a or AWS Y 7 / / EE re) PHYTOLOGIA An international journal to expedite botanical and phytoecological publication Vol. 63 September 1987 No. 5 CONTENTS ZDERO, C., BOHLMANN, F, & KING, R.M., Chemistry of the Barnadesiinae (Asteraceae) ...........2 cece cece cccccecs 313 ROBINSON, H., Some suggestions regarding the significance of chloroplast DNA variation in the Asteraceae ............. 316 SCHUSTER, R.M., & DAMSHOLT, K., Some new taxa of SMART DIMM RECs die ie LIP. fsa oie wale pins atp.bis aie Seve ov we 0% 0 325 OCHOA, C., Solanum longiusculus (Sec. Petota), nova ETE IY UELIMIN EN a0 aha pare as. seu sieais ete a wia/ace deo Piece nia 329 PASSINI, M.-F, & PINEL, N., Morphology and phenology ey I IRIAN Nts Goce a dis 'a wieie om Ga nee seb de Slee oe 331 PASSINI, M.-F, The endemic pinyon of Lower California: Pinus lagunae M.-F: Passini ..............2200ceeeeeeeeeee 337 ST. JOHN, H., Diagnosis of Delissea species (Lobeliaceae) from Kauai: Hawaiian Plant Studies 145 ................... 339 ST. JOHN, H., Diagnoses of Clermontia species (Lobeliaceae): Hawaiian Plant Studies 146 ................. 350 _ ST. JOHN, H., MEDEIROS, A.C., A Haleakala variety of . Lobelia (Lobeliaceae): Hawaiian Plant Studies 147 .......... 366 ST. JOHN, H., Diagnoses of Rollandia species (Lobeliaceae): Hawartan Piont Studies T4800. ce cet ence wenenss 367 ST. JOHN, H., Diagnoses of Panicum species (Graininae): Hawaiian Plant Studies 149 ............. 0c cece eee eens 368 TURNER, B.L., A new species of Perymenium (Asteraceae- Heliantheae) from Tamaulipas, Mexico .............++.+++: 396 FOOTE, M., The algae of New Jersey (U.S.A.). XIII Chlorophyta (Green Agae). D. Zygnematales (Zygnemataceae and Mesotaeniaceae) ..............+++++- 399 SANCHEZ V., PE., Mirtaceas Nicaraguenses I: Eugenia sanjuanensis sp. nov. ..........2...000eeeeeeeeees 402 SANCHEZ V., P-E., & ORTEGA T., L.M., Nueva especie de Eugenia L. (Myrtaceae) de Veracruz, Mexico ............. 404 ROBINSON, H., Studies in the Liabeae (Asteraceae). XVIII. A new species of Munnozia from Bolivia ................+.. 407 REED, C.F, New combinations required for the Flora of Central Eastern United States ............20cccccceceeees 410 Published by Harold N. Moldenke and Alma L. Moldenke 590 Hemlock Avenue N.W. Corvallis, Oregon 97330-3818 U.S.A. Price of this number $3.00; for this volume $16.00 in advance or $17.00 after close of the volume; $5.00 extra to all foreign addresses and domestic dealers; 512 pages constitute a complete volume; claims for numbers lost in the mail must be made immediately after receipt of the next following number for free replacement; back volume prices apply if payment is received after a volume is closed. CHEMISTRY OF THE BARNADESIINAE (ASTERACEAE) C. dero}, F. Bohlmann! and R. M. King? Institute for Organic Chemistry, Techpical University, D-1000 Berlin 12, F.R.G. Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560. Cladistic analysis [1] as well as chloroplast DNA systematics [2] has led to the suggestion that the subtribe Barnadeiinae of the Asteraceous tribe Mutisieae represents the most ancient surviving devergent element of the present-day members of the family. Separation of the subtribe from the Mutisieae has been suggested, and the group has been proposed as a subfamily [1]. Although the Asteraceae as a family is notable for a rich secondary metabolite chemistry, all repre- sentatives of the Barnadesiinae thus far surveyed have only triterpenes, a type of secondary metabolites (Table 1) which is widespread in all higher plant families. The aerial parts of nearly all studied species afforded lupeol, lupeyl acetate and further triterpenes which have not been investigated in detail. While most species of genera placed in the other subtribes of the Mutisieae contain characteristic compounds [4,5], some genera of the Gochnatiinae (Chimantaea Maquire, Steyerm. & Wurdack, Cyclolepis Don, Hyalis Don, Oldenburgia Less., Quelchia N.E.Brown, Stenopadus Blake, Stomatochaeta (Blake) Maquire & Wurdack, Urmenetea Phil., Table 2) also gave only triterpenes, but most representatives of the genera of this subtribe afforded sesquiterpene lactones [6]. It seems of interest in view of the proposed paraphyletic nature of this subtribe [2]. Table 1 - Investigated species of the Barnadesiinae Species (voucher and location in parenthesis) Barnadesia arborea H.B.K. (RMK 7762, Ecuador). B. dombeyana Less. (RMK 9047, Peru). B. polyacantha Wedd. (RMK 7476, Bolivia). IBRAR) am 9 OCT 19 1987 INE V\ PAE Cae BOTANICAL GARD B. wurdackii Ferreya (RMK 9259, Peru). j ore 313 314 PHY T-6-L3G/ GTA Vol. 63, No. Chuquiraga avallanedae Lorentz (RMK 9389, Argentina). C. erinacea Don (RMK 9436, Argentina). C. hystrix Don (RMK 9416, Argentina). C. jussievi Gmel. (Solomon 16358, Bolivia). C. oppositifolia Don (RMK 9404, Argentina). C. parvifolia (Griseb.) Hieron. (Solomon 16347, Bolivia). C. rosulata Gaspar (RMK 9417, Argentina). C. straminea Sandwith (RMK 9378, Argentina). Dasyphyllum diacanthoides [3]. and as Chuquiragua leucoxylon Poepp. ex Less. (Dr. Nufiez, Valdivia, Chile). D. sprengelianum (Gardn.) Cabrera (RMK 8045, Brazil). D. velutinum (Baker) Cabrera (RMK 8357, Brazil). Schlechtendahlia luzulaefolia Less. (grown from seeds, Montevideo). Details are reported elsewhere [7] and the triterpenes obtained were identified by comparing the 400 MHz ly NMR spectra with those of authentic compounds. Table 2 - Investigated species of the Gochnatiinae which have triterpenes. Achnopogon virgatus Maguire, Steyerm. & Wurdack (O.Huber 8690, Venezuela). Chimantaea mirabilis Maguire, Steyerm. & Wurdack (O.Huber 8577, Venezuela). C. similis Maguire, Steyerm. & Wurdack (O.Huber 8696, Venezuela). Cyclolepis genistoides Don (RMK 9357, Argentina). 1987 Zdero et al., Chemistry of Barnadesiinae 315 Hyalis argentia Don (RMK 9326, Argentina). Oldenburgia arbuscula E.Meyer C71) "54s Setth Africa). uelchia bracteata Maguire, Steyerm. & Wurdack (O.Huber 8678, Venezuela). Stenopadus sp. (O.Huber 9009, Venezuela). Urmenetea extracamensis Phil. (M. Silva, Chile, unpubl. ). Acknowledgements - We thank the Fonds der Chemischen Industrie, Frankfurt, for financial support, Dr. 0. Huber, Apartado 8040, Caracas 1080-A, Venezuela, Dr. J. Solomon, Herbario Nationale, La Paz, Bolivia and Dr. J. Nufiez, Universidad Austral de Chile, Valdivia, Chile, for plant material. REFERENCES l. Bremer, K., XIV. International Botanical Congress, Berlin 1987, Abstract p. 296. 2, Venesen. Ra Ko, balmez, J. Dal and Mach aet sis 0 He Jas XIV. International Botanical Congress, Berlin US87, sAbstract p. 27.5. 3. Hoeneisen, M. and Silva, M. (1986) Rev. Latinoamer. Quuaimseya los 4 Zdero, C., Bohimann, F., King, R. M., and Robinson, H. (1986). Phytochemistry 25, 509. >) 2dero, C., Bohlmann, F., King, R. Mo and Robinison, B. (1986). Phytochemistry 25, 2873 (and it. cited therein). 6. Bohlmann, F., Zdero, C., Schmeda-Hirschmann, G., Jakupovic, J., Dominquez, X. A., King, R. M. and Robinson, H. (1986). Phytochemistry 25, 1175 (and lit. cited therein). i Bohlmann, Fo, 2dero,. C.,. Kine. oR. Mand) Robin sion, H. (1984). Phytochemistry 23, 1979. SOME SUGGESTIONS REGARDING THE SIGNIFICANCE OF CHLOROPLAST DNA VARIATION IN THE ASTERACEAE. Harold Robinson Department of Botany National Museum of Natural History Smithsonian Institution, Washington, D.C., 20560. Recent discoveries of a unique inversion in chloroplast DNA in most Asteraceae has led to some speculation on a possible ancient evolutionary split in the family. The data involved offers interesting reenforcement for some ideas that have been developing during recent decades, but some detailed suggestions seem to conflict with structural evidence of relation- ships within one tribe involved, the Mutisieae. A slightly altered view is offerred here along with the suggestion of a possible correlated evolutionary factor in the chemistry of the plants. Jansen and Palmer (1987) describe an inversion in the chloroplast DNA of most Asteraceae that is unique to that family and which differs from the evident original form seen in other Dicotyledonous families. The families lacking the inversion are listed by Jansen and Palmer as Apiaceae, Araliaceae, Brunon- iaceae, Campanulaceae, Caprifoliaceae, Dipsacaceae, Goodeniaceae, Rubiaceae, Stylidiaceae, Valerianaceae, in addition to the Solanaceae. In contrast, 15 tribes or groups treated as tribes of the Asteraceae show only the inverted form of the chloroplast DNA in all of their genera tested by Jansen and Palmer: Arctoteae (3 genera); Cardueae (3 genera); Lactuceae (3 genera); Liabeae (2 genera); Vernonieae (4 genera); Eupatorieae (3 genera); Heliantheae (8 genera inclu- ding 2 Tageteae); Anthemideae (5 genera including Cotula and Ursinia); Astereae (5 genera); Calenduleae (3 genera); Inuleae (3 genera); and Senecioneae (3 genera). The interest centers on the tribe Mutisieae where three of the subtribes have the inversion: Gochnatiinae (4 genera); Mutisiinae (5 genera); Nassauviinae (3 genera); while one subtribe lacks the inversion: Barnadesiinae (3 genera). In its lack (of the inversion, the Barnadesiinae is like the other Dicotyledonous families, and an obvious initial con- clusion places that subtribe at the base of the evolu- tion of the presently known Asteraceae. There is no reason to doubt the validity of the findings of the DNA inversion by Jansen and Palmer, but there might be a question regarding the full extent of their evolu- tionary conclusions. 316 1987 Robinson, Chloroplast DNA variation in Asteraceae 317 The general suggestion by Jansen and Palmer (1987) that something in the Mutisieae is primitive within the Asteraceae finds support in other studies of the family. For decades the field has been pro- gressing from the crude assumption that the Heli- antheae is the primitive group in the family (Cron- quist 1955). Carlquist (1961) was first to suggest that the Mutisieae were at least coequal with the Heliantheae as a primitive element of the Asteraceae. Poljakov (1967) placed the series of tribes containing the Heliantheae, now known as the Asteroideae, in a more derived position in the family while the tribes including the Mutisieae, now known as the subfamily Cichorioideae, were shown as more primitive. Robinson (1981) has gone on to show that the Heliantheae are not even the primitive tribe in the subfamily Aster- oideae. Skvarla et al. (1977) noted that the pollen of the Mutisieae differed from that of most Asteraceae and most closely approached pollen of the probably rather closely related family Calyceraceae. Jeffrey (1977), in the same volume, noted the zygomorphic corolla form in the Mutisieae that caused him to regard the tribe as closest to the primitive form in the family. The findings of Jansen and Palmer (1987) reenforce this already growing body of evidence for primitiveness of the tribe. The problem of the evolutionary interpretation of Jansen and Palmer (1987) arises from the assumption that the one subtribe of the Mutisieae, the Barnade- Siinae, is actually so divergent from the remainder of the Mutisieae that all the other Cichorioideae and all the Asteroideae could have arisen from within the evolutionary gap (Fig. 1). The three genera of the Barnadesiinae tested, Barnadesia, Chuquiraga, and Dasyphyllum indeed have some odd characteristics. Dasyphyllum has no apical appendage on the anthers, a feature in contrast with the well-developed appendage characteristic of most Mutisieae. Nevertheless, Bar- nadesia and Chuquiraga of the same subtribe have an ordinary Mutisian apical appendage. Barnadesia seems almost unique in the whole subfamily Cichorioideae in the lack of spurred bases on the thecae of the anthers, but spurs like those of other Mutisieae are found in the subtribe in Chuquiraga and Dasyphyllum. Barnadesia is also unusual in the Mutisieae in the lophate form of its pollen (Fig. 2), but pollen of Chuquiraga (Fig. 3) and Dasyphyllum is much like that found in other subtribes of the Mutisieae (Fig. 3). The foregoing characters are mixed in the Barnadesi- inae with every unique character matched by an oppo- site condition within the subtribe. 318 PHYTOLOGIA Vol. 63, No. 5 ASTEROIDEAE Ww CICHORIOIDEAE Senecioneae Lactuceae Astereae Liabeae Anthemideae Vernonieae Inuleae Arctoteae Calenduleae Eremothamneae Heliantheae Cynareae Eupatorieae Mutisieae 3 subtribes Mt ad ~~ MUTISIEAE Barnadesiinae REST OF THE DICOTS CAMPANULALES Figure 1. Evolution of Asteraceae following proposal of Jansen and Palmer (1987). Groups below dashed line (B) with uninverted chloroplast DNA. Tribes above dashed line (W) with inverted chloroplast DNA. 1987 Robinson, Chloroplast DNA variation in Asteraceae 319 Figure 2. Barnadesia horrida Muschler, Pollen. 1. Coipar view. 2. Polar view. 3. Detail of crest. 4. Broken section of crest. 320 Pen Yr Or LO Gera Vol. 63, No.5 Figure 3. Chuquiraga jussieui Gmel. Pollen. 5. Polar view. 6. Colpar view. 7, 8. Broken edges of grains. Photographs of pollen prepared by Smithsonian Museum of Natural History SEM Laboratory. The microscope operated by Suzanne Braden. Stubs prepared by Mary Sangrey using facilities of Department of Botany Palynological Laboratory. 1987 Robinson, Chloroplast DNA variation in Asteraceae 32) One is not inclined to believe that the Barna- desiinae are so remote from other Mutisiinae that the remainder of the Asteraceae could evolve from within the gap. But there should be some explanation for the pattern of distribution of the chloroplast DNA in the family described by Jansen and Palmer (1987). The inversion cannot be regarded as a parallelism or the result of reversion. The detailed evidence of the DNA sequence offered by Jansen and Palmer precludes such a chance duplication. Jansen (pers comm.) has stated that the DNA segments are too large for transfer whole by viruses or bacteria, a conclusion that I must accept. One other mechanism, however, is possible. Hybridization could have the effect required, and hybridization is a common phenomenon in the Asteraceae where it seems to be a factor in the success of many tribes (Robinson 1983). The chloroplast DNA is trans- mitted cytoplasmically and is carried only through the female line which prevents any recombination with other cytoplasmic characters, but the maternal limit- ation does not prevent recombination with various characters transmitted in the nuclear DNA. It seems unavoidable that a chloroplast DNA inversion that starts in one chloroplast of one plant would initially become distributed unevenly in any derived population of sexually reproducing plants. The mixture could easily be perpetuated if the derived plants continued to be as capable of interspecific and even inter- generic hybridization as many present-day Asteraceae. The present assumption is that the more immediate descendents of the earliest Asteraceae with the in- verted chloroplast DNA carried the two forms mixed in their populations through many generations, and that the mixture spread across much of the initial diver- Sity of the family. It was from this mixed ancestral population that present Asteraceae were derived, but the older form of DNA was retained in only one sub- tribe of the comparatively primitive Mutisieae, the Barnadesiinae (Fig. 4). The conclusions given above accept the primitive Mature of the chloroplast DNA in the Barnadesiinae. An interesting question remains as to what other characters of the Barnadesiinae or Mutisieae might also be primitive. No others can be certain. The pollen of Barnadesia, although distinctive and present in the subtribe, is almost certainly not a primitive form. The more typical Mutisian type seen in Chu- guiraga and especially Dasyphyllum is more like the Calyceraceae (Skvarla et al. 1977). The lack of an anther appendage in Dasyphyllum also seems highly individual, and is unlike the vaste majority of the 322 PH Y.T 0 10 CL A Vol. 63, No. ASTEROIDEAE W CICHORIOIDEAE Senecioneae Lactuceae Astereae Liabeae Anthemideae Vernonieae Inuleae Arctoteae Calenduleae Eremothamneae Heliantheae Cynareae Eupatorieae Mutisieae vee 3 subtribes MUTISIEAE Barnadesiinae REST OF THE DICOTS CAMPANULALES Figure 4. Most probable evolution of variation in chloro- plast DNA in Asteraceae. Groups below dashed line (B) with uninverted chloroplast DNA. Tribes above dashed line (W) with inverted chloroplast DNA. 1987 Robinson, Chloroplast DNA variation in Asteraceae 323 Asteraceae of both presently accepted subfamilies. The appendaged condition seems to represent a cond- ition established in both subfamilies antidating the divergence of the Barbadesiinae. The nonspurred bases of Barnadesia might derive from ancestors that were more like the subfamily Asteroideae before the evol- ution of the trait in the Cichorioideae. The zygo- morphic bilabiate corollas noted by Jeffrey (1977) may be primitive as he has suggested. One particular set of characters found in the Barnadesiinae and many other Mutisieae offers an interesting possible insight into the early evolution of the Asteraceae. The habits of the Barnadesiinae are notably thorny as reflected in many of their names, Dasyphyllum ferox (Wedd.) Cabrera and Barna~- desia horrida Muschler. These plants and some other Mutisieae are also notable for their simple secondary metabolite chemistry compared to that of most other Asteraceae (Zdero et al. 1987). The impression is of a group that relies more on physical defenses instead of chemical ones. These plants are not the rich chemical factories that are seen in s0 many other tribes of the family. The simple chemistry seems very likely to be a survivor of a more primitive strategy in the Asteraceae. The simple chemistry of the Barnadesiinae raises one additional possibility. There has been little reason to assume that inversions of DNA sequences of the type seen in the Asteraceae necessarily have any Significant benefit. The success of the inverted form inthe family, however, might indicate that it is in some way favored. An actual positive benefit might not be involved but there may be a passive ability to better survive and function in a cell that has a richer chemistry. Cronquist (1977) has noted that a plant must be able to withstand its own repellents, and the unique inverted chloroplast DNA of most of the Asteraceae might be better able to withstand the pre- cursors of the numerous poisonous secondary metabo- lites of most tribes of the family. Even a slight effect could explain the success of the inverted form that is found in all the chemically richer members of the family. It is remotely possible that the more complex chemistry could not have evolved in the family without a less vulnerable inverted form of chloroplast DNA. 324 PSH Yo T*OrLy0 Ger A Vol. 63, No. Literature Carlquist, S. 1961. Comparative Plant Anatomy. Holt, Rinehart and Winston, New York. 146 p. Cronquist, A. 1955. Phylogeny and taxonomy of the Compositae. Amer. Midl. Naturalist 53: 478-511. - 1977. On the taxonomic significance of secondary metabolites in Angiosperms. Plant Syst. Evol., Suppl. 1, 179-189. Jansen, R. K. and J. D. Palmer. 1987. A chloroplast DNA inversion marks an ancient evolutionary split in the sunflower family (Asteraceae). Proc. Natl. Acad. 84(16): 5818-5822. Jeffrey, C. 1977. Chapter 6. Corolla forms in Compo- sitae-some evolutionary and taxonomic specula- tions. 111-118 In V. H. Heywood, J. B. Har- borne, and B. L. Turner, eds. The Biology and Chemistry of the Compositae. Academic Press, London, New York. Poljakov, P. P. 1967. Sistematika i proischozdenie sloznocvetnych. Alma-Ata: Nauka. Robinson, H. 1981. A revision of the tribal and subtribal limits of the Heliantheae (Asteraceae), Smithson. Contr. Botany 51: 1-102. - 1983. A generic review of the tribe Liabeae (Asteraceae). Smithson. Contr. Botany 54: 1-69. Skvarlsa, Ji‘ J., 3B. .L. Turner, Vv. Cc. Patel sand sane. Tomb. 1977. Chapter 8 Pollen morphology in the Compositae and in morphologically related families. 141-248 In V. H. Heywood, J. B. Harborne, and B. L. Turner, eds. The Biology and Chemistry of the Compositae. Academic Press, London, New York. Zdero, C., F. Bohlmann and R. M. King. 1987. Chemistry of the Barnadesiinae (Asteraceae). Phytologia 63(4): 313-315. SOME NEW TAXA OF JUNGERMANNIALES R. M. Schuster Cryptogamic Laboratory Hadley, Mass. 01035 and K. Damsholt Institute of Systematic Botany Copenhagen, Denmark I. New Taxa from South Greenland During the course of a joint expedition to South Greenland in the summer of 1982 we discovered a number of new taxa of Hepaticae. Although these are described at length, and illustrated, in a gener- al account by the senior author, in The Hepaticae of South Greenland [Nova Hedwigia, Beiheft 92, 1988], no Latin diagnoses are given in that work. In order to validate these taxa, the following brief La- tin diagnoses and type citations are offered. In addition, a brief Latin diagnosis of Scapania pseudocalcicola Schust. is appearing here. The basis for this new species is dis- cussed below. Lophozia hyperborea ssp. helophila Schust. & Damsh., ssp. n. Subspecies a ssp. hyperborea differens (a) praesentia pigmenta- tionis typi anthocyanini; (b) amphigastriis parvis minutisve; (c) praesentia gemmarum. Type. Tasiussaq Fjord (RMS & KD 82-1667). Schuster and Damsholt (1972) reported L. hyperborea first from South Greenland, on the basis of a single collection; the numerous collections we made in 1982 show rather conclusively that the South Greenland plants differ in both pigmentation patterns and in their ability to form gemmae from the species proper. Lophozia rubescens Schust. & Damsh., sp. n. Species a L. hatcheri differens (a) praesentia pigmentationis anthocyanini; (b) amphigastriis reductis; (c) cellulis magnis, 28-36 um in marginibus loborum; (d) praesentia guttarum olei (5-7)9-22(26) in omni cellula; (e) gemmis raris, 21-29 x 31-46 um. Type. Amit- suarssuk Fjord (RMS & KD 82-1192). L. rubescens is the fourth species in subg. Barbilophozia. Al- though there is some variability in cell size, this taxon clearly has much larger average-sized leaf cells than the other three taxa of the subgenus. In the small underleaves of mature plants L. rubescens is similar to the otherwise remotely allied L. barbata. Possibly a 325 326 Pol Yr 0710 Galva Vol. 63, No. 5 polyploid derivative of L. hatcheri or of L. lycopodioides is at hand. The often striking reddish pigmentation separates it from both. Lophozia debiliformis Schust. & Damsh., sp. n. Species L. elongatae (subg. Protolophoziae) atque L. alpestri (subg. Lophoziae) cognata. Distincta a prima: (a) inflorescentiis dioeciis; (b) praesentia gemmarum rubellarum; (c) cellulis (1)2-5 (6-14) guttas olei habentibus. Distincta a secunda: (a) cellulis multis solum (1)2-3(4) guttas olei permagnas habentibus; (b) amphi- gastria conspicuis; (c) foliis angulariter, saepe profundissime bi- lobatis. Type. Kangikitsoq Fjord (RMS & KD 82-1798). L. debiliformis includes a variable ensemble of very small Lo- phoziae that, on one hand, show similarities to L. longidens, on the other, to L. alpestris. The infinitely variable ensemble of pheno- types aggregated under this name has been the source of major diffi- culties. Virtually all phenotypes develop distinct to conspicuous underleaves. Lophozia debiliformis var. concolor Schust. & Damsh., var. n. Varietas a debiliforme typica distincta: (a) amphigastriis spo- radice permagnis, 350-700 um long.; (b) ora perianthii crenulato- denticulata, dentiis adsummum unicellulis. Type. Christian IV's 9, Tasiussaq (RMS & KD 82-1645). This paradoxical taxon, initially believed to be an autonomous species, seems to be a luxuriant extreme of L.debiliformis. How- ever, additional collections may show that an autonomous species is at hand. The collections came from a unique habitat in which var. debiliformis was not seen: the fine silt laid down from ice melt from upstream glaciers. Lophozia bicrenata var. immersa Schust. & Damsh., var. n. Varietas a var. bicrenata distincta: (a) pigmentatione pro- funde subpurpureo-brunnea; (b) cellulis leptodermatis etiam in lo- bis, omnibus (5)6-10(11) guttis olei habentibus, lamella media con- spicua atraque. Type. Kangikitsoq Fjord (RMS & KD 82-1810). Although the color, compactness and aspect recall L. excisa var. succulenta, which it was believed to be in the field, the perianth mouth, distinctly toothed, is as in L. bicrenata. It differs from phases of the latter in the thin-walled cells with fuscous middle lamellae, and in the retention of oil-bodies in all leaf cells. Marsupella spiniloba Schust & Damsh., sp. n. Species magnitudine et inflorescentiis paroeciis M. sprucei similis; distinct, autem: (a) marginibus foliorum revolutis; (b) lobis foliorum acute apiculatis; (c) foliis surcularum sterilium 1987 Schuster & Damsholt, Jungermanniales 327 pectinato-distichis. Type. Qornoq, Frederiksdal (RMS & KD 82-1297). M. spiniloba is distinct from its near ally, M. sprucei, in the non-capitate shoots; in the pectinate-distichous sterile shoots; and in the sharply apiculate leaf lobe apices. Cephaloziella mammillifera Schust. & Damsh., sp. n. Plantae autoeciae sed pseudodioeciae (gametetangiis 6 atque in axibus late remotis); amphigastria axium sterilium minuta aut vestigialia; folia basim versus 2-stratosa et superfacies abaxiales localiter mammillosae; caulis localiter armatus; cellulae oris peri- anthii (3)3.5-6(7):1. Type. Frederiksdal (RMS & KD 82-1040). This puzzling plant, which we could not “wedge” into any other, is allied to C. uncinata, but has the stem locally armed with cellu- lar protrusions and the abaxial surface of the bistratose leaf bases bear tumid or mammilliform abaxial surface protrusions. Cephaloziella uncinata var. brevigyna Schust. & Damsh., var. n. Varietas a varietate uncinata distincta: (a) lobis foliorum 8- 10(11-12) cellulis latitudine; (b) cellulis majoribus, 13-17 um lat. ad bases loborum; (c) lobis foliorum atque bractearum fere numquam hamatis. Type. Qornoq, S. of Frederiksdal (RMS & KD 88-1258). The var. brevigyna is a puzzling plant, assigned with some mis-— givings to C. uncinata s. lat., from which it differs in the wider leaf lobes, the larger leaf cells, the non-hamate lobe apices of bracts, and the tendency for gynoecia to form on abbreviated inter- calary axes. Cephaloziella byssacea var. polystratosa Schust. & Damsh., var. n. Varietas a var. byssacea distincta quod (a) folia 2(3)-stra- tosa; (b) folia latissima, lobis saepe 13-15 cellulis latitudine. Type. Ordlerit (RMS & KD 82-2132). The var. polystratosa is the only one of the innumerable vari- ants of C. byssacea in which leaves are 2(3)-layered in a large ba- sal field. They bear an analogous relationship to typical C. bys- sacea as L. opacifolia does to L. incisa and, possibly, should be regarded as forming an Arctic subspecies. II. A New Species of Scapania subg. Kaalaasia Scapania pseudocalcicola Schust., sp. n. Species S. calcicolae similis, differens valde, autem: (a) ora perianthii tenuiter dentata, dentis humilibus 1(2)-cellulis; (b) perianthio vix plicato. Type. Newfoundland: N. of Daniels Harbour, N. Pen. (RMS 68-1454). 328 PHYTOLOGIA Vol. 63, No. 5 S. pseudocalcicola was attributed in Schuster (1974, p. 316) to S. calcicola (Arn. & Perss.) Ingham. It differs from this in the short-serrulate to denticulate perianth mouth (cf. Schuster, l. c., figs. 358:4, 359:7-8). At the time this attribution was made the perianth of true S. calcicola was unknown, although that of ssp. ligulifolia (Schust.) Damsh. & Long had been shown to bear a lobu- late-dentate mouth, much as in S. gymnostomophila (cf. Schuster, 1. c., fig. 356:2, 13-16). Damsholt and Long (1979) assumed that the minute, low teeth of the perianths of S. pseudocalcicola (S. calci- cola in Schuster, 1l.c.) were due to poor development, associated with lack of fertilization. This assumption foots on an error: all taxa assigned to subg. Kaalaasia are known only from plants with un- fertilized gynoecia. Plants figured in Schuster (l.c., figs. 356:2 and 358:4) are from populations where no @ plants grew nearby; both bear unfertilized perianths. Although even in the absence of ferti- lization the perianths show some growth subsequent to archegonium maturation (and decay) it is very rare to see perianths develop to the point sometimes seen in S. gymnostomophila (cf. fig. 354:4 in Schuster, l.c.). Furthermore, as is very well documented, perianth maturation is via a basal meristem: even on the youngest perianths the apices undergo maturation, whereas the basal meristem copiously proliferates cells only subsequent to fertilization. Assuming that if fertilization had occurred, the elaboration of teeth of the per- ianth mouth would have continued, has no basis in fact: in all three of the species of Kaalaasia for which perianth mouths are shown in Schuster (l.c., figs. 354:5, 356:2, 13-16, 358:4, 359:7- 8), comparable stadia in development are illustrated. Acknowledgement: We are indebted to Dr. Hannah Croasdale for her aid with the Latin diagnoses. References Damsholt, K. & D. G. Long. 1979. The perianth of Scapania calci- cola (Arn. & Perss.) Ingham (Hepaticae) and the relationship to Scapania ligulifolia (Schust.) Schust. Lindbergia 5(2):73-76. Schuster, R. M. 1974. The Hepaticae and Anthocerotae of North America Vol. III, pp. i-xiv, 1-880, figs. 302-475. Columbia Univ. Press, N.Y. Schuster, R. M. & K. Damsholt. 1972. Lophozia (Orthocaulis) hyper- borea (Schust.) Schust. in southwest Greenland. Lindbergia 1 (3-4):166-68, 1 map. Schuster, R. M. & K. Damsholt. 1974, The Hepaticae of West Green- land from ca. 66°N. to 72° N. Meddel om Grgénland 199(1):1-373, figs. 1-33, 80 maps. SOLANUM LONGIUSCULUS (SECT. PETOTA), NOVA SPECIE PERUVIANA by C. Ochoa* Herbaceum, tuberiferum, 70-80 cm altum, ramificatum. Caults erectus, tenuis, basi, 5-7 mm crassus, glabrescens, pilis sparsissimis vix perspiciendis, conspersus; internodia, 6-8(-10) cm longa. Stolones 150 cm vel plus longi, tubercula parva, 1.0-1.5 cm diam., alba. Folia caule anguste decurrentes 16-20 cm longa, 7.5-10.0 cm lata, 5-juga et 10-14 foliola interjecta, parvula et sesstlia, rhachis pilis brevibus obtec- tus, pili simplices cum pilts glanduliferis tetralobulatis intermixti. Foliola, 4.0-4.5 cm longa, 1.3-1.6 cm lata, anguste elliptico-lanceolata, apice acuta, basi tnaequilatera, rotundata, petioluli 3-5 cm longi; foliola supra plus minusque dense pilosa, pilts brevibus in petiolulis densioribus, subtus dense pilosa, praesertim in venis venultsque. Foliola pseudostipulacea late subfalcata, 8-12 mm longa, 4-6 mm lata. Inflorescentia cymosa, 7-10(-14)-flora. Pedunculus 10 cm longus, bast 1.8 mm crassus puberulentus tam quam pedicelli et calyx. Pedicelius, 20-25 mm longus, prope calycem articulatus; pedicellus superior almost 4-5 mm longus. Calyx symetricus vel non, 7-8 mm longus, lobi anguste elliptico-lanceolatt, apice acuminati, acumina, 2.0-2.5 mm longa, acuta. Corolla rotato-pentagonalis, 3 cm diam., azureo-violacea. Columna antherarum cylindrico-conica, antherae, 5.5 mm longae, basi cordatae, filamenta, 0.5-1.0 mm longa, glabra. Stylus 10 mm longus, glaber; stigma parvum, ovale-compressum. Baccae ovales, 15 mm longae, viridae, vel cum, 1-2 stris longttudinalibus violaceis ornatae. Ad seriem Tuberosa pertinet. Numerus cromosomatum: 2n = 2x = 24. Typus: PERU, departamenti Apurimac, provincit Grau, Ranra, 3400 m supra marc, in itinere Ayrithuanca-Pamparackay. C. Ochoa 4125, Martius 1973 (holo- typus, OCH; isotypus, US). Affinitas: Habitu, pedunculi longi et multiflori, et corolla forma et colore affinitatem cum S. marinasense habet, sed valde diversa videtur speciatim forma et disectio folio- rum, forma et colore fructus. Habitat: In loca frigidis, puna et subpuna, 3400-3500 m alt., ad saxa tn locis humidis, inter arbusculus Cassia et herbac erectae Cajophora. * International Potato Center, P.O. Box 5969, Lima-Peru 329 330 PaHey £0) LOG aiaA Vol. 63, No. 5 Solanum tongiusculus Ochoa. Holotypus OCH-4125, ca. x 1/2 MORPHOLOGY AND PHENOLOGY OF PINUS LAGUNAE M.-F. Passini Marie-Francoise Passini and Nicole Pinel Université Pierre et Marie Curie 12 rue Cuvier 75005 Paris Pinus laguaae(Passini, 1987) was described, (Robert-Passini, 1981), as “a tree with an upright trunk, 12-15 meters high, leaves principally 3, sometimes 2 per fascicle, grey green; stomates on each surface; two resin ducts. Conelets slightly pedunculate. cones globular or sub-globular.” It is found in the Sierra de La Laguna, in the far southern tip of the Lower California Peninsula. This study describes the morphology and phenology of Praus /agunde . METHODS Several morphological characteristics were studied in growing trees in various localities : La Laguna, San Antonio, San Juanito on the eastern slope, experimental plot (1 hectare), Palo-Extrafo, La Chuparrosa and San Francisquito (Tab. 1). Tab. 1 - Geografic origin of the trees Site Longitude Latitude Elevation N‘of Trees La Laguna 110°58 23°33 1700 m 1-2-3-13 Picacho 110°01° 233% 1900 m 4 San Antonio 109° 58° 23° 32° 1950 m 5 Palo-Extrano 109°57' 23°31 1650 m 14-15 -16 Experimental 109°56' 23°31 1756 m 6-7 San Juanito 109°55' 23°31" 1400 m 8 La Chuparrosa 109°55' 23°31 1725 m 9 San Francisquito 109°57' 23°29 1400 m 10 For each tree, the eight and diameter of the trunk, the eight of the first sensitive branch, the length of growth units in 1982, 1983, 1984 (4 terminal branches from the main and lateral axis) were measured. Number of needles (300 to 1300 fascicles by tree), needle length (20 needles from each main and lateral axis) were measured. Height, weight, number of fertiles scales, number of two seed scales were noted for 30 matures cones by tree The length, width of seeds and seed wall thickness were measured (20 seeds by tree). A phenological study of trees from the two localities of La Laguna and plot experimental was made. Morever, four lots of seeds from La Laguna were put to germinate in november and december 1984, in april and may 1985, in identical conditions Heri covered Petri boxes, kept moist with distilled water, temperature 25 + 331 332 Poo Y -1)0"%L70°G) TA Vol. 63, No. RESULTS Trunk and branches Pines can grow from 3 to 21 meters. The pine height and the height of the first needle clad branches in La Laguna are significantly different when compared to heights of the experimental plot. No significative difference is found between diameter of trees in the two localities. La Laguna pines, growing 6,7 to 8 meters apart, are not so tall as those of the experimental plot, but have a more widespread habit. On the contrary, the more closely packed trees on the experimental site (3 to 5 meters apart) have a taller trunk and a sparser crown. Finally, pines in southern and south westerly localities are not so tall as those growing in the afore mentioned localities In 1983, the average branche length growth was 6, 9 - 2,3 cm and 6,3 - 1,5 cm in 1984. The difference between 1983 and 1984 is not significant in any locality . However. the difference between length from one locality to another is significant (Tab. 2). In particular, branches from trees in La Chuparrosa and San Juanito revealed far less growth than pine branches from La Laguna. Three localities : La Chuparrosa, San Juanito and San Francisquito, have a south westerly exposure, therefore with more sun and also drier soil. Tab. 2. - Length of growth units : comparison of La Laguna pines with other localities pines Site or N° of trees 1983- Significance” 1984 Significance* La Laguna 6.9 + 23 63+ 15 4 6,1 + 07 ns 78 + 1,3 ns 5 3,6 + 0,3 ns 46+ 05 0,05 6 49 + 01 ns 55 + 00 ns 7 6,0 + 06 ns 26 + 0,7 ns 8 50 + 2.1 ns 39+ 1,1 0.05 9 41 + 0.4 ns 33+ 0.4 0,001 10 40 + 00 0,05 24+ 00 0.001 * Student-test Foliage characteristics. No significative difference appears between year to year length of needles from fascicles of the main axes : Year Length in cm Number of fascicles 1982 45 480 1983 5.9 820 1984 5.5 820. However. a comparison of average needle lengths of the pines of San Francisquito and La Chuparrosa with those of La Laguna reveals a significant difference. A positive correlation exists between growth unit lengths (x) and needle lengths (y) for 1982 and 1983 on the one hand, and for 1984 on the other. 5 1987 Passini & Pinel, Pinus lagunae 333 The equation differs according to whether the branche belongs to a main growth axis; y =3,37+0,31 x r=0,54 DL=25 or to a lateral axis . y = 3,37+0,32x r=055 DL=26 The growth is therefore greater on the main axis than on lateral axis. Mature pines have communly 3 needles per fascicle, a smaller number has 2 needles per fascicle, and a few four needles, one needle per fascicle are un commun. In all mature trees, in 1983, there are 91% three needles per fascicle and merely 82 %, in 1984. Therefore, the number of two or three needles fascicles varies from year to year. Moreover, there is a higher percentage of two needles per fascicle in San Francisquito than in La Laguna (Tab. 3). Tab. 3. - Comparison of needle lengths and number of needles per fascicle of La Laguna pines and other localities pines Site or Needle length Percent 2 needles/ fascicle N° of trees (mm) 1983 S* 1984 S* 1983 S* 1984 S* LaLaguna 6,3 :1,1 58 +08 3% 4% 6 41 +03 005 46:0 0,05 7 ne 8 ns 7 6 +02 as 5,1 + 0,3 ns 0 ms 6 as 8 46 +1 ns 5.1 +03 ns 2 ey 4 ns ) 46 +0,4 001 44 +03 0,01 15 ns 25 as 10 46 +0 005 43:90 005: ,22 001 48 001 * Student-test (S=significance) A positive correlation (logaritmic) links up the annual growth unit and the percentage of three needle fascicles. Lagunae pine needles have ventral and dorsal stomatal lines : from one line to three lines on the dorsal surface, and four to six lines on the ventral surface. The ratio between surface lines (FV) and dorsal surface lines (FD) is slightly higher in localities south of the Sierra de La Laguna (San Francisquito and San Juanito). But no positive correlation appears neither between FV/FD and needle length nor FV/FD and percentage of needles per fascicles. The needles grow longer in June and are fully grown in october or november, they are a three year life span. Cones and seeds characteristics. The pedoncule of the seed cones are 0,2- 1,2 cm long. They are decidues. A mature seed cone is 3-6 cm long (mean length 3,9+0.5 cm). The scale number is about 9. Only median and upper scales bear seeds. In all the cones, 63% of scales have 2 fertile seeds and 37% one seed. The seed cones from San Francisquito are smaller than those of La Laguna and have fewer two seed scales (Tab. 4). 334 Paha er OL {0 (Car vA: Vol. 63, No. Tab. 4- Cone characteristics, comparison of pines of La Laguna with those of Palo-Extrano (14,15,16), experimental plot (6,7) and San Fran- cisquito (10) Site or Number N° of trees LaLaguna 228 14 31 15 13 16 21 6 21 7 31 10 23 Mean Cone length S* (cm 43 +05 15 +06 3.9 +04 44 +0.4 4,3 +05 46 +05 3,2 +03 3,9 +05 * Student-test (S=significance) ) 0,001 ns 0,001 ns 0,05 0.001 2 seeds scale 6,6 + 3.2 S* Pinus laguaae has large seeds coffee coloured or ligth brown mottled with dark brown. Mean length seed of all the localities are 12,9 + 1,3 mm long, 7,6 + 0,5 mm width. The shell seeds have 0,5 + 0.2 mm thick. Seeds from La Laguna are significantly longer than all of Palo-Extrafio, experimental plot, and San Francisquito. Moreover, certain trees of this localities produce seeds with thinner shelles than those of La Laguna (Tab. 5). Tab. 5.- Seed characteristics Site or Seedlength S* N° of trees (mm) LaLaguna 138 +1,3 14 12,445 15 13,1 + 0,7 16 15,0 +1 7 13,1 +0,7 10 115 +0.4 Mean 12,9 +12 0,001 0,02 0,001 0,02 0,001 * Student-test (S=significance) Seed width 5S* (mm) 7.9+0.9 84+05 0,02 74+06 0,02 § +05 aos 74+06 0,02 7.3206 0,001 76+05 Thickness shell 5S* (mm) ees s Germination. The four lots of seeds from La Laguna proved to have a high germinative capacity :90% germination within a five day response delay (Fig 1). These seeds, as those of Pinus catarinae (Passini, 1981), display no dormancy period. The mean number of cotyledons is 12,3 (150 germinations). 1987 Passini & Pinel, Pinus lagunae 335 Fig. 1.- Seed germination Cumulated frequencies % 100 7) Nov 84 @ 50 seeds Dec 84 A 68 seeds 30 Avr 85 0 41 seeds 25 Mai 85 4 50 seeds A Number of days b. Zura eee 9 Oa In the mountain, seeds germinate from September onwards and. germination percentage is also very high. Seeds were sown on the spot. in the forest, but only 30% of the initial plantlets survived after six months. March had the harviest losses. Phenology. The vegetative buds, that enter in dormancy in November, become active in late March At bud bursting time, the bud, 1-2 cm long, swelles at the base and the scales open up. The terminal bud stretches and lengthens during April, forming an inflorescence which will be fully developed at the end of May. Male branches grow will not begin until after pollination, at the end of June Female strobili can be seen as from late April onwards. Their scales are open mid May. In 1985, pollination began towards 20th May and carried on until 10th June. A slight delay in the dehiscence of male inflorescences was observed in La Laguna, compared with those in Palo-Extranio All the trees observed, in 1985, in the collecting localities of La Laguna and the experimental plot flowered Male inflorescences, generally abundant, were on the lower branches. On the contrary, female strobili developed in the upper part of the tree, although some were occasionally found on the lower branches. Two trees, three meter tall, about 30 years old (age determined by number of verticils) only bore female strobili Seed cones begin growing in March during the second year, they are ripe in August. In late August, early september, the hanging cones open and Jet fall their ripened seeds. Heavy late summer hurricanes often speed up the process, hurtling cones and seeds to the ground. Small mammals, such as Peromyscus trues, the “raton de los pines’, and many birds feast upon these seeds The birds include As/oma serrana and Malanerpes formicivorus which build up large stocks of these seeds inside dead tree trunks 336 PHY r O-L, 0 Cl A Vol. 63, No. CONCLUSION In this study, carried out in Sierra de la Laguna, from October 1984 to July 1985 (Pinel, 1985), it appears that morphological characteristics, especially the number of needles per fascicle. vary throughout the Piaus /agunae formation. Apart from varying number of needles per fascicle from one specimen to another in La Laguna, previously indicated by Passini (1981), it was noted that the number of needles per fascicle increases with altitude, between 1 400 and1 700m. Also, the positive correlation betwen annual growth unit and the percentage of three needles per fascicle has to be noted. We also draw attention to the fact that very long needles (longer than the average needle length of Piaus cembroies Zucc., Zavarin & Snajberk, 1986)) are to be found at La Laguna, and shorter needles in southern collection localities Characteristics of description type : three needles per fascicle, ventral and dorsal stomatal lines, average number od cotyledons (12,6) were confirmed But the average heigth oftrees is greater than that given in the description. Moreover, considerable variation was noted at the San Francisquito locality which is further south than that of La Laguna : the trees, there, are smaller, but at present, our studies do not eneble us to say whether this variation affects all the trees of the locality. Two types of seeds were found : a thin shell seed of 0.2 to 05 mm anda medium thick shell 0,5 to 0,9 mm. This study enable us to conclude that two varieties of Piaus /eguane . Further studies will specify their relations and ecological requirements. REFERENCES Passini M.-F., 1987. - The endemic pinyon of Lower California . Pinus /egunae M.-F. Passini. 1987. Passini M.-F., 1981.- Les foréts de Axaus cembroides au Mexique, étude phyto- géeographique et écologique. Ed. Recherches sur les Civilisations, 374 p. Pinel N., 1985. - La formation a Piaus cembrosdes var. Jegunae dans la Sierra de la Laguna, Basse Californie, Mexique. Rapport stagede DE.A., Toulouse. Robert-Passini M.-F., 1981. - Deux nouveaux pins pignons du Mexique, Bull. Mus. Hist. Natn., Paris, 4, 3, sect.B, Adansonia ,1,61-73. Zavarin & Snajberk, 1985. - Monoterpenoid and morphological differenciation within Piaus cembroides Biochem. Syst. and Ecol. 13, 2. 89-104. 5 THE ENDEMIC PINYON OF LOWER CALIFORNIA . P/NUS LAGUNAE M.-F. PASSINI Marie-Francoise Passini Université Pierre et Marie Curie, Laboratoire de Botanique tropicale 12 rue Cuvier, 75005 Paris Mexico has been an important center of Pious diversification, in particular of pines belonging to the sub-section cembroides which counts, now, 12 species My study of ecology and distribution of pines from the cembroides group(Passini, 1981) led me to describe, the pine in Sierra de la Laguna, Lower California as a variety of Piaus cembroides Zucc. (Robert-Passini 1981). In 1983, Bailey decided to give it subspecific rank. The morphological observations made by Pinel (1985) and the results obtained by Zavarin and Snajberk (pers. comm.) justify raising the rang to species PINUS LAGUNAE (M -F Robert-Passini, DK Bailey) MF Passini, coms. 2ov Pinus cembroides subsp. /aguaae (Robert-Passini) D.K Bailey, Phytologia 54,2, 89-99, 1983 Pinus cembroides var. Jagunae M.-F. Robert-Passini, Adansonia ser. 4, 3, sec. B, n° 1, 64-66, 1981 HOLOTYPE : -P, Sierra La Laguna, Delegacién Todos Santos, Baja California Sur, 23°34 N, 109°55 W, cristalline rock, 1650 m., 15.02.1978, M.-F Robert 10021 (HOLO- : P; ISO- : MPU, TLJ, ENCB, INIF) We have some several additional caracters to complete the description of the variety, using samples collected in february 1978 and july 1985. Araus /egunaehas an upright trunk, generally 12-15 meters high but can grow up to 21 meters. In open surroundings the habit is pyramid shaped whereas in a closed environnement the crown is sparse. The bark of mature trees is fissured and exhibits thick regular plates. The grey branchlets bear 3 needle fascicles (sometimes 2, seldom 4), 4-9 cm long (average length is 6,9 cm) soft to the touch and grey green in colour). Dorsal and ventral surfaces have stomacal! lines with more on the ventral surface, 4-8 lines, than on the dorsal surface, 1-3 lines The sub-globular cones are pedunculate, their average length is 3,9 cm and they grow singly or in twos. The peduncle can be 0,2 to 1.2 mm long and comes away with the cone The apterous seeds are 10 to 16 mm long, 6 to 10 mm wide with a 0,2 to0.9 mm thick shell. The endosperm is pink coloured. The following morphological characters distinguish Pious /egunae from Pinus cembroides si. : longer, more slender needles, longer cone peduncle, higher number of cotyledons (12,62), doubly quick-growing plantlets and saplings. But the biochemical characteristics revealed by Zavarin and Snajberk (1985) also bear this difference. Piaus cembrosdes wood is high: in «- pinene . 875% (min. 64, max. 9,7 %), low in sabinene : 3.4% (05-102 %) and in terpinolene : 2,1% (0,7-10,2 %). On the contrary Piaus /agunae is low in « -pinene : 135% (min 10,6 max. 16,1%), high in sabinene : 31,7% (14,5-45,7%) and in terpinolene : 27.2% (19,6-42,1%). 357, 338 POH, Yo 10) Ove eres Vol... 63, Nom > The monoterpene biosynthesis chains of these two taxa are quite distinct The difference between Praus /Jagunae and Pious cembroides is greater than that between Piaus remota and Pinus cembroides(SnajberK and Zavarin, 1986) The monoterpene constituents of Piaus /agunae are more akin to those of Praus discolor than to Piaus cembroides . Like Pinus lagunae , Pinus discolor synthesizes sabinene and terpinolene as well as a fair amount of pcymene (12,4% average). The latter only present in very small quantities in Piaus /aguaaewood (1,7% In addition to these characteristics pointed out by Passini, Bailey, Zavarin and Snajberk, the existence, in Sierra de 1a Laguna, of two varieties (Pinel, 1985) - one with a thin shell, 0,25-0,5 mm, the other with a thick shell (06-1 mm), has led me to raise Piaus cembroides subsp. /egunae (M.-F Robert-Passini) D.K Bailey to specific status. Differentiating a taxon distinct from Araus cembroides in Sierra de la Laguna was facilitated by the long term isolation which the mountains in the far southern tip of the Lower California peninsula underwent throughout the Tertiary period. This geografical isloation was accentuated in the Miocen by the La Paz-Todos Santos north-south fault This endemic species of Lower California, which adapts to chalkly parent rock soils, offers many advantages for retimbering dry, eroded areas, since growth rate is rapid (Passini, 1981). On these grounds, genetic studies will have to be pursued. ACKNOWLEDGEMENTS My grateful thanks go to Dr. Zavarin for having allowed me to use the results of analyses performed in this laboratory and for his helpful suggestions. REFERENCES Bailey D.K., 1983. - A new allopatric segregate from and a new combination in Pinus cembroides Zucc. at its southern limits. Phytologia 54, 89-99 Passini M.-F., 1981. - Les foréts de yous cembroides au Mexique, étude phytogéographigue et écologique. Ed. Recherches sur les Civilisations, 374 p Pinel N., 1985. - La formation a Piaus cembroides var. /agunae dans la Sierra de la Laguna, Basse Californie, Mexique. Rapport stage de DEA. Toulouse, 58 p Robert-Passini M.-F., 1981. - Deux nouveaux pins pignons du Mexique, Bull. Mus. Hist. natn, Paris, 4 3, sect. B, Adansonia 1 61-73. Snajberk K. & E. Zavarin, 1986. - Monoterpenoid differenciation in Relation to the morphology of Pinus remota. Biochem. Syst. and Ecol. 14 2, 155-163. Zavarin E. & K. Snajberk, 1985. - Monoterpenoid and morphological differeciation within Aiaus cembrosdes. Biochem Syst. and Ecol, 13 2, 89-104. DIAGNOSES OF DELISSEA SPECIES (LOBELIACEAE) FROM KAUAI HAWAIIAN PLANT STUDIES 145 Harold St. John Bishop Mus2um, Box 19000A, Honolulu Hawaii 96817, USA. It was known that the species of Delissea (or Cyanea) were numerous on the island of Kauai. Now, due to the in- tensive exploration and the collections made by Charles Christensen, it is rev@ajed that they are multidudinous. The types, unless otherwise indicated, are in the Bishop Museum, Honolulu. Delissea alba sp. nov. Frutex ramosus, ramulis glabris, petiolis 2.7-4 cm longis glabris, laminis 42.5-50.5 XK 9.6-12 cm oblanceo- latis subacuminatis basi cuneata et decurrenti infra in nervis scabre puberulis, racemis 7-9 cm longis, pedunc- ulo 3-4 cm longo puberulo, bracteis 5-6 mm longis lanceo- latis, pedicellis 9-12 mm longis, lobis calycis 5-6 mm longis lanceolatis, corollis 40 mm longis albis purpureo- lineatis puberulis, tubo filamentaru 31 mm longo glabro, antheris superis 9.5 mm longis glabris. Typus: Kauai I., Lumahai, C. Christensen 210, D. albilineata sp. nov. Frutex est, caule simpilci, petiolis 1.8-2 cm longis puberulis, laminis 24-34 3.5-5.4 cm oblanceolatis sub- acuminatis basi cuneata et decurrentiinfra in nervis pub- erulis, racemis 8 cm longis 10-floriferis, pedunculo 18 mm longo, bracteis 4-5 mm longis lanceolatis puberulis, pedicellis 13-15 mm longis puberulis, lobis calycis 6 mm longis lanceo-deltoideis puberulis, corollis 39 mm longis purpureis albi-lineatis puberulis, tubo filamentarum 30 mm longo glabro, antheris superis 10 mm longis glabris. Typus: Kauai I., Kahili Mt., C. Christensen ‘300. D. brevipedicellata sp. nov. Frutex est, caule simplici, petiolis 9-10.5 cm long- is glabris, laminis 37-42 K 9.7-10.8 cm oblanceolatis acutis basi cuneata et decurrenti infra in nervis prin- cipalibus puberulis, racemis 5-7 cm longis puberulis, pedunculo 1.5-2 cm longo, bracteis 2 mm longis lanceo- latis, pedicellis 8-12 mm longis pubervlis, lobis cal- ycis 1.8-2.5 mm longis deltoideis puberulis, corollis 37 mm longis. Typus: Kauai I., Waipa Valley C. Chris- tensen 239. D. cataracta sp. nov. Frutex ramosus est, petiolis 3-3.5 cm longis, laminis 45-54 X13.5-16.7 cm. oblanceolatis basi cun- ata et decurrenti infra nervis puberulis nervulis pil- osulis, pedicellis 8-17 mm longis, lobis calycis 6-7 339 340 PH Wet 07L OG. yA Vol: 63, No2 5 mm longis lanceolatis, corollis 35 mm longis,anthers sup- eris 10 mm longis. Typus: Kauai I., Limahuli Valley, S. Perlman & Wuchman 453. D. chartacea sp. nov. Frutex est, caule simplici, petiolis 5-6 cm longis, laminis 28.5-30 ¥7.4-8.3 cm oblanceolatis acutis basi cuneata et decurrent infra in nervis puberulis, racemis 5-9 cm longis 9-ll-floriferis puberulis, pedunculo 2-2.5 cm longo, bracteis 3-4 mm longis ellipticis ovatisve, pedicellis 6-8 mm longis puberulis, lobis calycis 2-3 mm longis lancei-ovatis puberulis, corollis 37 mm longis purpureis puberulis, antheris superis 11 mm longis glabris. Typus: Kauai I., Waipa Valley, C. Christensen 275. D. Christensenii sp. nov. Frutex est, caule simplici,petiolis 8-10 cm longis glabris, laminis 34-39 & 11-13 cm glabris oblanceolatis, pedicellis 12-14 mm longis puberulis, lobis calycis 3-3.5 mm longis deltoideis puberulis, corollis 52 mm longis purpureis puberulis, tubo filamentarum 28 mm longo glabro, antheris superis 11 mm longis glabris. Typus: Kauai I., Lumahai Valley, C. Christensen 201. D. coriacea A. Gray, var, deltoidea var. nov. Petiolis 6-9.5 cm longis glabris, laminis 20-30 X 5.2-9 cm ellipticis acutis basi cuneata infra midnervo in lateribus puberulis, racema 6-8 cm longa, pedunculo 2-3 cm longo, pedicellis 10-12 mm longis, lobis calycis 1-1.3 mm longis deltoideis, corollis 30 mm longis albis. Typus: Kauai I., Limahuli Valley, S. Perlman 1. D. coriaceae A. Gray, var. haupuensis var. nov. Laminis anguste oblanceolatis, corollis 20-23 mm longis. Typus: Kauai I., Laaukahi, C. Christensen 91. D. coriacea A. Gray, var. lumahaiensis var. nov. Petiolis 8-9 cm longis, laminis 4.5-30X 4.8-8.7 cm oblanceolatis acutis infra in midnervo in lateribus hirsutis, racemis 3-12 cm longis, pedunculo 2-9 cm longo, pedicellis 12-17 mm longis, lobis calycis 0.5-0.7 mm longis deltoideis, corollis 20-23 mm longis purpureis. Typus: Kauai I., Hanalei, C. N. Forbes 464.K. D. decumbens sp. nov. Frutex decumbens est, caule glabro, petiolis 3 cm longis puberulis, laminis 31-34 K 8.3-9.2 cm oblanceo- latis acuminatis basi cuneata infra in nervis puberulis, racemis 5-6.5 cm longis 7-floriferis puberulis, pedunculo 2-2.5 cm longo, bracteis 3 mm longis lanceolatis, ped- icellis 7-12 mm longis, lobis calycis 3.5-5 mm longis deltoideis, corollis 35 mm longis purpureis. Typus: Kauai I., Wainiha Valley, C. Christensen 271. D. deltoidea sp. nov. Frutey ramosus est, petiolis 20-27 mm longis hirsutulis, laminis 22.5-30.2 K 5.4-6.7 cm. oblanceo= _ 1987 St. John, Hawaiian Delissea 341 latis subacuminatis basi cuneata infra in nervis princi- palibus puberulis, racemis 6 cm longis 3-4-floriferis, pedunculo 8-14 mm longdpuberulo, bracteis 4-5 mm longis lanceolatis puberulis, pedicellis 12-17 mm longis puber- ulis, lobis calycis 2 mm longis deltoideis puberulis, corollis 37-38 mm longis violaceis puberulis, tubo fil- amentarum 24 mm longo, antheris superis 8.7-9 mm longis glabris. Typus: Kauai I., Wainiha Valley, C. Christensen 305 “Dp. denticulata sp. nov. Frutex ramosus est, ramulis glabris, petiolis 3 cm longis puberulis, laminis 45-49 }’ 12.5-15 cm oblanceola- tis, subacuminatis basi cuneata infra in nervis puber- ulis, racemis 8-9 cm longis 5-1ll-floriferis puberulis, pedunculis 15-25 mm longis, bracteis 12-15 mm longis ligulatis acutis, pedicellis 15-20 mm longis, lobis calycis 11-12 mm longis ligulatis acutis puberulis, corollis 35 mm longis violaceis puberulis, Typus: Kauai I., C. Christensen 237. D. divergens sp. nov. Frutex est, caule glabrdunico, foliis glabris, petiolis 6.5-8 cm longis, laminis 33-38.5 K 10-11.3 cm oblanceolatis acutis basi cuneata et decurrenti, rac- emis 5-7 cm longis 5-13-floriferis puberulis, pedunc- ulo 1.5-4 cm longo, bracteis 5 mm longis lanceolatis, pedicellis 7-10 mm longis, lobis calycis 4-5 mm longis ovati-deltoideis puberulis, corollis 38 mm longis lavandulis puberulis, tubo filamentarum 30 mm longo glabro, antheris superis 8.5 mm longis glabris. Typus: Kauai I., Waipa Valley, C. Christensen 234. D. duploserrata sp. nov. Frutex ramosus est, caule glabro, petiolis 2-5 cm longis puberulis, laminis 35-38 10-12.3 cm oblanceo- latis subacuminatis basi cuneata infra in nervis puber- ulis, racemis 10-14 cm longis{l1-27-floriferis puberulis, pedunculis 3-6 cm longis, bratteis 5-6 mm longis oblongis acutis, pedicellis 9-11 mm longis puberulis, lobis calycis 6-7 mm longis lanceolatis ciliolatis, corollis 40 mm longis puberulis albis et roseo-lineatis, tubo filamentarum 26 mm longo glabro, antheris superis 7 mm longis glabris. Typus: Kauai I., Wainiha Valley, C. Christensen 310. D. eleeleensis sp. nov. Frutex est, ramis glabris, foliis glabris, petiolis 5-8.5 cm longis, laminis 39-40.5 X 11.4-11.9 cm oblanco- latis acutis basi cuneata et decurrenti, racemis 10-14 cm longis puberulis, bracteis 2.5-3 cm longis lanceolatis, pedicellis 10-17 mm longis, lobis calycis 4-5 mm longis ovatis acutis puberulis, corollis 40 mm longis purpureis, tubo filamentarum 34 mm longo, antheris superis 11 mm longis glabris. Typus: Kauai I., Wainiha Valley, 342 PHY LO L Orc TWA Vol. 635) Now > C. Christensen 261. D. excurrens sp. nov. Frutex est, caule simplici glabro, petiolis 8.5-10.5 cm longis glabris, laminis 30-38.5 Xx 7.8-10.8 cm glabris elliptici-oblanceolatis subacuminatis basi cuneata, racem- is 7-9 cm longis 5-9-floriferis puberulis, pedunculis 2.5- 3.5 cm longis, bracteis 2 mm longis lanceolatis, pedicellis 15-18 mm longis, lobis calycis 2.5-3 mm longis puber- ulis, corollis 37 mm longis albis sed purpure-lineatis. Typus: Kauai I., Wainiha Valley, C. Christensen 311. D. Fauriei (H. Lévl.) comb. nov. Cyanea Fauriei H. Lévl., Fedde Repert spec. Nov. Regni Veg. 107) 156, 29177 , C. coriacea (A. Gray) Hillebr., var, Fauriei (H. Levl.) E. Wimm., Engler, Pflanzenr. 276b (106): 72, 1957. D. 2rutEvcosa sp.) nov. Frutex est, caulj.. glabro decumbenti, foliis glabris, petiolis 8-8.5 cm longis, laminis 52 X 14.5-15.3 cm oblan- ceolatis subacuminatis basi cuneata et decurrenti, rac- emis 9-10 cm longis 11-13-floriferis puberulis, pedunculo 3.5-4 cm longo, bracteis 4-5 mm longis lancei-ligulatis, pedicellis 7-10 mm longis, lobis calycis 3.5-4 mm longis deltoideis puberulis, corollis 45-47 mm longis puberulis, tubo filamentarum 31 mm longo glabro, antheris superis 8 mm longis glabris, Typus: Kauai I., Lumahai Valley, C. Christensen 247. D. glabrifolia sp. nov. Frutex est, caule simplici, petiolis 5-7 cm longis glabris, laminis 61-65.5 X 14-14.7 cm glabris oblanceolatis acutis basi cuneata, raemeis 8-10 cm longis puberulis, pedunculo 3-4 cm longo, bracteis 9-11 mm longis ligul- atis acutis, pedicellis 20-23 mm longis, lobis calycis 7-8 mm longis lineari-lanceolatis puberulis, corollis 47- 48 mm longis purpureis, tubo filamentarum 32 mm longo, glabro, antheris superis 12 mm longis glabris. Typus: Kauai I., Waioli-Waipa, C. Christensen 192. D. habenata sp. nov. Frutex ramosus glaber est, petiolis 3.5-7 cm longis, laminis 14-213 2.1-3 cm ligulatis ambitu acutis, rac- emis 3-6 cm longis 5-13-floriferis, pedunculo 1-3 cm longo, pedicellis 6-10 mm longis, lobis calycis 0.2 mm longis deltoideis, corollis 24-25 mm longis albis, tubo fil- amentarum 20 mm longo, antheris superis 4 mm longis. Typus: Kavai I., Limahuli Valley, S. Perlman 224. D. humilis (H. Wawra) comb. nov. Cyanea humilis H. Wawra, Flora 31: 47, 1873, (p. 32 in reprint). D. hypoleuca sp. nov. Frutex est, petiolois 3-6.5 cm longis glabris, lam- inis 23.5-29€ 6-6.8 cm glabris oblong-oblanceolatis- - acutis basi cuneata, racemeis 6-15 cm longis 7-13- 1987 St. John, Hawaiian Delissea 343 floriferis puberulis, pedunculo 3-8 cm longo, bracteis 1 cm longis lanceolatis, lobis calycis 2-2.5 mm longis del- toideis, corollis 35 mm longis puberulis, anthers super- is 8-9 mm longis glabris. Typus: Kauai I., Wahiawa Valley, S. Perlman 482. D. iliahiatilis sp. nov. Frutex est, caule simplici, foliis glabris, petiolis 4-4.2 cm longis, laminis 37-41 X 12.5-14.8 cm oblanceola- tis acuminatis basi cuneata, racemis 7-9 cm longis 11-19- floriferis puberulis, pedunculo 3-3.8 cm longo, bracteis 4-5 mm longis lanceolatis, pedicellis 10-16 mm longis, lobis calycis 3-4 mm longis deltoideis puberulis, corollis 35 mm longis purpureis albi-lineatis. Typus: Kauai I., Lumahai Valley, C. Christensen 249. D. impedicellata sp. nov. Frutex ramosus est, foliis glabris, petiolis 3.5-6.5 cm longis, laminis 15-18 X 3.8=4.8 cm oblanceolatis acutis basi cuneata et decurrenti, spicis 2.5-3 cm longis 10- 20-floriferis hirsutulis, pedunculo 3-6 mm longo, bract- eis 7-13 mm longis linearibus, floribus sessilibus, lobis calycis 2-2.5 mm longis deltoideis, corollis 28-32 mm longis puberulis. Typus: Kauai I., Waiai River, F. R. Warshauer 3,382. D. inermis sp. nov. Frutex ramosus glaber est, petiolis 3.5-12 cm longis, laminis 6.5-12.5 & 1.7-4 cm fusiformibus, racemsis 5-8 cm longis 3-5-floriferis, pedunculo 1.5-4.5 cm longo, ped- icellis 10-18 mm longis, lobis calycis 0.3 mm longis del- toideis, gemmis 13 mm longis albis. Typus: Kauai I., Wahiawa Stream, S. Perlman 486. D. kahiliensis sp nov. Frutex ramosus est, foliis glabris, petiolis 4.5-6 cm longis, laminis 11.8-12.4 cm anguste ellipticis ambitu acutis, racemis 5 cm longis glabris, pedunculo 2 cm longo, pedicellis 10-12 mm longis, lobis calycis 0.5 mm longis deltoideis, corollis 20 mm longis albis, tubo filamentar- um glabro, antheris superis 5.5 mm longis connectivis puberulis. Typus: Kauai I., Kahili, ‘C. Corn. D. keaensis sp. nov. Frutex ramosus est, petiolis 3.5-4 cm longis puber- ulis, laminis 37-43 & 10.7-11.6 cm oblanceolatis subacum- inatis basi cuneata et decurrenti infra puberulis, racemis 8-9 cm longis 9-15-floriferis puberulis, pedunculo 5-6.2 cm longo, bracteos 4-5 mm longis lanceolatis, pedicellis 8-10 mm longis, lobis calycis 5-6 mm longis deltoideis, corollis in gemma 29 mm longisalbis puberulis. Typus: Kauai I., Wainiha Valley, C. Christensen 306. D. kealawelaensis sp. nov. Frutex est, calule simplici glabro, foliis glabris, petiolis 4.5-6 cm longis, laminis 45-47 X 13-14.5 cm 344 PH xertO) Ev OnG elas Vol. 63, Nov 5 oblanceolatis acutis basi cuneata et decurrenti, racemis 4.5-9.5 cm longis 9-13-floriferis puberulis, pedunculo 1.2-3 cm longo, pedicellis 6-11 mm longis, lobis calycis 1.3-3 mm longis deltoideis, corollis 37 mm longis purpur- eis sed albi-lineatis, Typus Kauai I., Wainiha Valley, C. Christensen 268. D. kolekoleensis sp. nov. Frutex glaber est, petiolis 5-10 cm longis, laminis 19-26 4.4-5.5 cm ellipticis subacutis basi cuneata, racemis 19 cm longis, pedunculo.14 cm longo, pedicellis 15-25 mm longis, lobis calycis 1.5-2 mm longis deltoideis, corollis 50 mm longis rosaceis puberulis. Typus: Kauai I., Wahiawa Valley, S. Perlman 498. D. latibasilaris sp. nov. Frutex ramosus est, ramulis glabris, foliis sessili- bus vel subsessilibudet cum petiolis ad 2 cm longis alat- is, laminis 36-41 & 9.7-12 cm oblanceolatis subacumin- atis basi cuneata et decurrenti infra puberulis, racemis 8-9 cm longis 11-15-floriferis puberulis, pedunculo 1-2 cm longo, bracteis 5-6 mm longis ligulatis acutis sed basi hemisphaerica, pedicellis 12-13 mm longis, lobis calycis 6-7 mm longis lanceolatis glabris, corollis 35 mm longis puberulis albis sed magenta-lineatis. Typus: Kauai I., Lumahai Valley, C. Christensen 228. D. leiophylla sp. nov. Frutex est, caule simplici glabro, foliis glabris, petiolis 6-6.5 cm longis, laminis 35.5-43 > 13-17 cm oblanceolatis acutis basi cuneata, racemis 5-7 cm longis 9-17-floriferis puberulis, pedunculo 1.5-2.5 cm longo, bracteis 3 mm longis lanceolatis, pedicellis 6-11 mm longis, lobis calycis 1-2 mm longis lanceolatis puber- ulis, corollis 35 mm longis puberulis. Typus: Kauai I., Waioli, C. Christensen 278. D. ligulata sp. nov. Frutex est, caule simplici, foliis glabris, pet- iolis 3.5-4.5 cm longis, laminis 34.5-40 & 8.8-11.6 cm oblanceolatis subacuminatis basi cuneata, racemis 7-8 cm longis 7-10-floriferis puberulis, pedunculo: 2-4 cm longo, bracteis 4-6 mm longis lanceolatis, pedicel- lis 10-13 mm longis, lobis calycis 3-4 mm longis del- toideis, corollis 42-43 mm longis maroonis. Typus: Kauai I., Lumahai Valley, C¢. Christensen 205. D. limahuliensis sp, nov. Frutex est, caule simplici, foliis sessilibus 45-47 & 18-18.5 cm oblanceolatis subacuminatis basi cuneata infra puberulis, racemis 11 cm longis 17-25- floriferis, pedunculo 4-5 cm longo, bracteis 7 mm longis ligulatis subacutis, pedicellis 12-14 mm longis puberulis, lobis calycis 8.5-10 mm longis lanceolatis marginibus. puberulis, corollis 35 mm longis purpureis, tubo filamentarum 15 mm longo glabro, antheris super- 1987 St. John, Hawaiian Delissea 345 is 8 mm longis glabris. Typus: Kauai I., Limahuli Valley, S. Perlman 4. D. longicalyx sp. nov. Frutex est, caule simplici, petiolis 3-3.5 cm longis pilosulis, laminis 32-48 X 8.5-11 cm, oblanceolatis acutis basi cuneata et decurrenti infra puberulis, racemis 7-1l cm longis 7=15-floriferis puberulis, pedunculo 2-5 cm longo, bracteis 8 mm longis ligulatis acutis, pedicellis 15-18 mm longis puberulis, lobis calycis 9-18 mm longis lanceolatis puberulis, corollis 35 mm longis puberulis, tubo filamentarum 25 mm longo, antheris superis 9 mm longis glabris. Typus: Kauai I., Waioli-Waipa, C. Christensen 191. D. longiantherae sp. nov. Frutex est, caule simplici, foliis glabris, petiolis 7-7.5 cm longis, laminis 47-55 X% 14-16 cm oblanceolatis acutis basi cuneata, racemis 7-10 cm longis puberulis, pedunculo 1.5-2 cm longo, bracteis 3 mm longis lanceolatis, pedicellis 10-17 mm longis, lobis calycis 2 mm longis deltoideis, corollis 45 mm longis purpureis puberulis, tubo filamentarum 33 mm longo glabro, antheris superis 10 mm longis glabris. Typus: Kauai I., Waioli-Waipa, C. Christensen 215. D. lumahaiensis sp. nov. Frutex est, caule simplici, foliis subsessilibus, petiolis 12 mm longis glabris, laminis 35-44 & 9.2-13 cm oblanceolatis acutis basi cuneata et decurrenti infra puberulis, racemis puberulis, pedunculo 2-3 cm longo, rhachidibus 1.5-2 cm longis, bracteis 3-4 mm longis lan- ceolatis, pedicellis 8-10 mm longis, lobis calycis 3-5 mm longis deltoideis glabris, corollis 36 mm longis purpureis puberulis. Typus: Kauai I., Lumahai Valley, C. Christensen 204. D. Lydgatei nom. nov. Cyanea undulata Forbes, Occas. Papers B. P. Bishop Mus. S(1) 3 22<13,, wrth Fig.,, 19:12, non..Gaud.. (1e26y)., D. multiramosa sp. nov. Frutex ramosus est, petiolis 2-2.5 cm longis pil- osulis, laminis 26-31 \¥ 8-10.5 cm elliptici-oblanceo- latis subacuminatis basi cuneata infra in nervis pilosuls, racemis 5 cm longis 11l-floriferis puberulis, pedunculo 2 cm longo, bracteis 5 mm longis lanceolatis, pedicellis 6-7 mm longis, lobis calycis 5-6 mm longis ovatis acutis vel lanceolatis puberulis, corollis 32-35 mm longis pub- erulis. Typus: Kauai I., Waioli, C. Christensen 279. D. napaliensis sp. nov. Frutex est, caule simplici, petiolis 6-7 cm longis glabris, laminis 43.5-45.5 K 10.8-11 cm oblanceolatis subacuminatis basi cuneata infra puberulis, racemis 6-7 cm longis puberulis, pedunculo 1.5-3 cm longo, bracteis 5-6 mm longis lanceolatis, pedicellis 8-12 mm longis, 346 PHY T,0.L.0,CG 1 A Vol, 63,. Ho. obis calycis 4-4.5 mm longis ovatis acutis puberulis, cor- ollis 45 mm longis puberulis. Typus: Kauai I., Hanakapiai, C. Christensen 188. D. nigra sp. nov. Arboriformis est, caule simplici, foliis glabris, petiolis 6.5-8 cm longis, laminis 38-41 X¥ 4.9- 5.6 cm. oblongi-linearibus acutis basi decurrenti, racemis 27-28 cm longis 16-28-floriferis pendentibus glabris, pedunculo 5-10 cm longo, bracteis 1-3 mm longis lanceo- latis, pedicellis 10-13 mm longis, lobis calycis 1 mm longissubulatis vel lanceolatis, corollis 27-32 mm longis purpurei-nigris, tubo filamentarum 24-28 mm longo, antheris superis 9-10 mm longis glabris. Typus: Kauai I., Hanakapiai, C. Christensen 2. D. paliensis sp. nov. Frutex est, caule puberulo, petiolis 4-8 cm longis puberulis, laminis 20-34 X 5-12 cm glabris oblanceolatis acutis basi cuneata, racemis 5-7 cm longis 5-9-floriferis puberulis, bracteis 3-15 mm longis ligulatis acutis, pedicellis 9-11 mm longis, lobis calycis 1.8-2.2 mm longis deltoideis puberulis, corollis 43 mm longis, tubo filamen- tarum 30 mm longo, antheris superis 8 mm longis glabris. Typus: Kauai I., Limahuli Valley, S. Perlman 221. D. parva sp. nov. Frutex est, caule simplici, petiolis 5-5.5 cm longis glabris, laminis 60-62.5 X 11.7-12.7 cm oblanceolatis acutis basi cuneata infra in nervis puberulis, racemis 9-11 cm longis puberulis, pedunculo 2-2.5 cm longo, bracteis 12-13 mm longis ligulatis acutis, pedicellis 20- 30 mm longis, lobis calycis 7-9 mm longis lanceolatis puberulis, corollis 45 mm longis puberulis, antheris sup- eris 10 mm longis glabris. Typus: Kauai I., Waioli-Waipa, Cc. Christensen 213. ; D. Perlmannii sp. nov. Frutex est, caule simplici, petiolis 6-10 cm longis glabris, laminis 19-27 2 6.5-9 cm ellipticis basi cuneata et decurrenti infra in lateribus midnervi pilosulis, rac- emis 6-9 cm longis glabris, pedunculo 2-3 cm longo, ped- icellis 10-14 mm longis, lobis calycis 0.7-0.8 mm longis deltoideis, corollis 32 mm longis albis, tubo filamentarum 25 mm longo, antheris superis 6.5 mm longis glabris. Typus: Kauai I., Lumahai Valley, S. Perlman l. DD. plurifilora sp. nov- Frutex ramosus est, ramis glabris, petiolis 0-20 mm longis, foliis subsessilibus, laminis 30-40 KX 8-10.7 cm oblanceolatis subacuminatis basi cuneata et decurrenti infra puberulis, racemis 10-12 cm longis 15-30-floriferis puberulis, bracteis 6-8 mm longis ligulatis acutis, ped- unculo 15-25 mm longo, pedicellis 10-15 mm longis, lobis’° calycis 6-8 mm longis deltoideis, corollis 35 mm longis 1987 St. John, Hawaiian Delissea 347 puberulis albis sed vinaceo-lineatis, tubo filamentarum 24 mm longo glabro, antheris superis 9 mm longis glabris. Typus: Kauai I., Lumahai, ¢€. Christensen 223. D. purpurea sp. nov. Arbor 6.6 m alta est, caule simplici puberulo, fol- iis glabris, petiolis 3-4 cm longis, laminis 24-29 ® 10.5- 13 cm oblanceolatis acutis basi cuneata et decurrenti, racemjs 3-5 cm longis 5-1ll-floriferis puberulis, ped- unculo 6-15 mm longo, bracteis 3-4 mm longis lanceolatis, pedicellis 6-8 mm longis, lobis calycis 2-3 mm longis deltoideis puberulis, corollis 40 mm longis purpureis puberulis, tubo filamentarum 20-23 mm longo glabro, an- theris superis 8 mm longis glabris, Typus: Kauai I., Waioli-Waipa, C. Christensen 217. D. ramosSa sp. nov. Frutex ramosus est, ramulis glabris, foliis junior- ibus puberulis, petiolis 2-4 cm longis, laminis 28-40.5 & 7-10 cm oblanceolatis acutis basi cuneata et decurrenti, racemis 8-9 mm longis 11l-floriferis puber- ulis, pedunculo 3.5 cm longo, bracteis 7-8 mm longis ligulatis acutis, pedicellis 10-12 mm longis, lobis calycis 7-8 mm longis lanceolatis puberulis, corollis 45 mr longis laévancdulis puberulis. Typus: Kauai l., Waica, C. Christensen 24]. D. Robinsonii sp. nov. Frutex ramosus est, ramis glabris, petiolis5-6 cm longis puberulis, laminis 49.5-56.5 ®14.2-15 cm cboanceolatis subacuminatis basi cuneata et decurrenti infra in nervis puberulis, racemis 6-S cm longis 9-19-floriferis puberulis, pedunculo 10-17 mm longo, lobis calycis 5-6 mm longis deltoideis puberulis, corollis 44-45 mm longis purpureis, tubo filamentarum 35 mm longo glabro, anthers superis 10 mm longis glabris. Typus: Kauai I., Wainiha, C. Christensen 269. Di. sScopuli sp. nov. Frutex est, caule glabro, petiolis 4-7 cm longis puberulis, laminis 32-59 K 11.5-19 cm oblanceoatis subacuminatisbasi cuneata infra in nervis scabriter puberulis, racemis 12 cm longis 30-floriferis puberulis, pedunculo 15 mm longo, bracteis 5-7 mm longis ligulatis acutis, pedicellis 15 mm longis, lobis calycis 3-4.5 mm longis ovatis subacutis puberulis, corollis 47 mm longis purpureis puberulis, tubo filamentarum 30 mm longo, antheris sugtis 9 mm longis glabris. Typus: Kauai I., Limahuli, S. Perlman 222. D. serratifolia (Rock) comb. nov. Cyanea coriacea (A. Gray) Hillebr., var. serratifoli, Rock, Occas Papers B. P. Bishop Mus. 2245), 65 1857. 348 Pan ey 1. OFL, ORG Vol. 63, Noe D. simples sp. nov. Frutex est, caule simplici, petiolis 8-11.5 cm longis puberulis, laminis 38-53 X 10.5-16 cm oblanceolatis acutis basi cuneata et decurrenti infra hirsutulis, racemis 6-10 cm longis 7-1l-floriferis puberulis, pedunculo 1.5- 4.5 cm longo, bracteis 3-4 mm longis lanceolatis, pedicellis 10-23 mm longis, lobis calycis 1.8-2.5 mm longis deltoid- eis, corollis 35-40 mm longis purpureis puberulis, tubo filamentarum 25-28 mm longo glabro, antheris superis 8.5- 9 mm longis glabris. Typus: Kauai I., Ke'e, C. Christensen 3h rapt Simplex St. John, forma maculata forma nova. A species differt in laminis magenta-maculatis. Typus: Kauai I., Ke'e, C. Christensen 13a. D. subacuminata sp. nov. Frutex est, caule simplici, foliis glabris, petiolis 6-9 cm longis, laminis 30-33.5 10-11 cm oblanceolatis subacuminatis, racemis 7-9 cm longis 7-9-floriferis puberulis, pedunculo 2.5-3 cm longo, bracteis 3 mm longis lanceolatis, pedicellis 8-12 mm longis, lobis calycis 2.5- 4 mm longis lanceolatis, corollis 43 mm longis purpur- eis puberulis, antheris superis 9 mm longis glabris. Typus: Kauai I., Waioli-Waipa, C. Christensen 219. D. subintegra sp. nov. Frutex est, caule glabro, petiolis 6 cm longis glabris, laminis 33-34 &K 9.3-10.5 cm oblanceolatis acutis basi cuneata et decurrenti infra puberulis, racemis 5.5-6 cm longis 3-5-floriferis puberulis, pedun- culo 1.5-2cm longo, bracteis 3-4 mm longis lanceolatis, pedicellis 7-8 mm longis, lobis calycis 2.5-3 mm longis deltoideis puberulis, coroilis 30-35 mm longis albis, tubo filamentarum 28 mm longo glabro, antheris superis 8 mm longis glabris. Typus: Kauai I., Waioli C. & H. Christensen 276. b. subsessilis sp. nov. Frutex est, caule simplici, foliis subsessilibus, laminis 29-33 X 7.6-8.7 cm oblanceolatis subacuminatis basi decurrenti infra puberulis, racemis 5-6 cm longis 3-7-floriferis puberulis, pedunculo 1-2 cm longo, bracteis 8-9 mm longis ligulatis acutis, pedicellis 10-12 mm longis, lobis calycis 7-8.5 mm longis lanceolatis puber- ulis, corollis 33 mm longis puberulis albis sed purpureo-lineatis. Typus: Kauai I., Waioli-Waipa, C. Christensen 193. D. umbroSa sp. nov. Frutex ramosus decumbens est, foliis glabris, pet- iolis 8-9 cm longis, laminis 33-34 K 10.5-11.1 cm ob- lanceolatis acuminatis basi rotundata, racemis 5-7- floriferis puberulis, pedunculo 27-30 mm longo, bract- eis 2 mm longis deltoideis, pedicellis 10-13 mm longis, lobis calycis 2.3-2.8 mm longis deltoideis puberulis, - 1987 St. John, Hawaiian Delissea 349 corollis in gemma 22 mm longis puberulis. Typus: Kauai I., Wainiha Valley, C. Christensen 291. D. virgata sp. nov. Frutex ramosus est, foliis glabris, petiolis 3.5-5 cm longis, laminis 29.5-41.5 X 9.7 -—4.6 cm oblanceolatis subacuminatigdbasi cuneata et decurrenti, racemis 8-10 cm longis 3-7-floriferis puberulis, pedunculo 3-5 cm longo, bracteis 4 mm longis lanceolatis, pediceliis 10-15 mm longis, lobis calycis 4-5 mm longis puberulis, corollis 47 mm longis puberulis purpureis, tubo filamentarum 34 mm longo glabro, antheris superis 12 mm longis glabris. Typus: Kauai I., Wainiha, C. Christensen 308. D. waioliensis sp. nov. Frutex est, caule simplici, foliis glabris, petio- lis 6 cm longis, laminis 47.5 X 13 cm oblanceolatis acutis basi cuneata, racemis 5-9-fioriferispuberulis, bracteis 3-5 mm longis lanceolatis, pedicellis 8-12 mm longis, lobis calycis 3-4 mm longis deltoideis puberulis; coroliis 45 mm longis purpureispuberulis, tubo filamentar- um 33 mm longoglabro, antheris ‘superis 10 mm longis glab- ris. Typus: Kauai I., Waipa Valley, C. Christensen 68. D. waipaensis sp. nov. Frutex est, caule simplici, petiolis 5 cm longis glabris, laminis 43.5 X 11.4 cm oblanceolatis acutis basi cuneata infra in nervis puberulis, racemis 6 cm longis puberulis, pedicellis 6 mm longis, lobis calycis 5 mm longis deltoideis puberulis, corollis 30 mm longis purpureis puberulis, tubo filamentarum 27 mm longo glabro, antheris superis 8 mm longis glabris. Typus: Kauai I., Waipa Valley C. Christensen 67. DIAGNOSES OF CLERMONTIA SPECIES (LOBELIACEAE) Hawaiian Plar+t Studies 146 Harold St. John Bishop Museum, Box 19000A, Honolulu, Hawaii 96817, USA. The types are in the Bishop Museum, unless elsewhere located. Clermontia albimontis sp. nov. (sect. Clermontia), Fig. l. Diagnosis Holotypi: Ramulae puberulae sunt, petiolis puberulis, laminis 12-20 % 5.3-6.3 cm oblancei-elliptici subacutis serrulatis infra albe puberulis, cymis puberulis 2-floriferis, pedunculo 20 mm longo, pedicello 20 mm longo, gemmis grandis 4 KX 1.5 cm pallide viridibus puber- ulis, anthers superis 12 mm longis. Typus: Hawaii I., Laupahoehoe, Ha'akoa Stream, F. R. Warshauer et al. 1,289. C. bicolorata sp. nov. (sect. Clermontioideae). Fig. 2. Frutex 3 m altus glaber est, petiolis 25-37 mm longis, laminis 10-16 X¥ 2.2-4.8 cm fusiformi-ellipticis, cymis -floriferis, pedunculo 12 mm longo, lobis calycis 12-16 2.5-3 mmviridibus ligulatis, corolla in alabastro Z cm longa bicolorata. Typus: Hawaii I., Kawaihae, Kehena Ditch, F .R. Warshauer & McEidowney 2,469. €.- epilosa’ sp: nov: (sect. Clermontia) . Fig iss Diagnosis Holotypi: Frutex glaber est, laminis 13-17.6 XK 2,6-3.3 cm coriaceis anguste oblanceolatis serrulatis, cymis 8-10 cm longis 2-floriferis, pedunculo 2 cm longo, pedicellis 8-11 mm longis, perianthio 4.5 cm longopbscur- e rubris, antheris superis 13 mm longis. Typus: Hawaii I., Kaukini Honokane, G. Gillett 1,917. C. glabra sp. nov. (sect. Clermontia), Fig. 4. Diagnosis Holotypi: Frutex gliaber est, petiolis 3.5- 4.5 cm longis, laminis 19~22.5 5-6 cm oblanceolatis, cymis 12 cm longis 2-floriferis, pedunculo 45 mm longo, pedicellis 15 mm longis, lobis calycis petaloideis ad corollam aequantibus, corollis 45 mm longis obscure rubris, tubo filamentaru,"glabro, anthers glabris luteis eis superis 11 mm longis iliis inferis 8 mm longis. Typus: Maui I., State Park, road to Hana, G. W. Gillett 1,686. Cc. gracilis sp. nov. (sect...Clermontia),.Figs 5% Diagnosis Holotypi: Frutex 2 m altus est, ramis resinosis hirsutis pilis cum apicibus rubris, petiolis 10-22 mm longis hirsutulis, laminis 4.5-1l1 K1.8-3.4 cm subcoriaceis oblanceolatiscrenatis supra remote minute puberulis et midnervo hirsutulo infra nervis hirsutulis, cymis 2-floriferis, pedunculo 9-11 mm longo minute puberulo, pediceliis 15-17 mm longis glabris, hypanthio cylindrico, lobis calycis petaloideis22-27 mm longis 350 1987 St. John, Hawaiian Clermontia S5il tubo 4 mm diametro rosaceis, corollis (immaturis) 15-20 mm. longis. Typus: Maui I., Kipahulu, P. K. Higashino Setal 29,278 C. hualalaiensis sp. nov. (Sect. Clermontia), Fig. 6. Diagnosis Holotypi: Laminis 10-12% 2-3 cm anguste oblongi-ellipticis acutis crenulatis infra proxima mid- nervum pilosulis, cymis 8.5-10 cm longis 2-floriferis, pedunculo 25-32 mm longo, corollis 5 cm longis, antheris superis 14.5 mm longis. Typus: Hawaii i., Hualalai, G. W., Gillett. 1,704. C. kahuaensis sp. nov. (sect. Clermonioideae), Fig. 7. Frutex 5 m altus est, petiolis 2.5-3 cm longis puberulis, laminis 11-14 Xk 3.2-3.8 cm fusiformi-ellip- ticissubacuminatis crenulatis infra nervis puberulis, cymis 2-floriferis, pedunculo 1 cm longo, pedicellis 8 mm longis, lobis calycis 4-4.5 mm longis anguste lanceo- latis, corollis in gemma 2.5 cm longis puberulis. Typus: Hawaii I., Kahua 2, Pu'a Iki, R. L. Stemmermann et al. 4,000. C. kakeana Meyen, var rosea (Hillebr.) comb. nov. C. macrocarpa Gaud., var. rosea Hillebr., Fl. Haw. is-;i241, L868, C. kipahuluensis sp. nov. (sect. Clermontia), Fig. 8. Diagnosis holotypi: Frutex est, ramulis puberulis, petiolis 15-28 mm longis sparse puberulis, laminis 5.1-8.5 * 2-3.8 cm coriaceis ellipticis late acuminatis basi cuneata supra glabris infra midnervo hirsutulo, cymis 5-7-floriferis, pedunculo 8-11 mm longo puberulo, pedicellis 10-17 mm longis, calycibus petaloideis, peri- anthio 6 cm longo hamate recurvatoglabro albo, antheris glabris eis inferis 10 mm longis in apice penicillatis cum setis 1.5 mm longis. Typus: Maui I., Kipahulu, F. R. Warshauer et al. 2,886. Clermontia mauiensis sp. nov. (sect. Clermontia), Fig. oe Diagnosis Holotypi: Frutex est, ramis glabris, pet- iolis 12-19 mm longis glabris, laminis 10.5-14 x 3.5-4.5 cm subcoriaceis elliptici-oblanceolatis acutis cuneatis serrulatis infra in nervis minute puberulis, cymis 2-floriferis, pedunculo 20 mm longo, pedicellis 12 mm longis, lobis calycis petaloideis simulantibus circa 4.5 cm longis, petalis 4.5 cm longis, antheris superis 10 mm longis. Typus: Maui I., Hana, Ke'anae, P, K. Higashino et al...) , 089. C. pluriflora sp. nov. (sect. Clermonioideae), Fig. 10. Diagnosis Holotypi: Petiolis 18 mm longis, lamin- is 22-25 xX 5.5-7.8 cm fusiformi-ellipticis infra glaucis et nervis hirsutulis, cymis 3-6-floriferis pilosulis, lobis calycis 3 mm longis deltoideis, cor- Ollis 4 cm longis, tubo filamentarum pilosulis, 352 PRHeY TOF ONG eA Vol. 63, Now5 antheris superis 10.5 mm longis glabris. Typus: Kauai I., Alakai, L. M. Cranwell & Skottsberg 2,983, C. spatulata sp. nov. (sect. Clermontia). Diagnosis Holotypi: Frutex 2 m altus cum 45-50 ramis glabris est, petiolis 1-2 cm longis glabris, laminis 3.5-8 X 1.2-2.2 cm glabris spatulatis obtusis in dimidio ap- icali marginibus crenatis infra subalbis, inflorescentia glabra cernua 2-florifera, pedunculo 3.5-5 cm longofili- formi, pedicellis 3-3.5 cm longis, lobis calycis petaloid- eis, perianthio 5.5 cm longo pallide viridi et purpur- eo-tincto, tubo filamentarum glabro, antheris inferis 9 mm longis aliter glabris sed in apice penicillatis cum setis 1.5mm longis. Typus: Maui I., Honokawai, F. R. Warshauer & McEldowney 3,033. C. subteralbulus sp. nov. (sect. Clermontia). Diagnosis Holotypi: Frutex glaber est, petiolis 15- 20 mm longis, laminis 7-10 >*<1.6-3.4 cm coriaceis ob- lanceolatis acutis basi cuneata, cymis 6 cm longis 2-floriferis, pedunculis 15 mm longis, pedicellis 15 mm longis, lobis calycis 5 cm longis petaloideis, lobis cor- Ollae 5 cm longis viridibus, floribus arcuate decur- vatis, baccis 2 cm longis. Typus: Hawaii I., Makaopuhi, H. Hoshide & S. Kawaloa 2-5. C. viridis sp. nov. (sect. Clermontia). Fig-" 7m Diagnosis Holotypi: Frutex est, caule hirsutulo, petiolis 4-4.5 cm longis hirsutulis, laminis 21-23 6.5-7.5 cm chartaceis ellipticis cuneatis serrulatis infra in nervis puberulis, cymis 11 cm longis 2-flor- iferis puberulis, pedunculo 4 cm longo, pedicellis 3 cm longis, lobis calycis in gemma 4.5 cm longis petaloideis decurvatis. Typus: Hawaii I., Makakupu, P.. Conant (sacob2 no- 630.)- C. waikoluensis sp. nov. (sect. Clermontioideae), ELqe 2. Diagnosis Holotypi: Frutex ramosus est, laminis 9-15 X 2-3.6 cm oblongo-ellipticis, inflorescentia glabra 2-florifera, pedunculo 10-22 mm longo, pedicellis 25-42 mm longis, lobis calycis 2 mm longis deltodideis, corolla 6.5-7.5 cm longa subviridi, baccis 25 mm vel ultra diametro.Typus: Molokai I., Waikolu, H. St. John 25 p22 2e LEGEND Fig. 1. Clermontia albimontis St. John. Fig. 2. Clermontia bicolorata St. John. Fig. 3. Clermontis epilosa St. John Fig. 5. Clermontia gracilis St. John. il 2 3 Fig. 4. Clermontia glabra St. John. 5 Fig. 6. Clermontia hualalaiensis St. John. 1987 Fig. Fig. Fig. Fig. Fig. Fig. St. John, Hawaiian Clermontia 353 7. Clermontia kahuaensis St. Jobn. 8. Clermontia kipahuluensis St. John. 9. Clermontia mauiensis St. John. 10. Clermontia pluriflora St. John. 11. Clermontia viridis St. John. 12. Clermontia waikoluensis St. John Vol. 63, Nores Pi hoe Ts Oy 170) Golo A 354 12cm Clermontia albimontis St. John ie Fig. 359 St. John, Hawaiian Clermontia 1987 John © Clermontia bicolorata St. 2 Fig. 5 Vol. 63, No. FeHeyY 10 LOG) ExA 356 1cm Clermontia epilosa St. Jonn By Fig. 1987 St. John, Hawaiian Clermontia 357 bo Cre Rae ees bes, Dyin TAT Sau ee pease Gee: Hruww, Sane - wen ° Simm Fig. 42/Clermontta’ oqlabra St. John 5 Vol. 63, No. PHY. TO} OGL CA 358 ey eo ee San ars wre : oo BAC aA ot " nog ’ ‘ i T) “ -—_—— = bees Ree Dy) daa ce) acm = 2 he Py, EUnet as Veen esa % 3 ae senan siieea gh asi ESS 5 eh C oo ae ele itd os Sry Sea = Sage e Sars Bes N eset Ce ery: “neste, ae Je ENG “er -5. Clermontia, gracilis St. John 1987 St. John, Hawaiian Clermontia 359 Fag.) 6.7Clermontia, hualalavensis St. doan 360 Pony 1, OP OTC rr A Vol. 63; No? Fig. 7. Clermontia kahuaensis St. John 1987 Fig. B. St. John, Hawaiian Clermontia 361 Clermontia kipahuluensis St. John 362 P BYT, 0. L0G) 1A Vol. 63, Now5 Fag. 9). Cc 5S mm pL Clermontia mauiensis St. John 1987 St. John, Hawaiian Clermontia LO. Clernontiapluratilora iSt. 363 John 364 PLAY EO LO Gila Vol. 63, No. Fig. Ll! Clermontia viridis St.) ‘Joma St. John, Hawaiian Clermontia 365 Lo Clermontia waikoluensis St. John A HALEAKALA VARIETY OF LOBELIA (LOBELIACEAE) HAWAIIAN PLANT STUDIES 147 Harold St. John & A. C. Medeiros =Bishop Museum, Box 19000A, Honolulu, Hawaii 96817, USA Lobelia Gaudichaudii A.P.DC., var. albiflora var. nov. Diagnosis Holotypi: a var. longibracteata Rockdiffert in pedicellis hypanthioque hirsutulis, corollis 9-9.2 cm longis albis. Holotypus: Hawaiian Islands, Maui Island, Haleakala, n. w. rift, above Mid- Camp bog, on bog margin, 5.600 ft alt., 6/13/82, A... Medeiros. 4..(BISH). Comparison: Of the new variety, the closest relative is L. Gaudichaudii, var. longibracteata which has a stat- ure of 3-5 m; blades 18-20% 3-3.5 cm, elliptic oblong, bracts of spike 6-7 KX 0.6-0.9 cm, apparently greenish; pedicels glabrous; hypanthium glabrous; calyx lobes puberulous ciliate near the base; corolla 6.5-7.5 cm long, cream-colored with purplish streaks; anthers all penicillate with bristles 1.5-1.7 mm long. The var. albiflora has a stature of 2.6-3.3 m; blades 21-22 XK 2.5-2.9 cm, narrowly oblanceolate; bracts of spike 6.2-6.5 ¥ 0.9-1 cm, white; pedicels hirsute; hypanthium hirsutulous; calyx lobes puberulous ciliolate through- out; corolla 9-9.2 cm long, white; anthers all penic- illate with bristles 2-2.5 mm long. The new epithet is formed from the Latin words alba, white, and flora, flower, and it is chosen with reference to the flower color. 366 DIAGNOSES OF ROLLANDIA SPECIES (LOBELIACEAE) HAWAIIAN PLANT STUDIES 148 Harold St. John Bishop Museum, Box 19000A, Honolulu, Hawaii 96817, USA The types of the following species are in the Bishop Museum ROoliandia alba St. John & W. Takeuchi, sp. nov. Diagnosis Holotypi: Laminae 45.5 X 17 cm glabrae chartaceae oblanceolatae deltoideae sunt, cymis 8-10 cm longis 7-10-floriferis, pedunculo 12-22 mm longo, ped- icellis 12-15 mm longis puberulis, lobis calycis 3-4 mm longis late ovatis, corollis 5 cm longis albis, antheris superis 9 mm longis. Typus: Oahu I., Waianae Mts., Puu Hapapa, D. Paquin. Discussion: The nearest relative is R. longiflora Wawra, of Oahu, a species with the petioles 2.5 cm long; blades 30-40 6-8 cm, fusiform, entire or dentate to laciniate towards the base; raceme 3 cm long; calyx lobes 1-3 cm long; and the corolla 6-6.5 cm long. R. alba _ has the pet- ioles 6 cm long; blades 45.5% 17 cm, oblanceolate; racem- es 8-10 cm long; calyx lobes 3-4 mm long; and the corolla 5 cm long. Roliandia Obatae sp. nov. Diagnosis Holotypi: Petiolae 3-5 cm longae sunt, laminis 36-41.5 X 6-8 cm angustiore ellipticis acutis cuneatis infra puberulis, racemis 2-4 cm longis puberulis, pedicellis 12-17 mm longis, lobis calycis 2-3 mm longis Ovatis puberulis, corollis 7.5 cm longis, antheris glabris eis superis 15 mm longis. Typus: Oahu I., Mt. Kaala, J. K. Obata 77-316. Discussion: The most similar relative is R. calycina (Cham.) G. Don, var. kaalae (Wawra) E. Wimm., but it has the calyx lobes 4-6 mm long, rounded, glabrous or pub- erulent;corolla 5-5.5 cm long; anthers pubeulous on the sutures; and the blades bellow rusty scabrous short puberulous. 367 DIAGNOSES OF PANICUM SPECIES (GRAMINEAE) HAWAIIAN PLANT STUDIES 349 Harold St. John Bishop Museum, Honolulu, Box 19000A, Hawaii, USA The following Hawaiian species of Panicum are novelties. Unless otherwide located, all the types are in the Bishop Museum, Honolulu. Panicum annuale sp. nov. (sect. Depauperata). Planta annua 7-12 cm alta caespitosa est, vaginis pilosulis, laminis 1 mm latis, spiculis 1.5-1.8 mm longis, gluma exteriori 1.5-1.8 mm longa glabra. Typus: Maui I., Kahakuloa, R. Sylva. P. assurgens sp. nov. (sect. Depauperata). Planta annua 36 cm alta est, vaginis pilosulis, laminis 3-4 mm latis pilosulis, spiculis 1.5-2 mm longis glabris, gluma exteriori 1.5-2 mm longa lanceolata. Typus: Lanai I., Kaumalapau Hbr., R. Hobdy 1,344. Panicum baltodes sp. nov. (sect. Virgata), Figs. 1, 2. Diagnosis Holotypi: Planta perennis subcaespitosa 15 cm alta est, rhizomatibus pluribus, vaginis 10-20 mm longis pilosis, panicula 3-4 cm longa, ramis puberulis, Spiculis 2-2.3 mm longis glabris 2-floriferis, gluma se- cunda 1.9 mm longa 7-nervosa, lemma 2 mm longa 7-nervosa. Typus: Typus: Kauai I., Alakai Swamp, v. Balgooy 4,213 (L), isotype (BISH). Panicum bifurcatum sp. nov. (sect. Depauperata). Planta annua 37 mm alta est, vaginis pilosis, laminis 2-2.5 mm latis involutis pilosis, spiculis 2.3- 2.5 mm longis glabris, gluma exteriori 2.3-2.5 mm longa lanceolata. Typus: Molokai I.,Pohakumauliuli, N. Pekelo Jr. 31. Panicum conjugens Skottsb., Acta Horti Gothob. 15: 296, £10. 0G; 1944, Lectotype: Kauai, bog along trail from Lihue Maka- noi toward Kilohana, 13/8/38, 0. Selling 2,886 (BISH). Skottsberg published this species based on two and a possibel third collections. The number 2,886 is here chosen as the lectotype. Panicum Cookei sp. nov. (sect. Depauperata). Diagnosis Holotypi: Planta annua perennisve 60 cm alta est, culmis 30 cm altis pilosulis, vaginis pilosis, laminis 4 mm latis planis infra pilosulis, paniculis 15-20 cm longis ramis pilosulis, spiculis 3-3.3 mm longis lanceoloideis glabris, gluma exteriori longissima 3-nervosa. Typus: Molokai I., Kaunakakai, E. Y. Hosaka 1.861. 368 1987 St. John, Hawaiian Panicum 369 Panicum ekeanum sp. nov. (sect. Turfosa). Fig. 3. Diagnosis Holotypi: Planta perennis caespitoSa est, culmis 6-7 cm altisglabris, vaginis 4-7 mm longis mar- ginibus apicalibus ‘ciliolatis, laminis 0.8-1.3 mm latis supra in basi pilosulis, racemis 2-5-floriferis, ped- icellis 2-3 mm longis pilosulis, spiculis 1.8-2.3 mm longis, gluma exteriori 0.5-0.6 mm longaovata, gluma interiori 1.8-2.3 mm longa elliptica acuta 7-nervosa. Typus: Maui i., Mt. Eke, O. Degener 10,830. Panicum furtivum sp. nov. (sect. Depauperata). Diagnosis Holotypi: Planta annua 11-19 cm alta est, vaginis 3-4 cm longis, laminis 2-3 mm latisbilosulis, spiculis 1.7-1.9 mm longis, gluma exteriori 1.7-1.9 mm longa lanceolata. Typus: Hawaii I., Kealakahiki Pt?) Wo Ghar, 76,079 - Panicum Hobdyi sp. nov. (sect. Turfosa), Figs. 4, 5A. Diagnosis Holotypi: Planta perennis caespitosa est, culmis 5-10 cm altis, vaginis 3-10 mm longis glabris, laminis 0.8-1 mm latis involutis supra pilosulis, inflorescentia 7-15 mm longa pilosula 1-4-florifera, spiculis 1.9-2.3 mm longis gluma exteriori 0.8-1.2 mm longa ovata3-nervosa. lemma fertili 1.7 mm longa 3=nervosa,. Typus: Maud 1.., Eke trail, oR. Hobdy 707. Panicum honokowaiense sp. nov. (sect. Monticola?), BAGS) Obie Oe Planta perennis lignosa est, culmis 70 cm longis decumbentibus glabris, vaginis 3-6.5 cm longis glabris, laminis 7-11 mm latis , paniculis 22 cm longis, spiculis 1.6-1.8 mm longis lanceoloideis glabris, gluma exteriori 1.6-1.7 mm longa lanceolata 5-nervosa, illa interiori 1.8-1.9 mm longa 7-nervosa. Typus: Maui I., Honokowai, R. Hobdy 777. Panicum infraventale sp. nov. (sect. Turfosa), Fig. 7. Planta perennis caespitosa est, vaginis sub- glabris, laminis 1.4-4.5 mm latis subglabris, spiculis 1.5 mm longis ellipsoideis glabris, gluma exteriori 0.6 mm longa. Typus: Kauai I., Kahoaluamanu, C. N. Forbes 402.K. Panicum kahiliensis sp. nov. (sect. Turfosa?). Diagnosis Holotypi: Planta perennis caespitosa30-38 cm alta est, vaginis ciliatis, laminis 4-5 mm latis, spiculis 2.5-3 mm longis ellipsoideis glabris, gluma exteriori 1 mm longa ovata. Typus: Kauai I., Kahili Mb Bae Oconee no D 4. Panicum kahoolawense sp. nov, (sect. Depauperata). Diagnosis Holotypi: Planta annua 6-8 cm alta est, vaginis subglabris, laminis 1-1.2 mm latis ciliatis et villosis, spiculis 1.2-1.3 mm longis ellipsoideis, gluma exteriori 1.2-1.3 mm longa in apice puberula. Typus: Kahoolawe I., L. W. Cuddihy 343, - = 370 PHY, 7 01 0) Gra Vol. 63, No. 5 Panicum kanaioense sp. nov. (sect. Trichoidea). Diagnosis Holotypi: Planta annua 10 cm alta est, vag- inis pilosis, laminis 3-3.5 mm latis glabris, spiculis 1.8-2 mm longis pilosis, gluma exteriori 1.8-2 mm longa. Typus: Maui I., Kanaio, R. Hobdy 1,273. Panicum kaonohuaense sp. nov. (sect. Depauperata). Diagnosis Holotypi: Planta annua 50 cm alta est, vaginis pilosis, laminis 4-5 mm latis infra villosis, spiculis 2.5-3 mm longis lanceoloideis glabris, gluma exteriori 2.5-3 mm longa. Typus: Maui I., Konohua Gulch, R. Hobdy 805. Panicum Knudsenii sp. nov. (sect. Turfosa). Diagnosis Holotypi: Planta perennis 16 cm alta est, vaginis pilosulis, laminis 4-7.5 mm latis ciliatis, spiculis 2-2.2 mm longis ellipsoideis glabris, gluma exteriori 0.7-0.8 mm longa. Typus: Kauai I., Alakai, GaN. Forbes v6 /o.Ke Panium Kokeeense sp. nov. (sect. Virgata). Diagnosis Holotypi: Planta perennis glabra caespit- osa 3-4 cm alta est, vaginis ciliatis, laminis 3-5 mm latis, spiculis 2-2.2 mm longis ellipsoideis, gluma ex- teriori 0.5 mm longa ovata. Typus: Kauai I., Kauluweki bog, A. C. Medeiros 502. Panicum kukaiwaaense sp. nov. (sect. Depauperata). Diagnosis Holotypi: Planta 3.5-9 cm alta pulvin- osa breve vivens est, culmis vaginis laminis panic- ulisque puberulis, laminis involutis in aspecu 0.3-0.5 mm latis, paniculis 10-20 mm longis, spiculis 2-2.3 mm longis lanceolatis, gluma exteriori 2-2.3 mm longa et in 3 nervis microscopicis adpresse puberula. Typus: Molokai I., Kukaiwaa, R. Hobdy 2,184. Panicum lihauense sp. nov. (sect. Depauperata), Bigs. 18) 9 Diagnosis Holotypi: Planta perennis 15-20 cm alta caespitosa est, basi dura incrassata, culmis glabris, vaginis 13-20 mm longis, laminis 2.5-2.7 mm latis ligulatis glabris arte involutis, panicula 8-12 cm longa subcompactasparse puberula, spiculis 2.6-3 mm longis lanceoloideis pilosis, gluma exteriori 2.6-3 mm longa. Typus: Maui I., Lihau Peak, R. Hobdy 825. Panicum lineale sp. nov. (sect. Virgata), Figs. 10, ll. Diagnosis Holotypi: Planta perennis caespitosa est, culmis 48 cm altis glabris, foliis omnibus basalibus, vaginis 8,5-15 cm longis glabris except in apice pil- osulis, laminis 5 mm latis involutis glabris, paniculis 22-34 cm longis scabris. spiculis 3.8-4 mm longis lanceo- latis glabris, gluma exteriori 2 mm longa 5-nervosa. Typus: Kauai I., Kulanalilia, C. Christensen 324. Panicum longivaginatum sp. nov. (sect. Virgata), Figs. 12, (SA. 7a Diagnosis Holotypi: Planta 62-71 cm alta fere om- nino glabra licet oerennis est, vaginis 5-11 cm longis 1987 St. John, Hawaiian Panicum 371 marginibus superis pilose ciliatis, laminis 10-13 mm latis panicula 11-13 cm longa, spiculis 2.8-3.1 mm longis fusi- formi-ellipsoideis, gluma exteriolri 0.3-0.6 mm longa flabellata. Typus: Hawaii I., Upper Waiakea Forest, R. Gustafson 2,400. Panicum malikoense sp nov. (sect. Depauperata). Diagnosis Holotypi: Planta annua 5-10 cm alta est, culmis glabris, vaginis 8-12 mm longis puberulis, laminis 1-1.2 mm latis involutis glabris, spiculis 1.8-2.1 mm longis lanceoloideis glabris, gluma exteriori 1.8-2.1 mm longa ovat> lanceolata. Typus: Maui I., Maliko Gulch, £2/19176, KR. Sylva. Panicum mokuleiaense sp. nov. (sect. Depauperata?). Figs. 4,25. Planta perennis est, culmis 21-30 cm altis in basi tuberosis, vaginis 4.5-6 cm longis glabris, laminis 3-4 mm latis involutis, panicula 9-12 cm longa, spiculis 2.3- 3 mm longis lanceolatis villosis, gluma exteriori 2.5- 3 mm longa. Typus: Oahu I., Mokulej;a, H. St. John 23,692. Panicum nephelophilum Gaud., var. levius var. nov. Figs) 13B;, 15. Diagnosis Holotypi: Planta perennis 68 cm alta est, caudici incrassata, vaginis 5-9 cm longis glabris, laminis caulinis 6-9 mm latis glabris, panicula 20 cm longa, spiculis 2-2.3 mm longis lanceolatis, gluma exter- iori glabra alteram excedenti. Typus: Maui I., Paunau-Kuhua, H. St. John et al. 17,702. Panicum ninoleense sp. nov. (sect. Depauperata).Fig. 17. Diagnosis Holotypi: Planta annua 6-12 cm alta est, vaginis 1-2 cm longis puberulis, laminis 1 mm latis involutis supra pilosulis infra puberulis, panicula 1.5-3.5 cm longa, spiculis 1.5-2 mm longis pilosulis et villosis, gluma exteriori 1.5-2 mm ionedidates: ovata. Typus: Hawaii I., Ninole, H. St. John et al. 23,954. Panicum ooense sp. nov. (sect. Virgata). Diagnosis Holotypi: Planta perennis 25-30 cm alta est, vaginis pilosis, laminis 6-7 mm latis ciliatis, spiculis 2.8-3 mm longis ellipsoideis glabris, gluma exteriori 0.6-1.3 mm longa ovata. Typus: Hawaii I., Puu Oo, C. N. Forbes 811.H. Panicum pepeopaeense Stat,. nov. (sect. Turfosa). P. imbricatum Hillebr., var. molokaiense Skottsb., Acta Horti Gothob. 15: 290, Figs. 52-66, 1944, non P. molokaiense Deg. & Whitney in Dea. 1936. Lectotypus: Molokai I., Pepeopae, L. M. Cranwell 3,816 (GB). Panicum simplex sp. nov. (sect. Depauperata). Diagnosis Holotyp3: Planta annua 25-36 cm alta est, 372 PH Y “E07 OG. 1 A Vol. 63, No. 5 vaginis pilosis, laminis 2.4-3 mm latis pilosis, spiculis 2.2-2.5 mm longis glabris, gluma exteriori 2.2-2.5 mm longa. Typus: Lanai I., Hulupoe Bay, L. Stemmermann 3,711. Panicum subglabrum sp. nov. (sect. Depauperatum). Diagnosis Holotypi: Planta annua 58-"100" cm alta est, vaginis puberulis, laminis 5-8 mm latis puberulis, spiculis 3.3-4.2 mm longis lanceolatis glabris, gluma esteriori longiora. Typus: Lanai I., Poopoo Islet, R. Hobdy 1,357. Panicum Sylvanum sp. nov. (sect. Depauperata, Fig. 18. Diagnosis Holotypi: Planta annua 5-10 cm alta ramosa est, culmis minute puberulis, nodis puberulis, folinis caulinaribus multis, vaginis 6-11 mm longis minute puber- ulis, laminis 0.8-1.2 mm latis involutis minute puber- ulis, paniculis 15-25 mm longis pilosulis, spiculis 1.7-2 mm longis lanceoloideis glabris, gluma exteriori 1.7 mm longa elliptica 5-nervosa. Typus: Maui I., Maliko Bay, R. Sylva. Panicum waikoloaense sp. nov. (sect. Depauperata}), Fig. 19. Diagnosis Holotypi: Planta annua 9-18 cm alta eramosa est, nodis pilose cinctis, folia basali ca 23 cm longo, 3-4 foliis caulinibus 25-35 mm longis, laminis 1.5-2 mm latis pilose ciliatis supra pilosis infra sparse pilosulis, paniculis 2-5 cm longis pilosulis, spiculis 1.5-2 mm longis ellipsoideis acutis in dimidio basali pilosulis, gluma exteriori 1.5-2 mm longa late ovata 3-nervosa. Typus: Hawaii I., Puu Hononaohae, K. M. Nagata 2,432. Panicum waimeaense sp. nov. (sect. Turfosa). Diagnosis Holotypi: Planta perennis caespitosa 4-5 cm alta est, culmis exsertis sparse pilosulis, vaginis sparse pilosulis, laminis 1-1.5 mm latis, racemis 4-7- floriferisinfra pilosulis supra puberulis, spiculis 1.7- 1.8 mm longis glabris, gluma exteriori 0.4-0.5 mm longa hemisphaerica umbonata. Typus: Kauai I., Waimea, CC: Ne Forbes 1, 707ask. Panicum Wilburi sp. nov. (sect. Turfosa), Figs. 20, 21. Diagnosis Holotypi: Planta perennis 25-29 cm alta est, culmis glabris, foliis caulinaribus multis, vaginis internodos excedentibus pilose ciliatis, laminis 4-7 mm latis, paniculis 4-5.5 cm longis, spiculis 2.2-2.3 mm longis glabris, gluma exteriori 0.8-1 mm longa ovata l-nervosa. Typus: Kauai I., Alakai Swamp, H. St. John et als 23,053). Panicum wiliwilinuiense sp. nov. (sect. Depauperata). Diagnosis Holotypi: Herba perennis 15-20 cm alta est, vaginis glabris, laminis 5-9 mm latis glabris, spiculis 1.2-1.5 mm longis ellipsodeis, gluma exteriori 1.2-1.5 mm longa elliptica. Typus: Oahu I., Waialae Iki, Ys. Kondo. 1987 St. John, Hawaiian Panicum 373 LEGEND Fig. 1. Panicum baltodes St. John, a, habit, X 1; b, sheath and ligule, X 20; c, panicle, X 4. Fig. 2. Panicum baltodes St. John, d, node and shoot. X 10; e, spikelet, X 5; £, lst glume, X 15; g, 2nd glume, Kis: h, sterzie Jemma, 7x 15; 2, fertile iemma, «x 25; Jj, Pal€agixis. Fig. 3. Panicum ekeanum St. John, a, habit, X 1; b, panicle, X 4; c, sheath and ligule; xX 5; d, basal leaves, X 10; e, spikelet, X 15; £, 1st glume, X 15; g, 2nd glume, X 15; kh, lemma, X 15; i, palea and grain, X 15. Fig. 4. Panicum Hobdyi St. John, a, habit, X 1; b, raceme, X 4; c, spikelet, X 15. Pe Fig. 5A. Panicum Hobdyi St. John, d, stem and sheaths, x 107, e, Ist glume; x i57 £, 2nd glume, X 15; g, sterile lemma, X 15; h, fertile, lemma,»X 15; i, palea, Sees: j, prstil, x 15. Fig. 5B. Panicum honokowaiense St. John, k, sheath and ligule, X/10;, 1, fertile lemma; ‘X°157¢4m,;palea, .x.15. Fig. 6. Panicum honokowaiense St. John, a, habit, X %; b, panicle branch, X 4; c, spikelet, X 15; d, lst glume, xX 15; e7 2nd glume, X15; £;sterile temma,, X15. Fig. 7. Panicum infraventale St. John, a, habit, X 1; b sheath and ligule, X 10; c, sheath and ligule, xX 4; d, raceme, X 4; e, spikelet, 15; EE, »lstyglume;. x15; og) * 2nd glume, X 15; h, fertile lemma, X 15; i, palea, X 15; dy PL1SGil, EX.15. Fig. 8» Panicum lihavense St.,.John; a, habit, x 1; b, panicle, x4; cc; Ist glume, xX 15. Fig. 9. Panicum lihauense St. John,'d, sheath and ligule, X 10; e, sheath, ligule, and blade, X 10; f, spikelet, X 15; 9g, lst. glume, X 15; h,,fertile lemma, xX 15); ay paléa, X15. Fig. 10. Panicum lineale, St. John, a, habit, X %; b, spikelet, X 15; ¢, Ist, glume, x 15; d, 2nd glume ,//xv15; e, sterile lemma, X 15; £, fertile lemma, X 15; g, palea, xX 15% Fig. 11. Panicum lineale St. John, h, sheaths and lig- ule), X 10; 3, Jagule, X 15s 3, ‘panicle branch; wx-4; k, prstal, xs Si. Fig. 12. Panicum longivaginatum St. John, a, habit, X 1/3; b, panicle branch, X 4; c, spikelet, X 15; a; 1st glume, X 15; e,2nd glume, X 15; £, fertite denma, xX 15+ hag. aA Panicum longivaginatum St. John, g, palea, X 15;\-h, pistil, X15; 2, blade base, xX 10: Fig. 13B. Panicum nephelophilum Gaud., var. levius st. John, £, Ist glume, X 15; g, 2nd glume, x i5; 4h, fertile lemma, X 15; i, palea,; XK 155.5, Sheath and ligule, X 10. 374 PH YD eO\sis (OuG viva Vol... 63, No. “5 Fig. 14. Panicum mokuleiaense St. John, a, habit, X 1; b, fertile lemma, X 15; c, pistil, X 15. Fig.15. Panicum mokuleiaense St. John, d, sheath, ligule, and blade, X 10; e, panicle branch, X 4; £, spikelet, X 153g, 1st glume, X 15; h, 2nd glume, X 157 1; *Steresie lemma, X 15. Fig. 16. Panicum nephelophilum Gaud. var. levius St. John, a, ‘habie, x 1/4: b, sheath, ligules and blades, X 10; c, panicle branch, X 4; d, spikelet, X 15; e, anther, X 15. Fig. 17. Panicum ninoleense St. John, a, habit, X 1; b, sheath and ligules, X 10; c, sheath and ligules, X 10; d, panicle, X 4; €, spikelet, X*157°£)"ise glume, X 15; g, 2nd glume, X 15; h, sterile lemma, X 15; 1,/ fertile lemma, X 15. Fig. 18. Panicum Sylvanum St. John, a, habit, X 1; b, sheaths, X 10; c, sheath and ligule, X 10; d, panicle, X 4; e, spikelet, X 15; £, lst glume X 15: g, 2nd glume, X 15; h, sterile lemma, X 15;'\1, fertile lemma, seeis, j, paiea,’ X 15;°K, anthers, xX 15- Fig. 19. Panicum waikoloaense St. John, a, habit, X 1; b, panicle branch, X 4; c, leaf and sheath, X 10; d, sheath and ligule, X 10; e, lst glume, X 15; £, 2nd glume, X 15; gq, fertile’ lemma, X 15; h, palea,/” x 15; i, grain, “X 15; 7, stamens, “X'15. Fig. 20. Panicum Wilburii St. John, a, habit, K 4% b, sheath and blade, X 10; c, lst glume, X 15; d, 2nd glume, X 15; e, sterile lemma, X 15; f, fertile lemma, x oS: Fig. 21. Panicum Wilburii St. John, g, panicle branch, X 4; h, spikelet, X 15; i, palea, X 15; Jj, stamen, X 15. St. John, Hawaiian Panicum 37/5 1987 John Panicum baltodes St. i Eig. 5 Vol. 63, No. Roney 10 0) (GEA 376 John Panicum baltodes St. 2s Figns 1987 St. John, Hawaiian Panicum = 4™mM [PE alee eee Sees etm es! 3. Panicum ekeanum Si Sep ne lalal 1Cm Sy Vol. (63: Nowes Pi Yel. O G) tA 378 ZB Wi VA Yj, ‘ie Ki Viger Hd Li Fig, 4, Panicum Hobdyi St. John 1987 St. John, Hawaiian Panicum 379 Fig. SA. Panacum)Hobdyi St. John fo) 2mm \\ Pe ie ee Se ee pou wWU tiie k (6) 4MM | Ey een eae eee | Fig. SE. Panicum hHonokoOwalense St. John 380 Poy Teo EO Guia A Vol. 63, Noe. 5 2MM Fig. 6. Panicum honokowaiense St.John 381 St. John, Hawaiian Panicum 1987 Panicum infraventale St. John ee gnc me | i | 1987 WW 7 St. John, Hawaiian Panicum Panicum lihauense: St. John 383 Fig. 10. Panicum lineale St. demm St. John, Hawaiian Panicum Panxeum "ineate Sit. 385 1CM ee ee eee John 386 Pag. Ds St/ John Po HLY 5.0 L 0.6 TA Vol. 63, No. WWT fo} Panicum longivaginatum 5 1987 St. John, Hawaiian Panicum 387 pe | ) WW 7 | | | | 4 | eS ee ee Pig. PSB. Panrteum nephelophilum Gaud. Vain, ‘Lev Lws «Sb gohn 1987 Vis St. John, Hawaiian Panicum tte ye N Panicum mokuleiaense St. John 389 390 Pes PibYot sO) JOuei A Vol. 63, No. 6 Mh | ! nt LAY ; AY Wa eae Mt ' y ul ig fo) 2 MM fe) 41MM "i fii Gye ha Baa] ites ; See Panicum nephilophilum Gaud. Var, levius St. John 1987 St. John, Hawaiian Panicum 391 Pah lf 8 Panicum ninoleense St. Jenn 392 Pol YT O° 0 G.rnA Vol. 63, ‘No.5 393 St. John, Hawaiian Panicum 1987 TDP nae ~ = a eee Panicum waikoloaense St. John POs Fig. 394 P HY. T Oe O..C TA Vol. 63, Now \ SAR fi < / : LLL LT * , P5 ; , PS Nes het Lala ie ff ff y ‘ af rear Aah : ae A AD ay gr ; / 3 a3 feels Lae ad / : “alas aoe y, ¥ tae y if SIA e = Jf Hf. 1987 Fig. 2s St. John, Hawaiian Panicum Panicum Wilburni “St. 3 MM John 395 A NEW SPECIES OF PERYMENIUM (ASTERACEAE-HELIANTHEAE) FROM TAMAULIPAS, MEXICO B. L. Turner Department of Botany, Univ. of Texas, Austin, TX 78713 Treatment of the Asteraceae of Mexico (Turner & Nesom, in prep.) necessitates description of the following species of Perymenium: PERYMENIUM TAMAULIPENSE B. Turner, sp. nov. Fig. l P. grande Hemsl. simile sed habitu perenni, petiolis brevioribus (2-5 mm longis), et capitulis paucioribus (2- 4) in pedunculis longioribus (5-6 cm longis) differt. Perennial herb to 1 mm high. Stems terete, striate, strigillose to glabrate. Leaves opposite, 6-10 cm long, 3-5 cm wide; petioles 2-5 mm long; blades deltoid, 3- nervate from the base, appressed-strigose above and beneath, rough to the touch, the margins serrulate. Heads 2-4 in terminal lax cymes, the ultimate peduncles strigose, 5-6 cm long. Involucre campanulate, 7-8 mm high, 7-9 mm wide, 3-4 seriate, imbricate; bracts broadly ovate, 3-8 mm long, strigillose with ciliate margins, the apices obtuse to rounded. Chaff linear-lanceolate, scarious, yellowish. Ray florets 10-1ll, pistillate; corollas yellow, the ligules 7-10 mm long. Disk florets 30-40; corollas yellow, glabrous, 4-5 mm long. Achenes (immature) pubescent, ca 1.5 mm long; pappus of an elongate bristle 3-4 mm long, and 8-10 smaller bristles, 1 mm long or less. TYPE: MEXICO. TAMAULIPAS: Camino de cd. Victorisa al Molino, 30 km al W de Victoria, Bosque de Pinos Encino, ca 880 m, 23 Sep 1985, Manuel Yanez 463 (TEX; isotype UAT). Fay (1978) cites not a single species from the state of Tamaulipas. In his treatment the present collection will key to those species with an evidently imbricate involucre. Among these it appears to be closest to P. grande (in characters of the involucre) but has the habit of P. gymnoloides. The latter species occurs along the Gulf Coastal region of eastern Mexico to near the Tamaulipan border but it is readily distinguished from P. tamaulipense by its smaller, more numerous, heads on much shorter peduncles. 396 sy a 1987 Turner, Perymenium tamaulipense sp. nov. 397 ACKNOWLEDGEMENTS Guy Nesom provided the Latin diagnosis and Linda Vorobik prepared the excellent illustration. LITERATURE CITED Fay, J.J. 1978. A revision of Perymenium (Asteraceae- Heliantheae) in Mexico and Central America. Allertonia 1:235-296. 398 P HeY,T O.L/O.G.T A Vol. 63, Norgs Perymenium tamaulipense, from holotype. THE ALGAE OF NEW JERSEY (U.S.A.) XIII. CHLOROPHYTA (GREEN ALGAE). D. ZYGNEMATALES (ZYGNEMATACEAE AND MESOTAENIACEAE ) MaryAnn Foote College of Mount St. Vincent Rivercale, New York 10471 This is the thirteenth paper in the series examining the distribution of algae in the state of New Jersey. The genera are listed alphabeticaily and the collection dates chronclogically within them. Further ecological data may be obtained from the original sources. Publication was made possible by a Faculty Development Grant, College of Mt. St. Vincent. CHLOROPHYTA GREEN ALGAE ZYGNEMATALES Mougeotia capucina (Bory) Ag state (3 Mougeotia genuflexa (Diilw) Ag state Mougeotia nummuloides (Hass) DeToni state (3 Mougeotia quadranguiata Hass state (3 Mougeotia robusta (De Bary) Wittr state Mougeotia scalarils Hass state (3) Mougeotia sphaerocarpa Wolle ponds (1); Sussex Co (2); state (3) Mougeotia tenuis (Cleve) Wittr State (3) Mougeotia veridis (Kutz) Wittr State (3); D/R canal (6) Pleurodiscus purpureus (Wolle) Lager state Spirogyra bellis (Hass) Cleve streams and ponds (2); state (3) Spirogyra calospora Cleve streams and ponas (2) 399 400 PHY TOLOCGTHLA Vol. 63, No. Spirogyra cleveana Transeau New Brunswick golf course ditch (7) Spirogyra communis (Hass) Kutz Streams and ponds (2); state (3) Spirogyra condensata (Vauch) Kutz streams and ponds (2); state (3) Spirogyra crassa Kutz streams and ponds (2); state (3) Spirogyra fluviatilis Hilse streams ana pon Spirogyra grevilleana (Hass) Kutz streams and ponds (2); state (3) Spirogyra inflata (Vauch) Kutz Streams and ponds (2); state (3) Spirogyra insignis (Hass) Kutz State (1,3); streams and ponds (2) Spirogyra longata (Vauch) Kutz streams and ponds (2); state (3) Spirogyra majuscula Kutz streams and ponds (2) Spirogyra nitida (Dillw) Link state (2, Spirogyra porticolis (Muller) Cleve state (3 Spirogyra protecta Wood state (3) Spirogyra punctata Cl Pleasant Miils (1,2); state (3) Spirogyra quinina (Ag) Kutz streams and ponds (2) Spirogyra rivularis (Hass) Rab Streams and ponds (2) Spirogyra setiformis (Roth) Kutz Streams ana ponds (2); state (3) Spirogyra tenuissima var rugosa Transeau state (5 5 1987 Foote, New Jersey algae XIII Spirogyra varians (Hass) Kutz streams and ponds (2); state (3) Spirogyra weberi Kutz state (3) zygnema cruciatumm (Vauch) Ag streams and ponds (2); state (3) Zygnema insigna (Hass) Kutz Streams and ponds (2); state (3) zygnema pectinatum (Vauch) Ag state zygnema purpureum Wolie common in ponds (1); streams and ponds (2) zygnema stellinum (Muller) Ag streams ana ponds (2); state (3) Zygnema tenue Rab state Zygnema vaucheria Ag streams and ponds (2) REFERENCES 1. Wolle, F. 1887. Freshwater Algae of the United States. Vol 1. The Commenius Press. Bethlehem, Pa. 364 p. 2. Britton, W. 1889. Catalogue of Plants Found in New Jersey. John L. Murphy Pub Co. Trenton, N.J. 3. Collins, F.S. 1928. Green Algae od North America. G.E. Stechert & Co. N.Y. 400 p. 4. 5 1928. Supplemental Paper. G.E. Stechert & Co. N.Y. 47 p. Dy . 1928. Second Supplememtal Paper. G.E. Stechert & Co. 106 p. 6. Renlund, R.W. 1953. Ph.D. dissertaticn. Rutgers University. New Brunswick, N.J. 7. Keller, J.M. 1954. Ph.D. Dissertation. Rutgers University. New Brunswick, N.J. 401 MIRTACEAS NICARAGUENSES I. Eugenia sanjuanensis Sp. nov. Pablo E. Sanchez Vindas Museo Nacional de Costa Rica Departamento de Historia Natural Apdo. 749, San José Costa Rica Eugenia sanjuanensis P.E. Sanchez 4p. nov. Fautex glaber, 2-4 metralis; ramulis glabris, teretibus. Folia glabra subtus glauca, oblongo-elliptica vel oblongo-Lanceolata, 23.5-35.8 cm Longa, apicem acutus, basim subauriculata vel rotun- data. Inflorescentia umbelliformis, axillaris; pedicekli dense porphyreus tomentosi; bracteolae 2, separatae, 2-2.5 mm Longae; cakycis Lobi 4, elliptico-triangularis, porphyreus tomentos., 3-5 mm Longi. Fructus obLongo-ellipsoideus, 1.2-2.5 cm Longus; Semina 1; embryo homogeneus. Arbusto de 2-4 m de altura; ramitas cilindricas, glabras, exfo liadndose en placas papiraceas y amarillentas. Hojas verde-oscuro en la haz, glaucas en el envés, oblongo-elipticas u oblongo-lan- ceoladas, 23.5-35.8 cm de largo, 6-11 cm de ancho, coriaceas, gla bras en ambas superficies, Apice agudo, base subauriculada o re- dondeada, nervio central escasamente elevado y con una sobresa- liente arista central en la haz, prominente y ferrugineo en el en vés, nervios laterales 11-15 de cada lado, muy separados entre si, prominentes y pardo-rojizos en el envés, nervio marginal arquea- do entre los laterales e igual de prominente que éstos, 5-12 mm del margen, a partir de éste los nervios terciarios forman un ner vio submarginal a 2-3 mm del margen; pecfolo cilindrico, glabro, engrosado, 8-10 mm de largo, hasta de 5 mm de ancho. Flores desco nocidas. Infructescencias con el eje no elongado por lo que los frutos parecen grupos umbeliformes, axilares; frutos 4-6; bracteas densamente pardo-rojizo tomentosas externamente, glabras interna- mente, de ovado-deltoides a ovado-redondeadas, 1-2 mm de largo; pe dicelos densamente pardo-rojizo tomentosos, 7-10 mm de largo, has- ta de 2 mm de ancho; bracteolas separadas, densamente pardo-rojizo tomentosas externamente, glabras internamente, ovado-redondeadas, 2-2.5 mm de largo; lébulos del caliz 4, céncavos, denso pardo-ro- jizo tomentosas externamente, glabras internamente, ovado-deltoi- des u eliptico-deltoides, 3-5 mm de largo, 2.5-4 mm de ancho; dis co usualmente redondeado, glabro excepto en la base del estilo, 3.5-4 mm de ancho. Frutos oblongo-elipsoidales, amarillos, de pardo-rojizo tomentosos a glabrados en la madurez, 1.2-2.5 cm de largo, 6-10 mm de ancho; semilla 1, embrién homogéneo con los co- tiledones y radficula no discernibles. 402 1987 Sanchez, Mirtaceas Nicaraguenses 403 TIPO: NICARAGUA. DPTO. RIO SAN JUAN: Quebrada Santa Crucita, 50 m. s.n.m; P. Moreno 23444 (Holotipo MO; isotipos: CR, F, NY, US). MATERIAL ADICIONAL EXAMINADO: NICARAGUA. DPTO. RIO SAN JUAN: La Gloria 3.5 Kms al NE del poblado de Boca de SAabalo, 70 m.s.n.m. P. Moreno 25431 (MO). Eugenia sanjuanensis parece esta relacionada con E. traunciflo- nya (Cham. & Schl.) O. Berg que se distribuye en Veracruz (México). Las principales diferencias entre ambas especies son las siguien- tes: E. sanjuanensis E. tunciflora Hojas 23.5-35.8 cm de (11.4-) 14-25.2 cm largo de largo inflorescencia axilar cauliflora indumento pardo-rojizo tomen- palido-pubescente toso bracteolas separadas unidas frutos pardo-rojizo tomen glabros, 2.2-3.5 cm tosos, 1.2-2.5 cm de largo de largo AGRADECIMIENTOS: A Douglas Stevens por su apoyo y facilidades en la revisi6n del material botanico de las Mirtadceas de Nicaragua; a la Bi61l. Luz Marfa Ortega por el mecanografiado del manuscrito. NUEVA ESPECIE DE EUGENIA L. (MYRTACEAE) DE VERACRUZ, MEXICO Pablo E. Sanchez Vindas Museo Nacional de Costa Rica Departamento de Historia Natural Apdo. 749, San José Costa Rica Luz Maria Ortega Torres Instituto Nacional de Investigaciones sobre Recursos Bidticos Apdo. 63, Xalapa, Veracruz México Eugenia uxpanapensis P.E. Sanchez & L.M. Ortega 4p. nov. Arbor ca. 15-20 m alta. Folia gkabna, elliptica vel elliptica- ovata, (3.9-) 4.5-8.3 cm Longa, apicem caudatus, acuminis falcatis, costa inmersa Subtus prominens; petioli glabro, 7-10 mm Longs. In- fLorescentia omnis dense cupreus-tomentulosus praeter stylus; ra- cemus udsque ad 10 mm Longus; flores 4-8; pedicellki 1.5-4 mm Longs; bracteolae separatae, 0.8-1 mm Longae; cakycis Lob 4, 1.5-2 mm Longi; ovarium bikocukare; ovula in quoque Loculo 5-8. Fauctus obkatus, 1-1.3 cm Longus, 1-1.5 cm diametro; semina |. Arbol de 15-20 m de altura, hasta de 30 cm de d.a.p.; ramitas jOvenes cilindricas, hispidulosas, con los pelos cortos y erectos, variando de padlidos a cobrizos; pelos de las ramitas y la inflo- rescencia diferentes, los de la inflorescencia muy densos, encris pados, cobrizos o rojizos, 0.2-0.5 mm de largo. Hojas verde-oscu- ras y opacas en el haz, verde-palidas en el envés, amplio-elipti- cas u ovado-elipticas, raramente obovadas, (3.9-) 4.5-8.3 cm de largo, 2.4-3.5 cm de ancho, con puntos glandulosos principalmente visibles en el envés, glabra en ambas superficies, 4pice caudado- acuminado, por lo general el acumen falcado y hasta 1.5 cm de lar- go, base de aguda a cuneiforme, con los margenes decurrentes sobre el peciolo, nervio central profundamente acanalado y glabro en el haz, prominente, glabro y amarillento en el envés, nervios la- terales 11-18 de cada lado, incluyendo algunos intermedios, para- lelos entre si y con los intermedios, escasamente elevados en el haz y en el envés, nervio marginal escasamente arqueado entre los laterales e igual de prominente que éstos, paralelo al margen, 1-1.5 mm del margen; peciolo profundamente acanalado, glabro, ama rillento, 7-10 mm de largo, 1-1.5 mm de ancho. Inflorescencias un racimo hasta de 10 mm de largo, 1-2 por axila; ejes densamente co brizo-tomentulosos, 1.5-7 mm de largo, produciendo 4-8 flores opuestas y decusadas; brdcteas de cobrizo-tomentulosas a glabra- das, deltoides, 0.5-1.2 mm de largo; pedicelos densamente cobri- zo-tomentulosas, 1.5-4 mm de largo; bracteolas separadas, densa- 404 1987 Sanchez & Ortega, Eugenia uxpanapensis sp. nov. 405 mente cobrizo-tomentulosas, ovado-deltoides, muchas veces inconspi cuas por el denso tomento del hipanto, 0.8-1 mm de largo; yemas globosas, densamente cobrizo-tomentulosas, 4-4.5 mm de largo, 4- 4.5 mm de ancho en el globo de los pétalos; hipanto densamente co- brizo-tomentuloso, campanulado o en forma de copa, 1.5-2.5 mm de largo; lébulos del caliz densamente cobrizo-tomentulosos externa e internamente, ovado-redondeados, céncavos, 1.5-2 mm de largo, 1.5-2 mm de ancho; pétalos ciliados, con glandulas céncavas, pardo- rojizas y prominentes, ovados; disco redondeado, palido o cobrizo- pubescente en el anillo estaminal, 2-4 mm de ancho; estambres 80- 100, de 4-6 cm de largo; ovario bilocular; 6vulos 5-8 en cada 16- culo; estilo glabro, 4-5 mm de largo. Frutos glabros, muy glandulo sos, de globosos a oblatos, 1-1.3 cm de largo, 1-1.5 cm de diame- tro, coronados en el apice por los cobrizo-tomentulosos lébulos del caliz; semilla 1. TIPO: MEXICO. VERACRUZ: Mun. Hidalgotitlan, Rio Soloxuchil, SO del Campamento Hnos. Cedillo, Brigada Vazquez 293 (Holotipo XAL; isotipos: CR, MEXU, MO) MATERIAL ADICIONAL EXAMINADO: MEXICO. VERACRUZ: Mun. Hidalgoti- tlan, Km 2-3 del camino a Plan de Arroyos a Alvaro Obregén, BAiga- da Donantes 2735 (F, MEXU, MO, XAL); Mun. Hidalgotitlan, Brecha Hnos. Cedillo-La Escuadra, Brigada Vazquez 99] (MEXU); Mun. Hidal- gotitlan, camino Cedillo-La Escuadra, Donrantes et al 3572 (F); Mun. Hidalgotitlan, Camino a La Escuadra, Km 11 de Cedillo, Donan- tes et ak 3698 (F, MEXU); Mun. Hidalgotitlan, alrededores del Campamento Hnos. Cedillo, Donantes et al 4110 (F); Mun. Hidalgoti- tldn, Rio Soloxuchil, SO del Campamento Hnos. Cedillo, Ponce 248 (F, MEXU, MO, XAL). Esta especie ha sido confundida con E. crenularis Lundell, que se distribuye en los estados de México, Jalisco, Guerrero y Naya- rit. Las principales diferencias entre ambas especies son las si- guientes: E. uxpanapensis E. crenularis margen de la entero crenulado hoja nervios late- 11-18 de cada lado 8-10 de cada lado rales peciolo 7-10 mm de largo 3-6 mm de largo eje de la in- densamente cobrizo- esparcidamente pu- florescencia tomentuloso bescente o glabro 406 pedicelo bracteolas hipanto disco estaminal fruto PRY. TO) LONG LA densamente cobrizo- tomentuloso densamente cobrizo- tomentulosas 1.5-2.5 mm de largo densamente cobrizo- tomentuloso cobrizo-pubescente globoso u oblato, 1-1.3 cm de largo, 1-1.5 cm de ancho Vol. 63, No. 5 escasamente pubes- cente o glabro ciliadas hasta de 1 mm de lar go, glabro o escasa- mente puberulento glabro elipsoide, 9-10 mm de largo, hasta 7 mm de diametro Se distribuye principalmente en selva alta perennifolia. Su flo racién ha sido observada de abril a agosto; su fructificaci6n de agosto a noviembre y algunos en enero. Referencia: McVaugh, R. 1963. Tropical American Myrtaceae II: Notes on generic concepts and descriptions of previously unrecognized Fieldiana, Bot. 29:436-37. species. STUDIES IN THE LIABEAE (ASTERACEAE). XVIII. A NEW SPECIES OF MUNNOZIA FROM BOLIVIA. Harold Robinson Department of Botany National Museum of Natural History Smithsonian Institution, Wasington, D.C., 20560. Collections by A. Fournet in Bolivia include two specimens of a previously undescribed species of Munnozia. The species is named here after the collector. MUNNOZIA FOURNETII H. Robinson, sp. nov. Plantae herbaceae perennes ca. 1 m altae (7) lacticiferae. Caulis hexagonales rufescentes dense breviter hirsuti et stipitate glanduliferi. Folia opposita base late perfoliata, petiolis ad 7 cm longis et in alis ad 2 cm latis; laminae late deltoideae ad 18 cm longae et 15 cm latae base abrupte truncato- hastatae fere ad eadem planam valde trinervatae Margine in partibus omniis iregulariter dentatae apice vix vel breviter acuminatae supra viridibus dense tuberculatae et evanescentiter albe arachnoideo- tomentosae subtus dense albo-tomentosae in nervis majoribus fulvo hirsutulae. Inflorescentiae terminales laxe opposito-ramosae; pedunculae 10-12 cm longae dense brunneo-hirsutae et stipitate glanduliferae. Capitula late campanulata; involucra ca. 1 cm alta et 1.5 cm lata; squamae involucri ca. 25 et ca. 3-seriatae subaequilongae vel in exterioribus longiores lanceolatae ca. 10-12 mm longae et 1-2 mm latae apice attenuatae exteriores extus dense hirsutae et stipitate glanduliferae. Flores radii 30-35; corollae flavae, tubis ca. 4 mm longis pilosulis inferne vix glanduliferae; limbis 11-12 mm longis et ca. 2 mm latis; flores disci ca. 60; corollae flavae ca. 1 cm longae, tubis ca. 6 mm longis pilosulis et sparse glandulo-punctatis, faucibus ca. 2 mm longis anguste campanulatis, lobis ca. 2 mm longis et 0.6 mm latis apice glandulo-punctatae et pauce piluliferis; filamenta in parte superiore ca. 0.2 mm longa; thecae ca. 1.9 mm longae; appendices antherarum ovato- oblongae ca. 0.3 mm longae et 0.17 mm latae; scapi stylorum in partibus superioribus hispidulis ca. 3 mm longi; rami stylorum ca. 1 mm longi. Achaenia ca 1.5 mm longa 10-costata dense setuliferae; setae pappi longiores ca. 25 plerumque ca. 5 mm longae apice non incrassatae, setae exteriored tenuiores ca. 0.7-1.0 mm longae. Grana pollinis in diametro 33-35 pm. 407 - 408 PHY TOL 0 Gy ivA Vol. 63, No. TYPE: BOLIVIA: Dept. La Paz: Prés de Unduavi, ancienne route de Chulimani, 1.5 km, alt. 2950 m Herbacee de 1 m de haut. Feuilles opposées, sessiles, hirsutes, de 20 cm de long, triangulares, dentelées, bords du limbe brunatre. Latex blanc. Fluers jaunes, petales nombreux, sepales vert fonce bords bruns. 24/4/1986. A. Fournet 623 (Holotype, US). PARATYPE: BOLIVIA: Dept. La Paz: Yungas, prés de Unduavi, 1 km sur vieille route, alt. 2950 m. Herbacée de plus de 1 m de haut avec latex blanc. Feuilles opposées, sessiles, triangulaires, de 25 cm de long, de 15 cm de large, dentelées, hirsutes, vert vif. Tiges hirsutes. Fleurs jaunes en panicules, avec pédoncule de 10-12 cm. 24/4/1986. A. Fournet 616 (US). The new species is similar to some others known from the Bolivian area, but it is most distinct inthe broadly winged petiole of the leaf and in the narrow and glanduliferous involucral bracts. Munnozia gigan- tea (Rusby) Rusby and M. glandulosa (0.Kuntze) Rusby, which are possibly one species, have narrowly winged petioles and broader involucral bracts bearing whitish tomentum on their outer surfaces. Munnozia trinervia Ruiz & Pavon of southeastern Peru differs from the new species by its more narrowly winged petioles and by the graduated lengths of its narrow involucral bracts. 1987 Robinson, New Monnozia from Bolivia 409 ss hed Ee FOS HEMMER TRATOR TERK PP RA REESE PRY Se We tes share ceases 3959732 RASHBAY KHERBRRR Paes Se Pt Ripe Custertiqn Oo PPE BESS Det Soe Fa SOS Remen: Meare Munnozia fournetit H. Robinson, Holotype. Photos by Victor E. Krantz, Staff Photographer, National Museum of Natural _ History. Inset showing enlarged head. New Combinations required for the Flora of Central Eastern United States Clyde F, Reed In addition to the new combinations published in 1982 (see Phytologia 50(7): 461-462) in the preparation of the Flora of Cen- tral Eastern United States (Maryland, Delaware, Virginia and West Virginia, including the District of Columbia) ,the following nomen- clatural changes need to be made, Orchidaceae 1, Galearis spectabilis forma albiflora (Ulke) Reed, comb, nov, Based on Orchis spectabilis forma albiflora Ulke, Castanea 3: 70. 1938, Type: Virginia. Gramineae 2, Phragmites australis var, berlanderi (Fourn,) Reed, comb, nov. Based on Phragmites berlanderi Fourn,, Bull, Soc. Bot. France, 243 178. LS ii7e 3. Schizachyrium scoparium var, polycladum (Scribn, & Ball) Reed, comb. nov. Based on Andropogon scoparius var. polycladus Seniba: & Ball, Bur. U.S. Div. Agrost., 24: 40. 1900. Polygonaceae 4, Bilderdykia cilinodes forma erecta (Peck) Reed, comb. nov. Based on Polygonum cilinode (var.) erectum Peck, N.Y. State Mus., Rept. 46: 129. 1893, 5. Persicaria caespitosa vars Longiseta, (DeBruyn) Reed, comb. nov. Based on Polygonum longisetum © DeBruyn 1n Miq., Pl. Jungh, 307. 1854. Ros aceae 6. Malus angustifolia var. spinosa (Rehd.) Reed, comb. nov. Based on Malus ioensis var. spinosa Rehd., Trees and Shrubs, 2: 231, 1913. 7. Malus coronaria var. lancifolia (Rehd.) Reed, comb. noc. Based ~on Malus lancifolia Rehd. in Sargent, Trees and Shrubs, 2: Let, ey 258. tall Lc. 22° 229, 1983, Leguminosae 8. Amphicarpaea bracteata var. comosa (L.) Reed, comb. nov. Based on Glycine comosa L., Sp. Pl. 2: 754. 1753. 9. Chamaecrista fasciculata var, depressa (Pollard) Reed, comb nov. Based on Cassia depressa Pollard, Bull. Torr. Bot. "Club, Ds DED. ‘ts 2o25) L895 10, Ghansecctske fasciculata var. macrosperma (Fern.) Reed, comb. nov. Based on Cassia fasciculata var. macrosperma Fern., Rhodora 42: 455. 1940. 410 1987 Reed, New combinations for Eastern U.S. 411 11. Chamaecrista nictitans var, hebecarpa (Fern.) Reed, comb.nov. Based on Cassia nictitans var. hebecarpa Fern., Rhodora 38: LDS eo Sole 12. Chamaecrista nictitans var. leiocarpa (Fern.) Reed, comb. nov. Based on Cassia nictitans var. leiocarpa Fern., Rhodora 38: [RON Fe, CVI en Gyn MAS EIA. 13. Senna hebecarpa var. longipila (E.L.Braun) Reed, comb. nov. Based on Cassia hebecarpa var. longipila E.L.Braun, Rhodora 42: 49. 1940. Oxalidaceae 14. Oxalis dillenii subsp. recurva (Ell.) Reed, comb. nov. Based on Oxalis recurva Ell., Sketch Bot. S.C. & Ga., 1: 526. 1821. Anacardiaceae 15. Toxicodendron toxicarum forma elobatum (Fern.) Reed, comb. nov. Based on Rhus toxicodendron forma elobatum Fern., Rhodora (No\e" esis a olsbyge eo weg. eVAIIC 16. Toxicodendron toxicarum forma leiocarpum (Fern.) Reed, comb. nov. Based on Rhus toxicodendron forma leiocarpa Fern., Rhodora 43: 599. 1941. Umbelliferae 17. Sium suave var. floridanum (Small) Reed, comb. nov. Based on Sium floridanum Small, Man. Southeast Fl., 976. 1933. Ericaceae 18. Rhododendron periclymenoides forma glandiferam(Porter) Reed, comb. nov. Based on Azalea nudiflora (£.) glandifera Porter, Bull. Torr. Bot. Club, 27: 508. 1900. Syn.: Rh. nudiflorum £. glandiferum (Porter) Fern., Rhodora 43: 619. 1941. 19. Rhododendron periclymenoides forma album (Ait.) Reed, comb. nov. Based on Azalea nudiflora (f.) alba Ait., Hort. Kew., 1: 202. 1789. Syn.: Rh. periclymenoides var. album Pursh, Fl. Amer. Sept., 1: 152. 1814; Rh. nudiflorum forma album (Ait.) Rehd. in Wilson & Rehd., Monogr. Azal. (in text). 1921. Primulaceae 20. Dodecatheon pulchellum forma margaritaceum (Fassett) Reed, comb. nov., Based on D. amethystinum forma margaritaceum Fassett, Amer. Midl. Nat., 31: 475. 1944. Wisc. 21. Dodecatheon pulchellum forma strictlerae (Fern.) Reed, comb. nov. Based on D. meadia forma strictlerae Fern,, Rhodora 39: 320. 1937. Syn.: D. amethystinum forma strictlerae (Fern. ) Fassett, Amer. Midl. Nat., 31: 476. 1944. Pa. Contolvulaceae 22. Calystegia spithamaea var. pubescens (A.Gray) Reed, comb. nov. Based on Calystegia sepium var. pubescens A.Gray, Man. Bot. U.S., Ed. 5, 376. 1867. Syn.: Convolvulus spithamaetis var. pubescens (A.Gray) Fern., Rhodora 51: 70. 1949._ 412 PPheye TOM, OnG? THA Vol. 63, No. 5 Labiatae 23. Clinopodium vulgare (subsp. vulgare) var. neogaeum (Fern.) Reed, comb. nov. Based on Satureja vulgaris var. neogaea Fern., Rhodora 46: 388. 1944. Rubiaceae 24. Houstonia canadensis var. setiscaphia (L.G.Carr) Reed, comb. nov. Based on Houstonia setiscaphia L.C.Carr, Rhodora 46: 307. 1944. (Lee Co., Virginia), Compositae 25.Ageratina aromatica var. incisa (A.Gray) Reed, comb. nov. Based on Eupatorium aromaticum var. incisum A.Gray, Syn. Fl. N. Amer., 1(2): 101. 1884. Reed Herbarium 10105 Harford Rd,, Baltimore, Maryland 21234 Inasmuch as we do no editing, papers accepted for publication must be submitted in exactly the form that the author wants to have them published. They will then be photographed and printed by photo-offset in exactly the form as submitted except that we will add page numbers and running-heads. Typescripts should be prepared single-spaced on clean white heavy bond smooth and opaque paper. Elite type is probably the most space-economical. Typescript text must not exceed a rectangle 5% inches wide (horizontal) by 8° inches high (vertical), not including the running-head and page number. The title of the paper should be typed in all uppercase (capital) letters with 2 blank lines above the title and one beneath; then the name of the author in ordinary upper- and lower-case letters, along with his address (if so desired); followed by 2 blank lines; then the first line of text. It is usually best to leave a blank line between paragraphs. All scientific plant and animal names and group names should be typed either in italic type (if available) or underscored. Any corrections in the text made by the author must be complete and neat as they will be photographed as they are. The finished typescript as submitted by the author will be reduced from the 85, x 55/e inch size as submitted to 6% x 4 inches by the printer. It is therefore advisable to place a centimeter or millimeter scale on all text figures and plates included. waa ' Use a new heavily inked black typewriter ribbon and be sure to clean the type on the typewriter after each several pages of typing. Cost of publication at present is $12.00 US per page, with no subsequent rebates, but this rate may vary depending on inflation and costs, so it is best to ! ; inquire as to current rates. The page charges are due with the typescript and no paper will be published before payment is received in full. Each author will receive | gratis a proportionate share of the printed copies remaining after paid subscrip- tions are filled, but if separates (reprints or offprints) are desired, these will be charged extra in accord with the current rate for offprints provided by the printer. The cost of all such separates ordered must also be paid for in advance at the time the typescript is sent. No orders for separates will be accepted later, nor can additions or corrections be accepted. é Authors are asked to indicate in light pencil on the reverse side of each page q of their typescript the page number so that no mistakes in sequence occur. All manuscripts accepted will be published in the next issue, so that the size of the numbers may vary greatly. A volume will contain 512 pages. The plan insures prompt publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain limitations of size and proportion. An extra charge will be made for halftones, depending on their size, as fixed by the engraver. Se eae ee eT Sh Kee ea Tea ake s Articles dealing with research in all lines of botany and plant ecology, in any ‘ * e 2 reasonable length, biographical sketches, and critical reviews and summaries of if PA: literature will be considered for publication.