Ifi» THE THOMAS SAY FOUNDATION OF THE ENTOMOLOGICAL SOCIETY OF AMERICA ESTABLISHED 1915 LA FAYETTE, INDIANA 1916 VUnPHEVBIVIHS CO. PPESS, LAFAYETTE, INO. SARCOPHAGA AND ALLIES IN NORTH AMERICA by J. M. ALDRICH of the United States Bureau of Entomology Genitalium disquisitio abomi?iabilis displicet. — A maxim of Linnaeus, quoted by Fabricius in Illiger's Magazin fiir Insektenkunde, ii, p. 5, 1803. La Fayette, Indiana 1916 S3b' 'I INTRODUCTION The genus Sarcophaga was established by Mei- gen in 1826; he described thirty species, but in the manner of his time did not single out one of these as type of the genus. Westwood in 1840 designated or mentioned the Musca carnaria of Linnasus (in- cluded by Meigen) as the type species. Some ob- jection to Westwood's designations in general has been raised by Hopkins, on the ground that he lim- ited himself to British species. However, the same designation in Sarcophaga was made twice more within a few years, by Zetterstedt and Rondani. Carnaria has been universally accepted as type since that period.* Although Meigen in founding the genus gave a fairly full and accurate account of the genitalia of the male, these were little studied or entirely ignored for many years afterward. The numerous European species were never clearly differentiated until Pan- delle published his remarkable revision in 1896; in this he identified 63 species and varieties and recogniz- ably described them, including the genitalic char- acters of each. Even then, the lack of illustrations in his work prevented iimnediate recognition of the ])rilliant success of the method, and it was a decade before it made an appreciable impression. Since that time, however, the group has been thoroughly studied in the same manner by otlier European dipterists, and its classification for that fauna is on a far more satis- *References for this paragraph : Meisen, Svsteinatlsclie Beschreibung eiirop. Zwelfl. Ins., vol. v, 14, 1S26. Westwood. Tntrndnction to the Modern Classification of Insects, ii, Appendix. 140. l.S-IO. Hopkins. Procopdin^rs U. S. National Musenni. vol. 48, 115, 1914. Zetterstedt. Diptora Soandinavine. iv, p. 1281, 1S4.5. Rondani, Dipterologiae italieae Pi-odonuis, i, S6, 1S56. 6 THOMAS SAY FOUNDATION factory basis than that of any other Muscoid group. Bottcher in particular has given a beautiful con- densed synopsis of the European species, with dia- grams of the male genitalia; he made out about 80 species and varieties. Through his efforts and those of Villeneuve and Kramer, most of the old collections have been re-examined and the types of the early descrij)tions elucidated.* In the United States since the beginning of any indigenous dipterology about 1880 there has been a virtual blockade in this group, due to the vast num- ber of unrecognizable old descriptions, of which the types were scattered in European museums or lost. On account of this condition, Williston and Coquil- lett described new species only under some special stress of circmnstances. Dr. Hough, with character- istic energy, gathered up all the available material in collections, added to it largely himself, and projected a revision ; but before he had made much progress, in 1903, he decided that his medical practice required his undivided attention. So he sold his collection, includ- ing the Sarcophagidffi, to the University of Chicago. Van der Wulp had in the meantime added a list of Mexican and Central American forms. The writer's Catalogue of North American Diptera, published in 1905, shows 106 nominal species of Sarcophaga from North America, of which it is safe to say that hardly more than half a dozen were ever recognizably de- scribed. In the absence of any clear understanding of the specific characters, this looked like the most hopeless field imaginable. The first American worker to attack the group by the new genitalic method was Dr. Ralph R. Par- ker; he began in the winter of 1912-13 a study of the New England representatives. His first paperf con- tains a thorough and admirable discussion of the ♦References for this paragraph: Pandelle, Revue entomologiqiie, xr. 173-207, 1897. Boettcher. Deutsche ent. Zeitschrift, 1912, 525-544 and 705-736; and 1913, 1-16, 115-130, 239-254, .351-377. tProceedings Boston Sec. Nat. Hist., 35. 1-77, 8 plates, 1914. SARCOPHAGA AXD ALIJES 7 morphologj% together with full descriptions of the males of seven species in the genera (or subgenera) Ravinia and Bottcheria. At the request of the late F. M. Webster, then in charge of the section of Cereal and Forage Insect Investigations of the United States Bureau of Entomology, the writer undertook, in December, 1913, the project of a revision of the North American Sarcophagidfe. The necessity for this arose from fre- quent rearings of Sarcophagas as important parasites of grasshoppers, and the desire to get these parasites named and classified so that they might be properly studied. Limits of the Group. No distinct limits to the family Sarcophagid^e have appeared in recent literature. On the one hand, and most commonly, they have been lumped with Tachinidge ; on the other the family is extended to in- clude Metopia and Senotainia, which from the bare or nearly bare arista have been usuallv included in Tachinidfe. In the present work, I have endeavored to in- clude all Muscoid species which agree in having gen- eral gray or silvery, tessellated or changeable polli- nose coloration, the fourth vein ending considerably before the apex of the wing and with an almost angu- lar bend, the parafacials hairy, the arista plumose above and below for half its length or a little more. None of the species have discal macrochaet^e on tlie abominal segments, hairy eyes, long proboscis, rudi- mentary palpi, or more than a single pair of discal scutellar bristles. Species ordinarily considered Tach- inid are excluded by the bare arista ; most Dexiids bv the greater extent of the plumosity of the arista and bv discal abdominal or scutellar bristles, or combina- tions of these characters; Muscid^e (sens, str.) by the arista being plumose to the tip, the color often metal- lic, the a])domen less bristly, the fourth vein often with rounded bend, or combinations of these; Calliphora and Cynomyia, which have an arista like Sarcophaga, 8 THOMAS SAY FOUNDATION are sufficiently distinguished by tlie metallic abdomen, although there are other differences. Among the species included in the present paper, all have been assigned to Sarcophaga except a few aberrant forms with arista short-plumose, or with vi- brissse high u\) and approximated, or with a few other analogous characters. See table of genera for these. Characters of one sex are not used for ultimate sep- aration of genera, though they are often of much j)ractical value. The genus Sarcophaga is undoubtedly a large and rather homogeneous group, a natural large genus, as much so as Tabanus, Rhamphomyia, Culex, Bem- bidium, Formica, etc. Even granting the separation of some of its components in accordance with the views of recent writers, there remains a mass of several hun- dred species, scattered all over the world. After much consideration, I have decided not to attempt the separation of the genera Bliesoxipha, Ra- vinia, and Bottcheria, but to regard them as sub- genera or merely groups having some common char- acters. Blffisoxipha (Loew, Wiener Ent. Monatsch., v, 384, 1861) was established on a single female speci- men (European) described as grylloctona n. sp. in the same place. The only character differing from Sarcophaga as announced at the time was the pres- ence of a long larvipositor curved downward and for- ward. The group of species possessing more or less of this character is very complex and closely related in Europe as well as in North America; it has not yet been well worked out in Europe. Species num- bered from about 52 to 66, of the present paper, and a few others, are more or less of this type in the fe- male ; but the amount of development of the sternites is often slight, usually so. Figures 61b and 61c show the extreme form, from which a series might be made of species showing less and less development of larvi- positor, down to an ordinary Sarcophaga. Further- more, the species are so similar in the middle part of SARCOPHAGA AND ALLIES 9 this series that they can be separated only by the male genitalia, and probably a dozen of them can scarcely be separated at all in the female. It would seem like an over refinement of classification if we could deter- mine the genus only by the female genitalia, and the species only by those of the male; still more so as in this group it is almost impossible to tell which male goes with which female. We must take the species as nature makes them, but fortunately nature rarely makes genera, leaving that usually to the convenience of the taxonomist. I am not aware of any diagnostic characters for Bltesoxipha in the male sex. Ravinia (Desvoidy, Dipt. Env. Paris, ii, 434, 1863; Parker, Proc. Bost. Soc. Nat. Hist., 35, 52, 1914) has for its principal character the course of the frontal bristles, which diverge anteriorly but little, or not at all in the females of some species. The type is hcematades of Europe, and the nearest allies in the present paper are Nos 49, 117-121, 143-145. The character of the frontal rows is less developed in males, and there are many species of Sarcophaga out- side the group which show it; however, with some other rather minute characters, it marks off a good subgenus. Bottcheria (Parker, Proc. Bost. Soc. Nat. Hist., 35, 65, 1914) has for its principal character the pres- ence of a third (middle) series of long bristles on the outer side of the hind femur; the ventral plates also do not become successively narrower. Mr. Parker excluded the females from consideration in his pajDcr, hence did not develop the fact that these are male characters. I do not find any tangible characters which can be applied to both sexes. The type of the genus is Intisteima, and the other related forms are cimhicis, bisetosa, taunts, and parkeri. The femoral bristles are variable in cimhicifi males. While the five species are undoubtedly allied, I doubt if the fact is best ex])ressed by raising tliem to generic rank. Helico])ia was ])roposcd by Coquillett (Proc. Acad. Nat. Sci. Pliil., 1895, 317) for the species with 10 THOMAS SAY FOUNDATION hairy first vein, taking helicis as type. While I have found no case of inconstancy of this character in the North American species, such are said to exist in Europe ; and at the best this character cuts across the others which seem to have subgeneric or even group value, in such a way that it is clearly unavailable for generic separation in Sarcophaga. This has been rec- ognized by several dipterists. A survey of the present status of the Muscoid Diptera indicates unmistabably that our great need is not more genera, but a more complete knowledge of species. The tendency of extreme generic special- ization is to erect about itself impenetrable barriers of names, as is well illustrated by Desvoidy and Brauer. Genera conceived in this mental atmosphere are likely to be almost wholly subjective creations, which make no impression on the minds of others. The argument that the classification must express the relations has its dangers, partly from the subjective elements in- volved, and partly from the fact that the classification has also another important function, — to provide us with names for our species, which should be as stable as possible if they are to have any usefulness. Furth- ermore, to separate a species from the genus in which it has been located expresses one relation, that of dif- ference ; but it obscures another, that of resemblance, which may be more important. One of the main ob- jects of the present work is to make the identification of species as simple and certain as possible, and thus to attract biological and economic workers into the group. This object would inevitably be defeated by the erection of a considerable number of indistinctly separated genera. These are the governing consid- erations which have prompted the preservation as nearly as practicable of the old genus Sarcophaga. Spfcial Taxonomic Characteks. It is assumed that entomologists using the pres- ent paper will be tolerably familiar with terms used in describing Diptera; the papers of Parker and Bottcher, and Williston's widely used Manual of SARCOPHAGA AND ALLIES 11 North American Diptera, seem to render a lengthy discussion unnecessary here. But a few special terms, or special limitations of terms, are explained. Head. — The width of the front is a character of some importance, though subject to a moderate amount of variation. The width is expressed in a decimal, obtained by dividing the number of mi- crometer units in the narrowest width of the front by the number in the entire width of the head. Both of these distances are susceptible of rather exact meas- urement, hence the decimal is significent at least to two places, and has regularly been carried to three, though the variations in different specimens make the third figure of little importance. Where there were specimens enough, five males and five females were regularly measured, and all the separate measure- ments are given for each sex, as well as their average. The term hucca is applied in Dr. Hough's sense (Proc. Acad. Nat. Sci. Phil., 1898, 168) to the lat- eral edge of the mouth cavity, except that I extend the region up to the lower edge of the eye, including the narrow and in this part generally very indistinct transverse impression, which was excluded by Hough. Thus the height of the bucca can be roughly expressed as a fraction of the height of the compound eye, al- though on account of its slanting position it is im- practicable to measure this relation with any pre- cision. On the back of the head, a very distinct suture extends outward and downward from the central foramen, marking off the lowest part, which is the metacephalon; the suture continues as a perceptible groove or line to the edge of the mouth, cutting off the back part of tlie bucca. This line often separates the black and pale hairs of the bucca, the pale being behind the metacephalic suture. A careful study of this detail often aids in distinguishing the females of species. An even more useful aid for tlie purpose is the number of rows of black hairs beliind tlie eye on the outer part of the back of the head. The meta- 12 THOMAS SAY FOUNDATION cephalon is strikingly developed backward in the genus Sarothromyia (Fig. 7a). Tlioraoc. — The chsetotaxy is explained, for con- venience of reference, in connection with figure 14G, in the explanation of the plates. The expression "3-5 black stripes" means that there are three principal, distinct stripes, and outside of these one shorter, less distinct, on each side. Abdomen. — The genitalic characters are ex- plained in the next section. Legs.- — A male character of some importance is the frequent presence of a comb on the lower hind side of the middle femur near its tip ; this consists typically of a row of sliort, flattened, blunt bristles, close to- gether; there may be only five or six bristles of this character. It diminishes through a series of species so gradually that it is hard to say where it ceases to exist. None of the females possess it. JVings. — At the extreme base of the costa are found two scales, one so close to the thorax as to be somewhat difficult to see in many specimens. These are epaulet and subepaulet. The latter, which is the outer, is easily seen, and is always pale; the former is usually quite black, but in the liavinia group it is reddish-yellow, a minute but seemingly good char- acter. The costal segments are numbered, following Bottchcr, so that the first segment is before the hu- meral crossvein, the sixth and last measures the open- ing of tlie first posterior (or apical) cell. The costal spine, when present, is at the tip of the auxiliary vein. The Genitalic Chaiiacteks. One could hardly believe without experience how easy and how positive the identification of species becomes in the Sarcophagida^ by the aid of the male genitalia. The prime object of the present paper is to impress this fact and open a new field for taxo- nomic workers. There is no reason why in the course of a few years there may not be dozens of correctly SAECOPHAGA AND ALLIES 13 identified local collections of Sarcophagidee in the United States, and dozens of workers who are compe- tent to identify material and describe new species. Since the genitalia have this supreme importance for taxonomy in this group, it is necessary to describe their structure before passing to the analytical tables. The abdomen in Sarcophaga has four main seg- ments, beyond which in males are two smaller seg- ments of different appearance, called the hypopyg- ium, or genital segments; the first of these may be somewhat retracted into the fourth segment. The anus is located at the end of the second hypopygial segment. On the two sides of the anus or just below it are attached the forceps, two well chitinized organs lying side by side and usually curving somewhat for- ward below; they are generally visible in large part without special preparation. They do not have lat- eral motion, hence are not like true forceps. Their shape is highly characteristic for each species, and in many of them is very striking (compare figures No. 21, 24, 36, 45, 68, 75). They are the hindmost of the series of organs to be described. Assuming now that the genitalia have been spread ( a process to be de- scribed a little farther on), the next important part is the accessory plate, a more or less triangular sclerite lying just in front of the outer side of the forceps where the free part joins the segment. It offers fewer characters than the forceps, but is often pe- culiarly modified. The typical form is shown in fig- ures 21, 22 and 23; in Fig. 26 it is developed into a fingerlike process, and in 27 it is long and large, with hooks on the posterior side; compare also 9 and 52. Next in front of the accessory plates, and lying nearer the median line, we come to two hooklike or- gans, the posterior and anterior claspers, well sliown in figure 23. Each of these occurs in various forms, long and short, straight and curved, as an inspection of the figures will show. The forceps, accessory plates and claspers are all paired organs, but the remaining one is unpaired, — ■ 14 THOMAS SAY FOUNDATION the penis, lying in the middle line in front of the for- ceps and between the claspers. It is the most complex organ of all, and is of prime importance. A complete homology of its various forms would occupy a volmne and require years of study. It is usually two- jointed, the basal joint being a simple support. The distal segment is in most cases fairly well chitinized, and is essentially a swollen tube bearing various flat plates and slender processes, a further account of which will be found in the specific descriptions. The two segments of the hypopygium are deeply grooved beneath to receive these organs when they are in repose. The penis and claspers are supported on a firm mass of tissue which is attached on each side well forward, at the front lower corner of the second segment. This enables the penis and claspers to move up and down, as a consequence of which they seem sometimes to be longer in comparison with the for- ceps than they seem at other times, or in other speci- mens,— a point to be remembered in comparing speci- mens with each other or with figures. The penis bends forward and back freely, and is figured either way. On the ventral side of the abdomen, anterior to the genitalia, is another structure of importance, the fifth abdominal sternite. It will be apparent on in- spection that there is at the base of the abdomen a short sternite before the one that matches the first visible tergite (indicating that the apparent first ter- gite is a combination of the first and second) ; count- ing from this basal sternite, the fifth lies partly con- cealed within the ends of the (apparent) fourth ter- gite. It is deeply forked behind in the shape of a V or U, and often shows characters of specific import- ance, either brushes of very short spines, or blunt pro- cesses, sinuosities of outline, etc. sarcophaga and alijes 15 Spreading the Genitalia. This indispensable operation involves just about as much effort as spreading the wings of Lepi- doptera, and like it, may be performed either upon fresh specimens or upon dried ones that have been relaxed in a moist chamber. With fresh ma- terial the custom of the writer is to take a dissecting needle with a minute hook or right angle at the tip, hold the specimen under the low power of the binocu- lar dissecting microscope between the thumb and finger, and draw the forceps and inner organs as far back as they will go without tearing the tissues; stretching the nmscles and ligaments in this way a few times, they do not retract the organs so strongly. After this is done, lay the pinned fly with its side against the surface of a piece of cork, by sticking its pin into the edge of a second piece of cork that has been attached to the surface of the first. It is then a simple matter to again draw out the genitalia (under the binocular) and hold them so by sticking pins into the cork. Leaving the specimen to dry, a matter of two or three days, finishes the process. Dry material will need to be placed in the relax- ing jar for 24 hours, longer with very old specimens. Wliere time cannot be allowed for relaxing, boil the specimen in water for a minute and proceed; the wings will be spoiled, but the pollen of the body sur- face generally comes back to its original appearance better than when merely warm water is used. Modifications of the described method will natur- ally suggest themselves. A binocular microscope is not indispensable, though it is a very great advantage. Professor Melander's illuminating apparatus for the binocular (Jour. N. Y. Ent. Soc, xxi, 227, 1913) has been used througliout the present work, and cer- tainly doubles the usefulness of the microscope. 16 thomas say foundation Habits. The species of Sarcopliaga range in larval habit from scavengers to parasites of warm-blooded ani- mals. Many have been reared from excrement of mammals, a few from carrion, many from dead in- sects. Identifications recently made connect Sarcoph- aga hcemorrhoidalis with intestinal myiasis in man, and even suggest that many of the cases may be caused by this widespread species. Several species deposit their larvce in the cups or pitchers of the pitcher plant, Sarracenia, and are not known to have any other larval habit than to devour the decomposing insects that occur in the water of these cups. See discussion of this series under Sar- copliaga sarracenice. There appears to be a very interesting transition from the habit of devouring dead insect to parasitism upon living ones, in regard to which much more study of a critical nature will be required. Some species apparently may be called facultative parasites, others are obligates, being unable to live on dead tissues. To the latter class seem to belong all those in which the female possesses a more or less developed larvi- positor ; but the habit is probably not limited to these species. Without exception as far as known, the species are larviparous. The larvae of several species have been studied by Nathan Banks, who has given (Bull. 22, Technical Series, Bureau of Entomology) many figures showing larval structures in this and allied families, with table for their separation. Almost no specific characters are known in the larvae, so the identification can go no further than "Sarcophaga sp." unless the adult is reared. The minute booklets on the integument which serve for locomotion com- pletely encircle the segments in Sarcophaga, which is the main larval differential character. SARCOPHAGA AND ALLIES 1/ Pantel (La Cellule, xxi, 1 fasc., 105, 1910) states that the larvae of Sarcophaga which are para- sitic in Ortlioptera do not show any specialization for this mode of life, and simply lie among the tissues oi' the host, doing little damage until nearly fullgrown. and obtaining oxygen from the body fluid of the host. Several entomologists liave noted a peculiar larvipositing habit in species attacking grasshoppers, the female fly dashing at the flying host and attach- ing the sticky larva in midair. Evidently several species of Sarcophaga do this. Many rearing records are enumerated in the fol- lowing pages, most of which have been accumidated w^ithin recent years by workers of the Bureau of P^iuto- mology; each year sees a large increase in the number of such records, and it is to be hoped that there will be an increasing carefulness to eliminate the possi- bility that the supposed parasite did in fact larviposit after the death of the host. The general absence of precaution on this point vitiates a large part of exist- mg records, except in Orthoptera, M'hicli the Sarcoph- agas parasitize very freely and commonly. Besides the accumulated records of this paper, attention is directed to a summary by the writer in the Journal of Economic Entomology, viii, 242-24'(), 191.5. ACKXOWI^EDGKMKXTS. In addition to the National Museum collection (inclusive of recent additions from the Bin-eau of En- tomology), the writer has described the material in the University of Chicago (the Hough collection) ; the American Museum of Natural History; the Ujiiver- sity of Kansas, and that belonging to many individual dipterists who are mentioned in the ])roper places. Professor Mine's and Mr. A. H. Sturtevant's collec- tions, and that of Mr. H. S. Ilarbeck. have been espe- cially im])()i-tant in containing undescril)cd s])ecies. Dr. Ralph K. l*arker assisted very materiallv bv 18 THOMAS SAY FOUNDATION identifying several European species occurring in our fauna, and by the donation of paratypes of his species. Summary. One hundred and forty-five species and varieties, belonging to sixteen genera, are described in the pres- ent paper, and the genitalia are figured in 138 cases; other figures bring the total number on the plates to 170. A list of the included species is given in the ex- planation of the plates. The number assigned to a species in the text is in every case the same as the number of the figure or figures illustrating it. In a large number of the species, the female has not been described ; and it is possible that some errors have occurred in identifying females with the wrong male, as but few pairs collected in copula exist in col- lections. 'Not only is there much to be done in studying the females, but there are beyond doubt many spe- cies yet to be described, especially from tropical and subtropical regions. Nothing is ever final in tax- onomy. Only species seen by the writer are included. The numerous old names which cannot be elucidated at present (their types being lost or scattered in Euro- pean museums) are to be found in the Catalogue of 1905, and are not repeated here. It is impossible to follow any other course than to ignore them, or else w^ait many years with the group in liopeless confusion. The latter course seems especially undesirable since a real key to the species has l)een found in the genitalic characters. CLASSIFICATION Table of Genera of Sarcophagidje* 1. Abdomen wholly shining black; arista long-plu- mose (tropics). Phrisso podia Macq. Abdomen with three rows of shining black spots on the second, third and fourth segments; these are sometimes confluent, but at least the intervening angles are densely pollinose ; arista pubescent (widespread north) Wohlfahrtia B and B. Abdomen pollinose, at most the hinder part of the segments shining black in certain lights, tessellated in others 2. 2. Veins 1, 3, and 5 hairy; flrst posterior cell closed and short petiolate (tropical). Jolinsonia Coq. Vein 5 always bare 3. 3. Male with one orbital bristle, proclinate (the uppermost frontal sometimes simulating an- other, but reclinate) 4. Male without orbital 6. 4. Front protuberant; vibrissf\3 approximated and high above oral margin (Louisiana) Camptops n .g. Front not ])rotuberant, vibrissse as usual 5. .5. ^Nletaceplialon greatly developed, lience the lowxr edge of head remarkably long, pro- duced backward (tropical) Sarothromf/ia B. and B. jVIetacephalon not greatly developed (tro})ical) Sarcophagida V. d. AV. ♦Note. — In order to utilize the most snlionl differences, tlie tnble is based to some degree on male cliaracters, altlionnii it is lielieved tliat all the genera are rer'ognizal)le in the female sex. Only genera studied by the writer are included, but notes on others supposed to be allied are given at llic end. 20 THOMAS SAY FOUNDATION 6. Arista sliort-jilimiose 7. Arista loiig-pluiuose 10. 7. Parafaeial witli a single inac-rocliaeta below, pro- cliiiate (Louisiana, Texas) Camptopijga n. g. Parafaeial with a row of hairs. 8. 8. No costal spine (widespread north) A grid Desvoidy. Costal spine strongly developed 1). 9. Back of head with only black hair; anterior acrostichals very large (northeastern) Harhcckia n. g. Back of head with some pale hair; no ant acr (widespread) Hijpopclta n. g. 10. Two large posterior dorsocentrals, two equally large anterior ones (tropical) Notochcvta n. sp. Generally three or four ps dc, if only two the anterior ones are much reduced 11. 11. Vibrissa^ approximated and high above the oral margin (the distance ecpialing the length of the second antennal joint) (widespread) Emblem (ISO ma n. g. Vibrisste normal, or if somewhat approximated they are not strikingly removed from the oral margin 12. 12. Parafacials quite bare; first vein hairy (Louis- iana) Sthcnopi/ga n. g. Parafacials M'ith hairs, sometimes delicate. 1.3. 13. Ocellars absent; frontal rows not noticeably di- vergent below% reaching base of antennae (tropical) Harpagop/jga n. g. Ocellai's present; if absent, the frontal rows are abruptly divergent below and reach a])out to the middle of second antennal joint 14. 14. Fifth sternite of male undivided; epaulet yel- low; back of head with only black hair (northeastern) Thclodlscus n. g. Fifth sternite of male dee])ly divided (wide- spread) Sarcophaga Meig. sakcorhaga and allies 21 Notes ox Other Gexera Microchcctina Van der Wulp, I5iologia, Dipt., ii, 240, 1891, is a Dexiid, as stated by the describer. Braiier and von Bergenstamni placed it with their SarcophagidcV, and their arrangement was followed by the writer in his 1905 Catalogue. Adams, however, in Williston's Manual, 1908, properly replaced the genus in DexiicUe. Paramintlio Van der Wul]) (ibid., 20.5) is also a Dexiid, from its very small antenna% and discal nia- crocha?ta3 on segments 2-4 of the abdomen. Onesia Desv. This Eiuro])ean genus was reported from Mexico City by Van der Wulp (O. luc'dioidcs, ibid., 288, pi. vii, f. 11). The genus is close to Cyn- omyia, and from its bright metallic green coloration is readily distinguished from the group treated in the present paper. Ilespcromjfid 15rauer und von Bergenstamnu Zweifl, Kais. INIus., iv, 114, 1889; v, 3.59, 1891 ; vi, 1(52, 1893. One species, eri/fhroccra, from Texas (ibid.). The genus is allied to Senotainia, but has the first posterior cell closed and petiolate; it does not come within the scope of the present paper. The arista is almost bare. Erifthrandrd l^rauer und von Bergenstamm, ibid., V, 3(J8, 1891; vi, 10.5, 1893; one species, piclpcs, from Georgirt, U. S. A. The principal characters given are as follows: Parafacials with only very deli- cate hairs; arista pubescent, or at base short-plumose; no costal spine; first ]K)sterior cell open; male witlioul orbitals; third antennal joint 2-2l/^) times the second; ])ucca wide: fVont of male very narrow; vi})riss{u a little ap])r()\imated. From the data given, I am un- a])le to distinguish the sj^ecies, but it would prol)al)ly fall within the genus Sai'co|)haga as here accepted. 22 THOMAS Sx\Y FOUNDATION Genus PHRISSOPODIA Macq. Macquart, Nist. Nat. Dipt., ii, 222, 1835. Type designated as Peckia imperialis Desvoidy, which is stated to be without doubt the same as Sarcophaga praeceps Wied. Macquart, Dipteres Exotiques, ii, pt. iii, 253 (sep. 96), spelled Phrissopoda and Phryssopoda. Bauer und von Bergenstamm, Zweifl. Kais. Mus. iv, 1889 Phrissopoda); vi, 163, 1893 (id.). This genus has by several writers been regarded as a synonym of Peckia Desvoidy (Myodaires, 1830, 335 ) . The latter contained five species, no type be- ing designated; only one {imperialis) had the shining- black abdomen, the others apparently not differing from Sarcophaga as taken in the present work. Mac- quart was entirely within his rights in isolating ivi- perialis in a new genus, and the only defect in his pro- cedure as viewed by the most critical modern was that he failed to designate a type for Peckia from among the four species remaining in it. But it was not the fashion in those days for an entomologist to designate a type for another man's genus, if indeed he did so ■for his own. Macquart's genus Pln-issopodia was a valid genus at the time ; it contained besides imperialis ( ^praeceps) one other species evidently congeneric in the modern sense, also with a shining black abdo- men. Brauer and Bergenstamm in 1889 gave a rather full description of j^f'^iecejJS Wied., apparently from the types. Generic characters: Very large, robust species, with shining black abdomen and the male having on the middle and hind legs very abundant, bushy vil- losity, both on femora and tibiae. Head as in Sarcophaga; front narrowed in male, without ocellars or orbitals (B. B. however mention two pairs of vestigial orbitals in the male) ; third an- tennal joint more than three times the second in the male, nearly three times in female; epistoma some- what produced forward and downward in the male, where the vibriss^e are a little above the oral magrin; facialia hairy halfway up. SARCOPHAGA AND ALLIES 23 Thoracic cha^totaxy (from specimen) ; ps dc only two pairs, far back; ant dc 0; ant acr 0; prsc 1 me- dium; hum 3 (one is small) ; phum 1 or 2 small; prs 1 ; npl 2 large ; sa 2 ; inta 1 ; pa 2 ; stpl 2 with some large hairs; scutellum with 2 marginal and a stout, some- what smaller pair of decussate apicals, no discal or subapical. Abdomen shining black; hypopygium large, of similar type to Sarcophagas (to the tropical group w^ithout ocellars, including liillifera, etc. ) . No. 1. Phrissopodia praeceps Wied. Wiedemann, Auss. Zweifl. Ins., ii, 355, 1830 {Sarcoph- aga) . — Cuba. Desvoidy, Myodaires, 335, 1830 {Pcckia iinperialis). — Cuba [Macq.] Macquart, Hist. Nat. Dipt., ii, 223, desc. and probable syn. , Dipt. Exot., ii, pt. 3, 252, note. Walker, Trans. Ent. Soc. Lond., n. ser., v. 310 (Sarcoph- aga fortipes). — Hayti. [G.-T.] Brauer und von Bergenstamm, Zweifl. Kais. Mus., iv, 125, 1889; vi, 163, 1893, type redesc. — Cuba and Chile. Williston, Trans. Amer. Ent. Soc, xiii, 307, 1886, oc. in San Domingo {Pcckia). Giglio-Tos, Ditt. del Mess., iii, 67, 1894, syn. and oc. — Mexico. Williston, Manual of N. A. Dipt., 1908, 350, half-tone illustration {Pcckia). [F. H. Snow, Kans. Univ. Sci. Bull., ii, 345, 1904, re- ported the species from Arizona, but on examining his material I found it to be Trixodes obcsa Coq.] Male. Front .200 of head; parafrontals and parafacials white pollinose, not very glistening, the latter with an irregular row of fine but long hairs be- low near the eye ; frontal stripe blackisli, the anterior end twice as wide as near the ocelli; frontal bristles about 13, the lowest three ])airs abruptly diverging, and reaching almost to the middle of the second anten- nal joint; antennas blackish-brown, third joint slen- der, reaching five-sixths of the way to the vibrissit; arista with rather erect and long plumosity for three- fourths its length; palpi and proboscis black, ordin- 24 THOMAS SAY FOUXDATIOX ary; bucca one-half the eyeheight, with long, soft, mostly ])a\e hairs; back of head with two or three ir- regular rows of black hairs, the rest pale, abundant and conspicuous; outer vertical wanting. Thorax brownish-black, with 4-6 gray pollinose stripes, of which the subniedian pair are divided be- yond the suture; scutelluni uniform brownish-black, convex; dorsum with tine erect black hair, longer on the humeri, mesopleura, sternopleura, postalar callosi- ties, and sides of scutelliun ; calypters dark brown. Abdomen shining black; first and second seg- ments with only lateral bristles; third with a stout median marginal pair; fourth with a strong, irregular marginal row of some 20; fifth sternite blackish, ex- cised U-shape, the arms with some inconspicuous black hairs but not much exposed. Hypopygium large; first segment shining black, with fine erect hair and no bristles; second segment dark red, the sides brownish, hair erect and soft, no bristles ; forceps longj slender, straight, shining black, the sharp tips curving gently forward; from behind they are well separated, slender, parallel. Accessory plate shining black, prominent and rather slender, W'ith almost sharp tip, the hind edge thin and notched near tip; posterior clasper shining black, erect, long, gently curved, with small but distinct apical hook; anterior clasper robust, red at ])ase but shining black apically, strongly curved forward, the tip obliquely truncate to a second shorter point below. Penis red, the division between the basal and distal segments not very distinct; the back plate is deeply cleft at apex into two side plates, which have a minute tooth at tip and a long, curved one forming the anterior apical angle ; below this, on the front of the segment, are two lobes bearing pale hair, and between them is ^isilyle the l)lunt tip of a chitinous organ. Legs black; anterior tibia? without bristles except at tip, and on its front side for nearly the whole lengtli with a delicate brush of reddisli hairs; middle tibia longer than its femur and a little curved, with one SARCOPHAGA AND ALLIES 2.3 bristle on the outer front side, tlie villosity extends ujDon the hasitarsus; hind tibia with only two bristles before the tip ; hind femur and tibia botli swollen and curved away from each other in the middle ; claws and pulvilli all long. Wings l)rownish-gray, the base strongly black- ened; no costal spine; third costal segment twice the fifth and sixth together; fifth hardly twice tlie sixth; first vein bare, third with a few hairs; fourth vein bent in an acute angle, barely appendiculate. Length 22 mm. One male, Santiago, Cuba, Oct. 2.5. 19().*J (Capt. W. Kobinson), in tlie National Museum. Brauer and von Bergenstamm state tliat in the female the arista is short-plumose, and for only half its length; as they have added Chile to the type local- ity, I suspect that they have the M^rong female, from that country. Type. — In the Imperial Museunu Vienna. Genus WOHLFAHRTIA B. and B. Brauer und \on Bergenstamm, ZweiH. Ivais. Mus.. iv, 123, 1889; vi, 1C5, 1893; X'erhandl. Zool.-Bot. Ges., xliii, 1893, 501. Portschinsky, Hor?e Ent. Soc. Ross., ix, 1-180, 1875, 3 pi. and figs. (Sarcoplilia), biology of jinu/iiifica. the type species, etc. ; ibid., xviii, 1884, 1-69, man\- figs., mono- graph on the same, 1884. All in Russian. Osten vSacken, Berl. F,nt. Zeitsch., xxxi, 1887. 17. resume of Portschinsky's papers in luiglish : ]>. 18. habits of magnifica. Coquillett, Jour. N. Y. V.ut. Soc, iii, ]05. 1895 (Para- pliyfo); Rev. Tachin., 1897, 122 (id.). | bihnson. P.syche, xix, 103, 1912.I This genus was se])arated from Sarco])lii]a by its authors on characters which seemed very siiglit excejjt as to the arista, whicli was said to })e })ubcscent in Wohli'alirtia. short-j)ilose in Sarcophila. I doubted the validity of the genus until (juite recently (and lience referred opaca to Sarco])liila in Knt. News, xxiv, 21.5. 19i;3) ; Mr. Coquillett in 11)10 (Type Spe- 26 THOMAS SAY FOUNDATION cies, Proc. U. S. N. M., 620) held the same opmion. After examining a pair of the European Sarcophila latifrons in Professor Melander's collection, I see that Wohlfahrtia is very distinct from that genus, in which the hucca is narrow, the front not protuberant, and the head of entirely different shape, not to speak of other differences. Brauer and von Bergenstanmi included two spe- cies in the genus, Sarcophila magnifica and ineigeiiii of Schiner; in the third paper mentioned above, they named the former as type. In the generic descrip- tion, they attribute two pairs of orbital bristles to the male in both species ; in one male of magnifica in the Hough collection there is only a single orbital pair, and in two males of mei genii (all these are European, presumably named by Strobl) there are none, as in our North American species. As Professor Bezzi has pronounced one of our species identical with meigenii, it is evident that an error was made in the original generic description. Generic characters: Front and epistoma pro- duced, giving the head a rather square outline in pro- file; front wide in both sexes, but more so in the fe- male; third antennal joint not much longer than the second (except in the European W. maxima) ; arista pubescent; bucca high; parafacial wuth a few small hairs not in rows; ocellars and outer verticals present in both sexes ; back of head with only black hairs. Thoracic chaetotaxy {magnifica, from specimen in the Hough collection) : ps dc 4; ant dc 3-4; ant acr about 4 pairs present; post acr 2 pairs large in male, one in female; hum 3; posthum 2; prs 1 and an inner very small; stpl 2 and a small below the front one; scutellum with 4 marginals, the fourth pair close to- gether, no small apicals between them; subapicals, 2 rather large on one side and one on the other in the specimen; pa 2; sa 2. Abdomen densely, uniformly pollinose, with per- manent black shining or subshining spots in three longitudinal rows. SARCOPHAGA AND ALLIES 27 Wing as in Sarcopliaga; third vein hairy; third costal segment longer than fifth and sixth together. Hypopygimn inconspicnous, turned under the abdomen. The type species is in habit, according to Osten Sacken, the European analogue of our North Amer- ican Screw- Worm Fly, Chri/somyia macellaria F., its larva frequently getting into the sores of animals and not rarely attacking man, in Eastern Europe. Table of Species. — Males. Western species, the abdomen mostly pollinose, its black spots small No. 2. mei genii Schiner. Eastern species, the black spots large and con- fluent, the pollinose paTt not very conspicu- ous No. 3. vigil Walk. Several other species assigned to Paraphyto by Coquillett do not belong to Wohlfahrtia; the types appeared to the writer to belong to Dexiid genera, hence are not further considered here. But the type of Paraphyto undoubtedly belongs to Wohlfahrtia. No. 2. Wohlfahrtia ineigenii Schiner. Schiner, Fauna Austriaca, Dipt., i, 567 (SarcopJiila). — Europe. Meigen, Syst. Beschr., v, 17, pi. xliii, f. 9, female (as Sar- cophaga ruralis Fall.). [Schiner.] Portschinsky, Horse Soc. Ent. Ross., xviii, pi. ii (Sar- cophila). Megnin, ibid., xv, v (SarcopJiila ruralis). Brauer und von Bergenstamm, Zweifl. Kais. Miis.. iv, 123; vi, 165. Villeneiive, Bull. Soc. Ent. France, 1900, 363 (ruralis). Coquillett, Revis. Tachin., 1897, 122 (Paraphyto opaca).- — Colo, and New Mexico. Johnson, Psyche, xix, 103, 1912, notes; probable syn. of opaca. — Colorado and Utah. F. H. Snow, Kans. Univ. Sci. Bull., ii, 344, oc. in Arizona (Paraphyto opaca). Aldrich, Ent. News, xxiv, 215, oc. on flowers at Brigham. Utah (SarcopJiila opaca). 28 THOMAS SAY FOUNDATION The synonymy of Parapht/fo opaca rests upon the authority of Professor Murio Bezzi, of Turin, Italy, to whom North American specimens were sent by the writer; he reported them to he identical with Wohlfahrtia meigcnii (letter, June, 1915). Male. Front .830 of head (average of five, — .319, .321, .326, .337, .349); parafrontals and para- facials broad, silvery pollinose, glistening; frontal stripe broad, its upper part glistening and hardly dis- tinguishable from the parafrontal; frontal bristles not diverging below, reaching the base of the second an- tennal joint; transverse impression and middle of face reddish; antenna^ red, third joint blackish apically, not much longer than the second, and reaching more than two-thirds of the way to the vibrisstc, which are at the oral margin; palpi yellow; bucca about four-fifths the eyeheight. Thorax with thin gray pollen and three ill-de- fined black stripes; chaetotaxy as in magnifica (see generic description) except in the following points: ps dc 3, but room for another behind the suture; 1 pair prsc ; prs 1 ; posthum 1 or 2 ; sc with 1 pair subapical, which as in magnifica nn'ght be considered discal. Abdomen with three rows of shining black spots on densely pollinose ground; the pollen extends only to tlie edge, leaving the venter shining; first and sec- ond segments with only lateral bristles; third and fourth with an interrupted marginal row; fifth ster- nite not divided, yellow, at the middle of the hind edge a slight protuberance, Hypopygium black; first segment pollinose at base, shining at apex, with a row of bristles near base; second segment shining, globose, not bristly. Forceps black, nearly straight in profile ; from behind they are well separated })ut not divergent; accessory plate blackish, elongated, with a wide, bare, rounded free end farther forward tlian usual, beside the penis; posterior clasper yellow, very short and low, bent back but the ti]) curved forward, ordinarily cpiite invisible; anterior clasper yellow, erect and cur\'ed forward, the SAKCOPHAGA AND ALLIES 29 blunt tips ordinarily lying between tlie arms of the penis like a part of that organ; the claspers and penis seem to coalesce at base, and all lie deep in the groove of the under side of the hypopygium. Penis with but one segment, yellow, simple, much like that of Sar- cophaga communis'; it has a pair of conspicuous straight i)lack anterior plates. Legs l)lack, middle fenun- with comb; middle tibia with two bi-istles on the outer front side, and with long villosity in the inner, which is strikingly long at the very tip; hind tibia also with villosity, but not so long at the tip; botli of these tibi;c are curved, and the hind femur is thickened and curved. Female. Front .413 of head (average of three, — .405, .412, .421) ; Genital segment brown and shin ing above, the orifice transverse. Femora and ti})iae simple, but the hind femora are a little curved. Length TV^, to 14 mm. Five males and three females: four males and three females were taken at Moscow, Idaho, by the writer, some on flowers of goldenrod and parsnips, at dates from Jime 24 to Sept. 4. The other male is from Koehler, Xew Mexico (W. K. Walton). Coquillett's ty})e of Paraplnjto opdCd is in the Xational Muscuul and was examined by tlic writer. Type. — In the Imperial Museum. Vienna. No. 3. Wohlfahrtia vigil Walk. Walker, List of Dipterous Insects in the l>ritish Museum, iv, 831 (Sarco/^haga). — Nova Scotia. Coquillett, jour. N. Y. Rnt. Soc, iii, 105 (Farapliyto chittendeni) ; Revis. Tachin., 122 (id.).^ — N. V. ; Michigan. My attention was called to Walker's SarrojjJiaga vigil l)y C\ W. Jolmson, who had identified it; as I had just been studying tlic ty])e of ParapJuflo cJiii- tendcni, it was easy to detect the synonymy. Walker's description a])])lies \'ei-v well, and the s])ecies is so unicjue in coloi-ation as to leave little doubt of its cor- rect identification. 30 THOMAS SAY FOUNDATION It differs from mcigenii in practically nothing but the more broadly expanded shining black spots ol' the abdomen, which coalesce wholly or partly. The figure shows the extreme of coalescence, but even the minimum that I have seen looks very different from the western form. One male has the fourth abdominal segment and hypopygium mostly red. The number of ps dc evidently varies from three to four, but the fourth (just behind the suture) is small if present. The front measures in the males .333, .350, and .353 of the head (average .345, slightly more than in mei- geiiii), and in the females .417 and .424 (average .420). Three males and two females : two males and one female are from LudlowA^ille, New York, in the Car- negie Museum; one male and one female Rochester Junction, N. Y., collected by M. D. Leonard. Type. — Male, in British Museum. Vigil antedates mcigenii by thirteen years (1849, 18(32), and would replace the latter if it were ever shown that they are one species. The distinct geo- graphical boundaries, and the apparent absence of the vigil form in Europe, added to the obvious color char- acters, seem to assure the distinctness of the species. Genus JOHNSONIA Coq. Coquillett, Proc. Acad. Nat. Sciences Phil , 1S95, 316, fe- male. Head rather rounded, front broad in both sexes, with one proclinate orbital bristle in each; frontals only four or five, strong and somewhat reclinate, the rows not diverging below and reaching hardly to the base of the antenn oc. at Lake Worth, Fla. ; Psyche, xv, 1908, 76, oc. in Brahamas, Jamaica, and Florida ; Bull. Amer. Mus. Nat. Hist., xxxii, 75, oc. at Lake Worth, Miami, and St. Augus- tine, Fla. Male. Front .352 of head (average of two, — .344 and 3.59) ; head cinereous pollinose, not silvery; antenuce black, third joint twice the second and reach- ing four-fifths of the way to the a ibrissiu, which are very stout, slightly approximated, and almost at oral margin; palpi long, somewhat clavate; proboscis black, rather short, with small labella; bucca one- fourth the eyeheight; back of head with several rows of black hairs, none pale except a few below the neck. Tliorax with dense, rather uniform pollen of whitish-gray color, the stripes indistinct even when viewed from behind, bristles large. Abdomen concolorous, with black median stripe and a changeable one each side; second segment with long, appressed black hairs on hind edge; third with ]'ow of 8 bristles mixed with a few of the same hairs; fourth with row of about 16. Hypopygium black, gray pollinose, strikingly similar to that of Sarcophagula occidua. First seg- ment w^ith row of four small bristles before hind margin; second with two large and several small bristles; forceps black, slender and tapering, straight; accessory plate very large, suddenly narrowing to a distal prolongation wliich is wider, shorter, and more 38 THOMAS SAY FOUNDATIOX bent forward than in occidua; claspers as in that spe- cies ; jDenis with the tubular central part more swollen and shorter, and the terminal process much larger. Fifth sternite only showing on the middle, apparent- ly undivided, with black short brush. Legs black, femora rather stout; middle femora without combs, middle tibia with an irregular row of bristles on outer front side, one of which is large; hind tibiae without villositv. Front tibiae and tarsi with peculiar erect hairs forming a fringe on the upper and outer side, the tarsi a little swollen and the segments somewhat shortened, the claws and pulvilli shorter tlian in the other tarsi. Wing hyaline; veins yellow toward base, third with hairs almost to small crossv^ein; a small costal spine; third costal segment half as long as fifth. Female. Front .369 of head (average of two, — .361 and .376). Genital segment black, retracted. Front tarsi normal. Length, 5 to 6 mm. Two males, two females, Nassau, Bahamas, Dec. 11 and 14, 1912 (F. Knab), from the National Mu- seum. Type. — In the Imperial Museum, Vienna. The females appear to be indistinguishable from those of the following variety; they are merely asso- ciated from being collected together. As Schiner descrilied only females originally, tlie identification depends on Brauer and a^ou Bergenstamm, who de- scrilied the fringes of the male front tarsi. Schiner's description of the pollen as yellowish in color would make the identification very doubtful if Brauer and von Bergenstamm had not had his types for com- parison. Possibly Schiner was misled bv the wide front and orbital bristles, and overlooked some males in his material. SAKCOPHAGA AND AI.IJES 39 No. 8. Sarothromyia femoralis var. simplex n. var. Differs from the typical form only in having plain front tibicV? and tarsi in the male. The material examined shows no intermediate forms. I cannot separate the females, except by associating them with males taken at the same time or place. Four males, three females, Miami, Florida, Nov. 8 (C. H. T. Townsend), in the National Mnsemn; one female. Ft. Worth, Fla., collected by Mrs. Slos- son, probably belongs here. Holotype.— Male, No. 20,-t88, U. S. N. M. Allotype.— Female, No. 20,488, U. S. N. M. Genus SARCOPHAGULA Van der Wulp. Van der Wulp, Tijdsch. v. Ent., xxx, 173, 1887; Biologia, Dipt., ii, 289, 1896. Brauer und von Bergenstamm, Zweifl. d. Kais. Mus., vi. 164, 1893. Front wide, about one-third the head in both sexes, the middle stripe as wide as one side. Frontals reduced to fom* pairs, not divergent, the anterior a little below tlie base of the antenna\ One large or- bital present in both sexes, the posterior frontal might be considered a reclinate orbital, as it is far from the next frontal and much closer to the orbit. Outer vertical present in ])oth sexes. Parafacial narrow, with a single row of minute liairs. Antennje as in Sarcophaga, but tlie third joint only twice the sec- ond; arista long-plumose on basal half, tlie last joint much tliickencd at its base. Vibrissjc not approx- imated, almost at oral margin. Fourtli vein with rounded, oblitpie angle, the first posterior cell open a little ])efore the a])cx; third costal segment very short; hind crossvein rather erect, usually at a})oiit the middle between the anterior crossvein and the bend. Type of genus, SarcopJiac/a occ'tdua Fabricius, by designation of Cofjuillctt, Pi-oc. V. S. N. M., xxxvii. (102, 1 <)!(). 40 THOMAS SxVY FOUNDATION No. 9. Sarcophagula occidua Fabr. Fabricius, Ent. Syst , iv. 315, 1794; Syst. Antl, 288, 1805 (both Musca). — West Indies. Wiedemann, Auss. Zweifl. Ins., ii, 368, 1830 (Sarcoph- aga), type redescribed. Van der Wulp, Biologia, Dipt., ii, 289, pi. vii, f. 12, 1896 {imhecilla, canuta and tenuis). — Mexico. Williston, Trans. Ent. Soc. Lond., 1896, 361, pi. xii, f. 120 bis (Sarcophilodes puclla). — St. Vincent, W. I. Coquillett, Proc. U. S. N. M., xxii, 254 (Sarcophaga), occurrence in Porto Rico. A very small gray tropical species. Male. Front .358 of head (average of five,— .354, .354, .355, .359, .367) ; parafrontals and para- facials gray, not silvery ; frontal stripe red anteriorly ; antennae black, tip of second joint and base of third reddish, the third reaches three-fonrths of the way to the vibrissce; palpi and proboscis black, ordinary; bucca about one-sixth the eyeheight. Thorax yellowish gray pollinose, with three black stripes when viewed from behind; ps dc 3; ant acr 0; presc very small; postalar 2, the outer very small ; intra-alars 2 ; supra-alars 1 large and two min- ute ; hum 2 ; intrahum 1 ; presutural 1 large ; stpl 2 ; npl 2; scutellum with 2 marginals and 1 discal, no apical s. Abdomen black with changeable yellowish-gray pollen, which shows three shifting black stripes; first segment with only marginal bristles; second with some long depressed hairs along tlie margin, the mid- dle pair of which are usually more erect and have the size of small bristles; third with a row of 6; fourth with a row of ten. Hypopygium black, opaque ; first segment large, declivous, with a row of four bristles near tip ; second segment bent forward, with two large ascending and divergent bristles behind; forceps black, straight and tapering to a slender point, not hooked nor divergent ; accessory plate brown, very large and wide, with a long slender tip which has several distinct barbs on the front side; posterior clasper brown, with slender SARCOPHAGA AXD ALLIES 41 hooked tip and a long hair on the front side ; anterior clasper brown, moderately slender and hooked; penis with basal joint indistinct, the distal rather tubular, swollen and black in the middle, with a tongue-like process protruding from the end, and a brown ex- pansion attached to the front side. Fifth sternite showing very slightly. Legs black; middle femora without comb; mid- dle tibia with two bristles on the outer front side, one of them large; hind tibia of male not villous; claws and pulvilli all short. Wings hyaline; costal spine small; third vein with rather long hairs nearly to crossvein ; tliird costal segment hardly half the fifth. Female. Front .382 of head (average of five, — .360, .369, .391, .394, .397). Scutellum with a small marginal ]3air between the two large ones. Genital segment black, much retracted. Length 2.8 to 4,4 mm. Nmnerous specimens of both sexes: 56 ai-e from Havana, Cuba, (C. F. Baker) ; 5 Key West, Fla., Jan. 1, 1869 (C. V. Riley) ; 2 Porto Rico (Busck), determined by Coquillett as occidua; 1 Aguada Pas, Cuba (Sturtevant) ; 2 Panama, P. R. ; 1 Jamacia (Hough) ; 1 St. Augustine and 1 Lake Worth, Fla. (Hgh.); 1 Miami, Fla. The synonymy of Van der Wulp and Williston is from their descriptions only. Type. — Probably in the Imperial Museum, Vi- enna, if still in existence. CAMPTOPYGA New Genus. {Ka/jLTTTos, curved; Twyq, the hinder end) Rather bare, l)lack, medium sized species, with large, bulky hypopygium; the head rather convex behind, vibrissa* at oral margin and almost in the lower line of the head; face vertical; frontals only six in a row, but reaching to middle of second anten- nal joint, tlie last strongly diverging; one macro- 42 THOMAS SAY FOUXDATIOX cliceta at lower edge of parafacial next to eye, just above the facial impression; above this bristle only a few very small hairs; antenna^ nearly as long as the face; arista long and slender, the penultimate joint as long as wide, the last joint somewhat thickened at base, M^th very short plumosity for hardly half its length, the hairs but little longer than the greatest thickness of the arista; palpi normal; proboscis short, fleshy; eyes bare. Thorax with rather few and strong bristles, nearly bare otherwise, the usual small hairs few and scattered ; the scutellum has but four bristles, — two pairs of marginals. Type. — Camptopi/ga aristata new species. No. 10. Camptopyga aristata n. sp. Male. Front of considerable width, .260 of the head (average of three, — .252, .257, .271) ; the stripe velvety black, wider than either side; face and front with plumbeous pollen, not glistening when viewed from below; antenna' black, third joint one and two- tliirds times the second, reaching over five-sixths of the way to the vibrissa; facial ridges bare; bucca one- fifth the eyeheight, a little bristly along mouth; palpi and proboscis black; back of head with three rows of black hairs and not much pale beard; outer vertical bristles large, more than half as long as the inner. Thorax gray, thinly pollinose, three black stripes on dorsum; 3 ps dc; 3 stpl; no ant acr; presc very weak; 1 prothoracic; 1 on lower anterior corner of mesopleura; 3 humeral; 2 ant dc (one small). Abdomen blackish, club-shaped; when viewed from l)ehind the second, third and fourth segments each have a row of four silvery spots lying against the preceding segment, very distinctive; first and second segments with only a single long lateral ])ristle each side; third with a very large pair of median mar- ginals, and two lateral each side; fourth witli mar- ginal row of 12. Hypo])ygium large; first segment brown-black, with row of 6 stout bristles across hind part; second SAKCOPHAGA AND AIJJES 43 browii-blat'k, ^'e^y hirge, with four large and some smaller bristles directetl baekward and curving up- ward; fifth sternite broad and conspicuous, black, with only a linear fissure, forming a sort of hood for the genitalia in repose; forceps black, smallish, e^'en- ly incurved and tapering in profile, the tips not di- vergent ; hind claspers long and slender, curved ; front claspers heavier, darker and more strongly curved; penis blackish, slender nearly to the tip, which is bent squarely forward and has several small anterior pro- cesses. Legs very l)lack; middle femur without combs; middle tibia with an increasing row of three bristles on outer front side; hind tibia not villous. Wings yellowish; costal spine very long; third costal segment shorter than the fifth; fourth vein with square bend, either slightly rounded or pro- vided with a slight ap2)endix, ending at a distance be- fore the extreme apex which rather exceeds one half the fifth costal segment; third vein with stout hairs to the crossvein; first vein bare. Female. Front .331 of head, with two normal orbitals; second abdominal segment with a pair of erect median marginals; otherwise exceedingly like the male throughout ; genital segment black, retract- ed, with transverse apertin*e. Length of male (>, of female .5 mm. Five males and one female: one of each sex from Tifton, Ga., Oct. (3, 189(> (Hough coll.); one male from Opelousa, I^a., June, 1897 (ITough coll.) ; one male Alta Loma, Texas, Dec. 29, 1914 (Bisliopp, No. 3.521) ; one male Anglesea, N. .1. (C. AV. Joliii- son) ; the other male Slidell, La., July 2-6, 190.5 (Prof. Jas. S. Iline). Ilolotypc—Male, No. 20,489, V. S. N. M., from Alta lioma, Tex. Allotype.— Feiuale, No. 20489, U. S. N. IM. 44 THOMAS SAY FOUNDATION Genus AGRIA Robineau Desvoidy. Robineau-Desvoidj^ Myodaires, 1830, p. 376, established the genus with the following definition: "Antennis paulisper incrassatis, cheto villoso; abdo- men punctatum." He explained that he uses the term villoso as meaning subplumose. He described six species, all but one of which are regarded as syno- nyms of Musca aijinis Fallen by Bezzi in vol. iii of the Palaearctic Catalogue; the sixth he recognizes as a Sarcophaga; a finis thus becomes the type of the genus, and is so designated by Coquillett, Type-Species, 1910. Macquart, Hist. Nat. Dipt., ii, 229, 1835, recognized af/inis as belonging to this genus and as being the prior name for at least one of DesYoidy's species ; he howcYcr included a number of diverse forms which later caused some confusion in restricting the genus. Affinis occurs in Xorth America, having been identified for Dr. Parker by Dr. Bottcher, and for me by Parker. Its principal generic characters are as follows: Front in male narrowed in front of the ocelli, in female rather wide, in both with the median stripe wider than the side ; in the female the parafrontal has numerous rather large hairs, continuous with those of the parafacial, which are much the same as in Sar- cophaga. Frontals about 12, widening gradually be- low, reaching nearly to middle of second antenna 1 joint. Antennae reaching nearly to oral margin, the arista with short and rather dense plumosity extend- ing about half its length, vibrissa at oral margin; eyes bare; palpi normal, proboscis short and fleshy; occiput without any pale beard. Fourth vein at bend with a square angle and a slight fold; its end is as far before the apex as half the length of the fifth costal segment. Hypopygium large, mostl}^ occupied witli the first segment, which has a pronounced hump be- fore its middle, forming the apex of the 'abdomen in profile; a row of bristles before the hump; second segment smiall. SAKCOPHAGA AND ALLIES 45 No. 11. Agria affinis Fall. Fallen, Acta Holmiae, 1816, 230; Dipt. Suec. Miiscides, 1820, 39 (both Musca). Pandelle, Rev. Ent., xv, 173 (Sarcophila biclitnis), 1896. Bezzi, Pal. Kat., iii, 469, bibliography, etc., 1907. Male. Front narrowed below ocelli, .124 of head (average of four, — .117, .120, .125, .132) ; face and front silvery, liairs of parafacials distinct but varying in number; antennae blackish, third joint twice the second, reaching four-fifths of the way to the vi- brissse; bucca nearly one-third the eyeheight; outer vertical not differentiated. Thorax blackish, but Avlien viewed from behind gray with three black stripes, the middle of which extends to the end of the scutellum; 3 ps dc; 3 stpi; ant acr 2 pairs, large; prsc rather large; hum 3; npl 2; scutellum with three marginal pairs and a smaller discal, no apical (or the inner marginal might be re- garded as a very large apical pair). Abdomen rather silvery pollinose; viewed from behind there is a median black stripe and a large l)lack spot each side on segments two, three and four ; first and second segments with only lateral small bristles; third with a small median marginal pair; fourth with marginal row of about 18. Hypopygium black; segments as described above; forceps black, ^ery short and blunt; accessory ])lates developed into a large yellow forceps-like or- gan on each side, as long as tlie true forceps ;posterior claspers yellow, very low in position; anterior ones upriglit, with a hook on the posterior distal angle, the anterior point not much curved; penis mostly black, very thick and short, with a snout-like process in front turning down toward the base ahnost to the bodj^ of the liypopygial segment. Legs deep ]))ack; middle femora witli coml) at tip below on hind side; midcHc ti})ia witli two smallish ])ristles on outer front side: liind tibia» not villous; front basitarsus witli some l)ristles below at base. 46 THOMAS SAY FOUNDATION Wings hyaline; hind crossvein nearly parallel with wing-margin ; first vein bare, third bristly almost to crossvein; no costal spine; third and fifth costal segments equal. Female. Front .308 of head, the middle stripe three times as wide as one side, two orbitals as usual, outer vertical well developed; scutellum bristled as in male; genital segment black, with a dense row of small In-istles at edge, showing another complete end of a segment Avithin, and another within this of which the sternite is modified into a short, blunt larviposi- tor; the first sternite of the abdomen is fidly as long as the first tergite. Length, 7-8 mm. Seven males, one female: one pair North ]Mt., Pa., June 7 and Sept. 1 (no collector) ; two males, North Saugus, Mass., May 18, 1911 (II. E. Smith) : one male, Las Vegas Mts., N. M., 11,000 ft., June, 1901 (Cockerell) ; two males, A-^ictoria, B. C, bred from larvae of Vanessa antiopa by J. 1{. Anderson, in the Canadian National Collection; one male, White Mts. [N. II.] (Morrison), in the C. V. Riley collection, U. S. National Museum. Type.— Probably in the Academy of Sciences in Stockholm. HARBECKIA New Genus. The principal generic characters are the pro- tuberant front, sliort fronto-orl)ital rows, rather strong parafacial bristles in a long diagonal row. vibrissge at oral margin but mucli above lower line of head, wide bucca, strong acrosticlials, and the four equally-spaced and equally large bristles on the posterior margin of the scutellum in botli sexes. The hypopygium of the male is Sarcophagid in type, while the protuberant front and strong de- velopment of the bristles suggest a Dexiid relation- ship. Against the latter on the other hand might be SARCOPHAGA AXD ALIJES iT urged the small niuiiber of bristles on the seutelluin, and the partial plumosity of the arista. Xanied in honor of H. S. Harbeck, the Phila- (lelpliia dipterist, who placed at my disposal for the present work the riches of his private collection. Type. — Harhcckia tessellata, n. sp.; according to Mr. W. R. Walton, who looked nj) the matter for me, Brachifcoma pubiconm Coq. (Proc. U. S. X. M., XXV, 116) is congeneric. It differs chiefly in not pos- sessing median marginals on the second abdominal segment. No. 12. Karbeckia tessellata n. sp. Male. Head rather convex behind, front prom- inent, face vertical, lower line of head sloping up con- siderablj^ to meet it. Front narrow, .114 of head (average of two, — .09.5 and .132) ; middle stripe brown, much wider above than parafrontals, the lat- ter and the parafacials blackish, with a thin coating of lustrous gray pollen; frontals 10 to 12, hardly reaching base of antenna? and not strikingly diverg- ent below; parafacials with a row of ratlier strong bristles, ascending diagonally along the expanding facial impression; antenutC dark brown, third joint one and one-half times the second and reaching two- thirds of the way to the vibrisste, which are at the oral margin thougli mucli above the lower line of the head; arista thickened at base, penultimate joint very short, ])lum()sity shorter tlian in a normal Sarcopliaga and dwindling beyond tlie middle, so that from a third to almost a half of the length is bare ; facial ridges witli small hairs extending a])out. a third of the way up ; facial impression reddish-brown, conspicuous, wide toward the vil)rissa?; bucca over two-fifths of the eye- height, very bristly below; back of licad and bucca blackish, Avith tliin cincrous ])ollen, tlie former with four or five rows of bhick hair and almost no pale beard; outer vertical bristle quite small but larger 48 THOMAS SAY FOUNDATION than adjacent hairs; palpi black, ordinary; proboscis black, very short, fleshy. Thorax strongly bristled, subshining black all over, but when viewed from behind showing the three main stripes as in Sarcophaga, dark with pollinose interspaces ; 3 ps dc ; 3 stpl ; anterior acrostichals very strong, 3 pairs; prescutellars also large, equal to the dc beside them; scutellum with three pairs of mar- ginals, of which the basal as usual are rather on the upper edge of the margin, the other four bristles are of equal length and equally spaced around the apical Y>aYt; a large discal pair also on scutellum; notopleu- ral two large and two small. Abdomen rather cylindrical, black with almost silvery pollen in a tessellated pattern, leaving an in- distinct median black stripe and somewhat shining hind margins ; second segment with a large, erect pair of median marginal bristles; third and fourth with each a marginal row of 18 or 20. Hypopygium of smallish size, subopaque, black; first segment with a row of small bristles ; second witii rather numerous erect black hairs ; forceps black, quite straight and tapering, with long black semi-recum- bent hair behind except apically; accessory plate blackish, ending in an acute angle; posterior clasper smaller than anterior, deeply excised in front and ])earing a long hair; penis rather short and stout, the distal segment bent forward with a blunt angle be- hind, below it in front a pair of black projecting points and basad of these a pair of long processes rather rolled in at tip ; sternites sliglitly decreasing in width, the fifth concealed. Legs black, strong and bristly; middle femur with long bristles but no comb below; middle tibia with two bristles on outer front side near middle ; hind tibi^ without villosity. Wings subliyaline, slightly yellowisli ; a large costal spine; fourth vein witli rounded bend, ending moderately (but not vxry) far before tip, the first posterior cell rather narrowly open in the two males ; SARCOPHAGA AND ALLIf^S 4-9 first vein bare, third witii several hairs; third costal segment shorter than the fifth. Female. Front .313 of head (average of five, — • .289, .307, .309, .329, .330), orbitals present; outer verticals well developed; bucca fully half the eye- height; scutellum as in male; genital segment black, with quite long bristles, orifice rounded ; middle tibiie with three bristles on outer front side ; costa with two spines, one of striking size ; first posterior cell usually closed in or just before the margin, sometimes as widely open as in the two males. Length O^-S^/o mm. Two males, Germantown, Pa., July 2, 190-t, and May 20, 1905, collected by Mr. H. S. Harbeck. Nineteen females: 12 from Germantown, Pa., in summer of 1904 and 1905; 2 from Manahawkin, N. J., Sept. 5, 1909, and July 11, 1910; 3 from Tren- ton, N. J., Aug. 21, 1909, and Aug. 5, 1910 (all from Harbeck; one from Melrose Highlands, ]Mass., June 1, 1911; and one from "N. Andover Col." Mass., July 16, 1911 (H. E. Smith). Holotype.— Male, No. 20,490, U. S. X. M., from Germantown, Pa. Allotype. — Female, U. S. N. jNL, same number and locality. HYPOPELTA New Genus. (vTTo, below; irekTr], shield) Allied to Camptopyga, but differing in having no macrocha}t^ on sides of face and with different genital characters in both sexes. Black species with about tlie usual Sarcopliagid appearance ; the male without orbitals, and with small ocellars and distinct outer verticals; paral'acial with row of hairs near the eye below, tlie lower ones rather coarse; frontals diverging rapidly below, reaching as far as middle of second antennal joint; third anten- nal joint twice the second, witli long, slender arista which is short-plimiose for two-tliirds its lengtli. Vi- brissa? at oral margin, facialia very flat, bare except 50 THOMAS SAY FOUNDATION for a few hairs below; palpi normal, proboscis short. Eyes bare. Thoracic cheetotaxy: ps dc 3; ant dc 2; hum 3; intrahum 1; presut 1; ant acr 0; ps acr 1 (prescutel- lar) ; stpl 3; scutellmn with 1 marginal and 1 apical, both long, the latter nearer to each other than to the former. Abdomen of male very large and clubbed api- cally, more than in Camptopyga; first segment de- clivous, with crescent-shaped row of 8 bristles near lower border, a deep notch between the two segments, the second bulging abruptly from the suture into a hump, on which are several long, upcurved macro- chsetai. First four abdominal segments short on ven- ter, the sternites covered with erect long hair ; the fifth sternite standing out abruptly as a large protuberance which opens V-shaped behind; a sixth sternite stands out in plain sight as a lobe projecting from underneath the fourth at the side of the fifth, the end of this lobe almost or quite rests upon the forceps in repose. The female has the genital segment much larger than in Camptopyga; wing as in that genus. Tj^pe. — Hyiwpclta scrofa, new species. No. 13. Hypopelta scrofa n. sp. Male. Front .214 of head (average of four, — .197, .217, .221, .231) ; middle stripe more than twice the width of one side above; parafrontals and para- facials decidedly silvery; antennae and palpi black; back of head with three rows of black hairs and only a little pale beard. Thorax with 3-5 black stripes ; the aj^ical scutel- lar bristles reach to the third abdominal segment. Abdomen black with changeable pollen; seg- ments 2-4 with a transverse row of four silvery spots against the front margin, giving the effect of three shifting black longitudinal stripes between them; first and second segments with only one moderate lateral bristle; third with a pair of erect median marginals SAECOPHAGA AND ALLIES 51 and two laterals; fourth with a marginal row which extends down the sides in full strength and numbers about 24 bristles. Hypopygium as partly described; first segment black, second red. Forceps straight, hardly at all di- vergent, blackish, behind with dense hair slanting back and outward; at the tip a very minute tooth. Accessory plates large, broad, anterior edge shining black. Posterior clasper long, stout, and strongly curved; anterior clasper surprisingly modified, — it is reversed in position so that it closes backward, and greatly elongated, lying inside the margin of the cav- ity so as to be almost invisible even basally ; the great- ly elongated and enlarged tip is so far forward that it cannot be seen in ordinary spread specimens, but the hypopygium must be taken apart to reveal it; it is as large as the forceps of most species. A row of reddish hairs along its more slender basal part are plainly visible anterior to the hind clasper. Penis with a long, swollen, bright red basal segment, nar- rowing to a neck where it joins the distal one, the junction close and not hinged; distal segment with a rather small central mass, dorsally dividing just be- yond this into two long thin plates, which diverging slightly curve over the end and partly conceal the inner parts; these are several incurved processes, basad of which is a flat transverse plate with three lobes; caudad of this a brown thin margin expanded from the side of the segment. Legs black; middle femora without comb; mid- dle tibia witli two bristles on outer front side; hind tibia without villosity. Wings subhyaline; costal spine large; third cos- tal segment much shorter than fifth; the distance of the tip of the foin-th A^ein from the extreme apex is about two-thirds the fifth costal segment. Female. Front .312 of head; palpi sliglitly cla- vate; genital segment deep red, conspicuous, not re- tracted, witli a fi-inge of stout bristles; below it has 52 THOMxVS SxVY FOUNDATION a row of appressed black hairs at base and another in middle ( sixth sternite coalesced with Hi). Length 6^^-8l/o mm. Thirteen males and three females; se^en males are from LaFayette, Ind., collected by the anthor; two males and two females Opelousas, La., in the Hough collection; one male Tifton, Ga., also in the Hough collection; one male each from Georgesville and Kent, Ohio, in Professor Hine's collection; one male Florida (C. W. Johnson) ; the third female is in the Hough collection, collected in Wisconsin by Professor Wm. M. Wheeler. Holotype.— Male, No. 20,491, U. S. X. M., from LaFaj^ette, Ind. Allotype.— Female, No. 20491, U. S. X. M.. from Opelousas, La. NOTOOH^TA New Genus. {vuiTos, back; xcitri?, bristle) Eyes bare; front not ^ery prominent, narrow in male; upper frontals strong, several of the upper pair reclinate in a decreasing degree, the rows mod- erately divergent below and reaching almost to the middle of the second antennal joint; third antennal joint long, arista long and with very long plumosity almost to its tip ; vibrisste at oral margin, not approx- imated ; hairs of paraf acials minute or wanting ; palpi normal; proboscis short, fleshy; back of head nearly flat. Thorax witli uncommonly long bristles, as fol- lows: Ps dc 2 (the anterior not much more than half way to the suture, usually a coarse hair close in front of it) ; ant dc 2 large; ant acr 0; prsc 1 small; hum 2; npl 2; inthum 1 ; prsut 1 large; pa 2; inta 2 or 1 (the anterior small or absent) ; supa 3 (the middle one large) ; stpl 3; scutellum with 3 or 4 marginals (only the first and last large), one small apical, and some- times a minute subapical. SARCOPHAGA AND ALLIES 53 Abdomen largely shining; marginals only on third and fonrth segments, no discals; hypopygium small. Legs without villosity, not very bristly. Wings without costal spine; third costal seg- ment shorter than fifth; first vein bare, third hairy near base; apical cell open far before the tip of the wing, widened past its middle by a sinuosity of the third vein; apical crossvein forming a right angle with or without a small appendage. Type. — Notochacta suhpolita n. sp.; Van der "Wulp's Sarcophaga plumigera, as I identify it, also goes here. Table of Species. Facialia ( facial ridges ) hairy to much above the middle No. 1.5. plumigera'V. d. W. Facialia hairy only a little above vibriss^e No. 14. suhpolita n. sp. No. 14. Notochaeta suhpolita n. sp. Male. Front .147 of head (three gave identical measurements) ; parafacials and parafrontals silvery pollinose, but with a yellow tinge, the former with a few almost imperceptible hairs; frontal stripe black- ish, almost half the front in width above, widening below; frontal bristles about 12, erect and long, three or four of the upper pairs reclinate in some degree; antennae black, third joint 3 times the second, reacli- ing four-fifths of the way to the vibrissje ; facialia low and bare, face flat; palpi and proboscis black; bucca one-fiftli tlie eyeheight; hairs of back of head and of bucca all black; ocellars smallish, outer vertical absent. Thorax witli briglit yellowisli gray pollen alter- nating with 3-.5 sliining black stripes, tlie median of whicli is narrowed in front; pleunu with satiny gray pollen, in a certain light tliere is a black stripe from the humerus to the front part of tlie stcrnopleura ; anterior intraalar present but small. Abdomen largely shining black, ])ut the anterior part of tlie segments with thin but l)right, changeable 54 THOMAS SAY FOUNDATION submetallic gray pollen, leaving a distinct, shining and metallic median black stripe; first and second segments with only lateral bristles; third with a roAv of 10 strong, erect marginals; fourth with a row of 10 or 12. Fifth sternite black, V-shaped, very incon- spicuous, with slender, erect hairs on the exposed edge. Hypopygium black, opaque, small ; first segment with a row of small bristles behind ; second with erect black hairs, hardly bristly, in profile rather humped near base. Forceps small, black, straight, hardly di- vergent, uniformly tapering. Accessory plate brown, rather long, and narrow at tip; posterior clasper small, slender, erect, blackish, with hooked tip and a long hair from the base in front ; anterior clasper red- dish toward tip, slender, rather sinuous. Penis with short, brownish basal segment; distal segment black- ish, its basal portion slender, then rapidly widening; behind, it has a prominent angle, beyond which is a concavity bounded by two spreading ridges which converge and blend in a rounded apex ; in front, there is a mandible-like process and several pairs of small hooks on the main part, suggesting the head of an animal with open mouth. Legs black ; middle femur without comb ; middle tibia with one bristle on outer front side; hind tibia not villous, but ciliated on the outer side with several rows of noticeable hairs. Wing subinfuscated in front and along the veins, almost hyaline behind. Female. Front .268 of head (average of two, — ■ .263 and .274) ; with the usual ocellars and outer ver- tical, but only 8 frontals; scutellum with the same bristles as in the male, including apicals (in both sexes the apicals seem to arise a trifle before the hind edge) ; genital segments black, small. Length 8 mm. Three males and two females, Los Amates, Guatemala, Jan. 16-20, 1905, collected by Professor Jas. S. Hine. SARCOPHAGA AND ALLIES 55 Holotype. — Male, in Professor Hine's collec- tion. Allotype. — Female, in same place. No. 15. Notochaeta plumigera V. d. W. Van der Wulp, Biologia, Dipt., ii, 273, pi. vii, f. 6, 6a {Sarcophaga). — Tabasco, Mex. Male. Front .187 of head; parafrontals and parafacials golden pollinose; frontal stripe red; ocel- lars present, outer vertical absent; frontals 8 or 9; antennae brownish-black, third joint about five times the second, reaching six-sevenths of the way to the vibrissa?; arista longer and witli longer plumosity than in suhpolita; facialia ridged, hairy more than half way up; back of head with three rows of black hairs, on the middle and below to the metacephalic suture with pale hairs. Thorax and abdomen as in suhpolita, fifth stern - ite not visible. Hpyopygimu in the described specimen with brown first segment, the second yellowish (Van der Wulp gives them as black), but they seem to be bleached. Forceps black, short, concave behind for half their length, then excised on inner side and end- ing in slender point, which in profile is nearly straight, with a small claw ; accessory plate brown, rather long and narrow; posterior clasper brown, slender and gently curved, hooked at tip, and with a long hair far back from the tip; anterior clasper longer, slender, brown, with hooked tip. Penis stout, black and blunt at tip, the basal part yellowish; on the front side are two diverging plates near the apex, their front edges minutely serrate ; the back plate makes a pronounced angle at tlie tip. Legs and wings as in suhpolita. Length 9 mm. One male, Punta Arenas, Guatemala, jNIarch 14, 1915, collected by Professor Hine. Type. — In the Godman and Salvin collection, London. Not seen by the writer. 56 THOMAS SAY FOUNDATION EMBLEMASOMA New Genus. (efx(3}\.T]iJLa, mosaic; crco/xa, body) Differs from Sarcophaga mainly in having the vibrissa} quite high above the oral margin (as far as the length of the second antennal joint) and consid- erably approximated, so as to narrow the facial plate below more than usual. Type, Emblemasoma crro, the following species; a second species is E. jacialc, and the two agree in the following characters among others : Parafacials with several rows of hairs, none bristle-like, encroaching on the facial impression; parafrontals also with small hairs; frontals reaching to about middle of second antennal joint, not sud- denly diverging below, still with one or more extra bristles just outside the lowest in the row, evidently somewhat variable; antenutE short, third joint less than twice the second, and reaching three-fifths to three-fourths the way to the vibriss^e; bucca very wide, about half the eyeheight; scutellum with 3 or 4 marginal pairs, together with one discal and a small apical; 4 dc; ant acr long but not very distinct from the adjacent long, erect hairs; stpl 3; npl 4; sa 3; inta 2; pa 2; epaulets black; subepaulets almost white. Table of Species. — Males. Legs, palpi, proboscis and antennie black (wide- spread erTO n. sp. Legs, palpi, proboscis and antennae yellow (Georgia) faciale n. sp. No. 16. Emblemasoma erro n. sp. ]Male. Front .206 of head (average of three, — .200, .203, .214) ; parafrontals and parafacials with yellow satiny pollen, with brown reflections opposite antenncE ; frontal stripe brown, w ide and well-defined ; ocellars present, outer vertical not; antennse brown- ish-black, small, the second joint rather long, third hardly longer than second, lacking ahiiost its own length of reaching vibriss^e ; palpi and proboscis black, SAKCOPHAGA AND ALLIES ' O/ ordinary ; back of head with about four rows of black hairs, the inner irregular, and with considerable pale yellow hair about the neck and below, which does not extend forward on the bucca. Thorax gray pollinose, with 3-5 black stripes; the subniedian gray stripes di^ided farther than the suture. Abdomen gray pollinose, witli black median stripe and two shifting lateral ones; fourth segment red behind more or less broadly, and a faint reddish tinge showing on the sides of the third and even the second segment; tirst and second with only lateral bristles ; third with a stout median marginal pair and several laterals, making an interrupted row; fourth with a row of about a dozen. Hypop5"gium red; first segment of medium size, subshining, with row of bristles behind ; second rather small, with some erect bristly hairs; forceps red on basal half, not large, tapering to. middle, then with a sudden enlargement almost like a barb behind, not nmch tapering beyond, the tip blunt and rounded. Accessory plate red, finger-like; posterior clasper red, with brown flat tip not nuich hooked; anterior clasper black from middle, with flat rather straight tip. Penis with short, red basal segment much re- tracted; distal segment becoming black, rounded but not greatly enlarged at tip, bearing a striking pair of large red outstanding arms on the front side beyond the middle. Fifth sternite red, inconspicuous, V- shaped, with only small hairs on inner side and very few l)ristles on edge. Legs black; middle femur witli two rows of sj)ines below, the posterior forming a comb; middle tibia ^vitli two bristles on outer front side; hind femur M'ith two rows of long bristles below; hind tibia not vilknis, with long bristles on hind side. Wings su})-liyaline; no costal spine; third costal segment longer than fifth and six together; first vein Ijare; tliird hairy at base for a sliort distance. I^ength 13-1.5 mm. 58 THOMAS SAY FOUNDATION Three males: one from Russell Co., Kans., alti- tude 1,350 feet, July 26, 1912 (F. X. Williams), in the University of Kansas; one from Manahawkin, N. J., Aug. 21, 1911 (Harbeck) ; and one from Rio Janeiro, Brazil, collected in November by H. H. Smith, and given me by Professor Williston. Holotype.— Male, No. 20,492, N .S. N. M., from Manahawkin, N. J. No. 17. Emblemasoma faciale n. sp. Male. Front .209 of head (one specimen) ; para- facials and parafrontals gray, frontal stripe brown; frontals 13, not widely divergent below, on one side there is an additional bristle outside the lowest front- al; antennse red, third joint brown toward tip, one and two-thirds times the second, reaching three-fourths of the way to the vibrissse; palpi yellow, slender, darker at tip; proboscis yellow, short; back of head as in erro. Thorax gray, with 3-5 black stripes; the sub- median gray stripes wide and rather widely divided past the suture. Abdomen as in crro, fourth segment about half red. Hypopygium red, very similar to that of erro, but smaller; first segment retracted, with row of bristles on hind edge; second small; forceps entirely red, small, shaped as in erro; accessory plate red, long and finger-like; posterior clasper red, slender, straight, with dark, hooked tip; anterior clasper red, low, rather straight, excised on inner side near tip; penis as in eiTO. Fifth sternite as in erro. Legs red, tarsi black; bristles of mid femur comblike both in front and behind in the lower rows ; otherwise the bristles of the legs are as mentioned for erro. Wings not in good condition, but as far as the characters are discernible they are as in eri'o; this ap- plies to absence of costal spine, long third costal seg- ment, and hairy third vein. SARCOPHAGA AND ALLIES 59 Length 11 mm. One male, Tifton, Ga., Aug. 27, 1896, in the Hough collection of the University of Chicago. Plolotype. — In the Hougli collection. STHENOPYGA New Genus. (adeuos, strong; -rrvyq, the hinder end) Type, Sthenopyga glohosa, new species, de- scribed below. The characters are given in full in the description; those of principal generic importance appear to be the narrow parafacials, almost entirely destitute of even minute hairs, the large eye and nar- row" bucca; only one intra-alar, the posterior one; front protuberant above the insertion of the antenna;. No. 18. Sthenopyga globosa n. sp. Male. Front .188 of head (average of five, — .168, .184, .188, .195, .204) ; the eyes not diverging as rapidly below as in most species; face and front sil- very with slight yellow tinge; frontals not suddenly diverging laterally at lower end of the row, but end- ing higher up, the lowest barely beyond the base of the second antennal joint, and not diverging more than the lower half of the row as a whole ; parafacials entirely bare of hairs, densely pollinose; vibrissse at oral margin, only a few small hairs above them; an- temicC brown, third joint slender, fully twice the sec- ond, reaching three-fourths of the way to the vibrissse ; arista of medium length, long-plumose on basal two- thirds; bucca one-fifth the eyeheight; palpi dark yel- low, apical two-fifths brown; proboscis short, fleshy; outer vertical not differentiated; back of head witli only a few pale hairs about the neck. Tliorax densely gray pollinose, not shining in any part; when viewed from behind the usual 3-5 black stripes are visible but slender, in front view the 3 are brown; 3 ps dc; st])l 2; no ant acr; prescutellar very small; scutellum with two marginals (one very small between them), no subapicals, no apicals. 60 THOMAS SAY FOUNDATION Abdomen gray, densely gray pollinose, tessel- lated, the fourth segment mostly red ; first and second with only lateral, third with row of 10 or 12 marginal bristles; fourth with row of about 18. Hypopygium red ; first segment rather pollinose, with a row of small bristles at hind margin; second segment red, rather small and flattened; forceps small, red except tip, rather evenly pointed, posterior side basally with hair sloping away from apex; ac- cessory plates yellow, finger-shaped, bare; both clasp- ers of ordinary form ; penis with globose, blackish dis- tal segment, the apical portion of which curves for- ward and is divided mesially into two flat pieces side bj^ side, the side plates extending considerably for- ward; fifth sternite concealed, pale yellow, the branches slender, bearing a few black hairs. Legs yellow, or the femora partially infuscated, middle tibia with one bristle on outer front side ; mid- dle femur with moderate bristles below on hind side, not forming comb; hind tibia with very sparse villos- ity, only a few erect hairs, mostly on inner side ; claws and pulvilli long, the latter yellow. Wings hyaline, third and fifth costal segments about equal; first vein with only a few coarse hairs; third vein hairy almost to crossvein ; no costal spine. Female. Front .252 of head (average of five, — .231, .244, .246, .258, .282) ; orbitals and outer ver- ticals present; lower frontals as in male, or a very little more diverging than in the male, but not ex- tending any further down; parafacials with a very few hairs, minute and mostly pale; bucca one-sixth the eyeheight; scutellum with three marginal (the middle small), one pair subapicals (almost apical in position) ; genital segment red with roundish or tri- angular opening; middle tibia with 2 bristles on outer front side; middle femur without combs; first vein with numerous hairs . Length of male Tl/o to 8l^, of female 6 to 8 mm. Six males, seven females, Opelousas, La., March, April and May, 1897, all in Hough collection. One SAKCOPHAGA AND ALLIES (31 female lias a pair of median marginals on second ab- dominal segment. Another female, ]Miami, Florida, Dec. 19, 1912, shows no other apparent differences but has large median marginal bristles on second ab- dominal segment, in fact a complete but slightly n'- regular marginal row. It will require more material to explain these differences. Holotype.— Male, No. 20,493, U. S. X. JNI. Allotype.— Female, No. 20493, U. S. N. M. HARPAGOPYGA New Genus. (apTayr], hook; Tuyr], the hinder end) Parafacials with a few very delicate almost im- perceptible hairs; vibrissa? slightly above oral margin and slightly approximated (as far apart as two-thirds the width of the front, in the male) ; ocellar bristles entirely absent; orbitals none in male, one in female; frontal bristles descending to base of antenmu; arista as in Sarcophaga; third antennal joint about twice the second, slender. Thoracic cha^totaxy exactly as in Camptops, except that there are five or six bristles in the hypopleural row (this is probably somewhat variable in all species). First vein bare, fourth end- ing not very far before the apex (half the fifth costal segment ) . Agrees with Camptops and Sthenopyga in hav- ing the face destitute of the row of bristly hairs near the eye, and in the fourth vein ending not far before the apex of the wing. In this series of genera, Camp- tops is at once separated by having the vibriss'lilan