A Stereo-Atlas of Ostracod Shells edited by R. H. Bate, D. J. Horne, J. W. Neale, and David J. Siveter Volume 11, Part 1; 29th June, 1984 Published by the British Micropalaeontological Society, London Editors Dr R.H. Bate, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking, Surrey GU23 7EF. Prof. J.W. Neale, Department of Geology, The University, Hull HU6 7RH. Dr D.J. Horne, Department of Geology, City of London Polytechnic, Walburgh House, Bigland Street, London El 2NG. Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH. Editorial Board Dr G. Bonaduce, Stazione Zoologica, 80121 Napoli, Italy. Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles, France. Dr P. De Deckker, Research School of Pacific Studies, Australian National University, PO Box 4, Canberra ACT 2600, Australia. Dr D. van Harten, Universiteit van Amsterdam, Geologisch Instituut, Nieuwe Prinsengracht 130, Amsterdam, The Netherlands. Dr I. Purper, Departamento de Paleontologia e Estratigrafia, UFRGS, 90 000 Porto Alegre RS, Brazil. Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55, D 2000 Hamburg 13, West Germany. Officers of the British Micropalaeontological Society Chairman Dr R.H. Bate, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking, Surrey GU23 7EF. Secretary Dr P.P.E. Weaver, Institute of Oceanographic Sciences, Brook Road, Wormley, Godaiming, Surrey GU8 5UB. Tel: 042-879 4141. Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History), Cromwell Road, London SW7 5BD. Tel: 01-589 6323. Journal Editor Dr. L.M. Sheppard, SSI (U.K.) Limited, Tannery House., Tannery Lane, Send, Woking, Surrey GU23 7EF. Newsletter Editor Dr R.L. Austin, Department of Geology, The University, Southampton S09 5NH. Tel: (0703) 559122/557941. Conodont Group Chairman Dr R.L. Austin, Department of Geology, The University, Southampton S09 5NH. Secretary Dr H. A. Armstrong, Department of Geology, The University, Newcastle-upon-Tyne NE1 7RU. Tel: (0632) 328511. Foraminifera Group Chairman Dr M.D. Brasier, Department of Geology, University of Hull, Hull HU6 7RX. Secretary Dr J.V. Weston, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking GU23 7EF. Tel: (0483) 223902. Microplankton Group Chairman Dr G.C. Wilkinson, Britoil, 150 St. Vincent Street, Glasgow G2 5LJ. Secretary Dr S.G. Molyneux, British Geological Survey, Ring Road, Halton, Leeds LS15 8TQ. Tel: (0532) 605343. Ostracod Group Chairman Dr J. Athersuch, B.P. Research Centre, Chertsey Road, Sunbury-on-Thames, Middlesex TW16 7LN. Secretary Mr. I.P. Wilkinson, British Geological Survey, Nicker Hill, Keyworth, Nottingham NG12 5GG. Tel: (06077) 6111. Palynology Group Chairman Dr M.C. Boulter, Palynology Research Unit, N.E. London Polytechnic, Romford Road, London E15 4LZ. Secretary Mr N. Hooker, Britoil, 150 St. Vincent Street, Glasgow G2 5LJ. Tel: 041-204 2525. Calcareous Nannofossil Group Chairman Dr M.K.E. Cooper, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking GU23 7EF. Secretary Miss H. Stowe, Micropalaeontology Unit, University College, Gower Street, London WC1E 6BT. Tel: 01-387 7050. Instructions to Authors Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited. Format should follow the style set by the majority of papers in this issue. Descriptive matter apart from illustrations should be cut to a minimum; preferably each plate should be accompanied by one page of text only. Blanks to aid in mounting figures for plates may be obtained from any one of the Editors or Editorial Board. Completed papers should be sent to Dr David J. Siveter. The front cover shows a female left valve of Hemicythere villosa (Sars, 1866) Printed in the UK by UPS Blackpool Ltd., Stanley Road, Blackpool Stereo-Atlas of Ostracod Shells 11 (1) 1-4 (1984) Hippula turris (1 of 4) 595.336.13 (113.312) (430.2 : 161.013.54) : 551.35+ 552.55 ON HIPPULA (CETONA) TURRIS (SCHALLREUTER) by Roger E. L. Schallreuter (University of Hamburg, German Federal Republic) Hippula (Cetona) turris Schallreuter, 1967 1967 Oecematobolbina (Cetona) turris sp. n. R. E. L. Schallreuter, Neus . Jb. Geol. Palaont. Mh . , 1967 (7), 445, 446, fig. 7.3-4. 1970 Hippula turns ', R. E. L. Schallreuter, Hercynia N. F., 6 [1969] (3), tab. 2 (294/5). 1973 Hippula (Cetona) turris (Schallreuter); W. Neben & H. H. Krueger, Staringia, 2, pi. 94, fig. 6. 1977 Hippula turris ', R. E. L. Schallreuter, Palaont, Z., 51 (1/2), 38. 1980 Hippula (Cetona) turris (Schallreuter); R. E. L. Schallreuter & M. Kruta, Neus. Jb. Geol. Palaont. Mh., 1980 (8), 506. 1982 Hippula turns', R. E. L. Schallreuter, Palaeontographica (A), 178 (1/3), 27. Holotype: Sektion Geologische Wissenschaften, University of Greifswald, German Democratic Republic; no. 29/15, posteriorly incomplete 9LV. Type locality : Beach at Dornbusch, Isle of Hiddensee (Baltic Sea), Germany; lat. 54°36'N, long. 13° 7' E, Backsteinkalk erratic boulder (no. and type 1 B 1 ; equivalent of the upper Viruan Skagen Limestone of Central Sweden), middle Ordovician. Figured specimens: Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) nos. 2739 ( 9RV: PI. 11, 2, fig. 1), 2740 (cTLV: PI. 11, 2, fig. 2) and 2741 (posterodorsally incomplete 9LV: PI. 11, 4, figs. 1, 2). Nos. 2739 and 2740 are from Backsteinkalk erratic boulder no. Gis 30 (type 1B14; equivalent of the upper Viruan upper Dalby Limestone of Central Sweden), from the beach of Gislovshammar, SE Scania, Sweden; lat. 55° 29' N, long. 14° 18' E; coll, by the author in 1978. No. 2741 is from Backsteinkalk erratic boulder no. Stal (type 1B1), from the beach at Staberhuk, Isle of Fehmarn (Baltic Sea), Germany ; lat. 54° 24.5' N, long. 11° 19' E; coll, by the author in 1980. Explanation of Plate 11, 2 Fig. 1, 9RV, ext. lat. (GPIMH 2739, 1.13 mm long excluding spines); fig. 2,cfLV, ext. lat. (GPIMH 2740. 1.05 mm long excluding spines). Scale A (100 /nm; x 72), fig. 1; scale B (100 /tm; x 91), fig. 2. Stereo- Atlas of Ostracod Shell 11, 3 Hippula turris (3 of 4) Diagnosis: Species of Hippula (Cetona) with S2 distinct, long and sigmoidal, dorsally rather deep. Preadduc- torial node rather distinct; a relatively broad, conical bulb. Posteroventral lobe strongly developed, stituated ventrocentrally, ending in a long reticulate spine. Velar flange in males forming an angle with the lateral surface of about 90°, in females rather obtuse. Torus very broad (transverse), nearly (9) or as broad (tf) as velar flange. Adult valves c. 1.00-1.13 mmm long (excluding spines). Length: height ratio 1.60-1.80. Remarks: Ivanova ( Trudy-paleont . Inst., 172, 36, 1979) considered the cavities of the “histial” flange of the females of Oecematobolbina as a prototype of the locular type of dimorphism. This is considered impossible because loculi are always formed on the inner side of the dolon (the antrum), not the outer side. Copeland (Bull. geol. Surv. Can., 347, 16, 1982) considered Schallreuter’s subgeneric distinc- tion within Hippula untenable in the light of his material from the District of Mackenzie, Canada. In H. varicata he described the “histium” as consisting of three fluted flanges on the heteromorphic valve, and a single flange on the tecnomorphic valve. He could not observe this phenomenon in the European material. As shown herein (PI. 11. 2, fig. 2) the males also possess the characteristic torus with an entervening row of diamond-shaped spaces. Perhaps the North American forms exhibit a tendency to reduce the tori. Parahippula ventrospinosa (Kraft, Mem. geol. Soc. Am., 85, 1962) resembles Hippula in having radiating furrows of the velar flange in both dimorphs (op. cit., pi. 11, figs. 14a, 15, 17, pi. 12, figs. 4, 5, 7 cf. PI. 11, 2, figs. 1, 2). Such furrows could be the rudiments of the cavities. The missing tori in the tecnomorphs of H. varicata could be explained as an example of proterogenesis. Distribution: Known only from upper Viruan Backsteinkalk erratic boulders of northern Germany and southern Sweden (types 1B1, 1B3, IB 14, cf. R. E. L. Schallreuter, op. cit., 1970), middle Ordovician. Explanation of Plate 11, 4 Figs. 1-2, posterodorsally incomplete 9LV (GPIMH 2741, 1.07mm long excluding spines): fig. 1, ext. lat.; fig. 2, ext. ventrolat. Scale A (100/u.m; x 82), figs. 1, 2. Stereo- Atlas of Ostracod Shells 11,4 Hippula turns (4 of 4) Stereo-Atlas of Ostracod Shells 11 (2) 5-8 (1984) Schallreuteria lippensis (1 of 4) 595.336.13 (113.312) (430.1:161.010.54) : 551.35 + 552.55 ON SCHALLREUTERIA (LIPPEA) LIPPENSIS SCHALLREUTER subgen. et sp. nov. by Roger E. L. Schallreuter (University of Hamburg, German Federal Republic) Genus SCHALLREUTERIA Siveter, 1982 1982 Schallreuteria gen. nov.; D. J. Siveter, Stereo-Atlas of Ostracod Shells, 9, 93-100, pis. 9, 94; 9, 96; 9, 98; 9, 100. Subgenus LIPPEA subgen. nov. Type-species: Schallreuteria (Lippea) lippensis sp. nov. As for the type-species. Subgenus of Schallreuteria with only one sulcus (S2) and less pronounced quadrilobation. The type-species of Schallreuteria, S. superciliata (Reed, 1910), from the Longvillian (Caradoc Series) of England, is distinctly quadrilobate (cf. Siveter, op. cit.) and is thus considered a more conservative form. It belongs to the main line of the genus from which S. ( Lippea ) is separated by its loss of quadrilobation which occured at the latest by the lower upper Viruan (= Hamagian/Soudleyan). The characteristic dorsal spines of lobes LI and L3 of S. superciliata are also present in 5. lippensis (cf. Siveter, op. cit., PI. 9, 96, fig. 2). The main differences between S. (Schallreuteria) and 5. (Lippea) are the same as in the genera Rakverella and Pectidolon . Therefore, Pectidolon is now considered to be a subgenus of Rakverella. Rakverella is characterized by special cristae which are not present in Schallreuteria (Siveter, 95). Schallreuteria (Lippea) lippensis sp. nov. 1970 Rakverella pectinata-, R. E. L. Schallreuter, Hercynia N. F., 6, (3), 289, tab. 2 (292, 293) (pars). 1973 Rakverella pectinata (Oepik); W. Neben & H. H. Krueger, Staringia, 2, pi. 92, fig. 5. Explanation of Plate 11, 6 Figs. 1-4, ?LV (holotype, GPIMH 2900, 1353 ;u.m long): fig. 1, ext. lat. ; fig. 2, ext. vent, obi.; fig. 3, ornament of lateral surface (pillars, removed pillars, and reticulation); fig. 4, detail of surface reticulation. Scale A (250 ^.m; x 58), figs. 1-2; scale B (50 /xm; x 290), fig. 3; scale C (50 gm; x 225), fig. 4. Derivation of name: Diagnosis: Remarks: Stereo- Atlas of Ostracod Shells 11, 7 Schallreuteria lippensis (3 of 4) 1976 Rakverella pectinata (Opik); R. E. L. Schallreuter, Palaeontographica (A), 153 (4/6), 203-205 (pars), pi. 6(39), figs. 1, 2. 1983 Rakverella pectinata (Opik); R. E. L. Schallreuter, Palaeontographica (A), 180 (4/6), 165, 166, 179 (pars), pi. 11, fig. 3. Holotype: Type locality: Derivation of name: Figured specimens: Diagnosis: Remarks: Distribution: Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH), no. 2900,? LV. [Paratypes: nos. 2902-2904]. Lower upper Viruan (middle Ordovician) 14B2-type Backsteinkalk erratic boulder of the Baltic group (cf. R. E. L. Schallreuter 1970, op. cit., 287), no. Lipl, from the beach at Lippe, Hohwacht Bay, Baltic Sea, Germany; lat. 54° 20.5' N,long. 1(> 38.5' E. After the type locality. GPIMH nos. 2900 ( ? LV : PI. 11, 6, figs. 1-4; PI. 11, 8, fig. 2), 2902(?RV:P1. 11, 8, fig. l)and 2901 (?RV : PI. 11, 8, fig. 3). Nos. 2900 and 2902 are from the type locality; boulder collected by the author in July 1983. No. 2901 is from Backsteinkalk erratic boulder no. G29 (cf. Schallreuter 1983, op. cit., 165); lat. 57° 18' N,long. 18° 8' E. Species of S. (Lippea) with LI as a strong, broad dorsal spine, L2 a slender dorsal spine; weak pre- adductorial node and weak anteroventral node; L3 an elongate lobe with strong dorsal spine and weak posteroventral lobe-like elevation; L4 a slender dorsal spine and very weak ventral inflation. Only weak laterovelar furrow. Special reticulation (net standing on pillars), different on lateral surface and dolon. Females 1.30-1.39 mm long. The material was formerly assigned by Schallreuter (1983) to Rakverella pectinata (Opik), which was considered by Henningsmoen (1953) and Sarv (1959) as synonymous with R. bonnemai (Schallreuter 1976, op. cit., 204). The more abundant and better preserved material now in hand shows that the material does not belong to R. pectinata and that Henningsmoen and Sarv were apparently correct in considering that species synonymous with R. bonnemai. The holotype of R. pectinata is a steinkern, but the size and arrangement of the posterior spines is similar to that of R. bonnemai. Backsteinkalk erratic boulders (14B2-type) of northern Germany and Sweden (Isle of Gotland, Baltic Sea). Boulders Lipl, 14B2, 812, G14, G29 and G39; middle Ordovician. Explanation of Plate 1 1, 8 Fig. 1, ?RV, ext. lat. (paratype, GPIMH 2902, 1314 long); fig. 2, ?LV, detail of ornament of dolon (holotype, GPIMH 2900); fig. 3, 9RV, int. lat. (GPIMH 2901, 1390 pm long). Scale A (250/xm; x 59.5), figs. 1, 3; scale B (50/xm; x 350), fig. 2. Stereo-Atlas of Ostracod Shells 11. 6 Schallreuteria lippensis (2 of 4) Stereo-Atlas of Ostracod Shells 11,8 Schallreuteria lippensis (4 of 4) Stereo-Atlas of Ostracod Shells 11 (3) 9-12 (1984) Duringia spinosa (1 of 4) 595.336.13 (113.313) (430.2 : 161.011.50) : 551.35 + 552.55 ON DURINGIA SPINOSA (KNUPFER) by Roger E. L. Schallreuter ( University of Hamburg, German Federal Republic) Genus DURINGIA gen. nov. Type-species: Eurychilina spinosa Kniipfer, 1968 Derivation of name: Duringia, an old name for Thiiringen, the country of the type locality of the type-species. Diagnosis: A medium-sized, possible genus of the Piretellinae, with S2 long and sigmoidal and preadductorial node as a distinct bulb. Tecnomorphic velum appears as a row of spines. Females with a strongly convex tubulose dolon, having a row of spines sited at the border of the lateral and marginal surfaces. Lateral surface spinose. Remarks: The presumably tubulose velum places the genus within the Eurychilinacea. The female velum mostly resembles that of Piretella (cf. Schallreuter, Palaeontographica (A) 149 (4/6), pi. 26(5), figs. 1, 4, 6, pi. 27(6), fig. 1) but the dolonal spines are missing in Piretella. Another distinguishing feature is the long sigmoidal S2 of Duringia which is present elsewhere notably in the Oepikiidae (Schallreuter, op. cit., 175). The assignment of Duringia to the Piretellinae (Eurychilinidae) is therefore questionable. With respect to the tecnomorphic velar spines and the female dolonal spines Duringia strongly resembles Hithis (cf. Schallreuter, Palaeontographica (A), 144(1/3), pi. 17, figs. 1-3, 1973; Schallreuter & Siveter, StereoAtlas Ostracod Shells, 9 (2) 15, 85-88, 1982), Piretia (Schallreuter, op. cit., pi. 17, fig. 7, pi. 18, fig. 1) and Bromidella (Copeland, Bull. geol. Surv. Can., 347, pi. 2, figs. 21, 22, 1982), but in all these cases the dolon exhibits no tubules. Explanation of Plate 11, 10 Fig. 1, dors, and posterovent. incomplete ?9LV, ext. lat. (GPIMH 2727, 1.19mm long); fig. 2, fragmentary ?LV, int. obi. (GPIMH 2728); fig. 3, posterovent. incomplete tecnomorphic LV, ext. lat. (GPIMH 2729, 0.83 mm long excluding spines). Scale A (100 /am; x 65), figs. 1, 2; scale B (100 pm; x 90), fig. 3. Stereo- Atlas of Ostracod Shells 11, 11 Duringia spinosa (3 of 4) Duringia spinosa (Kniipfer, 1968) 1963 Eurychilina', H.Blumenstengel et. al., Geol. Ges. DDR Exkursionsfuhrer zur Herbsttagung, 1963, 6. 1968 Eurychilina spinosa n. sp. J. Kniipfer, Freiberger ForscHft. (C), 234, 9, 10, 24, 25, pi. 4, figs. la-b. 1973 Piretia ? spinosa (Kniipfer); R. E. L. Schallreuter , Palaeontographica (A), 144 (1/3), 89. 1980 Piretia ? spina (Kniipfer); G.Qvale, Norsk geol. Tidsskr. 60 (2), 94. Holotype: Type locality: Figured specimens: Diagnosis: Remarks: Distribution: Geologisches Institut, Bergakademie Freiberg, Sachsen, German Democratic Republic; no. 45/1024, larval RV. Middle adit of the Iron-ore mine Gebersdorf, Thuringia; lat. 50° 32' N, long. 11°17'E; limestone layer of the upper layer of the Upper Ore Horizon (Oberes Lager des Oberen Erzhorizontes = uppermost Caradoc, zone 13), Grafenthal series. Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) nos. 2727 ( 9LV: PI. 11, 10, fig. 1), 2728 (fragmentary 9 LV: PI. 11, 10, fig. 2), 2729 (larval LV: PI. 11, 10, fig. 3), 2730 (larval RV: PI. 11, 12, fig. 1), 2731 (larval RV: PI. 11, 12, fig. 2) and 2732 (larval RV: PI. 11, 12, fig. 3). All the figured specimens are from the limestone layer of the Upper Ore Horizon of the former open-pit iron-ore mine Wittmannsgereuth, on the ‘Breiten Berg’ near Saalfeld, Thuringia, German Democratic Republic (H. Blumenstengel et. al., op. cit., 5-7, fig. 1); lat. 50° 39' N,long. 11° 19.5' E; coll, by the author in about 1965. All the material is coarsely silicified. As for the genus. Kniipfer (1968) in his original description had in hand only tecnomorphs. He assigned the material to Eurychilina apparently based only on its similarity withE. multipustulosa Swain (7. Paleont., 36 (4), 727, 1962), a species now placed in synonymy with Bromidella spiveyi (Copeland, Bull. geol. Surv. Can., 347, 32, 1982). The latter species possesses an S2 sulcus developed as a pit (as in Uhakiella ) and a dolon without tubules (op. cit., pi. 6, figs. 3-8, 20-22). Known only from the type stratum in Thuringia, German Democratic Republic (localities given above). Caradoc Series, Ordovician. Explanation of Plate 11, 12 Fig. 1, posterodors. incomplete larval RV, ext. lat. (GPIMH 2730, 0.61 mm long excluding spines); fig. 2, larval RV, ext. lat. (GPIMH 2731, 0.70 mm long); fig. 3, early larval RV, ext. lat (GPIMH 2732, 0.47 mm long). Scale A (100 pm-, x 100), fig. 1; scale B (100 jam; x 92), fig. 2; scale C (100 jam; X 135), fig. 3. Stereo-Atlas of Ostracod Shells 11, 10 Duringia spinosa (2 of 4) Stereo- Atlas of Ostracod Shells 11, 12 Duringia spinosa (4 of 4) Stereo-Atlas of Ostracod Shells 11 (4) 13-16 (1984) Duringia triformosa (1 of 4) 595.336.13 (113.312) (429 : 162.005.51) : 551.35 + 552.55 ON DURINGIA TRIFORMOSA JONES sp. nov. by C. R. Jones ( University of Leicester, England) Duringia triformosa sp. nov. Holotype: Type locality: Derivation of name: Figured specimens: Brit. Mus. (Nat. Hist.) no. OS 12261; 9LV. Old quarry about 300 m south of Cwm Agol Farm, c. 7 km west of Llandeilo, Dyfed, Wales; approx, lat. 51°51'N, long. 4°05'W (Nat. Grid. Ref. SN 56552070). Llandeilo ‘Flags’, Llandeilo Series, middle Ordovician. Latin, formosus, beautifully formed; alluding to the three distinct morphological forms. Brit. Mus. (Nat. Hist.) nos. OS 12261 (holotype, 9 LV: PI. 11, 14, fig. 5), OS 12260 (juv. LV: PI. 11, 14, fig. 4), OS 12263 (cT? RV: PI. 11, 16, figs. 1, 2), OS 12262 (juv. RV: PI. 11, 16, figs. 3, 4), OS 12264 ( 9LV: PI. 11, 16, figs. 5, 6). One specimen ( ?LV: PI. 11, 14, figs. 1-3) was broken after photography. All the figured specimens are from the type locality and horizon, except for OS 12264, which comes from Capel Dewi quarry, 350m west of Ffynnon-Dewi, c. 15.5km west of Llandeilo, Dyfed (N.G.R. SN 47472063); Llandeilo ‘Flags’, lower Llandeilo, middle Ordovician. Explanation of Plate 11, 14 Figs. 1-3, ?LV (now broken, 1.30mm long): fig. 1, ext. lat; fig. 2, ext. ant; fig. 3, ext. vent. Fig. 4, juv. LV, ext. lat. (OS 12260, 1.02 mm long). Fig. 5,9LV, ext. lat. (holotype, OS 12261, 1.23 mm long). Scale A (250 /xm; x 42), figs. 1, 3; scale B (250 /xm; x 47), fig. 2; scale C (250 /xm; x 47), fig. 4; scale D (250/xm; x 45), fig. 5. Stereo- Atlas of Ostracod Shells 11, 15 Duringia triformosa (3 of 4) Diagnosis: Remarks: Distribution: Duringia with diminutive preadductorial node. Narrow depression (= remnant S3?) from postero- central region to dorsum. Females with weakly convex tubulose dolon, serrated distally. Tecno- morphic velum as row of spines (juveniles), or ventral flange (males?). Lateral surface spinose and granulose. D. triformosa is only the second described species of the genus. The younger type-species, D. spinosa (Kniipfer, 1968) (see Schallreuter, Stereo-Atlas of Ostracod Shells, 11 (3), 9-12, 1984), from the uppermost Caradoc of Thuringia, closely resembles/), triformosa but for the presence of its dolonal spines, more convex dolon, and lack of a remnant S3. Only one complete female valve of/), spinosa is known (1.19mm long), which falls within the large size variation displayed by D. triformosa (females from Llandeilo Series: 1.08-1.3+ mm long). Like the type-species, D. triformosa has a tubulose velum, justifying inclusion of the genus within the Eurychilinacea. However, the familial assignment of Duringia is questionable (Schallreuter, op. cit.). Schallreuter provisionally placed Duringia in the Piretellinae (Eurychilinidae) because the dolon of D. spinosa is similar but it also has dolonal spines not normally present in piretellines.Z). triformosa would support this assignment as it too lacks dolonal spines. However, the sigmoidal S2 of Duringia remains strikingly opikiid-like. The recognition of a remnant S3 (?) inD. triformosa may indicate quadrilobate ancestry; its familial assignment is therefore still uncertain. Llandeilo Series, and Costonian Stage, basal Caradoc Series, Dyfed, Wales and Harnagian Stage, Caradoc of Shropshire, England. Explanation of Plate 1 1, 1 6 Figs. 1, 2, ?d'RV (OS 12263, 1.27 mm long): fig. 1, ext. lat; fig. 2, ext. post. Figs. 3,4, juv. RV (OS 12262, 1.12 mm long): fig. 3, ext. ant.; fig. 4, ext. lat. Figs. 5, 6, large 9 LV, broken posteriorly (OS 12264, 1.3+ mm long): fig. 5, ext. ant. obi.; fig. 6, serrated terminations of the tubulose dolon. Scale A (250 /xm; X 43), fig. 1; scale B (250/xm; x 45), fig. 2; scale C (250/xm; x 44), fig. 3; scale D (250 /im; x 46), fig. 4; scale E (300 /xm; x 35), fig. 5; scale F (100 /xm; x 80), fig. 6. Stereo-Atlas of Ostracod Shells 11, 14 Stereo- Atlas of Ostracod Shells 11, 16 Duringia triformosa (2 of 4) Duringia triformosa (4 of 4) I Stereo-Atlas of Ostracod Shells 11 (5) 17-20 (1984) Hamanella implexa (1 of 4) 595.337.14 (118.15/118.21) (794 : 162.120.34) : 551.353 + 552.52 ON HAMANELLA IMPLEXA FINGER by Kenneth L. Finger ( Chevron Oil Field Research Company, La Habra, California, U.S.A.) Genus HAMANELLA Finger, 1983 Type-species (by original designation): Hamanella implexa Finger, 1983 Diagnosis: A genus of Trachyleberididae, subovate to subtrapezoidal in lateral view with uniformly coarse and extensive reticulum, anterior and posterior ends narrow and comparatively smooth; in dorsal view the carapace is bullet-shaped with short marginal extensions at anterior and posterior termina. Muscle scar pattern consisting of V-shaped frontal scar and vertical row of four ovate adductor scars ; dorsalmost adductor scar acutely angled posteriorly to those below it. Hingement holamphidont. Remarks: The inclination of the dorsalmost adductor scar suggests an affinity with the Rocaleberidini, from which Hamanella is otherwise morphologically distinct. Hamanella implexa Finger, 1983 1983 Hamanella implexa sp. nov. K. L. Finger, Micropaleontology , 29 (1), 94, pi. 8, figs. 1-9, pi. 10, fig. 3. Holotype: United States National Museum of Natural Flistory, Washington, coll. no. USNM 332113, ? car. [Paratypes: United States National Museum of Natural History coll. nos. USNM 332114, USNM 332115]. Explanation of Plate 11, 18 Figs. 1, 2, 2car. (holotype, USNM 332113, 700 pm long): fig. 1, ext. It. lat. ; fig. 2, ext. rt. lat. ; fig. 3, ?RV, int. lat. (paratype, USNM 332114, 670 pm long). Scale A (250 ^m; x70), figs. 1-3. Stereo-Atlas of Ostracod Shells 11, 19 Hamanella implexa (3 of 4) Lower Rincon Formation, Los Sauces Creek, Santa Barbara County, California, U.S.A. ; lat. 34° 22' N, long. 119° 25' W. Latest Zemorrian (Oligo-Miocen e), Hanzawaia crassisepta Zone; thin- bedded calcareous mudstones interpreted by Finger {Ibid.) as distal-fringe turbidites deposited at > 2000 m depth. United States National Museum coll. nos. USNM 332113 (holotype,? car.: PI. 11, 18, figs. 1, 2; PI. 11, 20, fig. 4), USNM 332114 (paratype, 9 RV: PI. 11, 18. fig. 3), USNM 332115 (paratype, d car.: PI. 11, 20, figs. 1-3). All from the type locality. As for the genus. Hamanella is presently regarded as a monotypic genus. The reversed valve overlap seen in all specimens obtained from Santa Barbara County is of questionable taxonomic value. I have examined an unnamed form from the Saucesian (lower Miocene) of adjacent Kern County that displays normal valve overlap and is otherwise not too dissimilar from H. implexa. Currently known only from the latest Zemorrian (Oligo- Miocene) of the Santa Barbara Embay - ment, California, U.S.A. Text-fig. 1 Composite drawing of//, implexa, ? RV, internal view (x 70). Explanation of Plate 11, 20 Figs. 1-3, d car. (paratype, USNM 332115, 840 pm long): fig. 1, ext. It. lat; fig. 2, ext. rt. lat.; fig. 3, ext. dors.; fig. 4,?car., ext. dors. (holotype, USNM 332113, 700 /xm long). Scale A (250 /xm; x 70), figs. 1-4. Type Locality: Figured specimens: Diagnosis: Remarks: Distribution : Stereo- Atlas of Ostracod Shells 11 (6) 21-24 (1984) Sagmatocythere paracercinata (1 of 4) 595.337.14 (118.22) (420 : 162.005.50) : 551.351 + 552.52 ON SAGMATOCYTHERE PARACERCINATA WHATLEY & MAYBURY sp. nov. by R. C. Whatley & C. Maybury ( University College of Wales, Aberystwyth) Sagmatocythere paracercinata sp. nov. Holotype: Type locality: Derivation of name: Figured specimens: Brit. Mus. (Nat. Hist.) no. OS 12116, $RV. [Paratypes: Brit. Mus. (Nat. Hist.) nos. OS 12117 - OS 12120]. Blue Clay, Sample No. 29, N W corner Vicarage Pit, St. Erth, Cornwall, England (Nat. Grid Ref. SW 556352); Upper Pliocene Latin, reflecting the close morphological and possible ancestral relationship of this species to Sagmatocythere cercinata (Bonaduce, Masoli & Pugliese, 1976) ( Pubbl . Staz. zool. Napoli, 40, 394-395, pi. 11, figs. 6-9) from the Recent of the Gulf of Aqaba, Red Sea. Brit. Mus. (Nat. Hist.) nos. OS 12116 (holotype, ?RV: PI. 11, 22, fig. 1), OS 12117 (crLY: PI. 11, 22, fig. 2), OS 12118 (cCRV: PI. 11, 22, fig. 3), OS 12120 (c/RV: PI. 11, 24, fig. 1, 3, 4), OS 12119 (cfLY: PI. 11, 24, fig. 2). All from the type locality and type horizon. Explanation of Plate 1 1, 22 Fig. 1,9RV, ext. lat. (holotype, OS 12116, 400 ^.m long); fig. 2,d'LV, ext. lat. (paratype, OS 12117, 450/xm long); fig. S.cfRV, ext. lat. (paratype, OS 12118, 440 pm long). Scale A (lOO^im; x 140), figs. 1-3. Stereo- Atlas of Ostracod Shells 11, 23 Sagmatocythere paracercinata (3 of 4) Diagnosis: Remarks: Distribution: A species of Sagmatocythere with a prominent alar process overhanging the ventral margin. Reticulae irregular in the alar region and immediately anterior of the eye spot. Free marginal areas flat and without reticulation. The close similarity between S. cercinata and the new species has already been indicated, the major differences being those of size and ornamentation; S. paracercinata is larger (5. cercinata , 360 pm long) and has relatively and absolutely smaller fossae. The Loxoconchidae are very diverse in the Upper Pliocene of N W France and St. Erth with 75 species/subspecies belonging to 1 2 genera. Sagmatocythere, represented by 18 species/subspecies is the most diverse genus of the family. S. paracercinata is abundant in the St. Erth beds (204 adult valves and 530 juvenile valves) but only one male right valve and two juvenile valves have been recovered from France. Upper Pliocene; St. Erth, Cornwall, England; Le Temple du Cerisier, S W of Rennes and from Borehole 1549 at Saint- Jean-la-Poterie, S W of Redon, N W France. Explanation of Plate 1 1, 24 Figs. 1, 3-4,cfRV (paratype OS 12120, 460 p,m long): fig. 1, int. lat.; fig. 3, ant. hinge element; fig. 4, post, hinge element. Fig. 2,d'LV, int. lat. (paratype, OS 12119, 470 pm long). Scale A (100 pm- x 140), figs. 1, 2; Scale B (20 pm- x 700), figs. 3, 4. Stereo-Atlas of Ostracod Shells 11 (7) 25-28 (1984) Sagmatocy there pseudomultifora (1 of 4) 595.337.14 (118.22) (420 : 162.005.50) : 551.351 + 552.52 ON SAGMATOCYTHERE PSEUDO MULTIFORA MAYBURY & WHATLEY sp. nov. by C. Maybury & R. C. Whatley (University College of Wales, Aberystwyth) Sagmatocythere pseudomultifora sp. nov. 71981 Loxoconcha sp.; H. Hagn, H. Malz & E. Martini, Geologica bav., 82, 270, pi. 2, figs. 1, 2. Holotype: Type locality: Derivation of name: Figured specimens: Brit. Mus. (Nat. Hist.) no. OS 12121, 9RV. [Paratypes: Brit. Mus. (Nat. Hist.) nos. OS 12122 -OS 12126]. Blue Clay, Sample No. 29, N W corner Vicarage Pit, St. Erth, Cornwall, England (Nat. Grid Ref. SW 556352); Upper Pliocene. Latin, reflecting its close similarity to Sagmatocythere multifora (Norman, 1865) (In: G.S. Brady, Nat. Hist. Trans. Northumberland & Durham, 1, 18-19, pi. 6, figs. 13-16). Brit. Mus. (Nat. Hist.) nos. OS 12121 (holotype, $RV: PI. 11, 26, fig. 1), OS 12122 (c/LV: PI. 11, 26, fig. 2), OS 12123 (d-RV: PI. 11, 26, fig. 3), OS 12124 (9LV: PI. 11, 28, fig. 1), OS 12125 (?car.: PI. 11, 28, fig. 2), OS 12126 (d'car.: PI. 11, 28, fig. 3). All from the type locality and horizon. Explanation of Plate 11, 26 Fig. 1,$RV, ext. lat. (holotype, OS 12121, 460 jam long); fig. 2,d'LV, ext. lat. (paratype, OS 12122, 510/un long); fig. S^RV, ext. lat. (paratype, OS 12123, 510 pm long). Scale A (100 /xm; x 125), figs. 1-3. Stereo- Atlas of Ostracod Shells 11, 27 Sagmatocythere pseudomultifora (3 of 4) Diagnosis: A species of Sagmatocythere with an inconspicuous alar process and strong reticulation, the fossae being particularly well-developed in the posterior and alar regions. A wide frill-like flange surrounds the free marginal areas which are strongly laterally compressed. Remarks: This species is probably synonymous with S. sp. (Hagn, Malz & Martini, 1981, op. cit.); but the small illustrations and lack of a formal description for this species render this possibility tenuous. S. pseudomultifora resembles S. multifora (Norman) in shape and gross morphology of ornament, but differs in size (Norman’s species is smaller, only 390-395 /xm long) and in its pos- session of a more regular reticulum and more strongly developed and acute alae. The two species constitute what are in the authors’ opinion more typical members of the genus than the type-species, 5. napoliana (Puri, 1963) (Athersuch, Stereo-Atlas of Ostracod Shells, 3 (21), 117-124, 1976). Distribution: S. pseudomultifora occurs abundantly in the Upper Pliocene St. Erth beds and is also present in the deposits of three localities in N W France of Redonian age: Le Bosq d’Aubigny, Le Bosq d’ Aubigny (Manche) and Le Temple du Cerisier. See J.-P. Margerel, Les Foraminiferes du Redonien. System- atique, Repartition stratigraphique , Paleoecologie, Nantes I, 1-207, 1968, for geographical and stratigraphical details of the French localities. The species probably also occurs in the Miocene of S Germany (Hagn, Malz & Martini, op. cit.). Explanation of Plate 1 1, 28 Fig. 1,9LV, int. lat. (paratype, OS 12124, 460 /xm long); fig. 2, ?car., ext. dors, (paratype, OS 12125, 470 /xm long); fig. 3, d'car., ext. dors, (paratype, OS 12126, 510/xm long). Scale A (100 /xm; x 125), figs. 1-3. Stereo-Atlas of Ostracod Shells 11 (8) 29-36 (1984) 595.337.14 (118.15) (430.1 : 161.006.51) : 551.351 + 552.51 Cytheridea muelleri muelleri (1 of 8) ON CYTHERIDEA (CYTHERIDEA) MUELLERI MUELLERI (V. MUNSTER) by Roseline H. Weiss (Geological Institute, University of Cologne, Germany) Genus CYTHERIDEA Bosquet, 1852 Subgenus CYTHERIDEA Bosquet, 1852 Type-species (by original designation): Cy there Mullerii v. Munster, 1830 Diagnosis: The typical subgenus of Cytheridea with a special type of hinge: left valve with terminal loculate sockets and a somewhat oblique median element, the anterior part of which is elevated while the posterior part is depressed with respect to the dorsal margin. Cytheridea (Cytheridea) muelleri muelleri (v. Munster, 1830) ? 1830 Cythere Mullerii Nob. v. Munster, Jb. Min. Geogn. Geol. Petref.-Kunde, 1, 63. 1838 Cytherina MiXlleri (v. Munster); F. A. Roemer, Neues Jb. Min. Geogn. Geol. Petref.-Kunde, 1838, 516, pi. 6, fig. 6. non 1852 Cytheridea MiXlleri (v. Munster); J. A. H. Bosquet, Mem. cour. mem. sav. etrang., 24 (1850-1851), 39, pi. 2, figs. 4a-f. ? 1894 Cytheridea MiXlleri (v. Munster; E. Lienenklaus, Z. dt. geol. Ges., 46, 220. non 1936 Cytheridea (Cytheridea) miXlleri (v. Munster); M. B. Stephenson, J. Paleont., 10, 699, pi. 94, figs. 1, 2, 7. 1952 Cytheridea miXlleri (v. Munster); F. Goerlich, Senckenbergiana , 33 (1/3), 188, figs. 6-12. 1955 Cytheridea miXlleri (v. Munster); H. J. Oertli & A. J. Key, Bull. Verein. schweiz. Petrol. -Geol. -Ing., 22(62), 21, pi. 1, figs. 15, 16. Explanation of Plate 11, 30 Fig. 1, $car., ext. dors. (GIK 932-1704, 830 /xm long): fig. 2,q'car., ext. dors. (GIK 932-1702, 800 /xm long). Length includes marginal spines. Scale A (100 /xm; x 113), figs. 1, 2. Stereo-Atlas of Ostracod Shells 11, 31 Cytheridea muelleri muelleri (3 of 8) 1956 Cytheridea mullerii (v. Munster); H. J. Oertli, Schweiz, palaeont., Abh., 74 (1), 36, pi. 2, figs. 39-41. ? 1958 Cytheridea mullerii (v. Munster); F. Goerlich, Fortschr. Geol. Rheinld. Westf, 1, 216. 1958 Cytheridea mullerii (v. Munster) ; E. Triebel, in : H. Freund (Ed .),Handbuch der Mikroskopie in der Technik, Frankfurt, II, 3, 122, fig. 23. non 1975 Cytheridea mullerii (v. Munster); M. Faupel, Gottinger Arb . Geol. Paldont., 17, 23, pi. 8, figs. 2a-b (= Cytheridea [C. ] pernota Oertli & Key, 1955). ? 1980a Cytheridea muelleri muelleri (v. Munster); H. Uffenorde, Neues Jb. Geol. Paldont. Mh., 119. 1981 Cytheridea (Cytheridea) muelleri (v. Munster); H. Uffenorde, Palaeontographica Abt. A, 172 (4-6), 137, pi. 1, figs. 5, 8, 1983b Cytheridea (Cytheridea) muelleri muelleri (v. Munster); R.H. Weiss, Palaeontographica, Abt. A, 182 (4-6), 89, pi. 19, figs. 1, 2, 4, 5, 8, pi. 20, figs. 1-8, pi. 21, figs. 1-6, text-fig. 11. Neotype: Type locality: Figured specimens: The Ostracoda from the collections of v. Munster are presumed lost. The neotype is housed at Forschungsinstitut Senckenberg, Frankfurt: X/e 1885, c/LV. [Paraneotypes: Forschungsinstitut Senckenberg, X/e 1886-1890]. Astrup near Osnabriick, Germany. Upper Oligocene. Geological Institute, University of Cologne, nos. 932-1008 (c^LV: PI. 11, 34, fig. 1), 932-1701 (c/car.: PI. 11, 32, fig. 1), 932-1702 ( tfcar.: PI. 11, 30, fig. 2), 932-1704 ( ?car.: PI. 11, 30, fig. 1), 932-1705 (c/RV.: PI. 11, 36, fig. 2), 932- 1706 (d/LV.: PI. 11, 32, fig. 2), 932-1707 ( ?RV; PI. 11,36, fig. 1), 932-1713 (rfRV: PI. 11, 34, fig. 2). All specimens were collected by Prof. E. K. Kempf in 1961 at a depth of 54.2-55.5 m from shaft Tonisberg near Krefeld, Germany (German Nat. Grid Ref.: R 34033, H 97555; long. 6°29'E, lat. 51°25' N); Upper Oligocene, Sphenolithus ciperoensis zone (NP25) according to Benedek & Muller ( Neues Jb. Geol. Paldont. Mh., 1974, 388); fine sand (grain size 0.2-0.06 mm = 92.5%) according to Kempf ( Niederrhein , 35, fig. 2, 1968) ; shallow marine (5-20 m water depth) according to Goerlich ( Fortschr . Geol. Rheinld. Westf., 1, 220, 1958). Explanation of Plate 11, 32 Fig. Ed' car., ext. vent. (GIK 932-1701, 825 /xm long); fig. 2,dXV, int. lat. (GIK 932-1706; 838 /xm long). Length includes marginal spines. Scale A (100/xm; x 113), figs. 1, 2. Cytheridea muelleri muelleri (2 of 8) Stereo- Atlas of Ostracod Shells 11, 30 Stereo- Atlas of Ostracod Shells 11, 32 Cytheridea muelleri muelleri (4 of 8) i Stereo- Atlas of Ostracod Shells 11, 33 Cytheridea muelleri muelleri (5 of 8) Sex N X L (yam) Min Max X H (yam) Min Max X L/H Min Max 99RV 3 771 763 775 421 413 425 1.832 1.794 1.879 d'd' RV 3 784 763 800 396 375 413 1.980 1.939 2.033 99 LV 13 773 738 800 426 413 438 1.815 1.771 1.848 rfrfLV 13 784 750 825 408 375 438 1.923 1.854 2.035 L (/am) W (yam) L/W Sex N X Min Max X Min Max X Min Max 9$ car. 16 774 738 813 355 338 388 2.179 2.138 2,257 d'd'car. 16 784 750 813 329 313 338 2.385 2.293 2.461 Table 1. Measurements on specimens (N = no. of specimens; x = mean; L = length without marginal spines; H = height; W = width); A = valves, B = carapaces. Diagnosis: Carapace ovate to subquadrangular in dorsal view; subtrapezoidal in lateral view, anterior end obliquely rounded, posterior end obliquely truncated, narrowly rounded posteroventrally. Dorsal margin straight to slightly convex; ventral margin straight (left valve) to very slightly concave in posterior half (right valve). Surface of the valves pitted. Along the free margin the pits are arranged in subparallel rows. Anterior end with marginal denticulations (right valve: up to 15; left valve: 7). Posterior end of the right valve provided with three marginal spines. Anteriorly and posteroventrally narrow vestibules are present. Explanation of Plate 11, 34 Fig. l,c/LV, ext. lat. (GIK 932-1008, 825 gm long); fig. 2,d'RV, ext. lat. (GIK 932-1713, 813 yam long). Length includes marginal spines. Both valves are from the same carapace. Scale A (100 yam; x 113), figs. 1, 2. Stereo-Atlas of Ostracod Shells 11, 35 Cytheridea muelleri muelleri (7 of 8) Remarks: Distribution : Acknowledgement: Sexual dimorphism pronounced, the males being lower and, in dorsal view, narrower than the females, but of the same length. Differences discussed in detail by Weiss (1983b, op. cit.). The hinge of each valve is divided into three elements. The terminal elements are dentate plates (right valve) or loculate sockets (left valve) ; the median element is oblique with respect to the dorsal margin and the left valve provided with different types of toothlets. Normal pores are moderately numerous, scattered and of the sieve - type. The sieve - plates are perforated (approx. 6-12 small pores along the diameter); the setal perforation is eccentric. Marginal pore - canals are numerous (approx. 40 anteriorly). They reach the shell surface distally of the flange in two parallel lines. Occasionally a lip is developed around the openings. Line of con- crescence and inner margin are slightly separated anteriorly and posteroventrally. The investigated specimens from Tonisberg agree absolutely with Goerlich’s material from the type-locality of Astrup. Many of the previous records described under this name by various authors (Reuss 1850, Bosquet 1852, Jones 1857, Speyer 1863, Stanceva 1962, Faupel 1975) repre- sent different species. Upper Oligocene: Astrup near Osnabriick, Germany (v. Munster 1830, Lienenklaus 1894, Stephenson 1936, Goerlich 1952, Oertli & Key 1955, Oertli 1956, Triebel 1958, op. cit.); Shaft Rossenray (Lower Rhine Basin), Germany (Goerlich 1958, op. cit.) ; Shaft Tonisberg (Lower Rhine Basin), Germany (Weiss 1983b, op. cit.); Niedersachsen (borings), Germany (Uffenorde 1980, 1981, op. cit.). Thanks are due to the Deutsche Forschungsgemeinschaft for providing the Cambridge Stereoscan 180. Explanation of Plate 11, 36 Fig. 1,$RV, int. lat. (GIK 932-1707, 800 (am long); fig. 2,d'RV, int. lat. (GIK 932-1705, 838 yam long). Length includes marginal spines. PI. 11, 32, fig. 2 and PI. 11, 36, fig. 2 represent both valves of a single carapace. Scale A (100 yam; x 113), figs. 1, 2. Stereo-Atlas of Ostracod Shells 11, 36 Cytheridea muelleri muelleri (8 of 8) Stereo-Atlas of Ostracod Shells 11 (9) 37-44 (1984) Cytheridea muelleri toenisbergensis (1 of 8) 595.337.14 (118.15) (430.1 : 161.006.51) : 551.351 + 552.51 ON CYTHERIDEA (CYTHERIDEA) MUELLERI TOENISBERGENSIS WEISS by Roseline H. Weiss ( Geological Institute, University of Cologne, Germany) Cytheridea (Cytheridea) muelleri toenisbergensis Weiss, 1983 1983b Cytheridea (Cytheridea) muelleri toenisbergensis subsp. nov. R. H. Weiss, Palaeontographica, Abt. A, 182 (4-6), 94, pi. 22, figs. 1-10, text-fig. 12. Holotype: Geological Institute, University of Cologne, Germany, GIK 932-1722, 9 car. [Paratypes: Geological Institute, University at Cologne, GIK 932: 1709-1712, 1714-1719, 1722-1727], Type locality: Shaft Tonisberg near Krefeld, Germany (German Nat. Grid Ref. : R 34033, H 97555 ; long. b° 29' E, lat. 51°25'N). Type horizon: Depth range 54.2-55.5 m; Upper Oligocene; Sphenolithus ciperoensis zone (NP25) according to Benedek & Muller (Neues Jb. Geol. Paldont., Mh., 1974, 388 §); fine sand (grain size 0.2-0.06 mm = 92.5%) according to Kempf (Niederrhein, 35, fig. 2, 1968); shallow marine (5-20 m water depth) according to Goerlich (Fortschr. Geol. Rheinld. Westf, 1, 220, 1958). Figured specimens: Geological Institute, University of Cologne, nos. 932-1709 ( 9RV: PI. 11, 44 fig. 1), 932-1714 ($LV: PI. 11, 42, fig. 1), 932-1716 (d-LV: PI. 11. 42, fig. 2), 932-117 ( dRV: PI. 11, 44, fig. 2), 932- 1718 (