine woods: throughout Mexico except Tabasco and the Yucatan Peninsula. 898. Piptochaetium leianthum (Hitchc.) Beetle * Mexico, endemic. Mountain grassland, Coahuila (type loc.), Nuevo Leon, Tamaulipas and Puebla. 899. Pintochaetium mexicanum (Hitchc.) Beetle Mexico, endemic. Central mountains: Mexico and Hidalao, Puebla. 900. Piptochaetium pringlei (Beal) Parodi U.S. and Mexico, native. Bocky slopes, northern border states south to Oaxaca 901. P. stipoides (Trin. & Rupr.) Hack. Mexico and South America, native. Pine woods: Nuevo Leon, Tamaulipas, San Luis Potosi, Jalisco & Guanajuato. 902. Pinotochaetium virescens (HBK) Parodi Mexico and Guatemala, native. Pine woods: Coahuila, Nuevo Leon south to Chianas. 902a. Piptochaetium virescens var. arseni (Hack.) Beetle Mexico, endemic. Described fram Michoacan. 282 PHY TO °LJ0 GeIcA Vol. 63, No. POA L. Festficea One hundred fifty species, termerate regions, wor ldwide. 903. BP. albescens Hitchc. Mexico, endemic. Mountains: Chihuahua (tvne locality). 904. P. annua L. Introduced. Weedy: common throughout Mexico exceot for Tabasco and the Yucatan neninsula. 905.) 2: Se Vasey and Scribn. U. S. and Mexico, native. Ditches: Baja California Norte, Baja California Sur, Sonora, Chihuahua, Coahuila and Nuevo Leon. 906. Poa coahuilensis Beetle Mexico, endemic. Dry slopes: Coahuila (type loc.) 907. BP. conglomerata Rupr. Mexico, endemic. Central mountains: Mexico, Puebla and Veracruz 908. P. fendleriana (Steud.) Vasey U. S. and Mexico, native. Dry slopes: Sonora, Chihuahua, Coahuila 909. P. oariffithsii Hitchc. Mexico, endemic. Dry slopes: Sonora (tyne loc.). 910. S song ital Scribn. & Williams . and Mexico. ee peel Baja California Norte. 911. P. orcuttiana Vasey California and Mexico, native. Dry slopes: Baja California Norte, Baja California Sur. 912. P. orizabensis Hitchc. Mexico, endemic. Central mountains: Nuevo Leon, Puebla and Mexico; Chianas. 913. P. pratensis L. TIntr p Cultivated: Northern, border States, also Mexico and Veracruz. 914. P. ruprechtii Rupr. Mexico, endemic Coahuila, Nuevo Leon, Tamaulipas and Mexico (type loc.). 915. BP. scabrella (Thurb.) Benth. U. S. and Mexico, native. Dry woods: Baja California Norte. 916. P. seleri Pilger Mexico and Guatemala, native. Mountain: Oaxaca, Chianas. 917. BP. sharvii Swallen Mexico, endemic. Shary soil: Veracruz, S.L.P. Mexico, Chianas. 918. P. strictiramea Hitchc. Mexico, endemic. Ledges: Chihuahua (type loc.). 4 1987 919. 920. Beetle, Mexican grasses 283 Poa tacanae Swallen Mexico, and Guatemala. Reported fram Chiapas. P. villaroeli Phil. Mexico and Chile, native. Mountains: Mexico and Puebla. POLYPOGON Desf. Aveneae 921. 922) 823%. 924. 925. Ten species, temmerate, worldwide. Polynogon australis Brongn. Intr ed from Argentina and Chile. Reported from Baja California Norte. P. elongatus HBK. Mexico to Argentina, native. Ditches: Scattered localities throughout Mexico except for Baja California and Yucatan Peninsula. Le ukempes HBK. Canada south to Argentina, native. Ditches: Baja California Norte, Baja California Sur, Nuevo Leon and the central montains. P. maritimus L. Introduced. Ditches: Baja California Norte and Sonora. P. monsneliensis (L.) Desf. In fluced . Ditches: Baja California Norte to Coahuila; Puebla, Jalisco. PRINGLEOCHLOA Scribn. Chlorideae 926. Monotypic. P. stolonifera (Fourn.) Scribn. Mexico, endemic. Known only fram Puebla. PSEUDECHINOLAENA Stapf Paniceae Bails 928. Monotynpic. P. polystachya (HBK) Stapf Mexico to Paraguay; tropical Africa. Weedy: Querrero, Mexico, Hidalao, Puebla, Oaxaca, Veracruz, Chiapas and Tabasco. Raddia strictiflora (Fourn.) Chase Mexico, endemic. Described from "Arroyo Sellero" REEDEROCHLOA Soderstrom and Decker Aeluropodeae o257 Monotypic. R. eludens Soderstrom and Decker Mexico, endemic. Inland salt flats: Durango (type loc.) and San luis Potosi. REIMAROCHLOA Hitchc. Paniceae 930. Four species in the American tropics. R. oligostachya (Munro) Hitchc. Florida, Cuba and Mexico, native. Ditches: Colima and Tabasco. 284 P HY T .0LLO.G IA Vol. 63, No. RHIPIDOCLADUM McClure Bambuseae Eleven species in the American tropics. 931. Rhinidocladum bartlettii McClure Mexico anc Guatemala and Honduras. Reported from Chianas. 932. R. pittieri (Hackel) McClure Mexico Guatemala, native. Tropical wood margins: Chianas and Carmmeche. 933. Rhinidocladum racermiflorum (Fourn.) McClure Mexico and Central America. Reported fram Sinaloa, Durango, Tamauliras, Nayarit, Oaxaca, Michoacan, and Veracruz. RHYNCHELYTRUM Nees Paniceae About 35 species, mostly African. 934. R. repens (Willd.) C. E. Hubb. Introduced. Cammon roadside weed throuchout Mexico. 935. Rottboellia exaltata L.f. Introduced frar tronical Asia. SACCHARUM L. Andropocgoneae About 10 species in the Old World tronics. 936. S. officinarum L. Introduced. Cultivated in moist lourland areas. SACCTOLEPIS Nash Paniceae Thirty species, tropical, worldwide. 937. S. myuros (Lam.) Chase Mexico and Cuba south to Brazil, native. Marshes: Nayarit, Jalisco, Veracruz, Mexico, Oaxaca and Chianas. 938. Sacciolepis striata (L.) Nash Southeastern U.S. and Mexico. Renorted from Tabasco. SCHAFFNERA Nash Chlorideae San Luis Potosi (tvne loc.). 940. Schismus arabicus Nees Introduced Weedy, Paja California Norte and Sonora. 941. S. barbatus (L.) Thell. Introduced. Weedy: Baja California Norte and Sonora. SCLEROPOGON Phil. Eragrosteae Monotypic. 942. S. brevifolius Phil. Mexico; also Chile and Argentina, native. Dry flats: San Luis Potosi south to Puebla. 1987 Beetle, Mexican grasses 943. Scleropocon longisetus Beetle United States and northern Mexico. Northern border states south to Guanajuato. SECALE L. Hordeae Five species in temmerate Eurone and Asia. 944, S. cereale L. Introduced. Occasionally cultivated: Nuevo Leon, Mexico, Tlaxcala. SETAPIA Beauv. Paniceae One hundred species, both temperate and tropical, worldwide. 945. Setaria anceps Stapf Introduced from Africa Renorted for Tabasco and Yucatan. 946. Setaria arizonica Roamingen Southwestern U.S. (Arizona) and Reported fram Sonora 947. S. (Panicum) charmani (Vasey) Pilger U. §., Bahamas and Mexico, native. Coastal sand: Yucatan. 948. S. geniculata (Lam.) Beauv. Subtropics, worldwide, native. Weedy: cammon throughout Mexico. 949. S. grisebachii Fourn. U. S. and Mexico, native. Vieedy: common throughout Mexico. 949a. S. grisebachii Fourn. var. yucatana (Herrm.) Romingen 950. S. latifolia (Scribn.) Herr. Mexico, endemic. Brush: Durango (type loc.) Jalisco. 951. S. leucopila (Scribn. & Merr.) K. Schum. U. S. and Mexico, native. Banks: northern border states (tyne loc.: Coahuila) south to Puebla. 952. S. Liebmannii Fourn. U. S. and Central America, native. Weedy: Baja California Sur, Sonora and Chihuahua south to Chianas. 953. S. longipila Four. Mexico and Central America, native. Woods: Nayarit, Durango, Jalisco, Colima and Guanajuato 954. S. lutescens (Wiegel.) Hubb. Introduced. Reported throuchout Mexico 955. S. macrosperma (Scribn. & Merr.) Schum. Sin, Sisp Bahamas and northern Mexico, native. Moist banks: Tamaulipas, Nuevo Leon and Duranco. 956. S. macrostachya HBK U. S. and Mexico, native. Moist banks: throughout Mexico (excent Baja California Norte and the Yucatan Peninsula). 957. S. maqna Griseb. U.S., Mexico, Carribean and Costa Rica, native. Coastal: Yucatan. 286 958. 959. 960. 961. 962. 963. 964. 965. 966. 967. 968. 968a. 969. 970. 971. 972. PHY TODOCGATA Vol. 63, No. S. palmeri Henrard Mexico, endemic. Desert brush: Baia California Sur. Ss. palmifolia (Koen. ) Stanf Introduced. Cultivated: Sonora and Baja California Sur. Ss. iculifera (Steud.) Fourn. Viest Les southern Mexico to Colombia, native. Pine woods: San Luis Potosi, Veracruz, Puebla, Oaxaca, Chianas and Tabasco. s. iretiana (Schultes) Kunth Mexico, Brazil, Peru and Bolivia, native. Moist banks: Tamaulipas, San Luis Potosi, Hidalgo, Puebla, Veracruz, Oaxaca and Chiapas. S. ramiseta (Scribn.) Pilger U. S. and Mexico, native. Brush: Tamaulipas, Nuevo Leon and Coahuila. S. rariflora Mikan Mexico and West Indies to Brazil, native. Tamaulipas. S. scandens Schrad. Mexico and Central America south to Argentina, native. Wet banks: Mexico, Oaxaca, Veracruz, Chiapas, Campeche and Yucatan. §. scheelei (Steud.) Hitchc. U. S. and Mexico, native. Chihuahua to Tamaulinas and south to Michoacan, Hidalgo and Veracruz. S. setosa (Sw.) Beauv. Introduced? Reported only from Nuevo Leon. Setaria sphacetate (Schum.) Stapf Introduced. Reported: Tabasco & Jalisco. S. tenax (L.Rich.) Desv. Mexico and West Indies, Central America south to Aroentina, native. Pine woods: Nuevo Leon, San Luis Potosi, Jalisco, south to Yucatan. S. tenax var. antrorsa Pomingen Mexico, co, endemic. Brush: Yucatan (type loc.) and Veracruz. Shady banks: Tamaulipas and Nuevo Leon. Setaria variifolium (Swallen) Mexico, endemic. Yucatan (type loc.) and Quintana roo. S. verticillata (L.) Beauv. Introduced. Common throughout, S. villosissima (Scribn. & Merr.) K. Schun. Southwestern U. S. and adj. Mexico, native. Igneous rocks: Sonora, Coahuila, Nuevo Leon and Tamaulipas. 1987 Beetle, Mexican grasses 287 973. S. viridis (L.) Beauv. Introduced? Reported fram Sonora, Chihuahua, San Luis Potosi and Veracruz. 974. S. vulpiseta (Lam.) R. & S. Mexico and Caribbean to South America, native. Reported from San Luis Potosi, Veracruz, Oaxaca, Chianas and Yucatan. SETARIOPSIS Scribn. Paniceae Two species in Mexico and northern South America, 975. S. auriculata (Fourn.) Scribn. Mexico ana Central America, Colombia and Venezuela, native. Grassy plains: Sonora and Chihuahua south to the Yucatan Peninsula. 975a. S. latiqlumis (Vasey) Scribn. México, endemic. Chihuahua (type loc.) south to Chianas. 975b. S. auriculata var. depauperata (Fourn.) 3eetle SITANION Raf. Hordeae Six species in North America. 976. S. loncifolium J. G. Smith U. S. and Mexico, native. Deserts and mountains: northern border states scuth to Puebla. 977. Soderstromia mexicana (Scribn.) Mexico, endemic. Ravine: Guerrero. 978. Sohnsia filifolia (Fourn.) Airv-Shaw Mexico, endemic. Brush: San Iwis Potosi. SORGHASTRUM Nash Andronoconeae Fifteen species, temmerate and tropical America and Africa. 979. Sorghastrum albescens (Fourn.) Beetle Mexico, emic. Reported fron Veracruz, Nayarit, Jalisco, Morelos and Chiapas. 980. S. brunneum Swallen Mexico and Guatemala, native. Oak woods: Jalisco, Hidaloo, Guerrero, Oaxaca and Chianas. 981. Sorghstrum elliottii (Mohr) Nash U. S. Mexico, native. Reportes fran Navarit and Jalisco. 982. S. oaleotii Fourn. Mexico, endemic. Reported only from Jalisco, Mexico, Veracruz and Chiapas. 983. S. incompletum (Presl) Nash Mexico to Colombia and Venezuela; tropical Africa, native. Pine woods: Nayarit sonth to Oaxaca. 984. S. nudipes Nash Mexico, endemic. Pine woods: Chihuahua (tipe loc.) an Sonora. 288 Pony 20 LOG A Vol. 63, No. 4 985. S. nutans (L.) Nash Canada south to Mexico; South America, native. Grassy slopes: northern border states south to Chiavas. 986. S. setosum (Griseb.) Hitch. Tamaulipas, S. L. S., Veracruz, Oaxaca, Chiavas, and Tabasco. SORGHUM Moench. Andronogoneae 987. Thirty-five snecies, mostly African. S. almum Parodi. Introduced. Cultivated in Sonora, Chihuahua, Coahuila, Nuevo Leon, Jalisco and Colima. y 983. S. bicolor (L.) Moench. Introduced. Cultivated throughout Mexico 989. S. halepense (L.) Pers. Introduced. Common weed thrcughout Mexico. 990. S. sudanense (Piner) Stapf Introduced, cultivated. 991. S. trichocladum (Rupr.) Kuntze Native, Mexico and Central America. Oak forests: Sinaloa south to Oaxaca (type loc.) SPARTINA Chlorideae 992. About 16 species, mostly American. S. cynosuroides (L.) Roth. U. S. and Mexico, native. Atlantic coastal marshs: reported onyl from Tamaulipas. 993. OS. folsosay iran: U. S. and Mexico, native. Pacific coastal marshes: Baja Norte and Baja Sur. 994. S. junciformis Engelmann U. S., Mexico, Central America south to Argentina, native. Coastal and inland marshes: Coahuila, Nuevo Leon and Tamaulivas south to the Yucatan Peninsula. 995. S. patens (Ait.) Muhl. U. S., West Indies and Mexico, native. Atlantic coastal marshes: Tamaulipas, Veracruz, Tabasco, Campeche and Quintana Roo. SPHENOPHOLIS Scribn. Aveneae Five species in North America. 996. S. obtusata (Michx.) Scribn. Alaska and Hudson Bay south to Mexico, native. Moist places: northern border states south to Oaxaca. 997. S. airoides (Torr.) Torr. var. airoides U. S. and Mexico, native. Alkali flats: northern border states south to Durango and San Luis Potosi. 997a. S. airoides var. minor (Vasey) Beetle U. S. and Mexico, native. Alkali flats: northern border states south as far as Mexico and Hidalgo. 998. S. asper (Michx.) Kunth U. S. and Mexico, native. 1987 Beetle, Mexican grasses 289 998. Sporobolus asper (Michx.) Kunth U. S. and Mexico, native. Reported for Coahuila 999. Sporobolus asperifolia Nees & Mev Bicentric, Canada, U.S. and Mexico; also southern Argentina, native. Moist flats: Baja California Norte, Chihuahua, Coahuila, Duranoo and San Luis Potosi. 1000. S. atrovirens Kunth Mexico, endemic. Baja California Sur, Durango to Tamauliras and south to the Yucatan Peninsula. 1001. S. buckleyi Vasey U. S. and Mexico; British Honduras, native. Shady flats: Nuevo Leon and Tamaulipas south to the Yucatan Peninsula. 1002. Smorobolus coahuilensis Valdez Mexico, endemic. Desdribed from Coahuila. 1003. S. contractus Hitchc. U. S. and Mexico, native. Gravel slopes: Baja California Norte, Sonora, Chihuahua, Coahuila and San Luis Potosi. 1004. S. cryrtandrus (Torr.) A. Gray U. S. and Mexico, native. Sandy soils: northern border states and Baja California Sur. 1005. S. cubensis Hitchc. Mexico, West Indies; Central America south to Bolivia, native. Oaxaca, Chiapas, Veracruz and Tabasco. 1006. S. damingensis (Trin.) Kunth Florida, Caribean and Mexico, native. Beaches: Yucatan and Quintana Poo. 1007. Sporobolus erectus Hitchc. Mexico, endemic. Reported from Veracruz (type loc.) and Chiapas. 1008. S. flexuosus (Thurb.) Rydb. U. 5. and northerm Mexico, native. Mesas: Northern border states. -1009. S. giganteus Nash U. S. and Mexico, native. . Mesas: Chihuahua and Coahuila. 1010. Sporobolus hintoni Hartley Mexico, endemic. Described from the State of Mexico. iol. Sh andicuss@.)ieR. Br. Mexico and West Indies to Colombia and Brazil, native. Sandy pine uplands: throughout Mexico. 1012. Ss. jacquerontii 1013. S. jumceus (Michx.) Kunth U. S. and Mexico, native. Pine barrens: Veracruz and Chianas. 1014. S. macrospermus Scribn. Mexico and Guatemala, native. Pine-oak: forests: Sinaloa, Nayarit, Zacatecas, Jalisco (tyne loc.) south to Chiapas. 290 1015. 1016. 1017. 1018. 1019. 1020. 1021. 1022. 1023. 1024, 1025. 1027. 1027a. PH Y T O*L'O'o WA Vol. 63, No. S. nealleyi Vasey U. S. and Mexico, native Gypvsophilous soils: Coahuila, Nuevo Leon and San Luis Potosi. S. palmeri Scribn. Mexico, endemic. Alkaline soils: Durango, Coahuila and San Luis Potosi. S. patens Swallen U. S. and Mexico, native. Known only from Sonora. S. pulvinatus Swallen U. S. and Mexico, native. Wet oravels: northern border states south to Caxaca. S. purpurascens (Sw.) Hamilt. U. S., West Indies, Mexico; Central America south to Peru, native. Salt Flats: Revillagiaedo Islands, Tamaulivas, Veracruz, Oaxaca and Chiapas. S. pyramidatus (Lam.) Hitchc. U. S., West Indies, Mexico; Central America south to Argentina, native. Salt flats: common throughout Mexico. S. regis I.M. Johnston Mexico, endemic. Salt flats: known only from Coahuila. S. spiciformis Swallen Mexico, endemic. Known only from Coahila. Sporobolus tenuissimus (Schrank) Kuntze Mexico, Virgin Tslands and Brazil. Reported from Michoacan. lus ii Hitchc. Texas and adjacent Mexico, native. Reported only for Tamaulipas. S. trichodes Hitchc. Mexico, endemic. Chihuahua, Zacatecas, Aguascalientes, Guanajuato, Jalisco (type loc.), Michoacan, Guerrero, Mexico, Morelos and Veracruz. S. virginicus (L.) Kunth var. virginicus Tropic subtropical coasts, worldwide. Coastal dunes and flats: Baja California, Sonora; Tamaulipas south to the Yucatan Peninsula. S. virginicus var. major Drege STENOTAPHRUM. Trin. Paniceae 1028. Seven species, trovical and subtropical, worldwide. S. secundatum (Walt.) Ktze. Tropics and subtropics, worldwide, native. Commonly cultivated throughout Mexico. 1987 Beetle, Mexican grasses 291 STIPA 1. Stipeae One hundred fifty species, temperate, worldwide. 1029. S. acuta Swallen México, endemic. Rocky soils: Carneras Pass, Coahuila. 1030. S. alta Swallen México, endemic. Desert snrub: known only from Coahuila. 1031. S. bracteata Swallen México, endemic. Known only from Baja California Norte. 1032. S. clandestina Hack. México, endemic. Coahuila and Nuevo Le6n, San Luis Potosi, Zacatecas, Aguascalientes, Michoac&n, Guanajuato and México. HOSS) oe CON Sina cCanHatehe. México, endemic. Rocky slopes: Hidalgo (type loc.), Nuevo Le6én and Tamaulipas south to Oaxaca, Veracruz and México. 1034a S. coronata Thurb. var. coronata U.S. and México, native. Rocky slopes: Reported only from Baja California Norte. 1034b S. coronata var. depauverata (Jones) Hitchc. U.S. and México. Revorted from Baja California Norte. 1035. S. diegoensis Swallen U.S. and México, native. Rocky slopes: Reported only from Baja California Norte. 1036. S. editorum Fourn. México, endemic. Rocky slopes: Coahuila, Nuevo Le6n and Tamaulipas. South to Puebla. 1037. S. eminens Cav. U.S. and México, native. Rocky slopes: Northern border states, south to Oaxaca. 1038. S. ichu (Ruiz & Pav.) Kunth México south to Argentina, native. Rocky slopes: Coahuila, San Luis Potosi south to Chiapas. 1039), S. Yepida Hitche- U.S. and México, native. Rocky slopes: reported only from Baja California Norte. 1039a S. lenida var. andersonii Vasey 1040. S.leucotricha Trin. & Rupr. U.S. and México, native. Grassland: Coahuila, Nuevo Le6n and Tamaulivas south to Oaxaca. 1041. 5. "iinearitolia Fourn. México and Guatemala, native. Central mountains: type from México "vrope Tacubaya" 1042. S. linearis Swallen México, endemic. Rocky slopes: known only from Nuevo Le6n. 1043. S. lobata Swallen U.S. and México, native. Revorted from Coahuila and Chihuahua. 292 PHY fT OsL OG Es Vol. 63, No. 1044. S. mucronata HBK Mexico south to Argentina and Chile, native. Mountains: Chihuahua, Coahuila and Nuevo Leon south to Chiapas. 1045. S. multinodis Scribn. Mexico, endemic. Mountains: Chihuahua, Coahuila and Nuevo Leon south to Puebla. 1046. S. neamexicana (Thurb.) Scribn. U. S. and Mexico, native. Sonora, Chihuahua, Coahuila, Nuevo Leon ana San Luis Potosi. 1047a. S. pulchra Hitchc. var. chra U. S. and Mexico, native. Dry slopes: reported only for Baja California Norte. 1047b. stip plc BHitchc. var. Cernua (Stebbins & Love) Beetle Tofsrud U. S. and Mexico, native. Rerorted from Baja California Norte. 1040a. S. robusta (Vasey) Scribn. U. S. and Mexico, native. Rocky slopes: Coahuila, Nuevo Leon and Tamaulipas. 1040b. S. saxicola Hitchc. Mexico, native, endemic. Described from Puebla. 1049. S speciosa Trin. & Rupr. URS Mexico, southern South America, native. Rocky slopes: reported onlv from Baja California Norte. 1050. S. tenuissima Trin. Mexico; also Chile and Aroentina, native. Rocky slopes: Coahuila, Nuevo Leon, San Luis Potosi, Guanajuato, Veracruz, Puebla and Oaxaca. 1051. S. virletti Fourn. Mexico, endemic. Known only fram San Luis Potosi. STIPORYZOPSIS Johnson Stipeae Two or three species in North America. 1052. S. bloameri (Bol.) Johnson Introduced. Cultivated: reported only from Coahuila. STREPTOCHAETA Schrad. Olyreae Two species in tropical America. 1053. S. sodiroana Hack. Mexico south to Ecuador, native. Tronical forest: reported only from Chianas. 1054. S. spicata Schrad. Mexico south to Brazil, native. Tropical forest: Veracruz and Chiapas. STREPTOGYNE Beauv. Streptoayneae One species in the American trorics 1055. 6&. americana Hubb. Mexico and Trinidad south to Brazil, native. Tropical woods: Veracruz and Chiapas. 1987 Beetle, Mexican grasses 293 THRASYA HBK Paniceae Twenty species in the American tropics 1056. 1. campylostachya (Hack.) Chase Mexico south to Bolivia, native. Pine forestes: Veracruz, Oaxaca and Chiapas. TRACHYPOGON Nees Androrogoneae Fifteen species in American tropics. 1057. T. ini Fourn. Mexico, endemic (introduced in Cuba). Veracruz (type loc.). 1058. TT. karwinskvi (Hack.) Nash Mexico, endemic. known only from the type locality: "Mexico" 1059. T. montufari (HBK) Nees Mexico and Ecuador, native. Zacatecas and San Luis Potosi south to Chiapas. 1060. T. palmeri Nash Mexico, endemic. Known only fram Jalisco. 1061. T. plumosus (H.&B.) Nees U. S. south to tropical South America, native. Reported only from Veracruz. 1062. T. secundus (Presl) Scribn. U. S. and Mexico south to Argentina, native. Baja California Norte, Sonora and Chihuahua south to Chiapas and Tabasco. 1063. Trach on stenonhyllus (R. & S.) Hitchc. Mexico and Central America, native. Grasslands: San Luis Potosi, Veracruz, Oaxaca, Chiapas, Tabasco and Camneche. TRAGUS Hall Zovsieae Three species in tropics and subtropics. worldwide. 1064. T. berteronianus Schult. Introduced. ; Northern border states south to Oaxaca. 1065. Trichoneura eleaans Swallen Southern Texas and northern Mexico, native. Reported for northern Tamaulipas. TRIDENS R. & S. . ERAGROSTEAE Sixteen North American species. 1066. T. albescens (Vasey) Woot. & Standl. U. S. and Mexico, native. Swales: Chihuahua, Coahuila, Nuevo Leon and Tamaulipas. 1067. T. eragrostoides (Vasey & Scribn.) Nash UL Say Mexico, native. Swales: Nuevo Leon, Oaxaca and Yucatan. 1068. T. flavus (L.) Hitche. U. S. and Mexico, native. Swales: reported only for Nuevo Leon. 294 POH Y T Oxb)0)G) vA Vol. 63, No. 1069. T. muticus (Torr.) Nash U. S. and Mexico, native. Swales: northern border states south to Durango, Zacatecas and San Luis Potosi. 1070. T. texanus (S. Wats.) Nash U.S. and Mexico, native. Swales: Coahuila, Nuevo Leon, Taraulipas and San Luis Potosi. TRINIOCHLOA Hitchc. Aveneae Three American species. 1071. T. laxa Hitchc. Ravine: reported only for Chihuahua. 1072. T. micrantha (Scribn.)Hitchc. Mexico, endemic. Reported only for Mexico and Morelos (tyne loc.), native. 1073. T. stipoides (HBK)Hitchc. Mexico south to Bolivia, native Pine woods: San Luis Potosi south to Chianas. 1074. Triplasis caribensis (Pohl) Beetle Mexico and Honduras. Sandy beach, Tabasco. TRIPOGON Roth Eragrostae 1075. T. spicatus (Nees) Ekman U. S., Cuba, Mexico and South America, native. Dry banks: Durango, Aguascalientes, San Luis Potosi and Veracruz. TRIPSACUM L. Andropogoneae Nine species in the Americas. 1076. Tripsacum bravum Grav Mexico, native, endemic. Mexico (Type collection) and Jalisco. 1077. T. dactyloides (L.) L. U. S., West Indies and Mexico, native. Grasslands: Coahuila and Nuevo Leon south to Guerrero. 1078. Tripsacum fasciculatum Trim. Mexico, West Indies, Central and South America, native. Barrancas, especially near cultivated vlots, Nayarit, Jalisco, Veracruz, Guerrero, Oaxaca and Chiapas. 1078a. Trivsacum floridanum Porter and Vasey Mexico and Florida, native. Pire words Tamaulipas 1079. T. lanceolatum 8upr. Mexico, Honduras and Panama, native. Pine woods: common throughout Mexico. 1080. T. maizar Hernandez X and Randolnh Mexico, endemic. Reported only for San Luis Potosi. 1081. T. vilosum Scribn. and Merr. Mexico, endemic. Forest margin: Chihuahua south to Chianas. 1987 Beetle, Mexican grasses 1082. Trivsacum zonilatanse Hernand. and Ranc. Mexico, native, endemic. Revorted from Michoacan and Guerrero. TRISETUM Pers. Aveneae Seventv-five species in temperate regions, worldwide. 1083. T. deyeuxioides (HBK) Kunth Mexico Central America south to Ecuador, native. Mountain meadows: Chihuahua and Nuevo Leon south to Chianas. 1083a. T. deyeuxiostis var. nubescens Scrijsner Mexico, endemic. Nuevo Leon, San Luis Potosi, Jalisco, Michoacan, Mexico, Guerrero and Veracruz. 1084. T. filifolium Scribn. Mexico, endemic. Pine, oak woods: Chihuahua (tyne loc.) and Duranco. 1085. T. interrmtun Fourn, U. S. and Mexico, native. Reported for Baja Norte (Type loc. for T. californicum). 1086. T. iramiense (Kuntze) Hitchc. Merits... Mexico and Central America south to Ecuador, native. Pine woods: fan Luis Potosi south to Chianras. 1087. Trisetum mexicanum (Swallen) Koch. Mexico, native, endemic. Described from State y Mexico 108€. T. palmeri Hitchc. Mexico, endemic. Sonora, Ccahuila, Sinaloa, Durango (tyme loc.) and Jalisco. 1089. TT. pringlei (€cribn)Hitchc. Mexico south to Panama, native. Meadows: Oaxace (tvpe loc.) and Chianas. 1090. TT. rosei Scribn. and Merr. Mexico and Guatemala, native. Mountains: Hidalao, Mexico, Puebla and Chiapas. 1091. T. spicatum (L.) Richt. Brtic-alpine and south in the mountains, native. Mountains: Coahuila and Nuevo Leon south to Puebla. 1092. TT. viride (HBK) Kunth. Mexico, endemic. Mountains: Queretaro (Tyne loc.), Mexico and Oaxaca. 1093. 7. virletii Fourn. "Mexico, endemic" Mountains :Nuevo Leon, San Luis Potosi (tyne loc.), Michoacan, Guerrero, Mexico, Morelos, Puebla and Veracruz. TRISTACHYA Nees AVENEAE Five snecies in Mexico and South America 1094, TT. angustifolia Hitche. Mexico, endemic. Known only from Nayarit. 1095. T. avenacea (Presl)Scribn. & Merr. Mexico and Guatemala, south to Brazil, native. Forest margin: Jalisco, Michoacan, Mexico, Oaxaca and Chianas. 296 PHY TOLOCGIA Vol. 63, No. 4 1096. T. laxa Scribn. & Merr. Mexico, endemic. Durango (tyne loc.) and Sinaloa. 1097. Trystachya penilosa Guzman Mexico, native, endemic. Described from Nayarit. TRITICUM L. Hordeae Many cultivated types. 1098. T. aestivum L. Introduced. Commonly cultivated. UNIOILA L. Centotheceae Ten snecies in the American (cf. Chasmanthium Link) U. S. and Mexico, native. Reported onlv for Nuevo Leon 1099 U. naniculata L. U. S. and Carribean and Mexico, native. Coastal dunes: Tamaulinas, Veracruz, Tabasco and Chianas. 1100. U. nittieri Hack. Mexico and Central America south to Ecuador, native. Sea beaches: Baja California, Sonora, Sinaloa, 1101. Vetiveria zizanioides (L.) Nash Introduced from Asia. Cultivated and escaned, renorted fram Chiapas VULPIA Gnel. Festuceae Thirty svecies, temnerate Europe and North and South America. 1102. V. bromoides (L.) Gray Introduced. Weedy: reported from Mexico and Chiaras. 1103. V. microstachys Munro var. ciliata 1103a. V. microstachys Munro var. nauciflora (Beal) Leonard & Gould 1104. V. var. hirsuta Hack. Tatreduced? Weedy: Baja California Norte, Sonora and Coahuila. 1104a. V. octoflora (Walt.) Rybd. PEAT. Lo ee Andronoconeae Cne or two American srecies under cultivation. 1105. Z. mays L. Subtropical American, native. Commonly cultivated throughout Mexico. 1106. Z. perennis (Hithc.) Reeves and Mancels. Mexico, endemic. — Described fram Jalisco. 1987 Beetle, Mexican grasses 297 ZEUGITES P. Br. Eragrosteae Twelve species in the American tropics. 1107. Z. americana Willd. var. mexicana (Kunth)Urban Mexico south to Bolivia, native. Tropical forest, S.L.P., Hidalgo, Guerrero, Veracruz and Chiapas. 1108. Z. auriculata Tenori Mexico, native, endemic. Tropical forest or oak woods, Jali§o, Michoacan, Mexico, Moralos and Guerrero. 1109. Z. capillaris (siitenc. ) Swallen exico, endemi-. ropical forest; Jalisco and Colima (type loc.). 1110. Z. hackelii Swallen Mexico, endemic. Tropical forest, Jalisco (type loc.) and Mexico. 1111. Z. latifolia (Fourn, )Hemsl. Mexico, endemic. Tropical forest; Jalisco, Guerrero, Oaxaca (type loc.) and Chiapas. ZOYSIA Willd. 5 species, mostly Asian. 1112. Z. matrella (L. )Nerr. Introduced from Asia. Cultivated for lawns; Nayarit. 1113. Z, tenuifolia Willd. Introduced from Asia. Cultivated for lawns, baja California Norte. ZIZANIOPSIS Doell and Asch. Three or 4 species in the Americas. Z. miliacea Doell and Asch. U.S.A.. and Mexico, native. Aquatic, Veracruz. CYPERACEAE TRINITENSES NOVAE C. D. ADAMS British Museum (Natural History), London, England Rhynchospora ierensis C. Adams, Spe nov. Herba perennis. Rhizoma torulosa breviter ramosa. Culmi plures dense caespitosi 12-40 cm alti filiformes compresso-trigoni costati glabri laevesque. Folia basalia caulinaque scapo fere dimidio breviora 0.5-0.8 mm lata basi plana medio canaliculata apice triquetra necnon scab- ridiuscula cetera glabra; vaginae truncatae striatae marginibus supernis hyalinis. Inflorescentia e corymbis 2-3-paucis culmo medio supernoque constructa; corymbus terminalis usque ad 2 cm altus et 3.5 cm latus decompositus paulo apertus radiis primariis circa 8 inaequalibus usque ad 2.5 cm longis; radii graciles bracteolis filiformibus vel setaceis basi membranaceis suffulti; corymbi laterales parviores compositi plerumque latiores quam longiores ped- unculis filiformibus. Spiculae singulae binatae vel ad 8-10-nim fasciculatae ramorum primariorum extremitates collocatae in anthelae formantes 2-3 mm longae ovoideo- lanceolatae teretes acutae pedicellatae (ubi singulare) vel subsessiles curvatae 2-florae 1-nucigerae; glumae circa 6 imus brevissimae ovatae 1-nervatae muticae vel minute setuloso-mucronatae membranaceae marginibus latis hyalinis infuscatae striis patentibus rubro-brunneis; stamina 3; stylus profunde 2-fidus. Achenium circa 0.8 mm longum ubique 0.9 mm latum suborbiculare biconvexum breviter stipitatum grosse transverso-rugulosum porcis 4-6 minute verticali-lineolatis et marginibus crassis cinerascenti-flavum; rostrum valde depressum circa 0.1 mm altum O.5 mm latum ad dimidium apicis extendens vix bilobatum minute apiculatum album vel cinerascens; setae nullae. Trinidad. Erin Savanna, 11.viii.1977, Philcox & Ramcharan 8146 (holotype K; isotype TRIN 23229). Additional collections: Erin Savanna, 28.vii.1980, Alkins sen. (BM, TRIN 24035); Icacos, road to the savanna, 26.11.1866, Finlay ? (TRIN 1891). Endemic to south-western Trinidad. 298 1987 Adams, Cyperaceae trinitensis novae 299 This species is distinguished from R. tenuis Link and R. emaciata (Nees) Boeckeler by the curved spikelets and smaller broader achenes with the beak strongly depressed. Scleria orchardii C. Adams, spe nov. Herba perennis. Rhizoma brevis torulosa. Culmi approximati 70-100 cm alti prope basim 2-4 mm lati erecti triquetri glabri. Folia ad basim elaminata vel laminis redactis; vaginae laxiusculae ad 13 cm longae alis sursum divergentibus ad 4 mm latis marginibus minute antrorse scabridiusculis cetera glabrae; contraligula ovato-deltata circa 5 mm longa nervosa margine ciliato; foliorum laminae (8-)20-50 mm longae 10-18(-23) mm latae lineares post apicem (2-)7-12 cm abrupte angustatae et sursum gradatim decresc- entes marginibus et costis abaxialibus non nisi prope apicem extremum antrorse scabridiusculis. Inflorescentiae 1-2, terminales et interdum e folii summi axilla ortae, termin- alis 4-8 cm longa 3-5 cm lata composita lateralis breviora pyramidalis ramis rigidis patentibus rubro-purpureis bract- eis foliaceis suffultis; rhachis principalis rami primarii ramulique acute angulati praeter interdum distales sparse scabridiusculi glabri haud passim puberuli; bracteolae setaceae ad basim longi-ciliatae. Spiculae spicatae vel racemosae; pistillatae circa 5 mm longae sessiles vel sub- sessSiles, vulgo spicula staminata 3.5-4 mm longa pedicell- ata concomitata pare bracteola filiformi ad basim lata valde ciliata suffulta. Hypogynium trilobatum lobis circa 2 mm longis 3 mm latis circa dimidiis versus basim irregulariter fissis eburneum apicibus atrantibus. Achenium 4 mm longum 3 mm latum ovoideo-ellipsoideum exapiculatum laeve nitens ad apicem puncto albo basim azureo-lacticolor cetera atro- purpureum. Trinidad. Las Lapas Trace, off Arima-Blanchisseuse road, 21.11.1981, Adams 14806 (holotype K; isotypes BM, TRIN 29929). Additional collections: Morne Bleu, 31.xii.1925, Williams sen. (TRIN 11495); Las Lapas Trace, 6.ix.1973, Adams 13603 (TRIN 20848). Endemic to the Northern Range of Trinidad. This species is closest to S. latifolia Swartz. It is 300 PHT 0. 0 Gi A Vol. 63, No. 4 distinguished by its ovoid-ellipsoid achenes, narrower leaves and differences of indumentum. The likelihood of its having arisen as a hybrid between that species and S. melaleuca Reichb. ex Schldl. & Cham. should be considered. The name is given to commemorate Robert Orchard Williams, economic botanist, who served with the Government of Trinidad between 1916 and 1934, and was, with Professor E.E.Cheesman, responsible for the preparation of early parts of the Flora of Trinidad and Tobago. Torulinium macrocephalum (Liebmann) C. B. Clarke var. davidsei C. Adams, var. nov. Panes Inflorescentia radiis primariis secundariisque evol- utis. Spicae 2-5 cm longae lineares plerumque simplices racemosae bracteis secundariis foliosis subtentis. Spiculae 5-8(-10) mm longae; flosculi plerumque 4-8; glumae 2-2.5 longae. Trinidad. 1 mile W of Blanchisseuse at Marian(n)e River crossing, 31.vii.e1970, Davidse 2477 (holotype MO). Additional collections: Diego Martin, Mahault, 26.viii.1916, Broadway s.n. (TRIN 8459); River Estate, 2.iii.1920, Britton & Hazen 187 (K, NY); Bande de L'Est, Mayaro, ? Finlay in 1866 (TRIN 2085). Typical T. macrocephalum has an inflorescence without evident rays, the spikes being congested into a usually solitary rounded compound head. Other varieties are rad- iate, IT. macrocephalum var. eggersii having ovoid sessile spikes in clusters at the ends of primary rays and this new variety having linear peduncled spikes in short racemes. The name of the new variety eponymises Dr. Gerrit Davidse, Curator, Missouri Botanical Garden. A NEW ARIZONA PURSHIA (ROSACEAE) Clark G. Schaack Department of Biological Sciences Deaver Herbarium Northern Arizona University Flagstaff, AZ 86011 Recent critical morphologic analysis of the extreme varfation included within the central Arizona species, Purshia (Kearney) Henrickson (Henrickson, 1986) and similar varfation observed on lectotype and {isolectotype preparations of the basfonym (Schaack, 1987) indicate that B. subintegra was based on material of hybrid origin. Morphology and phenology displayed in central Arizona hybrid residuum suggests that this nothospecies was formed via past hybridization between Pursbia stansburiana (Torr. ) Henrickson and an unnamed species previously included within P. subintegra. This unnamed species, now confined to calcareous substrate in Graham County, Arizona, is here described. Data to support the hybrid origin of B. subintegra will be presented elsewhere. PURSHIA PINKAVAE Schaacks sp. nov. Species haec et B. ericifolia (Torr.) Henrickson folia linearia vel angustissime spatulata integra valde revoluta speciebus aliis Burshiae differunt. E PB. ericifolia brachyblastis cylindricalibus lignosis, follis dispositis alternatim helicaleque cum apicibus rotundatis minute mucronatis, hypanthiis normaliter eglandulosis et tempore fructificante infundibuliformibus praecoce deciduisy lobis calycis non-apiculatis pubescentibus (superne sericeis, subtus plus minusve glabris) distinguenda. TYPE: U.S.A.: Arizona: Graham Co., along US highway 70, ca. 16 km northwest of Bylas, 11 Apr 1987, Schaack 2046 & Schaack (holotype ASC; isotypes to be distributed). Intricately branched shrubs to 1.4 m tall, multistemmed or with a very short single trunk; bark of older stems dark gray, fissured and shredding; upper branches light gray and smooth; woody spurs light gray, round in cross sections branched or unbranched, {in part composed of, and roughened by» lignified stipularpodia. Leaves helically alternate, in clusters on young growth or at the apex of woody spurs. Leaf blades borne on herbaceous~corfaceous ciliate-margined stipularpodia, green or brownish-green with age, perennial, herbaceous-corfaceous, linear-very narrowly spatulate, largest blade per woody spur 6-15 times as long as wide; uppe™ 301 302 PHY TD 68.10 Grivs Vol. 63, No. 4 surface loosely arachnose and scabrescent when young, glabrate- glabrous in ages not punctate; lower surface lanate when young; margins entire, revolute to strongly so; apex minutely mucronate or obtuse. Flowers 1 per spur or spur branch, sessile or pedicellate; pedicels 1-7 (-13) mm longs normally eglandulars moderately villous-lanate; hypanthium narrowly funnelform, ca. twice as long as wide, normally eglandular, moderately villous-lanate, funnelformbroadly funnelform and soon deciduous in fruit; calyx 5- lobed, sepals ovate and obovate, 2.0-5.5 mm longs eglandular, moderately villous-lanate, not apiculate; petals normally 5, ochroleucous (white?) and occasionally red or pink=-tinged, obovate- spatulate, largest petal (1.3-) 1.5-2.0 times as long as wide, glabrous or occasionally pubescent apically; stamens many, 2 (-3) series, inserted on the hypanthium; pistils sessile, distinct, 2-4 (-5) per flower. Achenes at maturity ca. 5-7 mm longs 2.8-3.3 mm wide, brown, + lanceolate, strongly striate with ca. 14-20 striae, sericeous above and + glabrous below; fruiting styles terminal, plumose, (17-) 20-50 (-55) mm long; fruiting stigmas 1-2 mm long or less. po = 9. Anthesis late March-mid April. Plants restricted to late Tertiary calcareous, lacustrine deposits ca. 16-21 km northwest of Bylas, Graham County, Arizona. ADDITIONAL SPECIMENS EXAMINED. U.S.A.: Arizona: Graham Co.,s US highway AG ca. 16 km ™ of Bylas, 9 Apr 1984, 11 Apr 1985, ca (ASU); ca. 18 km NW of Bylas, 17 Mar 1982, Clark 1506 & Parfitt (ASU); milepost 284 NW of Bylas, 18 Sept 1976, McGill] & Lehto 120683 (ASU); ca. 20 km NW of Bylas, 7 Sept 1968, Pinkavae Keil & Lehto 113397 (ASU); ca. 15.3 km NW of Bylas, 7 Apr 1969, Pinkavas, Kei] & Lehto 115583 (ASU); ca. 17.7 km NW of Bylas, 16 June 1974, Roberts & Keil 10126 (ASU); ca. 21 km NW of Bylas, 12 Apr 1986, Schaack 1742-175] & Schaack (ASC); ca. 16 km NW of Bylas, 16 May 1987, Schaack 2066-2068 & Schaack (ASC); between milepost 283 & 282 NW of Bylas, 16 May 1987, Schaack 2069 4 2070 & Schaack (ASC). This species is named in honor of Donald J. Pinkavas Professor of Botany at Arizona State University, student of Purshia, and the Arizona flora, in particular Cactaceae and Asteraceae. He was among those first to collect and report (Pinkava et. al.» 1970) the Graham County material northwest of Bylas. The linear-very narrowly spatulate, entire, strongly revolute leaf blades of B. pinkavae and B. ericifalia contrast strongly with, and readily distinguish these species from, the obovate (in outline) lobed, or toothed blades of other Burshia. The cylindrical woody spurs, helically alternate leaves with minutely mucronate-rounded apices, 2-4 (-5) achenes per flower, normally eglandular hypanthia, funnelform and soon deciduous fruiting hypanthia and eglandular non-apiculate sepals of PB. pinkavae distinguish this species from PB. gricifolia. Purshia ericifolia 1987 Schaak, A new Arizona Purshia 303 has flattened and often twisted woody spurs, distichous alternate leaves with aristate tips, 8-17 achenes per flower, normally stipitate glandular hypanthia, campanulate-broadly campanulate persistent fruiting hypanthia and stipitate glandular apiculate (in bud and at anthesis) sepals. Specific relationships in Burshia (By pinkavae and P. ericifolis) and the importance of hybridization as a speciation mechanism in Purshia will be topics for future investigations. ACKNOWLEDGEMENTS I thank Drs. David J. Keil» James Rominger and particularly Donald J. Pinkava for comments and criticisms on an earlier draft of this manuscript. Dr. David J. Keil kindly provided the Latin translation for the diagnosis of B. pinkavae and I gratefully acknowledge his assistance. LITERATURE CITED Henrickson, J. 1986. Notes on Rosaceae. Phytologia 60: 468. Pinkavas D.J.» E. Lehto and D. Keil. 1970. Plants new to the Arizona Flora - III. Jour. Ariz. Acad. Sci. 6: 134. Schaack,s C. 1987. Lectotypification of Cowania subintegrs Kearney» basionym of Burshia subintegra (Kearney) Henrickson (Rosaceae). Taxon 36: 452-454. TAXONOMIC IDENTITY OF LEUCAENA LEUCOCEPHALA (LAM.) DE WIT, WITH A NEW COMBINATION. Sergio Zarate Pedroche Universidad Nacional Auténoma de México, Herbario Nacional (MEXU). Presently: Graduate student supported by CONACYT, Department of Biology, Indiana University, Bloomington, IN, 47405. A considerable amount of agronomic research has been produced about Leucaena leucocephala (Lam.) de Wit (1961), well known in the tropics both as a weed and for its multiple uses (Dijkman, 1950; National Academy of Science, 1977). De Wit (1961) published this combination based on a description by Lamarck (1783), of which a type specimen exists in Lamarck’s Herbarium in Paris (P.LA). The validation of Lamarck’s epithet, in replacement of the Linnaean epithet glauca, generated a series of controversial papers (Isely, 1986, and references therein); however, the systematic identity of the binomial requires further clarification. A native to Mexico and Central America, this species was introduced by the Spaniards to the Philippine Islands during the XVI century; from there it expanded to its present pantropical distribution. Study of the Mexican populations of L. leucocephala led to the recognition of two distinct, but closely related taxa, and subspecific rank was proposed for them (ZArate P., 1982). Confusion exists as to the correct names of the three recognized horticultural varieties: ‘Hawaii’, ‘Salvador’ and ‘Peru’. The ‘Hawaii’ type was characterized from naturalized Hawaiian populations as a ramose shrub, flowering in all seasons, pantropically established as a successful colonizer (Brewbaker, 1975). The descriptions of L. leucocephala from Hawaii (Degener, 1946), as well as from the Yucatan Peninsula and the Tehuantepec Isthmus (Zarate P., 1982), clearly match Lamarck’s (1783) Mimosa leucocephala. The second agronomic type, known as ‘Salvador’ or ‘Hawaiian Giant’, despite the fact that it originated in Mexico (Brewbaker, 1975), is a fast growing, tall tree, flowering annually. The name ‘Salvador’, alluding to Central American origin, led to confusion with L. salvadorensis (Brewbaker, 1978, 1984), a different species (Zarate P., 1987). Both the ‘Salvador’ and ‘Peru’ types, the latter characterized by its ramose, arborescent habit, belong to a distinct taxon distributed in Mexico and Central America at altitudes from sea level to ca. 1000 m. It was described by J.N. Rose (1897) as L. glabrata, and is thus combined: L. leucocephala 304 1987 Pedroche, Identity of Leucaena leucophylla 305 (Lam.) de Wit subsp. glabrata (Rose) Zdrate P., comb. et stat. nov., L. glabrata Rose, Contr. U.S.Natl. Herb. 5:140-141, 1897. Mexico: Guerrero: ‘near Acapulco’, Palmer 368 (Holotype, US; Isotypes, A!, GH!) Both subspecies are cultivated for their edible seeds in Mexico, where their domestication probably occurred independently. L. Jleucocephala subsp. glabrata predominates as a cultigen, while subsp. leucocephala is an isolated domesticate (ZArate P., unpublished). Despite this, it is the latter subspecies that is now found naturalized in the Philippines, to where it must have been carried by the colonists; the explanation for this is not clear. It could have been an accidental dispersal, e.g. with the sand used as ballast in the ship journey from Mexico, or perhaps both subspecies were introduced, and selection was the cause for the establishment of subsp. lJleucocephala, better adapted to calcareous substrates such as seashores and coral reefs. In commercially propagated lineages (e.g. ‘PERU' & K67), segregation for pubescence of legumes and leaves has been observed (Soto & Zarate P. 52, 53 MEXU). In the Tehuantepec Isthmus region subsp. leucocephala has atypical glabrescent legumes but the usual canescent leaf buds and ciliate leaflets. These facts suggest that despite the species’ known self-compatibility (Brewbaker, 1982), gene exchange between the subspecies is possible and could have had a role in the development of the naturalized taxon, as well as in artificial selection of cultigens both recently and during the early domestication of the species. The taxa may be distinguished as follows: Small trees or shrubs 1-6 m tall. Foliar buds canescent; petiolar gland 1.5-2.5 mm long; pinnae 6-9 pairs; leaflets 9-18 pairs, 9-12 mm long, ciliate. Flowering buds canescent. Legume 13-16 x 1.2-1.5 cm, the stipe 7-10 mm long, velutinous or glabrescent. Seeds 5-8 mm long, 4-5 mm wide. Flowering in response _to water. Ce subsp. leucocephala Trees or shrubs 3-12 m tall, or taller. Foliar buds glabrate; petiolar gland 1-4 mm long; pinnae 3-8 pairs; leaflets 11-24 pairs, 8-15 mm long, glabrous. Flowering buds glabrous. Legume 11-18 x 1.2-2.3 cm, the stipe 10-20 mm long, glabrous. Seeds 6-10 mm long, 3-6 mm wide. Flowering once a year. subsp. glabrata S) 5 Ss) Clw)w)\6 6) {@) 6) 6) 6: 6 68, e! 0) @ (0 ¢! 8) © «'6] 8 (6) us) ACKNOWLEDGEMENTS The Instituto de Biologia, U.N.A.M. through the Herbario Nacional (MEXU) provided the funds for field work. M. en C. Mario Sousa S., former curator of MEXU, made avilable to me the type specimens loaned from A and GH, as well as his expertise. 306 PHY 16,0 0°¢ rs Vol. 63, No. Silvia E. Purata V. photographed the type specimens in P.LA for me. To Prof. Charles B. Heiser, Jr. for his ideas on plant dispersion to the Philippines. George Yatskievych reviewed and improved the manuscript. LITERATURE CITED Brewbaker, J.L. 1975. ‘Hawaiian Giant’ Koa Haole. Hawaii Agricultural Experiment Station. Miscellaneous Publication no. £25: 1978. Guide to the Systematics of the Genus Leucaena (Mimosaceae). Mimeograph, C.1.A.T., Cali. 1982. Systematics, Self- Incompatibility, Breeding Systems, and Genetic Improvement of Leucaena Species. Leucaena Research in the Asian-Pacific Region. Proc. of Workshop Nov. 1982. IDRC, Ottawa. p. 17-22. : 1984. Revision of the Genus Leucaena (Mimosoideae: Leguminosae). Unpublished paper, Honolulu. Degener, O. 1946. Flora Hawaiiensis 2nd. Ed. Honolulu. De Wit, H. 1961. Typification and correct names of Acacia villosa Willd. and Leucaena glauca (L.) Benth. Taxon 10: 50-54. Dijkman, M. 1950. Leucaena, a promising soil erosion control plant. Econ. Bot. 4:337-349. Isely, D. 1986. Notes on Mimosoideae (Leguminosae). Castanea 51(3) :202-206. Lamarck, J.B. 1783. Encycl. Method. Bot. 1:12. 4 National Academy of Science. 1977. Leucaena, Promising Forage and Tree Crop for the Tropics. Washington, DC. Zdrate, P.S. 1982. Las especies de Leucaena Benth. de Oaxaca con notas sobre la sistematica del género para México. Thesis. Facultad de Ciencias. U.N.A.M. México. 1987. Clarification of the Taxonomy of L. salvadorensis Standley ex Britton & Rose. Ann. Missouri Bot. Gard. 74(2), in press. A NEW SPECIES OF SABAZIA (ASTERACEAE, HELIANTHEAE) FROM DURANGO, MEXICO B. L. Turner Dept. of Botany, University of Texas, Austin, TX 78713 A forthcoming taxonomic treatment of the Asteraceae of Mexico (Turner and Nesom, in prep.) necessitates description of the following new species from Durango, Mexico. Sabazia gonzalezae B. L. Turner, sp. nov. S. liebmanii Klatt simile sed pedunculis brevioribus glanduli-pubescentibus et capitulis angustioribus campanulatis differt. Perennial herbs 40-50 cm high, the stems single and arising from slender nodose rhizomes, the root system fibrous at each node. Stems stiffly hirsute below, this grading upwards into a dense vestiture of glandular- trichomes. Leaves opposite, 3-5 cm long, 1.5-2.5 cm wide; petioles 2-5 mm long; blades ovate, 3-nervate, pubescent above and beneath, the leaves 6-8 dentate along each side, the apices acute to obtuse. Heads in a cluster of 3-4 in terminal corymbs, the latter on peduncles 9-11 cm long, the ultimate peduncles 0.8-2.0 cm long. Involucre campanulate, 3-4 seriate, more or less imbricate, 4-5 mm high, ca 6 mm wide; bracts ovate to elliptic, 3-5 mm long, 2-3 mm wide, somewhat scarious, the apices obtuse or rounded. Receptacular pales trifid. Ray florets 5-8; corollas white, the tube ca 2.5 mm long, densely pubescent, the ligules, 3-lobed, 5-8 mm long, ca 4 mm wide. Disk florets 20-30; corollas yellow, 4-5 mm long, the tube 1.0-1.5 mm long, the lobes ca 0.4 mm long. Achenes black, glabrous, striate, clavate, somewhat dorsally compressed, 2.0-2.5 mm long; pappus absent. TYPE: MEXICO. DURANGO: Mcpio. de Mesquital, 20 km de La Guajolota por al camino a Sn. Fco. Ocotan, 2440 m, 14 Nov 1985, M. Gonzalez et al. 1882 (holotype TEX; isotypes to be distributed) Additional specimen examined: DURANGO: Mpio. de Mezquital, 7 km al N La Guajolota, rumbo a Las Papas, 27 Aug 1985, I Solis 362 (TEX). Sabazia gonzalezae is related to the more southern S. liebmannii Klatt and, except for its slender rhizomes, would key to that species in Longpre's (1970) treatment 307 308 PHYTOLOGIHIA Vol. 63, No. 4 of the genus. McVaugh (1984), however, notes that the rhizomes of the latter may be thick or thin, which seems to be so, making the relationship between these two taxa fairly close. Within S. liebmannii the present taxon would key to the var. ovatifolia Longpre (confined to Oaxaca), largely because the leaves are less than twice as long as wide. Actually, S. liebmannii is a wide- ranging variable taxon and I can see little or no justification for the recognition of 4 varieties as accorded the species by Longpre. This might also be inferred from McVaugh's treatment of S. liebmannii, for he not only refers to Longpre's var. michoacana as a "form", he also sinks into synonymy under S. liebmannii the very different S. tridacoides Urbatsch & B. Turner. Actually the latter species is more closely related to taxa belonging to the recently resurrected Alloispermum (Robinson, 1979) than it is to Sabazia liebmannii. Unfortunately, McVaugh does not comment upon the generic problems relating to Alloispermum (a segregate from Calea) and I can only conclude that he has not understood the variation in this group, in spite of his observation that the chararcters of S. tridacoides ‘4are. mot Significantly different from those recorded for S. liebmannii." Inclusion of the present taxon in Sabazia is provisional. It might be that this genus will ultimately reside in synonymy under the older Alloispermum (Robinson, 1979). If so, however, it must relate to the "Sabazioid" species within this group and not to those centering about A. caracasanum (H.B.K.) H. Rob., its generotype. , its a pleasure to name this species for Dr. M. Gonzalez who has assembled a fine series of Asteraceae from the Mezquital region of Durango. ACKNOWLEDGEMENTS I am grateful to Dr. Guy Nesom for the Latin diagnosis and to Dr. Linda Vorobik for the excellent illustration. LITERATURE CITED Longpre, E.K. 1970. The systematics of the genera Sabazia, Selloa and Tricarpha (Compositae). Publ. Mus. Michigan State Univ. 4:287-381. McVaugh, R. 1984. Calea, in Flora Novo-Galiciana 12:187- 200. ; Robinson, H. 1979. Additions to Alloispermum, Galinsoga and Tridax. Phytologia 44:425-435. 1987 Turner, A new species of Sabazia 309 Fig.!. Sabazia gonzalezae, from holotype. TWO NEW FORMS OF HELIANTHUS AGRESTIS Eileen R.Wolde Helianthus agrestis Pollard f. almae E. R. Wolde Forma haec ab forma typica floribus solum usque 2 cm latis, dimidio inferno ligularum luteo, superno rufo-fusco, stigmatibus virescenti-luteis, foliis caulibusque juventute viridibus, ante anthesin purpureo- rufis, vetustate fuscescentibus, caulibus non glaucis differt. This form differs from the typical form by the flowers only to 2 cm wide, by the lower half of the ligules yellow, the upper red-brown, by the stigmas greenish-yellow, by the leaves and stems in youth green, before anthesis purple-red, with age darkening, by the stems not glaucous. This new form was collected by the author on November 2, 1986, 460 feet southof Bayshore Road, half-~- way between Crescent Lake and Magnolia Drives, North Fort Myers, Lee County, Florida, and is deposited in the Lundell Herbarium (L.L.) at the University of Texas, Austin, Texas. This sunflower is named in honor of Alma Molidenke, teacher, author and co-publisher of Phytologia for the past 35 years, witha lifelong dedication to biological education and to environmental and conservation causes. Helianthus agrestis Pollard f. oswaldii E.R. Wolde Forma haec ab forma typica floribus solum usque 25 mm latis, stigmatibus pallide viridibus, foliis cauli- busque primo in juventute viridibus, tandem purpureo- rufescentibus, caulibus non glaucis differt. This form differs from the typical form by the fiowers only to 25 mm wide, by the stigmas pale green, by the leaves and stem first in youth green, then be- coming purple-red, by the stems not glaucous. This new form was collected by Frederick W. Oswald, in whose honor it is named, on October 19, 1986, 440 feet south of Bayshore Road half-way between Crescent Lake and Magnolia Drives, North Fort Myers, Lee County, Florida, and is deposited in the Lundell Herbarium (L.L.) at the University of Texas, Austin, Texas. 310 BOOK REVIEWS Alma L. Moldenke "SYMBIOSIS - An Introduction to Biological Associations" by Vernon Ahmadjian and Surindar Paracer, xii + 212 pp., 70 black/white fig. incl. 62 photo., University Press of New England, Hanover, New Hampshire 03755. 1986. $32.50. Herein symbiosis is interpreted as first defined by H. Anton de Bary in 1879 as a living together of different types of organisms to include not only mutualism but also commensalism and parasitism. Such incidents occur widely in all five kingdoms - monerans, protoctists, fungi, plants and animals - from the invasions of some monerans into others to produce particulate cells with chloroplasts and mitotically dividing nuclei, to lichen associations, to alternate hosting of parasites and pathogenic relationships, and to elaborate co-evolutionary development of certain insect pollinators and specialized floral development. This book should encourage the teaching of courses in symbiosis. The language of this excellent text is clear, the concepts effectively developed, the reading stimulating, the reference sources and illustrations well chosen and the end of chapter review questions cover the topics meaningfully. There is a very well organized appendix on "Historical Landmarks in Symbiosis". In a dream I am readying to ask the head of department to let me teach such a course with this text. "FLORA HAWAIIENSIS Book 7 or New Illustrated Flora of the Hawaiian Islands" by Otto Degener and Isa Degener ca. 300 pp., 80 black/white botanical plates, 1 map, 4 pl + 7 botanist black/white photos. Privately published by the authors in Volcano, Hawaii 96785, 1986. $10.00 paperbound. It is wonderful that the authors are able to present serially their life-time botanical studies in this very convenient form of general and historical descriptions, family descriptions and keys to genera, species and subspecific units, special tributes with photographs to such leading Hawaiian botanists as Marie C. Neal, and a Hawaiian Plant Names List with the botanical and English equivalents prepared also by Noah Pekelo, Jr. The binding breaks easily - intentionally - so that the flora pages which are already provided with three looseleaf holes can be rearranged phylogenetically, alphabetically with the previously published parts or however is most efficient for the student or scholar of Hawaiian botany. We Moldenkes, along with innumerable other botanists, are grateful for the authors' long time friendship and their valuable scientific contributions. sil 312 PHY f.0-L 0G TA Vol. 63, No. 4 "IKEBANA - FRUITS AND VEGETABLES" by Toshie Yokoi, 104 pp., 82 color photo. and 4 black/white fig. Charles E. Tuttle Co., Inc. Rutland, Vermont 05701-0410. 1987. $24.95. The gifted artist-author is married to a professor of agriculture and is a pioneer in using farm and field flowering and fruiting products for ikebana so that village folks in the agricultural and rural areas can develop and express their appreciation for the beauties of nature. There are depicted exquisite arrangements of sunflowers and vine tomatoes, sweet potatoes and their attached vines, brussel sprouts and daffodils, and colored small rice cake balls fastened onto dogwood twigs. These and the many others illustrated in beautiful color photography display every bit as much beauty as the florist- purchased plants in the more usual forms of ikebana. Mrs. Fay Kramer, president emeritus of Ikebana International, could have been even more helpful in checking the English. "PLAIN OL' CHARLIE DEAM - Pioneer Hoosier Botanist" by Robert C. Kriebel x + 183 pp., 19 black/white photos. Purdue University Press, West Lafayette, Indiana 47907. 1987. $12.95 paperbound. The author is an appreciative family friend of this self- taught Hoosier botanist who logged more than 100,000 miles in his Native state during his field collecting and studying and who provided more than half of the 130,000 herbarium specimens in the Indiana University Deam Herbarium. Deam's "Flora of Indiana" is a masterpiece admired by so many field and taxonomic botanists. These are the climaxes of Deam's long, interesting, vital life which the author describes so naturally and effectively over the years along with the ideas, current events, and people from all walks of life - amateur naturalists, professional botanists, foresters, politicians, family that interacted with hin. "THE ADVENTURES OF CHARLES DARWIN" by Peter Ward, 108 pp., 14 color drawings, 1 color map, 11 black/white drawings, Cambridge University Press, Cambridge and London, U. K. and New York, N. Y. 10022, clothbound 1982, $10.95 & paperbound 1986 $5.95. "This book is a (juvenile) adventure story based closely on the diaries of Darwin's exploits on his famous round-the-world voyage. The young Charles Darwin was a likeable bold person with a great thirst for travel and new experiences." It is through the eyes and ears of an invented raw likeable cabin boy George that readers learn of many of Darwin's observations, collections, temporary land trips and cogitations aboard H.M.S. Beagle starting in 1831 and lasting for five years. Even adults would like the book for quick reading. Inasmuch as we do no editing, papers accepted for publication must be submitted in exactly the form that the author wants to have them published. They will then be photographed and printed by photo-offset in exactly the form as submitted except that we will add page numbers and running-heads. Typescripts should be prepared single-spaced on clean white heavy bond smooth and opaque paper. Elite type is probably the most space-economical. Typescript text must not exceed a rectangle 5° inches wide (horizontal) by 8% inches high (vertical), not including the running-head and page number. The title of the paper should be typed in all uppercase (capital) letters with 2 blank lines above the title and one beneath; then the name of the author in ordinary upper- and lower-case letters, along with his address (if so desired); followed by 2 blank lines; then the first line of text. It is usually best to leave a blank line between paragraphs. All scientific plant and animal names and group names should be typed either in italic type (if available) or underscored. Any corrections in the text made by the author must be complete and neat as they will be photographed as they are. The finished typescript as submitted by the author will be reduced from the 85s x 552 inch size as submitted to 6% x 4 inches by the printer. It is therefore advisable to place a centimeter or millimeter scale on all text figures and plates included. Use a new heavily inked black typewriter ribbon and be sure to clean the type on the typewriter after each several pages of typing. Cost of publication at present is $12.00 US per page, with no subsequent rebates, but this rate may vary depending on inflation and costs, so it is best to inquire as to current rates. The page charges are due with the typescript and no paper will be published before payment is received in full. Each author will receive gratis a proportionate share of the printed copies remaining after paid subscrip- tions are filled, but if separates (reprints or offprints) are desired, these will be charged extra in accord with the current rate for offprints provided by the printer. The cost of all such separates ordered must also be paid for in advance at the time the typescript is sent. No orders for separates will be accepted later, nor can additions or corrections be accepted. Authors are asked to indicate in light pencil on the reverse side of each page of their typescript the page number so that no mistakes in sequence occur. All manuscripts accepted will be published in the next issue, so that the size of the numbers may vary greatly. A volume will contain 512 pages. The plan insures prompt publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain limitations of size and proportion. An extra charge will be made for halftones, depending on their size, as fixed by the engraver. Articles dealing with research in all lines of botany and plant ecology, in any reasonable length, biographical sketches, and critical reviews and summaries of literature will be considered for publication. "q : a “a j =" Te ORR SE eee ee Ee Ee a; >