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BIX S^NCY*"*3- SYSID -BL- EW SERIES Systematic Review of Southeast Asian Longtail M&cm Macaca fascicular™ (Raffles, [11 590 . 5 BIX FI N.S. 31 KJ0" Publication 1470 PUBLISHED BY F IELD i Confabs :d for puti FIELDIANA Zoology NEW SERIES, NO. 81 Systematic Review of Southeast Asian Longtail Macaques, Macaca fascicularis (Raffles, [1821]) Jack Fooden Research Associate Division of Mammals Department of Zoology Field Museum of Natural History Chicago, Illinois 60605-2496 Accepted January 13, 1995 Published November 30, 1995 Publication 1470 PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY © 1995 Field Museum of Natural History Library of Congress Catalog Card Number: 95-61092 ISSN 0015-0754 PRINTED IN THE UNITED STATES OF AMERICA Table of Contents Abstract 1 Introduction 1 Geographic Distribution 2 Pelage 3 General Characterization 3 Sex and Age Variation 11 Geographic Variation 11 Dorsal Pelage Color Saturation 11 Dorsal Pelage Erythrism 13 Crown Color 15 Lateral Facial Crest Pattern 18 External Measurements and Propor- tions 21 Sex and Age Variation 21 Geographic Variation 22 Head and Body Length 22 Tail Length 23 Cranial Characters 30 Sex and Age Variation 30 Geographic Variation 33 Molecular Biology and Genetics 35 Mitochondrial DNA 35 Nuclear DNA 39 Multiple Alpha-Globin Genes 39 Highly Repeated Restriction Patterns ... 41 Blood Proteins 42 Blood Groups 45 Serum Cholesterol Response 46 Karyology 47 Diseases 47 Malaria 47 Simian T-Lymphotrophic Retrovirus, Type 1 49 Susceptibility to Attenuated Polio Virus ... 49 Natural History 49 Habitats 49 Arboreality/Terrestriality 50 Swimming 51 Troop Size and Composition 51 Home Range, Day Range 51 Density 53 Diet 54 Predators 54 Intertroop Behavior 56 Interspecific Behavior 57 Reproduction 58 Fossils and Subfossils 63 Systematics 65 Subspecific Taxonomy 65 Key to Recognized Subspecies 69 Subspecies Accounts 70 Macaca fascicularis fascicularis (Raffles, [1821]) 70 Macaca fascicularis aurea I. Geoffroy, [1831] 77 Macaca fascicularis philippinensis I. Geoffroy, [1843] 79 Macaca fascicularis umbrosa Miller, 1902b 79 Macaca fascicularis fusca Miller, 1903a 81 Macaca fascicularis lasiae (Lyon, 1916) 81 Macaca fascicularis atriceps Kloss, 1919c 82 Macaca fascicularis condorensis Kloss, 1926 84 Macaca fascicularis tua Kellogg, 1944 86 Macaca fascicularis karimondjawae Sody, 1949 86 Macaca fascicularis subspecies undeter- mined 87 Evolution and Dispersal 87 Stage I, > 1 Ma 88 Stage II, ca. 160 Ka 88 Stage III, > 18 Ka 90 Stage IV, ca. 18 Ka 95 Stage V, < 18 Ka 95 Stage VI, ca. 4.5 Ka 96 Acknowledgments 97 Literature Cited 98 Appendix 1 : Specimens Examined 1 20 Appendix 2: Gazetteer of Non-Philippine Macaca fascicularis Localities 1 28 Appendix 3: Dorsal Pelage Color Satur- ation in Fringing-Island Samples of Macaca fascicularis Compared with Saturation in Core-Area Reference Samples 181 Appendix 4: Dorsal Pelage Color Erythrism in Fringing-Island Samples of Macaca fascicularis Compared with Erythrism in Core-Area Reference Samples 1 84 Appendix 5: Crown Color Pattern Frequency in Samples of Macaca fascicularis That Include Dark-Crowned Specimens 187 Appendix 6: Geographic Variation of Lateral Facial Crest Pattern in Samples of Macaca fascicularis .... 188 Appendix 7: Head and Body Length in in Fringing-Island Samples of Macaca fascicularis compared with head and Body Length in Core-Area Reference Samples Collected at Similar Latitudes 1 90 Appendix 8: Tail Length in Fringing- Island Samples of Macaca fascicu- laris Compared with Tail Length in Core-Area Reference Samples Collected at Similar Latitudes 193 Appendix 9: Relative Tail Length in Fring- ing-Island Samples of Macaca fas- cicularis Compared with Relative Tail Length in Core-Area Reference Samples Collected at Similar Latitudes 196 Appendix 10: Blood-Protein Allele Fre- quencies at Polymorphic Loci in 1 1 Geographic Samples of Macaca fascicularis 198 Appendix 1 1 : Monthly Distribution of Recorded Reproductive Events in Natural Populations of Macaca fascicularis 200 Appendix 12: Subspecific Variation of Head and Body Length, Relative Tail Length, and Greatest Length of Skull in Macaca fascicularis .... 203 Subject Index 204 Index to Scientific Names 206 List of Illustrations 1 . Known locality records of Macaca fasci- cularis 3 2. Detail maps of non-Philippine localities of Macaca fascicularis A. Northwestern quadrant 4 B. Southwestern quadrant 6 C. Southeastern quadrant 9 3. Approximate extent of dry land in South- east Asia during last glacial maximum, ca. 18 Ka 12 4. Core area, shallow-water fringing islands and deep-water fringing islands inhabited by Macaca fascicularis 13 5. External characters in Macaca fascicu- laris 14 6. Dorsal pelage color saturation extremes in Macaca fascicularis 15 7. Crown color pattern in dark-crowned Macaca fascicularis 18 8. Infrazygomatic and transzygomatic lat- eral facial crest patterns in Macaca fas- cicularis 19 9. Geographic variation of lateral facial crest pattern in samples of Macaca fascicularis collected in Indochinese Peninsula, Isth- mus of Kra, Malay Peninsula, and neigh- boring islands 21 10. Latitudinal variation of head and body length in adult core-area specimens of Macaca fascicularis 24 1 1 . Latitudinal variation of head and body length in adult shallow-water fringing-is- land specimens of Macaca fascicularis compared with that in adult core-area specimens 26 12. Latitudinal variation of head and body length in adult deep-water fringing-island specimens of Macaca fascicularis com- pared with that in adult core-area speci- mens 27 1 3. Latitudinal variation of tail length in adult core-area specimens of Macaca fascicu- laris 29 14. Latitudinal variation of relative tail length in adult core-area specimens of Macaca fascicularis 30 15. Latitudinal variation of tail length in adult shallow-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens 31 16. Latitudinal variation of tail length in adult deep-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens 32 17. Latitudinal variation of relative tail length in adult shallow-water fringing-island specimens of Macaca fascicularis com- pared with that in adult core-area speci- mens 34 1 8. Latitudinal variation of relative tail length in adult deep-water fringing-island speci- mens of Macaca fascicularis compared with that in adult core-area specimens 35 1 9. Skull of adult female Macaca fascicularis, near topotype 36 20. Skull of adult male Macaca fascicularis, near topotype 37 21. Latitudinal variation of greatest length of skull in adult core-area specimens of Ma- caca fascicularis 40 22. Latitudinal variation of greatest length of skull in adult shallow-water fringing- island specimens of Macaca fascicularis compared with that in adult core-area specimens 41 23. Latitudinal variation of greatest length of skull in adult deep-water fringing-island specimens of Macaca fascicularis com- pared with that in adult core-area speci- mens 43 24. Blood-protein dendrograms for geograph- ic samples of Macaca fascicularis: Den- drogram that excludes Mauritius sample compared with dendrogram that includes Mauritius sample; consensus dendrogram 47 25. Type localities and limits of distribution of recognized subspecies of Macaca fas- cicularis and type localities of synony- mous nominal species or subspecies ... 64 26. Variation of greatest length of skull in adult Macaca fascicularis fusca (Indonesia: Pu- lau Simeulue) compared with variation in adult Sumatran M. f fascicularis 83 27. Tail length vs. head and body length in Macaca fascicularis fusca and M. f la- siae 83 28. Craniometric comparisons of adult Ma- caca fascicularis atriceps, M. f condor- ensis, and mainland Indochinese M. fas- cicularis 85 29. Latitudinal variation of adult head and body length in samples of Macaca fasci- cularis collected in the southern part of the Indochinese Peninsula and adjacent Isthmus of Kra, north of 10°N, compared with that in samples of parapatric hi. mu- latta collected in the northern part of the Indochinese Peninsula 92 30. Latitudinal variation of adult greatest length of skull in samples of Macaca fas- cicularis collected in the southern part of the Indochinese Peninsula and adjacent Isthmus of Kra, north of 10°N, compared with that in samples of parapatric M. mu- latta collected in the northern part of the Indochinese Peninsula 93 3 1 . Latitudinal variation of adult tail length in samples of Macaca fascicularis col- lected in the southern part of the Indo- Chinese Peninsula and adjacent Isthmus of Kra, north of 10°N, compared with that in samples of parapatric M. mulatta col- lected in the northern part of the Indo- Chinese Peninsula 94 List of Tables 1 . Dorsal pelage color saturation in core-area Macaca fascicularis: summary of varia- tion 16 2. Dorsal pelage color saturation in core-area Macaca fascicularis: geographic variation of mean value in 2-degree latitude-lon- gitude blocks 16 3. Dorsal pelage erythrism in core-area Macaca fascicularis: summary of varia- tion 17 4. Dorsal pelage erythrism in core-area Ma- caca fascicularis: geographic variation of mean value in 2-degree latitude-longitude blocks 17 5. Crown color pattern in Macaca fascicu- laris: summary of variation 18 6. Lateral facial crest pattern in Macaca fas- cicularis: summary of variation 20 7. External measurements and proportions in age/sex classes of wild-collected Ma- caca fascicularis 22 8 . Head and body length in core-area Macaca fascicularis: summary of variation 23 9. Summary comparison of head and body length in fringing-island and core-area samples of Macaca fascicularis 25 10. Tail length and relative tail length in core- area Macaca fascicularis: summary of variation 28 1 1 . Summary comparison of tail length and relative tail length in fringing-island and core-area samples of Macaca fascicularis 33 1 2. Cranial measurements and proportions in age/sex classes of wild-collected Macaca fascicularis 38 13. Schedule of emergence of deciduous teeth in captive Macaca fascicularis 38 14. Schedule of emergence of permanent teeth in captive Macaca fascicularis 39 1 5 . Greatest length of skull in core-area Macaca fascicularis: summary of variation 39 16. Summary comparison of greatest length of skull in fringing-island and core-area samples of Macaca fascicularis 42 17. Frequency of mtDNA types revealed by five restriction enzymes in six geographic samples of Macaca fascicularis 44 18. Frequency of alpha-globin gene haplotypes in geographic samples of Macaca fascicu- laris compared with frequencies in samples of M. mulatta and M. fuscata 45 19. Blood proteins: summary of frequencies of major alleles at polymorphic loci in 1 1 geographic samples of Macaca fascicu- laris 46 20. Summary of allele frequencies at the vita- min D binding protein locus in geographic samples of Macaca fascicular is 47 2 1 . Allele frequencies at the complement C6 locus in three imprecisely localized geo- graphic samples of Macaca fascicularis 48 22. Geographic variation of A-B-O and Lewis blood group phenotype frequencies in Macaca fascicularis 48 23. Geographic distribution of malaria spe- cies identified as natural parasites in Ma- caca fascicularis samples 49 24. Known vectors of malaria species that have been identified as natural parasites in Macaca fascicularis 49 25. Simian T-lymphotrophic retrovirus, type 1 : frequency of Macaca fascicularis in- dividuals or groups positive for anti- bodies 50 26. Recorded habitats of Macaca fascicu- laris 52 27. Recorded size of nonprovisioned troops of Macaca fascicularis 53 28. Ratio of sexually mature males to sexually mature females reported in nonprovisioned troops of Macaca fascicularis 54 29. Estimated home range and day range in troops of Macaca fascicularis 55 30. Foods reported eaten by nonprovisioned troops of Macaca fascicularis 56 3 1 . Daily waking-hour time budget estimates for Macaca fascicularis 57 32. Copulatory behavior data in two Macaca fascicularis groups 60 33. Macaque fossils or subfossils collected within geographic range of Macaca fas- cicularis 62 34. Summary comparison of external char- acters in recognized subspecies of Macaca fascicularis 66 35. Subspecies of Macaca fascicularis recog- nized as valid by Chasen (1940a) and re- cent authors 68 36. Macaca fascicularis aurea: external and cranial variation in adult specimens ex- amined 80 37. Sexual dimorphism of greatest length of skull compared in Macaca fascicularis fusca and Sumatran M. f fascicularis . . 82 38. Comparison of cranial proportions in In- dochinese Macaca fascicularis, M.f atri- ceps, and M. f condorensis 84 39. Hypothetical stages in evolution and dis- persal of Macaca fascicularis 88 Systematic Review of Southeast Asian Longtail Macaques, Macaca fascicularis (Raffles, [1821]) Jack Fooden Abstract The longtail macaque, Macaca fascicularis (Raffles, [1821]), is systematically reviewed, based on examination of 2,049 museum specimens, study of relevant literature, and observation of natural populations. Macaca fascicularis inhabits tropical Southeast Asia, where its natural range extends from southernmost Bangladesh to the Philippines and from islands west of Sumatra to Timor. This review of M. fascicularis includes analyses of geographic variation in pelage characters, external measurements and proportions, cranial characters, molecular and genetic characters, and disease susceptibility. Evidence concerning natural history, reproduction, and paleontology also is investigated. Ten subspecies of M. fascicularis are recognized, and a key to these subspecies is provided. Subspecies accounts present synonymies, type and type locality information, geographic distributions, and diagnoses. Six hypothetical stages in the evolution and dispersal of this species are proposed and discussed. An annotated gazetteer lists approximately 1,150 localities at which M. fascicularis has been collected or observed. Introduction Macaca fascicularis (Raffles, [1821]) is widely distributed in mainland and insular Southeast Asia from approximately 2 1°N to 1 0°S latitude and from 92°Eto 1 2 6°E longitude (Fig. 1). Vernacular names applied to this species include crab-eating, cynom- olgus, kra, and longtail macaque. The natural abundance of M. fascicularis is indicated by the large number of specimens in museum collections (Appendix 1), which greatly exceeds that of any other species of macaque. The existence of this species was first made known to Western science by the English seaman W. Dampier (1697, p. 220) and the German mis- sionary G. J. Camel (in Petiver, 1705, p. 2199), both of whom observed M. fascicularis in the Phil- ippines. The name "macaque" was first applied to this monkey by Buffon (in Buffon & Daubenton, 1766, p. 190), based on an imported specimen of unknown origin. The present review of Macaca fascicularis is based on examination of 2,049 museum speci- mens (Appendix 1 ), survey of relevant literature, and observation of natural populations in Thai- land and Bali. Specimens examined are preserved in the following institutions, which hereafter are cited by means of the following abbreviations: aiuz Anthropologisches Institut der Uni- versitat Zurich amnh American Museum of Natural His- tory, New York ansp Academy of Natural Sciences, Phil- adelphia bm(nh) British Museum (Natural History), London bnhs Bombay Natural History Society, Bombay ctnrc Centre for Thai National Reference Collections, Thailand Institute of Scientific and Technological Re- search, Bangkok fmnh Field Museum of Natural History, Chicago irsn Institut Royal des Sciences Natu- relles de Belgique, Brussels mcz Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts FIELDIANA: ZOOLOGY, N.S., NO. 81, NOVEMBER 30, 1995, PP. 1-206 Bell Museum of Natural History, University of Minnesota, Minne- apolis Museum National d'Histoire Na- turelle (Mammiferes), Paris Museum Zoologicum Bogoriense, Bogor Naturhistorisches Museum, Basel Naturhistorisches Museum, Bern Naturhistoriska Riksmuseet, Stockholm Natur-Museum Senckenberg, Frankfurt Philippine National Museum, Ma- nila Primate Research Institute, Kyoto University, Inuyama Royal College of Surgeons, Odon- tological Museum, London Rijksmuseum van Natuurlijke His- toric, Leiden Simian Conservation, Breeding & Research Center, Inc., Tanay, Lu- zon Sarawak Museum, Kuching Staatliches Museum fur Tierkunde, Dresden Museum of Zoology, University of Michigan, Ann Arbor University of the Philippines at Los Banos, College of Forestry, Los Banos, Luzon University of the Philippines at Los Banos, Zoology Department, Los Banos, Luzon National Museum of Natural His- tory, Washington, D.C. Zoological Laboratory of Utrecht University, Utrecht Zoologisches Museum des Hum- boldt-Universitat, Berlin Zoologisches Museum der Univ- ersitat Zurich Zoological Reference Collection, Department of Zoology, National University of Singapore Zoologisches Sammlung des Bay- erischen Staates, Munich Zoological Survey of India, Na- tional Zoological Collection, Cal- cutta Geographic names used in this review generally are officially approved names. However, the fol- lowing conventional names are retained because MMNH MNHN NHMBa NHMBe NHRM NMS PNM rcs(om) RMNH SICONBREC SMK SMTD UMMZ UPLBCF UPLBZD USNM ZLUU ZMB ZMUZ ZRC ZSBS ZSI of their greater familiarity (alternative names in parentheses): Burma (Myanmar), Java (Jawa), Lesser Sunda Islands (Nusa Tenggara), and Su- matra (Sumatera). Cited geographic names fre- quently include the following generic terms: gun- ong (Malaysian) or gunung (Indonesian) = moun- tain; ko (Thai) = island; pulau (Malaysian and Indonesian) = island; and teluk (Indonesian) = bay. This review begins with a general account of the geographic distribution of M. fascicular is. Follow- ing are discussions of (1) pelage characters, (2) external measurements and proportions, (3) cra- nial characters, (4) molecular and genetic char- acters, and (5) disease susceptibility; these discus- sions emphasize patterns of geographic variation. Subsequent sections summarize available infor- mation concerning natural history, reproduction, and paleontology. A discussion of subspecific tax- onomy, supplemented by a key to recognized sub- species, is based on material presented previously. For each recognized subspecies, a formal subspe- cies account provides the following: annotated synonymy; information concerning types, type lo- cality, and distribution; subspecific diagnosis; and summary of specimens examined (complete list in Appendix 1). The final section of this review pre- sents a hypothetical interpretation of the evolution and dispersal of M. fascicularis. Geographic Distribution Macaca fascicularis inhabits tropical Southeast Asia (Figs. 1, 2). Its natural geographic range ex- tends from southernmost Bangladesh and south- ern Burma southward and eastward through the southern part of the Indochinese Peninsula (south of 1 7°N), the Isthmus of Kra, the Malay Peninsula, Sumatra, Borneo, Java, and the Lesser Sunda Is- lands as far east as Timor, the Philippine Islands, and numerous small adjacent islands, including the southernmost three Nicobar Islands. The presence of M. fascicularis in some islands in Southeast Asia has been attributed to prehis- toric human introduction (see below, Fossils and Subfossils; Evolution and Dispersal). During his- toric times, successful introductions have occurred in Mauritius Island (I.), east of Madagascar, prob- ably in the sixteenth century (Sussman & Tatter- sall, 1986, p. 30); southwestern Sulawesi, date un- known (Miiller, [1840], p. 17; M. Weber, 1890b, p. 102; Dammerman, 1929, p. 6); Anguar Island, north of New Guinea, early twentieth century FIELDIANA: ZOOLOGY a Specimens examined 0 Literature records 10 100 110 120 Fig. 1. Known locality records of Macaco, fascicular is. For details, see Figure 2A-C. (Poirier & Smith, 1974, p. 264); Hong Kong, ca. 1945 (Marshall, 1967, p. 44; Southwick & Manry, 1 987, p. 48); and Pulau (P.) Tinjil, south of western Java, 1988-1990 (Kyes, 1993, p. 78). Physiographically, the geographic range of M. fascicularis includes mainland Southeast Asia, shallow- water islands on the Sunda Shelf, and deep- water islands at or beyond the edge of the Sunda Shelf (Figs. 3, 4). During the most recent glacial maximum, ca. 18 Ka, sea level was reduced by about 1 20 m, which exposed most of the Sunda Shelf and extended mainland Southeast Asia to include Sumatra, Borneo, Java, and other shallow- water islands (Heaney, 1986, p. 131; 1991a, p. 56). During the preceding glacial maximum, ca. 160 Ka, sea level may have been reduced by about 1 70 m, which exposed more of the Sunda Shelf but still left most deep-water islands isolated. To facilitate zoogeographic analysis, in subse- quent discussions of geographic variation the range of M. fascicularis is subdivided into three com- ponents: (1) a core area of distribution that com- prises mainland Southeast Asia, Sumatra, Borneo, and Java, four large landmasses that were inter- connected by dry land ca. 1 8 Ka; (2) shallow-water fringing islands— small islands within the 120-m bathymetric line that have been isolated from the core area for less than 1 8 K years; and (3) deep- water fringing islands— islands beyond the 120-m bathymetric line that have been isolated from the core area at least since the beginning of the pre- ceding interglacial, ca. 1 20 Ka (Van Couvering & Kukla, 1988, p. 461) and that may never have been joined to the core area. Pelage General Characterization (Fig. 5) Dorsal pelage color in M. fascicularis varies from buffy to yellowish gray to golden brown to reddish brown to blackish. Individual hairs generally are conspicuously marked by one or two pale subter- minal bands that vary in color from pale yellowish to golden to rufescent. The crown usually is more brightly colored than the back (pale subterminal bands more conspicuous in crown hairs), but in some specimens the crown is contrastingly darker and duller than the back (pale subterminal bands reduced or absent in crown hairs). The anterior edge of the crown is bordered by a transverse blackish supraorbital streak; hairs at the vertex FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS d Specimens examined 0 Literature records a-m Absence reports 240 25 ^ ,9 „/ \\.\j "I Vietnam (V) L $r-^3 * Cambodia Fig. 2A. Detail map of non-Philippine localities of Macaca fascicularis, northwestern quadrant. (Philippine localities are mapped in Fooden, 1991, p. 2.) For documentation, see Gazetteer (Appendix 2). Abbreviations in parentheses are those used in Gazetteer locality codes. Bangladesh (Ba) 1. Bilasodia; Bimirdia; Ghorardia; Ochodia; Ruku- modia. 2. Jolir dia. Burma (Bu) 1. 2. 3. 4. 5. 6. 7. Lai char. Myengun Kyun. Arakan Division. Hainggyi Kyun. Preparis Island. Desertion Creek. Rangoon. 8. Pegu. 9. Wimpong. 10. Haungtharaw. 1 1 . Ataran River; Ye Forest. 12. Moscos Island Game Sanctuary. 13. Tavoy River. 14. Taungbyauk Chaung. 15. Kathema Kyun; Mibya Kyun. 16. Kaser Doo Wildife Sanctuary. 17. Kadan Kyun. 18. Great Tenasserim River; Mergui. 19. Tagoot; Tenasserim; Thagyet. 20. Letsok-aw Kyun. 2 1 . Lanbi Kyun. 22. Ban Sadein; Pakchan River, near Bankachon; Pak- chan River, near Maliwun. 23. Ru, Pulo. 24. Zadetkyi Kyun. Cambodia (C) 1. Siemreab. 2. Pang Roloem-Sur Sdei area. 3. Laem Ngop-Phumi Cham Yearn (Cambodia or Thailand). Laos (L) 1. Thateng, Muang. Nicobars (N) 1 . Ol-kolo-kwak vicinity. 2. Little Nicobar. 3. Kopenheat; Pulo Nyur. 4. Campbell Bay vicinity. Thailand (T) 1 . Ban Nam Lai Tai. 2. Ban Mae Na Ree. 3. Wong, Nam Mae, 53 mi (= 85 km) E of Um Pang. 4. Ping, Mae Nam; Wat Khao Noh. 5. Ban Pak Nam Pho; Wat Krieng Krai Klang. 6. Wat Khao Wong Kot. FIELDIANA: ZOOLOGY often form an irregular tuft or crest. Hairs on the side of the head are pale ochraceous gray to pale ochraceous brown and form a variably prominent lateral facial crest that usually extends from near the angle of the jaw to the crown, but may be restricted to the mandibular region. The face gen- erally is thinly haired; the facial skin is brownish to pinkish, except for the upper eyelids, which are sharply defined whitish. On the limbs, the proxi- mal part of the outer surface is approximately the same color as the adjacent surface of the trunk. More distally, the outer surface of the limbs be- comes paler, pale grayish to pale golden brown at the wrists and ankles. The dorsal surface of the tail is golden brown to blackish basally, becoming paler (pale grayish to grayish brown) distally. Pel- age on the ventral surface of the trunk, limbs, and tail is thin and pale gray to whitish. 9. 10. 11. 12. 13. 14. 15. 16. 17. 18. 19. 20. 21. 22. 23. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34. 35. 36. 37. 38. 39. 40. 41. 42. 43. 44. 45. 46. 47. 48. 49 Sarn Pra Karn. Wat Khao Sompoad. Lat Bua Khao. Ban Sakaerat. Aranyaprathet. Kosumphi Forest Park. Wat Koo Pra Kona. Ban Kosum. Phu Phan. Don Poo Tao; Prang Koo. Ban Wan; Wat Ban Kan Yai. Wong, Nam Mae, 40 mi (= 65 km) E of Um Pang. Kata Taek; Sap Khao; Thap Salao, Huai. Ban Tamrong Phato; Khwae Noi, Mae Nam. Ban Phu Toie. Khao Phatowee. Wat Noi Chompoo. Wat Phra Buddha Chai. Pak Chong, Sathani. Ban Huai Maenam Noi. Siam, 13°45'N, 99°25'E. Khao Suan Luang; Tham Chomphon; Wat Cha-Am Kiri; Wat Huai Takhaeng; Wat Khao Chong Phran; Wat Khao Khan Hok; Wat Khao Yod Thong; Wat Ngern Rung Sawang; Wat Ratch Singkhorn; Wat Tham Kunchhorn. Wat Tham Sala. Bangkok; Chao Phraya, Mae Nam. Khangkhao, Ko; Samuk, Khao; Si Chang, Ko; Si Racha vicinity. Wang Kaew. Laem Sing mountains. Ven-Ven. Chang, Ko. Kut, Ko. Khao Wang; Phet Buri; Samut Songkhram vicinity; Wat Ban Rai Don; Wat Bun Thawi; Wat Khao Ban- dai It; Wat Khao Takhrao; Wat Khao Tamon; Wat Kut. Pran Buri, Mae Nam.; Wat Khao Takieb. Khram Yai, Ko; Klet Kaeo, Ko. Khao Sam Roi National Park. Prachuap Khiri Khan, few miles north; Wat Tham- mikaram Varaviharn. Wat Tham Khao Phlu. Wat Tha Mai Lai. Chumphon, Khlong. Phayam, Ko. Laem Son National Park. Chiew Larn Reservoir. Thung Thong Waterfowl Reserve. Khlong Pah Yie, Ko Samui. 50. Phangan, Ko. 51. Khao Lampi-Hat Thai Muang National Park; Phangnga Bay National Park; Wat Tham Suwan Khuha. 52. Na Ka Yai, Ko; Yao Noi, Ko. 53. Rang Yai, Ko. 54. Phi Phi Don, Ko. 55. Lanta Yai, Ko. 56. Ban Phra Muang; Hat Chao Mai National Park; Kantang; Muk, Ko; Talibong, Ko. 57. Mu Ko Phetra National Park. 58. Butang, Ko. 59. Telok Wau, Ko Tarutao. 60. Ban Nong Kok; Ban Nong Put; Ban Thap Plik, 1 km NE; Hat Noppharat Thara-Mu Ko Phi Phi Na- tional Park; Tham Horn; Wat Tham Sua. 61. Tyching. 62. Khao Rang Kai. 63. Nakhon Si Thammarat. 64. BanNa. 65. Wat Khuha Sawan; Wat Suwankuha. 66. Nang Kham, Ko. 67. Khao Noi/Khao Tangkuan. 68. Nong Chik region; Pattani; Pho, Laem; Yaring, tidal creeks near; Yaring region. 69. Ban Sai Kau; Kampong Biserat; Wat Khuha Phi- muk. Vietnam (V) 1. Sontra Peak. 2. Lac Giao. 3. Xa Trang Bom. 4. Ho Chi Minh City. 5. Tay Ninh. 6. Phu Quoc Dao. 7. Airfield building vicinity, Con Son; Ben Dam vi- cinity, Hon Ba; Con Son. Absence reports a. North Andaman Island. b. Middle Andaman Island. c. Henry Lawrence Island. d. Barren Island. e. South Andaman Island. f. Little Jolly Boy Island. g. Rutland Island. h. Little Andaman Island. i. Car Nicobar Island. j. Tillanchong Island, k. Camorta Island. 1. Nancowry Island; Trinkat Island, m. Phai, Ko. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS d Specimens examined 0 Literature records a- n Absence reports """W-Maffn*. Java (J) Fig. 2B. Detail map of non-Philippine localities of Macaca fascicularis, southwestern quadrant. (Philippine localities are mapped in Fooden, 1991, p. 2.) For documentation, see Gazetteer (Appendix 2). Abbreviations in parentheses are those used in Gazetteer locality codes. Indian Ocean (IO) 1. We, Pulau. Simeulue, Pulau: 2. Dalam, Lhok; Sibaboh, Lugu; Survey Site LB; Sur- vey Site LS; Survey Site SEM. 3. Ajer Dingin; Labuhanbajau; Sinabang. 4. Lasia, Pulau. 5. Tuangku, Pulau. Nias, Pulau: 6. Lafau. 7. Soliga. 8. Hilisimaetano. 9. Samasama. 10. Siaba, Teluk. 1 1 . Mursala, Pulau. 12. Tanahmasa, Pulau. 13. Tanahbala, Pulau. Java (J), including nearby islands 1. Cilacap. 2. Kalipucang; Pangandaran. 3. Tjeringin. 4. Tasikmalaya. 5. Tilu, Gunung. 6. Malabar; Preanger district. 7. Bandung, 2000 ft; Bandung, near. 8. Ciwangi; Singkil, Gunung; Takokak Reserve. 9. Bantargebang; Pelabuhanratu, Teluk. 10. Jasinga. 11. Cihara. 12. Sarongen; Tamandjaija. 13. Camara. 14. Handeuleum, Pulau; Niur; Panaitan, Pulau; Peu- cang, Pulau; Ujungkulon, Suaka Margasatwa. 15. Danau, Rawa. 16. Gunung Halimun Reserve. 17. Bogor; Depok; Salak, Gunung. 18. Gede, Gunung; Pangrango, Gunung. 19. Cikarang forest. 20. Cikujang; Pasir Carolina. 21. Indramayu; Jatibarang. 22. Majalengka; "P. Sember." 23. Cirebon; Ciremay, Gunung; Linggajati. 24. Baturaden; Kaligoea. South China Sea (SCS) Pinang, Pulau [2]; Redang, Pulau. Acheh, Pulau. Pulau Tioman: Camp II, Juara side; Juara, Telok; Kampong Mukut; Nipah, Telok, vicinity; Sedagong; Tekek. Pemanggil, Pulau. Aur, Pulu. Tinggi, Pulau. Sentosa Island; Singapore Island: Bukit Timah Na- ture Reserve; Changi; MacRitchie Reservoir Nature Reserve; northwestern part; Pandan, Sungai; Pung- gol; Sembawang, Sungai; Singapore Botanical Gar- dens; western part. FIELDIANA: ZOOLOGY 8. Pulau Bintan: north coast; Pasir Panjang; Sungei Biru. 9. Mentigi, Pulau Mapur. 10. Tanjong Sauh, Pulau Batam. 1 1 . Galang, Pulau; Nguwal, Pulau. 12. Bakung, Pulau; Sebangka, Pulau. 13. Lingga, Pulau. 14. Selayar, Pulau. 15. Bulan, Pulau, south. 16. Durian, Pulau; Durian-kecil, Pulau; Sugi, Pulau; Ungar, Pulau. 17. Karimun-kecil, Pulau; Selatbliat, Pulau Kundur; Pulau Karimun: Mensuda Bay; Monos; Pemeral; Pongka, Kampung. Bangka, Pulau: 18. Jebus; Muntok; Pamuja, Tanjung; Rengsam, Tan- jung; Simpang. 19. Sungaiselan. 20. Batu, Tanjung, Pulau Belitung. 21. Serutu, Pulau. 22. Pai, Teluk, Pulau Karimata. 23. Pelapis Tengah, Pulau; Penebangan, Pulau. 24. Lemukutan, Pulau. 25. Pejantan, Pulau. 26. Benua, Pulau; Jela, Pulau; Selintang, Pulau; Tam- belan Besar, Pulau; Uwi, Pulau. 27. Airabu, Pulau; Piling, Pulau. 28. Jemaja, Pulau. 29. Telaga, Pulau. 30. Matak, Pulau; Mubur, Pulau; Siantan, Pulau. 31. Laut, Pulau. Natuna Besar, Pulau: 32. Ulu, Sungai. 33. Binjai, Sungai. 34. Lagong, Pulau. 35. Subi-kecil, Pulau. 36. Serasan, Pulau. Strait of Malacca (SM) 1. Burau, Pulau; Langkawi, Pulau. 2. Pulau Pinang [1]: Pantai Krachut; Penang Botanical Gardens; Penang Hill; Telok Bahang. 3. Pangkor, Pulau. 4. Pintu Gedong, Pulau. 5. Rupat, Pulau. 6. Kapos Tinggi, Pulau Bengkalis; Padang, Pulau, north coast. Sumatra (S) 1. Meulaboh vicinity. 2. Meulaboh, ca. 35 km NW. 3. Meulaboh, ca. 60 km NW. 4. Banda Aceh, ca. 40 km S and 45 km S. 5. Banda Aceh, near and ca. 20 km SSW. 6. Banda Aceh, ca. 30 km ENE. 7. Banda Aceh, ca. 85 km ESE. 8. Takengon, ca. 40 km NNW. 9. Bur ni Bebuli; Takengon, ca. 15 km NNW. 10. Lhokseumawe, 80 km ESE. 1 1 . Lesten. 12. Gunung Leuser Reserve. 1 3. Alas, Lae, between Agusan and Ketambe Research 14. 15. 16. 17. 18. 19. 20. 21, 22. 23. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34. 35. 36. 37. 38. 39. 40. 41 42, 43, 44 45 46 47 Station; Goenoengsetan-Meloewak; Gumpang, near; Ketambe Research Station; Ketambe, ca. 5 km S and 10 km S. Ketambe, ca. 30 km SSE. Alas, Lae, between Muara Setulen and Lae Renun, between Lae Renun and Bengkong River, and be- tween Bengkong River and Gelombang; Bukit La- wang, ca. 35 km SW; Renun, Lae. Alas, Lae, between Gelombang and mouth of river. Singkel vicinity. Tamiang. Medan, ca. 75 km NW; Pangkalansusu; Serangjaya- hilir; Sikundur. Bukitlawang; Bukitlawang, Area I and Area II; Bun- gara, Area III; Ober Langkat district; Serapit Tand- jung; Tandjung Bringin; Tandjung Butus; Unter Langkat district. Batangkuis; Belawan; Deli, Sungai; Labbuhandeli vicinity; Medan, forest near, near sea level, and vi- cinity; Medan, 20 km N; Medan-Siantar; Tanjung- morawa; Terbanjawan. Serdang district. Batu Bara district; Lauttador; Padang/Bedagei dis- trict; Paguruan; Tandjung; Tebingtinggi vicinity, ca. 25 km SSE and ca. 35 km ESE. Tebingtinggi, ca. 55 km ESE and ca. 65 km and 75 km SE. Bukitlawang, ca. 65 km S. Medan, ca. 100 km SSW. Dolok Tinggi Radja Reserve. Medan, ca. 105 km SSE. Tebingtinggi, ca. 75 km SSW. Pematangsiantar; Tebingtinggi, ca. 60 km SSW. Batang Garut. Tapanuli, Teluk. Padangsidempuan vicinity. Padangsidempuan, ca. 45 km NNW. Rantauprapat vicinity. Telukpanji. Bangko, ca. 15 km, 20 km, and 25 km ENE, ca. 20 km N, and ca. 30 km NNW. Rokan-kanan, Sungai. Bangko, ca. 80 km S. Bangko, ca. 90 km SSE; Pagansan; Pap-ka; Siak Co- patta; Sumatra, east-central. Tapung-kanan, Sungai. Gasip, Sungai, ca. 10 km and 20 km above mouth; Mandau, Sungai; Mempura; Perawang, 4 km NW; Siak, Sungai, near Pekanbaru, ca. 1 5 km, 30 km, 45 km, 60 km, and 75 km below Pekanbaru, ca. 60 km, 48 km, 36 km, 24 km, and 1 2 km above mouth, and near mouth. Lubuksikaping; Pinagar; Pinagar, ca. 5 km SW and ca. 20 km E. Bukit Cangang. Bukittinggi and vicinity and ca. 20 km S; Maninjau; Payakumbuh. Pajo. Lubukminturun area; Meru, Gunung; Padang; Pangkalan; Pasar Usang area; Sungailundang; Ta- rusan, Teluk. Kambang. Siulakderas. Sandaran Agong. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 51. 52. 53. 54. 55. 56. 57. 58. 59. 60. 61. 62. 63. 64. 65. 66. 67. 68. 69. 70. 71. 72. 73. 74. 75. 76. 77. 78. 79. 80. 81. 82. 83. 84. 85. 86. 87. Telukkayubutih vicinity, ca. 15 km E, ca. 15 km NNE, ca. 1 5 km and 30 km WNW, ca. 30 km and 45 km ESE, and ca. 60 km SE. Kerinci, Gunung, northern foot; Pakan Selasa and ca. 10 km SSE. Sijunjung. Landai vicinity. Japura. Indragiri district. Kateman, Sungai. Indragiri, Sungai. Jambi, ca. 60 km NNW. Jambi, ca. 90 km ENE. Musi, Air, ca. 30 km and 1 5 km above Palembang, near Palembang, and ca. 12 km, 24 km, 36 km, 48 km, 60 km, 72 km, and 84 km below Palembang; Palembang district; Palembang, ca. 30 km, 40 km, 45 km, 50 km, 65 km, 70 km, 80 km, and 100 km N, ca. 30 km and 60 km NNE, ca. 40 km, 60 km, 65 km, and 70 km NNW, ca. 45 km and 60 km NW, and ca. 35 km SW. Harileko, Batang, ca. 12 km, 24 km, 36 km, 48 km, 60 km, 72 km, 84 km, and 96 km above mouth; Musi, Air, ca. 15 km, 30 km, 45 km, 60 km, 75 km, and 90 km above mouth of Batang Harileko; Musi, Air, ca. 50 km, 70 km, and 85 km above Palembang. Palembang, ca. 95 km and 100 km SW and ca. 90 km WSW. Lubuklinggau and vicinity and ca. 1 5 km N. Lubuklinggau, ca. 40 km NNW. Barisan Selatan. Kelabong, Bukit, vicinity. Jeleket vicinity and ca. 10 km W. Bengkulu. Bengkulu, ca. 40 km ENE. Sanggul, Bukit. Lubuklinggau, ca. 60 km SE. Baturaja, ca. 70 km WNW and ca. 75 km NW; Lahat. Baturaja, ca. 20 km and 30 km NNE. Palembang, ca. 60 km, 65 km, and 70 km SW. Muaradua. Suka Bandjar, NE and SE. Bukit Barisan Selatan, Taman Nasional. Wonosobo. Lampung, Propinsi. Kotabumi. Moro Batin and SSE; Telukbetung. Sukadana. Sungsan, ca. 50 km N; Kambas, Wai. Sungsan, ca. 20 km NNE. Kiambang. Kalianda. West Malaysia (WM) 1 . Kampong Titi Tinggi. 2. Perlis, Sungai. 3. Perangin, Bukit. 4. Pong. 5. Larut, Daerah; Ulu Ijok. 6. Hantu, Tanjong. 7. Cameron Highlands; Kinta, Daerah. 8. Telom, Sungai. 9. Berangkat, Gunong, east. 10. Terengganu, Sungai, vicinity. 1 1 . Kenyam, Sungai; Negara, Taman; Tahan, Sungai; Tembeling, Sungai. 1 2. Sungai Tekam Forestry Concession. 13. Berapit, Bukit, east; Kampong Cherok Paloh; Kam- pong Tanah Puteh; Kuantan; Pekan vicinity. 14. Kertau, Bukit. 1 5. Benom, Gunong; Benom, Gunong, NE slope; Kuala Lompat; Kuala Lompat Post, 0-1 km NW and 0- 2 km W; Lompat, Sungai, ca. 3 km and 4 km ENE of Kuala Serloh; Lompat, Sungai, ca. 3 km, 4 km, and 6 km W of Kuala Lompat Post; Patong, Bukit, ca. 1 km W; Tens, Sungai. 16. Keroh Forest Reserve; Telok Anson. 17. Morib. 18. Changkat Mentri; Cherakah, Bukit; Dusun, Sungai; Kampong Rantau Panjang; Kampong Sungai Buloh; Kuala Selangor; Kuala Selangor estuary; Lima Belas Estate; Pacific Tin; Port Swettenham vicinity; Tengi, Sungai; Tunggal, Bukit. 19. Bunga Buah; Damansara; Dusun Tua; Gaik Liew Estate; Kelang Road; Kuala Lumpur vicinity; Nan- as, Bukit; Templer Park; Ulu Gombak Forest Re- serve. 20. Pasoh Forest Reserve. 21. Lesong. 22. Endau, Sungai, vicinity; Kuala Rompin; Tanjong Panjair. 23. Telapak Burok, Gunong. 24. Menyala, Sungai; Port Dickson; Tanjong Tuan. 25. Kuala Pilah vicinity; Nuri Valley. 26. Melaka. 27. Nyalas. 28. Selai, Sungai. 29. Sekol, Sungai. Absence reports a. Berhala, Pulau. b. Jarak, Pulau. c. Sembilan Kepulauan. d. Babi, Pulau. e. Bangkaru, Pulau. f. Siberut, Pulau. g. Sipura, Pulau. h. Pagai Utara, Pulau. i. Pagai Selatan, Pulau. j. Enggano, Pulau. k. Anak Krakatau, Pulau; Rakata, Pulau; Rakata- kecil, Pulau; Sertung, Pulau. 1. Sebesi, Pulau. m. Datuk, Pulau. n. Temaju, Pulau. Absence report not represented because of map con- gestion: Sanglang-besar, Pulau (0°36'-0°38'N, 103°41'- 103°42'E). FIELDIANA: ZOOLOGY Sabah (Sab) Fig. 2C. Detail map of non-Philippine localities of Macaca fascicularis, southeastern quadrant. (Philippine lo- calities are mapped in Fooden, 1991, p. 2.) For documentation, see Gazetteer (Appendix 2). Abbreviations in parentheses are those used in Gazetteer locality codes. Brunei (B) 1. Bandar Seri Begawan; Brunei Bay area; Kampong Menuggol. Unmapped Brunei locality: Kuala Belalong Field Studies Centre (4°33'N, 1 15°08'E; for details, see Gaz- etteer, Appendix 2). Java (J), including nearby islands 1-24. See Figure 2B. 25. Candiroto. 26. Semarang. 27. Karimunjawa, Pulau; Kemujan, Pulau. 28. Pangonan. 29. Gedangan. 30. Ngawi. 31. Tamansari. 32. Tulungagung. 33. Kawarasan; Kediri district; Margomulio, Gunung; Manggis. 34. Wonokojo. 35. Meru Betiri Nature Park. 36. Mendit; Tengger, Pegunungan. 37. Pulau Doem area. 38. Bawean, Pulau. 39. Madura, Pulau. 40. Kangean, Pulau. 4 1 . Baluran Game Reserve. 42. Banyuwangi. 43. South Banyuwangi Nature Park. Kalimantan (K), including nearby eastern islands 1 . Cabang Panti Research Station; Gunung Palung Na- ture Reserve; Kampung Baru Study Area. 2. Ambawang, Sungai. 3. Senoeang. 4. Perbuah. 5. Roema Manoeal. 6. Semitau. 7. Sintang. 8. Riam. 9. Sekonyer Kan an River. 10. Tanjung Puting Reserve. 1 1 . Katingan, Sungai, right bank, 1 40 km above mouth. 12. Parit. 13. Katingan, Sungai, left bank, 160 km above mouth. 14. Katingan, Sungai, left bank, 200 km above mouth. 15. Kahayan, Sungai, right bank, 180 km above mouth of Sungai Rungan. 16. Liang Koeboeng. 17. Putussibau. 18. Poelau. 19. Busang, Sungai, left bank, 6 km above mouth, and FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS right bank, 1 km above mouth; Julai, Sungai, left bank, 1 km below mouth of Sungai Busang and 2 km and 4 km above Muarajuloi; Julai, Sungai, right bank, 1 km above mouth of Sungai Busang, 3 km below mouth of Sungai Busang, and 6 km above Muarajuloi; Murung, Sungai, left bank, 1 km below mouth of Sungai Danau, 2 km above mouth of Sun- gai Beriwit, 2 km above mouth of Sungai Turusan, 2 km below mouth of Sungai Turusan, and 8 km above Muarajuloi; Murung, Sungai, left bank, at mouth of Sungai Beriwit, 1 km below mouth of Sungai Beriwit, 5 km above mouth of Sungai Tu- rusan, and 1 km below mouth of Sungai Turusan; TelukJolo, 12 km N. 20. Kahayan, Sungai, left and right banks, 1 20 km above mouth of Sungai Rungan. 2 1 . Kahayan, Sungai, left bank, 60 km above mouth of Sungai Rungan. 22. Rungan, Sungai, right bank, 50 km above mouth. 23. Rungan, Sungai, right bank, above mouth; Kahay- an, Sungai, left bank, above and below mouth of Sungai Rungan, and right bank, below mouth of Sungai Rungan. 24. Kapuas, Sungai, right bank, 25 km above mouth. 25. Barito, Sungai; Kambang, Pulau; Kaget, Pulau. 26. Pelaihari; Pleihari Tanah Laut Game Sanctuary. 27. Karangintan. 28. Rantau. 29. Hantakan; Telang. 30. Tanjung. 31. Buntok. 32. Kampong Hadjak; Muaratewe. 33. Purukcahu. 34. Tibang, Mt. 35. Badang. 36. Sungai Kayan-Sungai Mentarang Nature Reserve. 37. Sembakung, Sungai. 38. Long Peleben; Long Pangian. 39. Kaboerau. 40. Karangtigau, Tanjung. 41. Maratua, Pulau. 42. Berau, Sungai; Birang, Sungai. 43. Merah. 44. Karangan, Sungai; Pelawan, Sungai. 45. Kariorang. 46. Kutai Nature Reserve, northeast corner; Sengata; Sengata, Sungai, Kutai Reserve; Sengata, Sungai, at Mentoko camp and 1 km, 2 km, 3 km, 5 km, 7 km, 9 km, 1 1 km, and 1 3 km below; Sengata, Sungai, below Sengata village. 47. Juyan, Sungai, right bank, near mouth; Mentoko Research Center; Sengata, Sungai, right bank, 1 km below mouth of Sungai Nubung. 48. Kembangjanggut. 49. Goson Djerong, near; Jembayan, Sungai; Karang- mumus, Sungai; Loa Bambam; Mahakam, Sungai, left bank, 1 km and 4 km above Sebulu; Mahakam, Sungai, north bank, above Samarinda; Mahakam, Sungai, right bank, 8 km above Sebulu and 2 km below Sebulu; Tangarveng Island. 50. Sepaku, Sungai. 5 1 . Klumpang, Teluk. 52. Sebuku, Pulau. 53. Matasiri, Pulau. Lesser Sundas (LS) Bali, Pulau, including nearby islet: 1. Bali Barat National Park; Banjoe Wetan; Gilima- nuk; Pulaki, Tanjung; Sendang; Trima, Teluk. 2. Jembrana. 3. Bangli, near; Batur, Gunung; Botanical Gardens; Bratan, Danau; Bratan, Gunung; Buyan, Danau- Danau Bratan region; Catur, Gunung; Desa Poet- jang; Gitgit; Kukuh; Sangeh; Ubud. 4. Penida, Nusa. Lombok, Pulau: 5. Pengsong, Gunung; Pusuk, Gunung; Suranadi. 6. Kuta. 7. Pusuk forst; Rinjani, Gunung; Sewela. Sumbawa, Pulau, including nearby islet: 8. Ai Beta; Batudulang; Maman; Semongkat. 9. Moyo, Pulau. 10. Ampang area. 1 1 . Dompu; Oo vicinity. 12. Ntori; Raba; Rite. Flores, Pulau, including nearby islets: 13. Ginggo, Teluk, Pulau Rinca; Kode, Nusa; Mangia- tan, Pulau; Mbura, Pulau Flores; Nanga Look, Pulau Flores; Seraya Besar, Pulau. 14. Sano, Wai. 15. Rana Mese. 16. Bari. 17. Reo. 18. Pota. 19. Sika. 20. Solor, Pulau. 21. Adonora, Pulau. Sumba, Pulau: 22. Payeti-Kambaniru. 23. Mao Marroe. Timor, Pulau, including nearby islets: 24. Kambing, Pulau; Oeassa vicinity, Pulau Semau. 25. Amarassie. 26. Benu; Bokong; Kuatnana; Lelogama; Timau, Fatu. 27. Nikiniki. 28. Mutis, Gunung. 29. Fatuboi. Philippines Unnumbered locality: Bongao Peak, Bongao Island (5°01'N, 1 19°45'E; for details, see Gazetteer, Appendix 2). Sabah (Sab) 1 . Padas Bay. 2. Rayoh. 3. Papar. 4. Bongkabong; Rugading. 5. Talibang; Tuaran; Tuaran-Kampong Tenghilan Road. 6. Keningau. 7. Garau; Kampong Bundu Tuhan; Kampong Kiau; Kampong Kiau-Tenampok Pass; Kenokok; Kiau- lan; Kinabalu, Mount; Kinabalu National Park; Lumu Lumu; Ranau; Tempasuk, Sungai; Tenom- pok; Tinonkok. 10 FIELDIANA: ZOOLOGY Sex and Age Variation Geographic Variation Dorsal pelage color is generally similar in adult, subadult, and juvenile males and females, but pel- age in adult males tends to be longer and sleeker than in other age/sex classes. In newborn infants, dorsal pelage is blackish (on head, trunk, limbs, and tail), and the facial skin is bare, unpigmented, and pinkish (Dang, 1977, p. 11; Aldrich-Blake, 1980, p. 147; Crockett & Wilson, 1980, p. 170; Fittinghoff & Lindburg, 1980, p. 189; Wheatley, 1982, p. 205; Koyama, 1985, p. 106). The gradual transition from blackish neonatal pelage to grayish or brownish postneonatal pelage begins at about age 2-3 months, which is the age when deciduous canines and first molars erupt (Table 1 3) and in- fants begin to obtain some of their food indepen- dently (see below, Reproduction). The change of pelage color occurs first on the appendages, then on the trunk, and last on the crown. In specimens examined, the transition is complete well before eruption of the permanent first molars (age ca. 15 mo; Table 14). Dorsal Pelage Color Saturation— To in- vestigate geographic variation in dorsal pelage col- or saturation in M. fascicularis, specimens ex- amined have been compared with standard skins, as in a previous study of Philippine M. fascicularis (Fooden, 1991, p. 5). In the previous study, the range of standard saturation index (SI) values ex- tended from 1 .0 (pale yellowish brown) to 3.0 (dark brown). In the present study, this range extends from 0.5 (buffy; fmnh 9965 1 , Thailand: Kata Taek) to 4.0 (blackish; usnm 114168, Indonesia, P. Si- meulue: Sibaboh, Lugu) (Fig. 6). Skins of infants (specimens with deciduous teeth only) are exclud- ed from the analysis. Also excluded is the aber- rantly albinistic (pale ochraceous) skin of an adult female (amnh 107094) collected at Perbuah, northwestern Kalimantan; eight other specimens from the same locality are normally colored. In the core area of distribution of M. fascicularis (see above, Geographic Distribution), dorsal pel- age color in 640 postinfantile specimens varies 8. Balambangan, Pulau. 9. Banggi, Pulau, south, and Karakit; Maliangin Besar, Pulau. 10. Sabor, Pulau Banggi. 1 1 . Malawali, Pulau. 12. Sabatik, Pulau. 13. Ulu Segama Reserve. 14. Bukit Garam. 15. Lokan, Sungai. 16. Betotan; Lungmanis Station. 17. Labuk Road; Sandakan, 8 mi (= 13 km) W; Sibuga Besar, Sungai. 18. Sapagaya Forest Reserve. 19. Abai; Kinabatangan, Sungai; Trusan Kinabatangan. 20. Dewhurst Bay; Kretam Besar, Sungai; Kretam Kechil, Sungai. 21. Lahad Datu; Segama, Sungai. 22. Darvel Bay. 23. Tawau Hills National Park. 24. Tawau. Sarawak (Sar) 1. Samunsam Wildlife Sanctuary. 2. Penrissen, Gunung. 3. Paku Cave. 4. Bako National Park; Kuching; Kuching, 10th mile; Sarawak, Sungai, mouth. 5. Bukar, Sungai; Entawa, Tanjong. 6. Pelandok, Sungai. 7. Simunjan, Sungai. 8. Lingga. 9. Paku, Saribas. 10. Jumpit. 1 1 . Lanjak-Entimau Orang-Utan Sanctuary (proposed). 12. Belaga. 13. Dulit, Bukit. 14. Selikan, Bukit. 15. Similajau National Park. 16. Niah Caves. 17. Niah National Park. 18. Baram, Batang; Baram district; Miri; Miri, Sungai; Miri district. 19. Long Ekang. 20. Kalulong, Bukit. 2 1 . Gunong Mulu National Park; Melinau Gorge; Mulu, Gunung. 22. Punang, Sungai. Absence reports a. Manuk Manka Island, Philippines (see Fooden, 1991, p. 40). b. Masalembu Besar, Pulau. c. Sangeang, Pulau. d. Banta, Pulau; Bugis, Gili; Lawadarat, Gili; Lawa- laut, Gili; Longos, Nusa; Misa, Pulau; Papagaran- besar, Pulau; Sababi, Pulau; Sabajor Besar, Pulau; Sebolon Besar, Pulau; Siaba Besar, Pulau; Tatawa, Pulau. e. Kelapa, Pulau; Komodo, Pulau; Langkoi, Pulau; Lengah, Pulau; Mbarapu, Nusa; Motang, Gili; Pa- dar, Pulau; Padar-kecil, Pulau. f. Lomblen, Pulau. g. Pantar, Pulau. h. Alor, Pulau. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 11 120 Fig. 3. Approximate extent of dry land in Southeast Asia during last glacial maximum, ca. 18 Ka; estimate based on present-day 120-m bathymetric line (Heaney, 1986, p. 137; 1991b, p. 147; U.S. Defense Mapping Agency bathymetric charts Nos. 63020, 1993; 63400, 1990; 71000, 1975a; 71006, 1975b; 93036, 1985; 94004, 1986). from buffy (SI = 0.5) to dark brown (SI = 3.0) (Tables 1 , 2); relatively few of these specimens (4.5%) are darker than medium brown (SI = 2.0). Pelage color in core-area specimens averages pal- est in the Indochinese Peninsula (mean SI = 0.99, n = 93), becomes darker southward and reaches maximum saturation in Sumatra (mean SI = 1.38, n = 178) and Borneo (mean SI = 1.47, n = 239), and then becomes somewhat paler farther south- ward in Java (mean SI = 1.26, n = 77). Within the Indochinese Peninsula, pale specimens are particularly concentrated near the northwestern margin of the species range, north of 1 4°N between 98°E and 102°E (mean SI = 0.73, n = 42). In all parts of the core area, variation in dorsal pelage saturation is relatively great, and overlap in sat- uration between samples from different parts of the core area is extensive. For example, a pale golden brown adult male collected in western Thailand (fmnh 99642, Ban Tamrong Phato) al- most perfectly matches an adult male collected in southern Sumatra (amnh 102765, Muaradua), and a pale yellowish brown adult male collected in east-central Sumatra (usnm 1131 69, Indragiri, Sungai) matches an adult male collected in western Java (usnm 156291, Bantargebang). Fringing-island specimens examined include 49 1 postinfantile skins— 234 collected in 71 shallow- water islands and 257 collected in 27 deep-water islands (Appendix 3). Although dorsal pelage sat- uration in most fringing-island specimens broadly overlaps that in core-area specimens, mean SI in fringing-island samples tends to exceed that in neighboring core-area samples (53 of 71 shallow- water fringing-island samples, 23 of 27 deep-water fringing-island samples). Strikingly dark fringing-island samples (con- spicuously darker than any other M. fascicularis samples examined) have been collected in six deep- water islands that constitute three geographic units: (1) Nicobar Islands (Katchall, Little Nicobar, Great Nicobar), northwest of Sumatra (mean SI = 3.9, n = 8); (2) P. Simeulue and P. Lasia, west of Su- matra (mean SI = 4.0, n = 15); and (3) P. Maratua, 12 FIELDIANA: ZOOLOGY Fig. 4. Core area, shallow-water (< 120 m) fringing islands, and deep-water (> 120 m) fringing islands inhabited by Macaca fascicularis (cf. Fig. 3). For names of islands, see Figure 2 and Fooden (1991, p. 2). east of Borneo (mean SI = 4.0, n = 4). The pe- ripheral and disjunct distribution of these intense- ly saturate populations of M. fascicularis was not- ed previously by Kellogg (1944, p. 76). Dorsal pelage also is relatively dark in many islands of the Philippine Archipelago (deep-water); in west- ern, northern, and eastern Philippine islands, mean SI varies from 2.4 to 3.0 (cf. Fooden, 1991, p. 5). A brief, equivocal comment alluding to pelage sat- uration in M. fascicularis in deep-water P. We, north of Sumatra, has been published by Scheff- rahnetal. (1994, p. 136). The darkest shallow-water fringing-island sam- ple was collected in P. Karimunjawa, north of Java (mean SI = 2.7, n = 6; cf. Sody, 1949, p. 132). A shallow-water fringing-island sample from Ko Khram Yai, in the Inner Gulf of Thailand, also is conspicuously darker (mean SI = 1 .8, n = 10) than neighboring core-area specimens (cf. Kloss, 1919c, p. 347). Inconclusively small shallow-water island samples of dark specimens have been collected in P. Redang, east of the Malay Peninsula (mean SI = 2.5, n = 2), Ko Na Ka Yai, west of peninsular Thailand (SI = 2.5, n = 1), and P. Belitung, be- tween Sumatra and Borneo (mean SI = 2.3, n = 2). Dorsal Pelage Erythrism— Dorsal pelage er- ythrism in M. fascicularis has been investigated by comparing specimens examined with selected standard specimens, in a procedure parallel to that used in investigating saturation (see above). The range of standard erythrism index (EI) values ex- tends from 1.0 to 3.0, as follows: EI = 1.0, noner- ythristic, color of pale hair bands pale yellowish (fmnh 99651, Thailand: Kata Taek); EI = 2.0, moderately erythristic, pale hair bands golden (fmnh 33505, Vietnam: Ho Chi Minh City); and EI = 3.0, intensely erythristic, pale hair bands ru- fescent (fmnh 62913, Philippines: Palawan, Puer- to Princesa). Specimens marginal in erythrism be- tween primary standards have been assigned EI values of 1 .5 or 2.5. Infant skins are excluded from the analysis. In core-area samples, erythrism in M. fascicu- laris averages relatively high in Sumatra and in the adjacent Malay Peninsula, south of 10°N (Ta- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 13 Fig. 5. External characters in Macaca fascicularis— left, Thailand: Lop Buri, Sarn Pra Karn, adult male; right, Indonesia: P. Bali, Sangeh, adult female and adult male. bles 3, 4). Of 645 postinfantile core-area skins ex- amined, 24 are intensely erythristic (EI = 3.0); all of these intensely erythristic specimens originated either in Sumatra or in the adjacent Malay Pen- insula. Within Sumatra and the Malay Peninsula, intense erythrism apparently is an individual vari- able, not a local variable; in local samples (n > 1) that include intensely erythristic specimens, the frequency of such specimens varies from 0. 1 2 (Su- matra: Batangkuis; n = 8) to 0.50 (West Malaysia: Port Dickson; n = 4). Intense erythrism (EI = 3.0) occurs in relatively pale core-area specimens (zrc 4-075, West Malaysia: Port Dickson, SI = 1.0) as well as in darker specimens (zsbs C- 1 76, Sumatra: Kampong Baru, SI = 2.5). The high incidence of intense erythrism in M. fascicularis in the Malay Peninsula has been indicated previously by Kloss (1919c, p. 347) and Chasen (1940a, p. 67). Aggi- marangsee (1992, p. 119) reported on erythrism in a provisioned population of M. fascicularis at Kosumphi Forest Park, northeastern Thailand. In fringing-island samples, erythrism data are available for 234 postinfantile specimens collected in 7 1 shallow-water islands and 2 1 8 postinfantile specimens collected in 25 deep-water islands (Ap- pendix 4). In shallow-water fringing-island sam- ples, mean EI tends to exceed that in neighboring core-area samples (mean EI greater in 48 fringing- island samples, mean EI less in 1 6 fringing-island samples, mean EI equal in 7 fringing-island and core-area samples). Geographic variation of ery- thrism in shallow-water fringing-island samples generally parallels that in neighboring core-area samples: 1 3 intensely erythristic specimens (EI = 3.0) have been collected in 10 shallow- water fring- ing islands (Bintan, Bulan, Durian, Karimun, and Nguwal, in the Riau Archipelago, east of Sumatra; Redang, Siantan, and Tioman, east of the Malay Peninsula; Pinang [1] and Pintu Gedong, west of the Malay Peninsula; and Singapore); all 10 shal- low-water fringing islands from which intensely erythristic M. fascicularis specimens have been collected are adjacent to Sumatra or the Malay Peninsula, where intensely erythristic core-area specimens have been collected. In these 1 0 fringing islands, the frequency of intensely erythristic spec- imens varies from 0.09 (Tioman, n = 1 1) to 1.00 (Redang, n = 2). 14 FIELDIANA: ZOOLOGY Fig. 6. Dorsal pelage color saturation extremes in Macaca fascicular is— fmnh 99651 (left), adult male, Thailand: Kata Taek, buffy (saturation index value, 0.5); usnm 1 14168 (right), adult male, Indonesia: P. Simeulue, Lugu Sibaboh, blackish (SI value, 4.0). Scale bar 10 cm. In deep-water fringing islands, erythrism is rel- atively high in most islands of the Philippine Ar- chipelago (Appendix 4) and is particularly high in Palawan, as noted previously (Fooden, 1991, p. 8). Erythrism is low in the very dark insular pop- ulations west of Sumatra (Appendix 4), and ery- thrism also is generally low in the deep-water Less- er Sunda Islands, as in nearby Java (Tables 3, 4). Crown Color— As previously indicated, the crown in M. fascicularis usually is more brightly colored than the back; however, in 63 of 1,103 postinfantile specimens examined (Table 5), the crown is darker than the back (Fig. 7) as a result of reduction or absence of pale bands in crown hairs. Darkness of the crown in these specimens varies from a dilute blackish wash to an irregular blackish streak to a clearly defined blackish patch. In dark-crowned specimens, the periphery of the crown patch usually is darker than the center. Of 648 core-area postinfantile specimens ex- amined, dark crowns are clearly defined in 1 and weakly defined in 13 (Table 5, Appendix 5). Thir- teen of these 14 dark-crowned specimens, includ- ing the one with a clearly defined dark crown, were FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 15 Table 1. Table 2). Dorsal pelage color saturation in core-area samples of Macaca fascicularis1 : summary of variation (cf. N Frequencies (%) at SI values Mean SI Sample area 0.5 1.0 1.5 2.0 2.5 3.0 value Indochinese Peninsula2 93 41.9 29.0 18.3 10.8 0.99 Malay Peninsula3 53 11.3 34.0 45.3 9.4 1.26 Sumatra 178 4.5 23.6 67.4 1.7 1.7 1.1 1.38 Borneo 239 0.8 43.9 24.3 21.8 9.2 1.47 Java 77 10.4 41.5 35.1 10.4 2.6 1.26 All 640 9.8 35.0 38.5 12.2 4.2 0.3 1.33 1 Excludes infants. 2 Includes Isthmus of Kra north of 1 0°N. 3 Includes Isthmus of Kra south of 10°N. collected in the Indochinese Peninsula or adjoin- ing Isthmus of Kra north of 10°N (n = 94); one, with a weakly denned dark crown, was collected in Borneo (n = 242). The Indochinese specimen with a clearly defined dark crown (bm(nh) 1881. 6. 30. 2, adult male) was collected in southern Vietnam ("Cochin China"; cf. Elliot, 1909, p. 252; Kloss, 1926, p. 358). In fringing-island samples, most dark-crowned specimens also were collected in the vicinity of the Indochinese Peninsula and northern part of the Isthmus of Kra (Table 5, Appendix 5). Of 452 Table 2. Dorsal pelage color saturation in core-area Macaca fascicularis (n = 506)1 : geographic variation of mean value in 2-degree latitude-longitude blocks (sample size indicated by italicized figures in parentheses) (cf. Table 1). Longitude (°E) 96- 98- 100- 102- 104- 106- 108- 110- 112- 114- 116- 118- Latitude 98 100 102 104 106 108 110 112 114 116 118 120 16-18 (°N) 1.0 (1) 0.6 (14) 1.0 (1) 1.5 (1) 14-16 0.8 (21) 0.8 (7) 1.0 (1) 12-14 1.1 (22) 1.0 (2) 1.0 (3) 0.5 U) 1.0 (1) 10-12 1.6 (6) 1.5 (12) 8-10 0.8 (8) 1.0 (1) 6-8 1.1 (6) 1.0 (3) 1.3 (20) 4-6 1.0 (1) 1.5 (6) 1.0 (1) 1.6 (8) 1.4 (16) 1.2 (29) 2-4 1.5 1.4 1.4 1.4 1.6 1.6 1.5 (1) (66) (76) (11) (12) (8) (2) 0-2 1.3 (2) 3.0 (1) 1.5 (32) 1.8 (6) 1.5 (1) 1.7 (3) 0-2 (°S) 1.4 (76) 1.0 (l) 1.5 (1) 1.2 (5) 2.1 (4) 1.7 (3) 2-4 1.0 (1) 1.2 (9) 1.8 (2) 1.4 (26) 1.4 (4) 4-6 1.1 (17) 6-8 0.5 (1) 1.1 (23) 1.3 (32) 1.3 (4) 1.0 (/) 8-10 1.0 (3) Excludes 1 34 imprecisely localized specimens that are included in Table 1 . 16 FIELDIANA: ZOOLOGY Table 3. Dorsal pelage erythrism in core-area Macacafascicularis1: summary of variation (cf. Table 4). N Frequencies (%) at EI values Mean EI Sample area 1.0 1.5 2.0 2.5 3.0 value Indochinese Peninsula2 93 69.9 5.4 23.6 1.1 1.28 Malay Peninsula3 53 37.7 17.0 30.2 5.7 9.4 1.66 Sumatra 180 23.3 8.3 56.7 1.1 10.6 1.84 Borneo 241 46.9 34.4 18.3 0.4 1.36 Java 78 57.7 23.1 19.2 1.31 All 645 44.2 20.2 30.8 1.1 3.7 1.50 Excludes infants. Includes Isthmus of Kra north of 10°N. Includes Isthmus of Kra south of 10°N. fringing-island postinfantile specimens examined, dark crowns are clearly denned in 1 7 and weakly denned in 32. Thirty-seven of the 49 dark-crowned specimens were collected in shallow-water islands adjacent to the Indochinese Peninsula or Isthmus of Kra. Of the 1 7 specimens with clearly defined dark crowns, 16 originated in three islands adja- cent to the Indochinese Peninsula: Ko Khram Yai, in the Inner Gulf of Thailand (n = 10: dark crown clearly defined, 7; weakly defined, 3); Con Son, in the South China Sea (n = 9: clearly defined, 8; weakly defined, 1); and Hon Ba, an islet adjacent to Con Son (n = 3: clearly defined, 1; weakly de- fined, 2) (cf. Kloss, 1916b, p. 32; 1919c, p. 347; Table 4. Dorsal pelage erythrism in core-area Macacafascicularis (n = 5 10)1: geographic variation of mean value in 2-degree latitude-longitude blocks (sample size indicated by italicized figures in parentheses) (cf. Table 3). Longitude (°E) 96- 98- 100- 102- 104- 106- 108- 110- 112- 114- 116- 118- Latitude 98 100 102 104 106 108 110 112 114 116 118 120 16-18 (°N) 1.0 V) 1.4 (14) 2.0 (1) 2.0 (1) 14-16 1.3 (21) 1.1 (7) 2.0 (1) 12-14 1.1 (22) 1.0 (2) 1.3 (3) 1.0 (1) 1.0 (/) 10-12 1.4 (6) 1.5 (12) 8-10 1.3 (8) 1.5 (D 6-8 1.4 (6) 2.0 (3) 1.1 (20) 4-6 2.0 (D 1.6 (6) 2.0 (1) 1.4 (8) 1.4 (16) 1.1 (29) 2-4 2.0 1.8 2.1 1.6 1.5 1.4 1.3 (D (68) (16) (11) (12) (8) (2) 0-2 1.8 (2) 2.0 (D 1.5 (32) 1.1 (6) 1.0 (/) 1.2 (3) 0-2 (°S) 2.1 (16) 1.3 (1) 2.0 (1) 1.4 (5) 1.4 (4) 2.0 (3) 2-4 1.5 (/) 1.6 (9) 2.5 (2) 1.5 (26) 1.4 (5) 4-6 1.5 (17) 6-8 1.0 (1) 1.2 (24) 1.2 (32) 1.1 (4) 1.5 (1) 8-10 1.0 (3) 1 Excludes 135 imprecisely localized specimens that are included in Table 3. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 17 Table 5. Crown color pattern in Macaca fascicularis] : summary of variation. For details, see Appendix 5. Frequencies of crown color patterns Crown with diffuse Crown with clearly Crown colored like blackish streak defined blackish N back, or brighter or wash patch2 Sample area N % N % N % Core area 648 634 97.8 13 2.0 1 0.2 Indochinese Peninsula3 94 81 86.1 12 12.8 1 1.1 Malay Peninsula4 53 53 100.0 Sumatra 179 179 100.0 Borneo 242 241 99.6 1 0.4 Java 80 80 100.0 Shallow-water fringing islands 234 190 81.2 28 12.0 16 6.8 Deep-water fringing islands 221 216 97.7 4 1.8 1 0.5 Totals 1,103 1,040 94.3 45 4.1 18 1.6 1 Excludes infants. 2 Cf. Figure 7. 3 Includes Isthmus of Kra north of 1 0°N. 4 Includes Isthmus of Kra south of 10°N. 1921, p. 76; 1926, p. 358). In Ko Khram Yai specimens, the crown patch is distinctively narrow (Fig. 7) and is notably darker peripherally than centrally (see below, M.f. atriceps). The only non- Indochinese fringing-island specimen with a clear- ly defined dark crown is one of two specimens (usnm 144666) collected in Basilan I., a deep-wa- ter island in the south-central Philippines. Al- though the crown is blackish in M. fascicularis specimens collected in P. Simeulue and P. Lasia, deep-water islands west of Sumatra, the color of the crown in these specimens does not contrast with the color of the back, which also is blackish. Lateral Facial Crest Pattern (Fig. 8)— The Fig. 7. Crown color pattern in dark-crowned Macaca fascicularis— usnm 2366 1 8 (left), adult male, Thailand: Ko Khram Yai, M.f. atriceps; usnm 357241 (right), adult male, Vietnam: Hon Ba, 3.1 km W and 0.6 km S of Ben Dam, M. f condorensis. Scale bar 2 cm. FIELDIANA: ZOOLOGY INFRAZYGOMATIC TRANSZYGOMATIC Gervais, 1854 Pocock, 1939 Fig. 8. Infrazygomatic and transzygomatic lateral facial crest patterns in Macaco, fascicularis, as depicted by Gervais (1854, p. 87) and Pocock (1939, PI. V). In both depictions, note posteriorly directed preauricular hairs in the infrazygomatic pattern and anteriorly directed preauricular hairs in the transzygomatic pattern. pale lateral facial crest in M. fascicularis is formed by the convergence of variably elongated poste- riorly directed hairs of the anterolateral facial re- gion and anteriorly directed hairs of the postero- lateral facial region. In most specimens examined, this crest sweeps upward from near the angle of the jaw to the lateral margin of the crown, passing between the eye and ear (transzygomatic crest pat- tern); less commonly, the crest is restricted to the mandibular region and terminates superiorly in a whorl low on the cheek (infrazygomatic crest pat- tern) (cf. Pocock, 1939, p. 78). In the infrazygo- matic pattern, unlike the transzygomatic pattern, hairs of the temporal region are smoothly directed posteriorly from the posterior margin of the eye to the anterior margin of the ear (anteriorly di- rected hairs are absent in this region); as a result, the infrazygomatic crest lacks the upper (preaur- icular) portion of the transzygomatic crest. Al- though the infrazygomatic pattern is relatively rare in M. fascicularis, it is common in M. mulatta, where the transzygomatic pattern occurs infre- quently (Stewart, 1933, p. 30). Transzygomatic and infrazygomatic patterns can be distinguished in the hair tracts of infants as well as in postinfants. In 819 specimens examined, the lateral facial crest pattern is transzygomatic in 728, infrazygo- matic in 83, and asymmetric (transzygomatic on FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 19 Table 6. Lateral facial crest pattern in Macacafascicularis1: summary of variation. For details, see Appendix 6. Frequencies of lateral facial crest patterns Asymmetric: infrazygomatic/ N Infrazygomatic Transzygomatic transzygomatic Sample area N % N % N % Core area 396 59 14.9 332 83.8 5 1.3 Indochinese Peninsula2 98 57 58.1 37 37.8 4 4.1 Malay Peninsula3 48 1 2.1 47 97.9 Sumatra 50 49 98.0 l4 2.0 Borneo 163 l5 0.6 162 99.4 Java 37 37 100.0 Shallow-water fringing islands 218 ll6 5.0 204 93.6 V 1.4 Deep-water fringing islands 205 138 6.3 192 93.7 Totals 819 83 10.1 728 88.9 8 1.0 1 See Figure 8; tabulation includes infants. 2 Includes Isthmus of Kra north of 10°N. 3 Includes Isthmus of Kra south of 10°N. 4 Collected at Kotabumi (sample size = 1). 5 Collected at Mt. Tibang, Kalimantan (sample size = 1). 6 Includes 10 specimens collected in islands west of the Indochinese Peninsula and Isthmus of Kra (see Appendix 6) and one geographically anomalous specimen collected in Indonesia: P. Bintan (sample size = 8), southeast of Singapore. 7 Collected in islands west of the Isthmus of Kra and Malay Peninsula (see Appendix 6). 8 Collected in the Nicobar Islands, P. Simeulue, and P. Lasia (see Appendix 6). one side and infrazygomatic on the other) in 8 (Table 6, Appendix 6). Of the 83 specimens with the infrazygomatic pattern, 5 1 originated in a re- stricted area bordering the Bay of Bengal, at the northwestern margin of the species range (Fig. 9). No specimens with the transzygomatic pattern have been collected in this area, which includes south- ernmost Bangladesh, southern Burma, nearby shallow- water islands in the Mergui Archipelago, and a small part of west-central Thailand. The eastern boundary of the restricted area that is ho- mogeneous for the infrazygomatic pattern gener- ally coincides with the mountain ranges that form the border between Burma and Thailand. Of the remaining 32 specimens with the infra- zygomatic pattern, 1 7 originated in two areas ad- jacent to the restricted area homogeneous for the infrazygomatic pattern (Fig. 9); both of these ad- jacent areas are heterogeneous for the lateral facial crest pattern. Nine of these specimens originated at five scattered localities in the central and eastern parts of the Indochinese Peninsula, immediately east of the area homogeneous for the infrazygo- matic pattern, along the northern margin of the range of M. fascicularis; seven specimens with the transzygomatic pattern also have been collected in this area (central and eastern Indochinese Pen- insula), which includes two localities where spec- imens with both crest patterns have been collected together. Eight specimens with the infrazygomatic pattern originated at three localities in the eastern and southern parts of the Isthmus of Kra, im- mediately southeast of the area homogeneous for the infrazygomatic pattern; 1 1 specimens with the transzygomatic pattern also have been collected in this area (eastern and southern Isthmus of Kra, two adjacent west-coast islands). The remaining 1 5 specimens with the infrazy- gomatic pattern originated at localities distant from the restricted area homogeneous for the infrazy- gomatic pattern. Three of these specimens origi- nated in the Nicobar Islands, northwest of Su- matra, together with five specimens with the transzygomatic pattern. Ten specimens originated in P. Simeulue and P. Lasia, west of Sumatra, together with five specimens with the transzygo- matic pattern. One specimen originated in P. Bin- tan, southeast of Singapore, together with seven specimens with the transzygomatic pattern. The final specimen with the infrazygomatic pattern originated in north-central Borneo, at Mt. Tibang, Kalimantan. 20 FIELDIANA: ZOOLOGY V ' / 1 i 1 1 Area homogeneous for infrazygomatic pattern \ N \ _/ 20 - V^ S (' ; Areas heterogeneous for lateral facial crest pattern _ ^s» \ e#^ N\ \ \ \ \ \ \ \ \ V* • v» N. • N /(»Ol 2 3 • <4sr!-(MO] --^ "No 2 / 0» i / O 3\ l f 6 x J v^ [•OI\ 15 \ /_°2 \(«X>> \2 °2 • 1 •• 1 2 (CO)! 11 °v '111 0 1 0 t— s 2°/ */' j 10 0 X £ \ / 9 IX 6> 3 Lateral facial crest pattern l^i\ • Infrazygomatic 2 *\ v~_ o Transzygomatic °° \ 3 \»» 2 si c Asymmetric 80\ ° , V°2 5 l „— ..J ^-_ I — 95 100 105 Fig. 9. Geographic variation of lateral facial crest pattern in samples of Macaca fascicularis collected in the Indochinese Peninsula, Isthmus of Kra, Malay Peninsula, and neighboring islands (see Appendix 6). Numerals indicate number of individuals represented by symbols, where this number exceeds 1 ; heterogeneous samples collected at single localities are indicated by parentheses. Of the eight specimens with the asymmetric lat- eral facial crest pattern, seven originated in the two areas heterogeneous for the lateral facial crest pattern that are adjacent to the restricted area ho- mogeneous for the infrazygomatic pattern (Fig. 9). Two of these specimens originated in the central part of the Indochinese Peninsula and five origi- nated in the eastern part of the Isthmus of Kra or in islands adjacent to the southern part of the west coast of the Isthmus of Kra. The remaining spec- imen with the asymmetric lateral facial crest pat- tern originated, somewhat incongruously, in southeastern Sumatra, at Kotabumi. External Measurements and Proportions Sex and Age Variation Collectors' measurements of wild-collected specimens of M. fascicularis indicate that head and body length (HB) in adult males (mean = 465.6 mm, n = 238) averages approximately 13% more than in adult females (mean = 412.0 mm, n = 161) (Table 7). Weight in adult males (mean = 5.36 kg, n = 69) averages 49% more than in adult females (mean = 3.59 kg, n = 46). Relative length of appendages is similar in adult males and fe- males: mean relative tail length (T/HB x 100) is 1 17.0 (extremes 66.7-149.5) in 392 adults, mean relative hind foot length (HF/HB x 100) is 28.7 (extremes 23.2-37.8) in 365 adults, and mean rel- ative ear length (E/HB x 100) is 8.0 (extremes 4.8-11.1) in 301 adults. (Note: Karrer [1970, p. 171] reported that the tail is semiprehensile in captive M. fascicularis.) Relative length of the tail and of the hind foot in infants and juveniles exceed that in fetuses, sub- adults, and adults (Table 7); similarly, relative length of the ear in infants and juveniles exceeds that in subadults and adults (unknown in fetuses). This implies that, during prenatal and early post- natal life, the growth rate of the tail, hind foot, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 21 Table 7. External measurements and proportions in age/sex classes of wild-collected Macaca fascicular is. Age/sex class2 Head and body length (mm) Relative tail length3 (T/HB x 100) Relative hind foot length3 (HF/HB x 100) Relative ear length (E/HB x 100) Weight3 (kg) Fetuses 82.5 78.8 23.7 — — 54-111 64.8-92.8 19.4-27.9 (2) (2) (2) Infants 254.8 ± 54.00 125.8 ± 24.52 33.6 ± 5.89 11.9 ± 2.51 0.76 ± 0.3424 153-380 61.8-167.3 15.3-43.7 7.6-18.3 0.24-1.36 (52) (50) (51) (42) (13) Juveniles 361.3 ± 51.90 125.7 ± 15.83 32.0 ± 2.95 8.9 ± 1.74 2.36 ± 0.917 240-490 60.3-167.8 23.6^0.0 5.1-14.1 0.90-4.54 (208) (205) (197) (183) (75) Subadults — 120.4 ± 17.66 29.7 ± 2.24 7.9 ± 1.20 — 59.7-164.1 24.7-35.7 4.8-11.1 (106) (97) (81) Subadult females 397.6 ± 32.33 120.9 ± 17.63 29.8 ± 1.88 7.6 ± 1.40 3.41 ± 0.731 326^175 59.7-159.7 26.5-33.5 4.9-9.6 2.50-4.88 (39) (39) (36) (29) (14) Subadult males 453.8 ± 44.36 120.1 ± 17.81 29.6 ± 2.43 8.0 ± 1.06 5.15 ± 0.967 356-600 63.7-164.1 24.7-35.7 4.8-11.1 3.86-7.26 (67) (67) (67) (52) (23) Adults — 117.0 ± 14.50 28.7 ± 2.42 8.0 ± 1.34 — 66.7-149.5 23.2-37.8 4.8-11.1 (392) (365) (301) Adult females 412.0 ± 36.86 116.4 ± 15. 555 28.8 ± 2.43 7.8 ± 1.28 3.59 ± 0.6906-7 315-545 70.4-148.4 23.5-36.2 5.0-10.5 2.35-5.44 (161) (159) (153) (131) (46) Adult males 465.6 ± 42.48 117.6 ± 13.378 28.7 ± 2.42 8.1 ± 1.37 5.36 ± 1.4386-7 370-630 69.2-149.5 23.2-37.8 4.8-11.1 3.40-12.00 (238) (232) (212) (170) (69) 1 Mean ± SD (where n > 2), extremes, and sample size (italicized figures in parentheses). 2 Dental specifications: infants, deciduous teeth only; juveniles, some permanent teeth erupted; subadults, M3 in females or C in males incompletely erupted (cf. Spiegel, 1952, p. 129); adults, all permanent teeth completely erupted. 3 Cf. Spiegel (1985, pp. 27, 37, 53), who recorded postnatal growth of tail length, hind foot length, and body weight in captive M. fascicularis; cf. Shimizu et al. (1994, p. 175), who reported on effect of dietary restriction on growth in captive M. fascicularis. 4 In captivity, mean birth weight is 0.34 kg for females (n = 156) and 0.37 kg for males (n = 166) (Dang et al., [1993], p. 150; cf. Berkson, 1968, p. 352). 5 Excludes bobtailed specimen zrc 4-127, West Malaysia: P. Tioman, Telok Juara (HB = 409, T = 215, T/HB x 100 = 52.6). 6 Cf. J. Suzuki and Varavudhi (1989, p. Ill) and Varavudhi et al. (1989a, p. 225), who have reported higher mean weights in artificially provisioned troops studied in Thailand (adult females, 4.03-5.86 kg; adult males, 6.80-9.33 kg; cf. Aggimarangsee, 1992, p. 139). 7Cf. Bakaretal. (1981, p. 12). 8 Excludes bobtailed specimens mzb 6483, Java: Cikujang (HB = 540, T = 250, T/HB x 100 = 46.3); and bm(nh) 1955.1511, Thailand: Ko Butang (HB = 403, T = 269, T/HB x 100 = 66.7). and ear exceeds the growth rate of the head and body and that subsequently— probably beginning in late infancy— the growth rate of the head and body exceeds that of the appendages (cf. Lumer & Schultz, 1941, p. 284; Karrer, 1970, p. 172; Yosh- idaet al., 1993, p. 438). Geographic Variation Head and Body Length— In 184 core-area adults, geographic variation of HB is generally similar in females and males (Table 8, Fig. 10). Surveying this variation from south to north, HB 22 FIELDIANA: ZOOLOGY Table 8. Head and body length in core-area Macaca fascicularis: summary of variation (cf. Fig. 10). Adult females Adult males Sample area N Mean ± SD Extremes N Mean ± SD Extremes Indochinese Peninsula1 16 449.1 ± 50.26 354-545 3 531.0 ± 98.02 434-630 Malay Peninsula2 11 393.6 ± 21.75 360-432 14 445.5 ± 46.62 370-519 Sumatra 5 420.6 ± 27.69 380-458 20 484.9 ± 37.04 420-551 Borneo 43 408.0 ± 27.34 354-469 42 473.1 ± 43.05 400-590 Java 11 435.1 ± 33.30 350-480 19 512.7 ± 46.29 434-610 All 86 418.0 ± 37.22 350-545 98 481.0 ± 49.24 370-630 1 Includes Isthmus of Kra north of 10°N. 2 Includes Isthmus of Kra south of 10°N. in both sexes is relatively great in Java (ca. 7°S); generally declines northward and reaches a min- imum at 0°-5°N in the Malay Peninsula, Sumatra, and Borneo; increases to about 12°N in the Isth- mus of Kra; and decreases farther northward to about 1 6°N, the latitude of the northernmost mea- sured specimens in the Indochinese Peninsula (cf. Aimi et al., 1982, p. 53). At similar latitudes in the Malay Peninsula, Sumatra, and Borneo, HB in each sex is approximately equal. The pattern of geographic variation of HB in core-area M. fas- cicularis parallels that previously reported for skull length in core-area samples of this species (Fooden & Albrecht, 1993, p. 526); for both HB and skull length, size variation in core-area M. fascicularis generally follows Bergmann's rule (cf. Mayr, 1963, p. 3 1 9), except for the aberrant northward decrease of size north of 1 2°N in the Indochinese Peninsula. One adult female collected at 7°01'S in Java ap- pears abnormally small (mcz 12757, Ban targe- bang, HB = 350 mm); a subadult female (usnm 156292, HB = 380 mm) collected at the same locality is larger. Fringing-island HB data are available for 215 adult specimens collected in 59 shallow- water is- lands and 2 1 deep-water islands (Appendix 7). In most of these islands, sample size is too small to determine conclusively whether or not insular HB differs from core-area HB at similar latitudes; however, mean HB in shallow-water fringing-is- land samples generally is less than in correspond- ing core-area samples (Table 9, Fig. 1 1). This pat- tern of HB variation in shallow-water fringing is- lands suggests a tendency toward insular dwarfism, as previously also observed for skull length vari- ation (Fooden & Albrecht, 1993, p. 525). HB is particularly small, compared with HB in core-area reference samples, in Pinang [1] (west of the Malay Peninsula) and Bali (east of Java). Among deep-water fringing-island samples, only those from the Lesser Sunda Islands show a clear tendency toward insular dwarfism (Table 9, Fig. 12, Appendix 7); in this respect, these samples resemble the sample from nearby Bali (see above), the shallow-water member of the Lesser Sunda group. In other deep-water fringing islands, HB in male samples tends to exceed that in core-area reference samples, and in Mindanao (Philippines) HB in both sexes conspicuously exceeds that in core-area reference samples. In one of four adult females collected in P. Simeulue (west of Sumatra; usnm 121513, HB = 325 mm), HB appears to be abnormally small; the skull and recorded body weight of this female are not particularly small (greatest skull length = 102.4 mm, weight = 3.74 kg). Tail Length— Collectors' measurements of tail length (T) are available for 183 core-area adults and 222 fringing-island adults (Table 10, Appen- dix 8). Although T averages less in females than in males, relative T (T/HB x 100) is approxi- mately equal in the two sexes, as indicated above (Table 7). Because geographic variation of T is not always concordant with geographic variation of HB, T variation and relative T variation are con- sidered separately in the following discussion. In core-area specimens of each sex, T tends to be negatively correlated with latitude (Fig. 1 3), in accord with Allen's rule (Mayr, 1963, p. 323). T averages greatest near the equator in the Malay Peninsula, Sumatra, and Borneo, somewhat less in Java (6°-8°S), and least at 15°-17°N in the In- dochinese Peninsula. In the Malay Peninsula, Su- matra, and Borneo, T— like HB (see above)— is approximately equal in each sex at similar lati- tudes. In Java, T is highly variable. T reduction in Indochinese M. fascicularis has been noted pre- viously (Fooden, 1964, p. 363; 1971, p. 29). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 23 650 600 550- 500 ~ 450- E E O) c O X) T3 C CO ca CD X 400- 350 550 500 450- 400- 350 300 Males D oo $ + 01 o| -e-j O x x ..+_OJ< 4 00 o D o I o o &£ x.. O.l « CA > of FI > CA> islands CA1 FI2 islands CA1 FI2 1 . FI sample includes both sexes a. Sexes in FI sample deviate concordantly from those in CA sample b. Sexes in FI sample deviate discordantly from those in CA sample 1) Females, FI > CA; males, CA > FI 2) Females, CA > FI; males, FI > CA 2. FI sample includes only one sex a. Females b. Males Totals 13 10 1 0.5 0.5 1 0.5 0.5 5 2.5 2.5 5 2.5 2.5 7 2 5 2 1 1 33 7 26 8 5 3 59 15 44 21 10 11 1 FI sample mean exceeds CA sample mean. 2 CA sample mean exceeds FI sample mean. p. 1 00) for two core-area specimens identified by these authors as M. fascicularis; these specimens were collected in 1965 and 1967 at Sontra Peak, Vietnam (16°07'N, 108°18'E). The problematic measurements (in millimeters) are the following: adult male, total length = 730, T = 222 (HB = 508, relative T = 43.7%), and immature female, total length = 665, T = 230 (HB = 435, relative T = 52.9%). The skull of the male and the skin and skull of the female are preserved in the col- lection of the usnm (No. 356979, adult male; No. 356968, subadult female). Collectors' field notes, kept in the archives of the Division of Mammals, usnm (photocopies provided by Linda Gordon and David F. Schmidt), confirm the accuracy of measurements reported by Van Peenen et al. (1 97 1) for the male, but these notes indicate that total length of the female is 6 1 5 (HB = 385, relative T = 59.7%), not 665, as re- ported by Van Peenen et al. The collector's no- tation on the skin tag of the female likewise in- dicates that the total length is 615. Although both of these Sontra Peak specimens are identified by Van Peenen et al. (1971, p. 134; cf. 1968, p. 609) as M. fascicularis, the field notes indicate that they were originally identified by the collectors as M. mulatta. Dorsal pelage of the available skin of the subadult female (usnm 356968) matches that of M. fascicularis fmnh 33505 (Vietnam: Ho Chi Minh City) and therefore confirms the identification made by Van Peenen et al. Because the skin of the adult male (usnm 356979) is lacking, its identification cannot be in- dependently verified. This specimen may be either M. fascicularis or M. mulatta; Sontra Peak is near the border between the ranges of these parapatric species (Fooden, 1971, p. 28). At my request, P. F. D. Van Peenen recently reexamined his files, and he reports that he no longer has notes that would resolve this ambiguity (pers. comm., 27 Feb. 1992). Because of the unresolved ambiguity, data for the male (usnm 356979) are excluded from the present analysis of characters of M. fascicularis. In any event, the short tail of the subadult female (usnm 356968, relative T = 59.7%) is in accord with the pattern of tail reduction in M. fascicularis northward in the Indochinese Peninsula (see above). If the adult male (usnm 356979, relative T = 43.7%) had pelage characters of M. fascicu- laris, it would indicate that even more extreme tail reduction has occurred in this species at Sontra Peak. (Note: The tail also is relatively short in two infants and one juvenile— amnh 87266, ansp 15136, 15138— collected at nearby Muang Tha- teng, Laos.) In a core-area adult female (bm(nh) 1939.181) collected in 1 938 in West Malaysia at Bukit Nanas (formerly known as Weld's Hill; 3°09'N, 101°42'E), the tail is unusually short (T = 3 1 5 mm, relative T = 78.8%), considering the latitude of collection (Fig. 13). The tail also appears short in an adult male collected (date unknown) at the same locality (bm(nh) 1939.180, no collector's measurements; estimated relative T = ca. 75%, based on mea- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 25 650 600 550 500- ~ 450 E E B 400 c O 350- X) C 550 n CO ■a co CD X 500-1 . Males ■ o o 0 400 350 300 O o o o a o 0 <■ o o o <* o Females b ' OB ^ ■ o 10°S 10 Latitude |e o. . ■' LA j 0. 15°N ■ Shallow-water fringing-island specimens Core-area specimens Fig. 1 1 . Latitudinal variation of head and body length in adult shallow-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (cf. Table 9 and Appendix 7). surements of dry skin). In another adult male col- lected in 1 9 1 3 at Bukit Nanas, T is normal (bm(nh) 1955.1512, T = 577 mm, relative T = 126.8%). Two additional short-tailed monkeys were ob- served at Bukit Nanas in 1966 (Bernstein, 1966, p. 1559; 1968a, p. 121); these two monkeys, which were associated with a troop of normal-tailed M. fascicularis, were inferred to be hybrids of M. fas- cicularis and M. nemestrina. In fringing islands, T data are available for small 26 FIELDIANA: ZOOLOGY 650 600 550 500 ~ 4504 E E ■g) 400 c o > o 350 X3 c 550 CO "O co CD X 0 0 o 0 0s 0 ^0 0 o o o oo 0 o 0° A. ■ o ol i&o <>: ■- o 0 o O I . -0 ■ o . 9 I A _. 0 500 450 400- 350 300 10°S 10 15°N Latitude ■ Deep-water fringing-island specimens c Core-area specimens Fig. 12. Latitudinal variation of head and body length in adult deep-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (cf. Table 9 and Appendix 7). samples of adults collected in 57 shallow- water lected at similar latitudes (Fig. 15, Table 1 1); this islands and 22 deep-water islands (Appendix 8). parallels the general reduction of HB in shallow- In shallow-water fringing islands, T is generally water fringing islands relative to HB in core-area reduced relative to T in core-area samples col- reference samples (see above). In deep-water fring- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 27 Table 10. 13, 14).' Tail length and relative tail length in core-area Macaca fascicularis: summary of variation (cf. Figs. Tail length Relative tail length2 Sample area N Mean ± SD Extremes N Mean ± SD Extremes Adult females Indochinese Peninsula3 16 434.4 ± 56.53 334-538 16 97.1 ± 10.95 76.1-114.7 Malay Peninsula4 11 481.5 ± 63.44 315-556 11 122.6 ± 16.78 78.8-143.3 Sumatra5 5 519.6 ± 12.18 508-535 5 124.0 ± 8.28 112.4-134.2 Borneo 43 507.3 ± 34.64 455-628 43 124.7 ± 9.78 110.2-144.0 Java 11 468.6 ± 52.37 345-555 11 108.5 ± 16.21 75.5-140.0 All 86 486.2 ± 52.91 315-628 86 117.2 ± 16.01 75.5-144.0 Adult males Indochinese Peninsula3 3 513.3 ± 50.14 456-549 3 97.9 ± 11.28 84.9-105.1 Malay Peninsula4 13 579.3 ± 41.02 515-640 13 132.6 ± 12.30 102.0-149.5 Sumatra5 20 556.6 ± 38.59 490-623 20 115.3 ± 10.78 99.4-137.9 Borneo6 43 576.7 ± 45.93 470-680 42 122.4 ± 11.32 103.4-147.2 Java7 18 556.5 ± 84.86 360-715 18 110.1 ± 21.73 69.2-148.9 All 97 567.2 ± 54.33 360-715 96 119.2 ± 15.71 Both sexes 69.2-149.5 Indochinese Peninsula3 19 97.2 ± 10.68 76.1-114.7 Malay Peninsula4 24 128.0 ± 15.07 78.8-149.5 Sumatra 25 117.0 ± 10.76 99.4-137.9 Borneo 85 123.6 ± 10.57 103.4-147.2 Java 29 109.5 ± 19.52 69.2-148.9 All 182 118.3 ± 15.84 69.2-149.5 1 Cf. J. Suzuki and Varavudhi (1989, pp. 1 1 1-1 13), who reported tail length in artificially provisioned troops of M. fascicularis in Thailand (cf. Aggimarangsee, 1992, p. 104) 2 Relative tail length = tail length/head and body length x 100. 3 Includes Isthmus of Kra north of 10°N. 4 Includes Isthmus of Kra south of 1 0°N. 5Cf. Bakaretal. (1981, p. 12). 6 One male specimen with tail length measurement lacks head and body measurement. 7 Excludes bobtailed specimen mzb 6483, collected at Cikujang; HB = 540, T = 250, T/HB x 100 = 46.3. ing islands, there is no general pattern of tail re- duction (Fig. 16, Table 11). In deep-water P. Si- muelue and P. Nias (both west of Sumatra) and in the deep-water Lesser Sunda Islands, T tends to be less than in core-area reference samples, but in the Nicobar Islands (northwest of Sumatra) and in many of the Philippine Islands, T tends to be greater than in core-area reference samples (Ap- pendix 8). The greatest reduction of T in fringing-island samples, relative to core-area reference samples, is evident in both sexes in P. Karimun and P. Bintan (both shallow-water islands east of Su- matra), P. Tioman (shallow- water, east of the Ma- lay Peninsula), and P. Simeulue and P. Nias (deep- water, west of Sumatra) (Figs. 15, 16; Appendix 8). T reduction may also be large in P. Pintu Ge- dong (west of the Malay Peninsula), P. Bangka (east of Sumatra), Ko Samui (east of the Isthmus of Kra), and P. Belitung (west of Borneo), shallow- water islands known from male samples only. The tail is unusually long in a male specimen collected in P. Acheh (shallow-water, east of the Malay Pen- insula, T = 698 mm; cf. Fig. 15). In P. Pinang [1] (shallow- water, west of the Malay Peninsula), one member of a troop of M. fascicularis had a de- formed tail (Lee Chin Thuan, 1964, p. 172). Relative T in shallow-water fringing-island samples is generally similar to relative T in core- area samples collected at similar latitudes (Fig. 1 7, Appendix 9); this implies that HB and T tend to be similarly reduced in shallow-water fringing-is- land populations, relative to HB and T in corre- sponding core-area populations. Conspicuous ex- ceptions occur in P. Tioman (east of the Malay Peninsula), where T is more reduced than HB, and, conversely, in Ko Khram Yai (south of the In- dochinese Peninsula), where HB is more reduced than T (cf. Figs. 11, 15). Relative T in deep-water fringing-island sam- ples frequently deviates from relative T in core- area samples collected at similar latitudes (Fig. 1 8, 28 FIELDIANA: ZOOLOGY 700 650 600 550 .".oo 450 400 c o 'c0 350 650 600 350 500 450 400- 350 ol □ 0 | D 1 i La Males ° ° I °i $ *s x i + 0 v \> + | J 1 ^ 00 i x X ! $ t+? %> x |o * i+ % ! O x*x |§ S{ + o °! o x t !# n ' + Ol + x X 1+ i ! o X D a a + |o° X 1 1 D 300 10°S Females D 1 ° x x ° i+ + i i 0 00 + X X a v x<0 D 1 r i I j D x x X i X* D 1 ! 1 x ; i ' i ' i ' j X 10 Latitude 15°N O Borneo □ Java x Mainland + Sumatra Fig. 13. Latitudinal variation of tail length in adult core-area specimens of Macaca fascicularis (cf. Table 10). Appendix 9). Relative T in P. Simeulue and P. Nias (west of Sumatra) is significantly less than at corresponding latitudes in West Malaysia, Su- matra, and Borneo, and relative T in Mindoro and Luzon (northern Philippines) is significantly great- er than at corresponding latitudes in the Indo- chinese Peninsula. In P. Simeulue and P. Nias, T is more reduced than HB, relative to HB and T in West Malaysia, Sumatra, and Borneo; in Min- doro and Luzon, both HB and T are increased— the latter more than the former— relative to HB and T in the Indochinese Peninsula, where both FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 29 150 5 Latitude 15"N O Borneo □ Java x Mainland + Sumatra Fig. 14. Latitudinal variation of relative tail length in adult core-area specimens of Macaca fascicularis (cf. Table 10). of these measurements decrease latitudinally (cf. Figs. 10, 13). In the deep-water Lesser Sundas, as in Bali (cf. Fig. 17), HB and T tend to be iso- metrically reduced, relative to HB and T in Java. Cranial Characters Sex and Age Variation (Figs. 19, 20) In wild-collected M. fascicularis, greatest length of skull (GL) in adult males (1 18.7 mm, n = 454) averages approximately 1 8% greater than in adult females (100.4 mm, n = 439) (Table 12). Zygo- matic arches are relatively narrow in both sexes; relative zygomatic breadth (ZB/GL x 100) aver- ages 67.9 in 440 males and 65.9 in 428 females. The rostrum is relatively long and narrow, partic- ularly in males; the rostral-postrostral ratio (R/ PR x 100) averages 56.7 in 316 males and 47.6 in 24 1 females. Supramaxillary ridges and lateral maxillary concavities vary from weakly to strongly defined in both sexes. Canines in males are rela- tively large. A median sagittal crest, formed by progressive convergence of the temporal lines, is prominent in many old males. During development from infancy to adult- hood, zygomatic breadth increases slightly more rapidly than skull length, and rostral length in- creases much more rapidly than postrostral length (Table 12; cf. N. Fujiwara, 1963, p. 57; Morimoto, 1982, p. 98; Nanda et al., 1987, p. 217). Recent studies indicate that the conspicuous sexual di- morphism of R/PR in adult M. fascicularis prob- ably is a result of both a higher rate and a greater duration of allometric growth of the rostrum in males (Ravosa, 1991, p. 408; Richtsmeier et al., 1993a, p. 28); prenatal and postnatal growth pat- terns contribute differentially to development of adult cranial morphology (Richtsmeier et al., 1 993b, p. 322). Fluctuating asymmetry of the skull generally increases with age (Halgrimsson, 1993, p. 431). The sequence and timing of dental emergence 30 FIELDIANA: ZOOLOGY 700 650 600 550- 500- 450 400 350 0 | 0 1 ol 0 | ■ ■ Males 0 o 1 0 \ 0 ■ o| 0 CO | 0 o ^> o _ o o.. 0 ■ ■ M • a%% 9 & -j ■ ■ s ■o <£*> o ■ ■ » ■ §■■ ■ : f'T" t « ■ ■ 0 |o 0 ■ ol 7<>i 1 ■ i 0 ■ 1 Bo | 0 0 0 1 ■ ■ V * ■ ■■ I ■■ ■ ■ ■ !o° ■ ■ 1 0 ■ ■ 1 ■ o en c 9 650-i 600 J 550 500 450- 100 350- 300 Females o lo' 0 o o ?$°l Od> ■ o d> o o <0 0" o % ■ ■ ■ ■ I 10°S 10 15°N Latitude ■ Shallow-water fringing-island specimens o Core-area specimens Fig. 15. Latitudinal variation of tail length in adult shallow-water fringing-island specimens ofMacacafascicularis compared with that in adult core-area specimens (cf. Table 1 1 and Appendix 8). in captive M. fascicularis have been investigated p. 250). Results of these investigations are in gen- by Spiegel (1952, p. 127), Berkson (1968, p. 354), eral agreement (Tables 13, 14; cf. Richtsmeier et Bowen and Koch (1970, p. 118), B. H. Smith et al., 19.93a, p. 4); however, Bowen and Koch (1970, al. (1994, pp. 214, 226), and Ostyn et al. (1995, p. 1 14) cautioned that the timing of dental emer- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 31 700 15°N Latitude ■ Deep-water fringing-island specimens Core-area specimens Fig. 16. Latitudinal variation of tail length in adult deep-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (cf. Table 1 1 and Appendix 8). gence in captivity may not be identical to that in and the last deciduous tooth (m2) apparently natural populations. emerges before age 1 0 months (Table 1 3); no sex- In captivity, the first deciduous tooth (i,) may ual differences have been observed in timing or emerge (i.e., penetrate the gingiva) before birth, sequence of emergence of deciduous teeth (Spiegel, 32 FIELDIANA: ZOOLOGY Table 11. Summary comparison of tail length and relative tail length (T/HB x 100) in fringing-island (FT) and core-area (CA) samples of Macaca fascicularis.1 For details, see Appendixes 8 and 9. Shallow-water FI samples Deep-water FI samples Categories of FI samples Number of islands FI > CA2 CA > FI3 Number of islands FI > CA2 CA > FI3 Tail length 1 . FI sample includes both sexes a. Sexes in FI sample deviate concordantly from those in CA sample b. Sexes in FI sample deviate discordantly from those in CA sample 1) Females, FI > CA; males, CA > FI 2) Females, CA > FI; males, FI > CA 2. FI sample includes only one sex a. Females b. Males Totals All samples (including both sexes) 15 11 2 1 1 0.5 1 0.5 2 0 1 0 1 0 8 31 57 0 6 11.5 8 25 45.5 2 10 22 1 3 8 1 7 14 Relative tail length 57 25 32 21 8 13 1 Samples for tail length differ slightly from those for relative tail length because both head and body length and tail length are not available for all specimens. 2 FI sample mean exceeds CA sample mean. 3 CA sample mean exceeds FI sample mean. 1952, p. 129; 16 females, 22 males, not all teeth studied in all specimens). The first permanent tooth (M,) may emerge as early as age 1 .2 years, and the last permanent tooth (M3) may not emerge until age 9.5 years (Table 14). The permanent canines of males emerge approximately 1 year later (C,, age 4.4 yr; C1, age 4.8 yr) than the much smaller permanent canines of females. Spiegel (1952, p. 128) indicated that the interval between initial emergence of a tooth and complete eruption to its final height is 1-2 months for most teeth, some- what longer for third molars, and several years for male canines. Geographic Variation In core-area specimens examined, skull length varies latitudinally (Table 15, Fig. 21), mostly in accord with Bergmann's rule, as previously re- ported by Fooden and Albrecht (1993, p. 525). In both sexes, skull length is high in Java (ca. 7°S), decreases to a minimum in Sumatra and Borneo (ca. 0°), increases to a second peak in the northern part of the Isthmus of Kra (ca. 1 2°30'N), and de- creases farther northward — contrary to Berg- mann's rule— in the Isthmus of Kra and Indo- chinese Peninsula (northernmost measured spec- imens collected ca. 1 7°N). Within each sex, skull length is approximately equal in specimens col- lected at similar latitudes in the Malay Peninsula, Sumatra, and Borneo; a similar latitudinal ho- mogeneity of skull length in these three disjunct landmasses was previously noted in M. nemestri- na (Fooden, 1975, p. 85). Four core-area skulls stand out as aberrantly large for their latitudes of collection (Fig. 21). Of these, one female skull (bm(nh) 1939.181, GL = 1 19.2 mm) and one male skull (bm(nh) 1939. 180, GL = 1 32.0 mm), both collected in West Malaysia at Bukit Nanas (3°09'N), belong to specimens that were previously cited for their aberrantly short tails (see above, External Measurements and Pro- portions); another adult male (bm(nh) 1955.1512, GL = 1 16.9 mm) collected at Bukit Nanas has a skull and tail of normal length. The other two aberrantly large skulls were collected at widely sep- arated localities in Borneo— a female (fmnh 68700, GL = 115.8 mm) in Sabah at Kretam Besar (5°32'N) and a male (usnm 521837, GL = 131.0 mm) in Kalimantan at Hantakan (2°38'S); the tail is of normal length in both of these specimens. Collectors' measurements of head and body length are available for three of the four specimens with aberrantly large skulls (all except bm(nh) 1 939. 1 80); in these three specimens, HB is large, but not aber- rantly so. In shallow-water fringing islands, geographic FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 33 50 140 I -jo 20- m i t no .c c V 100 ® 90 to DC 80 70- 60 Both sexes o ■ 0 o . Si 0° •■■&■ ■ o. si i * * if ; ° 10°S 0 5 10 Latitude 15°N ■ Shallow-water fringing-island specimens o Core-area specimens Fig. 17. Latitudinal variation of relative tail length in adult shallow-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (cf. Table 1 1 and Appendix 9). variation of skull length also is in general accord with Bergmann's rule, but skull length, like HB (see above), tends to be somewhat smaller in these islands than in core-area populations at similar latitudes (Fig. 22, Table 16); this indicates a gen- eral tendency toward insular dwarfism in shallow- water fringing islands (cf. Fooden & Albrecht, 1 993, p. 525). For four shallow-water fringing islands, samples are sufficiently large to establish that both females and males are significantly smaller than in core-area reference samples; these islands are P. Bintan (Indonesia: Riau Archipelago, 1°09'N), P. Tioman (West Malaysia: east coast, 2°48'N; in- cluding one aberrantly small male, usnm 101744, GL = 97.4 mm), Ko Kut (Thailand: southeast coast, 1 1°40'N), and Bali (Indonesia: Lesser Sunda Islands, 8°15'S). In five shallow-water fringing-is- land samples, skull length may be larger than in core-area reference samples; four of these islands are in the Java Sea, between Borneo and Java (Indonesia: P. Karimunjawa, 5°51'S; P. Kemujan, 5°51'S; P. Bawean, 5°48'S; P. Matasiri, 4°47'S), and one is north of Borneo (Sabah: P. Banggi, 7°09'N). In deep-water fringing islands, a tendency to- ward insular dwarfism is evident only in the Lesser Sunda Islands (8°32'-9°42'S) (Fig. 23, Table 16); skull length variation in deep-water Lesser Sunda Islands is similar to that in Bali, the only shallow- water Lesser Sunda Island (cf. Fig. 22). Skull length in deep-water fringing islands tends to be greater than in core-area reference samples in the Nicobar Islands (6°58'-8°00'N), northwest of Sumatra; in P. Simeulue (2°39'N) and nearby P. Lasia (2°10'N), west of Sumatra; and probably in P. Maratua (2°15'N), east of Borneo (cf. Fooden & Albrecht, 1993, p. 531). In the Philippine Islands (5°24- 1 8°3 1 'N), skull length tends to increase with lati- tude, extending the Bergmannian trajectory evi- dent in core-area populations between the equator and about 12°30'N (Fig. 23; Fooden, 1991, p. 10); variation of skull length in Philippine samples col- lected north of about 12°30'N does not follow the anti-Bergmannian trajectory evident in core-area 34 FIELDIANA: ZOOLOGY 150 140 3 130 O X 120 CO I 1 — t 110- o 100 0) 90 & 80 70- 60 O 0 0 ■<£ |o 10°S Both sexes vo :0o .o°x> 1 0 a^i * o ^t , \ M * lo = '. o- 00 0 5 10 Latitude 15°N ■ Deep-water fringing-island specimens o Core-area specimens Fig. 18. Latitudinal variation of relative tail length in adult deep-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (cf. Table 1 1 and Appendix 9). samples collected at similar latitudes in the Isth- mus of Kra and Indochinese Peninsula. Skull length is aberrantly small in one male specimen collected at Mahayahaya (13°55'N; GL = 109.3 mm), Lu- zon, Philippines. Molecular Biology and Genetics Mitochondrial DNA In a unique and valuable series of studies, Ha- rihara et al. (1986, p. 357; 1988, p. 1 18; 1991, p. 611) have employed five restriction enzymes to investigate restriction fragment length polymor- phism in the mitochondrial DNA (mtDNA) of six geographic samples of M. fascicularis (Table 1 7). One of these samples consists of five captive-born individuals— one born to a female imported from Vietnam and four born to females imported from Cambodia. Two samples consist of a total of 98 individuals livetrapped in Thailand; one of these samples originated north of the Isthmus of Kra (< 7 localities), and the other originated south of the Isthmus of Kra (2 localities). Each of the other three samples consists of 48 individuals, imported, respectively, from Malaysia, Indonesia, and the Philippines. For the last three samples, no further details are available concerning the regions or is- lands of origin of individuals in the samples or concerning possible geographic heterogeneity within the samples (Harihara et al., 1986, p. 357; 1988, p. 125); this is unfortunate, because Malay- sia, Indonesia, and the Philippines each include widely dispersed landmasses inhabited by isolated populations of M. fascicularis. In Table 1 7, the six sample areas of Harihara et al. are arranged in approximate geographic order. They are: (1) Vietnam and Cambodia, both east of (2) Thailand north of the Isthmus of Kra, which obviously is north of (3) Thailand south of the Isthmus of Kra; the latter sample area is north of (4) West Malaysia and northwest of East Malaysia FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 35 Fig. 19. Skull of adult female Macacafascicularis, near topotype— amnh 102764, Indonesia: Sumatra, Muaradua. (Photographs by James L. Balodimas, Division of Photography, fmnh.) (Sarawak and Sabah, both in Borneo); West and East Malaysia, together considered one sample area, are mostly north of (5) Indonesia; and Ma- laysia and Indonesia are southwest of (6) the Phil- ippine Islands. In the absence of information con- cerning whether any members of the Malaysian and/or Indonesian samples originated in Borneo, the geographic relationship of the Philippine sam- ple to the Malaysian and Indonesian samples re- mains ambiguous. For the four initially studied non-Thai samples, treatment of M. fascicularis mtDNA with each of the five restriction enzymes yielded three to six distinctive digestion patterns (enzyme morphs), as follows: BamHl and EcoRl, three morphs each; Hpal and Sstl, four morphs each; and Hi ndlll, six morphs (Table 17). In the 149 individuals in- cluded in these four samples, 21 different com- binations (mtDNA types) of these enzyme morphs were detected. In the 98 individuals included in the two subsequently studied Thai samples, four additional enzyme morphs and three additional 36 FIELDIANA: ZOOLOGY Fig. 20. Skull of adult male Macacafascicularis, near topotype— amnh 106566, Indonesia: Sumatra, Bukit Sanggul. (Photographs by James L. Balodimas, Division of Photography, fmnh.) mtDNA types were detected; at each locality in these two samples, all individuals were uniform in their mtDNA type (Harihara et al., 1991, p. 611). Except for the peculiar mtDNA type in the Thai- land south of the Isthmus of Kra sample (Harihara et al., 1991, p. 612), the distribution of enzyme morphs and mtDNA types in samples generally conforms to the geographic relationships of sam- ples (Table 17; cf. Melnick & Hoelzer, 1993, p. 3). Although the small Indochinese sample shares nei- ther of its two mtDNA types with other samples, it resembles the Malaysian sample in its high fre- quency ofSstl morph 2. Mitochondrial DNA types in the Thailand north of the Isthmus of Kra sample are closely related to those in the Indochinese and Malaysian samples (Harihara et al., 1 99 1 , p. 6 1 2). The Indonesian sample (10 mtDNA types), which FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 37 Table 12. Cranial measurements1 and proportions in age/sex classes of wild-collected Macaca fascicularis.2 Relative zygomatic Postrostral Greatest length breadth length Rostral- -post rostral Age/sex class3 (mm) (ZB/GL x 100) (mm) ratio (R/PR x 100) Fetuses 60.8 56.7 52.0 23.7 U) (1) (/) V) Infants 75.9 ± 6.21 63.1 ± 2.48 64.2 ±4.11 25.7 ± 3.60 56.0-93.7 57.0-68.8 50.6-70.8 18.2-34.1 (80) (78) (57) (57) Juveniles 93.3 ± 9.40 64.6 ± 2.44 71.8 ± 4.08 40.6 ± 6.28 72.2-130.0 57.0-78.1 62.7-85.4 28.3-55.5 (482) (476) (287) (287) Subadult females 97.6 ± 4.54 65.6 ± 2.54 73.3 ± 3.12 44.0 ± 3.23 89.7-109.9 60.2-72.9 67.2-80.2 37.7-51.5 (70) (69) (40) (40) Subadult males 111.5 ± 6.70 66.9 ± 2.72 78.4 ± 3.52 53.0 ± 4.28 95.7-128.0 59.8-75.4 70.2-88.1 42.3-63.4 (123) (122) (92) (92) Adult females 100.4 ± 5.63 65.9 ± 2.40 74.0 ± 3.26 47.6 ± 4.21 84.0-119.2 59.4-74.1 64.3-87.2 36.9-61.0 (439) (428) (241) (241) Adult males 118.7 ± 7.29 67.9 ± 2.66 80.9 ± 4.02 56.7 ± 3.94 97.4-140.1 59.8-76.4 70.6-93.1 41.9-66.3 (454) (440) (316) (316) 1 For definition of measurements, see Fooden (1969, p. 40). 2 Mean ± SD (where n > 2), extremes, and sample size (italicized figures in parentheses). 3 Dental specifications: infants, deciduous teeth only; juveniles, some permanent teeth erupted; subadults, M3 in females or C in males incompletely erupted (cf. Spiegel, 1 952, p. 1 29); adults, all permanent teeth completlely erupted. may be geographically intermediate between the Malaysian and Philippine samples (see above), shares 2 mtDNA types with the Malaysia sample (9 mtDNA types) and shares 2 other mtDNA types with the Philippine sample (4 mtDNA types). The Philippine sample shares no mtDNA types with any sample other than that from Indonesia (cf. Lawler et al., 1995; p. 137). Known genetic dis- Table 13. Schedule of emergence of deciduous teeth in captive Macaca fascicularis3; emergence age in days. Spiegel (1952, p. 131) Berkson (1968, p. 355) Bowen and Koch (1970, p. 118) Tooth Mean Extremes N Median4 Extremes N Mean Extremes N ii 14 0-41 37 30 15-75 10 18 7-35 18 i1 18 5-39 37 30 15-75 10 20 7^*2 18 la 26 0-58 37 30 30-75 10 36 14-49 18 i2 40 18-77 37 45 30-75 10 47 21-63 18 c, 71 28-187 37 75 45-105 10 72 35-98 18 c1 77 28-173 36 75 45-105 10 75 35-91 16 m, 1 m1 J 66 35-98 37 90 90 75-120 75-120 10 10 74 72 56-91 56-98 20 17.5s m2 154 98-252 36 165 150-210 10 176 119-287 18 m2 168 105-280 36 180 150-225 10 189 119-238 18 12 Designate positions of teeth. 3 Cf. B. H. Smith et al. (1994, p. 214), who cite the following mean emergence ages (days) without specifying sample sizes or extreme values: i,, 14; i\ 21; i2, 28; i2, 33; c„ 76; c1, 76; m„ 76; m1, 76; m2, 167; m2, 198. 4 Estimated to nearest half month. 5 Examined unilaterally in one subject. 38 FIELDIANA: ZOOLOGY Table 14. Schedule of emergence of permanent teeth in captive Macaca fascicularis5; emergence age in years. Spiegel (1952, p. 134)6 Bowen and Koch (1970, p. 119) Females Males Both sexes Tooth Mean Extremes N Mean Extremes N Mean Extremes N7 M, M1 I, I1 I2 I2 M2 M2 P3 P3 P4 P4 C, c M3 M3 1.38 1.54 2.38 2.38 2.62 2.85 3.38 3.54 3.62 3.92 3.38 3.85 5.54 6.23 1.23-1.69 1.31-1.92 2.08-3.23 2.31-3.00 2.38-2.92 2.38-3.15 3.00-3.69 3.23-3.92 3.23^.38 3.46-4.62 3.00-3.92 3.23-4.38 5.31-7.69 5.92-9.54 13 13 10 10 10 10 10 8 10 10 10 3 3 1.54 1.69 2.46 2.54 2.62 2.92 3.38 3.46 4.00 4.38 4.77 6.00 6.46 1.38-1.85 1.46-2.08 2.08-2.69 2.31-2.92 2.15-2.08 2.85-3.38 2.92-3.69 3.00-3.85 3.15-5.08 3.85-4.69 4.08-5.15 5.38-6.46 5.69-6.62 10 10 1.34 1.46 2.40 2.41 2.54 2.68 3.68 3.83 3.68 3.60 3.68 3.79 3.52 3.94 1.17-1.50 1.17-1.67 1.92-2.75 1.92-2.83 2.08-2.83 2.17-3.08 3.33-3.83 3.33^1.08 3.33-3.83 3.08-3.83 3.33-3.83 3.25-4.08 3.00-4.08 3.33-4.25 16 15 10 10.5 6.5 6.5 3 1.5 3 3 3 1.5 3 1.5 1-4 Designate positions of teeth. 5 Cf. B. H. Smith et al. (1994, p. 226), who cite the following mean emergence ages (yr) in females without specifying sample sizes or extreme values: M„ 1.50; Ml, 1.75; I„ 2.50; I1, 2.50; I2, 2.50; I2, 2.50; M2, 3.25; M2, 3.25; P3, 3.50; P3, 3.25; P4, 3.50; P4, 3.25; C„ 3.00; C, 3.25; M3, 5.50; M3, 6.00. Also cf. Ostyn et al. (1995, p. 250), who provide the following emergence ages (yr; mean ± SD) for six teeth: females (n = 3)— M,, 1.33 ± 0.09; M1, 1.46 ± 0.09; I,, 2.54 ± 0.16; I1, 2.70 ± 0.08; I2, 2.70 ± 0.08; I2, 2.85 ± 0.19; males (n = 6, except I2, n = 4)-M„ 1.35 ± 0.11; M1, 1.53 ± 0.10; I„ 2.48 ± 0.21; I1, 2.50 ± 0.23; I2, 2.57 ± 0.24; I2, 2.70 ± 0.30. 6 In the present table, Spiegel's data are converted from "Messjahres" to calendar years [calendar years = (Messjahre x 48/52) + (Messmonate x 4/52)]; cf. Spiegel (1952, p. 127, footnote 2). 7 Decimal fractions indicate unilateral examination of subjects. tances between sample pairs also generally con- form to geographic relationships (Table 17, foot- note 1). Further interpretation of the taxonomic and phylogenetic significance of these mtDNA data is hindered by the imprecise provenance and pos- sible geographic heterogeneity of the samples from Malaysia, Indonesia, and the Philippines. Nuclear DNA Multiple Alpha-Globin Genes— Restriction- endonuclease analysis of M. fascicularis DNA (n = 168) indicates that individuals of this species have from one to four alpha-globin genes per hap- loid genome and that the frequency of alpha-glo- Table 15. Greatest length of skull in core-area Macaca fascicularis: summary of variation1 (see Fig. 21) Adult females Adult males Sample area N Mean ± SD Extremes N Mean ± SD Extremes Indochinese Peninsula2 22 105.0 ± 6.50 95.7-116.2 10 123.9 ± 5.12 116.4-132.3 Malay Peninsula3 13 101.9 ± 7.33 92.6-119.2 20 115.0 ± 7.05 106.6-132.0 Sumatra 39 98.4 ± 3.42 93.1-103.9 80 116.2 ± 4.37 107.4-128.2 Borneo 71 99.1 ± 4.40 89.7-115.8 69 115.6 ± 5.36 103.2-133.5 Java 40 104.3 ± 4.26 97.0-114.0 44 126.3 ± 5.76 115.9-138.1 All 185 101.0 ± 5.40 89.7-119.2 223 118.2 ± 6.81 103.2-138.1 1 Includes specimens without precise localities that are excluded in Fooden and Albrecht (1993, p. 537). 2 Includes Isthmus of Kra north of 10°N. 3 Includes Isthmus of Kra south of 10°N. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 39 145 ~ 140 E E 135 X 130 e> Z 125 111 -J 120 - _l ^ 115 (A LU 110 " _l < S 105 h 100 _ 120 - E E 115 - I H O 2 LU D CORE AREA no h 105 100 h 95 UJ 90 - _J < 85 - 80 B = BORNEO M = MAINLAND J = JAVA S = SUMATRA M — M~~~ M ^ \ ""M "^"-^^ M FEMALES 10°S 8°S 6°S 4°S 2°S 0° 2°N 4°N 6°N 8°N 10°N 12°N 14°N 16°N 18°N 20°N LATITUDE Fig. 2 1 . Latitudinal variation of greatest length of skull in adult core-area specimens of Macaca fascicularis (Fooden & Albrecht, 1993, p. 526). Solid lines indicate third-order polynomial regressions of skull length on latitude for females and males; dashed lines indicate 95% confidence limits. bin gene haplotypes varies geographically (Table 1 8 and below, Blood Proteins; Barnicot et al., 1966, p. 241; 1970, p. 380; D. G. Smith & Ferrell, 1980, p. 558). Judging from available evidence, the sin- gle-gene haplotype is rare, having been detected (heterozygously) in only 1 of 1 4 west-central Thai individuals and 1 of 20 Indonesian individuals. The double-gene haplotype is most common; it is very frequent (0.88-1.00) in Thailand north of the Isthmus of Kra; less frequent (0.52-0.62) in Thai- land south of the Isthmus of Kra, in Malaysia, and in Indonesia; and, somewhat incongruously, it is very frequent (0.98) in the Philippines. The triple- gene haplotype, conversely, is relatively rare (0- 0.12) in Thailand north of the Isthmus of Kra; moderately frequent (0.28-0.43) in Thailand south of the Isthmus of Kra, Malaysia, and Indonesia; and absent in the Philippines. The quadruple-gene 40 FIELDIANA: ZOOLOGY I F CD z LU D I F CD D XL CO lu 90 - _l < ^ 85 UJ LL 80 45 40 35 30 - 25 - 20 " 15 - 10 - 05 - 00 20 h 15 10 - 05 - 00 95 h SHALLOW-WATER FRINGING ISLANDS MALES REGRESSIONS AND CONFIDENCE LIMITS TAKEN FROM CORE-AREA ANALYSES FEMALES J I I L 10°S 8°S 6°S 4°S 2°S 0°20N40N60N8°N 10°N 12°N 14°N 16°N 18°N 20°N LATITUDE Fig. 22. Latitudinal variation of greatest length of skull in adult shallow-water fringing-island specimens of Macaca fascicularis compared with that in adult core-area specimens (Fooden & Albrecht, 1993, p. 528). For key to shallow- water fringing-island symbols, see Fooden and Albrecht (1993, p. 537). Solid lines indicate third-order polynomial regressions for core-area specimens; dashed lines indicate 95% confidence limits (see Fig. 21). haplotype is relatively rare (0.02-0.08) and is re- stricted to Malaysia, Indonesia, and the Philip- pines. In M. mulatta and M. fuscata, two other members of the fascicularis species group (Food- en, 1991, p. 2), the frequency of alpha-globin gene haplotypes apparently is similar to that in Thai- land north of the Isthmus of Kra. Highly Repeated Restriction Patterns— A study by Crovella et al. (1994, p. 66) of highly repeated nuclear DNA sequences has revealed that, for nine enzymes employed, restriction patterns are indistinguishable in Sumatran, Javan, and Philippine M. fascicularis. Eight of the nine en- zymes— Alul, BamHI, EcoRl, Hindlll, Hinfl, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 41 Table 16. Summary comparison of greatest length of skull in fringing-island (FI) and core-area (CA) samples of Macaca fascicularis.1 For details, see Fooden and Albrecht (1993, Tables 2, 3). Shallow-water FI samples Deep-water FI samples Categories of FI samples Number Number of FI > CA > of FI > CA > islands CA' FP islands CA" FF 1 . FI sample includes both sexes a. Sexes in FI sample deviate concordantly from those in CA sample b. Sexes in FI sample deviate discordantly from those in CA sample 1) Females, FI > CA; males, CA > FI 2) Females, CA > FI; males, FI > CA 2. FI sample includes only one sex a. Females b. Males Totals 17 12 12 3 1.5 1.5 1 0.5 0.5 1 0.5 0.5 2 1.0 1.0 9 4 5 3 1 2 33 12 21 11 7 4 63 23 40 29 16.5 12.5 FI sample mean exceeds CA sample mean. CA sample mean exceeds FI sample mean. Hpal, Pstl, and Pvull— also failed to distinguish the three geographic samples of M. fascicularis from Indian and Chinese samples of M. mulatta. One enzyme— Xmn\— distinguished the M. fas- cicularis samples from the M. mulatta samples. Blood Proteins The most important body of data on blood- protein allele frequencies in natural populations of M. fascicularis is the product of a long-term research project that has been pursued since 1979 by Dr. Y. Kawamoto and associates (Kawamoto, 1982, p. 68; Kawamoto & Ischak, 1981, p. 240; Kawamoto & Suryobroto, 1985, p. 36; Kawamoto et al., 1981, p. 20; 1982b, p. 275; 1984, p. 135; 1987, p. 98; 1988, p. 172; 1989, p. 97; 1991, p. 600; Kondo et al., 1991, p. 20; 1993, p. 171; for references to other studies of blood-protein allele frequencies in this species, see Fooden & Lanyon, 1 989, p. 235). Dr. Kawamoto has generously made available an interim summary of his group's cur- rent data on M. fascicularis, partitioned into 1 1 regional or insular samples (Table 19, Appendix 1 0). Of the 2 1 polymorphic loci studied by this group, 14 exhibit relatively limited variation, with the same major allele common to all 1 1 samples; the 7 more variable loci are Alb, CA-II, Dia, HbA- II, IDH, Pi, and Tf (for key to abbreviations, see Table 19, footnote 1). As noted previously (cf. Fooden & Lanyon, 1989, p. 223; Kawamoto et al., 1991, p. 600), blood-protein polymorphism tends to be reduced in insular populations. Two dendrograms (Fig. 24a) of blood-protein relationships among these 1 1 samples have been published by Dr. Kawamoto's group (Kawamoto et al., 1991, p. 600; Kondo et al., 1993, p. 177); one of these dendrograms includes a sample of the introduced Mauritius population of M. fascicu- laris. A consensus tree (Fig. 24b) of stable clusters common to both dendrograms delineates group- ings that frequently are discordant with current and Late Pleistocene (ca. 1 8 Ka) land connections of the regions or islands where the respective sam- ples were obtained (cf. Fig. 3). The most conspic- uous of these discrepancies is the grouping of sam- ples from Thailand north of the Isthmus of Kra with samples from Bali (a shallow-water island that was connected to Java during the Late Pleis- tocene glacial maximum) and Lombok (a deep- water island that was connected to Sumbawa dur- ing the Late Pleistocene glacial maximum). Allele frequencies at two loci (HbA-II and Pi) tend to link the samples from Thailand north of the Isthmus of Kra with those from Bali and Lom- bok, and allele frequencies at one locus (Tf) tend to link the Bali and Lombok samples with each other but not with samples from Thailand north of the Isthmus of Kra (Table 1 9). None of the relevant HbA-II, Pi, or Tf alleles are exclusive to samples from Thailand north of the Isthmus of Kra, Bali, or Lombok. Similar allele frequencies 42 FIELDIANA: ZOOLOGY 145 _ 140 E E 135 ** X 130 i- n z 125 UJ —i _i 120 _i 73 * 115 CO UJ -1 110 < 5 105 100 DEEP-WATER FRINGING ISLANDS MALES REGRESSIONS AND CONFIDENCE LIMITS TAKEN FROM CORE-AREA ANALYSES 120 E \ b E 115 \ \ X __ \ \ c - ^Z Q L^ L X 110 \ \ — " x Bu O- M L H O z UJ _i _i _i 105 100 \ \ ^- \ \ ~ «" T X Bv x *kjw&-i — ** ^^^ \ 95 i ! \ E ^X" ^ \ V> « v^ ^"X \ UJ 90 ^ •*' \ _l *~ — — ■ < 2 UJ 85 FEMALES LL 80 ■ i i i i i i i i i i i i i i 10°S 8°S 6°S 4°S 2°S 0° 2°N 4°N 6°N 8°N 10°N 12 nit nit.fi |v iia Cambodia (98 > n of Kra Malaysia nesia pines no. Ba Ec Hi Hp Ss (n - 53) > 65") (n < 334) (n = 48) (n = 48) (n = 48) 20 1 1 1 2 2 80.0 0 0 0 0 0 21 1 1 4 2 2 20.0 0 0 0 0 0 225 235 7 ? ? ? 7 ? 7 7 ? 7 o } 100.0 0 0 0 0 0 0 0 0 246 ? ? 7 ? ? 0 0 100.0 0 0 0 6 1 1 3 1 3 0 0 0 10.4 0 0 7 1 2 1 1 2 0 0 0 10.4 0 0 15 1 1 1 1 4 0 0 0 2.1 0 0 16 3 1 1 1 2 0 0 0 2.1 0 0 17 1 1 3 1 2 0 0 0 2.1 0 0 18 1 1 5 1 2 0 0 0 2.1 0 0 19 1 1 1 3 2 0 0 0 2.1 0 0 1 1 1 1 1 2 0 0 0 60.4 6.3 0 3 1 1 1 1 3 0 0 0 8.3 37.5 0 5 1 2 2 2 1 0 0 0 0 29.2 0 8 1 2 2 1 1 0 0 0 0 8.3 0 9 1 3 1 2 1 0 0 0 0 8.3 0 10 1 1 4 1 3 0 0 0 0 2.1 0 11 1 1 1 4 3 0 0 0 0 2.1 0 12 1 3 6 2 1 0 0 0 0 2.1 0 2 1 1 1 1 1 0 0 0 0 2.1 52.1 4 1 2 1 1 1 0 0 0 0 2.1 43.8 13 1 2 1 2 1 0 0 0 0 0 2.1 14 2 2 1 1 1 0 0 0 0 0 2.1 1 Genetic distances between non-Thai sample pairs: Indochina-Malaysia, 0.00679 ± 0.001 15; Indochina-Indonesia, 0.01284 ± 0.00157; Indochina-Philippines, 0.02073 ± 0.001 10; Malaysia-Indonesia, 0.00848 ± 0.00184; Malaysia- Philippines, 0.01110 ± 0.00068; Indonesia-Philippines, 0.00270 ± 0.00058 (Harihara et al., 1988, p. 124). 2 Enzyme abbreviations: Ba = BamHl; Ec = EcoRl; Hi = Hindlll; Hp = Hpal; Ss = Sstl. 3 Vietnam, n = 1; Cambodia, n = 4. 4 Cf. Kawamoto et al. (1989, p. 95) and Harihara et al. (1991, p. 611). s Similar to mtDNA type nos. 1-21, particularly those in continental Southeast Asian samples (Harihara et al., 1991, p. 612). 6 Widely distinct from mtDNA type nos. 1-23 (Harihara et al., 1991, p. 612). Asia, Sumatra, Borneo, or Java. The Mauritius population, which apparently was introduced dur- ing the sixteenth century, probably originated from monkeys imported from Java. Computed genetic distances indicate remarkable divergence — ca. 100 times greater than expected— between allele fre- quencies in the Mauritius population and those in Asian populations of M. fascicularis; Kondo et al. (1993, p. 1 79) suggested that this may be the result of a genetic bottleneck (cf. Lawler et al., 1995, p. 138). Allele frequencies at the vitamin D binding pro- tein (DBP) locus have been studied by Tanaka et al. (1989, p. 104; 1991, p. 126) in 251 livetrapped Thai M. fascicularis and in 529 imported or cap- tive-bred Malaysian, Indonesian, and Philippine M. fascicularis (Table 20). Based on frequencies of the three most common alleles at this locus, the sample areas may be subdivided into three groups: (1) Thailand north of the Isthmus of Kra— DBP1 very high (0.86-1.00), DBP2 very low (0-0.01), DBP3 very low (0-0.01); (2) Thailand south of the Isthmus of Kra, Malaysia, and Indonesia— DBP1 moderately high (0.48-0.57), DBP2 low (0-0.19), DBP3 moderately high (0.30-0.40); and (3) Phil- ippines—DBP1 moderately high (0.40), DBP2 moderately high (0.59), DBP3 absent. Allele frequencies at the complement C6 locus have been studied by Omoto et al. (1991, p. 603) in 150 imported or captive-bred Malaysian, In- donesian, and Philippine M. fascicularis (Table 21). The Malaysian and Indonesian samples are 44 FIELDIANA: ZOOLOGY Table 18. Frequency of alpha-globin gene haplotypes in geographic samples of Macaca fascicularis compared with frequency in samples of M. mulatta and M.fuscata (O. Takenaka et al., 1989, p. 87; A. Takenaka et al., 1991, p. 324; A. Takenaka & Takenaka, 1991, p. 632). Region or island N Alpha-globin gene haplotypes Country Single Double Triple Quadruple M. fascicularis Thailand North of Isthmus of Kra East-central and southeast 30 0 1.00 0 0 West-central 14 0.04 0.96 0 0 Southwest 36 0 0.88 0.12 0 South of Isthmus of Kra 22 0 0.61 0.39 0 Malaysia Unknown 22 0 0.52 0.43 0.05 Indonesia Unknown 20 0.02 0.62 0.28 0.08 Philippines Unknown 24 M. mulatta 0 0.98 0 0.02 India Unknown 13 0 0.96 0.04 0 China Unknown 14 M . fuscata 0 0.93 0.07 0 Japan Unknown 30 0.02 0.96 0.02 0 similar in allele frequencies and differ from the Philippine sample (0.025 > P > 0.01), particularly in their lower frequency of alleles A and M4. The Philippine sample is notably less polymorphic than the Malaysian and Indonesian samples. In a recent study, Scheffrahn et al. ( 1 994, p. 135) investigated variation of blood-protein allele fre- quencies in M. fascicularis in Sumatra (nine groups representing four local populations) and four near- by islands (one shallow-water, three deep-water). Based on examination of five to seven polymor- phic loci, this study finds that (1) differentiation among six adjacent social groups in Sumatra is relatively great, (2) differentiation between these six adjacent groups and three other local popula- tions in Sumatra is roughly proportional to geo- graphic distance (25-350 km), (3) allele frequen- cies in a shallow-water fringing-island population in P. Tuangku (area ca. 300 km2) are similar to those in the four local populations in Sumatra, (4) populations in the deep-water fringing islands P. Simeulue (ca. 2,000 km2) and P. Nias (ca. 4,000 km2), both west of Sumatra, cluster together, and (5), differentiation is greatest in P. We (ca. 250 km2), a small deep-water fringing island north of Sumatra. Genetic drift is inferred to be the most important factor in blood-protein differentiation in insular populations. Although blood-protein al- lele frequencies in these M. fascicularis popula- tions in Sumatra and nearby islands are variably differentiated, all of these populations share one or more alleles at each of the five to seven loci. Blood Groups Limited information is available concerning geographic variation of human-type blood group frequencies in M. fascicularis (Table 22). In a study of the A-B-O system in M. fascicularis saliva and serum, type B was much less frequent in Malaysian (26.1%) and Indonesian (15.8%) samples than in a Philippine sample (76.9%); in an earlier study limited to A-B-O agglutinins in serum, type B ap- parently was similarly less frequent in Thai and Malaysian samples than in a Philippine sample. In M. mulatta, a species closely related to M. fas- cicularis and parapatric with M. fascicularis in Thailand (Fooden, 1980, p. 5), the frequency of type B is 97.0% (n = 200; Socha & Ruffie, 1983, p. 47; cf. Nakajima et al., 1970, p. 246); contrary to expectation, this is closer to the frequency in Philippine M. fascicularis than to that in Thai, Malaysian, and Indonesian M. fascicularis. Lewis group frequencies in M. fascicularis are similar in Thai, Malaysian, and Philippine samples. In a preliminary report on geographic variation of simian-type blood group H0* in M. fascicularis, Honjo et al. (1984, pp. 73-74; cf. Terao, 1985, p. 5 1) indicated that the gene frequency of H1 is mod- erately high in a Malaysian sample (n = 295), lower in an Indonesian sample (n = 284), and zero in a Philippine sample (n = 201). In simian-type blood group T^, the gene frequency of T2 is represented (graphically) as lower in the Malaysian sample than in the Indonesian and Philippine samples. Nu- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 45 Table 19. Blood proteins: summary of frequencies (%) of major alleles at polymorphic loci in 1 1 geographic samples of Macaca fascicularis (sample size indicated by italicized figures in parentheses). For details, see Appendix 10. Sampl e areas Thailand North of Isthmus of kra South Central and of south- South- Isthmus West Lom- Sum- Philip- Loci1 i and east west of Kra Malaysia Sumatra Java Bali bok bawa Timor pines2 alleles (727) (124) (33) (140) (276) (222) (136) (35) (81) (7) (142) Plasma proteins Alb A 84 20 52 94 90 100 100 100 100 93 100 B 16 80 48 6 8 0 0 0 0 7 0 Alp A 100 > 99 100 100 100 100 99 100 100 100 97 ChEs 1 100 100 100 100 100 99 100 100 100 100 100 Pi B 18 1 95 54 82 69 22 0 79 64 92 C 82 88 5 46 18 31 78 100 21 36 8 TBPA F 89 98 100 85 94 97 100 100 52 100 94 Tf C3 4 24 0 4 2 10 0 0 60 57 0 Dl 2 8 62 74 62 58 < 1 3 40 43 99 D2 32 13 0 2 0 1 0 0 0 0 0 G2 < 1 1 5 < 1 2 14 98 83 0 0 0 Erythrocyte proteins Acp A 97 99 97 96 > 99 100 100 100 100 100 100 ADA 2 100 92 100 100 100 100 100 100 100 100 99 AK 1 96 100 97 98 100 100 100 100 100 100 92 CA-I a 99 95 100 96 98 99 100 100 100 100 100 CA-II a 61 54 18 39 46 16 0 0 11 0 61 b 39 46 82 61 54 84 100 100 89 100 39 CellEs 1 100 100 100 99 > 99 97 100 100 80 86 95 Dia A 25 48 7 8 1 < 1 0 0 0 0 0 C 72 44 93 91 99 > 99 100 100 100 100 100 EsD 1 100 100 100 > 99 100 100 100 100 100 100 94 HbA-I 1 100 100 95 99 > 99 100 100 100 100 100 100 HbA-II 0 99 100 33 38 13 69 99 97 0 0 0 2 1 0 67 62 87 31 1 3 100 100 100 HbB 1 100 100 100 100 100 100 97 100 100 100 100 IDH 1 47 39 51 80 90 65 15 57 100 86 99 2 53 61 49 19 9 35 85 43 0 14 1 LDHA 1 94 97 100 100 98 100 100 100 100 100 100 PGD A 98 93 97 92 90 99 100 100 100 100 100 PHI 1 100 83 100 99 98 100 98 97 100 100 81 1 Abbreviations: Acp = acid phosphatase, ADA = adenosine deaminase, AK = adenylate kinase, Alb = albumin, Alp = alkaline phosphatase, CA = carbonic anhydrase, CellEs = esterase, ChEs = cholinesterase, Dia = NADH- diaphorase, EsD = esterase D, HbA = hemoglobin alpha-chain, HbB = hemoglobin beta-chain, IDH = isocitrate dehydrogenase, LDHA = lactate dehydrogenase-A, PGD = 6-phosphogluconate dehydrogenase, PHI = phosphohex- ose isomerase, Pi = protease inhibitor, TBPA = thyroxin-binding prealbumin, Tf = transferrin. 2 Island or islands unknown. merical values of these gene frequencies are not reported in this publication. Serum Cholesterol Response The effect on serum cholesterol level of two cho- lesterol-containing diets— one screening diet and one test diet— was studied in samples of adult male "Malayan" (?West Malaysian) and Philippine M. fascicularis (Taub & Bond, 1982, p. 339, abstract only, sample sizes not specified). In response to both diets, serum cholesterol level was signifi- cantly lower in the Malayan sample (screening diet, 235 mg/dl; test diet, 370 mg/dl) than in the Phil- ippine sample (screening diet, 353 mg/dl; test diet, 46 FIELDIANA: ZOOLOGY ■c 3 a. Dendrogram excluding Mauritius sample compared with dendrogram including Mauritius sample. Thailand, Cent. & SE1 Thailand, SW1 Bali Lombok Timor Sumbawa Sumatra W Malaysia Philippines Thailand, S2 Java Mauritius c D- rC 4: Consensus dendrogram. Thailand, Cent. & SE1 Thailand, SW1 Bali Lombok Thailand, S2 W Malaysia Sumatra — Philippines •• Java Sumbawa Timor -:.'-'.' -€ North of Isthmus of Kra. 2South of Isthmus of Kra. Fig. 24. Blood-protein dendrograms for geographic samples of Macaca fascicular is: a. Dendrogram that ex- cludes Mauritius sample (Kawamoto et al., 199 1 , p. 600) compared with dendrogram that includes Mauritius sample (Kondo et al., 1993, p. 177). b. Consensus den- drogram. 487 mg/dl). In response to the screening diet, high- density lipoprotein levels were significantly higher in the Malayan sample (64 mg/dl) than in the Phil- ippine sample (5 1 mg/dl), and serum lipid levels also were consistently and significantly different; in response to the test diet, serum lipid levels were less consistently different. Karyology Hirai et al. (1991, p. 619) studied the chromo- somal G-banding pattern in 297 imported or cap- tive-born M. fascicularis individuals that origi- nated in Vietnam, Cambodia, Malaysia, Indone- sia, and the Philippines; the sample size for each country is not specified. No significant differences in banding pattern were detected among samples from these five countries, although one female from Vietnam exhibited possible interstitial hetero- chromatin in one autosome. The chromosome number of M. fascicularis was confirmed to be 42. Diseases Malaria Of the seven species of malaria {Plasmodium) that have been recorded as natural parasites in macaques, five species naturally infect hi. fasci- cularis (Fooden, 1994, p. 576). Of these, P. inui is quartan (72-hr asexual erythrocytic cycle); P. cy- nomolgi, P. fieldi, and P. coatneyi are tertian (48- hr cycle); and P. knowlesi is quotidian (24-hr cy- cle). Natural infections with all five of these species have been recorded in West Malaysia, which ap- parently is the center of malaria infection in M. fascicularis (Table 23). For P. fieldi, natural infec- tions are known only in West Malaysia. For P. coatneyi, infections also have been reported in the southern Philippine islands of Palawan (question- able) and Cebu. For P. knowlesi, infections have Table 20. Summary of allele frequencies at the DBP locus in geographic samples of Macaca fascicularis (Tanaka etal., 1989, p. 106; 1991, p. 131). Region or island N Alleles Country DBP1 DBP2 DBP3 DBP* DBP16 Other Thailand North of Isthmus of Kra East-central and southeast 46 1.000 0 0 0 0 0 West-central 57 0.860 0 0 0 0.140 0 Southwest 118 0.970 0.013 0.013 0 0.004 0 South of Isthmus of Kra 30 0.567 0 0.400 0.033 0 0 Malaysia Unknown 204 0.476 0.130 0.380 0.007 0 0.007 Indonesia Unknown 167 0.491 0.192 0.305 0.003 0 0.009 Philippines Unknown 158 0.405 0.592 0 0 0 0.003 FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 47 Table 21. Allele frequencies at the complement C6 locus in three imprecisely localized geographic samples of Macaca fascicularis (Omoto et al., 1991, p. 604). Alleles Hetero- Country N A Al A2 B Bl M2 M3 M4 Other1 zygosity Malaysia 50 0.48 0.03 0.06 0.06 0.06 0.08 0.14 0.03 0.06 0.7298 Indonesia 50 0.43 0.05 0.01 0.08 0.02 0.08 0.18 0.05 0.10 0.7622 Philippines 50 0.66 0 0 0 0 0.10 0.09 0.14 0.01 0.5266 Total 150 0.52 0.03 0.02 0.05 0.03 0.09 0.14 0.07 0.05 - Seven minor alleles. been reported in Tioman, Java (questionable), Pa- lawan, and Cebu. For P. cynomolgi, infections have been reported from Cambodia in the north to Java in the south and from the Nicobar Islands in the west to Cebu in the east. Similarly, for P. inui, infections have been reported from Thailand in the north to Java in the south and from the Nic- obar Islands in the west to Mindanao in the east. Negative screening results suggest that malaria in- fections may be absent in M. fascicularis popu- lations that inhabit Luzon (n = 24) and Bali (n = 60). In malarious areas, the mean incidence of natural malaria infections in M. fascicularis is about 0.18 (n > 2,300); individual monkeys often are simultaneously infected with two or more species of malaria. Natural malaria infections in M. fas- cicularis and other macaques are relatively benign. The known vectors of malaria infections in M. fascicularis are five species of mosquitoes that be- long to the Leucosphyrus Group of the genus Anopheles (Table 24); geographic ranges of these five species of mosquitoes are partly overlapping (cf. Fooden, 1994, p. 581). One of these species, A. hackeri, is a proved vector of all five species of malaria that infect M. fascicularis. Individual mosquitoes sometimes harbor two or more species of malaria simultaneously. Although natural malaria infections in M. fas- cicularis are benign, untreated experimental infec- tions with one species of malaria, P. knowlesi, are Table 22. Geographic variation of A-B-O and Lewis blood group phenotype frequencies in Macaca fascicularis. Country Region or island Sample size and frequency (%) A-B-O groups (saliva, serum)1 N B AB Malaysia Unknown Indonesia Unknown Philippines Unknown A-B-O agglutinins (serum)2 N Anti-A Anti-B Anti-A + B Lewis groups (erythrocytes)2-3 N Le"- positive O 245 45.4 26.1 26.9 1.6 165 55.1 15.8 27.9 1.2 130 2.3 76.9 19.2 1.6 None Thailand Southwest (north of Isthmus of Kra) 146 26.7 26.0 35.0 12.3 Malaysia West Malaysia 232 24.6 32.3 30.2 12.9 Philippines Mindanao 60 88.3 5.0 5.0 1.7 N Leb- positive Thailand Southwest (north of Isthmus of Kra) Malaysia West Malaysia Philippines Mindanao 149 4.0 86 4.6 231 4.8 181 10.5 60 3.3 60 8.3 1 Reference: Terao et al., 1981, p. 76 (cf. Honjo et al., 1984, p. 73). 2 References: Nakajima et al., 1970, pp. 246, 248; Omoto et al., 1970, p. 217. 3 Tested by direct agglutination with human anti-Lea serum and rabbit anti-Leb serum. 48 FIELDIANA: ZOOLOGY Table 23. Geographic distribution of malaria species identified as natural parasites in Macacafascicularis samples. For details, see Fooden (1994, p. 576). Geographic origin of M. fascicularis sample Malaria species (Plasmodium) cynomolgi fieldi coatneyi knowlesi sp. Thailand X Cambodia X West Malaysia X X Tioman X Nicobar Islands X X Sumatra Java X X Borneo ? Palawan X X Cebu X X Mindanao X almost invariably fatal in samples of M. fascicu- laris from the Indochinese Peninsula and possibly from the northern part of the Isthmus of Kra, both outside of the natural range of P. knowlesi (Food- en, 1994, p. 585); in continental Southeast Asia, P. knowlesi is not known to occur north of West Malaysia. Conversely, untreated experimental in- fections with P. knowlesi are relatively benign in samples of M. fascicularis from Cebu, which is within the natural range of this species of malaria. This suggests that populations of M. fascicularis that inhabit areas within the natural range of P. knowlesi have evolved partial resistance to the del- eterious effects of this parasite. Simian T-Lymphotrophic Retrovirus, Type 1 Samples of M. fascicularis populations in sev- eral areas have been tested for antibodies to reveal infections with simian T-lymphotrophic retrovi- rus, type 1 (STLV-1) (Table 25). Infections with STLV- 1 are relatively frequent in M. fascicularis in Indonesia, particularly in P. Sumbawa (Lesser Sunda Islands, between P. Lombok and P. Flores), and rare or absent in other natural populations sampled. STLV-1 infections also are fairly fre- quent in the introduced population of M. fasci- cularis in Mauritius. Susceptibility to Attenuated Polio Virus A retrospective analysis of the frequency of spi- nal cord lesions induced by inoculation with var- ious dilutions of two types of polio vaccines in- dicates that M. fascicularis captives imported from the Philippines were up to 1 ,000 times more sus- ceptible to induction of lesions than M. fascicularis captives imported from Indonesia, Malaysia, and the Indochinese Peninsula (Chino et al., 1992, p. 11). Natural History Habitats Although M. fascicularis is most commonly en- countered at low elevations, its known altitudinal range in the core area of its distribution extends to at least 1 200 m in West Malaysia (Gunong Be- nom; possibly to 1500 m in Cameron Highlands), to 2000 m in Sumatra (Bur ni Bebuli), to 1 800 m in Borneo (Lumu Lumu), and to 2000 m in Java (Gunung Salak); in the northern part of the core area (altitudinal data available for Bangladesh, Vi- etnam, Cambodia, and Thailand), this species has not been recorded above 400 m (Lac Giao, Viet- Table 24. Known vectors of malaria species that have been identified as natural parasites in Macaca fas- cicularis. For details, see Fooden (1994, p. 581). Malaria species (Plasmodium) Vector species cyno- coat- know- (Anopheles) inui molgi fieldi neyi lesi balabacensis ? ? ? ? dims, sensu lato x x hacked x x x x x introlatus x x leucosphyrus, sensu lato x FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 49 Table 25. Simian T-lymphotrophic retrovirus, type 1: frequency of Macaca fascicularis individuals or groups positive for antibodies. Data for individuals Data for groups Individuals Groups Refer- Origin N positive % N positive % ence Blood samples obtained from monkeys in country of origin Thailand, north of Isthmus ofKra 220 0 0 7 0 0 1 Thailand, south of Isthmus ofKra 30 0 0 2 0 0 1 Thailand 117 4 3.4 — — — 2 West Malaysia 199 0 0 — — — 2 Indonesia 245 34 13.9 23 8 34.8 3 Sumatra — — — 4 1 25.0 Java — — — 7 2 28.6 Bali — — 0 4 0 0 Lombok — — 0 3 0 0 Sumbawa — — ca. 50 5 5 100.0 Blood samples obtained from imported laboratory monkeys Malaysia 113 4 3.5 — — — 4 Indonesia 103 20 19.4 — — — 4 Philippines 83 0 0 - - - 4 Blood samples obtained from introduced population Mauritius 102 15 14.7 6 - - 5 1 Key to references: /, Ishida & Varavudhi, 1992, p. 164. 2, Ishida et al., 1985, pp. 840-842. 3, Hayami et al. 1983, p. 620. 4, Hayami et al., 1984, p. 181.5, Matsubayashi et al., 1992, p. 284. nam). In shallow-water fringing islands, the alti- tudinal range extends to 900 m in P. Tioman (Se- dagong) and 1 200 m in Bali (Gunung Bratan). In deep-water fringing islands, where M. fascicularis often is the only native monkey, the altitudinal range extends to 1 700 m in Lombok (Gunung Rin- jani), 2100 m in Timor (Gunung Mutis), 1950 m in Mindanao (Mt. McKinley), 1800 m in Negros (Canlaon Volcano), 1 900 m in Mindoro (Mt. Hal- con), and 2300 m in Luzon (Mt. Data). At low elevations, the preferred habitats of M. fascicularis apparently are seashore, riverbanks, and swamp forest (Table 26). Along the seashore, mangrove forest is reported as a frequent habitat of this species from Bangladesh in the north (M. A. R. Khan & Wahab, 1983, p. 102) to Flores in the south (Auffenberg, 1 98 1 , p. 242): the only oth- er species of monkeys that regularly inhabit man- grove forest in Southeast Asia are two colobines, Trachypithecus cristatus and Nasalis larvatus. Ma- caca fascicularis also has been reported inland in lowland secondary and primary forest and, less frequently, in upland secondary and primary for- est. At nearly 2000 m in northern Sumatra, Volz (1912, p. 88) reported, "To my astonishment, up here in this cool damp high-altitude vegetation, I still observed several monkeys, namely Macacus cynomolgus [= M. fascicularis]" (translated from German). This species also inhabits heavily dis- turbed areas, including the immediate vicinity of villages and farms; following logging, the density of M. fascicularis sometimes increases (Johns, 1992, p. 438). Arboreality/Terrestriality On the seacoast and riverbanks, M. fascicularis often forages on the ground (25% of encounters in Sumatra [Crockett & Wilson, 1980, p. 165], 26.5% of encounters in Kalimantan [Fittinghoff & Lind- burg, 1980, p. 189]). In inland forests, M. fasci- cularis is predominantly arboreal (98% in West Malaysia [Bernstein, 1967, p. 204; Aldrich-Blake, 1980, p. 158], ca. 97% in Kalimantan [Wheatley, 1980, p. 216]). This species apparently favors the lower and middle forest strata; in more than 50% of forest sightings, troops were in branches less than 20 m above the forest floor (West Malaysia [Aldrich-Blake, 1980, p. 159; cf. Harrison, 1961, p. 14], Sumatra [Rijksen, 1978, p. 127], Kaliman- tan [Rodman, 1978, p. 476]). Observations in Sumatra indicate that smaller groups of M. fasci- cularis tend to occur higher in the canopy (van 50 FIELDIANA: ZOOLOGY Schaiketal., 1983b, p. 216; D. R. Vosetal., 1992, p. 388). In response to danger, troops may flee either on the ground (Burma [Tickell, 1854-1875, p. [17], Great Nicobar I. [Das & Ghosal, 1977, p. 265], Sumatra [Volz, 1912, p. 369; Ulmer in Mil- ler, 1942, p. 127], Lesser Sunda Islands [B. Rensch, 1930, p. [17]; Mertens, 1936, p. 320]), in the can- opy (Thailand [Fooden, [1975], p. 100], West Ma- laysian [Bernstein, 1967, p. 204], P. Simeulue [Su- gardjito et aL, 1989, p. 200], Sumatra [Crockett & Wilson, 1980, p. 165], Kalimantan [Kurland, 1973, p. 250; Wheatley, 1980, p. 216]), or partly on the ground and partly in the canopy (Thailand [Food- en, [1975], p. 100]). Accidental falls of this species out of trees have been observed in northern Su- matra (Karssemeijer et al., 1990, p. 286). Details of locomotor behavior are discussed by Kurland (1973, p. 250), Fleagle (1980, p. 198), Cant (1988, p. 31), and Cannon and Leighton (1994, p. 512). At night, M. fascicularis usually sleeps in tall, rel- atively bare trees near a river (Singapore [Ridley, 1906, p. 142], Sumatra [Moszkowski, 1909, p. 24], Sarawak [Hornaday, 1910, p. 358], and numerous later accounts, particularly Fittinghoff & Lind- burg, 1980, p. 190 -Kalimantan). Swimming Observers agree that M. fascicularis frequently enters bodies of water, apparently often for pleas- ure, and that it is an excellent swimmer. In Burma, a wounded male that was attempting to escape from humans swam about 50 m underwater (Tick- ell, 1854-1875, p. [18]). In a captive insular group of M. fascicularis in Japan, a young male that was defeated in a dominance fight apparently swam 100 m to a breakwater (Furuya, 1965, p. 325). However, during the course of 3 years in the same captive insular group, most of about 60 individ- uals that were driven away by dominant group members died by drowning; none apparently reached the next nearest island, which is about 250 m distant and which may not have been vis- ible to the escaping monkeys. Drownings also have been reported in natural populations in coastal Thailand (Aggimarangsee, 1992, p. 130). Troop Size and Composition Mean size of nonprovisioned troops of M. fas- cicularis generally varies from about 12 to 25 (Ta- ble 27). Mean troop size is exceptionally small (ca. 5) in a highly disturbed area in Vietnam and ex- ceptionally large in western Java (ca. 47, n = 5 troops) and Mauritius (77.5, n = 2 troops, intro- duced population). Crockett and Wilson (1980, p. 149) suggested that troop size in Sumatra may be smallest in mangrove forest, larger in inland pri- mary forest, and largest in inland secondary forest (cf. Ruiter, 1993, pp. 90-91), but the observed differences are acknowledged not to be statistically significant. In provisioned troops in Thailand, troop size (mean = 76.9,n = 33 troops) generally exceeds that in nonprovisioned troops (Aggimar- angsee, 1992, p. 150). Solitary males, not associ- ated with any troop, have been reported in Thai- land (Fooden, 1971, p. 24), Singapore (Furuya, 1965, p. 294), Sumatra (Wilson & Wilson, 1973, p. 5; Rijksen, 1978, p. Ill; Norikoshi, 1984, p. 5), Sabah (Kurland, 1973, p. 253), Java (Hoog- erwerf, 1970, p. 408), and Mindanao (Hoogstraal, 1951, p. 45). In sample areas studied, the pooled ratio of sexually mature males to sexually mature females varies from 0.22 to 0.90 (Table 28); within troops, known sex ratios vary from 0.14 to 1.67. During a 10-year period of investigation at Kua- la Lompat, West Malaysia, the size of local troops tended to remain relatively constant (Chivers & Raemaekers, 1980, p. 25 1). However, troops often temporarily subdivide into smaller units while for- aging (West Malaysia [Aldrich-Blake, 1980, p. 147], Sumatra [van Schaik et al., 1983b, p. 214], Kali- mantan [Kurland, 1973, p. 253; Rodman, 1978, p. 470; Wheatley, 1978, p. 348]). At each of two localities in Sumatra, a large troop has been ob- served to permanently subdivide into two un- equal-sized daughter troops (van Schaik et al., 1983a, p. 174; Norikoshi, 1984, p. 6). Home Range, Day Range Reported home-range area in nonprovisioned troops varies from 1 2.5 ha in P. Simeulue to more than 300 ha in West Malaysia (Table 29). Overlap between home ranges of adjacent troops generally is slight, but the home range of a daughter troop may overlap extensively with that of the troop from which it split (van Schaik & van Noordwijk, 1988, p. 80), and home ranges of troops in some small islands also may overlap extensively (Angst, 1973, p. 625). Although home ranges of some ad- jacent troops are separated by rivers (van Noord- wijk & van Schaik, 1 985, p. 850), the home ranges of a pair of adjacent troops in Borneo overlapped on both sides of a river (Fittinghoff & Lindburg, 1980, p. 207). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 51 Table 26. Recorded habitats of Macaca fascicular is. Sample area Lowland forest Upland1 forest _, , . ( nlti- Popu- lated Man- Other Riv- Sec- Sec- vated area; Refer- grove coastal erine ondary Primary ondary Primary area temple ence2 Core area X 7 X X X X 2 X X X X X X X X X 3 4 X X X X X X 5 X X X X X X X X 6 X X X X X XXX X 7 Bangladesh Burma Thailand, > 10°N Vietnam Thailand, < 10°N West Malaysia Sumatra Borneo Java Mergui Archipelago Con Son Tioman Singapore Riau-Lingga Archipelago3 Kepulauan Natuna4 Banggi group5 Bali Nicobar Islands6 Simeulue Nias Lesser Sundas7 Philippines Shallow-water fringing islands Deep-water fringing islands 10 11 12 13 14 15 16 17 18 19 20 21 22 1 Altitude > 500 m. 2 Key to references: 1, M. A. R. Khan & Wahab, 1983, p. 102. 2, Heifer, 1838, p. 858; Phayre in Blyth, 1844, p. 475; Mason, 1851, p. 220; Tickell, 1854-1875, p. [17]; Shortridge in Wroughton, 1915, p. 700, 3, Gairdner, 1914, p. 28; Kloss, 1916b, p. 32; Gyldenstolpe, 1917b, p. 6; Fooden, 1971, p. 15; Lekagul & McNeely, 1977, p. 293; Eudey, 1980, p. 75; Aggimarangsee, 1992, p. 119. 4, Van Peenen et al., 1971, p. 134; Lippold, 1977, p. 521. 5, Robinson in Bonhote, 1903, p. 4; Fooden, [1975], p. 100; Boonratana, 1988, pp. 73 fF.; Aggimarangsee, 1992, p. 137; Eudey, 1994, p. 273. 6, Kelsall, 1894b, p. 16; Rower, 1900, p. 316; Shebbeare, 1940, p. 57; Tweedie & Harrison, 1954, p. 16; Furuya, 1965, p. 287; Bernstein, 1967, p. 198; Harrison, 1969, p. 176; Medway, 1972b, p. 120; Southwick & Cadigan, 1972, p. 8; Olivers & Davies, 1979, pp. 19-20; Lim & Sasekumar, 1979, p. 106; J. R. MacKinnon & MacKinnon, 1980, pp. 176, 186; Mah & Aldrich-Blake, 1980, p. 354; J. R. Marsh & Wilson, 1981, p. 232; Johns, 1986b, p. 207. 7, Snelleman, 1887, p. 10; Hagen, 1890, p. 80; Miller, 1902a, p. 158; Schneider, 1905, p. 72; Moszkowski, 1909, p. 24; Volz, 1912, p. 88; H. C. Robinson & Kloss, 19 18, p. 6; F. A. Ulmer, Jr., in Miller, 1942, p. 129; Wilson & Wilson, 1973, p. 5; Rijksen, 1978, p. Ill; van Schaik & van Noordwijk, 1985a, p. 141; Ghiglieri, 1986, p. 108; Bismark, 1992, p. 13; Yanuar & Sugardjito, 1993, p. 34. 8, Wallace, 1869, pp. 82, 326; Hose, 1893, p. 8; J. Buttikofer in Jentink, 1897, p. 39; Hornaday, 1910, p. 358; Shelford, 1916, p. 10; Mjoberg, 1930, p. 25; Allen & Coolidge, 1940, p. 147; Burgess, 1961, p. 146; Davis, 1962, p. 57; Yoshiba, 1964, p. 25; Kawabe & Mano, 1972, p. 216; Kurland, 1973, p. 247; Banks, 1978, pp. 166 ff.; Mittermeier, 1980, p. 252; Joines, 1981, p. 9; Chivers & Burton, [1991], p. 140; Rodman, 1991, p. 364; Cannon & Leighton, 1994, p. 509, 9, J. J. Menden, 3 Jan.-8 Apr. 1933, amnh 101811, 102015, 102017-102022 (Linggajati; Ciremay); Hoogerwerf, 1970, p. 408; Bismark, 1992, p. 13. 10, Carpenter, 1887, p. 53. 11, Van Peenen et al., 1970, p. 421. 12, Medway, 1966, p. 16; D. W. Lee, 1977, p. 21. 75, Ridley, 1895, p. 24; Chasen, 1924a, p. 78 (cf. zrc 4-089); Chuang, 1973, p. 3; Lucas & Corlett, 1991, p. 203. 14, W. L Abbott, 23 Jul. 1899, usnm 101603 (P. Lingga); Abbott in Miller, 1906c, p. 281. 75, A. Everett in Thomas & Hartert, 1894, p. 654. 16, Shukor Md. Nor, pers. comm., 1 1-20 Jul. 1991. 17, Anonymous, 1931, p. 457; Angst, 1975, p. 372; Wheatley, 1988, p. 517; Wheatley & Harya Putra, 1994, p. 247. 75, W. L Abbott, 25-27 Feb., 23 Mar. 1901, usnm 1 1 1795-1 1 1797, 1 1 1799 (field catalog); Kloss, 1903a, pp. 1 14, 150; Abdulali, 1967, p. 143; Kalra, 1980, p. 50; Das & Ghosal, 1977, p. 265. 19, W. L. Abbott, 18 Nov. 1901, usnm 1 14162 (field catalog); van Schaik & van Noordwijk, 1985a, p. 140; Sugardjito et al., 1989, p. 201. 20, F. A. Ulmer, Jr., in Miller, 1942, p. 129. 21, Mertens, 1936, pp. 318-320; Pfeffer, 1959, p. 198; Auffenberg, 1981, p. 242; Kitchener et al., 1990, p. 98. 22, Cf. Fooden, 1991, p. 20. 3 Information available for P. Sugi and P. Lingga. 4 Information available for P. Serasan and P. Natuna Besar. 52 FIELDIANA: ZOOLOGY Table 27. Recorded size of nonprovisioned troops of Macaca fascicular is. Troop size Number of troops Sample area Mean Minimum Maximum Reference1 Core area Bangladesh 16.0 9 29 6 1 Burma ca. 15 5 20 ? 2 Thailand ca. 25 < 10 ca. 100 16 3 Vietnam ca. 5 3 ca. 10 ? 4 West Malaysia ca. 25 14 ca. 70 > 8 5 Sumatra ca. 20 6 ca. 65 63 6 Borneo < 20 < 10 30 > 33 7 Java ca. 47 39 Deep-water fringing islands 58 5 8 Little Nicobar I. ca. 25 ca. 20 ca. 30 ? 9 Great Nicobar I. ca. 18 6 30 6 10 Simeulue 12.5 10 15 10 11 Nias ca. 12 ? ? ? 12 Flores 15 8 32 4 13 Philippines ca. 18 ca. 5 ca. 50 > 7 14 Introduced populations Mauritius 77.5 66 89 2 15 Angaur ca. 15 ca. 10 ca. 20 9 16 1 Key to references: 1, M. A. R. Khan & Wahab, 1983, p. 104. 2, Tickell, 1854-1875, p. [17]; Yin, 1967, p. 9. 3, Fooden, 1971, pp. 15, 25; [1975], p. 100; Lekagul & McNeely, 1977, p. 293; Eudey, 1980, pp. 75-76. 4, Van Peenen et al., 1971, p. 134; Lippold, 1977, p. 521. 5, Furuya, 1965, p. 291; Bernstein, 1967, p. 199; Aldrich-Blake, 1980, p. 149; J. R. MacKinnon & MacKinnon, 1980, p. 168; Johns, 1986a, p. 685; cf. Flower, 1900, p. 316. 6, Snelleman, 1887, p. 10; F. Kurt & W. Sinaga in Angst, 1975, p. 332; Crockett & Wilson, 1980, p. 157; van Schaik & van Noordwijk, 1986, p. 297; Ruiter, 1993, p. 90. 7, Davis, 1962, p. 58; Yoshiba, 1964, p. 25; Kurland, 1973, p. 251; Macdonald, 1982, pp. 63, 71; Chivers & Burton, [1991], p. 141; A. Suzuki, 1991, p. 53. 8, Hoogerwerf, 1970, p. 408; Angst, 1975, p. 337. W. L. Abbott, 27 Feb. 1901, usnm 111797 (field tag). 10, Das & Ghosal, 1977, p. 266. 11, van Schaik & van Noordwijk, 1985a, p. 142. 12, F. A. Ulmer, Jr., in Miller, 1942, p. 129. 13, Auffenberg, 1981, p. 242. 14, Fooden, 1991, p. 18. 15, Sussman & Tattersall, 1986, p. 36. 16, Matsubayashi et al., 1987, p. 84. Reported mean day-range length varies from 325 m to 1,900 m (Table 29). In Sumatra, day- range length is positively correlated with troop size (van Schaik et al., 1983a, p. 176). Following a severe forest fire in Kalimantan, the mean day- range length in one troop declined from about 1 ,450 m to 1,300 m (Berenstain, 1986, p. 260). Density In a broad survey of M. fascicularis in Sumatra conducted in 1971-1973, reported density varied from 0.60 troops/km2 (11.2 individuals/km2) in hill scrub-grassland to 7.71 troops/km2 (143.4 in- dividuals/km2) in Rhizophora mangrove swamp (Crockett & Wilson, 1980, p. 160); overall mean density was 2.98 troops/km2 (55 individuals/km2) in 1 1 1 .45 km2 surveyed. Roughly similar densities have been recorded in Bangladesh (M. A. R. Khan & Wahab, 1983, p. 104), West Malaysia (Chivers & Davies, 1979, p. 17; J. R. MacKinnon & MacKinnon, 1980, p. 168; Marsh & Wilson, 1981, p. 232; cf. Southwick & Cadigan, 1972, p. 8), Su- matra (J. R. MacKinnon, 1973, p. 240; Rijksen, 1978, p. 122; van Schaik & van Noordwijk, 1985a, p. 142; Bismark, 1992, p. 14; Yanuar & Sugardjito, 1993, p. 35), Kalimantan (Kurland, 1973, p. 251; Wilson & Wilson, 1975, p. 254; Wheatley, 1982, p. 205; cf. Rodman, 1978, p. 472; A. Suzuki, 1991, p. 53), and Java (Bismark, 1992, p. 14). Investigators who have studied the density of this species in primary and secondary forest 5 Information available for P. Balambangan, P. Banggi, P. Malawali, and P. Maliangin Besar. 6 Information available for Katchall I., Little Nicobar I., and Great Nicobar I. 7 Information available for P. Lombok, P. Sumbawa, P. Rintja, and P. Flores. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 53 Table 28. Ratio of sexually mature males to sexually mature females reported in nonprovisioned troops of Macaca fascicularis. Number of Number of troops sexually mature individuals Troop sex ratios Pooled sex ratio Sample area Minimum Maximum Reference1 Bangladesh 6 55 0.17 0.25 0.22 7 Burma ? ? 0.20 0.25 0.22 2 West Malaysia 1 13 — — 0.62 3 Sumatra 7 116 0.69 1.67 0.90 4 Borneo 6 44 0.44 1.00 0.57 5 Java 4 99 0.62 1.00 0.83 6 Simeulue 10 85 0.14 1.00 0.55 7 1 Key to references: 1, M. A. R. Khan & Wahab, 1983, p. 104. 2, Tickell, 1854-1875, p. [17]. 3, Aldrich-Blake, 1980, p. 149. 4, van Noordwijk & van Schaik, 1985, p. 850; Ruiter, 1993, pp. 98, 99. 5, Kurland, 1973, p. 251; Macdonald, 1982, p. 63; Cannon & Leighton, 1994, p. 509. 6, Angst, 1975, p. 337. 7, Sugardjito et al., 1989, p. 202. agree that the density is greater— by a factor of 1.3-9.8— in secondary forest. In P. Simeulue, a deep-water island west of Sumatra, density (> 100 individuals/km2) reportedly is about twice that in comparable habitats in Sumatra (van Schaik & vanNoordwijk, 1985a, p. 142). Density reportedly also is high (> 400 individuals/km2) in P. Pe- utjang, a small shallow- water island off the western tip of Java (Angst, 1975, p. 329). In provisioned troops at Ubud, Bali, density is about 1,600 in- dividuals/km2 (Wheatley, 1989, p. 65). Based on rough estimates of population density and remaining available habitat, K. S. MacKinnon (1986, p. Ill) calculated provisionally that the population of M. fascicularis in Indonesia at that time was 3,726,860, and J. R. MacKinnon and MacKinnon (1 987, p. 1 89) calculated that the pop- ulation in mainland Southeast Asia north of West Malaysia was 309,360. If these provisional cal- culations are reasonable, the total population of this species about 1 0 years ago in its entire natural range— which, in addition to Indonesia and main- land Southeast Asia north of West Malaysia, in- cludes the Nicobar Islands, Malaysia, Brunei, and the Philippines-may have been approximately 5 million. Diet Fruits, of at least 185 species (Lucas & Corlett, 1991, p. 205; 1992, p. 45; cf. Ungar, 1994, p. 217; 1995, p. 232), apparently are the main food of M. fascicularis (Table 30). Ingestion of fruit frequent- ly (ca. 57% of observations) involves use of the incisors (Ungar, 1994, p. 210). When M. fasci- cularis feeds on fruit, seeds that are more than about 4 mm wide are separated from the sur- rounding pulp and rejected, either manually or by spitting, whereas smaller seeds usually are swal- lowed and defecated intact (Corlett & Lucas, 1 990, p. 167). Macaca fascicularis also consumes leaves and other plant parts, invertebrates, and small verte- brates. Widely reported invertebrate prey includes crustaceans, bivalves, and snails, along the sea- coast and riverbanks, and insects inland. Oysters sometimes are broken open by smashing with a stone that a monkey may transport up to 75 m for this purpose (Carpenter, 1887, p. 53). The capture and consumption of an adult sparrow by a captive M. fascicularis in the Calcutta Zoo is described by Mandal (1990, p. 435). This species often raids cultivated crops (Kloss, 1 903a, p. 1 28; Miller, 1 903a, p. 438; Das & Ghos- al, 1977, p. 266; Wilson & Wilson, 1977, p. 211). In Sumba, it has learned to eat the fruit of Opuntia elatior, an introduced American cactus (Dam- merman, 1928, p. 301). Relative proportions of various components of the natural diet vary seasonally and in response to severe environmental perturbations, such as forest fires (Aldrich-Blake, 1980, p. 160; Beren- stain, 1986, p. 258). These proportions also vary according to the monkey's age and sex (van Schaik & van Noordwijk, 1986, p. 305). Feeding is esti- mated to occupy 1 3-5 5% of the daily waking hours of M. fascicularis (Table 31). Predators Known natural predators on M. fascicularis in- clude crocodiles (^2 species), the Komodo giant monitor, the python, the Philippine eagle, leop- 54 FIELDIANA: ZOOLOGY Table 29. Estimated home range and day range in troops of Macaca fascicularis. Home range Day range Number Sample area Habitat (ha) (m) of troops Reference1 Nonprovisioned troops Bangladesh Mangrove 80 325 6 / West Malaysia Mangrove 200 — 1 2 Lowland forest 46.2 1,400 ? 3 Lowland forest 41 1,062 1 4 Lowland forest 14 100 5 5 Lowland forest > 300 — ? 5 Sumatra Mangrove >25 — ? 6 Nonmangrove 75 — ? 6 Lowland forest 50 1,500 7 7 Borneo Lowland forest 80 700 1 8 Lowland forest 125 1,900 1 9 Lowland forest 112.5 1,4502 1 10 Lowland forest 60 — 11 11 Simeulue Lowland forest 12.5 Semiprovisioned troops ~ < 10 12 West Malaysia Mangrove 35 — 1 13 Singapore Lowland forest 33 Urban troop 1 14 West Malaysia Golf course 7.2 1,009 1 4 1 Key to references: /, M. A. R. Khan & Wahab, 1983, pp. 104, 107. 2, Furuya, 1965, p. 288. 3, J. R. MacKinnon & MacKinnon, 1980, p. 168. 4, Mah & Aldrich-Blake, 1980, pp. 355-356. 5, Caldecott, 1986a, p. 155. 6, Crockett & Wilson, 1980, p. 168. 7, van Schaik et al., 1983a, p. 176; 1983b, p. 213. 8, Kurland, 1973, p. 252. 9, Wheatley, 1980, p. 233. 10, Berenstain, 1986, pp. 257, 260. 11, A. Suzuki, 1991, p. 53. 12, van Schaik & van Noordwijk, 1985a, p. 141; Sugardjito et al., 1989, p. 201. 13, Lim & Sasekumar, 1979, p. 107. 14, Lucas & Corlett, 1991, p. 203. 2 Mean for 1 1 mo prior to a major forest fire; the mean for 4 mo after the fire is 1,300 m. ards (2 species), and the tiger. Crocodiles (Croc- odylus sp., Tomistoma schlegeli) have been ob- served preying on M. fascicularis in Thailand (Mouhot, 1864, p. 152), Sumatra (Volz, 1912, p. 369), and Borneo (Shelford, 1916, p. 10; Mjoberg, 1930, p. 25; Galdikas & Yeager, 1984, p. 50). The Komodo giant monitor (Varanus komodoensis) is known to prey on M. fascicularis in Flores and nearby P. Rintja (Hoogerwerf, 1955, p. 26; Dar- evsky & Kadarsan, 1964, p. 1358; Pfeffer, 1959, p. 231; Auffenberg, 1981, pp. 228, 243; Sumardja, 1981, p. 4). A python {Python sp.) was observed to take a juvenile M. fascicularis in northern Su- matra (van Schaik et al., 1983b, p. 220). The Phil- ippine eagle (Pithecophaga jefferi) has been ob- served preying on monkeys in Samar and Min- danao (Fooden, 1991, p. 18). In western Java, leopards (Panthera pardus) are reported to prey on M. fascicularis in Ujung Kulon National Park and at Gunung Pangrango (Bartels, 1929, p. 81; Hoogerwerf, 1970, p. 402), and in eastern Java, leopards and tigers (P. tigris) are reported to prey on this monkey in Meru-Betiri Reserve (Seiden- sticker, 1983, p. 324); clouded leopards (Neofelis nebulosa) are reported to prey on M. fascicularis in Borneo (Banks, 1931, p. 77). Domestic dogs also attack and sometimes kill M. fascicularis (Ridley, 1895, p. 25; Seidensticker, 1983, p. 325), and humans are known to hunt this species for food (Labang & Medway, 1979, p. 56; Aggimar- angsee, 1992, p. 154). Under experimental conditions, hi. fascicularis exhibited fear of a stuffed python and a snake model (van Schaik & Mitrasetia, 1990, p. 105; Vitale et al., 1991, p. 281). In nature, M. fascicularis has been observed to mob a python (van Schaik & Mitrasetia, 1990, p. 106) and to engage in antag- onistic behavior directed toward crocodiles, Ko- modo giant monitors, and domestic dogs (Mou- hot, 1864, vol. 1, p. 152; Hagen, 1890, p. 81; Auffenberg, 1981, p. 243; Wheatley, 1991, p. 172). Field studies indicate that predator detection is a major determinant of social behavior and group size in M. fascicularis (van Schaik et al., 1983b, p. 220; van Schaik & van Noordwijk, 1985a, p. 139). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 55 Table 30. Foods reported eaten by nonprovisioned troops of Macacafascicularis; dietary proportions are indicated where available. Foods eaten Crustaceans, Other Fruit, Other Insects, mollusks, animal Refer- Sample area Leaves seeds plant parts spiders fish prey ence1 Core area Bangladesh 20% 4% 25% 51% ; Burma X X X X 2 Thailand X X X 3 West Malaysia X X X X X X2 4 16.1% 52.4% 8.3% 23.3% 5 24.0% 63.7% 8.8% 4.4% X3 6 Sumatra X X X X X 7 4.4% 63.3% 18.1% 84 Borneo X X X X X X' 9 8% 87% 3.5% 1% 10 Shallow-water fringing islands Mergui Archipelago X X 11 Singapore X X X 12 Banggi X 13 Deep-water fringing islands Nicobar Islands X X 14 Lesser Sundas X X X X 15 Philippines X X X* 16 Introduced population Mauritus 9% 70% 14% 5%7 17* 1 Key to references: 1, M. A. R. Khan & Wahab, 1983, p. 106. 2, Heifer, 1838, p. 858; A. P. Phayre in Blyth, 1844, p. 475; Mason, 1851, p. 220; Tickell, 1854-1875, p. [17]; Blyth, 1859, p. 275; G. C. Shortridge in Wroughton, 1915, p. 700; H. C. Smith, 21-30 Apr. 1936, bm(nh) 1936.9.10.8, 9.10.10, 9.10.14 (field tags); Pocock, 1939, p. 82. 3, H. C. Robinson in Bonhote, 1903, p. 4; Gairdner, 1914, p. 28; Gyldenstolpe, 1914, p. 3; Fooden, 1971, p. 24; [1975], p. 99; McNeely, 1977, p. 10; Lekagul & McNeely, 1977, p. 293. 4, Tweedie & Harrison, 1954, p. 16; Harrison, 1961, p. 7; Furuya, 1965, p. 289; Lim & Sasekumar, 1979, p. 109 (semiprovisioned troop); Lambert, 1990, p. 455. 5, Aldrich-Blake, 1980, p. 160. 6, J. R. MacKinnon & MacKinnon, 1980, p. 178. 7, Volz, 1912, p. 369; F. A. Ulmer, Jr., in Miller, 1942, p. 127; Rijksen, 1978, p. 112; Crockett & Wilson, 1980, p. 164; Whitten et al., 1984, p. 136; van Schaik & van Noordwijk, 1988, p. 82. 8, Ungar, 1994, p. 210. 9, Shelford, 1916, p. 10; Mjoberg, 1930, p. 25; Kern, 1964, p. 185; Roedelberger & Groschoff, 1967, p. 30; Kurland, 1973, p. 254; Rodman, 1978, p. 469; Fittinghoff & Lindburg, 1980, p. 190; Wheatley, 1980, p. 216; Joines, 1981, p. 9; Macdonald, 1982, p. 63; Leighton & Leighton, 1983, p. 185; Berenstain, 1986, p. 257; 10, Wheatley, 1978, p. 348; reported diet also includes 0.5% clay. 11, Carpenter, 1887, p. 53; H. C. Smith, 20-23 Apr. 1936, bm(nh) 1936.9.10.4-6, 1936.9.10.11. 12, Ridley, 1895, p. 24; Chasen, 1924a, p. 79; Lucas & Corlett, 1991, p. 203 (semiprovisioned troop). 13, Shukor Md. Nor, pers. comm., 11 Jul. 1991. 14, Kloss, 1903a, p. 129; Das & Ghosal, 1977, p. 266; Kalra, 1980, p. 50. 75, Dammerman, 1928, p. 301; Pfeffer, 1959, p. 198; Auffenberg, 1981, p. 242. 16, Fooden, 1991, p. 18. 17, Sussman & Tattersall, 1981, p. 200. 2 Sipunculid worms; lizards. 3 Frogs. 4 14.2% of diet is unspecified. 5 Birds' eggs or nestlings. 6 Earthworms; lizards; birds. 7 Includes all invertebrates. 8 2% of diet is unspecified. Intertroop Behavior Judging from reports of long-term observations, direct encounters between adjacent nonprovi- sioned troops are relatively rare (West Malaysia [Aldrich-Blake, 1 980, p. 1 5 1], Sumatra [van Schaik & van Noordwijk, 1988, p. 80], Kalimantan [Fit- tinghoff & Lindburg, 1980, p. 206], Java [Angst, 1973, p. 626]), which suggests mutual avoidance. In Sumatra, however, small daughter troops fre- quently encounter the large troops from which they have split (van Schaik & van Noordwijk, 1988, p. 56 FIELDIANA: ZOOLOGY Table 31. Daily waking-hour1 time budget estimates (%) for Macaca fascicularis.2 Activity Sample area Feeding Travel Other Reference Bangladesh Ebb tide Full tide West Malaysia Kalimantan Mauritius (introduced) 55 22 35 13 32 15 3 20 45 4 30 75 453 42 64 M. A. R. Khan & Wahab, 1983, p. 106 Aldrich-Blake, 1980, p. 161 Wheatley, 1980, p. 221 Sussman & Tattersall, 1981, p. 197 1 Estimates of total daily waking hours: West Malaysia, 13 hr (J. R. MacKinnon & MacKinnon, 1980, p. 185); Sumatra, 12.5 hr (van Schaik & van Noordwijk, 1988, p. 85); Kalimantan, 1 1.4 hr (Wheatley, 1980, p. 222). 2 Cf. Lim and Sasekumar (1979, p. 1 1 1), J. R. MacKinnon and MacKinnon (1980, p. 185), and Leon et al. (1993, p. 177). 3 Computed by subtraction. 80), and in P. Peutjan, an islet off western Java that is densely populated by M. fascicularis (see above), adjacent troops are often intermixed (Angst, 1 973, p. 626). Intertroop displays that may function to maintain intergroup spacing have been reported in West Malaysia and Kalimantan. In P. Simeulue, west of Sumatra, M. fascicularis has a unique rapid bark that probably functions in in- tertroop communication (van Schaik & van Noordwijk, 1985a, p. 143). During intertroop en- counters in Sumatra, copulations between mem- bers of different troops have been observed infre- quently (van Noordwijk, 1985, p. 289). In Brunei, northern Borneo, a larger troop was observed to displace a smaller troop (Macdonald, 1982, p. 71). In artificially provisioned troops, intertroop fight- ing is frequent (Furuya, 1 965, p. 288; Angst, 1975, p. 373; Norikoshi, 1984, p. 1 1; Koyama, 1984, p. 25; Wheatley, 1991, p. 171). Macaca fascicularis males have been observed to leave one troop and take up residence in another troop in northern Sumatra (Rijksen, 1978, p. Ill; van Noordwijk & van Schaik, 1985, p. 85 1), west- ern Sumatra (Norikoshi, 1 984, p. 2; Koyama, 1 984, p. 20; 1985, p. 1 19), eastern Kalimantan (Wheat- ley, 1982, p. 207; Berenstain, 1986, p. 258), and Bali (Koyama et al., 1981, p. 8). Such intertroop movement by males presumably is universal in natural populations of this species, as in other spe- cies of macaques. Males apparently leave their na- tal troop as late juveniles or subadults, before about age 7 years, and most males probably change troops several more times during their life; intertroop movement apparently precludes close inbreeding in natural troops (Ruiter et al., 1992, p. 186). In six troops studied in northern Sumatra, 40 of 44 adult and subadult (immigrant) males that were present in January 1 980 had emigrated by March 1984. In western Sumatra, the mean duration of troop residence of an adult male was estimated to be 1 .2 years. Most intertroop movements by males in northern Sumatra occurred near the beginning of the peak copulatory season. The maximum re- corded interval between a male's emigration from one troop and immigration into another troop was 2 months (semisolitary interval). Immigration of a new male often is followed by changes in a troop's dominance hierarchy. Females also occasionally move from one troop to another, but apparently at a much lower frequency than males (5 of 57 recorded migrations in northern Sumatra; 1 of 7 in western Sumatra; 0 of 1 1 in eastern Kalimantan; 0 of 2 in Bali; total, 6 of 77 = 7A Interspecific Behavior Interactions have been reported between M. fas- cicularis and M. nemestrina, leaf monkeys (Pres- bytis spp., Trachypithecus spp.; cf. Groves, 1993, p. 270), Nasalis larvatus, Hylobates spp., and Pon- S° pygmaeus. In a 3 -km2 study area in eastern Kalimantan, Rodman (1973, p. 657) found that the local occurrence of M. fascicularis was nega- tively correlated with that of M. nemestrina, Pres- bytis aygula (= P. hosei), Hylobates moloch, and Pongo pygmaeus. Noncompetitive associations of M. fascicularis and M. nemestrina have been observed in West Malaysia (Bernstein, 1967, p. 201), and seven mixed troops of these species have been recorded in Sumatra (Rijksen, 1978, p. Ill, single M. fas- cicularis individuals in six M. nemestrina troops; Crockett & Wilson, 1980, p. 175, M. nemestrina individual in M . fascicularis troop). These two spe- cies, however, generally occupy different ecologi- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 57 cal niches; M. fascicularis is rare in primary ev- ergreen rain forest, which is favored by M. ne- mestrina (Rodman, 1973, p. 656; 1978, p. 476; 1991, p. 364; J. R. MacKinnon & MacKinnon, 1978, p. 316; Crockett & Wilson, 1980, p. 162). Macaca fascicularis and Trachypithecus crista- tus frequently have been observed together in West Malaysia (Furuya, 196 1-1 962b, p. 56; Bernstein, 1968b, p. 8; Lim & Sasekumar, 1979, p. 108), Sumatra (Wilson & Wilson, 1973, p. 5), and Java (Hoogerwerf, 1970, p. 408, including three inter- specific mixed groups). Relations between these species reportedly are generally peaceful, but M. fascicularis often displaces T. cristatus. Hooger- werf (1970, p. 408) observed one violent inter- specific fight in Java, during which an immature T. cristatus ". . . fell to the ground torn to pieces." Associations of M. fascicularis and Presbytis melalophos have been observed in West Malaysia (Bernstein, 1967, p. 201; J. R. MacKinnon & MacKinnon, 1978, p. 319) and Sumatra (Snelle- man, 1887, p. 10; Koyama, 1985, p. 105). Most accounts indicate that these associations are peaceful . Koyama (1985, p. 105) reported that these two species often share the same sleeping tree, but J. R. MacKinnon and MacKinnon (1978, p. 319) indicated that M. fascicularis displaces P. mela- lophos from feeding and sleeping trees. Bernstein (1967, p. 201) reported noncompeti- tive association of M. fascicularis with Trachypi- thecus obscurus in West Malaysia, and Kurland (1973, p. 254) reported noncompetitive associa- tion with Presbytis aygula (= P. hosei) in Kali- mantan. Ridley (1895, p. 25) reported that leaf monkeys in West Malaysia fought for more than 2 hours with M. fascicularis over occupation of a feeding tree ("biting them [M. fascicularis] and throwing them out of the tree and into the river"). Banks (1978, p. 188) reported that M. fascicularis often prevented leaf monkeys in Borneo from en- tering sleeping trees. Peaceful association of M. fascicularis and Nasalis larvatus in mangrove swamps in Borneo has been observed frequently (Davis, 1962, p. 57; Kern, 1964, p. 185; Kawabe & Mano, 1972, p. 216; Kurland, 1973, p. 254; Macdonald, 1982, p. 117). Noncompetitive association of M. fascicularis and Hylobates lar in West Malaysia was reported by Bernstein (1967, p. 201). Macaca fascicularis and Pongo pygmaeus have been observed together on 32 occasions in Sumatra (Rijksen, 1 978, p. 112) and on two occasions in Kalimantan (Fittinghoff & Lindburg, 1980, p. 201); these observations in- clude peaceful interspecific sharing of feeding and sleeping trees and rudimentary interspecific threats— a few by P. pygmaeus and one by M. fascicularis. In West Malaysia, racket-tailed drongos (Dicru- rus paradiseus) have been observed following M. fascicularis through the canopy and catching in- sects stirred up by the monkeys' activity (Ridley, 1901, p. 105; cf. Fooden, 1969, p. 52). Reproduction Although copulations, pregnancies, and births have been recorded throughout the year in natural populations of M. fascicularis, long-term studies indicate that all three of these reproductive events exhibit seasonal peaks in this species (Appendix 1 1). The timing of these reproductive peaks ap- parently varies geographically and varies between years at the same locality. In a study in northern Sumatra, the annual birth peak tended to occur during or shortly after the local fruiting peak; in years when fruit production was relatively low, the birth peak tended to be delayed by 2 or 3 months (van Schaik & van Noordwijk, 1985b, p. 538). In Thailand, on the other hand, fertile matings— not births— tend to peak shortly after the fruiting peak, which occurs in April-June (Varavudhi et al., 1989a, pp. 221-222; cf. Tangpraprutigul & Var- avudhi, 1982, p. xci; Aggimarangsee, 1992, p. 129). In a 2- to 3-year study of captive M. fascicularis, annual variation of sperm concentration in semen collected by electroejaculation was investigated (Okamoto, 1994, p. 27); results were inconclusive. In natural populations of M. fascicularis, fe- males probably become reproductively active at about age 3.5 years. A pregnant juvenile female, aged about 3.5 years, was collected in western Thailand (fmnh 99646, with erupting C,, P3, and M2; cf. Table 14; Fooden, 1 97 1 , p. 24), and females in Sumatra reportedly produce their first infants at about age 4 years (Koyama, 1985, p. 106; van Noordwijk & van Schaik, 1987, p. 586; cf. Var- avudhi et al., 1989a, p. 222). In captivity, females may copulate as early as age 2.5 years (Chance et al., 1 977a, p. 6 1 9), and one captive female is known to have given birth at age 3 years (Timmermans et al., 1981, p. 121); however, fertility in captive females usually does not begin until about age 4 years (Spiegel, 1954, p. 230; Dang, 1983, p. 38). Mean age at menarche in 43 captive females was 2.5 ± 0.7 years (Honjo et al., 1984, p. 69; cf. Dang, 1983, p. 36). Mean length of the menstrual 58 FIELDIANA: ZOOLOGY cycle in 28 captive females was 30.9 days (SD = 4.8 days, extremes = 19-43 days, mode = 28 days, n = 595 cycles) (Dukelow, 1977, p. 34; cf. Joach- imovitz, 1928, p. 462; Comer, 1932, p. 404; Spie- gel, 1954, p. 235; Fujiwara et at, 1967, p. 506; Kerber & Reese, 1 969, p. 976; Valerio et al., 1 969, p. 287; MacDonald, 1 97 1 , p. 374; Nawar & Hafez, 1972, p. 45; Dang, 1977, p. 3; Goodman et al., 1977, p. 480; Zumpe & Michael, 1 983, p. 58; Hon- jo et al., 1984, p. 63; Wallis et al., 1986, p. 87). Males in natural populations usually begin to copulate at age 5-6 years, as subadults, after they have left their natal troops (van Noordwijk, 1985, p. 288; van Noordwijk & van Schaik, 1985, p. 852; cf. Varavudhi et al., 1989a, p. 222). As pre- viously indicated (see above, Natural History), close inbreeding probably is rare or absent in nat- ural populations of M. fascicularis. In Sumatra, subadult males that were still in their natal troops accounted for only 5 of 664 observed copulations. In Bangladesh, copulation attempts by immature males were actively thwarted by adult males (M. A. R. Khan & Wahab, 1983, p. 108). In captive males, testicular descent occurs at age 2.5 years, and adult testicular size is achieved at age 4.0-4.5 years (Cho et al., 1973, p. 408; Chance et al., 1 977a, p. 6 1 9). Fertile copulations have been recorded for three captive males at ages 3.5 years (1 male; Honjo et al., 1984, p. 68) and 3.8 years (2 males; Spiegel, 1954, p. 230). At or near the onset of sexual maturity in fe- males, the bare skin between the root of the tail and the anus swells prominently and the skin around the vulva frequently reddens (Joachimov- itz, 1928, p. 464; Corner, 1932, p. 408; Spiegel, 1 954, p. 232; Nawar & Hafez, 1 972, p. 49; Fooden, [1975], p. 100; Emory et al., 1980, p. 250; van Noordwijk, 1985, p. 281; Meishvili & Chalyan, 1986, p. 14; C. M. Anderson & Bielert, 1994, p. 285). This pubertal swelling and reddening ap- parently reaches its maximum about midway through the menstrual cycle and diminishes a few days later. In postpubertal females, sexual swelling and reddening generally are less conspicuous and are highly variable in size and duration; in many fully fertile females, sexual swelling and reddening are slight or absent (cf. Pocock, 1 906, p. 558; 1 939, p. 78; Kurland, 1973, p. 258; Wilson & Wilson, 1977, p. 211; Fittinghoff & Lindburg, 1980, p. 189). However, in a carefully studied population in northern Sumatra, sexual swellings generally were larger during the peak copulation season (Jan- Jun.), and copulations and consortships of most females tended to increase with increasing size of their sexual swelling or increasing redness of their perivulval skin (van Noordwijk, 1985, pp. 281, 287). In some females, sexual swelling or redden- ing and copulations were observed during preg- nancy (cf. Deputte & Goustard, 1980, p. 96); fol- lowing parturition, swelling was not observed for several months. Variably complete and generally concordant ac- counts of copulatory behavior in M. fascicularis are available from natural populations (Tickell, 1854-1875, p. [17]; Furuya, 196 1-1 962a, p. 76; Kurland, 1973, p. 258; M. A. R. Khan & Wahab, 1983, p. 104; Koyama, 1984, p. 31; 1985, p. 107; van Noordwijk, 1985, p. 281), introduced free- ranging populations (Poirier & Smith, 1974, p. 300; Sussman & Tattersall, 1981, p. 203), colony groups (Spiegel, 1954, p. 238; Goustard, 1961, p. 313; 1963, p. 710; 1968, p. 464; Furuya, 1965, p. 313; de Benedictis, 1973, p. 1470; Angst, 1974, p. 52; Chevalier-Skolnikoff, 1975, p. 207; Emory & Harris, 1978, p. 227; Deputte & Goustard, 1980, p. 85; Emory et al., 1980, p. 251; Shively et al., 1982, p. 375), and laboratory test pairs (Kanagawa et al., 1972, p. 453; Kanagawa & Hafez, 1973, p. 234; Michael & Zumpe, 1988, p. 379; Zumpe & Michael, 1983, p. 58; 1990, p. 148). Copulations in this species have been observed throughout the day, but evidence from troops in northern Su- matra suggests that copulations may be more fre- quent in the morning (ca. 60%) than in the after- noon (van Noordwijk, 1985, p. 286; cf. de Bene- dictis, 1973, p. 1473; Sussman & Tattersall, 1981, p. 203). Copulations occur both in trees and on the ground (Kurland, 1973, p. 258; Angst, 1974, p. 53; Sussman & Tattersall, 1981, p. 203; Koy- ama, 1985, p. 117). Copulations in M. fascicularis may be initiated by either sex. A female initiates copulation by star- ing at or approaching a male and presenting her hindquarters or by stereotyped head or arm move- ments; in laboratory test pairs, female use of head or arm movements to initiate copulation peaked near the middle of the menstrual cycle (Michael & Zumpe, 1988, p. 381). A male initiates copu- lation by staring at or approaching a female and raising her tail. Both female- and male-initiated copulations usually proceed with examination by the male of the perineum of the female; this ex- amination may be visual, digital, olfactory, or, rarely, lingual. If the female is acceptable (cf. de Benedictis, 1973, pp. 1479-1480), the male mounts the female, by gripping her hips with his hands and gripping her calves with his feet, and begins pelvic thrusting. In laboratory test pairs, mounts FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 59 Table 32. Copulatory behavior data in two Macaca fascicularis groups. Provisioned natural group, Bali Colony group, USA Ejaculatory copulations observed (N) SME copulations (N) Duration of SME mount1 (sec) Thrusts per SME mount1 MME copulations (N) Duration of MME mount1 (sec) Thrusts per MME mount1 Duration of MME sequence1 (min) Mounts per MME sequence1 Reference 169 139(82.2%) 7.1 (2.6-12.0) 16.4(5-32) 30(17.8%) 6.4 (3.0-9.4) 15(6-31) 7.1 (2-35) 2.4 (2-6) Wheatley, 1991, p. 171 17 8(47.1%) 8.5 (6-14) 17(14-21) 9 (52.9%) 6(2-11) 13(4-29) 41 (2-85) 3(2-6) Shively et al., 1982, p. 376 Mean and extremes. and mounting attempts tended to increase slightly during the first half of the menstrual cycle and tended to decrease more precipitously during the second half (Zumpe & Michael, 1983, p. 66). Ejaculation in M. fascicularis may occur either as a result of a single mount (single-mount ejac- ulations [SME]) or as a result of a series of mounts separated by dismounts (multiple-mount ejacu- lation [MME]). In a provisioned natural group in Bali, 82% of observed ejaculatory copulations were SME and 1 8% were MME; in a seminatural colony group, 47% of observed ejaculatory copulations were SME and 53% were MME (Table 32; cf. Fu- ruya, 196 1-1 962a, p. 76; van Noordwijk, 1985, p. 281). In laboratory tests, the frequency of SME was higher in first copulations of test pairs than in second copulations (27% of 191 first copula- tions, 6% of 154 second copulations) (Zumpe & Michael, 1983, p. 60; cf. Kanagawa et al., 1972, p. 453). In the natural and colony groups, the du- ration of both SME and MME mounts averaged about 7 seconds, and the number of pelvic thrusts per mount averaged about 15 (cf. Furuya, 1965, p. 313; Zumpe & Michael, 1983, p. 61); similar mount durations and thrust/mount rates have been reported for natural populations of M '. fascicularis in Bangladesh (10-15 seconds, 8-10 thrusts/mount; M. A. R. Khan & Wahab, 1983, p. 108) and Ka- limantan (5-7 sec, 4-12 thrusts/mount; Kurland, 1973, p. 258) and for a free-ranging introduced population in Angaur Island ("brief," 8-10 thrusts/ mount; Poirier & Smith, 1974, p. 300). The num- ber of mounts in an MME sequence averaged about 3 in the natural and colony groups; the duration of an MME sequence averaged about 7 minutes in the natural group and about 4 1 minutes in the colony group. Copulation in M. fascicularis usually is accom- panied by a characteristic staccato female vocal- ization and sometimes by a characteristic male vocalization (Goustard, 1963, p. 710; 1968, p. 470; Furuya, 1965, p. 313; Angst, 1974, p. 61; Poirier & Smith, 1974, p. 300; Chevalier-Skolnikoff, 1975, p. 207; Deputte & Goustard, 1980, p. 83; Wheat- ley, 1 982, p. 206; 1 984, p. 391; 1991, p. 171; Zumpe & Michael, 1983, p. 60; van Noordwijk, 1985, p. 281). At or near the time of ejaculation, the male pauses, and the female usually grasps one of the male's arms or legs and turns her face toward his (cf. Kanagawa et al., 1972, p. 453). Shortly after ejaculation, the male dismounts and the male and female move apart. After copulation, a pair may remain associated in a consortship for a variable period, ranging from about 1 hour to 3 weeks (Fu- ruya, 1965, p. 314; Poirier & Smith, 1974, p. 301; Sussman & Tattersall, 1981, p. 203; van Noord- wijk, 1985, p. 283); during a consortship, most, but not all, copulations of consortship partners are with each other. In natural groups, females may copulate as often as 3-4 times per hour, through- out the day, often with many different partners (Ruiter et al., 1992, p. 177). Dominant males in M. fascicularis apparently participate in a disproportionately large share of copulations. In northern Sumatra, the highest ranking 2 of 7 males engaged in 52.7% of 659 copulations (van Noordwijk, 1985, p. 285); in western Sumatra, 2 of 7 males engaged in 60.9% of 46 ejaculatory copulations (Koyama, 1984, p. 35; cf. Norikoshi, 1984, p. 12); in eastern Kali- mantan, 1 of 6 males engaged in 50.0% of 48 cop- ulations (Wheatley, 1982, p. 207; alpha status shifted during this study); and in Bali, 1 of 4 males engaged in 50.9% of 126 copulations (Wheatley, 1991, p. 171). Similar high frequencies of copu- lation by dominant males have been reported in 60 FIELDIANA: ZOOLOGY colony groups (Furuya, 1965, p. 315; Shively et al., 1982, p. 377). In natural and colony groups, some low-ranking adult males were never ob- served to copulate. In laboratory test pairs, the copulatory performance of a subordinate male was strongly inhibited by close proximity of a sepa- rately confined dominant male (Zumpe & Mi- chael, 1990, p. 154). In West Malaysia and Kali- mantan, aggression between males reportedly in- tensified shortly before periods of increased sexual activity (Aldrich-Blake, 1980, p. 151; Wheatley, 1982, p. 210). In troops observed in northern and western Sumatra, subordinate males frequently at- tempted to interfere with the copulations of dom- inant males, who, in turn, responded aggressively (Norikoshi, 1984, pp. 9, 12; Koyama, 1985, p. 107; van Noordwijk, 1985, p. 283). The copula- tory partners of high-ranking males in Sumatra generally tended to be females who were high rank- ing, not young, and not nulliparous (van Noord- wijk, 1985, p. 289). Paternity tests, using blood-protein analysis and DNA fingerprinting, have been conducted for 45 offspring in three troops in northern Sumatra (Rui- ter et al., 1992, p. 184; cf. Ruiter, 1993, p. 99; Ruiter & van Hooff, 1993, p. 518; Ruiter et al., 1994, p. 211); these tests indicate that dominant males sire most of the offspring in natural troops, as would be expected from observed copulation frequencies. Of 42 offspring for which probable fathers could be determined, 31 probably were sired by alpha males, 7 by beta males, and only 4 by approximately 10 lower ranking males. In ar- tificial colony groups, dominant males apparently do not maintain this reproductive advantage; pa- ternity tests in one colony revealed that, despite their higher frequency of copulations, dominant males did not sire a disproportionately large num- ber of offspring (Shively & Smith, 1985, p. 131). In a large captive group of M. fascicularis, mean gestation length for 1,141 live births was 163.5 ± 5.8 (SD) days (Honjo et al., 1984, pp. 63-64); in other captive groups, reported mean gestation lengths vary from 160.0 to 168.0 days (extremes 141-225 days) (Spiegel, 1954, p. 245; T. Fujiwara & Imamichi, 1966, p. 226; Valerio et al., 1969, p. 295; MacDonald, 1971, p. 374; Dang, 1977, p. 2; 1983, p. 41; Dukelow et al., 1979, p. 44; Vara- vudhi et al., 1989a, p. 222). Mean birth weight in the large captive group was 318.2±45.2gfor563 female infants and 347.5 ± 55.2 g for 600 male infants. The ratio of male infants to female infants tends to be greater for high-ranking captive fe- males than for low-ranking females (van Schaik et al., 1989, p. 151). Most births probably occur be- tween sunset and sunrise; in a captive group main- tained in a 14-hour light/ 10-hour dark environ- ment, 90% of 1 52 observed births occurred during dark hours, 1900-0500 (M. T. Suzuki et al., 1990, p. 252; cf. Ridley, 1906, p. 142; Spiegel, 1954, p. 248; Erwin, 1977, p. 358; Banks, 1978, p. 188; Kemps & Timmermans, 1982, p. 84). In captivity, 23 infants were nursed for an av- erage of 15.1 months (SD = 3.0 mo, extremes = 9-22 mo) (Spiegel, 1954, p. 262; cf. Chance et al., 1977b, p. 31); following 9 of these 23 nursing pe- riods, the mother's lactation continued without interruption through the nursing of one or more succeeding infants. In natural populations, infants begin to obtain some of their food independently about age 3 months (Karssemeijer et al., 1990, p. 288). The mean duration of postpartum amen- orrhea in captive nursing mothers was 6.6 months (SD = 2.6 mo, extremes = 2-13 n = 28) (Spiegel, 1954, p. 262; cf. Dang, 1979, p. 377; Cho, 1981, p. 254; Honjo et al., 1984, p. 66; Varavudhi et al., 1989a, p. 221). In northern Sumatra, in six to seven nonpro- visioned troops that included 42-53 adult females, the annual birth rate (births/adult females) from 1980-1981 to 1983-1984 was 0.45, 0.82. 0.30, and 0.75; the overall annual birth rate for these 4 years was 0.58 (van Schaik & van Noordwijk, 1985b, p. 538). In western Sumatra, in three pro- visioned troops that included 22-3 1 adult females, the annual birth rate from 1980 to 1984 was 0.73, 0.48, 0.8 1 , 0. 1 1 , and 1 .00, with an overall average of 0.63 (Koyama, 1985, p. 119). High and low annual birth rates tend to occur in alternate years in both the nonprovisioned and provisioned troops. The limited available data suggest that a biennial birth cycle may be common in these Sumatran troops and that most females may tend to produce their young during the same alternate years. In provisioned troops at Ubud, Bali, where the number of adult females varied from 27 to 46, the annual birth rate reported in 1986 and 1990-1992 was 0.59, 0.32, 0.41, and 0.33 (Wheatley & Harya Putra, 1994, p. 248). In four free-ranging provi- sioned groups of M. fascicularis introduced in P. Tinjil, south of Java, the annual birth rate is es- timated to be 0.56 (Kyes, 1 993, p. 8 1). In captivity, M. fascicularis females are capable of producing young every year (Ridley, 1895, p. 24; Chance et al., 1977a, p. 612; cf. Hadidian & Bernstein, 1979, p. 440; Timmermans et al., 1981, p. 120; Dang, 1983, p. 41; Honjo et al., 1984, p. 66). Effective fertility in females probably ceases at FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 61 Table 33. Macaque fossils or subfossils collected within geographic range of Macaca fascicularis. Number of fossils or Estimated subfossils reported M. fasci- M. nemes- Macaca Refer- Locality Epoch age (Ka) cularis trina sp. ence1 West Malaysia Bukit Chintamani Holocene ? 1 1 Kota Tongkat Holocene < 4 1 1 Gua Cha, Trench 79 A Holocene ? > 1 2 Gua Cha, Trench 79B Holocene < 3 > 1 2 Gua Cha, Trench 79B Holocene 6.3 > 1 2 Gua Madu Holocene Sumatra 7 > 1 3 Djamboe Cave Holocene/Pleistocene2 ? 11 4 Lida Ajer Cave Holocene/Pleistocene2 ? 1 6 1 4 Sibrambang Cave Holocene/Pleistocene2 ? 5 74 10 4 Unspecified cave Holocene/Pleistocene2 ? 1 11 4 4 Borneo: Sarawak Paku Cave ?Holocene ? 43 5 Unspecified cave ?Holocene ? l3 5 Niah Cave Holocene4 5 21 4 6 Niah Cave Pleistocene4 15 37 6 Niah Cave Pleistocene4 25 7 6 Niah Cave Pleistocene4 Java 35 5 1 6 Sampung Holocene 55 3 7 Goea Djimbe Holocene 10 26 4 Goea Ketjil Holocene 10 1 4 Cave near Wajak Holocene 10 l6 4 Punung Pleistocene 1202 16 8 Ngandong Pleistocene7 < 800 l8 9 Sangiran (?Kabuh) Pleistocene7 800 1 1 10 Sangiran Pleistocene7 ?800 2 11 Glagahombo (Kabuh) Pleistocene7 800 1 12 Kali Brangkal (Kabuh) Pleistocene7 800 1 12 Ndangklampok (Kabuh) Pleistocene7 800 1 12 Saradan Pleistocene7 ?800 1 13 Trinil Pleistocene7 1,000 7 4 Trinil Pleistocene7 1,000 1 14 Bangle Pleistocene7 Flores ? 1,000 2 4 Liang Toge Holocene Timor10 3.59 1 15 Bui Ceri Uato Holocene < 1.5 2 16 Uai Bobo 1 Holocene 1.6 2 16 Lie Siri Holocene <4.3 l3 16 Uai Bobo 2 Holocene 4.5 4 16 1 Key to references: 1, Matthews, 1961, pp. 18, 41; Dunn, 1975, p. 122. 2, Adi, 1985, pp. 66-67. 3, Tweedie, 1940, p. 7. 4, Hooijer, 1962b, pp. 50, 54-55, 58. 5, P. H. Napier, 1981, pp. 9, 20. 6, Hooijer, 1962a, p. 440. 7, Dammerman, 1934, p. 492. 8, Badoux, 1959, p. 88; J. de Vos, 1983, p. 421; J. de Vos et al., 1994, pp. 132, 134. 9, Aziz, 1989, p. 52. 10, Aimi, 1981, p. 409. 11, Hooijer, 1964, p. 76. 12, Aimi & Aziz, 1985, pp. 161-163. 13, Deninger, 1910, p. 1; Hooijer, 1962b, p. 54; Aimi, 1981, p. 412. 14, Stremme, 1911, p. 140; Hooijer, 1962b, p. 54. 15, Hooijer, 1967, p. 160. 16, Glover, 1986, pp. 78, 121, 158, 192. 2 See J. de Vos (1983, p. 422); J. de Vos et al. (1994, p. 132). 3 Minimum number of individuals; species identification tentative. 4 For tabulation details, see Fooden (1975, p. 61). 5 See Bell wood (1985, p. 200). 6Cf. Brink (1982, p. 180). 62 FIELDIANA: ZOOLOGY about age 20 years (Spiegel, 1954, p. 265; Angst & Thommen, 1977, p. 212; van Noordwijk & van Schaik, 1987, p. 587), which suggests that an av- erage female in a natural population may produce about eight to nine infants during her reproductive life (age ca. 4—20 yr). The maximum known life span in captive M. fascicular is is 37 years 1 month (M. L. Jones, 1982, p. 1 17; cf. Dumond, 1967, p. 203; Angst, 1975, p. 350; van Noordwijk & van Schaik, 1988, p. 30). In northern Sumatra, neonatal infant mortality (deaths before age 3 mo/births) from 1980-1981 to 1983-1984 was 0.21, 0.12, 0.25, and 0.06, with an overall average of 0.14 (van Schaik & van Noordwijk, 1985b, p. 538); neonatal infant mor- tality was high in years when the birth rate was low (see above). The average infant death rate be- fore age 1 year was 0.22 in northern Sumatra (van Noordwijk & van Schaik, 1 987, p. 585); this infant death rate is comparable to that reported in a pro- visioned free-ranging group at Ubud, Bali (0.25; Wheatley & Harya Putra, 1994, p. 248), and in a large captive group of M. fascicularis (> 0. 1 5; Lu- der, 1993, p. 142). Infanticide, committed by adult males, probably is one of the important causes of infant deaths (cf. Thompson, 1967, p. 18; Wash- burn & Hamburg, 1968, p. 473; Angst & Thom- men, 1977, p. 208; Erwin, 1977, p. 359; Tim- mermans et al., 1981, p. 120; Wheatley, 1982, p. 211; Pallaud, 1984, p. 92; Koyama, 1985, p. 106; van Noordwijk & van Schaik, 1 987, p. 585; Ruiter etal., 1994, p. 218). In the Ketambe study area, northern Sumatra, a natural nonprovisioned population of M. fas- cicularis increased from 70-75 in 1972 to more than 190 in 1986 (van Schaik & van Noordwijk, 1988, p. 94), an annual rate of increase of about 7%; at this rate, which obviously is not indefinitely sustainable, the population would double every 10.5 years. The introduced nonprovisioned pop- ulation in Angaur increased in less than 75 years from a few individuals to 480-600 in 1 973 (Poirier & Smith, 1974, pp. 264, 271; cf. Kawamoto et al., 1988, p. 176; Matsubayashi et al., 1989, p. 54). In Mauritius, another introduced nonprovisioned population apparently increased in less than 500 years from a few individuals to about 30,000 (Sussman & Tattersall, 1986, pp. 30, 38; Lawler et al., 1995, p. 138). In Florida, a seminatural menagerie group increased from 6 individuals (4 females, 2 males) in 1933 to about 150 individuals in 1966 (Dumond, 1967, p. 203), an annual rate of increase of about 1 0%. Fossils and Subfossils Known fossils indicate that M. fascicularis has inhabited the Sunda Shelf since at least later Early Pleistocene, approximately 1 Ma (Table 33). All of the earliest macaque fossils in this area, dating from 0.8 to 1.0 Ma, have been collected in Java, where, at this time, M. fascicularis apparently was sympatric with M. nemestrina. In Java, M. fas- cicularis fossils of this age outnumber those of M. nemestrina, but most of these fossils consist of isolated teeth or small jaw fragments, and some may be misidentified. The long period from 800 Ka to 35 Ka has yielded no macaque fossils within the present geo- graphic range of M. fascicularis. However, fossils and subfossils of M. fascicularis and M. nemes- trina have been collected in subsequent Late Pleis- tocene and Holocene deposits in West Malaysia, Sumatra, Borneo, and Java (M. fascicularis fossils only). In these deposits in Borneo and Java, M. fascicularis fossils and subfossils outnumber those of M. nemestrina. In deposits in West Malaysia and Sumatra, by contrast, M. nemestrina fossils and subfossils outnumber those of M. fascicularis. This is anomalous, because, at present, M. fasci- cularis generally outnumbers M. nemestrina in the broad area of overlap of their respective ranges, except in the interior of primary evergreen rain forest (Fooden, 1975, p. 61; above, Natural His- tory). The explanation for the anomalous relative frequencies of fossils and subfossils of these species in Sumatra and West Malaysia is unclear. In the Lesser Sunda Islands, Holocene subfossils of M. fascicularis, consisting of human food re- 7 Age estimates from Theunissen et al. (1990, p. 51; cf. Swisher et al., 1994, p. 1 1 19; J. de Vos & Sondaar, 1994, p. 1726; Swisher, 1994, p. 1727). 8 Cf. Koenigswald (1951, p. 219) and Medway (1972c, p. 80). 9SeeMusser(1981,p. 72). 10 Age estimate indicates age of earliest fossil collected at each locality; number of fossils indicates minimum number of individuals collected at each locality. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 63 Fig. 25. Type localities and limits of distribution of recognized subspecies of Macaca fascicularis and type localities of synonymous nominal species or subspecies (cf. Fig. 1; Fooden, 1982, p. 576); in the Philippine Islands, the indicated intersubspecific contact zone includes an area for which the subspecies is undetermined (cf. Fooden, 1991, p. 25). Because the geographic origins of the holotypes of Macacus cristatus Gray, 1870, and Macacus cynomolgus . . . var. Cumingii Gray, 1870, are inadequately known, these two synonyms of M. fascicularis are excluded from this figure (see Fooden, 1991, pp. 23, 24). Key to type localities of synonyms of recognized subspecies: M. f. aurea 1 . Pithecus vitiis Elliot, 1910; Burma, Letsok-aw Kyun. M. f. fascicularis 2. Pithecus validus Elliot, 1909; Vietnam: "Cochin China." 3. Pithecus capitalis Elliot, 1910; Thailand, Trang Province: Ban Phra Muang. 4. Macaca irus argentimembris Kloss, 1911b; Malay- sia, West Malaysia, P. Pinang [2]. 5. Pithecus agnatus Elliot, 1910; Indonesia, P. Tuang- ku. 6. Macacus phaeur a Miller, 1 903b; Indonesia, P. Nias: Teluk Siaba. 7. Pithecus mansalaris Lyon, 1916; Indonesia, P. Mur- sala. 8. Macacus irus I. Geoffroy, 1 826; Indonesia, Sumatra. 9. Macacus carbonarius I. Geoffroy, 1826; Indonesia, Sumatra (technical name also mistakenly applied to M. f. aurea and M. f. umbrosa). 10. Pithecus alacer Elliot, 1909; Indonesia, P. Kundur: Selatbliat. 11. Pithecus karimoni Elliot, 1909; Indonesia, P. Kar- imun: Monos. 12. Pithecus dollmani Elliot, 1909; Singapore, Singa- pore I.: Changi. 13. Pithecus laetus Elliot. 1909: Malaysia, West Malay- sia, P. Tinggi. 14. Pithecus bintangensis Elliot, 1909; Indonesia, P. Bintan: Sungei Biru. 15. Pithecus impudens Elliot, 1910; Indonesia, P. Sugi. 16. Pithecus lingae Elliot, 1910; Indonesia, P. Lingga. 17. Pithecus lapsus Elliot, 1910; Indonesia, P. Bangka: Tanjung Pamuja. 18. Pithecus lautensis Elliot, 1910; Indonesia, P. Laut. 1 9. Pithecus lingungensis Elliot, 1910; Indonesia, P. La- gong. 20. Pithecus sirhassensis Elliot, 1910; Indonesia, P. Ser- asan. 2 1 . Macacus pumilus Miller, 1 900; Indonesia, P. Benua. 22. Pithecus mandibularis Elliot, 1910; Indonesia, Bor- neo, Kalimantan: Sungai Ambawang, near Pontia- nak. 23. Pithecus carimatae Elliot, 1910; Indonesia, P. Kar- imata: Teluk Pai. 64 FIELDIANA: ZOOLOGY mains, have been collected in caves in Timor and Flores, both of which are east of Wallace's Line (passing between Bali and Lombok). In four care- fully excavated caves in Timor, where basal de- posits date back to ca. 13.5 Ka, the oldest M. fascicularis subfossils do not appear until ca. 4.5 Ka (Glover, 1986, p. 212). At lower levels, the only mammals included among the human food remains are murid rodents and bats, of which thousands of skeletal fragments were collected; these lower levels also yield stone tools. At ap- proximately the same level where M. fascicularis first appears, remains of palm civet and cuscus also appear, along with remains of domestic goat, pig, and dog and pottery. Glover (1986, p. 159) concluded that "... there is strong presumptive evidence that these species [M. fascicularis, palm civet, and cuscus] were introduced into Timor di- rectly by, or through the agency of man, between about 4000-5000 years ago." For M. fascicularis, a less plausible alternative hypothesis is that it was present in Timor earlier but was not hunted by the preceramic cave inhabitants, possibly because they lacked suitable projectile weapons (cf. Adi, 1985, p. 65). In Flores, a subfossil molar of M. fascicularis was collected in cave deposits, dated ca. 3.5 Ka, that also contain human artifacts and human skel- etal remains. Other mammals represented in these deposits are rat species, fruit bat, porcupine, and pig (Hooijer, 1967, p. 160). Based on available zoogeographic evidence, including Glover's ar- cheological data from Timor (see above), Musser ( 1 98 1 , p. 133) concluded that M. fascicularis, por- cupine, and pig probably were introduced into Flo- res by humans. Systematics Subspecific Taxonomy The geographic range of M. fascicularis is broad and encompasses mainland Southeast Asia and numerous large and small islands, both on and beyond the Sunda Shelf (Figs. 3, 4). It is therefore not surprising that this species exhibits great in- dividual, local, and geographic variation in a di- verse array of characters (see above); variation among populations of M. fascicularis ranges from trivial, to statistically significant, to locally dis- continuous, to totally discontinuous. Based on this variation, 50 specific or subspecific names have been proposed for populations of this species (Fig. 25). The evaluation of intraspecific variation for the purpose of defining taxonomically useful sub- species presents serious theoretical and practical difficulties, as previously discussed by Chasen (1940a, p. 66; cf. Table 35) in his classic review of M. fascicularis. In the present taxonomic analysis, subspecific status is accorded only to those geographic pop- ulations in which variation of at least one character is completely— or nearly completely— discontin- uous with variation of the same character in the nominotypical subspecies, M. f fascicularis. The reason for selecting character-state discontinuity as the critical requisite for subspecific recognition is that, without such discontinuity, unambiguous diagnosis of subspecies becomes difficult or im- possible. The requirement of character-state dis- continuity denies subspecific recognition to local populations that are distinct from adjacent pop- ulations of M. f fascicularis but that fall within 24. Macaca resima Thomas & Wroughton, 1909b; In- donesia, Java: Tasikmalaya. 25. Macaca modax Thomas & Wroughton, 1909c; In- donesia, Java: Cilacap. 26. [Simia] Aygula Linnaeus, 1758; Indonesia, Java. 27. Pithecus baweanus Elliot, 1910; Indonesia, P. Baw- ean. 28. Pithecus cupidus Elliot, 1910; Indonesia, P. Mata- siri. 29. Macaca irus submordax Sody, 1949; Indonesia, P. Bali: Desa Poetjang. 30. Macaca irus sublimitus Sody, 1932; Indonesia, P. Sumba: Payeti-Kambaniru and Mao Marroe. 31. Pithecus fascicularis limitis Schwartz, 1913; Indo- nesia, P. Timor: Lelogama. 32. Cynomolgus cagayanus Mearns, 1905; Philippines, Cagayan Sulu I. 33. Cynomolgus suluensis Meanrs, 1905; Philippines, Jolo I.: foot of Crater Lake Mountain. M. f. fascicularis/ M. f. philippinensis contact zone 34. Cynomolgus mindanensis apoensis Mearns, 1905; Philippines, Mindanao I.: Mt. Apo. 35. Cynomolgus mindanensis Mearns, 1905; Philip- pines, Mindanao I.: Pantar. M. f. philippinensis 36. Pithecus mindorus Hollister, 1913; Philippines, Mindoro I.: Alag River. 37. Macacus palpebrosus I. Geoffroy, 1851; Philippines, Luzon I.; "forets de Manille." 38. Macacus fur Slack, 1867; Philippines, Luzon I. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 65 Table 34. Summary comparison of external characters in recognized subspecies of Macaca fascicularis. x Dorsal Lateral Head and Relative pelage Crown facial crest Thigh (outer body length,4 tail length,4 Subspecies2 color color pattern3 surface), color ad. males ad. males aurea Buffy to me- Golden Infrazyg. Similar to dors. 560 ± 61 95 ± 9 dium brown pel. col. 520-630 85-104 brown N = 3 atriceps Buffy to me- Dark brown Transzyg. Similar to dors. 452 ± 17 118 ± 7 dium to blackish pel. col. 425-465 110-128 brown (narrow)5 N = 5 condorensis Buffy to me- Dark brown Transzyg. Similar to dors. 450 ± 18 115 ± 7 dium to blackish pel. col. 435^180 109-127 brown (broad)5 N = 5 fascicularis Buffy to dark Golden Transzyg. Similar to dors. 462 ± 43 118 ± 15 brown brown pel. col. 370-610 N= 193 69-150 N = 189 karimondjawae Dark grayish Dark brown 7 Similar to dors. 501 108 brown to diffusely blackish pel. col. N = 1 philippinensis Dark brown Dark golden Transzyg. Similar to dors. 480 ± 33 114 ± 9 brown pel. col. 410-530 N = 101-129 13 tua Blackish Yellowish brown Transzyg.6 Brownish gray 440 N = 131 1 umbrosa Blackish Yellowish Infrazyg. or Pale brownish gray 502 ± 25 115 ± 1 brown transzyg.7 475-525 N = 470 115-116 3 118 lasiae Blackish Blackish8 Infrazyg. or Similar to dors. transzyg. pel. col.9 N = 1 fusca Blackish Blackish10 Infrazyg. or Similar to dors. 478 ± 14 97 ± 6 transzyg. pel. col." 460-495 N = 90-105 6 1 Abbreviations: ad. = adult; infrazyg. = infrazygomatic; transzyg. = transzygomatic. 2 Listed in order of increasing pelage color saturation. 3 See Figure 8. 4 Entries indicate mean ± SD, extremes, and sample size; relative tail length = tail length/head and body length 100. 5 See Figure 7. 6 Subauricular hairs pale ochraceous-buff, conspicuously elongated. 7 Subauricular hairs pale brownish, not elongated. 8 Crown hairs inconspicuously annulated. 9 Outer surface of shanks pale grayish brown. 10 Crown hairs conspicuously annulated with pale yellowish. 1 ' Outer surface of shanks blackish. the range of variation of more distant populations of this subspecies. This is not intended to depre- ciate the theoretical importance of locally deviant populations, which often have great zoogeographic or evolutionary significance (see below); appreci- ation of this significance does not require formal subspecific recognition. A total of 1 0 subspecies of M. fascicularis are recognized here (Fig. 25, Table 34, Appendix 12). These are M.f. fascicularis (core area, many shal- low-water fringing islands, some deep-water fring- ing islands); M. f aurea (core area northwest of M.f fascicularis, adjacent shallow-water fringing islands); M. f atriceps, M. f condorensis, and M. karimondjawae (shallow-water fringing islands); and M. f umbrosa, M. fascicularis fusca, M. f lasiae, M. f tua, and M. f philippinensis (deep- water fringing islands). Characters and distribu- tions of these subspecies are briefly reviewed be- low and are treated in greater detail in subsequent subspecies accounts. 1 . M.f. fascicularis. The type locality of M. fas- cicularis—and, hence, of M. f fascicularis— is Bengkulu, southwestern Sumatra, a locality that fortuitously lies within the previously defined core area of distribution of the species. Judging from 66 FIELDIANA: ZOOLOGY available evidence, variation of characters in Su- matra is continuous with that in the Malay Pen- insula (including the Isthmus of Kra north to ca. 1 0°N), Borneo, and Java (see above); all of these parts of the core area are therefore included within the geographic range of M. f. fascicularis, as here defined. In the southern part of the Indochinese Peninsula, character-state variation in M. fasci- cularis also is continuous with that in the Malay Peninsula, Sumatra, Borneo, and Java, with the possible exception of variation of mtDNA types (Table 17) and resistance to experimental infec- tions with Plasmodium knowlesi malaria (see above; Fooden, 1994, p. 585). Because evidence for discontinuity of variation of mtDNA types and P. knowlesi resistance is incomplete, the southern part of the Indochinese Peninsula is provisionally retained within the geographic range of M. f. fas- cicularis. Although now disjunct, the range of M. f fascicularis in the southern part of the Indo- Chinese Peninsula was connected by dry land to the range of this subspecies in the Malay Peninsula during the last glacial maximum, ca. 1 8 Ka (Fig. 3). In most shallow-water fringing-island popula- tions of M. fascicularis, variation is continuous with that in core-area M. f fascicularis (Appen- dixes 3-9; Fooden & Albrecht, 1993, p. 537); therefore, these islands are included within the geographic range of M. f fascicularis. In eight of these shallow-water fringing-islands, populations are distinct from nearby core-area populations of M.f fascicularis but fall within the range of vari- ation of more distant populations of this subspe- cies; as indicated above, these populations are not recognized here as separate subspecies, although some have been recognized as subspecies by other authors. These eight populations, sorted according to characters that distinguish the fringing-island populations from nearby core-area populations of M.f. fascicularis, inhabit the following islands: P. Mursala (west of Sumatra), P. Redang (east of the Malay Peninsula), P. Lagong (west of Borneo), and P. Uwi (west of Borneo)— distinguished by dorsal pelage erythrism (Appendix 4); P. Tioman (east of the Malay Peninsula) and P. Belitung (west of Bor- neo)—distinguished by T and/or relative T (Ap- pendixes 8, 9); and Ko Kut and Phu Quoc Dao (both south of the Indochinese Peninsula)— dis- tinguished by skull length (Fooden & Albrecht, 1993, p. 528). In many other fringing-island pop- ulations, character-state variation is statistically significantly different from, but continuous with, variation in nearby core-area populations (Ap- pendixes 3-9); some of these populations also have been recognized as separate subspecies by other authors. The deep-water fringing-island population of M. fascicularis in P. Nias (west of Sumatra) resembles the eight shallow-water fringing-island popula- tions listed above in being distinct— in tail col- oration and T— from nearby core-area popula- tions of M. f fascicularis in Sumatra but falling within the range of variation of more distant pop- ulations of this subspecies. Although the proxi- modorsal surface of the tail is more intensely blackish in P. Nias than in Sumatra, as empha- sized by Miller (1903b, p. 63) in his description of the P. Nias monkey as Macacus phaeura, tail coloration in M. fascicularis in P. Nias broadly overlaps that in specimens collected in other parts of the species range (cf. mcz 36030, usnm 121871 — P. Nias; usnm 196823— Kalimantan; mcz 37415 — Sabah; usnm 124710-P. Bangka; usnm 124969, 124970-P. Belitung). T and relative T in P. Nias similarly overlap T and relative T in other parts of the species range (Figs. 15, 17; Appendixes 8, 9). Lacking character-state discontinuity, M. fas- cicularis in P. Nias is regarded here as subspecif- ically inseparable from M. f fascicularis. Deep- water fringing-island populations of M. fascicu- laris in the Lesser Sundas and southern Philippines are retained within M.f. fascicularis because known variation in these populations is continuous with that in nearby core-area populations of M. f fas- cicularis (Appendixes 3-5, 7-9; Fooden, 1991, p. 21; Fooden & Albrecht, 1993, p. 537). 2. M. f aurea. In the western part of the In- dochinese Peninsula and western part of the Isth- mus of Kra south to about 1 0°N, variation of the lateral facial crest pattern (infrazygomatic) is al- most completely discontinuous with variation of this character (transzygomatic) in the rest of the core area (Figs. 8, 9). This warrants recognition of the subspecies M. f aurea (type locality, Pegu, southern Burma). In all samples available from shallow-water islands in the Mergui Archipelago, west of the northern part of the Isthmus of Kra, the lateral facial crest pattern also is infrazygo- matic; these islands are therefore included within the geographic range of M. f aurea. Two areas immediately east of the range of M. f aurea are heterogeneous for the lateral facial crest pattern (Fig. 9). The more southern of these two areas, in eastern and southern parts of the Isthmus of Kra and adjacent southwestern islands, may be regarded as an intersubspecific contact zone between the ranges of M. f fascicularis and M. f FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 67 Table 35. Subspecies of Macaca fascicularis recognized as valid by Chasen (1940a) and recent authors; dash (— ) indicates that subspecies is extralimital in the cited work. Author and date w. c. o. P. H. Groves and Chasen Hill Napier Scott Weitzel Fa This Subspecies (1940a) (1974) (1981) (1982) (1988)» 1989 paper argentimembris X X X X X atriceps X2 X X X X X aurea X3 X X X X baweana X X X X — X bintangensis X X cagayana — X — — capitalis X X X X condorensis X2 X X X3 X cupida X X X — — X fascicularis (= irus) X X X X X X X fusca X X X — — X X impudens X — karimondjawae X X X — X laeta X X X X X X lasiae X — — X X limitis — X X X — X lingungensis X X mandibularis X X mindanensis — — — X mindora — — — X mordax X X X X X X phaeura X X X — X3 X philippinensis — X X — — X X pumila X X X X — X sirhassensis X X sublimitis — X X — X submordax X — tua — — X X umbrosa X2 X X — — X X valida — X X X3 X Subsp. (Borneo, north) X X Subsp. (P. Banggi) X Subsp. (P. Siantan) X In Weitzel et al. (1988, pp. 5, 96). Subspecies recognized, name not applied. Subspecies recognized provisionally. aurea. The more northern of these two areas, in central and eastern parts of the Indochinese Pen- insula, along the northern margin of the species range, may be either an intersubspecific contact zone between the ranges of M. f. fascicularis and M. f. aurea or an interspecific contact zone be- tween the ranges of M. fascicularis and M. mulatta (cf. Fooden, 1971, p. 24); future study of variation in M. mulatta may contribute to resolution of this ambiguity. 3. M. f. atriceps. The distinctive narrow dark crown patch warrants recognition of this subspe- cies in shallow-water Ko Khram Yai in the Gulf of Thailand (Fig. 7, Table 5, Appendix 5). 4. M.f. condorensis. The distinctive broad dark crown patch warrants recognition of this subspe- cies in shallow-water Con Son and nearby Hon Ba, both in the South China Sea (Fig. 7, Table 5, Appendix 5). 5. M. f karimondjawae. Variation of dorsal pelage color saturation in the population of M. fascicularis in P. Karimunjawa and presumably also in nearby P. Kemujan, both shallow- water islands north of Java, overlaps minimally with that in M. f fascicularis (Appendix 3). The dark grayish brown dorsal pelage color of this popula- tion is the basis for recognition of M. f. kari- mondjawae. 6. M.f. umbrosa. The population of M. fasci- cularis in the Nicobar Islands, northwest of Su- 68 FTELDIANA: ZOOLOGY matra, is one of four deep-water fringing-island populations with distinctively blackish dorsal pel- age (Appendix 3). The crown is yellowish brown and the thighs are pale brownish gray in M. f. umbrosa. 7. M. fascicularis fusca. In the population of M. fascicularis in deep-water P. Simeulue, west of Su- matra, the back, crown, and thighs are blackish, and the tail is shorter than usual in M. fascicularis (Figs. 16, 18; Appendixes 3, 8, 9). 8. M.f lasiae. In the population in deep-water P. Lasia, near P. Simeulue, the back, crown, and thighs are blackish, as in M. fascicularis fusca, but the tail is of normal length (Figs. 16, 18, 27; Ap- pendixes 3, 8, 9). 9. M.f. tua. In the population in deep-water P. Maratua, east of Borneo, the back is blackish, the crown is yellowish brown, and the thighs are brownish gray (Appendix 3). In M. f tua, unlike M. f. umbrosa, subauricular hairs are elongated and pale ochraceous-buff. 10. M. f philippinensis. In populations of M. fascicularis in western, northern, and eastern is- lands of the Philippine Archipelago, dorsal pelage color is distinctively dark brown (Appendix 3; Fooden, 1991, p. 3). In eastern and central Min- danao, southern Negros, and perhaps in some nearby islands, mixed populations of dark and pale individuals occur; this area of mixed populations is regarded as a contact zone between M. f phi- lippinensis and M. f fascicularis. Brief casual remarks have been published con- cerning dorsal pelage coloration in M. fascicularis in P. We, a deep-water island off the northern tip of Sumatra (Scheffrahn et al., 1994, p. 136). These remarks are inadequate for determination of the subspecific status of this population. The present subspecific classification of M. fas- cicularis differs from recent classifications pro- posed by W. C. O. Hill (1 974, p. 504), R. H. Napier (1981, p. 12), Scott (1982, unpubl. M.A. thesis, p. 182), Weitzel et al. (1988, p. 96), and Fa (1989, p. 54) (Table 35). P. H. Napier, whose list of sub- species follows that of W. C. O. Hill with a single exception (nonrecognition of subspecies cagayana Mearns, 1905), explicitly acknowledges that her classification is unsatisfactory and that many of the recognized subspecies cannot be diagnosed. Scott's classification is based on a sample of 148 specimens (including 1 34 skins and 69 skulls), more than 90% of which are in two museums (mzb, zrc); this sample, which apparently is almost identical with that studied by Chasen (1940a, p. xix), pro- vides an inadequate representation of variation in M. fascicularis. Weitzel et al. indicate that their classification is largely based on Scott's research. Fa, without further explanation or documentation, lists 2 1 subspecies of M. fascicularis that he rec- ognizes as valid; most, but not all, of these sub- species are the same as those recognized by authors cited above. Key to Recognized Subspecies The 10 subspecies of Macaca fascicularis recognized here are distinguished in the following key, which is based on external characters of postinfants: 1 . General color of dorsal surface of trunk buffy to yellowish gray to medium brown to dark brown, variably erythristic 2 General color of dorsal surface of trunk blackish 7 2. Preauricular hairs directed posteriorly, partly covering ears (lateral facial crest infrazygomatic) .... aurea Preauricular hairs directed anteriorly, forming part of lateral facial crest, not covering ears (lateral facial crest transzygomatic) 3 3. General color of dorsal surface of trunk buffy to yellowish gray to medium brown, variably erythristic 4 General color of dorsal surface of trunk dark brown,1 variably erythristic 6 4. Crown conspicuously darker than back 5 Crown not conspicuously darker than back fascicularis 1 Note: Approximately 5% of M.f. fascicularis specimens examined are dark brown and, if of unknown geographic origin, probably would be misidentified as either M. f. karimondjawae or M. f. philippinensis. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 69 5. Dark crown patch narrow, extending laterally approximately as far as middle of each eye, sharply delimited laterally and posteriorly atriceps Dark crown patch broad, extending laterally to or beyond lateral margin of each eye, not sharply delimited laterally and posteriorly condorensis 6. Pale annulations of dorsal hairs pale yellowish karimondjawae Pale annulations of dorsal hairs golden to rufescent philippinensis 1 . Outer surface of thighs blackish, approximately same color as dorsal surface of trunk 8 Outer surface of thighs brownish gray, contrastingly paler than dorsal surface of trunk 9 8. Outer surface of shanks blackish; tail length < 1 12% of head and body length fusca Outer surface of shanks pale grayish brown; tail length > 1 1 2% of head and body length . . . lasiae 9. Subauricular hairs pale ochraceous-buff, elongated (4-6 cm), extending to apex of lateral facial crest forming a conspicuous pale lateral facial patch; tail length > 1 22% of head and body length . . . tua Subauricular hairs brownish gray to pale gray, not elongated (2-4 cm), not extending to apex of lateral facial crest, relatively inconspicuous; tail length < 122% of head and body length umbrosa Subspecies Accounts Macaca fascicularis fascicularis (Raffles, [1821], p. 246) Jawanska Markattor: Osbeck, 1757, p. 99— external characters and behavior of monkeys collected and ob- served in Java by P. Osbeck, Jul. 1751; specimens not preserved. [Simia] Aygula Linnaeus, 1758, p. 27 — based on Ja- wanska Markattor: Osbeck, 1757, and supplementary information in unpublished letters from Osbeck to Linnaeus (P. H. Napier & Groves, 1 983, p. 1 1 7). Tho- mas & Wroughton, 1909a, p. 373 — misidentified as a species of leaf monkey. P. H. Napier & Groves, 1983, p. 118— shown to be a senior synonym of Macaca fascicularis (Raffles, [1821]); suppression requested. International Commission on Zoological Nomencla- ture, 1986, p. 229— specific name officially suppressed. [Cercopithecus] aygula: Erxleben, 1777, p. 39— external and cranial characters. Cercop[ithecus] Aigula: Schinz, 1821, p. 108— incorrect spelling of [Simia] Aygula Linnaeus, 1758. Schinz, 1825, p. 256 — said to be indistinguishable from Inuus cynamolgus [— Macaca fascicularis]. Macaca aygula: Daudin in Lacepede & Daudin, [1802], p. 148— listed as "Variete A" of Macaca cynomolgus [= M. fascicularis]. Pithecus aygula: E. Geoffroy, 1 803, p. 24— external char- acters. Cercocebus aygula: E. Geoffroy, 1812, p. 99— external characters. Macaque: Buffon in Buffon & Daubenton, 1766, p. 190, pis. 20, 22-24— based on specimen of unknown origin, not preserved; species said to inhabit "Congo & des autres parties de l'Afrique meridionale." F. Cuvier, 1819, liv. 3, p. 1, 2 pis. — illustrations of captives des- ignated as Irus by F. Cuvier, 1818; habitat, "vra- isemblablement du Senegal ou de la cote de Guinee." Aigrette: Buffon in Buffon & Daubenton, 1766, p. 190, pi. 21— based on specimen of unknown origin, not preserved; regarded as "variete" of Macaque: Buffon in Buffon & Daubenton, 1766. Simia Cynamolgos: Schreber, [1774], p. 91, pi. 13 (part; not Linnaeus, 1758, p. 28)— misidentification of Ma- caque: Buffon in Buffon & Daubenton, 1 766. Schlegel, 1876, p. 101 —name said to have been based on mis- identification. Blanford, [1888a], p. 624— name said to have been based on misidentification. Cercopithecus (Cynamolgus): Zimmermann, 1780, p. 186 (part; not Linnaeus, 1758, p. 28)— misidentification of Macaque: Buffon in Buffon & Daubenton, 1766. Cercocebus cynamolgos: Schlegel, 1876, p. 101 (not Lin- naeus, 1758)— taxonomic comparisons. [Cercopithecus] Cynomolgus: Erxleben, 1 777, p. 28 (part; not Linnaeus, 1766, p. 38)— misidentification of Ma- caque: Buffon in Buffon & Daubenton, 1766. Simia Cynomolgus: Audebert, 1798-1799, p. 5 (part; not Linnaeus, 1766, p. 38)— misidentification of Ma- caque: Buffon in Buffon & Daubenton, 1766. Simia cynomolgos: G. Cuvier, 1798, p. 98— incorrect spelling of [Simia] Cynomolgus: Linnaeus, 1766; mis- identification of Macaque: Buffon in Buffon & Dau- benton, 1766. Macaca cynomolgus: Daudin in Lacepede & Daudin, [1802], p. 148 (not Linnaeus, 1766)— misidentifica- tion of Macaque: Buffon in Buffon & Daubenton, 1766. [M]acacus cynomolgus: Desmarest, 1820, p. 65 (part; not Linnaeus, 1766)— misidentification of Macaque: Buffon in Buffon & Daubenton, 1766. Blanford, 1888b, pp. 21, 23— binomial said to be based on misidenti- fication, accepted as valid. Macacus ' cynomolgus' Auct.: Miller, 1900, p. 239 — specimens collected in P. Lingga and P. Tioman. Mil- ler, 1903a, p. 476 — specimens collected at Teluk Ta- panuli, Sumatra, and in P. Tuangku and P. Mursala. Cyn[ocephalus] cynomolgus: Latreille, 1 804, p. 292 (part; not Linnaeus, 1766, p. 38)— misidentification of Ma- caque: Buffon in Buffon & Daubenton, 1766. Cercocebus cynomolgus: E. Geoffroy, 1812, p. 99 (not Linnaeus, 1766)— external and cranial characters. [P]ithecus cynomolgus: Desmarest, 1817, p. 323 (part; not Linnaeus, 1766, p. 38)— misidentification of Ma- caque: Buffon in Buffon & Daubenton, 1766. Pithecus Cynomolgos: Blainville, [1839], pi. 7 — incorrect spelling of [Simia] Cynomolgus: Linnaeus, 1 766. Pith[ecus] (Mac[acus]) cynomolgus: Dahlbom, 1856, p. 1 18 (not Linnaeus, 1766, p. 38)— external characters. 70 FIELDIANA: ZOOLOGY I[nuus] cynomolgus: Wagner, [1839], p. 135 (not Lin- naeus, 1766, p. 38)— external characters; distribution; taxonomic comparisons. Irus: F. Cuvier, 1818, p. 120— unavailable name, not published in combination with generic name; pro- posed as substitute specific technical name for Ma- caque: Buffon in Buffon & Daubenton, 1766 (cf. I. Geoffrey, 1826, p. 588; [1831], p. 56: Gervais, 1854, p. 85; Miller, 1942, p. 127); habitat "vraisemblable- ment du Senegal ou de la cote de Guinee" (corrected to "du midi de l'Asie, et particulierement de Sumatra" by F. Cuvier, 1825, liv. 52, p. 1). Simia fasciculcris Raffles, [1821], p. 246— for details concerning holotype, see below. Cantor, 1846, p. 176 — cited as a synonym of Cercopithecus cynomolgus. H. C. Robinson, 1916, p. 63— type locality restriction. Semn[nopithecus]l fascicularis: [Vigors], 1830, p. 642— geographic distribution; type history; "doubtful whether it is a true Semnopithecus." Semnopithecus fascicularis?: Waterhouse, 1838, p. 4— geographic distribution. S[emnopithecus] fascicularis: Martin, 1838, p. 435— ex- ternal characters. Macacus fascicularis: Bonhote, 1903, p. 3— taxonomic history. Macaca fascicularis: Miller, 1906b, p. 65— specimens collected in P. Karimata. Wroughton, 1918, p. 556— taxonomic history. Miller, 1942, p. 126— taxonomic history. Hooijer, 1962b, p. 44— subspecific taxonomy. Macaca fascicularis group: Thomas, 1928, p. 832— spec- imens collected in Vietnam. [Cynomolgus] fascicularis: Trouessart, 1904, p. 16— geo- graphic distribution. Pith ecus fascicularis: Elliot, 1913, p. 233— taxonomic comparison. [Silenus] fascicularis: Stiles & Nolan, 1929, p. 529— parasites. P[ithecus] f[ascicularis] fascicularis: Schwarz, 1913, p. 297— taxonomic comparison. M[acaca] fascicularis fascicularis J. R. Napier & Napier, 1967, p. 403— geographic distribution. Macaca mulatto fascicularis: Fooden, 1964, p. 364— taxonomic comparison. Kra Buku: Raffles, [1821], p. 247— vernacular name of "smaller species" than Simia fascicularis Raffles, [1821], p. 246 [apparently juvenile Macaca fascicu- laris], specimens apparently not obtained; habitat, Su- matra and "other Malay islands." Macacus irus I. Geoffroy, 1826, p. 588— based on Ma- caque: Buffon in Buffon & Daubenton, 1 766, and liv- ing captives reported by F. Cuvier (1818, p. 112; 1819, liv. 3, p. 1); specimens not preserved, origin unknown; binomial incorrectly attributed to F. Cuvier, 1818. I. Geoffroy, [1831], p. 56— cited as a synonym of Ma- cacus cynomolgus Desmarest, 1820; taxonomic his- tory. Blanford, [1888a], p. 624— taxonomic history. Cabrera, 1910, p. 620— taxonomic history; type lo- cality, Sumatra. H. C. Robinson & Kloss, 1914, p. 394— type locality, probably Malacca (= Melaka). Miller, 1942, p. 127— cited as a synonym of Macaca fascicularis (Raffles, [1821]); binomial attributed to Blanford, [1888a]. Fooden, 1976, p. 226-cited as a synonym of Macaca fascicularis (Raffles, [1821]); bi- nomial attributed to I. Geoffroy, 1826. Macaca irus: H. C. Robinson & Kloss, 1915a, p. 130 — specimens collected in Ko Samui and Ko Phangan. [Macaca] zVus-group: Miller, 1933, pp. 5, 6— taxonomic history. [Macaca] ira: Weinman & Wiratmadja, 1969, p. 498 — trypanosomes. Pithecus irus: Elliot, 1913, p. 229— taxonomic compar- ison. [Silenus] irus: Stiles & Nolan, 1929, p. 530— parasites. Cynomolgus irus: Furuya, 1962, p. 377— dermatoglyph- ics. Macaca irus irus: H. C. Robinson & Kloss, 1918, p. 6— collected in Sumatra; taxonomic history. Chasen, 1940a, p. 66— taxonomic comparisons. Kellogg, 1945, pp. 116, 129— taxonomic comparisons. Sody, 1949, p. 130— taxonomic comparison. Carbonarius: F. Cuvier, 1825, liv. 52, p. 2, pi.— un- available name, not published in combination with generic name; based on living captive, origin un- known; specimen not preserved. Macacus carbonarius I. Geoffroy, 1826, p. 588— based on Carbonarius: F. Cuvier, 1825; external characters; taxonomic comparison. Lesson, 1827, p. 42— said to inhabit Sumatra. I. Geoffroy, [1831], p. 63— doubt- fully distinct. Eydoux & Souleyet [& Gervais], 1841, p. 6— cited as a synonym of Macacus aureus I. Geof- froy, [1831] [= Macaca fascicularis aurea]. Blyth, 1875, p. 7— cited as a synonym of Macacus cynomolgus [= Macaca fascicularis]. S[imia] carbonaria: Fischer, 1829, p. 26— habitat, Su- matra. C[ynamolgus] carbonarius: Reichenbach, 1862, p. 136 — taxonomic comparison. Semnopithecus kra Lesson, [1830], p. 20— replacement name for Simia fascicularis Raffles, [1821], p. 246. Semnopithecus Buku Martin, 1 838, p. 435 —specific name proposed provisionally for Kra Buku: Raffles, [1821], p. 247. J. Anderson, 1879, p. 74— cited as a synonym of Macacus cynomolgus [= Macaca fascicularis]. Inuus cercopithecus Mason, 1851, p. 220— unavailable name attributed to Blyth, cited in synonymy, never treated as available. Macacus cancrivorus Mason, 1 85 1 , p. 22 1 —unavailable name attributed to Blyth, cited in synonymy, never treated as available. C[ynamolgus] cynocephalus: Reichenbach, 1862, p. 133 (not Linnaeus, 1766, p. 38)— natural history. Macacus assamensis: Gray, 1 870, p. 3 1 (not McClelland in Horsfield, [1840], p. 148)— misidentification; tax- onomic comparison. Macacus Sinicus: Morice, 1875, p. 41 (not Linnaeus, 1 77 1 , p. 52 1)— misidentification of monkeys observed in Vietnam. Macacus pumilus Miller, 1900, p. 241 —holotype, usnm 101639, adult male, skin and skull, collected at Pulo Bunoa (= Pulau Benua), Kepulauan Tambelan, In- donesia, by W. L. Abbott, 6 Aug. 1899; paratypes, usnm 101638 (juvenile female, P. Benua, 5 Aug.), usnm 101666 (adult female, P. Uwi, 13 Aug.), and usnm 101711 (subadult male, P. Siantan, 8 Sep.), skins and skulls, collected in Indonesia by W. L. Abbott, 1899. Lyon & Osgood, 1909, p. 284— holotype cataloged. Weitzel et al., 1988, p. 112— topoparatypic specimens (P. Siantan) referred to M. fascicularis sirhassensis (El- liot, 1910). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 71 [Cynomolgus] pumilus: Trouessart, 1904, p. 16— distri- bution. Pithecus pumilus: Elliot, 1 9 1 3, p. 252— external and cra- nial characters. Macaca irus pumilus: Chasen & Kloss, 1928b, p. 29 — taxonomic comparison. Macaca irus pumila: Chasen, 1940a, p. 69— geographic distribution. Poole & Schantz, 1942, p. 245 — holotype cataloged. Weitzel et al., 1988, pp. Ill, 1 13— cited as name previously applied to specimens referred to M. fascicularis lingungensis (Elliot, 1910) and M. f. sir- hassensis (Elliot, 1910). M[acaca] f[ascicularis] pumila: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Macacus phaeura Miller, 1903a, p. 63 — holotype, usnm 121870 (Coll. No. 2399), adult male, skin and skull, collected at Teluk Siaba, P. Nias, Indonesia, by W. L. Abbott, 20 Mar. 1903; paratypes, usnm 121868 (Coll. No. 2393, adult male, 16 Mar.), usnm 121869 (Coll. No. 2395, adult female, 18 Mar.), and usnm 121871 (Coll. No. 2400, adult male, 20 Mar.), skins and skulls, collected at Teluk Siaba, P. Nias, Indonesia, by W. L. Abbott, 1903. Lyon & Osgood, 1909, p. 284-holo- type cataloged. Macaca phaeura: Lyon, 1906, p. 606— specimens col- lected in P. Bangka and P. Belitung. Miller, 1942, p. 128 — specimens collected in P. Nias. Pithecus phaeurus: Elliot, 1913, p. 243— external and cranial characters. Pithecus phaeura: Lyon, 1916, p. 458— geographic dis- tribution. Macaca irus phaeura: Chasen, 1940a, p. 69— geographic distribution. Poole & Schantz, 1942, p. 245— holotype cataloged. M[acaca] /[ascicularis] phaeura: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Cynomolgus suluensis Mearns, 1905, p. 430— holotype, usnm 125324 (Coll. No. 5750), adult male, skull only, collected at foot of Crater Lake Mountain, Jolo I., Philippine Islands, by E. A. Mearns, 16 Nov. 1903. W. C. O. Hill, 1974, p. 522-cited as a synonym of Macaca irus philippinensis I Geoffroy, [ 1 843]. Fooden, 1991, p. 22— cited as a synonym of Macaca fascicu- laris fascicularis (Raffles, [1821]). Macaca suluensis: Lyon & Osgood, 1909, p. 284— ho- lotype cataloged. Macaca suluana: Alcasid, [1970], p. 24— incorrect spell- ing of Cynomolgus suluensis Mearns, 1905. Macaca sulvensis: Chiarelli, 1972, p. 2 1 3 — incorrect spelling of Cynomolgus suluensis Mearns, 1905. [Macacus] suluensis: Raven, 1935, p. 237— geographic distribution. Pithecus suluensis: Hollister, 1912, p. 37 — listed. Elliot, 1913, p. 252— specific status indeterminate. M[acaca] p[hilippinensis] suluensis: Rabor, 1986, p. 1 38 — geographic distribution. Cynomolgus cagayanus Mearns, 1905, p. 431— holo- type, usnm 125325 (Coll. No. 5771), adult male, skin and skull, collected in Cagayan Sulu I., Philippine Is- lands, by E. A. Mearns, 25 Feb. 1904. P. H. Napier, 1981, p. 13— cited as a synonym of Macaca fascicu- laris philippinensis I. Geoffroy, [1843]. Fooden, 1991, p. 22— cited as a synonym of Macaca fascicularis fas- cicularis (Raffles, [1821]). Macaca cagayana: Lyon & Osgood, 1909, p. 283 — ho- lotype cataloged. Poole & Schantz, 1942, p. 241 — holotype cataloged. [Macacus] cagayanus: Raven, 1935, p. 236— geographic distribution. Pithecus cagayanus: Hollister, 1912, p. 36— geographic distribution. Macaca irus cagayana: W. C. O. Hill, 1974, p. 525 — external and cranial characters. M[acaca] p[hilippinensis] cagayanus: Rabor, 1986, p. 138— geographic distribution. Cynomolgus fuscus: Moszkowski, 1909, pp. 143, 302 (not Miller, 1903a, p. 476)— misidentification of spec- imens collected in Sumatra. Pithecus validus Elliot, 1909, p. 252 — holotype, bm(nh) 1881.6.30.2, adult male, skin and skull, collected in "Cochin China," Vietnam, by M. Boucard, before 1 882. Macaca validus: Kloss, 1919c, p. 348— taxonomic com- parison. M[acaca] i[rus] validus: Kloss, 1921, p. 75— taxonomic comparison. Macaca irus valida: Kellogg, 1945, p. 1 19— geographic distribution. M[acaca] f[ascicularis] valida: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. P. H. Napier, 1981, pp. 13, 19— external characters; holo- type cataloged. Macaca fascicularis validus: Van Peenen et al., 1971, p. 134 (part)— female specimens collected at Sontra Peak; geographic variation. Pithecus alacer Elliot, 1909, p. 253— holotype, bm(nh) 1909.4.1.36 (Coll. No. 1454), adult male, skin and skull, collected at Bliah (= Selatbliat), P. Kundur, Ke- pulauan Riau, Indonesia, by H. C. Robinson and E. Seimund, 18 Aug. 1908; paratype (available but not explicitly cited in original description), bm(nh) 1909.4.1.37 (Coll. No. 1495, adult female, skin and skull), collected at Selatbliat, P. Kundur, Indonesia, by H. C. Robinson and E. Seimund, 20 Aug. 1908. H. C. Robinson & Kloss, 1914, pp. 393, 394-type his- tory. Chasen, 1925, p. 93— cited as a synonym of Ma- caca irus F. Cuvier (I. Geoffroy, 1 826). Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cu- vier (I. Geoffroy, 1826). P. H. Napier, 1981, pp. 13, 15— cited as a synonym of Macaca fascicularis fasci- cularis (Raffles, [1821]); holotype cataloged. [Macacus] alacer: Raven, 1935, p. 236— geographic dis- tribution. M[acaca] i[rus] alacer: Dammerman, 1926b, p. 316 — geographic distribution. Pithecus karimoni Elliot, 1909, p. 254— holotype, bm(nh) 1909.4.1.34 (Coll. No. 1662), adult male, skin and skull, collected at Monos, P. Karimun, Kepulauan Riau, Indonesia, by H. C. Robinson and E. Seimund, 30 Aug. 1908; paratype, bm(nh) 1909.4.1.35 (Coll. No. 1636, adult female, skin and skull), collected at Mon- os, P. Karimun, Indonesia, by H. C. Robinson and E. Seimund, 29 Aug. 1908. H. C. Robinson Kloss, 1914, p. 393, 394— type history; taxonomic comparison. Chasen, 1925, p. 93— cited as a synonym of Macaca irus F. Cuvier (I. Geoffroy, 1826). Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier (I. Geoffroy, 1826). P. H. Napier, 1981, pp. 13, 15- cited as a synonym of Macaca fascicularis fascicularis (Raffles, [1821]); holotype cataloged. [Macacus] karimoni: Raven, 1935, p. 236— geographic distribution. 72 FIELDIANA: ZOOLOGY M[acaca] i[rus] karimoni: Dammerman, 1926b, p. 316— geographic distribution. Pithecus laetus Elliot, 1909, p. 255— holotype, bm(nh) 1909.4.1.21 (Coll. No. 849), subadult male, skin and skull, collected in P. Tinggi, West Malaysia, by H. C. Robinson, 25 Jun. 1908; paratype (available but not explicitly cited in original description), bm(nh) 1909.4.1.27 (Coll. No. 844, juvenile female, skin and skull), collected in P. Tinggi, by H. C. Robinson, date unspecified, presumably Jun. 1908. Elliot, 1913, p. 236— geographic distribution, P. Tinggi and P. Tio- man, West Malaysia. H. C. Robinson & Kloss, 1914, p. 393— type history; taxonomic comparisons. Macaca laetus: Kloss, 191 la— taxonomic comparison. [Macacus] laetus: Raven, 1935, p. 236— geographic dis- tribution. Macaca irus laetus: H. C. Robinson, 1919, p. 325 — zoogeography. Macaca irus laeta: Chasen, 1940a, p. 69— geographic distribution. Macaca fascicularis laeta: Medway, 1966, p. 16— natural history. P. H. Napier, 1981, pp. 13, 18— external char- acters; holotype cataloged. M[acaca]f[ascicularis] laeti.J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Pithecus dollmani Elliot, 1909, p. 256— holotype, bm(nh) 1909.4.1.20 (Coll. No. 1065), late subadult male, skin and skull, collected at Changi, Singapore I., by H. C. Robinson and E. Seimund, 22 Jul. 1908 (cf. P. H. Napier, 1981, p. 15). H. C Robinson & Kloss, 1914, p. 393— type history. Kloss, 1919c, p. 347— taxonomic comparison. Chasen, 1924b, p. 59— taxonomic com- parison. Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier (I. Geoffroy, 1826). P. H. Napier, 1981, pp. 13, 15— cited as a synonym of Ma- caca fascicularis fascicularis (Raffles, [ 1 82 1]); holotype cataloged. [Macacus] dollmani: Raven, 1935, p. 236— geographic distribution. [Macaca irus] dollmani: Sody, 1949, table 1— external and cranial characters. Pithecus bintangensis Elliot, 1909, p. 257— holotype, bm(nh) 1909.4.1.23 (Coll. No. 812), adult male, skin and skull, collected at Sungei Biru, P. Bintan, Kepu- lauan Riau, Indonesia, by H. C. Robinson, date un- specified, presumably Jun. 1908; paratypes, bm(nh) 1909.4.1.24 (Coll. No. 780, adult female, Sungei Biru, P. Bintan, 1 1 Jun.), bm(nh) 1 909.4. 1 .25 (Coll. No. 784, subadult female, Sungei Biru, P. Bintan, 12 Jun.), bm(nh) 1 909.4. 1 .26 (Coll. No. 746, adult female, Pasir Panjang, P. Bintan, 8 Jun.), and bm(nh) 1909.4.1.29 (Coll. No. 870, adult female, Tanjong Sauh, P. Batam, 10 Jun.), skins and skulls, collected in Kepulauan Riau, Indonesia, by H. C. Robinson and/or E. Seimund, 1908. Elliot, 1913, p. 246— type series information. Robinson & Kloss, 1914, pp. 393, 394— type history; taxonomic comparison. H. C. Robinson, 1916, p. 62— cited as a synonym of Pithecus fascicularis (Raffles, [1821]). Chasen, 1924b, p. 59— cited as a synonym of Macaca irus F. Cuvier (I. Geoffroy, 1826). Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier (I. Geoffroy, 1826). P. H. Napier, 1981, pp. 13, 15— cited as a synonym of Macaca fascicularis fascicularis (Raffles, [1821]); type history. [Macacus] bintangensis: Raven, 1935, p. 236— geo- graphic distribution. M[acaca] i[rus] bintangensis: Dammerman, 1926b, p. 3 1 6 —geographic distribution. M[acaca] f[ascicularis] bintangensis: Groves & Weitzel in Weitzel et al., 1988, pp. 5, 96, 99-external char- acters; geographic distribution. Macaca mordaxThomas & Wroughton, 1909c, p. 380— holotype, bm(nh) 1909.1.5.27 (Coll. No. 613), sub- adult male, skin and skull, collected at Tjilatjap (= Cilacap), sea level, W Java, Indonesia, by G. C. Shor- tridge, 19 Oct. 1907; paratype, bm(nh) 1909.1.5.28 (Coll. No. 783, juvenile female, skin and skull), col- lected at Cilacap, sea level, W Java, Indonesia, by G. C. Shortridge, 12 Nov. 1907 (cf. Thomas & Wrough- ton, 1909a, p. 373). Miller, 1933, p. 8 -possible phys- iological differentiation. [Macacus] mordax: Raven, 1935, p. 236— geographic distribution. Pithecus mordax: Elliot, 1913, p. 232— geographic dis- tribution; external and cranial characters. P[ithecus]f[ascicularis] mordax: Schwarz, 1913, p. 296— taxonomic comparison. Macaca fascicularis mordax: Hooijer, 1962b, p. 46— cranial characters; taxonomic comparisons. P. H. Na- pier, 1981, pp. 13, 18— external characters; holotype cataloged. Macaca irus mordax: Dammerman, 1928, p. 300— spec- imens collected in Sumba; taxonomic comparisons. Sody, 1929, p. 165— included in fauna of Java. Sody, 1933, p. 93— specimens collected in Bali; taxonomic comparisons. Mertens, 1936, p. 315 — specimens col- lected in Bali, Lombok, Sumbawa, and Flores; taxo- nomic comparisons. Sody, 1949, p. 131— specimens collected in Java; taxonomic comparisons. Macaca resima Thomas & Wroughton, 1909c, p. 381 — holotype, bm(nh) 1909.1.5.31 (Coll. No. 1219), late subadult male, skin and skull, collected at Tasikmalaja (= Tasikmalaya), 1 145 ft (= 350 m), Preanger (region), W Java, Indonesia, by G. C. Shortridge, 18 Jan. 1908. Dammerman in Chasen, 1940a, pp. 68, 70— cited as a synonym of Macaca irus mordax Thomas & Wroughton, 1909c. P. H. Napier, 1981, pp. 13, 18- cited as a synonym of Macaca fascicularis mordax Thomas & Wroughton, 1 909c; holotype cataloged. [Macacus] resimus: Raven, 1935, p. 237— geographic distribution. Pithecus resimus: Elliot, 1 9 1 3, p. 224— external and cra- nial characters; taxonomic comparison. Pithecus lapsus Elliot, 1910, p. 343— holotype, usnm 124863 (Coll. No. 3418), adult male, skin and skull, collected at Tanjung Pamuja, P. Bangka, Indonesia, by W. L. Abbott, 1 9 Jun. 1 904; paratypes, usnm 1 24969 (Coll. No. 35 19, adult male, skin and skull fragments, 1 9 Jul.) and usnm 1 24970 (Coll. No. 352 1 , adult male, skin and skull, 20 Jul.), collected at Tanjung Batu, P. Belitung, Indonesia, by W. L. Abbott, 1904. Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier, 1818 (I. Geoffroy, 1826). Poole & Schantz, 1942, p. 243 -holotype cataloged. P. H. Napier, 1981, p. 13— cited as a synonym of M. fascicularis fascicu- laris (Raffles, [1821]). [Macacus] lapsus: Raven, 1935, p. 236— geographic dis- tribution. Macaca irus lapsus: Sody, 1937, p. 247— specimens col- lected in P. Bangka; external and cranial characters; taxonomic comparison. Pithecus agnatus Elliot, 1910, p. 344— holotype, usnm FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 73 1 14409 (Coll. No. 1471), adult male, skin and skull, collected in P. Tuangku, Kepulauan Banyak, Indo- nesia, by W. L. Abbott, 26 Jan. 1 902; paratypes, usnm 1 14408 (Coll. No. 1464, late subadult male, skin and skull, 24 Jan.), usnm 111410 (Coll. No. 1472, adult male, skin and skull, 26 Jan.), usnm 1 1 44 1 1 (Coll. No. ?, subadult male, skull only, 28 Jan.), and usnm 1 14643 (Coll. No. ?, late juvenile male, skull only, 25 Jan.), collected in P. Tuangku, Kepulauan Banyak, Indo- nesia by W. L. Abbott, 1902. Chasen, 1940a, p. 66- cited as a synonym of Macaca irus irus F. Cuvier, 1818 (I. Geoffroy, 1826). Poole & Schantz, 1942, p. 241- holotype cataloged. P. H. Napier, 1981, p. 13— cited as a synonym of M. fascicularis fascicularis (Raffles, [1821]). [Macacus] agnatus: Raven, 1935, p. 236— geographic distribution. [Macaca irus] agnatus: Sody, 1949, Table 1— external and cranial characters. Pithecus lingungensis Elliot, 1910, p. 344— holotype, usnm 104853 (Coll. No. 492), adult male, skin and skull, collected in Pulo Lingung (= P. Lagong), Ke- pulauan Natuna, Indonesia, by W. L. Abbott, 19 Jun. 1900. Chasen, 1940a, p. 69— cited as a synonym of Macaca irus pumila Miller, 1 900. Poole & Schantz, 1942, p. 243-holotype cataloged. P. H. Napier, 1981, p. 14— cited as a synonym of M. fascicularis pumila Miller, 1900. [Macacus] lingungensis: Raven, 1935, p. 236— geo- graphic distribution. [Macaca irus] lingungensis: Sody, 1949, Table 1— ex- ternal and cranial characters. M[acaca] /[ascicularis] lingungensis: Groves & Weitzel in Weitzel et al., 1988, pp. 5, 96, 99— external char- acters; geographic distribution. Pithecus lautensis Elliot, 1910, p. 345— holotype, usnm 104854 (Coll. No. 614), adult male, skin and skull, collected in P. Laut, Kepulauan Natuna, Indonesia, by W. L. Abbott, 9 Aug. 1900. Chasen, 1940a, p. 69- cited as a synonym of Macaca irus pumila Miller, 1900. Poole & Schantz, 1942, p. 243— holotype cat- aloged. P. H. Napier, 1 98 1 , p. 1 4— cited as a synonym of M. fascicularis pumila Miller, 1900. [Macacus] lautensis: Raven, 1935, p. 236— geographic distribution. [Macaca irus] lautensis: Sody, 1949, Table 1— external and cranial characters. Pithecus sirhassensis Elliot, 1 9 1 0, p. 345 — holotype, usnm 104852 (Coll. No. 468), subadult male, skin and skel- eton, collected in Sirhassen I. (= P. Serasan), Kepu- lauan Natuna, Indonesia, by W. L. Abbott, 8 Jun. 1900. Chasen, 1940a, p. 69— cited as a synonym of Macaca irus pumila Miller, 1 900. Poole & Schantz, 1942, p. 245 -holotype cataloged. P. H. Napier, 1981, p. 14— cited as a synonym of M. fascicularis pumila Miller, 1900. [Macacus] sirhassensis: Raven, 1935, p. 237— geograph- ic distribution. [Macaca irus] sirhassensis: Sody, 1949, Table 1— exter- nal and cranial characters. M[acaca] f[ascicularis] sirhassensis: Groves & Weitzel in Weitzel et al., 1988, pp. 5, 96, 99— external char- acters; geographic distribution. Pithecus carimataeEWiot, 1910, p. 346— holotype, usnm 125101 (Coll. No. 3646), adult male, skin and skull, collected at Teluk Pai, P. Karimata, Indonesia, by W. L. Abbott, 24 Aug. 1 904; paratype (available but not explicitly cited in original description), usnm 125102 (Coll. No. 3661), adult male, skin and skull, collected at Teluk Pai, P. Karimata, Indonesia, by W. L. Abbott, 27 Aug. 1904. Lyon, 1911, p. 137-type history. Chas- en, 1935b, p. 2— taxonomic comparison. Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier, 1818 (I. Geoffroy, 1826). Poole & Schantz, 1942, p. 242— holotype cataloged. P. H. Napier, 1981, p. 13— cited as a synonym of M. fascicularis fascicu- laris (Raffles, [1821]). [Macacus] carimatae: Raven, 1935, p. 236— geographic distribution. [Macaca irus] carimatae: Sody, 1949, Table 1 —external and cranial characters. Macaca irus? carimatae: Sody, 1 949, p. 1 3 1 —specimens collected in P. Pelapis Tengah and P. Serutu, Kepu- lauan Karimata, Indonesia. Pithecus mandibularis Elliot, 1910, p. 347— holotype, usnm 142225 (Coll. No. 4196), late subadult male, skin and skull, collected at Sungei Sama (= Sungai Ambawang), near Pontianak, Kalimantan, Indonesia, by W. L.Abbott, 18 Jun. 1905. Lyon, 1911, p. 137- type history. Chasen & Kloss, 1931, p. 10— said to be based on individual variables. Poole & Schantz, 1 942, p. 244— holotype cataloged. P. H. Napier, 1981, p. 13— cited as a synonym of M '. fascicularis fascicularis (Raffles, [1821]). [Macacus] mandibularis: Raven, 1935, p. 236— geo- graphic distribution. Macaca irus mandibularis: Gyldenstolpe, 1920, pp. 3, 14— specimen collected at Kaboerau, Kalimantan, In- donesia; said to be probably "not [valid] as a distinct race." Sody, 1949, p. 130— taxonomic comparison. [Macaca irus] mandibula: Sody, 1949, Table 1— incor- rect spelling of Pithecus mandibularis Elliot, 1910: ex- ternal and cranial characters. M[acaca]f [ascicularis] mandibularis: Groves & Weitzel in Weitzel et al., 1988, pp. 5, 96, 99— external char- acters; geographic distribution. Pithecus baweanus Elliot, 1910, p. 347 — holotype, usnm 151829 (Coll. No. 5565), adult male, skin and skull, collected in P. Bawean, Java Sea, Indonesia, by W. L. Abbott, 24 Nov. 1907; paratype (available but not explicitly cited in original description), usnm 151830 (Coll. No. 5566), adult female, skin and skull, collected in P. Bawean, Indonesia, by W. L. Abbott, 24 Nov. 1907. Lyon, 1911, p. 137-type history. [Macacus] baweanus: Raven, 1935, p. 236— geographic distribution. Macaca irus baweana: Chasen, 1940a, p. 70— geographic distribution. Poole & Schantz, 1 942, p. 24 1 — holotype cataloged. Macaca irus baweanus: Sody, 1949, p. 132 — specimens collected in P. Bawean; taxonomic comparison. M[acaca]f [ascicularis] baweana: J . R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Pithecus cupidus Elliot, 1910, p. 348— holotype, usnm 151831 (Coll. No. 5584), adult male, skin and skull, collected in P. Matasiri, Java Sea, Indonesia, by W. L. Abbott, 8 Dec. 1907; paratype (available but not explicitly cited in original description), usnm 154368 (Coll. No. 6388), adult male, cranium only, collected 74 FIELDIANA: ZOOLOGY in P. Matasiri, Indonesia, by W. L. Abbott, presum- ably 1907-1908. Lyon, 1911, p. 137— type history. [Macacus] cupidus: Raven, 1935, p. 236— geographic distribution. Macaca irus cupida: Chasen, 1940a, p. 70— geographic distribution. Poole & Schantz, 1942, p. 242— holotype cataloged. [Macaca irus] cupidus: Sody, 1949, Table 1— external and cranial characters. M[acaca /[ascicularis] cupida: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Pithecus lingae Elliot, 1910, p. 349— holotype, usnm 101603 (Coll. No. ?), subadult male, skin and skull, collected in Linga I. (= P. Lingga), Kepulauan Lingga, Indonesia, by W. L. Abbott, 23 Jul. 1899; paratype, usnm 101602 (Coll. No. ?), late juvenile male, skin and skull, collected in P. Lingga, Indonesia, by W. L. Abbott, 23 Jul. 1899. Chasen, 1925, p. 93 -cited as a synonym of Macaca irus F. Cuvier (I. Geoffroy, 1 826). Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier, 1818 (I. GeofTroy, 1826). Poole & Schantz, 1942, p. 243— holotype cataloged. P. H. Na- pier, 1981, p. 13— cited as a synonym of M. /ascicu- laris f ascicularis (Raffles, [1821]). [Macacus] lingae: Raven, 1935, p. 236— geographic dis- tribution. M[acaca] i[rus] lingae: Dammerman, 1926b, p. 316 — geographic distribution. Pithecus impudens Elliot, 1910, p. 350— holotype, usnm 115675 (Coll. No. 1956), adult male, skin and skull, collected in P. Sugi, Kepulauan Riau, Indonesia, by W. L. Abbott, 24 Aug. 1902. Chasen, 1925, p. 93- cited as a synonym of Macaca irus F. Cuvier (I. Geof- froy, 1826). Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier, 1 8 1 8 (I. Geoffroy, 1826). Poole & Schantz, 1942, p. 242— holotype cataloged. P. H. Napier, 1981, p. 13— cited as a synonym of M. /ascicularis /ascicularis (Raffles, [1821]). M[acaca] impudens: Raven, 1935, p. 236— geographic distribution. M[acaca] i[rus] impudens: Dammerman, 1926b, p. 316— geographic distribution. Macaca irus? impudens: Sody, 1949, p. 129— specimens collected in P. Durian, Kepulauan Riau, Indonesia. Pithecus capitalis Elliot, 1910, p. 350— holotype, usnm 83271 (Coll. No. ? ), adult male, skin and skull, col- lected at Trong (= Ban Phra Muang, Trang Province), peninsular Thailand, by W. L. Abbott, 7 Mar. 1896; paratype cited in original description, usnm 83272 (Coll. No. ?, adult male, skin and skull), collected in Telibon I. [= Ko Telibong], Thailand, by W. L. Ab- bott, 27 Feb. 1896; paratypes available but not ex- plicitly cited in original description, usnm 83273 (Coll. No. ?, late juvenile male, skin and skull, Ban Phra Muang, 8 Mar.), usnm 83274 (Coll. No. 122, adult female, skin and skull, Tyching, 19 May), and usnm 83275 (Coll. No. ?, infant, skin and skull, Tyching, 19 May), collected in Trang Province, peninsular Thai- land, by W. L. Abbott, 1896. Kloss, 1919c, p. 347- taxonomic comparison. Weitzel et al., 1988, p. 109— cited as a synonym ofMacaca/ascicularis bintangensis (Elliot, 1909). Macaca capitalis: Gyldenstolpe, 1919, p. 131— geo- graphic distribution. Macaca irus capitalis: Chasen, 1940a, pp. 68-69— geo- graphic distribution; said to be possibly a junior syn- onym of [Pithecus] vitiis Elliot, 1 9 1 0, p. 346 (= Macaca /ascicularis aurea I. Geoffroy, [1831]). Poole & Schantz, 1942, p. 241— holotype cataloged. M[acaca] /[ascicularis] capitalis: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Macaca irus argentimembris Kloss, 191 la, p. 1 16— ho- lotype, bm(nh) 1949.426 (Coll. No. 3814, SM [= Se- langor Museum] 2068/10), adult male, skin and skull, collected in P. Pinang [2], West Malaysia, by C. B. Kloss, 3 Sep. 1910; paratypes, bm(nh) 1955.1523 (Coll. No. 3491, SM 2069/10, adult female, P. Pinang [2], 3 Sep.), zrc 4-054 (Coll. No. 3488, SM 2070/10, late subadult male, P. Redang, 2 Sep.), and zrc 4-055 (Coll. No. 3833, SM 2071/10, latejuvenile male, P. Redang, 5 Sep.), skins and skulls, collected in West Malaysia by C. B. Kloss, 1910. J. E. Hill, 1960, p. 31 -type history. Weitzel et al., 1988, p. 109— cited as a syn- onym of M. /ascicularis bintangensis (Elliot, 1 909). [Macacus] irus argentimembris: Raven, 1935, p. 236— geographic distribution. Macaca /ascicularis argentimembra: Kloss, 1911a, pp. 177, 181 —external and cranial characters; taxonomic comparison; type series information. Macaca /ascicularis argentimembris: J. R. Napier & Na- pier, 1967, pp. 349, 403— geographic distribution. P. H. Napier, 1981, pp. 13, 16— external characters; ho- lotype cataloged. Macaca [nemestrinus-Gruppe] aff. adusta: Elbert, 1912, p. 101 (not Miller, 1906a, p. 559)— misidentification of specimens collected in P. Sumbawa. Schwarz, 1912, p. 304— misidentification of specimens collected in P. Sumbawa. Pithecus /ascicularis aff. limitis Schwarz, 1912, p. 304— nomen nudum. Pithecus /ascicularis limitis Schwarz, 1913, p. 296— ho- lotype, zsbs 1 9 1 1 /2 1 04 (Coll. No. 1 9), adult male, skin and skull, collected at Lelogama, P. Timor, Indonesia, by C. B. Haniel, 28 May 1911; paratypes, zsbs 1911/ 2101 (Coll. No. 15, juvenile male, skin and skull, Fatu Timau, NE, 1200 m, 20 May), zsbs 191 1/2102 (Coll. No. 16, adult female, skin and skull, Lelogama, 22 May), zsbs 191 1/2103 (Coll. No. 17, adult male, skin only, Lelogama, 27 May), zsbs 1911/2105 (Coll. No. 21, infant male, skin and skull, Lelogama, 29 May), zsbs 1 9 1 1/2 1 06 (Coll. No. 22, juvenile male, skin only, Lelogama, 30 May), zsbs 1911/2107 (Coll. No. 24, infant male, skin only, Lelogama, 3 Jun.), zsbs 1911/ 21082 (Coll. No. 63, adult male, skin and skull, Kuat- nana, 300 m, 20 Jul.), zsbs 191 1/2109 (Coll. No. 69, adult male, skull only, Bokong, 180 m, 26 Jul.), and zsbs 1911/2110 (Coll. No. 68, adult male, skin only, Bokong, 180 m, 24 Jul.) collected in P. Timor, In- donesia, by C. B. Haniel, 1911. Schwarz, 1914, p. 117— type series information. M[acaca] /[ascicularis] limitis: J. R. Napier & Napier, 1967, pp. 349, 404— geographic distribution. [Macacus] irus limitis: Raven, 1935, p. 236— geographic distribution. Macaca irus limitis: Kellogg, 1945, p. 119— geographic distribution. Macaca irus limitus: Sody, 1949, p. 133, Table 1— in- correct spelling of Pithecus /ascicularis limitis Schwarz, 1913; taxonomic comparison. Pithecus mansalaris Lyon, 1916, p. 452— holotype, usnm FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 75 114560 (Coll. No. 1639), adult male, skin and skull, collected in Pulo Mansalar (= P. Mursala), west of Sumatra, Indonesia, by W. L. Abbott, 10 Mar. 1902; paratypes, usnm 1 14559 (Coll. No. 1624, adult male, skin and skull, 7 Mar.) and usnm 1 14561 (Coll. No. ?, adult female, skull only, 9 Mar.), collected in P. Mursala, Indonesia, by W. L. Abbott, 1902. Chasen, 1940a, p. 66— cited as a synonym of Macaca irus irus F. Cuvier, 1818 (I. Geoffroy, 1826). Poole & Schantz, 1942, p. 244— holotype cataloged. P. H.Napier, 1981, p. 13— cited as a synonym of Macaca fascicularis fas- cicularis (Raffles, [1821]). [Macacus] mansalaris: Raven, 1935, p. 236— geographic distribution. [Macaca irus] mansalaris: Sody, 1949, Table 1 —external and cranial characters. Pithecus sp.: Thomas, 1928, p. 832 — misidentification of juvenile male collected at Tay Ninh, Vietnam. Macaca mulatta: Thomas, 1928, p. 832 (not Zimmer- mann, 1780, p. 195)— misidentification of specimens (Coll. Nos. 777, 778) collected at Phu Quoc (not Phu- Qui), Vietnam. Macaca irus sublimitis Sody, 1932, p. 338— based on five specimens, unseen by Sody, reported by Dam- merman (1928, p. 300) as Macaca irus mordax Tho- mas & Wroughton, 1909c; type series, mzb 865 (adult female, Kambera [= Payeti-Kambaniru], 26 Mar.), mzb 866 (adult male, Kambera, 27 Mar.), mzb 867 (juvenile male, Mao Marroe, 4 May), mzb 868 (late juvenile female, Mao Marroe, 8 May), and mzb 869 (subadult female, Mao Marroe, 10 May), collected in P. Sumba, Indonesia, by P. F. Franck and/or Denin, 1925. [Macaca irus] sumbae: Sody, 1933, p. 94— lapsus for Macaca irus sublimitis Sody, 1932. Macaca irus sublimitus: Sody, 1 949, p. 1 34, Table 1 — incorrect spelling of Macaca irus sublimitis Sody, 1 932; taxonomic comparisons, distinctiveness of subspecies "still waits for thorough confirmation"; geographic distribution; type series information. W. C. O. Hill, 1974, p. 529— recognized provisionally, said to be probably a synonym of Macaca irus mordax Thomas & Wroughton, 1909c. M[acaca] /[ascicularis] sublimitis: J. R. Napier & Na- pier, 1967, pp. 349, 404— geographic distribution. Macaca irus submordax Sody , 1949, p. 133, Table 1 — holotype, rmnh unnumbered (Coll. No. E85), adult male, skin and skull, collected at Desa Poetjang, G. Agoeng (= Gunung [Mountain] Agung), E Bali, In- donesia, by H. J. V. Sody, date unspecified, apparently 1927-1931; paratypes, rmnh (Coll. No. E34, juvenile male, skin and skull, Sendang), rmnh (Coll. No. E64, adult male, skin and skull, Sendang), rmnh (Coll. No. E74, adult male, skin and skull, Sendang), rmnh (Coll. No. El 36, adult male, skull only, Jembrana, Negara district), and rmnh (Coll. No. El 39, subadult female, skin and skull, Jembrana, Negara district), collected in Bali, Indonesia, by H. J. V. Sody, apparently 1927— 1931; mzb 2001 (Coll. No. 1), adult female, skin and skull, collected at Batu Meringgit, Bali, Indonesia, by P. F. Franck, 8 Oct. 1928; mzb 6521 (Coll. No. 17/ 92/33, juvenile male), mzb 6522 (Coll. No. 15/93/33, adult male), and mzb 6523 (Coll. No. 16/04/33, adult male), skins and skulls, collected at Banju Wetan, Bali, Indonesia, by J. J. Menden, 18 Jul. 1933; and one unlocated additional female specimen cited by Sody in Table 1. W. C. O. Hill, 1974, p. 525 -cited as a synonym of Macaca irus mordax Thomas & Wrough- ton, 1909c. P. H. Napier, 1981, p. 13 -cited as a syn- onym of Macaca fascicularis mordax Thomas & Wroughton, 1909c. Simia mauritius: W. C. O. Hill, 1974, p. 507 (not Grif- fith, 1821, p. 58; an "entirely black" monkey)— cited as a synonym ofMacaca irus irus. P. H. Napier, 198 1, p. 13— cited as a synonym of Macaca fascicularis fas- cicularis. Holotype— Not preserved. Simia fascicularis Raffles, [1821] (p. 246) apparently is based on a single specimen, probably an adult female ("The body is about twenty inches long, and the tail a little more. . . . Canines short."). This specimen was part of a collection made in Sumatra for Raf- fles by P. Diard and A. Duvaucel during the period March 1819-March 1820 (T. S. Raffles in S. Raf- fles, 1830, pp. 703, 713); the original description of Simia fascicularis is included in Raffles's pub- lished catalog of this collection. The fate of the holotype is unclear. Part of the Raffles/Diard/Duvaucel Sumatra collection, possibly accompanied by the manu- script of Raffles's catalog, was sent to England in March 1820, and another, larger part was sent to England in or before June 1 820 (T. S. Raffles in S. Raffles, 1830, pp. 440, 447, 453, 715; Watson et al. in S. Raffles, 1830, p. 716). The text of Raf- fles's catalog was read at a meeting of the Linnaean Society of London on 5 December 1820. Three specimens of M. fascicularis (sexes unspecified) sent by Raffles from Sumatra ultimately reached the museum of the Zoological Society of London (Waterhouse, 1838, p. 7; cf. Vigors, 1830, p. 642), and two additional specimens sent by Raffles from Sumatra or Java reached the museum of the East India Company, London (Horsfield, 1851, pp. iii- iv, 17). When these two museums were disband- ed—the former in 1852-1860 and the latter in 1879— their collections were transferred to the bm(nh) (Thomas, 1906, pp. 40, 63), but none of Raffles's specimens of M. fascicularis are now pre- served in the bm(nh) (cf. P. H. Napier, 1981, p. 16). Another part of the Sumatra collection was transferred by Raffles to Diard and Duvaucel and subsequently was sent to France by Duvaucel in 1820-1821 (T. S. Raffles in S. Raffles, 1830, p. 720; Lacaze, 1856, col. 536). Included in this part of the collection was an adult female specimen of M. fascicularis (mnhn 362/233), which coinciden- 76 FIELDIANA: ZOOLOGY tally is a paralectotype of M. f. aurea I. Geoffroy, [1831] (see below). This female, however, almost certainly is not the holotype of Simla fascicularis; Raffles considered the specimens that he trans- ferred to Diard and Duvaucel to be "duplicates," not the "originals," which he cataloged and sent to England. Type Locality— Sumatra, restricted to "neigh- bourhood of Bencoolen [= Bengkulu]" by H. C. Robinson (1 9 1 6, p. 63). Bengkulu, in southwestern Sumatra, was the headquarters of Raffles, Diard, and Duvaucel (cf. Steenis-Kruseman, 1 950, p. 425). Distribution (Fig. 25)— The broad geographic range of M. f. fascicularis in Southeast Asia in- cludes the following continental and insular com- ponents: the southern part of the Indochinese Pen- insula (including southern Vietnam, Cambodia, and southern Thailand) and adjacent islands (ex- cluding the Con Son group and Ko Khram Yai); the northeastern part of the Isthmus of Kra; the Malay Peninsula and adjacent islands; Sumatra and adjacent islands (excluding the Nicobar Is- lands, the P. Simeulue group, Kepulauan Menta- wai, and P. Enganno); Borneo and adjacent islands (excluding P. Maratua), Java and adjacent islands (excluding P. Karimunjawa and P. Kemujan); Lesser Sunda Islands from Bali to Timor (exclud- ing P. Komodo and many nearby islets); and the south-central Philippines (including Cagayan Sulu, Sulu Archipelago, and western Mindanao). At the margins of the subspecific range are three inter- subspecific or interspecific contact zones, inhab- ited by mixed-phenotype populations, as follows: (1) central and eastern parts of the Indochinese Peninsula— an intersubspecific contact zone be- tween ranges of M. f. fascicularis and M.f aurea, or an interspecific contact zone between ranges of M. fascicularis and M. mulatta; (2) eastern and southern parts of the Isthmus of Kra and adjacent southwestern islands— an intersubspecific contact zone between ranges of M. f. fascicularis and M. f aurea; and (3) eastern and central Mindanao, southern Negros, and possibly nearby islands— an intersubspecific contact zone between ranges of M. f. fascicularis and M. f philippinensis (see below, M. f. aurea— Distribution; M. f. philippinensis— Distribution). diagnosis— General color of dorsal surface of trunk buffy to yellowish gray to medium brown (825 of 868 postinfantile specimens examined), occasionally dark brown (43 specimens; cf. Table 1 , Appendix 3); erythrism variable, pale hair an- nulations pale yellowish to golden to rufescent (cf. Table 3, Appendix 4); crown same color as back, or brighter (847 of 873 postinfantile specimens examined), occasionally with an indistinct black- ish wash (24 specimens), rarely contrastingly darker than back (2 specimens; cf. Table 5, Appendix 5); preauricular hairs directed anteriorly, forming part of lateral facial crest (crest transzygomatic, 501 of 504 specimens examined), rarely directed poste- riorly, partly covering ears (crest infrazygomatic, 2 specimens; crest asymmetric, 1 specimen; cf. Table 6, Appendix 6); T 67-150% of HB in 311 adult specimens examined (cf. Table 10, Appendix 9). Remarks— For comments on nine quasi-dis- tinct populations that are here included within M. f fascicularis, see above (Subspecific Taxonomy). Specimens Examined— Total, 1,550: skins and skulls, 739; skins only, 285; skulls only, 526 (see Appendix 1). Macaca fascicularis aurea I. Geoffroy, [1831], p. 58 Macacus aureus I. Geoffroy, [1831], pp. 58, 76, pi. 2— for details concerning type series, see below. Eydoux & Souleyet [& Gervais], 1841, p. 6, pi. 2— taxonomic comparisons. I. Geoffroy, [1843], p. 566— taxonomic history. Gray, 1849, p. 4— cited as a synonym of Ma- cacus cynomolgus [= Macaca fascicularis]. I. Geoffroy, 1851, p. 27— taxonomic comparison; type series in- formation. Blyth, 1875, p. 7— cited as a synonym of Macacus cynomolgus [= Macaca fascicularis]; said to be based on "a casual individual variety from Pegu." J. Anderson, 1879, p. 73— cited as a synonym of Ma- cacus cynomolgus [= Macaca fascicularis]; type said to be probably a market specimen. Elliot, 1913, p. 229— cited as a synonym of Pit hecus irus [= Macaca fascicularis]. Rode, 1938, p. 223— cited as a synonym of Macaca irus (F. Cuvier, 1818; I. Geoffroy, 1826); holotype cataloged. Macacus auratus Miiller, [1840], p. 49 (not E. Geoffroy, 1812, p. 93)— incorrect spelling of Macacus aureus I. Geoffroy, [1831]. Blyth, 1863, p. 9-cited as a syn- onym of Macacus carbonarius F. Cuvier (I. Geoffroy, 1826). I[nuus] aureus: Wagner, [1839], p. 138 (part)— external characters. Pith[ecus] (Mac[acus]) aureus: Dahlbom, 1856, p. 118 — external characters. Macaca irus aurea: Pocock, 1939, p. 79— taxonomic comparison; type locality restricted to Pegu. [Macaca irus] aureus: Sody, 1 949, Table 1 —external and cranial measurements. M[acaca] f[ascicularis] aurea: J. R. Napier & Napier, 1967, pp. 349, 403— distribution. Cercopithecus Cynosurus: Heifer, 1838, p. 858 (not Sco- poli, 1786, p. 44)— misidentification; natural history. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 77 M[acacus] cynomolgus(?): Blyth, 1844, pp. 474, 476 (not Linnaeus, 1766, p. 38)— specimens collected in Ara- kan. Macacus cynomolgus: Blyth, 1875, p. 7 (part; not Lin- naeus, 1766)— taxonomic comparisons. M\acacus] carbonarius: Blyth, 1847, p. 732 (not I. Geof- froy, 1826)— said to be common in Arakan. C\ynamolgus] mulatta: Reichenbach, 1862, p. 136 (part; not Zimmermann, 1780, p. 195)— misidentification; taxonomic history. Pithecus vitiis Elliot, 1910, p. 346— holotype, usnm 124176 (Coll. No. 3076), subadult male, skin and mandible (cranium lost Jan. 1938), collected at Domel I. (= Letsok-aw Kyun), Mergui Archipelago, Burma, by W. L. Abbott, 26 Jan. 1904; paratypes, usnm 104442 (Coll. No. 287, late juvenile male, Sullivan's I. [= Lanbi Kyun], 30 Jan.) and usnm 111898 (Coll. No. 757, adult female, St. Matthews I. [= Zadetkyi Kyun], 9 Dec), skins and skulls, collected in Mergui Archi- pelago, Burma, by W. L. Abbott, 1900. Pocock, 1939, p. 79— cited as a synonym of Macaca irus aurea. Chas- en, 1 940a, p. 68 — said to be possibly a senior synonym of Pithecus capitalis Elliot, 1910, p. 350 (= M. fasci- cularisfascicularis (Raffles, [1821 ])). P. H. Napier, 1981, p. 13— cited as a synonym of Macaca fascicular is au- rea. Macaca vitiis: Poole & Schantz, 1942, p. 246 — holotype cataloged. [Macaca irus] vitiis: Sody, 1949, Table 1— external and cranial measurements. Pithecus fascicularis: Wroughton, 1915, p. 699 (part; not Raffles, [1821], p. 246)— taxonomic history. Type Series — Macacus aureus I. Geoffroy, [1831, pp. 58, 76] is explicitly based on four spec- imens of Macaca fascicular is: (1) mnhn 362/234, Type Cat. No. 58a (lectotype, designated by Rode, 1938, p. 223), subadult or adult male, mounted skin only (skull measured by Elliot, 1913, p. 231; reported absent by Rode, 1938, p. 223), obtained in "Bengale" (now partly in eastern India and part- ly in Bangladesh) by L. T. Leschenault de la Tour, presumably in 1819-1820, acquired by the mnhn in Jul. 1822 (cf. Leschenault de la Tour, 1820, p. 359; 1822, p. 262; I. Geoffroy, 1851, p. 27; Rode, 1938, p. 223; Steenis-Kruseman, 1950, p. 321); (2) unpreserved specimen (not listed in catalog of I. Geoffroy, 1 85 1 , p. 27), sex unspecified, obtained in "Pegou" (= Pegu, Burma) by A. A. M. Reynaud, 1827-1828 (cf. de Rossel et al., 1829, p. 603; de Rossel, 1829, p. 609); (3) mnhn 362/233, Type Cat. No. 58b, adult female, mounted skin only, obtained in Sumatra (no further locality infor- mation available) by A. Duvaucel, 1 8 1 9-1 820, ac- quired by the mnhn 7 Sep. 1821 (cf. I. Geoffroy, 1851, p. 27; Rode, 1938, p. 223; de Lacaze, 1856, col. 535); and (4) unpreserved specimen (listed by I. Geoffroy, 1851, p. 29; not listed by Rode, 1938, p. 223), male, skin ("poils uses," I. Geoffroy, [ 1 843], p. 567) with or without skull, obtained in Java (no further locality information available), by P. Diard, 1821. I. Geoffroy ([1831], pp. 58, 76) indicated that dorsal pelage in all four of these specimens was conspicuously erythristic ("Dessus du corps d'un beau roux tiquete de noir") and cited this as the primary diagnostic character of Macacus au- reus, "Le Macaque Roux Dore." However, the infrazygomatic lateral facial crest (see above, Pel- age) that is now regarded as diagnostic of Macaca fascicularis aurea is clearly figured by I. Geoffroy ([1831], PI. 2) in his illustration of either the "Ben- gale" or Pegu specimen and is alluded to in his detailed description of the illustrated specimen ("longs poils blancs diriges en arriere . . . cachent en partie les oreilles," p. 77). No holotype is designated in the original de- scription of Macacus aureus I. Geoffroy, [1831]. In I. Geoffroy's (1851, p. 27) catalog of primates in the mhnh, the "Bengale" specimens ("L 'un des types de respece") and the Sumatra specimen {"L'un des types") are treated as coordinate type specimens of this taxon; the Java specimen is transferred to another taxon (p. 29), and the Pegu specimen is not mentioned. Ambiguous state- ments by J. Anderson (1879, p. 76) and Elliot (1913, p. 231) may or may not constitute desig- nations of the "Bengale" specimen as lectotype. The "Bengale" specimen is decisively designated as lectotype by Rode (1938, p. 223; cf. Interna- tional Code of Zoological Nomenclature, 1985, Article 74). Type Locality— Pegu, Burma (restricted by Pocock, 1 939, p. 79; cf. International Code of Zoo- logical Nomenclature, 1985, Article 72h). The lec- totype of Macacus aureus I. Geoffroy, [183 1], ob- tained in "Bengale," probably was a captive pur- chased in Calcutta (J. Anderson, 1879, p. 76); sim- ilar captive specimens in the Calcutta market, or nearby, have been reported by C. Belanger (in I. Geoffroy, [1831], p. 77), Eydoux and Souleyet [& Gervais] (1841, pp. xiv, 6), and J. Anderson (1879, p. 76). The original provenance of the lectotype is unknown. Based on the locality of one of the para- lectotypes (see above), Pocock (1939, p. 79) amended the type locality to Pegu. Distribution (Figs. 9, 25) — Southernmost Bangladesh, southwestern and southern Burma (including Preparis I.), adjacent islands in the Mer- gui Archipelago, and a small area in west-central Thailand; most of the geographic range is west of the mountain chains that form the border between 78 FIELDIANA: ZOOLOGY Burma and Thailand. Adjacent to the geographic range of M.f. aurea are two areas— one in central and eastern parts of the Indochinese Peninsula and the other in eastern and southern parts of the Isth- mus of Kra— where specimens with the diagnostic lateral facial crest character of M. f. aurea and specimens with the diagnostic lateral facial crest character of M. f. fascicularis have been collected, sometimes at the same locality. The area in eastern and southern parts of the Isthmus of Kra may be regarded as a M.f. aurea/ M.f. fascicularis contact zone; the area in central and eastern parts of the Indochinese Peninsula may be either a M.f. aurea/ M.f. fascicularis contact zone or a M. fascicularis/ M. mulatta contact zone (see below, Evolution and Dispersal). Diagnosis— General color of dorsal surface of trunk buffy to medium brown, pale hair annula- tions pale yellowish to golden, rarely rufescent; crown usually brighter than back, occasionally with an indistinct blackish wash; preauricular hairs di- rected posteriorly, partly covering ears (lateral fa- cial crest infrazygomatic); T 76-104% of HB in 12 adult specimens examined. For external and cra- nial measurements, see Table 36. Specimens Examined— M. f aurea, 53: skins and skulls, 42; skins only, 8; skulls only, 3. Inter- subspecific or interspecific contact zones, 45: skins and skulls, 41; skins only, 3; skull only, 1 (see Appendix 1). Macaca fascicularis philippinensis I. Geoffroy, [1843], p. 568 For synonymy, see Fooden (1991, pp. 22, 24). Holotype— Macacus philippinesis I. Geoffroy, [1843, p. 568], is based solely on mnhn 373 (265), albino adult male, mounted skin only (omitted from type catalog published by Rode, 1938, p. 222). This monkey was obtained alive in Manila by A. Chenest and presented by him to the me- nagerie of the mnhn on 6 Aug. 1841; it died on 29 Aug. 1 842 (cf. I. Geoffroy, 1 85 1 , p. 29). A figure of the holotype accompanies the original descrip- tion (pi. 5). Type Locality— Philippine Islands, probably Luzon ("Chenest . . . l'a acquis a Manille, et il le croit originaire de cette ile," I. Geoffroy, [1843], p. 570). Distribution (Fig. 25)— Western, northern, and eastern islands of the Philippine Archipelago, in- cluding Balabac, Palawan, Culion, Mindoro, Lu- zon, Samar, Leyte, and probably other islands north of about 10°N (Fooden, 1991, pp. 6, 23). Based on two available skins, northeastern Mindanao formerly was also included in the range of this subspecies (Fooden, 1 99 1 , p. 23); however, in June 1991 one of the critical specimens (bm(nh) 1877.10.6.2, Butuan River [= Agusan River]) was discovered to consist of a late infant or early ju- venile skin mismatched with a subadult female skull, which weakens the evidence for retaining northeastern Mindanao in the subspecific range. In eastern and central Mindanao, southern Ne- gros, and possibly in nearby islands, between the ranges of M. f philippinensis and M.f. fascicularis, is a contact zone inhabited by mixed-phenotype populations. Diagnosis— General color of dorsal surface of trunk dark brown, pale hair annulations golden to refescent (pale golden in 1 subadult male, fmnh 87718); crown brighter than back (indistinct blackish streak in 1 adult male, bm(nh) 1877.10.6.1); preauricular hairs directed anteri- orly, forming part of lateral facial crest (crest transzygomatic); T 101-138% of HB in 14 adult specimens examined. For further details, see Fooden (1991, pp. 3 ff). Specimens Examined— M. f philippinensis, 98: skins and skulls, 53; skins only, 15; skulls only, 30. Probably M. f philippinensis, 15: skin only, 1; skulls only, 14. M.f. philippinensis/ M. f fascicu- laris contact zone, 110: skins and skulls, 82; skins only, 5; skulls only, 23. Probably M. f philippi- nensis/'M.f fascicularis contact zone, 12: skins and skulls, 2; skin only, 1 ; skulls only, 9 (see Appendix 1). Macaca fascicularis umbrosa Miller, 1902b, p. 789 Cercopithecus cynomolgus: Cantor, 1846, p. 176 (part; not Linnaeus, 1766, p. 38)— Nicobar Islands included in distribution. Macacus cynomolgus: Barbe, 1846, pp. 365 (not Lin- naeus, 1766)— sight records. Blyth, 1846, p. 367 (part) — zoogeography. M[acacus] cynomolgos: Blyth, 1863, p. 9 (part; not G. Cuvier, 1798, p. 98)— specimen collected in "Nico- bars." Cercocebus carbonarius:Ze\eboT, [ 1 869], p. 7 (not I. Geof- froy, 1826, p. 588)— cited as "Var. a" of Inuus cy- nomolgus; specimen collected in Great Nicobar I. M[acacus] carbonarius: J. Anderson, 1881, p. 65 (part; FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 79 Table 36. Macaca fascicularis aurea: external and cranial variation in adult specimens examined. Latitude Adult females Adult males HB T/HB GL HB T/HB GL Locality (N) (mm) x 100 (mm) (mm) x 100 (mm) Haungtharaw 16°30' 125.7 Ye Forest 16°10' 108.4 Wong, 65 km E 15°55' 400 96.3 99.0 Wong, 85 km E 15°55' 460 76.1 98.5 Ban Tamrong Phato 14°54' 486 90.7 113.0 529 103.8 128.2 Taungbyauk Chaung 13°45' 116.2 Kathema Kyun 13°39' 126.3 Kadan Kyun 12°30' 520 95.2 129.8 Mergui 12°26' 128.7 Tenasserim1 12°05' 480 490 512 95.8 94.9 94.7 104.4 114.0 109.1 630 84.9 132.3 Zadetkyi Kyun 9°57' 470 89.4 114.3 Mean 471.1 91.1 108.54 559.7 94.6 128.50 SD 35.4 7.1 6.62 61.1 9.4 2.41 N 7 7 9 3 3 6 1 Summary statistics for female sample: HB, mean = 494.0, SD = 16.3; T/HB x 100, mean = 95.1, SD = 0.6; GL, mean = 109.17, SD = 4.80. not I. Geoffrey, 1 826)— cited as "Smaller var." of Ma- cacus cynomolgus; external characters of specimen collected in Nicobar Islands. Macacus umbrosus Miller, 1902b, p. 789 — for details concerning type series, see below. Lyon & Osgood, 1909, p. 285— holotype cataloged. Macaca umbrosa: Miller, 1933, p. 8— possible physio- logical differentiation. Poole & Schantz, 1942, p. 246— holotype cataloged. [Macacus] umbrosa: Raven, 1935, p. 237— geographic distribution. [Cynomolgus] umbrosus: Trouessart, 1904, p. 16— geo- graphic distribution. Pithecus umbrosus: Elliot, 1913, p. 228— external and cranial characters. [Silenus] umbrosus: Stiles & Nolan, 1929, p. 537 — par- asites. Macaca irus umbrosa: Pocock, 1 939, p. 82— external and cranial characters. Kellogg, 1944, p. 76— taxonomic comparisons; zoogeography. M[acaca]f[ascicularis] umbrosa: J. R. Napier & Napier, 1967, pp. 349, 402— geographic distribution. Type Series— Holotype (by original designa- tion), usnm 111795 (Coll. No. 888), adult male, skin and skull, collected in Little Nicobar I., Ni- cobars, India, by W. L. Abbott, 25 Feb. 1901; paratypes, usnm 1 1 1796 (Coll. No. 889, subadult male, Little Nicobar I., 26 Feb.), usnm 111797 (Coll. No. 893, adult male, Little Nicobar I., 27 Feb.), usnm 1 1 1 792 (Coll. No. 9 1 8, subadult male, Great Nicobar I., 8 Mar.), usnm 111793 (Coll. No. 929, subadult female, Great Nicobar I., 12 Mar.), usnm 111799 (Coll. No. 939, subadult female, Great Nicobar I., 23 Mar.), usnm 111801 (Coll. No. 886, adult male, Katchall I., 21 Feb.), and usnm 111802 (Coll. No. 887, juvenile male, Katchall I., 21 Feb.), skins and skulls, collected in Nicobars, India, by W. L. Abbott, 1901. Type Locality— Little Nicobar I., Nicobars, India. Distribution (Fig. 25)— Katchall I., Little Nic- obar I., and Great Nicobar I., Nicobars, India. Kloss (1903a, p. 114) reported that monkeys are absent in the Nicobars north of Katchall I. Diagnosis— General color of dorsal surface of trunk blackish (dark brownish gray in 1 adult male, usnm 1 1 1801), pale hair annulations pale yellow- ish; crown yellowish brown, hairs conspicuously annulated with pale yellowish; preauricular hairs directed anteriorly, forming part of lateral facial crest (crest transzygomatic, 5 of 8 specimens ex- amined), or preauricular hairs directed posteri- orly, partly covering ears (lateral facial crest in- frazygomatic, 3 specimens); subauricular hairs pale brownish gray, not elongated; outer surface of thighs and shanks pale brownish gray; T 115-11 6% of HB in 3 adult specimens examined. For external and cranial measurements, see Appendixes 7-9 and 12 and Fooden and Albrecht (1993, p. 538). Specimens Examined— Total 8: skins and skulls, 8 (see Appendix 1). 80 FIELDIANA: ZOOLOGY Macaca fascicularis fusca Miller, 1903a, p. 476 Macacus fuscus Miller, 1903a, p. 476 — for details con- cerning type series, see below. Lyon & Osgood, 1909, p. 283— holotype cataloged. Macaca fusca: Thomas, 1923, p. 591— specimen col- lected in P. Simeulue. [Cynomolgus] fuscus: Trouessart, 1904, p. 16— geo- graphic distribution. Pit hecus fuscus: Elliot, 1913, p. 228— external and cra- nial characters. Pithecus fuscus fuscus: Lyon, 1916, p. 457— geographic distribution. Macaca irus fusca: Chasen, 1940a, p. 69— geographic distribution. Poole & Schantz, 1 942, p. 242— holotype cataloged. Kellogg, 1944, p. 76— taxonomic compar- isons; zoogeography. M[acaca] f[ascicularis] fusca: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Type Series— Holotype (by original designa- tion), usnm 1 14164 (Coll. No. 1348), adult male, skin and skull, collected at Telok Dalam (= Lhok Dalam), east coast of Simalur I. (= P. Simeulue), Indonesia, by W. L. Abbott, 20 Nov. 1901 (for collecting locality details, see collector's field cat- alog, usnm); paratypes, usnm 114162 (Coll. No. 1346, adult female, P. Simeulue, 18 Nov. 1901), usnm 1 14163 (Coll. No. 1347, adult male, P. Si- meulue, 18 Nov. 1901), usnm 114165 (Coll. No. 1349, adult female, P. Simeulue, 20 Nov. 1901), usnm 114166 (Coll. No. 1350, adult female, P. Simeulue, 20 Nov. 1901), usnm 114167 (Coll. No. 1354, adult male, P. Simeulue, 26 Nov. 1901), usnm 114168 (Coll. No. 1387, adult male, Siba- boh, P. Simeulue, 17 Dec. 1901), usnm 114169 (Coll. No. 1396, adult male, Labuan Badjan Bay [= Labuhanbajau], P. Simeulue, 1 Jan. 1902), usnm 114247 (Coll. No. 1397, late juvenile male, P. Lasia, 4 Jan. 1902), and usnm 1 14248 (Coll. No. 1398, adult male, P. Lasia, 5 Jan. 1902), skins and skulls, collected in Indonesia by W. L. Abbott. Type Locality— Telok Dalam (= Lhok Dal- am), east coast of Simalur Island (= P. Simeulue), west of northern Sumatra, Indonesia. Distribution (Fig. 25)— P. Simeulue, west of northern Sumatra, Indonesia. Diagnosis— General color of dorsal surface of trunk blackish, pale hair annulations pale yellow- ish; crown blackish, hairs conspicuously annulated with pale yellowish; preauricular hairs usually di- rected posteriorly, partly covering ears (lateral fa- cial crest infrazygomatic, 9 of 13 specimens ex- amined), occasionally directed anteriorly, forming part of lateral facial crest (crest transzygomatic, 4 specimens); subauricular hairs pale gray, not elon- gated; outer surface of thighs and shanks blackish; T 90-108% of HB in 10 adult specimens exam- ined. For external and cranial measurements, see Table 37, Figures 26 and 27, and Appendix 12. Remarks— For comments on T and crown hair annulation in M. fascicularis fusca, see below (M. f. lasiae— Remarks). Behavior in M. fascicularis fusca has been compared with that in Sumatran M. f. fascicularis by van Schaik and van Noord- wijk (1985a, p. 141) and Sugardjito et al. (1989, p. 197). These authors suggest that sexual dimor- phism of size may be less in M. fascicularis fusca than in Sumatran M. f. fascicularis; however, re- duced sexual dimorphism is not apparent in avail- able cranial measurements of M. fascicularis fusca (Table 37, Fig. 26). Specimens Examined— Total, 19: skins and skulls, 12; skins only, 7 (see Appendix 1). Macaca fascicularis lasiae (Lyon, 1916, p. 453) Macacus fuscus: Miller, 1903a, p. 476 (part)— specimens collected in P. Lasia; taxonomic comparisons. Pithecus fuscus lasiae Lyon, 1916, p. 453— for details concerning type series, see below. W. C. O. Hill, 1 974, p. 5 1 1 —cited as a synonym of Macaca irus fusca Mil- ler, 1903a. P. H. Napier, 1981, p. 13-cited as a syn- onym of Macaca fascicularis fusca Miller, 1903a. Macaca irus lasiae: Chasen, 1940a, p. 69— geographic distribution. Poole & Schantz, 1942, p. 242— holotype cataloged. Kellogg, 1944, p. 76— taxonomic compar- isons; zoogeography. Type Series— Holotype (by original designa- tion), usnm 1 14248 (Coll. No. 1398), adult male, skin and skull, collected in P. Lasia, southeast of P. Simeulue, Indonesia, by W. L. Abbott, 5 Jan. 1902; paratype, usnm 114247 (Coll. No. 1397), late juvenile male, skin and skull, collected in P. Lasia, Indonesia, by W. L. Abbott, 4 Jan. 1902. These two specimens also are paratypes of M. fas- cicularis fusca Miller, 1903a, p. 476 (see above; cf. International Code of Zoological Nomencla- ture, 1985, Article 72d). Type Locality— Palau Lasia, southeast of P. Simeulue, west of northern Sumatra, Indonesia. Distribution (Fig. 25)— Pulau Lasia, west of northern Sumatra, Indonesia. Diagnosis— General color of dorsal surface of trunk blackish, pale hair annulations pale yellow- ish; crown blackish, hair annulations inconspic- uous; preauricular hairs directed posteriorly, part- ly covering ears (lateral facial crest infrazygomatic, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 81 Table 3 7 . Sexual dimorphism of greatest length of skull (mm) compared in Macacafascicularisfusca and Sumatran Macaca fascicularis fascicularis (cf. Fig. 26). Adult females Adult males Latitude Mean SD N Mean SD N Ratio1 M. fascicularis J usca 2°00'-2°59'N 102.22 2.81 4 121.77 1.88 7 1.19 Sumatran M. f. fascicularis 5°00'-5°59'S — — 0 120.15 4.58 4 — 4°00'-4°59'S 103.10 — 1 120.65 4.96 6 1.17 3°00'-3°59'S 98.35 2.90 2 117.64 3.61 7 1.20 2°00'-2°59'S 103.40 — 1 — — 0 — 1°00'-1°59'S 94.90 — 1 114.90 — 1 1.21 0°00'-O°59'S 100.25 4.04 4 111.16 1.24 7 1.11 0°00'-0°59'N — — 0 119.00 — 1 — 1°00'-1°59'N 95.03 1.82 3 116.27 3.99 3 1.22 2°00'-2°59'N 95.30 0.71 2 115.40 — 1 1.21 3°00'-3°59'N 97.64 2.16 10 115.86 3.42 21 1.19 4°00'-4°59'N 103.40 — 1 120.63 4.92 3 1.17 Total 98.18 3.31 25 116.65 4.39 54 1.19 Male mean divided by female mean. 1 of 2 specimens examined), or directed anteriorly, forming part of lateral facial crest (crest transzy- gomatic, 1 specimen); subauricular hairs pale gray- ish brown, not elongated; outer surface of thighs blackish, becoming pale grayish brown on outer surface of shanks; T 1 1 8% of HB in 1 adult spec- imen examined. For external and cranial mea- surements, see Figure 27 and Appendix 12. Remarks— Although M.f. lasiae, endemic to P. Lasia, is known from only 2 specimens (1 late juvenile male, 1 adult male), T in these 2 speci- mens is clearly longer— both relatively and abso- lutely—than in 13 specimens of M. fascicularis fusca (Fig. 27), which inhabits nearby P. Simeulue and which M. f. lasiae otherwise strongly resem- bles; the difference in T was noted previously by Miller (1903a, p. 477) and Lyon (1916, p. 453). Judging from the general pattern of T variation in M. fascicularis, the longer tail in M. f. lasiae prob- ably is primitive, and the shorter tail in M. fas- cicularis fusca probably is derived (Figs. 16, 18; Appendix 12). Crown hair annulations are less conspicuous in the 2 specimens examined of M. f lasiae than in 1 3 specimens examined of M. fascicularis fusca. The blackish dorsal pelage of M. f lasiae in deep-water P. Lasia (and M. fascicularis fusca in deep-water P. Simeulue) contrasts strikingly with the yellowish brown to golden brown dorsal pelage of M. f fascicularis in shallow- water P. Tuangku (usnm 1 14408-1 14410), an island that is only 50 km east of P. Lasia but that is on the Sumatran shelf (Figs. 3, 4). The blackish dorsal pelage of M. f lasiae and M. fascicularis fusca also contrasts strikingly with the brownish dorsal pelage of M. f. fascicularis in deep-water P. Nias, which is about 95 km southeast of P. Lasia (Appendix 3). Specimens Examined— Total, 2: skins and skulls, 2 (see Appendix 1). Macaca fascicularis atriceps Kloss, 1919c, p. 347 Macaca irus atriceps Kloss, 1919c, p. 347— for details concerning type series, see below. Kloss, 1921, p. 76 — external and cranial characters; taxonomic compari- sons. Kloss, 1926, p. 358— external and cranial char- acters; taxonomic comparisons. Poole & Schantz, 1 942, p. 243— holotype cataloged. Weitzel et al., 1988, p. 105— cited as a synonym of Macaca fascicularis bin- tangensis (Elliot, 1 909) (= M. f fascicularis). M[acaca] f[ascicularis] atriceps: J. R. Napier & Napier, 1967, pp. 349, 403— geographic distribution. Type Series— Holotype (by original designa- tion), usnm 236622 (Coll. No. 2283), adult male, skin and skull, collected in Koh Khram I. (= Ko Khram Yai), near Cape Liant (= Laem Samae San), southeastern Thailand, by C. B. Kloss, 30 Oct. 1916; paratypes, bm(nh) 1939.891 (Coll. No. 2284, adult male), bm(nh) 1939.892 (Coll. No. 2287, adult female), usnm 236618 (Coll. No. 2282, 82 FIELDIANA: ZOOLOGY ■ 125- ■ ■ ■ ■ ■ A 1 ■ ■ A 120- 3 M VI 1 ■ ■ ■ ■ ■ ■ A ■ f I ■ s -• — ■ ■ I ■ length of © ■ ■ ■ * ■ Greatest 8 D a H A D 0 100 a 0 A % 95- D 00 0 0 1 a a 90 1 0 600 6°S 4 2 0 2 4°N Latitude o ■ Sumatra: females, males A A Simeulue: females, males Fig. 26. Variation of greatest length of skull in adult Macacafascicularisfusca (Indonesia: P. Simeulue) com- pared with variation in adult Sumatran M.f.fascicularis (cf. Table 37). adult male), usnm 236619 (Coll. No. 2286, adult female), usnm 236620 (Coll. No. 2289, subadult female), usnm 236621 (Coll. No. 2290, juvenile male), zrc 4-012 (Coll. No. 2288, adult female), zrc 4-013, skull, and zrc 4-733, skin (Coll. No. 2285, adult male), skins and skulls, collected in Ko Khram Yai, Thailand, by C. B. Kloss, 30 Oct. 1916. Type Locality— Ko Khram Yai, near Laem Samae San, southeastern Thailand. Distribution (Fig. 25)— Ko Khram Yai, Thai- land. Diagnosis— General color of dorsal surface of trunk buffy to medium brown, pale hair annula- tions pale yellowish; crown with a narrow, sharply defined dark brown to blackish patch (Fig. 7) that extends laterally as far as middle of each eye and posteriorly as far as vertex or (rarely) occiput; preauricular hairs directed anteriorly, forming part of lateral facial crest (crest transzygomatic); T 1 09- 500 ^450 ♦ juv.* A A » ' A subad. ?ageA ^ * A A A A 1 i 1 ■ ■ 400 350 300 350 400 450 500 550 Head and body length (mm) ♦ Lasia: males A A Simeulue: females, males Fig. 27. Tail length vs. head and body length in Ma- cacafascicularisfusca (Indonesia: P. Simeuleu) and M. f lasiae (Indonesia: P. Lasia); specimens are adult, except as otherwise indicated (three specimens). 1 28% of HB in 9 adult specimens examined. For external and cranial measurements, see Table 38, Figure 28, and Appendix 12. Remarks- Kloss (1919c, p. 348; 1921, p. 76; 1926, p. 358) indicated that M. f. atriceps differs cranially from M. f. condorensis and Indochinese M . fascicularis in size of the rostrum, supraorbital ridges, orbits, zygomatic arches, mandible, and molars and in shape of the tooth rows and palate. As evidence, Kloss cited measurements of some of these structures in 9 specimens of M.f. atriceps (4 adult females, including 1 young adult regarded by Kloss as a subadult, and 5 adult males) and in 3 specimens of M. f. condorensis ( 1 adult female and 2 adult males); no measurements are cited for specimens of Indochinese M. fascicularis. New data, derived from 5 additional adult spec- imens of M.f. condorensis (1 female, 4 males) and 32 adult specimens of Indochinese M. fascicularis (22 females, 10 males), permit reevaluation of the six size characters specified by Kloss (Table 38, Fig. 28). For each of these six cranial charac- ters, size variation in M. f. atriceps overlaps that in M. f. condorensis and Indochinese M. fascicu- laris. Relative sizes of the rostrum, supraorbital FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 83 Table 38. Comparison of cranial proportions in Indochinese (I-C) Macaca fascicularis, M. f. atriceps, and M. J condorensis (cf. Fig. 28; Kloss, 1919c, p. 348; 1921, p. 76; 1926, p. 358). Adult females Adult males Population Mean SD Extremes N Mean SD Extremes N Rostral -postrostral ratio (R/PR x 100) I-C M. fascicularis 49.5 4.6 41.1-56.4 20 60.8 3.1 55.5-65.1 7 M. f. atriceps 51.0 2.8 48.1-54.5 4 58.4 3.6 54.0-63.9 5 M. f condorensis 50.4 _ 46.6-54.1 2 Relative biorbital breadth (BB/GL1 59.8 x 100) 3.4 55.3-63.3 5 I-C M. fascicularis 51.9 2.7 45.3-55.9 18 50.0 2.1 47.2-54.0 10 M. f atriceps 51.0 2.1 48.5-53.1 4 50.3 2.2 48.4-53.9 5 M. f condorensis 48.9 _ 48.0-49.7 2 Relative orbital height (OH/GL x 51.2 100) 4.4 46.5-56.8 6 I-C M. fascicularis 20.6 1.7 17.5-24.6 19 18.4 1.1 16.9-20.9 10 M. f atriceps 19.4 1.1 18.3-20.7 4 17.2 1.3 15.8-19.2 5 M. f condorensis 21.5 - 20.4-22.5 2 Relative zygomatic breadth (ZB/GL 19.1 x 100) 1.2 17.8-20.6 6 I-C M. fascicularis 67.8 2.4 63.2-71.3 22 68.9 1.7 66.4-71.2 10 M. f atriceps 66.8 2.4 63.1-68.4 4 68.4 2.3 66.8-71.8 4 M. f condorensis 65.7 - 64.9-66.6 2 Relative mandibular length (ML/GL 67.8 x 100) 2.6 65.1-71.3 6 I-C M. fascicularis 73.0 2.1 70.5-77.6 20 75.8 3.1 71.5-81.3 9 M. f. atriceps 72.4 0.5 71.8-72.9 4 75.6 1.6 73.4_77.9 5 M. f condorensis 73.2 - 72.4-74.0 2 74.9 1.6 72.6-76.4 6 Relative length of maxillary molar row (Ml M3/GL x 100) I-C M. fascicularis 19.9 1.1 18.2-22.6 22 17.4 0.8 16.2-18.6 10 M. f atriceps 21.6 0.7 20.9-22.2 4 19.8 1.1 18.9-21.3 5 M. f condorensis 20.3 - 19.8-20.8 2 18.3 1.4 15.7-19.8 6 1 Greatest length of skull, excluding incisors. ridges, zygomatic arches, and mandible are essen- tially similar in these three populations; relative size of the orbits tends to average smaller in M. f. atriceps than in the other two populations, as sug- gested by Kloss, and relative size of the molars tends to average slightly greater in M. f. atriceps than in the other two populations, as also sug- gested by Kloss. Specimens Examined— Total 11: skins and skulls, 11 (see Appendix 1). Macaca fascicularis condorensis Kloss, 1926, p. 357 monkies: King, 1784, p. 462 — said to be abundant in Jan. 1780. Macaca irus Cuv., subsp.: Kloss, 1 92 1 , p. 75 —specimens collected in Pulo Condore (= Con Son); taxonomic comparisons. Macaca irus condorensis Kloss, 1 926, p. 357 — for details concerning type series, see below. Gibson-Hill, 1949, p. 172— type history. J. E. Hill, 1960, p. 31— typ history. Macaca fascicularis condorensis: Van Peenen et al., 1970 p. 420— specimens collected in Con Son and Hon Ba external and cranial characters. P. H. Napier, 1981 pp. 13, 17— external characters; holotype cataloged. Type Series— Holotype (by original designa tion), bm(nh) 1947.1498 (CBK [C. B. Kloss] No 269 1), adult male, skin and skull, collected in Pule Condore (= Con Son), offCochin-China (= south ern Vietnam), by "Dr. Malcolm Smith's collec tor," 20 Sep. 1919; paratypes (implicitly cited ir original description; explicitly cited by Kloss, 1 92 1 p. 75), bm(nh) 1939.893 (CBK No. 2692, adul male, 23 Sep.) and bm(nh) 1939.894 (CBK No 2693, adult female, 19 Sep.), skins and skulls, col lected in Con Son, Vietnam, by "Dr. Malcolrr Smith's collector," 1919. Type Locality— Con Son, Vietnam. Distribution (Fig. 25)— Con Son and nearby Hon Ba, two islands in the South China Sea, south east of southern Vietnam. 84 FIELDIANA: ZOOLOGY I ■ I - v- • D n_ fD^ ■ nO s D A^pB ru 0° 90 100 110 120 130 Greatest length of skull (mm) 60 a ~ 50 a fl 45- | 40 (n ^ 35 30 65 - A ■ A ♦ Q« ■ ■ ■ ■ D * ♦ A S3 D □ □ A 0. A D D 0 ♦ D D o £ a ♦ a a n f D A D □ d a ■ ■ ■ ■ ■ □1 D A 90 Afl r? A Euo D D°An D a 70 75 80 85 90 Postrostral length (mm) 90- a a 5 so- ts ed 0) ■O I ■ ■ A ■ ■ ■ A a □ J D a n a a n cfl '0 £ o bO N 60- a a 0d a n * Ai , ■ O O D 0 ♦D 18- ■ , 1 , 1 1 1 1 — 1 100 110 120 130 Greatest length of skull (mm) Females Males O ♦ M. L atri.ceps A A M. f. can dor crisis □ ■ Indochinese M. fascicularis Fig. 28. Craniometric comparisons of adult Macaca fascicularis atriceps (Thailand: Ko Khram Yai), M. f. con- dorensis (Vietnam: Con Son), and mainland Indochinese M. fascicularis (cf. Table 38). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 85 Diagnosis— General color of dorsal surface of trunk buffy to medium brown (with blackish mid- dorsal streak in 2 juveniles, zrc 4-017, 4-018), pale hair annulations pale yellowish to golden; crown with a broad dark brown to blackish patch (Fig. 7) that extends laterally as far as lateral mar- gin of each eye and posteriorly as far as vertex or occiput, margin of crown patch not sharply delim- ited; preauricular hairs directed anteriorly, form- ing part of lateral facial crest (crest transzygo- matic); T 109-127% of HB in 6 adult specimens examined. For external and cranial measure- ments, see Table 38, Figure 28, and Appendixes 7-9 and 12 and Fooden and Albrecht (1993, p. 537). Remarks — See above (M. / atriceps — Re- marks). Specimens Examined— Total, 16: skins and skulls, 1 1 ; skins only, 2; skulls only, 3 (see Ap- pendix 1). Macaca fascicularis tua Kellogg, 1944, p. 75 Macaca irus tua Kellogg, 1944, p. 75 — for details con- cerning type series, see below. W. C. O. Hill, 1974, p. 507— cited as a synonym of Macaca irus irus F. Cu- vier, 1818 (I. Geoffroy, 1826). P. H. Napier, 1981, p. 13— cited as a synonym of Macaca fascicularis fasci- cularis (Raffles, [1821]). M[acaca]f[ascicularis] tua: J. R. Napier & Napier, 1967, pp. 349, 404— geographic distribution. Type Series— Holotype (by original designa- tion), usnm 197663 (Coll. No. 626), adult male, skin and skull, collected in Pulo Muara Tua (= P. Maratua), east of Borneo, Kalimantan, Indonesia, by H. C. Raven, 21 May 1913; paratypes, usnm 197660 (Coll. No. 623, juvenile female), usnm 197661 (Coll. No. 624, adult female), and usnm 197662 (Coll. No. 625, subadult male), skins and skulls, collected in P. Maratua, Kalimantan, In- donesia, by H. C. Raven, 21 May 1913. Type Locality— Pulau Maratua, Kalimantan, Indonesia. Distribution (Fig. 25)— Pulau Maratua, east of northeastern Kalimantan, Indonesia. Diagnosis— General color of dorsal surface of trunk blackish, pale hair annulations pale yellow- ish; crown pale yellowish brown, hairs conspicu- ously annulated with pale yellowish; preauricular hairs directed anteriorly, forming part of lateral facial crest (crest transzygomatic); subauricular hairs pale ochraceous-buff, conspicuously elon- gated; outer surface of thighs brownish gray, be- coming pale brownish gray on shanks; T 1 26-1 3 1 % of HB in 2 adult specimens examined. For external and cranial measurements, see Appendix 12. Specimens Examined— Total, 4: skins and skulls, 4 (see Appendix 1). Macaca fascicularis karimondjawae Sody, 1949, p. 132 Macacus cynomolgus: Willink, 1905, p. 175 (part, not Linnaeus, 1766, p. 38)— "P. Karimon-Djawa (P. Ka- moedian)" included in geographic distribution. Macaca irus karimondjawae Sody, 1949, p. 132, Table 1 —for details concerning type series, see below. W. C. O. Hill, 1974, p. 528— external and cranial characters. Macaca fascicularis karimondjawae: P. H. Napier, 1981, p. 13— type locality information. Type Series— Holotype (by original designa- tion), rmnh 10608 (mzb 2719), adult male, skin and skull, collected in Karimon Djawa (= P. Kar- imunjawa), Java Sea, Indonesia, by W. Rom- swinckel, 28 Nov. 1930 (incorrectly given as "28.VI.1930" in original description); paratypes (cf. Sody, 1949, p. 132, Table 1), mzb 1454 (adult male, cranium only, 7 May), mzb 1455 (adult fe- male, skin and skull, 9 May), mzb 1456 (juvenile female, skin and skull, 10 May), mzb 1457 (adult female, skin and skull, 11 May), and mzb 1458 (infant male, skin and skull, 14 May), collected in P. Karimunjawa, Indonesia, by K. W. Dammer- man, P. F. Franck, and/or Denin, 1926; mzb 2717 (juvenile male) and mzb 2718 (late juvenile fe- male), skins and skulls, collected in P. Karimun- jawa, Indonesia, by W. Romswinckel, 26 Nov. 1930; and mzb 2720 (adult female, cranium only) collected in P. Kemujan, < 1 km northeast of P. Karimunjawa, Indonesia, by W. Romswinckel, 25 Nov. 1930. Type Locality— Karimon Djawa (= P. Kari- munjawa), Java Sea, Indonesia. Distribution (Fig. 25)— Pulau Karimunjawa and, presumably, nearby P. Kemujan (known from one cranium only), Java Sea, 60 km north of cen- tral Java, Indonesia. Diagnosis— General color of dorsal surface of trunk dark grayish brown, pale hair annulations pale yellowish; crown frequently with a blackish wash; preauricular hair direction not studied; T 101-1 12% of HB in 3 adult specimens examined. For external and cranial measurements, see Ap- pendix 12 and Fooden and Albrecht (1993, p. 538). 86 FIELDIANA: ZOOLOGY Specimens Examined— Total 9: skins and skulls, 6; skin only, 1; skulls only, 2 (see Appendix 1). Macaca fascicularis Subspecies Undetermined Macacus cristatus Gray, 1870, p. 30— holotype, bm(nh) 1858.4.28.9, late juvenile male, skin (albinistic) and skull, acquired in 1858 from collection of Th. G. van Lidth de Jeude, Utrecht, provenance unknown. Schle- gel, 1876, p. 101— cited as a synonym of Cercocebus cynamolgos: Schlegel (= Macaca fascicularis (Raffles, [1821])). Elliot, 1913, p. 249-cited as a synonym of Pithecus philippinensis (I. Geoffroy, [1843]). W. C. O. Hill, 1974, pp. 477, 522— cited as a synonym of Ma- caca irus philippinensis I. Geoffroy, [1843]. P. H. Na- pier, 1981, pp. 13, 20— cited as a synonym of Macaca fascicularis philippinenesis I. Geoffroy, [1843]. Food- en, 1 99 1 , p. 24— subspecies not determined; taxonom- ic history. Macacus albus: P. H. Napier, 1981, p. 20— unavailable manuscript name, author unknown, written on tag of holotype of Macacus cristatus Gray, 1870. Evolution and Dispersal Available information concerning the distri- bution, variation, natural history, and paleontol- ogy of M. fascicularis provides a basis for hypo- thetical interpretation of the evolution and dis- persal of this species. Because most of the geo- graphic range of M. fascicularis is insular, a large part of this interpretation concerns water gaps and their effect on dispersal. Four major factors are pertinent to dispersal in the insular part of the range of M. fascicularis. 1. Eustatic changes in sea level. Since the be- ginning of the Pleistocene, ca. 1.75 Ma, worldwide sea level has been strongly controlled by, and in- versely correlated with, the cyclic growth of con- tinental glaciers (Van Couvering & Kukla, 1988, p. 459; Heaney, 1991a, p. 56; Cande & Kent, 1 992, pp. 13,936, 13,938). Sea level has been approxi- mately as at present since ca. 5 Ka (Clark et al., 1978, p. 283; Fairbanks, 1989, p. 639). It rose sigmoidally to this level from about 1 20 m lower during the last glacial maximum, ca. 1 8 Ka, when the Sunda Shelf was exposed and the Indochinese Peninsula, the Malay Peninsula, Sumatra, Borneo, Java, and shallow-water fringing islands were unit- ed to form a single large landmass (Fig. 3). During the preceding interglacial, ca. 120 Ka, sea level was high, and components of this large landmass were isolated, approximately as at present. During the penultimate glacial maximum, ca. 1 60 Ka, sea level was about 160 m lower than at present, there- by adding some deep-water fringing islands to the temporarily consolidated large Sunda Shelf land- mass. Similar cyclic sea-level changes, of impre- cisely known magnitude, have occurred with a pe- riodicity of ca. 100 Ka since the beginning of the Pleistocene. Tectonic forces also have affected the distribu- tion of land and sea within the range of M. fas- cicularis. Prominent examples of such effects are the approximately 500-m uplift of the north coast of P. Sumba (Pirazzoli et al., 1991, p. 1835), the approximately 500-m subsidence and subsequent reelevation of P. Maratua (Kuenen, 1947, p. 8), and the massive volcanic explosions of Rakata (= Krakatau) and Toba (Dammerman, 1948, p. 4; Chesner et al., 1 99 1 , p. 200; Rampino & Self, 1993, p. 269). However, the zoogeographic implications of Pleistocene tectonic changes in this area are more localized than those of Pleistocene eustatic changes. 2. Swimming. The swimming distance limit of M. fascicularis probably is about 1 00 m (see above, Natural History). 3. Natural rafting. Monkeys may be transport- ed across water gaps on floating trees or larger entwined masses of vegetation (cf. Wallace, 1895, p. 74). The tendency for M. fascicularis to exploit coastal habitats (see above, Natural History) makes this species a prime candidate for such passive dispersal on natural rafts. A specific instance is reported by Hickson (1889, p. 190): "Some days after the eruption of Krakatoa in 1883 a female green monkey [M. fascicularis] was found floating on some drifting timber in the Sunda Straits. She was terribly scorched, but completely recovered, and is, I believe, still alive." The average surface current speed in the Sunda Shelf area is about 1 km/hr (E. G. W. Smith, 1974, Fig. 2.3-1; Couper, 1989, p. 50). Arbitrarily as- suming that some members of a small group of M. fascicularis could survive on a natural raft for up to about 3 days, such a group potentially could colonize a new island about 75 km distant from its original habitat; during monsoons and other storms, the current would be stronger and the dis- tance covered would be greater. The reproductive capability of a small group of M. fascicularis in- dividuals to multiply and populate islands not pre- viously inhabited by this species is demonstrated by the success of introduced populations in Mau- ritius and Angaur (Poirer & Smith, 1 974, p. 264; Sussman & Tattersall, 1986, p. 30). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 87 Table 39. Hypothetical stages in evolution and dis- persal of Macaca fascicularis. Stage I, > 1 Ma Dispersal of M. fascicularis into Sunda Shelf area. Stage II, ca. 160 Ka Dispersal and isolation of progenitors of strongly dif- ferentiated deep-water fringing-island popula- tions. M. f. umbrosa: Nicobar Islands M. f. fusca, M. f. lasiae: Simeulue and Lasia M. f. tua: Maratua M. f. philippinensis: western, northern and eastern Philippines Stage III, > 18 Ka Differentiation of progenitors of populations in the Indochinese Peninsula and northern part of the Isthmus of Kra. M. f. aurea: Indochinese Peninsula, Isthmus of Kra, Mergui Archipelago M. f. fascicularis: Indochinese Peninsula, Isthmus of Kra Stage IV, ca. 18 Ka Dispersal and isolation of progenitors of weakly dif- ferentiated deep-water fringing-island popula- tions. M. f. fascicularis: Nias M. f fascicularis: southern Philippines Stage V, < 18 Ka Isolation of progenitors of shallow-water fringing-is- land populations and populations in Penida and Lombok (deep-water). M. f karimondjawae: Karimunjawa, Kemujan M. f. atriceps: Khram Yai M. f. condorensis: Con Son, Hon Ba M. f fascicularis: other shallow-water fringing is- lands, excluding Bali M. f fascicularis: Bali, Penida, Lombok Stage VI, ca. 4.5 Ka Dispersal and isolation of progenitors of populations in eastern Lesser Sunda Islands (deep-water). M. f fascicularis: Sumbawa-Timor 4. Transport by humans. Interisland transport of captive M. fascicularis by humans is reported by Wallace (1869, p. 294) and presumably has a long, if undocumented, history in Southeast Asia. Homo erectus reached Java by 1-2 Ma, and H. sapiens reached Borneo and Australia by ca. 40 Ka (Bellwood, 1985, pp. 29, 89, 98; cf. Swisher et al., 1994, p. 1119). Although much of the dispersal of hominids in the Sunda Shelf area probably oc- curred over dry land during glacial periods of low sea level, the dispersal of H. sapiens to Australia must have been over water, which implies that some means of over- water transport was available to humans in this area at least 40 Ka (cf. Sondaar etal., 1994, p. 1261). Available evidence indicates a minimum of six major stages in the evolution and dispersal of M. fascicularis (Table 39). These are discussed below. Stage I, > 1 Ma The earliest record of M '. fascicularis is at Trinil, east-central Java, ca. 1 Ma (Table 33); the Trinil fossils are part of the Dubois collection that also includes Homo erectus and two species of leaf monkeys, Presbytis comata and Trachypithecus auratus (Hooijer, 1962b, p. 5; Theunissen et al., 1990, p. 41; Groves, 1993, p. 273). Assuming that M. fascicularis, H. erectus, and leaf monkeys dis- persed to Java from mainland Southeast Asia, they probably reached Java by dry land during one of the Early Pleistocene periods of glacial advance and low sea level (cf. J. de Vosetal., 1994, p. 131). The earliest record of M. nemestrina also is in east-central Java but is somewhat later (ca. 800 Ka) than that of M. fascicularis (Table 33). Despite this later date, zoogeographic evidence indicates that M. nemestrina probably preceded M. fasci- cularis in the Sunda Shelf area (Fooden, 1975, p. 70). Macaca fascicularis is absent in Sulawesi and Kepulauan Mentawai, faunistically distinctive is- lands off the Sunda Shelf that are inhabited by M. nemestrina or its close relatives. Since M. fasci- cularis is otherwise much more widely distributed thanM nemestrina and its relatives (Fooden, 1980, p. 4), it appears that the intrinsic ability of M. fascicularis to disperse is not inferior to that of M. nemestrina. This suggests that the M. nemestrina stock dispersed from the Sunda Shelf to Sulawesi and Kepulauan Mentawai at some time prior to 1 Ma, before M. fascicularis had arrived on the Sunda Shelf. During Late Pleistocene or Holocene, M. fas- cicularis may have been less abundant than M. nemestrina in the Malay Peninsula and Sumatra (see above, Fossils and Subfossils). If so, this would require explanation, because M . fascicularis is now generally more abundant than M. nemestrina in the Malay Peninsula, Sumatra, and elsewhere (Fooden, 1975, p. 61). Stage II, ca. 160 Ka The most strongly differentiated populations of M. fascicularis inhabit four deep-water islands or island groups: (1) Nicobar Islands, northwest of Sumatra; (2) P. Simeulue and P. Lasia, west of FIELDIANA: ZOOLOGY Sumatra; (3) P. Maratua, east of Borneo; and (4) Philippines, excluding southern islands (Fig. 25). All of these populations are very dark (Appendix 3), and their deep-water island habitats are sym- metrically arrayed adjacent to the northwestern and northeastern margins of the Sunda Shelf (cf. Kellogg, 1 944, p. 76). Assuming that dark dorsal pelage is derived in M. fascicularis, the similar pelage saturation and geographic relationships of these populations suggest that they may have had a similar evolutionary history. These populations appear to have been isolated longer than popu- lations of M. fascicularis inhabiting shallow- water fringing islands, which were isolated by rising sea level subsequent to the last glacial maximum, ca. 1 8 Ka. A working hypothesis is that the progen- itors of these strongly differentiated populations reached their deep-water habitats during the pen- ultimate glacial maximum, ca. 1 60 Ka, when sea level was about 1 60 m lower than at present, and that these populations have been isolated at least since the subsequent interglacial, ca. 1 20 Ka. Fu- ture new evidence concerning the paleontology, morphology, or molecular biology of these pop- ulations may provide a test of this hypothesis. Nicobar Islands— This group includes 19 is- lands, of which 1 1 are larger than about 2 km2. M. fascicularis inhabits only Great Nicobar I., Lit- tle Nicobar I., and Katchall I. (Kloss, 1903a, p. 1 14)— the southernmost 3 of the 1 1 larger islands. Forest cover in the Nicobars is restricted to these three islands and Tillanchong I., one of the more northern islands (Kloss, 1903a, p. 109). Monkeys, including M. fascicularis, are absent from the An- daman Islands, a forested group of deep-water is- lands that lies between the Nicobar Islands and Burma (Kloss, [1928], p. 802; Chaturvedi, 1980, p. 134). Great Nicobar is separated from Sumatra by an ocean trench that is 1 50 km wide and more than 200 m deep; for half of its width, this trench is more than 1,000 m deep. The channel between Great Nicobar and Little Nicobar is 6 km wide and less than 100 m deep, which indicates that these two islands were united during the last glacial maximum. Narrow channels (< 10 km wide) be- tween Little Nicobar and Katchall are more than 1 20 m deep, which probably indicates that these two islands were not united during the last glacial maximum. The nonvolant mammal fauna of the Nicobars is limited to one species each of shrew, tree shrew, and pig, four to six species of rats, M fascicularis, and humans (Miller, 1902b, p. 792; Kloss, [1928], p. 802; Chaturvedi, 1980, p. 133; Musser & Carle- ton, 1993, pp. 651, 660); no information is avail- able concerning the date of first arrival of humans in the Nicobars (Fuchs, 1973, p. 285; Gratton, 1 992, p. 209). The poverty of the fauna, together with the deep ocean trenches that surround this group of islands, strongly implies that these islands have never had a land connection to Sumatra or any other source of the rich mammalian fauna of Southeast Asia. This in turn implies that M. fas- cicularis reached the Nicobars by over-water dis- persal; the absence of this species in the Andaman Islands indicates that it dispersed to the Nicobars from Sumatra, the Malay Peninsula, or the Isth- mus of Kra— not from Burma. As noted above, the degree of differentiation of M. fascicularis in the Nicobars suggests that this species may have reached these islands during the penultimate gla- cial maximum, ca. 1 60 Ka; at this early date, dis- persal by natural rafting would be more likely than dispersal by human introduction. Following the arrival and isolation of M. fas- cicularis in the Nicobars, dorsal pelage in the im- migrant population apparently darkened, and skull size increased, resulting in differentiation of the subspecies M. f umbrosa. The present morpho- logical uniformity of populations of this subspe- cies in the three islands that it inhabits implies relatively recent genetic exchange among these populations; this exchange presumably occurred during the last glacial maximum, ca. 1 8 Ka, when Great Nicobar was joined to Little Nicobar and the gap between Little Nicobar and Katchall was reduced to narrow channels (< 10 km wide). The lateral facial crest pattern is infrazygomatic in three of eight M. f umbrosa specimens exam- ined (one of three Great Nicobar specimens, two of three Little Nicobar specimens, zero of two Katchall specimens) (see Appendix 6). Three al- ternative interpretations would account for the rel- atively high frequency of the infrazygomatic pat- tern in M. f. umbrosa: (1) it may indicate that the progenitors of M. f umbrosa dispersed to the Ni- cobars from the Malay Peninsula or Isthmus of Kra, in or near the geographic range of M.f aurea (Figs. 9, 25); (2) it may indicate that the range of M. f aurea extended to Sumatra during the pen- ultimate glacial maximum, when progenitors of M. f umbrosa are hypothesized to have dispersed to the Nicobars; or (3) it may be the result of independent local mutation. P. Simeulue and P. Lasia— These two islands are approximately 1 20 km west of northern Su- matra (Fig. 25). Pulau Simeulue is relatively large FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 89 (ca. 100 x 25 km) and is about 20 km northwest of the much smaller P. Lasia (ca. 6x3 km). Al- though M. fascicularis inhabits both of these is- lands, it is absent in P. Babi (ca. 8x7 km), which is only 3 km south of P. Lasia (Abbott in Miller, 1903a, p. 479). Deep trenches (> 200 m) separate P. Simeulue and P. Lasia from Sumatra and from each other. The deep trench between P. Simeulue and Sumatra is about 1 5 km wide, and that between P. Simeulue and P. Lasia is about 4 km wide. The narrow channel between P. Lasia and P. Babi probably is less than 1 00 m deep. The nonvolant mammal fauna of the Simeulue group, as reported by Kloss ([1928], p. 802; cf. Sugardjito et al., 1989, p. 197; Musser & Carleton, 1993, pp. 652, 659), is limited to one species each of civet and pig, two species of rats, M. fascicularis, and humans. This fauna, like that of the Nicobars (see above), presumably dispersed over water. Judging from the degree of differentiation of M. fascicularis in P. Simeulue and P. Lasia, this spe- cies may have dispersed to these islands— presum- ably by natural rafting— during the penultimate glacial maximum. Following isolation, the pelage of the immigrant population darkened. Genetic exchange between the populations in P. Simeulue and P. Lasia probably occurred during the last glacial maximum. During subsequent postglacial reisolation, the tail apparently shortened in the P. Simeulue population (Fig. 27; cf. above, M.f las- iae— Remarks), resulting in differentiation of M. fascicularis fusca and M.f. lasiae. Because P. Lasia probably was joined to P. Babi during the last glacial maximum, a dark population of M. fasci- cularis probably also inhabited P. Babi at that time but subsequently became locally extinct. The lateral facial crest pattern is infrazygomatic in 9 of 1 3 M. fascicularis fusca specimens exam- ined and in 1 of 2 M.f. lasiae specimens examined (Appendix 6). The geographic location of P. Si- meulue and P. Lasia strongly indicates that the progenitors of M. fascicularis fusca and M.f. lasiae dispersed to these islands from Sumatra, not from the Malay Peninsula or Isthmus of Kra (i.e., in or near the present range of M.f. aurea; Fig. 25). This implies either that the range of M. f aurea for- merly extended to Sumatra or that there has been independent mutation in these islands from the transzygomatic crest pattern to the infrazygomatic crest pattern. P. Maratua— This U-shaped island (area ca. 25 km2) is a raised coral atoll 50 km east of north- central Borneo (Fig. 25; Kuenen, 1947, p. 5). It is separated from the Bornean shelf by a deep trench (> 180 m) that is 17 km wide; the water gap is narrowed by the presence of a stepping-stone is- land, P. Kakaban (area ca. 5 km2), which inter- venes in the middle of the trench. The only mammal collector known to have vis- ited P. Maratua is H. C. Raven, who worked there in August 1912 and May 1913 (Deignan, [1960], p. 267). In P. Maratua, Raven collected M. fas- cicularis, two species of rats, possibly a few other murine rodents, and one species of bat (Kellogg, 1944, p. 75; Musser & Carina, 1982, p. 6; M. D. Carleton, usnm, letter 16 Dec. 1993). Judging from the degree of differentiation of M. fascicularis in P. Maratua, it may have dispersed to this island from Borneo during the penultimate glacial max- imum, ca. 1 60 Ka; the dispersal probably was over water— presumably by natural rafting. In isola- tion, dorsal pelage in the founder population dark- ened, resulting in differentiation of the subspecies M.f. tua. In all four M.f. tua specimens examined, the lateral facial crest pattern is transzygomatic, as in Bornean M. fascicularis. Philippines, Excluding Southern Islands— As previously discussed (Fooden, 1 99 1 , p. 24), the progenitors of current populations of M. fascicu- laris in western, northern, and eastern islands of the Philippine Archipelago may have dispersed to these islands during the penultimate glacial max- imum, ca. 1 60 Ka. Following isolation, dorsal pel- age in the founder population became darker and also tended to become erythristic, resulting in dif- ferentiation of M. f philippinensis. Stage III, > 18 Ka Indochinese Peninsula and Northern Part of Isthmus of Kra— Although zoogeographic ev- idence, cited below, indicates that M. f aurea ex- isted as a differentiated subspecies in the western part of the Indochinese Peninsula and northern part of the Isthmus of Kra prior to the last glacial maximum, ca. 18 Ka, the evolutionary history of this subspecies and that of neighboring popula- tions of M.f. fascicularis remain enigmatic. Equiv- ocal evidence concerning variation in M. fasci- cularis in the Indochinese Peninsula and Isthmus of Kra is presented in the following paragraphs. The relationship between variation in M. fasci- cularis and variation in M. mulatta—a. closely re- lated species that replaces M. fascicularis in the 90 FIELDIANA: ZOOLOGY northern part of the Indochinese Peninsula (Food- en, 1982, p. 576)— also is indicated. 1. Dorsal pelage color averages paler and less erythristic in the Indochinese Peninsula and northern part of the Isthmus of Kra than in the Malay Peninsula and Sumatra (Tables 1, 3). The pattern of pelage color resemblances is indepen- dent of local land connections; the Indochinese Peninsula and Isthmus of Kra are continuous with the Malay Peninsula, which is separated by a water gap from Sumatra. The latitude of pelage color transition is about 10°N, near the southern end of the Isthmus of Kra. The pale dorsal pelage color in M. fascicularis north of 10°N generally ap- proaches that in neighboring allopatric popula- tions of M. mulatta; however, the posterior part of the dorsal surface is strongly erythristic in M. mulatta— and not in Indochinese M. fascicularis. 2. In the Indochinese Peninsula north of about 1 3°N, which is at the northern end of the Isthmus of Kra, head and body length and skull length begin to decline northward in M. fascicularis (Fig. 10; Fooden & Albrecht, 1993, p. 532). This de- cline, which is anti-Bergmannian, is retrograde to the general relationship between size and latitude in core-area M. fascicularis and also is retrograde to the relationship between size and latitude in M. fascicularis in the Philippines, at approximately the same latitude as the Indochinese Peninsula. The northward decline of size in Indochinese M. fascicularis tends to bring the size of this species near to that of neighboring populations of M. mu- latta (Figs. 29, 30). 3. Tail length declines rapidly in the Indo- Chinese Peninsula north of about 1 3°N (Fig. 1 3). This is an acceleration of the general decline of tail length in core-area M. fascicularis north of the equator (Allen's rule); there is no comparable ac- celerated northward decline of tail length in Phil- ippine M. fascicularis. Parallel to the trend noted above for head and body length and skull length, the accelerated northward decline of tail length in Indochinese M. fascicularis tends to bring the tail length of this species near to that of neighboring populations of M. mulatta (Fig. 31; cf. Fooden, 1971, p. 29). 4. The subspecies M. f aurea inhabits a re- stricted area bordering the Bay of Bengal, mostly west of the mountain ranges that form the border between Burma and Thailand, along the western side of the Indochinese Peninsula and Isthmus of Kra, south to about 10°N (Figs. 9, 25). This sub- species is distinguished by its lateral facial crest pattern, which is infrazygomatic and which con- trasts with the transzygomatic pattern that pre- dominates in M. f fascicularis. Adjacent to the geographic range of M. f aurea are two areas— one in central and eastern parts of the Indochinese Peninsula and the other in eastern and southern parts of the Isthmus of Kra— inhabited by hetero- geneous populations that include some individuals with the infrazygomatic pattern, some with the transzygomatic pattern, and some with the asym- metric pattern; occasionally, heterogeneous pop- ulations have been sampled at a single locality. The southern part of the Indochinese Peninsula is inhabited by a now-disjunct population of M. f fascicularis (homogeneous for the transzygomatic pattern). The lateral facial crest pattern in shallow-water fringing-island populations of M. fascicularis matches that in adjacent mainland populations in Southeast Asia; the crest is infrazygomatic in the Mergui Archipelago— shallow- water islands west of the Isthmus of Kra, and the crest is transzy- gomatic in shallow- water islands east of the Isth- mus of Kra and adjacent to the Malay Peninsula (Fig. 9). This implies that M. f aurea had already differentiated, and was approximately restricted to its present geographic range, prior to the last glacial maximum (ca. 1 8 Ka), when M. fascicularis pre- sumably dispersed from mainland Southeast Asia to adjacent shallow-water islands. Pelage color, head and body length, tail length, and skull length in populations of M. f aurea are generally similar to those in populations of M. f fascicularis that inhabit similar latitudes in the Indochinese Peninsula and Isthmus of Kra. The lateral facial crest pattern in M. f aurea is similar to that in M. mulatta, where the infrazygomatic pattern is predominant (Stewart, 1933, p. 30). 5. The frequencies of mtDNA types, blood- protein alleles, and resistance to Plasmodium knowlesi malaria in populations of M. fascicularis north of the Isthmus of Kra differ markedly from those in populations south of the Isthmus of Kra (Tables 17, 19; Fooden, 1994, p. 585). The fre- quency off. knowlesi resistance in populations of M. fascicularis north of the Isthmus of Kra is sim- ilar to that in M. mulatta. Based on this evidence, the following conclu- sions may be drawn. Although suggestive, these conclusions are not adequate to explain the evo- lutionary history of M. fascicularis in the Indo- Chinese Peninsula and northern part of the Isth- mus of Kra. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 91 650 -r- 600 550 500 4-50 Males L00J £ 550 -r 500 4-50- 4-00 550 ■•* * ; v v Females IV V V jV -7 IV V 10° ! 12 ! 14- i 16 I ' ! 18 20 Latitude [N] 22 24- 26° x M. fascicularis 7 M. mulatta Fig. 29. Latitudinal variation of adult head and body length in samples of Macaco, fascicularis collected in the southern part of the Indochinese Peninsula and adjacent Isthmus of Kra, north of 10°N (cf. Fig. 10), compared with that in samples of parapatric M. mulatta collected in the northern part of the Indochinese Peninsula (Burma, Thailand, Laos, and Vietnam; specimens in amnh, bm(nh), bnhs, fmnh, mnhn, usnm, zrc, and zsi). 1 . Macaca f. aurea and M. f. fascicularis be- came differentiated in the Indochinese Peninsula and northern part of the Isthmus of Kra sometime before the last glacial maximum; this differentia- tion probably occurred on opposite sides of the mountain ranges that now form the border be- tween Burma and Thailand. Each of these sub- species probably dispersed from the mainland to shallow- water islands adjacent to its respective range during the last glacial maximum (ca. 1 8 Ka), when these islands were connected to the main- land by dry land. The population heterogeneous for the lateral facial crest pattern that inhabits cen- tral and eastern parts of the Indochinese Peninsula may indicate introgression or intergradation be- tween M. f. aurea and M. f. fascicularis, or it may indicate introgression or intergradation between M. fascicularis and M. mulatta; the population heterogeneous for the lateral facial crest pattern that inhabits eastern and southern parts of the Isthmus of Kra presumably indicates introgression or intergradation between M. f. aurea and M. f. fascicularis. The disjunct population of M. f. fas- cicularis in the southern part of the Indochinese 92 FIELDIANA: ZOOLOGY 135 130 — 125- 120- 115 110- o 105 120 Q3 "5 115 o 110- 105- 100 95- Males 90 10° 12 I x X ! V X X : - X X I ; I V *x X T IV T V j ! ;V V i V " p *■ V X X j j X T i v I V xx x X X ;V ;X % | V < i V ! ; j V i j f Females ■ i ' i ■ i V . ■ j i | , j H 16 18 20 Latitude (N) 22 24 26c M. fascicularis v M. mulatta Fig. 30. Latitudinal variation of adult greatest length of skull in samples of Macaco fascicularis collected in the southern part of the Indochinese Peninsula and adjacent Isthmus of Kra, north of 10°N (cf. Fig. 21), compared with that in samples of parapatric M. mulatta collected in the northern part of the Indochinese Peninsula (Burma, Thailand, Laos, and Vietnam; specimens in amnh, bm(nh), bnhs, fmnh, mcz, mnhn, usnm, zrc, and zsi). Peninsula was geographically continuous with the population of the same subspecies in the Malay Peninsula during the last glacial maximum (Fig. 3). 2. Within M.f. fascicularis, a strong but incom- plete barrier to gene exchange has existed in the region of the Isthmus of Kra. This barrier has had a more profound effect on differentiation of pop- ulations of this subspecies north and south of the Isthmus of Kra than the Strait of Malacca (age 5- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 93 550- 500- 450- 400- 550- 300- 250- ?oo- x X ! V V Males v i v .7) ! v ! 1 Vj y V ' % 550 c •g 500 i— 450 400 350- 300- 250 200 150 Females xx; •v-r 10° 12 U 16 18 20 Latitude (N) 22 24 26c x M. fascicularis v M. mulatta Fig. 31. Latitudinal variation of adult tail length in samples of Macaco, fascicularis collected in the southern part of the Indochinese Peninsula and adjacent Isthmus of Kra, north of 10°N (cf. Fig. 13), compared with that in samples of parapatric M. mulatta collected in the northern part of the Indochinese Peninsula (Burma, Thailand, Laos, and Vietnam; specimens in amnh, bm(nh), bnhs, fmnh, mnhn, usnm, zrc, and zsi). 18 K yr) has had on populations in the Malay the Isthmus of Kra, which approximately marks Peninsula and Sumatra. Many other vertebrates— the boundary between the Indochinese and In- including amphibians, reptiles, birds, and mam- domalayan faunal subregions (Chasen, 1940a, p. mals— also are differentiated north and south of x; Corbet & Hill, 1992, p. 3). 94 FIELDIANA: ZOOLOGY 3. Populations of M. f. aurea and M. f. fasci- cularis in the northern part of the Isthmus of Kra and Indochinese Peninsula tend to be transitional between M. fascicularis in the Sunda Shelf area and M. mulatta, which replaces M. fascicularis in the northern part of the Indochinese Peninsula. Stage IV, ca. 18 Ka P. Nias— Although P. Nias is a deep-water is- land, its population of M. fascicularis is not strong- ly differentiated (see above, Subspecific Taxono- my), which suggests relatively recent isolation of this population. The strait between P. Nias (area ca. 3,500 km2) and the west coast of northern Su- matra is about 90 km wide, but the deep channel (120-180 m) in this strait is only about 20 km wide. The nonvolant mammal fauna reported for P. Nias by Kloss ([1928], p. 802; cf. Musser & Califia, 1982, pp. 11, 18; Musser &Carleton, 1993, pp. 614, 633, 652) is limited to M. fascicularis, one species each of tree shrew, binturong, pig, trag- ulid, and pangolin, two species of deer, and five species of murine rodents. Macaca fascicularis may have dispersed to P. Nias from Sumatra during the last glacial maxi- mum, ca. 1 8 Ka, when the intervening water gap was reduced to about 20 km. During subsequent isolation, the proximodorsal surface of the tail in the P. Nias population apparently became darker and the tail apparently became shorter. In all 12 specimens examined of M. fascicularis from P. Nias, the lateral facial crest pattern is transzygo- matic, as in Sumatran M. fascicularis. Philippines, Southern Islands— As previous- ly noted (Fooden, 1991, p. 6), dorsal pelage color in M. fascicularis in southern islands of the Phil- ippine Archipelago is paler than in M. f philip- pinensis, which inhabits western, northern, and eastern islands in this archipelago. The pale south- ern populations, which are allocated to the nom- inotypical subspecies M. f fascicularis, are indis- tinguishable in pelage color from populations in Borneo. This strongly suggests that progenitors of the southern populations dispersed from Borneo to the Philippines more recently— possibly during the last glacial maximum— than progenitors of M. f philippinensis (see above). Present distributions indicate that the relatively recent dispersal of M. f fascicularis to the southern Philippines occurred via the Sulu Archipelago (Fooden, 1991, p. 28). In Negros and Mindanao, a zone of mixed pop- ulations that include pale, intermediate, and dark individuals may be the result of interbreeding be- tween early-arriving M. f philippinensis and late- arriving M. f fascicularis. Stage V, < 18 Ka Shallow-Water Fringing Islands, Ex- cluding P. Bali— During the last glacial maxi- mum, ca. 1 8 Ka, sea level was approximately 1 20 m lower than at present, and shallow-water fring- ing islands— together with Sumatra, Borneo, and Java— were part of a single large landmass that extended from the Indochinese Peninsula to P. Bali (Fig. 3; Heaney, 1991a, p. 55). This permitted dispersal, or redispersal, of M. fascicularis over dry land to what are now shallow-water islands; as a result, populations of M. fascicularis in all of these present-day islands were genetically contin- uous. Subsequently, between 1 8 Ka and 5 Ka, sea level rose to its present level, and present-day shal- low-water insular populations of M. fascicularis became isolated. Following isolation, populations of M. fascicu- laris in many shallow-water fringing islands ap- parently have tended to become slightly darker, more erythristic, and/or smaller than their ances- tral stock, assuming that this stock is represented by core-area populations (Table 9; Appendixes 3, 4; Fooden & Albrecht, 1993, p. 533). The darkest shallow-water fringing-island population, M. f karimondjawae in P. Karimunjawa and presum- ably also in P. Kemujan, may have become iso- lated somewhat earlier than most other shallow- water fringing-island populations (Appendix 3). The distinctive dark crown patches of M. f atri- ceps in Ko Khram Yai and M. f condorensis in Con Son and Hon Ba presumably also developed in isolation following elevation of sea level since the last glacial maximum. During the same inter- val, dorsal pelage erythrism in M.f fascicularis in Borneo apparently differentiated somewhat from that in the same subspecies in Sumatra and the Malay Peninsula (Tables 3, 4). Despite postglacial isolation, many shallow-wa- ter island populations of M. fascicularis appear to have retained ancestral characters. Size is similar at similar latitudes in currently isolated popula- tions in the Malay Peninsula, Sumatra, and Bor- neo (Fig. 10; Fooden & Albrecht, 1993, p. 533). Dorsal pelage erythrism and tail length are more similar in populations in the Malay Peninsula and FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 95 Sumatra, which are now isolated, than in popu- lations in the Malay Peninsula and the Indo- chinese Peninsula, which are connected by dry land (Tables 3, 10). The infrazygomatic lateral facial crest pattern apparently has persisted in popula- tions in the Mergui Archipelago despite the post- glacial isolation of these islands from the Isthmus ofKra(Fig. 9). Lesser Sunda Islands: P. Bali, Nusa Penida, and P. Lombok— Pulau Bali is the only shallow- water fringing island in the Lesser Sunda group (Fig. 3; Kitchener et al., 1990, p. 1 1 1). The strait between Java and P. Bali is approximately 3 km wide and has a maximum depth of about 65 m. The strait between P. Bali and Nusa Penida is about 10 km wide and has a depth of more than 1 20 m for about half of its width. The strait be- tween Nusa Penida and P. Lombok is approxi- mately 23 km wide and has a depth of more than 1 20 m for almost all of its width. All three of these islands are inhabited by M. fascicularis (Fig. 2C). The only other nonhuman primate known to inhabit the Lesser Sundas is one species of leaf monkey, Trachypithecus auratus, the range of which includes P. Bali and P. Lombok. In these two islands, T. auratus is represented by an endemic subspecies, T. a. kohlbruggei (P. H. Napier, 1985, p. 56; Weitzel & Groves, 1985, p. 402; Groves, 1993, p. 273). During the last glacial maximum, ca. 1 8 Ka, P. Bali was joined to Java, but Nusa Penida and P. Lombok were separate islands (Kitchener et al., 1990, p. lll;Heaney, 1991a, p. 55). At that time, populations of M. fascicularis and T. auratus in P. Bali presumably were genetically continuous with those in Java. It is unclear whether or not the last glacial maximum was the time of initial dis- persal of M. fascicularis and T. auratus from Java to P. Bali; these species may have dispersed from Java during a previous glacial cycle, and their in- sular populations may merely have been reunited during the last glacial maximum. In either event, sometime before 5 Ka, rising sea level separated P. Bali from Java and, conse- quently, separated populations of M. fascicularis and T. auratus in P. Bali from those in Java. Fol- lowing this separation, M. fascicularis in P. Bali apparently underwent reduction in body size (Figs. 1 5, 22; Appendix 7) and also underwent a marked shift in allele frequencies at blood-protein loci HbA-II, Pi, and Tf (Table 19). Concurrently, T. auratus in P. Bali apparently became subspecifi- cally distinct from T. auratus in Java. The characters that distinguish M. fascicularis and T. auratus in P. Bali from their respective conspecifics in Java link populations of these spe- cies in P. Bali to those in P. Lombok. This strongly suggests that, subsequent to their differentiation in P. Bali, populations of both species dispersed— over water— from P. Bali to P. Lombok, as pre- viously proposed by Kawamoto and Suryobroto (1985, p. 39); the dispersal of M. fascicularis to Nusa Penida, located between P. Bali and P. Lom- bok, presumably occurred at the same time. Whether the dispersal of M. fascicularis and T. auratus from P. Bali to P. Lombok was by natural rafting or by human transport is uncertain (cf. Ev- erett in Hartert, 1896, p. 593; Kitchener et al., 1990, p. 112). Stage VI, ca. 4.5 Ka Lesser Sunda Islands: P. Sumbawa-P. Tim- or— Macaca fascicularis is known to inhabit 13 of the deep-water Lesser Sunda Islands east of P. Lombok (listed in west-east order): P. Sumbawa, P. Moyo, P. Sumba, P. Rinca, Nusa Kode, P. Man- giatan, P. Seraya Besar, P. Flores, P. Adonara, P. Solor, P. Semau, P. Kambing, and P. Timor (Fig. 2C). This species reportedly is absent in P. Ko- modo and P. Padar (between P. Sumbawa and P. Rinca, north of P. Sumba), and in many nearby islets, and also is absent in P. Lomblen, P. Pantar, and P. Alor (east of P. Adonara and P. Solor, north of P. Timor) (for documentation, see Gazetteer, Appendix 2). No other nonhuman primate inhab- its any of the Lesser Sunda Islands east of P. Lom- bok. None of these islands were connected to P. Bali during the last glacial maximum, although some of these islands were interconnected at that time to form four larger islands (Lombok-Sumbawa- Moyo, Sumba, Komodo-Padar-Rinca-Kode- Mangiatan-Seraya Besar-Flores-Alor-Solor, and Semau-Kambing-Timor) (Heaney, 1991a, p. 55). The isolation of these deep-water islands is reflect- ed in their relatively poor nonvolant mammal fau- nas (Laurie & Hill, 1954, pp. 13 ff.). In P. Flores (area ca. 20,000 km2), for example, Musser (1981, p. 134) listed 16 species of nonvolant mammals (shrews, 3; M. fascicularis; civet, 1; pigs, 2; deer, 1; porcupine, 1; rats, 5; mice, 2), of which some probably were introduced by humans. This con- trasts with 38 species of nonvolant mammals re- corded for P. Bangka (area ca. 1 2,000 km2), a shal- 96 FIELDIANA: ZOOLOGY low-water island on the Sunda Shelf (Heaney, 1 984, P. 12). Four items of partly contradictory evidence seem most relevant to attempting an interpretation of the history of M. fascicularis in deep-water Lesser Sunda Islands east of P. Lombok: (1) Dorsal pelage color in these islands is essentially similar to dorsal pelage color in Java, P. Bali, and P. Lombok (Ap- pendixes 3, 4). (2) External and cranial size in these islands is similar to size in P. Bali and P. Lombok (Figs. 15, 16, 22, 23); size is greater in Java. (3) Blood-protein allele frequencies at loci HbA-II, Pi, and Tf in these islands are similar to allele fre- quencies in Java (Table 1 9); frequencies of these alleles are divergent in P. Bali and P. Lombok. (4) Subfossils of M. fascicularis in P. Timor cave de- posits appear relatively late (ca. 4.5 Ka), concur- rently with the appearance of remains of domestic animals and pottery (see above, Fossils and Subfossils). The evidence of dorsal pelage color and Timor cave deposits suggests relatively recent dispersal of M. fascicularis to deep-water Lesser Sunda Is- lands east of P. Lombok. The blood-protein evi- dence indicates that the differentiated populations of M. fascicularis in P. Bali and P. Lombok prob- ably were not directly involved in dispersal of this species to islands east of P. Lombok. Assuming that the differentiated populations in P. Bali and P. Lombok inhabited those islands before M. fas- cicularis dispersed to islands east of P. Lombok (see above), the implication is that populations of M. fascicularis east of P. Lombok were derived from a source in Java, or elsewhere west of P. Bali. This geographic incongruity, together with the ar- cheological evidence of the Timor cave deposits, strongly supports the hypothesis that the popu- lations of M. fascicularis in the Lesser Sundas east of P. Lombok are the result of human introduc- tion, beginning ca. 4.5 Ka; this hypothesis has been previously proposed by Musser (1981, p. 133), Kawamoto and Suryobroto (1985, p. 39), and Glover (1986, p. 159). Multiple introductions would be required to es- tablish populations of M. fascicularis in all of the islands east of P. Lombok that it now inhabits. If Java was the source of these introductions, exter- nal and cranial size became convergently reduced in these islands. This is inconvenient to the hy- pothesis, but not impossible, considering the fre- quency of insular dwarfing in M. fascicularis (see above, External Measurements and Proportions; Cranial Characters). Auffenberg (1981, p. 242) suggests that the ab- sence of M. fascicularis in P. Komodo (area 393 km2) and P. Padar (13 km2) may be attributable to an insufficient year-round supply of fruit in these islands. This is puzzling, because small islands im- mediately east of P. Komodo and P. Padar support populations of M. fascicularis (P. Rinca, 278 km2; Nusa Kode [= P. Oewada Sami], 9.6 km2; P. Ser- aya Besar [= P. Seraja], 3.6 km2; P. Mangiatan [= P. Mengjatan], 1 km2). The insular distribution of M. fascicularis in this area may be affected in some way by the distribution of the Komodo giant mon- itor, a known predator of these monkeys (see above, Natural History) that is restricted to the following seven islands (listed in west-east order): P. Ko- modo, Nusa Mbarapu (0.6 km2, occasionally vis- ited by giant monitors), P. Padar, P. Rinca, Nusa Kode, Gili Motang (= Gili Mota), and western P. Flores (Auffenberg, 1981, p. 40). M. fascicularis coexists with the giant monitor only in three of the more easterly of these islands (P. Rinca, Nusa Kode, and P. Flores). The relatively recent dispersal of M. fascicularis to deep-water Lesser Sunda Islands, indicated by evidence cited above, is somewhat surprising. This species has been present in east-central Java since ca. 1 Ma (see above), and, during the maximum Pleistocene glaciation, the widest water gap be- tween Java and Sumbawa may have been only 400 m or less (Kitchener et al., 1 990, p. 1 1 0). However, as indicated by Heaney (1991a, p. 57), during gla- cial maxima, rainfall in the Lesser Sundas was much lower than at present. Perhaps forest cover was inadequate to sustain M. fascicularis in these islands during glacial maxima, when low sea level and narrow water gaps might otherwise have fa- cilitated the eastward dispersal of this species. Acknowledgments The generous cooperation of curators and staff members of institutions in which specimens ex- amined are preserved (see Introduction) has been indispensable to the conduct of this research and is gratefully acknowledged. For valuable supple- mentary advice and assistance, I also thank the following: H. Abdulali (bnhs), N. Aggimarangsee (Chiang Mai University, Thailand), M. Aimi (pri), G. H. Albrecht (University of Southern California, Los Angeles), W. Y. Brockelman (Mahidol Uni- versity, Bangkok), M. D. Carleton (usnm), S. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 97 Chakraborty (zsi), C. Fleming (amnh), W. K.-H. Fuchs (amnh), L. Gordon (usnm), C. P. Groves (Australian National University, Canberra), L. R. Heaney (fmnh), P. D. Jenkins (bm(nh)), Y. Ka- wamoto (pri), J. C. Kerbis Peterhans (fmnh), J. W. Koeppl (fmnh), S. M. Lanyon (fmnh), G. G. Musser (amnh), Shukor Md. Nor (fmnh), B. D. Patterson (fmnh), D. P. Schmidt (usnm), S. Un- nithan (bnhs), and P. F. D. Van Peenen (U.S. Navy, retired, Chicago). I am also grateful to three re- viewers, who painstakingly scrutinized the manu- script and provided numerous helpful suggestions. Literature Cited Abdulali, H. 1967. The birds of the Nicobar Islands, with notes on some Andaman birds. Journal of the Bombay Natural History Society, 64: 139-190. Adi Haji Taha. 1985. The re-excavation of the rock- shelter of Gua Cha, Ulu Kelantan, West Malaysia. Federation Museums Journal, n.s., 30: i-xi, 1-134. Aggimarangsee, N. 1992. Survey for semi-tame col- onies of macaques in Thailand. Natural History Bul- letin of the Siam Society, 40: 103-166. Aimi, M. 1981. Fossil Macaca nemestrina (Linnaeus, 1766) from Java, Indonesia. Primates, 22: 409-413. Aimi, M., and F. Aziz. 1985. Vertebrate fossils from the Sangiran Dome, Mojokerto, Trinil and Sambung- macan areas, pp. 155-168. In Watanabe, N., and D. Kadar, eds., Quaternary Geology of the Hominid Fos- sil Bearing Formations in Java. Special Publication No. 4. Geological Research and Development Centre, Indonesia. Aimi, M., A. Bakar, and J. Supriatna. 1982. Mor- phological variation of the crab-eating macaque, Ma- caca fascicularis (Raffles, 1821), in Indonesia. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 2: 51-56. Alcasid, G. L. [1970]. Checklist of Philippine Mam- mals. National Museum, Manila, 5 1 pp. [For date of publication, see Heaney, L. R., and D. S. Rabor, 1982, Occasional Papers of the Museum of Zoology, Uni- versity of Michigan, 699: 29.] Aldrich-Blake, F. P. G. 1 980. Long-tailed macaques, pp. 147-165. In Chivers, D. J., ed., Malayan Forest Primates: Ten Years' Study in Tropical Rain Forest. Plenum, New York. Allen, G. M., and H. J. Coolidge, Jr. 1 940. Mammal and bird collections of the Asiatic Primate Expedition. Mammals. Bulletin of the Museum of Comparative Zoology at Harvard College, 87: 131-166. Anderson, C. M., and C. F. Bielert. 1 994. Adolescent exaggeration in female catarrhine primates. Primates, 35: 283-300. Anderson, J. 1879. Anatomical and Zoological Re- searches: Comprising an Account of the Zoological Results of the Two Expeditions to Western Yunnan in 1868 and 1875, vol. 1. Bernard Quaritch, London, 985 pp. [For date of publication, see Corrigenda, be- tween pp. xii and xiii.] 1881. Catalogue of Mammalia in the Indian Museum, Calcutta. Part I. Primates, Prosimiae, Chi- roptera, and Insectivora. Indian Museum, Calcutta, xv + 225 pp. Andrews, C. W. 1900. Mammalia, pp. 22-33. In An- drews, C. W., ed., A Monograph of Christmas Island (Indian Ocean): Physical Features and Geology, with Descriptions of the Fauna and Flora. British Museum (Natural History), London. Andrews, R. C. 1911. Around the world for the Mu- seum. American Museum Journal, 11: 21-24. . 1943. Under a Lucky Star: A Lifetime of Ad- venture. Viking Press, New York, 300 pp. Angst, W. 1973. Pilot experiments to test group tol- erance to a stranger in wild Macaca fascicularis. Amer- ican Journal of Physical Anthropology, 38: 625-630. . 1974. Das Ausdrucks vernal ten des Javaneraf- fen Macaca fascicularis Raffles 1821. Fortschritte der Verhaltensforschung, 15: 1-91. 1975. Basic data and concepts on the social organization of Macaca fascicularis, pp. 325-388. In Rosenblum, L. A., ed., Primate Behavior: Develop- ments in Field and Laboratory Research, vol. 4. Ac- ademic Press, New York. Angst, W., and D. Thommen. 1977. New data and a discussion of infant killing in Old World monkeys and apes. Folia Primatologica, 27: 198-227. Annandale, N., and H. C. Robinson. 1903. Itinerary in Perak, Selangor, and the Siamese Malay states, pp. i-xlii. In Annandale, N., and H. C. Robinson, eds., Fasciculi Malayensis: Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States, 1 90 1-1 902. Anthropology, Part I. Long- mans, Green & Co., London. Anonymous. 1851. Opsomming der thans bekende zoogdieren van den Indischen Archipel, getrokken uit de Zoology of the Voyage of H. M. Ship Samarang (Lond. 1850). Natuurkundig Tijdschrift voor Neder- landsch Indie, 2: 443-455. . 1931. Photographing wild beasts alive: "Sit- ters" in Bali and Java. Illustrated London News, 88: 457. . 1977. Sundarban Forest— Bangladesh. Tiger- paper, 4(2): 13-15. 1994. Kaser Doo Wildlife Sanctuary, Burma. Asian Primates, 3(3-4): 1 1-12. Arnold, G. 1959. Longhouse and Jungle: An Expe- dition to Sarawak. Chatto and Windus, London, 206 pp. Audebert, J. B. 1798-1799. Histoire Naturelle des Singes et des Makis. Quatrieme Famille, pp. 1-10. Desray, Paris. Auffenberg, W. 1 98 1 . The Behavioral Ecology of the Komodo Monitor. University Presses of Florida, Gainesville, 406 pp. Aziz, F. 1989. Macaca fascicularis (R[a]ffles) from Ngandong, East Java. Publication of the Geological Research and Development Centre, Indonesia, Pale- ontology Series, 5: 50-56. 98 FIELDIANA: ZOOLOGY Azuma, S., A. Suzuki, and Y. Ruhiyat. 1984. The distribution of primates in Sebulu and R. Mahakan. Kyoto University Overseas Research Report of Stud- ies on Asian Non-Human Primates, 3: 45-54. Badoux, D. M. 1959. Fossil mammals from two fis- sure deposits at Punung (Java), with some remarks on migration and evolution of mammals during the Qua- ternary in South East Asia. Thesis, Rijksuniversiteit te Utrecht, Kemink en Zoon, Utrecht, 1 5 1 pp. Bakar, A., M. Amir, and Marshal. 1981. Morpho- logical studies on the crab-eating monkey in Indone- sia. Kyoto University Overseas Research Report of Studies on Indonesian Macaque, 1: 1 1-14. Banks, E. 1931. A popular account of the mammals of Borneo. Journal of the Malayan Branch of the Royal Asiatic Society, 9(2): 1-139. . 1978. Mammals from Borneo. Brunei Museum Journal, 4(2): 165-241. Barbe, P. 1 846. Notice of the Nicobar Islands. Journal of the Asiatic Society of Bengal, Calcutta, 15: 344- 367. Barnicot, N. A., E. R. Huehns, and C. J. Jolly. 1966. Biochemical studies on haemoglobin variants of the irus macaque. Proceedings of the Royal Society of London, ser. B, 165: 224-244. Barnicot, N. A., P. T. Wade, and P. Cohen. 1970. Evidence for a second haemoglobin a-locus duplica- tion in Macaca irus. Nature, 228: 379-381. Bartels, M., Jr. 1929. lets over het voedsel van den panter. Tropische Natuur, 18: 81-83. Beccari, O. 1904. Wanderings in the Great Forests of Borneo: Travels and Researches of a Naturalist in Sa- rawak. Arnold Constable & Co., London, 424 pp. Bellwood, P. 1985. Prehistory of the Indo-Malaysian Archipelago. Academic Press, Sydney, x + 370 pp. Berenstain, L. 1986. Responses of long-tailed ma- caques to drought and fire in eastern Borneo: A pre- liminary report. Biotropica, 18: 257-262. Berkson, G. 1968. Weight and tooth development during the first year in Macaca irus. Laboratory An- imal Care, 18: 352-355. . 1970. Defective infants in a feral monkey group. Folia Primatologica, 12: 284-289. Bernstein, I. S. 1 966. Naturally occurring primate hy- brid. Science, 154: 1559-1560. . 1967. Intertaxa interactions in a Malayan pri- mate community. Folia Primatologica, 7: 198-207. 1968a. Social status of two hybrids in a wild troop of Macaca irus. Folia Primatologica, 8: 121- 131. . 1968b. The lutong of Kuala Selangor. Behav- iour, 32: 1-16. Bismark, M. 1992. Peranan hutan di luar kawasan pelestarian alam dalam konservasi populasi Macaca fascicularis. Bulletin Penelitian Hutan, 549: 9-18. Blatnville, H. M. D. de. [1839]. Osteographie des primates. Sur les primates en general et sur les singes (Pithecus) en particulier, pp. 1-52. In Blainville, H. M. D. de, Osteographie . . . des Mammiferes, vol. 1 and atlas. J. B. Bailliere et Fils, Paris. [For date of publication, see C. D. Sherborn, 1898, Annals and Magazine of Natural History, 7th ser., 2: 76.] Blanford, W. T. [1888a]. Critical notes on the no- menclature of Indian mammals. Proceedings of the Zoological Society of London, 1887: 620-638. [For date of publication, see Duncan, F. M., 1937, Pro- ceedings of the Zoological Society of London, ser. A, 107: 74.] . 1888b. The Fauna of British India, Including Ceylon and Burma. Mammalia. Taylor and Francis, London, xx + 617 pp. [For date of publication, see Preface, p. hi.] Bleeker, P. 1851. Zoogdieren van Banka. Natuurkun- dig Tijdschrift voor Nederlandsch Indie, 2: 527-528. Blyth, E. 1 844. Notices of various mammalia with descriptions of many new species. Part I.— The Pri- mates, Lin. Journal of the Asiatic Society of Bengal, Calcutta, 13: 463-494. . 1846. Notes on the fauna of the Nicobar Is- lands. Journal of the Asiatic Society of Bengal, Cal- cutta, 15: 367-379. . 1847. Supplementary report of the curator of the Zoology Department. Journal of the Asiatic So- ciety of Bengal, Calcutta, 16: 728-737. . 1859. Report of Curator, Zoological Depart- ment, for February to May Meetings, 1859. Journal of the Asiatic Society of Bengal, Calcutta, 28: 27 1- 298. . 1863. Catalogue of the Mammalia in the Mu- seum Asiatic Society. Asiatic Society, Calcutta, 187 + xiii pp. 1875. Catalogue of mammals and birds of Bur- ma. Journal of the Asiatic Society of Bengal, Calcutta, 44(2), extra number: 1-64. Bonhote, J. L. [ 1 90 1 ]. On the mammals collected dur- ing the "Skeat Expedition" to the Malay Peninsula, 1899-1900. Proceedings of the Zoological Society of London, 1900: 869-883. [For date of publication, see Duncan, F. M., 1937, Proceedings of the Zoological Society of London, ser. A, 107: 75.] . 1903. Report on the mammals, pp. 1-45. In Annandale, N., and H. C. Robinson, eds., Fasciculi Malayensis: Anthropological and Zoological Results of an Expedition to Perak and the Siamese Malay States, 1901-1902. Zoology, Part I. Longmans, Green & Co., London. Boonratana, R. 1988. Survey of mammals in South Thailand parks. Natural History Bulletin of the Siam Society, 36:71-84. Bowen, W. H., and G. Koch. 1 970. Determination of age in monkeys (Macaca irus) on the basis of dental development. Laboratory Animals, 4: 1 13-123. Brink, L. M. van den. 1982. On the mammal fauna of the Wajak Cave, Java (Indonesia). Modern Qua- ternary Research in Southeast Asia, 7: 177-193. Brockelman, W. Y. 1981. Field research on primates in Thailand. Journal of the Science Society of Thai- land, 7:9-17. Buffon, G. L. L. de, and L. J. M. Daubenton. 1 766. Histoire Naturelle, Generate et Particuliere, avec la Description du Cabinet du Roi, vol. 14. L'Imprimerie Royal, Paris, 41 1 pp. Burgess, P. F. 1961. Wild life conservation in North Borneo, pp. 143-151. In Wyatt-Smith, J., and P. R. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 99 Wycherley, eds., Nature Conservation in Western Ma- laysia, 1961. Malayan Nature Society, Kuala Lumpur. Burke, D. S., R. R. Graham, and G. B. Heisey. 1981. Hepatitis A virus in primates outside captivity. Lan- cet, 1981(2): 928. Burkhill, H. M. 1961. Protection of wild life on Sin- gapore Island. Malayan Nature Journal, Special Issue: 152-164. Buttikofer, J. 1897. Zoological results of the Dutch Scientific Expedition to Central Borneo. Introduction. Notes from the Leyden Museum, 19: 1-25. Cabrera, A. 1910. On the scientific names of certain primates. Annals and Magazine of Natural History, 8th ser., 6: 617-621. Cadigan, F. C, and Lim Boo Liat. 1975. The future of Southeast Asian nonhuman primates, pp. 83-93. In Bermant, G., and D. G. Lindburg, eds., Primate Utilization and Conservation. John Wiley & Sons, New York. Caldecott, J. O. 1986a. A summary of the ranging and activity patterns of the pig-tailed macaque (Ma- caca n. nemestrina) in relation to those of sympatric primates in peninsular Malaysia, pp. 1 52-1 58. In Taub, D. M., and F. A. King, eds., Current Perspectives in Primate Social Dynamics. Van Nostrand Reinhold, New York. . 1986b. An ecological and behavioural study . 1931. The herpetology of Mt. Kinabalu, North Borneo, 13,455 ft.: Introduction. Bulletin of the Raf- fles Museum, Singapore, 5: 3-7. . 1935a. On a collection of mammals from the of the pig-tailed macaque. Contributions to Prima- tology, 21: i-xiii, 1-259. Cande, S. C, and D. V. Kent. 1992. A new geomag- netic polarity time scale for the Late Cretaceous and Cenozoic. Journal of Geophysical Research, 97(B10): 13,917-13,951. Cannon, C. H., and M. Leighton. 1 994. Comparative locomotor ecology of gibbons and macaques: Selection of canopy elements for crossing gaps. American Jour- nal of Physical Anthropology, 93: 505-524. Cant, J. G. H. 1988. Positional behavior of long- tailed macaques {Macaco, fascicular is) in northern Sumatra. American Journal of Physical Anthropology, 76: 29- 37. Cantor, T. 1846. Catalogue of Mammalia inhabiting the Malayan peninsula and islands. Journal of the Asi- atic Society of Bengal, Calcutta, 15: 171-203. Carpenter, A. 1887. Monkeys opening oysters. Na- ture, 36: 53. Chance, M. R. A., G. R. Emory, and R. G. Payne. 1 977a. Status referents in long- tailed macaques (Ma- caca fascicularis): Precursors and effects of female re- bellion. Primates, 18: 611-632. Chance, M. R. A., E. Jones, and S. Shostak. 1977b. Factors influencing nursing in Macaca fascicularis. Fo- lia Primatologica, 27: 28-40. Chasen, F. N. 1924a. A preliminary account of the mammals of Singapore Island. Singapore Naturalist, 3: 76-86. . 1924b. Notes on the fauna of Pulau Bulan, Rhio Archipelago. Journal of the Malayan Branch of the Royal Asiatic Society, 2: 58-62. 1925. Notes on the fauna of Pulau Galang, Natuna Islands, South China Sea. Bulletin of the Raf- fles Museum, Singapore, 10: 5^42. . 1 935b. On some mammals from the Karimata Islands and Dutch West Borneo. Treubia, 15: 1-7. 1 940a. A handlist of Malaysian mammals. Bul- letin of the Raffles Museum, Singapore, 15: i-xx, 1- 209. . 1940b. The mammals of The Netherlands In- dian Mt. Leuser Expedition 1937 to North Sumatra. Treubia, 17: 479-502. Chasen, F. N., and C. B. Kloss. 1 928a. On a collection of birds from the Anamba Islands, South China Sea. Journal of the Malayan Branch of the Royal Asiatic Society, 6: 43-63. . 1982b. On a collection of mammals from the Anamba Islands, South China Sea. Journal of the Ma- layan Branch of the Royal Asiatic Society, 6: 28-42. . 1931. On a collection of mammals from the Rhio Archipelago. Journal of the Malayan Branch of the Royal Asiatic Society, 3: 92-97. lowlands and islands of North Borneo. Bulletin of the Raffles Museum, Singapore, 6: 1-82. Chaturvedi, Y. 1980. Mammals of the Andamans and Nicobars: Their zoogeography and faunal affinity. Records of the Zoological Survey of India, 77: 127- 139. Cheang, K. C. 1962. Monkeys at Waterfall Gardens, Penang. Malayan Nature Journal, 16: 73. Chesner, C. A., W. I. Rose, A. Deino, R. Drake, and J. A. Westgate. 1991. Eruptive history of Earth's largest Quaternary caldera (Toba, Indonesia) clarified. Geology, 19: 200-203. Chevalier-Skolnikoff, S. 1975. Heterosexual copu- latory patterns in stumptail macaques {Macaca arc- toides) and in other macaque species. Archives of Sex- ual Behavior, 4: 199-220. Chiarelli, A. B. 1972. Taxonomic Atlas of Living Primates. Academic Press, London, 362 pp. Chtno, F., K. Eto, T. Ohkawa, T. Muto, and T. Komatsu. 1992. Different susceptibilities to atten- uated poliovirus of cynomolgus monkeys from the Philippines and other Southeast Asian countries. Jap- anese Journal of Medical Science and Biology, 45: 9- 17. Chtvers, D. J. 1971. The Malayan siamang. Malayan Nature Journal, 24: 78-86. Chivers, D. J., and K. M. Burton. [1991]. Some ob- servations on the primates of Kalimantan Tengah, In- donesia. Primate Conservation, 9: 138-146. [For date of publication, see Asian Primates, 1991, 1: 1.] Chivers, D. J., and G. Da vies. 1979. Abundance of primates in the Krau Game Reserve, peninsular Ma- laysia, pp. 9-36. In Marshall, A. G., ed., The Abun- dance of Animals in Malesian Rain Forests. Depart- ment of Geography, University of Hull, and Institute of South-East Asian Biology, University of Aberdeen, [Hull]. Chivers, D. J., and J. J. Raemaekers. 1980. Long- term changes in behaviour, pp. 209-260. In Chivers, 100 FIELDIANA: ZOOLOGY D. J., ed., Malayan Forest Primates: Ten Years' Study in Tropical Rain Forest. Plenum, New York. Cho, F. 1981. Breeding performance of the cynom- olgus monkey at Tsukuba Primate Center for Medical Science. Japanese Journal of Medical Science & Bi- ology, 34: 252-255. Cho, F., T. Fujtwara, S. Honjo, and K. Imaizumi. 1 973. Sexual maturation of laboratory-bred male cynom- olgus monkeys (Macaca fascicular is). Experimental Animals, Tokyo, 22: 403-409. Chuang, S. H. 1973. Introduction, pp. 1-6. /nChuang, S. H., ed., Animal Life and Nature in Singapore. Sin- gapore University Press, Singapore. Clark, J. A., W. E. Farrell, and W. R. Peltier. 1978. Global changes in postglacial sea level: A numerical calculation. Quaternary Research, 9: 265-287. Collins, W. E., McW. Warren, J. C. Skinner, and D. W. Alling. 1970. Plasmodium inui: Serologic re- lationships of Asian isolates. Experimental Parasitol- ogy, 27: 507-515. Coolidge, H. J., Jr. 1940. Mammal and bird collec- tions of the Asiatic Primate Expedition. Introduction. Bulletin of the Museum of Comparative Zoology at Harvard College, 87: 1 2 1-1 30. Corbet, G. B., and J. E. Hill. 1992. The Mammals of the Indomalayan Region: A Systematic Review. Natural History Museum Publications and Oxford University Press, Oxford, 488 pp. Corlett, R. T., and P. W. Lucas. 1990. Alternative seed-handling strategies in primates: Seed-spitting by long-tailed macaques (Macaca fascicular is). Oecolo- gia, 82: 166-171. Corner, G. W. 1932. The menstrual cycle of the Ma- layan monkey, Macaca irus. Anatomical Record, 52: 401-410. Couper, A. 1989. The Times Atlas and Encyclopedia of the Sea. Times Books, London, 272 pp. Cranbrook, Earl of. 1993. Research and manage- ment of the Batu Apoi Forest Reserve, Temburong, Brunei: The Universiti Brunei Darussalam/Royal Geographical Society Rainforest Project 1991/92. Global Ecology and Biogeography Letters, 3: 267-276. Crawfurd, J. 1828. Journal of an Embassy from the Governor-General of India to the Courts of Siam and Cochin China, Exhibiting a View of the Actual State of Those Kingdoms. Henry Colburn, London, 598 pp. Crockett, C. M., and W. L. Wilson. 1980. The eti- ological separation of Macaca nemestrina and M.fas- cicularis in Sumatra, pp. 148-181. In Lindburg, D. G., ed., The Macaques: Studies in Ecology, Behavior and Evolution. Van Nostrand Reinhold, New York. Crovella, S., M. P. Bigatti, G. Ardito, M. Del Pero, D. Montagnon, and L. Lamberti. 1 994. The high genetic homology of three Macaca fascicularis and two Macaca mulatto subspecies on the basis of their highly repeated DNA restriction patterns. Human Evolution, 9: 63-71. Cuvier, F. 1818. Du macaque de Buffon. Memoires du Museum d'Histoire Naturelle, 4: 109-120. . 1819. Le macaque, liv. 3, pp. 1-4. In Geoffrey Saint-Hilaire, E., and F. Cuvier, eds., Histoire Natu- relle des Mammiferes, vol. 1 . A. Belin, Paris. . 1825. Macaque a face noir, liv. 52, pp. 1-2. In Geoffroy Saint-Hilaire, E., and F. Cuvier, eds., His- toire Naturelle des Mammiferes, vol. 5. A. Belin, Paris. Cuvier, G. 1798. Tableau Elementaire de FHistoire Naturelle des Animaux. Baudouin, Paris, xiv + 710 PP. C. V. Primates (Indonesia). Ca. 1993. Deli Island breeding program [advertising leaflet]. Tangerang, Java, Indonesia, 2 pp. Dahlbom, A. G. 1856. Studia Zoologica . . ., vol. 1, fasc. 1. Berlingianis, Lund, 240 pp. Dammerman, K. W. 1926a. Een toch naar Soemba. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 86:27-122. . 1926b. The fauna of Durian and the Rhio- Lingga Archipelago. Treubia, 8: 281-326. — . 1928. On the mammals of Sumba. Treubia, 10:299-315. 1929. On the zoogeography of Java. Treubia, 11: 1-88. . 1 934. On prehistoric mammals from the Sam- poeng cave, central Java. Treubia, 14: 477-486. 1948. The fauna of Krakatau, 1883-1933. Ve- handlelingen der Koninklijke Nederlandsche Akade- mie van Wetenschappen, Afdeeling Natuurkunde, Tweede Sectie, 44: 1-594. Dampier, W. 1697. A New Voyage Round the World. Argonaut Press, London, 376 pp. (Reprint, 1927.) Dang, D.-C. 1977. Observations recueillies au cours de Felevage artificiel des nouveau-nes de macaque (Macaca fascicularis). Cahiers d' Anthropologic, 4: 1- 36. . 1979. Return of postpartum menstruation and fertility in laboratory Macaca fascicularis. Annales de Biologie Animale, Biochimie, Biophysique, 19: 375- 383. . 1983. La puberte femelle chez le singe Macaca fascicularis eleve en laboratoire: Menarche— copula- tion—gestation— fertilite. Cahiers d' Anthropologic et Biometrie Humaine, 1: 33—45. Dang, D.-C, G. Chaouat, A. Delmas, J. Hureau, G. Hidden, and J. -P. Lassau. [1993]. Assessment of factors responsible for variability of birth weight in the long-tailed macaque (Macaca fascicularis). Folia Primatologica, 59: 149-156. [For date of publication, see front cover of journal.] Darevsky, I. S., and S. Kadarsan. 1964. On the bi- ology of the giant Indonesian monitor ( Varanus ko- modoensis Ouwens). Zoologicheskii Zhurnal, 43: 1 355- 1360. (In Russian, with English summary.) Das, P. K., and D. K. Ghosal. 1977. Notes on the Nicobar crab-eating macaque. Newsletter of the Zoo- logical Survey of India, 3: 264-267. Davtes, G. [1983]. Distribution, abundance and con- servation of simian primates in Borneo, pp. 122-148. In Harper, D., ed., Conservation of Primates and Their Habitats, vol. 1. University of Leicester, Leicester. [For date of publication, see Current Primate Refer- ences, 1985 (May): 9.] Davis, D. D. 1 962. Mammals of the lowland rain- forest of North Borneo. Bulletin of the Singapore Na- tional Museum, 31: 1-129. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 101 de Benedictis, T. 1973. The behavior of young pri- mates during adult copulation: Observations of a Ma- caco, irus colony. American Anthropologist, 75: 1469— 1484. Deignan, H. G. [I960]. Henry Cusheir Raven's travels in eastern Borneo. Sarawak Museum Journal, n.s., 13- 14: 267-269. [For date of publication, see Medway, 1977, p. 154.] Delacour, J. 1928. Quatrieme expedition en Indo- chine. L'Oiseau, 9: 257-269. . 1 929. On the birds collected during the Fourth Expedition to French Indo-China. Ibis, 12th ser., 5: 193-220. Deninger, K. 1910. Ueber einen AfTenkiefer aus den Kendengschichten von Java. Centralblatt fur Miner- alogie, Geologie und Palaontologie, 1910: 1-3. Deputte, B. L., and M. Goustard. 1980. Copulatory vocalizations of female macaques (Macaca fascicu- laris): Variability factors analysis. Primates, 21: 83- 99. de Rossel, E. P. E. 1829. Compte rendu au Ministre de la marine . . . des operations faites pendant la cam- pagne de la Chevrette, commandee par M. Fabre, et des fruits qu'on peut tirer de cette campagne. Annales Maritimes et Coloniales, Paris, II. e Partie, 1: 607-6 1 8. de Rossel, E. P. E., C. L. Mathieu, and D. F. Arago. 1829. Rapport fait a l'Academie des sciences . . . sur les travaux relatifs aux sciences mathematiques qui ont ete executes pendant le voyage de la Chevrette. Annales Maritimes et Coloniales, Paris, II. e Partie, 1: 600-607. Desmarest, A. G. 1817. Macaque, pp. 319-329. In Nouveau Dictionnaire d'Histoire Naturelle, Nouvelle Edition, vol. 18. Deterville, Paris. . 1820. Mammalogie ou Description des Es- peces de Mammiferes, pt. 1 . Agasse, Paris, viii + 276 pp. Dukelow, W. R. 1977. Ovulatory cycle characteristics in Macaca fascicularis. Journal of Medical Primatol- ogy, 6: 33-42. Dukelow, W. R., J. Grauwiler, and S. Bruggemann. 1979. Characteristics of the menstrual cycle in non- human primates. I. Similarities and dissimilarities be- tween Macaca fascicularis and Macaca arctoides. Journal of Medical Primatology, 8: 39^47. Dumond, F. V. 1967. Semi-free-ranging colonies of monkeys at Goulds Monkey Jungle. International Zoo Yearbook, 7: 202-207. Duncan, J. F., P. F. D. Van Peenen, and P. F. Ryan. 1 970. Somatic chromosomes of eight mammals from Con Son Island, South Vietnam. Caryologia, 23: 173- 181. Dunn, F. L. 1975. Rain-forest collectors and traders: A study of resource utilization in modern and ancient Malaya. Monographs of the Malaysian Branch, Royal Asiatic Society, 5: 1-151. Edeson, J. F. B., and D. G. Davey. 1953. Isolation of a virulent strain of Plasmodium knowlesi Sinton and Mulligan, 1932. Transactions of the Royal Society of Tropical Medicine and Hygiene, 47: 259-260. Elbert, J. 1911. Die Sunda-Expedition des Vereins fur Geographie und Statistik zu Frankfurt am Main, vol. 1. Hermann Minjon, Frankfurt am Main, 274 pp. . 1912. Die Sunda-Expedtion des Vereins fur Geographie und Statistik zu Frankfurt am Main, vol. 2. Hermann Minjon, Frankfurt am Main, 373 pp. Elliot, D. G. 1 906. Descriptions of an apparently new species of monkey of the genus Presbytis from Su- matra, and of a bat of the genus Dermanura from Mexico. Proceedings of the Biological Society of Washington, 19: 49-50. . 1 909. Descriptions of apparently new species and subspecies of monkeys of the genera Callicebus, Lagothrix, Papio, Pithecus. Cercopithecus, Erythro- cebus, and Presbytis. Annals and Magazine of Natural History, 8th ser., 4: 244-274. 1910. Descriptions of some new species of monkeys of the genera Pithecus and Pygathrix col- lected by Dr. W. L. Abbott and presented to the United States National Museum. Proceedings of the United States National Museum, 38: 343-352. . 1913. A Review of the Primates, vol. 2. Amer- ican Museum of Natural History, New York, xviii + 382 + xxvi pp. [For date of publication, see Correc- tion, between pp. ii and iii.] Emory, G. R., and S. J. Harris. 1978. On the direc- tional orientation of female presents in Macaca fas- cicularis. Primates, 19: 227-229. Emory, G. R., S. J. Harris, and R. G. Payne. 1980. Sexual solicitation by females in long-tailed macaques {Macaca fascicularis). Biology of Behaviour, 5: 249- 252. Erwtn, J. 1977. Infant mortality in Macaca fascicu- laris: Neonatal and post-neonatal mortality at the Re- gional Primate Research Center Field Station, Uni- versity of Washington, 1967-1976. Theriogenology, 7: 357-366. Erxleben, J. C. P. 1777. Systema Regni Animalis per Classes, Ordines, Genera, Species, Varietates cum Synonymia et Historia Animalium. Classis I. Mam- malia. Impensis Weygandianis, Lipsiae, xlviii + 636 pp. Eudey, A. A. 1979. Differentiation and dispersal of macaques {Macaca spp.) in Asia. Dissertation, De- partment of Anthropology, University of California, Davis, 241 pp. . 1980. Pleistocene glacial phenomena and the evolution of Asian macaques, pp. 52-83. In Lindburg, D. G., ed., The Macaques: Studies in Ecology, Behav- ior and Evolution. Van Nostrand Reinhold, New York. . 1994. Temple and pet primates in Thailand. Revue d'Ecologie (Terre & Vie), 49: 273-280. Eydoux, J. F. T., and L. Souleyet [and P. Gervais]. 1841. Voyage Autour du Monde Execute Pendant les Annees 1836 et 1837 sur la Corvette la Bonite: Zool- ogie, vol. 1. Arthus Bertrand, Paris, xxxix + 328 pp. [For name of third author, see pp. ii, 3.] Eyles, D. E., R. H. Wharton, W. H. Cheong, and McW. Warren. 1964. Studies on malaria and Anopheles balabacensis in Cambodia. Bulletin of the World Health Organization, 30: 7-2 1 . Fa, J. E. 1989. The genus Macaca: A review of tax- onomy and evolution. Mammal Review, 18: 45-81. Fairbanks, R. G. 1989. A 17,000-year glacio-eustatic sea level record: Influence of glacial melting rates on 102 FIELDIANA: ZOOLOGY the Younger Dryas event and deep-ocean circulation. Nature, 342: 637-642. Fischer, J. B. 1829. Synopsis Mammalium. J. G. Cot- ta, Stuttgart, xlii + 527 pp. FlTTINGHOFF, N. A., JR., AND D. G. LlNDBURG. 1980. Riverine refuging in East Bornean Macaca fascicular is, pp. 182-214. In Lindburg, D. G., ed., The Macaques: Studies in Ecology, Behavior and Evolution. Van Nos- trand Reinhold, New York. FrrziNGER, L. J. 1861. DieAusbeutederosterreichisch- en Naturforscher an Saugetherien und Reptilien wah- rend der Weltumsegelung Sr. Majestat Fregatte No- vara. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaf- tliche Classe, 42: 383^16. Fleagle, J. G. 1980. Locomotion and posture, pp. 191-207. In Chivers, D. J., ed., Malayan Forest Pri- mates: Ten Years' Study in Tropical Rain Forest. Ple- num, New York. Flower, S. S. 1900. On the Mammalia of Siam and the Malay Peninsula. Proceedings of the Zoological Society of London, 1900: 306-379. Fooden, J. 1964. Rhesus and crab-eating macaques: Intergradation in Thailand. Science, 143: 363-365. . 1969. Taxonomy and evolution of the mon- keys of Celebes (Primates: Cercopithecidae). Biblio- theca Primatologica, 10: 1-148. — . 1971. Report on primates collected in western Thailand January-April, 1967. Fieldiana: Zoology, 59: 1-62. . 1975. Taxonomy and evolution of liontail and pigtail macaques (Primates: Cercopithecidae). Field- iana: Zoology, 67: 1-168. [ 1 975]. Primates obtained in peninsular Thai- land June-July, 1973, with notes on the distribution of continental Southeast Asian leaf-monkeys (Pres- bytism Primates, 17: 93-1 18. [For date of publication, see outside back cover.] 1976. Provisional classification and key to liv- ing species of macaques (Primates: Macaca). Folia Pri- matologica, 25: 225-236. 1980. Classification and distribution of living macaques (Macaca Lacepede, 1 799), pp. 1-9. In Lind- burg, D. G., ed., The Macaques: Studies in Ecology, Behavior and Evolution. Van Nostrand Reinhold, New York. . 1982. Ecogeographic segregation of macaque species. Primates, 23: 574-579. 1991. Systematic review of Philippine ma- caques (Primates, Cercopithecidae: Macaca fascicu- laris subspp.). Fieldiana: Zoology, n.s., 64: 1-44. — . 1944. Malaria in macaques. International Journal of Primatology, 15: 573-596. Fooden, J., and G. H. Albrecht. 1993. Latitudinal and insular variation of skull size in crab-eating ma- caques (Primates, Cercopithecidae: Macaca fascicu- laris). American Journal of Physical Anthropology, 92: 521-538. Fooden, J., and S. M. Lanyon. 1989. Blood-protein allele frequencies and phylogenetic relationships in Macaca: A review. American Journal of Primatology, 17: 209-241. Forbes, H. O. 1885. A Naturalist's Wanderings in the Eastern Archipelago. Sampson, Low, Marston, Searle & Rivington, London, 536 pp. Frechkop, S. 1934. Notes sur les mammiferes. XVI. Les mammiferes rapportes, en 1932, de Extreme-Ori- ent par S. A. R. le Prince Leopold de Belgique. Bulletin du Musee Royal d'Histoire Naturelle de Belgique, 10: 19-37. Fuchs, S. 1973. The Aboriginal Tribes of India. St. Martin's Press, New York, 308 pp. FujrwARA, N. 1963. Age changes in the skull of the crab-eating monkey. Primates, 4: 53-77. Fujiwara, T., and T. Imamichi. 1966. Breeding of cynomolgus monkeys as an experimental animal. Jap- anese Journal of Medical Science & Biology, 19: 225- 226. Fujiwara, T., I. Uchino, S. Honjo, K. Imaizumi, and T. Imamichi. 1967. Normal range of the menstrual cycle of cynomolgus monkeys under laboratory con- ditions. Japanese Journal of Medical Science & Biol- ogy, 20: 505-507. Furness III, W. H. 1 896. Glimpses of Borneo. Pro- ceedings of the American Philosophical Society, Phil- adelphia, 35: 309-320. Furuya, Y. 196 1-1 962a. On the ecological survey of the wild crab-eating monkeys in Malaya. Primates, 3: 73-76. . 1 96 1-1 962b. The social life of the silvered leaf monkeys Trachypithecus cristatus. Primates, 3: 41-60. . 1962. Studies on the dermatoglyphics of the macaques. Proceedings of the Japan Academy, 38: 377- 386. . 1965. Social organization of the crab-eating monkey. Primates, 6: 285-336. Gairdner, K. G. 1914. Notes on the fauna and flora of Ratburi and Petchaburi districts. Journal of the Nat- ural History Society of Siam, 1: 27-40. Galdikas, B. M. F., and C. P. Yeager. 1984. Croc- odile predation on a crab-eating macaque in Borneo. American Journal of Primatology, 6: 49-5 1 . Garnham, P. C. C. 1963. A new subspecies of Plas- modium knowlesi in the long-tailed macaque. Journal of Tropical Medicine and Hygiene, 66: 156-158. Geoffroy SATNT-HrLAiRE, E. 1803. Catalogue des Mammiferes du Museum National d'Histoire Natu- relle. Paris, 272 pp. . 1812. Tableau des quandrumanes, ou des an- imaux composant le premier ordre de la classe des mammiferes. Annales de Museum d'Histoire Natu- relle, 19: 85-122. Geoffroy Saint-Hilaire, I. 1 826. Macaque, pp. 584- 590. In Bory de Saint- Vincent, ed., Dictionnaire Clas- sique d'Historie Naturelle. Rey et Gravier and Bau- douin Freres, Paris. . [1831]. Mammiferes, pp. 1-160. In Belanger, C, Voyage aux Indes-Orientales .... Arthus Ber- trand, Paris. [For date of publication, see Sherborn, C. D., and B. B. Woodward, 1901, Annals and Mag- azine of Natural History, 7th ser., 7: 390.] . [1843]. Description des mammiferes nouveaux ou imparfaitement connus .... Premier memoire. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 103 Famille des singes. Archives du Museum, 2: 485-592. [For date of publication, see Geoffroy, I., 1851, p. vj.] . 1851. Catalogue Methodique de la Collection des Mammiferes .... Gide et Baudry, Paris, xv + vji + 96 pp. Gervais, P. 1854. Histoire Naturelle des Mammiferes . . . , vol. 1 . L. Curmer, Paris, [420] pp. Ghiglieri, M. P. 1986. A river journey through Gun- ung Leuser National Park, Sumatra. Oryx, 20: 104- 110. Gibson-Hill, C. A. 1 949. Bird and mammal type spec- imens formerly in the Raffles Museum collections. Bulletin of the Raffles Museum, Singapore, 19: 133— 198. Glover, I. 1986. Archaeology of eastern Timor. Terra Australis, 11: i-xxi, 1-241. Goodman, A. L., C. D. Descalzi, D. K. Johnson, and G. D. Hodgen. 1977. Composite pattern of circu- lating LH, FSH, estradiol, and progesterone during the menstrual cycle in cynomolgus monkeys Proceedings of the Society for Experimental Biology and Medicine, 155:479^181. Goustard, M. 1961. La structure sociale d'une colonie de "Macaca irus." Annales des Sciences Naturelles, Zoologie, 12th ser., 3: 297-322. . 1963. Introduction a l'etude de la communi- cation vocale chez Macaca irus. Annales des Sciences Naturelles, Zoologie, 12th ser., 5: 707-747. 1968. La sequence d'activites liees a la copu- Svenska Vetenskapsakademiens Handlingar, 50(8): 1- 76. . 1914. Mammals collected, or observed by the lation chez le macaque crabier Macaca irus. Annales des Sciences Naturelles, Zoologie, 12th ser., 10: 463- 474. Gratton, N. E. 1992. Nicobarese, pp. 208-210. In Hockings, P., ed., Encyclopedia of World Cultures, vol. III. South Asia. G. K. Hall & Co., Boston. Gray, J. E. 1849. Vertebrata, pp. 1-43. In Adams, A., ed., The Zoology of the Voyage of H. M.S. Samarang .... Reeve, Benham, and Reeve, London. . 1870. Catalogue of Monkeys, Lemurs, and Fruit-Eating Bats in the Collection of the British Mu- seum. British Museum, London, 137 pp. Griffith, E. 1821. General and Particular Descrip- tions of the Vertebrated Animals . . . Order Quadru- mana. Baldwin, Cradock, and Joy and Rodwell and Martin, London, 143 pp. Griswold, J. A. 1939a. Up Mount Kinabalu. I. White man and natives begin ascent. Scientific Monthly, 48: 401^14. . 1939b. Up Mount Kinabalu. II. Camping and collecting in Lumu Lumu and beyond. Scientific Monthly, 48: 504-518. Groves, C. P. 1993. Order Primates, pp. 243-277. In Wilson, D. E., and D. M. Reeder, eds., Mammal Spe- cies of the World: A Taxonomic and Geographic Ref- erence, 2nd ed. Smithsonian Institution Press, Wash- ington, D.C. Gurmaya, K. J. 1 989. Ecology, behavior and sociality of Thomas' leaf monkey in North Sumatra. Compar- ative Primatology Monographs, 2: 53-170. Gyldenstolpe, N. 1913. Birds collected by the Swed- ish Zoological Expedition to Siam 191 1-12. Kungliga Swedish Zoological Expedition to Siam 1911-1912. Arkiv for Zoologi, 8(23): 1-36. -. 1916. Zoological results of the Swedish Zoo- logical Expeditions to Siam 191 1-1912 & 1914-1915. IV. Birds II. Kungliga Svenska Vetenskapsakademiens Handlingar, 56(2): 1-160. 1 9 1 7a. On birds and mammals from the Malay Peninsula. Arkiv for Zoologi, 10(26): 1-31. 1 9 1 7b. Zoological results of the Swedish Zoo- logical Expeditions to Siam 191 1—1912 & 1914-1915. V. Mammals II. Kungliga Svenska Vetenskapsaka- demiens Handlingar, 57(2): 1-59. 1919. A list of the mammals at present known to inhabit Siam. Journal of the Natural History Society of Siam, 3: 127-175. . 1920. On a collection of mammals made in eastern and central Borneo by Mr. Carl Lumholtz. Kungliga Svenska Vetenskapsakademiens Handlin- gar, 60(6): 1-62. [For date of publication, see p. 62.] Hadidian, J., and I. S. Bernstein. 1979. Female re- productive cycles and birth data from an Old World monkey colony. Primates, 20: 429-442. Hagen, B. 1 890. Die Pflanzen- und Thierwelt von Deli auf der Ostkiiste Sumatra's: Naturwissenschlaftliche Skizzen und Beitrage. Tijdschrift van het Koninklijk Nederlandsch Aardrijkskundig Genootschap, 2nd ser., 7: 1-240. Halgrimsson, B. 1993. Fluctuating asymmetry in Ma- caca fascicularis: A study of the etiology of develop- mental noise. International Journal of Primatology, 14:421-443. Harihara, S., N. Aoto, M. Hirai, K. Terao, F. Cho, S. Honjo, and K. Omoto. 1986. Polymorphism in the mitochondrial DNA of cynomolgus monkeys. Pri- mates, 27: 357-361. Harihara, S., T. Inanishi, N. Saitou, K. Omoto, P. Varavudhi, and O. Takenaka. 1991. Phylogenetic analysis of Macaca fascicularis in Thailand, using data of mitochondrial DNA, pp. 611-612. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwamoto, eds., Pri- matology Today: Proceedings of the XHIth Congress of the International Primatological Society, Nagoya and Kyoto, 18-24 July 1990. Elsevier Science, Am- sterdam. Harihara, S., N. Saitou, M. Hirai, N. Aoto, K. Terao, F. Cho, S. Honjo, and K Omoto. 1988. Differen- tiation of mitochondrial DNA types in Macaca fas- cicularis. Primates, 29: 117-127. Harrison, J. L. 1961. The natural food of some Ma- layan mammals. Bulletin of the National Museum, State of Singapore, 30: 5-18. . 1969. The abundance and population density of mammals in Malayan lowland forests. Malayan Na- ture Journal, 22: 174-178. Harrison, J. L., and J. R. Hendrickson. 1963. The fauna of the islands of the Straits of Malacca, pp. 543- 555. In Gressitt, J. L., ed., Pacific Basin Biogeography: A Symposium. Bishop Museum Press, Honolulu. Harrisson, T. H. 1933. The Oxford University Ex- 104 FIELDIANA: ZOOLOGY pedition to Sarawak. Geographical Journal, 82: 386- 410. Hartert, E. 1 896. List of a collection of birds made in Lombok by Mr. Alfred Everett. Novitates Zoolo- gicae, 3: 591-599. Hayami, M., K. Ishikawa, A. Komuro, Y. Kawamoto, K. Nozawa, K. Yamamoto, T. Ishida, and Y. Hinuma. 1983. ATLV antibody in cynomolgus monkeys in the wild. Lancet, 1983(2): 620. Hayami, M., A. Komuro, K. Nozawa, T. Shotake, K. Ishikawa, K. Yamamoto, T. Ishida, S. Honjo, and Y. Hinuma. 1984. Prevalence of antibody to adult T-cell leukemia virus-associated antigens (ATLA) in Japanese monkeys and other non-human primates. International Journal of Cancer, 33: 179-183. Heaney, L. R. 1984. Mammalian species richness on islands on the Sunda Shelf, Southeast Asia. Oecologia, 61: 11-17. . 1986. Biogeography of mammals in SE Asia: Estimates of rates of colonization, extinction and spe- ciation. Biological Journal of the Linnaean Society, 28: 127-165. . 1 99 la. A synopsis of climatic and vegetational change in Southeast Asia. Climatic Change, 19: 53- 61. . 1991b. An analysis of patterns of distribution and species richness among Philippine fruit bats (Pter- opidae). Bulletin of the American Museum of Natural History, 206: 145-167. Helfer, J. E. 1838. Notes on animal productions of the Tenasserim Provinces. Journal of the Asiatic So- ciety of Bengal, Calcutta, 7: 855-863. Hellmayr, C. E. 1914. Die Avifauna von Timor, pp. 1-112. In Haniel, C. B., ed., Zoologie von Timor .... I. Lieferung. Kommissionsverlag der E. Schweizer- bartschen Verlagsbuchhandlung, Nagele und Dr. Sproesser, Stuttgart. Heurn, J. W. C. van. 1932. Vluchtige zoologische waarnemingen, gedaam tijdens een kort verblijf op Timor en Flores, Apri-Mei, 1930. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 92: 64-82. Hickson, S. J. 1889. A Naturalist in North Celebes .... John Murray, London, 392 pp. Hill, J. E. 1 960. The Robinson collection of Malaysian mammals. Bulletin of the Raffles Museum, Singapore, 29: 1-112. Hill, W. C. O. 1 974. Primates: Comparative Anatomy and Taxonomy. VII. Cynopithecinae: Cercocebus, Macaca, Cynopithecus. John Wiley & Sons, New York, xxi + 934 pp. Hiller, H. M. 1896. A brief report of a journey up the Rejang River in Borneo. Proceedings of the American Philosophical Society, Philadelphia, 35: 321-328. Hirai, M., K. Terao, F. Cho, and S. Honjo. 1991. Chromosome studies on cynomolgus monkeys (Ma- caca fascicularis), pp. 619-622. In Ehara, A., T. Ki- mura, O. Takenaka, and M. Iwamoto, eds., Prima- tology Today: Proceedings of the XHIth Congress of the International Primatological Society, Nagoya and Kyoto, 18-24 July 1990. Elsevier Science, Amster- dam. Hodgkin, E. P. 1950. The Anopheles umbrosus group (Diptera: Culicidae). Part II: Biology and transmission of malaria. Transactions of the Royal Entomological Society of London, 101: 319-334. Hohmann, G., and W. P. Peter. 1983. Ujung-Kulan- Nationalpark. Zeitschrift des Kolner Zoo, 26(2): 39- 47. . 1986. Samunsam Wildlife Sanctuary— EinRe- servat fur NasenafTen (Nasalis larvatus). Zeitschrift des Kolner Zoo, 29(3): 87-93. Hollister, N. 1912. A list of the mammals of the Philippine Islands, exclusive of the Cetacea. Philip- pine Journal of Science, ser. D, 7: 1-64. . 1913. A review of the Philippine land mam- mals in the United States National Museum. Pro- ceedings of the United States National Museum, 46: 299-341. Honjo, S., F. Cho, and K. Terao. 1984. Establishing the cynomolgus monkey as a laboratory animal. Ad- vances in Veterinary Science and Comparative Med- icine, 28: 51-80. Hoogerwerf, A. 1941. The birds of The Netherlands Indian Mt. Leuser Expedition 1937 to North Sumatra. 3. Itinerary. Treubia, 18(suppl.): 5-8. . 1954. Nature protection in Indonesia. Oryx, 2: 221-227. . 1955. Le lezard geant de Komodo. Science et Nature, 9: 22-27. . 1970. Udjung Kulon: The Land of the Last Javan Rhinoceros. E. J. Brill, Leiden, 512 pp. 1974. Report on a visit to wildlife reserves in East Java, Indonesia (August to November, 1 97 1 ), pp. 1-5 1 . In Nederlandsche Commissie voor Internation- ale Natuurbescherming, Mededelingen No. 21. Aus- terlitz, Holland. Hoogstraal, H. 1951. Philippine Zoological Expe- dition 1946-1947: Narrative and itinerary. Fieldiana: Zoology, 33: 1-86. Hoouer, D. A. 1962a. Prehistoric bone: The gibbons and monkeys of Niah Great Cave. Sarawak Museum Journal, n.s., 11: 428-449. . 1962b. Quaternary langurs and macaques from the Malay Archipelago. Zoologische Verhandelingen Uitgegeven door het Rijksmuseum van Natuurlijke Historie te Leiden, 55: 1-64. . 1964. New records of mammals from the Mid- dle Pleistocene of Sangiran, central Java. Zoologische Mededelingen Uitgegeven door het Rijksmuseum van Natuurlijke Historie te Leiden, 40: 73-88. 1967. Appendix II. Mammalian remains from Liang Toge, Flores, pp. 160-161. In Jacob, T., Some Problems Pertaining to the Racial History of the In- donesian Region. Thesis, Utrecht University, Utrecht. Hoopes, C. 1984. Morphological variation, selection, and the transferrin locus in Macaca fascicularis. Amer- ican Journal of Physical Anthropology, 63: 171. (Ab- stract only.) Hornaday, W. T. 1910. Two Years in the Jungle: The Experiences of a Hunter and Naturalist in India, Cey- lon, the Malay Peninsula and Borneo. Charles Scrib- ner's Sons, New York, 5 1 2 pp. Horsfield, T. [1840]. List of Mammalia and birds FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 105 collected in Assam by John McClelland. Proceedings of the Zoological Society of London, 1839: 146-167. [For date of publication, see Duncan, F. M., 1937, Proceedings of the Zoological Society of London, ser. A, 107: 79.] 1851. A Catalogue of the Mammalia in the tives in Primate Social Dynamics. Van Nostrand Reinhold, New York. -. 1 992. Vertebrate responses to selective logging: Museum of the Hon. East-India Company. Honour- able East-India Company, London, vi + 212 pp. Hose, C. 1893. A Descriptive Account of the Mammals of Borneo. Edward Abbott, London, 78 pp. Hubrecht, A. A. W. [1895]. Embryologisch onder- zoek van zoogdieren uit Nederlandsch-Indie .... Natuurkundig Tijdschrift voor Nederlandsch-Indie, 54: 25-90. [For date of publication, see Ruch, T. C, 1941, Bibliographica Primatologica, Charles C Tho- mas, Springfield, Illinois, p. 16.] Hume, A. O., and W. Davison. 1878. A revised list of the birds of Tenasserim, Stray Feathers, 6: i-viii, 1-524. International Code of Zoological Nomenclature, 3rd ed. 1985. International Trust for Zoological No- menclature, London, xx + 338 pp. International Commission on Zoological Nomenclature. 1986. Opinion 1400. Simia fasci- cularis Raffles, 1821 (Mammalia, Primates): Con- served. Bulletin of Zoological Nomenclature, 43: 229- 230. Ishida, T., and P. Varavudhi. 1 992. Wild long-tailed macaques {Macacafascicularis) in Thailand are highly infected with gamma herpes virus but not with simian T-lymphotropic retrovirus of type 1 . Folia Primato- logica, 59: 163-168. Ishida, T., K. Yamamoto, G. Ishimoto, T. Shotake, O. Takenaka, K. Nozawa, M. Hayami, and Y. Hinuma. 1985. A field study of infection with human T-cell leukemia virus among Asian primates. Micro- biology and Immunology, 29: 839-846. IUCN. 1971. United Nations List of National Parks and Equivalent Reserves, 2nd ed. Hayez, Brussels, 60 1 pp. Jacobson, E. 1913. Simaloer van thans. Tijdschrift van het Koninklijk Nederlandsch Aardrijkskundig Gen- ootschap, 2nd ser., 30: 356-357. . 1917. Het eiland Simaloer. Tijdschrift van het Koninklijk Nederlandsch Aardrijkskundig Genoot- schap, 2nd ser., 34: 265-277. Jentink, F. A. 1897. Zoological results of the Dutch Scientific Expedition to Central Borneo. The mam- mals. Notes from the Leyden Museum, 19: 26-66. Joachimovitz, R. 1928. Studien zu Menstruation, Ovulation, Aufbau und Pathologie des weiblichen Genitales bei Mensch und Affe (Pithecus fascicularis mordax). Biologia Generalis, 4: 447-540. Johns, A. D. 1981. The effects of selective logging on the social structure of resident primates. Malaysian Applied Biology, 10: 221-226. . 1986a. Effects of selective logging on the be- havioral ecology of West Malaysian primates. Ecology, 67:684-694. . 1986b. The effects of commercial logging on a West Malaysian primate community, pp. 206-2 1 1 . In Taub, D. M., and F. A. King, eds., Current Perspec- Implications for the design of logging systems. Philo- sophical Transactions of the Royal Society of London, ser. B, 335: 437^42. Joines, S. 1981. Sarawak's Samunsam Sanctuary. Zoonooz, San Diego, 54(5): 5-9. Jones, F. W. 1910. Corals and Atolls .... Lovell Reeve & Co., London, ix + 392 pp. Jones, M. L. 1982. Longevity of captive mammals. Zoologische Garten, n.s., 52: 1 13-128. Kalra, N. L. 1980. Emergence of malaria zoonosis of simian origin as natural phenomenon in Greater Ni- cobars, Andaman & Nicobar Islands— A preliminary note. Journal of Communicable Diseases, 12: 49-54. Kanagawa, H., and E. S. E. Hafez. 1973. Copulatory behavior in relation to anatomical characteristics of three macaques. American Journal of Physical An- thropology, 38: 233-240. Kanagawa, H., E. S. E. Hafez, M. M. Nawar, and S. Jaszczak. 1972. Patterns of sexual behavior and anatomy of copulatory organs in macaques. Zeitschrift fur Tierpsychologie, 31: 449-460. Karrer, R. 1 970. The use of the tail by an Old World monkey. Primates, 11: 171-175. Karssemeuer, G. J., D. R. Vos, and J. A. R. A. M. van Hooff. 1990. The effect of some non-social factors on mother-infant contact in long-tailed macaques {Macaca fascicularis). Behaviour, 113: 273-291. Kavanagh, M. 1982. Good news about orang-utans in Sarawak. Oryx, 16: 320-321. Kavanagh, M., and E. Laursen. 1984. Breeding sea- sonality among long-tailed macaques, Macaca fasci- cularis, in peninsular Malaysia. International Journal of Primatology, 5: 1 7-29. Kawabe, M., and T. Mano. 1972. Ecology and be- havior of the wild proboscis monkey, Nasalis larvatus (Wurmb), in Sabah, Malaysia. Primates, 13: 213-227. Kawamoto, Y. 1982. A reexamination of electro- morphs of plasma transferrin in the Indonesian crab- eating macaque {Macaca fascicularis). Kyoto Univer- sity Overseas Research Report of Studies on Asian Non-Human Primates, 2: 65-73. Kawamoto, Y., and Tb. M. Ischak. 1981. Genetic differentiation of the Indonesian crab-eating macaque {Macaca fascicularis): I. Preliminary report on blood protein polymorphism. Primates, 22: 237-252. Kawamoto, Y., Tb. M. Ischak, and J. Supriatna. 1982a. Gene constitution of crab-eating macaques {Macacafascicularis) on Lombok and Sumbawa. Kyo- to University Overseas Research Report of Studies on Asian Non-Human Primates, 2: 57-64. . 1984. Genetic variations within and between troops of the crab-eating macaque {Macaca fascicu- laris) on Sumatra, Java, Bali, Lombok and Sumbawa, Indonesia. Primates, 25: 131-139. Kawamoto, Y., T. Ishida, J. Suzuki, O. Takenaka, and P. Varavudhi. 1989. A preliminary report on the genetic variations of crab-eating macaques in Thai- land. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 7: 94-103. 106 FIELDIANA: ZOOLOGY Kawamoto, Y., K. Nozawa, and Tb. M. Ischak. 1981. Genetic variability and differentiation of local popu- lations in the Indonesian crab-eating macaque (Ma- caco fascicular is). Kyoto University Overseas Re- search Report of Studies on Indonesian Macaque, 1: 15-39. Kawamoto, Y., K. Nozawa, Tb. M. Ischak, J. Supri- ATNA, B. SURYOBROTO, AND P. VARAVUDHI. 1991. Evolution and genetic differentiation of the crab-eating macaque, pp. 599-600. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwamoto, eds., Primatology Today: Proceedings of the XHIth Congress of the Interna- tional Primatological Society, Nagoya and Kyoto, 1 8- 24 July 1 990. Elsevier Science, Amsterdam. Kawamoto, Y., K. Nozawa, K. Matsubayashi, and S. Gotoh. 1987. Reports on the crab-eating monkey in Angaur. III. Genetic variation. Kyoto University Overseas Research Report of Studies on Asian Non- Human Primates, 6: 97-102. . 1988. A population-genetic study of crab-eat- ing macaques {Macaca fascicularis) on the island of Angaur, Pulau, Micronesia. Folia Primatologica, 51: 169-181. Kawamoto, Y., T. Shotake, and K. Nozawa. 1982b. Genetic differentiation among three genera of family Cercopithecidae. Primates, 23: 272-286. Kawamoto, Y., andB. Suryobroto. 1985. Gene con- stitution of crab-eating macaques on Timor. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 4: 35-40. Kellogg, R. 1 944. A new macaque from an island off the east cost of Borneo. Proceedings of the Biological Society of Washington, 57: 75-76. . 1945. Macaques, pp. 113-134. In Aberle, S. D., ed., Primate Malaria. National Research Council, Division of Medical Science, Office of Medical Infor- mation, Washington, D.C. Kelsall, H. J. 1894a. Account of a trip up the Pahang, Tembeling, and Tahan rivers, and an attempt to reach Gunong Tahan. Journal of the Straits Branch of the Royal Asiatic Society, 25: 33-49. . 1894b. A journey on the Sembrong River from Kuala Indau to Batu Pahat. Personal account of the journey. List of mammals collected or observed during trip. Journal of the Straits Branch of the Royal Asiatic Society, 26: 1-17. Kemps, A., and P. Timmermans. 1 982. Parturition be- haviour in pluriparous Java-macaques {Macaca fas- cicularis). Primates, 23: 75-88. Kerber, W. T., and W. H. Reese. 1969. Comparison of the menstrual cycle of cynomolgus and rhesus mon- keys. Fertility and Sterility, 20: 975-979. Kern, J. A. 1964. Observations on the habits of the proboscis monkey, Nasalis larvatus (Wurmb), made in the Brunei Bay area, Borneo. Zoologica, 49: 183— 192. Khajuria, H. [1955]. Catalogue of mammals in the Indian Museum (Zool. Surv.). II. Primates: Cercopi- thecidae. Records of the Indian Museum, 52: 101— 127. [For date of publication, see Khajuria, H., 1956, Records of the Indian Museum, 52: 195.] Khan, M. A. R. 1981. The non-human primates of Bangladesh. Tigerpaper, 8(1): 12-15. . 1985. Mammals of Bangladesh. Nazma Reza, Dhaka, 92 pp. Khan, M. A. R., and M. A. Wahab. 1983. Study of eco-ethology of the crabeating macaque, Macaca fas- cicularis in Bangladesh. Journal of the Asiatic Society of Bangladesh (Science), 9: 101-109. Khan, M. K. B. M. 1988. Animal conservation strat- egies, pp. 251-272. In Cranbrook, Earl of, ed., Key Environments: Malaysia. Pergamon, Oxford. King, J. 1784. A Voyage to the Pacific Ocean . . . Per- formed under the Direction of Captains Cook, Clerke, and Gore . . . , vol. 3. Lord Commissioners of the Admiralty, London, 558 pp. Kitchener, D. J., Boeadi, L. Charlton, and Ma- haradatunkamsi. 1990. Wild Mammals of Lom- bok Island: Nusa Tenggara, Indonesia: Systematics and Natural History. Western Australian Museum, Perth, 129 pp. Kloss, C. B. 1903a. In the Andamans and Nicobars .... John Murray, London, xiii + 373 pp. . 1903b. Notes on a cruise in the Southern China Sea. Journal of the Straits Branch of the Royal Asiatic Society, Singapore, 41: 53-80. . 1908. Some visits to Batam Island. Journal of the Straits Branch of the Royal Asiatic Society, Sin- gapore, 50: 61-71. . 1911a. On a collection of mammals and other vertebrates from the Trengganu Archipelago. Journal of the Federated Malay States Museums, 4: 175-212. 1911b. On new mammals from the Trengganu Archipelago. Annals and Magazine of Natural History, 8th ser., 7: 115-119. . 1915a. On two new squirrels from the Inner Gulf of Siam. Journal of the Natural History Society ofSiam, 1: 157-162. . 1915b. On two new rats from the Inner Gulf of Siam. Journal of the Natural History Society of Siam, 1: 221-224. . 1916a. On a collection of mammals from Siam. Journal of the Natural History Society of Siam, 2: 1- 32. . 1 9 1 6b. On a collection of mammals from the coast and islands of Siam. Proceedings of the Zoolog- ical Society of London, 1916: 27-75. . 1917. On a third collection of Siamese mam- mals. Journal of the Natural History Society of Siam, 2:288-318. . 1919a. On a fourth collection of Siamese mam- mals. Journal of the Natural History Society of Siam, 3: 49-56. . 1919b. On birds from South Annam and Co- chin China. Part I. Phasianidae— Campophagidae. Narrative of the journey. Ibis, 1 1th ser., 1: 392-402. . 1919c. On mammals collected in Siam. Journal of the Natural History Society of Siam, 3: 333-407. 1921. The Pulo Condore group and its mam- mals. Journal of the Natural History Society of Siam, 4: 73-83. 1 926. Mammals from Pulo Condore, with de- scriptions of two new subspecies. Journal of the Siam Society, Natural History Supplement, 6: 357-359. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 107 . [1928]. Spolia Mentawiensia.— Mammals. In- troduction. Proceedings of the Zoological Society of London, 1927: 797-807. [For date of publication, see Proceedings of the Zoological Society of London, 1928: 268.] 1930. On mammals from the Raheng district, Western Siam. Introduction. Journal of the Siam So- ciety, Natural History Supplement, 8: 61-63. Koenigswald, G. H. R. von. 1951. Morphology of Solo man. Introduction. Anthropological Papers of the American Museum of Natural History, 43: 21 1-221. Kohlbrugge, J. H. F. 1896a. Bijdragen tot de na- tuurlijke geschiedensis van menschen en dieren. III. Zoogdieren van zuid-oost Borneo. Natuurkunding Tijdschrift voor Nederladsch-Indie, 55: 1 76-200. . 1896b. Bijdragen tot de natuurlijke geschie- densis van menschen en dieren. IV. Zoogdieren van den Tengger. Natuurkundig Tijdschrift voor Neder- ladsch-Indie, 55: 261-298. . 1902. Schadelmasse bei Affen und Halbaffen. Zeitschrift fur Morphologie und Anthropologic, 4: 3 1 8- 344. Kondo, M., Y. Kawamoto, K. Nozawa, K. Matsubayashi, T. Watanabe, O. Griffiths, and M.-A. Stanley. 1991. A report on the genetic vari- ability and constitution of crab-eating macaques (Ma- caca fascicularis) on the island of Mauritius. Kyoto University Overseas Research Report of Studies of Asian Non-Human Primates, 8: 17-25. . 1993. Population genetics of crab-eating ma- caques (Macaca fascicularis) on the island of Mauri- tius. American Journal of Primatology, 29: 167-182. Kool, K M. 1992. The status of endangered primates in Gunung Halimun Reserve, Indonesia. Oryx, 26: 29- 33. Koyama, N. 1984. Socio-ecological study of the crab- eating monkeys at Gunung Meru, Indonesia. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 3: 1 7-36. . 1985. Socio-ecological study of the crab-eating monkeys at Gunung Meru, West Sumatra. Kyoto Uni- versity Overseas Research Report of Studies on Asian Non-Human Primates, 4: 105-126. Koyama, N., A. Asuan, and N. Natsir. 1981. Socio- ecological study of the crab-eating monkeys in Indo- nesia. Kyoto University Overseas Research Report of Studies on Indonesian Macaque, 1: 1-10. Kuenen, P. H. 1 947. Two problems of marine geology: Atolls and canyons. Verhandelingen der Koninklijke Nederlandsche Akademie van Wetenschappen, Af- deeling Natuurkunde, Tweede Sectie, 43(3): 1-69. Kuntz, R. E. 1969. Vertebrates taken for parasitolog- ical studies by U. S. Naval Medical Research Unit No. 2 Expedition to North Borneo (Malaysia). Quarterly Journal of the Taiwan Museum, 22: 191-206. Kurland, J. A. 1973. A natural history of kra ma- caques (Macaca fascicularis Raffles, 1 82 1 ) at the Kutai Reserve, Kalimantan Timur, Indonesia. Primates, 14: 245-262. Kurt,F. 1973. Der Gunung Leuser Survey 1970. Zeit- schrift des Kolner Zoo, 16(2): 59-74. Kyes, R. C. 1993. Survey of the long-tailed macaques introduced onto Tinjil Island, Indonesia. American Journal of Primatology, 31: 77-83. Kyes, R. C, D. Sajuthi, and A. Lelana. 1991. Pre- liminary survey of the cynomolgus macaques on Tinjil Island. American Journal of Primatology, 24: 114. (Abstract only.) Labang, D., and Lord Medway. 1979. Preliminary assessments of the diversity and density of wild mam- mals, man and birds in alluvial forest in the Gunong Mulu National Park, Sarawak, pp. 53-66. In Marshall, A. G, ed., The Abundance of Animals in Malesian Rain Forests. Department of Geography, University of Hull, and Institute of South-East Asian Biology, University of Aberdeen, [Hull]. La Caille, N. L. de. 1 763. Journal Historique du Voy- age Fait au Cap de Bonne-Esperance. Guillyn, Paris, xxxvj + 356 pp. Lacaze, A. de. 1856. Duvaucel (Alfred), columns 534- 539. In Hoefer, M. le Dr., ed., Nouvelle Biographie Generale Depuis les Temps les Plus Recules Jusqu'a Nos Jours . . . , vol. 15. Rosenkilde et Bagger, Copen- hague. (Reprint, 1965.) Lacepede, B. G. E., and F. M. Daudin. [1802]. Tab- leau des divisions, sous-divisions, ordres et genres des mammiferes . . . avec l'indication de toutes les especes decrites par Buffon, et leur distribution dans chacun des genres, pp. 144-195. In Lacepede, B. G. E., ed., Histoire Naturelle par Buffon, vol. 14. P. Didot and Firmin Didot, Paris. [For date of publication, see Hus- son, A. M., and L. B. Holthuis, 1953, Zoologische Mededelingen Uitgegeveven door het Rijksmuseum van Natuurlijke Historie te Leiden, 32: 215.] Lambert, F. 1990. Some notes on fig-eating by arbo- real mammals in Malaysia. Primates, 31: 453-458. Latreille, P. A. 1804. Tableau methodique des singes, pp. 275-298. In Sonnini, C. S., ed., Histoire Naturelle Generale et Particuliere, par Leclerc de Buffon, vol. 36. Dufart, Paris. Laurie, E. M. O., and J. E. Hill. 1954. List of Land Mammals of New Guinea, Celebes and Adjacent Is- lands, 1758-1952. British Museum (Natural History), London, 175 pp. Lawler, S. H., R. W. Sussman, and L. L. Taylor. 1995. Mitochondrial DNA of the Mauritian ma- caques (Macaca fascicularis): An example of the founder effect. American Journal of Physical Anthro- pology, 96: 133-141. Lee Chin Thuan. 1 964. Monkey with a deformed tail. Malayan Nature Journal, 18: 172. Lee, D. W. 1977. Animals of Pulau Tioman, pp. 20- 22. In Lee, D. W., B. C. Stone, M. Ratnasabapathy, and Khoo Teng Tiang, eds., The Natural History of Pulau Tioman. Merlin Samudra Tioman, [?Kuala Lumpur]. Legendre, S. J. 1932. Adventures on hunting trails of Indo-China. Natural History, 32: 481-496. . 1936. Land of the White Parasol and the Mil- lion Elephants: A Journey through the Jungles of Indo- China. Dodd, Mead & Company, New York, 3 1 5 pp. Leighton, M., and D. R. Leighton. 1983. Vertebrate responses to fruiting seasonality within a Bornean rain forest, pp. 181-196. In Sutton, S. L., T. C. Whitmore, and A. C. Chadwick, eds., Tropical Rain Forest: Ecol- 108 FIELDIANA: ZOOLOGY ogy and Management. Blackwell Scientific Publica- tions, Oxford. Lekagul, B., and J. A. McNeely. 1977. Mammals of Thailand. Association for the Conservation of Wild- life, Bangkok, 758 pp. Leon, S., L. Taylor, and R. W. Sussman. 1993. Ac- tivity patterns of captive and free-ranging long-tailed macaques {Macaco, fascicularis). American Associa- tion of Zoological Parks and Aquariums, Regional Conference Proceedings, 1993: 171-178. Leschenault de la Tour, L. T. 1820. Extract d'une lettre de M. Leschenault a M. de Jussieu .... Me- moires du Museum d'Histoire Naturelle, 6: 359-364. . 1822. Relation abregee d'un voyage aux Indes Orientales. Memoires du Museum d'Histoire Natu- relle, 9: 245-274. Lesson, R.-P. 1827. Manuel de Mammalogie, ou His- toire Naturelle des Mammiferes. Roret, Paris, xv + 442 pp. . [1830]. Histoire Naturelle Generate et Parti- culiere des Mammiferes et des Oiseaux Decouverts depuis la Mort de Buffon. Pourrat Freres and Roret, Paris, 564 pp. [For date of publication, see I. Geoffroy, 1851, p. xji.] Lim Boon Hock, and A. Sasekumar. 1979. A prelim- inary study on the feeding biology of mangrove forest primates, Kuala Selangor. Malayan Nature Journal, 33: 105-112. Lindsay, H. M. 1926. Bombay Natural History So- ciety's Mammal Survey of India, Burma and Ceylon. Report No. 39: Mergui Archipelago. Journal of the Bombay Natural History Society, 31: 42-48. Linnaeus, C. 1758. Systema Naturae, vol. 1, 10th ed. Impenisis Direct. Laurentii Salvii, Holm, 824 pp. . 1766. Systema Naturae, vol. 1, 12th ed. Im- penisis Direct. Laurentii Salvii, Holm, 532 pp. . 1771. Mantissa Plantarum Altera Generum danau, and Billiton Islands, between Sumatra and Bor- neo. Proceedings of the United States National Mu- seum, 31: 575-612. . 1907. Mammals collected in western Borneo Editionis VI et Specierum Editionis II. L. Salvii, Holm, 143-588 pp. Lippold, L. K. 1977. The douc langur: A time for conservation, pp. 513-538. In Prince Rainier III and G. H. Bourne, eds., Primate Conservation. Academic Press, New York. Lowe, W. P. 1932. The Trail That Is Always New. Gurney and Jackson, London, 271 pp. . 1933. A report on the birds collected by the Vernay Expedition to Tenasserim and Siam. Ibis, 1 3th ser., 3: 259-283. Lucas, P. W., and R. T. Corlett. 1 99 1 . Relationship between the diet of Macaca fascicularis and forest phe- nology. Folia Primatologica, 57: 201-215. . 1992. Notes on the treatment of palm fruits by long-tailed macaques {Macaca fascicularis). Prin- cipes, 36: 45-48. Luder, H. U. 1993. Hazard rates and causes of death in a captive group of crab-eating monkeys {Macaca fascicularis). American Journal of Primatology, 30: 139-147. Lumer, H.,andA. H. Schultz. 1941. Relative growth of the limb segments and tail in macaques. Human Biology, 13: 283-305. Lyon, M. W., Jr. 1906. Mammals of Banka, Men- by Dr. W. L. Abbott. Proceedings of the United States National Museum, 33: 547-572. . 1908. Mammals collected in eastern Sumatra by Dr. W. L. Abbott during 1903, 1906, and 1907, with descriptions of new species and subspecies. Pro- ceedings of the United States National Museum, 34: 619-679. . 1909. Additional notes on mammals of the Rhio-Linga Archipelago, with descriptions of new spe- cies and a revised list. Proceedings of the United States National Museum, 36: 479-49 1 . . 1911. Mammals collected by Dr. W. L. Abbott on Borneo and some of the small adjacent islands. Proceedings of the United States National Museum, 40: 53-146. . 1916. Mammals collected by Dr. W. L. Abbott on the chain of islands lying off the western coast of Sumatra, with descriptions of twenty-eight new species and subspecies. Proceedings of the United States Na- tional Museum, 52: 437-462. Lyon, M.W., Jr., and W.H.Osgood. 1909. Catalogue of the type-specimens of mammals in the United States National Museum, including the Biological Survey collection. Bulletin of the United States National Mu- seum, 62: i-x, 1-325. Macdonald, D. 1982. Expedition to Borneo: The Search for Proboscis Monkeys and Other Creatures. J. M. Dent & Sons, London, 180 pp. MacDonald, G. J. 1971. Reproductive patterns of three species of macaques. Fertility and Sterility, 22: 373-377. MacKinnon, J. R. 1971. The orang-utan in Sabah today: A study of a wild population in the Ulu Segama Reserve. Oryx, 11: 141-191. . 1973. Orang-utans in Sumatra. Oryx, 12: 234- 242. MacKinnon, J. R., and K. S. MacKinnon. 1978. Comparative feeding ecology of six sympatric pri- mates in West Malaysia, pp. 305-321. In Chivers, D. J., and J. Herbert, eds., Recent Advances in Prima- tology, vol. 1 . Behaviour. Academic Press, London. . 1980. Niche differentiation in a primate com- munity, pp. 167-190. In Chivers, D. J., ed., Malayan Forest Primates: Ten Years' Study in Tropical Rain Forest. Plenum, New York. . 1987. Conservation status of the primates of the Indo-Chinese Subregion. Primate Conservation, 8: 187-195. MacKinnon, K. S. 1986. The conservation status of nonhuman primates in Indonesia, pp. 99-126. In Be- nirschke, K, ed., Primates: The Road to Self-Sustain- ing Populations. Springer- Verlag, New York. Mah,Y.L.,andF.P.G.Aldrich-Blake. 1980. Rang- ing behaviour of Macaca fascicularis (Raffles) in two different habitats in peninsular Malaysia, pp. 353-365. In Furtado, J. I, ed., Tropical Ecology and Develop- ment. International Society of Tropical Ecology, Kua- la Lumpur. Mandal, A. K. 1990. Crab-eating macaque Macaca FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 109 fascicularis (Raffles) feeding on house sparrow Passer domesticus (Linnaeus). Journal of the Bombay Natural History Society, 86: 435. Marsh, C. W., and W. L. Wilson. 1981. Effects of natural habitat differences and disturbance on the abundance of Malaysian primates. Malaysian Applied Biology, 10: 227-249. Marshall, P. 1967. Wild Mammals of Hong Kong. Oxford University Press, Hong Kong, 64 pp. Martens, E. v. 1 876. Die Preussische Expedition nach Ost-Asien. Nach Amtlichen Quellen. Zoologischer Theil, vol. 1 . Koniglichen Geheimen Ober-Hofbuch- druckerei, Berlin, 412 pp. Martin, W. C. L. 1838. A monograph of the genus Semnopithecus. Magazine of Natural History, n.s., 2: 320-326,434-^41. Mason, F. 1851. Tenasserim: Or Notes on the Fauna, Flora, Minerals, and Nations of British Burmah and Pegu .... American Mission Press, Maulmain, 712 pp. Matsubayashi, K., S. Gotoh, Y. Kawamoto, K. Nozawa, and J. Suzuki. 1989. Biological charac- teristics of crab-eating monkeys on Angaur Island. Pri- mate Research (Reichorui Kenkyu), 5: 46-57. Matsubayashi, K., S. Gotoh, Y. Kawamoto, T. Wa- tanabe, K. Nozawa, M. Takasaka, T. Narita, O. Griffiths, and M.-A. Stanley. 1992. Clinical ex- aminations on crab-eating macaques in Mauritius. Primates, 33: 281-288. Matsubayashi, K., S. Gotoh, K. Nozawa, and J. Suzuki. 1987. Reports on the crab-eating monkey in Angaur. I. Circumstances and morphology. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 6: 8 1—89. Matsubayashi, K., and D. Sajuthi. 1981. Microbi- ological and clinical examinations of cynomolgus monkeys in Indonesia. Kyoto University Overseas Research Report of Studies on Indonesian Macaque, 1: 47-56. Matthews, J. 1961. A check-list of "Hoabinhian" sites excavated in Malaya 1860-1939. Papers on Southeast Asian Subjects, 3: 1-59. Mayr, E. 1963. Animal Species and Evolution. Belk- nap Press of Harvard University Press, Cambridge, Massachusetts, 797 pp. McNeely, J. A. 1977. Mammals of the Thai man- groves. Tigerpaper, 4(1): 10-15. Mearns, E. A. 1905. Descriptions of new genera and species of mammals from the Philippine Islands. Pro- ceedings of the United States National Museum, 28: 425^60. Medway, Lord. 1958. Food bone in Niah Cave ex- cavations, (—1958). A preliminary report. Sarawak Museum Journal, 8: 627-636. . 1 966. Observations on the fauna of Pulau Tio- man and Pulau Tulai. 2. The mammals. Bulletin of the National Museum, Singapore, 34: 1 5-32. 1972a. The Gunong Benom Expedition 1967. I. Introduction. Bulletin of the British Museum (Nat- ural History), Zoology, 23: 1-7. 1972b. The Gunong Benom Expedition 1967. and mammals on Gunong Benom. Bulletin of the Brit- ish Museum (Natural History), Zoology, 23: 105-154. 1972c. The Quaternary mammals of Malesia: A review, pp. 63-98. In Ashton, P., and M. Ashton, eds., The Quaternary Era in Malesia. Department of Geography, University of Hull, and Institute of South- East Asian Biology, University of Aberdeen, [Hull]. 1977. Mammals of Borneo. Monographs of the Malaysian Branch of the Royal Asiatic Society, 7: 1-172. Medway, Lord, and D. R. Wells. 1971. Diversity and density of birds and mammals at Kuala Lompat, Pahang. Malayan Nature Journal, 24: 238-247. Megantara, E. N. 1989. Ecology, behavior and so- ciality of Presbytis femoralis in eastcentral Sumatra. Comparative Primatology Monographs, 2: 171-301. Meishvili, N. V., and V. G. Chalyan. 1986. Repro- ductive parameters and sexual behaviour of Java ma- caques (M. fafscjicularis). Vestnik Akademii Medit- sinskikh Nauk SSSR, 3: 14-16. (In Russian, with En- glish abstract.) Melnick, D. J., and G. A. Hoelzer. 1993. What is mtDNA good for in the study of primate evolution? Evolutionary Anthropology, 2: 2-10. Mertens, R. 1930. Die Amphibien und Reptilien der Inseln Bali, Lombok, Sumbawa und Flores. Abhan- dlungen der Senckenbergischen Naturforschenden Ge- sellschaft, 4: 115-344. . 1 936. Die Saugetiere der Inseln Bali, Lombok, Sumbawa und Flores. Zoologische Jahrbuch, Abtei- lung fur Systematik, Okologie und Geographie der Tiere, 68: 273-324. Michael, R. P., and D. Zumpe. 1988. A review of sexual initiating behavior by male and female cynom- olgus monkeys and some species comparisons. Pri- mates, 29: 375-393. Miller, G. S., Jr. 1900. Mammals collected by Dr. W. L. Abbott on islands in the [South] China Sea. Proceedings of the Washington Academy of Sciences, 2: 203-246. . 1901. Mammals collected by Dr. W. L. Abbott on the Natuna Islands. Proceedings of the Washington Academy of Sciences, 2: 1 1 1-138. 1 902a. Mammals collected by Dr. W. L. Ab- bott in the region of the Indragiri River, Sumatra. Proceedings of the Academy of Natural Sciences of Philadelphia, 1902: 143-159. . 1902b. Mammals of the Andaman and Nic- obar islands. Proceedings of the United States Na- tional Museum, 24: 751-795. . 1903a. Mammals collected by Dr. W. L. Ab- bott on the coast and islands of Northwest Sumatra. Proceedings of the United States National Museum, 26: 437^84. . 1903b. Seventy new Malayan mammals. Smithsonian Miscellaneous Collections, 45(1-11): 1- 73. 1 906a. The monkeys of the Macaca nemes- 6. The distribution and altitudinal zonation of birds trina group. Proceedings of the United States National Museum, 29: 555-563. . 1906b. Mammals collected by Dr. W. L. Ab- bott in the Karimata Islands, Dutch East Indies. Pro- 110 FIELDIANA: ZOOLOGY ceedings of the United States National Museum, 31: 55-66. . 1 906c. The mammals collected by Dr. W. L. Abbott in the Rhio-Linga Archipelago. Proceedings of the United States National Museum, 31: 247-286. . 1933. The groups and names of macaques, pp. 1-9. In Hartman, C. G., and W. L. Straus, Jr., eds., The Anatomy of the Rhesus Monkey (Macaca mu- latto). Williams & Wilkins, Baltimore, Maryland. 1942. Zoological results of the George Van- derbilt Sumatran Expedition, 1936-1939. Part V.— Mammals collected by Frederick A. Ulmer on Su- matra and Nias. Proceedings of the Academy of Nat- ural Sciences of Philadelphia, 94: 107-165. Mittermeier, R. A. 1980. Conservation in Brunei. Brunei Museum Journal, 4: 251-261. Mjoberg, E. 1929. Durch die Insel der Kopfjager: Abenteuer im Innern von Borneo. F. A. Brockhaus, Leipzig, 331 pp. . 1930. Forest Life and Adventures in the Malay Archipelago. George Allen & Unwin, London, 20 1 pp. Modigliani, E. 1889. Intorno ai mammiferi dell'isola Nias. Annali del Museo Civico di Storia Naturale di Genova, 2nd ser., 7: 238-245. Moore, J. C, and G. H. H. Tate. 1965. A study of the diurnal squirrels, Sciurinae, of the Indian and In- dochinese Subregions. Fieldiana: Zoology, 48: 1-351. Morice, A. 1875. Coup d'Oeil sur la Faune de la Coch- inchine Francaise. H. Georg, Lyon, 101 pp. . 1 876. Voyage en Cochinchine pendant les An- nees 1872-73-74. H. Georg, Lyon, 44 pp. Morimoto, M. 1982. A study of the cranial length of Catarrhina {Macaca fascicularis, Cercopithecus ae- thiops) by factor analysis. Journal of the Anthropo- logical Society of Nippon, 90: 97-107. (In Japanese, with English summary.) Morris, R. C. 1 936. To Malaya for a rhinoceros. Jour- nal of the Bombay Natural History Society, 38: 438- 446. Moszkowski, M. 1909. Auf neuen Wegen durch Su- matra. Dietrich Reimer (Ernst Vohsen), Berlin, xvii + 328 pp. Mouhot, H. 1864. Travels in the Central Parts of Indo- China (Siam), Cambodia, and Laos, during the Years 1858, 1859, and 1860. John Murray, London, 303 pp. (vol. 1), 301 pp. (vol. 2). Muller, S. [1840]. Over de Zoogdieren van den In- dischen Archipel, pp. 1-57. In Temminck, C. J., ed., Verhandelingen over de Natuurlijke Geschiedenis der Nederlandsche Overzeesche Bezittingen .... Zoolo- gie, Last van den Koning, Leiden. [For date of pub- lication, see Husson, A. M., and L. B. Holthuis, 1955, Zoologische Mededelingen Uitgegeven door het Rijks- museum van Natuurlijke Historie te Leiden, 34: 22.] Musser, G. G. 1981. The giant rat of Flores and its relatives east of Borneo and Bali. Bulletin of the Amer- ican Museum of Natural History, 169: 70-175. Musser, G. G., and D. Califia. 1982. Results of the Archbold Expeditions. No. 106. Identities of rats from Pulau Maratua and other islands off East Borneo. American Museum Novitates, 2726: 1-30. Musser, G. G, and M. D. Carleton. 1993. Family Muridae, pp. 501-755. In Wilson, D. E., and D. M. Reeder, eds., Mammal Species of the World: A Tax- onomic and Geographic Reference, 2nd ed. Smith- sonian Institution Press, Washington, D.C. Nadchatram, M. 1971. Templer Park: A brief ac- count of the natural history of Taman Rimba Templer (Templer Park). Malayan Nature Journal, 24: 139— 147. Nakajima, H., T. Tanaka, H. Nigi, and W. Prychodko. 1970. Human-type ABO, MN, and Lewis blood groups, and Gm and Inv factors in several species of macaques. Primates, 11: 243-253. Nakhasathten, S. 1989. Chiew Larn Dam wildlife res- cue operation. Oryx, 23: 146-154. Nanda, R., R. M. Baume, K. Tanne, and J. Sugawara. 1987. Longitudinal study of craniofacial growth in Macaca fascicularis. American Journal of Physical Anthropology, 73: 215-225. Napier, J. R., and P. H. Napier. 1967. A Handbook of Living Primates: Morphology, Ecology and Behav- iour of Nonhuman Primates. Academic Press, Lon- don, 456 pp. Napier, P. H. 1981. Catalogue of Primates in the Brit- ish Museum (Natural History) and Elsewhere in the British Isles. Part II: Family Cercopithecidae, Subfam- ily Cercopithecinae. British Museum (Natural Histo- ry), London, 203 pp. . 1985. Catalogue of Primates in the British Mu- seum (Natural History) and Elsewhere in the British Isles. Part III: Family Cercopithecidae, Subfamily Co- lobinae. British Museum (Natural History), London, 111pp. Napier, P. H., and C. P. Groves. 1983. Simia fasci- cularis Raffles, 1821 (Mammalia, Primates): Request for suppression under the plenary powers of Simia aygula Linnaeus, 1758, a senior synonym. Bulletin of Zoological Nomenclature, 40: 117-118. Nawar, M. M., and E. S. E. Hafez. 1972. The repro- ductive cycle of the crab-eating macaque (Macaca fas- cicularis). Primates, 13: 43-56. Nieuwenhuisen, J. T., and H. C. B. von Rosenberg. 1863. Verslag omtrent het eiland Nias en deszelfs bewoners. Verhandelingen van het Bataviaasch Gen- ootschap van Kunsten en Wetenschappen, 30: 1-153. Noordwuk, M. A. van. 1985. Sexual behaviour of Sumatran long-tailed macaques (Macaca fascicularis). Zeitschrift fur Tierpsychologie, 70: 277-296. Noordwuk, M. A. van, and C. P. van Schaik. 1985. Male migration and rank acquisition in wild long- tailed macaques (Macaca fascicularis). Animal Be- haviour, 33: 849-861. . 1987. Competition among female long-tailed macaques, Macaca fascicularis. Animal Behaviour, 35: 577-589. . 1988. Male careers in Sumatran long-tailed macaques (Macaca fascicularis). Behaviour, 107: 24- 43. Nordin, M. 1981. Voluntary food intake and digestion in Malaysian primates with special reference to the intake of energy and protein. Malaysian Applied Bi- ology, 10: 163-175. Norikoshi, K. 1984. Sociological study of the crab- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 111 eating monkeys (Macaca fascicularis) on Gunung Meru in Indonesia. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 3: 1-15. Oberholser, H. C. 1917. The birds of the Anamba Islands. Bulletin of the United States National Mu- seum, 98: 1-75. . 1919. The birds of the Tambelan Islands, South China Sea. Proceedings of the United States National Museum, 55: 129-143. Oi, T. 1986. Socio-ecological study on pig-tailed ma- caques {Macaca nemestrina) in Sumatra. Kyoto Uni- versity Overseas Research Report of Studies on Asian Non-Human Primates, 5: 71-78. Okamoto, M. 1994. Annual sperm concentration vari- ation in semen collected by electroejaculation in the cynomolgus monkey (Macaca fascicularis). Experi- mental Animals, Tokyo, 43: 25-3 1 . (In Japanese, with English abstract.) Omoto, K., S. Harada, T. Tanaka, H. Nigi, and W. Prychodko. 1970. Distribution of the electropho- retic variants of serum alpha, -antitrypsin in six species of macaques. Primates, 11: 215-228. Omoto, K., M. Hirai, N. Saitou, S. Harihara, and K. Terao. 1991. Extensive polymorphism of comple- ment C6 among crab-eating monkeys, pp. 603-604. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwa- moto, eds., Primatology Today: Proceedings of the XHIth Congress of the International Primatological Society, Nagoya and Kyoto, 1 8-24 July 1 990. Elsevier Science, Amsterdam. Osbeck, P. 1757. Dagbok ofver en Ostindisk Resa .... Lor. Ludy. Grefing, Stockholm, 376 pp. Osgood, W. H. 1932. Mammals of the Kelley-Roos- evelts and Delacour Asiatic expeditions. Field Mu- seum of Natural History Publication 312, Zoological Series, 18: 193-339. . 1941. Hunting the rare and elusive green pea- cock in the Indo-China jungle. Field Museum News 12(11): 1-2. Ostyn, J. M., J. C. Maltha, and F. P. G. M. van der Linden. 1995. Age assessment in infant crab-eating monkeys (Macaca fascicularis) based on tooth devel- opment. Journal of Zoology, London, 235: 247-252. Pallaud, B. 1984. Consequences d'un changement de hierarchie et de territoire dans un groupe de macaques crabiers (Macaca fascicularis). Biology of Behaviour, 9: 89-99. Parr, J. W. K., N. Mahannop, and V. Charoensiri. 1993. Khao Sam Roi Yot— One of the world's most threatened parks. Oryx, 27: 245-249. Pendlebury, H. M. 1936. An expedition to Korinchi Peak, Sumatra, carried out in 1914 by Messrs H. C. Robinson and C. Boden Kloss. Journal of the Feder- ated Malay States Museums, 8: 1-31. Petiver, J. 1705. De Quadrupedibus Philippensibus tractat. a Reverendo Georg. Jos. Camel. Philosophical Transactions, Royal Society, London, 25: 2197-2204. Pfeffer, P. 1959. Observations sur le varan de Ko- modo, Varanus komodoensis Ouwens 1912. La Terre etla Vie, 106: 195-243. PlRAZZOLI, P. A., U. Radtke. W. S. Hantoro, C. Jouannic, C. T. Hoang, C. Causse, and M. Borel Best. 1991. Quaternary raised coral-reef terraces on Sumba Island, Indonesia. Science, 252: 1834-1836. Plessen, V. von. 1936. Bei den Kopfjagern von Bor- neo. Schiitzen-Verlag, Berlin, 158 pp. Pocock, R. I. 1 906. Notes upon menstruation, gesta- tion, and parturition of some monkeys that have lived in the Society's Gardens. Proceedings of the Zoological Society of London, 1906: 558-570. . 1939. The Fauna of British India, including Ceylon and Burma. Mammalia. —Vol. I. Primates and Carnivora (in part), Families Felidae and Viverridae. Taylor and Francis, London, xxxiii + 463 pp. Poirer, F. E., and E. O. Smith. 1974. The crab-eating macaques (Macaca fascicularis) of Angaur Island, Pa- lau, Micronesia. Folia Primatologica, 22: 258-306. Poole, A. J., and V. S. Schantz. 1942. Catalog of the type specimens of mammals in the United States Na- tional Museum, including the Biological Surveys col- lection. Bulletin of the United States National Mu- seum, 178: i-xiii, 1-705. Price, D. L. 1959. Dirofilaria magnilarvatum n. sp. (Nematoda: Filarioidea) from Macaca irus Cuvier. I. Description of the adult filarial worms. Journal of Par- asitology, 45: 499-504. Rabor, D. S. 1986. Guide to Philippine Flora and Fauna: Birds, Mammals. Natural Resources Manage- ment Center, Ministry of Natural Resources, and Uni- versity of the Philippines, Quezon City, Philippines, 211pp. Raffles, S. 1830. Memoir of the Life and Public Ser- vices of Sir Thomas Stamford Raffles .... John Mur- ray, London, 723 pp. Raffles, T. S. [1821]. Descriptive catalogue of a a zoological collection, made on account of the Hon- ourable East India Company, in the island of Sumatra and its vicinity .... Transactions of the Linnean So- ciety of London, 13: 239-340. [For date of publication, see Horsfield, T., 1821, 'Tapirus Malayanus,' p. 2, in Zoological Researches in Java, and the Neighbouring Islands; Kingsbury, Parbury and Allen, London; cf. Jones, G. S., and D. B. Jones, 1976, A Bibliography of the Land Mammals of Southeast Asia, 1699-1969, Bernice P. Bishop Museum, Honolulu, p. 127.] Ratn, A. N., J. W. Mak, and R. Zamri. 1993. Simian malaria infection in wild caught Macaca fascicularis and Presbytis spp[.] in Malaysia. Southeast Asian Jour- nal of Tropical Medicine and Public Health, 24: 386- 387. Rampino, M. R., and S. Self. 1993. Climate- volca- nism feedback and the Toba eruption of ~ 74,000 years ago. Quaternary Research, 40: 269-280. Raven, H. C. 1935. Wallace's Line and the distribution of Indo-Australian mammals. Bulletin of the Ameri- can Museum of Natural History, 68: 179-283. Ravosa, M. J. 1991. The ontogeny of cranial sexual dimorphism in two Old World monkeys: Macaca fas- cicularis (Cercopithecinae) and Nasalis larvatus (Co- lobinae). International Journal of Primatology, 12: 403- 426. Reichenbach, L. 1862. Die Vollstandigste Naturges- chichte der Affen. Expedition der vollstandigsten Na- turgeschichte, Dresden, 204 pp. 112 FIELDIANA: ZOOLOGY Rensch, B. 1 930. Ein Biologische Reise nach den Klei- nen Sunda-Iseln. Gebriider Borntraeger, Berlin, 236 pp. Rensch, I. 1934. Farn und Barlappe der Sunda- Ex- pedition Rensch. (Unter Einbeziehung einer Aufsa- mmlung G. Steins von Timor.) Hedwigia, 74: 224- 256. RlCHTSMEIER, J. T., J. M. Cheverud, S. E. Danahey, B. D. Corner, and S. Lele. 1993a. Sexual dimorphism of ontogeny in the crab-eating macaque {Macaca fas- cicularis). Journal of Human Evolution, 25: 1-30. RlCHTSMEIER, J. T., B. D. CORNER, H. M. GRAUSZ, J. M. Cheverud, and S. E. Danahey. 1993b. The role of postnatal growth pattern in the production of facial morphology. Systematic Biology, 42: 307-330. Ridley, H.N. 1895. The mammals of the Malay Pen- insula. Natural Science, 6: 23-29. . 1 90 1 . Habits of the drongo. Journal of the Straits Branch of the Royal Asiatic Society, 35: 105. — . 1906. The Menagerie at the Botanic Gardens. Journal of the Straits Branch of the Royal Asiatic So- ciety, 46: 133-194. Rtjksen, H. D. 1978. A fieldstudy on Sumatran orang utans (Pongo pygmaeus abeli Lesson 1827): Ecology, behaviour and conservation. Mededelingen Land- bouwhogeschool Wageningen, Nederland, 78(2): 1- 420. Riley, J. H. 1930. Birds from the small islands off the northeast coast of Dutch Borneo. Proceedings of the United States National Museum, 77(12): 1-23. . 1938. Birds from Siam and the Malay Pen- insula in the United States National Museum collected by Drs. Hugh M. Smith and William L. Abbott. Bul- letin of the United States National Museum, 172: 1- 581. Robinson, A. H., and Y. Rustandi. 1982. Bali Barat National Park. Parks, 7(2): 13-14. Robinson, H. C. 1915. The zoology of Koh Samui and Koh Pennan. I. Introduction. Journal of the Federated Malay States Museums, 5: 128-129. . 1916. A collection of mammals and birds from Pulau Panjang or Pulau Mapor, Rhio-Lingga Archi- pelago. Journal of the Federated Malay States Muse- ums, 7: 59-72. . 1917. On a collection of birds from Pulau Langkawi and other islands on the north-west coast of the Malay Peninsula. Journal of the Federated Ma- lay States Museums, 7: 129-191. . 1919. Notes on the vertebrate fauna of the Pahang-Johore Archipelago. Journal of the Federated Malay States Museums, 7: 325-329. — . 1921. The birds of South- West and Peninsular Introduction. Journal of the Natural History 15. Siam Society of Siam, 5: 1 Robinson, H. C, and C. B. Kloss. 1910. On birds from the northern portion of the Malay Peninsula, including the islands of Langkawi and Terutau; with notes on other rare Malayan species from the southern districts. Ibis, 9th ser., 4: 659-675. . 1914. Some remarks on Dr. D. G. Elliot's 'Re- . 1915a. The zoology of Koh Samui and Koh Pennan. II. Mammals. Journal of the Federated Malay States Museums, 5: 130-139. . 1915b. List of a small collection of mammals and birds from the Krau River, western Pahang. Jour- nal of the Federated Malay States Museums, 5: 169— 175. . 1918. Mammals of Korinchi. Journal of the Federated Malay States Museums, 8: 1-72. 1919. On a collection of birds from the prov- ince of Puket, peninsular Siam. Journal of the Natural History Society of Siam, 3: 87-119. Rode, P. 1 938. Catalogue des types de mammiferes du Museum National d'Histoire Naturelle. I. Ordre des Primates. A.— Sous-ordre des simiens. Bulletin du Museum National d'Histoire Naturelle, Paris, 2nd ser., 10:202-251. Rodman, P. S. 1973. Synecology of Bornean primates. I. A test for interspecific interactions in spatial distri- bution of five species. American Journal of Physical Anthropology, 38: 655-659. . 1978. Diets, densities, and distributions of Bornean primates, pp. 465-478. In Montgomery, G., ed., The Ecology of Arboreal Folivores. Smithsonian Institution Press, Washington, D.C. . 1991. Structural differentiation of microhabi- view of the Primates.' Annals and Magazine of Natural History, 8th ser., 13: 389-399. tats of sympatric Macaca fascicularis and M. nemes- trina in East Kalimantan, Indonesia. International Journal of Primatology, 12: 357-375. ROEDELBERGER, F. A., AND V. I. GROSCHOFF. 1967. Wildlife of the South Seas. Constable, London, 216 pp. Rosenberg, H. von. 1882. Die Affen von Insulinde. Zoologische Garten, 23: 1 1 1-1 15. Rothchild, G. 1971. Animals in Bako National Park. Malayan Nature Journal, 24: 163-169. Rothchild, W. 1894. First glimpses of the zoology of the Natuna Islands. I. Introduction. Novitates Zool- ogicae, 1: 467-468. Rubio, R. P., P. L. Alviola III, and M. R. Felizardo. 1990. CITES Study A: Biological and Economic Im- plications in the Trade and Commercial Exploitation of the Philippine Long-Tailed Macaques. World Wild- life Fund (USA), Haribon Foundation for the Con- servation of Natural Resources, and Department of Environment and Natural Resources, [?Manila], 135 pp. Ruhiyat, Y. 1986. Preliminary study of proboscis monkey {Nasalis larvatus) in Gulung Palung Nature Reserve, West Kalimantan. Kyoto University Over- seas Research Report of Studies on Asian Non-Human Primates, 5: 59-69. Ruiter, J. R. de. 1 993. Capturing wild long-tailed ma- caques {Macaca fascicularis). Folia Primatologica, 59: 89-104. Ruiter, J. R. de, and J. A. R. A. M van Hooff. 1 993. Male dominance rank and reproductive success in pri- mate groups. Primates, 34: 5 1 3-523. Ruiter, J. R. de, J. A. R. A. M van Hooff, and W. Scheffrahn. 1994. Social and genetic aspects of pa- ternity in wild long-tailed macaques {Macaca fasci- cularis). Behaviour, 129: 203-224. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 113 RUITER, J. R. DE, W. SCHEFFRAHN, G. J. J. M. TROM- MELEN, A. G. UlTTERLINDEN, R. D. MARTIN, AND J. A. R. A. M van Hooff. 1992. Male social rank and reproductive success in wild long-tailed macaques, pp. 175-191. In Martin, R. D., A. F. Dixson, and E. J. Wickings, eds., Paternity in Primates: Genetic Tests and Theories. Implications of Human DNA Finger- printing. Karger, Basel. Salter, R. E., and K. M. Aken. 1983. The proboscis monkey in Bako National Park, Sarawak. Tigerpaper, 10(3): 6-8. Samilchuk, E., and S. Sarsaniya. 1 994. Genetic poly- morphism of alpha- 1 -antitrypsin in macaque species. Primates, 35: 353-360. Sarker, S. U., and N. J. Sarker. 1984. Mammals of Bangladesh— Their status, distribution and habitat. Tigerpaper, 11(1): 8-13. Satmoko, R. K. P. 1 96 1 . Udjung-Kulon Nature Park, Java, pp. 107-124. In Wyatt-Smith, J., and P. R. Wycherley, eds., Nature Conservation in Western Ma- laysia, 1961. Malayan Nature Society, Kuala Lumpur. Schaik, C. P. van, and T. Mitrasetia. 1990. Changes in the behaviour of wild long-tailed macaques (Ma- caca fascicularis) after encounters with a model py- thon. Folia Primatologica, 55: 104-108. Schaik, C. P. van, W. J. Netto, A. J. J. van Ame- rongen, and H. Westland. 1989. Social rank and sex ratio of captive long- tailed macaque females (Ma- caco, fascicularis). American Journal of Primatology, 19: 147-161. Schaik, C. P. van, and M. A. van Noordwuk. 1 985a. Evolutionary effect of the absence of felids on the so- cial organization of macaques on the island of Simeu- lue (Macaca fascicularis fusca, Miller, 1 903). Folia Pri- matologica, 44: 138-147. . 1985b. Interannual variability in fruit abun- dance and the reproductive seasonality in Sumatran long-tailed macaques (Macaca fascicularis). Journal of Zoology, London, ser. A, 206: 533-549. 1 986. The hidden costs of sociality: Intra-group ceedings of the Academy of Natural Sciences of Phil- adelphia, 91: 311-368. 1 940b. Zoological results of the George Van- variation in feeding strategies in Sumatran long-tailed macaques (Macaca fascicularis). Behaviour, 99: 296- 315. . 1988. Scramble and contest in feeding com- petition among female long-tailed macaques (Macaca fascicularis). Behaviour, 105: 77-98. Schaik, C. P. van, M. A. van Noordwuk, R. J. de Boer, and I. den Tonkelaar. 1983a. The effect of group size on time budgets and social behaviour in wild long- tailed macaques (Macaca fascicularis). Behavioral Ecology and Sociobiology, 13: 173-181. Schaik, C. P. van, M. A. van Noordwuk, T. van Bragt, and M. A. Blankenstein. 1991. A pilot study of the social correlates of levels of urinary Cortisol, prolactin, and testosterone in wild long-tailed macaques (Ma- caca fascicularis). Primates, 32: 345-356. Schaik, C. P. van, M. A. van Noordwuk, B. Warsono, and E. Sutriono. 1983b. Party size and early de- tection of predators in Sumatran forest primates. Pri- mates, 24: 211-221. Schauensee, R. M. de, and S. D. Ripley. 1940a. Zoo- logical results of the George Vanderbilt Sumatran Ex- pedition, 1936-1939. Part I.-Birds from Atjeh. Pro- derbilt Sumatran Expedition, 1936-1939. Part III.— Birds from Nias Island. Proceedings of the Academy of Natural Sciences of Philadelphia, 91: 399^13. Scheffrahn, W., J. R. DE RUITER, AND J. A. R. A. M van Hooff. 1994. Genetic relatedness within and between populations of Macaca fascicularis on Su- matra and off-shore islands, pp. 131-152. In Ruiter, J. de, ed., Behaviour and genes in natural populations of long- tailed macaques (Macaca fascicularis). Thesis, Faculty of Biology, Utrecht University, Utrecht. Schinz, H. R. 1821. Das Thierreich . . . , vol. 1. J. G. Cotta'schen Buchhandlung, Stuttgart, xxxviii + 894 pp. . 1825. Das Thierreich ..., vol. 4. J. G. Cott- a'schen Buchhandlung, Stuttgart, xiii + 793 pp. Schlegel, H. 1876. Monographic 40: Simiae. Revue Methodique, Museum d'Histoire Naturelle des Pays- Bas, 7: 1-356. Schmitt, J., D. Graur, and J. Tomiuk. 1990. Phy- logenetic relationships and rates of evolution in pri- mates: Allozymic data from catarrhine and platyrrhine species. Primates, 31: 95-108. Schneider, G. 1905. Ergebnisse zoologischer For- schungsreisen in Sumatra. Erster Teil. Saugetiere (Mammalia). Zoologische Jarbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 23: 1- 172. Schreber, J. C. D. [1774]. Die Saugthiere in Abbil- dungen nach der Nature mit Beschreibungen. Erster Theil. Wolfgang Walther, Erlangen, 190 pp. [For date of publication, see Sherborn, C. D., 1892, Proceedings of the Zoological Society of London, 1891: 588.] Schwarz, E. 1912. Die Saugetierausbeute der Sunda- Expedition, pp. 303-304. In Elbert, J., ed., Die Sunda- Expedition des Vereins fur Geographie und Statistik zu Frankfurt am Main, vol. 2. Hermann Minjon, Frankfurt am Main. . 1913. Two new mammals from the Malay Ar- chipelago. Annals and Magazine of Natural History, 8th ser., 11: 296-298. 1914. Saugetiere von Timor, pp. 1 13-135. In Haniel, C. B., ed., Zoologie von Timor ... II. Liefer- ung. Kommissionsverlag der E. Schweizerbartschen Verlagsbuchhandlung, Nagele und Dr. Sproesser, Stuttgart. Scopoli, I. A. 1786. Deliciae Florae et Faunae Insub- ricae Seu Novae, aut Minus Cognitae Species quas in Insubria Austriaca Tarn Spontaneas, quam Exoticas Vidit, Descripsit, et Aeri Incidi Curavit. Monasterii S. Salvatoris, Ticino, 87 pp. Scott, G. G. 1982. South-East Asian macaques: A study of skin and skull variation. M.A. Thesis, Aus- tralian National University, Canberra, 234 pp. Seidensticker, J. 1983. Predation by Panthera cats and measures of human influence in habitats of South Asian monkeys. International Journal of Primatology, 4: 323-326. Shebbeare, E. O. 1940. Malayan mammals. Malayan Nature Journal, 1: 54-59. 114 FIELDIANA: ZOOLOGY Shelford, R. W. C. 1916. A Naturalist in Borneo. T. Fisher Unwin, London, 33 1 pp. Shimizu, T., H. Narita, F. Ohkubo, T. Yoshida, F. Cho, and Y. Yoshikawa. 1994. A feeding experi- ment on laboratory-bred male cynomolgus monkeys. I. Morphometric study. Experimental Animals, To- kyo, 43: 173-180. (In Japanese, with English sum- mary.) Shively, C, S. Clarke, N. King, S. Schapiro, and G. Mitchell. 1982. Patterns of sexual behavior in male macaques. American Journal of Primatology, 2: 373- 384. Shively, C, and D. G. Smith. 1985. Social status and reproductive success of male Macaca fascicularis. American Journal of Primatology, 9: 129-135. Sinha, S., and A. Gajanana. 1984. First report of natural infection with quartan malaria parasite Plas- modium shortti in Macaca fascicularis umbrosa (=irus) of Nicobar Islands. Transactions of the Royal Society of Tropical Medicine and Hygiene, 78: 567. Slack, J. H. 1 867. Mammalogical notices. Proceedings of the Academy of Natural Sciences of Philadelphia, 19: 34-38. Small, M. F. 1 994. Macaque see, macaque do. Natural History, 103(3): 8-11. Smith, B. H., T. L. Crummett, and K. L. Brandt. 1994. Ages of eruption of primate teeth: A compen- dium for aging individuals and comparing life histo- ries. Yearbook of Physical Anthropology, 37: 177— 231. Smith, D. G., and R. E. Ferrell. 1 980. A family study of hemoglobin polymorphism in Macaca fascicularis. Journal of Human Evolution, 9: 557-563. Smith, E. G. W. 1974. Handbook of Marine Science, vol. 1. CRC Press, Cleveland, 627 pp. Snelleman, J. F. 1887. Zoogieren en vogels, pp. 1-58. In Veth, P. J., ed., Midden-Sumatra. Reizen en On- derzoekingen der Sumatra-Expeditie, Uitgerust door het Aardrijkskundig Genootschap, 1877-1879, vol. 4, pt. 1. E. J. Brill, Leiden. Socha, W. W., and J. Ruffie. 1983. Blood Groups of Primates: Theory, Practice, Evolutionary Meaning. Alan R. Liss, New York, 266 pp. Sody, H. J. V. 1929. Naamlijst van de zoogdieren van Java (met korte beschrijvingen van twee niuwe sub- species). Natuurkundig Tijdschrift voor Neder- landsch-Indie, 89: 160-166. . 1932. Four new mammals from Bali and Soemba. Natuurkundig Tijdschrift voor Neder- landsch-Indie, 92: 334-340. . 1933. On the mammals of Bali (with a note on the races of Callosciurus notatus of Java). Natu- urkundig Tijdschrift voor Nederlandsch-Indie, 93: 56- 95. . 1937. On the mammals of Banka. Temminck- ia, 2: 221-250. F. Aziz, J. de Vos, and U. L. Batu. 1994. Middle Pleistocene faunal turnover and colonization of Flores (Indonesia) by Homo erectus. Comptes Rendus de l'A- cademie des Sciences, Sciences de la Terre et des Pla- netes, ser. II, 319: 1255-1262. Southwick, C. H., and F. C. Cadigan, Jr. 1 972. Pop- ulation studies of Malaysian primates. Primates, 13: 1-18. Southwick, C. H., and D. Manry. 1987. Habitat and population changes for the Kowloon macaques. Pri- mate Conservation, 8: 48-49. Southwick, C. H., and B. Rosenbaum. 1992. The primate community of Pulau Kaget, Kalimantan. Ab- stracts, XlVth Congress of the International Prima- tological Society, p. 88. (Abstract only.) Southwick, C. H., and K. L. Southwick. 1983. Poly- specific groups of macaques on the Kowloon Penin- sula, New Territories, Hong Kong. American Journal of Primatology, 5: 17-24. Spencer, C. 1975. Interband relations, leadership be- haviour and the initiation of human-oriented behav- iour in bands of semi-wild free ranging Macaca fas- cicularis. Malayan Nature Journal, 29: 83-89. Spiegel, A. 1952. Weitere Beobachtungen fiber den zeitlichen Ablauf der Bezahnung u. des Zhanwechels bei Javamakaken. Zeitschrift fur Saugetierkunde, 18: 123-135. . 1954. Beobachtungen und Untersuchungen an Javamakaken. Zoologische Garten, n.s., 20: 227-270. — . 1985. Postnatal growth of Macaca fascicularis . 1 949. Notes on some primates, carnivora, and the babirusa from the Indo-Malayan and Indo-Aus- tralian Regions (with descriptions of 10 new species and subspecies). Treubia, 20: 121-190. Sondaar, P. Y., G. D. van den Bergh, B. Mubroto, born in captivity. Primate Report, 13: 1-55. Steenis-Kruseman, M. J. van. 1950. Malaysian plant collectors and collections, being a cyclopedia of bo- tanical exploration in Malaysia and a guide to the concerned literature up to the year 1950. Flora Ma- lesiana, 1st ser., 1: lxxvi-clii, 1-639. Stewart, T. D. 1933. The skin and its appendages, pp. 28-35. In Hartman, C. G., and W. L. Straus, Jr., eds., The Anatomy of the Rhesus Monkey (Macaca mulatto). Williams & Wilkins, Baltimore, Maryland. Stiles, C. W., and M. O. Nolan. 1929. Key catalogue of primates for which parasites are reported. U.S. Hy- gienic Laboratory Bulletin, 152: 409-601. Storer, P. J. 1978. A biological survey of a lowland evergreen scrub forest and meadowland in southern Thailand. Natural History Bulletin of the Siam Soci- ety, 27: 93-1 14. . 1979. A preliminary biological survey of Khao Khieo Wildlife Sanctuary. Natural History Bulletin of the Siam Society, 28: 25-46. Stott, K, Jr. 1964. Lungmanis: Peace in a forest pri- meval. Zoonooz, San Diego, 37(10): 10-15. Stremme, H. 1911. Die Saugetiere mit Ausnahme der Proboscidier, pp. 82-150. In Selenka, M. L., and M. Blanckenhorn, eds., Die Pithecanthropus-Schichten auf Java .... Wilhelm Engelmann, Leipzig. Stresemann, E. 1938. Vogel vom Fluss Kajan (Nor- dost-Borneo) gesammelt von Baron Victor von Pies- sen. Temminckia, 3: 109-136. Sugardjito, J., C. P. van Schaik, M. A. van Noord- wuk, and T. Mitrasetia. 1989. Population status FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 115 of the Simeulue monkey {Macaca fascicularis fused). American Journal of Primatology, 17: 197-207. Sumardja, E. A. 1981. First five national parks in Indonesia. Parks, 6(2): 1-4. SUPRIATNA, J., B. O. MANULLANG, AND E. SOEKARA. 1986. Group composition, home range, and diet of the maroon leaf monkey (Presbytis rubicunda) at Tan- jung Puting Reserve, central Kalimantan, Indonesia. Primates, 27: 185-190. Sussman, R. W., and I. Tattersall. 1981. Behavior and ecology of Macaca fascicularis in Mauritius: A preliminary study. Primates, 22: 192-205. . 1986. Distribution, abundance, and putative ecological strategy of Macaca fascicularis on the island of Mauritius, southwestern Indian Ocean. Folia Pri- matologica, 46: 28-43. Suzuki, A. 1986. The ecological survey on the effects of the forest fires and droughts in 1982-83, and the distributions and populations of primates along the middle-upper streams of Sungai Sengata in Kutai Na- tional Park, East Kalimantan, Indonesia. Kyoto Uni- versity Overseas Research Report of Studies on Asian Non-Human Primates, 5: 13-22. . 1989. Socio-ecological studies of orangutans and primates in Kutai National Park, East Kalimantan in 1988-89. Kyoto University Overseas Research Re- port of Studies on Asian Non-Human Primates, 7: 1- 42. . 1991. Forest fires' effects on the population of primates in Kutai National Park, East Kalimantan, Indonesia, pp. 51-54. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwamoto, eds., Primatology Today: Proceedings of the XHIth Congress of the Interna- tional Primatological Society, Nagoya and Kyoto, 1 8- 24 July 1 990. Elsevier Science, Amsterdam. Suzuki, J., and P. Varavudhi. 1989. Somatometric data of crab-eating macaques in Thailand. Kyoto Uni- versity Overseas Research Report of Studies on Asian Non-Human Primates, 7: 110-116. Suzuki, M. T., T. Ono, M. Kohno, H. Ogawa, and F. Cho. 1990. Hour of delivery in cynomolgus mon- keys under indoor individually-caged conditions. Pri- mates, 31: 251-255. Swisher III, C. C. 1994. Response to J. de Vos and P. Sondaar, Dating hominid sites in Indonesia. Science, 266: 1727. Swisher III, C. C, G. H. Curtis, T. Jacob, A. G. Getty, A. Supruo, and Widiasmoro. 1 994. Age of the ear- liest known hominids in Java. Science, 263: 111 8— 1121. Takenaka, A., and O. Takenaka. 1991. Multiplica- tion of a-globin genes in higher non-human primates, pp. 631-632. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwamoto, eds., Primatology Today: Proceed- ings of the Xlllth Congress of the International Pri- matological Society, Nagoya and Kyoto, 18-24 July 1990. Elsevier Science, Amsterdam. Takenaka, A., S. Ueda, K. Terao, and O. Takenaka. 1991. Multiple a-globin genes in crab-eating ma- caques (.Macaca fascicularis). Molecular Biology and Evolution, 8: 320-326. Takenaka, O., A. Takenaka, M. Arakawa, T. Ishida, J. Suzuki, Y. Kawamoto, and P. Varavudhi. 1 989. The multiple a-globin genes in the crab-eating ma- caques {Macaca fascicularis) and geographical distri- bution in Thailand. Kyoto University Overseas Re- search Report of Studies on Asian Non-Human Pri- mates, 7: 81-93. Tanaka, H. 1991. Distribution of the a, -antitrypsin (a, -AT) variants detected with isoelectric focusing in crab-eating macaques {Macaca fascicularis). Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 8: 43-50. Tanaka, H., Y. Kawamoto, T. Ishida, J. Suzuki, O. Takenaka, and P. Varavudhi. 1989. Polymor- phism of the vitamin D binding protein (DBP) in Thai- land crab-eating macaques {Macaca fascicularis). Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 7: 104-109. Tanaka, H., Y. Kawamoto, and K. Terao. 1991. Ge- netic polymorphism of the vitamin D-binding protein (DBP) in crab-eating macaques {Macaca fascicularis). Journal of Medical Primatology, 20: 126-132. Tangpraprutigul, P., and P. Varavudhi. 1982. Lack of breeding seasonality in the Macaca fasciciularis studied in Bangkok environment. Journal of Steroid Biochemistry, 17: xci. (Abstract only.) Taub, D. M., and M. G. Bond. 1982. Differences among Malayan and Philippine M. fascicularis in se- rum lipid responses. International Journal of Prima- tology, 3: 339. (Abstract only.) Terao, K. 1985. Genetic control of the cynomolgus monkey breeding colony by the use of blood groups. Japanese Journal of Medical Science & Biology, 38: 49-52. Terao, K, K. Fujimoto, F. Cho, and S. Honjo. 1981. Inheritance and distribution of human-type A-B-O blood groups in cynomolgus monkeys. Journal of Medical Primatology, 10: 72-80. THEUNISSEN, B., J. DE VOS, P. Y. SONDAR, AND F. AZIZ. 1 990. The establishment of a chronological frame- work for the hominid-bearing deposits of Java; a his- torical survey. Geological Society of America, Special Paper, 242: 39-54. Thomas, O. 1886. On the mammals presented by Allan O. Hume, Esq., C. B., to the Natural History Museum. Proceedings of the Zoological Society of London, 1886: 54-79. . 1 906. Mammals, pp. 3-66. In The History of the Collections Contained in the Natural History De- partments of the British Museum, vol. 2. British Mu- seum, London. -. 1923. On some mammals from Simalur Island, west of Sumatra, collected by Mr. E. Jacobson. Annals and Magazine of Natural History, 9th ser., 12: 591- 593. 1928. The Delacour Exploration of French Indo-China.— Mammals. III. Mammals collected dur- ing the Winter of 1927-28. Proceedings of the Zoo- logical Society of London, 1928: 831-841. Thomas, O., and E. Hartert. 1 894. First glimpses of the zoology of the Natuna Islands. III. List of the first collection of mammals from the Natuna Islands. Nov- itates Zoologicae, 1: 652-660. Thomas, O., and R. C. Wroughton. 1909a. On a collection of mammals from western Java presented 116 FIELDIANA: ZOOLOGY to the National Museum by Mr. W. E. Balston. Pro- ceedings of the Zoological Society of London, 1909: 371-392. . 1909b. On mammals from the Rhio Archi- pelago and Malay Peninsula collected by Messrs. H. C. Robinson, C. Boden Kloss and E. Seimund, and presented to the National Museum by the Govern- ment of the Federated Malay States. Journal of the Federated Malay States Museum, 4: 99-129. . 1909c. Two new macaques from W. Java. An- nals and Magazine of Natural History, 8th ser., 3: 380- 381. Thompson, N. S. 1967. Primate infanticide: A note and a request for information. Laboratory Primate Newsletter, 6(3): 18-19. Thonglongya, K. 1967. Note on the type locality of some birds and mammals collected by Count Nils Gyldenstolpe in Thailand. Natural History Bulletin of the Siam Society, 22: 185-187. Tickell, S. R. 1854-1875. Mammals of India. Un- published manuscript in library of Zoological Society of London. . 1859. Itinerary, with memoranda, chiefly to- pographical and zoological, through the southerly por- tions of the district of Amherst, Province of Tenas- serim. Journal of the Asiatic Society of Bengal, 28: 421^56. TlMMERMANS, P. J. A., W. G. P. SCHOUTEN, AND J. C. M. Krunen. 1981. Reproduction of cynomolgus monkeys (Macaca fascicularis) in harems. Laboratory Animals, 15: 119-123. Tomiuk, J. 1989a. Gene duplication and polymor- phism at the amylase loci of Macaca fascicularis and Macaca mulatta. Primates, 30: 89-94. . 1989b. Hemoglobin polymorphism in ma- caques with reference to the evolution of Macaca fas- cicularis and Macaca mulatta. Primates, 30: 95-102. Trouessart, E.-L. 1904. Catalogus Mammalium Tam Viventium Quam Fossilium, suppl. 5. R. Friedlander & Sohn, Berlin, 929 pp. Tweedie, M. W. F. 1940. Report on excavations in Kelantan. Journal of the Malayan Branch of the Royal Asiatic Society, 18(2): 1-22. Tweedie, M. W. F, and J. L. Harrison. 1954. Ma- layan Animal Life. Longmans, Green and Co., Lon- don, vii + 237 pp. Ungar, P. S. 1994. Patterns of ingestive behavior and anterior tooth use in sympatric anthropoid primates. American Journal of Physical Anthropology, 95: 197— 219. . 1995. Fruit preferences of four sympatric pri- mate species at Ketambe, northern Sumatra, Indo- nesia. International Journal of Primatology, 16: 221- 245. U.S. Board on Geographic Names. 1952. India, 2 vols. Washington, D.C., 787 pp. . 1964. Northern Vietnam. Washington, D.C., 311 pp. . 1970. Malaysia, Singapore, and Brunei, 2nd ed. Washington, D. C, 1014 pp. 1971a. Cambodia, 2nd ed. Washington, D.C., 392 pp. . 1971b. South Vietnam. Washington, D.C., 337 pp. 1973. Laos, 2nd ed. Washington, D.C., 348 pp. . 1976. Bangladesh. Washington, D.C., 526 pp. . 1982. Indonesia, 3rd ed., 2 vol. Washington, D.C., 1529 pp. U.S. Defense Mapping Agency. 1975a. Bathymetric chart No. 71000. Washington, D.C. . 1975b. Bathymetric chart No. 71006. Wash- Bathymetric chart No. 93036. Wash- Bathymetric chart No. 94004. Wash- Bathymetric chart No. 63400. Wash- Bathymetric chart No. 63020. Wash- ington D.C. 1985. D.C. ington 1986. D.C. ington 1990. D.C. ington 1993. D.C. ington Valerio D. A 1966a. Burma. Washington, D.C, 725 pp. 1966b. Thailand. Washington, D.C, 675 pp. A. J. Pallotta, and K. D. Courtney. 1969. Experiences in large-scale breeding of simians for medical experimentation. Annals of the New York Academy of Sciences, 162: 282-296. Van Couvertng, J. A., and G. Kukla. 1988. Pleis- tocene, pp. 459-464. In Tattersall, I., E. Delson, and J. A. Van Couvering, eds., Encyclopedia of Human Evolution and Prehistory. Garland, New York. Van Peenen, P. F. D., M. L. Cunningham, and J. F. Duncan. 1970. A collection of mammals from Con Son Island, Vietnam. Journal of Mammalogy, 51: 419- 424. Van Peenen, P. F. D., H. Hoogstraal, J. F. Duncan, and P. F. Ryan. 1968. Hematozoa from mammals of South Vietnam. Journal of Protozoology, 15: 608- 614. Van Peenen, P. F. D., S. W. Joseph, Ansari Saleh, R. H. Light, S. Sukeri, and R. See. 1974. The Indo- nesian Developmental Area Study: Observations on mammals from South and East Kalimantan (Borneo). Southeast Asian Journal of Tropical Medicine and Public Health, 5: 390-397. Van Peenen, P. F. D., R. H. Light, and J. F. Duncan. 1971. Observations on mammals of Mt. Sontra, South Vietnam. Mammalia, 35: 126-143. Van Peenen, P. F. D., P. F. Ryan, and R. H. Light. 1969. Preliminary Identification Manual for Mam- mals of South Vietnam. United States National Mu- seum, Smithsonian Institution, Washington, D.C, vi + 310 pp. VARAVUDHI, P., C EkAVIPASTI, U. YODYINGYUAD, T. NOOTPRAPAND, V. YODYINGYUAD, Y. ChAISEHA, W. Set[t]heetham, and K. Suwanprasert. 1989a. Growth and reproductive potential of free-ranging cy- nomolgus monkey (Macaca fascicularis) in 9 selected regions of Thailand. Journal of the Science Society of Thailand, 15: 221-227. Varavudhi, P., K. Suwanprasert, and W. Setthee- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 117 tham. 1992. Reproductive endocrinology of free- ranging adult cynomolgus monkeys (Macaca fascicu- laris) in Thailand, pp. 337-345. In Matano, S., R. H. Tuttle, H. Ishida, and M. Goodman, eds., Topics in Primatology, vol. 3. Evolutionary Biology, Reproduc- tive Endocrinology, and Virology. University of To- kyo Press, Tokyo. Varavudhi, P., J. Suzuki, U. Yodyingyuad, T. NOOTPRAPAND, V. YODYINGYUAD, Y. CHAISEHA, K. SUWANPRASERT, AND W. SETTHEETHAM. 1989b. Mother-infant relationship on patterns of thyroxine and tri-iodothyronine production obtained from 4 major regions in Thailand. Kyoto University Overseas Research Report of Studies on Asian Non-Human Pri- mates, 7: 76-80. Vigors, N. A. 1830. Catalogue of zoological speci- mens, pp. 633-697. In Raffles, S., Memoir of the Life and Public Services of Sir Thomas Stamford Raffles .... John Murray, London. [For name of author of catalogue, see p. 629.] VlTALE, A. F., E. VlSALBERGHI, AND C. DE LlLLO. 1991. Responses to a snake model in captive crab-eating macaques {Macaca fascicularis) and captive tufted capuchins (Cebus apella). International Journal of Pri- matology, 12: 277-286. Volz, W. 1912. Nord-Sumatra . . . , vol. 2. Dietrich Reimer (Ernst Vohsen), Berlin, 428 pp. Vorderman, A. G. 1900. Inlandsche namen van ee- nige Madoereesche planten en simplicia. Natuurkun- dig Tijdschrift voor Nederlandsch-Indie, 59: 140-143. Vos, D. R., G. J. Karssemeuer, and J. A. R. A. M van Hooff. 1992. Ecological constraints on the behav- iour of mother long-tailed macaques (Macaca fasci- cularis). Behavioral Ecology and Sociobiology, 31: 385- 391. Vos, J. de. 1983. The Pongo faunas from Java and Sumatra and their significance for biostratigraphical and paleo-ecological interpretations. Proceedings of the Koninklijke Nederlandse Akademie van Weten- schappen, ser. B, 86(4): 417-425. Vos, J. de, and P. Sondaar. 1994. Dating hominid sites in Indonesia. Science, 266: 1726-1727. Vos, J. de, P. Y. Sondaar, G. D. van den Bergh, and F. Aziz. 1 994. The Homo bearing deposits of Java and its ecological context. Courier Forschungs-Institut Senckenberg, 171: 129-140. Wagner, J. A. [1839]. Die Saugthiere in Abbildungen nach der Natur mit Beschreibungen, suppl. vol. 1. Ex- pedition des Schreber'schen Saugthier- und des Es- per'schen Schmetterlingswerkes, und in Commission der Palm'schen Verlagsbuchhandlung, Erlangen, 551 pp. [For date of publication, see Sherborn, CD., 1892, Proceedings of the Zoological Society of London, 1891: 591.] Wallace, A. R. 1 869. The Malay Archipelago vol. 1. Macmillan and Co., London, 478 pp. . 1895. Island Life .... Macmillan and Co., London, 563 pp. Wallis, J., B.J. King, and C. Roth-Meyer. 1986. The effect of female proximity and social interaction on the menstrual cycle of crab-eating monkeys (Macaca fascicularis). Primates, 27: 83-94. Warren, McW. 1966. Observations on the fauna of Pulau Tioman and Pulau Tulai. 12. Primate malaria. Bulletin of the National Museum, Singapore, 34: 1 50- 158. Warren, McW., W. H. Cheong, H. K. Fredericks, and G. R. Coatney. 1970. Cycles of jungle malaria in West Malaysia. American Journal of Tropical Medi- cine and Hygiene, 19: 383-393. Washburn, S. L., and D. A. Hamburg. 1968. Ag- gressive behavior in Old World monkeys and apes, pp. 458-478. In Jay, P. C, ed., Primates: Studies in Adaptation and Variability. Holt, Rinehart and Win- ston, New York. Waterhouse, G. R. 1838. Catalogue of the Mammalia Preserved in the Museum of the Zoological Society of London, 2nd ed. Richard and John E. Taylor, London, 68 pp. Weber, B. E. 1972. A parks system for West Malysia. Oryx, 11: 461-469. Weber, M. 1890a. Einleitung, pp. i-xii. In Weber, M., ed., Zoologische Ergebnisse Einer Reise in Niederlan- disch Ost-Indien, vol. 1. E. J. Brill, Leiden. . 1890b. Mammalia from the Malay Archipel- ago. I. Primates, Prosimiae, Galeopithecidae, Carniv- ora, Artiodactyla, Edentata, Marsupialia, pp. 93-1 14. In Weber, M., ed., Zoologische Ergebnisse Einer Reise in Niederlandisch Ost-Indien, vol. 1. E. J. Brill, Lei- den. Weinman, D., and N. S. Wiratmadja. 1969. The first isolates of trypanosomes in Indonesia and in history from primates other than man. Transactions of the Royal Society of Tropical Medicine and Hygiene, 63: 497-506. Weitzel, V., and C. P. Groves. 1985. The nomen- clature and taxonomy of the colobine monkeys of Java. International Journal of Primatology, 6: 399-409. Weitzel, V., C. M. Yang, and C. P. Groves. 1988. A catalogue of primates in the Singapore Zoological Ref- erence Collection, Department of Zoology, National University of Singapore (formerly the Zoological Col- lection of the Raffles Museum). Raffles Bulletin of Zo- ology, 36: 1-166. Wharton, R. H., D. E. Eyles, McW. Warren, and W. H. Cheong. 1 964. Studies to determine the vectors of monkey malaria in Malaya. Annals of Tropical Medicine and Parasitology, 58: 56-77. Wheatley, B. P. 1978. Foraging patterns in a group of longtailed macaques in Kalimantan Timur, Indo- nesia, pp. 347-349. In Chivers, D. J., and J. Herbert, eds., Recent Advances in Primatology, vol. 1. Behav- iour. Academic Press, London. . 1980. Feeding and ranging of East Bornean Macaca fascicularis, pp. 215-246. In Lindburg, D. G., ed., The Macaques: Studies in Ecology, Behavior and Evolution. Van Nostrand Reinhold, New York. -. 1982. Adult male replacement in Macaca fas- cicularis of East Kalimantan, Indonesia. International Journal of Primatology, 3: 203-219. 1 984. Copulatory calls of wild macaques, Ma- caca fascicularis, in Kutai Nature Reserve, East Ka- limantan, Indonesia. International Journal of Prima- tology, 5: 391. (Abstract only.) 118 FIELDIANA: ZOOLOGY . 1988. Cultural behavior and extractive for- aging in Macaca fascicularis. Current Anthropology, 29: 516-519. . 1989. Diet of Balinese temple monkeys, Ma- caca fascicularis. Kyoto University Overseas Research Report of Studies on Asian Non-Human Primates, 7: 62-75. 1991. The role of females in inter- troop en- counters and infanticide among Balinese Macaca fas- cicularis, pp. 169-172. In Ehara, A., T. Kimura, O. Takenaka, and M. Iwamoto, eds., Primatology Today: Proceedings of the XHIth Congress of the Interna- tional Primatological Society, Nagoya and Kyoto, 1 8- 24 July 1990. Elsevier Science, Amsterdam. Wheatley, B. P., A. Fuentes, and D. K. Harya Putra. 1993. The primates of Bali. Asian Primates, 3(1-2): 1-2. Wheatley, B. P., and D. K. Harya Putra. 1994. The effects of tourism on conservation at the Monkey For- est in Ubud, Bali. Revue d'Ecologie (Terre & Vie), 49: 245-257. Whitmore, T. C. 1972. The Gunong Benom Expedi- tion 1 967. 2. An outline description of the forest zones on north-east Gunong Benom. Bulletin of the British Museum (Natural History), Zoology, 23(2): 11-15. Whitten, A. J., S. J. Damanik, J. Anwar, and N. Hisyam. 1984. The Ecology of Sumatra. Gadjah Mada University Press, Yogyakarta, xiv + 583 pp. Willink, H. D. T. 1905. Mammalia voorkomende in Nederlandsh-Indie. Natuurkundig Tijdschrift voor Nederlandsch-Indie, 65: 153-345. Wilson, C. C, and W. L. Wilson. 1973. Final Report: Census of Sumatran Primates, Unpublished. Regional Primate Research Center, University of Washington, Seattle, 24 pp. . 1975. The influence of selective logging on primates and some other animals in East Kalimantan. Folia Primatologica, 23: 245-274. 1977. Behavioral and morphological variation among primate populations in Sumatra. Yearbook of Physical Anthropology, 20: 207-233. Wroughton, R. C. 1915. Bombay Natural History Society's Mammal Survey of India, Burma and Cey- lon. Report No. 17. Journal of the Bombay Natural History Society, 23: 695-720. . 1918. Summary of the results from the Indian Mammal Survey of the Bombay Natural History So- ciety. Journal of the Bombay Natural History Society, 25: 547-598. Yanuar, A. 1 994. Survey of the banded leaf monkey (Presbytis femoralis rhionis) on Bintang Island, In- donesia. Asian Primates, 3(3-4): 1-2. Yanuar, A., and J. Sugardjito. 1993. Population survey of primates in Way Kambas National Park, Sumatra, Indonesia. Tigerpaper, 20(3): 30-36. Yin, U Tun. 1954. Wildlife preservation and sanctu- aries in the Union of Burma. Journal of the Bombay Natural History Society, 52: 264-284. . 1967. Wild Animals of Burma. Rangoon Ga- zette, Rangoon, 301 pp. Yoshiba, K. 1964. Report of the preliminary survey on the orang-utan in North Borneo. Primates, 5: 1 1- 26. Yoshida, T., T. Shimizu, F. Cho, and N. Goto. 1993. Relative growth of physiques in laboratory-bred cy- nomolgus monkeys: A longitudinal study during the first 6 years of life. Experimental Animals, 42: 435- 441. Zelebor, J. [1869]. Saugethiere, pp. 1-42. In Reise der osterreichischen Fregatte 'Novara' um die Erde .... Zoologischer Theil, Bd. I. Kaiserlich-Koniglichen Hof- und Staatsdruckerei, Wien. [For date of publication, see Zoological Record, 1871: 2.] Zimmermann, E. A. W. 1780. Geographische Ge- schichte des Menschen und der vierfussigen Thiere, vol. 2. Weygandschen Buchhandlung, Leipsig, 432 pp. Zollinger, H. 1845. Land- en volkenkunde. Een uits- tapje naar het eiland Balie. Tijdschrift voor Neerland's Indie, 7(4): 1-56. . 1 847. Land- en volkenkunde. Het eiland Lom- bok. Tijdschrift voor Neerland's Indie, 9(2): 177-205. Zuckerman, S. [1933]. The menstrual cycle of the pri- mates. — Part VI. Further observations on the breeding of primates, with special reference to the suborders Lemuroidea and Tarsioidea. Proceedings of the Zoo- logical Society of London, 1932: 1059-1075. [For date of publication, see Proceedings of the Zoological So- ciety of London, 1933: 229.] Zumpe, D., and R. P. Michael. 1983. A comparison of the behavior of Macaca fascicularis and Macaca mulatta in relation to the menstrual cycle. American Journal of Primatology, 4: 55-72. . 1990. Effects of the presence of a second male on pair-tests of captive cynomolgus monkeys {Macaca fascicularis): Role of dominance. American Journal of Primatology, 22: 145-158. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 119 Appendix 1: Specimens Examined (Total 2,049) Macaca fascicularis fascicularis, Total 1,550 Skins and Skulls, 739 CAMBODIA, 2. Mainland '(2), Siemreab— mnhn 1929.460; no locality— mnhn 1961.61 1. INDONESIA, 456. Bali (21), Banjoe Wetan- mzb 6521-6523; Batoe-meringgit— mzb 2001; Bratan, Gunung, amnh 107555-107557; Desa Poetjang— rmnh Coll. No. E85 (external measure- ments from Sody, 1933, p. 94); Gilimanuk— amnh 107561-107568; Gitgit-NMS 16632; Jembrana- rmnh Coll. No. El 39 (external measurements from Sody, 1933, p. 94); Sendang— rmnh Coll. Nos. E34, E64, E74 (external measurements for all three from Sody, 1933, p. 94); Bangka (4), Pamuja, Tan- jung— usnm 124863; Rengsam, Tanjung— usnm 1247 10; no locality-RMNH Coll. Nos. Bk52, Bk53; Batam (4) "Tanjong Sauh"-BM(NH) 1909.4.1.29, bnhs 5080; Tanjong Turut-BM(NH) 1909.4.1.27, 1909.4.1.28; Bawean (8), no locality-MZB 1841- 1846, usnm 151829, 151830; Belitung (2), Batu, Tanjung-USNM 124969, 124970; Bengkalis (1), Kapos Tinggi— usnm 143582; Benua (2), no lo- cality-usNM 101638, 101639; Bintan (8), north coast— usnm 115676; Pasir Panjang— bm(nh) 1909.4.1.26, usnm 115677; Sungei Biru-BM(NH) 1 909.4. 1 .23-1 909.4. 1 .25; no locality-ZRC 4-093, 4-094; Borneo: Kalimantan (93), Ambawang, Sun- gai — usnm 142225; Badang— amnh 103730, 103731, 106024-106026, mzb 6513; Berau, Sun- gai— usnm 196817-196822; Birang, Sungei— usnm 196825 (includes skinned head in fluid); Buntok— bm(nh) 1910.4.5.23; Goson Djerong — usnm 196814; Hantakan — usnm 521837; Karangin- tan— bm(nh) 1910.4.5.22; Karangmumus, Sun- gai— usnm 196816; Karangtigau, Tanjung— usnm 196824; Kembangjanggut— mzb 8130; Liang Koe- boeng— rmnh 160, 178; Loa Bambam— usnm 196815; Long Peleben-AMNH 106027, mzb 6514; Mahakam, Sungai — usnm 196813; Merah — mzb 1164; Muaratewe — bm(nh) 1910.4.5.19, 1910.4.5.20; Parit-AMNH 103644-103658, 103660-103663, mzb 6515-6517, zmb 48005, 48006, 92191; Pelawan, Sungai-USNM 198301; Perbuah-AMNH 107091-107096, 107098, 107099, mzb 6519, 6520; Putussibau-ANSP 14038, 14039, 14044; Riam- amnh 106283-106285, mzb 6518; Roema Manoeal— rmnh 23, 24; Sembak- ung, Sungai— mzb 576, 6512; Semitau— rmnh 6; Sintang— ansp 14037, 14042, 14043; Tangarveng I. -usnm 1 96804; Telang— usnm 521838; Tibang, Mt.— mcz 22277; no locality— ansp 14041, bm(nh) 1856.9.3.6, zsbs Coll. Nos. 272, 738, 788, 1227, 1354, 1373, 1527;5w/a«(3),south-usNM 144419; no locality— zrc 4-098, 4-099; Durian (2), no lo- cality—mzb 241, 242; Flores (7), Mbura— mzb 2384-2387; Rana Mese-NMS 1 6630; Sano, Wai- mzb 2388; no locality— bm(nh) 1864.4.12.1; Gal- ang (4), no locality— zrc 4-100-4-103; Java (84), Bandung, near— usnm 521839; Bantargebang— mcz 12757, usnm 156291, 156292, 156295; Ban- yuwangi— rmnh 959; Batoeraden— rmnh Coll. No. 89C; Bogor— mzb 1884; Camara— mzb 6482; Can- diroto— rmnh Coll. No. 107B; Cihara— usnm 156456, 156458; Cikujang-MZB 6483, 6485; Cil- acap-BM(NH) 1909,1.5.27, 1909.1.5.28; Cire- bon— amnh 101891, 102016; Ciremay, Gunung - amnh 102015, 102017-102022; Ciwangi-BM(NH) 1909.1.5.29, 1909.1.5.30; Depok-Mcz 12755 (ex- ternal measurements from usnm field cat.), 1 2756; Gedangan— rmnh Coll. Nos. 74, V9, V10; Indra- mayu— mzb 2961, 2962; Jasinga— mzb 2052, 3189; Java, West— rmnh 2b, 3c, 4h; Kaligoea— mzb 574, 575; Kalipucang-BM(NH) 1909.1.5.19, 1909.1.5.32, 1909.1.5.33; Linggajati-AMNH 101811; Majalengka-AMNH 101808-101810; Pangandaran— bm(nh) 1909.1.5.24; Pangonan— rmnh Coll. No. M40; Pasir Carolina— mzb 6484; Pelabuhanratu, Teluk— usnm 156457, zrc 4-134; Salak, Gunung— rmnh Coll. No. A 147; Saron- gen— mcz 12759; Singkil, Gunung— rmnh 5919o, 5919p, 5919q; Tamadjaija— mcz 12758; Taman- sari— bm(nh) 1954.51, zrc 4-133; Tasikmalaya— bm(nh) 1909.1.5.31, mzb 1604-1609; Tiloe, Goenoeng— rmnh 5919n; Tjeringin— rmnh Coll. Nos. 13f, 25f, 26f; Wonokojo— rmnh Coll. Nos. Won3, Won4, Won5; no locality— nms 16628, rmnh 2505a, 2505b, 2527, 3094, 3095, 3097, zmb 113, zmuz 11627 (skin)/11630 (skeleton); Kan- gean (4), no locality — bm(nh) 1 9 10.4.6. 1 — 1910.4.6.4; Karimata (2), Pai, Teluk — usnm 125101, 125102; Karimun (7), Mensuda Bay- usnm 122849; Monos-BM(NH) 1909.4.1.34, 1909.4.1.35; Pemeral-BM(NH) 1909.4.1.30- 1909.4.1.33; Kundur (2), Selatbliat-BM(NH) 1909.4.1.36, 1909.4.1.37; Lagong (1), no locali- ty—usnm 104853; Laut (1), no locality— usnm 104854; Lingga (2), no locality— usnm 101602, 101603 (external measurements from Miller, 1906c, p. 276); Lombok (3), Pussuk forest— mzb 6528; Sewela-NMS 16634, zmb 92306; Mapur(l), Mentigi— zrc 4-097; Matasiri (1), east coast— usnm 151831; Mursala (2), no locality— usnm 120 FIELDIANA: ZOOLOGY 114559, 114560; Natuna Besar (2), Binjai, Sun- gai— zrc 4-139, 4-140; Nguwal (6), no locality— zrc 4-104-4-109; Nias (1 1), Hilisimaetano— ansp 20404-20406; Lafau-USNM 141372; Samasa- ma— usnm 141371; Siaba, Teluk— usnm 121868- 121871; Soliga-Mcz 36029, 36030; Pelapis Ten- gah (1), no locality— mzb 2914; Penida (3), no lo- cality— amnh 107558-107560; Serasan (2), no lo- cality—usnm 104852, zrc 4-144; Serutu (1), no locality— mzb 29 1 5; Siantan (3), no locality —usnm 101711, zrc 4-136, 4-137; Subi-kecil (3), no lo- cality—zrc 4-141-4-143; Sugi (1), no locality— usnm 115675; Sumatra (123), Batangkuis— zsbs Coll. No. 9; Bungur-Buikt-ZMB A25.09; Deli, Sungai— ansp 20217, 20219; Goenoengsetan- Meloewak— mzb 6481; Indragiri, Sungai— usnm 113169; Kalianda- amnh 1 02904-1 02906; Kam- bang— zmb A25.09; "Kg. Baru"— zsbs Coll. Nos. 156, 176, 180; Kotabumi-ZRC 4-124; Lesten- mzb 6480; Lubuklinggau— amnh 102211; Me- dan-ANSP 20221, zsbs Coll. Nos. 1/10, 5, 6, 15, 16, 21, 22, 26, 30-32, 35, 36, 42, 45-48, 51-53, 68 (Bruegel), 68 (Widnmann), 76, 79, 82, 89, 91, 92, 101, 105, 134, W2,Wll;Medan, forest near- zsbs Coll. No. 11; Muaradua— amnh 102763- 102770; Padang-RMNH 518, 981, 1051; Padang highlands— rmnh e2; Pajo— bm(nh) 1879.6.28.4; Pematangsiantar— mcz 35937, 35938; Sandaran Agong— zrc 4-110; Sanggul, Bukit— amnh 106565-106569, mzb 6477-6479; Serdang-ZSBS Coll. No. 134;Siulakderas-BM(NH) 1919.11.12.8; Sukadana— amnh 102907-102910; Sumatra, [east- central]— zmb A 14.08/37; Sumatra, east coast— zsbs Coll. Nos. 105, 135, 156, 166-168, 174; Su- matra, west coast— bm(nh) 1882.7.24.1; Tanjung- morawa— zsbs Coll. Nos. 7 1 [b], 72, 1 37; Tapanuli, Teluk— usnm 114505, 114506; Tapung-kanan, Sungai— zmb A14.08; Tarusan, Teluk— usnm 141145; Telukbetung— fmnh 14805; Telukpan- ji— zrc 4-111; no locality— NHMBa 3347, 3348, rmnh 1479, usnm 271018, zsbs Coll. Nos. 28, 35, 36, 38, 58, 64, 67, 93, 107, 112, 135, 157, 184, 1 85, 203-205, 2 1 2, 232; Sumba (8), Langgaliroe- mzb 6526, 6527, zmb 92075; Mao Marroe— mzb 867-869; Payeti-Kambaniru— mzb 865, 866; Sumbawa (8), Batudulang— mzb 6529, 6530, zmb 92307, 92308; Dompu-NMS 1025; Kambing, Pu- lau— mzb 6531; Ntori— zmb 92309; Oo vicinity— nms 1026; Tanahbala (2), no locality— usnm 121802, 121803; Timor (9), Kuatnana-ZSBS 1911/ 2108; Lelogama-ZSBS 1911/2102, 1911/2104, 1911/2105; Mutis, Gunung— mzb 6524, zmb 92 1 3 1 , 92 1 32; Nikiniki-MZB 6525; Timau, Fatu- zsbs 1911/2101; Tuangku (3), north coast— usnm 114408-114410; Uwi (1), no locality-uSNM 101666. MALA YSIA, 1 63. MALA YSIA: SABAH, Bang- gi (5), Banggi, Pulau, south— zrc 4-116-4-118; Karakit-FMNH 140939; Sabor-FMNH 140938; Borneo (74), Abai-Mcz 35600, 35606, 35611- 35613, 35629, 35641, 35642, 35693, 35694, 35700, 35701, 35722, 35724, 35726, 35727, 35729, 35731, 35732, 35744, 35761; Betotan,ZRC 4-115; Bongkabong-Mcz 37409, 37411, 37412; Darvel Bay— zmb A86.10; Garau— mcz 37349, 37351, 37414, 37418; Kampong Bundu Tuhan- usnm 292555-292558; Kampong Kiau — mcz 37348, 37352, zrc 4-1 20, 4-121 ; Keningau-AMNH 188343; Kenokok zrc 4-122; Kiaulan— mcz 37350, 37353; Kinabatangan, Sungai — usnm 19193; Kretam Besar, Sungai— fmnh 68700; Kre- tam Kechil, Sungai— fmnh 68699; Lahad Datu— zmb A89.04; Papar— usnm 317190; Ranau— mcz 37346, usnm 3 1719 1-3 17 195; Rayoh-ZRC 4-1 19; Rugading— mcz 37357; Sapagaya Forest Re- serve—fmnh 68701, 68702; Sibuga Besar, Sun- gai—fmnh 33545; Talibang— mcz 37354, 37406, 37413; Tawau— amnh 31288; Tempasuk, Sun- gai—mcz 37417; Tenompok— mcz 37415, 37416; Tinonkok— mcz 37347; Tuaran-Kampong Tengh- ilan Road-Mcz 37355, 37356, 37405, 37408; no locality-uSNM 344989-344992; Sebatik (2), no locality— usnm 175896 (skin and skull)/ amnh 30620 (postcranial skeleton), usnm 175897. MA- LAYSIA: SARAWAK, Borneo (15), Baram, Ba- tang-BM(NH) 1894.6.12.2, USNM 83945; Belaga or Kalulong, Bukit-BM(NH) 1951.66, 1951.68; Bukar, Sungai— zrc 4-113; Entawa, Tanjong— bm(nh) 1955.710; Kuching-SMK 092/3, 092/8; Kuching, probably— zmb 4939; Long Ekang— fmnh 88582; Paku, Saribas-BM(NH) 1955.711, 1955.712; Pelandok, Sungai-ZRC 4-112; Punang, Sungai— zrc 4-114; Selikan, Bukit— usnm 83944. MALA YSIA: WEST MALA YSIA, mainland (29), Benom, Gunong— zrc 4-065, 4-066; Changkat Mentri— bm(nh) 1955.1510, zrc 4-064; Cherak- ah, Bukit-BM(NH) 1955.1516, zrc 4-069; Dusun Tua— zrc 4-068; Gaik Liew Estate — bm(nh) 1955.1515; Hantu, Tanjong -zrc 4-063; Kelang Road-BM(NH) 1971.2611; Melaka — bm(nh) 1875.3.29.1, rmnh 41g; Nanas, Bukit— bm(nh) 1939.180, 1939.181, 1955.1512; Nyalas-ZRC 4-078 (external measurements from Weitzel et al., 1988, p. 107); Pong-BM(NH) 1934.7.18.1; Port Dickson -bm(nh) 1939.229 (skin)/RCS(OM) A85.4 (skull), zrc 4-073-4-075; Tanjong Panjair— bm(nh) 1955.1514, zrc 4-067; Tanjong Tuan, Keramat— zrc 4-072; Telom, Sungai-BM(NH) 1934.7.18.4, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 121 1934.7.18.5; Ulu Ijok-BM(NH) 1934.7.18.2, 1934.7. 18.3; no locality-usNM 301 758; Acheh(l), no locality — bm(nh) 1955. 1 5 IS; A ur (2), no local- ity-bm(nh) 1955.1520, zrc 4-080; Burau (1), no locality— bm(nh) 1955.1519; Langkawi (1), no lo- cality— bm(nh) 1 909. 11.1.1; Pemanggil (2), no lo- cality-BM(NH) 1955.1508, usnm 1 12500; Pinang flj(8), Pantai Krachut-ZRC 4-062; Telok Bahang bm(nh) 1955.1521, zrc 4-056-zrc 4-061; Pinang [2] (2), no locality -bm(nh) 1949.426, 1955.1523; Pintu Gedong (3), no locality— bm(nh) 1955.1517, zrc 4-070, 4-071; Redang (2), no locality— zrc 4-054, 4-055; Tinggi (5), no locality— bm(nh) 1909.4.1.21, 1909.4.1.22, 1955.1527, zrc 4-131, 4-132; Tioman (11), Juara, Telok — bm(nh) 1908.1.25.1, 1908.2.25.1, 1955.1524, 1955.1525, zrc 4- 1 25-4- 1 27; Sedagong, zrc 4- 1 29, 4- 1 30; no locality-BM(NH) 1955.1526 (skin)/zRC 4-128 (skull), usnm 101744. PHILIPPINES, 43. Basilan (3), Basilan I., east end— usnm 144666; Camp No. 4 and Camp No. 5, between— usnm 144665; Isabela— usnm 125326; Cagayan Sulu (1), no locality— usnm 125325; Mindanao (39), Bucong— fmnh 65440-65444; Canibongan— fmnh 67737-67739; Gubat— fmnh 67725, 67729-6773 1; Libu-FMNH 67727, 67728; Mamara— fmnh 67717, 67718; Pulunbato, Mt.— bm(nh) 1880.1 1.24.1; San Ramon-FMNH 33507- 33511; Sigayan-FMNH 67722, 67732, 67740- 67742; Situbo-FMNH 67719-67721, 67723, 67724; Tacuta — fmnh 67733-67735; Tampa- Ian— fmnh 67736; Zamboanga— usnm 144667, 144668, 144698, SINGAPORE, 1. Singapore (7), Botanical Gar- den-ZRC 4-082, 4-084; Changi-BM(NH) 1909.4.1.20; Punggol-BM(NH) 1955.1513, zrc 4-086; no locality— zmb 5444, zrc 4-090. THAILAND, 60. Mainland (32), Aranyaprath- et— mcz 35931; Ban Huai Maenam Noi— fmnh 99649, 99650; Ban Nong Kok-ZRC 4-034; Ban Nong Put— ctnrc (catalog number unknown); Ban Pak Nam Pho-Mcz 23812, 23813, zrc 4-031; Ban Phra Muang-USNM 83271, 83273; Ban Phu Toie-FMNH 99639, 99640; Ban Sai Kau-BM(NH) 1903.2.6.3; Ban ThapPlik, 1 kmE-FMNH 105654; Ban Thap Plik, 1 km NE-fmnh 105653; Kam- pong Biserat-BM(NH) 1903.2.6.1, 1903.2.6.4; Kantang— zrc 4-032; Kata Taek— fmnh 99651- 99656; Khao Rang Kai— ctnrc (catalog number unknown), fmnh 105689; Laem Sing mountains— usnm 251662; Lat Bua Khao, Sathani— usnm 236631, zrc 4-033; Tham Hom-FMNH 105649; Tyching-USNM 83274, 83275; Butang(l), no lo- cality— bm(nh) 1955.1511; Chang (3), no locali- ty-BM(NH) 1915.1 1.4.7, usnm 201551, zrc 4-039; Kut (8), no locality-BM(NH) 1915.11.4.8- 1 9 1 5. 1 1 .4. 1 0, 1 976. 1 826 (skin)/zRC 4-040 (skull), usnm 201552, 201553 (external measurements from Kloss, 1916a, p. 67), 254741, zrc 4-041; Phangan (2), southwest— zrc 4-044, 4-045; Phi Phi Don (4), no locality —zrc 4-046-4-049; Samui (2),west-BM(NH) 1955. 1522, zrc 4-050; Talibong (2), no locality— usnm 83272, zrc 4-051; Tarutao (6), Sungei Adang-BM(NH) 1909.11.1.2, 1909.1 1.1.3; Telok Wau-BM(NH) 1955. 1509, zrc 4-053; no locality-usNM 123990, 123991. VIETNAM, 8. Mainland (6), Cochin China- bm(nh) 1881.6.30.2, mnhn 1882.2; Ho Chi Minh City— fmnh 33505; Ho Chi Minh City, Zoological Garden-MNHN 1929.459; Sontra Peak, 3.9 km W, 0.3 km S-usnm 356968; Xa Trang Bom-ZRC 4-020 (external measurements from Weitzel et al., 1988, p. \ 14); Phu Quoc(2)-bm(nh) 1928.7.1.10, mnhn 1929.455. Skins only, 285 CAMBODIA, 1. Mainland (1), no locality - RMNH 38. CAMBODIA OR VIETNAM, 2. Mainland (2), "Camboja or Cochinchine"— bm(nh) 1 878.6. 1 7.4, 1878.6.17.5. INDONESIA, 241. Bali (1), Bratan, Danau- bm(nh) 1913.3.6.7; Bangka (1), Simpang— zsbs Coll. No. 25; Borneo: Kalimantan (82), Borneo, southeast — rmnh 35v (skull inside); Borneo, southwest— rmnh 32s, 33t, 34u (skulls inside); Kampong Hadjak— irsn 9796; Karangtigau, Tan- jung— usnm 196823; Muaratewe — bm(nh) 1910.4.5.21; Parit-AMNH 103659 (skull not lo- cated, cranial measurement from G. H. Albrecht, pers. comm., Oct. 1991), zmb 92190; Pelaihari— rmnh 36w (skull inside); Perbuah— amnh 107097; Poelau— rmnh 229 (cf. Hooijer, 1962b, p. 46, no. 48); Rantau— rmnh 4617 (cf. Hooijer, 1962b, p. 44, no. 11); Riam— amnh 106282; Senoeang— amnh 1 07 1 00 (male skin mismatched with female skull); Sintang— ansp 14036, 14040; no locality— zmb (unnumbered, 24 Apr. 1902), zsbs Coll. Nos. 3PS,4PS,7PS, 10-12, 1 5 Guvenile male), 15 (adult male), 16, 21, 36(450), 48, 157, 201, 205, 276, 277, 297, 298, 306, 372, 373, 445(316), 587, 616- 618, 694, 701, 709, 710, 730, 740, 741, 743, 749, 764-766, 771, 780, 1013, 1014, 1228, 1358, 1359, 1372, 1378, 1386, 1387, 1414, 1437, 1438, 1452, 1464, 1465, 1477, 1478, 1488, 1505, 1510, 1534, unnumbered (Breugel), unnumbered (Breugel, "von Zengen"); Java (25), Bantargebang— mcz 12760; Gedangan— rmnh (unnumbered); Indra- mayu— mzb 2939, 2960; Kawarasan— rmnh 2067; 122 FIELDIANA: ZOOLOGY Pangandaran— bm(nh) 1909.1.5.26; Pangrango, Gunung— rmnh 2067, 2067/II (skulls inside); Se- marang— zsbs (unnumbered); Tasikmalaya— mzb 1610; Java, West— rmnh 1, 5e, 6, 7g (skulls in- side), 8; no locality— NHMBa 3131, 3161, 3352, rmnh 9i, lOj, Ilk, 121, 13m, 14 (skulls inside); smtd (unnumbered); Kambing (1), no locality— rmnh 30p (skull inside); Laut (1), no locality— bm(nh) 1939.1054 (skull inside); Natuna Besar (6), Sinubing-BM(NH) 1939.1053 (skull inside); Ulu, Sungai— zrc 4-138; no locality— bm(nh) 1939.1055, 1964.428 (skulls inside), mcz 6660, 6661 (skulls inside); Mas (1), no locality— zrc 4-135 (skull not located); Pejantan (1), no locali- ty—zsbs I; Sumatra (114), Batangkuis— zsbs Coll. Nos. 3, 1 3, 1 5, 38, 47, 88, 96, 97, 1 97, 200; "Bran- dan?"-ZMB 38542; Deli, Sungai— ansp 20218, 20220 (fetuses in fluid); "I. Lendung"— zsbs Coll. Nos. 32, 74, unnumbered juvenile; "Kg. Baru"— zsbs Coll. Nos. 63, 77, 79, 81, 113; Medan-ZSBS Coll. Nos. 1, 6/II, 7, 7/II, 8/II, 9/II, 14, 24, 28 (juvenile), 29, 42/111, 43, 47/11, 52, 58, 60, 62, 67, 67/11, 71, 74, 75, 77, 80, 81, 81/11, 82, 90, 93, 94, 98, 99/11, 100/11, 102, 103, 104, 106, 111, 115, 1 16/VIII, 119, 124/III; Menam-ZMB 40820; Pa- dang— rmnh 44x, 45y, 46z (skulls inside), 47a, 48b, 49c, 50d (skulls inside), zmuz 1 1625, 1 1626; Palembang, Kotamady— zsbs Coll. Nos. 22, 23; Pangkalansusu— usnm 502457 (infant in alcohol); Sumatra, east coast— zsbs Coll. Nos. 1 58, 1 70, 1 7 1 , 189; Sumatra, south-ZMB 38540, 38541; Tan- jungmorawa— zsbs Coll. Nos. 60, 73[b], 75[b]; no locality— mnhn 362/233, zmb Coll. Nos. 9, 25, unnumbered juvenile, unnumbered male, zmuz 11619 (skull inside), zsbs Coll. Nos. 14, 27, 52, 61, 68, 70, 78, 78/1905, 95, 111, 115, 117, 123, 141, 155, 198, 199, 201, 202, 216, 235, 236, 242; Sumbawa (1), Batudulang— nms 16631 (cranial measurements from Mertens, 1936, p. 319, and G. H. Albrecht, pers. coram., Oct. 1991); Timor (6), Amarassie— rmnh 29o (skull inside); Bo- kong— zsbs 1911/2110; Lelogama— zsbs 1911/ 2103, 1911/2106, 1911/2107; Nikiniki-ZMB 92133; island unknown (1), Sunda Islands— zmb 111. INDONESIA OR MALAYSIA, 1. Borneo (1), Sendung— zmb (unnumbered). MALAYSIA, 24. MALAYSIA: SABAH, Borneo (2), Abai-Mcz 35695; Tuaran-Mcz 37407 (male skin mismatched with female skull); Sebatik (1), no locality- amnh 30619. MALAYSIA: SARA- WAK, Borneo (15), Baram, Batang— nms 4689, smk 092/1 3 (skull inside); Belaga or Kalulong, Bu- kit— bm(nh) 1 95 1 .67 (female skin mismatched with male skull); Dulit, Bukit— smk 092/9 (skull in- side); Kuching— smk 092/5 (skull inside); Ku- ching, 10th mile— smk 092/12 (skull inside); Ling- ga-SMK 092/2, 092/7 (skulls inside); Miri-Mcz 6662 (skull inside); Miri district— NHMBe 817; Penrissen, Gunong-SMK 092/4, 092/1 1/A3.312 (skulls inside); Punang, Sungai— smk 092/6, 092/ 1 0; Segobang, Sungai -smk 092/ 1 1/47 1 3 (skull in- side); Dindding(l), no locality— bm(nh) 1905.3. 1.3. MALAYSIA: WEST MALAYSIA, mainland (4), Melaka-MNHN 1848.401, 1848.402 (skulls in- side), rmnh 40, 42 (skull inside); Aur (1), no lo- cality—zrc 4-079. PHILIPPINES, 3. Mindanao (1), Zamboanga del Sur Prov.— siconbrec 1586 (living captive); Tawitawi (2), no locality— siconbrec 1225, 1475 (living captives). SINGAPORE, 5. Singapore (5), Botanical Gar- den—zrc 4-081 (skull inside), 4-083; Punggol— zrc 4-087, 4-088; Sembawang, Sungai— zrc 4-089. THAILAND, 3. Mainland (3), Bangkok— mnhn 1219 (skull inside); Ban Thap Plik, 1 km E— fmnh 105655 (infant in fluid); Kampong Biserat— bm(nh) 1903.2.6.2. VIETNAM, 5. Mainland (5), Ho Chi Minh City-NMS 5786, 5787; Ho Chi Minh City, Bo- tanical Gardens— bm(nh) 1928.7.1.9, mnhn 1929.457; Tay Ninh-BM(NH) 1928.7.1.7. Skulls only, 526 INDONESIA, 434. Bali (2), Bratan, Danau- bm(nh) 1913.3.6.5 (possibly skull of bm(nh) 1913.3.6.7, skin only); Jembrana— rmnh Coll. No. E136 (external measurements from Sody, 1933, p. 94); Bangka (173), Jebus-RMNH Coll. No. Bk7; no locality— rmnh Coll. No. Bk74, zluu 1, 2, 4- 16, 18, 20, 22, 25-30, 32-37, 39^13, 45-62, 64- 67, 73-75, 77-105, 106 (juvenile cranium only), 106 (adult mandible only), 107-112, 113, (juve- nile cranium only), 113 (adult mandible only), 1 1 4- 116, 117 (cranium and mandible), 1 17 (mandible only), 118, 119, 120 (cranium only), 120 (man- dible only), 121-134, 136, 137, 140-165, 167- 171, 173-178, 179 (mandible only), 181-184 (mandibles only), 185, 187-189 (mandibles only), zsbs 1964/232; Bintan (2), Pasir Panjang— zrc 4-092, Sungei Biru-ZRC 4-091; Borneo: Kali- mantan (24), Borneo, southeast— rmnh p, zmb 38543; Gosong Djerong— usnm 196827; Jembay- an, Sungai-USNM 199181, 199183, 199184, Kar- angan, Sungai— usnm 198300; Kariorang— zmb 48494, unnumbered; Loa Bambam— usnm 196826; Long Peleben— mnhn 1974; 144; Purukcahu— bm(nh) 1910.4.5.18; Tanjung— NHMBa 3747; no locality— amnh 107100 (female skull mismatched FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 123 with male skin), rcs(om) G 1 66.3 1 , zsbs 1 944/3009, Coll. Nos. 12-4-12-6, 20, 363/368, 748, 848, 864; Java (123), Bandung— rmnh 3438, zluu (unnum- bered); Cirebon — nms 6974, rcs(om) A85.6, A85.61-A85.65; Gedangan-RMNH Coll. VI, un- numbered adult males (2), unnumbered infant (1); Java, West— rmnh cat. ost. b-cat. ost. e (4); Jock- aboemi— zmb A34.09; Malabar-ZLUU (unnum- bered); Manggis-ZSBS 1909/4062, 1909/4601, unnumbered specimens (7), Margomulio, Gun- ung— zsbs (unnumbered); Ngawi— rmnh 4660; Pangandaran bm(nh) 1909.1.5.25 (with mounted skin, not examined); Pangrango, Gunung— nhrm Z2923-Z2925, Z3146-Z3150, Z3152-Z3154, Z3165, Z3166, Z3292, Z3293, Z3295, Z3314, Z3316, Z3320-Z3322, Z3324 (adult female), Z3324 (subadult female), Z3325, Z3328-Z3330; Salak, Gunung— rmnh Coll. Nos. A 137, A221; Tjeringin— rmnh Coll. No. 12f; Tulungagung— rmnh 4650-4655; no locality— ansp 12, NHMBa (unnumbered), nhrm Z2698-Z2700, Z2702, Z2709-Z2711, Z2718-Z2720, Z2726-Z2729, Z2730 (mandible only), Z273 1 , Z2739, Z3 1 5 1 , nms 1044, 1100, rmnh 2615i, 2651b-2651h, 2683, 3087-3089, 3099, 3106, 3107, 3109, 31 10, 31 13- 3115, 3177-3179, 3188, 3229, 4644, 6684, cat. ost. a, cat. ost. i-cat. ost. 1 (4), cat. ost. n, zmb 43867; Karimun (1), Pongka, Kampung— usnm 122848; Kundur(2), no locality-ZRC 4-095, 4-096; Lombok (2), Suranadi— pri 1400, no locality— bm(nh) 1 920. 1 .26.3; Matasiri(l), east coast— usnm 154368;Mwrsfl/tf(l)-ushJMll4561;Mas(5),Sia- ba, Teluk-usNM 121872-121874; no locality - zmb 48417, 48419; Padang{\), north coast— usnm 143583; Sumatra (87), Balbalan-ZMB 34005; Ba- tangkuis— zsbs Coll. Nos. 110, 114; Bengkulu— rmnh 4648; Bukittinggi— zmb 34241; Indragiri district— NHMBa 3774-3776, 3778-3781, zmuz 11668; Kateman, Sungai— usnm 123147; "Kg. Baru"— zsbs Coll. Nos. 62 (adult male), 113; La- hat— NHMBa 1805, 4829; Lampung, Propinsi— zsbs (unnumbered); Lubuksikaping— rmnh 4659; Me- dan— mcz 41 167; zsbs Coll. Nos. 28 (adult male), 67, 151; Mempura— zmb A 14.08 (subadult male), A 14.08/02 (infant); Padang-ZMUz 1 1652; Pagan- san-ZMB A 14.08 (subadult female), A 14.08/03- A 14.08/05; Pangkalan-RMNH 4645-4647; Pang- kalansusu— usnm 536025; Pap-ka— zmb A 14.08; Payakumbuh— rmnh 4649; Rawas— zmb 34025; Rokan-kanan, Sungai-ZMB A 14.08/08, A 14.08/ 1/07, A 14.08/11/01; Serangjaya-hilir-RMNH Coll. Nos. F144, F145; Siak Copatta-SMB A14.08/53; Sijunjung— rmnh w; Soekaranda— zmb A22.05; Sumatra, [east-central]— zmb A 14.08 (infant), A 14.08 (juvenile female), A 14.08 (subadult male); Sumatra, east coast— zsbs Coll. Nos. 157, 165, 169, 175; Tamiang— rmnh Coll. No. F143; Tanjung- morawa— rmnh q, z, unnumbered skeleton, zsbs Coll. Nos. 71 (adult male), 73 (subadult male), 75 (adult female); Wonosobo— rmnh Coll. No. 60; no locality-NMS 1045, 1046, 1101, 16635/IE.2.a, rmnh 1839, 1872a (female), 1872a (male), 3322, 4658, zmb Coll. Nos. 6 Guvenile), 6 (adult male), 15, unnumbered female, unnumbered male, zsbs Coll. Nos. 33, ?5 1 (adult male), 62 (subadult male), 93, 105, 109, 112, 115-117, 120, 202; Sumbawa (4), Dompu— nms 1027, 16633; Semongka— pri 1399; no locality— NHMBa 5341; Tanahmasa (1), no locality— usnm 121836 (external measure- ments recorded on skull tag); Timor (2), Bokong— zsbs 1911/2109; no locality — NHMBa 2703; Tuangku (2), north coast— usnm 1 1441 1, 1 14643 (external measurements recorded on skull tag); is- land unknown (1), Sunda Islands— NHMBa 2981. MALAYSIA, 82. MALAYSIA: SABAH, Borneo (65), Abai-Mcz 25699, 25711, 35569, 35571, 35576, 35578, 35587, 35608, 35619, 35622, 35623, 35626, 35633, 35634, 35643, 35651, 35652, 35655, 35656, 35658, 35661, 35673, 35677, 35681, 35725, 35730, 35734-35736, 35738, 35739, 35741, 35742, 35746, 35748- 35755, 35758 (cranium embedded in plaster), 35759, 35764-35767, 35768/491, 37663, 37664 (external measurements for 50 of these 5 1 speci- mens, all except mcz 35622, are listed on expe- dition record cards); Kinabalu, Mt.— mcz 37781— 37786, 37787-37789 (three mandibles only); Kin- abatangan, Sungai— usnm 19192; Sandakan, 8 mi W— fmnh 33547; Segama, Sungai— zmb A85.10; Talibang— mcz 57836; Tuaran— mcz 37407 (fe- male skull mismatched with male skin). MALA Y- SIA: SARAWAK, Borneo (12), Baram, Batang- ansp 6149, bm(nh) 1894.6.12.13; Belaga or Ka- lulong, Bukit— bm(nh) 1951.67 (male skull mis- matched with female skin), 1 95 1 .69, 1 95 1 .70; Du- lit, Bukit— zmb (unnumbered); Jumpit— zmb A 1870; Kuching-SMK Coll. Nos. 3-315, 3-320; Melinau Gorge— bm(nh), unnumbered mandibu- lar fragment; Mulu, Gunong— bm(nh) 1 894.6. 1 2. 1 ; Puram-ANSP 6113. MALAYSIA: WEST MA- LAYSIA, mainland (5), Benom, Gunong, north- east slope-BM(NH) 1979.2869; Nyalas-ZRC 4-076 (external measurements from skull tag), 4-077 (ex- ternal measurements from Weitzel et al., 1988, p. 107); Telok Anson-NHRM 3259, 3260. PHILIPPINES, 2. Jolo (1), Crater Lake Moun- tain—usnm 1 25324; Mindanao (I), Zamboanga— zmb A2920 (external measurements from Mar- tens, 1876, p. 206). 124 FIELDIANA: ZOOLOGY SINGAPORE, 1. Singapore (1), Botanical Gar- den— zrc 4-085. THAILAND, 4. Mainland (3), Ban Sakaerat- nhrm Coll. No. 1/3; Kampong Biserat— smtd B4346; Pattani— smtd B4348; Tarutao (1), no lo- cality— usnm 123992. VIETNAM, 3. Mainland (3), Cochin China- mnhn 1869.297; Ho Chi Minh City, Botanical Gardens— mnhn 1962.1445; Ho Chi Minh City, Zoological Garden— mnhn 1962.1443. Supplementary information has been derived from 1 2 unexamined specimens of M. f. fascicu- laris collected at the following localities: INDO- NESIA, Borneo: Kalimantan, Kaboerau (Gyld- enstolpe, 1920, pp. 14-15); Loa Bambam (Coll. No. 10, H. C. Raven field catalog, usnm); Long Pangian (Gyldenstolpe, 1920, pp. 13-14); Pelai- hari (Kohlbrugge, 1 896a, p. 185); Java, Kediri dis- trict— zsbs 191 1/2363, 191 1/2364 (specimens not seen, cranial measurements from G. H. Albrecht, pers. comm., Oct. 1991); Tengger, Pegunungan (Kohlbrugge, 1896b, p. 280); Sumatra, Labbu- handeli vicinity (Hagen, 1890, p. 82); Sumbawa, Dompu— nms 16629 (not seen, cranial measure- ments from Mertens, 1936, p. 319, and G. H. Albrecht, pers. comm., Oct. 1991). MALAYSIA: SABAH, Borneo, Abai— mcz 35768/492 (not seen, external measurements from expedition record cards). PHILIPPINES, Mindanao, Zamboanga- 2 specimens (Martens, 1876, p. 206). Macaco fascicularis aurea, Total 55 Skins and Skulls, 44 BURMA, 33. Mainland (23), Ban Sadein— amnh 54968; Haungtharaw-zsi 1 1989; Tagoot, bnhs 5073, 5075; Taungbyauk Chaung-BM(NH) 1936.9.10.8, 1936.9.10.9, 1936.9.10.12, 1936.9.10.13; Tavoy R.-bm(nh) 1936.9.10.10, 1936.9.10.14; Tenas- serim-BM(NH) 1914.12.8.15-1914.12.8.18, bnhs 5070, 5076, 5079, fmnh 82804, zrc 4-02 1-4-023; Wimpong— bm(nh) 1885.8.1.15; Ye Forest— bm(nh) 1910.12.24.1; Kadan (1), no locality— bm(nh) 1925.7.2.1; Kathema (4), no locality— bm(nh) 1936.9.10.4-1936.9.10.7; Lanbi (1), no locality— usnm 104442; Letsok-aw (1), no locali- ty—usnm 124176 (cranium missing); Mibya (2), no locality-BM(NH) 1936.9.10.3, 1910.9.10.11; Zadetkyi (1), no locality— usnm 1 1 1898. THAILAND, 11. Mainland (1 1), Ban Tamrong Phato— ctnrc (catalog number unknown), fmnh 99641, 99642, 99644-99648; Wong, Nam Mae, 65 km E of Um Pang— amnh 54677, bm(nh) 1924.9.2.8; Wong, Nam Mae, 85 km E of Um Pang— amnh 54679. Skins Only, 8 BURMA, 6. Mainland (6), Arakan Div.— zsi 1 1990; Haungtharaw-zsi 4389, 4390, 4392 (skulls inside all three); Mergui, bnhs 5078, zsi 5238. COUNTRY UNKNOWN, 2. "Bengale"-MNHN 362/234 (lectotype), 368/247. Skulls Only, 3. BURMA, 3. Mainland (2), Mergui— bm(nh) 1856.5.6.16; Tenasserim-BM(NH) 1972.1312; Lanbi (1), no locality— usnm 1241 12. Supplementary information has been derived from a published photograph of M. f. aurea ob- served at BANGLADESH, mainland, Whykeong Union Council (M. A. R. Khan, 1985, fig. 15). Macaca fascicularis aurea/ Macaca fascicularis fascicularis Contact Zone or Macaca fascicularis/ Macaca mulatta Contact Zone, Total 45 Skins and Skulls, 41 BURMA, 11. Mainland (10), Pakchan R., near Maliwun— amnh 54972; Pakchan R., near Ban- kachon-BM(NH) 1914.12.8.11, 1914.12.8.14; Thagyet-BM(NH) 1914.12.8.12, 1914.12.8.13, bnhs 5071, 5072, 5074; fmnh 82805, zrc 4-024; Ru (1), no locality— bmnh 5077. LAOS, 3. Mainland (3), Thateng, Muang— amnh 87266, ansp 15136, 15138 (external measure- ments from amnh catalog). THAILAND, 26. Mainland (22), Ban Mae Na Ree-FMNH 99657-99659; Ban Na-535154; Ban Nam Lai Tai— ctnrf (catalog number unknown), fmnh 99660-99663, 99666, 99667; Chumphon, Khlong-ZRC 4-029, 4-030; Nakhon Si Tham- marat— usnm 251661, zrc 4-035-4-037; Pak Chong, Sathani— zrc 4-025, 4-026; Phu Phan— usnm 3077 14 (skin)/307732 (skull); Pran Buri, Mae Nam-ZRC 4-027, 4-028; Naka Yai (1), no local- ity—zrc 4-043; Phayam (1), no locality— zrc 4-038; Rang Yai (1), no locality— zrc 4-042; Yao Noi (1), no locality— zrc 4-052. VIETNAM, 1. Mainland (I), Lac Giao-FMNH 46523. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 125 Skins Only, 3 LAOS, 2. Mainland(2), Thateng, Muang— amnh 87270,87271. THAILAND, 1. Mainland (1), Ban Nam Lai Tai— fmnh 99664 (infant in fluid). Skull Only, 1 THAILAND, 1. Mainland (1), Nakhon Si Thammarat— usnm 251660. M. / philippinensis, Total 98 (Skins and Skulls, 53; Skins Only, 15; Skulls Only, 30) Probably M. /. philippinensis, Total 15 (Skin Only, 1; Skulls Only, 14) M. / philippinensis/ M. f. fascicularis Contact Zone, Total 110 (Skins and Skulls, 82; Skins Only, 5; Skulls Only, 23) Probably M. /. philippinensis/ M. f. fascicularis Contact Zone, Total 12 (Skins and Skulls, 2; Skin Only, 1; Skulls Only, 9) For detailed lists of M. f philippinensis and M. f. philippinensis/ M. f. fascicularis contact zone specimens examined, see Fooden (1991, p. 32). Five revisions have been made to the previously published lists, as follows: 1. bm(nh) 1877.10.6.2, Mindanao: Agusan River, is now recognized as a composite consisting of the skin of an infant or juvenile mismatched with the skull of a subadult female (see above, M. f philippinensis account, Distribution). This skin and skull, formerly assigned to M. f phi- lippinensis, are now tentatively reassigned, as two separate specimens, to "Probably M.f phi- lippinensis/M. f fascicularis Contact Zone." 2. bm(nh) 1 877. 1 0.6. 1 , skin and skull, Mindanao: Surigao, collected near the Agusan River (see above) and formerly assigned to M. f philip- pinensis, is now also tentatively reassigned to "Probably M.f. philippinensis/ M.f fascicularis Contact Zone." 3. bm(nh) 1872.8.20.5, Negros I., S (M.fphilip- pinensis/M. f fascicularis Contact Zone), for- merly known only from its skin, is now known from both skin and skull. 4. bm(nh) 1872.9.20.1, skin and skull, Negros I., S (M.f. philippinensis/M.f fascicularis Contact Zone), not included in previous lists, was ex- amined after those lists were prepared. 5. fmnh 65453, skin only, formerly listed as of unknown origin and undetermined subspecies, is now known to have been collected at Negros: Amio riverbank (M.f. philippinensis/M.f. fas- cicularis Contact Zone). Supplementary information has been derived from two unexamined specimens of M. f philip- pinensis collected at the following localities: PHILIPPINES, Luzon, Nagpartian — usnm 144677 (transferred to Bureau of Science, Manila; external measurements from Hollister, 1913, p. 330); Palawan, Brookes Point— fmnh 62916 (in- fant in fluid, not seen; external measurements from field catalog). Macaco fascicularis umbrosa, Total 8 Skins and Skulls, 8 INDIA, 8. Great Nicobar(3), Kopenheat— usnm 111799; no locality-usNM 111792, 111793; Katchall (2), no locality-usNM 1 1 1801, 1 1 1802; Little Nicobar (3), no locality— usnm 1 1 1795 (ho- lotype), 111796, 111797. Macaca fascicularis fusca, Total 19 Skins and Skulls, 1 2 INDONESIA, 1 2. Simeulue (12), Ajer Dingin- bm(nh) 1923.10.7.1; Dalam, Lhok-usNM 1 14162, 114163, 1 14164 (holotype), 114165-114167; La- buhanbajau — usnm 114169; Sibaboh, Lugu— usnm 114168, 121511-121513. Skins Only, 7 INDONESIA, 7. Simeulue (7), Ajer Dingin- bm(nh) 1923.10.7.2; Sinabang— rmnh Coll. Nos. 20, 140, 1160, 1161;nolocality-RMNH574, 1330. Macaca fascicularis lasiae, Total 2 Skins and Skulls, 2 INDONESIA, 2. Lasia (2), no locality -usnm 114247, 114248 (holotype). 126 FIELDIANA: ZOOLOGY Macaco fascicular is atriceps, Total 1 1 Skins and Skulls, 1 1 THAILAND, 11. Khram Yai(l 1), no locality - bm(nh) 1939.891, 1939.892, usnm 236618- 236621, 236622 (holotype), zrc 4-012, 4-013 (skull)/4-733 (skin), 4-015, 4-016. For external measurements of zrc specimens, cf. Kloss (1921, p. 77) and Weitzel et al. (1988, p. 105). Macaca fascicular is condorensis. Total 16 Skins and Skulls, 1 1 VIETNAM, 11. Ba (3), Ben Dam, 1.2 km W, 0.1 km S-usnm 357242; Ben Dam, 1.7 km W- usnm 357239; Ben Dam, 3.1 km W, 0.6 km S— usnm 357241; Con Son (8), Airfield building, 2.5 km E, 0.7 km N— usnm 357014; Airfield building, 3.3 km E, 0.6 km N— usnm 357349; no locality— bm(nh) 1939.893, 1939.894, 1947.1498 (holo- type), 1955.1528, zrc 4-018, 4-019. Skins Only, 2 VIETNAM, 2. Con Son (2), no locality— mnhn 1882.5 (skull inside), zrc 4-017. Skulls Only, 3 VIETNAM, 3. Ba (1), Ben Dam, 1.7 km W- usnm 357240. Con Son (2), Airfield building, 0.5 km N, 4 km E— usnm 357401; no locality— mnhn 1882.1. Macaca fascicularis tua, Total 4 Skins and Skulls, 4 INDONESIA, 4. Maratua (4), no locality— usnm 197660-197662, 197663 (holotype). Macaca fascicularis karimondjawae, Total 9 Skins and Skulls, 6 INDONESIA, 6. Karimunjawa (6), no locali- ty-MZB 1455-1458, 2717, 2718. Skin Only, 1 INDONESIA, 1. Karimunjawa (1), no locali- ty— rmnh 10608 (holotype). The skull of this spec- imen was subsequently located and examined by G. H. Albrecht (pers. comm., Oct. 1991), who kindly provided the measurement of its greatest length, excluding incisors. Skulls Only, 2 INDONESIA, 2. Karimunjawa (1), no locali- ty— mzb 1454; Kemujan(l), no locality— mzb 2720. Macaca fascicularis, Subspecies Undetermined, Total 95 Skins and Skulls, 1 5 MAURITIUS, 3. Mauritius (3), Plaine Sophie - bm(nh) 1950. 1458; no locality -bm(nh) 1861.6.1.6, 1909.3.12.1. PHILIPPINES, 10. See Fooden (1991, p. 33). THAILAND, 2. Mainland (2), locality un- known-BM(NH) 1860.4.20.2, zsbs Coll. No. 95. Skins Only, 20 1BURMA, 3. Mainland (3), "Rangoon"— rmnh 51/j, 52/k; no locality— bnhs Coll. No. 7. MAURITIUS, 2. Mauritius (2), Kanaka - bm(nh) unnumbered female; no locality— rmnh L-2 (skin inside). PHILIPPINES, 9. See Fooden (1 99 1 , p. 33) and above, M. f. philippinensis/M. f. fascicularis Con- tact Zone, fmnh 65453. THAILAND, 5. Mainland (5), Phet Buh-mnhn 1 220 (lateral facial crest not examined); no local- ity— amnh 37208, mnhn 462, 767, zsbs Coll. No. 89. COUNTRY UNKNOWN, 1. No locality-ZRC 4-123. Skulls Only, 60 1BURMA, 2. Mainland (2), "Bengalen"— nms 1043, zmb 116. MAURITIUS, 1. Mauritius (1), no locality - bm(nh) 1955.12.26.43. PHILIPPINES, 52. See Fooden (1991, p. 34). THAILAND, 5. Mainland {5), Ping, Nam Mae- amnh 5468 1 (external measurements from amnh catalog); "13°45'N, 99°25'E"-bm(nh) 1914.8.22.3, 1914.8.33.4; no locality-zsBS Coll. Nos. 27, 123. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 127 Appendix 2: Gazetteer of Non-Philippine Macaca fascicularis Localities Except as noted previously (see Introduction), locality names listed as primary entries in this gaz- etteer preferentially are official names approved in gazetteers published by the U.S. Board on Geo- graphic Names (USBGN; Bangladesh, 1976; Bur- ma, 1966a; Cambodia, 1971a; India, 1952; In- donesia, 1982; Laos, 1973; Malaysia, Singapore, and Brunei, 1970; Thailand, 1966b; Vietnam, 1964,1971 b); in addition to providing lists of place- names and coordinates, USBGN gazetteers also contain useful glossaries of generic geographic terms for each country. Macaque localities that are not listed in USBGN gazetteers are spelled here as in the original sources. Secondary entries, with cross- references to corresponding primary entries, in- dicate variant spellings or alternate locality names that appear on specimen tags, in published litera- ture, or in unpublished manuscripts on macaques. No secondary entries are supplied for variant spellings of Indonesian place-names that contain "oe" in place of "u." For a gazetteer of Philippine macaque localities, see Fooden (1991, p. 34). The sequence of information presented in pri- mary entries is as follows: 1 . Locality name. 2. Altitude, if reported by collector or observer. 3. Name of island (for noncontinental localities), italicized. 4. Name of country, in capital letters. 5. Coordinates of locality (principal sources— USBGN gazetteers; published or unpublished field notes of collectors or observers). 6. Date of collection or observation. 7. Name of collector or observer. 8. Bibliographic reference (in parentheses) to published or unpublished field notes, if any. 9. Abbreviated name of museum (see Introduc- tion) where specimens are preserved. 10. Number of specimens available (with indi- cation of part preserved, if skin and skull are not both present). 1 1 . Abbreviated subspecific identification: ATR = M.f. atriceps; AUR = M. f. aurea; AUR/ FAS/MUL = M. f aurea/M. f. fascicularis contact zone or M. fascicularis/M. mulatta contact zone; CON = M.f. condorensis; FAS = M.f. fascicularis; FUS = M.ffusca; KAR = M. f karimondjawae; LAS = M. f lasiae; TUA = M.f. tua; UMB = M. f umbrosa. 1 2. Locality code as indicated in distribution maps (Fig. 2A-C). Abai; Borneo, MALAYSIA: SABAH; 5°42'N, 118°23'S; collected 2 Jun.-3 Aug. 1937 by S. L. Washburn and A. H. Schultz (see Coolidge, 1940, p. 124); mcz, 73 (including 1 skin only, 51 skulls only). FAS; C:Sab-19. Acheh, Pulau, MALAYSIA: WEST MALAYSIA; 2°39'-2°41'N, 103°46'-103°47'E; collected 18 Jul. 1915 by H. C. Robinson; bm(nh), 1. FAS; B:SCS-2. Adenare. See Adonara, Pulau. Adonara, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°14'-8°25'S, 123°00'-123°20'E; re- ported present before 1 937 by R. Mertens (1936, p. 319). FAS;C:LS-21. Agung, Gunung, See Desa Poetjang, Gunung Agung. Ai Beta; Pulau Sumbawa, Lesser Sunda Islands, INDONESIA; ca. 8°33'S, 1 17°25'E; blood sam- ples taken Jul. 1981-Jan. 1982 by Y. Kawa- moto, Tb. M. Ischak, and J. Supriatna (1982a, p. 58). FAS; CLS-8. Airabu, Pulau, INDONESIA; 2°43'-2°48'N, 106°12'-106°15'E; observed 18 Aug. 1899 by W. L. Abbott (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 68). FAS; B:SCS-27. Airfield building, 0.5 km N, 4 km E, 80 m; Con Son, VIETNAM; 8°44'N, 106°40'E; collected 5 Mar. 1968 by M. L. Cunningham (see Duncan et al., 1970, p. 180); usnm, 1 (skull only). CON; A:V-7. Airfield building, 2.5 km E, 0.7 km N, [0-590 m]; Con Son, VIETNAM; 8°44'N, 106°37'E; col- lected 28 Feb. 1968 by J. F. Duncan (see Van Peenenetal., 1969, pp. 100,285; 1970, p. 421); usnm, 1. CON; A:V-7. Airfield building, 3.3 km E, 0.6 km N, ca. 160 m; Con Son, VIETNAM; 8°44'N, 106°37'E; col- lected 12 Mar. 1968 by M. L. Cunningham (see Van Peenen et al., 1969, p. 100, 285; 1970, p. 421); usnm, 1. CON; A:V-7. Ajer Dingin; Pulau Simeulue, INDONESIA; 2°29'N, 96°23'E; collected 10 Mar. 1913 by E. Jacobson (1917, p. 276); bm(nh), 2 (including 1 skin only). FUS; B:IO-3. Akyab harbor. See Myengun Kyun. Alas, Lae, between Agusan and Ketambe Research Station; Sumatra, INDONESIA; ca. 3°48'N, 97°33'E; observed Oct. 1984 by M. P. Ghiglieri (1986, p. 106). FAS;B:S-13. Alas, Lae, between Bengkong River and Gelom- bang; Sumatra, INDONESIA; ca. 2°53'N, 128 FIELDIANA: ZOOLOGY 97°53'E; observed Oct. 1984 by M. P. Ghiglieri (1986, p. 108). FAS;B:S-15. Alas, Lae, between Gelombang and mouth of riv- er; Sumatra, INDONESIA; 2°32'N, 97°49'E; observed Oct. 1984 by M. P. Ghiglieri (1986, p. 108). FAS; B:S-16. Alas, Lae, between Lae Renun and Bengkong Riv- er; Sumatra, INDONESIA; ca. 3°02'N, 97°55'E; observed Oct. 1984 by M. P. Ghiglieri (1986, p. 108). FAS;B:S-15. Alas, Lae, between Muara Setulen and Lae Renun; Sumatra, INDONESIA; ca. 3°10'N, 97°55'E; observed Oct. 1984 by M. P. Ghiglieri (1986, p. 106). FAS; B:S-15. Alas Kedaton. See Kukuh. Alas River. See Alas, Lae. Alor, Pulau, Lesser Sunda Islands, INDONESIA; 8°08'-8°28'S, 124°20'-125°08'E; M.fascicularis reported absent before 1937 by J. J. M. F. Sy- mons (Mertens, 1936, p. 319). C:h. Amarassie; Pulau Timor, Lesser Sunda Islands, INDONESIA; 10°20'S, 1 2 3°40'E; collected Jun. 1829 by S. Muller and H. C. Macklot; rmnh, 1 (skin only, skull inside). FAS; GLS-25. Ambawang, Sungai, near Pontianak; Borneo: Ka- limantan, INDONESIA; 0°02'S, 109°42'E; col- lected 1 8 Jun. 1 905 by W. L. Abbott (field catalog; in Lyon, 1907, p. 548); usnm, 1. FAS; CK-2. Ampang area; Pulau Sumbawa, Lesser Sunda Is- lands, INDONESIA; ca. 8°47'S, 1 18°00'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Ka- wamoto (1982, p. 67). FAS; GLS-10. Amphoe Fhang. See Ban Na. Anak Krakatau, Pulau, INDONESIA; 6°06'S, 105°25'E; primates reported absent 1928-1934 by K. W. Dammerman (1948, p. 61); island emerged in 1928. B:k. Anamba. See Siantan, Pulau. Angaur Island, MICRONESIA; 6°53'-6°55'N, 134°08'-134°09'E; introduced population, ob- served Jun.-Aug. 1973 by F. E. Poirier and E. O. Smith (1974, p. 258). Blood samples taken Nov.-Dec. 1986 by Y. Kawamoto, K. Nozawa, K. Matsubayashi, and S. Gotoh (1988, p. 170). Subspecies uncertain; not mapped. Arakan Division; BURMA; 17°22'-21°33'N, 92°10'-94°54'E; collected in 1871 by museum collector; zsi, 1 (skin only, skull inside). Two skulls (2 ID, 2 IE) presented in 1843 by A. P. Phayre (see J. Anderson, 1881, p. 63) are not now in the zsi collection. AUR; A:Bu-3. Aranyaprathet; THAILAND; 13°41'N, 102°30'E; collected 6 May 1937 by J. A. Griswold, Jr. (see Allen & Coolidge, 1940, p. 147); mcz, 1. FAS; A.T-11. Arranya. See Aranyaprathet. Ataran. See Ye Forest, Ataran district, Moulmein region. Ataran River, between Podowk and Kya-eng; BURMA; ca. 16°02'N, 98°00'E; observed 3 Feb. 1 859 by S. R. Tickell (1 859, p. 426). AUR; A:Bu- 11. Atjeh. See Lesten, Daerah Istimewa Aceh. Aur, Pulau, MALAYSIA: WEST MALAYSIA; 2°26'-2°28'N, 104°30'-104°33'E; collected 14 Jun. 1912 by H. C. Robinson; bm(nh), 1. Col- lected 1-8 May 1927 by N. Smedley; zrc, 2 (1 skin only). FAS; B:SCS-5. Babi, Pulau, INDONESIA; 2°03'-2°07'N, 96°37'- 96°42'E; primates reported absent 7-14 Jan. 1902 by W. L. Abbott (see Miller, 1903a, p. 479). B:d. Badang, 400-600 m; Borneo: Kalimantan, IN- DONESIA; 2°45'N, 1 15°43'E; collected 21 May- 1 1 Jun. 1935 by V. von Plessen (see Stresemann, 1938, p. 109); amnh, 5; mzb, 1. FAS; C:K-35. Bahau, Sungai. See Badang. Bako National Park; Borneo, MALAYSIA: SA- RAWAK; ca. 1°40'N, 110°26'E; reported pres- ent in 1971 by G. Rothchild (1971, p. 169). Observed Sep.-Dec. 1980 by R. E. Salter and K. M. Aken (1983, p. 8). FAS; C:Sar-4. Bakong. See Bakung, Pulau. Bakung, Pulau, INDONESIA; 0°0r-0°10'N, 104°23'-104°30'E; reported present 15-22 Jul. 1903 by W. L. Abbott (see Miller, 1906c, p. 283). FAS; B:SCS-12. Balambangan, Pulau, MALAYSIA: SABAH; 7°10'-7°21'N, 1 16°51'-1 17°01'E; observed Feb.- Jun. 1991 by Shukor Md. Nor (pers. comm., 20 Jul. 1991). FAS; C:Sab-8. Balbalan; Sumatra, INDONESIA; not located, 5°38'N-5°57'S, 95°12'-106°05'E; collected 4 Jul. 1904 by W. Volz; zmb, 1 (skull only). FAS; not mapped. Bali, Pulau, Lesser Sunda Islands, INDONESIA; 8°04'-8°48'S, 114°26'-115°42'E; reported pres- ent ca. May 1845 by H. Zollinger (1845, p. 46). FAS; GLS-1 through LS-3. Bali Barat National Park; Pulau Bali, Lesser Sunda Islands, INDONESIA; ca. 8°08'S, 1 14°30'E; re- ported in 1982 by A. H. Robinson and Y. Rus- tandi (1982, p. 14). Observed May-Jun. 1990 by A. Fuentes (in Wheatley et al., 1993, p. 1). FAS; GLS-1. Balipor. See Pajo, Danau Singkarak, Balipor dis- trict. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 129 Baluran Game Reserve; Java, INDONESIA; ca. 7°51'S, 11 4°22'E; observed Sep. 1969-Jan. 1971 by W. Angst (1975, p. 326). FAS; CJ-41. Banda Aceh, ca. 20 km SSW; Sumatra, INDO- NESIA; ca. 5°27'N, 95°16'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-5. Banda Aceh, ca. 30 km ENE; Sumatra, INDO- NESIA; ca. 5°36'N, 95°35'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-6. Banda Aceh, ca. 40 km S; Sumatra, INDONESIA; ca. 5°10'N, 95°19'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1 980, p. 156). FAS;B:S-4. Banda Aceh, ca. 45 km S; Sumatra, INDONESIA; ca. 5°07'N, 95°18'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-4. Banda Aceh, ca. 85 km ESE; Sumatra, INDO- NESIA; ca. 5°17'N, 96°02'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-7. Banda Aceh, near; Sumatra, INDONESIA; ca. 5°34'N, 95°20'E; blood samples taken Nov.-Dec. 1986 by J. R. de Ruiter (1993, p. 91). FAS; B:S-5. Bandar Seri Begawan, near; Borneo, BRUNEI; ca. 4°53'N, 114°56'E; observed ca. 1980 by R. A. Mittermeier (1980, p. 252). FAS; C:B-1. Bandjar. See Tjeringin, near Banjar, Preanger re- gion, east. Bandung; Java, INDONESIA; 6°54'S, 107°36'E collected 29 Apr. 1938 by N. Gosselaar; rmnh 1 (skull only). FAS; B:J-7. Bandung, 2000 ft (= 600 m); Java, INDONESIA ca. 6°54'S, 107°36'E; collected in 1936 by D. P Bosscha Erdbrink; zluu, 1 (skull only); FAS B:J-7. Bandung, near, 700 m; Java, INDONESIA "6°54'S, 107°37'E"; collected 7 Oct. 1971 by NAMRU Djakarta Detachment; usnm, 1. FAS B:J-7. Banggi, Pulau, south, MALAYSIA: SABAH; ca. 7°06'N, 1 17°04'E; collected 4-5 Sep. 1927 by F. N. Chasen and C. B. Kloss (1931, p. 1); zrc, 3. FAS; C:Sab-9. Bangka, Pulau, INDONESIA; r30'-3°07'S, 105°06'-106°51'E; reported present before 1852 by P. Bleeker (1851, p. 527). Collected in 1898 by Blonk; zsbs, 1 (skull only). Collected in 1 900 by A. A. W. Hubrecht (see Kohlbrugge, 1902, p. 322); zluu, 170 (including 158 skulls only, 12 mandibles only). Collected ca. 1936 by H. J. V. Sody (1937, p. 248); rmnh, 3 (including 1 skull only). Reported present Nov. 1971-Jan. 1 973 by C. L. Darsono (Crockett & Wilson, 1 980, p. 156). FAS; B:SCS-18, SCS-19. Bangkaru, Pulau, Kepulauan Banyak, INDO- NESIA; 2°00'-2°07'N, 97°04'-97°09'E; mon- keys reported absent 16-21 Jan. 1902 by W. L. Abbott (see Miller, 1903a, p. 479). B:e. Bangko, ca. 1 5 km ENE; Sumatra, INDONESIA; ca. 1°49'N, 100°55'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-37. Bangko, ca. 20 km ENE; Sumatra, INDONESIA; ca. 1°49'N, 100°58'E; observed Nov. 1971-Jan. 1973 by CM. Crockett and W.L.Wilson (1980, p. 156). FAS; B:S-37. Bangko, ca. 20 km N; Sumatra, INDONESIA; ca. 1°55'N, 100°51'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1 980, p. 156). FAS; B:S-37. Bangko, ca. 25 km ENE; Sumatra, INDONESIA; ca. 1°41'N, 101°02'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson ( 1 980, p. 156). FAS;B:S-37. Bangko, ca. 30 km NNW; Sumatra, INDONESIA; ca. 2°03'N, 100°46'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson ( 1 980, p. 156). FAS; B:S-37. Bangko, ca. 80 km S; Sumatra, INDONESIA; ca. 1°00'N, 100°49'E; reported Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-39. Bangko, ca. 90 km SSE; Sumatra, INDONESIA; ca. POO'N, 101°05'E; reported Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-40. Bangkok; THAILAND; 13°45'N, 100°31'E; col- lected in 1869 by M. Boucourt; mnhn, 1 (skin only, skull inside). FAS; A:T-30. Bangli, near; Pulau Bali, Lesser Sunda Islands, INDONESIA; ca. 8°27'S, 1 15°21'E; observed ca. 1993 by B. P. Wheatley, A. Fuentes, and D. K. Harya Putra (1993, p. 1). FAS; QLS-3. Banguey Island. See Banggi, Pulau, south. Ban Huai Maenam Noi, ca. 15 km SE, ca. 75 m; THAILAND; 14°25'N, 98°51'E; collected 16- 1 7 Feb. 1 967 by J. Fooden ( 1 97 1 , p. 1 6); fmnh, 2. FAS; A:T-26. Banjak Islands. See Tuangku, Pulau and Bang- karu, Pulau, Kepulauan Banyak. Banjar. See Tjeringin, near Banjar, Preanger re- gion, east. Banjoewangi. See Banyuwangi. Banjoe Wetan (? = Banyuwedang), sea level; Pulau 130 FIELDIANA: ZOOLOGY Bali, Lesser Sunda Islands, INDONESIA; ?8°08'S, 114°36'E; collected 18 Jul. 1933 by J. J. Menden; mzb, 3. FAS; OLS-1. Banka. See Bangka, Pulau. Bankachon. See Pakchan River, near Bankachon. Ban Kosum; THAILAND; 15°50'N, 104°00'E; re- ported present before 1978 by B. Lekagul and J. A. McNeely (1977, pp. 293, 725). Subspecies uncertain; A:T-14. Ban Mae Na Ree, ca. 200 m; THAILAND; 16°25'N, 99°23'E; collected 13 Mar. 1967 by J. Fooden ( 1 97 1 , p. 1 8); fmnh, 3. AUR/FAS/MUL; A:T-2. Ban Me Thuot. See Lac Giao. Ban Na; THAILAND; ca. 8°10'N, 100°10'E; col- lected 11 Feb. 1954 by B. Lekagul; usnm, 1. AUR/FAS/MUL; A:T-64. Ban Nam Lai Tai, ca. 5 km W, ca. 300 m; THAI- LAND; 16°10'N, 99°20'E; collected 4-5 Apr. 1 967 by J. Fooden ( 1 97 1 , p. 1 9); ctnrc, 1 ; fmnh, 7 (including 1 infant in fluid). AUR/FAS/MUL; A:T-1. Ban Nong Kok; THAILAND; 8°06'N, 98°52'E; collected 1 1 Jan, 1918 by local collectors em- ployed by H. C. Robinson and C. B. Kloss (1919, p. 8 7); zrc, 1. FAS; A:T-60. Ban Nong Put, ca. 75 m; THAILAND; 8°11'N, 98°53'E; collected 5 Jun. 1973 by J. Fooden ([1975], p. 98); ctnrc, 1. FAS; A:T-60. Ban Pak Nam. See Chumphon, Khlong, mouth. Ban Pak Nam Pho, ? 1 50 m; THAILAND; 1 5°43'N, 100°09'E; collected 2 Apr. 1924 by J. H. Cham- brai [and K. G. Gairdner] (see Kloss, 1919a, p. 49; 1930, p. 61); zrc, 1. Collected 29-30 Aug. 1926 by C. J. Aagaard; mcz, 2. FAS; A:T-5. Ban Palian. See Khao Rang Kai. Ban Phra Muang; THAILAND; 7°18'N, 99°28'E; collected 7 Mar. 1896 by W. L. Abbott (see Riley, 1938, p. 12); usnm, 2. FAS; A:T-56. BanPhuToie, 100-200 m; THAILAND; 14°42'N, 99°07'E; collected 31 Jan. 1967 by J. Fooden (1971, p. 14); fmnh, 2. FAS; A:T-21. Ban Sadein; BURMA; 10°20'N, 98°32'E; collected 18 Feb. 1928 by Faunthorpe-Vernay Expedi- tion; amnh, 1. AUR; A:Bu-22. Ban Sai Kau, Nong Chik region; THAILAND; 6°38'N, 101°08'E;captivesobtained 17 Jun. 1902 by N. Annandale and H. C. Robinson (1903, p. xxxvii; cf. Bonhote, 1903, p. 3); bm(nh), 1. FAS; A:T-69. Ban Sakaerat; THAILAND; 14°30'N, 101°56'E; collected 8 Jan. 1 9 1 2 by N. Gyldenstolpe (1913, p. 3; 1914, p. 1; cf. Thonglongya, 1967, p. 187); nhrm, 1 (skull only). FAS; A:T-10. Banta, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°24'-8°29'S, 119°16'-119°18'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Bantam region. See Camara; Cihara; Cikujang; Pe- labuhanratu, Teluk; and Tamandjaija. Ban Tamrong Phato, ca. 100 m; THAILAND; 14°54'N, 98°31'E; collected 10-1 1 Feb. 1967 by J. Fooden (1971, p. 15); ctnrc, 1; fmnh, 7. AUR; A:T-20. Bantargebang; Java, INDONESIA; 7°01'S, 106°40'E; collected 21-23 Oct. 1909 by O. Bry- ant and W. Palmer; mcz, 2 (including 1 skin only); usnm, 3. FAS; B:J-9. Banten area. See Danau, Rawa. Ban Thap Plik. See Ban Nong Put; Tham Horn, 4 km W of Ban Thap Plik. Ban Thap Plik, 1 km NE, ca. 75 m; THAILAND; 8°1 l'N, 98°53'E; collected 4-5 Jun. 1973 by J. Fooden ([1975], p. 98); fmnh, 3 (including 1 infant in fluid). FAS; A:T-60. Ban Wan; THAILAND; 15°23'N, 104°H'E; ob- served 12-14 Jul. 1989 by N. Aggimarangsee (1992, pp. 109, 120; pers. coram., Oct. 1993). Subspecies uncertain; A:T-17. Ban Wang Kalang. See Khwae Noi, Mae Nam. Ban Wang Phato. See Ban Tamrong Phato. Banyuwangi; Java, INDONESIA; 8°1 2'S, 1 14°2 1 'E; collected 16 Dec. 1919 by unknown collector; rmnh, 1. FAS; CJ-42. Baram, Batang; Borneo, MALAYSIA: SARA- WAK; 4°35'N, 114°00'E; collected 1884-1898 by C. Hose and E. Hose (see Medway, 1 977, pp. 3-4); ansp, 1 (skull only); bm(nh), 2 (including 1 skull only); nms, 1 (skin only); smk, 1 (skin only, skull inside); usnm, 1. Collected Jul. 1896 by W. H. Furness III (1896, p. 310) and H. M. Hiller; ansp, 1 (skull only). FAS; C:Sar-18. Baram district; Borneo, MALAYSIA: SARA- WAK; ca. 4°35'N, 1 13°58'E; collected date un- known by unknown collector; University Mu- seum of Zoology, Cambridge, 1 (skeleton only; not seen, data from P. H. Napier, 1981, p. 14). FAS; C:Sar-18. Baramfluss. See Baram, Batang. Bari; Pulau Flores, Lesser Sunda Islands, INDO- NESIA; 8°21'S, 120°H'E; collected 21 Nov. 1888-9 Jan. 1889 by M. Weber (1890a, p. viii; 1890b, p. 102); museum unknown, 1 (skull only, not seen). FAS; C:LS-16. Barisan Selatan; Sumatra, INDONESIA; ca. 3°00'S, 102°15'E; observed in 1983 by K S. MacKinnon (1986, p. 1 1 2). Observed Jun. 1988 by M. Bismark (1992, p. 11). FAS; B:S-66. FOODEN: SYSTEMATIC REVIEW OF MAC AC A FASCICULARIS 131 Barito, Sungai. See Purukcahu, Sungai Barito. Barito, Sungai, right bank, above Banjarmasin; Borneo: Kalimantan, INDONESIA; ca. 3°13'S, 1 14°32'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chi vers & Burton, [1991], p. 140). FAS; CK-25. Barren Island, Andaman Islands, INDIA; 12°15'- 12°17'N, 93°50'-93°52'E; primates reported ab- sent before 1903 by G. S. Miller, Jr. (1902, p. 792; cf. Kloss, [1928], p. 802; Chaturvedi, 1980, p. 134). A:d. Batam, Pulau, Kepulauan Riau, INDONESIA; 0o58'-l°12'N, 103°54'-104°10'E; reported pres- ent 15-17 Sep. 1 905 by W. L. Abbott (see Miller, 1906c, p. 282). FAS; B:SCS-10. Batang Garut; Sumatra, INDONESIA; 2°23'N, 99°38'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-31. Batangkuis; Sumatra, INDONESIA; 3°40'N, 98°44'E; collected 1 906-1 9 10 by C. Bruegel; zsbs, 1 3 (including 1 0 skins only, 2 skulls only). FAS; B:S-21. Batoe-meringgit; Pulau Bali, Lesser Sunda Is- lands, INDONESIA; not located, 8°04'-8°55'S, 114°26'-115043'E; collected 8 Oct. 1928 by P. F. Franck; mzb, 1 . FAS; not mapped. Battam. See Batam, Pulau. Batu, Tanjung; Pulau Belitung, INDONESIA; ca. 2°56'S, 107°32'E; collected 19-20 Jul. 1904 by W. L. Abbott (in Lyon, 1906, p. 608); usnm, 2. FAS; B.SCS-20. Batu Apoi Forest Reserve. See Kuala Belalong Field Studies Centre. Batu Bara district; Sumatra, INDONESIA; ca. 3°00'N, 99°15'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-23. Batudulang, 800-900 m; Pulau Sumbawa, Lesser Sunda Islands, INDONESIA; 8°36'S, 1 17°19'E; collected 5-8 May 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 79; Mertens, 1930, p. 133; I. Rensch, 1934, p. 226); mzb, 2; nms, 1 (skin only); zmb, 2. FAS; C:LS-8. Batu Islands. See Tanahbala, Pulau; Tanahmasa, Pulau. Batur, Gunung, inside cone; Pulau Bali, Lesser Sun- da Islands, INDONESIA; 8°14'S, 115°23'E; un- confirmed report ca. 1993 by unspecified infor- mant (Wheatley et al., 1993, p. 1). FAS; CLS-3. Baturaden, Gunung Slamet, 700 m; Java, IN- DONESIA; ca. 7°19'S, 109°14'E; collected 9 Oct. 1929 by H. J. V. Sody; rmnh, 1. FAS; B:J-24. Baturaja, ca. 20 km NNE; Sumatra, INDONE- SIA; ca. 4°00'S, 104°17'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-74. Baturaja, ca. 30 km NNE; Sumatra, INDONE- SIA; ca. 3°56'S, 104°20'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-74. Baturaja, ca. 70 km WNW; Sumatra, INDONE- SIA; ca. 3°47'S, 103°39'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p.156). FAS;B:S-73. Baturaja, ca. 75 km NW; Sumatra, INDONESIA; ca. 3°42'S, 103°42'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-73. Bawean, Pulau, INDONESIA; 5°43'-5°53'S, 112°33'-112°44'E; collected 24 Nov. 1907 by W. L. Abbott (in Lyon, 191 1, p. 61); usnm, 3. Collected 6-12 May 1928 by Denin and P. F. Franck; mzb, 6. FAS; C:J-38. Belaga; Borneo, MALAYSIA: SARAWAK; 2°42'N, 1 13°37'E; collected Oct.-Nov. 1932 by Oxford University Expedition to Borneo (see Harrisson, 1933, p. 402; P. H. Napier, 1981, p. 1 4); bm(nh), undetermined portion of 6 speci- mens (including 1 skin only and 3 skulls only) collected at Belaga and/or Kalulong (C:Sar-20). FAS;C:Sar-12. Belawan; Sumatra, INDONESIA; 3°47'N, 98°41'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Kawamoto (1982, p. 67). FAS; B:S-21. Belitung, Pulau, INDONESIA; 2°32'-3°16'S, 107°32'-108°17'E; reported present before 1906 by T. Willink (1905, p. 175). FAS; B:SCS-20. Ben Dam, 1.2 km W, 0.1 km S; Hon Ba, VIET- NAM; 8°39'N, 106°33'E; collected 2 Mar. 1968 by J. F. Duncan (in Van Peenen et al., 1970, p. 421); usnm, 1. CON; A:V-7. Ben Dam, 1.7 km W, ca. 20 m; Hon Ba, VIET- NAM; 8°39'N, 106°33'E; collected 2 Mar. 1968 by P. F. D. Van Peenen (in Van Peenen et al., 1970, p. 421); usnm, 2 (including 1 skull only). CON; A:V-7. Ben Dam, 3.1 km W, 0.6 km S, 100 m; Hon Ba, VIETNAM; 8°39'N, 106°33'E; collected 2 Mar. 1968 by M. L. Cunningham (see Van Peenen et al., 1970, p. 421); usnm, 1. CON; A:V-7. "Bengale"; country unknown; acquired by L. T. Leschenault de la Tour (1820, p. 359; 1822, p. 262), probably in Calcutta, Sep. 1 8 1 9-Jan. 1 820; mnhn, 1 (skin only). Captive acquired 5-27 Apr. 1837 at Diamond's Harbor, near Calcutta, by J. F. T. Eydoux and L. Souleyet (1841, p. xiv, 6); mnhn, 1 (skin only). AUR; not mapped. 132 FIELDIANA: ZOOLOGY "Bengalen"; country unknown; acquired in 1852 by E. Riippell; nms, 1 (skull only). Collected date unknown by Frank; zmb, 1 (skull only). Sub- species uncertain; not mapped. Bengkalis, Pulau, INDONESIA; l015'-l°37'N, 101°59'-102°31'E; reported present 24 Feb.-3 Apr. 1906 by W. L. Abbott (in Lyon, 1908, p. 624). FAS; B:SM-6. Bengkulu; Sumatra, INDONESIA; 3°48'S, 102°16'E; colelcted 1887-1941 by E. Dubois; rmnh, 1 (skull only). FAS; B:S-69. Bengkulu, ca. 40 km ENE; Sumatra, INDONE- SIA; ca. 3°38'S, 102°35'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-70. Bengkulu, Propinsi (province); Sumatra, INDO- NESIA; 2°23'-5°00'S, 101°06'-103°50'E; blood samples taken Jan.-Nov. 1 979 by Y. Kawamoto and Tb. M. Ischak (1981, p. 238). FAS; not mapped. Benom, Gunong; MALAYSIA: WEST MALAY- SIA; ca. 3°50'N, 102°06'E; collected 8 Nov. 1913 by museum collector (see H. C. Robinson & Kloss, 1915b, p. 174); zrc, 2. FAS; B:WM-15. Benom, Gunong, 4000 ft (= 1220 m); MALAY- SIA: WEST MALAYSIA; ca. 3°50'N, 102°06'E; observed 1968-1972 by unspecified informant (Medway, 1972b, p. 120). FAS; B:WM-15. Benom, Gunong, NE slope, 700 ft (= 2 10 m); MA- LAYSIA: WEST MALAYSIA; 3°52'N, 102°1 l'E; collected 1 Feb.-15 Apr. 1967 by Lord Medway (1972a, p. 5; 1972b, p. 120; cf. Whit- more, 1972, p. 12); bm(nh), 1 (skeleton only). FAS;B:WM-15. Benu, Pulau Timor, Lesser Sunda Islands, IN- DONESIA; 9°56'S, 123°59'E; observed 24 Apr. 1930 by Jhr. W. C. van Heurn (1932, p. 65). FAS; C:LS-26. Benua, Pulau, INDONESIA; 0°55'-0°58'N, 107°25'-107°28'E; collected 6 Aug. 1899 by W. L. Abbott (in Miller, 1900, p. 243; cf. Kloss, 1903b, p. 61; Oberholser, 1919, p. 129); usnm, 2. FAS; B:SCS-26. Berangkat, Gunong, east; MALAYSIA: WEST MALAYSIA; ca. 5°10'N, 102°06'E; reported present in 1968 by D. Chivers (1971, p. 80). FAS; B:WM-9. Berapit, Bukit, east; MALAYSIA: WEST MA- LAYSIA; ca. 4°05'N, 103°20'E; reported present in 1968 by D. Chivers (1971, p. 80). FAS; B:WM- 13. Berau, Sungai, north bank; Borneo: Kalimantan, INDONESIA; 2°09'N, 117°29'E; collected 29 Jul. 1912 by H. C. Raven (see Deignan, [1960], p. 267); usnm, 6 (external measurements re- corded in field catalog). FAS; GK-42. Berhala, Pulau, INDONESIA; 3°46'N, 99°31'E; primates reported absent in 1953 by J. L. Har- rison and J. R. Hendrickson (1963, p. 548). B:a. Betotan, ca. 22 mi (= 35 km) SW of Sandakan, 31 m; Borneo, MALAYSIA: SABAH; 5°48'N, 1 1 7°50'E; collected 1 3 Aug. 1927 by F. N. Chas- en and C. B. Kloss (1931, p. 1; cf. Davis, 1962, p. 126;Weitzeletal., 1988, p. 133); zrc, l.FAS; C:Sab-16. Bettotan. See Betotan. Big Kretam River. See Kretam Besar, Sungai. Big Tambelan Island. See Tambelan Besar, Pulau. Bilasodia, Naf River; BANGLADESH; ca. 2 1°05'N,92°12'E; observed Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-l. Billiton. See Belitung, Pulau and Batu, Tanjung; Pulau Belitung. Bima. See Kambing, Pulau. Bimirdia, Naf River; BANGLADESH; ca. 21°05'N,92°12'E; observed Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-l. Binjai, Sungai; Pulau Natuna Besar, INDONE- SIA; 3°47'N, 108°14'E; collected 27 Aug. and 5 Sep. 1928 by F. N. Chasen (1935a, p. 5); zrc, 2. FAS; B:SCS-33. Bintan, Pulau, Kepulauan Riau, INDONESIA; 0°48'-l°13'N, 104°13'-104°34'E; collected 1 1 Jun. 1930 by G. Nunong; zrc, 2. FAS; B:SCS-8. Bintan, Pulau, north coast, Kepulauan Riau, IN- DONESIA; ca. TIO'N, 104°30'E; collected 12 Aug. 1902 by W. L. Abbott (field catalog; cf. Miller, 1906c, p. 282); usnm, 1. FAS; B:SCS-8. Bintang, Pulau. See Bintan, Pulau. Birang, Sungai; Borneo: Kalimantan, INDONE- SIA; 2°1 l'N, 1 17°27'E; collected 3 Oct. 1912 by H. C. Raven (see Deignan, [1960], p. 268); usnm, 1 (skin, skinned head in fluid; external mea- surements recorded in field catalog). FAS; GK- 42. Biserat. See Kampong Biserat. Blambangan-Purwo Nature Park. See South Ban- yuwangi Nature Park. Bliah. See Selatbliat. Bodo, Gili. See Sababi, Pulau. Boegies. See Bugis, Gili. Boekit Sanggul (Benkoelan). See Sanggul, Bukit, Propinsi Bengkulu. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 133 Boeloengan. See Badang; Long Peleben; Sembak- ung, Sungai. Bogor; Java, INDONESIA; 6°35'S, 106°47'E; col- lected 23 Sep. 1928 by G. Schiffer; mzb, 1. FAS; B:J-17. Bohorok district. See Bungara; Bukitlawang. Bokong, 180 m; Pulau Timor, Lesser Sunda Is- lands, INDONESIA; 9°58'S, 124°04'E; collected 24-26 Jul. 191 1 by C. B. Haniel (see Hellmayr, 1914, p. 5); zsbs, 2 (including 1 skin only, 1 skull only). FAS; CLS-26. Bolongs, Outer. See Myengun Kyun. Bolovens, Plateau des. See Thateng, Muang, Pla- teau des Bolovens. Bongao Peak, 70-300 m; Bongao Island, Tawitawi Prov., PHILIPPINES; 5°01'N, 119°45'E; 4 troops observed 24-26 Jan. 1990 by R. P. Ru- bio, P. L. Alviola III, and M. R. Felizardo ( 1 990, p. 29). FAS; C: not numbered (new record, not in Fooden, 1991, p. 2). Bongkabong. See Rugading, near Bongkabong. Bongkabong, near sea level; Borneo, MALAYSIA: SABAH;ca. 5°59'N, 1 16°04'E; collected 21 Aug. 1937 by J. A. Griswold, Jr. (see Coolidge, 1940, p. 123); mcz, 3. FAS; C:Sab-4. Borneo: Kalimantan, INDONESIA; 4°25'N- 4°10'S, 108°50'-119°00'E; collected in 1851 by J. H. Croockewit (see Medway, 1977, p. 3); rmnh, 1 (skin only, skull inside). Collected be- fore 1857 by Verreaux; bm(nh), 1. Collected ca. 1 897 by A. Harrison, Jr., and H. M. Hiller; ansp, 1 . Collected 24 Apr. 1 902 by Dr. Pagel; zmb, 1 (skin only). Collected 1907-1910 by C. Bruegel; zsbs, 67 (including 60 skins only). Collected date unknown by C. Bruegel; zsbs, 13 (including 4 skins only, 9 skulls only). Collected before 1948 by unknown collector; rcs(om), 1 (skull only). Collected 8 Nov. 1961-6 Apr. 1962 by R. E. Kuntz; usnm, 4. Collected date unknown by un- known collector; amnh, 1 (skull only; female skull mismatched with male skin). FAS; not mapped. Borneo: Kalimantan, southeast, INDONESIA; 4°25'N-4°10'S, 108°50'-119°00'E; collected in 1851 by J. H. Croockewit (see Medway, 1977, p. 3); rmnh, 1 (skull only). Collected date un- known by unknown collector; zmb, 1 (skull only). FAS; not mapped. Borneo: Kalimantan, southwest, INDONESIA; 4°25'N^lo10'S, 108°50'-119°00'E; collected in 1836 by S. Muller (see Medway, 1977, p. 3); rmnh, 3 (skins only, skulls inside). FAS; not mapped. Botanical Gardens, near Danau Buyan and Danau Bratan; Pulau Bali, Lesser Sunda Islands, IN- DONESIA; ca. 8°15'S, 115°09'E; observed ca. 1993 by B. P. Wheatley, A. Fuentes, and D. K. Harya Putra (1993, p. 1). FAS; C:LS-3. "Brandan?"; Sumatra, INDONESIA; not located, 5°38'N-5°57'S, 95°12'-106°05'E; collected date unknown by W. Volz; zmb, 1 (skin only). FAS; not mapped. Bratan, Danau, ca. 2500 ft (= 750 m); Pulau Bali, Lesser Sunda Islands, INDONESIA; 8°16'S, 1 15°1 l'E; collected Jan. 191 1 by E. Stresemann; bm(nh), 1 (skin only). Collected date unknown by unknown collector; bm(nh), 1 (skull only; possibly skull of preceding). FAS; C:LS-3. Bratan, [Gunung], 1200 m; Pulau Bali; Lesser Sunda Islands, INDONESIA; 8°15'S, 1 15°12'E; collected 3-8 Feb. 1 938 by V. von Plessen; amnh, 3. FAS; GLS-3. Brunei Bay area; Borneo, BRUNEI; ca. 4°50'N, 115°05'E; observed Nov. 1962-Mar. 1963 by J. A. Kern (1964, p. 185). FAS; C:B-1. Btg. Kwis. See Batangkuis. Bugis, Gili, Lesser Sunda Islands, INDONESIA; 8°30'S, 119°35'E; monkeys reported absent 1 969-1 973 by W. Auffenberg ( 1 98 1 , p. 40). C:d. Buitenzorg. See Bogor. Bukar, Sungai; Borneo, MALAYSIA: SARA- WAK; ca. 1°16'N, 110°28'E; collected 2 Nov. 1919 by C. Ulok; zrc, 1. FAS; C:Sar-5. Bukit Barisan Selatan, Taman Nasional, Lam- pung; Sumatra, INDONESIA; ca. 5°15'S, 104°15'E; observed Jun. 1988 by M. Bismark (1992, pp. 11, 13). FAS; B:S-78. Bukit Cangang; Sumatra, INDONESIA; ca. 0°10'S, 100°05'E; observed 8 Sep. and 20-23 Oct. 1980 by N. Koyama, A. Asuan, and N. Natsir (1981, p. 1). Blood samples taken Jul.-Oct. 1980 by Y. Kawamoto, K. Nozawa, and Tb. M. Ischak (1981, p. 16). FAS; B:S-44. Bukit Cheraka. See Cherakah, Bukit. Bukit Garam; Borneo, MALAYSIA: SABAH; ca. 5°28'N, 1 17°52'E; observed 25 May-23 Jul. 1963 by K. Yoshiba (1964, p. 25). FAS; C:Sab-14. Bukitlawang; Sumatra, INDONESIA; ca. 3°30'N, 98°06'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-20. Bukitlawang, Area I, Bohorok district; Sumatra, INDONESIA; ca. 3°34'N, 98°07'E; 2 troops ob- served 11 Nov. 1980-18 Apr. 1982 by K. J. Gurmaya (1989, p. 66). FAS; B:S-20. Bukitlawang, Area II, Bohorok district; Sumatra, 134 FIELDIANA: ZOOLOGY INDONESIA; ca. 3°33'N, 98°07'E; observed 1 1 Nov. 1980-18 Apr. 1982 by K. J. Gurmaya (1989, p. 66). FAS; B:S-20. Bukitlawang, ca. 35 km SW; Sumatra, INDO- NESIA; ca. 3°15'N, 97°55'E; reported before 1981 by M. Borner (Crockett & Wilson, 1980, p. 156). FAS;B:S-15. Bukitlawang, ca. 65 km S; Sumatra, INDONE- SIA; ca. 2°55'N, 98°07'E; reported Nov. 1971- Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-25. Bukit Timah Nature Reserve; Singapore Island; SINGAPORE; ca. 1°21'N, 103°48'E; observed before 1901 by S. S. Flower (1900, p. 316). Re- ported present in 1941 (Burkhill, 1961, p. 162). Observed 25 Jul. 1986-6 Nov. 1987 by P. W. Lucas (Lucas & Corlett, 1991, p. 202). FAS; B:SCS-7. Bukittinggi; Sumatra, INDONESIA; 0°19'S, 100°22'E; collected date unknown by unknown collector; zmb, 1 (skull only). FAS; B:S-45. Bukittinggi, ca. 20 km S; Sumatra, INDONESIA; ca. 0°23'S, 100°25'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-45. Bukittinggi vicinity; Sumatra, INDONESIA; ca. 0°19'S, 100°22'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-45. Bulan, Pulau, Kepulauan Riau, INDONESIA; 0°55'-l°02'N, 103°51'-103°58'E; collected 5-10 Apr. 1924 by F. N. Chasen (1924b, p. 59); zrc, 2. FAS; B:SCS-15. Bulan, Pulau, [south], Kepulauan Riau; INDO- NESIA; ca. 0°57'N, 103°56'E; collected 22 Mar. 1907 by W. L. Abbott (field catalog; cf. Lyon, 1909, p. 479); usnm, 1. FAS; B:SCS-15. Bulang, Pulau. See Bulan, Pulau. Boloh. See Kampong Sungai Buloh. Bundu Tuhan. See Kampong Bundu Tuhan. Bunga Buah, 1000 m; MALAYSIA: WEST MA- LAYSIA; 3°25'N, 101°45'E; reported as prob- ably present Jul. 1 978-Jun. 1 98 1 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM- 19. Bungara, Area III, Bohorok district; Sumatra, IN- DONESIA; ca. 3°32'N, 98°12'E; 3 troops ob- served 10 Aug. 1983-18 Apr. 1984 by K. J. Gurmaya (1989, p. 66). FAS; B:S-20. Bunguran Island. See Binjai, Sungai; Sinubing; and Ulu, Sungai. "Bungur-Buikt"; Sumatra, INDONESIA; not lo- cated, 5°38'N-5°57'S, 95°12'-106°05'E; collect- ed 2 Dec. 1908 by [H.] Schoede; zmb, 1. FAS; not mapped. Bunoa, Pulo. See Benua, Pulau. Buntok, Sungai Barito, 20 ft (= 6 m); Borneo: Ka- limantan, INDONESIA; 1°42°S, 1 1 4°48°E; col- lected 6 Oct. 1909 by G. C. Shortridge; bm(nh), l.FAS;C:K-31. Burau, Pulau, MALAYSIA: WEST MALAYSIA; 6°21'N, 99°41'E; collected 14 Dec. 1916 by H. C. Robinson (see H. C. Robinson & Kloss, 1910, p. 664); bm(nh), 1. FAS; B:SM-1. Buraw. See Burau, Pulau. [7BURMA]; ?10°00'-21°00'N, 92°00'-98°30'E; collected date unknown by unknown collector; bnhs, 1 (skin only). Subspecies uncertain; not mapped. Bur ni Bebuli, north of Danau Laut Tawar, ca. 2000 m; Sumatra, INDONESIA; ca. 4°40'N, 96°52'E; observed 2 Nov. 1905 by W. Volz (1912, pp. 88, 369). FAS; B:S-9. Busang, Sungai, left bank, 6 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 0°06'S, 1 1 3°57'E; observed 2-4 Sep. 1 986 by D. J. Oliv- ers and K. M. Burton ([1991], p. 143). FAS; CK-19. Busang, Sungai, right bank, 1 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 0°08'S, 1 1 3°58'E; observed 2-4 Sep. 1 986 by D. J. Oliv- ers and K. M. Burton ([1991], p. 143). FAS; CK-19. Butang, Ko, THAILAND; 6°32'N, 99°12°E; col- lected 4 Apr. 1911 by H. C. Robinson; bm(nh), 1. FAS; A:T-58. Buyan, Danau, -Danau Bratan region; Pulau Bali, Lesser Sunda Islands, INDONESIA; ca. 8°15'S, 1 15°09'E; observed ca. 1993 by B. P. Wheatley, A. Fuentes, and D. K. Harya Putra (1993, p. 1). FAS; CLS-3. Cabang Panti Research Station, Gunung Palung National Park; Borneo: Kalimantan, INDO- NESIA; 1°13'S, 110°08'E; observed Jan.-Jun. 1988 by C. H. Cannon and M. Leighton (1994, p. 509). FAS; C:K-1. Cagar Alam Lembah Anai; Sumatra, INDONE- SIA; not located; observed Feb. 1990 by M. Bismark (1992, p. 11). FAS; not mapped. Camara, Bantam region; Java, INDONESIA; 6°36'S, 105°37'E; collected 27 Jul. 1932 by P. F. Franck; mzb, 1. FAS; B:J-13. CAMBODIA; 10°24'-14°41'N, 102°20'-107°38'E; collected [1858-1 86 1] by H. Mouhot (1 864, map at end of vol. 2; cf. Schlegel, 1876, p. 104); rmnh, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 135 1 (skin only). Collected 1 Dec. 1960 by unknown collector; mnhn, 1. FAS; A:C-1 through C-3. "Camboja" or "Cochinchine"; CAMBODIA or VIETNAM; 8°33'-14°41'N, 102°21'-107°39'E; collected ca. 1 878 by M. Pierre (cf. P. H. Napier, 1981, p. 19; undocumented restriction of local- ity to "Tay Ninh"); bm(nh), 2 (skins only). FAS; not mapped. Cameron Highlands, 1 500 m; MALAYSIA: WEST MALAYSIA; 4°28'N, 101°22'E; reported as probably present Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-7. Camorta Island, Nicobar Islands, INDIA; 8°0 1 '- 8°15'N, 93°28'-93°34'E; primates reported ab- sent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, 1903a, p. 1 14). A:k. Camp II, Juara side; Pulau Tioman, MALAYSIA: WEST MALAYSIA; ca. 2°48'N, 104°12'E; ob- served Mar.-Apr. 1 962 by Lord Medway (1966, p. 16). FAS; B:SCS-3. Campbell Bay vicinity; Great Nicobar Island, IN- DIA; ca. 6°59'N, 93°55'E; observed 27 Feb.-8 Mar. 1986 by R. Grubh (see Abdulali, 1967, p. 143). UMB; A:N-4. Candiroto, 600 m; Java, INDONESIA; 7°10'S, 110°03'E; collected 20 Jun. 1929 by H. J. V. Sody; rmnh, 1. FAS; GJ-25. Cape Patani. See Pho, Laem. Cape Rachado Forest Reserve. See Tanjong Tuan, Keramat. Car Nicobar Island, INDIA; 9°07'-9°16°N, 92°44'- 92°51'E; primates reported absent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, 1903a, p. 114). A:i. Catur, Gunung; Pulau Bali, Lesser Sunda Islands, INDONESIA; ca. 8°14'S, 1 15°1 l'E; observed ca. 1993 by B. P. Wheatley, A. Fuentes, and D. K. Harya Putra (1993, p. 1). FAS; CLS-3. Celebes. See Sulawesi, Pulau. Chang, Ko, THAILAND; 1 1°57'-12°09'N, 102°16'-102°28'E; collected 9-11 Dec. 1914 by C. B. Kloss (1916b, p. 28); bm(nh), 1; usnm, 1; zrc, 1. FAS; A:T-35. Changi; Singapore Island, SINGAPORE; 1°23'N, 103°59'E; collected 22 Jul. 1908 by H. C. Rob- inson (in Thomas & Wroughton, 1909b, p. 102) and E. Seimund; bm(nh), 1. FAS; B:SCS-7. Changkat Budiman. See Keroh Forest Reserve. Changkat Mentri; MALAYSIA: WEST MALAY- SIA; 3°44'N, 101°15'E; collected 19-23 Sep. 1918 by C. B. Kloss (see P. H. Napier, 1981, p. 14); bm(nh), 2. FAS; B:WM-18. Chao Phraya, Mae Nam, below Bangkok; THAI- LAND; ca. 13°40'N, 100°33'E; observed Apr.- May 1912 and 1914-1915 by N. Gyldenstolpe (1914, p. 3; 1917b, p. 6). FAS; A:T-30. Cheraka Klang, Bukit. See Cherakah, Bukit. Cherakah, Bukit; MALAYSIA: WEST MALAY- SIA; 3°14'N, 101°23'E; collected 16-17 Nov. 1910 by museum collector; bm(nh), 1; zrc, 1 FAS; B:WM-18. Cheribon. See Cirebon. Cherok Paloh. See Kampong Cherok Paloh. Chiang Mai, northwest; THAILAND; ca. 19°15'N, 98°45'E; unconfirmed report, source unspeci- fied, cited by Varavudhi et al. (1992, p. 338; cf. Fooden, 1971, p. 28). Not mapped. Chiew Lam Reservoir; THAILAND; ca. 9°05'N, 98°40'E; rescued from flooding 1986-1987 by Royal Forest Deparment (Nakhasathien, 1989, p. 150). AUR/FAS/MUL; A:T-47. Chittagong Hill tracts; BANGLADESH; ca. 22°30'N, 92°20'E; improbable report (M. A. R. Khan, 1981, p. 13; 1985, p. 30). Not mapped. Christmas Island, AUSTRALIA; ca. 10°23'- 10°34'S, 105°34'-105°46'E; primates reported absent 1897-1898 by C. W. Andrews (1900, p. 22). Not mapped. Chumphon, Khlong, mouth; THAILAND; 10°22'N, 99°10'E; collected 7 Jul. 1917 by W. J. F. Williamson and M. A. Smith (see Kloss, 1917, p. 289); zrc, 2. AUR/FAS/MUL; A:T- 44. Cihara, Bantam region; Java, INDONESIA; 6°52'S, 106°06'E; collected 1 1 Nov. 1909 by O. Bryant (field catalog); usnm, 2. FAS; B:J-1 1. Cikarang forest; Java, INDONESIA; 6°15'S, 107°09'E; collected 9 May 1858 by J. Zelebor ([1869], p. 7); museum unknown, 1 (not seen). FAS; B:J-19. Cikujang, Bantam region; Java, INDONESIA; 6°41'S, 107°03'E; collected 8 Aug. 1932 by P. F. Franck; mzb, 2. FAS; B:J-20. Cilacap, sea level; Java, INDONESIA; 7°44'S, 109°00'E; collected 19 Oct. 1907 by G. C. Shor- tridge (see Thomas & Wroughton, 1 909a, p. 373); bm(nh), 2. FAS; B:J-1. Cirebon; Java, INDONESIA; 6°44'S, 108°34'E; collected 18 Dec. 1927 by K. Fritsche; nms, 1 (skull only). Collected before 1948, possibly by J. J. Menden; rcs(om), 6 (skulls only). FAS; B:J- 23. Cirebon, 600 m; Java, INDONESIA; ca. 6°44'S, 108°34'E; collected 18 Feb. 1933 by J. J. Men- den; amnh, 2. FAS; B:J-23. Ciremay, Gunung, 600 m (1 specimen) and 800 m (6 specimens); Java, INDONESIA; 6°54'S, 136 FIELDIANA: ZOOLOGY 108°24'E; collected 11 Mar.-8 Apr. 1933 by J. J. Menden; amnh, 7. FAS; B:J-23. Ci Tanduy. See Kalipucang. Ciwangi, 4000 ft (= 1200 m); Java, INDONESIA; 7°04'S, 107°02'E; collected 25 Sep.-l Oct. 1907 by G. C. Shortridge (see Thomas & Wroughton, 1909a, p. 373); bm(nh), 2. 1 FAS; B:J-8. Cochin China [region]; VIETNAM; 8°33'-12°17'N, 104°30'-107°39'E; collected before 1882 by M. Boucard; bm(nh), 1. Collected before 1883 by M. Germain; mnhn, 1 . FAS; A:V-3 through V-5. Cocos Islands. See Tikus, Pulu. Condor, P. See Con Son. Condore, P. See Con Son. Con Son, VIETNAM; 8°40'-8°47'N, 106°32'- 106°39'E; observed 20-28 Jan. 1780 by J. King (1784, p. 462). Observed 22-23 Aug. 1822 by J. Crawfurd (1 828, p. 199). Collected before 1 893 by M. Germain, mnhn, 2 (including 1 skin only, 1 skull only). Collected 19-23 Sep. 1919 by M. A. Smith; bm(nh), 3. Collected 6-18 Nov. 1920 by W. J. F. Williamson and C. Hose (see KJoss, 1926, p. 357; Weitzel et al., 1988, p. 110); bm(nh), 1; zrc, 3 (including 1 skin only). CON; A:V-7. Con Son, west shore, VIETNAM; ca. 8°43'N, 106°34'E; observed 20-25 Mar. 1969 by P. F. D. Van Peenen, M. L. Cunningham, and J. F. Duncan (1970, p. 421). CON; A:V-7. Cox's Bazar Forest Division. See Bilasodia, Bi- mirdia, Ghorardia, Ochodia, and Rukumodia. Dalam, Lhok; Pulau Simeulue, INDONESIA; 2°40'N, 96°08'E; collected 18-26 Nov. 1901 by W. L. Abbott (see Miller, 1903a, p. 437); usnm, 6. FUS; B:IO-2. Damansara; MALAYSIA: WEST MALAYSIA; 3°08'N, 101°38'E; laboratory animals obtained before 1982 by unknown collector (Nordin, 1981, p. 164). FAS; B:WM-19. Dampit. See Wonokojo (?= Wonokerto), Dampit district, southern Malang region. Danau, Rawa; Java, INDONESIA; 6°09'S, 105°59'E; blood samples taken in 1979 by Y. Kawamoto and Tb. M. Ischak (1981, p. 238). FAS;B:J-15. Darvel Bay; Borneo, MALAYSIA: SABAH; 4°50'N, 1 18°30'E; collected 21 Apr. 1909 by Dr. Pagel; zmb, 1. FAS; C:Sab-22. Dasun Tua. See Dusun Tua. Datu, Pulau. See Datuk, Pulau. Datuk, Pulau, INDONESIA; 0°09'-0°10'N, 108°38'-108°39'E; monkeys reported absent 2- 4 May 1907 by W. L. Abbott (in Lyon, 1911, p. 59). B:m. Deli. See Medan. Deli, Pulau, INDONESIA; 7°00'-7o01'S, 105°31'- 105°34'E; introduced in 1987 by C. V. Primates (Indonesia) (advertising leaflet, ca. 1993). Sub- species uncertain; not mapped. Deli, Sungai, between Belawan and Labuhandeli, sea level; Sumatra, INDONESIA; 3°46'N, 98°41'E; collected 17 Mar. 1939 by F. Ulmer (in Miller, 1 942, p. 1 27; cf. Schauensee & Rip- ley, 1940a, p. 311); ansp, 4 (including 2 fetuses in alcohol) FAS; D:S-21. DeLisle Island. See Phayam, Ko. Deli Terbanjawan. See Terbanjawan. Depok; Java, INDONESIA; 6°24'S, 106°50'E; col- lected 17 Jun. and 6 Jul. 1909 by O. Bryant and W. Palmer (see field catalog, usnm); mcz, 2. FAS; B:J-17. Depok, small forest near; Java, INDONESIA; ca. 6°24'S, 106°50'E; observed 1932-1957 by A. Hoogerwerf (1970, p. 408). FAS; B:J-17. Desa Poetjang, Gunung Agung; Pulau Bali, Lesser Sunda Islands, INDONESIA; ca. 8°21'S, 115°30'E; collected ca. 1930 by H. J. V. Sody (1933, p. 93); rmnh, 2. FAS; CLS-3. Dewhurst Bay area; Borneo, MALAYSIA: SA- BAH; ca. 5°35'N, 1 18°35'E; observed May-Jun. 1950 by D. D. Davis (1962, p. 57). FAS; C:Sab- 20. Dindding I.; Borneo, MALAYSIA: SARAWAK; not located, 0°43'-4°58'N, 109°34'-115°38'E; collected before 1906 by H. C. Robinson (see P. H. Napier, 1981, p. 14); bm(nh), 1 (skin only). FAS; not mapped. Dindings. See Hantu, Tanjong, Dindings. Dirk de Vries Bay. See Pangandaran, Teluk Parigi. Desertion Creek, Elephant Point, bank of Irra- waddy River, near Rangoon; BURMA; ca. 16°29'N, 96°20'E; collected before 27 Jan. 1876 by J. Armstrong (see J. Anderson, 1881, p. 63; Khajuria,[1955],p. 109).zsi, 1 (not seen). AUR; A:Bu-6. Djambajan, Sungai. See Jembayan, Sungai. Djapura. See Japura. Djasinga. See Jasinga. Djeboes. See Jebus. Djembrana. See Jembrana. Dolok Tinggi Radja Reserve; Sumatra, INDO- NESIA; ca. 3°07'N, 98°45'E; reported present before 1972 by IUCN (1971, p. 276). FAS; B:S- 27. Domel Island. See Letsok-aw Kyun. Dompu, 66 m; Pulau Sumbawa, Lesser Sunda Is- lands, INDONESIA; 8°32'S, 1 18°28'E; collected 22 Dec. 1909 and in 1910 by J. Elbert (1912, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 137 p. 98); nms, 2 (including 1 skull only). Collected 28 May 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 93; Mertens, 1930, p. 135; I. Rensch, 1934, p. 226); nms, 2 (skulls only; including 1 not examined, measurements from Mertens, 1 936, p. 3 1 9, and G. H. Albrecht, pers. comm., Oct. 1991). FAS; CLS-11. Don Poo Tao; THAILAND; 15°45'N, 104°22'E; observed 16-17 Jul. 1989 by N. Aggimarangsee (1992, pp. 109, 120; pers. comm., Oct. 1993). Subspecies uncertain; A:T-16. Dulit, Bukit, 4000 ft (= 1200 m); Borneo, MA- LAYSIA: SARAWAK; 3°21'N, 114°11'E; col- lected Sep. 1891 and Mar. 1894 by C. Hose (1893, p. 8; altitude cited as 5000 ft); smk, 1 (skin only, skull inside); zmb, 1 (skull only). FAS; C:Sar-13. Durai. See Durian, Pulau. Durian, False. See Sanglang-besar, Pulau. Durian, Pulau, Kepulauan Riau, INDONESIA; 0°42'-0°45'N, 103°42'-103°45'E; reported pres- ent Jun.-Aug. 1903 by W. L. Abbott (see Miller, 1906c, p. 279). Collected 18 Jun. 1923 by P. F. Franck(see Dammerman, 1926b, pp. 282, 302); mzb, 2. FAS; B:SCS-16. Durian-kecil, Pulau, Kepulauan Riau, INDO- NESIA; 0°43'-0°45'N, 103°39'-103°42'E; re- ported present 6-9 Jul. 1903 by W. L. Abbott (see Miller, 1906c, p. 280). FAS; B:SCS-16. Dusun, Sungai, 10 m; MALAYSIA: WEST MA- LAYSIA; 3°40'N, 101°20'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-18. Dusun Tua; MALAYSIA: WEST MALAYSIA; 3°08'N, 101°50'E; collected 3 Oct. 1906 by E. Seimund (see Thomas & Wroughton, 1 909b, p. 101); zrc, 1. FAS; B:WM-19. Eagle's Nest Trail, Kowloon Reservoir Area; UNITED KINGDOM: HONG KONG; ca. 22°21'N, 114°09'E; introduced population, ob- served Jan.-Feb. 1987 by C. H. Southwick and D. Manry (1987, p. 48). Subspecies uncertain; not mapped. Elephant Point. See Desertion Creek, Elephant Point. Empang area. See Ampang area. Endau, Sungai, vicinity; MALAYSIA: WEST MALAYSIA; ca. 2°40'N, 103°38'E; observed 16 Oct.-2 Nov. 1892 by H. W. Lake and H. J. Kelsall (see Kelsall, 1894b, p. 16). FAS; B:WM- 22. Engano Id. See Enggano, Pulau. Enggano, Pulau, INDONESIA; 5°17'-5°32'S, 102°05'-102°24'E; primates reported absent be- fore 1928 by C. B. Kloss ([1928], p. 802; cf. Lyon, 1916, p. 460). Reported present Nov. 1971-Jan. 1973 by unspecified informants (Crockett & Wilson, 1980, p. 156). Pending fur- ther information, Kloss's report is accepted as valid. B:j. Entawa, Tanjong, Sungai Samarahan; Borneo, MALAYSIA: SARAWAK; 1°17'N, 110°29°E; collected 21 Nov. 1919 by H. C. Robinson; bm(nh), 1. FAS; C:Sar-5. E. Siam. See "Siam." False Durian. See Sanglang-besar, Pulau. Fatuboi; Pulau Timor, Lesser Sunda Islands, IN- DONESIA; ca. 8°50'S, 126°20'E; observed 28 Apr.-3 May 1883 by H. O. Forbes (1885, p. 471). FAS;C:LS-29. Flores, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°14'-8°58'S, 119°48'-123°02'E; collected 1854-1862 by A. R. Wallace; bm(nh), 1. FAS; C:LS- 13 through LS- 19. Fort de Kock. See Bukittinggi. Gaik Liew Estate, Damansara; MALAYSIA: WEST MALAYSIA; 3°07'N, 101°37'E; collect- ed 13 Feb. 1921 by "T. H. S."; bm(nh), 1. FAS; B:WM-19. Galang, Pulau, Kepulauan Riau, INDONESIA; 0°42'-0°48'N, 1 04° 1 0'-l 04° 1 8 'E; collected 1 Jan. 1925 by F. N. Chasen; zrc, 4. FAS; B:SCS-1 1. Galang-baru, Pulau. See Nguwal, Pulau. Gantang. See Kantang. Garau, Kampong Kiau region, Mount Kinabalu, ca. 3000 ft (= 900 m); Borneo, MALAYSIA: SABAH; ca. 6°02'N, 116°31'E; collected 18-19 Aug. 1937 by J. A. Griswold, Jr. (1939a, p. 410); mcz, 4. FAS; C:Sab-7. Gasip, Sungai, ca. 1 0 km above mouth; Sumatra, INDONESIA; ca. 0°32'N, 101°44'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Gasip, Sungai, ca. 20 km above mouth; Sumatra, INDONESIA; ca. 0°37'N, 101°43'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Gedangan, Semarang district, 65 m; Java, IN- DONESIA; 7°11'S, 110°41'E; collected Jan. 1929-Oct. 1931 by H. J. V. Sody; rmnh, 7 (in- cluding 4 skulls only). Collected 6 Oct. 1931 by F. A. T. H. Verbeek; rmnh, 1 (skin only). FAS; CJ-29. Gede, Gunung, Pangrango; Java, INDONESIA; 6°47'S, 106°59'E; observed Jan. 1991 byM.Bis- mark(1992, pp. 11, 13). FAS; B:J-18. 138 FIELDIANA: ZOOLOGY Ghirbi. See Ban Nong Kok. Ghorardia, Naf River; BANGLADESH; ca. 21°05'N,92°12'E; observed Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-l. Gili Bodo. See Sababi, Pulau. Gili Lawa Darat. See Lawadarat, Gili. Gili Lawa Laut. See Lawalaut, Gili. Gilimanuk; Pulau Bali, Lesser Sunda Islands, IN- DONESIA; 8°10'S, 114°26'E; collected 8-10 Apr. 1938 by V. von Plessen; amnh, 8. FAS; CLS-1. Gili Mota. See Motang, Gili. Gilla, Pulo. See Jela, Pulau. Ginggo, Teluk; Pulau Rinca, Lesser Sunda Islands, INDONESIA; ca. 8°4 1 'S, 1 1 9°39'E; observed 26 May-6 Jul. 1953 by A. Hoogerwerf (see Auffen- berg, 1981, p. 242). FAS; QLS-13. Gitgit, 530 m; Pulau Bali, Lesser Sunda Islands, INDONESIA; 8°1 l'S, 1 15°08'E; collected 1 Aug. 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 210; Mertens, 1930, p. 145; I. Rensch, 1934, p. 227); nms, 1. FAS; CLS-3. Gn. Telapa Burok. See Telapak Burok, Gunong. Goenoengsetan-Meloewak, 325-520 m; Sumatra, INDONESIA; 3°45'N, 97°40'E; collected 21 Jan. 1937 by A. Hoogerwerf (1941, p. 5; cf. Chasen, 1940b, p. 485); mzb, 1. FAS; B:S-13. Goson Djerong, near, Sungai Makaham, south bank; Borneo: Kalimantan, INDONESIA; ca. 0°29'S, 117°02'E; collected 15 Jun. 1912 by H. C. Raven (see Deignan, [1960], p. 267); usnm, 2 (including 1 skull only). FAS; CK-49. Grabi. See Ban Nong Kok. Great Natuna Island. See Natuna Besar, Pulau. Great Nicobar Island, INDIA; 6°45'-7°14'N, 93°38'-93°57'E; captive obtained Mar. 1858 by J. Zelebor ([1869], p. 7; cf. Fitzinger, 1861, p. 389). Collected 8-12 Mar. 1901 by W. L. Ab- bott; usnm, 2. Observed Apr.-May 1975 by P. K. Das and D. K. Ghosal (1977, p. 265). UMB; A:N-3, N-4. Great Redang Island. See Redang, Pulau. Great Tenasserim River, mouth; BURMA; ca. 12°24'N, 98°37'E; observed before 1852 by F. Mason (1851, p. 220). AUR; A:Bu-18. Gulf of Siam, coast. See Si Racha vicinity. Gumpang, near; Sumatra, INDONESIA; ca. 3°5 1 'N, 97°33'E; blood samples taken Nov.-Dec. 1986 by J. R. de Ruiter (1993, p. 91). FAS; B:S- 13. Gunong Mulu National Park; Borneo, MALAY- SIA: SARAWAK; ca. 4°00'N, 114°55'E; re- ported as food of local residents Jan. 1978 by D. Labang and Lord Medway (1979, p. 56). FAS; C:Sar-21. Gunung Halimun Reserve; Java, INDONESIA; ca. 6°15'S, 106°30'E; observed Jul. 1989 by K. M. Kool (1992, p. 32). FAS; B:J-16. Gunung Leuser Reserve; Sumatra, INDONESIA; ca. 3°45'N, 97°H'E; observed Apr.-Aug. 1970 by F. Kurt (1973, p. 64). FAS; B:S-12. Gunung Palung Nature Reserve; Borneo: Kali- mantan, INDONESIA; ca. 1°13'S, 1 10°08'E; re- ported present before 1 983 by G. Davies ([ 1 983], p. 148). FAS;C:K-1. Hainggyi Kyun, BURMA; 1 5°98'-l 6°0 1 'N, 94°1 T- 94°22'E; reported present before 1880 by J. An- derson (1879, p. 76). AUR; A:Bu-4. Halimun, Gunung. See Gunung Halimun Reserve. Handeuleum, Pulau, INDONESIA; 6°45'S, 105°25'E; 3 individuals captured before 1974 by W. Angst (1973, p. 627). FAS; B:J-14. Hantakan, 3 km N of Pagat, 50 m; Borneo: Ka- limantan, INDONESIA; 2°38'S, 115°27'E; col- lected 2 Feb. 1971 by NAMRU 2 Djakarta De- tachment (see Van Peenen et al., 1974, p. 391); usnm, 1. FAS;C:K-29. Hantu, Tanjong, Dindings; MALAYSIA: WEST MALAYSIA; 4°19'N, 100°34'E; collected 19 Jul. 1918 by unknown collector; zrc, 1. FAS; B:WM-6. Harileko, Batang, ca. 12 km above mouth; Su- matra, INDONESIA; ca. 2°52'S, 104°01'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 24 km above mouth; Su- matra, INDONESIA; ca. 2°48'S, 103°57'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 36 km above mouth; Su- matra, INDONESIA; ca. 2°45'S, 103°52'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 48 km above mouth; Su- matra, INDONESIA; ca. 2°42'S, 103°47'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 60 km above mouth; Su- matra, INDONESIA; ca. 2°38'S, 103°42'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 72 km above mouth; Su- matra, INDONESIA; ca. 2°34'S, 103°37'E; ob- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 139 served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 84 km above mouth; Su- matra, INDONESIA; ca. 2°31'S, 103°32'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Harileko, Batang, ca. 96 km above mouth; Su- matra, INDONESIA; ca. 2°28'S, 103°27'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Hat Chao Mai National Park; THAILAND; ca. 7°20'N, 99°25'E; reported present 17-21 Dec. 1987 by park employees (Boonratana, 1988, p. 76). FAS; A:T-56. Hat Noppharat Thara-Mu Ko Phi Phi National Park; THAILAND; ca. 8°02'N, 98°47'E; re- ported present 9-12 Dec. 1 987 by local residents (Boonratana, 1988, p. 76). FAS; A:T-60. Haungdarau. See Haungtharaw. Haungtharaw; BURMA; 16°30'N, 98°13'E; col- lected 10 Dec. 1880 by Mr. Limborg and J. Anderson; zsi, 4 (including 3 skins only, skulls inside). AUR; A:Bu-10. Haut Padang. See Padang highlands. Henry Larence Island, Andaman Islands, INDIA; 12°05'-12°13'N, 93°03'-93°07'E; primates re- ported absent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, [1928], p. 802; Cha- turvedi, 1980, p. 134). A:c. Hilisimaetano; Pulau Nias, INDONESIA; 0°38'N, 97°44'E; collected 5-6 Jun. 1939 by F. Ulmer (in Miller, 1942, p. 129; cf. Schauensee & Rip- ley, 1940b, p. 399); ansp, 3. FAS; B:IO-8. Ho Chi Minn City; VIETNAM; 10°45'N, 106°40'E; collected in 1929 by T. Roosevelt (see Osgood, 1932, p. 208); fmnh, 1. Collected date unknown by H. Zeltmann; nms, 2 (skins only). FAS; A:V-4. Ho Chi Minh City, Botanical Gardens; VIET- NAM; 10°45'N, 106°40'E; collected Oct. 1926- Sep. 1927 by J. Delacour and W. P. Lowe (see Thomas, 1928, p. 832); bm(nh), 1 (skin only); mnhn, 2 (including 1 skin only, 1 skull only). FAS; A:V-4. Ho Chi Minh City, Zoological Garden; VIET- NAM; 10°45'N, 106°40'E; collected 26 Jul. 1926 and 1 1 Oct. 1926 by unknown collectors; mnhn, 2 (including 1 skull only). FAS; A:V-4. Hong Kong. See Eagle's Nest Trail, Kam Shan Entrance, Kowloon Reservoir Area, and Lower Taipo Road. Huai Ong Sit. See Ban Phu Toie. Huaytakaeng. See Wat Huai Takhaeng. Huay Takang. See Wat Huai Takhaeng. Hutan Lindung Lintau Buo; Sumatra, INDO- NESIA; not located; observed Aug. 1990 by M. Bismark (1992, p. 11). FAS; not mapped. "I. Lendung"; Sumatra, INDONESIA; not locat- ed, 5°38'N-5°57'S, 95°12'-106°05'E; collected data unknown by C. Bruegel; zsbs, 3 (skins only). FAS; not mapped. Indau River. See Endau, Sungai, vicinity. Indragiri, Sungai; Sumatra, INDONESIA; ca. 0°22'S, 103°26'E; collected 21 Sep. 1901 by W. L. Abbott (see Miller, 1902a, p. 143); usnm, 1. FAS; B:S-58. Indragiri district; Sumatra, INDONESIA; 0°10'- 0°55'S, 101°50'-103°30'E; collected 6 Dec. 1898 by G. Schneider (1905, p. 72); zmuz, 1 (skull only). Collected in 1 905 by H. Kummer; NHMBa, 7 (skulls only). FAS; B:S-56. Indragiri (Djapura). See Japura. Indramayu;7ava, INDONESIA; 6°20'S, 108°19'E; collected 30 Jul. 1930 and 29 Mar. 1931 by J. J. Menden; mzb, 4 (including 2 skins only). FAS; B:J-21. Indramojoe. See Indramayu. Irrawaddy River, right bank, northwest of Man- dalay; BURMA; ca. 22°05'N, 96°00'E; captive (presumably introduced) obtained before 1880 by Dr. Marfels (see J. Anderson, 1879, p. 74). Subspecies uncertain; not mapped. Isle de France. See Mauritius Island. Jalor. See Kampong Biserat. Jambi, ca. 60 km NNW; Sumatra, INDONESIA; ca. 1°05'S, 103°31'E; reported before 1981 by M. Borner (Crockett & Wilson, 1980, p. 156). FAS; B:S-59. Jambi, ca. 90 km ENE; Sumatra, INDONESIA; ca. 1°16'S, 104°21'E; reported before 1981 by M. Borner (Crockett & Wilson, 1980, p. 156). FAS; B:S-60. Jambu, tidal creeks near. See Yaring, tidal creeks near. Japura; Sumatra, INDONESIA; ca. 0°19'S, 102°21'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of speci- mens unknown (possibly zmuz 11668, skull only). FAS; B:S-55. Jarak, Pulau, MALAYSIA: WEST MALAYSIA; 3°59'N, 100°06'E; primates reported absent 1950-1960 by J. L. Harrison and J. R. Hen- drickson (1963, p. 548). B:b. Jasinga, Bogor district; Java, INDONESIA; 6°29'S, 106°27'E; collected 10 Apr. 1929 and 4 Sep. 1931 by P. F. Franck; mzb, 2. FAS; B:J-10. Jatibarang; Java, INDONESIA; 6°28'S, 108°17'E; 140 FIELDIANA: ZOOLOGY clinical examination in 1980 by K. Matsubay- ashi and D. Sajuthi (1981, p. 48). FAS; B:J-21. Java, INDONESIA; 5°53'-8°47'S, 105°07'- 114°37'E; collected in 1827 by van Swindern; nms, 1 . Collected in 1 832 by unknown collector; NHMBa, 1 (skull only). Collected in 1 836 by Ouv- ermer-Fischer; nms, 2 (skulls only). Two cap- tives obtained in 1858 by J. Zelebor or von Frauenfeld (see Fitzinger, 1861, pp. 385, 388). Acquired before 1 859 by C. J. Temminck; rmnh, 2 (skulls only). Collected 10 Jan. 1866 by Kok; rmnh, 1 (skin only, skull inside). Collected in 1898 by Bartels; nhrm, 17 (including 16 skulls only, 1 mandible only). Collected date unknown by Bartels; nhrm, 1 (skull only). Collected be- fore 1924 by Tucker; zmuz, 1 (skin, No. 1 1627/ skeleton, No. 1 1630). Collected before 1925 by unknown collector; NHMBa, 3 (skins only). Col- lected 14 Jan. 1932 by L. Heinrath; zmb, 1 (skull only). Collected before 1935 by unknown col- lector; rmnh, 2 (skulls only). Collected in 1936 and 1937 by C. Blazer; rmnh, 30 (including 24 skulls only). Collected before 1942 by E. Du- bois; rmnh, 1 (skull only). Collected date un- known by van Aken; rmnh, 1 (skull only). Col- lected date unknown by Hecht; zmb, 1 . Collected date unknown by T. B. Wilson; ansp, 1 (skull only). Collected date unknown by unknown col- lector; rmnh, 9 (including 6 skins only [skulls inside], 3 skulls only); smtd, 1 (skin only). FAS; not mapped. Java, west, INDONESIA; 5°52'-7°48'S, 105°12'- 108°52'E; collected 1818-1826 by C. L. Blume; rmnh, 2 (including 1 skin only [skull inside], 1 skeleton only). Collected 1 820-1 82 1 by H. Kuhl and J. C. van Hasselt; rmnh, 1 (skeleton only). Collected 1826-1837 by S. Muller and H. C. Macklot; rmnh 2 (skins only, skulls inside). Col- lected in 1863 by P. Diard; rmnh, 5 (including 1 skin only, 1 skeleton only). Collected date un- known by P. Diard; rmnh, 1 (skeleton only). Collected in 1 870 by unknown collector; rmnh, 1 (skin only, skull inside). Examined for para- sites before 1970 by D. Weinman and N. S. Wiratmadja (1969, p. 499). FAS; not mapped. Jebus; Pulau Bangka, INDONESIA; 1°44'S, 105°29'E; collected ca. 1935 by H. J. V. Sody (1937, p. 248); rmnh, 1 (skull only). FAS; B:SCS- 18. Jela, Pulau, INDONESIA; 0°57'-0°58'N, 107°29'- 107°30'E; observed 3-7 Aug. 1899 by W. L. Abbott (in Miller, 1 900, p. 243; cf. Kloss, 1 903b, p. 60). FAS; B:SCS-26. Jeleket, ca. 10 km W; Sumatra, INDONESIA; ca. 3°27'S, 102°15'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-68. Jeleket vicinity; Sumatra, INDONESIA; ca. 3°27'S, 102°20'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-68. Jemaja, Pulau, INDONESIA; ca. 2°49'-3°03'N, 105°41'-105°51'E; observed 16-28 Sep. 1899 by W. L. Abbott (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 74). FAS; B:SCS-28. Jembayan, Sungai; Borneo: Kalimantan, INDO- NESIA; 0°35'S, 117°00'E; collected 6-14 May 1914 by H. C. Raven (see Deignan, [1960], p. 269); usnm, 3 (skulls only). FAS; C:K-49. Jembrana, Negara district; Pulau Bali, Lesser Sun- da Islands, INDONESIA; 8°22'S, 1 14°39'E; col- lected ca. 1930 by H. J. V. Sody (1933, p. 93); rmnh, 2 (including 1 skin not examined). FAS; C:LS-2. Jering region. See Yaring region. Jessleton (= Kota Kinabalu). See Papar; Talibang; Tuaran. Jimaja. See Jemaja, Pulau. Jockaboemi; Java, INDONESIA; not located, 5°93'-8°47'S, 105°12'-1 14°36'E; collected 7 Jun. 1909 by Dr. Biedermann; zmb, 1 (skull only). FAS; not mapped. Johore Archipelago. See Pemanggil, Pulau. Jolir dia, Naf River; BANGLADESH; ca. 20°50'N, 92°18'E; reported present 1982-1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-2. Joloi River. See Julai, Sungai. Juara, Telok; Pulau Tioman, MALAYSIA: WEST MALAYSIA; 2°48'N, 1 04°1 3'E; collected 1 0 Jun. 1906 by H. C. Robinson; bm(nh), 1. Collected 9-10 Sep. 1907 by museum collector; bm(nh), 2. Collected 4-22 Jun. and 1-2 Jul. 1915 by H. C. Robinson; bm(nh), 2; zrc, 2. FAS; Ba:SCS-3. Julai, Sungai, left bank, 1 km below mouth of Sungai Busang; Borneo: Kalimantan, INDO- NESIA; ca. 0°09'S, 1 13°59'E; observed 2-A Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; QK-19. Julai, Sungai, left bank, 2 km above Muarajuloi; Borneo: Kalimantan, INDONESIA; ca. 0°1 l'S, 1 14°03'E; observed 30 Aug.-8 Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; CK-19. Julai, Sungai, left bank, 4 km above Muarajuloi; Borneo: Kalimantan, INDONESIA; ca. 0°10'S, FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 141 1 14°03'E; observed 30 Aug.-8 Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS;C:K-19. Julai, Sungai, right bank, 1 km above mouth of Sungai Busang; Borneo: Kalimantan, INDO- NESIA; ca. 0°1 l'S, 1 13°58'E; observed 1-2 Sep. 1 986 by D. J. Chivers and K. M. Burton ([1 99 1], p. 143). FAS; C:K-19. Julai, Sungai, right bank, 3 km below mouth of Sungai Busang; Borneo: Kalimantan, INDO- NESIA; ca. 0°08'S, 1 14°00'E; observed 30 Aug.- 1 Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; GK-19. Julai, Sungai, right bank, 6 km above Muarajuloi; Borneo: Kalimantan, INDONESIA; ca. 0°09'S, 1 14°02'E; observed 30 Aug.-8 Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; C:K-19. Jumpit [?= Jumbit]; Borneo, MALAYSIA: SA- RAWAK; ?1°07'N, 1 1 P26'E; collected date un- known by Rupert; zmb, 1 (skull only). FAS; C:Sar-10. Juyan, Sungai, right bank, near mouth, Kutai Re- serve; Borneo: Kalimantan, INDONESIA; ca. 0°27'N, 1 17°12'E; observed 13 Oct. 1985 by A. Suzuki (1986, p. 16). FAS; C:K-47. Kaban, Pulau. See Acheh, Pulau. Kaboerau [?= Keburau]; Borneo: Kalimantan, IN- DONESIA; ca. 2°4 1 'N, 1 1 7°07'E; collected 9 Jan. 1914 by C. Lumholtz (see Gyldenstolpe, 1920, p. 3); Zoological Museum, Christiana, 1 (not seen). FAS; CK-39. Kadan, Kyun, 500 ft (= 150 m), BURMA; 12°18'- 12°42'N, 98°18'-98°29'E; collected 3 Oct. 1923 by C. Primrose (see Lindsay, 1926, p. 42); bm(nh), 1. AUR; A:Bu-17. Kaddamayan River. See Tempasuk, Sungai. Kaget, Pulau; Borneo: Kalimantan, INDONESIA; ca. 3°25'S, 1 14°30'E; observed in 1983 by K. S. MacKinnon (1986, p. 112). Observed 1988- 1991 by C. H. Southwick and B. Rosenbaum (1992, p. 88). FAS; CK-25. Kahayan, Sungai, left bank, 60 km above mouth of Sungai Rungan; Borneo: Kalimantan, IN- DONESIA; ca. 1°38'S, 113°56'E; 3 troops ob- served Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; C:K-21. Kahayan, Sungai, left bank, 1 20 km above mouth of Sungai Rungan; Borneo: Kalimantan, IN- DONESIA; ca. 1°08'S, 1 13°54'E; observed Jul. 1984-May 1986 by K M. Burton (see Chivers & Burton, [1991], p. 140). FAS; CK-20. Kahayan, Sungai, left bank, above mouth of Sun- gai Rungan; Borneo: Kalimantan, INDONE- SIA; ca. 2°08'S, 1 1 3°56'E; 2 troops observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; GK-23. Kahayan, Sungai, left bank, below mouth of Sun- gai Rungan; Borneo: Kalimantan, INDONE- SIA; ca. 2°15'S, 114°00'E; observed Jul. 1984- May 1986 by K M. Burton (see Chivers & Bur- ton, [1991], p. 140). FAS; CK-23. Kahayan, Sungai, right bank, 1 20 km above mouth of Sungai Rungan; Borneo: Kalimantan, IN- DONESIA; ca. 1°08'S, 113°52'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; C:K-20. Kahayan, Sungai, right bank, 1 80 km above mouth of Sungai Rungan; Borneo: Kalimantan, IN- DONESIA; ca. 0°54'S, 1 13°12'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; C:K-15. Kahayan, Sungai, right bank, below mouth of Sun- gai Rungan; Borneo: Kalimantan, INDONE- SIA; ca. 2°15'S, 114°00'E; observed Jul. 1984- May 1986 by K. M. Burton (see Chivers & Bur- ton, [1991], p. 140). FAS; CK-23. Kajan, Sungai. See Long Peleben. Kalianda, 100 m; Sumatra, INDONESIA; 5°45'S, 105°38'E; collected 5 Aug. 1 934 by J. J. Menden; amnh, 3. FAS; B:S-87. Kaligoea, Gunung Slamet, 1350 m; Java, IN- DONESIA; ca. 7°14'S, 109°12'E; collected 18 Jan. 1917 by Denin; mzb, 2. FAS; B:J-24. Kalipoetjang. See Kalipucang. Kalipucang, Ci Tanduy, sea level; Java, INDO- NESIA; 7°39'S, 108°44'E; collected 10 Mar. 1908 by G. C. Shortridge (see Thomas & Wroughton, 1909a, p. 373); bm(nh), 3. FAS; B:J-2. Kalulong, [Bukit]; Borneo, MALAYSIA: SARA- WAK; 3°14'N, 114°39'E; collected Oct.-Nov. 1932 by Oxford University Expedition to Bor- neo (see Harrisson, 1933, p. 402; P. H. Napier, 1981, p. 14); bm(nh), undetermined portion of 6 specimens (including 1 skin only and 3 skulls only) collected at Kalulong or Belaga (C:Sar- 1 2). FAS; C:Sar-20. Kambang; Sumatra, INDONESIA; 1°42'S, 100°42'E; collected 19 Nov. 1908 by [H.] Schoede; zmb, 1. FAS; B:S-48. Kambang, Poulo. See Kambing, Pulau. Kambang, Pulau; Borneo: Kalimantan, INDO- NESIA; 3°19'S, 1 14°32'E; photographed before 1968 by Le Roux (see Roedelberger & Gros- choff, 1967, p. 30). FAS; C:K-25. Kambaniru. See Payeti-Kambaniru. Kambas, Wai; Sumatra, INDONESIA; ca. 5°02'S, 142 FIELDIANA: ZOOLOGY 105°52'E; observed in 1983 by K. S. Mac- Kinnon (1986, p. 112). Observed Jun. 1988 by M. Bismark (1992, p. 1 1). Observed Jan.-Mar. 1989 by A. Yanuar and J. Sugardjito (1993, p. 34). FAS; B:S-84. Kambera. See Payeti-Kambaniru. Kambing, Pulau; Pulau Sumbawa, Lesser Sunda Islands, INDONESIA; 8°27'S, 118°42'E; col- lected 28 Jul. 1 927 by Sunda-Expedition Rensch (see Mertens, 1930, p. 144; M. fascicularis re- portedly introduced in Pulau Kambing from Bima by Sultan of Bima); mzb, 1. FAS; C:LS- 12. Kambing, Pulau, Lesser Sunda Islands, INDO- NESIA; 10°14'S, 123°26'E; collected Jun. 1829 by S. Muller and H. C. Macklot; rmnh, 1 (skin only). FAS; CLS-24. Kamoedian, Pulau. See Kemujan, Pulau. Kamorta Island. See Camorta Island. Kampong Biserat; THAILAND; 6°32'N, 101°14'E; collected 16 May 1901 by N. Annandale and H. C. Robinson (1903, p. xxv; cf. Bonhote, 1903, p. 3); bm(nh), 3 (including 1 skin only); smtd, 1 (skull only). FAS; A:T-69. Kampong Bundu Tuhan, Mount Kinabalu; Bor- neo, MALAYSIA: SABAH; 5°59'N, 116°32'E; collected 30 Jun.-l 8 Jul. 1 95 1 by D. H. Johnson (see Coolidge, 1940, p. 124; Davis, 1962, p. 126); usnm, 4. FAS; C:Sab-7. Kampong Cherok Paloh; MALAYSIA: WEST MALAYSIA; 3°37'N, 103°23'E; laboratory an- imals obtained 1954-1959 by unknown collec- tor (Price, 1959, p. 499). FAS; B:WM-13. Kampong Hadjak, Muaratewe district; Borneo: Kalimantan, INDONESIA; ca. 0°57'S, 1 14°53'E; collected 5 May 1932 by S. A. R. le Prince Le- opold (see Frechkop, 1934, p. 25); irsn, 1 (skin only). FAS; CK-32. Kampong Kiau. See Garau; Kiaulan; Tempasuk, Sungai; Tinonkok. Kampong Kiau, 3000 ft (= 900 m); Borneo, MA- LAYSIA: SABAH; ca. 6°02'N, 116°31'E; col- lected 24-29 Apr. 1929 by F. N. Chasen (1931, p. 6); zrc, 2. FAS; C:Sab-7. Kampong Kiau, ca. 2200 ft (= 670 m); Borneo, MALAYSIA: SABAH; ca. 6°02'N, 1 1 6°3 1 'E; col- lected 13-23 Aug. 1937 by J. A. Griswold, Jr. (1939b, p. 514; mcz Asiatic Primate Expedition specimen list); mcz, 2. FAS; C:Sab-7. Kampong Kiau-Tenampok Pass, trail between, [ca. 1100 m]; Borneo, MALAYSIA: SABAH; ca. 6°00'N, 116°30'E; observed 3 Jun. 1932 by J. A. Griswold, Jr. (1939a, p. 410). FAS; C:Sab-7. Kampong Menuggol; Borneo, BRUNEI; 4°53'N, 114°59'E; observed in 1972 by D. Macdonald (1982, pp. 62, 71). FAS; C:B-1. Kampong Mukut, hillsides above; Pulau Tioman, MALAYSIA: WEST MALAYSIA; ca. 2°43'N, 104°11'E; observed Mar.-Apr. 1962 by Lord Medway (1966, p. 16). FAS; B:SCS-3. Kampong Punkah. See Pongka, Kampung. Kampong Rantau Panjang; MALAYSIA: WEST MALAYSIA; ca. 2°53'N, 101°29'E; examined for malaria before 1 95 1 by E. P. Hodgkin ( 1 950, p. 326). Observed Jul.-Sep. 1960 by Y. Furuya (1965, p. 287). FAS; B:WM-18. Kampong Sungai Buloh, 1 50-300 ft (= ca. 70 m); MALAYSIA; WEST MALAYSIA; 3°15'N, 101°18'E; observed 1947-1957 by J. L. Harri- son (1969, p. 176). FAS; B:WM-18. Kampong Tanah Puteh, Pekan district; MALAY- SIA: WEST MALAYSIA; 3°35'N, 103°22'E; ex- amined for malaria ca. 1961-1962 by R. H. Wharton, D. E. Eyles, McW. Warren, and W. H. Cheong (1964, p. 58). FAS; B:WM-18. Kampong Tenghilan. See Tuaran-Kampong Tenghilan Road, new bridge. Kampong Titi Tinggi; MALAYSIA: WEST MA- LAYSIA; 6°38'N, 100°1 5'E; captive obtained in 1964 by local trappers (Collins et al., 1970, p. 509). FAS;B:WM-1. Kampon Kadjak. See Kampong Hadjak. Kampung Baru Study Area; Borneo: Kalimantan, INDONESIA; 1°06'S, 110°10'E; reported pres- ent 2 Apr.-9 May 1986 by Y. Ruhiyat (1986, p. 60). FAS; CK-1. Kam Shan Entrance, Kowloon Reservoir Area; UNITED KINGDOM: HONG KONG; ca. 22°22'N, 114°09'E; introduced population, ob- served Jan.-Feb. 1987 by C. H. Southwick and D. Manry (1987, p. 48). Subspecies uncertain; not mapped. Kanaka; Mauritius Island, MAURITIUS; 19°59'- 20°31'S, 57°18'-57°48'E; introduced popula- tion, collected 29 May 1975 by R. L. Ciochan; bm(nh), 1 (skull not examined). Subspecies un- certain; not mapped. Kangean, Pulau, 4 ft (= 1 m), INDONESIA; 6°49'- 7°00'S, 1 1 5°1 2'-l 1 5°34'E; collected 1 5-23 Nov. 1909 by G. C. Shortridge; bm(nh), 4. FAS; C:J- 40. Kantan, See Kantang. Kantang; THAILAND; 7°25'N, 99°31'E; collected 1 1 Jan. 1 9 1 8 by C. B. Kloss (see H. C. Robinson & Kloss, 1910, p. 668; Weitzel et al., 1988, p. 104; W. L. Abbott, 1897, unpubl. map in usnm archives); zrc, 1. FAS; A:T-56. Kapos Tinggi; Pulau Bengkalis, INDONESIA; ca. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 143 1°27'N, 102°18'E; collected 22 Mar. 1906 by W. L. Abbott (field catalog; in Lyon, 1908, p. 623); usnm, 1. FAS; B:SM-6. Kapuas, Sungai. See Semitau, Sungai Kapuas. Kapuas, Sungai, right bank, 25 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 2°48'S, 1 14°17'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chi vers & Burton, [1991], p. 140). FAS; C:K-24. Karakit, Pulau Banggi, MALAYSIA: SABAH; 7°07'N, 117°05'E; collected 22 Jun. 1991 by Shukor Md. Nor (pers. comm., 17 Jul. 1991); fmnh, 1. FAS; C:Sab-9. Karangan, Sungai; Borneo: Kalimantan, INDO- NESIA; 1°19'N, 117°55'E; collected 1 Nov. 1913 by H. C. Raven (see Deignan, [1960], p. 269); usnm, 1 (skull only). FAS; CK-44. Karangintan, hilly country, Sungai Martapura, 30 ft (= 1 0 m); Borneo: Kalimantan, INDONESIA; 3°26'S, 1 14°55'E; collected 19 Aug. 1909 by G. C. Shortridge; bm(nh), 1. FAS; CK-27. Karanginton. See Karangintan. Karangmumus, Sungai, near Samarinda; Borneo: Kalimantan, INDONESIA; 0°30'S, 117°09'E; collected 3 Jul. 1912 by H. C. Raven (field cat- alog); usnm, 1. FAS; GK-49. Karang Tigan. See Karangtigau, Tanjung. Karangtigau, Tanjung; Borneo: Kalimantan, IN- DONESIA; 2°26'N, 1 18°00'E; collected 4 Aug. 1 9 1 2 by H. C. Raven (field catalog; cf. Deignan, [1960], p. 267); usnm, 2 (including 1 skin only). FAS; C:K-40. Karawassen. See Kawarasan. Karimata. See Pai, Teluk; Pulau Karimata. Karimon Anak. See Karimun-kecil, Pulau. Karimon Djawa. See Karimunjawa, Pulau. Karimon-Djawa, Pulau, (P. Kamoedian). See Ke- mujan, Pulau. Karimunjawa, Pulau, INDONESIA; 5°50'-5°53'S, 1 10°25'-1 10°29'E; collected 7-14 May 1926 by K. W. Dammerman, Denin, and P. F. Franck; mzb, 5 (including 1 skull only). Collected 26-28 Nov. 1930 by W. Romswinckel; mzb, 2; rmnh, 1 (skull not examined). KAR: C:J-27. Karimun-kecil, Pulau, Kepulauan Riau, INDO- NESIA; 1°07'-P10'N, 103°22'-103°25'E; re- ported present Jun.-Aug. 1903 by W. L. Abbott (see Miller, 1906c, p. 277). FAS; B:SCS-17. Kariorang; Borneo: Kalimantan, INDONESIA; 0°50'N, 117°52'E; collected in 1903 by M. Schmidt; zmb, 2 (skulls only). FAS; C:K-45. Kaser Doo Wildlife Sanctuary; BURMA; 13° 15'- 13°24'N, 98°45'-99°00'E; reported present ca. 1994 (Anonymous, 1994, p. 12). Subspecies un- certain; A:Bu-16. Kata Taek, ca. 200 m; THAILAND; ca. 15°28'N, 99°23'E; collected 2-3 Mar. 1967 by J. Fooden (1971, p. 17); fmnh, 6. FAS; A:T-19. Katchal Island. See Ol-kolo-kwak vicinity; Katch- all Island. Katchall Island, Nicobar Islands, INDIA; 7°52'- 8°02'N, 93°18'-93°27'E; observed before 1847 by P. Barbe (1846, p. 365). UMB; A:N-1. Kateman, Sungai, < 2 ft (= < 1 m); Sumatra, INDONESIA; 0°12'N, 103°20'E; collected 8 Sep. 1903 by W. L. Abbott (in Lyon, 1908, p. 625); usnm, 1 (skull only). FAS; B:S-57. Kathema Kyun, BURMA; 13°39'N, 98°12'E; col- lected 20-22 Apr. 1 936 by H. C. Smith; bm(nh), 4. AUR; A:Bu-15. Katingan, Sungai, left bank, 1 60 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 1°47'S, 1 13°19'E; observed Jul. 1984-May 1986 by K M. Burton (see Chivers & Burton, [1991], p. 140). FAS;C:K-13. Katingan, Sungai, left bank, 200 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 1°30'S, 1 13°10'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS;C:K-14. Katingan, Sungai, right bank, 1 40 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 2°04'S, 1 13°25'E; observed Jul. 1984-May 1986 by K M. Burton (see Chivers & Burton [ 1 99 1 ], p. 1 40). FAS;C:K-11. Kawarasan; Java, INDONESIA; 7°49'S, 1 12°07'E; collected 20 Apr. 1895 by unknown collector; rmnh, 1 (skin only). FAS; CJ-33. Kayan, Sungai. See Long Peleben. Kebun Percobaan Haurbentes, Areal; Java, IN- DONESIA; not located; obseved Oct. 1989 and Jan. 1991 by M. Bismark (1992, p. 11). FAS; not mapped. Kediri. See Manggis; Margomulio, Gunung. Kediri district; Java, INDONESIA; ca. 7°48'S, 1 12° 15 'E; collected 13 Nov. 1910 and 25 Nov. 1911 by V. Arnim; zsbs, 2 (skulls only; not seen, data from G. H. Albrecht, pers. comm., Oct. 1991). FAS; GJ-33. Kelabit uplands, 4000 ft. See Dulit, Bukit, 4000 ft. Kelabong, Bukit, vicinity; Sumatra, INDONE- SIA; ca. 3°13'S, 102°26'E; reported prsent Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:S-67. Kelang Road, 7.25 km; MALAYSIA: WEST MA- 144 FIELDIANA: ZOOLOGY LAYSIA; ca. 3°05'N, 101°40'E; collected 2 Dec. 1953 by Scrub Typhus Research Unit; bm(nh), 1. FAS; B:WM-19. Kelapa, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°39'-8°42'S, 119°13'-119°14'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:e. Kembangjanggut; Borneo: Kalimantan, INDO- NESIA; 0°08'N, 116°22'E; collected 28 Nov. 1956 by A. M. R. Wegner and Saan; mzb, 1. FAS; CK-48. Kemujan, Pulau, INDONESIA; 5°47'-5°51'S, 1 10°27'-1 10°30'E; reported present before 1906 by T. Willink (1905, p. 175). Collected 25 Nov. 1930 by W. Romswinckel; mzb, 1 (skull only). KAR; C:J-27. Kenepai, Gunung. See Roema Manoeal, south foot of Gunung Kenepai. Keningau, 800 ft (= 250 m); Borneo, MALAYSIA: SABAH; 5°20'N, 1 16°10'E; collected Sep. 1960 by R. E. Kuntz (1969, p. 193); amnh, 1. FAS; C:Sab-6. Kenokok, 3300 ft (= 1000 m); Borneo, MALAY- SIA: SABAH; ca. 6°04'N, 116°28'E; collected 25 Apr. 1929 by F. N. Chasen (1931, p. 6); zrc, 1. FAS;C:Sab-7. Kenyam, Sungai, 100 m; MALAYSIA: WEST MALAYSIA; 4°31'N, 102°28'E; observed Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-1 1. Keramat. See Tanjong Tuan, Keramat. Kerinci. See Sandaran Agong, Kerinci region; Siu- lakderas, Kerinci region. Kerinci, Gunung, northern foot; Sumatra, IN- DONESIA; 1°40'S, 101°20'E; observed Jun. 1985-Mar. 1986 by T. Oi (1986, p. 73). FAS; B:S-52. Keroh Forest Reserve; MALAYSIA: WEST MA- LAYSIA; ca. 4°13'N, 101°07'E; observed 2 Jun. 1935 by R. C. Morris (1936, p. 440). FAS; B:WM-16. Kertau, Bukit; MALAYSIA: WEST MALAYSIA; ca. 3°27'N, 102°37'E; examined for malaria 1965-1969 by McW. Warren, W. H. Cheong, H. K. Fredericks, and G. R. Coatney (1970, p. 386). FAS; B:WM-14. Ketambe, ca. 5 km S; Sumatra, INDONESIA; ca. 3°38'N, 97°40'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-13. Ketambe, ca. 10 km S; Sumatra, INDONESIA; ca. 3°36'N, 97°40'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-13. Ketambe, ca. 30 km SSE; Sumatra, INDONESIA; ca. 3°27'N, 97°50'E; reported present before 1 98 1 by M. Borner (Crockett & Wilson, 1980, p. 1 56). FAS; B:S-14. Ketambe Research Station, Gunung Leuser Na- tional Park, ca. 350 m; Sumatra, INDONESIA; 3°40'N,97°40'E; observed Jun. 1971-Aug. 1974 by H. D. Rijksen (1978, p. 1 1 1). Hormonal study Oct.-Nov. 1987 by C. P. van Schaik, M. A. van Noordwijk, T. van Bragt, and M. A. Blanken- stein (1991, p. 347). FAS; B:S-13. "Kg. Baru"; Sumatra, INDONESIA; not located (USBGN Gazetteer of Indonesia, 1982, p. 483, lists 2 populated places named "Kampungbaru" in Sumatra), 5°38'N-5°57'S, 95°12'-106°05E; collected in 1906 and 1910 by C. Bruegel; zsbs, 10 (including 5 skins only, 2 skulls only). FAS; not mapped. Khangkhao, Ko, THAILAND; 13°06'-13°07'N, 100°48'-100°49'E; reported present 15 Nov. 1990 by local boatman (Aggimarangsee, 1992, pp. Ill, 130; pers. comm., Oct. 1993). Subspe- cies uncertain; A:T-3 1 . Khao Khieo Wildlife Sanctuary; THAILAND; 13°13'N, 101°04'E; introduced population, re- leased before Jan. 1977, observed Jun. 1977- Jun. 1978 by P. J. Storer (1979, pp. 28, 46). Subspecies uncertain; not mapped. Khao Lampi-Hat Thai Muang National Park; THAILAND; ca. 8°23'N, 98°20'E; reported present 21-25 Nov. 1987 by local residents (Boonratana, 1988, p. 76). AUR/FAS/MUL; A:T-51. Khao Naw. See Wat Khao Noh. Khao Ngu. See Wat Ratch Singkhorn. Khao Noh. See Wat Khao Noh. Khao Noi/Khao Tangkuan; THAILAND; 7°1 3'N, 100°36'E; blood samples taken Aug.-Sep. 1988 by P. Varavudhi, J. Suzuki, U. Yodyingyuad, T. Nootprapand, V. Yodyingyuad, Y. Chaiseha, K. Suwanprasert, and W. Settheetham (1989b, p. 77; cf. Kawamoto et al., 1989, p. 95). Ob- served 9-14 Apr. 1989 by N. Aggimarangsee (1992, pp. 109, 139; pers. comm., Oct. 1993). FAS; A:T-67. Khao Noi-Khuo Tanguan. See Khao Noi/Khao Tangkuan. Khao Paskowee. See Khao Phatowee. Khao Phatowee; THAILAND; ca. 15°28'N, 99°45'E; observed 25 Feb. 1967 by J. Fooden (1971, p. 16). Observed 1973-1974 by A. A. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 145 Eudey (1979, p. 90; 1994, p. 274). FAS; A:T- 22. Khao Rang Kai, 4 km E. of Ban Palian, 75 m; THAILAND; 7°19'N, 99°48'E; collected 6 Jul. 1973 by J. Fooden ([1975], p. 98); ctnrc, 1; fmnh, 1. FAS; A:T-62. Khao Sam Muk. See Sammuk, Khao. Khao Sam Roi National Park; THAILAND; 12°05'-12°17'N, 99°53'-100°02'E; reported present before 1994 by J. W. K. Parr, N. Ma- hannop, and V. Charoensiri (1993, p. 245). AUR/FAS/MUL; A:T-40. Khao Suan Luang; THAILAND; 1 3°35'N, 99°45'E; observed 24 Dec. 1990 by N. Aggimarangsee (1992, pp. Ill, 128; pers. comm., Oct. 1993). FAS; A:T-28. Khao Wang; THAILAND; 13°06'N, 99°56'E; ob- served 12-13 Aug. 1989 and 9-10 Feb. 1991 by N. Aggimarangsee (1992, pp. 109, 112, 130); pers. comm., Oct. 1993). Subspecies uncertain; A:T-37. Khram Yai, Ko, THAILAND; 12°40'-12°43'N, 100°45'-100°48'E; collectd 30 Oct. 1916 by C. B. Kloss (1919c, p. 335); bm(nh), 2; usnm, 5; zrc, 2. Collected 1 Jun. 1917 by W. J. F. Wil- liamson; zrc, 2. ATR; A:T-39. Khulna disrict, Sundarbans; BANGLADESH; 22°20'N, 89°27'E; improbable report (Anony- mous, 1977, p. 14; cf. Sarker & Sarker, 1984, p. 9; M. A. R. Kahn, 1985, p. 30). Not mapped. Khwae Noi, Mae Nam, right bank, ca. 10 km be- low Ban Wang Kalang; THAILAND; ca. 15°03'N, 98°30'E; observed 3 Feb. 1967 by J. Fooden (1971, p. 15). AUR; A:T-20. Kiambang; Sumatra, INDONESIA; 5°27'S, 105°44'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-86. Kiau. See Garau; Kampong Kiau; Kampong Kiau- Tenampok Pass, trail between; Kiaulan; Tem- pasuk, Sungai; and Tinonkok. Kiaulan, Kampong Kiau region, ca. 2200 ft (= 670 m); Borneo, MALAYSIA: SABAH; ca. 6°02'N, 116°31'E; collected 17 Aug. 1937 by J. A. Gris- wold, Jr. (1939b, p. 514); mcz, 2. FAS; C:Sab-7. Kinabalu, Mount; Borneo, MALAYSIA: SABAH; ca. 6°05'N, 1 16°33'E; collected 31 May-28 Aug. 1937 by J. A. Griswold, Jr. (1 939a, p. 402; 1 939b, p. 504); mcz, 9 (including 6 skulls only, 3 man- dibles only). FAS; C:Sab-7. Kinabalu National Park; Borneo, MALAYSIA: SABAH; ca. 6°05'N, 1 16°35'E; reported present before 1983 by G. Davies([1983],p. 148). FAS; C:Sab-7. Kinabatangan, Sungai; Borneo, MALAYSIA: SA- BAH; ca. 5°42'N, 1 18°23'E; collected 19-23 Dec. 1887 by C. F. Adams (see Medway, 1977, p. 4); usnm, 2 (including 1 skull only). FAS; C:Sab- 19. King's Island. See Kadan, Kyun. Kinta, Daerah (district); MALAYSIA: WEST MALAYSIA; ca. 4°30'N, 101°12'E; collected before 1 90 1 by unknown collector (Rower, 1 900, p. 316); specimens reportedly in Taiping Mu- seum (not seen). FAS; B:WM-7. Klaeng. See Wang Kaew. Klang Road, 4V2 miles. See Kelang Road, 7.25 km. Klang Straits. See Pintu Gedong, Pulau. Klet Kaeo, Ko, THAILAND; 12°46'N, 100°51'E; trapped spring 1967 by G. Berkson (1970, p. 286). Subspecies uncertain; A:T-39. Kloet, Goenoeng. See Manggis, Gunung Kelud; Margomulio, Gunung. Klong Pah Yie; Ko Samui, northwest, THAI- LAND; ca. 9°32'N, 99°57'E; collected 7 May 1913 by H. C. Robinson (1915, p. 129) and E. Seimund; zrc, 1. FAS; A:T-49. Klumpang, Teluk; Borneo: Kalimantan, INDO- NESIA; ca. 3°00'S, 1 16°12'E; observed 8 Jan.- 13 Mar. 1908 or 18-19 Apr. 1909 by W. L. Abbott (in Lyon, 191 1, p. 58). FAS; GK-51. Kode, Nusa, Lesser Sunda Islands, INDONESIA; 8°47'-8°49'S, 1 19°37'-1 19°40'E; reported pres- ent 1969-1973 by W. Auffenberg (1981, p. 40). FAS;C:LS-13. Koh Alang Yai. See Rang Yai, Ko. Koh Chang Island. See Chang, Ko. Kohhang. See Pran Buri, Mae Nam, mouth. Koh Kram Island. See Khram Yai, Ko. Koh Kut Island. See Kut, Ko. Koh Lak. See Prachuap Khiri Khan. Koh Lang. See Rang Yai, Ko. Koh Naka Yai. See Naka Yai, Ko. Koh Nam Kam. See Nang Kham, Ko. Koh Pennan. See Phangan, Ko. Koh Piam. See Phayam, Ko. Koh Pipidon. See Phi Phi Don, Ko. Koh Samui. See Samui, Ko. Koh Si Chang. See Si Chang, Ko. Ko Khanghkao. See Khangkhao, Ko. Ko Khangkao. See Khangkhao, Ko. Ko Klet Kaeo. See Klet Kaeo, Ko. Komodo, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°26'-8°46'S, 119°22'-119°34'E; mon- keys reported absent Jul. 1969-Jun. 1970 by W. Auffenberg (1981, p. 242). C:e. Kopenheat; Great Nicobar Island, INDIA; 6°58'N, 93°43'E; collected 23 Mar. 1901 by W. L. Ab- 146 FIELDIANA: ZOOLOGY bott (field catalog; cf. Miller, 1902b, p. 751; Kloss, 1 903a, map facing p. 8); usnm, 1 UMB; A:N-3. Korinchi. See Sandaran Agong. Ko Sichang. See Si Chang, Ko. Kosumpee. See Kosumphi Forest Park. Kosumphi Forest Park; THAILAND; 16°15'N, 103°05'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Tak- enaka, and P. Varavudhi (1989, p. 95). Ob- served 21 Jul. 1989 and 12 Jan. 1991 by N. Aggimarangsee (1992, pp. 109, 111, 119;pers. comm., Oct. 1993). Subspecies uncertain; A:T- 12. Kosumphisai. See Kosumphi Forest Park. Kotabumi, Propinsi Lampung, 23 m; Sumatra, INDONESIA; 4°50'S, 104°54'E; collected 6 May 1940 by Vogelpol; zrc, 1. FAS; B:S-81. Kota Kinabalu. See Papar; Talibang; Tuaran. Kotapinang. See Telukpanji, Kotapinang region. Kotawaringin. See Riam, Kotawaringin district. Kowloon Reservoir Area; UNITED KINGDOM: HONG KONG; ca. 22°21'N, 114°09'E; intro- duced population, reported present ca. 1967 by P. Marshall (1967, p. 44). Subspecies uncertain; not mapped. Krakatau, Pulau. See Rakata, Pulau. Krakatau Ketjil, Pulau. See Rakata-kecil, Pulau. Krau Game Reserve. See Kuala Lompat. Krau River. See Benom, Gunong. Kretam Besar, Sungai, Kinabatangan district, 70 m; Borneo, MALAYSIA: SABAH; 5°32'N, 1 1 8°32'E; collected 25 May 1 950 by D. D. Davis ( 1 962, p. 1 1 ; field catalog); fmnh, 1 . FAS; C:Sab- 20. Kretam Kechil, Sungai, Kinabatangan district, 70 m; Borneo, MALAYSIA: SABAH; 5°31'N, 1 1 8°33'E; collected 1 3 May 1 950 by D. D. Davis ( 1 962, p. 1 1 ; field catalog); fmnh, 1 . FAS; C:Sab- 20. Kroh Reserve. See Keroh Forest Reserve. Kuala Belalong Field Studies Centre; Borneo; BRUNEI; ca. 4°33'N, 115°08'E; reported pres- ent 1991-1992 by Earl of Cranbrook (1993, p. 274). FAS; not mapped (record discovered after Fig. 2C was prepared). Kuala Binjai. See Binjai, Sungai. Kuala Indau. See Endau, Sungai, vicinity. Kuala Lompat; MALAYSIA: WEST MALAY- SIA; ca. 3°42'N, 102°17'E; observed 1969-1970 by D. Chivers (1971, p. 80). Observed 1969- 1970 by Lord Medway and D. R. Wells (1971, p. 246). FAS;B:WM-15. Kuala, Lompat, 50 m; MALAYSIA: WEST MA- LAYSIA; 3°43'N, 102°17'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). Observed Mar. 1984-Oct. 1986 by F. Lambert (1990, p. 455). FAS; B:WM-15. Kuala Lompat Post, 0-1 km NW; MALAYSIA: WEST MALAYSIA; ca. 3°43°N, 102°17'E; ob- served Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 19). Observed Jul. 1974-Jan. 1976 by F. P. G. Aldrich-Blake (1980, p. 147). FAS; B:WM-15. Kuala Lompat Post, 0-2 km W; MALAYSIA: WEST MALAYSIA; ca. 3°42'N, 102°17'E; ob- served Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 19). FAS; B:WM-15. Kuala Lumpur. See Nanas, Bukit, Kuala Lumpur. Kuala Lumpur vicinity; MALAYSIA: WEST MA- LAYSIA; ca. 3°10'N, 101°42'E; observed 1901- 1 902 by N. Annandale and H. C. Robinson (see Bonhote, 1903, p. 4). Observed Aug.-Dec. 1970 by C. H. Southwick and F. C. Cadigan, Jr. ( 1 972, p. 13). Observed Jun. 1976-Jul. 1977 by Y. L. Mah and F. P. G. Aldrich-Blake (1980, p. 354). Laboratory animals obtained 1 979-1 980 by un- known collectors (Burke et al., 1981, p. 928). FAS; B:WM-19. Kuala Pilah vicinity; MALAYSIA: WEST MA- LAYSIA; ca. 2°44'N, 102°15'E; examined for malaria before 1994 by A. N. Rain, J. W. Mak, and R. Zamri (1993, p. 386). FAS; B:WM-25. Kuala Rompin, 5 m; MALAYSIA: WEST MA- LAYSIA; 2°50'N, 103°26'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-22. Kuala Selangor, 0-40 m; MALAYSIA: WEST MALAYSIA; 3°21'N, 101°15'E; observed Feb. 1965-Jun. 1966 by I. S. Bernstein (1968b, p. 8). Observed Aug.-Dec. 1970 by C. H. Southwick and F. C. Cadigan, Jr. (1972, p. 10). FAS; B:WM- 18. Kuala Selangor, sea level; MALAYSIA: WEST MALAYSIA; 3°20'N, 101°17'E; reported pres- ent before 1973 by B. E. Weber (1972, p. 468). Observed Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM- 18. Kuala Selangor estuary, south bank; MALAYSIA: WEST MALAYSIA; ca. 3°21'N, 101°15'E; ob- served Mar. -Apr. 1977 by Lim Boon Hock and A. Sasekumar (1979, p. 106). FAS; B.WM-18. Kuantan; MALAYSIA: WEST MALAYSIA; 3°48'N, 103°20'E; captive obtained ca. 1963 by unspecified collectors (Garnham, 1963, p. 156). FAS; B:WM-13. Kuatnana, 300 m; Pulau Timor, Lesser Sunda Is- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 147 lands, INDONESIA; ca. 9°50'S, 124°10°E; col- lected 20 Jul. 1911 by C. B. Haniel (see Hell- mayr, 1914, p. 5); zsbs, 1. FAS; QLS-26. Kuching; Borneo, MALAYSIA: SARAWAK; 1°33'N, 1 10°20'E; collected 23 Sep. 1893 by un- known collector; smk, 1. Collected Mar. 1898 by unknown collector; smk, 1. Collected 25 Jul. 1 898 by C. Ulok; smk, 1 (skin only, skull inside). Collected date unknown by unknown collector; smk, 2 (skulls only). FAS; C:Sar-4. Kuching, 10th mile; Borneo, MALAYSIA: SA- RAWAK; ca. 1°33'N, 1 10°20'E; collected 16 Jan. 1895 by unknown collector; smk, 1 (skin only, skull inside). FAS; C:Sar-4. [Kuching, probably]; Borneo, MALAYSIA: SA- RAWAK; 1°33'N, 110°20'E; collected 1865- 1866 by G. Doria (see Beccari, 1904, p. 30); zmb, 1. FAS; C:Sar-4. Kuda. See Poelau (?= Kuda), Sungai Sibau. Kukuh; Pulau Bali, Lesser Sunda Islands, IN- DONESIA; ca. 8°31'S, 1 15°12'E; blood samples taken in 1980 by Y. Kawamoto, K. Nozawa, andTb. M. Ischak(1981,p. 16). Observed Jul. - Sep. 1993 by M. F. Small (1994, p. 10). FAS; QLS-3. Kundur, Pulau, Kepulauan Riau, INDONESIA; 0°38'-0°53'N, 103°21'-103°30'E; reported pres- ent Jun.-Aug. 1 903 by W. L. Abbott (see Miller, 1906c, p. 279). Collected 19 Aug. 1908 by E. Seimund; zrc, 2 (skulls only). FAS; B:SCS-17. Kuprakona. See Wat Koo Pra Kona. Ku Prakonna. See Wat Koo Pra Kona. Kut, Ko, THAILAND; 1 1°33'-1 1°46'N, 102°32'- 102°37'E; collected 23-29 Dec. 1914 by C. B. Kloss (1916b, p. 28); bm(nh), 3; bm(nh) (skull)/ zrc (skin), 1; usnm, 2; zrc, 1. Collected 1 Jun. 1917 by H. M. Smith (see Riley, 1938, p. 2); usnm, 1. FAS; A:T-36. Kuta; Pulau Lombok, Lesser Sunda Islands, IN- DONESIA; 8°55'S, 116°17'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Kawamoto (1982, p. 66). FAS; QLS-6. Kutai Nature Reserve, northeast corner; Borneo: Kalimantan, INDONESIA; ca. 0°30'N, 117°30'E; observed 1 May 1970-31 Jul. 1971 by P. Rodman (1973, p. 655). Observed Oct. 1974-Jun. 1976 by B. P. Wheatley (1978, p. 347). FAS; CK-46. Kute. See Kuta. Kya-eng. See Ataran River. Labuan Badjan Bay. See Labuhanbajau. Labuhanbajau; Pulau Simeulue, INDONESIA; 2°24'N, 96°28'E; collected 1 Jan. 1902 by W. L. Abbott (see Miller, 1903a, p. 437); usnm, 1. FUS; B:IO-3. Labuhandeli. See Deli, Sungai, between Belawan and Labuhandeli. Labuhandeli vicinity; Sumatra, INDONESIA; 3°45'N, 98°4 1 'E; collected 1881-1883 by B. Ha- gen (1890, p. 82); museum unknown, 1 (not seen). FAS; B:S-21. Labuk Road, Sepilok Forest Reserve; Borneo, MALAYSIA: SABAH; ca. 5°53'N, 118°00'E; observed 1968-1969 by M. Kawabe and T. Mano (1972, p. 216). FAS; C:Sab-17. Lac Giao, 400 m; VIETNAM; 12°40'N, 108°03'E; collected 18 Mar. 1937 by W. H. Osgood (1941, p. 1; field catalog); fmnh, 1. AUR/FAS/MUL; A:V-2. Lacon. See Nakhon Si Thammarat. Lacon Stritamarat. See Nakhon Si Thammarat. La Datu. See Lahad Datu. Laem Ngop-Phumi Cham Yeam, river between; CAMBODIA or THAILAND; ca. 12°00'N, 102°47'E; observed in 1914 by C. B. Kloss (1916a, p. 32). FAS; A:C-3. Laem Sing mountains; THAILAND; 12°29'N, 102°04'E; collected 7 Jun. 1926 by H. M. Smith (see Riley, 1938, p. 9); usnm, 1. FAS; A:T-33. Laem Son National Park; THAILAND; ca. 9°17'N, 98°31'E; observed 25-28 Nov. 1987 by R. Boonratana (1988, p. 76). AUR/FAS/MUL; A:T-46. Lafau; Pulau Nias, INDONESIA; 1°23'N, 97°13'E; collected 24 Mar. 1905 by W. L. Abbott (cf. Lyon, 1916, p. 458); usnm, 1. FAS; B:IO-6. Lagong, Pulau, INDONESIA; 3°34'-3°38'N, 108°04'-108°08'E; collected 19 Jun. 1900 by W. L. Abbott (see Miller, 1901, p. Ill); usnm, 1. FAS; B.SCS-34. Lahad Datu; Borneo, MALAYSIA: SABAH; 5°02'N, 118°19'E; collected 10 Nov. 1903 by Dr. Pagel; zmb, 1. FAS; C:Sab-21. Lahat; Sumatra, INDONESIA; 3°48'S, 103°32'E; collected in 1875 by unknown collector; NHMBa, 2 (skulls only). FAS; B:S-73. Lai char; BURMA; ca. 21°05'N, 92°12'E; ob- served Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1985, p. 30). AUR; A:Bu-l. Lampung. See Wonosobo, Propinsi Lampung. Lampung, Propinsi (province); Sumatra, INDO- NESIA; ca. 5°10'S, 104°45'E; collected in 1908 by J. Elbert; zsbs, 1 (skull only). Blood samples taken Jan.-Nov. 1 979 by Y. Kawamoto and Tb. M. Ischak (1981, p. 238). FAS; B:S-80. 148 FIELDIANA: ZOOLOGY Lampungs. See Lampung, Propinsi (province). Lamukotan, Pulau. See Lemukutan, Pulau. Lanbi Kyun, BURMA; 10°42'-10°59'N, 98°02'- 98°18'E; collected 30 Jan. 1900 and 6 Jan. 1904 by W. L. Abbott; usnm, 2 (including 1 skull only). AUR; A:Bu-21. Lancang Kuning; Pulau Bintan, Kepulauan Riau, INDONESIA; not precisely located, 0°48'- 1°13'N, 104°13'-104°34'E; observed 18 Jan.-2 Feb. 1992 by A. Yanuar (1994, p. 2). FAS; B:SCS-8. Landai vicinity; Sumatra, INDONESIA; ca. 0°01'S, 100°37'E; reported Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-54. Landak. See Perbuah, Sungai Landak. Langgaliroe; Pulau Sumba, Lesser Sunda Islands, INDONESIA; not precisely located, 9° 17'- 10°19'S, 1 18°57'-120°50'E; collected 24-29 May 1932 by G. Stein; mzb, 2; zmb, 1. FAS; C:LS- 22, LS-23. Lang Island. See Rakata-kecil, Pulau. Langkawi, Pulau, MALAYSIA: WEST MALAY- SIA; 6°15'-6°28'N, 99°38'-99°55'E; collected 10 Feb. 1909 by museum collector (see H. C. Rob- inson & Kloss, 1910, p. 664; H. C. Robinson, 1917, p. 130); bm(nh), 1. FAS; B:SM-1. Langkawi, Pulau, 50 m, MALAYSIA: WEST MA- LAYSIA; 6°15'-6°28'N, 99°38'-99°55'E; ob- served Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:SM-1. Langkoe. See Langkoi, Pulau. Langkoi, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°44'S, 1 1 9°22'E; monkeys reported ab- sent 1969-1973 by W. Auffenberg(1981, p. 40). C:e. Lanjak-Entimau Orang-Utan Sanctuary (pro- posed); Borneo, MALAYSIA: SARAWAK; ca. 1°30'N, 11 2°05'E; observed Aug.-Nov. 1981 by Sarawak Forest Department— World Wildlife Fund team (Kavanagh, 1982, p. 320). FAS; C:Sar-ll. Lankawi, P. See Langkawi, Pulau. Lanta Yai, Ko, THAILAND; 7°28'-7°4rN, 99°02'-99°08'E; collected 9-12 Jan. 1917 by H. C. Robinson and E. Seimund (H. C. Robinson, 1917, p. 135); museum unknown, 1 (not seen). FAS; A:T-55. Larut, Daerah (district); MALAYSIA: WEST MALAYSIA; ca. 4°55'N, 100°47'E; collected before 1901 by unknown collector (Flower, 1900, p. 316); specimens reportedly in Taiping Mu- seum (not seen). FAS; B:WM-5. Lasia, Pulau, INDONESIA; 2°08'-2°12'N, 96°37'- 96°40'E; collected 4-5 Jan. 1902 by W. L. Ab- bott (see Miller, 1903a, p. 438); usnm, 2. LAS; B:IO-4. Lat Bua Kao. See Lat Bua Khao, Sathani. Lat Bua Khao, Sathani; THAILAND; 14°52'N, 101°36'E; collected 10 Oct 1916 by C. B. Kloss (1919c, p. 335); usnm, 1; zrc, 1. FAS; A:T-9. Laut, Pulau, INDONESIA; 4°40'-4°46'N, 107°56'- 108°02'E; collected Jul. 1 894 by C. Hose; bm(nh), 1 (skin only). Collected 9 Aug. 1900 by W. L. Abbott (see Miller, 1901, p. 11 1); usnm, 1 . FAS; B:SCS-31. Lauttador; Sumatra, INDONESIA; 3°18'N, 99°15'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of speci- mens unknown (not seen). FAS; B:S-23. Laut Tawar, Danau. See Bur ni Bebuli. Lawadarat, Gili, Lesser Sunda Islands, INDO- NESIA; 8°28'-8°29'S, 119°33'-119°34'E; mon- keys reported absent 1969-1973 by W. Auffen- berg(1981,p. 40). C:d. Lawalaut, Gili, Lesser Sunda Islands, INDONE- SIA; 8°27'-8°28'S, 119°33'-119035'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Lelogama, 845 m; Pulau Timor, Lesser Sunda Is- lands, INDONESIA; 9°44'S, 123°57'E; collected 22 May-3 Jun. 191 1 by C. B. Haniel (see Hell- mayr, 1 9 1 4, p. 5); zsbs, 6 (includes 3 skins only); FAS; CLS-26. Lem Sing Mts. See Laem Sing mountains. Lemukutan, Pulau, INDONESIA; 0°43'-0°47'N, 108°42'-108°44'E; reported present 7-10 May 1907 by W. L. Abbott (in Lyon, 191 1, p. 59). FAS; B:SCS-24. Lengah, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°4 l'S, 1 1 9°28'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:e. Lesong, 100 m; MALAYSIA: WEST MALAY- SIA; 2°44'N, 103°09'E; reported as probably present Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM- 21. Lesten, Daerah Istimewa Aceh, 700 m; Sumatra, INDONESIA; 4°10'N, 97°40'E; collected 19 Mar. 1937 by A. Hoogerwerf (1941, p. 7; cf. Chasen, 1940b, p. 485); mzb, 1. FAS; B:S-11. Letsok-aw Kyun, BURMA; 1 1°27'-1 1°48'N, 98°10'-98°20'E; collected 26 Jan. 1904 by W. L. Abbott; usnm, 1 (skin and mandible only). AUR; A:Bu-20. Leuser, Gunung. See Gunung Leuser Reserve. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 149 Lhokseumawe, ca. 80 km ESE; Sumatra, INDO- NESIA; ca. 4°55'N, 97°48'E; reported present Nov. 1971-Jan. 1973 by local residents (Crock- ett & Wilson, 1980, p. 156). FAS; B:S-10. Liang Koeboeng; Borneo: Kalimantan, INDO- NESIA; 0°37'N, 1 13°08'E; collected 10-18 Apr. 1894 by J. Buttikofer (1897, p. 16); rmnh, 2 (see Jentink, 1897, p. 39, who also lists a third spec- imen). FAS; C:K-16. Lima Belas Estate, ca. 50-120 m; MALAYSIA: WEST MALAYSIA; ca. 3°46'N, 101°21'E; ob- served Jul. 1965-Jun. 1966 by I. S. Bernstein (1967, p. 199). Observed Jan. 1980-May 1981 by J. O. Caldecott (1986b, p. 21). FAS; B:WM- 18. Lima Bias Estate. See Lima Belas Estate. Linga Island. See Lingga, Pulau. Lingartjati. See Linggajati. Lingga; Borneo, MALAYSIA: SARAWAK; 1°21'N, 111°10'E; collected Nov. 1899 by mu- seum collectors; SMK, 2 (skins only, skulls in- side). FAS; C:Sar-8, Lingga, Pulau, INDONESIA; 0°02'N-0°22'S, 104°26'-105°00'E; collected 23 Jul. 1899 by W. L. Abbott (see Miller, 1900, p. 242; 1906c, p. 284; Kloss, 1903b, p. 54); usnm, 2. FAS; B:SCS- 13. Linggajati, Cirebon district, 600 m; Java, IN- DONESIA; 6°52'S, 108°28'E; collected 3 Jan. 1933 by J. J. Menden; amnh, 1. FAS; B:J-23. Linglung, Pulo. See Lagong, Pulau. Lingung, Pulo. See Lagong, Pulau. Little Andaman Island, INDIA; 10°31'-10°54'N, 92°22'-92°36'E; primates reported absent be- fore 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, [ 1 928], p. 802; Chaturvedi, 1 980, p. 1 34). A:h. Little Condor Island. See Ben Dam. Little Durian. See Durian-kecil, Pulau. Little Jolly Boy Island, Andaman Islands, INDIA; 11°31'-11032'N, 92°36'-92°37'E; primates re- ported absent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, [1928], p. 802; Cha- turvedi, 1980, p. 134). A:f. Little Karimon. See Karimun-kecil, Pulau. Little Kretam River. See Kretam Kechil, Sungai. Little Nicobar Island, INDIA; 7°13'-7°25'N, 93°37'-93°45'E; observed before 1847 by P. Barbe (1 846, p. 365). Collected 25-27 Feb. 1 90 1 by W. L. Abbott (see Miller, 1902b, p. 751; Kloss, 1903a, pp. 122, 128); usnm, 3. UMB; A:N-2. Little Tenasserim River. See Thagyet, Little Ten- asserim River. Loa Bambam; Borneo: Kalimantan, INDONE- SIA; 0°29'S, 1 17°02'E; collected 1-15 Jun. 1912 by H. C. Raven (see Deignan, [1960], p. 267); usnm, 2 (including 1 skull only; third specimen listed in field catalog). FAS; CK-49. Lo Bon Bon. See Loa Bambam. Loeboe; Sumatra, central, INDONESIA; not lo- cated, 5°38'N-5°57'S, 95°12'-106°05'E; ob- served 1877-1879 by J. F. Snelleman (1887, p. 10). FAS; not mapped. Loeboek Linggan. See Lubuklinggau. Loeboek Sikaping. See Lubuksikaping. Logo. See Lengah, Pulau. Loho Gringgo. See Ginggo, Teluk. Loka; Pulau Sulawesi, INDONESIA; ca. 5°26'S, 1 19°54'E; introduced, reported present 21 Oct.- 15 Nov. 1888 by Malay hunters employed by M. Weber (1890a, p. vii; 1890b, p. 102). Sub- species uncertain; not mapped. Lokan, Sungai; Borneo, MALAYSIA: SABAH; ca. 5°32'N, 1 17°33'E; observed 1969-1970 by P. S. Rodman (1991, p. 359). FAS; C:Sab-15. Lomblen, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°H'-8°35'S, 123°13'-123°55'E; M. fascicularis reported absent before 1937 by J. J. M. F. Symons (Mertens, 1936, p. 319). C:f. Lombok, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°13'-8°58'S, 115°50'-116°43'E; re- ported present before 1 848 by H. Zollinger (1847, p. 203). Reported present May-Jul. 1896 by A. Everett (in Hartert, 1896, p. 593). Purchased ca. 1920 by W. G. Wallace; bm(nh), 1 (skull only). FAS; CLS-5 through LS-7. Lompat, Sungai, ca. 3 km ENE of Kuala Serloh; MALAYSIA: WEST MALAYSIA; ca. 3°41'N, 102°1 l'E; observed Jul.-Dec. 1977 by D. Oliv- ers and G. Davies (1979, p. 20). FAS; B:WM- 15. Lompat, Sungai, ca. 3 km W of Kuala Lompat Post; MALAYSIA: WEST MALAYSIA; ca. 3°42'N, 102°16'E; observed Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 20). FAS; B:WM-15. Lompat, Sungai, ca. 4 km ENE of Kuala Serloh; MALAYSIA: WEST MALAYSIA; ca. 3°41'N, 102°12'E; observed Jul.-Dec. 1977 by D. Chiv- ers and G. Davies (1979, p. 20). FAS; B:WM- 15. Lompat, Sungai, ca. 4 km W of Kuala Lompat Post; MALAYSIA: WEST MALAYSIA; ca. 3°42'N, 102°15'E; observed Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 20). FAS; B:WM-15. Lompat, Sungai, ca. 6 km W of Kuala Lompat 150 FIELDIANA: ZOOLOGY Post; MALAYSIA: WEST MALAYSIA; ca. 3°42'N, 102°14'E; observed Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 20). FAS; B:WM-15. Long Ekang; Borneo, MALAYSIA: SARAWAK; 3°55'N, 1 14°27'E; collected 26 Feb. 1956 by T. A. Chavasse (see Arnold, 1959, p. 204); fmnh, l.FAS;C:Sar-19. Long Ikang. See Long Ekang. Longo, Pulau. See Longos, Nusa. Longos, Nusa, Lesser Sunda Islands, INDONE- SIA; 8°20'-8°21'S, 120°07'-120°09'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Long Pangian (?= Long Pangean); Borneo: Kali- mantan, INDONESIA; 2°42'N, 116°42'E; col- lected 24 Apr. 1914 by C. Lumholtz (see Gyld- enstolpe, 1920, pp. 3, 14-15); Zoological Mu- seum, Christiana, 1 (not examined). FAS; C:K- 38. Long Pelban. See Long Peleben. Long Peleben; Borneo: Kalimantan, INDONE- SIA; 2°47'N, 116°35'E; collected 30 Jun. 1935 by V. von Plessen (1936, p. 100; cf. Stresemann, 1938, p. 1 10); amnh, 1; mzb, 1. Collected Aug. 1956 by P. Pfeffer; mnhn, 1 (skull only). FAS; CK-38. Lontar, Pulau. See Lanta Yai, Ko. Lopburi. See Sarn Pra Kara, Lopburi. Lower Taipo Road, Kowloon Reservoir Area; UNITED KINGDOM: HONG KONG; ca. 22°21'N, 114°09'E; introduced population, ob- served Jul.-Aug 1980 and Jul.-Aug. 198 1 by C. H. Southwick and K. L. Southwick (1983, p. 19). Subspecies uncertain; not mapped. Luang Ko. See Rang Yai, Ko. Lubuklinggau, ca. 15 km N; Sumatra, INDO- NESIA; ca. 3°10'S, 102°50'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-64. Lubuklinggau, ca. 40 km NNW; Sumatra, IN- DONESIA; ca. 2°58'S, 102°43'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-65. Lubuklinggau, ca. 60 km SE; Sumatra, INDO- NESIA; ca. 3°40'S, 103°15'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-72. Lubuklinggau, sea level; Sumatra, INDONESIA; 3°18'S, 102°52'E; collected 21 Nov. 1933 by J. J. Menden; amnh, 1. FAS; B:S-64. Lubuklinggau vicinity; Sumatra, INDONESIA; ca. 3°18'S, 102°52'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-64. Lubukminturun area; Sumatra, INDONESIA; ca. 0°52'S, 100°23'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Kawamoto (1982, p. 67). FAS; B:S-47. Lubuksikaping; Sumatra, INDONESIA; OWN, 100°10'E; collected 1 Aug. 1930 by E. Jacobson; rmnh, 1 (skull only). FAS; B:S-43. Lumu Lumu, 6000 ft (= 1830 m); Borneo, MA- LAYSIA: SABAH; ca. 6°02'N, 116°35'E; cap- tives acquired Jun. 1932 by J. A. Griswold, Jr. (1939b, p. 506). FAS; C:Sab-7. Lungmanis Station, Sungai Segaliud; Borneo, MA- LAYSIA: SABAH; ca. 5°40'N, 117°45'E; ob- served in 1958 by K. Stott, Jr. (1964, p. 13). FAS; C:Sab-16. Macarah [?= Moearah] Doewa. See Muaradua. MacRitchie Reservior Nature Reserve; Singapore Island; SINGAPORE; ca. 1°21'N, 103°50'E; re- ported present before 1974 by S. H. Chuang (1973, p. 3). FAS; B:SCS-7. Madgalengka. See Majalengka. Madura, Pulau, INDONESIA; 6°52'-7°15'S, 1 1 2°4 1 '-1 1 4°07'E; reported present before 1 90 1 by A. G. Vorderman (1900, p. 143). FAS; C:J- 39. Maenam Chao Phaya. See Chao Phraya, Mae Nam, below Bangkok. Mae Nam Khwae Noi. See Khwae Noi, Mae Nam. Mahakam, Sungai, left bank, 1 km above Sebulu; Borneo: Kalimantan, INDONESIA; ca. 0°16'S, 1 1 7°00'E; observed 29-30 Aug. 1983 by S. Azu- ma, A. Suzuki, and Y. Ruhiyat (1984, p. 48). FAS; C:K-49. Mahakam, Sungai, left bank, 4 km above Sebulu; Borneo: Kalimantan, INDONESIA; ca. 0°17'S, 1 16°58'E; observed June 1972 by C. L. Darsono (see Wilson & Wilson, 1975, p. 257). Observed 29-30 Aug. 1983 by S. Azuma, A. Suzuki, and Y. Ruhiyat (1984, p. 48). FAS; QK-49. Mahakam, Sungai, north bank, above Samarinda; Borneo: Kalimantan, INDONESIA; ca. 0°30'S, 1 17°05'E; collected 31 May 1912 by H. C. Ra- ven (field catalog); usnm, 1. FAS; QK-49. Mahakam, Sungai, right bank, 2 km below Sebulu; Borneo: Kalimantan, INDONESIA; ca. 0°17'S, 1 1 7°00'E; observed 29-30 Aug. 1 983 by S. Azu- ma, A. Suzuki, and Y. Ruhiyat (1984, p. 48). FAS; QK-49. Mahakam, Sungai, right bank, 8 km above Sebulu; Borneo: Kalimantan, INDONESIA; ca. 0°18'S, 1 16°57'E; observed 29-30 Aug. 1983 by S. Azu- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 151 ma, A. Suzuki, and Y. Ruhiyat (1984, p. 48). FAS; CK-49. Mahasarakham. See Kosumphi Forest Park. Majalengka, Cirebon district, 600 m; Java, IN- DONESIA; 6°50'S, 108°13'E; collected 22 Dec. 1932 by J. J. Menden; amnh, 3. FAS; B:J-22. Malabar, Pengalengan district, 6000 ft (= 1 800 m); Java, INDONESIA; 7°07'S, 107°36'E; collected in 1937 by D. P. Bosscha Erdbrink; zluu, 1 (skull only). FAS; B:J-6. Malacca. See Melaka. Malakka. See Melaka. Malang. See Wonokojo (?= Wonokerto), Dampit district, southern Malang region. Malawali, Pulau, MALAYSIA: SABAH; 7°01'- 7°05'N, 117°16'-117°22'E; observed Feb.-Jun. 1991 by Shukor Md. Nor (pers. comm., 20 Jul. 1991). FAS; C:Sab-ll. Malaya. See MALAYSIA: WEST MALAYSIA. MALAYSIA: WEST MALAYSIA; 1°16'-6°43'N, 100°07'-104°18'E; collected date unknown by unknown collector; usnm, 1 . FAS; not mapped. Maliangin Besar, Pulau, MALAYSIA: SABAH; 7°04'-7°05'N, 1 17°02'-1 17°03'E; observed Feb.- Jun. 1991 by Shukor Md. Nor (pers. comm., 20 Jul. 1991). FAS; C:Sab-9. Maliwun. See Pakchan River, near Maliwun. Maman; Pulau Sumbawa, Lesser Sunda Islands, INDONESIA; ca. 8°37'S, 1 17°29'E; blood sam- ples taken Jul. 1981-Jan. 1982 by Y. Kawa- moto, Tb. M. Ischak, and J. Supriatna (1982a, p. 58). FAS; C:LS-8. Mandalay. See Irrawaddy River, right bank, northwest of Mandalay. Mandau, Sungai; Sumatra, INDONESIA; ca. 0°48'N, 101°47'E; observed ca. 1907 by M. Moszkowski (1909, p. 143, map following p. 192). FAS;B:S-42. Mandiri River. See Bantargebang. Manggis, Gunung Kelud, Kediri district; Java, IN- DONESIA; 7°49'S, 1 12°14'E; collected in 1909 by V. Arnim; zsbs, 9 (skulls only). FAS; C:J-33. Mangiatan, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°33'S, 1 1 9°4 1 'E; reported present 1 969- 1 973 by W. Auffenberg (1 98 1 , p. 40). FAS; C:LS- 13. Maninjau; Sumatra, INDONESIA; 0°18'S, 100°14'E; collected Mar.-Sep. 1888 by M. We- ber (1890a, p. iii; 1890b, p. 102); museum un- known, 1 (skeleton only; not seen). FAS; B:S- 45. Mansalar, Pulo. See Mursala, Pulau. Mao Marroe, 450 m; Pulau Sumba, Lesser Sunda Islands, INDONESIA; 9°58'S, 120°30'E; col- lected 4-10 May 1925 by P. F. Franck and Den- in (see Dammerman, 1926a, p. 91); mzb, 3. FAS; GLS-23. Mapor, Pulau. See Mentigi. Marah. See Merah. Maratua, Pulau, INDONESIA; 2°10'-2°19'N, 1 1 8°33'-l 1 8°39'E; collected 2 1 May 1 9 1 3 by H. C. Raven (in Riley, 1930, p. 2); usnm, 4. TUA; CK-41. Margo-Molio, Gunung. See Margomulio, Gun- ung. Margomulio, Gunung, Gunung Kelud group, Ke- diri district; Java, INDONESIA; 7°55'S, 1 1 2°1 5'E; collected Aug. 1 909 by V. Arnim; zsbs, 1 (skull only). FAS; C:J-33. Masalembu Besar, Pulau, INDONESIA; 5°32'- 5°36'S, 114°24'-114°27'E; monkeys reported absent 4-6 Dec. 1 907 by W. L. Abbott (in Lyon, 1911, p. 61). C:b. Mata, Pulo. See Matak, Pulau. Matak, Pulau, INDONESIA; 3°13'-3°23'N, 106°14'-106°1 8'E; observed 24 Aug.-5 Sep. 1 899 by W. L. Abbott (in Miller, 1900, p. 245; cf. Kloss, 1903b, p. 72). FAS; B:SCS-30. Matasiri, Pulau, east coast; INDONESIA; ca. 4°47'S, 115°50'E; collected 8 Dec. 1907 and 1907-1908 by W. L. Abbott (field catalog; in Lyon, 1911, p. 61); usnm, 2 (including 1 skull only). FAS; C:K-53. Maung district. See Phu Phan, Maung district. Mauritius Island, MAURITIUS; 19o59'-20o31'S, 57°18'-57°48'E; introduced population, ob- served 19 Apr. 1753 by M. l'Abbe de la Caille (1763, p. 230). Collected before 1 862 by E. New- ton; bm(nh), 1. Collected 20 Jul. 1875 by J. Au- debert; rmnh, 1 (skin only, skull inside). Col- lected before 1 9 1 0 by Franco-British Commis- sion; bm(nh), 1. Collected before 1956 by un- known collector; bm(nh), 1 (skull only). Observed 1977-1984 by R. W. Sussman and I. Tattersall (1986, p. 34). Blood samples taken Oct. 1989 by M. Kondo, Y. Kawamoto, K. Nozawa, K. Matsubayashi, T. Watanabe, O. Griffiths, and M.-A. Stanley (1993, p. 168). Subspecies un- certain; not mapped. Mbarapu, Nusa, Komodo area, Lesser Sunda Is- ands, INDONESIA; 8°37'S, 1 19°32'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:e. Mbura; Pulau Flores, Lesser Sunda Islands, IN- DONESIA; 8°34'S, 1 19°52'E; collected 17 Oct. 1929 by J. K. de Jong; mzb, 4. FAS; GLS-13. Medan, ca. 20 km N; Sumatra, INDONESIA; ca. 3°47'N, 98°42'E; observed Nov. 1971-Jan. 1973 152 FIELDIANA: ZOOLOGY by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-21. Medan, ca. 75 km NW; Sumatra, INDONESIA; ca. 4°04'N, 98°09'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-19. Medan, ca. 100 km SSW; Sumatra, INDONESIA; ca. 2°45'N, 98°1 8'E; reported present before 1 98 1 by H. D. Rijksen (Crockett & Wilson, 1980, p. 156). FAS;B:S-26. Medan, ca. 105 km SSE; Sumatra, INDONESIA; ca. 2°40'N, 98°42'E; reported present Nov. 197 1- Jan. 1 973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:S-28. Medan, forest near; Sumatra, INDONESIA; ca. 3°35'N, 98°40'E; collected in 1906 by C. Brue- gekzsBS, 1. FAS; B:S-21. Medan, near sea level; Sumatra, INDONESIA; 3°35'N, 98°40'E; collected in 1905 by C. Brue- gel, zsbs, 1 (skin only). Collected 1906-1910 by C. Widnmann; zsbs, 78 (including 41 skins only, 3 skulls only). Collected 21 May 1939 by F. Ulmer (in Miller, 1942, p. 127; cf. Schauensee & Ripley, 1 940a, p. 3 1 6); ansp, 1 . Collected date unknown by Mrs. P. W. Cremer-Jansen; mcz, 1 (skull only). FAS; B:S-21. Medan-Siantar; Sumatra, INDONESIA; ca. 3°35'N, 98°40'E; observed Mar.-May 1939 by F. A. Ulmer, Jr. (in Miller, 1942, p. 127). FAS; B:S-21. Medan vicinity; Sumatra, INDONESIA; ca. 3°35'N, 98°40'E; reported present Nov. 1971- Jan. 1 973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:S-21. Melaka; MALAYSIA: WEST MALAYSIA; 2°12'N, 102°15'E; collected 1818-1863 by P. Diard; rmnh, 1 (skin only). Collected before 1 849 by M. de Montigny; mnhm, 2 (skins only, skulls inside). Reported present before 1852 (Anony- mous, 1851, p. 444). Collected before 1876 by Gerrard; bm(nh), 1. Collected in 1875 by Schneider; rmnh, 2 (including 1 skin only, skull inside). FAS; B:WM-26. Melinau Gorge, rock shelter near RGS Camp 5, Mulu National Park, ca. 200 m; Borneo, MA- LAYSIA: SARAWAK; 4°08'N, 114°54'E; col- lected 24 Aug. 1977 by Lord Medway; bm(nh), 1 (mandibular fragment only). FAS; C:Sar-21. Meloewak. See Goenoengsetan-Meloewak. Mempura; Sumatra, INDONESIA; 0°43'N, 101°59'E; collected ca. 1907 by M. Moszkowski (1909, p. 151); zmb, 2 (skulls only). FAS; B:S- 42. Menam; Sumatra, INDONESIA; not located, 5°38'N-5°57'S, 95°12'-106°05'E; collected 25 Jul. 1928 by W. Comberg; zmb, 1 (skin only). FAS; not mapped. Mendawai, Sungai. See Katingan, Sungai. Mendit; Java, INDONESIA; ca. 8°00'S, 1 12°50'E; clinical examination in 1980 by K. Matsubay- ashi and D Sajuthi (1981, p. 48). FAS; C:J-36. Mengjatan, Pulau. See Mangiatan, Pulau. Mensuda Bay; Pulau Karimun, northeast, Kepu- lauan Riau, INDONESIA; 1°07'N, 103°23'E; collected 29 May 1903 by W. L. Abbott (field catalog; cf. Miller, 1906c, p. 277); usnm, 1. FAS; B:SCS-17. Mentarang, Sungai. See Sungai Kayan-Sungai Mentarang Nature Reserve. Mentigi; Pulau Mapur, west, Kepulauan Riau; IN- DONESIA; ca. 1°00'N, 104°48'E; collected 6 Jun. 1915 by H. C. Robinson (1916, p. 62); zrc, 1. FAS; B:SCS-9. Mentoko Research Center, Kutai National Park, ca. 50 m; Borneo: Kalimantan, INDONESIA; 0°24'N, 11 7°06'E; observed Jun. 1982-Sep. 1983 by L. Berenstain (1986, p. 258). FAS; C:K-47. Menyala, Sungai; MALAYSIA: WEST MALAY- SIA; ca. 2°29'N, 101°55'E; collected in 1972 by unknown collector; Department of Anatomy, Cambridge, 1 (in fluid; not seen, data from P. H. Napier, 1981, p. 14). FAS; B:WM-24. Me Ping River. See Ping, Mae Nam. Merah; Borneo: Kalimantan, INDONESIA; 0°50'N, 116°48'E; collected 25 Nov. 1925 by Madzoed; mzb, 1. FAS; C:K-43. Mergui; BURMA; 12°26'N, 98°36'E; collected be- fore 1857 by Prof. Oldham; bm(nh), 1 (skull only). Collected 25 Dec. 1881 by J. Anderson, zsi, 1 (skin only). Collected 1 Mar. 1914 by G. C. Shortridge (in Wroughton, 1915, p. 696); bnhs, 1. AUR; A:Bu-18. Meru, Gunung; Sumatra, INDONESIA; ca. 1°06'S, 100°27'E; 2 troops observed 21 Jun.-28 Oct. 1980 by N. Koyama, A. Asuan, and N. Natsir (1981, p. 1). Observed 27 Aug. 1984-5 Jan. 1985 by N. Koyama (1985, p. 106). FAS; B:S-47. Meru Betiri Nature Park; Java, INDONESIA; ca. 8°28'S, 1 1 3°48'E; observed 24 Aug.-1 4 Sep. 1 97 1 by A. Hoogerwerf (1974, p. 13). Observed in 1976 by J. Seidensticker (1983, p. 324). FAS; CJ-35. Meulaboh, ca. 35 km NW; Sumatra, INDONE- SIA; ca. 4°21'N, 95°54'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-2. Meulaboh, ca. 60 km NW; Sumatra, INDONE- SIA; ca. 4°30'N, 95°44'E; observed Nov. 1971- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 153 Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-3. Meulaboh vicinity; Sumatra, INDONESIA; ca. 4°09'N, 96°08'E; reported present Nov. 1971- Jan. 1 973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-1. Mewong River. See Wong, Nam Mae. Mibya Kyun, mouth of Tavoy River, BURMA; 13°36'N, 98°12'E; collected 23 Apr. 1936 by H. C. Smith; bm(nh), 2. AUR; A:Bu-15. Middle Andaman Island, INDIA; 12°05'-12°54'N, 92°42'-93°00'E; primates reported absent be- fore 1928 by C. B. Kloss ([1928], p. 802; cf. Chaturvedi, 1980, p. 134). A:b. Miri; Borneo, MALAYSIA: SARAWAK; 4°23'N, 113°59°E; collected Jun. 1890 by E. Hose and C. Hose; mcz, 1 (skin only, skull inside). Col- lected date unknown by unknown collector; Duckworth Laboratory of Physical Anthropol- ogy, Cambridge, 1 (skull and limb bones only; not seen, data from P. H. Napier, 1981, p. 14). FAS; C:Sar-18. Miri, Sungai; Borneo, MALAYSIA: SARAWAK; ca. 4°23'N, 113°58'E; collected date unknown by unknown collector; Duckworth Laboratory of Physical Anthropology, Cambridge, 1 (skel- eton only; not seen, data from P. H. Napier, 1981, p. 14). FAS; C:Sar-18. Miri district, Batang Baram; Borneo, MALAY- SIA: SARAWAK; ca. 4°23'N, 113°59'E; col- lected Jan. 1895 by C. Hose; NHMBe, 1 (skin only). FAS;C:Sar-18. Misa, Pulau, Lesser Sunda Islands, INDONESIA; 8°31'S, 119°45'E; monkeys reported absent 1969-1973 by W. Auffenberg(1981, p. 40). C:d. Mobur, Pulo. See Mubur, Pulau. Moeara Tewech. See Kampong Hadjak, Muara- tewe district. Moera Tewe. See Kampong Hadjak, Muaratewe district. Moeria, Mt. See Pangonan, Gunung Muria. Moeriah. See Pangonan, Gunung Muria. Mokut. See Kampong Mukut. Monos; Pulau Karimun, Kepulauan Riau; IN- DONESIA; 1°08'N, 103°23'E; collected 29-30 Aug. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, pp. 105, 107; cf. H. C. Rob- inson & Kloss, 1914, p. 394) and E. Seimund; bm(nh), 2. FAS; B:SCS-17. Morib; MALAYSIA: WEST MALAYSIA; ca. 3°50'N, 100°50'E; examined for malaria before 1994 by A. N. Rain, J. W. Mak, and R. Zamri (1993, p. 386). FAS; B:WM-17. Moro Batin; Sumatra, INDONESIA; 5°26'S, 105°06'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-82. Moro Batin, south-southeast; Sumatra, INDO- NESIA; ca. 5°35'S, 105°10'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-82. Moro Besar. See Durian, Pulau. Moro Kechil. See Durian-kecil, Pulau. Moscos Island Game Sanctuary; Moscos Islands, BURMA; 13°47'-14°27'N, 97°47'-97°56'E; re- ported present 1938-1939 by H. A. Maxwell (see Yin, 1954, p. 281). AUR; A:Bu-12. Mota, Gili. See Motang, Gili. Motang, Gili, Lesser Sunda Islands, INDONESIA; 8°48'-8°49'S, 119°46'-119°48'E; monkeys re- ported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:e. Moulmein. See Ye Forest, Ataran district, Moul- mein region. Moyo, Pulau, Lesser Sunda Islands, INDONESIA; 8°09'-8°23'S, 1 17°29'-1 17°42'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Kawamoto (1982, p. 67). FAS; CLS-9. Mrengoo Island. See Myengun Kyun. Mt. Setan-Meloewak. See Goenoengsetan-Mel- oewak. Mt. Sontra. See Sontra Peak. Muaradua, Palembang district, 100 m; Sumatra, INDONESIA; ca. 4°32'S, 1 04°0 5 'E; collected 10- 17 Jun. 1934 by J. J. Menden; amnh, 8. FAS; B:S-76. Muaratewe, Sungai Barito, 65 ft (= 20 m); Borneo: Kalimantan, INDONESIA; 0°57'S, 114°53'E; collected 21 Sep. 1909 by G. C. Shortridge; bm(nh), 3 (including 1 skin only). FAS; C:K-32. Muaratewe vicinity; Borneo: Kalimantan, IN- DONESIA; ca. 0°57'S, 1 14°53'E; observed 1979- 1986 by J. Marshall, D. J. Chivers, and K. M. Burton (see Chivers & Burton, [1991], p. 141). FAS; C.K-32. Muara Teweh. See Muaratewe. Muara Tua, Pulo. See Maratua, Pulau. Mubur, Pulau, INDONESIA; 3°18'-3°23'N, 106°10'-106°14'E; observed 24 Aug.-5 Sep. 1899 by W. L. Abbott (in Miller, 1900, p. 245; cf. Kloss, 1903b, p. 72). FAS; B:SCS-30. Mugitriman, Hak Pegusahaan Hutan (HPH), Jam- bi; not located; Sumatra, INDONESIA; ob- served Mar. 1989 by M. Bismark (1992, p. 11). FAS; not mapped. Muk, Ko, THAILAND; 7°22'N, 99° 1 8'E; observed 154 FIELDIANA: ZOOLOGY 4-8 Jan. 1917 by H. C. Robinson and E. Sei- mund (H. C. Robinson, 1917, p. 134). FAS; A:T-56. Mu Ko Phetra National Park; THAILAND; ca. 6°57'N, 99°42'E; observed 21-23 Dec. 1987 by R. Boonratana (1988, pp. 76, 81); also observed on unspecified adjacent islands. FAS; A:T-57. Mulu, Gunong. See Gunong Mulu National Park. Mulu, Gunung, 2000 ft (= 600 m); Borneo, MA- LAYSIA: SARAWAK; 4°03'N, 114°56'E; col- lected Oct. 1893 by C. Hose; bm(nh), 1 (skull only). FAS; C:Sar-21. Muntia, Pulau. See Muk, Ko. Muntok; Pulau Bangka, INDONESIA; 2°04'S, 105°10'E; collected in 1900 by A. A. W. Hu- brecht([1895], pp. 33, 88; cf. Kohlbrugge, 1902, p. 322; Zuckerman, [1933], p. 1062); zluu, un- specified portion of Pulau Bangka collection (see Bangka, Pulau). FAS; B:SCS-18. Mursala, Pulau, INDONESIA; 1°37'-1°42'N, 98°27'-98°36'E; collected 7-10 Mar. 1 902 by W. L. Abbott (see Miller, 1903a, p. 438); usnm, 3 (including 1 skull only). FAS; B:IO-ll. Murung, Sungai, left bank, 1 km below mouth of Sungai Danau Tolong; Borneo: Kalimantan, IN- DONESIA; ca. 0°10'S, 114°13'E; observed 4-7 Sep. 1986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, left bank, 2 km above mouth of Sungai Beriwit; Borneo: Kalimantan, INDO- NESIA; ca. 0°09'S, 1 14°17'E; observed 4-7 Sep. 1 986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, left bank, 2 km above mouth of Sungai Turusan; Borneo: Kalimantan, INDO- NESIA; ca. 0°1 l'S, 1 14°1 l'E; observed 4-8 Sep. 1986 by D.J. Chivers and KM. Burton ([1991], p. 143). FAS; QK-19. Murung, Sungai, left bank, 2 km below mouth of Sungai Turusan; Borneo: Kalimantan, INDO- NESIA; ca. 0°12'S, 1 14°08'E; observed 7-8 Sep. 1986 by D.J. Chivers and KM. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, left bank, 8 km above Muara- juloi; Borneo: Kalimantan, INDONESIA; ca. 0°10'S, 1 14°08'E; observed 7-8 Sep. 1986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, left bank, at mouth of Sungai Beriwit; Borneo: Kalimantan, INDONESIA; ca. 0°10'S, 1 14°15'E; observed 4-7 Sep. 1986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, right bank, 1 km below mouth of Sungai Beriwit; Borneo: Kalimantan, IN- DONESIA; ca. 0°09'S, 1 14°15'E; observed 4-7 Sep. 1986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, right bank, 1 km below mouth of Sungai Turusan; Borneo: Kalimantan, IN- DONESIA; ca. 0°1 l'S, 1 14°09'E; observed 7-8 Sep. 1986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Murung, Sungai, right bank, 5 km above mouth of Sungai Turusan; Borneo: Kalimantan, IN- DONESIA; ca. 0°09'S, 114°12'E; observed 4-8 Sep. 1986 by D. J. Chivers and K M. Burton ([1991], p. 143). FAS; CK-19. Musi, Air, ca. 1 2 km below Palembang; Sumatra, INDONESIA; ca. 2°58'S, 104°51'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 15 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°54'S, 103°58'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 1 5 km above Palembang; Sumatra, INDONESIA; ca. 3°02'S, 104°37'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 24 km below Palembang; Sumatra, INDONESIA; ca. 2°53'S, 104°54'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 30 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°53'S, 103°51'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 30 km above Palembang; Sumatra, INDONESIA; ca. 3°04'S, 104°29'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 36 km below Palembang; Sumatra, INDONESIA; ca. 2°47'S, 104°56'E; observed Nov. 1 97 1-Jan. 1 973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 45 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°49'S, 103°44'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 48 km below Palembang; Sumatra, INDONESIA; ca. 2°41'S, 104°56'E; observed FOODEN: SYSTEMATIC REVIEW OF MAC AC A FASCICULARIS 155 Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 50 km above Palembang; Sumatra, INDONESIA; ca. 2°59'S, 104°19'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 60 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°45'S, 103°39'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 60 km below Palembang; Sumatra, INDONESIA; ca. 2°35'S, 104°56'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 70 km above Palembang; Sumatra, INDONESIA; ca. 3°03'S, 104°10'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 72 km below Palembang; Sumatra, INDONESIA; ca. 2°28'S, 104°56'E; observed Nov. 1 97 1-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 75 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°44'S, 103°32'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 84 km below Palembang; Sumatra, INDONESIA; ca. 2°22'S, 104°54'E; observed Nov. 1 97 1-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Musi, Air, ca. 85 km above Palembang; Sumatra, INDONESIA; ca. 2°57'S, 104°05'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, ca. 90 km above mouth of Batang Har- ileko; Sumatra, INDONESIA; ca. 2°43'S, 103°25'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-62. Musi, Air, near Palembang; Sumatra, INDONE- SIA; ca. 2°59'S, 104°45'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Mutis, Gunung, 2100 m; Pulau Timor, Lesser Sunda Islands, INDONESIA; 9°34'S, 124°14'E; collected 28 Feb.-22 Mar. 1932 by G. Stein; mzb, 1; zmb, 2. FAS; C.LS-28. MyengunKyun, BURMA; 19°50'-20°05'N, 92°55'- 93°02'E; collected in 1 847 by S. R. Tickell (1 854- 1875, p. [18]); museum unknown (not seen). AUR; A:Bu-2. Naka Yai, Ko, THAILAND; 8°03'-8°04'N, 98°28'- 98°29'E; collected 4 Feb. 1 9 1 8 by local collectors employed by H. C. Robinson and C. B. Kloss (1919, p. 87);zrc, 1. AUR/FAS/MUL; A:T-52. Nakhon Si Thammarat; THAILAND; 8°26'N, 99°58'E; collected 27 Mar. 1924 and 19-29 Sep. 1926 by H. M. Smith (see Riley, 1938, p. 10); usnm, 2 (including 1 skull only); zrc, 3. AUR/ FAS/MUL; A:T-63. Nakhorn Sawan. See Wat Khao Noh. Nakon Sritammarat. See Nakhon Si Thammarat. Nam Kam, Koh. See Nang Kham, Ko. Nanas, Bukit, Kuala Lumpur; MALAYSIA: WEST MALAYSIA; 3°09'N, 101°42'E; collected 11 Nov. 1913, 6 Jun. 1938, and date unknown by museum collectors; bm(nh), 3. Observed Mar.- Jun. 1966 by I. S. Bernstein (1966, p. 1559; 1968a, p. 121). Observed Aug.-Dec. 1970 by C. H. Southwick and F. C. Cadigan, Jr. (1972, p. 11). FAS;B:WM-19. Nancowry Island, Nicobar Islands, INDIA; 7°55'- 8°02'N, 93°30'-93°34'E; primates reported ab- sent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, 1903a, p. 1 14). A:l. Nang Kham, Ko; THAILAND; ca. 7°20'N, 100°22'E; collected 12 Apr. 1899 by R. Evans and F. F. Laidlaw (see P. H. Napier, 1981, p. 17; Bonhote, [1901], p. 870); University Mu- seum of Zoology, Cambridge, 1 (skull only, not seen). FAS; A:T-66. Nanga Look, near Nggoer; Pulau Flores, Lesser Sunda Islands, INDONESIA; ca. 8°42'S, 119°48'E; observed 1969-1973 by W. Auffen- berg (1981, p. 242). FAS; GLS-13. Nankauri Island. See Nancowry Island. NatunaBesar, Pulau, INDONESIA; 3°37'^°13'N, 107°59'-108°25'E; collected 17 Oct. 1893 by A. Everett (in Thomas & Hartert, 1894, p. 654; cf. W. Rothchild, 1894, p. 468); bm(nh), 1 (skin only, skull inside). Collected Aug. 1 894 and Jan. 1896 by C. Hose and E. Hose; bm(nh), 1 (skin only, skull inside); mcz, 2 (skins only, skulls in- side). FAS; B:SCS-32, SCS-33. Naunchik. See Nong Chik region. Nawngchik, State. See Nong Chik region. Negara, Taman; MALAYSIA: WEST MALAY- SIA; ca. 4°42'N, 102°28'E; reported present ca. 1986 by M. K. B. M. Khan (1988, p. 267). FAS; B:WM-11. Negris Island. See Hainggyi Kyun. Ngarai Sianok, Bukit Tinggi; Sumatra, INDO- NESIA; not located; observed Feb. 1990 by M. Bismark (1992, p. 11). FAS: not mapped. Ngawi; Java, INDONESIA; 7°24'S, 1 1 1°26'E; col- 156 FIELDIANA: ZOOLOGY lected 1887-1941 by E. Dubois; rmnh, 1 (skull only). FAS; OJ-30. Nggoer. See Nanga Look, near Nggoer. Nguwal, Pulau, Kepulauan Riau, INDONESIA; 0°38'-0°40'N, 104°13'-104°15'E; collected 3 Jan. 1925 by F. N. Chasen (1925, p. 93); zrc, 6. FAS; B:SCS-11. Niah Caves; Borneo, MALAYSIA: SARAWAK; 3°49'N, 1 13°47'E; reported present before 1959 by Lord Medway (1958, p. 634). FAS; C:Sar- 16. Niah National Park; Borneo, MALAYSIA: SA- RAWAK; ca. 4°00'N, 114°00'E; reported as probably present before 1983 by G. Davies ([1983], p. 148). FAS; C:Sar-17. Nias, Pulau, INDONESIA; 0°33'-l°32'N, 97°03'- 97°56'E; observed before 1864 by J. T. Nieu- wenhuisen and H. C. B. von Rosenberg (1863, p. 19). Collected before 1890 by E. Modigliani ( 1 889, p. 239); museum unknown, 1 8 specimens (not seen). Collected Nov. 1 904 by W. L. Abbott (cf. Lyon, 1916, p. 458); zrc, 1 (skull not ex- amined). Reported present before 1906 by T. Willink (1905, p. 175). Collected date unknown by H. Raap and R. Knuth; zmb, 2 (skulls only). Reported present Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). Blood samples taken from captives 1984-1989 by W. Scheffrahn, J. R. de Ruiter, and J. A. R. A. M. van Hooff (1994, p. 135). FAS; B:IO-6 through IO- 10. Nicobar Islands, INDIA; 6°45'-8°02'N, 93°20'- 93°57'E; collected ca. 1846 by H. Lewis (see Blyth, 1846, p. 366; J. Anderson, 1881, p. 65; Khajuria, [1955], p. 109); formerly preserved in zsi (not seen). Examined for malaria parasites in 1979 by L. Kalra (1980, p. 50). Examined for malaria parasites ca. 1984 by S. Sinha and A. Gajanana (1984, p. 567). UMB; A:N-1 through N-4. Nikiniki, [720 m]; Pulau Timor, Lesser Sunda Is- lands, INDONESIA; 9°49'S, 124°28'E; collected 28 Mar. 1932 by G. Stein (see Hellmayr, 1914, p. 5); mzb, 1; zmb, 1 (skin only). FAS; CLS-27. Nipah, Telok, vicinity; Pulau Tioman, MALAY- SIA: WEST MALAYSIA; ca. 2°44'N, 104°08'E; observed Mar.-Apr. 1962 by Lord Medway (1966, p. 16). FAS; B:SCS-3. Niur, Suaka Margasatwa Ujungkulon; Java, IN- DONESIA; 6°40'S, 105°21'E; observed 20 Aug. 1941 by A. Hoogerwerf (1970, p. 408). FAS; B:J-14. Noesa Penida. See Penida, Nusa. Nong Chik region; THAILAND; ca. 6°48'N, 101°12'E; collected 17 Sep. 1901 by N. Annan- dale and H. C. Robinson (1903, p. xxxiii; cf. Bonhote, 1903, p. 3); museum unknown, 1 (not seen). FAS; A:T-68. Nongkok. See Ban Nong Kok. North Andaman Island, INDIA; 12°53'-13°35'N, 92°48'-93°05'E; primates reported absent be- fore 1928 by C. B. Kloss ([1928], p. 802; cf. Chaturvedi, 1980, p. 134). A:a. North Borneo. See Sabah. North Natuna Island. See Laut, Pulau. North Pagi Island. See Pagai Utara, Pulau. Ntori, Sape district, 420 m; Pulau Sumbawa, Less- er Sunda Islands, INDONESIA; 8°33'S, 118°52'E; collected 2 Jun. 1927 by Sunda-Ex- pedition Rensch (see Mertens, 1930, p. 136); zmb, 1. FAS;C:LS-12. Nu Pau. See Nang Kham, Ko. Nuri Valley; MALAYSIA: WEST MALAYSIA; ca. 2°34'N, 102°15'E; captive obtained ca. 1953 by J. F. B. Edeson and D. G. Davey (1953, p. 259). FAS; B:WM-25. Nusa Mbarapu. See Mbarapu, Nusa. Nusa Pinda. See Pinda, Nusa. Nusa Rondong. See Rondong, Nusa. Nusa Vadju. See Vadju, Nusa. Nyalas; MALAYSIA: WEST MALAYSIA; 2°26'N, 102°28'E; collected 23 Oct. 1910 by museum collector; zrc, 3 (including 2 skulls only; exter- nal measurements recorded on skull tag and in Weitzel et al., 1988, p. 107). FAS; B:WM-27. Ober Langkat district; Sumatra, INDONESIA; ca. 3°45'N, 98°20'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-20. Ochodia, Naf River; BANGLADESH; ca. 2 1°05'N, 92°12'E; observed Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-l. Oeassa (?= Oebuah) vicinity; Pulau Semau, Lesser Sunda Islands, INDONESIA; ca. 10°07'S, 123°27'E; observed ca. May 1 859 by A. R. Wal- lace (1869, p. 294). FAS; GLS-24. Oebuah. See Oeassa. Oewada Sami, Pulau. See Kode, Nusa. Ok Yam. See Laem Ngop-Phumi Cham Yearn. Ol-kolo-kwak vicinity; Katchall Island, Nicobar Islands, INDIA; 7°52'-8°02'N, 93°18'-93°27'E; collected 21 Feb. 1901 by W. L. Abbott (see Miller, 1902b, p. 751; Kloss, 1903a, p. 114); usnm, 2. UMB; A:N-1. [Oo vicinity], Pulau Sumbawa, Lesser Sunda Is- lands, INDONESIA; ca. 8°25'S, 118°35'E; col- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 157 lected 27 Nov. 1909 by J. Elbert (1912, p. 58); nms, 1. FAS;C:LS-11. Outer Bolongs. See Myengun Kyun. Pacific Tin; MALAYSIA: WEST MALAYSIA; ca. 3°25'N, 101°25'E; examined for malaria ca. 1 96 1-1 962 by R. H. Wharton, D. E. Eyles, McW. Warren, and W. H. Cheong (1964, p. 58). FAS; B:WM-18. Padang; Sumatra, INDONESIA; 0°57'S, 100°21'E; collected in 1836 by S. Miiller; rmnh, 7 (skins only, skulls inside). Collected in 1890 by H. Meyer; zmuz, 2 (including 1 skin only). Col- lected date unknown by unknown collector; rmnh, 3; zmuz, 1 (skull only). Clinical exami- nation Aug.-Nov. 1980 by K. Matsubayashi and D. Sajuthi (1981, p. 48). FAS; B:S-47. Padang, Pulau, north coast, INDONESIA; ca. 1°22'N, 102°20'E; collected 27 Mar. 1906 by W. L. Abbott (field catalog; cf. Lyon, 1908, p. 624); usnm, 1 (skull only; external measurements re- corded on skull tag). FAS; B:SM-6. Padang/Bedagei district; Sumatra, INDONESIA; ca. 3°20'N, 99°10'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-23. Padang highlands; Sumatra, INDONESIA; ca. 0°57'S, 100°21'E; collected in 1878 by J. F. Snelleman(1887, p. 10); rmnh, 1. FAS; B:S-47. Padangsidempuan, ca. 45 km NNW; Sumatra, INDONESIA; ca. 1°44'N, 99°07'E; reported present Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:S- 34. Padangsidempuan vicinity; Sumatra, INDONE- SIA; ca. 1°22'N, 99°16'E; reported present Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:S-33. Padar, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°38'-8°42'S, 119°32'-119°37'E; monkeys reported absent Jul. 1 969-Jun. 1 970 by W. Auf- fenberg(1981,p. 242). C:e. Padar-kechil, Pulau. See Padar-kecil, Pulau. Padar-kecil, Pulau, Lesser Sunda Islands, IN- DONESIA; 8°4 l'S, 1 1 9°32'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:e. Padas Bay; Borneo, MALAYSIA: SABAH; ca. 5°13'N, 115°34'E; observed Jul.-Oct. 1968 by T. Mano (see Kawabe & Mano, 1972, p. 215). FAS;C:Sab-l. Padeco, Hak Pengusahaan Hutan (HPH); Suma- tra, INDONESIA; not located; observed Dec. 1988 by M. Bismark (1992, p. 11). FAS; not mapped. Pagai Selatan, Pulau, INDONESIA; 2°47'-3°21'S, 100°10'-100°28'E; M. fascicularis reported ab- sent before 1 928 by C. B. Kloss ([ 1928], p. 802). B:i. Pagai Utara, Pulau, INDONESIA; 2°30'-2°52'S, 99°58'-100°13'E; M. fascicularis reported ab- sent before 1 928 by C. B. Kloss ([ 1 928], p. 802). B:h. Pagansan; Sumatra, INDONESIA; ca. 1°00'N, lOTOO'E; collected ca. 1907 by M. Moszkowski (1909, maps following pp. 192, 328); zmb, 4 (skeletons only). FAS; B:S-40. Pagat. See Hantakan, 3 km N of Pagat. Pagurawan. See Paguruan, Sungai. Paguruan, Sungai; Sumatra, INDONESIA; ca. 3°26'N, 99°20'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-23. Pai, Teluk; Pulau Karimata, INDONESIA; 1°32'- 1°40'S, 108°48'-108°59'E; collected 24-27 Aug. 1904 by W. L. Abbott (see Miller, 1906b, p. 55); usnm, 2. FAS; B:SCS-22. Paillin. See Pang Roloem-Sur Sdei area. Pajakombo. See Payakumbuh. Pajeti. See Payeti-Kambaniru. Pajo, Danau Singkarak, Balipor district; Sumatra, INDONESIA; 0°37'S, 100°30'E; collected be- fore 1880 by C. Bock (see P. H. Napier, 1981, p. 16); bm(nh), 1. FAS; B:S-46. Pakan Selasa; Sumatra, INDONESIA; 1°34'S, 101°08'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-52. Pakan Selasa, ca. 10 km SSE; Sumatra, INDO- NESIA; ca. 1°39'S, lOTlO'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-52. Pakchan River, near Bankachon; BURMA; ca. 10°09'N, 98°36'E; collected 29 Jan. 1914 and Dec. 1913-Apr. 1914 by G. C. Shortridge (in Wroughton, 1915, p. 696); bm(nh), 2. AUR/ FAS/MUL; A:Bu-22. Pakchan River, near Maliwun; BURMA; 10°14'N, 98°37'E; collected 28 Feb. 1928 by Faunthorpe- Vernay Expedition; amnh, 1 . AUR/FAS/MUL; A:Bu-22. Pak Chong, Sathani; THAILAND; 14°42'N, 101°25'E; collected 29 Feb. 1924 by H. M. Smith (see Kloss, 1916a, p. 2; Riley, 1938, p. l;Weitzel et al., 1988, p. 106); zrc, 1. Collected 21 May 1 93 1 by C. J. Aagaard; zrc, 1 . AUR/FAS/MUL; A:T-25. Pak Jong. See Pak Chong, Sathani. Pak Klong Pran. See Pran Buri, Mae Nam, mouth. 158 FIELDIANA: ZOOLOGY Pak Nam Chumphon. See Chumphon, Khlong, mouth. Paknampoh. See Ban Pak Nam Pho. Paku. See Pelandok, Sungai, Paku, Saribas. Paku, Saribas; Borneo, MALAYSIA: SARAWAK; 1°27'N, 1H°27'E; collected Mar. 1917 by C. Chunggat and H. C. Robinson (see P. H. Napier, 1981, p. 14); bm(nh), 2. FAS; C:Sar-9. Paku Cave; Borneo, MALAYSIA: SARAWAK; 1°25'N, 1 10°1 l'E; collected 1870 and 1879, prob- ably by A. H. Everett; bm(nh), Subdepartment of Anthropology, Department of Palaeontology, 2 (1 skeleton only, 1 skull only; not seen, data from P. H. Napier, 1981, p. 20). FAS; C:Sar-3. Palembang, ca. 30 km N; Sumatra, INDONESIA; ca. 2°43'S, 104°46'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 30 km NNE; Sumatra, INDO- NESIA; ca. 2°44'S, 104°51'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 35 km SW; Sumatra, INDONE- SIA; ca. 3°13'S, 104°33'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 40 km N; Sumatra, INDONESIA; ca. 2°38'S, 104°48'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 40 km NNW; Sumatra, INDO- NESIA; ca. 2°39'S, 104°34'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 45 km N; Sumatra, INDONESIA; ca. 2°36'S, 104°43'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 45 km NW; Sumatra, INDO- NESIA; ca. 2°42'S, 104°28'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 50 km N; Sumatra, INDONESIA; ca. 2°32'S, 104°48'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1 980, p. 156). FAS;B:S-61. Palembang, ca. 60 km NNE; Sumatra, INDO- NESIA; ca. 2°27'S, 104°51'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 60 km NNW; Sumatra, INDO- NESIA; ca. 2°32'S, 104°26'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 60 km NW; Sumatra, INDO- NESIA; ca. 2°37'S, 104°21'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 60 km SW; Sumatra, INDONE- SIA; ca. 3°20'S, 104°21'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-75. Palembang, ca. 65 km N; Sumatra, INDONESIA; ca. 2°24'S, 104°41'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 65 km NNW; Sumatra, INDO- NESIA; ca. 2°27'S, 104°32'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 65 km SW; Sumatra, INDONE- SIA; ca. 3°20'S, 104°16'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-75. Palembang, ca. 70 km N; Sumatra, INDONESIA; ca. 2°20'S, 104°42'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1 980, p. 156). FAS;B:S-61. Palembang, ca. 70 km NNW; Sumatra, INDO- NESIA; ca. 2°29'S, 104°23'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-61. Palembang, ca. 70 km SW; Sumatra, INDONE- SIA; ca. 3°23'S, 104°16'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-75. Palembang, ca. 80 km N; Sumatra, INDONESIA; ca. 2°15'S, 104°48'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 90 km WSW; Sumatra, INDO- NESIA; ca. 3°19'S, 104°01'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-63. Palembang, ca. 95 km SW; Sumatra, INDONE- SIA; ca. 3°28'S, 104°05'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-63. Palembang, ca. 100 km N; Sumatra, INDONE- SIA; ca. 2°05'S, 104°47'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-61. Palembang, ca. 100 km SW; Sumatra, INDO- NESIA; ca. 3°26'S, 103°58'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-63. Palembang district; Sumatra, INDONESIA; ca. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 159 2°55'S, 104°45'E; collected in 1905 by B. Hagen; zsbs, 2 (skins only). FAS; B:S-61. Palung, Gunung. See Gunung Palung Nature Re- serve. Pamuja, Tanjung; Pulau Bangka, INDONESIA; ca. 1°40'S, 105°20'E; collected 19 Jun. 1904 by W. L. Abbott (in Lyon, 1906, p. 607); usnm, 1. FAS;B:SCS-18. Panaitan, Pulau, INDONESIA; 6°32'-6°46'S, 105°07'-105°16'E; observed 1932-1957 by A. Hoogerwerf (1970, p. 408). FAS; B:J-14. Pandan, Sungai; Singapore Island; SINGAPORE; ca. 1°19'N, 103°45'E; observed before 1901 by S. S. Flower (1900, p. 316). FAS; B:SCS-7. Pandang, Sungei. See Pandan, Sungai. Panebangan, Pulau. See Penebangan, Pulau. Pangadaran. See Pangandaran. Pangandaran, Teluk Parigi; Java, INDONESIA; 7°41'S, 108°39'E; collected 4-11 Apr. 1908 by G. C. Shortridge (see Thomas & Wroughton, 1909a, p. 373); bm(nh), 3 (including 1 skin only, 1 skull only [with mounted skin, not examined]). Clinical examination in 1980 by K. Matsubay- ashi and D. Sajuthi (1981, p. 48). FAS; B:J-2. Pangerango, Mt. See Pangrango, Gunung. Pangkalan; Padang highlands; Sumatra, INDO- NESIA; ca. 0°57'S, 100°21'E; collected ca. 1889 by E. Dubois; rmnh, 3 (skulls only). FAS; B:S- 47. Pangkalansusu, Sumatra, INDONESIA; 4°07'N, 98°13'E; collected 15 Jun. 1972 by P. F. D. Van Peenen; usnm, 1 (infant in fluid). Collected be- fore 1979 by NAMRU 2 Djakarta Detachment; usnm, 1 (skull only). FAS; B:S-19. Pangkor, Pulau, MALAYSIA: WEST MALAY- SIA; 4°12'-4°16'N, 100°32'-100°35'E; reported present ca. 1963 by J. L. Harrison and J. R. Hendrickson (1963, p. 548). FAS; B:SM-3. Pangonan, Gunung Muria; Java, INDONESIA; ca. 6°36'S, 1 10°53'E; collected 19 Dec. 1928 by H. J. V. Sody; rmnh, 1. FAS; CJ-28. Pangrango, Gunung; Java, INDONESIA; 6°46'S, 106°57'E; collected in 1899 by Bartels; nhrm, 27 (skulls only). Collected 4 Jul. 1902 and Feb. 1 903 by unknown collectors; rmnh, 2 (skins only, skulls inside). FAS; B:J-18. Pangrolim. See Pang Roloem-Sur Sdei area. Pang Roloem-Sur Sdei area; CAMBODIA; ca. 12°50'N, 102°45'E; observed Oct. 1962 by D. E. Eyles, R. H. Wharton, W. H. Cheong, and McW. Warren (1964, p. 9). FAS; A:C-2. Panjaer, Tanjong. See Tanjong Panjair, Sungai Rompin. Panjang, Pulau. See Mentigi. Pantai Krachut; Pulau Pinang [1], MALAYSIA: WEST MALAYSIA; 5°27'N, 100°1 l'E; collect- ed 24 Mar. 1911 by E. Seimund (see Weitzel et al., 1988, p. 1 14); zrc, 1. FAS; B:SM-2. Pantar, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°11'-8°33'S, 123°55'-124°19'E; M.fasci- cularis reported absent before 1937 by J. J. M. F. Symons (Mertens, 1936, p. 319). C:g. Papagaran. See Papagaran Besar, Pulau. Papagaran Besar, Pulau, Lesser Sunda Islands, INDONESIA; 8°34'-8°35'S, 1 19°43'-1 19°44'E; monkeys reported absent 1969-1973 by W. Auf- fenberg(1981,p. 40). C:d. Papar, 20 mi (= 32 km) SW of Kota Kinabalu; Borneo, MALAYSIA: SABAH; 5°44'N, 1 1 5°56'E; collected 1 2 Sep. 1 960 by R. E. Kuntz (1969, p. 193); usnm, 1. FAS; C:Sab-3. Pap-ka; Sumatra, INDONESIA; ca. TOO'N, 101WE; collected Apr. 1908 by M. Mosz- kowski (1909, maps following pp. 192, 328); zmb, 1 (skull only). FAS; B:S-40. Parepare; Pulau Sulawesi, INDONESIA; 4°01'S, 1 19°38'E; captive (presumably introduced) ob- served 6-15 Oct. 1888 by M. Weber (1890a, p. v; 1890b, p. 102). Subspecies uncertain; not mapped. Parit, primary swamp forest, sea level; Borneo: Kalimantan, INDONESIA; 2°10'S, 113°00'E; collected 15 Jun.-16 Jul. 1935 by J. J. Menden; amnh, 20 (incuding 1 skull not examined, mea- surements from G. H. Albrecht, pers. comm., Oct. 1991); mzb, 3; zmb, 4 (including 1 skin only). FAS; C:K-12. Pasar Besar, Hak Pengusahaan Hutan (HPH); Su- matra, INDONESIA; not located; observed Jul. 1989 by M. Bismark (1992, p. 11). FAS; not mapped. Pasar Usang area; Sumatra, INDONESIA; ca. 1°00'S, 100°25'E; blood samples taken Jan. 1 979-Dec. 1 98 1 by Y. Kawamoto ( 1 982, p. 67). FAS; B:S-47. Pasir Carolina, Purwakarta district; Java, IN- DONESIA; ca. 6°34'S, 107°26'E; collected 17 Aug. 1932 by Kloster; mzb, 1. FAS; B:J-20. Pasir Panjang; Pulau Bintan, Kepulauan Riau, IN- DONESIA; ca. 1°10'N, 104°30'E; collected 7 Aug. 1902 by W. L. Abbott (field catalog; cf. Miller, 1906c, p. 282); usnm, 1. Collected 8 Jun. 1 908 by H. C. Robinson (in Thomas & Wrough- ton, 1909b, p. 104); bm(nh), 1; zrc, 1 (skull only). FAS; B:SCS-8. Pasoh Forest Reserve, 50 m; MALAYSIA: WEST MALAYSIA; ca. 2°57'N, 102°22'E; observed Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. 160 FIELDIANA: ZOOLOGY Wilson (1981, p. 232). Observed Jan. 1979-Jul. 1980 by J. O. Caldecott (1986b, p. 21). FAS; B:WM-20. Patani. See Pattani. Patani, Cape. See Pho, Laem. Patong, Bukit, ca. 1 km west; MALAYSIA: WEST MALAYSIA; ca. 3°43'N, 102°02'E; observed Jul.-Dec. 1977 by D. Chivers and G. Davies (1979, p. 22). FAS; B:WM-15. Pattani; THAILAND; 6°52'N, 101°16'E; collected 2 Jun. 1901 by H. C. Robinson (see Annandale & Robinson, 1903, p. xxi; Bonhote, 1903, p. 3); smtd, 1 (skull only). FAS; A:T-68. Payakumbuh, Padang highlands; Sumatra, IN- DONESIA; ca. 0°14'S, 100°38'E; collected ca. 1889 by E. Dubois; rmnh, 1 (skull only). FAS; B:S-45. Payeti-Kambaniru; Pulau Sumba, Lesser Sunda Islands, INDONESIA; 9°39'S, 120°18'E; col- lected 26-27 Mar. 1925 by P. F. Franck (see Dammerman, 1926a, p. 38); mzb, 2. FAS; C:LS- 22. Pegou. See Pegu. Pegu; BURMA; 17°20'N, 96°29'E; obtained in 1827-1828 by A. A. M. Reynaud (see de Rossel et al., 1829, p. 603; de Rossel, 1829, p. 609; I. Geoffroy, 1831, pp. 58, 77); museum unknown, 1 (not seen). Observed before 1889 by W. T. Blanford (1888b, p. 22). AUR; A:Bu-8. Pejantan, Pulau, INDONESIA; 0°07'-0°09'N, 107°12'-107°15'E; observed 1-2 Aug. 1899 by W. L. Abbott (in Miller, 1900, p. 243; cf. Kloss, 1903b, p. 59). Collected 15 Mar. 1907 by C. Bruegel; zsbs, 1 (skin only). FAS; B:SCS-25. Pekan vicinity; MALAYSIA: WEST MALAY- SIA; ca. 3°30'N, 103°25'E; observed 25-28 Jun. 1891 by H. N. Ridley, W. Davison, and H. J. Kelsall (see Kelsall, 1894a, p. 34). FAS; B:WM- 13. Pelabuhanratu, Teluk, 100 ft (= 30 m); Java, IN- DONESIA; 6°58'S, 106°32'E; collected 18 Mar. 1920 by C. B. Kloss (see Weitzel et al., 1988, p. 142); zrc, l.FAS;B:J-9. Pelabuhanratu, Teluk, Bantam region; Java, IN- DONESIA; ca. 6°58'S, 106°32'E; collected 10 Nov. 1909 by O. Bryant (see field catalog of W. Palmer); usnm, 1. FAS; B:J-9. Pelaihari; Borneo: Kalimantan, INDONESIA; 3°48'S, 1 14°45'E; collected 20 Aug. 1866 by J. Semmelink; rmnh, 1 (skin only, skull inside). Collected before 1897 by J. H. F. Kohlbrugge (1896a, p. 184); museum unknown, 1 (not ex- amined). FAS; C:K-26. Pelandok, Sungai, Paku, Saribas; Borneo, MALAY- SIA: SARAWAK; 1°32'N, 1 10°44'E; collected 15 Aug. 1916 by H. C. Robinson (see Weitzel et al., 1988, p. 142); zrc, 1. FAS; C:Sar-6. Pelapis Tengah, Pulau, INDONESIA; 1°17'- 1°20'S, 109°08'-109o10'E; reported present 29 May-1 June 1907 by W. L. Abbott (in Lyon, 1 9 1 1 , p. 60). Collected 2 1 Mar. 1 93 1 by L. Coo- mans de Ruiter and Madzoed (see Chasen, 1935b, p. 2); mzb, 1. FAS; B:SCS-23. Pelawan, Sungai; Borneo: Kalimantan, INDO- NESIA; 1°11'N, 117°55'E; collected 18 Dec. 1913 by H. C. Raven (field catalog; cf. Deignan, [1960], p. 269); usnm, 1. FAS; C:K-44. Pelehari. See Pelaihari. Peling, Pulo. See Piling, Pulau. Pemanggil, Pulau, MALAYSIA: WEST MALAY- SIA; 2°34'-2°36'N, 104°18'-104°21'E; collected 12 Jun. 1901 by W. L. Abbott (see Riley, 1938, p. 14); usnm, 1. Collected 12 Jul. 1915 by H. C. Robinson; bm(nh), 1. FAS; B:SCS-4. Pemangil Island. See Pemanggil, Pulau. Pematangsiantar, Sumatra, INDONESIA; 2°57'N, 99°03'E; collected Jul. 1937 by B. Lawrence; mcz, 2. FAS; B:S-30. Pemeral; Pulau Karimun, Kepulauan Riau, IN- DONESIA; 1°00'N, 103°25'E; collected 1 1 Aug. 1 908 by H. C. Robinson (in Thomas & Wrough- ton, 1909b, pp. 105, 107) and E. Seimund; bm(nh), 4. FAS; B:SCS-17. Penang Botanical Gardens; Pulau PinangflJ, MA- LAYSIA: WEST MALAYSIA; ca. 5°25'N, 100°20'E; observed 1940-1961 by K. C. Cheang (1962, p. 73). Observed Jul.-Sep. 1960 by Y. Furuya (196 1-1 962a, p. 76). Observed in 1974 by C. Spencer (1975, p. 83). FAS; B:SM-2. Penang Hill; Pulau Pinang [1], MALAYSIA: WEST MALAYSIA; ca. 5°26'N, 100°16'E; re- ported present before 1 976 by C. Spencer (1975, p. 85). FAS; B:SM-2. Penang Island. See Pinang, Pulau [1]. Penang Waterfall Gardens. See Penang Botanical Gardens. Penebangan, Pulau, INDONESIA; l°10'-ri3'S, 109°13'-109°16'E; observed 16 May-21 Sep. 1907 by W. L. Abbott (in Lyon, 1911, p. 60). FAS; B:SCS-23. Pengiki, Pulo. See Pejantan, Pulau. Pengsong, Gunung; Pulau Lombok, Lesser Sunda Islands, INDONESIA; ca. 8°36'S, 116°06'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Kawamoto (1982, p. 66). FAS; C:LS-5. Penida, Nusa, 300 m, Lesser Sunda Islands, IN- DONESIA; 8°43'-8°48'S, 115°30'-115°36'E; FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 161 collected 26 Feb. 1938 by V. von Plessen; amnh, 3. FAS; C:LS-4. Penrissen, Gunung, 4000 ft (= 1 200 m); Borneo, MALAYSIA: SARAWAK; 1°07'N, 110°13'E; collected May 1899 by E. A. W. Cox and R. W. L. Shelford (see Shelford, 1 9 1 6, p. 2 1 7; cf. Med- way, 1 977, p. 7 1 , who cited altitude as 4500 ft); smk, 2 (skins only, skulls inside). FAS; C:Sar-2. Penuba. See Selayar, Pulau. Pepidon, Ko. See Phi Phi Don, Ko. Perak, Sungai, upper. See Pong, upper Sungai Per- ak. Perangin, Bukit, 75-360 m; MALAYSIA: WEST MALAYSIA; 6°22'N, 100°31'E; reported as probably present Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-3. Perawang, 4 km NW; Sumatra, INDONESIA; ca. 0°40'N, 10 1°35'E; observed Sep.-Nov. 1981 and Apr. 1983-Sep. 1984 by E.N. Megantara(1989, p. 183). FAS;B:S-42. Perboewah, Kp. See Perbuah, Sungai Landak. Perbuah, Sungai Landak, 900 m; Borneo: Kali- mantan, INDONESIA; 0°53'N, 110°10'E; col- lected 27 Jul. -10 Aug. 1937 by J. J. Menden; amnh, 9 (including 1 skin only); mzb, 2. FAS; C:K-4. Perlis, Sungai, sea level; MALAYSIA: WEST MA- LAYSIA; 6°24'N, 1 00° 10'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-2. Petchaburi district. See "Siam." Peucang, Pulau, INDONESIA; 6°43'-6°45'S, 105°14'-105°16'E; observed 1932-1957 by A. Hoogerwerf (1970, p. 408). Observed Sep. 1969- Jan. 1971 by W. Angst (1973, p. 626; 1975, p. 326). FAS; B:J-14. Peutjang, Pulau. See Peucang, Pulau. Pexabury. See Phet Buri. Phai, Ko, THAILAND; 12°55'-12°57'N, 100°40'- 100°41'E; primates reported as apparently ab- sent Jan. 1915 by collectors employed by C. B. Kloss (1915a, p. 157; 1915b, p. 221). A:m. Phangan, Ko, southwest, THAILAND; ca. 9°40'N, 100°00'E; collected 27 May 1913 by H. C. Rob- inson (1915, p. 129) and E. Seimund; zrc, 2. FAS; A:T-50. Phangnga Bay National Park; THAILAND; ca. 8°25'N, 98°37'E; observed 17-21 Nov. 1987 by R. Boonratana (1988, p. 76); also seen on un- specified adjacent islands. Subspecies uncertain; A:T-51. Phattalung. See Wat Suwankuha. Phayam, Ko, THAILAND; 9°42'-9°46'N, 98°23'- 98°26'E; collected 20 Feb. 1919 by H. C. Rob- inson and C. B. Kloss; zrc, 1 . AUR/FAS/MUL; A:T-45. Phet Buri; THAILAND; 13°06'N, 99°57'E; col- lected in 1862 by M. Boucourt; mnhn, 1 (skin only). Subspecies uncertain; A:T-37. Phi Phi Don, Ko, THAILAND; 7°43'-7°47'N, 98°46'-98°48'E; collected 3-5 Feb. 1919 by H. C. Robinson (1921, p. 5) and C. B. Kloss; zrc, 4. FAS; A.T-54. Pho, Laem; THAILAND; ca. 6°57'N, 101°16'E; reported present Jun. 1901 by N. Annandale and H. C. Robinson (1903, p. xl). FAS; A:T-68. Phu Phan, Maung district; THAILAND; 1 6°42'N, 104°25'E; collected date unknown by R. E. El- bel; usnm, 1 (skin No. 307714, skull No. 307732). AUR/FAS/MUL; A:T-15. Phu-Qui. See Phu Quoc, Dao. Phu Quoc, Dao, VIETNAM; 10°00'-10°27'N, 103°50'-104°05'E; observed 1873-1874 by A. Morice (1876, p. 31). Collected 30 Dec. 1927 by J. Delacour, W. P. Lowe, [and P. Jabouille] (in Delacour, 1928, p. 265); bm(nh), 1; mnhn, l.FAS; A:V-6. Phu-Quoi. See Phu Quoc, Dao. Piling, Pulau, INDONESIA; 2°44'-2°46'N, 106°10'-106°12'E; observed 17 Aug. 1899 by W. L. Abbott and C. B. Kloss (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 68). FAS; B:SCS-27. Pinagar; Sumatra, INDONESIA; OWN, 99°55'E; observed Nov. 1971-Jan. 1 973 by C. M. Crock- ett and W. L. Wilson (1980, p. 156). FAS; B:S- 43. Pinagar, ca. 5 km SW; Sumatra, INDONESIA; ca. 0°07'N, 99°52'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-43. Pinagar, ca. 20 km E; Sumatra, INDONESIA; ca. OWN, 100°04'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-43. Pinang, Pulau [1], up to 2000 ft (= 600 m) MA- LAYSIA: WEST MALAYSIA; 5°15'-5°29'N, 100°11'-100°20'E; reported present before 1847 by T. Cantor ( 1 846, p. 1 76). Observed before 1 90 1 by S. S. Flower (1900, p. 316). FAS; B:SM-2. Pinang, Pulau [2], MALAYSIA: WEST MALAY- SIA; 5°44'-5°45'N, 103°00'-103o01'E; collected 3 Sep. 1910 by C. B. Kloss (1911a, p. 177); bm(nh), 2. FAS;B:SCS-1. Pinda, Nusa, Komodo area, Lesser Sunda Islands, INDONESIA; not located, ca. 8°30'S, 1 19°30'E; 162 FIELDIANA: ZOOLOGY monkeys reported absent 1 969-1 973 by W. Auf- fenberg (1981, p. 40). Not mapped. Ping, Mae Nam; THAILAND; ca. 16°00'N, 99°50'E; collected 7 Mar. 1924 by A. S. Vernay (see Lowe, 1932, p. 202; 1933, p. 260); amnh, 1 (skull only; external measurements recorded in amnh catalog). Subspecies uncertain; A:T-4. Pintu Gedong, Pulau, MALAYSIA: WEST MA- LAYSIA; 2°53'-2°56'N, 101°14'-101°16'E; col- lected 2-3 Nov. 1912 by museum collector; bm(nh), 1; zrc, 2. FAS; B:SM-4. P. Kemoedjan. See Kemujan, Pulau. Plaine Sophie; Mauritius Island, MAURITIUS; 19°59'-20°31'S, 57°18'-57°48'E; introduced population, collected 25 Apr. 1949 by R. d'Un- ienville; bm(nh), 1. Subspecies uncertain; not mapped. P. Lankawi. See Langkawi, Pulau. Plateau des Bolovens. See Thateng, Muang, Pla- teau des Bolovens. Pleihari Tanah Laut Game Sanctuary; Borneo: Ka- limantan, INDONESIA; ca. 3°48'S, 114°45'E; reported present before 1983 by G. Da vies ([1983], p. 148). FAS; GK-26. Pleyharie. See Pelaihari. Podowk. See Ataran River. Poelau (?= Kuda), Sungai Sibau; Borneo: Kali- mantan, INDONESIA; ca. 1°02'N, 112°59'E; collected 19 Jun. 1894 by J. Buttikofer (1897, p. 20); rmnh, 1 (skin only). FAS; GK-18. Pong, upper Sungai Perak, 900 ft (= 275 m); MA- LAYSIA: WEST MALAYSIA; 5°36'N, 101°02'E; collected 16 Jan. 1932 by A. B. Hol- loway and A. S. Vernay (see P. H. Napier, 1981, p. 14); bm(nh), 1. FAS; B:WM-4. Ponggol. See Punggol. Pongka, Kampung; Pulau Karimun, Kepulauan Riau, INDONESIA; 1°06'N, 103°24'E; collect- ed 25 May 1903 by W. L. Abbott (field catalog; cf. Miller, 1906c, p. 277); usnm, 1 (skull only). FAS;B:SCS-17. Pontianak. See Ambawang, Sungai, near Pontia- nak. Port Dickson; MALAYSIA: WEST MALAYSIA; 2°31'N, 101°48'E; collected 6 Jan.-2 May 1937 by E. Seimund; zrc, 3. Collected before 1940 by F. Colyer; bm(nh) (skin)/RCS(OM) (skull), 1. FAS; B:WM-24. Port Swettenham vicinity; MALAYSIA: WEST MALAYSIA; ca. 3°00'N, 101°24'E; reported present Aug.-Dec. 1970 by unspecified inform- ants (Southwick & Cadigan, 1972, p. 10). FAS; B:WM-18. Pota; Pulau Flores, Lesser Sunda Islands, IN- DONESIA; 8°20'S, 120°46'E; observed Apr.- May 1930 by Jhr. W. C. van Heurn (1932, p. 65). FAS; GLS-18. Poulo Condor. See Con Son. Prachuap Khiri Khan, few miles north; THAI- LAND; ca. 1 1°50'N, 99°48'E; observed Nov. 1914-Feb. 1915 by N. Gyldenstolpe (1916, p. 10; 1917b, p. 6). AUR/FAS/MUL; A:T-41. Prahmon. See Ban Phra Muang. Pran Buri, Mae Nam, mouth; THAILAND; 12°24'N, 100°00'E; collected 28 Jun. 1917 by W. J. F. Williamson and M. A. Smith (see Kloss, 1917, p. 289); zrc, 2. AUR/FAS/MUL; A:T- 38. Prang Koo; THAILAND; 15°40'N, 104°23'E; ob- served 17-18 Jul. 1989 by N. Aggimarangsee (1992, pp. 109, 120; pers. comm., Oct. 1993). Subspecies uncertain; A:T-16. Preanger. See Ciwangi; Tasikmalaya, Preanger re- gion; Tjeringin, near Banjar, Preanger region. Preanger district; Java, INDONESIA; 6°36'- 7°48'S, 106°22'-108°46'E; collected 12 Nov. 1890-8 Feb. 1891 by A. A. W. Hubrecht (see Zuckerman, [1933], p. 1062); museum un- known (not seen). FAS; B:J-6. Preparis Island, BURMA; 14°51'-14°53'N, 93°40'- 93°42'E; reported present before 1929 by C. B. Kloss ([1928], p. 806). AUR; A:Bu-5. "P. Sember"; Java, INDONESIA; ca. 6°50'S, 108°15'E; clinical examination in 1980 by K. Matsubayashi and D. Sajuthi (1 98 1 , p. 48). FAS; B:J-22. Pulaki, Tanjung; Pulau Bali, Lesser Sunda Islands, INDONESIA; 8°07'S, 114°35'E; observed Sep. 1969-Jan. 1971 by W. Angst (1975, p. 326). Blood samples taken in 1980 by Y. Kawamoto, K. Nozawa, and Tb. M. Ischak (1981, p. 16). FAS; C:LS-1. Pulau Doem area; Java, INDONESIA; ca. 7°35'S, 112°30'E; blood samples taken 1979-1981 by Y. Kawamoto (1982, p. 67). FAS; GJ-37. Pulau Kaban. See Acheh, Pulau. Pulau Lontar. See Lanta Yai, Ko. Pulau Misa. See Misa, Pulau. Pulau Muntia. See Muk, Ko. Pulau Panjang. See Yao Noi, Ko. Pulau Panjang Anak. See Yao Noi, Ko. Pulau Sulawesi, southern, INDONESIA; ca. 5°00'S, 120°00'E; introduced, reported present before 1883 by H. von Rosenberg (1882, p. 1 14). Re- ported present before 1930 by K. W. Dammer- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 163 man (1929, p. 6). Subspecies uncertain; not mapped. Pulau Telebong. See Talibong, Ko. Pulau Telibun. See Talibong, Ko. Pulo Beng Kalis. See Kapos Tinggi. Pulo-Condore. See Con Son. Pulo Gilla. See Jela, Pulau. Pulo Mata. See Matak, Pulau. Pulo Mobur. See Mubur, Pulau. Pulo Nyur; Great Nicobar Island, west coast, IN- DIA; ca. 6°55'N, 93°40'E; observed 2 1 Mar. 1 90 1 by C. B. Kloss (1903a, p. 150). UMB; A:N-3. Pulo Peling. See Piling, Pulau. Pulo Pengiki. See Pejantan, Pulau. Pulo Piling. See Piling, Pulau. Pulo Riabu. See Airabu, Pulau. Pulo Selindang. See Selintang, Pulau. Pulo Terutau. See Tarutao, Ko. Pulo Tioman. See Tioman, Pulau. Pulu Tikus. See Tikus, Pulu. Punang, Sungai; Borneo, MALAYSIA: SARA- WAK; 4°54'N, 1 15°21'E; collected 16 Apr. and 14-15 May 1939 by G. Nunong; smk, 2 (skins only); zrc, 2. FAS; C:Sar-22. Punggol; Singapore Island, SINGAPORE; 1°25'N, 103°54'E; collected 8-11 Nov. 1910 by A. H. Wong; bm(nh), 1; zrc, 4-086, 1. Collected 15- 1 6 Dec. 1 9 1 0 by unknown collector; zrc, 2 (skins only). FAS; B:SCS-7. Punoebo. See Selayar, Pulau. Puram; Borneo, MALAYSIA: SARAWAK; not located, 0°43'^°58'N, 109°34'-115°38'E; col- lected Jul. 1896 by W. H. Furness III (1896, p. 310) and H. M. Hiller (1896, p. 321); ansp, 1 (skull only). FAS; not mapped. Pura Uluwatu, Bukit Peninsula; Pulau Bali, south- western tip, Lesser Sunda Islands, INDONE- SIA; not located, 8°04'-8°48'S, 114°26'- 115°42'E; observed Jul.-Sep. 1993 by M. F. Small (1994, p. 8). FAS; not mapped. Purukcahu, Sungai Barito; Borneo: Kalimantan, INDONESIA; 0°35'S, 114°35'E; collected be- fore 191 1 by G. C. Shortridge; bm(nh), 1 (skull only). Observed 1979-1986 by J. Marshall, D. J. Chivers, and K. M. Burton (see Chivers & Burton, [1991], p. 141). FAS; QK-33. Pussuk forest. See Pusuk forest. Pusuk, Gunung; Pulau Lombok, Lesser Sunda Is- lands, INDONESIA; ca. 8°27'S, 1 16°07'E; blood samples taken Jan. 1979-Dec. 1981 by Y. Ka- wamoto (1982, p. 66). FAS; CLS-5. Pusuk forest; Pulau Lombok, Lesser Sunda Is- lands, INDONESIA; 8°25'S, 1 1 6°3 1 'E; collected 1 1 Apr. 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 46; Mertens, 1930, p. 129; I. Rensch, 1934, p. 227; Elbert, 1911, map 2); mzb, 1. FAS; GLS-7. Putussibau; Borneo: Kalimantan, INDONESIA; 0°50'N, 112°56'E; collected 19 Nov.-17 Dec. 1 897 by A. Harrison, Jr., and H. M. Hiller; ansp, 3. FAS; CK-17. Raba; Pulau Sumbawa, Lesser Sunda Islands, IN- DONESIA; 8°27'S, 118°46'E; blood samples taken Jul. 1981-Jan. 1 982 by Y.Kawamoto, Tb. M. Ischak, and J. Supriatna (1982a, p. 58; cf. Kawamoto, 1982, p. 66; USBGN Gazetteer of Indonesia, 1982, p. 1036). FAS; C:LS-12. Rachaburi. See Tham Chomphon, Wat Huai Tak- haeng, and Wat Ratch Singkhorn. Rakata, Pulau, INDONESIA; 6°08'-6°10'S, 105°25'-105°28'E; primates reported absent 1883-1934 by K. W. Dammerman (1948, p. 315); volcanic explosion in 1883. B:k. Rakata-kecil, Pulau, INDONESIA; 6°05'-6°06'S, 105°26'-105°27'E; primates reported absent 1883-1934 by K. W. Dammerman (1948, p. 55); volcanic explosion in 1883. B:k. Rana Mese, 1250-1400 m; Pulau Flores, Lesser Sunda Islands, INDONESIA; 8°38'S, 120°34'E; collected 19 Jun. 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 148; Mertens, 1930, p. 139; I. Rensch, 1934, p. 227); nms, 1. FAS; GLS-15. Ranau, ca. 1500 ft (= 450 m); Borneo, MALAY- SIA: SABAH; 5°57'N, 116°41'E; collected 23 July 1937 by J. A. Griswold, Jr. (see Coolidge, 1940, p. 123);mcz, 1. Collected 19-23 Sep. 1960 by R. E. Kuntz (1969, p. 193); usnm, 5. FAS; C:Sab-7. "Rangoon"; BURMA; 16°47'N, 96°10'E; captive acquired 16 Dec. 1870 by unknown collector; rmnh, 1 (skin only). Captive acquired 8 May 1 875 by unknown collector; rmnh, 1 (skin only). Subspecies uncertain; A:Bu-7. Rangoon, near. See Desertion Creek, Elephant Point. Rang Yai, Ko, THAILAND; 7°57'N, 98°26'E; col- lected 8 Feb. 1 9 1 8 by local collectors employed by H. C. Robinson and C. B. Kloss (1919, p. 87); zrc, 1. AUR/FAS/MUL; A:T-53. Rantau; Borneo: Kalimantan, INDONESIA; 2°56'S, 1 15°09'E; collected 10 Mar. 1916 by F. C. E. van der Putten (see Hooijer, 1962b, p. 44); rmnh, 1 (skin only). FAS; CK-28. Rantau Panjang. See Kampong Rantau Panjang. Rantauprapat vicinity; Sumatra, INDONESIA; 164 FIELDIANA: ZOOLOGY ca. 2°06'N, 99°50'E; reported present Nov. 1 97 1- Jan. 1 973 by local residents (Crockett & Wilson, 1980, p. 156). FAS;B:S-35. Ranun River. See Renun, Lae. Ratburi district. See "Siam." Rawa Danau. See Danau, Rawa. Rawas; Sumatra, INDONESIA; not located, 5°38'- 5°57'S, 95°12'-106°05'E; collected in 1906 by W. Volz; zmb, 1 (skull only). FAS; not mapped. Rayoh; Borneo, MALAYSIA: SAB AH; 5°15'N, 1 15°50'E; collected 24 Jun. 1928 by G. Nunong (see Davis, 1962, pp. 10, 127; Weitzel et al., 1988, p. 143); zrc, 1. FAS; C:Sab-2. Rayong. See Wang Kaew. Redang, Pulau, MALAYSIA: WEST MALAY- SIA; 5°45'-5°49'N, 102°59'-103°03'E; collected 2-5 Sep. 1910 by C. B. Kloss (1911a, p. 177); zrc, 2. FAS; B:SCS-1. Renau. See Ranau. Rengsam, Tanjung; Pulau Bangka, INDONESIA; 2°07'S, 105°35'E; collected 21 May 1904 by W. L. Abbott (in Lyon, 1 907, p. 607); usnm, 1 . FAS; B:SCS-18. Renun, Lae; Sumatra, INDONESIA; ca. 3°05'N, 97°55'E; observed May-Dec. 1971 by J. R. MacKinnon (1973, p. 240). FAS; B:S-15. Reo; Pulau Flores, Lesser Sunda Islands, INDO- NESIA; 8°19'S, 120°30'E; observed 21 Nov. 1888-9 Jan. 1889 by M. Weber (1890a, p. viii; 1890b, p. 102). Observed Apr.-May 1930 by Jhr. W. C. van Heurn (1932, p. 65). FAS; C:LS- 17. Riabu, Pulo. See Airabu, Pulau. Riam, Kotawaringin district, 300 m; Borneo: Ka- limantan, INDONESIA; 1°50'S, 111°54'E; col- lected 29 Oct-3 Nov. 1 935 by J. J. Menden; amnh, 4 (including 1 skin only). Collected 31 [Oct.] 1935 by J. J. Menden; mzb, 1. FAS; C:K-8. Rinca, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°37'-8°48'S, 119°37'-119°47'E; observed 26 May-6 Jul. 1953 by A. Hoogerwerf (1954, p. 225; 1955, p. 26). Observed 18 Apr.-14 Jul. 1956 by P. Pfeffer (1959, p. 231). FAS; C:LS- 13. Rinjani, Gunung, 1 700 m; Pulau Lombok, Lesser Sunda Islands; INDONESIA; ca. 8°24'S, 1 16°28'E; observed 23 Sep.-31 Oct. 1987 by D. Kitchener, Boeadi, L. Charlton, and Mahara- datunkamsi (1990, p. 98). FAS; CLS-7. Rintja, Pulau. See Rinca, Pulau. Rite; Pulau Sumbawa, Lesser Sunda Islands, IN- DONESIA; ca. 8°33'S, 1 1 8°53'E (cf. Raba); blood samples taken Jul. 1981-Jan. 1982 by Y. Ka- wamoto, Tb. M. Ischak, and J. Supriatna (1982a, p. 58). FAS;C:LS-12. Roema Manoeal, south foot of Gunung Kenepai; Borneo: Kalimantan, INDONESIA; 0°40'N, 111°42'E; collected 27 Dec. 1893 by J. Biitti- kofer (1897, p. 12); rmnh, 2. FAS; C:K-5. Roi-ed. See Wat Koo Pra Kona. Roi-et. See Wat Koo Pra Kona. Rokan-kanan, Sungai; Sumatra, INDONESIA; 1°23'N, 100°56'E; collected ca. 1907 by M. Moszkowski (1909, map following p. 328); zmb, 3. FAS; B:S-38. Rokankunan. See Rokan-kanan, Sungai. Rompin, Sungai. See Tanjong Panjair, Sungai Rompin. Rondong, Nusa, Komodo area, Lesser Sunda Is- lands, INDONESIA; not located, ca. 8°30'S, 1 19°30'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). Not mapped. Ru, Pulo, BURMA; 9°56'-9°57'N, 98°32'-98° 34'E; collected 13 Nov. 1913 by G. C. Shortridge (in Wroughton, 1915, p. 696); bnhs, 1. AUR/FAS/ MUL; A:Bu-23. Rugading, near Bongkabong, near sea level; Bor- neo, MALAYSIA: SABAH; ca. 5°59'N, 1 16°04'E; collected 22 Aug. 1937 by J. A. Gris- wold, Jr. (see Coolidge, 1940, p. 123); mcz, 1. FAS; C:Sab-4. Rukumodia, Naf River; BANGLADESH; ca. 21°05'N,92°12'E; observed Sep. 1982-Feb. 1983 by M. A. R. Khan and M. A. Wahab (1983, p. 104; cf. M. A. R. Khan, 1981, p. 13; 1985, p. 30). AUR; A:Ba-l. Rungan, Sungai, right bank, 50 km above mouth; Borneo: Kalimantan, INDONESIA; ca. 1°47'S, 1 13°42'E; observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; CK-22. Rungan, Sungai, right bank, above mouth; Borneo: Kalimantan, INDONESIA; ca. 2°09'S, 1 13°53'E; 2 troops observed Jul. 1984-May 1986 by K. M. Burton (see Chivers & Burton, [1991], p. 140). FAS; CK-23. Rupat, Pulau, INDONESIA; 1°41'-2°08'N, 101°23'-101°47'E; reported present 24 Feb.-3 Apr. 1906 by W. L. Abbott (in Lyon, 1908, p. 624). FAS; B:SM-5. Rupat Tinggi. See Kapos Tinggi. Rutland Island, Andaman Islands, INDIA; 11*21'- 1 1°31'N, 92°35'-92°42'E; primates reported ab- sent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, [1928], p. 802; Chaturvedi, 1980, p. 134). A:g. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 165 Sababi, Pulau, Lesser Sunda Islands, INDONE- SIA; 8o21'-8022'S, 120°01'-120°02'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Sabah; Borneo, MALAYSIA: SAB AH; 4°06'- 7°02'N, 1 15°26'-1 19°17'E; collected 24 Aug.-4 Oct. 1960 by R. E. Kuntz (1969, p. 193); usnm, 4. FAS; not mapped. Sabajor. See Sabajor Besar, Pulau. Sabajor Besar, Pulau, Lesser Sunda Islands, IN- DONESIA; 8°29'-8031'S, 119°43'-119044'E; monkeys reported absent 1 969-1 973 by W. Auf- fenberg (1981, p. 40). C:d. Sabalan. See Sebolon Besar, Pulau. Sabor; Pulau Banggi, MALAYSIA: SABAH; 7°14'N, 117°14'E; collected 19 Jun. 1991 by Shukor Md. Nor (pers. comm., 17 Jul. 1991); fmnh, 1. FAS; C:Sab-10. Saigon. See Ho Chi Minh City. Saint Matthew's Island. See Zadetkyi Kyun. Sakerat. See Ban Sakaerat. Salak, Gunung, 2000 m; Java, INDONESIA; 6°42'S, 106°44'E; collected Nov. 1931 and date unknown by H. J. V. Sody; rmnh, 3 (including 2 skulls only). FAS; B: J- 17. Salikan, Mt. See Selikan, Bukit. Sama, Sungei. See Ambawang, Sungai, near Pon- tianak. Samarinda. See Karangmumus, Sungai, near Sa- marinda, and Mahakam, Sungai, north bank, above Samarinda. Samasama; Pulau Nias, INDONESIA; 0°58'N, 97°54'E; collected 2 1 Feb. 1 905 by W. L. Abbott (field catalog; cf. Lyon, 1916, p. 458); usnm, 1. FAS; B:IO-9. Samui, Ko, center of island, 300 ft (= 90 m), THAILAND; ca. 9°30'N, 100°00'E; collected 15 May 1913 by H. C. Robinson (1915, p. 129; cf. P. H. Napier, 1981, p. 17) and E. Seimund; bm(nh), l.FAS; A:T-49. Samuk, Khao; THAILAND; 13°19'N, 100°54'E; reported present in 1981 by W. Y. Brockelman (1981, p. 13). Observed 10-16 Nov. 1990 by N. Aggimarangsee(1992, pp. 1 10, 129; pers. comm., Oct. 1993). FAS; A:T-31. Samunsam Wildlife Sanctuary; Borneo, MALAY- SIA: SARAWAK; ca. 2°00'N, 109°38'E; re- ported present in 1980 by S. Joines (1981, p. 9). Observed before 1987 by G. Hohmann and W. P. Peter (1986, pp. 89, 92). FAS; C:Sar-l. Samut Songkhram vicinity; THAILAND; ca. 13°24'N, 100°00'E; observed Mar. 1909 by K. G. Gairdner ( 1 9 1 4, p. 28). Subspecies uncertain; A:T-37. Samut Song Kram. See Samut Songkhram. Sandakan, 8 mi (= 13 km) W; Borneo, MALAY- SIA: SABAH; ca. 5°50'N, 118°00'E; collected 29 Aug. 1 929 by F. C. Wonder (see Davis, 1 962, p. 10); fmnh, 1 (skull only). FAS; C:Sab-17. Sandaran Agong, Kerinci region, 2450 ft (= 750 m); Sumatra, INDONESIA; 2°07'S, 101°32'E; collected 6 Jun. 1914 by H. C. Robinson and C. B. Kloss (1918, p. 6; in Pendlebury, 1936, p. 27);zrc, l.FAS;B:S-50. Sangeang, Pulau, Lesser Sunda Islands, INDONESflA; 8°08'-8°16'S, 1 19°00'-1 19°07'E; monkeys reported absent 1 969-1 973 by W. Auf- fenberg (1981, p. 40). C:c. Sangeh; Pulau Bali, Lesser Sunda Islands, IN- DONESIA; 8°28'S, 115°14'E; observed Sep. 1969-Jan. 1971 by W. Angst (1975, p. 326). Observed 21 Aug. 1973 by J. Fooden. Observed in 1980 by A. Bakar, M. Amir, and Marshal (1981, p. 11). Observed Jul.-Sep. 1993 by M. F. Small (1994, p. 8). FAS; CLS-3. Sanggul, Bukit, Propinsi Bengkulu, 500 m; Su- matra, INDONESIA; 3°50'S, 102°37'N; col- lected 15-19 Aug. 1936 by J. J. Menden; amnh, 5;mzb, 3. FAS;B:S-71. Sanglang- besar, Pulau, Kepulauan Riau, INDO- NESIA; 0°36'-0°38'N, 103°41'-103°42'E; mon- keys reported absent 10-1 1 Jul. 1903 by W. L. Abbott (see Miller, 1906c, p. 280). Not mapped. Sanglar. See Sanglang-besar, Pulau. Sano, Wai, 650 m; Pulau Flores, Lesser Sunda Islands, INDONESIA; 8°42'S, 120°00'E; col- lected 18 Nov. 1929 by Denin; mzb, 1. FAS; QLS-14. Sapagaya Forest Reserve, near sea level; Borneo, MALAYSIA: SABAH; 5°37'N, 118°04'E; col- lected 9 Aug. 1950 by D. D. Davis (1962, p. 1 1); fmnh, 2. FAS; C:Sab-18. Sapeh. See Ntori, Sape district. Sap Khao, 280 m; THAILAND; 15°35'N, 99°18'E; observed 29 Nov. 1973 and 25 Jan. 1975 by A. A. Eudey (1979, pp. 90, 198). Subspecies un- certain; A:T-19. Sarawak; Borneo, MALAYSIA: SARAWAK; 0°52'-4°58'N, 109°33'-115°38'E; collected in 1879, probably by A. H. Everett; bm(nh), Sub- department of Anthropology, Department of Palaeontology, 3 (1 maxillary fragment, 2 iso- lated molars; not seen, data from P. H. Napier, 1981, p. 20). FAS; not mapped. Sarawak, Sungai, mouth; Borneo, MALAYSIA: SARAWAK; ca. 1°38'N, 110°27'E; observed 1897-1912 by R. W. C. Shelford (1916, p. 10). FAS; C:Sar-4. 166 FIELDIANA: ZOOLOGY Saribas. See Pelandok, Sungai, Paku, Saribas. Sarn Pra Karn, Lopburi; THAILAND; 14°48'N, 100°37'E; observed 24 Feb. and 18 Apr. 1967 by J. Fooden (1971, p. 16). Observed before 1976 by F. C. Cadigan and Lim Boo Liat (1975, p. 86). Observed 22 Jun.-9 Jul. 1989 by N. Ag- gimarangsee (1992, pp. 109, 124; pers. comm., Oct. 1993). Subspecies uncertain; A:T-7. Sarongen; Java, INDONESIA; ca. 6°27'S, 106°00'E; collected 13 Nov. 1909 by O. Bryant (see field catalog in usnm); mcz, 1. FAS; B:J-12. Sarongge. See Sarongen. Sebang. See Sebangka, Pulau. Sebangka, Pulau, INDONESIA; 0°01'-0°15'N, 104°29'-104°43'E; reported present 26-31 Jul. 1903 by W. L. Abbott (see Miller, 1906c, p. 284). FAS; B:SCS-12. Sebatik, Pulau; MALAYSIA: SAB AH; 4° 10'- 4°17'N, 117°38'-117°53'E; collected 4 Jan. 1910 by R. C. Andrews (1911, p. 22; 1943, p. 69); amnh, 1 (skin only); usnm, 2. FAS; C:Sab-12. Sebattick Island. See Sebatik, Pulau. Sebattik Island. See Sebatik, Pulau. Sebesi, Pulau, INDONESIA; 5°55'-5°58'S, 105°28'-105°32'E; primates reported absent Sep. 1920-Jan. 1922) by K. W. Dammerman (1948, p. 68). B:l. Sebesy Island. See Sebesi, Pulau. Sebolon Besar, Pulau, Lesser Sunda Isllands, IN- DONESIA; 8°24'S, 1 19°49'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Sebuku, Pulau, INDONESIA; 3°22'-3°38'S, 116019'-116°27'E; reported present 31 Dec. 1907-5 Jan. 1908 by W. L. Abbott (in Lyon, 1911, p. 62). FAS;C:K-52. Sechawa. See Bakung, Pulau. Sedagong, 900 m; Pulau Tioman, MALAYSIA: WEST MALAYSIA; ca. 2°48'N, 104°1 l'E; col- lected 14-20 May 1927 by N. Smedley: zrc, 2. FAS; B:SCS-3. Segaliud River. See Lungmanis Station, Sungai Se- galiud. Segama, Sungai; Borneo, MALAYSIA: SABAH; 4°43'-5°25'N, 1 17°38'-1 18°47'E; collected 22 Aug. 1907 by Dr. Pagel; zmb, 1 (skull only); FAS; C:Sab-21. Segobang, Sungai; Borneo, MALAYSIA: SARA- WAK; 2 rivers by this name in Sarawak (mouth of one at 1°23'N, 110°E; mouth of other at 4°47'N, 1 14°58'E); collected 18 Sep. 1892 by E. Bartlett; smk, 1 (skin only, skull inside). FAS; not mapped. Sekol, Sungai, 100 m; MALAYSIA: WEST MA- LAYSIA; 1°54'N, 103°39'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-29. Sekonyer Kanan River (?= Sikunir, Sungai), Tan- jung Puting National Park; Borneo: Kaliman- tan, INDONESIA; ca. 2°50'S, 111°47'E; ob- served 1977-1983 by B. M. F. Galdikas and C. P. Yeager (1984, p. 50). FAS; CK-9. Selai, Sungai; MALAYSIA: WEST MALAYSIA; 2°16'N, 103°25'E; observed Jul. 1978-Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-28. Selama. See Ulu Ijok. Selatbliat; Pulau Kundur, Kepulauan Riau, IN- DONESIA; ca. 0°53'N, 103°22'E; collected 18- 20 Aug. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, pp. 105, 107) and E. Sei- mund; bm(nh), 2. FAS; B:SCS-17. Selayar, Pulau, INDONESIA; 0°17'-0°20'S, 104°23'-104°29'E; reported present 2-6 Aug. 1903 by W. L. Abbott (see Miller, 1906c, p. 280). FAS; B:SCS-14. Selikan, Bukit; Borneo, MALAYSIA: SARA- WAK; 3°31'N, 1 14°04'E; collected July 1895 by E. Hose and C. Hose; usnm, 1. FAS; C:Sar-14. Selindang, Pulo. See Selintang, Pulau. Selintang, Pulau, INDONESIA; 0°57'-0°58'N, 107°28'-107°29'E; observed 3-7 Aug. 1899 by W. L. Abbott (in Miller, 1900, p. 243; cf. Kloss, 1903b, p. 60). FAS; B:SCS-26. Semao Island. See Oeassa. Semarang; Java, INDONESIA; 6°58'S, 1 10°25'E; collected 1907-1908 by C. Bruegel and/or U. Schaule; zsbs, 1 (skin only). FAS; C:J-26. Sembakung, Sungai; Borneo: Kalimantan, IN- DONESIA; ca. 3°50'N, 1 17°00'E; collected Jul. 1912 by Mohari (see Sody, 1949, p. 180); mzb, 2. FAS; QK-37. Sembaloen. See Pusuk forest. Sembawang, Sungai; Singapore Island, SINGA- PORE; 1°27'N, 103°50'E; collected 21 May 1922 by F. N. Chasen; zrc, 1 (skin only). FAS; B:SCS-7. Sember, P. See "P. Sember." Sembilan Kepulauan, MALAYSIA: WEST MA- LAYSIA; ca. 4°02'N, 100°33'E; primates re- ported absent in 1953 by J. L. Harrison and J. R. Hendrickson (1963, p. 548). B:c. Sembrong River. See Endau, Sungai, vicinity. Semitau, Sungai Kapuas; Borneo: Kalimantan, INDONESIA; 0°33'N, 111°58'E; collected 15 Dec. 1893 by J. Biittikofer (1897, p. 11); rmnh, 1. FAS; C:K-6. Semongkat; Pulau Sumbawa, Lesser Sunda Is- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 167 lands, INDONESIA; 8°35'S, 1 17°20'E; collected 31 Oct. 1981 by M. Aimi (pers. comm., 3 Aug. 1990; cf. Kawamoto et al., 1984, p. 132); pri, 1 (skeleton only). Blood samples taken Jul. 1 98 1- Jan. 1982 by Y. Kawamoto, Tb. M. Ischak, and J. Supriatna (1982, p. 58). FAS; C:LS-8. Sendang; Pulau Bali, Lesser Sunda Islands, IN- DONESIA; 8°08'S, 1 14°38'E; collected ca. 1930 by H. J. V. Sody (1933, p. 93); rmnh, 3. FAS; C:LS-1. Sendung; Borneo, country unknown; not located, 7°02'N-4°11'S, 108°51'-119°17'E; collected 24 Apr. 1902 by Dr. Pagel; zmb, 1 (skin only). FAS; not mapped. Sengata; Borneo: Kalimantan, INDONESIA; 0°28'N, 1 17°33'E; observed summer 1971 by J. A. Kurland (1973, p. 250). FAS; CK-46. Sengata, Sungai, 1 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°31'N, 117°22'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 2 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°31'N, 117°23'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 3 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°32'N, 117°24'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; C:K-46. Sengata, Sungai, 5 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°32'N, 117°25'E; 2 troops observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 7 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°33'N, 117°26'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 9 km below Mentoko camp; Bor- neo: Kalimantan, INDONESIA; ca. 0°33'N, 1 17°27'E; 2 troops observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 1 1 km below Mentoko camp; Borneo: Kalimantan, INDONESIA; ca. 0°30'N, 117°28'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, 13 km below Mentoko camp; Borneo: Kalimantan, INDONESIA; ca. 0°30'N, 117°29'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, at Mentoko camp; Borneo: Ka- limantan, INDONESIA; ca. 0°30'N, 117°22'E; observed 1983-1989 by A. Suzuki (1989, p. 32). FAS; CK-46. Sengata, Sungai, below Sengata village; Borneo: Kalimantan, INDONESIA; ca. 0°27'N, 1 17°35'E; 12 troops observed 14 Apr. 1973 by C C Wilson and W. L. Wilson (1975, p. 257). FAS; CK-46. Sengata, Sungai, Kutai Reserve; Borneo: Kali- mantan, INDONESIA; 0°32'N, 117°28'E; ob- served 2-6 Apr. 1973 by C C Wilson and W. L. Wilson ( 1 975, p. 257). Observed 26 Jan. 1 973- 31 Jan. 1974 by N. A. Fittinghoff, Jr., and D. G. Lindburg (1980, p. 185). FAS; CK-46. Sengata, Sungai, right bank, 1 km below mouth of Sungai Nubung, Kutai Reserve; Borneo: Ka- limantan, INDONESIA; ca. 0°32'N, 117°07'E; observed 13 Oct. 1985 by A. Suzuki (1986, p. 16). FAS; CK-47. Sengatta River. See Sengata, Sungai. Senimba Bay; Pulau Batam, Kepulauan Riau, IN- DONESIA; 1°05'N, 103°56'E; observed 22 Mar. 1906 by C. B. Kloss (1908, p. 65). FAS; B:SCS- 10. Senoeang, 500 m; Borneo: Kalimantan, northeast, INDONESIA; ca. TOO'N, 11 OWE; collected 29 Aug. 1937 by J. J. Menden; amnh, 1 (skin only; male skin mismatched with female skull); FAS, CK-3. Sentosa Island, SINGAPORE; 1°14'-1°16'N, 103°48'-103°51'E; observed before 1901 by S. S. Rower (1900, p. 316). FAS; B:SCS-7. Sepaku, Sungai, ca. 350 m NE of camp, 15-50 m; Borneo: Kalimantan, INDONESIA; ca. 0°53'S, 116°43'E; observed 11-13 Mar. 1973 by C C Wilson and W. L. Wilson (1975, pp. 248, 254). FAS; CK-50. Sepilok Forest Reserve. See Labuk Road, Sepilok Forest Reserve. Seraja, Pulau. See Seraya Besar, Pulau. Serang Daja. See Serangjaya-hilir. Serang Djaja. See Serangjaya-hilir. Serangjaya-hilir; Sumatra, east coast, INDONE- SIA; 4°15'N, 98°12'E; collected Oct.-Nov. 1931 by H. J. V. Sody; rmnh, 2 (skulls only). FAS; B:S-19. Serapit Tandjung; Sumatra, INDONESIA; ca. 3°33'N, 98°20'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-20. Serasan, Pulau, INDONESIA; 2°28'-2°33'N, 108°58'-109°07'E; observed Sep. 1893 by A. Everett (in Thomas & Hartert, 1894, p. 654). Collected 8 Jun. 1 900 by W. L. Abbott (see Mil- ler, 1901, p. Ill); usnm, 1. Collected 27 Aug. 1931 by P. M. de Fontaine (see Chasen, 1935a, p. 5); zrc, 1. FAS; B:SCS-36. Seraya Besar, Pulau, Lesser Sunda Islands, IN- 168 FIELDIANA: ZOOLOGY DONESIA; 8°22'-8°24'S, 1 19°50'-1 19°52'E; re- ported present 1969-1973 by W. Auffenberg (1981, p. 40). FAS; QLS-13. Serdang district; Sumatra, INDONESIA; ca. 3°30'N, 99°00'E; collected 2 Feb. 1905 by H. Diirk (see Schneider, 1 905, map 1); zsbs, 1 . FAS; B:S-22. Sertung, Pulau, INDONESIA; 6°03'-6°07'S, 105°22'-105°24'E; primates reported absent 1883-1934 by K. W. Dammerman (1948, p. 50); volcanic explosion in 1883. B:k. Serutu, Pulau, INDONESIA; 1°42'-1°44'S, 108°41'-108°48'E; collected 25 Mar. 1931 by L. Coomans de Ruiter and Madzoed (see Chasen, 1935b, p. 2); mzb, 1. FAS; B:SCS-21. Sewela; Pulau Lombok, Lesser Sunda Islands, IN- DONESIA; 8°32'S, 116°35'E; collected 22-26 Mar. 1927 by Sunda-Expedition Rensch (see B. Rensch, 1930, p. 12; Mertens, 1930, p. 127; I. Rensch, 1934, p. 228); nms, 2. FAS; C:LS-7. S'gobang. See Segobang, Sungai. Shanghai; CHINA; 31°14'N, 121°28'E; captive (introduced) obtained Jul. 1 858 by J. Zelebor or von Frauenfeld (Zelebor, [1869], p. 8; cf. Fit- zinger, 1861, p. 389). Subspecies uncertain; not mapped. Siaba. See Siaba Besar, Pulau. Siaba, Teluk; Pulau Nias, INDONESIA; 1°31'N, 97°24'E; collected 16-20 Mar. 1903 by W. L. Abbott (see Lyon, 1916, p. 458); usnm, 7 (in- cluding 3 skulls only, 2 with external measure- ments recorded on skull tags); FAS; B:IO-10. Siaba Besar, Pulau, Lesser Sunda Islands, IN- DONESIA; 8°32'-8°33'S, 119°41'-119042'E; monkeys reported absent 1 969-1 973 by W. Auf- fenberg (1981, p. 40). C:d. Siaboh. See Sibaboh, Lugu. Siak, Sungai, ca. 12 km above mouth; Sumatra, INDONESIA; ca. 1°14'N, 102°10'E; observed Nov. 197 1-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 15 km below Pekanbaru; Su- matra, INDONESIA; ca. 0°35'N, 101°34'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 24 km above mouth; Sumatra, INDONESIA; ca. 1°07'N, 102°10'E; observed Nov. 1 97 1-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 30 km below Pekanbaru; Su- matra, INDONESIA; ca. 0°42'N, 101°39'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 36 km above mouth; Sumatra, INDONESIA; ca. 1°01'N, 102°07'E; observed Nov. 1 97 1-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 45 km below Pekanbaru; Su- matra, INDONESIA; ca 0°47'N, 101°44'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 48 km above mouth; Sumatra, INDONESIA; ca. 0°55'N, 102°05'E; observed Nov. 197 1-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 60 km above mouth; Sumatra, INDONESIA; ca. 0°48'N, 102°05'E; observed Nov. 197 1-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 60 km below Pekanbaru; Su- matra, INDONESIA; ca 0°45'N, 101°51'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, ca. 75 km below Pekanbaru; Su- matra, INDONESIA; ca. 0°48'N, 101°58'E; ob- served Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak, Sungai, near mouth; Sumatra, INDONE- SIA; ca. 1°20'N, 1 02° 10'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-42. Siak, Sungai, near Pekanbaru; Sumatra, INDO- NESIA; ca. 0°32'N, 101°27'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-42. Siak Copatta; Sumatra, INDONESIA; ca. 1°00'N, 101°00'E; collected ca. 1907 by M. Moszkowski (1909, map following p. 192); zmb, 1 (skull only). FAS; B:S-40. Siam. See THAILAND. "Siam"; THAILAND; "13°45'N, 99°25'E"; col- lected 18 Jun. 1913 and date unknown by K. G. Gairdner (1914, pp. 34-36; cf. P. H. Napier, 1 98 1 , p. 20, who cited this locality as "E. Siam"); bm(nh), 2 (skulls only). Subspecies uncertain; A:T-27. Siantan, Pulau, INDONESIA; 3°04'-3°13'N, 106°12'-106°17'E; collected 8 Sep. 1899 by W. L. Abbott (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 69; Oberholser, 1917, p. 2); usnm, 1. Collected 14 Sep. and 3 Oct. 1925 by F. N. Chasen (see Chasen & Kloss, 1 928a, p. 43); zrc, 2. FAS; B:SCS-30. Sibabo. See Sibaboh, Lugu. Sibaboh, Lugu; Pulau Simeulue, INDONESIA; 2°43'N, 96°06'E; collected 17 Dec. 1901 and 26 Oct. 1902 by W. L. Abbott (field catalog; cf. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 169 Miller, 1903a, p. 437; Lyon, 1916, p. 457);usnm, 4. FUS, B:IO-2. Sibau, Sungai. See Poelau (?= Kuda), Sungai Si- bau. Siberut, Pulau, INDONESIA; 0°55'-l°48'S, 98°35'-99°18'E; M. fascicularis reported absent before 1928 by C. B. Kloss ([1928], p. 802). B:f. Sibuga Besar, Sungai; Borneo, MALAYSIA: SA- BAH; ca. 5°56'N, 118°03'E; collected 21 Aug. 1929 by F. C. Wonder (field catalog; cf. Davis, 1962, p. 10); fmnh, 1. FAS; C:Sab-17. Sibugal River. See Sibuga Besar, Sungai. Si Chang, Ko, THAILAND; 13°07'-13°1 l'N, 100°48'-100°50'E; primates reported as appar- ently absent Jan. 1915 by C. B. Kloss (1915a, p. 157; 1915b, p. 221). M. fascicularis reported present 15 Nov. 1990 by local residents (Aggi- marangsee, 1992, pp. 1 10, 130). Pending further information, Aggimarangsee's record is accept- ed as valid. Subspecies uncertain; A:T-3 1 . Sidjoendjoeng. See Sijunjung, Padang highlands. Siemreab, 1 6 km S, 1 50 ft (= 45 m); CAMBODIA; 13°22'N, 103°51'E; collected 16 Dec. 1927 by J. Delacour (1929, p. 195) and W. P. Lowe; mnhn, 1. FAS; A:C-1. Siem-Reap. See Siemreab. Sijunjung, Padang highlands; Sumatra, INDO- NESIA; 0°42'S, 100°58'E; collected 21 Jul. 1877 by J. F. Snelleman (1887, p. 10; cf. Hooijer, 1962b, p. 44); rmnh, 1 (skull only). FAS; B:S- 53. Sika; Pulau Flores, Lesser Sunda Islands, INDO- NESIA; 8°45'S, 122°12'E; observed 21 Nov. 1888-9 Jan. 1889 by M. Weber (1890a, p. ix; 1890b, p. 102). FAS; GLS-19. Sikka. See Sika. Sikundur; Sumatra, INDONESIA; ca. 4°00'N, 98°00'E; observed May-Dec. 1971 by J. R. MacKinnon (1973, p. 240). FAS; B:S-19. Simalur Island. See Simeulue, Pulau. Simeulue, Pulau, INDONESIA; 2°19'-2°55'N, 95°41'-96°30'E; collected 31 Jul. 1924 and 23 Dec. 1 925 by unknown collectors; rmnh, 2 (skins only). Reported present before 1981 by J. T. Marshall (Crockett & Wilson, 1980, p. 156). Blood samples taken from captives 1984-1989 by W. Scheffrahn, J. R. de Ruiter, and J. A. R. A. M van Hooff(1994, p. 135). FUS; B:IO-2, IO-3. Similajau National Park; Borneo, MALAYSIA: SARAWAK; ca. 3°38'N, 1 13°20'E; reported as •probably present before 1983 by G. Davies ([1983], p. 148). FAS; C:Sar-15. Simpang; Pulau Bangka, INDONESIA; 1°54'S, 105°26'E; collected in 1905 by B. Hagen; zsbs, 1 (skin only). FAS; CSCS-18. Simujan River. See Simunjan, Sungai. Simunjan, Sungai; Borneo, MALAYSIA: SARA- WAK; ca. 1°17'N, 110°49'E; observed Aug. 1855 by A. R. Wallace (1869, pp. 69, 82). Collected 1876-1878 by W. T. Hornaday (1910, p. 358); museum unknown, 2 (not seen). FAS; C:Sar-7. Simunjon River. See Simunjan, Sungai. Sinabang; Pulau Simeulue, INDONESIA; 2°29'N, 96°23'E; collected 3-14 Feb. 1913 by E. Jacob- son (1913, p. 356; 1917, p. 276) and W. C. V. Heurn; rmnh, 2 (skins only). Collected Jul. 1913 by E. Jacobson (1913, p. 356; 1917, p. 276); rmnh, 2 (skins only). FUS; B:IO-3. Singapore Botanical Gardens; Singapore Island, SINGAPORE; 1°18'N, 103°49'E; collected 2 Apr. 1893 by unknown collector; zrc, 1 (skin only, skull inside). Observed before 1 90 1 by S. S. Flower ( 1 900, p. 3 1 6). Collected 26 Jul. 1912 by unknown collector; zrc, 1 (skin only). Col- lected 23 Mar. 1925 by museum collector; zrc, 1. Collected Apr. 1925 by unknown collector; zrc, 1 (skull only). Collected 11 Feb. 1944 by Oshita; zrc, 1. Observed Jul.-Sep. 1960 by Y. Furuya (1961-1962a, p. 76; 1965, p. 289). Ob- served 29 Jul. 1973 by J. Fooden. FAS; B:SCS-7. Singapore Island, northwestern part, mangroves, SINGAPORE; ca. 1°25'N, 103°44'E; reported present before 1974 by S. H. Chuang (1973, p. 3). FAS; B:SCS-7. Singapore Island, western part, mangroves, SIN- GAPORE; ca. 1°20'N, 103°38'E; reported pres- ent before 1974 by S. H. Chuang (1973, p. 3). FAS; B:SCS-7. Singapore Island, SINGAPORE; 1°16'-1°28'N, 103°38'-104°00'E; captive acquired Apr. 1858 by J. Zelabor ([1869], p. 8; cf. Fitzinger, 1861, p. 389) or von Frauenfeld. Collected [ 1 870-1 87 1 ] by A. B. Meyer (see Steenis-Kruseman, 1950, p. 358), zmb, 1. Collected 22 Mar. 1903 by un- known collector; zrc, 1. FAS; B:SCS-7. Singkarak, Lake. See Pajo, Danau Singkarak, Bal- ipor district. Singkel vicinity; Sumatra, INDONESIA; ca. 2°1 7'N, 97°49'E; blood samples taken Nov.-Dec. 1986 by J. R. de Ruiter (1993, p. 91). FAS; B:S- 17. Singkil. See Singkel vicinity. Singkil, Gunung; Java, west, INDONESIA; not located, 5°52'-7°48'S, 105°12'-108°52'E; col- lected 19-20 Jul. 1909 by H. W. van der Weele; rmnh, 3. FAS; B:J-8. Sintang; Borneo: Kalimantan, INDONESIA; 170 FIELDIANA: ZOOLOGY OWN, 1 1 1°30'E; collected 14 Oct.^l Nov. 1897 by A. Harrison, Jr., and H. M. Hiller; ansp, 5 (including 2 skins only). FAS; C:K-7. Sinubing; Pulau Natuna Besar, INDONESIA; not precisely located, 3°37'-4°13'N, 107°58'- 108°25'E; collected Jul. 1 894 by C. Hose; bm(nh), 1 (skin only, skull inside). FAS; B:SCS-32, SCS- 33. Siolak Daras. See Siulakderas, Kerinci region. Sipora Island. See Sipura, Pulau. Sipura, Pulau, INDONESIA; 2°02'-2°24'S, 99°32'- 99°52'E; M. fascicularis reported absent before 1928 by C. B. Kloss ([1928], p. 802). B:g. Si Racha vicinity; THAILAND; ca. 13°10'N, 1 00°56'E; observed Apr.-May 1 9 1 2 by N. Gyld- enstolpe (1914, p. 3). FAS; A:T-31. Sirhassen Island. See Serasan, Pulau. Sittwe harbor. See Myengun Kyun. Siulakderas, Kerinci region, 3000 ft (= 900 m), Sumatra, INDONESIA; 1°55'S, 101°18'E; col- lected 27 Mar. 1914 by H. C. Robinson and C. B. Kloss (1918, p. 6; in Pendlebury, 1936, p. 9); bm(nh), 1. FAS; B:S-49. Slamet, Mt. See Baturaden; Kaligoea. Smitau. See Semitau, Sungai Kapuas. Soekadono. See Sukadana. Soekaranda; Sumatra, INDONESIA; not located, 5°38'N-5°57'S, 95°12'-106°05'E; collected Aug. 1894 by Ude; zmb, 1 (skull only). FAS; not mapped. Soengei Slan. See Sungaiselan. Soengi Han. See Sungaiselan. Soliga; Pulau Nias, INDONESIA; ca. 1°02'N, 97°33'E; collected 25 Jul. 1937 by B. Lawrence (see Schauensee & Ripley, 1 940a, map following p. 368); mcz, 2. FAS; B:IO-7. Solombo, Pulau. See Masalembu Besar, Pulau. Solor, Pulau, Lesser Sunda Islands, INDONESIA; 8°26'-8°37'S, 122°53'-123°11'E; reported pres- ent before 1937 by R. Mertens (1936, p. 319). FAS; CLS-20. Songkhla. See Khao Noi/Khao Tangkuan. Sontra Peak, 0-300 m; VIETNAM; ca. 16°07'N, 108°18'E; observed Jun.-Aug. 1974 by L. K. Lippold (1977, p. 521). FAS; A:V-1. Sontra Peak, 3.9 km W, 0.3 km S, 150-600 m; VIETNAM; ca. 16°07'N, 108°18'E; collected 14 Sep. 1967 by P. F. D. Van Peenen (see Van Peenenetal., 1969, pp. 100,290; 1971, p. 134); usnm, 1. FAS; A:V-1. South Andaman Island, INDIA; 1 1°29'-12°15'N, 92°31'-92°48'E; primates reported absent be- fore 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, [ 1 928], p. 802; Chaturvedi, 1 980, p. 1 34). A:e. South Banyuwangi Nature Park; Java, INDO- NESIA; ca. 8°37'S, 114°28'E; observed 17-28 Sep. 1971 by A. Hoogerwerf (1974, p. 24). FAS; C:J-43. South Island. See Pelapis Tengah, Pulau. South Pagi Island. See Pagai Selatan, Pulau. South Sumatra Province. See Sumatera Selatan, Propinsi. Sriracha. See Si Racha vicinity. St. Barbe Island. See Pejantan, Pulau. Straits of Malacca. See Langkawi, Pulau. Suaka Margasatwa Padang Sugihan; Sumatra, IN- DONESIA; not located; observed Sep. 1988 by M. Bismark (1992, p. 1 1). FAS; not mapped. Subi-kecil, Pulau, INDONESIA; 3°01'-3°03'N, 108°51'-108°53'E; collected 5 Aug. 1931 by P. M. de Fontaine (see Chasen, 1935a, p. 5); zrc, 3. FAS; B:SCS-35. Sugi, Pulau, Kepulauan Riau, INDONESIA; 0°45'- 0°53'N, 103°43'-103°5rE; collected 24 Aug. 1902 by W. L. Abbott (see Miller, 1906c, p. 281); usnm, 1. FAS; B:SCS-16. Suka Bandjar, northeast; Sumatra, INDONESIA; ca. 4°55'S, 104°02'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1 980, p. 156). FAS;B:S-77. Suka Bandjar, southeast; Sumatra, INDONESIA; ca. 5°00'S, 103°57'E; observed Nov. 1971-Jan. 1 973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-77. Sukadana, 100 m; Sumatra, INDONESIA; 5°05'S, 105°33'E; 29-30 Jul. 1934; collected by J. J. Menden; amnh, 4. FAS; B:S-83. Sulawesi, Pulau, INDONESIA; 1°45'N-5°43'S, 1 18°45'-125°15'E; introduced, reported present before 1841 by S. Miiller ([1840], p. 17). Sub- species uncertain; not mapped. Sullivan's Island. See Lanbi Kyun. Sumatera Selatan, Propinsi (province); Sumatra, INDONESIA; 1°39'-4°53'S, 101°57'-106°1 l'E; blood samples taken Jan.-Nov. 1 979 by Y. Ka- wamoto and Tb. M. Ischak(1981,p. 238). FAS; not mapped. Sumatra, [east-central], INDONESIA; ca. 1°00'N, 101°00'E; collected ca. 1907 by M. Moszkowski (1909, maps following pp. 192, 328); zmb, 4 (including 3 skulls only). FAS; B:S-40. Sumatra, east coast, INDONESIA; 5°38'N-5°57'S, 95°12'-106°05'E; collected date unknown by H. Diirk; zsbs, 1 5 (including 4 skins only, 4 skulls only). FAS; not mapped. Sumatra, INDONESIA; 5°38'N-5°57'S, 95°12'- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 171 106°05'E; collected 7 Sep. 1 82 1 by A. Duvaucel; mnhn, 1 (skin only). Collected in 1889 by Dr. Moesch; zmuz, 1 (skin only, skull inside). Col- lected 1896-1897 by unknown collector; nms, 2 (skulls only). Collected 10 Feb. and 5 Jun. 1897 by Petersen; zmb, 2 (skulls only). Collected date unknown by Petersen; zmb, 7 (including 4 skins only, 3 skulls only). Collected 1905-1910 by C. Bruegel; zsbs, 52 (including 22 skins only, 1 1 skulls only). Collected date unknown by C. Bruegel; zsbs, 1 (skin only). Collected in 1924 by P. Wirz; NHMBa, 2. Collected 10 Apr. 1926- 18 Aug. 1930 by C. Blazer; rmnh, 4 (including 3 skulls only). Collected 3 Feb. 1938 by W. Groenvelt, rmnh, 1 (skull only). Collected be- fore 1942 by L. Coomans de Ruiter; rmnh, 1 (skull only). Collected 3 May 1943 by unknown collector; usnm, 1 . Collected date unknown by Dr. Porter; zsbs, 1 (skull only). Collected date unknown by unknown collector; nms, 2 (skulls only). FAS; not mapped. Sumatra, south, INDONESIA; 5°38'N-5°57'S, 95°12'-106°05'E; collected date unknown by W. Volz; zmb, 2 (skins only). FAS; not mapped. Sumatra, west coast, INDONESIA; 5°38'N-5°57'S, 95°12'-106°05'E; collected before 1883 by F. von Faber; bm(nh), 1 . FAS; not mapped. Sumbawa, Pulau, Lesser Sunda Islands, INDO- NESIA; 8°05'-9°07'S, 116°42'-119012'E; re- ported present before 1 906 by T. Willink ( 1 905, p. 175). Collected in 1914 by Dr. Pannekoek; NHMBa, 1 (skull only). FAS; C:LS-8, LS-10 through LS- 12. Sumbawang, Sungei. See Sembawang, Sungai. Sunda Islands; INDONESIA; 5°38'N-10°23'S, 95°1 2'-127°l 8'E; collected in 1 899 by P. Sarasin and F. Sarasin; NHMBa, 1 (skull only). Collected date unknown by von Altenstein; zmb, 1 (skin only). FAS; not mapped. Sundarbans. See Khulna district, Sundarbans. Sungai Kayan-Sungai Mentarang Nature Reserve; Borneo: Kalimantan, INDONESIA; ca. 3°00'N, 115°30'E; reported present before 1983 by G. Davies ([1983], p. 148). FAS; GK-36. Sungai Kinabatangau. See Abai. Sungailundang; Sumatra, INDONESIA; 1°05'S, 100°29'E; morphological study Jun.-Nov. 1980 by A. Bakar, M. Amir, and Marshal (1981, p. 11). FAS; B:S-47. Sungoxseldin; Pulau Bangka, INDONESIA; 2°24'S, 105°59'E; collected in 1900 by A. A. W. Hu- brecht([1895], pp. 33, 88; cf Kohlbrugge, 1902, p. 322; Zuckerman, [1933], p. 1062); zluu, un- specified portion of Pulau Bangka collection (see Bangka, Pulau). FAS; B:SCS-19. Sungai Tekam Forestry Concession, 250-350 m; MALAYSIA: WEST MALAYSIA; 4°10'N, 102°40'E; observed 1979-1981 by A. D. Johns (1981, p. 222). FAS; B:WM-12. Sungei Adang; Ko Tarutao, THAILAND; not pre- cisely located, 6°3 1 '-6°44'N, 99°36'-99°42'E; collected 10-11 Mar. 1909 by museum collec- tors (see H. C. Robinson & Kloss, 1910, p. 666); bm(nh), 2. FAS; A:T-59. Sungei Bernam. See Changkat Mentri. Sungei Biru; Pulau Bintan, INDONESIA; ca. 1°12'N, 104°33'E; collected Jun.-Sep. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, pp. 100, 104); bm(nh), 3; zrc, 1 (skull only). FAS; B:SCS-8. Sungei Buloh. See Kampong Sungai Buloh. Sungei Hdang. See Sungei Adang. Sungei Pandang. See Pandan, Sungai. Sungsan, ca. 20 km NNE; Sumatra, INDONESIA; ca. 5°15'S, 105°50'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-85. Sungsan, ca. 50 km N; Sumatra, INDONESIA; ca. 4°58'S, 105°48'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L.Wilson (1980, p. 156). FAS; B:S-84. Suranadi; Pulau Lombok, Lesser Sunda Islands, INDONESIA; 8°33'S, 116°15'E; collected 10 Dec. 1981 by M. Aimi (pers. comm., 3 Aug. 1990; cf. Kawamoto et al., 1984, p. 132); pri, 1 (skeleton only). Morphometric study Dec. 1981 by M. Aimi, A. Bakar, and J. Supriatna (1982, p. 52). Blood samples taken Jul. 1981-Jan. 1982 by Y. Kawamoto, Tb. M. Ischak, and J. Su- priatna (1982, p. 58). FAS; GLS-5. Sur Sdei. See Pang Roloem-Sur Sdei area. Survey Site LB, 5-50 m; Pulau Simeulue, IN- DONESIA; ca. 2°35'N, 96°10'E; 2 troops ob- served Jan. 1982 and 1 1 Mar.-3 Apr. 1984 by J. Sugardjito, C. P. van Schaik, N. A. van Noordwijk, and T. Mitrasetia (1989, p. 198). FUS; B:IO-2. Survey Site LS, 10-120 m; Pulau Simeulue, IN- DONESIA; ca. 2°44'N, 95°54'E; 2 troops ob- served Jan. 1982 and 11 Mar .-3 Apr. 1984 by J. Sugardjito, C. P. van Schaik, N. A. van Noordwijk, and T. Mitrasetia (1989, p. 198). FUS; B:IO-2. Survey Site SEM, 210 m; Pulau Simeulue, IN- DONESIA; ca. 2°45'N, 96°02'E; 5 troops ob- served Jan. 1982 and 1 1 Mar.-3 Apr. 1984 by J. Sugardjito, C. P. van Schaik, N. A. van 172 FIELDIANA: ZOOLOGY Noordwijk, and T. Mitrasetia (1989, p. 198). FUS; B:IO-2. Swela. See Sewela. Syonan. See Singapore Island, SINGAPORE. Tabanan. See Kukuh. Tagoot (?= Tagu), Great Tenasserim River; BUR- MA; 12°15'N, 99°03'E; collected 18 Apr. 1914 by G. C. Shortridge (in Wroughton, 1915, p. 697; cf. Moore & Tate, 1965, p. 332); bnhs, 2. AUR; A:Bu-19. Tahan, Sungai; MALAYSIA: WEST MALAY- SIA; ca. 4°35'N, 102°18'E; observed Jul.-Sep. 1960 by Y. Furuya (1965, p. 287). FAS; B:WM- 11. Tahang River. See Tahan, Sungai. Takengon, ca. 15 km NNW; Sumatra, INDO- NESIA; ca. 4°45'N, 96°47'E; reported present Nov. 1971-Jan. 1973 by local residents (Crock- et & Wilson, 1980, p. 156). FAS; B:S-9. Takengon, ca. 40 km NNW; Sumatra, INDO- NESIA; ca. 4°58'N, 96°42'E; reported present Nov. 1971-Jan. 1973 by local residents (Crock- ett & Wilson, 1980, p. 156). FAS; B:S-8. Takokak Reserve; Java, INDONESIA; ca. 7°03'S, 106°59'E; reported present before 1972 by un- specified informant (iucn, 1971, p. 276). FAS; B:J-8. Tale Sap. See Nang Kham, Ko. Talibang, near Kota Kinabalu, near sea level; Bor- neo, MALAYSIA: SAB AH; ca. 6°H'N, 116°14'E; collected 15-24 Aug. 1937 by J. A. Griswold, Jr. (see Coolidge, 1940, p. 123); mcz, 4 (including 1 skeleton only). FAS; C:Sab-5. Talibong. See Talibang. Talibong, Ko, northwest, THAILAND; ca. 7°17'N, 99°23'E; Collected 3 Jan. 1917 by H. C. Rob- inson (1917, p. 132); zrc, 1. FAS; A:T-56. Talibong, Ko, THAILAND; 7°12'-7017'N, 99°22'- 99°27'E; collected 27 Feb. 1 896 by W. L. Abbott (see Riley, 1938, p. 12); usnm, 1. FAS; A:T-56. Tamandjaija, Bantam region; Java, INDONESIA; 6°52'-7°00'S, 105°12'-106°33'E; collected 15 Jan. 1910 by O. Bryant; mcz, 1. FAS; B:J-12. Taman Negara. See Negara, Taman. Taman Rimba Templer. See Templer Park. Tamansari, 1600 ft (= 490 m); Java, INDONE- SIA; 7°57'S, 1 11°31'E; collected 20 Jan. 1920 by C. B. Kloss (see Weitzel et al., 1988, p. 144); bm(nh), 1; zrc, 1. FAS; C:J-31. Tambelan Besar, Pulau, INDONESIA; 0°57'- 1°02'N, 107°32'-107°36'E; observed 8-15 Aug. 1899 by W. L. Abbott (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 62). FAS; B:SCS-26. Tambelan Islands. See Benua, Pulau; Uwi, Pulau. Tamiang; Sumatra, northeast, INDONESIA; ca. 4°25'N, 98°16'E; collected Nov. 1931 by H. J. V. Sody (see Hooijer, 1962b, p. 45); rmnh, 1 (skull only). FAS; B:S- 18. Tana Bala. See Tanahbala, Pulau. Tanahbala, Pulau, INDONESIA; 0°15'-0°37'S, 98°17'-98°30'E; collected 4 Feb. 1903 by W. L. Abbott (see Lyon, 1916, p. 459); usnm, 2. Re- ported Nov. 1971-Jan. 1973 by local residents (Crockett & Wilson, 1980, p. 156). FAS; B:IO- 13. Tanah Laut. See Pleihari Tanah Laut Game Sanc- tuary. Tanahmasa, Pulau, INDONESIA; 0°01'N-0°23'S, 98°17'-98°34'E; collected 19 Feb. 1903 by W. L. Abbott; usnm, 1 (skull only). FAS; B:IO-12. Tanah Puteh. See Kampong Tanah Puteh. Tandjong. See Tanjung. Tandjung; Sumatra, INDONESIA; ca. 3°17'N, 99°19'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of speci- mens unknown (not seen). FAS; B:S-23. Tandjung Bringin; Sumatra, INDONESIA; ca. 3°47'N, 98°24'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-20. Tandjung Butus; Sumatra, INDONESIA; ca. 3°47'N, 98°24'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-20. Tandjung Laut. See Tandjung. Tangarveng Island, Sungai Mahakam; Borneo: Kalimantan, INDONESIA; ca. 0°24'S, 1 16°58'E; collected 2 1 Jun. 1 9 1 2 by H. C. Raven (see Deig- nan, [1960], p. 267); usnm, 1. FAS; CK-49. Tanjang Sau. See Tanjong Sauh. Tanjang Turut. See Tanjong Turut. Tanjong Batu. See Batu, Tanjung. Tanjong Pamuju. See Pamuja, Tanjung. Tanjong Panjair, Sungai Rompin; MALAYSIA: WEST MALAYSIA; ca. 2°49'N, 103°29'E; col- lected 2 Sep. 1 9 1 9 by museum collector; bm(nh), 1;zrc, 1. FAS; B:WM-22. Tanjong Rengsam. See Rengsam, Tanjung. Tanjong Sauh; Pulau Batam, east coast, Kepu- lauan Riau, INDONESIA; ca. 1°07'N, 104°09°E; collected 10-11 Jul. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, p. 104) and E. Seimund; bm(nh), 1; bnhs, 1. FAS; B:SCS- 10. Tanjong Tuan, Keramat; MALAYSIA: WEST MALAYSIA; 2°24'N, 1 0 1°52'E; collected 6 Apr. 1920 by unknown collector; zrc, 1. Observed FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 173 Aug.-Dec. 1970 by C. H. Southwick and F. C. Cadigan, Jr. (1972, p. 13). FAS; B:WM-24. Tanjong Turut; Pulau Batam, east coast, Kepu- lauan Riau, INDONESIA; ca. 1°07'N, 104°09°E; collected 12 Jul. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, p. 104) and E. Seimund; bm(nh), 2. FAS; B:SCS-10. Tanjung; Borneo: Kalimantan, southeast, IN- DONESIA; 2°1 l'S, 1 15°23'E; collected in 1905 by A. Buxtorf; NHMBa, 1 (skull only). FAS; C:K- 30. Tanjungmorawa, Kabupaten Deli Serdang; Su- matra, INDONESIA; 3°31'N, 98°49'E; collect- ed in 1882 and 1883 by B. Hagen (1890, p. 81); rmnh, 3 (including 2 skulls only, 1 skeleton only). Collected in 1 906 and date unknown by C. Brue- gel; zsbs, 8 (including 3 skins only, 3 skulls only). Collected 9 Jan. 1908 by L. Weigand; zsbs, 1. FAS; B:S-21. Tanjung Puting National Park; Borneo: Kaliman- tan, INDONESIA; 2°45'-2°48'S, 1 1 1°57- 112°01'E; reported present Dec. 1974-Jul. 1975 by J. Supriatna, B. O. Manullang, and E. Soe- kara (1986, p. 186). Reported present before 1983 by G. Davies ([1983], p. 148). Observed in 1983 by K. S. MacKinnon (1986, p. 112). FAS; :K-10. Tapanuli, Teluk; Sumatra, west, INDONESIA; 1°38'N, 98°45'E; collected 21 Feb. and 19 Mar. 1902 by W. L. Abbott (see Miller, 1903a, p. 438); usnm, 2. FAS; B:S-32. Tapanuli Bay. See Tapanuli, Teluk. Tapung-kanan, Sungai; Sumatra, INDONESIA; 0°46'N, 101°06'E; collected ca. 1907 by M. Moszkowski (1909, p. 149); zmb, 1. FAS B:S- 41. Tarusan, Teluk; Sumatra, INDONESIA; 1°13'S, 100°25'E; collected 27 Dec. 1904 by W. L. Ab- bott; usnm, 1 . FAS; B:S-47. Tarusan Bay. See Tarusan, Teluk. Tarutao, Ko, THAILAND; 6°3 1 '-6°44'N; 99°36'- 99°42'E; collected 19 Nov. 1903 and 10 Apr. 1904 by W. L. Abbott (see H. C. Robinson & Kloss, 1910, p. 666; Riley, 1938, p. 15); usnm, 3 (including 1 skull only). FAS; A:T-59. Tasikmalaja. Tasikmalaya. Tasikmalaya; Java, INDONESIA; 7°20'S, 108°12'E; collected Apr -Aug. 1 926 by Dr. Kopstein; mzb, 7 (including 1 skin only). FAS; B:J-4. Tasikmalaya, Preanger region, 1 145 ft (= 350 m); Java, INDONESIA; ca. 7°20'S, 108°12'E; col- lected 18 Jan. 1908 by G. C. Shortridge (see Thomas & Wroughton, 1909a, p. 373); bm(nh), 1. FAS; B:J-4. Tatawa, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°30'-8°31'S, 119°38'-119039'E; monkeys reported absent 1969-1973 by W. Auffenberg (1981, p. 40). C:d. Taungbyauk Chaung, Tavoy River, Tavoy dis- trict; BURMA; ca. 13°45'N, 98°26'E; collected 29-30 Apr. 1936 by H. C. Smith; bm(nh), 4. AUR; A:Bu-14. Tavoy River, mouth, Tavoy district; BURMA; 14°02'N, 98°12'E; collected 21 Apr. 1936 by H. C. Smith; bm(nh), 2. AUR; A:Bu-13. Tawao. See Tawau. Tawar-See. See Bur ni Bebuli. Tawau; Borneo, MALAYSIA: SABAH; 4°15'N, 117°54'E; collected 2 Jan. 1910 by R. C. An- drews (1943, p. 69); amnh, 1. FAS; C:Sab-24. Tawau Hills National Park; Borneo, MALAYSIA: SABAH; ca. 4°20'N, 1 18°00'E; reported present before 1983 by G. Davies ([1983], p. 148). FAS; C:Sab-23. TayNinh, 100 m; VIETNAM; 1 1°18'N, 106°06'E; collected 19 Jan. 1928 by J. Delacour (1928, p. 268; 1929, p. 196) and W. P. Lowe; bm(nh), 1 (skin only). FAS; A:V-5. Tebingtinggi, ca. 25 km SSE; Sumatra, INDO- NESIA; ca. 3°07'N, 99°15'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-23. Tebingtinggi, ca. 35 km ESE; Sumatra, INDO- NESIA; ca. 3°10'N, 99°25'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-23. Tebingtinggi, ca. 55 km ESE; Sumatra, INDO- NESIA; ca. 3°04'N, 99°36'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-24. Tebingtinggi, ca. 60 km SSW; Sumatra, INDO- NESIA; ca. 2°54'N, 99°00'E; reported present Nov. 1971-Jan. 1973 by local residents (Crock- ett & Wilson, 1980, p. 156). FAS; B:S-30. Tebingtinggi, ca. 65 km SE; Sumatra, INDONE- SIA; ca. 2°58'N, 99°38'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-24. Tebingtinggi, ca. 75 km SE; Sumatra, INDONE- SIA; ca. 2°54'N, 99°42'E; observed Nov. 1971- Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-24. Tebingtinggi, ca. 75 km SSW; Sumatra, INDO- NESIA; ca. 2°43'N, 98°53'E; reported present Nov. 1971-Jan. 1973 by local residents (Crock- ett & Wilson, 1980, p. 156). FAS; B:S-29. Tebingtinggi vicinity; Sumatra, INDONESIA; ca. 3°20'N, 99°09'E; observed Nov. 1971-Jan. 1973 174 FIELDIANA: ZOOLOGY by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-23. Tekek; Pulau Tioman, MALAYSIA: WEST MA- LAYSIA; ca. 2°49'N, 104°10'E; observed Mar.- Apr. 1962 by Lord Medway (1966, p. 16). FAS; B:SCS-3. Telaga, Pulau, INDONESIA; 3o01'-3o05'N, 105°58'-105°59'E; observed 13-1 6 Sep. 1899 by W. L. Abbott (in Miller, 1900, p. 245; cf. Kloss, 1903b, p. 73). FAS; B:SCS-29. Telang, 20 m; Borneo: Kalimantan, INDONESIA; 2°30'S, 115°24'E; collected 15 Feb. 1971 by NAMRU 2 Djakarta Detachment (see Van Pee- nen et al., 1974, p. 392); usnm, 1. FAS; C:K- 29. Telapa Burok, Gunong. See Telapak Burok, Gun- ong. Telapak Burok, Gunong, 605-1060 m; MALAY- SIA: WEST MALAYSIA; 2°49'N, 102°04'E; re- ported as probably present Jul. 1978-Jun. 198 1 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-23. Telibon Island. See Talibong, Ko, THAILAND. Telok Anson, near sea level; MALAYSIA: WEST MALAYSIA; 4°02'N, 101°01'E; collected 27 Apr. 1915 by N. Gyldenstolpe (1917a, p. 5); nhrm, 2 (skulls only; skins reported by Gyld- enstolpe, 1917a, p. 25). FAS; B:WM-16. Telok Bahang; Pulau Pinang [1], MALAYSIA: WEST MALAYSIA; 5°28'N, 100°13'E; collect- ed 12 Mar -3 Apr. 1911 by E. Seimund (see Weitzel et al., 1988, p. 102); bm(nh), 1; zrc, 6. FAS; B:SM-2. Telok Betong. See Telukbetung. Telok Dalam. See Dalam, Lhok. Telok Paanji. See Telukpanji, Kotapinang region. Telok Wau; Ko Tarutao, west coast, THAILAND; ca. 6°35'N, 99°35'E; collected 25-28 Dec. 1916 by H. C. Robinson (1917, p. 132); bm(nh), 1; zrc, 1. FAS; A:T-59. Telom, Sungai, 400 ft (= 120 m); MALAYSIA: WEST MALAYSIA; ca. 4°18'N, 101°48'E; col- lected 4 Mar. 1932 by F. A. B. Holloway and A. S. Vernay; bm(nh), 2. FAS; B:WM-8. Teluk Anson. See Telok Anson. Telukbetung; Sumatra, INDONESIA; 5°27'S, 105°16'E; collected before 1907 by W. Denna (see Elliot, 1906, p. 49); fmnh, 1. Clinical ex- amination Aug.-Nov. 1 980 by K. Matsubayashi and D. Sajuthi (1981, p. 48). FAS; B:S-82. Teluk Jolo, 12 km N; Borneo: Kalimantan, IN- DONESIA; ca. 0°18'S, 1 14°03'E; observed 27- 28 Aug. 1 986 by D. J. Chivers and K. M. Burton ([1991], p. 143). FAS; CK-19. Telukkayubutih, ca. 15 km E; Sumatra, INDO- NESIA; ca. 1°09'S, 102°06'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 15 km NNE; Sumatra, IN- DONESIA; ca. 1°06'S, 102°02'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 1 5 km WNW; Sumatra, IN- DONESIA; ca. 1°06'S, 101°52'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 30 km ESE; Sumatra, IN- DONESIA; ca. 1°12'S, 102°13'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 30 km WNW; Sumatra, IN- DONESIA; ca. 1°06'S, 101°44'E; observed Nov. 1971-Jan. 1973 by C M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 45 km ESE; Sumatra, IN- DONESIA; ca. 1°18'S, 102°18'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih, ca. 60 km SE; Sumatra, IN- DONESIA; ca. 1°23'S, 102°21'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS; B:S-51. Telukkayubutih vicinity; Sumatra, INDONESIA; 1°11'S, 101°59'E; observed Nov. 1971-Jan. 1973 by C. M. Crockett and W. L. Wilson (1980, p. 156). FAS;B:S-51. Telukpanji, Kotapinang region; Sumatra, IN- DONESIA; 2°02'N, 100°14'E; collected 6 Jun. 1937 by M. Boogaarts; zrc, 1. FAS; B:S-36. Teluk Terima. See Trima, Teluk. Temaju, Pulau, INDONESIA; 0°28'-0°31'N, 108o51'-108°52'E; monkeys reported absent 5- 6 May 1907 by W. L. Abbott (in Lyon, 1911, p. 59). B:n. Tembeling, Sungai; MALAYSIA: WEST MA- LAYSIA; ca. 4°30'N, 102°15'E; observed Jul.- Sep. 1960 by Y. Furuya (1965, p. 287). FAS; B:WM-11. Tempasuk, Sungai, Kampong Kiau region, Mount Kinabalu, ca. 2000 ft (= 600 m); Borneo, MA- LAYSIA: SABAH; ca. 6°05'N, 116°27'E; col- lected 9 Aug. 1937 by J. A. Griswold, Jr. (1939b, p. 514); mcz, 1. FAS; C:Sab-7. Templer Park, west section, near waterfall; MA- LAYSIA: WEST MALAYSIA; ca. 3°18'N, 101°36'E; observed before 1972 by M. Nadcha- tram (1971, p. 147). FAS; B:WM-19. Tenasserim; BURMA; 12°05'N, 99°01'E; collect- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 175 ed 4 Mar.-9 Apr. 1914 by G. C. Shortridge (in Wroughton, 1915, p. 696); bm(nh), 5 (including 1 skull only); bnhs, 3; fmnh, 1; zrc, 3. AUR; A:Bu-19. Tenasserim River. See Great Tenasserim River and Tagoot (?= Tagu), Great Tenasserim River. Tengger, Pegunungan, 4000 ft (= 1200 m); Java, INDONESIA; 7°55'S, 112°55'E; collected be- fore 1897 by J. H. F. Kohlbrugge (1896b, p. 280); museum unknown, 1 (not examined). FAS; GJ-36. Tenghilan Road. See Tuaran-Kampong Tenghilan Road. Tengi, Sungai, 5 m; MALAYSIA: WEST MA- LAYSIA; 3°3 1 'N, 101°17'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-18. Tenompok, 4700 ft (= 1430 m) and 4900 ft (= 1490 m); Borneo, MALAYSIA: SABAH; ca. 6°00'N, 116°32'E; collected 6-29 Jun. 1937 by J. A. Griswold, Jr. (1939a, p. 410); mcz, 2. FAS; CSab-7. Tenompok Pass. See Kampong Kiau-Tenampok Pass, trail between. Terbanjawan; Sumatra, INDONESIA; ca. 3°32'N, 98°38'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of speci- mens unknown (not seen). FAS; B:S-21. Terengganu, Sungai, vicinity; MALAYSIA: WEST MALAYSIA; ca. 5°00'N, 103°00'E; reported present in 1968 by D. Chivers (1971, p. 80). FAS; B-.WM-10. Terima, Teluk. See Trima, Teluk. Teris, Sungai, ca. 3 km S of Bukit Tapah; MA- LAYSIA: WEST MALAYSIA; ca. 3°35'N, 102°1 l'E; observed Jul.-Dec. 1977 by D. Chiv- ers and G. Davies (1979, p. 20). FAS; B:WM- 15. Terutau, Pulo. See Tarutao, Ko. Thaget. See Thagyet, Little Tenasserim River. Thagyet, Little Tenasserim River; BURMA; 12°06'N, 99°07'E; collected 25 Mar. 1914 by G. C. Shortridge (in Wroughton, 1915, p. 697); bm(nh), 2; bnhs, 3; fmnh, 1; zrc, 1. AUR/FAS/ MUL; A:Bu-19. THAILAND; 5°37'-17°48'N, 97°21'-105°39'E; collected 1858-1861 by H. Mouhot (1864, map at end of vol. 2); bm(nh), 1. Collected in 1862 and 1869 by M. Boucourt; mnhn, 2 (skins only, skulls inside). Collected in 1 906 by C. Bruegel; zsbs, 4 (including 1 skin only, 2 skulls only). Collected date unknown by Dr. Bulkley; amnh, 1 (skin only). Subspecies uncertain; not mapped. Tham Chomphon; THAILAND; 13°37'N, 99°36'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Tak- enaka, and P. Varavudhi (1989, p. 95). Ob- served 20 Dec. 1 990 by N. Aggimarangsee (1992, pp. Ill, 127, pers. comm, Oct. 1993). FAS; A:T-28. Tham Chompol. See Tham Chomphon. Tham Horn, 4 km W of Ban Thap Plik, ca. 75 m; THAILAND; ca. 8°11'N, 98°53'E; collected 3 Jun. 1973 by J. Fooden ([1975], p. 98); fmnh, 1. FAS; A:T-60. Thap Salao, Huai, 200 m; THAILAND; 15°38'N, 99°18'E; observed 15 Jul. 1977 by A. A. Eudey (1979, pp. 90, 198). Subspecies uncertain; A:T- 19. Thateng, Muang, Plateau des Bolovens; LAOS; 15°26'N, 106°23'E; collected 29 Jan.-13 Feb. 1932 by T. D. Carter (see Legendre, 1932, p. 495; 1936, p. 251); amnh, 3 (including 2 skins only); ansp, 2 (external measurements recorded in amnh catalog). AUR/FAS/MUL; A:L-1. Thung Thong Waterfowl Reserve; THAILAND; 8°50'N, 99°1 5'E; observed 22 Jul. 1 976-1 1 Mar. 1977 by P. J. Storer (1978, p. 113). AUR/FAS/ MUL; A:T-48. Tibang, Mt.; Borneo: Kalimantan, INDONESIA; 1°39'N, 114°34'E; collected Nov. 1925 by E. Mjoberg (1929, p. 117); mcz, 1. FAS; C:K-34. Tikus, Pulu, Cocos Islands, AUSTRALIA; 12°06'S, 96°54'E; introduced Jun. 1905-Sep. 1906, spe- cies identification tentative (F. W. Jones, 1910, p. 298). Subspecies uncertain; not mapped. Tillanchong Island, Nicobar Islands, INDIA; 8°26'-8°35'N, 93°37'-93°39'E; primates report- ed absent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, 1903a, p. 114). A:j. Tilu, Gunung; Java, west, INDONESIA; 7°33'S, 107°57'E; collected 6 May 1910 by H. W. van der Weele; rmnh, 1. FAS; B:J-5. Timau, Fatu, northeast, 1200 m; Pulau Timor, Lesser Sunda Islands, INDONESIA; ca. 9°35'S, 1 23°55'E; collected 20 May 1 9 1 1 by C. B. Haniel (see Hellmayr, 1914, p. 5); zsbs, 1. FAS; C:LS- 26. Timor, Pulau, Lesser Sunda Islands, INDONE- SIA; 8°19'-10°23'S, 123°27'-27°18'E; obtained ca. 1846 by W. H. Benson (Blyth, 1846, p. 367; J. Anderson, 1881, p. 65); skin and skull for- merly preserved in zsi (not seen). Reported pres- ent ca. 1856 by A. R. Wallace (1869, p. 326). Collected in 1 890 by B. Hagen; NHMBa, 1 (skull only). FAS; GLS-25 through LS-29. 176 FIELDIANA: ZOOLOGY Tinggi, Pulau, MALAYSIA: WEST MALAYSIA 2°17'-2°19'N, 104°06'-104°08'E; collected 24- 25 Jun. 1908 by H. C. Robinson (in Thomas & Wroughton, 1909b, p. 103; cf. H. C. Robinson, 1919, p. 325) and E. Seimund; bm(nh), 3; zrc, 1. Collected 17 Jun. 1915 by V. Knight; zrc, 1. FAS; B:SCS-6. Tingilan Road. See Tuaran-Kampong Tenghilan Road. Tinjil, Pulau, INDONESIA; 6°57'-6°58'S, 105°46'- 105°49'E; introduced Feb. 1988-Jun. 1990(Kyes et al., 1991, p. 114; 1993, p. 78). Subspecies uncertain; not mapped. Tinonkok, Kampong Kiau region, Mount Kina- balu, ca. 2200 ft (= 670 m); Borneo, MALAY- SIA: SABAH; ca. 6°02'N, 116°31'E; collected 23 Aug. 1937 by J. A. Griswold, Jr. (1939b, p. 514); mcz, 1. FAS; C:Sab-7. Tioman, Pulau, MALAYSIA: WEST MALAY- SIA; 2°43'-2°53'N, 104°07'-104°13'E; collected 4 Oct. 1899 by W. L. Abbott (see Miller, 1900, pp. 203, 246; Riley, 1938, p. 14); usnm, 1. Col- lected 20 Jun. 1 9 1 6 by C. B. Kloss; bm(nh) (skin)/ zrc (skull). Examined for malaria Sep. 1 96 1 and Apr. 1962 by McW. Warren (1966, p. 156). FAS; B:SCS-3. Tjarmara. See Camara. Tjandiroto. See Candiroto. Tjempaga. See Parit. Tjerimai. See Ciremay, Gunung. Tjeringin, near Banjar, Preanger region, east; Java, INDONESIA; ca. 7°22'S, 108°32'E; collected date unknown by H. J. V. Sody; rmnh, 4 (in- cluding 1 skull only). FAS; B:J-3. Tjihara. See Cihara; Pelabuhanratu, Teluk, Ban- tam region. Tjikaran, Walde. See Cikarang forest. Tjikoedjang. See Cikujang. Tji-Tandoei River. See Kalipucang, Ci Tanduy. Tji Wangie. See Ciwangi. Tk. Nipah. See Nipah, Telok, vicinity. Toeroek Tjahoe. See Purukcahu, Sungai Barito. Trang Bom. See Xa Trang Bom. Trima, Teluk; Pulau Bali, Lesser Sunda Islands, INDONESIA; 8°08'S, 1 14°32'E; blood samples taken in 1980 by Y. Kawamoto, K. Nozawa, and Tb. M. Ischak (1981, p. 16). FAS; GLS-1. Trinkat Island, Nicobar Islands, INDIA; 8°02'- 8°08'N, 93°34'-93°37'E; primates reported ab- sent before 1903 by G. S. Miller, Jr. (1902b, p. 792; cf. Kloss, 1903a, p. 114). A:l. Trinkut Island. See Trinkat Island. Trong. See Ban Phra Muang; Tyching. Trusan Kinabatangan; Borneo, MALAYSIA: SA- BAH; ca. 5°49'N, 1 18°20'E; observed May-Jun. 1950 by D.D.Davis (1962, pp. 57, 127, pi. 21). FAS;C:Sab-19. Tuangku, Pulau, INDONESIA; 2°02'-2°14'N, 97°07'-97°22'E; blood samples taken Nov.-Dec. 1986 by J. R. de Ruiter (1993, p. 91). FAS; B:IO-5. Tuangku, Pulau, north coast; INDONESIA; ca. 2°13'N, 97°14'E; collected 24-28 Jan. 1902 by W. L. Abbott (field catalog; cf. Miller, 1903a, p. 438); usnm, 5 (including 2 skulls only; external measurement recorded on tag of one of these skulls). FAS; B:IO-5. Tuan Keramat, Tanjong. See Tanjong Tuan, Ker- amat. Tuanku, Pulau. See Tuangku, Pulau. Tuanku, Pulo. See Tuangku, Pulau. Tuaran, near Kota Kinabalu, near sea level; Bor- neo, MALAYSIA: SABAH; 6°11'N, 116°14'E; collected 24 Jul. 1937 by J. A. Griswold, Jr. (see Coolidge, 1 940, p. 1 23); mcz, 2 (male skin mis- matched with female skull). FAS; C:Sab-5. Tuaran-Kampong Tenghilan Road, new bridge, near sea level; Borneo, MALAYSIA: SABAH; ca. 6°08'N, 116°15'E; collected 29 Jul.-8 Aug. 1937 by J. A. Griswold, Jr. (see Coolidge, 1940, p. 123); mcz, 4. FAS; C:Sab-5. Tulungagung; Java, INDONESIA; 8°04'S, 1 1 1°54'E; collected ca. 1 895 by E. Dubois; rmnh, 6 (skulls only). FAS; C:J-32. Turn Chompol. See Tham Chomphon. Tunggal, Bukit, 10 m; MALAYSIA: WEST MA- LAYSIA; 3°32'N, 10 1°27'E; observed Jul. 1978- Jun. 1981 by C. W. Marsh and W. L. Wilson (1981, p. 232). FAS; B:WM-18. Tyching; THAILAND; 7°33'N, 99°35'E; collected 19 May 1896 by W. L. Abbott (see Riley, 1938, p. 12); usnm, 2. FAS; A:T-61. Ubud; Pulau Bali, Lesser Sunda Islands, INDO- NESIA; 8°30'S, 115°16'E; observed 21 Sep.-3 Oct. 1980 by N. Koyama, A. Asuan, and N. Natsir (1981, p. 1). Observed in 1986, 1990, 1991, and 1992 by B. P. Wheatley and D. K. Harya Putra (1994, p. 246). FAS; CLS-3. Udon Thani, S; THAILAND; ca. 17°10'N, 102°50'E; unconfirmed report, source unspeci- fied, cited by Varavudhi et al. (1992, p. 338; cf. Fooden, 1971, p. 28; 1982, p. 576; Aggimar- angsee, 1992, p. 119). Not mapped. Udjung Kulon National Park. See Ujungkulon, Suaka Margasatwa. Ujungkulon, Suaka Margasatwa; Java, INDO- FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 177 NESIA; 6°39'-6°53'S, 105°12'-105°30'E; ob- served Sep. 1940-Jul. 1955 by A. Hoogerwerf (1970, p. 408). Reported present ca. 1961 by R. K. P. Satmoko (1961, p. 117). Observed Sep. 1969-Jan. 1971) by W. Angst (1973, p. 626; 1975, p. 326). Observed summer 1982 by G. Hohmann and W. P. Peter (1983, p. 44). FAS; B:J-14. Ulu, Sungai; Pulau Natuna Besar, INDONESIA; 3°53'N, 108°24'E; collected 25 Apr. 1909 by V. Knight (see Chasen, 1935a, p. 5); zrc, 1 (skull not examined). FAS; B:SCS-32. Ulu Gombak Forest Reserve; MALAYSIA: WEST MALAYSIA; ca. 3°18'N, 101°47'E; observed Aug. 1962 by R. H. Wharton, D. E. Eyles, McW. Warren, and W. H. Cheong (1964, p. 59). FAS; B:WM-19. Ulu Ijok, 400 ft (= 120 m); MALAYSIA: WEST MALAYSIA; 5°08'N, 100°48'E; collected 27- 30 Dec. 1930 by A. B. Holloway and A. S. Ver- nay (see P. H. Napier, 1981, p. 14); bm(nh), 2. FAS; B:WM-5. Ulu Sebol. See Sekol, Sungai. Ulu Segama Reserve; Borneo, MALAYSIA: SA- BAH; ca. 5°10'N, 1 17°54'E; observed Jun.-Oct. 1968 and Oct. 1969-Oct. 1970 by J. R. MacKinnon (1971, p. 153). FAS; C:Sab-13. Um Pang. See Wong, Nam Mae. Ungar, Pulau, Kepulauan Riau, INDONESIA; 0°37'-0°42'N, 103°28'-103°32'E; reported pres- ent Jun.-Aug. 1 903 by W. L. Abbott (see Miller, 1906c, p. 279). FAS; B:SCS-16. Unter Langkat district; Sumatra, INDONESIA; ca. 3°55'N, 98°20'E; collected 1897-1899 by G. Schneider (1905, p. 72); museum and number of specimens unknown (not seen). FAS; B:S-20. Uwi, Pulau, INDONESIA; 1°04'-1°07'N, 107°22'- 107°25'E; collected 13 Aug. 1899 by W. L. Ab- bott (in Miller, 1900, p. 244; cf. Kloss, 1903b, p. 65; Oberholser, 1919, p. 129); usnm, 1. FAS; B:SCS-26. Vadju, Nusa, Komodo area, Lesser Sunda Islands, INDONESIA; not located, ca. 8°30'S, 1 19°30'E; monkeys reported absent 1 969-1973 by W. Auf- fenberg (1981, p. 40). Not mapped. Ven-Ven (?= Mae Nam Welu); THAILAND; ca. 12°20'N, 102°20'E; observed 1-4 Mar. 1859 by H. Mouhot(1864, vol. 1, p. 152). FAS; A:T-34. Verlaten Island. See Sertung, Pulau. Victoria Island. See Ru, Pulo. Wai, Pulo. See Uwi, Pulau. Wai Sano. See Sano, Wai. Walde Tjikaran. See Cikarang forest. Wang Kaew; THAILAND; ca. 12°47'N, 101°39'E; blood samples taken Aug.-Sep. 1988 by Y. Ka- wamoto, T. Ishida, J. Suzuki, O. Takenaka, and P. Varavudhi (1989, p. 95). FAS; A:T-32. Wat (= Temple) Ban Kan Yai; THAILAND; 15°21'N, 1 04° 13'E; observed 14 Jul. 1989 by N. Aggimarangsee(1992,pp. 109, 122;pers. comm., Oct. 1993). Subspecies uncertain; A:T-17. Wat Ban Muang Khan. See Wat Ban Kan Yai. Wat Ban Rai Don; THAILAND; 1 3°07'N, 99°50'E; reported present 20 Aug. 1 989 by local nun (Ag- gimarangsee, 1992, pp. 110, 132; pers. comm., Oct. 1993). Subspecies uncertain; A:T-37. Wat Bun Thawi; THAILAND; 13°08'N, 99°56'E; observed 18 Aug. 1989 and 1 1 Feb. 1991 by N. Aggimarangsee (1992, pp. 109, 112, 131; pers. comm., Oct. 1993). Subspecies uncertain; A:T- 37. Wat Cha-Am Kiri; THAILAND; 13°48'N, 99°53'E; reported by local residents as exter- minated ca. 1 980 (Aggimarangsee, 1992, pp. 110, 134; pers. comm., Oct. 1993). FAS; A:T-28. Wat Huai Takhaeng; THAILAND; 13°35'N, 99°46'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Tak- enaka, and P. Varavudhi (1989, p. 95). Ob- served 20 Dec. 1 990 by N. Aggimarangsee (1 992, pp. Ill, 128; pers. comm., Oct. 1993). FAS; A:T-28. Wat Khao Bandai It; THAILAND; 13°06'N, 99°56'E; observed 10-1 1 Aug. 1989 and 1 1 Feb. 1991 byN. Aggimarangsee (1992, pp. 109, 112, 131; pers. comm., Oct. 1993). Subspecies un- certain; A:T-37. Wat Khao Chong Phran; THAILAND; 13°43'N, 99°46'E; observed 20 Dec. 1990 by N. Aggi- marangsee (1992, pp. Ill, 127; pers. comm., Oct. 1993). FAS; A:T-28. Wat Khao Khang. See Wat Ngern Rung Sawang. Wat Khao Khan Hok; THAILAND; 13°50'N, 99°38'E; observed 22 Dec. 1990 by N. Aggi- marangsee (1992, pp. Ill, 126; pers. comm., Oct. 1993). FAS; A:T-28. Wat Khao Noh; THAILAND; 15°57'N, 99°53'E; observed 10 Mar. 1967 by J. Fooden (1971, p. 17). Blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Takenaka, and P. Varavudhi (1989, p. 95). Reported pres- ent 22 Feb. 1991 by N. Aggimarangsee (1992, pp. 112, 123; pers. comm., Oct. 1993). Subspe- cies uncertain; A:T-4. Wat Khao Phlu. See Wat Tham Khao Phlu. Wat Khao Sompoad (?= Som Phot); THAILAND; ca. 15°10'N, 101°17'E; observed 11 Jan. 1991 178 FIELDIANA: ZOOLOGY by N. Aggimarangsee ( 1 992, pp. Ill, 1 23). Sub- species uncertain; A:T-8. Wat Khao Takhrao; THAILAND; 13°13'N, 99°56'E; observed 21 Aug. 1989 and 11 Feb. 1991 by N. Aggimarangsee (1992, pp. 1 10, 1 12, 132; pers. comm., Oct. 1993). Subspecies un- certain; A:T-37. Wat Khao Takieb; THAILAND; 1 2°3 1 'N, 99°59'E; observed 16-17 Sep. 1 990 by N. Aggimarangsee (1992, pp. 110, 134; pers. comm., Oct. 1993). Subspecies uncertain; A:T-38. Wat Khao Tamon; THAILAND; 13°07'N, 99°56'E; reported present 4 Feb. 1991 by N. Aggimarangsee (1992, pp. 112, 132; pers. comm., Oct. 1993). Subspecies uncertain; A:T-37. Wat Khao Wong Kot; THAILAND; 15°01'N, 100°33'E; reported present 24 Jun. 1989 by un- specified informants (Aggimarangsee, 1992, pp. 109, 115, 124; pers. comm., Oct. 1993). Sub- species uncertain; A:T-6. Wat Khao Yod Thong; THAILAND; 13°28'N, 99°43'E; reported present 25 Dec. 1990 by N. Aggimarangsee (1992, pp. Ill, 127; pers. comm., Oct. 1993). FAS; A:T-28. Wat Khuha Phimuk; THAILAND; ca. 6°31'N, 101°14'E; observed 21-22 Sep. 1990 by N. Ag- gimarangsee (1992, pp. 110, 139; pers. comm., Oct. 1993). FAS; A:T-69. Wat Khuha Sawan; THAILAND; 7°37'N, 100°05'E; observed 20-21 Sep. 1990 and 26-28 Feb. 1991 by N. Aggimarangsee (1992, pp. 1 10, 112, 138; pers. comm., Oct. 1993). FAS; A:T- 65. Wat Koo Pra Kona; THAILAND; 15°33'N, 103°49'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Tak- enaka, and P. Varavudhi (1989, p. 95). Ob- served 14-16 Jul. 1989 and 17-18 Jan. 1991 by N. Aggimarangsee (1992, pp. 109, 112, 120; pers. comm., Oct. 1993). Subspecies uncertain; A:T- 13. Wat Krieng Krai Klang; THAILAND; 15°44'N, 100°11'E; observed 22 Feb. 1991 by N. Aggi- marangsee (1992, pp. 112, 123; pers. comm., Oct. 1993). Subspecies uncertain; A:T-5. Wat Kut; THAILAND; 13°09'N, 99°57'E; ob- served 1 9 Aug. 1 989 by N. Aggimarangsee (1992, pp. 110, 131; pers. comm., Oct. 1993). Subspe- cies uncertain; A:T-37. Wat Ngern Rung Sawang; THAILAND; 13°42'N, 99°46'E; observed 20 Dec. 1990 by N. Aggi- marangsee (1992, pp. Ill, 128; pers. comm., Oct. 1993). FAS; A:T-28. Wat Noi Chompoo; THAILAND; 14°36'N, 100°09'E; reported present 7 Jul. 1989 by local monk (Aggimarangsee, 1992, pp. 109, 125; pers. comm., Oct. 1993). Subspecies uncertain; A:T- 23. Wat Phra Buddha Chai; THAILAND; 14°27'N, 100°57'E; observed 11 Jan. 1991 by N. Aggi- marangsee (1992, pp. Ill, 125; pers. comm., Oct. 1993). Subspecies uncertain; A:T-24. Wat Ratch Singkhorn; THAILAND; 13°34'N, 99°47'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ishida, J. Suzuki, O. Tak- enaka, and P. Varavudhi (1989, p. 95). Ob- served 24 Dec. 1 990 by N. Aggimarangsee (1992, pp. Ill, 129; pers. comm., Oct. 1993). FAS; A:T-28. Wat Suwankuha (cf. Wat Khuha Sawan); THAI- LAND; ca. 7°37'N, 100°05'E; blood samples taken Aug.-Sep. 1988 by Y. Kawamoto, T. Ish- ida, J. Suzuki, O. Takenaka, and P. Varavudhi (1989, p. 95). FAS; A:T-65. Wat Tha Mai Lai; THAILAND; 10°30'N, 98°58'E; observed 18 Sep. 1990 by N. Aggimarangsee (1992, pp. 110, 137; pers. comm., Oct. 1993). AUR/FAS/MUL; A:T-43. Wat Tham Khao Phlu; THAILAND; 10°44'N, 99°19'E; reported present 1-2 Dec. 1990 by lo- cal residents (Aggimarangsee, 1 992, pp. Ill, 137; pers. comm., Oct. 1993). AUR/FAS/MUL; A:T- 42. Wat Tham Kunchhorn; THAILAND; 13°29'N, 99°42'E; observed 19 Dec. 1990 by N. Aggi- marangsee (1992, pp. Ill, 126; pers. comm., Oct. 1993). FAS; A:T-28. Wat Thammikaram Varaviharn; THAILAND; 1 1°49'N, 99°48'E; observed 17 Sep. 1990 by N. Aggimarangsee (1992, pp. 1 10, 134; pers. comm., Oct. 1993). AUR/FAS/MUL; A:T-41. Wat Tham Sala; THAILAND; 13°49'N, 100°07'E; observed 12 May 1989 and 15-19 Dec. 1990 by N. Aggimarangsee (1992, pp. 109, 1 1 1, 125; pers. comm., Oct. 1993). FAS; A:T-29. Wat Tham Sua; THAILAND; 8°07'N, 98°55'E; observed 1 9-20 Sep. 1 990 by N. Aggimarangsee (1992, pp. 110, 138; pers. comm., Oct. 1993). FAS; A:T-60. Wat Tham Suwan Khuha; THAILAND; 8°26'N, 98°28'E; observed 17-21 Nov. 1987 by R. Boonratana (1988, p. 75). Reported present 19 Sep. 1990 by local residents (Aggimarangsee, 1992, pp. 110, 137; pers. comm., Oct. 1993). Subspecies uncertain; A:T-51. Waw, Telok. See Telok Wau. Wawo. See Ntori, Sape district. Way Kambas. See Kambas, Wai. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 179 We, Pulau, INDONESIA; 5°47'-5°53'N, 95°13'- 95°23'E; reported present before 1981 by J. T. Marshall (Crockett & Wilson, 1980, p. 156). Blood samples taken Nov.-Dec. 1986 by J. R. de Ruiter (1993, p. 91). Subspecies uncertain (see Scheffrahn et al., 1994, p. 136); B:IO-l. Weh, Pulau. See We, Pulau. Weld's Hill. See Nanas, Bukit, Kuala Lumpur. West Bali National Park. See Bali Barat National Park. West Bali Wildlife Reserve. See Bali Barat Na- tional Park. Whykeong Union Council. See Bilasodia; Bimir- dia; Ghorardia; Ochodia; Rukumodia. Wimpong, limestone rocks; BURMA; ca. 16°53'N, 97°28'E; collected ca. 1877-1878 by W. Davison (see Hume & Davison, 1878, p. 524; Thomas, 1886, pp. 65, 66); bm(nh), 1. AUR; A:Bu-9. Wong, Nam Mae, 40 mi (= 65 km) E of Um Pang, 1000 ft (= 300 m); THAILAND; ca. 15°55'N, 99°10'E; collected 9 Feb. 1924 by A. S. Vernay (see Lowe, 1932, p. 197; 1933, p. 260); amnh, 1;bm(nh), 1. AUR; A:T-18. Wong, Nam Mae, 53 mi (= 85 km) E of Um Pang, 800 ft (245 m); THAILAND; ca. 15°55'N, 99°25'E; collected 18 Feb. 1924 by A. S. Vernay (see Lowe, 1932, p. 197; 1933, p. 260); amnh, 1. AUR; A:T-3. Wonokerto. See Wonokojo (?= Wonokerto), Dampit district, southern Malang region. Wonokojo (?= Wonokerto), Dampit district, southern Malang region; Java, INDONESIA; ca. 8°13'S, 112°35'E; collected Apr. 1930 by H. J. V. Sody; rmnh, 3. FAS; C:J-34. Wonosobo, Propinsi Lampung; Sumatra, IN- DONESIA; 5°30'S, 104°30'E; collected 28 Dec. 1924 by H. J. V. Sody (see Hooijer, 1962b, p. 44); rmnh, 1 (skull only). FAS; B:S-79. Wynkoops Bay. See Pelabuhanratu, Teluk, Ban- tam region. Xa Trang Bom; VIETNAM; 10°57'N, 107°01'E; collected 1 Jun. 1918 by C. B. Kloss (1919b, p. 401; cf. Weitzel et al., 1988, p. 146); zrc, 1. FAS; A:V-3. YaoNoi, Ko, THAILAND; 8°05'-8°l l'N, 98°34'- 98°38'E; collected 30 Jan. 1918 by local collec- tors employed by H. C. Robinson and C. B. Kloss (1919, p. 87); zrc, 1. AUR/FAS/MUL; A:T-52. Yaring, tidal creeks near; THAILAND; ca. 6°52'N, 101°22'E; observed Jun. 1901 by N. Annandale and H. C. Robinson (1903, p. xxxix; in Bonhote, 1903, p. 4). FAS; A:T-68. Yaring region; THAILAND; ca. 6°52'N, 101°22'E; collected 19 Jun. 1899 by R. Evans and F. F. Laidlaw (see P. H. Napier, 1981, p. 17; cf. Bon- hote, [1901], p. 870); University Museum of Zoology, Cambridge, 1 (skull only, not seen). FAS; A:T-68. Ye Forest, Ataran district, Moulmein region; BURMA; ca. 16°10'N, 98°00'E; collected Nov. 1 9 1 0 by G. F. B. R. Thurling (see Pocock, 1939, p. 81); bm(nh), 1. AUR; A:Bu-l 1. Zadetkyi Kyun, BURMA; 9°50'-10°03'N, 98°07'- 98°18'E; collected 9 Dec. 1900 by W. L. Abbott; usnm, 1. AUR; A:Bu-24. 180 FIELDIANA: ZOOLOGY Appendix 3: Dorsal Pelage Color Saturation in Fringing-Island Samples of Macaco, fascicularis Compared with Saturation in Core-Area Reference Samples Island2 Fringing-island samples Core-area reference samples N 0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 Mean Mean N Frequencies at SI values FI> CA> CA3 FI4 P* Shallow-water islands West of Sumatra Tuangku 3 1 2 Mursala 2 2 Tanahbala 2 1 1 West of Isthmus of Kra and Malay Peninsula Kathema 4 2 1 1 Mibya 2 2 Kadan 1 1 Letsok-aw 1 1 Lanbi 1 1 Zadetkyi 1 1 Ru 1 1 Phayam 1 1 Na Ka Yai 1 Yao Noi 1 1 Rang Yai 1 1 Phi Phi Don 4 2 2 Talibong 2 2 Tarutao 6 1 5 Butang 1 1 Langkawi 1 1 Burau 1 1 Pinang [1] 8 6 1 1 Pintu Gedong 3 1 2 East of Sumatra Bengkalis 1 1 Singapore 11 3 5 3 Karimun 7 6 1 Kundur 2 1 1 Durian 2 1 1 Sugi 1 1 Bulan 3 1 2 Batam 3 2 1 Galang 4 1 3 Nguwal 6 1 3 2 Bintan 8 1 2 4 Mapur 1 1 Lingga 2 2 Bangka 3 2 1 East of Isthmus of Kra and Malay Peninsula Phangan 2 2 Samui 2 1 1 Redang 2 Pinang [2] 2 2 Tioman 11 2 7 2 Acheh 1 1 Pemanggil 2 1 1 Aur 3 1 2 Tinggi 5 2 3 1.3 1.4 70 x > 0.50 1.5 1.4 70 X >0.50 1.2 1.5 19 x > 0.20 1.4 1.1 22 X >0.20 1.5 1.1 22 X >0.20 2.0 1.1 22 X >0.10 1.5 1.1 36 X >0.40 1.5 1.1 36 X >0.40 2.0 1.1 20 X 0.20 1.5 1.1 20 X >0.20 1.0 1.1 20 x > 0.20 2.5 1.1 20 X 0.10 2.0 1.1 20 X 0.20 1.5 1.0 18 X >0.20 1.3 1.0 18 X >0.10 1.0 1.0 18 — - > 0.20 1.4 1.0 18 X < 0.02* 1.5 1.0 18 X >0.20 1.5 1.0 18 X >0.20 1.5 1.0 18 X >0.20 1.2 1.4 24 x > 0.05 1.8 1.4 15 X >0.10 1.5 1.5 18 - > 0.20 1.5 1.4 26 X >0.50 1.6 1.4 26 X >0.20 1.8 1.5 18 X >0.20 1.8 1.5 18 X >0.20 1.5 1.4 26 X >0.50 1.8 1.4 26 X >0.05 1.7 1.4 26 X >0.20 1.4 1.4 26 — - > 0.50 1.6 1.4 26 X >0.20 1.8 1.4 26 X < 0.05* 1.5 1.4 26 X >0.50 1.5 1.3 46 X >0.20 1.7 1.2 28 X < 0.05* 1.0 1.0 18 - > 0.20 1.5 1.0 18 X >0.20 2.5 1.3 15 X 0.02* 1.5 1.3 15 X >0.20 1.5 1.4 26 X >0.50 1.5 1.4 26 X >0.50 1.5 1.4 26 X >0.50 1.8 1.4 26 X >0.05 1.3 1.4 26 : < > 0.40 FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 181 Appendix 3: Continued Fringing-island samples Island2 Core-area reference samples N 0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 Mean Mean N Frequencies at SI values FI > CA > CA3 FI4 South of Indochinese Peninsula Khram Yai 10 4 6 Chang Kut 3 8 2 6 1 1 Phu Quoc Con Son 2 8 2 1 5 2 Ba 3 1 1 West of Borneo Siantan 3 2 1 Laut 2 1 Natuna Besar 6 4 1 Lagong Subi-kecil 1 3 1 2 Serasan 2 1 1 Uwi 1 Benua 2 1 Pejantan Karimata 1 2 1 1 1 Serutu 1 1 Belitung 2 1 1 South of Borneo Karimunjawa Bawean 6 6 3 1 1 4 1 Matasiri 1 1 Kangean Bali 1 19 3 7 1 3 5 1 Northeast of Borneo Banggi Cagayan Sulu Sebatik 5 1 3 1 2 1 1 1 1 De vp-\ West of Sumatra Katchall 2 Little Nicobar 3 Great Nicobar 3 Simeulue 13 Lasia 2 Nias6 12 2 7 1 2 Lesser Sunda Islands Penida 3 3 Lombok 2 2 Sumbawa 9 6 3 Flores 7 2 3 1 Sumba 8 2 6 Kambing Timor 1 11 1 8 3 Philippines and Maratua Balabac 1 Palawan 12 3 Culion 6 5 Busuanga Mindoro 1 7 1 2 1.8 1.3 1.0 1.0 1.7 1.3 1.7 1.3 1.6 1.5 1.3 2.0 1.0 1.3 2.0 2.0 2.0 2.3 2.7 1.5 1.5 1.5 1.3 2.1 2.0 1.2 1.1 1.0 1.0 1.3 1.3 1.3 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.5 1.3 1.2 1.4 1.2 1.2 27 28 28 16 15 15 33 33 32 32 32 32 33 33 33 38 38 38 37 40 30 40 40 1.3 20 1.2 49 1.3 45 1 3.5 1.4 68 3 4.0 1.4 68 3 4.0 1.4 68 13 4.0 1.4 68 2 4.0 1.4 68 1.6 1.4 69 1.5 1.2 40 1.5 1.2 40 1.2 1.2 40 1.3 1.2 40 1.4 1.2 40 1.0 1.2 40 1.1 1.2 40 3.0 1.3 65 2.9 1.3 65 2.6 1.3 65 2.0 1.3 65 2.9 1.3 65 < 0.001*** >0.20 > 0.90 > 0.20 > 0.20 >0.20 >0.50 >0.50 > 0.50 > 0.90 > 0.50 > 0.20 > 0.20 > 0.50 >0.20 >0.20 >0.20 > 0.05 < 0.001*** >0.20 >0.50 > 0.40 >0.50 0.05 >0.10 > 0.50 < 0.02* < 0.01** < 0.01** < 0.001*** < 0.02* >0.10 > 0.10 > 0.20 > 0.50 > 0.50 > 0.20 > 0.50 >0.50 >0.05 < 0.001*** < 0.001*** > 0.10 < 0.001*** 182 FIELDIANA: ZOOLOGY Appendix 3: Continued Fringing-island samples Island2 Core-area reference samples N 0.5 1.0 1.5 2.0 2.5 3.0 3.5 4.0 Mean Mean N Frequencies at SI values FI> CA> CA3 FI4 Luzon 24 Samar 3 Leyte 5 Negros 47 Mindanao 65 Balut 1 Basilan 3 Tawitawi 2 Maratua 4 32 8 16 12 16 9 20 3 5 12 4 1 2.9 3.0 3.0 2.4 1.6 3.0 1.7 1.0 1.3 1.3 1.3 1.3 1.3 1.3 1.3 1.3 65 65 65 65 65 65 65 65 4.0 1.3 42 < 0.001*** < 0.01** < 0.001*** < 0.001*** < 0.02* >0.05 >0.10 >0.20 < 0.01** 1 Core-area reference samples, except those compared with Philippine samples, consist of specimens (n > 15) collected in 2-degree latitude-longitude blocks (Table 2) situated nearest to the respective fringing islands; all Philippine samples are compared with a core-area reference sample that consists of 65 specimens collected in northern Borneo, in three 2-degree blocks that border the Sulu Sea. 2 For details, see Appendix 2 and Fooden (1991, p. 34). 3 Fringing-island (FI) sample mean exceeds core-area (CA) reference sample mean. 4 Core-area reference sample mean exceeds fringing-island sample mean. 5 Mann-Whitney (7-test, two-tailed; * P < 0.05, ** P < 0.01, *** P < 0.001. 6 Cf. Scheffrahn et al. (1994, p. 136). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 183 Appendix 4: Dorsal Pelage Color Erythrism in Fringing-Island Samples of Macaca fascicularis Compared with Erythrism in Core- Area Reference Samples Fringing-island samples Core-area reference Frequencies at erythrism N index values Mean samples Mean N FI> CA3 CA> FI4 Island2 1.0 1.5 2.0 2.5 3.0 p* Shallow-water islands West of Sumatra Tuangku 3 1 1 1 2.0 1.8 72 X >0.40 Mursala 2 2 2.5 1.8 72 X < 0.05* Tanahbala 2 1 1 1.8 2.1 19 X > 0.20 West of Isthmus of Kra and Malay Peninsula Kathema 4 3 1 1.3 1.1 22 X > 0.50 Mibya 2 2 1.0 1.1 22 X >0.50 Kadan 1 1 2.5 1.1 22 X >0.05 Letsok-aw 1 1 1.5 1.2 36 X >0.20 Lanbi 1 1 2.0 1.2 36 X > 0.10 Zadetkyi 1 1 2.0 1.4 20 X > 0.20 Ru 1 1 2.0 1.4 20 X >0.20 Phayam 1 1 1.0 1.4 20 X >0.20 Na Ka Yai 1 1 2.0 1.4 20 X >0.20 Yao Noi 1 1 2.0 1.4 20 X >0.20 Rang Yai 1 1 1.5 1.4 18 X >0.20 Phi Phi Don 4 4 1.5 1.4 18 X >0.20 Talibong 2 1 1 1.8 1.4 18 X >0.20 Tarutao 6 3 2 1 1.8 1.4 18 X >0.05 Butang 1 1 2.0 1.4 18 X >0.20 Langkawi 1 1 1.5 1.4 18 X >0.20 Burau 1 1 2.5 1.4 18 X >0.20 Pinang [1] 8 2 3 2 1 2.1 1.9 24 X > 0.20 Pintu Gedong 3 2 1 2.0 2.0 15 - - > 0.20 East of Sumatra Bengkalis 1 1 2.5 2.1 18 X >0.20 Singapore 11 3 1 3 2 2 2.0 1.9 26 X > 0.50 Karimun 7 1 2 3 1 2.3 1.9 26 X >0.10 Kundur 2 2 2.0 2.1 18 X >0.20 Durian 2 1 1 2.8 2.1 18 X >0.20 Sugi 1 1 2.0 1.9 26 X >0.50 Bulan 3 2 1 2.0 1.9 26 X >0.90 Batam 3 1 2 1.8 1.9 26 X >0.50 Galang 4 1 1 1 1 1.8 1.9 26 X > 0.50 Nguwal 6 3 2 1 2.3 1.9 26 X >0.50 Bintan 8 1 5 1 1 2.1 1.9 26 X >0.20 Mapur 1 1 2.5 1.9 26 X > 0.20 Lingga 2 2 2.0 1.8 46 X > 0.20 Bangka 3 1 2 1.7 1.6 28 X >0.50 East of Isthmus of Kra and Malay Peninsula Phangan 2 2 1.0 1.4 18 X >0.20 Samui 2 1 1 1.8 1.4 18 X >0.20 Redang 2 2 3.0 1.6 15 X 0.05 Pinang [2] 2 2 2.5 1.6 15 X >0.05 Tioman 11 3 3 3 1 1 1.7 1.9 26 X >0.50 Acheh 1 1 2.5 1.9 26 X >0.20 Pemanggil 2 1 1 1.3 1.9 26 X >0.10 Aur 3 2 1 1.5 1.9 26 X >0.40 Tinggi 5 2 2 1 1.9 1.9 26 - - >0.90 South of Indochinese Peninsula Khram Yai 10 9 1 1.1 1.1 27 — _ > 0.50 Chang 3 1 2 1.3 1.1 28 X >0.05 184 FIELDIANA: ZOOLOGY Appendix 4: Continued Fring inn-island samples Core-area reference Frequencies at jrythrism N index values Mean samples Mean N FI> CA3 CA> FI4 Island2 1.0 1.5 2.0 2.5 3.0 P5 Kut 8 5 1 2 1.3 1.1 28 X >0.10 Phu Quoc 2 1 1 1.3 1.4 16 X >0.20 Con Son 8 3 3 2 1.4 1.4 15 — — >0.20 Ba 3 1 2 1.7 1.4 15 X >0.20 West of Borneo Siantan 3 2 1 2.3 1.5 32 X < 0.05* Laut 2 1 1 1.8 1.5 32 X >0.20 Natuna Besar 6 1 2 3 2.1 1.5 31 X < 0.02* Lagong 1 1 2.5 1.5 31 X >0.10 Subi-kecil 3 3 1.0 1.5 31 X >0.05 Serasan 2 1 1 2.3 1.5 31 X >0.05 Uwi 1 1 2.5 1.5 32 X >0.05 Benua 2 1 1 2.0 1.5 32 X >0.20 Pejantan 1 1 1.5 1.5 32 — — >0.90 Karimata 2 1 1 1.3 1.5 38 X >0.40 Serutu 1 1 1.0 1.5 38 X >0.20 Belitung 2 2 1.5 1.5 38 - - >0.50 South of Borneo Karimunjawa 6 6 1.0 1.2 37 X >0.05 Bawean 6 3 2 1 1.3 1.2 40 X >0.20 Matasiri 1 1 1.5 1.5 31 — — >0.90 Kangean 1 1 1.5 1.2 40 X >0.20 Bali 19 14 2 3 1.1 1.2 40 X >0.20 Northeast of Borneo Banggi 5 3 2 1.2 1.1 20 X >0.20 Cagayan Sulu 1 1 1.5 1.1 49 X >0.20 Sebatik 3 1 2 1.3 1.2 43 X >0.20 Deep-water islands West of Sumatra Katchall 2 2 1.0 1.8 70 X >0.05 Little Nicobar 3 3 1.0 1.8 70 X < 0.05* Great Nicobar 3 3 1.0 1.8 70 X < 0.05* Simeulue 13 12 1 1.0 1.8 70 X < 0.001*** Lasia 2 2 1.0 1.8 70 X >0.05 Nias 12 2 8 2 2.0 1.8 71 X >0.05 Lesser Sunda Islands Penida 3 1 2 1.8 1.2 40 X < 0.02* Lombok 2 2 1.0 1.2 40 X >0.20 Sumbawa 9 7 2 1.1 1.2 40 X >0.40 Flores 7 6 1 1.1 1.2 40 X >0.20 Sumba 8 5 3 1.4 1.2 40 X >0.40 Kambing 1 1 1.5 1.2 40 X >0.20 Timor 11 5 4 2 1.4 1.2 40 X >0.10 Philippines and Maratua Balabac 1 1 2.0 1.2 65 X >0.10 Palawan 9 5 4 2.7 1.2 65 X < 0.001*** Culion 2 2 2.5 1.2 65 X < 0.02* Busuanga 1 1 2.0 1.2 65 X >0.10 Mindoro 3 1 2 2.3 1.2 65 X < 0.01** Luzon 10 5 5 2.3 1.2 65 X < 0.001*** Samar 3 1 2 1.8 1.2 65 X < 0.02* Negros 45 1 32 12 2.1 1.2 65 X < 0.001*** FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 185 Appendix 4: Continued Fringing-island samples Frequencies at erythrism index values Core-area reference samples Island2 N 1.0 1.5 2.0 2.5 3.0 Mean Mean N FI > CA3 CA > FI4 Mindanao 60 4 Balut 1 Basilan 3 1 Maratua 4 4 29 22 1.7 2.5 1.3 1.0 1.2 1.2 1.2 1.2 65 65 65 42 < 0.001*** >0.05 >0.20 >0.20 1 Core-area reference samples, except those compared with Philippine samples, consist of specimens (n > 15) collected in 2-degree latitude-longitude blocks (Table 4) situated nearest to the respective fringing islands; all Philippine samples are compared with a core-area reference sample that consists of 65 specimens collected in northern Borneo, in three 2-degree blocks that border the Sulu Sea. 2 For details, see Appendix 2 and Fooden (1991, p. 34). 3 Fringing-island (FT) sample mean exceeds core-area (CA) reference sample mean. 4 Core-area reference sample mean exceeds fringing-island sample mean. 5 Mann-Whitney U-test, two-tailed; * P < 0.05, ** P < 0.01, *** P < 0.001. 186 FIELDIANA: ZOOLOGY Appendix 5: Crown Color Pattern Frequency in Samples of Macaco, fascicularis That Include Dark-Crowned Specimens1 (cf. Table 5) N Crown color pattern Sample area Crown colored like back, or brighter Crown with Crown with diffuse blackish clearly defined streak or wash blackish patch2 Core area Burma Taungbyauk Thagyet Ban Sadein 4 5 1 3 3 1 2 1 Thailand Ban Tamrong Phato Lat Bua Khao Chumphon, Khlong 7 2 2 6 1 1 1 2 Vietnam Sontra Peak Tay Ninh Xa Trang Bom Ho Chi Minh City "Cochin China"3 1 1 1 1 2 1 1 1 1 1 1 Indonesia: Kalimantan Pelaihari 1 1 Total 28 14 13 1 Burma Shallow-water fringing islands Lanbi Zadetkyi 1 1 1 1 Thailand Khram Yai4 Kut Talibong 10 8 2 1 1 3 7 7 1 Vietnam Phu Quoc Con Son5 Ba 2 9 3 2 1 8 2 1 Malaysia, West Aur 3 2 1 Indonesia Pejantan Bangka Belitung Karimunjawa6 Bali 1 3 2 6 19 2 2 18 1 1 2 4 1 Total 70 26 28 16 Philippines Mindanao Basilan Island(s) unknown Deep-water fringing islands 60 58 3 2 5 3 2 1 2 Total 68 63 4 1 1 Excludes infants. 2 Cf. Figure 7. c Cf. Elliot (1909, p. 252) and Kloss (1926, p. 358). 4Cf. Kloss (1919c, p. 347). 5 Cf. Kloss (1921, p. 76; 1926, p. 358). 6Cf. Sody(1949, p. 132). FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 187 Appendix 6: Geographic Variation of Lateral Facial Crest Pattern in Samples of Macaco, fascicularis (Including Infants) Collected or Observed in the Indochinese Peninsula, Isthmus of Kra, Malay Peninsula, Neighboring Shallow-Water Fringing Islands, Nic- obar Islands, P. Simeulue, and P. Lasia (cf. Table 6; Figs. 8, 9) A. Samples Homogeneous for Lateral Facial Crest Pattern 1 . Indochinese Peninsula, Isthmus of Kra, Malay Peninsula a. Area homogeneous for the infrazygomatic pattern, n = 41. BANGLADESH, Whykeong (21°05'N, 90°12'E), 1 (cf. M. A. R. Khan, 1985, p. 29). BURMA, ArakanDiv.(19°27'N, 93°31'E), l;Wimpong (16°53'N, 97°28'E), 1; Haungtharaw (16°30'N, 98°13'E), 4; Ye Forest (16°10'N, 98°00'E), 1; Tavoy River (14°02'N, 98°12'E), 2; Taungbyauk (13°45'N, 98°26'E), 4; Mergui (12°26'N, 98°36'E), 2; Tagoot (12°15'N, 99°03'E), 2; Tenasserim (12°05'N, 99°01'E), 11; Ban Sadein (10°20'N, 98°32'E), 1. THAILAND, Wong, Nam Mae, 65 km E of Urn Pang (15°55'N, 99°10'E), 2; Wong, Nam Mae, 85 km E of Urn Pang(15°55'N, 99°25'E), 1; BanTamrongPhato(14°54'N, 98°31'E), 7; COUNTRY UNCERTAIN, "Bengale" (coordinates unknown), 1. b. Area homogeneous for the transzygomatic pattern, n = 66 (excludes 2 specimens with transzy- gomatic crests that were collected at unspecified localities in Thailand). THAILAND, Pak Nam Pho (15°43'N, 100°12'E), 3; Kata Taek (15°28'N, 99°23'E), 6; Lat Bua Khao (14°52'N, 101°36'E), 2; Ban Phu Toie (14°42'N, 99°07'E), 2; Ban Huai Maenam Noi (14°25'N, 98°5 l'E), 2; Aranyaprathet (13°41'N, 102°30'E), 1; Laem Sing Mountains (12°29'N, 102°06'E), 1; Ban Thap Plik (8°11'N, 98°53'E), 2; Tham Horn (8°1 1 'N, 98°53'E), 1 ; Ban Nong Kok (8°06'N, 98°52'E), 1 ; Tyching (7°33'N, 99°35'E), 2; Kantang (7°25'N, 99°30'E), 1; Khao Rang Kai (7°19'N, 99°48'E), 1; Ban Phra Muang (7°21'N, 99°28'E), 2; Ban Sai Kau (6°38'N, 101°08'E), 1; Kampong Biserat (6°32'N, 101°14'E), 3. VIETNAM, Sontra Peak (16°07'N, 108°21'E), 1; Tay Ninh (1 1°18'N, 106°06'E), 1; Xa Trang Bom (10°57'N, 107°00'E), 1; Ho Chi Minh City (10°45'N, 106°42'E), 2; Cochin China (coordinates uncertain), 1. VIETNAM OR CAMBODIA, no locality, 2. WEST MALAYSIA, 16 localities, 27. 2. Neighboring shallow-water fringing islands a. Island group homogeneous for the infrazygomatic pattern, n = 10. BURMA, Mergui Archipelago, Kathema(13°39'N, 98°12'E), 4; Mibya(13°36'N, 98°12'E), 2; Kadan(12°30'N, 98°22'E), 1; Letsok- aw (11°37'N, 98°15'E), 1; Lanbi (10°50'N, 98°15'E), 1; Zadetkyi (9°58'N, 98°13'E), 1. b. Islands homogeneous for the transzygomatic pattern, n = 102. THAILAND, Khram Yai (12°42'N, 100°47'E), 1 1; Chang (12°00'N, 102°23'E), 3; Kut(l 1°40'N, 102°35'E), 8; Phangan(9°45'N, 100°00'E), 2; Samui (9°30'N, 100°00'E), 2; Phi Phi Don (7°45'N, 98°47'E), 4; Talibong (7°15'N, 99°29'E), 2; Tarutao (6°35'N, 99°40'E), 6; Butang (6°32'N, 99°12'E), 1. VIETNAM, Phu Quoc (10°13'N, 103°57'E), 1; Con Son (8°43'N, 106°35'E), 9; Ba (8°39'N, 106°33'E), 3. WEST MALAYSIA, 11 islands, 39. SINGAPORE (1°23'N, 103°50'E), 11. FIELDIANA: ZOOLOGY B. Samples Heterogeneous for Lateral Facial Crest Pattern Latitude (N) Longitude (E) N Frequencies of lateral facial crest patterns Sample area Infra- Trans- Asym- zygomatic zygomatic metric 1 . Indochinese Peninsula, Isthmus of Kra Thailand Phu Phan Ban Mae Na Ree Ban Nam Lai Tai Pak Chong Pran Buri Chumphon Nakhon Si Thammarat BanNa 16°42' 16°25' 16°10' 14°42' 12°24' 10°22' 8°26' 8° 10' 104°24' 99°23 99°20 101°28' 100°00' 99° 10' 99°58' 100°12' 1 3 6 2 2 2 4 1 1 1 2 1 2 3 1 1 1 1 1 2 4 Laos Thateng 15°26' 106°24' 5 4 1 Vietnam Lac Giao 12°40' 108°00' 1 1 Burma Thagyet Pakchan River, Maliwun Pakchan River, Bankachon 12c06' 10°14' 10°09' 99°07' 98°37' 98°36' 7 1 2 5 2 1 1 1 Total 37 17 16 4 2. Neighboring shallow- water fringing islands Thailand Ru Phayam NaKaYai Yao Noi Rang Yai 9°57' 9°44' 8° 12' 8°07' 7°57' 98°32' 98°25' 98°31' 98°37' 98°26' 1 1 1 1 1 1 1 1 1 1 Total 5 2 3 Deep-water fringing islands: Nicobar Islands, P. Simeulue, P. Lasia India: Nicobar Islands Katchall 8°00' 93°22' 2 Little Nicobar 7°19' 93°41' 3 Great Nicobar 6°59' 93°46' 3 Total 8 3 5 Indonesia: west of Sumatra Simeulue Lasia 2°39' 2° 10' 96°08' 96°38' 13 2 9 1 4 1 Total 15 10 5 FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 189 Appendix 7: Head and Body Length (mm) in Fringing-Island Samples of Macacafas- cicularis Compared with Head and Body Length in Core-Area Reference Samples Collected at Similar Latitudes1 (cf. Table 9; Figs. 11, 12) Fringing-island samples Core-area sample statistics FI > CA Latitude3 Sample statistics > Island2 (°N) N Mean ± SD4 Extremes N Mean ± SD CA5 FI 6 pi A. Adult females Shallow-water islands West of Sumatra Mursala 1.67 1 390 18 407.3 ± 35.19 X >0.20 West of Isthmus of Kra and Malay Peninsula Zadetkyi 9.97 1 470 18 432.5 ± 52.83 X >0.20 Na Ka Yai 8.20 1 418 12 405.4 ± 34.97 X >0.20 Rang Yai 7.95 1 375 11 403.2 ± 35.76 X > 0.20 Phi Phi Don 7.75 2 420.5 415^26 10 394.0 ± 19.77 X > 0.10 Tarutao 6.58 2 414.5 379^150 19 393.4 ± 14.38 X > 0.20 Pinang [1] 5.47 1 340 15 399.1 ± 20.66 X 0.20 East of Sumatra Singapore 1.42 1 381 16 411.4 ± 35.26 X > 0.20 Karimun 1.06 2 335.0 315-355 17 411.9 ± 34.21 X < 0.05* Kundur 0.88 1 348 17 411.9 ± 34.21 X 0.20 Bulan 0.96 1 339 17 411.9 ± 34.21 X 0.20 Batam 1.12 2 382.0 379-385 17 411.9 ± 34.21 X > 0.20 Nguwal 0.65 1 375 18 416.8 ± 35.36 X >0.20 Bintan 1.15 2 353.5 320-387 17 411.9 ± 34.21 X >0.10 East of Isthmus of Kra and Malay Peninsula Phangan 9.75 2 395.5 391^00 18 432.5 ± 52.83 X > 0.20 Pinang [2] 5.73 1 395 19 396.3 ± 19.38 X >0.20 Tioman 2.80 6 399.3 ± 26.71 365^435 16 410.9 ± 36.29 X >0.20 South of Indochinese Peninsula Khram Yai8 12.70 4 418.8 ± 11.09 410-435 15 452.3 ± 50.22 X > 0.20 Kut 11.67 2 401.0 392^110 17 448.4 ± 48.54 X 0.20 Con Son 8.72 1 380 12 405.4 ± 34.97 X >0.20 West of Borneo Subi-kecil 3.03 1 393 19 406.1 ± 35.57 X >0.20 Uwi 1.10 1 394 17 411.9 ± 34.21 X > 0.20 South of Borneo Karimunjawa -5.85 2 443.5 417-470 12 434.5 ± 32.08 X >0.20 Bawean -5.80 2 418.0 398-438 12 434.5 ± 32.08 X > 0.20 Bali9 -8.25 4 430.8 ± 7.80 422-441 11 435.1 ± 33.30 X > 0.10 Northeast of Borneo Banggi 7.15 1 413 10 394.0 ± 19.77 X > 0.20 Deep-water islands West of Sumatra Simeulue10 2.66 4 398.8 ± 50.72 325^40 16 410.9 ± 36.29 X > 0.20 Nias 1.22 3 372.0 ± 19.16 350-385 16 411.4 ± 35.26 X > 0.05 Lesser Sunda Islands Sumbawa9 -8.56 2 415.0 390-440 11 435.1 ± 33.30 X >0.20 Flores9 -8.59 2 385.0 360-410 11 435.1 ± 33.30 X >0.10 Sumba9 -9.70 2 428.0 406-450 11 435.1 ± 33.30 X >0.20 Timor9 -9.57 1 490 11 435.1 ± 33.30 X 0.20 190 FIELDIANA: ZOOLOGY Appendix 7: Continued Fringing-island samples Core-area sample Latitude3 Sample statistics statistics FI > CA > Island2 (°N) N Mean ± SD4 Extremes N Mean ± SD CA5 FI 6 pi Philippines and Maratua Palawan 9.67 1 420 18 432.5 ± 52.83 X >0.20 Busuanga 12.07 1 420 18 445.7 ± 48.46 X >0.20 Luzon11 16.44 1 420 14 447.1 ± 52.16 X >0.20 Negros 9.42 5 425.8 ± 23.88 400-462 18 432.5 ± 52.82 X >0.20 Mindanao 7.12 5 438.6 ± 17.24 420-465 10 394.0 ± 19.77 X < 0.01** Tawitawi 5.17 1 394 16 398.5 ± 20.04 X >0.20 Maratua 2.25 1 420 B. Adult males 16 410.9 ± 36.29 X >0.20 Shallow-water islands West of Sumatra Tuangku 2.22 2 424.0 423^25 20 456.1 ± 44.97 X >0.20 Mursala 1.67 2 442.5 440-445 18 453.7 ± 44.74 X > 0.20 Tanahmasa -0.18 1 435 15 482.9 ± 49.87 X >0.20 Tanahbala -0.43 1 415 20 484.8 ± 43.38 X 0.20 West of Isthmus of Kra and Malay Peninsula Kadan12 12.50 1 520 9 477.6 ± 73.41 X >0.20 YaoNoi 8.12 1 440 19 457.1 ± 29.93 X >0.20 Phi Phi Don 7.75 1 450 19 457.1 ± 29.93 X >0.20 Talibong 7.25 2 476.0 470-482 19 457.1 ± 29.93 X >0.20 Tarutao 6.58 4 458.8 ± 16.52 440-480 19 457.1 ± 29.93 X >0.20 Butang 6.53 1 403 19 457.1 ± 29.93 X 0.10 Langkawi 6.37 1 430 19 457.1 ± 29.93 X >0.20 Burau 6.35 1 440 19 457.1 ± 29.93 X >0.20 Pinang[l] 5.47 5 403.4 ± 18.13 37^423 18 454.3 ± 28.06 X 0.002** Pintu Gedong 2.92 2 408.0 405^11 16 439.9 ± 40.62 X >0.20 East of Sumatra Bengkalis 1.45 1 445 16 450.1 ± 45.21 X >0.20 Padang 1.37 1 412 16 450.1 ± 45.21 X >0.20 Singapore 1.42 2 451.0 432-470 16 450.1 ± 45.21 X >0.20 Karimun 1.06 4 426.2 ± 34.37 392^74 20 449.4 ± 41.78 X >0.20 Kundur 0.88 1 483 19 450.3 ± 42.69 X >0.20 Durian 0.73 1 430 15 472.0 ± 44.10 X > 0.20 Sugi 0.82 1 420 16 459.6 ± 38.48 X >0.20 Bulan 0.96 1 472 20 449.4 ± 41.78 X >0.20 Galang 0.75 1 425 17 466.1 ± 45.88 X > 0.20 Bintan 1.15 4 449.0 ± 30.99 410-485 14 453.8 ± 47.23 X >0.20 Mapur 1.00 1 395 19 450.3 ± 42.69 X >0.20 Bangka -2.03 3 420.0 ± 32.23 383^*42 17 488.1 ± 46.40 X 0.02* East of Isthmus of Kra and Malay Peninsuh i Samui 9.50 2 430.0 423-431 20 465.8 ± 48.41 X >0.20 Pinang [2] 5.73 1 443 18 454.3 ± 28.06 X >0.20 Tioman 2.80 3 452.3 ± 18.45 432-468 19 444.4 ± 39.73 X >0.20 Acheh 2.67 1 498 20 445.7 ± 39.09 X >0.20 Pemanggil 2.58 1 415 20 445.7 ± 39.09 X >0.20 Aur 2.45 1 427 18 444.2 ± 40.68 X >0.20 Tinggi 2.30 1 538 17 446.2 ± 40.99 X 0.20 South of Indochinese Peninsula KhramYai12 12.70 5 452.0 ± 17.18 425^165 9 477.6 ± 72.28 X >0.20 Kut12 11.67 3 441.3 ± 19.50 419^*55 9 477.6 ± 72.28 X >0.20 PhuQuoc12 10.22 1 435 9 477.6 ± 72.28 X >0.20 Con Son 8.72 3 454.0 ± 23.30 435-480 19 457.1 ± 29.93 X >0.20 Ba 8.65 2 443.5 440-447 19 457.1 ± 29.93 X >0.20 FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 191 Appendix 7: Continued Fringing-island samples Core-area sample statistics Sample statistics Latitude3 FI > CA > Island2 (°N) N Mean ± SD4 Extremes N Mean ± SD CAS FI6 P7 West of Borneo Siantan 3.13 2 450.0 440-460 18 440.5 ± 38.25 X >0.20 Laut 4.72 1 457 19 453.7 ± 23.98 X >0.20 Natuna Besar 3.78 1 445 18 440.5 ± 38.25 X >0.20 Lagong 3.60 1 419 18 440.5 ± 38.25 X > 0.20 Subi-kecil 3.03 1 453 20 441.7 ± 36.36 X > 0.20 Benua 0.95 1 432 16 459.6 ± 38.48 X >0.20 Karimata -1.60 2 460.0 460-460 17 488.1 ± 46.40 X >0.20 Serutu -1.72 1 464 17 488.1 ± 46.40 X >0.20 Belitung -2.93 2 467.5 450-485 17 510.5 ± 35.46 X >0.10 South of Borneo Karimunjawa -5.85 1 501 15 495.3 ± 46.66 X >0.20 Bawean -5.80 1 472 15 495.3 ± 46.66 X >0.20 Matasiri -4.78 1 435 20 501.4 ± 41.32 X >0.20 Bali9 -8.25 6 464.7 ±29.19 425-495 19 512.7 ± 46.28 X < 0.05* Northeast of Borneo Banggi 7.15 2 505.0 485-525 19 457.1 ± 29.93 X > 0.05 Deep-water islands West of Sumatra Katchall 8.00 1 525 19 457.1 ± 29.93 Little Nicobar 7.32 2 490.0 475-505 19 457.1 ± 29.93 X > 0.20 Simeulue 2.66 6 478.3 ± 14.38 460-495 20 445.7 ± 39.09 X < 0.05* Lasia 2.17 1 470 20 456.1 ± 44.97 X >0.20 Nias 1.22 7 455.1 ± 21.45 430-490 17 450.4 ± 43.79 X >0.20 Lesser Sunda Islands Penida9 -8.75 1 458 19 512.7 ± 46.28 X >0.20 Lombok9 -8.48 2 450.0 430-470 19 512.7 ± 46.28 X > 0.05 Sumbawa9 -8.56 3 480.0 ± 10.00 470-490 19 512.7 ± 46.28 X > 0.10 Flores9 -8.59 1 423 19 512.7 ± 46.28 X 0.10 Sumba9 -9.70 1 452 19 512.7 ± 46.28 X > 0.20 Philippines and Maratua Balabac 7.90 1 450 19 457.1 ± 29.93 X > 0.20 Palawan 9.67 4 470.5 ± 49.10 410-530 20 465.8 ± 48.41 X > 0.20 Culion12 11.85 2 479.5 463^196 9 477.6 ± 72.28 X > 0.20 Mindoro12 13.17 3 482.0 ± 35.38 450-520 9 477.6 ± 72.28 X > 0.20 Luzon12 16.44 3 501.7 ± 2.89 500-505 9 477.6 ± 72.28 X > 0.20 Negros 9.42 2 502.5 466-539 20 465.8 ± 48.41 X > 0.20 Mindanao 7.12 5 486.6 ± 24.45 445-508 19 457.1 ± 29.93 X > 0.05 Tawitawi 5.17 1 386 20 453.4 + 26.74 X 0.10 Maratua 2.25 1 440 17 446.2 ± 40.99 X > 0.20 1 Mean latitude of core-area reference sample equals mean latitude of respective fringing-island sample; preferred size of core-area reference sample is 10-20. 2 For details, see Appendix 2 and Fooden (1991, p. 34). 3 Mean latitude of fringing-island sample localities (cf. Figs. 11, 12); negative numbers indicate latitudes south of the equator. 4 Standard deviation specified where n > 2. 5 Fringing-island (FI) sample mean exceeds core-area (CA) reference sample mean. 6 Core-area reference sample mean exceeds fringing-island sample mean. 7 Mann- Whitney [/-test, two-tailed; * P < 0.05, ** P < 0.01. 8 For measurement of zrc 4-012, cf. Kloss (1919c, p. 349) and Wetizel et al. (1988, p. 105). 9 Core-area reference samples consist of all measured female or male specimens collected in Java. 10 Excludes aberrantly large female (rmnh Coll. No. 1161, HB = 505) included in Figure 27. 11 Core-area reference sample consists of all measured female specimens collected north of 12.00°N. 12 Core-area reference samples consist of all measured male specimens collected north of 5.70°N. 192 FIELDIANA: ZOOLOGY Appendix 8: Tail Length (mm) in I ringing- Island Samples of Macaca fascicular is Compared with Tail Length in Core-Area Reference Samples Collected at Similar Latitudes1 (cf. Table 11; Figs. 15, 16) Fringing-islands samples Core-area sample statistics FI> CA CA5 FT Lati- tude3 (°N) Sample statistics > Island2 N Mean ± SD4 Extremes N Mean ± SD i jrr A. Adult females Shallow-water islands West of Sumatra Mursala 1.67 1 545 18 513.2 ± 62.30 X >0.20 West of Isthmus of Kra and Malay Peninsula Zadetkyi 9.97 1 420 NaKaYai 8.20 1 430 RangYai 7.95 1 420 Phi Phi Don 7.75 2 540.0 Tarutao 6.58 2 508.5 Pinang[l] 5.47 1 464 528-552 505-512 18 12 11 10 19 15 489.6 ± 35.59 495.3 ± 34.50 495.8 ± 36.14 501.9 ± 31.61 496.0 ± 28.38 495.3 ± 29.79 X X X X X X 0.20 0.20 0.20 0.20 >0.20 >0.20 East of Sumatra Singapore Karimun Kundur Bulan Batam Nguwal Bintan 1.42 1.06 0.88 0.96 1.12 0.65 1.15 1 2 1 1 2 1 3 445 423.5 475 503 470.5 505 451.0 ± 8.57 412-435 456-485 420-503 16 17 17 17 17 18 17 516.4 ± 65.06 517.5 ± 63.15 517.5 ± 63.15 517.5 ± 63.15 517.5 ± 63.15 527.5 ± 36.84 517.5 ± 63.15 X X X X X X X >0.20 0.05 >0.20 >0.20 0.10 >0.20 0.05 East of Isthmus of Kra and Malay Peninsula Phangan 9.75 2 426.5 Pinang[2] 5.73 1 550 Tioman1 2.80 5 418.2 ± 23.29 411^42 395^50 18 19 16 489.6 ± 35.59 496.8 ± 31.12 512.2 ± 66.43 X X X 0.05 0.20 0.002** South of Indochinese Peninsula Khram Yai Kut Con Son 12.70 11.67 8.72 4 2 1 473.8 ± 6.29 442.0 470 465^180 432^52 15 17 12 436.7 ± 57.74 446.2 ± 60.83 495.3 ± 34.50 X X X >0.20 >0.20 >0.20 West of Borneo Subi-kecil Uwi 3.03 1.10 1 1 465 514 19 17 509.6 ± 62.09 517.5 ± 63.15 X X >0.20 >0.20 South of Borneo Karimunjawa Bawean Bali8 -5.85 -5.80 -8.25 2 3 4 471.0 526.7 ± 48.05 441.5 ± 46.77 467^175 475-570 375-480 12 12 11 475.7 ± 52.68 475.7 ± 52.68 468.6 ± 52.37 X X X >0.20 0.20 >0.20 Northeast of Borneo Banggi 7.15 1 492 10 501.9 ± 31.61 X >0.20 Deep-water islands West of Sumatra Simeulue9 Nias 2.65 1.22 4 3 412.5 ± 53.31 427.0 ±21.28 345^75 400-450 16 16 512.2 ± 66.43 516.4 ± 65.06 X X < 0.02* 0.02* Lesser Sunda Islands Sumbawa8 Flores8 Sumba8 Timor8 -8.56 -8.59 -9.70 -9.57 2 2 2 1 430 417.5 406.0 345 400-460 390-445 404-408 11 11 11 11 468.6 ± 52.37 468.6 ± 52.37 468.6 ± 52.37 468.6 ± 52.37 X X X X >0.20 0.20 >0.10 >0.20 FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 193 Appendix 8: Continued Fringing-islands samples ^ |irn_'irii.i i»mi\!.i Lati- tude3 (°N) Sample statistics N statistics Mean ± SD FI> CA5 CA FI > Island2 N Mean ± SD4 Extremes 6 pi Philippines and Maratua Busuanga 12.07 1 530 18 443.6 ± 60.01 X >0.20 Luzon10 16.44 2 547.5 515-580 14 430.4 ± 58.58 X 0.05 Negros 9.42 5 491.8 ± 41.62 455-538 18 489.6 ± 35.59 X > 0.20 Mindanao 7.12 5 510.8 ± 56.31 445-564 10 501.9 ± 31.61 X > 0.20 Tawitawi 5.17 1 504 16 496.4 ± 29.12 X > 0.20 Maratua 2.25 1 530 B. Adult males 16 512.2 ± 66.43 X > 0.20 Shallow-water islands West of Sumatra Tuangku 2.22 2 505.0 480-530 19 572.5 ± 36.65 X > 0.05 Mursala 1.67 1 500 18 573.2 ± 37.59 X 0.20 Tanahmasa -0.18 1 570 15 589.9 ± 48.71 X > 0.20 Tanahbala -0.43 1 550 20 585.6 ± 44.51 X > 0.20 West of Isthmus of Kra and Malay Peninsula Kadan" 12.50 1 495 9 550.7 ± 59.48 X >0.20 Yao Noi 8.12 1 530 20 563.8 ± 48.52 X > 0.20 Phi Phi Don 7.75 1 570 20 563.8 ± 48.52 X > 0.20 Talibong 7.25 2 557.5 556-559 20 563.8 ± 48.52 X > 0.20 Tarutao 6.58 4 566.3 ± 16.01 550-580 20 563.8 ± 48.52 X > 0.20 Langkawi 6.37 1 570 20 563.8 ± 48.52 X >0.20 Burau 6.35 1 525 20 563.8 ± 48.52 X >0.20 Pinang [1] 5.47 5 519.4 ± 17.57 491-539 19 560.1 ± 46.78 X 0.05 Pintu Gedong 2.92 2 505.0 495-515 15 570.7 ± 38.82 X < 0.05* East of Sumatra Bengkalis 1.45 1 495 16 576.3 ± 37.44 X 0.20 Padang 1.37 1 470 16 576.3 ± 37.44 X 0.20 Singapore 1.42 2 565.0 533-597 16 576.3 ± 37.44 X > 0.20 Karimun 1.06 4 506.5 ± 45.18 472-572 20 580.3 ± 45.16 X 0.02* Durian 0.73 1 590 15 586.2 ± 51.04 X >0.20 Sugi 0.82 1 522 16 575.7 ± 46.85 X >0.20 Bulan 0.96 1 530 20 580.3 ± 45.16 X >0.20 Galang 0.75 1 580 17 581.2 ± 50.80 X >0.20 Bintan 1.15 4 500.8 ± 19.47 485-525 14 574.3 ± 37.64 X < 0.01** Mapur 1.00 1 535 19 578.0 ± 45.23 X > 0.20 Bangka -2.03 3 480.7 ± 42.85 435-520 17 586.2 ± 45.27 X < 0.01** East of Isthmus of Kra and Malay Peninsula Samui 9.50 2 480.5 459-502 21 562.4 ± 47.71 X < 0.05* Pinang [2] 5.73 1 620 19 560.1 ± 46.78 X 0.20 Tioman 2.80 3 483.7 ± 14.01 470-498 18 568.2 ± 37.25 X 0.002** Acheh 2.67 1 698 19 567.5 ± 36.33 X 0.10 Pemanggil 2.58 1 544 19 567.5 ± 36.33 X >0.20 Aur 2.45 1 575 17 565.3 ± 32.74 X >0.20 South of Indochinese Peninsula Khram Yai11 12.70 5 532.0 ± 18.24 510-550 9 550.7 ± 59.48 X >0.20 Kut" 11.67 3 484.3 ± 1.15 483^85 9 550.7 ± 59.48 X > 0.20 PhuQuoc11 10.22 1 445 9 550.7 ± 59.48 X 0.20 Con Son 8.72 3 509.0 ± 27.22 489-540 20 563.8 ± 48.52 X <0.10 Ba 8.65 2 530.0 500-560 20 563.8 ± 48.52 X > 0.20 West of Borneo Siantan 3.13 2 542.5 520-565 17 575.6 ± 39.07 X >0.20 Laut 4.72 1 533 20 567.9 ± 45.66 X > 0.20 194 FIELDIANA: ZOOLOGY Appendix 8: Continued Fringing- island samples Lati- tude3 (TV) Core-area sample Sample statistics statistics FI> CA5 CA FI > 1 Island2 N Mean ± SD4 Extremes N Mean ± SD P1 Natuna Besar 3.78 1 480 17 575.6 ± 39.07 X 0.20 Lagong 3.60 1 559 17 575.6 ± 39.07 X > 0.20 Subi-kecil 3.03 1 500 19 572.4 ± 38.98 X 0.10 Benua 0.95 1 559 16 575.7 ± 46.85 X > 0.20 Karimata -1.60 2 642.5 630-655 17 586.2 ± 45.27 X > 0.10 Serutu -1.72 1 591 17 586.2 ± 45.27 X > 0.20 Belitung -2.93 2 462.5 445^180 17 577.0 ± 35.28 X 0.02* South of Borneo Karimunjawa -5.85 1 544 14 570.5 ± 74.78 X >0.20 Bawean -5.80 2 580.0 580-580 14 570.5 ± 74.78 X > 0.20 Matasiri -4.78 1 540 19 555.6 ± 47.15 X > 0.20 Bali8 -8.25 6 517.8 ± 23.49 485-546 18 556.5 ± 84.86 X > 0.10 Northeast of Borneo Banggi 7.15 2 607.5 595-620 20 563.8 ± 48.52 X > 0.20 Deep-water islands West of Sumatra Katchall 8.00 1 605 20 563.8 ± 48.52 X > 0.20 Little Nicobar 7.32 2 565.0 550-580 20 563.8 ± 48.52 X > 0.20 Simeulue 2.66 6 465.8 ± 25.18 435-490 19 567.5 ± 36.33 X < 0.002** Lasia 2.17 1 555 19 572.5 ± 36.65 X > 0.20 Nias 1.22 7 505.3 ± 28.99 475-560 17 575.6 ± 36.35 X < 0.002** Lesser Sunda Islands Penida8 -8.75 1 462 18 556.5 ± 84.86 X > 0.20 Lombok8 -8.48 2 447.5 445^50 18 556.5 ± 84.86 X 0.05 Sumbawa8 -8.56 3 506.7 ± 35.12 470-540 18 556.5 ± 84.86 X > 0.20 Flores8 -8.59 1 500 18 556.5 ± 84.86 X > 0.20 Sumba8 -9.70 1 503 18 556.5 ± 84.86 X > 0.20 Philippines and Maratua Balabac 7.90 2 527.0 510-544 20 563.8 ± 48.52 X > 0.20 Palawan 9.67 4 526.8 ± 34.19 487-560 21 562.4 ± 47.71 X > 0.10 Culion" 11.85 3 526.7 ± 27.23 496-548 9 550.7 ± 59.48 X > 0.20 Mindoro" 13.17 3 570.7 ± 16.17 552-580 9 550.7 ± 59.48 X > 0.20 Luzon" 16.44 6 561.2 ± 41.81 510-625 9 550.7 ± 59.48 X > 0.20 Leyte 11.08 2 540.0 490-590 9 550.7 ± 59.48 X > 0.20 Negros 9.42 3 572.3 ± 7.37 564-578 21 562.4 ± 47.71 X > 0.50 Mindanao 7.12 5 552.4 ± 38.25 510-600 20 563.8 ± 48.52 X > 0.20 Tawitawi 5.17 1 507 20 562.2 ± 46.59 X > 0.20 Maratua 2.25 1 575 16 565.6 ± 33.79 X > 0.20 1 Mean latitude of core-area reference sample equals mean latitude of respective fringing-island sample; preferred size of core-area reference sample is 10-20. Excluded from this table are bobtailed specimens zrc 4-127 (female, West Malaysia: P. Tioman, Telok Juara, T = 215 mm) and bm(nh) 1955.151 1 (male, Thailand: Ko Butang, T = 269 mm). 2 For details, see Appendix 2 and Fooden (1991, p. 34). 3 Mean latitude of fringing-island sample localities (cf. Figs. 15, 16); negative numbers indicate latitudes south of the equator. 4 Standard deviation specified where n > 2. 5 Fringing-island (FI) sample mean exceeds core-area (CA) reference sample mean. 6 Core-area reference sample mean exceeds fringing-island sample mean. 7 Mann- Whitney U-test, two-tailed; * P < 0.05, ** P < 0.01. 8 Core-area reference samples consist of all measured female or male specimens collected in Java. 9 Excludes aberrantly large female (rmnh Coll. No. 1 161, HB = 505, T = 440) included in Figure 27. 10 Core-area reference sample consists of all measured female specimens collected north of 12.00°N. 11 Core-area reference samples consist of all measured male specimens collected north of 5.70°N. FOODEN: SYSTEMATIC REVIEW OF MAC AC A FASCICULARIS 195 Appendix 9: Relative Tail Length (T/HB x 100) in Fringing-Island Samples of Macaca fascicularis Compared with Relative Tail Length in Core- Area Reference Samples Col- lected at Similar Latitudes1; Samples Include Both Sexes (cf. Table 11; Figs. 17, 18) Fringing-island samples Lati- Sample statistics Core-area sample statistics tude3 (°N) N Mean ± SD FI > CA5 CA FI4 > Island2 N Mean ± SD4 Extremes pi Shallow-water islands West of Sumatra Tuangku 2.22 2 119.1 112.9-125.3 19 133.4 ± 11.80 X >0.10 Mursala 1.67 2 126.7 113.6-139.7 18 124.2 ± 9.76 X >0.20 Tanahmasa -0.18 1 131.0 18 125.6 ± 9.83 X >0.20 Tanahbala -0.43 1 132.5 18 124.6 ± 10.43 X > 0.20 West of Isthmus of Kra and Malay Peninsula Kadan 12.50 1 95.2 12 98.0 ± 8.48 X > 0.20 Zadetkyi 9.97 1 89.4 12 104.1 ± 14.94 X > 0.20 Na Ka Yai 8.20 1 102.9 18 124.3 ± 13.39 X > 0.20 Yao Noi 8.12 1 120.5 18 124.3 ± 13.39 X > 0.20 Rang Yai 7.95 1 112.0 17 124.9 ± 13.54 X > 0.20 Phi Phi Don 7.75 3 127.8 ± 1.54 126.7-129.6 16 127.3 ± 9.92 X >0.20 Talibong 7.25 2 117.1 115.4-118.9 16 127.3 ± 9.92 X >0.10 Tarutao 6.58 6 123.5 ± 7.60 112.2-135.1 14 127.7 ± 9.90 X >0.20 Langkawi 6.37 1 132.6 13 127.9 ± 10.27 X >0.20 Burau 6.35 1 119.3 13 127.9 ± 10.27 X >0.20 Pinang [1] 5.47 6 130.3 ± 8.52 122.1-144.1 26 122.6 ± 9.79 X >0.10 Pintu Gedong 2.92 2 123.8 122.2-125.3 18 130.3 ± 17.96 X > 0.20 East of Sumatra Bengkalis 1.45 1 111.2 18 124.2 + 9.76 X > 0.20 Padang 1.37 1 114.1 18 124.2 ± 9.76 X >0.20 Singapore 1.42 3 122.4 ± 5.18 116.8-127.0 18 124.2 ± 9.76 X >0.20 Karimun 1.06 6 121.8 ± 11.93 101.7-136.5 19 124.9 ± 9.94 X >0.20 Kundur 0.88 1 136.5 19 125.2 ± 10.95 X >0.20 Durian 0.73 1 137.2 20 124.5 ± 9.82 X >0.20 Sugi 0.82 1 124.3 20 124.5 ± 9.82 X >0.20 Bulan 0.96 2 130.3 112.3-148.4 15 122.5 + 9.39 X > 0.20 Batam 1.12 2 123.2 118.4-128.0 19 124.9 ± 9.94 X >0.20 Galang 0.75 1 136.5 20 124.5 ± 9.82 X >0.20 Nguwal 0.65 1 134.7 20 124.5 ± 9.82 X > 0.20 Bintan 1.15 6 114.9 ± 8.77 108.2-131.3 19 124.9 ± 9.94 X < 0.05* Mapur 1.00 1 135.4 14 122.4 ± 9.73 X > 0.20 Bangka -2.03 3 115.0 ± 14.89 98.4-127.2 20 117.8 ± 6.84 X > 0.20 East of Isthmus of Kra and Malay Peninsula Phangan 9.75 2 107.9 102.8-113.0 14 106.9 ± 15.44 X >0.20 Samui 9.50 2 111.7 108.5-114.9 14 106.9 ± 15.44 X >0.20 Pinang [2] 5.73 2 139.6 139.2-140.0 24 123.1 ± 10.00 X < 0.05* Tioman1 2.80 8 106.2 ± 8.38 94.7-118.4 19 130.1 ± 17.49 X < 0.002** Acheh 2.67 1 140.2 18 130.2 ± 17.99 X >0.20 Pemanggil 2.58 1 131.1 16 136.2 ± 10.48 X >0.20 Aur 2.45 1 134.7 16 136.2 ± 10.48 X > 0.20 South of Indochinese Peninsula Khram Yai 12.70 9 115.8 ± 5.72 109.2-128.2 18 97.1 ± 10.97 X < 0.002** Kut 11.67 5 110.1 ± 4.85 105.4-115.3 17 101.3 ± 14.52 X >0.05 Phu Quoc 10.22 1 102.3 14 106.9 ± 15.03 X >0.20 Con Son 8.72 4 115.0 ± 6.15 109.4-123.7 18 124.3 ± 13.39 X >0.05 Ba 8.65 2 119.6 111.9-127.3 18 124.3 ± 13.39 X > 0.20 West of Borneo Siantan 3.13 2 120.7 113.0-128.4 19 129.3 ± 17.39 X >0.20 Laut 4.72 1 116.6 12 127.7 ± 8.64 X >0.20 196 FIELDIANA: ZOOLOGY Appendix 9: Continued Fringing-island samples Lati- tude3 . (°N) Core- Sample statistics statistics FI> CAS CA FP > Island2 N Mean ± SD4 Extremes N Mean ± SD F Natuna Besar 3.78 1 107.9 15 125.4 ± 16.62 X >0.20 Lagong 3.60 1 133.4 20 127.0 ± 15.77 X >0.20 Subi-kecil 3.03 2 114.3 110.4-118.3 19 130.1 ± 17.49 X >0.10 Uwi 1.10 1 130.5 19 124.9 ± 9.94 X >0.20 Benua 0.95 1 129.4 15 122.5 ± 9.39 X >0.20 Karimata -1.60 2 139.7 137.0-142.4 22 117.9 ± 7.03 X < 0.05* Serutu -1.72 1 127.4 21 117.4 ± 6.86 X >0.10 Belitung -2.93 2 98.9 98.9-99.0 18 117.5 ± 6.29 X 0.02* South of Borneo Karimunjawa -5.85 3 107.2 ± 5.59 101.1-112.0 20 114.1 ± 20.44 X >0.20 Bawean -5.80 3 129.4 ± 11.96 122.1-143.2 20 114.1 ± 20.44 X >0.20 Matasiri -4.78 1 124.1 16 111.9 ± 8.34 X >0.20 Bali8 -8.25 10 108.1 ± 9.98 87.2-125.8 29 109.5 ± 19.52 X >0.90 Northeast of Borneo Banggi 7.15 3 120.0 ± 2.41 118.1-122.7 16 127.3 ± 9.92 X >0.10 Deep-water islands West of Sumatra Katchall 8.00 1 115.2 17 124.9 ± 13.54 X >0.20 Little Nicobar 7.32 2 115.3 114.9-115.8 16 127.3 ± 9.92 X >0.10 Simeulue9 2.66 10 99.9 ± 5.99 89.9-108.0 18 130.2 ± 17.99 X < 0.002** Lasia 2.17 1 118.1 20 132.7 ± 11.98 X >0.20 Nias 1.22 10 112.3 ± 7.40 103.9-128.6 18 124.2 ± 9.76 X < 0.002** Lesser Sunda Islands8 Penida -8.75 1 100.9 29 109.5 ± 19.52 X >0.40 Lombok -8.48 2 99.6 95.7-103.5 29 109.5 ± 19.52 X >0.20 Sumbawa -8.56 5 104.8 ± 6.22 97.9-114.9 29 109.5 ± 19.52 X >0.40 Flores -8.59 3 112.3 ± 15.13 95.1-123.6 29 109.5 ± 19.52 X >0.50 Sumba -9.70 3 100.5 ± 10.34 90.7-111.3 29 109.5 ± 19.52 X >0.20 Timor -9.57 1 70.4 29 109.5 ± 19.52 X >0.10 Philippines and Maratua Balabac 7.90 1 113.3 16 127.3 ± 9.92 X >0.20 Palawan 9.67 4 112.5 ± 9.42 104.3-124.4 14 106.9 ± 15.44 X >0.20 Culion 11.85 2 107.6 107.1-108.1 18 101.2 ± 13.35 X >0.20 Busuanga 12.07 1 126.2 20 99.8 ± 13.62 X 0.20 Mindoro 13.17 3 118.8 ± 9.02 111.5-128.9 18 97.1 ± 10.97 X 0.002** Luzon10 16.44 4 120.6 ± 15.44 101.0-138.1 19 97.2 ± 10.68 X < 0.01** Negros 9.42 7 115.5 ± 7.75 105.1-126.4 14 106.9 ± 15.44 X >0.10 Mindanao 7.12 10 115.3 ± 12.00 95.7-130.3 16 127.3 ± 9.92 X < 0.05* Tawitawi 5.17 2 129.6 127.9-131.3 29 123.4 ± 9.61 X >0.20 Maratua 2.25 2 128.4 126.2-130.7 19 133.4 ± 11.80 X >0.20 1 Mean latitude of core-area reference sample equals mean latitude of respective fringing-island sample; preferred size of core-area reference sample is 10-20. Excluded from this table are bobtailed specimens zrc 4-127 (female, West Malaysia: P. Tioman, Telok Juara, T = 215, HB = 409, T/HB x 100 = 52.6) and bm(nh) 1955.151 1 (male, Thailand: Ko Butang, T = 269, HB = 403, T/HB x 100 = 66.7). 2 For details, see Appendix 2 and Fooden (1991, p. 34). 3 Mean latitude of fringing-island sample localities (cf. Figs. 17, 18) negative numbers indicate latitudes south of the equator. 4 Standard deviation specified where n > 2. 5 Fringing-island (FI) sample mean exceeds core-area (CA) reference sample mean. 6 Core-area reference sample mean exceeds fringing-island sample mean. 7 Mann-Whitney [/-test, two-tailed; * P < 0.05, ** P < 0.01. 8 Core-area reference samples consist of all measured specimens collected in Java. 9 Excludes aberrantly large female (rmnh Coll. No. 1 161, HB = 505, T = 440, T/HB x 100 = 87.1) included in Figure 27. 10 Core-area reference sample consists of all measured specimens collected north of 10.00°N. FOODEN: SYSTEMATIC REVIEW OF MACACA FASCICULARIS 197 Appendix 10: Blood-Protein Allele Frequencies at Polymorphic Loci in 11 Geographic Samples of Macaca fascicularis Sample areas Thailand North of Isthmus of Kra South Central and of West south- South- Isthmus Malay- Sum- Philip- Loci2 and east west of Kra sia Sumatra Java Bali Lombok bawa Timor pines4 alleles3 (121) (124) (33) (140) (276) (222) (136) (35) (81) (7) (142) Plasma proteins Alb A 0.840 0.201 0.518 0.939 0.903 1.000 1.000 1.000 1.000 0.929 1.000 B 0.160 0.799 0.482 0.061 0.081 0 0 0 0 0.071 0 C 0 0 0 0 0.008 0 0 0 0 0 0 D 0 0 0 0 0.008 0 0 0 0 0 0 Alp A 1.000 0.997 1.000 1.000 1.000 1.000 0.989 1.000 1.000 1.000 0.972 B 0 0 0 0 0 0 0.011 0 0 0 0.028 * 0 0.003 0 0 0 0 0 0 0 0 0 ChEs 1 1.000 1.000 1.000 1.000 1.000 0.993 1.000 1.000 1.000 1.000 1.000 5 0 0 0 0 0 0.007 0 0 0 0 0 Pi5 A 0 0 0 0 0.008 0.005 0 0 0 0 0 B 0.182 0.011 0.954 0.543 0.816 0.686 0.221 0 0.790 0.643 0.919 C 0.818 0.883 0.046 0.457 0.176 0.309 0.779 1.000 0.210 0.357 0.081 D 0 0.106 0 0 0 0 0 0 0 0 0 TBPA F 0.894 0.978 1.000 0.846 0.935 0.973 1.000 1.000 0.519 1.000 0.944 S 0.106 0.022 0 0.154 0.065 0.027 0 0 0.482 0 0.056 Tf6 * 0.034 0.089 0 0 0 0 0 0 0 0 0 Bl 0 0 0 0.011 0.016 0 0 0 0 0 0 B2 0 0 0.067 0 0.004 0.023 0 0 0 0 0 CI 0 0 0 0 0.002 0 0 0 0 0 0 C2 0 0 0 0.057 0.201 0.111 0.004 0 0 0 0.007 C3 0.039 0.237 0 0.043 0.024 0.097 0 0 0.605 0.571 0 * 0.191 0.157 0.033 0 0 0 0 0 0 0 0 * 0 0.004 0 0 0 0 0 0 0 0 0 * 0.005 0 0 0 0 0 0 0 0 0 0 Dl 0.015 0.085 0.617 0.736 0.616 0.579 0.004 0.029 0.395 0.429 0.993 D2 0.324 0.127 0 0.025 0 0.007 0 0 0 0 0 * 0.069 0 0 0 0 0 0 0 0 0 0 El 0.025 0 0.017 0.047 0.002 0 0 0 0 0 0 E2 0.172 0.237 0 0 0.007 0.011 0 0 0 0 0 * 0.025 0.008 0 0 0 0 0 0 0 0 0 Fl 0 0 0 0.004 0 0.016 0.011 0.143 0 0 0 Gl 0.088 0.017 0.200 0.036 0.011 0.005 0 0 0 0 0 G2 0.005 0.013 0.050 0.004 0.016 0.138 0.982 0.829 0 0 0 HI 0.010 0.025 0.017 0.036 0.101 0.014 0 0 0 0 0 H2 0 0 0 0.004 0 0 0 0 0 0 0 Erythrocyte proteins Acp A 0.969 0.993 0.971 0.964 0.996 1.000 1.000 1.000 1.000 1.000 1.000 C 0.031 0.007 0.029 0.036 0.004 0 0 0 0 0 0 ADA 2 1.000 0.915 1.000 1.000 1.000 1.000 1.000 1.000 1.000 1.000 0.993 5 0 0 0 0 0 0 0 0 0 0 0.007 * 0 0.085 0 0 0 0 0 0 0 0 0 AK 1 0.962 1.000 0.972 0.979 1.000 1.000 1.000 1.000 1.000 1.000 0.919 2 0.038 0 0 0.004 0 0 0 0 0 0 0 3 0 0 0 0 0 0 0 0 0 0 0.081 4 0 0 0 0.011 0 0 0 0 0 0 0 5 0 0 0.028 0.007 0 0 0 0 0 0 0 CA-I a 0.991 0.951 1.000 0.964 0.975 0.993 1.000 1.000 1.000 1.000 1.000 b 0.010 0.049 0 0.021 0.004 0.002 0 0 0 0 0 c 0 0 0 0.014 0.021 0.005 0 0 0 0 0 198 FIELDIANA: ZOOLOGY Appendix 10: Continued Sample areas Thailam I North of Isthmus of Kra South Central and of West south- South- Isthmus Malay- Suni- Philip- Loci2 and east west of Kra sia Sumatra Java Bali Lombok bawa Timor pines4 alleles3 (121) (124) (33) (140) (276) (222) (136) (35) (81) (7) (142) CA-II a 0.612 0.536 0.177 0.392 0.463 0.164 0 0 0.111 0 0.609 b 0.388 0.464 0.823 0.608 0.537 0.836 1.000 1.000 0.889 1.000 0.391 CellEs 1 1.000 1.000 1.000 0.989 0.996 0.968 1.000 1.000 0.803 0.857 0.954 3 0 0 0 0.011 0.004 0.032 0 0 0.074 0 0 4 0 0 0 0 0 0 0 0 0.124 0.143 0.046 Dia A 0.247 0.477 0.071 0.082 0.010 0.005 0 0 0 0 0 C 0.715 0.439 0.929 0.914 0.990 0.995 1.000 1.000 1.000 1.000 1.000 H 0 0 0 0.004 0 0 0 0 0 0 0 * 0.039 0.084 0 0 0 0 0 0 0 0 0 EsD 1 1.000 1.000 1.000 0.996 1.000 1.000 1.000 1.000 1.000 1.000 0.944 4 0 0 0 0.004 0 0 0 0 0 0 0 5 0 0 0 0 0 0 0 0 0 0 0.056 HbA-F 1 1.000 1.000 0.951 0.989 0.996 1.000 1.000 1.000 1.000 1.000 1.000 4 0 0 0.049 0.007 0.004 0 0 0 0 0 0 5 0 0 0 0.004 0 0 0 0 0 0 0 HbA-IF 0 0.991 1.000 0.332 0.378 0.129 0.694 0.989 0.971 0 0 0 2 0.010 0 0.668 0.622 0.872 0.306 0.011 0.029 1.000 1.000 1.000 HbB 1 1.000 1.000 1.000 1.000 1.000 1.000 0.967 1.000 1.000 1.000 1.000 6 0 0 0 0 0 0 0.033 0 0 0 0 IDH 1 0.474 0.394 0.513 0.800 0.902 0.648 0.154 0.571 1.000 0.857 0.993 2 0.526 0.606 0.487 0.193 0.087 0.350 0.846 0.429 0 0.143 0.007 3 0 0 0 0 0.004 0 0 0 0 0 0 4 0 0 0 0.007 0.007 0.002 0 0 0 0 0 LDHA 1 0.941 0.972 1.000 1.000 0.979 1.000 1.000 1.000 1.000 1.000 1.000 3 0.059 0.028 0 0 0.021 0 0 0 0 0 0 PGD A 0.983 0.929 0.971 0.921 0.904 0.986 1.000 1.000 1.000 1.000 1.000 C 0 0.071 0 0.079 0.096 0.012 0 0 0 0 0 I 0 0 0 0 0 0.002 0 0 0 0 0 * 0.017 0 0 0 0 0 0 0 0 0 0 * 0 0 0.029 0 0 0 0 0 0 0 0 PHI 1 1.000 0.829 1.000 0.986 0.977 1.000 0.985 0.971 1.000 1.000 0.813 5 0 0 0 0.011 0.023 0 0 0 0 0 0.187 13 0 0 0 0 0 0 0.015 0 0 0 0 14 0 0 0 0 0 0 0 0.029 0 0 0 19 0 0 0 0.004 0 0 0 0 0 0 0 * 0 0.171 0 0 0 0 0 0 0 0 0 1 Based on data provided by Dr. Y. Kawamoto (letter, 5 May 1992); cf. Tomiuk (1989a, p. 90) and Schmitt et al. (1990, p. 96). Sample size indicated by italicized figures in parentheses. 2 Abbreviations: Acp = acid phosphatase, ADA = adenosine deaminase, AK = adenylate kinase, Alb = albumin, Alp = alkaline phosphatase, CA = carbonic anhydrase, CellEs = esterase, ChEs = cholinesterase, Dia = NADH- diaphorase, EsD = esterase D, HbA = hemoglobin alpha-chain, HbB = hemoglobin beta-chain, IDH = isocitrate dehydrogenase, LDHA = lactate dehydrogenase-A, PGD = 6-phosphogluconate dehydrogenase, PHI = phosphohex- ose isomerase, Pi = protease inhibitor, TBPA = thyroxin-binding prealbumin, Tf = transferrin. 3 Asterisks (*) indicate electrophoretically distinct alleles that have not yet been designated by letter or number. 4 Island or islands unknown. 5 Cf. Tanaka (1991, p. 47) and Samilchuk and Sarsaniya (1994, p. 357). 6Cf. Hoopes(1984, p. 171). 7 Cf. Tomiuk (1989b, p. 96). 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