S&3S3 THE GIFT OF FLORENCE V. V. DICKEY TO THE UNIVERSITY OF CALIFORNIA AT LOS ANGELES THE DONALD R. DICKEY LIBRARY OF VERTEBRATE ZOOLOGY THE FAUNA OF BRITISH INDIA, CEYLON AND BURMA. PUBLISHED UNDER THE AUTHORITY OF THE SECRETARY OF STATE FOR INDIA IN COUNCIL. EDITED BY W. T. BLANFOED. EEPTILIA and BATKACHIA. GEORGE A. BOULENGER. LONDON: TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET. CALCUTTA: I BOMBAY: THACKEK, SPIU'K, & CO. THACKER & CO., LIMITED. BERLIN : U. FIUEDLANDER & SOHN, 11 CARLSTEASSE. 1890. PRINTED BY TAYLOR AND FRANCIS, RED LION COUUT, FLEET STREET. QL 30") PKEFACE. Two classes of Vertebrata, the Reptiles and the Batrachians or Amphibians, are described in this volume of the ( Fauna of British India/ These two classes have often been associated in zoological works, and were, by many writers, until recently, not regarded as distinct. The study of both is usually prosecuted by the same observers, and no practical advantage would result from publishing the descriptions of the two separately. Mr. Boulenger, to whom Indian naturalists are indebted for the present work, has already classified and described all known Crocodiles, Tortoises, Lizards, Chamaeleous, and Batrachians in Catalogues of the British Museum that have appeared during the last eight years. Many of the characters of genera and species now published are taken from those Catalogues, but numerous additions have been made and several alterations have been inserted. Wherever the com- plete synonymy is given in the Catalogues it has been considered unnecessary, in the present work, to insert more than a reference to the original description of the species, to the page in the British Museum Catalogue where full details can be found, and to the general works on Indian Reptiles by Giinther and Theobald. 713616 "* The classification of the Snakes, which comprise nearly one half of the Reptilian species known to occur in India, is new, and all the descriptions of families, genera, and species have been prepared expressly for the present work. As there is no recent publication with a complete synonymy of the Ophidia, somewhat fuller references to the literature of the subject have been rendered necessary than in the other suborders of Reptiles and Batrachians. Two general works on Indian Reptiles have been published before the present volume. The first was ' The Reptiles of British India/ by Dr. A. Giinther, which appeared in quarto and was issued by the Ray Society in 1864; and the second, an octavo ( Descriptive Catalogue of the Reptiles of British India/ by Mr. W. Theobald, published in 1876. In the first, which was founded on the earlier publications of Russell, Cantor, Gray, Blyth, Jerdon, Kelaart, and others, largely supplemented by the author's own researches, the Batrachia were included ; but both the Batrachia and the Hydrophiinee or marine snakes were omitted in Mr. Theobald's book, a great part of which was virtually an abridgment of Giinther's, but with the numerous discoveries and observa- tions of Stoliczka, Beddome, Anderson, and of the author himself added to those of Giinther, many additions by the latter having been made after the publication of his large work on Indian Reptiles. In the present volume the principal additions have been due to the collections of Beddome in Southern India, Fea and Davison in Burma, and of Murray and the Editor of this work in Western India and Baluchistan. The limits of the fauna described in Giinther's ' Reptiles of British India ' wrere wider than those adopted in Theobald's work and in the present, and comprised all South-eastern Continental Asia. The area here accepted as that of the ' Fauna of British India ' has been defined in the Introduction to the Mammalia of the present series, and may be briefly described as British India with Burma and all other depen- dencies,, and with the addition of Ceylon— Baluchistan, Kash- mir, Ladak, and all Cis- Himalayan States being included, and Afghanistan, Tibet Proper, China, Siam, and the Malay Peninsula south of Tenasserim excluded. The following comparison of the number of species de- scribed, from within the limits adopted in the present work, in Giinther's ' Reptiles of British India 'with those enumerated in Theobald's Catalogue and in the present volume will serve roughly to indicate the additions to our knowledge of the Reptilia and Batrachia of India in the last quarter of a century. As already noticed, one subfamily of Snakes, the Hydrophiinre, was omitted by Theobald, Giinther, Theobald, Boulenger, REPTILIA. EMIDOSATJEIA (Crocodiles) .... CHELONI A (Tortoises & Turtles) SQTJAMATA. Lacertilia (Lizards) 1864. 4 .. 30 . . 95 1876. 4 54 173 1890. 3 43 225 lihiptoglossa (Chameleons) Ophidia (Snakes) .... .. 1 . . 180 1 225 1 264 310 457 536 BATRACHIA. ECAUDATA (Frogs and Toads). . CATJDATA (Newts &c.) APODA (CaDcilians) . . 37 0 2 •• 124 1 5 39 130 In cases where the numbers have been reduced the reduc- tion is due to the union of species formerly supposed to be distinct. The additions in the present volume are much more numerous than the preceding comparison indicates. By permission of the Trustees of the British Museum, through the kindness of Professor Flower and Dr. Giinther, VI PREFACE. several figures from the Catalogues of the National Collection prepared by Mr. Boulenger and from the Guide to the Reptile Gallery have been inserted in the present work. One cut lias also, by permission of the Zoological Society, been taken from its 'Proceedings/ The Fishes, Reptiles, and Batrachiaus of the f Fauna of British India ' have now appeared, with one volume of Birds and half a volume of Mammals, leaving two volumes of Birds and half a volume of Mammals still in hand. W. T. BLANFORD. July, 1890. SYSTEMATIC INDEX. REPTILIA "T Order I. EMYDOSAURIA . . 1 Faui. I. CROCODIUDJE . . 2 1 . Gavialis, Opp 3 1. gangeticus (Gm.) .... 3 2. Crocodilus, Law 4 1. porosus, Schn 4 2. palustris, Less 5 Order II. CHELONIA .... 6 Subord. I. THECOPHORA. 9 Superfam. A. Trionychoidea 9 Fam. I. TRIONYCHID^E 9 10 11 12 12 13 14 14 16 1. Trionyx, Geoff 1. subplanus, Geoff r 2. gangeticus, Cuv 3. leithii, Gray 4. hurum, Gray 5. formosus, Gray 6. phayrii, Theob 7. cartilagineus (Bodd.) 2. Pelochelys, Gray 15 1. cantoris, Gray 3. Chitra, Gray . .' 1. iiidica, Gray 4. Emyda, Gr 1. granosa 2. vittata, Ptrs. IS 10 16 16 17 17 3. scutata, Ptrs 18 Superfam. B. Cryptodira . . 18 Fam. II. TESTUDINUXE 18 1. Testudo, Z 19 1. .elongate, Blyth 20 2. leithii, Gthr 20 3. elegans, Schoepff 21 4. platynota, Blyth 22 5. emys, Schleg. 8f Midi. 22 2. Geoemyda, Gray 23 1. spinosa, Gray 25 2. grandis, Gray 25 3. depressa, Anders 25 3. Nicoria, Gray 26 1. trijuga (Scliweigg.) . . 27 2. tricarinata (Blyth) 28 4. Cyclemys, Bell 28 1. platynota, Gray 30 2. dhor, Gray 30 3. mouhoti, Gray 31 4. amboinensis (Daud.) 31 5. Bellia, Gray 32 1. crassicollis, Gray .... 32 6. Danionia, Gray 34 1. hitmiltonii, Gray 34 7. Morenia, Gray 35 1. oceUata(Z). #5.).... 35 2. petersii, Anders 36 8. Hardella, Gray 36 1. thurgi, Gray 36 9. Batagur, Gray 37 1. baska, Gray 38 10. Kachuga, Gray 38 1. lineata, Gray 40 2. trivittata (D. # B.) . . 41 3. dhongoka (Gray) 41 4. smitlii'i (Gray) 42 5. sylhetensis (Jerd.) .... 42 6. intermedia, Blanf. 43 7. tectum (Gray) 43 Fam. III. PLATYSTEUNID.S: . . 44 1. Platysternum, Gray .... 44 1. megacephalum, Gray . 44 SYSTEMATIC INDEX. Page Fam. IV. CHELONms: 47 1. Chelone, Brongn 48 1. mydas (L.) 48 2. imbricata (L.) 49 2. Thalassochelys, Fitz 49 1. caretta (L.) 50 Subord. II. A T H E c M Fam. V. SPHARGIDJE . 1. Dermochelys, Blainv 1. coriacea (L.) Order III. SQUAMAT A .... 52 Subord. I. LACERTILIA .. 52 Fam. I. GECKONID^: 54 1. Stenodactylus, Fitz 56 1. orientalis, Blanf. 57 2. lurnsdenii, Blgr 58 2. Alsophylax, Fitz 58 1. tuberculatus (Blanf.) 59 3. Gymnodactylus, Spix .... 59 1. fedtschenkoi, Strauch 61 2. scaber (Hiipp.) 62 3. brevipes, Blanf. 63 4. kachhensis, Stol 63 6. stoliczkee, Steind. 63 6. lawderanus, Stol 64 7. nebulosus, Bedd 64 8. jeyporensis, Bedd 65 9. deccanensis, Gthr. . . 66 10. albofasciatus, Blyr. . . 66 11. oldhami, Theob 67 12. triedrus, Gthr 67 13. frenatus, Gthr 68 14. khasiensis (Jerd.) 68 15. rubidus (Blyth) 69 16. pulchellus (Gray) 69 17. variegatus (Blyth) 70 18. fasciolatus (Blyth) .... 71 4. Agamura, Blanf. 71 1 . cruralis, Blanf. 71 5. Pristurus, Rilpp 72 1. rupestris, Blanf. 72 6. Gonatodes, Fitz 73 1. indicus (Gray) 74 2. wynadensis (Bedd.) . . 75 3. sisparensis (Theob.) . . 75 4. ornatus (Bedd.) 75 5. marmoratus (Bedd.). . 75 6. mysoriensis (Jerd.) . . 77 7. kandianus (Kel.) .... 77 8. gracilis (Bedd.) 78 9. jerdonii (Theob.) .... 78 10. littoralis (Jerd.) 79 7. Calodactylus, Bedd. 80 1. aureus, Bedd 80 8. Ptyodactylus, Gray .... 81 1. homolepis, Blanf. .... 81 9. Heinidactylus, Gray 82 1. reticulatus, Bedd. .... 84 2. gracilis, Blanf. 84 3. frenatus, D.$B 85 4. gleadovii, Murray. . . . 86 5. turcicus (L.) 87 6. persicus, Anders 88 7. maculatus, D. Sf B. . . 88 8. triedrus (Daud.) .... 89 9. subtriedus, Jerd. .... 90 10. depressus, Gray 90 11. leschenaultii, D. 8f B. 91 12. cocteei, D. $ B 92 13. giganteus, Stol. 92 14. bowringii (Gray) .... 93 15. karenorum (Theob.) . . 93 16. garnoti, I). $ B 94 17. platyurus (Schneid.) . . 95 10. Teratolepis, Gthr 96 1. fasciata (Blyth) 96 11. Gehyra, Gray 96 1. mutilata ( Wiegm.) . . 96 12. Lepidodactylus, Fitz 97 1. ceylonensis, Blgr 98 2. aurantiacus (Bedd.) . . 98 3. lugubris (D. $ B.) . . 99 13. Hoplodactylus, Fitz 100 1. duvaucelii (D.$B.).. 100 2. anainallensis (Gthr.). . 101 14. Gecko, Laur 101 1 . verticillatus, Laur. . . 102 2. stentor (Cantor) 103 3. monarchus (D. # B.) . 103 15. Ptychozoou, Fitz 104 1. hornalocephalum (Crev.) 104 16. Phelsuma, Gray 105 1. andamanense, Blyth . . 106 Fam. II. EUBLEPHABID.E . . 107 1. Eublepharis, Gray 107 1. hardwickii, Gray .... 107 2. macularius (Blyth) . . 108 SYSTEMATIC INDEX. Fam. III. AGAMID^E , Page 109 1. Draco, Linn Ill 1. maculatus (Gray) 112 2. blanfordii, Blgr 112 3. dussumieri, D. $ B. . . 113 4. teeniopterus, Gthr. . . 114 2. Sitana, Cuv 114 1. ponticeriana, Cuv 114 3. Otocryptis, Wiegm 115 1. bivittata, Wiegm 115 2. beddomii, Blgr 116 4. Ptyctolaemus, Ptrs 116 1. gularis, Ptrs 117 5. Cophotis, Ptrs 117 1. ceylanica, Ptrs 118 6. Ceratophora, Gray 118 1. stoddartii, Gray 119 2. tennentii, Gthr 120 8. aspera, Gthr 120 7. Lyriocephalus, Merr 121 1. scutatus (L.) 121 8. Gonyocephalus, Kaup .... 122 1. subcristatus (Blyth) . . 122 2. humii (Stol.) 123 3. belli! (D. # B.) 123 4. grandis (Gray) 124 9. Acanthosaura, Gray .... 124 1. armata (Gray) 125 2. crucigera, Blyr 125 3. lamnidentata, Blgr. . . 126 4. minor (Gray) 127 5. kakhienensis (Anders.) 127 6. major ( Jerd.) 128 7. tricarinata (Blyth) . . 129 10. Japalura, Gray 129 1. variegata, Gray 130 2. planidorsata, Jerd. 130 11. Salea, Gray 131 1. horsneldii, Gray .... 131 2. anamallayana (Bedd.) . 132 12. Calotes, Cuv. 132 1. microlepis, Blgr 134 2. cristatellus (Kuhl) 134 3. jubatus (D. 8?B.) 135 4. versicolor (Daud.) . . 135 5. maria, Gray 136 6. jerdonii, Gthr 137 7. emrna, Gray 137 8. mystaceus, D. % B. .. 138 9. grandisquamis, Gthr. . 138 10. nemoricola, Jerd. .... 139 11. ceylonensis, Blyr 139 12. liolepis, Blgr 140 13. ophioinachus (Merr.) . 140 Page 14. nigrilabris, Ptrs 141 15. liocephalus, Gthr 141 16. rouxii, D. 8f B 142 17. ellioti, Gthr 142 18. fete, Blgr 143 13. Charasia, Gray 144 'I. dorsalis (Gray) 144 2. blanfordiana, Stol. . . 145 3. ornata (Blyth) 145 14. Agama, Daud. 146 1. isolepis, Blgr 147 2. rubrigularis (Blanf.). . 147 3. tuberculata, Gray 148 4. dayana (Stol.) 148 5. hirnalayana (Steind.). . 149 6. agrorensis (Stol.) 149 7. melanura (Blyth) 150 8. lirata (Blanf.) 150 9. nupta, De Fil 151 10. caucasica (Eichw.) . . 151 15. Phrynocephalus, Kaup . . 152 1. olivieri, D. # B 153 2. theobaldi, Blyth .... 153 3. caudivolvulus (Pall.).. 154 4. ornatus, Blgr 154 5. maculatus, Anders. . . 155 6. luteoguttatus,.%r. . . 155 16. Liolepis, Cuv 156 1. belliana (Gray) 156 17. Uromastix, Merr 157 1. hardwickii, Gray .... 157 Fain. IV. 158 1. Ophisaurus, Daud. 159 1. gracilis (Gray) 159 Fam. V. VARANID^: 160 1. Varanus, Merr 161 1. griseus (Daud.) 163 2. tiavescens (Gray) 164 3. bengalensis (Daud.) . . 164 4. nebulosus (Gray) .... 165 5. dumerilii (Schleg.) . . 165 6. salvator (Laur.) 106 Fam. VI. LACERTID^; 167 1. Tachydromus, Daud 168 1. sexlineatus, Daud. ... 169 2. tachydromoides (Schkg.) 169 SYSTEMATIC INDEX. Page 2. Aoanthodactyliis, Wieym. 170 1, 'cantoris, Gthr 170 2, micropholis, Blanf. . . 171 3. Cabrita, Gray 171 1. leschenaultii (M.- Edw.) 172 2. jerdonii, Bedd. 173 4. Ophiops, Men&r 173 1, ierdonii, Blyth 174 2. .beddomii (Jerd.) 174 3, elegans, Menetr 175 4. microlepis, Blanf.. . . . 175 5. Eremias, Wieym. .' 176 1. guttulata (Licht.) 177 2. brevirostris (Blanf.).. 177 3. velox (Pall.) 178 4. fasciata, Blanf. 179 6. Scapteira, Wiegm 179 1. acutirostris, Blyr 179 Fam. VII. SCINCIDJE 180 1. Mabuia, Fitz 183 1. bibronii (Gray) 184 2. doriae, Blgr 184 3. dissimilis (Hallow.) . . 185 4. septemtteniata (Reuss) 185 5. innotata (Blanf.) .... 186 6. novemcarinata {Anders.) 187 T.'beddoruii (Jerd.) 187 8. vertebralis, Blyr 188 9. carinata (Schn.) 188 10. macularia (Blyth) . . 189 11. rugifera (Stol.) 190 12. moltifasciata(Zu^).. 191 13. tytieri, Blgr 191 14. quadricarinata, Blyr. . 192 2. Lygosoma, Gray 192 T. indicum (Gray) 195 2. zebratum, Blgr 195 3. iriaculatum (Blyth) . . 196 4. dussumieri, D.$B... 197 5. olivaceum (Gray) 197 6. kakhienense, Blyr. . . 198 7. melanostictum, Blyr. . 199 8. sikkimense (Blyth) . . 199 9. Mmalayanum (Gthr.) . 200 10. doriae, "Blgr 201 11. ladacense (Gthr.) 201 12. laterimaculatum, Blyr. 202 13. bilineatum (Gray) ..202 14. beddomii, Blyr 203 16. travancoricum (Bedd.) 204 I Page 10. inacrotyujpanum(£fo/.) 204 17. maerotis (Steind.) 205 (Mocoa formosa, Blyth.) 18. taprobanense (Kel.) . . 200 19. fallax, Ptrs 206 20. lineolatum (Stol.) 207 21. coinotti, Blgr 207 22. albopunctatum (Gray) 208 23. punctatum (L.) 208 24. guentheri (Ptrs.) 209 25. fea3, Blgr 210 26. cyaneUum (Stol.) 210 27.' anguinum (Theub.) . . 211 28.' calamus, Blyr 211 29. lineatum (Gray) 212 (Euprepes halianus, Nevill.) 3. Ablepharus, Fitz 213 1. brandtii, Strauch 213 2. grayanus (Stol.) 214 4. Ristella, Gray 214 1. rurkii, Gray 215 2. travancorica, Bedd. . . 216 3. guentheri, Blgr 216 4. beddomii, Blgr 216 5. Tropidophorus, D. 8f B. . . 217 1. berdmorii (Blyth) . . 217 2. yunnanensis, Blyr. . . 217 6. Euineces, Wieym 218 1. scutatus (T/ieob.) 218 2. taeniolatus (Blyth) .. 219 3. sclineideri (Daud.) . . 219 4. blythianus (Anden.).. 220 7. Scincus, Laur 220 1. arenarius, Murr 221 8. Ophiomorus D. $ B 221 1. tridactylus (Blyth) . . 222 2. blanfordii, Blyr 222 9. Clialcides, Laur 223 1. ocellatus (Forsk.) 224 2. pentadactylus (Bedd.) 224 10. Sepophis, Bedd. 225 1. punctatus, Bedd. 225 11. Chalcidoseps. Blgr 226 1. thwaitesii (Gthr.) 226 12. Acontias, C'uv 226 1. burtonii (Gray) 227 2. inonodactylus ( Gray) . 228 3. layardii, Kel. 228 4. sarasinorurn, F. Mull. . 228 Fam. VIII. DIBAMIDJE 229 1. Dibamus, 1). $ B 229 1. novae-guineae, D. Sf B. 230 SYSTEMATIC INDEX. ii Page Subord. II. RHIPTOGLOSSA. 230 Fam. I. CHAMJELEONTID^E. . . 230 1. Chamseleon, Law 232 1. calcaratus, Merr 232 Subord. III. O P H i D i A. , 232 Fam. I. TYPHLOPIDJE 234 1. Typhlops, Schneid 1. braniinus (Daud.). . 2. beddomii,^//- 3. leucomelas, Blgr. . . 4. jerdoni, Blgr 5. oatesii, Blyr 6. diardi, Sclileg 7. bothriorhynchus, Gthr 8. porrectus, Stol 9. theobaldiauus, Stol. 10. mirus, Jan 11. andamanensis, Stol. 12. tenuicollis(P*rs.).. 13. acutus (D. $ B.) . . Fam. II. GLAUCONIID^; 1. Glaucoma, Gray 1 . blanfordii, Blgr 235 236 237 237 238 238 238 239 240 240 240 241 241 241 242 243 243 Fam. HI. BOIDJE 244 Subfam. I. Pythonince 245 1. Python, Daud 245 l". reticulatus (Schneid.). . 246 2. molurus (L.) 246 Subfam. II. Boinee 246 2. Gongylophis, Wagl. 246 1 . conicus (Schneid.) .... 247 3. Eryx, Daud 247 1. johnii (Russ.) 248 Fam. IV. ILYSIID.E 242 1. Cylindrophis, Wagl. 249 1. rufus (Laur.) 251 2. maculatua (L.) 251 Fam. V. UBOPELTID^: 251 1. Uropeltis, Cuv 253 1. grandis, Kel 254 2. Rhinophis, Hempr 254 1. oxyrhynchus (Schneid.) 255 2. punctatus, M till 255 3. planiceps, Ptrs 255 4. trevelyanus (Kel.) 256 5. sanguineus, Bedd. .... 256 6. blythii, Kel. 253. 3. Silybura, Gray 257 1. melanogaster (Gray). . 258 2. pulneyensis, Bedd. . . 260 3. grandis (Bedd.), 261 4.. petersii, Bedd 261 5. maculata, Bedd. .... 261 6. liura, Gthr 262 7. ocellata, Bedd 262 8. nigra, Bedd. 263 9. nitida, Bedd. 263 10. dindigalensis, Bedd... 264 11. broughami, Bedd 264 12. macrorhynchus, Bedd. 264 13. beddomii, Gthr 265 14. ellioti, Gray 265 15. phipsonii, Mason .... 266 16. rubrolineata, Gthr. . . 266 17. .myhendrfe, Bedd 267 18. madurensis, Bedd. . . 267 19. rubromaculata, Bedd. . 268 20. arcticeps, Gthr 268 21. brevis, Gthr 268 22. macrolepis, Ptrs 269 4. Pseudoplectrurus, Blgr. . . 270 1. canaricus (Bedd.) .... 270 5. Plectrurus, D. $ B 270 1. perroteti, D. $ B 271 2. davisonii, Bedd 271 3. guentheri, Bedd. 271 4. aureus, Bedd 272 6. Melanophidium, Gthr. . . 272 1. wynadense (Bedd.) . . 272 2. punctatum, Bedd. 273 3. bilineatum, Bedd. 273 7. Platyplectrurus, Gthr. . . 273 1. trilineatus (Bedd.) .... 274 2. madurensis, Bedd 274 3. sanguineus (Bedd.) . . 274 Fam. VI. XENOPELTID^. . . 275 1. Xenopeltis, Reinw 276 1. unicolor, Reinw 276 SYSTEMATIC INDEX. Fam. VII. COLUBRIDJE Series A. Aglypha . . . ..s* 279 Subfam. I. Colubrince 279 1. Cakmaria, Boie 281 1. pavimentata, D. $ B. . 282 2. oatenata, Blyth 282 2. Xylophis, Bedd 283 1. perroteti (D. $ B.) .. 283 2. 8tenorhvnchus((37Ar.) . 284 3. Trachischiuiu, Gthr 284 1. fuscum (Blyth) 285 2. guentheri, Blgr 285 3. tenuiceps (Blyth) 286 4. monticola(ftmi5.) 286 5. rubriventer (Jerd.) . . 286 4. Blythia, Theob 287 1. reticulata (Blyth) .... 287 5. Aspidura, Wagl. 288 1. brachyorrhos (Boie) . . 289 2. copii, Gthr 289 3. guentheri, Fergus. . . 290 4. trachvprocta, Cope . . 290 6. Haplocercus, Gthr 290 1. ceylonensis, Gthr 291 7. Lycodon, Boie 291 1. striatus (Shaw) 292 2. jara (Shaw) 292 3. anamallensis, Gthr. . . 293 4. travancoricus (Bedd.) . 293 6. aulicus (L.) 294 6. atropurpureus (Cant.) 294 7. septentrionalis (Gthr.) 295 8. fasciatus (Anders.) . . 295 9. gammiei (Blanf.) 296 10. carinatus (Kuhl) 296 subfuscus, Cant 297 sanguiventer ( Cant.) . . 297 8. Hydrophobus, Gthr 297 1." nympha (Daud.) 298 2. gracUis (Gthr.) 298 3. davisonii (Blanf.). ... 299 9. Pseudocyclophis, Bttg. . . 299 1. walteri, Bttg 300 2. olivaceus (Bedd.) .... 300 3. bicolor (Blyth) 300 10. Polyodontophis, Blgr. . . 301 1. collaris (Gray) 302 2. subpunctatus (D. 8f B.) 303 3. Sagittarius (Cant.) 303 4. bisarigatus (Gthr.) . . 304 PaKe 11. Ablabes, D. $ B 304 1. calamaria (Gthr.) 305 2. scriptus, Theob 305 3. frenatus (Gthr.) 306 4. dorise, Blgr 306 5. rappii, Gthr 307 6. nicobariensis, Stol. 307 7. porphyraceus (Cant.) . 308 12. Coronella, Laur 308 1. brachyura (Gthr.) 309 13. Simotes, D. $ B 309 1. splendidus, Gthr 310 2. cvclurus (Cant.) 311 3. albocinctus ( Cant.) . . 312 4. violaceus (Cant.) 312 5. octolineatus (Schn.) ..313 6. arnensis (Shaw) 314 7. beddomii, Blgr 314 8. theobaldi, Gthr 315 9. cruentatus, Gthr 315 10. torquatus, Blgr 316 1 1 . plauiceps, Blgr 316 14. Oligodon, Boie 317 1. venustus (Jerd.) 317 2. travancoricus, Bedd. . . 318 3. affinis, Gthr 318 4. brevicauda, Glhr 319 5. dorsalis (Gray) 319 6. teinpletonii, Gthr 320 7. sublineatus, D.$B.. 320 8. ellioti, Gthr 321 9. subgriseus, D.fyB... 321 15. Lytorhynchus, Ptrs 322 1. paradoxus (Gthr.) 323 16. Zamenis, Wagl 323 1. korros (Schleg.) 324 2. mucosus (L.) 324 3. ventrimaculatus ( Gray) 325 4. ladacensis, Anders. . . 326 5. kareKnii (Brandt) .... 326 6. gracilis, Gthr 327 7. fasciolatus (Shaw) 327 8. diadema (Schleg.) 328 9. arenarius, Blgr 329 17. Zaocys, Cope 329 1. niorromarginatus (Blyth) 329 18. Coluber, L 330 1. heleua, Daud 331 2. reticularis, Cant 332 3. hodgsonii (Gthr.) .... 332 4. tseniurus (Cope) 333 5. radiatus, Schleg 333 6. melanurus, Schleg 334 SYSTEMATIC INDEX. 7. prasiuus, Blyth 8. frenatus (Gray) .... 9. oxycephalus, Boie .... 19. Xenelaphis, Gthr Page 334 335 335 336 336 337 337 337 338 338 339 339 339 340 340 341 343 343 344 344 345 345 346 346 347 347 348 348 349 350 350 350 351 352 352 352 353 353 354 354 355 355 355 356 356 357 358 359 359 360 Page 5. multimaculata, Schleg. 360 6. hexagonotus, Blyth . . 361 7. cyanea (D. $ B.) 361 8. forsteni(D. 8fB.) .... 362 29. Elachistodon, Reinh 362 1. westermanni, Reinh. . . 363 30. Psammodynastes, Gthr... 363 1. pulverulentus (Boie). . 363 31. Psammophis, Boie 365 1. hexagonotus (Cant.). . 20. Dendropnis, Boie 1. grandoculis, Blyr 2. pictus, Gm 3. subocularis, Blgr 4. bifrenalis, Blyr 1. leithii, Gthr 365 5. caudolineolatus, Gthr, . 21. Dendrelaphis, Blgr. .... 1. caudolineatus (Gray) . 22. Pseudoxenodon, Blyr. . . 1. macrops (Blyth) 23. Tropidonotus, Kuhl 2. condanarus (Metr.) . . 366 3. longifrons, Blgr 366 32. Dryophis, Boie 367 1. perroteti (D. # B.) . . 368 2. dispar (Gthr.) 368 3. fronticinctus, Gthr. . . 368 4. prasinus, Boie . . 369 1. modestus, Gthr 2. platyceps, Blyth .... 3. khasiensis, Blgr 5. mycterizans (Daud.) . . 370 6. pulverulentus (D.Sf B.) 371 33. Chrysopelea Boie 371 5. parallelus, Blyr 6. chrysargus, Schleg . . . . 7. nigrocinctus, Blyth . . 1. ornata (Shaio) . . 371 Subfam. IV. Homalopsina . . 372 34. Homalopsis, Kuhl 373 1. buccata (L.) . . 374 9. subrnimatus, Schkg. . . 10. hinmlayanus, Gthr. . . 11. monticola, Jerd 12 stolatus (L ) 35. Cerberus, Cuv. . . . 374 1. rhynchops (Schn.) . . 374 36. Hypsirhina, Wagl. 375 13 pi^cator (Schn ) 14 bellulus Stol 1. plumbea (Boie) 376 15. sancti-johannis, Blgr. . 16. punctulatus, Gthr. . . 17. plumbicolor, Cant. . . . angusticeps, Blyth . . 24. Helicops, Wayl 2. enhydris (Schn.) 376 3. blanfordii, Blgr 377 4. sieboldii (Schleg.) .... 377 37. Fordonia, Gray 378 1. leucobalia (Schleg.) . . 378 38. Gerardia, Gray 379 1. schistoaus (Daud.) . . 25. Xenochrophis, Gthr 1. prevostiana (Eyd. $ Gerv.) . 379 1. cerasogaster (Cant.) . . Subfam. II. Acrochordince . . 26. Stoliczkaia, Jerd. 1. khasiensis, Jerd 27. Chersydrus, Cuv 39. Cantoria, Gir 380 1. violacea, Gir 380 40. Hipistes, Gray 381 1. hydrinus (Cant.) .... 382 Series C. Proteroglypha . . 382 Subfam. V. Elapince 382 1. granulatus (Schn.) . . Series B. Opisthoglypha . . Subfani. III. Dipsadince 28. Dipsas, Boie 41. Callophis, Gray 383 1. trimaculatus (Daud.) . 384 2. maculiceps, Gthr 384 3. nigrescens, Gthr 384 4. macclellandii (Reinh.) 385 6. bibronii (Jan) 386 42. Adeniophis, Ptrs. . . 386 1. trigonata (Schn.) 2. barnesii, Gthr 3. ceylonensis, Gthr 4. gokool, Gray 1. intestinalis (Laur.) . . 386 SYSTEMATIC INDEX. 43. Bungarus, Daud 387 1. fasciatus (Schn.) 388 2. ceylonicus, Gthr 388 3. cfflruleus (Schn.) 388 4. bungaroides (Cant.) . . 389 5. lividus, Cant 389 44. Naia, Lour 390 1. tripudians, Merr 391 2. bungarus, Schleg 392 Subfam. VI. Hydrophiince . . 393 45. Platurus, Daud 394 1. laticaudatus (L.) 395 2. colnbrinus (Schn.) 395 46. Enhydris, Merr 396 1. curtus (Shaw) 396 2. hardwickii(G'ray) 397 47. Hydrus, Schneid 397 1. platurus (L.) 397 48. Hydrophis, Daud. 398 1. schistosus, Daud. 399 2. cserulescens (Shaw) . . 400 3. nigrocinctus, Daud. . . 401 4. mamillaris (Daud.) . . 401 5. spiralis (Shaw) 401 6. latifasciatus, Gthr. . . 401 7. coronatus, Gthr 402 8. torquatus, Gthr 402 9. qbscurus, Daud. 403 10. fasciatus (Schn.) 404 11. gracilis (Shaw) 404 12. cantoris, Gthr 405 49. Enhydrina, Gray 405 1. yalakadien (Boie) 406 50. Distira, Lacep 407 1. stokesii (Gray) 408 2. jerdonii (Gray) 408 3. robusta (Gthr.) 409 4. tuberculata (Anders.) 409 5. cyanocincta (Daud.) . . 410 6. b*i tuberculata (Ptrs.) . 411 7. ornata (Gray) 411 crassicollis (Anders.) . 411 8. lapemidoides (Gray) . . 412 9. yiperina (Schmidt) 413 Fam. VIII. AMBLYCEPHALIDJE .... 414 1 . Amblycephalus, Kiihl. ... 414 1. monticola (Cant.) 415 2. l«eyis, Boie 415 3. andersonii, Blgr 416 4. modestus (Theob.) 416 5. macularius (Theob.) . . 416 Fam. IX. VIPERID^E Subfam. 1. Viperince, 1. Azemiops, Blgr. 1. fese, Blgr. 2. Vipera, Laur l.russellii (Shaw) 2. lebetina (L.) 8. Echis, Merr 1. carinata (Schn.) . . . Page 417 418 418 418 419 420 421 422 Subfam. II. Crotalince 423 4. Ancistrodon, Beauv. . . . 1. hinialayanus (Gthr.) 2. hypnale (Merr.) . . . 5. Trimeresurus, Lacep. . . . 1. monticola, Gthr. . . . 2. strigatus, Gray 3. jerdonii, Gthr 4. mucrosquamatus (Cant.) 5. cantoris (Blyth) , 6. purpureomaculatus (Gray) 7. gramineus (Shaw) 8. anamallensis, Gthr. . . 9. trigonocephalus (Daud.) 10. macrolepis, Bedd 423 424 424 425 426 427 427 428 429 429 430 431 431 BATRACHIA 432 Order I. ECAUDATA. . . . 432 Suborder I. PHANEROGLOSSA. . 433 Series A. Firmisternia 433 Fam. I. RANIDJE 435 1 . Oxyglossus, Tsch 43(5 l.'lima (Gravh.) 436 2. laevis, Gthr 437 2. Rana, L 438 1 . hexadactyla, Less 441 2. cyanophlyctis, Schn. . . 442 3. corrugata, Ptrs 443 4. kuhlii, D. $ B 443 5. laticeps, Blgr 444 6. khasiana (Anders.) . . 444 7. liebigii, Gthr 445 8. sternosignata, Murr . . 445 SYSTEMAT Page 9 fees Blgr. . . 446 1C 1XDBX. 9. cruciger (Blyth) .... 10. eques (Gthr ) .... XV Page 476 476 477 477 477 478 478 479 479 480 480 481 481 482 483 483 484 484 485 485 485 486 486 •187 487 487 488 488 489 489 489 490 490 491 491 491 492 493 493 494 494 494 495 495 496 497 497 497 498 498 Ana 10. andersonii, Blgr. . . 447 1 1 dorise, Blgr 447 11, nasutus (Gthr.) 12. reticulatus (Gthr.) . . 13. dubius, Blgr 12. macrodon, D. <§• B. 448 13, verrucosa, Gthr. . . 448 14. tigrina, Daud 449 15. limnocharis, Wiegm 450 16. rufescens (Jerd.) .... 451 17. breviceps Schn. . . 451 14, nanusfG-7/i/-.) 15. jerdonii (Gthr.) 16. pleurostictus (Gthr.) . 17. microtvmpanum '(Gthr.) 18, stictomerus (Gthr.) . 19. fergusonii, Blgr 20. cavirostris (Gthr.) . . 8. Ixalus, D. # B. . . 18. dobsonii, Blgr 452 19. strachani (Murr.) .... 452 20. leithii, Blgr 453 21. beddomii (Gthr.) 453 22. semipaluiata, Blgr. . . 454 23. leptodactyla, Blgr. .. 454 . 24. diplosticta (Gthr.) . . 455 25. pbrynoderma, Blgr. . . 455 26. macrodactyla (Gthr.) . 455 27. gracilis, Gravh 456 1. o.xyrbynchus, Gthr. . . 2, leucorhinus, Mart. . . 3. schmardanus(Xe/.) .. 4. nasutus, Gthr. .... §. hvpomelas, Gthr 28. malabarica, D.8f B... 456 29 lateralis Blgr. . . 457 6. vittatus, Blgr 7. pulcher, Blgr 8. femoralis, Gthr 9. beddomii, Gthr 10. adspersus, Gthr 11. variabilis, Gthr 12 si°"natus, Blgr 30. curtipes, Jerd 458 31 tytleri ( Theob ) 458 32. nicobariensis (Stol.) . . 459 33. temporalis (Gthr.) . . 459 34. rnargariana (Anders.) 459 35. humeralis, Blgr 460 36. erytbrsea (Schleg.) . . 460 37. monticola (Anders.) . . 461 38. livida (Blyth) 462 39. latopalmata, Blgr. . . . 462 40. himalayana, Blgr. . . . 463 41. formosa (Gthr.) 463 3. Micrixalus, Blgr 464 1 saxicola (Jerd ) 465 13. flaviventris, Blgr 14. chalazodes, Gthr 15. gland ulosus, Jerd. . . . Fam. II. ENGYSTOMATID-ZE . . 1. Melanobatracbus, Bedd. . 1. indicus, Bedd. . 2. Calopbrynus, Tsch 2. sarasinoruui (F. Mull.) 465 3. opistborhodus (Gthr.) 465 4. fuscus (Blgr.) 466 5. silvaticus (Blgr.) 466 4. Nyctibatracbus, Blgr. . . 467 1. pygmseus (Gthr.). . . . 467 1 . pleurostigma, Tsch. . . 3. Microhyla, Tsch 2. ornata (D. # B.) 3. berdmorii (Blyth) 4. Callula, Gray 1. obscura, Gthr 5. Nannobatracims, Blgr. . . 468 1. beddomii, Blgr 468 6 Nannopbrys Gthr 469 2. pulcbra, Gray 3. macrodactyla, Blgr. . . 4. variegata, Stol. 5. triangularis, Gthr. . . 5. Cacopus, Gthr. . 1. cevlonensis, Gthr. . . 469 2. giientberi, Blgr 469 7. Kbacopborus, Kuhl 470 1. niaximus, Gthr 472 2. bimaculatus, Blgr. . . 472 3. malabaricus, Jerd. . . . 473 4. beddomii, Blgr 473 5 lateralis, Blgr 473 1. systoma (Schn.) 2. globulosus, Gthr 6. Glyphoglossus Gthr 1. molossus, Gthr Fam. III. DYSCOPHID^; 1 Calluella, Stol . . 6. tuberculatus (Anders.) 474 7. leucoiuystax (Gravh.). 474 XVI SYS1 Series B. Arcifera PEMAT 4*99 499 500 500 500 501 502 602 502 503 503 504 504 504 505 505 505 507 507 507 508 508 509 509 509 1C INDEX. Fain. VI. PELOBATIDJE . . . 1. Leptobrachium, Tsch.. . 1. monticola (Gthr.) . . 2. hasseltii, Tsch Page .. 510 .. 510 .. 510 .. 511 Fam. IV. BUFONHXS: 1. Nectophryne, Buchh. fy Ptrs 1. tuberculosa (Gthr.) . . 2. Bufo, Laur. 3. carinense, Blgr. . 4. fese, Blgr Order II. CAUDATA. . . . Fam. I. SALAMANDIUDJE . . . . 611 .. 512 . . 513 .. 513 513 2. kelaartii Gthr. . . 3. hololius, Gthr 5. beddomii, Gthr 6. latastii, Blgr 7. viridis, Laur 8. andersonii, Blgr 9. olivaceus, Blanf. .... 10. himalayanus, Gthr. . . 11. microtympanum, Blgr. 12. melanostictus, Schn. . . 13. parietalis, Blgr 14. biporcatus, Gravh 15. asper, Gravh 1. verrucosus, Anders. Order III. APOD A .. 514 .. 515 Fam. I. C./ECILIID^E .. 515 515 1. glutmosus (L.) 2. monochrous (Blkr.) 2. Urseotyphlus, Ptrs 1. oxyurus (D. $ B.) 2. malabaricus (Bedd.) 3. Gegenophis, Ptrs 1. carnosus (Bedd.) . . .. 515 .. 517 .. 517 .. 517 .. 518 .. 518 .. 518 1. sikkimensis (BlytK) . . Fam. V. HYLIDJE 1. Hyla, Laur 1. annectens (Jerd.) .... LIST OF THE PRINCIPAL WORKS QUOTED IN THE SYNONYMY. A. M. N. H. Annals and Magazine of Natural History. London, 1838-1889. Anders. An. Zool. Res. Anatomical and Zoological Eesearches, comprising an Account of the Zoological Results of the two Expeditions to Western Yunnan in 1868 and 1875. By John Anderson. London, 1879. Blanf. E. Pers. Eastern Persia, an Account of the Journeys of the Persian Boundary Commission, 1870-71-72.— Vol. II. The Zoology and Geology. By W. T. Blanford. London, 1876. Bouleng. Cat. Batr. Grad. Sec. Catalogue of the Batrachia Gradientia s. Caudata and Batrachia Apoda in the Collection of the British Museum. 2nd Edition. By G. A. Boulenger. London, 1882. Bouleng. Cat. Batr. Sal. Catalogue of the Batrachia Salientias. Ecaudata in the Collection of the British Museum. 2nd Edition. By G. A. Boulenger. London, 1882. Bouleng. Cat.Chelon. &c. Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum. New Edition, by G. A. Boulenger. London, 1889. Bouleng. Cat. Liz. Catalogue of the Lizards in the British Museum. 2nd Edition, by G. A. Boulenger. 3 yols. London, 1885-1887. Daud. Kept. Histoire Naturelle des Reptiles. By F. M. Daudin. 8 vols. Paris, 1802-1803. Dum. So Bibr. Erp. Gen. Erpetologie Generate ou Histoire naturelle complete des Reptiles. By A. M. C. Dumeril et G. Bibron. 9 vols. Paris, 1834-1854. Fayrer, Thanatoph. The Thauatophidia of India, being a Description of the Venomous Snakes of the Indian Peninsula, with an account of the influence of their poison on life, and a series of experiments. 2nd Edition, by J. Fayrer. London, 1874. Gray, 111. Ind. Zool. Illustrations of Indian Zoology, chiefly selected from the collection of Major-General Hardwicke. By J. E. Gray. 2 TO!S. London, 1830-1834. Gray, Cat. Liz. Catalogue of the Specimens of Lizards in the Collection of the British Museum. By J. E. Gray. London, 1845. Gray, Cat. Sh. Kept. i. Catalogue of Shield Reptiles in the Collection of the B ritish Museum.— Part I. Testudinata. By J. E. Gray. London, 1855. Gray, Cat. Sh. Kept. i. Suppl. Supplement to the Catalogue of Shield Reptiles.— Part I. Testudinata. By J. E. Gray. London, 1870. Gray, Cat. Sn. Catalogue of the Specimens of Snakes in the Collection of the British Museum. By J. E. Gray. London, 1849. 6 XV1U WORKS QUOTED. Gray, Cat. Tort. Catalogue of the Tortoises, Crocodiles, and Amphis- banians in the Collection of the British Museum. By J. E. Gray. London, 1844. Gunth. Cat. Batr. Sal. Catalogue of the Batrachia Salientia in the Col- lection of the British Museum. By A. Giinther. London, 1858. Gunth. Cat. Col. Sn. Catalogue of the Colubrine Snakes in the Collection of the British Museum. By A. Gimther. London, 1858. Gunth. R. B. I. The Reptiles of British India. By A. Giinther. London (Ray Society), 1864. J. A. S. B. Journal of the Asiatic Society of Bengal Calcutta, 1831-1889. Jan, Icon. Ophid. Iconographie G6nerale des Ophidiens. By G. Jan. Milan, 1860-1881. Kelaart, Prod. Prodromus Faunae Zeylanicse ; being Contributions to the Zoology of Ceylon. By E. F. Kelaart. Ceylon, 1852. Laur. 83711. Rept. Specimen medicum, exhibens Synopsin Reptilium emenda- tum. By J.N. Laurenti. Vienna, 1768. Linn. Syst. Nat. Systema Naturae. Ed. duodecima reformata. By C. A. Linne. Stockholm, 1776. MB. Ak. Berl. Monatsberichte der koniglich Preussischen Akademie der Wissenschaften zu Berlin. Berlin, 1856-1881. Merr. Tent. Tentamen Systematis Amphibiorum. By B. Merrem. Marburg, 1820. Murray, Zool. Sind. The Vertebrate Zoology of Sind. By James A. Murray. London and Bombay, 1884. P. A. Si B. Proceedings of the Asiatic Society of Bengal. Calcutta, 1865- 1887. P. Z. S. Proceedings of the Zoological Society of London. London. 1830- 1889. Russell, Ind Serp. An Account of Indian Serpents, collected on the Coast of Coromandel. By P. Russell. 2 vols. London, 1796-1801. Schleg. Phys. Serp. Essai sur la Physionomie des Serpens. By H. Schlegel. 2 vols. The Hague, 1837. Schneid. Hist. Amph. Historia Amphibiorum naturalis et literaria. By J. G. Schneider. Jena, 1799-1801. Theob. Cat. Descriptive Catalogue of the Reptiles of British India. By W. Theobald. Calcutta, 1876. Wagl. Syst. Amph. Natiirliches System der Amphibien. By J. Wagler Munich, 1830. REPTILIA. REPTILES may be briefly defined as cold-blooded Vertebrates breathing by lungs throughout their existence, and having the body covered with scales or scutes. A basioccipital bone is present in the skull, which articulates with the vertebral column by a single coudyle. The class Eeptilia is divided into a considerable number of Orders, which are, however, mostly extinct. Eecent Eeptiles belong to four Orders, viz. : — I. EMYDOSAUEIA, Crocodiles. II. CHELONIA, Tortoises and Turtles. III. SQTJAMATA, Lizards and Snakes. IV. EH YNCHOCEPH ALIA, a primitive type, probably ances- tral to the three others, and of which a single existing representative is known, the Tuatera of New Zealand. Order I. EMYDOSAURIA. Quadrate bone immovably united to cranial arches ; two hori- zontal bony temporal arches. Cervical and dorsal ribs mostly two-headed, the middle dorsals articulating with the transverse processes of the neural arch. Sternum and tnterclavicle present ; seven or eight transverse series of " abdominal ribs," not connected with the vertebral ribs, each series composed of four ossicles, and forming an angle directed forwards. Teeth present in the jaws, implanted in alveoli. Ventricle of the heart divided by a complete septum. Pectoral and abdominal cavities separated from each other by a muscular diaphragm. Anal opening longitudinal. Copulatory organ present, single. CROCODILTDjE. Family I. CROCODILID^. Nares anterior ; choanse opening far back between the pterygoids, which, like the palatines and maxillaries, are produced into osseous plates joining on the median line of the. palate; orbit communi- cating with the infratemporal fossa. Vertebrae proccelous ; ribs with unciuate processes. No clavicles. Pubis excluded from acetabulum. Manus with five well-developed digits, pes with four. A dorsal armour of bony scutes. Ears with movable opercles. Fig. A. Side view. atig. Angular. dr. Articular. bo. Basioccipital. d. Dentary. co. Exoccipital. /. Frontal. j. Jugal. I Lachrymal. 1.— Skull of Crocodilus pahistris. B. Upper view. m. Maxillary. n. Nasal. p. Parietal. pi. Palatine. pm. Prsemaxillary. prf. Procfrontal. pt. Pterygoid. C. Lower view. ptf. Postfrontal. q. Quadrate. qj. Quadratojugal. s.ang. Supra-angular. so. Supraoccipital. sq. Squamosal. tp. Transpalatine. GAVIALIS. 3 Tongue entirely attached to the floor of the mouth. Pupil vertical. Only the three inner digits clawed. The Crocodiles are at the same time the most highly organized and the largest of existing Eeptiles. They are thoroughly aquatic and carnivorous. Their eggs, which have a hard shell, are deposited on the banks of the water, in a hollow slightly covered up with sand. 23 or 24 living species are known from Africa, Southern Asia to the Tang-tse-Kiang, North Australia, and the tropical and warm parts of America. In the fauna with which the present work deals, Crocodiles are represented by two genera, thus distin- guished : — Snout extremely elongate ; 27 teeth or more on each side of upper jaw GAVIALIS, p. 3. Snout moderate; 17 to 19 teeth on each side of upper jaw CUOCODILTJS, p. 4. Genus GAVIALIS, Oppel, Ordn. Kept. p. 19, 1811. 27 to 29 upper and 25 or 26 lower teeth on each side, anterior largest, laterals subequal, not received into interdental pits ; the first, second, and third mandibular teeth fitting into notches in the upper jaw. Snout extremely narrow and elongate, dilated at the end ; nasal bones comparatively short, widely separated from the prsmaxillaries ; nasal opening smaller than the supratemporal fossae ; lower anterior margin of orbit (jugal) raised. Mandibular symphysis extremely long, extending to the 23rd or 24th tooth, comprising the splenial bones. A dorsal shield formed of four longitudinal series of juxtaposed, keeled, bony scutes. A single species, only found in rivers. 1. Gavialis gangeticus. Lacerta gangetica, Gmel. Syst. Nat. i, p. 1057. Gavialis gangeticus, Gilnth. Kept. B. 1. p. 63; Bouleng. Cat. Chd. $c. p. 275. Gharialis gangeticus, Theob. Cat. p. 37. Snout 3£ (adult) to 5| times (young) as long as broad at the base. Nuchal and dorsal scutes forming a single continuous shield, composed of 21 or 22 transverse series ; an outer row of soft, smooth or feebly-keeled scutes in addition to the bony dorsal scutes ; two small postoccipital scutes. Median fingers one-third, outer toes two-thirds webbed. A strong crest on the outer edge of the forearm, leg, and foot. Adult dark olive above ; young pale olive, with dark brown spots or cross-bands. Hob. Indus, Ganges, and Brahmaputra rivers and their larger tributaries ; also Mahanadi of Orissa, and Koladyne river, Arrakau, but not the Nerbudda, Tapti, Godiivari, Kistna, Irrawaddy, or other rivers of India or Burma. The Gharial reaches a length of 20 feet ; it feeds entirely upon fish, for seizing which its narrow jaws are specially adapted. B2 CEOCODILID.E. Genus CROCODILUS, Laur. Syn. Kept. p. 63 (part.), 1768. 17 to 19 upper and 15 lower teeth on each side ; fifth maxillary tooth largest ; the fourth mandibular usually fitting into a notch in the upper jaw. Snout more or less elongate ; nasal bones ex- tending to the nasal aperture, which is undivided and larger than the suprateinporal fossae. Splenial bones not entering the rnan- dibular symphysis, which does not extend beyond the eighth tooth. A dorsal shield formed of four or more longitudinal series of juxta- posed, keeled, bony scutes. Distribution. Africa, Southern Asia, North Australia, Tropical America. 11 or 12 species appear to be well established. Synopsis of Indian Species. Snout* 1| to 2 £ times as long as broad C. porosus, p. 4. Snout 1| to 1£ times as long as broad C. palustris, p. 5. 2. Crocodilus porosus. Crocodilus porosus, Schneid. Hist. Amph. ii, p. 159 ; Giinth. Rept. B. I. p. 62 ; Theob. Cat. p. 36 ; Boukny. Cat. Chel. Sfc. p. 284. Oopholis pondicherianus, Gray, A. M. N. H. (3) x, 1862, p. 268. Crocodilus pondicerianus, Giinth. Rept. B. I. p. 62, pi. vii ; Theob. Cat. p. 36. 17 to 19 upper teeth on each side. Snout ] | to 2| times as long as broad at the base ; head rough, snout with a more or less strong ridge on each side in front of the eye, slightly converging towards its fellow ; mandibular symphysis extending to the fifth tooth ; praemaxillo-inaxillary suture, on the palate, directed backwards or \V-shaped ; nasal bones separating the prsemaxiUaries above. Four large nuchal scutes forming a square, with one or two smaller ones on each side ; postoccipital scutes usually absent, sometimes small and irregular. Dorsal shield well separated from the nuchal, the scutes forming 16 or 17 transverse and 4 to 8 longitudinal series ; scales on sides and limbs smooth or feebly keeled. Fingers webbed at the base ; outer toes extensively webbed. A serrated fringe on the outer edge of the leg. Adult dark olive above ; young pale olive, with large black spots on the body and tail and dots on the head. Hob. Bengal, East coast of India, Ceylon, Burma, and Southern China to North Australia and the Solomon and Fiji Islands. Entering salt water and frequently occurring out at sea. Not recorded from West coast of India. It is not certain that this species is found far above the tideway in rivers. A specimen 33 feet long is on record. * The snout is the portion of the head in front of the orbits. CROCOUILUS. 5 3. Crocodilus palustris. Crocodilus palustris, Lesson in B6lang. Voy. 2nd. Or., Zool. p. 305 ; Kelaart, Prodr. p. 183 ; Giinth. Sept. B. I. p. 61, pi. viii, fig. A ; Theob. Cat. p. 36 ; Bouleny. Cat. Chel. Sfc. p. 285. Crocodilus trigonops, Gray, Cat. Tort. 8fC. p. 62. 19 upper teeth on each side. Snout 1^ to 1^ as long as broad at the base ; head rough but without any ridges ; mandibular synaphysis extending to the level of the fourth or fifth tooth ;, pree- maxillo-uiaxillary suture, on the palate, transverse, nearly straight, or curved forwards ; nasal bones separating the praemaxillaries above. Four large nuchals forming a square, with a smaller one on each side ; two pairs of smaller nuchals on a transverse series behind the occiput. Dorsal shield well separated from the nuchal, the scutes usually in 4, rarely in 6, longitudinal series, those of the two median usually considerably broader than long ; 16 or 17 transverse series. Scales on limbs keeled. Fingers webbed at the base ; outer toes extensively webbed. A serrated fringe on the outer edge of the leg. Adult blackish olive above: young pale olive, dotted and spotted with black. The largest specimen in the British Museum measures 12 feet, but individuals are said to grow much larger. Hub. India, Ceylon, Burma, Malay Peninsula and Archipelago. This is the common Crocodile of India, found in rivers, marshes, and ponds, and extending west to Sind and Baluchistan. This species is easily distinguished from C. porosus by its shorter snout and by the presence of five teeth in each pra?- maxillary bone, whilst the adult of the latter has but four. Order II. CHELONIA. Quadrate bone immovably united to cranial arches. Dorsal ribs single-headed, the anterior attached between two vertebrae. Body encased in a bony shell. A plastron formed of a small number of elements in addition to the clavicles. No sternum. No teeth ; jaws covered with horny sheaths. Anal opening round or longitudinal. Copulatory organ present, single. Oviparous ; eggs with a hard shell, except in the CTielonidce. The Tortoises and Turtles of India fall into five Families*. Suborder I. THECOPHORA. Dorsal vertebrae and ribs immovably united and expanded into bony plates forming a carapace. Parietals prolonged downwards towards the pterygoids. Superfam. A. TEIONYCHOIDEA. Articulation between the last cervical and the first dorsal vertebra by the zygapophyses only. Marginal bones absent or forming an incomplete series. Carapace and plastron without epidermal shields, covered with a soft skin Fam. 1. Trionychidae. "Superfam. B. CETPTODIEA. Centrum of the last cervical vertebra articulating with the centrum of the first dorsal. A complete series of marginal bones, connected with the ribs. Digits short ; tail short, with precocious ver- tebrae Fam. 2. Testudinidae. Digits short ; tail long ; caudal vertebras mostly opisthocoalous Fam. 3. Platysternidae. Limbs paddle-shaped, with only one or two claws. Marine . Fam. 4. Chelonidae. Suborder II. ATHECM. Vertebras and ribs free, separated from a bony exoskeleton. Skull without descending processes of the parietal bones. Limbs paddle-shaped, clawless. Marine Fam. 5. Sphargidae. * These five families, so far as Indian genera are concerned, may be recog- nized from external characters by use of the following key:— Limbs with four or five claws. Tail short Testudinidae. Tail nearly as long as the shell Platysternidae. Limbs with three claws Trionychidae. Limbs with one or two claws Chelonidae. Limbs without claws , Sphargidae. CHELOXIA. For the illustration of the osteological characters employed in the systematic arrangement, the following figures are appended of a skull, and of a shell stripped of the horny epidermal shields. A. Fig. 2.— Skull of Trionyx yangeticus. A. Side view. B. Upper view. C. Lower view. ar. Articular. j. Jugal. pt. Pterygoid. bo. Basiocoipital. m. Maxillary. ptf. Postfrontal. bp. Basiphenoid. opo. Opisthotic. q. Quadrate. c.a. Columella auris. p. Parietal. qj. Quadratojugal. cor. Coronoid. pi. Palatine. s.ang. Supra-angular. d. Dentary. pm. Prfemaxillary. so. Supraoccipital. eo. Exoccipital. prf. Praefrontal. sq. Squamosal. /. Frontal. pro. Prootic. v. Vouier. 8 CHELONIA. The arrangement of the bones or plates of the shell, as shown in fig. 3, differs entirely from that of the overlying shields, both in the upper shell (carapace) and lower (plastron) ; but the impres- sions of the shields are nearly always distinct, as represented in figures 8, 9, 10, 11, 12, 14. In these figures crosses are inserted to indicate the position of the anterior (axillary) and posterior (inguinal) processes or buttresses of the plastron, which, if present, anchylose with the lower surface of the carapace. Figure 4 refers to the terminology of the epidermal shields. Fig. 3. — Shell of Morenia ocellata, showing bony plates. A. Carapace. B. Plastron. co. Costal. typp. Hypoplastron. nu. Nuchal. enp. Entoplastron. m. Marginal. py. Pygal. epp. Epiplastron. ne. Neural. xyp. Xiphip hyop. Hyoplastron. xyp. Xiphiplastron. The investigation of the bony plates presents no great difficulty, and need not cause the slightest damage to the specimen, whether dry or preserved in spirit ; for the removal of one or two epidermal shields with a penknife requires but a little care, and after the examination the horny shields may be stuck on again with gum which is not soluble in alcohol. There is much greater variety, apart from individual variations, in the disposition of the bony plates than in that of the overlying shields, and much greater profit is to be derived from the study of the former in determining the relationship of genera of Tortoises. TBIONYCHIDJE. The connexion between the carapace and plastron is called the bridge. Fig. 4.— Shell of Chelone mydas, with epidermal homy shields. A. Carapace B. Plastrou. a. Abdominal. h. Humeral. n. Nuchal. an. Anal. iff. Intergular. p. Pectoral. o. Costal. im. Inframarginal. sc. Supracaudal. /. Femoral. m. Marginal. v. Vertebral. g. Gular. The anterior inframarginal is also termed axillary, and the posterior inguinal. Suborder I. THECOPHOEA. Superfam. A. TRIONYCHOIDEA. Neck bending by a sigmoid curve in a vertical plane ; cervical vertebrae without welt-developed transverse processes ; articulation between the last cervical and the first dorsal vertebra by the zyga- pophyses only. Mandible with articulary concavity ; outer border of tympanic cavity notched ; pterygoids not narrowed posteriorly, separated from each other, the basisphenoid joining the palatines. Pelvis not anchylosed to the carapace and plastron. Fourth digit with four or more phalanges. Epiplastra separated from the hyoplastra by the A-shaped entoplastron. Marginal bones absent or forming an incomplete series, not connected with the ribs. Family I. TRIONYCHIOE. Carapace and plastron without epidermal shields, covered with a soft skin. Jaws concealed under fleshy lips ; snout ending in a proboscis. Head and neck completely retractile. Ear hidden. Only the three inner digits clawed. Tail very short. 10 Distribution. Kivers of Asia, Africa, and North America. Of the six known genera, four are represented in the Indian Region. The Trionychoids are thoroughly aquatic Turtles, living chiefly in rivers ; Pelochelys even enters the sea. Specimens of Emyda, however, are occasionally found at some distance from water, according to the observations of Kelaart and Blanford. All are carnivorous, and their ferocious habits, together with the extreme rapidity with which they project their necks, render the larger indi- viduals dangerous to men bathing. Of no commercial value. Synopsis of Indian Genera. A. Plastron without cutaneous valves ; hyoplastroii distinct from hypoplastron ; no marginal ossi- fications. a. Orbit nearer the temporal than the nasal fossa ; bony choanae between the orbits TBIONYX, p. 10. b. Orbit nearer the nasal than the temporal fossa ; bony choanae between the orbits PELOCHELYS, p. 15. c. Orbit nearer the nasal than the temporal fossa ; bony choanae behind the orbits CHITRA, p. 16. B. Plastron with a cutaneous femoral valve, under which the hind limb may be con- cealed ; hyoplastron coossified with hypo- plastron ; marginal bones present EMYDA, p. 10. Genus TRIONYX, Geoftr. Ann. Mus. xiv, 1809, p. 1 (part.). Outer extremities of the nuchal plate overlying the second dorsal rib ; neural plates well developed. Limbs completely exposed. Hyoplastron distinct from hypoplastron ; not more than live plastral callosities. Bony choanse between the orbits ; jaws strong; postorbital arch narrower than the diameter of the orbit ; posterior border of pterygoids free, without ascending process. Distribution. Africa, Asia, North America. 15 species are recognized, 7 of which are found in India and its dependencies. The dimensions given are those of specimens in the British Museum : individuals may be found considerably larger, some attaining probably 5 feet in length of carapace. Synopsis of Indian Species. A. All the costal plates separated on the median line by the neural plates ; head very large . T. subplanus, p. 11. B. Last pair of costal plates in contact on the median line. a. Two neural plates between the first pair of costals; alveolar surface of lower jaw without a strong longitudinal ridge at the symphysis. TRIONYX. 11 a'. Inner edge of alveolar surface of mandible strongly raised, forming a sharp ridge, which sends off a short perpendicular process at the symphysis ; interorbital space, in the adult skull, narrower than the external nasal opening ; head with oblique black lines, converging for- wards ; no ocellar spots T. yangcticus, p. 12. I'. Inner edge of alveolar surface of mandible not or but very slightly raised, sym- physis flat or with a longitudinal furrow ; interorbital space narrower than the external nasal opening; head with oblique black lines, converging for- wards ; young with small ocellar dorsal spots T. leithii, p. 12. c'. Inner edge of alveolar surface of mandible not or but very slightly raised, sym- physis flat or with a longitudinal fur- row ; interorbital space, in the adult skull, broader than the external nasal opening; head black-marbled, with a large yellow spot behind each eye; young with large ocellar dorsal spots . . T. hurum, p. 13. b. A siiigle neural plate between the first pair of costals; alveolar surface of lower jaw with a strong longitudinal symphysial ridge. a'. Epiplastra widely separated from each other; plastral callosities well deve- loped and largely pitted T.formosus, p. 14. b'. Epiplastra in contact in front of the ento- plastron ; plastral callosities very feebly developed, finely sculptured T. phayrii, p. 14. c'. Epiplastra in contact in front of the ento- plastron ; plastral callosities well deve- loped and largely pitted T. cartilagineits^, 15. 4. Trionyx suhplanus. Trionyx subplanus, Geoffr. Ann. Mus. xiv, 1809, p. 11, pi. v, fig. 2 ; Qiinth. Kept. B. I. p. 49 ; Bouleng. Cat. Chel. $c. p. 246. Dogania giintheri, Gray, P. Z. S. 1802, p. 265. Trionyx giintheri, Gilnth. Rept. B. 1. p. 49, pi. vi, fig. A. Carapace very flat ; costal plates eight pairs, the last well deve- loped and separated by the last neural plate ; a single neural between the first pair of costals ; these plates very finely granulate and vermiculate ; a large fontanelle, till late in life, between the nuchal and dorsal plates. Epiplastra narrowly separated from each other in front of the entoplastron, which forms an obtuse or a right angle ; plastral rugosities scarcely developed, on the hyo- and hypoplastra and on the xiphiplastra. Dorsal skin of young with interrupted longitudinal ridges. Head very large ; snout (on the skull) about as long as the diameter of the orbit, which is more than double the interorbital width; postorbital arch extremely 12 TRIONYCHID.E. narrow, forming a keel on its outer surface; mandibular symphysis shorter than the diameter of the orbit, without median ridge. Brown above, yellowish beneath and on the sides of the neck ; head and neck with yellowish dots ; some young with an oblique dark streak behind each eye. Length of dorsal disk 10 inches. Hob. Mergui, Malay Peninsula, Sumatra, Borneo, Java. 5. Trionyx gangeticus. Trionyx gangeticus, Cuvier, Oss. Foss. v, pt. 2, pp. 186. 222. pi. xi, figs. 5-8; Bouleng. Cat. CM. $c. p. 248. Trionyx javanicus (non Geoff")-.), Gray, IN. Ind. Zool. ii, pi. Ixv ; Ounth. Kept. B. I. p. 48. Aspilus gataghol, Gray, A. M. N. H. (4) x, 1872, p. 339. Trionyx gangeticus, part., Theob. Cat. p. 28. Costal plates eight pairs, the last well developed and in contact throughout on the median line; two neurals between the first pair of costals ; plates coarsely pitted and vermiculate. Epiplastra narrowly separated from each other in front of the entoplastron, which forms an obtuse or a right angle ; plastral callosities very large, hyo-hypoplastral, xi phi pi astral, and, in old specimens, ento- plastral. Dorsal skin of young with longitudinal ridges of small tubercles. Head moderate ; snout (on the skull) about as long as the diameter of the orbit ; interorbital region, in the adult, con- siderably narrower than the nasal fossa ; postorbital arch one third to one half the greatest diameter of the orbit ; mandible with the inner edge strongly raised, forming a sharp ridge, which sends off a short perpendicular process at the symphysis ; the diameter of the mandible at the symphysis does not exceed the diameter of the orbit. Olive above ; back of young vermiculated with fine black lines, but without ocelli ; head with a black longitudinal streak from between the eyes to the nape, intersected by two or three A- shaped black streaks ; lower parts yellowish. Length of dorsal disk 2 feet. Hab. Gangetic river-system ; Indus ? 6. Trionyx leithii. Trionyx leithii, Gray, A. M. N. H. (4) x, 1872, p. 334 ; Bouleng. Cat. Chel. 8fc. p. 249. ? Trionyx gangeticus, Murray, Zool. Sind, p. 342. Intermediate between T. gangetiats and T. Tiurvm. Agrees with the former in the width of the interorbital space, the comparatively short mandibular symphysis, and the markings of the head ; with the latter in the longer and more pointed snout, the absence of a strong ridge on the inner alveolar surface of the mandible, and in the presence, in the young, of four or more dorsal ocelli, which are, however, smaller than in T. hurum. Hab. Kistna River, Deccan, and Nelambar Eiver, Malabar. This will probably prove to be a Peninsular form, ranging from the Indus to Madras and Malabar. TK1ONYX. 13 7. Trionyx hurum. Trionyx hurum, Gray, Syn. Rept. p. 47, pi. x ; id. III. Ind. Zool. ii, pi. Ixvi ; Bouleng. Cat. Chel. Sfc. p. 249. Tnonyx ocellatus, Gray. III. Ind. Zool. i. pi. Ixxviii : Theob. P. A. S. B. 1875, p. 174, pi. iv ; id. Cat. p. 29. Trionyx gangeticus, Giinth. Rept. B. I. p. 47. Trionyx buchanani, Theob. P. A. S. B. 1874, p. 78. Trionyx gangeticus, part., Theob. Cat. p. 28. Fig. 5. — Trionyx hurum (young). Costal plates eight pairs, the last well developed and in contact throughout on the median line; two iieurals between the first pair of costals ; plates coarsely pitted and vermiculate. Epiplastra narrowly separated from each other in front of the eutoplastron, which forms an obtuse or a right angle ; plastral callosities very large, hyo-hypoplastral, xiphiplastral, and in old specimens ento- plastral. Dorsal skin of young with longitudinal ridges of small tubercles. Head moderate ; snout (on the skull) a little longer than the diameter of the orbit ; iuterorbital region, in the adult, as broad as the nasal fossa; postorbital arch somewhat more than half the diameter of the orbit in the adult ; inaudible without strongly raised inner edge or longitudinal symphysial ridge ; the diameter of the mandible at the symphysis exceeds the diameter of the orbit. Olive-brown above and beuealh ; head, neck, limbs, and 14 TBIONYCHIDjE. border of shell light-dotted ; head with large yellow spots, viz. across the snout, behind the orbit, at the angle of the mouth, on the tympanic region, and on the chin ; these spots become indistinct in the adult, the head of which is more or less closely spotted or vermiculated with black ; dorsal disk in the young with dark marblings and four or six very large ocelli. Length of dorsal disk 16 inches. Hob. Ganges and its tributaries. 8. Trionyx formosus. Trionyx formosus, Gray, P. Z. S. 1869, p. 217, pi. xv, fig. 1 ; Theob. Cat. p. 31 ; Bouleng. Cat. Chel. Sfc. p. 250. Trionyx peguensis, Gray, Suppl.Cat. Sh.Rept. \, p. 99 ; Theob. Cat. p.31. Trionyx grayii, Theob. P. A. S. B. 1875, p. 176, pi. iii ; id. Cat. p. 31. Costal plates eight pairs, the last well developed and in contact throughout on the median line ; a single neural between the first pair of costals ; plates coarsely pitted and vermiculate. Dorsal skin of young with longitudinal ridges of small tubercles. Epi- plastra separated from each other ; entoplastron forming an obtuse or right angle ; plastral callosities well developed, hyo-hypoplastral and xiphiplastral, and coarsely sculptured like the carapace. Head moderate : snout (on the skull) about as long as the diameter of the orbit ; interorbital region, in the adult, as broad as the nasal fossa ; postorbital arch, in the adult, a little less than half the diameter of the orbit ; mandible with a strong ridge along the symphysis, the diameter of which equals or a little exceeds that of the orbit. Olive-brown above, white below ; head of young ornamented with yellow black-edged markings, the largest being a cross-band on each side of the back of the head ; these markings disappear in the adult, the head of which is closely spotted with black above and uniform white inferiorly ; young with four large dorsal ocelli, as in T. hurum. Size of T. hurum. Hab. Irrawaddy, Sittoung, and Salween rivers, Burma. 9. Trionyx phayrii. Trionyx phayrei, Theob. Journ. Linn. Soc. x. 1868, p. 18 ; id. Cat. p. 30; Bouleng. Cat. CM. #e. p. 251. Trionyx jeudi, Gray, P. Z. 8. 18«9, p. 217, figs. Trionyx cariniferus (non Gray), Theob. P. A. S. B. 1874, p. 80, pi. iv. Costal plates eight pairs, the last well developed, with slightly concave posterior border, and in contact on the median line ; a single neural between the first pair of costals ; plates rather coarsely pitted and vermiculate. Epiplastra in contact in front of the entoplastron, which forms an obtuse or right angle ; plastral callosities hardly developed, the hyoplastra bearing a mere trace of sculpture. Head moderate; skull as in T. formosus, but snout a little longer in the adult. Head of adult spotted or vermiculated with black above and uniform whitish inferiorly, as in T. formosus. Length of dorsal disk 20 inches. Hab. Pegu, Arrakan range, Tenasserim and Sumatra. PELOOHELYS. 15 10. Trionyx cartilagineus. Testudo cartilaginea, Boddaert, Epistola ad W. Roell, 1770. Trionyx ornatus, Gray, P. Z. S. 1861, p. 41. pi. v ; Gilnth. Kept. B. I. p. 48, pi. vi, fig. B. Trionyx stellatus, var. javanicus (Geofr.), Theob. P. A. S. B. 1874, p. 79, pi. iii ; id. Cat. p. 30. Trionyx ephippium, Theob. P. A. S. B. 1875, p. 177, pi. v. Trionyx cartilagineus, Bouleng. Cat. Chel. fyc. p. 253. Costal plates eight pairs, the last well developed and forming a median suture behind the last neural ; a single neural between the first pair of costals ; dorsal plates coarsely pitted and vermiculate. Dorsal skin of young with longitudinal ridges of small tubercles. Epiplastra in contact, or narrowly separated, in front of the entoplastron, which forms an obtuse or a right angle ; plastral callosities well developed in the adult, hyo-hypoplastral and xiphiplastral, and coarsely sculptured like the carapace. Head moderate ; snout (on the skull) considerably longer than the diameter of the orbit ; iuterorbitai region, in the adult, as broad as the nasal fossa, or a little narrower ; postorbital arch very narrow, one third or one fourth the diameter of the orbit ; mandible with a strong ridge along the symphysis, the diameter of which equals or exceeds the diameter of the orbit. Olive-brown above and on the chin ; head and chin with numerous yellow spots, the largest of which are below the temple ; dorsal disk of young light-dotted, and usually with a few large blackish spots ; lower parts white. Size of T. hurum. Hob. Pegu, Tenasserim, Siam, Camboja, Malay Peninsula, Sumatra, Borneo, Java. Genus PELOCHELYS, Gray, P. Z. S. 1864, p. 89. Outer extremities of the nuchal plate overlying the second dorsal rib ; neural plates well developed. Limbs completely exposed. Hyoplastron distinct from hypoplastron ; not- more than five plastral callosities. Bony choanso between the orbits ; jaws weak ; postorbital arch as broad as the diameter of the orbit ; posterior border of pterygoids free, without ascending process. A single species. 11. Pelochelys cantoris. Chitra indica, part., Gray, Cat. Tort. p. 49 ; Gunth. Rept. B. I. p. 50, pi. vi. fig. C. Gymnopus indicus, Cantor, J. A. S. B. xvi, 1847, p. 616. Pelochelys cantorii, Gray, P. Z. S. 1864, p. 90, figs. ; Theob. Cat. p. 28 ; Bouleng. Cat. Chel. 8fe. p. 26S. Costal plates eight pairs, the last well developed and forming a median suture ; a single neural between the first pair of costals ; plates coarsely pitted and vermiculate. Dorsal skin of young 16 TRIONYCHIDjE. tuberculate. Epiplastra small and widely separated ; entoplastron forming a right or an acute angle; plastral callosities largely developed. Head moderate; snout very short and broad; pro- boscis very short ; interorbital space broader than the greatest diameter of the orbit ; mandible narrowest at the symphysis. Olive above, uniform or spotted with darker ; lips and throat of young olive, speckled with whitish ; plastron whitish. Length of dorsal disk 2 feet. Hob. Ganges, Burma, Malay Peninsula, Borneo, Philippines. Genus CHITRA, Gray, Cat. Tort. p. 49, 1844. Outer extremities of the nuchal plate overlying the second dorsal rib ; neural plates well developed. Limbs completely exposed. Hyoplastron distinct from hypoplastrou ; not more than five plastral callosities. Bony choanse behind the orbits, which are situated quite at the anterior extremity of the skull ; jaws weak ; postorbital arch at least twice as broad as the diameter of the orbit ; posterior border of pterygoids free, without ascending process. A single species. 12. Chitra indica. Trionyx indicus, Gray, Syn. Kept. p. 47. Trionyx aegyptianus, var. indicus, Gray, III. Ind. Zool. i, pi. Ixxx. Chitra indica, part., Giinth. Rept. B. I. p. 50. Chitra indica, Gray, P. Z. S. 1864, p. 91, figs. ; Theob. Cat. p. 27 ; Bouleng. Cat. Chel Sfc. p. 264. Costal plates eight pairs, the last well developed and forming a median suture ; a single neural between the first pair of costals ; plates coarsely pitted and vermiculate. Epiplastra widely sepa- rated ; entoplastron forming an acute angle ; plastral callosities largely developed. Head small, with extremely short snout ; interorbital space considerably narrower than the greatest diameter of the orbit. Olive or green above, whitish interiorly ; dorsal disk oE young with dark vermiculations ; head and neck with dark longitudinal lines. Length of dorsal disk 2 feet. Htib. Ganges and Irrawaddy ; Indus ? Genus EMYDA, Gray, Syn. Rept. p. 49, 1831. Nuchal notched at each outer end, which underlies the first costal plate ; carapace large, with a series of bony plates in the posterior cutaneous border and a pranuchal marginal bony plate ; neural plates well developed, seven or eight in number, forming a EMTDA. 17 continuous series ; eighth pair of costals large in the adult, and, like the penultimate, forming a median suture. Plastron with a cutaneous femoral valve, under which the hind limb may be con- cealed ; hyoplastron coossified with hypoplastron ; seven plastral callosities (in the adult). Bony choanae between the orbits ; jaws strong ; postorbital arch moderate, much narrower than the dia- meter of the orbit ; posterior border of pterygoids with a median ascending process forming a suture with the opisthotic. Three species are distinguished, all being Indian or Burmese. Synopsis of the Species. Head with yellow spots ; entoplastral callosity small. E. granosa, p. 17. Head without yellow spots ; first marginal plate much larger than second E. vittata, p. 17. Head without yellow spots ; first marginal plate not larger than second E. scutata, p. 18. 13. Emyda granosa. Testudo granosa, Schoepff, Test. p. 127, pi. xxx, A & B. Trionyx granosa and T. punctata, Gray, III. Ind. Zool. ii, pis. Ixiii, Ixiv. Emyda granosa, Giinth. Sept. B. I. p. 45 ; Theob. Cat. p. 32 ; Bouleng. Cat. Chel. $c. p. 269. Eniyda dura, Anders. Journ. Linn. Soc. xii, 1876, p. 514. Carapace and plastral callosities finely and uniformly granulate ; two neural plates between the first pair of costals ; anterior marginal very large, much larger than the others; seven plastral callosities, of which the entoplastral is much the smallest ; the extent of these callosities varies considerably, not only with age, but according to individuals. Head moderate ; snout short. Skin of dorsal disk, in the young, longitudinally plaited. Olive-brown above, with round yellow spots on the head and back, which become less distinct in adult specimens; plastron and margin of carapace yellowish white. Length of dorsal disk 10 inches. Hub. Indus and Ganges plains, in rivers, canals, &c. 14. Emyda vittata. Emyda punctata (non Gray), Kelaart, Prodr. p. 179. Emyda vittata, Peters, MB. Ak. Berl 1854, p. 216 ; Giinth. Rept. B. I. p. 46 ; Theob. Cat. p. 32 ; Bouleng. Cat. Chel. §c. p. 269. Emyda ceylonensis, Gray, Cat. Sh. Rept. i, p. 64, pi. xxix A ; Giinth. Rept. B. I. p. 45. Under this name may be provisionally grouped such specimens as agree with E. granosa except in the uniform brown colour of the upper parts (with or without darker bands on the head and neck), and usually in the larger entoplastral and xiphiplastral callosities. The constancy of these characters and the correlation 18 between thorn, however, still require to be tested upon a larger number of specimens. Hab. Ceylon ; Southern India ; Grodavari, Mahiinadi, and their tributaries. 15. Emyda scutata. Emyda scutata, Peters, MB. Ak. Berl. 1808, p. 449 ; Theob. Cat. p. 32 ; Boukng. Cat. Chel $c. p. 270. Emyda fusconiaculata, Gray, A.M.N. H. (4) xi, 1873, p. 308. Very closely allied to E. yranosa and E. vittata, but anterior marginals less enlarged, less unequal in size, the first not larger than the second. Entoplastral callosity large or moderate. Brown above, carapace spotted or reticulated with darker. Hab. Irrawaddy. Superfam. B. CRYPTODIRA. Neck bending by a sigmoid curve in a vertical plane ; cervical vertebrae without transverse processes or with mere indications of them ; centrum of the last cervical articulating with the centrum of the first dorsal. Mandible with articulary concavities ; outer border of tympanic cavity incompletely bony ; pterygoids narrow in the middle, in contact on the median line. Pelvis not anchy- losed to the carapace and plastron. Digits with not more than three phalanges. Epiplastra in contact with hyoplastra ; ento- plastron, if present, oval, rhomboidal, or T-shaped. A complete series of marginal bones, connected with the ribs. Family II. TESTUDINID^E. Shell covered with epidermal shields. Caudal vertebra) pro- coelous. Neck completely retractile within the shell. Parietal bones widely separated from squamosals. Digits short or mo- derately elongate ; phalanges with condyles ; claws four or five. This family, which contains the bulk of recent Chelouians, is distributed over every part of the world except Australia. Ten genera are represented in the Indian Eegion. Synopsis of Indian Genera. A. Limbs club-shaped ; alveolar surface of upper jaw with a median ridge TESTUDO, p. 19. B. Digits distinct, with at least a rudiment of web ; alveolar surface of upper jaw without median ridge. a. Head covered with undivided skin. a'. Plastron not hinged ; no bony temporal arch. GEOEMYDA, p. 23. V. Plastron not hinged ; a bony temporal arch. NICOEIA, p. 26. c'. Plastron hinged, movable ; a bony temporal arch CYCLEHYS, p. 28. TESTUDO. 19 b. Hinder part of head covered with small shields. a'. Choanae between the eyes BELLIA, p. 32. V. Choanae behind the level of the eyes .... DAMONIA, p. 34. C. Digits distinct, extensively or entirely webbed ; alveolar surface of upper jaw with one or two median ridges. a. Fourth vertebral shield not longer than third, embracing three neural bones. a'. Axillary and inguinal buttresses of plastron short; choanse behind the level of the eyes MOBENIA, p. 35. b'. Axillary and inguinal buttresses much de- veloped, the former connected with the first rib or nearly reaching it. a". Choanee behind the level of the eyes ; al- veolar surface of upper jaw with a single median ridge HARDELLA, p. 36. b". Choana? behind the level of the eyes ; alveolar surface of upper jaw with two median ridges; fore limb with four claws only BATAGUR, p. 37. b. Fourth vertebral shield elongate, embracing four or five neural bones KACHUGA, p. 38. Genus TESTUDO, Linn. Syst. Nat. i, p. 350 (part.), 1766. Neural plates usually alternately tetragonal and octagonal ; costal plates alternately narrower and wider. Suture between the mar- ginal and costal bones and shields usually corresponding ; supra- caudal shield usually single. Plastron extensively united to the carapace by suture, with short axillary and inguinal buttresses, which do not reach, or just reach, the costal plates ; entoplastrou usually anterior to the humero-pectoral suture. Skull with weak or moderately wide postorbital and temporal arches ; alveolar surface of upper jaw with one or two median ridges ; choanae between the eyes. Head shielded above. Limbs club-shaped, covered with large scales or tubercles. Tail short. This genus, of which some 40 species are known, is represented in every part of the world except Australia. Habits terrestrial and herbivorous. Synopsis of Indian Species. A . Supracaudal shield single. a. Nuchal shield present ; forehead with large shields. a'. Anal shields forming a very short suture, or entirely separated by the anal notch. . T. elongata, p. 20. b'. Suture between the anals at least as long as that between the abdominals T. leithii, p. 20. C2 20 TESTUDINID^. b. No nuchal; forehead with small irregular shields. «'. Heel and hinder side of thighs with spur- like tubercles ; plastron with dark radia- ting lines T. eleyans, p. 21 . b'. Heel and hinder side of thighs without spur-like tubercles ; plastron without ra- diating lines T. platynota, p. 22. B. Supracaudal shields two T. emys, p. 22. 16. Testudo elongata. Testudo elongata, Blyth, J. A. S. B. xxii, 1853, p. 639 ; Guntli. Rept. B. /. p. 8 ; Theob. Cat. p. 3 ; Boukny. Cat. Chel. $c. p. 173. Shell considerably depressed, more than twice as long as deep, with flat vertebral region ; anterior and posterior margins slightly reverted, strongly serrated in young, feebly in old specimens ; shields concentrically striated, except in old specimens ; nuchal present (rarely absent), narrow and elongate ; supracaudal undivided, more or less incurved ; first vertebral usually nearly as long as broad in the adult, the others broader than long and nearly as broad as the costals. Plastron large, truncate anteriorly, deeply notched poste- riorly; suture between the pectoral shields as long as or longer than that between the humerals : suture between the gulars as long as or a little shorter than that between the pectorals ; anals forming a very short suture, or entirely separated by the anal notch ; axillary and inguinal moderate. Head moderate; a pair of large pnefrontal shields, usually followed by a nearly equally large frontal ; beak feebly hooked, tricuspid ; alveolar ridge of upper jaw short and rather feeble. Anterior face of fore limbs with moderate, unequal- sized, imbricate scales, which are largest towards the outer side ; no enlarged tubercles on the thighs ; tail ending in a claw-like horny tubercle. Carapace and plastron greenish yellow, each shield with an irregular black spot or blotch, which may occupy its greater portion, or may be much broken up or indistinct. Length of shell 10| inches. Hab. Bengal (Chaibassa), Assam, the whole of Burma, Camboja, Cochin China. 17. Testudo leithii. Testudo leithii, Giittth. P. Z. S. 1869, p. 602, figs. ; Theob. Cat. p. 3 ; Bouleng. Cat. Chel. tips. p. 175. Shell very convex, not twice as long as deep, deeply notched anteriorly ; nuchal shield triangular ; posterior border of supra- caudal forming an angle, projecting beyond the marginals ; second vertebral shield longest ; shields concentrically striated. Plastron large, hind lobe notched behind and movable in the adult ; suture between the pectoral shields a little shorter than that between the humerals ; suture between the anals longer than that between the TESTTJDO. 21 feniorals ; axillary and inguinal moderate. Head moderate ; a pair of large praefrontal shields, followed by a very large frontal ; beak scarcely hooked ; alveolar ridge of upper jaw short and feeble. Anterior face of fore limb covered with a few extremely large im- bricate bony scutes disposed in three longitudinal and five trans- verse series. Carapace yellow, each shield bordered anteriorly and laterally with blackish brown; plastron yellow, each abdominal shield with a dark brown triangular or cuneiform marking. Length of shell 5 inches. Hcib. Lower Egypt and the neighbouring part of Syria. The single specimen upon which the species was founded was said to be from Sind, where, however, this tortoise does not appear to have been rediscovered during the last twenty years. 18. Testudo elegans. Testudo elegans, Schoepff, Test. p. Ill, pi. xxv; Giinth. Rept. B. I. p. 4 ; Theob. Cat. p. 2 ; Bouhnq. Cat. Chel. tyc. p. 161. Testudo megalopus, Myth, J. A. 8. B. xxii, 1853, p. 640. Fig. 6. — Testudo elegans. Carapace very convex, dorsal shields often forming humps ; lateral margins nearly vertical ; posterior margin somewhat expanded and strongly serrated; no nuchal ; supracaudal undivided, incurved in the male ; shields strongly striated concentrically ; first vertebral longer than broad, the others broader than long, third at least as broad as the corresponding costal. Plastron large, truncate'd or openly notched in front, deeply notched, bifid behind ; suture between the humerals much longer than that between the femorals ; suture be- tween the pectorals very short : axillary and inguinal rather small. Head moderate ; forehead swollen, convex, and covered with rather small and irregular shields ; beak feebly hooked, bi- or 22 TESTUDINID.E. tricuspid; edge of jaws denticulated; alveolar ridge of upper jaw strong. Outer-anterior face of fore limb with numerous un- equal-sized, large, imbricate, bony, pointed tubercles ; heel with large, more or less spur-like tubercles ; a group of large conical or sub- conical tubercles on the hinder side of the thigh. Carapace black, with yellow areolas from which yellow streaks radiate ; these streaks usually narrow and very numerous : plastron likewise with black and yellow radiating streaks. Length of shell 10 inches. Hob. India (except Lower Bengal), extending west to Sind ; and Ceylon. 19. Testndo platynota. Testudo platynota, Blyth, J. A. S. B. xxxii, 1863, p. 83 ; Theob. Cat. p. 2 ; Bouletig. Cat. Chel 8fc. p. 162. Very closely allied to the preceding, but carapace more regularly oval ; posterior border not or but very slightly expanded, and more feebly serrated ; vertebral shields flat. Tellow rays on the dorsal shields wider and fewer, usually three to the right and three to the left of each shield ; plastron yellow, with some large black blotches but without radiating streaks. According to Anderson, the scales on the fore limb are fewer and all rounded, the sharp scales on the heel are absent, and the conical scales on the posterior portion of the thigh are very much smaller and not at all prominent. Hob. Upper Burma and Northern Pegu. 20. Testndo emys. Testudo emys, Schleg. 8f Milll. in Temminck, Ver. Nat. Nederl Ind., Rept. p. 34, pi. iv ; Boulmg. Cat. Chel. $c. p. 158. Geoemyda spinosa, part., Cantor, J. A. S. B. xvi, 1847, p. 607. Manouria fusca, Gray, P. Z. S. 1852, p. 134. Testudo phayrei, Blyth, J. A. S. B. xxii, 1853, p. 639. Manouria emys, Gilnth. Rept. B. 1. p. 10 ; Theob. Cat. p. 4. Testudo (Scapia) falconeri, Gray, P. Z. S. 1869, p. 169, figs. Shell considerably depressed, its depth not half its length ; ante- rior and posterior margins reverted, more or less strongly serrated ; nuchal present ; supracaudal shields two ; dorsal shields concentri- cally striated, often concave ; vertebrals much broader than long and at least as broad as costals. Plastron large, gular region some- what produced and usually notched, hind lobe deeply notched ; the pectoral shields may be widely separated from each other, or from a short median suture; axillary shield very small, inguinal large. Head moderate ; two large prafrontal shields and a large frontal ; beak not hooked ; jaws feebly denticulated, the alveolar surface of the upper jaw with a strong median ridge. Fore limb anteriorly with very large, bony, pointed, imbricate tubercles, forming four or five longitudinal series ; hind limb with very large bony tubercles on the plantar surface, with others larger, conical, and spur-like on GEOEMYDA. the heel, and a group of still larger conical tubercles on each side on the back of the thighs. Adult dark brown or blackish ; carapace of young yellowish brown, with dark-brown markings. Length of shell 18 inches. Hob. Assam, Burma, Siam, Malay Peninsula, Sumatra. Genus GEOEMYDA, Gray, P.Z. S. 1834, p. 100 (part.). Neural plates mostly hexagonal, hinder lateral margins shortest. Plastron extensively united to the carapace by suture, with axillary Fig. 7. — Skull of Geoemyda grandis. and inguinal peduncles just reaching the first and fifth costals ; entoplastron intersected by the huniero-pectoral suture. Skull 24 TESTUDINID^l. without bony temporal arch ; alveolar surfaces narrow, without median ridge. Upper surface of head covered with undivided skin. Digits with a short web. Tail very short, not longer in the young than in the adult. Distribution. Burma, Malay Peninsula and Archipelago. Three species are known. Habits terrestrial or semiaquatic. Fig. 8.— Shell of Geoemyda grandis. Synopsis of the Species. A. Anterior margin of shell serrated ; second verte- bral shield at least as broad as the second costal, much broader than long O. spinosa, p. 25. B. Anterior margin of shell not serrated; second vertebral shield narrower than the second costal, not much broader than long. a. Carapace arched or tectiform in a transverse section O. grandis, p. 25. b. Carapace much depressed, flat on the vertebral region G. depressa, p. 25. GEOEMYDA. 25 Emys spinosa, Gray, Syn. Kept. p. 20; id. III. Ind. Zool. i, pl.lxxvii, figs. 1, 2. Geoemyda spinosa, Gray, P. Z. S. 1834, p. 100 ; id. III. Ind. Zool. ii, pi. Ivii, fig. 2 ; Giinth. Rept. B. I. p. 18 ; Theob. Cat. p. 5 ; Bouleng. Cat. CM. Sfc. p. 137. Carapace of adult much depressed, with broad flat vertebral region and an uninterrupted obtuse keel ; anterior and posterior margins serrated ; carapace of young more regularly arched, with a short keel or spinose tubercle on each costal shield, and the border strongly serrated all round, each marginal being produced in a spine, with or without smaller denticulation on each side ; the vertebral keel stronger than in the adult. Vertebral shields (2-4) much broader than long, at least as broad as the costals ; nuchal shield longer than broad. Plastron large ; the width of the bridge about equals the length of the posterior lobe, which is angulai'ly eniar- ginate ; as regards size, the plastral shields take the following order commencing with the largest: — abdominals, pectorals, femorals, numerals, anals, gulars ; the suture between the abdominals not or but slightly longer than that between the pectorals ; axillary and inguinal shields present. Upper jaw bicuspid. Carapace brown above, reddish in the young, the vertebral keel lighter ; each plas- tral shield with yellow and dark-brown radiating streaks ; soft parts dark brown, with a yellowish spot on each side of the neck, near the ear. Length of carapace 8 inches. Hob. Tenasserim, Malay Peninsula, Sumatra, Borneo. 22. Geoemyda grandis. Geoemyda grandis, Gray, A. M. N. H. (3) vi. 1860, p. 218 ; Giinth. Rept. B. I. p. 19, pis. i & ii, figs. A & B ; Theob. Cat. p. 5 ; Bouleng. Cat. Chel. $c. p. 138. Carapace arched or tectiform, not flattened on the vertebral region, which bears a strong but obtuse keel ; only the posterior margin serrated. Vertebral shields (1-3) not or but slightly broader than long and narrower than the costals ; nuchal longer than broad. Plastron similar to that of G. spinosa • the length of the suture between the pectoral shields at least two thirds that of the abdominals. Dark brown or blackish above ; plastron and lower surface of marginals with black and yellow rays, either the black or the yellow predominating. Length of carapace 16 inches. Hob. Pegu, Tenasserim, Siarn. 23. Geoemyda depressa. Geoemyda depressa, Anders. A. M. N. H. (4) xvi, 1875, p. 284 ; id. An. Zool. lies. Yunnan, p. 721, pis. Iv, Ivi, & Ixxv B, figs. 1-5; Bouleng. Cat. Chel. $c. p. 139. Geoemyda arakana, Theob. Cat. p. vii. 26 Shell much depressed, the depression increasing from before backwards, the shell being somewhat expanded across the inguinal region ; only the posterior border serrated ; second vertebral about as long as broad, narrower than the costals ; nuchal longer than broad. Plastral shields as in G. spinosa and G. grandis. Carapace light brown ; plastron and lower surface of marginals yellow, the plastral shields with a few broad black rays ; the interval between the axillary and inguinal notches black. Head leaden, neck pale yellowish brown ; limbs of the latter colour, with the large scales dark, almost black. Length of carapace 9| inches. Hob. Hills near Akyab, Arrakan. Genus NICORIA, Gray, Cat. Sh. Kept, i, p. 17 (1855). Neural plates mostly hexagonal, with the hinder lateral margins Fig. 9.— Shell oiNicoria trijuga. shortest, or alternately tetragonal and octagonal. Plastron more or less extensively united to the carapace by suture, with short axillary NICORIA. 27 buttress, just reaching the first costal plate ; entoplastron intersected by the huruero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces without median ridge ; choanse between the eyes. Upper surface of head covered with undivided skin. Digits with a short web or nearly free. Tail short, not longer in the young than in the adult. Distribution. Oriental Eegion, Central and South America. Two species occur within the limits of the present fauna. Habits terrestrial or semiaquatic. Synopsis of Indian Species. Digits webbed ; plastron brown or black .... N. trijuga, p. 27. Digits nearly free ; plastron yellow N. tricarinata, p. 28. 24. Nicoria trijnga. Emys trijuga, Schiocigg. Prodr. p. 41 (370) ; Giinth. Kept. E. I. p. 29. Melanochelys trijuga, Gray, P. Z. S. 18(i9, p. 187 ; Thtob. Cat. p. 12. Emys trijuga, var. maderaspatana, Anders. An. Zool. Res. Yunnan, p. 729. Nicoria trijuga, Bouleng. Cat. Chelon. fyc. p. 121. Carapace moderately depressed, tricarinate. Vertebral shields in adult as long as broad or longer than broad, and much narrower than the costals; nuchal moderate or small, sometimes absent. Plastron large ; the width of the bridge equals or a little exceeds the length of the posterior lobe, which is broadly notched ; front lobe truncated. Axillary and inguinal shields present. Upper jaw not hooked, notched mesially ; the width of the mandible at the symphysis is less than the diameter of the orbit. Digits distinctly webbed. Tail shorter than the head. Shell brown, with a yellow stripe on each side of the plastron ; keels sometimes yellowish. Head brown, in the young reticulated with yellowish (in spirit), and with a yellowish line from the eye to above the tympanum. Length of sheU 9 inches. Hub. Peninsula of India and Punjab. Var. thermalis. Emys thermalis, Lesson, Cent. Zool. p. 86, pi. xxix. Geoclernmys seba, Gray, Cat. Sh. Rept. \, p. 18. Emys trijuga, Kelaart, Prodr. p. 177 ; Giinth. Rept. B. I. pi. ii, fig. C. Melanochelys sebae, Theib. Cat. p. 12. Emys sebae, Anders. An. Zool. Res. p. 729. Shell dark brown in the young, with a broad yellow border to the plastron ; keels often yellowish ; shell of adult deep black, with a narrow yellow border to the plastron. Soft parts blackish ; head of young and halfgrown with irregular bright-orange spots, which disappear in the adult. Length of shell 9 inches. Hab. Ceylon. 28 TESTUDINIDJB. Var. edeniana. Melanochelys edeniana, Theob. Cat. p. 12. Emys trijuga, var. burmana, Anders. An. Zool. Res. p. 729, pis. Ivii & Iviii. A larger race, with the shell deep black, the plastral border, and sometimes also the keels, yellowish. Light markings on the head, if present, small. Length of shell 16 inches. Hob. Burma. 25. Nicoria tricarinata. Geoemyda tricarinata, Blyth, J. A. S. B. xxiv, 1855, p. 714. Chaibassia tricarinata, Theob. Cat. p. 6. Chaibassia tricarinata and theobaldi, Anders. An. Zool. Res. Yunnan, pp. 717, 718 ; Bouleng. Cat. Chel 8?c. pp. 139, 140. Nicoria tricarinata, Lydekker, J. A. S. B. Iviii, 1889, p. 327, fig. Allied to the preceding species, but of more terrestrial habits, as indicated by the more convex carapace, abruptly descending pos- teriorly, and the very indistinct web between the digits. Scales on the limbs larger. The hypoplastron is frequently attached to the carapace by ligament; no inguinal shield. Carapace dark brown or black, with the three keels yellow; plastron uniform yellow ; head black, with a broad yellow or reddish band on each side passing over the eye and the tympanum; a narrow similar band below the angle of the mouth, along the inferior margin of the lower jaw ; neck and limbs blackish. Length of shell 5 inches. Hob. Bengal (Chaibassa) and Assam (Bisthnath Plain). I have recently had an opportunity of examining specimens of 0. tricarinata and theobaldi, and regard them as belonging to one species, which cannot be generically separated from N. trijuga. Genus CYCLEMYS, Bell, P. Z. S. 1834, p. 17. Neural plates mostly hexagonal, posterior lateral margins shortest. Plastron united to carapace by ligament, and more or less distinctly divided into two lobes moving upon a ligamentous hinge between the hyo- and the hypoplastra; both the latter bones contributing to the formation of the bridge; entoplastron intersected by the humero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces without median ridge ; choanae between the eyes. Upper surface of head covered with undivided skin. Digits webbed or nearly free. Tail short, not longer in the young than in the adult. Distribution. The greater part of the Oriental Eegion, east of the Bay of Bengal. Terrestrial or semiaquatic. CYCLEMYS. The ligameutous transverse hinge, dividing the plastron into an anterior and a posterior moiety, may be indistinct, and is nearly always so in quite young specimens. These may, however, be readily distinguished from young of the preceding genus by the hooked upper jaw. Fig. 10. — Shell of Cyckmys amboinensis. Synopsis of Burmese Species. A. Plastron emarginate posteriorly ; posterior margin of carapace serrated. a. Suture between pectoral shield and mar- ginals longer than that between abdomi- nal and marginals C. platynota, p. 30. b. Suture between pectoral and marginals shorter than that between abdominal and marginals. a'. Carapace unicarinate C. dhor, p. 30. b'. Carapace tricarinate ; C. motihoti, p. 31. B. Plastron not notched posteriorly ; carapace tricarinate in the young ; posterior margin not serrated C. amboinensis, p. 31. 30 TESTTTDINIDJE. 26. Cyclemys platynota. Emys platynota, Gray, P. Z. S. 1834, p. 54 ; id. III. Ind. Zool. ii, pi. Ivii, tig. 1. Cyclemys platynota, Gray, Cat. Sh. Kept. i. p. 43; Bculeng. Cat. CM. $c. p. 130. Notochelys platynota, Ovmth. Kept. B. I. p. 17; TJieob. Cat. p. 10. Carapace depressed, angular, the vertebral region flat, with an interrupted obtuse keel ; posterior margin serrated ; six, rarely seven, vertebral shields, the small additional shield intercalated between the fourth and fifth ; vertebrals (except the first) much broader than long, at least as broad as the costals. Plastron narrower than the opening of the shell, hind lobe openly emarginate, connected with the carapace by a distinct bridge, the width of which is contained twice and a half to twice and two thirds in the length of the plas- tron ; the hyo- and hypoplastral bones contribute to an equal degree to the formation of the bridge ; no distinct transverse hinge ; a large inguinal shield ; the suture between the pectoral shield and the marginals longer than that between the abdominal and the marginals ; the median suture between the pectoral shields as long as or a little longer than that between the abdominals. Hook of the upper jaw bicuspid, divided by a median groove. Digits exten- sively webbed ; claws long, curved, sharp. Front part of arm with broad, band-like transverse shields. Carapace reddish brown, or yellowish brown with radiating brown lines ; young with a pair of round black spots on each vertebral shield, and one or two similar spots on each costal ; plastron yellow with brown spots, or brown with the sutures between the shields yellow. Head yellowish brown ; neck brown, with irregular yellowish streaks, the broadest of which extends from the posterior corner of the eye. Length of shell 9| inches. Hob. Mergui, Malay Peninsula, Sumatra, Borneo. 27. Cyclemys dhor. Emys dhor, part., Gray, Syn. Rept. p. 20. Emys dentata, Gray, III. Ind. Zool. ii, pi. Iviii, fig. 2. Cvclemys oldhamii, Gray. P. Z. S. 1863. p. 178 ; Gilnth. Kept. B. I. p. 15, pi. v, fig. B. Cyclemys dentata, Theob. Cat. p. 8. Cyclemys dhor, Bouleng. Cat. CM. #c. p. 131. Carapace depressed, with a single, obtuse keel ; posterior margin serrated ; vertebral shields as broad as long or broader than long, narrower than the costals. Plastron narrower than the opening of the shell, hind lobe angularly notched, connected with the carapace by a distinct bridge, the width of which is contained twice and three fourths to three times in the length of the plastron ; the hyo- and hypoplastral bones contribute in a nearly equal degree to the formation of the bridge ; the transverse hinge between the hyo- and hypoplastrals, which does not correspond with the curved suture between the pectoral and abdominal shields, is only deve- CTCLEMTS. 31 loped in adult specimens ; a small inguinal shield may be present ; the suture between the pectoral shield and the marginals shorter than that between the abdominal and the margiuals ; the longest median suture is that between the pectorals. Hook of upper jaw bicuspid. Digits distinctly webbed, with sharp claws. Front part of arm with broad transverse scales. Carapace brown, or yellowish brown spotted with dark brown ; plastron yellowish or pale brown in the young, with dark brown spots or radiating lines ; in the adult, dark brown. Neck with dark and light longitudinal lines. Length of shell 7| inches. Hob. Malay Archipelago and Peninsula, Carnboja, Siam, Burma (Pegu, Tenasserim). 28. Cyclemys mouhoti. Cyclemys mouhotii, Gray, A. M. N. H. (3) x, 1862, p. 157 ; Bvuleng. Cat. Chel. $c. p. 132. Pyxidea mouhotii, Giinth. JRept. B. I. p. 16, pi. iv, fig. D ; Theob. Cat. p. 9. Carapace depressed, angular, the vertebral region flat, with three strong parallel keels ; posterior margin serrated ; vertebral shields much broader than long and as broad as the costals ; first vertebral as long as broad or broader than long, narrower than second. Plastron a little narrower than the opening of the shell, hind lobe angularly emarginate; bridge broad and very short, its width about one third the length of the plastron ; the hypoplastral bone and the abdominal shield have a much greater share in the forma- tion of the bridge than the hyoplastral and pectoral ; the longest median suture is that between the abdominals. Hook of the upper jaw strong, entire. Digits nearly free ; limbs covered with large scales. Carapace yellowish or reddish brown ; plastron yellowish, with or without a few large dark-brown spots ; neck of young brown, spotted with yellow. Length of shell 6| inches. Hab. Cachar, Siam, Cochin China. 29. Cyclemys amboinensis. Testudo aniboinensis, Daud. Kept, ii, p. 309. Cistudo amboinensis, Gray, III. Ind. Zool. i, pi. Ixxvii, figs. 3, 4. Cuora amboinensis, Gray, Cat. Sh. Rept. i, p. 41 ; Giinth. Kept. B. 1. p. 12, pi. iv, figs. A, B ; Theob. Cat. p. 7. Cyclemys aniboinensis, Bouleng. Cat. Chel. 8>c. p. 133. Carapace depressed and tricarinate in the young, usually very convex and without or with a single keel in adult females ; verte- bral shields as long as broad or a little longer than broad in the adult, broader in the young, much narrower than the costals. Plastron as large as the opening of the shell in the adult, without distinct bridge; no anal notch; the line of junction between the hyo plastron and the carapace shorter than that between the hypo- plastron and the carapace ; pectoral shields as long as the abdo- minals or shorter, as long as the humerals or slightly longer ; axillary and inguinal small or absent. Head rather small ; upper jaw scarcely hooked, without any emargination. Front part of arm with large transverse scales. Digits moderately webbed, with sharp claws. Carapace brown or blackish ; plastron yellow with large black spots, or dark brown with the suture between the shields yellow ; in the very young the black spots of the plastron are confluent into a broad longitudinal zone, the borders of the plastron being yellowish ; head and neck brown above, yellow beneath ; a yellow band borders the head and neck superiorly, meeting its fellow above the nostrils ; a second yellow band passes through the eye and is separated from the upper jaw by a dark- brown band ; ear yellow. Length of shell 8 inches. Hob. Pegu, Tenasserim, Siam, Malay Peninsula and Archipelago, eastward to the Moluccas. Genus BELLIA, Gray, P. Z. S. 1869, p. 197. Neural plates hexagonal, anterior lateral margins shortest. Plastron extensively united to the carapace by suture, with long axillary and inguinal buttresses, the latter anchylosed between the fifth and sixth costal plates ; entoplastron intersected by the humero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces without median ridge ; choanae between the eyes. Skin of hinder part of head divided into small shields. Digits webbed. Tail very short, not longer in the young than in the adult. Distribution. Burma, Siam, Malay Peninsula and Archipelago. Semiaquatic. 30. Bellia crassicollis. Emys crassicollis, Gray, Syn. Rept. p. 21 ; id. III. Ind. Zool. i, pi. Ixxvi, fig. 2 ; Cantor, J. A. S. B. xvi, 1847, p. 609 ; Giinth. Rept. B. I. p. 28, pi. iv, fig. E. Emys nigra, Blyth, J. A. S. B. xxiv, 1855, p. 713, and xxxii, 1863, p. 81. Bellia crassicolli.s, Gray, P. Z. S. 1869, p. 197; Souleiio. Cat. Cliel. #c. p. 98. Bellia crassilabris, Theob. Cat. p. 10. Carapace moderately depressed, tricarinate in the young, the keels, especially the laterals, becoming indistinct in old specimens ; vertebral region flattened in the adult male ; posterior border serrated ; nuchal small, broadest behind ; posterior side of first vertebral not half the length of the shield in the adult ; vertebrals 2 to 4, as long as broad or a little broader than long, much nar- rower than the costals, narrowly in contact with each other in old BELLIA. 33 specimens ; in the latter, the autero-lateral sides are convex, the postero-lateral longer and concave. Plastron smaller than the opening of the shell, truncate anteriorly, angularly notched pos- teriorly, feebly concave in the males, strongly angulated laterally Fig. 11.— Shell otBellia crassicollis. in young specimens, feebly in old ones ; the width of the bridge about equals the length of the hind lobe ; relative size of plastral shields very variable; abdominals usually forming the longest median suture, humerals the shortest ; axillary and inguinal shields present. Head rather large ; snout very short, not prominent ; upper jaw strongly curved on each side, not hooked ; the width of the mandible at the symphysis is a little less than the diameter of the orbit. Digits webbed to the claws ; front of forearm and outer side of leg with large transverse band-like scales. SheU dark brown or black, plastron usually with yellowish variegation, or bands following the sutures of the shields, or a broad band along each side. Soft parts dark brown or black ; some large yellow spots on the head, viz. above the eye, above the ear, below the ear, D 34 TESTUDINID^E. and on each side of the mandible ; these spots become indistinct in old specimens. Length of shell 7 inches. Httb. Tenasserirn, Siam, Malay Peninsula, Sumatra. Genus DAMONIA, Gray, P. Z. S. 1869, p. 193. Neural plates hexagonal, anterior lateral margins shortest. Plas- tron extensively united to the carapace by suture, with long axillary and inguinal peduncles ; the latter anchylosed between the fifth and sixth, exceptionally fourth and fifth, costal plates ; entoplastron intersected by the humero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces very broad, without median ridge ; choanse behind the level of the eyes. Skin of posterior part of head divided into small shields. Digits webbed. Tail short or moderate. Distribution. East Indies, China, Japan. Semiaquatic. A single species is found in India. 31. Damonia hamiltonii. Emys hamiltonii, Gray, Syn. Eept. pp. 21, 72 ; Giinth. Rept. B. I. p. 32. Emys guttata (non Schn.), Gray, III. Ind. Zool. i, pi. Ixxvi, fig. 1. Damonia hamiltonii, Gray, P. Z. S. 1869, p. 195 ; Theob. Cat. p. 11 ; Bouleng. Cat. Chel. $u. p. 93. Melanochelys pictus, Murray, A. M. N. H. (5) xiv, 1884, p. 107. Carapace much elevated, with three interrupted keels or series of nodose prominences corresponding to the vertebral and costal shields ; posterior border strongly serrated in the young, feebly in the adult ; nuchal moderate, broader posteriorly than anteriorly ; first vertebral not or scarcely broader anteriorly than posteriorly ; second and third vertebrals broader than long in the young, nearly as long as broad in the adult, narrower than the costals. Plastron large, angulate laterally, truncate anteriorly ; posterior lobe much narrower than the opening of the shell, nearly as long as the width of the bridge, deeply notched posteriorly. Head rather large ; snout very short, not projecting ; upper jaw emarginate mesially ; the width of the mandible at the symphysis neai'ly equals the horizontal diameter of the orbit ; a large shield, sometimes divided into three, covers the upper surface of the snout and the crown ; one shield round the upper jaw and one on each side between the eye and the ear. Digits webbed to the claws. Tail extremely short. Shell dark brown or blackish, elegantly marked with yellow spots and radiating streaks. Soft parts dark brown or blackish, with round yellow spots, largest on the head and neck. Length of shell 9 inches. Hal. Bengal, Punjab, Upper Sind. 35 Genus MORENIA, Gray, Suppl. Cat. Sh. Eept. i, p. 62, 1870. Neural plates hexagonal, anterior lateral margins shortest. Plas- tron extensively united to the carapace by suture, with short axil- lary and inguinal buttresses, just reaching the first and fifth costal plates ; entoplastron anterior to the humero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces very broad, that of upper jaw with a strong tuberculate median ridge ; edge of jaws strongly toothed ; choanse behind the level of the eyes. Upper surface of snout and crown covered with a single shield, behind which the skin is corrugated. Digits extensively webbed. Tail short. Distribution. Northern India and Burma. Aquatic ; herbivorous. Synopsis of the Species. Suture between abdominal shields longer than that between pectorals .......................... M. ocellata, p. 3". Suture between abdominal shields not longer than that between pectorals ...................... M. petersii, p. 36. 32. Morenia ocellata. Emys ocellata, Dum. Sf Bibr. Erp. Gen. ii, p. 329, pi. xv, fig. 1 ; Giitith. Eept. B. 1. p. 22. Emys berdmorei, Blyth, J. A. S. B. xxvii, 1858, p. 281. Morenia berdmorei, 'Gray, Suppl. Cat. Sh. Reft, i, p. 62 ; Theob. Cat. p. 17. Batagur (Morenia) ocellata, Anders. An. Zool. Res. Yunnan, p. 755, pis. Ix & Ixi. Morenia ocellata, Bouleng. Cat. Chel. $c. p. 66. Carapace moderately depressed, with a strong, interrupted, tubercular keel in the young, which becomes fainter in the adult ; margin not serrated ; nuchal narrow ; first vertebral not or but little broader anteriorly than posteriorly, lateral borders usually sinuous ; second, third, and fourth vertebrals broader than long or as long as broad, narrower than the costals. Plastron large, angulate laterally, truncate anteriorly, notched posteriorly ; the width of the bridge exceeds the length of the hind lobe ; the longest median suture is that between the abdominals ; suture bet\veen the gulars as long as that between the numerals or shorter ; axillary and inguinal large. Head moderate ; snout short, obtuse ; upper jaw notched mesially, denticulated ; lower jaw strongly serrated, flat beneath, its width at the symphysis equalling the diameter of the orbit. Scales on limbs small ; digits broadly webbed. Tail very short. Carapace brown, each shield of the disk with a large central blackish ocellus encircled with yellowish ; lower surface uniform yellow. Head olive, with yellow D2 markings ; a yellow streak running above the canthus rostralis, supraciliary edge and temple, and another from behind the eye to above the ear. Length of shell : d1 6 inches, § 9 inches. Hab. Burma. 33. Morenia petersii. Emys ocellata (non D. &• B.), Blytli, J. A. S. B. xxvii, 1858, p. 281. Morenia ocellata, Theob. Cat. p. 18. Batagur (Morenia) petersi, Anders. An. Zool. Res. Yunnan, p. 701, pi. lix. Morenia petersii, Bouleng. Cat. Chel. $c. p. 08. Very closely allied to the preceding. Snout much more pointed and relatively longer. Suture between the pectorals not shorter than that between the abdominals. Carapace black, each vertebral with a narrow yellowish mesial line ; the last four vertebrals with a yellowish linear horseshoe mark with the ends directed forwards ; each costal with an ocellus placed rather low and formed by a narrow yellowish line, above which are some irregular looped lines of similar colour ; the nuchal and each marginal with a vertical narrow yellowish mesial streak ; plastron yellow. Three yellow streaks on each side of the head, the lower extending from below the nostril to the angle of the mouth. Length of shell : 3 5 inches, $ 8 inches. Hab. Bengal. Genus HARDELLA, Gray, Suppl. Cat. Sh. Kept, i, p. 58, 1870. Neural plates elongate, hexagonal, anterior lateral margins shortest. Plastron extensively united to the carapace by suture, with extremely developed axillary and inguinal buttresses, the for- mer connected with the first rib, the latter anchylosed between the fifth and sixth costal plates ; entoplastron anterior to the humero- pectoral suture. Skull with a bony temporal arch ; alveolar surfaces very broad, that of upper jaw with a strong tubercular median ridge ; edge of jaws strongly toothed ; choana) behind the level of the eyes. LTpper surface of snout and crown covered with a single shield, behind which the skin is corrugated. Digits extensively webbed. Tail short. Distribution. Northern India. Thoroughly aquatic ; herbivorous. 34. Hardella thurgi. j, Syn. Kept. p. 2 Emys thurgii, Giinth. Rept. B. I. p, Emys thurjii, Gray, Syn. Rept. p. 22 ; id. 111. Ind. Zool i, pi. Ixxiii. BAT AGUE. 37 Batagur thurgii, Theob. Cat. Rept. Mus. As. Soc. 1868. p. 12 : id. Cat. p. 23. Kachuga oldhami, Gray, P. Z. 8. 1869, p. 200, fig. 14. Hardella thurgi, Gray, Suppl Cat. Sh. Rept. i. p. 58 ; Bouleng. Cat. CM. $c. p. 63. Hardella indi, Gray, Suppl. Cat. Sh. Rept. i, p. 58. Carapace moderately depressed, with an interrupted tubercular keel ; margin not serrated ; nuchal narrow, broadest posteriorly ; first vertebral usually narrower anteriorly than posteriorly, lateral borders usually slightly sinuous ; second, third, and fourth verte- brals considerably broader than long in the young, nearly as long as broad in the adult. Plastron large, angulate laterally, truncate anteriorly, angularly notched posteriorly ; the width of the bridge exceeds the length of the posterior lobe ; the longest median suture is that between the abdominals ; suture between the gulars as long as that between the humerals or a little shorter (at least three fourths) ; inguinal large, axillary smaller. Head moderate ; snout short, obtuse ; upper jaw notched mesially, serrated ; lower jaw very strongly dentated, flat inferiorly, its width at the symphysis a little less than the diameter of the orbit. Limbs with transversely enlarged, band-like scales. Carapace dark brown or black ; plas- tron yellowish, with a large blackish spot occupying the greater part of each shield, or entirely black. Soft parts dark brown ; a yellow band across the prsefrontal region, extending on each side, through the eye, to above the ear ; another from below each eye, passing below the nostrils. Length of shell 18 inches ( $ ). Males much smaller. Hub. Northern India (Ganges and Indus systems). Genus BATAGUR, Gray, Cat. Sh. Rept. i, p. 35, 1855. Neural plates elongate, hexagonal, anterior lateral margins shortest. Plastron extensively united to the carapace by suture, with extremely developed axillary and inguinal buttresses, the for- mer connected with the first rib, the latter anchylosed between the fifth and sixth costal plates ; entoplastron anterior to the humero- pectoral suture. Skull with a bony temporal arch ; alveolar surfaces very broad, that of the upper jaw with two strong, slightly den- ticulated median ridges ; edge of jaws denticulated ; choanse be- hind the level of the eyes. Limbs somewhat approaching the paddle-shape, very broadly webbed, with four claws. Tail very short. Distribution. Bengal, Burma, Malay Peninsula. Thoroughly aquatic ; herbivorous. TESTUDINID.E. 35. Batagnr baska. Emys batagur, Gray, Syn. Rept. p. 23 ; id. 111. Ind. Zool. ii, pi. lix. Emys baska, Gray, Syn. Rept. p. 24 ; id. 111. Ind. Zool. i, pi. Ixxv. Tetraonyx baska, Dum. $ Bibr. Erp. Gen. ii. p. 341 ; Theob. Cat. p. 25. Tetraonyx batagur, Gray, Cat. Tort. p. 29. Tetraonyx affinis, part., Cantor, J. A. S. B. xvi, 1847, p. 612. Batagur baska, Gray, Cat. Sh. Rept. i, p. 35, pi. xvi; Giinth. Rept. B. I. p. 37, pi. iii, figs. B, B'; Bouleng. Cat. Chel. fyc. p. 01. Carapace moderately depressed, with a vertebral keel in the young, which keel disappears in the adult ; nuchal broader than long ; first vertebral as broad in front as behind, or a little broader ; vertebrals 2 to 4 subequal, much broader than long in the young, nearly as long as broad and as broad as the costals in the adult ; the postero-lateral border of the third vertebral strongly concave. Plastron large, strongly angulate laterally in the young, convex in the adult, truncate anteriorly, angularly notched posteriorly ; the width of the bridge exceeds the length of the posterior lobe ; the longest median suture is that between the abdominals, the shortest that between the gulars, the latter never more than half that be- tween the hutnerals ; inguinal large, axillary smaller. Head rather small ; snout pointed, produced, directed upwards ; jaws with denticulated edge, upper feebly notched mesially ; the width of the lower jaw at the symphysis nearly equals the diameter of the orbit. Limbs with transversely enlarged, band-like scales. Upper surface of shell and soft parts olive-brown, lower surface yellowish. Length of shell 21 inches. Hab. Bengal, Burma, Malay Peninsula. Genus KACHUGA, Gray, Cat. Sh. Rept. i, p. 35, 1855. Fourth vertebral shield elongate, embracing four or five neural plates ; neural plates hexagonal, anterior lateral margins shortest. PJastrou extensively united to the carapace by suture, with ex- tremely developed axillary and inguinal buttresses, the former con- nected with or nearly reaching the first rib, the latter anchylosed between the fifth and sixth costal plates ; entoplastron anterior to the humero-pectoral suture. Skull with a bony temporal arch ; alveolar surfaces very broad, that of upper jaw with a median ridge ; edge of jaws denticulated ; choanae oa a level with the posterior border of the eyes, or behind it. Upper surface of head covered with undivided skin. Digits very broadly webbed. Tail very short. Distribution. India and Burma. Thoroughly aquatic ; her- bivorous. KACHUGA. 3! Synopsis of tlie Species. A. Third vertebral shield forming a broad suture with the fourth, which embraces four neural plates. a. Second vertebral shield with straight trans- verse posterior border. a'. Alveolar surface of upper jaw extremely broad, the median ridge nearer the outer than the inner margin ; bony choanae be- hind the orbits K. lineata, p. 40. b'. Alveolar surface of upper jaw narrower, the median ridge nearer the inner than the outer margin ; bony choanae between the orbits K. trivittata, p. 41 . Fig. 12.— Shell of Kachuga trivittata. b. Second vertebral shield pointed or produced behind, entering the third K. dhongoka, p. 41. B. Fourth vertebral shield pointed in front, nar- rowly in contact with the third, embracing five neural plates. a. Third vertebral quadrangular, much longer than broad, posterior border straight or slightly convex K. smithii, p. 42. 40 TESTUDINID^!. b. Third vertebral shield pentagonal, pointed be- hind, a'. Second vertebral shield much shorter than third, embracing two neural plates. a". Posterior margin of carapace strongly ser- rated ; 26 marginal shields (including caudals) K. sylhetensis, p. 42. b". Posterior margin not serrated K. intermedia, p. 43. b'. Second vertebral shield as long as broad, embracing three neural plates K. tectum, p. 43. 36. Kachuga lineata. Emys dhor, part., Gray, Syn. Kept, p. 20. Emys dentata, part, Gray, Syn. Rept. errata, & pis. via, ix ; id. III. Ind. Zool. ii, pi. Iviii, fig. 1. Emys lineata, Gray, Syn. Rept. p. 23. Einvs kachuga, Gray, III. Ind. Zool. i, pi. Ixxiv. Batagur lineata, Gray, Cat. Sh. Rept. i, p. 35, pi. xvii ; Gilnth. Rept. B. I. p. 39. Batagur ellioti, Gray, P. Z. S. 1862, p. 264 ; Gilnth. Rept. B. 1. p. 40, pi. iii, figs. A, A'. Kachuga hardwickii, Gray, P. Z. S. 1869, p. 202. ? Kachuga fusca, Gray, Suppl. Cat. Sh. Rept. i, p. 56. Kachuga liueata, Gray, Suppl. Cat. Sh. Rept. i, p. 56 ; Boideng. Cat. Chel. Sfc. p. 54. Batagur kachuga, Theob. Cat. p. 19. Carapace of young strongly keeled, the keels tubercular poste- riorly on the second and third vertebral shields, posterior margin strongly crenulated ; the marginal serrature disappears in adoles- cent specimens, and the vertebral keel, after being reduced to a series of low knobs, vanishes entirely in the full-grown, the cara- pace of which is very convex. Nuchal shield small, trapezoid, broadest posteriorly ; first vertebral as broad in front as behind, or broader ; second vertebral longer than third, with which it forms a straight transverse suture ; fourth longest and forming a broad suture with the third ; second vertebral broader than long in the young, as long as broad in the adult. Plastron angulate laterally in the young ; anterior and posterior lobes rather narrow and shorter than the width of the bridge, truncate anteriorly, openly notched posteriorly ; the longest median suture between the abdominals, the shortest between the gulars, which equals about one half that between the humerals ; the suture between gulars and humerals forms an obtuse angle, and so does that between humerals and pectorals; inguinal large, axillary smaller. Head moderate ; snout obtuse, moderately prominent ; jaws with denti- culated edge, upper not notched mesially ; alveolar surfaces very broad, the median ridge of the upper jaw being somewhat nearer the outer than the inner margin ; choanse behind the line of the posterior borders of the orbits ; the width of the lower jaw at the symphysis equals the diameter of the orbit. Limbs with trans- KACHUGA. 41 versely enlarged, baud-like scales. Brown above, yellowish below ; nape with red longitudinal lines. Length of shell 15 inches. Hab. Northern Peninsular India ; Ganges, Kistna, and Godavari rivers; Burma. 37. Kachuga trivittata. Emys trivittata, Dum, $ Bibr. Erp. Gen. ii, p. 33 1. Batagur trivittata, Theob. Jour. Linn. Soc. x, 1808, p. 14, and Cat. p. 20. Kachuga peguensis et trilineata, Gray, P. Z. S. 1869, p. 200. Batagur iravadica, Anders. An. Zool. Res. Yunnan, p. 736, pis. Ixiv, Ixv, Ixviii, & Ixix. Kachuga trivittata, Bouleng. Cat. Cliel. SfC. p. 55. Differs from the preceding in the much narrower alveolar sur- face of the jaws, the median ridge of the upper jaw being nearer the inner than the outer margin ; choanae between the orbits ; the width of the lower jaw at the symphysis is less than the dia- meter of the orbit. Considerable uncertainty prevails in the dis- tinction of isolated shells of this species from the preceding. The male B. trivittata is characterized by three black longitudiinal bands on the carapace, which are absent in the female according to Theobald, whose opinion I am disposed to endorse rather than that of Anderson, who makes a distinct species (B. iravadica) for the specimens (females and young males) without bands. Length of shell 22 inches. Hab. Irrawaddy river from Bhamo to Bangoon ; Tenasserim. 38. Kachuga dhongoka. Emys dhongoka, Gray, III. Ind. Zool. ii, pi. Ix. Batagur dhongoka, Gray, Cat. Sh. Rept. i, p. 36, pi. xviii ; Gunth. Rept. B. I. p. 42 ; Theob. Cat. p. 22. Pangshura leithii, Gray, Suppl. Cat. Sh. Rept. i, p. 60. Kachuga dhongoka, Bouleng. Cat. Chel. fyc. p. 56. Carapace much depressed, keeled in the young, the keels reduced to a knob on each of the anterior vertebrals in the adult ; posterior margin crenulated. Nuchal shield small, trapezoid, broadest behind ; first vertebral usually narrower in front than behind, with sinuous lateral borders ; second vertebral broader than long in the young, longer than broad in the adult, posterior border pointed or more or less produced, fitting into an emargination of the third vertebral, which is shorter than either the second or the third. Plastron angulate laterally in the young ; front lobe truncate, hind lobe angularly notched, shorter than the width of the bridge ; the longest median suture is between the abdominals ; suture between the gulars as long or nearly as long as that between the humerals ; the suture between gulars and humerals forms a right angle, and 42 TESTUDINID^E. that between humerals and pectorals a straight transverse line ; inguinal large, axillary smaller. Jaws and soft parts as in K. trivittata. Brown above, yellowish below ; carapace with three more or less distinct black longitudinal bands. Length of shell 14 inches. Hob. Northern India (Granges and Indus) ; and Poona in the Deccan, if a specimen obtained by Dr. Leith was correctly labelled. 39. Kachuga smitMi. Balagur smithii, Gray, P. Z. S. 1863, p. 253. Pangshura smithii, Oiinth. Rept. B. I. p. 36 ; Theob. Cat. p. 15. Kachuga smithii, Bouleng. Cat. Chel. fyc. p. 57, pi. i. Carapace much depressed, feebly keeled. Nuchal shield small, trapezoidal, broadest posteriorly ; first vertebral with sinuous lateral borders, usually a little narrower in front than behind ; second vertebral shortest, broader than long, usually with straight or slightly convex posterior border ; third vertebral considerably longer than broad, subquadrangular, posterior border straight or slightly convex ; fourth vertebral longest, tapering anteriorly and forming a narrow suture with the third ; fifth vertebral much broader than the others. Plastron feebly angulated laterally, large ; front lobe rounded, hind lobe angularly notched and as long as or a little shorter than the width of the bridge ; the longest median suture is that between the abdominals, which about equals the length of the front lobe ; gulars usually shorter than the suture between the humerals, their suture with the latter shields forming a right angle ; inguinal large, axillary smaller. Head moderate ; snout short, obtuse, feebly prominent ; jaws with denticulated edge, upper not notched mesially ; alveolar surface of upper jaw broad, the median ridge nearer the inner than the outer border ; bony choanso between the orbits ; the width of the lower jaw at the symphyeis is less than the diameter of the orbit. Fore limbs with large transverse scales. Pale olive-brown above ; dorsal keel usually blackish ; plastral shields and lower surface of mar- ginals dark brown, bordered with yellow. Length of shell 8^ inches. Hob. Upper Gauges and Indus with their tributaries. 40. Kachuga sylhetensis. Pangshura sylhetensis, Jerdon, P. A. S. B. 1870, p. 69 ; Theob. Cat. p. 15. Kachuga sylhetensis, Bouleng. Cat. Chel. fyc. p. 57. Carapace elevated, tectiform, the keel ending in an elevated pointed nodosity on the third vertebral shield, forming a strong ridge on the last two vertebral s ; posterior margin very strongly serrated ; 26 marginal shields, instead of 24 as in all other species of the genus ; first vertebral shield much broader in front than behind in the half-grown specimens, equally broad in front and KACHUGA. 43 behind in the adult ; second vertebral shortest, broader than long, with straight posterior border ; third vertebral considerably longer than broad, pointed behind, and forming a very narrow suture with the fourth, which tapers anteriorly and equals in length the first and second together ; fifth vertebral not broader than fourth, pointed posteriorly. Plastron large, feebly angulate laterally, truncate anteriorly, angularly notched posteriorly ; the longest median suture is that between the abdominals ; suture between the gulars as long as or shorter than that between the numerals; suture between gulars and humerals forming a right angle ; inguinal and axillary large. Carapace olive-brown, usually with a lighter streak along the keel ; plastral shields dark brown with a broad yellow border. Length of shell 7 inches. Hob, Sylhet and Assam. 41. Kachuga intermedia. Emys (Pangshura) tectum, var. intermedia, Blanf. J. A. S. £. xxxix, 1870, pt. 2, p. 339, pi. xiv, and xlviii, 1879, p. 110. Kachuga intermedia, Bouleng. Cat. Chel. fyc. p. 58. Shell as in K. tectum, but second vertebral shield shortest and broader than long. Carapace brown ; plastral shields black, with yellow anterior and lateral margins. Head dull oh' ve, paler below ; a ferruginous spot behind the eye, and three others, less well marked, in a convex line on the occiput ; limbs uniform olive. Length of shell 4| inches. Hob. Hasdo river, Bilaspur, Central Provinces ; Godavari. This Tortoise is only known to me from Blanf ord's description. 42. Kachuga tectum. Emys tecta, Gray, Syn. Kept. p. 23, pi. v ; id. III. Ind. Zool. i, pi. Ixxii. Emys tentoria, Gray, P. Z. S. 1834, p. 54. Pangshura tecta, Gunth. Rept. B. 1. p. 33 ; Theob. Cat. p. 13. Pangshura tentoria, Gunth. Rept. B. 1. p. 34, pi. iv, figs. C, C'; Theob. Cat. p. 14. Pangshura flaviventer, Gunth. Rept. B. I. p. 35 ; Theob. Cat. p. 15. Kachuga tectum, Bouleng. Cat. Chel. $c. p. 58. Carapace elevated, tectiform, the keel ending in a nodosity on the third vertebral shield ; posterior margin not or but very slightly serrated ; nuchal shield small, square or trapezoidal ; first vertebral very variable in shape, usually with straight lateral borders diverg- ing forwards in the half-grown specimens, narrower in front and with sinuous lateral borders in the adult ; second vertebral as long as or a little longer than second, frequently obtusely pointed behind ; third vertebral pointed behind, in contact with the point of the very elongate fourth ; fifth vertebral broader than the others. Plastron large, strongly angulated laterally in the young, truncate anteriorly, angularly notched posteriorly ; proportions of 44 PLATYSTEENID^E. plastral shields very variable ; suture between gulars and humerals forming a right angle ; axillary and inguinal large. Head mode- rate ; snout short, rather pointed and prominent ; jaws with den- ticulated edge, upper not notched mesially ; alveolar surface of upper jaw with the median ridge nearer the inner than the outer border ; bony choanse between the orbits ; the width of the lower jaw at the symphysis is less than the diameter of the orbit. Fore limbs with large transverse scales. Carapace olive, that of young with some black spots, especially on the posterior edge of the first three vertebrals ; an orange vertebral band and a narrow yellow margin ; carapace more uniform in the adult ; plastron orange or red, with black spots, or brown with a yellowish anterior and lateral border to each shield, in one specimen uniform yellow. Head blackish ; jaws and sides of crown orange ; neck with numerous yellow lines on a blackish ground ; limbs dark olive, spotted with yellow. Length of shell up to nearly 9 inches. Hah. Ganges and Indus systems. Family III. PLATYSTERNID.E. Shell covered with epidermal shields. Caudal vertebras mostly opisthocoelous. Neck completely retractile within the shell. Parietal bones widely separated from squamosals ; temporal region completely roofed over. Digits moderately elongate ; phalanges with condyles ; claws four or five. A single genus and species, remarkable for its large head and long tail. Genus PLATYSTERNUM, Gray, P. Z. S. 1831, p. 106. Plastron moderately large, connected with the carapace by liga- ment ; a series of inframarginal shields separates the plastral shields from the marginals. Head very large, covered above with an undivided horny shield; jaws very strong, hooked, without additional alveolar ridges. Digits feebly webbed, all except the outer toe clawed. Tail very long, subcylindrical, compressed at the end, with rings of squarish shields. A single species. 43. Platystenmm megacephalum. Platysteraum megacephalum, Gray, P. Z. S. 1831, p. 107 ; id. Ill Ind. Zool. ii, pi. Ixii ; Giinth. Rept. B. I. p. 43 ; Bouleng. Cat. Chel. 8fc. p. 46. Platysternon peguense, Gray, Suppl. Cat. Sh. Rept. i, p. 70 ; TJieob. Cat. p. 26. PLATYSTEENUM. Carapace much depressed, oval, with a feeble median keel poste- riorly ; anterior border broadly emarginate. Nuchal shield very small, broader than long ; vertebrals broader than long, as broad as the costals or a little narrower, much broader in the young, in which the posterior border of the carapace is distinctly serrated. V Fig. 13. — Skull of Platysternum megaccphal; Front lobe of plastron squarish, posterior angularly emarginate ; the width of the bridge contained four or five times in the length of the plastron ; the longest plastral shields are the anals, femorals, and humerals ; the shortest the gulars, which occupy, however, the whole width of the front border of the plastron ; frequently a small intergular shield between the gulars and the humerals. Upper surface of head and temple with an undivided horny scute. 46 PLATYSTERXIDJE. Throat with round flat tubercles. Anterior surface of arm with large squarish scutes. Hinder side of thighs with large conical tubercles. Tail at least as long as the shell. Carapace and soft parts of adult olive-brown ; plastron yellowish brown. Young more elegantly marked : — Upper surface of head and neck and carapace olive-brown, with a few dark brown dots on the crown and one in the centre of each costal scute ; the edge of the carapace yellow ; jaws yellow ; a yellow black-edged streak on each side along the temple ; sides and lower surface of neck and limbs whitish ; upper surface of limbs blackish : plastron yellow, with a symmetrical black marking along the middle ; tail blackish above, yellow with a black median line beneath. Head and neck 3 inches, carapace 6, tail 6^. Fig. 14.— Shell of Platysternum megacephalum. A rare tortoise, found in streams in Burma and Siam ; the speci- mens on record are mostly from Pegu and Tenasserim. It is also known from Southern China, where, according to Svvinhoe (P. Z. S. 1870, p. 409), it occurs in the western parts of Kwangtung Province and in Kwangse. CHELOHTD.B. 47 Family IV. CHELONID^. Shell covered with epidermal shields. Caudal vertebrae pro- coelous. Neck incompletely retractile ; cervical vertebrae short. Fig. 15.— Skull of Chelone imlricata. Temple completely roofed over ; parietal bone in contact with the squamosal. Limbs paddle-sbaped ; phalanges without condyles : claws OTIC or two. Tbe Marine Turtles belong to three species, falling into two 48 CHELONID^E. genera, viz. :— The Green Turtle, Chelone mydas, the Havvksbill, Chelone imbricata, and the Loggerhead, Thalassochelys caretta, all three of which occur in the Indian Ocean. The Green Turtle is valuable as an article of food, the Hawksbill for the tortoise-shell (the epidermal horny shields) ; whilst the Loggerhead is of no commercial value, or produces but a very inferior quality of tortoise- shell. The Chelonidae are thoroughly marine turtles, approaching the shores only for the purpose of depositing their eggs, which are very numerous, soft-shelled, and buried in the sand. As these eggs are large and excellent eating, they are much sought for. The genera are thus distinguished : — Costal shields 4 pairs CHELONE, p. 48. Costal shields 5 or more pairs THALASSOCHELYS, p. 49. Genus CHELONE, Brongniart, Bull. Soc. Philom. iii, 1800, p. 89. Carapace with persisting fontanelles between the costal and marginal plates. Costal shields four pairs. An iutergular shield and a series of inframargiual plastral shields. Two species are known. Synopsis of Species. Upper jaw not hooked C. mydas, p. 48. Upper jaw hooked C. imbricata, p. 49. 44. Chelone mydas. Testudo mydas, Linn. Syst. Nat. i, p. 350. Chelonia mydas, Schweigy. Prodr. p. 22; Bouleng. Cat. Chel. $c. p. 180. Chelonia virgata, Schweir/g. Prodr. p. 22 ; Cantor, J. A, S. B. xvi, 1847, p. 617 ; Gunth. liept. B. I. p. 53 ; Theob. Cat. p. 33. Carapace feebly unicarinate in the young (sometimes with slight indication of lateral keels), arched or subtectiform in the adult ; dorsal shields juxtaposed ; margin not or but indistinctly serrated, formed of 25 shields. Snout very short ; jaws not hooked ; horny sheaths of upper jaw with feebly denticulated edge and striated inner surface, of lower jaw with strongly denticulated edge; alveolar surface of upper jaw with two strong denticulated ridges ; symphysis of lower jaw short ; one pair of prsefrontal shields. Limbs usually with a single claw; the second digit sometimes provided with a distinct claw in young specimens. Young dark brown or olive above, the limbs margined with yellow : yellow beneath, with a large dark brown spot on the hand and foot. Carapace of adult olive or brown, spotted or marbled with yellowish. Length of shell 4 feet. Hob. Tropical and subtropical seas. Eather rare in the Bay of Bengal. Herbivorous ; edible, but occasionally poisonous. THALASSOCHELYS. 49 45. Chelone iiubricata. Testudo imbricata, Linn. Syst. Nat. i, p. 350. Chelonia imbricata, Schweigg. Prodr. p. 21 ; Bouleng. Cat. Chel. $c. p. 183. Eretmochelys squamata, Ayassiz, Conti: N. H. U. S. i, p. 382. Caretta squamata, Giinth. Rept. B. I. p. 54 ; Theob. Cat. p. 33. Fig. 16. — Chelone imbricata (young). Carapace tricarinate in the young, with the shields strongly imbricate, the vertebrals rhomboidal ; dorsal shields of adult smooth, of old specimens juxtaposed ; margin formed of 25 shields, more or less strongly serrated posteriorly. Snout compressed in the adult ; jaws hooked, not or but very feebly denticulated on the margin ; symphysis of lower jaw long ; two pairs of shields anterior to the azygous frontal. Limbs with two claws. Young pale brown above, blackish below. Carapace of adult marbled yellow and dark brown, plastron yellow ; shields of head and limbs dark brown with yellow borders. Length of shell 3 feet. Hab. Tropical and subtropical seas. Plentiful on the coasts of Ceylon and the Maldives. Carnivorous. Genus THALASSOCHELYS, Fitzinger, Ann. Wien. Mus. i, 1835, p. 121. Carapace completely ossified in the adult. Costal shields five pairs or more. A series of infrarnargmal plastral shields. A single species. E 50 SPHAEGIDJE. 46. Thalassochelys caretta. Testudo caretta, Linn. Syst. Xat. i, p. 351. Chelonia caouanna, SeJttDeigg. Prodr. p. 22. Chelonia olivacea, Eschsch. Zool. Ail. p. 3, pi. iii ; Cantor, J. A. S. B. xvi, 1847, p. 619. Caouana olivacea, Giinth. Rept. B. I. p. 52 ; Theob. Cat. p. 33. Thalassochelys caretta, Bouleng. Cat. Chel. $c. p. 184. Carapace of young with three strong keels, of adult arched or subtectiform ; dorsal shields juxtaposed ; margin serrated pos- teriorly in the young, formed of 27, or rarely 25 shields. Intergular shield very small or absent. Head large, with very strong, hooked jaws ; symphysis of lower jaw very long ; alveolar surface of horny beak with a median ridge, of jaws without ; two pairs of shields anterior to the azygous frontal. Limbs of young usually with two claws, of adult frequently with but one. Young uniform dark brown or blackish ; adult brown above, yellowish inferiorly. Length of shell 4 feet. Hob. Tropical and subtropical seas ; Mediterranean ; an acci- dental visitor in the North. Not rare in the Bay of Bengal. Carnivorous, feeding chiefly ou crustaceans and niollusca. Suborder II. ATHEC^E. Family V. SPHARGLD^E. Exoskeleton consisting of mosaic-like juxtaposed plates. Plastral elements eight ; no entoplastroii. Limbs paddle-shaped, clawless, the digits of the fore limb much elongate ; phalanges without condyles. This family contains a single genus and species, the well-known Leathery Turtle. Genus DERMOCHELYS, Blainville, Journ. de Phys. Ixxxiii, 1816, p. 259. Dorsal shield completely, ventral shield incompletely bony in the adult, the former with seven, the latter with five keels. Beak with two triangular cusps, between three deep notches ; no enlarged alveolar surface, the jaws simply sharp-edged. Head covered with small shields. 47. Dermochelys coriacea. Testudo coriacea, Linn. Syst. Nat, i, p. 350. Sphargis coriacea, Gray, Syn. Rept. i, p. 51. Dermatochelys coriacea, Giinth. Rept. B.I. p . 55 ; Theob. Cat. p. 34. Dermochelvs coriacea. Boulenq. Cat. Chel.