A Stereo-Atlas of Ostracod Shells

edited by R. H. Bate, D. J. Horne, J. W. Neale,

and David J. Siveter

Volume 12, Part 1; 31st May 1985

Published by the British Micropalaeontological Society, London

Editors

Dr R.H. Bate, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking, Surrey GU23 7EF.

Dr D.J. Horne, Department of Geology, City of London Polytechnic, Walburgh House, Bigland Street,
London El 2NG.

Prof. J.W. Neale, Department of Geology, The University, Hull HU6 7RH.

Dr David J. Siveter, Department of Geology, The University, Leicester LEI 7RH.

Editorial Board

Dr G. Bonaduce, Stazione Zoologica, 80121 Napoli, Italy.

Dr J.-P. Colin, Esso Production Research - European, 213 Cours Victor Hugo, 33321 Begles,
France.

Dr P. De Deckker, Research School of Pacific Studies, Australian National University, PO Box 4,
Canberra ACT 2600, Australia.

Dr D. van Harten, Universiteit van Amsterdam, Geologisch Instituut, Nieuwe Prinsengracht 130,
Amsterdam, The Netherlands.

Dr I. Purper, Departamento de Paleontologia e Estratigrafia, UFRGS, 90 000 Porto Alegre RS, Brazil.
Dr R.E.L. Schallreuter, Universitat Hamburg, Geologisch-Palaontologisches Institut, Bundesstrasse 55,
D 2000 Hamburg 13, West Germany.

Officers of the British Micropalaeontological Society

Chairman Prof. B.M. Funnell, School of Environmental Sciences, University of East Anglia, Norwich
NR4 7TJ.

Secretary Dr P.P.E. Weaver, Institute of Oceanographic Sciences, Brook Road, Wormley, Godaiming,
Surrey GU8 5UB. Tel: 042-879 4141.

Treasurer Dr J.E. Whittaker, Department of Palaeontology, British Museum (Natural History),
Cromwell Road, London SW7 5BD. Tel: 01-589 6323.

Journal Editor Dr. L.M. Sheppard, SSI (U.K.) Limited, Tannery House., Tannery Lane, Send, Woking,
Surrey GU23 7EF.

Newsletter Editor Dr R.L. Austin, Department of Geology, The University, Southampton S09 5NH.
Tel: (0703) 559122/557941

Conodont Group Chairman Dr R.J. Aldridge, Geology Department, University of Nottingham,
Nottingham NG7 2RD.

Secretary Dr H.A. Armstrong, Department of Geology, The University, Newcastle-upon-Tyne NE1 7RU.
Tel: (0632) 328511.

Foraminifera Group Chairman Dr M.D. Brasier, Department of Geology, University of Hull, Hull HU6 7RX.
Secretary Dr J.V. Weston, SSI (UK) Ltd., Tannery House, Tannery Lane, Send, Woking GU23 7EF.
Tel: (0483) 223902.

Microplankton Group Chairman Dr G.C. Wilkinson, Britoil, 150 St. Vincent Street, Glasgow G2 5LJ.
Secretary Dr J.B. Riding, British Geological Survey, Keyworth, Nottingham NG12 5GT.

Ostracod Group Chairman Dr J. Athersuch, B.P. Research Centre, Chertsey Road, Sunbury-on-Thames,
Middlesex TW16 7LN.

Secretary Mr. I.P. Wilkinson, British Geological Survey, Nicker Hill, Keyworth, Nottingham NG12 5GG.
Tel: (06077) 6111.

Palynology Group Chairman Dr M.C. Boulter, Palynology Research Unit, N.E. London Polytechnic,
Romford Road, London E15 4LZ.

Secretary Mr N. Hooker, Britoil, 150 St. Vincent Street, Glasgow G2 5LJ. Tel: 041-204 2525.
Calcareous Nannofossil Group Chairman Mr E.M. Finch, BP Research Centre, Chertsey Road, Sunbury-
on-Thames, Middlesex TW16 7LN.

Secretary Miss H. Stowe, Micropalaeontology Unit, University College, Gower Street, London WC1E 6BT.
Tel: 01-387 7050.

Instructions to Authors

Contributions illustrated by scanning electron micrographs of Ostracoda in stereo-pairs are invited.
Format should follow the style set by the majority of papers in this issue. Descriptive matter apart
from illustrations should be cut to a minimum; preferably each plate should be accompanied by one
page of text only. Blanks to aid in mounting figures for plates may be obtained from any one of the
Editors or Editorial Board. Completed papers should be sent to Dr David J. Siveter.

The front cover shows a right valve (927 /xm long) of Newnhamia petiola De Deckker, 1979, in life position;
from the type locality, Pine Tree Creek Lagoon, near Hughenden, Queensland, Australia. This species
swims upside down and can stick to the water surface tension with its flat ventral area.

Stereo-Atlas of Ostraeod Shells 12 (1) 1-6 (1985) Eucythere declivis (1 of 6)

595.337.14 (119.9) (420 : 162.005.50 + 261.26 : 161.000.57 + 420 : 162.002.54) : 551.351

ON EUCYTHERE DECLIVIS (NORMAN)

by David J. Horne & John E. Whittaker
(City of London Polytechnic & British Museum ( Natural History), London)

Genus EUCYTHERE Brady, 1868

1866 Cytheropsis gen. nov. G. O. Sars, Forh. VidenskSelsk. Krist. , 1865, 57-58 (= junior homonym of Cytheropsis M’Coy, 1849).

1868 Eucythere nom. nov. G. S. Brady, Trans. Linn. Soc. Lond., 26, 429.

Type-species: Cythere declivis Norman, 1865 (subsequent designation by Brady & Norman, Scient. Trans. R. Dubl. Soc.,

ser. 2, 4, 178, 1889)

Diagnosis: Carapace sub-triangular in lateral view with greatest height at or in front of mid-length. Anterior

margin broadly rounded, dorsal and ventral margins strongly convergent posteriorly. Greatest width
at or behind mid-length. External surface smooth or weakly ornamented. Inner lamella broad
anteriorly, narrow ventrally and posteriorly. Anterior and posterior vestibula present. Marginal
pore-canals straight, usually 10-15 anteriorly and about 3 posteriorly. Hinge lophodont, running
from about mid-length to the posterior margin. Four adductor muscle-scars in an arcuate row, the
lowermost being relatively large and crescent-shaped. Frontal scar relatively large, asymmetrically
U- or V-shaped. Prominent fulcral point. Normal pores conspicuous, sieve-type. Dimorphic, male
more elongate than female. Antennula with five articulated podomeres. Maxillular palp and
masticatory processes slender, innermost one much reduced. Legs slender. Male brush-shaped
organ relatively large, spatulate, with numerous distal setae. Male copulatory appendage relatively

small.

Explanation of Plate 12, 2

Fig. 1 , 6 RV, ext. lat. (paralectotype, 1984. 194, 590^m long) ; fig. 2, 9 R V, ext. lat. (lectotype, 1984. 193, 620/^m long) ; fig. 3, 9 LV, int.
lat. (1.13.33, 610/xm long).

Scale A (100^m; x95), figs. 1-3.

Stereo- Atlas of Ostraeod Shells 12, 3 Eucythere declivis (3 of 6)

Eucythere declivis (Norman, 1865)

Cythere declivis sp. nov. A. M. Norman, in : G. S. Brady (Ed.), Nat. Hist. Trans. Northumberland and Durham , 1 (1865-
1867), (1), 16-17, pi. 5, figs. 9-12.

Cythere declivis , Norman (n.sp.); A. M. Norman, in : G. S. Brady, Rept. Brit. Assoc. Adv. Sci., 1864 : 192 (identical description
to above but without illustrations).

Cytheropsis tenuitesta sp. nov. G. O. Sars, Forh. VidenskSelsk. Krist., 1865, 59.

Eucythere declivis (Norman); G. S. Brady, Trans. Linn. Soc. Lond. , 26, 430-431 (pars), pi. 27, figs. 22-26, 54-55 only (non
pi. 25, figs. 49-50, pi. 27, figs. 52-53).

Eucythere declivis (Norman); G. W. Muller, Fauna Flora Golf. Neapel., 21, 363, pi. 29, figs. 5, 13.

Eucythere declivis (Norman); G. O. Sars, An account of the Crustacea of Norway , 9,Ostracoda, pts. 9, 10, 163-164, pi. 75,
fig. 2, Bergen Museum.

Eucythere declivis (Norman) ; C. W. Wagner, Sur les Ostracodes du Quaternaire recent des Pays-Bas et leur utilisation dans
T etude geologiques des depots holocenes, 43-44 (pars), pi. 15, fig. 5 only (non figs. 1-4), Mouton & Co., The Hague.
Eucythere declivis (Norman); A. Rosenfeld, Meyniana, 29, 19-20, pi. 3, fig. 42.

Lectotype: Brit. Mus. (Nat. Hist.) no. 1984.193, $ RV + LV.

[Paralectotype: Brit. Mus. (Nat. Hist.) no. 1984.194, <3 RV + LV],

Type locality: Plymouth, SW England (approx, lat. 50°23'N, long. 04° 09' W); Recent.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1984.194 (paralectotype, 6 RV : PI. 12, 2, fig. 1), 1984.193 (lectotype,

9 RV: PI. 12, 2, fig. 2, Text-fig. la), 1984.195 (9 car. + appendages; appendages: Text-figs, lb-c),
1984.196 (c? car. + appendages; LV: PI. 12, 4, fig. 1; RV: PI. 12, 4, fig. 3; appendages: Text-figs,
lg-h). Hancock Mus., University of Newcastle, no. 1.13.33 (9 car. + appendages; LV: PI. 12, 2,
fig. 3, PI. 12, 4, fig. 2; RV: PI. 12, 4, fig. 4; appendages: Text-figs. ld-f).

The lectotype and paralectotype were taken from slide no. 1911.11.8 M 3496 in the Norman
collection, labelled “ Eucythere declivis (Norman) Types, Plymouth, Mr Barlee”. Nos. 1984. 195 and

Explanation of Plate 12, 4

Figs. 1 , 3, 6 ( 1984. 196, 560/um long); fig. 1, LV, ext. lat. ; fig. 3, RV, dors. ; figs. 2, 4, 9 (1.13.33, 610/um long): fig. 2, LV, ext. lat. ; fig. 4,
RV, dors.

Scale A (lOO^um; x 95), figs. 1-4.

1865

1865

71866

1868

non 1894
71925

1957

1977

Stereo-Atlas of Ostracod Shells 12, 2

Eucythere declivis (2 of 6)

Stereo-Atlas of Ostracod Shells 12, 5 Eucythere declivis (5 of 6)

Figured specimens: 1984.196, both from the North Sea off Aberdeen (approx, lat. 57° N, long. 00°) were provided by

(contd.) J. E. Robinson. Hancock Mus. no. 1.13.33, from 8 miles off the Durham Coast between Seaham and
Sunderland (approx, lat. 54°55' N, long. 01° 10' W), depth 20-30 fath. (36-55m), was taken from
slide no. 2.11.36 in the Brady collection.

Diagnosis: Carapace finely pitted in dorsal and median areas, smooth or with faint concentric ribbing in anterior

and ventral areas. Posteroventral marginal area strongly compressed in male, weakly so in female.
Anterior vestibulum relatively broad. Distal process of male copulatory appendage relatively large,
semicircular.

Remarks: In his “Monograph of the Recent British Ostracoda” Brady (1868, op. cit. ) recognised two forms in

addition to the trueE. declivis. One of these he described as E. anglica sp. nov. in the appendix of the
monograph, while the other was later described by Brady & Robertson (Ann. Mag. nat. Hist., ser. 4,
3, 370-371, pi. 21, figs. 12-14, 1869) as£. declivis var .prava ; both taxa have been largely ignored
by subsequent authors. Our examination of their respective type specimens in the Hancock Museum
has shown conclusively that they are both valid species (see Horne & Whittaker, Stereo-Atlas of
Ostracod Shells 12, 7-10 and 12, 1 1-14, 1985).£\ anglica is less elongate and more strongly pitted
thanE. declivis, while E. prava is easily distinguished by its posteromedian sulcus. The type slides of
E. declivis in the Norman collection contain specimens of both E. declivis and E. prava. We have
chosen as a lectotype and paralectotype two specimens which correspond closely to the original
description and illustrations of E. declivis.

The Mediterranean records oiE. declivis of G. W. Muller (1894, op. cit.) and most subsequent
authors may be referred to Eucythere curta Ruggieri, 1975 (Revta esp. Micropaleont. , 6, 433-434,
fig. 6 \q.v. for full synonymy). As far as we can ascertain E. declivis does not live in the Mediterranean.

The form illustrated by Sars (1925, op. cit.) appears to lack the posteroventral compressed
area exhibited by British specimens and also differs in minor details of carapace outline and the
male copulatory appendage; we are therefore in some doubt as to whether it is conspecific with
E. declivis. Of Wagner’s (1957, op. cit. ) illustrations of E. declivis, only one can be confidently
assigned to that species, while the others should be referred to E. argus (Sars, 1866).

Distribution: Records of E. declivis require careful re-examination in the light of the above-mentioned confusion

with other species. It appears to be fairly common in marine sublittoral waters around Britain and in
the North Sea. Rosenfeld (1977, op. cit.) recorded it in the Baltic Sea in salinities of 25-30%o.

Stereo-Atlas of Ostracod Shells 12, 6 Eucythere declivis (6 of 6)

Text-fig. 1. Eucythere declivis. a: 9 RV seen in transmitted light (lectotype, 1984.193). b-h: appendages; b, c, $ (1984.195), b: antennula.
c: antenna; d-f, 9 (1.13.33), d: maxillula, e: first leg, f: second leg; g, h, 6 (1984.196), g: third leg, h: copulatory appendage.

Stereo-Atlas of Ostracod Shells 12 (2) 7-10 (1985)
595.337.14 (119.9) (420 : 162.002.54) : 551.351

Eucythere anglica (1 of 4)

ON EUCYTHERE ANGLICA BRADY

1868

1868

Type locality:
Figured specimens:

by David J. Horne & John E. Whittaker
(City of London Polytechnic & British Museum (Natural History), London)

Eucythere anglica Brady, 1868

Eucythere declivis (Norman); G. S. Brady, Trans. Linn. Soc.Lond., 26, (pars), 431, pi. 25, figs. 49, 50 only ( non 430, pi. 27, figs.
22-26, 52-55) (non Cythere declivis Norman, 1865).

Eucythere anglica sp.nov. G. S. Brady, Trans. Linn. Soc.Lond., 26, 475, pi. 25, figs. 49, 50.

Lectotype: Hancock Museum, University of Newcastle, no. 1.54.20, 9 RV.

[Paralectotype: Hancock Museum no. 1.54.21, 6 RV],

Off Seaham, Durham coast, NE England (approx, lat. 54° 50'N, long. 01° 10'W); Recent.
Hancock Mus. nos. 1.54.20 (lectotype, 9 RV: PI. 12, 8, fig. 2), 1.54.21 (paralectotype, 6 RV: PI. 12,
8, fig. 1, PI. 12, 10, fig. 3), 1.54.22 (9 car. + appendages; LV: PI. 12, 8, fig. 3, PI. 12, 10, fig. 1,
Text-fig. lb; RV: PI. 12, 10, figs. 2, 4, Text-fig. la; appendages: Text-fig. lc-d). Nos. 1.54.20 and
1.54.21 were taken from slide no. 1.02.35 in the Brady collection, labelled “ Eucythere anglica, off
Seaham”. 1.54.22 is from off Hartlepool, NE England (approx, lat. 54° 41 'N, long. 01° 08' W) and
was taken from slide no. 2.13.04 in the Brady collection.

Carapace small (< 500pm long), conspicuously pitted in median and dorso-median areas, with
concentric ribbing in the anterior and ventral marginal areas. Posteroventral corner somewhat
compressed. Anterior vestibulum relatively broad.

Brady (1868, op. eft.) initially mentioned and illustrated this species as a form of E. declivis, but then
described it as a new species in the appendix of the same publication. It is smaller, less elongate and
more strongly pitted than E. declivis (see Horne & Whittaker, Stereo-Atlas of Ostracod Shells 12,
1-6, 1985).

Explanation of Plate 12, 8

Fig. 1 , 6 RV, ext. lat. (paralectotype, 1.54.21, 460 pm long); fig. 2, 9 RV, ext. lat. (lectotype, 1.54.20, 480pm long); fig. 3, 9 LV, ext. lat.
(1.54.22, 480pm long). Scale A (100pm; x 125), figs. 1-3.

Diagnosis:

Remarks:

Stereo- Atlas of Ostracod Shells 12, 9 Eucythere anglica (3 of 4)

Distribution: The Brady collection contains several examples of this species from marine sublittoral localities

around the British Isles. The absence of more recent records is probably due to confusion with

Explanation of Plate 12, 10

Figs. 1 , 2, 4, 9 ( 1.54.22, 480pm long): fig. 1 , LV int .lat. ; fig. 2, RV int.lat. ; fig. 4, RV dors.; fig. 3, <3 RV, dors, (paralectotype, 1.54.21,
460pm long). Scale A (100pm; x 125), figs. 1-4.

Stereo-Atlas of Ostracod Shells 1 2, 1 0

Stereo-Atlas of Ostracod Shells 12 (3) 11-14 (1985) Eucythere prava (1 of 4)

595.337.14 (119.9) (415 : 162.010.53 + 420 : 162.007.49 + 420 : 162.005.50) : 551.351

ON EUCYTHERE PRAVA BRADY & ROBERTSON

by David J. Horne & John E. Whittaker
(City of London Polytechnic & British Museum ( Natural History), London)

Eucythere prava Brady & Robertson, 1869

1868 Eucythere declivis (Norman); G. S. Brady, Trans .Linn .Soc .Lond . , 26, 430 (pars), pi. 27, figs. 52-53 only (non pi. 27, figs. 22-26,
54-55, pi. 25, figs. 49-50) ( non Cythere declivis Norman, 1865).

1869 Eucythere declivis var .prava, G. S. Brady & D. Robertson , Ann. Mag. nat. Hist., (ser. 4), 3, 370-371, pi. 21, figs. 12-14.

Lectotype:
Type locality:
Figured specimens:

Diagnosis:

Remarks:

Hancock Museum, University of Newcastle, no. 1.39.33, 9 carapace.

Westport Bay (= Clew Bay), Co. Mayo, W Ireland (approx, lat. 53° 50'N, long. 09° 40' W); Recent.
Hancock Mus. nos. 1.39.33 (lectotype, $ car.: PI. 12, 12, fig. 1), 1.39.34 (d car + appendages; LV:
PI. 12, 12, fig. 3, Text-fig. la; RV: PI. 12, 14, fig. 2; copulatory appendages; Text-fig. lc-d), 1.39.35
(c? car. -I- appendages; appendages: Text-fig. le-f). Brit. Mus. (Nat. Hist.) no. 1984.187 (9 car.; LV:
PI. 12, 12, fig. 2, PI. 12, 14, figs. 1, 4, Text-fig. lb; RV: PI. 12, 14, fig. 3). The lectotype was taken
from slide no. 2.03.16 in the Brady collection. Hancock Mus. nos. 1.39.34 and 1.39.35, from off
St. Mary’s, Isles of Scilly (approx, lat. 49° 55'N, long. 06° 15'W), depth 20 fath. (36m), were taken
from slide no. 2. 12.40 in the Brady collection. Brit. Mus. (Nat. Hist.) no. 1984. 187, from the English
Channel, S of the Eddystone Lighthouse (approx, lat. 50° 02 'N, long. 04° 22' W), depth approx.
75 m, was provided by S. Sturrock.

Posterior half of carapace with an irregular longitudinal posteromedian sulcus, anterior half faintly
reticulate. Surface smooth or very finely pitted. Anterior vestibulum relatively narrow. Male
copulatory appendage with a lemon-shaped distal process.

Brady (1868, op.cit.) originally mentioned and illustrated this species as a form of E. declivis
(Norman, 1865) (see Horne & Whittaker, Stereo-Atlas of Ostracod Shells 12, 1-6, 1985); it was later

Explanation of Plate 12, 12

Fig. 1, $ car., rt.lat. (lectotype, 1.39.33, 510/um long); fig. 2, 9 LV, ext. lat. (1984.187, 500/um long); fig. 3, 3 LV, ext. lat. (1.39.34,
480/u.m long). Scale A (100/u.m; x 125), figs. 1-3.

Stereo- Atlas of Ostracod Shells 12, 13 Eucythere prava (3 of 4)

Remarks (contd.): formally described as a variety of E. declivis by Brady & Robertson (1869, op. cit.) and is here raised
to specific status. E. prava is easily distinguished from other Eucythere species by its characteristic
postero-median sulcus.

Distribution: E. prava appears to be fairly widespread in marine sublittoral waters around the British Isles (Brady
and Norman collections and herein).

Text-fig. 1. Eucythere prava. a-b: valves seen in transmitted light; a: 3 LV (1.39.34); b: 9 LV (1984.187). c-f: appendages; c, d: 3
copulatory appendages (1.39.34); e, f: 3 antennula and antenna (1.39.35).

Explanation of Plate 12, 14

Figs. 1,3,4, 9 (1984.187, 500/um long): fig. 1 : LV, ext. lat.; fig. 3, RV, dors.; fig. 4, LV, ext. lat., detail of median area showing external
trace of central muscle-scar field; fig. 2, 3 RV, dors. (1.39.34, 480/am long).

Scale A (100/xm; x 125), figs. 1-3; scale B (50/am; x 350), fig. 4.

Stereo-Atlas of Ostraeod Shells 12, 12

lacy there prava (2 of 4)

Stereo-Atlas of Ostraeod Shells 12, 14 Eucythere prava (4 of 4)

Stereo-Atlas of Ostracod Shells 12 (4) 15-18 (1985) Piretia commasulcata (1 of 4)

595.336.13 (1 13.313) (430.1 : 161.008.54) : 551.35 + 552.55

ON PIRETIA COMMASULCATA SCHALLREUTER sp. nov.

by Roger E. L. Schallreuter
( University of Hamburg, German Federal Republic)

Piretia commasulcata sp. nov.

Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH), no. 2938;
9 LV, anterodorsally and posterodorsally incomplete.

[Paratypes: GPIMH nos. 2939-2942],

Upper Harjuan (upper Ordovician) Ojlemyrflint erratic boulder no. Sy60 of the upper Kaolinsand
(lower Pleistocene) from near Braderup, Isle of Sylt (N Frisian Is, N Sea), West Germany; lat. 54°
56'N, long. 8° 21'E.

Alluding to the comma-shaped sulcus.

GPIMH nos. 2938 (holotype, incomplete 9 LV: PI. 12, 16, figs. 1, 3), 2939 (paratype, juv. LV:
PI. 12, 16, fig. 2, PI. 12, 18, fig. 1), and 2940 (paratype, nearly complete juv. RV: PI. 12, 1 8, fig. 2).
All specimens are from the type locality; boulder collected by Ulrich von Hacht (Hamburg) in 1978.

Holotype:

Type locality:

Derivation of name:
Figured specimens:

Explanation of Plate 12, 16

Figs. 1,3, anterodorsally and posteriorly incomplete 9 LV (holotype, GPIMH 2938, 683 /am long): fig. 1, ext. lat.; fig. 3, ext. anterovent.

Fig. 2, juv. LV, ext. vent, (paratype, GPIMH 2939, 585/um long).

Scale A (100/am; x 110), figs. 1, 3; scale B (100/am; x 170), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 17 Piretia commasulcata (3 of 4)

Diagnosis:

Remarks:

Distribution:

Species of Piretia with short, comma-shaped sulcus near the dorsal margin. Preadductorial node
bulb-like but flattish. Shape (‘Gestalt’) rather long to long; shape of domicilium very long. A narrow,
low, ridge-like plica occurs along the dorsal border. Velum forms a ridge or narrow flange which
terminates posteroventrally in a short spine. Female has a very strongly convex dolon forming a false
brood pouch. A ridge occurs along the border of lateral and ventral surfaces of the pouch. Lateral
surface reticulate. Adults about 0.8 mm long.

P. commasulcata is distinguished from congeneric species mainly by its elongate shape and its
diagnostic short, comma-shaped sulcus. Piretia erinacea Schallreuter, 1964 (lower Upper Viruan
Backsteinkalk erratic boulders of Northern Germany) differs further in having a row of short spines
instead of a ridge at the border of the lateral and ventral surfaces of the female dolon, its tecno-
morphic velum is developed as a spinose ridge or a row of spines and its surface is spinose and
granulose (Schallreuter, Palaeontographica (A), 144, pi. 18, fig. 1, pi. 17, figs. 5-7, 1973).

Piretia reticulata Qvale, 1980 ( Norsk Geol. Tidsskr., 60, 94), from the Caradoc Series of the
Oslo region, resembles P. commasulcata in having reticulation and a velar flange with an abrupt
posterior termination but is distinguished by its much better developed sulcus, its centrodorsal spine
and by its tecnomorphic velum which consists of a row of spines recalling that ofP. erinacea (Qvale,
op. cit., figs. 2, 3).

Known from the type locality and from Ojlemyrflint erratic boulders of the Isle of Gotland, Baltic
Sea (boulder no. G30 of Schallreuter collection).

Explanation of Plate 12, 18

Fig. 1 , juv. LV, ext. lat. (paratype, GPIMH 2939); fig. 2, slightly incomplete juv. RV, ext. lat. (paratype, GPIMH 2940, 689/um long).
Scale A (100/um; x 170), fig. 1; scale B (100 /urn; x 140), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 18

Piretia commasulcata (4 of 4)

Stereo-Atlas of Ostracod Shells 12 (5) 19-22 (1985) Kroemmelbeinia valensis (1 of 4)

595.337.2 (113.313) (485 : 161.018.57) : 551.35 + 552.55

ON KROEMMELBEINIA VALENSIS SCHALLREUTER sp. nov.

by Roger E. L. Schallreuter
( University of Hamburg, German Federal Republic)

Kroemmelbeinia valensis sp. nov.

Geologisch-Palaontologisches Institut und Museum, University of Hamburg (GPIMH) no. 2943;
a carapace.

[Paratypes: nos. 2944-2947],

Upper Harjuan (upper Ordovician) Ojlemyrflint erratic boulder no. G287 from the beach at Vale,
NE Gotland (Baltic Sea), Sweden; lat. 57°48'N, long. 18°26'E.

After the type locality, Vale, Gotland.

GPIMH nos. 2944 (paratype, LV : PI. 12, 20, fig. 1), 2945 (paratype, RV ; PI. 12, 20, fig. 2), 2943
(holotype, car.: PI. 12, 22, fig. 1) and 2946 (paratype, LV; PI. 12, 22, fig. 2). All specimens are from
the type locality; boulder collected by the author in 1976.

Holotype:

Type locality:

Derivation of name:
Figured specimens:

Explanation of Plate 12, 20

Fig. 1, LV, ext. lat. (paratype, GPIMH 2944, 1505/u.m long); fig. 2, RV, ext. lat. (paratype, GPIMH 2945, 1805/xm long).
Scale A (250/u.m ; x 67), fig. 1 ; scale B (250/u.m ; x 55), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 21 Kroemmelbeinia valensis (3 of 4)

Diagnosis:

Remarks:

Distribution:

Species of Kroemmelbeinia with the ‘anteroventral corner’ situated near the longitudinal middle line
and only weakly pointed. Posteroventral corner pointed but not spinose. Adults up to 2.00mm long.
The middle Ordovician type -species, Kroemmelbeinia ala Schallreuter ( Geologie 18, 211, 1969)
(= Beecherellita ordovica Neckaja, 1973; Schallreuter, Geol. For Stockh . Forh. 97, 387, 1975), has
the same shape (length : height ratio) but is much smaller (adults 0.82mm long). Moreover, the
anteroventral corner in K. ala is more pointed and lies more ventrally than in K. valensis, and its
posteroventral corner forms a short spine whereas in K. valensis it is only pointed.

K. valensis clearly displays two stop-pegs in the larger valve (PI. 12, 22, fig. 2). In the last
decade this feature has proved to be very important in the higher taxonomy of Ordovician
podocopes. It is characteristic for Ordovician metacopes (e.g., see Stereo-Atlas Ostracod Shells 5,
56, fig. 1 ; 7, 76, fig. 2; 7, 80, figs. 1-2; Schallreuter, Proc. VII Internat. Symp. Ostracodes Belgrade
1979, pi. 1, figs. 2, 6, 8, 9, pi. 2, fig. 5), and is taxonomically more important than the general outline
and other features. Thus, it has helped unmask excellent examples of homeomorphy in ostracodes ;
for example, Kroemmelbeinia spina Schallreuter, 1969 has a distinct inner lamella but lacks stop-
pegs and was therefore later placed in Spinobairdia. A second example involves ‘Platyrhomboides’
minimus (with broad inner lamella) and ‘P.’ breviclaustrum (with two stop-pegs) (Schallreuter,
1979, op. cit., pi. 1, figs. 7, 8 and pi. 2, figs. 3, 4).

Known only from the type locality; upper Ordovician.

Explanation of Plate 12, 22

Fig. 1, car., ext. rt. lat. (holotype, GPIMH 2943, 1765yu.m long) ; fig. 2, LV, int. lat., slightly obi., showing stop-pegs (paratype, GPIMH
2946, 2000m long).

Scale A (250p,m; x 59), fig. 1 ; scale B (250/u.m; x 51), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 20

Krocmmelbciiuu valensis (2 of 4)

Stereo-Atlas of Ostracod Shells 12, 22

Kroemmclbeiniu valensis (4 of 4)

.........

Stereo- Atlas of Ostracod Shells 12 (6) 23-30 (1985) Glyptocythere raasayensis (1 of 8)

595.337.14 (116.222) (411.162.007.57) : 551.35

ON GLYPTOCYTHERE RAASAYENSIS STEVENS sp. nov.

by Graham W. Stevens
(University of Hull, England)

Glyptocythere raasayensis sp. nov.

University of Hull no. HU.212.J. 1, a 9 carapace.

[Paratypes: nos. HU.212. J.2-6].

Stream section at Brae (Nat. Grid. Ref.: NG 515418), Isle of Raasay, Scotland. Garantiana Clay,
garantiana Zone, Bajocian, Jurassic.

From the type locality, Isle of Raasay.

University of Hull, nos. HU.212. J.l (holotype, 9 car.: PI. 12, 24, fig. 1, PI. 12, 26, fig. 1), HU.212. J. 2
(c? RV: PI. 12, 24, fig. 2, PI. 12, 28, fig. 1 ; specimen broken subsequent to photography), HU.212. J. 3
(6 car.: PI. 12, 26, fig. 2), HU.212. J.4 (9 LV: PI. 12, 28, fig. 2, PI. 12, 30, fig. 2), HU.212. J.5 (<J LV:
PI. 12, 30, fig. 1). All the specimens are from the type locality and horizon.

A species of Glyptocythere in which the carapace shows well developed reticulation formed by fine
anastomosing ribs over most of the lateral surface. The postero-dorsal, dorsal and antero-dorsal
marginal areas are smooth and there are prominent pore conuli in the postero-dorsal area. Five
prominent longitudinal ribs are developed ventrally. Sexual dimorphism distinct, the presumed
females being proportionally higher and wider than the males. Hinge and muscle scar pattern typical
of the genus.

Explanation of Plate 12, 24

Fig. 1, 9 car., rt.lat. (holotype, HU.212.J.1, 750/im long); fig. 2, d RV, ext.lat. (HU.212. J.2, 700yu.m long).
Scale A (100/u.m; x 120), fig. 1; scale B (lOO^im; x 130), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 25 Glyptocythere raasayensis (3 of 8)

Remarks: Whereas the early Bajocian and Bathonian ostracod faunas of Britain are well known, there is a

dearth of information on higher Bajocian faunas and so a lack of material with which the present
species can be compared. However, on the continent the upper Bajocian Glyptocythere species are
well known; the type-species, G .tuber odentina Brand & Malz, 1962 ( Senckenberg.leth ., 43,
433-435), comes from the Obere Parkinsonienschichten of NW Germany which lies above the
garantiana Zone. There are clearly similarities between the present species and the type-species in
the general pattern of ornamentation but in G. raasayensis the reticulation lacks the accentuation of
the vertical element seen in G .tuber odentina. General shape is similar in the females, but the male
G .raasayensis has a narrower caudal projection. The Scottish species also has five longitudinal
ventral ribs compared with three or four in the type-species. G.regulariformis Brand & Malz, 1962
from the Garantianenschichten in Germany, of comparable age to the Scottish deposit, is much
closer to the present species in the form of ornamentation but has a much more convex ventral
margin. There is nothing else comparable from the garantiana Zone. The slightly earlier
G.praecursor Brand & Malz, 1966 from the Subfurcatenschichten shows similar but coarser
ornamentation and a more convex ventral margin. Other species are less close. Among the earlier
British species G.scitula Bate, 1 965 from the humphresianum Zone is closer than any of the German
species and G. raasayensis could well be a later derivative. The latter differs most obviously in the
better-defined postero-dorsal angle and the straight or slightly convex dorsal margin.

Explanation of Plate 12, 26

Fig. 1, $ car., dors, (holotype, HU. 212. J.l, 750/a.m long); fig. 2, d car., dors. (HU.212.J.3, 740/u.m long).
Scale A (100/i.m; x 130), figs. 1, 2.

Holotype:

Type locality:

Derivation of name:
Figured specimens:

Diagnosis:

’I'T""!

i.-...;

Stereo-Atlas of Ostracod Shells 12, 24

(ilypiocyihcrc rcuisaxcnsis (2 of 8)

Stereo-Atlas of Ostracod Shells 12, 27

Glyptocythere raasuyensis (5 of 8)

Remarks (contd.): The type locality, near Storab’s Grave, is regarded by Morton (Scott .J .Geol .. 12, 24, 1 976) as

the best in the Hebrides. Here the Garantiana Clay is approximately 2 m thick and lies directly
above the Bearreraig Sandstone Formation. It consists of poorly bedded, structureless medium-grey
clay which is highly fossiliferous and contains the ammonites Garantiana (G.) filicosta and Garantiana
(G.) baculata as well as a large number of small bivalves including prolific A vicula. The microfauna
is restricted to three foraminiferal species (Ammobaculites agglutinans, Frondicularia oolithica and
Lenticulina limbata) and this single ostracod species. The ammonites and all the other fauna suggest
normal marine salinities, but the pyritisation of the ammonites, the small size (less than 4 mm) of the
bivalves together with their pyrite skin and the restricted nature of the microfauna suggest generally
inimical conditions and perhaps somewhat deoxygenated bottom waters. The general aspect of the
fauna suggests a shallow shelf sea with a depth of about 20m.

Distribution: So far known only from the type locality.

Explanation of Plate 12, 28

Fig. 1, 6 RV, int.lat. (HU.212.J.2, 700/am long); fig. 2, 9 LV, int.lat. (HU.212.J.4. 660/um long).
Scale A (100/xm; x 140), figs. 1, 2.

Stereo-Atlas of Ostracod Shells 12, 29 Glyptocythere raasayensis (7 of 8)

Text-fig. 1 . Simplified geological map of central and southern Raasay showing the outcrop of the Garantiana Clay and the type locality at
Brae.

Text-fig. 2. Muscle scar pattern of G. raaysayensis.

Explanation of Plate 12, 30

Fig. 1, 6 LV, ext.lat. (HU.212. J.5, 575/u.m long); fig. 2, 9 LV, ext.lat. (HU.212.J.4, 660/u.m long).
Scale A (100 /am; x 160), fig. 1; scale B (100/um; x 150), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 28

Glyptoey there raasayensis (ft of 8)

Glyptocy there raasayensis (8 of 8)

Stereo-Atlas of Ostracod Shells 12, 30

Stereo-Atlas of Ostracod Shells 12 (7) 31-38 (1985) Limnocythere shixiaensis (1 of 8)

595.337.14 (119.3) (510 : 161.114.40) : 552.52

ON LIMNOCYTHERE SHIXIAENSIS (WANG)

by Wang Qiang

(Tianjin Institute of Geology and Mineral Resources, China)

Limnocythere shixiaensis (Wang, 1980)

1980 Leucocythere shixiaensis sp.nov. Q.Wang, Bull. Chinese Acad. geol.Sci., ser. 6, 1 (2), 127-134, pi. 1, figs. 1-5, text-fig. 2.

Neotype:
Type locality:

Figured specimens:

Diagnosis:

Tianjin Institute of Geology and Mineral Resources no. os.84.7-1, a 9 carapace.

Basal portion of the Xiashagou section, Yangyuan County, Hebei Province, China. Nihewan
Formation, Lower Pleistocene.

Tianjin Inst. Geol. & Min. Res. nos. os.84.7-1 (neotype, 9 car.: PI. 12, 32, figs. 1-4), os.84.7-2
(plesiotype, 9 car.: PI. 12, 34, figs. 1-4), os.84.7-3 (plesiotype, 9 car.: PI. 12, 36, figs. 1-4), os.84.7-4
(plesiotype, 9 juv. car.: PI. 12, 38, figs. 1-4), os.84.7-5 (plesiotype, 9 RV: PI. 12, 32, fig. 5; PI. 12,
34, fig. 5; Text-figs, la-d), os.84.7-6 (plesiotype, 9 LV: PI. 12, 36, fig. 5; PI. 12, 38, fig. 5; Text-fig. 2).
All the specimens are from the type locality and horizon.

Carapace reniform in lateral view with broadly rounded anterior margin; posterior margin more
squarish. Asymmetrical ridges on both valves; a ridge follows the valve margin in the posterior area
of the right valve and a wavy ridge occurs in the ventral area of the left valve. Two nodes occur
dorsally, one on either side of a dorso-median sulcus, with the anterior one developing a faint ridge
parallel to the antero-dorsal margin.

Explanation of Plate 12, 32

Figs. 1-4, $ car. (neotype, os.84.7-1, 860/um long): fig. 1, rt. lat. ; fig. 2, dors.; fig. 3, It. lat. ; fig. 4, vent.; fig. 5, 9 RV, int. lat. (os.84.7-5,
820/j.m long). Scale A (400/am; X 60), figs. 1-5.

Stereo- Atlas of Ostraeod Shells 12, 33 Limnocythere shixiaensis (3 of 8)

Remarks: Since all the original type specimens of this species have been destroyed, the specimens illustrated

herein are designated as a neotype and plesiotypes. Originally this species was thought to belong to
the genus Leucocythere (Wang 1980, op.cit.), but since it appears to be impossible to differentiate
between Leucocythere and Limnocythere on carapace characters alone, it is now decided to place it
in the latter genus. It is considered that the unusual morphological features of this species (asym-
metrical valves and ridges) do not simply result from a salinity change as was originally suggested
(Wang 1980, op.cit.). It should be mentioned that Limnocythere inderica Sharapova, as described
by Kazmina ( Trudy Inst. Geol. Geofiz. sib. Otd., 264a, 1-108, 1975), has a right valve with a peripheral
ridge and a left valve with a ventral ridge, or vice versa, but unfortunately she did not illustrate the
shape of the carapace. In the Danangou section in Yuxian County, Hebei Province, the author found
the carapace of a species with a posterior peripheral ridge in both valves (therefore symmetrical
ones), which was determined as Leucocythere burangensis Huang (Huang Baoren et. al., 1982; in:
Series of the Scientific Expedition to the Qinghai-Xizang Plateau -Palaeontology ofXizang - Book 4,
326-348); again the carapace was not illustrated in that book.

Distribution: Mr. Zhao Lianbi collected 125 carapaces and other single valves from the Nihewan Formation in the

Nihewan-Shixia village section in 1 964 ; the exact location remains unknown. The author collected a
few similar specimens from a yellow-green silt-clay and a green-yellow silt at the type locality
(Xiashagou section) in 1979. The Xiashagou section is located between Nihewan and Shixia and
yielded the famous Nihewan vertebrate fauna of Middle-Upper Villafranchian age (P. T. de Chardin
et al.,Annls Paleont., 19, 8, fig. 2, 1930).

Explanation of Plate 12, 34

Figs. 1-4, 9 car. (os.84.7-2, 820 /u,m long): fig. 1, rt. lat.; fig. 2, dors.; fig. 3, It. lat.; fig. 4, vent.; fig. 5, 9 RV dors, (os.84.7-5, 820/u.m long).
Scale A (400/u.m; x 60), figs. 1-5.

Stereo- Atlas of Ostracod Shells 12, 32

Limnocy there shixiaensis (2 of 8)

Stereo- Atlas of Ostracod Shells 12, 34

Limnocythere shixiaensis (4 of 8)

•! ?*

Stereo-Atlas of Ostracod Shells 12, 35 Limnocythere shixiaensis (5 of 8)

Text-fig. 1. 9 RV (os.84.7-5) of L. shixiaensis: a, dors.; b, int. lat.; c-d, details of anterior and posterior hinge teeth.

Explanation of Plate 12, 36

Figs. 1-4, 9 car. (os.84.7-3, 810/xm long): fig. 1, rt. lat.; fig. 2, dors.; fig. 3, It. lat.; fig. 4, vent.; fig. 5, 9 LV int. lat. (os.84.7-6, 820A<.m long).
Scale A (400 /urn; x60), figs. 1-5.

Text-fig. 2. 9 LV (os.84.7-6) of L. shixiaensis: int. lat.

Explanation of Plate 12, 38

Figs. 1-4, 9 juv.car. (os.84.7-4, 740/u.m long): fig. 1, rt. lat.; fig. 2, dors.; fig. 3, It. lat.; fig. 4, vent.; fig. 5, 9LV dors, (os.84.7-6, 820/j.m long).
Scale A (400/rm; x 60), figs. 1-5.

Stereo-Atlas of Ostracod Shells 12, 36

Stereo-Atlas of Ostracod Shells 12, 38

Limnocy there shixiaensis (8 of 8)

Limnocythere shixiaensis (6 of 8)

Stereo-Atlas of Ostracod Shells 12 (8) 39-44 (1985) Hiltermannicythere emaciata (1 of 6)

595.337.14 (119.9) (261.2 : 162.007.53 + 560) : 551.351

ON HILTERMANNICYTHERE EMACIATA (BRADY)

by John Athersuch & David J. Horne
(B.P. Research Centre, Sunbury-on-Thames & City of London Polytechnic)

Hiltermannicythere emaciata (Brady, 1867)

1867 Cythere emaciata sp. nov. G. S. Brady, Rept .Brit Assoc .Adv .Sci ., (for 1866), 210.

1868 Cythere emaciata Brady; G. S. Brady, Trans. Linn. Soc.Lond., 26, 414-415, pi. 31, figs. 31-37.

1874 Cythere emaciata Brady; G. S. Brady, H. W. Crosskey & D. Robertson, Palaeontogr.Soc. [Monogr.] 28, 161, pi. 9, figs. 14-17.

1940 Cythereis emaciata (Brady); O. Elofson, J .mar. Biol., 24, 498-499, figs. 6-8.

Type specimens: Not present in the Brady collection at the Hancock Museum, Newcastle or in the British Museum

(Natural History) and therefore presumed lost.

Type locality: In his original description Brady ( op.cit .) gave the occurrence of this species as “Hebrides (locality

doubtful), and many other places in Great Britain and Ireland”.

Figured specimens: Brit. Mus. (Nat. Hist.) nos. 1984, 184 (9 car.; LV : PI. 12, 40, fig. 1 ; PI. 12, 42, fig. 3; RV: PI. 12, 40,

fig. 2; PI. 12, 42, fig. 2; appendages: Text-figs, la-c, 2a-c), 1984.185 (c? car.; LV: PI. 12, 40, fig. 3;
RV: PI. 12, 42, figs. 1, 4; copulatory appendage: Text-fig. 2d). Both specimens are from a sample
collected by N. Ainsworth in Dublin Bay, Eire (approx, lat. 53° 18'N, long. 06° 05'W), 1 km off
Sandy Cove Harbour, depth 15 m.

Explanation of Plate 12, 40

Figs. 1, 2, $ car. (1984.184, 900 /um long): fig. 1, LV, ext. lat.; fig. 2, RV, ext. lat.; fig. 3, 6 RV, ext. lat. (1984.185, 830/u.m long).
Scale A (200/xm; x 65), figs. 1-3.

Stereo- Atlas of Ostracod Shells 12, 41 Hiltermannicythere emaciata (3 of 6)

Diagnosis: Carapace elongate, tapering posteriorly; posterior margin with a pointed extremity at about mid-

height. Reticulate ornament consists of relatively large, equal-sized, rounded and polygonal inter-
costal fossae, a few of which coalesce. Reticulum dominated by anterior concentric costa and three
distinct longitudinal costae. Ventral and median costae linked posteriorly by oblique costa. Male
carapace asymmetric; right valve compressed postero-ventrally, with ventral costa not developed
posteriorly. Distal process of male copulatory appendage lamellar, subtriangular, truncated
terminally.

Remarks: H. rubra (G. W. Muller, 1894), from the Mediterranean, is very similar to H .emaciata . However,

H. rubra is more quadrate in carapace outline, more truncate posteriorly and the ventral costa is
more prominent and not linked posteriorly to the median costa. The sexual dimorphism of the male
right valve is strong in H. emaciata but only weakly developed in H. rubra. The distal process of the
male copulatory appendage of H. rubra is distally pointed, while that of H. emaciata is relatively
broader and distally truncate (this comparison is based on Elofson’s ( op.cit . ) illustration of the male
copulatory appendage of H .emaciata as well as on our single, damaged specimen). For further
discussion of H. rubra see Athersuch & Horne {Stereo-Atlas of Ostracod Shells 12, 45-48, 1985).

Specimens identified as Falunia (Hiltermannicythere) emaciata from the Miocene of Turkey
by Doruk (1973, unpublished Ph.D thesis. University of Leicester) correspond well to our Recent
examples of the species and her material is regarded as conspecific.

Distribution: The distribution of both living and fossil species of Hiltermannicythere in NW Europe needs

reappraisal. H. emaciata appears to live in shallow sublittoral waters around the coasts of England,
Wales and Ireland. Recent Mediterranean records should probably be referred to H. rubra (G. W.
Muller). H. emaciata has also been recorded from the Pleistocene of Great Britain (Brady, Crosskey
& Robertson, op. cit.) and the Miocene of Turkey (Doruk, op.cit.).

Explanation of Plate 12, 42

Figs. 1 , 4, <S (1984. 185, 830 /urn long): fig. 1, LV, ext. lat.; fig. 4, LV, int. lat.; figs. 2,3,9 (1984.184, 900/u.m long): fig. 2, RV, dors.; fig. 3,
LV, dors. Scale A (200/nm; x 65), figs. 1-4.

Stereo-Atlas of Ostracod Shells 12, 40

Hiltermannicythere enmciata (2 of 6)

Stereo-Atlas of Ostracod Shells 12, 42

Hiltermannicythere emaciata (4 of 6)

Stereo- Atlas of Ostracod Shells 12, 43

Hiltermannicythere emaciata (5 of 6)

Stereo- Atlas of Ostracod Shells 12, 44

Hiltermannicythere emaciata (6 of 6)

Text-fig. 2. ? appendages (1984.184) of H. emaciata : a, first leg; b, second leg; c, third leg; d, dcopulatory appendage (damaged; 1984.185).

Stereo- Atlas of Ostracod Shells 12 (9) 45-48 (1985)
595.337.14 (119.9) (262 : 161.034.33 + 161.034.34)

Hiltermannicythere rubra (1 of 4)

551.351

1894

1971

1976

1979

ON HILTERMANNICYTHERE RUBRA (MULLER)

by John Athersuch & David J. Horne
(B.P. Research Centre, Sunbury-on-Thames & City of London Polytechnic)

Hiltermannicythere rubra (Muller, 1894)

Cythereis rubra sp. nov. G. W. Muller, Fauna Flora Golf.Neapel, 21, 372, pi. 28, figs. 21, 26, pi. 31, figs. 2, 3.
Carinocythereis sp.; P. J. Barbeito-Gonzalez, Mitt. hamb.zool.Mus. Inst. , 67, 281, pi. 14, figs, lc, 2c, 3c.

Hiltermannicythere aff.// .rubra (Muller); G. Bonaduce, G. Ciampo & M. Masoli .Pubbl.Staz-Zool. Napoli, 40 (1), (for 1975), 49,
pi. 28, figs. 1-5.

Hiltermannicythere rubra (Muller); J. Athersuch, J.nat.Hist., 13, 140, fig. 2 (13).

Type specimens:

Type locality:
Figured specimens:

Athersuch (Pubbl.Staz.zool. Napoli 40 (2), 344-348, 1976) reported specimens in both sections of
the G. W. Muller collection; 15 specimens at the Institut fiir Spezielle Zoologie und Zoologisches
Museum der Humboldt Universitat zu Berlin, E Germany and 13 specimens at the Zoologischen
Institut, Greifswald, E Germany.

Bay of Naples, Italy (lat. 40° 40'N, long. 14° 10'E); Recent.

Brit. Mus. (Nat. Hist.) nos. 1984. 186 (? car.; LV: PI. 12, 46, fig. 1;RV:P1. 12, 46, fig. 2), 1984.187
(d car.; RV: PI. 12, 46, fig. 3; LV: PI. 12, 48, figs. 1, 4; copulatory appendage: Text-fig. 1), 1984.188
(2 car.; RV: PI. 12, 48, fig. 2; LV: PI. 12, 48, fig. 3). All collected by J. Athersuch from localities in
Cyprus during November, 1973. 1984.186 and 1984.188 are from sand in a small cove just North of
Cape Greco (lat. 33° 55'N, long. 34° 10'E), water depth 10m, salinity 39.4%0, temperature 21.5°C.
1984.187 was found in fine sand in Ayia Napa Harbour (lat. 34° 58'N, long. 34° 00'E), water depth
3 m, temperature 20°C.

Carapace subquadrate, tapering slightly towards posterior; posterior margin truncate with ventral
part bearing four distinct marginal denticles. Numerous intercostal fossae of varying size, often

Explanation of Plate 12, 46

Figs. 1, 2, $ (1984.186, 720/um long): fig. 1, LV, ext. lat.; fig. 2, RV, ext. lat.; fig. 3, 6 RV, ext. lat. (1984.187, 760/cm long)
Scale A (200 /xm; x 80), figs. 1-3.

Diagnosis:

Remarks:

Stereo-Atlas of Ostracod Shells 12, 47 Hiltermannicythere rubra (3 of 4)

forming small clusters and frequently coalescing. Reticulum dominated by median and ventral
longitudinal costae, which are not linked posteriorly or only weakly so. Seen internally, hinge line
slightly below dorsal margin. Distal process of male copulatory appendage triangular, lamellar, with
a curved, pointed prolongation at its anterior corner.

H. rubra closely resembles H.emaciata (Brady, 1867) but differs in details of shape and ornament,
particularly the disposition of the costae, and in the nature of the adult sexual dimorphism (see
Athersuch & Horne, Stereo-Atlas of Ostracod Shells 12, 39-44, 1985). The sexual dimorphism of the
right valve, conspicuous in H.emaciata, is only weakly developed in H. rubra. Furthermore, on the
basis of the limited number of specimens of the two species that we have seen, it appears that in
H. rubra the male is the larger dimorph, while in H.emaciata the reverse is true.

Falunia capsula Uliczny, 1969 (Hemicytheridae und Trachyleberidae (Ostracoda) aus dem
Pliozdn der Insel Kephallinia , Dissertation, Univ. Munich) and Cythereis rubra pontica Caraion,
1967 [ Fauna Republicii Socialiste Romania, 4, Crustacea, pt. 10 (Ostracoda)-, Fam. Cytheridae
(Ostracode marine §i salmastricole), 1-164, Bucarest] also appear to be very similar to H. rubra but
require detailed examination before they can be assigned with certainty to this species. We consider
that the larger size of the specimens reported by Bonaduce et.al. (1976, op.cit.) is not a sufficient
difference to preclude them from being referred to this species.

Recent of the Mediterranean: Cyprus (herein),

Greece (Barbeito-Gonzalez, op.cit.), Italy (Muller;

Bonaduce et al., op.cit.)-, ? Black Sea (Caraion,
op.cit.). ?Pliocene of Greece (Uliczny, op.cit.).

Because of the similarity of this essentially Mediter-
ranean species to H.emaciata, all the records of both
species require further investigation before their full
geographical and stratigraphical distribution can be
ascertained.

Distribution:

Text-figure 1. Male copulatory appendage

(1984.187).

Explanation of Plate 12, 48

Figs. 1, 4, 6 (1984.187, 760 long); fig. 1, LV, ext. lat.; fig. 4, RV, int. lat.; figs. 2, 3, 9 (1984.188, 720 ,um long); fig. 2, RV dors.; fig. 3,
LV dors. Scale A (200/u.m; x 80), figs. 1-4.

Stereo-Atlas of Ostracod Shells 12, 46

Hiliermannicythere rubra (2 of 4)

Stereo-Atlas of Ostracod Shells 12 (10) 49-56 (1985) Brephocharieis complicata (1 of 8)

595.336.13 (113.312) (429 : 162.005.51) : 551.35 + 552.55

Derivation of name:
Diagnosis:

Remarks:

ON BREPHOCHARIEIS COMPLICATA (Salter)

by David J. Siveter
(University of Leicester , England)

Genus BREPHOCHARIEIS gen. nov.

Type-species: Beyrichia complicata Salter, 1848

Greek brephos, embryo + charieis, beautiful; resemblance of LI + L2 to a foetus. Gender feminine.
Quadrilobate tallinnelline. Lobes stout, equally elevated except for depressed dorsal half of L4,
confluent with an evenly curved, rounded connecting lobe sited very close to velum in lateral view;
laterovelar furrow virtually lacking. L2 short, straight, slopes towards the end of the posteriorly
reflexed dorsal part of LI. L3, LI and, to a lesser extent L4 have cusps above the dorsum. Sulci
(S1-S3) well developed, as is infravelar antral dimorphism. Female dolon moderately wide, extends
from near anterior cardinal corner to below S2, continues posteriorly (as in tecnomorphs) as a fine
ridge along ventral part of the valve.

The combined nature of its adventral structure, dimorphism (resembling that of the type-genus
Tallinnella) and lobation indicate the tallinnelline affinities of Brephocharieis. However, it is readily
distinguished from other, mainly middle Ordovician Baltic genera of the Tallinnellinae Schallreuter,
1976 ( Palaeontographica (A), 153, 165; also R. E. L. Schallreuter, Geologie, 15, 200, 1966). Like
Tetrada, brephocharieis lacks the distinctive laterovelar furrow (and very prominent velum) found
in Tallinnella Opik (type-species: T. dimopha Opik.j Publ. Geol. Inst. Univ. Tartu., 50, 24, 1937).
Brephocharieis is distinguished from Tetrada Neckaja (type-species: T. memorabilis (Neckaja,
1953); see R. E. L. Schallreuter Palaeontographica (A), 153, 166. 1976) by its more clearly differ-
entiated quadrilobation and connecting lobe, more prominent antral dimorphism and its ventrally
present velum in both dimorphs .Homeokiesowia Shallreuter ( Stereo-Atlas of Ostracod Shells , 6. 75-
78, 1979) differs in having dissolved lobes, no connecting lobe and a wide, distinct velum.

Explanation of Plate 12, 50

Fig. 1, 9 RV, ext. lat. (OS 12576, 2150/zm long); figs. 2-5, $ LV (OS 12577, 2270/xm long): fig. 2, ext. lat. ; fig. 3, ext. ant.; fig. 4 int.
vent. obi. showing antrum; fig. 5, ext. vent. Scale A (500/^m; x 25), fig. 1; scale B (500/a.m; x 25), figs. 2-5.

Stereo-Atlas of Ostracod Shells 12, 51

Brephocharieis complicata (3 of 8)

Brephocharieis complicata (Salter, 1848)

1848

1851

1852

1855

1868

1869

1881

1890
non 1892
1908
1934
1938
1947
1963
1966
1966

1977

Beyrichia complicata, Salter; J. W. Salter in : J. Philips & J. W. Salter, Palaeontological appendix, Mem. Geol. Surv. U.K., 2.
234 (pars), 352 (pars), pi. 8, figs. 16, 16a.

Beyrichia complicata (Salt.) ; F. McCoy in: A. Sedgwick & F. McCoy, A synopsis of the classification of the British Palaeozoic
Rocks . . . (1), 1851, 136 (pars), non pi. IE, figs. 3, 3a (= Sedgwick Museum, Cambridge, no. A. 16696 = Harperopsis scripta
(Harper, 1947)).

Beyrichia complicata-, J. W. Salter in \ A. Sedgwick & F. McCoy, Ibid., appendix A, ii (pars), non pi. IE, figs. 3, 3a
(= Sedgwick Museum, Cambridge, no. A 16696 = Harperopsis scripta (Harper, 1947)).

Beyrichia complicata , Salter; T. R. Jones, Ann. Mag. nat. Hist., (2), 16, 163-165 (pars), pi. 6, figs. 1-4, non fig. 5 (= on
British Geological Survey London, slab no. 49449 = Harperopsis scripta (Harper, 1947)).

Beyrichia complicata Salter; T. R. Jones & H. B. Holl.Ann. Mag. nat. Hist., (4), 2, 59 (pars).

B. complicata, Salter; T. R. Jones, On the Palaeozoic Bivalved Entomostraca , 15, text-fig. 22, non 11, Hertford.
Beyrichia complicata, Salter; J. W. Salter & R. Etheridge in : A. C. Ramsey, Geology of North Wales (2nd ed.), 3, Mem.
Geol. Surv. U.K., 487 (pars), pi. 19, fig. 9, non 106, London.

Tetradella complicata, Salter; E. O. Ulrich, Jl. Cincinn. Soc. nat. Hist., 13, 112.

Beyrichia complicata , Salt.; T. E. Marr, Geol. Mag., 108.

Tetradella complicata (Salter); E. O. Ulrich & R. S. Bassler, Proc. U.S. natn. Mus., 35, 306.

Tetradella complicata (Salter); R. S. Bassler & B. Kellett, Spec. Pap. geol. Soc. Am., 1, 190, 480 (pars).

Beyrichia complicata Salter; C. J. Stubblefield, Summ. Progr. geol. Surv., pt. 2 (for 1936), 35.

Tetradella complicata (Salter); J. C. Harper, Geol. Mag., 84, 346, pi. 10, fig. 3.

Tallinnella complicata (Salter); N. Spjeldnaes, Palaeontology , 6, 255, pi. 36, figs. 9-13, text-fig. 1.

Gunnaropsis complicata ; R. E. L. Schallreuter, Geologie, 7, 853.

Cerninella (Cerninella) complicata (Salter, 1848); A. Pfibyl ,Cas. Narodm'ho Musea, odd, ph'rod., 135,206-207 (pars)-, non
pi. 2, figs. 4-9, text-figs. 4a-c.

Tetradella complicata Salter; F. M. Swain in: F. M. Swain (ed.), Stratigraphic Micropaleontology of Atlantic Basin &
Borderlands , 29, fig. 3 (34), Elsevier, Amsterdam.

Explanation of Plate 12, 52

Fig. 1, 6 RV, ext. lat. (OS 12574, 2125 Atm long) ; figs. 2-5, 3 LV (OS 12575, 2125 Atm long) : fig. 2, ext. lat. ; fig. 3, ext. ant.; fig. 4, detail of
ext. vent, showing ornament and velum; fig. 5, ext. vent. Scale A (500 Atm; x 25), figs. 1-3, 5; scale B (200 Atm; x62), fig. 4.

Stereo-Atlas of Ostracod Shells 12. 50

Brephocharicis complicate i (2 of 8)

Stereo-Atlas of Ostracod Shells 1

Stereo-Atlas of Ostracod Shells 12, 53 Brephocharieis complicata (5 of 8)

1978 ‘Tallinnella’ complicata (Salter, 1848); D. J. Siveter m : R. H. Bate & E. Robinson (eds.), A Stratigraphical Index of British
Ostracoda, Geol. J. special issue 8, 41, 43, 45, 48, pi. 1, figs. 7, 8.

1979 Cerninella complicata (Salter, 1848); A. Pribyl, Sbornik Ndrodniho Musea (B), 33 (for 1977), 67-68 (pars), pi. 3, figs. 1,2,
non pi. 3, figs. 3-7, text-figs. 3(3-5), 10 (1), 11 (4), 16 (2).

Lectotype: Designated herein [the neotype designation of Pribyl (1966, 206) is invalid], A left valve

tecnomorph ( not heteromorph as stated by Pribyl 1979, 143) ; internal and external moulds on rock
pieces of British Geological Survey Museum, London, nos. GSM 24525 and GSM 24526 (= part
and counterpart) respectively. The internal mould was figured by Jones 1855 (pi. 6, fig. 3) and Pribyl
1979 (pi. 3, fig. 1). The original of ‘Beyrichia’ complicata of Jones 1855, pi. 6, fig. 4 (= a
tecnomorphic right valve internal mould) is also on GSM 24525. From Llan Mill (see ‘Type locality’).

The specimens of Jones on GSM 24525 and 24526 are part of Salters 1848 syntype
collection: they were considered as “lectosyntypes” by Stubblefield (op. cit.) and Harper (op. cit) ;
furthermore, the pieces were catalogued in 1848 (Museum labels) and Jones (1855, 163, 164)
acknowledges Salters’ help and the use of ‘Museum of Practical Geology’ (= GSM) specimens.

The specimens of ‘Beyrichia’ complicata of Jones 1855, pi. 6, figs. 1, 2 are wax casts of left
and right valve tecnomorphs respectively, now in the British Museum (Nat. Hist.), London, no.
I 6323; the original external moulds, from Llan Mill, have not been identified but the probability is
that they were also taken from amongst the many external moulds on the original slabs held by the
British Geological Survey. Topotype rock piece GSM 24527, containing the original of Harper 1947
(op. cit., pi. 10, fig. 3 = a heteromorphic left valve internal mould), is also possibly syntype material
(Stubblefield 1938). GSM 24525, 24526 and 24527 contain tens of moulds of B. complicata.

The original of ‘Beyrichia’ complicata of Jones 1855, pi. 6, fig. 5, is a small tecnomorphic left
valve from Harnage, Shropshire, on British Geological Survey, London, slab no. 49449 and belongs
Xo Harper op sis scripta (Harper, 1947); see C. R. Jones & D. J. Siveter, Stereo-Atlas of Ostracod
Shells , 10, 5-12, 1983. Museum label and catalogue information claim the original of Jones 1855
pi. 6, fig. 5, is British Museum (Nat. Hist.) no. I 6325, a single larval quadrilobate palaeocope also
from Harnage. As Jones states (1855,1 64) that the original is in the ‘Museum of Practical Geology’,
the specimen in the British Museum is considered not to be the figured specimen.

Explanation of Plate 12, 54

Fig. 1, 9 LV, ext. lat. (OS 12581 ; 2025 pm long); figs. 2, 3, 8 RV (OS 6665, 2180pm long): fig. 2, ext. lat. ; fig. 3, ext. post.; fig. 4, $ RV,
ext. lat. (OS 6666, 2050pm long). Scale A (500pm; x 26), figs. 1, 4; scale B (500pm; x 25), figs. 2, 3.

Stereo- Atlas of Ostracod Shells 12, 55 Brephocharieis complicata (7 of 8)

Type locality: Llandeilo ‘Flags’ at Llan Mill (= Lann Mill of Salter 1848), 3km E of Narbeth, Dyfed, S Wales;

Nat. Grid Ref.: SN 139 143. The precise locality and horizon is unknown but the hillside quarry
200m NW of Llan Mill Farm is the probable locality; it lies within the Bryn-glass Limestone
Member, Lampeter Velfrey Formation, Narberth Group of Addison 1974, Unpublished Ph.D.
thesis. Queen’s University, Belfast) and is of probable upper Llandeilo age.

Figured specimens: British Museum (Nat. Hist.) nos. OS 6665 (8 RV : PI. 12, 54, figs. 2, 3), OS 6666 (9 RV : PI. 12, 54,

fig. 4), OS 12573 (tecnomorphic LV: PI. 12, 56, figs. 2-4), OS 12574 (c? RV: PI. 12, 52, fig. 1),
OS 12575 (8 LV: PI. 12, 52, figs. 2-5), OS 12576 (9 RV: PI. 12, 50, fig. 1), OS 12577 (9 LV: PI. 12,
50, figs. 2-5), OS 12578 (tecnomorphic LV: PI 12, 56, figs. 5, 6), OS 12579(9 RV: PI. 12, 56, fig. 1),
OS 12581 (9 LV: PI. 12, 54, fig. 1).

All specimens are silicified; obtained by acid preparation of limestone from the old quarry
c. 300m S of Cwm Agol Farm,c. 8km W of Llandeilo, Dyfed, S Wales; appox. lat 51°51'N, long.
4°05'W (Nat. Grid Ref.: SN 5655 2070). Llandeilo ‘Flags’, Llandeilo Series, Ordovician.

As for the genus. Brephocharieis is at present monotypic.

‘Beyrichia ’ complicata was the first ostracode species erected from the British Ordovician (Siveter
1978, 41). Within the silicified material described herein there is notable size variation (mixed
chronodemes/ecodemes?) of females (length, including dolon: c. 1900-2270pm) within a single,
acid prepared sample (e.g. PI. 12, 54, figs. 1, 3 cf. PI. 12, 56, fig. 1). Moreover, some of the smaller
females (PI. 12, 56, fig. 1) appear to have relatively more constricted lobes (possibly a factor of size
or preservation). A tendancy to develop more constricted lobes (not quite cristation) is also seen in
younger instars (e.g. PI. 12, 56, fig. 5).

Llandeilo Series around Narberth and Llandeilo, S Wales. The range has now been extended (C. Jones,
in prep.) to include the upper Llanvirn and the Costonian Stage, Caradoc Series of that area.

Explanation of Plate 12, 56

Fig. 1, 9 RV, ext. lat. (OS 12579; 1950pm long) ; figs. 2-4, tecnomorphis LV (OS 12573; 1625 pm long): fig. 2, ext. lat., fig. 3, ext. vent. ;

fig. 4, detail anterodors. ; figs. 5, 6, tecnomorphic LV (OS 12578; 1500pm long): fig. 5, ext. lat.; fig. 6, ext. vent.

Scale A (500pm; x 25), figs. 1-3, 5, 6; scale B (200pm; x75), fig. 4.

Diagnosis:

Remarks:

Distribution:

Stereo-Atlas of Ostracod Shells 12, 54

Brephocharieis complicate/ (6 of 8)

Stereo-Atlas of Ostracod Shells 12, 56

Brephocharieis complicate/ (8 of 8)

Stereo-Atlas of Ostracod Shells 12 (11) 57-60 (1985) Henningsmoenia gunnari (1 of 4)

593.336.13 (113.312) (430.1 : 161.011.55) : 55.35 + 552.55

ON HENNINGSMOENIA GUNNARI (THORSLUND)

by R. J. Orr

(Queen’s University of Belfast, Northern Ireland)

Genus HENNINGSMOENIA Schallreuter, 1964
1964 Henningsmoenia gen. nov.; R. E. L. Schallreuter, Ber. geol. Ges. DDR, 2, 91.

Type-species (by original designation): Winchellatia gunnari Thorslund, 1948
Diagnosis: Unisulcate Steusloffiinae with a long sigmoidal S2. Preadductorial node a low bulb or indistinct.

Plica weak. Posteroventral lobe with a ridge-like crista; crista short or long. Female velum with a
S-like step anterocentrally and a ridge-like torus; tecnomorphs with or without torus. Antrum
botulate and dolonal. Lateral surface tuberculate or pustulate and also maybe finely reticulate.

Henningsmoenia gunnari (Thorslund, 1948)

1948 Winchellatia gunnari sp. nov. P. Thorslund Bull. geol. Instn. Univ. Upsala, 23, 368, pi. 20, figs. 2, 3.

1976 Henningsmoenia gunnari (Thorsland); R. E. L. Schallreuter, Palaeontographica (A) 153, (4/6), 199, pi. 7, figs. 1-8.

1982 Henningsmoenia gunnari (Thorsland); L. K. Gailfte,/«: R. Z. Ulst, L. K. Gaillte & V. 1. Yakovleva , Ordovician Latvia , 119, 124,
130, 191. Riga.

1983 Henningsmoenia gunnari (Thorsland); R. E. L. Schallreuter, Palaeontographica (A) 180, (4/6), 178, pi. 11, fig. 7 (q. v. for full
synonomy).

1984 Henningsmoenia gunnari (Thorslund); R. E. L. Schallreuter, Geol. For. Stockh. Forh., 101, 95.

Holotype: Palaeontological Institute, University of Uppsala, Sweden; tecnomorphic left valve (listed as a right

valve in Thorslund 1948, 373, pi. 20, fig. 2).

Type locality: Kullatorp core (depth 65.05m), Kinnekulle, Vastergotland, Sweden; approx, lat. 58° 30'N,

long. 13° 25'E. Skagen Limestone (=Johvi Stage), middle Ordovician.

Explanation of Plate 12, 58

Figs. 1, 3, $ LV (K 10034, 1020/i.m long): fig. 1, ext lat.; fig. 3, ext. obi. vent. Figs. 2, 4, $ RV (K 10035, 1030p.m long): fig. 2, int. ant.
showing part of antrum; fig. 4, int. lat. Scale A (250p.m; x 65), figs. 1,3,4; scale B (100/um; x 86), fig. 2.

Stereo- Atlas of Ostracod Shells 12, 59 Henningsmoenia gunnari (3 of 4)

Figured specimens:

Diagnosis:

Remarks:

Distribution:

Ulster Museum, Belfast, nos. K 10034 (9 LV: PI. 12, 58, figs. 1, 3), K 10035 (9 RV: PI. 12, 58, figs.
2, 4; PI. 12, 60, fig. 2), K 10036 (tecnomorphic RV: PI. 12, 60, figs. 1, 3) and K 10037 (juv. RV: PI.
12, 60, fig. 4). All specimens are silicified, from Backsteinkalk erratic boulder no. Stb 1, from
Staberhuk, Isle of Fehman, near Puttgarden, West Germany. Backsteinkalk boulder Type 1B1;
source = ? Skagen Limestone, Sweden (Schallreuter, Palaeontographica (A), 44, (1/3), 65, 1973).
Sample courtesy of Dr. R. E. L. Schallreuter.

Henningsmoenia species with a small, bulb-like preadductorial node. Plica weak, only developed
either side of sulcus. Posteroventral crista short, posterior end spine-like, merges with valve
posterior to sulcus. Adult male and female similar in lateral view. Anterocentrally female velum
follows a S-like course. Female torus strong; broad dolonal botulate antrum; tecnomorphic torus
weak to absent. Right valve with a narrow ridge proximal to the free margin. Primary surface
ornament pustulate; second-order ornament a fine reticulation, often missing.

The fine reticulation of Henningsmoenia gunnari is illustrated for the first time herein. In early
juvenile specimens the crista is represented only by a weak posterior spine. This juvenile crista is
similar to that seen in the female of Henningsmoenia billingensis Schallreuter (op. cit., 95, fig. 3,
1984) from the Upper Dalby Limestone (= Idavere Stage). In Henningsmoenia costa Orr (Stereo-
Atlas of Ostracod Shells , 12, 61-68, 1985), from the upper Ordovician, the crista is complete to the
anterodorsal margin.

Middle Ordovician of the Baltic and Scandinavia: Skagen Limestone (Johvi [DiJ Stage), Siljan
district and Vastergotland, Sweden; Lower Chasmops Shale (Idavere [Cm] Stage), Oslo Region,
Norway; Auleliai Member (upper Cm, lower Di), Lithuania; Adze and Blidene formations (Cm,
Di and Keila [Du] Stage, Western Latvia; Svjantusk Member (Cm) and Auleliai Member, Eastern
Latvia.

Explanation of Plate 12, 60

Figs. 1 , 3, tecnomorphic RV (K 10036, 860 Aim long): fig. 1, ext. lat.; fig. 3, ext. obi. vent. fig. 2, 9 RV, int. vent., detail of torus, velum and
margin (K 10035); fig. 4, juv. RV, ext. lat. (K 10037, 520Aim long).

Scale A (250Aim; x78), fig. 1; scale B (250auti; x 82), fig. 3; scale C (50auti; x 115), fig. 2; scale D (250Aim; x 125), fig. 4.

Stereo-Atlas of Ostracod Shells 12, 58

Henningsmoenia gunneri (2 of 4)

Stereo-Atlas of Ostracod Shells 12, 60

Henningsmoenia gunneri (4 of 4)

> i

.

■>;V ' ' V 5:

.:v/. '■ ■

Stereo-Atlas of Ostraeod Shells 12 (12) 61-68 (1985) Henningsmoenia costa (1 of 8)

595.336.13 (113.313) (415 : 162.007.53): 551.35 + 552.55

ON HENNINGSMOENIA COSTA ORR sp. nov.

by R. J. Orr

(Queen’s University of Belfast, Northern Ireland)

Holotype:
Type locality:

Derivation of name:
Figured specimens:

Diagnosis:

Henningsmoenia costa sp. nov.

Ulster Museum, Belfast, no. K 10024; 9 RV.

SW facing cliff, small cove about 230m S W of renovated Martello Tower, 800m SE of Portrane,
Co. Dublin, Eire; approx, lat. 53° 29'N, long. 6° 06'W (Nat. Grid.: 0 257504). Bed 7, Locality CN,
Lower Limestones Member, Portrane Limestone; Cautleyan, Ashgill Series, upper Ordovician.
Latin costa , rib; referring to the prominent ridge.

Ulster Museum, Belfast, nos. K 10024 (holotype, 9 RV: PI. 12, 62, fig. 1), K 10025 (9 RV: PI. 12,
62, fig. 2; PI. 12, 66, figs. 1, 3), K 10026(9 RV: PI. 12, 62, fig. 3; PI. 12, 66, fig. 2), K 10027 (d LV:
PI. 12, 62, fig. 4; PI. 12, 64, fig. 5; PI. 12, 68, fig. 1), K 10028 (9 LV: PI. 12, 62, fig. 5), K 10029
(d LV: PI. 12, 64, figs. 1, 2), K 10030 (juv. RV: PI. 12, 64, fig. 3), K 10031 (d LV: PI. 12, 64, fig. 4),
K 10032 (tecnomorphic car.: PI. 12, 68, fig. 2) and K 10033 (9 LV: PI. 12, 68, fig. 3).

Specimen K 10026 is from Bed 9 and K 10027 and K 10031 are from Bed 22 of loc. F, S W
facing cliff, about 145 m S W of type locality, Lower Limestones Member, Portrane Limestone. The
other specimens are from the type locality. All the material is coarsely silicified.

Species of Henningsmoenia in which the posteroventral crista continues to the anterodorsal border.
Plica weak to absent. Female velum with diagnostic S-like step anterocentrally. No torus in tecno-
morph, possible torus in female. Marginal sculpture: a narrow marginal ridge in right valve and a
narrow admarginal ridge in left valve.

Explanation of Plate 12, 62

Fig. 1, 9 RV, ext. lat. (holotype, K 10024, 1 1 10/am long); fig. 2, $ RV, int. ant. velum and antrum (K 10025, 1005 yum long); fig. 3, 9 RV,
ext. vent. (K 10026, 1 101 /am long); fig. 4, d LV, int. ant. velum (K 10027, 1200 jam long); fig. 5, 9 LV, int. lat. showing antrum
(K 10028, 930/am long). Scale A (200/am; x65), figs. 1, 3, 5; scale B (lOOjam x85), fig. 2; scale C (100/am-; x70), fig. 4.

Stereo- Atlas of Ostraeod Shells 12, 63 Henningsmoenia costa (3 of 8)

Remarks: Henningsmoenia costa is the youngest recorded species of the genus. The older species assigned to

the genus display a progressive elongation of the crista on the wing-like posteroventral structure. In
H. billingensis Schallreuter (Geol. For. Stockh. Forh., 106, 95, 1984), from the upper Member of the
Dalby Limestone (Idavere [Cm] Stage), only a short posterior spine is present. In the type-species,
H. gunnari (Thorslund, 1948) from the Skagen Limestone (Johvi [ Di ] Stage; overall stratigraphic
range = Idavere to Keila [Dn] Stage), the short crista extends from the posterior spine to posterior
of the sulcus (see Orr, Stereo-Atlas of Ostraeod Shells, 12, 57, 1985). In H. costa this restricted
crista has developed into a long ridge which extends to the anterodorsal border. On first examination
this ridge might appear to be histial in position. However, a closer examination of some specimens
(see PI. 12, 64, figs. 4, 5, and PI. 12, 68, figs. 1, 2) reveals that between the ridge and the velum there
is a bend in the histial position.

The oldest specimens of H. costa are from the lower Member of the Portrane Limestone
(loc. F, bed 48) of Pirgu [Fic] age. Thus, there is a long stratigraphic interval between the youngest
record of//, gunnari and the introduction of//, costa. In terms of its lateral morphology and strati-
graphic distribution (Nabala [Fia] to Lower Pirgu [Fic] Stage) ‘Ordovicia’ pictis Neckaja ( Trudy
VNIGR1, 115,1 958) would appear to partly fill this gap. However, even though the exact nature of
its female dimorphic structures are unclear, both Schallreuter (Palaeontographica , (A), 178, (1/3),
34, 1982) and Gailfte ( Ordovician Latvia, 120, 1982, Riga) assign the species to Sigmobolbina
Henningsmoen. According to this interpretation ‘O’ .pictis represents an homeomorph oiHennings-
moenia.H. costa is homeomorphic with other, approximately contemporaneous species such as the
Baltic Naevhithis naevus Schallreuter ( Stereo-Atlas of Ostraeod Shells, 8, 137-140, 1981) and
? Pelecybolbina kolkaensis Gailite (In: Grigelis, A. A. (Ed.): The fauna and stratigraphy of Palaeo-
zoic and Mesozoic of Baltic and Byelorussia, 61, 1975, Vilnius). The female of H. costa has the
anteroventral step to the velum also seen in H. gunnari and N. naevus. However, A. naevus lacks the
crista and torus which are indicative of Henningsmoenia .

Explanation of Plate 12, 64

Fig. 1,2,3 LV (K 10029, 1200 /am long): fig. 1 , ext. lat.; fig. 2, ext. vent.; fig. 3, juv. RV, ext. lat. (K 10030, 620 /am long); fig. 4, 3 LV, int.

vent. (K 10031, 1010/am long); fig. 5, 3 LV, int. vent, detail of margin and velum (K 10027, 1200/am long).

Scale A (200/am; x 55), figs. 1,2; scale B (200/am; x 100), fig. 3; scale C (200/am; x 72), fig. 4; scale D (50/am; x 140), fig. 5.

Stereo-Atlas of Ostracod Shells 12, 6

Henningsmoenia costa (2 of 8)

Stereo-Atlas of Ostracod Shells 12, 64

Henningsmoenia costa (4 of 8)

■ -

mm

- ■ v v *»:

- :\

' f % *

Stereo-Atlas of Ostracod Shells 12, 65 Henningsmoenia costa (5 of 8)

Remarks (contd.): In//. gunnari and H. billingensis the distal edge of the velum is split ventrally to provide a ridge-like

torus. In H. costa a clear corresponding feature is not immediately apparent, but in female specimens
the distal end of the velum is much thicker than in the tecnomorphs (see especially PI. 12, 66, fig. 1).
This thickened velar border probably corresponds to a separate velum and torus, but because of the
coarseness of the silicification the individual ridges have coalesced. In one specimen (PI. 12, 68,
fig. 3) three narrow contiguous ridges are visible and may represent one velar and two toric ridges.

A further difference between//, costa and H. gunnari is the nature of the contact margin. In
H. gunnari the right valve has a contact list and an admarginal ridge and the left valve an inner semi-
sulcus. By contrast in H. costa the right valve has a marginal ridge and the left valve an admarginal
ridge (see Text-fig. 2). The precise contact structure present in H. billingensis is unknown.

‘Sigmobolbina’ cuneata Abushik & Sarv ( Paleontologija Drevnego Palaozoja Pribaltiki i
Podolii, 107, pi. 2, figs. 6-8, 1983), from the Molodovo Stage (= Nabala Stage), may also belong to
Henningsmoenia , though in this species there is no distinct crista and it more likely belongs in
Naevhithis .

Distribution: See text-fig. 1. Lower Limestones Member (Iocs. F, C, CN), Main Limestones Member (loc. B),

Upper Limestones Member (Iocs. B, K, A), Portrane Limestone, Portrane, Eire: Cautleyan to
Lower Rawtheyan, Ashgill Series, upper Ordovician.

Explanation of Plate 12, 66

Figs. 1, 3, 9 RV (K 10025, 1050/xm long): fig. 1, int. vent., ant. detail of antrum and velum.; fig. 3, int. lat. showing antrum; fig. 2, 9 RV,
ext. ant. (K 10026, lOlOjum long).

Scale A 100 /xm; x 140), fig. 1; scale B (200 /xm; x 86), fig. 2; scale C (200/xm; x 65), fig. 3.

Stereo- Atlas of Ostracod Shells 12, 67 Henningsmoenia costa (7 of 8)

H. gunnari

nm^auzzm

RV LV

H. costa

Text-fig. 1. Localities for H. costa (CN is the type locality).

Text-fig. 2. Contact sculptures in Henningsmoenia.

Explanation of Plate 12, 68

Fig. 1, <3 LV, int. lat. (K 10027, 1200 p.m long); fig. 2, tecnomorphic car., ext. vent. (K 10032, 960/um long); fig. 3, 9 LV, ext. vent, detail of
antrum and velar edge (K 10033, 1220p,m long).

Scale A (200/xm; x 60), fig. 1; scale B (200/xm; x 60), fig. 2; scale C (100/xm; x 140), fig. 3.

Stereo-Atlas of Ostracod Shells 12. 66

Henningsmoenia costa (6 of 8)

Stereo-Atlas of Ostracod Shells 12, 68 Henningsmoenia costa (8 of 8)

Stereo-Atlas of Ostracod Shells 12 (13) 69-72 (1985) Tyrrhenocythere amnicola (1 of 4)

595.337.14 (119.9) (262.2 : 161.022.38) : 551.313.1

ON TYRRHENOCYTHERE AMNICOLA (SARS)

by Timothy R. Maness & Roger L. Kaesler
(University of Kansas, Lawrence, Kansas, USA)

Genus TYRRHENOCYTHERE Ruggieri, 1955

Type-species (by original designation): Tyrrhenocythere pignattii Ruggieri, 1955 (= Cythere amnicola Sars, 1888)
1955 Tyrrhenocythere gen. nov. G. Ruggieri; J. Paleont., 29 (4), 698.

Diagnosis: A genus of Hemicytheridea, subrectangular, subtrapezoidal, or auriform in lateral view. Eye tubercle

distinct. Anterior vestibule partitioned into pockets; each pocket is the base of a fan-like bundle of
radial pore canals. Muscle-scar pattern consists of a row of four adductor scars, dorsomedian scar
divided, ventromedian scar usually divided; lone, sometimes divided, scar located dorsomedially ;
oblique row of three frontal scars anterior to adductors.

Remarks: The peculiar vestibule is the outstanding distinguishing feature of this genus. The appendages are

typically hemicytherid and were well illustrated by Sars (1888).

Tyrrhenocythere amnicola (Sars, 1888)

1888 Cythere amnicola sp. nov. G. O. Sars, Arch. Math. Naturv., 12, 305-312, pi. 15, figs. 8-10, pi. 10.

1902 Cythere sicula sp. nov. G. S. Brady, Trans, zool. Soc. Lond., 16, pt. 4, 198, pi. 25, figs. 1-7.

1947 Hemicy there sicula (Brady) ; Z. S. Bronstein,Faw/ia SSSR. Rakoobraznye2 , l.Ostracodapresnykh vod., 293, text-figs. 194-197,
Moskva. AN SSSR.

1955 Tyrrhenocythere pignattii sp. nov. G. Ruggieri, J. Paleont., 29 (4), 698-699, text-figs. 1-5.

1977 Tyrrhenocythere amnicola (Sars); N. Krstic, in H. Loffler & D. Danielopol (eds.), Aspects of Ecology and Zoogeography of
Recent and Fossil Ostracoda, 395-405, pi. 1, fig. 4, Junk, The Hague (q.v. for full synonymy).

Explanation of Plate 12, 70

Fig. 1, 8 RV, ext. lat. (1101251, 1250/u.m long); fig. 2, $ RV, ext. lat. (1101253, 1070/am long); fig. 3, <J RV, int. lat. (1101257, 1330/xm
long). Scale A (400/um; x48), figs. 1-3.

Stereo-Atlas of Ostracod Shells 12, 71 Tyrrhenocythere amnicola (3 of 4)

Type specimens: Uncertain ; either Zoological Museum, Oslo, or lost. Ruggieri deposited a paratype (of T. pignattii),

no. OT-167, with the Department of Micropaleontology, American Museum of Natural History,
New York.

The Cyane River near Siracusa, Sicily (approx, lat. 37°04'N, long. 15°17'E); Recent.

Mus. of Invert. Paleo., Univ. of Kansas, USA, nos. 1101250 (<3 LV: PI. 12, 72, fig. 1), 1101251
(<J RV: PI. 12, 70, fig. 1), 1101253 (9 RV: PI. 12, 70, fig. 2), 1101256 (< S LV: PI. 12, 72, fig. 3),
1101257 (c? RV: PI. 12, 70, fig. 3), 1 101269 (9 LV: PI. 12, 72, fig. 2). All from the Gulf of Corinth,
Greece, lat. 38°21'40"N, long. 22°25'35"E, water depth 73.2 m, from an 8.95 m long core taken by
the Lamont-Doherty Geological Observatory.

Surface, densely finely pitted; faint reticulation near free margin.

Strongly dimorphic; males more elongate and laterally compressed; females more auriform.

We are grateful to Dr. N. Krstic for her taxonomic suggestions based on study of our
specimens and to the American Museum of Natural History for lending us Ruggierri’s paratype.
Brackish- water deposits of the
Mediterranean Region. Krstic (op. cit.)
presents the following distribution:

Recent: Caspian, Pontian, Ionian, and
Tyrrhenian basins. Upper Pliocene-
Pleistocene: Caspian Basin.

Type locality:
Figured specimens:

Diagnosis:

Remarks:

Distribution:

Text-fig. 1. Camera-lucida drawing of Tyrrhenocythere
amnicola, 8 RV, int. view.

100 /urn

Explanation of Plate 12, 72

Fig. 1, 8 LV, ext. lat. (1101250, 1210^™ long); fig. 2, 9 LV, ext. lat. (1101269, 1100/am long); fig. 3, 8 LV, int. lat. (1101256,
1310/am long). Scale A (400/am; x48), figs. 1-3.

Tyrrhenocy there amnicola (2 of 4)

Stereo-Atlas of Ostracod Shells 12, 70

Stereo-Atlas of Ostracod Shells 12, 72

Tyrrhenocy there amnicola (4 of 4)

Stereo-Atlas of Ostraeod Shells 12 (14) 73-76 (1985) Paranotacythere magnified (1 of 4)

595.337.14 (116.312) (427.4 : 162.001.54) : 551.351 + 552.52

on PARANOTACYTHERE (PARANOTACYTHERE) MAGNIFICA LOMAX sp. nov.

by Allistair Lomax
( University of Hull, England)

Paranotacythere (Paranotachthere) magnified sp. nov.

Holotype:

Type locality:

Derivation of name:
Figured specimens:

Diagnosis :

University of Hull coll. no. HU.307.C. 1 ; 6 carapace.

[Paratypes: University of Hull nos. HU.307.C.2, HU.307.C.3, HU.307.C.4.1-10],

Bed C9A of coastal section of Speeton Clay, Speeton, E Yorkshire, England; lat. 54° 10'N, long.
0° 14' 40" W. Lower Hauterivian, lower Cretaceous.

Latin magnified , magnificent; in reference to its excellent ornamentation and preservation.
University of Hull coll. nos. HU.307.C. 1. (holotype, 6 car.: PI. 12, 74, fig. 1; PI. 12, 76, fig. 2),
HU.307.C.2 (9 LV: PI. 12, 74, fig. 2) and HU.307.C.3 (9 LV: PI. 12, 76, fig. 1). All specimens are
from the type locality and horizon.

A species of Paranotacythere ( Paranotacythere ) with strong reticulation, a large postero-dorsal
tubercle and in which the uppermost blade-like ventral rib tends to end at the anterior of the two
postero-ventral tubercles or only continues between them in much reduced fashion.

Explanation of Plate 12, 74

Fig. 1, 6 car., ext. It. lat. (holotype, HU.307.C. 1, 560/am long); fig. 2, 9 LV, ext. lat. (paratype, HU.307.C.2, 530/am long).
Scale A (100 /am; x 162), fig. 1; scale B (100/xm; x 181), fig. 2.

Stereo- Atlas of Ostraeod Shells 12, 75 Paranotacythere magnifica (3 of 4)

Remarks: The lower Hauterivian Paranotacythere (P.) magnifica differs from the upper Hauterivian P.costata

(Kaye) in the absence of thick and rounded ventral ribbing. P.costata also has a much less prominent
dorsal tubercle and reduced reticulation compared with P. magnifica. P. ramulosa (Sharapova),
which is first found in Bed C3, upper Hauterivian at Speeton, differs only in having two large
tubercles in the anteroventral region which replace the uppermost ventral rib. This suggests that
P. ramulosa is related to P. magnifica and is probably derived from it by loss of the ventral rib and
addition of the two anteroventral tubercles.

Distribution: P.(P.) magnifica is rare and has so far only been obtained from Bed C9A in the Speeton Clay of the

coastal section, E Yorkshire, England. Over two dozen specimens have been recovered. The
lithology is a dark grey, sideritic clay and common ostraeod associates include Schuleridea punctatula
(Roemer), Protocythere triplicata (Roemer), Protocythere hechti Triebel and Acrocythere
hauteriviana (Bartenstein).

Explanation of Plate 12, 76

Fig. 1, 9 LV, int. lat. (paratype, HU.307.C.3, 610/cm long); fig. 2, 6 car., ext. dors, (holotype, HU.307.C.1, 560/am long).
Scale A (100/xm; x 1 59), fig. 1 ; scale B (100/xm; x 175), fig. 2.

Stereo-Atlas of Ostracod Shells 12, 74

Stereo-Atlas of Ostracod Shells 12, 76

Paranoiac) there magnified (2 of 4)

Paranotacythere magnified (4 of 4)

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Stereo- Atlas of Ostracod Shells: Vol. 12, Part 1

CONTENTS

12 (1) 1-6
12 (2) 7-10

12 (3) 11-14

12 (4) 15-18
12 (5) 19-22
12 (6) 23-30
12 (7) 31-38
12 (8) 39-44
12 (9) 45-48
12(10) 49-56
12(11) 57-60
12(12) 61-68
12(13) 69-72
12(14) 73-76

On Eucythere declivis (Norman); by D. J. Horne & J. E. Whittaker
On Eucythere anglica Brady; by D. J. Horne & J. E. Whittaker
On Eucythere prava Brady & Robertson; by D. J. Horne & J. E.
Whittaker

On Piretia commasulcata Schallreuter sp. nov.; by R. E. L. Schallreuter
On Kroemmelbeinia valensis Schallreuter sp. nov. ; by R. E. L. Schallreuter
On Glyptocythere raasayensis Stevens sp. nov.; by G. W. Stevens
On Limnocythere shixiaensis (Wang); by Q. Wang

On Hiltermannicythere emaciata (Brady); by J. Athersuch & D. J. Horne
On Hiltermannicythere rubra (Muller); by J. Athersuch & D. J. Horne
On Brephocharieis complicata (Salter); by D. J. Siveter
On Henningsmoenia gunnari (Thorslund); by R. J. Orr
On Henningsmoenia costa Orr sp. nov.; by R. J. Orr
On Tyrrhenocythere amnicola (Sars); by T. R. Maness & R. L. Kaesler
On Paranotacy there (Paranotacy there) magnifica Lomax sp. nov.; by
A. Lomax

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