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a ACOOUNT OF THE RESULTS GAINED FROM A RECONNAISSANCE, Ps
‘TRIP MADE IN THE SPRING AND EARLY SUMMER OF 1899. — = : ie

PREPARED UNDER THE DIRECTION OF THE ENTOMOLOGIST = =
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BULLETIN No. 21, New SERIES.

Us DEPARTMENT OF AGRICULTURE

DIVISION OF ENTOMOLOGY.

PRELIMINARY REPORT

ON THE

INSECT ENEMIES OF FORESTS IN. THE NORTHWEST.

| AN ACCOUNT OF THE RESULTS GAINED FROM A RECONNAISSANCE
TRIP MADE IN THE SPRING AND EARLY SUMMER OF 1899.

PREPARED UNDER THE DIRECTION OF THE ENTOMOLOGIST
BY

A. D. HOPKINS, Ph. D

5)

Vice-Director and En tomologist of the West Virginia
Agricultural Experiment Station.

WASHINGTON:
GOVERNMENT PRINTING OFFICE.
L339 9:

LETTER OF TRANSMITTAL.

U. 8. DEPARTMENT OF AGRICULTURE,
Division oF ENTOMOLOGY,
— Washington, D. C., October 18, 1899.

Sir: I have the honor to transmit herewith manuscript of a prelimi-
nary report upon forest conditions as relating to insect work in
portions of the States of Oregon, Washington, and Idaho, which has
been prepared by Dr. A. D. Hopkins, of the West Virginia Agricul-
tural Experiment. Station, and which is based upon observations made
by him under commission from this Division during April, May, and
June of the present year. Dr. Hopkins was chosen for this work on
account of his great knowledge of forest insects, a subject of which
he has made a special study for a number of years, his previous obser-
vations having been made largely in the State of West Virginia. The
writer considers himself fortunate in being able to secure Dr. Hopkins’s
services for this investigation, and feels sure that the results obtained,
even as displayed in this preliminary report, will be of value to the
forest interests of the Northwest. The publication of this report as
Bulletin No. 21, New Series, is recommended.

Respectfully, pl OL TowARD:
Entomologist.
Hon. James Wixson,
Secretary of Agriculture.

Go

PREFATORY NOTE.

The accompanying preliminary report of my investigations of insects
injurious to forests in California, Oregon, Washington, and Idaho is
made in accordance with the authorization of the Secretary of Agri-
culture, dated March 30, and the letter of the Entomologist, dated
Avorilialsle 1899:

Before proceeding further, I wish to express grateful acknowledg-
ments to all those who have in any way assisted me in this work, among
whom the following deserve special mention: Messrs. Pergande and
Busck, for the excellent care of the breeding material sent in from the
field; Mr. Hine, for the excellent and rapid manner in which the reared
and collected insects were mounted; Mr. Clifton, for assistance along
other lines, and especially Mr. Schwarz, who devoted much time out-
side of office hours in helping me to make the preliminary investigation
and classification of alcoholic material and the determination of the
Coleoptera.

In addition to the above I should mention my obligations to United
States Senator McBride for letters of MD EOUCL OE | to persons in Oregon
who gave me valuable information.

Among the entomologists and other members of the staff or faculty
of the several universities and experiment stations visited whose hos-
pitality and interest in furthering the objects of the journey deserve
special mention were Dr. Hilgard and Professors Woodward and
Davy, of the University and Experiment Station at Berkeley, Cal.;
Professors Cordley and Lake, of the Agricultural College and Experi-
ment Station at Corvallis, Oreg.; Professor Kincaid, of the State Uni-
versity of Washington; Professor Piper, of the Agricultural College
of Washington, and Professors Aldrich and Henderson, of the Univer-
sity of Idaho.

ADE Sre

CON EN Ts:

Ontlhimesoistnip,andelocalities visited . 1252. ..5. 52.22. I
SUMMIT Ole WOL Kaan deresulitges 20 ecns 25 soos. ie sae neee. eee
Information on the habits of Scolytids made the prime object -..........----
Principal Scolytid enemies of forests of the Northwest.................------
Destructive Buprestid enemies of Hemlock, red fir, and Noble fir.......-.---
HOTest-oneeKG C1OlIatONeme ee woes yh oe) See Se oe ee

General observations on forest trees and forest conditions........-.-..-------
General observations on some forest problems of the Northwest......-..-----
@onsideration: of preventives and. remedies. ..........2.....---+22 2222 22222:
Sumuiamyennderecommendatlonsss: 25. ..8. 552.22 lee. 8 ee ee

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PRELIMINARY REPORT ON THE INSECT ENEMIES OF
, BORESTS IN4AHE NORTHWEST.

OUTLINE OF TRIP, AND LOCALITIES VISITED.

I left Morgantown, W. Va., April 9, and Washington, D. C., April
12, and, going by the southern route through Texas, New Mexico,
Arizona, and California, arrived in San Francisco on the evening of
April 17. The following day (April 18) was spent in visiting the
University of California, at Berkeley, where I called on Professor
Woodworth, entomologist, and Prof. J. B. Davy, botanist, of the
Agricultural Experiment Station, who offered every facility and gave
valuable assistance in my study of the native forest trees represented
in the university arboretum. Here I collected numerous species of
Scolytids from the pines and other conifers.

On April 19 I proceeded to Guerneville, Cal., some 70 miles north
of San Francisco, to investigate the insect enemies of the California
redwood (Sequoia sempervirens), and, in addition to several other spe-
cies of insects collected from the bark and wood of this tree, I found
that Phlwosimus cristatus was a very common enemy.

On April 20 considerable time was spent in Golden Gate Park, San
Francisco, studying the indigenous trees of the Pacific coast repre-
sented there, and in collecting specimens of insect enemies of the park
trees. I also called on Mr. N. Hoffman, who is a dealer in imported
and native woods, and is well informed on the timber interests of the
Pacific coast, and obtained from him some valuable information.

I left San Francisco to pursue my investigations in the vicinity of
McCloud and Sisson, Cal., where I had been informed some timber
was dying, and arrived at the former place, a small village in the
heart of the yellow and sugar pine forests, on the southern slopes of
Mount Shasta, April 21. Among the more important observations
made here was the discovery of a bark-beetle of the genus Den-
droctonus killing the yellow pine, and another of the genus Scolytus
doing great damage to living fir trees. In addition to this a large
number of heretofore unrecorded facts were noted regarding the insect
enemies of the spruce, fir, cedar, pines, etc. The owners of the timber
in this locality expressed themselves as much interested and pleased
that these subjects were being investigated, and they kindly offered

(

.

every facility for prosecuting the work, furnishing livery, tools,
guides, ete.

April 22, I arrived at Grants Pass, Oreg., and investigated the con-
dition of the forests in relation to insect depredations in the moun-
tains north, northeast, and southwest of the city. The most important
feature found here was the prevalence of the same species of Dendro-
etonus discovered at McCloud, and many hundreds of trees which had
evidently died from its attack. In one place a group of over thirty
trees were dying. These were carefully investigated. The living and
dying bark were found to be infested with different stages of the
Dendroctonus, and quite conclusive evidence was found that the
death of these trees was primarily due to the attack of this bark-
beetle. In another section it was found that the same species had
attacked the sugar pine, Pinus lambertiana, which, with Pinus ponder-
osa, are among the most valuable timber trees of the Northwest. The
discovery was also made here that this Dendroctonus was not attracted
to felled trees, as is the case with most other Scolytids, but is attracted
to, and freely breeds in, trees that have been girdled by the settlers
and farmers in the process of clearing the land; and that these trees
form nuclei for the multiplication and spread of the pest to the healthy
trees in adjoining forests.

Leaving Grants Pass on the 27th, I spent a few hours profitably col-
lecting specimens near Albany and Corvallis, arriving at the latter
place in the evening. In accordance with instructions I visited the ~
agricultural college and experiment station here on the 29th, and
called on the entomologist, Prof. A. B. Cordley.

The next day, April 30, accompanied by Professor Cordley, I pro-
ceeded to Newport, Oreg., at the mouth of Yaquina Bay, where I
spent the day investigating the insect enemies of the trees of this most
interesting section. Among the important observations made here
was my discovery of a Scolytid enemy of the white alder (Alnus rhom-
bifolia), which is very common and quite injurious tothistree. Another
species of Scolytid, representing an undescribed Hylesinus, was found
to be common in the bark of recently felled hemlock. These observa-
tions had the added interest of being the first record of Scolytid bark-
beetles infesting alder and hemlock. An undescribed Dendroctonus
was found in the bark of Picea sitchensis, and several other Scolytids,
which are undescribed, were found in this spruce and in Pinus contorta.
I was also fortunate in finding. in a cone of the last-named tree Mr.
Schwarz’s rare and interesting species, Pityophthorus coniperda, the
occurrence of which on this coast is most remarkable. Taking it all in
all, this locality proved to be one of especial scientific and economic
interest. |

May 1, we returned to Corvallis, and the following day, accompanied
by Professor Lake, horticulturist and botanist of the college and sta-

g

tion, we proceeded into the Cascade Range, stopping at Detroit, near
the terminus of the railroad, where two days were spent in profitable
observations of the forest conditions and insect enemies of the cedars,
firs, spruces, pines, hemlock, maple, ete.

May 4, two guides were secured at Berry, Oreg., and we entered the
Cascade Forest Reserve. Here I made some important observations
regarding a trouble which had caused the death of large numbers of
noble firs (Adzes nobilis) and hemlock. While I was not able to
secure specimens of the insects which doubtless caused the death of
these trees (owing to the fact that the trees had been dead too long), I
concluded from the character of their work on the outer sapwood that
it must be a Buprestid, possibly belonging to the genus Melanophila.
Many had healed over before the trees died, thus presenting con-
clusive evidence that they were attacked while living and probably
while in healthy condition.

While on my way to Portland, May 5, I stopped at Salem and called
on Mr. E. C. Giltner, to whom I had a letter of introduction from
United States Senator McBride. Mr. Giltner was very kind in giving
information and in introducing me to persons who were familiar with
the forest conditions of certain sections of the State.

May 7, I proceeded to St. Helen, Oreg. Here investigation was made
of a serious trouble affecting the hemlock and red fir, which proved
to be similar to, if not identical with, that affecting the hemlock and
noble fir in the Cascade Reserve, near Berry. The work of a
Buprestid larva was found in healthy, dead, and dying trees, and I
was fortunate in securing a few larvee, also some nice specimens of
healed-over galleries. The latter presented conclusive evidence that
the attack had been made on healthy, growing trees. It seems that
the trees succumb only after several years of successive attack by this
insect. The extensive cutting of timber for fuel carried on here does
not seem to answer the usual purpose of attracting the insects away
from the living trees. Therefore, this is a problem of considerable
interest, and, under more favorable conditions for study, should receive
attention, in order to determine some method of preventing the rav-
ages, which may prove to be even more extensive than at present.
In the evening I called on Mr. W. H. Dolman, with a letter of
introduction from Senator McBride, and that gentleman, from his
experience and practical knowledge, was able to give me valuable
information regarding the forest conditions in that section of the State.

Leaving St. Helen May 8, I arrived at Astoria, Oreg., the next morn-
ing, and made observations and collections in this most interesting
locality.

May 9, I proceeded to Seaside, Oreg., where I learned of a serious
trouble affecting the timber some 14 miles back on the Coast Range in
the vicinity of Ahlers. I at once secured a guide and horses and pro-

10.

ceeded some 7 miles that night. The next morning we followed a trail
through the dense and magnificent forest here to Ahlers, where the
timber, as far as could be seen in all directions, was dead. The trees
had been dead eight or nine years; hence it was not possible to per-
sonally investigate the cause of the trouble. I was particularly fortu-
nate, however, in obtaining from the postmaster, Mr. Herman Ahlers,
a most interesting account of the beginning and ending of the trouble
as observed by him at the time. It seems that the trees were defoli-
ated, probably by a geometrid larva, in 1890-91, and that nearly all of
the hemlock and Tideland spruce on an area lying between 450 and
1,200 feet above sea level in the southern half of Clatsop and the
northern half of Tillamook counties died as a direct result of this
defoliation.

Returning to Seaside May 11, I did some collecting there from scrub
pine, Tideland spruce, and red fir, securing several species of insects
new to the collection. At Hillsboro, Oreg., I called on Congressman
Tongue, of the First district of Oregon, who heartily approved of the
work in which I was engaged and expressed the hope that good results
would follow.

May 12th proceeded to northern Washington to continue investiga-
tions in that region, stopping one day at Seattle, and arriving at Port
Angeles May 15. Here two days were spent in active work, and many
additional facts were determined and valuable material added to the
collection. In addition to many of the same set of enemies of the
common forest trees observed at other points visited, I found here a
lepidopterous larva, as yet undetermined, which was quite common,
feeding on the leaves of red fir.

Returning from Port Angeles, I stopped two days (May 17, 18) at
Port Williams, Wash., where a special investigation was made to
determine the cause of the death of a large number of white fir and
red fir on a small military reserve there. It was found that the
trees were being attacked by the same Scolytus as the one observed
near McCloud, Cal., and also by a new Hylesinus. The red fir was
infested with Dendroctonus similis, and it was quite evident that this
species was the cause of their death. Some of the trees were felled
for the purpose of making a thorough study of the insects inhabiting
them. Some important facts were learned, and a large series of speci-
mens of the insects and their work was collected.

Returning to Seattle on the 19th, I spent the next day with Professor
Kincaid, entomologist at the State University, and several hours were
employed in collecting on the extensive and partly forested university
campus.

Leaving Seattle May 20, I proceeded to eastern Washington and
northern Idaho, where nearly two weeks were spent in the vicinity of
Spokane and Buckeye, Wash., and Sand Point and Kootenai, Idaho,

11

making investigations. Here I found quite different conditions pre-
vailing from those observed in western Washington and Oregon in
the prevalence of quite a different set of forest trees, notably the
lodge-pole pine (%nus murrayana), the mountain white pine (Pinus
monticola), and the Western larch (Larix occidentalis), all of which
yielded a large amount of new material and furnished new problems
for study.

Among the more important observations made here was the common
occurrence of the same Dendroctonus enemy of the yellow pine
observed in northern California and southern and central Oregon.
Many trees were found in the vicinity of Buckeye, Wash., which were
dying or had died, evidently from the attack of this insect, since in
every tree, of a large number examined, abundant examples of the
insect and its work were found. Jalso found that a serious trouble
had prevailed among the white pine in the vicinity of Kootenai, where
a large amount of this most valuable timber had died within the past
eight or ten years. Upon investigation I found quite conclusive evi-
dence that the damage had been caused by another undescribed Den-
droctonus allied to D. frontalis, but much larger than this or the one
found killing the Western yellow pine.

Dendroctonus similis was found quite abundant in larch, excavating
galleries and depositing eggs in living bark in recently felled trees.
Its habits and the common occurrence of its work in standing dead
trees indicated that it was one of the principal enemies of this kind of
timber.

In addition to the above observations many others were made on the
enemies of the pines, spruces, cedars, larches, birches, poplars, willows,
etc., and a large series of valuable specimens was added to the collection.

After making quite a thorough investigation of the conditions found
in this section of Washington and Idaho, I proceeded, on June 3, to
Pullman, Wash., where I visited the Agricultural College and Experi-
ment Station and called on the entomologist, Professor Piper.

June 4 Professor Piper took me into the forest at the base of Cedar
Mountain, Idaho, where a few hours were spent in successful collection
of insect enemies of the spruce, fir, pine, and other trees in this region.
Many new observations were made here and much new material col-
lected.

June 5 was spent with Professor Aldrich, entomologist of the Idaho
University and Agricultural Experiment Station, who was naturally
much interested in the object of my trip, since it was partly through
his suggestion that this work was undertaken.

On June 6 I made another excursion into the forests of Cedar Moun-
tain in company with the botanist of the University of Idaho, Pro-
fessor Henderson, and three students from the botanical and entomo-

12

logical classes. Among the additional facts obtained on this excursion
was the discovery that the Dendroctonus enemy of the yellow pine
attacks trees defoliated by the pine butterfly (Veophasca menapia),
and that it is quite intimately associated with the destruction of a
large amount of timber only partly defoliated by the butterfly larva.
Owing to the earliness of the season, no observations could be made on
this insect which has been in previous years so enormously abundant
and destructive in eastern Washington and in Idaho and Montana.
Indeed, it was hardly necessary that I should do so, since the subject
has received considerable attention at the hands of Professor Aldrich.
Further information, however, on the relations of Scolytids to the
subsequent death of trees only partly defoliated by it is a subject of
considerable importance which I believe has not previously been taken
into consideration.

Leaving Moscow on the 7th of June and going via Pendleton, Oreg.,

and Shoshone, Idaho, I arrived at Hailey, Idaho, June 9, where some
successful collecting was made from red fir logs at a sawmill, and
some valuable information was obtained from Messrs. Watt and
Peacock regarding the forest conditions and timber interests of that
section of Idaho. Learning of nothing else of especial importance
requiring my attention, I started on my return journey to Washington,
D. C., where I arrived on the morning of June 17, 1899.
- The total distance traveled from Washington, D. C., and return by
rail, boat, wagon, and on foot was about 9,000 miles; the time occu-
pied, sixty-six days; the number of localities visited, twenty-six.
The number of specimens collected is as follows:

Scolytidesmounted {(oumted))) ees sya o ee ee pean reece 1, 074
Scolkytidesimalcohol: (combed) sees aS see te ee 2, 003
Miscellaneous, principally Coleoptera, in aleohol (counted) -.-.-----.-------- 286
Miscellaneous, mounted and in alcohol (estimated) -......-.-.----.--/------ 300
Specimens of work of insects (counted)o 2 2 5-2 ee ee 650

Totals vosso3 oe. os be sk i See ee ale lee ee 4, 363
Separate motes im notebook. 2.22 asco Soe ae are 760

While the number of specimens is not as large as might have been
secured by general collecting, the manner in which nearly all were
collected from their host plants, and the large series of notes on their
habits, makes them all the more valuable.

SUMMARY OF WORK AND RESULTS. ~

The above outline of the trip will indicate the character and amount
of work done and the success attained in efforts to carry out instruc-
tions.

The distance traveled, the number of localities visited, and the lim-
ited time occupied in the trip from Washington and return left but

13

little time for actual work in the field. Realizing that this would be
the case, I devoted my attention to the investigation of a few leading
features which promised to yield the best results.

INFORMATION ON THE HABITS OF SCOLYTIDS MADE THE PRIME OBJECT.

Previous to this trip little had been published regarding the host
plants and habits of the Scolytide of the Northwest, and scarcely any
observations had been made on the relation of this destructive class of
insects to the unhealthy conditions of the timber. I felt, therefore,
that with my previous experience in the study of these insects in the
forests of West Virginia, it was my first duty to obtain as much infor-
mation as possible regarding the habits of those found in the localities
visited, and I spared no pains or labor to accomplish this end. As a
result of this effort specimens of some sixty species of Scolytids,' rep-
resenting some twenty genera, were secured. This includes nearly all
of the species heretofore known from northern California, Oregon,
Washington, and Idaho. It also includes a large number Heretorore
unknown from this region, with representatives of two or three new
genera. ‘The host plants and some features of the habits and life his-
tory of every species were determined. A large series of biological
material of both the undescribed and described species was also col-
lected.

PRINCIPAL SCOLYTID ENEMIES OF THE FORESTS OF THE NORTHWEST.

The genus Dendroctonus.—TVhis genus, which has been found to con-
tain some of the most destructive enemies of the conifers of the East,
was found to be represented in the Northwest by three described and
two or three undescribed species. They were found here, as in the
Kast, to be the principal enemies of pine, spruce, and larch; and from
what was observed of their habits it is plain that under favorable con-
ditions most, if not all, of them are capable of attacking and killing
healthy timber.

One species which I have provisionally identified as Dendroctonus
brevicomis Lec.” was found to be a most destructive enemy of the

?This is the number of the species determined by Mr. Schwarz and myself in a
preliminary study of the entomological material. Subsequent identification of the
mounted specimens may show that there are a few more or a few less species and
genera.

? Heretofore this species had not been recognized since it was first described by
LeConte. Dr. Dietz, in his notes on species of Dendroctonus of boreal America, states
that he does not recognize any difference in typical specimens of D. frontalis Zimm.°
and D. brevicomis, Lec., and consequently concludes that they are identical. The
examples I found are undoubtedly distinct from D. frontalis, although very closely
allied to this species in specific characters and habits. It agrees so closely with
Le Conte’s description of D. brevicomis, however, that I think it must we recognized
as such until a comparison With the type proves the contrary.

14

yellow pine (Pinus ponderosa) in northern California, southern and
eastern Oregon, northeastern Washington, and western Idaho. A large
amount of some of the finest timber in all of these localities had died
within the past seven or eight years, evidently as a direct result of
attacks by this bark beetle. It was also found to attack and prevent
the recovery of trees injured by defoliating insects and other causes.
Its habits and the character of its galleries appear to be identical with
those of Dendroctonus frontalis, which is noted for its destruction of
vast quantities of pine and spruce timber in West Virginia and adjoin-
ing States between 1890 and 1893. It is killing the Western yellow
pine, just as D. frontal’s commenced to kill the Eastern yellow pine
(Pinus echinata) before it spread to all the other pines and spruce.
Therefore, just as ). frontalis has proven to be the most destructive
enemy of Eastern conifers, the Western representative of this species
will doubtless prove to be, under similarly favorable conditions, equally
as destructive to the Western forests in which the conifers predominate.

Among the most important features observed regarding the habits of
this beetle was the fact that it is attracted to trees girdled by settlers
and farmers in the process of clearing land, and that in the bark of
such trees it breeds and multiplies in sufficient numbers to enable it to
attack and kill the timber in adjoining healthy forests. Indeed, my
observation leads me to conclude that a considerable number of girdled
pine trees may easily form a nucleus for a destructive invasion by it.

A Dendroctonus allied to the one just mentioned, but evidently unde-
scribed, was found to be a special and dangerous enemy of the sugar
pine (Pinus lambertiana) and the mountain white pine (Pinus monti-
cola), especially of the latter. It was frequently met with in the
vicinity of Grants Pass, Oreg., in sugar pine, and was found abundant
in the bark of dying and dead standing and felled white-pine trees in
the vicinity of Sand Point and Kootenai, Idaho, where a large amount
of timber had died, evidently as a result of its attack. This is the
same beetle as the one sent to the Division of Entomology by Mr.
Ayers from Columbia Falls, Mont., and subsequently sent to me, with
other Dendroctonus material, for study. It is undoubtedly capable of
attacking and killing great quantities of white and sugar pine, but may
possibly be prevented from doing so in the future, in all regions where
extensive timber cutting is carried on, by its being attracted to the
stumps, logs, and tops of trees felled for lumber and fuel.

Another species, at present recognized as Dendroctonus terebrans,
was commonly met with in the bark of living, dying, and dead _ stand-
ing trees and the stumps of recently felled Pinus ponderosa, P. lam-
bertiana, P. monticola, P. murrayana, P. contorta, and P. radiata, in
all of the localities where these species of pine grow. It was found
to be quite a common enemy of the Monterey pine on the university
campus at Berkeley, Cal., and I was informed by Professor Davy that

15

the unhealthy condition: of this pine in its native home (Monterey
County, Cal.) was probably due to the attack of this or of a closely
allied bark beetle. The fact that it was not found in spruce, fir, or
larch indicates that in the Northwest, as in the East, this large repre-
sentative of the genus almost exclusively infests the pine. Its habit of
attacking trees at the base is the same as in cae East, as are also the
social habits of the larvee.

The information I was able to gather of the habits and different
stages of the Western representatives of D.terebrans, of which nothing
had been recorded, will aid greatly in arriving at definite conclusions
regarding the identity of numerous doubtful forms collected on this
trip, and those among the large series of material in the national col-
lection.

Another Dendroctonus, referred to in my notes as ** Dendroctonus sp.,
near rufipennis,’ proved upon comparison with identified specimens in
my collection to be the same as examples labeled 2. semzlis.* It appears
that nothing has heretofore been recorded regarding the host trees or
habits of this insect, yet I found it to be one of the commonest bark
beetles of the region traversed, and a special enemy of the red fir
(Pseudotsuga taxifolia). It was especially abundant wherever this
tree was being cut for fuel or lumber. Partially developed broods
were in the bark of trees felled the previous year, and some had emerged
and were entering the bark of logs and stumps of recently felled ones,
in the living bark of which they were excavating galleries and depos-
iting eggs. Ona small military reservation at Port Williams, Wash.,
J found it associated with the death of large numbers of red fir,
and in eastern Washington and northern Idaho it was found to be quite
as common an enemy of the Western larch (Larix occidentalis) as of
the red fir. The fact that it was found so common in these two kinds
of trees, and not in Picea or Abies, suggests that there is possibly a
closer affinity between Pseudotsuga and Larix than has heretofore
been recognized by botanists.

An undescribed Dendroctonus, near 1). rujipennis, was found in the
bark of Tideland spruce near Newport, Oreg. This beetle is repre-
sented in the national collection by a few large, black examples from
Queen Charlotte Island, and in my collection by a large series recently
sent to me for identification from the same island by the Reverend
Keen. This is very likely a special enemy of the Tideland spruce,
and, judging from the habits of other species of the genus, will doubt-
less prove at times quite injurious or destructive.

The genus Scolytus.—This genus, which is represented in the East by
three common species which depredate on deciduous trees (viz, Scolytus
rugulosus on fruit trees, S. guadrispinosus on hickory, and S. mutzcus

1 This may, after all, prove to be D. obesus Mann., which LeConte believed to be
synonymous with D. rujipennis. (Rhynchophora of North America, 1876, p. 389.)

16_

on hackberry), is represented in the West by several species which
infest conifers. The genus is of especial interest on account of the
destructive powers of some of its representatives, which are capable
of attacking and living in the bark of healthy trees. One or more
unrecognized species allied to S. preceps were found to be primarily
destructive to the wood of living white fir trees in all of the localities
visited. The beetles attack the bark of healthy, vigorous trees and
excavate their primary, or egg-galleries transversely through the inner
bark and through the outer surface of the wood. When the attack is
not sufficient to kill the trees, these wounds heal over, but in the mean-
time a decay often sets in at these injured places, which extends through
the heart wood and for several feet above and below the wound, thus
rendering the wood worthless for lumber, and often for fuel. That
these trees were in a vigorous, growing condition when attacked was
proven by the fact that in many instances a normal growth of wood
and twigs had formed the year the attack was made, and when the
injury was not sufficient to weaken the vitality of the tree a normal
growth was made the next and succeeding years until the wounds were
healed. At Port Williams, Wash., many trees were dying, all of which
were infested with larve and pup of one or more of these Scolytus
enemies, and it was quite evident that they had been the primary cause
of the trouble. ne

Another species, at present recognized as S. wiispinosus, was found
to be a very common enemy of red fir along the coast, and of the red fir
and Western larch in Idaho and eastern Washington. No evidence was
found, however, that this species had caused the death of any of the
trees.

The genus Tomicus.—This genus is represented in the collection by
some ten species, four of which appear to be new. While I found
some evidence to indicate that a species recognized as Zomicus pine
was the primary cause of the death of numerous examples of the lodge-
pole pine in the vicinity of Spokane and of yellow pine near Moscow,
Idaho, the other species appeared to occur only as secondary enemies
or as allies of the more aggressive Dendroctonus.

Other Scolytids.—Among the other numerous species of Scolytids
collected, none were recognized as specially destructive enemies of
trees, except LeConte’s Hylesinus aspericollis, which is nota Hylesinus,
but appears to represent an undescribed genus, characterized by a
seven-jointed antennalfunicle. This I found infesting the living bark of
white alder (Alnus rhombifolia), with evidence that it not only hastens
the death of injured trees, but that it may be the primary cause of the
death of healthy ones.

Examples of larve and pupe, with a few dead adults, of an unde-
scribed Hylesinus were found quite abundant in the bark of a felled
hemlock at Newport, Oreg. This and the one found in alder are of

if,

special interest from the fact that there appears to be no previous
record of a Scolytid bark-beetle infesting either Alnus or Tsuga.

Another Scolytid, which does not seem to be referable to any known
genus, was found in the thin bark of young dying sugar pine trees in
the vicinity of Grants Pass, Oreg.; also in other pines and in the
red fir.

The numerous other Scolytids represented in the collection made on
this trip can not be mentioned in this preliminary report, but will form
the subject of several pages of original matter in my final report on the
results of this investigation.

DESTRUCTIVE BUPRESTID ENEMIES OF HEMLOCK, RED FIR, AND
NOBLE FIR.

The evidence found in living, dying, and dead noble fir and hemlock
in the Cascade Forest Reserve, at Berry, Oreg., and in hemlock and
red fir along the Columbia River, in the vicinity of St. Helen, indicated
that much loss of valuable timber had been due to the work of
Buprestid larve in the living bark of these trees. The conditions
at the time were not favorable for a study of this class of enemies or
for a special investigation of the damage which seemed to be due to
their attacks. It is, however, a subject of special importance, which
will demand considerable attention in the future. Specimens of larve
collected from the bark of living trees were provisionally identified
by Mr. Schwarz as belonging to the genus Melonophila, probably J/
drummondi, which is closely allied to JL fulvoguttata, a destructive
enemy of the tie.nlock in West Virginia. An undetermined Ceram-
bycid bark »orer was found associated with the Buprestid larve in
hemlock a.d red fir, but the evidence pointed to the Buprestid as
having ~nade the first attack.

FOREST TREE DEFOLIATORS.

While I did not have an opportunity, owing to the earliness of the
season, to investigate the insects which are injurious and destructive
to the foliage of forest trees, I found abundant evidence of this class
of depredators, especially in the work of the pine butterfly on the
yellow pine of eastern Washington and western Idaho, and the
Geometrid of Clatsop County, Oreg., on Tideland spruce and hemlock.

The work of the pine butterfly has been previously recorded by
Mr. Stretch, in Papillio (Vol. Il, pp. 103-110), and subsequently
copied in the Fifth Report of the United States Entomological Com-
mission (pp. 762-767). The subject has also received special attention
within recent years at the hands of Professor Aldrich, of the Idaho
station; therefore the only observation made by the writer on this trip
which it seems proper to mention in this connection is that relating to

8602—No. 21 2

13

Dendroctonus, Tomicus, and other Scolytids attacking the defoliated
and partly defoliated trees. My observations led me to conclude that
were it not for the secondary attack of the Scolytids many of these
trees would have recovered from the injuries by the butterfly larvee.
Indeed, it seems to me that this association of bark beetles with defo-
liating caterpillars in causing the death of trees isan important feature
to be taken into consideration in any study of troubles of this character.

The work of the supposed geometrid larva, found near Ahlers,
Oreg., has not, I believe, been previously recorded, and, while I did
not have an opportunity to see specimens of this insect, which has not
attracted attention since the summer and fall of 1890, I obtained some
valuable information from Mr. Ahlers regarding the character of un
invasion, the insect, and the extent of its depredations.

Mr. Ahlers, postmaster at Ahlers, took a course in entomology at
Wernigerode am Harz, Province of Saxony. Hence, his observations
are of especial interest. Mr. Ahlers said:

The worms commenced to attract a little attention here in 1889. A few moths
were seen in the fall of the same year. In July, 1890, the worms appeared in great
numbers, the first on the hemlock, feeding on the base of the leaves and cutting
them off. When standing beneath the trees the droppings from the insects and the
falling leaves sounded like rain. When all of the leaves were eaten from the trees
the worms would let themselves down by means of webs, and, if they were not full
grown, would feed on the leaves of all kinds of shrubs and trees, except the Douglas
spruce and cedar. The worms were observed during July and August, and disap-
peared in the latter month, probably going into the ground to pupate. In October
the moths began to come out. They were grayish white with dark markings on the
wings. The wings spread about 13 inches. The moths would appear on the wing
about 3 o’clock in the afternoon in enormous swarms around the tops of the trees,
resembling a white cloud. They continued to fly for about three weeks, at the end
of which time the ground was covered with the dead insects, and the small streams
were filled with them, in some places forming dams. The next year, 1891, they
attacked the Sitka spruce and defoliated the trees, which died the following fall or
winter. * * * When the worms were coming down from the trees the webs
made the trees look as if they were covered with a grayish veil. The worms were
about 13 inches long, the sides grayish green, with darker gray zigzag markings
on the back. When traveling they measured their way. This trouble extended
over parts of Clatsop and Tillamook counties, killing all the hemlock and Sitka

spruce in a belt between an elevation of about 450 and 1,200 feet above tide. The
Douglas spruce and red cedar were not injured.

At the time of my visit much of the area covered by dead timber
had been frequently burned over by forest fires, so that in some places
nearly all of the dead timber had disappeared, but in other places,
where the conditions had not been so favorable for the spread of the
fire, the dead, barkless trunks of the trees were still standing, the tops
broken and the branches fallen, the scene presenting at once a most
impressive example of the destructive powers of an insect enemy of
forests and of subsequent devastation by forest fires.

When it is considered that some of the spruce which grow here are
10 to 13 feet in diameter, 300 feet tall, and will yield more lumber

19

than many an acre of timber in the Eastern States, one can have some
conception of the vast amount of timber which had been destroyed.
While the caterpillars which were primarily to blame for this destruc-
tion have not attracted attention within recent years, they will doubtless
make their appearance again in destructive numbers and attack the
remaining timber, which is becoming each year more accessible to
lumbering operations, and hence rapidly increasing in value.

BENEFICIAL INSECTS AND DISEASES.

A large series of predaceous and parasitic insect enemies of the
principal destructive species was collected, and many features regard-
ing their occurrence, distribution, and habits noted. The predaceous
enemies of Scolytids were quite common in nearly all of the localities
visited, but the Chalcidid, Braconid, and Ichneumonid parasites of
Scolytids and other bark and wood-infesting insects were rare as com-
pared with the same class of insects in the East.

Numerous examples of diseases of both injurious and_ beneficial
insects were found, and many insects had died from their effects. It
was not possible, however, with the limited time at my disposal to
determine the exact relation of these factors to the injuries caused by
the principal enemies.

GENERAL OBSERVATIONS ON FOREST TREES. AND FOREST CONDITIONS.

Forest trees recognized and studied.—Some 22 species of conifers and
14 of deciduous trees were recognized on this trip and their insect
enemies studied, the following of which deserve special mention on
account of their common occurrence and commercial importance.

COMMON NAMES. BOTANICAL NAMES.
ediwOode seen S22 2e* ee ag et Bao fa te Sequoia sempervirens.
Niccrethayvellow=apine-: = 52 e5 228 ll Sl lek eels. e ls e Pinus ponderosa.
SIUZEE | UIE eee pp eel rr phe Aueye SONNET ch Oe Pinus lambertiana.
Silver pine (Western white pine) - ----- A es ehh aig olan gee Pinus monticola.
ves ebomelas comuce) 220.00) o.oo eee te eee Pseudotsuga taxifolia.
fivele lame sa iticemaeerer ea oes oo ba 6 Sele SO oes Picea sitchensis.

Teel GEG GE 225 Sa AS eee ee eer Thuja plicata.

BMC ctCEMe bend OQCKMe eos 2 | Nas Moo. os Se ee ee ee et ee Tsuga heterophylla.
anlcticus inition a. 87 809s ol ees oe a Abies grandis.
Ion remiss (EeMlakOh” eo oka 35 joss ort eee Se ees Abies nobilis.

VP SSSIET, GWE, oA RR ee es ee eT epee ee Larix occidentalis.

Each of the timber trees named in this list was found to have one or
more insect enemies. My observations on these trees and the relation
of their enemies to the general condition of the timber may be briefly
summarized as follows:

The redwood.—This appears to be an exceedingly hardy tree, and so
far as I observed in the short time spent in the forests of this species in
the vicinity of Guerneville, Cal., is in a healthy condition. It has,

20

however, in /’lwosinus cristatus an enemy which, under especially
favorable conditions, might prove quite destructive to standing timber.
At present this insect is attracted to the bark of felled trees in which
it breeds in great numbers. A Cerambycid bark borer, Callidium
janthinum, was also found under the bark of a felled tree, and may
be a common enemy.

The Western yellow pine.—This is widely distributed through north-
ern California, Oregon, Washington, Idaho, and many other places
wherever the soil and climatic conditions are favorable to its growth.
Indeed, it is one of the commonest and most valuable timber trees of
the region. From the evidence found it has probably a greater num-
ber of insect enemies than any of the other trees mentioned, and at
present is suffering far more from their ravages. It has in Dendroc-
tonus brevicomis a most pernicious enemy, which penetrates, and
excavates winding galleries through, the living bark of the finest
trees, thus speedily causing their death. Very many trees have died
and are dying from this cause, and the dead ones are contributing to
the spread of forest fires. Its next greatest enemy is the pine but-
terfly, which has, from time to time, defoliated and caused the death
of much of the best yellow-pine timber in eastern Washington and in
Idaho.

There are many secondary enemies of greater or lesser importance
among the Scolytid genera Pityophthorus, Pityogenes, Xyloterus,
Tomicus, Hylastes, and Hylurgops, which contribute to the death of
trees primarily injured by defoliating and other insects, fire, and other
causes. Numerous Buprestid and Cerambycid enemies of the wood
and bark contribute to the unhealthy condition of the timber and the
destruction of the wood. A lepidopterous larva infests the terminal
twig of young trees near Moscow, Idaho, and one or more Curculionid
beetles breed in the bark at the base of young and old trees.

The sugar pine.—W hile this, one of the most valuable timber trees
of California and southern Oregon, is in a more healthy condition
than the yellow pine, it has a dangerous enemy in an undescribed
Dendroctonus as well as in the destructive enemy of the latter. It
has also a host of other enemies which infest the twigs, bark, wood,
and roots. i

The silver, or western white pime.—This is a very common and most
valuable tree in eastern Washington and in Idaho, and has a special
enemy in the same undescribed Dendroctonus that infests the sugar
pine. A vast amount of dead timber was found in groups and scat-
tered through the forests in the vicinity of Kootenai and Sandpoint,
Idaho, which showed good evidence, in the great abundance of gal-
leries of this insect in the bark, that they had been killed by it. A
Tomicus, closely allied to confusus, was also found to be a very com-
mon enemy, entering the living bark of recently felled trees; as was

21

also a Monohammus larva, which was found mining in, and destructive
to, the wood of standing and felled trees.

The red fir.—This common and hardy timber tree of the North-
west, is of especial value both for its lumber and fuel product.
While it is generally in a healthy condition, many dead and dying
trees were found in all the sections visited, and in some places
sufficient evidence was found to show that it had dangerous ene-
mies in certain Buprestid and Cerambycid bark borers, which are
capable of living in the healthy bark of growing trees. It has also a
special enemy in Dendroctonus similis, which is everywhere common
in the bark of dead and dying standing trees and in the dead and living
bark of logs and stumps of felled ones. The evidence found in the
dying trees on the military reservation at Port Williams indicates that
this bark beetle may, under favorable conditions, be the primary cause
of the death of the trees. Scolytus uwnispinosus is also a very common
enemy, infesting the bark on the main stem of small trees, and on the
branches of larger ones, which have been previously injured or felled.
A lepidopterous larva was found feeding on the foliage near Port
Angeles, and its common occurrence and habits of feeding indicate that
it might easily become quite destructive to the foliage.

The Tideand spruce.—This is a magnificent tree, found in great
forests along the coast in Oregon and Washington. It is attacked by
several pernicious enemies, the principal of which is the undetermined
Geometrid, mentioned on another page, which has destroyed immense
quantities of timber and will doubtless be a menace to timber interests
here in the future. The undescribed Dendroctonus found in the bark
and trees of logs at Newport, Oreg., is a special enemy, as are also
Dryocetes affaber, Hylurgops rugipennis, Xyloterus bivittatus, and
some other bark beetles which were found mining in the partly living
bark or wood near Ahlers, Oreg.

Red cedar.—This is a common tree throughout Oregon and Wash-
ington, and is of especial value for shingles, telegraph poles, and
lumber. Its principal enemies: are one or more species of Phlceo-
sinus and the Cerambycid, Callidium janthinum, which infests the
living bark of recently felled trees and those injured by fire or other
causes. The heartwood of living trees is also seriously injured by
ants, which extend the damage started by Cerambycid heartwood
borers, and a fungous disease. Numerous dead trees were observed in
all sections of the forest, many of which had evidently died from
injuries by one or more of these enemies.

The hemlock.—This is also a common hardy tree. It is attacked by
the same destructive bark borers as those mentioned under red fir as
well as by the Geometrid mentioned under Tideland spruce. In addi-
tion to these it has an undescribed Scolytid enemy which infests the
bark. Many dead trees were found in the Cascade Reserve and both

22.

dying and dead ones in the forests near St. Helen, all of which showed
evidence of having been killed or injured by a Buprestid larva, while
the great numbers of dead hemlock in Clatsop County, Oreg., were
sufficient evidence of the destructive powers of the Geometrid.

The white or grand jir.—This is a very common tree, which would
be of special value for the manufacture of pulp but for serious injuries
by one or more species of Scolytus and Hylesinus, which infest the
living trees, causing a decayed condition of the wood of some and
the death of many others, so that the fir is often in an unhealthy
condition and the dead and decaying trees contribute to destructive
forest fires. It also has a number of other Scolytid, Buprestid, and
Cerambycid enemies which infest the wood and bark. The same
caterpillar that was found attacking the foliage of the red fir was also
found feeding on the leaves of this fir.

The noble jir.—This is, as its name indicates, a most noble tree,
found in its perfection at moderate altitudes. It has a serious enemy
in the Buprestid, similar to, if not identical with, that which attacks
the red fir and hemlock. While I did not have an opportunity,
owing to deep snow, to make extended observations regarding the
condition of this timber, it would appear from what was seen in the
Cascade Reserve that many of the finest trees have died and that an
unhealthy condition prevails.

The Western larch.—TYhis larch, unlike its Eastern representative,
grows chiefly in dry sandy or gravelly soils and attains a diameter of 3
or 4 feet. It appears to me to be a very hardy tree, often retaining
its vitality for months after it is felled. It has a common enemy in
Dendroctonus similis, which, as previously stated, is capable of doing
considerable harm to this tree and the red fir. Many dead trees
were found showing evidence of having been killed by it, and the
beetles were common, entering the living bark of recently felled trees
in the vicinity of Kootenai, Idaho. A Buprestid bark borer, probably
Melanophila, was found in all stages of the bark in felled trees, and
Scolytus unispinosus was common in the inner bark of the tops and
branches of trees felled in the summer of 1898. The number of dead.
trees observed and the common occurrence of dead branches on living
ones indicated that considerable damage had been done by its insect
enemies.

GENERAL OBSERVATIONS ON SOME FOREST PROBLEMS OF THE
NORTHWEST.

My interest in forest subjects in general and previous studies of
the forest conditions in West Virginia led me to make some general
observations on the forest problems of the Northwest which are in
any way related to insect depredations, such as the influence of farm-

-™ rr

23

ing and lumbering methods, mining operations, forest fires, and
diseases of trees.

Influence of farming methods.—1 found that, in saddition to the
girdled timber contributing to the aml slentien of dangerous insect
enemies of forests, the clearings made in the midst of primitive for-
ests by the settlers and farmers expose the trees on the borders of
such clearings to abnormal influences, which weaken the vitality of the
matured timber. ‘The destructive insects and their allies, breeding in
the girdled trees, emerge and attack these weakened ones, and with
this additional material, and most favorable conditions for their multi-
plication and spread into the healthy forest, a trouble is often started
which results in the destruction of vast quantities of timber.

Influence of lumbering methods.—The abundance of timber in this
ereat forested area and the cost of transporting the products to market
leads in most cases to wasteful lumbering methods, only the best timber
suitable for special purposes being taken. The remainder, which in a
short time would be just as valuable for other purposes, is left with
scarred trunks, broken tops and branches, and deprived of the pro-
tective environments under which it had developed. The ground is
strewn with tops, branches, and trunks of the felled timber, which,
with the injured standing trees, furnish the most attractive breeding
places for all bark and wood infesting insects. If the cutting is con-
tinued from year to year in the same locality the insects, upon emere-
ing from the débris of the old cuttings, migrate to that of the new, and
do not invade the healthy or undisturbed forests. If, on the other
hand, cutting is discontinued and the débris is not destroyed by fire,
there is great danger of the insects, upon emerging, attacking the
healthy timber in the surrounding forests and continuing their rav-
ages there.

The relation of forest ve ‘es to depredations by insects.—This is a sub-
ject to which little attention has heretofore been given. My observa-
tions in the fire-swept areas in the Northwest has, in addition to
previous observations in West Virginia, convinced me that this is a
problem of considerable importance. Trees dying from injury by
fires furnish favorable breeding places for vast numbers of destructive
and other bark and wood infesting insects. Those slightly injured
and weakened in vitality are prevented from recovery by the attack
of bark beetles, which are ever ready to take advantage of any oppor-
tunity to overcome the resisting forces of their favorite host plants.
Thus the most favorable conditions are offered for the multiplication
of insect depredators, which, through their augmented power in
increased numbers, are capable of extending their ravages into the
healthy forests. Fire wounds at the bases of living trees also lead to
serious trouble and losses of the most valuable timber. These wounds
give entrance to Cerambycid and Buprestid heartwood borers, black

Jae

ants, and wood-decaying fungi, all of which extend from year to year
_ their obscure yet destructive work. Nor does the trouble end here.
The dead trees and decaying wood of the living ones, as indicated on
previous pages, offer most favorable conditions for the starting and
spread of forest fires. It is therefore plain that in considering methods
of preventing depredations by insects those relating to the prevention
of forest fires are, under certain conditions, of special importance.

In the nonreserved forests, where extensive timber operations are
carried on, the prevention of forest fires as a means of controlling
insect depredations is far less important than in the reserves. In fact
the débris in cut-over areas serves as attractive traps for the insects,
where they may be effectually destroyed, or their migration to healthy
timber prevented, by the almost inevitable conflagrations which follow
these cuttings. Therefore under such conditions it is not so much the
problem of prevention of fires as that of their control which should
receive attention, since if this débris can be burned at a time when the
conditions are least favorable for the fire to spread into the standing
timber the desired good, in the destruction of insects, would be attained
and the evil effects of the fires averted. On the other hand, every pos-
sible effort should be made to prevent outbreaks of fires In virgin and
reserved forested areas, not only to prevent the vast destruction by
fire alone, but as a precaution against destructive ravages by insects.

Relation of insect enemies of trees to forest jires.—This is a subject on
which nothing appears to have been published, yet it requires only a
little observation in the Western forest to make it clear that the trees
which have been killed by insects furnish, in their fallen branches,
standing and fallen partly decayed trunks, and dry bark, a most favor-
able condition for the starting, rapid spread, and perpetuation of forest
fires.

The relation of diseases of trees to insect enemies of forests.—lt is
well known that forest trees weakened by disease contribute to the
multiplication of their insect enemies. It is also known that insects
will attack healthy trees, and that diseases of the bark and wood fol-
low asa result of such injuries. Therefore, in the investigations of
unhealthy ‘conditions of forests it is often exceedingly difficult, with-
out some previous knowledge of the habits of the diseases and insects
found associated with them, to decide which is to blame for the pri-
mary injury. Our present knowledge of the subject, however, indi-
cates that as a rule unhealthy forest trees, like unhealthy animals,
present characteristic symptoms, which indicate quite clearly the pri-
mary cause of the trouble. The evidence I have been able to gather in
the forests of the East and Northwest makes it plain to me that, of the
two causes, while many small trees are killed by root diseases, the
unhealthy condition of the larger trees is more often due to primary

PD,

attacks by insects. Indeed, it appears that insects contribute more to

25

the spread of fungous diseases of the bark and wood of the main trunk
than do such diseases to the spread and ravages of insects.

The most striking example of insects contributing to the spread of
fungous diseases was observed in the white fir throughout the region
traversed; the heartwood of which is commonly rendered worthless
by decay as the result of wounds in the living bark made by Scolytus
bark beetles.

Interrelations of forest fires, insects, and fJungous diseases.—It will be
seen from the foregoing references to the relation of forest fires to
insect ravages, insects to forest fires, diseases of trees to insects, and
insects to fungous diseases, that there is a close interrelation, and, to a
certain extent, interdependence of all of these factors in the destruc-
tion of valuable forest products. While I had made some observations
along this line in the forests of West Virginia, I had little conception
of the magnitude and importance of the problem previous to my trip
through the forests of the Northwest, where quite conclusive evidence
was found that had it not been for previous depredations by insects
and consequent rapid decay of the standing and felled timber, far less
destruction by forest fires, or need for trouble and expense in efforts
toward the prevention and control of this prevalent evil, would have
occurred. It was also evident froin the conditions observed in some
sections that practical means of greatly mitigating the destructive
effects of these separate and combined causes of prevailing troubles
could be had through a better knowledge of some of the leading
features of this problem.

CONSIDERATION OF PREVENTIVES AND REMEDIES.

The object of the trip was to study insect enemies of forests and
causes of prevalent unhealthy conditions, rather than the subject of
preventives and remedies. Indeed, much additional and detailed
investigation relative to the life histories ‘and habits of the more
destructive enemies, and of the conditions which affect them is neces-
sary in order to obtain evidence on which to base reliable conclusions.

The fact that the Western destructive pine-bark beetle is attracted
to girdled trees, suggested girdled and felled trap trees as a practical
method of preventing and checking destructive outbreaks of this and
other insects with similar habits, especially in forest reserves, and I
think some experiments should be conducted in order to determine
whether or not this would prove effectual.

The discontinuance, so far as possible, of the common practice of
settlers and farmers girdling pine and spruce trees and leaving them
standing in the clearing from year to year was suggested by the con-
ditions observed as a most important precaution against the multipli-
cation and destructive outbreaks of certain dangerous enemies ot
conifers. On the other hand if such girdled conifers could be felled

200"

and the bark removed before the broods of bark-beetles emerge, vast
numbers of these pests would be effectually destroyed.

The prevention and intelligent control of forest fires is also believed,
as previously stated, to be a most important precaution against the
extension of insect depredations, as is also the prevention of insect
ravages, and the proper management of timber killed by them, a
necessary precaution against forest fires.

The introduction of natural enemies of destructive insects as a means
of controlling them might prove effectual under certain favorable con-
ditions, but before this or any other method is recommended for the
prevention and control of forest pests, each separate trouble, and the
attendant physical and other conditions prevailing in the region or
forest in which it prevails, should be thoroughly studied.

SUMMARY AND RECOMMENDATIONS.

Before concluding this preliminary report it seems proper to call
attention to some of the insects and subjects relating to their destrue-
tive ravages which the results of my investigation suggest as the most
important for investigation. The insects mentioned in the following
list were found to be of special economic importance on account of
their destructive habits, and therefore demand primary consideration:

Dendroctonus brevicomis Lec., a widely distributed and common
barkbeetle, destructive to the Western yellow pine and the sugar
pine; observed in California, Washington, Oregon, and Idaho.

Dendroctonus n. sp., a common and destructive enemy of the white
pine and the sugar pine; observed in Oregon and Washington, and
reported from Montana and South Dakota.

Dendroctonus similis Lee., a very common and widely distributed
enemy of the red fir and Western larch; observed in California,
Oregon, and Idaho.

Scolytus preceps Lec. and closely allied species, one or more of
which are very common, widely distributed, and destructive enemies
of the fir; observed in California, Oregon, and Idaho.

Geometrid larve, destructive to the foliage and causing the death of
vast quantities of Tideland spruce and hemlock in Clatsop and Tilla-
mook counties, Oreg.; the result of its work observed. .

Melanophila drummondi Kirby, or a closely allied species, a common
and destructive enemy of the red fir, hemlock, and noble fir; larve
and work observed in Oregon, Washington, and Idaho.

Neophasia menapia Feld., the pine butterfly, a common and destruc-
tive enemy of the Western yellow pine in Idaho and eastern Washing-
ton; its destructive work observed.

The first five Insects named in this list are probably the five worst
insect enemies of the Western forest. The evidence found indicates
that they have caused the death of many million dollars’ worth of the

2

finest timber and that they are capable of causing greater losses in the
future. Nothing whatever seems to have been published regarding.
the habits of any of them, and little or nothing was known of their
economic importance previous to the investigation here reported.

The interrelation of forest fires, insect enemies, diseases of trees, and
human agencies in the destruction of forests, is a problem of especial
importance, the investigation of which will, no doubt, yield most valu-
able results. |

I would also suggest the importance of conducting experiments with
girdled and felled trap trees in some of the forest reserves threatened
by bark beetles, to determine their value in preventing and controlling
the ravages of destructive bark-infesting insects. Experiments in
partially burning dead timber and the débris left by timber cutters, at
a time of year when the conditions are least favorable for its spread, is
also a line of work which seems to be promising in results.

A study of certain prevailing physical conditions in the Northwest
and their relation to distribution, common or rare occurrences, and
destructive habits of the principal insect enemies of the forest, will
also surely lead to results of special interest and value.

In addition to results of economic importance, the investigation of
the subjects mentioned, involving as they do technical studies of the
insects and of broad problems relating to their life, should lead to
valuable contributions to science.

In conclusion, may I suggest that the increasing public interest in
forest protection, the vast forested domain included in reserves and
national parks, the aggregate public and private wealth in undeveloped
forest resources, and the extensive destruction of the best timber each
year by insects and related evils seem to warrant giving this heretofore
neglected feature of the forest problems of this country especial at-
tention. It is a line of scientific research which, in consideration of
its relation to their work and interests can: not fail to receive the
hearty support and cooperation of other divisions of the public serv-
ice—such as the Division of Forestry, the General Land Office, the
State experiment stations and forestry schools.

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