FIELD MUSEUM OF NATURAL HISTORY FOUNDED BY MARSHALL FIELD, 1893 PUBLICATION 259 BOTANICAL SERIES VOL. IV, No. 7 SPERMATOPHYTES, MOSTLY PERUVIAN 1. The Status of Rynchospora. Some Peruvian Sedges . . 165 2. New Capers from Peru 168 3. Some Peruvian Annonaceae with a New Guatteria . . . 171 4. New and Renamed Peruvian Melastomes 172 5. Variants of Ostrya and a Texas Tephrosia 192 BY J. FRANCIS MACBRIDE B. E. DAHLGREN Acting Curator, Department of Botany EDITOR THE LIBRARY OF THE JUL 4 1943 CHICAGO, U. S. A. July 5, 1929 Dup. U. of C, Oh. FIELD MUSEUM OF NATURAL HISTORY FOUNDED BY MARSHALL FIELD, 1893 PUBLICATION 259 BOTANICAL SERIES VOL. IV, No. 7 SPERMATOPHYTES, MOSTLY PERUVIAN 1. The Status of Rynchospora. Some Peruvian Sedges . . 165 2. New Capers from Peru 168 3. Some Peruvian Annonaceae with a New Guatteria . . . 171 4. New and Renamed Peruvian Melastomes 172 5. Variants of Ostrya and a Texas Tephrosia 192 BY J. FRANCIS MACBRIDE B. E. DAHLGREN Acting Curator, Department of Botany EDITOR THE LIBRARY OF THE CHICAGO, U. S. A. July 5, 1929 PRINTED IN THE UNITED STATES OF AMERICA BT FIELD MUSEUM PRESS FB " Z- SPERMATOPHYTES, MOSTLY PERUVIAN FROM THE CAPTAIN MARSHALL FIELD EXPEDITIONS TO PERU J. FRANCIS MACBRIDE i. THE STATUS OF RYNCHOSPORA. SOME PERUVIAN SEDGES In considering the South American species of this well-known group of Cyperaceae in connection with my own collections from Peru, I was impressed with the similarity of certain members to species of Dichromena Michx. Fl. Bor. Am. i : 37. 1803. On investiga- tion I found that Rynchospora Vahl, Enum. 2: 229. 1806 (Rhyncho- spora of authors) has frequently been amended to include Dichromena and that authors who have maintained both genera have often done so with evident misgivings. It seems sufficient to refer to Bentham and Hooker, Gen. PI. 3: 1047. 1883 for a resume of the treatments of authors to that date: Bockeler merged the two genera in 1872, Linnaea 37: 525, and the recent distinguished student of the Cyper- aceae, Kiikenthal, Bot. Jarhb. 56: Beiblatt 125: 16. 1921, followed suit. C. B. Clarke, Urb. Symb. Ant. 2: 99. 1900, kept Dichromena but remarked: "Genus a Rynchospora Sect. Psilocarya non nisi differt spiculis capitatis, saepe stramineis vel cinnamomeis, nuces paucas (1-3) maturantibus ; forsan cum cl. Boeckeler in Rynchospora potius mergendum." Yet this same authority retained in Rynchospora, cf. Kew Bull. Misc. Inf. Add. Ser. 8: 117 et seq. 1908, species that exhibit in one or more respects the salient characteristics upon which Dichromena has been maintained as a genus, for instance, the characters of few nutlets, or no setae, or simple capitate inflorescence, or preseence of foli- aceous bracts. Among others mention may be made of R. Weber- baueri Clarke, maturing 1-3 nutlets, of R. barbata (Vahl) Kunth and R. rigida (Kunth) Bcklr. with involucrate capitate and pale inflo- rescences but developed setae, of R. ebracteata (Standl.) Pfeiff. with 165 i66 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV bracts reduced to subobscurity but in other respects with the char- acters of Dichromena. The list could be extended of species such as these that collectively at least show a dovetailing of all characters upon which Dichromena has rested. These "connecting" species are all tropical American : the botanist writing on an area outside their range might well find it convenient for himself and for his readers to recognize both genera, for there, in aspect alone, they seem different enough. But from the stand- point of sound taxonomy, all species considered, there is only one generic concept, for which the proper name is Dichromena. Ryncho- spora stands as a conserved name only for those who regard it as distinct from Dichromena. Probably the segregate genus Pleurostachya Brongn. should also be included in Dichromena. It was regarded as a section only of Rynchospora by Bentham and Hooker. There is an excellent review of these and allied genera by Pfeiffer, Rep. Spec. Nov. 23: 342. 1927, and his synopsis of the characters shows clearly their relative distinct- ness, and the weakness of the particular segregates considered here. The following species have been recorded as growing in Peru with the exception of the first, which is cited as a representative North Temperate species of the Section Rynchospora. Dichromena alba (L.), comb. nov. Schoenus albus L. Sp. PI. 44. 1753. Rynchospora alba (L.) Vahl, Enum. 2: 236. 1806. Dichromena alta (Bcklr.), comb. nov. Rynchospora globosa (HBK.) R. & S. Syst. 2: 89. 1817, not D. globosa (HBK.) R & S., I.e. 90. Chaetospora globosa HBK. Nov. Gen. and Sp. i: 230. 1815. Rynchospora alta Bcklr. Cyp. Nov. Heft 2: 20. 1890, ace. Clarke. DICHROMENA BLEPHAROPHORA Presl, Rel. Haenk. 1:351. 1830. D. ciliata Presl, I.e. 197, t. 32, f.2, 1828, not Vahl. DICHROMENA BRUNNEA Bcklr. Linnaea, 37: 593. 1873. DICHROMENA CILIATA Vahl, Enum. 2: 240. 1806. DICHROMENA DISSITIFLORA Steud. ex Bcklr. Linnaea, 37: 598. 1873 Dichromena glauca (Vahl), comb. nov. Rynchospora glauca Vahl, Enum. 2: 233. 1806. Rynchospora ferruginea (HBK.) R. &. S Syst. 2: 85. 1817. DICHROMENA GLOBOSA (HBK.) R.&.S. Syst. 2: 90. 1817. Schoenus globosus HBK. Nov. Gen. and Sp. i: 229. 1815. If I may judge from a specimen of Andre referred here, from near Vista, Nov. Granat., this species differs from D. ciliata as indicated by HBK. and appears to merit recognition, although reduced by Clarke, Urb. Symb. Ant. 2: 101. 1900. SOME PERUVIAN SEDGES 167 Dichromena Kiikenthalii (Pfeiff.), comb. nov. Rynchospora Kukenthalii Pfeiff. Rep. Spec. Nov. 17: 237. 1921. R. Uleana Kiikenth. Bot. Jahrb. 56: Beibl. 125: 17. 1921, not Bcklr. Allg. Bot. Zeit. 2: no. 1896. Dichromena Lechleri (Steud.), comb. nov. Rynchospora Lechleri Steud. ex Becklr. Linnaea, 37: 633. 1873. Dichromena macrochaeta (Steud.), comb. nov. Rynchospora macrochaeta Steud. ex Bcklr. Linnaea, 37: 632. 1873. Dichromena Mandonii (Clarke), comb. nov. Rynchospora Man- donii Clarke, Kew Bull. Add. Ser. 8: 38. 1908. Dichromena polyphylla (Vahl), comb. nov. Rynchospora poly- phylla Vahl, Enum. 2: 230. 1806. Schoenus polyphyllus Vahl, Eclog. Amer. 2:5. 1798. DICHROMENA PULCHELLA Poepp. & Kth. in Kunth, Enum. 2: 277. 1837. DICHKOMENA RADicANs Schl. & Cham. Linnaea, 6: 28. 1831. Dichromena Ruiziana (Bcklr.), comb. nov. Rynchospora Ruiziana Bcklr. Linnaea, 37: 641. 1873. DICHROMENA RUIZIANA (Bcklr.) Macbr., var. triceps (Bcklr.), comb. nov. Rynchospora triceps Bcklr. Linnaea, 37: 642. 1873. ^- Ruiziana Bcklr., var. triceps K. Schum. ex Clarke in Urb. Symb. Ant. 2: 115. 1900. Dichromena semiinvolucrata (Presl), comb. nov. Rynchospora semiinvolucrata Presl, Rel. Haenk. i: 198. 1830. Dichromena umbraticola (Poepp. & Kth.), comb. nov. Ryncho- spora umbraticola Poepp. & Kth. in Kunth, Enum. 2: 300. 1837. Dichromena Weberbaueri (Clarke), comb. nov. Rynchospora Weberbaueri Clarke, Bot. Jahrb. 37: 518. 1906. Cyperus Schraderi, nom. nov. C. Martianus Schrad. ex Nees in Mart. Fl. Bras. 21: 32. 1842, not Schult. Mant. 2: 108. 1824. According to Weberbaur, Veg. der Erde 12: 238. 1911, this Brazilian species has been found in southern Peru, Valley of Sandia, Dept. of Puno. Stenophyllus arenarius (Nees), comb. nov. Isolepis arenaria Nees, Linnaea 9: 291. 1834. Bulbostylis arenaria (Nees) Lindm. Bihang K. Sv. Vet. Akad. Handl. 26, Afd. 3, No. 9. 19. 1900. Sten- ophyllus tenuifolius (Rudge) Britton, var. latesquamata Pfeiff. Bot. Archiv6: 189, 193. 1924. There seems to be good precedent for regarding this as a species. Although it may be closely related to 5. tenuifolius, its suborbicular scales are suggestive of those of 5. scaber. Originally Brazilian, it is now known from the Departments of Ancash and Junin, Peru. STENOPHYLLUS CAPILLARIS (L.) Britton, var. ciliatus (Presl), comb. nov. Stenophyllus eu-capillaris ciliata (Presl) Pfeiff. Bot. Archiv6: 187, 193. 1924. 168 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV Originally collected near Huanuco, Peru, by Presl and apparently not recollected in Peru, this plant seems to be better known in Brazil. Scleria spicata (Spreng.), comb. nov. Rhynchospora spicata Spreng. Syst. i: 194. 1825. Scleria pleostachya Kunth, Enum. 2: 355- 1837. According to Weberbaur this Brazilian species has been found in the Sandia Valley, Dept. of Puno, Peru. 2. NEW CAPERS FROM PERU Gynandropsis Herrerae, spec, nov., annua, herbacea, simplex subglabra vel ubique minutissime pulverulento-pubescens; caulibus circa 3 dm. altis; foliis intermediis summisque ternatis, petiolis gracilibus striatis 4-7 cm. longis; foliolis petiolatis ovatis vel ovato- lanceolatis basi cuneatis mediocriter obliquis inaequalibus, apice plus minusve abrupte acutis vel acuminatis submembranaceis plerumque 6-8 cm. longis, 2.5-3 cm. latis; racemis laxifloris ebract- eatis vel bracteis caducissimis ; pedicellis fructiferis refracto-paten- tissimis circa 2 cm. longis; laciniis calycinis ovatis acutis circa 3 mm. longis; petalis 10-12 mm. longis, lamina elliptico-spathulata, ungue subaequilongo ; staminibus nonnihil vel vix corollam excedentibus ; toro incluso tantum 7 mm. longo; siliqua (nondum perfecte matura) lineari-cylindracea utrinque acuta circa 6 cm. longa, thecaphorum pedicello duplo brevius. — PERU: forest, alt. 2 — 2300 m., near Rio Yanamayo, below "Pillahuata," Dept. of Cuzco, May, 1925, Francis W. Pennell 14073 (TYPE, Field Museum). This species is entirely different from any so far recorded from Peru and apparently has no known close relative. The collector recorded the flower as "flesh ocher" in color. It is named for Dr Herrera of the University of Cuzco, the distinguished authority on the plants of the region. Cleome Figueroae, spec, nov., inermis, subherbacea vel suffrutes- cens; caulibus striatis subsimplicibus 1-1.5 m- altis glabris vel obscure subadpresse strigillosis ; foliis 5~7-sectis utrinque minute pubescentibus cum pilis crispulis vel hispidulis subtus pallidioribus et nerviis prominente purpurascentibus; petiolis 8-10 cm. longis plus minusve strigillosis etiamque cum pilis longioribus parce inter- mixtis vel subpilosis imprimis ad apicem; foliolis petiolatis ovato- lanceolatis utrinque acuminatis 5-8 cm. longis 1.5-3 cm- latis; racemis elongatis (2.5 dm. plus minusve); bracteis simplicibus suborbiculari-cordatis aliquid scabris; sepalis rotundo-ovatis acutis vix 5 mm. longis; petalis oblongo-ellipticis fere 2 cm. longis; pedi- cellis fructiferis subreflexis circa 2 cm. longis; siliquis (immaturis) glabris lineari-cylindraceis obscure torulosis circa 6 cm. longis, longe stipitatis; stipes 8 cm. longis. — PERU: Huacachi, near Muna, Dept. of Huanuco, May 2o-June i, 1923, Macbride 4173 (TYPE, Field Museum). NEW CAPERS FROM PERU 169 Probably this species most resembles C. longifolia Presl which, however, is described as having lanceolate acuminate sepals and much shorter (4 cm. long) pods. It is named for Reyes Figueroa of Mito, Peru, in recognition of his loyalty and interest while serving as guide on the Captain Marshall Field Botanical Expeditions of 1922 and 1923. Cleome Herrerae, spec, nov., ut videtur peraffinis C. longifolia; caulibus, foliis subtus, petiolisque plus minusve strigillosis ; sepalis ovato-lanceolatis, basi haud attenuatis, apice sensim subacuminatis, dense subadpresse strigillosis, 6-7 mm. longis; petalis oblongo- lanceolatis, basi anguste attenuatis, circa 17 mm. longis; ovario glabro; pedicellis fructiferis solum 1.5 cm. longis; siliquis (immaturis) circa 5 cm. longis; stipes 3.5 cm. longis. — PERU: Valle de San Miguel, Cedrobamba, Dept. of Cuzco, 20 de Julio de 1928, F. L. Herrera 1992 (TYPE, Field Museum). C. longifolia Presl, Rel. Haenk. 2: 84. 1831, the only Peruvian species to which this beautiful Cleome is closely related, is described as having sepals that are attenuate at the base, petals about 12 mm. long, pedicels 2.5 cm. long and pods about 4 cm. long. It does not seem desirable at this time to regard these differences, definite as they are, as representing merely a variation of this species. Dr. Herrera reports the common name as "Facma." Cleome limoneolens, spec, nov., C. chilensis peraffinis; basi indurato-suffrutescens vel fruticosa, ramosa; caulibus aliquot dm. altis adscendentibus flexuosis mediocriter basi ad apicem dense sordide glanduloso-pubescentibus cum pilis hirsutulis vel crispulis; foliis 5-7-foliolatis utrinque granuloso-glandulosis vel paullo scab- ridis; petalis tantum 5-6 mm. longis. — PERU: loose soils of river- canon slopes, San Rafael, Dept. of Junin. April 4, 1923, Macbride 3145 (TYPE, Field Museum); steep rocky grassland, Huacachi, near Muna, Dept. of Huanuco, May 2o-June i, 1923, Macbride 4088. This may prove to be only a variety of C. chilensis DC., which is not uncommon in Peru, but no intermediate plants have been seen. Its shrubby base and small flowers seem, therefore, to be distinctive characters. The plants were noticeably lemon-scented. Cleome mono chroma, spec, nov., subherbacea stricta haud ramosa aliquot dm. (vel i m. ?) alta; caulibus minute pulverulentis fere basi ad apicem vel plus minusve villosis et parce cum glandulis stipitatis glanduliferis imprimis parti superiori; foliis membranaceis longi-petiolatis (petiolis circa 1.5 dm. longis) 7-foliolatis utrinque molliter subadpresse strigillosis; foliolis oblongo-ellipticis vel oblongo- lanceolatis utrinque attenuatis, — apice acuminatis basi extenuato- cuneatis — sessilibus plerumque circa 8-12 cm. longis 2-3.5 latis; 170 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV floribus 4 cm. longis plus minusve villosis et parce glandulosis ut videtur sepalis petalisque albo-viridibus ; sepalis anguste lineari- acuminatis 2-2.5 cm- longis; petalis paullo latioribus subobtusis; pedicellis fructiferis 4-5 cm. longis; stipes 5-6 cm. longis siliquiis obscure puberulis pendulis vel suberectis anguste oblongo-cylind- raceis 10-14 cm. longis fere i cm. latis basi subacutis apice acumina- tis. — PERU: sandy trail edge, La Merced, Dept. of Junin, Aug. 10-24, 1923, Macbride 5386 (TYPE, Field Museum). COLOMBIA: Rio de la Honda near Arbelaez, Feb. 19, 1876, Ed. Andre 1589. MEXICO: Misantla, Vera Cruz, Aug., 1912, Pur pus 5873. No Peruvian species resembles this unusually well-marked Cleome. Its strict habit, large greenish flowers with linear sepals and petals, and long oblong-clavate pods are characteristics that combined make it most distinctive. Yet I have not been able to find a name for it — and the material cited from Colombia and Mexico was distributed unnamed. Cleome eosina, n. nov. C. microcarpa Hass. Rep. Spec. Nov. 12: 254. 1913, not C. microcarpa Ule, Bot. Jahrb. 42: 201. 1908. Since there is already a Cleome Hassleriana this Paraguayan species may be called eosina as it suggested to its author several species, C. af finis DC., C. diffusa Banks and C. aculeata L. Capparis Schunkei, spec, nov., arbor parva; ramulis glabris; foliis submembranaceis fortiter reticulato-venosis ellipticis vel oblon- go-ellipticis plerumque circa 15 cm. longis et 5 vel 6 cm. latis, apice subabrupte breviter acuminatis basi subcuneatis, petiolatis (petiolis circa 1.5 cm. longis), supra glabris (vel juventute minute stellato- pubescentibus) lucidis, subtus ramulisque mflorescentiis obscure subferrugineo-stellato-pubescentibus; racemis ramosis aliquid pani- culatis foliis brevioribus paucifloris; pedicellis gracilibus 1.5-2 cm. longis; bracteis minutis caducissimis; calycibus petalisque mediocriter externe fulvo-pubescentibus cum pilis stellatis; sepalis oblongo- lanceolatis subacutis reflexis 3 mm. longis; squamulis circa i mm. longis; petalis late ovatis subobtusis 5 mm. longis; staminibus circa 20 pistilloque longe exsertis; ovario ovoideo glabro. — PERU: Chan- chamayo Valley, Dept. of Junin, 1924-1927, Carlos Schunke 418 (TYPE, Field Museum); Hacienda Schunke, La Merced, Macbride 5701. No species of the subgenus Colicodendron — to which this plant belongs — appears to approach C. Schunkei except possibly C. lepi- dota (Turcz.) Knuth of Venezuela from which it differs in character of pubescence, shape of leaves and number of stamens. In foliage only it suggests two glabrous Peruvian species, C. laurina HBK. and C. Sprucei Eichl. both of which, however, belong in another subgenus. PERUVIAN ANNONACEAE 171 C. Schunkei is a small tree with spreading branches and pale greenish-yellow flowers. It is named with pleasure for the collector whose hospitality I enjoyed when on the Captain Marshall Field Botanical Expedition to Peru in 1923. 3. SOME PERUVIAN ANNONACEAE WITH A NEW GUATTERIA Guatteria Rusbyi, nom. nov. G. lucida Rusby, Mem. N. Y. Bot. Gard. 7: 245. 1927, not G. lucida Presl, Rel. Haenk. 2: 78. 1831. This recently described species from Bolivia may appropriately bear the name of the author who evidently overlooked the much earlier use of the term lucida for a distinct Peruvian species. Guatteria pachypetala (Diels), comb. nov. Oxandra pachypetala Diels, Notizb. 10: 173. 1927. If one may judge from the original characterization, this Peruvian tree is a Guatteria rather than an Oxandra because it has "stamina numerosa" and "connectivum vix productum." According to all authorities, even including Engler and Diels, Notizb. 3: 49. 1900, Oxandra has few (6-18) stamens and the connective is produced above the anther. If these characters do not "hold," Oxandra can hardly be maintained as a distinct genus. Guatteria socialis, spec, nov., liana; ramulis foliisque glabris; foliorum petiolo prope i cm. longo supra nonnihil sulcato crasso, lamina chartacea oblongo-vel ovato-elliptica plerumque circa 1 5 cm. longa 4-7 cm. lata utrinque sensim angustata vel apice subabrupte acuminata (acumine subobtusato) supra pallide-glauca nitidula subtus viridi-nitidiore, nervis lateralibus primariis utrinque 9-11 angulo acuto adscendentibus subprominulis, venis reticulatis pro- minulis; pedunculis glabris solitariis axillaribus subgracilibus 2.5-3 cm. longis, basi squamuloso-bracteolatis vix articulatis vel medio quandoque unibracteolatis ; bracteolis minute ciliatis; sepalis orbi- culari-ovatis subacutis solum minutissime ciliatis; petalis glabris ut videtur subaequalibus subovatis vel oblongo-ellipticis circa 12 mm. longis, 6-8 mm. latis; baccis ignotis. — PERU: Dept. Junin, Chanchamayo Valley, 1500 m., Oct. 1924-27, Carlos Schunke jp5 (TYPE, Field Museum). This was noted in the field by the observant collector to be a liana and apparently is only the second scandent Guatteria to be recorded. G. scandens Ducke has pubescent flowers. Dr. Diels has described a climbing Annona (A. scandens Diels). In addition to the habit this species seems to differ in size and shape of leaves, elongate 172 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV peduncles, etc., from the comparatively few Guatterias with com- pletely glabrous foliage and flowers. Its relationship may be with G. jurensis Diels. Fusaea rhombipetala (R. & P.), comb. nov. Anona rhombipetala R. & P. ex G. Don, Gard. Diet, i: 87. 1831. Safford has recently established the genus Fusaea, Contrib. U. S. Nat. Herb. 18:64. 1914, and evidently for the most excellent reasons. He has also indicated that the Peruvian species cited above is in fact a Fusaea rather than an Annona but he did not make the transfer. Duguetia odorata (Diels), comb. nov. Aberemoa odorata Diels, Notizb. 10: 171. 1927. Duguetia pedunculata (Diels), comb. nov. Aberemoa pedunculata Diels, Bot. Jahrb. 37: 409. 1906. Duguetia peruviana (R. E. Fries), comb. nov. Aberemoa peru- viana R. E. Fries, Kgl. Sv. Vet. Akad. Handl. 34, no. 5: 22, pi. 3, fig. 1-3. 1900. Duguetia St. Hil. Fl. Braz. Mer. i: 35. pi. 7. 1825 is "conserved" according to the International Rules of Botanical Nomenclature in place of Aberemoa Aubl. PI. Guian. 1:610. 1775. There seems always to have been some question, however, as to the generic identity of Aublet's plants with St. Hilaire's, and Safford has recently pointed out (Contrib. U. S. Nat. Herb. 18: 60-6 1. 1914) that they can scarcely be congeneric. The use of the name Aberemoa, therefore, should remain restricted to the original and very imperfectly known species A. guianensis Aubl. 4. NEW AND RENAMED PERUVIAN MELASTOMES Ernestia Sprucei (Cogn.), comb. nov. E. tenella (Bonpl.) DC., var. Sprucei Cogn. Bull. Acad. Belg. 3. 14: 929. 1887. Cogniaux described this plant, to distinguish it from typical E. tenella, as a branched shrub 1.5-2 m. high with purple flowers and densely setose (at apex) ovary. These characters seem to me specific : E. tenella is subherbaceous or woody below, 0.5 m. high and has white flowers and a scarcely puberulent ovary. Brachyotum callosum, spec, nov., fruticosum 0.5-1.5 m. altum, dense ramosissimum ; ramis teretibus flexuosis adpresse setulosis inferne demum glabratis; ramulis dense subadpresse setulosis plerumque i dm. longis; foliis valde approximatis 3-nerviis oblongo- PERUVIAN MELASTOMES 173 ellipticis utrinque obtusis 8-10 mm. longis, 4-5 mm. latis subplanis vel mediocriter revolutis; petiolis (circa i mm. longis) foliisque subtus adpresse densequ e setulosis, supra callis conicis apice breviter setiferis dense onustis; floribus pedunculatis plerumque ternis 5- meris (semper?) breviter pedicellatis ; bracteis tarde deciduis; calycis tubo conico vel anguste campanulato parce adpresseque strigoso circa 8 mm. longo, lobis triangulari-ovatis acutis circa 5 mm. longis; petalis late obovatis circa 12 mm. longis plus minusve breviter ciliatis; antheris oblongo-linearibus, connective brevissime elongate non tuberculato. — PERU: in scattering hillside thickets, 15 miles northeast of Huanuco, June 12-22, 1922, Macbride & Feather stone 2181 (TYPE, Field Museum). This species is a member of the section Adesmiae and apparently is most nearly related to B. Trianaei Cogn. Melast. 167. 1891, not- withstanding it 5-merous flowers. It can be distinguished from Cogniaux's plant by the entirely different pubescence of the leaves and calyx. In some respects it resembles both B. lutescens (R. & P.) Triana and B. microphyllum Triana but the pubescence of the former is hispid and strigose and that of the latter sparsely strigose and setulose. In most if not all species of Brachyotum the flowers may be either 4- or 5-merous. For instance, B. lutescens was described as sometimes 5-merous and so are my Peruvian specimens. B. callosum therefore will probably be found to vary in the same way. Its flowers (apparently yellowish) were not fully grown so the petals may become longer than described. Brachyotum Figueroae, spec, nov., fruticosum, circa 1 m. altum; ramis gracilibus tarde glabratis ut videtur 2-3 dm. longis; ramulis mediocriter approximatis plerumque i dm. longis dense subadpresse- que hirsuto-setulosis; petiolis foliisque subtus dense hirsutis, foliis supra crassis conicis apice setiferis dense tuberculatis, late ovatis subplanis circa 9 mm. longis, 5-6 mm. latis; floribus breviter pedun- culatis solitariis vel 3-4, 4- vel 5-meris; calycis rubescentibus sparse setulosis, tubo plus minusve anguste campanulato circa i cm. longo, lobis ovatis circa 4 mm. longis; petiolis rotundato-obovatis breviter ciliatis circa 12 mm. longis; connective non producto antice brevissime 2 -tuberculato. — PERU: stream bank, Catuc, pueblo 15 miles east of Huaraz, Dept. of Ancash, Oct. 4, 1922, Macbride & Feather stone 2504 (TYPE, Field Museum). Only two other species of the section Dicentrae seem to be closely related to this, namely B. Maximowiczii Cogn. and B. rosmarinifolium (R. & P.) Triana. B. Figueroae may be distinguished from the former of these by its much shorter calyx-lobes in proportion to the tube and whitish flowers, and from the latter by the broadly ovate leaves that are hirsute rather than setulose beneath, as well as by the color 174 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV of the flowers and proportionately shorter calyx-lobes. It is named for Reyes Figueroa, my helpful companion on many collecting journeys in 1922 and 1923. Brachyotum tyrianthinum, spec, nov., fruticosum, 1-2 m. altum, solum mediocriter ramosum ; ramis gracilibus glabris paullo vel hatid excoriatibus; ramulis petiolis foliisque supra plus minusve dense adpresse setulosis; petiolis circa 3 mm. longis; foliis ovatis plerumque 2 cm. longis et circa i cm. latis, 3-nerviis, subtus parce hirtello- setulosis; floribus solitariis cernuis 4-meris (semper?) " violaceo- purpureis; calycis tubo adpresse setulosis campanulato 7 mm. longo, lobis parce setulosis oblongo-lanceolatis circa i cm. longis; petalis subrotundato circa 15 mm. longis minute ciliatis; connective obscure vel breviter tuberculato. — PERU: slender rather open bush on shrubby southwestern slope, Mito, Dept. of Huanuco, July 8-22, 1922, Macbride & Feather stone 1438 (TYPE, Field Museum). This species, definitely referable to the section Dicentrae, appa- rently resembles both B. Grisebachii Cogn. and B. Naudinii Triana. It differs from the former in its ovate rather than oblong leaves and in its shorter and broader petals. The longer leaves and calyx-lobes and rounded petals separate it from the latter. The pubescence on the upper leaf-surfaces is peculiar in that each bristle emanates from an elongate callous that scarcely exceeds the bristle in width and is attached to or is a part of the leaf-surface. TIBOUCHINA LONGIFOLIA (Vahl) Baill., var. simulans, var. nov., foliis interdum 7-nerviis; sepalis plus minusve setoso-glandulosis ; petalis basi ad apicem valde ciliatis; connective basi breviter pro- ducto. — PERU: La Merced, Dept. of Junin, Aug. 10-24, 1923, Macbride 5300 (TYPE, Field Museum) ; Pozuzo, Dept. of Huanuco, June 10-22, 1923, Macbride 4556. I should be inclined to describe this variant as a new species were it not for the apparent variability of T. longifolia and the fact that the numerous segergates based on relative length of connective and character and degree of pubescence when better known will very likely be found to represent extremes only in different directions of variation. The above variety resembles also T. stenopetala Cogn. except that the petals are not narrowed at base. Tibouchina pleromoides (Naud.), comb. nov. Lasiandra plero- moides Naud. Ann. Sci. Nat. 3. 13: 131. 1850. Pleroma maurocarpum Triana, Trans. Linn. Soc. 28: 47. 1871. Tibouchina maurocarpa (Triana) Cogn. Melast. 260. 1891. PERUVIAN MELASTOMES 175 Aciotis cordata (Veil.), comb. nov. Melastoma cor data Veil. Fl. Flum. 178. 1825 et Ic. 4: tab. 114. 1827. Spennera dysophylla Benth. Journ. Bot. 2: 296. 1840. Aciotis dysophylla (Benth.) Triana, Trans. Linn. Soc. 28: 52. 1871. Triana, I.e. 151, refers the plant of Velloso to that of Bentham without question and as it is the earlier known its name may be used. Monochaetum canescens (Bonpl.), comb. nov. Rhexia canescens Bonpl. Rhex. 47. tab. 18. 1823. R. Bonplandii Kunth in index, Rhex. M. Bonplandii (Kunth) Naud. Ann. Sci. Nat. 3. 4: 51. tab. 2. f.iB. 1845. Monochaetum subditivum, spec, nov., M. dicranantherum peraffine; petiolis plerumque 8-10 mm. longis; foliis anguste ovatis 3-4 cm. longis circa 1.5 cm. latis, apice mediocriter attenuato-acutis, 5-plinerviis; cymis axillaribusque terminalibus, 2-4-floris; calyce setis patulis elongatis plerumque glandulosis hirsuto; lobis oblongo- lanceolatis definite acutis; staminibus minoribus caudatis caudis linearibus antheris paullo brevioribus. — PERU: an open 3-4 ft. shrub of montana slopes. Villcabamba, hacienda on Rio Chinchao, Dept. of Huanuco, July 12-26, 1923, Macbride 5194 (TYPE, Field Museum). This typical species of Monochaetum on casual observation would readily pass for M. dicranantherum (R. & P.) Naud. but careful examination discloses a number of differences that make its refer- ence to the latter unsatisfactory. The calyx-hairs are nearly all tipped with a gland; otherwise the pubescence is exactly that of M. dicranantherum. Also the leaves are longer and more gradually acute, and the appendage of the smaller anthers almost equals the anther in length. These differences may be found to be relative but at present there is no evidence to this effect. M. subditivum bears some resemblance to M. mllosum Gleason, Bull. Torr. Club, 52: 335, but the leaf-pubescence of the latter is evenly distributed over the upper surfaces. In both M. dicranan- therum and the species proposed here it is confined to lines between the veins. There is also a difference in the relative size of the smaller anthers and their appendages. Two other species that are related are at once distinct because of their 7 -nerved leaves, namely M. pauciflorum Triana and M. glanduliferum Triana. Marshallfieldia, gen. nov. Flores 5-meri. Calycis tubus infundib- ularif ormi-campanulatus ; limbus brevis, haud dilatatus, subsimplex, lobis rotundatis. Stamina valde inaequalia filamentis distincte alatis; maiorum antherae tenues, apice arcuatae, anguste uniporosae; 176 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV minorum antherae haud rostratae, obtusae, rectae. Connective infra loculus non producto, antice inappendiculato, postice processu elongate fere erecto apice biaristato instructo. Ovarium liberum. — Frutices scandentes vel ut videtur sarmentosi vel pseudoparasitici. Folia petiolata, 5 nervia. Flores mediocres, in racemos (vel pani- culas?) paucifloros terminales depositi. — Affinis videtur Merianiae. Marshallfieldia corallina, spec, nov., scandens; ut videtur caule indiviso flexuoso dense rufo-villoso ; foliis late ovalibus vel rotundis, basi rotundatis, apicem versus sensim angustatis, acutis, membrana- ceis, supra primo leviter villosis imprimis ad nervos demum glaber- rimis, subtus mediocriter dense rufo-pilosis praecipue ad nervos, circa n cm. longis et 8 cm. latis; petiolis etiam dense rufo-villosis, 2-2.5 cm. longis; floribus 2 vel 3, breviter pedicellatis ad apicem ramorum subcongestis ; calycis tubo parce adpresse villoso-setosis vel subglabris, basin versus cuneatim angustato, angusto, circa 5 mm. longo, limbo non dilatato, lobis rotundatis, abrupte acutis, margine tenuiter membranaceis minute serrulatis, circa 1.5 mm. longis; petalis oblongo-obovatis, circa 12 mm. longis, 6 mm. latis; staminibus valde inaequalibus ; filamenta maiorum 9 mm. longa, antherae fere 8 mm. longae, appendices circa 3 mm. longae; minorum filamenta fere n mm. longa, antherae et appendices 4.5 mm. longae; ovario glabro, stylo 8 mm. longo. — PERU: climbing a tree-trunk, Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27-Sept. i, 1923, Macbride 5642 (TYPE, Field Museum). This interesting new genus of the Melastomaceae was collected on the second of the botanical expeditions to Peru sponsored by Captain Marshall Field. It is an attractive plant with creeping stem and somewhat fan-shaped leaves densely covered on the under side with soft rusty-red hairs. The upper leaf -surface is dark green and smooth. The flowers are a coral-pink with yellow anthers. Marshallfieldia is not satisfactorily referable to any of the tribes of its family as these have been characterized by Cogniaux in his monograph, Melast. 1891. Although only flowering material is known it appears to be allied to the Merianiae in which, however, there are no genera with very unequal stamens. Notwithstanding the unequal stamens its generic relationship is apparently with Adelobotrys which it resembles in habit and foliage and exactly matches in character of the anther-appendages but its calyx is lobed like that of some species of Meriania. Meriania Weberbaueri, spec, nov., arbor 8 m. alta; ramis ad nodos satis incrassatis plus minusve obscure tetragonis dense fur- furaceo-puberulis vel demum fere glabris; foliis late ovalibus vel elliptico-ovatis, obtusis, basi aliquid attenuatis, margine inferne integerrimis superne valde repando-serrulatis, circa 2 dm. longis, PERUVIAN MELASTOMES 177 i dm. latis (superioribus multo reductis) submembranaceis supra glabris, subtus minute denseque pubescentibus cum pilis stellatis et pilosis intermixtis, 7-plinerviis; nervis mediocriter prominentibus; nervulis transversalibus subtus tenuissime ramuloso-reticulatis ; cymis paucifloris; calycis lobi triangulates, tubo aequales vel longi- ores, dentibus exterioribus vix 3 mm. longis; tubo dense furfuraceo- puberulis, campanulato 6 mm. longo, superne 8 mm. lato, obscure costato distincte tuberculato, limbo haud dilatato; petalis roseis late obovatis circa 2 cm. longis; staminibus magnis, antheris valvis valde crispato-undulatis, connective basi valde incrassato, in ap- pendicem brevem minute bilobatam producto. — PERU: valley of the Rio Masamerich, tributary of the Rio Pangao, Dept. of Junin, May 7, 1913, Weberbauer 6659 (TYPE, Field Museum). This beautiful species, although clearly a member of the section Umbellatae, is not closely related to any of the known species. Its stamens resemble most those of the otherwise entirely different M. speciosa (Bonpl.) Naud. except that the shorter appendages are shortly bilobed instead of acute and hooked. Macrocentrum peruvianum (Cogn.), comb. nov. M. fasciculatum (DC.) Cogn., var. peruvianum Cogn. Bot. Jahrb. 42: 138. 1908. This simple-stemmed procumbent plant with fruiting calyces nearly 1.5 cm. long can scarcely be regarded as only a variety of M. fasciculatum (DC.) Cogn. which is known only from French Guiana and is an erect branched herb with much longer petioles and leaves and much shorter — only 6 mm. long — fruiting calyx. Tococa undabunda, spec, nov., fruticosa circa 1.5 m. alta, mac- rophylla, anisophylla; ramis subteretiusculis plus minusve setulosis demum glabris ; ramulis flexuosis petiolisque undique dense setulosis, setis elongatis, rigidis, patentibus; foliis longe petiolatis, elliptico- ovatis basi subrotundatis vel quandoque leviter attenuatis apice subabrupte attenuato-acuminatis vel caudatis, margine obscure undulatis et longiuscule satis denseque ciliatis, 5-rierviis, supra parce subadpresseque setulosis subtus ad nervos subdense hirtellis caeterum glabris, majoribus plerumque 2-2,5 dm. longis et 1-1.4 dm. latis, in petiolo vesciculiferis, minoribus aliquando vescicula destitutis; vesicis ut videtur ovoideis; nerviis subtus et supra prominentibus, nervulis transversalibus ubique insigne undulatis, subtus ramuloso- reticulatis; petiolo (in foliis majoribus) 5 vel 6 cm. longo; capitulis terminalibus circa 30- vel 4o-floris breviter pedunculatis ; floribus sessilibus ebracteolatis ; calyce inferne papilloso, superne parce setuloso, tubo oblongo-campanulato circa 5 mm. longo distincte 5-lobato, lobis late ovatis longe setulosis; petala ut videtur alba, immatura. — PERU: open shrub of montana slope, Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27-Sept. i, 1923, Macbride 5734 (TYPE, Field Museum). 178 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV Only a few species of Tococa have sessile flowers borne in a capitate inflorescence. Of these T. unddbunda most suggests T. spadiciflora Triana, Trans. Linn. Soc. 28: 132. 1871 because of the position of the vescicles on the petioles although that species has 5-plinerved leaves and bracteate flowers. If the occurrence of vescicles is disregarded our plant appears to be more nearly related to other species from all of which it is readily distinguished by characters of pubescence, leaves, petioles and calyx. The calyx- pubescence was pink. Sometimes but not usually the leaves beneath the inflorescence are greatly reduced to nearly linear long-acuminate bracts. The name refers to the flexuous branchlets, wavy leaf- margins and cross-veins. Blakea Sawadae, spec, nov., fruticosa 2 m. alta; ramis teretibus vel obtuse subtetragonis; foliis 5-nerviis oblongo- vel ovato-ellipticis, basi plus minusve acutis, apice subacutis et abrupte caudato- acuminatis, plerumque 1.5 dm. longis et 7 vel 8 cm. latis, medio- criter coriaceis, supra glabris sub lente minute denseque puncticulatis, subtus ad nervos nervulosque leviter furfuraceis, caeteris glabris; petiolis robustis vix furfuraceis 2-3 cm. longis; floribus 2-4-fasciculatis subsessilibus 3 cm. latis; pedicellis 2-5 mm. longis; bracteis valde inaequalibus, exterioribus base definite connatis, dense furfuraceis ovatis longe acutis inferne dorso carinatis, 1-1.5 cm- longis, interiori- bus subrotundatis fere glabris vel satis furfuraceis setoso-ciliatis circa i cm. longis; calyce subglabro 12-17 mm. longo, tubo campanu- lato, limbo regulariter 6-lobato, lobis ovatis apice dilatatis subobtusis; petalis albis subrotundalis circa i cm. longis; antheris dolabriformis caeruleis. — PERU: open shrub in hillside thickets, Pampayacu, Dept. of Huanuco, July 19-25, 1923, Macbride 5058 (TYPE, Field Museum). This species appears to be most nearly related to B. caudata Triana, Trans. Linn. Soc. 28: 148. 1871, from which it notably differs, especially in its acutish leaves, longer petioles, shorter ped- icels and regularly lobed calyx. Its fleshy white flowers with con- trasting blue anthers were very attractive. Its -name commemorates fittingly the hospitality and aid in collecting extended by Mr. Masaho Sawada of Huanuco, at the hacienda at Pampayacu. Blakea chanchamayensis, spec, nov., fruticosa-liana ; ramis teretibus junioribus dense rubiginoso-furfuraceis demum glabris; foliis 5-nerviis oblongo ellipticis basi apiceque gradatim attenuatis, apice acuminatis plerumque circa 1.5 dm. longis et 5 vel 6 (-7) cm. latis, chartaceo-subcoriaceis, supra glabris, subtus ad nervos paullo furfuraceis vel demum glabris; petiolis mediocriter gracilibus 2-2.5 cm. longis; floribus solitary's vel 2, subsessilibus; pedicellis 2-3 mm. longis ; bracteis subaequalibus, exterioribus basi leviter connatis plus PERUVIAN MELASTOMES 179 minusve dense furfuraceis fere rotundatis subabrupte acutis circa i cm. longis; interioribus fere glabris circa 8 mm. longis; calyce glabro circa 1.5 cm. longo, tubo campanulato, limbo valde 6-lobatis, lobis late ovatissubacutis; floribusignotis. — PERU: Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27-Sept. i, 1923, Macbride 5676 (TYPE, Field Museum); Chanchamayo Valley, Dept. of Junin, Dec. 1924-1927, Carlos Schunke 299; also Oct., 438. This plant may be either a Topobea or a Blakea, so far as can be determined from the fruiting material available. It seems unlikely, however, that it is referable to the former genus since it most nearly approaches in appearance two species of the latter, namely Blakea caudata Triana and B. Holtonii Hochr. It differs from the former in the narrower gradually acuminate leaves and the subsessile flowers and from the latter in the 5 -nerved leaves and subsessile flowers. Blakea incerta, spec, nov., P. Spruceana perafnnis; ramis glabris vel paullo furfuraceis et setulosis ad nodos; foliis inconspicue 7- nerviis; nervis mediano multo crassiore, 2 exterioribus submsr^ln- alibus vix distinctis; petiolis 1.2-2.5 cm. longis; pedicellis axillaribus binis gracilibus 12-18 mm. longis; bracteis subrotundatis haud acutis demum glabris 12-15 mm. longis; calycis lobis late ovatis 4-5 mm. longis; petalis roseis vix 1.5 cm. longis. — PERU: forest shrub or tree 5-7 m. high, Cushi, Dept. of Huanuco, June 19-23, 1923, Macbride 4850 (TYPE, Field Museum). Except for the fact that the flowers are borne in twos this speci- men matches well the plate of B. Spruceana Cogn. Fl. Bras. 14*: 560. tab. 121. f.I. 1888. However, besides the disagreement in the the number of flowers, B. incerta departs from the description of B, Spruceana in a number of ways: the leaves, although indistinctly are actually 7 -nerved, the petioles average longer, the pedicels are shorter, the bracts are not at all acute and the calyx-lobes are longer. It approaches B. latifolia (R. & P.) D. Don in some respects but is at once distinct by its terete calyx-tube. Miconia Wagneri, spec, nov., arbuscula foliosa 1.5-2.5 m. alta vel interdum arbor; ramulis petiolis pedunculisque pilis longiusculis adpressis inferne simplicibus apice stellato-multifidis submolliter denseque f ulvo-pubescentibus ; foliis membranaceis integerrimis vel remote calloso-repando-denticulatis, late ovato-ellipticis, basi apice- que sub- vel rotundatis vel apice acutis, 1.5-2.5 dm. longis, 8-12 cm. latis, nervulo obscuro marginali praetermisso valde 5-nerviis, supra glabris paullo nitidulis, nervis mediocriter prominulis, subtus ad nervos dense, caeteris parce, stellatis, nervis cum nervulis trans- versalibus prominentibus, venis conspicue ramuloso-reticulatis ; petiolis i.5-2\cm. longis; pedunculis mediocribus plerumque i dm. i8o FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV longis; floribus sessilibus ad apices ramulorum plus minusve glomer- ulatis vel interdum solitariis vel axillaribus, 4-meris; calycis tubo primum minute stellate demum glabro circa 1.5 mm. longo, limbo chartaceo in prima juventute clauso, in anthesi irregulariter lacero; petalis viridibus fere 2 mm. longis, oblongo-obovatis ; antheris, sublinearibus superne paullo attenuatis, apice minute i-porosis, basi postice brevissime calcarato, solum 1.5 mm. longis; stylo 5 mm. Dept. of Junin, Aug. 2;-Sept. i, 1923, Macbride 5718 (TYPE, Field Museum); Chanchamayo Valley, Dept. of Junin, Dec., 1924-1927, Carlos Schunke 355. Only one other species of the section Laceraria has 5 -nerved leaves, namely M. Duckei Cogn. and it has 5-merous flowers and a densely hirtellous calyx. In other respects M. Duckei seems, from description, to resemble M. Wagner i. This interesting species is named for Mr. William Wagner of Lima whose hospitality enabled me to make extensive collections in the vicinity of his delightful hot springs resort at Viso, Peru. Miconia Adrieni, spec, nov., arbuscula usque 2 m. alta, fere glaberrima; ramulis flexuosis, plus minusve sulcatis, subacute tetragonis, superne (etiam petiolis paniculisque) praecipue ad nodos rninutissime parceque squamuloso-furfurascentibus; foliis subchart- aceis utrinque glabris ellipticis vel oblongo-ellipticis, ut videtur apice attenuatis et acutis, basi satis angustatis, in petiolum brevem decur- rentibus, 14-18 cm. longis, 5.5-7.5 cm. I atis, definite 3-plinerviis; nervis supra mediocriter conspicuis, subtus cum nervulis transversali- bus prominentibus, 3 interioribus circa i cm. supra basin folii abeuntibus, 2 lateralibus exterioribus a basi; petiolis 1-2.5 cm- longis; paniculis plus minusve foliosis fere 1.5 dm. longis, ramulis acute tetragonis; floribus subsessilibus, s-meris; calyce 4 mm. longo acute lobato; petalis albis, circa 4 mm. longis; antheris i-porosis, vix 3 mm. longis, paullo attenuatis, anteriore minute bituberculatis; stylo circa 5 mm. longo, superne leviter incrassato; stigmate capi- tellato. — PERU: clump shrub at sunny edge of hillside thicket, Pampayacu, Dept. of Huanuco, July 19-25, 1923, Macbride 5077 (TYPE, Field Museum). This showy-flowered member of the section Eumiconia belongs to the group of species typified by M. prasina (Sw.) DC. and appears to resemble most M. juruensis Pilger. The latter, however, is a tree 15 m. high with much smaller leaves that are very shortly 3-plinerved, and with longer anthers and style. M. Adrieni is named for M. Paul Adrien of the Huaron Mining Company of Shelby, Peru whose friendly aid in my collecting work, together with that of his associates, M. Marcel Tuillier and Mons. Andre" Porret and Auguste Berrier so abundantly merits this men- tion and recognition. PERUVIAN MELASTOMES 181 MICONIA NERVOSA (Sm.) Triana, var. mediana, var. nov., foliis longe petiolatis, inf erne longe attenuates; petiolis 2-4 cm. longis; petalis albis. — PERU: open 2 m. bush of montana, Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27-Sept. i, 1923, Macbride 5811 (TYPE, Field Museum); 3 m. shrub, La Merced, Aug. 10-24, 1923, Macbride 5507. The widely distributed typical form of this well-marked species has subsessile leaves with the petioles often only 0.5 cm. long, and red flowers. The specimens cited are intermediate in leaf-character between M. nervosa (Sm.) Triana and M. pseudo-nervosa Cogn. and suggest that the latter also should be treated as a variety of the former. M. pseudo-nervosa differs from M. nervosa chiefly in its long-petioled leaves that are obtusish at base. Its flowers are white. Miconia Malatestae, spec, nov., robusta, parce ramosa, 3 m. alta, fere glaberrima; ramis obtuse tetragonis, superne sulcatis glabris vel minutissime furfuraceis, ad nodos firme annulatis; foliis glabris, carneo-coreaceis intense viridibus supra subnitidulis, in sicco flavescentibus, integerrimis vel obscure et remote ciliato-denticula- tis, oblongo- vel ovato-ellipticis, obtusis vel breviter acutis, basi satis attenuatis et in petiolum brevem crassum decurrentibus, interdum valde disparis, plerumque circa 2 dm. longis et i dm. latis, nervulo marginali praetermisso 3-nerviis; nervis supra vix notatis subtus valde conspicuis sed nervulis transversallibus non prominulis; petiolis crassis, 1.5-2 cm. longis; paniculis late pyramidatis 1-2 dm. longis; floribus brevissime pedicellatis, 5-meris; calyce glabro breviter acute lobato circa 4 mm. longo; petalis albis late obovatis 3 mm. longis; staminibus 4.5 mm. longis; antheris linearibus ut videtur i-porosis, basi breviter bituberculatis fere 3 mm. longis; connective infra antheram circa i mm. producto; stylo circa 5 mm. longo; stigmate peltato; ovario apice inappendiculata ; fructu 6 mm. crasso, purpureo. — PERU: coarse-stalked shrub of montana, Hacienda Villcabamba on Rio Chinchao, Dept. of Huanuco, July 17-26, 1923, Macbride 5176 (TYPE, Field Museum). It is unusual for a species of Miconia to have the anther-connec- tive greatly prolonged but there is no doubt that such is the case here: the filament itself is linear, not tapering, and shorter than the connective. The relationship of M. Malatestae may be with M. stipularis Naud. or M. scutata Gleason (of the section Amblyarrhena) which it resembles vegetatively but its anthers are more nearly characteristic of the section Eumiconia. The leaves, or the stamens and ovary of the species mentioned are entirely different from those of our plant. M. Malatestae is named for Sr. Enrique Malatesta of Huanuco in grateful recognition of the courtesies extended by him to the 182 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV members of the Captain Marshall Field Botanical Expeditions to Peru in 1922 and in 1923 particularly when enjoying the hospitality of his Hacienda Villcabamba where this interesting and handsome- foliaged plant was collected. Miconia modica, spec, nov., arbuscula circa 8 m. alta; ramulis obtuse tetragonis sulcatis, petiolis paniculis calycibusque pilis fur- furaceo-stellatis fulvescentibus breviter denseque obtectis; foliis breviter 5-6-plinerviis submembranaceis integerrimis vel obscure remoteque calloso-repando-denticulatis, ovato-ellipticis, basi obtusis vel subrotundatis apice sensim breviter obtuseque acuminatis, plerumque circa 2 dm. longis et i dm. latis, supra glabris, nervis non prominentibus, subtus plus minusve fulvo-stellato-furfuraceis praecipue ad nervos nervulosque, nervis cum nervulis transversali- bus prominentibus, mediano multo crassiore, 3 interioribus saepius 4-8 mm. supra basin folii abeuntibus, 2-4 lateralibus exterioribus plus minusve distincte a basi; petiolis 4-7 cm. longis; paniculis multifloris; pedicellis 1-2 mm. longis; fioribus 5-meris; calyce circa 3 mm. longo; petalis albis, 4 mm. longis, puberulis; filamentis stylo- que glanduloso; antheris oblongo, haud ventricosis, vix 2 mm. longis; stylo 4-5 mm. longo; stigmate capitellato, i mm. lato. — PERU: evergreen forest, Choimacota Valley, Prov. of Huanta, Dept. of Ayacucho, Feb. 28-March 10, 1926, Weberbauer 7552 (TYPE, Field Museum). M. floribunda (Bonpl.) DC., to which this species is closely related, has more densely pubescent leaves borne on petioles 2-4 cm. long, longer anthers bulged at the base and a broadly peltate stigma. Miconia Pulgari, spec, nov., arbuscula, fere glaberrima; ramis glabris, gracilibus, teretibus; ramulis petiolis pedunculisque glabris vel interdum sparse cum pilis crispulis paullo pubescentibus; foliis membranaceis integerrimis vel minutissime adpresseque ciliatis, oblongo-lanceolatis, subabrupte obtuse acuminatis, basi acutis, supra glabris, subtus ad nervos nervulosque parce piloso-hirtellis caeteris glabris, plerumque circa i dm. longis et circa 3 cm. latis, 3-plinerviis (nervulo marginali praetermisso) ; nervis lateralibus exterioribus a basi, interioribus circa 5 mm. supra basin folii abeuntibus, nervis supra vix notatis, subtus cum nervulis transversalibus et venulis bene prominentibus; petiolis 8-10 mm. longis; paniculis densifloris 4-6 cm. longis; fioribus 5-meris; pedicellis circa i mm. longis; calyce glabro vel minute parceque granuloso-stellato, denticulate, fere 4 mm. longo; petalis late obovatis, circa 2.5 mm. longis; antheris anguste obovoideis, apice paullo arcuatis, minute i-porosis, circa 1.5 cm. longis; stylo paullo exserto, circa 5 mm. longo, apice sub- capitallato. — PERU: Huacachi, near Muna, Dept. of Huanuco, May 2o-June i, 1923, Macbride 4145 (TYPE, Field Museum). This straggling shrub with pinkish-white flowers belongs in the section Amblyarrhena and somewhat resembles several species but PERUVIAN MELASTOMES 183 is entirely distinct by virtue of its triplinerved leaves the few other species having this character being altogether different in other respects. Sr. Francisco Pulgar of Panao, by his friendly interest and influence assisted materially in the successful culmination of the difficult trip to Muna and Pozuzo on which journey this fine bush was collected. Miconia sphagnophila, spec, nov., arbuscula parce ramosa circa i m. alta; ramis ramulis petiolis paniculis foliisque subtus setis longiusculis patulis plerumque basi valde incrassatis dense rufo- pubescentibus ; foliis breviter s-y-plinerviis, rigidis, supra bullis conicis longe setuliferis dense obtectis, ovato-ellipticis, basi rotunda- tis, apice obtusis vel paullo acutis, plerumque 6 cm. longis et 3.5-4 cm. latis; petiolis 8-10 mm. longis; paniculis confertifloris; floribus subsessilibus, 5-meris;calyce 4 mm. longo; petalis albis circa 2.5 mm. longis; filamentis glanduloso-pilosis; antheris fere 2 mm. longis, minute i-porosis; stylo peltato. — PERU: sphagnum-montana, Play- apampa, Dept. of Huanuco, June 16-24, 1923, Macbride 4503 (TYPE, Field Museum). This species is related to M. hamata Cogn., M. hirta Cogn. and other allies of M. capitellata Cogn. of the section Amblyarrhena, from all of which it at least differs in the much longer calyx and smaller leaves. Its pubescence seems to resemble that of M. radula Cogn. rather closely but that species has 5 -nerved leaves. Miconia trichrona, spec, nov., ut videtur arbuscula; ramis ramu- lis petiolisque pilis crispis firmiusculis patulis plus minusve dense obtectis; foliis rigidis tenuissime crenulatis, ovatis vel elliptico- ovatis, basi rotundatis, apice aliquid angustatis subrotundatis vel late obtusis, haud acutis, plerumque 8 cm. longis et 4-5 cm. latis, supra bullis crispulis setuliferis, subtus brevissime denseque crispe villoso-hirtellis, s-y-nerviis; nervis utrinque mediocriter prominen- tibus; petiolis 1.5-2 cm. longis; paniculis congestifloris circa 7 cm. longis; floribus sessilibus, ad apices ramulorum glomerulatis ; calyce breviter denseque setuloso, circa 2 mm. longo, minute denticulato; petalis fere 2 mm. longis; filamentis glabris; stylo 4 mm. longo, parce piloso; stigmate capitellato. — PERU: above Tabaconas, Prov. of Taen, Dept. of Cajamarca, May, 1912, Weberbauer 6309 (TYPE, Field Museum). Three members of the section Amblyarrhena seem to be about equally related to this species, namely M . scabra Cogn., M. asperrima Triana, and M. Ruizii Naud. M. trichrona differs from the first and second in the quality of the pubescence, from the second and third in the obtuse leaves and also from the third in the obscurely lobed calyx. 184 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV MICONIA GLABERRIMA (Schlecht.) Naud., var. australis, var. nov., foliis basi abrupte obtusis vel subemarginatis ; paniculis 3-4 cm. longis, minutissime parceque furfuraceo-puberulis. — PERU: open, 1-2 m. shrub, Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27- Sept. r, 1923, Macbride 5769 (TYPE, Field Museum); also 57/7. Typical M. glabenima (Schlecht.) Naud. of Mexico and Central America has leaves that are acutish or at least somewhat narrowed at base and often a rather large and glabrous panicle. The variety australis is not well-marked but in view of its geographic separation its apparent differences seem worth recording. Miconia Miles-Morgani, spec, nov., arbuscula, 0.5-1 m. alta; ramis gracilibus fere teretibus plus minusve furfuraceo-stellatis demum glabris; ramulis petiolis pedunculisque breviter denseque furfuraceo-stellatis; foliis membranaceis obscure undulato-crenula- tis late ovatis vix vel haud acutis, basi rotundatis vel leviter corda- tis supra minutissime parceque stellatis vel vetustioribus glabris et plus minusve reticulato-bullatis, subtus subsparse stellatis ad ne vos nervulosque etiam conspicue reticulatis, 5-nerviis, plerumque circa 5 cm. longis et 3 cm. latis; nervis paullo prominentibus praecipue 2 marginalibus ; petiolis gracilibus 1-1.5 cm- longis; paniculis anguste pyramidatis circa 8 cm. longis; pedicellis i mm. longis; floribus interdum cernuis, 5-meris; calyce fere glabro, obscure denticulate circa 2 mm. longo; petalis albis circa 2 mm. longis; antheris sub- cuneiformis, late 2-porosis 1.5 mm. longis; stylo 2 mm. longo; stig- mate peltato. — PERU: on trail to Tambo de Vaca from Muna, Dept. of Huanuco, June 5-7, 1923, Macbride 4318 (TYPE, Field Museum). This attractive bush with fragrant white flowers is closely related to M. peruviana Cogn. (Sect. Cramaniwri) but the latter has narrower acute leaves and hirtellous rather than stellate pubescence. Mr. Miles Morgan of the Cerro de Pasco Copper Corporation and other officials repeatedly aided in the work of the members of the Captain Marshall Field Botanical Expeditions to Peru in 1922 and 1923, and appreciation of their help is recorded gratefully. Mr. Morgan in particular gave generously of his personal time and his interest may be commemorated fittingly in the name of this new species. Miconia opacifolia, spec, nov., arbuscula circa 3 m. alta, laminis foliorum supra scabris exceptis glaberrima; ramis obtuse tetragonis superne plus minusve sulcatis, subgracilibus ; foliis sessilibus mem- branaceis in sicco supra flavescentibus, subtus aliquanto purpur- ascentibus, obscure et remote undulato-denticulatis, ovato-oblongis vel -lanceolatis, basi cordatis, apice sensim acuminatis, 1.5-2 dm. longis, plerumque 7 vel 8 cm. latis, 5-7-nerviis; nervis supra leviter PERUVIAN MELASTOMES 185 impressis, subtus mediano valde prominentibus, exterioribus caeteris multo gracilioribus ; paniculis pyramidatis, circa i dm. longis; flori- bus 5-meris, minutis, brevissime pedicellatis; calyce breviter lobato vix i mm. longo; petalis ut videtur albis; antheris obovoideis, 2- porosis, truncatis; stylo incluso. — PERU: Hacienda Villcabamba on Rio Chinchao, Dept. of Huanuco, July 17-26, 1923, Macbride 5197 (TYPE, Field Museum). This species belongs to the section Cremanium and most re- sembles M. lilacina Triana which has smaller glabrous entire leaves and larger flowers. The leaves of M. opacifolia, notwithstanding the minute scabrosity on the upper surface, were velvety in appear- ance before drying. Miconia Ottikeri, spec, nov., arbuscula circa i m. alta; ramis ramulisque glabris acute tetragonis subquadrialatis ; foliis coraceis minute calloso-cilato-denticulatis, ovatis, acuminatis, basi subrotun- datis, supra glabris, subtus flavescentibus ad nervos (imprimis mediano) nigro-pilosis caeteris glabris, plerumque circa 7 cm. longis et 3-3.5 cm. latis, definite 3-nerviis, nervulo marginalo fere obsoleto praetermisso ; nervis supra leviter impressis, subtus prominentibus, mediano multo crassiore, venis ramuloso-reticulatis in foliis juniori- bus conspicuis sed demum obsoletis; petiolis 1.5-2 cm. longis, supra cum pilis nigris longiusculis firmis pubescentibus; paniculis anguste laxis circa i dm. longis; pedicellis circa 2 mm. longis; calyce glabro, subhemisphaerico, breviter 5-lobato, 3-4 mm. longo; floribus ignotis; bacca rubra, 5-6 mm. crassa. — PERU: Tambo de Vaca, Dept. of Huanuco, June 10-24, 1923, Macbride 4430 (TYPE, Field Museum). In spite of the lack of flowers I have little hesitancy in proposing this shrub as a new species because its general characteristics so obviously ally it to M. flavescens Cogn. and other members of the section Chaenopleura, to none of which, it can be referred. Its sharply angled branches and the curious restriction of the pubescence to the leaf -veins and the petioles are unusual characters for the group. In naming this neat shrub for Mr. Ottiker, an official of the Ferrocarril Central del Peru, I am recording with pleasure his very welcome aid in the matter of transport from Lima to La Oroya. Miconia ramosipila, spec, nov., arbuscula vel interdum arbor; ramis superne obtuse tetragonis, sulcatis, plus minusve cum pilis plumosis breviter pubescentibus; petiolis pedunculis ramulisque in- florescentiarum pilis plumoso-penicellatis stellulatisque fulvescenti- bus intermixtis dense obtectis; foliis integerrimis vel parce adpresse- que breviter ciliatis, submembranaceis, supra glabris, nitidulis et obscure bullatis, subtus ad nervos cum pilis fulvo-plumoso-penicella- tis dense pubescentibus, ad nervulos transversalibus et etiam caeteris mediocriter fulvo-stellatis, elliptico-ovatis, subabrupte acutis, 1 86 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV basi subrotundatis vel emarginatis, 1.5-2 dm. longis, 8-10 cm. latis, 7-nerviis; nervis supra vix notatis, subtus 5 valde prominentibus, reliquis 2 marginalibus cum transversalibus parum prominulis; petiolis usque 3-7 cm. longis; paniculis anguste pyramidatis, parce ramosis, ramulis spicatis; floribus glomerulatis, 5-meris; calyce glabrato, s-lobato, vix 1.5 mm. longo; petalis vix i mm. longis; antheris minutissme porosis, sublinearibus vix 0.5 mm. longis, conn- ective distincte breviterque producto; stylo i mm. longo; stigmate peltato. — PERU: in montana, trail from Muna to Tambo de Vaca, Dept. of Huanuco, June 5-7, 1923, Macbride 4321 (TYPE, Field Museum). Apparently this is a species of the section Cremanium and is related in a general way to M. valida Cogn. and allies, in spite of the rather narrow anthers. No species of Eumiconia seems, from description, to resemble M. ramosipila at all closely. Miconia saxatilis, spec, nov., arbuscula circa 3 m. alta; ramis obtuse, ramulis acute tetragonis, ad nodos incrassatis; ramis ramulis petiolis pedunculis ramulisque inflorescentiarum sordide denseque stellatis ; foliis integerrimis vel superne undulato-subdenticulatis late ovato-ellipticis basi et apice subabrupte angustatis, apice etiam breviter acuminatis plerumque 15-18 cm. longis et 7.5-9 cm. latis, membranaceis, supra primum glabris viridibusque, subtus pallide viridibus et etiam tenuiter ad nervos venasque satis stellatis, breviter subtriplinerviis (nervulo marginale praetermisso) ; nervis lateralibus exterioribus a basi, interioribus 4-8 mm. supra basin folii abeuntibus, nervis interioribus praecipue subtus cum nervulis transversalibus mediocriter prominentibus; petiolis 2-4.5 cm- longis; paniculis densifloris circa 8 cm. longis; floribus minutissimis 5-meris, brevis- sime pedicellatis ; calyce glabrato, obtuse denticulate vix i mm. longo; petalis vix 0.5 mm. longis; antheris subcuneiformiis fere 0.5 mm. longis, 2-porosis; stylo circa i mm. longo; stigmate peltato. —PERU: wet rocky upland, Playapampa, Dept. of Huanuco, June 16-24, 1923, Macbride 4881 (TYPE, Field Museum). This slender shrub with tiny flowers perhaps is most nearly re- lated to M. micrantha Cogn. of the section Cremanium which, how- ever, has glabrous leaves. Miconia sulcata, spec, nov., arbor parva tota glaberrima; ramis junioribus acute tetragonis et valde quadrisulcatis, robustis; foliis membranaceis integerrimis vel obscure remoteque repando-denti- culatis, ovato-ellipticis, apice subrotundatis obtusis, basi plus minusve sensim attenuatis, ut videtur plus minusve disparis, majoribus 3 dm. longis et fere 1.5 dm. latis, supra viridibus, subtus aliquid purpurascentibus, 5-nerviis; nervis supra haud prominentibus, subtus 3 interioribus valde prominentibus, reliquis 2 marginalibus parum prominulis, nervulis transversalibus gracilibus, ramuloso-reticulatis ; PERUVIAN MELASTOMES 187 petiolis 5-8 cm. longis; paniculis late pyramidatis, multifloris ; floribus minutis, subsessilibus, immaturis ut videtur 5-meris; calyce minute denticulate, circa i mm. longo; antheris evidente obovoideis; stylo ut videtur punctiformis. — PERU: Chanchamayo Valley, Dept. of Junin, Jan., 1924-1927, Carlos Schunke 423 (TYPE, Field Museum). It may seem unwise to describe as new a Miconia that is only in bud so that the floral characters are not entirely discernible but the anthers are sufficiently developed to enable one to place it in either the section Cremanium or Chaenopleura. No described species of either section, apparently, has such remarkably sulcate branches, large glabrous leaves and extremely small flowers. MICONIA THEAEZANS (Bonpl.) Cogn., var. longifolia (Cogn), comb. nov. M. theaezans (Bonpl.) Cogn., subsp. viridis Cogn., var. longifolia Cogn. Fl. Bras. 14*: 421. 1888. MICONIA THEAEZANS (Bonpl.) Cogn., var. tetragona (Cogn.), comb. nov. M. theaezans (Bonpl.) Cogn., subsp. flavescens Cogn., var. tetragona Cogn. 1. c. 421. The above variants of this variable species of Miconia are better treated, it seems to me, as varieties of the species rather than as varieties of two subspecies. This simpler classification is certainly more practical in ordinary usage whatever its disadvantages, if any, in monographic work. Miconia trichogona, spec, nov., ramis ramulisque teretibus plus minusve flexuosis ad nodos conspicue setosis caeteris glabris; foliis ovatis obscure denticulatis et subadpresse ciliatis, breviter acumina- tis, basi subcordatis, 9-13 cm. longis, 4-5 cm. latis, supra minute parceque scabris et ad nervos plus minusve pilosis, luteo-viridibus, subtus glabris saepius rufescentibus, 5-7-nerviis; petiolis supra sulcatis et dense setoso-pilosis, 1.5-2 cm. longis; paniculis pyramida- tis satis multifloris circa i dm. longis; pedicellis circa i mm. longis; bracteis subscariosis ad apicem ciliatis circa 5 mm. longis; floribus ignotis; calyce distincte 5-lobato; bacca circa 5 mm. crassa. — PERU: in montana, Hacienda Villcabamba, Rio Chinchao, Dept. of Huanuco, July 17-26, 1923, Macbride 5178 (TYPE, Field Museum). There are only a few species of Miconia with glabrous branches setose only at the nodes and none of them apparently except M, setinodis (Bonpl.) Naud. of the section Cremanium approaches our Peruvian species closely and it has 6-8-merous flowers and smaller 3-5-nerved leaves. Miconia vitiflora, spec, nov., arbuscula, circa 2 m. alta; ramis ramulisque inflorescentiarum obtuse tetragonis vel subteretibus, primum praecipue ad nodos longe plumoso-pubescentibus demum glabris; petiolis plerumque 6-n cm. longis cum pilis crispulis plumosis i88 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV pallide fulvescentibus plerumque 3-5 ram. longis dense pubescenti- bus; foliis submembranaceis supra intense viridibus, glabris, subtus plus minusve purpurascentibus, ad nervos nervulosque breviter stellato-plumoso-pubescentibus, subrotundatis vel subcordatis, apice abrupte breviter obtuseque acuminatis, plerumque 1.5-2 dm. longis et 1-1.5 dm. latis, y-nerviis; nervis supra leviter impressis, subtus cum nervulis transversalibus ramuloso-reticulatis prominentibus, mediano multo crassiore; paniculis multifloris, late pyramidatis, circa 1.5 dm. longis; floribus 5-meris, ut videtur subdioicis; pedi- cellis vix i mm. longis; calyce glabro, acute denticulate; petalis vix i mm. longis; antheris obovoideis, late biporosis, connectivo breviter producto, antice minute tuberculato; stylo i mm. longo; stigmate peltato; bacca fere 4 mm. crassa. — PERU: montana slope at Hacienda Villcabamba, Dept. of Huanuco, July 17-26, 1923, Macbride 5165 (TYPE, Field Museum). Apparently the only species of the section Cremanium related to M. vitiflora is M. polygama Cogn. and it has smaller leaves that are hirsute on both sides. Miconia brevis, spec, nov., arbuscula tota glaberrima; ramis teretibus vel junioribus obscure tetragonis; foliis subcoraceis obscure crenulato-denticulatis ovato-ellipticis, basi acutiusculis vel obtusis, apice obtusis interdum ad apicem breviter abrupteque angustatis sed haud acutis, plerumque circa 4 cm. longis et 2 cm. latis, 5-nerviis vel subtriplinerviis, nervulo marginal! praetermisso ; nervis gracili- bus supra 3 interioribus satis impressis, 2 exterioribus vix notatis, subtus 3 prominentibus reliquis 2 marginalibus parum prominulis, venis anguste reticulatis; petiolis gracilibus, 1-2 cm. longis; paniculis plus minusve cernuis, 4-5 cm. longis; pedicellis vix i mm. longis; calyce obscure 5-denticulato, 2 mm. longo; petalis 5, ut videtur albis irregulariter suborbicularis, vix 2 mm. longis; staminibus 10; fila- mentis triangularibus, 1.5 mm. longis; antheris late obovoideis, vix i mm. longis; stylo glabro, fere 5 mm. longo; stigmate capitellato. — PERU: between Huancabamba and Ayavaca, Dept. of Piura, May, 1912, Weberbauer 6334 (TYPE, Field Museum). Other species of the section Chaenopleura which most resemble this shrub vegetatively, notably M. alpina Cogn. and M. fruti- culosa Cogn., have 4-merous flowers. M. brevis seems to be well- marked, apart from its 5-merous flowers, by its short 5-nerved leaves, short pedicels, calyx, petals and stamens. Miconia Griffisii, spec, nov., arbuscula satis ramosa et foliosa, 1-1.5 m. alta; ramis teretibus vel superne subtetragonis petiolisque breviter denseque setuloso-hirtellis haud furfuraceis vel stellatis; foliis mediocriter coriaceis integerrimis vel obscure et remote calloso- ciliatis oblongo- vel ovato-ellipticis breviter obtuseque acuminatis vel acutis, basi obtusis vel vix acutis, supra glabris, subtus ad nervos nervulosque parce hirtellis caeteris glabris, plerumque circa 8 cm. PERUVIAN MELASTOMES 189 longis et 3-3.5 cm. latis, nervulo marginali praetermisso 3-nerviis; nervis nervulisque supra satis impressis, subtus 3 valde prominenti- bus, 2 marginalibus vix notatis, nervulis conspicue ramuloso-reticula- tis; petiolis 7-12 mm. longis; paniculis densifloris circa i dm. longis; floribus 4-meris; pedicellis circa 5 mm. longis; calyce turbinato- campanulato, glabrato, 4 mm. longo, distincte lobato; lobis sub- rotundatis, extus tuberculatis ; petalis albis, suborbicularis, circa 3 mm. longis; antheris obovoideis, circa 1.5 mm. longis; stylo 5-6 mm. longo; stigmate peltato. — PERU: Tambo de Vaca, Dept. of Huanuco, June 10-24, 1923, Macbride 4382 (TYPE, Field Museum). The anthers of this species are entirely characteristic of the section Chaenoplertra no member of which resembles M. Griffisii closely unless possibly the poorly described M. thyrsoidea (D. Don) Naud. which, however, is said to have tomentose branches, leaves pilose beneath on the nerves and smaller flowers. This handsome shrub with showy white flowers is named for Mr. C. N. Griffis of Lima the well-known editor and English Secre- tary to the President. He extended most helpfully the courtesies of the government to the Captain Marshall Field Botanical Ex- peditions to Peru in 1922 and 1923. Miconia trichocaula, spec, nov., arbuscula valde ramosa; ramis ramulis petiolisque dense rufo-pubescentibus cum pilis patentibus firmiusculisque 2-3 mm. longis et pilis brevisque furfuraceo-plumosis intermixtis; foliis numerosissimis obscure repando-denticulatis sub- membranaceis subrotundato-ellipticis haud acutis, supra glabris laevibusque, subtus solum ad nervos paullo furfuraceis, plerumque circa 2 cm. longis et 1.5 cm. latis, 3-nerviis; petiolis 5-8 mm. longis; racemis 1-2 cm. longis, pseudoaxillaribus, paucifloris; pedicellis 2-3 mm. longis; floribus 4-meris, plus minusve cernuis; calyce glabro 4-denticulato, fere 2 mm. longo; petalis suborbicularis vix 1.5 mm. longis; antheris obovoideis; stylo incluso vix 1.5 mm. longo; stigma peltato.— PERU: above Huancabamba, Dept. of Piura, April, 1912, Weberbauer 6094 (TYPE, Field Museum). This is an addition to the section Chaenopleura and from descrip- tion is related to M. bullata (Turcz.) Triana. The leaves of the latter, however, are characterized as bullate-tuberculate above and pilose on the veins beneath. Miconia dichrophylla, spec, nov., arbor 5 m. alta; ramulis acute tetragonis, superne valde compressis, junioribus petiolis pedunculis paniculisque dense squamuloso-pubescentibus; foliis submembran- aceis obscure undulatis vel inferne integerrimis, oblongo-ovatis vel -lanceolatis plerumque circa 1.5 dm. longis et 5 cm. latis, basi acutis, apice in caudiculam (acumine 1-2 cm. longo) subabrupte contracto, supra viridibus laevibusque, subtus argenteis pilis lepidotis adpressis i go FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV ad medium (praecique ad nervos nervulosque) saepius fulvo-puncta- tis dense vestitis, 3-plinerviis; petiolis 5-7 mm. longis; paniculis anguste pyramidatis paullo ramosis 0.5-1 dm. longis; paniculae ramis simplicibus vel inferne bi-trifidis 1-2 cm. longis, subsecundi- floris; floribus 5-meris sessilibus; calyce campanulato circa 2 mm. longo, leviter 5-costato, minutissime dentato, squamulis fulvis et albidis lepidotis utrinque tecto; petalis suborbicularis circa 1.5 mm. longis, glabris; antheris oblongo-linearibus, apice paullo attenuatis, ut videtur minute uniporosis circa i mm. longis, connective infra loculos non producto, basi antice obscure bigibboso, postice breviter calcarato; stylo vix clavato, truncato, vix 1.5 mm. longo. — PERU: slender tree of montana, La Merced, Dept. of Junin, Aug. 10-24, 1923", Macbride 5495 (TYPE, Field Museum). The flowers of this species are not fully developed and conse- quently I am not positive as to their characters : certainly, however, the anthers are spurred posteriorly and this characteristic apparently eliminates the possisibilty of referring M. dichrophylla to either M. lepidota DC. or M. fulva (Rich.) DC. to both of which it bears at least some superficial resemblance. Miconia nectaria, spec, nov., arbor 5-7 m. alta, glaberrima; ramis ramulisque teretibus vel superne plus minusve compressis; foliis subcoriaceis ubique pallide viridibus integerrimis, anguste vel oblongo-ovato-lanceolatis, apice sensim et longe acuminatis, basi acutis vel subobtusis, 1.5-2 dm. longis, 4-6.5 cm. latis, 3-nerviis vel subtriplinerviis ; nervis supra leviter impressis, subtus cum nervulis transversalibus satis prominentibus ; petiolis i vel 2 cm. longis; petiolo etiam lamina basin versus 3-5 nectariis cupiliformis stipita- tis instructo; nectariis 2-5 mm. latis; paniculis mediocribus, pyrami- datis, 1-1.5 dm- longis; pedicellis circa 2.5 mm. longis; floribus ignotis; calyce 5-lobato; lobis ovatis; bacca subglobosa, 3 mm. crassa; semina pyramidata. — PERU: slender tree of montana slope, Hacienda Schunke, La Merced, Dept. of Junin, Aug. 27-Sept. i, 1923, Macbride 5738 (TYPE, Field Museum). This species and the following are exceptionally well-marked by the nectar-like appendages that are borne with uniform regularity on the petioles and leaf-bases. Miconia Aspiazui, spec, nov., M . nectaria affinis; ramis ramulisque satis compressis; foliis membranaceis integerrimis ellipticis, basi breviter acutis vel attenuatis, apice, subabrupte longo-acuminatis vel fere caudatis 1.5-2 dm. longis, 8-12 cm. latis, 5-nerviis vel sub-5- plinerviis; petiolis circa 3 cm. longis; paniculis 1.5-2 dm. longis, multifloris; ramulis spiciformis; floribus ignotis; calyce ut videtur minute 5-denticulato; bacca subglobosa, circa 5 mm. crassa. — PERU: La Merced, Dept. of Junin, Aug. 10-24, 1923, Macbride 5586 (TYPE, Field Museum). PERUVIAN MELASTOMES 191 This species has the same curious appendages on the petioles that are described as occuring on the probably closely related M. nectaria. M. Aspiazui, however, is clearly distinct by virtue of its very different foliage. It is named for Dr. R. Aspiazu, distinguished physician and surgeon of Lima, and enthusiastic student of the local flora who generously aided the work of the Captain Marshall Field Botanical Expeditions to Peru. Icaria, gen nov. Flores 4-meri. Calycis turbanato-campanulatus ; limbus dilatatus, lobatus. Stamina aequalia, breviuscula; filamentis late alatis; antherae breves, ut videtur longitudinaliter i-2-rimosae, connective sub loculis nullo vel non elongate, lateraliter in appendi- cem latam dilatatam emarginatam fere erecto producto. Ovarium liberum. Bacca sicca coriacea. — Frutex glabratus, caule elongate, tortuoso, adscendente, teretiusculo. Folia petiolata, 3-nervia. Flores mediocres, pedicelati in paniculas parvas terminates depositi. — Affinis videtur Miconiae. Icaria fictilis, spec, nov., fruticosa laxe ramosa; ramis ut videtur suberecto-patentibus flexuosis teretibus glabris, vel junioribus ramulis petiolisque obscure furfuraceis; foliis integerrimis vel minu- tissime sparseque ciliatis, glabris vel subtus indestincte furfuraceis, late ovato-ellipticis, basi subcordatis, apice vix acutis, plerumque 3-4 cm. longis et 2-2.5 cm- latis, 3-nerviis; nervis subtus paullo prominentibus, nervulis transversalibus haud ramuloso-reticulatis ; petiolis gracilibus 1-1.5 cm- longis; paniculis paucifloris racemiformis, 5-7 cm. longis; pedunculis ramulisque inflorescentiarum breviter stellato-furfuraceis ; floribus longe pedicellatis ; calyce circa 7 mm. longo, subtiliter stellato-furfuraceo, tubo turbinato, limbo dilatato, 4-lobato, extus minutissime tuberculato; lobis late ovatis; petalis circa 5 mm. longis et 6 mm. latis; staminibus fere 6 mm. longis; filamentis 2 mm. latis; antheris obovoideis; appendicibus 1.5 mm. longis, i mm. latis, apice irregulariter bidentatis; stylo 4 mm. longo; stigmate peltato; bacca (immatura) 5 mm. crassa, calycis lobis persistentibus coronata. — PERU: in patches on steep shady slope, Tambo de Vaca, Dept. of Huanuco, June 10-24, !923> Macbride 44 j i (TYPE, Field Museum). The aspect of this shrub is that of several species in the section Chaenopleura of Miconia but the broadly winged filaments and the curious wing-like erect appendages at either side of the anther at its base are characters that are entirely at variance with the generic character of that genus. The fruit, however, is undoubt- edly baccate which places the plant in the tribe Miconieae although in this tribe highly specialized stamineal development is most unusual. I am indebted to Dr. Paul C. Standley for suggesting the generic name. 192 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. IV 5. VARIANTS OF OSTRYA AND A TEXAS TEPHROSIA OSTRYA VIRGINIANA (Mill.) K. Koch, forma glandulosa (Spach), comb. nov. O. mrginica (Moench.) Willd., var. glandulosa Spach, Ann. Sci. Nat. 2 se'r. 16: 246. 1841. 0. mrginiana (Mill.) K. Koch, var. glandulosa (Spach) Sarg. Bot. Gaz. 62: 216. 1919. My attention has been called to this form — characterized by the presence, in greater or less abundance, of stiffish glandular trichomes intermixed with the usual substrigose pubescence of the petioles, young branchlets and peduncles — by my own collection, number 7000 from Trout Park, Elgin, 111., Aug. 30, 1926. Examina- tion of the series of specimens in the Illinois Herbarium of the Field Museum indicates that this variant is not at all uncommon and apparently grows together with the typical eglandular state of the species, as observed for Indiana by Mr. Deam, Trees of Ind., Dept. Conserv. Publ. 13: 80. 1921. As there appear to be no other differences associated with the glandulosity, it seems better to regard it as merely a glandular form. House, N. Y. State Mus. Bull. 254: 267. 1924 lists it for western New York with the remark "is scarcely worthy of varietal rank." It seems to grow with the typical form in general throughout the northern portion of the species' range but the specimens of forma glandulosa that I have seen from New England, New York and Pennsylvania are usually less glandular than many, at least, of the collections from farther west. In this connection I have studied the specimens in the herbarium of the Field Museum from the southern area of the species' range and have reached the conclusion of Winkler, Pflanzenr. 4. 61: 22. 1 904, that the Mexican and Central American plants with somewhat more pubescent more nearly oblong more or less doubly serrate leaves are too nearly approached by material from the southern states and accordingly are better regarded as constituting a geo- graphical variety, that even then is often obscurely marked. Stand- ley, Contrib. U. S. Nat. Herb. 23: 169. 1920, considers the Mexican plant, under the name O. guatemalensis (Winkl.) Rose, "very closely related to O. mrginiana but seems fairly distinct." However, he describes the leaves as "ovate" which is indicative of their vari- ability since the term is applicable particularly to the form proposed as O. mexicana Rose, very properly referred by Standley to 0. guatemalensis. If one regards 0. mrginiana as distinct from the Old World 0. italica and the Mexican plant as a variety of the former, its name and synonymy are as follows: PERUVIAN MELASTOMES 193 OSTRYA VIRGINIANA (Mill.) Koch, var. guatemalensis (Winkl.), comb. nov. O. italica Scop., subsp. virginiana (Mill.) Winkl., var. guatemalensis Winkl. Pflanzenr. 4: 61 : 22. 1904. O. guatemalensis (Winkl.) Rose, Contrib. U. S. Nat. Herb. 8: 292. 1905. O. mexicana Rose, I.e. TEPHROSIA ONOBRYCHOIDES Nutt. Journ. Acad. Phil. 7: 104. 1834; T. & G. Fl. N. Am. i: 292. 1838. — TEXAS: Dallas, June, 1877, Reverchon; Fayette Co., 1892, E. W. Crawford 47; Houston, June1 17, 1926, G. Eifrig. INDIAN TERRITORY: Limestone Gap, July 4, 1877, Geo. D. Butter 115. LOUISIANA: Dr. Hale. TEPHROSIA ONOBRYCHOIDES Nutt., var. texana (Rydb.), comb, nov. Cracca texana Rydb. N. Am. Fl. 24: 176. 1923. — TEXAS: Hemstead, July i, 1872, E. Hall 119; Hockley, 1890, W. F. Thurow; Fayette Co., 1891, H. Wurzlow; Houston, July 13, 1926, G. Eifrig. LOUISIANA: Alexandria, Hale. Dr. Rydberg's Cracca texana, I.e., is a fairly well marked herb- arium species by virtue of the appressed strigose character of its pubescence especially on the calyx and under surfaces of the leaves. The pubescence of typical T. onobrychoides is more pilose in quality and mostly spreading. However, there is some intermixture of strigose hairs. Furthermore, as shown by the above citations, col- lections have been made in the same localities that, except for this somewhat variable difference in tjie nature of the pubescence, appear to be indistinguishable. Accordingly it seems that the character of T. onobrychoides can be most logically defined by including C. texana in it as variety. THE LIBRARY OF THE JUL241943 UNIVERSITY OF ILLINOIS UNIVERSITY OF ILUNOIS-URiANA 30112049897561