Historic, archived document Do not assume content reflects current scientific knowledge, policies, or practices. Ds, DEPARTMENT OF AGRICULTURE. BOTANICAL DIVISION. SO ree ENE AN oO Ss. ln tC: @) il) Or SOME OF THE WORK OF THE DIVISION, INCLUDING EXTRACTS FROM CORRESPONDENCE AND OTHER COMMUNICATIONS. PREPARED BY Dr. GEO. VASEY AND B. T. GALLOWAY, - UNDER THE DIRECTION OF THE COMMISSIONER OF AGRICULTURE. WASHINGTON: GOVERNMENT PRINTING OFFICE. 1889. 11244—No. 8——1 | LETTER OF SUBMITTAL. Str: We have the honor of presenting a bulletin representing, in part, the work of this Division, which we trust will be received as a con- tribution of some value in advancing the mutual helpfulness and inter- dependence which should exist between the Department of Agriculture and the workers and thinkers in the line of agricultural and botanical science. Respectfully, GEO. VASEY. B. T. GALLOWAY. Hon. NoRMAN J. COLMAN, Commissioner of Agriculture. INTRODUCTION. - In the progress of the work of the botanical division many facts and observations are brought to light which hardly seem appropriate to be incorporated in the annual reports, but which will be of interest to many readers, and which will surely be welcomed in some form by special in- vestigators in botanical science. In the great work which the Government has undertaken, of furnish- ing the means of an advanced practical education through the agricult- ural colleges and experiment stations, new appliances and new facil- ities for instruction are becoming a necessity. It has seemed to the officers of this division that much valuable material can be furnished in this direction by the occasional issuance of a bulletin especially adapted to help forward the line of botanical work for teachers and students. In the hope that this may be the result, the present paper is dedices to the agricultural colleges of the country. pes. i te des al eraadl Ce BOTANICAL DIV FSlON: PREPARED BY DR. GEO. VASEY. ee BOTANICAL DIVISION. Prepared by Dr. GrorGr VASEY. GRASS EXPERIMENT STATIONS. For several years past the Department of Agriculture has been making special investigation of the grasses of the West, particularly those of the arid districts, with a view of ascertaining what are the prevailing or prominent species, what is their range of distribution as to locality and soil, what their value for grazing purposes, and what kinds offer a_ reasonable prospect of being valuable for culiivation. Great interest has been felt in this subject both by farmers and cattlemen, and Gov- ernment aid has been invoked for the purpose of conducting suitable experiments, and such aid has been strongly recommended by the Com- missioner of Agriculture. In response to these requests, Congress, a few months ago, made a moderate appropriation for the purpose of establishing grass experiment | stations. It was determined that one of these stations should be located. west of the 100th meridian. Commissioner Colman immediately ap- pointed Dr. Vasey, botanist of the Department, to proceed to the West to select a location. After a careful investigation he made a selection at Garden City, in southwestern Kansas, where a public spirited citi- zen made a free lease to the Government, for five years, of 160 acres of land on the high prairie two miles north of the city. Thisis so located that irrigation can be employed on such portion of the land as may be desirable. .The chief object of the station is to experiment with grasses and forage plants, both native and foreign—with any kinds, in fact, which give promise of utility and adaptation to the climatic conditions of the arid piains, and furnish a substitute for the scanty pasturage now existing. This isa great necessity. In order to secure the perma- nent settlement of this portion of country, it should be ascertained what can be depended upon in the occupation and cultivation of the dry uplands where irrigation can not be applied. It is a vital question, af- fecting the interests of thousands of settlers over a great extent of country. | Upon the location of the station the Commissioner appointed Prof. J. A. Sewall, of Denver, as superintendent of the station; and he has up to the present time had 80 acres fenced, 40 acres plowed, a quan- 9 1G Gs tity of grass sods transplanted into prepared ground; has put up a small seed-house, and made everything in readiness for more exten- sive work next spring. The results of these experiments will be watched with great interest; but definite results can not be expected without the allowance of a considerable period of time. The need of some grasses suitable to the climate of the dry region lying mainly west of the 100th meridian has been long felt and ae- knowledged. It is estimated that there are in eastern Colorado, west- ern Kansas, western Nebraska, and southern Wyoming 120,000 square miles, or over 76,900,000 acres, of this arid, elevated country. This region was known to be covered principally by short grasses called “ buffalo” and ‘ mesquit,” in some places thickly covered, in others more and more sparsely, while in some portions the amount of grass was exceedingly small. During the greater part of the last twenty- five years this region has been occupied for cattle and sheep ranches, for which purpose alone it was thought to be adapted. But even for this purpose its ability to support the domestic grazing animals was very small compared with its vast extent. This arises partly from the fact that only such portions as were within reasonable distance from water could be utilized, and partly from the scanty supply of grass. The estimates as to the supporting capacity of these plains vary much according to localities, some statements being that 40 to 50 acres are re- quired to support one animal, others that 20 or 30 acres are sufficient, and yet others that 10 to 15 acres are enough. All these estimates are correct as to certain localities, and it may be considered probable that over the entire region the average amount re- quired would be 15 acres to the head of stock. This, however, was under the wasteful ranch system. A limitation of the range and the size of the herds, so that they could be properly cared for as to water, shelter, and winter food, would greatly increase the grazing capacity of the country. But during the past five or six years there has been such an over- whelming influx of settlers into that district that the ranchmen have been nearly all forced to abandon the country in favor of homesteaders, who are seeking to make farms of their quarter and half-sections. A series of favorable seasons had occurred, during which the few scat- tered settlers in the country had been successful in their agricultural efforts, and then began a rush of immigration which soon absorbed a great portion of the most desirable locations. The settlers were full of the most sanguine expectations, and an army of boomers invaded the country, located town sites and city sites, and a general inflation of values occurred. A few well located towns will perhaps retain their marvelous development, but many others have come to a stand-still, and many will retrograde, for the reason that numerous thriving towns can not exist in an agricultural country without the improvement and successful cultivation of the surrounding land, and to this end the resi- dents of that region must now direct their efforts. ig Sufficient time has not yet elapsed to determine what will be the ulti- mate success of general agriculture in this section, but there can be no doubt that the country is eminently adapted to pastoral uses, and the settlers would do well to bestow their attention largely on stock-raising and dairying. But it is evident that in order to make this business profitable for small farmers or men of limited acres, some means must be devised for securing an increased production of grass upon the land. This is the present most important problem for the arid districts. It is asserted that the commonly cultivated grasses will not be successful in these arid lands except where irrigation can be applied. The native grasses of the region, which are mostly “ buffalo grass” and “ grama grass,” are acknowledged to be very nutritious, but the yield is so hgnt as to require a large area for cattle to range over to obtain support. The inquiry naturally occurs, “Can not some grasses be obtained which will not only endure the aridity of the climate, but also make a more vigorous growth and a more abundant production ?” It may be safely laid down as a principle that no great improvement in this respect can be expected without a cultivation of the soil. With this even the buffalo and grama grasses may be expected to double their production. Theinquiry is sometimes made, ‘“* Which of our native grasses might be reasonably expected to be adapted to cultivation in the said region ?” We have already stated that the “ buffalo grass” (Buchloe dactyloides) and ‘“‘grama grass” (Bouteloua oligostachya) were the prevailing na- tive species on the prairie or upland districts. But there are others. which deserve consideration, and some of them may be found to pos’ sess the qualities desired. Andropogon provincialis, commonly called blue-joint, is a grass. of wide range, being found from Massachusetts to Texas, and to the Rocky Mountain region. It is also, sometimes, but improperly, called * blue- stem” and “blue-grass.” Even the name “ blue-joint” is inappropriate, for the joints are rarely if ever blue, and the most conspicuous color about the grass is its purplish top, by which it may be distinguished at a great distance. It commonly grows from 4 to 6 feet high, rather sparsely on the ground, but with an abundance of long leaves, so that it produces a large crop of hay. It was formerly plentiful on the prai- ries of Illinois and Iowa, and still occurs largely in most of the states west of the Mississippi, and is universally esteemed and cut for hay in its wild state. It thrives best on rich bottom-land, not too moist nor too dry, but is capable of considerable variety in that respect. It is well known all over Kansas, being abundant in the eastern part of the siate, and the general testimony is that it has been spreading westward since the settlement of the country, and its extension in the West is always regarded with pleasure. It starts growth late in the season, and is chiefly valuable for summer and fall grazing, and for hay-mak- ing. It is said that close pasturing for a few years kills it out. 12 2. Andropogon Hallii.—This species much resembles the preceding, but the culms are stouter, the leaves thicker and more succulent, the flower spikes are larger, and the flowers more hairy. It prevails in very sandy soil, and is most frequent in western Kansas and Colo- rado, alsoin Nebraska and northward along the Missouri River. The leaves and stem are commonly of alight bluish-green color. This will provably be well adapted to light sandy soils. 3. Andropogon scoparius.—This is very common on prairies, and is sometimes called short or small blue-joint, sometimes bunch-grass, and sometimes wire-grass. It grows in dense tufts cr bunches, both on low land andin thin soils on bluffs and hills. It is in most places a constituent of prairie hay, and if cut early makes good feed. 4. Stipa spartea, called porcupine grass, arrow-grass, and devil’s knit- ting-needles, from the long, stiff, twisted awns inclosing the seed. These awns are said to be injurious to stock, and particularly to sheep, as their sharp points penetrate the wool and sometimes the flesh. But the seeds ripen early (about the 1st of July, according to Professor Crozier), and during the rest of the season the grass is well marked by the tall culms bearing the bleached, empty, oat-like panicles. But the long root-leaves continue to grow and furnish valuable feed until late in the season. It should receive attention in any experiments for a past- ure grass. It is the buffalo grass of the Saskatchawan region. 5. Sporobolus heterolepis, called bunch-grass, and wire-grass, from the abundant, long, wiry leaves and stems. I found it a considerable ele- ment in the prairies of southern Dakota, and it occurs southward to Texas. It was also common on the prairies of [linois and Wisconsin before the incoming of settlements. West of the 100th meridian, how- ever, especially in sandy soils, this species is replaced by two others of the same genus, viz: Sporobolus cryptandrus and S. airoides. All these species should receive attention. 6. Deyeuxia Canadensis, frequently called red-top, and sometimes also blue-joint. Professor Crozier, who spent some time in northwestern Iowa and adjacent parts of Minnesota and Dakota, in studying the na- tive grasses, says: ‘*This is considered by some to produce the best hay, for cattle, of all the native grasses. It is very leafy and stands re- markably thick upon the ground. The seed ripens early in July, but the leaves remain green until winter. It is probably hardly equal to some of the upland grasses in quality, but it gives a larger yield, and is undoubtedly worthy of cultivation. It is usually found upon the margins of ponds; it will thrive, however, on land that is only slightly moist, and often occurs along the banks of roadside ditches. On rather low land which has been broken and allowed to go back it frequently comes in, and after a few years occupies the land to the exclusion of all other vegetation.” This species, athough promising for moist land, would perhaps fai: on arid soil, but there are other species of the genus prevailing in the 13 mountain region, which are accustomed to dry soil, and may be found serviceable for cultivation, particularly such as D. confims and D. syl- vatica. : 7. Ammophila longifolia, a tall, coarse-stemmed grass occurring in very sandy districts, or on sandy river-banks, is in some localities a useful grass as aresort for cattle late in the autumn, and in winter it is said to save many range cattle from starvation. Its long, creeping root-stocks penetrate deeply into the soil and give it good staying qual- ities, for which reason it may be valuable in some localities. 8. Oryzospis cuspidata.—This is one of the so-called bunch-grasses, which occurs near the mountains, ia sandy soil, and has been called indian millet. It will grow in very dry, sandy soil, furnishes a con- siderable foliage, and large, spreading panicles of very nutritious seeds. It is said that cattle keep fat upon the grass in the mountain 4dis- tricts. 9. Panicum virgatum, sometimes called switch grass and sometimes wild red-top. It occurs mostly in low or moist ground, usually near streams, and varies in height from 2 to 4 feet, with long leaves and a wide, spreading panicle. Professor Crozier says it yields two to four tons peracre, and should be cut early to be of good quality. Its value has been recognized in some parts of Colorado, and around Greeley it has been considerably cultivated, with satisfactory resulf. It should receive a fair trial in the experiments. 10. Another perennial species, Panicum obtusum, occurs in some parts of Colorado and New Mexico which ought to receive attention. In. New Mexico and Texas if is called vine mesquit and running mesquit, from its habit of throwing out long runners, sometimes 6 to 10 feet long, which at intervals form thickened, woolly joints, which sometimes take root. I found patches of it growing in southern Colorado, making an even, grassy surface, and appearing as if it would cut two tons per acre. T also saw it occurring in considerable quantity in hay brought into the Pueblo market. It deserves attention with reference to its agricult- ural value. 11. Chrysopogon nutans is related to the Andropogons, particularly to the tall A. provincialis. It often grows 6fect high, and has a drooping panicle somewhat resembling oats, and hence, has been called wild oats. It is of frequent occurrence on the prairies and isa nutritious grass, but should be cut early, as at full maturity the stems are coarse and are rejected by cattle. 12. Agropyrum glaucum.—This is known by different names, as Colo- rado blue-stem, blue-grass, wheat-grass, wild quack-grass, and gumbo- grass. It is closely related to the quack-grass or couch-grass, so com- mon and so much dreaded by some in the older-settled parts of the ccuntry. It has a stiff stem and leaves, which are usually of a bluish- green color. On hard, dry soil its growth is low and sparse, only here 14 and there a scattered stalk with a flower spike, somewhat like a starved, beardless head of wheat; but in low, moist ground it often grows with great vigor 2 or 3 feet high, and is considered valuable for hay. On the borders of ditches and on irrigated ground it yields a heavy cut- ting. It has great persistence in the ground on account of its strong, running root-stock. Whether it will, in cultivated ground become as (lifficult to eradicate as its Eastern namesake can not now be predicted, but when a persistent, nutritious grass is the great want of a country it is worth while to take some risks. 13. Agropyrum tenerum.—Another species of this family is found in Colorado, western Kansas, and Nebraska, which may prove valuable for some qualities. It has a narrow, slim spike of flowers and stiff culms. It does not have running root-stocks, but grows in clumps of variable size. It furnishes a large quantity of foliage and matures early, and for these qualities it may be desirable to cultivate it in a mixt- ure of grasses for pasture. It is common on banks of ditches and on the waste grounds in the city of Denver. 14. Several species of Elymus, particularly Elynius canadensis and E. virginicus, are common and known by the name of rye-grass. They are considered nutritious grasses, and are commonly cut for hay. They are coarse and probably would not bear pasturage, but may be worth cultivating for forage. The above are the principal native perennial grasses which have oe- curred to me as having promise of utility for the arid districts which are now under consideration. Other species will come in for attention, particularly some whose range is more restricted to the neighborhood of the mountains. I have already stated that the grama-grass should be placed under cultivation, not only the most common species, Bouteloua oligostachya, but others, as the B. racemosa. -There are also several strong-growing species in Mexico which shouid be tried. The tall-growing buneh- grasses of Arizona might also be experimented with. These include several species of Sporobolus and Epicampes. Panicum bulbosum, a vigorous grass of Arizona and New Mexico, hay- ing thickened, bulbous root-stalks, should be tried, and if it would en- dure the winters would probably be a valuable pasture grass. There are also several species of Poa belonging to the mountain district which might prove valuable in cultivation, as Poa tenuifolia, P. undina, P. nemoralis, and others. There are also some native species of Festuca which merit attention, such as Festuca scabrella, and several large forms of Festuca ovina. Also several species of Bromus. Some attention must also be given to annual grasses suitable for for- age and hay. The necessary supply of hay for winter feeding of cattle may sometimes be more profitably procured from the annual grasses, as millet, Hungarian grass, and the various forms of sorghum. In this 15 class of annual grasses there is a wider field for selection than in the perennials, and they are obtainable from our own and other countries. There are also many fodder plants, other than grasses, which need eareful trial. The family of leguminous plants furnishes a large num- ber of these valuable species, such as the clovers, alfalfa, medicago, meli- lotus, sainfoin or esparsett, vetch, peas, lotus, lupines, serradella, ete. There are also some native forage plants which deserve experiment. We have several species of Lupinus in the Rocky Mountain region which are vigorous growers and of large size, apparently more productive than the usually cultivated foreign ones. Of Vicia and Lathyrus we have also promising native kinds. Some of the native species of Airi- plex are well known to furnish valuable winter forage for cattle, and it may be well to ascertain what will be their value under cultivation, especially for unusually sterile or sandy soil. Attention should be given to a combination of grasses for pastures, some of which should be of early growth and some of later, so as to se- cure a succession of feed during the entire season. The judicious blend- ing of five or six species may be made to accomplish this result. By using such a mixture the ground may be more uniformly covered and there will be a better succession of tender foliage. Some recent experiments at the Missouri Agricultural College fully bear out this conclusion. A 50-acre lot was sowed with five varieties of pasture grass and three of clovers. Upon this 50 acres fifty-two head of cattle grazed throughout the season, without making use of an adjacent pasture, a part of which contained excellent blue-grass, except at long periods, and then only cursorily passing over it. In England great attention has been given to combining several kinds of grasses in meadows and pastures, and it is claimed that the practice is better for the land, and gives a larger yield than when only one variety is employed. In any plan for experiments with grasses for the purpose above indi- cated the element of time must receive large consideration. It must be remembered that the seeds of wild species have first to be obtained in small quantities and to be collected by hand, and that two years will be consumed in getting fairly started. Having once obtained preliminary beds from which a stock of seed may be easily obtained, the trials can be made on a larger scale. Sev- eral acres of each kind of grass or fodder plant should be secured, with careful note of their relative merits as to quality and yield. These experiments can not be completely satisfactory without ascer- taining the conduct of the grasses in actual use as pastures, for it is well known that some grasses will bear the tramping and grazing of cattle, while others will not. The least attempt at a series of experiments, therefore, should cover a period of five years, and there is great reason to expect that they may be profitably continued for twice that time, oo 16 Experiments of this kind are difficult and expensive, and can not well be made by private individuals. The great extent of country interested in, and needing such work renders it highly expedient that the Govern- ment should undertake it. It is expected that another station for the trial of new grasses will be located next spring, probably in one of the Southern States. NOTES ON GRASSES. On the south bank of the Arkansas River, at Garden City, in south- western Kansas, there is a long range of sand hills or sand bluffs. The citizens of the place say that a few years ago these bluffs were destitute of vegetation, and the loose, white sand was blown about in clouds by ° the strong prevailing winds, but that recently they have become cev- ered with grass. I crossed the river to ascertain what were the grasses that had secured a foothold in the shifting sand beds, and found that they are now sparsely covered principally with Adropogon Halltt and Redjieldia flecuosa. The first grass is aclose relative of A. provincialis, but with stouter culms, shorter and more succuient leaves, larger flow- ers, and the whole plant of a glaucous-green color. Its roots are strong and penetrate deeply in the sand to reach moisture. On sandy ridges and prairies it often takes the place of A. provincialis, and is a promising grass for such situations. The other grass, Redfieldia flexu- osa, formerly called Graphephorum flexuosum, has not been collected by botanists for many years, its known localities being very few. It was therefore a great pleasure to rediscover it here. Its roundish, eylin- drical leaves, one and one half to two feet long, were sparsely scattered, and could with difficulty be pulled up from the long, deep, underground root-stocks, which evidently were very efficient in holding the sand in place. : Unfortunately there were few flowering culms visible, and they were immature and varied from the typical plant in having but one flower developed in the spikelets. Whether this variation is constant in this locality or is dependent upon the unusual dryness of the present sea- son can not now be determined, but is deserving of future investigation. Agropyrum glaucum and A. tenerum are both prevalent inthe dry dis- tricts of Kansas, Colorado, and Dakota, particularly where the native sod has been disturbed, as in old, abandoned wagon-roads and in the neighborhood of ditches. Both kinds are conspicuous on roadsides in the city of Denver, near the shallow irrigating ditches. A. tenerum grows in clumps and does not have running root-stocks. It is anearly grass, maturing in July, and afterwards is conspicuous by its whitened culms and leaves. In several places in Kansas, Colorado, and Dakota an crect, much. branched, bushy form of Muhlenbergia glomerata was observed. Its ap- 17 pearance of thrift and greenness has been noticed, and my attention was called to it by farmers and observant citizens. It would under cultivation produce a large amount of forage, and it will be given trial at the grass station. What are the common grasses of the prairies ? Near Wichita, Kans., the examination of a piece of native upland prairie showed Stipa spartea, Panicum virgatum, Keleria cristata, Andro- pogon provincialis, and Andropogon scoparius as the principal species. Here and there occurred a little of Bouteloua racemosa and B. oligosta- chya, with occasionally small patches of Buchle dactyloides. With these were mingled Amorpha canescens, Psoralea floribunda, Petalostemon viola- ceum, a Helianthus, Aplopappus rubiginosus, and Huphorbia marginata. On lower ground were Panicum virgatum of larger: growth, Agropyrum glaucum, Spartina cynosuroides, with Elymus virginicus and HL. canadensis. A ride of 20 miles over the prairies of Butler County showed that the principal upland grasses are the five species first mentioned. A very intelligent farmer said they would yield a cutting of 1 to 24 tons per acre. He had tried various tame grasses, but without much satisfac- tion, as they mostly winter-killed, and he doubted if he could obtain any grasses better than those of the virgin prairie. Old settlers say that buffalo grass used to be very abundant, but it has nearly disap- peared from this section. Blue-grass (Poa pratensis) is coming in, and he thinks it will take the country. In southwestern Minnesota and southern Dakota there yet re- main extensive stretches of primitive prairie, and a careful examina- tion showed that on the uplands the common grasses were mainly the ones mentioned as prevailing in eastern Kansas, viz: Stipa spartea, Andropogon provineialis, A. scoparius, Keleria cristata, together with patches of Sporobolus heterolepis and Bouteloua racemosa. On moister ground was Panicum virgatum, Hlymus canadensis, and in wet ground - Spartina cynosuroides. Wherever the ground has been broken and not cultivated, as on railroad embankmentsand roadsides, Agropyrum glau- cum and A. tenerum had taken possession. According to the observa- tion of Professor Crozier the native prairies of northwestern fowa are characterized by the same species as above mentioned. Having been familiar with the prairie vegetation in northern Illinois forty years ago, I was struck with the similarity, the same species being then prevail- ing ones in that region. The Stipa spartea is an early grass and ripens its seeds about the first of July. Later in the season it is recognized by the persistent bleached culms and empty glumes of the spreading panicle. But the radical tufts of leaves continue growing green and vigorous throughout the summer, frequently being 2 feet long. Al- though somewhat coarse they make very good hay. 11244—No, 8——2 18 BOTANICAL NOTES. There is great latitude in the popular use of the term “ mesquit grass.” On the plains of western Kansas, New Mexico, Colorado, and Nebraska it is generally applied to the prevailing grass of that region, viz, Bouteloua oligostachya, although the same grass is also called buf- falo grass, particularly northward. | A correspondent in Texas sends three grasses under the name of mes- quit, viz, Bouteloua texana, which he says is the common mesquit (this species seems to be mainly confined to Texas); (2) Hilaria cenchro- ides var. texana, which is running mesquit; and (3) Buchle dactyloides, which there is called curly mesquit. There are several other species of Bouteloua, common in New Mexico and Arizona, which are also gen- erally recognized as mesquit grasses. Strangely, however, in the South- ern States, according to Professor Phares, Holcus lanatus is sometimes ealled velvet mesquit grass, and he states that so far as has come to his knowledge nine tenths of all the so-called mesquit grass planted in the Southern States is this Holcus. Mr. Il. W. Anderson, of Great Falls, Mont., states that in that sec- tion Lygodesmia juncea is becoming a bad weed in cultivated ground. The somewhat related Huropean Chondrilla juncea is a very trouble- some weed in Maryland and Virginia, extremely difficult to eradicate. Tt was introduced at an early period in the settlement of the country. P.S. Clark, Hempstead, Tex., writes that Sesbania vesicaria has cov- ered the entire prairies this sammer, and caused the destruction of the grass. Considerable has been said recently in Florida agricultural papers about a grass called maiden cane, which there is considered a great nuisance. This grass is Panicum curtisti, or a variety of it. In swampy land it grows 3 or 4 feet high, with strong culms and coarse running reot-stocks and is almost invariably sterile. In dry, sandy fields it is lower in stature, and has a branching panicle. It has been particularly brought to notice by Mr. Simpson, of Manatee, and Professor Curtiss, of Jacksonville (not the Curtis for whom the species was named, Rev. M. A. Curtis, of North Carolina), has called it Simpson grass. Polish wheat, Triticum polonicum, is frequently sent from Montana and the Northwest, where it is partially naturalized under the name of wild-goose wheat and Montana rye. A notion prevails that the seed was dropped by wild geese. It has been sometimes cultivated, and is very hardy and productive, but not of very good quality. Mr. F. A. Swinden, Brownwood, Tex., writes that alfalfa (Medicago sativa) seems in that locality to do well without irrigation. He will plant 100 acres of it in the spring. He has an orchard of 400 acres of pecan trees (Carya olivaformis). R. Maitre, New Orleans, says that Panicum crus-galli delights in moist ditches or furrows, and has the appearance of thrifty oat-stalks, and it 19 is eagerly eaten by horses, mules, or cattle. Professor Phares states that in Louisiana, Mississippi, and some other States it is mowed annu- ally, some farmers asserting that they harvest four or five tons per acre. In one county in Mississippi hundreds of acres are annually mowed on single farms. The genuine Ranunculus repens, undoubtedly introduced from Europe, according to J. G. Fish, Whitney’s Point, N. Y., is becoming a danger- ous enemy to the farmer. When it once gets thoroughly rooted in the soil, and particularly in meadow lands, clover, timothy, red-top, and June grass are perfectly impotent to resist it. — W. H. Williams, Crook City, Dak., sends specimens of two grasses, (Agropyrum tenerum and Elymus canadensis) which he says are superior to any kinds he has seen. They are found mostly on land that has been broken and allowed to stand uncultivated. The growth is remarkably heavy and the hay is of superior quality. W. A. Sanders, Sanders P. O., California, sends a bunch of grass (Hpicampes rigens) cut from his pasture, and says it is closely grazed off wherever stock can get at it. He says it grows all summer without water, and under excessive dryness cures into hay and makes palatable, nutritious feed until spoiled (as all hay spoils) by autumn rains. It is our best native dry land grass. Dr. M. McKenzie, Centreville, Mo., sends specimens of Triodia ses- lerioides, a tall, showy grass, which he introduced into his pastures and finds that it is likely to become a pest. Mr. E. W. Reasoner, of Manatee, Fla., sends specimens of a grass, a species of Pennisetum, the seed from Northwest India, which he has had in cultivation, and which he says is very luxuriant and grows to the height of 6 or 7 feet, One plot 10 feet wide and 100 feet long aver- ages 64 feet high. It was planted in June, and part of it was cut twice. He has not yet determined whether it is annual or not, but considers it @ promising grass. Paspalum distichum.—Several species of Paspalum have received atten- tion in the South, as being useful pasture grasses and very durable from their creeping and rooting habit. Paspalum distichum is one of these species. It grows principally in low, moist ground. Its stems and culms are mostly prostrate and running, sending up here and there a few flower-bearing culms. It is found in the Southern States and Texas, thence to California. Farther south it is found in most tropical coun- tries. Mr. W. A. Sanders, of Fresno County, Cal., writes recently as follows: “Are you aware of the value of Paspalum distichum for seed- ing pond-holes that dry up or nearly so in autumn? Such ponds are usually spots of bare, stinking mud, but when well set to this grass will yield all the way up to 80 tons (in the green state) of autumn feed for stock, especially valuable for cows first, then follow with sheep till every vestige is devoured. Surely it has an immense food value in such places.” = 20 THE GENUS PANICUM IN THE UNITED STATES. This genus is one of great difficulty to the student, and the present monograph, it is hoped, will be of essential benefit in the prosecution of botanical investigation. PANICUM, Linn. Spikelets with one perfect terminal flower, and usually a second one which is male only or rudimentary. Outer empty glumes two, ene of which is smaller than the other, often very small, or rarely wanting, membranaceous. The glume of the perfect flower with its palet are usually of a coriaceous texture, and obtuse or obtusish. The second flower, when present, is membranaceous. Inflorescence spicate or pan- iculate. . SECTION 1.—Digitaria.—Panicles of several spikes or spike-like branches, which are digitate or approximate at the top of the cul; spikelets usually small and in alternate pairs along one side of the flat- tened rhachis, one sessile and one short-pedicelled; empty glumes two, or sometimes three, the lower one then being minute. 1. P. glabrum, Gaud. 2. P. sanguinale, Linn. 3. P. filiforme, Linn. 4, P. serotinum, 7rin. SEcTION II.—Trichachne.—Spikelets densely silky-hairy, or fringed with long hairs, sessile or short-pedicelled, in pairs or clustered along the rhachis of the numerous simple, erect, spikes or branches of the pan- icle, acute; lower glume very minute. 5. P. leucopheum, A. B. K. 6. P. lachnanthum, Torr. SECTION IIJ.—Subspicata.—Panicle linear or narrow, the branches short, sessile, erect or appressed, or the apex spicate. Branches densely flowered : 7. P. paspaloides, Pers. 8. P. Curtisil, Chap. 9. P. obtusum, H. B. K. 10. P. reticulatum, Torr. 11. P. Texanum, Buckl. Branches lax flowered: 12. P. Chapmani, Vasey. 13. P. Reverchoni, Vasey. 14. P. subspicatum, Vasey. 15. P. stenodes, Gris. : SECTION IV.—Platyphylla.—Panicle of few (three to six), simple, mostly sessile, spreading branches, the spikelets sessile in two rows on 21 one side of a flattened rhachis, as in Paspalum, but with a small lower glume. 16. P. platyphyllum, Munro. 17. PB. plantagineum, Zink. SECTION V.—Brachiaria. Branches erect, orsomewhat spreading, not appressed, mostly approximate and sessile. 18. P. prostratum, Lam. 19. P. cxespitosum, Sicz. 20. P. fasciculatum, Stwvz. 21. P. grossarium, Linn. SECTION VI.—Polystachya.—Branches of the panicle numerous, Spreading, single or subfasciculate, simple, or with some short branch- lets, lowering to the base (except in P. hians), spikelets sessile or very short pedicelled. . P. barbinode, 7rin. P. gymnocarpon, HUI. . repens, Linn. P. anceps, Mich. (some forms). .laxum, Swe. . P. hians, Hil. SECTION VII.—Hupanicum.—Branches of the panicle subdivided, single or fascicled, spreading and usually naked below, with the spike- lets pedicelled, sometimes long-pedicelled. Panicle small, narrow; branches few, erect. 28. P. ciliatissinum, Buckl. 29. P. xanthophysum, Gray. 30. P. pedicellatum, Vasey. dl. P. depauperatam, Muhi. 32. P. angustifolium, HUl., non Chapman. Panicle small but diffuse, oval or oblong: ITI RE TSS) 1S) NG) I oo Ki “1 SD Ot rg 33. P. mtidum, Lam. o4. P. laxiflorum, Lam. 30. P. dichotomum, Linn. 36. P. neuranthum Gris. of. P. Joorii, Vasey. 38. P. nudicaule, Vasey. 39. P. consanguineum, Kth. 4). P. scoparieum, Lam. 41. P. Wilcoxianum, Vasey. Panicle larger, oval, or oblong: 42, P. sperocarpon, Hl. 43. P. microcarpon, Muhl. 44, P. viscidum, Hl. 45. P. commutatum, Schultz (P. nervosum, Hl.). 46. P. clandestinum, Linn. 22 ES Panicle larger, oval, or oblong—Continued. 47. P. latifolium, Linn. 48. P. scabriuseulum, £ll.? Chap. Panicle effuse; branches capillary and much subdivided, except in P. sparsiflorum (P. angustifolium Chap., non EHIL): 49. P. capillare, Linn. 50. P. autumnale, Bosc. of Ptah; Vas: 52. P. proliferum, Lam. 53. P. miliaceum, Linn. 54. P. verrucosum, Jluhl. 55. P. sparsiflorum (P. angustifohum, Chap., non £Il.). Tall grasses, with ample and diffuse panicle : 6 P. anceps., Miche. . P. agrostoides, Miuhl. 59. P. maximum Jacq. (P. jumentorum, Pers.). 60. P. amarum, Hl. 61. P. virgatum, Linn. 62. P. Havardii, Vasey. SEcTION VIII.—Frutescentes.—Culms frutescent, spikelets large and turgid, empty glumes loose. 63. P. divaricatum, Linn. SECTION LX.— Villifora.—Panicle diffuse, spikelets very large, the glumes very silky-villose. 64. P. Urvilleanum, Ath. Section J.—DIGITARIA. 1. P. glabrum, Gaud. (Digitaria humifusa Pers)—Culms spreading, prostrate or erect, 5 to 12 inches long; leaves 1 to 2 inches long, smooth or pubescent; spikes 2 to 6, spreading, 1 to 2 inches long, approxi- mate; spikelets oblong, about 1 line long, obtusish; empty glumes, nasty two (the lower one generally wanting), pubescent, both equaling the flower. In cultivated and waste grounds less common than the following. Variety Mississippiensis, Gattinger.—Taller (6 to 20 inches high), spikes longer (14 to 34 inches), spikelets acutish. Vicinity of Nashville, Tenn. (Dr. Gattinger). 2. P. sanguinale, Linn. (Crab-grass).—Culm erect or decumbent, usually rooting at the lower joints, leaves and sheaths smooth or hairy, variable in length and breadth; spikes 3 to 10 or 15, spreading, digi- tate or approximate, 3 to 6 inches long; spikelets oblong-lanceolate, about 14$ lines long, acute, smoothish, pubescent; first glume minute, second half to two-thirds as long as the spikelet, third equaling the spikelet. 23 Variety ciliare (P. ciliare etz). Naturalized everywhere. 3. P. fliforme, Linn.—(Digitaria filiformis Muhl.).—Culms very slender, 1 to 3 feet high, erect, simple, or Sparingly branched; leaves narrowly linear, 3 to 10 inches long, erect, somewhat pubescent, and with sheaths hairy, spikes 2 to 8, erect or somewhat spreading, 2 to 6 inches long; spikelets 1 line long or less, oblong, acute; empty glumes two, the lower slightly shorter and the upper equaling the flower. Sandy or graveily ground; Maine to Florida and westward. 4, P. serotinum, Zrin. (Digitaria serotina Mr.; D. villosa Hll.).—Per- ennial, extensively creeping, much branched, making a thick carpet; flowering culms 6 to 10 inches ; leaves short, erect, hairy, 1 to 245 inches long, Sheaths villous; spikes 3 to 5, about 2 inches long, narrow; spike- lets less than 1 line long, acute, two empty glumes, the first one-quarter to one third as long, and the second nearly as long, as the spikelet. North Carolina to Florida and westward. Section I].—TRICHACHNE. 5. P. leucopheum, 7. B. K.—Culms stout, 2 to 4 feet high, crect; leaves broadly linear, 1 foot long, scabrous above, ligule ciliate; pan- icle racemose, contracted, dense, 6 to 10 inches long, the simple branches numerous, erect, fasciculate, 2 to 4 inches long; spikelets linear-lanceo- late, acuminate, about 2 lines long; lower glume minute, second nearly as long as, and the third equaling, the perfect flower, which is linear-lan- ceolate and mucronate-pointed. Low, rich ground, South Florida, (Chapman, Curtiss, eitc.). 6. P. lachnanthum, Torr.—Culms rather slender, 2 to 3 feet high; leaves much narrower and shorter, panicle shorter, more slender, with fewer and shorter spikes; spikelets smaller, 1 to 14 lines long, ovate- lanceolate, the perfect flower ovate-lanceolate, acute and mucronate- pointed. - Mostly on dry hills, Texas, Arizona, and New Mexico. Section IJI.—SuBSPICATA. 7. P. paspaloides, Pers.—Culms 2 to 3 feet high, decumbent and root- ing below, smooth, stout, leafy; leaves narrow, 6 to 10 inches long, be- coming somewhat involute, and with the sheaths smooth; panicle 5 to 10 inches long, strict, of 10 to 20 appressed, sessile, simple branches, the lower ones 1 to 14 inches long, above gradually shorter; spikelets ovate- oblong, obtusish, smooth, 1 to 14 lines long, imbricate in two rows on the narrow axis, the lower glume truneate-obtuse, one-third as long, and second glume two-thirds as long as the spikelets (also thin and obtuse). In ditches or water, Florida to Texas and Mexico. 8. P. Curtisii, Chap.—Culms 3 to 4 feet high, often rooting at the lower joints; leaves linear-lanceolate, plane, rigid, 6 to 9 inches long, 6 to 9 lines wide, smooth, sheaths smooth or hairy; panicle slender, 24 spike-like, 6 to 8 inches long, of numerous appressed slender, single, or fascicled branches, the lower oues remote; spikelets ovate-lanceolate, acutish, about 1 line long, lower glume acute, keeled, half as long as the spikelet, the second a little shorter than the third, which equals the . acutish perfect flower; second and third glumes strongly three to five nerved, sterile flower triandrous. Closely resemuples the preceding. Ponds and swamps, Florida to Texas. 9. P. obtusum, H. B. A.—Cuims 14 to 2 feet high, firm, leafy; send- ing off long (sometimes 8 to 10 feet) runners from the base; leaves 4 to 6 inches long, 2 to 3 lines wide; panicle 3 or 4 inches long, nar- row, composed of four to seven approximate, appressed, sessile branches, the lower ones about 1 inch long, the upper becoming very short, densely flowered ; spikelets mostly in alternate pairs of which one is sessile, the other short-pedicelled, forming three to four rows, turgid, oblong, obtuse, and 14 lines long, lower glume two-thirds as long as the spikelet, five-nerved, second glume seven-nerved, third five-nerved, a little longer than the fertile flower; sterile flower triandrous. Mexico, Texas, New Mexico to Colorado. 10. P. reticulatum, Torr.—Culmns 19 to 15 inches high, much branched below, leafy; leaves 14 to 3 inches long, acute, scabrous or pubescent, as ao are the sheaths; panicle 2 to 3 inches long, lateral ones shorter, close or dense; branches single or fascicuiate, unequal, 14 or 2 inches long, the longer with interrupted spikelets; about 14 lines long, sin- gle or in pairs, one sessile and one pedicelled, obovate, acutish; first glume one-fourth to one-third as long as the spikelet, acutish, second and third strongly five to seven nerved, and conspicuously reticulately veined. Texas to Arizona. 1i. P. Texanum, buckley.—Culms stout, 2 to 4 feet high, branehing from a decumbent and sometimes rooting base, leafy, smooth; leaves lanceolate, 4 to 10 inches long, 6 to 12 lines wide, smoothish or scab- rous above and on the margins, sheaths softly and sparsely pubescent, nodes pubescent; panicle (the base generally inclosed in the upper sheath) 4 to 8 inches long, one-half to 1 inch wide, of 5 to 10 erect, rather slender, simple, erect branches, 1 to 3 inches long, closely fereet rhachis scabrous-pubescent; spikelets single or in pairs, alternate mene the angular rhachis, oblong, lanceolate, acute, 2 to 24 lines long, smooth, or finely pubescent; firstglume broadly ovate, half to two-thirds as long as the spikelets, acute, five nerved, second and third glumes equaling the spikelet, strongly five to seven nerved, perfect flower ovate or ob- long-ovate, acutish, transversely wrinkled with fine reticulated strize. Cultivated for a forage grass. Texas. 12. P. Chapmani, Vasey. (P. tenuiculmum, Chap. non Meyer),—Culms filiform, 14 to 24 feet high, erect, simple; leaves chiefly radical, linear, half as lene as the culm, aboune 2 lines wide, upper ones shorter; ama 6 to 12 inches long, of ete to twelve remote, slender, erect spikes, the ; 25 upper ones short and few (three’to six) flowered, sessile, one-half inch long, the lower longer, sometimes subdivided, and with long, sheathed peduncles; rhachis flexuous, bristle-like at the apex, spikelets about one line long, ovate, lower glume one-half as long as the spikelet, ob- tuse, upper glume nine-nerved. Florida (Chapman, Curtiss, Garber). 13. P. Reverchoni, Vasey. (distributed by Reverchon as Setaria uni- seta).—Culms filiform from a thick, strong rootstock, erect, 10 to 18 inches high, simple; leaves chiefly radical, long and narrow; half as long as the culm; panicle 3 to 6 inches long, slender, an interrupted spike, at least above, below with short, three to five-flowered, remote, fascicu- late branches; spikelets 14 lines long, plano-convex, obtusish; lower, glume one-half as long as the spikelet, three-nerved, second and third seven-nerved; rhachis terminated by a short bristle, sometimes also a bristle below the spikelets. Texas (Reverchon),. 14. P. subspicatum, Vasey.—Culms 10 to 18 inches high, leafy; upper leaves longer than the lower (lowest 1 inch, highest 5 inches), 1 to 2 lines wide, rather rigid, light green, acute; panicle spicate at apex, below with five or six short, approximate, appressed, rather lax branches, one-half inch to 1 inch long; spikelets 1 line long, a little curved, lower glume one-half as long as the spikelet, second glume seven-nerved, third glume five-nerved; rhachis with a short bristle at apex and Sometimes a bristle below the spikelets. eaves shorter and more rigid, spike shorter and closer, and spikelets smaller than in the pre- ceding. Texas (Buckley, Nealley). These three (12, 13, and 14) form a natural group. 15. P. stenodes, Gris. (P. anceps var. strictum Chap.).--Culms 2 to 3 feet high, wiry, smooth; leaves erect, rigid, narrow (1 to 2 lines), 3 to 4 inches long; panicle narrow, 2 to 3 inches long, of five to six slender, Short, appressed branches, sessile or nearly so, mostly single, some- times two or three together; spikelets about 1 line long, lanceolate, acute; first glume half as long as the spikelet; second and third five- nerved, acute. In ponds and water. — Florida to Alabama (Dr. Mohr) and Texas (Nealley). Section IV.—PLATYPHYLLA. 16. P. platyphyllum, Munro.—Culms decumbent and rooting at.the lower joints, 1 to 2 feet high, branching below, pale green in color; leaves lanceolate, 3 to 4 inches long, 4 to 6 lines wide, smooth, firm, the base clasping; margins scabrous, pale green; sheaths smooth, striate, the upper one inclosing the base of the panicle, which consists of three to five simple, sessile, alternate, slightly spreading, distant branches, each 13 to 3 inches long; spikelets sessile, and alternate on a flat- tened rhachis; lower glume thin, obtuse, one-third as long as the sec- 26 ond and third, which are five-nerved, smooth, and one-third longer than the obtuse fertile flower; sterile flower of a glume only. Texas (HK. Hall, G. C. Nealley). 17. P. plantagineum, ZLink.—Much like the preceding, but stcuter and larger, with larger and wider leaves (6 to 10 inches by 6 to 9 lines), the spikes more remote, longér (2 to 4 inches), and the lower peduncled in the sheaths; spikelets much like the preceding. Introduced on ballast ground at Philadelphia. Section V.—BRACHIARIA. 18. P. prostratum, Zam. (P. cespitosum Siz ?).—Culms creeping and rooting at the base, then ascending, 1 to 2 feet long, with rather distant leaves, which are 14 to 2 inches long, lanceolate or cvate-lance- olate, acute, clasping at the ciliate base, sheaths shorter than the inter- nodes; panicle exserted, short, about 2 inches long, of five to eight simple or subsimple branches, the lowest of which are 1 to 13 inches long, closely flowered; rachis frequently with stiff hairs; spikelets three- quarters of a line long, smooth, first glume one-quarter as long as the spikelet, second and third equal; perfect flower striate-punctulate, mu- cronate-pointed. Our specimens agree with Trinius, Fig. 185 A. 7 Louisiana (Langlois), Texas (Nealley). 19. P. cespitosum, Swz.?—Culms numerous, succulent, and thickish, rooting at the base, ascending to 1 or 2 feet long; leaves crowded below, erect, linear-lanceolate, 2 to 4 inches long, 3 to 4 lines wide, ciliate on the margins, striate, lower sheaths inflated, panicle about 2 inches long, of five to eight branches becoming long exserted, rhachis and branches hairy and angular, lower branches about 1 inch long; spike- lets 14 to 14 lines long, ovate-oblong, acute, lower glume one-third as long as the spikelet, perfect flower as in the preceding but larger. The whole plant pale green. robably in Texas and New Mexico, certainly in Chihuahua, Mexico. (Pringle, No. 375.) 20. P.fuscum, Swz. (P. fasciculatum Sw.).—Culms 1 to 3 feet high, de- cumbent and branching at base; leaves linear or lanceolate-linear, 3 to 10 inches long, glabrous or pubescent; panicle 3 to 6 inches long, spread- ing, the branches simple or more or less fasciculated below, the luwer about 2 inches long; rhachis hairy; spikelets in pairs or in small clust- ers, 14 lines long, acutish, lower glume about one-third as long as the spikelet, deltoid, second and third glumes five to seven nerved, and cross-nerved. Texas. Variety major; culms stouter, 2 to 3 feet high, branched, sheaths and leaves scabrous-punctulate or smoothish, leaves broader, acumi- nate; panicle fuller, 3 to 6 inches long, branches more humerous (up to twenty or thirty), scattered and fasciculate, rhachis and branches scabrous, and with seattered hairs. Mexico (Dr. HK. Palmer). 27 Variety fasciculatum (P. fasciculatum, Swz.); leaves longer and wider (8 to 10 iines wide), panicle 4 to 5 inches long, the branches very numer- ous, Spikelets becoming dark brown. Southern Florida. 21. P. grossarium, Linn. Specimens from ballast ground at Philadelphia have been referred to this species, but if correct it seems too near P. fuscum. Section VI.—POLYSTACHYA. [Small forms of P. anceps might come in this group. | 22. P. barbinode Trin. (Para-grass).—Culms 2 to 6 feet high, stout, ascending; nodes vil:ous; leaves linear-acuminate, glabrescent; panicle lax, 6 to 8 inches long, branches mostly simple or a few, fascicled, spreading, about 2 inches long, somewhat distant, spikelets 1 to 14 lines long, glabrous, one-sided, spreading, lowest glume deltoid, one- fourth as long as the spikelet, one-nerved, second and third equaling the spikelet, five-nerved; fertile flower ellipsoidal, bluntish. Introduced and cultivated in the South. 23. P. gymnocarpon, Hll.—Perennial. Culms 2 to 4 feet high, erect, rigid, smooth; leaves lanceolate, smooth, 1 foot long or more, 1 inch or more wide, cordate at base, sheaths shorter than the internodes, striate, nodes black; panicle large, 9 to 15 inches long, branches 4 to 6 inches long, erect-spreading, scattered or 2 to 3 together, racemose, spikelets in nearly sessile clusters of 3 to 6, appressed along the branches, some- what one sided, 245 to 3 lines long, lanceolate, outer glumes nearly alike, lanceolate-subulate, rough-keeled, two to three times longer than the perfect flower, first glume three-nerved, a little to one-third shorter than the second and third, which are about five-nerved, the third glume or neutral flower with a palet one-half as long. TIlorida to Texas. 24. P. repens, £.—Culms 14 to 2 feet high, from strong creeping rhi- zomas, leafy, covered below with loose sheaths with short or deficient blades, the blades becoming longer upward, conduplicate or involute, the uppermost 3 to 4 inches long, sometimes pubescent at the base; panicle erect, thin, 3 to 4 inches long, the branches alternate, naked at base, erect or slightly spreading, the lower ones each with two or three rather long, appressed branchlets; spikelets short-pedicelled and ap- pressed, interrupted or racemose along the slender branches, 1 line long, oval, acute, smooth, the lower glume broad, obtuse, one-third as long as the spikelet, second and third about seven-nerved. Sandy shores of the Gulf, Florida to Texas. Variety confertum, Vasey. Culms shorter, leaves very divergent and rigid, panicle more con- densed, the branches shorter and spikelets more crowded, lower glume rather longer and less obtuse. Mobile, Ala., to Vera Cruz, Mex. (2177 F. Muller.) 28 25. P. anceps, Mich.—See No. 56. 26. P.laxum Sv. of this group, common in the West Indies, is prob- ably also in southern Florida, but I have seen no authentic specimens. 27. P. hians, £ll—Perennial, culms slender, smooth, 6 to 18 inches high, simple, from slender, wiry, creeping root-stocks; leaves linear, 3 to 5d inches long, 1 to 2 lines wide, smooth, erect; panicle rather small, 3 to 5 inches long, the slender, racemose branches 1 to 2-inches long, erect-spreading, mostly single and distant, the lower third naked; spikelets about 1 line long, in small, nearly sessile, approximate clus- ters, outer glumes ovate, acute, generally three-nerved, the lower one- half as long as the spikelet, third glume longer than the seeond and having a thick, rigid, obovate palet rather longer than its glume, and spreading apart from it, hence probably the name hians, from hio, to gape. North Carolina to Texas. 28. P. ciliatissimum, Buckl.—Culms procumbent and much branched, often rooting at the joints, at first short-jointed and much condensed, with leaves 1 to 14 inches long, and the panicles short and invaginate, becoming elongated, with long-exserted panicles (lateral and terminal) which become 14 to 2 inches long, narrow, with a few short, few-flow- ered, appressed branches, and linear-lanceolate, acuminate leaves, 2 to 4 inches long, ciliate on the margins below, with sparsely ciliate and hairy sheaths, the nodes white-woolly; spikelets ovate, acute, 2 lines long, pubescent or villous; the lower glume lanceolate, acute, three- nervéed; smooth, except at the base, two-thirds as long as the spikelet; second, eleven to thirteen nerved ; pubescent to densely villous, with a smooth, acute, hardened point; the third one, or flowering glume of the sterile flower a little shorter than the second, five-nerved, ciliate on the margins, its palet equally long, ovate, thin, the fertile flower one- fourth to one-third shorter than the largest glume, obtusish, minutely furrowed. Texas, Section- VII.—EUPANICUM. 29. P. Xanthophysum, Gray.—Culms erect, 1 to 2 feet high, simple or branched near the base; leaves lanceolate, acuminate, 4 to 6 inches long, 5 to 10 lines wide, smooth except the scabrous margins, strongly nine to eleven nerved, rather contracted at the ciliate, clasping base; panicle long-exserted, 2 to 4 inches long, of a few simple, erect or appressed, few-flowered branches; spikelets obovate, 14 lines long, minutely downy; lower glume about half the length of the spikelet, second and third about nine-nerved. Plant yellowish-green. Canada to Pennsylvania and Wisconsin. 30. P. pedicellatum, Vasey.—Culms 1 to 2 feet high, slender, branching below, smooth ; culin leaves 2 to 3 inches long, 2 to 3 lines wide, gradu- ally tapering to an acute point, erect, somewhat rigid ; main panicle long exserted, the lateral ones less so, all small and few flowered, 13 to 2 oe 29 inches long, of four or five alternate short branches each only two to three flowered; spikelets long-pedicelled, oblong-obovate, obtuse, 14 lines long, sparsely pubescent, the lowest glume at some distance from the others, ovate, one-nerved, nearly half as long as the spikelet, second and third glumes oblong, obtuse, seven-nerved. Texas, J. Reverchon. | Resembles P. angustifolium, H#Il., but is smaller, with smaller, fewer- flowered panicle, and rather smaller spikelets. 31. P.depauperatum, Muhl.—Culms erect, simple, tufted, 6 to L6inches high; leaves of the culm about three, erect, narrowly linear, 3 to 6 inches long, sheaths usually pubescent; panicle small, 1 to 2 inches long, contracted, sometimes overtopped by the upper leaves ; spikelets 1 to 14 lines long, obovate, lower glume one-third the length of the seven to nine nerved upper ones. Common east of the Mississippi. Variety laxa.—Weaker stemmed, panicle with longer and more spreading branches (14 to 2 inches) the lower ones single, or verticil- late; spikelets smaller. Virginia, Florida, Texas, Arkansas, Missouri. 32. P. angustifolium, Hil., non Chapman.—Culms weak, slender, smooth, 1 to 2 feet: long, diffusely branched from the base; leaves linear, 3 to 5 inches long, 1 to 2 lines wide; sheaths short, nodes thick- ened; panicles simple, terminal and lateral, 3 to 5 inches long; the few branches single, capillary, distant, 1 to 3 inches long, bearing each two to six spikelets which are mostly in pairs at the end of the branchlets ; spikelets oblong-obovate, acute, 14 lines long, the outer glumes papil- | lose-hispid, the lower one minute, acutish. South Carolina to Texas. 33. P. nitidum, Lam.—Culms 10 to 20 inches high, slender but firm, rarely branched, smooth; leaves mostly at the base of culm, lanceolate or lance-linear, 1 to 2 inches long, rather rigid, and with the sheaths smooth, the culm leaves few and smaller; panicle ovate or oblong, 14 to 24 inches long; branches verticillate below, the lower ones, 1 to 14 inches long, spreading, capillary, numerously flowered ; purplish spike- lets on pedicels 2 to 4 times aslong, oval, one-haif to two-thirds line long, pubescent, the lowest glume one-fourth to one-third the length of the spikelet. (a) Variety minor (P. ramulosum of Chapman’s supplement, but I think cannot be the plant of Michx.)—Lower than the type (6 to 8 inches high); tufted; leaves narrower and smaller ; panicle 1 to 14 inches long ; rhachis smooth ; spikelets smooth or nearly so; second and third glumes five-nerved. Florida. (b) Variety ensifolium (P. ensifolium Bald.).—Radical leaves linear lanceolate, 14 inches long, smooth, those of the slender (8 to 12 inches long) culm, distant, small (about one-half inch), and horizontally diver- 30 gent; panicle depauperate, less than an inch long, the rays single and few flowered. Described from Dr. Chapman’s specimens. Florida. | (c) var. major.—Here I could place several variable forms, mostly smooth, with stouter culms, a more open, less dense panicle, with fewer and stouter branches (sometimes reflexed), and fewer and larger spikelets. | 34, P. laxiflorum, Lam.—Culms erect, weak, 6 to 18 inches high, rarely branched, smooth below the panicle ; leaves linear or lanceolate-linear, 3 to 7 inches long, 3 to 4 lines wide, mostly radical, the 2 or 3 of the culm little shorter, acuminate, nearly smooth except on tne ciliate mar- gins, pale green or yellowish-green; sheaths mostly shorter than the leaves, sparsely and retrorsely villose; ligule a ring of soft, white hairs, panicle capillary, 2 or 3 inches long, branches sparingly divided from the base, mostly single, alternate and few flowered, spreading; the © rhachis and branches sparsely long-hairy; spikelets long-pedicelled, 1 ‘line long, oval, or elliptical, obtuse, strigosely pubescent, lower empty glume ovate, one nerved, one-quarter as long as the spikelet; second and third glumes seven to nine nerved; fertile flower as long as the spikelet, acutish. In sandy woods. Var. pubescens.—Culm rather stouter, and pubescent, 12 to 15 inches high; leaves hairy, shorter, lanceolate, the lower 2 to 24 inches long, 3 to 5 lines wide, the upper about 1 inch, upper sheath elongated ; pani- cle 2 to 34 inches long, the branches and spikelets more numerous, and spikelets smaller; lowest giume roundish ovate, one-third as long as the spikelets; second and third glumes five to seven nerved. Closely ap- proaches P. nitidum, and is perhaps the P. nitidum, var. ciliatum, Torr. 35. P. dichotomum, Linn.—Culm 10 to 30 inches high ; at first mostly simple, smooth, or pubescent, bearing a more or less exserted, spread- ing, compound panicle, 1 to 4 inches long, and lanceolate, flat leaves; those of the culm larger above, 3 to 4 inches long; those at the root tufted, usually ovate-lanceolate, short, and thick ; the culms commonly branching later in the season, the branches often clustered, and bearing smaller, mostly included panicles; spikelets about 1 line long (two- thirds of a line to 1 line), oblong-obovate, pubescent or downy, lower glume one-third as long as the spikelet. (a) Variety barbulatum.—With the nodes barbed, and with the sheaths smooth or pubescent. (b) Variety viride —Smooth all over, leaves light green and narrower. (c) Var. divaricatum.—E ect, 6 to 9 inches high, slender, smooth, very much branched above the base, caespitose, the radical leaves very short and tufted, those of the culm and branches very narrowly linear or in- volute, acuminate, about 1 inch long, divaricate, the numerous panicles small, racemose, few flowered ; the spikelets on alternate, filiform ped- icels one-half inch to 1 inch long; spikelets three-fourths of a line long, oval-oblong, smooth, the lower glume one-third as long as the spikelet; second glume one-third shorter than the spikelet, five-nerved ; third ol glume equaling the spikelet, seven-verved; fertile flower as long as the spikelet. (d) Variety villosum (P. villosum Hll.2—Culms decumbent, branching from the first, leaves scabrous above, ciliate, sheaths and culms villose, panicle thinner and fewer-flowered. (e) Var. elatum.—Culms tall (2 to 3 feet), erect, smooth; cauline leaves 4 to 5 inches long, 5 to 6 lines wide, smooth, very acute; pan- icle 4 inches long, smooth, the branches long, erect-spreading, rather sparsely flowered; spikelets a little more than 1 line long, smooth, oblong, acutish, the lower glume ovate, acute or acutish, nearly half as long as the spikelet, fertile flower shorter than the spikelet. 36. P. neuranthum Gris.—Erect or ascending, about 1 foot high, slender, smooth; leaves linear-acuminate, 2 to 4 inches long, firm, flat or subinvolute; terminal panicle long exserted, 1 to 2 inches long, with few-flowered, divergent branches, the lateral panicles numerous, narrow, short (less than 1 inch long), oval, pubescent, lower glume mi- nute, deltoid, nearly nerveless, one-third to one-fourth as long as the spikelet, second and third seven-nerved, equaling the fertile flower, which is ovoid, obtuse, and smooth. (a) Variety ramosum Gris.—Ascending, much branched, leaves flat or involute, very narrow; spikelets 1 line long, elliptical, obtuse. South Carolina, Florida to Texas. ov. P. Joorii, Vasey.—Culms about 1 foot high, much branched above, very leafy; leaves 4 to 6 inches long, 6 to 10 lines wide, thinnish, smooth, those of the extreme branches smaller; panicles numerous, small, inclosed in the sheaths, few-flowered; spikelets oblong, over 1 line long, sparsely pubescent. Louisana (Dr. J. F. Joor); Mississippi (Professor Tracy). This is very unusual in its abundant, large leaves, and condensed form, growing in dense tufts. o8. P. nudicaule, Vasey.—Culms 14 to 2 feet high, slender, with 2 or 3 long internodes; leaves mostly near the base, linear lanceolate 2 to 4 inches long, 2 to 5 lines wide, smooth, light green, the 2 to3 culm-leaves distant, narrow, giving the culm a naked appearance; panicle long-ex- serted, small, smooth, 2 to 3 inches long, 1 inch wide, sparsely flowered, branches alternate; spikelets elliptical-oblong, acute, 14 lines long, smooth, the lower glume very small ‘one-fifth as long as the spikelet), second glume seven nerved, third, five nerved, fertile flower one-fourth shorter than the spikelet. Swamps, Santa Rosa County, Fla. (A. H. Curtis). 39. P. consanguineum, Ath. (P. villosum #Ul.).—Culms about 14 feet high, erect, branching in age; leaves linear-lanceolate, mostly about 3 inches long, 2 to 5 lines wide, the later ones almost filiform, somewhat pubescent, sheaths villose; panicle 2 to 3 inches long, open, rather thin; spikelets 1 to 14 lines long, pubescent. South Carolina to Florida. 32 = 40. P. scoparium Lam. (P. pane Hll.).—Culms erect, becom- ing branched and reclining, 1 to 2 feet high, somewhat ScHIEDES: leaves erect, about 2 inches long, 4 to 5 lines wide, faintly nerved, ceroans on the margins, hairy-fringed near the base, the sheaths bristly with stiff, spreading hairs, or smoothish; the panicle 2 to 3 inches long, spreading, open, branches smooth, mostly simple, alternate, rather few flowered ; spikelets 1$ to 12 lines long, oval to obovate, hairy or smoothish, lower glume about one-third as long as spikelet. New England to California and southward. A widespread species. (a) Var. major (P. scoparium, /li.).—This differs from the preceding in its stouter, rougher culm, larger leaves, 4 to 6 inches long, three- fourths of an inch wide (1 to 13 inches wide, Elliott), which are soft- pubescent below, and its spikelets 2 lines long. We have only seen this from South Carolina ine Ravenel). (b) Var. angustifolium.—Culms tall and erect, 2 to 3 feet high, branch. ing; leaves longer and narrower than the faba: 3 or 4 inches long, 3 to A lines wide, rather rigid, sheaths sparsely pubescent; panicle long-ex- serted, 3 to 4 inches long. South Carolina (Dr. Ravenel); Ilinois (Dr. Schenck); Fortress Mon- roe, Va. (Vasey). (c) Var. Liebergii (P. Liebergii, Scrib.).—Culms tall and erect, un- branched, 2 feet high; leaves lanceolate, distant, 3 to 4 inches long, 6 to 9 lines wide, scabrous or papillose, hairy, about eleven-nerved, sheath shorter than the internodes, ciliate, or the margins Somewhat scabrous and sparsely hairy, panicle about 3 inches long, 1 ineh wide, the fewish branches erect-spreading and few-flowered; spikelets obovate, obtuse, 13 lines long, the outer glumes papillose, hairy, lower glume ovate or ovate-lanceolate, pointed, nearly half as long as spikelet, second glume seven-nerved, third glume nine-nerved, equaling the flower, having a hyaline palet of nearly its own length, and inclosing three stamens. Plymouth County, Lowa (John Lieberg). 41, P. Wilcoxianum, Vasey.—Culms about 6 inches high, entire or sparingly branched and with the leaves more or less white-hairy; leaves linear-lanceolate, erect, acuminate, 2 to 3 inches long, hairy both sides, rather rigid, sheaths striate, hairy, mostly longer than the internodes, ligule obsolete; panicle oblong, rhachis zigzag, 1 inch long, lower branches subverticillate, short, spikelets about 14 lines long, pubescent, lower glume ovate, one-fourth as long as the spikelet, second aud third glumes with five toseven broad nerves, the third hardly as long as the flowering glume. Nebraska (Dr. T. E. Wilcox). 42, P.spherocarpon, 4/l.—Culms 15 to 24 inches long, dto 7 lines wide, lanceolate, acute, obscurely nerved, smooth except at the ciliate-cordate base, sheaths shorter than the joints, smooth; panicle 3 to 4 inches long, spreading, spikelets oval, five-eighths to three fourths of a iine long, 30 smoothish, lower glume one-third as long as the spikelet. Differs from P. dichotomum in its more rigid, smooth, pale leaves, with cordate base. Dry or moist ground, of same range as the next species. Var. Floridanum.—Culms 13 to 24 feet high, rigidly erect, leaves rigid, erect, six to seven on the culm, lanceolate, smooth, about 3 inches long, 5 to 6 lines wide, the margins near the base, and sheaths ciliate ; panicle oval to oblong, 3 to 5 inches long, and spikelets one-half line long, pubescent, the lower glume one-fourth or one-fifth as long as the spikelet. Florida. — Described by Dr. Chapman as P. sphaerocarpon Hll., but it hardly agrees with Elliott’s description. It isintermediate between P. micro- carpon and P. spheerocarpon. 43. P. microcarpon, Muhl. (P. multiflorum, #l.).—Culms 2 to 3 feet high, stout, erect, smooth ; leaves 4to 7 inches long, 9 to 12 lines wide, lanceolate, gradually tapering to a slender point, with nine to eleven nerves, roughish above and, on the margin, sometimes bristly-ciliate at the rounded, clasping base, smooth on the under side, sheaths mostly longer than the internodes, smooth except on the margins; ligule nearly obsolete, panicle becoming long-peduneled, 3 to 7 inches long, 1 to 2 inches broad, spreading, multiflorous, branches mostly verticil- late, very numerous, flowering to the base, spikelets mostly long-pedi- celed, oval, five-eighths of a line long, nearly smooth, lower glume one- fourth as long as spikelet, second glume 7-nerved. Low ground, Massachusetts to Texas. 44, P. viscidum, Hll.—Culms 2 to 4 feet high, stout, becoming much branched, leafy, velvety-downy all over, except a narrow ring below each node; sheaths soft-downy, the hairs spreading or reflexed and often viscid, leaves downy or smoothish, lanceolate, 4 to 8 inches long, diffuse, the branches numerous and much subdivided, spikelets fully 1 line long, oblong-ob: vate, pubescent, lower glume one-fourth to one-fifth as long as the 7 to 9 nerved upper one. Common. 45. P. commutatum, Schultz (P. nervosum, Muhl.).—Culms 2 to 23 feet long, erect, unbranched, rather siender, smooth; leaves at the base rigid, ovate-lanceolate, the three or four culm leaves firm but not rigid, mostly 3 to 4 inches long, 8 to 12 lines wide, acuminate, cordate at base, smooth except on the margins, sheatas smooth or somewhat pubescent, much shorter than the nodes; panicle peduncled, 3 to 5 inches long, 2 to 3 inches wide, open, rather thin, branches smooth, single or subverticillate; spikelets rather long-pedicelled, oblong, sparsely hairy, 1 to 14 lines long, lower glumes one-third as long as the spikelet, second about five, and third seven nerved. A fine species, in 11244—No. 8 3 34 appearance like P. clandestinum, but more slender, with shorter leaves, and smaller spikelets, which are early deciduous. Pennsylvania to Texas. V (a) Var. minor, differs from the preceding in its rather smaller size (id to 2 feet) and smaller panicle, and the spikelets more persistent. Southern States. 46. P. clandestinum, Zinn.—Culms rigid, 2 to 3 feet high, leafy to the top, rarely branched, upper nodes smooth, the lower often hairy; sheaths smoothish, or the lower ones often rough with papillose, bristly hairs, shorter than the internodes, leaves large, lanceolate ‘to oblong- lanceolate, 4 to 8 inches long, 9 to 15 lines wide, with a cordate, clasping base, very taper-pointed, smooth except the rough margins ; panicle at first partially sheathed, becoming long-pedunculate, somewhat oblong in outline, 4 to 6 inches long, open and becoming spreading, branches verticiilate ; spikelets 14 lines long, slightly pubescent or smooth, ob- long-ovate to obovate, the lower glume acutish, nearly one-half as long as the spikelet, second and third equal, many nerved; perfect flower ob- jong, pointed. New England to Virginia, North Carolina, and westward to Illinois. 47. P. latifolium, Linn.—Culms 1 to 23 feet high, smooth, mostly simple, the nodes (generally) and the collar of the sheath soft wooly- hairy, sheaths smooth or more or less pubescent; leaves broadly lance- olate from a cordate, clasping base, 3 to 4 inches long, about three- fourths to 14 inches wide, tapering to a point, roughish above, smooth, or sparsely hairy below, eleven to thirteen nerved; panicle more or less exserted, becoming long-peduneled, 2 to 4 inches ieng, the branches mostly alternate, spreading, rather few flowered; spikelets 14 to 2 lines long, obovate, sparsely pubescent, the lower glume one-third, or less than one-half as long as the spikelet, upper one about nine nerved. Maine to Texas. (a) Variety molle; downy all over. Virginia to Louisiana. (b) Var. australis; leaves more contracted at base, and spikelets lar- ger, 2 to 24 lines. Alabama to Texas. 48. P. scabriusculum, Ell. ? Chap. (P. Nealleyi, Vasey).—Culms 3 to 4 feet high, firm, smooth, rarely branched; leaves linear-lanceolate, 4 to 8 inches long, one-half inch wide, long-pointed, firm, mostly smooth except near the base, sheaths shorter than the joints, mostly smooth, or sometimes sparsely pubescent; panicle larger, 5 to 8 inclaes long, diffuse, smooth, branches mostly alternate, much divided, the larger ones 3 inches long; spikelets ovate, acutish, sparingly pubescent when young, becoming smooth, lower glume small, second glume strongly seven to nine nerved. North Carolina to Texas. This is the P, scabriusculum described by Chapman, but does not ap- 35 . pear to me to answer that of Elliott, who compares his with P. virga. tum. 49. P. capillare, Lina.—Culms 10 to 20 inches high, mostly branched at the base, rather robust, erect or ascending, sheaths and usually the leaves copiously hairy or hirsute; panicles, lateral and terminal, half to two-thirds the length of the plant, vaginate below, much branched, the branches divaricate when old; spikelets ovoid to oblong or lanceo- late, obtusish, acute or even acuminate, three-fourths to nearly 2 lines long, smooth, on long capillary pedicels, lower glume one-half to two- thirds as long as the spikelet, obtuse to acuminate, fertile flower ob- tusisb, rather shorter than the spikelet. All over the continent. a. Variety campestre, Gattinger.—Culms slender, 14 to 2 feet high, much branched, leaves and sheaths sparsely hirsute, panicle, rather pyramidal, much shorter than in the type (4 to 6 inches long, more open and thinly flowered, spikelets three-fourths of a line long, smooth, oblong, rather acute, lower glume obtusish. Tennessee, Dr. Gattinger. b. variety flexile, Gatlinger.—Culms slender, 13 to 24 feet high, branched below, leaves linear, 6 to 10 inches long, narrow, erect, smooth or smoothish, sparsely ciliate on the margins and sheaths, pan- icle longer and narrower than in the preceding, 5 to 9 inches long, 2 to 4 inches wide, branches slightly spreading, single or in twos, smooth; spikelets 14 lines long, lanceolate-acuminate, lower glume acute, per- fect flower one-third shorter than the spikelet. Tennessee, Dr. Gattinger. 50. P.autumnale, Bose. (P. divergens, Muhl.).—Culms 12 to 18 inches high, erect or decumbent below; leaves comparatively small, 2 or 3 inches long, 2 to 3 lines wide, smooth except on the scabrous or undu- late margins, sheaths smooth or the lower ones, sparsely hairy, ligule membranaceous, obtuse, conspicuous; panicle very effuse, one-third the length of the culm or more, the capillary branches long and sparingly divided, often reflexed at maturity, and somewhat scabrous, the axils sometimes sparsely hairy, the branchlets or pedicels long, naked, and terminated by a single spikelet, which is spindle-shaped or obovate, L to 14 lines long, acute, the lower glume very minute (one-sixth to one eighih as long as the spikelet), second and third glumes acute, ciliate near the apex, or in southwestern specimens (variety pubiflorum), pu- bescent all over, but little longer than the acute flowering glume. Illinois to Texas and the Southern States. 51. P. Hallii, V. @ S—Culms slender, 1 to 2 feet high, branching; leaves slender, 4 to 6 inches long; sheaths sparsely pubescent or smooth; panicle open, diffuse, 4 to 5 inches long, sparsely flowered ; spikelets 13 lines long, acute, smooth, lower glumes half as long as spikelets, perfect flower a little sherter. More slender than P. capil- 36 lare, with smoother culms, smaller panicles, stouter branches, and rather larger spikelets. Texas. This species has resemblanee to P. capillare on one side and to P. proliferum on the other. From the first, it is distinguished by its more erect slender culms, never more than sparsely pubescent, smaller pan- icle, with erect spreading stouter. branches, and usually larger or thicker spikelets; from the second, by its smaller size, more erect culins, and smaller panicles, with shorter branches. - 52. P. proliferum, Lam.—Annual. Culms usually thickish, often succulent, branching, geniculate at the decumbent or procumbent base, 14 to 3 feet long, glabrous; leaves linear, 6 to 12 inches long, sheaths flattened, glabrous, ligule ciliate; panicles terminal and lateral, 4 to 12 inches long, the long, slender, primary branches at length spreading and diffuse ; spikelets 1 to 14 lines long, pale green, sometimes purplish, appressed, short pedicelled, lower glume broad, obtusish, about one. fourth as long as the spikelet, perfect flower a little shorter than the spikelet. Damp places, Maine to Texas. (a) Var. geniculatum (P. geniculatum, HIl.).—Culms 3 to 6 feet high, succulent, Sometimes nearly an inch thick at the base, bent and branch- ing at the joints, leaves sometimes 2 feet long and 6 to 10 lines wide; sheaths much inflated when young, sometimes a little hairy at the base; panicle sometimes 2 feet long, very diffuse. A valuable grass. Southern States to Texas. 53. P. miliaceum, Linn.—Culms 2 to 4 feet high, erect, branched, pu- bescent; sheaths loose, striate, hirsutely pubescent; leaves flat, linear- lanceolate, 6 to 10 inches long, 4 to 8 lines wide, smoothish above, sparsely pubescent below; panicle oblong, nodding, 6 to 10 inches long, branches verticillate, erect-spreading (not diffuse), solitary or in pairs, angular, hispid, or scabrous; spikelets ovate, acuminate, glabrous, 2 lines long, lower glume broad, very acuminate, five-nerved, about one- half as long as the spikelet, the third glume seven to nine-nerved, a little longer than the oval, acute, biconvex, perfect flower. Cultivated, and rarely escaped from cultivation. Valuable for forage. 54. P. verrucosum, Muhl.— Culms slender, 1 to 3 feet long, somewhat branching, smooth; leaves linear-lanceolate, 3 to 4 inches long, 2 to 3 lines wide, and with the sheaths glabrous; panicles mostly terminal, 6 to 10 inches long, diffusely-spreading, branches mostly single, capillary, rather few flowered; spikelets three-fourths to 1 line long, obovate or oval, obtuse or abruptly acute, the outer glumes roughened with fine warts, the lower one about one-fourth as long as the spikelet; perfect flower, acute, about equaling the spikelet. New England to Florida and Mississippi. , 55. P. sparsiflorum, Vasey. (P. angustifolium, Chap. non Zll).— Culms weak, slender, smooth, 1 to 2 feet long, diffasely branched from . oT the base; leaves linear, 5 to 5 inches long, 1 to 2 lines wide; Ser ules short; nodes thickened ; panicles Shines Aonuma and lateral, 3 to 5 inches long, the few branches single, capillary, distant, 1 to 3 inches long, bearing each 2 to 6 spikelets, which are mostly in pairs at the end of the branchlets; spikelets oblong-obovate, acute, 14 lines long, the outer glumes papillose-hispid, the lower one minute, acutish. South Carolina to Texas. 56. P. anceps, Aichx.—Culms flattened, erect, 2 to3 feet high, less stout than in P. agrostoides, from strong creeping root-stocks; leaves rather broadly linear, a foot or more long, 4 to 5 lines wide, smooth or the lower ones with the sheaths pubescent; panicles contracted, lateral and terminal, 6 to 12 inches long, the branches from single to verticil- late, rather distant, particularly below, generally very abundantly flow- ered; spikelets 1 to 14 lines long, ovate to lanceolate, pointed, the apex a little curved, lower glume about one-half as long as the spikelet, ke elume five to seven-nerved. VY (a) Var. pubescens.—Leaves and sheaths more or less ube conte panicle and branches slender; spikelets ovate-lanceclate, acute, little more than 1 line long. Mobile, Ala. (Dr. Mohr). v (b) Var. angusta.—Lower sheaths villose; leaves very long and nar- row, erect, rigid; spikelets narrowly lanceolate, acuminate, 14 lines long, the lower glume one-third as long as the spikelet. Texas (G. C. Nealley). ¥ (ec) Var. densiflorum.—Leaves broad; panicle glomerate and inter- rupted, the branches densely flowered; spikelets lanceolate, 14 lineslong. Texas (J. FY. Riggs). | 57. P. agrostoides, Spreng.—Culms flattened, erect, stout, 2 to 4 feet high, usually in thick clumps, glabrous, much branched above, the branches erect; leaves linear, 1 to 2 feet long, 3 to 4 lines wide, smooth, as are the compressed sheaths; panicles terminal and lateral, 4 to 12 inches long, oblong, becoming pyramidal, very full, the lower branches 2 to 3 inches long, much subdivided to the base; spikelets racemose, very short pedicelled, crowded and mostly one-sided on the branches, ovate-oblong to lanceolate, three-fourths of a line to 14 lines long, lower glumes acute or acutish, half as long as the spikelet, second and third glumes five-nerved, half longer than the perfect flower, which is alittle bearded at the apex. In wet ground. Varies in the size and form of the spikelets, which in the Western form seldom exceed a iinelong; there is an Eastern form in which the spike- lets are lanceolate, about 14 lines long, and the perfect flower small and conspicuously stalked. 58, P. bulbosum, H. Bb. #.—Rhizoma creeping, thick, bulbous; culms 4 to 5 feet high, smooth, stout, with long internodes; leaves long and narrow (1 to 2 feet by 2 to 4 lines), smooth, or the lower ciliate below ; panicle about 1 foot long, spreading, copious, pale green, branches 38 scattered and verticillate, the lower 4 to 5 inches long, much subdivided nearly to the base, branchlets slender and numerously flowered; spikelets sometimes in pairs or racemose, 14 lines long, smooth, obtuse or acutish, not acuminate, lower glume half or rather more than half as long as the spikelet, broad, three-nerved, acutish, second and third glumes equal, five-nerved, the third with a narrow palet, perfect flower as long as the spikelet, oblong, lanceolate, abruptly acute, the point sometimes a little roughened. | Texas, New Mexico, and Arizona. Probably P. avenaceum /th. is not different. V (a) Var. minor (P. maximum, var. bulbosum, Munro), a smaller form, about 2 feet high, with the panicle much reduced. In same region. 59. P. maximum, Jacq. (P. jumentorum Pers.).—Culms erect, 3 to 5 feet high, glabrous; nodes softly pubescent; leaves linear, 1 to 14 feet long, 9 to 12 lines wide, rather rigid, margin scabrous, sheaths smooth ; panicle 14 feet long, much branched, hispid or scabrous, the branches verticillate, long and contracted, racemose, scabrous; spikelets mostly short-pedicelled, 15 lines long, ovate, smooth, acute; lower glumes about one-third as long as the spikelets; second glume shorter than the spikeiet; perfect flower acute. Introduced and cultivated in the South. 60. P. amarum, H/l.—Culms 2 to 3 feet high, from a stout running root-stock, thick, columnar, nearly one-half inch in diameter; leaves nearly flat, almost coriaceous, glaucous, 1 to 14 feet long, 4 to 6 lines wide, tapering to a long, slender point; sheaths glabrous, striate; pan- icle 1 foot or more in length, appressed, or becoming somewhat spreading, the lower branches verticillate, 6 inches long, subdivided nearly to the base, narrowly paniculate; spikelets rather racemose, ovate, 24 lines long, acute; glumes thick, the lower glume about two- thirds the length of spikelet, acute; second glume the longest, seven- nerved, acuminate; third glume rather shorter and obtusish; perfect flower linear-oblong, obtuse, one-fourth shorter than the spikelet. Grows among the sand-hills on the seashore. South Carolina to Florida. v (a) Var. minor, V. & S.—Leaves involute; panicle shorter, narrower, distantly branched, comparatively few-fiowered, the branches rather glomerate and unequal; spikeletsrather larger (24 to 3 lines); the lower glumes longer, or nearly as long as the spikelet. Fortress Monroe, Va., and northward, near the coast. 61. P. virgatumn, Linn.—Stout, erect, unbranched, 3 to 5 feet high, from strong, creeping root-stocks ; leaves flat, very long (1 foot or more), 3 to 4 lines wide, smooth; panicle compound, 6 to 18 inches long; branches single to verticillate, generally very numerous and becoming diffuse, spreading or drooping; spikelets on rough pedicels, 14 to 2 lines long, ovate to ovate-lanceolate, pointed, smooth ; lower glume more than half the length of the spikelet (sometimes two-thirds), five-nerved; sec- 39 ond, five-nerved; third, five to seven-nerved, acute to acuminate, larger than the perfect flower. Sterile flower with3stamens. Widely diffused and quite variable. The Atlantic form has spikelets about 14 lines long. The Western form has spikelets about 2 lines long, the glumes acuminate. A form on the Atlantic coast has the panicle quite close and rather oblong, and may be called variety conferta; another form (var. elongata) has a very long and narrow panicle, with spikelets even 24 lines long. Maine to Florida, and throughout the interior of the country. 62. P. Havardii, Vasey. (P. virgatum var. macrospermum, V.).—With the habit of western forms of P. virgatum, but much stouter; culms 5 to 6 feet high, with rigid, involute, long-pointed, glaucous leaves, li- gule a ring of short hairs; panicle 14 feet long, diffuse, the branches less abundantly flowered; spikelets 3 lines long; lowest glume one half as long as the spikelet, prominently five to seven-nerved; second glume nine-nerved, ovate, acuminate; third glume five to seven-nerved, its thick palet nearly as long; fertile flower about one-fourth shorter than the spikelet. Guadaloupe Mountains, Texas; Dr. Havard, G. C. Nealley. Section VII.—FRUTESCENTES. 63. P. divaricatum, Zinn.—Shrubby, smooth; culms 6 to 8 feet high with short and spreading branches; leaves lanceolate, acuminate, 2 to 4 inches long, 2 to 6 lines wide, deciduous from the persistent sheaths ; panicles terminating the branches, i to 3 inches long, nearly simple or branched, the branches rather distant and diverging, half to 1 inch long, rather sparsely subdivided; spikelets smooth, 2 lines long, tumid, obo- vate, nodding, on pedicels as long or longer; lower glume triangular- ovate, one-third as long as the spikelet; second and third glumes as long as the spikelet, very broad, nine to eleven nerved ; apex of the fertile flower downy-tipped. Southern Florida. Section [X.—VILLIFLORA. 64. P. Urvilleanum, Ath.—Culms 13 to 2 feet high, from a strong running root-stock, the whole plant densely soft-hairy ; near the base short-jointed and covered with weather-worn leaves or sheaths; leaves rigid, 12 to 18 inches long, 3 lines wide at the base, becoming con- volute, and ending in a long, setaceous point, the upper ones exceeding the panicle, which is 6 to 10 inches long, open, but rather contracted, the branches semi-verticillate or scattered, the shorter ones flowering to the base ; the longer ones (3 to 4 inches long) racemosely subdivided and flower bearing towards the ends or throughout; spikelets 24 to 3 lines long, ovate, acute; the empty glumes siiky-villous; lower glume seven-nerved, two-thirds as long as the fifteen-nerved second glume}; third glume rather shorter, eleven to thirteen-nerved; perfect flower oblong, glabrous, except on the margins, which are long-ciliate. In sand, southern California to Arizona. LAL 2 i: SECTION OF VEGETABLE PATHOLOGY. PREPARED BY B. T. GALLOWAY. CONTENTS. I. Potato Scab. JI. Gum Disease, or Foot-rot of the Orange. ilI. Parasitic Fungi of Missouri. IV. Extracts from Correspondence: . Remedies for apple scab. . Apple rust. . Bitter-rot of apples. . Pear blight. . Remedy for black-rot of grapes. . Black-knot of the grape. . Melon rust. . Anthracnose of the bean. . ** Malarial germ.” 43 SECTION OF VEGETABLE PATHOLOGY. POTATO SCAB. During the past year the Section has received frequent inquiries from different parts of the country relative to this disease, and while it is not our purpose in this paper to fully answer these, it is hoped that what is here stated may lead to a better understanding of the nature of the malady among those who are particularly interested. Potato seab is not a new disease by any means, nor is it peculiar to America, for we find frequent mention of its occurrence in Hurope; in fact, most of the important papers on the subject are to be found in German works on plant diseases.! The cause of scab is a question that has long been discussed. Some of the older writers held that it was due to the attacks of fungi, while others attributed it to the depredations of animal parasites. At pres- ent, however, it is the general belief of those who have given the mat- ter careful study that it is not, as a general thing, due to either of the foregoing causes, but is the result of certain physiological changes that take place in the tuber when the latter is grown under certain condi- tions. | The potato is in reality an underground stem, and, like similar parts of other plants, its outer covering consists of a thin, tough membrane, which serves as a protection to the more tender parts within. This en- veloping coat is made up of minute cells, the walls of which consist of a substance known as suberine, or cork. The corky membrane is pro- vided with numerous minute structures known as lenticels, by means of which an exchange of gases takes place between the interior and exterior of the tuber.2, In the presence of an excess of moisture the lJenticels ereatly increase in size, and often appear as small, woolly tufts, scattered here and there over the surface. At the same time the cork layer be- comes thickened at these points, and as a final result of this process small wart-like projections are formed. Where these occur the skin is weakened, and, if the conditions which favor this development continue, decay soon sets in. In its efforts to heal the wound thus produced the tuber gives rise to new layers of cork cells beneath the diseased parts, and, as a result of this continued dying of the outer cells and the form- ation of a new growth beneath, a scab is produced. Wherever the corky membrane is wounded the tuber immediately makes an effort to repair theimjury. It would seem reasonable, there- 1 See Bibliography. 2 Stahl, Bot., Zeit. 1873. 45 46 fore, to believe that in some cases at least the primary injury is due to the depredations of grubs, myriapods or other small animals,! or it may be brought about by the irritating or corroding action of some substance — in the soil. In any event the final result is practically the same as where the original damage is caused by an enlargement of the lenticels,. All who have thoroughly investigated the disease agree in the opin- ion that extreme humidity favors its development, but what influence beyond this the character of the soil has upon the malady is not posi- tively known. There have been many theories advanced havin ga bear- ing upon this question, but they are so conilicting that nothing of value can be derived from them. During the vear 1887 a number of trials were made at the New York Agricultural Experiment Station with a view of testing (1) influence of the soil, (2) effects of excessive moisture, (3) use of scabby seed, (4) effects of color of skin, (5) effects of fungicides, and (6) effects of chemical fertilizers and stable manure upon the devel- opment of seab. It was shown by these experiments that an excess of moisture and the use of fresh stable manure materially increased the number of scabby potatoes. The data obtained are summed up as fol- lows: ‘¢(1) The scab is not primarily caused by a fungus. (2) It is not due to the work of insects. *©(3) In nearly every instance an increased yield was accompanied by an increased percentage of scabby potatoes. “6(4) Any marked change in the rapidity of growth, either an increase or a decrease, tends to an increased production of scab. (5) A continuous growth from the time of first vegetation until the tubers are fully matured appears to be the condition least favorable to the production of scabby potatoes.” BIBLIOGRAPEY. Arthur: Sixth Report of the New York Agricultural Experiment Station, 1887, p. 344. Beckwith: Sixth Report of the New York Agricultural Experiment Station, 1887, p. 307. Frank: Krankheiten der Pflanzen, 1880, p. 140. Plowright : Gardener’s Chronicle, vol. 23, 1885, p. 60. Report of the Connecticut Agricultural Experiment Station, 1877, p. 67; 1886, p. 79. Sorauer : Handbuch der Pflanzen Krankheiten, Ist ed., 1884, p. 84; 2d ed., 1886, p. 227. 1 Mr. Alfred Rose, writing to the Rural New Yorker under date of March 7, 1885, says that he has experimented largely for forty years on potato culture, and he thinks the wire worm (Iulus) causes scab; that these ‘‘ worms” are more prevalent and the scab much worse on rather moist soils than on those light and sandy; that where scab prevails, the sooner the potatoes are dug the better, and that anything that for- wards the crop aids in escaping the scab. He has found that the use of lime slaked with water which has been saturated with salt and into which 2 pounds of sulphur have been stirred for each bushel of lime has effectually driven out these ‘‘ worms ”’ and prevented scab. He thinks that land infested with these worms should be plowed in the fall and have 250 pounds of kainit and 5 bushels of lime, prepared as above described, sown broadcast per acre and well cultivated in. on POTATO SCAB.! This disease, which is known to practical gardeners as scab or scurf, may be recognized by the brown, irregular concavities, with ragged, often upturned edges, that make their appearance in the tuber. These concave places spread gradually and become filled with brown, decayed re-aains of cells, excrements of mites, mycelium threads, ete. The deeper these places eat into the flesh of the healthy tuber the greater is its loss of sound tissue, and consequently of food value; but even if the scabby portions are not very deep and the decrease in food value unimportant, the potatoes affected with scab spots lose consider- ably in market value on account of their discased appearance. If the small projections on the edge of the scab are examined they will be found to consist of parenchyma cells, which once formed a part _of the healthy tissue of the tuber, but are now dark colored, and mostly without starch grains, in place of which the other cell contents are massed together in a spherical shape. From these dead, suberified cells there is either a gradual transition into the healthy, starch- bearing tissue, or they are separated from the latter by a layer of true tabular cork cells which have arisen, at the expense of the starch, from the cell layers bordering on the scab spot. The danger to the potato lies in the fact that as long as it remains in the ground the edges of the scab very often begin to degenerate again and the spot becomes deeper. If vhe potatoes are once dug, no further penetration of the seab spot can be discerned, even when buried for the winter. In case of early varieties that are gathered at different times, a section of the scab spots shows that they are no deeper in those po- tatoes dug at the end of the season than in the ones gathered some weeks earlier. I conclude from this circumstance that the extension of scab spots is not continuous,even while the potatoes remain in the ground, but is confined to different periods, and perhaps in many cases to one. IT consider the times when hard showers follow a long continued drought as such scab forming periods. This view is founded upon the changes which occur when a potato is brought into a very moist atmosphere or part of it into direct contact with dripping water. By this means the development of the lenticels becomes so increased from their normal condition that they are easily recognized by the naked eye as small, white, woolly warts. Nearly all parts of plants which are covered by a persistent cork layer possess interruptions on the surfacein form of len- ticels or bark-pores, in which there is a loose, corky tissue provided with air conducting intercellular spaces. This tissue usually arises beneath the stomata and shares in their physiological work, 7 e., the exchange of atmospheric air between the interior and exterior of the plant. In 1Translation of paper by P. Sorauer, Handbuch der Pflanzen-Krankheiten, 1386, D. 227. 48 the wild cherry,’ privet, elder, ash, and willow, the lenticels arise under one stoma, and in the black walnut aud poplar under several, by the separation of the parenchyma cells underneath and surrounding the breathing pores. In other plants—for example, barberry, broom-corn, eurrants, and other species of Ribes—tie lenticels originate in the cork cambium (phellogen). In the potato the rind is composed of tabular cork cells (Fig. 1 *), and under this, as well as below the stomata? the first beginnings of the lenticel formation oceur in the form of irregular cells containing but little protoplasm. (Fig. 1a). The formation of these cells is continually penetrating farther in, while the cells first formed absorb water, swell up, and burst the cork rind, thus forming the seab-producing lenticel, from which the loose cells within (Fig. 1 /) emerge in a whitish, moist, flour-like mass. These cells soon degener- ate, and the process of degeneration extends inward, so that the com- pact, united lenticel cells (fig. 1 v) must always be looked for deeper and deeper in the flesh, and the starch (Fig. 1 st) disappears propor- tionally from the surrounding tissue. We have, therefore, two processes to consider in the formation of scab; the first is the growth of lenticel cells, which proceeds until the discharge of the flour-like cell mass; and the second consists of the de- LE. Stahl, Entwicklungsgeschichte und Anatomie der Leuticellen, Bot. Zeit., 1873, Nr. 36-39. 2 Caspari in Sitzungsberichte der niederrheinischen Gesellschaft fiir Natur- und Heil- kunde, v. 8, Jan., 1857, cited in Bot. Zeit., 1857, p. 116. 49 generation of the cells in the interior of the lenticel, and the consequent, extension of cork formation into the interior of the tuber. That this hypertrophied formation of lenticels is dependent upon an unusual amount of moisture we conclude from the following facts: During a long continued season of wet weather the lenticels of alder trees stand out in the form of thick white protuberances; by immers- ing a piece of cherry stem in water a luxuriant growth of cork can be artificially produced ; finally, in ease of the potato, these cork warts may be actually produced by keeping the potato for a long time in moist air, Nobbe’ found, by a water culture of the potato, that the tubers grownin water produced small warts whilestill very young. These warts were caused by a local increase in growth in the cork formation, and were not present in tubers produced in the air. The view that superfluous moisture at an unseasonable time is the cause of the increased growth of cork that produces scab is indorsed by Caspari? and Schacht? as well by Frank. A later work by Stahl es- tablishes the fact that if transpiration 1s stopped lenticels develop beneath the stomata.? On the contrary, farmers assert that in the ma- jority of cases the adding of lime and marl to the soil, and manuring® with dung and street-sweepings causes the disease. Kuhn agrees with Wallroth*‘ in the opinion that the formation of scab is caused by a fun- gous growth (Hrysibe subterranea, Wailr.). On the other hand, iron, when present ina low state of oxidation, is considered injurious. In a field of potatoes at Newmarket three rows which had been marled with light colored earth mixed with iron oxide were found free from scab, while the rows that had received mar! that was dark colored from the presence of ferrocyanate of peroxide of iron were very scabby.® The assertion has frequently been made by practical farmers that fresh animal manure causes seab, especially in the thin-skinned varieties,° and that the disease makes its appearance after an application of soap- suds, while after a strong application of potash, which had proved very injurious at another time, the tubers were free from scab." 1 Versuchsstationen, 1864, p. 58. 2 Bot. Zeitung, 1857, p. 116. 3 Bericht an,d. Kel. Landes-Oecomomie-Collegium iiber die Kartoffelpflanze und deren Krankheiten, Berlin, p. 15. 4 Die Krankheiten der Pflanzen, 1880, p. 141. 5 Biedermann’s Centralbl., 1879, p. 714. 6 Landwirthsch. Zeitschr. f. Westphalen und Lippe, 1864. 7 Linnea, 1812, p. 332. Der Knollenbrand der Kartoffel, von Hofrath Dr. Wallroth. He says: ‘‘Themuch discussed disease of the potato known in economic writings under the name of potato scab, potato wart, scab disease, brown stain, and potato rot, I have known for a long time as a kind of vegetable rust (Uredo, Ustilago, and Cwoma of authors, Erysibe Theophr., Adans, Murr., Wallr., not DC.). 8 Landwirth, 1875, p. 352. 9Landwirth, 1875, p. 319 and 352. 10 Janovski, Kartofflenbauversuche. Prager Landw. Wochenbl., cited in Central bl. f. Agrik.-Chemie, 1876, I, p. 430. 11244—-No, 8——4 50 I believe that Wallroth’s opinion is erroneous. It is true we find fungous formations upon the dead parts, but none that attack healthy tissues. Many have objected to the view that scab is caused by adding lime, marl, or dung to the soil, but when it comes to final definite re- sults the reports vary in regard to secondary circumstances. For ex- ample, one person Says that seab made its appearance after the appli- cation of marl, but that the greatest development of the disease did not occur the first year,' during which the potatoes were perfectly healthy, but that the disease steadily increased during the ten following years. The same account claims that beets are similarly affected by marl. It is interesting to note that where the marl was not used the potatoes showed no scab. In Posnia, in the seventeenth century, marl was fre- quently added to the loose soil of large estates for the purpose of raising Medicago media. This was generally preceded by two crops of potatoes, which were always marled. In spite of this, scab never made its ap- pearance if the marl was drawn before winter and thoroughly mixed with the soil.2 The same experiment was recorded a year earlier in Saxony in the Zeitschrift des landwirthsch. Centralvereins (p. 219). Heiden * reports a very thorough experiment at Pommritz in regard to the use of lime. 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