4 ae n sive 1 . Can 4s, yt ue sora ets y ’ ‘ WAN via ’ AGYEN MG yok CR ere ran A ayers ys , . wed past ‘ ‘ tne . ’ vee ‘ vein 7 . rag 104 Beive fs ata Weide ' ‘ yah wee hay, Was tate, Rea ates pepe puree # * Ran Bs Searhces re \ ud toh = & ~ . Peles cigamectouee eentens : sy hey Es Sa oa kerk een Sober ye se Dek Dee ke eT en ok me Vee eis ses Netty 3 Ge SaMe Reeser soe oh avis eae ‘ . SAG AR REE SS NS ty be pagetona Bed hehe BA Aline : n atta os t >> ‘ vee ‘ a neh ee SUA SA ARES ’ a ' ‘ A . Aye Satay Nore; . , . ‘ Pa pies “ te Nees hat that Rae Ms , nee : . ina aha f rege hea get og ep Amp Ra ong wee : ttre Vases rain thay Sete baton ey a ‘ te dry at age y See bet ete eae P . en See kre Se era oe Cae erence " eae btiekenal see? 4 ‘ . te de , . wos . oo A ‘ 5 7 at eee y - Fin? % : re vse s 3 ; ot 7 ic ba is fs Ae f : ‘ 7 . ‘ , ‘ oe weg ’ ’ ‘ el ‘ ake . ‘ J / we hi. , - or end , "4 , ‘ + . " , oa ; eg / : s ’ ofr f= is . ’ days ‘ A retin. = Sentrae a . . ‘ , . ‘ : , { red gers pyt . Pa gt nie pry Une ea . a ghee tif SP BY . Ss ahaa Sy hy a" Evdd gmt amas ‘ : FAO ANE Be age ph ul Rhy ah PT penny 23 . a a7 z pare yas oraiabatier alae te acto , ‘ ’ td a Ney ga Fy tibet gas ogee oat Geb p Bata net ete eR Pbede ‘ than iy abet er a Saask ‘hed tier t Heth D eid pegs 4 pee Se ge wT 7+ 7) 2 BAA BT? bay ad? LEAS Seah perp HME es sien mal age ee te: owes, : ‘ F 4 : ae ’ is 3 Na ‘ ‘ ‘8 Bin Ok Poot & Si vgrceet ae oe ' i ‘ PaO : iz 4 H ¥ $i sa +h ben ‘ ‘ Pa) bee ms ~ ‘ aay a Hi te i PD rf ‘ Wek we pee ys m4 ‘i ri WE paettes, ‘ he, ot ‘ , 4 ? : vie yt ogi ao bs tb RO OPeN aT | he ‘ qué Ys aii eis vos : ' vt A | | i All i Ml vail | i Ia, 4 D> Vp ral wh “> x a = AMA ph Ae ae hfs ry A nui i] h jh b : Healt tike'd ith i all abt A iy b | ay 2 I~ 2 y, %; eM § 5N aes oi a Ts. ‘s iss Vy i “, = ene, 4 % iy j fe { _———— * ne 9 a a Le Ls | a hi hy ge iy | O te, yes F Sy dy SE” ih “7 BN

_>_ Studies of Neotropical caddis flies, VI: On a collection from moruawestemm: Mexico 022 a ee Ee ea The first record of the Paduniellini in the New World (Trichop- tera: Psychomyiidae) 2... se ee eee FOOTE, B. A.: See NOVAK, J. A. FRANCLEMONT, J. G.—Two new species of noctuids (Lepidoptera, Noc- tuidae, .Noctiinae)) 2... 1 3 2 be FROESCHNER, R. C.: See DRAKE, C. J. GAGNE, R. J.—Tinconaria albescentis, a new North American genus and species of Cecidomyiidae found in seeds of Triodia albescens Vasey ( Diptera ) Notes on the genus Thecodiplosis Kieffer in North America, and some generic reassignments (Diptera: Cecidomyiidae) — GALINDO, P. and H. TRAPIDO—Description of Haemagogus aeritinctus, ii 225 73 97 155 338 a new species from British Honduras, with a note on the validity of Haema- gogus lucifer (H. D. and K.) (Diptera: Culicidae) _... 103 GHOSH, A. K.: See MEDLER, J. T. GURNEY, A. B.—A substitute name for the genus Nicuesa in cockroaches eDictyopteras blattanias vblattellidae)); 022 ee) ee 9387 See WALKER, T. J. HERRING, J. L. and H. C. CHAPMAN—A new species of Omania from Micronesia «@klemiptend: eS alaigdac))) eles tobe ano a eT Ve 8 NY 354 HIBAERIG Ke iwAt——Sexa ratloswOlelatypiuse se eee ee eee ee ee 997 HODGES, R. W.—The trivial name of the potato tuberworm (Lepidoptera: (Celechitd acy) mess eS ee See Se eas See Pd alread er cota x 52 JAMES, M. T.—The blow flies of Dominica (Diptera: Calliphoridae) _. 368 JOHNSTON, D. E.—On the occurrence of two species of Palaeacarus in the eastem, United. States, (Acari: Acaniformes))\_.2 =. 4 A 301 Observations on Oribatei. The discovery of Haplochthonius simplex (Willmann) in North America (Acari: Acariformes) — 365 JOY, S. J.: See McDANIEL, B. KINGSOLVER, J. M.—On the genus Rhipibruchus Bridwell, with descrip- tions of a new species and a closely related new genus (Coleoptera: SYAEYCUTIUG EYEE IBV abil arb Vel) |! eesee tee ime! ay A eee see, Ms Ne Seer Owes eee ee eee 318 and R. E. WHITE—A review of the genus Aulonium from thes United: States, (Coleoptera: Colyditdae)) = 149 KRAMER, J. P.—New Neotropical Neocoelidiinae with keys to the species of Coelidiana, Xenocoelidia and Nelidina (Homoptera: Cicadellidae) —_ 31 A key to the species of Lystridea Baker with description of a new species from California (Homoptera: Cicadellidae: Errhomenellini) _ 292 LEONARD, M. D.—Macrosiphon rosae (Linnaeus) on Ilex (Homoptera: EXSOLAIVGICG EEYE® es ce ete ae Deere NR eS oe Oe em, |e 59 A list of the aphids of Cyprus (Homoptera: Aphididae) _._.. 259 LIN, C. S.—Bionomics of Cerceris simplex graphica at Lake Texoma (Hy- menopterass «sphecidae:, Philanthinae)) V2 ee ee 312 LIN, N.—Linear copulation in Xylocelia franclemonti Krombein (Hymenop- PETA MO PMECICAG )) ee aes ss WES Pee 2. AS Es 343 MARINKELLE, C. J.—Cimex hemipterus (Fabr.) from bats in Colombia, South: America (Hemiptera: Cimicidae) 179 MARSH, P. M.—The Nearctic Doryctinae, IV. Notes on the genus Dendro- soter Wesmael and description of one new species (Hymenoptera: Bra- (SOLTIG FG) ote es an re aie sal AD. ba ae A ee Re, rc A a a a eR 246 The Nearctic Doryctinae, V. The genus Leluthia and comments on the status of the tribe Hecabolini (Hymenoptera: Braconidae) —_ 359 McDANIEL, B.—Lectotype designation from three syntypes of Radfordia sebuliger liwing (Acarma:: Myobiidae)). 183 . J. P. SHOEMAKER and S. J. JOY—The discovery of Listro- phorus leuckarti Pagenstecher on Microtus pennsylvanicus from North paaenca. ( Acamma: Jisttophomoaae) ise Fe Pe 340 MEDLER, J. T.—A note on Ceropales maculata fraterna Smith in Wiscon- Sink li vmMenopteras POmpmiGae))) 08 2 sf ee 92 and A. K. GHOSH—An alate aphid with an extra cornicle Gilomoptera i ecAphididae))s | ssa¥s") be se wk aT) at te) eh BA 366 MENDEZ, E.—Description of a new genus and species of Trimenoponidae from, Panama .(Mallophagays |... eae ee ee eee MOCKFORD, E. L.—Some Psocoptera from plumage of birds — MUESEBECK, C. F. W.—A new braconid parasite of the potato tuberworm (Hymenoptera: 352 ee ee eee NOVAK, J. A., W. B. STOLTZFUS, E. J. ALLEN, and B. A. FOOTE—New host records for North American fruit flies (Diptera: Tephritidae) OLIVE, A. T.—Chromosomes of the aphid genus Dactynotus (Homoptera) — PEYTON, E. L.: See SCANLON, J. E. PIPKIN, S. B.—A new member of a sibling set belonging to the Drosophila tripunctata group (Diptera: Drosophilidae) —... POLHEMUS, J. T.—Notes on North American Saldidae (Hemiptera) — A new saldid from California (Hemiptera: Saldidae) —- PRICE, R. D.—The status of Ratitiphagus batocina Eichler (Mallophaga: Menopomidae) et. a 2 8 he ee oe ee eb ee and K. C. EMERSON—Additional synonymies within the amblyceranAbird lice (Mallophaga) ....._ = =. ee eee RATTANARITHIKUL, M.: See BRAM, R. A. ROBINSON, H.—New species of Dolichopodidae from the United States and Mexicoe Diptera): 2.0. b A ee, ee ee Neoparentia, a new genus of American Dolichopodidae (Diptera) New species of Micromorphus from the United States and Mexico (Diptera: Dolichopodidae)! __.- eee eee RUSSELL, L. M.—A list of the species of Macrosiphoniella Del Guercio described from North America (Homoptera: Aphididae) SABROSKY, C. W.—Notes on the tachinid genus Cylindromyia in North Americal\(Diptera)i 220") 8 a eee SCANLON, J. E. and E. L. PEYTON—Anopheles (Anopheles) tigertti, a new species of the aitkenii group from Thailand —--_-----______ SCHOEPPNER, R. F.: See WHITSEL, R. H. SHENEFELT, R. D.—Protection of insects being transported in liquid ___ SHOEMAKER, J. P.: See McDANIEL, B. SLATER, J. A.—Synonymy in the Lygaeidae (Hemiptera) SMILEY, R. L.—Further studies on the Tarsonemidae (Acarina) — Dolichomotes navei, a new genus and new species of pyemotid mitey;(Acarina: Pyemotidae) = se Eee SMITH, D. R.—Extended distribution for Pristiphora abbreviata (Hartig) (iymenoptera:,.. Tenthredinidae) 2... eee Two sawflies new to North America (Hymenoptera: Ten- thredinidae) (i. ee. ee | eee New synonymy in Pleroneura Konow (Hymenoptera: Xyeli- A review of the subfamily Heterarthrinae in North America (Hymenoptera: Tenthredinidae) |. *. .. § a eee SPANGLER, P. J.—A new Brazilian Berosus and a description of the female of B. spectatus D’Orchymont (Coleoptera: Hydrophilidae) — STEYSKAL, G. C.—A key to the species of Tritoxa Loew (Diptera: Otitidae) iv 287 307 Ua 146 303 1 24 346 367 248 114 252 329 184 Palaeosepsis species described by Melander and Spuler (Diptera: “SSIS EG) aa aa a eR ae oe Replacement names for preoccupied specific names in Dolicho- PICS (Dips Fem as) a f Anthomyia procellaris Rondani in North America (Diptera: Akad raroyatlers) ae Sa ae Te ee ee ee ee eee A new subgenus for Leucopis luteicornis Malloch, a predator on mealybugs in India (Diptera: Chamaemyiidae) —....- Distinguishing Amoebaleria defessa (Osten Sacken) from A. saekeni Garrett (Diptera:. Heleomyzidac) 22.2 ee Something better than polyporus or pith for double mounts __ 2 STOLTZFUS, W. B.: See NOVAK, J. A. STONE, A.—A synoptic catalog of the mosquitoes of the world, supplement Me GDinteriaGulicidae))). 2s. ae Bae ye ee ee ee STRASSEN, R. ZUR—New locality records and a new species of Chiro- Latins Halidayss(ihysanoptera:s Lbripidae)) #22") TOWNES, H.—Taxonomic notes on Kentrotryphon, Ktenostilpnus and Pa- rentypoma (Hymenoptera: Ichneumonidae) _...-- Notes on the Motschoulsky types of Ichneumonidae ( Hymenop- A new Ateleute from the United States (Hymenoptera: Ich- ME MINGTMICAG:)/ cet. eR RRM ot. RAM eee Re eae ee = TRAPIDO, H.: See GALINDO, P. WALKER, T. J. and A. B. GURNEY—tThe metanotal gland as a taxonomic character in Oecanthus of the United States (Orthoptera: Gryllidae) WEBER, N. A.—Synonyms of Trachymyrmex bugnioni Forel and Trachymyr- mex diversus Mann (Hymenoptera: Formicidae) _.. WHITE, R. E.—Ptilinus friendi, a nomen nudum (Coleoptera: Anobiidae) _ See KINGSOLVER, J. M. WHITSEL, R. H. and R. F. SCHOEPPNER—Parasitic trombidiid mites on Dasyhelea mutabilis (Coquillett) (Diptera: Ceratopogonidae) WILSON, N.—Ectoparasites of Canadian birds and mammals _ “a 275 296 For more than half a century, Cyanamid has consistently led the chemical industry in developing new products and application techniques that have helped immeasurably to bolster our na- tional farm economy. ™@ Topping the list of Cyanamid ‘firsts’ is Malathion LV* Concentrate, introduced commercially last year for the control of boll weevils on cotton after two years of use on more than 1 million treatment acres in cooperation with the U.S.D.A.’s Agricultural Research Service. m Malathion LV Con- centrate is also being used extensively to combat grasshoppers, cereal leaf beetles, corn rootworm beetles, mosquitoes, blue- berry maggots, flies and beet leafhoppers. m Watch for progress reports of new tests conducted against many other pests with both aerial and ground equipment. Data being processed daily show clearly that Malathion LV Concentrate is fast making all other methods of insect control obsolete! m@ Before using any pesticide, stop and read the label. *Trademark SERVES THE MAN WHO MAKES AMERICAN CYANAMID COMPANY A BUSINESS OF AO aaa PRINCETON, NEW JERSEY ‘ol. 69 MARCH 1967 No. 1 95+ 10678 sels PROCEEDINGS of the ENTOMOLOGICAL SOCIETY of WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 PUBLISHED QUARTERLY (Continued on back, cover) | BOYD, E. M.—Deutonymphs as endoparasites of the “Eastern belted king- fisher and the Eastern green heron in North America (Acarina: Sarcopti- PRTMTN ER EaT ENO LEC LAG AG) meee er cs ee eh ie eee) BRAM, R. A. and M. RATTANARITHIKUL—Six new species of the Culex (Lophoceraomyia) mammilifer group from Thailand (Diptera: Culicidae) 1 BURKS, B. D.—A new polyembryonic parasite of the pepper moth (Hy- (oo unpl eet gs) EP air iT rcp TS ll so IG gs es Sho mene CORNELL, J. F.—New Coleoptera records from North Carolina DENNING, D. G. and J. W. BEARDSLEY—tThe collection of Cheumato- psyche analis in Hawaii (Hydropsychidae: Trichoptera) — 56 DRAKE, C. J. and R. C. FROESCHNER—Lacebugs of the Galapagos meeneneiaem -( temipteras Lingidae) (2.2 oe 82 DUCKWORTH, W. D.—A new species of Atteva from Central America Peerumantera Yoonomentidae) 21.0 a FLINT, O. S., JR.—A new synonymy in the Trichoptera HODGES, R. W.—The trivial name of the potato tuberworm (Lepidoptera: Gelechiidae ) KRAMER, J. P.—New Neotropical Neocoelidiinae with keys to the species of Coelidiana, Xenocoelidia and Nelidina (Homoptera: Cicadellidae) ENTOMOLOGICAL SOCIETY OF WASHINGTON OrGANIZED Marcu 12, 1884 OFFICERS FOR 1967 a ra, =. Mr. Louis G. Davis, President Dr. Jon L. Hernine, Editor ARS, USDA, Rm. 466 c/o Department of Entomology Federal Center Bldg. U.S. National Museum Hyattsville, Maryland 20782 Washington, D.C. 20560 Dr. RicHarp H. Foote, President-Elect Mr. RoBERT SMILEY, Custodian c/o Department of Entomology Entomology Research Division, USDA U.S. National Museum Washington, D.C. 20250 Washington, D.C. 20560 Dr. Raveu A. Bram, Recording Secretary Mr. Victor ADLER, Program Committee Chairman Department of Entomology Entomology Research Division, ARS, USDA U.S. National Museum ARC, Beltsville, Maryland 20750 Washington, D.C. 20560 Mr. Davi R. Smitn, Corresponding Secretary Capr. Witt1AM B. Hutt, Membership Committee c/o Department of Entomology Chairman U.S. National Museum 746 Beall Ave. Washington, D.C. 20560 Rockville, Maryland 20850 Dr. ARTHUR K. BuroitT, JRr., Treasurer Dr. H. H. SHeparp, Delegate to the Washington — ARS, USDA, Rm. 125 Academy of Sciences North Bldg. PPA-ASCS Beltsville, Maryland 20705 U.S. Department of Agriculture Washington, D.C. 20250 Honorary President Dr. T. E. Snyper, U.S. National Museum Honorary Members Mr. C. F. W. MurEseseck, U.S. National Museum Dr. F. W. Poos Dr. E. N. Cory in the Proceedings. MEMBERSHIP.—Members shall be persons who have demonstrated interest in the science of entomology. Annual dues for members are $5.00; initiation fee is $1.00 (U.S. currency). PROCEEDINGS.—Published quarterly beginning with March by the Society at Washington, D.C. Members i in good standing are entitled to the Proceedings free of charge. Nonmember subscriptions are $6.00 per year, both domestic and foreign (U.S. currency), payable in advance. All remittances should be made payable to) The Entomological Society of Washington. ft The Society does not exchange its publications for those of other societies. dq ; STATEMENT OF OWNERSHIP Title of Publication: Proceedings of the Entomological Society of Washington. Frequency of Issue: Quarterly (March, June, September, December). Location of Office of Publication, Business Office of Publisher and Owner: The Eaton Society “ Washington, c/o Department of Entomology, U.S. National Museum, Washington, D.C. 20560 Editor: Dr. Jon L. Herring, same address as above. | Managing Editor and Known Bondholders or other Security Holders: none. This issue mailed March 28, 1967 Second Class Postage Paid at Lawrence, Kansas, U. S. A. ALLEN PRESS, INC. Cauley.) LAWRENCE, KANSAS US, A. PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Vol. 69 MARCH 1967 No. 1 SIX NEW SPECIES OF THE CULEX (LOPHOCERAOMYIA) MAMMILIFER GROUP FROM THAILAND (Dierera: CuLicmaAer)! RaupuH A. BRAM and MANop RATTANARITHIKUL South East Asia Mosquito Project, Department of Entomology, Smithsonian Institution, Washington, D. C. 20560? The most recent and complete taxonomic treatment of the Culex subgenus Lophoceraomyia in any area of South East Asia was that of Colless (1965). This study described 14 new species, revalidated five synonyms, and generally placed the taxonomy of this difficult group on a sound basis for future investigations. The present study describes six new species from Thailand, all of which belong to the mammilifer subgroup of the mammilifer group as defined by Colless (1965). The mammilifer group of the subgenus Lophoceraomyia is distinguished from the fraudatrix group mainly by the presence of a mammiliform protuberance on the inner surface of the male antennal torus. In the male terminalia, members of the mammilifer group exhibit both an internal process and a spinose or toothed dorsal process which compose the lateral plate of the phallosome; members of the fraudatrix group possess a toothless dorsal process only. In the larval stage members of the mammilifer group have head hairs 4-C distinctly longer than the distance between their bases, in contrast to the fraudatrix group in which these hairs are shorter than the distance between their bases. The mammilifer subgroup of the mammilifer group is recognized by the presence of one lower mesepimeral bristle in the adult, by the larval antenna with hairs 2 and 3 well separated from the apex of the shaft, and by the general container habitats of the larvae; the brevipalpus subgroup of the mammilifer group lacks a lower mesepimeral bristle in the adult, hairs 2 and 3 of the larval antenna are inserted at the apex of the shaft, and the larval habitat is generally restricted to pitcher plants (Nepenthes spp. ). ‘This work was supported by Research Contract No. DA-49-193-MD-2672 from the U. S. Army Medical Research and Development Command, Office of the Surgeon General. Immediate publication secured by full payment of page charges—Editor. 2 Present address of second author—Public Health Division, USOM, Bangkok, Thailand. PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, bo —TL HWANG 1967 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 3 In the following descriptions, terminology of the adult habitus and the immature stages conforms primarily to the format of Belkin (1962); terminology applied to the male antenna and male terminalia follows Colless (1965), except as noted below. Colless referred to the first flagellar division of the male antenna as segment III; in our terminology the first flagellar division is called flagellomere I, rec- ognizing only three true antennal segments (namely, the scape, the pedicel or torus, and the flagellum). Structures of the male terminalia recognized as coxite and style of coxite by Colless are termed basimere and distimere, respectively, in this study. Material utilized in this study is housed in the U. S. National Museum and consists principally of collections made by the Department of Medical Entomology, South East Asia Treaty Organization Medical Research Laboratory, Bangkok, Thailand, and to a lesser extent, the D. C.—E. B. Thurman mosquito collection. Culex (Lophoceraomyia) spiculosus, species novum The adult female is very similar to that of C. traubi Colless and cannot be separated with certainty. The male is recognized by the absence of conspicuous tufts on antennal flagellomeres V and IX, the submarginal setae of the basimere are arranged in a regular row, and the internal process of the lateral plate of the phallosome does not project beyond the dorsal process. The fourth stage larva is characterized by possessing a long siphon with 4 pairs of strong, long, subventral tufts, having head hair 5-C bifid or trifid, 16, 17-C absent, and having the thoracic and abdominal integument covered with numerous, prominent spicules. Female. Proboscis dark brown with a pair of prominent basoventral bristles and a pair of smaller setae between them; palpus similar in color to the proboscis; antenna longer than the proboscis; decumbent scales of the vertex narrow and dark, somewhat broader and pale immediately adjacent to the orbital line; erect scales forked, uniformly dark brown. Scutum and midlobe of scutellum sparsely covered with dark scales; pleural integument uniformly pale, tinged with green in fresh specimens; one lower mesepimeral bristle present. Wings normal, outstanding scales narrow, including those on veins Re and Rs. Anterior surface of hind femur predominantly pale, with a narrow dark stripe along the dorsal border and a dark band at the apex; hind tibia and tarsus com- pletely dark; mid- and fore-legs uniformly dark scaled, although in some specimens the anterior surface of the mid-femur may exhibit a somewhat paler basoventral area on the anterior surface. Abdominal terga covered with dark brown scales; sterna slightly lighter than the terga. < Fig. 1, Culex (Lophoceraomyia) spiculosus sp. nov. Dorsoventral view of the fourth stage larva, dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. voL. 69, No. 1, MARCH, 1967 ENT. SOC. WASH., PROC. TL HWANG PROG. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 5 Male. Similar in general appearance to the female; length of palpus slightly greater than the length of the proboscis. Flagellomeres V through IX as illustrated in figure 1; flagellomere V with a group of from 6 to 8 very narrow, acute scales whose apices do not reach beyond flagellomere VII; flagellomere VIII with an internal tuft of long and rather straight, broad, dark brown setae; flagellomere IX without a tuft of obviously modified setae. Terminalia as illustrated in figure 1; basimere with from 6 to 8 strong submarginal setae inserted in a straight row; subapical lobe of the basimere with the three rods subequal in length and width, gently curved and pointed apically; internal leaflet rodlike, but broad and bent; external leaflet oblong-ovate, pointed apically and striate; accessory proc- esses narrow, setae-like; distimere normal, the apex without annulations; lateral plate of the phallosome with the internal process distinctly curved and pointed distally, not projecting beyond the apex of the dorsal process; dorsal process with an apical knob and approximately 10 short, but distinct, teeth on the lateral margin. Larva. Chaetotaxy and structure as illustrated in figure 1. Head _ lightly pigmented, antenna concolorous with head capsule but with a dark basal ring and progressively darker beyond insertion of antennal hair 1-A; 1-A large, multiple, pectinate, but with the lateral hairlets on only one side of the individual branches. Head hair 1-C darkly pigmented, tapering to a sharp point, its length greater than half the distance between bases of the pair; 4-C single, simple; 5-C with 2 or 3 branches, pectinate, 6-C bifid, subequal in length; 16, 17-C absent. Thoracic and abdominal integument densely spiculose throughout, the spicules of the thorax slightly more prominent than those of the abdomen; thoracic hairs 1, 2-P single, pectinate, subequal in length; 3-P single, pectinate, shorter and slenderer than 1, 2-P; 4-P bifid, pectinate; 5, 6-P single, pectinate; 7-P bifid, pectinate; 8-P single, pectinate; 14-P bifid, simple. Comb consisting of from 35 to 45 fan- shaped scales arranged in a broad, triangular patch. Siphon index variable, ranging from 7:1 to 11:1 (average, 8.8:1); 4 pairs of subventral tufts inserted in a line on the siphon; individual tufts 2 to 5 branched, finely pectinate, their length greater than the width of the siphon at the point of insertion; pecten consisting of from 11 to 14 teeth restricted to approximately the basal fourth of the siphon; individual pecten tooth with a prominent distal spine and approximately 6 to 8 lateral barbs, the proximal 2 or 3 barbs very coarse. Type Data. Holotype male with associated larval and pupal skins and terminalia and antennae slide mounted from Doi Sam Sao, Tak Province, Thailand, deposited in the U. S. National Museum, No. 69151. Paratypes: 1 male and 2 females with associated larval and pupal skins with the same data as the holotype. The specific name applies to the characteristic thoracic and abdominal spiculation of the fourth stage larva. Distribution. Known only from the following Provinces in Thai- land: Tak, Nakhon Nayok, and Chiang Mai. Eleven individual < Fig. 2, Culex (Lophoceraomyia) peytoni sp. nov. Dorsoventral view of the fourth stage lava, dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. 6 PROG. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Zo BA ’ \ aN \ —TL. HWANG Fig. 3, Culex (Lophoceraomyia) eukrines sp. nov. Dorsoventral view of the fourth stage larva, dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 7 rearings have been examined as well as 6 males, 3 females, and 50 larvae. Taxonomic Discussion. This species and peytoni sp. nov. clearly fall within the sibling species complex of C. traubi-C. ganapathi rec- ognized by Colless (1965). Its closest affinity, both in anatomical features and larval biology, is with traubi from which it can be separated as follows: in the male terminalia of spiculosus sp. nov. the internal process of the phallosome does not project beyond the dorsal process, a condition which does exist in traubi; in the larva, hair 2 of the anal segment is single in spiculosus sp. nov. as opposed to bifid in traubi. Additional distinguishing features may be found in the male antennae. Biology. Larvae of this species have been collected on 16 occasions from primary rain forests; once from a bamboo stump, once from a bamboo internode, once from a stump hole, and on all other occasions from large tree holes. Collections were made during July, August, and September. Habits of the adults are unknown. Culex (Lophoceraomyia) peytoni, species novum The female is indistinguishable from spiculosus sp. nov. The adult male may be recognized by the presence of only a few yellowish long hairs on flagellomere V, an internal tuft of at least 5 strongly sigmoid setae on flagellomere VII, a tuft of obviously modified, dark, strong setae on flagellomere IX, and by the internal rod of the subapical lobe of the basimere slightly expanded subapically, followed by a rather broad, gently curved apex. The fourth stage larva possesses a long siphon with 4 pairs of strong, long, subventral tufts; the thoracic integument is covered with fine spicules, but the abdominal integument is mostly smooth and the lateral denticles of the individual pecten teeth are very broad proximally. Female. No features have been found which differ consistently from the description presented for spiculosus sp. nov. Male. Similar to spiculosus sp. nov. except as noted below. Flagellomeres V through IX as illustrated in figure 2; flagellomere V with a group of approximately 6 narrow, setaelike scales which extend to approximately the middle of flagellomere IX; flagellomere VIII with an internal tuft of from 6 to 9 gently curved, dark brown setae; and flagellomere IX with a small internal tuft of 3 strong, rather long, dark setae. Terminalia as illustrated in figure 2; basimere with 4 or 5 strong submarginal setae inserted in a straight row; subapical lobe of the basimere with the internal rod slightly expanded subapically, then bent and with a rather broad, lightly sclerotized apex; the central and external rods sub- equal in length, sharply hooked apically; accessory processes narrow, but some- what broader than those found in spiculosus sp. nov.; internal leaflet rod- like, gently bent; external leaflet slender, striate; distimere with 1 or 2 additional setae on the proximal third; dorsal process of the lateral plate of the phallosome with a subapical knob and 5 short, but distinct teeth on the lateral margin. Larva, Chaetotaxy and structure as illustrated in figure 2, similar to spiculosus sp. nov. except as noted below. Head hairs 16, 17-C represented by minute spicules which are easily overlooked even in good preparations. Thoracic integu- ment sparsely spiculate, the spicules fine and rather long; abdominal integument 8 PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 Fig. 4, Culex (Lophoceraomyia) pholeter sp. nov. Dorsoventral view of the fourth stage larva, dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 9 glabrous. Thoracic hair 7-P single or bifid, pectinate. Comb consisting of a broad, triangular patch of from 35 to 50 elongate, fan-shaped scales. Siphon index variable, ranging from 6.6:1 to 9.2:1 (average, 8:1); 4 pairs of subventral tufts inserted in a line on the siphon; individual tufts 3 to 5 branched, their length greater than the width of the siphon at the point of insertion; pecten consisting of from 10 to 14 teeth restricted to the basal fourth of the siphon; individual pecten tooth with an extended, fine distal spine, and from 6 to 10 lateral barbs, the basal barb (sometimes the proximal 2 or 3 barbs) very large and rather rounded apically. Type Data. Holotype male with associated larval and pupal skins and terminalia and antenna slide mounted from Phatthalung Province, Thailand, 34 kms. on Phatthalung to Trang Road, 15. X. 1964, K. Mongkolpanya, from tree hole 12 feet above ground, deposited in the U. S. National Museum, No. 69182. Paratypes: 1 male and 1 female with associated larval and pupal skins with the same data as the holotype, and 2 males from Ko Chang Island, Ranong Province, Thailand, 16. IX. 1964, K. Mongkolpanya, from a tree stump hole with associated larval and pupal skins. This species is named in honor of SFC E. L. Peyton of the SEATO Medical Research Laboratory. Distribution. The authors have seen specimens from the following Provinces in Thailand: Trang, Chiang Mai, Sara Buri, Narathiwat, Phatthalung, Chumphon, Ranong, and Chanthaburi. Sixteen indi- vidual rearings have been studied, as well as 4 additional males and 70 larvae. Taxonomic Discussion. As indicated under spiculosus sp. nov., this species falls within the traubi—ganapathi sibling species complex. Its anatomical affinity seems closest to ganapathi, but larval biology is rather intermediate between the two. The adult females cannot be distinguished from the three other members of the complex. The male may be separated from ganapathi primarily by the presence of a tuft of modified setae on flagellomere IX as well as the presence of 1 or 2 setae on the proximal third of the distimere; the male differs from traubi and spiculosus sp. nov. by the presence of a tuft of long, narrow, tapering scales on flagellomere V whose apices reach to approximately the middle of flagellomere IX. The larva may be distinguished from ganapathi by the form of the pecten tooth in which the lateral denticles are very broad basally and by the fact that head hair 1-C is never expanded on the basal half as frequently found in ganapathi; the larva is distinguished from traubi and spiculosus sp. nov. by the glabrous abdominal integument and the undeveloped median caudal filament. Biology. Larvae have been collected on 33 occasions from tree holes, root holes, bamboo internodes, and bamboo stump holes in primary rain forests. One collection was made from a rock hole in dense forest. Habits of the adults are unknown. 10 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 : 14 ov ™ vill 5 Mayslidg OF a8 —JT71. HWANG PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 1] Culex (Lophoceraomyia) eukrines, species novum The adult female is virtually indistinguishable from spiculosus sp. nov. and its relatives, but does exhibit slight differences in the scaling patterns on the anterior surface of the hind femur. The adult male exhibits an internal tuft of 3 gently curved setae on flagellomere VII and a tuft of obviously modified, dark, strong setae on flagellomere IX. The larva may be immediately recognized by the head hair 6-C which is always single and stout. Female. Similar to spiculosus sp. nov. except for the following difference. Anterior surface of hind femur completely pale on the proximal half, with a narrow apical dark band which extends proximally along the dorsal border. Male. Similar to spiculosus sp. nov. except as noted below. Flagellomeres V through IX as illustrated in figure 3; flagellomere V with a tuft of 6 very narrow, pale setae extending beyond flagellomere IX; flagellomere VII with an internal tuft of from 3 to 5 gently sigmoid, specialized setae and several shorter, straight setae; flagellomere IX with a tuft of from 3 to 5 long, strong, dark setae. Terminalia as illustrated in figure 3; similar to peytoni sp. nov. but with 5 or 6 submarginal setae on the basimere, without setae on the basal third of the distimere, and with approximately 15 teeth on the lateral margin of the dorsal process of the lateral plate of the phallosome. Larva. Chaetotaxy and structure as illustrated in figure 3, similar to spiculosus sp. nov. except as noted below. Head hair 5-C usually bifid, sometimes single, pectinate; 6-C always single, pectinate, somewhat stouter than the individual branches of 5-C; 16, 17-C represented by minute spicules. Thoracic and abdominal integument glabrous; thoracic hairs 5, 6, 7, 8-P single, pectinate, but 7-P occasionally bifid. Comb consisting of from 35 to 50 elongate, fan-shaped scales arranged in a broad triangular patch. Siphon index variable, ranging from 7:1 to 10:1 (average, 8.2:1); 4 pairs of subventral tufts inserted in a line on the siphon; individual tufts with from 2 to 4 branches, their length greater than the width of the siphon at the point of insertion; pecten consisting of from 8 to 12 teeth restricted to the basal third to fourth of the siphon; individual pecten tooth with a prominent distal spine and from 5 to 9 lateral barbs, the proximal 2 or 3 barbs more prominent than the apical barbs. Type Data. Holotype male with associated larval and pupal skins and terminalia and antenna slide mounted from Huai Bong Ti, Kanchanaburi Province, Thailand, 2. VI. 65, K. Mongkolpanya, from rock pool in bamboo grove, deposited in the U. S. National Museum, No. 69183. Paratypes: 1 male and 2 females with associated larval and pupal skins from Khao no Chang, Kanchanaburi Province, Thailand, 6. VI. 65, E. L. Peyton and K. Mongkolpanya; and 1 male with associated larval and pupal skins from Ban Sai Yok, Kanchanaburi Province, Thailand, 26. V. 1965, S. Chumchulcherm, from a rockhole in a bamboo grove in a mountainous area. The specific name is derived from the Greek adjective meaning clear or distinct. < Fig. 5, Culex (Lophoceraomyia) fuscosiphonis sp. nov. Dorsoventral view of the fourth stage larva, dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. 12; PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 ——— Oe area JL. HWANG Fig. 6, Culex (Lophoceraomyia) incomptus sp. nov. Dorsal view of the male terminalia, and lateral view of the antennal flagellomeres V through IX. Distribution. The authors have examined 24 collections from Kanchanaburi Province, Thailand and one collection each from Nakhon Nayok and Trang Provinces. Fourteen individual rearings were studied, as well as 17 additional males, 21 additional females, and 120 larvae. Taxonomic Discussion. The adult stage (both male and female) demonstrates affinity to the traubi—ganapathi sibling species com- plex but the larva appears sufficiently distinctive to exclude it from this complex. The female differs from the species previously men- tioned only on the basis of the scale patterns of the anterior surface of the hind femur, and this character is frequently difficult to evaluate. The adult male is differentiated from the members of the traubi— ganapathi complex by the presence of an internal tuft of approximately 3 gently sigmoid specialized setae on flagellomere VII; it differs from peytoni sp. nov. by lacking setae on the basal third of the disti- mere and by possessing approximately 15 teeth on the dorsal margin of the outer process of the lateral plate of the phallosome; it differs from spiculosus sp. noy. on the basis of the internal rod on the subapical lobe of the basimere which is slightly expanded sub- apically then bent and with a rather broad, lightly sclerotized PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 13 apex. The larva is primarily distinguished from the members of the traubi—ganapathi complex by the glabrous thoracic and abdominal integument and by the fact that head hair 6-C is always single, rather stout, and apparently somewhat flattened. Biology. The type habitat of this species was a rock hole located on the face of a cliff in a secondary rain forest which contained about two gallons of water. Larvae of eukrines sp. nov. have also been collected from flood pools in a primary rain forest environment, bamboo internodes in a primary rain forest, a coconut shell in a secondary rain forest, a wheel track in a bamboo grove, a teak tree hole, and a pandanus axil in a primary bamboo grove. Most of the collections, however, were made in mountainous areas from bamboo internodes, or rock pools or other container habitats in close association with bamboo groves. Nothing is known of the adult biology. Culex (Lophoceraomyia) pholeter, species novum The female is virtually indistinguishable from spiculosus sp. nov. The adult male is distinguished by flagellomere V possessing a conspicuous tuft which has from 1 to 4 long, broad scales, by the presence of 3 long, prominent submarginal setae on the basimere, and by the shape of the internal process of the lateral plate of the phallosome. The fourth stage larva exhibits a long siphon with 4 pairs of long, rather fine, usually bifid subventral tufts and thoracic hair 8P is branched and reduced in size. Female. The adult female is virtually indistinguishable from spiculosus sp. nov., but the abdominal sterna are very pale in contrast to the condition found in spiculosus sp. nov. Male. Similar in general appearance to spiculosus sp. nov. except as noted below. Flagellomeres V through IX as illustrated in figure 4; flagellomere V with at least one very broad scale which tapers to a sharp point and extends to flagellomere VIII, as well as several more narrow, but equally long setae-like scales; flagellomere VIII with a tuft of 5 or more strong, broad, dark setae which exhibit a prominent median bend; flagellomere IX with an internal tuft of 3 to 5 long, rather straight, strong setae. Terminalia as illustrated in figure 4; basimere with 3 strong submarginal setae inserted in a straight row; subapical lobe of the basimere with the internal rod tapering to a sharp point and somewhat shorter than the central and external rods which are subequal in length and_ broadly bent apically; internal leaflet rod-like, fine; external leaflet oblong-ovate, bluntly rounded apically and striated; accessory processes narrow, setae-like; distimere minutely annulate on the apical third of the convex surface; dorsal process of the lateral plate of the phallosome with an apical knob and approximately 7 short, but distinct teeth on the lateral margin. Larva. Chaetotaxy and structure as illustrated in figure 4, similar to spiculosus sp. nov. except as noted below. Head hair 1-C darkly pigmented, tapering to a sharp point, its length less than half the distance between bases of the pair; 16, 17-C represented by small, but distinctive, spicules. Thoracic and abdominal integument glabrous. Thoracic hair 8-P with 2 or 3 branches, simple, very short and fine; 14-P bifid or trifid. Comb consisting of a broad, triangular patch of from 60 to 80 elongate, fan-shaped scales. Siphon index variable, ranging from 14 PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 6:1 to 7:1; 4 pairs of subventral tufts inserted in a line on the siphon; individual tufts usually bifid (occasionally trifid or single), their length greater than the width of the siphon at the point of insertion; pecten consisting of from 8 to 11 teeth restricted to approximately the basal third to fourth of the siphon; individual pecten tooth with a prominent distal spine and from 7 to 10 lateral barbs, the proximal two barbs very coarse, the distal barbs fine. Type Data. Holotype male with associated larval and pupal skins and terminalia and antenna slide mounted from Khao Mai Ha Wa, Chon Buri Province, Thailand, 19. VII. 65, E. L. Peyton, from a crab hole in a mountainous secondary rain forest, deposited in the U. S. National Museum, No. 69184. Paratypes: two females with associated larval and pupal skins and one male from the type locality and habitat, 30 VI. 65. The specific name applies to the larval habitat and is taken from the Greek noun meaning one who lurks in a hole. Distribution. This species is known only from the type locality and from Khao Mai Kaeo, Chon Buri Province, Thailand. Three individ- ual rearings were studied as well as 3 additional males and 4 larvae. Taxonomic Discussion. The adult male appears to have its closest affinity to C. mammilifer (Leicester), due to the presence of the conspicuous tuft on flagellomere V; however, it may be separated from mammilifer on the basis of having only 3 prominent submarginal setae on the basimere and the internal process of the lateral plate of the phallosome does not project beyond the apex of the dorsal process. The larva is also similar to that of mammilifer, but can be recognized by the presence of 4 pairs of subventral tufts on the siphon; the form of the thoracic hair 8-P is also distinctive, and hair 6 of abdominal segment I is bifid rather than 3 or 4 branched. Biology. The type habitat of pholeter sp. nov. is a small crab hole in a secondary rain forest in a mountainous area. Another collection was made from a crab hole in a tapioca plantation near the type habitat. Collections were made during June, August, and October. Habits of the adults are unknown. Culex (Lophoceraomyia) fuscosiphonis, species novum The adult female is indistinguishable from spiculosus sp. nov. The adult male is recognized by flagellomere V lacking a conspicuous tuft of long, board scales, by having the prominent submarginal setae inserted in two irregular rows on the basimere, and by the internal process of the lateral plate of the phallosome projecting beyond the apex of the dorsal process. The fourth stage larva exhibits a long, darkly pigmented siphon with 4 pairs of fine, rather short subventral tufts and thoracic hair 8-P single. Female. Indistinguishable from spiculosus sp. nov. Male. Similar to spiculosus sp. nov., except as noted below. Flagellomeres V through IX as illustrated in figure 5; flagellomere V with an internal tuft of about 5 narrow, acute setae whose apices do not reach beyond flagellomere VII; flagellomere IX with a tuft of from 3 to 5 strong, rather long, dark setae. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 15 Terminalia as illustrated in figure 5; basimere with approximately 9 submarginal setae inserted in two parallel but, irregular rows; subapical lobe of the basimere with the internal rod rather broad, constricted at the apex and filamentous beyond the constriction; the central and external rods subequal in length, gently curved apically; lateral plate of the phallosome with the internal process rather narrow, distinctly curved and pointed distally, projecting beyond the apex of the dorsal process; dorsal process with an apical knob and approximately 10 short teeth on the lateral margin. Larva. Chaetotaxy and structure as illustrated in figure 5, similar to spiculosus sp. nov. except as noted below. Head moderately pigmented, antenna slightly darker with a distinctly darker basal ring. Thoracic and abdominal integument covered with a sparse pattern of extremely fine, rather long spicules; thoracic hair 7-P single or bifid, pectinate; 14-P usually bifid, occasionally single, simple. Comb consisting of from 30 to 45 elongate, fan-shaped scales arranged in a broad triangular patch. Siphon index variable, ranging from 10:1 to 12.6:1 (average, 11.8:1); 4 pairs of subventral tufts inserted in a line on the siphon; individual tufts fine, with from 2 to 4 branches, their length equal to, or greater than the width of the siphon at the point of insertion; pecten consisting of from 7 to 10 teeth restricted to approximately the basal fourth or less of the siphon; individual pecten tooth gently curved, with a prominent distal spine and with from 6 to 10 lateral barbs; distal barbs fine, proximal barbs coarse, and conspicuous; siphon and anal segment very darkly pigmented, more so than in any other species examined. The association of the larvae with the male is presumptive: larvae were collected along with a pupa from which the holotype male was reared. Type Data. Holotype male with terminalia and antennae slide mounted from Muang, Phatthalung Province, Thailand, 15. X. 1964; S. Chumchulcherm, from a tree hole in a primary forest, deposited in the U. S. National Museum, No. 69185. Paratypes: one female with its associated larval and pupal skins from the type locality; one male from Muang, Trang Province, 9. X. 1964, S. Chumchulcherm, reared from a root hole. The specific name is of Latin origin and applies to the exceptionally dark siphon of the fourth stage larva. Distribution. In addition to the type locality, this species has also been collected in Thailand from Trad and Trang Provinces. The following material was examined in addition to the types: 24 ¢ reared from pupae; 12 with associated larval and pupal skins; and 13 larvae. Taxonomic Discussion. The adult male of this species is extremely similar to C. demissus Colless, and on the basis of Colless’ description of the unique holotype specimen, the terminalia appear to be insepa- rable; however, the internal tuft of antennal flagellomere V in fus- cosiphonis sp. nov. does not exhibit the distinctively long setae which are said to be present in demissus. The larva is rather similar to that of C. wilfredi Colless but may be distinguished on the basis of the very darkly pigmented siphon and anal segment in fuscosiphonis sp. noy. and by hair 2 of the anal segment which is bifid in the case of fuscosiphonis sp. nov. and trifid in the case of wilfredi. 16 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Biology. The type habitat of this species was a tree hole located in a primary rain forest. Specimens have also been collected from root holes in mountainous primary rain forests and one collection was made from a tree hole in a secondary scrub stand near the sea coast. The biology of the adults is unknown. Culex (Lophoceraomyia) incomptus, species novum The adult male may be recognized by the presence of 2 rows of submarginal setae on the basimere, by the 6 acutely pointed scales on flagellomere V, and by the characteristically bulbous setae on flagellomere VIII. Female. Unknown. Male. Similar to fuscosiphonis sp. nov., except as noted below. Flagellomeres V through IX as illustrated in figure 6; flagellomere V with an internal tuft of approximately 6 acute, striated setae which extend almost to flagellomere VII (these setae are somewhat stronger than those found in fuscosiphonis sp. nov.); flagellomere VIII with an internal tuft of 7 strong, dark, gently curved setae, the distal 2 of these setae with a characteristic bulbous, median expansion; flagellomere IX with an internal tuft of 3 short, slender setae and 3 longer, slenderer setae. Terminalia as illustrated in figure 6; basimere with 10 strong, submarginal setae inserted in two irregular rows; subapical lobe of the basimere with the internal rod rather robust, constricted at the extreme apex and pointed, the central and external rods subequal in length, hooked apically; internal leaflet rod-like, quite slender; accessory processes narrow and setae-like; distimere with minute annulations on the apical third of the convex surface; dorsal process of the lateral plate of the phallosome with an apical spiculate knob and 12 or more teeth on the lateral margin; internal process robust, distinctly curved, not projecting beyond the apex of the dorsal process. Larva. Unknown. Type Data. Holotype male with terminalia and antennae slide mounted from Doi Sutep, Chiang Mai Province, Thailand, 7. I. 53, D. C. and E. B. Thurman, deposited in the U. S. National Museum, No. 69186. Paratypes: two males with terminalia and antennae slide mounted and with the same data as the holotype, deposited in the U.S. National Museum. The specific name applies to the overall adult habitus and is derived from the Latin adjective meaning unadomed. Distribution. In addition to the type locality, one specimen was collected resting at Doi Tad Fah, Chiang Mai Province, Thaildan. Four males and their associated terminalia and antennae were studied. Taxonomic Discussion. This species demonstrates a close affinity to fuscosiphonis sp. nov. and demissus, but the bulbous expansion of setae on flagellomere VII and the shape of the processes on the subapical lobe of the basimere clearly separates this species from its close relatives. C. bandoengensis Brug, another member of the mam- milifer group which exhibits the submarginal setae in two rows, is PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Ibe easily separated on the basis of the shape and number of the subapical lobe of the basimere. Biology. Virtually nothing is known of the biology of this species. Larvae were collected (and the skins subsequently lost) from a tree hole on Doi Sutep Mountain, and one adult male was collected from a low plant under shady damp jungle on Doi Tad Fah Mountain. ACKNOWLEDGMENTS The authors are indebted to Dr. Botha de Meillon, SEAMP, Smithsonian Institution and Dr. Alan Stone, Agricultural Research Service, U. S. Department of Agriculture who reviewed the manuscript and offered many suggestions throughout the study. Lt. Colonel J. E. Scanlon, Walter Reed Army Institute of Research initiated the collections which led to these studies while at the South East Asia Treaty Organization Medical Research Laboratory, Bangkok. Personnel of the Department of Medical Entomology, SEATO Medical Research Laboratory, Bangkok, Thailand collected most of the material and prepared specimens for study. Included among these were: Dr. D. J. Gould, SFC E. L. Peyton, Mr. S. Esah, Mr. S. Maneewangse, Mr. S. Chumchulcherm, Mr. K. Mongkolpanya, Mrs. R. Rattanarithikul, Miss S. Sandhinand, and Miss P. Narawat. Technical assistance at SEAMP, Smithsonian Institution was provided by Miss C. C. Lipscomb who typed the manuscript and Miss J. L. Hwang who prepared the illustrations. Dr. P. F. Mattingly kindly permitted the first author to compare specimens with types in the British Museum (Natural History). LITERATURE CITED Belkin, J. N. 1962. The mosquitoes of the South Pacific. Univ. Calif. Press, Berkeley, 2 vols., 608 and 412 pp. Colless, D. H. 1965. The genus Culex, subgenus Lophoceraomyia, in Malaya (Diptera: Culicidae). Jour. Med. Ent. 2: 261-307. 18 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 ANOPHELES (ANOPHELES) TIGERTTI, A NEW SPECIES OF THE AITKENIT GROUP FROM THAILAND! Joun E, SCANLON? and E. L. PEyTon® During routine surveys of jungle mosquitoes in Southeastern Thai- land a small number of larvae and pupae of the Anopheles aitkenii group were found in water aspirated from crab holes along a small freshwater stream. Examination of the immature stages and the reared adults disclosed that this was a new species. The aitkenii group was most recently reviewed by Reid (1965), who listed seven valid species for Malaya and Borneo. According to the infrasubgeneric classification adopted by Reid and Knight (1961) the aitkenii species group belongs to the Anopheles series of the Angusticorn section of the subgenus Anopheles. The species of the aitkenii group are very similar in gross adult morphology, the adults being rather small, brownish, and with a somewhat culicine appear- ance. Identification is based chiefly on the male terminalia and larval and pupal chaetotaxy; the females of the group cannot be separated with certainty. Members of the group are largely restricted to hill forests in Southern and Southeast Asia. The adults are rarely taken in large numbers and their habits are very poorly known. The immature stages are collected more frequently and seem to favor small streams, seepages and springs under forest cover. There is practically nothing known of the biting habits of the females and nothing known of their possible role in the transmission of human or animal malaria. The most salient group characters outlined by Reid (1965) are as follows: very narrow upright forked occipital scales in the adults; wing scales uniformly brown in color; larva with branched antennal hair not over % the length of the antennal shaft; inner clypeal hairs rather far apart at the base for the subgenus Anopheles, saddle hair often branched. Reid (1965) also indicated that abdominal seta 5 in the pupa is considerably shorter than the abdominal segments, but the present species differs from other pupae of the aitkenii group in this respect. In the following description and accompanying illustrations the terminology and numbering of the setae are essentially those of Belkin (1962). This paper is number 165 from the Army research program on malaria. Supported in part by contract number DA-49-193-MD-2672. Immediate publica- tion secured by full payment of page changes—Editor. 2 Lieutenant Colonel, Medical Service Corps, US Army. Walter Reed Army Institute of Research, Washington, D. C. Southeast Asia Mosquito Project, Smithsonian Institution, Washington, D. C. ’ Master Sgt., US Army, Medical Component—Southeast Asia Treaty Organiza- tion, Bangkok, Thailand. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 19 Anopheles (Anopheles) tigertti, sp. n. (Figs. 1, 2) Female. A rather small (wing 2.9 mm) delicate species, uniformly brownish in color. No characters have been found which will permit its separation from other members of the aitkenii group. Head. Proboscis long, thin, uniformly dark, labellum dark brown; palpus similarly colored, slightly shorter than proboscis; antennal flagellomeres brown, without scales, but with well developed whorls of setae; vertex with a small number of forward projecting chetae and thin upright forked scales. Thorax. Propleural setae obscured in the single female available; wing with dark scales only, fringe unicolored, anterior fork-cell index 1.25, posterior fork-cell index 0.55; halter knob dark scaled, stem paler; legs without distinct markings, femora somewhat lighter ventrally than dorsally; scutum with well developed achrostichal and dorso-central bristles, a median dark line from the anterior promentory to the pre-scutellar space, single lateral lines on either side of the median line, from the anterior margin of the scutum to the scutellum; scutellar bristles strongly developed; anterior pronotal lobe with eight setae, sternopleuron with three setae, three upper mesepimeral setae. Abdomen. Dull olive brown above, slightly paler ventrally, heavily covered with fine setae; no distinctive markings. Male. Very similar in general appearance to female. Head. Proboscis long, thin, dark; palpus with elongated apical knob; antennal flagellomeres lighter in color than in the female, with more strongly developed setal tufts; scales of vertex similar to female. Terminalia. General structure as in figure 2. Basimere short, conical, with well developed setae; two basal spines tapered, curved apically; subapical spine curved; distimere evenly curved, with 5-6 short setae near the apex; dorsal lobe of the harpago divided, outer portion with two strong flattened setae, inner portion with a flattened spine having a club-like apex; ventral lobe of the harpago also divided, outer portion with two spines, inner rounded portion with one long seta and numerous smaller setae, tip of the phallosome without leaflets, body of the phallosome with several small spines on dorsal and ventral surfaces and near the apex. Larva. Chaetotaxy and general appearance as in figure 1. Head. Antenna with well developed spicules on basal two-thirds, smaller number on apical third, antennal hair (1—A) dorsal, 5-9 branches; hair 2-C divided into 3-4 branches about 7% from the base, bases of hairs 2-C separated by less than the distance be- tween 2-C and 3-C on either side; hair 4-C with 3-4 branches, inserted slightly external to the base of 2-C, hair 3-C divided into 2 branches about *4 from base. Thorax. Hair 1-P with 8-10 branches; 2-P longer, with a more strongly developed basal tubercle. One long hair of each meso- and meta-thoracic pleural group (9-12) 2 branched beyond the middle; metathoracic palmate hair (3-T) well developed, with flattened leaflets and terminal filaments. Abdomen. Hair 1-I poorly developed, filamentous hair 1-II with flattened leaflets, but without definite shoulders or terminal filaments; palmate hairs well developed on seg- ments III-VII hair 6-III with 6-12 branches; pecten with 14-18 teeth, fairly uniform in length, but occasionally with 1-2 shorter teeth; hair 1-X with 1-3 branches. Pupa. Chaetotaxy and general appearance as in figure 2. Cephalothorax. Trumpet tubular, with a deep cleft; hair 12-C with numerous branches. Abdomen. Hairs 1, 3 and 5 well developed on segments II to VII; hair 5 long on most PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 Dorsoventral view of the , enlarged view of placement of clypeal setae; B, enlarged Anopheles (Anopheles) tigertti sp. nov. fourth stage larva. A view of hair 1-IV lenyex IIe PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 21 segments particularly IV and V, reaching or exceeding the posterior margins of the succeeding segments; hair 9 long on segments IV—VIII. Type Data. Holotype male with associated larval and pupal skins and terminalia slide mounted with the following data; Thailand, Prahinburi Province, Ban Bu Phram, 150 m, 20 January 1966, Peyton and Kol collectors, deposited in the U. S. National Museum, No. 69239. Paratypes: 1 male with associated pupal skin and terminalia, 1 male with associated terminalia, one female and six larvae having the same data as the holotype to be deposited in the British Museum and the U. S. National Museum. The species is named for Colonel William A. Tigertt, Director of the Walter Reed Army Institute of Research who has done much to promote the study of mosquitoes and malaria in Southeast Asia in recent years. Distribution. In addition to the type locality (14°16’N., 101°53’ E.) the junior author has identified larval specimens from Khao Sai Dao, Chantanburi Province (12°51’N., 102°13’E.). This is a heavily forested mountain near the Cambodian border and approximately 100 miles southeast of the type locality. Taxonomic Discussion. The aitkenii species group forms a well defined and fairly homogeneous entity with the Anopheles series. The present species differs from other aitkenii group species in the male terminalia; most strikingly in having small spines or denticles on the surface of the phallosome. In A. insulaeflorum the phallosome has spicular processes, but these are limited to the sides of the phallosome toward the apex. The external portion of the dorsal lobe of the harpago has two spines, as in fragilis and bengalensis, rather than three as in aitkenii, insulaeflorum and _stricklandi. The larvae of tigertti have relatively few branches (6-12) on hair 6-III, and in this respect resemble insulaeflorum and stricklandi more than they do the other members of the group, in which this hair has 20-50 branches. In both insulaeflorum and _ stricklandi, however, the inner clypeal hairs (2-C) are simple and have their bases set very close together, while these hairs are multiple and inserted somewhat apart in tigertti. Among other species of the aitkenii group: pinjeurensis Barraud is known from a single male which has a remarkably elongated phal- losome without spicules. A. acaci Baisas and bornsensis McArthur have larvae with extensive dichotomous branching of hair 2-C. The larvae of palmatus have greatly enlarged tergal plates and_ short frontal hairs (5-7-C). Biology. The finding of the immature stages in holes bored by land crabs may have been fortuitous in that the larvae might have been stranded in the overflowing and subsequent drying of a nearby stream. However, two factors rule against this—the finding of larvae of tigertti in a similar niche some 100 miles to the southeast of the type locality bo 2; PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Fig. 2. Anopheles (Anopheles) tigertti sp. nov. Dorsoventral view of pupa and dorsal view of male terminalia. A, apical portion of harpago; B, phallosome. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 23 and the failure to collect larvae from the nearby stream. A significant number of mosquito species are known to inhabit crab holes, including species of the genus Deinocerites Theobald 1901 in the New World, and a number of forms in SE Asia, including Uranotaenia atra Theobald, 1905 Aedes (Rhinoskusea) longirostris (Leicester) 1908, and several species of Aedes (Cancraedes) Edwards 1929. Most of these, however, involve marine crabs and littoral areas. Terrestrial crabs are fairly common in the evergreen forests of Southeast Asia and it is possible that further collecting will show their burrows to be a prolific source of mosquitoes. The adult habits of tigertti are entirely unknown since all of the adults examined were reared from immature stages. Adults of the aitkenii group are rather rarely collected, usually being found resting in the jungle. There are reports of aitkenii group females attacking man (Christophers, 1933; Scanlon and Esah, 1965; Macdonald and Traub, 1960) and Reid (1965) specifically lists bengalensis as biting man in Malaya. However, the usual impossibility of distinguishing the fe- males of the group and their relative rarity in inhabited areas in SE Asia makes it difficult to make very definite statements in this regard. There seems to be little reason to believe that species of the aitkenii group play any role in the transmission of human malaria. The paucity of human biting records in jungle areas where extensive collections have been made probably indicates that females of the group usually feed on some forest vertebrate other than man. ACKNOWLEDGMENTS The specimens of A. tigertti discussed here were collected by Mr. Somboon Maneechai and Mr. Kol Mongkolpanya, under the direction of the junior author and Dr. Douglas J. Gould, Department of Medical Entomology, U. S. Army Medical Component—SEATO. Specimens were examined by Dr. John Reid, British Museum, and Drs. Alan Stone and Botha de Meillon of the SEAMP. The figures were prepared by Miss Jung Lea Hwang. REFERENCES Belkin, J. N. 1962. The mosquitoes of the South Pacific. Univ. Calif. Press, Berkeley. 2 vols., 608 and 412 pp. Christophers, S. R. 1933. The fauna of British India. Diptera. Vol. IV. Anophelini. Taylor and Francis, London, 371 pp. Macdonald, W. W. and R. Traub. 1960. An introduction to the ecology of the mosquitoes of the lowland dipterocarp forest of Selangor, Malaya. Stud. Inst. Med. Res. Malaya, No. 29: 79-109. Reid, J. A. 1965. A revision of the Anopheles aitkenii group in Malaya and Borneo. Ann. Trop. Med. Parasitol. 59(1): 106-125. , and K. L. Knight. 1961. Classification within the subgenus Anopheles (Diptera: Culicidae). Ann. Trop. Med. Parasitol. 55(4): 474- 488. Seanlon, J. E. and S. Eash. 1965. Distribution in altitude of mosquitoes in Northern Thailand. Mosq. News 25(2): 137-144. 24 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 NOTES ON NORTH AMERICAN SALDIDAE ( HEMIPTERA ) Joun T. Potnemus, 3115 So. York St., Englewood, Colorado The following notes affected the synonymy of two species of the genus Saldula and contain the descriptions of two new species of the genus Salda, one of which has long been confused with Salda littoralis (L.). Iam indebted to R. H. Cobben who first noted one of the new species and one of the errors in synonymy. Saldula severini Harris 1943 Saldula severini Harris 1943, J. Kans. Ent. Soc. 16(4): 152. Drake and Hottes (1954) synonymized Saldula severini with S. orbiculata (Uhler) on the basis of the pronotal margins, which were held to fall within the specific range of variation of the latter species. That this synonymy is wrong was first noticed by R. H. Cobben who suggested that I publish this information. Subsequent investigation by the author has revealed that the male parameres are very unlike, those of orbiculata (fig. 1) having a definite mammillate process caudad while severini lacks this (fig. 2). Also, severini can be separated from orbiculata by the length of the hairs of the hind tibia which are sub- equal to the width of the tibia in the former, and twice or more the width of the tibia in the latter. Saldula severini Harris is therefore restored to specific status. Saldula separata (Uhler) 1878 Salda separata Uhler 1878, Proc. Boston Soc. Nat. Hist. 19: 432-433. Saldula illinoiensis Drake 1949, Ark. for Zool. 42B(3): 2-3 (New Synonymy ) Drake and Hoberlandt in 1951 synonymized Saldula separata (Uhler) with S. pallipes Fabricius without giving a motivation. Prob- ably they were following the lead of Torre-Bueno (1924) who noted that Acanthia interstitialis Say is “known under two further names- pallipes Fabricius and separata Uhler.” Blatchley (1926) considered separata a good species and used the long black hairs on the dorsum as a key character. Uhler’s original description mentioned the “erect pubescence” several times, and I have long been convinced that separata is conspecific with Saldula illinoiensis Drake. A comparison of specimens of the latter species with the type of separata shows this to be true; therefore Saldula separata Uhler is restored to specific status, and Saldula illi- noiensis Drake is made a synonym thereof. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 5 Figs. 1-7. 1, Left male paramere of Saldula orbiculata Uhler. Two views A&B; 2, Left male paramere of Saldula severini Harris. Two views A & B; os 4, 5 and 6. Salda coloradensis n. sp.; 3, Left male paramere. Two views A & B; 4, Parandria; 5, Penis; 6, Median sclerotized structure of penis; 7, Median sclerotized structure of penis, Salda buenoi McDunnough; (Scale for all figures except 5, shown at bottom of Plate) 26 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 a os 4/5 * % > 1 | om Figs. 8-14. 8, 9, 10, and 11. Salda alta n. sp. 8, Left male paramere; 9, Penis; 10, Median sclerotized structure of penis; 11, Parandria; 12, Pronotum, Salda coloradensis, n. sp.; 13, Pronotum, Salda buenoi, McDunnough; 14, Pronotum, Salda alta, n. sp. The type of Saldula separata, in the Harris collection of the Museum of Comparative Zoology at Harvard, was compared by Dr. P. J. Darlington, Jr. and I am indebted to him for his labors in searching the collections, making the comparison, and furnishing the following information from Harris’s notebook. The type bears a number 77( 138 ) and under 77 in the notebook is the following: “Leptopus- Acanthia alternata, S.- Mss? Dr. Green. On salt marsh? May 15, 1837.” This | : ) PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 iT notation matches Uhler’s statement that the specimen was labelled with a number 77. Dr. Darlington notes that “..... the type is hardly in first class condition, for it is pinned through the scutellum in such a way as to spread the specimen slightly, and it is somewhat rubbed, so that the pubescence is partly missing... . . Salda coloradensis, n. sp. Large, elongate, general color black, macropterous. Head.—Black, very faintly shining, rugulose; preocellar spot yellowish white; covered with appressed golden pubescence and the usual three pairs of long hairs on frons and vertex, longer pubescence and scattered long hairs on labrum and anteclypeus; vertex slightly carinate between ocelli and eye on each side, the carina produced anteriorly around the front margin of the eyes; ocelli raised slightly, separated by the width of an ocellus; mandibular plates, maxillary plates, anteclypeus and center portion of labrum ochreus; rostrum blackish brown, be- coming lighter apically, extending past hind coxae. Thorax.—Pronotum black, rugulose, covered with thick appressed golden pubescence, feebly shining, narrow; lateral margins straight, narrowed somewhat anteriorly (100:67),! almost meeting the collar on a straight line; callus feebly to moderately raised, with small shallow impression in center; posterior lobe shorter than anterior lobe (13:30). Underparts black, covered with fine yellowish white pubescence; acetabulae broadly margined with ochreoleucus. Scutellum as wide (70) as long (68), black, rugose; vestiture as on pronotum. Wings.—Hemelytra fully developed; vestiture as on pronotum except for the cells of the membrane, the thick pubescence hiding the dorsal surface; black, rugulose and minutely pebbly surface, faintly shining; membrane deep fumose, opaque, minutely pebbly on the surface, with four cells. Extremities —Antennal segments 1 and 2 ochreus, segment 1 brown beneath on basal two thirds, segment 2 dark brown on apical tenth, stout; segments 3 and 4 dark brown, as thick or thicker than segment 1; all segments clothed with fine yellowish pubescence, segment 1 with a few black spines and hairs, the other segments with scattered long hairs; antennal proportions: ¢; segment 1, 25; segment 2, 70; segment 3, 50; segment 4, 53. 9; segment 1, 25; segment 2, 73; segment 3, 48; segment 4, 50. Legs ochreus; femora with usual browish spots; apex of tibia and second tarsal segment, third tarsal segment brownish black; coxae brownish black, margined basally and apically with ochreus; usual dark spines on tibia. Genital Structures.—Paramere, parandria, penis and median sclerotized struc- ture of the penis as shown in figures 3, 4, 5 and 6 respectively. Penis filum coiled three and one half times. Body Measurements.—Holotype ( 2), length 5.3 mm., width 2.3 mm. Allotype (2), length 5.5 mm., width 2.5 mm. Mean length of 106: 4.95 mm. (max. 5.5; min. 4.6). Mean width of 10¢: 2.16 mm. (max. 2.5; min. 2.0). Mean length of 10¢: 5.44 mm. (max. 5.9; min. 5.0). Mean width of 109: 2.42 mm. (max. 2.6; min. 2.2). 1 For all measurements, 60 units = Imm. 28 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Material—Holotype (¢), Sedalia, Colorado VI-29-1961, CL.56, J. T. Polhemus. Allotype (2), same data as holotype. Paratypes, all collected by J. T. Polhemus, as follows: 1 specimen, Sedalia, Colo- rado, CL. 56, VI-26-1961; 4 specimens, Sedalia, Colorado, CL. 56, VI- 29-1961; 1 specimen, Sedalia, Colorado, CL. 56, VII-1-1961; 30 speci- mens, Sedalia, Colorado, CL. 56, VII-18-1962; 8 specimens, Granby, Colorado, CL. 55, VI-24-1961; 8 specimens, 1 nymph, Hartsel, Colorado, CL. 120, VI-16-1962; 14 specimens, Hartsel, Colorado, CL. 120, VIII- 18-1962; 6 specimens, Silver Plume, Colorado, CL. 44, VIII-6-1961; 1 specimen, Silver Plume, Colorado, CL. 44, VIII-20-1961; 1 specimen, Sargents, Colorado, CL. 208, VIII-3-1963; 2 specimens, Georgetown, Colorado, CL. 44, VII-21-1962; 1 specimen, Georgetown, Colorado, CL. 36, VITI-6-1961; 3 specimens, Saguache, Colorado, CL. 149, VIII-19- 1962; 1 specimen, Deckers, Colorado, CL. 178, V-16-1963; 1 specimen, 2 nymphs, Tabermash, Colorado, CL. 129, VI-23-1962. The holotype, allotype and paratypes are in the collection of the author. Paratypes will be sent to the U. S. National Museum, California Academy of Science and University of Kansas. Habitat—Salda coloradensis inhabits the damp margins of ponds and damp meadows in the foothills and parks of the Rocky Mountains. It is quite local in both time and place, but is sometimes found in numbers, associating at times with Salda bouchervillei Provancher and Salda buenoi McDunnough. This species prefers sparse grassy cover, and may range onto almost dry soil. Near Granby, Colorado, speci- mens were taken in a marsh on granite boulders by pulling away the dense grass cover at the base, thus exposing the saldids resting on the vertical stone surface. Comparative Notes—Salda coloradensis is most closely allied to S. littoralis (L.) and S. buenoi. Concerning the latter, S. buenoi is usually more robust than coloradensis, and has more rounded lateral pronotal margins, especially the antero-lateral angles (Plate I, figures 12 and 13) and darker antenna. However the two are found together at Hartsel, Colorado and are virtually indistinguishable except on the basis of genitalia. The median sclerotized structure of the penis and parameres taken together will serve to separate the adults in difficult situations, as the sclerotized plates of the penis structure are somewhat flattened in buenoi (fig. 7) and vertical in coloradensis (fig. 6). The paramere of S. buenoi has been figured by Drake and Hottes (1951). The median sclerotized structure of the aedeagus of Salda littoralis (L.), figured by Cobben (1957), and coloradensis are quite different, and the two probably do not inhabit the same range. S. littoralis occurs across northern Europe, Siberia, and into Alaska. Drake and Hottes (1951) record it from Canada and the U. S., however it remains to be determined how far south it has invaded, as many of the existing PROG. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 29 records must be discarded as misidentifications, referring instead to coloradensis. Another closely allied species, Salda nevadensis Wagner (1960); has been studied. It was found to be very similar to S. littoralis, the genitalic structures being virtually the same, and is so far known only from Spain. Salda coloradensis is the only saldid known to me that is more easily separated from its congeners in the nymphal state than in the adult state. The nymphs are covered with hairs subequal in length to the width of the hind femora, while nymphs of buenoi and littoralis have only a few insignificant hairs. R. H. Cobben first noticed that this species is not identical to Salda littoralis (1.). Salda alta, n. sp. Large, elongate, general color black, macropterous. Head.—Black, barely shining, rugulose; preocellar spot yellowish white; generally covered with short, inconspicuous golden pubescence, longer near eyes, on labrum and anteclypeus; the usual three pairs of long hairs on frons and vertex; vertex depressed between eye and ocelli on each side, the depression produced anteriorly as a carina which continues around anterior margin of eyes; ocelli raised slightly, separated by more than the width of an ocellus; mandibular plates, maxillary plates, center of labrum and lower portion of anteclypeus ochreoleucus; rostrum brown, extending between hind coxae. Thorax.—Pronotum black, rugulose, covered with short, fine golden pubescence, feebly shining; lateral margins straight, narrowed moderately anteriorly (137:71), anterior angle curved moderately to meet collar (Plate II, fig. 14); callus moderately raised, with shallow impression in center; posterior lobe shorter than anterior lobe (15:35). Underparts black, covered with fine golden pubescence; acetabulae margined with ochreoleucus. Scutellum as wide as long (30:30), black, rugose; vestiture as on pronotum. Wings.—Hemelytra fully developed, black; with four yellowish spots along outer margin of inner corium, about equally spaced, the first one-fourth of the distance from base to apex, the last next to the membrane; one yellowish spot in center of outer corium, next to inner corium, and a similar spot near the apex of the clavus; vestiture as on pronotum, inner veins of membrane largely free of pubescence; rugulose, faintly shining, except membrane which is relatively smooth and dull; membrane fumose to light sub hyaline smoky brown; darker spot in center of each cell well defined; with four cells. Extremities—Antenna brown, dorsal surface of first segment ochreoleucus, long, slender; all segments clothed with brown pubescence, segment 1 with scattered dark spines, remaining segments with scattered long hairs; antennal proportions: 6; segment 1, 33; segment 2, 80; segment 3, 50; segment 4, 52. 9; segment 1, 33; segment 2, 83; segment 3, 55; segment 4, 52. Legs ochreus; femora brown beneath, with usual brown spots; apex of tibia and dorsum of third tarsal segment brown; coxae brown to blackish brown, margined with luteus; usual dark spines on tibia. Genital Structures——Paramere, penis, median sclerotized structure of penis and 30 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 parandria as shown on Plate II, figures 8, 9, 10 and 11 respectively. Penis filum coiled three and one half times. Body Measurements.—Holotype ( ¢), length 6.9 mm., width 3.1 mm. Allotype (2 ), length 7.4 mm., width 3.5 mm. Paratype ( ¢ ), length 6.9 mm., width 3.0 mm. Mean length of 59: 7.47 mm. (max. 7.9; min. 6.9). Mean width of 59: 3.53 mm. (max. 3.7; min. 3.4). Material—Holotype (é) Goose Lake, south of Creede, Colorado, Longitude 107°, Latitude 37°35’, Elevation 11,700 ft., CL. 291, VII-19- 1964, J. T. Polhemus. Allotype (2), and paratypes, one male, four females, three nymphs, same data as holotype. All material is in the collection of the author. Habitat.—Salda alta n. sp. was found on the shore of Goose Lake, which lies in a small depression on a bench at 11,700 ft. in the San Juan Range. The habitat is a typical, wet, high montane meadow with tussocks of grass, black spongy soil, and is ice free only a few months of the year. Only low brush and grass was present in the meadow where alta was found, however a good stand of timber was present on the opposite shore of the lake. The location has a southeastern ex- posure, with a granite wall rising to the west. Comparative Notes.—Salda alta most closely resembles Salda lugubris (Say) in the hemelytral markings and size, however the latter species has the lateral pronotal margins rounded, the first cell of the membrane does not project forwardly as far as in alta, and the genitalia (see Drake and Hottes 1951) are different. In the type series of alta the hemelytral markings vary considerably, being prominent in some speci- mens and showing additional spots at the middle apex and inner apex of the inner corium, and near the apex of the embolium. The type is the least marked of the series. REFERENCES Blatchley, W. S. 1926. Heteroptera of Eastern North America, Indianapolis, 1116 pp. Cobben, R. H. 1957. Beitrag Zur Kenntnis Der Uferwanzen (Hem. Het. Fam. Saldidae), Ent. Berichten, 17: 245-257. Drake, C. J. and F. C. Hottes. 1951. Saldidae of the Americas (Hemiptera), Great Basin Nat. 10(1-4): 51-61. 1954. Synonymic data and description of a new saldid (Hemip- tera). Occ. Papers, Mus. Zool. Univ. Michigan 553: 1-2. Drake, C. J. and L. Hoberlandt. 1951. Catalogue of Genera and species of Saldidae (Hemiptera). Acta Ent. Mus. Nat. Pragae 26(376): 9. Torre-Bueno, J. R. 1924. Three Canadian Acanthiidae (Saldidae, Heterop- tera), Canadian Ent. 56: 296-300. Wagner, E. 1960. Salda nevadensis nov. spec. (Hem. Het. Saldidae), Trabajos Museo Zool., Barcelona, Nueva Serie Zool. 2(3): 2-4. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 31 NEW NEOTROPICAL NEOCOELIDIINAE WITH KEYS TO THE SPECIES OF COELIDIANA, XENOCOELIDIA, AND NELIDINA (HoMoprTERA: CICADELLIDAE ) JAMEs P. KraMer Entomology Research Division ARS U.S. Department of Agriculture Washington, D. C. As far as known, the long-horned leafhoppers or Neocoelidiinae are unique to the Americas. Recently I published a generic revision of the group (Kramer 1964). In spite of a combination of distinctive features which at once set them apart from all other subfamilies of leafhoppers, Linnavuori (1965: 142) reduced the group to a tribe of the Delto- cephalinae on the basis of one character, the bifurcate anterior branches of the tentorium. This proposed change has slight merit in my opinion. It would reduce one of the most distinctive assemblages of leafhoppers to a tribe of the already excessively large and unwieldy Deltocephalinae. My treatment, as a subfamily, agrees with both Evans (1947: 196) and Oman (1949: 59). Included in this paper are keys to the species of the following genera: Coelidiana Oman (four n. spp.), Xenocoelidia Kramer (two n. spp.), and Nelidina DeLong (one n. sp.). Two new genera, Chin- chinota (one n. sp.) and Coelindroma (two n. spp.), are described. Single new species are described in Neocoelidia Gillette & Baker, Biza Walker, and Tozzita Kramer. Two species names are reduced to synonymy. Coelidiana OMAN For a generic description of Coelidiana, see my earlier paper (Kramer 1964: 274-5). The following key includes all known species except those three based on unique female types. The three species in question, coronata (Ball), distinctis- sima Linnavuori, and lurida Linnavuori, are actually of uncertain generic place- ment at this time. KEY TO THE SPECIES OF Coelidiana OMAN (Males ) 1. Aedeagus with one or two long basal processes _____.__-___________. 2 memerons. wath no basal processes. 22.2.2 2. 4 2. Aedeagus with a pair of basal processes (fig. 2) patrator, n. sp. Never pusewith single) basal process 1. a 3 3. Pygofer with tooth on ventral margin at apex Sig ania? & Heller 1961: pl. 2, fig. 2), apex of aedeagus with a moderate sized, posteriorly directed, beak (ibid. pl. 2, fig. 4) flavida Linnavuori & Heller Pygofer with tooth on ventral margin much basad of apex (Linnavuori 1965: fig. 4a), apex of aedeagus with a tiny anteriorly directed beak (ibid. fig. ON gy LES ena eee brasiliensis Linnavuori 4. Pygofer terminating dorsally with a sharp point of moderate length and Matha, short tooth on ventral margin (fig. S$) ... 5 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 32 DURATA 3 PATRATOR PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Se) Byporers variable. put mot -as aboves <2. ..2- 2.285 8 Pe i et 7 5. Aedeagal shaft crenulated ventrally (Kramer 1964: fig. 95) Ba Si, es I a Be OE ees eg ee LL eae eroceata (Osborn) Aedeagal shaft smooth ventrally (fig. 3) _-..- paaiees ae. 6 6. Aedeagal shaft not upturned near base, extreme apex usually bluntly rounded WhKaramaer MlOG4s fis. 99) 2 2 Ne rubrolineata (Baker ) Aedeagal shaft abruptly upturned near base, extreme apex not bluntly (CLERC ICH (Ghee 5) Aa eee Eee Oe eect ea vee fl. SeeeeUE durata, n. sp. Weeetiaietmbe with’ processes) 8 8 PUMAUALMOe LWVItNOUL PTOCessés: _LAle sy ee ee 11 8. Processes of anal tube long and conspicuous (figs. 4 and 6) — 9 Processes of anal tube short and inconspicuous —... =. 10 9. Pygofer with a stout decurved process on hind margin, first segment of anal fubesaboutas long as wider(fig. 4) ancora, n. sp. Pygofer without a stout decurved process on hind margin, first segment of anal tube much longer than wide (fig. 6) —. colens, n. sp. 10. Ventral margin of pygofer with two teeth, aedeagus simple (DeLong 1953: [Blk tejae Tabata") Pe ee ta SOA AE Ee em aaa ee IAS ace taebeott bidentata DeLong Ventral margin of pygofer with one tooth, aedeagus with a pair of short triangular processes near apex (Linnavuori 1965: fig. 4e) perme ET De sone Ne Ps ee ata ee oP Pg eee SS signaticeps Linnavuori 11. Dorsal margin of pygofer terminating with a long slender spine, without aecnOit toot, Ou wentral, margin... 2-9. tl Se 12 Dorsal margin of pygofer not terminating with a long slender spine, with a short tooth on ventral margin etre et". i eee ee ©) - a3 12. Both dorsal and ventral margins of pygofer terminating with slender spines Q@elong, 1053: pl Ss uftg ci), set te bimaculata (Baker) Only dorsal margin terminating with a slender spine (DeLong 1953: pl. 8, LTE, JG) ES Ds i ee i spina DeLong 13. Aedeagus comparatively slender, distal portion of style slender and necklike, plates longer than pygofer (fig. 7) undata (Linnavuori ) Aedeagus comparatively stouter, distal portion of style not as above, plates ONE EN ale DV OOLGR: fac) ee ee ee ee 14 14. Aedeagal shaft gradually upturned distally, posterior margin of pygofer manecate (Delong 1953: pl. 8, fig. 5) unipuneta (DeLong) Aedeagal shaft more abruptly upturned distally, posterior margin of pygofer rounded (Delong 1953: “pl.°8, fig. 6) 2 anomala (DeLong) Coelidiana patrator, n. sp. (Figs. 1-2) Length.—Males 6-6.25 mm. Structure —Crown broadly angular at apex; interocular width one and a half times median length. << Figs. 1-7. Coelidiana patrator, n. sp. Fig. 1, male genital capsule laterally; fig. 2, aedeagus posteriorly. Coelidiana durata, n. sp. Fig. 3, male genital capsule laterally. Coelidiana ancora, n. sp. Fig. 4, male genital capsule laterally. Coelidiana colens, n. sp. Fig. 5, head and thorax dorsally; fig. 6, male genital foe laterally. Coelidiana undata (Linnavuori). Fig. 7, male genital capsule aterally. 34 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 CHARISMA Bete sh =) x 44-94-44" oe, No LITURA 14 Figs. 8-17. Xenocoelidia charisma, n. sp. Fig. 8, male genital capsule lat- erally; fig. 9, subapical process of aedeagus extended in posterior view; fig. 10, aedeagal apex posteriorly; fig. 11, head and thorax dorsally; fig. 12, ventral cover of male genital capsule; fig. 13, right style ventrally. Xenocoelidia litura, n. sp. Fig. 14, male genital capsule laterally; fig. 15, ventral cover of male genital capsule; fig. 16, aedeagal apex dorsally; fig. 17, right style ventrolaterally. Coloration.—Ground color stramineous to yellow; legs and thoracic venter palest; face, crown, pronotum, and scutellum yellow; following portions stramineous to white: a pair of longitudinal oval areas flanking midline on crown from posterior margin to beyond eyes, narrow area on posterior margin of pronotum, a pair of longitudinal oval areas on scutellum flanking midline and narrow short area near base on each lateral margin; forewings pale milky hyaline. Male genitalia.—Genital capsule laterally (fig. 1) with short acute process at apex of dorsal margin, ventral margin thickened, terminating with a distinct tooth, stylar apex broadly hooked, aedeagus simple and gradually recurved distally with a pair of long simple processes near base, processes seen best in posterior view of aedeagus (fig. 2). Genital capsule ventrally with plates fused except for short distance at apex. Female genitalia—Female unknown. Types.—Holotype male (USNM Type No. 68934) and paratype male, Tingo Maria, Peru, 12 August 1960, O. Vargas. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 oD Notes.—The paired basal processes of the aedeagus set patrator apart at once from all other species of Coelidiana. Coelidiana durata, n. sp. (Fig. 3) Length.—Males 5.5-6 mm. Female 6 mm. Structure —Crown bluntly angular at apex; interocular width slightly more than median length. Coloration.—Identical to that of rubrolineata (Baker), see DeLong 1953: 127. Male genitalia —Nearly identical to that of rubrolineata (DeLong 1953: pl. 8, fig. 1) except for the style and aedeagus (fig. 3), style with distal portion broadly hooked and flattened dorsally, aedeagus moderately stout basally with shaft narrowed and abruptly upturned. Female genitalia —Posterior margin of pregenital sternum nearly truncate with a slight notch at middle. Types.—Holotype male (USNM Type No. 68935), Abancay, Peru, 13 July 1960, Vargas and Salazar, on Paspalum; allotype female and one paratype male with same data except 11 July 1960, S. Vargas, on barley; and one paratype male with same data except 8 July 1960, Young and Salazar, on alfalfa. Notes.—C. durata is very similar to rubrolineata, but it can be readily distinguished from it on the basis of the aedeagus as noted in the key to species. Coelidiana ancora, n. sp. (Fig. 4) Length.—Male 6.75 mm. Structure —Crown broadly angular at apex; interocular width slightly less than one and a half times median length. Coloration.—Ground color stramineous to yellow; legs and thoracic venter palest; face and crown stramineous, midline of crown faintly bright yellow; anterior portion of pronotum stramineous with rest bright yellow; scutellum stramineous with apex bright yellow; forewings yellowish hyaline fading to colorless hyaline broadly on costal margins. Male genitalia—Genital capsule laterally (fig. 4) with a long, irregular, decurved, stout process on posterior margin; a long distally scaly process from first segment of anal tube (Note: Both of these processes are paired but only one of each is shown in fig. 4); stylar apex strongly hooked; aedeagus stout but simple with shaft decurved and then recurved just before apex; gonopore apical. Genital capsule ventrally with plates fused except for short apical separation. Female genitalia—Female unknown. Type.—Holotype male (USNM Type No. 68936), Huanuco, Peru, Huallaga River Valley, tropical jungle, March 1954, Felix L. Woytkow- ski. Notes.—The shapes of the processes of the anal tube and pygofer provide the most obvious unique features of ancora. 36 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 18 SPORTULA 22 Figs. 18-24. Tozzita petulans, n. sp. Fig. 18, male genital capsule laterally; fig. 19, head and thorax dorsally; fig. 20, ventral cover of male genital capsule; fig. 21, aedeagal apex posteriorly. Nelidina sportula, n. sp. Fig. 22, male genital capsule laterally; fig. 23, ventral cover of male genital capsule; fig. 24, aedeagal apex posteriorly. Coelidiana colens, n. sp. (Figs. 5-6) Length—Males 5.5 mm. Females 6-6.5 mm. Structure —Crown (fig. 5) angular at apex; interocular width slightly less than median length. Coloration —Somewhat variable, ground color stramineous to bright yellow; venter, including legs and face, usually paler; crown and thoracic dorsum marked with three irregular longitudinal orange to red stripes (fig. 5), the unmarked areas PROG. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 on at times washed with pale green; forewings variable from milky hyaline to yellow hyaline and in females with commissural margins variably orange to red for entire length of clavus. Male genitalia —Genital capsule laterally (fig. 6) with a sclerotized band on pygofer from anterior dorsal angle to posterior ventral angle where it broadens along posterior margin; first segment of anal tube with a pair of long processes, processes decurve distally and terminate in sharp points; style distally with a sharp dorsal bend and a small apical hook; aedeagus with shaft gradually narrowed and upturned distally, extreme apex sharply narrowed, gonopore apical. Genital capsule ventrally with plates fused except at apex. Female genitalia.—Posterior margin of pregenital sternum produced and broadly wedge-shaped. Types——Holotype male (USNM Type No. 68937), allotype female, eight paratypes, six males and two females, Iquitos, Peru, 4 August 1960, J. Gonzalez, on pascae; one paratype male (California Academy of Sciences ), 67 miles east of Tingo Maria, Yurac, Peru, 11 December 1954, E. L. Schlinger and E. S. Ross. Notes.—The sclerotized band on the pygofer and the long pair of processes from the anal tube provide the distinguishing features of colens. Xenocoelidia KRAMER For a generic description of Xenocoelidia, see my earlier paper (Kramer 1964: 265). Interestingly enough, females of this genus have not yet been described. KEY TO THE SPECIES OF Xenocoelidia KRAMER (Males ) 1. Crown and pronotum without markings, dorsal ground color ivory white; apex of aedeagus in lateral view with a sharp tooth and a broad acute bladelike dorsal expansion (Kramer 1959: pl. Il, fig. 8) (Colombia) ceo) in A A SPOS 9 OD Dae he ED) Wee ee EE EOE seed pee ae youngi Kramer Either or both crown and pronotum with markings, dorsal ground color not imenanwhite: aedeagus) TG as. aboves 2. Ses a) ed 2 2. Head marked with two pairs of orange spots, pronotum with a pair of orange-margined black spots near anterior margin; apex of aedeagus in lateral view with a long crooked proximal process and a short slender distal process at apex (Kramer 1964: fig. 42) (Brazil) inflata (Osborn ) Head marked with orange but not as two pairs of spots, pronotum without black markings: aedeagus not as*above: 224 3 3. Central portion of pronotum without longitudinal stripes, anterior tibia with longitudinal brown stripe on outer edge; pygofer simple (Kramer 1959: eweleetiontly)> 1CColomintayir sl ts te 10 soy colombiana Kramer Central portion of pronotum with longitudinal stripes, anterior tibia uni- colorous; pygofer with a process or projection on dorsal or ventral margins SHEE eLipretiee oe, obt.cbie ss. Tt yn ven ue mebos. 2 (és wero nee) 2 ie te 4 4. Dorsal markings vivid; pygofer with process only at apex of ventral margin, aedeagal shaft with long sharp projection in basal half on proximal margin (fig. 8) (Peru) ab josh, Diy reine bs ated si abet ewaat a Ly, charisma, n. sp. 38 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Dorsal markings pale; pygofer with process or projection at apex of both dorsal and ventral margins, aedeagal shaft without a projection in basal half on proximal margin (fig. 14) (Peru) _........_.._..._ litura, n: sp. Xenocoelidia charisma, n. sp. (Figs. 8-13) Length.—Males 6.5—7 mm. Structure.—Head distinctly narrower than pronotum; crown longer at middle than next to eyes, anterior margin rounded (fig. 11). Coloration.—Venter, including legs and face, stramineous, at times lightly washed with pale green; antennae and a pair of poorly delimited spots below ocelli orange; ground color of crown, pronotum, and scutellum pale blue-green, marked (fig. 11) with orange-red as an inverted U-shaped band between eyes and as four broad longitudinal stripes on pronotum, scutellum marked mesally with orange-red and on each side with yellow to greenish yellow; forewing with ground color hyaline to somewhat milky, commissural margin fairly broadly and regularly brown to red-brown from scutellar apex to tip of wing, claval vein and area on either side of it variably washed with yellow to greenish yellow. Male genitalia.—Genital capsule laterally (fig. 8) with a short, slender, simple, and at times upturned, process at apex of ventral margin; hind margin of pygofer often turned in with anteriorly directed setae; style hooked apically with large preapical lobe; aedeagus a modified U-shape with paired processes at apex and single process or projection on proximal margin of shaft. Genital capsule ventrally (fig. 12) with a few setae on middle third. Style ventrally (fig. 13) with typical mesal bend and lateral lobe. Terminus of aedeagus posteriorly (fig. 10) with gonopore near apex. Female genitalia—Female unknown. Types.—Holotype male (USNM Type No. 68938), Tingo Maria, Peru, 12 August 1960, O. Vargas; four paratype males with same data except two each collected by M. Rojas and D. A. Young. Notes——The aedeagus of charisma is unique in the genus; the coloration is also distinctive and will help to identify this species as noted in the key. Xenocoelidia litura, n. sp. (Figs. 14-17) Length.—Males 6-6.4 mm. Structure —Head distinctly narrower than pronotum, crown longer at middle than next to eyes, anterior margin narrowly rounded. Coloration.—Venter, including legs and face, stramineous; antennal scape orange, rest brownish; ground color of crown, pronotum, and scutellum sordid white to stramineous; crown with a pair of longitudinal oval orange spots, one next to each eye; pronotum and scutellum marked much like those of charisma except the longitudinal orange stripes are narrower; forewing with ground color hyaline to somewhat milky, commissural margin marked like that of charisma except with orange instead of brown. Male genitalia—Genital capsule laterally (fig. 14) with a short toothlike projection at apex of ventral margin and a longer slender upright process at apex PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 39 of dorsal margin, hind margin setose, style strongly hooked apically and with a large preapical lobe, aedeagus modified U-shape with gonopore apical and three pairs of anteriorly directed processes (fig. 16) on distal third. Genital capsule ventrally (fig. 15) with a few setae on middle third. Style ventrolaterally (fig. 17) with typical mesal bend. Female genitalia—Female unknown. Types.—Holotype male and one paratype male (without head), Yurac, Peru, 67 miles east of Tingo Maria, 4 October 1954, E. L. Schlinger and E. S. Ross, in collection of California Academy of Sciences, San Francisco, California. Notes.—X. litura looks much like a pale or teneral form of charisma, but the many differences in the male genitalia leave no doubt that it is a very distinct species. Tozzita KRAMER The new species described below belongs in Tozzita, but the generic definition (Kramer 1964: 267) will have to be modified to include species with both asymmetrical and symmetrical aedeagi. Couplet 7 in my key to the genera of the genera of the Neocoelidiinae (ibid. 261) will be changed as follows: 7. Style well sclerotized, not wrinkled, and much shorter than aedeagus (fig. 47); aedeagus with apical elaborations (fig. 46) _.____________ Tozzita Style poorly sclerotized, wrinkled, and as long as aedeagus (fig. 60); aedeagus without apical elaborations (fig. 58) ss Xigqillliba Note: The figures cited above refer to illustrations which appeared in my 1964 paper. Tozzita petulans, n. sp. (Figs. 18-21) Length—Males 6.5-6.75 mm. Structure —Typical of genus in all respects. Coloration.—Venter including legs and face stramineous; abdominal segments brownish and only stramineous along posterior margins; at times with faint touches of pink on thoracic and facial sutures, and on scape of antenna; a distinct black spot at middle of extreme upper edge of face just below but touching transverse carina; ground color of crown, pronotum, and scutellum stramineous; sometimes vaguely washed with orange; marked with brown on their midlines and with orange laterally on scutellum as in figure 19; forewing mainly colorless hyaline but with a few touches of pale brown or milky hyaline areas, entire costal vein and area of forewing adjacent to scutellum washed with pink. Male genitalia—Genital capsule laterally (fig. 18) with a blunt projection at apex of pygofer, first segment of anal tube flanked by a pair of large rather foot- shaped processes (only one is visible in lateral view), connective short, style weakly hooked apically, symmetrical aedeagus slender and upcurved distally. Genital capsule ventrally (fig. 20) with plates entirely fused but suture visible on slender distal portion. Aedeagal apex posteriorly (fig. 21) with three divisions, lateral divisions acute apically, each with tooth on outer margin, middle division with two pairs of extremely slender appendages and apical gonopore. Female genitalia —Female unknown. 40 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 26 I | po 27 Zz 29 30 31 pes Figs. 25-31. Chinchinota styx, n. g. and n. sp. Fig. 25, head and thorax dorsally; fig. 26, male genital capsule ventrally; fig. 27, connective and _ style ventrally; fig. 28, aedeagus posteriorly; fig. 29, aedeagus laterally; fig. 30, male genital capsule laterally; fig. 31, style laterally. Types.—Holotype male (USNM Type No. 68939) and one paratype male, Rio Caraguata, Matto Grosso, Brazil, March 1953, F. Plaumann. Notes.—T. petulans can be separated at once from ips Kramer, the only other species in the genus, by the symmetrical aedeagus. There are many other details of the male genitalia which differ between the species. These details can be understood best by comparing the drawings of petulans (figs. 18-21) and ips (Kramer 1964: figs. 45-49). Nelidina DELONG For a generic description of Nelidina, see my earlier paper (Kramer 1964: 272). Females of this genus are, as yet, not known. Key TO THE SPECIES OF Nelidina DELONG ( Males ) 1. Length 4.5 mm., aedeagal processes ribbonlike and twisted (Kramer 1964: fos) Ae 2)) ii¢Eolombia’) 2d ee eee aan taeniola Kramer Length 7 mm. or more; aedeagal processes neither ribbonlike nor twisted 9 2. Aedeagal processes uniformly slender and about half as long as shaft, aedeagus without a slender tooth at apex (Kramer 1964: fig. 80) (CUE(= lp] Clea SS SO Oe A eye ee defila (Delong) Aedeagal processes broadened, only narrow at base and apex, and much less than half as long as shaft, aedeagus with a slender tooth on dorsal margin near apex ((fig.22) (Peru) eee sportula, n. sp. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 4] Nelidina sportula, n. sp. (Figs. 22-24 ) Length.—Males 7.5 mm. Structure —Typical of genus in all respects. Coloration.—Sordid stramineous to yellow with only markings consisting of embrowning in clavi near scutellar apex and single brown spot at apex of each clavus. Male genitalia.—Genital capsule laterally (fig. 22) with a broad, moderately long lobe distally on dorsal margin and a preapical notch on ventral margin; apex of style scarcely hooked; aedeagus long, slender, nearly straight, with a pair of subtriangular processes at apex and a small slender tooth dorsally near apex. Gonopore (fig. 24) ventral near apex. Plates (fig. 23) with a few macrosetae distally. Female genitalia —Female unknown. Types.—Holotype male (USNM Type No. 68940) and two paratype males, Callanga, Paucartambo Prov., Dept. Cusco, Peru, 19 February 1952, F. L. Woytkowski. Notes.—N. sportula may be separated from the other members of the genus by the features noted in the key to species. Chinchinota, n. gen. Type-species: Chinchinota styx, new species. Without a carina separating face and crown, clypellus expanded distally, clypeus narrow and tapering downward, crown in dorsal view broadly rounded and wider than long, crown smoothly rounded to face, ocelli on extreme upper edge of face and just barely discernable in dorsal aspect, head including eyes narrower than the short pronotum, length of antennae exceeding total length of body and forewings, venation of forewings highly obscure. Male genitalia: valve absent, plates fused, pygofer without processes or prolongations, anal tube simple, connective elongate and Y-shaped, not fused with aedeagus, style long and slender with mesal lobe highly elongated, aedeagus stout and somewhat U-shaped. Discussion.—This genus will trace to Chinaia in couplet 2 of my key to the genera of the Neocoelidiinae (Kramer 1964: 261). Chinchinota can be separated from Chinaia on the basis of the Y-shaped connective, the exceedingly long slender style, and the lack of elaborations on the distal portion of the pygofer. Chinchinota styx, n. sp. (Figs. 25-31 ) Length.—Male 6.5 mm. Structure.—Suture between clypellus and clypeus absent, antennal ledges oblique, pleural margins of pronotum without carinae. Coloration.—Legs and venter stramineous, washed variably with brown; ground color of face and antennae stramineous, lateral edges of clypellus and of clypeus, as far upward as antennal ledges, narrowly black to brownish, areas of genae around antennal bases broadly black to brownish, each lateral margin of clypeus from yellow ocellus downward with a broad orange red stripe along inner edge of frontal suture; ground color of crown, pronotum, and scutellum stramineous to sordid 42 PROC. ENT. SOC. WASH., VOL. 69, NO. 1, MARCH, 1967 i HA a FUNGOSA 33 32 FUNGOMA 30 Figs. 32-36. Coelindroma fungosa, n. g. and n. sp. Fig. 32, head and thorax dorsally; fig. 33, male genital capsule laterally; fig. 34, ventral cover of male genital capsule. Coelindroma fungoma, n. sp. Fig. 35, aedeagus laterally; fig. 36, aedeagal apex posteriorly. stramineous marked with orange to orange brown as in figure 25; forewing milky hyaline with costal vein, claval suture, and area of clavus along scutellum and commissural margin, orange brown. Male genitalia—Genital capsule laterally (fig. 30) with posterior margin of pygofer rounded and produced beyond plates, an oblique pigmented thickening in upper distal portion; genital capsule ventrally (fig. 26) with plates meeting to form a slight projection at middle of posterior margin, suture between plates visible on about distal third; connective ventrally (fig. 27) broadened at juncture with aedeagus and narrowed towards forked base; style ventrally (fig. 27) much longer than connective, with mesal lobe slender and lateral lobe shorter and _ stouter; style laterally (fig. 31) with mesal lobe hooked at apex and lateral lobe acute at apex; aedeagus laterally (fig. 29) wide with a pair of projecting, moderately long, simple, slender processes on distal margin near apex; aedeagus posteriorly (fig. 28) with processes arising from a membranous or weakly sclerotized area. Female genitalia —Female unknown. Type.—Holotype male (USNM Type No. 68941), Colombia, 1941, L. Richter. Notes.—Except for the comparatively somber coloration, the habitus of Chinchinota styx is much like that of members of the genus Chinaia. The male genital structures are much different, however. See dis- cussion under generic description. PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 43 Coelindroma, n. gen. Type-species: Coelindroma fungosa, new species. With a carina separating face and crown; ocelli on anterior margin of crown, one near each end of carina; clypellus parallel-sided; crown in dorsal view produced and bluntly angular apically with median length at least as great as interocular width; head narrower than pronotum; pronotum with posterior margin broadly and sharply indented; antennae over half as long as total length of body and forewings; venation of forewing obscure except apically. Male genitalia: valve absent, plates fused except at apex and with a few macrosetae distally, pygofer without dorsal processes but with a ventral tooth or hook, anal tube simple, connective Y-shaped with arms strongly divergent and not fused with aedeagus, style of moderate length with mesal lobe hooked, aedeagus elongate and not highly elaborated. Discussion.—This genus will trace to Neocoelidiana in couplet 15 of my key to the genera of the Neocoelidiinae (Kramer 1964: 262). Coelindroma can be separated from Neocoelidiana on the basis of the simple anal tube, the presence of macrosetae on the fused male plates, and the elongated aedeagus. Coelindroma fungosa, n. sp. (Figs. 32-34) Length.—Males 5.5-6 mm. Structure —Suture between clypellus and clypeus present, antennal ledges vertical, pleural margins of pronotum carinate. Coloration.—Venter including legs and face stramineous to sordid stramineous, only distinct marking a black spot at middle of extreme upper edge of face just below but usually touching transverse carina; ground color of crown, pronotum, and scutellum pale stramineous to yellow and typically marked as in figure 32; crown with a stalked black spot at apex and two discal, transverse, orange bands joined to a median longitudinal orange band on midline; pronotum marked with orange as an irregular transverse band behind anterior margin, and as irregular longitudinal bands on midline and near each lateral margin, scutellum with a pair of longitudinal orange bands near lateral angles; forewings milky hyaline with faint embrowning at claval apices. Male genitalia —Genital capsule laterally (fig. 33), pygofer with a large tooth at apex of short ventral margin, posterior margin oblique, anal tube short, connective and style about equal in length, style with a slender apical hook, aedeagus slender throughout length and recurving near apex, gonopore apical. Genital capsule ventrally (fig. 34) with a few macrosetae on either side distally. Female genitalia —Female unknown. Types.—Holotype male (USNM Type No. 68942), Tingo Maria, Peru, 12 August 1960, D. A. Young; one paratype male with same data except collector O. Vargas; two paratype males (California Academy of Sciences), Tingo Maria, Monson Valley, Peru, one 10 November 1954, the other 2 December 1954, E. L. Schlinger and E. S. Ross. Notes.—The coronal markings and the aedeagal shape distinguished Coelindroma fungosa. 44 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Figs. 37-40. Neocoelidia onca, n. sp. Fig. 37, head and thorax dorsally; fig. ! 38, male genital capsule laterally. Biza ava, n. sp. Fig. 39, male genital capsule laterally; fig. 40, aedeagus posteriorly. Coelindroma fungoma, nN. sp. (Figs. 35-36 ) Length.—Male 5.8 mm. Structure.—Not different from that of C. fungosa. Coloration.—Like that of C. fungosa except as follows: without a black spot at middle of extreme upper edge of face, crown with black spot at apex larger and not stalked, orange coronal markings encompassing most of crown behind anterior margins of eyes and then on each side extending along margin of crown between eye and ocellus to cover antennal ledge, markings of pronotum and scutellum much PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 45 less distinct, forewings very narrowly embrowned for entire length along com- missual margin. Male genitalia —Like that of C. fungosa except aedeagus (fig. 35) stouter and with a large preapical bump on dorsal margin. Female genitalia—Female unknown. Type—Holotype male, Tingo Maria, Monson Valley, Peru, 23 December 1954, E. L. Schlinger and E. S. Ross, in California Academy of Sciences, San Francisco, California. Notes.—C. fungoma is certainly very similar to fungosa. The dif- ferences in the markings of the head and thorax, as noted above, may or may not prove to be of specific value; but the aedeagus is quite different from that of fungosa. There is, of course, the possibility that this is an anomaly; however, it seems best to provide a name until this has been demonstrated. Neocoelidia onca, n. sp. (Figs. 37-38 ) Length—Male 5.5 mm. Structure.—Antennae more than half body length; crown (fig. 37) bluntly angular at apex, interocular width slightly more than median length, a weak carina on anterior margin between ocelli; pronotum faintly transversely rugulose. Coloration —Venter, including legs and face, yellow; ground color of crown, pronotum, and scutellum yellow to stramineous with darker markings (fig. 37); crown mesally tan from base anterior to near apex, extreme apex with black spot, a pair of lateral black spots at base and apex of tan area; pronotum mesally brown with poorly defined black spot flanking brown area near anterior margin on each side, humeral angles narrowly black; scutellum discally marked with light brown, a black spot at middle of each lateral margin, extreme apex darkened; forewing pale milky hyaline lightly washed with pale brown, commissural margin broadly brown with inner edge of brown area irregular, claval and costal veins black to brown. Male genitalia—The genital capsule laterally (fig. 38) is typical of all members of the genus (see description Kramer 1964: 262-263); aedeagus close to that of barretti (Baker) (DeLong 1953: pl. 4, fig. 13) but the dorsal shaft is slightly expanded preapically and the cleft ventral shaft is shorter, stouter basally, with left fork slightly shorter than right one; style distally with both a dorsal and ventral projection. Female genitalia —Female unknown. Type.—Holotype male (USNM Type No. 68943), Ignacio, Misiones, Argentina, May 1961, Krauss. Notes.—N. onca is readily distinguished from all other members of Neocoelidia by the distinctive markings on the crown, pronotum, and scutellum. This species provides the southernmost record for the genus in South America. Biza ava, n. sp. (Figs. 39-40) Length—Male 7.5 mm. Structure.—Typical of genus in all respects. 46 PROC. ENT. SOC. WASH., VOL. 69, No. 1, MARCH, 1967 Coloration.—Ground color yellow, thoracic venter and legs slightly paler, with- out markings on head or thorax, forewing yellow on basal two-thirds with a transverse brown band from near claval apex to costal margin, area beyond band pale brown hyaline with veins brown. Male genitalia—The genital capsule laterally (fig. 39) with apical portion of pygofer broad and tooth on ventral margin fairly large, stylar apex stout and hooked, aedeagus narrowed and recurved distally with a pair of avicephaliform processes on shaft (fig. 40). Female genitalia—Female unknown. Type.—Holotype male (USNM Type No. 68944), Summit, Panama, Canal Zone, July 1953, Krauss. Notes.—In my key to the species of Biza (Kramer 1962: 103), ava will trace to couplet 2. It can be separated from both crocea (ibid. fig. 7) and chinai (ibid. fig. 3) by the much shorter and uniquely shaped aedeagal processes. Chinaia BRUNER & METCALF This genus at the time of its revision (Kramer 1959) included eleven species. Additional material from Panama has shown that permista Kramer is merely a color variant of caprella Kramer. Hence, Chinaia permista is a synonym of Chinaia caprella Kramer, NEW SYNONYMY. The only species described since the revision is serrata Linnavuori (1965: 149) from Matto Grosso, Brazil. Xigilliba KRAMER Xiqilliba was described (Kramer 1964: 268) as a monobasic genus with the type-species bellator Kramer, a Brazilian species. Linnavuori (1965: 147) described a second species, coelidoides, from “Rioja, South America.” The two species are identical. Hence, Xiqilliba coelidoides Linnavuori is a synonym of Xigilliba bellator Kramer, NEW SYNONYMY. REFERENCES DeLong, D. M. 1953. A synopsis of the tribe Neocoelidinii in the Americas (Homoptera: Cicadellidae). Lloydia 16(2): 93-131. Evans, J. W. 1947. A natural classification of leaf-hoppers (Jassoidea, Homoptera). Part 3: Jassidae. Ent. Soc. London Trans. 98: 105-271. Kramer, J. P. 1959. An elucidation of the Neotropical genus Chinaia with a key to males and a new allied genus (Homoptera: Cicadellidae: Neocoelidii- nae). Proc. Biol. Soc. Washington 72: 23-32. . 1962. $<— STATE———_ Ch —— ae neg PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. 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When tables are necessary, each table should be prepared as a line drawing or typed on a separate page with heading at top and footnotes below. Number footnotes consecutively for each table. Use only horizontal rules. Illustrations—No extra charge is made for line drawings or halftones. Authors must plan their illustrations for reduction to the dimensions of the printed page and the individual figures must be mounted on suitable board. Proportions of full-page illustrations should closely approximate 45% X 6” (26 x 36 picas); this usually allows explanatory matter to appear on the same page. On the back of each illustration should be stated (1) the title of the paper, (2) the author’s complete name and address, and (3) the number of the illustration such as “Now! (of 3)* ete: Figure legends—Legends should be typewritten double-spaced on separate pages headed ExpLANATION OF FicurREs and placed following REFERENCES. Do not attach legends to illustrations. Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) LEONARD, M. D.—Macrosiphon rosae (Linnaeus) on Ilex (Homoptera: ATTIC AR: ieee een Ue aN ee Ne eee Re ook Woe ns 59 MEDLER, J. T.—A note on Ceropales maculata fraterna Smith in Wiscon- sin: (ymenopterey Pompilidae) 2-0 eet 92, POLHEMUS, J. T.—Notes on North American Saldidae (Hemiptera) — 24 SABROSKY, C. W.—Notes on the tachinid genus Cylindromyia in North PAierical (Piper) ets ie i da 60 SCANLON, J. E. and E. L. PEYTON—Anopheles (Anopheles) tigertti, a new species of the aitkenii group from Thailand —_------------- 18 SMITH, D. R.—Extended distribution for Pristiphora abbreviata (Hartig) (iymenoptera:| Tenthredinidaé) +02 0 ee 92, SMITH, D. R.—Two sawflies new to North America (Hymenoptera: Ten- Reem ICRe) gta Ga Ase 2 ee te eS 95 SPANGLER, P. J.—A new Brazilian Berosus and a description of the female of B. spectatus D’Orchymont (Coleoptera: Hydrophilidae) — 64 STEYSKAL, G. C.—A key to the species of Tritoxa Loew (Diptera: Otitidae) 94 STRASSEN, R. ZUR—New locality records and a new species of Chiro- this Halliday '( Thysasoptera: Thripidae). 47 TOWNES, H.—Taxonomic notes on Kentrotryphon, Ktenostilpnus and Pa- rentypoma (Hymenoptera: Ichneumonidae) -_..---._..----------------------------— 58 TOWNES, H.—Notes on the Motschoulsky types of Ichneumonidae (Hy- TRICHAOPIUE TA: )) ea ee aku Me See A eR ob) 93 INFORMATION FOR CONTRIBUTORS Cover 3 OO Vol. 69 JUNE 1967 No. 2 S95. 70673 PROCEEDINGS of the ENTOMOLOGICAL SOCIETY of WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 PUBLISHED QUARTERLY CONTENTS (Continued on back cover) FLINT, O. S., JR.—Studies of Neotropical caddis flies, VI: On a collection Praminnierciwestern Newco es. EN ee) ee FRANCLEMONT, J. G.—Two new species of noctuids (Lepidoptera, Noc- fC ee NOCUIINAG hse eee ee re es ee ee ee A 97 GAGNE, R. J.—Tinconaria albescentis, a new North American genus and species of Cecidomyiidae found in seeds of Triodia albescens Vasey MUDDY arse visa) meee ce te a 2 ee ee ee OAL 155 GALINDO, P. and H. TRAPIDO—Description of Haemagogus aeritinctus, a new species from British Honduras, with a note on the validity of Haema- gogzus lucifer (H. D. and K.)) (Diptera: Culicidae): —._.-._ 103 KINGSOLVER, J. M. and R. E. WHITE—A review of the genus Aulonium from the United States (Coleoptera: Colydiidae) 149 MARINKELLE, C. J.—Cimex hemipterus (Fabr.) from bats in Colombia, south America (Hemiptera: Gimicidae) 228 2) 179 McDANIEL, B.—Lectotype designation from three syntypes of Radfordia suboneer Fiwing ( Acarina:; Myobiidae) 0.0 MUESEBECK, C. F. W.—A new braconid parasite of the potato tuberworm LE bonetjct GLC) (21 | OMe Semee aie Se ap: Sean ARONA AN PINE Ue RPS COs ty eS ae oe vers NOVAK, J. A., W. B. STOLTZFUS, E. J. ALLEN, and B. A. FOOTE—New host records for North American fruit flies (Diptera: Tephritidae) ENTOMOLOGICAL SOCIETY OF WASHINGTON OrcANIzED Marcu 12, 1884 OFFICERS FOR 1967 Mr. Louis G. Davis, President Dr. Jon L. Hernine, Editor ARS, USDA, Rm. 466 c/o Department of Entomology Federal Center Bldg. U.S. National Museum Hyattsville, Maryland 20782 Washington, D.C. 20560 Dr. Ricuarp H. Foore, President-Elect Mr. RosperT SMILEY, Custodian c/o Department of Entomology Entomology Research Division, USDA U.S. National Museum Washington, D.C. 20250 Washington, D.C. 20560 Dr. Rautpeu A. BRAM, Recording Secretary Mr. Vicror ADLER, Program Committee Chairman Department of Entomology Entomology Research Division, ARS, USDA U.S. National Museum ARC, Beltsville, Maryland 20750 Washington, D.C. 20560 Mr. Davp R. Smitn, Corresponding Secretary Carr. WitL1AM B. Hutt, Membership Committee c/o Department of Entomology Chairman U.S. National Museum 746 Beall Ave. Washington, D.C. 20560 Rockville, Maryland 20850 Dr. Artruur K, Buropitt, Jr., Treasurer Dr. H. H. SHeparp, Delegate to the Washington ARS, USDA, Rm. 125 Academy of Sciences North Bldg. PPA-ASCS Beltsville, Maryland 20705 U.S. Department of Agriculture Washington, D.C. 20250 Honorary President Dr. T. E. SNypER, U.S. National Museum Honorary Members Mr. C. F. W. MuEsEBECK, U.S. National Museum Dr. F. W. Poos Dr. E. N. Cory MEETINGS.—Regular meetings of the Society are held in Room 43 of the U.S. National Museum on the first Thursday of each month from October to June, inclusive, at 8 P.M. Minutes of meetings are published regularly in the Proceedings. MEMBERSHIP.—Members shall be persons who have demonstrated interest in the science of entomology. Annual dues for members are $5.00; initiation fee is $1.00 (U.S. currency). PROCEEDINGS.—Published quarterly beginning with March by the Society at Washington, D.C. Members in good standing are entitled to the Proccedings free of charge. Nonmember subscriptions are $10.00 per year, both domestic and foreign (U.S. currency), payable in advance. All remittances should be made payable to The Entomological Society of Washington. The Society does not exchange its publications for those of other societies. STATEMENT OF OWNERSHIP Title of Publication: Proceedings of the Entomological Society of Washington. Frequency of Issue: Quarterly (March, June, September, December). Location of Office of Publication, Business Office of Publisher and Owner: The Entomological Society of Washington, c/o Department of Entomology, U.S. National Museum, Washington, D.C. 20560. Editor: Dr. Jon L. Herring, same address as above. Managing Editor and Known Bondholders or other Security Holders: none. ' ee This issue mailed August 30, 1967 Second Class Postage Paid at Lawrence, Kansas, U. S. A. ALLEN PRESS, INC. ee eo LAWRENCE, KANSAS U.S. > PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Vol. 69 JUNE 1967 No. 2 TWO NEW SPECIES OF NOCTUIDS (LepiwoprerA, Nocrurar, Nocrurnae)! Joun G. FRANCLEMONT Department of Entomology Cornell University Ithaca, New York 14850 The species of Abagrotis described here was set aside as new in the fall of 1960. Sometime later, Mr. John S. Buckett of the University of California, Davis, indicated that he intended to revise the genus Abagrotis. We agreed that I would publish the description of the new species from Madera Canyon, Arizona. A grant (No. 303-Johnson Fund) from the American Philosophical Society made possible the collecting in Madera Canyon during the summer of 1960, and'a grant (No. 3339-Penrose Fund) also from the American Philosophical Society made possible the collecting in Madera Canyon during the spring of 1963. This assistance is gratefully ac- knowledged. The drawings are by Mrs. Margaret A. Menadue. Abagrotis alampeta, n. sp. A dark, rather obscurely marked species, with the male genitalia somewhat similar to alcandola Smith, 1908, p. 288 (= tristis Barnes and McDunnough, 1912, p. 8, pl. 1, fig. 17), but easily separated by its simple antennae; those of the male of alcandola are serrate. Description: Male with head, thorax, and forewings above deep fuscous or brownish black, the scales pale tipped, giving a frosted appearance. Palpi with first and second segments, except apex of second, black; a broad black band from behind eye to base of forewing, a small triangular black patch in front of eye; the overall effect, a black band beginning on the palpi and ending at the base of the forewing. Patagia with a line of pale, whitish scales on the dorsal margin, a second line of pale scales about one-third below dorsal margin, immediately below this a band of black scales; anterior tuft of thorax low, apex black and a narrow band of black scales below this. Forewing with the markings somewhat indistinct; basal half-line pale with dark marginal lines; antemedial line pale with dark marginal lines, irregularly crenulate, outwardly oblique from costal margin to inner margin; postmedial line pale with dark marginal lines, slightly and irregu- 1~Immediate publication secured by full payment of page charges—KEditor. 98 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Figs. 1-2. Abagrotis alampeta. Scale 1 mm. 1, paratype, male genitalia, slide: JGF 4519; la, paratype, aedoeagus, slide: JGF 4517; 2, paratype, female genitalia, slide: JGF 4518. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 99 larly crenulate, excurved below costa, then incurved at Cui and Cuz, then almost straight to inner margin; orbicular circular with a vague, pale annulus; reniform moderately large, with a vague, pale annulus; subterminal vague, pale, somewhat irregular; subterminal area pale; terminal line a series of vague, blackish bars; fringe concolorous with subterminal area. Hindwing and dorsum of abdomen fuscous; hindwings paler toward base; veins dark scaled; fringe with a light yellowish fuscous line at base, followed by a dark fuscous line, outer half white (does not show well in the photographs). Wings below, light fuscous with a frosted appearance except on the disk of the forewings, some black scales inter- mixed, especially on the costal areas of the forewings. Anal tuft of abdomen reddish brown. The males show some variation in the amount of pale scaling on the basal and costal areas of the forewing and in the distinctness of the annuli of the reniform and orbicular. The general color may be reddish in some specimens. The females are generally paler than the males and with a decidedly yellowish gray cast; some are very reddish, but this color phase is not common; about ten to twelve percent of the Madera Canyon population exhibit it. Expanse: 33-38 mm. Male genitalia as figured. The adeoeagus with a group of short, stout spines at apex; the vesica armed with a single stout cornutus and a long, broad, ridged, sclerotized plate. Female genitalia as figured. Described from 167 males and 204 females from Madera Canyon in the Santa Rita Mountains of southeastern Arizona. TYPE: Male. Madera Canyon, 4880’, Santa Rita Mts., Santa Cruz Co., Arizona 6 May 1963, J. G. Franclemont. In Franclemont Collec- tion. PARATYPES: 166 Males: 51, Madera Canyon, 5600’; Santa Rita Mts., Santa Cruz Co., Arizona, 15 June-11 July 1960; 74, Madera Can- yon, 5600’; and 41, Madera Canyon, 4880’, Santa Rita Mts., Santa Cruz Co., Arizona, 20 April-23 May, 1963. 204 Females: 1, Madera Can- yon, 5600’, Santa Rita Mts., Santa Cruz Co., Arizona, 8 October 1959; 91, Madera Canyon, 5600’; Santa Rita Mts., Santa Cruz Co., Arizona, 15 June-15 July 1960; 61, Madera Canyon, 5600’; and 51, Madera Canyon, 4880’, Santa Rita Mts., Santa Cruz Co., Arizona, 20 April-27 May 1963; all collected by J. G. Franclemont. Paratypes will be dis- tributed to the United States National Museum, the British Museum (Natural History), the Canadian National Collection, the American Museum of Natural History and the Los Angeles County Museum of Natural History; the majority will be retained in the Franclemont Col- lection; two are at present in the Bauer—Buckett Collection, Davis, California and others will be deposited in that collection. The species has also been collected in Cave Creek Canyon in the Chiricahua Mountains of southeastern Arizona and in the Flagstaff area of north central Arizona. NO. 2, JUNE, 1967 > 69 PROC. ENT. SOC. WASH., VOL. 100 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 101 Figs. 5-7. Abagrotis alampeta. Scale approx. 1.4 x. 5, Abagrotis alampeta. TYPE; 6, Abagrotis alampeta, paratype male with pale scaling; 7, Abagrotis alampeta, paratype female; 8, Anicla cemolia, TYPE. Five female paratypes of barnesi Benjamin, 1921, p. 97, from the following localities are this species: Mohave., Co., Ariz. (2), Reding- ton, Arizona (1), and Palmerlee, Cochise Co., Ariz. (2). Anicla cemolia, n. sp. This species is very similar in appearance to the figure of Agrotis altes Druce, 1889, vol. 1, p. 285, vol. 3, pl. 27, fig. 12, described from Teapa, Tabasco, Mexico. Hampson, 1903, p. 525, treates altes as a junior synonym, the male, of Agrotis ornea Druce, 1889, vol. 1, p. 285, vol. 3, pl. 27, fig. 11, described from near Mexico City. The forewings of the new species are paler; there is no conspicuous black spot below 2A as figured by Druce for altes and ornea; and the hindwing is white, slightly hyaline in both sexes, not shaded outwardly with fuscous as in the male (altes) of ornea and not dark fuscous as in the female. This species differs from Anicla infecta (Ochs.) by its much smaller size, white hindwings, and narrowly pectinate male antennae. Description: Antennae of male narrowly pectinate, of female simple and sparsely ciliate. Head and thorax light gray with a slight yellowish cast, with a sparse < Figs. 3-4. Anicla cemolia. Scale 1 mm. 3, paratype, Windley Key, Monroe Co., Florida, male genitalia, slide: JGF 5276. 3a, paratype, aedoeagus, slide: JGF 5276. 4, paratype, Key Largo, Monroe Co., Florida, female genitalia, slide: JGF 5272. 102 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 scattering of black scales and with a small patch of black scales at middle of collar (patagia). Forewing concolorous with head and thorax, light gray with a yellowish cast and with a scattering and clustering of black scales; the lines obso- lescent; the basal line represented by a small black spot on costa near base of wing and a second black spot in fold below Cu; ante-medial line represented by a black spot on costa, a black spot in fold, and a diffuse cluster of black scales on the inner margin; postmedial line represented by a black spot on costa and vague line of black scales, the line excurved below costa, then slightly curved to inner margin; reniform a broad gray annulus with a pale central area and two black spots, the larger on the outer side, the smaller on the inner side; subterminal line diffuse, irregular, broad, blackish, area beyond subterminal line blackish, terminal line a series of black dots on the veins; fringe concolorous with ground color of wing. Hindwing white; two black spots near the apical angle represent- ing the terminal line; fringe white. Expanse: 25-28 mm. Male genitalia as figured. Female genitalia as figured. TYPE: Male. Tavernier, Monroe Co., Florida, October 21, 1965, J. N. Todd. In Franclemont Collection. PARATYPES: 14 males and 4 females, Tavernier, Monroe Co., Florida, August 16, 1955-October 23, 1955, J. N. Todd; 1 male in United States National Museum Collection, 2 males in Franclemont Collection, 11 males and 4 females in C. P. Kimball Collection; 8 males and 2 females, Key Largo, Monroe Co., Florida, March 27, 1965—April 27, 1965 and July 21, 1965-August 26, 1965, Mrs. Spencer Kemp; 3 males and 2 females in Franclemont Collection, 5 males in C. P. Kim- ball Collection; 1 male, Windley Key, Monroe Co., Florida, December 11, 1955, J. N. Todd; in Franclemont Collection; 1 female, Craig, Florida, April 16, 1956, J. N. Todd; in the United States National Mu- seum. Collection. There are twenty males and five females from Tavernier Key and one male from Key Largo also before me, but the condition of these specimens is too poor to include them in the type series. REFERENCES Barnes, W. and J. H. McDunnough. 1912. Fifty new species and varieties. Contributions to the natural history of the Lepidoptera of North America 1(5): 3-36. Benjamin, F. H. 1921. A study of the noctuid moths of the genera Lampra, Hbn, and Cryptocala, gen. noy. Bull. S. Calif. Acad. Sci. 20(3): 71-154. Druce, H. 1889. In Godman and Salvin, Biologia Centrali-Americana, Insecta, Lepidoptera—Heterocera 1 (text, i-xxxii, 1-490), 3 (Plates). Hampson, G. F. 1903. Catalogue of the Lepidoptera Phalaenae in the British Museum 4 (Agrotinae) i-xx, 1-689. Smith, J. B. 1908. Notes on the species of Rhynchagrotis Sm. with descriptions of new species. Can. Ent. 40: 221-228, 286-288. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 103 DESCRIPTION OF HAEMAGOGUS AERITINCTUS, A NEW SPECIES FROM BRITISH HONDURAS, WITH A NOTE ON THE VALIDITY OF HAEMAGOGUS LUCIFER (H. D. AND K.) (DiererRA: CULICIDAE) Pepro Gaxinpo! and Haroip TRAPIDO" * In 1955, while conducting investigations on the ecology of yellow fever in Middle America, the authors visited British Honduras for several days. The specific objective of this trip was to investigate the mosquito fauna of the area in relation to the possible transmission of yellow fever virus. At the time, there was an outbreak of jungle yellow fever moving west along the north coast of Honduras toward the east coast of Guatemala and British Honduras. We arrived at Belize several days after a devastating hurricane had passed through that territory, so that conditions were not favorable for the capture of adult mosquitoes. Therefore, efforts were concentrated on the finding of larvae in tree- holes within a variety of ecological situations. Collections were made in the vicinity of Belize and of Stann Creek, as well as at several points along the road between these two towns. The present report deals with descriptions of the adult, male terminalia, pupa and larva of a new species of Haemagogus found breeding in mangrove associations at two localities. A discussion is also included of the taxonomic relationships between the new species and related forms. From the evidence presented the authors conclude that Haemagogus lucifer (Howard, Dyar and Knab, 1912) is a valid species and not a synonym of H. regalis Dyar and Knab, 1906 as pro- posed by Komp (1954). Haemagogus (Haemagogus) aeritinctus, n. sp. MALE.—Head. Proboscis somewhat longer than the fore femur, uniformly pur- ple in color. Palpi very short, barely longer than the clypeus, clothed with purple scales. Clypeus naked, shiny black in color. Antennae about half as long as the proboscis, sparsely plumose; tori dark, bare. Vertex blue-scaled with a very nar- row line of silvery scales bordering the eyes. Occiput clothed with flat, broad, straw-colored scales. Mentum light-scaled. Thorax. Anterior pronotal lobes large, inner angle clothed with silvery scales, remainder of lobe blue-scaled. Mesonotum black, clothed with bright coppery scales which give a purplish reflection under oblique illumination, except for a small spot of blue-green scales above the roots of the wings. Scutellum with a mixture of blue-green and coppery scales. Pleuron mostly covered with bright silvery scales except for posterior pronotal lobe and meron which are bare and shiny black. Pleural chaetotaxy as follows: no propleurals, two or three black 1 Gorgas Memorial Laboratory, Apartado 6991, Panama, Republic of Panama. 1,2 Present address: The Rockefeller Foundation, Universidad del Valle, Facultad de Medicina, Apartado Aéreo 6555, Cali, Colombia. 104 PROC, ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Male terminalia of Haemagogus aeritinctus n. sp. Fig. 1, Basistyle; fig. 2, Disti- style; fig. 3, Mesosome; fig. 4, Eighth tergite; fig. 5, Claspette; fig. 6, Tenth ster- nite. setae on posterior pronotum, no post-spiraculars, no sternopleurals, two yellowish pre-alars, three or four upper mesepimerals. Postnotum with one seta near its posterior border. Coxae and trochanters yellowish with a patch of silvery scales. Femora clothed with purplish scales, base and undersides largely pale-scaled; hind femur also bearing a long patch of silvery scales on its anterior surface. PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 105 Tibiae and tarsi covered with purple scales; tarsal claw formula 1.1—1.1-0.0. Wings shorter than the abdomen; cell-Re much longer than its petiole. Halter with pale base and dark knob, tip silvery-scaled. Abdomen. Abdominal tergites clothed with purple scales and large basal seg- mental patches of silvery scales which become progressively smaller apically. Male terminalia. Eighth tergite (Fig. 4) bearing 42 large setae from outstand- ing sclerotized bases in irregular rows which are at most three setae deep. The setae of the posterior row appear to be very narrow, striated, modified scales and are flanked by three or four of the longest setae of the sclerite. Basistyle as figured (Fig. 1). Dististyle (Fig. 2) about two-thirds the length of the basistyle, expanded on distal third then tapering sharply to a rather acute point, giving the appearance of a narrow snout; appendage of the dististyle in- serted subapically, expanded distally, tongue-like. Stem of claspette (Fig. 5) rather narrow, slightly sinuate, bent sharply at right angles on distal third, clothed with sparse short pile on proximal half and bearing two stout setae from promi- nent insertions on its inner surface near the base. Filament a flat, widely ex- panded leaf inserted distally on the stem. Other parts as illustrated in figures. FEMALE.—Coloration as for the male; proboscis slightly shorter; tarsal claws unarmed. LARVA.—Head (Fig. 18). Globose, dark. Antennae glabrous, small, barely exceeding the anterior border of head; antennal hair single, short, inconspicuous, inserted slightly beyond the middle of antennae. Head chaetotaxy as figured. Thorax. Skin glabrous. Chaetotaxy as follows: Prothoracic hairs: Nos. 1, 2, 3 inserted in the same chitinized plate; No. 1 long, with 2-5 branches; No. 2 about equal to No. 1, single; No. 3 as long as No. 1, 4-8 branched; No. 4 slightly shorter than No. 1 with 3-7 branches; Nos. 5 and 6 equal in size, with coalescing sclero- tized bases, longer than No. 1; No. 7 of same size as Nos. 5 and 6, 3-4 branched; No. 8 slightly more than half as long as No. 4, with 3-8 branches; Nos. 9, 10, 11 and 12 inserted in same sclerotized tubercle, Nos. 9 and 11 always single; No. 10 with 1-5 branches; No. 12 single or double. Mesothoracic hairs: No. 1 with 2-8 branches; No. 2 slightly more than half as long as No. 1, single; No. 3 single, longer than No. 2; No. 4 about equal to No. 3, single; No. 5 single or double; Nos. 6 and 7 inserted on one sclerotized plate, both about equal to No. 5 in length; No. 6 with 4-8 branches; No. 7 single; No. 8 5-9 branched; Nos. 9, 10, 11 and 12 inserted on same sclerotized base, No. 9 single as long as No. 8; No. 10 6—8 branched; No. 11 single; No. 12 shorter than No. 4 with 1-3 branches; No. 13 multiple, about equal to No. 8; No. 14 with 3-7 branches. Metathoracic hairs: No. 1 3-7 branched; No. 2 longer than No. 1, single; No. 3 shorter than No. 2, 2-6 branched; No. 4 with 1-3 branches, shorter than No. 1; No. 5 about equal to No. 4 with 2-6 branches; No. 6 single, as long as No. 2; No. 7 a multiple tuft; No. 8 about as long as No. 5, multiple; Nos. 9, 10, 11, 12 inserted on same sclero- tized base; No. 9 single, No. 10 a multiple tuft, No. 11 single, No. 12 3-branched. Abdomen. Skin glabrous. Chaetotaxy as follows: Hair No. 6 with 3-5 branches on segments I and II, double or rarely triple on segments III to VI, single on segment VII; hair No. 7 single, or rarely double on segment I. Subdorsal hairs very variable, usually with 3-4 branches. Comb-scales 20 to 38 in number, ar- ranged in a patch 3 rows deep. Terminal segments as figured (Figs. 15, 16 and Ls 106 PROG. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Male terminalia parts of Haemagogus lucifer and H. regalis. Fig. 7, H. lucifer: 8th tergite; fig. 8, H. regalis: 8th tergite; fig. 9, H. lucifer: Dististyle; fig. 10, H. regalis: Dististyle; fig. 11, H. lucifer: Mesosome; fig. 12, H. regalis: Meso- some; fig. 13, H. lucifer: Claspette; fig. 14, H. regalis: Claspette. PUPA.—Trumpets short and stumpy, diameter of pinna about twice the length of the meatus; tracheoid small, about 4% as long as meatus. Abdominal chaetotaxy as figured (Fig. 19). Rudimentary spiracles on segments II-VII quite prominent and located laterally on the segments. PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 107 Type Material Holotype: Male, mounted on card point. Associated larval and pupal skins mounted on a slide. Terminalia stained, dissected and mounted on a second slide. Reared from larvae collected in a rot-hole of a red mangrove tree (Rhizophora mangle ), in the vicinity of Stann Creek, British Honduras, on 5 October, 1955. P. Galindo and H. Trapido, collectors. Allotype: Female, mounted on card point. Associated larval and pupal skins mounted on a slide. Same collecting data as holotype. Paratypes: Two males mounted on card-points. Associated larval and pupal skins and terminalia mounted on separate slides. Same col- lecting data as holotype. One male mounted on a card-point. Associated larval and pupal skins and terminalia mounted on separate slides. Reared from larvae taken in a rot-hole of a Ficus tree at the fringe of a mangrove swamp, near Belize, British Honduras, on 5 October, 1955. P. Galindo and H. Trapido, collectors. Eight males mounted on card points. Associated pupal skins and terminalia mounted on separate slides. Reared from pupae. Seven specimens with same collecting data as holotype, one taken at Belize, same collecting data as paratype above. Seven males mounted on card-points. Two with terminalia mounted on slides. Four with same collecting data as holotype, and three from Belize. Four females mounted on card-points. Associated larval and pupal skins on slides. Same collecting data as holotype. Twelve females, nine from type locality and three from Belize, mounted on card-points with associated pupal skins mounted on slides. Four females mounted on card-points taken while attempting to bite in a mangrove swamp near Belize, B. H., on October, 1955. P. Galindo and H. Trapido, collectors. Ten larval skins, eight from the type locality and two from Belize, mounted on slides. Disposition of type material. Holotype and allotype to be deposited in the U.S. National Museum. Paratype series to be divided between the U.S. National Museum and the mosquito collections of the Univer- sity of California at Los Angeles (U.C.L.A.) and of Gorgas Memorial Laboratory, Panama. Taxonomic discussion. This new species belongs to the group of Middle American species of Haemagogus sens. str. which are com- monly found breeding in mangrove swamps. These species are: regalis D. and K., lucifer H. D. and K., iridicolor Dyar, chalcospilans Dyar and boshelli Osorno. The peculiar coppery sheen of the mesonotal scales in aeritinctus n. 108 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 / ~ =—— ratte SAL ——————— W/m ty wee < = > Te eS = hy: Immature stages of Haemagogus aeritinctus n. sp. Fig. 15, Terminal segments of larva; fig. 16, Individual pecten tooth; fig. 17, Individual scale of comb of 8th segment; fig. 18, Head capsule of larva, fig. 19, Abdominal chaetotaxy of pupal skin. sp., differentiates it from all other species of Haemagogus, with the exception of uriartei Shannon and Del Ponte, which occurs in merid- ional South America and belongs to a different subgenus. The yel- lowish color of the coxae of aeritinctus is shared only by H. chalco- spilans, from which aeritinctus may be separated by the color of the mesonotal scales, the longer proboscis and the shape of the dististyle PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 109 and claspettes of the male terminalia. The male terminalia of aeritinctus shares characters with both regalis and lucifer. The shape of the disti- style is like the latter species while the claspettes and mesosome are much as in regalis. However, the three species may be separated by the number of setae with sclerotized insertions on the eighth tergite. In aeritinctus they range from 33 to 46 (in 20 specimens ) with a mean of 40.9; in regalis from 46-85 (in 59 specimens) with a mean of 64.5, and in lucifer from 62-104 (in 78 specimens) with a mean of 85.8. Komp (loc. cit.) sank H. lucifer (H. D. and K., 1912) a mosquito described from Panama, in the synonymy of H. regalis D. and K., 1906 with type locality in Sonsonate, El] Salvador, stating: “The writer has examined the male terminalia of H. lucifer on slide No. 1461 in the U.S. National Museum collection, which was made from a male of the type series . . . and finds that the terminalia of this specimen correspond in all particulars with the male terminalia on three slides . . . which are from the type series of H. regalis. The writer has many specimens of “H. lucifer’ from Panama, of which he has dissected and mounted the male terminalia. These agree with those of the three slides of the type series of H. regalis, noted above.” In further support of his case, Komp (loc. cit.) published four photomicrographs of the terminalia slides of H. lucifer and H. regalis discussed above, pointing out the apparent similarity of the different parts in these preparations. The authors have been fortunate in working with fresh material of H. regalis from El Salvador, Guatemala and Mexico, as well as with a long series of H. lucifer from various parts of Panama and Colombia and find these two taxons to differ in the following respects: a). The shape of the dististyle. In H. lucifer (Fig. 9) it swells be- yond the middle and then tapers to a sharp point, appearing narrowly snout-shaped. In H. regalis (Fig. 10) there is no appreciable swelling in the dististyle which is arcuate and tapers gradually and very slightly from base to apex. This character can be appreciated in the photo- micrograph of the terminalia of H. lucifer published by Komp (loc. cit.) but cannot be clearly distinguished in the photomicrograph of H. regalis, due to the orientation of the specimen. However, one of us (P.G.), has examined all the slides of the type series of H. regalis in the U.S. National Museum, and finds that, with the exception of the holotype, they all appear to agree well in this characteristic with the fresh material in the authors’ collection, despite the fact that they are rather poor preparations. The holotype lacks both dististyles so it could not be studied properly. b). The mesosome of H. regalis (Fig. 12) has a rather prominent subapical serrated carina along the ventral surface. In H. lucifer (Fig. 11) this carina is always missing although some slight serrations are sometimes present just below the apex. This character can only be 110 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 seen in preparations where the mesosome has been dissected out and mounted in lateral view. c). In H. lucifer (Fig. 13) the stem of the claspette swells medianly, thus appearing much stouter than in H. regalis (Fig. 14) which has slender and slightly sinuate claspette stems. This characteristic is ex- tremely constant and can be seen in Komp’s photomicrographs, but the difference was thought by him to be due to distortions during the preparation of the slides which are in poor condition. d). Both forms have thick, stiff setae inserted in sclerotized bases on the distal margin of the eighth tergite, but the mean number of these setae differs. In 59 specimens of H. regalis (Fig. 8) the number of setae range from 46 to 85 with a mean of 64.5. In 78 specimens of H. lucifer (Fig. 7) the number of thick setae on the eighth tergite range from 62 to 104 with a mean of 85.8. e). These two forms occupy widely different geographical areas. H. lucifer is known from Colombia, Panama and southeastern Costa Rica and is replaced in Costa Rica along both coasts by H. iridicolor, which extends into Nicaragua. H. regalis, on the other hand, occurs from the Pacific coast of Mexico in the state of Chiapas, south to the Gulf of Fonseca in the border area between El Salvador and Hon- duras. On the Atlantic coast it has been found only in the states of Veracruz and Tabasco in southern Mexico, being replaced to the south in British Honduras and Guatemala by the new species H. aeritinctus. The latter is in turn isolated from contact with H. lucifer by H. iridi- color which occupies all of the Atlantic littoral of Nicaragua and Costa Rica. About a year after his publication, the late W. H. W. Komp in a letter to the authors, stated: “I am now inclined to believe that lucifer is a subspecies of the more northerly occurring regalis.”. However, in view of the evidence presented above we see no reason at present to treat these two forms as only subspecifically distinct and therefore consider them as distinct species. Geographical distribution. Haemagogus aeritinctus is known only from Belize and Stann Creek, British Honduras, and from the east coast of Guatemala. Altitudinal distribution. Sea-level. Ecology. The authors have taken this species in mangrove swamp at Stann Creek, and among Ficus trees adjacent to mangrove at the outskirts of Belize. Larvae were extremely abundant in rot holes in the mangrove and Ficus, but adults attacking man were scarce. The scarcity of adults is probably not significant as the collecting was done only several days after a devastating hurricane in the area. The adults which approached attacked about the head. Careful search of the coastal mangrove of northern Honduras on several occasions failed to PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 ilstal reveal any Haemagogus there. We also made one search of the man- grove near Chetumal, Quintana Roo, Mexico, just north of British Honduras, but did not find Haemagogus. Dr. Jorge Boshell (personal communication ) collected the new species along the east coast of Gua- temala just north of Puerto Barrios. He captured adults attempting to bite in mangrove swamps and recognized the species on sight by the coppery sheen of the mesonotum, the yellowish coxae and the habit of females of attacking man about the head. It appears that this is a species of restricted range. It is morphologically close to, and the ecological equivalent of the littoral species H. regalis. Relation to yellow fever. Nothing is known, but probably not of significance in the epidemiology of sylvan yellow fever because of its restricted littoral range and habitat. The species is in part peridomestic however, and appears to be abundant at Belize. Thus it might con- ceivably become involved in the transmission of yellow fever from man to man if the disease were once introduced. Acknowledgment. The authors wish to express their appreciation to Professor Eustorgio Méndez, Gorgas Memorial Laboratory, for the illustrations. REFERENCES Komp, W. H. W. 1954. Haemagogus lucifer H. D. and K., 1912, a synonym of Haemagogus regalis D. & K., 1906. Proc. Ent. Soc. Wash. 56: 193-195. A NEW MEMBER OF A SIBLING SET BELONGING TO THE DROSOPHILA TRIPUNCTATA GROUP (DiprerA: DROSOPHILIDAE ) SARAH BEDICHEK PIPKIN Department of Zoology, Howard University, Washington, D.C. 20001 Drosophila leticiae Pipkin, n. sp. External characters of imagines: 4 9, Arista with 6 or 7 dorsal and 3 ventral branches in addition to terminal fork. Front dull yellowish brown, darker in ocellar triangle; ocelli pink; 7 inconspicuous frontal hairs on each side apex of frontal triangle; orbital hairs 5 or 6. Proclinate orbital 44 posterior reclinate; an- terior reclinate thin, about 4 proclinate. Face, ¢, white; 9, yellowish. Carina broad, flat, widening distally; white in male; yellowish brown in female. One prominent oral bristle; proboscis yellowish, shining, darker distally, with yellowish hairs. Cheek yellowish, absent behind; width from base of oral bristle to eye border 42 greatest diameter of eye. Orbits yellowish brown. Eye dull red, a little darker in the dorsal 44; pile straw-colored. Eye index 1.2. Palpi yellow, with one prominent subapical hair, 2 others on lateral margin of palpus in addition to small hairs. Acrostichal hairs in 6 rows; mesonotum shining brown; pleura shin- 112 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Fig. 1. Outline drawing of paired homologues of hybrids between D. leticiae, n. sp. and D. metzii showing translocations involving the 4 longer autosomes and some absence of pairing of portions of homologues. Only the X chromosomes (accidentally fragmented in making the preparation) and the shortest autosome, located beside the X, are not involved in the translocations. Ends of chromosomes are marked “E.” ing yellowish-brown; scutellum dark brown; anterior scutellars slightly divergent; halteres dusky yellowish. Anterior sternopleural 14 posterior sternopleural; mid- sternopleural thin, %4 anterior. Legs yellowish; 2 golden hairs proximal end basal metatarsus of leg 3. Wings brown; veins darker brown; posterior crossvein clouded; costal index, 3.1; 4th vein index, 1.5; 5x index, 1.25; 4c index, 0.75; Heavy bristles on the basal 14 of third costal section; one prominent hair at apex of first costal section. Abdomen ( ¢) yellowish brown with shining black apical bands on ter- gites 2-6 extending almost to lateral margins, with indistinct medial interruptions on tergites 2, 3, 4; apical band thickened medially on tergites 5, 6. Anal plates yellowish; flattened laterally to form a keel shaped projection; numerous bent PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 113 hairs on ventral margin. Sternites pale gray except most posterior one which is yellowish. Female abdomen yellowish brown; tergites 2, 3, 4, 5 with black shining apical bands fading at lateral margins; paramedial extensions on these tergites; tergite 6 with black medial trapezoid shaped mark; tergite 7 yellow; sternites pale. Body length (etherized) ¢ 2.5 mm; 2 3 mm. Wing length ¢ 2.25; 2 3 mm. Internal characters of imagines and genitalia: Anterior malpighian tubules branched basally at a distance the width of intestine, free, with distal ends turned back; posterior malpighian tubules apposed with continuous lumen; testes pale yellow; 3 inner coils (vas deferens) and 4 outer coils (testis proper); sperm pump with 2 posterior diverticula each scarcely the length of the greater diameter of the pump. Forceps with 11-13 primary teeth arranged in a sinuous row; no secondary teeth; the two forceps joined by a medially grooved plate; about 8 marginal bristles and 7 bristles on the upper surface of the forceps; toe rounded with about 9 bristles. A long medially directed bristle on the posterior border of each concha of the hypandrium. Apodeme of the phallus a slightly bent rod; head of phallus simple, bulbous distally with ventrally attached laterally project- ing ears. Spermatheceae with distal portion dark brown, less chitinized proximally; oval, no constriction on inner duct; ventral receptacle thin, tightly coiled, with about 20 gyres. Ovipositor golden brown, acuminate at tip; 19 teeth. Egg: with 4 slender filaments, each 0.7 the length egg. Puparium golden brown; aperture of anterior spiracles ringed with black stigmatic plates bearing about 27 amber filaments; horns including anterior spiracles a little less than 1% the length of the puparium; posterior spiracles black; apart. Chromosomes: Laboratory culture 3L5 shows salivary chromosomes with a medium length X, markedly heterochromatic at the right hand end; one long autosome, three medium length autosomes, and a very short autosome; chromo- center scant. Salivary chromosomes of hybrids between this species and its sibling, D. metzii (Barro Colorado Island strain), show the presence of translocations in- volving the 4 long autosomes and a lack of pairing of portions of all homologues (Fig. 1). Belongs in subgroup IV of the tripunctata species group of the sub- genus Drosophila, being a sibling of D. metzii Sturtevant 1921 and D. pellewae Pipkin and Heed, 1964. D. leticiae differs from metzii and pellewae in the distinctly smaller male body length, absence of sexual dimorphism of abdominal coloration, and possession of fewer primary teeth of forceps. D. leticiae further differs from metzii in the yellowish brown color of the female face, less chalky aspect of the male face; from pellewae in the white face of the male. Reciprocal crosses of D. leticiae and D. metzii yield some viable hybrids. Many of the hybrids were fertile inter se, but laboratory populations derived from them usually died out in F, or F3. D. pellewae, when used as the female par- ent, produced few or no progeny with D. leticiae males, though hybrids were obtained from the reciprocal cross. The latter were sometimes fertile inter se, but laboratory populations derived from them usually die out in F» or Fs. Distribution: Holotype ¢; 12 6, 12 2 paratypes from laboratory 1l4 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 culture 3L5 developed from 15 founder females netted over fallen cacao fruit in house yard on the Amazon River at El Marco, Brasil, near Leticia, Colombia, June 16-20, 1964 (U.S. National Museum); 10 ¢ and 10 & paratypes from laboratory stock 3L5, same data, (Drosophila Type and Reference Collection) Austin, Texas. REFERENCES Pipkin, S. B. and W. B. Heed. 1964. Nine new members of the Drosophila tripunctata species group (Diptera: Drosophilidae). Pacific Insects 6: 256-273. Sturtevant, A. H. 1921. The North American Species of Drosophila. Carnegie Institution of Washington Publication 301, New Era Printing Co., Lancaster, Pa. p. 78. NEW SPECIES OF DOLICHOPODIDAE FROM THE UNITED STATES AND MEXICO (DireTera ) Haroip Rosrnson, Dept. of Botany, Smithsonian Institution, Washington, D. C. The present paper is primarily intended to supplement the synopsis of the Dolichopodidae of the Southeastern United States (Robinson, 1964), and includes those undescribed species from the Eastern U.S. which have been encountered since that study. A number of species from the Western U.S. and some related or otherwise interesting spe- cies from Mexico and Central America are also described. Dolichopus crassitibia, n. sp. (Figs. 1-4) Male.—Length 3.0 mm; wing 3.0 mm by 1.0 mm. Face about a fourth as wide below as high, becoming gradually wider above, covered with silvery white pollen; front broad, broader above, metallic greenish, almost completely covered with white pollen: palpus small, pale; proboscis brown. Antenna black; segment 1 short, widened distally from a narrow base, with hairs above; segment 2 shorter and wider, very short below; segment 3 about as long as wide, blunt; arista from near middle of dorsal edge. Lower postocular setae pale and somewhat flattened. Thorax metallic green, slightly dulled with grayish pollen, a violet band be- tween rows of small acrostichals; 6 pairs of large dorsocentrals, 5th pair set in- ward from the rest of the line; scutellum with large pair of bristles and small erect hair on the lateral margin; a black bristle over the fore coxa. Legs black with black setae, all knees, tips of fore and middle tibiae, inner surface of fore tibia, and base of fore metatarsus yellowish. Fore and middle coxae with numerous bristles on anterior surface, distal ones large; middle and PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 115 Fig. 1-17. Dolichopodidae, males. Dolichopus crassitibia n. sp. 1, hind tibia; 2, wing; 3, hypopygial lamella; 4, hypopygium. Rhaphium steyskali n. sp. 5, an- tenna; 6, hypopygium. Diaphorus pseudopacus Robinson. 7, hypopygium; 8, tips of hypopygial appendages, ventral view. Diaphorus canus Robinson. 9, hypopyg- ium; 10, hypopygial appendages, ventral view. Diaphorus bakeri n. sp. 11, hy- popygium; 12, hypopygial appendages, ventral view. Diaphorus gibbosus Van Duzee; 13, hypopygium; 14, tips of hypopygial appendages, ventral view. Chryso- tus clypeatus n. sp. 15, hypopygium; 16, face and mouth, palpi turned edgewise; 17, antenna. 116 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 hind coxae with a large external bristle; middle and hind femora with a preapical anteriorly, hind femur with a few long pale hairs along middle of lower posterior surface; fore tibia with a pair of indistinct dorsals near base, a second pair near the middle, of which the posterodorsal is rather large, 2 small posterodorsals and 2 larger apicals; middle tibia with 2 large anterodorsals and 1 small posterodorsal in basal third, 1 anterodorsal, 1 posterodorsal, and 1 ventral near distal third, 5 apicals; hind tibia (fig. 1) greatly thickened, with 3 anterodorsals of which 2 including a small one are near the base, 4 large posterodorsals, numerous indis- tinct ventrals, 2 apicals. Lengths of segments of fore tarsus from the base as 8-4-3-2-2; middle tarsus as 10-5-4-3-3; hind tarsus as 8-8-6-4-3, segment 1 very thick with 2 large bristles above. Wing (fig. 2) rather oblong with a prominent anal angle and a large sinus behind the crossvein, hyaline with veins brown. Costa swollen at juncture of vein 1; veins 2 and 4 nearly parallel beyond the crossvein, vein 4 with slight double bend before middle of last part, vein 3 converging with vein 4 toward the tip; crossvein about as long as last of vein 5. Calypter, its setae, and knob of halter pale, stem of halter brown. Abdomen about as long as thorax, tapering, shining metallic green dulled with grayish pollen on sides. Hypopygium (fig. 4) two-thirds as long as preabdomen, black with greenish tinges; lamellae (fig. 3) rounded, whitish with a jagged brown border; inner appendages pale. Holotype ¢, from wasp nest, on shore of Potomac River by Plummers Island, Montgomery Co., Maryland, June 16, 1963, K. Krombein. Spec- imen in USNM, no. 69332. This is the “Dolichopus n. sp.” listed as prey of Ectemnius (Hypocrabro) paucimaculatus (Packard) by Krom- bein (1964, p. 86). The species would key to the group of species containing Dolichopus kansensis, D. ovatus, and D. acuminatus using my key to the south- eastern species. The new species differs from these and from most other species of Dolichopus by the greatly thickened hind tibia and the deep sinus in the hind margin of the wing. Rhaphium steyskali, n. sp. (Figs. 5-6) Male.—Length 5.3 mm; wing 4.0 mm by 1.5 mm. Face long and narrow, becoming wider above, covered with white pollen; front broad, slightly wider above, metallic green with thin whitish pollen; palpus ap- parently yellow with pale hairs; proboscis brown. Antenna (fig. 5) brown with segment 1 yellow below; segment 1 widened distally to as wide as long, bare above; segment 2 very short, about as wide as first, ringed by submarginal black setulae; segment 3 about as long as face, elliptical, covered with fine pale hairs, with darker strand along dorsal margin ending at insertion of arista; arista slightly preapical, about as long as segment 3. Upper postocular setae black, lateral and lower postocular surface covered with long white hairs. Thorax metallic green with mesoscutum very shining, thin whitish pollen evi- dent toward margins and on pleura. Setae black; acrostichals short, biseriate; 5 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 LILY) pairs of long dorsocentrals; scutellum with a pair of long widely-separated bristles, a smaller bristle on each lateral margin; proepisternum covered with short pale hairs. Coxae yellow, base of middle coxa slightly darkened externally, segments of fore and middle tarsi with brownish tips, dorsal margin of hind femur darkened distally, hind tibia brownish yellow and becoming brownish distally, hind tarsus dark brown, legs otherwise pale. Fore and middle coxae with numerous pale hairs on anterior surface, fore coxa without black setae, middle coxa with a cluster of black setae forming an apical thorn, middle and hind coxae with a long black external seta; posterior surface of fore femur with many rather long pale hairs, some pale hairs on hind surface of middle femur toward base, other setae of legs mostly black; femora without distinct preapicals; fore tibia with pair of small dorsals near basal fourth, small posterodorsal near middle, 1 distinct apical; middle tibia with 1 stout anterodorsal near base, smaller one beyond middle, 1 small posterodorsal near middle, 5 apicals; hind tibia with pair of short stout dorsals near middle, 1 anterodorsal near base, 1 posterodorsal near second fifth, 2 strong apicals. Lengths of segments of fore tarsus from base as 12-4-3-2-2; middle tarsus as 15-7-5-3-2; hind tarsus as 10-12-8-5-3. Wing elongate oval, hind margin rather oblique distally, surface tinged with brown especially distally, veins brownish; vein 2 reaching twice as far along costa as vein 1, nearly straight; vein 3 nearly straight and parallel with 2 but curving gradually backward near tip; vein 4 slightly offset forward in middle of last part, nearly parallel with 3 at tip; crossvein nearly perpendicular to base of vein 5, two-thirds as long as last of vein 5; vein 6 represented by a strong fold that bends backward in middle. Calypter, its long setae, and halter pale. Abdomen considerably longer than thorax, tapering, metallic green with slight whitish pollen on sides; long pale setae on sides of basal terga, long black setae along hind margins. Hypopygium (fig. 6) brown, rather small, extending slightly forward under preabdomen, tips prolonged into smooth light brownish rods; lamel- lae strap-like, acute apically, brown, with dark setae over outer surface, a few pale setae at tip. Holotype ¢, Sweetwater Creek near old “Camp Torreya,” Liberty Co., Florida, Mar. 23, 1954, George Steyskal. Specimen in USNM, no. 69333. The new species would key to Rhaphium insolitum Curran but dif- fers by having a much more massive third antennal segment and a much longer strap-like hypopygial lamella. The new species is the only Rhaphium known to me having a preapical arista. Diaphorus bakeri, n. sp. (Figs. 11-12) Male.—Length 2.7 mm; wing 2.7 mm by 1.1 mm. Face about as wide as long; front triangular below, obliterated in upper half; both face and front brown with slight grayish pollen; palpus brown with a few dark hairs and a black apical seta; proboscis brown. Antenna brown, all segments of nearly equal length; segment 1 distally about as wide as long, bare above; segment 2 slightly wider than first, ringed by small setulae of which one above 118 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 / Fig. 18-31. Dolichopodidae, males. Chrysotus picticornis Loew. 18, hind femur; 19, hypopygium. Chrysotus mexicanus n. sp. 20, hind femur; 21, hypopyg- ium. Chrysotus neopicticornis n. sp. 22, hind femur; 23, hypopygium. Telma- turgus costaricensis n. sp. 24, antenna; 25, fore tarsus. Medetera marylandica n. sp. 26, wing; 27, hypopygial appendages; 28, tip of aedeagus, ventral view. Chryso- timus arizonicus n. sp. 29, hypopygium. Chrysotimus obscurus n. sp. 30, hypo- pygium. Chrysotimus metallicus Parent. 31, hypopygium. is rather long; segment 3 nearly twice as wide as long, covered with numerous fine pale hairs, with arista inserted near upper margin and with distal margin slightly produced below arista. Lower postocular surface covered with long black hairs. Thorax brown, with slight yellowish pollen above, more grayish pollen on pleura. Mesoscutum rather strongly arched; bristles black; acrostichals small, biseriate; 5 pairs of dorsocentrals; 1 pair of long scutellars, a small seta on each lateral margin; a few small setae above the fore coxa. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 119 Legs brown. Hairs and bristles black; fore and middle coxae with numerous black hairs on the anterior surface, fore coxa with larger setae on distal margin, middle and hind coxae with a large external bristle; fore femur with hairs on ven- tral surface of thickest part rather erect, with a row of prominent setae along anteroventral surface, hind femur with numerous long hairs along posteroventral surface and a few stiff hairs anteroventrally near tip, preapical bristles lacking; fore tibia with 1 very small anterodorsal near basal third; middle tibia with 2 anterodorsals and 2 posterodorsals all small, 4 large apicals; hind tibia with 2 small but distinct anterodorsals, 4—5 posterodorsals, 4 apicals. Lengths of segments of fore tarsus from base as 14-7-5-4-3, with tibia as about 23, pulvilli nearly as long as segment 5; middle tarsus as 16-8-6-4-3, pulvilli scarcely enlarged; hind tarsus as 11-9-6-4-3, pulvilli not enlarged. Wing rather oval with basal part very broad, hind margin only slightly curved from middle to near tip of vein 4; surface with brownish tinge and brown veins; vein 1 reaching about half way from base to tip of vein 2; veins 2, 3, and 4 nearly straight and parallel beyond the crossvein, vein 3 only slightly more curved backward toward the tip; crossvein perpendicular to basal part of vein 4, about three-fifths as long as last of vein 5; vein 6 represented by slight fold. Calypter and halter brownish, setae of former black. Abdomen longer than thorax, brown with slight yellowish gray pollen toward the sides and base; hairs and bristles black, longer toward the sides. Hypopygium (figs. 11, 12) brown, small, capping tip of preabdomen, with 4 large bristles projecting posteriorly from near base; hypopygium bearing straight slender arms with a swelling on middle of inner surface, a flange above forming a short hood, with a narrow setiferous appendage and a median hooked appendage between the arms reaching about half of length of arms. Holotype ¢, from Olga, Washington, July 9, 1926, C. F. Baker. Spec- imen in USNM, no. 69334. The new species would key to Diaphorus nigricans in the treatment of southeastern Dolichopodidae, but differs most noticeably by the presence of prominent setae posteroventrally on the hind femur. These setae plus the longer first wing vein provide distinctions from D. fuscus, and the lack of enlarged pulvilli on the middle and hind tarsi provide a further distinction from D. snowi. I have included illustrations of hypopygia of some of the related species having yellow tibiae; D. pseudopacus Robinson, figs. 7-8; D. canus Robinson, figs. 9-10; D. gibbosus Van Duzee, figs. 13-14. Chrysotus neopicticornis, n. sp. (Figs. 22-23 ) Male.—Length 2.1 mm; wing 2.1 mm by 0.9 mm. Eyes essentially contiguous below the middle of the face, face narrow above mouth, covered with yellowish pollen; front broad, slightly broader above, me- tallic green with slight yellowish pollen; palpus yellow; proboscis yellowish brown. Antenna with first and most of second segment yellow, upper part of second and most of third segment brown; segment 1 bare above; segment 2 about as long as 120 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 but distally wider than first, with a submarginal ring of dark setulae; segment 3 longer than second, half again as long as wide, with arista in slight apical sinus. Lower postocular setae pale. Thorax metallic green dulled with yellowish pollen above, with thicker whitish pollen on pleura. Setae black; acrostichals biseriate; 5 pairs of dorsocentrals; a pair of large scutellars, a small scutellar hair on lateral margin; a pale seta above fore coxa. Legs mostly pale, middle coxa and base of hind coxa brownish. Setae mostly dark; many long pale setae on anterior surface of fore and middle coxae, hind coxa with a rather dark external bristle; fore femur with a series of prominent posteroventrals (distal ones longer), ventral setae paler toward base of femur; middle and hind femora with rather short pale ventral setae, those of hind femur evenly spaced, hind femur (fig. 22) with 2-3 brownish anteroventral bristles near tip; middle tibia with 1 strong anterodorsal and 1 weak posterodorsal near basal third, a very small bristle in each dorsal row beyond middle, 3 distinct apicals; hind tibia with 2 rather strong anterodorsals and 3 smaller posterodorsals rather evenly spaced, 2 apicals and 1 subapical. Lengths of segments of fore tarsus from base as 8-4-3-2-2; middle tarsus as 9-4-3-2-3; hind tarsus as 7-5-4-3-3. Wing rather oblong oval, clear with brownish veins; veins 3 and 4 nearly parallel beyond crossvein, vein 2 only slightly diverging; crossvein perpendicular to last of vein 4, almost half as long as last of vein 5; vein 6 represented by a slight fold. Calypter and halter pale, setae of former pale brownish. Abdomen only slightly longer than thorax, metallic green with slight pollen which is more yellowish above; setae mostly short with rather pale reflections, a few slightly longer and darker near tergal margins. Hypopygium (fig. 23) small, enclosed in tip of preabdomen; outer appendages small, pale; inner appendages primarily a pair of stout dark armatures with a blunt tip bearing 2 short but dis- tinct setae. Face of female wide, about half as wide as high; palpus with numerous black hairs. Setae on anterior surface of fore coxa much shorter, no long or distinctive setae ventrally on femora; setae of hind tibia generally stronger. Holotype ¢ and allotype °, from soil of earth dam, wooded area about 3 miles north of Spartanburg, South Carolina, May 8, 1961; 2 4 paratypes, soil by stream, University of Tennessee Farm, Knox Co., Tennessee, July 18 and Aug. 11, 1957, all H. Robinson; 1 2 paratype, Duke Garden, Durham Co., North Carolina, April 28, 1959, H. S. Daoud; 1 ¢, Canton, New York, July 27, 1931; 2 ¢, Vincennes, Indiana, June 6; 2 6, Lafayette, Indiana, June 1 and 27, all J. M. Aldrich. Holo- type (no. 69335), allotype, and the Aldrich collections in USNM; others presently in my collection. For discussion see following species. Chrysotus mexicanus, n. sp. (Figs. 20-21 ) Male.—Length ca. 2.0 mm; wing ca. 2.0 mm by 0.9 mm. Head, thorax, legs except hind femur, wing, and preabdomen essentially as PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 121 described for C. neopicticornis. Ventral margin of hind femur (fig. 20) with sparse pubescence, at base bearing a pair of distinct black setae that are very close and often adhere to each other. Hypopygium as illustrated (fig. 21). Holotype 4, allotype °, 6 4, 2 2 paratypes, from km 375, rt. 180, Veracruz, Aug. 7; 2 2 paratypes, soil by stream, near Santiago Tuxtla, Veracruz, Aug. 8, 1962; 1 ¢ and 2 2 paratypes, near Tierra Blanca, Veracruz, May 12, 1963; 3 é and 2 2 paratypes, edge of shaded stream, Tuxtepec, Oaxaca, May 12; 1 2, above Rio Valle Nacional, Oaxaca, May 14, 1963; 1 ¢, near Jaltepec River, rt. 185, Oaxaca, Aug. 8; 3 ¢ and 1 2, near km 220, rt. 185, Oaxaca, Aug. 9, 1962; 4 ¢ and 1 2, El Salto, San Luis Potosi, May 9; 7 ¢ and 1 2, near junction of rts. 190, 200, Chiapas, May 21; 2 4 and 1 2, just below crest of Sierra above Arriaga, Chiapas, May 22; 1 ¢, Rio Lajas, Chiapas, May 23, 1963; all from Mexico collected by H. Robinson; 3 ¢ and 1 2, Clarke Hall, Dominica, Jan. 22-23; 2 6, near Layou River mouth, Mar. 10, 1964, H. Robinson; 6 é and 3 2, Layou River mouth, Jan. 9; 1 ¢, same loca- tion, Mar. 8; 5 ¢ and 1 2, same location, Mar. 18-24; 7 6 and 2 2, Cabrit Swamp, Mar. 22-25; 4 6 and 1 2, Clarke Hall, Malaise and light traps, Jan. 8—Mar. 31; all 1964 collected by W. W. Wirth; 1 ¢, Clarke Hall, May 28; 1 6, Melville Hall Airport, July 14, 1966, George Steyskal; the latter all from Dominica collected during participation in the Bredin—Archbold-Smithsonian Biological Survey of Dominica. Holotype (no. 69336), allotype, and the Dominica collections in USNM,;; others presently in my collection. After careful examination of the Chrysotus picticornis complex I find three distinct species are involved. Those specimens having a distinct dark spot and four or more distinct black bristles near the tip of the hind femur and rather dense pale pubescence ventrally near the base of the femur, are C. picticornis. In this species the hypopygial arma- ture is very slenderly pointed. The two new species lack the distinctly dark tip on the hind femur and the anteroventral preapical setae are less distinct (sometimes only 1 present). Of the two new species, C. mexicanus is easily distinguished by the pair of black ventral bristles at the base of the hind femur. The hypopygial armature of C. mexi- canus seems rather variable, in one specimen from Chiapas (near Arriaga) the armature is very broad and is curved on the lower edge, in specimens from Dominica the armature is very slender. Though the range of C. picticornis in the Neotropics is not as extensive as pre- viously believed, I have collected specimens near Jitotol in Chiapas, Mexico. Chrysotus clypeatus, n. sp. (Figs. 15-17) Male.—Length 2.0 mm; wing 2.1 mm by 0.7 mm. Face (fig. 16) very broad, becoming broader below, terminating at suture with 122, PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 the broad and bluntly pointed clypeus greatly recessed from the plane of the face, covered with silvery white pollen; front slightly broader than upper part of face, dark brownish with thin yellowish pollen; palpus rounded-oblong, nearly as long as face, covered with silvery white pollen; proboscis small, brown. Antenna (fig. 17) brown; segment 1 very small, bare above; segment 2 larger but hardly longer, ringed by short brownish setulae; segment 3 triangular, slightly longer than wide, covered with fine pale pubescence, with arista borne just before apex. Lower postocular surface with many pale setae. Thorax dark brownish, with slight yellowish pollen above, whitish pollen toward sides and on pleura. Setae black; acrostichals small, biseriate; 6 pairs of dorso- centrals, hind pair long; 1 pair of long scutellars, a small seta on lateral margin of scutellum; 1-2 pale setae above fore coxa. Legs brown with fore and middle trochanters, all tibiae, and all but tips of basitarsi pale. Fore and middle coxae with pale setae on anterior surface; middle coxa with a brownish setae toward outside, hind coxa with black external seta; femora without preapical setae, a few prominent antero- and posteroventrals near tip of hind femur; fore tibia without distinctive setae; middle tibia with 1 strong anterodorsal near base, 1 very small ventral, 4 apicals; hind tibia with 2 small anterodorsals, 4 small posterodorsals, 4 stronger apicals. Lengths of segments of fore tarsus from base as 9-4-3-2-2; middle tarsus as 11-5-3-2-2; hind tarsus as 8-6-4-2-2. Wing rather oblong, clear, veins brown; veins 3 and 4 parallel, curving slightly backward; crossvein near midwing, perpendicular to anterior wing margin, about two-thirds as long as last of vein 5. Abdomen cylindrical, slightly longer than thorax, dark metallic brownish green; marginal setae of tergum 6 projecting, but not longer than setae of other terga. Hypopygium (fig. 15) small, embedded in tip of preabdomen, bare, brown; lamel- lae minute, brownish, pale pubescent. Holotype ¢, Port St. Joe, Gulf County, beach, Florida, Mar. 17, 1954, George Steyskal. Specimen in USNM, no. 69337. The new species seems most closely related to Chrysotus lamellifer Robinson, which has the same type of unusual clypeus, but C. lamel- lifer is a somewhat larger species with a short rather reniform third antennal segment and with a distinct ventral bristle on the hind basi- tarsus. Telmaturgus costaricensis, n. sp. (Figs. 24-25) Male.—Length 1.8 mm; wing 2.0 mm by 0.8 mm. Face very narrow, essentially obliterated in lower half; front broad, broader above, dark with violet reflections, with very slight pollen; palpus and proboscis brown. Antennal segments (fig. 24) 1 and 2 small, yellowish; segment 3 brown, triangular, half again as long as wide; arista dorsal from near base of third seg- ment, plain. Postocular setae small, brown. Thorax evenly rounded above, dark brownish above with violet reflections, only slightly pollinose; pleura yellowish. Acrostichals uniseriate, absent posteriorly; 4 pairs of large black dorsocentrals, posterior pairs largest, an anterior 5th pair very small; scutellum with 1 pair of bristles, no hairs. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 123 Legs yellowish with dark setae. Middle femur with preapical bristles anteriorly and posteriorly; hind femur with both upper and lower preapicals anteriorly; middle tibia with antero- and posterodorsals paired near basal third, 1 antero- dorsal near distal third, 2 apicals; hind tibia with a few short indistinct dorsals, 2 very small ventrals. Lengths of segments of fore tarsus (fig. 25) from base as 8-6-5-5-3, segment 1 slightly thickened, all segments bearing very long curved setae along antero- and posteroventral surfaces; middle tarsus as 11-9-6-4-4; hind tarsus as 6-11-7-4-3. Wing elongate oval, slightly brownish tinged, veins brown; veins 3 and 4 essen- tially parallel beyond the crossvein, vein 2 diverging slightly forward; crossvein a little over half as long as last of vein 5; vein 6 represented by slight fold. Calypter, its setae, and knob of halter brown. Abdomen slightly longer than thorax, cylindrical, brownish with slight violet reflections; setae dark. Hypopygium small, set on tip of preabdomen, brown with small brown appendages. Holotype ¢, La Suiza de Turrialba, Costa Rica, April 1922, Pablo Schild. Specimen in A. L. Melander collection in USNM, no. 69338. The new species differs from Telmaturgus parvus (Van Duzee) of the Eastern United States by the presence of 4 pairs of dorsocentrals; it differs from T. tumidulus (Raddatz) of Europe by the plain arista and the differently shaped fore tarsus. Lamprochromus occidentalis, n. sp. Male.—Length 2.0 mm; wing 2.0 mm by 0.8 mm. Setae mostly black. Face wide above, gradually becoming very narrow below, covered with silvery white pollen; front broad, broader above, shining dark green; palpus small, brown; proboscis brown. Antenna black, all segments short and about as long as wide; segment 1 bare above; segment 2 with a ring of setulae and 1 long dorsal seta; segment 3 pointed, with a dorsal arista. Lower postocular setae pale. Thorax bright metallic green with a violet stripe along middle of mesoscutum, a patch of velvety black above wing base; grayish pollen on pleura. Acrostichals biseriate; 4 pairs of dorsocentrals; 1 pair of strong scutellars, a small hair on lateral scutellar margin; proepisternum with a few pale setulae before anterior spiracle. Middle and hind coxae slightly brownish basally, fore and middle tarsi rather darkened distally, hind leg brown except basal third of femur, legs otherwise pale. Fore coxa with numerous small pale hairs and distally with black setae on anterior surface; middle and hind coxae with a large seta toward the outer surface; middle and hind femora with preapical bristles; fore tibia with 1 small anterodorsal near base; middle tibia with 2 small anterodorsals, 2 larger posterodorsals, 4 apicals; hind tibia with 2 strong anterodorsals, many large posterodorsals of which some are quite prominent, 3 apicals. Lengths of segments of fore tarsus from base as 9-4-3-2-2: middle tarsus as 11-7-5-3-3; hind tarsus as 9-10-5-3-5. Wing rather oval, slightly brownish; veins 2, 3, and 4 curving only very slightly backwards, veins 3 and 4 essentially parallel beyond crossvein; crossvein perpen- dicular to base of vein 4, two-thirds as long as last of vein 5; vein 6 represented by a slight fold; anal angle only moderately rounded. 124 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 > > ? > Abdomen longer than thorax, cylindrical; wholly metallic green. Hypopygium capping tip of preabdomen, dark with small brownish appendages. Holotype ¢, from Wells, Nevada, July 12, 1911; 1 ¢ paratype, from Juliaette, Idaho, July 5, 1930; both J. M. Aldrich. Specimens in USNM, no. 69339. The new species differs from others known from North America by the wholly metallic abdomen. It differs from European species by the short pointed antenna and the almost wholly brownish hind leg. Medetera marylandica, n. sp. (Figs. 26-28) Male.—Length ca. 2.0 mm; wing 2.0 mm by 0.7 mm. Face narrow below, metallic dark green, slightly dulled by yellowish white pollen; front broad, broader above, slightly more pollinose than face; head strongly excavated behind vertex; palpus and proboscis black. Antenna blackish, all seg- ments short, segment 3 truncate with an apical arista. Postocular setae black with pale reflections. Thorax strongly arched with strongly flattened posterior slope, dark metallic green dulled with grayish pollen. Acrostichals biseriate, absent posteriorly; 4 pairs of large black dorsocentrals becoming progressively smaller and grading into hairs anteriorly; 4 scutellar bristles, lateral pair small; proepisternum with a few small black setae below. Legs with fore and middle knees and tibiae, basal joints of fore and middle tarsi yellow, hind tibia yellowish brown, remainder of legs brown. Setae brown- ish, all but larger with pale reflections; middle femur with numerous longer setae along anteroventral surface in distal half; hind femur with 2 bristles toward tip in middle of anterior surface; middle tibia with small anterodorsal and smaller posterodorsal paired near basal third. Lengths of segments of fore tarsus from base as 10-4-3-2-4; middle tarsus as 14-6-4-3-2; hind tarsus as 8-10-6-3-4, basi- tarsus with a deep prominently rimmed pocket basally on inside surface. Wing (fig. 26) rather oval, clear, veins brown; vein 2 and last of vein 4 nearly parallel, vein 3 curving backward and ending near vein 4; base of vein 5 thick- ened for nearly two-thirds its length; crossvein slightly over half as long as last of vein 5; vein 6 represented by slight fold. Calypter and halter pale, setae of calypter brownish. Preabdomen about as long as thorax, strongly tapered, metallic blackish green with slight yellowish white pollen; terga bearing small black marginal bristles. Hypopygium pedunculate, black; appendages (fig. 27) yellow, lateral lamella deeply cleft, aedeagus (fig. 28) slender with a preapical enlargement. Holotype ¢, Bethesda, Montgomery Co., Maryland, July 24, 1965, George Steyskal. Specimen in USNM, no. 69340. The new species seems closest to Medetera freyi E. Thuneb. of Eu- rope in the shape of the genitalia, more like M. excellens Frey in leg color. Of these the former has a much shorter thickened area on the fifth vein and the latter has much paler pubescence. The only pre- viously known North American species with a thickened fifth vein, M. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 125 crassivenis Curran, has the last part of the fifth vein hardly half as long as the crossvein, and has a hind femur with more and longer hairs anteriorly but no bristles. Chrysotimus arizonicus, n. sp. (Fig. 29) Male.—Length 2.0 mm; wing 2.6 mm by 1.0 mm. All hairs and bristles pale. Face broad with sides nearly straight and parallel, metallic green with slight gray pollen; front broad with sides diverging from below, metallic green with yellowish pollen; palpus yellow with a number of hairs; proboscis brown. Antenna black, with all segments short; segment 1 bare above; segment 2 ringed with small setulae; segment 3 triangular, covered with small hairs, with arista inserted near middle of the oblique dorsal margin. Lower postocular setae in single series. Thorax metallic green except the yellow metepimeron, with yellowish pollen above, more grayish pollen on pleura; posterior slope of mesoscutum flattened. Acrostichals small, irregularly biseriate; 5 pairs of large dorsocentrals; 1 pair of long scutellars, a small seta on lateral scutellar margin; proepisternum with a few short setae above fore coxa. Legs yellow except the black fifth segments of the tarsi. Fore and middle coxae with numerous hairs on anterior surface, fore coxa with a series of bristles on distal margin; middle and hind coxae with a strong external bristle; middle and hind femora with a preapical bristle; fore tibia without evident bristles; middle tibia with 2 anterodorsals near base and middle, a smaller posterodorsal near each, 3 apicals; hind tibia with 2 anterodorsals and 3 posterodorsals, 2 small but dis- tinct apicals. Lengths of segments of fore tarsus from base as 9-4-3-2-2; middle tarsus as 1]1-5-3-2-2; hind tarsus as 8-6-4-3-3. Wing rather oval, clear with yellow veins; veins 2, 3, and last part of 4 all very slightly curved backwards and nearly parallel to each other and to front wing margin, vein 3 slightly more curved toward tip, vein 4 ending at or just behind tip of wing; crossvein perpendicular to base of vein 4, two-fifths as long as last of vein 5; vein 6 represented by a slight fold. Calypter and halter yellow. Abdomen only slightly longer than thorax; shining metallic green terga dulled with slight yellow pollen, sterna yellow. Hypopygium (fig. 29) brown, apparently usually partly concealed, extending forward nearly half length of preabdomen; appendages brown. Female very similar to male except in abdomen. First 4 segments of abdomen yellow, segment 5 broad and metallic green; middle of hind margin of tergum 4 slightly indented. Holotype ¢, allotype 2, and 1 2 paratype, from Grand Canyon Na- tional Park (north rim), Arizona, July 15, 1954, W. L. Downes, Jr. Holotype (no. 69341) and allotype in USNM, paratype in the Downes collection at the Department of Entomology, University of Illinois. The new species seems close to Chrysotimus luteopalpus Curran, but the antenna of the latter species is mostly yellow and the female abdomen has the fifth tergum yellow and margins of the other terga plain. 126 PROC, ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Chrysotimus schildi, n. sp. Male.—Length 1.5 mm; wing 2.0 by 0.8 mm. Hairs and bristles mostly yellow, verticals, ocellars, and anterior dorsocentrals brownish. Basic characters as in C. arizonicus but antennae yellow with an apical arista; only center of mesoscutum bright metallic green, thorax otherwise yellow; hind tibia with only 1 obvious anterodorsal, posterodorsals very small and indistinct; lengths of segments of fore tarsus from base as 13-6-5-3-3, middle tarsus broken off, hind tarsus as 11-9-6-4-3; wing vein 4 ending slightly behind wing tip; abdo- men yellowish on lateral margins of terga, pale brownish below; hypopygium brownish with greenish reflections, of structure similar to C. obscurus & C. metal- licus, bases of appendages yellow. Female abdomen with tergum 1, all but central posterior triangle of tergum 3, and sides of tergum 4 yellow. Holotype é and allotype ?, La Suiza de Turrialba, Costa Rica, April, July 1922, Pablo Schild. Specimens in A. L. Melander collection in USNM, no. 69342. The species is readily distinguished by the dorsal spot of metallic green on the mostly yellow thorax. Chrysotimus obscurus, n. sp. (Fig. 30) Male.—Length 1.8 mm; wing 2.3 by 1.0 mm. Similar to C. arizonicus but hairs and bristles of body brownish, those of legs more yellowish; head and thorax including metepimeron much darker metallic green, abdomen brownish with green reflections (hypopygium as in fig. 30); front and mesoscutum with light grayish pollen; palpus black; antennal segment 3 very blunt with an essentially apical arista; acrostichals distinctly biseriate; coxae ex- cept tips and hind femur except extreme base and tip brown; middle tibia with 2nd anterodorsal near distal third, 1-3 posterodorsals; lengths of segments of fore tarsus from base as 8-4-3-2-2, middle tarsus as 10-5-4-2-2:; hind tarsus as 8-6-4-3-2; wing veins brownish yellow, vein 4 ending slightly behind wing tip, crossvein per- pendicular to last of vein 4; setae of calypter brownish. Holotype ¢ and 1 ¢ paratype, from roadside foliage, rain forest, Sierra Juarez above Rio Valle Nacional, Oaxaca, Mexico, May 17, 1963, H. Robinson. Holotype in USNM, no. 69343; paratype presently in my collection. Chrysotimus obscurus seems close to such species as C. pusio Loew and C. metallicus Parent, but is distinct by the brownish coxae and hind femur and the blunt third antennal segment with an apical arista. Chrysotimus metallicus has brownish bristles as in C. obscurus, but the anterior wing margin is more curved and the second and fourth veins are less distinctly parallel. The female of C. pusio has a partly yellow abdomen as is common in the genus, but C. obscurus may prove to be more related to C. metallicus, in which the female abdomen is entirely metallic green. 1 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 12 REFERENCES Krombein, Karl VY. 1964. Natural History of Plummers Island, Maryland. XVIII. The Hibiscus wasp, an abundant rarity, and its associates (Hymenoptera: Sphecidae). Proc. Biol. Soc. Wash. 77: 73-112. Robinson, H. 1964. A synopsis of the Dolichopodidae (Diptera) of the South- eastern United States and adjacent regions. Misc. Publ. Ent. Soc. Amer. 4: 103-192. FURTHER STUDIES ON THE TARSONEMIDAE ( AcARINA ) Ropert L. SMILEY Entomology Research Division, ARS, U. S. Department of Agriculture, Washington, D. C. 20250 In a continuation of studies on the Tarsonemidae (Smiley, 1964), a family of mites of importance to agriculture, I here describe one species belonging to the genus Hemitarsonemus Ewing, two species to the genus Steneotarsonemus Beer, and two species to the genus Tarsonemus Canestrini as new from material in the U. S. National Museum collec- tion. All are from the Western Hemisphere. Also included in this study are five new species of tarsonemids collected from various areas in the South Pacific and submitted for determination by D. M. C. Manson of the Horticultural Research Centre, Department of Agricul- ture, Levin, New Zealand. This collection is represented by the follow- ing genera: Hemitarsonemus Ewing (one species), Tarsonemella Hirst, and Tarsonemus Canestrini (four species). The genus Neotarsonemus is here erected to include the two species, Hemitarsonemus latus (Banks) and H. beeri Smiley in which legs IV of the males possess ter- minal claws which are reduced to a ball-like process. The genus Fungi- tarsonemus Cromroy (1958) is here synonymized with the genus Hemi- tarsonemus. Ewing (1939) and Beer (1954) give an excellent histori- cal account of the confusion and errors made previously in the genus Hemitarsonemus and perpetuated by former acarologists. It is hoped that this present generic arrangement will alleviate past and future errors. A key is presented for the males of species belonging to Hemitarso- nemus. Tarsonemella is reviewed, and additional morphological char- acters are supplied for the genus as well as for T. beameri Beer. 128 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 129 Steneotarsonemus spinki, n. sp. (Figs. 1-9) The male of this species is characterized by the presence of a pair of dagger- shaped setae on femur and genu IV, and a short, stout, blunt spurlike seta on tibia III. Male. Body elongate and broadest in anterior region of hysterosoma. Dorsal propodosomal setae simple, short, stout, slightly dagger-shaped; third pair of propodosomals longest, one-third longer than first pair; second pair shortest; fourth pair slightly longer than first pair. Hysterosoma with three rows of simple setae; first row with single pair of lateral setae; second row with two pairs of setae; third row with single pair of setae; setae of first row longest, setae of other rows more or less equal in length. Ventral propodosomal and hysterosomal setae sub- equal in length; coxae I with single pair of setae, slightly shorter than single pair on coxae II; coxae III with two pairs of setae subequal in length, and slightly longer than setae of coxae I and II; coxae IV with pair of dagger-shape setae. Ventral apodemes as figured. Legs I and II similar in size and length; genu I with a rodlike solenidion similar to solenidion on tibia II but without spurlike seta; legs III longest, with spurlike seta stouter and longer than spurlike setae of legs I and II; leg IV as figured. Femur with large inner median lateral flange; inner an- terior and outer median setae short and of equal length; posterior inner seta about as long as segment, strong, dagger-shaped; genu with daggerlike ventral seta simi- lar to that on femur, and with distal rodlike solenidion; tibia—tarsus short, with two tiny inner setae and stout ventrally curved claw. Body 217 yu long by 121 yu wide. The female associated with the above male is similar in having a long rodlike solenidion dorsally on tarsus I and a bifurcate spurlike seta on the distal ventral surface. Female. Body elongate, broadest in region of hysterosoma. Propodosomal shield trapezoidal, bearing two pairs of setae; first pair slightly serrate, about one-third as long as second pair, second pair about two thirds as long as shield; pseudo- stigmatic organs ovoid, bearing small spurlike projections, pedicel as long as ex- panded distal portion. Dorsum of hysterosoma with five distinct transverse seg- ments; first segment with two pairs of setae, outer pair simple and longer than the inner serrate pair; second segment without setae; third segment without simple setae but with two inner serrate setae subequal to the pair of the first segment; fourth segment without setae; fifth segment with two pairs of serrate setae. First pair of ventral apodemes Y-shaped, converging with the anterior median apodeme; a pair of simple setae adjacent to the first pair of apodemes; apodemes II longer and stronger, with a pair of simple setae which are shorter than the anterior pair; posterior median apodeme not converging with apodeme III and transverse apo- deme; apodeme III obscure, and above these, a pair of long simple setae; apodeme < Steneotarsonemus spinki, new species. Fig. 1, dorsum, male; fig. 2, venter, male; fig. 3, dorsum, female; fig. 4, venter, female; fig. 5, ventral, tibia and tarsus I, male; fig. 6, dorsal, tibia and tarsus I, male; fig. 7, dorsal, tibia and tarsus I, fe- Hae fig. 8, ventral, tarsus and tibia II, female; fig. 9, pseudostigmatic organ, emale. 130 PROG. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 131 IV converging with genital plate, and with a pair of simple setae subequal in length to the pair on apodeme II. Legs robust, as figured. Body 274 u long by 108 u wide. The male holotype, USNM 3148, and a female paratype were col- lected on Sogata orizicola Muir, a planthopper, Baton Rouge, La., No- vember 18, 1960, by W. T. Spink, for whom this species is named. Steneotarsonemus friedmani, n. sp. (Figs. 10-16) The large empodium on legs I-III and the ventral body striations will separate this species from other known members of the genus. Male. Body elongate oval, broadest in anterior region of hysterosoma. Propo- dosomal shield trapezoidal; with four pairs of simple setae, first pair shortest; second pair two-thirds length of third pair, slightly blunt apically; third pair longest; fourth pair subequal to second pair, saberlike and pointed distally. Hysterosoma with two pairs of simple setae subequal in length located in the region of metapodosoma, about one-half length of fourth pair of propodosomals; posteriorly with a single pair of simple setae subequal to anterior pair. Ventral propodosomal setae and apodemes as figured; apodeme I converging with anterior median apodeme; apodeme II not converging with anterior median nor transverse apodeme; apodeme I, the inner coxal condyles, anterior median, and apodemes II forming heart-shaped figure; striation for the heart-shaped figure as figured. Venter of hysterosoma as figured; the striae dotlike and stronger in anterior region, with some longitudinal striation, becoming smaller and dense below this region and on coxae III. Legs robust as figured. Body 223 u long by 114 uw wide. Female. Body elongate, broadest at propodosomal and hysterosomal suture. Propodosomal shield trapezoidal, bearing two pairs of simple setae; first pair one- half length of second pair; second pair about two-thirds as long as shield; pseudo- stigmatic organs ovoid, bearing small spurlike projections; pedicel as long as ex- panded distal portion. Dorsal capitulum about as long as broad, with a pair of serrated setae. Dorsum of hysterosoma with four segments; first segment with two pairs of simple setae, outer pair one-fourth longer than inner median pair; second segment with a pair of inner median setae subequal to pair on the first segment; third segment with two pairs of simple setae (a lateral and median) subequal in length to those of segments I and II; fourth segment with only a pair of lateral simple setae subequal in length and size to those of segments II and III. Ventral surface as figured. Legs I-III strong and robust; with large empodia and claws; leg IV extending to margin of the body and terminating in a whiplike seta two times the length of the leg. Body 300 uw long by 121 uw wide. The male holotype, USNM 3167, four paratype males and seven females were collected in leaf sheath of unidentified grass, Van Cort- < Steneotarsonemus friedmani, new species. Fig. 10, dorsum, male; fig. 11, dorsal, tarsus and tibia I, male; fig. 12, dorsal, tarsus and tibia II, male; fig. 13, leg IV, male; fig. 14, venter, male; fig. 15, dorsum, female; fig. 16, venter, female. PROC, ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 132 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 133 landt Park, Bronx, New York, May 5, 1965, by W. Friedman, for whom this species is named. Hemitarsonemus leonardi, n. sp. (Figs. 17-19) Three distinctive characters readily separate this species from other known members of this genus. These are: the dorsal serrate setae, the long stout claw of leg IV, and the slightly sclerotized lateral plates on the venter of the hysterosoma. Only the male is known. Male. Body slightly elongate, broadest in region of the metapodosoma. Propodo- soma trapezoidal and bearing near lateral margins four pairs of serrate setae; first and fourth pair subequal in length, second pair slightly shorter than first and fourth pair, fourth pair subequal to first pair. Hysterosoma with two pairs of equal serrate setae, stouter and subequal in length to first pair of propodosomals; and with a pair of lateral simple setae subequal in length to the two pairs of serrate setae. Opisthosoma with a pair of serrate setae equal in length to the two hysterosomal pairs. First and second pairs of ventral propodosomals equal in length. Ventral apodemes of propodosoma and hysterosoma as figured. Venter of hysterosoma with three pairs of simple setae, equal in length to the two pairs of ventral propodosomal setae; and with a slightly sclerotized plate on each side, beginning slightly posterior to the propodosomal and hysterosomal suture and ex- tending almost to coxae III; lateral plate with a simple seta two-thirds longer than the ventral propodosomal and hysterosomal setae. Legs I and II similar in length. Leg III subequal in size and length to legs I and II. Leg IV as figured; femur with small inner median lateral protuberance bearing a simple seta and with two simple setae subequal in length below protuberance; tibia long and slender, with a short seta; tarsus short, with two simple setae and one whiplike seta longer than the leg, and with long strong, stout, curved claw. Body 185 uw long by 95 u wide. Female. Not known. The male holotype, USNM 3147, was collected on Citrus sinensis (L.), Jamaica, March 9, 1961, by J. B. Bache-Wiig. The species is named for Dr. M. D. Leonard, of Washington, D. C. Hemitarsonemus deleoni, n. sp. (Figs. 20-23) The male of this species keys out to Hemitarsonemus peregrinus Beer but it differs by having dorsal serrate setae. Male. Body slightly elongate, broadest in region of metapodosoma. Propodo- soma trapezoidal, slightly sclerotized and bearing near lateral margins four pairs co Hemitarsonemus leonardi, new species. Fig. 17, dorsum, male; fig. 18, leg IV, male; fig. 19, venter, male. Hemitarsonemus deleoni, new species. Fig. 20, dorsum, male; fig. 21, dorsal, tarsus and tibia I, male; fig. 22, leg IV, male; fig. 23, venter, male. 134 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 of serrate setae; first and fourth pairs subequal in length; third pair longest, longer than propodosomal shield; fourth pair longer than first and second pairs but less than one-half length of third pair. Hysterosoma with two pairs of equal serrate setae, stouter and subequal in length to fourth pair of propodosomals; with a pair of lateral simple setae subequal in length to fourth pair of propodosomals. Opistho- soma with a pair of serrate setae equal in length to the two hysterosomal pairs. Apo- demes I shorter than apodemes II; apodemes III and IV longer than anterior apo- demes and as figured, with a lightly sclerotized lateral plate on each side, begin- ning slightly posterior to the propodosomal and hysterosomal suture and extending almost to coxae III; lateral plate with simple seta two-thirds longer than the ven- tral propodosomal and hysterosomal setae. Legs I and IL similar in length, leg II with extremely large solenidion; legs III subequal in size and length to legs I and II. Leg IV as figured; femur with small inner, lateral protuberance bearing a simple seta, and a serrate seta; tibia with a long, slender sensory rod, about half the length of the tibia, tarsus about as long as broad and with large claw. Body 185 u long by 95 uw wide. Female. Not known. The male holotype, USNM 3168, was collected on Hibiscus sp.., Tonga Islands, June 15, 1965, by L. W. Burgess. This species is named for the late Dr. Donald DeLeon, who died the summer of 1966. KEY TO THE MALES OF Hemitarsonemus EWING 1. Without a subapical spurlike process on the inner margin of femur IV _ 9) With a subapical spurlike process on the inner margin of femur IV _- PREOREN h | Seh is O Eo a ae eee) Ae tepidariorum (Warburton) 2. Dorsal jnody setae, serrated: ee 3 Dorsal), body: setae: simple, =n ee 4 3. Tibia IV with a long, dorsal, tactile, rodlike seta => deleoni, n. sp. Tibia IV with a short, dorsal, tactile, rodlike seta leonardi, n. sp. 4, First and second pair of propodosomals not subequal in length 5 First and second pair of propodosomals subequal in length cocosi DeLeon 5. Second propodosomal seta not as long as the first pair of outer hysterosomal SCtAC aio acet ng ee ee eee 6 Second propodosomal setae about as long as the first pair of outer hystero- somal Seta SoS 3. eee a oe ea ee ee lodici DeLeon 6. First outer pair of dorsal hystersomals subequal in length to the third pair of PROPOUOSOMIAIS aes eee ie ee ee boringuensis (Cromroy ) First outer pair of dorsal hysterosomals twice the length of the third pair of WTO POCLOSOMANS 2:3 60a ee ee ee ee ee ee peregrinus (Beer) Genus Tarsonemella Hirst, 1923 This genus differs from the other genera in the family by the female possessing the following: Tibiotarsus I terminates in a stout strongly recurved claw which is recessed basally and truncated distally; tarsi II and III have empodia and an outer single lateral claw each; tarsi IJ, III, and IV each has a ventral spur dis- tally; the venter of the hysterosoma has three pairs of apodemal setae and three pairs of subequal genital setae. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 135 I have studied a slide with four females and six males, which I believe to be Tarsonemella beameri Beer. Neither Hirst nor Beer described a male, and the opportunity is here taken to point out generic and specific differences based on this sex. The generic characters for the male are as follows: Tarsi Il and III have strong ventral spurs; the pretarsal elements are normal and have two claws. The venter of the propodosoma has three pairs of setae and the hysterosoma has five pairs of setae. The opisthosoma has two pairs of lateral setae on a transverse line, the posterior pair having migrated inward. Apodeme II has two pairs of simple setae. Type species. Tarsonemus (Tarsonemella) africanus Hirst. Type data. Female, collected from—*“A hymenopterous insect (Agaon xystrum Waterston ); from Koforidua, Gold Coast (April 4, 1921).” Tarsonemella beameri Beer (Figs. 24-29) Tarsonemella beameri Beer. 1958. Jour. Kan. Entomol. Soc. 31: (2) 188-192. Female. The female is as illustrated and described by Beer except the following: The hysterosoma has six segments and is lightly sclerotized with two pairs of simple setae and six pairs of lobelike setae. The pseudostigmatic organs are mace- like. The ventral hysterosomal has three pairs of subequal genital setae and two pairs of subequal apodemal setae and a smaller pair below the suture adjacent to the transverse apodeme. The empodium of tarsi II and III has only a single outer lateral claw. Figure 25 represents my drawing of the tarsal claw on legs II and III, and figure 26 represents Beer’s drawing of the tarsal claw for the same leg. Male. Body elongate and broadest in region of the metapodosoma. Dorsal propodosomal setae simple, long, and slender; third pair of propodosomals longest, one-half again longer than others; first and second pair subequal in length; third pair slightly longer than first and second pair. Hysterosoma with a pair of simple lateral setae; two pairs of finely serrated setae postero-laterally, the outer pair longest, the inner pair about one-half the length of the outer pair; a pore located above this pair of setae; below these setae a pair of simple setae about one-half the length of the inner pair of finely serrated setae. Venter of gnathosoma and hysterosoma as figured; propodosoma with three pairs of setae; first pair shortest and adjacent to apodeme I; second pair, slightly longer than first pair, medially located on coxae II; coxae II with small lateral pore. Apodeme I forming Y-shaped juncture with anterior median apodeme; apodeme II curved, well defined, and converging with anterior median apodeme; apodeme III with two pairs of simple setae medially, and one pair anterior to apodeme IV; apodeme IV with three simple setae, one adjacent to apodeme III, another above the trochanter, these two being subequal in length; a shorter and smaller seta located posteriorly on coxae IV ; posterior median apodeme well defined; coxae III and IV slightly sclero- tized. Anal plate large and well defined, lying above the subcircular genital pa- pilla. Legs I and II robust, as figured; legs III smallest, tarsi II and III with ven- tral spurs; legs IV as figured; femur with inner protuberance bearing a simple seta; below this seta a large finely serrated seta. Body 185 uw long by 121 » wide. Type data. Holotype and paratypes: Females, Saipan, Marianas Is- PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 136 Lo Vg ae Toa PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 137 lands, November 14, 1947, W. H. Lange, from the beetle, Brontispa mariana, on coconut palm, Cocos nucifera L. Specimens examined. Three paratype females in the USNM Collec- tion, and four females and six males collected by R. F. Winch in Chest- nut, Cook Islands, April 10, 1964, which I believe to be the same species. Neotarsonemus, n. gen. (Figs. 30-31) This genus is established to include two species previously assigned to Hemi- tarsonemus Ewing. These species are H. latus (Banks) and H. beeri Smiley. The two species may be separated as follows: In the males, the hysterosomal setae are long in beeri and short in latus. The females may be separated in that the trans- verse apodeme connects with the anterior median apodeme in beeri, whereas in latus these two apodemes do not connect. The generic characters for the male are as follows: Tarsi II and III are with- out claws but with well defined bell-shaped empodia; tarsi I have claws. Leg IV is composed of five segments: the tarsus, tibia, femur, coxa, and the apodeme. The apodemal segment is defined by the fusion of the posterior median apodeme, apodeme IV, and what is here considered as the fifth apodeme, all connecting to the coxal segment. When these two segments merge with the posterior median apodeme, they form an archlike structure which is not present among males of the other genera. Femur IV has a spurlike process on the inner margin. Tarsus IV terminates with a knoblike claw. The propodosoma has four pairs of setae. The characters for the females are as follows: Tarsus I has a recurved claw which is recessed basally and truncated distally. Tarsi II and III are without claws but have well defined bell-shaped empodia; there is a pair of setae between coxae IV. Type species. Hemitarsonemus beeri Smiley, 1964. Tarsonemus bucheleri, n. sp. (Figs. 32-35) This species may be recognized by the presence of an extremely large solenidion on tibia II and by a serrate seta on genu I and II ventrally. Male. Body short, oval, broadest at metapodosoma. Dorsal body setae short, slender, as figured; third pair of propodosomals longest, one-third longer than first and fourth pairs; second pair shortest, about two-thirds length of third pair; fourth pair subequal in length to first pair. First three pairs of hysterosomals of equal length, about as long as first and fourth pairs of propodosomal setae. Opisthosoma with a single pair of simple setae, subequal to second pair of propodosomals. Ven- tral apodemes as figured, first pair shorter than second pair and not forming the Y-shaped juncture at the anterior median apodeme. Transverse apodeme slightly obscure and separating at the anterior median apodeme; setation as figured; first < Tarsonemella beameri. Fig. 24, dorsum, female; fig. 25, tarsus and tibia II, fe- male; fig. 26, distal segment of tarsus II (after Beer); fig. 27, venter, female; fig. 28, venter, male; fig. 29, dorsum, male. 138 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 139 ventral apodemal setae shorter than second pair; posterior median apodeme form- ing an M-shaped juncture with apodemes III and IV; setation adjacent to the two apodemes simple and subequal in length. Legs I and II subequal in length and size; legs III smaller than legs I and II; legs IV as figured; femur with short anterior inner seta and a longer, stronger distal, daggerlike seta; genu with whip- like seta and strong solenidion; tibia—tarsus short, with short, slightly curved claw. Body 159 u long by 83 uw wide. Female. Body broadly oval, broadest at midlength. Pseudostigmatic organs spherical, bearing small spurlike projections, pedicel as long as expanded distal portion. Propodosomal shield with two pairs of setae; anterior pair serrated, two- thirds length of second pair; second pair simple, long, and slender, one-tenth shorter than length of the entire propodosoma; with a pair of median simple setae, one-third length of first pair of propodosomals and subequal to lateral pair. Hystero- soma with four segments, first segment with a pair of dorsal simple setae, and a pair of lateral simple setae; second segment larger than first, with a pair of serrated setae; third segment with lateral simple setae, and with a pair of serrated setae subequal to those of second segment; fourth ventral segment with one pair ser- rated setae; apodeme I, short, converging with anterior median apodeme; apodeme II longer than I, and not converging with anterior median apodeme; a pair of small, simple setae adjacent and below apodeme I; apodeme II without setae, transverse apodeme U-shaped, apodemes III and IV with a pair of subequal simple setae; posterior median apodeme forming a Y-shape at nonconverging extremities of apo- deme III; apodeme IV converging with posterior median apodeme. Ventral seta- tion for legs and genital plate as figured. Body 217 u long by 114 uw wide. The male holotype, USNM 3169, and a female paratype were col- lected on Ananas sativus Schult., Cook Islands, December 20, 1963, by A. Bucheler, for whom the species is named. Tarsonemus hunti, n. sp. (Figs. 36-43 ) This species may be recognized by the presence of the finely serrated hystero- somal setae and by a solenidion on tibia II which is almost as long as the segment. Male. Body short, oval, broadest at metapodosoma. Propodosomal setae simple; first pair stronger and longer than others; second pair shortest; third pair slightly longer than second pair but not as long as fourth pair; fourth pair slightly shorter than first pair. Gnathosoma about as wide as long and as figured. Hysterosoma with three pairs of finely serrated setae, subequal in length; two pairs located transversely above suture; a single pair below suture; a pair of simple lateral setae present. Ventral propodosomal and hysterosomal setae subequal in length. An- terior median apodeme forming two distinct right angles with transverse apodeme; < Neotarsonemus, new genus. Fig. 30, diagrammatic presentation of venter of fe- male N. latus; fig. 31, diagrammatic presentation of venter of female N. beeri. Tarsonemus bucheleri, new species. Fig. 32, dorsum, male; fig. 33, venter, male; fig. 34, dorsum, female; fig. 35, venter, female. 140 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 7 m™ u E >» peat —— \ See /S Ye : ne al PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 141 sclerotization light above and below this apodeme. Venter of the hysterosoma as figured and with bowl-shaped plate above and adjacent to apodemes III and IV. Legs I and II similar in length. Leg II smaller in size and length to legs I and II. Leg IV as figured and terminating with a long slender curved claw. Body 127 u long by 70 uw wide. Female. Distinctive in having short, rodlike solenidion on tarsi I and II and a ball-shaped pseudostigmatic organ with spurlike projections. Body 165 u long by 95 uw wide. The male holotype, USNM 3170, and a female paratype were col- lected on Hibiscus sp., Tonga Islands, October 13, 1964, by P. C. Hunt, for whom the species is named. Tarsonemus stricketti, n. sp. (Figs. 44-45) This species may be recognized by the presence of the finely serrated hystero- somal setae and the ventral striation pattern. Male. Body broad, oval, broadest at sejugal suture. Dorsal propodosomal setae simple, long and slender; third pair of propodosomals longest, one-half as long as the other three pair; first pair longer than second and fourth pair; second pair shortest; fourth pair about two-thirds the length of first pair. Hysterosoma with three pairs of finely serrated setae; the first two pairs subequal in length, lying transversely near the posterior margin; a shorter pair caudad. Venter of propodo- soma as figured; apodemes I short, forming a Y-shaped juncture with anterior median apodeme; apodemes II long and strong, curving inward to anterior median apodeme; transverse apodeme strong, curving inward to anterior median apodeme to form apodemal plate II; apodemal plate I with lateral pore below coxal condyle, and with a simple seta; apodemal plate II without pore, but with a simple seta; striations for each plate as figured. Venter of hysterosoma as figured; with a pair of lateral plates each with a simple seta; apodemal plates III and IV with longi- tudinal striations; each plate with a pair of simple setae; seta on plate III longer than that on IV. Legs I and II subequal in length and size and with strong solenidion on each tarsus as figured; leg III smallest; leg IV with short, stout femur bearing a simple seta proximally and long slender saberlike seta distally; tibia IV with a dorsal strong solenidion and terminating with a short, strong tarsal claw. Body 172 u long by 96 u wide. The male holotype, USNM 3173, was collected from water chestnut, Hong Kong, January 13, 1965, by J. H. Strickett, for whom the species is named. Tarsonemus moseri, n. sp. (Figs. 46-50 ) This species is characterized by the short stubby femur of leg IV and by the length of the fourth pair of propodosomal setae. < Tarsonemus hunti, new species. Fig. 36, dorsum, male; fig. 37, venter, male; fig. 38, dorsum, female; fig. 39, venter, female; fig. 40, tibia and tarsus I, male; fig. 41, leg IV, male; fig. 42, leg IV, female; fig. 43, leg I, female. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 143 Male. Body oval, broadest in region of metapodosoma. Dorsal propodosomal setae simple, long and slender; fourth pair of propodosomals longest, one-half longer than second pair, and one-third longer than first and third pair; first and third pair subequal in length. Hysterosoma with three pairs of simple setae; first two pairs subequal in length, lying transversely near the posterior margin; a shorter pair caudad. Venter of propodosoma as figured; apodemes I short, forming a Y- shaped juncture with anterior median apodeme; apodeme II long and strong, curv- ing inward to anterior median; anterior median not well defined below apodeme II, converging with transverse apodeme and forming distinct right angles and apod- emal plate II; apodemal plate II large, with lateral pore medially to transverse apodeme and coxal condyle II. Venter of hysterosoma as figured; two lateral plates each with a simple seta; apodemal plate III with two simple setae, proximal seta one-third longer than distal seta; apodemal plate IV without setae. Legs I and II subequal in length and size; tarsus II with large solenidion twice the size of solenidion on tarsus I; leg III smallest; leg IV with short stubby femur, bearing a short simple seta proximally and long saberlike seta distally; tibia IV with a strong dorsal solenidion and a long whiplike seta, and terminating with short, strong tarsal claw. Body 134 u» long by 83 u wide. The male holotype, USNM 3172, and 7 male paratypes were col- lected from inner galleries of loblolly pine with Dendroctonus frontalis Zimmerman at Elizabeth, Louisiana, January 3, 1966, by John C. Moser, U. S. Forest Service, Southern Forest Experiment Station, U. S. Department of Agriculture, Pineville, Louisiana, for whom the species is named. Tarsonemus stegmaieri, 0. sp. (Figs. 51-55) This species may be recognized by the presence of the saberlike seta on femur IV and the ventral spur on tibia IV. Male. Body oval, broadest in region of metapodosoma. Dorsal propodosomal setae simple, long, and slender; third pair of propodosomals longest, almost one- half longer than second and fourth pairs; first pair one-third shorter than third pair; second and fourth pairs subequal in length. Hysterosoma with three pairs of simple setae; first two pairs subequal in length and lying transversely near the posterior margin; a shorter pair caudad. Venter of propodosoma as figured; apo- demes I short, forming a Y-shaped juncture with anterior median apodeme; apo- deme II long and strong, curving inward to anterior median apodeme; anterior median apodeme stronger below apodemes II and converging with transverse apo- deme to form apodemal plate II; apodemal plate II wider than long, whereas apodemal plate I is longer than wide; each plate with a simple seta; the setae on plate I shorter; plate I with lateral pore. Venter of hysterosoma as figured; apex of apodemes III and IV forming M-shaped juncture with posterior median apo- — Tarsonemus stricketti, new species. Fig. 44, venter, male; fig. 45, dorsum, male. Tarsonemus moseri, new species. Fig. 46, dorsum, male; fig. 47, tarsus and tibia I, male; fig. 48, tarsus and tibia II, male; fig. 49, leg IV, male; fig. 50, venter, male. 144 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 / | 51 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 145 deme; apodemal plate III with two simple setae, proximal seta one-half longer than distal seta; apodemal plate IV without setae. Legs I and II subequal in length; legs III smallest; tarsus III with a ventral spur; leg IV with short femur, bearing a short, simple seta and long daggerlike seta distally; tibia IV with strong dorsal solenidion and a short simple seta, and terminating with long, strong tarsal claw. Body 140 u long by 70 » wide. Female. Not known. The male holotype, USNM 3171, was collected from coconut in husk, Bahama Islands, August 31, 1964, by C. E. Stegmaier, for whom the species is named. Tarsonemus newkirki, n. sp. (Figs. 56-63 ) This species may be recognized by the third pair of propodosomal setae which are longer than legs I and II and by the heart-shaped figure formed by apodemes I and II, coxal condyles I, and the anterior median apodeme. Male. Body oval, broadest at suture separating propodosoma and hysterosoma. Dorsal propodosomal setae simple, long, and slender; third pair of propodosomals longest, one-half longer than first pair; first pair strong, one-third longer than second pair; second pair shortest; fourth pair subequal to first, but not as strong. Hysterosoma with three pairs of simple setae; first two pairs subequal in length and lying transversely near posterior margin; a shorter pair caudad. Venter of propodosoma as figured; apodemes I short, curving to coxal condyles, and form- ing Y-shaped juncture with anterior median apodeme; apodemes IJ strong, con- verging with anterior median apodeme; when viewed as a whole, these structures give a heart-shaped figure; anterior median apodeme not well defined below center of apodemes II, converging with transverse apodeme; apodemal plates I and II with a pair of setae subequal in length. Venter of hysterosoma as figured; pos- terior median apodeme forming M-shaped juncture with apodemes III; apodemal plate III with two pairs of subequal simple setae; apodemal plate IV without setae. Legs I and II subequal in length and size; leg III smaller, but subequal in length to legs I and II; leg IV as figured; with long femur bearing an inner simple seta proximally, a daggerlike seta medially and a simple seta dorso-distally; tibia IV with a strong solenidion and a saberlike seta; tarsus IV terminating with a short, strong tarsal claw. Body 127 u long by 76 uw wide. Female. Distinctive in having a heart-shaped figure formed by apodemes I and II, coxal condyles I, and the anterior median apodeme. Body 127 u long by 76 u wide. The male holotype, USNM 3174, and a female paratype were col- lected on semidecayed banana, Fiji Islands, September 22, 1963, New — Tarsonemus stegmaieri, new species. Fig. 51, dorsum, male; fig. 52, tibia and tarsus I, male; fig. 53, tibia and tarsus II, male; fig. 54, leg IV, male; fig. 55, venter, male. Tarsonemus newkirki, new species. Fig. 56, dorsum, male; fig. 57, tarsus and tibia I, male; fig. 58, leg IV, male; fig. 59, venter, male; fig. 60, dorsum, female; fig. 61, tarsus I, female; fig. 62, leg IV, female; fig. 63, venter, female. 146 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Zealand Department of Agriculture, Plant Quarantine No. A320. The species is named for Richard A. Newkirk, Survey and Detection, ARS, Plant Pest Control Division, U. S. Department of Agriculture. REFERENCES Beer, R. E. 1954. A revision of the Tarsonemidae of the Western Hemisphere (Order Acarina). Univ. Kansas Sci. Bull. 36 Pt. 2, No. 16: 1091-1387. 1958. The Genus Tarsonemella Hirst, with description of a new species (Acarina, Tarsonemidae). Jour. Kan. Ent. Soc. 31 (2): 188-192. Cromroy, H. L. 1958. A preliminary survey of the plant mites of Puerto Rico. Jour. Agr., Univ. Puerto Rico 42 (2): 39-144. Ewing, H. E. 1939. A revision of the mites of the subfamily Tarsoneminae of North America, the West Indies, and the Hawaiian Islands. U. S. Dept. Agr. Tech. Bull. 653, 1-63. Smiley, R. L.: 1964. Some Tarsonemidae from the Republic of the Congo. Proc. Ent. Soc. Wash. 66 (3): 145-150. NEW HOST RECORDS FOR NORTH AMERICAN FRUIT FLIES (DierERA: TEPHRITIDAE ) J. A. Novak, W. B. Stotrzrus,’ E. J. ALLEN,” and B. A. Foote Department of Biclogical Sciences, Kent State University, Kent, Ohio 44240 Although host plants have been reported for about 70% of the ap- proximately 240 species of Tephritidae known to occur in America north of Mexico, many of the published records are not based on actual rearings or are too fragmentary to be of much value. A few workers assumed that the larvae of a particular tephritid fed on a plant merely because adults were seen on the foliage. Another source of confusion has resulted from the changing taxonomic status of certain tephritid groups. For example, some previously well-known species have been shown subsequently to be really species groups (see Bush, 1966, on Rhagoletis). It is frequently difficult, if not impossible, to determine at a later date which species of the complex was actually reared from a particular host. This paper is based on rearings conducted in southern Idaho and northwestern Montana during the summers of 1965 and 1966 and in northeastern Ohio between 1961 and 1966. It is presented now so that the records can be included in a comprehensive list of host plants of 1 Present address: Department of Biology, Eastern Mennonite College, Harrison- burg, Virginia. 2 Present address: Department of Entomology, University of Idaho, Moscow. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 147 Nearctic Tephritidae being compiled by Miss Karen Cassens and Rich- ard H. Foote, Entomology Research Division, ARS, USDA. The fruit flies were identified by R. H. Foote. The host plants were determined by W. H. Baker of the University of Idaho, by J. Thomas of Stanford University, and by A. Cusick and T. S. Cooperrider of Kent State University. The Tephritidae listed below follow the arrangement given in the recently published Diptera Catalog (Stone, et al., 1965). For each included species are given the host plants, part of host attacked, loca- tion of rearing, and any other pertinent observations. Host RECORDS Procecidochares minuta (Snow). Chrysothamnus viscidiflorus (Hook.) Nutt.; buds. Idaho. Larvae form small, monothalamous bud galls at apices of small twigs. Eutreta frontalis Curran. Aster simplex Willd.; young shoots. Ohio. Eutreta sparsa (Wiedemann). Solidago altissima L.; rhizomes. S. rugosa Mill., Aster laevis L., Helianthus annuus L., H. giganteus L., Vernonia altissima Nutt.; stems. Ohio. Stem-dwelling larvae usually form small galls. Adults reared from different host plants differ in size, color, and wing pattern and may represent host races. Males bred from S. altissima used froth masses as part of their court- ship display (Stoltzfus and Foote, 1965). Eurosta elsa Daecke. Solidago juncea Ait.; rhizomes. Ohio. Larvae form large, rather soft galls on rhizomes just below the crown. Eurosta reticulata Snow. Solidago juncea; young shoots. Ohio. Larvae form large, terminal bud galls in crown. Jamesomyia geminata (Loew). Lactuca canadensis L.; flower heads. Ohio. Xenochaeta dichromata Snow. Hieracium cynoglossoides Arv-Touv.; flower heads. Montana. Icterica circinata (Loew). Bidens cernua L.; flower heads. Ohio. Foote (In Press) presented the life cycles and described the immature stages of the 2 Nearctic species of Icterica. Icterica seriata (Loew). Bidens aristosa (Michx.) Britt., B. coronata (L.) Britt., B. frondosa L., B. tripartita L.; flower heads. Ohio. Oxyna palpalis (Coquillet). Artemisia tridentata Nutt.; small branches. Idaho. Larvae form small, succulent, polythalamous galls. Although as many as 17 variously sized larvae were found in one gall, usually only 5 to 7 mature larvae were present. Euaresta bella (Loew). Ambrosia artemisiifolia L.; flower heads. Ohio. Each larva completes development and pupates within one achene. Foote (1965) discussed briefly the life histories of this and the following species. Euaresta festiva (Loew). Ambrosia trifida L.; flower heads. Ohio. Each larva completes development within one achene. Paroxyna albiceps (Loew). Aster laevis, A. novae-angliae L.; A. pilosus Willd., A. praealtus Poir., A. puniceus L.; flower heads. Ohio. Novak and Foote (In Press) discussed the natural history and described the immature stages of this species. 148 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Dioxyna picciola (Bigot). Bidens aristosa, B. cernua, B. tripartita; flower heads. Ohio. Trupanea bisetosa (Coquillet). Arnica latifolia Bong.; flower heads. Montana. Trupanea jonesi Curran. Arnica diversifolia Greene, Aster foliaceus var. parryi (D. C. Eaton) Gray; flower heads. Idaho, Montana. Trupanea radifera (Coquillet). Balsamorhiza sagittata (Pursh) Nutt., Chrysopsis villosa (Pursh) Nutt., Senecio hydrophiloides Rydb.; flower heads. Montana. Tephritis stigmatica (Coquillet). Aster canescens Pursh, Senecio crassulas Gray, S. triangularis Hook.; flower heads. Idaho, Montana. Neotephritis finalis (Loew). Balsamorhiza sagittata, Gaillardia aristata Pursh, Helianthus annuus; flower heads. Montana. Aciurina maculata (Cole). Chrysothamnus nauseosus (Pall.) Britt.; stems. Idaho. Larvae form small galls along stem. Aciurina semilucida (Bates). Chrysothamnus nauseosus; buds. Idaho. Larvae form small bud galls along branches. Valentibulla californica (Coquillet). Chrysothamnus nauseosus; flower heads. Idaho. Valentibulla munda (Coquillet). Chrysothamnus nauseosus; stems. Idaho. Larvae form large, polythalamous galls. Xanthaciura tetraspina (Phillips). Eupatorium maculatum L.; flower heads. Ohio. Stenopa vulnerata (Loew). Senecio aureus L.; young shoots. Ohio. Larvae form small galls near apices of lateral shoots arising from crown. Strauzia longipennis var. intermedia (Loew). Helianthus giganteus; stems. Ohio. Strauzia |. var. longitudinalis (Loew). Helianthus decapetalus L.; stems. Ohio. Strauzia l. var. perfecta (Loew). Ambrosia trifida; stems. Ohio. Strauzia l. var. typica (Loew). Helianthus annuus; stems. Ohio. Strauzia l. var. vittigera (Loew). Helianthus tuberosus L.; stems. Ohio. Tomoplagia obliqua (Say). Vernonia altissima; flower heads. Ohio. REFERENCES Bush, G. L. 1966. The taxonomy, cytology, and evolution of the genus Rhago- letis in North America (Diptera, Tephritidae). Bull. Mus. Comp. Zool., Har- vard Univ. 134: 431-562. Foote, B. A. 1965. Biology and immature stages of eastern ragweed flies (Tephritidae). Proc. No. Centr. Br., Ent. Soc. Am. 20: 105-106. Biology and immature stages of fruit flies: The genus Icterica Loew (Diptera: Tephritidae). Ann. Ent. Soc. Am. (In Press). Novak, J. A. and B. A. Foote. Biology and immature stages of fruit flies: Paroxyna albiceps (Loew) (Diptera: Tephritidae). J. Kans. Ent. Soc. (In Press ). Stoltzfus, W. B. and B. A. Foote. 1965. The use of froth masses in courtship of Eutreta (Diptera: Tephritidae). Proc. Ent. Soc. Wash. 67: 263-264. Stone, A. et al. 1965. A Catalog of the Diptera of America north of Mexico. Agr. Handbook No. 276, ARS, U. S. Dept. Agr., 1696 pp. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 149 A REVIEW OF THE GENUS AULONIUM FOR THE UNITED STATES (COLEOPTERA: COLYDIIDAE ) Joun M. Kincsotver and RicHarp E. WHITE Entomology Research Division, ARS, U. S. Department of Agriculture, Washington, D. C. 20560 Species in the genus Aulonium are small, flat, shining beetles char- acterized in most species by sexually dimorphic sculpturing on the pronotal disk. Because they are cryptic, they are seldom collected and are probably more common than museum collections and faunal lists indicate. Specimens are occasionally reared or collected by forest en- tomologists in connection with biological studies of Scolytidae, but they are considered of importance only for their association with and possible predation on economic forest insects. As is true for most Colydiidae, little is known of the habits of the species of Aulonium. Kleine (1909) included Aulonium trisulcum (Geoffroy ) in his list of predators of the bark beetle, Scolytus multi- striatus (Marsham) in Europe. Craighead (1920) reported that very young larvae of Aulonium tuberculatum LeConte feed on macerated woody tissue. DeLeon (1934) listed Aulonium longum LeConte as “probably a facultative predator” associated with the mountain pine beetle, Dendroctonus ponderosae Hopkins, in ponderosa pine (Pinus ponderosa Lawson) and lodgepole pine (Pinus contorta Douglas). Until more is known about the habits of Auloniwm spp., caution should be exercised in interpretation of tree host records. Relationships may lie with insect hosts rather than plant hosts. The genus Aulonium was erected in 1845 by Erichson for four spe- cies: Colydium bidentatum Fabricius, 1801; Colydium parallelopipe- dum Say, 1827 (both from the Western Hemisphere); Ips sulcata Oliver, 1790 (now a synonym of Aulonium trisulcus (Geoffroy) 1785); and Colydium bicolor Herbst, 1797 (now a synonym of Aulonium ruficorne (Oliver), 1790). Both of the latter species are European. No type- species was designated by Erichson, and no subsequent designation of a type-species has been discovered. Two generic names are listed as synonyms of Aulonium by Hetschko (1930). Redtenbacher, in 1845, proposed the first, Anoectochilus, for Colydium bicolor Herbst, but gave no description though he listed the genus in both a key to genera of Colydiidae and a catalog of species. In 1849, Redtenbacher himself synonymized Anoectochilus with Aulonium. Pascoe, in 1860, proposed the second, Gloeania, for his new species ulomoides from Brazil. Gemminger and Harold (1868), all subsequent papers and catalogs by various authors, and most recently, Hinton (1936), have placed the species ulomoides in Aulonium. Horn (1878), in a revision of the Colydiidae for the United States, 150 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 provided a key to the four species treated in the present paper but did not include illustrations. No new species have been described from the United States since that time, and no species have been trans- ferred to the genus Aulonium. Hetschko (1930) erroneously placed Colydium lineola Say in Aulonium. Various faunal papers have included treatments of one or more spe- cies of Aulonium for the respective regions in the United States. The genus is known from North and South America, Europe, and Asia. Genus Aulonium Erichson Aulonium Erichson, 1845, p. 275. Type-species, Colydium bidentatum Fabricius (present designation ). Anoectochilus Redtenbacher, 1845, p. 124. Type-species, Colydium bicolor Herbst (monobasic ). Gloeania Pascoe, 1860, p. 99. Type-species, Gloeania ulomoides Pascoe (mono- basic). Form depressed, elongate, subparallel. Polished brownish red to black. Pro- thorax and elytra subequal in width. Head depressed dorsally; epistoma slightly penetrating anterior margin of each eye; antenna 1l-segmented with 3-segmented club, basal insertion beneath epistoma adjacent to eye. Procoxae globular with cavities closed behind; outer apical angle of protibia finely denticulate. Pronotal disk usually sexually dimorphic. Elytra elongate; striae finely, serially punctate; intervals finely alutaceous to reticulose, lateral margins subparallel, apices con- jointly rounded. Key To Species OF Aulonium OF THE UNITED STATES 1. Pronotal disk with paired distinct umbones at anterior one-third separated by, shallow. channel) \(figs22) 4,7). .2. 2 2 Pronotal disk without umbones; median channel may be present 4 2. Anterior margin of pronotum with two rounded knobs (fig. 2) — 5 CN eR g COME Sano Stee OF 5, pn Lee Ue ie ie se 4 tuberculatum Aviterior margin of pronotum without knobs 9. ee 3 3. Anterior margin of pronotum with distinct emargination at middle (fig. 7); prostemum’ very finely’ punctate —22) 770) 2) eee é longum Anterior margin nearly straight (fig. 4); prosternum strongly punctate h 1s WEP URE SU Ate Aarne ombO Eanes CREA. Seas Tes ee FeSO Rha) é parallelopipedum 4. Apex of each sublateral carina of the pronotum extending beyond anterior corner of pronotum; median lines deeply impressed (fig. 1) — SEE LRP ee oe ee See CWE eeS eae eS Ree PL NS EE ee eee ee 6, 2 ferrugineum Apex of each sublateral carina not extending beyond pronotal corners; median lines lightly impressed or nearly obliterated _______________ 5 5. Median lines of pronotal disk distinct, approaching at middle but diverging at base and apex (fig. 5); prosternum strongly, evenly punctate —- 23 SAR) Rt Se MO ae AL ta i De ek ee des Re Eb on as Sor 2 parallelopipedum Lines of pronotum nearly obliterated on apical half of pronotum, subparallel in basal half; prosternum finely punctulate, with a tendency to be trans- versely striguldse*t..gube Ui welek oe el Sey eee eee ee Se 6 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 151 6. Anterior half of pronotal disk between sublateral carinae divided by broad, shallow channel into paired convexities corresponding to prominent um- benesvinné= (ie<6) = élytranmicolorous | 22 ees 2 longum Anterior half of disk not strongly channeled, usually flat (fig. 3); apical half of elytra usually much darker than basal half — 2 tuberculatum Aulonium tuberculatum LeConte (Figs. 2 and 3) Aulonium tuberculatum LeConte, 1863, p. 67; Horn, 1878, p. 574-575; Blatchley, 1910, p. 554. Male.—Body 3.6 times as long as wide. Reddish brown with apical half of elytra usually suffused with or distinctly piceous, suffusion usually extended along suture to scutellum. Epistoma slightly emarginate apically; vertex with prominent paired tubercles. Pronotum (fig. 2) slightly longer than wide (4:3.5), margins slightly arcuate, baso-lateral angles rounded; fine submarginal sulcus extending from antero-lateral angle to basal angle merging with transverse basal sulcus; an- terior margin sinuate, broadly emarginate medially between prominent vertical knobbed_ processes; sublateral carinae prominent at anterior margin becoming obsolete at middle of pronotal disk, replaced by paired sublateral sulci on basal half; paired umbones at apical one-fourth; surface punctulate, slightly coarser punctures in paired submedian basal lines. Elytra with striae faintly indicated, intervals punctulate. Prosternum convex, faintly punctulate to impunctate. Length 4.5 mm. Female.—Color as for male. Pronotal disk (fig. 3) smoother, lacking anterior tubercles and umbones; carinae less pronounced; anterior margin with a narrow, transverse, slightly raised bead between anterior extremities of sublateral carinae. Length 4.0-4.2 mm. Geographical Range.—Alabama, District of Columbia, Florida, Georgia, Mary- land, New Jersey, New York, North Carolina, Pennsylvania, South Carolina, Texas, Virginia, West Virginia. Plant Hosts——Pinus virginiana (Miller), Pinus rigida Miller. Aulonium longum LeConte (Figs. 6 and 7) Aulonium longum LeConte, 1866, p. 378; Horn, 1878, p. 574; Hatch, 1961, p. 246. Male.—Body about 3 times as long as wide. Reddish brown with piceous suf- fusion on margins of elytra. Head slightly convex above; vertex with two tubercles; epistoma evenly rounded except for very shallow apical emargination. Pronotum longer than wide (4.0:3.5), subparallel, lateral margins slightly convergent apically; anterior margin shallowly, bisinuately emarginate between apices of sublateral carinae, deeply emarginate between sublateral carinae and antero-lateral corners; pronotal disk as in fig. 7, surface sparsely punctulate; anterior umbones rounded, sublateral carinae sharp and prominent anteriorly, obsolete on basal two-thirds. Elytra subparallel in basal three-fourths, conjointly rounded apically; intervals flat, discal striae depressed basally, faintly indicated medially, merging with punctures of intervals apically. Prosternum strongly convex, shallowly, sparsely punctate. Length 5.5 mm. 152 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Pronota of Aulonium. Fig. 1, ferrugineum Zimm., é and @; fig. 2, tubercula- tum Lec., 6; fig. 3, tuberculatum Lec., 9; fig. 4, parallelopipedum (Say), 6; > fig. 5, parallelopipedum (Say), @; fig. 6, longum Lec., 9; fig. 7, longum Lec., é. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 153 Female.—Slightly smaller than male (4.5—4.75 mm). Pronotal disk convex but not tuberculate (fig. 6). Geographical Range.—Arizona, California, Colorado, Idaho, Montana, New Mexico, Oregon, South Dakota, Utah, Washington. Plant Hosts.—Pinus edulis Engelmann, Pinus jeffreyi A. Murray, Pinus ponderosa Lawson, Pinus strobiformis Engelmann. Aulonium parallelopipedum (Say ) (Figs. 4 and 5) Colydium parallelopipedum Say, 1826, p. 263. (From LeConte, 1859 ) Aulonium parallelopipedum, LeConte, 1859, p. 324; Horn, 1878, p. 574; Blatchley, 1910, p. 554. Aulonium aequicolle LeConte, 1859a, p. 84. Synonymized by Crotch, 1873, p. 46. Male.—Body 3 times as long as wide. Reddish piceous. Head nearly flat on vertex, slightly convex on dorsum of epistoma; vertex with two indistinct convexi- ties between eyes; epistoma shallowly emarginate apically; head densely punctulate, punctures deeper on vertex than on epistoma. Pronotum as long as wide, quad- rate, margins subparallel, slightly convergent anteriorly and posteriorly, postero- lateral corners angulate, anterior margin perceptibly emarginate between sub- lateral carinae, beaded, deeply emarginate between sublateral carinae and an- terior corners; pronotal disk (fig. 4) with paired umbones low, rounded; paired median sulci extending from base to apex of pronotal disk; surface, finely, evenly punctate. Elytra subparallel on basal three-fourths, conjointly rounded apically; intervals flat, striae composed of fine, close-set punctures distinct from base to apex. Prosternum slightly convex; punctation moderately dense. Length 5.5 mm. Female.—Slightly smaller than male (4.75 mm). Pronotal disk (fig. 5) not umbonate. Geographical Range.—Arkansas, District of Columbia, Florida, Georgia, Illinois, Kentucky, Louisiana, Maryland, Mississippi, Michigan, New Jersey, New York, North Carolina, Ohio, Pennsylvania, South Carolina, Tennessee, Texas, Virginia. Plant Hosts.—Castanea dentata (Marshall) Borkhausen, Carya spp., Liriodendron tulipifera Linnaeus, Quercus spp., Taxodium distichum (L.) Richard. Aulonium ferrugineum Zimmermann (mie, 11) Aulonium ferrugineum Zimmermann, 1869, p. 254; Horn, 1878, p. 574, 575. Male.—Body 3.6 times as long as wide. Uniformly reddish brown. Head flat with small, paired umbones on vertex; epistoma slightly emarginate. Pronotum longer than wide (7:5), margins slightly convergent toward base; posterior cor- ners rounded; submarginal sulcus fine, continuous with basal sulcus, the latter slightly arcuate medially; anterior margin deeply, evenly emarginate between apices of sublateral carinae, the latter extending beyond anterior corners; marginal bead of median emargination flat; sublateral carinae extending basad about half length of pronotal disk, continuing as a sublateral sulcus to base; median paired sulci convergent apically; surface punctulate, median sulci with elongate, irregular punc- tures. Elytra with striae indistinct; surface with irregular punctulation. Proster- 154 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 num convex, punctulate, with punctures tending to be transversely strigose. Length 3.75—4.0 mm. Female.—Color as for male. Pronotal disk same as that of male. Length 3.75—- 4.0 mm. Geographical Range.—Alabama, Texas, Florida, North Carolina, South Carolina. Plant Hosts.—Pinus echinata Miller, Pinus palustris Miller, Pinus rigida Miller, Pinus taeda Linnaeus. REFERENCES Blatchley, W. S. 1910. The Coleoptera or Beetles of Indiana. Indiana Dept. Geol. Nat. Res. Bull. 1. 1386 pp. Craighead, F. C. 1920. Biology of some Coleoptera of the Families Colydiidae and Bothrideridae. Proc. Ent. Soc. Wash. 22(1): 1-13. Crotch, G. R. 1873. Check List of the Coleoptera of America, North of Mexico. Salem, Mass. 136 pp. DeLeon, D. 1934. An Annotated List of the Parasites, Predators and Other Associated Fauna of the Mountain Pine Beetle in Western White Pine and Loblolly Pine. Can. Ent. 66: 51-61. Erichson, W. F. 1845. Naturgeschichte der Insecten Deutschlands. Abt. 1, Coleoptera, Bd. 3, Lief. 2: 161-320. Gemminger, M. and E. von Harold. 1868. Catalogus Coleopterorum Hucusque Descriptorum Synonymicus et Systematicus. Vol. 3: 753-978. Hatch, M. H. 1961. The Beetles of the Pacific Northwest. Part 3. Univ. of Wash. Press, Seattle. 503 pp. Hetschko, A. 1930. Coleopterorum Catalogus, Junk. Pars 107: 1-124. Hinton, H. E. 1936. Miscellaneous Studies in the Neotropical Colydiidae (Col. ) Rev. de Ent. 6(1): 47-97. Horn, G. H. 1878. Synopsis of the Colydiidae of the United States. Proc. Amer. Philos. Soc. 17: 555-592. Kleine, R. 1909. Die Europaischen Borkenkifer und ihre Feinde aus den Ordungen der Coleopteren and Hymenopteren. Ent. Blatt. 5(3): 41-50. LeConte, J. L. 1859. The Complete Writings of Thomas Say on the Entomology of North America. Vol. 2. Boston. 1859a. Catalogue of the Coleoptera of Fort Tejon, California. Acad. Nat. Sci. Phila. Proc. 1859: 69-90. 1863. New Species of North American Coleoptera. Smiths. Misc. Coll. 167. 86 pp. 1866. Additions to the Coleopterous Fauna of the United States, No. 1. Acad. Nat. Sci. Phila. 1866: 361-394. Pascoe, F. P. 1860. Notices of New or Little Known Genera and Species of Coleoptera. Journ. Ent. 1(2): 98-132. Redtenbacher, L. 1845. Die Gattung der deutschen Kaefer-Fauna nach der analytischen Methode bearbeitet nebst einem kurz gefassten Leitfaden zum Studium dieses Zweiges der Entomologie. Wien. 177 pp. 1849. Fauna Austriaca. Die Kafer, nach der analytischen Methode bearbeitet. Wien. 883 pp. Say, T. 1826. Descriptions of New Species of Coleopterous Insects Inhabiting the United States. Journ. Acad. Nat. Sci. Phila. 5(2): 237-284. Zimmermann, C. C. A. 1869. Synonymical Notes on Coleoptera of the United States, with Descriptions of New Species. Trans. Amer. Ent. Soc. 2: 243-259. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 ilts{s) TINCONARIA ALBESCENTIS, A NEW NORTH AMERICAN GENUS AND SPECIES OF CECIDOMYIIDAE FOUND IN SEEDS OF TRIODIA ALBESCENS VASEY ( Diptera ) RAYMOND J. GAGNE Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560 The genus described below belongs to the tribe Contariniini of the dipterous family Cecidomyiidae. It is close to Contarinia and Stenodiplosis on the basis of male and female genitalic characters, but it differs from them and all the other genera of the tribe by the following combination of characters: The 2-segmented palpus, the greatly enlarged tarsal claw in relation to the last tarsomere, and the equal length of the tibiae and tarsi. The wing is slightly narrower than other grass-infesting midges belonging to Contarinia and Stenodi- plosis. Tinconaria, n. gen. Antenna with 12 flagellomeres, the two nodes of each flagellomere subequal, each girdled by one circumfilum and a whorl of setae. Palpus 2-segmented, the segments short, seldom longer than twice the diameter. Wing narrow; Cu: and Cus evanescent on the distal half; area behind cubitus narrowed. Tarsi short, approximately equal to the tibiae in length; tarsal claws (fig. 2) simple, ap- proximately two-thirds the length of the last tarsomere. Male postabdomen (fig. 1) with the tenth tergum and sternum both bilobed, the lobes rounded; aedeagus not greatly produced, tapering distally; basimere stout; distimere tapering gradually beyond the middle, toothed at apex. Female ovipositor (8th abdominal segment to apex, inclusive) approximately twice as long as remainder of abdomen, longitudinally striated on distal half except for the sclerotized cerci (figs. 3, 4) which are elongate-attenuate and bear several setae. Type-species: Tinconaria albescentis n. sp. This genus is established for the species described below, which was reared from seed heads of Triodia albescens Vasey (Gramineae). Information about the antennae is from a specimen not included in the following description but discussed below. Tinconaria albescentis, n. sp. Antennae missing. Palpus: 2-segmented; first segment short and_ blunt, 0.020—0.030 mm. in length, 0.015—0.025 mm. in dia.; second segment short, tapering at apex, 0.028—0.033 mm. in length, 0.010—0.015 mm. in dia. Chaetotaxy: frontoclypeal setae, 9-15 (avg. of 8 observations, 12); dorsocentral, 9-14 (12); scutal, 11-16 (13); scutellar, 2-4 (3); subalar, 3-4 (3). Length of foretarsomeres (in mm.): I, 0.065-0.075 (avg. of 6 observations, 0.068); II, 0.200-0.225 (0.210); III, 0.190-0.200 (0.195); IV, 0.060-0.075 (0.065); V, 0.060-—0.070 (0.065). Per cent length of each tarsomere to total length of tarsus, from I to V; 13, 42, 19, 13, 13. Tarsi subequal in length to tibiae. Tarsal claw (fig. 2) 156 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Fig. 1. Male postabdomen, 400%. Fig. 2. Tarsomere V of foreleg, 300x. Fig. 3. Cerci of ovipositor, dorsal view, 450. Fig. 4. Same, lateral view, 450x. very long, simple, more than two-thirds the length of tarsomere V, 0.045—0.050 mm. in length. Empodium shorter than claws. Wing length 1.36-1.49 mm. (avg. of 8 observations, 1.43 mm.), width 0.47-0.53 mm. (0.50 mm.); ratio width to length (based on avgs.) 0.345. Cu: and Cuz evanescent on distal half; posterior area narrowed. Male postabdomen (fig. 1): tenth tergum and sternum bilobed, the sternum longer; aedeagus acute, as long as tenth sternum; basimere short, stout; distimere tapering gradually beyond middle, toothed at apex. Female postabdomen: cerci (figs. 3.4) long, thin, closely juxtaposed laterally, tapering at end, 0.065-0.075 mm. in length and with several setae scattered along the surface. Immatures stages unknown. Material examined. Holotype (on slide): ?; ex seeds of Triodia albescens; intercepted by U.S. Department of Agriculture, Plant Quarantine Division; Brownsville, Texas; V-16-1966; 66-14344; U.S. National Museum type no. 69358. Paratypes: 1¢, 622, with same data as holotype. Types all deposited in U.S. National Museum. Midges similar to T. albescentis have been previously reported in the literature but have been misidentified as Contarinia sorghicola (Coquillett), the sorghum midge. G. G. Ainslee (unpub.) reared several specimens from the heads of Triodia flava (L.) Smyth and identified them as C. sorghicola. Dean (1911), Walter (1941), and Barnes (1956) referred to those specimens under that name. In the U.S. National Museum there is one specimen of the midges reared by Ainslee. The label bears the following data: “Diplosis sorghicola; Web. No. 1874; Clemson Col., $.C.; 2 Oct. 08; bred from heads of PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 157 Triodia sesleroides; Geo. G. Ainslee.” T. sesleroides is a synonym of T. flava. Though it is obvious the specimen belongs to Tinconaria because of the 2-segmented palpus and the large tarsal claw, it is in very poor condition on a slide. Therefore I cannot be sure whether this specimen belongs to the species described above or is a different species. This Ainslee record does show, however, that Tinconaria is found on another species of Triodia. REFERENCES Barnes, H. F. 1956. Gall midges of economic importance. Vol. VII. Cereal Crops. Crosby Lockwood and Son Ltd., London. 261 pp. Dean, W. H. 1911. The sorghum midge. U.S. Dept. Agr. Bur. of Ent. Bull. 85: 39-58, 2 pls. Walter, E. V. 1941. The biology and control of the sorghum midge. U.S. Dept. Agr. Tech. Bull. 778: 1-26. THE METANOTAL GLAND AS A TAXONOMIC CHARACTER IN OECANTHUS OF THE UNITED STATES’ (OrTHOPTERA: GRYLLIDAE) T. J. WALKER? AND AsHLEY B. GuRNEY® Male crickets of the subfamily Oecanthinae have a large glandular cavity in the metanotum. This metanotal gland is sometimes called Hancock's gland because Hancock (1905) was the first to describe it adequately. The metanotal gland is mouthed (ie., “licked” or bitten) by the female before and after she receives the spermatophore from the male. Its secretions probably are important both to the suc- cessful transfer of the spermatophore and to the diversion of the female while the sperm pass from the spermatophore into the sperm receptacle. During this time, the male has the tegmina raised, ex- posing the gland, and the female is astride the male. The female removes and eats the spermatophore soon after leaving the male. A third function of the gland may be to keep the pair together until another spermatophore can be passed. Postcopulatory feeding at the gland may last for over an hour (65 minutes observed in Oecanthus pini by TJW), and a second spermatophore may be transferred within 70 minutes of the first (observed in O. argentinus by TJW). A fourth function may be that the secretions are specifically distinct and account 1 Study supported by NSF Grant GB4949. 2 Department of Entomology, University of Florida, Gainesville. ° Entomology Research Division, ARS, USDA, Washington, D.C. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 1.0 mm = PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 159 for the usual failure of females to respond to the courtship of hetero- specific males (Walker 1963). The morphology and histology of the oecanthine metanotal gland are described by Engelhardt (1914) for Oecanthus pellucens (Scopoli ) and by Fulton (1915) for an unspecified species of Oecanthus (prob- ably nigricornis or quadripunctatus ). Males of some species in other gryllid subfamilies have metanotal glands—Gryllinae, Discoptila fragosoi (Bolivar) (Boldyrey 1928); Phalangopsinae, Homoeogryllus (LeRoy 1964); Eneopterinae, Tafalisca, Hapithus (TJW). However, only in Oecanthinae are the glands large and complex and present in all species examined. The chief parts of the metanotal gland in Oecanthinae are a metascutum (Fig. 4, ms) and metascutellum (msl), henceforth referred to here as “scutum” and “scutellum.” A well-defined and distinctive posterior median lobe of the scutum (Fig. 4, pml) occurs in some species but not in others; it is henceforth termed “posterior median scutal lobe.” L. Chopard and A. B. Gurney were first to examine oecanthine metanotal glands for their taxonomic value: Chopard (1955, Fig. 30-32) pointed out that three similar species of African Oecanthus differed in features of the gland. Gurney, in 1951, examined many New World species of Oecanthus and drawings were made (including Fig. 1-6 of this paper). Because of uncertainties in the taxonomy and nomenclature of U. S. Oecanthus he did not publish his results. These uncertainties were subsequently dealt with by Walker (1962, 1963). As a result of Gurney’s work with Oecanthus, Walker (1967, Fig. 20-27) studied the metanotal glands of the oecanthine genus Neox- abea, discovered features of taxonomic value, and illustrated the glands of eight species. The present paper summarizes the early studies by Gurney and the later studies by Walker of the metanotal glands of U. S. Oecanthus. Methods. Five or more specimens of each U. S. species (except laricis) and closely related Latin American species were examined. Specimens from diverse localities were selected to maximize intra- specific variation. Pinned specimens were relaxed, and their metanotal — Fig. 1-9. Male metanotum of Oecanthus. Fig. 1-5, Dorsal view of scutum and scutellum (scale beneath Fig. 1). Fig. 1. niveus, specimen from Geneva, N. Y.; fig. 2, exclamationis, Johnson City, Tenn.; fig. 3, nigricornis, Glen Echo, Md.; fig. 4, californicus, Huachuca Mts., Ariz.; fig. 5, allardi, Ciudad Trujillo [Santo Domingo], Dominican Republic; fig. 6, Posterior median scutal lobe of rileyi (Locality of figured specimen uncertain, probably Corvallis, Oreg.); fig. 7-9, Caudal view of scutellum (small portions of scutum showing, scale as in fig. 1); fig. 7, exclamationis, Gainesville, Fla.; fig. 8, leptogrammus, Brownsville, Tex., fig. 9, niveus, Highlands Co., Fla. ms—metascutum (scutum), ms/—metascutellum (scutellum ), pml—posterior median lobe of scutum, set—setae on scutum at scutoscutellar suture, setl—setae on scutellum, st—scutal tubercle, w—width of posterior median scutal lobe. 160 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 glands were exposed by elevating and pulling forward the tegmina. They were then dried in this position. In alcohol-preserved specimens, the glands were exposed by amputating the tegmina. Results. Among U. S. species of Oecanthus the scutellum is not elaborately modified as in most species of Neoxabea (Walker 1967), nor is there, except in O. niveus, a well-developed median scutal tubercle (Fig. 1, st) such as Chopard (1955) found useful in identify- ing African Oecanthus. However, as the following key and accom- panying figures illustrate, features of the metanotal glands may be used to separate the species into the same species groups that have been recognized on the basis of other features such as antennal mark- ings, calling song, and stridulatory file (Walker 1962). Features of the metanotal gland, especially the lack of brushes or bundles of long setae on the scutum at the scutoscutellar suture or on the scutel- lum, indicate that species related to rileyi are not closely related to niveus. Therefore, the rileyi group of species is here recognized as distinct from the niveus group. Although Walker (1962) considered these to be a single group, features of the antennal markings, calling song, and stridulatory file support the present division. In general, species within a species group are difficult or impossible to separate on the basis of the metanotal gland, and other character- istics are adequate for identification. However, as indicated in coup- lets 4 and 5 of the key, within the niveus group the metanotal gland easily separates species that in a few instances are difficult to sep- arate by other means. Indeed, the differences in the glands within the niveus group are as great as the differences used to separate the other species into species groups. Oecanthus fultoni shows significant geographical variation in the structure of the metanotal gland. West Coast specimens have a pos- terior median scutal lobe similar to that of O. rileyi, and eastern speci- mens resemble the West Indian O. allardi in this respect. Key to U.S. Oeecanthus Species, BASED ON MALE METANOTAL GLANDS Il Brushes or bundles of long setae on scutum at scutoscutellar suture (Fig. 2, set) or on scutellum (Fig. 1, sett) —___ niveus group —--__--- 4 1 No setal brushes or bundles on scutum at scutoscutellar suture nor on Scutellum (Fig: (O)14, 5)! 2 2 ee — eee a 9. (1’), Scutum with posterior median lobe (Fig. 45.5) —.—.-. _. — =e 3 2! Scutum without posterior median lobe (Fig. 3) ----- nigricornis group, including argentinus Saussure, celerinictus T. Walker, laricis T. Walker, nigricornis F. Walker, pini Beutenmuller, and quadripunctatus Beuten- muller 3 (2) Width of posterior median scutal lobe (Fig. 4, w) twice length or more; anterior edge of lobe describing a shallow arc (Fig. 4) varicornis PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 161 group, including ecalifornicus Saussure, latipennis Riley, major T. Walker, and varicornis F. Walker By Width of posterior median scutal lobe (Fig. 5, w) twice length or less; anterior edge of lobe describing a semicircle (Fig. 5) or a circumflex (ES) YA CLITA 5H SS oA PRI RRS RN ee ee ee rileyi group, in- cluding allardi T. Walker & Gumey, fultoni T. Walker, and rileyi Baker 4(1) Pair of vertical, bent-tip, setal bundles on scutum at scutoscutellar STIULIREL (Ieee), eat ee ook eee eT Se Ee ete 3 exclamationis Davis 4’ No} setaliibundles ‘atiscutoscutellar ‘suture: 222 © ee es 5 5 (4’) Scutellum with a pair of forward-directed horn-like processes; pair of setal brushes on scutum at scutellar processes (Fig. 8) Ce as ee ee ee OS ee eee a See ae leptogrammus T. Walker 5! Scutellum without horn-like processes; pair of setal brushes on scutellum (TBE 0 alee». anil ine Bis r= 6a MULE. agri ID, «cites a Nees niveus (De Geer) REFERENCES Boldyrey, B. Th. 1928. Einige Episoden aus dem Geschlechtsleben von Dis- coptila fragosoi Bol. (Orthoptera, Gryllidae). Rev. Russe Ent. 22: 137-147. Chopard, Lucien. 1955. Orthoptera Ensifera et Tridactyloidea. South Afr. Animal Life, Results Lund Univ. Exped. 1950-1951. 2: 266-301. Engelhardt, V. V. 1914. Uber die Hancocksche Driise von Oecanthus pellucens Scop. Zool. Anz. 44: 219-227. Fulton, B. B. 1915. The tree crickets of New York: life history and bionomics. N. Y. Agr. Exp. Sta. Tech. Bull. 42. 47 p. Hancock, J. L. 1905. The habits of the striped meadow cricket (Oecanthus fasciatus Fitch). Amer. Nat. 39: 1-11. LeRoy, Yveline. 1964. Les caracteres sexuels et le comportement acoustique des males d’Homoeogryllus reticulatus Fabricius. Bull. Soc. Ent. France 69: 7-14. Walker, T. J. 1962. The taxonomy and calling songs of United States tree crickets (Orthoptera: Gryllidae: Oecanthinae). I. The genus Neoxabea and the niveus and varicornis groups of the genus Oecanthus. Ann. Ent. Soc. Amer. 55(3): 303-322. 1963. The taxonomy and calling songs of United States tree crickets (Orthoptera: Gryllidae: Oecanthinae). II. The nigricornis group of the genus Oecanthus. Ann. Ent. Soc. Amer. 56(6): 772-789. 1967. Revision of the Oecanthinae (Gryllidae: Orthoptera) of America south of the United States. Ann. Ent. Soc. Amer. 60: 784-796. 162 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 STUDIES OF NEOTROPICAL CADDIS FLIES, VI: ON A COLLECTION FROM NORTHWESTERN MEXICO OLIVER S. FLINT, JR. Smithsonian Institution, Washington, D.C. 20560 One of the more interesting and well-known zoogeographic facts is the intermingling of the Nearctic and Neotropical faunas in Mexico. It has been known for a long time that Nearctic elements enter Mexico via the central highlands, but become greatly reduced south of the trans-Mexican volcanic belt just south of Mexico, D. F., al- though a few elements keep reappearing on the high mountains as far south as Costa Rica. Conversely, Neotropical elements enter Mexico from the south and extend northward to greater or lesser degrees in the lowland regions of the Pacific and Gulf coasts. Complicating this pattern are the effects of rainfall, elevation, and the ecological re- quirements of the group involved. The Trichoptera show this intermingling of faunas in Mexico in a clear manner. The northern elements enter Mexico along the Sierra Madre Occidental, but the large desert area generally bars the caddis flies from the high region to the east of the mountains. In the warmer streams of the coastal lowlands just to the west of the mountains are found many species, probably of Neotropical origin, which are widely distributed in Central America and which are also found in many of the warmer streams in the western United States. Confusing this picture are certain genera, apparently of Neotropical origin (Ato- psyche and Marilia, at least) which have become cool-adapted, and extend northward along the same mountains which are inhabited by the Nearctic groups. The collection upon which this report is based, made by a group from the Canadian Department of Agriculture, is the most important collection made in the “Nearctic” section of northwestern Mexico. The material is mainly from the high mountains of Durango, and serves to link the relatively well-known areas in Arizona with the region near Mexico City. It is not surprising therefore that there are many records extending the known ranges of Arizonan species far to the south and a smaller number extending ranges from near Mexico City to the northwest. All specimens which are listed from the base camp without more specific locality data are from a site located at 9000 feet elevation, 10 miles west of El Salto, Durango, Mexico. They were collected pri- marily by J. E. H. Martin, with smaller numbers taken by J. F. Mc- Alpine, W. C. McGuffin, and H. F. Howden. A brief discussion of PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 163 the site, its weather and ecology may be found in Thomas (1966, Can. Ent. 98: 871-875). Holotypes and allotypes are deposited in the Canadian National Collection. Paratypes and material of the described species are di- vided between the Canadian National Collection and the U.S. National Museum. I wish to thank Dr. Fernand Schmid for making the collec- tion available for my study. In addition to the material specifically reported on here, there are females in the genera Atopsyche, Chimarra, Polycentropus, Cheu- matopsyche, Hydropsyche, and Lepidostoma which are not determined to species. It is not presently possible to be certain of the correct association of males and females for the majority of the species in these genera. RHYACOPHILIDAE Atopsyche boneti Ross and King Atopsyche boneti Ross and King, 1952, Ann. Ent. Soc. Amer. 45: 194. The species, described from Morelos and which I have collected near Cordoba, Ver., is here recorded from northwestern Mexico for the first time. Base Camp: 6 July 1964, 1é. Atopsyche calopta Ross and King Atopsyche calopta Ross and King, 1952, Ann. Ent. Soc. Amer. 45: 188. A single male of this species was collected at the Base Camp on 13 June 1964. The original material came from the states of Morelos, Oaxaca, and Chiapas. Atopsyche sperryi Denning Atopsyche sperryi Denning, 1949, J. Kans. Ent. Soc. 22: 88. Ross and King, 1952, Ann. Ent. Soc. Amer. 45: 190. This species was previously known only from Arizona. Base Camp: 20 April 1964, 16; 8 June 1964, 26; 9 June 1964, 3¢; 11 June 1964, 1¢; 13 June 1964, 24; 17 June 1964, 446; 18 June 1964, 16; 20 June 1964, 16; 25 June 1964, 24; 27 June 1964, 14; 1 July 1964, 24; 15 July 1964, 34; 29 July 1964, 34; 31 July 1964, 16; 5 Aug. 1964, 16; 8 Aug. 1964, 1¢. Durango, 5 miles west El Salto, 7 June 1964, 14. Atopsyche tripunctata Banks Atopsyche tripunctata Banks, 1905, Trans. Amer. Ent. Soc. 32: 17. Ross, 1947, Trans. Amer. Ent. Soc. 73: 128. Ross and King, 1952, Ann. Ent. Soc. Amer. 45: 190. Similarly to the preceding species, this species was known only from Arizona. Base Camp: 8 June 1964, 24; 16 June 1964, 14; 18 June 1964, 14; 26 June 1964, 1g; 28 June 1964, 34; 6 July 1964, 14; 7 July 1964, 14; 8 July 1964, 4é; 29 July 1964, 1¢; 31 July 1964, 16; 1 Aug. 1964, 246; 2 Aug. 1964, 16; 8 Aug. 1964, 12. 164 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 yi A Pee / 7 J fh adh Figs. 1-16. Atopsyche socialis, n. sp.: 1, male genitalia, lateral; 2, aedeagus, lateral. Chimarra bicoloroides, n. sp.: 3, male genitalia, lateral; 4, clasper, dorsal. Polycentropus palmitus, n. sp.: 5, male genitalia, lateral; 6, aedeagus, lateral. Limnephilus pollux, n. sp.: 7, male genitalia, dorsal; 8, aedeagus, lateral; 9, male genitalia, lateral; 10, female genitalia, lateral; 11, female genitalia, dorsal. Limnephilus ctenifer, n. sp.: 12, male genitalia, lateral; 13, aedeagus, lateral; 14, female genitalia, dorsal; 15, female genitalia, lateral; 16, male genitalia, pos- terior. PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 165 Atopsyche socialis, n. sp. (Figs. 1-2) This species is closely related to A. tripunctata Banks, from which it differs in the shape of the paracercus and in the structure of the aedeagus which lacks the bifurcate dorsal process but has a divided tip. Adult. Length of forewing, 10-11 mm. Color grayish brown; forewings mottled brown, paler in anal region. Sixth sternum with apicomesal process about a third of length of segment; seventh sternum with apicomesal process about a fifth of length of segment. Male genitalia: Filicercus elongate, clavate, shorter than paracercus. Paracercus slender, with a dorsal subapical tooth and dorsal and lateral apical teeth. Basal clasper segment long, inner surface with a shelf ending in a short apical process; apical segment trianguloid, dorsoapical margin inrolled. Aedeagus with a strong lateral spine, apex split into dorsal and ventral points, a simple internal rod present. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’, 11 Aug. 1964, J. E. H. Martin. Paratypes. Same, but 8 June 1964, 16; 17 June 1964, 3¢; 25 June 1964, 14; 8 July 1964, 1¢; 10 July 1964, 2¢. GLOSSOSOMATIDAE Glossosoma ventrale Banks Glossosoma ventrale Banks, 1904, Trans. Amer. Ent. Soc. 30: 109. Ross, 1956, Evol. & Classif. Mtn. Caddisfl., p. 153, fig. 301. Fischer, 1960, Trich. Catal. 24: There is a single female collected at the Base Camp on 6 Aug. 1964 ap- parently belonging to this species. This is the first record of the species, genus, and subfamily (Glossosomatinae ) in Mexico. PHILOPOTAMIDAE Wormaldia arizonensis (Ling ) Dolophilus arizonensis Ling, 1938, Pan-Pac. Ent. 14: 63. Ross, 1941, Trans. Amer. Ent. Soc. 67: 51. Fischer, 1961, Trich. Catal. 2: 46. Wormaldia arizonensis (Ling): Ross, 1949, Proc. Ent. Soc. Wash. 51: 155. Ross, 1956, Evol. & Classif. Mtn. Caddisfl., p. 62, fig. 74. These are the first records of this species in Mexico. Base Camp: 8 June 1964, 16 19; 13 June 1964, 19; 17 June 1964, 16 19; 1 July 1964, 1¢; 6 July 1964, 1¢; 10 July 1964, 19; 12 July 1964, 19. Durango: 5 miles west of El Salto, 7 June 1964, 19; 3 miles west of El Salto, 19 June 1964, 1¢. Chimarra bicolor ( Banks ) Philopotamus bicolor Banks, 1901, Trans. Amer. Ent. Soc. 27: 370. Fischer, 1961, Trich. Catal. 2: 6: Chimarra bicolor (Banks): Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 3. One female of this Mexican species from Sinaloa, 15 miles west of El Palmito, 5000’, 20 July 1964. 166 PROC, ENT, SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Chimarra bicoloroides, n. sp. (Figs. 3-4) In coloration this species is similar to C. bicolor (Banks), however, only the head and pronotum are orange whereas the whole body is orange in bicolor. The structure of the male genitalia places this species in a very isolated position. The presence and shape of the dorsal and lateral lobes of the tenth tergum together with the extremely long spines in the aedeagus render the genitalia very distinctive. Adult. Length of forewing, 9 mm. Head, pronotum, and forecoxae orange; rest of body, wings and appendages fuscous. Male genitalia: Ninth segment with a midventral keel. Clasper with a narrow dorsal prolongation ending in a sharp point, mesal surface with a distinctive central ridge dividing the thick basal from the thin apical portion. Tenth tergum divided into broad, rectangular dorsal plates, and more elongate lateral plates. Aedeagus (completely everted in type) with an extremely long internal spine overlying a long trough-like ventral spine; membranous apex with dorsal and ventral spiculate patches. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’, 28 July 1964, J. E. H. Martin. Chimarra embia Ross Chimarra embia Ross, 1959, Ent. News 70: 170. There is a single male of this common Mexican species. Sinaloa, 21 miles east of Villa Union, 25 July 1964. Chimarra mexicana (Banks ) Rhyacophila mexicana Banks, 1900, Trans. Amer. Ent. Soc. 26; 259. Chimarra mexicana (Banks): Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 3. This distinctively marked species is recorded from the northwestern part of Mexico for the first time. Base Camp: 11 June 1964, 1¢; 17 June 1964, 74 19; 25 June 1964, 2¢. Chimarra utahensis ( Ross ) Chimarrha utahensis Ross, 1938, Bull. Il. Nat. Hist. Surv. 21: 134. Chimarra utahensis (Ross): Ross, 1951, Proc. Cal. Acad. Sci. (4th Ser.) 27: 67. This species, known from the southwestern United States and Baja California, is recorded from Mexico proper for the first time. Durango, 7 miles west of Durango, 7000’, 26 July 1964, 1¢. PsYCHOMYIIDAE Polycentropus arizonensis Banks Polycentropus arizonensis Banks, 1905, Trans. Amer. Ent. Soc. 32: 85. Ross, 1938, Psyche 45: 13. Denning, 1956, Pan-Pac. Ent. 32: 80. Fischer, 1962, Trich. Catal. 3: 66. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 167 Known previously only from New Mexico and Arizona, this species is recorded from Mexico for the first time. Base Camp: 8 June 1964, 1¢; 13 June 1964, 14; 16 June 1964, 14; 20 June 1964, 1¢; 2 July 1964, 19; 10 July 1964, 19; 24 July 1964, 19; 31 July 1964, 1°. Polycentropus aztecus Flint Polycentropus aztecus Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3608, p. 9. The following are paratypes of this recently described species. Base Camp: 25 June 1964, 146; 15 July 1964, 14; 28 July 1964, 16; 29 July 1964, 26; 6 Aug. 1964, 1¢. Polycentropus casicus Denning Polycentropus casicus Denning, in Denning & Sykora, 1966, Can. Ent. 98: 1222. There is a topotypic pair of this common, widespread, Mexican species. Du- rango, 24 miles west of La Ciudad, 15 June 1964, 1¢ 19. Polycentropus halidus Milne Polycentropus halidus Milne, 1936, Studies N. Amer. Trich. 3: 86. Denning, 1948, Psyche 55: 23. Denning & Sykora, 1966, Can. Ent. 98: 1220. This species is known otherwise from New Mexico, Arizona, California and Mexico. Base Camp: 13 June 1964, 1¢; 17 June 1964, 2¢; 20 June 1964, 14; 25 June 1964, 24; 28 June 1964, 14; 5 July 1964, 1¢. Durango, 2 miles west of El Salto, 19 June 1964, 124. Polycentropus palmitus, n. sp. (Figs. 5-6) This species is close to P. halidus Milne, from which it differs in lacking the hooklike appendage from the inner lobe of the cercus, and in having longer, narrower claspers. Adult. Length of forewing, 6.5 mm. Color brown; body and appendages slightly paler, forewing spotted with patches of golden hair. Male genitalia: Cercus divided into 3 parts: a broad dorsolateral lobe, an inner slightly more elongate ventral lobe, and a dorsal saber-like process. Clasper long and slender, tip slightly upturned, inner face with a slight ridge, culminating in a_ basal tooth. Aedeagus broad, with an apicoventral process, trianguloid in posterior aspect, and dorsolateral plates. Holotype, male. Mexico (Sinaloa), 8 miles west of El Palmito, 19 July 1964, H. F. Howden. Polycentropus picana Ross Polycentropus picana Ross, 1947, Trans. Amer. Ent. Soc. 73: 136. Denning, 1966, Pan-Pac. Ent. 42: 232. This species, which is common and widely distributed in northeastern and central Mexico, is recorded from northwestern Mexico for the first time. Durango, 7 miles west of Durango, 7000’, 26 July 1964, 14 19. 168 PROC, ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 HypROPSYCHIDAE Smicridea fasciatella McLachlan Smicridea fasciatella McLachlan, 1871, J. Linn. Soc. Lond., Zool. 11: 136. Ross, 1944, Bull. Ill. Nat. Hist. Surv. 23: 85. Fischer, 1963, Trich. Catal. 4: 132. This common species is widespread in southwestern United States and northern Mexico. Sinaloa: 28 miles east Villa Union, 500’, 4 Aug. 1964, 29; 21 miles east Villa Union, 25 July 1964, 34 39. Smicridea utico Ross Smicridea utico Ross, 1947, Trans. Amer. Ent. Soc. 73: 144. This is another common species in Central America, Mexico, and the south- western United States. Sinaloa: 21 miles east of Villa Union, 25 July 1964, 12; 28 miles east of Villa Union, 500’, 4 Aug. 1964, 19. Cheumatopsyche gelita Denning Cheumatopsyche gelita Denning, 1952, Can. Ent. 84: 20. Known previously only from the type series collected in Arizona, the species is here recorded from Mexico. Base Camp: 25 June 1964, 1¢2. Cheumatopsyche zion Ross Cheumatopsyche zion Ross, 1947, Trans. Amer. Ent. Soc. 73: 141. The species is widely distributed over the southwestern United States, and is here recorded from Mexico for the first time. Base Camp: 4 July 1964, 19; 29 July 1964, 19; 6 Aug. 1964, 14. Hydropsyche solex Ross Hydropsyche solex Ross, 1944, Bull. Ill. Nat. Hist. Surv. 23: 271. This species seems to have a widespread distribution throughout the more arid areas of western United States, but has not been recorded from Mexico previously. Durango: 5 miles west of Durango, 6500’, 29 June 1964, 16 292; 7 miles west of Durango, 7000’, 26 July 1964, 49°. Hydropsyche oslari Banks Hydropsyche oslari Banks, 1905, Trans. Amer. Ent. Soc. 32: 13. Ross, 1938, Psyche 45: 18. Fischer, 1963, Trich. Catal. 4: 65. Denning, 1964, Pan-Pac. Bint. 403) 133% This common and widespread western North American species has been known in Mexico previously only from Baja California. Base Camp: 6 June 1964, 14; 8 June 1964, 34; 17 June 1964, 24; 25 June 1964, 24; 1 July 1964, 124; 2 July 1964, 2%, 29 July 1964, 1¢. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 169 Hydropsyche occidentalis Banks Hydropsyche occidentalis Banks, 1900, Trans. Amer. Ent. Soc. 26: 258. Ross, 1938, Psyche 45: 17. Fischer, 1963, Trich. Catal. 4: 58. Denning, 1964, Pan-Pac. Ent. 40: 133. This species has a range like that of the preceding. Base Camp: 5 June 1964, 13; 9 June 1964, 26; 2 July 1964, 14; 4 July 1964, 26; 17 July 1964, 14. LIMNEPHILIDAE Limnephilus rothi Denning Limnephilus rothi Denning, 1966, Pan-Pac. Ent. 42: 235. This species, recently described from southern Arizona, is recorded from Mexico for the first time. Base Camp: 9 June 1964, 1¢; 16 June 1964, 19; 17 June 1964, 19; 18 June 1964, 1¢ 19; 29 June 1964, 19; 10 July 1964, 14; 22 July 1964, 14; 25 July 1964, 19. Limnephilus pollux, n. sp. (Figs. 7-11) This species is related to morrissoni and castor; from both it is easily separated by the much reduced tenth tergites, and from castor by the more elongate claspers. Adult. Length of forewing, male 12-14 mm., female 13-15 mm. Forewings brown, with considerable white fenestration around anastamosis, and with oblique paler mark basad. Basal tarsal segment of foreleg about 11% times length of second segment. Spurs yellow, 1, 3, 4; foretibia and femur without dark spinulose bands. Male genitalia: Eighth tergum produced into a bilobate process covered with dark spicules. Ninth segment narrow ventrally; clasper completely united to posterior margin of segment, but dorsally produced into a long, finger-like lobe whose apex is directed mesad. Cercus elongate, directed laterad, margins subparallel, mesal face with a slightly produced, black, rugose stripe. Tenth tergite a compressed trianguloid plate, dorsal margin darkened apically. Aedeagus with lateral arms membranous apically, with dorsal margin bearing a dense fringe of setae. Female genitalia: Ninth sternum slightly produced apicolaterally, in ventral aspect broadly and shallowly emarginate mesally. Cercus fused to tenth segment, but produced as a pair of short acute lobes, tenth segment a short tube. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’. 17 July 1964, J. E. H. Martin. Allotype, female. Same data. Paratypes. Same but, 8 June 1964, 3¢; 17 June 1964, 12; 4 July 1964, 12: 8 July 1964, 1¢ 19. Limnephilus toussainti Banks Limnephilus toussianti Banks, 1924, Bull. Mus. Comp. Zool. 65: 439. Limnephilus toussainti Banks: Flint, 1963, Proc. Ent. Soc. Wash. 65: 211. The original spelling of the specific name is an obvious misspelling of the name of the Haitian general Toussaint, and was emended in 1963, but without comment. 170 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 The type of this species bears a label indicating it is from Port-au-Prince, Haiti, Wm. Mann. This is apparently a mislabelled specimen. During con- versations with Dr. Westfall I have learned that there are several Odonata with similar labels, but which are actually from central Mexico. The specimens are from the base camp on 13 June 1964, 14; 17 June 1964, 19; 29 June 1964, 146; 2 July 1964, 19. Limnephilus ctenifer, n. sp. (Figs. 12-16) This is a species belonging to the diversus group, but which is easily recognized by the shape of the clasper, cercus, and dorsal armature of the lateral arms of the aedeagus. Adult. Length of forewing, 9-10 mm. Color almost uniformly brown, fore- wings with darker flecks along veins, and with membrane faintly irrorate. Basal tarsal segment of forelegs subequal in length to second segment; forefemur with a shallow groove on ventral face, both this and ventral surface of foretibia densely covered with black spicules; apex of foretibia with a broad hoodlike process, if this is a true spur, then formula is 1, 3, 4. Male genitalia: Eighth tergum with a small triangular, apicomesal, spiculate patch. Ninth segment broad laterally, with clasper united to posterior margin and bearing an upright, darkened, pointed process. Cercus elongate, tapering to a mesally directed point, with posterior margin dark and dentate. Tenth tergite with a small, posteriorly- directed, dark point arising from a large flat base. Aedeagus with heavily sclerotized lateral arms bearing an apicodorsal comb of spines, and apicoventrally bearing slender setae. Female genitalia: Ninth segment continuous dorsally, gradually widened ventrally. Cercus in lateral aspect a pointed lobe, dorsally an almost continuous broad plate. Tenth segment a slightly longer tube, emarginate mesally. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’, 26 June 1964, W. C. McGuffin. Allotype, female. Same but, 25 June 1964, J. E. H. Martin. Paratypes. Same but, 17 June 1964, 12; 21 June 1964, 12; 25 June 1964, 2°; 29 June 1964, 26 12; 1 July 1964, 1é; 4 July 1964, 1¢; 7 July 1964, 12; 17 July 1964, 1¢. Limnephilus biparta Denning Limnephilus biparta Denning, 1966, Can. Ent. 98: 1223. This species, recently described from specimens taken at the base camp, is apparently rather common there. Base Camp: 9 June 1964, 1¢; 25 June 1964, 22; 28 June 1964, 26; 29 June 1964, 29; 30 June 1964, 19; 1 July 1964, 12 39; 2 July 1964, 29; 4 July 1964, 19; 6 July 1964, 46 19; 8 July 1964, 33 22; 10 July 1964, 1¢ 29; 11 July 1964, 19; 12 July 1964, 5¢ 59; 15 July 1964, 1g 29; 17 July 1964, 12; 18 July 1964, 1g 19; 23 July 1964, 34; 28 July 1964, 24 59; 29 July 1964, 84; 30 July 1964, 34; 31 July 1964, 34 19; 1 Aug. 1964, 36 19; 3 Aug. 1964, 16; 5 Aug. 1964, 1¢ 19; 8 Aug. 1964, 74; 9 Aug. 1964,5¢ 39; 10 Aug. 1964, 1¢4. PROC. ENT. SOG. WASH., VOL. 69, No. 2, JUNE, 1967 171 Limnephilus mexicanus, n. sp. (Figs. 17-21) This is a species of the assimilis group, close to L. frijole Ross. It is rec- ognized by the lack of a mesal tooth on the cercus and the presence of a dorsal comb of spines on the lateral arms of the aedeagus. Adult. Length of forewing, male 10-12 mm., female 11-14 mm. Color brown, forewing irrorate with darker brown, veins spotted with dark brown. Basal segment of foretarsus of male half length of second segment; femur and tibia of foreleg with mesal surface bearing a row of dark spicules; spurs 0, 2, 3. Male genitalia: Eighth tergum with an apicomesal patch of dark spicules. Ninth segment moderately expanded laterally. Clasper united to ninth segment, extending posteriad, about as broad as long, with dorsal margin flat. Cercus narrow and elongate, dorsomesal margin slightly inrolled, without mesal tooth. Tenth tergite about as long as broad, tapering to an upturned apical point. Aedeagus with lateral arms heavily sclerotized, dorsal margin with a comb of enlarged setae, ventrally with thin setae. Female genitalia: Ninth segment in- complete dorsally, almost twice as high as long. Cercus a poorly defined dorso- lateral swelling on tenth segment. Tenth segment tubular, slightly prolonged ventrally. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’. 8 June 1964, J. E. H. Martin. Allotype, female. Same data. Paratypes. Same but, 6 June 1964, 1¢ 2°; 8 June 1964, 8¢ 62; 11 June 1964, 22; 13 June 1964, 34 32; 16 June 1964, 34; 17 June 1964, 26 52; 18 June 1964, 34 2°; 20 June 1964, 3¢; 24 June 1964, 12; 26 June 1964, 14; 28 June 1964, 2°; 29 June 1964, 12; 1 July 1964, 22; 15 July 1964, 1¢. Durango: 3 miles west El Salto, 8500’, 19 June 1964, 1é 12; Buenos Aires, 10 miles west La Ciudad, 9000’, 11 June 1964, 1°. Clistoronia graniculata (Denning), n. comb. Limnephilus graniculata Denning, 1966, Can. Ent. 98: 1223. This species, recently described from specimens taken at the base camp, is clearly a member of the genus Clistoronia. The many small pale spots found on the forewings of the related species are nearly lacking in this species. The species is common at the base camp: 5 June 1964, 19; 6 June 1964, 49; 8 June 1964, 29; 9 June 1964, 19; 11 June 1964, 19; 16 June 1964, 39; 17 June 1964, 19; 20 June 1964, 19; 25 June 1964, 19; 26 June 1964, 14; 29 June 1964, 19; 1 July 1964, 19; 5 July 1964, 19; 6 July 1964, 146; 7 July 1964, 24; 9 July 1964, 146 19; 12 July 1964, 29; 17 July 1964, 19; 18 July 1964, 14; 22 July 1964, 2¢; 28 July 1964, 26 29; 29 July 1964, 14 19; 30 July 1964, 1¢ 19; 31 July 1964, 26 19; 1 Aug. 1964, 1¢ 49; 3 Aug. 1964, 1g; 4 Aug. 1964, 16; 7 Aug. 1964, 19; 8 Aug. 1964, 3¢ 19; 9 Aug. 1964, 1¢@; ll Aug. 1964, 19. Hesperophylax magnus Banks Hesperophylax magnus Banks, 1918, Bull. Mus. Comp. Zool. 62: 20. Ross, 1938, 172 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Figs. 17-27. Limnephilus mexicanus, n. sp.: 17, male genitalia, lateral; 18, male genitalia, dorsal; 19, aedeagus, dorsolateral; 20, female genitalia, lateral; 21, female genitalia, dorsal. Marilia “A”: 22, male genitalia, lateral. Marilia “B”: 23, male genitalia, lateral. Lepidostoma lacinatum, n. sp.: 24, male genitalia, lateral; 25, male genitalia, dorsal. Lepidostoma rectangulare, n. sp.: 26, male genitalia, dorsal; 27, male genitalia, lateral. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 1k7es% Psyche 45: 32. Banks, 1943, Bull. Mus. Comp. Zool. 92: 348. Ross, 1944, Bull. Dl. Nat. Hist. Surv. 23: 280. This species, known from the southwestern United States, is recorded from Mexico for the first time. Base Camp: 6 June 1964, 19; 13 June 1964, 14; 16 June 1964, 19; 17 June 1964, 19; 1 July 1964, 146; 4 July 1964, 19; 11 July 1964, 14; 17 July 1964, 12. ODpDONTOCERIDAE Marilia flexuosa Ulmer Marilia flexuosa Ulmer, 1905, Ann. Hofm. Wien 20: 70. Ross, 1941, Trans. Amer. Ent. Soc. 67: 100. Ross, 1951, Proc. Cal. Acad. Sci. (4th Ser.) 27: 71. Denning, 1964, Pan-Pac. Ent. 40: 134. Fischer, 1965, Trich. Catal. 6: 33. This species is widely distributed, being reported from Texas to Brazil. Pre- vious Mexican records are from Vera Cruz (Ross, 1941) and Baja California (Ross 1951, Denning 1964). Base Camp: 9 June 1964, 14; 4 July 1964, 19. Marilia species In Marilia in the Neotropical Region there exists an extremely con- fusing complex of “species” whose genitalia are all very similar. There are slight differences between each “species” not only in the genitalia, but also in the coloration. However, I had not been able to take the exact same “species” at two locations, which made me think that there might be only one (or a very few) species existing in rather isolated populations. However, there are two “species” in the collec- tion from the base camp, which make me wonder if they are not dis- tinct after all. In order to solve this question, it will take much more material than is now available. The following names are available for this group of species in southwestern North America and Central America: nobsca Milne, mexicana (Banks), and crea Mosely. The first “species” (A), fig. 22, is distinguished by the pale hairs of the forewing having a yellowish cast, having unicolorous antennae, slightly shorter and broader claspers, and a broader apex of the tenth tergum. Base Camp: 10 June 1964, 6¢. The second “species” (B), fig. 23, has the pale hairs grayish-white, annulate antennae, a more slender clasper, and a more constricted apex of the tenth tergum. Base Camp: 8 June 1964, 12; 12 July 1964, 12; 17 July 1964, 12; 6 Aug. 1964, 14; (also 29-30 June 1964, 36, Spangler coll. U.S.N.M.). CALAMOCERATIDAE Phylloicus ornatus (Banks ) Notiomyia ornata Banks, 1909, Ent. News 20: 342. Fischer, 1965, Trich. Catal. 6: 19. 174 PROG. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Phylloicus ornatus (Banks): Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 17. In coloration this specimen seems identical to the type from Texas. Sinaloa, 15 miles west of El Palmito, 5000’, 20 July 1964, 1°. Phylloicus aeneus ( Hagen ) Macronema aeneum Hagen, 1861, Syn. Neur. N. Amer., p. 285. Phylloicus aeneus (Hagen): Ulmer, 1905, Ann. Hofm. Wien 20: 78. Fischer, Trich. Catal. 6: 20. Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 17. These examples, which are the large dark-winged form generally called mexicana (Banks), belong to aeneus (Hagen) as I have recently defined the species. Base Camp: 10 June 1964, 29. Durango, 14 miles southwest of El Salto, 8000’, 30 June 1964, 19. LEPTOCERIDAE Oecetis disjuncta ( Banks ) Oecetina disjuncta Banks, 1920, Bull. Mus. Comp. Zool. 64: 351. Oecetis disjuncta (Banks): Fischer, 1966, Trich. Catal. 7: 154. Ross, 1951, Proc. Cal. Acad. Sci. (4th ser.) 27: 74. This species is extremely close to avara (Banks), but is generally larger and darker. It has been recorded (Ross, 1951) from Baja California. Base Camp: 8 June 1964, 29; 9 Jume 1964, 19; 11 June 1964, 19; 25 June 1964, 16; 1 July 1964, 16 19; 2 July 1964, 19; 4 July 1964, 29; 6 July 1964, 19; 8 July 1964, 29; 15 July 1964, 19; 29 July 1964, 19. Oecetis arizonica Denning Oecetis arizonica Denning, 1951, J. Kans. Ent. Soc. 24: 159. This species, which is known from southern Arizona, is here recorded from Mexico for the first time. Base Camp: 1 July 1964, 19; 2 July 1964, 14; 4 July 1964, 19; 20 July 1964, 19. Oecetis inconspicua ( Walker ) Leptocerus inconspicuus Walker, 1852, Catal. Neur. Brit. Mus. 1: 71. Oecetis inconspicua (Walker): Betten & Mosely, 1940, Walker Types Trich., p. 67. Fischer, 1966, Trich. Catal. 7: 149. This species is one of the commonest and most widespread New World caddis flies, being known from all over North America, the Greater Antilles, and higher elevations in Central America. Durango, 7 miles west of Durango, 7000’, 26 July 1964, 19. Leptocella dorsalis Banks Leptocella dorsalis Banks, 1901, Trans. Amer. Ent. Soc. 27: 368. Fischer, 1966, Trich. Catal. 7: 54. Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 21. This is one of the more common Central American species of the genus. Sinaloa, 21 miles east of Villa Union, 25 July 1964, 1¢. | PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 175 Leptocella gracilis Banks Leptocella gracilis Banks, 1901, Trans. Amer. Ent. Soc. 27: 369. Fischer, 1966, Trich. Catal. 7: 51. Flint, 1967, Proc. U.S.N.M., vol. 123, no. 3619, p. 21. The species seems common in the northern half of Mexico. The examples from Durango have a slightly darker ground color than the types, but otherwise seem identical. Durango, 7 miles west of Durango, 7000’, 26 July 1964, 1¢ 19. LEPIDOSTOMATIDAE Lepidostoma bakeri Flint Lepidostoma bakeri Flint, 1965, Proc. Ent. Soc. Wash. 67: 175. This species was described from southern Arizona and is here recorded from Mexico for the first time. Base Camp: 9 June 1964, 1¢. Lepidostoma bispinosa ( Ulmer) Atomyiodes bispinosa Ulmer, 1911, Ann. Soc. Ent. Belg. 55: 25. Lepidostoma bispinosa (Ulmer): Denning, 1962, Pan-Pac. Ent. 38: 39. Lepidostoma alexanderi Denning, 1962, Pan-Pac. Ent. 38: 37 (new synonymy ). This species is found from southern Arizona to Costa Rica in the mountainous regions. It is quite variable in details of the genitalia, especially the serrations along the posterior margin of the tenth tergite and exact shape of the apex of the clasper. I have seen examples with all intergradations in serration between the one with a uniform series of small teeth (alexanderi type) to the one with only a few, widely spaced, large teeth (bispinosa type). Often the two sides of the tenth tergum are not symmetrical in this regard, one side having more teeth than the other. There is 1¢ from the base camp, 17 June 1964, and 14 from Durango, 13 miles west of E] Salto, 18 June 1964. Lepidostoma knulli Ross Lepidostoma knulli Ross, 1946, Ann. Ent. Soc. Amer. 39: 280. Described from southern Arizona, this species is recorded from Mexico for the first time. Base Camp: 7 July 1964, 1¢; 8 July 1964, 1¢. Lepidostoma lacinatum, n. sp. (Figs. 24-25) This species is a member of the unicolor group, probably closest to the following new species. The large, serrate apicoventral lobe and the flattened, twisted apex of the clasper are diagnostic for this species. Adult. Length of forewing, 10-12 mm. Color brown, antennae and legs paler, forewing brown with scattered golden setae. Basal antennal segment 3 times as long as broad; maxillary palpus 1l-segmented, concave mesally with specialized setae; other secondary sexual modifications lacking. Male genitalia: Tenth tergite with apex upturned, dentate, halves widely separated mesally. 176 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Clasper rather broad, apex produced into a flattened plate twisted mesally, with a large serrate apicoventral lobe; basidorsal process with dorsal arm short, apical arm long and flattened, appressed to clasper, with apex produced into a hook. Aedeagus with a pair of pointed dorsal rods appressed to central tube. Holotype, male. Mexico (Durango), 10 miles west of El Salto, 9000’, 2 July 1964, J. E. H. Martin. Paratypes. Same data, 34; 6 June 1964, 1¢6; 8 June 1964, 2¢; 13 June 1964, 1¢; 17 June 1964, 3¢; 27 June 1964, 14; 7 July 1964, 146; 15 July 1964, 1¢; 29 July 1964, 1¢. Mexico (Sinaloa), El] Palmito, 15 July 1964, J. F. McAlpine, 1¢. Lepidostoma rectangulare, n. sp. (Figs. 26-27 ) This species is likewise a member of the unicolor group, and probably most closely related to the preceding species. The very broad apex and small basidorsal lobes of the clasper render this species very distinctive. Adult. Length of forewing, 10 mm. Color brown; legs and antennae slightly paler, forewing spotted with golden hairs. Basal antennal segment about 4 times as long as broad; maxillary palpus 1-segmented, concave mesally with specialized hairs; no other sexual modifications. Male genitalia: Tenth tergite rounded in lateral view, with posterior margin bearing small teeth, halves broadly separated in dorsal aspect. Clasper elongate, apex broad and truncate in lateral view, with a small subapical ventral process, apex bearing a mesal lobe in dorsal aspect; basidorsal processes short. Aedeagus with 2 stout dorsal processes whose tips are angled mesad. Holotype, male. Mexico (Durango), 10 miles west of El] Salto, 9000’, 25 July 1964, W. C. McGuffin. HELICOPSYCHIDAE Helicopsyche mexicana Banks Helicopsyche mexicana Banks, 1901, Trans. Amer. Ent. Soc. 27: 368. Ross, 1944, Bull. Ill. Nat. Hist. Surv. 23: 289. Denning, 1964, Pan-Pac. Ent. 40: 133. The species is widespread in Mexico and southwestern United States. Base Camp: 17 June 1964, 19; 18 June 1964, 19; 17 July 1964, 14. Sinaloa, 8 miles west of El Palmito, 19 July 1964, 1¢. Chihuahua, Mesa de Huracan (108°15’, 30°4’), 7400’, 21-25 July 1964, 19. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Wee, A NEW BRACONID PARASITE OF THE POTATO TUBERWORM ( HYMENOPTERA ) Cart F. W. Mueseseck, U. S. National Museum, Washington, D.C. 20560 Most of the species of the braconid genus Orgilus, all presumably internal parasites of lepidopterous larvae, are undescribed. I have undertaken a study of the North American species, in which the South American form described here is included because of its ap- parent establishment in California. Advance publication of this de- scription has been requested in order that a name may be available for use in a paper now in preparation at the Citrus Research Center, Riverside, California, that deals with the biology of the parasite. Orgilus lepidus, n. sp. This is rather similar to O. lateralis (Cresson), from which it may be dis- tinguished, however, by its smoother abdomen and by the poorly developed sub- basal tooth of the tarsal claw. From the only other known South American species of the genus, O. gossypii Muesebeck, it differs in its relatively shorter radial cell, its smoother abdomen, much weaker dorsal keels of the first tergite and its darker coloring. Female.—Length about 4 mm. Head not wider than thorax, in dorsal view 1.8 times as wide as long; face 1.2 times as wide as eye height, sharply punctate; malar space slightly more than one-third as long as eye height; temples gradually receding, 0.6 as wide as eyes, smooth and shining; occipital carina strong laterally, very weak medially; ocellocular line more than twice as long as diameter of an ocellus; vertex finely punctate; antennae 29- or 30-segmented in the available specimens, even the shortest segments of the flagellum slightly longer than broad. Thorax stout; mesoscutum shining and largely smooth, with only scattered punctures; notauli weakly foveolate anteriorly, more strongly so posteriorly; disc of scutellum rather flat, polished; propodeum evenly convex, largely alutaceous to granulose with some weak, indefinite rugulae medially; side of pronotum finely roughened below the impression, smooth and polished above it; mesopleuron smooth and polished, the longitudinal furrow sinuate and finely foveolate; meta- pleuron largely smooth. Hind coxa evenly coriaceous and rather mat; hind femur 1.5 times as long as hind coxa and 3.8 times as long as its maximum width; inner calcarium of hind tibia slightly more than half as long as metatarsus; tarsal claw with a subbasal angulation but without a distinct subbasal tooth as in lateralis. Radial cell on wing margin about 1.2 times as long as stigma; second abscissa of radius nearly on a line with intercubitus; stub of third abscissa of cubitus punctiform; nervulus clearly postfurcal; lower abscissa of basella a little less than half as long as mediella or than maximum width of hind wing. Abdomen a little narrower than thorax; first tergite nearly one and one-half times as long as broad at apex, evenly coriaceous, the dorsal keels weak and extending only slightly beyond spiracles; second tergite about 1.25 times as wide at base as long, more or less coriaceous or granulose medially, broadly 178 PROG. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 SN =< ae Figs. 1 and 2. Orgilus lepidus, n. sp. Fig. 1, part of fore wing; fig. 2, tarsal claw. smooth laterally; third and following tergites smooth and polished; ovipositor sheath just about as long as head, thorax and abdomen combined. Yellowish brown; antennae brownish black; palpi piceous; a spot on frons and vertex enclosing the ocelli, the mesopectus and propodeum black; tegulae and wing bases reddish yellow; wings hyaline; legs brownish yellow, the hind coxa more or less darkened inwardly toward base, and also the hind femur along its upper edge, the hind tibia blackish with a conspicuous pale annulus at extreme base, all tarsi darkened; abdomen with apex of first tergite and all of the second and third tergites usually brownish yellow, the remainder of the dorsum of the abdomen black. Male.—Essentially like the female but usually darker, especially on the coxae. The antennae are more slender and are 28- to 30 segmented in the available specimens. Holotype—USNM 69536. Described from 5 females (one, the holotype) and 5 males (one, the allotype) reared from the potato tuberworm, Phthorimaea operculella (Zeller), at Balcarce, Argentina; 6 females and 6 males labeled as having been reared from lepidopterous larvae on Chenopodium at Montivideo, Uruguay, in 1946 by H. L. Parker, and 1 female which is said to have been reared from P. operculella at Moreno, California, July 20, 1965, by E. R. Oatman. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 179 CIMEX HEMIPTERUS (FABR.) FROM BATS IN COLOMBIA, SOUTH AMERICA (HEMIPTERA: CIMICIDAE ) C. J. MarinxELLE, Depts. of Biology and Microbiology, Universidad de los Andes, Bogotd, Colombia (S.A. ) The common human bed bug, Cimex lectularius L., is known to parasitize a wide variety of mammals and domestic birds. Accord- ing to Usinger (1966) Cimex species originated phylogenetically from bats, but some adapted to other hosts. While C. hemipterus (Fabr.) is mainly a parasite of man in tropicopolitan areas, it has only been recorded a few times from bats: Scotophilus kuhli Leach and S. heathii (Horsfield) from India in 1908 and 1912, S. temmincki (Horsfield) from Java (Indonesia) in 1913, unidentified bat species from Java in 1932 (Usinger loc. cit.) and Myotis nigricans extremus Miller from Mexico in 1936 (Hoffman 1944). It has also been found on poultry in Jamaica and in nests of common swifts in India ( Usinger ocx cit.) During a survey of zoonotic diseases in Colombia, 3686 bats were examined for the presence of ectoparasites. Approximatively 400 of these were obtained from houses infested with C. lectularius or C. hemipterus. Two bats, Peropteryx macrotis (Wagner), collected from a hollow tree near the village of La Plata (Depart. Huila) were infested with numerous C. hemipterus. The parasites were firmly adhered to the skin near the dorsal part of the tail base and some of them were inflated with the hosts blood. None of the bats examined showed infestation with other species of Cimicidae. Some Cimicidae are capable of transmitting trypanosomes among bats (Van Den Berghe et al. 1963) and human bedbugs have been suspected of being able to transmit Trypanosoma cruzi. Because both man and bats often harbor T. cruzi in Colombia (Marinkelle 1966), the finding of C. hemipterus on bats may prove to be of importance in the epidemiology of south american trypanosomiasis. To my knowledge, this is the first case of C. hemipterus parasitizing a non-human host in the area south of Mexico to be reported. Acknowledgements: I am very grateful to Dr. G. M. Kohls (R. M. L., Hamilton, Montana) for his help in checking the literature and to Dr. C. O. Handley (U. S. Nat. Mus., Washington, D. C.) for the identification of the bat species. This. investigation was partially supported by U. S. Army Grant DA-ARO-49-092-66-G 109. REFERENCES Hoffmann, M. A. 1944. Ectoparasitos de murciélagos mexicanos. Tesis Univ. Nac. Mexico, 150 pp. 180 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Marinkelle, C. J. 1966. Observations on human, monkey and bat trypanosomes and their vectors in Colombia. Trans. Roy. Soc. Trop. Med. Hyg. 60(1): 109- 116. Usinger, R. L. 1966. Monograph of Cimicidae (Hemiptera—Heteroptera). The Thomas Say Foundation 7: 585 pp. Van Den Berghe, L., M. Chardome, and E. Peele. 1963. An African bat trypanosome in Stricticimex brevispinosus Usinger, 1959. J. Protozool. 10; 135- 138. PALAEOSEPSIS SPECIES DESCRIBED BY MELANDER AND SPULER (DireTEeRA: SEPSIDAE ) Three species described in the genus Sepsis by Melander and Spuler (1917) were placed by Duda (1926) in his subgenus Palaeosepsis, a taxon which I believe to be of generic rank. The Melander and Spuler type specimens are now in the United States National Museum collections. I have examined them with the following results. Full citations will appear in the pertinent part of the neotropical catalogue of Diptera, now in process of publication. Palaeosepsis armillata (Melander and Spuler), 1917: 18, 81 (fig. 11, not 6). Curran (1928:75) claimed that Duda (1926:92) confused this species with P. furcata (M. and S.). Examination of the type specimens from “Hayti,” one of which is hereby designated lectotype, shows that Duda’s identification is correct and that Melander and Spuler evidently intermixed the captions for the figures of the two species. P. furcata (M. and S.), 1917: 19, 81 (fig. 6, not 11). See above. One of the originally cited males is labelled as “type.” P. hoplicnema (M. and S.), 1917: 17, 81 (fig. 12). The unique male type shows that Duda’s figure (1926: 107, fig. 77) is more accurate than that of the original description. The species, as stated by Duda, is quite certainly a synonym of P. armata (Schiner). Literature Cited—Curran, C. H. 1928. Diptera or two-winged flies. N. Y. Acad. Sci., Sci. Surv. Porto Rico and the Virgin Is. 11 (pt. 1): 1-118; Duda, O. 1926. Monographie der Sepsiden (Dipt.), I. Ann. Naturhist. Mus. Wien 40; 1- 110; Melander, A. L., and A. Spuler. 1917. The dipterous families Sepsidae and Piophilidae. State College Wash. Agric. Expt. Sta. Bull. 143: 1-(103).— GrorcE C. StTEYSKAL, Entomology Research Division, ARS, U. S. Department of Agriculture, Washington, D. C. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 181 A NEW ATELEUTE FROM THE UNITED STATES (HyMENOPTERA, ICHNEUMONIDAE ) Henry Townes, American Entomological Institute, Ann Arbor, Mich. The genus Ateleute Foerster is represented by many species in the warmer parts of the Old World, where it has been reared several times from Psychidae. It has not yet been reported from the Western Hemisphere. Mr. Peter Hattenschwiler, of Uster, Switzerland, on a visit to the United States in 1963, reared a specimen of this genus at Greenville, South Carolina from a psychid, along with some other parasites of Psychidae from several localities in the Carolinas. Although Ateleute has not been reported previously from America, Mr. Hattenschwiler’s is not the first specimen known. Some years ago, I saw two male specimens in the U. S. National Museum that were collected in south central United States. In 1965, Mr. John Schmid found a male of the genus among Malaise trap material from Golden Pond, Kentucky, collected by Dr. Sam Breeland. These three males may be the same species as the female described below, but males of Ateleute are difficult to associate with their females, as there is strong sexual dimorphism. The genus occurs also in South America. I have seen three undescribed species from Peru. Ateleute is an isolated genus of the subfamily Gelinae. By “key characters” it belongs in the tribe Mesostenini, but the genus may be a derivative of the Chirotica group of genera in the Gelini, and thus may have independently developed mesostenine characters. For the present, I prefer to place it in the Mesostenini. It should be in a subtribe by itself: the Ateleutina. The more distinctive characters of Ateleute are: Postpectal carina complete, not interrupted in front of each middle coxa; mesopleural fovea absent; areolet moderately large, about twice as wide as high, the second intercubitus weak or absent; second recurrent vein vertical, with one bulla; tergite 1 often longitudinally striate, its spiracle at the midlength; propodeum long, its apical transverse carina usually dis- tinct, its basal transverse carina absent or indistinct, without distinct longitudinal carinae. Ateleute carolina, n. sp. Male: Unknown. Female: Front wing 3.7 mm. long; postero-median part of mesoscutum with longitudinal wrinkles; apical transverse carina of propodeum distinct, the basal carina obsolete; propodeum in front of apical transverse carina mat and with weak transverse wrinkling; second intercubitus faint; tergite 1 weakly mat, medially subpolished, with a median longitudinal impression, without any longi- tudinal striae; tergite 2 weakly mat, without punctures or wrinkles; ovipositor sheath 0.6 as long as front wing. 182 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Fulvous, with the head black. Clypeus, palpi, and propleurum not visible because glued next to card mount; mandible whitish; antenna black, the front half of scape ferruginous, apical 0.3 of flagellar segment 5, all of flagellar seg- ments 6-9, and basal 0.6 of flagellar segment 10 white; pronotum fuscous, its front part, lower part, and hind corner white; mesonotum and metanotum black, the lateral lobe of mesoscutum margined with white mesally and anterolaterally, the median lobe of mesoscutum broadly margined with white anterolaterally; apex of scutellum white; subtegular ridge whitish; mesopleurum with a fuscous area beneath front wing; legs pale fulvous, their fifth tarsal segments infuscate. Distinctive characters of this species are the lack of longitudinal striae on tergite 1, and the color pattern of the thorax. Type 2, Greenville, South Carolina, emerged from case of Astala confederata (Psychidae), Feb. 23, 1963, Peter Hiattenschwiler (Townes ). NEW SYNONYMY IN PLERONEURA KONOW (HYMENOPTERA: XYELIDAE) Davin R. Smitrn, Entomology Research Division, ARS, U. S. Department of Agriculture, Washington, D.C. 20560 Ross (1932) revised the North American Pleroneura Konow and included 7 species. He stated that the species had been described from very few specimens and that larger series would be necessary in order to determine how stable the characters are on which these species were based. In recent years, several large series of Pleroneura have been collected in the East and West, and it is now possible to evaluate some of the characters which were previously used for species separation. P. brunneicornis Rohwer and P. borealis Felt were both described from New York State and are the only described eastern forms. Ross separated these species from the western Pleroneura by the pale pos- terior margin of the pronotum and pale femora, then distinguished between them by veins r-m and m-cu of the fore wing being interstitial or not. In series of a species of Pleroneura collected in Maine, the position of veins r-m and m-cu of the fore wing was found to be variable; consequently, I consider these two species to be the same. The western species of the group, in which the body is primarily black, were separated by the dark or light coloration of the tibiae, femora, and clypeus. Several series of a species of Pleroneura which were collected in Oregon and California show a high degree of color variation within and between sexes. In some instances, the associated males and females will key to different species in Ross’ key. Therefore, PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 183 I believe that the 3 western species, californica Ashmead, schwarzi Rohwer, and aldrichi Ross, as recognized by Ross, are the same. The types of these species also fall within the range of variation which I have found. The following synonymy is proposed: Pleroneura brunneicornis Rohwer Pleroneura bruneicornis (!) Rohwer, 1910, Can. Ent. 42: 89, 9. Pleroneura borealis Felt, 1917, Can. Ent. 49: 220, 6, 9. New synonymy. Pleroneura californica (Ashmead ) Manoxyela californica Ashmead, 1898, Psyche 8: 214, 9°. Pleroneura fulvicornis Rohwer, 1910, Can. Ent. 42: 89, ¢. Synonymy by Ross, 1951, confirmed. Pleroneura schwarzi Rohwer, 1910, Can. Ent. 42: 220, 9. New synonymy. Pleroneura aldrichi Ross, 1932, Ann. Ent. Soc. Amer. 25: 158, ¢, 2. New synonymy. Two other western species, koebelei Rohwer and lutea Rohwer, were recognized by Ross, and they may be separated from californica by their predominately yellow coloration and genitalic characters. As yet, I have not seen additional specimens of these forms. P. brunneicornis and P. californica may be separated by genitalic characters, and the larvae of brunneicornis also differ from larvae which may be californica taken from the twigs of fir. The eastern brunneicornis has been reared from larvae boring in the twigs of fir, whereas californica is associated with fir only by adult collections and oviposition observations. The hosts and habits of koebelei and lutea are not known. REFERENCES Ross, H. H. 1932. The Hymenopterous family Xyelidae in North America. Ann. Ent. Soc. Amer. 25: 153-169. 1951. Xyelidae. in Muesbeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog. U. S. Dept. Agr., Agr. Monogr. 2: 5-7. LECTOTYPE DESIGNATION FROM THREE SYNTYPES OF RADFORDIA SUBULIGER EWING (AcARINA: MYOBIIDAE) The need for the designation of a lectotype for the species Radfordia subuliger Ewing became apparent when it was found that Ewing (1938, Proc. Ent. Soc. Wash. 40: 180-197) did not designate a holotype for R. subuliger Ewing. The type material upon which Ewing based his new description has been studied by the author by the loan of the three syntypes deposited in the United States Na- tional Museum. 184 PROC, ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 Syntype : Ov harvest mouse Radfordia : c.PuL pea ° Su buliger Co/lese Park jd. n- sr: Rag Greenfield, Coll. Zags Ty Pes Apr. 6, 1732 v, Lectotype Fig. 1. Radfordia subuliger Ewing, female lectotype designation on type slide No. 1280 USNM. Ewing (1938) described R. subuliger from two females which were mounted on a single slide, designated as Type No. 1280 USNM and from one female mounted on a separate slide with a label that read at the time of the description as Myobia subuliger n. sp. This slide is from a different host than that of the type slide and collected on a different date. Also this slide is in very poor condition. The selection of the lectotype by the author has been made from the slide bear- ing the Type No. 1280. Due to the possible destruction of the type material and the slide labels, no attempt has been made to remount onto single slides these two syntypes. Indication to the lectotype is possible by the deterioration of the slide median. A drawing showing the location of the lectotype and syntype on the Type No. 1280 USNM is shown in fig. 1. This drawing has been attached to the back of the type slide. B. McDANrIEL, Department of Entomology- Zoology, South Dakota State University, Brookings, South Dakota 57006. Approved by the Director of South Dakota Agricultural Experiment Station as Journal Series No. 763. A LIST OF THE SPECIES OF MACROSIPHONIELLA DEL GUERCIO DESCRIBED FROM NORTH AMERICA (HomMortTEeRA: APHIDIDAE) LoutsE M. Russet, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D. C. 20250 Twenty-one species assignable to Macrosiphoniella (sensu lato) have been described from North America, but only seven have been placed in this genus. In this paper, names of the 14 remaining species are associated with Macrosiphoniella to make new name com- PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 185 binations available. The 21 species are listed, and their original cita- tions and previous name combinations are given. One North Amer- ican species included in the genus because of a misidentification is discussed. Five species of Macrosiphoniella (absinthii (Linnaeus), leucanthemi (Ferrari), millefolii (De Geer), tanacetaria (Kalten- bach), and trimaculata Hille Ris Lambers) described from Europe also have been reported from North America. The repository of type specimens is given when known. Additional types of some species presumably are present in other collections. Macrosiphoniella Del Guercio 1911 is closely related to Dactynotus Rafinesque 1818. Indeed, Eastop in 1961 (A study of the Aphididae (Homoptera) of West Africa p. 19) included atra (Ferrari) 1872, the type-species of Macrosiphoniella, in Dactynotus. In 1966 (Austral. Jour. Zool. 14: 455) however, he recognized Macrosiphoniella as a valid genus. Takahashi and Moritsu (Mushi 37: 1, 1963) stated that Macrosiphoniella “can not be definitely separated from Dactynotus . . . but is here maintained as a valid genus, since the trend of the modifi- cation of the body structures is different from that of the latter genus.” Although most species may readily be placed in one group or the other, border forms are difficult to assign. Future studies doubtless will show whether Macrosiphoniella should be considered of generic rank. The genus is now accepted by workers though perhaps with- out critical consideration of the reasons for doing so. The salient characteristics of Macrosiphoniella are as follows: Last rostral segment slender, tapered, often stiletto-shaped with acute tip, with setae located on basal portion of segment longer than the 3 apical pairs (much longer if segment is stiletto-shaped). Cornicles reticulated at least near apex; nearly or actually flangeless; cylindrical, or enlarged basally and sometimes slightly so distally; slightly shorter than, to twice as long as, cauda; usually without, but sometimes with, a few setae. Frontal tubercles distinct, diverging. Setae on antennal segment III blunt or enlarged apically, usually not acute. Tergum membranous but scleroites often present around bases of setae; sclerites absent, or present anterior and/or posterior to the cornicles; 4 or more setae on tergum 8. Each Ist tarsal segment with 3 setae. Apterae with secondary sensoria on antennal segment III. Most species holo- cyclic on Anthemideae (Compositae ). Macrosiphoniella can be distinguished from Dactynotus by_ its tapered, often stiletto-shaped beak with the basal setae of the last rostral segment usually much longer than the apical ones; by its flangeless or almost flangeless cornicle; and by the presence of 3 setae on each lst tarsal segment. None of these conditions pertain to Dactynotus except the last, where the Ist tarsal chaetotaxy may be 3 but more often is 4 or 5. 186 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 In the following list, M. after the species and author name refers to Macrosiphoniella. albicornus (Knowlton and Allen) M., n. comb. Mucrotrichaphis albicornus Knowlton and Allen 1940, Ohio Jour. Sci. 40: 33. Macrosiphum albicornum (Knowlton and Allen), Palmer 1952, Aphids of the Rocky Mountain Region p. 290. Holotype—USNM. anomellus (Knowlton and Allen) M., n. comb. Macrosiphum anomellus Knowlton and Allen 1938, Can. Ent. 70: 75. Mucrotrichaphis anomellus (Knowlton and Allen), Knowlton and Allen 1940, Ohio Jour. Sci. 40: 34. Cotypes—USNM. artemisiae Cowen—see coweni Hunter. artemisiphilus (Knowlton and Allen) M., n. comb. Macrosiphum artemisiphilus Knowlton and Allen 1938, Can. Ent. 70: 75. Cotypes—USNM. artemisivulgaris (Knowlton and Allen) M. Macrosiphum artemisivulgaris Knowlton and Allen 1938, Can. Ent. 70: 76. Macrosiphoniella artemisivulgaris (Knowlton and Allen), Quednau 1966, Can. Ent. 98: 428. Cotypes—USNM. cefsmithi (Knowlton and Allen) M., n. comb. Macrosiphum cefsmithi Knowlton and Allen 1938, Can. Ent. 70: 76. Cotypes—USNM. chrysanthemi Oestlund—see sanborni (Gillette ). chrysanthemicolens Williams—see sanborni (Gillette). cinerescens Hille Ris Lambers, M. Macrosiphoniella cinerescens Hille Ris Lambers 1966, Hilgardia 37: 588. Holotype—Hille Ris Lambers colln.; paratypes—Univ. of California, Berkeley. coweni (Hunter) M., n. comb. Nectarophora artemisiae Cowen in Gillette and Baker 1895, Colo. Agr. Expt. Sta. Bul. (Tech. Ser. 1) 31: 123 (preoccupied). Nectarophora coweni Hunter 1901, new name, Iowa Agr. Expt. Sta. Bul. 60: 114. Macrosiphum coweni (Hunter), Sanborn 1906, Kans. Univ. Sci. Bul. 3: 232; Gillette and Palmer 1934, Ent. Soc. Amer. Ann. 27: 175 (selected neotype and paraneotypes ). Neotype and paraneotypes—USNM. essigi (Soliman) M., n. comb. Obtusicauda essigi Soliman 1927, Calif. Univ. Pubs. Ent. 4: 99. Macrosiphum essigi (Soliman), Knowlton and Allen 1938, Can. Ent. 70: 78. Dactynotus essigi (Soliman), Eastop 1961, A study of the Aphididae (Homop- tera) of West Africa p. 19. Cotypes—Univ. of California, Berkeley. filifoliae (Gillette and Palmer) M., n. comb. Macrosiphum filifoliae Gillette and Palmer 1928, Ent. Soc. Amer. Ann. 21: 5. Cotypes—USNM. PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 187 flavila (Knowlton and Allen) M., n. comb. Mucrotrichaphis flavila Knowlton and Allen 1940, Ohio Jour. Sci. 40: 35. Macrosiphum flavilum (Knowlton and Allen), Palmer 1952, Aphids of the Rocky Mountain Region p. 306. Cotypes—USNM. frigidae (Oestlund) M., n. comb. Siphonophora. frigidae Oestlund 1886, Minn. Geol. and Nat. Hist. Survey Ann. Rpt. 14: 20. Nectarophora frigidae (Oestlund), Oestlund 1887, Minn. Geol. and Nat. Hist. Survey Bul. 4: 83. Macrosiphum frigidae (Oestlund), Sanborn 1906, Kans. Univ. Sci. Bul. 3: 232. Obtusicauda frigidae (Oestlund), Soliman 1927, Calif. Univ. Pubs. Ent. 4: 99. Cotypes—Univ. of Minnesota, St. Paul. frigidicola Gillette and Palmer, M. Macrosiphoniella frigidicola Gillette and Palmer 1928, Ent. Soc. Amer. Ann. Pals Ss Macrosiphum frigidicola (Gillette and Palmer), Gillette and Palmer 1934, Ent. Soc. Amer. Ann. 27: 180. Cotypes—USNM. glabrum (Gillette and Palmer), M., n. comb. Macrosiphum glabrum Gillette and Palmer 1928, Ent. Soc. Amer. Ann. 21: 2. Cotypes—USNM. jonesi (Gillette and Palmer) M., n. comb. Macrosiphum jonesii Gillette and Palmer 1928, Ent. Soc. Amer. Ann. 21: 7. Cotypes—USNM. ludovicianae (Oestlund) M. Siphonophora ludovicianae Oestlund 1886, Minn. Geol. and Nat. Hist. Survey Ann. Rpt. 14: 23. Nectarophora ludovicianae (Oestlund), Oestlund 1887, Minn. Geol. Nat. Hist. Survey Bul. 4: 80. Macrosiphum ludovicianae (Oestlund), Sanborn 1906, Kans. Univ. Sci. Bul. ss PBR, Tritogenaphis ludovicianae (Oestlund), Oestlund 1922, Minn. State Ent. Rpt. 19: 142. Macrosiphoniella ludovicianae (Oestlund), Hille Ris Lambers 1931, Tijdschr. v. Ent. 74: 176. Cotypes—Univ. of Minnesota, St. Paul. pennsylvanica (Pepper) M. Macrosiphum pennsylvanicum Pepper 1950, Fla. Ent. 33: 3. Macrosiphoniella pennsylvanica (Pepper), Hille Ris Lambers 1966, Hilgardia 37: 588. Holotype, allotype, paratypes—USNM; paratypes—Pepper colln. sanborni (Gillette) M. Siphonophora_ chrysanthemicolens Williams 1891, Univ. Nebr. Dept. Ent. Spec. Bul. 1: 8, nomen nudum. Macrosiphum chrysanthemi (Oestlund), misidentification by Sanborn 1904, Kans-Univs Sei, Bulje3:ive: Macrosiphum sanborni Gillette 1908, Can. Ent. 40: 65. 188 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 Siphonophora chrysanthemicolens Williams 1911 (1910), Nebr. Univ. Studies 10: 75, described. Macrosiphoniella sanborni (Gillette), van der Goot 1917, Indian Mus. Records (Indian Sci. Jour.) 13: 183. Pyrethromyzus sanborni (Gillette) Boerner 1952, Mitt. Thiir. Bot. Ges. 3: 168. Dactynotus (Pyrethromyzus) sanborni (Gillette), Eastop 1961, A study of the Aphididae (Homoptera) of West Africa p. 19. Cotypes—USNM. tapuskae ( Hottes and Frison) M. Macrosiphum tapuskae Hottes and Frison 1931, Ill. Nat. Hist. Survey Bul. 19: Oe Macrosiphoniella tapuskae (Hottes and Frison), Hille Ris Lambers 1938, Temminckia 3: 37. Holotype and paratypes—lIllinois Nat. Hist. Survey, Urbana; paratypes—USNM. toti (Knowlton and Allen) M., n. comb. Mucrotrichaphis toti Knowlton and Allen 1940, Ohio Jour. Sci. 40: 33; Palmer 1952, Aphids of the Rocky Mountain Region p. 332. Palmer synonymized toti with Macrosiphum zerothermum Knowlton and Allen. I do not accept this synonymy. Holotype and paratypes—USNM. zerohypsi (Knowlton and Allen) M., n. comb. Mucrotrichaphis zerohypsi Knowlton and Allen 1940, Ohio Jour. Sci. 40: 34. Macrosiphum zerohypsi (Knowlton and Allen), Palmer 1952, Aphids of the Rocky Mountain Region p. 332. Cotypes—USNM. zerothermus (Knowlton and Allen) M., n. comb. Macrosiphum zerothermus Knowlton and Allen 1938, Can. Ent. 70: 83. Cotypes—USNM. Macrosiphoniella eupatorii (Williams) of Leonard 1963 is a misidentification because, though the identity of eupatorii is uncertain, the insects do not agree with the description of this species. The specimens (“sweeps” taken in 1946) reported as eupatorii by Leonard are Microparsus desmodiorum Smith and Tuatay. Williams’ eupatorii has been recorded as follows: Siphonophora eupatoris Williams 1891, Nebr. Univ. Dept. Ent. Spec. Bul. 1: 26, nomen nudum. Macrosiphum eupatorii (Williams), Sanborn, 1906, Kans. Univ. Bul. 3: 237- 238, nomen nudum. Siphonophora eupatorii Williams 1911 (1910), Nebr. Univ. Studies 10: 77, described. Macrosiphum eupatorii (Williams), Davis 1911, Nebr. Univ. Studies 11: 281 (Reprint Nebr. Univ. Dept. Ent. Contrib. 5: 29). Macrosiphoniella eupatorii (Williams), Leonard 1963, Kans. Ent. Soc. Jour. 36: Ta: Following his description of ewpatorii, Williams stated, “On white snakeroot (Eupatorium ageratoides), in company with Aphis ageratoides Oestlund, October, 1889. Apterous females and PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 189 pupae of a Siphonophora very close to, if not identical with the above, were collected September, 1890, on Eupatorium perfoliatum at Ashland not far from the place where that species was collected, differing only in being of a more uniformly greenish-brown color and having the tail concolorous with the body. Length of body, 1.75-2.00 mm.; antennae, 2.90 mm.; honey tubes 0.40 mm. The species has affinities with Myzus.” Williams’ 1911 paper was published 11 years after his death, and its introduction as well as other brief portions were prepared by some other person. Regarding eupatorii, it is stated (pp. 6, 78) that there were no specimens in the University of Nebraska or the U. S. National Museum collections. And Davis, after including eupatorii in Macrosi- phum, stated, “I have had no opportunity of studying this species since the types are lost.” There apparently are no specimens of the eupatorii described from Eupatorium ageratoides, October 1889, and only these can be con- sidered types of this species. However, two slides bearing apterous adults and nymphs collected from E. perfoliatum at Ashland, Sep- tember 1890, are in the U. S. National Museum. These specimens are Microparsus desmodiorum Smith and Tuatay. It is evident that they are a different species than those described from E. ageratoides be- cause specimens from the latter host are said to have both the cornicles and the cauda black, with the cornicles twice as long as the cauda. In the specimens from E. perfoliatum the cornicles are black and the cauda pale, and the cornicles are slightly longer than the cauda. Thus Siphonophora eupatorii Williams 1911, will remain a nomen dubium unless type specimens of the species are found. I have not been able to locate them in the collection of the U. S. National Mu- seum. It is possible that eupatorii belongs in Macrosiphoniella or Dactynotus, for the cauda is concolorous with the dark cornicles in several species of these genera, and in some species of each genus the cornicles are approximately twice as long as the cauda. A Micro- parsus with a black cauda is not known at present though this does not preclude the existence of such a species. A Microparsus from Eupatorium (Compositae ) would be of interest because species of this genus have been known with certainty only from Desmodium (Leguminosae). It seems questionable whether Williams would have confused Eupatorium ageratoides and E. per- foliatum with Desmodium, however, for he also described Sipho- nophora desmodii from Desmodium canescens. S. desmodii Williams 1911, whose types are in the USNM, was first described as Microparsus variabilis Patch 1909. 190 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 SOPHY I. PARFIN 1918-1966 Photo taken in January 1947 Sophy I. Parfin was born in New Haven, Connecticut, April 22, 1918, and died in New Britain, Connecticut, October 28, 1966. She had retired on disability September 15, 1960, from the Smithsonian In- stitution, Washington, D. C., where she was an entomologist on the staff of the U. S. National Museum, working on the systematics of Neuroptera and related insects. It is particularly tragic that her health failed at such an early age, before the full potential of a scientific career to which she was deeply devoted could be realized. Her parents, John Parfinowich and Anastia Hamego, were born in Russia; her mother died in 1919. Sophy preferred the shorter sur- name Parfin, and in 1950 obtained official court permission for the change. She seldom used her full middle name, Ivanovna, which by Russian custom means “daughter of John”. Following graduation from Meriden High School, she attended Bethel Junior College, St. Paul, Minnesota (1935-37), and was the Salutatorian of her class. She then attended Wheaton College, Whea- ton, Illinois, where she received honors and the B. S. degree in 1939, with a major in zoology. She spent an additional academic year at Wheaton taking education courses and obtaining a teacher's certificate. At both Bethel and Wheaton she studied Bible extensively. German and zoology, the latter including one course in entomology, were strongly represented at Wheaton. Sophy’s record shows a dedicated PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 191 interest in advancing her education. In New Jersey during 1943 she attended night classes, especially in English, at the Newark College of Engineering. Later, during her Washington, D. C. residence, she studied medical entomology and Russian at the U. S. Department of Agriculture Graduate School (1944); morphology, ecology, and physiology of insects at the University of Maryland (1946-47 ); mycol- ogy at George Washington University (1946-47). During her em- ployment at the Smithsonian Institution, she took an unpaid leave of absence to study entomology at the University of Minnesota (July 1, 1947-June 30, 1949); the original l-year leave was followed by 3 shorter extensions. Her studies at Minnesota led to an M. S. degree, also to her 1952 paper on the Megaloptera and Neuroptera of Minnesota. She was active in extra-curricula activities in college. She was a member of the glee club, and also played the violin, mandolin, and piano. On the rifle team she won medals as a sharpshooter; one year she was college table tennis champion. She was a good tennis player and swimmer, and was active in both sports after college. In Wash- ington she was chairman of the sports committee, American Associa- tion of University Women. Miss Parfin took pride as a woman in scientific work, and became secretary of the Washington Chapter of Sigma Delta Epsilon, a professional society of women scientists. She was also a member of the Entomological Society of Washington. Following her retirement, she was a member of the Calvary Baptist Church of Meriden. During parts of 1940-42 Miss Partin was a substitute teacher in Connecticut high schools, especially Meriden High School, by day, and worked nights at the New Departure Manufacturing Co. In September-November 1941 she taught biology and mathematics in high school at Orono, Maine. In 1942 she began work as an as- sistant mechanical engineer at the Western Electric Co., Kearney, N. J., where she assembled and inspected ball-bearing components; she also instructed new employees. The work at Western Electric was considered important to the war effort (World War II), and she had to obtain special permission from the War Manpower Com- mission to leave there and to join the staff of the Smithsonian Institu- tion. At that time the Smithsonian had many demands for the handling and identification of mosquitoes and other insects of medical impor- tance from overseas and domestic sources. Miss Parfin began Smith- sonian employment as a Scientific Aid in the Division of Insects, U. S. National Museum, on January 17, 1944, under a War Service Appointment. With the exception of two years at the University of Minnesota, she remained at the Smithsonian until her retirement, _ advancing through the ranks of Preparator and Junior Entomologist to Assistant Curator. 192 PROC. ENT. SOC. WASH., VOL. 69, No. 2, JUNE, 1967 At the Smithsonian Institution, Miss Parfin made identifications and provided consultation in several orders of Neuropteroid insects, expecially the Neuroptera, and curated them in the small but grow- ing collection. She took an avid interest in research, and it was her nature to inquire into all details of a problem insofar as information was available. Because of her limited prior experience in the tech- nicalities of Neuroptera systematics, and the time required for service responsibilities, completion of manuscripts was necessarily slow; a list of her publications follows. The most important paper is that on Sisyridae, small brown lacewinged Neuroptera which as larvae are parasitic on freshwater sponges. Most of her available research time for two years or more was given to the preparation of this paper. At least two years before Miss Parfin’s retirement, her health be- came affected by arthritis and associated disease. It became increas- ingly difficult for her to continue the physical requirements of han- dling museum drawers, making her usual painstaking dissections, and the like. Her associates and other friends were distressed by the burdens which detracted from her previous glad-hearted and en- thusiastic personality. In the six years of her retirement, during which she lived with her father and her devoted step-mother, her health continually worsened from the onslaught of arthritis and its com- plications. In closing this biographical account of a lamented colleague whose life and career were interrupted just as the fully productive years were approaching, we pay further tribute to her high standards of scholarship, ideals of research achievement, and dedicated interest in entomology. To her father, step-mother, and sister we extend warmest sympathy. AsHLEy B, Gurney and Luetta M. WALKLEY Entomological Publications of Sophy I. Parfin 1947. [Untitled note on Mecoptera from Oregon] Note given at May 1, meeting. Proc. Ent. Soc. Wash. 49: 258. 1952. The Megaloptera and Neuroptera of Minnesota. Amer. Midl. Naturalist 47: 421-434. 1955. Additional records for Brachypanorpa carolinensis (Banks) (Mecoptera, Panorpidae). Proc. Ent. Soc. Wash. 57: 204-205. 1956. Taxonomic notes on Kimminsia (Neuroptera; Hemerobiidae). Proc. Ent. Soc. Wash. 58: 203-209, 15. figs. 1956. The spongilla-flies, with special reference to those of the Western Hemi- sphere (Sisyridae, Neuroptera). Proc. U. S. Nat. Mus. 105: 421-529, 24 figs., 3 pls. (By Sophy I. Parfin and Ashley B. Gumey) 1958. Notes on the bionomics of the Mantispidae (Neuroptera: Planipennia). Ent. News 69: 203-207. 1959. Chapter 37, Neuroptera, pp. 973-980, 4 figs., in Freshwater Biology, Edited by W. T. Edmondson, 1248 pp. (By Ashley B. Gumey and Sophy I. Parfin ) PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 193 BOOK REVIEWS Systems Analysis in Ecology, edited by Kenneth E. F. Watt. Academic Press, New York, N.Y. xiii + 276 pp., illus. January, 1967. $11.50. Designed to survey problems and techniques of systems analysis in ecology, this is the first book to present new electronic and computer procedures being used in ecology and resource management. Professor Watt has brought together a fascinating array of papers that discuss the motives for using systems analysis in ecology, the electronic principles used in data acquisition systems, instrumented universes for studying animal behavior, the design of systems experiments, systems simulation, statistical aspects of data reduction by the use of computers, and a host of additional items applicable to computer usage in this complicated field of biology. The book will appeal especially to those ecologists whose respect for the modern-day computer reaches beyond its ability simply to reduce data. As aptly expressed by one of the contributors, Crawford S. Holling, “. . . if biology has told us anything, it is that complex systems are not simply the sum of their parts. There is an emergent principal when fragments act and interact in a whole system. The speed and large memory of modern digital computers for the first time allows the ecologist, in principle, to incorporate all the relevant actions and interactions of the fragments of complex ecological systems in an integrated manner.” One of the greatest values of this book might well be to show ecologists how they can take advantage of a very powerful tool in in- vestigations now under way or in the planning stage —RicHArp H. Foore, Entomology Research Division, ARS, U.S. Department of Agriculture, Washing- ton, D.C. 20560. Insektenstimmen. By S. L. Tuxen. 156 pp., 89 figs. 1967. Publisher: Springer- Verlag (Berlin, Heidelberg, New York). Price: DM 12.80 ($3.20). Man’s interest in sounds produced by insects is very old, but in recent years, especially since the appearance of a 1949 book “The songs of insects” by George Pierce, a Harvard physics professor, refined modern equipment has permitted the “songs” to be recorded, analyzed, and described to others in a way that was not available to Scudder, Allard, and other early observers. Consequently, there have been many advances which have placed the study of insect sounds on a thorough scientific basis, at the same time demonstrating how a knowledge of this aspect of behavior may contribute to systematics, ecology, and applied phases of entomology. The present book, the title of which would be “Insect sounds” in English, first appeared in Danish in 1964 and now is issued as the 88th in a series of small, handbook-size volumes on natural history published in German by Springer-Verlag. It is attractive, well printed on good paper, and sturdily bound. Dr. S. L. Tuxen is an entomologist of broad interests, best known professionally as a specialist of Protura, for editing a 1956 volume on insect genitalia, and for heading the Department of Entomology in the Zoological Museum in Copenhagen. In private life Dr. Tuxen is a gifted musician, an enthusiastic raconteur of observations in many foreign lands, and a charming host. The book is divided into sections or small chapters, the titles of which trans- 194 PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 late as follows, with a few explanatory words by this reviewer in parentheses: Do insects sing? A musician and his instrument; Song of the cricket, a little sound-theory; Song and mating of the cricket; Ranges and territories of crickets; Song of the tree grasshoppers (Tettigoniidae); A field grasshopper woos; A brief summary; The ear of the grasshopper; A little acoustics and a little electrophysi- ology; Direction hearing; Singing and hearing; The “near the knee” organ (tympanal organs); Hearing hairs and flight reaction; A little singing brain; Unorthodox grasshopper voices; Other stridulating insects; Underwater singers; Ghost tappers (death-watch, etc.); The happy cicadas; The deathshead moth and the sounds in the beehive; What does a male midge hear?; Moth and bat; Why do insects sing? The origin of songs, some theories. The 1961 book, “Insect sounds,” by Peter T. Haskell, is a somewhat more formal, near counterpart in English. Though Dr. Tuxen contributes some ob- servations of his own, the book is mainly a summary of work by other entomolo- gists, and the experiments and illustrations of Richard D. Alexander, Busnel, Haskell, Regen, Roeder, Treat, T. J. Walker and others have been drawn upon freely. The bibliography is quite short, the bulk of the literature having been assembled in 1960 by Hubert and Mabel Frings. The illustrations, of which about 10 are original, do much to make the book attractive and to clarify the text. For instance, fig. 27 shows Regen’s clever method of placing crickets in cages suspended from small free-flying balloons in order to demonstrate that hearing occurred and was not due to any vibrations through a solid substrate. For those persons with an interest in insects or general natural history, and who read German, this seems like an extremely pleasant and informative book. For serious students it provides a very readable, relatively non-technical summary of the highlights of recent contributions on insect sounds and their meaning. For graduate students studying German, the book might be useful as a practice reader. AsHLEY B. GurNry, Entomology Research Division, ARS, U. S. Depart- ment of Agriculture, Washington, D.C. 20560. BOOK NOTICE Clarification of the status of the type specimens of Diabroticites (Coleop- tera, Chrysomelidae, Galerucinae). By Ray F. Smith and John F. Lawrence. Univ. Cal. Pubs. Ento., vol. 45: 1-168, 4 pls., 1967. This work is a guide to the location of the type specimens of the over 900 valid species of this group and provides a nomenclatural base for future work. The Diabroticites is about equal to the content of the genus Diabrotica prior to 1947. PROC. ENT. SOC. WASH., VOL. 69, NO. 2, JUNE, 1967 195 SOCIETY MEETINGS 750th Regular Meeting—March 2, 1967 The 750th regular meeting of the Society was called to order by the president, Mr. Louis G. Davis, on March 2, 1967 at 8:00 p.m. in room 43, U. S. National Museum. Thirty-one members and twenty-seven guests were in attendance. Minutes of the previous meeting were approved as read. The names of Richard H. Whitsel, Cheng Shan Lin, Ralph E. Webb, and Gerald I. Stage were read for the first time as candidates for membership in the Society. J. Phillip Keathley and Gerald G. Elgert were received into the Society. R. H. Foote announced that the annual joint meeting with the Insecticide So- ciety of Washington will be held on June 1 at the University of Maryland. L. G. Davis reported that Ralph W. Sherman, a member of our Society, was pre- sented a distinguished service award by Kansas State University on January 30, 1967. K. V. Krombein exhibited a portion of a collection of over 89,000 insects from Ireland (mostly Hymenoptera) which had been donated to the U. S. National Museum by Mr. A. W. Stelfox. R. H. Foote reviewed and exhibited a new book, “Trap-nesting Wasps and Bees: Life Histories, Nests, and Associates”, by K. V. Krombein and published by the Smithsonian Press. G. G. Elgert ex- hibited a photograph which he had taken of an earwig, Anisolabis maritima (Géné), guarding its eggs. L. G. Davis displayed several new pamphlets pub- lished by the Plant Pest Control Division, USDA dealing with insects which do not occur in the United States. The speaker for the evening, Dr. Botha de Meillon, presented a most interesting and informative review of the entomology of malaria in Africa south of the Sahara. A lively discussion followed. Following the introduction of visitors, the meeting was adjourned by President Davis at 9:30 p.m. Ratpn A. Bram, Recording Secretary. 751st Regular Meeting—April 6, 1967 The 751st regular meeting of the Society was called to order by the President, Mr. Louis G. Davis, on April 6, 1967 at 8:00 p.m. in room 43, U. S. National Museum. Thirty-eight members and twenty visitors were in attendance. Minutes of the previous meeting were approved as read. Richard H. Whitsel, Chen Shan Lin, Ralph E. Webb, and Gerald I. Stage were received into the Society. Mrs. Helen Sollers-Riedel discussed the activities of the Teen International Entomology Group, an organization sponsored by Cornell University. R. I. Sailer reviewed and exhibited a recent book, “A Monograph of the Emesinae (Redu- viidae, Hemiptera)”, by P. W. Wygodzinsky and published as volume 133 of the Bulletin of the American Museum of Natural History, 1966. The first speaker of the evening, Dr. Ashley B. Gurney, related his experiences collecting insects in Ethiopia, and discussed the activities of some of the entomol- ogists working there. Dr. Oliver S. Flint reviewed his recent collecting expedition along the Antarctic. Impressive slide series accompanied each lecture. Following the introduction of visitors, the meeting was adjourned by President Davis at 9:50 p.m. RALpu A. Bram, Recording Secretary. For more than half a century, Cyanamid has consistently led the chemical industry in developing new products and application techniques that have helped immeasurably to bolster our na- tional farm economy. ™@ Topping the list of Cyanamid ‘firsts’ is Malathion LV* Concentrate, introduced commercially last year for the control of boll weevils on cotton after two years of use on more than 1 million treatment acres in cooperation with the U.S.D.A.’s Agricultural Research Service. m Malathion LV Con- centrate is also being used extensively to combat grasshoppers, cereal leaf beetles, corn rootworm beetles, mosquitoes, blue- berry maggots, flies and beet leafhoppers. m@ Watch for progress reports of new tests conducted against many other pests with both aerial and ground equipment. Data being processed daily show clearly that Malathion LV Concentrate is fast making all other methods of insect control obsolete! m Before using any pesticide, stop and read the label. *Trademark SERVES THE MAN WHO MAKES AMERICAN CYANAMID COMPANY SINE eS OF Oa PRINCETON, NEW JERSEY PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. Manuscripts for publication, proof and other editorial matters should be addressed to the Editor (for address, see inside front cover of this issue). Typing—All manuscripts must be typed on bond paper with double-spacing and ample margins. Carbon copies or copies on paper larger than 8% x 11 inches are not acceptable. Do not use all capitals for any purpose. Underscore only where italics are intended in the body of the text, not in headings. Number all pages consecutively. References to footnotes in the text should be numbered consecutively and typed on a separate sheet. First page—The page preceding the text of the manuscript should include (1) the complete title, (2) the order and family in parentheses, (3) the author’s name or names, (4) the institution with city, state and zip code or the author’s home city, state and zip code if not affiliated, (5) in the upper left hand comer, the complete name and address to which proof is to be sent. Names and descriptions of organisms—The first mention of a plant or animal should include the full scientific name with the author of a zoological name not abbreviated. Descriptions of taxa should be in telegraphic style. References—Citations in the text of papers longer than one printed page should be by author and date and should refer to a list of concluding REFERENCEs listed alphabetically in the following format: Muesebeck, C. F. W. 1963. Host relationships of the Euphorini (Hyme- noptera: Braconidae). Proc. Ent. Soc. Wash. 65(4): 306. and L. M. Walkley. 1951. in Muesebeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2: 90-184. In shorter articles, references to literature should be included in parentheses in the text. Tables—Tables are expensive and should be kept to an absolute minimum. In most cases, material in tables can be incorporated into the text. When tables are necessary, each table should be prepared as a line drawing or typed on a separate page with heading at top and footnotes below. Number footnotes consecutively for each table. Use only horizontal rules. Illustrations—No extra charge is made for line drawings or halftones. Authors must plan their illustrations for reduction to the dimensions of the printed page and the individual figures must be mounted on suitable board. Proportions of full-page illustrations should closely approximate 44g < 6” (26 X 36 picas); this usually allows explanatory matter to appear on the same page. On the back of each illustration should be stated (1) the title of the paper, (2) the author’s complete name and address, and (3) the number of the illustration such as BNO (oto) ete. Figure legends—Legends should be typewritten double-spaced on separate pages headed EXPLANATION OF FicurREs and placed following REFERENCEs. Do not attach legends to illustrations. Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) PIPKIN, S. B.—A new member of a sibling set belonging to the Drosophila tripunctata group (Diptera: Drosophilidae ) ROBINSON, H.—New species of Dolichopodidae from the United States and Mexico (Diptera) RUSSELL, L. M.—A list of the species of Macrosiphoniella Del Guercio described from North America (Homoptera: Aphididae ) SMILEY, R. L.—Further studies on the Tarsonemidae (Acarina) SMITH, D. R.—New synonymy in Pleroneura Konow (Hymenoptera: Xyeli- alchee NP yh! ewe ee hs occa he ee Se A STEYSKAL, G. C.—Palaeosepsis species described by Melander and Spuler (Ditpteras Sepsidae ) 2. ae ee TOWNES, H.—A new Ateleute from the United States (Hymenoptera: Ich- MeumMAielae ) Vy Se ad es A a ee WALKER, T. J. and A. B. GURNEY—The metanotal gland as a taxonomic character in Oecanthus of the United States (Orthoptera: Gryllidae) —. BOOK REVIEWS 23005000105 J 0 ee ee Ee eee BOOB: (VG TICE) cc ho ee ey eee OBITUARY—SOPHY I. PARFIN, 1918-1966 _ SOCIBEY: MEETINGS tt. 5 8 Lit 114 184 127 182 180 Y* fO6k3 LDraAse Vol. 69 ee SEPTEMBER 1967 No. 3 PROCEEDINGS of the ENTOMOLOGICAL SOCIETY of WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 NOV 2 196 BLAKE, D. H.—Some new and old species of Colaspis in the West Indies _.. 225 CARTWRIGHT, O. L.—A new Thyce from Georgia (Coleoptera: Scarabaei- CONTENTS (Continued on back cover) ere ene. oe Fe i Sh ee ene 2 EE ee ee CLARK, G. M.—A new nasal mite from the robin (Turdus migratorius ) eoearnae ‘Speleopmathinae) . 2 294 DELONG, D. M.—Studies of the Mexican Deltocephalinae: a new genus, Conversana, and three new species (Homoptera: Cicadellidae) — : DODGE, H. R.—Two new metallic species of Helina from Chile (Diptera: INETSCIC HE) eee eveet ee at 2 Te ee ee ee ce BA) Te Be ee : EVANS, H. E.—New generic records of Bethylidae from South America MPRRRTEMArS [AEE Pot Ws ae ee oe Os Free ee hs og ee en ee 269 GURNEY, A. B.—A substitute name for the genus Nicuesa in cockroaches iMmareevontera:: Blattariay Blattellidae) 0... : memteeie.) 1. A.—Sex ratios of Platypus. KRAMER, J. P.—A key to the species of Lystridea Baker with description of a new species from California (Homoptera: Cicadellidae: Errhomenellini) 25 LEONARD, M. D.—A list of the aphids of Cyprus (Homoptera: Aphididae ) MARSH, P. M.—The Nearctic Doryctinae, IV. Notes on the genus Dendro- soter Wesmael and description of one new species (Hymenoptera: Bra- TERE NUS AT Se Sg Sa DR 9 2 ee ARN Nace tes ee ENTOMOLOGICAL SOCIETY OF WASHINGTON OrGANIZED Marcu 12, 1884 OFFICERS FOR 1967 Mr. Louis G. Davis, President Dr. Jon L. HEerninc, Editor ARS, USDA, Rm. 466 c/o Department of Entomology Federal Center Bldg. U.S. National Museum Hyattsville, Maryland 20782 Washington, D.C. 20560 Dr. Ricuarp H. Foore, President-Elect Mr. Ropert SMILEY, Custodian c/o Department of Entomology Entomology Research Division, USDA U.S. National Museum Washington, D.C. 20250 Washington, D.C. 20560 Dr. Rautpu A, Bram, Recording Secretary Mr. Victor ADLER, Program Committee Chairme Department of Entomology Entomology Research Division, ARS, USDA U.S. National Museum ; ARC, Beltsville, Maryland 20750 Washington, D.C, 20560 Mr. Dav R. Smirn, Corresponding Secretary Capt. Wirtu1AM B. Hutt, Membership Committe c/o Department of Entomology Chairman U.S. National Museum 746 Beall Ave. Washington, D.C. 20560 Rockville, Maryland 20850 Dr. ArruuRr K. Burpitt, Jr., Treasurer Dr. H. H. SHEPARD, Delegate to the Washingte ARS, USDA, Rm. 125 Academy of Sciences North Bldg. PPA-ASCS Beltsville, Maryland 20705 U.S. Department of Agriculture Washington, D.C. 20250 Honorary President Dr. T. E. Snyper, U.S. National Museum Honorary Members Mr. C. F. W. MursEsreck, U.S. National Museum Dr. F. W. Poos Dr. E. N. Cory MEETINGS.—Regular meetings of the Society are held in Room 43 of the U.S. National Museum on the | Thursday of each month from October to June, inclusive, at 8 P.M. Minutes of meetings are published regul in the Proceedings. MEMBERSHIP.—Members shall be persons who have demonstrated interest in the science of entomolk Annual dues for members are $5.00; initiation fee is $1.00 (U.S. currency). PROCEEDINGS.—Published quarterly beginning with March by the Society at Washington, D.C. Member good standing are entitled to the Proceedings free of charge. Nonmember subscriptions are $8.00 per y both domestic and foreign (U.S. currency), payable in advance. All remittances should be made payabl The Entomological Society of Washington. The Society does not exchange its publications for those of other societies. STATEMENT OF OWNERSHIP Title of Publication: Proceedings of the Entomological Society of Washington. Frequency of Issue: Quarterly (March, June, September, December). Location of Office of Publication, Business Office of Publisher and Owner: The Entomological Society Washington, c/o Department of Entomology, U.S. National Museum, Washington, D.C. 20560. | Editor: Dr. Jon L. Herring, same address as above. Managing Editor and Known Bondholders or other Security Holders: none. This issue mailed October 13, 1967 1 Second Class Postage Paid at Lawrence, Kansas, U. S. A. | ALLEN PRESS, INC. eee LAWRENCE, KANSAS US. & PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Vol. 69 SEPTEMBER 1967 No. 3 A SYNOPTIC CATALOG OF THE MOSQUITOES OF THE WORLD, SUPPLEMENT III (DreTERA: CuLicipAE )* ALAN STONE, Entomology Research Division, ARS, U. S. Department of Agriculture, Washington, D. C. 20560 This paper is the third supplement to the Synoptic Catalog of the Mos- quitoes of the World by Stone, Knight, and Starcke (1959), the second supplement having appeared in Vol. 65 (1963), No. 2, pp. 117-140 of the Proceedings. The introductory remarks as to format and coverage in Supple- ment II apply to this supplement also. One new name of the genus-group and 104 new names of the species group are listed. I am _ particularly indebted to T. H. G. Aitken, M. Delfinado, B. de Meillon, J. C. Hitchcock, je. GC. lL. Hogue, Z. Liepa, J. P. Macnamara, Bek: Mattingly, J. A. Reid, J. E. Scanlon, H. Sollers-Riedel, and T. J. Zavortink for pertinent information. Introduction Page 1. With the additions in this supplement and taxonomic changes, the number of valid genera and subgenera stands at 119, the number of valid species becomes 2,680, and the total number of names of the species-group 4,319. Systematic Arrangement 6. The genus Mansonia (see p. 101) has been restricted to the two sub- genera Mansonia and Mansonioides, and Coquillettidia raised to generic rank with Rhynchotaenia as a subgenus of it. Kompia has been restored to subgeneric status in the genus Aedes. Catalog of the Family Culicidae 9. Taxonomic Anatomy.—1963b, Reid, 32-38 (pupal trumpet). General Taxonomy.—1966, Dodge, 337-393 (1st stage larvae). Australasian Region.—1966, Steffan, 179-237 (checklist, Papuan subregion ). Nearctic Region.—1963, Dodge, 796-813 (larvae; eastern North America). Neotropical Region.—1961, Fauran, 2-60 (catalog, French Guiana) ; 1962, Garcia and Ronderos, 105-212 (Anophelini, Argen- tina). Oriental Region.—1966, Delfinado, 1-252 (Culicini, Philippines). ‘Reprints are for sale by the Thomas Say Foundation, Entomological Society of America, 4603 Calvert Road, College Park, Maryland 20740. 198 10. ee 13. 14. 15. 16. Lif PROC, ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Genus BIRONELLA Theobald Subgenus BIRONELLA Theobald bironelli (Christophers ).—Transfer to p. 11 as synonym of gracilis Theobald. gracilis Theobald.—Transfer to p. 11 as valid name. confusa Bonne- Wepster.—Slooff, 1963, 138 (4*, 9, P*, L*). gracilis (Theobald).—I.C.Z.N., 1965, 164 (Validation under Plenary Powers ). bironelli (Christophers), ibid. (Placed on list of rejected and invalid specific names ). Genus ANOPHELES Meigen Important references: 1962, Service, 120-158 (W. Africa, keys) ; 1963, Quy, 26-34 (keys, Vietnam); 1963, various authors in Russell, West, Manwell, and MacDonald, 641-701 (keys). Subgenus ANOPHELES Meigen Important reference: 1965, Reid, 106-125 (key, aitkenii group). acaci Baisas.—Philippines, Borneo. Reid, 1965, 115 (L*; syn.). borneensis McArthur. aitkenii James. ee Reid, 1965, 107 (¢*; 9 Pie syn pallida dicen fragilis Theobald.—Transfer to p. 18 as valid species. ssp. treacherii Leicester—Transfer to p. 18 as synonym of fragilis Theobald. ssp. bengalensis Puri—Change volume number from 18 to 17; trans- fer to p. 16 as valid species. pallida Ludlow.—Transfer to p. 13 as synonym of aitkenii s. str. var. borneensis MacArthur.—Transfer to p. 13 as synonym of acaci Baisas. ssp. stantoni Puri—Transfer to p. 18 as synonym of fragilis Theobald; change BM to NE. var. djajasanensis (Brug).—Change to djajasanensis (Brug). See Reid reference under ssp. interruptus (syn.). ssp. interruptus Puri—Malaya. Reid, 1963a, 111 (taxonomy); change ssp. to var. annulipalpis Lynch Arribalzaga—Garcia and Casal, 1964 (1965). Rev. Soc. ent. Argentina 27: 5 (P*). asiaticus (Leicester).—Delete parentheses; insert as original citation “1903, J. trop. Med. 6: 291 (A); delete Lophoscelomyia. bengalensis Puri—Pakistan (Bengal), India (Assam), Burma, Thai- land, Indo China, Philippines, S$. China, Taiwan. Reid, 196d; 113 (79> P*, e*tosprstatusye brevipalpus Roper.—Change “Moluccas” to “Bangka Island.” petragnani Del Vecchio.—Transfer to p. 25 as valid species; insert “; as var.” after “E*,” missiroli Del Vecchio.—Insert “; as var.” after “E*”. collessi Reid—Malaya, Borneo. 1963a. Ann. trop. Med., Parasit.. 57: 103 (6%, 22 2p ieaeeriea: Type-loc: Ulu Gombak near Kuala Lumpur, Selangor, Malaya (BM). var. ziemanni Griinberg.—Insert “; as sp.” after “2. ”» 18. 19. 28. 29. 30. PROC. ENT. SOC. WASH., VOL. 69, No. 3, SEPTEMBER, 1967 199 fragilis Theobald—Malaya, Indonesia e. to Celebes and Borneo, Philippines. Reid, 1965, 109 (4*, 2, P*, L*; syn.). treacherii Leicester. stantoni Puri. var. formosus Ludlow.—Insert “; as sp.” after “2.” var. simlensis (James ).—Insert “; as sp.” after “Patagiamyjia.” var. oedjalikalahensis Nainggolan.—Change to oedjalikalah; insert in Bonne-Wepster and Swellengrebel after “L*,” “; as oedjalikalahensis.” var. udjalikalah Waktoedi.—Delete, merely a variant spelling of above. insulaeforum (Swellengrebel and Swellengrebel de Graaf).—Burma, Thailand, Taiwan. Delete “P Japan, ? New Guinea, ? Aus- tralia. 2 Reid. 1965.19 (Coss 2 Psa) lesteri Baisas and Hu.—Japan, China. letifer Sandosham.—Thailand. ssp. japonicus Yamada.—Insert “; as sp.” after “2 *” in first line. noniae Reid.— Malaya. 1963a. Ann. trop. Med. Parasit. 57: 108 (4, 2*, P*, L*). Type- loc: Ulu Langat, Selangor, Malaya (BM). pursati Laveran.—Transfer to p. 27 as valid sp. palmatus Rodenwaldt.—Delete “?” before Malaya. Reid, 1965, 117 (ie). petragnani Del Vecchio.—Mediterranean Region. Coluzzi, 1962, 1025 (resurrected from synonymy). ssp. francisecanus McCracken.—Change “SU” to “LACM”. willardi Vargas.—Delete “Ciudad Juarez.” stonei Vargas.—Insert “and USNM” after “ISET.” annulimanus Van der Wulp.—Change “North America” to “Wiscon- sin. pursati Laveran.—Indochina, Thailand, Malaya. Reid, 1963a, 98 (6*, 2*, P*, L*; resurrected from synonymy ). roperi Reid.—Assam. sintonoides Ho.—Malaya, Thailand. stricklandi Reid.—Malaya. 1965. Ann. trop. Med. Parasit. 59: 121 (¢*, 2, P*, L*). Type- loc: Johore, Malaya (BM). tasmaniensis Dobrotworsky.—Tasmania, Australia. See" Proc inn. Soc.N/S:W.. 91: 130963 P92 P*, LX, 1"). Type-loc: Birralee, Tasmania (CSIR). tibiamaculatus (Neiva).—Argentina. tigertti Scanlon and Peyton.—Thailand. 1967. Proc. ent. Soc. Wash. 69: 19 (¢*, 9, P*, L*). Type-loc: Ban Bu Phram, Prachinburi Province, Thailand (USNM). whartoni Reid.— Malaya. 1963a. Ann. trop. Med. Parasit. 57: 106 (é, 2*, P, L, E). Type- loc: Singgora, Maran, Pahang, Malaya (BM). Subgenus NYSSORHYNCHUS Blanchard albitarsis Lynch Arribalzaga.—Garcia and Casal, 1964 (1965). Rev. Soc. ent. Argentina 27: 6 (P*). 200 30. 36. 38. 39. 40. 41. 42. 43. 44. 45. 46. AT. 48. 49. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Subgenus KERTESZIA Theobald bambusicolus Komp.—Change to bambusicola. hylephilus Dyar and Knab.—Delete “Manoa, Orinoco River, Venezu- ela; Guayaquil, Ecuador and.” Belkin, Schick and Heine- mann, 1965, 44 (lectotype). Subgenus CELLIA Theobald apoci Marsh.—Arab, 1963, 73 (P*). brohieri Edwards.—Transfer, with synonym, to p. 44 as variety of hancocki Edwards. cavernicolus Abonnenc.—Change to cavernicola. cydippis De Meillon.—Lips, 1962, 108 (to sp. status). punjabensis James.—Insert “; as var.” after “Myzomyia.” melanocosta Newstead and Carter.—Insert “; as var.” after Pyreto- phorus.” domicolus Edwards.—Change to domicola; transfer to p. 46 as variety of longipalpis (Theobald). wardi Leeson and Theodor.—Insert “; as var.” after “L*.” errabundus (Swellengrebel).—Transfer to p. 32 as synonym of darlingi Root. Reid and Bonne-Wepster, 1966, 190 (sup- pression proposed ). subumbrosa Theobald.—Change to swbumbrosus, delete Myzomyia, and change reference to Mem. Lpool Sch. trop. Med. 10 (App. ): 4. gambiae Giles —Coluzzi, 1964, 197-232 (taxonomy). merus Donitz.—Transfer to p. 48 as valid species. litoralis Halerow.—Transfer to p. 48 as synonym of merus Donitz. var. melas (Theobald ).—Delete Pyretophorus and parentheses around Theobald. Change reference to Mem. Lpool Sch. trop. Med. 10(App.): 2. Place Bathurst in [ ]. Transfer to p. 48 as valid species. grenieri Grjebine.-—Madagascar. 1964. Bull. Soc. Path. exot. 57: 38 (L*). Type-loc: Ampandro- atraka, Anosibé Distr., Madagascar (IRSM). hancocki Edwards. var. brohieri Edwards——Adam, Hamon, Rickenbach and Lips, 1956, 756 (to var. status). indefinitus (Ludlow).—Malaya, Philippines, Indochina, China, Tai- wan, Mariana Islands. Reid, 1966, 327 (to sp. status; syn.). malayensis Hacker. jamesii Theobald.—Malaya. longipalpis (Theobald). var. domicola Edwards.—Adam, Bruce-Chwatt, and Hamon, 1956, 107 (to var. status). ssp. willmori (James ).—Change to willmorei, here emended. var. pitchfordi (Giles).—Insert “; as ssp.” after “Pyretophorus.” melas (Theobald ).—Coluzzi, 1964, 275 (to sp. status). merus Donitz.—Coluzzi, 1964, 225 (to sp. status; syn.). litoralis Halcrow. tangensis Kuhlow, 1962, Ztschr. f. Tropenmed. u. Parasit. 13: 448 (6, 2). Type-loc: Tanga, Tanganyika (BNI). ssp. flavirostris (Ludlow).—Delete “f.” after “Myzomyia” and insert asisp: 52. Do. 56. 57. 58. 59. 65. 68. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 201 ssp. rupicolus Lewis.—Change to rupicola. var. indefinitus (Ludlow) .—Transfer to p. 45 as valid species with var. malayensis Hacker as synonym. var. eydippis De Meillon.—Transfer with synonym to p. 41 as valid species. subpictus Grassi. var. vadakadiensis Doraisamy.—India. 1963. Bull. Ent. (Madras) 4: 42 (L*). Type-loc: Near Vadakadu, Rameswaram Island, Madras, India (MSI). swahilicus Gillies —Kenya, Tanzania. 1964. (Proc« BR: ent Second. 4(B)) So: 61, (6 %..0%, BP L*),. Type-loc: Goshi, Molindi Distr., Kenya (BM). var. orientalis Swellengrebel and Swellengrebel de Graaf. “Neomyzomyia’ before “punctulata.” theobaldi Giles —Change original reference to “1901. Ent. mon. Mag. ate 198." varuna lyengar.—Change IM to LU. Insert Genus TOXORHYNCHITES Theobald Important reference: 1962, Lima, Guitton, and Ferreira, 225-252 (Ankylorhynchus and Lynchiella, revision, key). Subgenus ANKYLORHYNCHUS Lutz eatharinensis (Lima, Guitton, and Ferreira ).—Brazil. 1962. Mem. Inst. Osw. Cruz 60: 230 (4*, 2, P*, L*; Ankylo- rhynchus). Type-loc: Not given, but probably Santa Catarina, Brazil (IOC). Subgenus LYNCHIELLA Lahille bambusicolus (Lutz and Neiva).—Change to bambusicola. mara Anduze.—Lima, Guitton, and Ferreira 1962, 243 (syn. sug- gested). guianensis Bonne-Wepster and Bonne.—Insert “; as ssp.” after “Mega- rhinus.” ssp. septentrionalis (Dyar and Knab).—Dodge, 1964, 46-53 (L*). mara Anduze.—Transfer to p. 59 as synonym of bambusicola. Subgenus TOXORHYNCHITES Theobald aurifluus (Edwards ).—Lien, 1965, 9 (4*, 9*, P*, L*). ssp. conradti Griinberg.—Corbet, 1963, 9 (L*, P*, E). kaimosi (Someren ).—Corbet, 1963, 11 (P*). manicatus (Edwards) .—Lien, 1965, 2 (¢*, 2*, P*, L*). Genus TRIPTEROIDES Giles Important reference: 1963b, Peters, 89-100 (New Guinea, key P). Subgenus TRIPTEROIDES Giles elegans Brug.—Peters, 1963b, 89 (P*, L*). lorengaui Peters—New Guinea. 1963. Proc. Ki ‘ent? Sores ond.” (B)" 32: 67 «¢6.*4 2, P*,. L*). Type-loc: Lorengau River, Manus Island, New Guinea (CSIR). PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 nissanensis Lee.—Peters, 1963a, 70 (P*). novohanoverae Peters.—New Guinea. 1963. Proe.: R. ent. Soe: Taonds(B)). 32s 63: (4%) 92 Peaaieee Type-loc: Noipuas, New Hanover, New Guinea (CSIR). szechwanensis Hsu.—China. 1964. Acta ent. Sinica 13: 298 (6*, L*). Type-loc: Chengtu, Szechwan, China (SMC). Subgenus RACHIONOTOMYIA Theobald marksae Dobrotworsky.—Australia. 1965. Mosq. of Victoria :58 (¢*, 9, L*). Type-loc: Cabbage Tree Creek, Victoria, Australia (NMM). perplexus Peters —New Guinea. 1963. Proc. R. ent. Soc. Lond. (B) 32: 91 (9, P*). Type-lce: Maprik, Sepik District, New Guinea (CSIR). Genus TRICHOPROSOPON Theobald Subgenus TRICHOPROSOPON Theobald 73. var. mogilasium (Dyar and Knab).—Insert “; as sp.” after “Joblotia.” 74-77. Running Heads. Change “TRICHOPOSOPON” to “TRICHO- 74. 1: Ue 79. 80. 84. 89. 90. PROSOPON.” Subgenus SHANNONIANA Lane and Cerqueira fluviatilis (Theobald ).—Change to fluviatile. Subgenus RUNCHOMYIA Theobald cotopaxensis Levi Castillo—Change to ecotopaxense. edwardsianum Lane and Cerqueira.—Cerqueira, 1961, 460 (¢*, @, P*, L*), lunatum (Theobald ).—Delete “Argentina.” paranensis Bréthes.—Argentina. Garcia and Casal, 1965, 14 (4%, resurrected from syn.); place after pallidiventer. Genus WYEOMYIA Theobald Subgenus WYEOMYIA Theobald culebrae Dyar.—Change “1926” to “1923.” labesba Howard, Dyar and Knab.—Change “Ancon” to “Tabernilla.” Subgenus DENDROMYIA Theobald belkini Casal and Garcia.—Argentina. 1966. Physis 26: 155 (4*, 9*, L*, P*). Type-loc: Igu, Misiones, Argentina (INM). Genus PHONIOMYIA Theobald lassalli (Bonne-Wepster and Bonne).—Change “LU” to “USNM.” Lectotype here selected bears labels: Trinidad, W.I./F.W. Urich Collector/ B12-6/ See slide No. 364/ @ of trinidad- ensis D. K. ident. 1906; terminalia on slide 364. Genus LIMATUS Theobald Simondella.—Change “Lavaran” to “Laveran.” PROC, ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 203 Genus SABETHES Robineau-Desvoidy Subgenus SABETHES Robineau-Desvoidy 91. kappleri Bonne.—Change “LU” to “NE.” Subgenus SABETHOIDES Theobald 93. rangeli Surcouf and Gonzales-Rincones.—Change “Sabethoides.” < ‘Sabethoides” to Genus TOPOMYIA Leicester 95. aureoventer Theobald.—Change “BM” to “IM”; only slide of wing and leg in BM. Genus FICALBIA Theobald Subgenus RAVENALITES Doucet 100. deguzmanae Mattingly.—Delfinado, 1966, 20 (é*, 2, P*, L*). Genus MANSONIA Blanchard 101. Genus MANSONIA Blanchard.—Transfer with following line to p. 104 preceding Subgenus MANSONIOIDES. Genus COQUILLETTIDIA Dyar Insert Genus COQUILLETTIDIA before Subgenus COQUIL- LETTIDIA. Important references: 1962, Wharton, 1-114 (Malaya); 1962 (1963), Ronderos and Bachmann, 48-51 (to generic status). Subgenus COQUILLETTIDIA Dyar crassipes (Van der Wulp).—Wharton, 1962, 6 (é6*, 2, P*, L*, E). 102. giblini (Taylor).—Delete “Australia, Philippines, Formosa, Malaya, Thailand, Burma.” nigrosignatus Edwards.—Transfer to p. 103 as valid sp. hodgkini (Wharton ).—Malaya, Indonesia. 1962. Inst. med. Res. Malaya, Bull. 11: 12 (¢*, 2*; Mansonia). Type-loc: Lamir (Pekan), Pahang, Malaya (BM). 103. nigrosignata (Edwards).—Malaya, Indonesia, Philippines. Wharton, 1962, 7 (6*, 9*, L*, E; resurrected from synonymy ). testaceus Van der Wulp.—Delete “[?] United States.” 104. samoaensis Stone.—Samoa. 1966. Proc. ent. Soc. Wash. 68: 331 (¢6*, 2). Type-loc: Taba- logi Ridge, Tau Island, Manua Group, Eastern Samoa (USNM). Subgenus RHYNCHOTAENIA Bréethes Insert Subgenus RHYNCHOTAENIA, its synonym, and all its species to follow species of Subgenus COQUILLET- TIDIA. Genus MANSONIA Blanchard Insert Genus MANSONIA Blanchard. Important references: 1962, Wharton, 1-114 (Malaya); 1962 (1963), Ronderos and Bachmann, 46-48. 204 105. 106. 107. 108. 109. 110. at, W112, 113. PROC, ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Subgenus MANSONIOIDES Theobald annulata Leicester.—Celebes. Wharton, 1962, 15 (¢*, 2*, P*, L*, FE). annulifera (Theobald).—Delfinado, 1966, 23 (6*, 9*). bonneae Edwards.—Wharton, 1962, 17 (3%, oF ,P*, | ig! 3) dives (Schiner ).—Wharton, 1962, 20 Cares OF, _P*, L*, E). papuensis (Taylor) _—Change “901” to “200” and “Melanoconion” to “Taeniorhynchus.” Subgenus MANSONIA Blanchard Important reference: 1963, Ronderos and Bachmann, 57-65 (keys). pseudotitillans (Theobald).—Ronderos and Bachmann, 1963, 61 (G4. oO". Baaice titillans (Walker) _—Ronderos and Bachmann, 1963, 61 (¢*, P*, L*, ihe Transfer Subgenus RHYNCHOTAENIA, its synonyms, and all its species to p. 104 as subgenus of COQUILLETTIDIA. Subgenus Uncertain aurata Dobrotworsky.—Australia. 1962. Proc. Linn. Soc. N.S.W. 87: 295 (2*). Type-loc: Cabbage Tree Creek, Victoria, Australia (NMM). variegata Dobrotworsky.—Australia. 1962. Proc. Linn. Soc. N.S.W. 87: 293 (6*, 2*). Type-loe: Cann River, Victoria, Australia (NMM). Genus URANOTAENIA Lynch Arribalzaga albescens Taylor.—Peters, 1964, 27 (A, P*, L). Delete Penn 1949b reference. albosternopleura Peters.—New Guinea. 1963. Proc. R. ent. Soc. Lond. (B) 32: 203 (é, 2*). Type-loc: Maprik, Sepik Distr., New Guinea (CSIR). amiensis Peters —New Guinea, Papua. 1963. Proc. R. ent. Soc. Lond. (B) 32: 207 (4*, 2*). Type-loc: Ami, 10 miles nw. of Maprik, Sepik Distr., New Guinea (CSIR). annandalei Barraud.—Philippines. Delfinado, 1966, 39 (4*, 2, P*, 1 boi arguellesi Baisas——Delfinado, 1966, 40 (¢*, 2, P*, L*). argyrotarsis Leicester.—Peters, 1964, 25 (¢*, 9, P, L). benoiti Wolfs.—Congo. 1964, in Benoit, Mus. Roy. Afr. Centr. Ann., Sci. Zool. 132: 309 (L*). Type-loc: Mwendua River, Kasongo, Congo (CMT). bertii Cova Garcia and Rausseo.— Venezuela. 1964. Rev. Sanid. Asist. soc. 29: 221 (6*, 2, P*, L*). Type-loc: Cumbre de Choroni, Aragua, Venezuela (DERM). bimaculata Leicester—Delete Philippines. cavernicola Mattingly.—Vattier and Adam, 1962 (1963), 911 (3%, Pek ye clara Dyar and Shannon.—Delfinado, 1966, 42 ( 6 *, P*, L*). hirsutifemora Peters.—Papua. 1964. Proc. Roy. ent. Soc. Lond. (B) 33: 21 (¢*, 2*). Type-loc: Port Moresby, Central District, Papua (CSIR). 115. 116. 107. 118. 119. 120. 121. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 205 ludlowae Dyar and Shannon.—Delfinado, 1966, 47 (¢, 2, L; syn.). reyi Baisas. maculipleura Leicester.—Wattal and Kalra, 1965, 312 ( ¢*). mendiolai Baisas.—Delfinado, 1966, 48 (¢*, 9, P*, L*). moresbyensis Peters.—Papua. 1963. Proc. R. ent. Soc. Lond. (B) 32: 204 (¢*). Type-loc: Port Moresby, Central Distr., Papua (CSIR). nivea Leicester.—Delfinado, 1966, 50 (6 *, 2, L*). novaguinensis Peters—New Guinea. 1963. Proc. R. ent. Soc. Lond. (B) 32: 141° (6*, 2, P*, L). Type- loc: Ami, 10 miles nw. of Maprik, Sepik District, New Guinea (CSIR). ssp. alticola Peters—New Guinea. 1963. Proc. R. ent. Soc. Lond. (B) 32: 142 (4, 2). Type- loc: Goroka, Eastern Highlands District, New Guinea (CSIR). obseura Edwards.—Third line, change “Edwards, 1928” to “Edwards and Given, 1928.” paralateralis Peters—New Guinea. 1964. Proc. R. ‘ent. Soc. Lond. (B) 33: 19 (6*, 2). Type-loc: Kavieng, New Ireland District, New Guinea (CSIR). paranovaguinensis Peters—New Guinea. 1963. Proc. R. ent. Soc. Lond. (B) 32: 135 (¢*, P*, L*),.. Type- loc: Maprik, Sepik District, New Guinea (CSIR). philippinensis Delfinado.—Philippines. 1966: J. med, Ent. 3: 36 (¢*,. 9, P*, L*).Type-loe: San Jose, Mindoro, Philippines (USNM). reyi Baisas.—Transfer to p. 115 as synonym of ludlowae Dyar and Shannon. rossi Delfinado.—Philippines. 1966. J. med. Ent. 3: 36 (6*, 2). Type-loc: San Jose, Mindoro, Philippines (USNM). testacea Theobald.—Delfinado, 1966, 53 (¢*, 2, P*, L*). tubanguii Baisas.—Delfinado, 1966, 55 (¢*, 2, P*, L*). Genus HODGESIA Theobald malayi Leicester.—Delfinado, 1966, 58 ( 6 *, P*, L*). Genus ORTHOPODOMYIA Theobald Thomasina Newstead and Carter—Change “Newstead and Carter” to “Newstead and Thomas” after Mansonia longipalpis. nigritarsis Leicester—Change “(NE)” to “(BM).” albipes Giles.—Delete entirely. Giles was not treating a Philippine species. nkolbissonensis Rickenbach and Hamon.—Cameroon. 1966, (1965), Bull. Soc: Path. exot.. 585 1112 (6%) 9)... Vype- loc: Nkolbisson, 8 km from Yaounde, S. Cameroon (IERT). Genus AEDEOMYIA Theobald catasticta Knab.—Delfinado, 1966, 72 (¢, 2, P*, L*). 206 128. 130. 135. PROC, ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Genus PSOROPHORA Robineau-Desvoidy Subgenus PSOROPHORA Robineau-Desvoidy ciliata (Fabricius).—Delete West Indies. Guedes et al., 1965, 15 ( * * cilipes (Fabricius ) —Guedes et al.; 1965, 15; 16 (dA ;24)e holmbergii (Lynch Ariibalzaga) —Guedes et al., 1965, LES lineata (Humboldt).—Delete Argentina, French Guiana, Brazil, Bo- livia. saeva Dyar and Knab.—Transfer below as valid species with genu- maculata as synonym. pallescens Edwards.—Guedes et al., 1965, 18 (2*). saeva Dyar and Knab.—Argentina, Brazil, Surinam, Trinidad, Vene- zuela, ?Bolivia, PFrench Guiana. Stone, 1967 (in press) (resurrected from synonymy). Subgenus JANTHINOSOMA Lynch Arribalzaga Important reference: 1964, Guedes and Souza, 471-486 (key). albigenu Peryassi.—Argentina, Bolivia, Brazil, Paraguay, Peru, Vene- zuela (northern limits of distribution uncertain). Guedes and Souza, 1964, 474 (6*, 2*, P*, L*; resurrected from synonymy). paraguayensis Strickland. bruchi Petrocchi. albipes (Theobald ).—Guedes and Souza, 1964, 476 ( 4*, P*, L*). circumflava Cerqueira.—Peru. Guedes et al., 1965, 18 (P*). cyanescens (Coquillett).—Guedes et al., 1965, 18 (6 *, 2*). diserucians (Walker ).—Guedes et al., 1965, 19 (6*, 2*). ferox (Humboldt ).—Guedes et al., 1965, TCS Fel pazosi Pazos.—Insert before “Cuba,” “Vuelta Abajo (Las Villas) .” fiebrigi Edwards. — Paraguay, Bolivia. Souza and Guedes, 1962, 249 (6*, 2*; resurrected from synonymy ). forceps Cerqueira. —Guedes et als, 1965, 20°( od *e2*): lanei Shannon and Cerqueira. Guedes et ale 1965, 18(fO")) lutzii (Theobald).—Change “Janthinosoma” to “Janhtinosoma.” Gue- des et al., 1965, 21 (3*, 9*). varipes (Coquillett).—Distribution and identity uncertain. Possibly Central American varipes is different from that in United States; most South American “varipes” is albigenu. albigenu Peryassi.—Transfer to p. 128 as valid species with paraguay- ensis and bruchi as synonyms. Subgenus GRABHAMIA Theobald cingulata (Fabricius ).—Guedes et al., 1965, 21, 22 (4*, 2*). neoapicalis Theobald.—Insert “; Culex” at end of second line, within parenthesis. confinnis (Lynch Arribalzaga) —Guedes et al., 1965, 23 (¢*, 2*). dimidiata Cerqueira.—Guedes et al., 1965, 23 (6*, 2*). discolor (Coquillett). —Change date from “1930” to “1903.” paulli Paterson and Shannon.—Guedes et al., 1965, 18 (@*). varinervis Edwards.—Guedes et al., 1965, 24 (6 *, 2*). Genus ERETMAPODITES Theobald tonsus Edwards.—Someren and Hamon, 1964, 82-84 (P*, L*) 137. 138. 145. 146. 147. 148. 149. 150. 151. 153. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 20 bo | Genus AEDES Meigen Subgenus OCHLEROTATUS Lynch Arribalzaga Aedes, subgenus Kompia Aitken.—Transfer to p. 157 before Sup- genus FINLAYA as valid subgenus. albifasciatus (Macquart).—Ronderos and Garcia 1962 (1963) a, 35 (ee angustivittatus Dyar and Knab.—Ronderos and Garcia, 1962 (1963)a, od (BE). meprai Martinez and Prosen.—Ronderos and Garcia, 1962 (USGS). so" (syns). = antipodeus (Edwards) .—Marks and Nye, 1963, 50 (¢*, 2, P*, L*). praeteritus Segiy.—Transfer to p. 150 as synonym of pulchritarsis. africanus Re aE eee “taeniorhynchus” to “Taeniorhyn- chus. borealis Ludlow.—Change “Culex” to “Culex.” communis (De Geer). ssp. nevadensis Chapman and Barr.—Nevada. 1964. Mosq. News 24: 444 (6*, 9, P*, L). Type loc: Lamoille Canyon, Elko Co., Utah, United States (USNM). patersoni Shannon and Del Ponte.—Transfer to p. 150 as valid sp. explorator Marks.—Australia. 1964. Pap. Dep. Ent. Univ. Qd. 2: 60 (2*, L*). Type-loc: Mt Olga, 300 mi. sw. of Alice Springs, Northern Territory, Australia (UQ). fryeri (Theobald).—Mattingly, 1963, 165 (L*). imperfectus Dobrotworsky.—Australia. 1962. \Proc, Linn Soe. N.S.W. 87: 296°(6*, 2, P*; L*). Type- loc: Woori Yallock, Victoria, Australia (USNM). lasaensis Meng.—China. 1962. Acta ent. Sinica 11: 159 (¢*, L*). Type-loc: Lasa [Tibet, China] (LU). linesi Marks.—Australia. 1964. Pap. Dep. Ent. Univ. Qd. 2: 63 (2). Type-loc: Robe, South Australia, Australia (CSIR). meprai Martinez and Prosen.—Transfer to p. 138 as synonym of angustivittatus. mombasaensis Mattingly— Kenya. 1963. Proc: R. ent. Soc. Lond. (B) 32:-.165 (¢, 9, L*). Type- loc: Mombasa, Kenya (BM). nigrithorax (Macquart).—Dobrotworsky, 1966, 135 (9*, L). patersoni Shannon and Del Ponte.—Ronderos and Garcia, 1962 (1963) a, 35 (P); (resurrected from syn.). phaecasiatus Marks.—Australia. 1964. Pap. Dep. Ent. Univ. Qd. 2: 64 (2). Type-loc: 5 miles from Maningrida, Amhem Land, Northem Territory, Australia (UQ). pulehritarsis (Rondani). praeteritus Seguy.—Coluzzi, in litt. (syn.). purpureipes Aitken.—Transfer to p. 157 under Subgenus KOMPIA. luteovittata Theobald.—Insert after “Culex” “nemorosus var.” sapiens Marks.—Australia. 1964, Pap. Dep. Ent: Univ. Od. 2: 66 (¢*, 9*). Type-loc: North Bourke, New South Wales, Australia (CSIR). mathist Neveu-Lemaire.—Change “French Guiana” to “Brazil.” 208 154. 155. 157. 165. 167. 168. 170. 171. 173. 175. 176. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 simanini Gutsevich.—U.S.S.R. (Uzbekistan). 1966. Zool. Zhur. 45: 457 (¢*, 2). Type-loc: Kokand, Fergana Valley, Uzbekistan (ZIL). dorso-vittatus Villeneuve.—Insert after “Culex”, “nemorosus var.” subalbirostris Klein and Marks.—Marks and Nye, 1963, 56 ( ¢*, 2%, Belts): Insert Subgenus KOMPIA Aitken. Mattingly, 1961, 16 (retumed to subgeneric status). purpureipes Aitken.—Insert from p. 151 under Subgenus KOMPIA. Subgenus FINLAYA Theobald kompi Vargas and Downs.—Arizona. mjoebergi (Edwards ).—Change to “mjdbergi.” occidentalis (Skuse). ssp. milsoni (Taylor).—Dobrotworsky, 1962, 301 (as sp.; P*, er. shintiensis.—Change “Lein” to “Lien.” sorsogonensis Banez and Jueco.—Philippines. 1966. Acta med. Philippina 2(4): 195 (¢*, 2*, L*). Type-loc: Sorsogon Province, [Luzon] (IHP). stanleyi Peters —New Guinea. 1963., Proc. R: ent. Soc; Lond. (B). 32:1 (¢¢ 19; B* ies) eeigae= loc: Minj, Western Highlands District, New Guinea (CSIR). subbasalis Dobrotworsky.—Australia. 1962. Proce. Linn. Soc..N:S.W. 87: 298) (6%, 25° P* ks eee loc: Ginninderra Falls, New South Wales, Australia (CSIR). tutuilae Ramalingam and Belkin.—Samoa. 1965. Contr. Amer. ent. Inst. 1(4): 3 (¢*, 9, P*, L*). Type-loc: Aoloau, Tutuila, American Samoa (USNM). Subgenus HOWARDINA Theobald alloteenon Kumm, Komp and Ruiz.—Add to type-locality, “Poas Volcano.” arborealis Bonne-Wepster and Bonne.—Change “USNM” to “ITH.” aurivittatus Cerqueira.x—Martinez, Prosen and Carcavallo, 1964, 54 (ieee cozumelensis Diaz Najera.—Mexico. 1966. Rev. Invest. Salud. publ. (Mex.) 26: 334 (4*, 9*, P*, L*). Type-loc: San Miguel de Cozumel, Quintana Roo, Mexico (ISET). Subgenus CHAETOCRUIOMYIA Theobald calabyi Marks.—Marks, 1964, 132 (2). elchoensis Taylor.—Marks, 1964, 134 (6*, 9, P*, L*). macmillani Marks.—Australia. 1964. Proc. Linn. Soc. N.S.W. 89: 138 (¢*, 2*). Type-toe: N.W. Dungog, Gummi Plain, Barrington Area, New South Wales (US). spinosipes Edwards.—Marks, 1964, 142 (2, E*). Subgenus HALAEDES Belkin australis (Erichson).—New Zealand. er: 180. 181. 182. 183. 184. 186. 187. 188. 189. 190. 194, PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 209 Subgenus GEOSKUSEA Edwards tonsus Edwards.—Mattingly, 1965, 23 (4¢*). Delete “? Celebes.” Subgenus STEGOMYIA Theobald albopictus (Skuse ).—Restore New Guinea, deleted in Suppl. I. arboricolus Knight and Rozeboom.—Change to arboricola. bambusicolus Knight and Rozeboom.—Change to bambusicola. Supplement II. montana Koidzumi. Change reference given to: “1918: Trans. nat.. Hist. Soc: Formosa 38; 141 (4; Stegomyia). Type-loc: Chikutoki, Kagi District, Formosa (LU)..” gardnerii (Ludlow). ssp. imitator (Leicester).—Malaya, Nepal, S. China. Mattingly, 1965, 36 (resurrected from synonymy; syn.). christianus Dyar. hensilli Farner.—Transfer to p. 187 as ssp. of scutellaris. laffooni Knight and Rozeboom.—Knight and Hull, 1951, 221 (L). paullusi Stone and Farner.—North Bormeo. seutellaris (Walker).—Delete Philippines. Colless, 1962, 313 ( é*, taxon). ssp. hensilli Farner.—Delete Singapore of Suppl. I. Colless, 1962, 314 (to ssp. status). ssp. malayensis Colless—Singapore, Malaya, Philippines. 1962. Proc. Linn. Soc. N.S.W. 87: 314 (¢*, 9). Type- loc: Pulau Hantu, Keppel Harbor, Singapore (CSIR). tabu Ramalingam and Belkin.—Tonga Islands. 1965. Contr. Amer. ent. Inst. 1(4): 1 (4*, 9, P*, L). Type-loc: Eua Island, Tonga (USNM). usambara Mattingly —Someren and Hamon, 1964, 80 (P*). vinsoni Mattingly.— Mattingly, 1963, 168 (¢@ ). imitator Leicester.—Transfer to p. 183 as ssp. of gardnerii. christianus Dyar.—Transfer to p. 183 as synonym of gardnerii imitator. Subgenus AEDIMORPHUS Theobald albocephalus Theobald.—Change reference to “Mem. Lpool Sch. trop. Med. 10(App.): 4. arabiensis (Patton).—Transfer with synonym to p. 198 as synonym of vexans (Meigen). dialloi Hamon and Brengues.—Dahomey. 1965. Bull. Soc. Path. exot. 58: 103 (4*). Type-loc: Tchakarkou, Natitingou Prefecture, Dahomey (IERT). ebogoensis Rickenbach and Ferrara.—Cameroon. 1966. Bull. Soc. Path. exot. 58: 26 (4*). Type-loc: Ebogo, Cameroon (IERT). hamoni Mattingly.— Nigeria. 1963. Proc. Ri ent; Soe ond:;(B)s32-166 (644 2 *)s 4lype- loc: Akute Village, Ogun River, near Iju waterworks, S. Nigeria (BM). lottei Hamon and Brengues.—Ivory Coast. 1965. Bull. Soc. Path. exot. 58: 101 (4*). Type-loc: Tiassalé, Ivory Coast (IERT). 210 195. 196. 198. 207. 209. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 niveoscutellum (Theobald) .—Delete asterisk after “¢.” nocturnus Theobald.—Hawaii. Correction to Suppl. II. Change “125° to “195.” nyounae Hamon and Adam.—Brengues and Hamon, 1965, 92 (P*, L* Me rickenbachi Hamon and Adam.—Rickenbach and Ferrara, 1964, 972, * * tauffliebi Rickenbach and Ferrera.—Cameroon. 1965. Bull. Soc. Path. exot. 58: 24 (6*). Type-loc: Mbéga, Dept. Dyong and Sanaga, Cameroon (IERT). vexans (Meigen).—Hamon et al., 1966, 373 (syn.). arabiensis Patton.—Insert in synonymy after montcalmi. sudanensis Theobald.—Insert in synonymy after eruthrosops. Subgenus NEOMELANOCONION Newstead Supplement II. Le Berre and Hamon reference inadvertently left out of “Literature Cited.” See that section in this Supplement. aurovenatus Worth—Supplement I. Change type locality from “Nduma” to “Ndumu.” palpalis (Newstead ).—Gambia. taeniarostris (Theobald ).—Bailly-Choumara, 1965 (1966), 671 (P*, Subgenus DICEROMYIA Theobald periskelatus (Giles ).—Qutubuddin, 1945, 34 (¢?). whartoni Mattingly —Malaya. 1965. Culic. Mosq. Indomalayan Area VI: 65 (8*, P*, L*). Type-loc: Ulu Gombak, Malaya (BM). Subgenus AEDES Meigen nubicolus Laffoon.—Change to nubicola. uneus (Theobald) .—Knight and Hull, 1951, 222 (L). Subgenus CANCRAEDES Edwards miachaetessa Dyar and Shannon.—Change “1935” to “1925.” lateralis Theobald.—Change “1919” to “1818.” Genus ARMIGERES Theobald Subgenus ARMIGERES Theobald baisasi Stone and Thurman.—Baisas 1935, 488 (¢*, 2; as kuchin- gensis); Delfinado, 1966, 80 (3 *; 2, P*, L*). . kuchingensis Edwards.—Transfer Baisas’s 1935b reference to baisasi Stone and Thurman. malayi (Theobald) .—Philippines. manalangi Baisas.—Delfinado, 1966, 86 (¢*, 2, P*, L*). papuensis Peters—New Guinea. 1963. Proce: R: .ent. Soe, Lond:2(B) 32: 4 (6) Cee eae Type-loc: Maprik, Sepik District, New Guinea (BM). setifer Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 87 (¢*, 2*). Type-loc: Pinigi- san, Mantalingajan, Palawan, Philippines (ZMC). bo bo i=) PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 PAIL Subgenus LEICESTERIA Theobald digitatus Edwards.—Delfinado, 1966, 88 (6 *, 2, P*, L*). flavus Leicester—Delfinado, 1966, 89 (4*, 2, P*, L*). magnus (Theobald) .—Delfinado, 1966, 91 ( ¢*, 2, P*, L*). omissus (Edwards ).—Delfinado, 1966, 92 (¢*, 2, P*, L*). Genus CULISETA Felt Important references: 1963, Barr, 324-330 (key pupae, Nearctic) ; 1964, Maslov, Ent. Obozr. 43: 193-217 (Ent. Rev. : 97-107) (taxon- omy). Subgenus CULISETA Felt alaskaensis (Ludlow) .—Barr, 1963, 328 (P*). ssp. indiea (Edwards).—India, Pakistan. Maslov, 1964, 203 (100) (to ssp. status). annulata (Schrank). ssp. subochrea (Edwards).—Maslov, 1964, 204 (100) (to ssp. status). atlantica (Edwards) .—Insert from p. 222. zottae (Ungureanu).—Change from ssp. to syn. impatiens (Walker) .—Barr, 1963, 329 (P). incidens (Thomson) .—Barr, 1963, 328 (P*). particeps (Adams) .—Barr, 1963, 328 (P*). sinensis (Meng and Wu) .—China. 1962. Acta ent. Sinica 11: 382 (6*, 2, P, L*; Theobaldia). Type-loc: China (LU). subochrea (Edwards) .—Transfer with its synonym to p. 218 as ssp. of annulata (Schrank). Subgenus CULICELLA Felt atra Lee.—Insert from p. 222. minnesotae Barr.—Transfer to p. 221 as ssp. of silvestris (Shingarev). morsitans (Theobald) .—Delete northern North America. ssp. dyari (Coquillett) —Northern North America. Maslov, 1964, 209 (103) (to ssp. status). brittoni Felt. ochroptera (Peus).—Place as ssp. of silvestris (Shingarev). silvestris (Shingarev). ssp. ochroptera (Peus).—Maslov, 1964, 209 (103) (to ssp. status). ssp. minnesotae Barr.—Maslov, 1964, 209 (103) (to ssp. status). ssp. amurensis Maslov.—China. 1964. Ent. Obozr. 43: 210 (¢*, 2, L). Type-loc: Cen- tral Amur Region, Ussuri Basin, Southem Maritime Province, Northeast China (LU). Subgenus CLIMACURA Howard, Dyar and Knab antipodea Dobrotworsky.—Australia. 1962; ‘Proc: kinn. SoexN:S-Wesie 29h (67%) 2. P*y i"). Type- loc: Cann River, Victoria (NMM). melanura (Coquillett ).—Barr, 1963, 327 (P*). weindorferi (Edwards ).—Dobrotworsky, 1966, 143 (é*, 2, P*, L*). PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Subgenus ALLOTHEOBALDIA Brolemann indica (Edwards) .—Transfer to p. 218 as ssp. of alaskaensis (Lud- low). Subgenus Uncertain atlantica (Edwards ).—Transfer to p. 218 in subgenus Culiseta. atra Lee.—Transfer to p. 220 in subgenus Culicella. Genus CULEX Linnaeus Subgenus LUTZIA Theobald . fuseanus Wiedemann.—Delfinado, 1966, 97 ( 6 *, 2, P*, L*). setulosus Doleschall—Change “1875” to “1857.” bimaculata Theobald.—Insert after “2” “; as var.” fusca Theobald.—Insert after “A” “; as var.” Subgenus NEOCULEX Dyar amaniensis Someren and Hamon.—Tanzania. 1964. J. ent. Soc. S. Afr. 27: 78 (¢*, 9*). Type-loc: Amani, Tanganyika (BM). 6. brevipalpis (Giles). —Delfinado, 1966, 125 (4*, 2, P*, L*). bo bo garioui Bailly-Choumara and Rickenbach.—Cameroon. 1966. Bull. Soc. Path. exot. 59: 144 (6*). Type-loc: Mbol, Yokadouma, E. Cameroon (IERT). laplantei (Hamon, Adam and Mouchet).—Change “Eboga” to “Ebogo.” nematoides Dyar and Shannon.—Insert “2.” pseudoandreanus Bailly-Choumara.—Cameroon. 1965 (1966). Bull. Soc. Path. exot. 58: 660 (4*). Type-loc: Ebogo, Arrondissement de Mbalmayo, 100 km. s. of Yacundé, Cameroon (IERT); Bailly-Choumara and Pajot, 1965 (1966): G65¢R* Ils): capensis De Meillon.—Insert after “?” “as var.” Subgenus MOCHTHOGENES Edwards chiyutoi Baisas.—Delfinado, 1966, 129 (¢*, P, L). culionicus Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 130 (¢*, 2). Type-loc: San Pedro, Culion I., Palawan, Philippines (USNM). latifoliatus Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 131 (6*, 2). Type-loc: Molawin Creek, College, Laguna, Philippines (USNM). laureli Baisas—Philippines. Delfinado, 1966, 132 (4*, 2, P*, L*; resurrected from synonymy ). malayi (Leicester).—Delete Philippines and remove laureli from synonymy. shrivastavii Wattal, Kalra and Krishnan.—Andaman Islands. 1966. Bul. Ind. Soc. Mal. Com. Dis. 3: 159 (4*). Type-loc: Ferrar Ganj, S. Andaman Islands (NCID). tricontus Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 133 (4*). Type-loc: Molawin Creek, College, Laguna, Philippines (USNM). (oe) PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 All uncinatus Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 134 (4*, 2). Type-loc: Osmena, Samar (USNM). Subgenus LOPHOCERAOMYIA Theobald Important reference: 1965, Colless, 216-307 (Malaya, keys). aculeatus Colless—Malaya. 1965. J. med. Ent. 2: 286 (4*, 9, L*). Type-loc: Bukin Tungal, Kg. Gajan Forest Reserve, Perak, Malaya (ANIC). acutipalus Colless.—Malaya. 1965. J. med. Ent. 2: 299 (4*, 9, L). Type-loc: Kg. Chantek Bahru, Singapore (ANIC). alphus Colless.—Malaya, Borneo. 1965. J. med. Ent. 2: 283 (4*, 9, L*). Type-loc: Kg-Sijankang, Selangor, Malaya (ANIC). atracus Colless—Change “BM” to “CSIR.” christiani Colless—Change “BM” to “CSIR.” bandoengensis Brug.—Java, Malaya. Colless, 1965: 292 (é*, 9, L; to sp. status). bengalensis Barraud.—India, Malaya, Hainan I. Colless, 1965, 293 Coes On 1. to spy status) bicornutus (Theobald).—Burma, Malaya, Botel Tobagu (nr. For- mosa). Colless, 1965, 291 (4*, L*; resurrected from syn.). brevipalpus (Theobald) .—Stone, 1957, 172 (resurrected from syn.) ; Colless, 1965, 301 (é*, 2, L). cinctellus Edwards.—Philippines. crassicornis Colless.—Malaya. 1965. J. med. Ent. 2: 296 (¢*). Type-loc: Ulu Gombak, Selangor Malaya (ANIC). cubitatus Colless—Malaya. 1965. J. med. Ent. 2: 273 (é6*, 2, L). Type-loc: Mandai Road, Singapore (ANIC). curtipalpis (Edwards).—Edwards, 1928, 279 (4*); Colless, 1965, SOTO 702 1L)k demissus Colless.—Malaya. 1965. J. med. Ent. 2: 296 (¢6*). Type-loc: Ulu Gombak, Selan- gor, Malaya (ANIC). eminentia (Leicester ).—Colless, 1965, 298 (¢*, L). brevipalpus Theobald.—Transfer to p. 232 as valid species. eukrines Bram and Rattanarithikul.—Thailand. 1967. Proc. ent. Soc. Wash. 69: 11 (6*, 9, L*). Type-loc: Huai Bong Ti, Kanchanaburi Province, Thailand (USNM). variata Leicester —Change “NE” to “BM.” Transfer to p. 235 as valid species. fuscosiphonis Bram and Rattanarithikul—Thailand. 1967. Proc. ent. Soc. Washington 69: 11 (¢*, 2, L*). Type- loc: Muang, Phattalung Province, Thailand (USNM). ganapathi Colless.—Malaya. 1965. J. med. Ent. 2: 294 (4*, 9, L*). Type-loc: Ulu Langat, Selangor, Malaya (ANIC). gibbulus Delfinado.—Philippines. 1966. Mem. Amer. ent. Inst. 7: 105 (6*). Type-loc: San Jose, Mindoro, Philippines (USNM). 214 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 hewitti (Edwards) .—Colless, 1965, 303 (¢*, 2, L). incomptus Bram and Rattanarithikul—Thailand. 1967. Proc. ent. Soc. Wash. 69: 16 (8*, 2). Type-loc: Doi Sutep, Chiang Mai Province, Thailand (USNM). inculus Colless——Malaya. 1965. J. med. Ent. 2: 280 (¢*, 2, L*). Type-loc: Kampong Sireh, Selangor, Malaya (ANIC). josephinae Baisas—Change to josephineae and transfer to p. 234 as valid species. . josephineae Baisas.—Philippines. Delfinado, 1966, 106 (4*; resur- rected from synonymy). luearis Colless—Malaya. 1965. J. med. Ent. 2: 299 (¢*). Type-loc: Ulu Pandan, Singa- pore (ANIC). bicornuta Theobald.—Transfer to p. 232 as valid species. macdonaldi Colless—Malaya, Philippines. 1965. J; med: Ent: 2:-276 (3*, L*). Type-loc: Singapore (ANIC); Delfinado, 1966, 108 ( ¢*, P*, L*). mammilifer (Leicester ).—Hainan I. Colless, 1965, 287 (4*, 2, L*; syn.); Delfinado, 1966, 109 ( 4 *, P*, L*). chiungchungensis Hsiii—1963. Acta ent. Sinica 12: 229 (4*, 9, L*). Type-loc: Chiungchungsang, Hainan Island, Kwang- tung Province, China (LU). minor (Leicester ).—Colless, 1965, 289 (6 *, 2, L*; syn.). nolledoi Baisas. var. bengalensis Barraud.—Transfer to p. 232 as full species. var. bandoengensis Brug.—Transfer to p. 232 as full species. 5. navalis Edwards.—Malaya. Colless, 1965, 304 (4*, 29, L). niger (Leicester ).—Colless, 1965, 282 (6*, 2, L). nolledoi Baisas.—Transfer to p. 234 as synonym of minor (Leicester). petersi Colless—Change “BM” to “CSIR.” pseudornatus Colless—Change “BM” to “CSIR.” peytoni Bram and Rattanarithikul.—Thailand. 1967. Proc. ent. Soc Wash. 69: 7 (¢*, 9, L*). Type-loc: Phatt- halung-Trang Road, Phatthalung Province, Thailand (USNM). pholeter Bram and Rattanarithikul.—Thailand. 1967. Proc. ent. Soc. Wash. 69: 13 (4*, 9, L*). Type-loc: Khao Mai Ha Wa, Chon Buri Province, Thailand (USNM). reidi Colless.—Malaya, Philippines. 1965. J. med. Ent. 2: 279 (¢*, 2, L*). Type-loc: West Coast Road, Singapore (ANIC). spiculosus Bram and Rattanarithikul.—Thailand. 1967. Proc. ent. Soc. Wash. 69: 3 (¢*, 2, L*). Type-loc: Doi Sam Sao, Tak Province, Thailand (USNM). traubi Colless.—Malaya. 1965. J. med. Ent. 2: 295 (4*, 2, L*). Type-loc: Ulu Gombak, 16th mile, Selangor, Malaya (ANIC). uniformis (Theobald) .—Philippines. variatus (Leicester).—Colless, 1965, 270 (6*, 2, L*; resurrected from synonymy ). whartoni Colless—Malaya. 1965. J. med Ent. 2: 275 (é*, 2, L*). Type-loc: Singapore (ANIC). 238. 239. 240. 241. 242. 243. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 215 wilfredi Colless—Malaya. 1965. J. med. Ent. 2: 297 (4*). Type-loc: Malaya (ANIC). Subgenus CULICIOMYIA Theobald pallidothorax Theobald.—Philippines. papuensis (Taylor) .—Philippines. ramakrishnii Wattal and Kalra.* 1965. Bull. Ind. Soc. Mal. Com. Dis. 2: 309 (6*, 2). Type-loc: Lachhiwala Reserve Forest, Dehra Dun, India (NICD). spathifurea (Edwards) .—Delfinado, 1966, 123 (é*, 2, P, L*). India. Subgenus THAIOMYIA Bram Culex, subgenus Thaiomyia Bram, 1966, Proc. ent. Soc. Wash. 68: 73. Orthotype: Culex dispectus Bram. dispectus Bram.—Thailand. 1966: Proe. ent. Soc. Wash. 68: 75 (6%, 2) P*, L*). Type-loc: Doi Sam Sao, Tak Province, Thailand (USNM). Subgenus CULEX Linnaeus Important reference: 1967, Bram, 1-222 (keys, New World). Lasioconops Theobald.—Change reference to Mem. Lpool Sch. trop. Med. 10 (App.): 8. abnormalis Lane.—Colombia. Bram, 1967, 20 (6 *, 2, L; syn.). scutatus Rozeboom and Komp. acharistus Root.—Argentina, Chile. Bram, 1967, 23 ( ¢*, L*). alani Forattini— Colombia. 1965. Entomologica Médica 2: 143 (é*). Type-loc: Catera a Gachancipa, Colombia (USNM); Bram, 1967, 23 ( 6*). alticola Martini—Transfer to p. 242 as synonym of apicinus Philippi. annulirostris Skuse.—Delfinado, 1966, 139 ( 6*, 2, L*; syn.). palmi Baisas. gambiensis Theobald.—Insert “; as var.” after “(2 ).” apicinus Philippi—Bolivia. Bram, 1967, 26 (6 *, 2, L; syn.). alticola Martini. archegus Dyar.—Peru, Colombia, Ecuador. Bram, 1967, 28 ( ¢*, L; resurrected from syn.). Place before argenteopunctatus. articularis Philippi—Type-loc: Casa Pangue, Llanquihue, Chile (USNM). Bram, 1967, 30 ( ¢*; neotype). azuayus Levi Castillo—Transfer to p. 252 as synonym of nigripalpus Theobald. beauperthuyi Anduze.—Transfer to p. 259 as synonym of saltanensis Dyar. bickleyi Forattini—Colombia. 1965. Entomologia Médica 2: 146 (4*). Type-loc: Bosque Cal- deron, Bogota, Colombia (USNM); Bram, 1967, 33 (¢*). bidens Dyar.—Argentina, Bolivia, Mexico, Venezuela. Bram, 1967, 35 (4 *; resurrected from syn., syn.). interfor Dyar. ? Under Article 31 of the International Code of Zoological Nomenclature this name is presumably not emendable; under the old Code it would have to be changed to ramakrishnani, as the species is named after Dr. Ramakrishnan. 246. 247. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 brethesi Dyar.—Bachmann and Casal, 1962, 78 ( 4 *, P*, L*). camposi Dyar.—Colombia, Ecuador, Peru. Bram, 1967, 49 (4%; raised to sp. status). cheni Ho.—China. 1963. Acta ent. Sinica 12: 368 (3*, 9, L*). Type-loc: Canton, China (CMC). chidesteri Dyar.—Argentina. Bram, 1967, 42 ( 6 *, L; syn.). deanei Correa and Ramalho. confusus Baisas.—Transfer to p. 251 as synonym of mimulus Ed- wards. coronator Dyar and Knab.—Bram, 1967, 46 ( 6 *, L; syn.). coronator mooseri Vargas and Martinez Palacios. ousqua Dyar.—Transfer to p. 253 as valid species. usquatus Dyar.—Transfer to p. 264 as valid species. usquatissimus Dyar.—Transfer to p. 264 as valid species. albertoi Anduze.—Transfer to p. 253 as synonym of ousqua Dyar. ssp. camposi Dyar.—Transfer to p. 244 as valid species. ssp. mooseri Vargas and Martinez Palacios —Change to synonym. covagarciai Forattini— Venezuela. 1965. Entomologia Médica 2: 128 (4). Type-loc: San Antonio de Caparo, Municipal Capitol, Venezuela (USNM); Bram, 1967, Do (C7415): deanei Correa and Ramalho.—Transfer to p. 244 as synonym of chidesteri Dyar. declarator Dyar and Knab.—Bram, 1967, 56 ( ¢*, L; syn.). forattinii Correa and Ramalho. bidens Dyar.—Transfer to p. 243 as valid species. fasyi Baisas—Change “[Davao], Mindanao” to “[Mountain], Luzon.” fernandezi Casal, Garcia and Cavalieri—Argentina. 1966. Physis 26: 185 (4*, 2, P*, L*). Type-loc: Quebrada de Piquirenda, Salta, Argentina (INM). forattinii Correa and Ramalho.—Transfer to p. 245 as synonym of declarator Dyar and Knab. gameti Bailly-Choumara.—Cameroon. 1966. Bull. Soc. Path. exot. 59: 148 (¢*). Type-loc: Mbol, Yoka- douma, E. Cameroon (IERT). habilitator Dyar and Knab.—Peru. incognitus Baisas.—Delfinado, 1966, 145 (4 *, 2, P, L*). interfor Dyar.—Transfer to p. 243 as synonym of bidens Dyar. kesseli Belkin.— Belkin, 1964, 236 (¢*, 2, P*). lahillei—Correction to Suppl. II. Change “Casals” to “Casal”; also index, p. 140. levicastilloi Lane.—Venezuela. Bram, 1967, 74 ( 6 *, L; syn.). tejerai Cova Garcia.—1962. Rev. Sanid. Asist. Soc. 27: 312 (4%, 9, P*, L*). Type-loc: Near La Cumbre de Choroni, Aragua, Venezuela (DERM). mimulus Edwards.—Delfinado, 1966, 150 (¢*, 2; syn.). confusus Baisas. mollis Dyar and Knab.—Bram, 1967, 80 ( é *, L; syn.). tisseuili Senevet. nigripalpus Theobald.—Bram, 1967, 84 ( 4 *, L; syn.). azuayus Levi-Castillo. 264. 275. 276. 278. bo — ~l PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 oswaldoi Forattini.—Brazil. 1965. Entomologia Médica 2: 167 (4*). Type-loc: Macaphyba, Natal, Brazil (USNM); Bram, 1967, 86 ( 6*). ousqua Dyar.—Mexico to Colombia. Bram, 1967, 50 (4 from syn.; syn.). albertoi Anduze. palmi Baisas.—Transfer to p. 241 as synonym of annulirostris Skuse. ssp. quinquefasciatus Say.—Restore to this status. trifurcatus Fabricius.—Delete “fatigans ssp.” pseudovishnui Colless.—Delfinado, 1966, 147 (P, L*). quinquefasciatus Say.—Return to p. 254 as ssp. (see Suppl. I). salinus Baisas.—Transfer to p. 260 as synonym of sitiens Wiedemann. saltanensis Dyar.—Brazil, Panama, Venezuela. Bram, 1967, 97 ( 4%, Leisyn:): beauperthuyi Anduze. scutatus Rozeboom and Komp.—Transfer to p. 242 as synonym of abnormalis Lane. sitiens Wiedemann.—Delfinado, 1966, 151 (¢*, 2, L*; syn.). salinus Baisas. spinosus Lutz.—Colombia, Venezuela. suborientalis Baisas—Change “[Davao], Mindanao” to “[Mountain], Luzon.” thriambus Dyar.—Dominican Republic, Costa Rica, Colombia. tisseuilli Senevet—Emended by Belkin, 1965, 24 to tisseuili. Trans- fer to p. 251 as synonym of mollis Dyar and Knab. siamensis Barraud and Christophers.—Insert “; as var.” after “4. usquatissimus Dyar.—Costa Rica to Ecuador and British Guiana. Bram, 1967, 51 ( 6 *; resurrected from synonymy). usquatus Dyar.—Mexico to Argentina. Bram, 1967, 49 (4*). *. resurrected ”» Subgenus MELANOCONION Theobald ocossa Dyar and Knab.—Change “Surinam” to “British Guiana.” elarki mE alae Casal, 1963, 317 (4*; resurrected from syn.). eastor Dyar.—Trinidad. implicatus Senevet and Abonnenc.—Transfer to p. 273 as synonym of nigrescens. nigrescens (Theobald ).—Casal, 1963, 317 ( 4*). implicatus Senevet and Abonnenc. clarki Evans.—Transfer to p. 268 as valid species. oedipus Root.—Garcia and Casal, 1965, 9 (P*). portesi Senevet and Abonnenc.—French Guiana, Trinidad, Brazil. Aitken and Galindo, 1966, 202 ( 4*, 2; resurrected). cayennensis Floch and Abonnenc. rabanicolus Floch and Abonnenc.—Change to rabanicola. portesi Cenevet and Abonnenc.—Transfer to p. 274 as valid species. eayennensis Floch and Abonnenc.—Change “PIG” to “NE”; transfer to p. 274 as synonym of portesi. ybarmis Dyar.—Trinidad. zeteki Dyar.—Trinidad. Subgenus MICROCULEX Theobald Important reference: 1962, Coutinho and Forattini, 209-234. bo (ee) i) 286. 290. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 intermedius Lane and Whitman.—Coutinho and Forattini, 1962, 217 (8*, 9, P*, L*). lanei Coutinho and Forattini—Brazil. 1962. Studia ent. 5: 210 (é*, 2, P*, L*). Type-loc: Bertioga (Itaguaré) , Santos, Sao Paulo, Brazil (FH). pleuristriatus Theobald.—Coutinho and Forattini, 1962, 223 oe PE OES): shopei Forattini and Toda.—Brazil. 1966. Studia ent. 9: 507 (4*, 2*, P*, L*). Type-loc: Utinga, Belem, Para, Brazil (FH). xenophobus Ronderos.—Venezuela. 1965. Physis 25: 249 (6*, 9*, P*, L*). Type-loc: San Bonifacio, Sucre, Venezuela (INM). Subgenus AEDINUS Bourroul americanus (Neveu-Lemaire).—Transfer to p. 283 as a nomen dubium. Change French Guiana to Brazil. bisculatus Coquillett—Trinidad, Antilles. Transfer to follow belemen- sis as a valid species with antillum-magnorum of authors as a synonym. canaanensis Lane and Whitman.—Change “1934” to “1943.” eauchensis Floch and Abonnenc.—Change “Moyenne-Mana” to iCaunx Subgenus CARROLLIA Lutz bihaicolus Dyar and Nufiez Tovar.—Change to bihaicola. Nomina Dubia americanus (Neveu-Lemaire).—French Guiana. Fauran, 1961, 45 (taxon. ). Genus DEINOCERITES Theobald pseudes Dyar and Knab.—Texas. Fossil Species loewi Giebel.—Place “Giebel” in parentheses; change “Culex” to “Toxorhynchites’; insert “Culex” after “A”; change SLi to SININ Gus Literature Cited Adam, Bruce-Chwatt, and Hamon. 1956. (See Catalog.) Adam, Hamon, Rickenbach, and Lips. 1956. (See Catalog.) Aitken, T. H. C. and P. Galindo. 1966. On the identity of Culex (Melanoconion) portesi Senevet and Abonnenc 1941 (Diptera: Culicidae). Proc. ent. Soc. Wash. 68: 198-208, illus. Aken (Supplement II.) Change “onderzbek” to “onderzoek.” Arab, B. Al-Tikrity. 1963. Recent studies on the ecology of Anophe- les apoci in Iraq. I. The geographical distribution of the breeding places and the morphology of Anopheles apoci pupae. Irak Bul. endemic Dis. 5: 73-81, illus. 293. 294. 296. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 219 Bachmann, A. O. and O. H. Casal. 1962. Notas s6bre Culex (Culex) Argentinés (Diptera: Culicidae). Ann. Inst. nac. Microbiol. 1: 77-81, illus. Bailly-Choumara, H. 1965 (1966). Description de la larve et de la nymphe d’Aedes (Neomelaniconion) taeniarostris Theobald, 1910. Observations sur une variation de coloration chez ladulte. Bull. Soc. Path. exot. 58: 671-676, illus. Bailly-Choumara, H. and P. X. Pajot. 1965 (1966). Contribution a l'étude des Culex (Neoculex) de la région éthiopienne (Diptera, Culicidae). Description de la larvae et de la nymphe de Culex (Neoculex) pseudoandreanus Bailly-Choumara. Bull. Soc. Path. exot. 58: 665-671, illus. Baisas, F. E. 1935. (See Catalog, 1935b.) Barr, A. R. 1963. Pupae of the genus Culiseta Felt. I. Descriptions and a key to the North American species (Diptera: Culicidae). Ann. ent. Soc. Amer. 56: 124-330, illus. Belkin, J. N. 1964. The adults and pupae of Culex (C.) kesseli from Tahiti and remarks on the atriceps group (Diptera: Culicidae). Ann. ent. Soc. Amer. 57: 236-239, illus. Belkin, J. N., R. X. Schick, and S. J. Heinemann. 1965. Mosquito studies (Diptera: Culicidae) V. Mosquitoes originally described from Middle America. Contr. Amer. ent. Inst. 1(5): 1-95. Bram, R. A. 1967. Classification of Culex subgenus Culex in the New World (Diptera: Culicidae). Proc. U. S. Nat. Hist. Mus. 120 (3557) > 1-122, illus. Brengues, J. and J. Hamon. 1965. Description de la larve et de la nymphe d’Aedes (Aedimorphus) nyounae Hamon et Adam, 1958. Bull. Soc. Path. exot. 58: 92-101, illus. Casal, O. H. 1963. Comentarios a proposito de la revalidacion de “Culex (Melanoconion) clarki” Evans (Diptera: Culicidae). Rev. Bras. Biol. 23: 317-320, illus. Cerqueira, N. L. 1961. Sobre Trichoprosopon (R.) edwardsianum (Lane & Cerqueira, 1942) (Diptera Culicidae). Studia ent. 4: 459-464, illus. Colless, Donald H. 1962. Notes on the taxonomy of the Aedes scut- ellaris group, and new records of A. paullusi and A. albopictus (Diptera: Culicidae). Proc. Linn. Soc. N.S.W. 87: 312-315, illus. . 1965. The genus Culex subgenus Lophoceraomyia in Malaya (Diptera: Culicidae). J. med. Ent. 2: 16-306, illus. Coluzzi, M. 1962. Le forme di Anopheles claviger Meigen indicate con i nomi missirolii e petragnanii sono due specie riproduttiva- mente isolate. R. C. Accad. Lincei 32: 1025-1030. . 1964. Morphological divergences in the Anopheles gambiae complex. Riv. de Malariol. 43: 197-232, illus. Corbet, P. S. 1963. Observations on Toxorhynchites brevipalpis con- radti Griinb. (Diptera, Culicidae) in Uganda. Bull. ent. Res. 54: 9-17, illus. Coutinho, J. O. and O. P. Forattini. 1962. Notas sobre Culicidae (Diptera). 4a—Contribuicao ao estudo do subgénero Microculex com a descricao duma nova espécie. Studia ent. 5: 209-234, illus. 297. Delfinado, M. D. 1966. The Culicine mosquitoes of the Philippines, tribe Culicini (Diptera, Culicidae). Mem. Amer. ent. Inst. 7: 1- 252, illus. 299. 300. 301. 303. 304. 306. 308. 309. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Dobrotworsky, N. V. 1962. Notes on Australian mosquitoes (Dip- tera, Culicidae). VI. Five new Victorian species and a descrip- tion of the larva of Aedes milsoni (Taylor). Proc. Linn. Soc. N.S.W. 87: 291-302, illus. 1966. Mosquitoes of Tasmania and Bass Straits Tela seeProns linn, Soc. NIS |W. Ole 121-146.allus: Dodge, H. R. 1963. Studies on mosquito larvae. I. Later instars of eastern North American species. Canad. Ent. 95: 796-813, illus. . 1964. Larval chaetotaxy and notes on the biology of Toxorhynchites rutilus septentrionalis (Diptera: Culicidae). Ann. ent. Soc. Amer. 57: 46-53, illus. 1966. Studies on mosquito larvae II. The first- stage larvae of North American Culicidae and of world Anophe- linae. Canad. Ent. 98: 337-393, illus. Edwards, F. W. 1928. (See Catalog. ) Fauran, P. 1961. Catalogue annoté des culicidés signalés en Guyane Frangaise. Arch. Inst. Pasteur Guiane Inini 465: 1-60. Garcia, M. and O. H. Casal. 1964 (1965). Descripcion de tres pupas de mosquitos neotropicales (Diptera, Culicidae). Rev. Soc. ent. Argentina 27: 5-10, illus. . 1965. Culicidae (Diptera) del Delta del Parana. II. Apuntes systematicos y biologicos. Delta del Parana 5(8): 5-16, illus. Garcia, M. and R. A. Ronderos. 1962. Mosquitoes de la Republica Argentina. I. Tribu Anophelini (Diptera—Culicidae—Culicinae) . An. Com. Cient. Prov. Bs. As. 3: 103-212, illus. Guedes, A. S. and M. A. de Souza. 1964. Sdbre Psorophora (Jan- thinosoma) albigenu Lutz, 1908 e Psorophora (Janthinosoma) albipes (Theobald, 1907) (Diptera, Culicidae). Rev. Bras. Malar. e Doengas trop. 16: 471-486, illus. Guedes, A. S., M. A. Souza, C. S. Maciel, and S. H. Xavier. 1965. Catalogo illustrado dos mosquitoes da colecao do Instituto Nacional de Endemias Rurais. I. Género Psorophora Robineau-Desvoidy, 1827. Rev. bras. Malar. e Doengas trop. 17: 3-24, illus. Hamon, J., M. Maffi, P. Grenier, C. S. Quédraogo, and D. Djimé. 1966. Notes sur les moustiques de la République Islamique de Mauritanie (Dipt. Culicidae) (Ile partie). Ann. Soc. ent. France 11: 371-383. International Commission on Zoological Nomenclature. 1965. Opinion 737. Bironella gracilis Theobald, 1905 (Insecta, Diptera) : Validated under the plenary powers. Bull. zool. Nomencl. 22: 164-166. Knight, K. L. and W. B. Hull. 1951. (See Catalog, 1951b.) Le Berre, R. and J. Hamon. 1960 (1961). Description de la larve, de la nymphe et de la femelle d’Aedes (Neomelanoconion) jamoti Hamon et Rickenbach, 1954, et révision des clés de détermination concernant le sous-genre Neomelanoconion en Afrique au Sud du Sahara. Bull. Soc. Path. exot. 53: 1054-1064, illus. Lien, J. C. 1965. Mosquitoes of Taiwan: Genus Toxorhynchites Theobald. J. med. Ent. 2: 1-16, illus. Lima, A. da Costa, N. Guitton, and O. Ferreira. 1962. Comentarios relativos as espécies da tribo Toxorhynchitini (Megarhinini) com a descricao de uma espécie nova de Lynchiella (Diptera, Culici- dae). Mem. Inst. Osw. Cruz 60: 225-252, illus. bo bo — PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Lips, M. 1962. Anopheles du Congo (ex Belge). 7. Quelques espéces des groupes Cellia et Neocellia. Références-récoltes-répartition et importance médicale actuelle. Riv. di Parassitol. 23: 107-134. 310. Marks, E. N. 1964. Notes on the subgenus Chaetocruiomyia Theo- oad (Diptera: Culicidae). Proc. Linn. Soc. N.S.W. 89: 131-147, illus. Marks, E. N. and E. R. Nye. 1963. The subgenus Ochlerotatus in the Australian Region (Diptera: Culicidae). VI. The New Zea- land species. Trans. R. Soc. N.Z., Zool. 4: 49-60, illus. 311. Martinez, A., A. F. Prosen and R. U. Carcavallo. 1964. El alotipo macho de Aedes (Howardina) aurivittatus Cerqueira. An. Inst. Med. Reg. 1962/63, 6: 53-58, illus. Maslov, A. V. 1964. On the systematics of bloodsucking mosquitoes of the group Culiseta (Diptera, Culicidae). Ent. Obozr. 43: 193- 217, illus. Ent. Rev. 97-107. Mattingly, P. F. 1961. The culicine mosquitoes of the Indomalayan Area. Part V. Genus Aedes Meigen, subgenera Mucidus Theobald, Ochlerotatus Lynch Ariibalzaga, and Neomelanoconion Newstead. British Museum, London, 62 pp., illus. . 1963. New and remarkable Aedes (Diptera: Culici- dae) from Africa. Proc. R. ent. Soc. Lond. (B) 32: 165-170, illus. . 1965. The holotype of Aedes (Geoskusea) tonsus See (Diptera: Culicidae). Proc. R. ent. Soc. Lond. (B) 34: 23, illus. . 1965. The culicine mosquitoes of the Indomalayan Area. Part VI. Genus Aedes Meigen, subgenus Stegomyia Theo- pad (Groups A, B, and D). British Museum, London, 67 pp., illus. 315. Peters, W. 1963a. Mosquitoes of New Guinea (Diptera: Culicidae). Part V. New species and previously undescribed pupae of the genus Tripteroides Giles. Proc. R. ent. Soc. Lond. (B) 32: 63-71, illus. . 1963b. Mosquitoes of New Guinea (Diptera: Cu- licidae). VI. The genus Tripteroides Giles: a new species of the subgenus Polylepidomyia Theobald, notes on the taxonomy and distribution, and a revised key to described pupae. Proc. R. ent. Soc. Lond. (B) 32: 89-100, illus. ————_———. 1964. Mosquitoes of New Guinea (Diptera: Cu- licidae). Part IX. Further new species of Uranotaenia Lynch Arribalzaga and notes on the genus. Proc. R. ent. Soc. Lond. (B) 33: 19-29, illus. 316. Qutubuddin, M. 1945. A description of the hitherto unknown female of Aedes (Diceromyia) periskeletus (Giles) 1902. Proc. nat. Acad. Sel, India 15(2): 33-35% Quy, Do Van. 1963. Clefs pour la détermination des 25 espéces les plus communes d’Anopheles (adultes, femeles et larves) du Sud- Viétnam. Rapp. ann. Fonct. tech. de Inst. Pasteur du Viet-Nam, Ann. 1962: 26-34, illus. 317. Reid, J. A. 1963a. Notes on anopheline mosquites from Malaya, with descriptions of three new species. Ann. trop. Med. Parasit. 57: 97-116, illus. . 1963b. A note on the structure of the pupal trum- pet in some mosquitoes. Proc. R. ent. Soc. Lond. (B) 38: 32-38, illus. 317. 324. 325. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 —_——_—_————.. 1965. A revision of the Anopheles aitkenii group in Malaya and Borneo. Ann. trop. Med. Parasit. 59: 106-125, illus. —_—__—_—_—_——.. 1966. A note on Anopheles subpictus Grassi and A. wigaieens Ludlow (Diptera: Culicidae). J. med. Ent. 3: 327-331, ius. Reid, J. A. and J. Bonne-Wepster. 1966. Anopheles errabundus (Swellengrebel, 1925) (Insecta, Diptera): proposed suppression as a specific name under the plenary powers. Z.N. (S.) 1760. Bull. zool. Nomencl. 23: 190. Rickenbach, A. and L. Ferrara. 1964. Description de la larve et de la nymphe de Aedes (Aedimorphus) rickenbachi Hamon et Adam, sae (Diptera: Culicidae). Bull. Soc. Path. exot. 57: 972-978, illus.. Ronderos, R. A. and A. O. Bachmann. 1962 (1963). A proposito del complejo Mansonia (Diptera-Culicidae). Rev. Soc. ent. Argen- tina 25: 43-51, illus. . 1963. Mansoniini neotropicales I (Diptera-Culici- dae). Rev. Soc. ent. Argentina 26: 57-65, illus. Ronderos, R. A. and M. Garcia. 1962 (1963)a. Description de pupas de Aedes argentinos (Diptera-Culicidae). Rev. Soc. ent. Argentina 25: 35-37, illus. ———————. 1962 (1963)b. Nota sinonimica en Culicidae (Diptera). Rev. Soc. ent. Argentina 25: 38. Russell, P. F., L. S. West, R. D. Manwell, and G. MacDonald. 1963. Practical Malariology, 2nd Ed., xv + 750 pp., London, Oxford University Press, illus. Service, M. W. 1962. Keys to the West African Anophelini. Acta Tropica 19: 120-158. ———————————.. 1963. The ecology of the mosquitoes of the north- ern Guinea savannah of Nigeria. Bull. ent. Res. 54: 601-632, illus. Slooff, R. 1963. Some notes on Bironella (Bironella) confusa Bonne- Wepster (Diptera; Culicidae; Anophelinae). Ent. Ber. Amst. 23: 137-142, illus. Someren, E. C. van and J. Hamon. 1964. A new species of Culex from Tanganyika, the descriptions of the pupa of Aedes usumbara Mattingly and the early stages of Eretmapodites tonsus Edwards. J. ent. Soc. S. Afr. 27: 78-85, illus. Souza, A. and A. 8S. Guedes. 1962. Sdbre Psorophora (Janthinosoma) fiebrigi Edwards, 1922 (Diptera—Culicidae). Rev. bras. Malariol. e Doengas trop. 14: 249-254, illus. Steffan, W. A. 1966. A checklist and review of the mosquitoes of the Papuan subregion (Diptera: Culicidae). J. med. Ent. 3: 179- 237, map. Stone, A., K. L. Knight, and H. Starcke. 1959. A synoptic catalog of the mosquitoes of the world. The Thomas Say Foundation, Ent. Soc. Amer., Vol. 6, 358 pp. Vattier, G. and J. P. Adam. 1962 (1963). Description de la nymphe d’Uranotaenia cavernicola Mattingly, 1954. Notes morphologiques et écologique sur la larve et l’adulte. Bull. Soc. Path. exot. 55: 911- 918, illus. Wattal, B. L. and N. L. Kalra. 1965. An entomological survey of Dehra Dun Valley (Uttar Pradesh). Parti II. Description of Culex (Culiciomyia) ramakrishnii sp. n. and allotype ( é ) of Uranotaenia bo (cw) PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 2 maculipleura Leicester, 1908. Bull. ind. Soc. Mal. Com. Dis. 2: 309-313, illus. Wharton, R. H. 1962. The biology of Mansonia species in relation to the transmission of filariasis in Malaya. Inst. med. Res. Fed. Malaya Bull. 11: 1-114, illus. Type Depositories 327. ANIC. Australian National Insect Collection, Canberra, Australia. BNI. Barnhard-Nocht-Instituts fiir Schiffs- und Tropenkrankheiten, Hamburg, Germany. CMC. Department of Parasitology, Chungshan Medical College, Canton. DERM. Laboratorio de Entomologiade de la Division de Endemias Rurales, Maracay, Venezuela. 329. NCID. National Institute of Communicable Diseases, Delhi, India. NMC. Naturwissenschaftlichen Museum der Coburger Landestiftung, Coburg, Germany. Australia (ANIC). 330. China (CMC, SMC), Germany (BNC, NMC), India (NICD), Vene- zuela (DERM). Index 336. cephasi—Change to cephasi. 338. decannus.—Change to deccanus. 344. josephinae.—Change to josephineae. karwari (Barraud ).—Change to karwari. 347. miachaetessus—Change to miachaetessa. 351. pennai.Change 151 to 150. 353. reptans Linnaeus. Change “Mon-mosquito” to “Non-mosquito.” 354. scutipunctatus—Change “Neivan” to “Neiva.” shintiensis—Change “Lein” to “Lien.” 355. stokesi—Change 196 to 197. Index to Supplement ITI This includes only those names not indexed in the original catalog or in Supplements I and II. The page numbers here refer to the number at the left in the preceding text, quivalent to the proper position in the original catalog. BculeatusmGolless) se eee 232 catharinensis (Lima, Guitton, and acutspalus\C@olless 2 2 282 Bierbeina’) 2. + See es ee ee 59 clamp bonabtimir tS a8. to DAO ri Ghent TAG) sete kt a oe hee 244 albosternopleura Peters 109 chiungchungensis Hsii _....___._____. 234 alpuusmC@olless* ee ee OOO. | Collesst eld ee il alticolane eters nate tee a ee /ecovacarcian MoOrattiini =a 245 amaniensis Someren and Hamon __ 225 cozumelensis Diaz Najera —-_- Wes: aMNIenSIS: Peters. 8.2 109 crassicornis Colless __.......... 232 AMMOTELSIS: Waslov a) 2 OI “icubitatus, Colless: 2.8 ee 232. antipodea Dobrotworsky _......... 221 culionicus Delfinado —..__ 231 aurata Dobrotworsky ____......... 0s clerics Collass 2 BB belkini Gasal and’ Garcia 84 dialloi Hamon and Brengues _____ 192 benoit Wolfs, acer UO) idispectuss, Bram. "...2 0 27 et) = 239 bertii Cova Garcia and Rausseo _. 110 ebogoensis Rickenbach and Ferrara 192 brekleyi Forattim 285 243 eukrines Bram and Rattanarithikul 233 224 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 explorator Marks 25 ae 145 peytoni Bram and Rattanarithikul — 235 fernandezi Casal, Garcia, and Cav- phaecasiatus Marks _..... 150 11 | ene eee ena Ve aoe 247 philippinensis Delfinado — 118 fuscosiphonis Bram and Rattanari- pholeter Bram and Rattanarithikul 235 thikul ___...__...___.._.__________._.__...... 233 pseudoandreanus Bailly-Choumara 228 gameti Bailly-Choumara 248 ramakrishnii Wattal and Kalra —__ 238 ganapathi Gollesss= e288 reidi Colless) EE 235 garioui Bailly-Choumara and FOSSIDelfina Coase 118 Rickenbachy seen 2 lee 227" ~samoaensis: Stone) =..2- = > See 104 gibbulus, Delfinado: 2 255: sapiens!’ Marks 0.500 6 eee 153 grenieri Grjebine AAW. setiten sD eltinad oye ee 213 hamomni Mato yee See 192 = shopei Forattini and Toda —_________- 280 hirsutifemoraPeters: =.) 113 © shrivastavii Wattal, Kalra and hodekinwe Wharton) 2s. 102 Koishnan. = eee 231 imperfectus Dobrotworsky — 147s simanini |\Cutsevich = 154 incomptus Bram and Rattanarithi- sinensis (Meng and Wu) __....__-- 219 kul a Fa 238 “sorsogonensis Banez and: jueco, 2 1%) inculus"@ollesste 8! ee 233 spiculosus Bram and Rattanarithi- lanei Coutinho and Forattini 280 lou 64 eer ee 235 Jashensis Nien 2 148. stanleyi-Peters.2.2 4) pla latifoliatus Delfinado 23) istrickland: Reid == 29 Jinesi Maxks"- 25 = ee 148 — subbasalis Dobrotworsky eral loreneaur eters) 68" ‘swahilicus Gillies) = 2 eee 56 lottei Hamon and Brengues GA szechtianensis Sty ee 71 lucamsa@olless) see e2 se eee 234 tabu Ramalingam and Belkin _— 188 macdonald Colless ._... +. 934 ‘tangensis Kuhlow —_... 48 macmillann, Marks - 175 tasmaniensis Dobrotworsky —- 29 malayensis @olless 22 ee 187 _ tauffliebi Rickenbach and Ferrara 198 marksae Dobrotworsky 2) eetejeran Cova Garciay = see 250 mombasaensis Mattingly 149 eaiihaiomyia Bram 2 eee 239 moresbyensis Peters __.......--- 116 tigertti Scanlon and Peyton 29 nevadensis Chapman and Barr _.. 143. traubisGolless) <.-2. 5 235 nkolbissonensis Rickenbach and tricontus Delfinado -... =. eaaoe JE GATTO VOSA: se eee seamen ce 123 tutuilae Ramalingam and Belkin _. 172 Howie wmeian fe 0 7t 2 . fs 24 “wmematus Delfinado’ = > = aeee 232 novaguinensis Peters _........______ 117 vadakadiensis Doraisamy _...... 55 novohanoverae Peters 69 variegata Dobrotworsky — 108 oswaldoi Forattini ......_.......... 252 whartoni Mattingly (Aedes) _____. 203 papuensis#leters;see = 213 whartoni Reid (Anopheles) —.__- 30 paralatexalis Peters) 118 whartoni Colless (Culex) 235 paranovaguinensis Peters Ss Swaliredi(Colless === eee 235 penmplexusseeterse — es 28 ee 72 xenophobus Ronderos 280 REPLACEMENT NAMES FOR PREOCCUPIED SPECIFIC NAMES IN DOLICHOPODIDAE (Diptera ) Medetera postminima nomen novum for Medetera minima Van Duzee, 1925, Psyche 32: 180 (California, U.S.A.), not Medetera minima (as Medeterus mini- mus) De Meijere, 1916, Tijd. Ent. 59: 259 (Java); also recorded from Sumatra by Hollis, 1964, Beaufortia 10: 260. Sympycnus pessimplex nomen novum for Sympycnus simplicipes De Meijere, 1916, Tijd. v. Ent. 59: 252 (Java), not Sympycnus simplicipes Becker, 1908, Mitteil. Zool. Mus. Berlin 4: 46 (Canary Islands, Europe). This species has recently been recorded from Sumatra by Hollis, 1964, Beaufortia 10: 269, under the De Meijere name.—Grorcr C. StTEysKAL, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington. D.C. 20560. bo Ol PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 2 SOME NEW AND OLD SPECIES OF COLASPIS IN THE WEST INDIES Doris H. BLAke, Department of Entomology, Smithsonian Institution, Washington, D.C. 20560 This paper is an attempt to straighten out the names of the blue or green species of Colaspis found in the West Indies as well as to describe several new species collected there. These rather large conspicuous beetles are of strikingly brilliant blue or green color, sometimes shot with rosy or golden lights, and so have attracted the attention of ento- mologists from the early days. Linnaeus described the first species from “America” in 1758, quite simply as “green with yellow legs and antennae’, and named it Chrysomela occidentalis. The name has been applied to a Jamaican species but the beetle was collected by Rolander, who, according to Henrikson (1928, Ent. Med. 15:97) collected not in Jamaica but in Surinam (Dutch Guiana). Later Linnaeus described another species that he named Cryptocephalus jamaicensis. This species his pupil, Fabricius, had already described a few years earlier in 1781 as Erotylus flavipes from Jamaica and his name takes prece- dence over Linnaeus’ jamaicensis. In 1792 Fabricius described Chry- somela luteicornis from the islands of “America meridionalis” and Olivier in 1808 described Colaspis unicolor and C. femoralis from the “Antilles” as well as Colaspis smaragdula from Santo Domingo. Since there is great similarity between these blue or green species of Colaspis, it is difficult to identify them from their brief old Latin descriptions except by the locality in which they were taken, and when the locality is simply “Antilles”, identification can only be made certain by extant specimens of the original beetles. A few old Fabrician specimens are still left; one, C. luteicornis, is at Kiel, and some day someone may identify it. I am unable to identify the Olivier species C. wnicolor or femoralis, or C. nigricornis Suffrian (changed by Weise to C. suffriana) or varicornis Suffrian, both from Cuba. All four species are described as having dark or partly dark antennae, and this character does not fit the blue green species of Colaspis from the West Indies except an un- described one from Jamaica. Lefévre’s description of C. chevrolati from Hispaniola and C. in- sidiosa from the “Antilles” are likewise difficult to interpret. F. Monrés spent a year studying the collections in France, and took many of his beetles there for comparison. Since I know that Monrés examined Lefevre’s collection I have taken the specimen that he labelled as C. chevrolati as probably correct. It is close to Olivier’s C. smaragdula. In the material studied I have found six species from Cuba, three species from Hispaniola, and two species from Jamaica, of these blue or green Colaspis. A single specimen has been collected in Puerto Rico, according to G. N. Wolcott, who described it briefly as “an 226 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 elongate brownish beetle with light green iridescence most marked on pronotum and head”. It was collected on “weeds in a clearing on El Yunque’. No one has been able to locate the specimen in the collec- tions in Puerto Rico, and it may possibly represent a new species. On the other hand, on the islands between Puerto Rico and South America there is a species of Colaspis which is a brownish beetle with faint green lustre that has been collected on the mainland of South America and also on the islands of St. Vincent and Grenada. It corresponds with Lefévre’s description of C. insidiosa from the “Antilles” pretty closely and it may be that this occurs as far north as Puerto Rico. Wolcott's brief description of the color of the Puerto Rico specimen matches the description of the color of C. insidiosa Lefévre. One other species of Colaspis has recently been collected on St. Lucia by O. S. Flint. It is almost piceous with a faint green or blue tint in the dried specimen and has pale yellow brown legs with the tarsi blue green. Colaspis cubensis, n. sp. Fig. 1 Between 6 and 7.5 mm. in length, oblong oval, very shiny, metallic green, rarely with a bluish lustre, coarsely and on the elytra geminately striate punctate, the intervals being slightly costate, legs and antennae yellowish brown. Head coarsely and rugosely punctate, a more or less distinct median line down occiput; shiny, brilliant green, with labrum and mouthparts yellowish brown. Antennae yellowish brown with tip of terminal joint dark piceous, very slender, extending to about middle of elytra. Prothorax with slightly angulate sides with a more or less distinct angle at about the middle, and a tooth at apical and basal angles, surface coarsely and somewhat irregularly punctate, with bare places, not very convex, brassy green. Scutellum small, shining green. Elytra coarsely and geminately striate punctate, the rows near apex becoming single and intervals on sides and at apex more costate, brilliant green, only occasionally bluish green. Body beneath shining green, legs pale yellowish brown, in male first tarsal joint of middle and anterior legs a little widened but not swollen as in other species. Length 6-7.5 mm.; width 3-4.2 mm. Type, male, and § paratypes, USNM 68322. Type locality —Cayamas, Cuba, collected by E. A. Schwarz. Other localities—Cuba: vicinity of Havana, T. Barbour; Havana, Baker; Santiago de las Vegas, on Solanum melongeum, P. Cardin; San Antonio de los Banos, Jose H. Pazos; Alquiza, on Solanum corvum, L. Scaramuzza; Jaronu, on eggplant, L. Scaramuzza; Santa Clara, R. Combs; Baragua, on eggplant, L. Scaramuzza; Nagua, Oriente Prov., S. C. Bruner; C. H. Ballou; San Blas, Santa Clara; Bahia Honda, Wick- ham; Soledad, Cienfuegos, P. J. Darlington, Jr. Loma del Gato, Cobra Range, about 3000 ft. alt., P. J. Darlington, Jr.; Banos de San Vicente, Pinar del Rio, C. Parsons; Buenos Aires, Trinidad Mts., C. Parsons; PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 227 Upper Yana Valley, L. Scaramuzza; Vinales, Pinar del Rio, W. J. Clench, P. Vaurie. Remarks.—This Cuban species has always been called C. smaragdula Oliv. ever since Suffrian identified it as that, but smaragdula was de- scribed from “St. Domingo” and the present species is confined to Cuba. It is distinguished from the species from Hispaniola in having costate elytra, and is distinguished from other species of Colaspis from the West Indies by its proportionately shorter prothorax, more gemi- nately striate punctate elytra, less swollen first tarsal joint in the male, and by the broadly rounded apex to the aedeagus that has only a very short blunt tip. Colaspis aleyonea Suffrian Fig. 13 Colaspis alcyonea Suffrian, Archiv. f. Naturg., vol. 32, 1866, p. 326. About 7 mm. in length, oblong oval, pronotum alutaceous, densely and deeply punctate with rounded sides, elytra shiny, densely, deeply and substriately punc- tate, without costae, dark blue green, blue or even purplish, legs and antennae yellow brown. Head densely and coarsely punctate with a depression in middle of front, dark blue green or even purplish blue, mouthparts yellowish brown. Antennae extend- ing almost to middle of elytra, yellow brown. Prothorax with rounded sides, sur- face alutaceous and not at all shiny, punctures deep, moderately coarse and dense. Scutellum dark. Elytra shiny, punctures deep, moderately coarse and dense. Elytra more shiny than pronotum with dense, coarse punctation tending to be striate and with some horizontal ridging, a little costate at apex. Body beneath shining blue or green, legs yellow brown, first tarsal joint of front and middle legs rotund in male. Length 6.5—7.3 mm.; width 3—-3.6 mm. Type, whereabouts unknown. Type locality —Cuba. Other localities—Cuba: Havana, Barbour; Buenos Aires and Mina Carlota, Trinidad Mts., C. Parsons, June and July 1939; Buenos Aires, P. J. Darlington, Jr., May 1936, at 2500-3500 ft. alt.; Soledad, Cien- fuegos, May 1936, P. J. Darlington, Jr. Remarks.—The deep blue or even purple coloring as well as the dense elytral punctation and lack of costae distinguish this from C. cubensis. The first tarsal joint of the male is rounded and wider than in C. cubensis. The aedeagus has an unusually long narrow tip. Colaspis orientalis, n. sp. Fig. 8 About 6.5 mm. in length, oblong oval, prothorax distinctly alutaceous and not shiny, elytra finely alutaceous but somewhat more shiny, prothorax deeply, densely and coarsely punctate, not very convex with rounded, not at all angulate sides, elytra coarsely and substriately punctate, without costate intervals, green, legs and antennae yellowish brown, in male the first tarsal joint of all legs wide. bo PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 sy (es Flavipes (Fabr.) tN 2 ©. farri nsp S. Cviridula (Suffrian) 7 C. barberi n.sp. . C.orientalis n.sp PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 229 L , 9. Colaspis darlingToni n.sp. i iS a s Se ff M™, ty (2.C. chevrolati Lefevre (Monris det) Rot auFrince Haiti 13. C. luciae n sp. I4: C.aleyonea Suffrian 230 PROC, ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Head coarsely and densely punctate, an obscure median line down occiput, surface dull, not shiny, dark green with the labrum and mouthparts deep reddish brown. Antennae not extending below middle of elytra, slender, yellow brown. Prothorax with rounded sides, not angulate, a tooth at basal and apical angles, surface alutaceous, not at all shiny, with dense, deep, and coarse punctures and in intervals very fine punctures, dark green. Scutellum dark green. Elytra a little wider than prothorax, slightly alutaceous, but more shiny than prothorax, with regular coarse and nearly striate punctures, without costae except for a slight trace at apex, deep green. Body beneath shiny dark green, legs yellow brown; in the male the anterior and middle first tarsal joint rotund and the cor- responding joint in hind feet nearly as well rounded. Length 6-6.5 mm.; width 3-3.1 mm. Type, male, MCZ 31203. Type locality —Soledad, Cienfuegos, Aug. 1920, N. Banks. Other locality —Baracoa, Oriente Province, Cuba, collected by S. C. Bruner and Leon Boucle, April 21-30, 1929. Remarks.—The dull dark green surface of the prothorax and lack of elytral costation distinguish this from C. cubensis. It is very closely re- lated to C. alcyonea Suffrian, which has similar elytral punctation but is more shiny and usually is blue or even purplish whereas this species is a dark green. The two species are difficult to separate except by comparison of the aedeagi. The aedeagus is wider and with a more slender tip in alcyonea. Only two specimens, both males, of this species are known, but in the two the aedeagi are alike and distinctly different from alcyonea. Colaspis fervida (Suffrian ) Fig. 4 Chalcophana fervida Suffrian, 1866, Archiv. f. Naturg. 32:330. Between 5 and 6 mm. in length, broadly oblong oval, lustrous green, elytra shining with golden and rosy lights, antennae and tarsal joints pale yellowish brown; prothorax with traces of angularity at sides and with scattered punctures denser on sides, elytra without costae, punctures substriate, not dense or coarse. Head usually with a slight median depression on occiput, otherwise smoothly polished and rather densely punctate, lustrous green with brown mouthparts. Antennae extending below humeri, yellow brown with the basal joint metallic green. Prothorax moderately convex with traces of angularity on sides, a strong apical tooth and smaller basal one, surface polished, lustrous green with scattered punctures, denser on sides and not coarse. Scutellum polished green. Elytra moderately convex, with small humeri and a faint trace of tranverse depression below, no signs of costate intervals, small substriate punctation, surface polished, shining with a golden or rosy sheen. Body beneath green, prosternum densely punctate, legs green except the pale yellow brown tarsal joints, sometimes these having a greenish lustre, in male inner side of hind tibiae serrate, first tarsal joint of anterior pairs of legs well rounded. Length 5-6 mm.; width 3-3.6 mm. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 231 Type, whereabouts unknown. Type locality —Cuba. Other localities —Habana, Baker; Cayamas, Schwarz. Remarks.—The rosy and golden lights of the polished green elytra make this species easily recognizable. It has much the same polished surface as C. smaragdula Oliv. from Hispaniola, but is a shorter, broader and more rounded beetle. This is one of three species of Colaspis, all of which occur in Cuba, that are shorter and more rotund than the other species of blue-green Colaspis from the West Indies. These three species bear a strong resemblance to all the species found in Hispaniola in having polished elytra with scattered and not very dense punctation and in being without costae. In all of them the sides of the prothorax show more or less degrees of angulation in spite of Suffrian’s statement that the sides are rounded. There is one peculiarity in the males of the group that the inner side of the hind tibiae is serrate. Colaspis viridula (Suffrian ) Fig. 5 Chalcophana viridula Suffrian, 1866, Archiv. f. Naturg. 32:331. About 4 mm. in length, oblong oval, lustrous green with pale yellow antennae and tarsal joints; prothorax with scattered punctures, elytra with substriate and not dense or coarse punctation, a transverse depression below basal callosity, no trace of costae. Head with a slight median depression on occiput, front without depressions, and with scattered, not dense or coarse, punctation; mouthparts dark brown. Antennae with basal joint having a faint metallic lustre, rest pale yellow except for a dark tip to terminal joint. Prothorax somewhat convex, with rounded sides, a trace of angulation along sides, a pronounced apical tooth and small basal one; surface polished green with scattered punctures, denser on sides, nowhere coarse. Scutellum shining green. Elytra without trace of costae, a tranverse depression below basal callosities, punctation tending to be striate, not coarse or dense. Body beneath shining green, prosternum densely punctate, legs green, tarsal joints yel- lowish brown with faint greenish lustre; hind tibiae in male with serrate edge on inner side and first tarsal joints of anterior pairs of legs much rounded. Length 4.2 mm.; width 2.6 mm. Type, whereabouts unknown. Type locality —Cuba. Remarks.—A single specimen from Loma del Gato, Sierra del Cobre, elevation 2600-3325 ft., Oriente Province, Cuba, was collected in Sep- tember 1935 by J. Acuna, S. C. Bruner and L. Scaramuzza which seems to fit Suffrian’s description of Chalcophana viridula. Suffrian wrote that it was smaller than C. fervida but very similar. It lacks the golden sheen of fervida being simply polished green. The coloring otherwise is much the same, and the serration on the hind tibiae in the male is the same. 232 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Colaspis barberi, n. sp. Fig. 7 About 5.5 mm. in length, oval, shining, the prothorax and head lightly and elytra more coarsely punctate, deep violaceous blue shining with greenish lights, tarsi and antennal joints except the basal one yellowish brown. Head with a faint trace of median depression on occiput, otherwise without depressions, polished, distinctly and not densely punctate, brilliant blue or purple with green lights; mouthparts brown. Antennae extending a little below humeri, slender, outer joints a little broader, basal joint with metallic blue lustre and tip of terminal joint dark, otherwise yellowish brown. Prothorax convex with sides rounded in one specimen and with a trace of angularity in other specimen, a tooth at basal and apical angles; surface brilliant dark blue with purplish or greenish lights, distinctly and not coarsely or densely punctate. Scutellum shining dark blue. Elytra broad and moderately convex, polished dark blue with purplish or greenish lustre, and with coarse and not dense punctation that tends to be irregu- larly striate, a slight depression below basal callosity. Body beneath shining dark blue, almost glabrous, legs dark blue with tibiae slightly hairy at apex, and tarsal joints a yellowish brown with faint metallic green lustre. Length 5.5 mm.; width 3.3 mm. Type, female, and one female paratype, USNM 68321. Type locality—Taken on “Musail cayes” at Baragua, Cuba, Nov. 5, 1926 by L. Scaramuzza. Remarks.—This is distinguished from C. fervida Suffrian by its deep purplish blue coloring and by the finer and not so densely punctate pronotum. No male of the species is known, so it is impossible to say whether there is serration on the hind tibiae, but it is likely since this is closely related to the group of species with that character, composed of C. fervida (Suffrian) and C. viridula (Suffrian). H. S. Barber had labelled this as a new species and set it aside to describe. Suffrian’s description of C. striata differs from this species in its smaller size, in the dark coloring of the tarsi, and in the striate punctation of the elytra. Colaspis smaragdula Olivier Figs. 10, 11 Colaspis smaragdula Olivier, 1808, Entomologie 6:883. Olivier’s description translated is as follows: Colaspis smaragdula, punctate, green, antennae and feet rufous, punctures of elytra substriate. Magnitude and form of C. aurata. Antennae pale rufous. Head punctate, green, eyes fuscous. Thorax punctate, green. Scutellum green. Elytra green with substriate punctures. Body green. Feet rufous. It has a gilded appearance. The antennae are of a pale yellowish brown, eyes are brown. All the body is of a beautiful brilliant green color, the head and thorax are punctate, the elytra are punctate, the punctures form almost geminate striae. The feet are yellowish brown. It is found in Santo Domingo. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 233 This short description easily fits all the Colaspis found on Hispaniola, as well as many elsewhere, and has led to the old Olivier name being applied pretty generally to most of the species with green coloration in the West Indies since Suffrian applied it to the Cuban species. However the habitat limits the species to Hispaniola. All the Colaspis that I have examined from Hispaniola are alike in being a brilliant polished green, without costae, except for traces near the apex, and with the punctures of the prothorax and elytra not very dense or coarse, and the elytral punctures tend to be striate or even geminate striate. Yet the aedeagi of these polished green beetles show considerable variation. In some specimens from the mountains of the central part of the Dominican Republic the aedeagi are totally different. The beetles themselves are larger with a larger prothorax. Since these beetles were collected in an inaccessible country in the interior, it is not so likely that this is the Olivier species and I am describing it as new. From the rest of the beetles I am illustrating two specimens with somewhat different aedeagi. One figure is of a specimen from Guay- mate, in the southeastern part of the Dominican Republic. The second figure is of a specimen from Source, Metalas, Haiti, which is in the low coastal region between Port-au-Prince and Ennery. Whether this variation in the aedeagi is subspecific I cannot determine as I have far too little material from which to form any conclusion. Colaspis chevrolati Lefévre fide F. Monrdés Fig. 12 Colaspis chevrolati Lefévre, 1891, Ann. Soc. Ent. Belg. 35:CCLVII. Lefévre’s description of Colaspis chevrolati translated is as follows: Colaspis chevrolati, suboblong ovate, convex, metallic green, with blue tints, shining, labrum, palpi, antennae and feet bright fulvous. Length 644-7 mm. St. Domingo. Head strongly and not densely punctate, between eyes a short longitudinal im- pression. Prothorax transverse with arcuate sides and in middle obsoletely sinuate, convex above, in middle of disc rather distantly and on sides more densely punctate with elongate (aciculatim) punctures. Scutellum smooth, apex subrotund. Elytra below humeri obsoletely transversely impressed, unevenly punctate, towards apex and especially near suture with substriate punctation, humeral callosity moderately rounded, smooth. This pretty species comes from Chevrolat’s collection where it was confused with C. smaragdula Oliv. of the same country. It resembles it in color but in form and punctation is very different. In the Monros collection are two specimens from Port-au-Prince, Haiti, that F. Monrds has identified as chevrolati Lefevre. Monrdés spent a year in Paris studying collections there and may have com- pared material with Lefevre types. The figure of the male herewith given has an aedeagus that differs only slightly from that of other specimens of the genus in Hispaniola. 234 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Colaspis darlingtoni, n. sp. Fig. 9 About 6.5 mm. in length, oblong oval, shining, prothorax and elytra distinctly but not coarsely nor densely punctate, punctures on elytra tending to be geminate striate; green or bluish green with antennae and legs yellowish brown. Head coarsely and densely punctate with an obscure median line down occiput, very shiny green with mouthparts dark reddish brown. Antennae not extending to middle of the elytra, slender, pale yellow brown with tip of terminal joint dark. Prothorax moderately convex with sides feebly angulate, especially a little below the middle, a tooth at basal and apical angles; surface shining with moderately dense, deep but not coarse punctures, green or bluish green. Scutellum shining blue or green. Elytra with a slight basal callosity and a depression below this in which punctures are a little coarser than on callosity, punctation irregularly gemi- nate striate, not very dense or coarse, without ridging between except feeble traces of costae at apex; green or bluish green. Body beneath shining blue or green or even with a purplish tinge, legs yellowish brown, first tarsal joint in anterior and middle legs roundly widened. Length 6.1-6.7 mm.; width 3.3-3.7 mm. Type, male and 2 paratypes, MCZ 31204. One paratype in the U.S. National Museum. Type locality —Foothills of the Cordillera Central, south of Santiago, Dominican Republic, June 1938, P. J. Darlington, Jr. Other locality Constanza, 3000-4000 ft., August 1938; 25 miles by road south of Puerto Plata, June 1938, P. J. Darlington, Jr. Remarks.—While the description of Colaspis smaragdula Olivier fits this species also, it does not appear so likely to be Olivier’s species be- cause it was taken in the mountains in the middle of the Dominican Republic in inaccessible country, where P. J. Darlington with his usual perspicacity found it. The aedeagus of this species is utterly un- like that of any of the rest of the species of the island. Colaspis flavipes (Fabricius ) Fig. 3 Erotylus flavipes Fabricius, 1781, Spec. Ins. 1:159. Cryptocephalus jamaicensis Gemelin ed. Linnaeus Systema Naturae, vol. 4, 1788, p. 1729. Colaspis flavipes Olivier, 1801, Entomologie 6:881. (Probably not the same as Fabricius’ species as Olivier gives the locality as French Guiana. ) Between 6 and 8 mm. in length, oblong oval, lustrous green, blue green or some- times brown with greenish lustre in the punctures or faintly green along margins, occasionally entirely brown. Legs and antennae pale yellow brown; coarsely punc- tate, punctures on elytra in single striate lines near suture, becoming semi-geminate from 5th row to side with little smooth space between lines of punctures on sides. Head at base of occiput finely punctate, punctures becoming coarser towards frontal tubercles which are impunctate, a depression in middle of front, labrum pale yellow brown. Antennae long and slender, pale yellow brown. Prothorax not twice as wide as long, rather flat, sides angulate at middle, with small tooth PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 235 at basal and apical angles; punctures irregular, not so dense in middle, contiguous on sides. Scutellum lustrous, impunctate. Elytra with short intra-humeral sulcus, a depression below scutellum in which coarse striate punctures are sunken; four rows of striate punctures from suture to middle of each elytron, more or less evenly striate with smooth spaces between rows, thence rows of striate punctures be- coming more geminate with intervening space less and more costate, on sides and at apex distinctly costate; lustrous green or in some specimens brown with green in punctures, occasionally entirely brown. Body beneath lustrous green or brown, impunctate except on prosternum, legs entirely yellow brown. Length 5.7-8.3 mm.; width 3-4 mm. Type possibly in British Museum (Natural History), “Museum D. Banks”. Type locality —Jamaica. Other localities—Manchester Parish, Mandeville, collected by Van Dyke; on Elephant’s ear and Susumbra, collected by J. M. Dale, 20 June 1950; St. Thomas Parish, 1 mile east of Lyssons, May 28, 1954, T. H. Farr; Corn Puss Gap, on Susumbra, Oct. 1, 1950, R. P. Bengry; Bath, 1937, Chapin and Blackwelder; St. Ann’s Parish; Monoagre, H. Morrison, Sept. 14, 1917; Mt. Diablo, June 13, 1965, T. H. Farr; Tre- lawny Parish, Windsor Estate, T. H. Farr, March 1960, R. P. Bengry, Aug. 30, 1955; Cockpit City, J. Maldonado, Dec. 28, 1961; Hope Gar- dens near Kingston, May 23, 1921, C. C. Gowdey; Kingston, near Liguana, on wild solanaceous plant, M. Kisluik, June 7, 1931. Remarks.—At first the entirely brown coloring of some specimens seemed to indicate a different species from C. flavipes, but further study of a series of specimens taken at one time by T. H. Farr at Mt. Diablo left no doubt that the species varies from lustrous bright green to entirely brown. The brown specimens have been taken in certain inland areas of Jamaica in Trelawny Parish and St. Ann’s Parish. Since this is the only blue-green Colaspis that occurs in Jamaica there is little doubt that the Fabrician name must be applied to it. Colaspis farri, n. sp. Fig. 2 About 5.5 mm. in length, oblong oval, lustrous dark blue with a bluish green lustre about elytral striate punctation; rugosely punctate on prothorax, lines of moderately coarse punctures on elytra, antennae and legs dark. Head with interocular space approximately half width of head, densely punc- tate throughout with a trace of median depression, shining blue green, labrum and mouthparts dark brown. Antennae extending below humeri, basal joints with a metallic bluish lustre, remainder dark brown. Prothorax slightly angulate at middle on sides with a tooth at apical angle and a less marked one at base, sur- face rugose with deepset and not too dense punctation, dark blue with purplish and greenish lustre. Scutellum dark, polished, impunctate. Elytra with regular rows of moderately coarse punctures, these rows having a blue or green lustre against the dark blue intervals, intervals towards apex becoming costate. Body 236 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 beneath dark with a bluish or greenish lustre, legs also dark. Length 5.5 mm.; width 2.9 mm. Type, female, USNM 68320. Type locality—Grove Place, Manchester Parish, Jamaica, collected 23 June 1960 by T. H. Farr. Remarks.—The small size and deep dark blue color distinguish this species. All the others treated in this paper have pale yellow legs or at least the tarsal joints are pale, and the antennae pale. This species alone has entirely dark legs and dark antennae. Colaspis insidiosa Lefévre Fig. 6 Colaspis insidiosa Lefévre, Ann. Ent. Soc. Fr. (5) VII, 1877, p. 145. Lefévre’s description of Colaspis insidiosa translated is as follows: Colaspis insidiosa, oblong ovate, fuscous aeneous, shining, body below sparsely pubescent, margins of prothorax and elytra obscurely green, labrum, antennae and legs rufous brown; head densely punctate, between eyes with a longitudinal impression, eyes large, dark, prothorax convex, unequally punctate with very minute, scarcely visible punctures in the interstices, sides with reflexed margins, rounded and widened at middle with obsolete bidentate edge; scutellum smooth, rounded at apex, elytra below base transversely strongly impressed, with prominent humeri, that are smooth. Male, smaller, elytra towards suture quite regularly subgeminate punc- tate, near sides more strongly and less regularly subrugose punctate, first joint strongly dilated in anterior tarsi. Length 714-8 mm.; latitude 34 mm. Female, longer and wider, elytra on interior disc regularly striately subgeminate punctate, towards apex and near suture quite profoundly punctate sulcate, interstices convex, smooth, near sides strongly costate, interstices tranversely rugose, first tarsal joint of anterior legs triangular. Type locality —*“Antilles”. Other localities —Venezuela: Caracas; Grenada: St. George (Lee- ward side), H. H. Smith, Becquia, H. H. Smith. St. Vincent: Yambu River Valley, on guava, M. Kisluik, C. E. Cooley; on cotton, W. N. Sands and H. A. Ballou. Mistique, on cotton leaves, W. H. Sands. Remarks.—Although this beetle was identified in the Bowditch col- lection as Colaspis fastidiosa Lefevre, I believe that it is rather Colaspis insidiosa Lefevre, the habitat of which is given as the “Antilles”. Lefeévre’s description of a brown beetle with greenish margins, and with geminate striate punctures on the elytra fits this beetle and like- wise his description of the rugose punctation and costae on the sides of the female applies to it. In the shape of the large prothorax it is unlike any other blue green Colaspis from the West Indies. Colaspis luciae, n. sp. Fig. 13 About 4.5 mm. in length, oblong oval, shining, almost piceous with a faint bluish green tint (when boiled to relax, beetle was distinctly dark bluish green), antennae PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 237 yellowish brown with last four joints darker brown; legs pale yellowish brown with tarsal joints bluish green; pronotum not so coarsely punctate as elytra, in which punctation in basal half is confused, becoming striate in apical half. Head with interocular space more than half width of head, coarsely punctate over entire head, punctures larger in lower front, a slight median depressed line down front. Antennae extending to middle of elytra, pale yellow brown with four apical joints dark brown. Prothorax with two angularities in middle of sides, and a tooth at each angle, densely and coarsely punctate. Scutellum small, rounded. Elytra with shining impunctate humeral prominences, otherwise coarsely, contigu- ously and confusedly punctate in basal half, the punctures being coarser than those on pronotum, and becoming striate in apical half with the intervals slightly costate. Body beneath dark bluish green, smooth with only the sides of prosternum punc- tate. Legs yellowish brown with tarsal joints bluish green, first joint in front and middle tarsi enlarged in male. Length 4.5 mm.; width 2.5 mm. Type, male, USNM 69217. Type locality —Marisule, St. Lucia, 30 July 1963, O. S. Flint. Remarks.—C. femoralis Olivier and C. unicolor Olivier, both from the “Antilles”, are described as bronzy, the latter even a little coppery, with antennae fuscous except at base. These descriptions do not apply to this species. There is no bronzy or coppery lustre at all to this beetle. It is almost black with a very faint blue green tint that may be more pronounced in the living beetle as it was distinctly dark blue green when boiled. It is smaller than most of the blue green species and the yellow brown legs have the tarsal joints blue green, unlike any other species from the West Indies. A SUBSTITUTE NAME FOR THE GENUS NICUESA IN COCKROACHES (DicryopTERA, BLATTARIA, BLATTELLIDAE ) In 1929 Hebard (Trans. Amer. Ent. Soc. 55: 397) described the genus Nicuesa, based on N. dichroa Hebard from Panama. It is preoccupied by Nicuesa Distant 1893 (Biol. Centr. Amer., Heteroptera 1 (Suppl.): 385) in Lygaeidae. Slater 1964 (Cat. Lygaeidae World, 1: 151) recognizes Nicuesa Distant as_ valid. No other name is available for Nicuesa Hebard, according to Princis 1965 (Orthopterorum Cat., Pars. 7: 392). Furthermore, Dr. K. Princis (Univ. Lund), in litt. 1967, agrees with the unavailability of a name. Therefore, I here propose Drabeha, new name, to replace Nicuesa Hebard. The name of the type species is Drabeha dichroa (Hebard), new combination. Drabeha is modified from the anagram of the name Hebard, as a tribute to my late friend Morgan Hebard for his many basic researches in the Blattaria. AsHLEY B. Gurney, Entomology Re- search Division, ARS, U. S. Department of Agriculture. 238 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 A NEW THYCE FROM GEORGIA (COLEOPTERA: SCARABAEIDAE ) O. L. Cartwricut, Curator of Coleoptera, Department of Entomology, U.S. National Museum, Washington, D.C. 20560 This is the first species of the genus Thyce to be recorded from east- ern United States, all others being from California, New Mexico, and Texas. It was first submitted for determination by Dr. Max R. Osburm of the U.S. Department of Agriculture pecan laboratory at Albany, Georgia, who also obtained all the additional specimens used in pre- paring the following description. I am pleased to name the species after Dr. Osburn, a fellow student and friend for over forty years. Thyce osburni Cartwright, new species Holotype male: Length 21 mm., width 10 mm. Elongate oval, dark blackish brown, finely pubescent, with pronotal midline, scutellum, inner side of sutural costae of the elytra and underside white. Labrum free, emarginate. Clypeus very broadly deeply recurved anteriorly, densely coarsely setigerously punctate, the setae rather sparse, more or less recumbent and directed toward midpoint of the distinct, sinuate, frontal suture, the punctures gradually finer and closer laterally and along the suture; remainder of head densely, coarsely punctate but with a covering of dense white recumbent setae interspersed with long erect hairs, the erect hairs about as long as the clypeus, the recumbent setae less than half as long as the erect hairs. Clypeus widest in font with broadly rounded lateral an- terior angles. Eyes large, separated by about 4 times their width. Antennae 10- segmented with 3-segmented club, the club subequal to stem, 3.5 times as long as terminal segment of maxillary palpus, and as long as basal width of clypeus. Pronotum 4.5 mm. long, 7 mm. wide, anterior angles rounded, posterior angles distinct, sides sinuate, widest at middle, the anterior half arcuate outward, pos- terior half arcuate inward; distinctly margined except medially at base where bead becomes fine and obsolete; surface densely, moderately finely punctate, the punctures with rim distinct anteriorly and obsolete posteriorly, many uniting to form lines, densely covered with fine semi-recumbent brownish hair, the hair and its puncture generally directed toward midpoint of base but some lateral hair forming a swirl around lateral fovea; narrow, white midline with dense white hair directed posteriorly; a few scattered, very long, erect brownish hairs along anterior half of lateral margin, the anterior margin, and inward slightly toward disc of pronotum at middle. Scutellum with dense recumbent white hair. Posterior edge of pronotum below beading also with dense white hair between pronotum and elytra. Elytra strongly margined laterally, somewhat depressed basally between humeri and anterior margin. Surface finely, densely covered with setigerous punctures, the fine hair as on pronotum but a trifle shorter, directed posteriorly, those on the sutural costae gradually becoming white along inner margin. Sterna with very dense long white hair as on the posterior margins of the femora. Abdominal segments and pygidium covered closely with short, recumbent white hair from dense fine punctures, the abdominal segments with a narrow PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 239 Thyce osburni, n. spi (63e) smooth shining area anteriorly at middle. Femora and tibiae with much less dense mixture of short recumbent and long erect hair. Outer claw on all tarsi with strong basal tooth, inner claw with much smaller tooth. Holotype collected May 19, 1966, in black-light trap in pecan orchard, 4% miles southeast of Albany, in Dougherty County, Georgia, by W. L. Tedders. USNM 69180. The 22 male paratypes from same locality by same collector, July 1, 240 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 1964 to May 19, 1966. Variation is negligible in the typical series, length 20 to 23 mm., width 9 to 11 mm. Thyce osburni, the only known species from eastern United States, differs from all species from western States by the very long, com- paratively narrow club of its antennae. The length of the antennal club of Thyce osburni equals the width of the head between the eyes and is nearly as long as the entire head. It is 3.5 times the length of the fourth segment of the maxillary palpus. This species is strikingly and beautifully marked as well, the white, very distinct, narrow median line of the pronotum, white scutellum, dense white hair between the pronotum and elytra and the white suture of the elytra, all against the uniform dark brown of the rest of the upper surface, presents a very handsome appearance. ANTHOMYIA PROCELLARIS RONDANI IN NORTH AMERICA (DieTERA, ANTHOMYIDAE ) Anthomyia procellaris Rondani, 1866, Atti Soc. Ital. Sci. Nat. 9: 147;—1866- 1871(?), Dipt. Ital. Prodr. 6: 154; Séguy, 1929, Encycl. Ent. (B2, Diptera) 5: 69;—1930, Mem. Soc. Sci. Nat. Maroc 24: 161; Tiensuu, 1935, Acta Soc. Faun. Flor. Fenn. 58(4): 13. Séguy figured the male genitalia of specimens he identified with this species. Tiensuu figured parts of the male genitalia of specimens from Finland and stated that at least in that country the characters of the males of A. pluvialis (L.) and A. procellaris Rond. were rather constant. It has been apparent to me for some time that a few species of Anthomyia occur in North America besides the 2 species so far recorded: A. pluvialis (L.) and A. oculifera Bigot (see Stone et al., 1965, Cat. Dipt. Am. north of Mex., U. S. Dept. Agr., Agr. Res. Surv., Agr. Handbook 276: 865). The identity of most of those species is not yet clear, but one of them agrees so well with the data of Séguy and Tiensuu concerning A. procellaris that I feel certain that the same species occurs in North America. I have examined specimens (all in USNM) from England as well as from the following North American localities: MICHIGAN; Livingston County, E. S. George Reserve, June 5, 1943 (G. Steyskal); MARYLAND: Bethesda, July 31, 1962 (C. W. Sabrosky), from puparia found in robin nest; WEST VIRGINIA: Natrium, July 3, 1961 (H. M. Kulman), from puparia found in tent of eastern tent caterpillar Malacosoma americanum (Fabr.); SOUTH CAROLINA: Clemson, December 8 and 14, 1966 (J. A. Payne), reared from pig carcass in tree—GrorcE C. STEYSKAL, Entomology Research Division, ARS, U. S. Department of Agriculture, Wash- ington, D. C. 20560. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 QAI TWO NEW METALLIC SPECIES OF HELINA FROM CHILE (DieTeRA: Muscripse )* H. Ropnery Donce, 385 Johnson Hall, Washington State University, Pullman, Washington In limited Chilean material from various sources two apparently new metallic Phaoniines have been found which seem best referred to Helina, though neither agrees fully with Malloch (1934) or other diagnoses of that genus. Helina violescens, New Species Length 7 mm. A blackish fly with thorax blue-black and abdomen blue-green. Legs black; eyes bare; squamae dark; parafacials and lunule silvery pollinose; 4 posterior dorsocentral bristles. Male.—Front 0.13 of head width; frontal rows of about 10 pairs, the upper 3 or 4 hair-like; frontal vitta velvety black, broadened to lunule; parafrontals linear, dark pollinose; ocellar bristles very strong, proclinate; verticals undiffer- entiated; parafacials narrow, silvery pollinose; eyes bare; antennae black, seg- ment 3 is 1.4 as long as 2; arista long, minutely pubescent; clypeus with antennal depressions; vibrissae slightly above oral margin; cheeks black, 0.2 of head height; occiput black-haired. Palpi black, slender. Thorax shining blue-black, with 2 faint pollinose vittae on anterior half of notum. Chaetotaxy: Acrostichals 3:1; dorsocentrals 3:4; intraalars 1:2; supraalars 1:2-3 (prealar strong); humerals 3; notopleurals 2, posterior callus 2; scutellars 3 marginal, 1 small discal, some setules laterally outside the marginal bristles; propleuron bare; prosternum bare; metasternum bare; pteropleuron and_ hypo- pleuron bare; beret hairs 1—2; sternopleurals 1:2 to 2:3; postalar declivity bare; infrasquamals none. Spiracles black. Wing hyaline, veins bare, dark brown; vein 3 ends in wing apex; vein 4 straight, posterior cross vein sinuous; costal spine vestigial; basicosta and epaulet black; squamae blackish, the margins darker; lower lobe extends 0.75 beyond upper. Halters black. Legs black. Middle tibia with 2 small posterior bristles and none on anterior side; hind tibia lacks calcar, but with a very long, erect PD at apical fourth and 2 AD, 2 AV bristles. Abdomen shining, blue-green, with scattered erect bristles on the sides of third and disc of fourth segments; venter black-haired; 1st sternum bare. Fifth sternum with a small notch; genital segments small, retracted. Female.—Similar to male in most respects. Front slightly narrowed to vertex, which is 0.33 of head width; ocellars more divergent; ocellar triangle large, reaching half way to lunule; inner and outer verticals well developed; orbital bristles absent; frontal rows of about 8 pair of bristles; parafrontals setulose; para- facials glabrous on upper half; costal spine longer, double in allotype, vestigial in most paratypes; tibial bristles identical to the male. Type Series—Holotype male, Las Condes, Santiago Prov., Chile, Oct. 6-12, 1958 (Luis Peas); Allotype and two male, one female para- 1 Scientific paper No. 2621, Washington Agricultural Experiment Station; Project No. 9043. 249 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 types, “Concep(cion), 1.Oct.(19)55, J. A. Coll.”; 1 female, Quintero, Oct. 15, 1951 (Maria Etcheverry ); 2 females, Angol, Chile, Nov. 9 and Sept. 19, 1950 (D. Cataldo); 1 female, Quillota, Valparaiso Prov., Chile, Dec. 30, 1954 (Luis Pefias); 5 males, Barrancas, Santiago, Chile, Mar. 14, 1955 (Luis Pefas); 2 males, 21 females, Dalcahue, Chiloe Prov., Chile, lat. 42°23’, Feb. 10-20, 1957 (Luis Peas); one male, Maullin, Chile. Holotype, allotype and paratypes at Washington State University; paratypes to be sent to Prof. Jorge Artigas, Univ. of Concepcion; Prof. Maria Etchevvery and the U. S. National Museum. This species keys to Helina (part) in Couplet 5b of Snyder (1940) because the hypopleuron is bare; however the presence of setules about the posterior notopleural bristle and on the outer margin of the scutellum would exclude it from that group in couplet 4. In Malloch’s key (1934) it runs to viola Malloch, but differs by the bare eyes, hyaline wings and smaller size. If the PD bristle of the hind tibia is interpreted to be the calcar, this species runs to Euphaonia ful- vohumeralis Malloch, which is obviously different. It seems better to refer it to Helina, at least for the present. Three of the five Barrancas males have the wings deeply yellowed but do not appear to differ otherwise. Some specimens may have three pairs of postsutural acrostichals, but only the prescutellar pair is strong. Helina veniseta, New Species Length 7 mm. Superficially very similar to the preceeding species but lunule and parafacials with blackish pollen, cruciate interfrontals present, acrostichals 2:3, all very strong, notopleuron with scattered setules about both bristles and vein 3 setulose. Female.—Head black; eyes extremely sparsely and minutely spiculose. Front parallel-sided, 0.33 of head width; frontal rows of 9-10 pair, the upper 2 pair reclinate; frontal vitta broad, velvety black, with a pair of cruciate interfrontals and the ocellar triangle reaching about 0.33 to lunule from the anterior ocellus; ocellar and vertical bristles very strong; no orbitals; parafrontals narrow, with a row of black setules; parafacials wider, polished on upper half, blackish pollinose below; antennae black, segment 3 is 1.4 as long as segment 2; arista very minutely pubescent to tip; vibrissa as in preceeding species; cheek 0.25 of head height; cheek, metacephalon and occiput entirely black-haired; palpi black, slightly swollen; haustellum short. The antennal axis is 0.64 and vibrissal axis 0.63 of the head height. Thorax bluish-black, faintly dusted along front margin; acrostichals 2:3; dorso- centrals 2:4; intraalars 1:2; supraalars 1:3, the prealar subequal to the posterior notopleural; humerals 3; notopleurals 2 plus many setules; posterior callus 2; scutellars 3 strong marginals and a weaker basal, 2-3 discals; propleuron and prosternum bare; sternopleurals 2:2; pteropleuron setuled on posterior half; hypo- pleuron setuled before the spiracle, bare below; beret hairs present; infrasquamals absent. Posterior spiracle large, oval, without bristles in lower margin; supra- spiracular convexity equal in size and shape to the spiracle, pilose. Legs black; PROC. ENT. SOC. WASH., VOL. 69, No. 3, SEPTEMBER, 1967 243 hind tibia with erect PD at apical fifth and 2 AD and 2-3 AV at about middle; hind femur with complete AD and AV rows; middle tibia with 3 P, no PV bristles; middle femur with a complete PV row, the basal 2-3 stout and erect; anterior tibia with only one outstanding bristle, dorsal and subapical. Wing hyaline, veins dark; vein 3 with dorsal setules at base, extending half way to the cross vein, ventrally bare on node, followed by 3 small setules, visible on one wing only; vein 3 apically rather strongly bowed; vein 4 straight on apical section; remigium bare; costal spine vestigial; basicosta and epaulet black; squama blackish with black margins, the lower lobe subtriangularly rounded, projecting about halfway beyond the upper lobe; halters black. Abdomen shining blue, the 4th segment concolorous, with scattered, erect hairs. Venter black-haired, the first sternum bare. Male.—Unknown. Holotype.—Female, Las Condes, Santiago Prov., Chile, Oct. 6-12, 1958 (Luis Penas), at Washington State University. The setulose vein 3 would exclude this species from Helina. How- ever, in Malloch’s key it does not fully agree with Myiospila because vein 4 is straight nor with Hydrotaea because of the absence of a proclinate orbital bristle, so it would key falteringly to Helina. Its resemblance to violescens may be more than superficial. In Curran’s (1934, 1965) Manual it runs to Mydaea, but it is excluded from that genus by the presence of anterior acrostichal and female interfrontal bristles. If keyed past Helina in Malloch it differs from the metallic Darwinomyia by the eye not conspicuously pilose, the presence of hypopleural hairs and absence of PV bristles on the middle tibia; from Euphaonia fulvohumeralis Malloch by the clavate palpi, from Psilo- chaeta by the presence of cruciate interfrontals in the female and from Brachygasterina by the hypopleural hairs. Furthermore, all of these genera are stated in Malloch’s key to have vein 3 bare. Helina violescens and veniseta do not agree with any of the metal- lic Muscidae described from Argentina by Shannon and del Ponte (1926). In Albuquerque’s (1952) key to metallic Neotropical Phaoni- inae they run best to Brachygasterina and differ from Xenomorellia Malloch and Dasymorellia Malloch by the remigium bare. LITERATURE CITED Albuquerque, D. de O. 1952. Sobre alguns generos metalicos de Phaoninae neotropicais e descrigao de uma Xenomorellia nova (Diptera: Musicidae). Bol. Mus. Nac. (Zool.) no. 116: 1—(21). Curran, C. H. 1934, 1965. The families and genera of North American Diptera. Ist ed., 1934, priv. publ., 512 pp.; 2nd ed., 1965, Henry Tripp, N. Y., 515 pp. Malloch, J. R. 1934. Muscidae. Dipt. Patagonia and S. Chile (Brit. Mus. [Nat. Hist.]), pt. 7, fasc. 2: 171-346. Shannon, R. C., and E. Del Ponte. 1926. Sinopsis parcial de los Muscideos Argentinos. Rev. Inst. Bacteriol. (Dept. Nac. Hig.) 4: (1)-44, pls. 1-2. Snyder, F. C. 1940. New and little known tropical Helina Desvoidy, allied to Ariciella Malloch (Diptera: Muscidae). Am. Mus. Novitates no. 1060: 1-9. 244 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 SYNONYMY IN THE LYGAEIDAE ( HEMIPTERA )* James A. SLATER, Department of Zoology and Entomology, University of Connecticut, Storrs, Connecticut Syzygitis reflexa Bergroth, 1921. Ann. Soc. Ent. Belg. 61:165. The taxonomic position of this monotypic genus has been an enigma for many years. Bergroth related it to Polychisme and Scudder (1962. Trans. Roy. Ent. Soc. Lond. 114:193) suggested that the two genera were “most probably” congeneric. In the Helsinki Museum is a female of Polychisme poecilus (Spinola ) which certainly is the type specimen of Syzygitis reflexa. It fits the Bergroth description in every particular and the locality data (“Chile, Concepe. (P. Herbst)” ) is identical. A holotype label has been placed on this specimen. Syzygitis is here formally placed as a junior synonym of Polychisme and S. reflexa a junior synonym of P. poecilus. New synonymy. The key to the two species of Polychisme given by Scudder (1962) is inaccurate as in a series of thirty-two specimens of poecilus from Chile the variation in number of corial punctures covers the entire range of numbers used by Scudder to differentiate poecilus from the Central American ferruginosus (Stal). P. poecilus, like most other ischnorhynchines, is usually found in the macropterous condition. I have in my collection, however, a series of six specimens that I take to be conspecific with poecilus that are strongly coleopteroid, with the clavus and corium distinct and the lat- ter strongly broadened and rounded laterally, more convex dorsally, and with the apical corial angle extending back to the antero-lateral corner of abdominal tergum seven. The membrane is thus correspond- ingly small, although attaining the apex of the abdomen. Such coleop- teroid conditions are frequently associated with relatively stable habitats and/or old stable land areas, and in the present case would seem to have great zoogeographic significance as these insects ap- parently are associated with Nothophagus in Chile. Lethaeus simpsoni Bergroth, 1912. Ann. Mag. Nat. Hist. (8):10:195. This species was originally described from “Badagri, Southern Nigeria (J. J. Simpson).” As usual Bergroth did not indicate type specimens or so label his material. However, the Helsinki Museum possesses a male specimen with the above locality data plus “1910-213., 1-2-10”, and the scientific name in Bergroth’s handwriting. This is cer- 1 This work was supported by a grant in aid from the National Science Founda- tion. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 245 tainly the type specimen and a red holotype label has been placed on the pin. L. simpsoni, despite Bergroth’s statement that it is “very distinct in several characters” appears to me to be indistinguishable from Lethaeus africanus Dallas, and is here formally synonymized. New synonymy. Apteroloides Slater, 1964. South African Animal Life 10:60-61. Unfortunately the five year period that the above paper was in press has resulted in a case of synonymy. The genus Apteroloides was de- scribed with Aspilocoryphus signatus Distant 1918 as type species and also included Apterola angusticollis Lindberg 1939. Seidenstucker (1964. Reichenbachia 2:56:199-207) described the genus Stenaptula with Apterola angusticollis Lindberg as type species. Since Seidenstuck- er’s paper appeared in January of 1964 it has priority and Apteroloides Slater becomes a junior synonym of Stenaptula Seidenstucker. New synonymy. Blissus obscurus Slater, 1964. South African Animal Life 10:102. This specific name is a junior homonym of Blissus doriae obscurus Reuter (1888. Rev. d'Ent. 7:223) described from Greece. I propose, therefore, the name Blissus latoides new name to replace Blissus ob- scurus Slater, preoccupied. Lasiosomus bellus Slater, 1964. South African Animal Life 10:159. Again the press time of my South African paper appears to have resulted in a case of synonymy. L. bellus had originally appeared in my manuscript as L. minutus, but was changed to bellus in the galley (by oversight it still appeared as minutus in the key) due to the appearance of the description of a Lasiosomus minutus by Scudder (1962. Mus. Roy. Afr. Cent. Sci. Zool. N. 110, p. 423). It now appears to me that bellus is actually synonymous with Lasiosomus minutus Scudder and the former becomes a junior synonym. New synonymy. Naphiellus Scudder, 1962. Can. Ent. 94:982. This genus was described by Scudder for Aphanus latus Distant from India. Slater (1964. S. Afr. Animal Life 10:189) synonymized latus with Rhyparochromus dilutus Horvath 1892. Therefore the cor- rect name for the taxon should be Naphiellus dilutus (Horvath). Pachymerus crudelis Haglund, 1895. Ofv. Vet. Akad. Forh. 7:462-3. This species was originally described by Haglund in the omnibus genus Pachymerus and subsequently placed in Lachnophoroides by Distant (1918. Ann. Mag. Nat. Hist. (9):2:260). However, this latter genus is said by Distant 1920 to pertain to the Oxycareninae. (I have not seen the type species L. ornatipennis Distant 1914 from New Cale- donia.) Crudelis Haglund is a rhyparochromine and belongs to the genus Dieuches. 246 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 THE NEARCTIC DORYCTINAE, IV. NOTES ON THE GENUS DENDROSOTER WESMAEL AND DESCRIPTION OF ONE NEW SPECIES (HYMENOPTERA, BRACONIDAE ) Paut M. Marsu, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560 Muesebeck (1938) revised the genus Dendrosoter for the United States, discussing generic synonymy and providing a key to the four known species. The present paper provides additional distribution records for two species and a description of a new one from the south- western United States. The records listed below are in addition to those found in the Hymenoptera Catalogue (Muesebeck and Walkley, 1951) and its two supplements (Muesebeck, 1958 and 1967). Dendrosoter protuberans (Nees ) Bracon protuberans Nees, 1834, p. 121. This species is being studied in Michigan and Ohio as a potential control agent for Scolytus multistriatus (Marsham), the smaller Euro- pean elm bark beetle. It is listed here in the likely event that it will become established in the United States. In Muesebeck’s key (1938, p. 282), protuberans will run to couplet 3 and agrees most closely with integer Muesebeck. However, the males of protuberans are easily rec- ognized by their swollen third segment of the maxillary palpus and by the short stigma in the hindwing (fig. 1); the females are distinguished by the shorter malar space which is less than one-half of the eye height. Dendrosoter suleatus Muesebeck Dendrosoter sulcatus Muesebeck, 1938, p. 284. I have seen specimens from Georgia, Louisiana, Mississippi, and New York. This species has previously been recorded from Florida, North Carolina, Virginia, Washington, D.C., Wisconsin, and Texas. The range of sulcatus is probably throughout the states east of the Great Plains. Dendrosoter chansleri Marsh, n. sp. Female.—Length 4.5 mm. Color dark brown except calli on head and basal antennal segments which are lighter brown. Head cubicle; frons deeply excavated between calli; temples and malar space equal to eye height; face, temples, and vertex finely rugulopunctate, frons rugolose with a weak median longitudinal carina from antennae to median ocellus; calli strongly rugose; antennae 25 seg- mented, inner edge of first and second flagellar segments concave, length of first flagellar segment nearly equal to second. Mesonotal lobes granular; notauli deep, meeting before scutellar furrow in a wide coarsely rugose area; mesopleural disc PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 QA7 Fig. 1. Dendrosoter protuberans (Nees), fore and hindwings, ¢. Fig. 2. D. chansleri Marsh, n. sp., fore and hindwings, °. finely reticulate; mesopleural furrow shallow, foveolate, slightly longer than one-half width of mesopleuron; scutellar furrow deep, with numerous cross carinae; scutellar disc granular; propodeum entirely rugose, carinae definite only on apical edge. Wings maculated, venation as in fig. 2. Hind coxae coarsely striate. First abdominal tergum as long as apical width, longitudinally striate, without a definite raised median area; tergum (2+ 3) longitudinally striate on basal one-third; re- mainder of abdominal terga smooth; ovipositor a little longer than abdomen. Male.—Unknown. Holotype Female—NEW MEXICO: Lincoln National Forest, J. F. Chansler, Hopk. U.S. 37240-G, Pinus ponderosa, in association with Dendroctonus adjunctus. USNM 68927. Paratypes.—3 ? ?, same data as holotype; all deposited in U.S. National Museum. This species runs to integer in Muesebeck’s key but differs in its larger size, color, coarser sculpturing, larger rugose area where notauli meet, deeper notauli, wing maculations rather than cross bands, more convex scutellum, and lack of definite carinae on the propodeum. REFERENCES Muesebeck, C. F. W. 1938. The genus Dendrosoter Wesmael in the United States. Proc. Ent. Soc. Wash. 40:281—287. 1958. Family Braconidae. in K. V. Krombein, Hymenoptera of America North of Mexico. Synoptic Catalogue. First Supplement. U.S. Dept. Agr., Agr. Monogr. 2:18-36. 1967. Family Braconidae. in K. V. Krombein, Hymenoptera of America North of Mexico. Synoptic Catalogue. Second Supplement. U.S. Dept. Agr., Agr. Monogr. 2:27-60. Muesebeck, C. F. W. and L. M. Walkley. 1951. Family Braconidae. in C. F. W. Muesebeck et al., Hymenoptera of America North of Mexico. Synoptic Cata- logue. U.S. Dept. Agr., Agr. Monogr. 2:90-184. Nees ab Esenbeck, C. G. 1834. Hymenopterorum Ichneumonibus affinium monographiae. Stuttgart, Tiibingen, Cotta 1: 1-320. 248 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 ADDITIONAL SYNONYMIES WITHIN THE AMBLYCERAN BIRD LICE (MALLOPHAGA )* Rocer D. Price® and K. C. EMERsSON® During the past year a number of new synonymies within the bird lice have come to our attention. These are as follows: Colpocephalum aculeatum Piaget, 1885, Pediculines Suppl.: 121. Colpocephalum olivaceae Price and Beer, 1965, Ann. Ent. Soc. Amer. 58: 119. New synonymy. Piaget’s species was based on a 6 ( Piaget Collection slide 1121 at the British Museum (Natural History) ) supposedly from a specimen of Columba iriditorques Cassin in the Museum de Leide. Subsequently, Hopkins and Clay (1952) stated that C. aculeatum was actually from some member of the Psittaciformes and they placed it in Psittacomeno- pon. Recent study has shown that C. aculeatum and C. olivaceae are conspecific, with the true host probably being Lampribis olivacea, a ciconiiform and the type-host of C. olivaceae. Coincidentally, Piaget (1885) also described Laemobothrion pallidum from the same host species as that for C. olivaceae, to substantiate that he had material available from that host. Colpocephalum angolensis Price and Beer, 1963, Can. Ent. 95: 750. Colpocephalum angolensis Tendeiro, 1964, Ann. Mus. Roy. l’Afr. Cent., Ser. 8, No. 132: 171. New synonymy. This is a situation in which the same name was given in 2 separate descriptions for what is obviously the same species of louse. Colpocephalum heterosoma Piaget, 1880, Pediculines: 572. Colpocephalum poopoensis Carriker, 1956, Rev. Brasil. Ent. 5: 140. New synon- ymy. Carriker (1956) described both C. heterosoma boliviana and C. poo- poensis from specimens of a single series taken from Phoenicopterus chilensis Molina. Price and Beer (1965a), without examination of the types, were able to determine that C. heterosoma boliviana was a synonym; and that C. poopoensis was also probably a synonym of C. heterosoma. We have now studied Carriker’s type-series of both, in- cluding the holotype é of each (USNM 68655 and 68656, respectively ), and believe they are morphologically inseparable from C. heterosoma found on P. antiquorum Temminck. Clay (1951) has pointed out the 1 Paper No. 6106, Scientific Journal Series, Minnesota Agricultural Experiment Station, St. Paul, Minnesota 55101. 2 Department of Entomology, Fisheries, and Wildlife, University of Minnesota, St. Paul, Minnesota 55101. ° 2704 North Kensington St., Arlington, Virginia 22207. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 249 great amount of variation within ¢ ¢ of C. heterosoma; Carriker’s holo- type of C. heterosoma boliviana is a specimen of the “small” form and his holotype of C. poopoensis is one of the “large” form. To add further to his confusion, Carriker included both a “small” and a “large” form within his paratypes of C. poopoensis. The é genitalia as illustrated by Carriker (1956: Figs. 24, 26, 27), and upon which he placed consider- able importance, are actually inseparable, the differences as shown either being due to his interpretation or to an oversight of parts. Colpocephalum nanum Piaget, 1890, Tijdschr. Ent. 33: 257. Neocolpocephalum buteonis Eichler, 1954, in Bach and Eichler, Monatsch. f. Veterinarmed. 9: 13. New synonymy. Price and Beer (1963) placed N. buteonis as a species sedis incertae due to its unrecognizable description. We recently obtained from the Zoologisches Museum of Hamburg 3 ¢ paratypes of N. buteonis (WEC 1748a, WEC 4072ab, WEC 4072v), all of which are typical C. nanum. The illustration of the dorsal ? terminalia by Eichler (Bach and Eichler, 1954: Fig. 1) is misleading and must either represent a misinterpreta- tion of certain details or possibly a distorted specimen. Since these 3 2 Qare the only specimens of the type-series known to be available for study, since they bear the same collection data as the holotype, and since they are from Buteo buteo (L.), a bird commonly infested with C. nanum, we can only conclude that N. buteonis is a junior synonym of C. nanum. Colpocephalum napiforme Rudow, 1869, Z. Naturwiss. 34: 395. Colpocephalum heterospizium Carriker, 1963, Mem. Soc. Cien. Natur. La Salle 23: 15. New synonymy. A @ paratype from the same collection as the holotype @ is inseparable from C. napiforme; the illustration of the 4 genitalia by Carriker (1963: Plate III, Fig. 3a) may likewise be construed as being similar to those of C. napiforme. Colpocephalum pectinatum Osborn, 1902, Ohio Nat. 2: 201. Colpocephalum ictiniae Carriker, 1963, Mem. Soc. Cien. Natur. La Salle 23: 13. New synonymy. The type-series of C. ictiniae composed of the ¢ holotype (USNM 68759), ¢ allotype, and 3 2 paratypes supposedly from a single collec- tion from Ictinia plumbea (Gmelin), a falconiform, are all typical owl Colpocephalum and compare favorably with material we have seen of C. pectinatum. Ciconiphilus decimfasciatus (Boisduval and Lacordaire, 1835), Faune Ent. En- virons Paris: 123. Ciconiphilus pilherodii Carriker, 1964, Rev. Brasil. Biol. 24: 102. New synonymy. Ciconiphilus agami Carriker, 1964, Rev. Brasil. Biol. 24: 103. New synonymy. 250 PROC. ENT. SOC. WASH., VOL. 69, No. 3, SEPTEMBER, 1967 A study of the 2 holotype (USNM 68869), 4 allotype, and a @ para- type of C. pilherodii and the ¢ holotype (USNM 68870) and ¢ allotype of C. agami has shown both series to agree well with specimens of C. decimfasciatus as delimited by Price and Beer (1965b). Ciconiphilus quadripustulatus (Burmeister, 1838), Handb. Ent. 2: 438. Colpocephalum sphenorhynchus Tendeiro, 1964, Ann. Mus. Roy. l’Afr. Cent., Ser. 8, No. 132: 173. New synonymy. Specimens from Sphenorhynchus abdimii (Lichtenstein), the type- host of C. sphenorhynchus, have been examined previously (Price and Beer, 1965b) and found to be conspecific with C. quadripustulatus. These specimens agree well with the description of C. sphenorhynchus. Cuculiphilus (Aegypiphilus) gypsis Eichler, 1944, Dtsch. Ent. Z. 1943: 57. Aegypiphilus contrastus Eichler and Zlotorzycka, 1963, Acta Parasitol. Polonica 11: 216. New synonymy. Aegypiphilus secundus Eichler and Zlotorzycka, 1963, Acta Parasitol. Polonica 11: 217. New synonymy. Scharf and Price (1965) discussed their reluctance to render an opinion on these names in view of material seen to that time. Since then, however, we have obtained for study the holotype ? of A. con- trastus and the holotype é of A. secundus. We have found no signifi- cant differences between the ¢ holotype and only specimen known to date of C. gypsis and ¢ ¢ of A. contrastus and no significant differences between the ¢ holotype and only specimen known to date of A. secundus and é @ of A. contrastus. Eichler and Zlotorzycka (1963) unfortunately provided no adequate separating characteristics for these lice; as a re- sult, we feel there is now no longer justification for recognizing these as separate species. Kurodaia caputonis (Carriker, 1966), Amer. Midl. Nat. 76: 77. Conciella clamatori Carriker, 1966, Amer. Midl. Nat. 76: 78. New synonymy. Conciella setosa Carriker, 1966, Amer. Midl. Nat. 76: 79. New synonymy. Conciella glaucidiae Carriker, 1966, Amer. Midl. Nat. 76: 79. New synonymy. Carriker (1966) described the above 4 species of Conciella consecu- tively. We have studied his type-series, including the holotype of each, and can find no means for separating them. Presumed differences cited in the descriptions are attributable primarily to distortions in the handling of the specimens. These specimens are extremely close to Kurodaia crassiceps (Piaget), and may eventually prove inseparable, but K. caputonis, with dimensions generally slightly smaller and with a narrower, more clearly defined, somewhat pointed penis, is main- tained here as a distinct species, with page priority over the 3 junior synonyms. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 251 Laemobothrion maximum (Scopoli, 1763), Ent. Carniolica: 382. Laemobothrion (Laemobothrion) grandiculus Tendeiro, 1964, Ann. Mus. Roy. l Afr. Cent., Ser. 8, No. 132: 185. New synonymy. No reliable difference has been demonstrated between specimens of Laemobothrion from Buteo rufofuscus (J. R. Forster), the type-host of L. grandiculus, and other series considered to represent L. maximum (see Nelson and Price, 1965). REFERENCES Bach, G. and W. Eichler. 1954. Federlinge in Federspulen. Monatsch. f. Vet- erinarmed. 9: 12-14. Carriker, M. A., Jr. 1956. Neotropical Mallophaga miscellany, No. 9. A new genus and species. Rev. Brasil. Ent. 5: 111-146. 1963. New and little known Mallophaga from Venezuelan birds (Part II). Mem. Soc. Cien. Natur. La Salle 23: 5—42. 1966. New species and records of Mallophaga (Insecta) from Neotropical owls( Strigiformes ). Amer. Mid]. Nat. 76: 74~99. Clay, T. 1951. Systematic notes on the Piaget collections of Mallophaga.—Part III. Ann. Mag. Nat. Hist. (12) 4: 1159-1168. Eichler, W. and J. Zlotorzycka. 1963. Studien ber Raubvogelfederlinge. IV. Bemerkenswerte Colpocephalidae von Geiern (Vulturidae und Aegypiidae). Acta Parasitol. Polonica 11: 199-221. Hopkins, G. H. E. and T. Clay. 1952. A Check List of the Genera & Species of Mallophaga. London: Brit. Mus. ( Nat. Hist. ), 362 p. Nelson, R. C. and R. D. Price. 1965. The Laemobothrion (Mallophaga: Lae- mobothriidae) of the Falconiformes. J. Med. Ent. 2: 249-257. Piaget, E. 1885. Les Pediculines, Essai Monographique, Supplement. Leide: E. J. Brill, xii + 200 p. Price, R. D. and J. R. Beer. 1963. Species of Colpocephalum (Mallophaga: Menoponidae ) parasitic upon the Falconiformes. Can. Ent. 95: 731-763. 1965a. The Colpocephalum (Mallophaga: Menoponidae) of the Ciconiiformes. Ann. Ent. Soc. Amer. 58: 111-131. 1965b. A review of Ciconiphilus Bedford (Mallophaga: Menoponi- dae). Can. Ent. 97: 657-666. Scharf, W. C. and R. D. Price. 1965. A taxonomic study of the genus Cuculi- philus (Mallophaga: Menoponidae). Ann. Ent. Soc. Amer. 58: 546-555. 252, PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 NEOPARENTIA, A NEW GENUS OF AMERICAN DOLICHOPODIDAE (DirTEra ) Haroip Rosinson, Dept. of Botany, Smithsonian Institution, Washington, D.C. During the century since the recognition of Sympycnus Loew, the genus has been relatively ill-defined and has been easily confused with other genera in various subfamilies such as Symbolia (Diaphorinae), Stolidosoma (Stolidosominae), and Peloropeodes (Rhaphiinae). The most critical efforts have left a large and variable genus, and attempts at further subdivision have depended excessively on characters limited to the male sex. Only recently it has been possible to clarify the status of one very natural and uniform segregate genus, Calyxochaetus, which differs from Sympycnus by the lack of hairs on the scutellum (Robin- son, 1964). Collecting in southern Mexico I have found a number of species of Sympycninae which lack hairs on the scutellum but otherwise in no way resemble Calyxochaetus. These species all possess long thread- like hypopygial lamellae, a character that led Parent (1954) to de- scribe a related species from Costa Rica as a Nematoproctus. For these species I propose the recognition of the following new genus. Neoparentia, gen. nov. Setae mostly black. Face very narrow, usually with eyes contiguous near the middle in the male; front broad, broader above, mostly metallic greenish or bluish; palpi small; proboscis brownish; antennal segment 1 bare above, segment 2 trun- cate apically, segment 3 with arista dorsal; lower orbital setae uniseriate, pale. Mesoscutum not flattened posteriorly; acrostichals uniseriate; 6 pairs of dorso- centrals; 2 scutellars, scutellum without hairs; proepisternum with only a few pale setae above. Middle and hind coxae with an external bristle; mid femur with both anterior and posterior preapical; hind femur with 1 or 2 preapicals anteriorly; front metatarsus of male not shortened. Wing elongate oval: veins 3 and 4 nearly straight and parallel to noticeably divergent beyond the crossvein; last part of vein 5 distinctly longer than crossvein; vein 6 represented by slight fold. Abdomen cylindrical in male, 6 tergites visible; hypopygium small usually rather globose, placed on tip of abdomen; lamellae filliform. Type species, Neoparentia bisetosa sp. n. The various hypopygia of Neoparentia are highly divergent in form but in no case show the conical shape that is typical of Calyxochaetus. Neither are the hypopygial lamellae short and inconspicuous as in that genus. In addition, Calyxochaetus usually has a shining violet front (obscured by pollen in C. nodatus), and usually has the front meta- tarsus of the male shortened (not shortened in C. metatarsalis). The face of Neoparentia is narrower in both sexes, % as wide below as high in the female compared % or more as wide below as high in the female of Calyxochaetus. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 255 The new genus contains two rather different elements, the smaller species showing little superficial resemblance to the larger ones. Ap- parently Neoparentia is itself a complex showing considerable evolu- tionary divergence. KEY TO THE SPECIES OF Neoparentia 1. Abdominal tergites 2 and 3 yellow on the side; sternite 5 sometimes with a pair of stout rodlike setae; hypopygial capsule forming a pair of long slender projections on the forward end; length usually 3 mm or more —__ 2 Abdominal tergites wholly dark, sternite 5 without rodlike setae; hypopygial capsule blunt or with only short projections on the forward end; length Fesshthiameoerninn 2 eee woes ctr val gener ere Rune ere Vek TE eRe ee oe Bate ae! 4 2. Metepimeron yellow; abdominal sternite 5 without large rodlike setae (Costa [RUG aces BS pt ee RS ed GP all, oA Ripe oe RE Rem Dyleie Peele SS bicolor (Parent ) Metepimeron dark; abdominal sternite 5 of male with large rodlike setae —__ 3 3. Length 2-3 mm; second joint of male fore tarsus somewhat thickened QU@ ali Oriel) ee ae ee ere ee ee caudata (Van Duzee) Length 3-3.5 mm; fore tarsus of male plain (Oaxaca, Chiapas) aS ESI TE Ere ee Ree ld RY Daye Se, SN Se ES _____ bisetosa sp. n. 4. Palpus white; acrostichals sparse or lacking posteriorly; wing of male with tip deformed, veins 3 and 4 strongly diverging apically (Oaxaca, Chia- [04 (Si) eae ae ee ne a SR nn Oe NS oe deformis sp. n. Palpus dark; acrostichals distinctly reaching onto posterior slope; wing of male not greatly deformed, veins 3 and 4 hardly diverging apically 5 5. Hind tibia with two anterodorsals; male fore tarsus with last joints modified, usually folded against each other (Oaxaca, Chiapas) tarsalis sp. n. Hind tibia with only one anterodorsal; fore tarsus plain 6 6. Lower pleural surface yellow; wing veins 3 and 4 slightly diverging apically ((@ostagRicar) sae ten 2. Be, Pere Te See Ae ee le ee schildi sp. n. Pleura totally dark; wing veins 3 and 4 parallel apically (Oaxaca, Chia- PES) ere aes a AS as Ne Rg el Ne obseura sp. n. Neoparentia bisetosa sp. n. (Figs. 1, 2) Male.—Length 3.4 mm; wing 3.2 mm by 1.1 mm. Face long and narrow with eyes nearly contiguous in lower half, gradually wider above, covered with yellowish-white pollen; front metallic bluish with slight yel- lowish pollen. Palpus pale. Antenna brown, first and part of second segment yellow, first rather narrow; second segment very short; third segment about as long as wide, rounded apically; arista basal. Mesonotum metallic greenish dulled with yellow pollen, darker lines following rows of bristles, grayer pollen on sides; metepimeron mostly darkened, pale below. Acrostichals reaching onto posterior slope. Legs mostly yellow; distal joints of tarsi, tip of hind femur and tibia, and all of hind tarsus dark; setae mostly black, pale setae anteriorly on fore and middle coxae and ventrally on femora. Middle femur with a row of short erect black setae near base on posteroventral surface; middle tibia with large anterodorsal and postero- dorsal near basal fourth, anterodorsal and crest of about eight ventrals near distal 254 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 co t rh we av oo 1 A nears “ttre "OIA pe TT OTR AAA MaATANNTATEN ii lr 2 a (\ [ ( A Trp, (tn; eit) MITT THT TUT IMAT RT TATNTTIT AT Trem MO 6 ny < Ty a it Atty PrpaganntTTIATN TO 10 Figs. 1-11, Neoparentia, males. Figs. 1-2, bisetosa sp. n.: 1, wing; 2, hypo- pygium. Fig. 3, bicolor (Parent), hypopygium (after Parent, 1954). Figs. 4-5, deformis sp. n.: 4, hypopygium; 5, wing. Fig. 6, obscura sp. n., hypopygium. Figs. 7-10, tarsalis sp. n.: 7—8, fore tarsi; 9, hypopygium; 10, wing. Fig. 11, schildi sp. n., hypopygium. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 255 third, one subapical ventrally, two or three apicals; hind tibia with two antero- dorsals, five or more posterodorsals, five or more indistinct ventrals, four apicals. Lengths of joints of fore tarsus from base as 9-5-4-3-2; middle tarsus as 12-7-6-4-3; hind tarsus as 8-12-8-5-4. Wing slightly brownish tinged; third and fourth veins curving slightly back- wards and converging slightly toward the tip, essentially parallel at the tip, cross- vein about half as long as last of fifth vein; anal margin slightly but distinctly rounded. Tip of calypter dark, its setae and the halter pale. Abdomen very elongate; second and third segment yellow with brown fore and hind margins, other segments brown; setae mostly black; fifth sternite bearing a pair of long stiff blunt bristles. Female face wider, as wide as third antennal segment, covered with gray pol- len; all segments of antennae short, third wider than long. Middle femur without distinct posteroventrals; middle tibia with only two ventrals, one near basal and one near distal third. Anal margin of wing slightly more prominent. Holotype male, allotype female, three male and ten female paratypes, from around puddles on trail, ridge above Pueblo Nuevo, Chiapas, Mexico, Dec. 24, 1962. Two female paratypes, from small pool in pine forest, near crest of Sierra Juarez above Ixtlan de Juarez, Oaxaca, Mexico, May 19, 1963. Holotype and allotype in U.S. National Mu- seum, others presently in the author’s collection. The species is the largest in size of those now known in the genus. The pair of rod-like appendages on the venter of the abdomen are rather distinctive, but recently Harmston and Miller (1966) have called attention to such structures in the following apparently related species from the Western United States. Neoparentia caudata (Van Duzee) comb. n. Sympycnus caudatus Van Duzee, 1917, Can. Ent. 49: 338. Parasyntormon caudatum (Van Duzee) Harmston and Miller, 1966, Proc. Ent. Soc. Wash. 68: 92. This California species differs from N. bisetosa by the smaller size, 2-3 mm, the more restricted yellow coloration at the base of the abdo- men, and the somewhat thickened second joint of the fore tarsus. Harmston and Miller (1966) call attention to the paired setae on the venter of the abdomen which are similar to those in many species of Parasyntormon. According to the illustration by Van Duzee (1930) the antenna of his species is of a Sympycnus or Neoparentia type and completely unlike Parasyntormon. Van Duzee describes the hypo- pygial lamellae as very long, yellow, fringed with long hairs. Though such lamellae are found in the genus Sympycnus, apparently paired rod-like setae on the abdominal sternites are not. Pubescence of the scutellum has not yet been noted. 256 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Neoparentia bicolor (Parent) comb. n. (Fig. 3) Nematoproctus bicolor Parent, 1954, Beitr. Ent. 4: 223. The species is most obviously distinct from Nematoproctus by the lack of numer- ous long pale hairs on the upper part of the proepisternum. Of particular note in Parent’s (1954) original description are the following: Length 3.0 mm; palpus creamy white, antenna yellow; a single series of acros- tichals; metepimeron, coxae, abdominal segments 2 and 3 and sides of 4 and 5 yellow; middle femur with series of 3—4 rigid setae ventrally near the base, middle tibia with 2 setae of remarkable length ventrally in the apical part; setae of calypter black. Neoparentia deformis sp. n. (Figs. 4, 5) Male.—Length 2.8-3.0 mm; wing 3.6 mm by 1.3 mm. Face long and narrow, gradually wider above, covered with yellowish-white pollen; front with metallic bluish ground color nearly obscured by dense yellowish- brown pollen. Palpus white with white hairs which are more prominent apically. First two antennal segments yellowish, third segment brown; first segment long and narrow; second very short; third longer than wide, triangular, rounded basally; arista nearly basal. Mesonotum metallic bluish nearly obscured by dense yellowish brown pollen; pleural surface dark with grayish pollen, lower half of metepisternum yellow. A series of acrostichal setae anteriorly, posteriorly acrostichals sparse or lacking. Legs almost wholly yellow, hind tarsus dark distally; setae mostly black, some pale setae anteriorly on fore and middle coxae and ventrally on femora. Middle tibia with antero- and posterodorsal near basal third, anterodorsal and ventral near distal third, four apicals; hind tibia with one anterodorsal, about four postero- dorsals, two or three small ventrals, four apicals. Lengths of joints of fore tarsus from base as 10-4-3-2-2; middle tarsus as 11-5-4-2-2; hind tarsus as 6-7-4-2-2. Wing clear, broadly rounded and slightly crinkled apically; third vein curving gradually forward near the tip, fourth curving gradually backward near tip and prominently divergent from third; crossvein three-fourths as long as last of fifth vein; anal margin slightly but distinctly rounded. Tip of calypter dark, its setae pale; knob of halter slightly to distinctly brownish. Abdomen very elongate, drooping, brown; hypopygium small for the genus. Female face as wide as antenna, grayish; third antennal segment only about as long as wide, subacute; hind tibia with three or four rather distinct ventrals; third and fourth wing veins only slightly divergent at tips, anal margin slightly more prominent. Holotype male, allotype female, and eight male paratypes, from small pool in pine forest, near crest of Sierra Juarez above Ixtlan de Juarez, Oaxaca, Mexico, May 19, 1963. One female paratype, by puddles on trail, ridge above Pueblo Nuevo, Chiapas, Mexico, Dec. 24, 1962. Holotype and allotype in U.S. National Museum, others presently in the author’s collection. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 257 Neoparentia obscura sp. n. (Fig. 6) Male.—Length 2.0 mm; wing 2.2 mm by 0.9 mm. Face long and very narrow, almost obliterated in lower part, covered with yel- lowish pollen; front metallic bluish dulled with grayish yellow pollen. Palpus dark. First two segments of antenna yellowish, first narrow, second shorter and broader; third segment brown, triangular, pointed, half again as long as wide; arista basal. Mesonotum metallic bluish green, dulled with grayish pollen anteriorly; pleura brownish, lower half of metapisternum yellow. Acrostichals reaching onto pos- terior slope. Legs wholly yellow with setae black. Middle tibia with two rather long antero- dorsals, first paired with a shorter posterodorsal near basal fourth, small ventral near distal third, three apicals; hind tibia with one anterodorsal near base, three or four shorter posterodorsals, two or three rather indistinct ventrals. Lengths of joints of fore tarsus from base as 8-5-4-3-2; middle tarsus as 10-5-4-3-2; hind tarsus as 5-7-5-3-3. Wing clear; third and fourth veins nearly straight and essentially parallel be- yond crossvein; crossvein about half as long as last of fifth vein; anal margin dis- tinctly rounded. Tip of calypter and its setae black; knob of halter brown. Abdomen slightly longer than thorax, brown. Female face as wide as first antennal segment; antennal segments all short, second shortest, third about as wide as long, rounded. Hind tibia with a series of indistinct ventrals. Anal margin of wing slightly more prominent. Holotype male, allotype female, five male and three female para- types, on foliage in rain forest, near 100 km marker, Rt. 175 above Rio Valle Nacional; five male and one female paratypes, near Rio Valle Nacional, Oaxaca, Mexico, May 16-18, 1963. One female paratype, near Puerto Vientes above Pueblo Nuevo, Chiapas, Mexico, May 24, 1963. Holotype and allotype in the U.S. National Museum, others presently in the author's collection. Neoparentia tarsalis sp. n. (Figs. 7-10) Male.—Length 2.0 mm; wing 2.2 mm by 0.9 mm. Face long and very narrow, eyes contiguous near middle, face covered with whitish pollen; front dark metallic bluish with slight brownish pollen. Palpus dark. Antenna brown; first segment narrow, about as long as third; second shorter; third slightly longer than wide, bluntly pointed; arista basal. Mesonotum metallic bluish green dulled with brownish pollen, more grayish on pleura; lower half of metapisternum yellow. Acrostichals reaching onto pos- terior slope. Legs wholly yellow with mostly black setae, some pale setae on anterior surface of fore coxa and ventrally on femora. Middle tibia with an antero- and postero- dorsal near basal third, anterodorsal and small ventral near distal third, four apicals; hind tibia with two strong anterodorsals, five or more prominent postero- dorsals, four or five small ventrals, four apicals. Lengths of joints of fore tarsus 258 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 from base as 8-5-2-2-3, last three joints usually tightly folded; middle tarsus as 11-5-4-3-2; hind tarsus as 5-7-4-3-2. Wing clear; third vein straight, fourth parallel with third for much of last part, bending slightly backward near tip; crossvein about two-thirds as long as last of fifth vein; anal margin slightly but distinctly rounded. Tip of calypter and its setae black; knob of halter pale brown. Abdomen very slightly longer than thorax, brown with slight yellow on side of second tergite and venter. Female face wider, as wide in middle as first antennal segment; antennal seg- ments all short, second shortest, third about as wide as long, rounded. Ventrals of hind tibia more prominent, joints of fore tarsus not folded. Anal margin of wing slightly more prominent. Holotype male, allotype female, six male and three female para- types, on foliage in rain forest, above Rio Valle Nacional, Oaxaca, Mexico, May 14-16, 1962; two male and one female paratypes, same locality, Dec. 20-21, 1962; two male and two female paratypes, near 100 km marker, Rt. 175, Sierra Juarez above Rio Valle Nacional, Oaxaca, Mexico, May 17-18, 1963. One male paratype, near Puerto Vientes above Pueblo Nuevo, Chiapas, Mexico, May 24, 1963. Holo- type and allotype in U.S. National Museum, others presently in the author's collection. The slightly divergent tips of the third and fourth wing veins seem to indicate relationship to Neoparentia deformis, but that species has a much more deformed wing and lacks the modified fore tarsus in the male. Neoparentia schildi sp. n. (Fig; 11) Male.—Length 2.5 mm; wing 2.7 mm by 1.0 mm. Face long and narrow, gradually wider above, covered with grayish brown pollen; front metallic bluish with slight brownish pollen. Palpus dark. Antenna brown, first two segments rather yellowish; first segment narrow; second shorter and wider; third half again as long as wide, triangular, bluntly pointed; arista basal. Mesonotum metallic bluish green dulled with brownish pollen; upper pleural surface brownish with gray pollen, lower pleural surface and metapisternum yel- lowish. Acrostichals reaching onto posterior slope. Legs wholly yellow with mostly black setae, some pale setae anteriorly on fore coxa and ventrally on femora. Middle tibia with antero- and posterodorsal near basal third, anterodorsal and ventral near distal third, one or more rather slender anteroventrals near tip, four apicals; hind tibia with one anterodorsal, about four posterodorsals, two small ventrals, four apicals. Lengths of joints of fore tarsus from base as 9-4-3-2-2; middle tarsus as 11-5-4-2-2; hind tarsus as 6-8-5-3-2. Wing slightly brownish tinged, rather pointed at tip of fourth vein; third vein straight, fourth vein slightly diverging from third near tip; crossvein two-thirds as long as last of fifth vein; anal margin slightly but distinctly rounded. Tip of calypter and its setae black; knob of halter very pale brownish. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 259 Abdomen slightly longer than thorax, brown. Female unknown. Holotype male and one male paratype, Turrialba, Costa Rica, Nov. 1922. Two male paratypes, La Suiza, Costa Rica, one dated April, 1922. All collected by Pablo Schild and in the A. L. Melander collec- tion in U.S. National Museum. Though Neoparentia schildi provides the southernmost known rec- ord for the genus, other species will undoubtedly be found in South America. REFERENCES Harmston, F. C. and L. S. Miller. 1966. New and little known Dolichopodidae from the Pacific Northwest and intermountain areas. Proc. Ent. Soc. Wash. 68: 88-93. Parent, O. 1954. Quelques Diptéres Dolichopodides (Deuxiéme article). Beitr. Ent. 4; 221-230. Robinson, H. E. 1964. capitalis (Norton) Messa Leach There are 3 or 4 species in North America. M. populifoliella (Town- send), a generally pale-colored species, and 1 or 2 other dark-colored species are leaf-miners of Populus. M. wuestneii (Konow ), a Holarctic species and known in North America only from Alaska and Northwest Territories, is a leaf-miner of Salix. Additional series of members of this genus will be needed before a revision is attempted; however, a new record for North America for a European species is given below. Messa nana (Klug) Tenthredo nana Klug, 1814, Mag. Gesell. Naturf. Freunde Berlin 8: 73. This species is quite distinct from all other members of this genus in North America and may be separated by the distinct ridges on the head enclosing an ocellar basin and supra-antennal basin, the lack of surface sculpture on the lobes of the mesonotum, the slender antenna, and the black coloration with the labrum, upper angles of the pro- notum, tegula, extreme base of each femur, and each tibia and tarsus whitish. It appears indistinguishable from the European specimens I have examined. The North American records for this species are as follows: MAINE: Scarsboro, 4 June 1966, L. J. Lipovsky (2¢ 2 ); Scarsboro, 11 June 1967, on birches in nursery, D. R. Smith (11 9 ¢). NEW YORK: Oswego Co., Oswego Township, from sticky board trap in pear orchard, June 3, 1966, R. W. Stelle (22 @ ). Profenusa MacGillivray Smith (1966) gave a key to the 6 North American species in this genus. P. platani Burks was included in that key; however, a new genus is described in this paper for this species. The 5 species now considered to belong in this genus are leaf-miners of Quercus, Betula, and Crataegus. P. lucifex (Ross) had been known only from the type specimen from Illinois, but, recently, I have seen additional specimens which were reared from red oak in Maine and from bur oak in Ontario. Bidigitus, n. g. Type-species.—Profenusa platani Burks. Description.—Antenna with 3rd segment longer than 4th segment; 2nd segment PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 281 longer than wide. Prepectus absent. Fore tarsus subequal in length to fore tibia. Fore wing with vein 2A and 3A straight at apex. Hind wing with cell R: open; anal cell present, with petiole much longer than cell. Tarsal claw with 2 long sub- equal outer teeth and a large acute basal lobe. This genus is close to Profenusa but differs by the unique tarsal claw and the short anal cell of the hind wing. In Profenusa, there is a single outer tooth on the tarsal claw, and the anal cell of the hind wing, when present, is much longer than its petiole. The genitalia of this species are also quite different from those of Profenusa. The single species in this genus, platani, is a leaf-miner of Platanus racemosa, and it is known only from southern California. Nefusa Ross The single species in this genus, N. anita (MacGillivray ), is known only from the eastern United States, where it is a leaf-miner of Viola. This species is unique in that it possesses a prepectus, a character ab- sent in all other North American Heterarthrinae and present only in the European genus Parna: otherwise, it is close to Profenusa. Although rarely collected, it was plentiful in Malaise trap collections in Mary- land. Setabara Ross The one species in this genus, S. histrionicus (MacGillivray), is known only from the western United States. All that is known of its biology is that a good series of adults was collected from Prunus at Talent, Oregon by L. G. Gentner. This species is probably a leaf- miner. The closed cell R, of the hind wing and the simple tarsal claw with an indistinct basal lobe will separate this genus from others of this tribe. Fenella Westwood Fenella Westwood, 1840, Introd. Mod. Class. Ins. 2: 54. Type-species.—Fenella nigrita Westwood. Monotypic. Fenella nigrita Westwood Fenella nigrita Westwood, 1840, Introd. Mod. Class. Ins. 2: 54. This is the first record of this species and genus for North America, and its discovery must be attributed to Dr. H. E. Milliron of the Ento- mology Research Institute, Canada Department of Agriculture, Ottawa. The specimens Dr. Milliron obtained were reared from larvae which were mining leaves of Potentilla at Ottawa, Canada. Recently, I have also seen several specimens of this species from East Lansing, Michi- gan. The species may be separated from other North American Fenu- sini by its small size (2 mm. or less in length) and by its 10- or 11- 282 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Fig. 1. Prolatus artus, n. sp., sheath. Fig. 2. Prolatus artus, n. sp. lancet. Fig. 3. Profenusa canadensis (Marlatt), tarsal claw. Fig. 4. Bidigitus platani (Burks), tarsal claw. Fig. 5. Prolatus artus, n. sp., tarsal claw. Fig. 6. Setabara histrionicus (MacGillivray ), tarsal claw. segmented antenna. The simple tarsal claws, absence of the anal cell in the hind wing, and the basal loop formed by vein 2A and 3A meet- ing 1A in the fore wing place this genus close to Fenusa. Fenusa Leach There are 3 species in this genus in North America. Benson (1951) gave a key to these 3 species and the following key is extracted from his. 1. Lateral lobes of mesonotum with fine surface sculpture; abdominal tergites dull with alutaceous surface sculpture; vein 2r of fore wing joins Rs before 3r-m; male rare; larva mines in leaves of Ulmus; eastern North America, probably introduced from Europe ulmi Sundevall Lateral lobes of mesonotum and abdominal tergites smooth, without surface sculpture; vein 2r of fore wing joins Rs beyond 3r-m ___..--_____-_ 2 2. Antenna longer than thorax; 3rd segment only 1% times as long as 4th; at least segments 4 and 5 more than twice as long as broad; male unknown; larva mines in leaves of Alnus; transcontinental and Holarctic —.__- MBE lar? NORA" De bree ereerode tnd, oY) 2 5 Bye ee Beh dohrnii (Tischbein ) Antenna shorter than thorax; 3rd segment twice as long as 4th; segments 4 and 5 less than twice as long as broad; males common; larva mines in leaves of Betula; eastern North America, California, Oregon, Utah, Wash- ington, probably introduced from Europe —----.--- pusilla (Lepeletier ) Prolatus, n. g. Type-species.—Prolatus artus, n. sp. Description.—Antenna with 3rd segment subequal to or only slightly longer PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 283 than 4th segment; 2nd segment slightly longer than wide. Prepectus absent. Fore tarsus long and slender, 144 times or more length of fore tibia; mid tarsus about 1% times length of mid tibia; hind tarsus shorter than hind tibia. Fore wing with vein 2A and 3A curving up and meeting 1A, forming a small basal cell. Hind wing with cell R: open; anal cell absent. Tarsal claw simple, long and slender. This genus is erected for the unique new species described below. It may be separated from other Heterarthrinae by the long, slender fore and mid-tarsi, the slender antenna with the 3rd and 4th segments subequal in length, simple tarsal claw, and the absence of the anal cell in the hind wing. It is apparently most closely related to Fenusa. Prolatus artus, n. sp. Female.—Length, 3.8 mm. Entirely black with clypeus, labrum, mandibles, tegula, upper angles of pronotum, extreme apex of each femur, and extreme base of each tibia light brown to whitish. Head and body, except for anterior and lateral lobes of mesonotum, covered with fine white hairs. Wings uniformly sub- hyaline. Characters as for genus. Clypeus truncate; malar space linear; genal carina ab- sent. Sheath slender, straight above and rounded below and at apex (fig. 1). Lancet as in fig. 2. Male.—See discussion. Holotype.—Female, Oregon, Jackson Co., 10 mi. N. W. Pinehurst, May 5, 1962, David R. Smith. USNM 69159. Host.—Unknown. Discussion.—I have seen one male which may be this species; how- ever, it is associated only by general structure and collection localities (both from Oregon ), and since the association is not definite, I hesitate to designate it as an allotype. The coloration of this male is similar to that of the female described except for the pronotum and tegulae, which are black. The fore and mid tarsi are also slightly longer in relation to the lengths of their respective tibia than those of the female. The data on this male are as follows: “Oregon, Benton Co., Scott’s Hill, 1 mi. S. W. Corvallis, moss and ground litter, March 10, 1960, J. D. Lattin.” It was undoubtedly collected from a Berlese sample. One structural character not mentioned in the generic or specific de- scription is the presence of a cross vein in the radial cell of the hind wing. This is extremely unusual and I know of no other sawflies with this extra vein. I am choosing only to mention this character since it may be an abnormality, a phenomenon very common in the wing venation of sawllies. This species is immediately recognized from all other Fenusini by the long slender fore and mid tarsi and the simple tarsal claws. REFERENCES Benson, R. B. 1951. Tenthredinidae. in Handbooks for the identification of British insects. Roy. Ent. Soc. London 6: 51—252. 284 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Burks, B. D. 1958. Tenthredinidae. in Krombein et al., Hymenoptera of America North of Mexico, Synoptic Catalog, Supplement 1, U. S. Dept. Agr., Agr. Monogr. 2: 8-18. 1967. Tenthredinidae. in Krombein et al., Hymenoptera of Amer- ica North of Mexico, Synoptic Catalog, Supplement 2, U. S. Dept. Agr., Agr. Monogr. 2: 15-26. Ross, H. H. 1936. The Nearctic sawflies of the genus Fenusa (Hymenoptera: Tenthredinidae). Trans. Illinois St. Acad. Sci. 29: 263-266. 1937. A generic classification of the Nearctic sawflies (Hymenop- tera: Symphyta). Illinois Biol. Monogr. 15: 1-173. 1951. Tenthredinidae. in Muesebeck et al., Hymenoptera of Amer- ica North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2: 22-82. Smith, D. R. 1966. A new Profenusa (Hymenoptera: Tenthredinidae) from red oak, with keys to the adults and known larvae of the Nearctic species. Ann. Ent. Soc. Amer. 59: 719-723. PARASITIC TROMBIDIID MITES ON DASYHELEA MUTABILIS (COQUILLETT) (DieTERA: CERATOPOGONIDAE ) R. H. Wuitset and R. F. ScHOEPPNER, Biologists, San Mateo County Mosquito Abatement District, Burlingame, California On August 13, 1965, a male ceratopogonid was collected and sub- sequently noted to be parasitized by three mites. The male gnat was collected in an oak-woodland area within Santa Clara County, Cali- fornia. Dr. Willis W. Wirth, Agricultural Research Service, United States Department of Agriculture, Washington, D.C., determined the host as Dasyhelea mutabilis (Coqg.), a common Nearctic gnat. Dr. Irwin M. Newell, University of California at Riverside, identified the mite larvae as belonging to the genus Valgothrombium. Identified specimens have been retained by each specialist. The position of the mites on the ceratopogonid is related in fig. 1. One mite was attached to the ventral side of the first abdominal seg- ment between the sternite and hind coxa. Another mite was attached between the second and third abdominal sternite. A third mite was dislodged during transport. Newell relates that these mites frequent moist situations where they crawl over the substrate in search of prey. Presumably, the Dasyhelea was parasitized immediately following pupation. According to Newell, it is not uncommon that these mites locate the pupae of the host and await ecdysis before attachment. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 285 Fig. 1. Valgothrombium mites attached to Dasyhelea mutabilis. Apparently, trombidiid mites are associated with a wide variety of insects as related by the literature. Newell (P. C., 1966) has records of Valgothrombium on Culicoides (Diptera: Ceratopogonidae). Odhiambo (1957) noted a trombidiid mite associated with Oxycarenus (Hemiptera: Lygaeidae) while Davis (1961) reports Allothrombium mitchelli as a predator on the balsam woolly aphid (Homoptera: Aphididae). Salman (1929) records a Trombidium species feeding on the eggs of Coleophora salmani (Lepidoptera: Coleophoridae). Severin (1944) relates that a Eutrombidium species serves as a predator of grasshoppers (Orthoptera: Locustidae). Records of trombidiid mites associated with diptera of medical importance are presented in an annotated list by Jenkins (1964). REFERENCES Davis, R. 1961. A mite, Allothrombium mitchelli, new to science, predator on _ the balsam woolly aphid. Proc. Ent. Soc. Wash. 63(4): 269-272. Jenkins, D. W. 1964. Pathogens, parasites and predators of medically impor- tant arthropods. Bull. World Health Organization, 30, Suppl., 150 pp. Newell, I. M. 1966. Personal correspondence. University of California, River- side, California. 286 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Odhiambo, T. R. 1957. The bionomics of Oxycarenus species (Hemiptera: Lygaeidae ) and their status as cotton pests in Uganda. Jour. Ent. Soc. So. Africa 20(2): 235-249. Salman, K. A. 1929. Notes on the immature stages and biology of a birch case-bearer. Ann. Ent. Soc. Amer. 22(3): 480-488. Severin, H.C. 1944. The grasshopper mite Eutrombidium trigonum (Hermann) an important enemy of grasshoppers. So. Dakota Agr. Expt. Sta. Tech. Bull. 3: 1-36. PROTECTION OF INSECTS BEING TRANSPORTED IN LIQUID At one time or another, most entomologists have had to try to work free with- out damaging them insects transported in liquid in vials into which wads or wisps of cotton had been inserted to protect the specimens against rubbing and breakage. The difficulties encountered in trying to separate the insects from the cotton are well known. To overcome the problems involved, at one time I tried substituting pieces of cloth for the cotton, thinking that one might be able to wash or float the insects from a single piece—but the result was just as bad or worse. Saran Wrap, however, can be used to protect the specimens and they are readily and quickly floated from the plastic. Saran was tested in several solvents over a period of about six months to de- termine whether or not it might prove to be generally useful for this purpose, with the following results: Solvent at full strength Effect on Saran 95% alcohol None evident. xylol No effect. glacial acetic acid No effect. dioxane Some dissolving of the Saran and the dioxane became slightly milky. Brittle after drying. chloroform No effect. Saran slightly stiffened after drying. ammonia No disintegration of the Saran but it became slightly browned. Saran flexible after drying. ethyl acetate No effect. As a result of the tests and our experience with the Saran, we are routinely substituting it for cotton or cloth, inserting a crumpled ball of it into the vials. Additional protection of specimens against mechanical damage may be ob- tained by placing them in more viscid media such as glycerine, or even heavy syrups like white Karo. Liquids of desired viscosity can readily be obtained by using methyl cellulose. When necessary, suitable agents, such as ethyl acetate, may be added to prevent decomposition. After they arrive at their destination speci- mens stored in such materials can easily be washed in water and then stored in alcohol—R. D. SHENEFELT, Department of Entomology, University of Wisconsin, Madison, Wisconsin 53706. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 287 DESCRIPTION OF A NEW GENUS AND SPECIES OF TRIMENOPONIDAE FROM PANAMA ( MALLOPHAGA ) Eustorcio MENDEZ, Gorgas Memorial Laboratory, Panama, R. P. The systematics of the family Trimenoponidae remains unsatisfac- tory. The group is comparatively small since only nine species are known, all of which are found in the American tropics associated mainly with rodents and marsupials. In his monograph, Werneck (1948) combines this particular group with the family Boopidae (found on mammals) and the families Ricinidae and Menoponidae (found on birds), in a single group, the family Ricinidae. However, this system of classification is not generally followed and many investigators con- sider the lice grouped in the family Trimenoponidae, as designated by Harrison (1915) and reviewed by Ferris (1922) as a separate, well- defined group restricted to mammal hosts. The most recent discussions regarding affinities and host-parasite re- lationships within the family Trimenoponidae have been presented by Hopkins (1949) and Vanzolini & Guimaraes (1955). To date this group contains five genera, namely: Cummingsia, Chinchillophaga, Harri- sonia, Philandesia and Trimenopon. Recently Dr. Phyllis T. Johnson has kindly submitted to me for study specimens of Trimenoponidae that apparently belong to an unknown genus and species and form the basis for the present paper. This discovery bears out the statement of Emerson (1964) to the effect that more collecting from probable hosts in South America could enlarge the family. Hoplomyophilus, n. gen. Generic diagnosis—Trimenoponidae lacking spinelike processes on ventral side of head; with two lateral spiniform setae at level of antenna. Prothorax subquad- rate, smaller than pterothorax; anterior thoracic stigmata conspicuous. Hoplomyophilus can be readily separated from the other known genera, with the exception of Philandesia and Trimenopon, by the absence of spinelike processes on the ventral region of head. The new genus can be separated from the latter two genera by structural fea- tures of the head and thorax. The following differences are outstand- ing: In Hoplomyophilis the head has two conspicuous lateral spiniform setae at same level with antenna. These spiniform setae are absent in Philandesia and Trimenopon. In Hoplomyophilus the prothorax is sub- quadrate, smaller than pterothorax, whereas both Philandesia and Trimenopon have a trapezoidal prothorax which is larger than ptero- thorax. Description—Head without spinelike processes on ventral region; with lateral and posterior margins sinuate; posterior margin with central sinus; with two dorsal, 288 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 Figs. 14, Hoplomyophilus nativus, n. gen., n. sp.: 1, dorsal-ventral view of male holotype; 2, genitalia of male holotype; 3, dorsal-ventral view of female allotype; 4, genitalia of female allotype. submarginal spiniform setae at the antennal level; clypeal region moderately pro- duced, not limited by distinct suture; eyes absent; maxillary palpi four-segmented, exposed; antennae four-segmented, exposed, not protected ventrally by a flap; antennal fossae deep; temporal lobes slightly prominent, truncate. Prothorax and pterothorax fused, both subquadrate; anterior thoracic stigmata conspicuous; pos- terior thoracic stigmata reduced; sternal plates fused into a single plate. Legs short, stout; with distinct pulvilli on the first tarsal segment of all legs. Abdomen subovate, with five pairs of abdominal stigmata; male genitalia of simple type. PROC, ENT. SOC. WASH., VOL. 69, No. 3, SEPTEMBER, 1967 289 Type species.—Hoplomyophilus nativus, new species. Affinities.—It is my opinion that Hoplomyophilus is closely related to the peculiar genus Harrisonia Ferris. Similarities are noticed par- ticularly in the morphology of the thorax and abdomen. Among the most important features common to the two genera are the complete union of prothorax and pterothorax, the fusion of the thoracic sternal plates into a single plate and the conspicuous anterior thoracic stig- mata. The fact that Harrisonia also parasitizes spiny rats, Proechimys semispinosus being its true host, seems to support this belief. Hoplomyophilus nativus, n. sp. (Figs. 1-4) Description —MALE (Figs. 1, 2). Head slightly wider than long, with clypeal region moderately produced, having anterior margin evenly convex, provided with minute and short marginal and submarginal setae. Lateral margins of head sinuate, slightly notched; each margin armed with two dorso-submarginal spiniform setae located at same level with the antenna. Posterior margin of head sinuate, with indentation at middle. Maxillary palpi four-segmented, exposed, with few short setae mostly concentrated on apical segment. Antennae four-segmented, exposed, bearing a few fine setae. Last antennal segment slightly swollen, semiglobular, larger than remaining segments. Antennal fossae deep, with very sclerotized, con- cave margin. Temples truncate, moderately projecting, with lower angle bearing a long dorsal seta that reaches middle of thorax. Both dorsal and ventral regions of head clothed with setae of different sizes and irregular distribution. Thorax longer than broad, bearing short and medium-size setae scattered on dorsal and ventral regions. Sternal plates apparently fused into a single plate extended along thorax, including ventral setae. Prothorax fused with pterothorax. This fusion apparently indicated by well chitinized longitudinal internal ridge in the middle of thorax. Prothorax subquadrate, smaller than pterothorax, provided with dorsal sublateral flap armed with conspicuous spiniform seta. Pterothorax subquadrate. Anterior thoracic stigmata prominent. Posterior thoracic stigmata reduced. Legs short and stout. First pair smaller than second and third pair, which are of about same size. All legs clothed with short setae sparsely distributed. Basal tarsal segment of all legs with small pulvilli. Apical tarsal segment of all legs ending in two claws. Abdomen slightly elongate, ovate, with sinuous lateral margins. Each tergum and sternum, except terminal ones, provided with no more than two irregular rows of short and medium size setae. Segments I-VI with two short and one long latero-marginal seta. The last reaches maximum length on segments IV—VI. Apical abdominal segments with few marginal and inner setae. Genitalia (Fig. 2) simple, with basal plate elongate, having posterior half dis- tinctly broad, provided with well chitinized walls; with acute sinus at middle of caudal margin. Anterior half gradually tapering to end on long, slender blade of about even width throughout. Parameres apparently absent. FEMALE (Figs. 3, 4). General morphology and chaetotaxy essentially as in the male. Differences between sexes are found in size and the terminal abdominal 290 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 PANAMA Fig. 5, Map of Panama showing the known range of Hoplomys gymnurus and the localities where Hoplomyophilus nativus has been collected. segments. The female genital area (Fig. 4) presents lateral gonopods provided with setae, preceded by latero-marginal group of setae. Caudal region separating gonopods having one marginal row of six medium-size setae and about two rows of minute setae accompanied by a few short ones. A patch of about ten small spicules is present at the middle of caudal area near its margin. Types.—Holotype male from Cerro Azul, Province of Panama, R. P., 21 March 1961; allotype female from Isla Escudo de Veraguas, Prov- ince of Bocas del Toro, 21 March 1964; one female paratype from same locality and date as allotype; one female paratype from Camp Pina, Canal Zone, 6 December 1960 and one female paratype from Rio Changena, Province of Bocas del Toro, 23 September 1961. All types collected by personnel of the Environmental Health Branch, Office of the Surgeon General, United States Army Caribbean, Fort Amador, Canal Zone. Lengths.—Male holotype, 1.27 mm; female allotype, 1.46 mm. Type host.—Hoplomys gymnurus (Thomas, 1897). Holotype and allotype will be deposited in the collections of the U.S. National Museum. One paratype will be deposited in the collec- tions of the British Museum (Natural History). Other paratypes will be deposited in the collections of Dr. Phyllis T. Johnson and the Gorgas Memorial Laboratory, respectively. Remarks.—It is of some interest to note that the specimens of the present new genus and species of mallophaga obtained on Isla Escudo de Veraguas, were collected from the subspecies Hoplomys gymnurus PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 291 wetmorei Handley, whereas the continental specimens were taken from Hoplomys gymnurus goethalsi Goldman. No morphological differences have been detected among the lice coming from these two different subspecies of host. It may be assumed that Hoplomyophilus nativus is rare since the examination of numerous specimens of the type host from several localities in Panama over a period of years has indicated that this rodent is more commonly parasitized by mallophaga of the genera Gyropus and Gliricola of the family Gyropidae. The terrestrial genus Hoplomys belongs to the spiny rat family Echimyidae and is monotypic. In Panama it has a wide distribution especially along the Caribbean Coast and has been collected in ever- green forest up to an altitude of 2100 feet. Besides Panama, the genus also is known to exist in the following countries: Costa Rica, Nicaragua, Colombia, Ecuador, Venezuela, Brasil and British Guiana. In figure 5 the range of Hoplomys gymnurus in Panama is indicated by the cross- hatched area. I wish to express my sincere appreciation to Dr. Phyllis T. Johnson for allowing me to study and describe this interesting mallophaga. I am also grateful to Dr. K. C. Emerson for advice and criticism given me during the preparation of this paper. REFERENCES Emerson, K. C. 1964. A new genus and species of Mallophaga. Ann. Mag. Nat. Hist. Ser. 13. 7:383-384. Ferris, G. F. 1922. The mallophagan family Trimenoponidae. Parasitology 14(1):75-86. Harrison, L. 1915. The respiratory system of Mallophaga. Parasitology 8(1): 101-127. Hopkins, G. H. E. 1949. The host-associations of the lice of mammals. Proc. Zool. Soc. Lond. 119( 2) :387—604. Vanzolini, P. E. and L. R. Guimaraes. 1955. Lice and the history of South American land mammals. Rev. Brasil. Ent. 3:13—46. Werneck, F. L. 1948. Os Maldfagos de Mamiferos. Parte I. Amblycera e Ischnocera (Philopteridae e parte de Trichodectidae). 243 pp. Revista Brasil. Biol., Rio de Janeiro. 292, PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 A KEY TO THE SPECIES OF LYSTRIDEA BAKER WITH DESCRIPTION OF A NEW SPECIES FROM CALIFORNIA (Homoptera: CICADELLIDAE: ERRHOMENELLINI) James P. Kramer, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560 The species of Lystridea Baker were last treated by Oman (1938: 178-180) in his revision of the Nearctic Errhomenellini, a tribe which he (1949: 61) later assigned to the Tettigellinae, now called Cicadel- linae. Metcalf (1963: 4) has subsequently transferred the tribe to the Aphrodinae. There is still question as to the correct subfamily place- ment of the tribe, but the resolution of the matter is beyond the scope of this paper. For a description of the tribe, see Oman (1949: 75). Lystridea can be separated from all other North American genera in the Errhomenellini by the following combination of characters: crown with numerous, fine, irregularly longitudinal striae; head short and broad, crown not strongly produced anteriorly; head as broad as or broader than pronotum, posterior margin of pronotum not incised. The species of the genus are large (5.5-10mm.), robust, and known only from our far western States of California, Oregon, and Nevada. In the key below, it should be noted, only the male genitalia provide characters which allow positive differentiation of species. While size is useful in separating females, there is some intergradation in this character. The female of the new species is unknown. The interested student is referred to Oman (1938: 178-180) for additional data on uhleri (Baker) and nuda Oman. KryY TO THE SPECIES OF Lystridea BAKER ERAN leg tes lee Sie he eR ee oe ee a Hemalesyes tt. = fin «sec Pe OG ate AE. el te 4 2. Aedeagus greatly enlarged distally and with long paired processes near apex (sce | ES cen ene eae ee ev Naber ENO DOI ne RUDE D PS wilkeyi, n. sp. Aedeagus not greatly enlarged distally and without long paired processes WEA ADCK eee bees ten eae eens ee ee ete pace eee 3 3. Aedeagus with many stout setae on apical portion (fig. 6) —. uhleri (Baker ) Aedeagus with minute teeth on apical portion (fig. 4) —___- nuda Oman 4. Length 8-10 mm., width of head 2.75—3.00 mm. -______ uhleri (Baker ) Length 6.75-8.25 mm., width of head 2.50-2.75 mm. —.________. nuda Oman Lystridea wilkeyi, n. sp. (Figs. 1-3) Length: Male 7.00-7.50 mm. Coloration: Ground color of venter sordid stramineous to reddish brown, usually marked with black; legs pale brown to reddish brown, variably touched with black and/or spotted with yellow; clypeus and clypellus brownish, usually with PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 293 wilkeyi _——_—__— \S nuda hi =~ ee a 6 Zi Figs. 1-3, Lystridea wilkeyi, n. sp.: 1, aedeagus laterally; 2, style laterally; 3, aedeagus ventrally. Figs. 4, 5, Lystridea nuda Oman: 4, aedeagus laterally; 5, style laterally. Figs. 6, 7, Lystridea uhleri (Baker): 6, apical portion of aedeagus laterally; 7, style laterally. Note: The aedeagi of nuda and uhleri are of the same general shape, but the aedeagus of uhleri is much larger. darker irrorations; genae and lora black, heavily marked with yellow as irregular spots on lora and as a crescentic band on each gena; antennal bases and surround- ing areas black. Crown, pronotum, and scutellum with ground color yellowish brown; crown and pronotum variably mottled with black, mottling usually strong- est on anterior coronal margin either side of apex and on pronotum just behind each eye; scutellum with basal angles and mesal area broadly black; forewings gray, variably mottled with dark brown, most veins marked with alternating tan and dark brown areas. Male genitalia: Aedeagus in ventral view (fig. 3) with shaft narrowed, a pair of sharp projections basally, and a longer pair of lateral appendages subapically; aedeagus in lateral view (fig. 1) with apical portion greatly enlarged; style (fig. 2) relatively simple. Female genitalia: Female unknown. Types: Holotype male, Lebec, California, Alt. 4,000 ft. May 13, 1928, J. O. Martin. Nineteen male paratypes as follows: 1, Bodie, Cali- fornia, 8,475 ft., July 1-7, Wickham; 3, Argus Mts., Inyo County, Cali- fornia, May 22, 1937; 9, Walker Pass, Kern County, California, May 20, 1960, R. P. Allen; 2, 29 miles west of Benton, California, June 24, 1960, R. P. Allen; 2, Junction 395-108, Mono County, California, June 14, 1959, R. P. Allen; 2, Topaz, California, July 4, 1917. Notes: The species is named for Mr. R. F. Wilkey of the California Department of Agriculture, Bureau of Entomology, Sacramento, Cali- 294 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 fornia. He sent me the first specimens of this species for identifica- tion. The holotype will be deposited in California Academy of Sciences and paratypes in the collections of the United States National Museum and the Bureau of Entomology of the California Department of Agri- culture in Sacramento. REFERENCES Oman, P. W. 1938. Revision of the Nearctic leafhoppers of the tribe Errhomen- ellini (Homoptera: Cicadellidae). Proc. U. S. Nat. Mus. 85: 163-180. 1949. The Nearctic leafhoppers (Homoptera: Cicadellidae). A generic classification and check list. Mem. Ent. Soc. Wash. 3: 1-253. Metcalf, Z. P. 1963. General Catalogue of the Homoptera. Fascicle VI Cicadel- loidea, Part 8 Aphrodidae: 1-268. Published by the U. S. Department of Agri- culture. A NEW NASAL MITE FROM THE ROBIN (TURDUS MIGRATORIUS) (ACARINA, SPELEOGNATHINAE ) Gorpon Marston Ciark, U.S. Department of Health, Education, and Welfare, Public Health Service, National Institutes of Health, National Institute of Allergy and Infectious Diseases, Rocky Mountain Laboratory, Hamilton, Montana In 1963 a series of speleognathid nasal mites was recovered from two robins (Turdus migratorius) in the Bitterroot Valley of western Montana. These mites were a new species which is herein described. All measurements are given in millimeters. Boydaia turdi, new species (Fig, 1) ADULT.—Conforms to the characteristics of the genus Boydaia as outlined by Fain (1963). LARVA (Fig. 1A).—Color milky white; shape broadly oval; weakly sclerotized; size, 0.34 in length by 0.3 in width, excluding gnathosoma which is 0.08 wide at base; with three-segmented palpi measuring 0.048 in length; palpal tarsus reduced to a stubby, subapical tibial thumb, bearing two barbelled and one bladelike seta. Dorsum: Anteriorly with a pair of slightly expanded, circular, finely barbelled sensillae 0.027 in length, preceded by a pair of short presensillar barbelled setae; dorsal setae barbelled, short and expanded, arranged in a 4-4-2-2-2 formula; dorsal cuticle very finely striated. Venter: With two pairs of short, barbelled sternal setae and two pairs of genital setae. Legs: Tarsus I specialized and characteristically modified, elongated, measures PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 295 Fig. 1, Boydaia turdi, larva: A, ventral and dorsal views; B, tarsus I, dorsal; C, tarsus I, ventral; D, head of claw, tarsus I. 296 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 (base of pulvillus to tibia) 0.04 long and 0.05 wide; claws equal in length, 0.065, characteristic in shape, one fine, with a shepherds crook curve apically, the other stouter with a “sea horse head” configuration apically (Fig. 1D); tarsal I setation indicated in Fig. 1B and C; tarsae of other legs relatively unmodified although the claws of tarsus II are elongate, blunt and equal (0.027); claws of tarsus III normal, short, curving, blunt (0.015). Numbers of setae present on leg segments similar to other Boydaia larvae. Diagnosis: Boydaia turdi larvae may be distinguished from all other known larvae of the genus Boydaia, particularly B. jordani van Eyndhoven, 1955, by the configuration of the claws of tarsus I. In B. turdi one claw is clubbed, the other has a fine hooked extremity while in B. jordani both claws are clubbed. Holotype: Larva USNM 3247 is deposited in the collection of the United States National Museum, Washington, D.C. Paratypes are deposited at the Rocky Mountain Laboratory. Type Host: Turdus migratorius Linnaeus. Type Locality: Lost Horse Canyon, Ravalli Co., Montana. REFERENCES Fain, A. 1963. Chaetotaxie et classification des Speleognathinae (Acarina: Trombidiformes ). Bull. Inst. Roy. des Sci. Nat. de Belgique 39(9): 1-80. Van Eyndhoven, G. L. 1955. Boydaia jordani sp. n., a new endoparasitic mite (Acarina: Speleognathidae). Trans. Roy. Ent. Soc. London: 203-208. DISTINGUISHING AMOEBALERIA DEFESSA (OSTEN SACKEN) FROM A. SACKENI GARRETT (DieTERA: HELEOMYZIDAE) Gill (1962, Proc. U.S. Nat. Mus. 113: 579) in his revision of this family, stated that these two species could be distinguished only by differences in the male postabdomen. The series of these species in the U.S. National Museum show the following differences; that in the venation of the wing is apparently valid for both sexes. Amoebaleria defessa (O. S.).—&, in addition to the postabdominal differences, the hind basitarsus lacks a claw or apicoventral production. In both sexes, the narrowest distance between the 2 branches of vein Rs, near tip of the wing, is distinctly greater than the difference at the same point between the anterior branch (Re,s) and the costa; the angle at the apex of cell R: is approximately 25°. Amoebaleria sackeni Garr.— é, hind basitarsus with distinct apicoventral pro- duction or incipient claw. In both sexes, the anterior branch of Rs is more curved, making the narrowest distance between the branches equal to or less than the distance between the anterior branch and the costa; the angle at apex of cell Ri is approximately 30°.—GrorcE C. StEysKaL, Entomology Research Div., ARS, U.S. Department of Agriculture, Washington, D.C. 20560. PROC. ENT. SOC. WASH., VOL. 69, No. 3, SEPTEMBER, 1967 297 SOMETHING BETTER THAN POLYPORUS OR PITH FOR DOUBLE MOUNTS All of the materials commonly used for making double mounts with minuten nadeln frequently give trouble by becoming loose, either on the main pin or the minuten. The silicone rubber that I have been using for microvial stoppers, etc. (Gurney, Kramer, and Steyskal, 1964, Ann. Ent. Soc. Am. 57: 240-242), obtain- able from plastics distributors in most large cities as General Electric RTV-11 or Dow Corning RTV-521, has excellent pinning qualities. A very delicate pin may be pushed into it, even with the blunt end foremost, and it will be firmly gripped and not come loose. I cast the material into approximately 3-mm-thick plaques in plastic boxes and cut it into cubes with a knife—GrorcE C. STEYSKAL, Entomology Research Division, ARS, U.S. Department of Agriculture, Wash- ington, D.C. 20560. SEX RATIOS OF PLATYPUS (CoLEOPTERA: PLATYPODIDAE ) In the use of ultraviolet light traps for determining populations of various forest insects, members of the ambrosia beetle genus Platypus are recovered and enu- merated. Sex ratios of two species in the genus are of interest. According to W. M. Blackman (Miss. Agric. Expt. Sta. Tech. Bul. 11:38, 1922), male Platypus beetles in galleries greatly outnumber the females. H. G. Hubbard (U.S.D.A. Bur. Ent. Bul. 7:14, 1897) states that “the female is frequently accom- panied by several males in the galleries”. Both of these workers indicate that males should be encountered with greater frequency than females. Although the light traps sample only the population of newly emerged adult Platypus beetles, moving from trees where they developed to other trees or stumps suitable for attack, there is good reason to believe that both sexes are equally attracted to the ultraviolet light source. In 1965 and 1966 the following recoveries were found in the trap collections. Platypus compositus Say, a species most frequently associated with hardwood trees: 194 males, 135 females. Platypus flavicornis Fabricius, a species most fre- quently associated with coniferous trees: 762 males, 1306 females. The abundance of P. flavicornis is attributable to operation of the traps in woodlands predomi- nantly stocked with pine trees. A preponderence of males is not indicated in these studies. A third species of the eastern United States, P. quadridentatus Oliver, has not been recovered in the trap collections in Florida.—L. A. Herrick, Depart- ment of Entomology, University of Florida, Gainesville 32601. 298 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 BOOK REVIEWS A Revision of the Genus Lobopoda (Coleoptera: Alleculidae) in North America and the West Indies. By John M. Campbell. Illinois Biological Monographs 37, 203 pp., 186 figs. University of Illinois Press, Urbana and London. 1966. The family Alleculidae has been neglected until just recently, and the genus Lobopoda, one of the most difficult, was in bad need of work. This revision by Campbell helps to make sense of Lobopoda and points to more good work in the family. Lobopoda now contains 195 described species, all from the New World. Of those, 87 are from the area covered in this revision. Campbell has arranged the 87 species (30 new) in 5 subgenera (3 new) and 33 species groups. Descrip- tions of all taxa, distributions, and illustrations of male genitalia are given. The males are keyed to species; females are keyed to subgenus but only in one sub- genus are they keyed to species. Phylogenetic trees give some idea of relationships between species, and the position of the genus in the family is discussed. Campbell is to be congratulated on his first major work—T. J. Spm_Man, Entomology Re- search Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560. A Bibliography of the Psocoptera (Insecta). Australian Zoologist XIII (2): 137-209, 1965. A Catalogue of the Psocoptera of the World. Ibid XIV (1): 1-145, 1967. By C. N. Smithers. Price: Of latter, $2.00; of former, not known. These publications are regarded as of such major importance as to merit general notice; for specialists of the Psocoptera (Corrodentia) they are invaluable. The bibliography, estimated to consist of more than 1,200 titles, was compiled with the active cooperation of 22 specialists and others. The catalogue includes 1,605 species regarded as valid, in 197 genera and 31 families. All valid species and synonyms, including those preserved in amber but not other fossils, are listed, with their distribution. However, precise type localities and most references sub- sequent to original descriptions are not shown. Mr. Smithers, Curator of Insects in the Australian Museum, Sydney, who has studied psocid systematics for some 12 years and is still a young man, in these two publications has earned warm thanks for retrieving the scattered literature of a much neglected group of insects. —AsHLEY B. Gurney, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560. PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 299 1967 MEMBERSHIP DRIVE ENTOMOLOGICAL SOCIETY OF WASHINGTON WASHINGTON, D. C. MEMBERS: Let’s increase the membership of our Society! If you know someone interested in the science of entomology who is not now a member please pass this application on to that person. If the blank is cut along the rule at left, this issue of the Proceedings will not be spoiled. However, it is not necessary to use this form as long as the requested information is furnished to the Chairman, Membership Committee or the Editor. For information concerning the Society and its officers, see the inside cover of this issue. MEMBERSHIP APPLICATION Date Application is hereby made for membership in the Entomological Society of Washington. Name Mailing Address Interest in and/or association with the science of entomology: Applicant's Signature 300 PROC. ENT. SOC. WASH., VOL. 69, NO. 3, SEPTEMBER, 1967 For more than half a century, Cyanamid has consistently led the chemical industry in developing new products and application techniques that have helped immeasurably to bolster our na- tional farm economy. ™ Topping the list of Cyanamid “firsts” is Malathion LV* Concentrate, introduced commercially last year for the control of boll weevils on cotton after two years of use on more than 1 million treatment acres in cooperation with the U.S.D.A.’s Agricultural Research Service. m Malathion LV Con- centrate is also being used extensively to combat grasshoppers, cereal leaf beetles, corn rootworm beetles, mosquitoes, blue- berry maggots, flies and beet leafhoppers. m Watch for progress reports of new tests conducted against many other pests with both aerial and ground equipment. Data being processed daily show clearly that Malathion LV Concentrate is fast making all other methods of insect control obsolete! ™ Before using any pesticide, stop and read the label. *Trademark SERVES THE MAN WHO MAKES AMERICAN CYANAMID COMPANY A BUSINESS OF AGH Cae PRINCETON, NEW JERSEY PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. 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Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) MENDEZ, E.—Description of a new genus and species of Trimenoponidae from Panama ( Mallophaga ) PRICE, R. D. and K. C. EMERSON—Additional synonymies within the amblyceran bird lice (Mallophaga) ROBINSON, H.—Neoparentia, a new genus of American Dolichopodidae ( Diptera ) SHENEFELT, R. D.—Protection of insects being transported in liquid SLATER, J. A.—Synonymy in the Lygaeidae (Hemiptera) SMITH, D. R.—A review of the subfamily Heterarthrinae in North America (Hymenoptera: Tenthredinidae ) STEYSKAL, G. C.—Replacement names for preoccupied specific names in Dolichopodidae (Diptera ) STEYSKAL, G. C.—Anthomyia procellaris Rondani in North America (Dip- tera: Anthomyiidae ) STEYSKAL, G. C.—A new subgenus for Leucopis luteicornis Malloch, a predator on mealybugs in India ( Diptera: Chamaemyiidae ) STEYSKAL, G. C.—Distinguishing Amoebaleria defessa (Osten Sacken) from A. sackeni Garrett (Diptera: Heleomyzidae ) STEYSKAL, G. C.—Something better than polyporus or pith for double mounts STONE, A.—A synoptic catalog of the mosquitoes of the world, supplement Ill (Diptera: Culicidae) WEBER, N. A.—Synonyms of Trachymyrmex bugnioni Forel and Trachymyr- mex diversus Mann (Hymenoptera: Formicidae ) WHITSEL, R. H. and R. F. SCHOEPPNER—Parasitic trombidiid mites on Dasyhelea mutabilis (Coquillett) (Diptera: Ceratopogonidae ) BOOK REVIEWS MEMBERSHIP APPLICATION BLANK Tol. 69 DECEMBER 1967 No. 4 PROCEEDINGS of the ENTOMOLOGICAL SOCIETY ot WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 PUBLISHED QUARTERLY CONTENTS (Continued on back cover) BRAM, R. A.—Lectotype assignments for several species of the genus Culex miGueneast Asia «( Diptera: \Culicidae) 2. 327 FLINT, O. S., JR.—The first record of the Paduniellini in the New World Mitcupmieras @sychomundae) o.oo he Be 310 GAGNE, R. J.—Notes on the genus Thecodiplosis Kieffer in North America, and some generic reassignments (Diptera: Cecidomyiidae) —-_-__-- 338 HERRING, J. L. and H. C. CHAPMAN—A new species of Omania from Piineonesrs | ( bemipters: salaiae)) 2 Ee 354 JAMES, M. T.—The blow flies of Dominica (Diptera: Calliphoridae) ——— 368 JOHNSTON, D. E.—On the occurrence of two species of Palaeacarus in the eastern United" States (Acar: Acarifonmes) #2. - 2 301 JOHNSTON, D. E.—Observations on Oribatei. The discovery of Haploch- thonius simplex (Willmann) in North America (Acari: Acariformes) 365 KINGSOLVER, J. M.—On the genus Rhipibruchus Bridwell, with descrip- tions of a new species and a closely related new genus (Coleoptera: Bric bidae- Ta Chinge)\o- 5 eae a ee oe ae Re ond ee ee Ne elt Pee 318 LIN, C. S.—Bionomics of Cerceris simplex graphica at Lake Texoma (Hy- menoptera:~ Splecidae: : Philanthimae) 20. 8s 312 LIN, N.—Linear copulation in Xylocelia franclemonti Krombein (Hymenop- COT Se CUTE 528) (Die ia ST a LS TSS (AREF Pe en SR pe Fo ee Re Oa OS ee SC ee 2. At least upper part of face and lower part of front covered mith dark brown mollen) wathowtlopviots. Stay border’ 2. a 6 Prnd: tipia, wathout evident posterodorsal, 2 3 Hind tibia with a small but distinct posterodorsal ______________________ 4 3. Hypopygium asymmetrical, strongly swollen on one side (Fig. 5) CON Tex )) peers as i eee a ll eel M. asymmetricus, n. sp. Hypopygium essentially cylindrical (Texas, Mexico) _.. M. fulvosetosus Parent 4. Apex of hypopygium oblique with upper edge sharply projecting, a tuft of stiff black hairs on apical surface (Fig. 4) (Mexico) __ M. lithophilus, n. sp. Apical surface not oblique, bearing only rather pale hairs — 5 5. Hypopygium bearing only a narrow pubescent appendage (Md., S.C., Fla., Mexico, Costa Rica, Panama, West Indies, Europe, New Zealand?) BS Bae ges BT her Nd oS SRE Shee a AR ips ee oc 8 ea M. albipes (Zetterstedt ) Hypopygium bearing a small but broad pubescent appendage distally (West nr CLiGS)) Same eee. cater eee Tel a eer ee 2h M. eaudatus ( Aldrich) 6. Face of male narrower than that of female, never whitish below, usually distorted in dry specimens; 3rd antennal segment slightly longer than broad with arista borne at middle of dorsal edge _...._.___-________ i 330 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 4 Fig. 1-5. Micromorphus, male hypopygia; 1. M. leucostoma, n. sp.; 2. M. knowltoni, n. sp.; 3. M. bifrons Robinson; 4. M. lithophilus, n. sp.; 5. M. asym- metricus, N. sp. Face of male very broad, as broad as in female, often whitish below, rarely distorted in dry specimens; 3rd antennal segment as broad as long with arista bome slightly beyond middle of dorsal edge ___-_-----__----------------- 8 7. Hypopygium without evident appendages, with a tuft of long hairs on tip CNBYeM@ Gebec) Res ts 30) sevens oy 2 5 ores M. minimus (Van Duzee) Hypopygium with a pair of very long slender lamellae, without long apical TYAS Cea OTT pIN acpi a Re oe M. longilamellatus Robinson 8. Face without white pollen on lower part; hypopygium with only small slen- der apical appendages (Fig. 2) (Utah) —__-_______. M. knowltoni, n. sp. Face with white pollen on lower part; hypopygium with a pair of short pubescent. apical appendages ..._.__.. ee 9 9. White pollen on face restricted to area above suture; hypopygial appendage broad, bearing many long setae (Fig. 3) (Iowa) — M. bifrons Robinson White pollen covering entire lower half of face; hypopygial appendage nar- row, bearing a few short hairs (Fig. 1) (Washington) M. leucostoma, n. sp. The following four species are previously undescribed. Micromorphus leucostoma, n. sp. (Fig. 1) Male.—Length 1.8 mm; wing 1.6 mm by 0.7 mm. Face very wide above, half as wide below, upper half covered with black velvety area which extends above antennae onto lower third of front, lower part of face covered with white pollen; front broad with sides diverging from base, with yellowish pollen above toward middle and grayish pollen toward side; palpus brown with gray pollen and numerous pale hairs; proboscis brown. Antenna black; segment 1 short, broadened toward tip, bare above; segment 2 slightly broader and nearly as long as first, ringed with small dark setulae; segment 3 half again as long as second, rather triangular, covered with fine hairs, with arista inserted a little beyond middle of dorsal edge. Lower postocular setae pale. Thorax rather gibbous with posterior slope of mesoscutum flattened; dark with yellow pollen on mesoscutum and scutellum, more grayish pollen on pleura, hind margin of scutellum yellowish. Bristles yellowish; acrostichals lacking; 6 pairs of dorsocentrals; 1 pair of scutellars; 1 rather small bristle above fore coxa. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 331 Middle and hind coxae infuscated at base, last segments of tarsi and hind tarsus from tip of segment 1 brown, legs otherwise yellow. Hairs and bristles mostly pale; fore and middle coxae with numerous hairs on anterior surface, fore coxa with a series of larger setae along distal margin, middle and hind coxae with a rather small external seta; middle and hind femora with a preapical seta; fore tibia without distinct bristles; middle tibia with 1 anterodorsal and a smaller posterodorsal near basal third, 3 distinct apicals; hind tibia with 1 anterodorsal and 1 posterodorsal near basal third, 1 posterodorsal near distal third, 2-3 apicals. Lengths of segments of fore tarsus from base as 7—3-2-2-2; middle tarsus as 9-43-99. hind tarsus as 5—5—3—-2-2. Wing rather oval, clear with brown veins; vein 3 nearly straight, vein 2 diverg- ing from it slightly, vein 4 nearly parallel to 3 beyond crossvein and ending slightly behind wing apex; crossvein nearly perpendicular to base of vein 4, about two-fifths as long as last of vein 5; vein 6 represented by slight fold. Calypter, its setae, and the halter pale. Abdomen only slightly longer than thorax, dark with slight grayish pollen toward sides and base; pubescence pale, marginal setae toward side of first tergum rather prominent. Hypopygium (fig. 1) small, brown, with only small appendages, with numerous fine hairs over the blunt apical surface. Female with head and parts of legs broken off, but otherwise showing dif- ferences only in genitalia. Holotype ¢ and allotype 2°, from O'Sullivan Dam, Grant Co., Washington, July 22, 1954, M. T. James. Specimens in USNM, No. 69344. The new species is one of three known to me having the face broad and apparently resistant to distortion as the specimen dries. Of the other two, Micromorphus knowltoni, described below, is distinct in the complete lack of white pollen on the face, and M. bifrons Robinson of Iowa has a smaller less distinct white pollinose area on the face. Each of the three species has distinctive hypopygia, that of M. leu- costoma being unusually short and stout. Micromorphus knowltoni, n. sp. (Brig, 84) Male.—Length 1.4 mm; wing 1.7 mm by 0.6 mm. Face very broad, about two-fifths as broad below as above, divergent margins continuing along sides of the broad front; brown velvety area covering face and extending onto lower margin of front; a few pale reflections above the clypeus, most of front with light brownish pollen; palpus and proboscis brown, former with a small black apical seta. Antenna brown, segment 3 broken off, remaining parts as in M. leucostoma. Lower postocular setae pale. Thorax mostly as in M. leucostoma, brown with yellowish pollen above and grayish pollen toward the sides and on the pleura; 1 strong bristle above fore coxa. Outer surfaces of middle and hind coxae and sometimes part of fore coxa infuscated, distal segments of tarsi brownish, legs otherwise pale. Most hairs and bristles of legs rather pale; coxae and femora as in M. leucostoma; fore tibia with- out evident bristles; middle tibia with a distinct rather dark anterodorsal above 332 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 the middle, 3 strong apicals; hind tibia with a strong anterodorsal near basal third, 2 small posterodorsals all rather dark, 2-3 small apicals. Lengths of segments of fore tarsus from base as L0—5—4—3-3; middle tarsus as 12—7—5-3-3; hind tarsus as 8— others broken off. Wing rather oval, clear with veins pale toward base; veins 2, 3, and 4 nearly straight and parallel beyond crossvein, vein 4 diverging slightly toward tip and ending at or just after wing tip; crossvein perpendicular to base of vein 4, two- fifths as long as last of vein 5; vein 6 represented by very slight fold. Calypter and halter yellow, setae of former pale. Abdomen about as long as thorax, greenish brown dulled with considerable eray pollen; pubescence pale. Hypopygium (fig. 2) brownish, small, with numer- ous pale hairs on flattened apical surface and a few very short slender brownish appendages toward the upper side, Female with face only slightly wider than in male, about half as wide below as above; antennal segment 3 triangular, about as long as broad, with arista inserted just beyond middle of dorsal margin; lengths of segments of hind tarsus as S—-9-6—4-4. A Holotype ¢ and 4 & paratypes, Clear Creek Canyon, Utah, July 14-16, 1946; allotype 2°, Long Valley, Utah, July 4, 1946, all G. K. Knowlton. Specimens in the Iowa State University collection, Ames, Iowa. The species is closely related to both Micromorphus leucostoma, described above, and M. bifrons Robinson of Iowa, both of which have similar wide faces. In M. knowltoni, however, there is essentially no white pollen on the face and the hypopygium bears only very small slender appendages. Micromorphus asymmetricus, 1. sp. (Fig. 5) Male—Length 1.4 mm; wing 1.4 mm by 0.6 mm. Face narrow below, less than a fourth as wide as above, with sides running nearly parallel slightly above the clypeus, the narrow white pollinose margins sharply demarcated from the central black triangle; front broad with diverging sides continuous with those of face, covered with light brownish pollen; palpus black, proboscis brown. Antennae and postocular area as in M. leucostoma, Thorax similar to M. leucostema, with brownish yellow pollen above, more grayish pollen toward side and on pleura; tip of scutellum rather pale; bristles light brownish, 1 large seta above fore coxa. Bases of coxae slightly infuscated, distal segments of tarsi brownish, legs other- wise pale. Fore and middle coxae with numerous pale hairs anteriorly the distal ones longer, middle and hind coxae with a rather pale external bristle, other hairs and bristles of legs mostly dark; middle and hind femora with a preapical seta; fore tibia without evident bristles; middle tibia with an anterodorsal and a posterodorsal near base, 3 apicals; hind tibia with 1 small anterodorsal near basal third, 3 very small apicals. Lengths of segments of fore tarsus from base as 12-6-4—3-2; middle tarsus as 16-S—5—4-3; hind tarsus as 8-10—-6-44. PROG. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 333 Wing rather oval, clear with brown veins; vein 3 nearly straight, vein 2 slightly divergent; vein 4 slightly curved, equally far from 3rd at crossvein and near tip, ending slightly behind wing tip; crossvein perpendicular to last of vein 4, a third to a half as long as last of vein 5; vein 6 represented by a slight fold. Calypter, its setae, and the halter rather pale. Abdomen about as long as thorax, tapered toward tip; mostly blackish with slight grayish pollen toward side and base; hairs and bristles rather yellowish brown. Hypopygium (fig. 5) small, dark brown, distinctly swollen near tip on left side; a few short, upwardly curved appendages and a small tuft of pale hairs at tip. Holotype ¢, Aug. 4, and 1 4 paratype, Aug. 5, 1962, from moist calcareous rock surfaces, in a small shaded ravine about 5 km south of Tamazunchale, San Luis Potosi, Mexico. Holotype in USNM No. 69345; paratype presently in my collection. The species is one of the narrow faced group related to Micro- morphus albipes Zetterstedt. The new species is quite distinct in the very asymmetric hypopygium and the hind tibia completely lacking posterodorsals. Micromorphus lithophilus, n. sp. (Fig. 4) Male.—Length 1.3 mm; wing 1.3 mm by 0.6 mm. Face narrow below, about a third as wide as above, with the diverging sides straight, covered with blackish velvety areas with paler reflections along the margins of the eyes; front broad with diverging sides continuous from face, covered with a dull brownish pollen; palpus and proboscis black. Antenna essen- tially as in M. leucostoma, segment 3 slightly broader than long. Lower postocular setae pale. Thorax as in M. leucostoma. Bases of middle and hind coxae and sometimes base of fore coxa infuscated, distal segments of tarsi dark, legs otherwise mostly pale. Fore and middle coxae with numerous pale hairs anteriorly, longer setae distally, middle and hind coxae with a rather dark external seta, otherwise hairs and bristles of legs dark; middle and hind femora with a preapical seta; fore tibia without evident bristles; middle tibia with a small anterodorsal and a small posterodorsal near basal third, 3 apicals; hind tibia with an anterodorsal and a posterodorsal near the basal third and a very small posterodorsal near distal third, 3 apicals. Lengths of segments of fore tarsus from base as 10—5-—4—3-3; middle tarsus as 12-6—-5-3-3; hind tarsus as 6-9-5—4—4. Wing rather oblong oval, clear with blackish veins; veins 3 and 4 nearly straight and parallel beyond the crossvein, vein 2 very slightly diverging; crossvein nearly perpendicular to last of vein 4, about half as long as last of vein 5; vein 6 represented by slight fold. Calypter and halter sometimes darkened, former with light brownish setae. Abdomen about as long as thorax, tapered to tip, blackish with grayish pollen toward base and sides; hairs and bristles brownish, paler toward sides of terga. Hypopygium (fig. 4) small, dark brownish, with apical surface oblique and bearing a tuft of stiff black setae in the middle, upper edge of the apex projecting. Female with face slightly wider, with a wider pale margin along the side. 334 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 Holotype ¢, allotype 2,9 ¢, and 6 2 paratypes, from moist shaded calcareous rock, in a small ravine about 5 km south of Tamazunchale, San Luis Potosi, Mexico, Aug. 4-5, 13-14, 1962, H. Robinson. Holotype ( No. 69347) and allotype in USNM; others presently in my collection. The species is very closely related to Micromorphus albipes Zetter- stedt but differs by the darker center of the face and by the more pointed hypopygium bearing a tuft of black setae. REFERENCES Parent, O. 1929. Etude sur les Dolichopodides exotiques de la collection von Réder. Ann. Soc. Sci. Brux. (B) 49: 169-246. Robinson, H. 1964. A synopsis of the Dolichopodidae (Diptera) of the South- eastern United States and adjacent regions. Misc. Publ. Ent. Soc. Amer. 4: 103- 192. DOLICHOMOTES NAVEI, A NEW GENUS AND NEW SPECIES OF PYEMOTID MITE (ACARINA: PYEMOTIDAE ) Rosert L. Smitey, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20250 The tarsonemine family Pyemotidae includes more than 120 known species, some of which are of medical or agriculture importance (Cross 1965). Krantz (1957) erected the genus Dolichocybe for a pyemotid mite that possesses unusual morphological characters: the hysterosoma which is divided into four segments with transverse striae separating the genital plate from the fourth pair of coxal plates; a pair of long posterior ventral whiplike setae; and an elongated gnathosoma. The genus here described is similar to Krantz’s and may be separated by the following characters: the reduction in body and leg setation, the longitudinal striation separating the coxae medially, and the absence of the pseudostigmatic organs. Dolichomotes, n. ¢. Type-species: Dolichomotes navei, new species. Gnathosoma of female oval and tapering; palpal tarsus possessing two distinct short claws and solenidion. Idiosoma long and slender, tapering, truncate posteriorly, and terminating with a pair of long strong simple setae ventrally. Legs I-IV similar in size and segmentation; each with two claws and empodium; tarsi I and II with a large striated disclike sensory seta (fig. 1); and tibiae I-IV similar by having a smaller sensory organ. Propodosoma without pseudostigmatic organs. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 335 Figs. 1-4, Dolichomotes navei, n. sp.: 1, tarsus I, dorsal, 9; 2, venter of gnathosoma, @; 3, venter, 2; 4, posterior ventral seta. Although this species lacks the globoid-shaped pseudostigmatic organs on the propodosoma, it has a pair of long slender sensilla arising from deep, specialized pits. The sensilla may be serving the same purpose as the pseudostigmatic organs. Cross (1965) reports that the pseudostigmata and the pseudostigmatic organs are not present in the genera Paracarophenax Cross, Acarophenax Newstead and Duvall, and Adactylidum Cross. 336 PROC, ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 — JZ) S\\or 3 =) | q oo Sw | Mt : 0 i J \ \4 \\ ray is Figs. 5-8, Dolichomotes navei, n. sp.: 5, dorsum, @; 6, leg I, 2; 7, lewis Fy, evel US 002 PROG. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 337 Dolichomotes navei, n. sp. (Figs. 1-8) Female. Body elongate and broadest in medial region of hysterosoma. Gnath- osoma as wide or slightly wider than long; palpal tarsus (fig. 2) with a pair of short curved claws, a microseta, and a solenidion; palpal tibia with long slender seta and microseta laterally. Propodosoma longer than wide; dorsally with two pairs of simple lateral setae subequal in length and a pair of longer slender simple sensory sensilla. Hysterosoma long and slender, tapering and becoming truncate posteriorly; with a pair of long strong simple caudal setae; dorsum divided into three distinct segments. First segment with two pairs of simple setae, a lateral and a medial pair. Second segment with one pair of lateral simple setae slightly longer than those of the first segment and medial setae consisting of base only and with a pair of anterior internal sclerotized structures. Third segment with two pairs of marginal simple lateral setae subequal in length and the remnants of a third pair medially; posterior and ventrally with a pair of long strong simple setae about as long as length of hysterosoma and a pair of microsetae at posterior lateral margin of genital plate. Ventral propodosomal and hysterosomal setae subequal in length. Coxal plates large and distinct, separated from each other medially by striae; transverse striae also in area between coxae IV and genital plate. Coxae I with a single pair of setae, slightly shorter than pair on coxae II; coxae III with single pair of setae slightly longer than pair on coxae IV. All tarsi (figs. 6-8) with two well developed claws and empodium. Tarsi I and II similar, with a large striated, disclike sensory seta, a microspur, a pair of simple dorsal setae distally, and a shorter pair of simple seta ventrally. All tibiae similar, each with a small, striated, disclike sensory seta similar to those on tarsi I and II. Legs III and IV similar, without striated disclike sensory seta on the tarsi. Body 145 uw long; 64 “ wide. Holotype. Female, U.S. National Museum No. 3226; collected on Areca catechu L., Hawaii, March 19, 1960 by R. Nave, for whom this species is named. REFERENCES Cross, E. A. 1965. The generic relationships of the family Pyemotidae (Acarina: Trombidiformes ). Univ. Kansas Sci. Bull. 45, No. 2: 29-275. Krantz, G. W. 1957. Dolicocybe keiferi, a new genus and new species of pye- motid mite, with a description of a new species of Siteroptes (Acarina: Pyemo- tidae). Ann. Ent. Soc. Amer. 50: 259-264. 338 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 NOTES ON THE GENUS THECODIPLOSIS KIEFFER IN NORTH AMERICA, AND SOME GENERIC REASSIGNMENTS (Dierera: CECIDOMYIIDAE ) RAYMOND J. GAcGNE, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560 No key to the North American genera of Cecidomyiidae will properly separate Thecodiplosis from Contarinia. Felt (1918, 1925, 1958) distinguished between the two genera on the basis of whether the costa is interrupted at its juncture with R;. The costa is invariably interrupted in species of both genera. The most reliable basis for the separation of the adults of the two genera lies in the female postabdomen: in both genera it is long and protrusible, but in Thecodiplosis brachyntera (Schwagrichen), the type-species, and in the species which I consider here to belong to the genus, it is not longitudinally striated, and the cerci are ovoid and fleshy; in Contarinia the postabdomen is longitudinally striated, and the cerci are long-attenuate, well-sclerotized, and closely juxtaposed laterally. Males of both genera are essentially alike: the tenth tergum and sternum are bilobed and subequal, and the aedeagus is triangular, distally attenuate, and subequal in length to the tenth segment. Foote (1965) listed nine species in the genus Thecodiplosis. Except for the recently named T. piniresinosae Kearby, that treatment of the group followed Felt (1918). Only four of the species actually belong there on the basis of the structure of the female postabdomen. These are: T. cupressiananassa (Osten Sacken), T. hudsonici (Felt), T. piniradiatae (Snow and Mills), and T. piniresinosae Kearby. Con- tarinia constricta Condrashoff, C. cuniculator Condrashoff and C. pseudotsugae Condrashoff, also belong to Thecodiplosis on the basis of the female ovipositors and larval characters. All the species now assigned to Thecodiplosis are associated with twigs or leaves of conifers. T. cockerelli Felt, reared from pine needles, is here transferred to Contarinia on the basis of the female characters discussed above. Thecodiplosis dulichii Felt fits well the description of Anthodiplosis Kieffer (Kieffer, 1913) and is here transferred to that genus. This species has been reared from seeds of a sedge, Dulichum sp. Antho- diplosis may be separated from all the other North American genera of Contariniini by the fact that the nodes on each flagellomere are unequal in size and the circumfilum on each node is rudimentary with very short loops. The female ovipositor is not protrusible, and each flagellomere of the female antenna has a distal neck two-thirds as long PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 339 as the node. The only other species of Anthodiplosis is A. rudimentalis Kieffer, known from Europe, and reared from heads of Artemisia vulgaris. T. liriodendri (Osten Sacken), causing ocellate leaf galls on the tulip tree, Liriodendron tulipifera L., is here transferred to Thomasin- iana because of the larval characters (Mohn, 1955). Adult males presumably reared from these galls by Felt, fit the description of the genus in Kieffer (1913). T. quercifolia (Felt), reared from an oak leaf gall made by Cynips sp., has three circumfila on each flagellomere and therefore belongs in another tribe, the Cecidomyiini. The proximal circumfilum on the distal node of each flagellomere is not developed into loops but girdles the node closely. The two other circumfila have well-developed loops, but the loops are much longer on one side of the node than on the other. The species is here referred to Bremia Rondani. T. zauschneriae Felt, reared from rosette galls on Zauschneria cali- fornica Presl., is here transferred to Contarinia on the basis of the female characters discussed above. REFERENCES Condrashoff, S. F. 1961. Three new species of Contarinia Rond. (Diptera: Cecidomyiidae ) in Douglas-fir needles. Can. Ent. 93: 123-130. Felt, E. P.. 1918. Appendix: a study of gall midges. VI. N.Y. State Mus. Bull. (1917) 202: 76-205. . 1925. Key to gall midges. (A resumé of Studies I-VII). N.Y. State Mus. Bull. 257: 1-239. . 1958. III. Subfamily Itonidinae (= Cecidomyiinae). Conn. State Geol. Nat. Hist. Survey Bull. 87:93-206. Foote, R. H. 1965. In Stone et al., A Catalog of the Diptera of America North of Mexico. U.S. Dept. Agr., Agr. Handbook 276: 241-295. Kieffer, J. J. 1913. Diptera. Fam. Cecidomyidae. Genera Insectorum, Fasc. 152: 346 pp., 15 pls. Mohn, E. 1955. Beitrage zur Systematik der Larven der Itonididae. Zoologica, Stuttgart 38(105): 1-247, 30 pls. 340 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 THE DISCOVERY OF LISTROPHORUS LEUCKARTI PAGENSTECHER ON MICROTUS PENNSYLVANICUS PENNSYLVANICUS FROM NORTH AMERICA‘ (ACARINA: LISTROPHORIDAE ) B. McDANIEL?, Jon P. SHOEMAKER® and Susan J. Joy® The mite species Listrophorus leuckarti Pagenstecher (the type species of Listrophorus Pagenstecher, 1861) was recorded from the meadow vole Microtus arvalis arvalis (Pallas). Recent material col- lected in Canada and the United States extends the distribution of this species to include the Nearctic region. A study of material from the Trouessart collection revealed the following distribution records: Argentina (on host Mus sylvaticus); India (on host Arvicola amphi- bius); and France (on host A. arvalis). All specimens had been collected by Rollinat and determined by Trouessart. No dates of collection were recorded. The French material from A. arvalis was reported by the late Dr. Marc Andre (personal communication ) to be homotypes of Trouessart, although not indicated as such. With these new records, L. leuckarti Pagenstecher has been found in the Palearctic, Nearctic, Neotropical, and Oriental realms. The following is the habitat range of L. leuckarti in Canada and the United States: Silverwater, Manitoulin Island, Canada, from Microtus pennsylvanicus pennsylvanicus collected by Tho Scholten on July 26, 1961, and Cabell and Mason Counties, West Virginia, United States, in August 1966, collected by Jon P. Shoemaker and Susan J. Joy. The material collected in Canada consists of a single male, six females and two larvae. The material from West Virginia are four males, three females and two larvae. All of these agree in structure with specimens from Argentina, France, and India. The following description and illustrations of L. leuckarti are made from Trouessart homotypes and specimens from West Virginia. Listrophorus leuckarti Pagenstecher Male—Body laterally compressed, elongate, narrowing posteriorly, dorsum striate (fig. 2). Legs well-developed. Hair-clasping organ of the typical listro- phorid type, with flap-like plates to clasp hairs of host. Opisthosomal length much less than distance between coxa of leg IV and coxa of leg I. Head well- developed, propodosomal and opisthosomal plates prominent. Head plate narrow- ing to a point at anterior end, extending beyond legs I; posterior portion projecting and partly enclosing coxa I. Propodosomal plates extending from dorsum of 1 Approved by the Director of the South Dakota Agricultural Experiment Station as Journal Series No. 766. * Department of Entomology-Zoology, South Dakota State University, Brookings, South Dakota. 3 Department of Zoology, Marshall University, Huntington, West Virginia. PROC. ENT. SOC. WASH., VOL. 69, No. 4, DECEMBER, 1967 341 Listrophorus leuckarti Pagenstecher. Fig. 1, lateral view of female; fig. 2, lateral view of male. body and covering region above second pair of legs but not connected to coxae II, separated from head plate by approximately the width of coxa II. Shape of prodosomal plates rhomboidal, with a single seta placed at the anterior section. Another seta of same shape and size as propodosomal setae between lower section of propodosomal plates and head plate on membranous portion of body. Middle section of dorsum strigose (fig. 2). Two setae similar in structure to propodosomal plate setae on striated region of body, placed between legs II and III, and the other between opisthosomal plate and coxa IV. Opisthosomal plate divided, each portion extending from anal region of body cephalad to coxa IV. The setae of each plate similar in structure to propodosomal plate setae, with a single seta on anterior portion and two on posterior portion of opisthosomal plate (fig. 2). Strial pattern of body around opisthosomal plates different from body region between legs II and anterior portion of opisthosomal plate (fig. 2). Venter of gnathosoma with listrophorid flap-like plates covering sternal area from coxae I to mouthparts. First pair of legs longer than second pair with typical listrophorid arrangement of segments, i.e., with the femur greatly elongated (longer than all other segments combined). Legs II smaller than others. All legs with sclerotized regions associated with apodemes, similar to the division of the propodosomal plate. Plates associated with legs II with a single seta, legs III having two pairs of setae; those with legs I and IV without setae. Two simple 342 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 setae between coxae I small, not visible when specimen mounted laterally. A pair of microseta; between apodemes of legs III and also between coxa IV. All legs with conicals. Male genitalia close to legs III on metapodosomal. Aedeagus distinct, well-developed, sickle-shaped, apex bent (fig. 2). Anal suckers small, associated with a sclerotized bar, the suckers and bars not plainly visible when specimen mounted laterally. Anal region with single pair of simple setae divided into two reduced lobes, heavily sclerotized; the divided nature not evident when specimen mounted laterally. Female—Twice as long as male, with head plate and propodosomal plate similar to male. Body laterally compressed, elongated. Distance between legs IV and posterior end of body subequal to distance between coxa I and coxae IV. Legs similar to those of male, with the first pair longer than second. Legs III and IV equal in length. Propodosomal plate same shape as male (fig. 1); setae similar in structure and placement. Propodosomal plate divided as in male, with two setae on main and largest division, one placed on most anterior section and the other at most posterior region. Striations of dorsum similar to male from propod- osomal plate to coxa IV. Striations from coxa of legs IV as shown in fig. 1. Dorsum with four pairs of setae, one pair placed near coxa IV, a second pair caudad of coxa IV approximately length of legs IV, and two pairs near anal section of body. Two pairs of simple setae associated with anal opening. Opis- thosomal region without dorsal plate. Genital area enclosed by apodemes of legs III, two small genital suckers present within arch of apodemes; all this region not visible when specimen mounted laterally. Coxal plates of legs III and IV sclerotized, well-developed. Entire venter with striations, similar to dorsum. All legs with conicals. The female may be distinguished by the shape of the propodosomal shield. Listrophorus leuckarti is separated from all other members of the genus by the shape of the aedeagus (fig. 2), the shape of the propod- osomal plate, type of body striations, and male opisthosomal plates. It should be noted that the females of the genus Listrophorus are less distinctive, and without the male, species identification is at times extremely difficult. However, the shape of the propodosomal plate and the dorsal striations are the most distinguishing characters of the females. In the key by McDaniel (1965), L. leuckarti was identified as L. americanus Radford. The separation of L. leuckarti and L. amer- icanus presently can be made only on the basis of the structure of the aedeagi. The senior author is presently working on the complex of the genus Listrophorus found on muskrats which involves L. validus, L. doxieri and L. americanus. Until this study has been completed no further distinctions in this complex are possible. ACKNOWLEDGMENTS The authors wish to express their appreciation to the late Professor Mare Andre, Laboratoire d’ Acarologie, France, for the loan of material of L. leuckarti from the Trouessart collection and to Th. Scholten, University of Victoria, Victoria, Canada, for specimens collected by him. The authors also wish to thank Drs., PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 343 R. J. Walstrom, E. U. Balsbaugh, Jr. and P. A. Jones of the Dept. of Entomology- Zoology, South Dakota State University for reading the manuscript; and giving of their time in order that this paper could be completed. This investigation was supported in part by a grant from the Marshall Research Foundation, Marshall University. REFERENCES McDaniel, B. 1965. The Subfamily Listrophorinae Gunther with a Description of a New Species of the Genus Listrophorus Pagenstecher From Texas ( Acarina, Listrophoridae ). Acarologia 7(4): 704-712. LINEAR COPULATION IN XYLOCELIA FRANCLEMONTI KROMBEIN ( HYMENOPTERA: SPHECIDAE ) NorMAN Lin, Department of Zoology and Entomology, The Ohio State University, Columbus, Ohio 43210 Linear copulation, in which both members of the pair are dorsal side up and form a straight or nearly straight line with heads facing away from each other, has seldom been reported in sphecid wasps. As far as I could determine such behavior has only been reported in the cicada killer wasp Sphecius speciosus (Drury) by Lin (1966), and Scullen (1965) has published a photograph of a pair of Cerceris frontata Say copulating in this position. Evans (1966) mentions only one other case of linear copulation among solitary wasps and that occurred in the eumenid Monobia quadridens (Linnaeus). The present account of copulation in Xylocelia franclemonti Krombein is another case of linear copulation and therefore seems worth reporting. Both the Raus (1918) and the Peckhams (1898) reported observing copulation in X. metathoracica Mickel and X. americana (Packard) respectively, but gave no description of the position. Three observers have noted instances of seemingly unsuccessful mating in species of Xylocelia. Thus, Krombein (1958) stated that several males of X. virginiana Rohwer attempted unsuccessfully to mate by pouncing on females on the ground. The Raus (1918) noted that such pouncings occurred frequently in X. metathoracica. Powell (1963) stated that X. occidentalis (Fox) pairs were often observed in what appeared to be unsuccessful mating attempts, but none were seen copulating. These encounters “consisted of a male riding on the back of a female as the latter walked about.” The wasps were oriented head over head, but contact of abdomens was not seen. These were at the most 10 or 20 second engagements and “usually terminated at some disturbance such as the pair falling from a grass stem or being bumped by another 344 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 male in flight.” Similar unsuccessful attempts at mating were observed in the present study. X. franclemonti (det. H. E. Evans) was found nesting in consider- able numbers from 1957 to 1966 in sandy tracts along baseball fields in the Parade Grounds, a huge field in Brooklyn, New York. Colonies of S. speciosus are located on these same tracts. The following observations were made during the summers of 1958, 1960, 1965, and 1966. A total of 14 unsuccessful copulatory attempts were observed on nine different days between 11:08 am and 5:34 pm and between June 18 and August 31. The single copulating pair was observed on July 25, 1965. As the data for X. metathoracica (Rau and Rau, 1918), X. virginiana (Krombein, 1958), and X. occidentalis (Powell, 1963) also suggest, unsuccessful attempts at copulation in X. franclemonti are seemingly more common than successful copulation. In all cases of apparent unsuccessful attempts at mating, a smaller male was mounted on, or mounted a larger female in the position described above by Powell. Five pairs in this position alighted on the ground, and two alighted on a fence. One of the pairs alighted on the ground several times and another on the fence several times. Two pairs were observed resting on plants and in the case of one, a still smaller male was mounted in the same position on the primary male. In seconds they flew away, the female carrying the two males. Five cases were observed of males in flight, pouncing on females to assume the above position. Once there was a brief struggle, the male assumed the usual position on the back of the female, and the pair then flew off, the female carrying the male. On a sixth occasion a pouncing male missed the female, landed next to her, and then flew away. Apparently the same female landed again about 30 seconds later and a male was successful in assuming the usual position on her back. At least three of the 14 pairs walked about considerably. Once a female walked up to a nest and, apparently because of the male on top of her, was unsuccessful in her attempted entry. She then flew, carrying the male with her. Unsuccessful attempts at copulation in X. virginiana were directed at females engaged in nesting activities (Krombein, 1958). A pair of X. franclemonti remained on the ground or some other surface for less than a minute on five occasions, and a pair was on a surface for more than a minute on two occasions, the longest being three or four minutes. This was considerably longer than the maximum of 20 seconds reported for X. occidentalis by Powell (1963). In five cases, a male X. franclemonti was observed to lash the female with his antennae while mounted on her. On two occasions males lashed the antennae of the female, on three occasions the head. In one of these latter cases the female repeatedly unfolded and folded her wings in what may have been an “attempt” to eject the male. She PROC. ENT. SOC, WASH., VOL. 69, NO. 4, DECEMBER, 1967 345 then flew off carrying the male with her, and they landed on a plant about eight inches away; they then flew off again. Seven pairs were observed to fly off together, the female carrying the male, and in some of these cases additional landings and take offs were observed. On one occasion a male alighted on a female, lashed her antennae with his, and after about one minute flew off. On a second occasion a male alighted on a female and then flew away. On July 25, 1965, a pair of X. franclemonti in linear copulation suddenly alighted on the ground in front of me. The male faced in a direction opposite that of the female. The male as in the previous cases was noticeably smaller and thinner than the female. The pair remained where they landed for about 30 seconds, when I placed my pencil next to them. They then climbed the pencil, the female pulling the male. The pair walked about the pencil considerably, the female continuing to pull the male behind her. The pencil was lifted four or more feet above the ground when the pair jumped off and in a spiral manner descended to the ground. Several aspects of apparent unsuccessful mating behavior have not previously been described in Xylocelia. These include the frequent carriage of the male in flight by the female, and the antennal lashing of the head or antennae of the female by the male. The precopulatory and copulatory behavior of X. franclemonti is similar to that of the cicada killer. Copulating cicada killers may fly. Females, usually noticeably larger than males, pull the males after them in walking and are never pulled by the males. Essentially the same precopulatory position is assumed as in X. franclemonti with the smaller male over the female, and mating may not take place at all. Nesting females will not mate as is also suggested in the present study in X. franclemonti and seems to be the case in X. virginiana (Krombein, 1958). Courtship includes the lashing of the female's antennae and head by the male’s antennae. A second male is some- times found in the precopulatory position on top of the first male. Flight frequently occurs in the precopulatory position, but the female is apparently unable to commence flight while carrying more than one male (Lin, 1963, 1966a, 1966b, and personal observation ). I am indebted to Dr. Howard E. Evans for identifying the Xylocelia. Funds for the summer of 1965 and 1966 were provided in part by a Sigma Xi Grant-in-Aid of Research. REFERENCES Evans, H. E. 1966. The behavior patterns of solitary wasps. Ann. Rev. Ent. 11: 123-154. Krombein, K. V. 1958. Miscellaneous prey records of solitary wasps. III (Hymenoptera, Aculeata). Proc. Biol. Soc. Wash. 71: 21-26. Lin, N. 1963. Territorial behavior in the cicada killer wasp Sphecius speciosus (Drury) (Hymenoptera: Sphecidae) I. Behaviour 20: 115-133. 346 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 1966a. Copulatory behavior of the cicada killer wasp Sphecius spe- ciosus. Animal Behaviour 14: 130-131. 1966b. Standard and emergency courtship in the cicada killer wasp Sphecius speciosus (Hymenoptera: Sphecidae). Amer. Zool. 6: 585. Peckham, G. W. and E. G. Peckham. 1898. On the instincts and habits of the solitary wasps. Wisc. Geol. Nat. Hist. Surv. Bull. 2, 245 pp. Powell, J. A. 1963. Biology and behavior of nearctic wasps of the genus Xylo- celia with special reference to X. occidentalis (Fox). (Hymenoptera: Spheci- dae). Wasmann Jour. Biol. 21: 155-176. Rau, P. 1935. The courtship and mating of the wasp Monobia quadridens. Ent. News 46: 57-58. and N. Rau. 1918. Wasp studies afield. Princeton University Press, Princeton, 372 pp. A NEW SALDID FROM CALIFORNIA (HEMIPTERA: SALDIDAE ) J. T. PoLnemus, 3115 S. York, Englewood, Colorado 80110 Several years ago, while studying the Usinger Collection, I separated a new species of Saldula. I had hoped to include it in a more com- prehensive study of the genus, but the need for the new name has prompted me to publish it alone. Saldula usingeri, n. sp. Of moderate size, moderately broad, general color black, macropterous. (For all measurements, 60 units = 1 mm.) Head: Black, feebly shining, surface almost obscured by dense, short, re- cumbent silvery pubescence; preocellar spot yellowish; with scattered long erect setae in addition to the usual three pairs of long hairs on frons and vertex; ocelli slightly raised and separated by the width of an ocellus; interocular space greater than the width of an eye (20:19); clypeus and anteclypeus testaceous medially, piceous elsewhere. Thorax: Pronotum black, feebly shining, rugulose, surface almost obscured by pubescence, vestiture as described for head; covered with long black erect setae, very evident at lateral margins when viewed from above; lateral margins very slightly convex to straight, narrowing moderately anteriorly; callus slightly raised, with deep pit in center; anterior lobe longer than posterior lobe (17:8, not including collar); collar wide (5); underparts black, clothed with fine silvery pubescence; scutellum equal in width and length (57:57), with vestiture similar to pronotum, depressed transversely across center. Wings: Hemelytra fully developed, pubescence and setae as on pronotum; ground color black; for most specimens, the inner corium and clavus are solid velvety black, and the outer corium is feebly shining and marked as in figure 1A; occasionally light spots are found on the inner corium along the corial suture and at the apex along the membrane; membrane fumose to deep fumose, with four cells. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 347 Yoram Yo mm Fig. 1, Saldula usingeri, n. sp.: A, left hemelytron; B and C, left male paramere, two views; D, parandria. Abdomen: Piceous, covered with short, decumbent silvery pubescence; in female, caudal portion of subgenital plate leucine, this plate produced caudad; parandria as shown in figure 1D. Extremities: Antennal segment 1 testaceous, with deep brown ventral basal stripe; segment 2 deep brown, testaceous on apical third or fourth; segments 3 and 4 deep brown to piceous; as thick as the apex of segment 2, relatively slender for the genus; all segments clothed with short pubescence and scattered longer hairs, the latter longest on segments 3 and 4; proportions as follows: Holotype ¢: 1;20 : II;42 : III;28 : IV:30. Allotype ¢: 1;20 : II;48 : III;29 : IV;28. Legs testaceous, coxae, base and apex of trochanters, base of femora flavous to flavo-testaceous; tibia and tarsal segments darkened apically, tibia with usual dark spines. Genitalia: Paramere as shown in figure 1B and 1C; filum gonopori coiled one and three fourths times; median sclerotized structures of aedeagus typically Saldula like. Measurements: Holotype ¢, length 3.65 mm., width 1.85 mm., allotype °, length 4.8 mm., width 2.4 mm. Mean length of 10 ¢ ¢: 3.84 mm.; min. 3.65 mm., max. 4.35 mm. Mean width of 10 ¢ ¢: 1.81 mm.; min. 1.65 mm., max. 2.0 mm. Mean length of 10 2 9: 4.20 mm.; min. 3.9 mm., max. 4.8 mm. Mean width of 10 2 @: 2.18 mm.; min. 1.9 mm., max. 2.4 mm. Material: Holotype, male, (R. L. Usinger Collection ), Calif., Colusa Co., Wilbur Spr., Aug. 16, 1960, R. L. Usinger collector; allotype, female, same locality but May 17, 1961, Wygodzinsky collector; 348 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 paratypes as follows: 9 ¢ 6,8 2 2, same data as holotype; 3 ¢ 3,4 22, 2 nymphs, same but May 17, 1961, Wygodzinsky collector; 1 ¢, 6 22, same but Sulphur Cr. at Wilbur Hot Spgs., Mar. 29, 1956, on shore in ‘Footbath’ pool area, H. B. Leech collector; 1 2, same as holotype but May 15, 1967, N. Ueshima collector; 3 ¢ ¢, 1 2, same as holotype but H. C. Chapman collector. Paratypes are in the collections of P. Wygodzinsky, R. L. Usinger, H. C. Chapman, J. T. Polhemus, and USNM. I am indebted to Drs. Wygodzinsky, Usinger and Chapman for the opportunity to study this material, and to the latter for helping in the diagnosis of this material. This species is named in honor of Dr. R. L. Usinger in recognition of his interest in and fine contributions to our knowledge of Saldidae. Comparative notes: Saldula usingeri n. sp. is close to Saldula pilosella (Thomson) of Europe. Both of these species have small eyes and a wide interocular space, the latter often exceeding the width of an eye in both species. In usingeri, however the front lobe of the pronotum is long compared to the posterior lobe (measured on a median line, not including the collar), the ratio for usingeri being anterior: posterior, 17.7: 10.8 (10 specimens) and for pilosella 12.9: 10.9 (10 specimens). In addition the pilosity of the paramere is much heavier in usingeri (fig. 1) and the paramere is of a different shape. The color of the specimens of the two species is quite different in the material examined, pilosella being much lighter. Cobben (1958) considers Saldula hirsuta (Reuter) a subspecies of S. pilosella. Material of hirsuta has not been examined, but as Cobben gives only the relative hairiness as the difference between the two forms of pilosella, it is clear that we are dealing with a different species in usingeri. Considering North American species, usingeri is closest to Saldula pexa Drake, however the parameres, parandria and eunomy are dif- ferent. The clavus always has a well defined apical spot in pexa, and never in usingeri, however in the lightest specimens of the latter there is a small, ill-defined light area. The most distinctive character of usingeri, however, is the wide interocular space due to the small eyes, particularly noticeable in large females where the head looks dis- proportionately small. Drake (1950) lists Saldula hirsuta (Reuter) as occurring in Cali- fornia, but almost certainly he was referring to either usingeri or pexa. REFERENCES Cobben, R. H. 1959. Notes on the classification of Saldidae, with the descrip- tion of a new species from Spain, Zool. Meded. Leiden 36: 303-316. Drake, C. J. 1950. Concerning North American Saldidae (Hemiptera), Bull. Brooklyn Ent. Soc. 45(1): 1-7. PROG. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 349 ECTOPARASITES OF CANADIAN BIRDS AND MAMMALS! Nrxon Wuson, Bernice P. Bishop Museum, Honolulu, Hawaii 96819 A small collection of ectoparasites from birds and mammals is listed with comments relative to host relationships and distribution in Canada. Important past references to each species in Canada are reviewed briefly. Eighteen species belonging to 7 orders are listed from 13 species of birds and mammals. Eight of these are new Canadian, provincial and/or host records. Specimens have been distributed between Joseph Moore Museum, Earlham College, Richmond, Indiana and Bernice P. Bishop Museum. Thanks are extended to Mr. W. W. Baker, Dr. R. E. Mumford and Mrs. G. L. Ward for providing most of the material for study. List of Ectoparasites Class ACARINA Order MrETASTIGMATA Family IxopmaEr Haemaphysalis leporispalustris (Packard ) 1 6,1 92, ex Lepus americanus Erxleben, New Brunswick, Charlotte Co., Kent I., 25.VI1.1963, I. L. Brisbin (FN 29). This is the tick encountered most often on leporids in North America. Gregson (1956) tabulated many records from Canada including four from New Brunswick. Ixodes angustus Neumann 1 2, ex Tamias striatus (Linnaeus), Ontario, Thessalon, 64 km N, 26.VII.1965, J. B. Cope (JBC 581). According to Gregson (1956) this species is best known from the western provinces but is found throughout Canada. He listed it from this host in Ontario. Order MrsosTIGMATA Family HAEMOGAMASIDAE Eulaelaps stabularis (Koch) 1 9, ex Sorex cinereus Kerr, Saskatchewan, Regina, 3 km SE, 28.VIII.1959, N. Wilson (E 1156); 1 9, ex Microtus pennsylvanicus (Ord), Saskatchewan, Regina, 3 km SE, 28.VIII.1959, N. Wilson ( E 1157). E. stabularis is a widespread species recorded mostly from rodents and insec- tivores. This is the first record from Saskatchewan and from S. cinereus. All previous Canadian records are from Ontario (Banks, 1909; Jameson, 1950; Judd, 1950). Haemogamasus ambulans (Thorell ) 1 9, ex Eutamias minimus (Bachman), Ontario, Chapleau, 32 km E, VII.1965, W. H. Buskirk and J. B. Cope. 1 This investigation was supported in part by Public Health Service Research Grant AI-01723 from the National Institutes of Allergy and Infectious Disease, National Institutes of Health, Public Health Service. 350 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 This species is known from the northern United States, Canada and Alaska under the name Haemogamasus alaskensis Ewing. Rodents and insectivores are the common hosts. It never has been reported from E. minimus but there are records from Ontario (Jameson, 1950; Keegan, 1951). The record of Haemo- gamasus hirsutus Berlese from Ontario by Judd (1950) may be this species. Family LAELAPIDAE Androlaelaps fahrenholzi ( Berlese ) 1 N, ex Microtus pennsylvanicus, Saskatchewan, Regina, 3 km SE, 28.VIII.1959, N. Wilson (E 1157). This is a widespread species occurring most frequently on rodents. It has been recorded from Canada by Strandtmann (1949), Jameson (1950), Judd (1950, 1953) and Scholten et al. (1962), but never from Saskatchewan. Laelaps alaskensis Grant 3 29, ex Microtus pennsylvanicus, Ontario, Attawapiskat, 7.VII.1965, J. B. Cope et al. (JBC 550). This rodent mite has been reported from Keewatin and Ellef Ringnes Island in Canada (Harper, 1956; McAlpine, 1964; 1965). Records of Laelaps muris (Ljungh) from Ontario by Judd (1950; 1953; 1954) are probably this species. Order PRrostTiGMATA Family TROMBICULIDAE Euschongastia sp. 40 LL, ex Microtus pennsylvanicus, Saskatchewan, Regina, 3 km SE, 27.VIII. 1959, N. Wilson (E 1155). All specimens lack sensillae making specific identification impossible. They are very close to E. campi Brown and Brennan and E. sciuricola (Ewing) both of which have been collected in Canada (Brown and Brennan, 1952; Farrell, 1956). Class INsECTA Order ANOPLURA Family HorLopLEURIDAE Hoplopleura acanthopus ( Burmeister ) 1 6, 2 29, ex Microtus pennsylvanicus, Saskatchewan, Regina, 3 km SE, 98.VIII.1959, N. Wilson (E 1157). H. acanthopus is a common ectoparasite of Microtus in North America and has been reported previously from Canada (Ferris, 1921; Harper, 1956; Scholten et al., 1962; McAlpine, 1964, 1965; Spencer, 1966). Order MALLOPHAGA Family MENOPONIDAE Myrsidea anaspila ( Nitzsch) 2, 2 9, ex Corvus corax Linnaeus, Ontario, Moosonee, 28.VI.1965, J. B. Cope; 2 99,1N, ex C. corax, Ontario, Moosonee, 15.VII.1965, J. B. Cope and W. H. Buskirk. Neither Spencer (1948, 1957), Stirrett (1952) nor Whitehead (1954) listed this species from Canada; however, it is to be expected on the common raven. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 351 Trinoton querquedulae (Linnaeus ) 1 ¢, ex Anas carolinensis Gmelin, Manitoba, Delta, 13.VIII.1964, D. A. Manuwal. This mallophagan occurs on several species of ducks and has been collected from this locality on this host (Buscher, 1965). There are other Canadian records from British Columbia (Spencer, 1948), Ontario (Stirrett, 1952) and Quebec (Whitehead, 1954). Family PHILOPTERIDAE Saemundssonia sternae (Linnaeus ) 1 @, ex Sterna hirundo Linnaeus, Manitoba, Delta, 13.VIII.1964, D. A. Manuwal (REM 4501). This species is expected to occur wherever the common tern does, although apparently it has not been collected in Canada. Order SIPHONAPTERA Family CERATOPHYLLIDAE Megabothris atrox (Jordan) 4 292, ex Mustela erminea Linnaeus, Ontario, Moosonee, 14.VII.1965, W. H. Buskirk (B). Holland (1949) mentioned only two records of this species; that of Jordan’s from Alberta and a new one from Northwest Territories. The above record extends the range about 2,300 km eastward in Canada. The few collections indicate this flea is a parasite of weasels. Three of four specimens varied from the original description in that they had fewer pronotal spines (23, 23, 24, 26). Jordan (1925) indicated there were 26 to 29 spines in the pronotal ctenidium of his specimens which suggested to him that it might be a bird flea. Megabothris quirini ( Rothschild ) 2 64,1 2, ex Mustela erminea, Ontario, Moosonee, 14.VII.1965, W. H. Buskirk (A); 1 4, ex M. erminea, Ontario, Moosonee, 14.VII.1965, W. H. Buskirk (133). This flea is normally a parasite of cricetid rodents and the present records may reflect predation by weasels on these rodents. This is a widespread species in Canada and Holland (1949) listed several records from Ontario. Monopsyllus eumolpi eumolpi ( Rothschild ) 1 g, ex Eutamias minimus, Ontario, Chapleau, 32 km E, VII.1965, W. H. Buskirk and J. B. Cope. This chipmunk flea probably occurs throughout the range of Eutamias in Canada. Holland (1949) and Hubbard (1949) each reported three records from Ontario. Monopsyllus vison (Baker ) 1 6, ex crawling on Homo sapiens Linnaeus, Ontario, Moosonee, 12.VII.1965, W. H. Buskirk. 2 ¢ 6, ex Mustela erminea, Ontario, Moosonee, 14.VII.1965, W. H. Buskirk (B). This species is widespread in Canada with Tamiasciurus hudsonicus (Erxleben ) the true host. The specimen from man probably was acquired while handling S52, PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 mammals and those from a weasel reflect the predatory nature of this animal. Holland (1949), Hubbard (1949) and Scholten et al. (1962) listed several records from southern Ontario. Orchopeas caedens durus (Jordan) 2466, 2 92, ex Mustela erminea, Ontario, Moosonee, 14.VII.1965, W. H. Buskirk (B). The subspecies durus has a widespread but discontinuous distribution in Canada. Its true host is T. hudsonicus and the above record probably indicates predation by the weasel on red squirrels. There are several records of this species from Ontario (Holland, 1949; Judd, 1954; Scholten et al., 1962). Family IscHNOPSYLLIDAE Myodopsylla insignis ( Rothschild ) 1g, 1 @, ex Myotis lucifugus (Le Conte), New Brunswick, Charlotte Co., Kent I., 17.VIII.1963, I. L. Brisbin (FN 54). M. insignis is widely distributed in Canada from inland British Columbia to New Brunswick, from which latter province the above record is the first. Bats of the genus Myotis, especially M. lucifugus, are recorded most frequently as hosts. Order DipTERA Family HippoBoscipAE Ornithomya fringillina Curtis 1 4, ex Zonotrichia albicollis (Gmelin), Ontario, Moosonee, 15.VII.1965, J. B. Cope and W. H. Buskirk; 1 9, ex Passerella iliaca (Merrem), Ontario, Moosonee, 16.VII.1965, J. B. Cope; 1 ¢, ex clothing, Ontario, Moosonee, 17.VII.1965, W. H. Buskirk; 1 9, ex Z. albicollis, Ontario, Moosonee, 21.VII.1965, J. B. Cope. This is a common hippoboscid of certain species of passeriform birds in temperate North America and there are numerous records from Canada, including Ontario, (Bequaert, 1954; Davies, 1958; Bennett, 1961). The fox sparrow apparently is not one of the preferred hosts as Bequaert (1954) published only one verified record. REFERENCES Banks, N. 1909. New Canadian mites. Proc. Ent. Soc. Wash. 11(3): 133-143. Bennett, G. F.. 1961. On three species of Hippoboscidae (Diptera) on birds in Ontario. Can. J. Zool. 39: 379-406. Bequaert, J. C. 1954. The Hippoboscidae or louse-flies (Diptera) of mammals and birds. Part II. Taxonomy, evolution and revision of American genera and species. Ent. Amer. 34(N.S.): 1-232. Brown, J. H. and J. M. Brennan. 1952. A note on the chiggers (Trombiculi- dae) of Alberta. Can. J. Zool. 30: 338-343. Buscher, H. N. 1965. Ectoparasites from anseriform birds in Manitoba. Can. J. Zool. 43: 219-221. Davies, D. M. 1958. Notes on avian hippoboscids. J. Parasit. 44(2): 239-240. Farrell, C. E. 1956. Chiggers of the genus Euschongastia (Acarina: Trombicu- lidae) in North America. Proc. U.S. Natl. Mus. 106 (3364): 85-235. Ferris, G. F. 1921. Contributions toward a monograph of the sucking lice. Part II. Stanford Univ. Publ., Univ. Ser., Biol. Sci. 2(2): 53-133. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 353 Gregson, J. D. 1956. The Ixodoidea of Canada. Can. Dept. Agr. Publ. 930, 92 pp. Harper, F. 1956. The mammals of Keewatin. Univ. Kans. Mus. Nat. Hist., Misc. Publ. No. 12, 94 pp. Holland, G. P. 1949. The Siphonaptera of Canada. Can. Dept. Agr. Publ. 817, Tech. Bull. 70, 306 pp. Hubbard, C. A. 1949. Fleas in the collection of the Royal Ontario Museum of Zoology. Can. Ent. 81(1):; 11-12. Jameson, E. W., Jr. 1950. The external parasites of the short-tailed shrew, Blarina brevicauda (Say). J. Mammal. 31(2): 138-145. Jordan, K. 1925. New Siphonaptera. Novit. Zool. 32: 96-112. Judd, W. W. 1950. Mammal host records of Acarina and Insecta from the vicinity of Hamilton, Ontario. J. Mammal. 31(3): 357-358. 1953. Mammal host records of Acarina and Insecta from the vicinity of London, Ontario. J. Mammal. 34(1): 137-139. . 1954. Some records of ectoparasitic Acarina and Insecta from mam- mals in Ontario. J. Parasit. 40(4): 483-484. Keegan, H. L. 1951. The mites of the subfamily Haemogamasinae (Acari: Laelaptidae). Proc. U.S. Natl. Mus. 101 (3275): 203-268. McAlpine, J. F. 1964. Arthropods of the bleakest barren lands: composition and distribution of the arthropod fauna of the Northwestern Queen Elizabeth Islands. Can. Ent. 96( 1-2): 127-129. . 1965. Insects and related terrestrial invertebrates of Ellef Ringnes Island. Arctic 18(2): 73-103. Scholten, T. H., K. Ronald, and D. M. McLean. 1962. Parasite fauna of the Manitoulin Island region. I. Arthropoda parasitica. Can. J. Zool. 40: 605-606. Spencer, G. J. 1948. Some records of Mallophaga from British Columbia birds. Proc. Ent. Soc. Brit. Columbia 44: 3-6. . 1957. Further records of Mallophaga from British Columbia birds. Proc. Ent. Soc. Brit. Columbia 53: 3-10. 1966. Anoplura from British Columbia and some adjacent areas. J. Ent. Soc. Brit. Columbia 63: 23-30. Stirrett, G. M. 1952. Mallophaga collected from birds in Ontario. Can. Ent. 84(7): 205-207. Strandtmann, R. W. 1949. The blood-sucking mites of the genus Haemolae- laps (Acarina: Laelaptidae) in the United States. J. Parasit. 35(4): 325-352. Whitehead, W. E. 1954. Avian Mallophaga from Quebec. Can. Ent. 86(2): 65-68. 354 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 A NEW SPECIES OF OMANIA FROM MICRONESIA (HEMIPTERA: SALDIDAE) Jon L. Herrinc and Haroip C, CHapMAn, Entomology Research Division, ARS, U.S. Department of Agriculture+ The following new species is the first member of the genus Omania Horvath from Micronesia. Previously, four species were known: coleoptratra Horvath, 1915 from the Sultanate of Oman and the Sinai Peninsula, marksae Woodward, 1958 from intertidal coral rocks of the Great Barrier Reef of Australia, samoensis Kellen, 1960 from American Samoa and satoi Miyamoto, 1963 from the Tokara Islands south of Kyushu, Japan. The present species was collected at Nauru Island in southeastern Micronesia. All of the members of the genus are extremely small and secretive and have thus far eluded discovery on most of the islands of Oceania. Both Woodward (1958) and Kellen (1960) have given excellent accounts of the life history, habits and adaptations of these saldids to a marine life in the intertidal zone. Drake (1961) has summarized the literature and included a splendid illustration of O. marksae. Key TO THE SpEcIES OF Omania HorvATH 1. Eyes small, width two-thirds or less the width of the vertex at level of ocelli, length subequal to length of pronotum; vertex with silvery pu- bescence; hemelytra almost unicolorous —....___»_-___»_ 2 Eyes large, width four-fifths or more the width of the vertex at level of ocelli, length greater than length of pronotum; vertex with golden pu- bescence; hemelytra with a contrasting pattern of bluish bloom and black — 4 2. Collum of head distinctly triangular; all antennal segments concolorous light brown; calluses of pronotum flat, poorly delimited; pronotum, scutellum, and hemelytra uniformly bluish gray; legs light brown _ coleoptrata Horvath Collum of head not triangular, anterior demarcation arcuate; antennal seg- ments III and IV dark brown or black; calluses of pronotum slightly raised and clearly demarcated; pronotum, scutellum and hemelytra velvety: blast." oie wee eee eee 3 3. Antennal segment III only one-half as long as IV, apices of segment I and II fuscus, all of III and IV black; posterior lobe of pronotum about twice as long as the anterior collar; hemelytra uniformly dull black _ satoi Miyamoto Antennal segment III approximately two-thirds as long as IV, apex of seg- ment II and all of III and IV brownish black; posterior lobe of the pronotum only slightly longer than anterior collar; hemelytra marked with a bluish bloom along costal margin and near middle of disk — ib Re ee Ree Ree ee ee i eee ee marksae Woodward 4. Length of eye almost one and a half times length of pronotum, its posterior margin extending beyond the posterior margin of the pronotal collar; 1 Washington, D.C. and Lake Charles, Louisiana, respectively. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 355 median fovea of callus deep, almost completely separating callus into two swollen lobes; scutellum very small, distinctly less than twice as wide as long; irregular silvery blue pattern occurring extensively over pronotum, Geeta lhonaay euavel InyercrslKyiade) oe nauruensis, nN. sp. Length of eye no more than one and a quarter times length of pronotum, its posterior margin not surpassing the posterior margin of the pronotal col- lar; median fovea of callus distinct but not dividing the callus into two separate lobes; scutellum twice as wide as long; irregular silvery blue pattern confined to sides of pronotum and three stripes on hemelytra Be iy a ta, Ree ee Ree samoensis Kellen Omania nauruensis, 0. sp. (Figs. 1-2) Male.—Head in front vertical, strongly deflexed, not projecting in front of eyes; over twice as wide as long, 40: 17°. Eyes large, four-fifths the width of vertex at level of ocelli; almost one and a half times length of pronotum on median line, 23: 16, their hind margins surpassing the hind margin of the pronotal collar; vertex, frons and clypeus shining black, finely rugulose; pubescence fine, golden. Ocelli raised, separated from each other by twice the distance between ocellus and eye. Anterior pair of cephalic trichobothria large, closely resembling ocelli but pale, bearing the usual pair of long, black setae. Posterior hairless collum on median line one-fourth total length of head, 4:17, with dull blue-gray pruinosity; anterior margin broadly arcuate. Clypeal region and underside of head with long, pale hairs. Rostrum testaceous, reaching middle coxae. Relative lengths of antennal segments I-IV, 9: 13: 11: 16, with numerous hairs; segments I and II pale, III and IV uniformly reddish brown. Pronotum hardly more than one-half as long as wide at base, 16: 33; lateral margins rounded and evenly tapered to anterior collar; posterolateral angles bluntly angled but not carinate or explanate; collar about one-sixth of total length, 2.5: 16; callus prominently elevated and clearly demarcated by a sulcus at sides and behind, median fovea continued as a sulcus which almost completely divides callus into two lobes; pronotal margins and median sulcus with blue-gray pruinosity. Scutellum small, over one and a half times wider than long, 16: 10; one-half as wide as base of pronotum, 16: 33; disk depressed on sides leaving a longitudinal carina joined to the transverse basal one, clearly defined by the blue-gray pruinosity. Forewings completely covering sides of abdomen and extending to level of sterna; dark, entirely coriaceous, elytron-like; not over-lapping and without a trace of membrane; costal margins broadly convex, apices rounded; surpassing abdomen, slightly diverging. Claval suture present, weakening apically, but clearly defined by preapical pruinose band. Blue-gray pruinose areas as follows: a large basal and preapical band on clavus, an irregular patch on base of corium extending narrowly down the lateral margin of hemelytra, widening at middle third and extending inward almost to commissure, a broad band at apical margin. Hind wings absent. (oo nits) —) lami: 356 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 Fig. 1, Omania nauruensis, n. sp., holotype ¢. Pronotum, scutellum and hemelytra impunctate, with rather sparse covering of short, curved hairs, these becoming longer laterally. Coxae and most of femora shining testaceous; apical fifth or sixth of each femur, all tibiae, tarsi stramineous with abundant long hairs; tibiae with some stiff spines. Genital capsule small with a pair of minute claspers (fig. 2). Total length 1.20 mm., greatest width of hemelytra 0.57 mm. Female very similar to male but larger. Length 1.36 mm., greatest width of hemelytra 0.68 mm. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 357 Fig. 2, é clasper. Holotype male (USNM 69554) and allotype female, Nauru Island, 432 miles west of the Gilbert Islands and just south of the equator, Nov. 25, 1966; 137 paratypes, same locality but Nov. 16, 19, and 25, 1966. All collected by H. C. Chapman. Holotype, allotype and 35 paratypes in the USNM. Paratypes in John T. Polhemus collection and H. C. Chapman collection. O. nauruensis can be distinguished from the other four members of the genus by its large, elongate eyes and the distinctive and apparently constant color pattern. CoLLectinc NOTES Omania nauruensis was collected at low tide from coral pinnacles that were common in the reef which completely surrounds Nauru Island. Most collections were made in the vicinity of Anibare Bay. The coral outcroppings varied in size from a foot to more than eight feet in height and many were very broad (fig. 3). Fig. 4 shows a coral pinnacle which generally produced specimens of this saldid. Speci- mens were collected by flushing or splashing water against the coral out-croppings just above the water where the surface was still moist from the previous high tide. The specimens were washed into the water where they were easily collected by a dipper or finger tip and then placed in a vial of alcohol. Because of the small size of this species, their collecting was facilitated by splashing pinnacles which were in the open sunlight and had white sand at their bases. Specimens appeared rather helpless in the water; if they were not collected, they eventually returned to a pinnacle. O. nauruensis apparently comes out of the interstices of the coral outcroppings at definite times during low tides. Collecting of this species was rather poor immediately following low tides. 358 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 Fig. 4, Coral pinnacle which often produced Omania nauruensis. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 359 Halovelia marianarum Usinger was the only other aquatic bug noted in the general habitat. This marine veliid was often seen around and on portions of the coral pinnacles. REFERENCES Drake, C. J. 1961. Hemiptera: Saldidae. Insects of Micronesia 7(6): 287-305. B. P. Bishop Museum. Horvath, G. 1915. Novum genus peculiare Acanthiidarum. Ann. Mus. Nat. Hung. 13: 598-599. Kellen, W. R. 1960. A new species of Omania from Samoa, with notes on its biology (Heteroptera: Saldidae). Ann. Ent. Soc. Amer. 53: 494-499. Miyamoto, S. 1963. New halophilus saldids from the Tokara Islands. Sieboldia 3: 39-48. Woodward, T. W. 1958. Studies on Queensland Hemiptera. Part III. A re- markable new intertidal saldid. Univ. of Queensland papers 1(7): 101-110. THE NEARCTIC DORYCTINAE, V. THE GENUS LELUTHIA AND COMMENTS ON THE STATUS OF THE TRIBE HECABOLINI (HYMENOPTERA, BRACONIDAE ) Paut M. Marsu, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20560 The Doryctinae have been divided into two tribes depending upon the presence of the second intercubitus in the forewing and, hence, three cubital cells (Doryctini), or the absence of the second inter- cubitus, hence, two cubital cells (Hecabolini). Tobias (1961) de- scribed a genus and species of Doryctinae which usually had two cubital cells but in which several specimens had three. After further study, and when he found no other characters to distinguish the Hecabolini and Doryctini, he concluded that “the presence or absence of the second intercubitus can not serve as a reliable criterion for the separation of these groups.” During my study of the genus Leluthia Cameron which normally has three cubital cells, I observed one distinct species and several specimens of another with only two cubital cells. Furthermore, one specimen has the right wing with two cells and the left with three. Otherwise, the specimens are identical and, I believe, congeneric. I agree with Tobias that this character of the forewing is not reliable 360 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 for tribal separation and that, therefore, the two tribes should be combined. Leluthia Cameron Leluthia Cameron, 1887, p. 392. Type-species: Leluthia mexicana Cameron. Desig. by Viereck, 1914. Russellia Muesebeck, 1950, p. 78. Preoce. by Vargas, 1943. Type-species: Hetero- spilus (?) astigma Ashmead. Orig. desig. New synonymy. Russellella Muesebeck and Walkley, 1951, p. 178. New name for Russellia Muesebeck. Distinguished by the following characters: head subcubical; first flagellar segment longer than second; forewings with two or three cubital cells, radial cell short, usually not nearly attaining apex of wing, recurrent vein entering first cubital cell, sometimes at extreme apex, subdiscoideus nearly on a straight line with discoideus, first brachial cell open at apex; foretibia with row of 5-10 small stout spines; tergum (243) with a tranverse strongly bisinuate groove and short shallow oblique grooves from base of tergum which set off a circular or oval area medially at base of tergum (fig. 10). Cameron (1887) included two species in Leluthia, mexicana and fuscinervis, the types of which are in the British Museum. Sub- sequently Viereck (1914) designated mexicana as the type-species of the genus. The other species, fuscinervis, is not congeneric with mexicana and, in fact, belongs to the genus Heterospilus. Muesebeck (1950) described the genus Russellia to include his species prosopidis and astigma Ashmead. These species are definitely congeneric with Leluthia, and prosopidis is identical with the type-species, mexicana. Key to Nearcric SpreciEs oF Leluthia Soap ere ae oe ieee eae EA 2 Malesia. ad ter Labs 2 ies et be! oe eee 5 2. Second intercubitus absent, forewing with two cubital cells — 3 Seco” intercubitus present, forewing with three cells 4 3. Ovip.:itor as long as abdomen; hindfemur about four times longer than its Pre neste welt es eee eel LT Sh eee floridensis, n. sp. Ovipositor shorter than abdomen; hindfemur about three times longer than RCAC RIVER eee ee astigma (Ashmead ) 4. Vertex transversly rugosopunctate; thorax stout, slightly longer than high Pee AU ORD et Le Re eT eS ee mexicana Cameron Vertex punctate or granular; thorax flattened dorsoventrally, usually about touiite sasilOmcewag abi ein a. 8 pits heel Peas te eee astigma (Ashmead ) 5. Stigma present in hindwing (fig. 8); if rarely absent, then vertex trans- VETSC IN TUPOSOPUNGCtALG oe A eee mexicana Cameron Stigma absent in hindwing; vertex punctate or granular _. 6 6. Second intercubitus absent, forewing with two cubital cells; hindfemur about four times longer than its greatest width _..-- floridensis, n. sp. Second intercubitus present; if rarely absent, then hindfemur about three times longer than greatest width __._____________ astigma (Ashmead ) PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 361 Figs. 1-3. Wings of Leluthia species: fig. 1, astigma, 2, fore and hindwing; fig. 2, astigma, 6, hindwing; fig. 3, floridensis, 2, fore and hindwing. Figs. 4-7. Male genitalia: fig. 4, astigma, genital capsule; fig. 5, astigma, ninth sternum; fig. 6, floridensis, genital capsule; fig. 7, floridensis, ninth sternum. Leluthia astigma (Ashmead), n. comb. (iar, Il, 25 2h by) Heterospilus (?) astigma Ashmead, 1896, p. 215. Holotype male, USNM 69555. Russellia astigma (Ashmead), Muesebeck, 1950, p. 78. Russellella astigma (Ashmead), Muesebeck and Walkley, 1951, p. 178. Female——Length of body, 3-6 mm, ovipositor, 1-2 mm. Color usually entirely dark brown or black with legs lighter; often legs and basal antennal segments bright honey yellow, and body marked with reddish brown. Head cubical, flat- tened dorsoventrally; entirely finely granular or punctate, lower part of temples occasionally smooth; malar space one-third eye height; temples about one-half eye width; antennae 25-32 segmented. Thorax flattened dorsoventrally, often very much so; pronotum short, somewhat swollen; propleuron granular; propleural groove distinct, weakly foveolate; mesonotum not sharply declivous anteriorly; mesonotal lobes finely granular; notauli weak, rugose, obscured behind by wide rugose area; scutellar furrow with 7 cross carinae; scutellar disc granular; meso- pleural disc and mesosternum granular; mesopleural furrow usually smooth; 362 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 subalar groove shallow, often rugose; propodeum coarsely granular, often longi- tudinally rugosopunctate medially. Hindcoxae and all femora granular; hindfemora about three times longer than greatest width. Wing venation as in fig. 1; stigma usually about four times as long as wide; second intercubitus occasionally absent. First abdominal tergum longer than wide, longitudinally rugosopunctate; tergum (2+3) with area in front of transverse groove short and broad, more often oval than circular; remainder of terga granular; ovipositor about two-thirds abdominal length. Male.—Essentially as in female; hindwing without stigma (fig. 2); genitalia as in fig. 4, ninth sternum narrowed and truncate anteriorly (fig. 5). Type Locality—Morgantown, West Virginia. Distribution.—Arizona, California, Iowa, Maryland, North Carolina, Ohio, Oklahoma, Pennsylvania, Quebec, Texas, Utah, Virginia, West Virginia, Wyoming; Mexico. Hosts.—Agrilus sp., A. difficilis Gory, (?) A. politus (Say). The specimens from Oklahoma, Texas, California, and Mexico are generally flatter and lighter than the other more northern specimens. However, I can find no significant differences and feel that they are members of this species. Leluthia floridensis, n. sp. (Figs.G, G6, 7) Female.—Length of body, 4-5 mm, ovipositor, 2-2.5 mm. Head, thorax, and first abdominal tergum, and tergum (2+3) medially at base, dark brown, often black; legs and remainder of abdomen brown. Head flattened dorsoventrally; eyes large; malar space about one-fourth eye height; temples about one-half eye width; vertex flat, not declivous posteriorly, transversely rugulopunctate; face, temples, and frons punctate; antennae 26-29 segmented. Thorax flattened dorsoventrally; prothorax punctate; propleural groove shallow, rugulose; mesonotal lobes punctate; notauli shallow, weakly foveolate; obscured behind by wide rugose area; scutellar furrow narrow, with 7 cross carinae; scutellar disc flat, punctate; mesopleural disc and mesosternum punctate; mesopleural furrow shallow, weakly foveolate, usually as long as mesopleuron; subalar groove shallow, wide, rugose; propodeum nearly horizontal, completely longitudinally rugose. Legs punctate; hindfemora usually about four times longer than wide. Wing venation as in fig. 3; second intercubitus always absent; radial cell somewhat longer than in other species but not reaching wing tip. First abdominal tergum about as long as apical width, longitudinally rugose, without a raised median area; tergum (2+3) granular behind transverse sinuate groove; remainder of abdominal terga granular; ovipositor equal to length of abdomen. Male.—Similar to female; length 3-4 mm; genitalia as in fig. 6; ninth sternum pointed anteriorly (fig. 7); hindwing without stigma. Holotype female.—Florida: Polk County, August 31, 1961, ex citrus limbs. USNM 69553. Paratypes.—Florida: Polk County, 6 22, 3 66, August 28 and September 5, 1961, ex citrus limbs; Lake Alfred, 1 2, August 21, 1961. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 363 Figs. 8-12. Leluthia mexicana: fig. 8, fore and hindwings, ¢; fig. 9, hindwing, 9; fig. 10, tergum (2+3); fig. 11, genital capsule, ¢; fig. 12, ninth sternum, ¢. Host.—One specimen was reared from citrus limbs infested with the weevil Cryptorhynchus fallax LeConte. This species is so similar morphologically and in general habitus to the type-species, mexicana, that I feel it undoubtedly belongs in Leluthia. The presence of only two cubital cells in the forewing would have placed this species in the tribe Hecabolini; but, in the light of the brief discussion above, this tribe has no validity. Although the male genitalia of floridensis are somewhat different, I do not think that this is enough to warrant a separate generic placement. 364 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 Leluthia mexicana Cameron (Figs. 8-12) Leluthia mexicana Cameron, 1887, p. 392. Holotype female in the British Museum. Russellia prosopidis Muesebeck, 1950, p. 78. Holotype female, USNM 59483. New synonymy. Russellella prosopidis (Muesebeck), Muesebeck and Walkley, 1951, p. 178. Female—Length of body, 3-6 mm, ovipositor, 1.5-3.5 mm. Color dark brown or black except lower part of head, prosternum, and legs which are often lighter brown. Head subcubical; vertex and frons transversely rugosopunctate, face rugulose, temples granular; malar space about one-third eye height; temples about three-fifths eye width; antennae 22-30 segmented. Thorax stout, not flattened; pronotum short, propleuron rugose; propleural groove not distinct; mesonotum declivous anteriorly; mesonotal lobes granular; notauli represented by wide shallow rugose lines, obscured posteriorly by wide rugose area; scutellar furrow with 7 cross carinae; scutellar disc granular; mesopleural disc punctate; mesopleural furrow deep, finely foveolate; subalar groove broad and strongly rugose; meso- sternum punctate; propodeum completely rugose, no indication of carinae. Hind- coxae and all femora granular. Forewing venation as in fig. 8; hindwing, fig. 9; stigma of forewing about three times as long as wide; second segment of radius short, sometimes much shorter than first segment. First abdominal tergum about as long as broad at apex, longitudinally rugose, without raised median area; tergum (2+3) finely rugosopunctate behind transverse groove (fig. 10); remainder of terga granular; ovipositor about as long as abdomen. Male.—Essentially as in female; length 2.5—4.5 mm; hindwing with a stigma, (fig. 8), rarely without one; genitalia as in fig. 11; ninth stemum narrowed and truncate (fig. 12). Type Locality—Northern Sonora, Mexico. Distribution—Arizona, California, New Mexico, Oklahoma, Texas; Mexico. REFERENCES Ashmead, W. H. 1896. Descriptions of new parasitic Hymenoptera (Paper No. 2). Trans. Amer. Ent. Soc. 23: 179-234. Cameron, P. 1887. Family Braconidae. In Biologia Centrali-Americana. In- secta. Hymenoptera 1: 312-419. Muesebeck, C. F. W. 1950. Two new genera and three new species of Braconi- dae. Proc. Ent. Soc. Wash. 52: 77-81. and L. M. Walkley. 1951. Family Braconidae. In Muesebeck et al., Hymenoptera of America North of Mexico. Synoptic Catalog. U.S. Dept. Agr., Agr. Monogr. 2: 90-184. Tobias, V. I. 1961. A new genus from the tribe Doryctini and its taxonomic importance. Zool. Zh. 40: 529-535. (In Russian. ) Vargas, L. 1943. Los subgeneros americanos de Anopheles. Rev. Inst. Salub. Enferm. Trop. Mex. 4: 57-77. Viereck, H. L. 1914. Type species of the genera of Ichueumon flies. Bull. U.S. Nat. Mus. 83: 1-186. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 365 OBSERVATIONS ON ORIBATEI. THE DISCOVERY OF HAPLOCHTHONIUS SIMPLEX (WILLMANN) IN NORTH AMERICA (Acari: ACARIFORMES )* Haplochthonius simplex (Willmann, 1930) is a small, presumed primitive, oribatid mite of the enarthronote family Haplochthoniidae. The species is of interest to students of comparative morphology of Acariformes because of (1) a presumed primitive notogastral chaetotaxy; (2) the absence of notogastral scissures in the immature stases; (3) the presence of 5 pairs of centrodorsal notogastral lyrifissures; (4) the hypertrophy of the postlabral apodeme; and (5) the presence of 2 pairs of genital gland ducts emptying into the genital vestibule. Haplochthonius simplex was originally described from a single specimen found in leaf litter in Holland. The species was redescribed in magnificent detail by Grandjean in 1947 (Ann. Sc. Nat. Zool., Paris, lle Ser., VIII:213-248) and it is this redescription on which my determination of the specimens mentioned below is based. Grandjean collected H. simplex in several countries in the Mediterranean region from extremely dry litter and, at Periguex, inside his house. He charac- terized H. simplex as a xerophilous species and suggested that its occasional appearance, in small numbers, in Berlese extracts of mosses and moist litter materials may be due to contamination of the sample after it was brought into the laboratory. Willmann (1952; Zeitschr. f. Parasitenk. 15:392-428) recorded a single specimen of H. simplex from the pelage of a vole, Microtus oeconomicus, in Germany. The mite has apparently never been found outside of the Palaearctic region. In a Berlese extract of some dry straw and debris from the roost of an English sparrow (Passer domesticus) one female and 2 tritonymphs of Haplochthonius simplex were found. The roost material was taken from my garage, 3 miles south of Wooster, Wayne Co., Ohio on 5 November 1964. A second collection made on 9 November from the same roost netted one protonymph, 2 deutonymphs, and one tritonymph. The number of mites taken is not impressive but does definitely establish the presence of this interesting mite in the new world. It seems doubtful that the mites can be regarded as laboratory contaminants of the samples as we have never before encountered H. simplex in our laboratory or in samples extracted there. Other mite inhabitants of the roost were Aeroglyphus robustus (Banks), Dermanyssus hirundinis (Hermann), Dermatophagoides sp. immatures (the latter two associates of the English sparrow) and, in small numbers, miscellaneous Cheyletidae, Tarsonemidae, and erythracarine Anystidae. In passing it may be mentioned that van der Hammen (1952; Zool. Verhandel., No. 17:22) reported that no specimen was present on the type slide of H. simplex in the Oudemans collection at the Rijksmuseum van Natuurlijke Historie, Leiden. During a brief visit to the Oudemans collection in September, 1962, I examined this type slide and can report that the specimen is indeed present in the prepara- tion—Dona.p E. Jounston, Institute of Acarology, The Ohio State University, Columbus, Ohio 43210. 1Tmmediate publication secured by full payment of page charges—Editor. 366 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 AN ALATE APHID WITH AN EXTRA CORNICLE (HomorrTerA: APHIDIDAE)+* A single specimen of an alate aphid bearing an extra cornicle was found among approximately 300,000 aphids that were examined during 1960-66 in connection with a research project on aphid migration (NC-67). Entomologists in the central United States who cooperated in the project collected aphids in yellow-pan water traps, wind traps, and suction traps, and submitted the material to an identifica- tion center at the Department of Entomology, University of Wisconsin. The abnormal specimen was a species of Macrosiphum taken in a suction trap at Lincoln, Nebraska, October 3, 1966. The extra cornicle arose from the right side of abdominal segment VI (fig. 1). The specimen had the tips of the cornicles and antennae broken off. As these bear diagnostic characters, specific identification was not possible. Only once previously has this interesting and rare occurrence been reported in North America. Leonard described an extra pair of cornicles on apterae of Aphis sambucifoliae Fitch from Long Island, New York (Proc. Ent. Soc. Wash. 68: 320, 1966). Fig. 1. Dorsal view of Macrosiphum sp., showing location of an extra cornicle on segment VI of the abdomen. In Europe, extra cornicles have been reported on apterae of Aphis sp. near esulae Borner and Megoura viciae Kalt. (Remaudiére, G., Rev. Path. Veg. et Entom. Agric. 43: 31-35, 1964). This appears to be the first record of an extra cornicle either on an alate form or in the genus Macrosiphum.—J. T. MEpLER AND A. K. Guosn, University of Wisconsin, Madison. 14 contribution from the Wisconsin Agricultural Experiment Station, as a collaborator under North Central Regional Project NC-67, Migration of Aphids and Noctuids. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 367 THE STATUS OF RATITIPHAGUS BATOCINA EICHLER (MALLOPHAGA: MENOPONIDAE ) Eichler (1949, Boll. Soc. Ent. Ital. 79: 11-13) erected the new genus Ratiti- phagus and included in it the single new species, R. batocina. The description was based only on the female, presumably taken from a specimen of Pterocnemia pennata pennata (d’Orbigny) (Order Rheiformes). Hopkins and Clay (1952, A Check List of the Genera & Species of Mallophaga, London, 362 pp.), in reference to this, stated: “There is no generic description and the description of the type-species is so grossly inadequate that we are unable to form a useful opinion as to whether the genus is good or not, especially as it is extremely probable that the occurrence . . . on the alleged host was due to contamination.” They suggest, without availability of material for study, that Ratitiphagus is probably a synonym of Colpocephalum. I have recently been able to borrow 2 paratype 2 2 (WEC slides 1179q and 1179r) of R. batocina from the Zoologisches Staatsinstitut und Zoologisches Museum, Hamburg; these specimens are from the type-host with collection data: Punta Arenas, Magalhaenstrasse [Chile], 23 April 1912. While they are not of optimal quality, they apparently possess no features significantly differing from those of 2 2 of Colpocephalum trispinum Piaget as described by Price and Beer (1965, Ann. Ent. Soc. Amer. 58: 111-131). I see no reason, on the basis of available material, for not considering R. batocina a junior synonym of C. trispinum, whose type-host is a member of the Neotropical Ciconiiformes. This undoubtedly is an instance of contamination of the rheiform host, thus leading to the incorrect assumption by Eichler that he was dealing with a uniquely new louse from P. pennata.—Rocer D. Price, Department of Entomology, Fisheries, and Wildlife, University of Minnesota, St. Paul, Minnesota 55101. PTILINUS FRIENDI, A NOMEN NUDUM (COLEOPTERA: ANOBIIDAE ) The name Ptilinus friendi has appeared in print in Dissertation Abstracts in the abstract of John B. Simeone’s thesis for the Ph.D. degree (1961, volume 21, number 7) and in the Canadian Entomologist (1961, volume 93, number 6, p. 428). In the first reference above the name is given as having been proposed in the thesis for a species described as new; in the second reference the name is cited as a synonym of P. pectinicornis (L.). Neither of these literature citations serve to validate the name. Ptilinus friendi has thus never been validly proposed and is not to be accepted into our nomenclature.—RicHARD E. Wuirer, Entomology Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 368 PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 THE BLOW FLIES OF DOMINICA?* (DiereRA: CALLIPHORIDAE ) The blow flies of the Bredin-Archbold-Smithsonian survey of the island of Dominica consisted of only two species, both common and of wide distribution in tropical America. Though some variation was noticeable within the series, it was no more than could be noted in the same species in many areas on the American mainland. Cochliomyia macellaria (Fabricius), the secondary screw-worm, was represented by 45 specimens from the following localities: Clarke Hall, 12 Oct. to 17 Nov., 1964, P. J. Spangler, 12 April, 1964, O. J. Flint, Jr., 8-13 Jan., 1965, malaise trap, W. W. Wirth, and 12 May and 6 June, 1966, human feces, G. Steyskal; Cabrit Swamp, 3 Nov., 1964, Spangler, and 23 Feb., 1965, Wirth; Fond Colet, 5-9 Oct., 1964, Spangler; S. Chiltern, 8-10 Nov., 1964, Spangler; Grand Bay, 13 April, 1964, Flint; and near Layou, 22 Jan., 1964, H. Robinson. The primary screw-worm, C. hominivorax (Coquerel), was not represented in the material; this species, however, is very rare in collections obtained other than through rearing. Phaenicia eximia (Wiedemann), the common greenbottle fly of tropical America, was represented by 16 females and 2 males, the latter not in good condition. The parafacials vary in color, as they do in some series from the mainland, some being not as pale as the descriptions of this species would indicate. Localities represented were: Clarke Hall, 12 Oct. to 22 Nov., 1964, Spangler, 12 April, 1964, Flint, and 27 April, 1966, R. J. Gagné; Pont Casse, 12 & 23 Nov., 1964, Spangler; Cabrit Swamp, 3 Nov., 1964, Spangler; and Trafalgar Falls, 1200 ft., 5-6 April, 1966, Gagné—Mavurice T. JAMEs, Department of Entomology, Washington State University, Pullman, Washington 99163. 1 Scientific Paper 3031, College of Agriculture, Washington State University. Work conducted under Project 9043. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 369 SOCIETY MEETINGS 743rd Regular Meeting—May 5, 1966 The 743rd meeting of the Society was called to order by the President, Miss Louise M. Russell, on May 5, 1966, at 8:00 p.m. in Room 43, U.S. National Museum. Twenty-eight members and sixteen guests were in attendance. Minutes of the previous meeting were approved as read. Four new members were received into the society: Jeffrey N. L. Stibick, Carl J. Mitchell, Botha DeMeillon, and John D. Lattin. There was one candidate for membership, Dale L. Jackson. Mr. Lou Davis reminded the Society of the annual dinner meeting held jointly with the Insecticide Society of Washington at the University of Maryland. Helen Sollers-Riedel announced that she had tickets for the dinner available for anyone interested. M. D. Leonard presented an interesting resume of the various speculations and investigations concerning the biblical food “manna.” T. L. Bissell exhibited leaves from American Holly badly injured by a dipterous leaf miner. He also exhibited vol. 1, no. 1 of the recently established Journal of the Georgia Entomological Society. Vic Adler announced the establishment of the Bussart Award sponsored by Velsicol Chemical Corporation in honor of the late J. Everett Bussart, distinguished entomologist and past president of the Entomological Society of America. A recently collected Periodical Cicada was exhibited by David Gilsey. Mr. Robert Kemp exhibited his entry in the Science Fair competition entitled “Ant reaction to light intensity.” Following his demonstration, President Russell presented him with a book on insects as a small token of the Society’s appreciation. The speaker for the evening, Dr. Reinhold A. Rasmussen, Walter Reed Army Medical Center, gave an interesting illustrated talk on the relationships between bees, scent and orchid pollination. Following the introduction of visitors, the meeting was adjourned at 9:35 p.m. —W. DonaLp Duckworth, Recording Secretary. 745th Regular Meeting—October 6, 1966 The 745th regular meeting of the Society was called to order by the president, Miss Louise M. Russell, on October 6, 1966, at 8:00 p.m. in room 43, U.S. National Museum. Thirty-seven members and twenty-five guests were in attend- ance. Minutes of the previous meeting were approved as read. One new member was received into the Society—Dale L. Jackson. There were eight candidates for membership—J. C. Lien, Thomas G. Floore, Joseph T. Koski, Clyde F. Smith, Linda Butler, D. F. Martin, D. G. Denning, and Robert W. Poole. President Russell announced the appointments to the following committees: Auditing Committee—J. E. Gilmore, J. A. Fluno, chairman, R. I. Sailer; Nominat- ing Committee—P. A. Woke, chairman, B. A. App, J. M. Kingsolver. T. J. Spilman spoke on the life of Frank A. McDermott, a specialist on the Lampyridae, who died on July 3, 1966. In addition, the second edition of the Lampyridae part of the Coleopterorum Catalogus Supplementa, written by Mr. McDermott, was reviewed. Paul A. Woke noted the Anniversary meetings of the School of Public and Environmental Health. 370 PROC, ENT. SOC, WASH., VOL. 69, NO. 4, DECEMBER, 1967 Lou Davis presented greetings from Miriam A. “Ma” Palmer. The program for the evening was an interesting illustrated symposium entitled “A Review of Egypt and its Entomology.” Participants included R. H. Foote, moderator, R. W. Hodges, P. M. Marsh, G. C. Steyskal, and E. W. Baker. After the Introduction of visitors, the meeting was adjourned at 10:00 p.m.— Paut M. Marsu, Acting Recording Secretary. 746th Regular Meeting—November 3, 1966 The 746th regular meeting of the Society was called to order by the president, Miss Louise M. Russell, on November 3, 1966, at 8:00 p.m. in room 43, U.S. National Museum. Twenty-one members and ten guests were in attendance. Minutes of the previous meeting were approved as corrected. Eight new members were received into the Society—J. C. Lien, T. G. Floore, J. T. Koski, C. F. Smith, L. Butler, D. F. Martin, D. G. Denning, and R. W. Poole. There were two candidates for membership—Robert F. Whitcomb and Dan M. Harmon. Paul A. Woke, chairman of the nominating committee, presented the following proposed slate of officers for 1967: President, Louis G. Davis; President-elect, Richard H. Foote; Recording secretary, Ralph A. Bram; Corresponding secretary, David R. Smith; Treasurer, Arthur K. Burditt, Jr.; Editor, Jon L. Herring; Custodian, Robert Smiley; Program Committee Chairman, Victor E. Adler; Membership Committee Chairman, William B. Hull. President Russell announced that the December meeting will be held on December 8th. Mr. Bissell noted that Thyridopterix ephemeroformis was causing damage to junipers at the cornerstone of the Mason Dixon marker in the southwest corner of Delaware. Dr. Hodges noted that volume one of Microlepidoptera Palaearctica was recently issued. It concerns the Crambinae and was written by S. Bleszynski. The program for the evening was an interesting talk entitled Insects Versus Computers, presented by Dr. Judson U. McGuire, Biometrical Services Staff, USDA. A lively period of discussion followed Dr. McGuire's talk. After the introduction of visitors, the meeting was adjourned at 9:30 p.m.— Pau M. Marsu, Acting Recording Secretary. 748th Regular Meeting—January 5, 1967 The 748th regular meeting of the Society was called to order by the president, Mr. Louis G. Davis, on January 5, 1967, at 8:00 p.m. in room 43, U.S. National Museum. Thirty-two members and twelve guests were in attendance. Minutes of the previous meeting were approved as read. Mr. C. John Lane was received into the Society. President Davis announced committee appointments for 1967 as follows: Membership Committee—William B. Hull (Elected Chairman), H. Ivan Rainwater, Ronald W. Hodges, D. H. Messersmith, A. S. Michel; Program Com- mittee—Victor E. Adler (Elected Chairman); Publications Committee—Jon L. Herring (Elected Chairman), James P. Kramer, Paul M. Marsh, W. Donald Duckworth; Advertising Committee—James R. Foster, John M. Kingsolver; Finance Committee (established by the bylaws)—Arthur K. Burditt, Treasurer PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 371 (Chairman), Jon L. Herring, Editor, Robert Smiley, Custodian, Victor E. Adler, Program Chairman; Representative to the Washington Academy of Sciences— Harold H. Shepard. President Davis announced the deaths of the following members during 1966: Dr. H. Ruckes, Dr. R. L. Webster, and Dr. G. Wolcott. Dr. Gurney added the name of Miss S. I. Parfin. Mr. T. L. Bissell noted that in 1966 Dr. E. F. Knipling, a member of the Society, had received the Rockefeller Public Service Award and the National Medal of Science for his outstanding contributions to science. Dr. B. D. Burks commented on investigations being carried out in Bolivia under British auspices on Triatoma vectors of Chagas disease. Dr. A. B. Gurney exhibited the striking horned male of Dynastes hercules (L. ) and a red-spotted evaniid, Hyptia johnsoni Ashmead, which were collected in the West Indies. Dr. W. H. Anderson reported briefly on the successful introduction of a flea beetle, Agasicles sp., from Argentina to control alligator weed in the southeastern United States. Miss Louise M. Russell presented the retiring president’s address, a most interesting illustrated review of the present state of Aphidology. This was followed by a stimulating illustrated discussion of the tropical horse tick as a vector of equine piroplasmosis by Dr. D. W. Anthony. Following the introduction of visitors, the meeting was adjourned by President Davis at 9:45 p.m.—Raupu A. Bram, Recording Secretary. 749th Regular Meeting—February 2, 1967 In the absence of the President, the 749th regular meeting of the Society was called to order by the President-elect, Dr. Richard H. Foote, on February 2, 1967 at 8:00 p.m. in room 43, U.S. National Museum. Forty-six members and twenty-two guests were in attendance. Minutes of the previous meeting were approved as read. The names of ENS J. Phillip Keathley, MSC, USNR and Mr. Gerald G. Elgert were read for the first time as candidates for membership in the Society. Dr. Foote regretfully announced the death on February 1, 1967 of Mrs. T. E. Snyder, wife of our Honorary President. C. W. Sabrosky exhibited an insect box which he had received from northern Nigeria and which was infested with the hyde beetle, Dermestes maculatus deGeer. G. G. Elgert exhibited a scarab beetle, Polyphyla variolosa Hertz, which he col- lected in a light trap on Assateague Island, Maryland as a new state record. V. E. Adler presented several current newspaper and magazine articles of entomological interest. A. B. Gumey exhibited a pair of Dryococelus australis (Montrouzier ), a very large and unusual walkingstick from Lord Howe Island, east of Australia. It had been thought to be extinct, due to depredations of introduced rats; how- ever, a recent note by McAlpine (Aust. Ent. Soc. News Bull. 2: 71-72, 1966) records its presence on Ball’s Pyramid, a rocky island about 14 miles from Lord Howe. G. C. Steyskal reviewed and exhibited the book, “The Amazing World of Insects” by A. T. Bandsma and R. T. Brandt, published by the Macmillan Company in 1963. F. Smith exhibited a new book, “Insect Colonization and Mass Production,” edited by C. N. Smith and published by the Academic Press. The first speaker for the evening, Dr. W. Donald Duckworth, presented a most interesting illustrated discussion of “Entomology in the Tropicc-—New Emphasis 372 PROC, ENT, SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 on an Old Subject.” Dr. Duckworth’s remarks concentrated on his experiences and knowledge of entomological programs in the New World tropics. The next speaker, Dr. Reece I. Sailer, reviewed the past and present research programs of the U.S.D.A. European Parasite Laboratory located outside of Paris, France. Dr. Sailer’s informative, illustrated discussion included vignettes drawn from his five years experience as director of the laboratory. Following the introduction of visitors, the meeting was adjourned by President- elect Foote at 10:10 p.m.—RAtpu A. Bram, Recording Secretary. PROC. ENT. SOC. WASH., VOL. 69, NO. 4, DECEMBER, 1967 373 INDEX TO VOLUME 69 Abagrotis alampeta, 97 Amblyceran bird lice, synonymy, 248 Amoebaleria defessa and sackeni, dif- ferences, 296 Anicla cemolia, 101 Anopheles ( Anopheles ) tigertti, 19 Anthomyia procellaris, in North Amer- ica, 240 Apenesia chilena, 271 Aphid, extra cornicle, 366 Aphids, of Cyprus, 259 Ardeacarus americanus, 77 Aspidepyris austrinus, 270 Ataleute carolina, 181 Atopsyche socialis, 165 Atteva zebra, 71 Aulonium, key to U. S. species, 150 Berosus (E.) latipalpus, 64; spectatus, female, 65 Bethylidae, new generic records, 269 Bidigitus, 280; platani, 281 Biza ava, 45 Blow flies, of Dominica, 368 Boydaia turdi, 294 Cerceris simplex graphica, bionomics, 312 Ceropales maculata, in Wisconsin, 92 Chamaemyiidae, key to genera, 275 Cheumatopsyche analis, in Hawaii, 56 Chimarra bicoloroides, 166 Chinaia caprella, 46; permista, 46 Chinchinota, 41; styx, 41 Chirothrips, new locality records, 47; kurdistanus, 49 Chrysotimus arizonicus, 125; obscurus, 126; schildi, 126 Chrysotus clypeatus, 121; mexicanus, 120; neopicticornis, 119 Cimex hemipterus, from bats, 179 Coelidiana, 31; key, 31; ancora, 35; colens, 36; durator, 35; patrator, 33 Coelindroma, 43; fungoma, 44; fungosa, 43 Colaspis barberi, 232; cubensis, 226; darlingtoni, 234; farri, 235; luciae, 236; orientalis, 227 Coleoptera, new records from North Carolina, 81 Conservana, 266; angustata, 268; con- versa, 266; reversa, 267 Copidosoma capsicum, 54 Corythaica darwiniana, 89; wolfiana, 87 Culex, lectotype assignments, southeast Asia spp., 327 Culex (Lophoceraomyia) mammilifer group, 1; eukrines, 11; fuscosiphonis, 14; inconcomptus, 16; peytoni, 7; pholeter, 13; speculosus, 3 Cylindromyia, in North America, 60; dosiades lobata, 62 Dactynotus, chromosomes, 303 Dendrosoter, notes, 246; chansleri, 246 Diaphorus bakeri, 117 Dicosmoecus frontalis, 55; aureoventris, Bo Dolichomotes, 334; navei, 337 Dolichopodidae, replacement 224 Dolichopus crassitibia, 114 Drabeha, 237 Drosophila leticiae, 111 Canadian names, Ectoparasites, birds and mammals, 349 Fenusa, key, 282 Fruit flies, new host records, 146 Gabucinia alcyon, 75 Haemagogus aeritinctus, 103; lucifer, 103 Haplochthonius simplex, in N. Amer., 365 Hecabolini, status, 359 Helina veniseta, 242; violescens, 241 Hemitarsonemus, key to males, 134; deleoni, 133; leonardi, 133 Heterarthrinae, host list, 277; key to Nearctic genera, 278 Hoplomyophilus, 287; nativus, 289 Ichneumonidae, notes on Motschoulsky types, 93 Indioleucopis, 275 Insects, protection of, in liquid, 286 Kentrotryphon, notes, 58 Ktenostilpnus, notes, 58 374 Lace bugs, of the Galapagos, 82 Lamprochromus occidentalis, 123 Leluthia, key to spp., 360; floridensis, 362 Lepidostoma lacinatum, gulare, 176 Leucopis luteicornis, new subgenus for, 275 Limnephilus ctenifer, 170; mexicanus, 171; pollux, 169 Listrophorus leuckarti, 340 Lygaeidae, synonymy, 244 Lystridea, key, 292; wilkeyi, 292 Macrosiphon rosae, on holly, 59 Macrosiphoniella, list of North Amer- ican species, 184 Marilia species, 173; (A), 173; (B), 173 Medetera marylandica, 124 Melisandra, new to North America, 95; morio, 95 Metallus, key, 280 Micromorphus, key to American spp., 329; asymmetricus, 332; knowltoni, 331; Jeucostoma, 330; 333 Mite deutonymphs, as endoparasites of birds, 73 Mosquito catalog, supplement III, 197; index, 223 Nelidina, key, 40; sportula, 41 Neocoelidia onca, 45 Neoparentia, 252; key, 253; bisetosa, 253; deformis, 256; obscura, 257; schildi, 258; tarsalis, 257 Neotarsonemus, 137; type-species, beeri, 137 Nicuesa, replacement name for, 237 175; rectan- lithophilus, Oecanthus, male metanotal glands, 157; key to U.S. species based on glands, 160 Omania, key to spp., 354; nauruensis, 356 Orgilus lepidus, 177 Paduniella nearctica, 310 Paduniellini, first new world record, 310 Palaeacarus, in eastern U. S., 301 PROC. ENT. SOC. WASH., VOL. 69, No. 4, DECEMBER, 1967 Palaeosepsis, Melander and_ Spuler species, 180 Parascleroderma pucallpa, 271 Parentypoma, notes, 58 Parfin, S. I., obituary, 190 Pectinibruchus, 324; longiscutus, 324 Phatnoma eremaea, 83 Phthorimaea operculella, 52; 52 Platypus, sex ratios, 297 Pleroneura, new synonymy, 182 Polycentropus palmitus, 167 Pristiphora abbreviata, extended dis- tribution, 92 Prolatus, 282; artus, 283 Psocoptera, from bird plumage, 307 Ptilinus friendi, nomen nudum, 367 terrella, Radfordia subuliger, lectotype designa- tion, 183 Ratitiphagus batocina, status, 367 Rhaphium steyskali, 116 Rhipibruchus, 318; prosopis, 322 Salda coloradensis, 27; alta, 29 Saldidae, North American, 24 Saldula usingeri, 346 Steneotarsonemus spinki, 129 Strongylogaster macula, new to North America, 95 friedmani, 131; Tarsonemella beameri, 135 Tarsonemus bucheleri, 137; hunti, 139; moseri, 141; newkirki, 145; steg- maieri, 143; stricketti, 141 Telmaturgus costaricensis, 122 Thecodiplosis, 338 Thyce osburni, 238 Tinconaria, 155; albescentis, 155 Tozzita, 39; petulans, 39 Trachymyrmex bugnioni synonymy, 273; diversus synonymy, 274 Tritoxa, key to species, 94 Trombidiid mites, parasitic on Dasy- helea mutabilis, 284 Xenocoelidia, key, 37; charisma, 38; litura, 38 Xiqilliba, 39, 46; bellator, 46; coelid- oides, 46 Xylocelia franclemonti, linear copula- tion, 343 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. Manuscripts for publication, proof and other editorial matters should be addressed to the Editor (for address, see inside front cover of this issue). Typing—All manuscripts must be typed on bond paper with double-spacing and ample margins. Carbon copies or copies on paper larger than 84% x 11 inches are not acceptable. Do not use all capitals for any purpose. Underscore only where italics are intended in the body of the text, not in headings. Number all pages consecutively. 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Ent. Soc. Wash. 65(4): 306. and L. M. Walkley. 1951. in Muesebeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2: 90-184. In shorter articles, references to literature should be included in parentheses in the text. Tables—Tables are expensive and should be kept to an absolute minimum. In most cases, material in tables can be incorporated into the text. When tables are necessary, each table should be prepared as a line drawing or typed on a separate page with heading at top and footnotes below. Number footnotes consecutively for each table. Use only horizontal rules. Illustrations—No extra charge is made for line drawings or halftones. Authors must plan their illustrations for reduction to the dimensions of the printed page and the individual figures must be mounted on suitable board. Proportions of full-page illustrations should closely approximate 4% x 6” (26 X< 36 picas); this usually allows explanatory matter to appear on the same page. On the back of each illustration should be stated (1) the title of the paper, (2) the author’s complete name and address, and (3) the number of the illustration such as “No. 1 (of 3)” etc. Figure legends—Legends should be typewritten double-spaced on separate pages headed ExpLANATION OF FicuREs and placed following REFERENCEs. Do not attach legends to illustrations. Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) McDANIEL, B., J. P. SHOEMAKER and S. J. JOY—The discovery of Listrophorus leuckarti Pagenstecher on Microtus pennsylvanicus from North America .( Acafina: Listrophoridae) se 340 MEDLER, J. T. and A. K. GHOSH—An alate Sern with an extra cornicle (Homoptera: Amhigidae ye qo 2 i ole MOCKFORD, E. L.—Some Psocoptera from plumage of birds —..-- 307 OLIVE, A. T.—Chromosomes of the aphid genus Dactynotus (Homoptera) — 303 PRICE, R. D.—The status of Ratitiphagus batocina Eichler (Mallophaga: Menoponidae) LORRI IRe 367 POLHEMUS, J. T.—A new saldid from California (Hemiptera: Saldidae) _.. 346 ROBINSON, H.—New species of Micromorphus from the United States and Mexico, (Diptera: Dolichopodidae) —~ ——.._..__..__.._ ee 329 SMILEY, R. L.—Dolichomotes navei, a new genus and new species of pyemotid mite (Acarina: Pyemotidae) _....._..._ 334 WHITE, R. E.—Ptilinus friendi, a nomen nudum (Coleoptera: Anobiidae) —. 367 WILSON, N.—Ectoparasites of Canadian birds and mammals 349 SOGIETY MEETINGS 2.0 eee eee 369 "7 INDEX, VOLUME 69 0.5.00... ee 373 EANTHSO Nigye FEB ~ 9 1968 J ipnacits PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF W ASHINGTON Volume 70 OFFICERS FOR THE YEAR 1968 wt _ ——— CO 2D ad ara EON rc SO a es a Richard H. Foote (ED SVD) 2 2 Ayes ll ele HE UE ak lS ol le Helen Sollers-Riedel Recording Secretary OR SARC. 5 eR RE ye Peet 2 Ralph A. Bram MERESTONCIN MS CCTCLOMY) ne David R. Smith OBERT ITO essen ete EE eee Arthur K. Burditt, Jr. SE cers 2 Tae e Sne RE Ss T SI SAA it tie Se a Paul M. Marsh (ESPs, eee ES specail Uda id apd ied le le tibiae Robert L. Smiley Program Committee Chairman Richard G. Oakley Membership Committee Chairman __.._.-_.-____»_»_ William B. Hull Delegate to the Washington Academy of Spence. eI et nakat oe eA, W. Doyle Reed PUBLISHED BY THE SOCIETY WASHINGTON, D.C. 1968 TABLE OF CONTENTS, VOLUME 70 BALDUF, W. V.—On the life of Vespula vulgaris (L.) and V. maculifrons (Buysson) “(Hymenoptera: Vespidae) —...... eee BICK, G. H. and J. C. BICK—Demography of the damselfly, Argia plana Calvert (Odonata: Coenagriidae) _.._........ BICK, J. C.: See BICK, G. H. BLAKE, D. H.—Ten new chrysomelid beetles from Dominica and Jamaica __ BLANTON, F. S.: See WIRTH, M. O. BOHART, R. M.—New Hoplisoides from the United States (Hymenoptera: Sphecidae yas tert) eee BRAM, R. A.—A ieileguaetim of Culex (Acalleomyia) obscurus (Leicester ) (Diptera Gulicidae)) = sess ee ee BRENDLE, J. J.: See BROOKS, R. B., JR. BROOKS, R. B., JR. and J. J. BRENDLE—Colonial behavior of Apis mel- lifera mellifera L. after exposure to gamma radiation (Hymenoptera: AIG ag ie fee 2 ee eee BURGESS, L. P., JR. and C. W. WINGO—tThe host and present status of Alysia Pathunda Say in Missouri (Hymenoptera: Braconidae) — BURKS, B. D.—New North American species of Ceratosmicra Ashmead (Hymenoptera: ‘Chaleididac) 2.2 T—E—eeeeeee CAPRILES, J. M.—A new genus and species of Deltocephalinae from Puerto Rico (Homoptera: ‘Cicadellidae) 2... Eee CHRISTENSON, D. M.: See GJULLIN, C. M. COOK, D. R.—New species of Neoacarus Halbert and Volsellacarus Cook from North America (Acarina: Neoacaridae) Water mites of the genus Stygomomonia in North America (Acarina: Momoniidae)). 2 3 ee Eee COONEY, J. C. and K. L. HAYES—A new locality record for Ixodes woodi Bishopp, (Acarina: -Ixodidne) sof... ee eee CRABILL, R. E., JR.—On the true identity of Chomatophilus with descrip- tion of a new species, and with key and catalogue of all sogonid genera (‘Chilopoda:Géophilomorpha: Sogonidae)) _...__. eee On the true identities of Tuoba and Nesogeophilus (Chilopoda: Geophilomorpha: Geophilidae) 2.0... a eee CUNLIFFE, F.—A new species of Pimeliaphilus from California (Acarina: Ptery@osomidae) .. <.2..8 ee ee DELFINADO, M. D. and E. R. HODGES—Three new species of the genus Tripteroides, subgenus Tripteroides Giles (Diptera: Culicidae) — EMERSON, K. C.—The host of Stachiella retusa martis Werneck (Mal- lophaga: 'Trichodectidae) ...5 nee and R. D. PRICE—A new species of Rhynonirmus from Thailand (Mallophaga: Philopteridae) 20. Eee FISK, F. W. and A. B. GURNEY—Neotropical cockroaches of the genus Xestoblatta: a new species from Costa Rica and notes on other species (Dietyoptera: Blattaria: Blattellidae) —...... Eee FOX, J. W.—Strepsiptera from the Smithsonian Institution’s survey of Dominica, with description of a new species of Elenchidae — FROESCHNER, R. C.—Telamona archboldi, a new treehopper from Florida ii 287 303 204 170 35 67 210 383 323 184 137 238 (Homoptera: Membracidae) — 154 Burrower bugs from the Galspacee rea: feollected ne hue 1964 expedition of the Galapagos Scientific Project (Hemiptera: Cydnidae) _ 192 Notes on the systematics and morphology of the lacebug subfamily Wanatacadenmnacy (demipterac, (Ringidae)) 4.2 ee tk 245 GAGNE, R. J.—Chrybaneura harrisoni, a new genus and species from Central America (Diptera: Cecidomyiidae) — 33 Authorship of Pachydiplosis oryzae, a rice See ae Reta nl Osiios Diptera. Cece Oni yild aes py 4 teas eee Be ie Brees 6 te 136 GENTRY, J. W. and E. J. GERBERG—Dipterous eggs as food for adult Coccinellidae (Coleoptera: Coccinellidae) per ae eee ee 142 GERBERG, E. J.: See GENTRY, J. W. GJULLIN, C. M., L. F. LEWIS, and D. M. CHRISTENSON—Notes on the taxonomic characters and distribution of Aedes aloponotum Dyar and Aedes communis (De Geer) (Diptera: Culicidae) Abie Batre 133 GLOVER, S.: See HUNTER, P. E. GORSKI, L. J.—A new species of Lygaeus from Peru (Hemiptera: Lygae- CIEE) Le er ra ee eee ee 74 GURNEY, A. B.: See FISK, F. W. HAYES, K. L.: See COONEY, J. C. HOCHMAN, R. H.: See PEYTON, E. L HODGES, E. R.: See DELFINADO, M. D. HUANG, Y.-M.—Neotype designation for Aedes (Stegomyia) albopictus ROkHSe) MuCMiptera Daas Byline” : with " ake wie a et wae oe 7) rey Pay ay t a4 , ¥ "White Tyan qe mr ink vf wy, Wet q is wae Vol. 70 MARCH 1968 No. 1 PROCEEDINGS of the ENTOMOLOGICAL SOCIETY of WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 PUBLISHED QUARTERLY CONTENTS BLAKE, D. H.—Ten new chrysomelid beetles from Dominica and Jamaica _. 60 BRAM, R. A.—A redescription of Culex (Acalleomyia) obscurus (Leicester ) OCS Tc Ly Ae a aS ee eee | aS Se 52 CAPRILES, J. M.—A new genus and species of Deltocephalinae from Puerto in Hawmamera= Gicndelinae). 0 Ss COOK, D. R.—New species of Neoacarus Halbert and Volsellacarus Cook from North America (Acarina: Neoacaridae) _... = CUNLIFFE, F.—A new species of Pimeliaphilus from California (Acarina: EEE DET TS ge EES SPOS eed Sa oe eae SE CN OE coke n= Seer Se 1 GAGNE, R. J.—Chrybaneura harrisoni, a new genus and species from Central Pomenica | Dinteras (Cecidoutynaae) 2 bi GORSKI, L. J.—A new species of Lygaeus from Peru (Hemiptera: Lygaei- HUNTER, P. E. and S. GLOVER—The genus Passalobia Lombardini, 1926, with description of a new species (Acarina: Diarthrophallidae) 38 KINGSOLVER, J. M.—A review of the obtectus group in Acanthoscelides Schlisky, with designations of lectotypes (Coleoptera: Bruchidae: Bru- Nn estes cere hol, Se ede eee ee Log eh KNUTSON, L. V. and J. ZUSKA—A new species of Pteromicra and of Euthycera from western North America (Diptera: Sciomyzidae) — —— (Continued on back cover) ENTOMOLOGICAL SOCIETY OF WASHINGTON ORGANIZED MarcuH 12, 1884 OFFICERS FOR 1968 Dr. Ricuarp H. Foore, President Dr. Pau, M. Marsn, Editor Ent. Res. Div., ARS, USDA Systematic Entomology Laboratory, USDA Plant Industry Station c/o U.S. National Museum ' Beltsville, Maryland 20705 Washington, D. C. 20560 Mrs. HELLEN SOLLER-RIEDEL, President-Elect Mr. RoBert L. SMiLEy, Custodian ; Agr. Res. Ser., USDA Ent. Res. Div., ARS, USDA Pe Room 466, Federal Center Bldg. Room 3139, South Bldg. 7 Hyattsville, Maryland 20782 Washington, D. C. 20250 ¥ Dr. RAaupu A. Bram, Recording Secretary Mr. RicHarp G. OAKkLEy, Program Committee Ent. Res. Div., ARS, USDA Chairman 5 Plant Industry Station 1914 Franklin Ave. " Beltsville, Maryland 20705 McLean, Virginia 22101 Ms Dr. Davin R. Smiru, Corresponding Secretary Carr. WILLIAM B. Hutt, Membership Committee Systematic Entomology Laboratory, USDA Chairman : c/o U.S. National Museum 746 Beall Ave. Washington, D. C. 20560 Rockville, Maryland 20850 Dr. ArtHuR K. Burpirrt, Jr., Treasurer Ent. Res. Div., ARS, USDA Plant Industry Station Beltsville, Maryland 20705 Honorary President Dr. T. E. SnypER, U.S. National Museum ee ee ee Honorary Members 7 Mr. C. F. W. MursEeseck, U.S. National Museum f Dr. F. W. Poos uF ba J Dr. E. N. Cory ; MEETINGS.—Regular meetings of the Society are held in Room 43 of the U.S. National Museum on the fir Thursday of each month from October to June, inclusive, at 8 P.M. Minutes of meetings are published regula in the Proceedings. MEMBERSHIP.—Members shall be persons who have demonstrated interest in the science of entomology} Annual dues for members are $5.00; initiation fee is $1.00 (U.S. currency). PROCEEDINGS.—Published quarterly beginning with March by the Society at Washington, D.C. Members i good standing are entitled to the Proceedings tree of charge. Nonmember subscriptions are $8.00 per yeal both domestic and foreign (U.S. currency), payable in advance. All remittances should be made payable t The Entomological Society of Washington. The Society does not exchange its publications for those of other societies. STATEMENT OF OWNERSHIP Title of Publication: Proceedings of the Entomological Society of Washington. Frequency of Issue: Quarterly (March, June, September, December). Location of Office of Publication, Business Office of Publisher and Owner: The Entomological Society ¢ Washington, c/o Department of Entomology, U.S. National Museum, Washington, D.C. 20560. ‘ Editor: Dr. Paul M. Marsh, same address as above. Managing Editor and Known Bondholders or other Security Holders: none. This issue mailed April 17, 1968 Second Class Postage Paid at Lawrence, Kansas, U. S. A. 66044 ALLEN PRESS, INC. wee LAWRENCE, KANSAS 66044 U.S. B PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Vol. 70 MARCH 1968 No. 1 A NEW SPECIES OF PIMELIAPHILUS FROM CALIFORNIA’ (ACARINA: PTERYGOSOMIDAE ) FREDERICK CUNLIFFE, Walpole, New Hampshire Specimens of an undescribed species of Pimeliaphilus collected from Trogaderma parabile in California were sent to the U.S. Department of Agriculture in Washington in 1966 for determination. They are here described as new. For detailed studies on the genus see Newell and Ryckmann (1966). Pimeliaphulus trogadermus, new species, is distinctive in having long dorsal body setae entirely barbed throughout, in that the short ventral body setae are entirely barbed, in lacking striae on the female dorsal shield, and in the shape of the dorsal shields of the male. Pimeliaphilus trogadermus, n. sp. (Figs. 1-10) Female. Length of idiosoma 574 u; length of gnathosoma 160 «. Propodosomal shield 177 long, broad anteriorly and narrowing posteriorly, 97 « wide between Ist pair of setae, 81 « between 2nd pair, and 77 uw between 3rd pair. First pair of shield setae 52 uw long, 2nd pair 97 uw long, 3rd pair 145 uw long; other dorsal body setae 150 uw long except for humerals which are 81 y long; all dorsal body setae strongly barbed throughout entire length. Chelicerae 129 w long; femoral setae of palps barbed and 32 uw long; femur and patella of palps not fused; ventral setae of gnathosoma sparsely barbed on basal half and tapering to point and located on level with insertions of palpi. Peritremes short and not reaching to margins of gnathosoma. Genital and anal setae much shorter than dorsal setae; ventral and coxal setae similar to genital setae; no setae on coxae IV; all setae strongly barbed. Femora I and IV divided into basifemur and telofemur; leg setae long and entirely barbed; tarsus I with long slender solenidion; tarsus II with short, stout solenidion; tarsus III with short, slender solenidion; tibiae I, II, and IV each with a long, slender bothridion. Length of leg segments as follows—tarsus to femur in microns: I, 129-94-80-113; II, 113-55-55-81; III, 135-74-65-81; IV, 171-90-74-97. Setal lengths in relation to leg segments are as figured. Male (allotype). Length of idiosoma 383 y; length of gnathosoma 96 uy. Propodosomal shield 145 yw long, parallel sided and rounded posteriorly; 48 yu between Ist, 2nd, and 3rd pair of shield setae, and 52 « between 4th pair. First pair of shield setae 48 u« long, 2nd pair 81 yu long, 3rd pair 84 uw long, and 4th pair 103 » long; humeral and other dorsal body setae 100-103 u long; all dorsal body setae strongly barbed; anal setae short and smooth. Chelicerae 81 ,» long; peri- ‘Immediate publication secured by full payment of page charges—Editor. 2 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Win Ny aS ) pre U7, ¥ “ fy rf W\- i Figs. 1-4, Pimeliaphilus trogadermus, n. sp.: 1, dorsal view, 2; 2, gnathosoma, @; 3, dorsal view, ¢; 4, ventral body seta, ¢. tremes short and not reaching margins of gnathosoma; femora and patella of palps not fused; femoral setae short, about 18 « long, and strongly barbed. With two pairs of hysterosomal plates, the anterior pair small, the posterior pair large and reaching ventrally; all plates bearing two pairs of setae. Ventral body and coxal setae as in female. Femora I and IV not divided; tibia I, III, and IV each with a slender bothridion, and tarsus I, II, and III each with a solenidion. Lengths of leg segments as follows—tarsus to femur in microns: I, 106—65-—58-90; II, 87-42— 42-48; III, 106-48—42-45; IV, 119-55-45-65. Lengths of leg setae in relation to segments are as figured. Holotype. Female, ex Trogoderma parabile, University of Califor- nia, Riverside, California, March 25, 1966 by Gregory J. Partida. Allotype. Male with the above data. Paratypes. Two males with the above date. Eight nymphs were also in the series. The holotype and allotype are in the U.S. National Museum (no. 3249); a male paratype and a nymph are in the collection in the Department of Biology, University of California, Riverside; a male paratype and a nymph are in the collection of the Bureau of Ento- mology, State of California, Sacramento, California. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 5-10, Pimeliaphilus trogadermus, n. s il, 2; 8, leg IV, 9; 9, leg I, @: 10, leg II, P...0, leg 1 oO: 6. Jeg Il, 9. 7. les é. 4 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 REFERENCE Newell, I. M. and R. E. Ryeckman. 1966. Species of Pimeliaphilus (Acari: Pterygosomidae) attacking insects, with particular reference to the species parasitizing Triatomae (Hemiptera: Reduviidae). Hilgardia 37(12):403—435. A REVIEW OF THE OBTECTUS GROUP IN ACANTHOSCELIDES SCHLISKY, WITH DESIGNATIONS OF LECTOTYPES (COLEOPTERA: BRUCHIDAE: BRUCHINAE) Joun M. Kincso.ver, Systematic Entomology Laboratory, U.S. Department of Agriculture, Washington, D.C. 20560 Much confusion has centered around the correct scientific name to be used for the bean weevil (or bruchid) now accepted as Acantho- scelides obtectus (Say). Bridwell furnished a list of synonymy for this species in Larson and Fisher (1938), but many of the names included were undoubtedly placed there from descriptions only and must be checked by examination of type specimens. In 1942, Bridwell pub- lished a resumé of the nomenclatural history of the bean bruchid and described two closely related new species in the Neotropical Region, A. obreptus and A. obvelatus, which also infest stored beans, but he did not include illustrations of them. The specimens upon which the descriptions were based are deposited in the U.S. National Museum Collection, but types unfortunately were never selected by Bridwell. In the same paper, he also gave a footnote reference to the true identity of A. obsoletus (Say), the name of which has often been applied to the bean bruchid but which belongs to a different species group. As early as 1892, C. V. Riley recognized the distinctness of A. obtectus and A. obsoletus which had been confused by Horn in 1873 and again by LeConte and Horn in 1877. However, this error by LeConte and Horn was perpetuated by Pic (1913) in his catalogue of Bruchidae, which caused further confusion. Acanthoscelides obso- letus has been reared only from seed of the genus Tephrosia (= Cracca) and has not been known to infest any other genus of plants. Say (1831) apparently mistakenly identified the host of obsoletus as Astragalus, because he also reared from the same seeds the curculionid Apion segnipes Say which is restricted in host to Tephrosia virginiana (L.) Pers. This combination of insect species has since been reared from T. virginiana. The purpose of the present paper is to designate lectotypes of Acanthoscelides obreptus and A. obvelatus, and to illustrate salient characters for ease in identification of A. obtectus, A. obreptus, and PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 5 A. obvelatus. The male genitalia of A. obsoletus are also illustrated for contrast. The obtectus group is described in detail by Bridwell (1942, p. 253), but the principal group characters are illustrated here. Those to be noted especially are the arcuate lateral margins of the pronotal disk (fig. 13), the shape of the antennae (essentially identical in both sexes, fig. 20), the shape of the head and eyes (fig. 18), the obsolete frontal carina (fig. 18), the regularity of the elytral striae (fig. 13), the shape and dentition of the hind leg (fig. 15), the lack of modification of the basal abdominal sternite in the male, the expanded apices of the lateral lobes that form a hoodlike cover above the apex of the median lobe (figs. 2, 5, 9), the shape of the ventral valve of the median lobe, and the arrangement of the armature of the internal sac (figs. 1, 4, 7). An additional character not mentioned by Bridwell is the presence of a small denticle at the extreme base of the third, fourth, and fifth striae. This character may be useful in relating this group to others in Acanthoscelides and in other genera. The following key is taken as published by Bridwell (1942). Key TO THE OxstrEctus Group oF Acanthoscelides Scuitsky, 1905 1. Antenna, legs, and pygidium entirely reddish; ovipositor strongly scle- rotized, ending in two closely applied, spinelike processes, margin narrowing before the apex with a dentiform tubercle on each side areca Ec | SS, 2 ee ER 9 Pe 2 eee obreptus Bridwell Antenna dark gray in the middle; ovipositor neither strongly sclerotized BE: ROVERS THO CVO UT Co) = eee Ee ee 8 ee er ee 2, 2. Antenna dark gray except at base beneath, segments longer than broad; pygidium and hind leg usually dark _.... obvelatus Bridwell Antenna with apical segment red, segments 7-10 broader than long; hind leg red except along lower margin of femur; pygidium red obtectus (Say ) The strongly sclerotized ovipositor of A. obreptus (fig. 12), as Brid- well has noted, is unlike that of any other known bruchid except the distantly related A. oblongoguttatus (Fahraeus) from Mexico. Acan- thoscelides obtectus is easily recognized by its red abdomen, red terminal antennal segment, and bronzy-gray elytra. Characteristics in the male genitalia are consistent and definitive when proper prepara- tions have been made. Acanthoscelides obreptus Bridwell (Bigss 7,08.) 910; 12) Acanthoscelides obreptus Bridwell, 1942, p. 256. Color.—Mostly reddish dorsally with darker borders on elytra, occasionally darker on disk, vestiture gray with bronzy sheen, with vague maculations of brown hairs, elongate whitish mark on third interval limited anteriorly and posteriorly by brown spots, thorax and middle of abdomen piceous ventrally, 6 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Vag? S N = Figs. 1-15, Acanthoscelides spp. Figs. 1-3, obtectus 6: 1, median lobe, ventral; 2, lateral lobes, ventral; 3, lateral lobes, lateral. Figs. 4-6, obvelatus 6: 4, median lobe, ventral; 5, lateral lobes, ventral; 6, lateral lobes, lateral. Figs. 7-10, obreptus @: 7, median lobe, ventral; 8, apex of internal sac; 9, lateral lobes, ventral; 10, lateral lobes, lateral. Fig. 11, obtectus 2, apex of ovipositor. Fig. 12, obreptus 9, apex of ovipositor. Fig. 13, obtectus, dorsal habitus. Fig. 14, obsoletus, apex of hind tibia. Fig. 15, obtectus, hind leg. Abbreviations: vv, ventral valve; is, internal sac;.ml, median lobe; gs, gonopore sclerites. PROC. ENT. SOC. WASH., voL. 70, No. 1, MARCH, 1968 7 lateral areas of abdomen, antenna, and legs red except for lower margin of hind femur and occasionally median portion of hind tibia. Pygidium reddish to piceous, covered with yellowish vestiture. Male genitalia—Ventral valve with lateral margins noticeably angulate, lateral lobes expanded at apices, overlapping apically, pedicels bowed, dense patch of fine spicules on mesal margin near apex of each lateral lobe (figs. 9, 10), internal sac with fine hairlike spicules only near apical orifice, remainder with ridges and small, blunt denticles nearly to gonopore sclerites, two rows of fine, acute denticles near gonopore sclerites, apex complex (figs. 7, 8). Female genitalia.—(fig. 12) apices of strengthening rods thickened, acute apically, papillae absent. This species is known from Mexico, Guatemala, Panama, Haiti, Trinidad, Venezuela, Colombia, Peru, and Chile. All specimens with recorded host plants are from Phaseolus lunatus L., except the series from Trinidad which was reared from Dolichos lablab L. The species has been intercepted numerous times in Plant Quarantine inspections. The male lectotype (USNM 69235) has been selected from a series of 75 specimens from Colonia Mendoza, Venezuela, reared April, 1938 from Phaseolus lunatus seed by C. H. Ballou. Acanthoscelides obvelatus Bridwell Gbigs.04 5556) Acanthoscelides obvelatus Bridwell, 1942, p. 257. Color.—Piceous, vestiture gray with indistinct brownish maculae on elytra, distinct elongate white mark at middle of third elytral interval, fore and middle legs and posterior margins of abdominal sternites occasionally with reddish suf- fusion; ventral side of basal four antennal segments reddish; pygidium piceous with gray vestiture condensed into fine median line. Male genitalia.—(figs. 4, 5, 6). Median lobe elongate, ventral valve triangular, base hoodlike; lateral lobes somewhat divergent, spatulate, with prominent ridge of fine spicules on mesal margin near apex of each lobe, in lateral aspect strongly angulate at base of apical expansion (figs. 5, 6); internal sac with fine spicules extending about half the distance from apical orifice to gonopore sclerites, then with broad ridges and denticulations in remaining half. Female genitalia—As in A. obtectus (fig. 11). This species was described from a series taken in San Cristobal, Chiapas, Mexico in 1931 in Phaseolus vulgaris L., and the male lecto- type (USNM 69236) was selected from this collection. Subsequent Plant Quarantine interceptions have been made in Phaseolus vulgaris varieties (principally black beans) from Guatemala and Colombia. Acanthoscelides obtectus (Say) Ghisgele ne. 35 ble lanl blige) Bruchus obtectus Say, 1831, p. 1. (LeConte Edition, Vol. 1, p. 259). For further synonymy, see Larsen and Fisher (1938, p. 4-5). Includes Bruchus irresectus Fahraeus, 1839, designated as type-species of Acanthoscelides Schilsky, 1905 by Bridwell, 1929, p. 42. A. obtectus was listed in synonymy of A. irresectus by Schilsky, 1905. 8 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 16-22, Acanthoscelides spp. Figs. 16-17, obsoletus ¢: 16, median lobe, ventral; 17, lateral lobes, ventral. Fig. 18, obtectus, front of head. Fig. 19, obsoletus, ¢ antenna. Fig. 20, obtectus, ¢ & 2 antenna. Fig. 21-22, obsoletus; 21, outline of lateral margin of prothoracic disk; 22, middle of basal abdominal sternite, ¢. Color.—Darker maculations of elytra brownish with bronzy tinge, lighter areas yellowish-gray with bronze overtones, middle of third interval of each elytron with characteristic elongate lighter mark; following areas red: pygidium, lateral areas of abdomen, posterior margin of each abdominal sternite, basal four segments and terminal segment of each antenna, fore and middle legs except occasionally darker at the extreme base of femur, hind legs except along ventral margin of each femur (fig. 15). Male genitalia—(figs. 1, 2, 3). Median lobe elongate; ventral valve triangular, slightly arcuate on lateral margins; lateral lobes (fig. 3) overlapping at apices, mesal margins nearly straight; internal sac with fine spicules extending more than half length of sac, cluster of coarser spicules near gonopore sclerites. Female genitalia —(fig. 11) membranous except at apex; strengthening rods elongate, slender. Acanthoscelides obtectus is an almost cosmopolitan pest of stored Phaseolus spp. bean seeds as well as other commercial seeds and is well known in agricultural regions around the world. Say’s holotype has been lost, but because the species is so universally known, I do not think it necessary to designate a neotype. The three species in the obtectus group can be differentiated by the external characters given in the key and by characters in the male genitalia. The similarity among the three species in the median lobe is apparent in the illustrations, but the most striking differentiating characters are to be found in the lateral lobes. In the female genitalia, those of obtectus and obvelatus are not greatly different from those found in other species now placed in the composite genus Acantho- PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 9 scelides, but the radical form found in A. obreptus suggests that dif- ferent oviposition habits might be found in this species. Acanthoscelides obsoletus (Say) (Figs. 14, 16, 17, 19, 21, 22) Bruchus obsoletus Say, 1831, p. 2. Whether A. obsoletus alone constitutes a species group or will be a member of a larger group awaits a revision of the genus for the New World, a study now in progress. For present purposes, the following characters differentiate A. obsoletus from the members of the A. obtectus group: lateral margins of prothorax nearly straight in dorsal aspect (fig. 21); third and fourth striae approximate at base, each ending basally in a polished black denticle slightly raised above the surrounding area; hind tibia with long, slender mucro (fig. 14); male antenna long, broadly serrate (fig. 19); basal abdominal sternite in male with median elongated depression fringed with incurving hairs (fig. 22); frontal carina evident but not prominent; ventral valve equilaterally triangular; apices of lateral lobes scarcely reaching apex of median lobe, not hoodlike (fig. 17); armature of internal sac composed of coarser spicules and denticles (fig. 16). The female antenna is similar to fig. 20. The hind femur of A. obsoletus is essentially as in the obtectus group (fig. 15). REFERENCES Bridwell, J. C€. 1929. The cowpea bruchid (Coleoptera) under another name —a plea for one kind of entomological specialist. Proc. Ent. Soc. Wash. 31(2): 39-44. 1942. Two new American bean bruchids (Coleoptera). Rev. Chilena Histor. Nat. 44( 1940) :249-258. Horn, G. H. 1873. Revision of the Bruchidae of the United States. Trans. Amer. Ent. Soc. 4:311-342. Larson, A. O. and C. K. Fisher. 1938. The bean weevil and the southern cowpea weevil in California. U.S. Dept. Agr. Tech. Bul. 593, 70 p. LeConte, J. L., G. H. Horn, and J. Leidy. 1877. Report on insects introduced by means of the International Exhibition. Proc. Acad. Sci. Phila. 1876:267-271. Pic, M. 1913. Coleopterorum Catalogus, pars 55, Bruchidae. Junk and Schen- kling, Berlin, 74 p. Riley, C. V. 1892. The pea and bean weevils. Rept. Sec. Agr., Rept. Ento- mologist, 1892:170-172, 2 pl. and L. O. Howard. 1893. On the nomenclature and on the ovi- position of the bean weevil. Insect Life 5:27-33. Say, T. 1831. Descriptions of North American Curculionides and an arrange- ment of some of our known species agreeably to the method of Schoenherr. New Harmony, p. 1-30. (LeConte Ed. 1:259-299) Schilsky, J. 1905. In Kiister and Kraatz, Die Kafer Europa’s, Heft 41 (Bruchi- dae), 101 p. 10 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 A NOTE ON THE SLEEPING BEHAVIOR OF PHILANTHUS GIBBOSUS (FABRICIUS) (HYMENOPTERA: SPHECIDAE ) NorMan Lin, Department of Zoology and Entomology, The Ohio State University, Columbus, Ohio 43210 The sleeping behavior of solitary bees and wasps has recently received considerable attention (Evans and Linsley, 1960; Linsley, 1962). However, our state of knowledge of this most interesting subject is at present quite fragmentary. Philanthus is no exception, with the sleeping behavior of most species being unreported. Several brief accounts, however, have appeared on the sleeping behavior of P. gibbosus (Fabricius) (Peckham and Peckham, 1898; Rau and Rau, 1918; Reinhard, 1921-22) and the present study supports these accounts. In the present study, a lone male of P. gibbosus was observed on July 4, 1966, in the Parade Grounds, a 40-acre field used for sandlot baseball in Brooklyn, New York. The male entered a hole or nest in the ground at 6:45 P.M. and was captured when it came out seconds later. This male was probably searching for a place to spend the night in accord with the observations of the Peckhams (1898). Later in the summer (August 9), a small colony of P. gibbosus was dis- covered in a different location in the Parade Grounds. Living wasps were last observed in this colony on September 2. On August 9, the colony consisted of a total of six nests devoid of any vegetational cover and located in the flat ground of a baseball diamond consisting of hard, bare, sandy soil, seemingly with a high clay content. Evans and Lin (1959) found nine of ten nests studied dug into the side of vertical sand banks, and the tenth into flat sand. The two closest nests in the Parade Grounds colony were approximately 18 inches apart and the two furthest were approximately nine feet apart. No nest was more than an estimated five feet ten inches from its nearest neighbor, and all but one were approximately four feet or less from their nearest neighbor. At times, the heads of females could be seen “peering” from the entrances of nests. When the nests were approached the females usually withdrew within the nests, only to again reappear at the entrances within minutes. Such behavior was noted on several days. It was possible by approaching such nests cautiously to capture females in the entranceway before they could withdraw. Evans and Lin (1959) also noted such “peering” behavior and found that it preceded flight. At 8:15 A.M. on August 30, the colony was again visited. Several nests were observed, all had outer closures, and three appeared to have been closed from the inside. A few other nests probably had permanent closures. Rectangular glass containers were placed over PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 11 the entrances of the three nests seemingly closed from the inside, so as to prevent wasps from entering or leaving the enclosed area. The observer then left the colony. When he returned at 10:30 A.M., he found wasps under two of the three glass traps. One trap contained a female and the other contained a male and a female. All three nests were now open. The male and female were captured for identifica- tion. The wasps had been in the nests the entire time and when they emerged between 8:15 A.M. and 10:30 A.M., they were detained by the traps. Presumably these wasps had spent the night in the nests. On August 31 at 7:35 A.M. three nests were found, and all were closed. They were covered with glass traps and when reexamined at 9:20 A.M. and 11:30 A.M. all were open. No Philanthus, however, were in the traps. The nests had apparently been closed from the inside where the wasps had spent the night, and after opening the nests, the trapped wasps apparently reentered without again closing them. At 8:50 A.M. on September 2, two nests (both closed ) were located, and then covered with glass traps. One nest had previously been covered on August 31. Between 8:50 and 8:52 a P. gibbosus was observed to land for a few seconds on paper, and two others were seen in flight, one in an orientation flight. Wasps were not under the traps at 10:35 A.M. and 12:15 P.M. On September 5 at 12:43 P.M. and 7:00 P.M. only one nest could be found in the colonial area and there was no evidence that it was recently active. Wasps were also not observed on September 9 at 1:30 P.M. nor September 18 at 12:23 P.M. A few traces of nests were found on September 9 and none on September 18. DIscussION According to the Peckhams (1898) and Reinhard (1921-22) both males and females of Philanthus gibbosus temporarily share a common nest which includes sleeping in the nest. One such nest described by the Peckhams (1898) housed three males and four females. The Peckhams maintained that such wasps living in a common nest were siblings, and Reinhard (1921-22) likewise suggested that this might be the case. These authors, however, do not present any data to support this view. The Raus (1918) also found that females of P. gibbosus spend the night in the nest, but presented no data on males. The present study supports the previous findings that males and females may spend the night together in the nests which are closed from the inside. In addition it was found that females apparently also spend the night by themselves in the nests which are likewise closed from the inside. No data, however, was obtained suggesting males and females using the same nest are siblings. Evans and Lin (1959) found that females of Philanthus politus politus Say and 12 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Philanthus bilunatus Cresson spend the night in the nest which is then closed from the inside. Closure of the nest at night by Philanthus probably is an adaptation for keeping certain parasites or predators from entering the nest and in addition may also function in maintain- ing a more stable microclimatic environment within the nest. Both sexes sleeping together in the nest appears to be relatively unknown among digger wasps. Another example is the nyssonine wasp Clitemnestra gayi (Spinola); the males and females are reported to spend the night together in the nest galleries (Janvier, 1928). Reinhard (1921-22) found that P. gibbosus had two generations a year in Maryland. The first generation emerges during May and June and are found to be in “full swing” by the end of June. The second generation begins to make its appearance during the middle of July, and the gradual emergence continues until almost the end of August. In the present study a male was captured on July 4 and another was captured on August 30. It is well known that the males of solitary wasps are generally shorter lived than the females, and the presence of males in early July, and late August or early September suggests more than one generation. Furthermore, the limited data fits the Reinhard (1921-22) distribution, suggesting two generations a year in the Parade Grounds in New York. Evans and Lin (1959), however, state that P. gibbosus probably has one generation a year in Ithaca, New York. According to Evans and Lin (1959), P. gibbosus makes its appearance in Ithaca in early July and remains active until late September. Of course the data in the present study is fragmentary and it is by no means conclusive that P. gibbosus has two generations a year in Brooklyn. I am indebted to Dr. Howard E. Evans for identifying the Philan- thus. Funds for the summer of 1966 were provided in part by a Sigma Xi Grant-in-Aid of Research. REFERENCES Evans, H. E. and E. S. Lin. 1959. Biological observations on digger wasps of the genus Philanthus (Hymenoptera: Sphecidae). Wassmann J. Biol. 17:115— 1382: Evans, H. E. and E. G. Linsley. 1960. Notes on a sleeping aggregation of soli- tary bees and wasps. Bull. S. Calif. Acad. Sci. 59:30-37. Janvier, H. 1928. Recherches biologiques sur les predateurs du Chili. Ann. Sci. Natur., Zool. (10)11:67—207. Linsley, E. G. 1962. Sleeping aggregations of Aculeate Hymenoptera—lII. Ann. Ent. Soc. Amer. 55:148—164. Peckham, G. W. and E. G. Peckham. 1898. On the instincts and habits of the solitary wasps. Wisc. Geol. Natur. Hist. Surv. Bull. 2:245 pp. Rau, P. and N. Rau. 1918. Wasp Studies Afield. Princeton University Press, Princeton, New Jersey. Reinhard, E. G. 1921-1922. The life history and habits of the solitary wasp Philanthus gibbosus. Ann. Rept. Smithsonian Inst., 363-376 pp. PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 13 A NEW GENUS AND THREE NEW SPECIES OF ERYTHRAEOIDEA (ACARINA: ERYTHRAEIDAE AND SMARIDIDAE ) Rospert L. SMILEY, Entomology and Research Division, ARS, U.S. Department of Agriculture, Washington, D.C. 20250 According to Baker and Wharton (1952), adults of the family Smarididae can be distinguished from those of the Erythraeidae by mouth parts that can be withdrawn into the body. Presently, this is the only practical character for separation; otherwise the resemblance between the families is marked. “Although in the adults the difference between these two families is striking—the presence of the armilla to the mouthparts being a character of undoubted family significance— in the larval Smarididae no feature can be seen in the mouthparts that can be considered of family significance” (Southcott, 1961). Southcott defines the armilla as the extensile collar bearing the gnathosoma in the Smarididae; the armilla is absent in the Ery- thraeidae. Also, Southcott (1946, 1948) used the tarsal claws to dis- tinguish larval smaridids from erythraeids and (1961) presented a detailed account of the validity of this character. Included in the present paper are descriptions of two erythraeids belonging to the genera Balaustium Heyden and Sphaerolophus Berlese. The location of the eyes and sensory pits and the shape of the palpal tarsi will separate the two genera. A new genus, Paraphanolophus, is being erected to include a larval form belonging to the family Smarididae. The new species of Balaustium and Sphaerolophus were sent to me by Dr. W. L. Putman, Research Branch, Canada Department of Agriculture, Vineland Station, Ontario, Canada. The Balaustium species was reared and used in an insemination study (Putnam, 1966). Balaustium putmani, n. sp. (Figs. 1-10) This species is characterized by the anterior portion of the crista being truncate in both sexes. Female. Palpi long and strong; palpal tibia with short, stout distal claw possessing a single tooth (fig. 1); palpal setae long and slender, lightly serrated, intermixed with simple setae; palpal tarsus elongate, longer than tibia, with 10-12 strong solenidia (fig. 2) apically. Anterior venter of gnathosoma (fig. 3) has a cupshaped structure, with fingerlike projections, and with a pair of spurs ventrally; medially with 5 pairs of short, lightly serrated setae and 7 pairs of similar but longer setae laterally and posteriorly. Anterior dorsum of gnathosoma (fig. 4) with a pair of short simple setae and a longer serrate pair. Setae (fig. 5) of dorsal propodosoma and hysterosoma serrated, subequal in length; those of venter (fig. 6) simple. A single pair of lenslike eyes and a single pair of sensory pits located at about the posterior level of the crista (fig. 9). Crista (fig. 8) 14 PROC. ENT. SOC. WASH., VoL. 70, No. 1, MaRcH, 1968 vil fy | \ /s ied Vay /\ b/ d | 1 \ it Hy a ee We ye Ae \/ . fire h\ = ji ‘ VAi\\ Pantin’ y SAN) oat en Figs. 1-8, Balaustium putmani, n. sp., 2: 1, left palpus; 2, palpal tarsus; 3, venter of gnathosoma; 4, dorsum of gnathosoma; 5, dorsal setation; 6, ventral setation; 7, striae above crista metopica; 8, crista metopica. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 15 long, narrow, lightly punctate, with a strongly sclerotized ribbonlike longitudinal median band; a pair of medium-length serrated sensory setae on the anterior and posterior area; posterior sensory setae arising from noselike projections of scutum; with 9 stout serrated setae anteriorly to the anterior sensory setae, 3 pairs of shorter setae, and a single subequal serrate seta between the anterior and posterior sensory setae. A fine distinct ornamentation of the striae (fig. 7) anteriorly to the crista. Legs I one-third longer than legs IV; legs II shortest; legs III slightly longer than legs II; legs IV slightly longer than legs III. Tarsus I (fig. 10) with simple setae dorsally (T); apically with few lightly serrated (A); laterally with solenidia (X) intermixed with many comblike setae (V) at the anterior ventral portion, extending to about midlength of tarsus; posteriorly and ventrally with few smaller serrated setae (Z). Tibia I dorsally with few serrated setae varying in length (A), and solenida (X); laterally with many simple setae (T). Tarsus I 186 uw long and tibia I 226 uw long. Length of body, not including gnathosoma, 1600 xu. Male. Crista metopica similar to that of female, long, narrow, lightly punctate, and with a strong sclerotized ribbonlike longitudinal median band; with a pair of slender serrated sensory setae on anterior and posterior area; with 5 stout serrated setae anteriorly to anterior sensory setae; 4 pairs of stout serrate setae between anterior and posterior sensory setae. Dorsal and ventral body setae similar to those of female. Length of body, not including gnathosoma, 840 u. Holotype. Female, Canadian National Collection No. 9451, was collected and reared from plum bark, Vineland Station, Ontario, Canada, by W. L. Putman, 28 August 1961 for whom the species is named. Paratypes. 2 males and 7 females with the above data. Three females and a male paratype are in the U.S. National Museum Collec- tion, and the rest are in the Canadian National Collection. Sphaerolophus canadensis, n. sp. (Figs. 11-19) This is the first species of this genus recorded from North America. The species is distinctive in that the posterior portion of the crista is beneath the cuticle. Female. Palpi strong; palpal tibia with short claw (fig. 11); palpal tarsus strong, globe-shaped, with subequal finely serrated setae dorsally, and with few shorter finely serrated setae apically; with annulated solenidia ventrally. Setae of the dorsal propodosoma (fig. 12) strong, serrated, and subequal. Venter of propodosoma (fig. 13) with serrated setae varying in length and size. Dorsally a single pair of lenslike eyes (fig. 14) located adjacent to middle of crista. Crista (fig. 15) long, narrow, slightly rounded anteriorly and pointed posteriorly; with a narrow, ribbonlike, longitudinal band; anteriorly with a pair of slender, lightly serrated sensory setae and 5, long, stout, strong serrate setae; posteriorly with a single pair of serrated setae slightly longer than anterior pair; median lateral area of crista intermixed with 4 pairs of stout serrated setae varying in length and with 5 pairs of shorter and smaller serrated setae. Striae (fig. 16) anterior to crista as figured. Venter of gnathosoma (fig. 17) anteriorly with fingerlike projections and with 10 pairs of serrated setae; subequal in length; posteriorly 16 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 Vv \ | | A Figs. 9 and 10, Balaustium putmani, n. sp., 2: 9, right eye; 10, right tarsus and tibia I. Figs. 11 and 12, Spaerolophus canadensis, n. sp., 2: 11, right palpus; 12, dorsal setation. qi PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 zg Wt, Wz, S& NV Wit A Soy 1°) GOUU0Y Figs. 13-18, Spaerolophus canadensis, n. sp., 2: 13, ventral setation; 14, right eye; 15, crista metopica; 16, striae above crista metopica; 17, venter of gnatho- soma; 18, dorsum of gnathosoma. 18 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Sy AY a 1% @ B\- AKG ee 3/7 (NE fi -— aV = ; 4/ N / Fig. 19, Spaerolophus canadensis, n. sp., 2: diagrammatic presentation of tarsi and tibiae I-IV. Figs. 20-23, Paraphanolophus metcalfei, n. sp., larva: 20, dorsum; 21, left palpus; 22, dorsum of gnathosoma; 23, venter of gnathosoma. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 19 1 TA i i ot) Ay rh ny nOSOOOPD Dag) WAIN owe ver ee AR ps ee A a ce =DAn, DODOIOSOOS Vee CeCe rece Figs. 24 and 25, Paraphanolophus metcalfei, n. sp., larva: 24, venter; 25, leg III. with 30 pairs of longer and stronger serrated setae. Dorsum of gnathosoma with 4 pairs of finely serrated seta (fig. 18). Legs I longer than body; legs II shortest; legs III slightly longer than leg II, but not as long as legs I; legs IV longest; leg setae serrated, long, and numerous. Tarsi (fig. 19) of all legs slightly oval. 20 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Tarsus I 253 uw long and tibia 493 uw. Length of body, not including gnathosoma, 1986 uw. Male. Not known. Holotype. Female, Canadian National Collection No. 9450, was collected from plum, Vineland Station, Ontario, Canada, 23 July 1964 by W. L. Putman. Paratypes. Two females with the above data. A paratype is in the U.S. National Museum Collection and the Canadian National Collec- tion. Paraphanolophus, n. gen. Type-species: Paraphanolophus metcalfei, n. sp. This genus is distinctive in having the following: the tarsal claws are sym- metrical and lightly ciliated along the ventral sides; the empodium is strong, falciform and nonpulvilliform; the scutal sensilla bases (anteriorly and posteriorly ) are close together; there are only 2 pairs of ventral setae in the area bounded by coxae II and III, and at the anterior level of coxae III and extending to the posterior of hysterosoma there are 45 pairs of serrate setae subequal in length. According to Southcott’s (1961) key to the genera of the larval Smarididae, this species is near the genus Phanolophus Andre but differs by having the fore- mentioned characters. Paraphanolophus metealfei, n. sp. (Figs. 20-25) Larva. Body (fig. 20) globe-shaped with 2 eyes on each side near coxae II. Dorsal setae serrated, consistent in size and length medially; but varying in length marginally. Scutum rounded, excavated anteriorly, and lightly sclerotized; with a pair of long posterior and a pair of shorter anterior sensillae plus 2 pairs of serrated setae near antero lateral margin of scutum; second pair of serrated setae about one-fourth longer than first pair. Previous to remounting this specimen had all 4 sensillae. Venter of hysterosoma (fig. 24) with 45 pairs of serrate setae, subequal in length, smaller in size than setae of dorsum. Dorsum of gnathosoma (fig. 22) as figured. Venter of gnathosoma (fig. 23) with fingerlike projections and, surrounding oral cavity, the ciliae form a flange; with a pair of strong chelicerae; a pair of simple setae posterior to flange. Palpi (fig. 21) strong; palpal femur with a stout serrated seta, palpal genu with a stout serrated seta subequal in length to seta of femur; palpal tibia with 1 serrated and 2 simple setae (the anterior simple seta longer and stronger than posterior simple seta), with stout bifurcate distal claw; palpal tarsus with 2 subequal solenidia, 3 simple setae subequal in length, and an eupathid. Legs I and II subequal in length; legs III (fig. 25) slightly longer; all legs with coarse serrated setae and a few finer subequal serrated setae distally. Each tarsus with a strong distal bothridon dorsally and ventrally. Tarsus III 172 u long and tibia III 346 uw long. Setal for- mula for legs I-III, coxae 1-1-1; trochanter 1-1-1; basifemora 4—4—4; telofemora 5-5-5; genua 9 + 2 — 8 — 8; tibiae 14 + 2-104 1-13 +4 1; tarsi 24-24-24. The formula includes tactile and solenidia. Body, excluding gnathosoma, 680 u long. Adults and Nymphs. Not known. PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 21 Holotype. Larvae, USNM 3227, was collected on Saccharosydne saccharivora Westwood, British Honduras, 20 January 1967 by J. R. Metcalfe for whom this species is named. Paratype. A larva with the above data is in the U.S. National Museum Collection. REFERENCES Baker, E. W. and G. W. Wharton. 1952. An Introduction to Acarology. The Macmillan Co., New York, N. Y., 465 pp. Putman, W. L. 1966. Insemination in Balaustium sp. (Erythraeidae). Acaro- logia 8(3) :424427. Southeott, R. L. 1946. On the family Smarididae (Acarina). Proc. Linn. Soc. N.S. W. 70(3—4):173-178. 1948. Larval Smarididae (Acarina) from Australia and New Guinea. Proc. Linn. Soc. N.S. W. 72 (5-6) :252-264. 1961. Studies on the systematics and biology of the Erythraeoidea (Acarina), with a critical revision of the genera and subfamilies. Austral. J. Zool. 9:367-610. A REPORT ON THE SALDIDAE COLLECTED BY THE GALAPAGOS INTERNATIONAL SCIENTIFIC PROJECT 1964 (HEMrIPTERA )+* Joun T. Potnemus, 3115 S. York, Englewood, Colorado 80110 The following report is based on the material collected by members of the Galapagos International Scientific Project of 1964.7 This material, exclusive of types, has been divided, as quantity permits, among the following individuals and institutions: California Academy of Sciences, R. L. Usinger, P. D. Ashlock, J. T. Polhemus, U.S. Na- tional Museum and B. P. Bishop Museum. To date only one saldid has been noted from the Galapagos, Pentacora sphacelata (Uhler). This report adds a new species of the genus Saldula and notes on a saldid taken at light on a ship off Ecuador. Pentacora sphacelata ( Uhler) Salda sphacelata Uhler, 1877, Bull. U.S. Geol. Geogr. Surv. 3:434-436 (Mas- sachusetts, Maryland). Salda rubromaculata Heidemann, 1901, Proc. Wash. Acad. Sci. 3:368-369 ( Albe- marle I., Galapagos). The specimens of this species from the Galapagos seem quite typical, with no apparent shift in characters from our mainland material. P. sphacelata has been collected along the west coast of the 1 Contribution No. 83 from the Charles Darwin Foundation for the Galapagos. 2 Supported in part by Grant GE-2370 from the National Science Foundation. 22, PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Americas as far south as Guatemala, and further collecting will prob- ably extend the range of this even further southward on the mainland. Specimens from the Galapagos material have been compared pre- viously by Cobben (1965) to sphacelata and a closely related species, Pentacora saratogae Cobben, and his view concurs with mine. Material: 6 66, 13 22, Galapagos Arch., Isla Baltra, 3-II-1964, D. Q. Cavagnaro; 14, Galapagos, Baltra Is., (S. Seymour), Feb. 3, 1964, P. D. Ashlock Collector; 1 ¢, 4 22, Galapagos Arch., Tortuga Bay, Santa Cruz Is., II-10-1964, Heleotropium curassavicum, G. Kus- chel Collector. Saldula sectilis (Hodgden) Salda sectilis Hodgden, 1949, Jour. Kansas Ent. Soc. 22:160-161 (Canal Zone). One specimen is present in the material, with the head and abdomen missing. The remains must be attributed to Saldula sectilis, as the unusual hemelytral pattern (fig. 2) is similar to the type of sectilis which I recently studied at the U.S. National Museum. Also, the coloration of the legs and shape of the pronotum match Hodgden’s description and my type notes. Material: One broken specimen, Ecuador, Guayaquil, Mar. 5, 1964. At light on ship, P. D. Ashlock Collector. Saldula galapagosana, n. sp. Of small size, moderately broad, general color piceous, macropterous. (For all measurements, 60 units equal 1 mm.) Head: Black, shining, frons and vertex rugulose; preocellar spot yellowish; covered with short golden hairs and usual three pairs of long erect black hairs on frons and vertex; ocelli raised slightly and separated by approximately the width of an ocellus; rostrum brown, extending between hind coxae; clypeus and anteclypeus testaceous. Thorax: Pronotum black, shining, rugulose, covered with short decumbent golden pubescence; lateral margins very slightly convex, narrowing moderately anteriorly; callus moderately raised, with deep depression in center, anterior lobe longer than posterior lobe (13/8, not including collar); underparts black, clothed with fine silver pubescence; scutellum width subequal to length (42/41), with vestiture similar to pronotum, depressed tranversely at center. Wings: Hemelytra fully developed, covered with short decumbent golden and black pubescence; ground color dark brown to black, dull, very finely rugulose; pattern of markings as shown in fig. 1A, the three light markings in the inner corium being bluish-white pruinose areas in holotype, much bluer in some other specimens; embolar region largely flavo-testaceous in holotype, much darker in some other specimens; light brown to greenish pruinose areas occurring in an irregular elongate pattern on the inner corium and a medial area on the outer corium next to the corial suture; membrane fumose, with darker fumose markings, with four cells. Abdomen: Piceous, covered with short, decumbent silver pubescence; caudal margins of segments sometimes narrowly margined with ochroleucus; in female, subgenital plate with caudal half ochroleucus to leucine, this portion being PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 23 'Aa_mm Fig. 1, Saldula galapagosana, n. sp.: A, left hemelytron; B, C, left ¢ paramere, two views; D, parandria; E, ¢ coupling plate; F, aedeagus, showing sclerotized structures. Fig. 2, S. sectilis (Hodgen), left hemelytron. produced posteriorly; parandria as shown in fig. 1D; male coupling plate very narrow, with thirteen stout pegs as shown in fig. 1E. Extremities: Antennal segment 1 flavo-testaceous, segment 2 testaceous, seg- ments 3 and 4 blackish brown; segments 3 and 4 somewhat thickened, subequal in diameter to segment 1, segment 2 slenderest; segments 1, 3 and 4 clothed with short yellowish hairs, segment 2 clothed with brown hairs equal in length to segment width; segments 2, 3 and 4 with scattered longer hairs; antennal segments proportioned as follows: holotype ¢, 14:24:22:26; allotype 2, 18:30:23:28. Legs leucine to testaceous; femora testaceous on apical half, becoming leucine or ochroleucus at base; tibia ochroleucus, testaceous at base, sometimes with testaceous band at center and at apex, with usual dark spines. 94 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Genitalia: Paramere as shown in figs. 1B and C, rugulose on main body and processus sensualsis; median sclerotized structure of aedeagus as in fig. 1F; filum gonopori coiled two and a half times. Measurements: Holotype ( é¢), length 3.05 mm, width 1.35 mm; allotype (2), length 3.35 mm, width 1.70 mm. Mean length of 10 ¢ ¢, 2.99 mm; min. 2.80 mm, max. 3.15 mm. Mean width of 10 ¢ 6, 1.39 mm; min. 1.35 mm, max. 1.60 mm. Mean length of 6 9 2, 3.26 mm; min. 3.14 mm, max. 3.35 mm. Mean width of 6 9 9, 1.67 mm; min. 1.60 mm, max. 1.75 mm. Material: Holotype, male (California Academy of Sciences), Gala- pagos Arch., Isla Santa Cruz, Grassland 750 M., IV-6, 1964, D. Q. Cavagnaro Collector; Allotype, female (California Academy of Sci- ences), Galapagos Arch., Isla Santiago, N.W. slope, 600 M., V-30, 1964, D. Q. Cavagnaro Collector. Paratypes as follows: 16 46,4 22, 1 nymph, Galapagos Arch., Isla Santiago, N.W. slope, 600 M., V-—30, 1964, D. Q. Cavagnaro Collector; 1 ¢, 2 22, Galapagos, N. of Academy Bay, Santa Cruz Is., Miconia Belt, 1300’ Elevation, I-20, 1964, P. D. Ashlock Collector; 1 2, 1 nymph, same island, collector and date, Grassland, 2100’ Elev.; 2 nymphs, same island and collector, Grassland, 1800’ Elev., II-18, 1964; 2 22, Galapagos Arch., Bella Vista, Santa Cruz I., 500 M., I-31—64, G. Kuschel Collector. Comparative Notes: Saldula galapagosana is most closely related to an undescribed species from Ecuador (contained in my collection), but also bears a resemblance to S. penningtoni Drake. The latter has male parameres of a very different form however; penningtoni stands very close to S. dentulata (Hodgden) in this respect, both species having the processus sensualsis produced. (An excellent figure of the paramere of S. dentulata is given by Cobben, 1960.) The whitish blue to blue pruinose areas on the inner corium will separate gala- pagosana from the closely related Saldula species known to me. According to Drake’s treatment of the saldids of Micronesia (Drake 1961), galapagosana does not closely resemble any of the species known from that region. Zoogeography: The saldid fauna of the Galapagos seems closely allied to that of central and south America, although our knowledge of the latter fauna is still rather poor. Saldula galapagosana is most likely a sibling species arising from the same stock as the undescribed species mentioned before. REFERENCES Cobben, R. H. 1960. The Heteroptera of the Netherlands Antilles—III. Saldi- dae. Studies on the Fauna of Curacao and other Caribbean Islands 9:44-61. The Hague. 1965. A new Shore-bug from Death Valley California. Pan-Pac. Ent. 41(3):180-185. Drake, C. J. 1961. Hemiptera: Saldidae. In B. P. Bishop Museum, Insects of Micronesia 7(6):287—305. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 25 NOTES ON NORTH AMERICAN PIOPHILIDAE. III.* (DreTERA ) GrorGE C. STEYSKAL, Systematic Entomology Laboratory, U.S. Department of Agriculture, Washington, D.C. 20560 The types of some species of Piophila subgenus Liopiophila de- scribed by Melander and Spuler (1917) and Melander (1924), now deposited in the U.S. National Museum, were dissected in order to examine the male postabdomen and thus more firmly establish their identity. Lectotypes have been selected for three species, the names of two of which are now in synonymy. Bibliographic citations for all names will be found in the Catalog of the Diptera of America north of Mexico (Stone et al., 1965). Piophila (Liopiophila) nigrimana Meigen (Fig. 1) Piophila occipitalis Melander and Spuler, 1917. New synonym. Piophila morator Melander, 1924. New synonym. Piophila privigna Melander, 1924. Synonymy confirmed. Examination of the male postabdomen of European and North and South American material of this species, as well as the types of P. occipitalis (lectotype from “Chicago, Illinois”), P. morator (unique male from “Pullman, Washington”), and P. privigna (lectotype from “Woods Hole, Massachusetts, 7-21-2”), has revealed a wide variation in general body color but a consistent and characteristic conformation of the postabdomen (fig. 1), with the bent peg on the sternum men- tioned in my last note in this series. The peg is best considered as situated on the margin of the 7th sternum, rather than on the 6th, though these two sclerites are more or less fused. The long, coiled aedeagus is bare for an appreciable distance basally, and a ligulate sclerite (likely the anterior gonite) bears a pair of anteriorly directed setae, usually dark in color. The hypandrium in ventral view (h) is rather broadly bridged across where the basal part of the basiphallus is invaginated to form the phallapodeme. The posterior margin of the 5th sternum is decidely sinuate. Some of the material examined has the characters of coloration ascribed to Piophila nigricornis Meigen, a rarely captured European species. In view of the variation in color in extra-European material, it is highly probable that P. nigricornis also will be found to be merely a color variant of P. nigrimana. The front of P. nigrimana is usually largely yellowish, the antenna often infuscate apically, and the fore coxa and femur are sometimes wholly black, though usually the fore coxa is yellow and the fore femur is yellow basally for some distance. 1 See these Proceedings 60:246, 1958 (paper I); 66:177-181, 1964 (paper II). 26 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Fig. 1. Piophila (Liopiophila) nigrimana Meigen, male, Chicago, [Illinois (lectotype of P. occipitalis Melander and Spuler), postabdomen. h = ventral view of hypandrium; fig. 2. Piophila (Liopiophila) nigricoxa Melander and Spuler, male, Pullman, Washington (lectotype), postabdomen. h = ventral view of hypandrium. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 DAT Piophila (Liopiophila) nigricoxa Melander and Spuler (Fig. 2) The postabdomen of P. nigricoxa is distinctly different from that of P. nigrimana; the peg of the 7th sternum is inclined but not bent, and the sclerotization of the 6th and 7th sterna differs in a number of ways. The long, coiled aedeagus is haired nearly to the base; the broad anterior gonite bears a long, apically directed, usually pale seta; and the hypandrium (h) is only narrowly bridged across where it becomes invaginated to form the phallapodeme. The posterior margin of the 5th sternum is virtually straight. External characters distinguishing P. nigricoxa from P. nigrimana are hard to find. In the available specimens, the front is black posteriorly and becomes abruptly yellowish at about % of the distance from the anterior ocellus to the anterior frontal margin; the antenna is bright orange-yellow; and the fore coxa and femur are black, except the extreme tip of the latter. At each side of the lower part of the back of the head in both species is a broad band of whitish pruinosity. A male from “Pullman, Wash.” has been selected as lectotype. REFERENCES Melander, A. L. 1924. Review of the dipterous family Piophilidae. Psyche 31:78-86. and A. Spuler. 1917. The dipterous families Sepsidae and Piophili- dae. Bull. Wash. Agr. Exp. Sta. 143:1-103. Stone, A., et al., eds. 1965. A catalog of the Diptera of America north of Mexico. U.S. Dept. Agr., Agr. Handbook 276:1—-1696. THE GENUS MICROPLATYMERUS OF THE SEYCHELLES! (COLEOPTERA: CURCULIONIDAE; COSSONINAE ) ExLwoop C. ZIMMERMAN, Bishop Museum, Honolulu, Hawaii In 1908, Hugh Scott collected a large series of a small, amber-colored weevil from leaf bases of the strange coco-de-mer palm (Lodoicea maldivica) on Praslin Island, Seychelles, Indian Ocean. These were described as a new genus and new species, Microplatymerus lodoi- ceivorus, by George Champion in his report on the Curculionidae collected by the Percy Sladen Trust Expedition to the Indian Ocean in 1905 [Trans. Linnean Soc. London (II-Zoology) 16(14):451—452, pl. 24, figs. 37, 37 a—-c, 1914], and since then the weevil has remained an enigma. Most unfortunately, numerous errors were made by Cham- 1 This is number 14 of a series of reports resulting from studies made possible by National Science Foundation Grant G-18933. Figs. 1-3. _Microplatymerus lodoiceivorus Champion, dorsal, lateral and ventral views. Total length: 2.1 mm. [B.M. (N.H.) photos. ] PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 29 pion, and he wrongly concluded that the weevil belonged to the Antliarhininae. The Antliarhininae is a group of strange weevils attached to cycads in south Africa, and Microplatymerus bears no relationship to that group. The Antliarhininae have straight antennae, and they belong to a different major section of the Curculionidae than does Microplatymerus which has geniculate antennae. In Coleopterorum Catalogus, pars 146, 1936, the four genera Ant- liarhinus, Platymerus, Microplatymerus and Hoplorhinus are listed by Klima in the Antliarhininae. Only Antliarhinus and Platymerus belong to the subfamily. Hoplorhinus was removed from the Antliarhininae later in 1936 by Csiki who placed it in Cossoninae: Trypetesini in pars 149 of Coleopterorum Catalogus. Much earlier, Champion, in Biologia Centrali-Americana (Insecta: Coleoptera) 4(4):277, 1903, had _ re- moved Hoplorhinus from the Antliarhininae and placed it in his “Group Hoplorrhina,” which he then considered distinct from the Trypetesinae. (It should be noted that Champion used the term “group” for what generally are called subfamilies today.) He altered his opinion, however, when he published his 1910 work in Biologia Centrali-Americana (Insecta: Coleoptera) 4(7):4, 5, and he concluded that Hoplorhinus should be assigned to the Trypetesinae. Whether Hoplorhinus is or is not a member of the Trypetesinae, and whether there is such a natural group as the Trypetesinae, are separate ques- tions beyond the scope of this report. The facts that Hoplorhinus is not a member of the Antliarhininae and is not related to Microplaty- merus are what concern us here. The remaining genus, of the group under discussion, is Microplatymerus, and that genus must be trans- ferred to the Cossoninae, as demonstrated below. It is rather difficult to understand what led Champion to place Microplatymerus in association with such an aberrant group as the Antliarhininae, because Microplatymerus has various typical features of the Cossoninae. Champion said that “Microplatymerus has the general facies of a small calandrid, and the broad flattened sterna and ascending mesothoracic epimera are suggestive of certain Madarides of the group Barina.” To me, it has the facies of a cossonid and bears little resemblance to the calandrids, excepting perhaps in a very vague way to some Sitophilus. Moreover, the mesepimera are in no sense ascending as they are in the Baridinae. The mouthparts are typically cossonid, the metepisternal suture bears the nearly diagnostic row of specialized squamae (“sclerolepidia”) of true cossonids, and the rectum has the sclerotized “loop” of the typical Cossoninae strongly formed, as my fig. 11 demonstrates. There is no doubt that Micro- platymerus belongs to the Cossoninae. There are various misleading or erroneous statements in the original description of the genus, and these require discussion. Champion said that the rostrum is short, but it is not unusually short. It is longer 30 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 4-8. Sketches of details of a male Microplatymerus lodoiceivorus Cham- pion. 4 and 5, dorsal and lateral views of aedeagus and associated parts; 6, outline of pygidium; 7, ninth sternite (“spiculum gastrale”) with a separate sketch of its cephalic end in broad view at 8. All drawn to same scale. than the head and one-half as long as the pronotum. The mandibles appear normal and not “scarcely visible” as stated by Champion. The submandibular, tooth-like projection of the gena is well developed. The postocular constriction is feeble and, as viewed from above, is contiguous with the posterior margins of the eyes; and, as viewed from the side, the median dorsal contour of head and rostrum is only gently depressed above the eyes. The antennae are inserted at only about one-half the length of an eye in front of the eyes, and the stout, strongly clavate scape extends slightly caudad of the lower posterior margin of the eyes when at rest. There is no suprascrobal sulcus (“secondary scrobe” ), and the scrobe curves rapidly beneath the eye. The scrobe is not “shallow” as originally stated. The antennae are typically cos- sonid; the club is subequal in length to the preceding four funicular segments, it is only slightly broader than the fifth funicular segment, and its shiny basal segment is more than twice as long as the remainder of the club. The flat scutellum is not “very small” as originally de- scribed, but its breadth is subequal to the combined breadths of the PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 iil ECZ O25 Figs. 9-16. Outline sketches of details of a female Microplatymerus lodoi- ceivorus Champion. 9, genitalia; 10, eighth sternite; 11, sclerotized “loop” of rectum (the male has a similarly formed loop); 12, pygidium; 13, ectal face of a metatibia, setae omitted; 14, metatarsus, setae omitted; 15, head and rostrum; 16, antenna, setae omitted. The scale line near 9 applies only to 9, that near 11 applies to 10, 11, and 12, that below 15 applies to 14, 15, and 16, and fig. 13 is drawn to a larger scale. first intervals of the elytra (measured across the suture at about the basal one-fourth). The decurved, golden hairs on the dorsum are especially conspicuous on the elytra and most are directed obliquely mesad; the two lateral intervals have only small hairs and are mostly bare. Elytral stria 10 is complete to the apex. The flattened ventral surface, with the pro-, meso- and metasternum forming a continuous level, together with the short, broad form of the weevil no doubt reflect its mode of life in the palm fronds. Although the sutures between the procoxae and the mesocoxae can each be seen through the transluscent derm as a dark line, the sutures do not indent the 32 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 smooth surface. The procoxae are separated by twice the transverse diameter of a coxa, and the distance between them is only slightly less than the distance between the mesocoxae. The metacoxae are separated by only a little more than one-half the separation of the mesocoxae, and this is a rather unusual character. The mesepimera are not unusual as was suggested by Champion, and they do not extend dorsad to truncate the elytral humeri. The mesepimeral suture is mostly obliterated. Champion misinterpreted the metepisternum when he said “meta- thoracic episterna rather broad, polished, fused with the sternum.” He evidently confused the internal structure, which can be seen through the transluscent derm, instead of using the true metepisternal suture as the external boundary. The suture is clearly marked by the line of peculiarly modified squamae or “sclerolepidia” along it, and the exposed metepisternum is in truth narrow. It is very narrow cephalad, gradually broadens caudad and at its broadest point is only about as broad as the lateral elytral interval. The intercoxal piece of the first abdominal ventrite is gently arcuate; the combined medial lengths of ventrites one and two is greater than the length of the remainder of the venter (30:25), and ventrites two plus three are shorter than five (about 10:15) which is more than twice as broad as long (35:15 on the female measured ). It is confusingly stated in the generic description that the stout tibiae are “unarmed at the tips, angulate at the outer apical angle, rounded within,” but it is said in the description of the species: “Inter- mediate and posterior tibiae each with two minute teeth at the outer apical angle.” It is obvious that the tibial armature has been mis- interpreted. The tibiae are short and stout and are adapted to the mode of life of the weevil on the host palm. The protibiae are quite cossonid-like, the uncus is distinct and strong, and there are two teeth on the outer tibial edge, one of which is evidently the mucro and the other a pre-mucral denticle on the angulation at the side of the tarsal articulating groove. The meso- and metatibiae each have four teeth on the anterior face of the apex, the bottom one of which appears to be the uncus, and what appears to be the mucro is present as another tooth at the lower apex of the tibia on the side opposite to the uncus. The three more dorsal teeth or spurs (see fig. 13) are unusual and recall similar spurs in the Scolytinae and as recently described by Kuschel for his group Araucariini [New Zealand Journal of Science 9(1):4, 1966]. It is possible that Champion did not have adequate optical equipment and hence could not see these small details. As noted by Champion, there are few external differences between the sexes of this weevil, and it is difficult to distinguish them. The kind cooperation of the British Museum (Natural History) made possible the completion of this report. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 33 CHRYBANEURA HARRISONI, A NEW GENUS AND SPECIES FROM CENTRAL AMERICA (DiprerA: CECIDOMYIIDAE ) RAYMOND J. GacNnr, Systematic Entomology Laboratory, U.S. Department of Agriculture, Washington, D.C. 20560 Specimens of this cecidomyiid were found among empty spider egg cases in Costa Rica and Panama by Dr. James O. Harrison of Mercer University, Macon, Georgia. The midges do not appear to be parasites of the spider eggs. The midges represent a new genus which belongs to the tribe Brachyneurini of the supertribe Oligot- rophidi and resembles closely the genus Brachyneura Rondani in the form of the male and female genitalia and in the wing venation. The antennae, however, differ greatly from those of Brachyneura and resemble more closely those of typical Oligotrophidi. In Chrybaneura, the antennal flagellomeres have two longitudinal circumfila, many closely set setae ventrally and a few dorsally, and no scales; in Brachy- neura, the flagellomeres have several very fine, wavy, longitudinal circumfila, do not possess setae, and have many long, narrow scales arranged in longitudinal rows. Chrybaneura, n. gen. Head with compound eyes broadly joined on vertex. Frontoclypeus and area behind eyes densely covered with scales. Antenna: scape short, narrowest at base; pedicel globular, as long as scape, both segments together subequal in length to first flagellomere; 10 flagellomeres, each with a long cylindrical node densely covered with minute setulae and a short, naked distal neck, the latter less than \% length of node, and shorter in the female; setae sparse dorsally, numerous and closely set ventrally; each node encircled by 2 transverse circumfila connected by 2 longitudinal strands. Palpus 3-segmented, each segment progressively longer than the preceding, cylindrical, longer than wide, tapering at ends, covered with scales, 4-5 setae present near apex of third segment, setae sparse elsewhere. Thorax: 2 dorsocentral rows of scales and setae on the mesoscutum, very wide at anterolateral angles, narrowing posteriorly, ending slightly anterior to the scutellum; 2 lateral scutal rows of setae beginning behind prescutal suture and ending above the wing bases; setae present on disc of scutellum and on anepim- eron. Wing (fig. 1): costa interrupted at juncture with R; proximal to wing apex; Ri short, less than 1% wing length; R; arched forward approximately 1% distance from base, a sensorium present distal to arch; Cu simple, not extending to edge of wing; entire wing covered with scales which are denser on veins than on membrane. Legs: 5 tarsomeres, first shorter than second; covered with scales; tarsal claws strongly curved, with well-developed teeth; empodium and _ pulvilli short. Male postabdomen: tenth tergum broad, bilobed, the lobes roughly trian- gular; tenth sternum about as long, but narrower than the tergum, bilobed, the lobes narrow and rounded at apex; basimere short, stout, with many setae; 34 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 1-6, Chrybaneura harrisoni, n. sp.: 1, wing (40X); 2, fifth flagellomere, é, dorsal view (225X); 3, same, ventral view (225X); 4, fifth flagellomere, 9, dorsal view (225X); 5, postabdomen, ¢, dorsal view (225X); 6, postabdomen, 9, lateral view (250X). claspettes absent; distimere short, constricted beyond the middle, and toothed at apex. Female genitalia: non-protrusible; 2 dorsolateral cerci present; a long median narrow lamella present below cerci; eighth sternum divided into 2 roughly triangular posteriorly projecting ventrolateral lobes. Type-species, Chrybaneura harrisoni n. sp. Chrybaneura harrisoni, n. sp. Antenna: length third male flagellomere (figs. 2-3), 0.06 mm, the stem 4 length of node; length third female flagellomere (fig. 4),°0.05 mm, the stem %& length of node; 2 transverse circumfila near ends of each flagellomere, arching toward middle to meet the 2 longitudinal connectives. Length of palpal segments I-III (in mm): I, 0.012-0.013; II, 0.019-0.021; III, 0.029-0.032. Chaetotaxy: dorsocentral setae, 23-27; scutal, 12-17; scutellar, 8-10; anepimeral, 7-9. Wing length, 0.53-0.83 mm. Length femur: tibia: tarsus approximately 9:1:1. Percent length of each tarsomere to total length of tarsus, from I to V: I, 14; II, 31; II, 20; IV, 17; V, 18. Male postabdomen (fig. 5) tenth tergum wide, broadly and triangularly emarginate, with several setae along lateral edges; tenth sternum as long as tergum, bilobed with emargination not as deep as that of tergum, and rounded, the resulting lobes narrow, each with a seta at apex; aedeagus narrow, as long as tenth segment; basimere stout, unlobed, with several setae along length; distimere broad at base, narrowing at middle, ending in a sclerotized tooth. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 35 Female postabdomen (fig. 6): cerci 0.07—0.08 mm in length, each with several setae; lower lamella, long, narrow, 0.035 mm in length; triangular lobes of eighth sternum prolonged posteriorly almost as far as cerci. Immature stages unknown. Material examined: Holotype (on slide): ¢, Panama: Changuinola, No. 358, VII-15-1965, from old egg shells of parasitized spider eggs [parasitized by Hymenoptera], coll. James O. Harrison, USNM 69510. Paratypes: 10 ¢ 4, 9 22, same data as holotype; 4 ¢6, 7 2°, Costa Rica : Ciudad Quesada, No. 385, VII-15-1966, from spider egg cases, coll. James O. Harrison. All types deposited in U.S. National Museum. A NEW GENUS AND SPECIES OF DELTOCEPHALINAE FROM PUERTO RICO (HoMoprTERA: CICADELLIDAE ) J. Matponapo Caprites, Department of Biology, Cayey Regional College, University of Puerto Rico, Cayey, Puerto Rico 00633 A new genus and species of leafhopper from the mountains of Puerto Rico is herein described. All efforts have failed to collect this species by sweeping from the ground or low level vegetation around the light traps in which all specimens have been caught. This seems to be a tree-dwelling species. I have great pleasure in dedicating this genus to my friend, Dr. James P. Kramer, of the U.S. Department of Agriculture, Washington, D. C., for his many contributions to the knowledge of the Cicadellidae. The types are deposited in the U.S. National Museum and in my collection. In the measurements that follow, 10 micrometer units correspond to 0.075 mm. To establish the systematic position of this new genus I have followed R. Linnavuoris “Revision of the Neo- tropical Deltocephalinae and some related Subfamilies,” Annales Zoologici Societatis Zoologicae Botanicae Fennicae “Vanamo,” Vol. 20, No. 1, 1959. Krameraxus, n. gen. Cicadellidae, Deltocephalinae, Euscelini. Relatively robust, not flattened body, about 5.9 mm long. Ocelli on anterior margin of crown contiguous to eyes. Episternum concealed. Dorsum without rough circular pits. Face flat, not hairy; frontoclypeus broadening upwards. Gena not visible from above. Crown very slightly longer medianly than next to eyes, nearly flat, smooth, rounded to face at anterior margin. Pronotum inconspicuously finely transversely rugose; narrower than head across eyes. Scutellum smooth, about 1.4 times as broad as long, apical half raised. Forewings well developed, with numerous false veins or broken pigmented lines, small irregular brownish and subcircular or subsquare whitish 36 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 1-10, Krameraxus leucornatus, n. gen. and n. sp.: 1, habitus; 2, head and part of thorax in lateral view, ¢; 3, face, ¢; 4, aedeagus in lateral view; 5, aedeagus in ventro-caudal view; 6, valve and ¢ plates; 7, style and connective in dorsal view; 8, connective in caudal view; 9, ¢ genital capsule in lateral view; 10, 2 seventh sternum and adjacent segments. or grayish areas; 4 apical cells, the fifth missing; apparently 3 subapicals, the crossvein closing the inner missing or perhaps obscured by the false veins. Face, crown, pronotum, and scutellum mostly yellowish-white or yellowish with a grayish sordid appearance and ornamented with pale or dark brown. Male genitalia: pygofer with membranous lateral fold and macrosetae. Plates triangular and with macrosetae. Style with a short curved apophysis and broad plate-like basal part. Connective U-shaped, probably a modification of the Y-shaped type (fig. 7), flattened. Valve and genital plate articulated, not fused. Aedeagus symmetrical, horizontal, spined apically, arising from a disc-shaped unsclerotized structure, a strong spine pointing caudad on middle of lower edge of disc-shaped structure. Type of genus: Krameraxus leucornatus, n. sp. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 37 Krameraxus leucornatus, n. sp. (Figs. 1-10) Characterized by its medium size, wedge-shaped body, and highly contrasting anterior yellowish parts with the brown forewings. Female—face, crown, pronotum, and scutellum mostly yellowish-white; eyes brownish or yellowish, crown with 2 brown anterior spots and two posterior dashes near median line; pronotum with a brown spot on disc in line with each inner margin of eye, pale brown irregular areas near anterior margin as illustrated; scutellum with a triangular brown area near each anterior angle, a small semi- circular brown area on each side near apex, 2 brown spots on disc anterior to brown transverse suture, 2 very small brownish areas anteriorly near median line (fig. 1). A dark spot below each ocellus. Anteclypeus yellowish-white on basal half. Apical half of anteclypeus, lorae, and genae heavily mottled with brown. Forewing mostly polished brown, with many darker spots, brown transverse false veins and ramose lines, with many subcircular or subsquare grayish areas; costal area yellowish and spotted with brown on middle third, first and last third brownish with lighter and darker brown spots, a hyaline area above the yellowish section of costa; inner margin of and a large round area near apex of clavus whitish. Pro- and mesofemur brown with yellowish irregular annuli; metafemur and tibiae stramineous with brown spots, especially along strongly spined margins; spines stramineous. Abdominal sterna grayish with pale brown margins. Last abdominal segment on middle of hind margin brown; pygofer mostly brown, yellowish near base of ovipositor; ovipositor brown at midlength, light colored basally and apically. Crown slightly longer near middle than near eye (3.7:3.2), slightly less than 4 times as wide as long near eyes (12:3.2). Head wider across eyes than pronotum (25:22). Pronotum twice as wide as long (22:11). Scutellum about 1.5 times as wide as long (17:11). Last abdominal sternum as in fig. 10; pygofers with macrosetae. Length 5.9 mm. Male—general coloration much as in female. Yellow of head and thorax with a brownish tinge, not so bright as in female; genal area lighter, with very scarce pale brown mottling; large whitish spot near apex of clavus present or with a squarish gray area in its place. Costal margin without yellowish middle area. Crown slightly longer at middle than near eye (3.1:3.0), slightly less than 4 times as wide as long near eye (11:3.0). Head wider across eyes than pronotum (23:20). Pronotum twice as wide as long (20:10). Scutellum about 1.3 times as wide as long (16:13). Male genitalia as illustrated. Length 5.2 mm. This genus runs to the Euscelini in Linnavuori’s key to the tribes of the Deltocephalinae. In his key to the genera of Euscelini, p. 153, it runs to couplet 11 if the connective is considered as of the Y-type and whether considered to have two or three subapicals. It differs strikingly from the keyed genera. Holotype, male, from El Verde Experimental Forest Station, near El Yunque, Puerto Rico, June 1967, in a light trap, collected by the author, USNM 69658. Allotype, female, same data, in the USNM. Paratypes, four: two same data as holotype, and two from Maricao State Forest, taken in a light trap operated in the Fish Hatchery, in the author's collection. 38 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 THE GENUS PASSALOBIA LOMBARDINI, 1926, WITH DESCRIPTION OF A NEW SPECIES (ACARINA;: DIARTHROPHALLIDAE )! Preston E. Hunter and SANDRA GLOovER,? Department of Entomology, University of Georgia, Athens, Georgia 30601 The genus Passalobia was erected by Lombardini (1926) for a new species, Passalobia quadricaudata, taken from under the elytra of a passalid beetle from Brazil. Lombardini (1938a, 1938b, 1951) later added the following species, all based on nymphs, to this genus: duodecimpilosa, 1938, major, 1938, and peritrematica, 1951. Womersley (1961) moved duodecimpilosa to Diarthrophallus Tragardh and erected a new genus, Passalana, for peritrematica leaving only two species, quadricaudata and major—both from passalid beetles from Brazil—in Passalobia. In this paper we are describing a new species of Passalobia taken from a small number of alcohol preserved passalid beetles from Costa Rica. Passalobia Lombardini, 1926 Lombardini, G. 1926. Boll. Soc. Entom. Ital. 63: 158, figs. 1-2. Womersley, H. 1961. Trans. Roy. Soc. S. Australia 84: 35. Womersley (1961) redescribed this genus and included a key to the species. The genus is distinct in having the tectum helmetlike with an apical spike, dorsum with one (subposterior) pair of long simple setae and the elongate body slightly constricted behind coxae IV. Passalobia dubinerae, n. sp. (Fig. 1A-K) The female is distinct from quadricaudata, the only known female, in that the perigenital ring is triangular in shape and completely posterior to coxae IV whereas in quadricaudata the perigenital ring is oval and extends to the middle of coxae II. The separation of these adults from major must await description of the adult of major. Although the nymphal stage described for major and quadricaudata was not indicated, it appears to be the deutonymph for both. If so, the following combination of characters are distinctive for the deutonymph of the new species: Idiosoma approximately three times as long as area from coxae I to IV; para-anal setae shorter than posterior dorsal setae; dorsal setae arising from the integument posterior to the dorsal plate. FEMALE. Idiosoma elongate, 626 » long, constricted behind legs IV, 193 u at greatest width anterior to coxae IV, 101 yu at narrowest part of constriction, 123 w at greatest width posterior to constriction (measurements are the average 1 University of Georgia College of Agriculture Experiment Station, Journal Series Paper No. 110, College Station, Athens. Partially supported by the National Science Foundation, Grant No. GB-3414. 2 Associate Professor and Graduate Student, respectively. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 39 i Y { eet. a \ eM Co) =a * | | { ~n fit ahhs) NLA Fig. 1. Passalobia dubinerae n. sp. Female: A, dorsum; B, venter; C, ventral view of caruncle III; D, dorsal view of tarsus I; E, tectum; F, ventral view of gnathosoma; G, chelicera. Deutonymph: H, dorsum; I, venter. Protonymph: J, dorsum; K, venter. of 10 specimens). Dorsum (fig. 1A). A single plate covering most of dorsum, plate constricted behind legs IV, posterior to constriction widens slightly; integu- ment visible around all margins of plate; 1 pair of long simple forwardly curving setae arising from posterior margin of plate, other setae on plate consisting of 9 pairs of very minute simple setae (setae smaller in relation to body size than shown in fig. 1A) in area above coxae; 1 pair of pores near posterior margin of plate; plate without reticulations. Venter (fig. 1B). Coxae I forming a V medially, the anterior margins not touching; 1 pair of pores in integument anterior to base of coxae. Sternal, endopodal, metasternal and ventral plates coalesced, 550 yu long, 88 » wide at level of coxa IV, expanding behind genital opening to 174 yu wide; extending posteriorly to and in contact with anal plate; coalesced plate bearing 3 pairs of pores and 5 pairs of small simple setae; location of pores—2 pairs just posterior to genital opening, 1 pair at anterior level of genital opening, 1 pair between coxae II and III, 1 pair at posterior margin of coxae IV, 1 pair midway between genital and anal opening and 1 pair at posterior margin of genital opening. Anal plate small, triangular shaped; separated from genital plate by a suture; anal setae simple, para-anals long, capitate at tip, post-anal no longer than length of anal opening. Genital opening surrounded laterally and anteriorly by perigenital ring; anterior margin of ring well posterior to coxae IV; genital plate tongue-shaped, bearing striaelike marks medially. Integument visible lateral 40 PROC, ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Table 1. Leg chaetotaxic formulae of deutonymph-female of Passalobia dubinerae n.sp. Total number of setae only given for tarsi. Segment Leg Coxa ‘Trochanter Femur Genu Tibia Tarsus I 04 ri coat fs ee 5 2 00 ll 00 11 0 00 me. VEAL Ol II 0—0 alt loa | eal! lek VI 1 ll Wal 00 Mal 0 10 gl ll 01 Ill 0—0 iil == I Kee IL ll 1 10 ll 00 ll IV oo a 0 it 1 Es 1? 4 1 Y 1 ba 0) 10 Wl 00 ll Prot ] ga 211%) 02h ee rotonymph: 10” 10”: 10 2: 00 2" of plate behind coxae IV and bearing 1 pair of simple, short setae near posterior of body. Peritreme dorsolaterally, at level of coxa III, approximately half width of coxa in length, no peritremal plate seen. Tritosternum bearing 2 spined lacinae; a pair of simple setae flanking base. Legs. Setae simple, all short except for a pair of longer setae on tarsi II-IV and a long whiplike seta on dorsum of genu I and tip of tarsi I. Chaetotaxy formulae given in Table 1. Pretarsi I-IV each bearing a large padlike caruncle and a pair of sclerotized T-shaped claws (fig. 1C), each claw attached to the caruncle by the middle bar of the T with the cross bar, which runs parallel to the caruncle, terminating at each end in a sharply pointed claw; distal to the attachment of each claw the caruncle bears a thickened padlike, claw-shaped structure. Posterodorsally tarsus I bears a subterminal 3-tined clawlike structure (fig. 1D). Gnathosoma (fig. 1E-G). Tectum (fig. LE) with a ventral groove—appearing as a longitudinal strip; margin smooth. Internal mali long, extending to middle of tibia, bearing centrally a thickened supporting structure; lateral and medial of supporting structure mali is membranous, lateral margin scalloped, medial margin bearing 2 spinelike projections. Venter of gnathosoma bearing only 3 pairs of setae, relative lengths as shown, internal posterior rostral setae absent; deutosternal groove present but without teeth. Palpal setae simple; all short except for 2 longer setae on distal margin of tibia and 1 on tarsus; tarsus bearing a thickened spinelike terminal setae. Chelicerae chelate, well sclerotized; movable digit with 1 strong tooth, fixed digit with 2 teeth (fig. 1G). DEUTONYMPH. Idiosoma 540 u« long, 115 « wide (average of 2 specimens ); constricted behind coxae IV. Dorsum (fig. 1H). Single dorsal plate 457 » long, following general contour of body, constricting to 48 « and expanding posterior of constriction to 90 «; not extending to margins of idiosoma nor to posterior margin of body; 4—5 pairs of minute setae (relative size smaller than illustrated ) above coxal region; integument bearing 1 pair of long simple setae posterior of dorsal plate. Stigmata dorsolateral above coxae III, visible dorsally. Venter (fig. II). Coxae I meeting medially. Single elongate plate 403 « long, 48 « at greatest width behind coxae IV, extending posteriorly almost to anal plate; bearing 4 pairs of setae and 2 pairs of pores positioned as shown; 2 pairs of setae—1 pair anterior PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 4] and 1 posterior—off plate; relative lengths of setae as shown; 1 pair large pores in integument medial of coxae II. Anal plate rounded; anal setae simple, para- anals long, length of post-anal about equal to length of anal opening. Trito- sternum as in female. Legs. Chaetotaxy as in female. Length of legs I-IV: II, 195 pw; Ill, 183 ww; IV, 185 uw. Gnathosoma of same general characteristics as described for female. PROTONYMPH. Idiosoma 453 y» long (average of 5 specimens); weakly constricted behind coxae IV. Dorsum (fig. 1J). Dorsal podosomal plate only; plate 168 » long, 93 w wide; shape as shown; integument visible around margin of plate; bearing 4 pairs of minute setae (relative size of setae much smaller in relation to body than shown). Posteriorly idiosoma bearing 1 pair of long simple setae. A pair of peritremes (?) arising from the dorsum near the posterior margin of the dorsal plate (these structures are free of the body and appear to connect to the stigmata which are dorsolateral in position; similar structures were not seen in the deutonymph or female.) Venter (fig. 1K). Plate 133 uw long, 35 yw at greatest width; bearing 2 pairs small simple setae positioned as shown. Integu- ment bearing 1 pair small setae just posterior to coxae I and a slightly larger pair near posterior of body; 2 pairs of pores—1l pair anterior to coxae I, second pair at level of coxae II. Anal plate not well sclerotized; bearing 3 simple setae, para-anals long; length of post-anal seta about equal to length of anal opening. A pair of simple setae at base of tritosternum; lacinae as in female. Legs. Chaeto- taxic deviations from adult discussed below; lengths of legs II-IV as follows: II, 180 y»; III, 167 w; IV, 155 uw. Gnathosoma. Hypostomal process, corniculi and ventral hypostomal setae of general facies of female. Male unknown. LEG CHAETOTAXY. The chaetotaxic formula following Evans (1963) is given in Table 1 for the deutonymph-female. Of particular interest, compared to other diarthrophallids and mesostigmatic mites, is the loss of the posterior seta on coxae II, III, and IV. The protonymphal leg chaetotaxy is the same as that of the deutonymph-female except for the absence of the following setae: trochanter III and IV—pl of each segment missing; femur IIJ—pv,: missing; femur IV—avi, ad: and pv: missing. Described from a series of 10 females, 2 deutonymphs and 5 pro- tonymphs. Holotype (female) data: San Vito, Costa Rica; 11-IJ- 1965; M. V. Truitt, coll.; from passalid beetle, genus Publius. All females, deutonymphs and 3 protonymphs with same data. Two protonymphs with same data except collected from passalid beetle of genus Passalus. Holotype, three female paratypes, one deutonymph and 2 protonymphs deposited in U.S. National Museum, Washington, D. C. Remaining material in collection of Department of Entomology, University of Georgia, Athens, Georgia. Remarks. The free peritremelike structure of the protonymph is unusual. Free peritremes occur in Passalana peritrematica (Lamb. ) which is known only from the nymphal stage. Neither Lombardini nor Womersley indicated which nymphal stage was represented. From the ventral chaetotaxy it appears likely the specimen was a_ pro- tonymph. The description of additional species of Passalobia would 42, PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 answer the question of whether all protonymphs of this genus have similar peritremelike structures and, if so, should Passalana retain separate generic standing. ACKNOWLEDGMENTS We would like to express our appreciation to Mrs. Virginia Truitt Mullen, Department of Entomology, University of Georgia, for the passalid beetles from which these mites were taken and to Dr. O. L. Cartwright, U. S. National Museum, Washington, D.C., for determinations of the beetles. REFERENCES Lombardini, G. 1926. Duo nova genera acarorum. Boll. Soc. Ent. Ital. 58: 158-161. 1938a. Acari novi. Mem. Soc. Ent. Ital. 17:44—46. - 1938b. Acari novi II. Mem. Soc. Ent. Ital. 17:118—120. - 1951. Acari novi. Redia 36:245-250. Womersley, H. 1961. On the family Diarthrophallidae (Acarina Mesostigma- Monogynaspida) with particular reference to the genus Passalobia Lombardini 1926. Trans. Roy. Soc. S. Australia 84:27-44. NEW GENERA AND SPECIES OF NEOTROPICAL BLISSINAE (HemMipTERA: LyYGAEDAE)? James A. SLATER and DaRLEEN B. Witcox, Systematics and Environmental Biology Section, University of Connecticut, Storrs, Connecticut 06268 During the course of recent revisional work on the Chinch Bug subfamily Blissinae we have described and treated three new genera, Patritiodemus (Slater and Ahmad, 1968), Praetorblissus (Slater, 1966 ) and Reticulatodemus (Slater and Wilcox, 1966). In the present paper we describe an additional new species in each of these genera from specimens which have become available since the original publication of the new genera, describe an additional South American genus Caveloblissus, and comment upon the hitherto unknown macropterous form of Heteroblissus anomilis Barber. Caveloblissus, n. gen. Body subelongate, robust, nearly parallel sided; head and anterior pronotal lobe above strongly shining, posterior pronotal lobe from anterior margin of transverse impression to and including entire humeral area and scutellum dull pruinose, dividing line on pronotum between anterior shining and posterior pruinose areas 1 This work was supported by a grant in aid from the National Science Foun- dation. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 43 Ss as Figs. 1-3, Caveloblissus americanus, n. sp.: 1, clasper; 2 and 3, sperm reservoir. sharp and straight transversely across pronotum; scutellum lacking shining areas even on median elevation; antennae relatively elongate with segments 2 and 3 almost completely terete, 2nd segment slightly enlarged at distal end, 4th segment strongly fusiform; hemelytra with apical corial margin concave basally, lateral corial margins conspicuously sinuate, membrane subhyaline, differing markedly in thickness and texture from clavus and corium; all femora moderately incrassate, mutic; fore coxal cavities open; head and thorax completely pruinose below; mesosternum bearing a deep median longitudinal furrow; metathoracic scent gland orifice small, rounded, auricular; fore tibiae linear, lacking a series of teeth, and completely non-fossorial, Ist and 2nd tarsal segments of nearly equal length, basal segment not enlarged and broadened; antenniferous tubercles bluntly trun- cate; claspers (fig. 1) short and stout, inner lobe set relatively far from base, sperm reservoir (figs. 2, 3) consisting of a central sclerotized rod and a pair of slender, elongate lobed lateral “wings.” Type-species: Caveloblissus americanus, n. sp. This new genus is closely related to Neoblissus Bergroth and to the other Western Hemisphere species such as leucopterus (Say) and its allies, currently placed in the genus Blissus Burm. The genus is sepa- rable, however, by virtue of the shining head and anterior pronotal lobe. In Neoblissus and Western Hemisphere “Blissus” species the head and entire pronotum are completely pruinose. It is interesting that this sharp division of the pronotum into shining and pruinose areas parallels that of Cavelerius Distant (Oriental) and Praeblissus Barber (W. Hemisphere). This relationship, however, seems to us to be the result of independent specialization from a more generalized td PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Fig. 4, Caveloblissus americanus, n. sp., dorsal view. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 45 ancestor and not to indicate direct phylogenetic derivation of the three taxa. The evolutionary relationships of this genus will be more fully discussed in a subsequent paper on generic relationships in the subfamily. Caveloblissus americanus, n. sp. (Fig. 4) Head, pronotum and scutellum black, pronotum becoming bright reddish brown across humeral area, tylus dark red-brown on anterior half; hemelytra sordid white, membrane subhyaline, clavus dark red-brown on basal third, corium with a very large dark red-brown apical patch, this dark coloration extending along entire apical corial margin and midway to base of cubital vein, an additional rectangular brown patch at inner angle of apical corial margin extending 4% way from apical margin inward, in contact both with cubital vein and claval suture and strongly contrasting with the pale flavescent corium adjacent; abdomen uniformly bright red-brown; all femora bright castaneous, tibiae and tarsi flaves- cent with a basal and apical brown band around tibiae, somewhat suffused with brown on fore tibiae; antennae reddish brown with proximal 24 of 2nd segment a light tan, apex of 3rd tarsal segment dark brown, strongly contrasting with flavescent color of preceding tarsal segments; head somewhat rugulose; pronotum and scutellum minutely indistinctly punctate; clothed above and below with a rather thick covering of decumbent sericeous hairs. Head non-declivent, conspicuously convex across vertex, tylus subacuminate, extending midway to distal end of Ist antennal segment, eyes somewhat produced laterally, set very slightly away from antero-lateral pronotal angles, length head 46 mm, width head .67 mm, interocular space .40 mm; pronotum moderately convex, lateral margins nearly straight from base to central area of calli, then strongly mesally arcuately curved to anterior margin, posterior margin nearly straight across base of scutellum with very slightly produced posteriorly directed lobes laterad of scutellar base, transverse impression complete, shallow, width across humeri equal to width across calli, length pronotum .87 mm, width pronotum 1.22 mm; scutellum with a low inconspicuous median elevation on distal %, length scutellum .40 mm, width scutellum .54 mm; hemelytra with lateral corial margins sinuately concave, membrane extending midway to posterior margin of 7th abdominal tergum, leaving abdominal connexiva exposed through- out; distance apex clavyus—apex corium .76 mm, distance apex corium—apex abdomen 1.63 mm; labium relatively elongate, extending well onto mesosternum, distal end of 1st segment approaching or attaining base of head, 2nd segment attaining or nearly attaining base of prosternum, length labial segments (male paratype) I .33 mm, II .40 mm, III .36 mm, IV .53 mm, length antennal segments I .15 mm, II .40 mn, III .34 mm, IV .55 mm; total length 4.68 mm. Holotype: Male. BRAZIL: Porto Alegre 25-VII-945 “on Bambusa sp.” (R. Gomex Costa). In La Plata Museum. Paratypes: BRAZIL: 1 female, same data as holotype; 1 male, 1 female, Sao Leopoldo (J. W. Stahl); 1 male Corumba. PARAGUAY: 1 female, Asuncion 4.X.16 (Vezenyi). In Hungarian National Museum, Stockholm Museum, U.S. National Museum (Drake coll.), and J. A. Slater collections. 46 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 23 A J Fig. 5, Praetorblissus gradus, n. sp., dorsal view. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 AT Praetorblissus gradus, n. sp. GEiew 5) General coloration a nearly uniform dark castaneous red-brown becoming fuscous on antennae, scutellum, anterior pronotal lobe in region of calli and exposed metanotum; legs and connexivum a contrastingly pale yellowish ochra- ceous; body in large part glabrous or with only a few scattered hairs, more numerous along lateral margins of pronotum; head and anterior pronotal lobe smooth and impunctate, or nearly so, extreme anterior area of pronotum with numerous large, coarse punctures placed in a collar-like position, posterior pronotal lobe coarsely and deeply punctate, scutellum and abdominal terga with numerous small scattered punctures. Head non-declivent, shallowly and irregularly rugulose, moderately convex, eyes sessile or nearly so, set very slightly away from antero-lateral pronotal angles, tylus extending to distal 4% of Ist antennal segment, length head .62 mm, width head .91 mm, interocular space .61 mm; pronotum prominently arcuate, the lateral margins nearly straight or even slightly expanded from humeral angles to area of calli, then sharply and convexly narrowing to anterior margin, calli strongly convex to form a depressed median furrow on anterior lobe, posterior margin nearly straight, at most very slightly concave, transverse impression obsolete, posterior lobe only % or slightly more than % length of anterior lobe, length pronotum 1.10 mm, width pronotum 1.67 mm; scutellum elevated and convex on basal half, with a low broad median elevation distally, length scutellum .61 mm, width scutellum 1.18 mm; hemelytra reduced to extremely tiny undif- ferentiated pads, extending along lateral margin only midway to posterior margin of metanotum, median angle reaching only midway to apex of scutellum, apical margin strongly concave, length wing pad .99 mm; metanotum completely exposed, coarsely and deeply transversely grooved; abdomen elongately ovoid, suture between abdominal terga 5 and 6 deeply curved caudad mesally, length abdomen 3.45 mm; metathoracic scent gland orifice broad, rounded distally, projected anteriorly as a broad finger-like process (fig. 6); fore femora strongly incrassate, armed below on distal third of each ventral surface with a large elongate sharp spine, additional short inconspicuous spines present distally, middle and hind femora mutic; labium slightly exceeding fore coxae, length labial segments I 42 mm, II .46 mm, III .42 mm, IV .34 mm. (approx.); antennae slender, terete, 4th segment very narrowly fusiform, length antennal segments I .23 mm, II .65 mm, III .65 mm, IV .84 mm; total length 4.68 mm. Holotype: Male. PERU: Utcuyacu and Agua Dulce, Prov. of Tarma, Dept. of Junin IIJ-1948 (F. Woytkowski) (donor Wm. Procter). In American Museum of Natural History. Paratypes: 2 males, same data as holotype. 1 male, BOLIVIA: Cochabamba (Germain) (coll. Noualhier 1898). In Paris Museum, American Museum of Natural History and J. A. Slater collections. This is the second species known in the genus Praetorblissus and it differs strikingly from the type species, gigas Slater. P. gradus is only about half the size of gigas and importantly it lacks spines on the middle and hind femora. This character is important in the phylogeny of the Blissinae and is often used to establish generic entities. How- 48 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 ever, in the present case it appears undesirable to place this new species in a distinct genus since it is so obviously closely related to the type species of Praetorblissus and the loss of spines on the middle and hind femora represents an evolutionary advance in the phylogeny of the genus which would be masked were we to not recognize the close relationship of these two species in every other respect. The condition of the head, pronotum, type of extreme wing brachyptery, shape of scent gland orifice (fig. 6), body texture, open coxal cavities, and even spines on the fore femur indicate the very close relationship between gigas and gradus, and there is no doubt that the two are very closely related to one another, gradus being the more recently evolved of the two. Slater (1966) has previously remarked on the possible relationships between Praetorblissus and Heteroblissus Barber. This new species makes this relationship even more compelling since gradus is about the size of Heteroblissus anomilis Barb., and both lack spines on the middle and hind femora. Nevertheless, gradus is not at all intermediate between gigas and Heteroblissus anomilis, being a typical Praetorblissus in all respects except the condition of the femoral spines mentioned above. Heteroblissus anomilis Barber Heteroblissus anomilis Barber, 1954, Rev. Brasil Biol. 14(2):221-3. Heteroblissus anomilis: Ashlock and Lattin, 1963, Ann. Ent. Soc. Amer. 56: 694-703. Heteroblissus anomilis: Slater, 1966, Univ. Conn. Occ. Pap., Biol. Sci. Ser. 1(1): 3-11. This species, the only representative of the genus, has previously been known only from extremely micropterous specimens. Since the nature of the front wings is important in establishing generic relation- ships it is pleasing to report the presence of two fully macropterous females in the Leningrad Museum from Misiones, Loreto, Argentina. The hemelytron of these macropters has the clavus and corium heavily sclerotized, strongly contrasting with the thin semi-hyaline membrane, the apical corial margin concave basally, but straight for a considerable portion of its length. The clavus bears a row of punc- tures along its inner margin and on either side of a raised calloused line near the claval suture. The corium bears a row of punctures on either side of the cubital vein and mesad of the strongly raised radial vein. The corium on the lateral half is of a contrasting thickness and composition to the inner half. The entire dorsal as well as the ventral surface is subshining, completely devoid of pruinose areas. The membrane is broadly rounded and extends distad almost to the posterior margin of the sixth abdominal tergum. The genus is an interesting one in that it possesses what appears to be a stridulatory area on the abdominal sterna (see Ashlock and Lattin, ibid.). PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 49 Fig. 6, Praetorblissus gradus, n. sp., scent gland orifice. As is so frequently the case with species exhibiting wing poly- morphism, the pronotal shape of the macropterous form is quite unlike that of the brachypterous specimens. In the former the pronotum is more elongate, broader across the humeri than across the calli and with at most a lateral trace of an obsolete transverse impression. H. anomilis was originally described from southern Brazil. In addition to the Loreto locality mentioned above we have examined material from San Antonio in Misiones, Argentina. Reticulatodemus orbiculoides, n. sp. Head, anterior pronotal lobe and scutellum black with apex of tylus, scutellar carina and posterior pronotal lobe bright red-brown to castaneous; clavus and corium testaceous, suffused with dusky coloration on apical 4% of corium, mem- brane broadly sordid white-margined with brown veins and a large elongate brown central macula covering greater portion of membrane disc, but not extend- ing to lateral or apical margins; abdomen uniformly red-brown; all femora bright reddish brown with tibiae and tarsi flavescent, antennal segments 1 and 2 red brown (3 and 4 missing); central area of head, large triangular and mesally contiguous patches on calli, posterior 4% of pronotum, and scutellar carina shining, remainder of dorsal surface and entire thoracic venter a strongly contrasting pruinose gray; pronotum punctate, posterior pronotal band smooth and glabrous; otherwise sparsely clothed with short, decumbent somewhat scale-like sericeous hairs. Head slightly convex across vertex, non-declivent, tylus prominent, consider- ably exceeding juga but short, extending half way to distal end of Ist antennal segment, eyes flattened against head surface to form nearly evenly curving lateral 50 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 margins, set well away from antero-lateral pronotal angles, antenniferous tubercles strongly hooked, length head .61 mm, width head .80 mm, interocular space .53 mm; pronotum moderately convex, area across humeri slightly more swollen than anterior lobe, lateral margins sinuately narrowing from humeral angles to anterior margin, transverse impression very shallow, broad, complete, posterior margin slightly and evenly concave, a weak, broad punctate median groove between calli on anterior lobe, length pronotum 1.06 mm, width pronotum 1.25 mm; scutellum with a prominent shining median carina, length scutellum .53 mm, width scutellum .58 mm; hemelytra with lateral corial margins nearly parallel, evenly narrowing to expose connexiva, membrane reaching anterior margin of 7th abdominal tergum, distance apex clavus—apex corium 1.14 mm, distance apex corium—abdomen 2.85 mm; labium obscured, apparently extending to fore coxae; fore femora strongly incrassate, ventral surface narrowing to a subacute expanded ridge, armed below on distal 4% with a single large prominent acute spine; antennae stout, 2nd segment terete, length antennal segments I .19 mm, II .49 mm; total length 7.08 mm. Holotype: Female. BRAZIL: Esp. Santo, 98 (Staud). In Hungarian National Museum. This species is very closely related to R. orbiculatus Slater and Wilcox, the two species being unique within the genus in possessing similar “flattened” eyes and an ovoid head shape. However, the head and eyes of orbiculoides are significantly broader than are those of orbiculatus, and this gives a less “hemispherical” bulbous appearance to the head and can be expressed as a ratio of head width to interocular width (1.51 orbiculoides, 1.44 orbiculatus). The two species have pronota almost identical in shape, color and texture. R. orbiculoides has the most minute and obscure wing reticulation of any species in this genus. There are tiny roundish cells visible on the membrane, and only vestiges of the anastomosing lines usually present on the clavus and corium. R. orbiculoides also lacks the black points that are present at the caudo-lateral angles of abdominal connexiva 4 through 7 in orbiculatus, and has the entire 2nd antennal segment dark red-brown rather than strongly darkened on the distal half as in orbiculatus. We have recently examined an additional female of orbiculatus in the Leningrad Museum from Rio Caqueta, Colombia VII-1926 (Woronov ). Patritiodemus delicatus, n. sp. Body moderately elongate, robust, sub-linear; dorsal surface uniformly pruinose, including entire humeral area of pronotum, an irregular “comma-shaped” trans- verse shining band on calli and another anterior to ocelli; head and anterior pronotal lobe dark gray, pronotal collar, posterior pronotal lobe, tylus and scutellum bright reddish tan, abdomen red brown with connexiva a strongly contrasting yellow, hemelytra pale testaceous, veins and apical 4 of corium suffused with darker tan, legs and antennae uniformly very light yellow, 3rd and 4th antennal segments very lightly suffused with light brown, head and pronotum PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 dl finely and evenly punctate; scutellum with irregular relatively coarse punctures laterad of median elevation, scattered punctures present on clavus and corium; sparsely clothed, particularly laterad, with long semi-erect yellowish hairs. Head broad, declivent, moderately convex across vertex, eyes protrudent, set well away from antero-lateral pronotal angles on shelf-like head extensions, juga short, blunt, tylus extending 14 distance to distal end of lst antennal segment, length head .60 mm, width head .91 mm, interocular space .52 mm; pronotum with lateral margins slightly sinuate, narrowing strongly and evenly from humeral angles to anterior margin, posterior margin moderately concave with shallow caudo-lateral lobes produced laterad of scutellum, transverse impression broad, shallow, nearly obsolete, length pronotum .99 mm, width pronotum 1.30 mm; scutellum with a low median elevation on distal 7%, length scutellum .51 mm, width scutellum .61 mm; hemelytra with lateral corial margins slightly sinuate, narrowing opposite claval commissure then abruptly widened posteriorly, mem- brane bluntly and evenly rounded at apex, covering anterior 4% of 7th abdominal tergum, distance apex clavus—apex corium 1.25 mm, distance apex corium— apex abdomen 2.04 mm; fore femora very slender, scarcely incrassate, armed below on distal 4% with a single short sharp spine; metathoracic scent gland orifice short, rounded, auricular, tilted slightly cephalad; labium reaching posterior margin of prosternum, 2nd segment surpassing base of head by % its length, length labial segments I .34 mm, II .30 mm, III .23 mm, IV .27 mm; antennae moderately elongate, slender, segments 2 and 3 terete, 4 narrowly fusiform, length antennal segments I .23 mm, II .69 mm, III .68 mm, IV .87 mm; total length 6.12 mm. Holotype: Female. BRAZIL: Chapada “Acc. No. 2966,” Oct. In Carnegie Museum. Paratypes: 1 female, same data as holotype; 1 female, same except “Nov.” In Carnegie Museum and J. A. Slater collections. In the key to species of Patritiodemus, P. delicatus runs to P. albo- maculatus, but is readily distinguishable from the latter by virtue of the much longer labium which reaches the base of the prosternum in delicatus, whereas it does not even attain the fore coxae in albo- maculatus. P. delicatus is, in fact, unique within the genus Pat- ritiodemus on this character, as all of the other species have the labium very short and at most attaining the fore coxae. The very delicate non-incrassate fore femora, lack of a diffuse central dark area on the hemelytra and generally pale coloration are all distinctive features. Nevertheless, the protrudent nearly “stalked” eyes, presence of a fore femoral spine, and completely pruinose pronotum seem to us to warrant its inclusion in the genus Patritiodemus. ACKNOWLEDGMENTS We wish to extend our sincere appreciation to the following individuals for their kindness in allowing us to study material in their various institutions: Dr. P. Wygodzinsky (American Museum of Natural History), Dr. G. Wallace (Carnegie Museum), Dr. A. Soos (Hungarian National Museum), Dr. B. Torres (La Plata 52 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Museum), Dr. I. M. Kerzhner (Leningrad Museum), Dr. A. Villiers (Paris Museum), Dr. E. Kjellander (Stockholm Museum) and Dr. R. C. Froeschner (U.S. National Museum). Our sincere thanks are due Miss Karen Stoutsenberger, Mrs. Mary Stock and Mr. Abdul Hamid ( University of Connecticut) for preparations of the illustrations. REFERENCES Ashlock, P. D. and J. D. Lattin. 1963. Stridulatory mechanisms in the Lygae- idae, with a new American genus of Orsillinae (Hemiptera: Heteroptera). Ann. Ent. Soc. Amer. 56:693-703. Slater, J. A. 1966. New micropterous Blissinae from South America (Hemip- tera: Lygaeidae). Univ. Conn. Occ. Pap., Biol. Sci. Ser. 1(1):3-11. Slater, J. A. and I. Ahmad. 1968. Neotropical Blissinae: Patritiodemus, a new genus with six new species. Acta Ent. Mus. Nat. Pragae. In press. Slater, J. A. and D. B. Wilcox. 1966. An analysis of three new genera of Neo- tropical Blissinae (Hemiptera: Lygaeidae). Ann. Ent. Soc. Amer. 59(1):61-76. A REDESCRIPTION OF CULEX (ACALLEOMYIA) OBSCURUS (LEICESTER) (DieTERA: CuLIcmAE)! Ratpu A. BRAM,” South East Asia Mosquito Project, Department of Entomology, Smithsonian Institution, Washington, D.C. 20560 Culex (Acalleomyia) obscurus (Leicester) has been poorly charac- terized since its original description. Although Leicester (1908) recognized the highly modified male terminalia as being one of the outstanding distinguishing features of this species, his description was rather vague and was not accompanied by an illustration. Edwards (1922) illustrated the male terminalia, but the specimen which he used was apparently mounted in a somewhat twisted position. As a result, the representation of most of the structures is either misleading, incomplete, or erroneous. No subsequent author has attempted to rectify these descriptions, nor have the immature stages been de- scribed. The following redescription is based on the type series in the British Museum (Natural History); on specimens received from Dr. S. Ramalingam, University of Kuala Lumpur, accessioned to the U.S. National Museum; on material collected by Dr. D. H. Colless on loan from the British Museum; and on specimens from the Institute for Medical Research, Kuala Lumpur, deposited in the U.S. National 1This work was supported by Research Contract No. DA-49-193-MD-2672 from he U.S. Army Research and Development Command, Office of the Surgeon General. 2 Present address: Entomology Research Division, Agricultural Research Service, United States Department of Agriculture, Beltsville, Maryland 20705. PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 53 Museum. Format and terminology conforms to that followed by Bram (1967). Culex (Acalleomyia) Obscurus (Leicester), 1908 (Fig, 1) Acalleomyia obscurus Leicester, 1908, Cul. Malaya: 194 (4, 9). Micraedes obscurus (Leicester): Edwards, 1913, Bull. Ent. Res. 4:237 (tax- onomy ). Culex (Acalleomyia) obscurus (Leicester): Edwards, 1922, Indian J. Med. Res. 10:285 ( ¢*); Macdonald and Traub, 1960, Stud. Inst. Med. Res. Fed. Malaya 29:105 (biology); Macdonald, Smith, and Webb, 1965, J. Med. Ent. 1:338 (distribution); Bram, 1967, Proc. Ent. Soc. Wash. 69:328 (lectotype designa- tion). The adult female is most difficult to recognize, but in well preserved specimens the rather broad, flat scales on the midlobe of the scutellum are diagnostic. The adult male is easily distinguished from other species of the genus by the short palpus and the unique terminalia. The fourth stage larva exhibits an incomplete saddle on abdominal segment X and characteristic comb scales. FEMALE. A rather small species without distinctive ornamentation. Head. Proboscis uniformly dark brown; palpus very short, similar in color to the pro- boscis; length of antenna slightly less than that of the proboscis; vertex with narrow, dark decumbent scales at the occiput, becoming somewhat broader and pale at the orbital line; erect scales of the vertex forked, uniformly brown. Thorax. Integument of the scutum dark brown, clothed with rather dense, bronze- brown scales; acrostichal bristles developed only at the extreme anterior margin; anterior and posterior dorsocentral bristles well developed, as are the other bristle patterns normally found in the genus; midlobe of the scutellum clothed with a rather dense patch of broad, flat scales; integument of the pleuron pale brown (or with a greenish tint in some specimens), without noticeable scale patches; 1 prominent posterior sternopleural and 1 lower mesepimeral bristle present, in addition to the prominent bristles on the prealar area. Wing. Normal for the genus, with the dorsal scales uniformly bronze-brown. Legs. Anterior surface of the hind femur predominantly pale, with a rather broad apical dark band which extends proximally along the dorsal margin; hind tibia and tarsus dark; fore and mid legs dark, without indications of pale patterns. Abdomen. Terga totally dark brown; sterna somewhat lighter, but not pale. MALE. Habitus similar to that of the female. Head. Palpus short, its length approximately 14 that of the proboscis; antenna sparsely plumose. Terminalia. (fig. 1C). Basimere well developed, clothed with both setae and scales (par- ticularly on the proximal half); a prominent basal tubercle present on the mesal margin upon which is inserted a strong, broad, long, and heavily sclerotized, blunt rod and a shorter, slenderer, pointed rod; subapical lobe developed as a strong, conicle tubercle at the extreme apex of the basimere; a strong, long, hooked rod, a shorter, slenderer hooked rod, and a similar but pointed rod inserted at the crown of the tubercle in addition to several accessory setae; a prominent, strongly sclerotized, Y-shaped structure basad of the rods, followed by a broad, truncate leaf and with a prominent, straight seta inserted at the extreme base of the tubercle; distimere expanded subterminally, then narrowing at the apex; with a small recurved spine opposite the distimere claw, and a larger PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 54 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 55 recurved spine on the margin of the expanded area; phallosome similar to species of the subgenus Culiciomyia; lateral plate of the phallosome with a basal tooth and with from 7 to 10 lateral denticles of similar size; proctiger with a tuft of fine, dorsomesal spines and prominent, spatulate, lateroventral spines; from 2 to 4 short, cercal setae present; basal sternal process short and truncate; tergum IX deeply indented, with from 3 to 5 strong setae on each of the 2 lobes. PUPA. Without outstandingly distinctive characteristics; similar to the pupae of species in the subgenus Culiciomyia. LARVA (figs. 1A, B). Head. Antenna normal for the genus, with a narrow, dark basal ring; head hair 1-C filamentous, its length considerably less than half the distance between the bases of the pair; 4-C single, simple; 5-C with from 4 to 6 pectinate branches; 6-C with 3 or 4 pectinate branches; 16, 17-C represented by minute spicules. Thorax. Integument glabrous; hairs 1, 2, 3-P single, pecti- nate, 3-P fine, considerably shorter than 1, 2-P; 4-P bifid or trifid, pectinate; 5, 6-P single, pectinate; 7-P single or bifid, pectinate; 8-P very short and fine, single or bifid, simple; 14-P single, simple. Abdomen. Integument glabrous; comb consisting of a broad, somewhat triangular patch of 70 or more fan-shaped scales; siphon index variable, ranging from 3.8:1 to 5.5:1 (average of 6 specimens, 4.4:1); 3 pairs of subventral tufts inserted on the siphon; individual tuft with from 2 to 5 simple branches, its length approximately equal to the width of the siphon at the point of insertion; pecten consisting of from 7 to 10 teeth restricted to the basal third or less of the siphon; individual pecten tooth rather short, with from 3 to 5 distinct basal barbs and a series of fine spicules at the truncate apex; saddle of segment X darkly pigmented, incomplete; ventral brush consisting of 8 hair tufts inserted on the grid; anal gills longer than the saddle, gradually tapering to a rounded apex. TYPE DATA. Lectotype female of obscurus from Kuala Lumpur is in the British Museum. Three lectoparatype females and 1 lecto- paratype male (which is badly damaged and with terminalia missing ) are also on deposit in the British Museum. Six individual rearings with associated larval and pupal skins have been examined from Malaysia as well as 14 additional males and nine females. DISTRIBUTION. This species was originally described from Kuala Lumpur, Selangor, MALAYSIA. Subsequently, Macdonald and Traub (1960) also collected specimens from the state of Selangor. Mac- donald, Smith, and Webb (1965) reported material from Kampong China, Kuching, Sarawak. Specimens have been examined in the British Museum collection from Tawau, Sabah. TAXONOMIC DISCUSSION. The monotypic subgenus Acalle- omyia demonstrates closest affinity to the subgenus Culiciomyia. The — Fig. 1, Culex (Acalleomyia) obscurus (Leicester). A, dorsoventral aspect of the head and thorax of the fourth stage larva; B, lateral aspect of the terminal gecorinal segments of the fourth stage larva; C, tergal aspect of the male terminalia. 56 PROC. ENT. SOC. WASH., voL. 70, No. 1, MARCH, 1968 phallosome of the male terminalia of obscurus is consistent with species of Culiciomyia in Southeast Asia and the fourth stage larva exhibits a number of points in chaetotaxy, particularly the ventral brush of abdominal segment X consisting of only eight hair tufts, which are suggestive of the subgenus Culiciomyia. However, the short male palpus is certainly not consistent with Culiciomyia, nor is the basimere of the male terminalia, which is clothed with scales as well as setae and which demonstrates a unique basomesal tubercle possessing two strong rods. In the fourth stage larva, head hair 1-C is exceptionally short and fine and the saddle of abdominal segment X is incomplete; otherwise, chaetotaxy and structure are quite similar to Culiciomyia. In the adult female of obscurus, the most noteworthy variation from Culiciomyia is the presence of a rather dense patch of broad, flat scales on the midlobe of the scutellum. As has been suggested for the subgenus Acallyntrum by Belkin (1962), it is quite possible that Acalleomyia represents an off-shoot of the Culiciomyia stem with specialized requirements for leaf axil or stump hole habitats. BIOLOGY. Macdonald and Traub (1960) collected larvae from coastal areas of Selangor in the axils and stumps of Nipa-palms. Specimens have been examined from Sabah which were collected from the saline habitat of a hole in a fallen mangrove tree. Ramalin- gam and associates (pers. comm. 1967) have collected larvae on eight occasions from the brackish water in Nipa-palm leaf axils in a secondary mangrove area and also collected a single female of this species biting man in the vicinity of the larval collections. Adults and larvae of obscurus have been collected during the months of July, October, December, and February. ACKNOWLEDGMENTS Dr. Botha de Meillon and Dr. Alan Stone kindly reviewed the manuscript and offered points of advice; Dr. P. F. Mattingly permitted examination of types in the British Museum (Natural History) and arranged for the loan of specimens; and Dr. S. Ramalingam forwarded individually reared specimens and data from Kuala Lumpur. To each of these men I extend my gratitude. The illustration was prepared by Miss Thelma L. Ford of the South East Asia Mosquito Project, and technical assistance was rendered by Miss M. T. Houck, also of SEAMP. REFERENCES Belkin, J. N. 1962. The mosquitoes of the South Pacific. Univ. Calif. Press, Berkeley. 2 vols., 608 and 412 pp. Bram, R. A. 1967. Contributions to the mosquito fauna of Southeast Asia. II. The genus Culex in Thailand (Diptera: Culicidae). Cont. Amer. Ent. Inst. 2(1):1—296. Edwards, F. W. 1922. A synopsis of adult oriental culicine (including mega- rhinine and sabethine ) mosquitoes. Part I. Indian Jour. Med. Res. 10:249-293. PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 57 Leicester, G. F. 1908. The Culicidae of Malaya. Stud. Inst. Med. Res. F. M. S. (3):18-261. Macdonald, W. W., C. E. G. Smith, and H. E. Webb. 1965. Arbovirus infec- tions in Sarawak: observations on the mosquitoes. Jour. Med. Ent. 1:335-347. Macdonald, W. W. and R. Traub. 1960. Malaysian parasites XXXVII. An introduction to the ecology of the mosquitoes of the lowland dipterocarp forest of Selangor, Malaya. Stud. Inst. Med. Res. Malaya 29:79-109. GRYLLOPROCIPHILUS FROSTI, NEW GENUS, NEW SPECIES, FROM THE EASTERN UNITED STATES (HoMopTERA: APHIDIDAE )! CrypE F. SmirH and JouHNn O. PEPPER? Grylloprociphilus frosti probably lives on the subterranean parts of its unknown host for the major portion of its life cycle as is customary with most species in related genera. It is named in honor of Dr. S. W. Frost who collected the majority of the known specimens. It is described in the hope that other entomologists might help unravel the life history of this interesting species which apparently produces sexuparae only during the shortest days of the year. Grylloprociphilus, n. gen. Antennae 6-segmented, secondary sensoria oval, ciliated. Fore wings with media simple. Hind wings with media and cubitus originating near the same point. Metafemora on sexuparae distinctly larger than pro- or mesofemora. Cauda rounded, indistinct. Grylloprociphilus differs from Pemphigus Hartig, 1839, Prociphilus Koch, 1857, and Stagona Koch, 1857, in having the metafemora enlarged. From Pemphigus it differs in having more setae on R IV + V and the secondary sensoria with long cilia. From Prociphilus and Stagona it differs in having spine-like setae at the apex of the tibiae. Type-species: Grylloprociphilus frosti, n. sp. Grylloprociphilus frosti, n. sp. ( Bigs? e423) Characteristics: Head without wax plates. Secondary sensoria distinctly ciliated, sensoria on a.s. III on distal half, a.s. VI with numerous, conspicuous setae. R IV + V bearing 4 to 8 accessory setae. * Paper number 2197 of the Journal Series of the North Carolina State University Agricultural Experiment Station, Raleigh, N. C. Professor, North Carolina State University at Raleigh, and Professor Emeritus, Pennsylvania State University, University Park, Pa., respectively. The authors express their appreciation to Louise M. Russell, U.S. Department of Agriculture, Washington, D. C.; D. Hille Ris Lambers, Bennekom, Netherlands, and H. L. G. Stroyan, Harpenden, Herts, England for their opinions and to the U.S. National Museum for loan of specimens. 58 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Fig. 1, Grylloprociphilus frosti, n. sp.: (A) antenna; (B) R IV + V; (C) cauda. (Drawings by John Graham) Wax plates on dorsum of thorax distinct, distance between wax plates being approximately equal to longest diameter of plates. Protarsomere I with one seta distinctly shorter and more peg-like than the other setae. A slide in the U.S. National Museum bears the following data, “Flying, Nov. 19. 15. Abd. yellowish green slightly covered with cottony excretion.” Sexupara: Head dark, without wax plates. Antennae dark; setae on a.s. III approximately equal to % diameter of segment, setae conspicuous, especially on a.s. VI; secondary sensoria on distal half of segments, distinctly ciliated, outer rim approximately % to 34 diameter of segment. R IV + V attaining second coxae and bearing 4 to 8 accessory setae. Pro-, meso-, and metathorax with distinct oval wax plates. Distance between plates approximately equal to greatest diameter of plate. Metafemora distinctly larger (wider) than pro- or mesofemora. All femora rugose dorsally, denticulate ventrally. Distal end of each tibia with 4 spine-like setae. Other setae on tibiae long, many of them being equal to or longer than diameter of tibia. Tibiae without denticulations. Chaetotaxy of tarsomeres I-III, 3 to 7, variable, usually 6-5-4. Protarsomere I bearing one seta distinctly shorter and more spine-like than the other setae. Plantar setae approximately 4% length of claw, pointed. Tarsomeres with faint denticulations. Abdomen with marginal wax plates which are nearly circular and increasing slightly in diameter toward the caudal end. Marginal wax plates bearing 1 to 6 setae, those nearest the caudal end having the greater number. Two round medial wax plates on dorsum of abdominal segment I and one elongated plate on segment VIII which bears 4 to 9 setae. Genital plate with approximately 12 setae on the caudal margin and 12 more scattered over the rest of the plate. Gonapophyses variable, but usually bearing 4-3-4 setae. Anal plate with 5 to 9 setae, about the same shape and length as the setae on the cauda. Cauda bearing 3 to 5 setae. Measurements:* Body length 2.43 (1.73-2.94). RIV + V, 0.15 (0.11-0.19); a.s. III, 0.24 (0.18-0.29); IV, 0.14 (0.09-0.15); V, 0.15 (0.11-0.19); VI, 0.16 (0.14-0.19) + 0.04 (0.03-0.04). Hind tibiae 0.88 (0.65-0.89); hind tarsi, 0.22 (0.16-0.25). Secondary sensoria. A.s. III 3-4 (3-5), IV, 2-3 (2-3), V, 2-3 (1-3). Embryos of the sexupara with vestigial mouth parts. Plantar setae longer than the claws and expanded at the tip. 3 All measurements are in millimeters. The first number refers to holotype, the numbers in parentheses show the range of 11 specimens measured. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 59 Fig. 2, Grylloprociphilus frosti, n. sp. (Photograph by A. T. Olive) Type locality: Lake Placid (Archbold Biological Station), Florida. Types: Holotype and some paratypes in the United States National Museum. Other paratypes in the collections of H. L. G. Stroyan, Harpenden, Herts, England; D. Hille Ris Lambers, Bennekom, Nether- lands; the British Museum of Natural History, London, England; the Canadian Museum of Natural History, Ottawa, Canada; A. N. Tissot, Gainesville, Florida; the North Carolina State University at Raleigh; and the authors. Collections: Data on 3 slides in the United States National Museum indicate this species was first collected “flying” November 18, °95 (probably 1895) (one specimen). No locality or collector recorded. In 1941 W. F. Turner collected a single specimen on January 29 in Peach County, Georgia. On February 3, 1932, P. W. Mason collected 3 specimens (one slide) “swarming” in Washington, D.C. Numerous specimens have been collected by S. W. Frost in a black light trap 60 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 at Lake Placid (Archbold Biological Station), Florida, 1958-1964 (Holotype and Paratypes), during the months of November through January. In a note Dr. Frost wrote “104 specimens taken night of Jan. 4, 1964, very few taken before or afterwards.” John Graham collected one specimen in a “Sticky band trap” in a “Beech woods” at Raleigh (Umstead Park), North Carolina, February 24, 1962, one specimen in a “cob web, Nov. 28, 1963” and one “in flight, Raleigh, N.C., Jan. 5, 1962.” J. O. Pepper and B. F. Coon collected one speci- men in a black light trap at Center Hall, Pa., Oct. 19-26, 1958. We do not know the host of this aphid. TEN NEW CHRYSOMELID BEETLES FROM DOMINICA AND JAMAICA Doris H. Buaxe, Department of Entomology, Smithsonian Institution, Washington, D.C. 20560 The following new species of Chrysomelidae, with one exception, were collected during the Bredin-Archbold Smithsonian Biological Survey in Dominica. A. B. Gurney on his way back from Dominica collected a new species of Oedionychus in Jamaica. Habrophora thelmae, n. sp. (Fig. 1) Elongate oblong oval, clothed with short, fine, appressed, yellow pubescence; head, prothorax and breast dull brown, elytra, abdomen and legs paler yellow brown, elytra with dark spot on basal umbone and a dark lateral stripe from humerus along side to middle, an irregular fascia across elytra at middle, also 3 inconspicuous costae on each elytron with dense striate punctures on each side of them, the intervals between confusedly punctate; antennae pale yellow brown with joints 7, 8, 10 and 11 dark. Head with interocular space half width of head, eyes emarginate, occiput and front with fine yellow pubescence, an inconspicuous dark median line, clypeus with finely punctate surface not completely hidden by pubescence, labrum pale, jaws piceous. Antennae extending to middle of elytra, long and slender, pale yellow brown with distal joints dark. Prothorax densely covered with pale pubescence that hides the dense punctation; a transverse depression in basal half, convex with rounded sides and very thin line marking lateral margin, prosternum taken up with coxal cavities, between these a moderately wide median area. Scutellum densely pubescent. Elytra elongate, thinly covered with appressed yellow pubescence and with 3 somewhat inconspicuous costae edged on either side with a dense row of striate punctures, intervals with confused punctures; PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 61 a prominent basal umbone on each elytron and small prominent humeri with intrahumeral sulci; basal umbones deep brown, humeri and sides to middle also brown, at middle an irregular brown fascia across. Femora slightly enlarged, tibiae long and slender, claws toothed. Length 5.5 mm, width 2.2 mm. Type, female, USNM 69718. Type locality—Pont Lolo, 1800 ft. alt., Dominica, Feb. 19, 1965, collected by J. F. G. and T. M. Clarke. Remarks.—Although this species certainly belongs in an Eumolpid genus close to Fidia, Xanthonia and Habrophora, and especially the last, it differs from them by having the elytral punctation not definitely striate. As in Habrophora there are three elytral costae on each elytron, each having on each side a row of dense striate punctures. Because of these costae edged with striate punctures, I am putting it in the genus Habrophora, but it differs from the other species of the genus that I have examined in having distinctly confused puncta- tion in the intervals and in having sparser elytral pubescence. I take pleasure in naming this after Mrs. J. F. G. Clarke, who did her share of the collecting on Dominica. Alethaxius dominicae, n. sp. (Fig. 7) Oblong oval, yellow brown with deeply and densely punctate head, prothorax and elytra, elytra with punctures somewhat confused in basal half, on sides and at apex striate punctures with costate intervals, much more pronounced in female. Head with interocular space approximately half width of head, a median line down front, surface densely and deeply punctate, clypeus dull and alutaceous and separated from front by somewhat indistinct sulcus. Antennae extending below humeri, joints 7-11 darker and wider. Prothorax moderately convex with 2 angularities on each side, dull yellow brown, deeply and moderately densely punctate. Scutellum dark. Elytra with confused punctures in basal half from about scutellum to middle, on sides and below the middle in apical half striate rows of punctures with costate intervals, much more developed in female. Body beneath reddish brown, legs yellowish brown, not toothed. Length 2.3-2.7 mm; width 1-1.5 mm. Type, male, USNM 69719. Type locality —Clarke Hall, Dominica, Feb. 4-7, 1965, H. E. Evans. Other locality —‘' mile west of Pont Lolo, 1800 ft. alt., Dominica, Feb. 19, 1965, J. F. G. and T. M. Clarke. Remarks.—In size and coloration this is very much like A. puerto- ricensis Blake, but lacks the knobby projections on the elytra, espe- cially noticeable in the female of that species, although even the male has a few knobs about the humeri. In this species the female has very strongly developed costae that in the male are noticeable on the sides and apex. The single female was collected farther up the mountain but in the same general area, and there is no question it is the same species. 62 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Metachroma gagnei, n. sp. (Fig. 6) Oblong oval, shining, yellow brown with reddish brown basal and 2 lateral spots on each elytron; striate punctures on elytra feeble over basal callosities, fading away on sides and at apex; pronotum impunctate; antennae with apical joints darker. Head with interocular space less than half width of head, occiput and front nearly impunctate, a transverse sulcus between antennal sockets, clypeus nearly impunctate. Antennae long and slender with apical joints thicker and darker. Prothorax evenly rounded but not very convex with rounded lateral sides and depression over head; mirror smooth, yellow brown. Scutellum reddish brown. Elytra with striate punctures nearly obsolete over basal callosities and indistinct on sides and at apex; a median long reddish brown basal spot on each elytron and 2 others along the side, one below humerus, the other, larger, at about the middle. Body beneath yellow brown, shining, legs pale, femora not toothed. Length 2.5 mm; width 1.3 mm. Type, female, USNM 69720. Type locality —Path to Cabrits, Dominica, 2 April 1966, R. J. Gagne. Remarks.—There is only one specimen of this small species which is about the size of M. testaceum Blatchley from the tip of Florida and Cuba. It is the smallest species of Metachroma from Dominica yet collected and rather distinctively marked with the reddish brown elytral spotting. Galerucella brevivittata, n. sp. (Fig. 2) Elongate with short prothorax and long, parallel-sided elytra, dull yellowish brown with dark head, prothorax with a diagonal dark mark on either side, elytra with 6 deeper brown vittae and brown sutural edges, the median vittae not reaching base or apex, all covered with very short fine pubescence. Head with wide set eyes and a median line down occiput to interantennal area, little evidence of tubercles or frontal carina, lower front short and somewhat depressed, occiput and mouthparts dark. Antennae extending below humeri, 2 basal joints rather inflated, 3rd joint longest of all, remainder diminishing and somewhat wider, with base pale, apex dark, apical 3 joints entirely dark. Pro- thorax transverse with sides and median anterior area depressed, at each angle a small tooth, dull yellow brown with piceous diagonal plaga on each side, surface finely and densely punctate with a few short hairs. Scutellum dark. Elytra elongate, parallel-sided with prominent humeri and short intrahumeral sulcus, depressed below scutellum, surface finely punctate and covered with short fine yellowish pubescence, closely appressed; yellow brown with darker sutural edges, and on each elytron 2 median dark brown vittae, 1 from the scutellum, the other from end of intrahumeral sulcus, not reaching base or apex, a 3rd from the humerus. Body beneath dark except for pale yellow brown prosternum, and somewhat darker legs. Length 5.2 mm; width 2.5 mm. Type, male, USNM 69721 and four paratypes. Type locality—Clarke Hall Estate, Dominica, May 31, 1966, G. Steyskal. Feb. 4-7, 1965, H. E. Evans. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 63 Other localities; —3 m. w. Pont Lolo, 1800 ft. alt. Feb. 19, 1965, J. F. G. and T. M. Clarke. Remarks.—This is another of the species of West Indian Galerucella with long, vittate elytra. In this one the median vittae do not reach entirely from base to apex. Other vittate species closely resembling it but differing in the vittae are G. jamaicensis Blake, G. melanocephala Blake, G. maculipes Blake, and G. oteroi Blake, the last three from Cuba. Oedionychus gurneyi, n. sp. (Fig. 3) Elongate oblong oval, shining, nearly impunctate, rather flat with wide explanate margin, yellow brown, pronotum with wide piceous vitta curving on the inner side on each side near explanate margin, elytra with a piceous marking covering humerus and extending below middle, across apex an irregular piceous band. Head with interocular space less than half width of head, yellowish brown with well developed frontal tubercles, a short carina and short lower front; fine punc- tures over upper face. Antennae yellow brown with apical joints deeper in color- ing, slender, not reaching middle of elytra. Prothorax nearly flat with wide explanate margin, yellow brown with a wide curving piceous vitta on each side; surface shining with almost imperceptible fine punctures. Scutellum triangular, pale. Elytra long, wider than prothorax, with wide explanate margin and about 5 times as long as prothorax; shining pale yellow brown with irregular piceous marking from humerus to beyond middle, before apex a narrow irregular band nearly across elytra. Body beneath and legs pale yellow brown. Length 6.8 mm; width 3.3 mm. Type, female, USNM 69722 and one paratype, female. Type locality—Green Hills, Portland Parish, ca. 3800 ft. alt., Nov. 13-20, 1966, A. B. Gurney. Remarks.—The dark piceous markings on the pronotum and elytra are unlike those of any other species of Oedionychus found in the West Indies, and because of the high mountain region in which it was collected, I believe it is endemic in Jamaica. I take pleasure in naming it for its collector, Dr. Gurney. Hadropoda xanthoura, n. sp. (Fig. 9) Elongate oblong oval, reddish or yellowish brown, densely covered with short fine golden and brown pubescence, prothorax only a little wider than long with nearly straight sides, elytra much wider than prothorax with striate punctation more or less covered by pubescence, an irregular brownish fascia below middle made by short brown pubescence, which is also present on basal callosities; apex densely clothed with yellow pubescence. Head brownish with interocular space less than half width of head, eyes large, face long, a deep linear depression above carina between absent frontal tubercles, carina between closely set antennal sockets very narrow and short, lower front depressed, occiput covered with short pale pubescence. Antennae filiform, basal and 2nd joints swollen, 3rd joint long and slender, longer than 4th or 5th which 64 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 > ? - are subequal, 6th shorter, 7-10th shorter and wider and tending to be darker. Prothorax only slightly wider than long, convex with a hump in middle and depressed in basal part; pale yellowish brown covered with fine, appressed, short pubescence. Scutellum pale and with fine pubescence. Elytra much wider than prothorax, regularly striate punctate, with strong basal callosities, surface covered more or less by fine, short, appressed pubescence that over basal callosities tends to be deep brown and also an irregular brown fascia of brown pubescence across elytra in apical half, rest of elytra with short golden pubescence. Body beneath and legs pale yellowish or reddish brown, less densely pubescent. Length 3.9—4.2 mm; width 3.6-1.8 mm. Type, male, USNM 69723. Type locality—2.5 m east of Pont Casse, Dominica, Jan. 16, 1965, W. W. Wirth. Other localities—Freshwater Lake, 21 Jan. 1965, W. W. Wirth; Springfield Estate, 20 July 1963, O. S. Flint, Jr.; S. Chiltern Estate, 20 Feb. 1965, W. W. Wirth. Remarks.—This is unlike any other species of Hadropoda in the markings. It resembles somewhat H. calva Blake from the Dominican Republic, but that species has the occiput devoid of pubescence. Lactica flinti, n. sp. (Fig. 4) Elongate oblong oval, shining, nearly impunctate, pale reddish brown with antennae except 2 basal joints and tarsi except 1st joint of hind one dark, a dark reddish brown spot on either side of pronotum. Head with interocular space more than half width of head, a large puncture surrounded by several smaller ones on either side near eye, frontal tubercles rather indistinct and a very short carina, lower front depressed. Antennae long and except for 2 basal joints dark. Prothorax with slightly rounded sides and a feebly marked basal sulcus, on either side a deeper reddish brown spot; surface shining, minutely punctate. Scutellum pale. Elytra elongate, with small sharp humeri, a short intrahumeral sulcus and faint umbone at base; shining, reddish brown, not visibly punctate. Body beneath shining reddish brown with few punctures and nearly glabrous, legs reddish brown except for black tarsi, and the first joint of hind tarsi not dark. Length 3.5-3.7 mm; width 1.8 mm. Type, male, and one paratype, female, USNM 69724. Type locality —.4 mile E. of Pont Casse, collected May 6 and June 23, 1964, O. S. Flint, Jr. Remarks.—Dr. Flint tells me these beetles were taken at about 2000 ft. altitude, at the beginning of the rain forest. I have already de- scribed one other pale species of Lactica from Dominica, collected at 800 ft. altitude by R. G. Fennah. L. flinti differs from this one in coloration having only the tarsal joints dark, and also in having two spots on the pronotum. This pronotal spotting is unlike any up to now found in these pale species from the West Indies. The basal sulcus on the pronotum is also feebler in comparison with the rest. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 65 Heikertingerella wirthi, n. sp. (Fig. 10) Ovate, shining yellow brown, finely punctate, last 5 joints of antennae deeper brown. Head with interocular space less than half width of head, eyes very large, interantennal area narrow and produced in a long carina extending down front, deep brown. Antennae with 2 basal joints swollen, 3rd short, remainder long, subequal and filiform, last 5 or 6 joints deeper brown. Prothorax yellowish or even reddish brown, minutely punctate, sides contracting in a straight line towards eye, basal margin sinuate over scutellum. Scutellum yellowish or reddish brown. Elytra convex, widest at middle, with small humeri, surface moderately shiny, finely punctate, yellowish brown. Body beneath and legs yellowish or reddish brown. Length 2.2-2.9 mm; width 1.5-1.8 mm. Type, male, USNM 69725 and four paratypes. Type locality—Freshwater Lake, Dominica, April 14, 1964, O. S. Flint, Jr. Other localities—Clarke Hall, 11-20 March 1965, W. W. Wirth; Tareau Cliffs, La Franchetti, April 13, 1966, R. J. Gagne. Remarks.—It is difficult to distinguish this species from H. dom- inicae Blake except by comparison of the aedeagi. In general H. dominicae is larger and wider and more rounded, and not so distinctly punctate. In my description of H. dominicae I wrote that two speci- mens collected by R. H. Foote on Dominica, both females, were more distinctly punctate than the type specimen. I believe now that these may be H. wirthi although there is no certainty because they are both females. Sidfaya philtata, n. sp. (Fig. 5) Ovate, moderately convex, shining black with yellowish brown mouthparts, antennae and legs, hind femora dark; densely and coarsely punctate; prothorax without depressions and with oblique anterior angles and an angularity at middle of sides. Head with smooth polished occiput and a deep line on either side from above eyes to well marked frontal tubercles; interantennal area broad and extending a short way down front, lower front short and mouthparts somewhat declivous and pale. Antennae extending below humeri, 2 basal joints somewhat swollen, joints 3-7 pale and slender, apical joints thicker and darker. Prothorax moderately convex, anterior angles widely oblique, an angle at middle of side, and a small basal tooth; surface shining, black with coarse but not dense punctation. Scutellum dark. Elytra with sharp prominent humeri and slight basal umbone having a faint transverse depression below; surface shining black with coarser and denser punctation than on pronotum. Body beneath and hind femora shining black, nearly impunctate and glabrous. Length 2.4 mm; width 1.4 mm. Type, female, USNM 69726 and two paratypes, both female. Type locality—a1.6 miles west of Pont Casse, Dominica, April 24, 1964, O. S. Flint, Jr.; April 10, 1966, R. J. Gagne. 66 PROG. ENT, SOC. WASH., VOL. 70, NO. ) S ]| i) L Habrophora thelmae 5. Sidfaya philtata 1, MARCH, 1968 6.Melachroma gagnei 7Alethaxius dominicae 8.Homoschema dominicae 9. Hadropoda xanthoura 10. Heikertinderella wirtht PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 67 Remarks.—This is the third species of the genus to be described, the first being S. polutima Blake from Cuba, the second, S. punctatissima Blake from Jamaica. In its short broad prothorax and coarse elytral punctation this resembles the Jamaican species, but in its ovate shape it is more like the Cuban species. All three species are shining black with coarse, confused punctation, and the prothorax has broad oblique angles anteriorly and in the middle of the side a distinctive angularity. Homoschema dominicae, n. sp. (Fig. 8) Lustrous, head, prothorax, legs and antennae orange yellow, elytra bluish or greenish, often purplish, very finely punctate. Head with wide set eyes, rounded occiput, a line of punctures from eyes to well developed frontal tubercles, a very short carina and short lower front of face. Antennae extending below humeri, basal joints swollen. Prothorax moderately convex with well rounded sides and oblique anterior angles and small tooth at basal angles, basal margin nearly straight; disc nearly impunctate, lustrous orange yellow. Scutellum dark brown. Elytra lustrous greenish or purplish blue, mod- erately convex with transverse depression below basal callosities; very finely and inconspicuously punctate. Body beneath with abdomen and sometimes _ breast brown; hind legs with swollen femora. Length 2.3-2.6 mm; width 1.3-1.5 mm. Type, male, and eight paratypes, USNM 69727. Type locality—Mero, Dominica, collected by O. S. Flint, Jr., 24 July 1963. Remarks.—This is one of the smaller species of Homoschema and with its dark abdomen akin to H. nigriventrum Blake from Puerto Rico, although the aedeagus is more like that of H. jamaicense Blake. NEW SPECIES OF NEOACARUS HALBERT AND VOLSELLACARUS COOK FROM NORTH AMERICA! (ACARINA: NEOACARIDAE ) Davip R. Coox, Department of Biology, Wayne State University, Detroit, Michigan 48202 Cook (1963) described four species of Neoacarus and the new neoacarid genus Volsellacarus from the interstitial waters of North America. The present study? adds two new species of Neoacarus and one new species of Volsellacarus, all from the ground water habitat. In presenting measurements in this paper, those of the holotype and * Contribution No. 189 from the Department of Biology, Wayne State University. 2 Supported by a grant (GB-3717) from the National Science Foundation. 68 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MaRcH, 1968 Figs. 1-5, Neoacarus occidentalis, n. sp.: 1, ventral shield, 2; 2, ventral shield, 4; 3, Ill-Leg-4, 5, 6, 4; 4, L-Leg-5 and 6, 2; 5, palp, ¢. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 69 allotype are given first. If a series of specimens is available, the range of variation is given in parentheses following the measurements of the primary types. Holotypes and allotypes will be placed in the Field Museum of Natural History (= Chicago Natural History Museum). Neoacarus occidentalis, n. sp. (Figs. 1-5) Male: Dorsal shield 502 uw in length, 349 uw in width; ventral shield, including first coxae, 562 w in length, 410 uw in width; ventral shield oval; first coxae fused medially without any trace of a suture line between them; third and fourth coxae separated medially by the genital bay; fourth coxae extending well posterior to the genital field (fig. 2); genital field long and narrow, 107 u in length; three pairs of elongated genital acetabula present, these occupying opening of gonopore; gonopore flanked by a pair of glandularia; eyes reduced, integumental pigment absent. Dorsal lengths of the palpal segments: P-I, 24 u; P-II, 38 uw; P-III, 19 »; P-IV, 42 u; P-V, 22 uw; ventral portion of P-IV expanded into a pointed projection, the distal margin of which bears 2 spine-like setae (fig. 5); dorsal lengths of the distal segments of the first leg: I-Leg-4, 80 uw; I-Leg-5, 97 yw; I-Leg-6, 110 uy; dorsal lengths of the distal segments of the third leg: III-Leg-4, 103 u; II-Leg-5, 134 uw; II-Leg-6, 176 uw; Il-Leg-5 expanded ventrally, gradually tapering towards distal end; fig. 3 illustrates the chaetotaxy of these segments; anterior surface of IlI-Leg-5 with a patch of swimming hairs (shown as stippled setal bases on fie. 3): Female: Dorsal shield 509 yu in length, 364 » in width; ventral shield, includ- ing projecting first coxae, 566 w in length, 426 w in width; ventral shield oval; first coxae fused medially without a trace of a suture line; second and third coxae slightly separated medially; fourth coxae forming a shallow genital bay; genital field 131 » in length, 129 uw in width; gonopore relatively large; 7 genital acetabula present on each side, these lying on elongated sclerites on their respective sides; acetabular sclerites only lightly fused with the ventral shield (fig. 1); eyes reduced in size, integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 19 «; P-II, 40 uw; P-III, 17 yu; P-IV, 42, u; P-V, 22 uw; structure and chaetotaxy of palp similar to that illustrated for the male; dorsal lengths of the distal segments of the first leg: I-Leg-4, 79 u; I-Leg-5, 93 uw; I-Leg-6, 100 u; Fig. 4 illustrates the proportions and chaetotaxy of I-Leg-5 and 6. Types: Holotype, adult male, taken in a gravel bar in Battle Creek adult female, same data as holotype. A newly metamorphosed male four miles west of Viola, Shasta Co., California, July 30, 1966; allotype, which appears to belong to this species was collected in a gravel bar in Bingham Creek approximately eight miles northeast of Satsop, Mason Co., Washington, August 5, 1966. It is not assigned to the type series. Discussion: This is the first species of Neoacarus known from the West Coast area of North America. The structure of the palp, with the projection on P-IV ventral rather than distoventral in position, will 70 PROC. ENT, SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Ce eee ne A aL aS c | | Figs. 6-10, Neoacarus minimus, n. sp.: 6, ventral shield, 2; 7, ventral shield, é; 8, Ill-Leg-4, 5, 6, ¢; 9, I-Leg-5 and 6, 2; 10, palp, 9. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 al. separate N. occidentalis from any previously described species. How- ever, the following species possesses a similar palp. See the discussion section under the latter. Neoacarus minimus, n. sp. (Figs. 6-10) Male: Dorsal shield 380 » (380-403 xu) in length, 273 uw (273-289 wu) in width; ventral shield, including first coxae, 456 u (456-470 «) in length, 304 u (304-327 «) in width; ventral shield oval; first coxae fused medially without any trace of a median suture line; third and fourth coxae separated medially by the genital field; fourth coxae extending only slightly posterior to the genital field; genital field long and narrow, 95 « (95-97 «) in length; 3 pairs of genital acetabula present, these occupying opening of gonopore; gonopore flanked by a pair of glandularia and 2 pairs of relatively long setae (fig. 7); eyes reduced, integu- mental pigment absent. Dorsal lengths of the palpal segments: P-I, 20 » (19-20 uw); P-II, 34 w (32-35 #); P-III, 18 » (17-18 uw); P-IV, 35 w (35-37 uw); P-V, 19 » (18-21 «); ventral portion of P-IV expanded into a pointed projection, the distal margin of which bears 2 small spine-like setae; dorsal lengths of the distal segments of the first leg: I-Leg-4, 69 uw (66-69 uw); I-Leg-5, 95 w (90-95 uw); I-Leg-6, 107 w (104— 110 «); dorsal lengths of the distal segments of the third leg: IlI-Leg-4, 93 u (87-93 w); Il-Leg-5, 107 » (104-107 ~); II-Leg-6, 135 w (131-140 ,); III- Leg-5 expanded ventrally, gradually tapering towards distal end; ventral expansion of III-Leg-5 pointing somewhat proximally; fig. 8 illustrates the chaetotaxy of these segments; anterior surface of III-Leg-5 bearing a small patch of swimming hairs (shown as stippled setal bases in fig. 8). Female: Dorsal shield 414 » (395-414 uw) in length, 304 uw (289-304 xu) in width; ventral shield, including first coxae, 486 u (464-486 uw) in length, 334 yu (326-334 w) in width; ventral shield oval; first coxae fused medially without a trace of a suture line; fourth coxae forming a relatively deep genital bay (fig. 6); genital field 121 » (117-121 w) in length, 103 » (98-103 ~) in width; gonopore relatively large; 7-8 (6-8) genital acetabula present on each side, these lying on elongated sclerites on their respective sides; acetabular sclerites fused with the ventral shield; eyes reduced, integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 24 w (21-24 uw); P-II, 35 w (34— 35 mw); P-III, 20 » (17-20 »); P-IV, 36 uw (36-37 u«); P-V, 20 uw (17-20 xu); fig. 10 illustrates the proportions and chaetotaxy of the palp; dorsal lengths of the distal segments of the first leg: I-Leg-4, 64 u (62-64 uw); I-Leg-5, 87 u (86-87 un); I-Leg-6, 103 » (96-103 »); fig. 9 shows I-Leg-5 and 6. Types: Holotype, adult male, collected in a gravel bar in a stream near Twin Mountain, Coos Co., New Hampshire, August 21, 1964; allotype, adult female, from gravel deposits in a stream near Griffen (on Route #8, three miles from Warren Co. line), Hamilton Co., New York, August 18, 1964. Paratypes: two males, same data as holotype; one female, same data as allotype; one male, collected in a gravel bar in a small stream near Warm Springs, Bath Co., Virginia, July 16, 1963. 72 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Figs. 11-15, Volsellacarus sabulonus, n. sp.: 11, ventral shield, 2; 12, palp, 9; 13, IlI-Leg-5 and 6, 6; 14, I-Leg-5 and 6, 9; 15, ventral shield, 6. Fig. 16, V. ovalis Cook: IlI-Leg-5 and 6, é. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 The Discussion: The present species resembles the preceding species, N. occidentalis, in structure of the palp but may be easily distinguished as follows. The female of minimus differs from all previously described species of the genus in possessing a deep genital bay (fig. 6). Males of minimus and occidentalis may be easily separated by the structure of the third leg (figs. 3 and 8) and the morphology of the ventral shield. Males of N. occidentalis have a much deeper genital bay. N. minimus is also the smallest known species in the genus. Volsellacarus sabulonus, n. sp. (Figs. 11-15) Male: Dorsal shield 638 u in length, 456 u» in width; ventral shield, including first coxae, 718 « in length, 517 » in width; ventral shield oval; first coxae fused medially without a trace of a suture line medially; third and fourth coxae separated medially by the genital field; fourth coxae extending slightly posterior to the genital field; 4 long setae present on each side immediately posterior to the insertions of the fourth legs (fig. 15); genital field long and narrow, 138 wu in length; 3 pairs of elongated genital acetabula present, these occupying the opening of the gonopore; gonopore flanked by a pair of glandularia; eyes reduced, integu- mental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 50 u; P-II, 88 uw; P-III, 52 u; P-IV (including distoventral projection), 240 »; P-V, 124 wu; structure of palp similar to that illustrated for the female; dorsal lengths of the distal segments of the first leg: I-Leg-4, 145 uw; I-Leg-5, 171 uw; I-Leg-6, 173 yu; dorsal lengths of the distal segments of the third leg: III-Leg-4, 180 y; IlI-Leg-5, 221 u; IlI-Leg-6, 201 u:; ventral side of III-Leg-5 with a slight bulge near proximal end, this bulge bearing a patch of short slightly-thickened setae (fig. 13); a few swimming hairs located at distal end of III-Leg-5. Female: Dorsal shield 745 uw (684-745 w) in length, 555 » (498-555 uw) in width; ventral shield, including first coxae, 836 » (764-836 ) in length, 616 u (578-616 «) in width; ventral shield oval; first coxae fused medially without a trace of a median suture line; second and third coxae slightly separated medially; fourth coxae forming a shallow genital bay; genital field 183 «4 (178-183 y) in length, 166 » (159-166 «) in width; gonopore relatively large; 6 pairs of genital acetabula present on each side, these lying on elongated sclerites on their respec- tive sides (fig. 11); acetabular sclerites fused with the ventral shield; eyes reduced, integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 67 » (64-67 «); P-II, 110 uw (104— 110 »); P-III, 64 « (62-64 «); P-IV (including distoventral projection), 322 u (312-322 w); P-V, 169 w (162-169 uw); fig. 12 illustrates the proportions and chaetotaxy of the palp; dorsal lengths of the distal segments of the first leg: I-Leg-4, 173 « (148-173 »); I-Leg-5, 190 « (180-190 »); I-Leg-6, 180 » (169- 180 «); fig. 14 shows I-Leg-5 and 6. Types: Holotype, adult male, taken in a sand bar in a small stream five miles west of Marietta (on Alternate US 50), Washington Co., Ohio, July 26, 1964; allotype, adult female, from a gravel bar in a small creek six miles north of Millboro Springs, Bath Co., Virginia, July 24, 1964; paratype, one female, same data as allotype. 74 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Discussion: This is the second known species in the genus Vol- sellacarus. It changes the original conception of the genus only in that the first coxae of the male may be pointed (fig. 15) rather than rounded as in V. ovalis. II-Leg-5 of the male, while without pro- nounced sexual dimorphism as in Neoacarus, does exhibit valuable specific characters. Figs. 13 and 16 illustrate I[I-Leg-5 and 6 of sabulonus and ovalis respectively. The new species is proportionally much narrower than V. ovalis. Two species of Neoacarus, originally described from the Ozark and Rocky Mountain area, are now known to occur in eastern North America. Neoacarus similis Cook: One male, from gravel deposits in a stream near Griffen (on Route #8, three miles from Warren Co. line), Hamilton Co., New York, August 19, 1964; one female, taken in a gravel bar in a small stream near Limestone, Victoria Co., New Brunswick, August 26, 1964; three males, one female, found in a gravel bar in the North Branch of the Meduxnekeag River at Monticello, Aroostook Co., Maine, August 28, 1964. Neoacarus ozarkensis Cook: One male, taken in a gravel deposit in Knapp’s Creek near Minnehaha Springs, Pocahontas Co., West Virginia, July 22, 1964; one male, two females, from gravel deposits in a stream near Griffen (on Route #8, three miles from Warren Co. line), Hamilton Co., New York, August 19, 1964. REFERENCES Cook, David R. 1963. Studies on the phreaticolous water mites of North Amer- ica: the family Neoacaridae. Ann. Ent. Soc. Amer. 56:481—487. A NEW SPECIES OF LYGAEUS FROM PERU (HEMIPTERA: LYGAEIDAE) LEON J. Gorski, Department of Zoology and Entomology, University of Connecticut, Storrs, Connecticut 062681 The genus Lygaeus Fabricius is the type genus of the family Lygaeidae and was originally used to include many species now in other heteropterous families as well as throughout the Lygaeidae. Attempts to better define the various genera of the Lygaeidae have resulted in the use of the phallus as a taxonomic character. Ashlock 1 Present address: Department of Biological Sciences, Central Connecticut State College, New Britain, Connecticut 06050. PROC. ENT. SOC. WASH., voL. 70, No. 1, MARCH, 1968 75 2 3 Figs. 1 and 3, Lygaeus slateri, n. sp.: 1, $6 paramere; 3, membrane, dorsal view. Fig. 2, L. inaequalis Walker, ¢ paramere. (1957) points out the value of the phallus for discrimination of the higher taxa in the Lygaeidae and as a secondary consideration the use of the sclerotized portions (parameres) for possible specific discrim- inations. It is the paramere or clasper characteristic which has been used primarily to define Lygaeus slateri n. sp. as a part of the revision of the entire genus. The parameres are bilaterally symmetrical and relatively simple in Lygaeus. Three regions are definable on each paramere: the basal shank, the proximal hook, and the apical hook. The apical hook is arm-like with a curvature of varying intensity. This portion is always hollowed out along its inner surface. The basal shank also shows a varying type of concavity. The inner margin of the shank portion of the paramere exhibits many of the specific modifications. Specific differences can most readily be seen when the left paramere is oriented with the inner concave side dorsal. In this position one can see the concavity of the arm and shank as well as the outline provided by the presence or absence of processes, figs. 1 and 2. Lygaeus slateri, n. sp. (Figs. 1 and 3) Head, anterior portion of pronotum, scutellum and distal abdominal segments grey-black; anterior half of clavus and posterior half of pronotum yellow-orange; 76 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Ui Fig. 4, Lygaeus inaequalis Walker, dorsum, ¢. PROC. ENT. SOC. WASH., vOL. 70, No. 1, MARCH, 1968 ad pro, meso and metapleura grey with exception of slight overlapping of yellow- orange from pronotum on upper margin of propleuron and with slight darkening along entire ventral midline; abdomen bright orange with yellow border along narrow edge of connexivum, also with 2 black spots on each segment of abdomen, 1 on the connexival border and 1 extending dorsally from the ventral midline. Ostiolar peritreme, antennae, legs, genital area and labium black. Eyes large, occupying most of lateral surface of head, a prominent orange fascia extending from base of vertex as a narrow parallel sided band, branching anterior to eyes and extending laterally to include antenniferous tubercles; tylus with short white pilosity; ocelli located in area between eye and narrow median orange band. Head length 2.32 mm; width across eyes, 3.04 mm; interocular space, 2.40 mm. Clavus with median black spot not attaining outer margin but merging with inner margin of clavus and extending to apex along inner margin, spot then meets outer margin of corium at the level of the claval apex. Scutellum with T-shaped carinae; length, 1.40 mm; width, 2.14 mm. Membrane dark brown suffusing to black, veins concolorous with background of membrane. In addition, the membrane has 2 discal white spots in the center at the level of the apices of the corium; membranal margin white. Antennae relatively large and moderately thick, segments III and IV fusiform, labium elongate attaining metacoxae; length of antennal segments: I, .92 mm; II, 2.08 mm; III, 1.72 mm; IV, 2.24 mm. Holotype: male; Contumefa, Peru-2700 M II-1950 (Weyrauch): USNM 69066. Paratypes, same data as holotype: 1 male, J. A. Slater Collection, University of Connecticut; 1 female, USNM. Close geographical and morphological relationship exists between Lygaeus slateri and Lygaeus inaequalis Walker 1872, (fig. 4). Exter- nally the latter species lacks a pair of white discal spots which are quite conspicuous on the membrane of slateri (fig. 3). However, the paramere structure of these two species presents the most reliable diagnostic characteristic for identification. The major difference is in the angle of the dorsal projection from the shank which in slateri is more acute (fig. 1). In the arm of the paramere in inaequalis there is also a notch on the dorsal surface which is absent in slateri (fig. 2). I have named this species for Dr. James A. Slater for sharing his knowledge of the Lygaeidae with me and also for extending both encouragement and criticism during this research. REFERENCES Ashlock, P. D. 1957. An investigation of the taxonomic value of the phallus in the Lygaeidae (Hemiptera-Heteroptera). Ann. Ent. Soc. Amer. 50(4):407—426. Walker, F. 1872. Catalogue of the specimens of Hemiptera-Heteroptera in the collection of the British Museum. London, Pt. 5: p. 49. 78 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 A NEW SPECIES OF PTEROMICRA AND OF EUTHYCERA FROM WESTERN NORTH AMERICA (DrererA: SCIOMYZIDAE)+ L. V. Knutson? and JAN Zusxa?® The species described here were discovered among material loaned to us by G. E. Shewell, Entomology Research Institute, Ottawa, Canada. Additional material was provided by B. A. Foote, Kent State University, Kent, Ohio. Pteromicra rudis, n. sp. (Figs. 1-4) Male and female. Body length, 2.08 to 2.72 mm. All bristles and hairs (except aristal and squamal) black. Head slightly higher than long. Frons somewhat narrowed and much lighter anteriorly; slightly wider than long, shiny, yellowish brown, with scattered hairs. Orbital plates and ocellar triangle dark brown to black. Cheeks broad, about 4% height of eye, with several strong hairs, especially on ridge above oral margin. Face and cheeks yellow, with variable amount of silver to yellowish pruinosity. Occiput shiny black, slightly pruinose; area between occipital grooves and circumcervical area with denser silver pruinosity. One pair of ocellar bristles; 2 pairs of frontorbitals, anterior pair only slightly shorter than posterior pair; 1 pair of inner and 1 pair of outer vertical bristles; 1 pair of divergent postverticals. First and second antennal segments yellowish to brownish; third segment yellowish basally, brownish apically. Basal 14 of arista thickened, somewhat lighter than dark brownish apical %; pilosity shorter than width of arista at base. Blackish orbito-antennal spot present. Proboscis black; labellae large, yellow; palpi yellow, with 1 or 2 long subapical bristles and several hairs. Thorax black, faintly pruinose. Propleural bristle strong. Humeral bristle usually small; posthumeral large; anterior notopleural well developed; posterior notopleural Y% to % as large; 1 supraalar; 1 postalar; no presutural dorsocentrals; posterior postsutural dorsocentrals well developed, anterior pair 1% as large, some specimens with 1 or 2 pairs of large, subequal hairs before anterior dorsocentral bristles. Basal and apical scutellars well developed, apicals twice as large as basals. Meso- pleuron bare, with dense yellowish or silver pruinosity around spiracle and remain- ing surface slightly pruinose (Colorado specimens) or very shiny (Montana speci- mens); pteropleuron with 2 or 3 weak to strong bristles; sternopleuron with many fine hairs. Prosternum yellowish white, densely pruinose. Fore coxae elongate, silvery, pruinose, mesially with scattered hairs, anteriorly with 1 bristle at or just below middle and 1 subapical bristle; middle and hind coxae black with silver pruinosity; hind coxae bare posteromesially. Fore femora 1 Research supported by National Science Foundation Grants GB-2415 and GB-5452. 2 Department of Entomology and Limnology, Cornell University, Ithaca, N. Y. Present address: Systematic Entomology Lab., USDA, c/o U.S. National Museum, Washington, D. C. 20560. 3 Central Research Institute of Food Industry, Prague, Czechoslovakia. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MaRcH, 1968 79 Figs. 1-4, Pteromicra rudis, n. sp.: 1, ¢ terminalia, ventral view, surstyli in natural position; 2, ¢ terminalia, caudal view, surstyli extended; 3, ¢ terminalia, lateral view of left side, surstyli slightly more extended than in fig. 2; 4, wing. 80 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 black, without pecten but with several strong hairs ventromesially at apex; middle and hind femora black, sometimes brownish apically; middle femora with strong bristle beyond middle on anterior surface. Fore tibiae black; middle and hind tibiae yellowish to brownish, sometimes darkened apically; each tibia with 1 strong, dorsal preapical bristle. Fore tarsi black; apical half of last segment yellowish; middle and hind tarsi yellowish, last 2 segments slightly darker. Wings (fig. 4) 1.92 to 2.28 mm long, mostly darkened, lighter basally and apically. First vein ending above or slightly before anterior crossvein; anal vein extending to margin, weak apically. Halteres yellow, squamae dusky with brownish hairs. Abdomen black, subshiny, with sparse silver pruinosity. Female cerci yellowish. Male genitalia, figs. 1-3. Anterior surstyli strongly flexed dorsomesially, usually not visible in lateral view. Holotype —Male. About 9 mi. SW Lima, Beaverhead Co., Montana; 14-VITI-1965; B. A. Foote. In Cornell University Collection, CU type number 4410. Allotype—Female. Same data. In Cornell University Collection, CU type number 4410. Paratypes—One male, one female; Doolittle Ranch, 9800’, Mt. Evans, Colorado; 3-VIII-1961; W. R. M. Mason. One male, one female; same locality; 15-VII-1961; C. H. Mann. One male; 5 mi. SW Idaho Springs, Colorado; 8600’; 27-VII-1961; C. H. Mann. In Cana- dian National Collection, Ottawa. The habitat near Lima, Montana where B. A. Foote collected this species is located at the edge of East Creek Campground (elevation 7,200 feet) and consists of small, partially shaded, grass-sedge stands scattered among dense growths of willows along a small creek. Other Sciomyzidae occurring there were Pherbellia argyra Verb., P. schoen- herri maculata (Cress.), P. vitalis (Cress.), Dictya montana Stey., Tetanocera rotundicornis Lw., and T. spirifera Mel. (B. A. Foote, in litt.). Pteromicra rudis apparently is the only known member of the genus with a somewhat patterned wing, the other species having at most infumated veins. This tiny, black species is similar to P. anopla Steyskal, P. perissa Steyskal, and P. inermis Steyskal and most ob- viously different from the remaining species in lacking a pecten at the apex of the fore femur. The yellow palpi, entirely black fore femora, and presence of anterior surstyli distinguish P. rudis from the brownish and larger P. anopla, which has black palpi, mainly yellow fore femora and lacks anterior surstyli. Pteromicra inermis is usually much larger than P. rudis and the basal % or %4 of the fore femora is yellowish. Pteromicra perissa, a very small species known from 1 male from Colorado, has the basal segments of the fore tarsi pale whereas in P. rudis they are black; the male genitalia of these two species are quite different. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 81 Pteromicra rudis has several atypical features. The cheeks are relatively much wider (% of the height of the eye) than any other described Pteromicra and in this regard are similar to several other genera of Sciomyzini. Although the shiny nature of the body, espe- cially of the mesopleuron and pteropleuron, has been used as a key character for Pteromicra by many workers, the thoracic pleura of some specimens of P. rudis (those from Colorado ) have a distinct pruinosity. Available data indicate that the described species of Pteromicra have two postsutural dorsocentral bristles. Some specimens of P. rudis have, in addition, one or two strong hairs or weak bristles between the thoracic suture and the anterior pair of postsutural dorsocentrals. Pteromicra rudis may be included in the recent key to Nearctic species by Foote, (1959) as follows. Couplets 1-2 are unchanged. 3. Fore femora yellowish on basal one-fourth to one-third inermis Steyskal orevtemorascomplctelys blnckse= = ems 2s ae 3a 3a. Fore tarsi pale basally; wing not patterned; posterior surstylus without ITO CESS OW AT CS Tek 0 Tee A Te ee perissa Steyskal Fore tarsi black basally; wing patterned; posterior surstylus with process OnFAn CET OTM aT Cnn ee eee rudis Knutson and Zuska Couplets 4-13 are unchanged. Subsequent to the revision of Pteromicra by Steyskal (1954), five additional North American species have been described in this genus, and one of Cresson’s species has been resurrected from synonymy (Steyskal, 1956, 1957; Foote, 1959; Fisher and Orth, 1966). A total of 16 species, including the new one described above, thus have been reported from North America. Three of these (P. glabricula (Fallén), P. leucopeza (Meigen), and P. pectorosa (Hendel) have an Holarctic distribution. Three more species are known only from the Palearctic Region. Euthycera mira, n. sp. (Fig. 5) Male. Body length, 8.0 mm. All bristles and hairs (except aristal and squamal ) black. Head slightly longer than high, conspicuously protrudent anteriorly. Frons slightly convergent posteriorly, about 1.5 times as wide as distance between anterior ocellus and anterior margin, mostly dull yellow, brownish around mid- frontal stripe. Midfrontal stripe shiny, concave, acuminate anteriorly, almost reaching anterior margin of frons. Frons with several short hairs near anterior margin, a few hairs on orbital plates. Two pairs of subequal frontorbital bristles; anterior pair situated on large, velvety-black oval spots which are about as long as widest part of midfrontal stripe. One pair each of ocellars, internal verticals, and external verticals; postocellars long, divergent. Ocellar triangle shiny, dark brown. Occiput with a few rather long bristles, patch of short hairs above cervix, and cordate black spot below postocellar bristles about twice as wide as greatest width of midfrontal stripe. Orbito-antennal spot well developed. Facies, para- facies, and cheeks pale yellow; facies subshiny, with sharp carina in upper part 82 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 Fig. 5, Euthycera mira, n. sp., holotype ¢@. and sharply delimited, shiny, black, cordate spot above middle, spot with apex directed dorsally and about 0.8 of maximum width of midfrontal stripe. Epistoma distinctly protrudent anteriorly. Parafacies with a few, fine, scattered hairs and row of sparse, short hairs on anteroventral margin. Cheeks about 0.6 times height of eyes; with very short, fine, scattered hairs and with 3 large bristles along lower margin. Eyes 1.6 times as long as high, maximum diameter slightly oblique. Lunula strongly exposed, shiny, brownish reticulate. Antennae yellowish, about two-thirds as long as head including lunula. First segment short, brownish yellow, with transverse row of tiny hairs dorsally. Second segment compressed laterally, 1.4 times as long as wide, yellowish ventrally, brownish reticulate dorsally; dorsal surface with rather long hairs and several long bristles; ventral surface with long hairs; dorsal margin sharp, almost straight, ventral margin smooth and distinctly convex. Third segment 0.8 times as long and 0.8 times as wide as second, oval, yellow basally, darkened toward apex, almost black dorso-apically. Arista sub- basal, yellow, about as long as antenna, basal fifth thickened, remainder hairy and with whitish proclinate hairs about as long as width of arista at base. Pro- PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 83 boscis yellowish brown; labellae large; palpi yellow, spatulate, with a few long hairs. Thorax brownish, mostly dull and strongly pruinose. Mesonotum in poster- odorsal aspect with 2 indistinct brown vittae separated by distance half as wide as frons. Pair of brown spots posteromesially from humeral calli, on bases of posthumeral and supraalar bristles, and smaller spots around bases of other mesonotal bristles. Mesonotum densely covered with very short, strong hairs situated on small brownish spots. Indistinct prescutellar spot mesially. Scutellum brown, shiny mesially, borders pruinose; pair of velvety blackish spots on postero- ventral margin. Prosternum and pleura, except for propleura and_ sternopleura, bare with whitish pruinosity. Propleura with a few hairs on lower surface, sternopleura with bristles on lower surface and hairs on upper surface. Humeral calli, upper part of mesopleura, and greater part of pteropleura dark brown, shiny to subshiny. One prescutellar pair of acrostichal bristles; 2 pairs of postsutural dorsocentrals situated on posterior half of postsutural part; 1 pair each of humerals, posthumerals, and supraalars; 2 pairs of notopleurals and postalars; 2 pairs of long scutellars, distance between basals and apicals slightly more than distance between apicals. No pleural or vallar bristles. Legs yellowish, brownish, and black. All coxae brownish yellow, pruinose. Femora brownish yellow, apices black. Posterior femora with irregular, suberect hairs and bristles ventrally. Tibiae yellowish, apices black. Preapical bristles of all legs about as long as apical width of tibiae. Basal 2 segments of tarsi translucent white, apical half or entire third segment and 2 apical segments blackish. Claws with whitish bases and black apices. Wings 8.5 mm long, densely patterned, white spots mostly subcircular, pattern slightly darker before rs than behind rz, white spots between r; and m in 3 rows. Halteres brownish yellow. Abdomen long, slender, in dorsal aspect only slightly tapering posteriorly, densely pruinose except for large, subshiny protandrium and small, subshiny epandrium. Median stripe indistinct; lateral stripes diffuse, interrupted. Fifth tergum half as long as fourth tergum. Holotype—Male. 14 mi. SW El Salto, Durango, Mexico; 8,000’; 26-VI-1964; J. F. McAlpine. In Canadian National Collection, Ottawa, Ontario. Euthycera mira can be distinguished easily from other known species of this genus by the large, sharply delimited, shiny black spot on the face, and by the unusual coloration of the legs. The differences between this new species and Euthycera arcuata, the other North American species, are as follows: Face without black spot. Black, cordate, occipital spot absent. Aristal hairs at least twice as long as basal width of arista. Legs yellow, without dis- tinct annulations. Wings with 2 rows of white spots between veins rs fen) ie) SS es ca gee ere ee ee Euthycera arcuata (Loew) Face with large, shiny black spot. Occiput with cordate black spot. Aristal hairs as long as basal width of arista. All femora, tibiae, and tarsi black or blackish apically. Wings with 3 rows of white spots between rs and 1) 2 eee eee Ye SA aS 8 Euthycera mira Knutson and Zuska 84 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 While only two species are now known from North America, in the Palearctic Region there are at least eight species referable to this genus and several poorly known species presently included under Limnia Robineau-Desvoidy also may belong to Euthycera. The pres- ence of vallar (subalar) bristles and mesopleural hairs in Limnia serve to distinguish that genus from Euthycera. REFERENCES Fisher, T. W. and R. E. Orth. 1966. A new species of Pteromicra from Western North America and resurrection of Pteromicra pleuralis (Cresson) (Diptera: Sciomyzidae). Pan-Pac. Ent. 42:307-318. Foote, B. A. 1959. A new species of Pteromicra reared from land snails, with a key to the nearctic species of the genus (Diptera: Sciomyzidae). Proc. Ent. Soc. Wash. 61:14-16. Steyskal, G. C. 1954. The genus Pteromicra Lioy (Diptera: Sciomyzidae) with especial reference to the North American species. Papers Michigan Acad. Sci., Arts, Let. 39:257-269. - 1956. New species and taxonomic notes on the family Sciomyzidae (Diptera, Acalyptratae). Papers Michigan Acad. Sci., Arts, Let. 41:73-87. - 1957. A new species of the genus Pteromicra associated with snails. Proc. Ent. Soc. Wash. 59:271-272. FURTHER RECORDS OF APHIDS FROM PLUMMERS ISLAND, MD. (HomMorTeRA: APHIDIDAE ) An annotated list of the aphids of Plummers Island, Maryland, was published by the writer in Proc. Biol. Soc. Wash. 79:117-126, 1966. The following records were not available for inclusion in this paper. Dactynotus (Dactynotus) ambrosiae (Thomas). This common and widespread species, the brown ambrosia aphid, was represented in the list only by McAtee’s collection in 1914 on Solidago sp. A number of alatae and apterae were collected by Geo. B. Vogt, 11 Aug. 1964, on Ambrosia trifida (A. T. Olive det.). Dactynotus (Lambersius) gravicornis (Patch). Not in the previous paper. A number of alatae and apterae were collected by Geo. B. Vogt, 11 Oct. 1964, on Solidago sp. (A. T. Olive det.) Second record for Maryland, the first being College Park, Sept. 1951 (W. E. Bickley coll—L. M. Russell det.; 2 slides in USNM). Dactynotus (Lambersius) pseudambrosiae Olive. Not in the previous paper; first record for Maryland. A number of alatae and apterae were collected by Geo. B. Vogt, 27 Sept. 1964, on Lactuca biennis and 18 Oct. 1964, on L. canadensis (A. T. Olive det.). Dactynotus (Uromelan) ruralis (Hottes & Frison). Alatae and apterae collected by Geo. B. Vogt, 16 Aug. 1964, on Actinomeris alternifolia (A. T. Olive det.). Drepanaphis saccharini Smith & Dillery. This is referred to in the previous paper as “Drepanaphis near parvus Smith, probably n. sp., 25 June 1964 on suckers of silver maple” (Geo. B. Vogt coll—C. F. Smith det. ).—Mortmer D. LEONARD, 2480 16th St., Washington, D.C. 20009. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 85 THE GENUS EULABIDOCARPUS LAWRENCE (AcARINA: LISTROPHORIDAE )* B. McDAniEL, Department of Entomology—Zoology, South Dakota State University, Brookings, South Dakota 57006 The genus Eulabidocarpus Lawrence (1948) was erected for the single species Labidocarpus compressus Ewing. In that publication detailed information as to the curious features of Ewing’s L. com- pressus was given. Lawrence, however, clearly stated that all of his studies were taken from two descriptions by Ewing (1910, 1929). The illustrations of L. compressus by Ewing are extremely generalized and as Lawrence so clearly stated, the species as described and illus- trated by Ewing makes it impossible to place this species in the genus Labidocarpus Trouessart. Only by reference to the holotype, which now appears to be lost, could E. compressus (Ewing) be conclusively recognized. The author has tried unsuccessfully to secure the holotype or other material of L. compressus Ewing. E. W. Baker (personal communication) has informed the author that the holotype or other material of L. com- pressus Ewing cannot be found, either in Ewing’s type collection or in the U.S. National Museum collection. It was apparent that Lawrence (1948) in establishing the genus Eulabidocarpus for Ewing’s L. compressus was removing from Labido- carpus this poorly illustrated and described species. It is also apparent that the genus Eulabidocarpus should remain monotypic, at least until additional material could be collected from the type host and the genus redescribed from actual specimens rather than the literature. In essence the single species E. compressus (Ewing) should be re- garded as a species inquirendae—i.e. a species, the taxonomic position of which is in doubt because of difficulties in identifying the species. Unfortunately this interpretation has not been followed by the work of Pinichpongse (1963c). He has added two new species to the genus Eulabidocarpus Lawrence. The genus Eulabidocarpus is redescribed by Pinichpongse (1963c) based on the two new species E. flexipes and E. rectipes. The structures which prompted Lawrence to establish Eulabidocarpus are not utilized. In Pinichpongse’s review of a subfamily which he called Chirodiscinae,’ he (1963a, b, c, d) apparently overlooked the work of McDaniel and Lawrence (1962) in which the genus Parakosa McDaniel and Lawrence was described. ' Approved by the director of the South Dakota Agricultural Experiment Station as Journal Series No. 796. * The use of the subfamily Chirodiscine is not valid due to the findings by McDaniel (in press). 86 PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 In the revised description of Eulabidocarpus by Pinichpongse (1963c) he refers to legs III and IV as having a “large denticulated spur” in addition to spurs. This is one of the distinctive structures used to characterize the genus Parakosa. Also in the redescription of Eulabi- docarpus it is stated that “Legs III and IV distinctly separated from legs I and II.” This statement is very misleading if it was meant to utilize Lawrence’s character of the third and fourth pair of legs form- ing a separate group far removed from the modified anterior legs, and placed near the posterior end of the body. In all members of the superfamily Listrophoroidea, legs III and IV are distinctly separated from legs I and II. Dusbabek and Cruz (1966) collected E. flexipes Pinichpongse from a subspecies of the type host, Molossus major tropidorhynchus from Cuba. In this work the authors placed E. flexipes Pinichpongse in the genus Parakosa McDaniel and Lawrence by comparing their material from Molossus major tripidorhynchus with the description of Parakosa. Also Dusbabek and Cruz (1966) rightfully pointed out the difference in the host relationship of E. compressus Ewing and E. rectipes Pinichpongse. They also illustrated and described the copulatory nymph and larva of Parakosa flexipes (Pinichpongse ). Through the courtesy of Drs. E. W. Baker and R. E. Crabill, the author was able to study the holotypes, allotypes and paratypes of E. flexipes Pinichpongse and E. rectipes Pinichpongse. From this study it became clear that E. rectipes also belongs in the genus Parakosa. Parakosa rectipes ( Pinichpongse ), n. comb. Eulabidocarpus rectipes Pinichpongse, 1963, Acarologia 5(3):400. The presence of the spine (thick seta of Pinichpongse) in addition to spurs on legs III and IV, the two long, thick subequal setae inserted above legs III, and two thick setae posterior to head plate, are found on P. rectipes. The removal of Parakosa rectipes (Pinichpongse ) makes Eulabido- carpus Lawrence monotypic. Until additional material can be obtained from the type host and the genotype E. compressus redescribed, it is the author’s strong opinion that Eulabidocarpus should remain mono- typic. The genus Parakosa McDaniel and Lawrence now contains three species, P. tadarida McDaniel and Lawrence, P. flexipes (Pinich- pongse ) and P. rectipes (Pinichpongse ). All are from the Neotropical region of the Western Hemisphere and found on bats of the family Molossidae of the suborder Microchiroptera. KEY TO THE SPECIES OF Parakosa MCDANIEL AND LAWRENCE 1. Spur of tarsus IV distally curved toward claw and with pointed ape xen ee Oe eae ee ls Ee P. flexipes (Pinichpongse) PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 87 Spur of tarsus IV not distally curved toward claw and either rounded or DOlted bil JAperess4 et eee hes cont wee begs ee «yee ee coe 2 2. Spur of tarsus IV broad, with inner margin curved, and beset with IRULIST GV Seater eee 8 Be SD P. rectipes (Pinichpongse ) Spur of tarsus IV slender, setae-like, and straight, without PUTTER WS eee er an a P. tadarida McDaniel and Lawrence REFERENCES Dusbabek, F. and J. De La Cruz. 1966. Nuevos Generos y Especies de Acares (Acarina: Listrophoridae ) Parasitos de Murcelagos Cubanes. Poeyana 31:1—20. Ewing, H. E. 1910. New Acarina from India. Trans. Acad. Sci., St. Louis 19: 113-121. 1929. A manual of external parasites. Baltimore, 225 pp. Lawrence, R. F. 1948. Studies of some parasitic mites from Canada and South Africa. Jour. Parasitol. 34:364—-379. McDaniel, B. and R. F. Lawrence. 1962. A new genus and species of parasitic mites from Mexico (Acarina, Listrophoridae). Jour. Parasitol. 48:463-466. McDaniel, B. (in press) The superfamily Listrophoroidea and establishment of some new families (Listrophoroidea: Acarina). Acarologia. Pinichpongse, S. 1963a. A review of the Chirodiscinae with descriptions of new taxa (Acarina: Listrophoridae). (1st series). Acarologia 5(1):81-91. 1963b. 90 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 Figs. 5-7. Baldulus tripsaci, n. sp. Fig. 5, right forewing; fig. 6, head, prono- tum, and scutellum in dorsal view; fig. 7, fifth instar nymph in dorsal view. land, 20 September 1967, J. P. Kramer and R. F. Whitcomb; 1 male and 5 females, Churchton, Maryland, 23 September 1967, R. F. Whit- comb; 4 males, 18 females, and 2 nymphs, Benton (Lowndes County ), Alabama, 12 October 1967, H. B. Cunningham and T. D. Eichlin. Entire type series collected on Tripsacum dactyloides, eastern gama- grass. Notes: The species is named for its host grass, Tripsacum dacty- loides. The name “tripsaci” is the genitive singular of Tripsacum. Baldulus tripsaci can be distinguished easily from all its congeners PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 91 tN Fig. 8. Distribution of eastern gamagrass, Tripsacum dactyloides, in the United States (after Hitchcock). by the fact that it lacks black or brown spots on the crown and brown stripes on the pronotum. The male genital structures are closest to those of the Mexican Baldulus bilineatus DeLong (DeLong 1950: plate III). The styles of B. tripsaci and B. bilineatus are nearly iden- tical, but the stalk of the connective is very much longer in B. tripsaci. The aedeagus of B. tripsaci is clearly U-shaped with the proximal pair of apical processes much shorter than the distal pair; the aedeagus of B. bilineatus is scarcely or not U-shaped, and the two pairs of apical processes are subequal in length. Because the connective is unusually long, the distal portion of the aedeagus often protrudes dorsally from the genital capsule in B. tripsaci. The aspect of the aedeagus shown in fig. 1 can be observed frequently by viewing the genital capsule from above without resorting to the usual clearing procedures. The fifth instar nymph (fig. 7), length 2.5-3.0 mm, resembles the adult but has a pair of irregular black spots dorsally on each of the last two abdominal segments and paired setae on all abdominal segments. Another leafhopper, Amplicephalus lawsoni (DeLong), was also found to breed abundantly on eastern gamagrass in Maryland. This species had been recorded previously from Kansas and Mississippi, but its host grass was unknown. A. lawsoni and B. tripsaci coexist on the same host grass, and the two species were often collected together. The infrequency with which A. lawsoni had been collected in the past and the discovery of B. tripsaci can be explained in part by a high 92 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 host plant specificity. Gamagrass is difficult to sweep, at least at the stage when the grass can be positively identified; and even though its distribution is extensive, gamagrass is not often common (Hitch- cock 1935: 767). Both leafhoppers belong to the exceedingly large subfamily Deltocephalinae, but they are in distinctly different tribes. A. lawsoni is a conspicuous, robust, distinctly marked, brown leaf- hopper in the tribe Deltocephalini; B. tripsaci is an inconspicuous, slender, fragile, unmarked yellowish leafhopper in the tribe Mac- rostelini. With the unaided eye alone, B. tripsaci could be mistaken easily for a typhlocybine leafhopper. PROBABLE DISTRIBUTION AND ECONOMIC IMPLICATIONS The distribution of the grass Tripsacum dactyloides (fig. 8) includes most of eastern United States westward to the eastern portions of the Great Plain states south of the Dakotas. This indicates that B. tripsaci will eventually prove to have a much wider distribution than is presently recorded and may occur wherever Tripsacum dactyloides is found. Eastern gamagrass, Tripsacum dactyloides, and corn, Zea mays, are known to be closely related because these two grasses hybridize (Mangelsdorf and Reeves 1931). Also, Dalbulus maidis (DeLong and Wolcott), whose host plant is corn, is closely related to B. tripsaci and is the major vector of corn stunt virus in many parts of the southern United States (Stoner 1965). However, some leafhoppers are known to change hosts: Whitcomb (1957) reported three species transferred from Elymus grasses to quackgrass, Agropyron repens, in the midwest. Thus on the basis of the close relationship between D. maidis and B. tripsaci and their host grasses and with the precedent for a change in host plants by other leafhoppers, B. tripsaci may eventually be implicated, either directly or indirectly, in the corn stunt virus problem. Tripsacum dactyloides could well harbor a virus that is intraspecif- ically transmitted by B. tripsaci and that may be transmitted inter- specifically to corn. REFERENCES DeLong, D. M. 1950. The genera Baldulus and Dalbulus in North America in- cluding Mexico (Homoptera: Cicadellidae). Bull. Brooklyn Ent. Soc. 45(4): 105-116. Hitcheock, A. S. 1935. Manual of the grasses of the United States. U.S. Dept. Agr. Misc. Publ. 200:1—1040. Mangelsdorf, P. C. and R. G. Reeves. 1931. Hybridization of maize, Trip- sacum, and Euchlaena. Jour. Heredity 22( 11) :328-343. Stoner, W. N. 1965. A review of corn stunt disease (Achaparramiento) and its insect vectors, with résumés of other virus diseases of maize. U.S. Dept. Agr., ARS 33-99: 1-35. Whitcomb, R. F. 1957. Host relationships of some grass-inhabiting leafhoppers. Unpublished Thesis, Univ. Illinois, Urbana. PROC. ENT. SOC. WASH., VOL. 70, No. 1, MARCH, 1968 93 SUMMARY REPORTS OF SOCIETY OFFICERS FOR 1966' CORRESPONDING SECRETARY (For the fiscal year 1 November 1965 to 31 October 1966) Membership on 1 November 1965 _ 484 Reductions: Resioneds 3252.4. 2 ET ABO ee 2 19 ID FOp pe cape Raa a he ho 6 | DIETCrSe cro UPR see en a cite des ends obit ee 4 PING bay) LES ere ks Ne EL eh EE EYOTA 0! 29 Increases: Elected to, Membership = .2 25 REMIStALCGL Sx ® eK. Seto Ps OVER 2 eo 2 (CLE) frees ees eee EEO esc: ee Oe ae See 27 Membership on 31 October 1966 ROL Pope a Sah A x Met ee 8 nek St 492, Classes of Membership: LD AaResst <1 Ab Ys bao ke Meena AB LEO Ye ee ee Oe 468 LLELYe 22 SiS SER, Seen Saas Aa A es IO ae eee 5 PETE Chapt teak viele) tee 6 Ue fem Ys oe tse 2S ey hr Saat ls ET ONO La yy ge ete Week een Sea ie ee Fabel ee eed 4 TNE Se SS Alea ea oon RL MEE «a ca Se ae 492 The membership is distributed among 47 states, the District of Columbia, 2 ter- ritories, and 22 foreign countries. Circulation of the Proceedings (September 1966 issue ) : State Senne eee 2 Re) ee ee ee 527 Distictuos (Columbia. st. 91 Tee EOSSESSIO TIS 0 ee Se 5 10 HOreionn Countricsee.c- eo eee ee 178 AIRC) tet Re Jt A hn res ES A I eel Re 806 Distribution of the Proceedings (September 1966 issue) : SOMIMC METS ieee ee A474 MOWSUDSCrIDENS hese. ee ee 8 332 TOE axe kop a RA an ies tei Pepe a OR as ee ey 806 The Proceedings go to members and subscribers in 50 states, the District of Columbia, 2 territories, and 51 foreign countries. D. M. ANDERSON, Corresponding Secretary. TREASURER (For the period 1 November 1965 to 31 October 1966) General Publications Total Fund Fund Fund Cash on Hand November 1, 1965 $1,126.02 $10,492.12 $11,618.75 Maries PReCetpisy cele a Ly es aie PN Leeks 6,493.63 708.67 7,202.30 Wigtalltee Bots 2... tien lawn e 2S $7,620.26 $11,200.79 $18,821.05 Minus Expenditures _......- 6,018.74 6,018.74 Cash on Hand October 31, 1966 $1,601.52 $11,200.79 $12,802.31 Respectifully submitted, ARTHUR K. BuroitT, Jr., Treasurer. 1Due to the loss of these reports in shipment to the printer, publication was withheld until now.—Ed. 94 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 CUSTODIAN (For the period 1 November 1965 to 31 October 1966) The value of stock sold by the Custodian’s office amounted to $1,352.20. Of these items, $170.20 was for 25 copies of the Memoirs, $4.00 for 2 copies of the Weld volumes, $678.00 for complete volumes and miscellaneous numbers of the Proceedings, and $500.00 for miscellaneous reprints. Sales of the Memoirs were: No. 1, 1 copy; No. 2, 2 copies; No. 3, 6 copies; No. 4, 9 copies; No. 5, 7 copies. A copy of the complete, detailed report is on file with the Recording Secretary. Respectfully submitted, Ropert L. SMmry, Custodian. EDITOR (For the calendar year 1966) Four numbers of the Proceedings were published in 1966. Of the pages pub- lished, four were devoted to advertising and 338 to scientific papers, notes, obituaries, book reviews, minutes of meetings, and announcements. Eighty-seven scientific papers and notes were published during the year. The Society and the Proceedings benefitted from five paid papers of 32 pages. This did not cause the articles of regular contributors to be postponed. Respectfully submitted, Jon L. Herrine, Editor. PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 95 SOCIETY MEETINGS 752nd Regular Meeting—May 4, 1967 The meeting was called to order by President Louis G. Davis at 8 p.m. at the regular meeting place, Room 43, United States National Museum. Twenty-two members and eighteen visitors were present. Minutes of the 75lst were approved as read. Three applications for membership were presented: Robert Hamman, Norman Lin and Elliot Krafsur. President Davis announced that Dr. Frank Favorite will speak at the Joint Dinner on June 1. Visitor David Gilsey announced the National Audubon camp in a nearby state during the summer. Under notes and exhibition of specimens, T. L. Bissell mentioned the collecting of two species of aphids, apparently common, on barley and wheat. L. G. Davis showed a piece of wood damaged by Coptotermes formosanus, the Formosan termite. He said this potentially serious pest is now established in two localities in the State of Louisiana. In Hawaii it is especially injurious to utility poles. As part of the program three high school students presented their science fair exhibits with which they had won awards in their respective areas. The students and their subjects were: Patrick Hughes, LaPlata, Maryland, “Relationships in Entomology”; George Roland, Brandywine, Maryland, “A 2-D Chromatography Study of Drosophila Pteridine Metabolism”; and Bruce C. Black, Falls Church, Virginia, “Effects of Mono, Di and Tri Saccharides on the Activities of Honey Bees.” Each explained his subject in well-prepared detail. The principal speaker of the evening was Dr. Adolph Krebs trained at Univer- sities of Heidelberg, Bonn and Frankfurt a. Main, and presently at the Walter Reed Army Institute of Research, Division of Nuclear Medicine. Dr. Krebs’ subject, “Ionizing Radiation and Ants,” was illustrated by a motion picture and slides. Interesting comparisons were made between ants and other organisms as to their reactions to radiation. Respectively submitted, THEo L. Bisset, Acting Recording Secretary. 753rd Regular Meeting—June 1, 1967 The 753rd meeting of the Society was held jointly with the Insecticide Society of Washington in the Fort McHenry Room of the Adult Education Center, University of Maryland, College Park. About 180 persons were present. After a delicious dinner served by University students, members and their friends were entertained by a group of four young men who performed “Eine Kleine Nacht- musik.” The principal speaker of the evening, Dr. Frank G. Favorite, Space Science Board, NAS-NRC, presented the fascinating story of space biology studies carried on by United States scientists. The meeting adjourned at 9:45 p.m. Respectfully submitted, RicHarp H. Foote. 754th Regular Meeting—October 5, 1967 The 754th regular meeting of the Society was called to order by the President, Mr. Louis G. Davis, on October 5, 1967 at 8:00 p.m. in room 43, U.S. National Museum. Twenty-four members and seven visitors were in attendance. Minutes of the previous meeting were approved as read. Elliot S. Krafsur, Robert E. Hammon, and Norman Lin were received into the Society. The names of Maynard 96 PROC. ENT. SOC. WASH., VOL. 70, NO. 1, MARCH, 1968 J. Ramsey, Cary J. Hansel, William B. Steltzfus, Alfredo D’Ascoli, and Donald R. Roberts were read for the first time as candidates for membership. Dr. F. M. Wadley exhibited his new book, “Experimental Statistics in Ento- mology,” published by the Graduate School Press, U. S. Department of Agriculture in 1967. Dr. R. I. Sailer discussed experiences collecting coccinellids along the sea shore at Rehobeth Beach, Delaware. Dr. R. A. St. George reported that an infestation of the Formosan termite had recently been discovered in Eastern South Carolina. The principal speaker of the evening, Dr. A. B. Park, delighted the audience with a well illustrated discussion of Stress Detection. Any stress exerted on a plant species results in modifications of its normal wavelength signature which can be detected by aereal surveillance. Some of the broad applications of these new techniques were discussed. Following the introduction of visitors, the meeting was adjourned by President Davis at 9:40 p.m. Respectifully submitted, RaLpH A. Bram, Recording Secretary. 755th Regular Meeting—November 2, 1967 The 755th regular meeting of the Society was called to order by President Louis G. Davis on November 2, 1967 at 8:00 p.m. in room 43, U.S. National Museum. Twenty-four members and eleven guests were in attendance. Minutes of the previous meeting were approved as read. Maynard J. Ramsay, Cary J. Hansel, William B. Steltzfus, Alfredo D’Ascoli, and Donald R. Roberts were received into the Society. The names of Richard A. Newkirk, F. E. Wood, and Sueo Nakahara were read for the first time as candidates for membership. Mr. R. H. Nelson announced the XIII International Congress of Entomology to be held in Moscow, U.S. S.R from 2-9 August 1968. Dr. William E. Bickley, Chairman of the Nominating Committee, presented the following slate of candi- dates for 1968: President, Richard H. Foote; President-elect, Helen Sollers-Riedel; Recording Secretary, Ralph A. Bram; Corresponding Secretary, David R. Smith; Treasurer, Arthur K. Burditt, Jr.; Editor, Paul M. Marsh; Custodian, Robert L. Smiley; Program Committee Chairman, Richard G. Oakley; Membership Com- mittee Chairman, William B. Hull. Dr. R. H. Foote reviewed “The Dictionary of the Biological Sciences” by Peter Gray and published in 1967 by Reinhold Press. Mr. T. L. Bissell reported that in July 1967 a one-acre spot in a 25-acre field of soybeans at Carmichael, Queen Anne’s County, Maryland was killed by feeding of adult weevils, Colomycterus setarius Roelfs. The spot adjoined a field of maturing wheat from which the insects may have come. This species was first found in Maryland in Baltimore County in 1935. It sometimes appears in large numbers and damages ornamental plants both herbaceous and woody or invades dwellings. This is the first record we have in the State of damage to a field crop. President Davis exhibited the Yearbook of Agriculture for 1967, “Outdoors U.S. A.” The principal speaker for the evening was Mr. Richard Oliver of the National Academy of Science, Division of Biology and Agriculture. During his discussion of the International Biological Program, Mr. Oliver outlined the broad objectives and structure of the Program and encouraged participation by members of the Society. Following the introduction of visitors, the meeting was adjourned by President Davis at 9:35. Respectfully submitted, RatpH A. Bram, Recording Secretary. PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. Manuscripts for publication, proof and other editorial matters should be addressed to the Editor (for address, see inside front cover of this issue). Typing—All manuscripts must be typed on bond paper with double-spacing and ample margins. Carbon copies or copies on paper larger than 81% x 11 inches are not acceptable. Do not use all capitals for any purpose. Underscore cnly where italics are intended in the body of the text, not in headings. Number all pages consecutively. References to footnotes in the text should be numbered consecutively and typed on a separate sheet. First page—The page preceding the text of the manuscript should include (1) the complete title, (2) the order and family in parentheses, (3) the author’s name or names, (4) the institution with city, state and zip code or the author's home city, state and zip code if not affiliated, (5) in the upper left hand corner, the complete name and address to which proof is to be sent. Names and descriptions of organisms—The first mention of a plant or animal should include the full scientific name with the author of a zoological name not abbreviated. Descriptions of taxa should be in telegraphic style. References—Citations in the text of papers longer than one printed page should be by author and date and should refer to a list of concluding REFERENCEs listed alphabetically in the following format: Muesebeck, C. F. W. 1963. Host relationships of the Euphorini (Hyme- noptera: Braconidae). Proc. Ent. Soc. Wash. 65(4): 306. and L. M. Walkley. 1951. in Muesebeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2: 90-184. In shorter articles, references to literature should be included in parentheses in the text. Tables—Tables are expensive and should be kept to an absolute minimum. In most cases, material in tables can be incorporated into the text. When tables are necessary, each table should be prepared as a line drawing or typed on a separate page with heading at top and footnotes below. Number footnotes consecutively for each table. Use only horizontal rules. Illustrations—No extra charge is made for line drawings or halftones. Authors must plan their illustrations for reduction to the dimensions of the printed page and the individual figures must be mounted on suitable board. Proportions of full-page illustrations should closely approximate 454g x 6” (26 36 picas); this usually allows explanatory matter to appear on the same page. On the back of each illustration should be stated (1) the title of the paper, (2) the author’s complete name and address, and (3) the number of the illustration such as “No. 1 (of 3)” ete. Figures should be numbered consecutively. Figure legends—Legends should be typewritten double-spaced on separate pages headed ExpLANATION OF FicurREs and placed following REFERENCES. Do not attach legends to illustrations. Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) KRAMER, J. P. and R. F. WHITCOMB—A new species of Baldulus from gamagrass in eastern United States with its possible implications in the corm stunt virus problem (Homoptera: Cicadellidae: Deltocephalinae) —_ LEONARD, M. D.—Further records of aphids from Plummers Island, Md. (Homoptera: * Aphididae) +... LIN, N.—A note on the sleeping behavior of Philanthus gibbosus (Fabricius ) (itymenopteras (Sphecidaé) 2 McDANIEL, B.—The genus Eulabidocarpus Lawrence (Acarina: Listrophor- POLHEMUS, J. T.—A report on the Saldidae collected by the Galapagos International Scientific Project 1964 (Hemiptera) — SLATER, J. A. and D. B. WILCOX—New genera and species of Neotropical Blissinae (Temiptera: Liygacidae) 0.0000 ee SMILEY, R. L.—A new genus and three new species of Erythraeoidea (Acaring» Exythracidae and Smarididae) —......._....... . . - SMITH, C. F. and J. O. PEPPER—Grylloprociphilus frosti, new genus, new species, from the eastern United States (Homoptera: Aphididae) BF STEYSKAL, G. C.—Notes on North American Piophilidae. III (Diptera) __ 25 STEYSKAL, G. C.—Priority of the generic names Tonnoiria Malloch and Toennoita Farent .( Diptera)’ ssc. ZIMMERMAN, E. C.—The genus Microplatymerus of the Seychelles (Cole- optera::' Cuveulionidae: Cossoninae) -...-. 1 27 SUMMARY REPORT OF SOCIETY OFFICERS SOGIES DIFETINGS - 0 ae ZEITHSO NS. MAY 7 1968 _ ZJppaniiS — Et tT Vol. 70 JUNE 1968 No. 2 hax... PROCEEDINGS of the EN'TOMOLOGICAL SOCIETY a WASHINGTON U.S. NATIONAL MUSEUM WASHINGTON, D.C. 20560 PUBLISHED QUARTERLY CONTENTS BURKS, B. D.—New North American species of Ceratosmicra Ashmead Baromenopterss ‘Ghaleididac) ....- 170 EMERSON, K. C.—The host of Stachiella retusa martis Werneck (Mal- Pe eolimienMoneetdae)), 22.20 PS fee 191 EMERSON, K. C. and R. D. PRICE—A new species of Rhynonirmus from fuatend (Mallophasa: Philopteridae) 00.0 184 FISK, F. W. and A. B. GURNEY—Neotropical cockroaches of the genus Xestoblatta: a new species from Costa Rica and notes on other species (Dictyoptera: Blattaria: Blattellidae) Atte eS ee kd 137 FROESCHNER, R. C.—Telamona archboldi, a new treehopper from Florida Miaueritera’, wiembracidae) 2242 154 FROESCHNER, R. C.—Burrower bugs from the Galapagos Islands collected by the 1964 expedition of the Galapagos Scientific Project (Hemiptera: LDA E) 0 alee MEET CAS Se ae es a veers Ieeeee - annette h 192 GAGNE, R. J.—Authorship of Pachydiplosis oryzae, a rice pest of Asia and ameter Diptera, Cecidompirdae ) 2-05 eh 136 GENTRY, J. W. and E. J. GERBERG—Dipterous eggs as food for adult Coccinellidae (Coleoptera: Coccinellidae) _......------ 142 GJULLIN, C. M., L. F. LEWIS and D. M. CHRISTENSON—Notes on the taxonomic characters and distribution of Aedes aloponotum Dyar and Aedes communis (De Geer) (Diptera: Culicidae) 133 (Continued on back cover) ENTOMOLOGICAL SOCIETY OF WASHINGTON OrGANIZED Marcu 12, 1884 OFFICERS FOR 1968 Dr. Ricwarp H. Foore, President Dr. Paut M. Manrsn, Editor Ent. Res. Div., ARS, USDA Systematic Entomology Laboratory, USDA Plant Industry Station c/o U.S. National Museum Beltsville, Maryland 20705 Washington, D. C. 20560 Mrs. HELLEN SOLLER-RIEDEL, President-Elect Mr. Roserr L. SMILEY, Custodian Agr. Res. Ser., USDA Ent. Res. Div., ARS, USDA Room 466, Federal Center Bldg. Room 3139, South Bldg. Hyattsville, Maryland 20782 Washington, D. C. 20250 Dr. Ratpeu A. Bram, Recording Secretary Mr. Ricuanrp G. OAKLEy, Program Committee Ent. Res. Div., ARS, USDA Chairman Plant Industry Station 1914 Franklin Ave. Beltsville, Maryland 20705 McLean, Virginia 22101 Dr. Davw R. Smiru, Corresponding Secretary Carr. WiLL1AM B. Hutt, Membership Committee — Systematic Entomology Laboratory, USDA Chairman c/o U.S. National Museum 746 Beall Ave. Washington, D. C. 20560 Rockville, Maryland 20850 Dr. AntTHUR K. Burpirt, JRr., Treasurer Ent. Res. Div., ARS, USDA Plant Industry Station ; Beltsville, Maryland 20705 ‘" Honorary President i Dr. T. E. Snyper, U.S. National Museum Honorary Members Mr. C. F. W. MuEsEseEck, U.S. National Museum Dr. F. W. Poos Dr. E. N. Cory MEETINGS.—Regular meetings of the Society are held in Room 43 of the U.S. National Museum on the firs Thursday of each month from October to June, inclusive, at 8 P.M. Minutes of meetings are published regular in the Proceedings. f MEMBERSHIP.—Members shall be persons who have demonstrated interest in the science of entomology Annual dues for members are $5.00; initiation fee is $1.00 (U.S. currency). fh PROCEEDINGS.—Published quarterly beginning with March by the Society at Washington, D.C. Members it good standing are entitled to the Proceedings free of charge. Nonmember subscriptions are $8.00 per year both domestic and foreign (U.S. currency), payable in advance. All remittances should be made payable to The Entomological Society of Washington. The Society does not exchange its publications for those of other societies. STATEMENT OF OWNERSHIP Title of Publication: Proceedings of the Entomological Society of Washington. Frequency of Issue: Quarterly (March, June, September, December). i Location of Office of Publication, Business Office of Publisher and Owner: The Entomological Society ol Washington, c/o Department of Entomology, U.S. National Museum, Washington, D.C. 20560. Editor: Dr. Paul M. Marsh, same address as above. ; Managing Editor and Known Bondholders or other Security Holders: none. This issue mailed June 29, 1968 Second Class Postage Paid at Lawrence, Kansas, U. S. A. 66044 ALLEN PRESS, INC. ee reo LAWRENCE, KANSAS 66044 U.S, Be PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Vol. 70 JUNE 1968 No. 2 STUDIES ON IDIOCERINAE LEAFHOPPERS: IV. A NEW SPECIES OF BALOCHA AND ONE OF PEDIOSCOPUS, MIMICS J. Mauponapo-Caprites, Department of Biology, University of Puerto Rico, Cayey Regional College, Cayey, Puerto Rico 00633 A new species of Balocha Baker and one of Pedioscopus Kirkaldy are herein described. The generic position of the latter is somewhat tentative; male specimens are needed for a definite decision. Both species were collected by C. F. Baker in Bormeo. Unfortunately there is no date or other information on the labels, except Sandakan, that would indicate that they were collected at the same time and place. The coloration of both species is quite similar to that of those species of Balocha that are striped with orange and yellow. This meager information suggests that the new species of Pedioscopus is a mimic of the new species of Balocha. The types are deposited in the United States National Museum and paratypes in my collection. In the measurements that follow, 10 micrometer units are equivalent to 0.05 mm. Pedioscopus balochoides, n. sp. (Figs. 1-3) Female—general coloration orange and yellow. Head yellowish with an orange transverse band on disk of vertex. Pronotum on anterior half yellowish, posterior half orange. Scutellum bright yellow. Thorax ventrally and laterally, legs, abdomen, and genital sclerites, except for brown tip of ovipositor, yellowish. Clavus orange on inner half, yellowish on outer half. Corium with a pale orange tinge, changing into brownish on apical cells; costal margin pale orange; veins on corium concolorous, barely visible, slightly brownish and better defined near apex of wing. Ocelli black. Vertex parallel-sided, shagreen, less than 4% as long as pronotum (4:10); pronotum about % as long as wide (10:28). Scutellum wider than long (21:15). Face as in fig. 1. Forewing as in fig. 2. Length 4.1 mm. Female genitalia—hind margin of last abdominal sternum angularly produced caudad and with truncate apex (fig. 3). Ovipositor slightly surpassing pygofers, without spines; pygofers smooth. Holotype, female, USNM 69531; paratypes, one female each in the USNM and in my collection; all from Sandakan, Borneo, C. F. Baker collector. 98 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Figs. 1-3, Pedioscopus balochoides, n. sp.: 1, head, frontal view; 2, forewing; 3, last abdominal segments, 9. Figs. 4-11, Balocha bicolor, n. sp.: 4, head, frontal view; 5, apex of forewing; 6, aedeagus, lateral view; 7, aedeagus, caudal view; 8, style, lateral view; 9, apex of style, dorsal view; 10, pygofer and valve, lateral view; 11, last abdominal segments, 2. Figs. 12-14, B. bicolor uniformis, n. ssp.: 12, apex of forewing; 13, last abdominal segments, 2; 14, aedeagus, lateral view. The venation of the forewing and the characters of the face in- dicate that this species is a Pedioscopus. It is the only known species of this genus that is banded with orange and yellow. Most of the species in this genus are conspicuously banded or spotted with black. PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 99 Others not so marked, like P. modestus Baker and P. maquilingensis Baker from the Philippine Islands, are ochraceous and definitely not banded with orange. Balocha bicolor, n. sp. (Figs. 4-11) Male—overall color orange and yellow. Vertex yellowish, the band on face barely visible from above; face pale orange, with an orange band from eye to eye above ocelli. Pronotum under margin of head yellow, anterior half orange, posterior half yellowish-orange or yellowish. Clavus with inner longi- tudinal half orange, outer half whitish-yellow. Corium contiguous to clavus orange, remainder translucent and more or less tinged with pale orange and turning into brownish on apical cells; costal margin narrowly pale orange; veins somewhat conspicuous, slightly deeper orange. Thorax ventrally and laterally, abdomen, legs, and genital capsule pale orange or slightly yellowish. Brown spot on forked cell extending to contiguous outer apical cell. Vertex very finely striate, about % as long as pronotum (3.2: 10); pronotum % as long as wide (10:25), posterior margin very slightly concave above scutellum. Scutellum wider than long (17:12). Face (fig. 4) slightly wider across eyes than long (27:25); lateral margin of frons (or postclypeus) roundly angled, lateral frontal suture not reaching ocellus, pointed well above ocellus of other side; clypeus (or anteclypeus) expanded apically, bell-shaped, as long as wide apically (5:5). Forked cell with peduncle shorter than cell (4:7) or slightly longer (7: 6), as in fig. 5. Length 4.1 mm. Genitalia typical of the genus, as in fig. 6-10. Female—coloration and shape much as in male; forked cell of forewing with peduncle slightly shorter (5:6) or longer (6:5) than cell; blackish spot as in male. Genitalia: last abdominal sternum rectangular, hind margin slightly convex, as in fig. 11. Holotype, male, USNM 69532; allotype female, in the USNM; six paratypes in the USNM, two in my collection; all from Sandakan, Borneo, C. F. Baker collector. This species is quite close to Balocha lucida Maldonado described from Sarawak. They can be separated, besides the characters in the short key below, by many small details of the aedeagi, different apex of the styles, not so bright orange in bicolor, etc. In my key to the studied species of the genus (1961), B. bicolor runs to the second part of couplet 3 and can be separated from B. lucida as follows: Peduncle three times as long as pedunculate cell; apical half of aedeagus thicker, with posterior spines arising from apex; ovipositor shiny brown “2 SoG! Sas Sees, A ee oe ee ee ee ee B. lucida Peduncle shorter, about as long as pedunculate cell; apical half of aedeagus thinner, with posterior spines arising below apex; ovipositor dull yellow ee Lee eaee Swe: YEN. ty) Ry A ee Fe ee ed J Se B. bicolor 100 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Balocha bicolor uniformis, n. ssp. (Figs. 12-14) Six specimens, from the same locality and perhaps collected at the same time, show slight differences in coloration of the pronotum, shape of the pedunculate cell, and genitalic characters. Although these characters seem to be constant there is intergrading between the extremes of this form and some specimens of the typical form, so I am calling these a separate subspecies. Field or laboratory observations would clarify the real rank of this group. General shape and coloration much as in the typical form; with the following differences: pronotum uniformly (hence the name) orange, instead of banded; pedicel of pedunculate cell equal in length to cell in most cases, slightly shorter or longer in some (fig. 12), spot of brown near apex of wing extending slightly both sides of pedunculate cell in most specimens. Female genitalia slightly different, as in fig. 13, last sternum rectangular, posterior margin slightly convex. Aedeagus of male more enlarged on apical half (fig. 14). Holotype, male, 69533. Allotype, female, in the USNM; three paratypes in the USNM, two in my collection. All from Sandakan, Borneo, C. F. Baker collector. The two forms of Balocha bicolor can be separated as follows: Pronotum with anterior half orange, posterior yellowish B. bicolor bicolor Pronotumiunitorm|lysoraneey a) es ne eee B. bicolor uniformis So far I have included seven species in Balocha, namely, B. astuta (Melichar), B. bicolor n. sp., B. lucida Maldonado, B. melichari (Baker), B. nacreatus (Baker), B. pallida Maldonado, and B. tricolor Distant. Idiocerinus baker Merino is either a Balocha or a Pedioscopus as Merino (1936, p. 326, 327) writes about “the reduction in size of the second apical cell” and “second outer reduced” in this species. The coloration would be unusual for a Balocha, and it agrees more with the coloration of other species of Pedioscopus. Dr. T. Ishihara, from Ehime University, Matsuyama, Japan, has kindly pointed out to me that “Balocha flavocapitata Kato is not an Idiocerus but a species closely related to Oncopsis mali (Matsumura).” The generic position of Idiocerinus stali Fieber and Balocha angulifera Walker remain to be established. REFERENCES Maldonado-Capriles, J. 1961. Studies on Idiocerinae Leafhoppers: I. Idio- cerinus Baker, 1915, synonym of Balocha Distant, 1908 and notes on the species of Balocha. Proc. Ent. Soc. Wash. 63(4): 300-308. 1964. Studies on Idiocerinae Leafhoppers: II. The Indian and Philippine species of Idiocerus and the genus Idioscopus. Proc. Ent. Soc. Wash. 66( 2) :89-100. Merino, G. 1936. Philippine Cicadellidae (Homoptera). Philippine Jour. Sci. 61(3) :307—400. PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 101 THE NEARCTIC DORYCTINAE, VI. THE GENERA ACROPHASMUS, GLYPTOCOLASTES, DORYCTINUS, AND A NEW GENUS, STENOCORSE (HYMENOPTERA: BRACONIDAE) Pau M. Marsu, Systematic Entomology Laboratory, Entomology Research Division, Agr. Res. Ser., USDA* The present paper deals with four genera of the Doryctinae in which the head is narrow behind the eyes and somewhat transverse. This shape of the head differs from that of the typical Doryctinae, as ex- emplified by Doryctes, and has influenced past workers to place these genera in several other supergeneric groups (see discussion in Marsh 1965). The species described below in Acrophasmus represent the first records of this genus for the Nearctic Region. Acrophasmus Enderlein Acrophasmus Enderlein, 1912, p. 16. Type-species: A. exilis Enderlein. Monotypic and original designation. Head transverse or subcubical, temples narrow; first flagellar segment longer than second; forewing (fig. 1) with 3 cubital cells; recurrent vein entering apex of first cubital cell; first brachial cell closed at apex; subdiscoideus leaving first brachial cell below middle of apical end; first segment of mediella short, one- fourth or one-fifth as long as second segment; hindwing of male with a stigma; foretibiae with a row of 5-12 stout spines on anterior edge; tergum (24+ 3) (fig. 4) with a transverse arcuate furrow behind middle and often with a second transverse furrow behind this curved in opposite direction; digitus of male genitalia usually with a tooth on inner side (fig. 6). The sculpturing of tergum (2+ 3), with its transverse grooves rather than oblique basal grooves, and the short first segment of the mediella in the hindwing will distinguish Acrophasmus from the other genera in this group. This genus has been previously recorded only from the Neotropical Region. One described species, secundus (Muesebeck and Walkley ), and several new species described below were placed in the genus Doryctinus in the collection of the U. S. National Museum but are congeneric with Acrophasmus. Key ro NEaArctTic SPECIES OF Acrophasmus 1. Length of first abdominal tergum usually equal to apical width, at most slightly longer, but then abdominal terga beyond (2+ 3) smooth; female antenna 26-segmented or less — 2 Length of first abdominal tergum at least 1. a times cvibaaae Ta apical width; abdominal terga beyond (2+ 3) sculptured; female antenna Di -SCoMeENteG sl Ola eI ONG meee eee ge nan Ie te Na OA Ee es SIR | 3 *Mail address: c/o U.S. National Museum, Washington, D.C. 20560. 102 PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 2. Radial vein ending on wing margin well before apex, radial cell short along wing margin; abdominal terga beyond (2 + 3) punctate; first abdominal tergum as long as apical width; ovipositor of female slightly shorter than ordyslengthret eet lee el ee is he butleri Marsh, n. sp. Radial vein reaching apex of wing, radial cell longer; abdominal terga smooth beyond (2+ 3); first abdominal tergum slightly longer than apical width; ovipositor of female equal to body length __ lyeti Marsh, n. sp. 3. Ovipositor of female at least 1.5 times longer than body; vertex rugoso- DUTICLALE IEE 2 pees eee ee 8 eee secundus (Muesebeck and Walkley ) Ovipositor at most 1.25 times longer than body, usually about equal to body lempth vertex tsttiatess oe.) fue eee OU 2 4. Vertex narrow, sharply declivous posteriorly; malar space San one-fourth eye height, temples one-third eye width; body color reddish brown or Nehitere Miia ines Ow ele for 9 oss) es) Fraebinns ferrugineus Marsh, n. sp. Vertex rounded, not sharply neuen posteriorly; malar space about one- third eye height, temples at least one-half eye width; body color brown OT. ane DTO Wi. ss 2. te retell asl pee Be arizonensis Marsh, n. sp. Acrophasmus arizonensis Marsh, n. sp. (Figs. 8, 9) Female.—Body length, 5-5.5 mm; ovipositor, 6-7 mm. Color brown to dark brown, head and mesonotum usually darker, legs beyond femora and antennae light brown. Head subcubical, vertex rounded, not declivous behind; temples one-half eye width; malar space one-third eye height; frons not excavated; head entirely finely striate except temples which are usually smooth below; antenna 30-segmented. Propleuron rugose along posterior margin, punctate above propleural groove; anterior mesonotal angles rounded; mesonotal lobes punctate; notauli foveolate, obscured behind by wide rugose area covering posterior one- half of median mesonotal lobe; scutellar furrow with 6 cross carinae; scutellar disc punctate, margined laterally; mesopleural disc and mesosternum punctate; mesopleural furrow shallow, narrow, weakly foveolate; subalar groove wide, rugose; basolateral areas of propodeum indicated by indistinct carinae, punctate basally; remainder of propodeum rugose. Foretibia with row of 7-8 spines; fore and midlegs nearly smooth, hindlegs finely punctate. Wing venation as in fig. 8; radiella absent or very indistinct. First abdominal tergum 1.5 times longer than apical width, longitudinally rugose, median area indicated basally by 2 indistinct carinae and apically by a raised short area, obscured medially; tergum (2+ 3) 1.25 times longer than wide, longitudinally rugose except apex which is smooth, both transverse grooves distinct; remainder of abdominal terga punctate on basal one-half, smooth on apical one-half; ovipositor a little longer than body. Male.—Essentially as in female; antenna 27-segmented; hindwing with a stigma (fig. 9). Holotype Female—ARIZONA: Bear Canyon, Hk. Hwy. mi. 12, Santa Catalina Mountains, July 14, 1961, Werner, Nutting, U.V. It. trap. USNM 69637. Paratypes.—1?, 14, same data as type. Female paratype deposited in collection of Univ. of Arizona, Tucson; male in U. S. Nat. Mus. PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 103 Acrophasmus butleri Marsh, n. sp. (Figs. 10, 11) Female——Body length, 4 mm; ovipositor, 3 mm. Color dark brown except head and forelegs which are lighter brown. Head somewhat transverse; vertex rounded, not declivous behind; malar space one-third eye height; temples about three-fifths eye width; frons and vertex striate, temples weakly striate dorsally, smooth ventrally, face indistinctly rugose; antenna 23-segmented. Propleuron rugosopunctate; propleural groove indistinct; anterior angles of mesonotum prominent, rounded; mesonotal lobes rugosopunctate; notauli indistinct, obscured behind by wide rugose area covering posterior one-half of median mesonotal lobe; scutellar furrow with 7 cross carinae; scutellar disc punctate, margined laterally; mesopleural disc rugosopunctate; mesopleural furrow deep, narrow, weakly foveolate; mesosternum punctate; propodeum entirely rugosopunctate. Foretibia with row of 5-6 spines (forelegs missing in female holotype). Wing venation as in fig. 10; radial cell short, radial vein reaching wing margin well before apex. First abdominal tergum as long as apical width, longitudinally rugosopunctate, raised median area distinct only at apex; tergum (2+ 3) as long as greatest width, longitudinally rugosopunctate before first transverse groove, rugae disappearing behind so that apical one-fourth of tergum is punctate, first transverse groove very indistinct, second absent; remainder of terga punctate entirely; ovipositor slightly less than body length. Male.—Essentially as in female; antenna 22-23 segmented; hindwing with a stigma (fig. 11). Holotype Female—ARIZONA: Santa Rita Range Reserve, Pima County, June 2, 1958, G. D. Butler, swept/mesquite. USNM 69638. Paratypes.—2 4 ¢, same data as type except one dated May 4, 1957. One ¢ paratype deposited in collection of Univ. of Arizona, Tucson; other in U.S. Nat. Mus. Acrophasmus ferrugineus Marsh, n. sp. , (Figs. 1, 2, 4-7) Female.—Body length, 4.5-7 mm; ovipositor, 4-6.5 mm. Color brownish red to reddish orange; head, legs, and venter of abdomen always reddish orange. Head slightly transverse; vertex sharply declivous behind; frons slightly depressed; eyes large; malar space about one-fourth eye height; temples about one-third eye width; vertex and temples striate, frons and face rugosopunctate; antenna 30-40 segmented. Pronotum short; propleuron rugose; propleural groove broad, foveolate; mesonotum sharply declivous anteriorly, anterior angles rounded; mesonotal lobes punctate; notauli foveolate anteriorly, obscured posteriorly by wide rugose area which covers posterior three-fourths of middle mesonotal lobe; scutellar furrow with 6 cross carinae; scutellar disc flat, punctate; mesopleuron rugosopunctate; mesopleural furrow narrow, weakly foveolate; mesosternum punctate; propodeum entirely rugose, rarely with some indication of a pattern of carinae. Foretibia with an irregular row of 8—12 stout spines; legs mostly punc- tate. Wing venation as in fig. 1. First abdominal tergum 1.75 times longer than apical width, longitudinally rugose with an indistinct raised median area which narrows apically; tergum (2 + 3) slightly longer than its greatest width, longi- 104 PROC, ENT, SOC, WASH,, VoL. 70, NO, 2, JUNE, 1968 ~ , Acrophasmus ferrugineus, n. sp.: 1, fore and hindwings, 2; 2 Figs. 1, 2, 4-7 hindwing, @; 4, abdominal tergum (2-++3), dorsal view; 5, @ genitalia; 6, di- gitus; 7, ninth sternum, ¢. Fig 3, A. secundus (Mues. & Walk.), abdominal ter- gum (2+-3), dorsal view. a tudinally rugose before first transverse groove, longitudinally rugosopunctate be- hind; second transverse groove present, usually weak; remainder of abdominal terga punctate; ovipositor nearly as long as body. Male.—Essentially as in female; length, 2.4-5.5 mm; antenna 21-32 seg- mented; head not always as sharply declivous behind; hindwing with a stigma (fig. 2); genitalia as in figs. 5-7. Holotype Female—NORTH CAROLINA: Durham, June 30, 1943, J. A. Beal colr., Host: Celtis laevigata Weed. USNM 69639. Paratypes—GEORGIA;: Atlanta, 1 °, June 23, 1938, P. W. Fattig. FLORIDA: Mariana, 4 °°, June 19, 1905, W. F. Fiske. MISSOURI: Jefferson City, 1 &°, April 22, 1958, A. C. Burrill, ex. Diospyros virginiana L, MISSISSIPPI: Doko (?), 1 &, 1 é, April 14, 1920, M. W. Blackman, NORTH CAROLINA: Durham, 5 ° °, June-August, 1943, C. laevigata, Quercus borealis var. maxima ( Marsh.) Ashe; Raleigh, L é, August 16, 1940, R. C. Barnes. PENNSYLVANIA: Linglestown, PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 105 4 29,1 6, A. B. Champlain, Hopk. U. S. 12214. SOUTH CARO- LINA: Ferguson, 1 2°, 5 46, ex Lyctus planicollis LeConte in Fraxinus; Charleston, 1 °, June 1, 1905, W. F. Fiske. TEXAS: Browns- ville,6 2 °,4 46, H.S. Barber; Cameron County, 1 6, August 3, 1928, R. H. Beamer; Victoria, 1 ?, June 11, 1907, R. A. Cushman; Winona, 229°,3 64, October, 1930, H. Baker, dead pecan wood. VIRGINIA: Falls Church, 1 3, June, 1922, R. A. St. George. All paratypes de- posited in U.S. Nat. Mus. In addition to the type series I have seen specimens of ferrugineus from Brazil, Guatemala, and the British West Indies. Host.—Several specimens were reared from Lyctus planicollis Le- Conte in Fraxinus, and from unknown hosts in Celtis laevigata, Quercus borealis var. maxima, and Diospyros virginiana. Acrophasmus lyeti Marsh, n. sp. (Figs. 12, 13) Female.—Body length, 2.5-4 mm; ovipositor, 2.5-4 mm. Color reddish brown except legs and sometimes apex of abdomen and venter of thorax which are honey yellow. Head subcubical; vertex rounded; eyes large; malar space nearly one-fourth eye height; temples one-third eye width; frons, vertex, and temples finely striate, temples often smooth on lower portion, face rugulose; antenna 18-26 segmented. Propleuron punctate; propleural groove broadening posteriorly, foveolate; mesonotal lobes punctate; notauli foveolate, obscured behind by wide rugose area which covers apical one-half to three-fourths of median mesonotal lobe; anterior mesonotal angles not prominent, rounded; scutellar furrow with 6 cross carinae; scutellar dise punctate; mesopleural disc and mesosternum punctate; mesopleural furrow narrow, foveolate; subalar groove rugose; propodeum mostly rugose, carinae weakly indicated but forming large areola medially, basolateral areas punctate. Foretibia with row of 5—7 spines. Wings as in fig. 12. First abdominal tergum one and one-third times longer than apical width, longitudinally rugose, raised median area distinct basally and apically, obscured medially; tergum (2-4 3) as long as widest part, longitudinally rugose, smooth at apex, first transverse groove distinct, sinuate, second groove indistinct; re- mainder of terga usually smooth and highly polished, at most very finely granular basally; ovipositor equal to length of body. Male.—FEssentially as in female; length, 2—3 mm; antenna 17-23 segmented; hindwing with a stigma (fig. 13). Holotype Female—FLORIDA: Indian River County, June 10, 1957. USNM 69640. Paratypes—FLORIDA: Crescent City, 11 299, 8 ¢4, reared from bamboo infested with Lyctus striatus Melsheimer; Indian River County, 8 22,1 6, June 10, 1957; Lake City, 1 2, 1 4, H. G. Hub- bard, ex Lyctus in bamboo. LOUISIANA: Tallula, 8 22, 15 44, October 5, 1939, M. B. Christian, ex Lyctus infested Pecan. NORTH CAROLINA: Watauga County, 3 °°, ex-hickory rake handle infested with powder post beetles. SOUTH CAROLINA: Ferguson, 3 2? ?, 106 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 1 eee Figs. 8-15, wings of Acrophasmus species: 8, arizonensis, n. sp., 2 fore and hindwing; 9, same, 6 hindwing; 10, butleri, n. sp., 2 fore and hindwing; 11, same, @ hindwing; 12, lycti, n. sp., 2 fore and hindwing; 13, same, ¢ hindwing; 14, secundus (Mues. & Walk.), 2 fore and hindwing; 15, same, ¢ hindwing. 12 64, W. F. Fiske, Hopk. U. S. 3486E. All paratypes deposited in U.S. Nat. Mus. Hosts.—Lyctus sp. and L. striatus in bamboo. Acrophasmus secundus ( Muesebeck and Walkley), n. comb. (Figs. 14, 15) Heterospilus texanus Ashmead, 1896, p. 214. Holotype male in U. S. Nat. Mus., no. 58860, preocc. in Doryctinus by Ashmead, 1889. Doryctinus secundus Muesebeck and Walkley, 1951, p. 178. New name for texanus Ashmead, 1896. Female.—Body length, 3.5-6 mm; ovipositor, 5-10 mm. Color dark brown, tarsi and antennae brown, tibiae with white annulus at base. Head _ nearly cubical; vertex rounded, not sharply declivous behind; temples two-thirds eye width; malar space about one-third eye height; vertex, frons, and face PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 107 rugosopunctate, frons sometimes appearing striate in smaller specimens, temples transversely striate, often smooth on lower portion; antenna 27-34 segmented. Propleuron rugose; propleural groove very broad; mesonotal lobes rugosopunctate, anterior mesonotal angles prominent but rounded, median mesonotal lobe slightly depressed anteriorly between angles; notauli foveolate anteriorly, ob- scured posteriorly by wide rugose area which covers posterior three-fourths of middle mesonotal lobe; scutellar furrow with 6 cross carinae; scutellar disc punctate, margined laterally; mesopleuron rugose; mesopleural furrow broad, foveolate; mesosternum striatopunctate; propodeum entirely rugose with lateral carinae and areola occasionally weakly indicated. Legs granular; foretibia with row of 8-10 stout spines. Wing venation as in fig. 14. First abdominal tergum 1.5 times longer than apical width, longitudinally rugose, often with an indistinct raised median area; tergum (2+ 3) 1.25 times longer than greatest width, longitudinally rugose except apex which is smooth; first transverse groove very distinct, second usually absent, occasionally indicated by a weak indentation; remainder of terga granular on basal three-fourths, smooth on apical one-fourth; ovipositor at least 1.5 times longer than body. Male.—Essentially as in female; hindwing with a stigma (fig. 15). Type Locality—Cypress Mills, Texas. Distribution Georgia, Louisiana, Massachusetts, Michigan, Mis- souri, New Jersey, Ohio, Pennsylvania, Rhode Island, Texas, Virginia; Mexico (Cuernavaca ). Glyptocolastes Ashmead Glyptocolastes Ashmead, 1900, p. 142. Type-species: Glyptocolastes texanus Ashmead. Monotypic and original designation. Glyptodoryctes Ashmead, 1900, p. 144. Type-species: Heterospilus caryae Ashmead. Monotypic and original designation. New synonymy. Head somewhat transverse, narrow behind the eyes; first flagellar segment longer than second; thorax stout; forewing with 3 cubital cells; second segment of radius often shorter than first intercubitus; recurrent vein entering apex of first cubital cell; subdiscoideus leaving first brachial cell below its middle; first segment of mediella at least equal to second segment, usually longer; foretibia with row of 6-10 stout spines; abdominal tergum (2+ 3) with oblique diverging grooves which set off basal corners of tergum (fig. 24), rarely these grooves are connected medially by a fine transverse line. Ashmead (1900) characterized Glyptocolastes and Glyptodoryctes only as he included them in his key to the Ichneumonoidea. Simi- larities in wing venation, sculpture of tergum (2+ 3), and general habitus leaves no doubt that the type-species of these genera are congeneric. In 1909, Crawford described the species bruchivorus and placed it in Glyptocolastes. However, because of differences in male genitalia, wing venation, and habitus, I feel that this species is not congeneric with texanus and caryae and is quite distinct from all 108 PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 genera of the Doryctinae, a fact which warrants a separate generic placement for it (see below ). Key to Species or Glyptocolastes tyes ee ee ee eee 2 ini fo Sy ae tt ray ae. a ge EME SE 3 2. First abdominal tergum shorter than apical width, at most equal; second segment of radius shorter than first intercubitus texanus Ashmead First abdominal tergum longer than apical width; second segment of radius usually equal to or longer than first intercubitus —___ earyae (Ashmead ) SLincwinh Aviat a Stuer, eee caryae (Ashmead ) Hindwime wirbout a-stigma texanus (Ashmead ) Glyptocolastes caryae (Ashmead ), n. comb. (Bigs 16. 177 20; 21,24) Heterospilus caryae Ashmead, 1896, p. 214. Holotype female in U. S. Nat. Mus., no. 69559. Glyptodoryctes caryae (Ashmead), 1900, p. 144. Hormiopterus claripennis Brues, 1907, p. 61. Holotype female in Milwaukee, Wisconsin, Public Museum. Female.—Body length, 2.5-6 mm; ovipositor, 2-6 mm. Color light to dark brown, legs usually brown or honey yellow. Head transverse, temples about two-fifths eye width, somewhat wider in smaller specimens; malar space about one-fourth eye height; face roughened except for raised smooth line down middle; temples usually smooth; frons and vertex transversely striate; vertex in larger specimens declivous behind; antenna 23-37 segmented. Pronotum very short; propleuron rugose; propleural groove deep, foveolate; mesonotum strongly declivous anteriorly, dorsal surface at right angles with anterior surface; mesonotal lobes punctate; notauli deep, foveolate, obscured behind by wide rugose area which is about as wide as scutellar disc; scutellar furrow with 7 cross carinae; scutellar disc punctate; mesopleural disc punctate; mesopleural furrow deep, foveolate; subalar groove wide, rugose; mesosternum punctate; propodeum strongly rugose, carinae usually not distinct but rugae occasionally forming areola medially. Hindcoxae punctate laterally, rugose dorsally; femora punctate. Wing venation as in fig. 16; second segment of radius usually longer than first intercubitus; first segment of mediella longer than second, usually much longer. First abdominal tergum about 1.5 times as long as apical width, strongly longitudinally rugose, with a weakly defined raised median area; tergum (2+ 3) slightly longer than wide, oblique grooves deep and occasionally connected across tergum by a semicircular fine line or groove (fig. 24); tergum (2+ 3) longitudinally rugosopunctate on nearly its entire length, smooth only at extreme apex; remainder of terga granular at least basally; ovipositor nearly as long as body. Male.—Essentially as in female; length, 2.5-4 mm; antenna 21-32 segmented; hindwing with a stigma (fig. 17); genitalia as in figs. 20, 21. Type Locality—Morgantown, West Virginia. Distribution —Arkansas, Florida, Georgia, Kansas, Maryland, Missis- PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 109 Paes 1 pe Figs. 16, 17, 20, 21, 24, Glyptocolastes caryae (Ashm.): 16, fore and hindwing, 2; 17, hindwing, ¢; 20, ¢ genitalia; 21, ¢ ninth abdominal sternum; 24, ab- dominal tergum (2+3), dorsal view. Fig. 18, G. texanus Ashm., fore and hind- wing, 2. Figs. 19, 22, 23, Stenocorse bruchivora (Cwfd.): 19, fore and hind- wings, 2; 22, 6 genitalia; 23, ¢ ninth abdominal sternum. 110 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 sippi, New York, North Carolina, Pennsylvania, South Carolina, Texas, Virginia, Washington, D. C., West Virginia. Host.—Scobicia bidentata (Hom). Glyptocolastes texanus Ashmead (Fig. 18) Glyptocolastes texanus Ashmead, 1900, p. 142. Holotype female in U. S. Nat. Mus., no. 14167. Female.—Body length, 2-4 mm; ovipositor, 1-2 mm. Color entirely brown or dark brown except legs, lower part of head, and mouthparts which are often light brown or honey yellow. Head somewhat transverse; temples about one- half eye width; malar space one-third eye height; vertex and frons transversely striate, face roughened, temples smooth at least on lower portion; vertex declivous behind in larger specimens; antenna 19-27 segmented. Pronotum short; propleuron rugosopunctate; propleural groove shallow, indistinctly foveolate; mesonotum strongly declivous anteriorly, anterior and dorsal surfaces at right angle or greater; mesonotal lobes punctate; notauli shallow, foveolate, obscured posteriorly by rugose area which is usually not as wide as scutellar disc; scutellar furrow with 7 cross carinae; scutellar disc punctate, margined laterally; meso- pleural disc rugosopunctate; mesopleural furrow shallow, weakly foveolate; subalar groove indistinct; propodeum rugose, basolateral areas often punctate. Hindcoxae punctate, often rugose dorsally; femora punctate. Wing venation as in fig. 18; second segment of radius usually shorter than or equal to first intercubitus; first segment of mediella longer than second. First abdominal tergum usually shorter than apical width, at most equal, longitudinally rugose, with narrow raised median area; tergum (2+ 3) wider apically than long, longitudinally rugosopunctate on basal three-fourths, oblique grooves shallow; remainder of terga punctate basally, smooth apically; ovipositor about as long as abdomen plus one-half of thorax. Male.—Essentially as in female; hindwing without a stigma. Type Locality—San Diego, Texas. Distribution —Arizona, Texas; Baja California. Host.—Dendrobiella quadrispinosa (Lec. ). Stenocorse Marsh, n. gen. Type-species: Glyptocolastes bruchivorus Crawford. Head transverse; temples narrow, equal to one-half eye width or less; thorax robust; notauli weakly or not at all defined; second segment of radius of forewing equal to or shorter than first; radius reaching wing margin before apex of wing; stigma broad; recurrent vein entering base of first cubital cell; subdiscoideus leaving first brachial cell near its base; nervulus postfurcal; radiella of hindwing weak or absent; first segment of mediella longer than second; postnervellus usually curved toward wing apex; hindwing of male without stigma; foretibia with row of 8-12 short spines on outer edge; femora and tibiae somewhat swollen; abdomen short, broadly oval; abdominal tergum (2+ 3) with a_ bisinuate transverse groove or weak line on basal one-half; ovipositor not longer than abdomen; digitus and cuspis of male genitalia truncate. PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 IIL As mentioned earlier, Glyptocolastes bruchivorus is quite distinct from most Doryctinae and certainly does not belong in that genus which is exemplified by G. texanus Ashmead. Therefore, the above new genus is proposed for this species and several, as yet undescribed, Neotropical ones. Of particular importance in distinguishing Stenocorse is the transverse head and narrow temples, the robust habitus, the short second segment of the radius, the sculpture of abdominal tergum (2+ 3), and the structure of the male genitalia. Species in this genus are apparently parasites of seed beetles in the family Bruchidae and are distributed from the southwestern United States to South America and the West Indies. The Nearctic Region contains one species, the type-species of the genus. Stenocorse bruchivora (Crawford), n. comb. (Figs. 19, 22, 23) Glyptocolastes bruchivorus Crawford, 1909, p. 203. Holotype female in the U. S. Nat. Mus., no. 12816. Female——Length of body, 3-4.5 mm; ovipositor, 0.75-1.5 mm. Thorax and abdomen reddish-brown, head and legs lighter brown, legs sometimes honey yellow, mesonotum, mesopleuron, and abdominal terga often marked with black, propodeum often entirely black; entire body with a thick covering of gold hair. Head rugose, vertex strongly so; frons slightly excavated; malar space one-third eye height; temples slightly less than one-half eye width; antennae 20-27 segmented. Thorax short and broad, nearly as broad between tegulae as high; pronotum extremely short; propleuron longitudinally rugose; mesonotum sharply declivous anteriorly, coarsely rugose; notauli weak, indicated by coarser rugae, meeting posteriorly in a wide rugose area; scutellar furrow with 5 cross carinae; scutellar disc flat, granular; mesopleural disc rugulopunctate; subalar groove wide, foveolate; mesopleural furrow wide, deep, weakly foveolate, about two-thirds mesopleural width; mesosternum granular, short transverse rugae on either side of median sternal groove; propodeum sharply declivous posteriorly, dorsal surface about as long as scutellar disc, strongly rugose, oc- casionally with granular basolateral areas and a short basal carina. Wing vena- tion as in fig. 19; second segment of radius never longer than first; nervulus postfurcal by about its own length. Femora and tibiae somewhat swollen, femora about three times as long as wide; tarsi short, hind basitarsus about as long as first flagellar segment; last segment of hindtarsus equal in length to second. Abdomen about as long as thorax, oval, strongly arched; first tergum longitudinally rugose, about one and one-half times wider at apex than long, abruptly declivous basally, with two longitudinal carinae converging toward apex; tergum (2+ 3) two-thirds as long as apical width, longitudinally rugose except for apical one-fourth which is granular; tergum (2+ 3) with a bisinuate transverse groove across basal one-half which is sometimes represented only by a weak line, lateral sections of this groove sometimes extend to base of tergum; remainder of abdominal terga granular; ovipositor at most equal in length to abdomen beyond first tergum. re, PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Male.—Essentially as in female; length of body, 2.5-4.5 mm; genitalia as in figs. 22, 23; digitus and cuspis truncate, ninth sternum broad. Type Locality —Victoria, Texas. Distribution.—Arizona, Texas; Mexico, Canal Zone, Peru; Hawaii. Hosts.—Algarobius prosopis (Lec.), Acanthoscelides quadridentatis (Schaffer), Mimosestes sallaei (Sharp), Megacerus sp. Doryctinus Roman Doryctinus Roman, 1910, p. 122. Type-species: Exothecus rugulosus Cresson. Monotypic and original designation. Roman proposed this genus on the basis of assumed topotypic material of Exothecus rugulosus. His brief description and illustra- tions indicate that this material was not conspecific, and probably not congeneric, with E. rugulosus. The type of rugulosus is, however, unique and apparently intermediate between Glyptocolastes and Acrophasmus. The first segment of the mediella in the hindwing of rugulosus is slightly shorter than in species of Glyptocolastes, but longer than in species of Acrophasmus. The sculpturing of tergum (2+ 3) combines the oblique basal grooves of Glyptocolastes and the transverse furrows of Acrophasmus. In general habitus, E. rugulosus appears similar to G. caryae. Other than the holotype female, I have seen only two females which can be referred to Doryctinus. Until I am able to see more material of this group, particularly males, I wish to retain it as a distinct genus. A second species referred to Doryctinus, secundus (Muesebeck and Walkley), is now placed in the genus Acrophasmus (see before). Doryctinus rugulosus (Cresson) Exothecus rugulosus Cresson, 1872, p. 190. Holotype female in U. S. Nat. Mus., no. 1620. Doryctes texanus Ashmead, 1889, p. 627. Holotype female in U. S. Nat. Mus., no. 2930 (same specimen as E. rugulosus Cresson). Female.—Body length, 6 mm; ovipositor, 5 mm. Head, legs, and abdomen brown, thorax darker brown. Head somewhat transverse; temple slightly less than one-half eye width; malar space about one-third eye height; vertex and frons striate, temples smooth, face rugulose except for smooth median longitudinal raised line; vertex narrow, declivous posteriorly; antenna broken in type, first flagellar segment longer than second. Pronotum short, nearly vertical; propleuron rugose, propleural groove deep, foveolate; mesonotum sharply declivous an- teriorly; mesonotal lobes punctate, median one with an indistinct median longitudinal foveolate line on anterior face; notauli foveolate anteriorly; scutellar disc granular; mesopleural disc and mesosternum punctate; mesopleural furrow foveolate; subalar groove wide, rugose; propodeum rugose, without carinae. Legs including coxae punctate. Second segment of radius about as long as first intercubitus; recurrent vein entering extreme apex of first cubital cell; radiellen PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 TS cell strongly narrowing toward apex, radiella reaching wing margin before wing apex; first segment of mediella equal to second. First abdominal tergum longer than apical width, longitudinally rugose; tergum (2+ 3) with 2 wide, shallow, longitudinal, diverging grooves on basal one-half and 2 narrow, deep, transverse grooves across middle; remainder of abdominal terga granular; ovipositor nearly as long as body. Type Locality.—Texas. REFERENCES Ashmead, W. H. 1889 (1888). Descriptions of new Braconidae in the collec- tion of the U. S. National Museum. Proc. U. S. Nat. Mus. 11:611-671. 1896. Descriptions of new parasitic Hymenoptera. Trans. Amer. Ent. Soc. 23:179-234. 1900. Classification of the ichneumon flies, or the superfamily Ichneumonoidea. Proc. U. S. Nat. Mus. 23:1-220. Brues, C. T. 1907. Notes and descriptions of North American parasitic Hymenoptera, III. Bull. Wisc. Natur. Hist. Soc. 5:54-62. Crawford, J. C. 1909. New parasitic Hymenoptera. Proc. Ent. Soc. Wash. 11:203-207. Cresson, E. T. 1872. Hymenoptera Texana. Trans. Amer. Ent. Soc. 4:153—292. Enderlein, G. 1912. Zur Kenntnis der Spathiinen und einiger verwandter Gruppen. Arch. Naturgesch. 78( A) (2) :1-37. Marsh, P. M. 1965. The Nearctic Doryctinae, I. A review of the subfamily with a taxonomic revision of the tribe Hecabolini. Ann. Ent. Soc. Amer. 58:668-699. Muesebeck, C. F. W. and L. M. Walkley. 1951. Family Braconidae. in Muesebeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2:90-184. Roman, A. 1910. Notizen zur Schlupwespen Sammlung des schwedischen Reichsmuseum. Ent. Tidskr. 31:109-197. THE SYNONYMY OF AMOEBALERIA SACKENI GARRETT (DireTERA: HELEOMYZIDAE) A series of 21 specimens, all taken in the same cave (Equality Cave, Saline County, Illinois, 22 May—15 October 1967) by Ronald A. Brandon, of Southern Illinois University, and kindly submitted by him for determination, shows variation between the conditions considered typical of Amoebaleria defessa Osten Sacken, 1877, and A. sackeni Garrett, 1925 (v. Gill, 1962, Proc. U.S. Nat. Mus. 113: 578-580; Steyskal, 1967, Proc. Ent. Soc. Wash. 69:296). The variation indicates that the form described as A. sackeni is no more than that assumed by the larger or more robust specimens of A. defessa. A. sackeni Garrett should, therefore, be considered a synonym of A. defessa Osten Sacken.—GrorcE C. STEYSKAL, Systematic Entomology Laboratory, Entomology Research Division, Agr. Res. Serv., USDA, c/o U.S. National Museum, Washington, D.C. 20560. 114 PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 THE GENUS BRACHYTREMELLA TRAGARDH, 1946, WITH DESCRIPTIONS OF THREE NEW SPECIES (ACARINA: DIARTHROPHALLIDAE )! Preston E. Hunter and SANpRA GLOVER,” Department of Entomology, University of Georgia, Athens, Georgia 30601 Tragardh (1946) erected Brachytremella for a single female, B. spinosa Tragardh, collected from a passalid beetle, Protomocerus sp., from New Guinea. Womersley (1961) redescribed the female and described the male and tritonymph from passalids collected in New Guinea. At the same time Womersley described two additional species—trdgdrdhi from Australia and bornemisszai from Australia and New South Wales—and gave a key to the three species. In this paper we are describing three new species taken from a small collection of alcohol preserved passalid beetles from Costa Rica and bringing to six the number of species in the genus. Brachytremella Tragardh, 1946 Tragardh, I. 1946. Ent. Medd. 24:386. New genus by designation. Womersley, H. 1961: Trans. Roy. Soc. S. Aust. 84:11. Adults of this genus may be recognized by the following characters: body oval in shape and bearing 6 pairs of long, heavy, barbed, capitate dorsal setae; metopodal plates absent or fused with ventral plate; genital plate of the female not separated posteriorly by a suture from the ventral plate; legs II of the male not modified. The protonymph has 5 pairs of dorsal setae of the same type as in the adult, whereas the deutonymph has 6 pairs of setae of this type. Brachytremella joanae, n. sp. (Fig. 1A-F) Known only from the female and deutonymph and may be recognized by the following characters: 2nd and 4th pairs of dorsal setae shorter than other dorsal setae, dorsum of femora III and IV bearing 2 heavy, spined setae; female ventral plate not enlarged behind coxae IV; in the deutonymph only sternal setae IV are on the ventral plate. FEMALE. Idiosoma 480 long, 370u wide. (Measurements are the average of 2 specimens.) Shape rounded oval. Dorsum (fig. 1A). Dorsal plate entire, not extending to posterior and posterolateral margins of body; plate with a number of pores located in the general arrangement of the setal pattern of mesostigmata. Six pairs of spined, heavy, tapering setae, all ending in a small but distinct knob; 4 pairs arising from dorsal plate, pairs 5 and 6 arising from integument; anterior (lst) pair 320u long, 2nd 210, 4th pair slightly longer * University of Georgia College of Agriculture Experiment Station, Journal Series Paper No. 23, College Station, Athens. Partially supported by National Science Foundation Grant No. GB-3414. 2 Associate Professor and Graduate Student, respectively. PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 1S Fig. 1. Brachytremella joanae, n. sp. Female: A, dorsum; B. venter; C, car- uncle II, ventral view; D, tectum; E. chelicera. Deutonymph: F, venter. than 2nd; median pair of setae arising from integument 290. long, lateral pair from integument 365 long. Venter (fig. 1B). Sternal, metasternal and ventral plates coalesced, extending posterior of coxae IV, not widening behind coxae IV; combined plate 400u long, 2154 between coxae III, 130” between coxae IV and 115u near the rounded posterior end of the plate; sternal area only semisclerotized medially, sclerotized at lateral margins; reticulations on sclerotized area of sternal, metasternal and genital plates; ventral area of plate not distinctly reticulated; 5 pairs of ventral setae; sternal setae I arising from 116 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Table 1. Deutonymph-adult leg chaetotaxy formulae of Brachytremella. Total number of setae only given for the tarsi. Deviations of B. crossi given in footnotes. Leg Coxa Trochanter Femur Genu Tibia Tarsus 0) I 21 el Ol I 0 0 J——1 1 il 1——1 1——1 7 setae® 2 0 i 00 11 0) 2, 11 Ol II 0——0 1——1 1 2 1 J1——1 1——1 13 setae 2) 2 It 00 leh 0) 1 22, iil Ol Ill 0——0O0 1——O0 1 1 J——1 ]——1 13 setae 2 , idl 00 WIL 0) 2 22, iLL Ol IV 0——0 1——0* 1 iP 1——1 ]1——1 13 setae 1 D, WI 00 JUL * B. crossi: Tat 2, » B. crossi: jee ial © B. crossi: 6 setae semisclerotized area of sternum, approximately 3 times length of other ventral setae; 5th pair of setae located at posterior level of coxae IV; positions of other setae and relative length of setae as shown; pores in plate and integument as shown. Genital plate elongate-rounded, 178u long, 1434 wide, fused posteriorly to ventral plate; thickened along free margins. Endopodal plates II fused to sternal plate anteriorly, free along posterior half; III free, curving around anterior of coxa III; IV encircling coxa IV, widening slightly posterior of coxa. Anal plate wider than long, bearing 2 spined, tapering, apically knobbed setae up to 395 long. Peritremal plate surrounding peritreme, enlarging lateral of stigmata and bearing a pore; peritreme length 3-4 times width of stigmata. Metopodal plates absent. Tritosternum of type illustrated in fig. 2D; one pair of fleshy, ciliated setae flanking base. Legs. Coxa I fragmented, medial setae arising from integument. Chaetotaxy, following Evans (1963), given in table 1. All setae simple except the subterminal clawlike structure on tarsus I and spined, heavy setae on the following: 1 seta on genua I, III and IV; 1 on femora I and II; 2 on femora III and IV, the posterior seta on each segment shortest. Tarsus I with 2 terminal setae and a subterminal clawlike structure; tarsi II-IV with well developed padlike caruncles (fig. 1C) bearing a pair of thickened structures medially giving the appearance of claws (these structures are on the ventral surface of the caruncle and each consists of a thickened central part which bears a series of individual, thickened striae running parallel to the long axis of the caruncle and extending to the margin of the caruncle, median striae heavier and longer than lateral ones; proximal of this structure is a smaller but similar shaped line of thickened striae; these structures undoubtedly aid the mite in clinging to the sclerotized body of the host); caruncle thickened proximally and along lateral margins. Lengths of legs, including coxa and caruncle: I, 227u; II, 3554; Il], 3704; IV, 3804. Gnathosoma. PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 ILAL7 Ventrally as in following species (see fig. 3D). Tectum (fig. 1D) consisting of 4 terminal parts, median pair thickest and rakelike in appearance; outer parts ciliated, shorter than median and each bearing a short fingerlike ciliated structure; dorsally a small ciliated pointed structure projects between base of median extensions. Gnathosoma bearing a pair of short, simple setae dorsolaterally midway between idiosoma and palpal trochanter. Chelicerae strongly chelate; excrescence trough-shaped, toothed along both margins, arising from fixed digit (fig. LE). Palps normal for genus; number of setae per segment as follows: tarsus 6, tibia 7, genu 4, femur 4, and trochanter 1; relative lengths of setae as shown. DEUTONYMPH. Shape as in female. Idiosoma 4804 long, 370u wide. Dorsum. General features as in female, setae approximately same lengths as in female. Venter (fig. 1F). Ventral plate 260u long, 115 at widest part; bearing 1 pair of setae; 1 pair pores between coxae II and 1 pair at margin of plate between coxae IV; striations distinct, general pattern as shown; 4 pairs of setae arising from integument, positions and relative lengths as shown. Endopodal plates II, III and IV present, shapes and positions as illustrated. Anal plate diamond shaped; setae 375u long, of same type as dorsal setae. Peritreme and plate, and tritosternum as in female. Legs and gnathosoma with general features as in female. Male, protonymph and larva unknown. Described from 2 females and 2 deutonymphs. Collection data for holotype (female): Costa Rica, Cartago Province; 13-VII-1965; J. Baird, coll.; from alcohol vial containing passalus beetles Passalus perparvulus Kuw. and Pseudacanthus tennis Kuw. Female paratype and deutonymphs with same date. Holotype and one deutonymph deposited in U. S. National Museum, Washington, D. C., remaining specimens in collection of Department of Entomolgy, University of Georgia, Athens, Georgia. Brachytremella crossi, n. sp. (Figs. 2A-J; 4B) This species is distinct in having the first pair of dorsal setae shorter than the second and in having only 1 heavy spined seta on the dorsum of femora and genua III and IV; adults with the sterni-geniti-ventral plate enlarged posterior to coxae IV and the posterior setae on this plate near the plate’s posterior margin; male with a ventral spur on trochanters II and III. FEMALE. Idiosoma oval, 530u long, 343u wide. (Measurements are the average of 3 specimens.) Dorsum (fig. 2A). Dorsal plate entire, not extending to posterior and posterolateral margins of body, shape similar to idiosoma. Dorsum with 6 pairs of heavy, spined setae which taper to a small but distinct knob at tip; first 4 pairs arising from margin of plate, last 2 pairs arising from integument behind dorsal plate; anterior pair shortest, up to 200u long, 2nd pair 298 long, median pair on integument 291 long, relative lengths of others as shown. Numerous pores in plate in position of normal mesostigmata setal pattern. Venter (fig. 2B). Sternal, metasternal and ventral plates coalesced, 497 long, 177 wide between coxae III, constricting to 1334 between coxae IV and 167 at widest point behind coxae IV; rounded posteriorly; medial area of anterior margin 118 PROC, ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Fig. 2. Brachytremella crossi, n. sp. Female: A, dorsum; B, venter; D, trito- sternum; E, chelicera; F, tectum. Male: C, venter of tarsus III; G, venter. Deutonymph: H, dorsum. Protonymph: I, dorsum; J, venter. semisclerotized; 5 pairs of simple setae arising from plate, first pair 2-3 times length of other setae and arising from the semisclerotized area, last pair thickest and located well posterior to coxae IV, positions of other setae as illustrated; light reticulate markings prominent on posterior half of plate; pores on plate as illustrated. Genital plate elongate oval shape, fused to ventral plate; 1874 long, extending from middle of coxae IV to middle of coxae II; sternal plate thickened around genital opening. Anal plate transversely diamond shaped, bearing 2 setae up to 270u long, of same type as dorsal setae. Endopodal plates II fused with sternal plate; III free, curving around anterior end of coxae III and extending posteromedial of coxa; IV free, encircling coxae IV. Metopodal plates absent. Peritreme short, length approximately 3 times width of stigmata; peritremal plate triangular shaped, surrounding peritreme, knoblike enlargment lateral of stigmata containing a small pore. Tritosternum with 2 flattened lacinae; lacinae membranous with small teeth on lateral margin, slightly thickened medially (fig. 2D); a pair of ciliated setae flanking base. Legs. Chaetotaxy given in table 1. Leg I 183u long; with 2 terminal setae, anterior seta as long as com- bined length of tarsus and tibia, posterior seta half that long; subterminal clawlike group of setae arising from posterior dorsal margin of tarsus (see fig. 3B); femur I and genu I each with a spined, heavy dorsal seta, other setae simple. Coxa I fragmented, medial seta arising from integument. Tarsi II-IV with a pair of PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 119 long slender ventral setae; large pad like membranous caruncle with thickened area resembling claws on ventral surface. Femora II, II] and IV and genua III and IV each with a heavy spined seta dorsally. Lengths, including coxa and caruncle, of legs II-IV: Il, 277»; II, 273u; IV, 2754. Gnathosoma. Tectum consisting of 2 lateral anterior extensions and a median extension which is split at the tip for about %4 its length, all extensions ciliated (fig. 2F). Palpal setae simple except for heavy, spined seta arising from dorsum of femur; remaining setae short except for 2 longer setae on tibia and 1 on tarsus; relative positions as shown in fig. 3D. Deutosternal groove without teeth; 3 pair of ventral setae, anterior hypostomal setae twice length of posterior pair; capitular setae about as long as posterior hypostomal setae; hypostomal process (internal malae) long, feathered, curving anterior to the long sclerotized corniculi. Chelicerae chelate, fixed digit bearing a trough-shaped excrescence which is smooth on the lateral margins and arrowhead-shaped at tip (fig. 2E). A short simple seta on each dorsolateral margin of gnathosoma about midway between idiosoma and palpal trochanter. MALE. Shape as in female, idiosoma 493u long, 3174 wide. (Measurements are the average of 3 specimens.) Dorsum. Plate entire, surrounded posteriorly and posterolaterally by integument as in female; furnished with 6 pairs of spined, tapering, capitate setae of type and arrangement as in female, anterior pair 1874 long, second pair 2804 long, median pair on integument 273 long. Venter (fig. 2G). Sternal, metasternal and ventral plates coalesced, 353, long, 1534 wide between coxae III, 924 between coxae IV, 167m at greatest width behind coxae IV; posterior margin of plate rounded, anterior margin irregular in outline, semisclerotized medially; plate lightly reticulated; 1 pair of pores at margin of plate at posterior level of coxae II; bearing 4 pairs of setae, anterior pair longest, relative lengths and positions as illustrated. Genital plate 148. long; transverse suture near posterior margin; internal kidney-shaped genital structure at each posterolateral margin of genital plate. Endopodal plates II, III and IV present and as in female. Anal plate wider than long, slightly bilobed from weak constriction medially; furnished with 2 setae of type occurring on dorsal plate; seta 257u long. Peritreme, peritremal plate and tritosternum as in female. Legs. Chaetotaxy as given in table 1. Leg I, 177u long. Leg II thickest, 2824 long; with a bilobed setigerous spur on venter of trochanter (fig. 2G). Leg III 273 long bearing prominent slightly curved spur on ventrum of trochanter with ventral seta of trochanter arising from the lateral margin of this spur (fig. 2G). Leg IV 290 long. Except for spurs on trochanter II and III chaetotaxy as in female. Tarsi II-I[V without claws; well developed padlike caruncle with basal part thickened, terminal part membranous; a pair of sclerotized clawlike structures at terminal end of tarsi (fig. 2C). Gnathosoma with general facies as in female. Movable digit of chelicera with excrescences as in female. DEUTONYMPH. Shape as in female; idiosoma 433u long, 291u wide. (Mea- surements are the average of 4 specimens.) Dorsum (fig. 2H). Plate entire, surrounded by integument; truncate posteriorly, otherwise with general shape of idiosoma. Dorsum bearing 6 pairs of spined, tapering, capitate setae, 2 pairs arising from dorsal plate, 4 pairs from integument; anterior pair of setae 159u long, second pair 260u long, relative lengths of remaining setae as shown. Venter (fig. 4B). Ventral plate 2534 long, 1124 at greatest width, shape as 120 PROC. ENT. SOC. WASH., VOL. 70, NO. 2, JUNE, 1968 Fig. 3. Brachytremella cartwrighti, n. sp. Female: A, venter; B. tarsus I, ven- tral view; C, tectum; D, ventral view of gnathosoma; E, chelicera. Male: F, venter. Protonymph: G, venter; H, dorsum. Larva: I, venter; J. dorsum. figured; anterior margin thickened and without reticulate pattern of remainder of plate; 1 pair of pores at margin where thickened area begins, a pair of porelike structures in plate behind coxae III; 5 pairs of ventral setae, fourth pair arising from ventral plate, remaining setae on integument, relative lengths as illustrated. Endopodal plates I], III and IV present; II widened anteriorly and bearing a pore, III rodlike, IV curving posteriorly around coxae IV. Integument bearing 5 pairs of pores, largest pair located medial of fragmented coxae I, position of others as shown. Anal setae 213 long; anal plate as illustrated. Tritosternum, peritreme and peritremal plate as in female. Legs. Chaetotaxy (see table 1) and caruncle as in female. Gnathosoma. General features as in female; chelicera with excrescence on fixed digit. PROTONYMPH. Idiosoma 400u long, 2804 wide (based on two specimens). Dorsum (fig. 21). Plate truncate posteriorly; not extending to margins of body. Bearing 5 pairs of spined, tapering, capitate setae; second and fourth pairs of setae on plate; first pair 140u long, second 250u long, relative lengths of other setae as shown. Venter (fig. 2J). Ventral plate without setae; shape as shown, 230u long, 1004 at greatest width; margins thickened and without reticulations which cover remainder of plate; one pair of pores at margin medial of coxae II. Integument bearing 5 pairs of pores and 4 pairs of setae, anterior pair of setae longest, posterior pair longer than second or third pairs. Endopodal plates as in deutonymph. Anal setae same type as on dorsum, 160u long; shape of anal PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 121 plate as illustrated. Tritosternum as in adults. Peritreme shorter than in adults, plate more oval, bearing 1 pore posterolateral of stigmata. Legs. Chaetotaxy discussed below. Pretarsi I-IV with caruncle as in female; genua and femora I, III and IV and femur II each with a single heavy, spined seta dorsally. Gnathosoma. General features as in adult. Palpal tibia without setae, genu with 1 dorsal setae (one less on each of these segments than in adults), ventrally no seta on trochanter, remaining segments as in deutonymph. Described from 3 females, 3 males, 4 deutonymphs and 3 pro- tonymphs. Holotype (female) data: Costa Rica, Heredia Province; 8-II-1965; J. Baird, coll.; from alcohol vial containing passalid beetles Passalus perparvulus Kuw. and Pseudacanthus tennis Kuw. One fe- male, 1 male, 4 deutonymphs, and 2 protonymph with same data as holotype. One female and 1 male from Turrialha, Costa Rica; March, 1965; M. V. Truitt, coll.; from passalid (Passalus) beetle. One deutonymph from Cerro Meurte, Costa Rica; 5-II-1965; M. V. Truitt, coll.; from passalid (Passalus) beetle. The male and female from Turrialha had longer dorsal setae and differed in other minor char- acters from Heredia Province specimens, but major taxonomic char- acters were as in the Heredia Province material. Holotype, male paratype, 2 deutonymphs and protonymph deposited in U. S. National Museum, Washington, D. C., remaining paratypes in collection of Department of Entomology, University of Georgia, Athens, Georgia. Brachytremella cartwrighti, n. sp. (Figs. 3A-J; 4A, C, D) This species is distinguished by having the second pair of dorsal setae shorter than other dorsal setae; in having 2 spined setae arising from femora III and IV; and in the adults the ventral plate widening slightly behind coxae IV with the last pair of setae on this plate located at the level of coxae IV. FEMALE. Oval shape; idiosoma 590 long, 4004 wide. (Measurements are the average of 2 specimens.) Dorsum (fig. 4C). Dorsal plate with a rounded extension between fourth pair of dorsal setae; surface of plate with faint polygonal reticulate patterns. Dorsal setae I-IV arising from margin of dorsal plate, the 2 remaining pairs arising from the integument posterior of the plate; second pair of setae shortest, 290u long, first pair 400u long, remaining setae approximately equal to first pair in length. Venter (fig. 3A). Sternal, metasternal and ventral plates coalesced, 430u long, 2184 at widest part between coxae III, 133u between coxae IV, 158u behind coxae IV; anterior sternal area semisclero- tized, sternal setae I arising from this area; 5 pairs of setae arising from the coalesced ventral plates, first pair longest, last pair next longest and remaining 3 pairs of equal length, positions of setae as shown; linear reticulations around genital opening, faint polygonal type pattern posteriorly; plate bearing 4 pairs of pores—1 pair between coxae II, a pair between coxae III and 2 pairs in the area of coxae IV. Genital plate rounded, 188 long, extending from level of anterior margin of coxae IV to middle of coxae IJ, thickened around free margins. Endopodal plate II fused to sternal plate for *4 its length, posterior %4 free; endopodal III 122; PROC. ENT. SOC. WASH., VOL. 70, No. 2, JUNE, 1968 Fig. 4. Brachytremella cartwrighti, n. sp. Deutonymph: = Se Date of of flies Alysia Aphaereta Aleochara percent puparia emerged ridibunda pallipes bimaculata _ parasitism 6/13 20 14 15.0 0.0 0.0 15.0 6/22 69 40 26.1 2.9 0.0 29.0 7/6 4 4 0.0 0.0 0.0 0.0 TANS 25 22, 12.0 0.0 0.0 12.0 po 38 29 7.9 ee 0.0 PALE IL THQ 8 5) 0.0 S45 0.0 Olan 8/25 143 ie: 7.0 27.3 5.6 39.9 9/13 ii, 3 0.0 42.9 0.0 42.9 9/28 1 0 0.0 0.0 0.0 0.0 10/4 5 0 0.0 40.0 60.0 100.0 Total 320 189 11.9 16.9 3.4 32.2 the temperature varied from 70 to 76 degrees F. and relative humidity was not controlled. Most of the parasites had died during the pupal stage and had to be dissected from the host puparia. Under field conditions, bucoprophilous fly larvae pupate in soil where the humidity is generally higher than laboratory air and is relatively constant. The change from field conditions to laboratory conditions may have had an adverse effect on A. ridibunda during the developmental period. Aphaereta pallipes (Say) and Aleochara bimaculata (Gravenhorst ) also used R. assidua as a host and may have limited the availability of R. assidua to parasitism by A. ridibunda. Table 1 shows parasitism of R. assidua by these three species of parasites. Field observations —A. ridibunda appears in the field at the end of the first week of June in central Missouri. The parasite was first ob- served in the field on June 10, 1966, and on June 7 the following year. A. ridibunda is active in the field from June to October, and specimens have been collected as late as October 10 in central Missouri. Activity of the parasite appears to end with cooler weather, probably after the first frost. This species probably overwinters as an immature form in the puparium of its host. Otherwise the adults would be expected to appear in the field earlier in the spring. Field observations showed that A. ridibunda spends considerable time searching for hosts. This species was observed on many occasions searching the manure, con- tinually probing the surface with the ovipositor, sometimes for two hours or more with little interruption. A. ridibunda was never observed in association with decaying meat and appeared to have little interest in this medium when adults were placed on meat in the laboratory. Lindquist (1940) stated that A. ridibunda at Uvalde, Texas was never observed on cow manure and appeared to have little interest in larvae living in dung when confined in cages. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 207 Present status of Alysia spp—Because A. ridibunda in Missouri did not occupy the same ecological niche as the species reported as A. ridibunda by Lindquist (1940), the status of specimens referable to A. ridibunda was investigated. In December, 1966, a series of 39 speci- mens collected in the Missouri studies was sent to the Systematic Entomology Laboratory of the USDA in Beltsville, Maryland for de- termination. They were identified as Alysia n. sp. near ridibunda Say, a species very similar to A. ridibunda. The determination was made by P. M. Marsh and C. F. W. Muesebeck. However, further correspond- ence with Marsh in November, 1967 revealed that the 1966 determina- tion was in error and that specimens from Missouri were Alysia ridibunda Say. The specimens in Texas reported on by Lindquist and Roberts were a distinct, but closely related species, recently described as A. cariosa by Marsh (1968). Specimens in the National Museum Collection under the name ridibunda were found to be a mixture of the two species and many of the determinations made previously from these museum specimens are incorrect. Refer to Marsh (1968 ) for distinguishing the two species. According to Marsh, it would appear that all literature references to A. ridibunda as a parasite of carrion-infesting flies are not that species, but that all references to A. ridibunda as a parasite of dung-feeding flies are actually ridibunda. Hence, Howard (1900), Cates (1964), and Thomas (1967) were re- ferring to Alysia ridibunda Say, while Lindquist (1932, 1940) and Roberts (1935) were referring to A. cariosa Marsh. State records for A. ridibunda in the National Museum Collection include the following: Missouri, Virginia, Maryland, District of Colum- bia, Arkansas, Georgia, Kansas, New Jersey, Illinois, Pennsylvania, Texas, and West Virginia. A. cariosa Marsh is represented by speci- mens from Texas and Arizona. There are also two specimens of A. ridibunda from Virginia reared from Ravinia assidua. SUMMARY During 1966 and 1967, larvae of necrophilous and bucoprophilous muscoid Diptera were collected to determine the host of Alysia ridibunda Say in Missouri. Although Lindquist (1932) had reported A. ridibunda as a blowfly parasite in the southwestern United States, this species was not recovered from carrion-infesting flies in Missouri. A. ridibunda, however, was recovered from Ravinia assidua (Walker ) reared in cow manure. During the summer of 1967, 38 A. ridibunda were recovered from 320 R. assidua. The highest parasitism of R. assidua in one collection was 26% on June 22. The average parasitism over the season was nearly 12%. R. assidua was the only host recorded for A. ridibunda. Because A. ridibunda in Missouri did not occupy the same ecological niche as the species reported as A. ridibunda by Lindquist, the status 208 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 of A. ridibunda was investigated. All references to A. ridibunda in the literature do not apply to the same species. The species in Missouri associated with cow manure is ridibunda. The species associated with carrion reported by Lindquist (op. cit.) and Roberts (1935) is a dis- tinct but closely related species. This is based mostly on differences in size and certain morphological features, but also on host and habitat preference. Acknowledgment.—We are grateful to Paul M. Marsh of the U. S. Department of Agriculture for identifying A. ridibunda and for the information regarding the two similar species of Alysia. REFERENCES Cates, M. D. 1964. Insect competition in and utilization of fresh field manure. Unpublished Master’s Thesis. The University of Missouri, Columbia. Howard, L. O. 1900. A contribution to the study of the insect fauna of human excrement. Proc. Wash. Acad. Sci. 2:541—604. Lindquist, A. W. 1932. Alysia ridibunda Say, parasitic on blowfly larvae. J. Econ. Ent. 25(2):414—415. 1940. The introduction of an indigenous blowfly parasite, Alysia ridibunda Say, into Uvalde County, Texas. Ann. Ent. Soc. Amer. 33(1):104—112. Marsh, P. M. 1968. A new species of Alysia parasitic on carrion-infesting flies. Proc. Ent. Soc. Wash. 70(3):208—-210. Roberts, R. A. 1935. Some North American parasites of blowflies. J. Agric. Res. 50(6) :479-494. Say, T. 1859. A description of some new species of Hymenoptera of the United States. (From Contributions of the Maclurian Lyceum to the Arts and Sciences v. 1, pp. 67-83 Philadelphia (1828). In Leconte, J. L., The com- plete writings of Thomas Say on the entomology of North America pp. 373- 375. illus. New York, London. Thomas, G. D. 1967. Natural enemies of the face fly Musca autumnalis De Geer in Missouri. Unpublished Ph.D. Dissertation. The University of Missouri, Columbia. A NEW SPECIES OF ALYSIA PARASITIC ON CARRION-FEEDING FLIES (HYMENOPTERA: BRACONIDAE) PauL M. Marsn, Systematic Entomology Laboratory, Entomology Research Division, Agr. Res. Serv., USDA? The following species is described in connection with studies being made on dung-feeding flies and their parasites by L. P. Burgess, Jr. and C. W. Wingo (see this issue, page 204) at Columbia, Missouri. This species, Alysia cariosa Marsh, is similar to A. ridibunda Say, and ‘Mail address: c/o U. S. National Museum, Washington, D.C. 20560. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 209 Fig. 1, Alysia cariosa, n. sp., wing venation. Fig. 2, A. ridibunda Say, wing venation. the two have been confused in past literature. A. ridibunda appears to parasitize dung-feeding flies, whereas A. cariosa attacks only car- rion-infesting flies. Hence, several earlier biological papers in which A. ridibunda was mentioned as a parasite of blow flies in carrion (e. g. Lindquist, 1932 and 1940; Roberts, 1935) actually apply to A. cariosa. A long series of the material used by Lindquist is in the U. S. National Museum and is included in the type series of A. cariosa. The most important characters which distinguish A. cariosa from A. ridibunda are its generally larger size, the weak or absent notauli, the tridentate rather than bidentate mandibles, the parallel rather than converging carinae on the first abdominal tergum, the wing venation (compare figs. 1 and 2), and completely orange-red abdomen in the male. The male of ridibunda has the abdomen red with the apex black. Alysia cariosa, n. sp. (Fig. 1) Female.—Length of body, 4-7.5 mm; ovipositor, 1-2.5 mm. Thorax and abdomen orange-red; head, including base of mandibles, deep maroon or nearly black, mandibles often lighter at apex; proepisternum and legs, including coxae, deep maroon to dark brown; wings completely dusky, veins dark brown, stigma often light brown. Head entirely smooth, nearly 1.5 times as wide as high and slightly wider than thorax; face with a weakly raised median line from antennae to clypeus; frons with a shallow pit between lateral ocelli; mandibles with 3 distinct teeth, rugose on at least basal 1%; antennae 39- to 49-segmented, first 2 flagel- lar segments about equal in length, but the second often slightly longer than the first, segments in apical % of flagellum about as wide as long. Thorax smooth and polished; notauli absent or, at most, represented by short, weak indentations anteriorly; scutellar furrow with 1 median cross carina; mesopleural and subalar furrows smooth; propodeum with strong median and lateral carinae, the median one forked apically; legs slender; last segment of tarsus longer than third and fourth combined; tarsal claws large. Wing venation as in fig. 1; stigma about 7 or 8 times longer than wide; radius arising from apical 4% of stigma; recurrent vein usually entering base of second cubital cell, often interstitial with first intercubitus; 210 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 nervulus interstitial with basal vein. Abdomen smooth and polished; first tergum about as long as apical width, with 2 median, parallel carinae which extend posteriorly from base to apical ¥% of tergum; ovipositor at most as long as abdomen beyond first tergum. Male.—Essentially as in female; length of body, 5.5-9 mm; first abdominal tergum occasionally rugose between carinae. Holotype Female—TEXAS: Uvalde, flesh and green bottle flies, Dec., ’35, A. W. Lindquist. USNM 69957. Paratypes.—ARIZONA: Duncan, 2 22, 1 ¢, September 22, 1931, parasite of blow fly, A. W. Lindquist, Bishop No, 17273. TEXAS: 23 22,57 44, same data as holotype. All paratypes are deposited in the U.S. National Museum. REFERENCES Lindquist, A. W. 1932. Alysia ridibunda Say, parasitic on blowfly larvae, J. Econ. Ent. 25:414-415. 1940. The introduction of an indigenous blowfly parasite, Alysia ridibunda Say, into Uvalde County, Texas. Ann. Ent. Soc. Amer. 33:104—112. Roberts, R. A. 1935. Some North American parasites of blowflies. J. Agr. Res. 50:479-494, WATER MITES OF THE GENUS STYGOMOMONIA IN NORTH AMERICA (Acarina, MOMONIDAE)* Davin R. Cook, Department of Biology, Wayne State University, Detroit, Michigan 48202 This paper treats eight species of Stygomomonia Szalay known from North America, six of which are described as new. The two previously described species are S. riparia Habeeb and S. moodyi Mitchell. Holo- types and allotypes will be deposited in the Field Museum of Natural History (= Chicago Natural History Museum). In presenting measure- ments in this paper, those of the holotype and allotype are given first. If a series of specimens is available, the range of variation is given in parentheses following the measurements of the primary types. Members of the genus Stygomomonia exhibit considerable intra- specific variation in number of heavy setae on the fourth coxae and in shape and distance apart of the suture lines of the third and fourth coxae. The structure of the palp is very similar in all species of Stygomomonia s. s. and is therefore nearly useless in defining species. The author has found proportions of the distal segments of the first leg, structure of the peripheral thickening of the dorsal shield, and _ ‘Contribution No. 200 from the Department of Biology, Wayne State University. Supported by Grants G-9042 and GB-3717 from the National Science Foundation. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 211 degree of fusion of the posterior glandularia platelets of the venter to be the most stable characters which seem to be of taxonomic value. Since most of the species diagnoses are based on a combination of characters, a key to species is provided as an aid in identification. The genus is primarily a group confined to interstitial waters, but where abundant, specimens are occasionally taken in “surface water” collections. Since the subsurface water habitat has been poorly sampled in many areas of the world, the true distribution of the genus is probably unknown. Species of Stygomomonia have been collected in Europe, Japan and North America. A rather unusual species from New Zealand has also been tentatively assigned to this genus. It is interesting that extensive collections of interstitial water mites made by the author in Western India were negative as far as Stygomomonia was concerned. It is here proposed that the genus Stygomomonia be divided into three subgenera, two of which have a Holarctic distribution and a third which is known only from New Zealand. Subgenus Stygomomonia Szalay Diagnosis: Characters of the genus Stygomomonia; anterior coxal groups (first and second coxae) separated by articular membrane from the third coxae; genital field of male subterminal in position; dorsal shield entire; excretory pore not fused with the posterior glandularia platelets. Subgenotype: Stygomomonia latipes Szalay. Discussion: The separated anterior coxal groups are diagnostic for the typical subgenus. Members of Stygomomonia s. s. are known from Europe, Japan and North America. Subgenus Allomomonia, n. subgen. Diagnosis: Characters of the genus Stygomomonia; anterior coxal groups (first and second coxae) fused with the third coxae; genital field of male located at extreme posterior end of ventral shield; dorsal shield entire; excretory pore not fused with the posterior glandularia platelets. Subgenotype: Stygomomonia moodyi Mitchell. Discussion: The fused condition of the coxae and the entire dorsal shield are characters which will separate Allomomonia from other subgenera of Stygomomonia. The new subgenus has representatives in Japan and North America. Subgenus Neomomonia, n. subgen. Diagnosis: Characters of the genus Stygomomonia; anterior coxal groups (first and second coxae) fused with the third coxae; dorsal shield divided into anterior and posterior sclerites; dorsal shield closely flanked by several pairs of platelets; excretory pore fused with the enlarged posterior glandularia platelets. Subgenotype: Stygomomonia torquipes Hopkins. Discussion: The new subgenus is based on a female specimen described by Hopkins (1966) from New Zealand. It appears to be a rather divergent member of the genus Stygomomonia but, until the male is known, its placement in this genus should be regarded as tentative. bo —_ bo PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Figs. 1-3, Stygomomonia moodyi Mitchell: 1, ventral shield, ¢; 2, ¢ palp; 3, dorsal shield, ¢. Fig. 4, S. separata, n. sp., distal segments, Ist leg, 6. Figs. 5, 6, 9, S. occidentalis, n. sp.: 5, distal segments, 1st leg, 2; 6, ventral shield, ¢; 9, dorsal shield, ¢. Fig. 7, S. neomexicana, n. sp., distal segments, Ist leg, @. Fig 8, S. pallida, n. sp., distal segments, Ist leg, @. a PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 213 Kry to NortH AMERICAN SPECIES OF Stygomomonia 1. Anterior coxal groups (first and second coxae) fused with the third coxae (fig. 1); claw of first leg short and decidedly recurved (fig 30) (subgenus PAU GINO Osta) meee cree 3a ew eee ee ae moodyi Mitchell Anterior coxal groups separated from the ‘hid coxae (fig. 6); claw of first leg relatively long and narrow, and only slightly recurved (fig. 4) (sub- PEMUSH OLY ZOMOMON IC) mene ee eeee ee ook eet te eee ee 9) 2. Four pairs of genital acetabula present (figs. 18, 25) — riparia Habeeb Three pairs of genital acetabula present (figs. 21, 23) 0 3 3. The two pairs of glandularia posterior to the genital field located on small sclerites which are either separated (fig. 10) or only lightly fused (fig. ASS) * emcee ats soa kD, alae lL uN Dee a cae cea 4 The two pairs of glandularia posterior to the genital field fgenteae on ache tively large sclerites which are broadly fused on their respective sides (ENG (Gi): Sintec d panic elec AE LR tag Ne es 5 4. Dorsal projection on basal uaéiem of I-Leg-6 somewhat proximal in Falatiog to attachment site of segment (fig. 4) — Aucune la separata, n. sp. Dorsal projection on basal portion of I-Leg-6 distal in relation to the attach- ment site of segment (fig. 7) pratt xo OO ofa otha rh neomexicana, Nn. sp. 5. Dorsal projection on basal portion of I-Leg-6 euioer proximal in relation to) attachment site of segment (fig. 16) —... plana, n. sp. Dorsal projection on basal portion of I-Leg-6 distal in relation to the attach- MENtESILGLOL SCOMENEN MOS Or GH oes se ee 6 6. I-Leg-4 greatly expanded at distal end (fig. 22) archers eS: I-Leg-4 only slightly expanded at distal end (figs. 5, 8) _------------ ii 7. I-Leg-5 more than twice the length of I-Leg-6 (fig. 5); peripheral thicken- ing of dorsal shield not extending medially towards posterior glandularia (Unies Gy)! ase te Be a acon i Ue LAE Se Ae _ occidentalis, n. sp. I-Leg-5 less than twice the length of I-Leg-6 (fig. 8); peripheral thicken- ing of dorsal shield extending medially towards posterior glandularia (hates OLS RS Wa ieee Meee” oes gee ree aes ee eet Se pallida, n. sp. Stygomomonia (Allomomonia) moodyi Mitchell (Figs. 1-3, 17, 30) Stygomomonia moodyi Mitchell, 1959, Trans. Amer. Microsc. Soc. 78:154. Male: Dorsal shield 426u-471u in length, 3124-348 in width; dorsal shield with ill-defined ridges near periphery (fig. 3); ventral shield, including posterior glandularia platelets 441u—471u in length, 3504-3954 in width; anterior coxal groups solidly fused with the third coxae; 2 small setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 124u-156u in width; 3 pairs of genital acetabula; genital field terminal (fig. 1); posterior glandularia of venter fused on their respective sides into elongated platelets which are slightly constricted in middle, these platelets 694-79 in greatest dimension; excretory pore without associated sclerotization; eyes reduced: integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 144-174; P-II, 42u-48; P-III, Q1p-24u; P-IV, 35u-39u; P-V, 29u-3lu; fig. 2 illustrates the proportions and chaetotaxy of the palp; dorsal lengths of the distal segments of the first leg: 214 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Figs. 10-12, Stygomomonia separata, n. sp.: 10, ventral shield, ¢; 11, dorsal shield, 4; 12, genital field region, 9. Figs. 13, 15, 16, S. plana, n. sp.: 13, ventral shield, 2; 15, dorsal shield, 2; 16, distal segments, Ist leg, 9. Fig. 14, S. occi- dentalis, n. sp., 6 palp. Fig. 17, S. moodyi Mitchell, genital field region, ?. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 2S I-Leg-4, 8lu—-90u; I-Leg-5, 104u-12lu; I-Leg-6, 6]lu-62u; structure of first leg similar to female; swimming hairs absent. Female: Dorsal shield 426u-483. in length, 3194-380 in width: dorsal shield similar to that illustrated for the male; ventral shield, including posterior glandularia platelets 4414-5174 in length, 3504-441 in width; anterior coxal groups solidly fused with the third coxae; typically with 2 (but in one instance 3) small setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield (fig. 17); genital field 1354-1874 in width; the individual acetabular plates 334-52 in width; 3 pairs of genital acetabula; posterior glandularia as described for male, these 73u—-80u in greatest dimension; excretory pore without associated sclerotization; eyes reduced; integumental pigment absent. Dorsal lengths of the palpal segments: P-I, 14u-16u; P-II, 45u-48u; P-III, 21u-24u; P-IV, 364-384; P—V, 28u—-3ly; structure of palp similar to male; dorsal lengths of the distal segments of the first leg: I-Leg-4, 83u-90u; I-Leg-5, 1184-133; I-Leg-6, 62u-66u; claw at tip of first leg recurved; fig. 30 illustrates the modification of the first leg; swimming hairs absent. Material Examined: 1 female, from a gravel bar in the Smith River (ap- proximately 18 miles east of Gardiner), Douglas County, Oregon, August 12, 1961; 33 males, 13 females, collected in a gravel bar in Ten Mile Creek approximately 7 miles of Yachats, Lane County, Oregon, August 13, 1961; 2 males, taken in a gravel bar in Satus Creek (approximately 3 miles south of Yakima County line), Klickitat County, Washington, August 18, 1967; 1 female, taken in bottom gravels in Battle Creek 4 miles west of Viola, Shasta County, California, July 30, 1966; 1 male, from gravel deposits in a small stream near Warm Springs, Bath County, Virginia, July 20, 1963. Mitchell originally recorded this species from Addison, Chittenden, and Washington Counties, Vermont. Discussion: The species S. kantoensis described by Imamura (1959) from Japan also has the anterior coxal groups fused with the third coxae. However, the latter species possesses numerous enlarged setae on the fourth coxae as illustrated in figs. 6 and 10, and the posterior glandularia of the venter are all separated from each other. S. moodyi possesses two very small setae on the fourth coxae (fig. 1) and the posterior glandularia of the venter are fused on their respective sides. The western populations of this species tend to be proportionally narrower and the genital field proportionally less wide than the eastern representatives. However, these differences are not clear cut and do not seem to justify subspecific separation. Stygomomonia (Stygomomonia) riparia Habeeb (Figs. 18, 20, 24, 25) Stygomomonia riparia Habeeb, 1957, Leaflets of Acadian Biology 15:8. Stygomomonia riparia: Mitchell, 1959, Trans. Amer. Microsc. Soc. 78:156. Male: Dorsal shield 410u-524u in length, 2894-3654 in width; peripheral thickening of dorsal shield extending medially towards posterior glandularia (fig. 20); ventral shield, including posterior glandularia platelets 4184-502 in length, 334u-408u in width; anterior coxal groups separated by articular membrane from 216 PROC. ENT. SOC. WASH., VOL. 70, No. 3, SEPTEMBER, 1968 Figs. 18, 20, 24, 25, Stygomomonia riparia Habeeb: 18, ventral shield, 6; 20, dorsal shield, ¢; 24, distal segments, Ist leg, 6; 25, genital field region, °. Fig. 19, S. pallida, n. sp., 3 palp. Figs. 21-23, 26, S. ozarkensis, n. sp.: 21, genital ae poe Q; 22, distal segments, Ist leg, ¢; 23, ventral shield, ¢; 26, dorsal shield, ¢. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 21 “I the third coxae; 5 to 7 enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 118y—-141y in width; typically with 4 pairs of genital acetabula but occasionally with only 3 acetabula on one side, 4 on the other side; posterior glandularia fused on their respective sides into oval platelets (fig. 18); these platelets 83u-lll in greatest dimension: excretory pore without associated sclerotization; eyes reduced; integumental pig- ment absent. Dorsal lengths of the palpal segments: P-I, 154-18; P-Il, 424-45; P-III, 23u—-27u; P-IV, 42u-52u; P—V, 29u—36y; structure of palp similar to that illustrated in fig. 19; dorsal lengths of the distal segments of the first leg: I-Leg-4, 70u—90u; I-Leg-5, 156u-194u; I-Leg-6, 76u-80u; greatest heights of distal segments of first leg: I-Leg-4, 28u-35u; I-Leg-5, 42u-52u; I-Leg-6, 38y-45u; claw at tip of first leg 48u-52u in length; fig. 24 illustrates the proportions and chaetotaxy of the distal segments of the first leg; swimming hairs absent. Female: Dorsal shield 456u—525u in length, 3194-358 in width; dorsal shield similar to male; ventral shield, including posterior glandularia platelets 4494-517 in length, 350u—418. in width; anterior coxal groups separated by articular mem- brane from the third coxae; 4 to 7 enlarged setae present on each side of the fourth coxae posterior to insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield (fig. 25); genital field 1384-159 in width; the individual acetabular plates 354-454 in width; typically with 4 pairs of genital acetabula, but one individual had only 3 acetabula on one side, 4 on the other side; posterior glandularia as described for male, 97u— 124u in greatest dimension; excretory pore without associated sclerotization; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 174-19»; P—II, 42u—52u; P-III, 23u-27u; P-IV, 42u-48u; P-V, 3lu—35u; dorsal lengths of the distal segments of the first leg: I-Leg-4, 76u-86u; I-Leg-5, 173u-201u; I-Leg-6, 76u-86u; great- est heights of distal segments of first leg: I-Leg-4, 29u-35y; I-Leg-5, 45y-59u; I-Leg-6, 38u-46u; claw at tip of first leg 52u-55u in lengths; structure of palp and first leg similar to male. Material Examined: (All collections from sand and gravel deposits of streams ) 5 males, 4 females, 2 nymphs, from the St. Francis River near Sam A. Baker State Park, Wayne County, Missouri, July 9, 1960; 8 males, 5 females, same data on June 24, 1961; 8 males, 9 females, same data on June 26, 1961; 3 males, 4 nymphs, from a stream at Patterson, Wayne County, Missouri, July 11, 1960; 4 females, 2 nymphs, same data on June 25, 1961; 12 males, 5 females, 1 nymph, collected in the Black River northwest of Piedmont, Reynolds County, Missouri, July 8, 1960; 5 males, 10 females, same data on June 26, 1961; 1 female, from the Current River at Big Spring State Park, Carter County, Missouri, July 12, 1960; 5 males, 3 females, 1 nymph, taken in the Jacks Fork River at Alley Spring State Park, Shannon County, Missouri, July 13, 1960; 11 males, 4 females, 1 nymph, same area where a cold spring joins the river, July 14, 1960; 2 females, 3 nymphs, from the Meramec River 11 miles northeast of Salem, Dent County, Missouri, July 19, 1960; 1 male, 2 females, same data on July 1, 1961; 4 males, collected in the Meramec River at Cook Station, Crawford County, Missouri, July 23, 1960; 1 male, 1 female, from the Eleven Point River near Greer Spring, Oregon County, Missouri, June 27, 1961; 2 females, from the Roaring River at Eagle Rock, Barry County, Missouri, July 6, 1961; 1 male, 1 nymph, taken in the King’s River, Carroll 218 PROG. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Figs. 27, 29, Stygomomonia pallida, n. sp.; 27, ventral shield, ¢; 29, dorsal shield, ¢. Figs. 28, 31, S. neomexicana, n. sp.: 28, ventral shield, 9; 31, dorsal shield, 2. Fig. 30, S. moodyi Mitchell, distal segments, Ist leg, 9. Fig. 32, S. plana, n. sp., genital field area, 6. Fig. 33, S. occidentalis, n. sp., genital field area, ¢. County, Arkansas, July 21, 1960; 8 males, 1 female, 1 nymph, found in a small stream 1 mile north of Albion, Pushmataha County, Oklahoma, July 9, 1961; 1 male, 1 female, from a small stream 4 miles south of Attica, Warren County, Indiana, July 12, 1964; 1 female, taken in a stream near Lynch, Harlan County, Kentucky, July 15, 1964; 1 female, collected in a stream near Fredonia, Licking County, Ohio, June 18, 1967; 1 male, from a stream at Cedar Falls State Park, PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 219 Hocking County, Ohio, June 18, 1967; 1 male, 3 females, from a large stream near Haynes, Hocking County, Ohio, June 18, 1967; 9 males, 6 females, taken in a tributary of the Jackson River approximately 1.5 miles south of Bacova, Bath County, Virginia, May 19, 1961; 1 male, 1 female, found in a stream near Griffen, Hamilton County, New York, August 19, 1964; 3 males, 1 female, from the North Branch of the Meduxnekeag River, Aroostook County, Maine, August 28, 1964; 1 female, collected in a small stream near Limestone, Victoria County, New Brunswick, August 26, 1964; 1 female, from the Agawa River near the southern border of Lake Superior Provincial Park, Algoma District, Ontario, August 28, 1965. Discussion: Habeeb (1957) originally described this species from the Salmon River, Victoria County, New Brunswick. Mitchell (1959) collected this species in Chittenden County, Vermont. The presence of four pairs of acetabula will distinguish this species from all other members of the genus. Stygomomonia (Stygomomonia) ozarkensis, n. sp. (Figs. 21-23, 26) Male: Dorsal shield 532 in length, 3954 in width; peripheral thickening of dorsal shield not extending medially in direction of posterior glandularia (fig 26); ventral shield, including posterior glandularia platelets 516u in length, 441, in width; anterior coxal groups separated by articular membrane from the third coxae; 5 to 7 enlarged setae present on each side of fourth coxae posterior to the insertions of the fourth legs; genital field 1284 in width; 3 pairs of genital acetabula; posterior glandularia fused on their respective sides into oval platelets (fig. 23); these platelets 104 in greatest dimension; excretory pore without associated sclerotization; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 17; P-II, 554; P-III, 274; P-IV, 48; P-V, 35y; structure of palp similar to that illustrated in fig. 19; dorsal lengths of the distal segments of the first leg: I-Leg-4, 8lu; I-Leg-5, 197u; I-Leg-6, 80u; greatest height of distal segments of first leg: I-Leg-4, 41u; I-Leg-5, 59u; I-Leg-6, 41y; claw at tip of first leg 48u in length; fig. 22 illustrates the propor- tions and chaetotaxy of the distal segments of the first leg; swimming hairs absent. Female: Dorsal shield 547 in length, 441 in width; dorsal shield similar to male; ventral shield, including posterior glandularia platelets 578 in length, 501 in width; anterior coxal groups separated by articular membrane from the third coxae; 5 to 6 enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield (fig. 21); genital field 1664 in width; 3 pairs of genital acetabula; posterior glandularia platelets as described for male; post- genital sclerite 201u in width; excretory pore without associated sclerotization; eyes reduced; integumental pigment absent. Holotype: Adult male, from a gravel bar in a small stream on Route 116 approximately 1 mile west of Colcord, Delaware County, Okla- homa, July 7, 1961. Allotype: adult female, same data as holotype. Discussion: The combination of three pairs of acetabula, lack of a 220 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 medial extension of the peripheral thickening of the dorsal shield, relatively short I-Leg-6 and greatly expanded I-Leg-4 are diagnostic for the present species. Stygomomonia (Stygomomonia) pallida, n. sp. (Figs. 8, 19, 27, 29) Male: Dorsal shield 517u(464y-517) in length, 350u4(3194-350u) in width; peripheral thickening of dorsal shield extending medially in direction of posterior glandularia (fig. 29), but not always as pronounced as in illustration; ventral shield, including posterior glandularia platelets 5174(456y4-517u) in length, 388u(373u—-410u) in width; anterior coxal groups separated by articular mem- brane from the third coxae; 5(4—-6) enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 1284 (1184- 130u) in width; 3 pairs of genital acetabula, these 114-17» in diameter; posterior glandularia fused on their respective sides into oval platelets (fig. 27); these platelets 1074(96u-107~) in greatest dimension; excretory pore without associated sclerotization; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 15u(154-18); P-II, 48u(46y— 52u); PIII, 27u(24u-28u); P-IV, 45u(43u-48u); P-V, 33u(33u-38u); fig. 19 illustrates the proportions and chaetotaxy of the palp; dorsal lengths of the distal segments of the first leg: I-Leg-4, 86u(80u-86u); I-Leg-5, 194u( 187-204, ); I-Leg-6, 104u (984-104); greatest height of distal segments of first leg: I-Leg-4, 34u(31u-34u); I-Leg-5, 52u(52u-55u); I-Leg-6, 45u(42u—-45u); claw at tip of first leg 69u(66u-69u) in length; fig. 8 shows the distal segments of the first leg; swimming hairs absent. Female: Unknown. Holotype: Adult male, taken in a sand and gravel bar in a tributary of the Jackson River approximately 1.5 miles south of Bacova, Bath County, Virginia, May 19, 1961. Paratypes: 5 males, same data as holotype. Discussion: The combination of three pairs of genital acetabula, presence of medial extensions of the peripheral thickening of the dorsal shield, and relatively long I-Leg-6 (over one-half length of I-Leg-5) will distinguish the present species from other members of the genus. Stygomomonia (Stygomomonia) plana, n. sp. (Bigss 1315. 16432) Male: Dorsal shield 456u in length, 312u in width; structure of dorsal shield similar to that illustrated and described for female; ventral shield, including posterior glandularia platelets 454 in length, 334 in width; anterior coxal groups separated by articular membrane from the third coxae; 6 or 7 enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 904 in width; 3 pairs of genital acetabula, these approximately 10u in diameter; posterior glandularia fused on their respective sides into oval glandularia platelets (fig. 32); these platelets 107 in greatest dimension; ex- PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Ds bo — cretory pore without associated sclerotization; eyes reduced; integumental pig- ment absent. Dorsal lengths of the palpal segments: P-I, 174; P-II, 484; P-III, 27u; P-IV, 45u; P-V, 33u; palp similar to that illustrated in fig. 14; dorsal lengths of the distal segments of the first leg: I-Leg-4, 694; I-Leg-5, 173; I-Leg-6, llly; claw at tip of first leg 694 in length; structure of first leg similar to female; swimming hairs absent. Female: Dorsal shield 445u(441u—502u) in length, 304u(289u-349) in width; peripheral thickened area of dorsal shield not extending medially in region of posterior glandularia (fig.15); ventral shield 464u(434u-517u) in length, 334u (3274-3804) in width; anterior coxal groups separated by articular membrane from the third coxae; 5 to 7 enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and post- genital sclerite separated from the ventral shield (fig. 13); genital field 118u(111u— 121) in width; the individual acetabular plates 3lu(28u-3lu) in width; 3 pairs of genital acetabula; posterior glandularia as described for the male; these glandularia platelets 1054(100u-114u) in greatest dimension; excretory pore without associated sclerotization; eyes reduced; integumental pigment absent. Dorsal lengths of the palpal segments: P-I, 154(154-17u); P-II, 45u(42u-51,); P-III, 24u(24u-29u); P-IV, 42u(42u—-46u); P-V, 3lu(3lu—-33u); dorsal lengths of the distal segments of the first leg; I-Leg-4, 78u(69u-80u); I-Leg-5, 170u (170u-2074); I-Leg-6, 1llu(104u-134u); greatest height of distal segments of first leg: I-Leg-4, 29u(28u-29u); I-Leg-5, 48u(48u-52u); I-Leg-6, 48u(45u— 48u); claw at tip of first leg 80u(69u—-93u); dorsoproximal portion of I-Leg-6 extending slightly proximally (fig. 16); swimming hairs absent. Holotype: Adult male, taken in gravel deposits in the South Fork of the Trinity River near Forest Glen, Trinity County, California, July 30, 1966. Allotype: adult female, same data as holotype. Paratypes: 1 female, same data as holotypes; 1 male, found in a gravel bar in Salmon Creek on Highway no. 1, Monterey County, California, July 23, 1966; 1 female, from gravel deposits in the Big Sur River at Big Sur Camp Grounds, Monterey County, California, July 23, 1966; 1 female, collected in a gravel bar in the Van Duzen River 16 miles east of Bridgeville, Humboldt County, California, July 30, 1966; 1 female, found in a gravel bar in Satus Creek (on US 97 approximately 3 miles south of the Yakima County line), Klickitat County, Washing- ton, August 18, 1961. Discussion: The combination of three pairs of acetabula, lack of medial extensions of the peripheral thickening of the dorsal shield, enlarged posterior glandularia platelets, and comparatively large I-Leg-6 will separate the present species from all other members of the genus. Stygomomonia (Stygomomonia) occidentalis, n. sp. (igsw55 659145 33)) Male: Dorsal shield 410u(410u-547u) in length, 289u(289u-395u) in width; peripheral thickening of dorsal shield not extending medially in region of posterior bo bo bo PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 glandularia (fig. 9); ventral shield, including posterior glandularia platelets 426u(426u-547u) in length, 3274(327u-449.) in width; anterior coxal groups separated by articular membrane from the third coxae; 3-5(3-6) enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 98u(98u-117) in width; 3 pairs of genital acetabula, these 10u(8u-10u) in diameter; posterior glandularia fused on their respective sides into oval platelets (fig. 6); excretory pore without associated sclerotization; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 14u( 144-18); P-II, 47u( 47-55); P-III, 22u(22u-27u); P-IV, 42u(42u-48u); P-V, 29u(29u-38u); fig. 14 illus- trates the proportions and chaetotaxy of the palp; dorsal lengths of the distal segments of the first leg: I-Leg-4, 694(69u-83u); I-Leg-5, 159u(159u-194,y); I-Leg-6, 73u(73u-93); greatest height of the distal segments of the first leg: I-Leg-4, 28u(28u-32u); I-Leg-5, 42u(42u-52u); I-Leg-6, 3lu(31lu-38); claw at tip of first leg 48u(48u-62u) in length; structure of first leg similar to female; swimming hairs absent. Female: Dorsal shield 441u(441u—547) in length, 304u(304~-395.) in width; dorsal shield similar to that of male; ventral shield, including posterior glandularia platelets 440u(440u-539u) in length, 3344(334u-440u) in width; anterior coxal groups separated by articular membrane from the third coxae; 5—-7(4-7) enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield (fig. 33); genital field 121u(121u-159u) in width; the individual acetabular plates 26u(26u-34u) in width; 3 pairs of genital acetabula, these 10u4(9u—-12u) in diameter; postgenital sclerite 1454(1454-200u) in width; posterior glandularia as described for the male; excretory pore without associated sclerotization; eyes re- duced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 13u(134-17); P-II, 46u(46u—-55,); P-III, 24u(24u-284); P-IV, 43u(43u-50u); P—V, 32u(32u-41u); palp similar to male; dorsal lengths of the distal segments of the first leg: I-Leg-4, 7lu(71lu— 90u); I-Leg-5, 1734(170u-21lu); I-Leg-6, 76u(76u-100u); greatest height of the distal segments of the first leg: I-Leg-4, 28u(28u-35u); I-Leg-5, 42u(42u— 55u); I-Leg-6, 3lu(3lu-38u); claw at tip of first leg 46u(46u-66u) in length; fig. 5 illustrates these segments; swimming hairs absent. Holotype: Adult male, taken in a sand bar in the Rogue River (1 mile south of the Douglas County line), Jackson County, Oregon, August 11, 1961. Allotype: adult female, same data as holotype. Paratypes: 1 female, same data as holotype; 3 males, from a gravel bar in Swauk Creek at Swauk Creek Camp Ground, Kittitas County, Washington, August 19, 1961; 2 males, 11 females, collected in a gravel bar in the Fisher River 30 miles southeast of Libby, Lincoln County, Montana, August 22, 1961; 1 male, 1 female, from a gravel bar in the Swan River approximately 6 miles southeast of Bigfork, Lake County, Montana, August 25, 1961; 1 male, from a gravel bar in Salmon Creek on Highway no. 1, Monterey County, California, July 23, 1966; 1 male, 2 females, taken in a gravel bar in Moccasin Creek (on State Road no. 49 six miles southeast of State Road no. 120), Tuolumne County, California, July 24, 1966. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 2: bo (ov) Discussion: The combination of three pairs of acetabula, lack of medially-directed extensions of the peripheral thickening of the dorsal shield, enlarged posterior glandularia platelets, and relatively short I-Leg-6 will separate S. occidentalis from all other species of the genus found in North America. Stygomomonia (Stygomomonia) separata, n. sp. (Figs. 4, 10-12) Male: Dorsal shield 562u(502u—562u) in length, 410u(380u-4104) in width; peripheral thickening of the dorsal shield not extending medially in region of the posterior glandularia (fig. 11); ventral shield, including posterior glandularia platelets 540u(494u-540u) in length, 464u(441u-464u) in width; anterior coxal groups separated by articular membrane from the third coxae; 4—-5(3-5) en- larged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; genital field 121u(1074-121u) in width; 3 pairs of genital acetabula; the 2 pairs of glandularia posterior to the ventral shield small and separated (fig. 10); excretory pore without associated sclerotization; eyes re- duced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P—I, 16u( 144-17); P-II, 55u( 484-55.) ; P-III, 27u(24u-27u); P-IV, 52u(48u-52u); P-V, 38u(32u-38u); structure of palp similar to that illustrated in fig. 14; dorsal lengths of the distal segments of the first leg: I-Leg-4, 9lu(80u-91u); I-Leg-5, 239u(2154-239u); I-Leg-6, 142u(128u-142u); greatest height of distal segments of first leg: I-Leg-4, 3lu(29u-31lu); I-Leg-5, 66u(56u-66u); I-Leg-6, 52u(48u-52u); dorsoproximal portion of I-Leg-6 extending somewhat proximally (fig. 4); claw at tip of first leg 100u in length; swimming hairs absent. Female: Dorsal shield 586u(532u-608.) in length, 456u(395u-456u) in width; dorsal shield similar to that of male; ventral shield, including posterior glandularia platelets 570u(525u-608u) in length, 502u(456u-524u) in width; anterior coxal groups separated by articular membrane from the third coxae; 5—7(4-7) en- larged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield (fig. 12); genital field 149u(142u-168u) in width; the individual acetabular plates 38u(36u-42u) in width; 3 pairs of genital acetabula; the posterior glandularia as described for the male; excretory pore without associated sclerotiza- tion; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P-I, 17u( 164-204); P-II, 62u(59u-62»); P-IlI, 35u( 29-35); P-IV, 55u(50u-55u); P-V, 42u(38u-43.); dorsal lengths of the distal segments of the first leg: I-Leg-4, 974(93u-100u); I-Leg-5, 2634 (231u-263u); I-Leg-6, 149u(142u-149u); greatest height of distal segments of the first leg: I-Leg-4, 35u(34u-35u); I-Leg-5, 72u(66u-72u); I-Leg-6, 594 (52u-59u); claw at tip of first leg 107u(93u-107u) in length; structure of palp and first leg similar to male; swimming hairs absent. Holotype: Adult male, found in a gravel bar in Satus Creek (on US 97 approximately 3 miles south of the Yakima County Line ), Klickitat County, Washington, August 18, 1961. Allotype: adult female, same 224 PROC. ENT. SOC, WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 data as holotype. Paratypes: 1 female, same data as holotype; 4 males, 12 females, from a gravel bar in Ten Mile Creek approximately 7 miles south of Yachats, Lane County, Oregon, August 13, 1961; 1 female, taken in a gravel bar in the Fisher River 30 miles southeast of Libby, Lincoln County, Montana, August 22, 1961. Discussion: There are three known species of Stygomomonia s. s. which possess two pairs of small glandularia platelets at the posterior end of the ventral shield. These include S. hachiojiensis described by Imamura (1959) from Japan, and the two North American species, S. separata and S. neomexicana. The present species differs in that I-Leg-6 is more than one-half the length of I-Leg-5. Stygomomonia (Stygomomonia) neomexicana, n. sp. (arbi Ti, 2toy, Bill) Female: Dorsal shield 562 in length, 4104 in width; peripheral thickening of the dorsal shield extending somewhat medially in direction of posterior glandularia (fig. 31); ventral shield, including posterior glandularia platelets 555u in length, 471 in width; anterior coxal groups separated by articular membrane from the third coxae; 5 enlarged setae present on each side of the fourth coxae posterior to the insertions of the fourth legs; acetabular plates and postgenital sclerite separated from the ventral shield; genital field 1454 in width; the individual acetabular plates 3lu in width; 3 pairs of genital acetabula; posterior glandularia platelets small, separated on one side, lightly fused on the other side (fig. 28); excretory pore without associated sclerotization; eyes reduced; integumental pigmentation absent. Dorsal lengths of the palpal segments: P—I, 214; P—II, 60u; P—III, 324; P-IV, 55u; P-V, 41y; structure of palp similar to that illustrated in fig. 14; dorsal lengths of the distal segment of the first leg: I-Leg-4, 100u; I-Leg-5, 2284; I-Leg-6, 98u; greatest height of the distal segments of the first leg: I-Leg-4, 38u; I-Leg-5, 59u; I-Leg-6, 45u; claw at tip of first leg 66u in length; figure 7 illustrates these segments; swimming hairs absent. Male: Unknown. Holotype: Adult female, taken in a gravel bar in Whitewater Creek approximately 5 miles northeast of Glenwood, Catron County, New Mexico, July 13, 1966. Discussion: The structure of I-Leg-6 with its dorsoproximal projec- tion extending distally (fig. 7) will separate the present species from both S. hachiojiensis and S. separata. REFERENCES Habeeb, H. 1957. A subterranean mite. Leaflets of Acadian Biol. 15:8. Hopkins, C. L. 1966. Two new species of water-mite (Acari, Hydrachnellae ) from New Zealand. Trans. Roy. Soc. New Zealand 8:111-117. Imamura, T. 1959. Water-mites (Hydrachnellae) of subterranean waters in Kanto District, Japan. Acarologia 1:426—451. Mitchell, R. D. 1959. A new water-mite of the genus Stygomomonia (Family Momoniidae). Trans. Amer. Microsc. Soc. 78:154—157. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 225 THE GENUS TYPOPSILOPA CRESSON IN THE WESTERN HEMISPHERE (DieTERA: EPHYDRIDAE ) Wiis W. Wirt, Systematic Entomology Laboratory, Entomology Research Division, Agr. Res. Serv., USDA‘ The genus Typopsilopa Cresson has only two recorded American species, atra (Loew), thought to range from Oregon to Ontario and south to California, Florida, and Central America, and _ flavitarsis Cresson, a Neotropical species ranging to Arizona and the southern states. Examination of the male genitalia of available specimens reveals that these species have been confused with each other, that two new North American species have been confused under atra, and that there are three new Neotropical species closely related to flavi- tarsis. The species are readily separated by examination of the male genitalia, but they are very difficult to recognize by using external characters. Cresson applied the name Psilopa nigra Williston to a species of Helaeomyia, but examination of Williston’s type shows that it is a Typopsilopa and identical with flavitarsis Cresson, which falls as a synonym. The types of the new species here described are in the U. S. National Museum (abbreviated USNM in this paper) in Washington, D.C., unless otherwise specified. I am greatly indebted to the fol- lowing persons and institutions for the opportunity to study material in their respective museums and collections: Paul H. Amaud, Jr., California Academy of Sciences, San Francisco, California (CAS); George W. Byers, University of Kansas, Lawrence (KANS); J. G. Chillcott, Canada Department of Agriculture, Ottawa, Canada (CAN); Saul Frommer, University of California, Riverside (Riverside); Harold J. Grant, Academy of Natural Sciences of Philadelphia, Pennsylvania (ANSP); R. O. Schuster, University of California, Davis (Davis); and Pedro W. Wygodzinsky, American Museum of Natural History (AMNH), New York, N. Y. Genus Typopsilopa Cresson Typopsilopa Cresson, 1916, Ent. News 27:147.—Cresson, 1947, Trans. Amer. Ent. Soc. 73:123 (syn.: Psilopina). Type-species, Typopsilopa flavitarsis Cresson, orig. des., = nigra ( Williston ). Psilopa, subg. Psilopina Becker, 1926, in Lindner, Flieg. Pal. Reg., fam. 56, p. 38. Type-species, Ephygrobia electa Becker, orig. des. Small black subshining flies with hyaline, unmarked wings, pale halter knobs, and legs except portions of the tarsi blackish; macrochaetae strong. Head moderately short and broad; breadth-height-length proportions as 3-6-8. 1Mail address: c/o U.S. National Museum, Washington, D.C. 20560 226 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Eyes bare. Frons short and sloping toward antennae, 0.41 as broad as head; mesofrons and orbits poorly differentiated; 1 pair of long reclinate orbitals, a proclinate seta arising slightly anterolaterad and about a third as long as orbital; inner and outer verticals long, both reclinate; a pair of long proclinate ocellars arising slightly behind level of anterior ocellus. Vertex convex but not sharply so; ocellar triangle equilateral; occiput with fine setae; cheeks setose and bearing 2-3 short bristles, the ventral one longer. Face nearly as broad as frons, slightly convex, with small foveae under the antennae; most prominent at level of the strong upper facial which is located at a third to half the distance from epistomal margin to antenna base; a second lower facial, a third to half as long as upper, and several fine setae in facial series. Epistomal margin slightly emarginate, clypeus moderately developed and exposed; palpi and proboscis normally de- veloped. Antenna with second segment bearing a long, stout, proclinate spine at anterodorsal corner; third segment short, 1.2-1.9 times as long as broad; arista with 8—12 long dorsal rays. Thorax moderately stout; mesonotum convex anteriorly, more flattened caudad; scutellum moderately flat; 2 pairs of notopleurals; 1 humeral; 1 presutural; 1 supra-alar; 1 intra-alar; 3 pairs of dorsocentrals, the anteromost located before the suture, the posterior pair (not a true dorsocentral) more closely approximated in front of scutellum; 2 pairs of marginal scutellars; all bristles long; surface of mesonotum and scutellum subshining but coarsely scoriaceous, also bearing numerous fine, somewhat appressed setae, those in acrostichal area forming 8 irregular rows. One strong and 1 weak mesopleural bristle and 1 strong sternopleural; pleura also with fine setae. Legs moderately stout, without special modifications; forefemur in both sexes with anteroventral comb of 15-20 short, flattened denticles; femora with a few enlarged bristle-like hairs; midtibia with- out erect extensor bristles but a long, ventral bristle at tip; tarsi pale at base, distal segments variably darkened. Wing with costa extending to fourth vein; second vein moderately long, costal index (length of second costal section divided by length of third) of American species 1.20-1.37; first posterior cell parallel-sided; fourth vein index (length of ultimate section divided by length of penultimate) 1.05-1.42. Abdomen moderately long and broad, slightly convex dorsally, subshining, provided dorsally with numerous short appressed setae and a few short bristles at posterior segment margins. Sixth tergum slightly exposed usually in female, not in male. The American species of Typopsilopa are readily separated into two groups on the basis of structure of the male genitalia. In the Atra Group, the ninth tergum is a narrow ribbon (though broader in arnaudi); the tenth sternum forms a narrow and ribbonlike genital arch mesad with the lateral ends expanded into setose lobes, which in arnaudi become elongated arms with a few heavy, black spines. The fifth sternum is a long setose plate with posterolateral corners expanded and lobelike, and the sixth sternum is a narrow bare plate with posterior end bifurcate. The anterior surstylus is a long slender process with slender tip bearing a few setae, arising with stouter base articulating between the anteroventral corner of the ninth tergum and the posterolateral process of the sixth sternum. On the midline PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 D2. between the surstyli, the anteroventral processes of the hypandrium are appressed as slender, fingerlike hyaline blades. Species of this group have the foretarsus black and the mid and hindtarsi with tarso- meres 1-2 pale or dark, and 3-5 dark. This group has a northern distribution and has relatives and counterparts in Asia and Africa, although not in Europe. The second group, which is exclusively Neotropical, is based on nigra (Williston), the type of the genus. In these species, the ninth tergum is broad, and in one species the cerci become modified. The greatest modifications in this group are found in the shape of the tenth stemum, which becomes elongated and massive, the antero- ventral lobes more approximated, and in manni, bearing dense, heavy spines. The lobes of the tenth sternum, along with the apices of the anterior surstyli and the ventral processes of the hypandrium, lie at rest in a heavily sclerotized box formed by the greatly broadened and dorsally arched sixth sternum, for which the fifth sternum forms a ventral setose margin. Species of the Nigra Group have tarsomeres 1-4 pale on all legs with only the fifth dark. Typopsilopa occupies an anomalous position, sharing characters which otherwise distinguish the subfamilies Psilopinae and Noti- philinae. For many years, it was thought to be related to Psilopa Fallén because of the close superficial resemblance in body form and color and most features of chaetotaxy. Cresson (1916), when he erected the genus Typopsilopa, compared it only with Psilopa and Clasiopella Hendel, pointing out that the generic importance of the presence or absence of the two true dorsocentrals may be questioned. It was not until 1946 that the true relationships of the genus were recognized when Cresson placed it in a new tribe of Notiphilinae, the Typopsilopini, giving the following tribal characters: “Eyes bare; lunule not developed; ocellars approximate, and strong, aligned with or caudad of line of anterior ocellus; antennal spine strong; dorsocentrals arranged 1: 1 or 0: 1; posterior notopleural not removed dorsad; mid _ tibiae without erect extensors; costa extending to vein IV.” Key TO THE AMERICAN SPECIES OF Typopsilopa (See also Table 1 ) 1. Tarsi dark brown, at least on forelegs; face flatter in middle; sternum VI of male (figs. 1, 8) narrow and ribbonlike, usually not much more than half as wide as distance across lobes of sternum X, not markedly concave Tarsi pale at least in part on all legs; face more tumid in middle; sternum VI of male (fig. 13) broader than distance across lobes of sternum X, heavily sclerotized and forming a broad concavity into which the genital mrocesses.. Gan be thrust) anterouorsally {0 2 2 4 2. Lobes of sternum X (figs. 10, 12) greatly elongated arms nearly as long as the transverse basal bridge of sternum, and each bearing distally 4 long, stout DlackoSpines:all tarsi black . > 2 arnaudi, n. sp. 2 228 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Lobes of sternum X (figs. 2, 5) low and flaplike and bearing distally only fine hairs or a few bristles; at least mid and hindtarsi slightly yellowish _ 3 3. Sternum VI (fig. 1) of male with the bifurcate posterior portion broader than the anterior end; lobes of sternum X (fig. 2) rounded with numerous fine hairs; surstyli (fig. 3) slender and rodlike distally; third antennal segment long, 1.9 times as long as greatest breadth ____ antennalis, n. sp. Sternum VI (fig. 9) of male with the bifurcate posterior portion narrower than the anterior end; lobes of sternum X angulate (fig. 5) with a few stouter bristles; surstyli (figs. 6, 11) stouter and more fingerlike distally; third antennal segment short, 1.5 times as long as basal breadth rpetatrmme seeree dee rete hie. dese SY oe ee atra (Loew) 4. Male cerci (fig. 18) produced on Fianterovential ends in form of a pointed, blackish, strongly sclerotized spine; sternum X with ventromesal margin between the lateral lobes slightly concave; foreleg with only tarsomere 1 ULI I oee etn lt I CD a th 8 oh tee ee __. archboldi, n. sp. Male cerci (fig. 14) normal, anteroventral ends not spinelike; sternum X with deeply emarginate anteroventral margin; foretarsi variously darkened Mg de es 5 5. Sternum X (fig. 13) eh lateral lobes widely separated, low and angular, not as long as the distance between their distal points; foreleg with tarsomeres 1—4" buicht yellow == eee nigra (Williston) Sternum X (figs. 15, 17) with lateral lobes much longer and broader than the mesal emargination between them; foreleg with only tarsomere 1 yellowish 22)! F ek et ee 6 6. Lobes of sternum al Gig: 15) with fin’ bristles only ye wae inea, nN. sp. Lobes of sternum X (fig. 17) covered distally with curved stout black SDUCS pas ret oe aoe a, pcs dbeusd-p Re oe Se ee manni, 0. sp. Typopsilopa atra (Loew) (Figs. 5-9, 11) Psilopa atra Loew, 1862, Monogr. N. Amer. Dipt. 1:143 (male; Middle States). Typopsilopa atra (Loew); Cresson, 1916, Ent. News 27:147 (comb. ).—Cresson, 1946, Trans. Amer. Ent. Soc. 72:240 (dist.: Mass. to Oreg., s. to Fla., Utah and Calif.).—Wirth, 1965, in Stone, et al., Cat. N. Amer. Dipt. p. 743 (syn.: scoriacea Loew ). Psilopa scoriacea Loew, 1862, Monogr. N. Amer. Dipt. 1:142 (female; N.Y.). Male, Female——Wing 2.25 mm. long by 0.88 mm. broad. Shining black; mesonotum and scutellum scoriaceous, with sparse coarse brown pollen; face with slight whitish dusting, especially on sides; antenna and palpus entirely black; tarsus brownish black on foreleg, tarsomeres 1 and 2 yellowish on mid and hind- legs; wings grayish hyaline, the veins yellowish; halter knob creamy white, stem dark. Antenna with third segment short, 1.5 times as long as greatest breadth; arista with 9 rays. Strong upper facial located about a third way from epistomal margin to antenna base; face nearly flat, only slightly convex. Wing with costal index 1.39; fourth vein index; hind crossvein 1.06 times as long as breadth of first posterior cell at same level. Male genitalia as in fig. 11. Sternum V (fig. 8) nearly twice as long as posterior breadth, with uniform sparse setose hairs on surface. Sternum VI PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 229 Table 1. Quantitative Characters of Typopsilopa species. Tarso- Tarso- meres Antenna meres Legs II, Wing Fourth Arista Ill Leg I Ill Length Costal Vein Rays (L/W) brown brown (mm. ) Index Index Atra Group atra (Loew ) 9 1.5 1-5 3-5 2.25 1.30 1.39 antennalis n. sp. 8 1.9 1-5 3-5 212 1.20 1.42 arnaudi n. sp. 9 iS 1-5 1-5 21S V2 127 Nigra Group nigra (Will. ) 10 i 5 5 2.86 1 e37/ 1.05 archboldi n. sp. 8 lea 9-5 5 2.56 1.26 1.08 manni nN. sp. 10 avs 2-5 5 2.56 11 349) 1.28 inca sp. 12 IES 2-5 5 2.62 1.25 lly (fig 9) a small, elongated, narrow sclerite without hairs or setae; anterior end broader and rounded, the ventral surface slightly concave, spoonlike, and tucked dorsally under the posterior end of sternum V; posterior end bifurcate with a pair of small lobes. Sternum X (fig. 5) with very slender mesal bridge, the lateral ends expanded in a pair of angulate lobes bearing a few stout, long, bristly hairs. Surstyli and (figs. 6, 11) processes of hypandrium (figs. 7, 11) each produced in a moderately slender fingerlike process, the former with 8-10 very fine distal setae; slenderer and straight in western specimens, and slightly curved in eastern in- dividuals. Distribution —North America from California to Ontario and New York, south to Georgia and Central America. Specimens Examined.—182 specimens, as follows: ARIZONA: Cat- alina Mts. (Wehrle); Wickenberg, Hassayampa R. (Wirth). AR- KANSAS: Washington Co. (Rouse). CALIFORNIA: Capistrano Hot Spr. (Melander ); Centerville, Fresno Co. (Yost); Lemore, Kings Co. (Hall); Live Oak Park (Melander); Panamint Springs, Inyo Co. (Howden); Riverside (Melander); San Juan Hot Spr. (Melander); Temecula (Melander); Sentenac Canyon, San Diego Co. (Bechtel); Victorville (Richards); Whitewater Fish Hatchery, Riverside Co. (Arnaud). COSTA RICA: La Caja, 8 km. w. San Jose (Schmidt). DISTRICT OF COLUMBIA: Chain Bridge (McAtee). GEORGIA: Billy’s Id., Okefenokee Swamp; Tifton (Bradley); Valdosta (Young). GUATEMALA: Ingenio Railroad Sta. (Aldrich). ILLINOIS: Al- gonquin; Champaigne Co. (Shackleford); Chicago (Melander); Dubois (Malloch); Glen Ellyn (Melander); Meredosia (Malloch); Muncie (Malloch); Pistakee Bay (Melander); River Forest (Melander). INDIANA: Dune Park (Melander); Lafayette (Aldrich); Michigan City (Aldrich). IOWA: Ames. KANSAS: Lawrence (Sanderson ); Manhattan (Sabrosky). KENTUCKY: Lexington (Aldrich). MARY- LAND: Chesapeake Beach (Aldrich, Malloch). MASSACHUSETTS: Woods Hole (Melander). MEXICO: El Salto, Durango, 9000 ft. (McAlpine); Mt. Colima, pine zone, se. slope (Leech, Ross); Sierra 230 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Figs. 1-4, Typopsilopa antennalis, n. sp., 6 genitalia; 1, sternum VI, ventral view; 2, sternum X, ventral view; 3, surstylus, lateral view; 4, process of hypan- drium, lateral view. Figs. 5-9, 11, T. atra (Loew), & genitalia: 5, sternum X, ventral view; 6 and 7, apices of surstylus and process of hypandrium, respectively, New York specimen, lateral view; 8, sternum V, ventral view; 9, sternum VI, PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 yl | Laguna, Big Canyon, Baja Calif. (Ross & Bohart); Tijuana, 60 km. s. (Melander). MICHIGAN: Detroit (Steyskal); Midland Co. (Dreis- bach); Nottawa (Sabrosky). MISSOURI: Atherton (Adams); Co- lumbia (Crosby). NEW JERSEY: Trenton. NEW YORK: Allegany St. Park (Wirth); Bergen (Beamer); Niagara (Melander); Sinclair- ville (Wirth). NORTH CAROLINA: Andrews Bald, Gr. Smokies (Melander); Highlands (Richards). OHIO: Wauseon. ONTARIO: Grand Bend (Shewell); Pt. Ryerse (Shewell). TEXAS: Austin (Melander); Brownsville (Hardy); Hunt, Guadalupe R. (Wirth); Kerrville, Henke Pond (Wirth); Dallas (Jones); Brownwood (Painter); Cibola R., Sutherland Spgs. (Nabours & Sabrosky ); Liberty (Tucker); San Antonio (Melander); Uvalde, Gamer State Park (Wirth). UTAH: St. George (Knowlton & Smith); Spanish Fort (Corbett). VIRGINIA: Alexandria (Aldrich); Fairy Stone St. Park (Vockeroth). WISCON- SIN: Devils Lake (Melander). Typopsilopa antennalis Wirth, n. sp. (Figs. 1-4) Male.—Indistinguishable externally from atra (Loew) except for length of third antennal segment which is 1.86 times as long as greatest breadth. Costal index 1.20; fourth vein index 1.42; hind crossvein 1.72 times as long as breadth of first posterior cell at same level. Wing 2.12 mm. long by 0.85 mm. broad. Genitalia as in figs. 1-4. Sternum X (fig. 2) with the lateral lobes bearing fine setose hairs only, no strong bristles, the lobes placed closer together with a broader mesal bridge than in atra; sternum VI (fig. 1) with posterior bifurcate portion at least as broad as the anterior half; surstylus (fig. 3) with distal end very slender, bearing 2—3 fine hairs as extreme tip; process of hypandrium (fig. 4) a slender fingerlike blade. Female.—Not distinguished with certainty from atra in the series recorded. Distribution—F lorida, Georgia, South Carolina. Types.—Holotype male, Ridgeland, South Carolina, 24 March 1954, A. Stone (USNM 69525). Paratypes, 8 males: FLORIDA: Royal Palm St. Park, 27 January 1939, A. L. Melander, 1 male. GEORGIA: Tifton, § June 1896, 3 males. SOUTH CAROLINA: Manning, 29-30 May 1914, W. Stone, 4 males (ANSP, USNM). Typopsilopa arnaudi Wirth, n. sp. (Figs. 10, 12) Male, female.—as in atra (Loew) except as follows: Face slightly more carinate and not quite so dull; tarsomeres 1-5 saturate black on all legs; costal index 1.32; fourth vein index 1.27; wing 2.13 mm. long by 0.80 mm. broad. < ventral view; 11, genitalia of Arizona specimen, lateral view. Figs. 10, 12, T. arnaudi, n. sp.: 10, é genitalia, lateral view; 12, sternum X, ventral view. Fig. 13, T. nigra, n. sp., 6 genitalia, ventral view. Abbreviations: as, anterior sur- stylus; c, cercus; hyp, hypandrium; st, sternum; t, tergum. bo 32 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Figs. 14, 15, Typopsilopa inca, n. sp.; 14, 6 genitalia, lateral view; 15, sternum X, ventral view. Figs. 16, 17, T. manni, n. sp., 6 genitalia: 16, surstylus, lateral view; 17, sternum X, ventral view. Fig. 18, T. archboldi, n. sp., & genitalia, lateral view. Abbreviations: as, anterior surstylus; c, cercus; hyp, hypandrium; st, sternum; t, tergum. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 233 Male genitalia as in fig. 10. Sternum X (fig. 12) with very slender mesal bridge, the lateral lobes each greatly produced ventrocephalad in a long arm bearing 4 long, black, stout, blunt spines. Sterna V and VI similar to those of atra, VI with slender bifurcate posterior end; surstyli slender and curved gradually to fingerlike tip bearing a few minute subapical setae; processes of hypandrium long slender blades. Distribution.—California and Oregon to Idaho, Utah, New Mexico, and Mexico. Types.—Holotype male, allotype female, Deep Creek, California, 25 October 1953, A. L. Melander (USNM 69526). Paratypes, 64 males, 91 females: ARIZONA: Granite Delta, 30 May 1935, P. W. Oman, 1 male; Indian Pine, 28 August 1964, M. E. Irwin, 1 male; Montezuma Wells Nat. Mon., 30 June 1953, W. W. Wirth, 3 females; Patagonia, 24 June 1933, P. W. Oman, 1 female; same, 27 June 1953, W. W. Wirth, 1 male; Portal, S. W. Res. Sta., 26 September 1964, 1 June 1965, V. D. Roth, 2 males (Riverside); Sunnyside Canyon, Huachuca Mts., 11 July 1940, D. G. Hall, 1 female (ANSP); same, 9 July 1940, D. E. Hardy, 1 male (KANS). CALIFORNIA: Aguanga, 4 mi. w. River- side, 24 April 1951, E. I. Schlinger, 1 female (Davis); Alamos Creek, San Luis Obispo Co., 22 June 1948, W. W. Wirth, 1 male, 1 female; Alpine Lake, 14 August 1957, A. L. Melander, 1 female; Barton Flats, August 1942, September 1950, 23 August 1952, A. L. Melander, 2 males, 3 females; Big Meadows, 30 July 1937, G. Spurlock, 2 females; Centerville, Fresno Co., 7 June 1965, P. Yost, 1 female (Riverside); Cisco, 5 June 1940; M. & H. James, 1 male, 1 female; Colfax, 20 May 1952, E. I. Schlinger (Davis); Crestline, 4 July 1942, A. L. Melander, 1 female; Deep Spring, Inyo Co., 16 July 1953, E. IL. Schlinger, 3 males, 1 female (Davis); Dutch Flat, Placer Co., 20 May 1952, E. I. Schlinger, 1 female (Davis); Fallen Leaf, Eldorado Co., 6500 ft., 13 July 1961, J. G. Chillcott, 1 female (CAN); Glen Alpine, Lake Tahoe, 1 September 1930, H. H. Keifer, 1 female; Green Valley, 26 July 1944, A. L. Melander, | male, 4 females; Idyllwild, 7 June 1942, A. L. Melander, 1 male; Jenks Lake, 14 July, 24 August 1950, A. L. Melander, 3 females; Jacumba Springs, 28 June 1917, J. M. Aldrich, 4 females; Julian, 5 May 1945, A. L. Melander, 1 female; Kaweah River, Tulare Co., 12 October 1947, R. Coleman, | male; Keen Camp, 7 June 1942, A. L. Melander, 1 male, 3 females; Lake Tahoe, 11 August 1940, L. J. Lipovsky & D. E. Hardy, 2 males; Onyx, 7 mi. e., Kern Co., 12 June 1961, H. F. Howden, 1 female (CAN); Ortega Hwy., Mariana River, 19 October 1944, A. L. Melander, 1 female; Pacific, 9 August 1940, D. E. Hardy, 1 male, 1 female; Palm Springs, 19 November 1943, 4 October 1946, A. L. Melander, 2 males, 2 females; Pinecrest, Tuolumne Co., 13 August 1948, P. H. Amaud, 1 male, 1 female; Riverton, 19 August 1953, E. I. Schlinger, 1 female (Davis); Siskiyou Co., 19 July 1948, W. W. Wirth, 1 male (CAS); Sardine 234 PROC, ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Creek, Mono Co., 6 July 1951, A. T. McClay, 2 females (Davis); Strawberry, Tuolumne Co., 20 July 1951, A. T. McClay, 1 male, 2 females; (Davis), Thorn, Mohave Desert, 30 May 1944; A. L. Melander, 1 male; Trinity River Camp, Trinity Co., 17 July 1953, A. T. McClay, 3 males (Davis); Tuolumne Meadows, | July 1940, D. E. Hardy, 1 male; Valyermo, 13 May 1934, A. L. Melander, 1 male, 4 females; Victorville, 12 May 1955, W. R. Richards & W. R. Mason, 2 males, 9 females (CAN); Whitewater Fish Hatchery, Riverside Co., 6 January 1953, P. H. Armaud, 1 male, 2 females; Willets, 19 June 1935, A. L. Melander, 1 male; Yosemite Nat. Park, 1 August 1940, L. J. Lipovsky, 1 male, 27 June 1947, A. L. Melander, 1 female; Yuba Pass, Sierra Co., 20 August 1953, E. I. Schlinger, 1 male, 1 female (Davis). IDAHO: Caldwell, 2372 ft., 9 July 1926, C. Wakeland, 1 male (ANSP). MEXICO: Baja California, Sierra San Pedro Martir, La Grulla, 6900 ft., 12-15 June 1953, P. H. Arnaud, 14 males, 12 females; Rancho Viejo, 7000 ft., 13 June 1953, P. H. Arnaud, 3 females (CAS); Trail La Joya to La Zanja, 10 June 1953, P. H. Arnaud, 1 male, 1 female (CAS); E] Salto, 9000 ft., 10 mi. w. Durango, 10 June 1964, J. F. McAlpine, 1 male, 3 females (CAN); same, 7 May 1961, Howden and Martin, 1 female (CAN). NEVADA: Beatty, 26 May 1940, G. E. Bohart, 1 female (CAS); Carson City, 25 May 1952, E. I. Schlinger, 1 female (Davis); Crystal Springs, Lincoln Co., 21 June 1953, A. B. Gurney, 1 male, 2 females; Reno, 20 October 1915, H. G. Dyar, 1 male. NEW MEXICO: Belen, 1 July 1947, L. D. Beamer, 1 male; Jemez Springs, June, September 1914, 2 males, 4 females (ANSP); Socorro, 1916 Williston Coll., 1 female; Rio Grande River, Taos Co., 6 July 1953, W. W. Wirth, 1 male. OREGON: Kerby, 18 September 1934, A. L. Melander, 2 males, 3 females (ANSP, USNM). UTAH: Kanab, 19 April 1935, A. L. Melander, 1 male. This species is named in honor of Dr. Paul H. Arnaud, Jr., of the California Academy of Sciences in recognition of his extensive contri- butions to the dipterology of western North America. It is readily distinguished from the other North American species by the saturate black tarsi and the heavy black spines on the male tenth sternum. Typopsilopa nigra (Williston), n. comb. (Fig. 13) Psilopa nigra Williston, 1896, Trans. Ent. Soc. London 1896:139, pl. 13, fig. 139 (female; St. Vincent I.; fig. head). Typopsilopa flavitarsis Cresson, 1916, Ent. News 27:147 (Ariz. ).—Cresson, 1946, Trans. Amer. Ent. Soc. 72:240 (dist.: Neotropical, Fla., Ga., La., and Ariz. ).— Wirth, 1956, Amer. Mus. Novitates no. 1817, p. 17 (Bahamas). New synonymy. Male, female.—Shining black; mesonotum and scutellum scoriaceous; tarsi bright yellow except tarsomere 5 brownish; wing grayish hyaline; halter knob white; frons and face less shining than in atra; opaque frontalia less differentiated; face PROC. ENT. SOC. WASH., VOL. 70, No. 3, SEPTEMBER, 1968 235 more or less irregularly wrinkled. Face longer than in atra, more tumid in middle, the long upper facials higher, about midway between epistomal margin and antenna. Antenna short, third segment only 1.2 times as long as greatest breadth; arista with 10 rays. Legs stouter than in atra, femora particularly stout; hairs on abdominal terga rather strong and dense. Second vein longer than in atra, costal index 1.37; fourth vein index 1.05. Wing 2.86 mm. long by 1.10 mm. broad. Male genitalia as in fig. 13. Stermnum VI heavily sclerotized and broad with distinct longitudinal wrinkles forming a boxlike ventral concavity within which the tenth sternum, surstyli, and processes of the hypandrium lie in repose. Sternum X large, slightly more than twice as broad as greatest length, the anteroventral margin with a broad rounded excavation mesad, the anterior margin on each side forming a broad, angulate lobe provided with fine, somewhat twisted hairs. Cerci normal, flaplike, setose. Surstyli gradually tapered to slender, bladelike processes, each with a row of minute setulae on ventral side; anteroventral processes of hypandrium slender blades lying between the surstyli on midline. Distribution.—Arizona to Georgia, south to Brazil. Types.—Psilopa nigra: 2 syntypes, St. Vincent I. (one is in the British Museum (Nat. Hist.), London and is here designated lecto- type). Typopsilopa flavitarsis: holotype male, Bill Williams Fork, Arizona, F. H. Snow (KANS); also 3 male paratypes, same data. Specimens Examined.—74 specimens, as follows: ARIZONA: Bill Williams Fork (Snow), holotype and 1 male paratype; Standfield (Butler). BAHAMAS: Andros I., Mangrove Cay (Hayden); Eleuthera I., New Portsmouth (Hayden and Giovannoli); Long I., Deadman’s Cay (Hayden); San Salvador I., near Cockburn Town (Hayden). BRAZIL: Amazon River near Obidos (Holt, Blake, and Agostini); Gavea, Rio de Janeiro (Sousa Lopez). BRITISH GUIANA: George- town Bot. Gard. Exp. Sta. (Squire), bred from rice stool. CALI- FORNIA: Needles (Kusche). COSTA RICA: Peralta (Calvert). DOMINICA: Cabrit Swamp (Wirth). DOMINICAN REPUBLIC: Samana (Sanchez). FLORIDA: Alachua Co. (Weems); Ft. Lauder- dale (Melander); Hendry Co. (Morse); Little River (Knab); Miami (Knab). GEORGIA: Tifton. LOUISIANA: New Orleans (Plank); Slidell. MEXICO: La Bolsa (McGovran). NICARAGUA: Potosi (Woke). PANAMA: Ancon, C. Z. (Greene); Taboga I. (Busck). PUERTO RICO: Mayaguez. VIRGINIA: Warsaw, Richmond Co. (Wirth). Discussion—I examined the female syntype of Psilopa nigra Willis- ton in the British Museum (Nat. Hist.) in 1957 and found that it was a Typopsilopa; at that time, a critical comparison with flavitarsis was not made. Characters given by Williston in the original descrip- tion of nigra which agree well with flavitarsis: shape of face and antenna; position of the facials midway between epistomal margin and antenna; arista with 10-12 rays; legs black, all the tarsi yellow, with the distal joints blackish; second section of the costal vein about one-third longer than the third section; ultimate section of the 236 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 fourth vein only a little longer than the penultimate section; length 3 mm. These characters fit flavitarsis much better than archboldi, n. sp., the only other Typopsilopa species whose known distribution would bring it into consideration as Williston’s species. Typopsilopa archboldi Wirth, n. sp. (Fig. 18) Male, female.—Similar to nigra (Williston) but differing as follows: Fore- tarsus with only tarsomere 1 yellowish, 2-5 dark brown; face more tumid in middle; third antennal segment longer, 1.7 times as long as greatest breadth; arista with 8 rays; costal index 1.26; fourth vein index 1.08. Wing 2.56 mm. long, 0.96 mm. broad. Male genitalia as in fig. 18. Sternum V short and bristly; sternum VI a massive, well sclerotized box-like concavity without longitudinal wrinkling; sternum X with narrow mesal bridge, anterior margin with low rounded bristly lateral lobes, the mesal excavation very gently rounded. Cerci large, each greatly prolonged ventrally in a heavily sclerotized, sharp spinelike point extending over mesal excavation of sternum X. Surstyli greatly bent near base in lateral view, distal por- tion slender and nearly straight, bearing 2 fine, distal setae and a few subapical hyaline spicules. Anteroventral process of hypandrium very stout, not divided, forming a subconical ventral protuberance. Distribution.—Dominica, Puerto Rico. Types.—Holotype male, allotype female, Cabrit Swamp, Dominica, 22-25 March 1965, W. W. Wirth, Bredin-Archbold-Smithsonian Bio- logical Survey of Dominica (USNM 69527). Paratypes, 9 males, 9 females: DOMINICA: 7 males, 7 females, same data as types. PUERTO RICO: San German, 23 Dec. 1962, P. & P. Spangler, 2 males, 2 females. This species is dedicated to Mr. John Archbold, in appreciation of his support of the Biological Survey of Dominica and his keen interest in the scientific exploration of the island. Typopsilopa manni Wirth, n. sp. (Figs. 16, 17) Male, female.—Similar to nigra (Williston) but differing as follows: third antennal segment longer, 1.7 times as long as greatest breadth; foretarsi darker, tarsomeres 2-5 brownish; costal index 1.22; fourth vein index 1.28. Wing 2.56 mm. long, 0.95 mm. broad. Male genitalia as in figs. 16, 17. Sternum V short, broad posteriorly, surface with short setose hairs; sternum VI heavily sclerotized, concaved ventrally forming a ventral pouch for the genital processes, narrowed posteriorly. Stermum X (fig. 17) greatly elongated, the bulbous lateral lobes as long as basal breadth of sternum, with a narrow, deep mesal cleft between the lobes nearly halfway to base of sternum; apices of lobes densely covered with short stout sharp spines on posteroventral side; anterodorsally each lobe drawn out in a finely setose flap ending in a distal point provided with 2 stout spines, this point not nearly as long as the rounded lobes. Cerci normal; flaplike. Surstyli (fig. 16) abruptly PROC. ENT. SOC. WASH., VOL. 70, No. 3, SEPTEMBER, 1968 237 bent near base, rather stout proximally, gradually narrowed to slender tip with small lateral point; anteroventral processes of hypandrium an appressed pair of slender pointed blades. Distribution Bolivia; Argentina, Paraguay. Types——Holotype male, Rosario, Bolivia, Rio Rocagua, No. 1921, W. M. Mann, Mulford Biol. Exped. (USNM 69528). Allotype female, Argentina, Formosa, Mission Laishi, 13-15 Dec. 1948. Paratypes, 3 males: PARAGUAY: Incarnacion, 16 Jan. 1927, F. & M. Edwards, 1 male. San Bernardino, 1908, Fiebrig, 1 male (ANSP); same, Babarczy coll., 1 male. This species is dedicated to Dr. William M. Mann in tribute to his early interest in entomology and his worldwide eminence as a zoologist. Typopsilopa inea Wirth, n. sp. (Figs. 14, 15) Male, female.—Similar to nigra (Williston) but differing as follows: Third antennal segment long, 1.9 times as long as greatest breadth; arista with 12 rays; foretarsi darker, tarsomeres 2—5 brownish; costal index 1.25; fourth vein index 1.17. Wing 2.62 mm. long, 1.06 mm. broad. Male genitalia as in fig. 14. Sternum V slightly longer than posterior breadth, surface with short setose hairs; sternum VI heavily sclerotized, concaved ventrally forming a ventral pouch for the genital processes. Sternum X (fig. 15) long and broad, quadrate in outline with a narrow anteromedian cleft extending to about a third of total length, the lateral lobes with broad truncate apical margins and bearing long bristly hairs on posteroventral and distal surfaces. Cerci normal. Surstyli slightly curved, tapering to slender tip bearing subapically 2-3 minute setae; anteroventral processes of hypandrium an appressed pair of hyaline blades slightly expanded distally in lateral profile. Distribution —Ecuador, Peru. Types.—Holotype male, allotype female, Canete Peru, 17 May 1941, P. A. Berry (USNM 69529). Paratypes, 7 males, 7 females: ECUA- DOR: Loya, Catamayo, 1500 meters, 24 March 1965, L. Pena (CAN). PERU: 6 males, 6 females, same data as types; Loreto, Yarinacocha, 9-18 April 1963, L. Pena, 1 female (CAN). This species is near manni n. sp. but differs markedly in the bristly instead of spinose armature of the more truncate lobes of the tenth sternum. The tenth sternum is more like that of nigra (Williston), but the lateral lobes are more truncate and the surstyli are slenderer, whereas nigra also differs in its pale tarsomeres 1-4 on the foreleg. REFERENCES Cresson, E. T., Jr. 1916. Descriptions of new genera and species of the dipterous family Ephydridae.—III. Ent. News 27:147-152. 1946. Synopses of North American Ephydridae (Diptera) II. The tribe Notiphilini of the subfamily Notiphilinae. Trans. Amer. Ent. Soc. 72:227-240. 238 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 STREPSIPTERA FROM THE SMITHSONIAN INSTITUTION’S SURVEY OF DOMINICA, WITH DESCRIPTION OF A NEW SPECIES OF ELENCHIDAE’ Jean WALKER Fox, Section of Insects and Spiders, Carnegie Museum, Pittsburgh, Pennsylvania 15213 Dr. Paul J. Spangler, Curator, Division of Coleoptera, United States National Museum (USNM) submitted for identification seven male Strepsiptera from the Smithsonian Institution’s biological survey of the Island of Dominica. Six were Elenchidae belonging to a new species of Pseudelenchus Ogloblin (1925) and the seventh specimen provides a new record of Corioxenos antestiae Blair (1936). The material preserved in fluid was first examined under magnifi- cation ranging from 10x to 80x since some of the structures can be more profitably observed before permanent mounting. This same range of magnification was employed for examination of the two dry mounts, one of which was in very poor condition, lacking both head and abdomen. Slides were studied with a Wild M20 compound microscope having 10x oculars and 3x, 10x and 20x objectives, and its camera lucida attachment was used for accurate drawings. I wish to acknowledge the professional help and encouragement in this study which has been given by my husband, Dr. Richard M. Fox. Both Mr. R. D. Pope of the British Museum (Natural History ) and Dr. Paul Spangler at USNM have assisted me by checking some specific problems. Richard L. Satterwhite, our staff artist, as usual pertected my drawings for publication and Joseph Y. Quil finished the manuscript copy. Family Elenchidae Pierce In a recent paper (1967) I mentioned the disagreement among authors as to the exact diagnostic characters of the Elenchidae, the major character under discussion being the number of antennal seg- ments. Five-segmented antennae were specified as diagnostic by Pierce (1907) who originally established the family with type-genus Elenchus Curtis, of which the type-species is E. walkeri (Curtis) (1831). I have not seen the holotype of E. walkeri which is in the Curtis Collection in Melbourne, Australia, but Mr. R. D. Pope of the British Museum (Natural History) kindly examined a paratype in the Dale Collection and reported (personal communication) that the antennae are clearly 5-segmented. Elenchus solomonensis Fox and Elenchinus eastopi Fox, which I described from the British Museum (Natural History) collection, both have five antennal segments. This ' Research supported by National Science Foundation Grant GB 5682. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 239 is also true of at least 11 of the 16 species previously described under various genera of Elenchidae. With 5-segmented antenna: Elenchus tenuicornis (Kirby) (1815). Elenchus walkeri (Curtis) (1829). Type-species of Elenchus by monotypy. Elenchus templetoni Westwood (1835). Deinelenchus australensis Perkins (1905). Type-species of Deinelenchus by monotypy. Liburnelenchus (= Mecynocera) koebeli Pierce (1909). Type-species of Li- burnelenchus by monotypy. Elenchus melanias Perkins 1910. Elenchus melanias silvestris Perkins 1910. Liburnelenchus heidmanni Pierce (1918). Elenchinus heidmanni Pierce (1918). Type-species of Elenchinus by monotypy. Elenchinus japonica Esaki and Hashimoto (1931). Sogatelenchus mexicanus Pierce (1961). Type-species of Sogatelenchus by monotypy. With 4-segmented antenna: Pseudelenchus carpathicus Ogloblin (1925). Type-species of Pseudelenchus by monotypy. Elenchinus delphacophilus Ahlberg (fide Lindberg, 1939). Elenchinus chlorionae Lindberg (1939). Elenchus falcipennis Luna de Cavalho (1956). Not known; publication unavailable: Elenchus maorianus Gourlay 1953 (fide Kifune, 1964). Under these circumstances it does not seem systematically sound to define the family, as did Bohart (1941), as having 4-segmented antennae, but rather to redefine it as possessing either 4 or 5, with species in at least two genera. Other diagnostic characters of the species described show too much similarity to justify establishment of another family. Since the species described below has 4-segmented antennae, it should be assigned to Pseudelenchus Ogloblin (1925), type-species P. carpathicus Ogloblin, 1925, which is the only genus in the family based on a species with 4-segmented antennae. Pseudelenchus spangleri, n. sp. (Figs. 1-6) Measurements are stated for the holotype and the averages for specimens seen are given in parentheses. Male: Body, light brown, 0.95 mm. long (0.84 mm.); hindwing, 1.0 mm. broad (0.95 mm.) with expanse of 2.2 mm. (2.0 mm.); width of head between eyes, 0.15 mm. Head: Squared vertex. 4-segmented antenna set laterally well in front of eyes; segment I prominent, about same size as II; III is branched with tip of 240 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 flabellum reaching slightly beyond middle of IV. Eye with 4 large ocelli visible dorsally on inner rim, probably 12-16 total. Mouthparts with very stout mandible somewhat rounded apically, a third longer than distal segment of maxillary palpus; 2-segmented palpus with proximal segment two-thirds length of distal one. Metathorax: prescutum with sharply triangular posterior and rounded anterior margin protruding beyond that of the scutum, well separated from scutellum. Scuti with defined sutures separating them from membranous portion above the broad, almost rectangular scutellum. Large, bandlike postlumbium. Postscutel- lum about three-fourths the length of anterior metathoracic sclerites. Forewing: Small paddle-shaped structure. Hindwing: 5 main veins. C and Sc coalesced for part of their length and appearing to be a single strong vein; R very short with single detached vein below; M: and Me about equal length; single A. Legs: All with 2-segmented tarsus. Midleg with slender femur longer than tibia (7:5). Hindleg with femur much stouter than, and equal in length to, tibia. First tarsal subsegment in all legs broader and twice the length of ter- minal one. Aedeagus: Quite thick for over half its length then tapering evenly to pointed apical tip; diagonal rows of tiny bristles on proximal third. Abdominal segment X: Short with tapered posterior margin. Female, larva and host unknown. Holotype male: Clarke Hall, Dominica, Malaise trap, 15 October 1964, Spangler; USNM 70069. 4 paratype males divided with two slides at CM, 2 slides and 1 dry mount at USNM. A damaged dry mount, probably this species, USNM, not included in type-series. Discussion: There are several characteristics which separate this species from P. carpathicus Ogloblin. P. spangleri is much smaller (0.8 mm.:1.5 mm.); the metathorax is longer than the abdomen; the postlumbium is not so broadly curved; the head is narrower with eyes closely set, not slightly pedunculate. Ogloblin states that there are only four main veins in the hindwing while P. spangleri shows five. However, his figure indicates that this apparent discrepancy is a matter of terminology since he seemed to consider C and Sc as a single vein. This species is named in honor of the collector, Paul J. Spangler. Corioxenos antestiae Blair (1936) Single male specimen, Cabrit Swamp, Dominica, 18 July 1964, O. S. Flint, USNM. The specimen was examined in fluid before mounting. Due to an accident, the material on the slide was disoriented and contaminated with dust, but efforts to repair it might have resulted in further damage to this sole specimen and were avoided. Basic structures can be observed, and although the aedeagus is broken, its tip shows under the edge of the left hindwing. There are no bristles along the aedeagus of this specimen, nor were PROC. ENT. SOC. WASH., VOL. 70, No. 3, SEPTEMBER, 1968 241 0.1mm i 6 TD, (ae, 0.5mm Fy po Figs. 1-6. Pseudelenchus spangleri, n. sp.: 1, body showing antenna and wing in dorsal aspect; 2, detail of antennal segments; 3, hindleg; 4, mandible; 5, maxillary palpus; 6, aedeagus. (Figs. 2-6 use 0.1 mm scale. ) there any found by Blair, but Kirkpatrick (1937) noted aedeagal bristles in the extensive material he studied. It is also smaller than the fresh African specimens studied by Kirkpatrick which is doubtless due to the rapid shrinkage of dead material. The wide gap in distri- bution between the slopes of Kilimanjaro and the swamps of Dominica may well be accounted for by the probability that antestiae’s host, a pest on coffee, was imported from Africa along with the coffee tree seedlings for Dominican plantations. REFERENCES Blair, K. G. 1936. A new genus of Strepsiptera. Proc. R. Ent. Soc. London (B) 5:113-117. Bohart, R. M. 1941. Revision of the Strepsiptera with special reference to the species of North America. Univ. California Publ. Ent. 7:91-160. Curtis, John. 1824-1839. British Entomology. 769 col. pls. with descriptive letterpress. London. Esaki, T. and S. Hashimoto. 1931. Report on the leaf-hoppers injurious to the rice plant and their natural enemies. Kyushu Imperial Univ., Dept. Agric., No. 2 (1930):1-59. [See Fox, J., 1967 for translation. ] 242, PROC, ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Fox, Jean W. 1967. Report on some Strepsiptera in the British Museum (Natural History), including new species of Halictophagidae and Elenchidae. Proc. R. Ent. Soc. London (B) 36:41-49. Kifune, Teiji. 1964. Notes on Strepsiptera (1). Insects of the Tsukushi Region. Tsukushi Ent. Soc., Fukuoka, Japan 10:1-14. Kirby, William. 1815. Addendum to the Strepsiptera (Stylops tenuicornis). Trans. Linn. Soc. London 11:233—234. Kirkpatrick, T. W. 1937. Studies on the ecology of coffee plantations in East Africa. II. Trans. R. Ent. Soc. London 86:247-343. Lindberg, Hakan. 1939. Der parasitismus der auf Chloriona-Arten (Homop- tera, Cicidina) lebenden Strepsiptere Elenchinus chlorionae n. sp. sowie die einwirkung derselben auf ihren wirt. Acta Zool. Fennica 22:1-179. Luna de Carvalho, E. 1956. Primeria contribuicéo para o estuda dos Estrep- sipteros angolenses. Pub. Cult., Co. Diamant. Angola (Lisbon) 29:5—54. Ogloblin, A. A. 1925. The Strepsiptera of the collections of the entomological department of the National Museum, Prague. Shornik Ent. Odd. Narod. Mus. Praze 3(24):171-176. Perkins, R. C. L. 1905. Leaf hoppers and their natural enemies. Bull. Ha- waiian Sugar Planters’ Assn., Exper. Sta. Bull. 1(3):90-111. Pierce, W. D. 1907. A preliminary review of the classification of the order Strepsiptera. Proc. Ent. Soc. Wash. 9:75-85. 1909. A monographic revision of the twisted-winged insects comprising the order Strepsiptera Kirby. Bull. U. S. Nat. Mus. 66:232 pp. 1918. The comparative morphology of the order Strepsiptera together with records and descriptions of insects. Proc. U. S. Nat. Mus. 54( 2242 ):391-501. 1961. A new genus and species of Strepsiptera parasitic on a leaf- hopper vector of a virus disease of rice and other Gramineae. Ann. Ent. Soc. Amer. 54(4):467-474. Westwood, J. O. 1835. Description of a new strepsipterous insect recently discovered in the Island of Mauritius. Trans. Ent. Soc. London 1:173-174. A NEW SPECIES OF THE GENUS MACROCEPHALUS SWEDERUS FROM MEXICO (HEMIPTERA : PHYMATIDAE ) NicHoLas A. KormiLev, 365 Lincoln Pl., Brooklyn, New York 11238 Through the kind offices of Dr. Joseph C. Shaffner, Associate Pro- fessor at the Texas A. & M. University, College Station, Texas, I have had the privilege of examining a small lot of Macrocephalus Swederus from the southern United States and Mexico, for which I express to him my sincere thanks. In this lot were six known and one new species. Five of the species are common in this area: M. cimicoides Swederus, five specimens PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 243 from Texas; M. incisus Stal, seven specimens from Mexico; M. pre- hensilis (Fabricius), 12 specimens from Mississippi, Missouri and Texas; M. stali Handlirsch, three specimens from Texas and Mexico. The rare Mexican species M. spiculosus Champion was represented by a lone specimen from Simojovel, Chiapas. The new species is described below. Macrocephalus (Lophoscutus) spiculiger, n. sp. (Fig. 1) Male. Closely related to M. lepidus Stal and running to it in the Handlirsch’s key for Macrocephalus species (1897:182). It is of the same shape and color, only the male has more black color, and less brown, and the size is slightly smaller (5.5 mm.). The main differences are: fore disc of pronotum in both sexes with small, sharp, erect, black spicules instead of round granules; antero- lateral-anterior borders of pronotum finely denticulate, bearing yellow spicules instead of rounded granules, the spicules slightly longer and not so sharp as those on fore disc. Other characters as in M. lepidus. Head cylindrical, brown, densely covered with fine, sharp, black granulation. Antennae of male with segments I and IV black, II and III brown with black spot; of female, I-III orange yellow, IV green. Pronotum with anterior border sinuate; antero-lateral angles acute, slightly divergent; antero-lateral-anterior borders straight, convergent forward, denticulate; lateral notch shallow, sinuate; antero-lateral-posterior borders convex, rounded, very finely granulate; lateral angles barely incised; postero-lateral borders deeply sinuate; posterior angles small, acute; posterior border convex, rounded medially. Fore disc moderately inflated, declivous, brown with dense, erect, sharp, black spicules; antero-lateral-anterior borders, and lateral notch, pale yellow; postero- lateral borders rimmed with white; rest of pronotum black. In the female, black color replaced by testaceous, only spicules on fore disc black. Hind disc roughly punctured. Scutellum tongue-shaped, reaching slightly over the tip of abdomen in both sexes. Median carina linear (Lophoscutus type), clearly visible, deeply depressed base pale yellow, elsewhere convex (in the female greenish-yellow). Disc very roughly punctured at base, punctures progressively diminishing toward tip, there one-fourth as large. Color: base, large elongate ovate spot in the middle of disc, transverse band, and apical portion, black, all 3 spots contiguous with one another; 2 (1 +1) lateral spots on fore half, and 2 (1+ 1), larger lateral spots on hind half, pale yellow, with exception of 6 (3+ 3) small spots on the latter, which also are black. In female, black color replaced by testaceous; lateral borders of hind half, and tip of disc testaceous with a few round, brown dots. Corium black, exterior border pale yellow. In female, testaceous and greenish yellow. Disc with fine, whitish granules. Abdomen strongly rounded laterally, more so in female. Connexivum wide and horizontal; connexivum II in both sexes with a narrow, reflexed edge slightly produced beyond outer border of connexivum III, other PE-angles of connexiva not protruding. Ventral side: head yellowish green, rostrum green, pleurae and sternum orange yellow, venter and legs pale yellow to green; fore tibiae, tips of median and hind 244 PROC. ENT, SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 eno Fig. 1. Macrocephalus (Lophoscutus) spiculiger, n. sp., 6. A = right paramere. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 245 tarsi, and spots on connexivum IV, blackish. Fore tarsi absent. Paramere of type in subgenus Lophoscutus. Measurements: first figure in ratios represents the length and the second the width of measured part, 25 units equal to 1 mm. Head: ¢—29:17.5, 92—30: 18; relative length of antennal segments, I to IV, are: 6 —9(5):5(3.5):6(3):14(6), 9—8(4.5):4(3.5):5.5(3):10(5.5), figures in parentheses represent maximum width of segment; pronotum: ¢—35:52 (maximum width across fore lobe 25), 9 —34:52.5 (maximum width across fore lobe 29); scutellum: ¢—73:40, 9— 73:40; abdomen: ¢6—75:68, 9—71:72 (maximum width in both sexes is across segment III). Total length: ¢—5.52, 2—5.40 mm.; width of pronotum: 4¢—2.08, 9—2.10 mm.; width of abdomen: ¢—2.72, 2—2.88 mm. Holotype: $, MEXICO, Guerrero, 6 mi W of Iguala—P. M. & P. K. Wagner collectors, 10.VII.1966; USNM 69955. Allotype: 2°, MEXICO, Guerrero, 12 mi S of Chilpancingo, same collectors, 12.VII.1966; in the U. S. National Museum collection. REFERENCE Handlirsch, A. 1897. Monographie der Phymatiden. Ann. Naturh. Hofmus. Wien 12( 2) :127-230. NOTES ON THE SYSTEMATICS AND MORPHOLOGY OF THE LACEBUG SUBFAMILY CANTACADERINAE (HEMIPTERA: TINGIDAE ) RicHARD C. FROESCHNER, Smithsonian Institution, Washington, D. C. 29560 Deposition of the Carl J. Drake collection of Hemiptera-Heter- optera in the U. S. National Museum, especially the remarkably complete accumulation of known taxa of lace bugs, provided a unique opportunity to prepare a needed key to the tingid genera of the world. Based on that collection, the recent Drake and Ruhoff (1965a ) catalogue, and appreciated help from NSF grant GB-791, such a key is being constructed. The necessity for changes and additions to the information in that catalogue and the desire to keep the generic key in a certain format leads me to early publication of changes and descriptions of new taxa. The subfamily Cantacaderinae appears to be a natural and valid group whose included genera fall into two categories characterized by Drake and Davis (1960:78) as the tribe Cantacaderini with a “stenocostal area” (the single outermost row of cells of the costal area 246 PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 set off by a thickened vein) and the tribe Phatnomini lacking the stenocostal area. My attempts to verify the correct tribal placement of the included genera revealed that the dorsal evidence of the stenocostal area was matched by a more complex development ventrally which could be present without evident dorsal expression. The ventral development of the stenocostal area (fig. 1) consists of the outer row of cells being confined between two thickened veins which subbasally curve abruptly mesad and continue a parallel course with a narrow groove between them. The inner end of this groove disrupts the hypocostal lamina (uninterrupted in other Tingidae) and is directly in line with the peritreme extending from the scent gland opening. Such morphological alignment and possible functional inter- dependence imply a complex genetic control during the formative period. This ventral expression of the “stenocostal area” confirmed all tribal assignments of genera in the Drake and Ruhoff (1965a) catalogue except that of Stenocader Drake and Hambleton whose lack of dorsal expression of the stenocostal area led to its placement in the Phatnomini. The ventral modifications are, however, very strongly developed on Stenocader so it is here transferred to the tribe Cantacaderini. Cyclotynaspis Montandon Cyclotynaspis Montandon, 1892:265. Cyclotynaspis acalyptoides Montandon Cyclotynaspis acalyptoides Montandon, 1892:265. This species was known only from the Singapore holotype for more than 70 years. Fortunately it recently was rediscovered by Dr. D. H. Murphy, University of Singapore, who found six adults and one nymph during his study of the forest litter fauna in Singapore. Study of this series confirms the Drake and Ruhoff placement of the genus in the tribe Phatnomini. The type and all adults studied were females. Gonycentrum Bergroth, rev. gen. Teleia Fieber 1844:56. Preoccupied by Teleia Hiibner, 1825, in Lepidoptera. Gonycentrum Bergroth, 1898:9. Malala Distant, 1910:101. New synonymy. Diagnosis: This is the only genus of the tribe Phatnomini with the combina- tion of the exposed scutellum and a long, slender, horizontal spine directed for- ward above each eye. Type species: of Teleia Fieber, T. coronata Fieber, monobasic; of Gonycentrum Bergroth, Teleia coronata Fieber, objective synon- ymy; of Malala Distant, M. bulliens Distant, monobasic and original designation. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 247 StA Fig. 1, Cantacader quadricornis (Le Peletier and Serville). Lateral view of head, thorax and bases of wing and abdomen. HcL = hypocostal lamina; SGO = scent gland opening; StA = stenocostal area. Comments: In the Drake and Ruhoff (1965a) catalogue, Gony- centrum contained 12 species from three different zoogeographic subregions: Ethiopian, Oriental and Australian. During the search for characters with which to construct a key to the species, it soon became evident that certain head structures, which are generically important throughout the family, were associated with the species from each zoogeographic subregion suggesting that this was a com- posite genus. The name Gonycentrum must follow the type species of the preoccupied name Teleia which it replaces. Thus the concept of Gonycentrum is returned to its original monobasic status containing one Oriental species. For the other two groups of species formerly assigned to the inclusive use of Gonycentrum, the name Sinalda Distant can be resur- rected from synonymy for the species of southern Africa, and a new genus must be proposed for those from the Australian Region; for the latter I propose below the name Carldrakeana. The genus Malala also becomes involved with Gonycentrum because both genera are based on the same species. Subsequent to their catalogue, Drake and Ruhoff (1965b) described Malala charieis from New Guinea. Unfortunately, this species cannot follow Malala bulliens Distant into Gonycentrum because it is generically distinct. In fact the lack of paired spines on the vertex combined with certain other structures prevents its assignment to any known genus. A new genus, Distocader, is proposed for it below. To summarize all the generic changes necessitated by this redefini- Q 248 PROC. ENT, SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 tion of Gonycentrum, a key to the pertinent genera is offered below and followed by the necessary generic discussions and descriptions: 1. Head with a prominent tubercle on median line between eyes —______- Ds Head without a tubercle on median line between eyes 3 Head with 3 tubercles on vertex, a median one and one above each eye (South Africa) _..... ; ae Sinalda Distant Head with one tubercle on Roe fhe median one, none fabore eyes (New Guinea) _. eee Distocader, n. gen. Vertex with a lone Jee spine arising near eee chong each eye and extending horizontally and anteriorly to or beyond anterior margin of eye (“Ostindien,” India, Ceylon) ——....____________ Gonycentrum Fieber Vertex without spines or tubercles above eyes (Australia, New Guinea) ~whiso” eae fesse eee see ete eter ne eeeta see Carldrakeanayenescens Lo (ee) Considering the good figure of Teleia coronata given with the original description by Fieber and the proximity of the type localities of coronata and Malala bulliens, it is quite surprising that Distant and all subsequent workers failed to recognize that both names belong to the same species. The fact that both Fieber’s and Distant’s excel- lent illustrations show the uncovered scutellum and the long, supra- ocular spines on the vertex leaves no other conclusion but synonymy as this combination is known in no other tingid. Drake and Ruhoff’s (1965b:244) placement of their species charieis in Malala appeared to result from the superficial similarity of their species to bulliens, but the head armature makes this untenable and forces the creation of a new genus for their species. List OF SPECIES OF Gonycentrum coronata (Fieber), 1844, p. 56—Teleia, [Bergroth, 1898, p. 9]. “Ostindien.” = Malala bulliens Distant, 1910, p. 101. New synonymy. Ceylon. Sinalda Distant, rev. gen. Sinalda Distant, 1904:426. Diagnosis: Among those genera of the tribe Phatnomini bearing a spine or prominent tubercle on the midline of the vertex, Sinalda may be recognized by also having paired spines or tubercles on the vertex and no lobular or spinelike projections on the horizontal or slightly oblique (not recurved) paranota. Type of genus: Sinalda elegans Distant, subsequently designated by Monte (1947:4). Sinalda was made a synonym of Gonycentrum by Drake (1950:165). When the present study found Gonycentrum to contain three groups of species distinguishable on certain head structures coupled with geographic distribution, Sinalda was available as the generic name for the species of southern Africa (see above treatment of Gonycentrum for details). Eight of the previously described species of lace bugs plus one new one belong to Sinalda. PROC. ENT, SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 249 Sinalda haplotaxis, n. sp. Diagnosis: Among the named macropterous forms within the genus, this species is distinguished by the costal area containing only a single row of subquadrate areolae (except in basal fifth) which are distinctly larger than the areolae of the subcostal area. It appears to be closest to thomasi (Drake) in which the costal area has two or three rows of areolae for full length. Characters: Holotype male, length 2.3 mm.; macropterous. Head dorsally with 8 stout cephalic tubercles: one pair each on juga, above clypeus, and above eyes, plus a single tubercle each medially on the vertex and clypeus; bucculae low (less than half the height of an eye), uniseriate, slightly surpassing and in contact beyond apex of clypeus; labium reaching between hind coxae; antennal segment I little stouter and longer than II, segments III and IV missing. Pronotum without inflated cysts, anterior margin weakly concave, slightly convex behind eye; disc with 3 longitudinal carinae plus vague, longitudinal cal- loused swelling above each humerus; paranota distinctly wider than_ tibial diameter, with 2 rows of areolae from humeri forward, outline constricted oppo- site calli; posterior margin transverse, nearly straight, exposing scutellum. Elytra nearly flat (most of left one missing, areas well defined by elevated prominent veins; costal and subcostal areas subdivided by two and four, respectively, elevated thickened crossveins; costal area narrow, with 1 regular row of sub- quadrate areolae (except in basal fifth) larger than areolae of subcostal area. Scent gland opening obscure, without elevated auricular peritreme. Sternal laminae straight, parallel on all three thoracic sterna. Abdomen somewhat longi- tudinally impressed mesoventrally near base. Holotype male: Transvaal, December 30, 1956, A. L. Capener (USNM 69913, in Carl J. Drake collection ). The species name is derived from two Greek words meaning “single” and “row” and reflects the virtually wholly uniseriate costal area. List OF SPECIES OF Sinalda aethiops (Distant) 1902, p. 238—Phatnoma. New combination. South Africa. afra (Drake and Ruhoff), 1961, p. 126—Gonycentrum. New combination. South Africa. angustata (Drake), 1956, p. 15—Gonycentrum. New combination. Tanganyika. elegans Distant, 1904, p. 428—Sinalda. Restored combination. South Africa. haplotaxis, new species (see above). Transvaal. nebulosa Distant 1904, p. 428—Sinalda. Restored combination. South Africa. reticulata Distant, 1904, p. 427—Sinalda. Restored combination. South Africa testacea (Distant), 1902, p. 238—Phatnoma. New combination. South Africa. thomasi (Drake), 1956, p. 14—Gonycentrum. New combination. Kenya. Distoeader, n. gen. Diagnosis: Among those genera of the tribe Phatnomini bearing a cephalic spine or tubercle on the midline of the vertex, this one can be recognized by having no pairs of spines on the vertex and no lobular or spinelike projection on the margin of the oblique paranotum. Characters: Head with 6 tubercles (a pair above base of clypeus, a pair of weak tubercles on juga, one each on midline of vertex and clypeus); eyes more 250 PROG. ENT, SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 than half as wide as interocular space; bucculae reaching apex of clypeus, not incurved anteriorly; labium reaching between hind coxae; antennae (missing from only available specimen, character derived from original illustration of type species) with segment III about 5 times as long as I plus II. Pronotum without inflated cysts; anterior margin weakly concave; disc tri- carinate, median complete, percurrent, dorsal outline angularly emarginate above calli, lateral simple, reaching to calli; paranota obliquely elevated, biseriate except around humeri where uniseriate; posterior margin transverse, nearly straight, exposing scutellum. Elytra conjointly weakly convex; areas separated by thick- ened veins; vein between discoidal and subcostal area elevated, containing a row of subquadrate areolae; subcostal area quadriseriate, costal area weakly oblique, biseriate for nearly full length; hypocostal lamina uniseriate. Sternal laminae present, low, areolate on all thoracic sterna, diverging on metasternum. Peritreme elevated, transversely oval auricle. Abdomen convex basomedially. Type of genus: Malala charieis Drake and Ruhoff, present desig- nation. Geographic distribution: The lone known specimen was from New Guinea. Comments: In the absence of any outstanding feature to be signaled by the generic name, the Latin word disto, meaning “be separate” or “differ” is combined with “cader” which is commonly used as a suffix in generic names in this subfamily. List OF SprEcIES OF Distocader charieis (Drake and Ruhoff), 1965:244—Malala. New combination. New Guinea. Carldrakeana, n. gen. Diagnosis: This genus may be recognized within the tribe by the combination of no interocular spines or tubercles, the narrow, straight, slightly oblique, areolate paranota, and the costa gradually widening from the base. Characters: Head with 4 dorsal cephalic spines (a pair above base of clypeus, one on each jugum); eyes almost half as wide as interocular space; bucculae far surpassing apex of clypeus, anteriorly incurved and usually touching each other; labium reaching base of abdomen; antennal segment III 3 to 4 times as long as I plus II. Pronotum without inflated cysts; anterior margin transverse, nearly straight; disc with 1 or 3 longitudinal carinae, median percurrent, lateral carinae, when present, percurrent, interrupted above calli; paranota flat, somewhat oblique, narrow, with 1 or 2 rows of areolae; posterior margin weakly convex to truncate across middle third, exposing scutellum. Elytra conjointly convex; areas distinctly delimited by elevated veins; discoidal and subcotsal areas usually interrupted by prominently elevated cross veins; costal area oblique, narrow, gradually widening from base, with 1 or 2 rows of areolae. Scent gland opening not visible, without elevated peritreme. Abdomen with midventral groove restricted to segments I and II, former with a spinelike prominence on each side of groove. Type of genus: Phatnoma tindalei Hacker, present designation. Geographic distribution: Carldrakeana occurs in Australia, Tas- mania, New Guinea, and New Zealand. PROC. ENT. SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 251 Comments: This taxon is that part of the genus Gonycentrum, in the broad sense, characterized by the absence of spines or tubercles between the eyes. For further discussion of this genus, see the com- ments under Gonycentrum. As is evident, the name proposed for this genus is a dedication to the late Dr. Carl John Drake who, in a half century of studying lacebugs, contributed many important papers and crowned his activi- ties on the group with coauthored elucidations on their morphology and phylogeny (Drake and Davis 1960) and a world catalogue (Drake and Ruhoff 1965a) of the taxa involved. Carldrakeana was misidentified as Cyperobia by Drake and Davis (1960:29, fig. 31) and Woodward (1961:155, figs. 18 and 19) who published illustrations of specimens under the name Cyperobia carec- torum. The lack of an inflated “vesicle” or cyst anteriorly on the pronotum, which was very clearly characterized in Bergroth’s original description of Cyperobia, prevents considering these figures as repre- sentatives of that genus. In fact, Drake and Davis’ sketch and Wood- ward's figure 19 both show Carldrakeana scoia, a conclusion which in part is confirmed by examination of two of Woodward's Stokes Valley specimens now at hand. Woodward's figure 18 clearly runs to Carl- drakeana but presents a combination of characters unlike that found in any of the three species known within the genus. List oF Species OF Carldrakeana engista (Drake and Ruhoff), 1961:127—Gonycentrum. New combination. New Guinea. socia (Drake and Ruhoff), 1961:128—Gonycentrum. New combination. Tasmania. tindalei (Hacker), 1928:177—Phatnoma. New combination. Australia. Minitingis Barber, rev. gen. Minitingis Barber, 1954:7. Diagnosis: Within the tribe Phatnomini, only this genus and Gonycentrum (in the restricted sense used above) possess seven spines on dorsum of head and a long labium reaching to second abdominal sternite. In Gonycentrum the spines above the eyes lie on the surface of the head and project horizontally and anteriorly to or beyond the anterior margin of the eye; in Minitingis these spines are much shorter and obliquely elevated. Type of the genus: Minitingis minusculus Barber, monobasic. Comments: Drake and Ruhoff (1960:28) synonymized Minitingis under Zetekella. Considering only the three species catalogued under that genus by Drake and Ruhoff (1965a) I tentatively accepted this synonymy. But the appearance of a second West Indies species agreeing with minusculus in the narrow form, head armature, long labium, paranotal development, and grooved abdomen created a bo ut bo PROC. ENT, SOC. WASH., VOL. 70, NO. 3, SEPTEMBER, 1968 Fig. 2, Minitingis elsae, n. sp. distinct morphological pattern of West Indies versus continental American species. This pattern appears to have true zoogeographical significance best represented at this time by generic recognition; so Barber’s genus is here resurrected for these two species. 9 PROC. ENT. SOC. WASH., VOL. 70, No. 3, SEPTEMBER, 1968 bo OL ioe) Minitingis elsae, n. sp. (Fig. 2) Diagnosis: The conspicuous black and white color pattern is distinctly unique within the subfamily. Structurally this new species can be separated from the only other member of the genus by the greater number or rows of areolae (4 in contrast to 2) in the costal area. Characters: Holotype male. Length 2 mm.; brachypterous. Head with 7 dorsal spines (one pair each above eyes, above base of clypeus and on juga, plus 1 medially at midlength of clypeus); antennophores anterolaterally pro- longed into prominent spines; bucculae slightly surpassing apex of clypeus; antennal segment II shortest, I slightly longer, HI about 5 times as long as I plus II, IV slightly thickened, about one-third as long as III; labium reaching apex of abdominal sternite II. Pronotum without inflated cysts; anterior margin concave; disc unicarinate, median carina percurrent, elevated, areolated, dorsal margin angularly incised between calli; discal surface including collar but not calli, densely, subcribrately punctured; paranota developed full length, narrow, 3 areolae wide opposite calli, lateral margin with a weak angulation subapically and submedially; posterior margin transverse, weakly convex medially, exposing small scutellum. Elytra con- jointly weakly convex; areas separated by thickened veins; vein between subcostal and discoidal areas laminately elevated and areolated almost to apex, vein sepa- rating discoidal and sutural areas similarly developed except basally between discoidal and claval regions where it becomes almost obsolete; discoidal and subcostal areas subdivided by thickened cross veins; costal and subcostal areas 4 areolae wide for full length; clavocorical suture obliterated, claval commissure distinct, elevated. Scent gland opening surrounded by strongly developed auricular peritreme. Sternal laminae on all segments uniseriate, straight, nearly parallel. Abdomen strongly impressed along basal third of midline. Head and body black; collar dorsally and laterally, apical third of paranota, sternal laminae, metapleura, including peritremes, ivory white; heleytron ivory white, marked with dark fuscous or black as follows: Narrow base and 3 or 4 quadrate areas in costal area; lines along subdividing cross veins and nearby veins of subcostal and discoidal areas; and veins of sutural areas. Antennae and legs brownish. Holotype ¢. Jamaica, intercepted on air freight being carried into the United States on September 19, 1966 (USNM 69914). This beautiful little lace bug is dedicated to my dear wife, Elsie Herbold Froeschner, for her years of understanding and cooperation in my entomological pursuits and her willingness to beautify and increase the usefulness of my papers with her skillfully executed illustrations. List oF SpEcIEs OF Minitingis elsae, new species (see above). Jamaica. minusculus Barber, 1945:7—Minitingis. Restored combination. Bahamas Is. REFERENCES Barber, H. G. 1954. ZY, a Wilh ING aa | St > ey | Neat hea r\f an A No \ / j a, j \ 1 ae 2 \ ] \ | fest se i AN iN Ke » fase} aoe K<-—=--C B 1X i 8 | m& A ee \ ae \ j ) D HY P= od aN | | ~~ / / \ ry west / (SCZ - e / Figs. 1-3, 5-7, Chomatophilus aphanistes, n. sp., holotype: 1, penult, ultimate and postpedal segments, ventral, A — penult sternite, B — coxopleural gland crypts, C = ultimate sternite, D = fused female gonopods, E = anal pore crypts, F = adanal laminae; 2, sixth sternite with flanking subcoxae, ventral, sternital porefield shown, those of subcoxae deleted: 3, right prehensor and adjacent prosternum, ventral, A = serrulate ventral edge, B = poison calyx, C = diastema, D = pleurogram; 5, tenth tarsus and pretarsus, anterior surface, A = paired anterior parungues (deflected proximad), B = single posterior parunguis; 6, first and second maxillae, ventral; 7, clypeus. Figs. 4 and 8, C. smithi Pocock: 4, right prehensor, ventral; 8, clypeus. 326 PROC. ENT. SOC. WASH., VoL. 70, NO. 4, DECEMBER, 1968 posteriorly by fragments. PREHENSORS. Basal articles short and broad, claw relatively long. Flexed, not attaining anterior head margin. Mesal denticles, basal tooth both absent. Ungular ventral edge smooth or irregularly serrulate; ungular dorsal and ventral surfaces not appressed. TRACHEATION. Apart from the usual dorsal oblique connectives with a subdorsal system of trans- segmental trunks running anteroposteriorly. Four thereof arising in segment II passing caudad and joined in Segment VI by four additional, all terminating variously somewhere anterior to body mid-length. PARATERGITES. Absent. LEGS. Short and robust, very sparsely setose. Each pretarsus with two anterior parungues and one posterior parunguis. STERNITES. All wider than long. Sculpturing coarsely areolate. Setae minute, sparse. Carpophagus-structures absent. Those more anterior with very shallow midlongitudinal sulci. Subcoxal porefields present, extensive. Sternital porefields: on one through penult; un- divided, each in the form of a transversely wide posterior band with its middle extended anteriorly in a broad arc, the extension on posterior segments much more extensive than on anterior segments. Formed anterolateral porefields absent, represented by irregularly scattered single spores. ULTIMATE PEDAL SEG- MENT. Pretergite entire, laterally not fissate. Tergite shieldshaped, narrowed, posteriorly; length about equal to greatest width. Presternite medially divided. Sternite: much wider than long; sides and rear margin together describing a broad semicircle. Coxopleuron: only slightly inflated, relatively short and small, not encroaching upon penult segment; each with two ventral cryptic gland cavities, these heterogeneous, composite, with inclusive canals and lumina irregularly formed by a distinctive papillate membrane, the anterior pit often partly concealed by penult sternite. Telopodite: somewhat longer than penult; articles more or less swollen depending upon sex and species; with two tarsalia; pretarsus distinctly unguiform, basally not filamentous.*, POSTEPEDAL SEGMENTS. Male gonopods widely separated, biarticular. Female gonopods each uniarticular, medially fused without demarcation. Anal pores notably large, each internally composite and heterogeneous with constituent canals and papillate membrane therein like the coxopleural crypts. Chomatophilus aphanistes, n. sp. The new species, although indisputably congeneric with smithi, is very different from it in many particulars of infrageneric rank. Grossly different from smithi, aphanistes is shorter and mesopodal; the Pocock form is distinctively polypodal and sizeable. Furthermore in smithi: (1) pedal pairs, 81-115. (2) Tarsungular ventral edge is mostly smooth, with shallow, irregular serrulations only proximally. (3) Clypeal setae robust and numerous; present anterolaterally. (4) Porefield of sternite one with posterolateral extensions. (5) Female ultimate legs notably inflated. (6) Second maxillary coxosternal lappets absent. (7) Tarsungular blade far exceeding length of basal articles. In aphanistes: (1) pedal pairs, 47-59. (2) Tarsungular ventral edge finely, regularly serrulate over entire length. (3) Clypeal setae delicate, less numerous; absent anterolaterally. (4) Porefield of sternite one strictly subcentral, without posterolateral extensions. (5) Female ’ Compare with Garrina, wherein rear leg pretarsi, including the ultimate, al- though typically unguiform, are basally conspicuously fibrous, which in the Geophilomorpha is most uncommon. 7 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 32 ultimate legs at most very slightly inflated. (6) Second maxillary coxosternal lappets present. (7) Tarsungular blade not greatly exceeding length of basal articles. There are additional differences, but the foregoing seem the most useful and obvious. Holotype: female. Mexico, San Luis Potosi, on route 80 4.3 miles east of Ciudad del Mais at 4,300’ elevation. August 10, 1966. G. B. Ball and D. R. Whitehead, legg. Deposited in the U. S. National Museum. GENERAL. Length, 30 mm. Pedal segments, 57. Body shape, anteriorly and posteriorly slightly attenuate. Color: antennae and dorsum pale yellow; venter sordid white. ANTENNAE. Length to head length, 8:3. Filiform, distally attenuate slightly, slightly flattened dorsoventrally. CEPHALIC PLATE. Greatest width to length, 9:8. Setae very short and sparse. Frontal suture barely visible as a band of weaker areolation. Prebasal plate slightly exposed medially. CLYPEUS. Paraclypeal sutures prominent and complete. Anterocentral fenestra and prelabral plagulae absent. Setae: relatively long and delicate; few in number, forming two groups, a central one of 4 girdled by a setal semicircle; none anterolaterally. LABRUM. Sidepieces merging with midpiece. Entire labral margin fringed with hyaline filaments, these very long on sidepieces but notably shorter on midpiece. FIRST MAXILLAE. Coxosternum with 15 setae dispersed irregularly in two files. Medial lobes relatively long and wide, their intervening diastema comparatively deep. Coxosternal lappets narrow, pointed, much shorter than telopodite lappets. Telopodites: bipartite, each with a thick scabrous lappet. SECOND MAXILLAE. Isthmus comparatively shallow; medially areolate, neither membranous, suturate, nor diastemate. Postmaxillary sclerites absent. Telopodite: prominent dorsal and ventral basal condyles present; rela- tively short and inflated; without denticles or protuberances; terminal claw short and weak, not exceeding neighboring setae. PROSTERNUM. Greatest width exceeding exposed length. Anteromedial diastema broad and relatively deep, denticles absent. Pleurograms prominent, complete to condyles. Pleuroprosternal sutures entirely absent. PREHENSORS. Flexed, not attaining anterior head margin. Mesobasally with a very low swelling. Ungular blade ventrally coarsely and irregularly serrulate; ungula not notably recurved, not uncinate. Poison calyx in lower tarsungula; poison gland terminating in trochanteroprefemur. STERNITES through penult. Setae short, very sparse. Each with a very shallow midlongitudinal depression. Carpophagus-structures absent. Porefields: present on first through penult; on anterior sternites (except first) each a posterior, trans- verse, uninterrupted band medially extended forward in a low, broad arc; on more posterior sternites the mediocephalad extensions become wide and very long eventually occupying most of the paramedian length of each sternite. LEGS. Setae very sparse and minute. Rear legs dorsally not flattened. Pretarsi: each with two anterior and one posterior parungues, essentially equal, minute. ULTI- MATE PEDAL SEGMENT. Pretergite very wide, not fissate laterally. Tergite greatest length about equal to greatest width. Presternite medially entirely sepa- rated by wide membranous area. Sternite: greatest width far exceeding length; sides and posterior margin conspicuously rounded, nearly semicircular. Coxo- pleuron: comparatively small, slightly inflated, a patch of setae on posteroventral surface, otherwise nearly glabrous; each with two deep cryptic gland cavities, 328 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 these of the heterogeneous type with many constituent canals, their lumina lined with a distinctive papillate membrane.‘ Telopodite: longer than penult; the articles but slightly swollen; the two tarsal articles equal in length and diameter; pretarsus clearly unguiform. POSTPEDAL SEGMENTS. Sparsely setose. Female gonopods uniarticulate and completely fused medially without demarcation. Anal pores, like coxopleural crypts, internally heterogeneous and lined with convoluted papillate membrane. Paratypes. There are 37 paratypes representing many localities in four Mexican States. All agree closely with the holotypic description, except that in the males the ultimate legs differ in being much swollen. Pedal segments, 47-59, mode 51. Lengths, 15-30 mm. U. S. National Museum collection. VERACRUZ. Slope of Vulcan San Martin north of San Andres Tuxtla, July 20-August 1, 1959, Keeton and Valentine. San Andres Mountains, Bastanal, 2500’—3000’, September 19, 1965, Ball and White- head. Lake Catemaco, Coyame, July 2, 1963, Whitehead. San Andres Mountains, 25 miles west of Contecompan, 100’, September 18, 1965, Ball and Whitehead. SAN LUIS POTOSI. 18 miles east of Ciudad del Mais, September 3, 1964, Awram and Whitehead. 24.7 miles east of Landa de Matamoros, 5000’, October 18, 1965, Ball and Whitehead. 4.3 miles east of Ciudad del Mais, July 10, 1966, 4300’, route 80, Ball and Whitehead. Ciudad del Mais, September 3, 1964, Whitehead. TAMAULIPAS. 8.1 miles west of Encino, 3100’, October 11, 1965, Ball and Whitehead Gomez Farias, Aqua Livida, 5800’, October 9, 1965, Ball and Whitehead. 8 miles northwest of Gomez Farias, Rancho del Cielo, 3800’, August 6, 1965, Ball and Whitehead. OAXACA. 21.8 miles north of Tuchatengo, July 18, 1966, Ball and Whitehead. Chomatophilus smithi Pocock Chomatophilus smithi Pocock, 1896, Biol. Centr. Amer., Chilopoda, p. 39. Chomatophilus smithi Pocock: Attems, 1929, Tierreich, Lief. 52, p. 349. (As Geophilomorpha incertae sedis. ) GENERAL. Length, to 90 mm. Pedal segments, 81-115. Shape, anterior and posteriorly attenuate. Color: antennae and dorsum yellowish-orange; venter sordid white. ANTENNAE. Length to head length ca. 11:4. Filiform, distally slightly attenuate, dorsoventrally slightly flattened. CEPHALIC PLATE. As wide as long to slightly longer than wide. Setae short and sparse. Frontal suture barely visible as a band of weaker areolation. Prebasal plate slightly exposed medially. CLYPEUS. Paraclypeal sutures prominent, wide, complete. Antero- central fenestra and prelabral plagulae absent. Setae: relatively long, notably robust and numerous; only on anterior half of clypeus; present anterolaterally. LABRUM. Sidepieces margin with midpiece without clear demarcation. Entire labral margin fringed with hyaline filaments, those of sidepieces much longer than those of midpiece. FIRST MAXILLAE. Coxosternum with some 30 robust setae, these irregularly disposed. Medial lobes long and very wide, the intervening * For further discussion of this membrane in sogonids and other taxa see Crabill, 1961, Proc. Ent. Soc. Wash. 63:132. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 329 cleft deep. Coxosternal lappets absent. Telopodites: bipartite, each with a robust scabrous lappet. SECOND MAXILLAE. Isthmus comparatively shallow; medially areolate, not hyaline or suturate or diastemate. Postmaxillary sclerites absent. Telopodite: prominent dorsal and ventral condyles present; relatively short and much inflated; without denticles or protuberances; terminal claw rather long, exceeding neighboring setae. PROSTERNUM. Greatest width exceeding visible length. Anteromedial diastema broad and deep, with low evanescent denticles in most. PREHENSOR. Flexed, not surpassing front of head. Without mesal denticles but tarsungula mesobasally swollen. Ungular blade smooth for most of its length (proximally with a few shallow serrulations). Poison calyx in lower tarsungula. Poison gland terminating in lower trochanteroprefemur. STERNITES through penult. Setae short and very sparse. Each with a very shallow midlongitudinal depression. Porefields: present on first through penult; on anterior sternites (including the first) each a very wide, transverse and un- interrupted band medially extending forward in a prominent arc; on posterior sternites the mediocephalad extension becoming very wide and long, eventually occupying most of the paramedian length of each sternite. LEGS. Setae very sparse, minute. Rear legs dorsally somewhat flattened. Pretarsi: each with two” anterior and one posterior parungues, these essentially equal and minute. ULTIMATE PEDAL SEGMENT. Pretergite wide, laterally not fissate. Tergite greatest width about equal to length. Presternite medially broadly divided. Sternite: greatest width far exceeding length; sides and rear conspicuously rounded, nearly semicircular. Coxopleuron: comparatively small, slightly inflated, with a patch of setae ventroposteriorly; each with two deep cryptic gland cavities, these of the heterogeneous type with numerous constituent canals, their lumina lined with a distinctive papillate membrane. Telopodite: longer than penult; the articles in both sexes notably inflated; distotarsus notably shorter than proximotarsus; pretarsus distinctly unguiform. POSTPEDAL SEGMENTS. Sparsely setose. Female gonopods uniarticular, medially broadly fused without demarcation; male gonopods biarticular, medially widely separated. Anal pores, like coxopleural crypts, internal heterogeneous with inclusive canals and lining papillate membrane. The holotype in the British Museum (number 1897.3.1.127) has no more precise locality than “Amula” on its ticket. In the original de- scription Pocock places Amula in the State of Guerrero at 6000 to 7000 feet elevation. Several very extensive Mexican gazetteers fail to reveal this place name, but they do give a number of listings for Amole, at least one of which is in Guerrero. It seems possible that Amula is a misspelling of Amole. I have examined the holotype, and in addition specimens collected in the following localities. SAN LUIS POTOSI. 19.3 miles northwest of Tamazunchale on route 85. Near Ciudad del Mais. NUEVO LEON. Chipinque mesa near Monterrey. QUERA- TARO. Near Pinal de Amoles. TAMAULIPAS. Near Gomez Farias, Racho del Cielo. ®> The presence of two anterior parungues, instead of the usual one, is most uncommon in the Geophilomorpha. Elsewhere I have encountered it only in the Schendylidae. 330 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Key to Soconi GENERA The following key is partly based upon Chamberlin’s published descriptions and keys, which portions remain to be verified through direct recourse to specimens. I assume responsibility for the placement and identification of those genera (signaled by asterisks) material of which I have studied. Those familiar with Chamberlin’s several sogonid keys (e.g. in Univ. Utah Biol. Series, VII (3):17, 1943) will note that, unlike him, I have not dichotomized the conditions of the ultimate pretarsus, that is, pretarsus a claw versus pretarsus a tubercle, which in my key might have been introduced in the fifth couplet. This is because I have found both conditions to be intragenerically variable within Garrina and Sogona. In fact, I have seen specimens of both genera wherein in the same specimen one ultimate pretarsus could be called tuberculate, the other unguiform. la. Ultimate tarsus consisting of one article; ultimate pretarsus absent; each coxopleuron with oneterypt 2 ee Timpina Chamberlin Ib. /Ultiniate’ tarsus consisting of ‘two articles: 0... 2 3 a ee 2a. Each coxopleuron with one crypt ds Gosipina Chamberlin 2b. Each coxopleuron with two ae sie tisenhel otto t le ede 3 3a. Ventral pores absent; ultimate pretarsus aigent _....... Oligna Chamberlin Sb Ventral: pores present) {ice se leno te bo Ay eer es 4a. Tarsungula with basal tooth —. ss s«~Portoricona Chamberlin* 4b. Tarsungula without basal tooth —__ aes eee = 15: 5a. Pleuroprosternal sutures entirely absent; Menate) steunite octenornn Greadis TOUUGCLEC fete eat aS te a Hac Se Chomatophilus Pocock* 5b. Pleuroprosternal sutures present, complete or broadly incomplete; ultimate sternite posteriorly not broadly rounded _.... = 22 1S eee 6 6a. Pleuroprosternal sutures widely snBorelete) not redehing anterior margin iY : : : woe et . (Carrina Chamberlin’ 6b. Bleiropiostennal = sutures Peeinicie. ee anterior marecin 2 pat eC eee = ee een Ye meri ciire) (Clivnienlavainlin™ CATALOGUE OF SOGONID GENERA The following names, not all of which belong there, have been referred to Sogonidae. An asterisk indicates I have seen material. Andenophilus Verhoeff (? = Oryidae ) Andenophilus Verhoeff, 1942, Titschack’s Fauna Perus, II, p. 62. Type-species: A. striatus Verhoeff, 1942. Monobasic. Remarks: Except for the mandible, Verhoeff has described a very typical oryid. Indeed, his species is otherwise so oryid that I can only believe him to have erred in describing the mandible. But apart from the question of what Andenophilus is, it is indubitably not a sogonid. Chomatophilus Pocock* Chomatophilus Pocock, 1896, Biol. Centr. Amer., Chilop., p. 39. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 331 Nuevona Chamberlin, q.v. New synonymy. Type-species: C. smithi Pocock, 1896. Monobasic. Garrina Chamberlin* Garrina Chamberlin, 1915, Bull. Mus. Comp. Zool. Harv. 59:506. Pycnona Chamberlin, q.v. New synonymy. Type-species: G. ochra Chamberlin, 1915. Original designation. Gosipina Chamberlin Gosipina Chamberlin, 1940, Pan.-Pac. Ent. 16:56. Type-species: G. bexara Chamberlin, 1940. Original designation, monobasic. Idiona Chamberlin (= Arctogeophilus, new synonymy ) Idiona Chamberlin, 1946, Can. Ent. 78:69. Type-species: I. shelfordi Chamberlin, 1946. Original designation, monobasic. Remarks: The genus can only be a junior synonym of the chilenophilid genus Arctogeophilus. Nuevona Chamberlin (= Chomatophilus, new synonymy ) Nuevona Chamberlin, 1941, Pan.-Pac. Ent. 17:185. Type-species: N. leonensis Chamberlin, 1941. Original designation and mono- basic. Oligna Chamberlin Oligna Chamberlin, 1943, Univ. Utah Biol. Ser. 7:19. Type-species: O. pueblana Chamberlin, 1943. Original designation and mono- basic. Remarks: I believe there is at least some reason for suspecting the genus to be a junior synonym of the dignathodontid Pagotaenia Chamberlin, but under the circumstances this cannot be proved without recourse to the types. Nonetheless, the reader is alerted to the possibility. Portoricona Chamberlin* Portoricona Chamberlin, 1950, Proc. Biol. Soc. Wash. 63:159. Type-species: P. adjunta Chamberlin, 1950. Original designation. Pyenona Chamberlin* (= Garrina, new synonymy ) Pycnona Chamberlin, 1943, Univ. Utah Biol. Ser. 7:18. Type-species: P. pujola Chamberlin, 1943. Original designation. Sogona Chamberlin* Sogona Chamberlin, 1912, Bull. Mus. Comp. Zool. Harv. 54:431. Type-species: S. minima Chamberlin, 1912. Original designation and mono- basic. Timpina Chamberlin Timpina Chamberlin, 1912, Bull. Mus. Comp. Zool. Harv. 54:433. Type-species: T. texana Chamberlin, 1912. Original designation and monobasic. SOL PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 ON THE LIFE OF VESPULA VULGARIS (L.) AND Vv. MACULIFRONS (BUYSSON) (HYMENOPTERA; VESPIDAE) W. V. BaLpur, Department of Entomology, University of Illinois, Urbana, Illinois 61803" The observations reported here pertain to living colonies of Vespula vulgaris (L.) and V. maculifrons (Buysson) taken at Eaglenest Lakes near Ely in northeastern Minnesota. They have to do with nests, seasonal development and certain interrelations of the two species. V. vulgaris and maculifrons are close relatives in the subgenus Vespula. LirE OF Vespula vulgaris The six colonies, identified by capital letters A-F, are described separately and treated in chronological order of the months and days on which they were removed from their nesting sites for analysis. Thus they afford data pertinent to the seasonal distribution of the several castes. A. The colony developed inside a small dark pumphouse, where the nest was attached to the door casing. Two combs were under construction on July 1, 1961. The central or discal zone of comb | had yielded 20 workers, of which only 14 were present when the colony was inactivated at night with cyanide gas. The mother queen also was taken. Outwardly from the discal zone there occurred suc- cessively (1) a zone of 41 capped cocoons containing pupal workers, (2) a zone of submature to small worker larvae and (3) a zone of peripheral cells containing eggs. Moreover, the second life cycle or brood generation was represented by eggs and small larvae in the renovated discal cells in which the workers had developed. The small apical or distal comb comprised about 51 incomplete cells that housed only eggs and small larvae. Previous observations on Vespula at Eaglenest Lakes indicate that colony A was founded on or about June 1. Development through the embryonic, larval and pupal stages of Dolichovespula maculata (L.) required 26-30 days (Balduf, 1954). Therefore it is estimated that the first worker vulgaris emerged from their cocoons shortly before June 28. B. The nest hung in full view from the ship-lap siding on the shaded south side of a cabin. First seen on June 19, 1959 (fig. 1), it was left intact until July 10. The process of applying paper pulp, by the sole worker present in the nest, was as described by Duncan (1939, p. 122). 1My thanks go to Messrs. Karl V. Krombein, U.S. National Museum and C. D. F. Miller, Entomology Research Institute, Ottawa, Canada, for assistance in determining the wasps. PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 333 However, one aspect of this procedure seems to have been overlooked. This I have seen frequently also in D. maculata and D. arenaria (Fabr.). It is as follows. When the vulgaris worker returned from the field bearing a mass of paper pulp destined for application to an incomplete layer of the nest envelope, it did not advance directly to apply the material, but invariably first entered the nest. After passing a few seconds there, it emerged, still holding the pulp intact, and chose a place on the envelope, where it then applied it in the meticulous manner described by Duncan. Having thus disposed of the load, the worker did not fly directly from the envelope to the field, but again entered the nest for another brief stay, after which it emerged to fly to the field. All activity had ceased in this nest before July 10. The envelope consisted of two almost complete outer layers, and two inner ones whose apical parts had been cut off. The single comb was circular, only 21 mm in diameter, and comprised 42 uniformly small cells. Workers had developed in 6 cocoons in the discal zone, while three contained mummied pupae. The remaining 33 outer cells remained incomplete. One worker lay dead in the nest. The queen seems to have been absent since before June 19. C. The oval shape of this nest conformed to the abandoned lair of a chipmunk (Eutamias) in which it had developed. The top remained visible through the oblique runway of the mammal, and the bottom rested on the floor of the cavity nine inches below the surface of the ground. When dug out on August 15, 1956, the nest comprised four roundish combs. Beside the 577 active workers captured, 550 white-capped pupal cells had been formed in the basal combs. These cells were of the same size as those from which workers had emerged, hence con- tained worker pupae. Also some prospective workers probably re- mained in the basal combs as second brood larvae, while some adult workers had already died. The mother queen, taken when the nest was disassembled, remained actively reproductive. Juvenile males and queens had not appeared but probably existed as larvae in combs 3 and 4. D. The nest occurred in a shallow subterranean cavity that had been formed by collapse of a rotting log. Latter was covered by a layer of small boulders and soil. Access to the nest was by a round hole % in. in diameter at the base of a quaking aspen. Four combs of irregular shape had been constructed. A total of 528 vulgaris was captured after dark of August 17-25, 1955. They comprised 524 workers, three juvenile males and the dead mother queen, that had concealed herself between combs | and 2 in the top of the nest. Pro- duction of adult reproductives had just begun. This colony also contained seven workers of V. maculifrons. 334 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Fig. 1. Aerial nest of Vespula vulgaris (L.), frontal view, somewhat diagram- matic: a, suspensorium; b, entrance; c, c, edge of nest; d, d, dashed lines represent margins of strips of nest envelope; e, groove where strips meet. E. The colony inhabited a cavity in a rotting stump: latter mea- sured six inches in diameter and 14 inches in height. When removed from its site on August 31, 1954, the nest comprised seven oval combs whose maximum diameters ranged from 4.5 down to 3.5 inches. Taken through September 1-5 were 472 workers, 13 young males and nine queens. The queen mother had disappeared. Pupae in the number of 147 from cocoons in comb 5 were identified as 126 males and 21 queens. The cocoons of the seven combs were of two distinct sizes, the smaller produced workers, the larger males or queens. Accordingly, combs 1, 2, 3 and about % the area of comb 4, comprised worker cells; whereas % of comb 4 and all of 5, 6 and 7 consisted of male and queen PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 335 Fig. 2. Mesoscutellum, metanotum and propodeum of Vespula maculifrons (Buysson): a, mesoscutellum; b, metanotum; c, c, pair of subquadrate yellow patches on propodeum. larvae or pupae. Combs 5 and 6 had produced some adults but most of the cells still housed pupae, with a few larvae in their peripheral zones. Comb 7 presented a solid block of 112 new white pupal cocoons of uniformly large size, and a marginal zone of larger larvae, but no eggs. The absence of eggs and the existence of some immature larvae indicate the mother queen had ceased functioning only a few days before September 5. Almost the entire potential of workers in combs 1-4 had been realized, whereas the component of the reproductive caste would have continued to increase through much of September. Total frosts in this interlake area may be delayed until October. F. The nest was built between two rafters on the north eave of a neighbor's pumphouse. Being broad and short, it protruded but little below the sloping overhang, and the entrance remained invisible because located close to a rafter. 336 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 On September 3, 1967, three days before the nest was taken down, the owner of the premises attempted to eliminate the wasps by directing a forcible stream of water at it, but only tore the fragile envelope off, hence left the combs exposed. On the night of Sep- tember 5, the combs had been covered again but thinly. Moreover, numerous workers continued at work on the envelope, illumined dimly by a distant porch light. Hence, my decision to delay taking the colony on the sunny morning of September 6. In two hours, use of a light net reduced the worker force to safe numbers. The fragile combs broke into pieces when they fell 9 feet to the ground. Four combs had been constructed. The total of 321 adult vulgaris captured comprised 247 workers, the living queen mother, 19 daughter queens and 54 young males. A few adults escaped when the nest crashed to the ground. Additional pupal reproductives remained in the combs. This vulgaris nest also contained 8 workers of V. maculifrons. Nesting material of vulgaris. In his review of materials employed by Vespine wasps in building their nests, Duncan (1939, p. 121) states that V. vulgaris “seems definitely to prefer thoroughly decayed wood in the brown rotten stage.” All my nests of this species confirm the Duncan conclusion, since all consisted of such fragile, friable or crumbly wood. This fragility necessitates extreme care in order to remove the nests undamaged from their niches. It may explain also why the strips that form the nest envelopes extend obliquely to vertically over the surface (fig. 1) rather than circumferentially as on the nests of other Eaglenest Vespulas. LIFE OF Vespula maculifrons Certain colonies of Vespula described in this article afford indications that V. maculifrons on occasions maintains three modes of life, as follows: (1) it lives as an independent free-breeding species, (2) as an inquiline of V. vulgaris and (3) interbreeds with vulgaris. 1. I have one colony (G) that consisted entirely of V. maculifrons. It nested in the attic of a cabin where it was both invisible and in- accessible to me. The queen was discovered on June 17, 1959 after the nest had been founded, and the first worker appeared on June 27. Juvenile workers loafed at the emergence hole in the cabin wall through much of the summer. From September 22-27, I netted workers as they arrived and departed from the wall. Adding several taken previously, my collection totalled 72. Although not seen after July 1, the queen performed her reproductive function most of the summer, as may be judged by the number and duration of the worker progeny. Juvenile males and queens were not seen. The 72 workers are remarkably uniform in color pattern, i.e. the black antennal scape, yellow outer orbits and the pair of large squarish PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Bor propodeal patches. On the other hand, the markings of the clypeus and the shapes of the yellow hind margin of the pronotum lean toward vulgaris. However, this form appears to be only one of several now included under the name maculifrons (Miller, 1961, p. 10), hence the final concept of the species remains to be determined. 2. Vulgaris colonies D and F were unique in housing small numbers of worker maculifrons. Colony D, taken through August 17-25, 1955, contained seven, and colony F,, netted on September 6, 1967, harbored eight worker maculifrons. These workers, like the 72 obtained from my free-living colony G above, bore the pair of large yellow propodeal spots (fig. 2). Miller (1961, p. 10) found this same xanthic form of maculifrons “rare and sporadically” distributed throughout the popu- lation of vulgaris where it made up less than 5% of each colony. Certain observed facts indicate the procedure by which this bilateral relation takes place. First, the two related species V. maculifrons and vulgaris coexist in the Eaglenest Lakes area. Here they also commonly develop independently. Since neither a mother queen, nor juvenile queens or males occurred in colonies D and F, this minority component of worker maculifrons can have originated only in a free-breeding colony. The workers in such colonies of maculifrons emerge from their cocoons approximately between late June and early September. It must be supposed that some workers, incident to foraging for food and paper pulp, discover a nest of vulgaris. By devious devices, a few become established in the nest guarded by somewhat hostile worker vulgaris. Establishment of this minority is most likely to take place during August when the worker caste is presumably at its numerical peak. This is suggested by the fact that colonies D and F contained maculifrons late in August and early September. Yet it is possible that this relation may be established as early as July. Considering that the mortality rate of workers is high, the composi- tion of the maculifrons element in vulgaris nests may change sig- nificantly during the season. This presumes that the maculifrons of vulgaris colonies are not relieved of acquiring their own food in the field. And while so engaged some may return to their native home. Conclusion: the small minorities of maculifrons found in colonies of vulgaris perform in the role of inquilines, temporary and occasional. 3. Colony H gives evidence of interbreeding between V. vulgaris and maculifrons. It nested beyond sight and access in the double wall of a cabin. The annoying workers suffered destruction from swatters and chemicals. It was estimated that 300 adults had been destroyed by August 15, 1961. I selected a sample of 13 workers in good condition for study, and recovered the battered queen vulgaris. That the male parent was a maculifrons is shown by the typical yellow outer orbits and the pair of large subquadrate yellow propodeal patches displayed by all the 13 workers. Other characters of the 338 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 workers varied widely from the maculifrons I have from colony G to the vulgaris from colonies A to F. The intergradation pertained par- ticularly to the following features: the clypeal color pattern ranged from immaculate to spotted to a short median stripe to fully anchor- shaped. Likewise the yellow hind margin of the pronotum varied from narrow-parallel as in vulgaris to very wide as in extreme maculifrons. A much larger sample from this colony could scarcely have shown more diversity in these characters than the 13 workers taken. SUMMARY Life of vulgaris. The overwintered queen founds her nest late in May to early June at Eaglenest Lakes, Minn. near Ely. The first workers emerge from their cocoons late in June. By mid-August, the number of workers reaches several hundreds, and continues to increase until about September 1, when colony D contained 524. Thereafter the worker force diminishes, since the queen mother begins laying male and queen eggs about August 1. Juvenile males appear somewhat before the young queens, and in larger numbers. A few males but no queens had appeared on August 25, and a sharp increase had re- sulted in colony E on September 5. Colony F, taken on September 6 contained 54 males and 19 daughter queens. Emergence of this caste may continue into late September. The queen mother of colony B had disappeared before July 1, while that of colony F remained active on September 6. The latter had a productive lifetime of more than three months. Since the above six colonies developed in six different years, the varied weather conditions obviously affected the dates of the events described. Life of maculifrons. Evidence is presented that shows maculifrons may perform as (1) a free-breeding species; (2) a small number of workers become established as inquilines in nests of vulgaris, and (3) maculifrons and vulgaris sometimes hybridize. In the latter in- stance the mother queen was a vulgaris. REFERENCES Balduf, W. V. 1954. Observations on the white-faced wasp, Dolichovespula maculata (Linn.) (Vespidae, Hym.). Ann. Ent. Soc. Amer. 47:445—458. Dunean, C. D. 1939. A contribution to the biology of North American Vespine wasps. Stanford Univ. Pubs., Univ. Ser., Biol. Sciences, 8( 1): 1-272. Miller, C. D. F. 1961. Taxonomy and distribution of nearctic Vespula. Can. Ent. 93( Suppl. 22) :1-52. PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 339 SNELLENIUS IN THE NEOTROPICAL REGION (HYMENOPTERA: BRACONIDAE: MICROGASTERINI ) R. D. SHENEFELT, Department of Entomology, University of Wisconsin, Madison, Wisconsin 53706 Snellenius was originally described by Westwood from specimens collected in New Guinea and has since been recorded from Borneo, China and Japan. In 1933 Brues described a species from the Oligocene Baltic amber which he included in the genus but the thoracic struc- ture is such that the insect is probably incorrectly placed. If this fossil is omitted, Snellenius currently includes five species, all of which appear to be uncommon. It was therefore quite surprising to find in material from South America several undescribed species which were relatively well represented by specimens. The generic placement of these insects proved to be difficult as the material would not fit into any of the tribes recognized by Telenga in 1955. The antennae and wing venation are definitely microgasterine in character, but the very deep notaulices and other features are aberrant. At the United States National Museum no identified comparable specimens were located, but Dr. Marsh remembered seeing a species of similar insects and kindly arranged to have the 24 specimens involved loaned to me for study. Examination of the Neotropical material has disclosed that the characterization of the genus must be altered somewhat. The features which seem to help follow: Eyes hairy; occiput immargined; antennal segments flattened to nearly terete; antennae 18-segmented; notaulices deeply impressed, crenulate; median lobe of mesoscutum distinctly higher than lateral lobes, with its side margins raised and a low longitudinal carina along the middle; the notaulices separated behind only by a high keel-like carina; prescutellar fovea broad and deep; scutellum margined along the sides by a distinct rounded carina; antero-laterad of scutellum and laterad of prescutellar fovea a pair of impressed ear-like lobes; band bordering scutellum broadly interrupted on posterior portion; sternauli deep, coarsely crenu- late; prepectal carina either complete and distinct from in front of sternauli across the venter, or incomplete and reduced to a mere ridge across the anterior end of the median sternal groove only; median sternal groove broad, trough-like and crenulated; propodeum with a prominent median carina, the remaining sculpture varying from coarsely foveolate with very high carinae to nearly reticulate-rugose, the dorsal and posterior faces either distinctly separated by an angle or indistinctly limited and the profile appearing rounded. Forewing with three cubital cells, the second (areolet) small (but large for Microgasterinae) and closed by second intercubitus; second intercubitus mostly transparent, joining second abscissa of radius; third abscissa of radius indicated by pigmented line which may be curved so it appears concave behind; nervulus from nearly interstitial (see West- wood’s drawing) to postfurcal to the point where the first abscissa of discoideus is at least two-thirds as long as the second. Cubitellan cell closed; nervellus not 340 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 sinuate, curved inwardly; vannal lobe differentiated. Hind coxa small, approxi- mately as long as first abdominal tergite; spurs of hind tibia very small, their length less than the apical width of the robust hind tibia. Abdomen sessile; first abdominal tergite with a narrow median sclerotized plate with nearly parallel sides and with broad membranous lateral portions in which the spiracles are located (these side portions may be folded downward in mounted specimens so the abdomen appears to be petiolate ). The Neotropical species of Snellenius run to Microplitis in the keys by Muesebeck 1922, Fahringer 1936, Morley 1936, and De Saeger 1944. In Nixon’s 1965 key they may not fit the first two characters given in couplet 11. The major features assisting in recognition of the genus are the following: the unusual thoracic structure (including the deep notauli- ces, margined scutellum, the median sternal groove, the large, crenu- lated sternauli, and the foveolate area beneath the forewing); the narrow parallel-sided median plate of tergite 1; the relatively large second cubital cell; the short tibial spurs and robust hind tibiae. The head may or may not be relatively small; the propodeum may or may not be sharply angled as seen in profile from the side; the antennal segments may be strongly flattened to nearly terete. The following key, which is practically that of Nixon in the first five couplets, should aid in recognition of the species included in the genus. 1. Disc of scutellum reduced to a small, blunt, erect cone; flagellar segments Stronely:tlattened =29% ps. oN we ee oe a ane | vollenhovii Westwood Disc of scutellum not cone-like but flattened and with strong raised lateral margin _______ poets tf A A Se ee ee 2 2. Notaulices shallow, flagellar segments weakly ened _ radiealis Wilkinson Notaulices very deep, crenulated, separated behind by ms a narrow, keel- like carina; middle lobe of mesoscutum decidely raised above lateral lobes ER 2 PN eR es ee ee A 3 3. Propodeum with carinae high and so coarsely reticulate that between lateral and median carinae not more than 3 or 4 hollows occur along a transverse |i e90 e n eE ROMDrE Ce re 4 Propodeum not so coarsely reticulate, the carinae lower and the number of hollows between the median carina and the lateral carina more numerous; median plate of first tergite a little more than twice as long as its apical width; notaulices narrowing anteriorly and lateral edges of median lobe of mesoscutum not upturned _.- pee ee theretrae Watanabe 4. Temples rugose; hind femur with well developed keel on apical two-fifths OM pIIME! SIC) oe 5 Temples smooth or scabrous-punctate; hind femur without keel 6 5. Head deeply emarginate between the almost angular corners of the temples; forewing nearly hyaline igo _ philippinensis (Ashmead ) Head weakly emarginate, temples without prominent angles; forewing deeply ITIMIISCALCG ee esee eit phe tod et oe en gelleus Nixon 6. Prepectal carina complete, extending from in front of sternauli across venter; PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 341 vertex and temple scabrous-punctate; plate of tergite 1 a little more than twice as long as broad at apex = U Prepectal carina incomplete, extending across median somal groove only; vertex and temple mostly smooth; plate of tergite 1 at least 3 times as long as broad at apex ~ Sub be pO SO Mk teats Oe = 8 7. Coloration involving red, black, and yellow; Pediclln fea as long as first abscissa of basella : _ tricolor, n. sp. Coloration involving only red ah Black! patna, oe three- fourths as long as first abscissa of basella is __ bicolor, n. sp. 8. General body color piceous; penultinate segment of maxillary palpus less than halfvas lone”asithe ultimate =. 2 __..... atratus, n. sp. General body color testaceus; penultimate ccomient Ge maxillary palpus at least half as long as the ultimate — = vee’ peruensis, n. sp. Snellenius tricolor, n. sp. Holotype: ¢°, bearing the following data on the label “Argentina: Horco Molle, Tuc. Apr. 3-10, 1966 L. A. Stange.” Deposited in the U. S. National Museum. Length: Head and thorax 1.7 mm, abdomen 1.3 mm, antenna 3.5 mm, forewing 3 mm. Black or piceous: Head, sides of thorax below, venter, scutellum and portions of propodeum. Antennae deep brown, mesoscutum (and other portions of thorax which are not dark) ferrugineous to light brown. Plate of abdominal tergite 1 plus hypopygium and tergites beyond 2, except on the sides, brown. Ovipositor sheaths brown. Sides of abdomen, second and third tergites (excepting a slightly browned area in the anterior central portion of tergite three), lateral thirds of tergite 4, and lateral fourths of tergite 5 contrasting yellow to yellow-testaceous. Maxillae and labium, including palpi, yellow. Legs primarily fuscous but distal portions of anterior femora, basal parts of tibiae and tarsi yellowish, and a band of proximal portions of mid and hind tibiae testaceous. Temple: 0.7 x as wide as eye in lateral view, eyes from in front one-fourth further apart above than at narrowest part of face, slightly emarginate opposite antennal bases. Reticulation of propodeum moderately coarse, with about 3-4 hollows along a transverse line from median carina to lateral carina. Plate of tergite 1 twice as long as its apical width, rugose on posterior third and with apex curved; remaining abdominal segments polished. Allotype: é, data same as on holotype. Differs from the holotype in being less dark. The scape slightly lighter in color above, the legs, including all coxae, mostly yellowish. The apical two-thirds of hind tibiae and posterior tarsi brown. Labial palpi small, their length about equal to the width of the labrum. Plate of tergite one yellowish with darkened edges. Deposited in U.S. National Museum. Paratypes: 22 4 8 from the same locality. Three collected in the same week as the holotype and the other 19 taken Mar. 7-13, 1966 by Mr. Stange. In collections of Instituto Miguel Lillo at Tucuman, U.S. National Museum and the author. Among the paratypes the color varies considerably, but the head is 342 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 always black, the mesoscutum has a distinct reddish cast and the anterior portion of the abdomen is yellowish. The coloration of the legs varies from nearly all yellow to nearly all brown. In most of the specimens the scape is ferrugineous above, contrasting with the brown in the remainder of the antenna. Snellenius bicolor, n. sp. Very similar to tricolor but differing in that the areas showing as yellowish in tricolor are ferrugineous (and unicolorous with the meso- scutum ); the temple is as broad or broader than the eye; the labial palpus in the ¢ is distinctly longer than the labrum is wide. Holotype: 9°, bearing the labels “Avispas, Peru IX-1962.” “Collec- tion R. D. Shenefelt.” Taken by L. Pefia in Malaise trap. In collection of author. Length: Head and thorax 1.8 mm, abdomen 1.3 mm, antenna 3.4 mm, fore- wing 3.1 mm. Head black. Eyes reddish. Thorax and legs ferrugineous (Maerz & Paul Plate 5 D-12). Antennae, all pretarsi, posterior tibiae and apical three-fourths of posterior femora brown. Abdomen yellow ochre (Maerz & Paul Plate 11 L-7) with the plate of tergite 1 darker and the membranous area paler than the re- mainder. Palpi not contrasting in color with anterior leg. Temple and eye subequal in width when viewed from the side. Eyes above 20 units apart, separated by 17 units at the narrowest part of the face, slightly emarginate opposite antennal bases. Reticulation of propodeum coarser than in tricolor, with 2-3 hollows along a transverse line from median carina to lateral carina. Plate of tergite 1 twice as long as wide at apex, rugose in apical half. Allotype: é, bearing the labels “Quincemil, Peru 10-15 XI-1962.” “Collection R. D. Shenefelt.”. Taken by L: Pefia in Malaise Trap. In collection of author. Agrees with the holotype in essential features. Base of scutellum and posterior edge of postnotum browned. Paratypes: 6 & @: two with same data as allotype; four from Quincemil, Peru IX-62. Two in U.S. National Museum and remainder in author's collection. Snellenius atratus, n. sp. Holotype: ¢, with labels “Avispas, Peru 20-30 IX-1962.” “Collec- tion of R. D. Shenefelt” on the pin. Collected by L. Pefia in Malaise Trap. In collection of author. Length: Head and thorax 1.6 mm, abdomen 1.3 mm, antenna 3.2 mm, fore- wing 3 mm. Black or very dark brown except as follows: antennae brown; palpi dark tan; middle coxae, all trochanters, proximal and distal areas on anterior femora, fore and middle tarsal segments pale brown (like the palpi). Wings uniformly in- fuscated. Temple smooth, %4 broader than eye; posterior margin of eye broadly and shallowly concave when eye is viewed from side; maxillary palpus longer than PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 343 height of head, space between eyes at narrowest point of face two-thirds the distance between eyes above. Labial palpi longer than width of labrum. Penultinate segment of maxillary palpi 0.4 times length of ultimate. Areolet of forewing unusually long, narrow, nearly triangular (the second abscissa of radius and second intercubitus forming practically a straight line with first abscissa of radius and the cell about twice as long as high). Third abscissa of radius indistinct. Mediella 13 units long, first abscissa of basella 22 and curved near base. Posterior side of cubitellan cell more than twice as long as the cell is wide at apex. Median plate of abdominal tergite one 314 times as long as wide at apex, with sharp carinae along sides on basal two-thirds, the carinae turning inwardly onto the plate near their ends and causing it to appear contracted in this area; rugose on apical third, excepting the smooth raised area at the very end. Propodeum strongly areolated with about three hollows along a transverse line between median and lateral carinae; not sharply angulate in profile. Spurs of hind tibiae ™%4 as long as hind basitarsis. Allotype: ¢, same data as associated with holotype. In collection of author. Agrees with the holotype except that second abscissa of radius and second intercubitus do not form straight line with first abscissa of radius, the areolet appearing four sided as usual. Median plate of tergite 1 nearly smooth on apical half. Mid-coxae brown on outer half and all trochanters brown. Eye relatively larger than in 9, temple only about 4o wider than eye. Paratypes: 4 2 2: 3 Avispas, Peru 1-15 X-1962; 1 Avispas, Peru IX-1962. One in collection of U.S. National Museum. Three in author's collection. 17 ¢ 4: 2 Avispas, Peru 20-30 IX-1962; 6 Avispas, Peru 1-15 X-1962; 3 Avispas, Peru IX-1962; 3 Quincemil, Peru IX-1962; 3 Quincemil, Peru 20-30 X-1962. Two in collection of U.S. National Museum. Remainder in collection of author. In the paratypes the coloration is very constant. The shape of the areolet varies as indicated under the notes regarding the allotype. The amount of roughness on the apical half of median plate of tergite one is variable. Snellenius peruensis, n. sp. Structurally very similar to atratus but the middle lobe of mesonotum more concave with the lateral margins raised higher and tending towards tuberculate on each side at a little behind the middle. Penulti- mate segment of maxillary palpi at least half as long as the ultimate. The median plate of tergite 1 with the lateral carinae extending nearly to the apex and the plate concave for most of its distance and usually with a central narrow to carinate raised area along the posterior portion just before the smooth terminal swelling. Spurs of hind tibiae relatively longer. Holotype: ¢, with the following labels on the pin “Avispas, Peru 344 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 IX-1962” “Collection of R. D. Shenefelt.”. Taken by L. Pefa in Malaise trap. In collection of author. Length: Head and thorax 1.8 mm, abdomen 1.4 mm, antenna 4.0 mm, fore- wing 3.8 mm. Fuscous, with the antenna beyond the scape, the distal halves of the hind tibiae and the posterior tarsi darker brown. Wings uniformly brown. Temple smooth, narrower than eye (eye 22, temple 18) receding more strongly than in atratus; space between eyes above 18, at narrowest part of face 15. Flagellar segments nearly terete. Penultimate segment of maxillary palpi 9 units long, ultima 18. Areolet 0.7 as high as long along cubitus (measurements made inside of veins), four-sided, the second abscissa of radius forming a slight angle with first abscissa and a definite angle with second intercubitus. Mediella 14 units long, first abscissa of basella 25 and not so strongly bent near base as in atratus. Wing veins broader (heavier) than in atratus. Median plate of tergite 6 wide at apex, 20 long, with side margins raised, mostly rugulose, with a high central carina located in the middle in front of a terminal smooth raised boss. Propodeum coarsely and strongly areolated with posterior and dorsal faces evident but not sharply angled in profile. Spurs of hind tibia 0.3 as long as posterior basitarsus. Allotype: 4, with data labels same as those on holotype. In collec- tion of author. Agrees well with holotype. The flagellar segments approximately oval in cross section, i.e., two times as wide as deep. Paratypes: 5 2 2°: three from Avispas, Peru IX-1962; 1 from Avispas 1-15 X-1962; and one from Quincemil, Peru 10-15 XI-1962. One in collection of U.S. National Museum. Remainder in collection of author. 13 4 ¢: 5 from Avispas, Peru IX-1962, 2 from Avispas 20-30 XI-1962, 6 from Quincemil, Peru 10-15 XI-1962. Two in U.S. National Museum. Remainder in collection of author. In two of the paratypes the body is much lighter brown than in the remainder but the wings and antennae are as dark as in the other specimens. REFERENCES Ashmead, W. H. 1904. A list of the Hymenoptera of the Philippine Islands, with descriptions of new species. Jour. N.Y. Ent. Soc. 12:1—-22. Brues, C. T. 1933. The parasitic Hymenoptera of the Baltic amber. Bernstein- Forsch. 3:95. De Saeger, H. 1944. Exploration du Parc National Albert. Mission G.F. de Witte (1933-1935). Microgasterinae, Fasc. 47:1-342. (Key pp. 23-24.) Fahringer, J. (1935) 1936. Opuscula Braconologica 4. Palaearktische Region. Lieferung 1—-3:1—276 Microgasterinae. (Key p. 11-15.) Morley, C. 1936. Notes on Braconidae XV.—Microgasterinae. Entomologist 69: 39-42, et seq. (Key p. 39.) Muesebeck, C. F. W. 1922. A revision of the North American ichneumon flies belonging to the subfamilies Neoneurinae and Microgasterinae. Proc. U.S. Nat. Mus. 61, Art. 15 (2436):1-76. (Key p. 8-9.) PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 345 Nixon, G. E. J. 1965. A reclassification of the tribe Microgasterini (Hymen- optera: Braconidae). Bull. Br. Mus. Natur. Hist., Ent. Suppl. 2:1-275. (Key to genera p. 11-17; to spp. of Snellenius pp. 270-271. ) Westwood, J. O. 1882. Descriptions of new or imperfectly known species of Ichneumones adsciti. Tijdschr. Ent. 25:17—48. ON THE TRUE IDENTITIES OF TUOBA AND NESOGEOPHILUS (CHILOPODA: GEOPHILOMORPHA: GEOPHILIDAE ) In his 1920 treatment of the Chilopoda of the Australian Region, Chamberlin (Bull. Mus. Comp. Zool. 64:35) described a curious new geophilomorph from the Solomons, referring it to the Gonibregmatidae, which is for many an emunctory catch-all of arcane genera, and called it Tuoba curticeps, new genus and species. And there it has reposed, unevoked and generically unidentifiable, for Chamberlin’s delineation, published without figures, entails several crucial errors that have led subsequent workers down error’s garden path. So when in 1924 Verhoeff (Nat. Hist. Juan Fernandex 3:413) proposed within Geophilus a new subgenus, Nesogeophilus, from Juan Fernandez, he forgivably failed to identify its two species as proper congeners of the unknowable Tuoba curticeps. Having studied all of the pertinent types, I can assert confidently that since laticollis (Attems), the type-species of Nesogeophilus (by Attems’ subsequent designation in 1929) is congeneric with curticeps Chamberlin, type-species of Tuoba (by original designation), it follows that Nesogeophilus 1924 is a junior subjective synonym of Tuoba 1920. I imagine Chamberlin assigned Tuoba to the Gonibregmatidae because, as _ is clear from his description, he failed to discern the actual condition of the coxopleural glands, which are not manifest as freely-opening surface pores as he suggested. Rather each coxopleuron has a single large, ventral, concealed, glandular crypt that is heterogeneous, multiglandular, and multicanaliculate. A second and most distinctive feature of the genus, that no one has yet detected, is the pretarsal anterior parunguis, which is both strictly spiniform and greatly elongate, being as long as the claw proper and much longer than the minute posterior parunguis. This extraordinary condition surely seems associated with the genus’ distinctively littoral preferences; it probably serves as a special hold-fast adaptation. Tuoba belongs where Attems and Verhoeff stationed Nesogeophilus, in Geophilidae. The genus, which is predominantly littoral and world-wide, is represented in the Australian Region by the following: curticeps Chamberlin, 1920, Solomons hartmeyeri (Attems, 1911) (= laticeps q.v., part, and sydneyensis q.v., part. New Synonymies and Combinations, Australia ) laticeps (Pocock, 1891), New Combination, Australia sydneyensis (Pocock, 1891), New Combination, Australia xylophagus (Attems, 1903) (= laticeps q.v., New Synonymy and Combina- tion, New Zealand ). R. E. Crasity, Jr., Smithsonian Institution, U. S. National Museum, Washing- ton, D. C. 20560. 346 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 A NOTE ON THE IDENTITY OF HYLEMYA VARIATA (FALLEN) AND H. VARIABILIS STEIN (DieTERA: ANTHOMYIIDAE ) H. C. Hucxett, Riverhead, New York 11901 The subgenus Hylemya of authors has been recorded in North America as containing two species, Hylemya alcathoe (Walker) and H. variata (Fallén). Recently Ackland (1967, p. 120) has observed that the species Hylemya variabilis Stein (1916, p. 155), as known to him, possessed diagnostic characters comparable to those illustrated by Huckett (1924, figs. 17, 68) for variata in North America. His ac- ceptance of Stein’s species was conditional, provided Stein was correct in regarding variabilis as apart from variata. Through the courtesy of Dr. Inge Persson of the Naturhistoriska Museum near Stockholm I have had the privilege and opportunity of examining the type-specimens of Musca variata Fallén. All such had a small red ticket with printed numbers 100 to 105 and 68 in ink respectively, and another series with numbers 493 to 496 and 67 in ink. In addition all specimens possessed a green label, on which was printed Riksmuseum, Stockholm. Of the ten specimens present I regard the four males and four females as belonging to variata, one female (No. 102) to Hydrophoria ambigua (Fallén), and one female (No. 495) to Hydrophoria annulata (Pandellé). I have designated the male No. 494, mounted on a coarse grayish pin, as the lectotype of Musca variata Fallén. The drawings of male copulatory appendages and genitalia of variata have been made from a male numbered 101, having a similar coarse pin. It should be noted that in this specimen one of the outer pair of parameres of the genitalia, as shown, has three filaments (fig. 5), the other having the normal two. The chief distinctions between the species variabilis and variata may be found in the structure of male copulatory appendages. In variata the cercal plate or cerci is longer than wide and gonostyli slender, lengthy and curved caudad at apex when viewed laterad (figs. 4,5) whereas in variabilis the cerci are as wide as long, and the gonostyli short, notched proximad and curved cephalad when viewed laterad (figs. 1, 2). A single comparison of the ovipositor failed to indicate any notable differences between the two species. I have yet to recognize variata (Fallén) as occurring in North America, examination having been made of males from Alaska, Yukon Territory, British Columbia, Manitoba, Ontario, Quebec, New York, Iowa and California. The character of the copulatory appendages exhibited by these specimens all served to confirm Stein’s concept of variabilis as apart from variata. It would appear that previous records of H. variata from North America are actually based upon misidentified PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 347 Figs. 1-6, copulatory appendages and genitalia, 6 ¢: 1-3, Hylemya variabilis Stein; 4—6, H. variata ( Fallén). specimens of H. variabilis, which is here recorded from North America for the first time. REFERENCES Ackland, D. M. 1967. Diptera from Nepal, Anthomyiidae. Bull. Brit. Mus. (Natur. Hist.), Ent. 20(4):107-139, 83 figs. Huckett, H. C. 1924. A systematic study of the Anthomyiinae of New York, with especial reference to the male and female genitalia. Mem. 77 N.Y. (Cornell) Agr. Exp. Sta., 1-91, 192 figs. Stein, P. 1916. Die Anthomyiden Europas. Arch. f. Naturgesch. (1915) 81A (10) :1-224. 348 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 THE PANAMANIAN ATTA SPECIES (HYMENOPTERA: FORMICIDAE ) Neat A. WEBER, Department of Biology, Swarthmore College, Swarthmore, Pennsylvania 19081 Three species of Atta are known from the Republic of Panama, including the Panama Canal Zone area (Weber, 1956). The present note illustrates these for the first time. One has become of particular interest in view of the biochemical studies of Martin et al. (1967 and manuscript ) and all have been used by me in various published and unpublished biological studies. All have been maintained in my laboratory and 1966 colonies of two are currently thriving. Of the three, Atta sexdens (L.) is widespread in South America and extends into Costa Rica. The common Atta cephalotes (L.) of South America is represented in Panama by the subspecies isthmicola Weber, and perhaps undescribed subspecies are found in Costa Rica and elsewhere in Central America. The distribution of sexdens and cephalotes in South America is figured in Weber, 1966. A narrow, isolated coastal strip in the state of Bahia, Brasil, that was inadvertently omitted from the cephalotes map was kindly called to my attention by Dr. Pedrito Silva. The distribution of the third, Atta colombica tonsipes Santschi, is Panamanian only, so far as known. The identifica- tion is based on the Santschi type material, which I have studied (Weber, 1958). It comes from two Panamanian localities where this ant still occurs. The three may be distinguished from one another as follows: 1. Head of soldier relatively smooth and shiny on the occiput; workers smooth 2. No pre-occipital spine or tubercle in soldier or worker; mostly a species of forest’... colombia tonsipesssausea Pre-occipital spine or tubercle in soldier and worker; grasslands or grass- land-forest margin ecotone ae sexdens (L.) Of the three, sexdens has the smallest male but all have large females. The weights in life of representative soldiers were 67-103 milligrams (mature isthmicola colony), 27-35 mg (tonsipes of 1% year colony ) and 23-64 mg (young sexdens colony). It appears that it takes more than two or three years to produce the largest soldier in tonsipes and a shorter period in the other two. Small soldiers are produced in all three in the second half of the first year of colony life. In young colonies the species may be easily distinguished by the smooth and shiny workers in isthmicola contrasted with the matte workers of the other two and the pre-occipital tubercle in sexdens being absent in tonsipes. PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 349 \ 5 Ps KX, . ne \ cae e) eri \ i = e @-=s an ay. ise ; \ 3 Be a q iy [Fes I Cy. : 2 } A Lf Wet th at = UANVT - es Ss S| ¢ - Pd vA Pe. : TWN oy 7 th A NAL f as Y \V HT ( i Ze {@ ) eos - 3 OS ~ : te v i 4 & - ) Ta foN OY ry (“ oN 7 : x pr cs 6 vy e Fig. 1, Atta sexdens (L.) worker, Panama. Figs. 2, 5-8, A. colombica tonsipes Santschi: 2, thorax length 2.70 mm, width of head 2.18 mm; 5, worker head width 2.25 mm, back of eyes, 1.80 mm from clypeal margin to mid-occipital impression; 6, worker head width 3.40 mm and length as above 2.40 mm; 7, worker head 1.60 mm and 1.50 mm; 8, alate female from above. Figs. 3 and 4, A. cephalotes isthmicola Weber: 3, cotype worker, thorax 2.70 mm, width of head 2.18 mm, ant size identical to tonsipes above, Canal Zone; 4, soldier, dorsal view, Canal Zone. ACKNOWLEDGMENTS David Klingener kindly determined the distribution of tonsipes in the Canal Zone for me in 1957 and Suzanne Tubby (now Dr. S. T. Batra) made the drawings. Aided by NSF Grant GB5346. REFERENCES Martin, M. M., G. A. Carls, R. F. N. Hutchins, J. G. MacConnell, J. S. Martin and O. D. Steiner. 1967. Observations on Atta colombica tonsipes (Hymen- optera: Formicidae). Ann. Ent. Soc. Amer. 60:1329-1330. ——., R. M. Carman and J. G. MacConnell. 1968. Nutrients derived from the fungus cultured by the fungus-growing ant, Atta colombica tonsipes (Hymen- optera: Formicidae). Unpublished manuscript. 350 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Weber, N. A. 1956. Symbiosis between fungus-growing ants and their fungus. 1955 Yearbook Amer. Phil. Soc., pp. 153-157. 1958. Nomenclatural notes on Proatta and Atta (Hym.: Formicidae). Ent. News 69:7-13. 1966. Fungus-growing ants. Science 153:587-604. STUDIES ON CALIFORNIA ANTS. 4. TWO SPECIES OF CAMPONOTUS (HYMENOPTERA: FORMICIDAE) Roy R. SNELiLING, Los Angeles County Museum of Natural History, Los Angeles, California 90007 Prior to the work of Creighton (1950) the genus Camponotus Mayr had assigned to it approximately 60 names for various components of the Nearctic fauna. Creighton attacked the problems posed by this superabundance of names and reduced them to 48. Since then little has been done: three new species have been described (Creighton, 1952, 1965; Smith, 1953), two have been redescribed (Creighton, 1965; Creighton and Snelling, 1966), one has been excluded from the Nearctic fauna (Creighton, 1952) and one subspecies has been trans- ferred from one species to another and back again (Brown, 1950; Gregg, 1963). While studying the ants of California and Baja Cali- fornia, Mexico, I became convinced that still further changes are in order. Two of the necessary changes affecting the species of Cali- fornia are proposed here. A large part of the material on which this study is based is in the collections of the Los Angeles County Museum of Natural History. An important collection from Baja California has been made available through the courtesy of E. L. Sleeper and E. M. Fisher, Long Beach State College, Long Beach, California. Other material, including important type specimens, was studied at the United States National Museum through the kindness of D. R. Smith. In several conversations I have had the benefit of the extensive experience of W. S. Creighton and M. R. Smith has made available for reference an unfinished manu- script revision of the subgenus Myrmentoma Forel. To each of these gentlemen my very sincere thanks for their cooperation. The figures for this paper were prepared by Ruth A. DeNicola to whom I remain grateful. Camponotus (Tanaemyrmex) festinatus (Buckley), new status Formica festinata Buckley, 1866, Proc. Ent. Soc. Phila. 6:164. % 9. Camponotus (Camponotus) fumidus pubicornis Emery, 1894, Zool. Jahrb., Abt. f. System. 7:668, 670. 3%. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Soll! Camponotus fragilis Pergande, 1894, Proc. Calif. Acad. Sci. 4:26. &. Camponotus fumidus var. festinatus: Wheeler, 1902, Trans. Texas Acad. Sci. 4:22. %. Wheeler, 1910. Ann. N. Y. Acad. Sci. 20:312-314. Camponotus fumidus var. fragilis: Emery, 1895, Zool. Jahrb., Abt. f. System. 8:336. %. Wheeler, 1910, Ann. N. Y. Acad. Sci. 20: 315. 3. Camponotus fumidus var. spurcus Wheeler, 1910, Ann. N. Y. Acad. Sci. 20:315. SQ. Camponotus fumidus subsp. festinatus: Creighton, 1950, Bull. Mus. Comp. Zool. 104:376. Although this ant has traditionally been considered a form of C. fumidus Roger I have departed from this treatment. Roger (1863) described his species from a major worker from an unspecified locality in Venezuela. While the original description is not completely ade- quate by current standards, one significant characteristic was de- scribed; Roger stated quite clearly that the scapes and tibiae of C. fumidus were without erect hairs (“. . . fehlt am Scapus und an den Schienen.”). Since a number of so-called subspecies have been assigned to C. fumidus which possess abundant erect hairs on the scapes and tibiae I have examined them and find that they differ specifically from one another and therefore, presumably, from C. fumidus as well. I have not seen true C. fumidus; the few specimens available to me under this name do not agree with the original description and I see no reason to consider them the same as Roger’s ant. All material which I have seen from Mexico and the United States has been assigned to C. festinatus. While there is a considerable amount of variation in these samples, they consistently possess erect hairs on the scapes and tibiae. Of the several forms currently assigned to C. fumidus, C. f. pubi- cornis, C. fragilis and C. f. spurca may be disregarded as outright synonyms of C. festinatus, as shown by Creighton (1950). Two West Indian forms, C. f. vittata Forel and C. f. lucayana Wheeler, are de- serving of consideration, since they bear somewhat on the status of our mainland form. Of C. f. vittata I have seen a few workers and majors and a single female from Brazil and a long series of workers and majors from the island of Dominica. Of C. f. lucayana I have seen several series, including all castes, from the Bimini Islands. These, together with C. festinatus, form a distinct group but each apparently should stand as a separate species. All agree in possessing erect hairs on the scapes and tibiae as well as on the cheeks. Although there is conspicuous variation in the color of these ants they offer an undeniable basic similarity. Basically the integument is pale yellow but it tends to be overlain by a distinct brownish infuscation, especially on the occipital region, the thoracic dorsum and on the gaster. In its greatest development (most material 352 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 3 3a 3b Fig. 1, Camponotus festinatus (Buckley), major worker, head, frontal view; la, major worker, lower half of head, lateral view; 1b, female, head, frontal view; lc, male, head, lateral view; 1d, female, petiolar scale, lateral view. Fig. 2, Camponotus lucayanus Wheeler, same. Fig. 3, Camponotus vittatus Forel, a, b, same; 3c, female, petiolar scale, lateral view. of C. vittatus) the yellow is almost completely obscured except for lateral spots on the gastric segments. The same pattern is present, too, in infuscated individuals of C. lucayanus and C. festinatus but not as well developed though occasional specimens show lateral spots. PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 353 All three species have the apico-median portion of the clypeus slightly prolonged, with its margin transverse. A median longitudinal carina is present, but the extent of its development varies from one species to the next. In C. vittatus (fig. 3a) the carina is high and sharp and extends the entire length of the clypeus in the workers and females. In C. lucayanus (fig. 2a) and C. festinatus (fig. la) the carina is much lower and rounded; indeed, it is hardly more than a median angulation of the clypeal disc. In these species, also, the carina extends only about two-thirds the length of the clypeus. In profile, the carina in C. lucayanus is relatively flat in the apical three-fourths and slopes abruptly in the basal one-fourth to the basal clypeal suture. By con- trast, that of C. festinatus presents a low contour which is evenly rounded; rarely is it slightly angled at the basal one-fourth. The above considerations apply to the workers, both major and minor, and to the females. Males of C. vittatus have not been avail- able, but I have seen numerous males of C. lucayanus and C. festinatus. The profile of the clypeus is dramatically different. In C. lucayanus (fig. 2c) the clypeus is either flat or with an angular transverse im- pression in the middle; the basal one-fifth falls steeply, sometimes vertically, to the basal suture. No such condition exists in C. festinatus males. In these the clypeal profile may be slightly convex, a little more rounded toward the base, or with a transverse median impression. But, in all specimens seen by me, the basal portion is evenly rounded toward the basal suture (fig. 1b). The major workers of C. lucayanus have a slightly shorter scape than do those of the other species, exceeding the occipital corners by about the length of the first funicular segment. In C. festinatus and C. vittatus the scapes extend beyond the occipital corners by a distance nearly equal to the combined lengths of the first two funicular seg- ments. The shape of the head, in full face view, differs among the three species. In C. vittatus the head of the majors is relatively longer and the convergence of the lateral margins below is more pronounced (fig. 3). In the minor workers of C. festinatus and C. lucayanus the margins of the head, below the level of the eyes, are parallel or slightly narrowed toward the mandibular insertions, while in C. vittatus the sides of the head are divergent below; i.e., the head is a little broader at the level of the mandibular insertions than at the level of the lower margins of the eyes. The shape of the head of the females differs but the differences are slight. That of C. vittatus (fig. 3b) is distinctly sinuate in full face view, the face being narrowed slightly below the level of the eyes. This, however, is based upon a single specimen and I believe it may not prove consistent, since I have seen a few females of C. festinatus with 354 SCAPE LENGTH (MM ) SCAPE LENGTH (MM) HEAD WIDTH (MM) PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Lz, 18 1g 2.0 21 2.2 2.3 24 2.5 HEAD LENGTH (MM ) ©- C. sayi - cotypes O- C.sayi-western (- C.sayi - eastern 1.8 19 2.0 21 2.2 23 24 2.5 2.6 HEAD LENGTH (MM) PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 S00 an indication of a similar narrowing. The head is longest and narrowest in C. vittatus, shortest and broadest in C. festinatus. The erect cephalic hairs differ and in this, also, C. vittatus is most distinctive. In both C. festinatus and C. lucayanus the longest hairs are shorter than the maximum diameter of the eyes, while in C. vittatus these hairs are distinctly longer. The latter species also has a greater number of erect hairs as indicated in the figures. Finally, the shape of the petiolar scale differs. In C. lwcayanus and C. festinatus the scale is variable in profile, but it is usually blunt above, with the summit rather evenly rounded (figs. ld, 2d). The scale of C. vittatus (fig. 3c) is distinctly cuneate in profile, the apex strongly narrowed and angular. There is no evidence that these forms are sympatric with one another and hence there is no opportunity for the intergrades one would expect if they were subspecies of a single polytypic species. This, of course, does not rule out the possibility that they may actually be subspecies, but I am convinced that the morphological evidence is against such an interpretation. That any of these could be subspecies of C. fumidus, which lacks the erect hairs so conspicuous on the scapes and tibiae of the other three, I seriously doubt. The southernmost of the three species, C. vittatus, is typically the most hirsute; specimens of C. festinatus from the southern parts of its range exhibit an increase in the density of erect hairs present. This seems to be precisely the opposite of what should be expected if this ant is, in fact, a subspecies of the Venezuelan C. fumidus. It seems more reasonable to accord this ant specific status until conclusive evidence to the contrary is forthcoming. Camponotus (Myrmentoma) sayi Emery Camponotus sayi Emery, 1894, Zool. Jahrb., Abt. f. System. 7:679. %. Wheeler, 1910, Ann: N. Y. Acad. Sci. 20:343. 3%. Camponotus sayi var. bicolor Pergande, 1894, Proc. Calif. Acad. Sci. 4:161. 3 Q ¢. Preoce. Camponotus fallax subsp. rasilis Wheeler, 1910, Jour. N. Y. Ent. Soc. 18:227. 394. New synonym. Camponotus sayi californica Emery, 1925, in Wytsman, Gen. Insect. 183:118. New name for C. sayi bicolor Pergande. New synonym. Camponotus (Myrmentoma) rasilis: Creighton, 1950, Bull. Mus. Comp. Zool. 104:389. Gregg, 1963. Univ. Colo. Press, Boulder, pp. 677-678. Camponotus (Myrmentoma) sayi: Creighton, 1950, Bull. Mus. Comp. Zool. 104:390. cs Fig. 4. Head length-scape length relationships in Camponotus sayi Emery major workers. Fig. 5, Cephalic index-scape length relationships in C. sayi. Fig. 6, Head length-head width relationships in populations of C. sayi major workers. 356 PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 Camponotus sayi was described by Emery from a limited series of specimens taken at Phoenix, Arizona. These ants were sent to him by Pergande who, typically, retained a portion for his own collection. Six specimens of this series (three major and three minor workers), marked as cotypes, are in the collections of the United States National Museum. These specimens, plus the cotypes of C. sayi bicolor, are the basis for the discussion which follows. Emery’s original description, while brief, is straightforward and con- tains sufficient information to permit recognition of the species. At this time the ant was compared to C. marginatus discolor (i.e., C. caryae discolor), but was noted to differ in the lack of foveiform punctures on the cheeks. Pergande described C. sayi bicolor from a long series of workers, two females and three males from Chuparosa in the Sierra Laguna and San Jose del Cabo, Baja California, Mexico. The name chosen by Pergande was preoccupied and Emery renamed it C. sayi californicus in 1925. Pergande characterized his ant as being distinctly larger than C. sayi from Arizona (of which he had part of the original series) but I do not think this is important since size is a notoriously poor character in this genus. Wheeler (1910a) described C. fallax rasilis from material taken in Texas, Arizona, Louisiana and Florida. The original description was so vague that any small Camponotus with reddish head and thorax and nonpilose cheeks could be assigned to it. The key provided by Wheeler was no better since separation of the various forms depended on color and “average size.” In the same year Wheeler published a second paper (1910b) on Camponotus. Here, C. fallax rasilis was merely listed, while C. sayi was redescribed, evidently from a few cotypes and a few strays taken by Wheeler at Phoenix (the type locality of C. sayi) and at Prescott, Arizona. Wheeler's redescription of C. sayi was reasonably detailed, but in the case of the major workers, it was evidently based upon maximum-sized individuals representing the ideal condition. Further commentary was provided: “This species, as Emery has remarked, is very similar to C. fallax discolor. It is even more like fallax rasilis, but the head and thorax are more robust, the head is more excised behind, the clypeal notch is smaller, the epinotum more angular and the sculpture is different, the punctures on the sides and front of the head being much smaller and the surface of the head and thorax somewhat more shining. These differences are not very pronounced and it may be necessary, when sayi is better known, to reduce it to the rank of a subspecies of fallax.” Another key was given; this key included C. sayi, not included in the earlier paper. Here, C. sayi was separated from C. fallax forms by its larger, broader head and the non-pubescent gaster. Creighton (1950) used a different method for separating C. sayi from C. rasilis (elevated there to specific rank). He utilized the PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 357 relatively shorter scape of C. sayi, said to fall short of the occipital comers. In C. rasilis the scape was said to extend beyond the occipital comers by at least the apical breadth of the scape. Since one of the cotype majors of C. sayi has the scape extending beyond the occipital comers, a character seen also is some cotypes of C. sayi bicolor, it seemed prudent to investigate this character in de- tail. All available cotype majors of C. sayi and C. sayi bicolor were measured for head length and scape length. Similar measurements were made on randomly selected majors of C. rasilis from Mississippi, Georgia, Texas and Oklahoma. The results are shown in fig. 4. It is evident that a poorly defined regression zone exists and that there is a tendency for the eastern populations to exhibit a relatively longer scape. However, it is equally obvious that there is a broad overlap in scape length. Interestingly, the cotypes of C. sayi fall very clearly in the median area. Since these data suggest at least a partial differentia- tion I next attempted to correlate scape length with the cephalic index (HL + HW x 100), the results of which are shown in fig. 5. Finally, the relationship of head length to head width is shown in fig. 6. These data confirm those indicated in fig. 4, that differentiation does exist but that it is neither significant nor consistent. On the basis of cephalic characters, it is clear that neither C. sayi bicolor = californicus nor C. rasilis is worthy of separation. I believe that all the differences cited to separate these forms are the results of allometry; relative scape length decreases, cephalic punctures become finer and sparser, the occipital excision becomes more pronounced, integument becomes shinier, angles become more exaggerated, with the increase in size. The only difference left is that of the supposedly non-pubescent gaster of C. sayi versus the condition of C. rasilis in which there are evident scattered fine ap- pressed hairs. These hairs are present in C. sayi, but least evident in the largest specimens. This is due, I think, not to a reduction in the number of such hairs, but rather to the more shining integument of these individuals. These hairs are obvious in C. rasilis and small speci- mens of C. sayi because they reflect light and hence are more apparent against the duller integument. But, in the case of specimens with a nearly polished integument, the entire surface is highly reflective and these fine, appressed hairs are merely more difficult to perceive. Because the differences which purportedly separate C. sayi from C. rasilis are correlated with allometric growth I see nothing to be gained by the continued separation of these insects. It should be further clear that, while the eastern and western populations tend to exhibit some differences, these are slight and of little practical value. The populations from Texas eastward appear to be more constant in their characters and the maximum head length of the majors is a little less than is true of the western populations (fig. 4). Coupled 358 PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 with this smaller size is a relatively longer antennal scape and a slightly duller integument (especially on the head). The western populations are highly plastic (perhaps in response to greater variation in habitats and climate) and individuals within a single colony sample may possess either a long or short scape, or one of intermediate length. REFERENCES Brown, W. L., Jr. 1950. The status of two common North American carpenter ants. Ent. News 61:157-161. Creighton, W. S. 1950. The ants of North America. Bull. Mus. Comp. Zool. 104: 1-585. 1952. Studies on Arizona ants (4). Camponotus (Colobopsis) papago, a new species from southern Arizona. Psyche 59:148—-162. 1965. Studies on southwestern ants belonging to Camponotus, sub- genus Myrmobrachys. Amer. Mus. Novit. 2239:1-9. and R. R. Snelling. 1966. The rediscovery of Camponotus yogi Wheeler. Psyche 73: 187-195. Gregg, R. E. 1963. The ants of Colorado. Univ. Colo. Press, Denver, xvi + 792 pp. Roger, J. 1863. Die neu aufgefiihrten Gattungen und Arten meins Formiciden- Verzeichnisses. Berlin Ent. Zeit. 7:131-214. Smith, M. R. 1953. A new Camponotus in California apparently inhabiting live oak, Quercus sp. Jour. N. Y. Ent. Soc. 61:211—214. Wheeler, W. M. 1910a. The North American forms of Camponotus fallax Ny- lander. Jour. N. Y. Ent. Soc. 18:216—-232. 1910b. The North American ants of the genus Camponotus Mayr. Ann. N. Y. Acad. Sci. 20:295-354. THE MAYFLY GENUS HEXAGENIA IN MEXICO (EPHEMEROPTERA: EPHEMERIDAE ) W. P. McCarrerty, Department of Zoology and Entomology, University of Utah, Salt Lake City, Utah 84112 All reported records of the genus Hexagenia Walsh in Mexico, Cen- tral and South America are of species of the subgenus Pseudeatonica Spieth, while those species known from America north of Mexico are all of the subgenus Hexagenia s.s. During the course of a generic study of the family Ephemeridae, specimens of Hexagenia (Hexagenia) bilineata (Say) and H. (H.) limbata (Serville) from Mexico have been studied as follows: H. bilineata: El Banito Valles, San Louis Potosi, June 26, 1940, H. Hoogstraal and K. Knight, 1 male imago in the collection of the University of Utah. H. limbata: Ajijic, Lake Chapala, Jalisco, August 23, 27, 28, 1966, Marion E. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 359 Smith, 2 males and 1 female imago in the collection of the University of Utah; Rio Guayalejo, Tamaulipas Province, December 22, 1939, Lewis Berner, 8 im- mature nymphs in the collection of Jay R. Traver. The above records represent a considerable extension of the reported range of both these species (see Spieth, 1941; Hamilton, 1959), yet the distributional extension appears to follow the pattern of many species. Spieth (loc. cit.) indicates localities in Texas where both species have been taken and also cites a New Mexico record for H. bilineata. H. limbata is also reported from Colorado, Utah, and California. Type material of Hexagenia (Pseudeatonica) mexicana Eaton were the only specimens of the genus previously reported to occur in Mexico, and Eaton (1883-1888) cites only “Mexico” as the type locality for this species. It is probable that H. (P.) mexicana is restricted to the Neotropical regions of Mexico and that there is a natural geographic separation of the respective subgenera. Additional collecting in both Mexico and Central America is needed before our understanding of this distributional problem is complete. It is of interest here to note that Kimmins (1960) regards Pseude- atonica as a full genus. This may well be the case; however, since the nymphs of Pseudeatonica are unknown at the present time, it re- mains a question as to which ranking of the taxon is correct. Edmunds and Allen (1966) have pointed out the importance of knowing the nymphal stage of mayflies before a proper classificatory arrangement can be constructed. Undoubtedly the nymphs, when known, will help clarify the status of Pseudeatonica. I would like to thank Dr. George F. Edmunds, Jr., Dr. Marion E. Smith and Dr. Jay R. Traver for supplying specimens collected in Mexico. REFERENCES Eaton, A. E. 1883-88. A revisional monograph of recent Ephemeridae or may- flies. Trans. Linn. Soc. Lond., Sec. Ser. Zool. 3:1—352. Edmunds, G. F., Jr. and R. K. Allen. 1966. The significance of nymphal stages in the study of Ephemeroptera. Ann. Ent. Soc. Amer. 59(2):300-303. Hamilton, E. W. 1959. A tabular summary of the biology of North American mayfly nymphs (Ephemeroptera). Iowa State Coll. Jour. Sci. 33(4):443-474. Kimmins, D. E. 1960. The Ephemeroptera types of species described by A. E. Eaton, R. McLachlan, and F. Walker. Bull. Brit. Mus. (Nat. Hist.) Ent. 9(4): 269-318. Spieth, H. T. 1941. Taxonomic studies on the Ephemeroptera, Il. The genus Hexagenia. Am. Midland Naturalist 26:233-280. 360 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 A CASE OF PARTIALLY REDUPLICATED ANTENNA IN MINETTIA OBSCURA (LOEW) (DieTERA: LAUXANIIDAE) A male specimen of Minettia obscura (Loew), taken on low vegetation along Cabin John Creek west of Bethesda, Maryland, on 27 May 1968, along with many M. lyraformis Shewell and a few M. americana Malloch, M. lobata Shewell, and other M. obscura, was seen to have an abnormal right antenna, as shown in the accompanying figure. The left antenna is normal, as apparently is the rest of the specimen. LUNULE B Fig. 1. Teratological right antenna of Minettia obscura (Lw.); A—profile; B—dorsomesal view of portion not obscured by left antenna (2, 3-—segments 2 and 3 resp. of basic part of antenna). The teratological antenna consists of a normal first segment, a dorsoventrally broadened second segment, and an extra third segment bearing an extra arista apparently proceeding from a groove in the lower side of the extra third segment. The lower, more normal third segment differs from that of its mate only in slightly different outline, being slightly broader and more angulate apically. The extra third segment bears on its mesal side a deep oval pit. The specimen has been deposited in the collections of the United States National Museum.—GeorceE C. STEYSKAL, Systematic Entomology Laboratory, Entomology Research Division, ARS, USDA, c/o U. S. National Museum, Washington, D. C. 20560. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 361 THREE NEW SPECIES OF THE GENUS TRIPTEROIDES, SUBGENUS TRIPTEROIDES GILES! ? (DreTERA: CULICIDAE ) Mercepbes D. DELFINADO® and ELAINE R. Honces, Southeast Asia Mosquito Project, Smithsonian Institution, Washington, D. C. 20560 INTRODUCTION The subgenus Tripteroides of the genus Tripteroides Giles is a complex group of about 50 species occurring from India and Ceylon, through Thailand and Viet-Nam, north to China, Taiwan and Japan, and south through the Philippines, Malaysia, Indonesia, New Guinea, the Solomon Islands, and Australia. Pending a revision of the subgenus we are describing three new species from Malaya and Thailand. We feel that these species draw attention to certain diagnostic characters in this subgenus. We are following the classification proposed by Belkin (1962) who recognized three subgenera as follows: “Tripteroides for the orna- mented species, Rachisoura for nonornamented species with predaceous larvae with incomplete, widely separated maxillary sutures, and Rachio- notomyia for all the other nonornamented species.” The larval and pupal chaetotaxy and terminology used is also that of Belkin (1962) for the most part. Following Knight (1968), we are labeling the larva] basal maxillary hair “bmh,” having found it to be of some taxonomic value. Type specimens will be deposited in the U.S. National Museum, Washington, D.C., and the British Museum ( Natural History ), London. Many of the Tripteroides (Tripteroides) species are difficult or impossible to separate using adult coloration and scaling alone. The species usually have brilliant blue scales on the vertex; silver patches on pleuron, abdomen, and femora; narrow dark scales on scutum; broad black scales on scutellum and anterior pronotum and narrow ones on scutum and posterior pronotum. Color of scutellar and pleural integu- ment is somewhat variable, though in a few species it seems constant enough to be helpful. Differences between some species may be noted in the extent of silver pleural scaling, the shape and extent of silver abdominal patches, and differences from the norm described above; however, in some species these characters tend to be somewhat variable also. A good many of the members of this subgenus characteristically have rows of long scales and diagonally striated setae at the apex of the 1 This work was supported by Research Contract No. DA-49-193-MD-2672 from the U.S. Army Medical Research and Development Command, Office of the Sur- geon General. 2 Immediate publication secured by full payment of page charges.—Editor. ’ Present address: Department of Entomology, University of Hawaii, Honolulu, Hawaii 96822. 362 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 male hind tibia and base of hind first tarsomere; additional specialized scaling of the male legs as well as claw structures present valuable spe- cific characters. The female terminalia have not provided useful char- acters for other workers and have not been used here. Male terminalia in some cases look similar superficially but on closer examination show good characters. The ninth tergum, distimere, and paraproct have traditionally been used, but we found Baisas and Ubaldo-Pagayon’s method (1953) of dissecting the aedeagus and studying it from lateral as well as dorsal views especially productive. The ventral projection of the aedeagus (Baisas calls it the ventral arm) can be seen clearly only in lateral aspect. Lien (1958, p. 15) illustrates his new Tripteroides (T.) cheni in lateral view to differentiate it from bambusa (Yamada), but few other authors have given illustrated descriptions of this structure. In the immatures there are many similarities among species and much specific variation. In pupae the length and character of abdom- inal hairs 3-VII and 5-II-VI are sometimes useful. Paddle shape and size and the location of paddle spicules, when present, may prove to be valuable characters. Because there are no striking differences among the pupae of the present species, they are not described in detail, and only tarsalis, n. sp. as a whole and paddles of malayi, n. sp. and denticu- latus, n. sp. are figured. Larval differences may be found in the basal maxillary hair (bmh), 14-C, possibly other head hairs, and, most useful, segment VIII, siphon, and anal segment. Larvae of the Southeast Asian species have mesothoracic tubercles protruding to a small point but never with a long pointed lobe which is present in some species from New Guinea, Solomon Islands, and Fiji [bimaculipes (Theobald), binotatus Belkin, brevipalpus Brug, distigma (Edwards ), lipovski Bel- kin, purpuratus (Edwards ), and quasiornatus (Taylor) ]. The three species that we are describing have an aedeagal type which generally resembles that of bambusa from Japan, Taiwan, China, Ryukyu-Retto and vicinus (Edwards) from Borneo, Malaya, China, Sumatra in having dorsal teeth and twin ventral projections, each of which has an anterior hump, sometimes faintly sclerotized, immedi- ately sternad of the main aedeagal tube. The descriptions are mainly based on the holotypes, but the extent of variation in all the specimens which were available for examination is noted. Tripteroides (Tripteroides) denticulatus, n. sp. (Figs. 1, 4, 5) Mae. Head. Vertex with bright blue decumbent scales; lateral surface with sil- very scales; occiput with erect dark brown scales; torus bare, yellowish or whitish- brown; clypeus bare, pale brown; proboscis about 1.5 longer than fore femur, underside of base of proboscis with a number of dark bristles; palpi uniformly dark, about 0.1 length of proboscis. Thorax. Scutal integument brown, darker at dorso- PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 363 ! watt (0 vill 4- l ‘ny > 1 NNN J > F aa tey whi MP 5 A ~~ 4 I< MALAY (ects la \ + : \ fo Ay, | s | 8 1.0mm 7 nedge \S \ 0.05mm aedeagus \ \ \ / : Pes I ~\ el all MALAYA Wann / (holotype) \ bie Dy foie f ‘\paraproct \ é ore IX tergum ez THAILAND DENTICULATUS 364 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 central portion, yellowish around anterior promontory, humeral and_ supraalar areas, with dark brown narrow scales and bristles; yellowish or beige prescutellar space and scutellum, the latter with broad dark brown or black scales; anterior pro- notum yellowish with broad dark brown scales and a number of strong bristles, posterior pronotum yellowish with narrow dark brown scales interspersed with a few broader ones, with 1 dark bristle; propleuron yellowish with 2 bristles; pleuron medium brown on postspiracular and spiracular areas, sternopleuron, paratergite and mesepimeron; meron and sometimes paratergite pale brown; sternopleuron with silvery broad appressed scales from below prealar to above mid coxa; a patch of translucent silvery scales on mesepimeron. Legs. Silvery scales on coxae; anterior surface of each femur with a silver line extending from base to one-third and 2 silver spots, 1 median and 1 subapical (not well defined on the fore femur of the holotype); legs otherwise dark anteriorly; dorsal edge of fore tibia with an inwardly inclined row of 14-18 more or less regularly spaced stiff, long semi-erect spines, each of these spines, on the whole, longer than the diameter of the tibia. Fore tarsomere V modified; a paratype slide preparation shows the plantar surface with 2 stout, sharply pointed basal spines mounted on tubercles plus 4 other tubercles each bearing a small seta (as in malayi, fig. 4). Mid tarsomere V unmodified. Hind leg with a group of long semi-erect setae and scales at apex of tibia and base of tarsomere I (fig. 4); these scales and setae are shown enlarged, setae with diagonal striations and scales with a central rib resembling a long nar- row feather; tarsomere V unmodified. Claws. (Fig. 4, drawn from paratype). Fore tarsal claws unequal, the larger claw with a strong median tooth projecting at about right angles to it; mid claws small, simple and equal; hind claws very small, simple and equal. Abdomen. Terga dark dorsally, with distinct postero-lateral patches of silvery-white scales on II-VII (patches larger on II and VII); I and VIII entirely dark. Abdominal maculation is generally similar to tarsalis (fig. 4) though lateral patches tend to be smaller in denticulatus; sterna pale gold. Terminalia. (Fig. 1, drawn from holotype and 2 Thailand paratypes). Basimere with long bristles on lateral distal margin; basal lobe with about 10 long and some short bristles; distimere incurved, attenuated at middle, with a few fine setae distally and an apical flattened spiniform appendage; aedeagus tapered distally with a variable number of strong middorsal teeth (4-10) and a ventral projection serrated finely on distal margin; paramere outwardly curved; paraproct with 6-8 teeth and 4-8 cercal setae; tergum IX with 6-9 apical bristles of equal length on each lobe. The aedeagus of the paratype shown in lateral view appears to differ slightly from that of the holotype. At this stage we feel that these differences are not of specific value, though this might prove to be so when more material is available for examination. Pupa. (Fig. 5, paddle drawn from holotype, 4 specimens examined.) Hair 3-VII reaching to or beyond posterior margin of VIII; paddles 1.5—2.0 times as long as wide, smooth and rounded apically, without spicules. Except for paddles, pupa resembles tarsalis as illustrated. Larva. (Fig. 1, drawn from paratype, 4 specimens examined.) Head. 4-C single, long, flattened and expanded at middle, tapering to a fine point; 7-C about as long as 4-C, 5-7 branched; 8-C normally with 2 weak branches (1-3); 9-C with 2-6 short, weak branches; mental plate (MP) with 17-19 teeth; strong basal max- illary hair (bmh) varying in size from much smaller to as large or larger than 14-C, usually with 3-11 branches; 14-C usually with 5 or fewer branches, about 3% length PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 365 Ny (paratype) IX tergum MALAY/ MALAYA 366 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 of antenna; 15-C usually with 5-6 branches (2-6). Thorax. 7-T a 2-branched heavy, barbed spiniform with serrated tips. Abdomen. 6-I-V double; 7-I-II double, 3-branched on one side of segment I of one specimen; segment VIII with 16-25 small comb scales (CS), dorsolateral comb scales pointed, ventral ones broadly flattened and fringed; 1-S with 3-5 (on one side of one specimen, 2) moderately long branches; la-S number 8-10 tufts, usually 2-branched (1-2); 3-5 widely spaced pecten teeth (PT) on each side; 2a-S number 7-8, scattered dorsally and laterally with 1-3 branches; saddle with 2—4 stout, relatively long and 2-5 weaker, short marginal spines; 1-X with 1-3 branches; 4-X with 3—4 short branches, about half length of anal papillae; anal papillae about 2% times length of saddle. Type DATA. Holotype male #0914/11, Lower Perak, Pulau Tiga, Mataya, 14-11-58 (W. W. Macdonald), terminalia, larval and pupal skins on slides; paratypes all males: #0914/9, same data as for holo- type, with cast skins mounted; #01281/24, 8 mi. Bantung Rd, Selangor, Mataya, 27-8-59 (W. W. Macdonald), terminalia, legs, larval and pupal skins mounted; #PU 24-35 and #PU 24-40, both collected in 1964 from Phatthalung, Muang, THamanp (S. Chunchulcherm), and both with terminalia mounted. Both Thailand specimens without associated cast skins. The holotype and 1 paratype with associated larval and pupal skins from Malaya will be deposited in the British Museum (Natural History ); the 2 paratypes from Thailand and 1 para- type with associated skins from Malaya are deposited in the U.S. National Museum. In addition there is 1 male from Malaya not labeled paratype which will be at the British Museum, namely, #0677/6 from Templer Park, Selangor, 25-6-58, with hind leg and larval and pupal skins mounted. DiIsTRIBUTION. MALAYA; THAILAND. Hasrrat. Tree holes. TAXONOMIC biscussion. T. denticulatus differs from similar species, bambusa, malayi, and tarsalis, in having a large tooth on the larger fore claw of the male, equal male mid claws, a simple male mid tarsomere V (lacking spines or tubercles), feather-like scales on male hind tarso- mere I, and small but distinct serrations on the distal margin of the aedeagal ventral projection. The aedeagal middorsal teeth of denticu- latus are confined to the central area, whereas those of bambusa extend to the lateral edges of the aedeagus (fig. 5). The male fore claw of bambusa (fig. 5) has a small triangular tooth, much smaller than that of denticulatus (fig. 4). There are no reliable differences among the larvae and pupae of these species other than the presence of spicules on the pupal paddles of tarsalis and bambusa which are not present in denticulatus. The species most resembling denticulatus is vicinus, also found in Malaya. The male of vicinus has a similar aedeagus, identical feather-like scales on the hind tarsomere I, an equally simple mid tarsomere V, and a fore claw with equally large tooth (fig. 5). However, this tooth arises at about a 70 degree angle from the claw, PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 367 sat THAILAND I (paratype) olosmm TARSALIS 368 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 whereas the same tooth of denticulatus projects at about a 90 degree angle. T. vicinus differs more significantly in that tergum IX has each broad lobe slanted so that the length of inner edge is about one-third length of outer edge and that each lobe bears as many as 28 bristles which are twice as long laterally as medially (fig. 5); the male mid claws are unequal; its pupal paddle is heavily spiculed, and pupal seta 3-VII extends no more than to half the length of VIII. The larva of vicinus differs from denticulatus in having comb scales on a plate which is sometimes incompletely formed and in having 4-X double, longer than the anal papillae, and almost as long as 1-X. Tripteroides (Tripteroides) malayi, n. sp. (Figs. 2, 4, 5) Mate. Head. Vertex with comparatively narrow anterior band of bright blue decumbent scales, lateral portion with silvery scales; the rest of the head with dark brown scales; occiput with row of erect dark brown scales; torus and clypeus grayish-brown; proboscis dark brown and about 1.3 as long as fore femur, several dark basal bristles present; palpi dark brown, 0.1 the length of proboscis. Thorax. Integument of scutum dark brown, yellowish on humeral area, with narrow dark brown scales (most are rubbed off); scutellum yellowish and heavily clothed with flat, broad black scales and 8 bristles; postnotum medium brown; anterior and posterior pronota yellowish; 8-9 black bristles and broad dark brown scales on anterior pronotum; narrow dark scales and 1 dark bristle on posterior pronotum; yellowish propleuron with 2 bristles; paratergite, postspiracular area, sternopleuron, mesepimeron, and meron dark brown; silvery broad appressed scales on sterno- pleuron from below prealar area to above mid coxa; a short line of silvery appressed scales between the sternopleuron and postspiracular area; mesepimeron almost bare except for a small patch of about 5 translucent silvery scales on anterior portion. Legs. Coxae with translucent scales; each femur with 2 distinct silvery spots—1 median and 1 sub-apical; mid femur also has silvery line running from just below base to one-third its length; hind tibia with diagonally striated setae on apical fourth and long spines and setae at apex; semi-erect scales on base of hind tarso- mere I; no long feather-like scales are apparent on any of the legs and no special- ized scales or setae exist on the mid legs; plantar surface of fore tarsomere V modified with 2 stout, sharply pointed basal spines plus 2—5 tubercles with small setae extending from them (fig. 4, holotype); plantar surface of mid tarsomere V modified with about 4 small tubercles, each with a small seta; hind tarsomere V unmodified. Claws. (Fore claws drawn from holotype; mid and hind claws drawn from paratype.) Fore and mid tarsal claws unequal, simple; hind claws small, simple, equal. Abdomen. (Fig. 4, holotype.) Terga dark dorsally; segments II and III with silvery lateral patches deeply emarginated along their apical borders (emargination more pronounced on III); lateral silvery patches on IV-VI divided into a larger and smaller patch on each segment; on segment IV of the paratype the patches are not as widely separated as on the holotype and are connected anteriorly by a single line of whitish scales; sterna pale gold. Terminalia. (Fig. 2, holotype.) Basimere with a few long strong bristles and many short ones; disti- mere slightly swollen in distal half with a few scattered fine setae and an apical spiniform appendage; aedeagus tapered distally, with strong middorsal teeth on PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 369 ay) = IE sie itp ee : LI gan : 2 LF 1.0mm tarsomeres “Y tarsomeres i; \ tarsomeres Hip tarsomeres MALAYA TARSALIS EGS oO SS SS TARSAL/S abdomen Ss MALAY/ abdomen rot Giclaws te ] MALAYA 0.05mm | DENTICULATUS DENTICULATUS 370 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 each side of center line; ventral projection as in fig. 2, without teeth; parameres curved outwardly; paraproct with 5-6 teeth, 4-6 cercal setae; tergum IX with 8 bristles of equal length on each lobe in the holotype, 10 and 12 on lobes of the paratype; each lobe broad and slightly expanded at apex. Pura. Chaetotaxy similar to tarsalis (fig. 5). 3-VII long, reaching beyond posterior margin of VIII. Paddles gently rounded apically, not as pointed as tarsalis, smooth, without spicules, 1.8—2.0 times as long as wide (paddle drawn from holotype, fig. 5). Larva. (Fig. 2, primarily holotype.) Head. 4-C single, expanded and flattened medianly, tapering to a fine point; 7-C 5-branched on the one specimen where it was present, a little longer than 4-C; 8-C 1-2 branched; 9-C very short branched tuft; 10-C single although with weak branch on one side of holotype and forked distally on one side of paratype; basal maxillary hair (bmh) strong, short, stellate; 14-C 2-3 branched; 15-C 3-5 branched; mental plate (MP) with 17-20 teeth. Thorax. 7-T spiniform, strongly barbed, single or double, longer than in most other species. Abdomen. 6-I-II 2-branched, 6-III-V 1-2 branched; 7-I single, 7-II 1-2 branched. Segment VIII with 10-12 comb scales (CS), spine-like except for 2-3 of the most ventral ones which have broad fringed apices; 1-S 3-branched; la-S total 9-10 tufts of 1-2 branches; 2a-S with 8-10 tufts scattered dorsally and laterally, 2-3 branched; siphon with 3 pecten teeth (PT) on each side; saddle with 2-3 long, strong marginal spines; 1-X single, very long; 4-X single, short and fairly strong; anal papillae longer than saddle. Type DATA. Holotype male #0613/27, Trengganu, Gunong Tabu, Kuanta, 20 mi. Pahang Rd, Mataya, 7.v.58 (W. W. Macdonald), ter- minalia, legs, larval and pupal skins on slides; paratype male #0613/20, same data as holotype, terminalia, legs, pupal and larval skins on sides. Holotype with associated mounts will be deposited in the British Museum (Natural History), London; paratype and associated mounts will be retained in the U.S. National Museum. Hasirats. No record. DiIsTRIBUTION. MALAYA. TAXONOMIC DISCUSSION. Despite aedeagal resemblances to vicinus, bambusa, tarsalis, and denticulatus, malayi can be differentiated by its simple untoothed male tarsal claws, the absence of unusual setae and scales on the male legs, ventral projection of aedeagus without serra- tions, middorsal teeth confined to centerline of aedeagus, and the sil- very lateral abdominal patches which are medianly emarginate on III and double on IV-VI. Tergum IX of malayi differs from that of vicinus in the same way as does this structure in denticulatus, although malayi has more tergal bristles (8-12) than denticulatus (see vicinus, fig. 5). The larva of malayi differs from the above mentioned species in having 4-X single and only 11-12 CS, but it resembles the larva of similis (Leicester), also found in Malaya, in these as well as in some other characters. However, similis has even fewer CS (6-7), a larger bmh, and a large stellate tuft for 14-C. The pupa of similis, like vicinus, differs from malayi in having spicules on its paddle and a short 3-VII, a PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 BAMBUSA SS x eg fore claws aS S: ae middorsal BAMBUSA TAIWAN V/C/NUS aedeagus dorso- lateral IX tergum VICINUS BORNEO S/MILIS MALAYA TARSALIS THAILAND MALAY/ MALAYA DENTICULATUS MALAYA 371 ore PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 reaching no more than one-half the length of VIII. There are clear dif- ferences between malayi and similis in the adult stage; similis has sil- very scaling on the anterior and posterior pronota, large and unbroken lateral abdominal silvery patches, and a tooth on the larger fore claw of the male. Its terminalia resembles nitidoventer (Giles) with markedly swollen distimere, tergum IX with short bristles becoming somewhat longer laterally, and aedeagal ventral projection bearing anterior teeth without hump (see similis, fig. 5). Adult characters of malayi which differentiate it from similis, in addition to those mentioned earlier, are black scaling on the pronotal lobes, a relatively slender distimere, long bristles on tergum IX, and an untoothed aedeagal ventral projection with anterior hump. Tripteroides (Tripteroides) tarsalis, n. sp. (Figs. 3, 4, 5) Mate. Head. Anterior half of vertex with bright blue decumbent scales, lateral portion with silvery scales; occiput with a row of erect dark brown scales; clypeus and torus bare, gray-brown; proboscis dark, measuring about 1.25 as long as fore femur, several dark basal bristles present; palpi dark, measuring about 0.1 length of proboscis. Thorax. Scutum yellowish-brown with narrow blackish scales, brownish supraalar and anterior promontory bristles; scutellum and postnotum yellowish- brown; anterior and posterior pronota and propleuron yellowish; anterior pronotum with broad dark scales and about 6 dark bristles; posterior pronotum with narrow dark scales interspersed with a few broad dark ones and 1 long, dark bristle; pro- pleuron with 2 brown bristles; scutellum with broad dark scales and 8 dark bristles; dark medium brown patch covering paratergite, postspiracular and subspiracular areas, sternopleuron, mesepimeron, and meron; silvery broad appressed scales on sternopleuron from below prealar area to above mid coxa, a row of a few silvery appressed scales between the sternopleuron and postspiracular area; mesepimeron largely covered with silvery broad scales. Legs. Coxae with translucent scales; femora with the usual anterior silvery spots, 1 median and 1 subapical; mid femur with silver line extending from base to about one-third; on fore femur this stripe appears to be composed of yellowish scales extending from base to half length of femur, and on hind femur the stripe is absent. Fore tarsomeres II-V modified; II with some semi-erect scales; III with many erect and semi-erect scales; IV about half length of V with a few semi-erect scales; V with a few plantar tubercles sur- mounted by strong setae and 2 small pointed spines near the base, a number of very long scales also on the plantar surface followed by a long seta extending from shortly below a much reduced empodium, the latter with unusually short setae projecting from its apex; from the apex of tarsomere V projects a long, slender pedestal bearing a club-shaped structure covered with very narrow scale-like setae with recurved tips; mid leg with tarsomeres II-V modified; II with some semi- erect scales; III with semi-erect scales and a prominent basal patch of long, narrow, sinuous striated scales, the longest of these reaching to near the apex of the tarso- mere; this patch of scales is visible in pinned specimens as a curved tuft which stands out at an angle as shown in fig. 4; mid tarsomere III also with a row of erect scales which in pinned specimens look like spines; mid tarsomere IV about two-fifths longer than V with a row of erect scales on basal half; mid tarsomere V ioe) ivy) PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 lacks any tubercles but is slightly modified in having its diameter narrowed medi- ally, presenting a curved appearance, and in having a greatly reduced empodium; hind leg is as usually seen in the subgenus, without any modifications. Claws. (Legs and claws drawn from holotype.) Fore tarsal claws elongated and slender, subequal, the smaller claw narrow and hairy to before tip, the larger claw expanded into a very setiferous lobe medianly with a long seta arising below the lobed por- tion and reaching nearly to the tip of the claw; mid claws unequal, narrow, the smaller one simple, the larger bearing many setae and a small setiferous lobe near the apex; hind claws very small, simple, equal. Abdomen. (As in fig. 4, paratype female.) Terga dark brown dorsally with silvery-white lateral bands on III-VI, subapical on II-IV, apical on V-VI, with silvery patches almost completely covering the sides of II and VII; none of the silvery patches or bands meet dorsally; seg- ments I and VIII all dark brown; sterna pale gold. Terminalia. (Fig. 3, holotype). Basimere with long strong bristles on lateral and distal margins; 4-6 long and many short bristles on basal lobe; distimere attenuated at middle, fine setae at distal portion and with an apical flattened spiniform appendage; aedeagus tapered distally with sharply projecting lateral shoulders, strong sharp middorsal teeth confined to each side of center line, a narrow ventral projection without teeth; paraproct with 5-7 teeth, usually with 4 cercal setae (2-5); parameres outwardly curved; tergum IX with 4-8 bristles on each lobe. FEMALE. Coloration as in male. Legs. Apex of hind tibia and basal fifth of tarsomere I with long semi-erect setae, many of which are diagonally striated, and with a few feather-like ones on the tibia like those drawn for the male denticulatus (fig. 4, enlargement of setae and scales). Claws. Simple, small, equal on fore and mid; simple, small and unequal on hind leg. Pupa. Chaetotaxy as in fig. 5 (not drawn from paratypes, 20 specimens exam- ined). 3-VII reaches to or beyond posterior margin of VIII; paddles 1.5-2.0 as long as wide, pointed apically and with fine spicules on apical portion, extending up inner margin. Larva. (Fig. 3, not drawn from paratypes, 17 specimens examined). Descrip- tion based on cast skins of paratypes. Head. 4-C single, long, expanded and flat- tened medially, tapering to a fine point; 7-C shorter than 4-C, with 3-5 branches; 8-C usually with 1-2 branches (1-3); 9-C about as long as 8-C, with 2-5 branches; mental plate (MP) with 15-18 teeth; bmh ranges from about half to three-fourths the length of 14-C, 8-16 branched; 14-C about two-thirds length of antenna, with 6-12 branches; 15-C 5-9 branched. Thorax. 6-M a barbed, stout spiniform, usu- ally single, but 2-3 branched in some specimens; 7-T normally a 2-branched, pointed, barbed spiniform, occasionally single or 3-branched. Abdomen. 6-I-IV with 2 branches, 6-V occasionally single; 7-I-II single, though 2-branched on one side of one specimen; segment VIII with 16-26 comb scales; the dorsolateral scales pointed, a few lateroventral ones flattened, fringed; 1-S with 3-5 branches; la-S 8-14 in number, 1-3 branched; 2a-S number 8-12, scattered dorsally and laterally, usually with 2 branches (1-3); siphon with 3-5 PT on each side; saddle with 3-5 long, 2-3 short marginal spines; 4-X normally with 3-4 branches (3-5); 1-X usually single, long; anal papillae more than twice the length of saddle. Type pata. Holotype male #689/3, Fraser’s Hill, W. Pahang, Mataya, Sept., 1954 (J. A. Reid), terminalia and legs on slide; 3 para- types from Ma.aya include 1 female #689/5, same data as holotype, 374 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 with hind leg and antenna, larval and pupal skins mounted; 1 male #0611/2, Gunong Tabu, Trengganu, 7-5-58 (W. W. Macdonald), ter- minalia, larval and pupal skins mounted. Sixteen male paratypes from THAILAND were collected by SEATO Lab personnel in 1963 and 1964 (Sgt. E. Peyton, P. Boonyakanit, $. Chunchulcherm, C. Diraphat, S. Esah, S. Maneechai, S. Maniwongse, K. Mongkolpanya): Six originate from the province Nakhon Nayok, THatLanp: #NY 37-20 and #NY 37-24 from Moh Sing To, both with larval and pupal skins mounted; #NY 37-22 also from Moh Sing To; #NY 122-32 from Phaklory Mai, terminalia and larval skin mounted; #NY 123-10 from Khaoyai, termi- nalia and legs mounted; #NY 123-11 from Khaoyai. Five specimens came from Chiang Mai: #T-2195-10L and #T-2123-1P from Doi Sutep; #T-2413-1P, #T-2753, #T-2761. Two specimens were collected in a rain forest, 1 mile in altitude, located at Doi Sam Sao, Tak: +00265-104 and #00265-110. One specimen is from each of the follow- ing provinces: Chanthaburi, Khao Sai Dao Tai, evergreen forest, 1 mile altitude, #00864-8, pupal and larval skins mounted; Nakhon Si Thammarat, Ban Thuan Lek, primary rain forest, 1,000 ft. altitude, #01082-104; Trang, #TG-102-33, terminalia mounted. The holotype with associated slides will be deposited in the British Museum ( Nat- ural History ), London; paratypes with associated slide mounts will be distributed between the British Museum (Natural History), London and the U.S. National Museum, Washington, D.C. Hasrrat. Tree holes. DisTRIBUTION. MALaya, THAILAND. In addition to the holotype and paratypes, 1 male with associated pupal skin from Selangor, MALAya, and 25 specimens from THAILAND were examined. The Thailand material contained 1 whole larva and 17 males with 9 associated larval and pupal skins from Chiang Mai; 3 males, 2 with associated pupal skins, and 1 female with associated pupal skin from Nakhon Nayok; and 2 males, 1 with associated larval and pupal skins, and 1 female with associated pupal skin from Chanthaburi. TAXONOMIC piscussion. T. tarsalis can be distinguished from other known species of the subgenus by its male tarsal and claw modifica- tions, especially by the prominent tuft of long scales on mid tarsomere III, erect scales on mid tarsomeres III and IV and on fore tarsomere III, and by its elongated hairy and lobed claws. The aedeagus of tar- salis differs from its nearest morphological relatives (bambusa, denticu- latus, malayi, vicinus) in having sharply defined lateral shoulders. Mid-dorsal teeth are confined to the center line of the aedeagus, in contrast to bambusa. The aedeagal ventral projection is without teeth, unlike denticulatus. The lobes of tergum IX tend to be narrower than those of malayi and differ from vicinus for the same reasons as does tergum IX of denticulatus (see vicinus, fig. 5). The pupa is distinguish- able from malayi and denticulatus in having paddle spicules, from PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 375 vicinus in having 3-VII long, reaching to or beyond posterior margin of VIII, whereas in vicinus 3-VII is much shorter. Pupae of tarsalis and bambusa are virtually indistinguishable, inasmuch as bambusa_ pos- sesses paddle spicules in the same positions as does tarsalis, as well as other similar characters. The tarsalis larva shows no reliable difference from bambusa and denticulatus. It differs from malayi in having a branched 4-X, a greater number of comb scales, and more branches in 14-C. The larva of vicinus differs from tarsalis in the possession of a complete or incomplete comb scale plate, and long double 4-X. ACKNOWLEDGMENTS We are very appreciative of the help given us on this group by Drs. Botha de Meillon and Alan Stone. REFERENCES Baisas, F. E. and A. Ubaldo-Pagayon. 1953. Notes on Philippine mosquitoes, XVI. Genus Tripteroides. Monogr. Inst. Sci. Tech., Manila, 198 pp., illus. Belkin, J. N. 1962. The mosquitoes of the South Pacific. Univ. Calif. Press, Berkeley. 2 vols., 608 and 412 pp., illus. Edwards, F. W. 1914. New Culicidae from Borneo and Hong Kong. Bull. Ent. Res. 5(2,):125-128. Giles, G. M. 1904. Notes on some collections of mosquitoes, etc., received from the Philippine Islands and Angola; with some incidental remarks upon classifica- tion. J. Trop. Med. 7:365-369, illus. Knight, K. L. 1968. Contributions to the mosquito fauna of Southeast Asia. IV. Species of the subgroup Chrysolineatus of group D, genus Aedes, subgenus Finlaya Theobald. Cont. Amer. Ent. Inst. 2(5):1—32, illus. Leicester, G. F. 1908. The Culicidae of Malaya. Stud. Inst. Med. Res. F. M. S. 3(3):18-261, illus. Lien, J.C. 1968. New species of mosquitoes from Taiwan (Diptera: Culicidae ) Part II. New species of Tripteroides, Orthopodomyia, Culiseta and Uranotaenia. Trop. Med. (Japan) 10(1):1-20, illus. Yamada, S. 1917. Nipponsan bunkwa no ni shinska (Two new species of Japanese Culicidae). Dobutsugaku Zassi (Zool. Mag.) 29 (No. 341):61-72 (in Japanese ). ioe) | (on) PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 A REVISED INTERPRETATION OF THE PROCTIGER OF MALE URANOTAENIA WITH A RELATED NOTE ON HODGESIA (DreTerRA: CULICIDAE )*'” E. L. PEyron and R. H. HocumMan, Southeast Asia Mosquito Project, Smithsonian Institution, Washington, D.C. 20560 Although the structure of the male terminalia has long been recog- nized as an important criterion in the classification of mosquitoes, their structure in the genus Uranotaenia Lynch Arribalzaga remains little studied and incompletely understood. The forms of the proctiger and the ninth tergite have received contradictory and confused treatment. Edwards (1920) proposed the term “lobes of the ninth tergite” for the more or less distinct prominences at the apical margin of the tergite. He indicated their various development in different genera, mentioning that in at least one species of Uranotaenia they were without bristles. Subsequent authors recognized Edwards’ lobes in other species of Uranotaenia (Dyar and Shannon, 1925; Matheson, 1944; King and Hoogstraal, 1946; Pratt, 1946; La Casse and Yamaguti, 1950; Galindo, Blanton and Peyton, 1954; Peters, 1963a, 1963b, and 1964). Lane (1943 and 1953) used the term “external bars” for these structures. Belkin (1953 and 1962) described the lobes of the ninth tergite with “ventrolateral sclerotization” in South Pacific Uranotaenia. The proc- tiger was regarded as almost completely or completely membranous. Freeborn (1924), however, studying terminalia of many culicid genera, concluded that in Uranotaenia the proctiger bore on its dorsal face a pair of “broad, well chitinized sclerites recurved over the summit,” to which he applied Crampton’s term “epiprocts.” He seems to have drawn this generalization from an examination of one species, U. geo- metrica Theobald, which he figured with a dorsal lobar plate of the proctiger, discontinuous with the ninth tergite and with no basolateral extension. Carpenter, Middlekauff, and Chamberlain (1946), while following the standard usage of Edwards’ terminology, parenthetically registered doubt that these were true lobes of the ninth tergite in Uranotaenia and suggested that the so-called “lobes” were productions of the tenth tergite or tenth sternite (proctiger). Yamaguti and La Casse (1951) and Carpenter and La Casse (1955) reiterated this inter- pretation. The most thorough examination of Uranotaenia terminalia was made by Dampf (1943) in his study of Uranotaenia syntheta Dyar and Shannon. His excellent, detailed figures of the terminal structures in lateral, posterior, and tergal views show basolateral sclerotization of 1 This work was supported by Research Contract No. DA-49-193-MD-2672 from the U.S. Army Medical Research and Development Command, Office of the Sur- geon General. 2 Immediate publication secured by full payment of page charges—Editor. ~l PROG. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 37 Figs. 1 and 2, Uranotaenia geometrica Theobald, ¢ terminalia: 1, lateral view; 2, anterior view. Figs. 3 and 4, Hodgesia malayi Leicester, ¢ terminalia: 3, lat- eral view; 4, anterior view. Abbreviations: AE, aedeagus; AM, anal membrane; BLS, basolateral sclerotization of proctiger; BM, basimere; BTL, basotergal lobe of proctiger; PT, paraproct; 9T, ninth tergite. the proctiger articulating with a pair of lobes which are fused with the apical margin of the ninth tergite. These lobes he regards as tongue- shaped “protuberancias” of the ninth tergite. Like Edwards, he asserts that corresponding lobes occur in many genera (“tan frecuentes en muchos culicidos” ). In the course of studies of the genus Uranotaenia in Southeast Asia, we have determined that the “lobes of the ninth tergite” of the above- mentioned authors are, in Uranotaenia, wholly continuous with the basolateral sclerotization of the proctiger, and thus they are on the contrary, lobes of the proctiger (figs. 1,2). Belkin and Page in work in progress on Jamaican Uranotaenia have independently arrived at this same conclusion and have proposed the term “basotergal lobes of the proctiger” for these structures (personal communication with Dr. Belkin ). 378 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 This paper presents techniques that allow careful study of terminalia, indicates the range of development of the proctiger lobes in Urano- taenia, and considers their possible phylogenetic significance. It is hoped others will be stimulated to investigate the proctiger develop- ment of additional species of Uranotaenia and allied genera. Further study of the variation in the form of the basotergal lobes of the proc- tiger with speciation will, we feel certain, increase our understanding of the phylogeny within the genus and with other genera. The experiences gained from the present study indicate the need to emphasize the absolute necessity of dissecting the terminalia of Urano- taenia, for an accurate interpretation of all the structures. Whole mounts are totally inadequate and should be made only after an ade- quate series of dissections has been prepared. In view of the obvious value, we present a detailed discussion of techniques we have found satisfactory for the preparation and dissection of the terminalia of Uranotaenia. Our method of preparation of terminalia is modified from that of Galindo, Blanton, and Peyton (1954). After the terminal segments of several specimens are clipped they are placed in small 5 cc test tubes containing 10% KOH. The tubes are then placed in a beaker of boiling water for approximately 20 minutes. The parts are transferred to a weak solution of acetic acid to neutralize the alkali. They are then dehydrated in alcohol and transferred to a staining solution of 1% acid fuchsen in 70% alcohol and allowed to remain for approximately 20 minutes. The staining dish should be as small as possible for the ter- minalia of the smaller species are extremely difficult to find after stain- ing. The dish should be covered to prevent rapid evaporation of the alcohol. The terminalia are transferred from the stain to a drop of liquid phenol on a concave slide for dissection. If clearing of the stain in the phenol appears too rapid, a small drop of copal-phenol can be added to arrest the clearing. The individually dissected parts are posi- tioned in minute drops of copal-phenol on a clean slide and then placed in a drying oven for a minimum of four days in order to drive off the phenol and harden the drops. If larger amounts of copal-phenol are used the required drying period must be increased, for any remaining phenol will crystalize as the slide ages and degrade the quality of the mount. After the copal-phenol is thoroughly dried, small bits of cover- slip glass are placed around the specimen and then covered with a drop of thin balsam and a small cover slip. If prepared in this manner the prepositioned parts will not move and the bits of glass will prevent crushing from the gradual settling of the cover slip. There are other satisfactory preparation methods, but all share two essential cautions. Due to the extremely small size of many Uranotaenia species and the generally weakly sclerotized structures of the terminalia, it is abso- lutely necessary to use a moderate stain after maceration in KOH. PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 379 ay RiGas U. brevirostris Edwards . U. obScura Edwards. U. rossi Delfinado, U. nivipleura leicester . AHosKnuws YU. subnormalis Martini . U. lateralis Ludlow Fig. 5, Tergal views of proctiger of Uranotaenia species. Abbreviations: BLS, basolateral sclerotization of proctiger; BTL, basotergal lobe of proctiger. 380 PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 Dissections should never be made in the staining solution or other dark media, for the structures and points of attachment can not be seen clearly under an ordinary dissecting microscope. Dissecting in the staining solution as suggested by Galindo, Blanton, and Peyton (1954), is the primary reason these authors failed to correctly interpret the lobes of the ninth tergite and the proctiger in the American Urano- taenia. It is strongly suggested that prior to mounting or dissecting any mosquito terminalia, a thorough examination be made from all angles in a clear non-volatile medium such as phenol, glycerin, or clove oil. Only in this manner can all the structures and points of attachment be correctly identified and the structures removed in a logical sequence without damage. With terminalia of Uranotaenia an examination should also be made to assure that the anal lobes are not everted. This condition has been observed in the majority of reared material exam- ined. As the presence and position of apical bristles and the overall configuration of the proctiger are of diagnostic value, an attempt to reshape the structure should be made. We have usually found it suffi- cient to insert a dissecting pin through the anterior orifice of the proc- tiger and gently push the lobes outward until they are fully distended. Our analysis of the development of the proctiger in Uranotaenia is based upon an examination of 30 species (26 Southeast Asian, three American, and one South Pacific). Two American species, U. geomet- rica and U. syntheta, were also examined since they are the only two species whose terminal structures have been treated in some detail by previous authors (Freeborn, 1924; Dampf, 1943). We find in geomet- rica that those dorsal structures, identified by Freeborn as epiprocts, are extensions of the basolateral sclerotization, slightly recurved, but well above the dorsal face of the proctiger—that is, they are the baso- tergal lobes (figs. 1,2). In syntheta the suture between the “protuber- ancias” and the basolateral sclerotization shown by Dampf in his posterior view is an interpretation not supported by our specimens. Furthermore, his lateral view of the basolateral sclerotization shows no such suture. Essentially two types of development are seen in the basotergal sclerotization. In type I the sclerotization takes the form of a single broad median lobe, with or without a shallow or deep median apical emargination (fig. 5: obscura Edwards, nivipleura Leicester, brevi- rostris Edwards, rossi Delfinado). In the extreme form of this type (fig. 5: nivipleura) the median emargination is quite deep and the tergal sclerotization gives the appearance of type II. However, this condition of type I can be distinguished from type II development by the presence of a completely sclerotized dorsal bridge connecting the differentiated corners. This type seems to characterize Group C of Edwards (1941). In type II the basolateral sclerotization terminates in more or less well defined lobes which have an incompletely sclero- PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 381 tized or wholly membranous dorsal bridge. The lobes may be finger- like and well separated (fig. 5: swbnormalis Martini) or extremely shallow and often closely approximated (fig. 5: lateralis Ludlow). This type seems to characterize Edwards’ Groups A and B. We have made a limited investigation of the development of the basolateral sclerotization in other genera. To our knowledge it is only in Uranotaenia and Hodgesia Theobald that the basolateral sclerotiza- tion assumes its peculiar tergal lobar modification. We have studied in detail two species of Hodgesia, malayi Leicester (figs. 3, 4) and bailyi Barraud. In Culex (Melanoconion) Theobald, the basolateral sclerotization is also highly modified but it is reasonably clear that the modification does not adapt it to the same function, even though the precise nature of their function may not be understood. The strong development of ninth tergal lobes as seen in most other genera is conspicuously absent in Uranotaenia and Hodgesia. It is likely that the basolateral lobes in these genera have evolved to assume the function of the ninth tergal lobes. Edwards (1932) erected the tribe Uranotaeniini to include Urano- taenia, Hodgesia, and Zeugnomyia Leicester. Belkin (1962) placed Uranotaenia and Hodgesia in separate monotypic tribes, indicating that the two genera were unrelated and without close relatives. It is beyond the scope of this paper to make tribal classifications; however, the unique specialization of the proctiger and the absence of developed ninth tergal lobes in both genera appear to suggest a closer affinity than hitherto understood. ACKNOWLEDGMENTS We are indebted to Dr. Alan Stone of the U.S. Department of Agriculture and to Dr. Botha de Meillon, Project Leader, Southeast Asia Mosquito Project, for helpful discussions, encouragement, and assistance during the course of the work and for examining and commenting on the manuscript. We also had the privilege of dis- cussing the work with Dr. J. N. Belkin, who kindly made his unpublished contribu- tion on the subject available to us. Our thanks are due to Lt. Col. Bruce F. Eldridge for specimens of U. geometrica from his collection. REFERENCES Belkin, J. N. 1953. Mosquitoes of the genus Uranotaenia in the Solomon Islands (Diptera: Culicidae). Pacif. Sci. 7:312-391. 1962. The mosquitoes of the South Pacific (Diptera, Culicidae ). Vol. 1. University of California Press, Berkeley and Los Angeles. 608 pp. Carpenter, S. J., W. W. Middlekauff, and R. W. Chamberlain. 1946. The mosquitoes of the southern United States east of Oklahoma and Texas. Amer. Midl. Nat. Monogr. 3, 292 pp. and W. J. La Casse. 1955. Mosquitoes of North America (north of Mexico). University of California Press, Berkeley and Los Angeles. vi + 360 pp. Dampf, A. 1943. Distribucion y ciclo anual de Uranotaenia syntheta Dyar & 382 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 Shannon en Mexico y descripcion del hipopigio masculino (Insecta, Diptera). Rev. Soc. Mex. Hist. Nat. 4:147-169. Dyar, H. G. and R. C. Shannon. 1925. The types of Philippine mosquitoes described by Ludlow and other notes on the fauna. (Diptera, Culicidae). Insec. Inscit. Menst. 13:66—89. Edwards, F. W. 1920. The nomenclature of the parts of the male hypopygium of Diptera Nematocera, with special reference to mosquitoes. Ann. Trop. Med. Parasit. 14:23—40. 1932. Genera Insectorum. Diptera. Fam. Culicidae. Fascicle 194. Belgium. 258 pp. 1941. Mosquitoes of the Ethiopian Region. III. Culicine adults and pupae. British Museum (Natural History), London. 499 pp. Freeborn, S. B. 1924. The terminal abdominal structures of male mosquitoes. Amer. J. Hyg. 4:188—212. Galindo, P., F. S. Blanton, and E. L. Peyton. 1954. A revision of the Urano- taenia of Panama with notes on other American species of the genus. Ann. Ent. Soc. Amer. 47:107—177. King, W. V. and H. Hoogstraal. 1946(1947). New species of New Guinea Uranotaenia of the tibialis group (Diptera: Culicidae). Ann. Ent. Soc. Amer. 39:585-596. La Casse, W. J. and S. Yamaguti. 1950. Mosquito fauna of Japan and Korea. Off. Surg., 8th U.S. Army, Kyoto, Honshu. 268 pp. Lane, J. 1943. Sobre o género Uranotaenia (Diptera, Culicidae, Culicini). Rev. Ent., Rio de J. 14:137-161. 1953. Neotropical Culicidae. Sao Paulo, Brazil. 1112 pp. Matheson, R. 1944. Handbook of the mosquitoes of North America. 2nd ed. Ithaca, N.Y. 314 pp. Peters, W. 1963a. Mosquitoes of New Guinea (Diptera: Culicidae). Part VII. A new species complex of Uranotaenia Lynch Arribalzaga, 1891. Proc. R. Ent. Soc. Lond. (B) 32:135-144. 1963b. Mosquitoes of New Guinea (Diptera: Culicidae). Part VIII. New species of the genus Uranotaenia Lynch Arribalzaga, 1891. Proc. R. Ent. Soc. Lond. (B) 32:203—209. 1964. Mosquitoes of New Guinea (Diptera: Culicidae). Part IX. Further new species of Uranotaenia Lynch Arribalzaga and notes on the genus. Proc. R. Ent. Soc. Lond. (B) 33:19-29. Pratt, H. D. 1946. The genus Uranotaenia Lynch Arribalzaga in Puerto Rico. Ann. Ent. Soc. Amer. 39:576—584. Yamaguti, S. and W. J. La Casse. 1951. Mosquito fauna of North America. Part II. Genera Megarhinus, Wyeomyia, Uranotaenia, and Culiseta. Off. Surg., Hq., Logistical Command, APO 343. 85 pp. eEEeEeEeEEEEeEeEeEeEeEeEeEeEeEeEEyEyEEEE——y PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 383 A NEW LOCALITY RECORD FOR IXODES WOODI BISHOPP’*” (ACARNIA: IxODIDAE) During a survey of ticks occurring in Alabama that extended from the spring of 1963 through the fall of 1966, one engorged female, of Ixodes woodi Bishopp was collected from a woodrat during the month of March. This specimen was referred to Dr. Glen M. Kohls of the U. S. Public Health Service, Rocky Mountain Laboratory, for confirmation of identification. It appears to represent the first record of the occurrence of I. woodi in this State or even in the Southeast. Accord- ing to the records of Kohls (personal communication ), I. woodi has only been re- ported from Texas, Indiana, Colorado, and California. Practically all of these collections were associated with woodrats. The single female specimen of I. woodi was taken on an eastern woodrat from a cave seven miles north of Flat Rock, Alabama, in Jackson County. Jackson County is in the extreme northeast corner of the State. Two specimens of Dermacentor variabilis (Say ) were taken from the same animal. Two subspecies of the eastern woodrat, Neotoma floridana (Ord.), occur in Alabama: N. floridana illinoensis How. and N. floridana magister Baird. The female of I. woodi was taken on the latter subspecies. Seventeen specimens of N. floridana magister were collected and six of N. floridana illinoensis; these 23 woodrats, however, yielded only one specimen of I. woodi. Of interest is the fact that the one specimen collected was taken from a woodrat trapped in a cave. —JosEpH C. Cooney and Kirpy L. Hays, Department of Zoology-Entomology, Auburn University Agricultural Experiment Station, Auburn, Alabama 36830. 1 This investigation was supported, in part, by Public Health Research Grant AI 04904 from the Institute of Allergy and Infectious Diseases. 2Immediate publication secured by full payment of page charges—Editor. NOTICE TO CONTRIBUTORS Effective July 1, 1968, all manuscripts submitted for publication in the Proceedings, including notes of one page or less, must be accompanied by an abstract suitable for publication. The abstract must be typed on a separate sheet, should be brief (not more than 3% of the original), and written in whole sentences, not telegraphic phrases. Abstracts are not printed with notes of one page or less but will be sent to Biological Abstracts when the notes are pub- lished. Therefore, precede the abstract for notes with a paragraph giving the author’s name, his affiliation in parentheses, the title of the article, and the abbreviated name of the journal. These abstracts will be printed beginning with the March, 1969, issue of the Proceedings. 384 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 A NEW MOSQUITO RECORD FOR THE UNITED STATES (DireTERA: CULICIDAE) A single male of Culex (Aedinus) latisquama (Coquillett) from Lee County, Florida, was recently found in the United States National Museum. The hand- written label is “Estero, Fla. J. B. Van Duzee.” This specimen was undated, but Junius B. Van Duzee collected at Estero from April to July of 1906. He found Wyeomyia mitchelli (Theobald) (= W. antoinetta Dyar and Knab), W. van- duzeei Dyar and Knab, and Culex pilosus (Dyar and Knab) (= C. floridanus Dyar and Knab). Estero is the type-locality for vanduzeei and the two synonymous species. Culex latisquama was described from Port Limon, Costa Rica, and has also been found in Panama, Honduras, and Surinam. It is the type species of the subgenus Tinolestes Coquillett, a synonym of Aedinus Bourroul, and it is the first record of the subgenus in the United States. The mosquito is small and similar in appearance to species of the subgenus Melanoconion, but the male palpus is only about half as long as the proboscis, segment 4 being reduced and segment 5 being minute. The terminalia of this specimen agrees very well with those of specimens from Panama. The species breeds in crabholes, and a search for larvae in crabholes in the vicinity of Estero might rediscover the species as well as extend the range of Deinocerites cancer Theobald.—ALan STONE, Systematic Entomology Laboratory, Entomology Research Division, ARS, USDA, c/o U.S. National Museum, Washington, D. C. 20560. PROC. ENT. SOC. WASH., VOL. 70, No. 4, DECEMBER, 1968 385 INDEX TO VOLUME 70 Acanthoscelides, key to obtectus group, 4: obreptus, 5; obsoletus, 9; obtectus, 7; obvelatus, 7; pyrrhomelas, 319 Acrophasmus, key to Nearctic species, 101; arizonensis, 102; butleri, 103; ferrugineus, 103; lyeti, 105; secun- dus, 106 Aedes albopictus, neotype designation, 297; aloponotum, communis, taxo- nomic characters and_ distribution, 133 Alethaxius dominicae, 61 Alysia eariosa, 209; ridibunda, host and present status, 204 Amoebaleria sackeni, synonymy, 113 Amynothrips, 175; andersoni, 179 Aphids, Plummers Island records, 84 Apis mellifera mellifera, colonial be- havior, 303 Argia plana, demography, 197 Atta, Panamanian species, key, 348 Balaustium putmani, 13 Baldus tripasci, 88 Balocha bicolor bicolor, 99; bicolor uniformis, 100 Blissinae, Neotropical, new genera and species, 42 Blissus antillus, slateri, 150 Brachytremella, 114; cartwrighti, 121; crossi, 117; joanae, 114 Burrower bugs, Galapagos Islands, 192 152; planus, 151; Camponotus (Myrmentoma) sayi, 355; (Tanaemyrmex ) festinatus, 350 Cantacaderinae, systematics and mor- phology, 245 Carldrakeana, 250; list of species, 251 Caulocampus, key to species, 126; aceri- caulis, 126; matthewsi, 128 Caveloblissus, 42; americanus, 45 Ceratosmicra, key to North American species, 170; campoplegicis, 172; debilis, 174; immaculata, 174; me- teori, 174; paya, 174; provancheri, 171 Chomatophilus, 324; aphanistes, 326; smithi, 328 Chrybaneura, 33; harrisoni, 34 Chrysomelidae, of Dominica & Jamaica, 60 Coccinellidae, dipterous eggs as food, 142 Corioxenos antestiae, 240 Cryptognathus barrasi, 313 Culex (Acalleomyia) obscurus, scription, 53; (Aedinus) latisquama, new record, 384 Culicoides, breeding places in Panama, 132 Culiseta, in Connecticut, 187 Cyclotynaspis, 246; acalyptoides, 246 rede- Dallasiellus murinus, 192 Damselfly, demography, 197 Deltocephalinae, from Puerto Rico, 35 Diarthrophallus quercus, 193 Distocader, 249; list of species, 250 Doryctinus rugulosus, 112 Elenchidae, of Dominica, 238 Erythraeoidea, new genus and species, 13 Eulabidocarpus, 85 Euthycera mira, 81 Euxoa, lectotypes selected some species of J. B. Smith, 263 Galerucella bervivittata, 62 Glyptocolastes, key to Nearctic species, 108; caryae, 108; texanus, 110 Gonycentrum, 246; list of species, 248 Grylloprociphilus, 57; frosti, 57 Habrophora thelmae, 60 Hadropoda xanthoura, 63 Heikertingerella wirthi, 65 Heteroblissus anomilis, 48 Hexagenia, in Mexico, 358 Hodgesia, note on male proctiger, 376 Homoschema dominicae, 67 Hoplisoides carinatus, 287; cazieri, 288; floridicus, 289; glabratus, 291; projectus, 290 Hoplocheylus pieckardi, 307 Hybomitra, new synonymy, 302 Hylemya variata and variabilis, note on identity, 346 386 PROC. ENT. SOC. WASH., VOL. 70, NO. 4, DECEMBER, 1968 > > > Idiocerinae leafhoppers, 97 Ixodes woodi, new record, 383 Krameraxus, 35; leucornatus, 37 Lactica flinti, 64 Lygaeus slateri, 75 Marcocephalus (Lophoscutus) spieuli- ger, 243 Metachroma gagnei, 62 Micracanthia sechuhi, 165 Microchrysa, key to North American larvae, 148; flavicornis, 148 Microplatymerus, of Seychelles, 27 Microvelia ashlocki, 129 Minettia obscura, partially reduplicated antenna, 360 Minitingis, 251; list of species, 253; elsae, 253 Mites, new species from pine, 307 Molothrognathus rosei, 309 Neoacarus minimus, 71; occidentalis, 69 Neophyllobius lorioi, 309 Neoraphignathus howei, 315 Nesogeophilus, true identity, 345 Oedionychus gurneyi, 63 Pachydiplosis oryzae, authorship, 136 Paraeupalopsellus hodgesi, 315 Parakosa, key to species, 86; rectipes, 86 Paraphanolophus, 20; metealfei, 20 Paratoxurinae, new subfamily name, 147 Passalobia, 38; dubinerae, 38 Patritodemus delicatus, 50 Pedioscopus balochoides, 97 Pentacora sphacelata, 21 Philanthus gibbosus, sleeping behavior, 10 Pimeliaphilus trogadermus, 1 Piophila (Liopiophila) nigricoxa, 27; nigrimana, 25 Podothrips, new name combination, 125 Polyergus, male genitalia and taxonomy, 156 Praetorblissus gradus, 47 Pseudelenchus spangleri, 239 Pteromicra rudis, 78 Reticulatodemus orbiculoides, 49 Rhinoecius bisetosus, 143; 143 Rhynonirmus kingi, 184 cavannus, Saldula galapagosana, 22; sectilis, 22 Secutobruchus, 280; ceratioborus, 283; gastoi, 285 Sennius willei, 318 Sidfaya philtata, 65 Sinalda, 248; list of species, 249; haplo- taxis, 249 Snellenius, 339; key to species, 340; atratus, 342; bicolor, 342; peruensis, 343; tricolor, 341 Sogonidae, key to genera, 330; cata- logue of genera, 330 Somatia, key to species, 256; papaveroi, 255; sehildi, 256; sophiston, 256 Sphaerolophus canadensis, 15 Stachiella retusa martis, host, 191 Stenocorse, 110; bruchivora, 111 Strepsiptera, of Dominica, 238 Stygomomonia, key to North American species, 213; (Allomomonia), 211; (Neomomonia), 211; moodyi, 213; neomexicana, 224; occidentalis, 221; ozarkensis, 219; pallida, 220; plana, 220; riparia, 215; separata, 223 Telamona arechboldi, 154 Tonnoiria, priority of names, 87 Tripteroides denticulatus, 362; malayi, 368; tarsalis, 372 Tuoba, true identity, 345 Turacoeca bedfordi, 260; leucotis, 262; scleroderma, 258; subrotunda, 261 Typopsilopa, 225; key to species of Western Hemisphere, 227; anten- nalis, 231; arehboldi, 236; arnaudi, 231; atra, 228; imea, 237; manni, 236; nigra, 234 Uranotaenia, note on male _proctiger, 376 Vespula vulgaris, maculifrons, life habits, 332 Volsellacarus sabulonus, 73 Xestoblatta, notes on Neotropical spe- cies, 137; eantralli, 137 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Information for Contributors ‘Publication (without charge) in the Proceedings is reserved for members only. However, immediate publication of papers by members (as well as non-members ) may be obtained after acceptance at a cost to the author of $15.00 per printed page. Regular papers are published in approximately the order that they are received. Manuscripts should not exceed 30 typewritten pages including illustra- tions. Papers of less than a printed page may be published as space is available at the end of longer articles. Manuscripts for publication, proof and other editorial matters should be addressed to the Editor (for address, see inside front cover of this issue). Typing—All manuscripts must be typed on bond paper with double-spacing and ample margins. Carbon copies or copies on paper larger than 8% x 11 inches are not acceptable. Do not use all capitals for any purpose. Underscore only where italics are intended in the body of the text, not in headings. Number all pages consecutively. References to footnotes in the text should be numbered consecutively and typed on a separate sheet. First page—The page preceding the text of the manuscript should include (1) the complete title, (2) the order and family in parentheses, (3) the author's name or names, (4) the institution with city, state and zip code or the author's home city, state and zip code if not affiliated, (5) in the upper left hand corner, the complete name and address to which proof is to be sent. Names and descriptions of organisms—The first mention of a plant or animal should include the full scientific name with the author of a zoological name not abbreviated. Descriptions of taxa should be in telegraphic style. References—Citations in the text of papers longer than one printed page should be by author and date and should refer to a list of concluding REFERENCEs listed alphabetically in the following format: Muesebeck, C. F. W. 1963. Host relationships of the Euphorini (Hyme- noptera: Braconidae). Proc. Ent. Soc. Wash. 65(4): 306. and L. M. Walkley. 1951. in Muesebeck et al., Hymenoptera of America North of Mexico, Synoptic Catalog, U. S. Dept. Agr., Agr. Monogr. 2: 90-184. In shorter articles, references to literature should be included in parentheses in the text. Tables—Tables are expensive and should be kept to an absolute minimum. In most cases, material in tables can be incorporated into the text. When tables are necessary, each table should be prepared as a line drawing or typed on a separate page with heading at top and footnotes below. Number footnotes consecutively for each table. Use only horizontal rules. Illustrations—No extra charge is made for line drawings or halftones. Authors must plan their illustrations for reduction to the dimensions of the printed page and the individual figures must be mounted on suitable board. Proportions of full-page illustrations should closely approximate 454g X 6” (26 X 36 picas); this usually allows explanatory matter to appear on the same page. On the back of each illustration should be stated (1) the title of the paper, (2) the author's complete name and address, and (3) the number of the illustration such as “No. 1 (of 3)” etc. Figures should be numbered consecutively. Figure legends—Legends should be typewritten double-spaced on separate pages headed ExpLANATION OF FicurEs and placed following REFERENCEs. Do not attach legends to illustrations. . Proofs, reprints and abstract—Proofs and forms for the abstract and reprint order will be sent to the authors by the printer with explicit instructions for their return. Major changes in proof will be charged to the author. CONTENTS (Continued from front cover) KINGSOLVER, J. M.—One previously described and one new species of South American Bruchidae injurious to commercial legume seed crops (Cole- optera ) McCAFFERTY, W. P.—The mayfly genus Hexagenia in Mexico (Ephemer- optera: Ephemeridae ) PECHUMAN, L. L. and A. STONE—A new synonymy in Hybomitra (Diptera: Tabanidae ) PEYTON, E. L. and R. H. HOCHMAN—A revised interpretation of the proctiger of male Uranotaenia with a related note on Hodgesia (Diptera: Culicidae ) SHENEFELT, R. D.—Snellenius in the Neotropical Region (Hymenoptera: Braconidae: Microgasterini) SMILEY, R. L. and J. C. MOSER—New species of mites from pine (Acarina: Tarsochelidae, Eupalopsellidae, Caligonellidae, Cryptognathidae, Raphi- gnathidae, and Neophyllobiidae ) SNELLING, R. R.—Studies on California ants. 4. Two species of Camponotus (Hymenoptera: Formicidae) STEYSKAL, G. C.—A case of partially reduplicated antenna in Minettia obscura (Loew) (Diptera: Lauxaniidae ) STONE, A.—A new mosquito record for the United States (Diptera: Culici- 318 358 “Fe 9 af el Oo : i Pat e & aa ea. rf ages 2, ly ss [7