HARVARD UNIVERSITY. LIBRARY OF THE MUSEUM OF COMPARATIVE ZOOLOGY GIFT OF S a/vvs^vji-ej( vH -i>vuLXv< aJJjr^ "pUL<,€^vJ^L/^ ^I,l1ia. SYNOPSIS REPTILIUM; OR SHORT DESCRIPTIONS OF THE SPECIES OF REPTILES. BY JOHN EDWARD GRAY, F.Z.S., F.G.S., F.R.G.S., M.R.S.L., CORRRSPOXDING MEMBER OF THE ACADEMY OF NATURAL SCIENCES AND MACLURIAN LYCEUM OF PHILADELPHIA, OF THE IMPERIAL SOCIETY OF THE OBSERVERS OF NATURE AT MOSCOW, AND HONORARY MEMBER OF THE SOCIETY OF NATURAL HISTORY AT PARIS, &C. &C. PART [.— CATAPHRACTA. TORTOISES, CROCODILES, AND ENALtOSAURIANS. LONDON : PUBLISHED BY TREUTTEL, WURTZ, and Co. SOHO-SQUARE; G. B. SOWERBY, 156, REGENT-STREET; AND W. WOOD, 37, TAVISTOCK STREET, COVENT GARDEN. 1831, MCZ LIBRARY HAIVARD UHIVERSITY. C'- ''DRIDGE. MA USA DIGGENS AND JONES, LEICESTER STREET. TO THOMAS BELL, Esa., F. R. S., &c. THESE FEW PAGES ARE DEDICATED, AS A MARK OF RESPECT AND ESTEEM, BY THE AUTHOR. PREFACE. The Collections of Reptiles of the British Museum, the College of Surgeons, and Mr. Bell, have furnished the basis of this work. The two first of these collections con- tain many of the species which have been described by Dr. Shaw ; the College of Surgeons contains the Tortoises which were in the Leverian Museum ; but in the part now published I am most indebted to the kindness of Mr. Bell, whose collection of Tortoises far exceeds that of any museum in Europe, and whose liberality in allowing me the use of it I cannot too highly appreciate. It is to be hoped that his Monograph, for which he has collected them, and for which he has kept and had drawn alive more than two-thirds of the known species, will shortly appear. To render the collection of species as complete, and the synonyma as correct, as possible, every opportunity has been taken, during my visits to the continental museums, to exa- mine and take notes of the individual specimens which have been described by the various foreign authors who have written on this subject. Amongst the continental cabinets that of the Garden of Plants, of Paris, must be first men- tioned, if not from its intrinsic value, from the fact that most of the modern original writers on this branch of natural history have used it as their type collection ; witness the works of La Cepede, Latreille, and Daudin, among the French ; and Oppel, Oken, and Schweiger, among the Germans. It is much to be regretted that many of the specimens described by these authors should not have been more particularly ticketed, and that most of the species VI. PREFACE. collected by the later expeditions, are not yet added to the public parts of the collections. I have to thank Baron Cuvier, M. F. Cuvier, and M. Dumeril for their kindness in permitting me to examine these subjects, and more especially the former, whose attention to me on each of my visits to Paris, has been highly flattering to my feel- ings. Besides the national Museum at Paris, by the kind- ness of M. de Blainville, I have been enabled to examine the Museum of the Ecole de Medicine, containing several curious Reptiles, especially some from California. The Royal Collection at Berlin having been recently re-arranged, and the Royal Museum of Leyden and the Museum of the Senckenbergers Society of Francfort having been formed within these few years, the greater part of the specimens are quite fresh and in the most perfect condition, and their history is generally known and accurately marked upon them. These museums are the more valuable, as each of them is peculicu: for having the most complete collec- tions from certain parts of the world. That of Berlin excels in those of Buchara, of Mexico, and of the Brasils ; while the Leyden Museum is richest in the productions of the Dutch colonies, as the Islands of the Indian Archipelago, the Cape, and Surinam. That of Francfort contains the most complete collections of the animals of Egypt and the rest of Northern Africa that was ever brought together, having been entirely formed by the exertions of Dr. Riippell, during his travels in those countries, and extended by specimens received from other museums in exchange for his duplicates ; yet this monument of the industry of an individual must rank very high amongst the museums of Europe. After having laid before the scientific public the novelties which he has discovered. Dr. Riippell has again left Europe (at his own cost) to extend still further the empire of science. PREFACE. Yll. I hardly know how sufficiently to express my thanks to Herr Temminck and Herr Schlegell, of Leyden ; to Professor Lichtenstein and Herr Deppe, of Berlin ; to Drs. Cretzschmarr and Riippell, and Senator Von Heyden, of Francfort, for the courtesy and attention which they shewed me during my visits to the various museums under their direction ; indeed with such liberality that it would be impossible, however desirable, to imitate them in our more populous town. In each of these museums all the specimens w-ere intrusted to me, to describe, draw, or examine them, as might best suit my purpose, without any restraint, except that, at Leyden, Herr Temminck requested I would indicate in what Museum I had seen it, and the name under which it was there described, a rule which I hope I have most faith- fully kept. At Francfort some specimens w^ere even sent to my hotel, that they might be examined more at my leisure. I cannot here omit to mention the names of Sir James Mac Grigor, and Dr. Burnet for their kindness in allowing me to examine the Museum of Fort Pitt Chatham and of Haslar Hospital, and to Dr. Horsfield for the faciUties which he gave me of seeing the Reptiles in the IMuseum of the India House, and more especially of comparing and copying the drawings made under the superintendence of Dr. Hamil- ton in India. Besides those w^ho have assisted me with specimens, I cannot forget the kindnesses shewn me by Prince Massena Baron Ferussac, and M. Deshayes, at Paris ; Professor Reinwardt, at Leyden ; Professors Kunth and Ehrenberg, at Berhn ; and Herrn Oken, Fischer, Otto, Boie, and numerous other German, Swedish, and Danish naturalists at Hamburgh, in whose society I spent one of the happiest weeks of my life. The opportunity of examining the museums of the north of Europe not occurring till the body of the monograph was printed, I have been reduced to the necessity of adding the VIU. PREFACE. remarks and additional species as an appendix. To this appendix have also been added descriptions of some drawings of Chinese species sent by Mr. Reeves to General Hard- wicke, which will be shortly figured in a work on the zoology of that country now in the press ; and also the synonyma of Dr. Wagler's System der Amphibien, which has but lately arrived in London. 1 have to regret that after every enquiry and considerable delay on its account, I have not been able to procure the last part of the Annals of the Lyceum of New^ York, in which I understand M. Le Conte has given descriptions of the American species of Tortoises.* The two other parts of this work are in a considerable state of forw'ardness, and the next part, containing the Saurian animals, I hope to be able to print by the end of this year ; but should any circumstance prevent it, a complete index has been added to this part, so as to make it a distinct work. It is due to the reader that some apology should be made for the roughness of the etchings which are added to the work. They are, in fact, the first attempts of some ama- teurs, (partly spoiled by myself,) but for whose kindness it ■would have gone without any; and if they afford any facilities to the student they have fulfilled their office. They add nothing to its cost. British Museum ; Jan. 1831. * While correcting this proof Mr. Children has kindly put into my hands the above paper. I do not find amongst the specimens I have described any that accurately agrees with the Test- concinna, {"Emys reticularia. Say,") the Test, rubritentris, {" Emys serrata, Say,") or the Test, floridiana of this author ; his Test, insculpta is the Emys speciosa; his Test, geographica is the Emys Lesiieurii ; his Test, reticulata is the Emys reticularia ; and his Test, serrata is the Emys scripta of this work. Several of his synonyma from European works are erroneous, as may be seen by referring to the synonyma of this work, which, in the instances where they differed from him, have again been examined. SYNOPSIS REPTILIUM. REPTILIA. Animalia vertebrata, pulmonibus respirantia, sanguine rubro frigido, corde unilocular! et biaurito, cute squamosa. Ovipara vel ovovipara. This class, the Reptilia of Brongniart, is part of the Am- phibia of Linne, and answers to the Pholidota of Merrem. The skin is clothed with horny imbedded plates or with imbricate scales, covered by a thin, often deciduous epidermis. The bones of the skull are usually divided by sutures, and the neck is furnished with several vertebrae. The ribs are perfect; they often surround the body like a ring, and are sometimes dilated on the sides and united together, so as to include the body as it were in a bony case. The animals respire by cellular lungs, which are furnished with a windpipe, strengthened by cartilaginous rings. The heart has a 5ingle ventricle, divided into two or more cells, giving origin to two arteries, and receiving the cold red blood by two veins from two auricles. The penis is always distinct, and both it and the vagina of the female are often forked. The eggs are sometimes hatched in the body of the mo- ther ; a process which, under peculiar circumstances, as the want of a convenient place to deposit them, will take place even in those species which usually lay them. When they are laid, they are covered with hard shells, and are furnished with a thick internal lining. The young are like the mother, and do not undergo any transformations in their growth. The Reptilia have been divided by Cuvier and other naturalists into three or four orders, from the development B CAT \PHR ACTA. of their feet ; but after careful study, I have been induced to distribute them into two leading divisions, as proposed by M. Latreille, and to subdivide ihese into six orders. Sjjnopsis Ordinum et Familianim. Sect. I.— CATAPHRACTA-. lingua brevis adnata; or- gsna generationis simplicia ; ossa quadrata in cranio inclusa. Fam. 1. Chelonii : vertebrae dorsi costse et sternum ag- gli.tinata immobilia, fornicem dorsalem sternalemque for- mantia ; os edentulum rostratum. Fam. 2. Emydosauri: vertebrae dorsi et costae mobiles liberae; os dentatum; pedes digitati. ? Fam. 3. Ichthvosauki : vertebrae dorsi et costae mobiles liberae; os dentatum ; pedes pinnilormes. Sect. I[. SQUAMATA : lingua libera; organa genera- tionis duplicia ; ossa quadrata e cranio discreta. Fam. 4. Saurt : os non dilatabile ; cutis squamis variis inaequalibus tecta. Fam. 5. Ophiosauri : os non dilatabile ; cutis squamis aequalibus similibus tecta. Fam. 6. Ophidii : os dilatabile ; ossa mastoidea e cranio discreta ; pedes nulli. Sect. 1. CATAPHRACTA, Latr. Lingua brevis adnata ; organa generationis simplicia; anus longitudinalis vel rotundus ; ossa quadrata et pterygoidea in cranio affixa. The reptiles of this division, which was originally proposed by M. Latreille, are distinguished by their tongues being very short and affixed to the sides of the mouth, so that they can scarcely be exserted. The males have a single penis and the females a single vagina, like most other vertebrated animals. Their vent is either a longitudinal slit or a round- ish hole. The quadrate bone and the pterygoid processes are included in and form part of the skull. The limbs and tail are covered with large bony scales, and the body is either protected by two shields, formed by the union of the ver- tebrae, ribs, and sternum into a bony case, and covered with a cartilaginous or horny skin, or by longitudinal rows of bony plates inserted in the skin, and covered with an epidermis- CHELONIi; 3 which falls oiF in small pieces. Their lungs are enveloped by the thickened peritoneum, which performs the part and has the appearance of a diaphragm. They are all oviparous, and none of them poisonous ; but some, from their large size and carnivorous habits, are dangerous even to man. Order 1. CHELONII, Latr. Vertebrae dorsi costae et sternum agglutinata immobilia fornicem dorsalem stern alem que formantia, cute coriacea vel cartilaginea tecta; vertebras colli 8-9 mobiles ; claviculse utrinque duee ; pulmones abdomen intrantes ; os edentulum rostratum; tympana aperta cute tecta; vesica urinaria; penis simplex imperforatus ; anus rotundatus plicatus. Linneeus considered the tortoises as forming a genus under the name of Testudo ; Brongniart first separated them as a distinct group under the name of Cheloniens, which has been adopted by Cuvier. Latreille has latinized it to Chelonii ; and Oppel and Merrem use for this group the name of Testudi- nata. These animals are peculiar at first sight from their body being included in a bony case [Testa], leaving only the head, neck, limbs, and tail free. These parts are covered with a scaly riigulose or wrinkly skin, and are generally capable of being withdrawn into the cavity of the shell for protection. The head is sometnnes covered with regular shields. The jaws are usually furnished with horny cases, often toothed at the edges in the place of true teeth. The upper one overlaps the other like the lid of a box. They are rarely, as in the genus Chdys, protected with fleshy lips. The tongue is short, blunt, with long filiform papillae. The eyes have distinct eyeUds ; and the tympanum of the ear is visible and nearly superficial. The neck varies greatly in length ; it is usually withdrawn into the body of the shell when the animal is at rest, but in those families {Chel//dce) which have long necks, it is bent on the side of the body under the margin of the shell. The case {(esta) consists of two bony shields [scuta) united by the margin. The upper shield or carapace [scutum dursale) is formed by the eight pairs of ribs, being more or less dilated on the sides, united together, and adherent to the vertebra of the back by a toothed seam, so as to prevent their having the slightest motion one on the 4 CHELONII. other. The extent of the dilatation of the ribs greatly in- creases with the growth of the animal ; but in some groups, as Trio7iycidce and the Sea Turtles, they have a part of the distal or outer end not dilated, and always free. The lower shield sternum or plastron [scutum ventrale) is formed of four pairs of bones, and an anterior central one, more or less firmly united together, forming a disk, ana- logous to the breast-bone or sternum of other animals. These bones, in the land and most of the fresh -water tor- toises, are united together by toothed sutures into one disk ; but in the genera Ckelydra and in the families Trlonycidce and Chelodlnidce they form a ring, and leave a space filled with cartilage in the centre ; the two lateral pairs being large and often united together, and forming the body of the sternum. These are connected together behind by means an arch formed by the hinder pair : and in front, in the turtle, by the front pair forming an arch with the central bones, as a small pointed process on their inner edge. In the Triovycidcn and Chelydrce, the furcate central bones are united with the linear front pair placed on their outer sides. In the latter family these bones, especially the two lateral pairs, are often covered externally with rugose cal- losities. This sternum, in those animals where the bones are closely soldered together, is always attached to the bones of the margin by a dentated bony suture, except in the genus Cistuda, where it is united by a ligamentous junction, admitting of a slight motion ; while in the others, where the bones form a ring (except in the genus Chelydra), they are only united to the margin, when it exists, by some cartilaginous processes. The bones of which the sternum is composed are generally so attached to each other as not to allow of any motion ; but in a few of the land and fresh-water tortoises, the junction between the second and third, and sometimes between the third and fourth pairs of bones is by a ligamentous suture, which allows either or both extremities to move on the third pair as a door does on its post, and they are therefore called box tortoises. The margin of the shell is usually formed of a series of small bones similar to the cartilages of the ribs in mammalia ; uniting together the ends of the ribs of which the back of the shell is composed, and forming the bond of union between them and the sternum. In some, as among the Trionyches, the marginal bones are reduced to small isolated portions. CHELONII. placed on the cartilaginous margin opposite the ends of the ribs ; while in other species of this genus they are entirely wanting. The case is sometimes covered with a cartilaginous (in the Trionijches) or coriaceous {Sphargis) continuous skin ; but more usually with regular symmetrical horny plates, called sciitella. These shields, when the tortoises are first hatched, are distinct and covered with minute rugosities; they enlarge in size as the animal grows, by the addition of new layers of hcrny matter on their under side, extending beyond their former margin. These additions are generally marked by con- centric grooves, and the rugosity marking the original size of the shield is called the areola. The shields adapt them- selves to the shape of the bones beneath ; and when the bones, as is often the case in the land-tortoises, are raised into conical projections, they assume the same form. They are placed on the bony case in a regular manner, the centre of the back being covered with thirteen plates {sciitella disci) placed in three rows. The central row consists of five shields, which being placed over the vertebrae, are called vertebral plates ; and the side rows of each are designated, for the same reason, from the bones on which they are placed, the costal plates. These are surrounded by 11 or 12 or rarely 13 pairs of smaller plates, called the marginal shields {sciitella margina- lia), added to which there is often a small plate in the centre of the front edge called the nuchal plate, (sciitellum niichale.) In all the water tortoises the pair of plates over the tail, called the caudal shields {sciitella caiidalia), are separate from each other. But in the land tortoises, they are united into a single plate, which is often incurved and vaulted. The sternum is covered with six pairs of plates, which for the sake of preventing circumlocution in the description of the species, have been named according to their position ; thus the first pair are called gular {sciitella Qularia) ; the second humeral, {scut, hiivieralia) ; the third pectoral (scut, pectoraliaj ; the fourth abdominal, {scut. ahdominaUa) ; the fifth femoral, {scut.femoralia) ; and the sixth anal, {scut, analia.) Some- times, as in the genus Sternotherus, the gular pair are united into a single plate ; and in others, as in the Chelijdrce and Chelofiice, there is a small plate called the intergular shield, fscuf. ifitergularej placed between them. The symphysis by which the sternum is attached to the upper shell is mostly covered by the outer sides of the abdominal plates (as in the genus Sternotherus) or of the pectoral and abdominal plates ; b CHELONII. but in the turtle it is covered with four square peculiar shields; and there is placed at each end of this suture a small plate. The front one of these is called from its position the axillary shield, {scid. axillare), and the hinder the inguinal shield, {scut, inguinale.) The legs are usually thick and short ; but the feet offer considerable variation, conformable to the habits of the ani- mals. In the land tortoises the toes are very short giving the feet a clublike appearance, and they are armed with short blunt claws ; while in the fresh water tortoises the fingers are elongated and flexible, the space between them is more or less hlled up with a web, and they are all generally provided with sharp claws. In the genus Trionyx, however, as in the crocodiles, the two outer toes are clawless. It is in the marine turtle that the fingers gain the maximum of elonga- tion, and become united together by a covering of thick coria- ceous skin into a compressed fin, which is sometimes armed with short blunt claws. The tail is generally short and thick, but in one species it is long and crested on its back, like that of a crocodile. When the animal is at rest it is generally bent on one side under the edge of the upper shell. The heart is composed of two auricles and one double celled ventricle ; the blood of the body enters the right auricle, and that of the lungs the left ; but the two kinds ot blood are more or less completely mixed in passing into the ventricle. The lungs are very large, and placed in the same cavity as the other viscera ; but the peritoneum covering them is thickened and assumes the appearance of a diaphragm. The jaws ate destitute of true teeth, but are usually covered with a horny skin, often denticulated, the upper one covering the lower like the lid of a box. The tongue is short, and covered with long papillae ; the stomach is simple, and the intestines moderately long, and sometimes furnished with a coecum. The bladder is large. The penis of the male is simple, rather large, and channelled. The eggs are covered with a hard shell, and the young, when hatched, are marked with a large umbilical slit in the centre of the sternum, which soon disappears. The age of a specimen is best determined by the solidity and completeness of the ossification of the bones of the shell, and by the smallness or indistinctness of the areola compared with the size of the shields. It is very difficult to determine the sex of these animals externally. Perrault, and most naturalists since his time. CHELONII, have considered the concavity of the sternum as a mark of the male sex ; but this concavity is only found in the land tor- toises, and cannot therefore be used for the purpose which Sir E. Home supposes, (see Phil. Trans.) Mr. Sowerby [Zool. Jour.) observes that a specimen in which this cha- racter was distinctly marked, proved, on dissection, to be a gravid female. Dr. Harlan appears to consider the length of the, tail as a character for this purpose, but I have not had the opportunity of verifying his remark, which is not very definite. Synopsis Familiarum et Generum. Fam. I. Testudinid.e. — Pedes clavati, ungues 5-4 obtusi, scutella caudalia unita. 1. Testudo. — Testa dorsalis sternique solida, scutella stern i 12. 2. Chersina. — Testa dorsalis sternique solida, scutella sterni II. 3. Kimjxis. — Testa dorsahs postice mobilis, scutella sterni 12. 4. Pyxis. — Sternum antice mobile. Fam. II. Emyd^. — Pedes palmati, ungues 5-4 acuti, scutella sterni 11 vel 12. 5. Cistuda — Sternum latum cum testa symphysi carti- laginea connexum, scutella sterni 12. 6. Emys — Sternum latum cum testa symphysi ossea connexum, scutella sterni 12. 7. Kinoslernon. — Sternum latum cum testa symphysi ossea connexum, scutella sterni 12. 8. Clielydra. — Sternum angustum cruciforme. Fam. in. Chelyd^. — Pedes palmati, ungues 5-4 acuti, scutella sterni 13. 9. Sternotkerus. — Maxillae corneae, sternum antice mobile. 10. Chelodina. — Maxillae corneae, sternum solidum, scu- tellum intergulare inclusum. 11. Hydraspis. — Maxillae corneae, sternum solidum, scu- tellum intergulare marginale. 12. Chelys. — Maxillae labiis carnosis tectae. Fam. IV. TRiONYCiDiE. — Pedes palmati, ungues 3-3 acuti, testa cute molli tecta. 8 TESTUDO. 13. Triojiyx. — Sternum angustum. 14. Emycla. — Sternum latum, lateribus valvis munitum. Favi. V. Chelomad^. — Pedes pinniformes, compressi. 15. Sfihargis — Testa cute coriacea tecta. 16. Chelonia. — Testa scutellis tecta. Fam. 1. TESTUDINID^. Pedes clavati, ungues 5-4 obtusi, maxillae corneae. Testa solidissima scutis cornels tecta, scutellis marginalibus 24, caudali lato inflexo. The head i:^ globular, shielded ; the jaws are covered with horny sheaths ; the neck is thick and retractile. The shell is more or less globular, very solid and thick, the ribs being united together quite to the margin when the animals are adult, and covered with horny shields, which are generally grooved, and exhibit their areola for a considerable time. The sternum is broad and very solid, the bones of which it is formed filling up the centre ; and it is firmly attached to the upper shell by a bony suture as long as the abdo- minal and pectoral plates. The marginal plates are 24 or 26 in number, and have often a nuchal one in front; the hinder pair are always united together, and form a single broad incurved plate. The feet are short and clubbed, very like those of an elephant in shape, and armed with short blunt claws. The tail is thick and short. They are slow in their motions, eating vegetables and roots, and living in woods or fields in warm parts of the world. In a cold cli- mate, they burrow and sleep during the winter. Gen. 1. TESTUDO. (Tortoise.j Scutum dorsale sternaleque solida, scutella sterni 12, gularia distincta. The sternum of this genus is generally quite solid and des- titute of any movement, but Mr. Bell has informed me that he has observed in the female of T. Grceca and T. areola, that just before they are about to deposit their eggs, the hinder lobe of the sternum becomes loosened and capable of a slight mobility. TESTUDO. 9 1. Test. Indica, [Indicm Tortoise.) — Testa nigra, lateribus rotundatis ; scutello nuchali plerumque nuUo, pectoralibus brevissimis. Testudo Indica, Gmel. fide Perrault Acad. Sci. iii. t. cop. Schoepf t. 22, f. a. Shaw, Zool. t. 3. — Chersina retusa Merrem. — Testudo Indica Vosmceri, Sclioepf, t. 22, f. 13. (Mus. Vosmaer,) cop. Shaw Zool. t. 3, f. 2. Test, elephan- topus, Harlan Jour. Acad. N. S. Phil. v. t. 11. — Test ni- gra, Quo}" et Gaim. Freyc. Voy. t. 37, (v. Mus. Par.) Test. Californica, Quov Bui. Sci. (v. Mus. Par.) Cuv. Os. Fos. V. t. 11, f. 17,20,' (Anat.) Junior. Testa nigra margine lateral! angulato, areolis mag- nis. Test. Dussumieri, Schlogel MSS. (v. x\Ius. Leyd.) — Pet. Gaz. t. 76, f. 4. Var. scutello nuchae angusto, costalibus anterioribus con- vexis. Testudo Indica Dekay Jour. Acad. X. S. P. (v. Mus. D. Bell et Par.) Scutello nuchali brevissimo costalibus subconvexis caudali producto subreflexo. (v. Mus. Col. Surg.) /3. Testa antice compressa, scutellis vertebralibus 5 gib- bosis, (v. ]Mus. Par.) Y. Testa supra coUum complanata, scutello postico mar- ginali cum vertebralibus 5 gibbosis. Long. 28 — Test, gi- gantea Schwe'igev 17. Hab. in Brasilia, (Mus. Lisbon, fide Schw.) Habitat in India Orientali, Gefi. Hardwicke, Insula Mau- ritiana. Insula Aldebra, jSI. Dussumiere, Galapagos, D.ifor- lan, Seychelles, (v. v. Hort. Zool., et t. Mus. Brit., Col. Chir., Par., et D. Bell.) A variable species in form, but always known by its uni- form black colour ; the margin is keeled when young, but becomes rounded on the sides in the adult animal. 2. Test. Hercules, (Hercules Tortoise). — Testa subqua- drata oblonga depressa, lateribus inflexis ; scutellis sulcatis, nigris, areolis flavis, nuchali nullo ; stern o postice rotundato bilobo. Testudo Hercules, Spix t. 14. Junior. Test, carhonaria, Spix t. 16. Test, talulata. Far. Fitz. Test.planata, Gmel. Test. Schweigeri, Fitz, ined.? (non vidi.) /3. truncata. Testa solidissima ; ossibus scutellisque ele- vatis, scutellis convexis sulcatis, areolis parvis truncatis luteis. (Mus. D- Bell.) c 10 TESTUDO. Habitat in Brasilia, (v. v. Hort. Zool., et t. Mus. Brit., Col. Chirurg., D. Bell. &c.) A large species, sometimes 24 inches long, (v. Mus. Col. Surg.) The shields are sometimes worn smooth in the adult animal, and the sides of the shell are often very much con- tracted, the areola is small, and rarely exceeds three quarters of an inch in width. The animal is more or less red or yellow spotted, and the tail varies in length, probably in the different sexes ; it may prove only a variety of the next. 3. Test, tahulata, (Tahiilur Tortoise.) Testa oblonga depressa, scutellis nigro-fuscis sulcatis (demum levigatis), areolis latis pallide fuscis, scutello nuchali nullo, sterno postice acute bilobo. Test, tabuiutu. Wall). Test. Grceco Hermann, Test, tes- selata. Schn. Schoepf t. 12 f. 2. t. 13. Pr. Max. t, (viv.) Test. Lutraria, Gmel. T. Hecate, Brown Jam. n. 5. Test. Casado, Spix t. 17. (scutellis l3evibus).Wiedeman Arch. Zool. ii. (Anat). Junior. Testa fusca marginibus subdenticulatis. areolis latis, punctulatis. Test, sculpta, Spix, Bras. t. 18. — Seba, i. t. 80 f. 2, cop. Shaw Zool. iii. t. 8. — Max. t. (viv.) Schoepf t. 14. f. 1-3. Pullus. Test.denticulata, Lin. Schoepf t. 28. f.l. (Mus. Lin.) Hab. in Brasilia, (v.v. Hort, Bell, et nost., t. Mus. D.Bell.) This species differs from the former in being more depressed and less contracted on the sides, in the areolsc of the shields being larger, rarely less than one inch wide, and the posterior extremity of the sternum being acutely lobed. 4. Test. Schweigeri, (Schreiger's Tortoise). — Testa ob- longa depressa, scutellis pallide fuscis obscure fusco radiatim punctatis, nuchali nullo, areolis parvis centralibus, sterno luteo fusco radiato, antice acute emarginato. Hab. (v. t. Mus. Col. Surg.) The shell light, probably not full grown, the front margin nearly straight with a slight central indentation, the an- terior and posterior lateral margin bluntly scalloped on the edge, the caudal scale broad, edge refiexed, the discal shields with distinct concentric grooves becoming deeper near the edge, the areola very small, central, not more than 6 or 7 lines wide, the vertebral plates flattened, the anterior one convex, the gular plates small, the pectoral ones short and broad, the inguinal and axillary plates wanting except TESTUDO. 1 I on one side, where they are very small : length 8-i, breadth G^ inches. 5. Test. Polyphemus, {Gopher Tortoise.) — Testa oblonga depressa ; scutellis pallide fuscis sulcatis nuchali, lato qua- drato, sterno antice. dilatato; cauda obsoleta, unguibus de- pressis quadratis. Test. Polyphemus Bartram, 18. Daud. Emys Poly- phemus, Schw. Test, depressa, Lesueur Mss. (Mus. Par.) Guerin Icon. Rept. t. If. I. Test. Carolina, Le Conte Mss. (Mus. Par.) Hab. in America Septentrionali (v. t. Mus. Par.) 6. Test, marginata [Bell-shaped Tortoise.) Testa oblonga ventricosa, marginibus posticis horizontaliter explanatis, scu- tellis sulcatis nigris flavo variegatis, areolis luteis parvis, scu- tello nuchali elongate gracili. Test, maiginata Schoepf t. 11 and t. 12 f. 1. Test. Grail, Herman. Test. Grceca Lacep, t. 5, f. 2, cop. Enc. Meth t. 5, f. 4. Test, campanulata Walb. Test. Lu- traria. Gmel. Spix Cephal, t. 4. f. 12 — 15. Anat. (caret) Hab. in Brasilia, Egyptia, Fitzinger, (v. v. Hort. Zool. et t. Mus. D. Bell.) The Animal is black brown, with large scales on the limbs. The sternum is nicked in front and birid behind. The Test, sulcata of Shaw, Miller, Cym. Phys. t. 26. cop. Test, calcarata, Schw. Abhand. t. and Bechst. Lacep. t. Cher- sina calcarata, (part) Merrem, may be intended for either this or the following species, but it is not good for either. 7. Test, radiata. {Radiated Tortoise.) — Testa hemisphe- rica, scutellis planis sulcatis nigris flavoradiatis, areolis rufis, scutello nuchali nuUo, sterno antice attenuate sinuato trun- cato postice bifido. Test, radiata, Shaw, Grew Mus. t. 3, f. 1, cop. Shaw, Zool. iii. t. 2, (Mus. Brit.) Test. Coui. Daud. t. 26 f. 1, 2, (Mus. Par.) Test. Madagascarensis, Commerson (Mus. Par.) T. calcarata, (part) Merrem, f. 11. Cuv. Os. Fos. v. 193, t.l2, f. 40, Anat. i. 12, f. 1, f. 6, f. 13, f .16, 17, f. 27, 28, f. 35. )3. Senegalensis, Testa pallide lutea ; scutellis sulcatis, areolis parvis rugosis, scutello caudali inflexo, pedibus squa- mis magnis compressis tectis, ano bicalcarato — long. 15-|-", Habitat in Africa OccidentaH, Insula Madagascar (v. v. Hort. nost.,ett. Mus, Brit. D. Bell, &c.) j3. Senegal (v. t. Mus. Pax.) 12 TESTUDO. 8. Testudo Pardalis, {Leopard Tortoise.) — Testa subhe- mispherica lutea nigro maculata, scutellis sulcatis, costalibus areola prope marginem superiorem posita, nuchali nuUo. Testudo Pardalis, Bell Zool. Jour. iii. t. 25, Test, armula, Boie Mss. (v. Mus, Lej^d.) Test. Boei Wagler, t. (non vidi.) Junior. Test, marginata and Test. Grceca, var. (v. t. Mus. Par.) Pullus, Testa flava, scutellis nigro -marginatis, areolis annulo fusco maculisque duabus nigris ornatis. Test. Lipunc- tata, Cuv. Mss. (v. t. Mus. Par.) Habit, ad Cap. Bon. Spei, D. R. Brown, (v. v. Hort., Bell, et t. Mus. D. Bell ; Junior, Mus. Brit.) 9. Test, stellata, [Starred Tortoise.) — Testa ovato subglo- bosa, scutellis convexis sulcatis nigris luteo radiatis, areolis magnis luteis prope marginem posteriorem positis, scutello nuchali nullo, cauda brevi, capite pedibusque nigris luteo ma - culatis. Test, acti/iodes Bell Zool. Jour. iii. t. 14. Junior. Testa nigra, scutellis luteo radiatis, (Mus. nost.) Test, stellata, Schweiger. 2'est. elegans, Schoepf, t. 26, cop. Daud. t. 25, f. 1. (v. t. Mus. Paris.) Pullus Testa flava, commissuris scutellorum macula ovata nigra notatis. Test, elegans, Shaw, Seba Thes. t. 79, f. 3, cop. [Elegant tortoise,) Shaw Zool, iii. t. 6. Habitat in India orientali, Ceylonia, (v. v. Hort,, Bell, et t. Mus. nost. et D. Bell.) Coromandel. 10. Test, geometrica. (Geometrical Tortoise. j Testa ob- longo globosa, scutellis nigris gibbosis sulcatis luteo radiatis, areolis flavis truncatis, scutello nuchali angusto elongato. Test, geometrica. Lin. S. N. &c. Gotwald, f. 13 & 16, — Seba., t. 80. f. 3, & 8. Piso Bras. iii. t. 105, f. ].— Knorr. Del. ii. t. 52, f. 1.— Lacep. H. 1, t. 3, f. 2. cop. Shaw. Zool. iii. t. 2. and Enc. M. t. 6. f. 1. Schoepf. t. 10. cop. Daud t. 25. f. 1. Var. )3. Tentoria : Scutello nuchali brevissimo subnuUo, scutellis disci ossibusque conicis acutis elevatis. Test, tentoria. Bell Zool. Jour. iii. t. 13. Junior, sine scutellis testa flava. Test, luteola, Daud. t. 25. Schw. (v.t. Mus. Par.) Hab. ad Cap. Bon. Spei. /3. D. R. Brown, (Mus. Brit.) Madagascar. jD. Farquhar. v. v. TESTUDO. 13 11. Test. Grceca, {Greek Tortoise.) Testa oblongo-globosa scutellis subgibbis nigro flavoque variegatis ; caudali hamato obtuso, nuchali gracili elongate. Testiido Grceca, Lin. nee Lacep. Test. Carolina, Her- mann. Test, geometrica. Brunnich; Test. Hermanni. Gniel. Seba, t. 80, f. I, cop. Shaw, iii. t. 1. — Schoepf, t. 8. — Blu- menb. Abbild. N. H. t- 66.— Knorr. del. iii. t 51.— Mayer, Zeit. t. 28. ex.. Scutellis posticis explanatis Schoepf. t. 9. /3 "Sterno lobo postico mobili," Bell. MSS. Habitat, in Europa Meridionali et Africa Septentrionali, Mauritania, Ef/w. Long. 7.", Lat. 6.", (v. v. et t. Mus. Brit. Par. et D. Bell.) T. Zolkafa, Forst., said to have no nuchal shield, may be a variety of this species, but I have never seen it. The hinder lobe of the sternum of the female, as has been observed by INIr. Bell, is sometimes moveable. 12. Test, areolaia, [Areolated Tortoise.) Testa oblonga convexa, scutellis elevatis suicatis luteis, commissuris pro- funde exaratis, areolis fulvis impressis, scuteilo nuchali gracili. Test, areolata, Thunb, Nov. Act. Suec. Seba, 1 t. 80, f. 6.; Var. monstrosa scutellis vertebralibus 6, Gotwald, f. 14, Schoepf. t. 23.; scuteUis marginalibus, 26 (v. Mus. D. Bell) ; Hab. ad Cap. Bon. Spei., D. R. Brown, Mus. Brit. j3 pallida. Testa hemispherica, scutellis paliide virescen- tibus, areolis nigescentibus, scuteilo caudali lato iticurvo, sterni lobo postico mobili, (v. v. Hort. D. Bell.) Apt to vary in the number of dorsal and marginal plates. T.fasciata, Daud. of Ceylon, from Van Ernest's notes, is perhaps a monstrosity, with 18 dorsal, and 27 marginal plates. Animal pale brown, beak strongly toothed, head with small scales and a large occipital plate ; fore legs with lanceolate tubercular scales ; tail short, conical, thickly covered with small scales. I should have been inclined at first sight to have consi- dered the variety as a distinct species ; but it exactly agrees in the sculpture of its shields, and in the peculiar scaling of the animal with the type species. 13. Test, signata, [Marked Tortoise.) Testa humili, scu- tellis disci plauiusculis flavescentibus lineolis punctisque nigris 14 CHERSINA. adumbratis, areolis nigricantibus depressis, scutellis margin nalibus 26 leviter dentatis, nuchali angustissimo. Test, signata, Walb. Chelon., ii f. cop. Schoepf. t. 28 ; Test, denticulata, Var. Gmel. Habitat in Africa, (v. t. Mus. Bell, et Par.) Test. Cafra, Daud. (v. Mus. Par.) is probably a mon- strosity of this species, with 1 5 dorsal plates ; and Test. Juvencella. Daud. (v. t. Mus. Par.) seems only a variety with convex scutella ; but it is impossible to determine it with accuracy, as all the three specimens in the Paris Museum are in very bad condition. There are also two specimens in the same collection, marked as a variety, with 22 marginal plates, said to come from Carolina. Species Fossiles, Cuv. Os. Fos. v. 244. Testudo Lamanonii. Tortue des environs d'Aix, Cuv. Os. Fos. v. 244. Lamanon Jour. Phys. xvi. t. 3, cop. Cuv. Os. Fos. t. 13, f. 9— 11. Fossil, Gypsum Rock Aix, considered as fossil human heads ; breadth six, hei-ht seven inches. Test. Neraudii, Tortue de L' Isle de France. Cuv. Os. Fos. V. 248, t. 15, f. 17, (humerus) f. 18 (tibia). Fossil, in the volcanic deposits of the Isle of France. Testudo squamata, Daud., from Bontius n. 82, is certainly a species of Manis, of which lUiger has formed a genus, under the name of Pamphr actus. Gen. 2. CHERSINA, (Chersina.) Scutum dorsale sternaleque solida ; scutella sterni 11, gularia connata. This genus only differs from the other Land Tortoises, in the gular or anterior pair of sternal shields being united into a single plate, and produced in front of the shell. It agrees in this character with the genus Kinosternon, amongst the Terrapins, or Emydes. Merrem gives the above generic name to all the Land Tortoises, keeping that of Testudo for the family. I have restricted it to the species with the above character. ^/ 3 ^.^.i-Z^ .^i^^. (£. ^ / .^ /: .'/'-' H "Wilis v,';*i'w \i]i"' ':'\ ■^s*^ ■e^ /Z^^^r^-t^^^^ ^^^"^ '<^-/ I ^ ^ v§ A, KINIXYS. 15 1. Chersina angulata, [Bell's Chersina.) Testa oblonga ventricosa nigra supra collum triangulariter excisa ; scu- tellis disci rectilineis sulcatis, areolis flavis depressis. Testudo angulata, Dumeril, Schw. 7 (v. t. Mus. Par,) Test- Bellii., Gray Spic. Zool. t. 3, f. 4. (v. t. Mus. Brit.) Knorr. Del. Nat. iii., t. 52. f. 2. Habitat ad Cap. Bonae Spei, D. R. Brown, (v. v. Hort. nost. et t. Mus. Brit. D. Bell.) The gular shields produced and truncated in front, the nuchal plate very narrow, the front half of the marginal plates yellow, the hinder half black ; sternum yellow, black rayed. Howit, (Mus. Liverpool, t. ) appears to figure a variety with the gular plates much produced. Test, pnsilla, Lin. and Daud. (not Edw. nor Grew's figure) and con- sequently Test, miniata, Lacep. appears to belong to this species, which is sometimes reddish beneath when alive ; as also Test. Africana, Herm., and Test, tahulata Africana, of Schweiger. Gen. 3. KINIXYS, Bell. [Kinixys.) Testae dorsi pars posterior mobilis, parti anteriori sutura cariosa connexa. Sternum solidum scutellis 12. This genus is easily distinguished from the other Tortoises in the adult state, by the hinder part of the upper shell being capable of considerable motion, and separated from the central and front part by a transverse, irregular, carious, and somewhat ligamentous suture. In the young state, they difter but slightly from the common tortoises, as the dorsal suture is then scarcely to be observed. The suture can scarcely be said to be sub -elastic. Baron Cuvier (Reg. Anim. ii. 10 note,) appears to consider this structure as merely an accidental circumstance, he having seen only one specimen ; but I have verified the fact of its being common in the group on 9 specimens, four of which I have seen alive. 1. Kinixys Homeana, [Home's Kinixys.) — Testa obi ongo subquadrata fusca, margine posteriore retlexo, scutello verte- bral! 5 margine superior! prominente, nuchali elongato gracili. Test. Homeana, Gray, (v.t. Mus. Brit.) Kinixys Homeana, Bell, Lin. Trans, xv. t. 17, (v.t. Mus. Bell.) /3 Scutello nuchali lato, (v. t. Mus. Par.) 16 • pvxis. Hab. in Demerara, (v. v. Hort., D. Bell.) /3 Guadaloupe. Peculiar for the upper edge of the fifth vertebral plate being prominent, giving the back of the shell an angular ap- pearance, while in the next the central part is convex. 2. Kinixys erosa. [Eroded Khiixys.) — Testa oblonga fusca, margine posteriore reflexo denticulato, scutello vertebrali 5 equaliter convexo. nuchali nullo, (v. Mus. Brit.) Junior. Testa dorsi postice rotundata. Khiixys castanea. Bell. Lin. Trans, xvii. t. 18. Phllus. Testa depressa dorso immobili, margine denticu- lato. Testudo denticulato, Shaw, Zool. iii. t. 13, (v. Mus. Col. Chir.) et Test, erosa, Schw. (from Shaw.) Hab. (v. V. et t. Mus. Brit.) When young, the shell is depressed, with a strongly denti- culated margin all round, and the back is not moveable. The shell graaually increases in weight and solidity, and the dorsal suture becomes distinct, but it is not till the adult age that the centre of the fifth vertebral plate becomes convex, and slightly produced. Gen. 4. PYXIS. Bell. (Box Tortoise.) Testa dorsi solida, sterni lobus anterior mobilis, sutura inter scutella humeralia et pectoralia. The front lobe of the sternum is moveable, like the Box Terrapins (Kinosternon) ; but here the suture is placed be- tween the humeral and pectoral sternal shields, while in those tortoises it is always placed between the pectoral and abdo- minal ones. This shell is distinguished at first sight from the last-named by its greater convexity and sohdity. The sternum has also twelve plates, while the other box tortoises have either eleven [Kinosternon) or thirteen [Sternotherus.) The feet are clavate, hke those of the other animals of this family. ]. Pyxis aranoides, [Radiated lox tortoise.) — Testa hemis- pherica, luteo nigroque variegata. Pyxis aranoides. Bell, Lin. Trans, xv. t. 16. Hab. (v. t. Mus. D. Bell.) Length six, breadth four inches ; varies greatly in the in- tensity of its colour in the three specimens which are in Mr. Bell's collection. CISTUDA. 17 Fam. IL EMYD.^ (or Emydidcc.) Pedes palmati, digitis distinctis, unguibus 5-4 longe acu- minatis, nares pervii, mandibulee corneee. Testa depressa, scutellis cornels tecta ; scutella marginalia 23-27, caudalia distincta, sterni 11 vel 12. The head of these animals is rather depressed and shielded; the neck is contractile into the body of the shell. The feet are depressed and expanded, with separate toes united toge- ther by a distinct web, and armed with sharp and rather long claws ; the tail is usually more or less exposed. The shell is generally depressed and solid, with a distinct bony margin, and covered with horny plates ; the dorsal plates 13, with twelve pairs of marginal plates, the caudal pair being separated by a distinct suture, and with a narrow nuchal one. The sterno -costal commissure is generally long, and usually fur- nished with a distinct and rather large axillary and inguinal plate. The sternum has eleven or twelve shields, the gular pair being sometimes united, but never, as in the next family, having an extra plate between them. The vertebrae of the neck bend in a perpendicular bow, and the bones of the pelvis are only attached to the vertebrae, and not to the bones of the sternum. They hve in ponds and ditches, in warm and temperate climates ; are rapid in their motions ; eat mollusca, worms, insects and carrion, and only take their food while in water : the eggs are oval and white. I have observed 36 species ; 18 of which have come from America, and 14 from the old world. The external characters to distinguish the sexes of these animals have not been satisfactorily made out. Dr. Harlan observes, that the females of some species are more keeled than the males. The concavity of the sternum, which only occurs in the first genus, does not appear to be peculiar to the males; as Mr. Sowerby, Zool. Jour. ii. 485, says that he has found eggs in the body of a specimen of Cistuda Carolina which had a very concave sternum. Gen. 1. CISTUDA. [Box Terrapin.) Sternum latum, extremitatibus rotundatis, symphysi liga- mentosa testae dorsali adnexum, in duos lobos per suturam transversam medio divisum, scutella 12. D 18 CTSTUDA. The form of the sternum of this genus is very pecuhar, for it is divided across by a cartilaginous suture between the pectoral and abdominal plates; and being only connected to the back shell by a cartilaginous suture, both the anterior and posterior lobes are moveable on the same axis, so as to be capable of entirely closing the cavity of the upper shell, and concealing the inclosed animal from view and danger. The axillary and inguinal shields are small and obscured by the margin ; the marginal plates vary in number from 23 to 27, including a distinct nuchal one. This group consists of four species, each found in different parts of the world. The European species, by its depressed form, slightly lobed sternum, and wider symphysis, appears to unite them to the true Emydes. This genus forms part of the genus Chtuda of Fleming and Say, and part of Terrapene and Sternotheriis of Mr. Bell. 1. Cistiida Carolina, [ylmerican Box Terrapin). — Testa subhemispherica fusca luteo-variegataleviter carinata scutellis subsulcatis, marginalibus 23 — 27. Animal nigrum luteo vel aurantio variesjatum. Test. Carolina, Lin. Daud. Test, brevicaudata, Lacep, (v. Mus. Par.) Test, invar cerata striata, Bonat. Terrapene Ca- rolina, Bell. (v. Mus. Bell.) Test, clausn, Daud. Cistiida clausa. Say. Test, virgulata Daud. ii. t. 23, (v. Mus. Par.) Test, incarcerata, Bonat. Test, carinata, Lin. Emi/s diihia, Schw. (v. Mus. Par.) Terrapene guttata. Bell, (v. Mus. '&^\\.) Emys Schneideri,'&c\\yN. Edw. t, 20.5, cop. Shaw Zool. iii. t. 7, and Seligman vi. t. 100, and Bechst. Lacep. t. f. — Schoepf. t. 7. — Bloch, Berl. Naturf, t. 1, f, 1, 2. — Grew. t. 3< f. 2. Junior, Testa fusca pallide nebulosa, Terrapene nehulosa. Bell, (v. Mus. Bell. Col. Surg.) /3. Scutello nuchali nullo (v. Mus. Bell.) Habitat in America Boreali. {v. v. Hort. nost, et Bell.) T have examined and kept several living specimens, and carefully compared the original specimens of most of the above synonyma, and have not been able to separate them from one another ; I beheve that Mr. Bell has himself given up the idea of this species being distinct, since he has been enabled to examine more living specimens. The young are often brown and pale clouded, but the pale spots are placed in the same situations as the yellow spots in the adult spe- cimen, and gradually pass into them. CISTt'DA. 19 2. Cistuda Amboinensis, (Amhoina Box Terrapin.) — Testa hemispherica obscure tricarinatanigrescente, margine lato ex- panse, scutello nuchali lineari, stenio luteo nigroque variegato; animal nigro luteo variegatum, capitis lateribus fasciis dua- bus luteis ornatis. Test. Amboinensis, Riche, Daud. Einys Amboinensis, Sch. Test, couro, Lesch. (v. Mus Par.) Emys couro, Schw. Ter- rapene bicolor. Bell Zool. Jour. ii. t. 14. Junior. Testa acute tricarinata, marginibus horizontaliter expansis, (v. Mus. Brit.) Cisluda Amboinensis, nob. Illust. Ind. Zool t. f. 2. /3. Leveriana, Testa ovata subcompressa. long. 74, lat. b^, pol. (v. t. Mus. Col. Surg.) Habit, in Java et Penang (v. v. et v. t. Mus. Brit., Par., et D. Bell.) Not in America. 3. Cistuda trifasciata. (Three banded Box Terrapin.) — Testa ovata, carinata, obscure lutea nigro rubroque variegata, fasciis tribus dorsalibus ornata, sterno postice bifido ; capite luteo lateribus fasciis duabus fuscis notatis, cauda longa. Sternotherus trifasciatus, Bell Zool, Jour. iii. t. 13, (v. t. Mus. Bell.) Hab. Differs from the other species by its ovate shape and the three dorsal streaks, the two bands on the side of the head unite over the occiput. 4. Cistuda Europcea. [European Box Terrapin.) — Testa depressa ovata fusca radiatim luteo punctata, sterno postice sub-bifido ; capite corporeque nigro luteo punctatis, cauda longa. Testudo Europcea, Sch^v. Schoepf, Test. t. 1, cop. Sturm, t. 2, 3, 4. Test.jiava, Lacep, t. 6. Test, punctata, Got- wald, f. 12. Meyer t. 29. Test, orbicularis, Lin. S. N. Wulf. Test. Lutraria, Herm. Marsigl. Dan. iv. t. 33, 34. Emys Lutraria, Merrem. Knorr, Delic. ii. t. 32, f. 5. Ter- rapene Europea, Bell ; Speckled Tortoise, Shaw, Cuv. Os, Fos. v. t. 12, f. 2, f. 7, f. 12, f. 18, 19, f. 27, 28 f. 2>Q.— Testa artijiciose polita. Test, meleagris, Shaw, Nat. t, 104, (v. t. Mus. Col. Surg.) Test. Lutraria, Lacep, (v. Mus. Par.) Junior. Testa depressa scutellis areohs magnis, marginibus luteo radiatis. Test, pulchella, Schoepf t. 26, Emys pul- chella, Merrem, (v. Mus. Par. et Brit.) Habit, in Europa Australi. (v. v. Hort. Bell et nost.) 20 EMYS. More depressed than the other species. M.OppelJn (1811) remarked the fact of the sternum being moveable, and it has since been observed by Schweiger, Bojanus, and Mr. Bell. For the Anatomy of this animal consult Bojanus's beautiful and accurate Work. Gen. 2. EMYS, (Terrapin). Sternum dilatatum solidum, antice truncatum postice bifi- dum, persymphysinosseamcum testa conjunctum, scutella 12. This genus only contains part of the Emijdes of former authors, it being limited to those Terrapins which have the sternum united to the back shell by a bony symphysis, and covered with six pairs of bony shields. The symphysis is generally about \ the length of the sternum, covered by the outer sides of the pectoral and abdominal shields, and fur- nished with small and partly exposed axillary and inguinal plates; the margin is always covered with 12 pairs of plates and a small nuchal one. A. Testa margine acute dentato, sterni laterilus rotundatis. Orbis Veteris Incolte. 1. Emys spinnsa, {Spinous Terrapin.) — Testa (pulli) suborbiculari depressa pallide fusca dorso obtuse carinato, margine explanato acute dentato, areolis punctatis spinis centralibus armatis, sterno pallide fusco brunneo radiato. Eniys spinosa, Bell. Mss. (v. Mus. Nost.) Gray, lUust. Ind. Zool. ii. t. f. 1. Habit, apud Penang. Capt. Hay, A large species only known from two specimens in a very young state, 4^ inches long and broad. The back is fur- nished with a broad flattened central keel, and the margin is deeply dentated and serrated all round. 2. Ejnys Dhor, {Dhor Terrapin.) — Testa (pulli) subor- biculari depressa, fusca nigro punctata, dorso obtuse cari- nato, margine explanato postice acute dentato, areolis mag- nis punctatis ; animal nigrescens lateribus colli gutturisque aurato-lineatis, pedibus aurantio maculatis. Emys Dhor. Gray, lUust. Ind. Zool t. , (v. Mus. Brit.) Emys Hasselti, Boie Mss. (v. Mus. Leyd.) Habitat in Bengal, called Dhor and Thum, General Hard- wicke, Java, Van Hasselt. (v. v. Hort. Bell et nost.) Like the former, only known from three young specimens. k^ 'S l^^Z^t.^-.^^ ^yi-^ -iT^ ''^ /^/. ^ EMYS. 21 one of which, given me by Mr. Bell, I kept alive for some time. The margin of the shell is much expanded, and recurved on the sides, and sharply but simply toothed be- hind. The sternum is pale blackish speckled, and the shields finely blackish rayed, rounded and crenated in front. The keel and the large size of the areola of the specimens would doubtless disappear as the animal grew older. 3. Emys Spengleri, {Spengler's Terrapin.) — Testa oblonga depressapallide fusca, tricarinata carinis continuis distantibus, margine postico profunde serrato, scutellis vertebralibus qua- dratis. Tei>iudo Spengleri, Walb. Berl. Naturf v. t. 3. Emys Spengleri Schw. 32. Testudo serrata Shaw, Zool. t. 9. (v. Mus. Col. Surg.) Test, serpentina, var. Latr. Test, tri- carinata, Bory St. Vincent Atlas, t. 37, f. 1. (bona.) Habit, in Africa, M. Bory St. Vincent. The shell pale brown, the hinder dentations are simple and acute, and the sides of the sternum are keeled, the tail is long and exserted. 4. Emys crassicollis, {Thick-necked Terrapin.) Testa ovata oblonga, leviter convexa, nigra, obscure tricarinata carinis approximatis. marginibus lateralibus revolutis, postico dilatato profunde serrato. Emys crassicollis, Bell, MSS. (v. t. Mus. Bell) ; lUust. Ind. Zool. t. . Junior, Testa nigra, tricarinata, (v. t. Mus. Bell.) Habit, in Sumatra, (Mus. Brit, et Bell.) Shell with the side margins narrow and revolute, the ser- ratures of the hinder margins entire, the dorsal keels near together, the first vertebral plate long narrow hexagonal, the sternum blackish, keeled on the sides, truncated before, and with a small roundish sinuosity behind; tail short, head and neck very thick, blackish. 6. Emys Hamiltonii, (Dr. Hamilton's Terrapin.) Testa oblonga, tricarinata, nigra luteo radiata, postice subdentata, sterno lateraliter carinato ; capite corporeque nigro-luteo maculatis. Emys guttata. Gray, lilust. Ind. Zool. i. t. ., non Schw, Habitat in India, Dr. Hamilton, Mus. Ind. From Dr. Hamilton's drawing, which is evidently from a young specimen, it is black, and the shields broadly yellow- ■ 22 EMYS. rayed ; the first vertebral plate is nearly square, and the second and third broad six-sided, the fourth long six-sided. The sternum is truncated before and nicked behind ; the tail short, length two and three-quarters, and breadth tw o inches. This may be the young of the preceding, as the vertebral plates alter their form by age ; but both the animal and shell are yellow-spotted, and the shell is less toothed behind, ■which is not the case with a young dry specimen of E. cras- sicoUis in Mr. Bell's Collection. 6. Emys Tlmrjii, (Tlutrgy Terrapin.^ Testa oblonga sub-convexa nigra, margine lutescente postice subdentato, sternonigro,lateribus subcarinatis, scutellis vertebralibusprimo quadrato, secundo et tertio late hexagonalibus ; capite nigres- cente, superciliis mentoque lineolis luteis ornatis, pedibus viridibus luteo maculatis. Testudo Thnrjii, Hamilton, Icon. ined. (v. Mus. Ind.) cop. Emijs Tlmrjii, n. Illust. Ind. Zool. t. Habitat in India, Dr. Hamilton and General Hardwicke. Length six, breadth four inches ; the head blackish, with a yellow band from the nostril over the eyes, and another from the chin to the back of the throat. Is this Test, mela- tiocephala Daud. from Van Ernest's notes? This may be the adult of E. crassicollis, but Mr. Bell's stuffed specimen of that species has no appearance of any yellow on the head. It was doubtless one of these species that is represented in the Indian Sculpture, formed out of Jade, formerly in possession of Dr. Flemming, and lately presented to the British Museum. 7.? Einys ocidifera, (Eyehearing Terrapin.) Testa puUi hemispherica depressa postice dentata, scutellis annulis nigris, suturalibus connexis, sterno luteo, maculis lineisque fuscis ornato. Test, oculij'era, Kuhl. Beytr. Shell hemispherical, strongly toothed behind, shields ele- vated, furrowed, granulate, grooved ; each two connected longitudinally by a largish round black spot, and horizon- tally by a largish black circle, consequently each displaying laterally two half eyes, and anteriorly and posteriorly two half spots, except in the three last ones, which show only the three half spots ; margin with a spot on each suture, the point of which corresponds with the spot on the costal shields, sternum yellow, with dark brown spots and streaks. ^» N £^^ Ji^iSC A 2^ EMYS. 23 Hab. Mus. Bed. (nonvidi.) only a very young specimen, B. Testa margine integro, sterni laterihus suhcarinatis, Orbis Veteris Incolae. 8. Emijs tecta, (Tented Terrapin.) — Testa ovato-oblonga solida, olivacea, dorso elevate angulariter compresso, scu- tellis vertebralibus elongate hexangulatis, 1, 2, et 3 cari- natis postice productis sub-tuberculatis, margine integerrimo. Emys tecta, Bell, MSS. (v. t. Mus. Bell.) Illust. Ind. Zool. t, Testudo Dura, et T. Katuya, Hamilton, MSS. (v. Icon. Mus. Ind.) Junior. Sterno rubro nigroque variegato. Testudo Pang- shure, et Test. Khagraskata ; Hamilton, MSS. (v. Icon. Mus. Ind.) Habitat in India, Dr. Hamilton, Gen. Hardwicke. The shell is very solid, with a strong internal rib on each side, just behind the back of the front opening ; the first, second, and especially the third vertebral plates with a keel, which is bright orange when alive, and produced into a peak at the end of each plate. The sternum is high and keeled on the sides, dark brown, pale -edged, and in the young shells variegated with black and red ; the head is greenish, red-spotted, and the neck yellow -lined. 9. Emys Vineata, {Lined Terrapin.) Testa oblonga olivacea leviter tuberculata-carinata, margine postico dilatato, scutellis vertebralibus primo quadrate, reliquis lenge hexagonalibus, sterno lutescente ; capite cinereo coeru- lescente, mento genisque lutee variegatis, superciliis nucha- que coccineo lineatis, cauda longa. Emys lineata, (v. Icon. Hard.) cop. Illust, Ind. Zool. t. Habitat in India, General Hardwicke. The first vertebral plate is nearly square, the rest are narrow hexagonal, and the sternum is keeled on the sides ; but the animal, in these plain species, affords the best cha- racter : it is greenish, with a bluish grey tint on the head, and marbled w-ith yellow on the chin and cheeks, with a bright orange line over each eyebrow, and six or eight on the back of the neck ; the chin has two yellow spots, and the tail is long. 10. Eynys Batagur, (Batagnr Terrapin.) Testa subor- biculari depressa leviter carinata pallide olivacea, margine 24 EMY«. integerrimo postice dilatato, scutellis vertebralibus prinio quadrato, secundo et tertio latis hexagonalibus ; corpore cinereo, mento labiisque pallide lutescentibus ; cauda bievi. Testudo Balagur. Hamilton, Icon. (v. Mus. Ind.) cop. Eynys Batagur ; lUust. Tnd. Zool. t. |3. Boska. Testa oblongo ovata, sterni lateribus subro- tundatis, long. 5, lat. 4 pol. Testudo Baska, Hamilton, Icon, ined. (v. Mus. Ind.) cop. Emi/s Baska, Illust. Ind. Zool. t. Habit, in India, Df. Hamilton. The hinder and more especially the hinder part of the lateral, margin dilated ; the length is four and a half, and the breadth four, inches. II. Emys villa ar is, [Common Terrapin.) Testa ovato- depressa obscure olivaceo viridi rugulosa nigro punctata, sterno piano nigrescente lateribus obtuse carinatis ; capite pedibus caudaque nigrescentibus aurantio-lineatis. Emys Lutraria,^c\i\v.'^. Bell. MSS. (v. Mus. Bell.) Test. Lutraria, Lin. ? ? ? Baud. ? Lacep. Ovip. t. 4. ? Jan. Testa ovata, plus minusve-tuberculato tricari- nata olivacea, maculis aurantiis nigro circumdatis ornata ; sterno nigro ; capite artubusque lineis aurantiis nigrisque variis, cauda longa. Emys Sigritzii. Michaux, MSS. (v. Mus. Leyd. Mus. Brit.) Habit, in Europa Australi? (v. v. Hort. Bell, nost. et t. Mus. Brit.,&c.) A common species, of which I have seen more than twenty living. The adult shell has a rounded keelless back, of a dirty green colour, and covered with irregular lurid blackish spots. The young shell has three, more or less distinctly elevated, equi-distant, irregular, tubercular, and crumpled keels ; the head is olive, and the cheeks, neck, and feet marked with black -edged orange lines. C. Testa margine integro, sterni laterihusrotundatis. Orbis Novae Incolae. 12. Emys scahra. {Rough Terrapin.) — Testa ovato ob- longa plana acute carinata, scutellis (" luteo nigroque va- riegatis" (Schw.) in juniori aetate albidis ; capite lineis su- perciliaribus luteis, postice furcatis. Test, scahra, Lin. Lacep. t. 8, f. 2. (v. Mus. Par.) cop. Enc. M. t. 6, f. 2. T. verrucosa, Walb. Test, orbicularis v. Grael. Test. Amhoinensis, Lin. e. Seba, i. t. G, f. 1 2. Emys t-^/. 4. t^ ^'^■' '^'/^-IL EMYS. 25 dorsata, Schoepf, Schw. n. 3. Cuv. Os. Fos. v. t. 12, f. 45, Anat. Emijs dorsal is, Spix. t. 9, f. I, 2. Habitat in America Meridionali. I have only seen a young specimen. Spix's figure is also from a young specimen , it only dift'ers from the spe- cimen next described in the spot on the side of the occiput not being joined to the superciliary band. He describes the legs as yellow, black-lined. 13. Emys punctularia, [Dotted Terrapin.) — Testa ob- longa convexa fusca obscure carinata, scutellis margine ni- grescente circumdatis, vertebralibus complanatis costalibus declivibus nuchali brevissimo angusto ; occipite nasoque ma- culis duabus, superciliisque lineis aurantiis ornatis, coUo nigricante luteo maculato. Testudo punctularia, Daud. (v. Mus. Par.) Emys piinc- tularia, Schw. n. 19. Habitat in America Meridionah. Cayenne. Richard, (Mus. D. Bell, V. V.) The tail is short and the head is blackish, with an orange spot over each nostril, a band over each eye-brow, and a large spot on each side the occiput. 14. Emys marmorea. {Marbled Terrapin) — Testa ovata depressa virescenti flavo nigroque variegata, scutellis flavo- mareinatis, nuchali subbrevi tetrao;ono ; animal olivaceo brun- neoque virescens subtus flavicans, capite lineolis fiavis in- signi, Cauda longa tenui. Emys marmorea, Spix t. 10. Habitat in Brasilia. Length of shell 4| — breadth, 3-i inches. Caup has referred this species to Emys picta, but neither the figure nor description agrees with that species. As I have not seen it, I have given the essential parts of M. Spix's description. The figure is most like Emys decussata. but the shields are smooth. 1 5. Emys Muhlenhergi, {Muhlenberg's Terrapin.) — Testa ovali-oblonga humili obscure carinata, lateribus contractis, scutellis leviter sulcatis castaneis luteo variegatis, nuchali lineari ; capite nigrescente occipite maculis 2 fulvis ornato. Testudo Muhlenbergii, Schoepf, t. 31. Einys Muhlen- bergii, Schw. n. 30. Emys bipunctata, Say, Jour. A.N. S. P. Emys fusra, Lesuenr Mss. (v. Mus. Par.) E 26 EMYS. Hab. in America Boreali. Length of shell 4, breadth 3 inches, (v. t. Mus. Par.) The shields are concentrically and radiately striated, the head is blackish varied with yellow, with two large irregular fulvous spots on the sides of the occiput. 16. Emys guttata, {Spotted Terrapin.) Testa ovata plana postice dilatata, scutellis laevibus nigro-fuscis, guttis flavis irro- ratis, nuchali lineari ; capite luteo-maculato. Testudo guttata, Schn. Berl. Gesells. Nat. Fr. iv. t.4 — Gotwald, fri5. Test, punctata, Bosc. Schoepf, t. 5. Test, anomjma, Schn. Seba, i, t. 80, f. 7, cop. Spotted Tortoise, Shaw, t. 10. Emys guttata, Schw. /3. Scutellis nigris, macuhs luteis centralibus notatis, (v. Mus. Bell.) Junior. Testa nigra obtuse carinata, scutellis nigris maculis luteis centralibus ornatis, (v. t. Mus. Bell.) Hab. in America Boreali. (v. t. Mus. Bell, Par.) 17. Emys picta, {Painted Terrapin.) Testa oblonga subcon- vexa, scutellis tenuibus levissimis olivaceo fuscis lato flavo marginatis ; capite gulaque nigris luteo-lineatis, cauda longa gracili. Testudo picta, Herm. Schoepf, t. 4, cop. Painted Tortoise, Shaw. Zool. t. 10. Emyspicta, Schw. n. 22, (v. t. Mus. Par.) Jun. Testa cinerea scutellis albido marginatis, (v. Mus, Brit.) Test, cinerea, Brown lUust. t. 48, (v. t. Mus. Brit.) cop. Cinereous Tortoise, Shaw, Zool. t. Schoepf. t. 3, f.2, 3, and Bechst. Lacep. t. — E?nys cinerea, Schw. n, 23. Hab. in America Boreali, (v. t. Mus. Brit., D. Bell, Par.) Caup has apparently without sufficient evidence referred Emys marmorata, Spix, t. 10. to this species. 18. Emys speciosa, {Specious Terrapin.) — Testa oblonga plana antice carinata, margine postico lateraliter subreflexo, scutellis radiatim et concentrice sulcatis, flavo nigroque minute punctatis, areolis parvis, nuchali gracili elongato, sterni lutei areolis nigris margine posteriori scutellorum impositis. Emys inscripta, Le Conte Mss. (fide t. Mus. Par.) Emys seeder a. Say, non Lin. /3. levigata, Scutellis levigatis radiis nigris flavisque ornatis. Emys speciosa. Bell, Mss. (v. t. Mus. Bell.) Hab. in America Boreali, New Jersey, (v. Mus. Bell et Par.) EMYS. 27 The shields elegantly and minutely dotted with black and yellow, and in the smooth variety beautifully black and yellow raved. Those of the sternum have a square black areola placed on their hinder margin According to Mr. Say, the skin of the anim.al is above of a uniformly dark greenish ijrown and beneath fulvous. 19. Emys conceiitr'ica, {Concentric Terrapin.) — Testa ovali subconvexa leviter tuberculato carinata, scutell^is ossibusque subtus pallide luteis, zonis fuscis concentricis ; cute albido nigro puiictato. Testudo centrata, Bosc. D. H.Nat, xxii. 264. (v. t. Mus. Par.) Test. Terrapin, Schoepf, t. 15. — Latr. Rept. t. 6, f. 2. Test palustris, Gmel. Eimjs centrata, Schw. a. Uvida. Testa livida obscure annulata. Emys livida, Bell, Mss. (v. t. ]Mus. Bell.) /3. polita. Testa margine revoluta, scutellis nigris politis profunde sulcatis, sterno lutescente macuhs magnis nigris notato, (v. t. Mus. Col. Surg, et D. Bell.) Junior. Testa carinis continuis, scutellis sterni nigro mar- ginatis ; cute albido cinereo nigro maculato, capite colloque lineis nigris notatis. Habit, in America Boreali. (v. v. Hort. Bell.) The head of the living animal is very broad and depressed, like the Hydraspes, and the neck thick ; they are slate-co- loured, black-speckled. In the young specimens the skin is bluish and black-speckled ; the forehead is marked with concentric black lines, and there are three curved black lines over each ear , the neck is also marked with short black strokes. The polished variety appears very different, but I have seen specimens which unite it with the common state of the species. Cuvier (R. A. ii. 11 note,) indicates an Emys concentrica of Le Conte, as distinct, but I do not find it described. 20. Emys reticularia, {Reticulated Terrapin.) Testa oblonga olivaceo fusca pallide reticulata, postice Integra, scutellis, fasciis radiatis annulisque pallidis nigro marginatis areolas circumdantibus, vertebralibus Imo. quadrato, reliquisquadrato- hexasionalibus, marginalibus subtus maculis nigris suturalibus 3 vel 4 lateralibus rotundatis aliisque posticis linearibus, asillaribus inguinalibusque macula ocellata ornatis ; sterno angusto lateribus subcarinatis. Test, reticularia, Latr. Test, reticulata, Bosc, Daud, t. 22, f. 3, (tig. accurata) Emys reticulata, Say. Emys geo- graphica, Mus. Bell, (v. Mus, Bell.) 28 EMYS. Hab, in America Boreali, Daud. Say, (v. t. Mus. D. Bell.) There are three or four round black spots placed in the sutures of the under sides of the lateral marginal plates, some linear spots in the sutures of the hinder ones, and also an oval-eyed spot on each end of the sterno-costal symphysis. 21. Emysvittata, [Banded-Necked Terrapin.) — Testaovato oblonga sub-convexa levi, obscure carinata, postice duplici- dentata, fusca lineolis inequalibus luteis variegata, lineis cen- tralibus sub-annulatis, lateralibus transversis, marginibus supra maculis quadratis luteo nigroque annulatis, subtus maculis ocellatis utrinque suturis impositis, sterno piano lutescente, scutellis axillaribus inguinalibusque lato nigro annulatis ; capite pedibusque luteo lineatis. Hab. in America Boreali. (Mus. Brit.) This is somewhat like Emysconcinna, Le Conte Mss. Cuv. R. A. Guerin Iconogr. t. f. which is also T. geometrica of Lesueur according to the same authority. Indeed the figure may be intended for this species. I did not observe any specimen under that name in the Paris Museum. The British Museum specimen is not quite full grown and has lost great part of its colour. The first vertebral plate is nearly square, and the others broadly hexagonal. 22. Kmys decnssala, [Decussated Terrapin.) Testa oblonga pallide fusca obtuse carinata postice subdentata, subtus lutescente, maculis subocellatis scutellis axillaribus inguinali- bus suturque marginalium impositis ; scutellis rugulosis irregulariterradiatim sulcatis ; animal virescens, genis gulaque obscure pallide lineatis. Emys decussata. Bell, MSS. (v. v. Mus. Bell.) Test. serrata. Daud. (non tig.) Emys Serrata. Scbw. ? var. Potter, dicta Harlan ? Junior. Testa obscure fusco variegata ; sterno fascia cen- trali irregulari viridi nigro marginata, annulisque duobus nigris antice ornato. Hab. in America Boreah, (v. v. Hort. nost. et t. Mus. Brit. &c.) This species is often brought alive to this country ; but it does not appear to have been as yet well described. It is easily distinguished by its uniform colour, and irregularly radiately grooved plates, decussated by small, irregular, concentric wrinkles. The young is obscurely varied with darker brown on the feMYS. 29 back, and the sternum is marked with a dark -edged, irregular, central, green line, with a ring on each side of the front lobe placed on the suture between the gular and humeral plates. 23. Emys scr'ipta, (Lettered Termpin.J Testa oblonga longitudina'liter rugulosa fusca, lateribus irregulariter luteo- fasciatis, postice subdentata irregulariter luteo annulata, scutellis vertebralibus obtuse carinatis, Imo urceolato, 4to et 5to longe hexagonalibus, marginalibus subtus maculis centralibus rotundatis ornatis, sterno convexo lutescente antice maculis 2, lateribusque utrinque 4, notato : capite gulaque linea centrali, temporibus utrinque macula lutea, variegatis. Test, serratu. Daud. t. 21, f. 1, (fig. mediocris) non Schw. nee Bell. Jun. Testudo scripta. Schoepf, t. 3, f. 5, (v Mus. Par.) cop. ; Lettered Tortoise, Shaw. Zool. t. , (v. Mus. Bell.) Hab. in America Boreali, Carolina Bosc. (v. t. Mus. Bell,&c.) The shell large, convex, brown, and irregularly yellow- lined ; the under edge has a series of black spots on the back edo-e of each marginal plate (not on the sutures, as represented by Daudin) ; the sternum yellowish, with a spot on the centre of each gular plate, and four on each side on the costo-sternal symphysis. The head with a central yellow line, another on the upper lip, a forked one on the centre of the chin, and a triangular spot behind each eye. 24. Emys serrata. {Serrated Terrapin.) Testa oblonga Ion - gitudinaliter rugulosa olivaceo fusca, fasciis pallidis irregu- iaribus transversis variegata, postice subdentata, scutellis vertebralibus obtuse carinatis, Imo longe urceolato, reliquis longe hexagonalibus, marginalibus subtus maculis subocel- latis ad suturas positis, sterno flavescente piano, lineis nigro marginata ornato. Emysreticidaria. Bell, MSS. (v. Mus. Bell.) Emysserrata, Say. Emys serrata, Var. Schw. Test, serrata, (5 Merrem. /3 Testa scutellis vertebralibus mediis complanatis, (v. M. Brit.) Hab. in America Boreah. (v. t. Mus. Bell, &c.) The under side is yellowish, with a series of subocellated spots on the sutures of the marginal plates, and a dark edo-ed pale line on the sutures, between the outer sides of the sternal and marginal plates. Some of the specimens 30 EMYS. show slight indications of a dark edged band along the centre of the sternum; and two black rings on the sutures, between the gular and humeral sternal plates. As Harlan appears to call this the red-bellied Terrapin. it may be Emys rahriventris of Mr. LeConte, cited in Cuvier's Rcgne Animal. 25. Emjjs or fiat a, (Ornamented Teirapin.J Testa oblonga longitudinaliter rugosa olivacea, scutellis vertebralibus irre- gulariter annulatis, Irno urceolato, secundo et tertio longe hexagonis, costalibus marginalibusque supra annulis pal- lidis pupillis marginibusque nigris donatis, infra ocellis suturalibus; sterno pallido, centro lateribusque fasciis nigro marginatis donatis. Junior. Viridis, scutellis costalibus annulis duobus aurantiis alUisque luteis notatis, capite luteo lineato, (v. v. et v. t. Mus. Brit.) Hab. in America Meridionali, (v. t. Mus. Col. Surg. Mazctland, A. ColHe, Esq.. Brit. Mus.) The costal and marginal shields are marked with black edged pale rings, having a black central dot, and the vertebral plates with irregular rings. The under side is pale yellow, with a dark edged line down the centre, and a double one the whole length of the suture, between the sternum and the marginal shields. The head of the young animal has yellow lines, with an interrupted orange streak on each side the occiput, and a forked one on the side of the throat. 26. Erm/s ri/gnsa, (Rugose Terrapin.) Testa ovata oblonga postice sub-serrata leviter tuberculato-carinata nigra punctis literisque luteis ornata, scutellis rugulosis areolis indistinctis, marginalibus luteis nigro punctatis, sterno luteo punctis lineolisque nigris ornato. Test, rvgosa, Shaw, Zool, iii. t. (v. Mus. Col. Surg.) )3. livida. Testa livida, nigro punctata, (v. Mus. Bell.) Hab. in America Septentrionali r The shell oval oblong, bhmtly and subtuberculately keeled, the sides slightly contracted, black dotted and lettered with yellow ; the shields longitudinally and somewhat radiately grooved ; the areola indistinct ; the first vertebral one long, urceolate 2d, 3d, and 4th long 6-sided, the 5th broad 6- sided, the marginal shields smoothish yellow lettered and dotted with black ; the five hinder pair deeply lobed in the centre, the nuchal plate long and narrow, the sternum yellow, black EMYS. 'A] dotted, truncated before and behind, the axillary plate small, the inguinal ones larger; length \\~, breadth, 6 inches. Shaw's figure scarcely shews the rugosities or the keel suffi- ciently distinctly. Mr. Bell's specimen may be discoloured, it difters in being livid grey, and the spots impressed and as if burnt into the horny coat. 27. Emys Lesiietir'i ,[Lesii€iir' sTerrapin.) — Testa ovatacon- vexa Isevi, antice tuberculato-carinata, postice profunde den- ticulata. olivaceo fusca, lineis pallidis nigro marginatis anas- tomosantibus ornata, sterno luleo, scutellis nigro marginatis, raarginibus subtus olivaceo fuscis lineis inequalibus subconcen- tricispallide nigro marginatis ornatis ; capite pedibusque lineis luteis angustis numerosis variegatis, temporibus macula trian- gulari notatis. "/3. Scutello vertebrali primo urceolato." Ejnj/s aeogra- phica, Lesueur, Jour. Acad. N. S. Phil. t. Emys pseiido- geographica, Lesueur Mss. (Mus. Paris.) Hab. in America Boreali. (v. t. IMus. Brit.) The first and fifth vertebral plates are broad and pentangular and the rest broad hexagonal. The sterno-costal suture and the under side of the margin is blackish olive, with broad and narrow dark-edged pale irregular somewhat concentric lines. Emys geographica of Lesueur agrees with the jNIuseum specimen, except in that the first vertebral plate is not urn- shaped, and Lesueur does not notice the triangular temporal spot. 28. Emys BeUi'i, {BeWs Terrapin.) — Testa oblonga centro depressa lateribus convexis olivacea, fasciis irregularibus viri- dibus nigropunctatis reticulata, subtus nigrescente punctis maculisque luteis ornata, marginibus antico posticoque luteis maculis nigris flavo punctatis ad suturas positis, sterno margine irregulari luteo circumdato. Tnhab. (v. t. Mus. Col. Surg.) The shell solid, oblong, the centre depressed, the sides rounded, the margin broad, centre slightly reflexed over the hinder legs, the nuchal shield long linear, the vertebral shields nearly square, the first urnshaped, the others six-sided, with the sides straight; above olive, varied with irregular pale greenish lines dotted and edged with black, placed on the marsin and across the middle of each of the shields, the 32 EMYS. centre band being most distinct on the marginal plates ; the under side of the margin black dotted with yellow on the sides, and yellow on the ends, with irregular yellow dotted black spots placed on the suture between each of the plates ; the symphysis with a broad longitudinal yellow spotted black band, separated from the margin, and divided down the middle by two pale yellow lines. The sternum nearly flat, its surface and upper edge black, dotted with yellow and surrounded by an irregular yellow edge; the ends truncated, the front one denticulated, the hinder lobe broad, rounded on the sides. Length 9, breadth 8-|-, inches. 29. Emys kinosternoides, {White spotted Terrayin.) Testa (puUi) oblonga depressa, pallide fusca obtuse albo carinata, margine albido seiTato subtus luteo immaculato, scutellis disci fasciis irregularibus albis nigro marginatis ornatis, sterno antice posticeque rotundato ; capite fusco fasciis albidis variegate subtus pallido. Hab. (v. t. Mus. Col. Surg.) Of this species I only know a very young specimen in spirits. The shell is oblong depressed with a broad low continuous central white keel ; the second and third verte- bral shields each have a brown edged white irregular cross band, and the costal plates are varied with irregular brown edged white spots ; the margin white, with a series of trian- gular brown spots placed round the inner edge on the suture between each of the plates ; beneath pale yellow not spotted. The sternum nearly flat, acutely romided in front and rounded hehind; the axillary and inguinal plates small, the head brown varied with a white band, beneath whitish ; the nape brown, with three brown edged pale bands. Length of shell 1^ inch. Besides the peculiarity of the colouring in this species, it is the only species that I am acquainted with that has both ends of the sternum rounded as in Kinosternon. 30. Emijs annulifera, [Ringbearing Terrapin.) — Testa (pulli) oblonga depressa scutellis punctatis pallide fuscis luteo lineatis, vertebralibus annulis fuscis ad suturas positis ; subtus pallida fusco marmorata. Hab. (v. t. Mus. Brit.) Shell oblong depressed, behind entire, the scales punctu- late pale brown lined and ringed with yellow, the vertebral series with a narrow central row of brown rings, with a larger EMVS. 33 . series on each side, the costal shields lined with brown and white, the marginal with concentric sub-eyed squarish brown rings placed on the sutures and each occupying two half shields; beneath pale with irregular brown spots and lines placed on the sutures; sternum before round, behind truncated; head with numerous unequal w'hite lines. Length, ]i inch. JEmydes Fossiles, Cuv. Os. Fos. v, 227. a. Emys Hugi, Testa ovali, leviter convexa, scutellis verte- bralibus latis, marginahbus ansustissimis Emys du Jura, Cuv. Os. Fos. V. 237, t. f. 4. 5,^7, testa. 8, 9, 10, 11, caput. Fossil in the Jura, in a nearly perfect state. Length 24. breadth 20 inches. b. Emys Mantelli. Emys de Sussex, Cuv. Os. Fos. v. 232. c. Emys Cuvieri. Emys des Molasses de la Dordogne, et de la Suisse. Cuv. Os. Fos. v. 232, t. 15, f. 19. AUied^'to Emys Serrata. d. Emys Parkinsonii. Emys de Sheppey. Cuv. Os. Fos. v. 234, t. 15, f. 14, 15. Parkinson Org. Rem. iii. t. 18, f. 3. and 2, cop. Cuv. Os. Fos. t. , f. 13. e. Emys Hugii. Testa depressa. Emyde du Jura, n. 2, Cuv. Os. Fos. V. 229. t. 20, f. 6. Fossil in the Jura. Length about 22 inches. f. Emys Triomichoides. Testa postice costis tribus diver- gentibus donata. Emyde du Jura, n. 3, Cuv. Os. Fos. v. 229', t. 21, f. 1. Fossil in the Jura, M. Hugi ; only a portion of the back of the shell. It is peculiar for the back ribs being united together without the intervention of the dorsal plate of the vertebrae, as in some of the Trionyches. 0-. Emys Camperi. Emyde de Bruxelles. Cuv. Os. Fos. v. 236, t. 15, f. 16, t. 13, f. 8. h. Emys Luc'ii. Emyde Deluc, Bourde Mem. Emyde des Sables d'Aste, Cuv. Os. Fos. v. 233. i. Emys Parisiensis. Emyde des Platrieres de Paris. Cuv. Os. Fos. iii. 329. v. 227. F 34 KINOSTERNON. Gen. 3. KINOSTERNON, Spix. [Kinosternon.) Sternum latum antice rotundatum, postice rotundatum vel sub-bilobum; lobus medius fixus, per symphysiu osseum cum testa conjunctus, anterior et posterior plerumque mobiles lobo medio ligamentis articulati ; scutella sterni 11, gulare unicum. This genus is at once distinguished from Emijs by the suture which unites the sternum to the shell being only the length of the abdominal plates, and its being covered by the long axillary and still longer inguinal plate ; the sternum is also peculiar for having only 1 1 shields, the front or gular pair being soldered into one, and for being divided across by two sutures into three lobes. The front and hinder lobes move freely on the central one. The sutures are placed be- tween the pectoral and abdominal, and the femoral and ab- dominal plates ; often one or both become obliterated by age, or other circumstances. The marginal plates vary from 20 to 23, or 25 ; the side ones are narrow, and the nuchal plate is generally distinct and slender. The chin of the animal is bearded like the Hydraspides, and the end of the tail is often clawed. It consists of part of the genus Terrapene, of Merrem ; Cistuda, of Fiemming ; and includes the genus Kinosternon, of Mr. Bell ; and also part of his genus Sterno- thenis. The species hitherto found all come from America. 1. K'tnost. scorpioldes, (^Tkree-keeled Kinosternon.) Testa oblonga sub-compressa tricarinata carinis continuis, scu- tellis dorsalibus longe hexagonis sub-imbricatis, sterno lato postice leviter bitido, lobis anticis mediisque equalibus, lobo postico sub-longiore. Pennsylvanian Tortoise, Var., Shaw, Zool. 1. 15, (v.t. Mus. Col. Surg.) Test, scorpioides, Lin. Lacep. Quad. Ov. t. (v. t. Mus. Par.) Chersina scorpioides, Merrem. Emys scorpioides, Schw. Test, tricarinata, Daud. Kinosternon Shavianum, Bell. Terrapene triporcata, Wiegmann? Kin. longicaudatum, Spix, t. 12.? a. Acuta, sterno postice acuto integro, t. f. (v. t. Mus. Brit.) Jun. Testudo Retzii, Daud. Emys Retzii, Schw. (v. t. Mus. Par.) Emj/s tricarinata, Schoepf. t. 22. cop. Shaw, Zool. ii. t. 11. Terrapene tricarinata, Merrem, 28, (v. Mus. Bell, Col. Surg ) N Tah. (K J y/ 4:>^/>-s/-/^^^>^H^ Jz-i^^^^'- C^A-^a ^ / ^ KINOSTERNON. 35 j3. Scutellis marginalibus lateralibus sub-latioribus, (v. t. Mus. Bell.) Hab. in America Meridionali; Mexico, lJ^eigman?i; Brasilia, SpLv. (v. t. Mus. Brit. Bell, &c.) The finest species of the genus. Shaw's figure is very good, but the anterior marginal plates which exist in his specimen have been omitted by the artist. Tt varies in regard to the breadth of the dorsal plates and the distance between the dorsal keels. Weigmann, (Isis, xxi. 364,) refers to Shaw's figure, and his description agrees excellently with our species, except that he says sterno antice mohili angustissimo, and areolJs steriii 12, neither of which correspond. The usually accurate Schweiger falls into the same error with regard to the sternal plates, in his description of both the old and young state, but this may originate in the gular plates being crossed by an elevated ridge in the usual place of the suture. All the specimens I have seen have only had eleven. Kinosternon hrevicaudatum, Spix, t. 13, appears only to differ in the tail and hinder lobe being rather shorter, perhaps a male ; both the species have 23 marginal plates in the figures, and 25 according to the text. 2. K'uiost. Pennsylvanicum, (JPennsylvanianKinosternoii.) Testa oblonga fuscaleviter convexa, scutellis dorsi complanatis, sterno parum angustiore quam apertura testae dorsalis, postice bifido, lobo centrali anteriore subbreviori ; capite pallida lineolato. Test. Pennsylvanica, Bosc. Emys Pennsylva?iica, Schw, Test, suhriifa, Lacep, Edw. t. 287, cop. Shaw Zool. iii. t. 17. Seligman, t. 37, and E. M. t. 5, f. 1. — Schoepf, t. 24, f. A. B.— Daud. t. 24, f. 1, 2. /3. Sterno solido. Test, glut'mata, Daud. t. 24, f. 4, (mala.) — Schoepf, t. 24, f. b. Emys glut'mata, Merrem, Bell. Y. Scutellis marginalibus 20. Terrapene Boscii, Merrem. Sternotherus Boscii, Bell. Hab. in America Boreali, (v. t. Mus, Brit. Bell, &c.) 3. Kinost. odoratum, {Musky Kinosternon.) Testa ovata carinata lateribus declivibus fusca, sterno angustissimo antice acuto postice acute bifido ; capite fusco linea laterali ornato. Test, odorata, Bosc, Daud. t. 24, f. 3 ? Emys odorata, Schw. (v. t. Mus. Brit.) Cistuda odorata. Say. Sternotherus odoratus, Bell. 36 CHELYDRA. (i. glutinala. Sterni lobis immobilibus (v. t. Mus. Brit.) Hab, in America Boreali. Gen. 4. CHELYDRA, Schw. Sternum angustissimum cruciforme centro coriaceo, scu- tella membranacea, symphysis gracilis elongata scutello pro- prio tecta. The head large and beak horny, like the other Terrapins, but the sternum is very narrow and cross-shaped. All the bones of which it is formed resemble those of the genus Trionyx and Chelonia ; they are only covered with five pairs of very thin skin-like sternal plates. The symphysis between the sternum and the back shell is very long and narrow, and covered with a peculiar plate which may represent the pair that are deficient in the centre of the sternum. The axillary and inguinal plates are distinct and equally thin. The limbs are large and strong, and the tail is long and longitudinally crested on the upper surface. This genus consists of only a single species, which, from its resemblance to the Alligator, has been called the Alligator Tortoise. It is Emj/s, section B. of Oppel, and it has been called Chelonura. by Fleming, Saurochelys, by Latreille, and Rapara, by myself ; but I have adopted here the oldest name, published in a paper which I could not find when T first wrote of these animals. Chelydra serpentina, {Alligator Terrapin.) Testd oblonga, medio depressa bicarinata, postice obtuse dentata. Test, serrata. Penn. Arct. ZooL not Shaw. Chelydra lacertina, Schw. (v. Mus. Brit.) Howit Liverpool Mus. t. Junior. Testa tricarinata postice acute dentata. Test, serpentina, Lin. Schoepf, t. 6, cop. t. Daud. t. 60, f. 2, and Shaw, Zool. t. 29. Chelydra serpentina, Schw. Chelonura serpentina, Say. Test, longicauda, Shaw, Mss. Mus. Col. Surg. (Pullus.) Hab. in America Boreali, (v. v. Exeter Change, v. t. Mus. Brit. Par. ct D. Bell), called coulta by the slaves. STERNOTHERUS. 37 Fam. ///.— CHELYD^, or ChelydidcE. Pedes palmati, digitis distinctis, unguibus 5-4 eloiigatis iicuminatis ; nares tubulosi ; mandibulse plerumque cornese ; testa depressa, scutis cornels tecta ; scutella marginalia 24-25, caudalia distincta, sterni 13. The head much depressed and broad, covered with regular large shields ; the neck long, broad, retractile under the side of the shell, and not into the centre of it as in the Emydes, the shell generally much depressed, the margin furnished with a small nuchal plate, and the caudal pair always separated by a distinct suture. The sternum is always attached to the upper shell by a bony suture. The symphysis is rather short and covered by the outer end of the pectoral and abdominal plates. The axillary and inguinal plates are usually very small and hidden by the margin. The sternal shields thirteen, the additional or intergular plate being situated between the pair of gular plates, or between the hinder angle of them and the front angle of the humeral plates. The vertebrae of the neck are so formed that the animal cannot withdraw the head into the shell, like the other tortoises : and the bones of the pelvis are united by a bony process to the hinder part of the ster- num as well as to the vertebrae, as observed by Cuvier in the skeleton of Chelys, (Os. Fos. v. 214, t. 12, f. 20) ; and by Mr. Bell in Hydraspis. This character may be known even when the pelvis is lost by the scar left by its attachment to the bone. They live in ponds and ditches in the warm regions of both Continents, eating MoUusca and worms, and feeding only in the water. The animals of this group require further examination and comparison. Of the sixteen species here indicated, two are Australasian, five African, and nine Tropical American. Gen. 1. STERNOTHERUS, {Sternotherus.) Sternum latum, lobus anterior mobilis sutura transversa ligamentosa ad lobum medium articulatus, scutellum inter- gulare marginale, nuchale nullum, maxillge cornege. 38 CHELODINA. The shell is roundish, rather convex ; the sternum is broad, rounded before, bifid behind. The front lobe (and sometimes the hinder one, according to Daudin) is separated from and moveable on the central lobe by a transverse carti- laginous suture. The suture of the front lobe is placed be- tween the pectoral and abdominal plates, and the front lobe is furnished with an internal bony process on each side, near the suture, serving as a hinge. The inlergular plate is in the margin, between the gular plates ; the margin is destitute of any nuchal plate ; the lateral marginal plates are very narrow^ and the two front ones arc very short and square. The inner surface of the hinder lobe of the sternum, in the specimens which I have examined, is marked with the scar of the attachment of the pubis. This genus consists of only part of the species of the genus established under the above name by Mr. Bell ; the other species belonging to the family of Emydce. 1 . Slcrnolherus castaneiis, (Chesnut SternotJierits.) — Testa ovata convexa acute carinata castanea, scutellis nigris levis- simis radiantibus radiatim rugosis ; areolis punctulatis rugosis. Einys casla/iea, Schweiger. Testiido siibiiigra> /3. Daud. (v.t. Mus. Par.) Stcrnotherus Leach'ianus, Bell Zool. Jour. iii. t. 14. (v. t. Mus. Bell.) Habit. (v.t. Mus. Bell et Paris.) 2. Stcrnnthen/s snlniger, [Black Sternotherus.) " Testa ovali convexa, scutellis nigris nitentibus in margine striatis ; verte- bralibus mediis planis leviter carinatis." Test, subnigra. Lacep. Daud. t. 7, f. "2. Test, nigricans, Merrem. Emys subnigra, Schw. Hab. (non vidi.) This species scarcely appears distinct from the former ; I did not observe it at Paris, and have therefore given Sch wel- der's specific character. According to Daudin the hinder lobe of the sternum is mobile ; but that may be a variety of age. This character is not noticed by Schweiger. Gen. 2. CHELODINA, [Chelodina.) Maxillae corner, sternum latum solidum, scutellum inter- gulare hexagonum inter angulos scutellorum gularium hume- raliumque, nuchale distinctum. HYDRASPIS. 39- This genus forms section c. of the genus Emys of Oppel, and part of the Hj'draspis of Mr. Bell. The shell is depressed, hroad, and covered with very thin smooth membranaceous shields, without any distinct areolae, the sternum is very broad, and rounded in front, with the intergular plate long, hexagonal, and placed in the angle between the gular and humeral plates. The neck is very long : the head as in the other animals of this family, is bent in under the side margin of the shell, as was first observed by Dr. Oppel, in 1811. 1. Chelodina lo?igicolUs, {Long-necked chelodina.) Testa oblongadepressa, scutellis fuscis.vertebralium primoelongato, sterno iuteo scutellis fusco-marginatis. Testudo longiadlis, Shaw, New Hol.t. 7, (v. t. Mus. Brit.) cop. Zool. iii. t. 6. Emys longicollis, Schw. (v. t. Mus. Par.) Habit, in Nova Hollandia, (v. v. et t. Mus. Brit.) Gen. 3. HYDRASPIS, [Hijdraspis.) Maxillae corneae, nares breves tubulares, sternum solidum angustum, scutellum inrergulare marginale. These shells are depressed and broad, covered with rather thin horny plates, generally in their youth marked by distinct areolae, which disappear by age. The margin is generally narrow on the sides, and often furnished with a nuchal plate. The sternum is narrow, truncated in front and bifid and narrower behind. The intergular plate is marginal between the gular plates. The head is broad, depressed, and covered with one or more large shields ; and the chin is furnished with two beards. The neck is large and warty. According to Spix, the Brasilian species feed on fruit and grass, and lay their eggs in the sand. Those I have had alive of the Cape species chose worms and fish-intestines, and would not touch vesietables. This genus only contains part of the Hydraspis of Mr. Bell ; the Chelodina of Fitzinger appears to be synonymous with it. 1. Hydraspis suhrufa, [Cape Hydraspis.) Testa oblonga humili unicolore pallide fusca, scutellis laevibus, vertebralibus complanatis, nuchali nullo, marginalibus laterahbus angustis- simis ; capite depresso laevi, scutellis magnis tecto. 40 HYDRASPIS. Test, sulrufa, Lacep. t. 12 ; cop. E. M. t. G, f. 5. (v. Mus. Par.) Emys subrufn, Schw. Test, hadia, Daud. Junior. Scutellis concentrice sulcatis. Aestudo galeata, Schoepf t. 3, f. 1, cop. Galeated tortoise, Shaw, t. 12. Test, scahra, Retz. Emys olivacea, Schw. (v. t. Mus. Brit.) Test. Se?iega/ensis, Daud. (v. t. Mus. Par.) Habitat in Africa, Cap. Bon. Spei, Senegal? (v. v. Hort. nost. et t. Mus. Brit., Par., et Bell.) I have never seen this tortoise from India, but being brought from the Cape by the Indian ships, they, as well as many other Cape animals, are often called Indian. This animal is pale brown, black speckled, and the shell, when alive, ash-coloured, black-spotted. 2. Hydraspis AdansoiiU, [Adansoii's Hydraspis.) Testa ovata humili postice latissima lutea nigro punctata, scutello nuchali nullo, vertebralibus carinatis, primo elongato pandu- riformi postice triangulari. Emys Adansonii, Schw. (v. t. Mus. Par.) Hab. in Nigritia. The Paris Museum specimen has lost most of its shields. 3. Hydraspis Macquarrii, [New Holland Hydraspis.) Testa ovata depressa antice contracta postice expansa dentata olivaceo-fusca, linea dorsali impressa, scutellis rugulosis, nu- chali angusto, sterno lutescente. Emys Macquarrii, Cuv. MSS. R. A. ii. 1 1 note. (v. t. Mus. Par.) Habit, in Nova Hollandia, Macquarrie River. The hinder marginal plates are nicked in the middle of their edge. Length 11, breadth 8 inches. 4. Hydraspis planiceps, [Flat-headed Hydraspis.) Testa oblonga nigro fusca dorso complanata lateribus declivi, mar- ginibus lateralibus reflexis, scutello nuchali lineari, sterno luteo marginato, cauda brevi. Test, planiceps et Te . ^ " '*"