vi.U hte 1/ FORTHE PEOPLE FOR EDVCATION FOR SCIENCE LIBRARY OF THE AMERICAN MUSEUM OF NATURAL HISTORY THE SPORTING FISH OF GREAT BRITAIN 1^ ■^ ^ v2. -' ^ ■g- o s ^-1 < * * UQ ■ w 4 i THE SPORTING FISH OF GREAT BRITAIN lA NOTES ON ICHTHYOLOGY ^ NOTES ON ICHTHYOLOGY BY I^CHOLMONDE LEY-PEN NELL LATE HER MAJESTY'S INSPECTOR OF SEA FISHERIES AUTHOR OF " Saititott and Trojit," " Pfke unci other Coarse Fish'^ *' The Modt'vn Practical Angler^* " The Angler-Natitralist^^ " The Book of the Pike," &c. Editor of the late ^' Fishcr- jnau's Magazine and Pericw.'* ILLUSTRATED BY SIXTEEN LITHOGRAPHS OF FISH IN GOLD, SILVER, AND COLOURS. LONDON SAMPSON LOW, MARSTON, SEARLE, AND RIVINGTON Crown Buii-DInos, i88, Fleet Street 1886 {All rights reserved) 1,11 I ■■ 1 1 I , I CIIISWICK press:— C. WHITTINGHAM and CO., TOOKS COl'RT, CHANCERY LANE. c^^^ ^™g l^^^^^a ' i^ ^^g f^^M '^^ v^^"^ ^^'^'> lis ^ ^R ks^SlR m^^ ^^ S^jjj PREFATORY NOTE. jHIS attempt to represent the principal fresh-water fish of the British Islands, at least approximately in their actual forms and colours, may not, perhaps, be unwelcome to lovers of Natural History. The Chapter on the Outlines of Ichthyology will, I hope, be useful to Anglers and students. To the author of Yarrell's " British Fishes," as also to Mr. J. Van Voorst, the publisher of that excellent and exhaustive treatise, I gladly take this opportunity of acknowledging my many obligations. H. C.-P. ^^^^^ @ 3^^ M ^m i^ i B i m CONTENTS. HE Sat.mon The Bull Trout The Sea Trout (or Salmon-Trout) . The Common Trout The Great Lake Trout The Grayling .... The Pike The Perch The Carp The Tench The Barbel The Bream The Roach The Dace and the Chul; The Bleak Outlines of Ichthvolooy 147 I 32 37 47 58 62 69 86 94 108 116 12 2 128 138 14+ .,85 LIST OF ILLUSTRATIONS OF FISH. FACE HE Salmon i The Bull Trout 32 The Sea Trout 37 The Common Trout 47 The Loch Leven Trout 52 The Great Lake Trout 5S The Grayling 62 The Pike 6(> The Perch 86 The Carp 94 The Tench .... 108 The Barbel 116 The Bream 122 The Roach 12S The Rudd 132 The Dace 13S The Chub 140 The Bleak 144 THE SALMON.' ^NTIL within the last fifty years very httle was known of the natural history of the Salmon, the united lore of those most interested in the fisheries amount- ing to little beyond the fact that the fish ascended the rivers to spawn during the spring and summer — spawned — and descended again to the sea within the followino- two or three months. Since the period referred to, however, and espe- cially during the last two or three decades, the researches of ichthyologists and the experiments which have been conducted on a large scale by enterprising and scientific men have thrown a flood of light upon the subject, converting doubts into certainties, theories into practice, and gene- rally advancing our knowledge of the subject to a point which has already produced practical results of great importance in the re-stocking of our ex- ' Salino sa/iir. Salwo, Lat. for a salmon ; sa/nr, (tqm the Latin sa/arins. of salt. K 2 THE SPORTING FISH hausted salmon rivers, and the increase of a valuable article of the national food-supply. I may perhaps be permitted to refer with some satisfaction to my own small share in bringing about the legislative reforms to which the present material prosperity of our fisheries is of course primarily attributable. It is now some years, since, in an article on this subject in the Saturday Revieiv, it was stated, " Mr. Pennell, Mr. Buckland, and a few others, have worked hard in the cause of the Salmon, in spite of territorial apathy, and if we are ever again to have this fish as plentiful as it was in the days of the mythical apprentices, it will be mainly owing to their exertions." The following sketch, which I originally pub- lished in the Times, gives in a summarised form the history of the Salmon as it was then known. It cannot, I think, be said that subsequent experi- ments and investigations have in any material point added to or taken from it. Proved Facts in the History of the Salmon. 1 . Salmon and Grilse invariably spawn in fresh water if possible, — both the eggs, and the young fry whilst in the Parr state, being destroyed by contact with salt water. 2. The eggs are usually deposited on gravelly shallows, wliere they hatch in from So to 140 days. OF GREAT BRITAIN. 3 according to the temperature of the water. Eggs remaining unhatched beyond the latter period will seldom hatch at all, possibly from having been destroyed by the low temperature. 3. The eggs deposited by the female will not hatch under any circumstances unless vivified, after exclusion, by the milt of the male ; and — at least up to the period of migration — there is no difference whatever in fry bred between Salmon only, between Grilse onl)', between Salmon and Grilse, between Salmon and Parr, or between Grilse and Parr. ' The female Parr cannot spawn ; but the male Parr possesses, and constantly exer- cises, the power of vivifying Salmon and Grilse eggs. \_Note '. — This is to be understood as referring to the specific characteristics of any of the Sahiion thus bred. It seems very probable that there may be in fry variations of size or development depending on their parentage or generation, as there are also known to be differences in the size of the eggs of different breeding fish dependent upon the size and age of the latter.] 4. The fry remain one, two, and, in some cases, three years in the rivers as Parr before going down to the sea, — about half taking their departure at one year, nearly all the others at two years, and the remainder (which are exceptional) at three years old. 5. All young Salmon-fry are marked with bluish 4 THE SPORTING FISH bars on their sides until shortly before their migra- tion, up to which period they are Parrs ; they then invariably assume a more or less complete coating of silvery scales and become Smolts, — the bars, or Parr-marks, however, being still clearly discernible on rubbing off the new scales. 6. The young of all the species here included in the genus Sahuo have at some period of their ex- istence these bluish bars ; and consequently such marks are not by themselves proofs that fry bearing them are the young of the true Salmon {Salmo salar). 7. Unless the young fish put on their Smolt dress in May or early in June, and thereupon go down to the sea, they remain as Parrs another year; and without Smolt scales they will not migrate, and cannot exist in salt water. & 8. The length of the Parr at six weeks old is about an inch and a half or two inches; and the weight of the Smolt before reaching the tidal wave from one to two ounces. 9. In at least many cases, Smolts thus migrating to the sea in May and June return as Grilse, some- times within five, generally within ten weeks, the increase in weight during that period varying from 2 to 10 lbs., the average being from 4 to 6 lbs. ; OF GREAT BRITAIN. 5 and these Grilse spawn about November or De- cember— go back to the sea — and (in many- cases) re-ascend the rivers the next spring as Salmon, with a further increase of 4 to 12 lbs. Thus, a fish hatched in April, 1854, and marked when migrating in May, 1855, was caught as a Salmon of 22 lbs. weight in March, 1856. 10. It appears certain, however, that Smolts do not always return during the same year as Grilse, but frequently remain nine or ten months in the sea, returning in the following spring as small-sized Salmon. \_NoU. — It will thus be seen that the fry of the Salmon are called Pans, or Parr, until they put on their migratory dress, when they become Sino/ts and go down to the salt water; Grilse if they return during the first year of their migration ; and at all other periods Sctlmon.^ 11. It has also been clearly proved that, in ge- neral, Salmon and Grilse find their way back to spawn to the rivers in which they were bred — some- times to the Identical spots, — spawn about Novem- ber or December, — and go down again to the sea as "spent fish," or " Kelts," in February or March, — returning, in at least many cases, during the following four or five months as " clean fish," and with an increase in weight of 7 to 10 lbs.' ' Shortly before spawning, and whilst returning to the sea as Kelts, or spent fish, Salmon are unfit for food, and their capture 6 TBE SPORTING FISH This, in a condensed form, is at the present time — as it was when first pubHshed — a fair statement of our positive knowledge as regards the leading facts in the history of the Salmon ; and it may be doubted if any subsequent discoveries — however they may add to or amplify — will be found to mate- rially modify them. In " Fact, No. lo" will be found the first statement of the now very generally ad- mitted theory of a double or divided migration. The several species belonging to the genus Salmo, and properly coming within the scope of these pages as British Sporting Fish, may be broadly divided into three groups : — 1. The Silver, or Migratory species {i.e. those migrating to or from the sea) ; 2. The Yellow, or Non-migratory species ; and, 3. The Charrs, or Orange- and Red-coloured species ; the general colouring of the first being always more or less greyish-silver, that of the second golden or yellow, and that of the third, espe- is then illegal. " Foul fish " before spawning are, if males, termed Red fish, from the orange-coloured stripes with which their cheeks are marked, and the golden-orange tint of the body ; the females are darker in colour, and are called Black fish. After spawning the males are called Kippers, and the females Shedders or Baggits. OF GREAT BRITAIN: 7 daily on the under part of the body, crimson and orange of various degrees of brilliancy. The species of the Silver, or Migratory group, are three in number ; which include, according to the opinions of ichthyologists, the whole of our migra- tory Salmonidcc, under whatever local names they may occur : The true Salmon (Parr, Smolt, Grilse) Salmo salar ; The Bull-Trout, also called Grey Trout, Sewin, and Roimdtail — Salmo criox ; and The Sea-Trout or Salmon-Trout (sometimes also called White Trout) — Salmo trutta. In the few further observations which limits of space permit of my offering on the general history of the true Salmon [Salmo salar), I shall adopt the sequence suggested by its natural habits, as likely to be that most easily followed and most conve- nient for reference. Thus, commencing with the ascent of the fish in the early spring and summer, its course will be briefly sketched from the tidal wave up the river to the spawning-ground, and back again to the sea, — the gradual changes ot colour and condition being at the same time pointed out. Returning then to the spawning- beds, where the ova should be approaching the 8 THE SPORTING FISH time of hatching, the growth and habits of the young fry will be traced from the ^g'g until their mieration to the salt water as Smolts ; their sub- sequent return as Grilse ; and, finally, as Salmon. Ascent of Salmon from the sea : fresh-run FISH. Sooner or later during the spring and summer months a proportion at least of the Salmon in the bays and estuaries of the coast make their way up the rivers for the purpose of spawning — their ge- neral colouring at this period being a brilliant sil- very white, merging into a bluish black with a few dark spots on the upper part of the body and head. When first ascending from the sea, Salmon are termed " fresh-run" fish, and are then in the most perfect condition both for the rod and the table. The marks by which a fresh-run fish may be known are, the bright, silvery hue of the belly and sides, and the comparatively loose adherence of the scales — this more ^particularly in the Grilse. In the case also of Salmon just fresh from the sea, a species of parasite, called " tide-lice," may be fre- quently found attached to the fish. These, how- ever, are killed by a few hours' contact with fresh water. In some rivers the ascent and spawning-time of the Salmon are much earlier than in others. This OF GREAT BRITAIN. 9 is often the case in rivers issuing from large lakes, in which the water has previously undergone a sort of filtering process and has become warmer, owing to the greater mass and higher temperature of its source ; whilst, on the other hand, streams which are liable to be swollen by the melting of snows, or cold rains, or which are otherwise bleak and ex- posed, are later in season, and yield their principal supply when the great lake rivers are beginning to fail. The order in which fish ascend rivers is — allow- ing for the variations already pointed out — gene- rally somewhat as follows : — First come the strong, early runners. These are succeeded by the Grilse, and by the small " Spring Salmon " which have probably never ascended at all as Grilse, but have remained in the sea since the Smolt state — a period of from eight to ten months, as noticed in " Proved Facts," No. 10. The scales of these spring Salmon are not easily rubbed off like those of the Grilse, and their tails are not so forked. A few of them generally appear with the early-running fish. As the season ad- vances, the larger fish and those heavy with spawn begin to work their way upwards from the mouths of the rivers and estuaries towards the hicrher reaches ; and such fish continue ascending from the sea until the close of the autumn, or, if the river be an early one, of the summer. Even as late as November and December, and lo THE SPORTING FISH the January and February following, a few fish continue to run which have been usually considered as " barren " — without capability of spawning ; but this notion was proved b)' the late Mr. Ffennell ^ to be erroneous, as upon dissection he found that the females had ova in them perfectly developed, although not larger than mustard-seed, whilst in the males a thin thread of milt was always dis- cernible. These fish, of which many ascend the Tay in November and the three following months, remain nearly a year in the fresh waters before spawning, and although their colours gradually be- come darker in consequence, they are to all intents and purposes "clean fish " — a term used to express the antithesis to " foul fish " — and are exceedingly good eatincr. The appearance of these so-called barren Sal- mon at a time when most fish are spawning, or are just recovering from the process, illustrates what I believe to be the most important fact connected with the history of the Salmon, and one which — until the publication of " Proved Facts," and my subsequent notes in the "Angler-Naturalist" in 1863 — does not appear to have been at all under- stood, although following as a natural corollary to the propositions of Mr. Ffennell, and hinted at by Mr. Brown in his account of the Stormontfield experiments', — viz., that the principle of a divided ' See Appendix to Report of Commission of House of Lords on the Salmon Fisheries. OF GREAT BRITAIN. ii misiration is not confined to the Parrs on cromcr to the sea, or to the Smolts on their return from it, some as Grilse and some as spring Salmon, but that it also extends to the old and adult fish after spawning — one portion of these latter coming back into the rivers during the following summer, and the rest not until the spring succeeding it ; in other words (and this is the gist of the whole), that at least a proportion of Salmon spaivn only on every alternate year. An analogous fact was observed by Dr. Davy with regard to the spawning of the common Trout. Dr. Davy was in the habit of opening the fish he caught, and by this means he discovered that, as the spawning-season approached, only about one half of the females had visible eggs, whilst in the other half there were no signs of the development of the ova. Charr, also, are frequently taken in Win- dermere in hicrh condition in October and Novem- ber, which is their regular spawning-season, — a fact which would seem to point to the possibility of the rule of alternate spawning-years holding good in the case of all fish here included under the genus Salmo. The design of this law or instinct — which, when once apprehended, will be found to explain many of the perplexities in the history of the Salmon — is intelligible enough, viz., to ensure as large a supply of clean fish throughout as large^ a portion of the year as possible, and to enable each river to 12 THE SPORTING FISH support the greatest stock, — a result which could only be obtained by such a provision as the above. It is also doubtless intended to ensure an equal dis- tribution of the fish throughout the whole length of the river. These Salmon, by ascending thus early, before their spawn is at all matured, are vigorous, and able to overcome the obstacles in their up- ward course to the extreme sources of the river — to which those fish which remain in the sea until heavy with spawn could never penetrate. But to return. During the early part of the season, the Salmon in the rivers, which do not at once ascend, remain in or near the mouths, most commonly advancing with the flood, and retiring with the ebb of the tide — unless captured by any of the contrivances hereafter mentioned ; but as the season pro- gresses they get gradually further into the fresh water beyond the influence of the sea, and at about this time will be found to be becominof full of roe, and more or less out of condition and unfit for food, according to their forward state as breeding-fish. In fact, the edible qualities of the Salmon when ascending rivers depend en- tirely upon the state of the development of the milt or roe. Even in the salt water this loss of condition follows upon the maturing of the spawn ; and when fish are precluded from entering rivers by want of a rise in the tide, or other cause, the development takes place as in the stream, and the OF GREAT BRITAIN. 13 Salmon assume the recldisli-coloured tints distinc- tive of spawning-fish. With the approach of the spawning-time, the anxiety of the Salmon to ascend increases. They shoot up rapids with the velocity of arrows, and make wonderful efforts to surmount cascades and other impediments, frequently clearing a height of two or three yards at a bound. It has been calcu- lated— though I do not vouch for the accuracy of the calculation — that, when swimming, or rather darting at full speed, the Salmon will glide through the water at the rate of about 1,500 feet per minute, or 2,160,000 feet (upwards of 400 miles) per day — a pace which, if it could be maintained, would speedily carry' the fish round the world. Marvellous stories are related of Salmon-jumps, some altogether incredible, others, to say the least of it, highly improbable. No doubt the depth of the water from which they take their spring materially influences its height ; but, as a general rule, the limit of the perpendicular leap certainly does not exceed 12 or 14 feet; or, if they rise higher than that, the effort is aimless, and they are dashed down again by the current before they have recovered their energy. Frequently they are killed by the exhaustive violence of their exertions, and some- times they alight upon the rocks and are captured. With the advance of the season the fish begin to gain the upper and shallower reaches, or spawning- grounds ; and at this time all the Salmon and 14 THE SPORTING FISH Trout species resident in fresh water, both migra- tory and non-migratory, acquire, in lieu of their brilliant spring tints, a dusky-yellowish exterior, accompanied by a considerable increase of mucus or slime, — the fins also becoming more muscular. As the important operation approaches, these colours undergo a still further deterioration, the general hue of the body in the males assuming a browner or more golden tinge, and the cheeks being marked with orange-coloured stripes ; the lower jaw elongates, and a gristly projection or horn turns upwards from the point, which is used by the Salmon as an organ of offence in its con- tests with other fish. In this state the males are called " Red fish," or are said to be " on the reds." The females are somewhat darker in colour, and are known by the name of " Black fish." The Spawning-beds. The usual time for spawning is from November until the latter end of January or the beginning of February ; but there are e.vceptional rivers both earlier and later, as already pointed out ; and it is probable that many of the so-called " barren fish," entering the fresh water in November and Decem- ber, spawn in the succeeding October. The process of spawning is as follows : — A pair of fish, male and female, select a gravelly shallow OF GREAT BRITAIN. 15 suitable for the purpose, which is generally occu- pied also by other spawners, both Salmon and Trout, as well as by a considerable number of male Parrs. These latter, as stated in " Proved Facts," No. 3, are perfectly qualified to con- tinue their species, and they perform a most important part in the reproductive process ; for the attentions of the male Salmon being con- stantly distracted by the necessity of protecting the spawning-bed from the intrusion of other fish, the ova of the' female are during these absences vivified by the milt of the Parrs.' The female deposits her eggs in shallow furrows in the gravel, to which they adhere by a thin coating of gluti- nous matter, the male at the same time shedding his milt over them. Whether these furrows are made conjointly by both spawners, or by the female fish only, and whether the snout or the tail is the organ used in the delving process, have been dis- puted points amongst naturalists. From the con- current testimony, however, of those who have had the best opportunities of observation, it now ap- pears certain that the trenches are made by the tail of the female fish only, and that the male takes ' According to the CNperiments of Mr. John Shaw (subse- quently confirmed by those at Stormontfield), male Parrs attain to the breeding-state in about eighteen months from the time of hatching. The females, it would appear, never become pro- lific whilst in the Parr state unless they are amongst the excep- tional fish, alluded to in " Proved Facts," No. 4, which remain over the third year in the rivers before migrating. 1 6 THE SrORTING FISH no share whatever in the more laborious portions of the parental duties. The only extra-matrimonial function that he performs consists in exerting an unwearied vigi- lance to protect his seraglio from the invasions of rival males, all of whom he assiduously endeavours to expel, — living, in fact, in a perpetual state of active hostilities. The female, regardless of the frequent absences of her lord during these contests, and probably satisfied with the presence of the male Parrs, proceeds with her operations by throw- ing herself, at intervals of a few minutes, upon her side, and whilst in that position, by a rapid action of the tail, she digs a receptacle for her eggs, a portion of which she on each occasion deposits, and, again turning on her side, covers it up by a renewed action of the tail ; thus alternately dig- ging, depositing, and covering ova until the whole are laid, — a process usually occupying a period esti- mated by different authorities at from three or four to twelve days. In the case of the male fish being captured or killed, the female retires to the nearest large pool in search of a fresh mate, with whom she returns and completes the process of depositing her eggs. This she will repeat several times if her partner be removed ; and it is mentioned as a fact by Mr. Young that nine male Salmon in succession have been thus killed from the side of a sinole female, who then brought back with her, as companion, a OF GREAT BRITAIN. 17 large yellow Trout I In consequence, we may sup- pose, of the arduous nature of his military duties and reddish colour at this period, the term " old soldier" is frequently used to designate the male Salmon after spawning. Return of Salmon to the sea as " spent fish " after spawning. After spawning, the adult fish are for some time in a very weak and exhausted state, and have not energy sufficient immediately to return to the sea. They usually drop down from the spawning- grounds, or " redds," to the first quiet deep, where they remain until their strength is somewhat re- cruited. They then continue falling back with the winter and spring floods, descending from pool to pool, and avoiding as much as possible weirs and rapid currents, until they reach the sea, where they quickly recover their condition, to ascend again (at least in many cases) in the autumn or succeed- ing spring for the same purpose as before, — always remaining, however, for a considerable period in the brackish water or tideway before making either decided change. After the conclusion of the spawning-process, Salmon are called " spent " or unclean fish, or " Kelts;" and at this time they are quite unfit for food — -indeed almost poisonous — and their capture is prohibited by law. Lately-spawned Kelts may 1 8 THE SPORTING FISH be recognized by their dark unhealthy colour, lanky, flaccid appearance, and by the enlargement of the vent. Their gills also are almost invariably found to be infested by a species of white worm (the Lcrncea Sabnonis of Linnaeus), which adheres firmly to the inside of the gill covers, and from which they are released by contact with the salt water, — a similar release from other parasites being obtained on passing to the fresh water from the sea. Within a period of five or six months after their return to the salt water, it has been proved that at least a proportion of Kelts find their way back to the upper reaches of the river as clean fish, having gained in weight during that time from 7 to 10 lbs. That spawned fish improve greatly in condition before they leave the fresh water there is no room to doubt, although they are never really fit for the table until their return from their sea-trip. A fish of this kind is known as a " well-mended Kelt," a term which is common amongst fishermen. Hatching of the Eggs and Growth of the YOUNG Fry. Leavine now the exhausted and more or less ill- conditioned Kelts to recruit themselves in their salt-water bath, we return to the spawning-bed OF GREAT BRITAIN. 19 where the eggs are approaching the time of hatchinof. Into this bed, during the preceding three months, a dozen females have each poured the germs of, say, from seventeen to twenty thousand Salmon, which, if they all arrived at maturity, would repre- sent in approximate figures some three million five hundred thousand pounds' weight of wholesome food, or a money- value of about _^ 160,000. Un- fortunately, however, the fry in fact added to the stock of the river are a mere fraction, and those that survive to return as Grilse a very trifling frac- tion of these numbers. The latter have been cal- culated by Mr. Brown at about one in every 1,000, and by Messrs. Ffennell and Ashworth at one in every six thousand, of the original deposit of ova. The causes of this destruction are numerous. From the first laying of the &gg until the plunge of the young Smolt into the tidal wave, and even after- wards in the broader waters of the estuary or open sea, a hundred wholesale depredators lie in wait for it. First there are the shoals of hungry fish of all kinds which prowl about the fords, pressing close behind the spawner, and ready to fight for the possession of her eggs almost before they are laid ; then come the voracious larvas of the May- fly and Stone-fly, and a host of kindred insects, which work their way in amongst the gravel and destroy, perhaps less ostentatiously, but not less certainly. io THE SPORTING FISH If the ^^^ escapes these perils, and having per- formed its protective mission releases its charge in due course, fresh dangers await the delicate and immature nursling. Again, the trout and the wild- duck, and even the parent salmon themselves, hunt it out in its sheltering creeks and crevices ; and hundreds of fry are daily sacrificed on a single spawning-bed by this means. Last of all comes man, who wantonly, either for amusement or for the sake of a single dish, fills his basket with pro- duce which, if allowed to pass to the sea, would have returned in a few weeks worth a pocketful of gold. With these various enemies besetting every period of their existence, it ceases to be a matter of surprise that the percentage of fry attaining the Grilse state should be as trifling as it is ; the only wonder is that it is not still smaller. Many causes of destruction, however, menace the ova of the Salmon besides those enumerated : a winter flood perhaps sweeps down the river, and buries a whole brood under a foot of sand-drift ; not only the duck and the grebe, but all sorts of water-fowl and amphibia perform their share of the work of depredation ; and though we know but little of the habits of the Smolts when once in the salt water, it may be conjectured that their adver- saries here are not less active and numerous than those of the river. To revert to the spawning-beds. In from 40 to 60 days after being first deposited OF GREAT BRITAIN. 21 in the spawning-bed, the ^^ A K u o I OF GREAT BRITAIN. 53 At Herdcott House, near Salisbury, there is preserved the skin of a Trout that turned the scale at 25 lbs., and was in length 4 feet 2\ inches; in girth it measured 2 feet i inch. Another fish is re- corded to have been taken in a small tributary of the Trent at Drayton Manor exceeding in weight 2 1 lbs. ; but there appears to be some doubt whether it was a bond fide Yellow Trout or a Salino trutta. The usual spawning-time of Trout is in the latter end of October or November, and thence up to the beginning of February (the operation, how- ever, in each particular fish continuing only about eight days) ; and at this period the under jaw in the old males exhibits in a modified degree the elongation and upward curving characteristic of the male Salmon at the same time. From the experiments of Dr. Davy, elsewhere commented upon, it appears probable that at least a proportion of Trout, like some Salmon, spawn only in alternate years. The situation chosen for, and the mode of conducting the spawning-pro- cess are also very similar to those noticed in the Salmon, — the eye, however, of the young fish be- coming visible in about three weeks, and the ^'g^ being usually hatched in from forty to fifty days. The number of eggs, in proportion to the weight of the fish, is about the same as in the case of the Salmon. The yolk-bag is absorbed in from three to five weeks ; and in six weeks or two months the young fry are about an inch long and able to shift 54 THE SPORTING FISH for themselves. From this time their growth is rapid or slow according to the nature and quality of their food and other local circumstances. In order to ascertain the relative nourishment of the different descriptions of food, some interesting experiments were made not many years ago. Trout were placed in three separate tanks, one of which was supplied daily with worms, another with live minnows, and the third with flies. The result was, that the fish fed with worms grew slowly and had a lean appearance, — those dieted on minnows became much larger, — whilst such as were fattened upon flies only, attained in a short space of time ex- traordinary dimensions, — w-eighing twice as much as both the others put together — the quantity of food eaten by them being actually less.' On another occasion Trout were kept for many years in a store stream, and tested with various kinds of diet, when it was ascertained that in some instances the increase in weight was as much as gibs, in four years (or from i to lo lbs.) Recent piscicultural experiments have demon- strated the great value of the Fresh-water Shrimp ('Gammari ') as an article offish diet, and for feeding young fry on, and I once had an exceptionally good opportunity of verifying the growth-rate of Trout when fed upon this insect. I quote from my volumes on Fishing in Messrs. Longman and Co.'s " Badminton Library of Sport :" — ' Stoddait, "Art of Ani^ding in Scotland." OF GREAT BRITAJN. 55 "At Encombe, in Dorsetshire, the seat of the Earl of Eklon, there is an artificial pond of two or three acres in extent facing the house. The pond is paved with marble at the bottom and sides, and is supplied with water from a small fountain fed from a spring in the neighbouring valley, carried by an artificial tunnel under some high hills. The pond is, for all practical purposes, stagnant ; the fountain's supply not being more than equivalent to the summer evaporation. From 1S62 to 1S64 this pond was drained off and left absolutely dry, in order to kill the weeds and clean the bottom. In 1864 the water was turned in again, and in August of that year a number of artificially reared Trout of the same season's hatching, about three- quarters of an inch long, were put into the pond. In August, 1866, the pond was again dried for cleansing purposes, when it was found that the Trout had grown in the space of two years to an amazing extent — four or five pounds being the smallest size, and a weight of six pounds ten ounces havinof been attained in several cases. "When visiting at Encombe in September of the following year, I examined the pond at Lord Eldon's request, with a view to ascertaining to what cause, in the absence of any artificial feeding, the extraordinary growth-rate was to be attributed. With the aid of a bucket and a rope, the explana- tion was not hard to tmd : the luholc pond zoas siviply swarDiing wiik ivalcr shrimps, and on ques- S6 THE SPORTING FISH tioning the keeper he assured me that when the water in the pond was let off there were Uterally cart-loads of these insects. My informant as to the facts and dates was Lord Eldon, who also exa- mined the keeper in my presence as to the circum- stances, and whose account was again confirmed in every respect by the corroborative testimony of Mr. Dickson, one of Lord Eldon's stewards, who was present and saw the fish weighed when caught. " The only outlet to the pond was a small drain at one end, up which nothing could practically pass, even if there were any Trout streams at hand with which it could be supposed to communicate. " It may be mentioned that the weight of the Trout at the end of the first year was from a quar- ter of a pound to half a pound." From this account and the experiments on the different modes of fattening Trout above referred to, it is evident that fish and grubs bear no com- parison with insect-food in point of nourishment, doubtless in consequence of the amount of phos- phate of lime contained in the latter. Of the insects specially contributing to the food of fish, probably the most nutritious of all are the May- flies, upon which, when arrived at maturity, the adult Trout wreak a signal vengeance for the de- struction effected by the larvae of the one amongst the eggs of the other. rrincipal Characteristics of t lie Coiinnon Trout (Salnio f;irio). (Taken from a llaiuiishire fish, 12 inches in length.) — Length OF GREAT BRITAIN. 57 of head compared to length of head and body, without tail-fin, as I to 4 ; depth of body rather greater than length of head. Profile of back and belly about equally convex. Head blunt ; lower jaw longest when the mouth is open, but shutting within the upper jaw on its being closed. Back-fin commencing be- tween point of nose and commencement of upper rays of tail- fin ; third ray of back-fin longest, and longer than base of fin. Small back-fin commencing halfway between origin of large back-fin and upper extremity of tail-fin ; pectoral fin two-thirds of length of head ; ventral fins under middle of first back-fin, and half-way between origin of pectoral fin and end of base of anal fin ; anal fin beginning halfway between origin of ventral fin and commencement of lower rays of tail-fin. Tail slightly forked, very gradually becoming square, or slightly convex in very old fish. Pyloric cajca ' seldom exceeding 46 in number. Teeth numerous, strong, and curving inwards, in six rows on upper surface of mouth and jaw, and four on lower. Teeth larger in males than in females. Number of scales above and below lateral line about 25. Vertebra: 56. The following were the full colours of a Dart- moor Trout taken by the author in May, 1885 : — The under line of belly (not seen when fish is looked at sideways), white ; sides, for one third of the way up, golden yellow ; two lines of red spots, one above and one below the lateral line : dark brown spots all over back, also above the lateral line, and a few below it just behind the gills; a few black or dark brown spots on the gill-cover {opcrailiiin). Head, gill-covers, and irides green bronzy-gold ; back and top of head, bronzy-green ; golden under the throat. Dorsal-fin green with black spots ; Pectoral- and ventral-fins orange ; anal-fin green-orange, edged with white ; adipose-fin having a bright margin of i)inky-red, the same colour as the red spots on the body ; tail-fin orange-green, top and bottom edges fringed with orangey-red. Fin-rays : D. 14 : P. 14 : V. 9 ; A. 11: C. 19. ' The pyloric csca are narrow pouches or ciik-de-sac, more or less numerous, attached to the pylorus or mouth of the intestines. THE GREAT LAKE TROUT.' |HE points in which this fish differs from the Common Trout, and also the dis- tinctions by whicli it maybe most easily recognized, have been already given in the chapter on Salmon. The Great Lake Trout, which is perhaps best known to anglers as the species for which Loch Awe has always been famous, is probably distri- buted throughout almost all the larger and deeper lochs of Scotland. It occurs to my knowledge in Lochs Ericht, Lochy, Garry, and Laggan, and it has also been recognized in Loch Shin, in Lochs Loyal and Assynt, and amongst some of the Orkney and Shetland Islands. In Ireland it ap- pears to be an inhabitant of all the best-known and most extensive lakes, having been found in Loughs Mask, Melvin, Erne, Corrib, and Neagh, where it is locally named Biiddagh, the younger ' Salnw fcrox. Salino, a Salmon or Trout ; fcrox, fierce, Lat. an y d H 1=3 o Hi W < Hi BRITISH SPORTING FISH. 59 and smaller-sized fish being termed DolacJunis. It is the Ullsivatcr Troiit and Grey Trout of the English Lake-districts, referred to by Dr. Hey- sham, and was at one time, it appears, erroneously considered to be identical with the Great Trout of the Lake of Geneva — a theory contradicted by Agassiz, who pronounced it to be distinct from any of the large Continental species. The specific name/crox has been given to this fish from its extraordinary fierceness and voracity, which are such that, having once seized a bait, it will allow itself to be dragged by its hold for forty or fifty yards, and when accidentally freed will im- mediately seize it again. The stomachs of the specimens that I have caught have been constantly found gorged with food. The only way of taking the larger fish is by spinning with a Loach, a small Parr, or other glittering bait trailed behind a boat, for which purpose very powerful tackle is required, as the fish is of immense strength, and its teeth are nearly as sharp as those of a Pike. The Great Lake Trout is almost wholly confined to lochs and deep extensive tracts of water, where it reigns in solitary grandeur, seldom venturing far up or down the streams, and never descending to the sea. It spawns in September. Principal Characteristics of the Great Lake Trout. — Length of head compared to total length of head, body, and tail-fin, about as i to 4^ ; depth of body less than length of head. Teeth large, strong, and numerous, arranged in six rows above 6o THE SPORTING FISH and four below — two of the upper rows extending along the vomer. Origin of back-fin halfway between point of nose and commencement of upper tail-fin rays ; third ray of back-fin longest, and equal to length of base of fin ; small back-fin half- way between last ray of large back-fin and end of tail-fin, and directly over origin of last ray of anal fin. Fins generally rather small and muscular. Colours when in season : upper parts and back-fin deep purplish brown, changing into reddish grey, and thence into fine orange on breast and belly ; whole body when fresh out of water as if glazed over with a tint of rich lake- colour. Gill-covers and back-fin marked with large dark spots, and whole body covered with markings of different sizes and varying in number in different individuals, being sometimes scattered and of large size, and at others thickly set and of small dimensions. Each spot surrounded by a paler ring occa- sionally of a reddish hue. Spots becoming more scattered below lateral line ; none on belly. Pectoral, ventral, and anal fins yellowish green, darker towards the e.xtremities. Tail of great breadth and power — broader than greatest depth of body, slightly forked when young, becoming square with age ; in very old fish slightly convex. Pyloric ca;ca from 34 to 49 in num- ber. Scales thin and flexible, different in form from those of the Trout, and more circular than those of any of the migra- tory species. Fin-rays in specimen \ from Loch Awe : ) D. 13: P; 14: V. 9: A. II : C. 19. In specimens from Lough Neagh, one ray less in D., P., and A. fins, and one ray more in C. fin. The only remaining British representatives of the genus Sabno^ — viz., the Charrs — being, hke the fcrox, exchisively inhabitants of Lakes, may appropriately receive in this connection a few words for the sake of completeness ; although, ' Or true Salmon and Trout species. OF GREAT BRITAIN. 6i unfortunately for the Angler, not coming strictly under the denomination of " Sporting Fish." In- deed, the species of this very beautiful group rarely take either bait or fly : so rarely that, amongst all the lakes, lochs, and loughs I have fished in all parts of the three kingdoms, I never actually caught an undeniable Charr but once, and that was in a little ghastly-looking, deep, black "tarn" on the top of a mountain in Inverness, during a snowstorm. At least five distinct species of Charrs, in- habiting different localities in the British Islands, have, however, been identified by Dr. Gunther, who has written a series of most interesting and beautifully illustrated papers on the subject. These species are — i. Sahno Cambncus, the Welsh Charr. 2. Sahno Wilhig/ibii, the Windermere Charr. 3. Saljuo Grayi, the Grey Charr, or Fresh-water Herring of Lough Melvin, Ireland. 4. Saliiio Alpinus, the Northern Charr, found in Lake Helier, Hoy, Orkneys, and probably in other Scotch waters ; and 5, Saltno Colli, found in Loughs Esk and Dan, Ireland. As a Division or Group the Charrs may gene- rally be distinguished from the two other sections of the same genus by the brilliant tints of the belly, which increase in vividness as the spawning- season approaches, and by the comparatively very minute size of their scales. THE GRAYLING.' ALTHOUGH a well-known and beauti- ful member of the Salmon family, the Grayling belongs to a different genus to the preceding groups, which all follow the true Salmon and Trout as their types. Its generic characteristics are : two back-fins ; the base of the first very long, with numerous rays ; the second small and adipose, without rays. Mouth small, with a squarish orifice ; teeth very small and conical ; air-bladder large ; body elongated. Gill- rays seven or eight. In the "Angler-Naturalist,"^ in my remarks on the Grayling, I observe that — " Whilst yielding to its sister species the Trout in the qualities of dash and obstinate courage, the Grayling is yet a sturdy and mettlesome fish — ' a foeman worthy of ' Thymalhis vulgaris. Thyiihil/its, "thymy" — ixom t/iyiutiiii, thyme, and rii/garis, common, Lat. - "A Popular History of British Fresh-water Fish," or the " Anglcr-Naturahst." London : Routledge and Sons. s-^ ^ I i w BRITISH SPORTING FISH. 63 our steel ;' and if the former is the liandsomer, the latter must, I think, be admitted to be the prettier species of the two. The Trout has, so to speak, a Herculean cast of beauty; the Grayling rather that of an Apollo — light, delicate, and gracefully symmetrical." Though abounding in some streams, the Gray- ling is a remarkably local and even comparatively rare fish, thriving best in rivers the bottoms of which are composed principally of sandy gravel or loam — a soil highly favourable to the production of the insect food on which it in a great measure subsists. Rocky or stony bottoms are very inimi- cal to its breeding; and this is probably the reason why, though flourishing in many Continental waters, so few exist in those of Ireland or Scot- land. Indeed, even in England, a dozen names or so include all our streams which have any right really to be considered as properly Grayling- waters ; and these, with hardly an exception, be- long to the southern and western portions of the island. The fact is no doubt accurately stated by Mr. Blaine when he says, " Grayling require other peculiarities of location besides those of tempera- ture, such as, for instance, the general character of the water they inhabit, and certain circumstances in the natureof its composition derived from its sources. It is probably owing to the abstraction of some of these requisites that the breeding of the fish in several rivers in which they have been attempted to 64 THE SPORTING FISH be naturalised has not been attended with much success. In some they soon disappeared ; in others they remained, but never thrived ; while in some waters, though they lived, and at first increased, yet they were afterwards observed to shift their quarters to different grounds, in most of which cases it proved, as in the Test of Hampshire, that they migrated from above downwards, probably in search of deeper and more tranquil waters : for the angler cannot fail to observe that Grayling do not, like Trout, affect very rapid shallows and the coldest torrents ; on the contrary, they seem to thrive best where milder currents alternate with deep and extensive pools." On these rapids, how- ever— or "stickles," as they are termed — small Grayling may frequently be found, but the large fish rarely, except in the spawning-season. The haunts of large Grayling are the deepish and slowly-running tails of streams or pools, a few yards before the formation of fresh shallows ; and here they will be found at all times, except when spawning. It has been asserted that the Grayling is not, like the Trout, indigenous to this country, but was introduced by the monks on account of its edible qualities — a supposition to which the peculiarity of the local distribution doubtless gave rise. The fol- lowing is a list of the best known Grayling waters, though I will not say that there may not be others with which I am unacquainted : — OF GREAT BR ITALY. 65 In Hampshire and Wiltshire the GrayUng" is found in the Test, the Itchen, and in both the Avons, &c. ; in Herefordshire, in the Teme, the Lug, the Wye, and the Arrow ; in Shropshire, in the Teme and Clun ; in Staffordshire, in the Hodder, the Trent, the Dove, the Blythe, and the Wye ; in Derbyshire, in the Dove ; in Merioneth- shire, in the Dee, between Curlen and Bala ; in Lancashire, in the Ribble ; in Yorkshire, in the Derwent, near Scarborough, in the Yore, the Wharfe, and in the Whiske, near Northallerton, in the Rye, Swale, Costa, and (Yorkshire) Dove, near Pickering ; in Berkshire, in the Kennet, at Hungerford ; and in Cumberland, according to Heysham (but this appears doubtful), in the Esk and the Eden. In Scotland the Grayling may now be considered to be established in the Clyde, and it has also been recently caught in the Annan, as recorded by a writer in the " Eishing Gazette." The Teme Grayling has the reputation of being the finest in England, and when in the height of condition — that is, in October or November — and just taken from the water, is certainly one of the most beautiful fish that can be imagined. At this time the back is of a deep purple colour, with small dark irregular spots on the sides ; the sto- mach is brilliantly white, with a fringe or lacing of gold ; the tail-, pectoral, and ventral fins are of a rich purplish tint. The dorsal fin is very large — almost disproportionately so — and is covered with F 66 THE SPORTING FISH scarlet spots and wavy lines upon a dark ground of reddish-brown. The little velvet back-fin near the tail is also dark brown or purple, and the whole body is shot with violet, copper, and blue re- flexions when seen in different lights. Properly to appreciate this colouring, the fish should be laid horizontally upon the hand to be looked at, in which position its varied tinting is seen to the ofreatest advantage. In addition to its delicate colouring, the Grayling has been supposed to possess a peculiar thymy smell, from which it takes its specific designation. The English name, " Grayling," is probably a modifica- tion of" Grey-lines," in reference to the longitudinal dusky-blue bars with which its body is marked. In size Grayling vary according to locality; but they rarely exceed 3 lbs. in weight, and by far the greater number of those taken are under i lb. Oc- . casionally, however, they are met with of even a larger size than that above named : Mr. T. Lister Parker took three fish in the Avon, near Ring- wood, which together weighed 12 lbs. ; a Grayling of 4nbs. weight was killed in the Test, and one of 5 lbs. is recorded to have been taken in the neigh- bourhood of Shrewsbury. No doubt the possible ultimate growth of the Grayling depends upon the amount and quality of its food, which, besides flies, worms, caterpillars, etc., consists of the larvae of dragon-flies, May-flies, and other ephemera. OF GREAT BRITAIN. 67 Unlike the rest of its congeners of the Salmon family generally, the Grayling never jumps out of water, and is apparently unable to surmount either natural or artificial obstructions or to stem very rapid torrents, being, in fact, much more prone to going down than up the stream. It has the power, however, of raising itself rapidly to the surface, and of descending again with stone-like velocity — a faculty which has been ascribed to the action of the large dorsal fin striking either up- wards or downwards against the current, but which is more probably attributable to the unusual size of the swimming- or air-bladder. The eggs are numerous but considerably smaller than those- of the Trout, being about the size of partridge-shot, and when viewed in the rays of the sun having very much the colour of the opal. The body of the embryo fish becomes distinctly visible in about nine days, and the egg itself hatches in fourteen or fifteen days from the date of deposit, — results obtained in the case of the eggs of the Trout in about thirty-five and fifty days respec- tively. The spawning-time is in April or the beginning of May, the fish getting into condition in July, and reaching its prime in October and November, when most of the other Salmonidce are going off. The Grayling has, moreover, according to Sir Humphry Davy, the advantage of rarely being so much out of season as to be unfit for food or un- 68 BRITISH SPORTING FISH. willing to take a bait if judiciously offered. This author, who has supplied a very fair history of the fish, considered that it might be taken at all times of the year, and that when there were flies in the water it would generally rise to them. '' The specimen from which the illustration is taken, was a beautiful Teme fish about a pound and a half in weight. Its proportions were : — The total length of the head, body, and tail-fin, being considered as 5^, the length of the head alone as one. Depth of body for about half its length slightly greater than the length of head ; tail forked. Number of scales in lateral line, 27. Further Cliaractcristics cf flic Grayling (as found in a speci- men 10 inches long). — Head small and pK)inted, flattened at the top. Between point of nose and commencement of first back-fin equal to one-third entire length, tail-fin rays excluded. Back-fin very long at the base, nearly equal to twice the length of its longest ray ; pectoral fins long, narrow, and pointed ; ventral fins commencing directly under centre of back-fin ; final fin commencing halfway between origin of ventral fins and fleshy part of tail. Opening of mouth when viewed in front squarish ; teeth small, numerous, and curved ; none on tongue, and only a few on end of vomer, or central bone in roof of mouth, and on the adjoining ends of the bones of the palate, situated in single rows on the jaws. Profile of back slightly convex, that of belly nearly straight. Scales large, seven in an oblique row above the lateral line. Vertebrae 58. Colour : sides marked with about 15 dusky longitudinal bands or bars; the general hue becomes darker with age, and about the spawning- season the pectoral fins are reddish with small black spots : for further remarks on colouring see p. 65. w THE PIKE.' |HE Pike, of which we have only one ^^ recognized species in this country and ^^ ^y on the Continent, is common to most """^ of the rivers and lakes of Europe and North America, and the INIascalonge {Esox estor) and the Northern Pickerel {Esox lucioidcs) — both inhabitants of the great lakes; the common Pickerel {Esox reticulatus) — indigenous to all the ponds and streams of the Northern and Midland States ; the Pickerel of Long- Island {Esox fasciatits) — probably confined to that locality ; the White Pickerel {Esox vittatus), the Black Pickerel {Esox niger), and Esox phaleratus — all inhabitants of the Pennsylvanian and Western waters, have been generally recognized as distinct species. Of these the first two are the types, the others following, more or less closely, the same formation as to comparative length of snout, formation of the lower jaw, dental system, gill covers, &c. As regards the European Pike, it seems pro- bable that there may be varieties yet to be dis- ' Esox Indus — Esox, a Pike ; luciiis, the Pike, Latin names. 70 THE SPORTING FISH covered, as Dr. Genzik, the well-known naturalist of Lintz, informed me that he had found some specimens which had teeth like the fangs of a viper — capable of being erected or depressed at pleasure, — a circumstance the more remarkable as the jaws also of the Pike are furnished with extra bones to increase the size of its gape, very similar to the corresponding bones in the Viper-con- formation. Indigenous in all climates which are not tropical, a chilly or even frigid latitude is apparently essen- tial to the well-being of the Pike. Thus in Nor- way and Sweden, Siberia, and the lakes of Canada and Lapland it reaches its full development, breed- ing in vast numbers, and commonly attaining the length of 4 or 5 feet, whilst it rapidly degenerates on approaching warmer latitudes — diminishing in geographical distribution with the spruce fir, and ceasing entirely in the neighbourhood of the Equator. Of the numerous names by which the Pike is known the common term " Pike," or " Pickerel," is probably the only one derived from our own language ; and this would appear to have origi- nated in the Sa.xon word ////', signifying " sharp- pointed," in reference doubtless to the peculiar form of the Pike's head — or possibly teeth } In Sweden the fish is named Giidda, and in Denmark (amongst other variations) Gcdde, appa- rently identical with the Lowland Scotch, Gcdd, OF GREAT BRITAIN. 71 though spelt somewhat differently. M. Valen- ciennes has published a long list of the names which the fish bears amongst the Sclavonic and Tartar races, but none of them seem to have any relation to those by which it is known on the western coasts of Europe. The Pike is the Brocket of the French, by whom it is also called JSrocketoii, Lance or Lanceron, and Becqiwl — the last having reference doubtless to the flattened or duck-bill-like form of the muzzle. The ancient classical name of the Pike was Ltuiiis^ under which it is mentioned by several old writers ; and from this root have doubtless sprung the terms Lnce or Lucie (the "White Lucie" of Shakespeare and of heraldry), as well as the Luccio or Luzzo of the Italians. To the ancient Greeks, so far as we are aware, the Pike was a stranger; or if known, has escaped notice in the writings of Aristotle. In the works of several Latin authors it is mentioned, and is stated to have been taken of very great size in the Tiber ; but it has been doubted by naturalists whether this fish — the Esox of Pliny— is synony- mous with the Esox, or Pike, of modern ichthyology. ' Nobbes suggests that the name Lucius is derived "either a luceiido, from ' shining in the waters,' or else (which is more probable) from Lukos, the Greek word for lupus : for as," says he, " the wolf is the most ravenous and cruel amongst beasts, so the Pike is the most greedy and devouring among fishes. So that Lupus Fiscis, though it be proper for the Sea-wolf, yet it is often used for the Pike itself, the Fresh-water Wolf." 72 THE SPORTING FISH Ausonius, however, living about the middle of the fourth century, clearly alludes to the Pike in his well-known lines : — " Lucius obscurus ulva lacunas Obsidet. Hie, nullos meusarum lectus ad usus, Fumat fumosis olido nidore popinis." The age to which the Pike will attain has been always a debated point amongst naturalists. Pen- nant mentions one ninety years old. Pliny con- sidered it as the longest-lived, and likely to reach the greatest age, of any fresh-water fish ; while Sir Francis Bacon, agreeing in this view, yet limited its probable maximum to forty years. In natural connexion with this part of the sub- ject— the limit of age in the Pike — occurs that of its probable growth and size when suffered to attain to full development. It has been the custom amongst modern writers to aftect a civil disbelief in the accounts of very large Pike handed down to us by numerous credible witnesses ; and the prevailing impression appears to be that a weight of 30 or 40 lbs. is about the real maximum attained. I could easily refer, however, to many attested examples of Pike having been taken in the British Islands up to the weight of 70, 80, and even 90 lbs. ; but a single instance, too well au- thenticated to admit of doubt, will suffice. I refer to the case of the Kenmure Pike — mentioned also by Daniel in his " Rural Sports," and by Dr. Grierson and other authors — the weight of which OF GREAT BRITAIN. 73 was 72 lbs. It was taken in Loch Ken, Galloway, a sheet of water belonging to the Castle of Ken- mure, where the head of the fish is still preserved, and may be seen by any one sufficiently curious or sceptical to desire ocular demonstration. To give a general idea of the size of the Pike, I may quote one measurement — that across the back of the head, — the width of which was nine inches. Dr. Genzik, who gave me some valuable in- formation concerning the Continental Pike, states that, in the fish-markets of Vienna, Lintz, and Munich, Pike are not unfrequently exposed for sale of 80 and 90 lbs. weight and upwards, — that the fishermen on the Danube, near Strudel and Wirbel, have legends of specimens 15 and 20 feet long, which break through all their nets, — and that at Traunkirchen, on the Gmiinden Water, there are still living some fishermen who declare that on one occasion when dragging the lake they enclosed a Pike longer than either of their boats, and that they began, as they expressed it, " to say their prayers, thinking the enemy was on their nets ; the Pike, however, with one spring, jumped over the nearest boat and escaped " ! At Oberneukirchen Dr. Genzik himself saw a Pike taken out of a large tank or preserve, which after being cleaned, weighed 97 lbs. and some ounces ; and an officer of Tyrolese Rifles informed him that whilst at Bregentz during the past au- 74 THE SPORTING FISH tumn (1862), he was present when a Pike was caught weighing upwards of 145 lbs. After this we may be indined to give more credence to Bloch's statement that he once exa- mined a portion of the skeleton of a specimen which measured eight feet. The rate of growth of the Pike has been by different authors variously estimated at from i to 5 lbs. a year; but these estimates do not, generally, appear to be based on any very reliable grounds, still less upon actual experiment, and therefore go but a little way towards advancing our knowledge of the subject. My own e.xperience leads me to believe that the growth-rate is susceptible of very great variation, depending upon the nature of the water and the amount of food supplied to the fish, but that in the open waters— at least in Eng- land— it seldom averages more than i lb. a year during the first two years, and from i-^- to 2 lbs. a year afterwards, decreasing again after eight or nine years to about the original ratio. This ave- rage cannot, of course, be taken as any index of what may be done by keeping Pike in rich pre- serves or fattening them in stews, as the capacity of the Pike for food is well nigh inexhaustible, and is in analogy with its digestion the marvellous rapidity of which has been aptly described as re- sembling the action of fire. Of one point, however, I have fully convinced myself, viz. that during the first year the maximum OF GREAT BRITAIN. 75 growth, in open waters, does not much exceed half a pound. The grounds of this conclusion are briefly as follow : — Pike spawn in March or April ; in June, when Pike-fishing commences, I have not unfrequently taken, and seen taken, with the net, small Jack of about an ounce, or a little more, in weight ; in September, again, I have constantly taken them, with a minnow, of ihrce or four ounces ; and in January and February speci- mens of from yft'^ to seven otinccs ; whilst I have never within my memory caught the smaller-sized fish at the later periods, or vice versa, — thus point- ing clearly to the inference that at these seasons there were young Jack of those respective sizes, and none others — in other words, that the different sizes represented the different stages of growth. These I believe to be the fish of about \ lb. of the following season. The Pike is a true cosmopolitan in his feeding. Fish, flesh, and fowl are alike acceptable to him. Animal, mineral, and vegetable — his rapacity de- vours them all. Lord Walsingham writes to me that he once caught a pike with a hare's foot, and on another occasion with a dead rat ! Nothing, in short, that he can by any means get into his stomach comes amiss to him ; and imperial man himself has on more than one occasion narrowly escaped being laid under contribution to his larder. His own species enjoy no immunity from this uni- versal rapacity; on the contrary, it has been asserted 76 THE SPORTING FISH that more young Jack are destroyed by their own parents than by any or perhaps all other enemies put together. A remarkable instance of this characteristic propensity was communicated to me by Mr. L — of Chippenham, Wiltshire, to whom the incident occurred. This gentleman had set a trimmer in the River Avon over night, and on proceeding the next morning to take it up he found a heavy Pike apparently fast upon his hooks. In order to extract these he was obliged to open the fish, and in doing so perceived another Pike of considerable size inside the first, from the mouth of which the line proceeded. This fish it was also found neces- sary to open ; when, extraordinary to state, a third Pike of about \ lb. weight and already partly digested was discovered in the stomach of the second ! Of the indiscriminating character of the Pike's appetite a more apt illustration could not perhaps be given than the following, kindly communicated to me by Lord Walsingham, who is himself an en- thusiastic angler and a close observer of nature : — " A newly cut ash-pole, with the bark peeled off for a few inches at the small end, was lying across the punt in which I was fishing, the pole was some- what bent, and owing to the wind and my move- ments in the boat, the end frequently dipped in the water. A Pike dashed at it and seized it, leaving the marks of his teeth distinctly visible in the wood." OF GREAT BRITAIN. ^^ Another amusing instance was related to me by an eye-witness : — Upon a piece of water belonging to Wandle House, Wandsworth, some toy vessels were being sailed, at the stern of one of which was attached a small boat fancifully decorated with green and gilding. As the little craft swept briskly across the pool, with her boat in tow, a Pike suddenly darted from the water and seized the boat in his jaws, well nigh capsizing the whole flotilla in his efforts to drag his capture to the bottom. To this task, however, his strength was apparently unequal, and a fresh breeze springing up, the submerged nautilus reappeared on the surface and continued her voyage, but had hardly got fairly under way when the Pike again dashed forward to the attack, seizing her as before, and repeating the performance vmtil the boat grounded on the opposite bank. It has been mentioned that in rare instances and when under the influence of either extreme anger or hunger. Pike have been very clearly con- victed of attempted manslaughter. Such being the case, it is hardly necessary to say that it is by no means uncommon for animals, often of large size, to be similarly attacked, and, in the case of the smaller species, devoured, by this fish. Ac- counts are on record of otters, dogs, mules, oxen, and even horses being assaulted. Poultry are constantly destroyed by the Pike, — " the dwellers in the ' Eely place,'" as Hood punningly says, 78 THE SPORTING FISH " having come to Picc-a-dilly." I have seen a Pike take a pigeon that I had myself shot from a pontoon at the Welsh Harp— the circumstance indeed was witnessed by a number of people. Lord Walsingham writes to me : — " Partridge shootintr near a large piece of water I wounded, late in the evening, a bird, which flew in the direction of the water ; the next day I was Pike fishing, and about fifty yards from the land, just opposite where the wounded bird must have reached the water, I caught a Pike of about 8 lbs. with a partridge in his stomach— undoubtedly my bird. I discovered it by seeing feathers sticking out of his throat while extracting the hooks." Sometimes the heads of swans diving for food encounter instead the ever- open jaws of this fish, and both are killed ; whilst among the frogs he is the very " King Stork" of the fable. He will even seize that most unsavoury of all morsels, the toad, although in this case the inherent nauseousness of the animal saves it from being actually swallowed, — its skin, like that of the lizard, containing a white highly acid secretion which is exuded from small glands dis- persed over the body. Pike will attack both the land- and water-rat ; occasionally pouching them, but more frequently treating them as in the case of the toad, — a fact confirmed by Captain Williamson, who adds : " But whether owing to the resistance that animal (the rat) makes, which I have witnessed to be very OF GREAT BRITAIN. 79 fierce — and that under water too — or whether owing to the hair or scent displeasing them, I know not ; but they do not appear to be very partial to the quadruped." Sometimes the Pike lies in ambush, protruding only its eyes and grim muzzle through the weeds, when the movement of seizing a victim is little beyond a quick turn of the body and an opening and shutting of the jaws ; but generally he takes his prey with a rush and a flash, emerging so sud- denly and with such startling energy, that I have, in more than one instance, known a Troller lite- rally drop his rod from the effects of sheer terror. Woe to the fish or flesh that gets once fairly enclosed between the jaws of a Pike ! From that more than Regulus-like incarceration it rarely effects its escape. Like the Remora, which will allow itself to be cut to pieces rather than relinquish its hold, the Pike exhibits an extraordinary tenacity of purpose and reluctance to quit its grasp of a prey once seized. A Pike will even constantly make a considerable fight, and actually allow itself to be dragged many yards by the obstinacy of its hold, without ever having been pricked by a hook — shaking the bait out of its mouth perhaps when almost in the net. I have witnessed the same thing to occur with a cork ball tied to a string and drawn across a stock-pond. This mixture of determination and ferocity 8o THE SPORTING FISH makes the Pike the undisputed master of our fresh waters, akhough it has been asserted that, from its superior momentum, a Trout or Sahnon of equal weight would have the advantage in a pitched battle. I doubt it much. What chance these fish have against the Pike is shown by the effect of introducing the latter into trouting and salmon waters, where the new-comer speedily dispossesses the rightful tenants. Witness, for instance, the ravages committed in the Canterbury River, in the Wandle, in the Colne near Draycot and Cow- ley, in the Teviot, and in Lochs Katrine, Lomond, Awe, and Jurit in Scotland ; and the same thing is known to have taken place in many of the best Irish waters, where the Pike is still continuing to spread and multiply, displacing by degrees the Trout and other indigenous races. Indeed, how Pike spread is a problem which it has perplexed naturalists to explain. A stream, or pond, or loch, reserved perhaps for centuries to the docile phlegmatic Carp, or " star-stoled Trout," suddenly begins to show symptoms of a falling off; the next year matters are worse ; the water is dragged, and the first fish to come up in the net is probably a Pike. How the Pike came there, or who put it there, remains unexplained ; but the cause of the depletion of the water is no longer a mystery. Some authors have accounted for these singular immigrations by supposing that the Pike, like the Eel, actually travels overland in wet OF GREAT BRITAIN. 8i weather from one pond to another ; and several curious circumstances, which have come to my knowledge, would appear to lend some colour to the supposition. In order to test the migratory theory, Mr. Newenham, an English resident at Antwerp, had two small ponds excavated near together, and stocked one with Pike, and the other with small Roach, Dace, &c. At the end of the second day he caused both ponds to be emptied, when it was found that several of the Pike from pond No. i had made their way by some means into pond No. 2, and had destroyed a great part of the fry. A singular fact, pointing indirectly to the same conclusion, once came under my own observation. A pool five or six yards square, for the reception of small fish, had been constructed close to a stew- pond containing Pike ; the work had been finished in the afternoon, and the pond left to fill. On visiting it the next morning, I was surprised to find it already occupied by a Jack of about \ lb. weight, which had contrived thus early to take possession. The sudden appearance of Pike at certain times is not less remarkable than their unaccountable disappearance at others. There is no doubt that in seasons of great heat or unusual drought, when ponds or reservoirs have become rapidly dried up, the Pike that were in them have vanished in a very extraordinary manner, and that upon the return of the water they have been immediately G 82 THE SPORTING FISH found in apparently undiminished numbers — a phenomenon, however, by no means confined to Esocidcr, as the same thing has been observed with regard to Carp and Tench ; and Eels it is well known frequently make comparatively long jour- neys " overland." Although from its vigorous and unsparing de- structiveness the Pike has many detractors and few apologists, it must not be supjDosed that it is alto- gether without any of the softer instincts. On the contrary, it has been known to exhibit under par- ticular circumstances a very decided amount of friendship, and even affection, especially in the conjugal form. The " one virtue," however, to which, amongst a thousand crimes, the name of the Pike has been specially linked is gratitude : it has been asserted that he never attacks his physician, the Tench. To this subject I have referred in the notice of the latter fish. The haunts of Pike vary considerably at diffe- rent times of the year, and also vary with the nature of particular waters ; but it usually prefers a still, unfrequented spot, plentifully supplied with weeds and flags, selecting if possible a gravelly or sandy bottom. The neighbourhoods of reeds, docks, bulrushes, and the broad-leaved water-lily are its favourite resorts ; and of these, a flooring of lilies, with from four to six feet of quiet current over it, and a wall of reeds at the side, springing from the bottom, is the best — OF GREAT BRITAIN. 83 " A league of grass washed by a slow broad stream Tliat, stirr'd with languid pulses of the oar, Waves all its lazy lilies and creeps on. . . ." Indeed, it may be said that the reed and the hly are to the Pike what the hollybush is to the wood- cock. In lochs and meres it commonly frequents the most shoal and weedy parts, small inlets and little bays, or the mouths of streams where minnows or other fry congregate ; and in rivers, back-waters and dam-heads, eddies between two streams, or in fact any water that is weedy, of moderate depth, and not too much acted upon by the current. As a general rule, Pike will be found during the summer in or close upon the streams, and in winter, after the first heavy flood, in the large eddies and deeps. At the latter season the fish feed best about mid- day, with a breeze and a warm sun ; and in the summer months at morninor and evenin"- with a cloudy sky and plenty of wind. A hot, sultry day is always inimical to success in Pike-fishing ; as also a muddy or flooded state of the water : a full water, however, if not discoloured, is very favour- able. When in high season, the general colour of the Pike is green spotted with bright yellow, whilst the gills are of a vivid red ; when out of season the green changes to a greyer tint, and the yellow spots become pale. It spawns about March or April, according to the climate, forwardness of the spring, and other local circumstances, — the young 84 THE SPORTING FISH females of three or four years old taking the lead and the dowagers following. For this purpose they quit the open waters in pairs, and retire into the fens, ditches, or shallows, where they deposit their spawn amongst the leaves of aquatic plants ; and during this period the male may often be ob- served following the female about from place to place, and attending upon her with much apparent solicitude. As many as 80,000 eggs have been counted in one fish. When the spawning-process is complete the fish return again into the rivers, and are then for some weeks in a state of partial stupefaction, and unfit for food. In streams they begin to be in condition again about June, and are in their best season in November; but in still waters the recuperative process is much slower. Characteristics of tlic Pilie Family. — Head depressed, large, oblong, blunt. Body elongated, rounded on the back ; sides compressed, covered with scales. Dorsal fin placed very far back, over anal fin. Jaws, palatine bones, and vomer furnished with teeth of various sizes. Principal Characteristics of the common Pike. — Body elon- gated, nearly uniform in depth from head to commencement of back-fin, then becoming narrower ; body covered with small scales ; lateral line indistinct. Length of head compared to total length of head, body, and tail, as i to 4. Back- and anal fins placed very far back, nearly opposite each other. From point of nose to origin of pectoral-fin, thence to origin of ventral fin, and thence to commencement of anal fin are three nearly equal dis- tances. Pectoral and ventral fins small ; rays of anal fin elongated. Tail somewhat forked. Shape of head long, flattened, and wide ; gape extensive. Lower jaw longest, with numerous small teeth round the front. The sides with five or six very large and sharp OF GREAT BRITAIN. 85 teeth on each side. Upper jaw somewhat duck-billed. Teeth on vomer small ; on the palatine bones larger and longer, parti- cularly on the inner edges : none on superior maxillary bones. Head covered with mucous orifices placed in pairs. Cheeks and upper parts of gill-covers covered with scales. Colour of head and upper part of back dusky olive-brown, growing lighter and mottled with green and yellow on sides, passing into silvery white on belly ; pectoral and ventral fins pale brown ; back-, anal-, and tail-fins darker brown, mottled with white, yellow, and dark green. THE PERCH/ ^UVIER has taken the Perch as the type of his leading genus. In other words, it is the first Species, of the first Genus, of the first family, of the first Order, of the first Series of fish in his systematic classification. In the present essay, however, I have found it on the whole more convenient not to adopt the strictly ichthyological sequence in re- ferring to the different species of sporting fish, and the Perch here takes his place not according to family ' descent,' but as naturally following after the Pike, with which in actual sport he is so constantly associated. The Perch — the common Perch of our ponds and rivers — is very generally distributed over almost the whole of Europe and Great Britain, and is one of the most generally recognized inhabitants of English waters from the Tweed to the Land's End. In Wales it is a somewhat local fish, and confined principally to stagnant waters; in Ireland more widely diffused, though still in distribution ^ rercafluviatilis. Perca, the Perch ; fliiviatilis, of the river, Lat. ^ W X BRITISH SPORTING FISH. 87 rather unequal ; in Scotland very general south of the Frith of Forth and comparatively rare to the north of it, ceasing entirely amongst the innumer- able waters of Sutherland and Ross, or, where observed, owing its introduction to very recent times. Of the British Perch, so far as my experience extends, the Thames produces the best in the matter of quality ; Windermere, and Slapton Ley, in Devonshire, the greatest show as to quantity ; and the Kennet from Hungerford to Reading, or perhaps the Avon below Sommerley, the finest specimens for general size and weight. The num- bers of Perch in Windermere and Slapton Ley are almost incredible : but their size is commonly in- significant. The Perch of the Thames, which is also notice- able for its fine colouring, probably owes its supe- rior gastronomic attractions to the great purity of the stream above locks as well as to the wide range in the choice of food, spawning-ground, &c., which it affords ; but in whatever waters the fish is bred it is seldom other than palatable as well as wholesome, and it is on this account a frequent item in the invalid's dietary. " More wholesome than a Pearch of Rhine," says a German proverb mentioned by Izaak Walton, who also quotes a learned authority to the effect that it possesses a small stone in the head thought to be very " medi- cinable." 88 THE SPORTING FISH The Perch selected for iUustration was a Kennet fish ot 3 lbs. weight — one of the most perfectly shaped specimens that I ever met with. The Perch lives long out of water (resembling in that respect Carp and Tench), and if carried with care and occasionally moistened will, in cold weather, exist for several hours in this condition. Yarrell says that Perch are constantly exhibited in the markets of Catholic countries, — where doubt- less they are a popular article of " fast ;" — and that on these occasions, when not sold, they are taken back to the [:)onds from which they came to be re- marketed at another opportunity. The size to which the Perch will attain is some- what doubtful. The largest specimen that I ever remember to have met with was preserved in a small inn in Perthshire. It was very imperfectly stuffed, and had no doubt shrunk considerably in the operation ; but the label stated that it weighed 7 lbs. when caught, and from its appearance it cer- tainly could not have weighed less than five or six. Perch of 2 and 3 lbs. are by no means uncommon, and specimens of even 4 lbs. are probably less rare than may be supposed. Such fish have been taken in the Pen Ponds, Richmond Park ; and Pennant records the capture of one in the Serpentine which weighed 8 lbs. Donovan speaks of a Perch of 5 lbs., caught in Bala Lake. One of 6 lbs. was taken by Mr. Hunt, of Brades, Staffordshire, from the Birmingham Canal ; and two fish of 8 lbs. each OF GREAT BRITAIN. 89 are stated to have been caught, the one in the Wikshire Avon, and the other in Dagenham Reach, Essex. One is recorded by Izaak Wakon as having been taken by a friend, which measured two feet in length ; and in the " Sure Angler's Guide," the author says that he saw the figure of a Perch drawn in pencil on the door of a house near Oxford, which was twenty-nine inches long, and he was informed that it was the outline of a living fish. It is not probable, however, that either of these sizes represents the maximum attainable by the Perch under favourable circumstances, even in British waters ; and it probably reaches a much ereater weight in Scandinavia and other northern countries. The Danube breeds enormous Perch ; and in the Church of Lulea, Lapland, the head of one is preserved which is nearly a foot long — • Solving the entire length of the fish at somewhere about 3-^ feet. After the Pike, the Perch is the most absolutely fearless of our sporting fish, often following a bait up to the very surface of the water or side of the boat, when with jaws open and spines erect he presents a really splendid appearance. He will often remain thus for several instants bristling like a porcupine, as if threatening reprisals on those who have baulked him of his prey — retiring at length in sullen wrath with his face still turned towards his assailants. 90 THE SPORTING FISH How far the spines of the Perch protect him from the assauks of other fish — especially of the Pike — is not quite certain. In many fishings both in England and Scotland small Perch are consi- dered the favourite baits for the Pike, which does not seem to be at all deterred by their spiky ap- pearance. Moreover, as the Pike always swallows his prey head-foremost, whilst the spines of the Perch are capable only of projecting backwards — shutting down like the props of an umbrella upon pressure from in front — it would appear that they ought rather to assist than impede the operation of swallowing. The favourite food of the Perch consists of min- nows, of which he can devour an extraordinary number ; but failing these, any small fry, worms (especially brandlings), insects, frogs, caterpillars, and grubs of all sorts are good baits. The extent to which he will gorge himself with minnows may be judged by the fact — within the knowledge of most anglers — that when he has filled his stomach so that he can positively swallow no more, he will still endeavour to bite and, if pos- sible, masticate others ; and it is by no means un- common under these circumstances to hook and capture a Perch with the tails of the minnows which he has already partially swallowed, and been unable to pouch, protruding from his gullet ! From this it may be inferred that fish, like many other animals — including man himself, in the first degree OF GREAT BRITAIN. 91 — find a gratification in the exercise of their pre- datory instincts even when the appetite of hunger has been fully satiated. Although having, it will thus be seen, the bump of destructiveness very liberally developed, Perch are by no means difiicult to tame : Mr. Jesse succeeded, after a few days only, in inducing them to feed from his hand. Bloch mentions having watched them depositing their ova in a vessel kept in his own room ; and I was informed by Mr. Bartlett, the eminent and indefatigable Superin- tendent of the Zoological Gardens in Regent's Park, that the Perch in the aquarium there in- crease their numbers by an annual shoal of young fry which are hatched and reared under the eyes of hundreds of visitors. A comical anecdote turning upon this faculty of ready acclimatization is related by the author of " Fishes and Fishing." A country gentleman, in order to add to the usual attractions of a Christmas visit, offered a London friend of piscatorial ten- dencies the temptation of a "day's fishing in his private water." The sportsman arrived, with a good assortment of rods, lines, &c., and was all- impatient to be at his work. After lunch his host in- troduced the visitor to " his water" — which proved to be an ornamental basin, in width about equal to the lencrth of one of the rods the angler had brought with him. The chagrin and disappoint- ment of the latter may be imagined ; but upon the 92 THE SPORTING FISH assurance that there really were fish in the pond, he put his tackle together and adjusted a bait. It had hardly touched the water before he hooked and landed a fine Perch ; another and another fol- lowed, and by the time his friend came to summon him to dinner he had thirty-five fish in his basket. "Well," said his host, " I am glad you have had such sport ; I caused three dozen to be put in the day before yesterday." " Indeed ?" was the reply, — "then I will go back and catch the thirty-sixth after dinner." Though attaining their greatest perfection in clear, sharp streams. Perch thrive well on clayey or sandy bottoms, preferring generally a water of a moderate depth and current, and frequenting holes, mill-dams, hollow banks, and, in summer, the under-currents of weirs. In winter their fa- vourite haunts are back-waters and eddies ; and at this time of the year they feed best about the middle of the day, more particularly if bright and warm. They usually swim in large shoals. Their spawning season is at the end of April or the be- ginning of May ; and so prolific are they, that a specimen weighing half a pound has been found to contain 280,000 ova. The eggs are deposited in strings which hang about the weeds and rocks, and when seen through a bright sunlight present a beautiful appearance almost resembling festoons of pearls. Of the Perch family only two fresh-water species OF GREAT BRITAIN. 93 are known to exist in this country, the common Perch, and the Ruffe or Pope ; the latter of Httle interest either for sport or for the table. The characteristics known to all the Percida are : — Fin- rays partly spinous and partly flexible. Body oblong, and covered with hard or rough scales. Bones of the gill-covers variously armed at the edges with notches or spinous points. Most of the species composing this family have the ventral fins under the pectoral. Principal Characteristics of tlic Perch. — The whole length of the fish, — head, body, and tail-fin— being considered as i, the length of the head alone, to the point of the gill-cover, is about 3A : depth of body at the deepest point as nearly as possible equal to length of head ; nape of neck and shoulder rising very abruptly, giving a general appearance of the fish being rather " hog-backed ;" belly-line less convex. Fleshy portion of tail very slender. Tail-fin rather small. Two back-fins, the first supported by strong pointed bony rays fourteen in number ; the second back-fin having sixteen soft rays, and commencing very nearly over the anal fin : the commencement of the first dorsal, pectoral, and ventral fins, very nearly in a line, slanting slightly backwards towards the belly; and the pectoral-fin com- mencing a very little more forward than the point of gill-cover. Colour : muzzle dark brownish green ; irides, cheeks, and gill- covers, a greenish bronze ; pectoral-fins nearly yellow of a lightish tint ; pectoral, anal, and tail fins, brilliant scarlet ; second back-fin greenish, with a slight tinge of scarlet or orange. Back dark olive green, becoming nearly brown in large speci- mens ; sides golden yellow marked with dark transverse bars of a greenish colour : belly from muzzle to tail quite white. Scales in lateral line about sixty-six. Lateral line convex. THE COMMON CARP.' H E family of Carps (Cyprinidcc) belongs to the first Order of fishes, Malacop- tcrygii, or those having almost all their fin-rays soft and flexible, and the genus Cyprimis, or true Carps {the first genus of this family), are distinguished by — the body being always covered with large scales ; a single long back-fin ; the lips fleshy ; mouth small ; teeth in the throat, but none in the jaws ; and three gill-rays. To the family Cypriiiida: belong no less than twenty-two species found by ichthyologists in our fresh waters; including the Roach, Dace, Chub, Barbel, and Bream, constituting some of the most prominent of the so-called " coarse fish," regarded as objects of interest from the sportsman's point of view. The common Carp is the typical fish, and there- fore occupies first place in right of precedence ichthyological. The specimen selected for illustra- tion was taken from the Obelisk ponds in Windsor ' Cyprimis carplo. Cypriims, a Carp ; carpia, the Carp, Lat. y an enlargement of the anterior, or more for- OUTLINES OF ICHTHYOLOGY. 151 ward, end of the nervous axis the brain is formed, with which the nerves of smell, sight, taste, and hearing are connected. Immediately under the spinal column, and divided from each other by their membranes, are situated the organs of respi- ration, circulation, digestion, excretion, and also, when it exists, the air- or swim-bladder. This type of structure is common to all vertebrate animals ; but the peculiar attribute of Fishes is, that in them the function of respiration, or oxygen- ation of the blood, is performed through the me- dium of zuatcr, not merely temporarily, but during the continuance of life. Duly to apprehend the various ' plans ' on which fish are constructed requires a thorough acquain- tance with piscine organization in all its forms. Even a condensed treatise on such a subject would necessarily occupy a volume. In the present out- line, therefore, I shall only attempt to bring before the reader the general characteristics and most remarkable features, his attention being more par- ticularly directed to those organs upon the modifi- tions of which the classification of the various families and species, especially of fresh-water fish, is based. The first and most important function of fish and all other animals is breathing. For the pur- pose of breathing fish are provided on each side of 152 OUTLINES OF ICHTHYOLOGY. the neck with gills {branchice, Latin), consisting of arches of bone, to which the filaments, or spongy- portions, of the gills are attached, and having their surfaces covered by a tissue of innumerable blood-vessels. Of these branchial or gill-arches, there are in fish five, of which four support the gills, the fifth being appropriated to the " dentiferous pharyn- geal arch," a long arch in the throat, bearing teeth. In breathing the water is taken in at the mouth, and, after passing through the gills, is allowed to escape behind by the gill-openings. In its passage through the gills and gill filaments, the water im- parts to the latter the oxygen of the air which it contains, and receives carbon in return, as in the lungs of an air-breathing animal. The gills do not decompose the water, so as to derive oxygen from it, but merely separate the oxygen from the com- mon air contained in the water. Hence, if water is deprived of this air, or impregnated with un- wholesome gases, fish cannot exist in it. The gills also possess the power of imbibing oxygen not only from the portion of the air which is mixed with the water, but also directly from the atmospheric air itself; and this process may fre- quently be noticed in a vivarium, or other recep- tacle for fish, where the water is foul, or not changed sufficiently often, when the fish will be observed to remain for long periods together with their noses protruding out of the water. The OUTLINES OF ICHTHYOLOGY. 153 great majority of fishes, however, cannot continue to breatlie in atmospheric air alone beyond a very limited period, as, without the aid of fluid, the leaves of the gills, or gill filaments, shortly become stuck together, when, of course, the air cannot pass between them, and they lose the power of imbibing oxygen. From this results the gasping of fishes out of water, which is the effort of nature to separate the gill filaments. The blood is propelled to the gills by the heart, which thus answers to the right ventricle of warm- blooded animals ; and from the gills it is sent to an arterial trunk, lying immediately under the backbone, forming the left or systemic ventricle of the heart, which sends the blood throughout the body. In intimate connection with the organs of res- piration occurs the question of the blood tem- perature. The consumption of oxygen in all fishes is comparatively small, which may be the cause of the cold nature of their blood. The temperature of the bodies of fish that swim near the bottom is seldom more than two or three degrees higher than that ofthe water at its surface. In surface-swimmers a temperature of 10° Fahr. above that ofthe water has been occasionally found ; and it may be re- ceived as a general law, that those fish which swim near the surface have a high standard of rcspira- 154 OUTLINES OF ICHTHYOLOGY. tion (and, therefore, of temperature), die rapidly on being taken out of the water, and have flesh prone to quick decomposition, and vice versa. Two better examples of these laws could not perhaps be given than those of the Bleak, a surface-swimmer, and the Tench, a ground-swimmer. Amongst sea- fish, Mackerel, Salmon, and Herrings may be quoted as instances of the former law, and Eels, Flat Fish, and Skate, of its converse. ' With a low standard of respiration, and tenacity of life, is connected the extraordinary power exhi- bited by some species of sustaining extremes of. high and low temperature. The Gold fish lives and thrives in water as high as 85° Fahr. Fishes exist in the hot springs and baths of various coun- tries, ranging from 113° to 120°; and in South America Humboldt states that he saw fish thrown up alive, and apparently in health, from the bottom of a volcano, along with water and heated vapour which raised the thermometer to 210°, or two degrees below boiling point. On the other hand, in parts of Europe and North America, Perch and Eels, and several species of Carp, are not uncom- monly frozen and thawed again, and even trans- ported from place to place in a frozen state, with- out injury to life. - Passing to the senses with which fish are en- dowed, we come, as first in importance, to the sense of Sight. The eyes of fish occupy entirely OUTLINES OF ICHTHYOLOGY. 155 different positions in different species, — in some, such as the Slra/ichicr, gills, and fixis, fixed.) — These fish have their gills fixed at the outer edges, with a separate opening through which the water from each gill escapes. They have also small cartilaginous arches suspended in the muscles opposite the gills, which may be called gill-ribs. They form two Families. The foregoing eight Orders are again subdivided into FamiHes, Genera, and Species, as already stated. The following List exhibits in a tabular form the classification above explained, with the addi- tion of the names of the various Families : — 2 o 1— 1 H < U 1— < l-H C/5 c/) W fe CO 5 < J o u w K in fe E.S -2 ^__ M s ; , Ob '^ ?^ -J:-; ?■! f/V W ffi rn t— < te 1 • C/J •— ' D CO n iz; a! »— ( (73 h4 t-H H Pi < U So," urs-^ •50? 5 K i! •HO2- gj-g S„-g -JEo: ^iTi c^^ -a li y.9-,% / 4) U.jC ■" > ^ '— r-i ,' /-- V -^ ^ •- ji<; 0 ^ a, — — ^ 1 ■-C •sits 1 -si^l^ 5-S ■^l^.i OQ-fc.'-" ^ -5 J:'>,^-3 HON ■/ac/ti- ■S 3 •^ ^- ■0 " - ' _ . r^^ = =":e.^ c2 „ ii2 1; 'O .~ '-J T d 0 V i i Jl V it ^^ M ^ 3 tn ^ rt -* ;; 1L> _^ X ^ ~; ^ -5 J3 ■i^|l^ §2 1 0 2 " -- ;5 " 2 « C rt UM f 13 Sj=j:i - ."2 bfl :: rt " c ^ S-C -^ -C -y; 1* U O S f-^-c-^ Otn > 4J -• ^i T ^-i' •£■6 ■203 amilies, rinidce rp fami T'i ■s 1 111 i '2 k^ i 4S 2< ^ « rn ■»■ ih rt S aj3 u 0. o X H < = a, E „ S -S •^ < ^i^i: Cs. U. !>; C/f ^ L^ C/5 11 N 178 OUTLINES OF ICHTHYOLOGY. The following is a classification, according to their families, genera, &c., of fish inhabiting British fresh waters. It will be observed that with the ex- ception of the Lampreys and Lamperns the whole of them belong to the first series, which we have seen consist of the True, or bony, fishes. The Lamperns and Lampreys belong to the second, or Cartilaginous, series of fish. In this list I do not include doubtful species, such as the so-called Loch Leven Trout ; Gold and Silver fish (the common gold fish of the aquaria) ; Golden Tench ; or Salino Fontmalis, and other species of an admittedly non-indigenous character, or only recently acclimatized. In regard to the technical names of the fish, the first indicates the genus, and the second the spe- cies, as, for example, Pcrca Jluviatilis — Perca re- presenting the genus, and Jluviatilis the species — the Perch of the river, — and so on. 1ST Series. TRUE, OR BONY, FISH. Order I. Spiiiy-finncd Fish. Family i. Perches — Percid.e. Species. Perch. {Perai Jiiivia/ilis.) Ruffe, or Pope. {Aicriiia vii/givis.) Family 2. Fisii with hard cheeks. Species. Bullhead, or Miller's Thumb. {Q'llt^s gobio.) OUTLINES OF ICHTHYOLOGY. 179 Rough-tailed Stickleback. (Gastcroslciis trachunis.') (Five other species.) Order II. Soft-fimied Fish with ventral fins on the belly. Family i. Carps — CvPRiNiDyE. Species. Common carp. {Cyprinus carj>w.) Prussian, or Gibel, Carp. {Cyfrinus f;ibi-lio.) Barbel. {Barbus vulgan's.) Gudgeon. ( Gohio fluviatilis. ) Tench. {Tiwa vulgaris.) Bream, or Carp Bream. {Abramis braiiia.) White Bream, or Bream-flat. {Abramis b/iaa.) Pomeranian Bream. [Abramis Buggcnhagii.) Dace. {Leuciscus vulgaris.) Roach. {Leuciscus riitilus.) Rudd, or Red-eye. {Leuciscus erythrophthalmus.) Chub. {Leuciscus cephalus.) Graining. {Leuciscus Lancastriciisis.) Bleak. {Leuciscus alburnus.) Minnow. {Leuciscus p/uwinus.) Loach. {Cobitis barbatula.) Spined Loach, or Groundling. {Cobitis taiiia.) Family 2. Pikes — Esocid.'e. Species. Common Pike, or Jack. {Esox lucius.) Family 4. Salmon and Trout — Salmonid.^. Species. Salmon. {Salmo salar.) Bull Trout. (Grey Trout, Sewin, or Roundtail.) {Salmo eriox.) Sea Trout (or Salmon Trout). {Saliuo trutla.) Common (or Yellow) Trout. {Salmo fa rio.) Great Lake Trout. {Salmo ferox.) Charrs. i8o OUTLINES OF ICHTHYOLOGY. Grayling. (^Thyviallus 'i-uh^iiris.) Fresli ^\■atel• Herrings. {Corcgoiii.) (Four species or varieties.) Order III. Soft-finncd Fish having lower arms. l"'amily i. Cod — G.vdid.e. Species. IJmbot, or Eel-pout. {Lota vn/gan's.) Order IV. Sflfi-Fiimcd Fish lacking ventral fins. Family i. Eels — MuR/ENID/E. Species. Sharp-nosed Eel. {Anginlla aciitiri'stris.) Broad-nosed Eel. {Aiigiii/ia laiirosiris.) Snig. {Angiiilla mcdiorostris. ) 2ND Series. CARTILAGINOUS FISH. Order II. Fish with Fixfd GUIs. Family i. T^ish with mouth formed in a sucker. CVCLOSTOMATA. Species. Lamprey, or Sea Lamprey. {Pdiomyzflii inariniisi) Lampern, or River Lamprey. iPctromyzimJliiviatilis.) Fringed-lipped Lampern. [Pctroiiiyzpn P/ancri.) To give a practical example of this classifica- tion : — The Common or River Perch is 2i species of the gcfuts Pcrca, of the Fajnily of the Pcrcidcr, of the Order Acanthroptcrygii, belonging to the True or Bony series of fish ; or, in other words, is the first .Species, of the first Genus, of the first Family, of the first Order, of the first Series. OUTLINES OF ICHTHYOLOGY. iSi In conclusion, a few suggestions for enabling the spoctsman more readily to identify his fish, and for the treatment of any supposed new species or varieties, may possibly not be altogether out of place. First, then, upon the capture of any such specimen, the best and simplest course is imme- diately to preserve it in methylated (or common) spirit of wine — or failing these, any other strong spirit — when it can be examined and pronounced upon at leisure. Dried or stuffed fish are compa- ratively useless for scientific purposes. If, how- ever, from any cause it should be found imprac- ticable to preserve the specimen, the next best thing is a correct description in writing ; and to give this with sufficient accuracy to make it of value to science — to render it, in short, such that a natu- ralist shall be justified in pronouncing upon it with reasonable certainty — all that is required is the clear comprehension and record in an intelligible form of a few simple facts. The first point to be noted is the nature of the Fins — whether generally soft and flexible as in the Salmon and Carp, or hard and spiny as in the Perch : upon this it depends in which of the two principal divisions of Bony Fishes the species should be placed. This distinction is so obvious and remarkable, that when once mentioned it cannot be overlooked by the least careful ob- server ; and as to one or the other of these divisions belongs ever)' fish, without exception, of which the 1 82 OUTLINES OF ICHTHYOLOGY. Sportsman takes cognizance, its importance is evi- dent. The spiny-finned fishes contain fifteen Families, but no great subordinate divisions. The soft-finned fish, on the contrary, are divided into three strongly-marked Orders founded upon the position, or absence, of the ventral fins; and to the first of these — vialacoptcrygii abdoniinalcs — belong the whole of our soft-finned sporting fish. The division in which any fish is to be classed having been thus decided, it remains only to deter- mine its family and species. As regards family no difficulty whatever can be experienced, there being only four families in all to which it can belong, and these being clearly re- cognizable by the most obvious signs. To commence with the spiny-finned fishes : — All the sporting fish of this order belong to one Family, the Percidce — of which the Perch is the type ; whilst those having soft fins are embraced in three families : the Cypi-inidcc, of which the Carp is the type ; the Esocidcc, of which the Pike is the type ; and the Salmonidcr, of which the Salmon is the type. In regard to these families, again, no difficulty can be found, every species of the Salmon possessing the peculiar characteristic of two back fins, and no other species of the same Order more than one, — the Pike, of which we have only a single variety, being totally dissimilar from every other family, — and the remaining species OUTLINES OF ICHTHYOLOGY. 183 without exception belonging to the Cyprinidae, or Carp family. To decide with certainty between different genera, or different species of the same genus, re- quires a more minute observation and comparison, according to the " characteristics " which will be found appended to each. The points upon which such minor distinctions depend, and which will consequently be those to be particularly observed and noted, are : — (a.) The shape of the body, which can be best ascertained by placing the fish straight on a sheet of white paper, and, when it has stiffened, carefully tracing the outline with a pencil. (b.) The relative positions of the yfw (see p. 170); and their form, more especially as regards the back- and tail-fins, whether forked, concave, square, or convex, (c.) The form of the head z.nA gill-covers (see p. 16S), and their length from the tip of the muzzle to the hinder margin, as compared with that of the whole body, measuring from the tip of the muzzle to the extremity of the tail, (d.) The nature of the tcctk and the names, if possible, of the boius on which they are placed (see pp. 171-174). (e.) The colouring of the body and fins of the fish, when first caught, (f.) The number of scales forming the lateral line. (G.) The number of 7?«-;i?)'^, or sup- ports, in each fin. These, in the tail fin, are counted from the first long ray out- side, either above or below; and in the other fins in all cases from the first ray nearest the head of the fish. The fin-rays are denoted, as in these pages, by placing the initial letter of the name of the fin before the numeral express- ing the number of rays it contains — as D. (for " Dorsal ") 9 : P. (for "Pectoral") 15 : &c. Take, for example, the fin-rays of the Gudgeon, which are particularly simple : — D. 9: P. 15 : V. 8: A. 7: C. igj. Occasionally there are short incumbent rays on the base of i84 OUTLINES OF ICHTHYOLOGY. the tail fin, above and below the longest rays, and these are frequently omitted from the reckoning ; when they are men- tioned, it is in general in the shape of a fraction ])laced after the number of the full-length rays, the numerator denoting those above the fin, and the denominator those below it, as in the example. When there is more than one dorsal fin, the rays are enumerated with a line between them — • D. 9 — 12 : the first numeral having reference to the first, or more forward, fin. In the fish having two dorsals, as the Perch, the first is usually wholly spinous, and this is indicated as in the last case, by a line between the two numbers ; when, however, there is but one dorsal, of which the fore part only is spinous, the sign + is placed between the spinous and soft rays. This is also the rule in regard to other fins partly spiny and partly soft : thus — (Fin-rays of Perch.) D. 15 — 1 + 13: P. 14: V. 1+5: A. 2+8: C. 17. Some of the soft-finned fish have an occasional spinous ray ; and when this is the case their position, &c., should be fully described — a perpendicular line after such spinous ray being drawn in the ray formulary. Take the fin-rays of the Common Carp for example : — D. 4/19: P. 17: V. 9: A. 3/5: C. 19. In this instance the fourth dorsal and third anal fin-rays are spinous. T\^>i form of the gill-covers has been mentioned as one of the points to be relied on for distin- guishing one species from another ; for whilst, owing to circumstances of food, water, or climate, different individuals of the same species will often differ widely from each other, and from their ori- ginal type (in so far at least as regards those ex- ternals which most readily strike an unscientific OUTLIXES OF ICHTHYOLOGY. 1S5 eye), the peculiarities of structure of the gill-cover, apparently of little moment, ma)-, from their im- portance and permanence, not unfrequently indi- cate totally distinct and immutable species. Any permanent and unvarying difference in the form of the head, formation of the gill-covers, and dental system, coupled with variations of colour, form, habit, or the like, which might otherwise be deemed casual, are held to constitute a distinct species. A good example of this difference is seen in the form of the gill-covers of the Salmon and the Sea Trout — the hinder marcjin of the oill-cover formin^r in the true Salmon almost a semicircle, whilst that of the Sea Trout approaches more nearly a right angle, and both differ somewhat from the gill- covers of the Bull Trout. CHISWICK PKE6S ;— C. WHITTINCHAM .AND CO., TOOKS COUKT, CHANCERY LANE. iSoofes on Angling FLOATING FLIES AND HOW TO DRESS THEM. A Treatise on tlic most Modern Methods of Dressing Artiticial Flies for 'iVout and Grayling. W'liU fail Illus- trated Directions, and containing Ninety Hand-coloured Engravings of the most Kiilnig Patterns, together with a few Hints to Dry-Fly Fishermen. tSy Fk^uekic M. IIai.i-iii-;i), '* Detached Badger '* of " The Field," Member of the '* Houghton Club," '' Fly Fishci.'," Club," &c. A large-paper edition, printed on Dutch hand-made paper, limited to loo copies for England and 50 for America, price ^os. {This Edition is exhausted.) Small-paper edilion — Demy 8vo, first edition limited to 500 copies, cloth, 15J., post /ret: THE SPORTING FISH OF GREAT BRITAIN. WITH NOTES ON ICHTHYOLOGY. My H. Cholm..ndki.ey-P££NNELL, Ute Inspector of Sea Fisheries. Author of ■■ Salmon and Trout." " Pike and other Coarse Fish, Phe Modern Practi^jal Angler/' "The Angler-Naturalist," "The Book of the Pike," Kditor of the late " Fisher- man's Maga?ine and Review." Illustrated by Sixteen Lithographs of Fi^h ia Gold, Silver, and Colours. .\ large-paper edition, printed on Dutch hand-made paper, limited to 100 copies for England and 50 for America. Small-paptr edition — Demy 8vo, first edition limited to 500 copies. iJiist Kiady. NORTH-COUNTRY TROUT FLIES. By T. E. Pritt. With Coloured Plates of all the best Flies. Second Edition, cloth, demy 8vo, 10^. 611'. Nijw RtADv. Small 410, 364 pages, cloth extra, loj. 6d. FLY-RODS AND FLY-TACKLE. Suggestions as to their Manufacture and Use. Uy HliNuv P. Welis. Illustrated. 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