BOSTON PUBLIC LIBRARY 3 9999 06317 800 6 HUNTING AND HARVEST TRENDS FOR MIGRATORY GAME BIRDS OTHER THAN WATERFOWL: 1964-76 J/X UNITED STATES DEPARTMENT OF THE INTERIOR FISH AND WILDLIFE SERVICE Special Scientific Report— Wildlife No. 218 Library of Congress Cataloging in Publication Data Martin, Elwood. Hunting and harvest trends for migratory game birds other than water- fowl, 1964-76. (Special scientific report— Wildlife; no. 218) Supt. of Docs, no.: I 49.1533:218 1. Fowling— United States. 2. Fowling— United States— Statistics. 3. Up- land game bird shooting— United States. 4. Upland game bird shooting- United States— Statistics. I. Title. II. Series: United States. Fish and Wildlife Service. Special scientific report— wildlife; no. 218. SK361.A256no. 218[SK313] 338.3'72'830973 79-607015 NOTE: Use of trade names does not imply U.S. Government endorsement of commercial products. HUNTING AND HARVEST TRENDS FOR MIGRATORY GAME BIRDS OTHER THAN WATERFOWL: 1964-76 By Elwood M. Martin * M J \ UNITED STATES DEPARTMENT OF THE INTERIOR FISH AND WILDLIFE SERVICE Special Scientific Report— Wildlife No. 218 Washington, D.C. • 1979 Hunting and Harvest Trends for Migratory Game Birds Other than Waterfowl: 1964-76 by ElwoodM. Martin U.S. Fish and Wildlife Service Office of Migratory Bird Management Laurel, Maryland 20811 Abstract Hunting activity and harvest estimates for 10 species or groups of migratory game birds other than waterfowl, based on data collected for 13 seasons (1964-76) in the Service's Annual Ques- tionnaire Survey of U. S. Waterfowl Hunters, are presented. The 1964-75 data, available in time for additional analysis, are discussed in terms of their usefulness as index values for detecting short-term changes and long-term trends and demonstrating regional differences. Species or groups for which estimates were obtained include the white-winged dove {Zenaida asiatica), band- tailed pigeon [Columba fasciata), mourning dove {Zenaida macroura), American woodcock {Philo- hela minor), common snipe (Capella gallinago), sandhill crane {Grus canadensis), sora {Porzana Carolina), other rails, gallinules, and American coot [Fulica americana). Among an average annual population of 2,013,300 duck stamp purchasers (1964-75), the mourning dove was the most popular of these game birds (averaging 478.000 hunters per year), followed by the coot (172,600 hunters), woodcock (168,700 hunters), snipe (74,600 hunters), band-tailed pigeon (25,400 hunters), white-winged dove (23,700 hunters), rails other than the sora (14,300 hunters), sora (6,300 hunters), gallinules (4,600 hunters), and crane (4,000 hunters). The average annual harvest index (unadjusted for reporting biases), for each species during this period was 12.05 million mourning doves, 1.17 million coots, 588,000 woodcock, 422,000 snipe, 314,000 white-winged doves, 192,000 band-tailed pigeons, 105,000 rails other than the sora, 32,100 soras, 26,400 gallinules, and 7,200 cranes. Several significant year-to-year changes were detected in mourning dove and woodcock average annual bag estimates at the management unit-flyway level. A number of long-term trends were noted as well, including significant increases in the proportions of duck stamp buyers also hunting mourning doves (about 1% annually), woodcock (3%), snipe (2%), cranes (8%). and rails other than the sora (6%). Significant long-term changes in average annual bag included a decrease for band-tailed pigeons in the three Pacific coast States and an increase in the Atlantic Flyway and nationally for rails other than the sora. In addition, a significant long-term increase averaging 2% per year was noted in the ratio of duck stamp sales to hunting license sales. High correlations obtained between certain results in the survey and the results of independent surveys are discussed. Various problems associated with using a waterfowl-hunter sampling frame as the basis for a survey of the hunting of migratory game birds other than waterfowl are evident throughout these analyses. The inescapable conclusion is that, although this survey of waterfowl hunters provides valuable data on the subject, a better sampling frame (or frames) for measuring the activity and success of all hunters of migratory game birds other than waterfowl is needed to meet the high standards now being set for the management of this valuable resource. Since 1964, the U. S. Fish and Wildlife Service has collected information on the hunting of most migra- tory game birds other than waterfowl through its Annual Questionnaire Survey of U.S. Waterfowl Hunters. Ten species or groups of such birds are now included on the questionnaire: white-winged dove (Zen- aida asiatica), band-tailed pigeon (Columba fasciata), mourning dove (Zenaida macroura), American wood- cock (Philohela minor), common snipe (Capella gal- linago), sandhill crane (Grus canadensis), sora (Por- zana Carolina), other rails, gallinules, and American coot {Fulica americana). In addition, the barnyard pigeon or rock dove (Columba livia) has been included since 1967 to reduce the apparent tendency to report them as band-tailed pigeons; rock dove harvest data are not included in this report. The population of duck stamp buyers is used in the absence of a more suitable sampling frame for hunters of migratory game birds other than waterfowl. It is incomplete in varying degrees depending on the geo- graphic area and species of interest. Therefore, for most of these species, hunting activity and harvest figures obtained are index values, useful primarily as indicators of changes and trends and not as measure- ments of total hunting activity and success. Mac- Donald and Martin (1971) summarized survey data for the first 5 years, and a detailed description of the survey will be found there. These early estimates, with some revisions and corrections, are included here together with estimates for an additional 8 seasons. Information on coot hunting activity and success, as obtained in the waterfowl harvest survey since its inception in 1952, was not examined by MacDonald and Martin (1971) but is included here. Annual changes and long-term (1964-75) trends are examined briefly (estimates for the 1976-77 season became avail- able after work on the 1964-75 data had been com- pleted) and, for several species, data from this survey and several independent surveys are compared. I gratefully acknowledge the help and advice of J. W. Artmann, U. S. Fish and Wildlife Service, in the planning and assembly of material for this report. Procedures and Limitations The survey and its analysis have remained essen- tially as reported by MacDonald and Martin (1971). Certain aspects, however, deserve closer examination or re-emphasis. The Sampling Frame The information source is the sample of duck stamp purchasers selected for the annual waterfowl harvest survey. M. F. Sorensen, Office of Migratory Bird Man- agement, Laurel, Maryland, in an administrative report on sandhill crane hunting dated 7 July 1977, found that 71% of the 1976-77 season crane hunting permit holders also bought duck stamps. In most States, for hunters 16 years old or older, probably at least 90% of the coot hunters, somewhat fewer (75 to 90%?) rail, gallinule, and snipe hunters, and still fewer woodcock, dove, and pigeon hunters purchased duck stamps. Although these figures are speculative, they reflect the inconsistency of the relationship between waterfowl hunting and other migratory game bird hunting, an indication that waterfowl hunters form a poor sampling frame for hunters of other migratory game birds. The possibility that some of the estimates presented here for waterfowl hunters may contain substantial response/nonresponse biases is a further complication. For example, both 1975 and 1976 surveys of all crane hunting permit holders (the best sampling frame one could hope to have) produced somewhat lower esti- mates of crane hunting activity and success than did the waterfowl hunter surveys (M. F. Sorensen and H. M. Reeves, Office of Migratory Bird Management. Laurel, Maryland, administrative report on sandhill crane hunting dated 9 July 1976; M. F. Sorensen, administrative report cited above). Such biases do not necessarily negate the value of data provided by waterfowl hunters as indicators of changes and trends, however. The relative importance of waterfowl hunting to all hunting in each State (Table A-l) is obviously an important consideration in using the data obtained in the survey. The average proportion of all licensed hunters also buying duck stamps in 1964-75 was 13%, ranging from less than 1% (West Virginia) to over 51% (North Dakota). Furthermore, these percentages in- creased significantly in about one-half the States during this period; there were no significant decreases. Obviously, waterfowl hunters will not be representa- tive of all hunters in a State, and the problem is com- pounded when waterfowl hunters from the various States are combined to produce totals for larger units. However, basic information on the relative importance of each species to hunters and to individual States and on trends in harvest, particularly changes in its size and distribution, should be almost as apparent from a sample of waterfowl hunters as from a sample of all hunters. Ruos and Tomlinson (1968) developed a complicated procedure for extrapolating total dove harvest from dove harvest by waterfowl hunters, but some problems remained, and it was not practical to repeat this procedure every year or use it for other species. Clark (1972:13-14) calculated that less than 50% of the total U. S. woodcock harvest can be attrib- uted to waterfowl hunters. In view of such problems, no attempt has been made to project estimates beyond the waterfowl hunter framework for any of the species included in this report. The Survey Except for the addition of several species, the ques- tionnaire design introduced in 1964 remained un- changed through 1968. A slightly revised design was tested concurrently with the original questionnaire in 1967 and 1968 and has been used exclusively since. Some differences in results attributable to ques- tionnaire differences were noted and appropriate adjustments (unpublished report by E. M. Martin, Office of Migratory Bird Management, Laurel, Mary- land, dated 17 December 1970) have been made to maintain comparability among years. Harvest estimates for coots have routinely appeared in Federal reports on waterfowl hunting. Application of the bias-adjustment procedure developed for water- fowl in the 1950's by E. L. Atwood (mimeographed report dated September 1959 on file at Migratory Bird Management Office, Laurel, Maryland) resulted in a sizeable reduction (averaging 35%, see Benning et al. 1975:64) for response bias in coot harvest figures through the years. For the present report, the data on coot hunting activity and success were re-examined for all years of the waterfowl harvest survey and recal- culated without bias adjustment; nor have bias adjust- ments been applied to data for other species. However, because coots have been handled like waterfowl since the survey began, coot data still differ somewhat from those for the other species included here. Active coot hunters were not identifiable, but successful coot hunters were. In addition, estimates of harvest by waterfowl hunters under 16 years; of age (conserva- tively, an additional 9%) and of unretrieved kill are available from this survey for coots but not for the other nonwaterfowl species. Analysis of Results To demonstrate potential uses of the survey results, I provide several comparatively simple examples of data analyses. These are intended to show the strengths and weaknesses of the data and point the way for further analysis by those interested in par- ticular aspects. The approaches used fall into two general categories: (1) seeking evidence of significant differences and changes or trends on the basis of this survey alone, and (2) evaluating the degree to which the results of this survey are supported by (correlated with) the results of independent surveys. The independent surveys examined include other Fish and Wildlife Service surveys and selected State surveys. Many States collect harvest data on at least some of the species treated here, and more detailed comparisons with these data are encouraged. A better under- standing of the harvest, and of each survey, should result. Survey Results and Discussion Activity and harvest estimates for the hunting of other migratory game birds by waterfowl hunters in the United States from 1964 through 1976 are sum- marized by species in Table 1. A more detailed sum- mary (by State, flyway, and management unit) of the 1964-75 data available for inclusion in the additional analyses noted above appear in Appendix A, and similar detail for the 1976-77 season, which became available later, are presented in Appendix B. Statis- tical examinations of differences and trends are sum- marized for each species in Tables 2 through 6. The results are discussed briefly in this section by species. White-winged Dove During 1966-75, in the five States in which white- winged dove hunting has been permitted, an average of 23,700 waterfowl hunters (7.7% of the duck stamp buyers) harvested about 313,500 white-winged doves (13.2 birds per hunter) annually (Table A-2). Both the percentage hunting white-winged doves and the average bag decreased slightly during the period, but neither change was statistically significant (Table 2). However, on an individual State basis, decreases in the percentage hunting in Texas and the average bag in Arizona were significant, if all statistical assumptions, which tend to be weakest for the less widely hunted species, are met. Band-tailed Pigeon These game birds have been hunted in seven States since 1970; only three States participated before 1968. In 1975, an estimated 19,200 (5.2%) of the waterfowl hunters in these seven States also hunted band-tailed pigeons, bagging about 107,900 or 5.6 each (Table A-3). The 1975 figures were the lowest obtained during the 1966-75 period. In the three States for which data for the entire period are available, both the percentage hunting and the average bag showed downward trends (Table 2). These trends were detected in all three States but were significant only in the Washington and combined area average bag figures. 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CD c _3 i © -r o © © CO o t~ O CO o ■* *o © 01 Q © o © © in © ^ © rH © © t- © a © od kO m © 00 o o © UD © CO | 3 o © CO © © © o CN © CO © id q © © CD T*. *f ^ © © UD CO •— i CO CO CO *-*j to q © fr- CO © © CO id fr- © © in m t-1 cm" ci CM ©* CN ©* m" O in* fr-* CO* rH eo* ©" ©* o Table 2. Summary of statistical evidence of long-term trends in the hunting of other migratory game birds by waterfowl hunters in those States that had an open season on the designated species every year during the period indicated. Percentage hunting each species Average bag Average Average Area Period annual Period annual Species and period included average8 change t value1' average3 change t valueb White-winged dove 5 States 7.75 -0.15 -1.44 13.15 -0.35 -1.59 (1966-75) Band-tailed pigeon 3 States 8.63 -0.13 -1.03 7.85 -0.20 -2.76* (1966-75) Mourning dove Eastern Manage. Unit 41.23 0.37 4.30** 28.13 -0.14 -1.43 (1964-75) Central Manage. Unit 44.79 0.51 3.66** 27.74 -0.13 -0.83 Western Manage. Unit 33.44 0.03 0.42 19.44 -0.19 -3.34** Total 39.80 0.37 4.69** 25.68 -0.09 -1.13 American woodcock Atlantic Fly way 20.26 0.50 6.77** 3.45 -0.02 -0.83 (1964-75) Mississippi Flyway 10.73 0.39 3.62** 3.58 0.01 0.33 Central Flyway 1.54 0.07 2.63* 2.58 -0.01 -0.10 Total 12.20 0.38 6.03** 3.51 -0.01 -0.37 Common snipe Atlantic Flyway 3.94 0.03 1.33 6.48 -0.09 -1.12 (1964-75) Mississippi Flyway 4.26 0.08 1.84 5.22 -0.05 -1.52 Central Flyway 2.39 0.10 2.52* 5.39 0.02 0.22 Pacific Flyway 3.87 0.18 5.28** 6.05 0.04 0.68 Total 3.78 0.09 3.15* 5.68 -0.04 -1.38 Sandhill crane 7 States 1.25 0.10 2.88* 1.79 0.12 1.37 (1968-75) Sora Atlantic Flyway 0.71 -0.04 -4.83** 6.52 0.13 0.94 (1964-75) Mississippi Flyway 0.36 0.01 1.66 4.32 -0.12 -1.20 Central Flyway 0.28 0.02 1.27 3.46 -0.32 -1.62 Total 0.44 -tr. -0.17 5.12 -0.09 -1.09 Other rails Atlantic Flyway 1.59 0.06 3.15* 9.66 0.38 2.88* (1964-75) Mississippi Flyway 0.89 0.08 4.11** 3.58 0.01 0.33 Central Flyway 0.48 0.02 0.75 3.85 0.19 1.83 Total 1.00 0.06 3.27** 7.20 0.23 2.37* Gallinules Atlantic Flyway 0.38 -0.03 -3.76** 5.32 -0.20 -1.54 (1964-75) Mississippi Flyway 0.30 0.03 3.31** 6.36 0.44 2.79* Central Flyway 0.15 0.02 2.63* 4.10 0.31 1.76 Pacific Flyway 0.14 tr. 1.15 5.53 0.11 0.42 Total 0.27 0.01 1.77 5.95 0.19 1.84 American coot1' Atlantic Flyway 6.60 -0.10 -0.81 6.38 0.02 0.45 (1964-75) Mississippi Flyway 10.44 -0.03 -0.21 7.92 -0.06 -0.71 Central Flyway 5.05 0.18 2.78* 4.02 0.04 1.55 Pacific Flyway 8.34 0.13 2.34* 7.41 0.01 0.08 Total 8.21 0.01 0.14 7.13 -0.04 -0.56 aAll years weighted equally in regression calculations. h*Indicates an average change significantly different from zero at the 95% confidence level; **denote the 99% confidence level. cCoot data based on successful coot hunters; data for other species based on active hunters of respective species. waterfowl hunters (24.2% of all duck stamp buyers) harvested slightly over 12 million mourning doves (25.2 doves per hunter) annually (Tables A-4, A-5). Almost 40% of the duck stamp buyers in those States in which dove hunting was permitted throughout this period hunted mourning doves. The proportion in- creased significantly (about 1% annually, a crude indi- cator of change obtained by expressing average annual change as a percentage of the period average) during this period in the Eastern and Central Management units and in the overall figures. In 13 of 30 States there were significant increases, but in 1 (Louisiana) a significant decrease was indicated (Table 3). Success rates were lowest (averaging 19.4 doves per > < 5 bo 2 3 C o 5 ffl > 5 -c •*■ co o > o c '3 u 3 ^ a; c ca > S J= < « o T3 CD •§ P c -o c 3 C a cv -a CD cu - E g « 0) CO ^ M- 3 CO -J OS CO ^ CO OS t- ID 00 O O C- in o n in w to n co w n h i-otNCNNOO'-'nddeN TT CD irt CO CM i-> CM 00 CD 00 O ^ O rH iD O w o N in CO CO OS CN •-< TT CO CM 00 O _ r-, ^ © P - ■ O O fc. 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CN CNOO-— < CO «— "COCOCOCOC-COCNCOO— i ^T OOOOOOOOOO-HCOO--' O CO CO OHO .— 'OOCS-HOiDCOCOCNOmaSiDCDCOCNCO coqcoooqoCH(NcD--qNqqcoHiN lodddHoiiOHXco'cNaJdcooo'dH mcoTiocoTconiomniNioiOTriN'r (NNhNOCOhCO OOOOOOOO COt-OSCNXCNt-OS Tros^oosoor- N O f O CD r- OS 00 CN if) 00 f CO ^ o ^h d oooooooo £ « co Jr Xi co "C -2 « -o> o S < Q tu CO CJ CO CO •sb-S 0)3 1> o O = « J S Sz ed "c CO C !tj o 0) 'c 'S > o CO CO •> c en B c O 3 cu c c .a CO JO CO CL tf c« e- > M 5 5 .5 f? bo -C r i c a 3 c 1 Q CO « Ul eu c >^ c _. c 03 & Si 0) 51 § > J= 5 a c 01 CO -: CO en s s s (•."O C < r hunter per season) in the Western Management, Unit (Table A-5), and decreased significantly (about 1% annually) during the period (Table 3). Elsewhere, Loui- siana showed a significant decrease while West Vir- ginia showed a significant increase. The overall down- ward trend was not statistically significant, but, as noted by Clark (1972:13) with reference to woodcock, such a decrease in average success as participation increases would not be surprising, since most of the additional hunters must be newcomers to the sport and relatively inexperienced. In year-to-year compari- sons, significant increases in average bag were indi- cated for the 1966-67 and 1972-73 seasons whereas a significant decrease was recorded for 1967-68 (Table 4). American Woodcock During 1964-75 an average of 168,700 waterfowl hunters (10.6% of the duck stamp buyers in the three eastern flyways; 12.2% in those States having a wood- cock season) harvested about 587,500 woodcock (3.5 per hunter) annually (Tables A-6, A-7). Waterfowl hunters in the Atlantic and Mississippi flyways participated in woodcock hunting in about equal numbers, but, although the rate of participation in the Atlantic Flyway has been about twice that in the Mis- sissippi Flyway (20% vs. 10%), success rates have dif- fered little (3.4 vs. 3.6 birds per hunter per season). The average annual harvest by waterfowl hunters ex- ceeded 50,000 in four States, of which Michigan was the leader (89,000). The proportion of waterfowl hunters also hunting woodcock has increased significantly since 1964 in all three flyways and in 19 of the 33 States in which wood- cock have been hunted regularly (Table 3); the increase has averaged about 3% per year. Significant long-term changes in average bag were indicated in seven States but not at flyway or U.S. levels. Indications of signifi- cant changes from the previous year in average annual bag (Table 4) were found in the Atlantic Flyway in 1969 (decrease) and the overall 1972 average (increase). Common Snipe Annually, an average of 3.7% (74,600) of the duck stamp purchasers in the United States hunted snipe during 1964-75, bagging about 422,000 snipe (5.7 per hunter) each year (Tables A-8, A-9). The average an- nual snipe harvest by waterfowl hunters was highest in Louisiana (96,000) and also exceeded 50,000 in two other States. The proportion of waterfowl hunters also hunting snipe increased significantly by more than 2% per year during this period, with significant increases recorded for 15 of 46 States; the increases appeared to become progressively larger from east to west across the country (Table 5). No long-term trend was evident in the average snipe bag. Significant increases were recorded in two States (Georgia, Mississippi), as were significant decreases (Nebraska, North Carolina). Sandhill Crane Currently, sandhill cranes are hunted in nine States, and this survey has included all crane hunting States since 1968 when seven States were involved. During 1968-75, duck stamp buyer participation in crane hunting in the seven States continuously involved averaged 1.2% (4,000 hunters), and they bagged an average of 7,200 cranes (1.8 per hunter) each season. Texas was the most important State for crane hun- ting, accounting for 39% of the hunters and 58% of the harvest during this 8-year period (Table A-10). The proportion of duck stamp buyers participating in the crane hunt in these seven States increased signifi- cantly since 1968 (averaging 8% annually); average bag has not changed significantly (Table 2). Sora An annual average of 0.4% (6,300) of the duck stamp buyers in the three eastern flyways hunted the sora during 1964-75, bagging about 32,100 birds (5.1 per hunter) each year (Tables A-ll, A-12). Harvests aver- aged over 2,000 birds per year in only five States in- cluding Iowa, which did not have a rail season until 1972. Louisiana and New Jersey were the top States, each with harvests of 5,300 birds per year by duck stamp buyers. Both percentage hunting and average bag become progressively smaller from east to west across the country. The only statistically significant long-term change during this period was a decrease of almost 6% per year in percentage hunting in the Atlantic Flyway (Table 2). Other Rails Through the years, this category has included the king rail {Rallus elegans), clapper rail (i?. longirostris), Virginia rail {R. limicola), yellow rail {Coturnicops noveboracensis), and black rail ILaterallus jamai- censis). In 1967, the black rail was omitted from the list of hunted species, followed a year later by the yellow rail, and these omissions continue. Very few birds of either species have been taken by hunters. Also, in most noncoastal States, the king rail has not been hunted since 1967. Even before then, the king rail was rarely, if ever, important in the rail harvest (Sanderson 1977:99-101). Thus, although hunters were not asked to report separately any rails but the sora. Table 4. Statistical evidence of year-to-year differences in the average annual bags of mourning doves and American woodcock by waterfowl hunters, 1964-75. Mourning dove American woodcock Eastern Central Western Mississippi Seasons Management Management Management Atlantic and Central compared Unit Unit Unit Total Flyway flyways Total 1964 vs. 1965 x"(n)a 0.04(16) 1.71(7) -1.56(7) 0.06(30) -0.09(17) -0.54(16) -0.31 (33) f-valueb 0.03 1.39 -2.07 0.07 -0.29 -2.12 -1.55 1965 vs. 1966 x(n) 2.31(16) 1.50(7) 1.04(7) 1.83(30) -0.05(17) 0.35(16) 0.14(33) f-value 3.05** 1.42 2.18 3.78** -0.34 1.89 1.23 1966 vs. 1967 x(n) -1.91 (16) -1.10(7) 0.0 (7) -1.27(30) 0.43(17) 0.23(16) 0.33(33) f-value -1.94 -1.06 0.0 -2.15* 2.07 0.80 1.91 1967 vs. 1968 x(n) 0.48(16) 0.62(8) -1.19(7) 0.14(31) 0.18(17) -0.08(17) 0.05 (34) f-value 0.68 0.38 -1.27 0.24 0.64 -0.17 0.20 1968 vs. 1969 x(n) 1.84(16) 0.10(8) -0.51 (7) 0.85(31) -0.63(17) 0.29(17) -0.17(34) f-value 1.47 0.08 -0.59 1.13 -2.77* 0.56 -0.59 1969 vs. 1970 x(n) -1.31 (16) -1.11(8) 0.29(7) -0.90(31) 0.02(17) -0.82(16) -0.39(33) f-value -1.66 -0.67 0.47 -1.50 0.07 -1.80 -1.47 1970 vs. 1971 x(n) -0.42(16) -0.20(8) -1.00(7) -0.49(31) -0.05(17) 0.16(16) 0.05(33) f-value -0.60 -0.10 -1.13 -0.76 -0.34 1.21 0.50 1971 vs. 1972 x(n) 2.32(16) 2.95 (8) 0.33(7) 2.04(31) 0.38(17) 0.31(16) 0.34 (33) f-value 2.35* 1.89 0.23 2.82** 1.83 1.30 2.23* 1972 vs. 1973 x(n) -2.64(16) -1.33(7) 0.94(7) -1.50(30) 0.05(17) 0.03(17) 0.04 (34) f-value -1.84 -1.25 2.35 -1.79 0.30 0.11 0.25 1973 vs. 1974 x(n) -0.15(16) -1.89(8) -1.39(7) -0.88(31) 0.04(17) 0.05(17) 0.04 (34) f-value -0.13 -1.38 -1.98 -1.26 0.19 0.24 0.31 1974 vs. 1975 x(n) -0.01(16) 2.39(8) 0.77(7) 0.79(31) -0.06(17) -0.05 117) -0.06 (34) f-value -0.01 2.03 0.86 1.50 -0.29 -0.32 -0.43 ax = average difference between State-level estimates of annual bag per waterfowl hunter; n = number of States. b* Indicates a significant correlation at the 95% confidence level; **denote the 99% level. some degree of separation is still possible. In most instances, rails reported from States not permitting hunting of large rails will be Virginia rails. However, harvest in these States has amounted to only 6% of the total. Clapper rails are probably dominant in the harvest in most other States although the degree of dominance will vary, and more knowledge of the sit- uation in each State is necessary should a separation be desired. An annual average of 0.9% (14,300) of the duck stamp buyers in the three eastern flyways hunted rails other than the sora during 1964-75, and bagged about 105,000 such birds (7.4 per hunter) annually (Tables A- 13, A-14). Louisiana and South Carolina were the most important harvest States for these birds, each har- vesting well over 20,000 annually and together ac- counting for 50% of the U. S. harvest by waterfowl hunters during this period. Outside Louisiana, the harvest of other rails was strongly concentrated along the Atlantic coast. Both the percentage hunting and the average bag showed significant increases in most areas (Table 2). Gallinules Included here are the common gallinule (Gallinula chloropus) and the purple gallinule (Porphyrula martin- ica). The latter is much more restricted in range, occur- ring primarily in parts of eight southeastern States. It has been excluded from Florida's list of hunted species since 1972. On the other hand, in Louisiana it appar- ently becomes numerous enough at times to be the subject of depredation control measures in rice fields. Some hunters undoubtedly fail to distinguish galli- nules (at least common gallinules) from coots. How- ever, in many areas, few, if any, gallinules are present when coots are being hunted, and vice versa, because of differences in migration dates (and therefore season dates), and main wintering grounds. A possible clue to the extent of this misidentification is found in the Service's Duck Wing Collection Survey which, since 1965, has also asked for coot (but not gallinule) wings. Common gallinule wings are received at the rate of about three per thousand coot wings (no purple galli- nule wings have yet been received), whereas waterfowl Table 5. Statistical evidence of long-term trends in the hunting of common snipe by waterfowl hunters in those States that had a snipe season every year during the period 1964-75. Percentage hunting snipe Average snipe bag Flyway and State Average Average Period annual t Period annual t average8 change valueb average" change valueb 1.91 0.03 0.54 2.72 -0.07 -0.82 2.57 -0.13 -1.48 3.78 0.14 0.74 20.62 0.67 3.47** 10.99 -0.03 -0.20 8.31 0.10 1.12 5.28 0.23 3.69** 2.65 0.12 1.72 3.74 -0.13 -1.04 0.89 -0.01 -0.31 3.56 0.05 0.33 1.97 0.12 2.33* 2.42 0.04 0.55 1.67 0.10 1.15 2.07 0.01 0.09 2.82 0.06 0.70 3.72 -0.18 -2.07 1.35 0.06 2.12 2.12 0.08 1.58 5.84 -0.08 -0.76 3.58 -0.16 -2.34* 1.33 0.05 1.24 2.20 -0.03 -0.85 3.12 0.27 3.36** 2.80 -0.03 -0.09 7.80 0.13 1.76 5.54 -0.09 -0.42 2.10 0.19 3.61** 3.00 -0.27 -1.54 3.82 0.01 0.18 3.29 -0.06 -1.02 3.20 0.36 1.82 2.30 -0.29 -1.09 3.94 0.03 1.33 6.48 -0.09 -1.12 9.27 0.36 2.31* 5.40 0.15 1.23 1.72 0.06 1.16 5.59 -0.22 -1.31 1.83 0.03 0.87 3.72 0.04 0.91 3.16 0.24 3.11* 2.48 -0.02 -0.24 3.78 0.27 6.20** 3.89 -0.03 -0.58 2.27 0.10 1.26 2.52 -0.05 -0.57 11.02 -0.16 -1.32 8.35 -0.02 -0.29 4.13 0.07 1.03 3.21 -0.04 -0.54 2.74 0.16 3.11* 3.38 0.07 0.77 6.14 0.12 1.24 5.12 0.23 2.49* 1.51 0.02 0.47 4.02 -0.08 -1.00 2.64 -0.03 -0.43 3.07 -0.06 -0.67 2.96 0.16 2.15 3.61 0.12 1.01 4.27 0.18 2.30* 3.21 0.01 0.16 4.26 0.08 1.84 5.22 -0.05 -1.52 1.11 0.12 4.76** 3.88 0.06 0.65 2.00 -0.01 -0.15 3.33 -0.18 -1.65 2.33 0.06 1.34 3.50 -0.18 -2.89* 1.92 0.07 2.14 5.81 0.02 0.10 1.64 0.15 2.20 3.63 0.12 1.00 1.12 tr. 0.02 3.88 -0.12 -0.83 3.81 0.20 2.40 6.58 0.09 0.74 2.39 0.10 2.52* 5.39 0.02 0.22 2.01 0.12 1.25 4.86 -0.32 -1.10 4.62 0.25 6.14** 7.12 0.10 1.12 1.41 0.11 2.64* 3.19 0.07 0.67 1.78 0.16 4.83** 3.88 -tr. -0.02 2.49 0.20 2.59* 4.55 0.05 0.31 3.92 0.25 3.99** 4.83 0.06 0.94 2.57 0.18 2.39* 3.48 0.12 1.34 4.41 0.07 1.12 5.84 0.01 -0.12 3.87 0.18 5.28** 6.05 0.04 0.68 3.78 0.09 3.15* 5.68 -0.04 -1.38 3.26 0.10 1.46 5.65 -0.02 -0.09 Atlantic Connecticut Delaware Florida Georgia Maine Maryland Massachusetts New Hampshire New Jersey New York North Carolina Pennsylvania Rhode Island South Carolina Vermont Virginia West Virginia Subtotal Mississippi Alabama Arkansas Illinois Indiana Iowa Kentucky Louisiana Michigan Minnesota Mississippi Missouri Ohio Tennessee Wisconsin Subtotal Central Colorado Kansas Nebraska North Dakota Oklahoma South Dakota Texas Subtotal Pacific Arizona California Colorado Idaho Nevada Oregon Utah Washington Subtotal Total Alaska (1965-75) "All years weighted equally in regression calculations. b*Indicates an average annual change significantly different from zero at the 95% confidence level; fidence level. denote the 99% con- 10 Table 6. Statistical evidence of long-term trends in the hunting of American coot by waterfowl hunters in each State during the period 1964-75. Percentage bagging coot Average coot bag Average Average Period annual t Period annual t Flyway and State average* change valueb average* change valueb Atlantic Connecticut 3.38 -0.02 -0.16 2.78 -0.05 -1.31 Delaware 3.54 -0.09 -0.55 5.01 0.25 1.11 Dist. of Columbia 2.68 -0.20 -2.16 3.23 -0.59 -2.01 Florida 21.65 -0.29 -1.15 9.82 0.07 0.68 Georgia 6.71 -0.07 -0.47 5.41 0.08 0.61 Maine 5.32 -0.21 -1.85 5.88 -0.06 -0.34 Maryland 2.04 0.10 0.13 4.10 -0.33 -2.08 Massachusetts 3.97 -0.07 -0.60 3.83 -0.15 -1.17 New Hampshire 2.45 -0.13 -0.97 2.68 -0.12 -0.90 New Jersey 5.25 -0.13 -1.14 3.60 -0.03 -0.53 New York 3.95 -0.13 -1.10 3.48 -0.11 -1.90 North Carolina 11.29 0.43 1.69 7.55 0.15 0.59 Pennsylvania 5.73 -0.09 -0.51 3.45 -0.06 -0.75 Rhode Island 5.19 -0.32 -1.79 3.21 0.08 0.67 South Carolina 10.76 -0.04 -0.22 5.74 -0.13 -1.62 Vermont 2.21 -0.13 -1.76 2.62 0.10 1.01 Virginia 9.42 0.88 3.65" 10.66 0.71 2.36* West Virginia 7.94 -0.08 -0.21 3.60 -0.33 -1.64 Subtotal 6.60 -0.10 -0.81 6.38 0.02 0.45 Mississippi Alabama 13.32 -0.17 -0.62 7.65 0.29 1.37 Arkansas 3.69 0.01 0.11 4.86 0.15 1.16 Illinois 8.13 -0.17 -0.77 5.19 0.06 0.81 Indiana 10.50 -0.06 -0.22 4.95 0.08 0.63 Iowa 6.46 0.05 0.35 4.12 -0.07 -1.14 Kentucky 5.09 0.01 0.04 3.79 0.09 0.67 Louisiana 16.81 0.45 1.34 17.85 -0.51 -1.14 Michigan 11.60 -0.17 -0.78 4.02 -0.01 -0.14 Minnesota 10.28 0.03 0.29 5.33 -0.05 -0.98 Mississippi 6.29 0.02 0.10 6.75 0.11 0.60 Missouri 4.17 0.12 1.59 3.66 0.15 1.73 Ohio 10.97 -0.10 -0.33 4.10 tr. 0.04 Tennessee 8.54 0.25 1.05 4.83 -0.21 -1.31 Wisconsin 13.17 -0.23 -1.17 7.36 -0.05 -0.55 Subtotal 10.44 -0.03 -0.21 7.92 -0.06 -0.71 Central Colorado 3.06 0.09 1.52 2.81 -0.02 -0.25 Kansas 5.35 0.05 0.66 3.69 0.04 0.55 Montana 3.90 0.09 1.31 3.01 0.21 3.25** Nebraska 4.40 0.25 2.31* 3.23 -tr. -0.01 New Mexico 6.02 0.26 1.50 3.24 0.12 1.31 North Dakota 5.12 0.09 0.62 5.31 -0.07 -0.35 Oklahoma 6.37 0.19 1.23 4.27 0.21 1.37 South Dakota 4.70 0.25 2.91* 4.56 0.09 0.92 Texas 5.51 0.26 4.52** 4.12 0.07 1.09 Wyoming 4.33 0.19 1.41 2.99 0.22 1.58 Subtotal 5.05 0.18 2.78* 4.02 0.04 1.55 Pacific Arizona 8.55 0.17 1.71 5.25 0.15 0.65 California 12.29 0.22 2.51* 9.13 0.04 0.23 Colorado 4.38 0.19 1.82 3.55 0.11 0.76 Idaho 5.06 -tr. -0.01 4.96 -0.01 -0.14 Montana 4.07 -0.02 -0.24 5.23 -0.03 -0.07 Nevada 5.87 0.12 0.97 6.57 0.08 0.55 New Mexico 6.40 -0.28 -1.14 3.29 -0.06 -0.32 Oregon 5.84 0.30 4.16** 5.68 0.15 1.35 Utah 8.07 0.38 2.71* 4.54 0.02 0.41 Washington 5.06 -0.06 -0.90 4.62 -0.10 -1.23 Wyoming 2.84 0.09 0.86 4.43 0.19 0.71 Subtotal 8.34 0.13 2.34* 7.41 0.01 0.08 Total 8.21 0.01 0.14 7.13 -0.04 -0.56 Alaska (1965-75) 1.66 -0.03 -0.84 4.24 -0.30 -1.97 •All years weighted equally in regression calculations. ■"Indicates an average annual change significantly different from zero at the 95% confidence level; "denote the 99% con- fidence level. 11 hunters report bagging about 22 gallinules per thou- sand coots. Thus, waterfowl hunters could be misiden- tifying as many as 12% of the gallinules they take as coots. An average of about 4,600 (0.2%) of the duck stamp buyers in the United States during 1964-75 hunted gallinules, taking an average of 26,400 gallinules (5.8 per hunter) annually (Tables A-15, A-16). About 31% of the gallinule hunters and 50% of the harvest by waterfowl hunters were attributed to Louisiana. Statistically significant long-term trends were detected in both percentage hunting (decrease in Atlantic Flyway, increases in Mississippi and Central flyways) and average bag (increase in Mississippi Flyway) at the flyway level, but there was no evidence of nationwide trends (Table 2). American Coot Coot hunting activity and harvest are undoubtedly overestimated in the New England-New York coastal areas where scoters (Melanitta spp.) are routinely referred to as coots and reported as such to an unknown degree by waterfowl hunters. However, this overestimate involves comparatively few birds (Table A-17) in areas where the American coot is relatively uncommon (Sanderson 1977:133). On the average, each year during 1964-75, about 8.2% (164,000) of the duck stamp buyers in the United States hunted coots successfully. Probably very few (assume 5%) who hunted coots were unsuccessful, so, for purposes of comparing hunting pressure placed on coots with that placed on other migratory game birds by duck stamp buyers, an estimate of 172,600 hunters (8.7%) seems reasonable. The harvest by all waterfowl hunters, including those under 16 years old and hunting without a duck stamp, averaged 1,175,000 coots (almost 7 per hunter) annually during the same period. Allowing for survey differences, a similar range in coot hunting activity and success was experienced during 1952-63 (Tables A-18, A-19). Over 58% of the harvest occurred in the Mississippi Flyway; Louisiana accounted for over one-fourth of the harvest in the United States (Table A-17). California and Wisconsin together accounted for another 25%. No long-term trends were evident in average bag, but average in- creases of 3.6% in the Central Flyway and 1.6% in the Pacific Flyway per year in the percentage of waterfowl hunters hunting coots during 1964-75 were statis- tically significant (Table 6). Selected Comparisons with Other Surveys Some of the results of this survey are compared, at the State level, with those of several other surveys (Table 7). The predominance of high correlations is encouraging and perhaps surprising, particularly since those for total hunters and total bag are as prominent as those for average bag. The latter, being less influ- enced by variability in duck stamp sales, was expected to be a better measure of the actual situation. The reader is reminded that a high correlation between surveys does not imply that the surveys yield the same results, only that the results tend to differ by a con- stant amount or factor which can be large or small, positive or negative, depending on the species. These high correlations look impressive but may be somewhat misleading. Often very wide ranges are involved (e.g., Nevada with 70,100 and California with 1,210,400 mourning doves bagged by duck stamp buyers in 1972), and such data may show substan- tial departures from a normal distribution. Even comparatively insensitive surveys might agree well enough in such situations to produce high correlations. Thus, the degree of agreement between surveys should be presumed to be exaggerated to some extent, partic- ularly when used for comparisons among States. Com- parisons among seasons tend to yield somewhat lower, though still often significant, correlations, and these results are probably more realistic and useful. The waterfowl hunter survey has the advantage that it is uniformly applied in all States whereas surveys conducted by individual States may lack this uniform- ity. The low correlations with band-tailed pigeon data from Oregon and Arizona, contrasting with the gen- erally high correlations found elsewhere, may indicate that the surveys of Oregon and Arizona were less effi- cient than those of the other States. Different survey methods can produce markedly different results. For example, a special mail questionnaire survey of white- winged dove hunters in Texas in 1976, based on an effi- cient sampling frame of white-winged dove stamp pur- chasers, produced activity and harvest estimates about 2.5 times larger than those produced by the tra- ditional check station survey (J. H. Dunks, Texas Parks and Wildlife Department, Austin, mimeo- graphed report on white-winged dove harvest in Texas dated 21 March 1977). Measurements of trends and other relationships may be much less affected, however. Simple questionnaire design changes can be important, too. With the elimination of the waterfowl calendar (for reporting daily success) from the Federal questionnaire in 1969 after 2 years of testing, it was necessary to reduce subsequent estimates of other migratory game bird hunting activity by 4%, success- ful coot hunters by 10%, and coot harvest by 16% to maintain comparability with results obtained in pre- vious years (E. M. Martin, Office of Migratory Bird Management, Laurel, Maryland, unpublished report on changes in the Federal questionnaire survey dated 12 Table 7. Correlation of estimates of total hunters, total bag, and average bag of migratory game birds from Federal questionnaire (this report) with estimates from other sources." Source Season(s) Category Correlation coefficient for: Total Total Average Species and State or area of datab used testedc hunters bag bag White-winged dove Lower Rio Grande (Texas) 1 1966-75 Seasons (10) — 0.60 — Arizona 2 1966-75 Seasons (10) — 0.66* 0.91* Band-tailed pigeon California 2 1966-75 Seasons (10) 0.76* 0.92** 0.87** Oregon 2 1966-75 Seasons (10) -0.28 trace 0.43 Washington 2 1966-75 Seasons (10) 0.74* 0.91** 0.75* Arizona 2 1968-75 Seasons (8) -0.37 0.31 0.57 New Mexico 3 1968-75 Seasons (8) 0.82* 0.64 0.44 Colorado 4 1970-75 Seasons (6) 0.92** 0.93** 0.76 Utah 2,5 1970-75 Seasons (6) 0.71 0.58 0.48 Mourning dove Arizona 2 1966-75 Seasons (10) 0.75* 0.70* 0.27 California 2 1966-75 Seasons (10) 0.68* 0.43 0.71** Idaho 2 1966-75 Seasons (10) 0.87** 0.84** 0.27 Nevada 2 1966-75 Seasons (10) 0.69* 0.69* 0.75** Oregon 2 1966-75 Seasons (10) 0.65* 0.57 0.78 Utah 2 1966-75 Seasons (10) 0.71* 0.73** 0.82** Washington 2 1966-75 Seasons (10) 0.38 0.76** 0.76** Management Units Eastern 6 1971 or 72 States (10) 0.55 0.69* 0.91** Central 6 1971 or 72 States (7) 0.97** 0.99** 0.91** Western 6 1971 or 72 States (7) 0.96** 0.97** 0.98** Total States (24) 0.86** 0.87** 0.94** Sandhill crane All states except Alaska 7 1975 States (8) 0.98** 0.99** 0.96** aAsterisks indicate significant correlation: * at the 95% confidence level and ** at the 99% level. bSource of data for comparisons with waterfowl hunter data from questionnaires: 1. Sanderson (1977:257) and telephone conversation with J. H. Dunks, Texas Parks and Wildlife Department, Austin. 2. Minutes of the Western Migratory Upland Game Bird Technical Committee, Report 15, dated June 1976. 3. Braun et al. (1975:14) and telephone conversation with J. L. Sands, New Mexico Department of Game and Fish, Albuquerque. 4. Braun et al. (1975:14) and telephone conversation with C. E. Braun, Colorado Division of Wildlife, Fort Collins. 5. Braun et al. (1975:14) and minutes of Western Migratory Upland Game Bird Technical Committee, Report 14, dated June 1975. 6. Sanderson (1977:286). 7. M. F. Sorensen and H. M. Reeves, Office of Migratory Bird Management, Laurel, Md., administrative report on sandhill crane hunting dated 9 July 1976. cSample sizes in parentheses. 17 December 1970). It is apparently coincidence that this design change and the significant upward trends in the percentages hunting several species occurred at about the same time, in part because, if they were related, then the number of significant decreases has also been underestimated, probably a much less likely occurrence. The results of some general comparisons of Federal questionnaire and wing collection surveys are shown in Table 8. Duck survey data are shown as a standard with which to compare woodcock survey data. Whereas the agreement between the two surveys appears increasingly good between 1971 and 1975 for ducks, it has remained generally poor for woodcock, well below the point of real usefulness. Although the waterfowl-hunter sampling frame is acknowledged to be least complete for woodcock, dove, and pigeon hunters, the generally high correlations obtained with dove and pigeon data (Table 7) led to the expectation of similar results for the woodcock. Possibly regional differences are a factor. Somewhat poorer agreement was found in dove data from the Eastern Management Unit where the lowest ratios of duck stamp buyers to hunters occur (Table A-l), and where nearly all wood- 13 Table 8. State-level comparisons of Federal questionnaire and wing survey estimates of the average seasonal success of duck and woodcock hunters. Flyway Number of States Correlation coefficient3 for x bag Species 1971-72 1972-73 1973-74 1974-75 1975-76 Ducks Atlantic 18 0.48* 0.71** 0.80** 0.82** 0.87** Mississippi 14 0.81** 0.88** 0.88** 0.93** 0.91** Central 10 -0.17 0.12 0.30 0.70* 0.78** Pacific 11 0.14 0.84** 0.72* 0.85** 0.82** U.S. 54 0.57** 0.81** 0.81** 0.87** 0.86** American woodcock Atlantic Mississippi and 17 0.71** 0.51* 0.42 0.39 0.33 Central 14 to 17 0.24 0.52* 0.35 0.41 0.43 Total 31 to 34 0.54** 0.53** 0.45** 0.44** 0.37* a*Indicates a significant correlation at the 95% confidence level; **denotes the 99% level. cock hunting activity occurs. Secondly, the incidence of mourning dove hunters in the States in which this bird has been hunted is much higher than the compara- ble figure for woodcock (40% vs. 12%), resulting in a much larger sample of mourning dove hunters. Thirdly, these wing surveys sample select groups of high success hunters whereas the questionnaire survey attempts to randomly sample all waterfowl hunters. Although adequate for most estimates related to waterfowl hunting, it appears that one or both of these sampling frames yield poor results for woodcock hunting. Although some statistical tests were applied to data at flyway, management unit, and U. S. levels (obtained by adding various State estimates), it should be recog- nized that an additional source of error is contained in such figures because of the inability in this survey to either sample the hunters of these various species pro- portionately or to apply appropriate weighting or expansion factors for making such combined-area esti- mates. General indications are probably still valid but it may be better, when applying the results in manage- ment programs, to place more emphasis on the figures for individual States. Undoubtedly, additional infor- mation of better quality on changes and trends can be extracted from the present survey with a more sophis- ticated statistical approach, but it is also apparent that, as concluded by MacDonald and Martin (1971:8), a better sampling frame for measuring the harvest of migratory game birds other than waterfowl is still needed. References Benning, D. S., M. M. Smith, and S. L. Rhoades. 1975. Waterfowl status report, 1973. U. S. Fish Wildl. Serv., Spec. Sci. Rep. -Wildl. 188. 105 pp. Braun, C. E., D. E. Brown, J. C. Peterson, and T. P. Zapatka. 1975. Results of the four corners cooperative band-tailed pigeon investigation. U. S. Fish Wildl. Serv.. Resour. Publ. 126.20 pp. Clark, E. R. 1972. Woodcock status report, 1971. U. S. Fish Wildl. Serv., Spec. Sci. Rep.-Wildl. 153. 47 pp. MacDonald, D., and E. Martin. 1971. Trends in harvest of migratory game birds other than waterfowl, 1964-65 to 1968-69. U. S. Fish Wildl. Serv., Spec. Sci. Rep.-Wildl. 142.29 pp. Ruos, J. L., and R. E. Tomlinson. 1968. Mourning dove status report, 1966. U. S. Fish Wildl. Serv., Spec. Sci. Rep.-Wildl. 115.49 pp. Sanderson, G. C, ed. 1977. Management of migratory shore and upland game birds in North America. International Association of Fish and Wildlife Agencies, Washington, D. C. 358 pp. 15 Appendix A The 1964-75 survey results, the subject of various analyses in this report, are summarized here at State, flyway or management unit, and national levels. Other investigators may wish to carry out additional anal- yses after referring to the preceding sections for back- ground information on the survey and the limitations of its results. 16 Table A-l . Summary of average duck stamp and hunting license sales and relationships between them at State, management unit, and flyway levels: 1964-75. 1964-75 average: Percentage buying duck stamps Flyway, Duck Hunting 1964-75 Average Management Unit, stamp license annual 5 b and State sales sales average change value Atlantic Flyway Eastern Management Unit Connecticut 12,262 76,875 15.76 0.43 4.33** Delaware 11,242 27,112 41.40 1.50 4.49** Florida 29,748 242,001 12.35 -0.26 -1.56 Georgia 11,965 323,265 3.72 0.07 1.14 Maine 15,518 203,998 7.58 0.24 4.34** Maryland 31,774 186,209 17.02 0.58 4.63** Massachusetts 24,193 121,761 20.08 0.80 4.34** New Hampshire 8,421 87,739 9.74 0.79 13.24** New Jersey 32,425 184,310 17.49 0.46 4.53** New York 88,414 728,452 12.19 0.58 2.93* North Carolina 25,780 418,782 6.18 0.15 2.00 Pennsylvania 61,977 1,109,284 5.53 0.17 2.14 Rhode Island 3,167 16,113 19.55 0.73 5.38** South Carolina 18,500 200,706 9.26 0.28 2.78* Vermont 6,886 139,669 4.93 0.31 7.20** Virginia 19,049 410,088 4.66 -0.01 -0.44 West Virginia 1,834 253,786 0.72 tr. 0.46 Mississippi Flyway Eastern Management Unit Alabama 14,247 339,408 4.20 -tr. -0.06 I llinois 71,007 470,355 15.12 0.34 2.01 Indiana 23,820 471,342 5.11 0.21 2.97* Kentucky 9,619 271,295 3.52 0.10 3.78** Louisiana 105,379 323,977 32.47 0.12 0.41 Michigan 98,476 896,244 11.06 0.34 2.51* Mississippi 22,025 288,926 7.59 0.20 2.01 Ohio 35,317 558,113 6.35 0.22 2.84* Tennessee 26,250 449,714 5.80 0.12 1.94 Wisconsin 125,821 624,214 20.16 0.15 0.71 Central Management Unit Arkansas 42,896 300,134 14.11 0.49 2.27* Iowa 53,870 327,737 16.36 0.35 2.41* Minnesota 149,520 418,497 36.33 0.51 0.79 Missouri 50,729 447,724 11.28 0.24 3.11* Central Flyway Central Management Unit Colorado (East)c 36,503 284,859 12.84 0.58 3.31** Kansas 50,735 220,705 22.72 1.16 6.77** Montana (East) Nebraska 40,959 195,586 20.97 1.06 4.72** New Mexico (East) 6,332 118,640 5.38 -0.04 -0.63 North Dakota 46,020 89,337 51.39 0.83 2.63* Oklahoma 29,896 237,170 12.65 0.23 1.29 South Dakota 40,263 156,892 25.82 0.16 0.68 Texas 115,196 745,854 15.42 0.04 0.30 Wyoming (East) 7,231 155,686 4.64 0.20 3.21** Pacific Flyway Central Management Unit Colorado (West) Montana (West) 26,646 186,846 14.46 -0.23 -1.55 New Mexico (West) Wyoming (West)c Western Management Unit Arizona 11,363 156,994 7.21 0.12 1.34 California 156,155 704,288 22.19 0.33 2.84* Idaho 30,444 210,393 14.49 0.11 1.26 Nevada 12,671 63,061 20.11 0.19 1.13 Oregon 51,585 349,733 14.76 0.25 2.14 Utah 33,698 218,195 15.47 0.21 1.48 Washington 72,402 325,053 22.29 0.14 0.92 Alaska 13,085 56,602 23.42 -0.04 -0.15 Management Unit Totals Eastern 935,115 9,423,741 9.90 0.23 2.51* Central 696,797 3,885,665 17.86 0.30 2.05 Western 368,319 2,027,718 18.15 0.18 1.85 Flyway Totals Atlantic 403,154 4,730,150 8.49 0.23 2.87* Mississippi 828,976 6,187,682 13.38 0.26 1.98 Central 373,135 2,204,728 16.82 0.37 2.97* Pacific 394,965 2,214,564 17.82 0.13 1.40 U. S. Total (with Alaska) 2,013,315 15,393,725 13.04 0.25 2.38* ? All years weighted equally in regression calculations. Asterisks indicate annual change significantly different from zero--* at the 95% confidence level and ** at the 99* confidence level. c Hunting license figures not separable by flyway; data for these States appear in the flyway where most stamp sales occurred. Table A-2. White-winged dove hunting activity and success by hunters purchasing duck stamps: 1966-75. 17 1966-67 1967-68 1968-69 1969-70 1970-71 1971-72 1972-73 1973-74 1974-75 1975-76 Average New Mexico Number hunting 330 350 310 320 580 440 320 290 440 670 400 Percent hunting 6.0 5.9 4.9 5.6 8.8 6.0 5.0 4.7 5.9 8.1 6.2 Total bag 4,400 5,600 6,400 3,600 4,200 4,400 3,800 3,500 3,900 11,100 5,100 Average bag 13.3 15.7 20.7 11.3 7.2 10.1 11.9 12.1 9.0 16.7 12.6 Texas Number hunting 11,400 9,900 8,900 7,200 9,100 9,400 9,700 7,200 8.100 9,400 9,000 Percent hunting 11.5 8.9 9.4 6.3 6.3 6.9 7.6 6.2 6.7 7.2 7.5 Total bag 221,600 182,400 106.500 98,200 139,900 85,400 132,100 103,100 130,900 147,100 134,700 Average bag 19.4 18.4 12.0 13.6 15.4 9.1 13.6 14.4 16.2 15.6 14.9 Arizona Number hunting 4,100 4,600 4,000 4,700 4,500 3,900 4,900 4,400 5,000 5,800 4,600 Percent hunting 47.0 44.5 39.9 37.1 31.8 25.6 41.9 39.9 37.5 50.5 38.6 Total bag 86,800 114,700 86,400 57,000 65,900 41,700 64,700 61,500 55,300 81,400 71 ,400 Average bag 21.4 25.2 21.2 12.2 14.7 10.6 13.1 14.0 11.1 14.0 15.6 Cal ifornia Number hunting 10,600 10,700 7,500 10.700 11,500 1 1 , 200 6,900 5,600 9,500 9,200 9,300 Percent hunting 7.0 7.1 4.7 6.2 6.2 6.6 4.6 4.0 6.7 6.7 6.0 Total bag 101,000 155,100 68,800 102,500 129,000 114,600 69,100 69,700 80,600 97,500 98,800 Average bag 9.6 14.4 9.2 9.6 11.2 10.2 10.1 12.4 8.5 10.6 10.6 Nevada Number hunting 360 530 370 310 500 340 280 220 290 350 360 Percent hunting 3.0 4.2 3.0 2.4 3.5 2.3 2.2 1.6 2.5 2.6 2.7 Total bag 3,900 4,300 2,000 4,200 3,000 5,000 4,200 1.500 3,000 3,500 3,500 Average bag 10.9 8.1 5.5 13.5 6.0 14.6 15.0 7.0 10.4 10.0 9.8 TOTAL Number hunting 26,800 26,100 21,100 23.200 26,200 25,300 22,100 17,700 23,300 25,500 23,700 Percent hunting 9.7 9.0 7.4 7.3 7.1 7.3 7.2 6.1 7.9 8.4 7.7 Total bag 417.700 462,100 269.100 265,600 342,000 251,100 274,000 239,400 273,700 340,600 313,500 Average bag 15.6 17.7 12.8 11.4 13.1 9.9 12.4 13.5 11.8 13.4 13.2 Table A-3. Band-tailed pigeon hunting activity and success by hunters purchasing duck stamps: 1966-75. 1966-67 1967-68 1968-69 1969-70 1970-71 1971-72 1972-73 1973-74 1974-75 1975-76 Average California Number hunting 11,400 10 ,000 11,100 9,600 9,900 9,700 13,500 8,100 8,800 6,400 9,900 Percent hunting 7.5 6.6 6.9 5.5 5.3 5.7 9.1 5.7 6.3 4.6 6.3 Total bag 109,000 64 ,600 97,400 70,900 82,700 86,400 142,100 67.000 74,600 39,300 85,400 Average bag 9.6 8.5 8.8 7.4 8.3 9.0 10.5 8.2 8.4 6.1 8.7 Oregon Number hunting 6,000 7 ,000 5,400 7,600 6,100 8,600 8,800 7,200 6,200 5,300 6,800 Percent hunting 12.3 14.6 11.3 13.8 10.0 14.8 17.1 14.2 12.1 9.8 13.0 Total bag 47,800 59 ,400 44,400 65,600 43,100 75,300 66,900 58,100 45.500 37,900 54,400 Average bag 8.0 _ 8.5 8.2 8.7 7.1 8.8 7.6 8.1 7.4 7.1 8.0 Washington Number hunting 9,200 6 ,100 8,000 8,100 8,600 9,400 7,000 7,500 6,400 5,800 7,600 Percent hunting 13.4 8.6 11.0 10.1 10.3 12.4 10.1 10.5 9.3 8.1 10.4 Total bag 71,000 45 ,400 77,000 54,600 59,900 65,700 38,900 47,500 42 , 1 00 25,100 32,700 Average bag 7.7 7.5 9.7 6.7 7.0 7.0 5.6 6.3 6.5 4.3 6.9 COASTAL STATE SUBTOTAL Number hunting 26,600 23 ,000 24,500 25,300 24,600 27,600 29,400 22,800 21,500 17,600 24,300 Percent hunting 9.9 8.5 8.7 8.2 7.4 9.1 10.9 8.6 8.2 6.7 8.6 Total bag 227,800 189 ,400 218,800 191.100 185,700 227,400 247.900 172,500 162,200 102.200 192,500 Average bag 8.6 8.2 8.9 7.6 7.5 8.2 8.4 7.6 7.6 5.8 7.9 Arizona Number hunting I' 600 510 450 770 510 330 640 640 440 Percent hunting 5.9 4.0 3.2 5.0 4.3 3.0 4.8 5.5 4.5 Total bag 2,200 790 1,660 2,280 540 1.620 2,140 2,440 1,370 Average bag 3.7 1.6 3.7 2.9 1.1 4.9 3.4 3.8 3.1 New Mexico Number hunting 120 60 210 360 320 250 360 350 200 Percent hunting 1.8 1.1 3.2 4.9 4.9 4.1 4.8 4.2 3.7 Total bag 270 270 340 2,050 1,290 790 830 1.970 780 Average bag 2.3 4.2 1.6 5.7 4.1 3.2 2.3 5.7 3.9 Colorado Number hunting 380 850 670 590 810 540 380 Percent hunting 0.9 1.8 1.6 1.4 1.9 1.2 1.5 Total bag 1,240 5,090 2,490 1,110 2,770 1.250 1,400 Average bag 3.3 6.0 3.7 1.9 3.4 2.3 3.6 Utah Number hunting 110 110 350 60 230 120 100 Percent hunting 0.3 0.3 1.1 0.2 0.6 0.3 0.5 Total bag 540 200 910 90 570 60 240 Average bag 5.0 1.8 2.6 1.3 2.4 0.5 2.4 INLAND STATE SUBTOTAL Number hunting 720 570 1,140 2,100 1,850 1,230 2,040 1 ,640 1,410 Percent hunting 4.3 3.1 1.2 2.0 2.0 1.3 2.0 1.6 1.8 Total bag Average bag 2,470 1,060 3,770 9,620 5,230 3,610 6,310 5,720 4,720 3.5 1.9 3.3 4.6 2.8 2.9 3.1 3.5 3.3 TOTAL Number hunting 26,600 23 ,000 25,200 25.900 25,700 29,700 31,200 24.100 23,500 19,200 25,400 Percent hunting 9.9 8.5 8.5 7.9 6.0 7.2 8.6 6.7 6.5 5.2 7.4 Total bag 227.800 189,400 221.300 192,100 189,500 237,000 253.200 176,100 168,500 107,900 196,300 Average bag 8.6 8.2 8.8 7.4 7.4 8.0 8.1 7.3 7.2 5.6 7.7 a/ Blank space indicates that the season was not open on band-tailed pigeons. 18 imcNW'-iri'-inoowiNajo a}(^(\jir>oiic t— <£siOWff.OMrr»a;c 1 iO CO r*> I — CT> O O O i"n O in t .— cm c^wninniflr .— VT> r- O CO »— 10 00 in o"i r>. or .— .— .— Cft IffiiCiOUJOU ■> CT> (*> CO CO <"1 f i CD (*1 tc WCO O (M in Oi p — >ON«iDicaiO OC^COr TCI O CT o ro Ps s O ^ -«'incr>r-.*3'OOr-.u ouiwicmio- coi- - co cfi co co IN coOCMCHf lO IDr-fl-lOr-lflCDO a> w • — in co m m . — m co .ooisontof i co p-mooinu looM^otn o "i cr» CO CO Oi iO CT* <■" J Ol C\J *3- CO c n \o i — fn in cri"! 3^ O cf> c ioovio nuii — ionico r co a- •— cm — u i •— co o - lO CT* C\J CH 10 r — CO gONMiOi — r«. 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Sandhill crane hunting activity and success by hunters purchasing duck stamps: 1966-75. 1966 -67 1967-68 1968-69 1969-70 1970-71 1971-72 1972-73 1973-74 1974-75 1975-76 Average Alaska Number hunting 590 310 290 340 470 250 290 540 560 350 400 Percent hunting 5.6 3.1 2.4 2.6 3.6 1.7 2.0 3.2 3.5 2.3 2.9 Total bag 460 440 340 380 720 160 100 1,440 1,050 290 540 Average bag 0.8 1.4 \ 1.2 1.1 1.5 0.6 0.3 2.7 1.9 0.8 1.4 Texas Number hunting -£/ 74C 1,110 1,630 1,100 1,630 1,390 1,570 1,800 2,110 1,450 Percent hunting - o.; ' 1.2 1.4 0.8 1.2 1.1 1.4 1.5 1.6 1 .2 Total bag - 1,390 1,110 3,860 2,560 4,130 2,270 7,500 4,700 7,010 3,840 Average bag - l.S 1.0 2.4 2.3 2.5 1.6 4.8 2.6 3.3 2.6 New Mexico Number hunting - - 380 210 600 710 130 290 180 410 360 Percent hunting - - 6.0 3.8 9.1 9.8 2.0 4.8 2.4 5.6 5.4 Total bag - - 1,430 430 1,120 1,260 180 420 220 710 720 Average bag - - 3.7 2.0 1.9 1.8 1.4 1.4 1.2 1.7 2.0 Colorado Number hunting b/ - no 330 410 750 360 310 670 350 410 Percent hunting - 0.3 0.9 1.0 1.6 0.9 0.7 1.6 0.9 1.0 Total bag - 0 230 590 500 370 610 760 80 390 Average bag - 0 0.7 1.4 0.7 1.1 1.9 1.1 0.2 1.0 North Dakota Number hunting 270 820 840 330 520 420 1,220 2,600 880 Percent hunting 0.7 1.9 1.7 0.6 1.0 0.8 2.2 4.4 1.8 Total bag 120 370 520 130 890 1,130 1,530 3,030 960 Average bag 0.4 0.4 0.6 0.4 1.7 2.7 1.3 1.2 1.1 Oklahoma Number hunting 130 170 160 120 80 450 490 410 250 Percent hunting 0.5 0.6 0.5 0.3 0.2 1.4 1.5 1.2 0.8 Total bag 30 no 80 120 0 420 460 1,080 290 Average bag 0.3 0.7 0.5 1.0 0 0.9 0.9 2.7 1.2 South Dakota Number hunting 0 0 20 90 250 30 220 0 80 Percent hunting 0 0 0.1 0.2 0.5 0.1 0.5 0 0.2 Total bag 0 0 50 90 120 90 380 0 90 Average bag -- — 2.0 1.0 0.5 3.0 1.7 — 1.2 Montana Number hunting 30 100 250 80 120 Percent hunting 0.1 0.4 0.9 0.3 0.4 Total bag 0 20 130 0 40 Average bag 0 0.2 0.5 0 0.3 Wyomi ng Number hunting 20 70 10 130 60 Percent hunting 0.2 0.8 0.1 1.8 0.7 Total bag 0 80 30 0 30 Average bag 0 1.2 2.1 0 0.5 TOTAL Number hunting ■1 2,300 3,500 3,600 3,900 3,000 3,800 5,400 6,400 4,000 Percent hunting Survey 0.9 1.2 1.1 1.1 0.9 1.1 1.5 1.7 1.2 Total bag incomplete 3,000 5,400 5,600 6,400 3,900 11,700 9,200 12,200 7,200 Average bag 1.3 1.5 1.6 1.7 1.3 3.1 1.7 1.9 1.8 a/ Dashes indicate that no data are available as questionnaire used did not include crane question, b/ Blank space indicates that the season was not open on cranes. 25 ooooooooooooooo • o o o OOOOOOOOOOOOOOO OOO ■ O O 'OO o ooooooooooooooocooo ooooooooooooooc .1 o u-i uj o <& tvicjoicoi—r g <£> <7> lO 0< 7IMr Of 300 OOOOO O OOOOOOOOOOOOOOO OOOOOOOOO o OOOOOOOOOOOOOOC OOO OOOOC OOOOOOOOOOOOOOOOOOO O c> f) f> if> o \o ^ c^j cr> cnj r^MCA CO n •£> cm O er» OOOOOOOOOOOOOOO OOO OOOOO o OOOOOOOOOOOOOOO OOOOOOOOO o O^ voiO cor-.r-.eO'S-.— co«3-r-.o Mcor^^M* r^. a, OOOOOOOOOOOOOOOOOOO CUkOc^O^^-dCT*' — in^^M^r^. ^ co OOOOOOOOOOOOOOOOOOO OOOOOOOOOOOOOOO OOOOO O O O o OOOOOOOOOOOOOOO OOOOO o o o o OOOOOOOOOOOOOOO OOOOO O O O O ^D 0"< i^ajnncoiDMOio r">.— o rOi-nmwiowOMiO'- -) co t— ro r-i c r l-> f> tsj CO •: ■ o o • o ■ o ■O -O ■ o ■ - o o o - o o ■ o ■ ■ o CO ^O ii"> ^ ' — ^ ■ o ■ • o o • o • ■ o "loo • . o ■ ■ o 10 • — i — (^ ui w <» i\no o • o ■ • o o o ■ • o o ■ o • • o u co m en o ■ o • ■ ■ o • vO «3- CO I-. f— t e^i m\Of -lfl(\JOi — , o *-> 't- -^ J u is ■•- f ■- >e u , c c =» >, aiJU-'-cnaj.— .u:£Os-.ci/ia).cO'r- " — CKt-Ui-l-C^I/l k CUM E DlWd. 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O HI i— s a. i— «a: 1— o- c >— C IO r— O 3 ID •r- CTCn in cu OJ (— T3 3 .O C O" ro fO C QJ 1— 4-» qj cn o T3 •+- ID ro O cn 1— I U U QJ S- U in co o 3 r OJ3 U in C ro "O O 33 o^-ocnooi onOcoom IT) . ,— -CO • O vO O <— O O-s CM • (Ti • r— . i— f— r— O - in -vo *m r— • crt • m • O 10 o in o in 0050«J-OU1 *i— »C0 "in o co o en o r^. i— i— CM CM ■— i— CO — •— o> CO ■— o> I— CM t— CM O O O 0> O •— OOOIOOl r- -in ■ 1*^ ■ CTl ■— ■— r— r Or-OlDO" OlDOOOi- r— - \£) -CO • ooioi-oro in ■ o ■ 10 ■ o r-* o co o -a- o o o in o r- rvi • r- -co ■ O ro o i— o CO Oinoojon o • <— • CO - i— CM CO i— CO CO r— r- CO OCOOMOCO o m o <— o r*"> "* ■ n -en • QC0O«O" O O O CO o T*- ro ■ r— - CM . i— CM CM o o o en o m CD O <— O *3" U3 . O IP • O U") O CO O CO OCftO>*OM <* ■ vO • O • 3TD CO ■M o o o >J o u 1 Oi£ C 3 •>- m 3 a. 'i3 (ur-P a. .Q U (O 1- (O 1- •■- i- +j i ZQ.h ^ Q- h- Q 0) cnjD c 3 J _Q O « t- fD O - ^ 1> +J d) +J u J 3 QJ O > O QJ : z a- I— <* h- Q- OVi- t- «J -t-> C C7>J3 C 3 -> -4 U (B t ID U - E t+J ()« l. - 3 CO O > O 4) J Z a. I— «£ I— O- iO +J 0">-0 C. 3 CT>.0 C 3 35 Appendix B Survey results for the 1976-77 season, which became available after work on the earlier data had been largely completed, are summarized in Tables B-l and B-2. It is suspected that the 1976-77 results, and perhaps to a lesser degree the 1975-76 results, were affected by the unavoidable addition of Privacy Act statements on survey forms. A further intrusion into the survey in the form of a prohibition on the proper testing of these changes has thus far frustrated efforts to assess comparability with the results of previous surveys. Response rates for the survey mailing list were depressed by about 30% in 1976 when the Privacy Act statement was implemented for this phase of the survey, and some reduction, though much smaller, in questionnaire response rates likely occurred in both 1975 and 1976. A previous questionnaire change which increased response rates inflated all activity and harvest estimates, particularly those for coots (E. M. Martin, Office of Migratory Bird Manage- ment, Laurel, Maryland, unpublished report on changes in the Federal questionnaire survey dated 17 December 1970). This latest change, by depressing response rates, can thus be expected to have decreased activity and harvest estimates, perhaps most notice- ably for coots. It is hoped that some evaluation can still be made and adjustments incorporated where necessary. In the meantime, the suspicion remains that activity and success figures for 1975 (to a slight degree) and 1976 (to a potentially very significant degree, at least for certain species) may be under- estimates. 36 Table B-l. Mourning dove, white-winged dove, band-tailed pigeon, and sandhill crane hunting activity and success by duck stamp purchasers during the 1976-77 season. Species, Stamp buyers participating % Total Birds harvested Species and State Stamp buyers participating % Total Birds harvested Management Unit, and State Per Total hunter (thousands) Per hunter Total Mourning Dove Eastern Alabama Delaware Florida Georgia Illinois Kentucky Louisiana Maryland Mississippi North Carolina Ohio Pennsylvania Rhode Island South Carolina Tennessee Virginia West Virginia Other States^ Unit Total Central Arkansas Colorado Kansas Missouri Nebraska New Mexico Oklahoma Texas Wyoming Other States3 Unit Total Western Arizona Cal i form' a Idaho Nevada Oregon Utah Washington White-winged dove 61.4 42.7 53.7 63.2 42.9 59.1 32.3 32.3 66.6 59.6 33.2 43.4 31.7 66.0 59.4 51.0 46.3 1.9 26.8 37.0 37.7 59.7 38.5 47.2 60.8 54.9 55.3 25.4 1.5 29.9 65.3 40.8 29.2 38.3 23.4 36.3 18.1 9,400 5,200 16,900 8,500 27,700 8,300 39,500 12,600 19,200 16,500 13,400 30,400 1,100 14,300 15,600 10,000 900 7,800 257,200 21,600 17,100 34,400 22,600 20,300 4,700 18,400 76,600 2,600 4,300 222,500 8,000 52,600 10,200 4,900 12,200 14,700 12,300 Unit Total 32.9 115,100 45.3 20.3 39.2 46.0 21.5 38.1 25.8 17.7 39.2 31.5 19.0 13.8 13.9 35.3 36.0 28.7 16.3 14.6 27.9 30.8 19.3 23.5 24.2 23.3 30.6 30.8 31.2 15.0 12.3 27.0 37.2 18.5 11.2 15.5 13.0 10.4 12.6 16.8 424.3 104.8 663.1 391.8 593.9 316.8 1,017.6 223.9 753.9 518.8 253.5 418.4 15.4 504.4 562.1 286.9 14.7 114.8 7,179.3 663.9 328.5 806.4 545.2 474.7 142.8 567.5 2,388.5 38.6 52.8 6,008.8 299.8 972.3 114.6 76.4 158.2 152.6 155.4 1,929.2 New Mexico Texas Arizona California Nevada Total 4.0 7.4 43.9 4.6 1.8 7.4 Band-tailed pigeon California 4.9 Oregon 11.9 Washington 7.9 Arizona 4.5 New Mexico 1 .2 Colorado 1.1 Utah 0.5 Total 5.4 Sandhill crane Alaska Texas New Mexico Colorado North Dakota Oklahoma South Dakota Montana Wyoming Total 5.5 2.5 6.5 1.1 2.4 0.9 0 0.2 0.9 2.0 310 10,200 5,400 6,000 240 22,200 6,300 6,200 5,400 560 100 480 210 19,300 1,020 3,420 500 510 1,350 300 0 70 100 7,300 2.9 15.5 13.4 11.0 12.3 13.6 5.7 5.6 5.0 4.8 0.9 5.1 3.5 5.4 1.1 1.9 0.5 0.3 0.3 2.4 0.5 2.0 1.3 880 158,600 72,300 65,900 2,910 300,700 36,100 34,900 27,100 2,640 90 2,430 740 104,000 1,080 6,660 250 130 380 730 0 30 190 9,500 Grand Total 29.0 594,800 25.4 15,117.3 aNo open season on mourning dove in State where duck stamp was purchased, and no data are available for reassignment of hunters to State of harvest. 37 o .ooooooooooopoppp ooooooopogopooo ooooooooooo nooooooooooo ouoooooooooooooooo ooooooooooooooo ooooooooooo ggoooooooooo - .— en _ C M r- r- o ■.coe\i*oi/>cncor' 0OfMU">*J-<-"--OJ0 IIOIOOOMDi- o o • o o o c o o o o o oc 3 O O O O O O O O O O C c; r- .— en - o ■ — tNjt\ja)Oco*oio'" » OCOCM (si — i- o o o o o o o .oooooooog • o - .o O i- ■ — O o o i. o oooooooooooooo ooooooooooo O O — OO OO O — — — . ■ -_» -— O-— -O i o i o ■ ■ i o o o 3 o i i wo i o o i co -ooooo • -oooooooo • • o — o j- !_— - o oo 1-1-S soooooooooooo oooooooooo o o — oc o oo ^ii"— — — 5 — 5 3 — o— - s- oo — o oo ooo ■ o • • o o ••■0''°-_; oooooooo-o- 3 o O- ■ r\j o o oo OO . .. . . . . .o- — 'O i. o o i. o ooooooooooooooo .oooooooooc ooo o o o o ooc -o — o o i o^^ iinnom inOffif^ Oi iOi i O ■ CO ^ooooooooooo -oo .oooooooooo )i-c3(-o cj ooo oooo o u o o i. . . o - ■■ o ■ -c o -OOOlOOW O (-- O u so • o o • o - o ooooooooooooooooooc oo— o ooo— o oooo o c — io m o «s- — - - oeo c^ mo'io'i -(-> CM CO f ooooo i-ooooooooooooo o o — O *J o o — o oooo o o — i\j co *r — - NNif>r>i — O >— w n «j tfi ic ui r-. oo dtxi c\i — dodo— * odd o dii d — — d o'ci^d ■ o ■ o o ■ • ■ o oopooopooopoooooooo oooooooooooooc - — o o ooooooooooooooo ooooooooooooooooooo ooooooooooooooo csjmi— o^,r~-i/iQO'^Cfifsi»j-f^f— m fi en r— r~> ij.j-(»iMfsnoc«— OCstNCOCOI^i'lIM^OOM'- in in OOOOOOOOOOOOOOO g°S°°H°s=555So5 iSSS-SBSS^ oooooooooo ,2 ^ Oi cnui O 9 ffi ' — Oo o c m o i t. >, a jm T3-r- rtJUrtlCUO >, CCS»J jaou.o^£E^^^zcLCct/i>>3 C C 3 O 2 •£ Sn As the Nation's principal conservation agency, the Department of the Interior has responsibility for most of our nationally owned public lands and natural resources. This includes fostering the wisest use of our land and water resources, protecting our fish and wildlife, preserv- ing the environmental and cultural values of our national parks and historical places, and providing for the enjoyment of life through out- door recreation. The Department assesses our energy and mineral resources and works to assure that their development is in the best interests of all our people. The Department also has a major responsi- bility for American Indian reservation communities and for people who live in island territories under U.S. administration. GPO 850- 079 UNITED STATES DEPARTMENT OF THE INTERIOR FISH AND WILDLIFE SERVICE EDITORIAL OFFICE AYLESWORTH HALL, QSU FORT COLLINS, COLORADO 80523 POSTAGE AND FEES PAID U.S. DEPARTMENT OF THE INTERIOR INT 423 NOTE: Mailing lists are computerized. Please return address label with change of address.