UNIVERSITY OF KANSAS miscellaneous MUSEUM OF NATURAL HISTORY publication No. 77 Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part II. Fossil Amphibians and Reptiles By H.-P. Schultze, L. Hunt, J. Chorn and A. M. Neuner UNIVERSITY OF KANSAS LAWRENCE 1985 December 3, 1985 Ernst Mayr Vbmy - v^ L. MMieum of Compam%v€ 7.ook)^ Harvard University UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY The University of Kansas Publications, Museum of Natural History, beginning with volume 1 in 1946, was discontinued with volume 20 in 1971. Shorter research papers formerly published in the above series are now published as Occasional Papers, Museum of Natural History. The Miscellaneous Publications, Museum of Natural History, began with number I in 1946. Longer research papers are published in that series. Monographs of the Museum of Natural History were initiated in 1970. All manuscripts are subjected to critical review by intra- and extramural specialists; final acceptance is at the discretion of the Director. Institutional libraries interested in exchanging publications may obtain the Occasional Papers and Miscellaneous Publications by addressing the Exchange Librarian, University of Kansas Library, Lawrence, Kansas 66045. Individuals may purchase separate numbers of all series. Prices for all publications of the Museum may be obtained from the Publications Secretary, Museum of Natural History, University of Kansas, Lawrence, Kansas 66045. The University of Kansas Museum of Natural History Miscellaneous Publication No. 77 December 3, 1985 /,l/f:i;l Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part II. Fossil Amphibians and Reptiles By H.-P. ScHULTZE, L. Hunt, J. Chorn and A. M. Neunep Museum of Natural History, University of Kansas Lawrence, Kansas 66045 U.S.A. The University of Kansas Lawrence 1985 University of Kansas Publications, Museum of Natural History Editor: Robert M. Mengel Managing Editor: Joseph T. Collins Miscellaneous Publication No. 77 i-iv; Pp. 1-66 Published December 3, 1985 MCZ LIBRARY JUL 2 3 2008 K .. ., vRD UNIVERSITY Museum of Natural History The University of Kansas Lawrence, Kansas 66045 U.S.A. Printed By University ok Kansas Printing Service Lawrence. Kansas CONTENTS INTRODUCTION 1 SHORT HISTORY OF THE COLLECTION 2 ACKNOWLEDGMENTS 4 CATALOGUE 5 Class Amphibia 5 Subclass Lepospondyli 5 Order Nectridea 5 Subclass Labyrinthodontia 5 Order Temnospondyli 5 Order Batrachosauria 7 Order Microsauria 8 Labyrinthodontia inc. sedis 9 Subclass Lissamphibia 9 Order Anura 9 Order Urodela 16 "Amphibia" 20 Cohort Amniota 20 Class "Reptilia" 20 Subclass Captorhinomorpha 20 Subclass Testudines 23 Order Casichelydia 23 Subclass Sauropsida 30 Superorder Lepidosauria 30 Order "Eosuchia" 30 Order Squamata 33 Superorder Archosauria 47 Order Thecodontia 47 Order Crocodylia 48 Order Pterosauria 48 Order Saurischia 49 Order Ornithischia 49 Superorder Euryapsida 50 Order Sauropterygia 50 Subclass Theropsida 57 Superorder Pelycosauria 57 LITERATURE CITED 58 APPENDIX: GENERA AND SPECIES LISTED 64 INTRODUCTION In 1982, part one (Fossil Fishes) of the Catalogue of Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History was published. Part two (Amphibi- ans and Reptiles) follows the same scheme. It includes type and figured fossil her- petological specimens regardless of whether or not they were relocated. Each cited speci- men is accompanied by the following infor- mation: currently used catalogue number and old number if cited in publication; name as published (the term "error" after a cited name indicates either misspelling in publica- tion or variance from spelling of name used in the catalogue); author and year of publica- tion (we were not able to cite all editions of text-books and popular books, however, an effort was made to cite the first edition in all cases); page of type description; figure; preserved anatomical part(s) of specimen; geologic stage, formation, member (see Zeller 1968); locality and collector (where known). Within the Cretaceous, the specific- ity of assignments to geologic stage varies for old localities depending on our knowl- edge of each locality. A locality can be assigned to stage if its location is well enough defined, because the areal distribu- tion of outcrops correlated with various stages is known. Within one species, the specimens are arranged first after their type category and second in order of the appear- ance of the figures within the publication. Every specimen is cited under the spe- cies to which it is assigned in the latest available publication, except where other- wise noted. Species are grouped according to the following classification: Class Amphibia Subclass Lepospondyli Order Nectridea Family Urocordylidae Subclass Labyrinthodontia Order Temnospondyli Suborder Rhachitomi Superfamily Trimerorhachoidea Family Saurerpetontidae Superfamily Eryopoidea Family Eryopidae Superfamily Dissorophoidea Family Dissorophidae Suborder Stereospondyli Superfamily Capitosauroidea Family Capitosauridae Order Batrachosauria Order Microsauria Suborder Tuditanomorpha Family Gymnarthridae Subclass Lissamphibia Order Anura Family Pelobatidae Family Bufonidae Family Ranidae Order Urodela Suborder Cryptobranchoidea Family Cryptobranchidae Family Ambystomatidae Cohort Amniota Class "Reptilia" (excluding Aves and Mammalia) Subclass Captorhinomorpha Suborder Captorhinoidea Family Captorhinidae Subclass Testudines Order Casichelydia Suborder Cryptodira Superfamily Chelonioidea Family Protostegidae Family Toxochelyidae Family Cheloniidae Superfamily Testudinoidea Family Chelydridae Family Emydidae Family Testudinidae Subclass Sauropsida Superorder Lepidosauria Order "Eosuchia" Family Petrolacosauridae Order Squamata Suborder Sauria Infraorder Iguania Family Iguanidae Infraorder Scincomorpha Family Scincidae Family Teiidae Infraorder Anguimorpha Superfamily Diploglossa Family Anguidae Superfamily Platynota MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Family Helodermatidae Family Mosasauridae Suborder Serpentes Suborder Amphisbaenia Family Amphisbaenidae Superorder Archosauria Order Thecodontia Suborder Pseudosuchia Infraorder Parasuchia Family Parasuchidae Order Crocodylia Suborder Mesosuchia Family Goniopholididae Order Pterosauria Suborder Pterodactyloidea Family Pteranodontidae Order Saurischia Suborder Sauropodomorpha Infraorder Sauropoda Family Camarasauridae Family Atlantosauridae Suborder Theropoda Infraorder Carnosauria Family Megalosauridae Order Ornithischia Suborder Ankylosauria Family Nodosauridae Suborder Ceratopsia Family Ceratopsidae Superorder Euryapsida Order Sauropterygia Suborder Plesiosauria Superfamily Plesiosauroidea Family Elasmosauridae Superfamily Pliosauroidea Family Polycotylidae Subclass Theropsida Order Pelycosauria Suborder Sphenacodontia Family Sphenacodontidae Suborder Edaphosauria Family Nitosauridae Family Edaphosauridae The classification follows Romer (1966) and Kuhn-Schnyder (1980) with the follow- ing changes: The Microsauria are excluded from the Lepospondyli because of their ver- tebral structure (Carroll and Gaskill 1978). The term Batrachosauria is used in the sense of Panchen (1977. 1980) replacing An- thracosauria. The terms "Reptilia" and "Archosauria" are used in the conventional way without including birds and mammals. The classification of Gaffney (1975) is adopted for the turtles. The order "Eosuchia" is retained for want of a better place to put Petrolacosaiinis (Peabody 1952, Reisz 1977). "Eosuchia" is not a monophyletic group (Reisz 1981). The clas- sification of Thecodontia is that of Chatter- jee (1982). thus the phytosaurs (Parasuchidae) have been placed within the order Pseudosuchia. SHORT HISTORY OF THE COLLECTION A short history of the fossil vertebrate collection as a whole is given in Part I (Schultze et al. 1982), therefore this discus- sion is restricted solely to the acquisition of fossil amphibians and reptiles. In this con- nection, we can distinguish three active peri- ods of publication, each connected with the collection of one of the following people: 1) S. W. Williston (1890-1902); 2) C. W. Hibbard (1928-1946); 3) F. E. Peabody (1948-1955). Before S. W. Williston joined the faculty of the University of Kansas, F. H. Snow was active in collecting and acquiring fossils for the University of Kansas. The fates of these early specimens cannot always be deter- mined. We do not know how many of the "41 saurians, 6 pterodactyles and 117 fishes" (Snow 1880) from the collecting parties of 1877 and 1878 are still in the collection; only a few specimens in our collection are definitely known to have been collected in those years. At an even earlier date the University of Kansas acquired an amphibian track way from the Upper Penn- sylvanian of eastern Kansas that was col- lected by B. F. Mudge in 1873 and is now on exhibit. The type specimen of the plesiosaur Trinacromerum anonymum was also col- lected by B. F. Mudge in 1873. The acquisi- tion date is unknown for these specimens, as it is for the oldest type specimen in the collection, Plesiosaurus giilo. The latter specimen belonged to the private collector Mr. J. Savage. Leavenworth, Kansas, when Cope (1872) reported on it at a meeting of the Academy of Natural Sciences, Phila- delphia. E. D. Cope (1886, 1889) described TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES another type in the collection, a series of dermal ossicles and a single phalangeal ele- ment of Geochehme orfhopygia, which he assigned to a new species of armadillo, Caryoderma snovianum. During his stay at the University of Kansas (1890-1902). S. W. WiUiston di- rected many active students and collectors: G. R. Allman, E. H. S. Bailey, J. W. Beede, Barnum Brown, E. C. Case, B. M. Dickin- son, L. L. Dyche, C. N. Gould, R. C. Gowell, J. B. Hatcher, B. H. Hill, G. Jewett, H. Kahl, V. L. Kellogg, W. C. Knight, W. T. Lee, W. N. Logan, H. T. Martin, T. Morse, T. R. Overton, A. T. Osborn, S. Prentice, E. S. Riggs, J. P. Sams, E. H. Sellards, E. E. Slosson, W. Snow, C. H. Sternberg, A. Stewart, E. P. West. They collected mostly in the Upper Cretaceous of western Kansas, and in the Upper Jurassic of Wyoming, but also in the Paleozoic of eastern Kansas. This was the period of most rapid growth of the fossil herpetological collection, especially that part from the Upper Cretaceous of western Kansas. Collecting slowed after Williston left in 1902, but the years 1910 and 1911 were notable exceptions with collecting in the Upper Cretaceous of western Kansas and in the Oligocene-Miocene of Colorado by large field parties (C. E. McClung and H. T. Martin with W. Bailey, C. Fairchild, R. E. Fox, W. Green, H. Gunthorp, J. Henshall, R. Kimball, Lindsay, P. McArdle, H. Mar- tin, R. L. Moodie, T. Morse, A. T. Osborn, G. Sternberg, E. H. Taylor). C. W. Hibbard's field activities (1928-1946) focused on the Pliocene in Meade County, western Kansas. He was chiefly interested in small mammals, but at the same time he and his field crews col- lected an immense amount of isolated am- phibian and reptile bones from deposits in western Kansas. His assistants were students and colleagues: B. P. Bagrowski, W. J. Baumgartner, M. Bear, C. D. Bunker, R. R. Camp, V. L. Carter, H. Cochran, L. Coghill, W. Dalquest, T. Downs, D. H. Dunkle, L. Eisley, M. K. Elias, H. Firner, V. Frechel, J. Frye, E. C. Galbreath, A. Graffham, B. George, M. Green, E. R. Hall, B. Hall, C. J. Hesse, J. E. Hill, H. Hildebrand, F. J. Holden, W. H. Hon, H. Jacob, M. Jewett, J. Kennedy, R. P. Ker- oker, W. Lane, B. Leonard, S. Lohman, T. McLaughlin, V. Mahon, M. Maldonado, D. Mallott, T. Moojen, W. K. McNown, F. Parks, M. Potter, E. S. Riggs, G. C. Rinker, R. Rinker, H. Setzer, H. T. U. Smith, L. H. Sprague, Stavely, F. Studer, E. H. and R. Taylor, J. Tihen, O. Tiemiere, J. Twente, C. C. Williams and W. Yost. E. H. Taylor based many new species on this isolated material, most of which were later syn- onymized by J. Tihen. F. E. Peabody (1948-1955) was a mem- ber of the Department of Zoology, and he could obtain museum support only by work- ing jointly with R. W. Wilson, the successor toC. W. Hibbard as curator of the collection of fossil vertebrates. Wilson was thus ac- tively involved in collecting in the Permian of Fort Sill, Oklahoma (1950), and in the Pennsylvanian of Garnett, Kansas (1953, 1954). Other workers at Garnett were V. Hogg, D. Horr, N. Hotton III, R. Weeks and two neighbors of the quarry. E. Gates and W. Slocum. After Peabody left the University of Kansas in 1955. R. R. Camp continued the excavation at Garnett in 1955. N. Hotton in, a member of the Department of Anatomy, was another vertebrate paleon- tologist at the University of Kansas inter- ested in Paleozoic amphibians and reptiles. He described Acroplous from the Speiser Shale at Keats, a locality discovered in the 1930's by M. Jewett. Hotton surveyed the exposures of the Speiser Shale, Lower Per- mian, throughout Kansas as described by Hattin (1957). Since then, the Speiser Shale exposures found continued interest. O. Bon- ner and L. D. Martin reopened the quarry at Keats in 1971 , and in the following years O. Bonner and S. Dart (1972), and R. W. Coldiron (1974) continued to collect there and at other localities. B. Foreman was active throughout 1976-1980 in the Speiser Shale at a new locality: Eskridge, Wa- baunsee County, Kansas; and in 1980 H.-P. Schultze began a study of fossil vertebrates of the Speiser Shale. It was then that D. Kirkpatrick discovered another locality rich in amphibians and lungfishes (near Junction City). Another Paleozoic locality (near Robinson, northeastern Kansas), known MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY since the 1890's was relocated by L. D. Martin and O. Bonner in 1974, and recol- lected by J. Chorn 1974-1979 with the help of K. Whetstone, C. Conley and H.-P. Schultze. Collecting in the Upper Cretaceous of western Kansas has continued since O. Bon- ner joined the Museum staff in 1968. Bonner also made important new collections of Kiowa Shale vertebrates from the Lower Cretaceous. L. D. Martin and E. Wiley participated in collecting the Kansas Cre- taceous, and L. D. Martin led parties into the Pierre Shale Cretaceous of Wyoming and South Dakota. Numerous students assisted in collecting the Cretaceous; the most nota- ble being J. D. Stewart who collected widely in the Niobrara Chalk. Others include: D. Bennett, M. Bonner, J. Chorn, D. Frailey, B. Foreman, J. Harrison, J. Junge, D. and P. Kopsick, M. Neuner, P. Overton, D. Nixon, G. Parker, E. Rickart. T. Setoguchi, B. Stein, and K. Whetstone. Reptile (and other significant specimens) material from the Niobrara Chalk was also obtained from Marion Bonner. In conclusion, the amphibians and rep- tiles from the Upper Pennsylvanian and Permian of eastern Kansas and reptiles from the Upper Cretaceous of western Kansas are the most significant parts of the fossil her- petological collection at the University of Kansas. The Pliocene amphibians widen the scope into the Cenozoic. The collection includes 72 holotypes, 1 1 paratypes, 32 syntypes, and 361 figured specimens; of those 2 holotypes, 3 paratypes, 18 syntypes, and 53 figured specimens could not be relo- cated. Fifty-five holotypes, and 9 paratypes characterize species stiH considered valid; 17 holotypes, and 2 paratypes have been synonymized. Eighteen holotypes represent the type-species of their genus. Abbreviations used in text are as follow: AMNH, American Museum of Natural His- tory; KUVP, Museum of Natural History, University of Kansas; MCZ, Museum of Comparative Zoology, Harvard University. ACKNOWLEDGMENTS We wish to express our thanks for help in various matters to O. Bonner, L. D. Martin, J. D. Stewart, and R. W. Wilson, all at the Museum of Natural History, University of Kansas, Lawrence, and to B. Foreman, In- dependence, Kansas. J. D. Stewart helped in assigning the Cretaceous localities to stage. We are especially grateful to Jan Elder and Coletta Spencer of the word processing serv- ice of the Division of Biology for their patience in typing a difficult manuscript. The compilation of the catalogue was supported by The National Science Founda- tion, Grant no. DEB-81 15956. TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES KUVP 350 KUVP 349 CATALOGUE CLASS AMPHIBIA Linnaeus 1758 Subclass Lepospondyli Zittel 1888 Order Nectridea Miall 1875 Family Urocordylidae Lydekker 1890 Crossotelos Case 1911 Crossotelos anmilatus Case 1911 (Type-species) Trimerorhachis leptorhynchus Case 1902 (nomen oblititm) Case described Trimerorhachis leptorhynchus in 1902 but in 191 IB dropped the taxon as indeterminate, suggesting that it might belong to the cricotid anthracosaur Cricotilhis Case 1902. Williston (1910A) suggested that Cricotillus brachydens Case was indistinct from the nectridean Crossotelos. Panchen (1970) and Olson (1967) both considered Cricotillus brachydens Case and Trimerorhachis leptorhynchus Case to be the same species, and Panchen (1970) placed Cricotillus brachydens Case in the order Batrachosauria, suborder Anthracosauria, incertae sedis. Olson (1970) considered Trimerorhachis leptorhynchus Case to be a nomen oblitum, but also considered Cricotillus brachydens Case to be closely related to the nectridean Diplocaulus, not Crossotelos as Williston (1910A) had thought. Olson retained Cricotillus brachydens Case in the Cricotidae. Milner (1978) followed Olson's conclusions on Tri- merorhachis leptorhynchus, but considered Cricotillus brachydens Case to be a subjective junior synonym of the urocordylid nectridean Crossotelos annulatus. Trimerorhachis leptorhynchus sp. nov.— Case 1902: p. 64, no figs; holotype. Trimerorhachis leptorhynchus Case— Olson 1970: text-fig. lA-E; pi. lA-B. Partial skull. Lower Permian, Wichita Group, Wellington Formation; two miles north- east of Orlando, SE'A, Sec. 31, T20N. RIW, Noble County, Oklahoma; collected by G. W. Stevens. Cricotillus Case 1902 Cricotillus brachydens Case 1902 (Type-species) Cricotillus brachydens g. et sp. nov.— Case 1902: p. 63, no figs; holotype. Cricotillus brachydens Case— Case 191 IB: text-tig. 20. Cricotillus brachydens Case— Olson 1970: text-fig. IF; pl. IC-D. Cricotillus brachydens Case— Milner 1978: text-fig. 6A-B; based on figures and information given by Olson (1970). Fragments of rostrum, including parts of the maxillaries, nasals, and vomers. Lower Permian, Wichita Group. Wellington Formation; two miles north- east of Orlando, SE'A, Sec. 31, T20N, RIW, Noble County, Oklahoma; collected by G. W. Stevens. Subclass Labyrinthodontia Owen 1860 Order Temnospondyli Zittel 1888 Suborder Rhachitomi Cope 1882 Superfamily Trimerorhachoidea Case 1946 Family Saurerpetontidae Chase 1965 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 9822 KUVP 28352 KUVP 49532 Acroplous Hotton 1959 Acroplous vorax Hotton 1959 (Type-species) Acroplous vorax n. sp.— Hotton 1959: p. 164, pi. 28, text-fig. 1-5; holotype; skull, mandible, vertebrae, limb elements, pectoral and pelvic girdles. Lower Permian, Council Grove Group, Speiser Shale Formation; NW'/4, SEV4, Sec. 36, T9S, R6E, Riley County, Kansas; Kitten Creek, just east of Keats, approximately 100 feet north of old U.S. Highway 24; collected by Mark Jewett in 1938. Acroplous vorax Hotton— Coldiron 1978: text-figs. 1-3, 4A-B; disarticu- lated specimen in situ, including skull roof, occiput, partial para- sphenoid, vertebrae, and scales. Collected from the type locality by L. Martin and O. Bonner in 1971. Acroplous— Bennett 1984: text-fig. 79; complete skull with interclavicle. Lower Permian, Council Grove Group, Speiser Shale Formation; SE'A, Sec. 36, T13S, RUE, Wabaunsee County, Kansas; approximately 1.5 miles northwest of Eskridge on State Highway 99; roadcut on north side; from mudstone above red shale; collected by Brian Foreman in 1978. Superfamily Eryopoidea Romer 1947 Family Eryopidae Cope 1882 Eryops Cope 1877 Eryops megacephalus Cope 1877 KUVP 348 Eryops megacephalus Cope— Williston 1899B: pi. 27, consists of figures of right scapulo-coracoid (inside view), which was erroneously cited as the left scapulo-coracoid, and a single tooth. Both figures are reversed. The tooth is figured as isolated but actual specimen is still in matrix. Plate 29; right scapulo-coracoid (outside view), erroneously cited as left scapulo-coracoid; figure is reversed. Plate 30, fig. 1, right mandible; erroneously cited as left mandible; figure is reversed. Plate 30, figs. 2,2a, listed as undetermined bones (identified by Moodie (1910) as the right clavicle). Orientation of both figures is correct. Eryops willistoni, new species— Moodie 1910: p. 243, pi. 49, figs. 1-5; pis. 50-52, pi. 53, figs. 1-8; holotype; parts of skull, right mandible, single maxillary tooth, forelimbs and pectoral girdle, vertebrae, and single rib. Figures 1 and 2 of pi. 49; pis. 50-51; and figures 1 and 2 of pi. 53 are based on the same material illustrated in Williston (1899B). Image orientation of all figures is correct. Lower Permian, Wichita Group, Wellington Formation, McCann Sand- stone; McCann quarry (earlier Nardin site, or Eddy site), on Deer Creek, 8 miles west and 4 miles south of Blackwell and 2.5 miles northeast of Eddy, NEV4, Sec. 16, and SW'A, Sec. 9, T26N, R2W, Kay County, Oklahoma; collected by C. N. Gould in 1897. Olson (1967) considered this species a synonym of E. megacephalus Cope. KUVP 5962 Superfamily Dissorophoidea Bolt 1969 Family Dissorophidae Boulenger 1902 Arkanserpeton Lane 1932 Arkanserpeton arcuatum Lane 1932 (Type-species) Arkanserpeton arcuatum genus et species nov.— Lane 1932: p. 315, pis. KUVP 17941 KUVP ? TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 26A-C; holotype; femur. This specimen was originally published as KUVP 1430. Upper Pennsylvanian, Allegheny Formation, a bed ot semianthracite coal within the Paris Shale; Eureka Mine, about one mile west of Paris, Logan County, Arkansas. This specimen was originally cited by Lane (1932) as KUVP 1430. The same number was later assigned to the holotype of Scaphiopus pliobatrachus Taylor 1937. The holotype of Arkanserpeton arcuatum Lane was later renumbered as KUVP 5962, while the holotype oi Scaphiopus pliobatrachus Taylor remained as KUVP 1430. Actiobates Eaton 1973 Actiobates peabodyi Eaton 1973 (Type-species) Actiobates peabodyi new species— Eaton 1973: p. 2, text-fig. 1; holotype; skull and skeleton. Upper Pennsylvanian, Missourian, Stanton Limestone Formation, Rock Lake Shale Member; approximately six miles northwest of Garnett, NWV4, NWV4, NE'/4, Sec. 5, T20S, R19E, Anderson County, Kansas. Suborder Stereospondyli Zittel 1888 Superfamily Capitosauroidea Save-Soderbergh 1935 Family Capitosauridae Save-Soderberg 1935 Mastodonsaurus Jaeger 1828 cf. Mastodonsaurus sp. indet.— Williston 1897F: pi. 21; single tooth; no KU number assigned; specimen not relocated. Lower Permian (cited as Upper Carboniferous); near Louisville, Pot- tawatomie County, Kansas; collected by Herbert Bailey in 1897. Eoscopus Daly 1976, nomen nudum Eoscopus lockardi Daly 1976, nomen nudum (Type-species) Eoscopus lockardi— Daly 1976: p. 100. Material in the collections of the Museum of Natural History, The University of Kansas, Lawrence, and in the collections of the Geological Department, Kansas State Univer- sity, Emporia. Upper Pennsylvanian, Virgilian, Shawnee Group, Topeka limestone; quarry two miles east of Hamilton, SE'A, Sec. 5, T24S, R12E. Green- wood County, Kansas. Daly (1976) cited distinguishing characters of a new unarmored dis- sorophoid, nevertheless the new name Eoscopus lockardi is not available because no figure (International Code Zoological Nomenclature, General Recommendations App. El 7) and no type specimen is given (International Code Zoological Nomenclature, Art. 72). Order Batrachosauria Efremov 1946, incertae sedis Batrachosauria incertae sedis Hesperoherpeton Peabody 1958 Peabody (1958) initially placed the genus Hesperoherpeton in the order Anthracosauria, family Cricotidae. Eaton and Stewart (1960) erected a new order (Plesiopoda) and family (Hesperoherpetonidae), based on more material. Romer (1966) did not follow Eaton and Stewart's conclusions and placed Hesperoherpeton in the order Anthracosauria, suborder ?Seymouriamorpha, incertae sedis. In placing Hesperoherpeton in the MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY order Batrachosauria. incertae sedis, without family designation, we follow Kuhn (1972). Hesperoherpeton garnettense Peabody 1958 (Type-species) KUVP 9976 Hesperoherpeton garnettense n. gen. and n. sp.— Peabody 1958: p. 571, text-fig. 1; holotype. Hesperoherpeton garnettense Peabody— Eaton and Stewart 1960: text-fig. 8. Right scapulo-coracoid and neural arch. Upper Pennsylvanian, Missourian, Stanton Limestone Formation; Hard- esty Farm, approximately six miles northwest of Garnett, NW'A, NW'A, NE'A. Sec. 5, T20S. R19E, Anderson County, Kansas; collected by F. E. Peabody in 1954. KUVP 10295 Hesperoherpeton garnettense Peabody— Eaton and Stewart 1960: text- figs. 1-7 and 9-11. Hesperoherpeton— Szarsk'i 1962: text-figs. 36 and 37; text-fig. 38 (after text-figs. 4, 2 and 1 1 in Eaton and Stewart 1960). Hesperoherpeton— Eaton 1962: text-fig. 3A (after text-fig. 11 in Eaton and Stewart 1960). Hesperoherpeton garnettense Peabody— Tatarinov, L. P. 1964 A: text-fig. lOOa-g (after text-figs. 1 , 2, 4, 5a, 7c, 7b, 1 1 respectively in Eaton and Stewart 1960). Hesperoherpeton garnettense Peabody— Kuhn 1965: text-fig. 16 (after figs. 1 and 4 in Eaton and Stewart 1960). Hesperoherpeton garnettense Peabody— Kuhn 1972: text-figs. 20.7-20.8, (after text-figs. 1 and 4 in Eaton and Stewart 1960). Various cranial elements, right mandible, vertebrae, right scapulo-cor- acoid, right clavicle, left forelimb. Collected from the type locality by R. R. Camp in 1955. Order Microsauria Dawson 1863 Suborder Tuditanomorpha Carroll and Gaskill 1978 Family Gymnarthridae Case 1910 Euryodus Olson 1939 Euryodus peabodyi (Carroll and Gaskill) 1978 Carroll and Gaskill (1978) determined that material previously referred to as Cardiocephahis cf. sternbergi by Gregory et al. (1956), from Fort Sill, Oklahoma, actually represented a new species, Cardiocephahis peabodyi. Schultze and Foreman (1981), in a review of the tuditanomorph micro- saurs, synonymized Cardiocephalus peabodyi into the genus Euryodus. KUVP 8967 Cardiocephalus cf. sternbergi— Gregory et al. 1956: text-figs. 8A-D; nearly complete skull with jaws. Lower Permian, Wichita Group, Wellington Formation, fissure fills within the Ordovician Arbuckle Limestone; Dolese Brothers limestone quarry at Richards Spur, just west of U.S. Hwys. 62 and 281, 10.5 miles south of Apache, 6 miles north of Fort Sill, SW'/:, Sec. 31, T4N, RllW, Comanche County. Oklahoma; collected by F. E. Peabody in 1950. Euryodus sp. KUVP 9923 Cardiocephahis cf. .sternbergi— Gregory et al. 1956: text-figs. 8E-G; isolated left lower jaw. KUVP 9928 Cardiocephalus cf. .sternbergi— Gregory et al. 1956: text-figs. 16E-G; isolated occipital-basioccipital complex. TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES KUVP KUVP KUVP KUVP KUVP KUVP KUVP 9929 9930 9931 9932 9933 9934 9923. Cardiocephalus sp.— Gregory et al. 1956: text-fig. 22; scapulo-coracoid. Cardiocephalus sp. — Gregory et al. 1956: text-tig. 22; scapulo-coracoid. Cardiocephalus sp.— Gregory et al. 1956 Cardiocephalus sp.— Gregory et al. 1956 Cardiocephalus sp.— Gregory et al. 1956 Cardiocephalus sp.— Gregory et al. 1956 9928-34 Lower Permian. Wichita Group. text-fig. 22; ischium, text-fig. 22; sacral rib. text-fig. 23; pelvis, text-fig. 23; pelvis. Wellington Formation, fissure fills within the Arbuckle Limestone; Dolese Brothers limestone quarry at Richards Spur, just west of U.S. Hwys. 62 and 281. 10.5 miles south of Apache. 6 miles north of Fort Sill, SW'A, Sec. 31, T4N. RllW. Comanche County, Oklahoma; collected by F. E. Peabody in 1950. Euryodus primus Olson 1939 KUVP 9922 Euryodus primus Olson— Gregory et al. 1956: text-fig. 14; isolated lower jaw. Specimen not relocated. KUVP 9926 Euryodus primus Olson— Gregory et al. 1956: text-fig. 16B-D; isolated atlas. KUVP 9936 Euryodus primus Olson— Gregory et al. 1956: text-fig. 15; sections of dentary teeth. KUVP 9922, 9926, 9936 Lower Permian, Wichita Group, Wellington Formation, fissure fills within the Ordovician Arbuckle Limestone; Dolese Brothers lime- stone quarry at Richards Spur, just west of U.S. Hwys. 62 and 281 . 10.5 miles south of Apache, 6 miles north of Fort Sill. SW'A. Sec. 31, T4N, RllW, Comanche County. Oklahoma; collected by F. E. Peabody in 1950. Euryodus bonneri Schultze and Foreman 1981 KUVP 47367 Euryodus bonneri n. sp.— Schultze and Foreman 1981: p. 3. text-figs. 1, 2; holotype; nearly complete, articulated skull with jaws, vertebrae, and humerus. Lower Permian, Middle Wolfcampian, Council Grove Group, Lower Speiser Shale Formation; road-cut on Kansas State Highway 99, 1.5 miles northwest of Eskridge, SE'A, Sec. 36, T13S. RUE. Wabaunsee County, Kansas; collected by Brian Foreman in 1976. Labyrinthodontia incertae sedis KUVP 24002 Unidentified labyrinthodont— Bolt 1974: text-fig. 1; left vomer, (er- roneously described as a right palatine). Unidentified labyrinthodont— Schultze and Chorn 1983: fig. la-c; left vomer. KUVP 65042 Unidentified labyrinthodont-Schultze and Chorn 1983: figs. 2, 3a-c; left vomer. Lower Permian, Wichita Group, Wellington Formation, fissure fills within the Ordovician Arbuckle Limestone; Dolese Brothers limestone quarry, at Richards Spur (Ft. Sill), SW'A, Sec. 31, T4N, RllW, Comanche County, Oklahoma. KUVP 82697 Labyrinthodont ?intercentrum— Schultze 1985: text-fig. 8.3. Lower Permian, Gearyan, Council Grove Group, Upper Speiser Shale Formation; road cut on KS Hwy. 38, SE'A, SE'A, Sec. 20, T32S. R7E, Cowley County. Kansas. Subclass Lissamphibia Haeckel 1866 Order Anura Laurenti 1768 Family Pelobatidae Stannius 1856 10 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Scophiopus Holbrook 1836 Scaphiopus pUobatrachus Taylor 1937 KUVP 1430 Scaphiopus pUobatrachus sp. nov.— Taylor 1937: p. 515, pi. 1, fig. 1; holotype. Scaphiopus pUobatrachus T?iy\or — Taylor 194 IB: text-fig. lA-B; coccyx with fused sacral vertebrae. KUVP 1434 Scaphiopus pUobatrachus TdLyXor — Taylor 1941B: text-ftg. 3A; fragment of radio-ulna. KUVP 1432 Scaphiopus pUobatrachus Taylor— Taylor 194 IB: text-fig. 3B; fragment of humerus. KUVP 1436 Scaphiopus pUobatrachus Taylor— Taylor 1941B: text-fig. 3C; fragment of ilium. KUVP 1431 Scaphiopus pUobatrachus Taylor— Taylor 1941B: text-fig. 3D-E; frag- ment of humerus. KUVP 1433 Scaphiopus pUobatrachus Taylor — Taylor 194 IB: text-fig. 3F; fragment of radio-ulna. KUVP 1430-1434, 1436 Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SWA, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by H. T. Martin and C. W. Hibbard in 1928. KUVP 1469 Scaphiopus antiquus n. sp.— Taylor 1941B: p. 184, text-fig. 2; holotype; coccyx with fused sacral vetebrae. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SWA, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by D. Dunkle and E. H. Taylor in 1935. Tihen (1960) considered Scaphiopus antiquus Taylor to be a synonym of Scaphiopus pUobatrachus Taylor. Scaphiopus studeri Taylor 1938 KUVP 1478 Scaphiopus studeri sp. nov. -Taylor 1938: p. 408, pis. 42-43; pi. 44, figs. 2, 4-6, 8, 10, 12-14; pi. 45, figs. 2-3, 6, 10-13; holotype. Scaphiopus studeri Taylor— Lane 1945B: text-fig. 7 (after pi. 42 in Taylor 1938). Scaphiopus— BQxxntiX 1984: text-fig. 81 (after pi. 42 in Taylor 1938). Skull and skeleton, part and counterpart. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); about one mile east of "Rhino Hill Quarry," near Wallace County line. Sec. 7, TllS, R37W, Logan County, Kansas; collected by Frank Studer in 1934. Scaphiopus diversus Taylor 1942 KUVP 6368 Scaphiopus diversus n. sp.— Taylor 1942: p. 202, pi. 15, tig. 2A-B; holotype; portion of combined sacral vertebrae and coccyx. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade. NE'A, SWA, Sec. 22, T33S, R29E. Meade County, Kansas; collected by C. W. Hibbard and party in 1938. KUVP 6367 Neoscaphiopus noblei n. gen. et n. sp.— Taylor 1942: p. 204, pi. 15, fig. 5A-B; holotype, portion of combined sacral vertebrae and coccyx. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S. R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Tihen ( 1960) considered Neoscaphiopus noblei Taylor to be a synonym of Scaphiopus diversus Taylor. Family Bufonidae Fitzinger 1826 Bufo Laurenti 1768 TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES KUVP 1446 KUVP ? KUVP 1438 KUVP 1439 KUVP 1440 KUVP 65019 KUVP 1449 KUVP 1450 KUVP 1448 KUVP 1447 KUVP 6397 KUVP 1452 Bufo hihhardi Taylor 1937 KUVP 1437 Bufo hihhardi sp. nov.— Taylor 1937: p. 517, pi. 1, fig. 3; holotype. Bufo hihhardi Taylor— Taylor 194 IB: text-fig. 5; erroneously cited as ' KUVP 1472. Bufo hihhardi Taylor— Tihen 1962: text-fig. 62 (after text-fig. 5 in Taylor 1941). Sacral vertebra. Bufo hihhardi sp. nov.— Taylor 1937: pi. 1. fig. 4; fragment of urostyle. Bufo hihhardi ? sp. nov.— Taylor 1937: pi. 1, fig. 5; sacral vertebra. Published as KUVP 1466a. Specimen not relocated. Bufo hihhardi sp. nov.— Taylor 1937: pi. 1, fig. 6; scapula. Bufo hihhardi sp. nov.— Taylor 1937: pi. 1, fig. 7; fragment of humerus. Bufo hihhardi s^ . nov.— Taylor 1937: pi. 1, fig. 8; fragment of radio-ulna. Bufo hihhardi sp. nov.— Taylor 1937: pi. 1, fig. 9; fragment of radio-ulna. " Published as KUVP 1440a. Bufo hihhardi sp. nov.— Taylor 1937: pi. 2, fig. 3; fragment of ilium. Bufo hihhardi sp. nov.— Taylor 1937: pi. 2. fig. 4; fragment of ilium. Specimen not relocated. Bufo hihhardi sp. nov.— Taylor 1937: pi. 2, fig. 13; fragment of tibia- fibula. Specimen not relocated. Bufo hihhardi sp. nov.— Taylor 1937: pi. 2, fig. 14; fragment of femur. Bufo hihhardi Taylor— Tihen 1962: text-fig. 36; fragment of ilium. Bufo arenarius sp. nov.— Taylor 1937: p. 521. pi. 1, fig. 2; holotype. Bufo arenarius Taylor— Taylor 194 IB: text-fig. 6A; erroneously cited as KUVP 1470. Bufo arenarius Taylor— Tihen 1962: text-fig. 62 (after fig. 6 A in Taylor 1941). Sacral vertebra. Bufo arenarius sp. nov.— Taylor 1937: pi. 1 Bufo arenarius sp. nov.— Taylor 1937: pi. 1 Bufo arenarius ? sp. nov.— Taylor 1937: scapula. Published as KUVP 1458a. Specimen not relocated Bufo arenarius sp. nov.— Taylor 1937: pi. 1 not relocated. Bufo arenarius sp. nov.— Taylor 1937: pi. 1 imen not relocated. Bufo arenarius sp. nov.— Taylor 1937: pi. 2. Bufo arenarius sp. nov.— Taylor 1937: pi. 2, Bufo arenarius sp. nov.— Taylor 1937: pi. 2 fibula. Bufo arenarius sp. nov. — Taylor 1937: pi. 2 fibula. Bufo arenarius sp. nov.— Taylor 1937: pi. 2 fibula. Published as KUVP 1464c. Specimen not relocated. Bufo arenarius sp. nov.— Taylor 1937: pi. 2, fig. 10; fragment of femur. Bufo arenarius sp. nov.— Taylor 1937: pi. 2, fig. 11; fragment of tibia- fibula. Published as KUVP 1459a. Specimen not relocated. Bufo arenarius sp. nov.— Taylor 1937: pi. 2, fig. 12; fragment of femur. 1446-50; 1455; 1457-61; 1464-66; 6397; 65019 Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SWV4, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by H. T. Martin and party in 1928. KUVP 1474 Bufo arenarius Taylor— Taylor 194 IB: fig. 6B; sacral vertebra; collected KUVP 1456 KUVP 1458 KUVP ? KUVP 1455 KUVP 1457 KUVP 1466 KUVP 1465 KUVP 1461 KUVP 1464 KUVP ? KUVP 1460 KUVP? KUVP 1459 KUVP 1437-40 , fig. 10; humerus. , fig. 11; scapula. pi. 1, fig. 12; fragment of fig. 13; humerus. Specimen , fig. 14; radio-ulna. Spec- . fig. 1; fragment of ilium. , fig. 2; fragment of ilium, fig. 6; fragment of tibia- fig. 7; fragment of tibia- fig. 9; fragment of tibia- 12 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY at the type locality by E. H. Taylor and D. Dunkle in 1935. Tihen (1962) considered Bufo arenahus Taylor to be a synonym of Bufo hihbardi Taylor. Bufo sp. KUVP ? Bufo sp.— Taylor 1937: pi. 2. fig. 5; fragment of tibia-fibula. Published as KUVP 1464a. Specimen not relocated. KUVP ? Bufo sp.— Taylor 1937: pi. 2, fig. 8; fragment not identified. Published as KUVP 1464b. Specimen not relocated. Tertiary, Late Miocene. Ogallala Formation (Hemphillian land mammal age); Edson Quarry. SWA, Sec. 25, TIOS, R38W. Sherman County, Kansas; collected by H. T. Martin and party in 1928. 6319 Bufo sp.?-Taylor 1942: pi. 17; figs. lOA-lOB; coccyx. 6334 Bufo sp.— Taylor 1942: pi. 19; fig. 12; ilium. 6335 Bufo sp.— Taylor 1942: pi. 19; fig. 13; ilium. 61319, 6334-35 Tertiary. Pliocene. Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade. NE'A, SWA. Sec. 22. T33S, R29E, Meade County. Kansas; collected by C. W. Hibbard and party in 1938. KUVP KUVP KUVP KUVP Family Ranidae Bonaparte 1831 Ratia Linnaeus 1758 Holman (1963) considered Anchylorana Taylor a synonym of Rana Linnaeus. Rana dubita (Taylor) 1942 KUVP 6377 Anchylorana dubita, n. sp. -Taylor 1942: p. 209, pi. 15, fig. 4A-B; holotype sacrum. Tertiary, Pliocene. Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade. NE'A. SWA. Sec. 22, T33S, R29E. Meade County. Kansas; collected by C. W. Hibbard and party in 1938. Rana ephippium Taylor 1942 KUVP 6370 Rana ephippium, n. sp. -Taylor 1942: p. 214. pi. 14. fig. lA-B; holotype; sacral vertebra with fragmentary diapophyses. Tertiary, Pliocene. Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A. SWA, Sec. 22, T33S. R29E, Meade County. Kansas; collected by C. W. Hibbard and party in 1938. Rana fayeae Taylor 1942 KUVP 6378 Rana fayeae, n. sp.— Taylor 1942: p. 212. pi. 14. fig. 4A-B; holotype; nearly complete sacral vertebra. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Rana meadensis Taylor 1942 KUVP 6376 Rana meadensis, n. sp.— Taylor 1942: p. 213, pi. 14, fig. 5A-B; holotype; nearly complete sacral vertebra. Tertiary, Pliocene. Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade. NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Rana moorei (Taylor 1942) KUVP 6375 Anchylorana moorei, n. sp.— Taylor 1942: p. 208, pi. 15, fig. 3A-B; holotype; fragmentary sacrum. TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 13 Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA. Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. Rana parvissima Taylor 1942 KUVP 6451 Rana parvissima, n. sp.— Taylor 1942: p. 217, pi. 16, fig. 2; holotype; fragment of a sacral centrum, with one coccygeal condyle and the right diapophysis present. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E. Meade County, Kansas; collected by C. W. Hibbard and party in 1941. Rana rexroadensis Taylor 1942 KUVP 6369 Rana rexroadensis, n. sp.— Taylor 1942: p. 215, pi. 14, fig. 3A-B; holotype; sacral vertebra, missing diapophyses. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S. R29E. Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Rana robustocondyla (Taylor 1942) KUVP 5106 Anchylorana robustocondyla, n. sp.— Taylor 1942: p. 211, pi. 15, fig. 1; holotype; sacral centrum and eighth vertebra. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E. Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Rana valida Taylor 1942 KUVP 5133 Rana valida, n. sp.— Taylor 1942: p. 216, pi. 14, fig. 2A-B; holotype; sacral vertebra, missing diapophyses. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Rana sp. KUVP 6379 Rana sp. -Taylor 1942: Pl- KUVP 6314 Rana sp. -Taylor 1942: pi. KUVP 5107 Rana sp. -Taylor 1942: pl. KUVP 65092 Rana sp. -Taylor 1942: pl. KUVP 6385 Rana sp ?-Taylor 1942: specimen is missing. KUVP 5134 Rana sp. ?-Taylor 1942 :pl KUVP 5105 Rana sp. ?-Taylor 1942 :pl KUVP 5103 Rana sp. ?-Taylor 1942 :pl KUVP 6364 Rana sp. -Taylor 1942: pi. KUVP 6365 Rana sp. -Taylor 1942: pi. KUVP 6360 Rana sp. -Taylor 1942: pi. KUVP 5121 Rana sp. -Taylor 1942: pi. KUVP 6317 Rana sp. -Taylor 1942: pi. KUVP 6354 Rana sp. -Taylor 1942: pi. KUVP 6345 Rana sp. -Taylor 1942: pi. KUVP 6362 Rana sp. -Taylor 1942: pi. KUVP 65093 Rana sp. -Taylor 1942: pi. KUVP 6349 Rana sp. -Taylor 1942: pi. KUVP 6310 Rana sp. -Taylor 1942: pi. KUVP 5103, 5105,5107, 5121, 5134, 6: HO, 16, fig. 1; sacral vertebra. 16, fig. 1 1; humerus. 16, fig. 12; tibio-fibula. 16, fig. 13; femur. Cited as KUVP 5107A. pl. 17, fig. lA-lB; scapula. Half of this 1. 17, fig. 2A-2B; scapula. . 17, fig. 3A-3B; scapula. . 17, fig. 4; scapula. 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 19, fig 6314, 6317, 6345, 6349. 6354, 6365, 6379, Hum. Specimen not relocated. Hum. Hum. Hum. Hum. Hum. "ium. Hum. Hum. Cited as KUVP 6364A. 10; ilium. 1 1; ilium. 14 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 6327 KUVP 6328 KUVP 6323 KUVP 5088 KUVP 6383 KUVP 6318 KUVP 6325 KUVP 5088, KUVP 6306 KUVP 6356 KUVP 6371 KUVP 6312 KUVP 6352 KUVP 6342 KUVP 6355 KUVP 6373 KUVP 6372 KUVP 6346 KUVP 6311 KUVP 6348 6385, 65092-3 Tertiary, Pliocene. Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Ranidae indeterminate (referable to Anchylorana) Ranidae indet.— Taylor 1942: pi. 17, fig. 6A-6B; fragment of coccyx. Cited as, "probably referable to Anchylorana/^ Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Ranidae indeterminate (referable to Rana) Ranidae indet.— Taylor 1942: pi. 17, fig. 7A-7B; fragment of coccyx. Cited as, "probably referable to Rana.'' Ranidae indet.— Taylor 1942: pi. 17, fig. 8A-8B; fragment of coccyx. Cited as, "probably referable to Rana." Ranidae indet.— Taylor 1942: pi. 17, fig. 9A-9B; fragment of coccyx. Cited as, "probably referable to Rana." Ranidae indet.— Taylor 1942: pi. 17, fig. 12A-12B; coccyx. Cited as, "probably referable to Rana." Ranidae indet.— Taylor 1942: pi. 17, fig. 13A-13B; coccyx. Cited as, "probably referable to Rana." Ranidae indet.— Taylor 1942: pi. 17, fig. UA-llB; coccyx. 6318, 6323, 6325, 6328, 6383 Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Anura indeterminate (referable to Rana) Anura indet.— Taylor 1942: pi. 18, fig. 1; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 2; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 3; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 4; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 5; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 6; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 7; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. Rana. Anura indet.— Taylor 1942: pi. Rana. Anura indet.— Taylor 1942: pi. 18, fig. 10; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18. fig. 11; ilium. Cited as referable to Rana. Anura indet.— Taylor 1942: pi. 18, fig. 12; ilium. Cited as referable to Rana. fig. 8; ilium. Cited as referable to fig. 9; ilium. Cited as referable to TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 15 KUVP 5086 KUVP6331 KUVP 6307 KUVP 6333 KUVP 6315 KUVP 6359 KUVP 6340 KUVP 6341 KUVP 6353 KUVP 6350 KUVP 6358 KUVP 6316 KUVP 6336 KUVP 6332 KUVP 6339 KUVP 6363 KUVP 6357 KUVP 6366 KUVP 6337 ?KUVP 6374 KUVP 6361 KUVP? KUVP 6347 KUVP 6309 KUVP? 8, fig. 13; ilium. Cited as referable to Anura indet.— Taylor 1942: pi. Rami. Anura indet.— Taylor 1942: pi. 18, fig. 14 Rami. Anura indet.— Taylor 1942: pi. 18, fig. 15 Rami. Anura indet.— Taylor 1942: pi. 18, fig. 16 Rami. Anura indet.— Taylor 1942: pi. 18, fig. 17 Rami. Anura indet.— Taylor 1942: pi. 18, fig. 18 Rana. Anura indet.— Taylor 1942: pi. 18, fig. 19 Rami. Anura indet. -Taylor 1942: pi. 18, fig. 20 Rami. Anura indet.— Taylor 1942: pi. 18, fig. 21 Rana. Anura indet.— Taylor 1942: pi. 18, fig. 22 Rana. Anura indet.— Taylor 1942: pi. 18, fig. 23 Rana. Anura indet.— Taylor 1942: pi. 20, fig. 1; ilium. Cited as, referable to Rana Anura indet.— Taylor 1942: pi. 20, fig. 2; ilium. Cited as referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet. — Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. Anura indet.— Taylor referable to Rana. ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to ilium. Cited as referable to 'mostly Anura indet.— Taylor referable to Rana. Anura indet.— Taylor 1942: pi. 20, fig. 3; ilium. Cited as, 1942: pi. 20, fig. 4; ilium. Cited as, 1942: pi. 20, fig. 5; ilium. Cited as, 1942: pi. 20, fig. 6; ilium. Cited as, 1942: pi. 20, fig. 7; ilium. Cited as, 1942: pi. 20, fig. 8, ilium. Cited as, 1942: pi. 20, fig. 9, ilium. Cited as, 1942: pi. 20, fig. 10, ilium. Cited as, 1942: pi. 20, fig. 11, ilium. Cited as. Erroneously listed as KUVP 6309, as it does not match the figure. Specimen not relocated. Anura indet.— Taylor referable to Rana. 1942: pi. 20, fig. 12; ilium. Cited as, 1942: pi. 20, fig. 13; ilium. Cited as. 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 'mostly 1942: pi. 20, fig. 14; ilium. Cited as, "mostly referable to Rana.'" Originally listed as KUVP 6336A. Specimen not relocated. 16 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY 15; ilium. Cited as, "mostly 16; ilium. Cited as, "mostly KUVP6351 Anura indet. -Taylor 1942: pi. 20, fig. referable to Rana/^ KUVP 6338 Anura indet. -Taylor 1942: pi. 20, fig. referable to Rana.'^ KUVP 6344 Anura indet. -Taylor 1942: referable to Rana." KUVP 6343 Anura indet.— Taylor 1942: referable to Rana.'' KUVP 6329 Anura indent. -Taylor 1942: referable to Rana." KUVP 5086, 6306-07, 6309, 6311-12, 6315-16, 6219, 6331-33, 6336-44, 6346-48, 6350-53, 6355-59, 6361, 6363, 6366, 6371-74. and all KUVP? Terti- ary, Pliocene, Rexroad Formation (Blancan land mammal age); 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. pi. 20, fig- 17; ilium. Cited as. "mostly pi. 20. fig- 18; ilium. Cited as. "mostly pi. 20 fig- 19; ilium. Cited as, "mostly Anura indeterminate KUVP 6384 Anura indet. -Taylor 1942: KUVP 5096 Anura indet. -Taylor 1942: KUVP 6320 Anura indet. -Taylor 1942: KUVP 6322 Anura indet. -Taylor 1942: KUVP 6321 Anura indet. -Taylor 1942: KUVP 6324 Anura indet. -Taylor 1942: KUVP 6382 Anura indet. -Taylor 1942: KUVP 6381 Anura indet. -Taylor 1942: KUVP 6326 Anura indet. -Taylor 1942: KUVP 5096. 6320-22, 6324, 6326, 6381-82, tion (Blancan land mammal SWV4, Sec. 22, T33S, R29E Hibbard and party in 1938. pi. 16, fig. 3A,3B; parasphenoid. pi. 16, fig. 4A,4B; radio-ulna. pi. 16, fig. 5A,5B; radio-ulna. pi. 16, fig. 6A,6B; radio-ulna. pi. 16, fig. 7A,7B; radio-ulna. pi. 16, fig. 8A,8B; radio-ulna. pi. 16, fig. 9A.9B; radio-ulna. pi. 16, fig. 10A,10B; radio-ulna. pi. 17, fig. 5A,5B; fragment of coccyx. 6384 Tertiary, Pliocene, Rexroad Forma- age); 16 miles southwest of Meade. NE'A. , Meade County, Kansas; collected by C. W. Order Urodela Baur 1888 Suborder Cryptobranchoidea Dunn 1922 Family Cryptobranchidae Cope 1889 Andhas Tschudi 1837 Andhas matthewi (Cook 1917) KUVP 12004d Andhas matthewi— Meszoely 1966: text-figs. 5a, 6b; trunk vertebra. Erroneously published as UK12004-C. Andhas matthewi— Estes 1981: text-fig. 2d (redrawn from Meszoely 1966. text-figs. 5a, 6b). Tertiary, Miocene, Pawnee Creek Formation (Hemingfordian land mam- mal age), Martin Canyon local fauna; "Quarry A," NE'A, Sec. 27, TUN, R53W, Logan County, Colorado. Family Ambystomatidae Hallowell 1856 Ambystoma Tschudi 1838 Ambystoma kansensis (Adams 1929) KUVP 5250 Plioambystnma kansensis gen. et sp. nov.— Adams, in Adams and Martin 1929: p. 505. no figs.; holotype. Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 32. Plioambystoma kansense (error) — Lane 1945B: text-fig. 6. KUVP 5250 has been cited as the holotype, however it is a composite of disarticulated pieces. Single elements have been described and figured in TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 17 Adams and Martin (1929, 1930), but they are not the same elements used to reconstruct the skull or skeleton of the holotype. The paratypes designated by Adams in Adams and Martin (1929), KUVP 5251-53, have not been relocated. On this basis, the elements figured in the 1929 paper by Adams and Martin are considered syntypes. Composite skull and skeleton. KUVP 65020 Plioambystoma kansensis—Adsim^, in Adams and Martin 1929: text-fig. 1; syntype; right premaxilla. KUVP ? Plioambystoma kansensis—Adsim^, in Adams and Martin 1929: text-fig. 2; syntype; left maxilla; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 3; syntype; right prefrontal; specimen not relocated. KUVP 65021 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 4; syntype; left parietal. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 5; syntype; right squamosal; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 6; syntype; right frontal; specimen not relocated. KUVP 65022 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 7; syntype; right orbitosphenoid. KUVP 65023 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 8; syntype; left otic capsule. KUVP 65024 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 9; syntype; left otic capsule. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 10; syntype; right stapes; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 11; syntype; right quadrate; specimen not relocated. KUVP 65025 Plioambystoma kansensis— Adams, in Adams and Martin 1929; text-fig. 12; syntype; left otic capsule. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 13; syntype; left quadrate; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 14; syntype; left maxillary teeth; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 15; syntype; left pterygoid; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 16; syntype; right dentary; specimen not relocated. Ambystoma kansense (error)— Estes 1981 : text-fig. 14d upper figure (after Adams and Martin 1929, text-fig. 16). KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 17; syntype; second basibranchial; specimen not relocated. KUVP 65026 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 18; syntype; left angular. Ambystoma kansense (error)— Estes 1981 : text-fig. 14d lower figure (after Adams and Martin 1929, text-fig. 18). KUVP 65027 Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 19; syntype; sacral vertebra. Ambystoma kansense (error)— Estes 1981: text-fig. 14e left (after Adams and Martin 1929, text-fig. 19). KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 20; syntype; left rib; specimen not relocated. KUVP ? Plioambystoma kansensis— Adams, in Adams and Martin 1929: text-fig. 18 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY 21; syntype; parasphenoid; specimen not relocated. KUVP ? Plioambystoma kansensis—Mdim?,, in Adams and Martin 1929: text-fig. 22; syntype; axis; specimen not relocated. Ambystoma kansense (error)— Estes 1981 : text-fig. 14e right (after Adams and Martin 1929, text-fig. 22). KUVP ? Plioambystoma kansensis—Kdam?,, in Adams and Martin 1929: text-fig 23; syntype; sagittal section of body vertebra; specimen not relocated KUVP ? Plioambystoma kansensis—JKddims. in Adams and Martin 1929: text-fig 24; syntype; third caudal vertebra; specimen not relocated. KUVP ? Plioambystoma kansensis—kddim?,, in Adams and Martin 1929: text-fig 25; syntype; left scapulo-coracoid; specimen not relocated. KUVP 65028 Plioambystoma kansensis—Mdims,, in Adams and Martin 1929: text-fig 26; syntype; right humerus. KUVP 65029 Plioambystoma kansensis—Msim?,, in Adams and Martin 1929: text-fig 27; syntype; right radius and ulna; right ulna not relocated. KUVP 65030 Plioambystoma kansensis— Addim?,, in Adams and Martin 1929: text-fig 28; syntype; right fibula. KUVP 65031 Plioambystoma kansensis—M?im?,, in Adams and Martin 1929: text-fig 29; syntype; right tibia. KUVP ? Plioambystoma kansensis—A.d?Lva^. in Adams and Martin 1929: text-fig 30; syntype; right ilium; specimen not relocated. KUVP 65032 Plioambystoma kansensis—kd?ixns, in Adams and Martin 1929: text-fig 31; syntype; ilium, ischium and sacral rib in position. KUVP 65033 Plioambystoma kansensis— A.d?ims, in Adams and Martin 1929: text-fig 32; syntype; left femur. KUVP 65034 Plioambystoma kansasenese (error) — Adams and Martin 1930: pi. 31, fig 1; right premaxilla. KUVP 65035 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, fig 2; right parietal. KUVP 65036 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 3; right frontal. KUVP 65037 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 4; right squamosal. KUVP 65071 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, fig 5; left pterygoid. KUVP ? Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 6; right maxilla; specimen not relocated. KUVP 65072 Plioambystoma kansasenese (error) — Adams and Martin 1930: pi. 31 , fig 7; right orbitosphenoid. KUVP 65073 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 8; right quadrate. KUVP 65074 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 . fig 9; right prefrontal. KUVP 65075 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 10; parasphenoid. KUVP ? Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 , fig 11; axis; specimen not relocated. KUVP 65076 Plioambystoma kansasenese (error) — Adams and Martin 1930: pi. 31, fig 12; body vertebra. KUVP 65077 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, fig 13; sacral vertebra. KUVP 65078 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, fig 14; left otic capsule. KUVP 65079 Plioambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, fig TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 19 15; caudal vertebra. KUVP 65080 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 16; second basibranchial. KUVP 65081 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 3 17; ilium. KUVP 65082 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 3: 18; ischium. KUVP ? PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 19; left scapulo-coracoid; specimen not relocated. KUVP 65083 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 20; left humerus. KUVP ? PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 21; left ulna; specimen not relocated. KUVP ? PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 22; left radius; specimen not relocated. KUVP 65084 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 23; right femur. KUVP 65085 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, 24; right tibia. KUVP 65086 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 25; left fibula. KUVP 65087 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 26; dentary. KUVP 65088 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 27; left rib. KUVP ? PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31. 28; left rib; specimen not relocated. KUVP 65089 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31, 29; left rib. KUVP ? PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 . 30; right rib; specimen not relocated. KUVP 65090 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 31 . 31; right rib. KUVP 65091 PUoambystoma kansasenese (error)— Adams and Martin 1930: pi. 3 1 . 32; right sacral rib. KUVP 5250; 65020-37; 65071-91; and all KUVP ?: Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SWA, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by H. T. Martin and party in 1928. Tihen (1958) considered the genus PUoambystoma Adams to be a synonym of Ambystoma Tschudi. Taylor (1941B), Lane (1945B) and Tihen (1958) erroneously listed the specific name of this taxon as kansense, rather than kansensis. KUVP 1468 Lanebatrachus martmi n. gen. and n. sp.— Taylor 1941B: p. 180, text-fig. 4A-B; holotype; portion of dentary and angular. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SWA, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by H. T. Martin and party in 1926. KUVP 1470 OgaUalabatrachus horahum n. gen. and n. sp.— Taylor 194 IB: p. 181, text-figs. 7A-C; holotype; dorsal vertebra. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Rhino Hill Quarry, SE'A, NE'A, Sec. 11, TllS, R38W, Wallace County, Kansas; collected by E. H. Taylor and D. Dunkle in 1935. 20 KUVP? MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Tihen (1958) considered Lanebatrachus martini Taylor and Ogallalabatrachus horahum Taylor to be synonyms of Plioambystoma { = Amby stoma) kansense { — kansensis). "Amphibia" Onychopus gigas gen. et sp. nov.— Martin 1922: p. 104, pis. 1 and 3; holotype; slab containing numerous tracks; specimen not relocated. Upper Pennsylvanian, Virgilian, Douglas Group, Stranger Formation, probably Tonganoxie Sandstone in Weston Shale Member; east bank of Wakarusa River at Dightman's Crossing, 5 miles east-southeast of Lawrence, NE'A, Sec. 14, T13S, R20E, Douglas County, Kansas; collected by R. and J. Coghill in 1919. This material was originally described by Martin as tracks of a giant amphibian, however, Baird (1963) referred to them as fossil tree stump holes. KUVP 35 1 COHORT AMNIOTA Haeckel 1866 CLASS "REPTILIA" Laurenti 1768 (excluding Aves and Mammalia) Subclass Captorhinomorpha Watson 1917 Suborder Captorhinoidea Vaughn 1955 Family Captorhinidae Case 1911 Captorhinidae indet. Pleuristion brachycoelus Case 1902, nomen dubium (Type-species) When first described. Case considered Pleuristion brachycoelus to be of indeterminate affinity within the Reptilia. In 1907 Case placed the genus in the suborder Pelycosauria, incertae sedis, but in 1911 A he included Pleuristion in the amphibian suborder Temnospondyli. Olson (1970) stated that Case, in considering Pleuristion to be a temnospondylous amphibian, was confusing the similarity in form between microsaurs and captorhinomorphs. Williston (1910B) referred Pleuristion back to the Reptilia. and in 1916, noted the similarities between it and the cap- torhinid, Captorhinus. Von Huene (1925) placed Pleuristion back in the Pelycosauria but later Romer and Price (1940), on the basis of more material, placed the genus in the order Cotylosauria. Later Romer (1956, 1966) placed the genus in the family Captorhinidae and Olson (1970) followed Romer's conclusions. Heaton (1980) considered the Cotylosauria (consisting of the seymouriamorphs, diadectids. and related forms), to be an order of advanced non-anthracosaurian amphibians in the subclass Batrachosauria, closely related to the Reptilia. According to Olson (1984) KUVP 351 is indeterminate within the Captorhinidae and P. brachycoelus is a nomen dubium. Pleuristion brachycoelus, gen. et sp. nov.— Case 1902: p. 67; no text- figures; holotype; a series of three cervical vertebrae. In the original description. Case failed to designate a holotype. Instead he used two specimens which Olson (1970) considered as representing two differ- ent individuals, an adult and a juvenile, designated KUVP 351a and 351b respectively. Olson referred to the adult specimen as the holotype. Also see KUVP 65275. Pleuristion brachycoelus— CasQ 1907: pi. 1, figs. 14-15. Pleuristion brachycoelus Case— Olson 1970: text-fig. 3A-C; pi. 2, fig. D. Type locality erroneously stated to be in Logan County, Oklahoma. Pleuristion brachycoelus— 0\son 1984: text-fig. 6A-C, lower figure (from TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 21 lOD. Erroneously listed as KUVP 291 (after text-fig. 2 in Peabody 188c, (after text-fig. 2 , 13 (after text-fig. 2 in Peabody in Peabody Olson 1970, text-fig. 3A-C). Type locality erroneously stated to be in Logan County, Oklahoma. KUVP 65275 Pleuristion bnichycoelus Case— Olson 1970: fig. 3J, pi. 2, fig. E. This specimen, along with KUVP 35 1 , is the original material used by Case in erecting the genus, and consists of eleven vertebrae plus rib fragments, a femur, and phalangeal elements from an immature individual. This specimen was designated by Olson as referred material and given the number KUVP 351b. Locality erroneously stated by Olson to be in Logan County, Oklahoma. KUVP 351, 65275 Lower Permian, Wichita Group, Wellington Formation; at Alva, two miles northeast of Orlando; SE'A, Sec. 31, T20N, RIW; Noble County, Oklahoma; collected by G. W. Stevens. Captorhinus Cope 1896 Captorhinus agiiti (Cope 1882) KUVP 8964 Captorhinus— Vtdihody 1951 : text-fig. 2; figure is a composite of the right tarsus, based on KUVP 8964, 8965, and 9783. The astragalus, tibiale, centralia, and distal tarsale 1, 2, and 3, in the figure are based on KUVP 8964 (also see KUVP 9783); pi. 1, fig. D, right astragalus and "navicular." Captorhinus sp.— Peabody 1952: tig 8694. Captorhinus— HwQWQ 1956: text-fig. 1951). Captorhinus — Romer 1956: text-fig. 1951), figure is reversed. Captorhinus— Carroll 1969: text-fig 1951). KUVP 8965 Captorhinus— Peabody 1951: text-fig. 2; a composite of the right tarsus based on KUVP 8964, 8965, and 9783. It is not possible to identify which element in the figure is based upon KUVP 8965. This number is represented by an incomplete pes with an associated tibia and femur. Captorhinus— Huene 1956: text-fig. 291 (after text-fig. 2 in Peabody 1951). Captorhinus— Romer 1956: text-fig. 188c (after text-fig 1951), figure is reversed. Captorhinus— Carroll 1969: text-fig. 13, (after text-fig. 1951). KUVP 9783 Captorhinus— Peabody 1951: text-fig. 2; a composite of the right tarsus based on KUVP 8964, 8965, and 9783. The astragalus and calcaneum in the figure is KUVP 9783. Plate 1, fig. B. left astragalus with associated calcaneum. Captorhinus— Huerxe 1956: text-fig. 291 (after text-fig. 1951). Captorhinus— Romer 1956: text-fig. 188c (after text-fig 1951), figure is reversed. Captorhinus— Carroll 1969: text-fig. 13 (after text-fig. 1951). KUVP 9782 Captorhinus— Peabody 1951: pi. 1, fig. A, left astragalus with associated calcaneum. KUVP 9785 Captorhinus-Peabody 1951: pi. 1, fig. C, left astragalus. KUVP 9784 Captorhinus-Peabody 1951: pi. 1, fig. E, two astragali, secfioned to show details of internal structure. KUVP 9610 Captorhinus-Peabody 1952: text-fig. lOA, atlantal centrum. in Peabody in Peabody in Peabody in Peabody in Peabody 22 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 9609 Captorhinus—Peabody 1952: text-fig. lOB, atlantal arch. KUVP 9608 Captorhinus—Peahody 1952: text-fig. IOC, astragalus. KUVP 14753 Captorhinus-Fox 1964: text-fig. 4; listed by Fox as KUVP 8963. Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 21.2; er- roneously listed as KUVP 14756. Posterior part of left mandibular ramus. KUVP 9978 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 1. Captorhinus aguti— Gaiinty and McKenna 1979: text-fig. 5. Skull. KUVP 14746 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 10.1-10.3. Right palatine. KUVP 14755 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 12.1-12.4. Captorhinus aguti— Knhn 1969: pi. 18.7-18.10 (after text-fig. 12.1-12.4 in Fox and Bowman 1966). Left epipterygoid. KUVP 14754 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 13.1-13.4. Captorhinus aguti— ¥.\\\\n 1969: pi. 18.13-18.16 (after text-fig. 13.1-13.4 in Fox and Bowman 1966). Right quadrate. KUVP 14750 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 18.1-18.3; incomplete basi-parasphenoid; erroneously cited as KUVP 14749. KUVP 8963 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 20.1-20.2; incomplete right mandibular ramus. KUVP 14756 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 21.1; poste- rior part of left mandibular ramus; erroneously listed as KUVP 14753. KUVP 14744 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 24; scapulo- coracoid, erroneously listed as KUVP 14757. KUVP 14749 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 25; right scapulo-coracoid. KUVP 14745 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 28.1-28.4. Captorhinus aguti-YM\\n 1969: pi. 19.5-19.8, (after text-fig. 28.1-28.4 in Fox and Bowman 1966). Left humerus. KUVP 14752 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 29.1-29.3. Captorhinus aguti— ¥.u\\n 1969: pi. 19.11-19.13, (after text-fig. 29.1-29.3 in Fox and Bowman 1966). Left ulna. KUVP 14748 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 32.1-32.2; left pelvis. KUVP 14747 Captorhinus aguti (Cope)— Fox and Bowman 1966: text-fig. 33.1-33.4. Captorhinus aguti— Kw\\n 1969: pi. 19.1-19.4, (after text-fig. 33.1-33.4 in Fox and Bowman 1966). Left femur. KUVP 8962 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 1A,B; anterior portion of interclavicle; text-fig. 1 E,F, reconstruction based on KUVP 8962 and 65541. KUVP 65541 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 1C,D; stem of inter- clavicle; text-fig. 1E,F, reconstruction based on KUVP 65541 and 8962, but only cited as KUVP 8962. KUVP 65542 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 3C,D; left scapulo- coracoid; text-fig. 5A,B, reconstruction based on MCZ 4345 and KUVP 65542; cited as KUVP 8962. KUVP 65544 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 8A-F; left humerus, cited as KUVP 8962. TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 23 KUVP 65545 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 9A-F; left radius; cited as KUVP 8962. KUVP 65546 Captorhinus aguti (Cope)— Holmes 1977: text-fig. 11 A; ulnare, cited as KUVP 8962. KUVP? Captorhinus aguti (Cope)— Holmes 1977: text-fig. IIB; intermedium. Specimen not relocated; cited as KUVP 8962. KUVP 65548 Captorhinus aguti (Cope)— Holmes 1977: text-fig. UC; radiale. Er- roneously cited as AMNH 2465. KUVP 8962-65, 9608-9610, 9783-85. 9978, 14745-50, 14752, 14754-56, 65541-42, 65544-48 Lower Permian, Wichita Group, Wellington Formation, fis- sure fills within the Ordovician Arbuckle Limestone; Dolese Brothers limestone quarry at Richards Spur just west of U.S. Hwys. 62 and 281, 10.5 miles south of Apache, 6 miles north of Fort Sill, SW'A, Sec. 31, T4N, Rl IW, Comanche County, Oklahoma; collected by F. E. Peabody, 1950, 1951, 1953. Subclass Testudines Linnaeus 1758 Order Casichelydia Gaffney 1975 Suborder Cryptodira (Cope 1868) Superfamily Chelonioidea Baur 1893 Family Protostegidae Cope 1873 Subfamily Protosteginae Wieland 1902 Protostega Cope 1871 Protostega gigas Cope 1871 KUVP 1202 Protostega— Case 1897: pi. 4; pi. 5, figs. 9-11 (enlargements of certain elements from pi. 4). Protostega gigas Cope— Hay 1908: text-fig. 250, (after pi. 4 in Case 1897). Incomplete plastron with nuchal plate and marginalia; specimen is now missing pieces of certain elements figured by Case. Upper Cretaceous, Campanian, Niobrara Formation, Smoky Hill Chalk Member; Charley Bennett Ranch, west of Russell Springs, T13S, R35W, Logan County, Kansas; collected by H. T. Martin. KUVP 4863 Protostega— Case 1897: pi. 6, figs. 12-15, 18; humerus, scapula, cora- coid, pubis, and femur, respectively. Specimens represented by figs. 17 and 19, if KUVP specimens, have not been relocated. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by H. T. Martin in 1894. KUVP 1208 Protostega— Case 1897: pi. 6, fig. 16; many pieces (may not all belong to one individual): ischium, humerus, ribs, plastron. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; 2.5 miles southeast of Elkader, Sec. 11, T15S, R32W, Logan County, Kansas; collected by T. R. Overton and H. T. Martin in 1895. KUVP 1201 Protostega— WiWiston 1902A: text-fig. 1, hind limb, missing some phalangeal elements. Upper Cretaceous, Campanian, Niobrara Formation, Smoky Hill Chalk Member; Charley Bennett Ranch, west of Russell Springs, T13S, R35W, Logan County, Kansas; collected by C. M. Sternberg in 1900. Subfamily Chelospharginae Zangerl 1953a Chelosphargis Zangerl 1953a 24 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Chelosphargis advena (Hay 1908) (Type-species) KUVP 1209 Protostega advena sp. nov.— Hay 1908: p. 199, text-figs. 256-259; holotype; plastron, twelve marginalia, two or three neuralia, and portions of two costal plates. Upper Cretaceous, Senonian or Niobrara Formation, Smoky Hill Chalk Member; exact locality doubtful, Logan County, Kansas. KUVP 1219 Chelosphargis advena (Hay)— Zangerl 1953A: text-fig. 21A-N. Chelosphargis a^v^na— Gaffney 1979: text-fig. 191A-E, (after text-fig. 21A-E in Zangerl 1953A). Skull roof, mandible, hyoid element, marginalia, costal plates, epiplastra, entoplastron, xiphiplastra, scapular fragment, coracoid, and cervical vertebrae, second right costal plate with connecting neural. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality in doubt, but probably collected in Gove County, Kansas, by S. W. Williston in 1891. KUVP 1258 Chelosphargis advena (Hay)— Zangerl 1953A: text-fig. 25A-B; mar- ginalia, posterior portion of carapace and middle part of carapacial disk of an immature individual. Upper Cretaceous, Coniacian or Santonian, Niobrara Formation. Smoky Hill Chalk Member; Martin's Canyon, Sees. 14-15, 22-23, 26, 34-35, T14S, R26W, Gove County, Kansas; collected by H. T. Martin in 1910. Family Toxochelyidae Hay 1902 Porthochelys Williston 1901 Porthochelys laticeps Williston 1901 (Type-species) KUVP 1204 Porthochelys laticeps, gen. et sp. nov.— Williston 1901B: p. 195. pi. 19, figs. 1-2, pis. 20-21, pi. 22, figs. 1-2. Porthochelys laticeps Williston— Hay 1908: text-figs. 231-234, pi. 31, figs. 2,3. Porthochelys laticeps Williston— Zangerl 1953B: text-fig. 83 (these are corrected illustrations after pi. 19, fig. 1-2 in Williston 1901B), text- fig. 84 (after pi. 21 and pi. 22, fig. 2, respectively, in Williston 1901B). Porthochelys laticeps Williston— Sukhanov 1964: text-fig. 419a and b (redrawn from text-figs. 231 and 232 in Hay 1908); text-fig. 419c (after text-fig. 234 in Hay 1908); text-fig. 419d (redrawn from pi. 31, fig. 3 in Hay 1908). Holotype; greater portion of the skull, dentary, entire left side of carapace, entire plastron (except epiplastra and entoplastron) and right humerus. Upper Cretaceous, Coniacian or Santonian, Niobrara Formation, Smoky Hill Chalk Member; on the Saline River, Trego County, Kansas; collected by E. H. Sellards in 1900. Toxochelys Cope 1873 Toxochelys hrachyrhinus Case 1898 KUVP 1215 Toxochelys latiremis Cope— Case 1898: pi. 79; greater portion of skull. Upper Cretaceous. Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by E. P. West. KUVP 8258 Toxochelys latiremis— Case 1898: pi. 80, figs. 1-2. Toxochelxs latiremis Cope— Hay 1908: text-fig. 206 (redrawn from pi. 80, figs. 1,2 in Case 1898). TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 25 KUVP KUVP 1227 1225 KUVP KUVP KUVP 65547 1225, KUVP 1216 KUVP 1212 KUVP KUVP 1203 1212. fig. 1; femur, lower jaw. , fig. 2, pi. 82, fig. 6; femur, pi. 83. figs. Smoky Toxoche Ivs—Williston 1914: text-fig. 113 (after text-fig. 206 in Hay 1908). Scapula and right coracoid. Toxochelys latiremis— Case 1898: pi. 81. Toxochelys latiremis— Case 1898: pi. 81 ischium, ilium, and other fragments. Toxochelys latiremis-Case 1898: pi. 81. figs. 3-5, 10-11 2-4; specimens not relocated. 1227, 8258 Upper Cretaceous, Senonian, Niobrara Formation, Hill Chalk Member; exact locality unknown, western Kansas. Toxochelys latiremis— Case 1898: pi. 81, figs. 6-8; proximal end of humerus, right ilium. Upper Cretaceous, Coniacian, Niobrara Formation, Smoky Hill Chalk Member; 3.5 miles southwest of Castle Rock, Sec. 22-23, T14S, R26W, Gove County, Kansas; collected by S. W. Williston in 1891. Toxochelys latiremis-Case 1898: pi. 82, figs. 1 (KUVP 1216b) and 2 (KUVP 1216a), four elements comprising the first and fourth or fifth finger, and other finger elements. Upper Cretaceous, Senonian. Niobrara Formation, Smoky Hill Chalk western Kansas; collected by H. '8: p. 378. pi. 84, fig. 1; KUVP 1244 Member; exact locality unknown, western Kansas; collected by H. T. Martin. Toxochelys brachyrhinus , sp. nov. — Case figure 2 is a reconstruction of the skull. Toxochelys brachyrhina (error) Case— Hay 1908: pi. 31, fig. 1. Toxochelys latiremis Cope— Zangerl 1953B: pi. 11. lower figures. Holotype; crushed skull with mandible. Zangerl (1953B) considered Toxochelys branchyrhina (error) Case to be a synonym of Toxochelys latiremis Cope. However, the type of T. latiremis is now thought to be from the Pierre Shale Formation and differs morphologically from Niobrara Toxochelys. T. brachyrhimis is used here as the valid name for all Niobrara Toxochelys (R. Hirayama, personal communication, 1985). Toxochelvs latiremis Cope— Williston 1901 B: pi. 18; skull. Toxochelys— Bennett 1984: text-fig. 83B (after pi. 18 in Williston 1901B). 1203 Upper Cretaceous, Coniacian, Niobrara Formation, Smoky Hill Chalk Member; 3.5 miles southwest of Castle Rock, Sees. 22-23, T14S, R26W, Gove County, Kansas; collected by S. W. Williston in 1891. Toxochelvs latiremis Cope— Zangerl 1953B: text-fig. 73, reconstruction of plastron, based on KUVP 1244; pi. 21. Toxochelys latiremis Cope— Mlynarski 1976: text-fig. 38 (after text-fig. 73 in' Zangerl 1953B). Carapacial and plastral elements, vertebrae, limb bones, and anterior portion of the skull of a juvenile individual, in matrix. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Toxochelys latiremis (Hay 1905) When originally described by Hay in 1905, T browni was placed in the genus Porthochelys Williston. Zangerl (1953B) considered it as belonging to the genus Toxochelys, as T browni (Hay). He also referred the following KUVP material, identified by Wagner (1898) as T latiremis Cope, to T. browni (Hay). According to R. Hirayama (personal communi- cation, 1985) the type material of T latiremis Cope 1873 is probably from 26 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY the Pierre Shale Formation and is indistinguishable from T. browni (Hay 1905). T. browni is therefore a junior synonym of T. latiremis. KUVP 1293 Toxochelys latiremis Cope— Wagner 1898: text-fig. 1 ; posterior portion of the skull (cited as KUVP 1221); not relocated. Upper Cretaceous, Campanian, Pierre Shale Formation, Sharon Springs Shale Member; Eagle Tail Creek, near Sharon Springs. T13S. R40W, Wallace County, Kansas; collected by S. W. Williston in 1891. KUVP 1294 Toxochelys latiremis Cope— Wagner 1898: text-t~ig. 2; fragment of the lower jaw (cited as KUVP 1222); not relocated. Upper Cretaceous, Campanian, Pierre Shale Formation, Sharon Springs Shale Member; near Lisbon, Sec. 8, T12S, R36W, Logan County, Kansas; collected by Sydney Prentice in the summer of 1898. Ctenochelys Zangerl 1953b Ctenochelys sp. KUVP 2050 Toxochelys procax Hay— Hay 1908: text-fig. 228; isolated mandible. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Kansas. Ctenochelys stenoporus (Hay 1905) Case (1898) identified the following material as Toxochelys serrifer Cope. Hay (1905) reexamined the specimens and determined that Case, by not having access to Cope's type material of T serrifer had misidentified the KUVP material. Hay considered it a new species. T stenoporus Hay and specified that any bones bearing number KUVP 2060 should be consid- ered the type material for this form. KUVP 1205 Toxochelys serrifer Cope— Hay 1898: text-figs. 1-3 (the anteriormost neural in text-fig. 2 is missing.) Toxochelys serrifer Cope— Case 1898: pi. 80, figs. 6 coracoid (lower half now missing), 8,9; specimen in fig. 8 not relocated; pi. 82, figs. 12,13, pi. 83, fig. 1. Ctenochelys stenopora (error) (Hay)— Zangerl 1953B: text-fig. 108. Fragmentary posterior left half of carapace, consisting of five costals, third and fifth through eighth neurals, marginalia, and pygals, coracoid, humerus, ilium, phalanges, carpal bones. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown; western Kansas. KUVP 2060 Toxochelys serrifer Cope-Case 1898: pi. 80, figs. 3,4,5 (specimen fig. 5 not relocated), pi. 82, figs. 3,4,5. Toxochelys stenoporus sp. nov.— Hay 1905: pi. 180, text-figs. 8-11. Hay erroneously stated that the marginalia figured by Case (1898), in pi. 83, fig. 1, bear the number KUVP 2060. These three bones bear the old number KUVP 1270, but actually belong to KUVP 1205. In his 1908 tome, he stated that plates 80, 82 and 83 in Case (1898) are a composite of KUVP 2060 and KUVP 1270, but that all of this material belongs to the same species. According to Hay the elements in pi. 80, fig. 4 and pi. 83, fig. 1, belong to KUVP 1270. Toxochelys stenopora (error) Hay— Hay 1908: text-figs. 214-217 (after text-figs. 8-1 1 in Hay 1905) text-figs. 218-220. Hay specified that any bone bearing the number KUVP 2060 should be regarded as belonging to the holotype of T stenopora (error) Hay. Ctenochelys stenopora (error) (Hay)— Zangerl 1953B: text-fig. 107 (re- construction of the plastron based on KUVP 2060, pi. 80, figs. 3-5 in Case (1898), and text-fig. 220 in Hay (1908)). TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 27 Holotype; fragmentary skull, mandible, dentary, hypoplastron, xiphiplastron, left hyoplastron, marginalia, nuchal, limb elements. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by S. W. Williston in 1891. KUVP 1228 Toxochelys sp. indet.-Hay 1908: pi. 30, figs. 1,2. Ctenochelys stenopora (error) (Hay)— Zangerl 1953B: text-fig. 108. Zangerl suggested this material may belong to Ctenochelys procax (Hay), since it differs substantially from KUVP 1205 (C. stenoporus). Fifteen marginalia, fragments of costals, and a series of neuralia, including two suprapygals. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; near Monument Rock, Gove County, Kansas; collected by H. T. Martin. Family Cheloniidae Gray 1825 Desmatochelys Williston 1894 Desmatochelys lowi Williston 1894 (Type-species) KUVP 1200 Desmatochelys Lowii Williston (error)— Williston 1894B: p. 5, text-figs, on pp. 12 (text-fig. 8a, 8c, 8-11 not relocated), 15 (not relocated), 16; pis. 2-4; pi. 5, figs. 1,2a, 2, 3 (specimens of figs. 1,2a not relocated), pi. 6, figs. l,la,lb,2-4. Desmatochelys lowii Williston (error)— Williston 1898C: pis. 73-75 (after pis. 2-4 in Williston 1894); pi. 76, fig. 1 (after text-fig. on p. 16 in Williston 1894), fig. 2 (after text-fig. on p. 15 in Williston 1894, not relocated), figs. 8, 8a-c, 9-11 (after text-figs, on p. 12 in Williston 1894, only elements in figs. 8,8b relocated); pi. 77, figs. 1-3, fig. 1 is inverted (after pi. 5, figs. 1-3 in Williston 1894, elements of figs. 1,2a not relocated); pi. 78, figs. l,la-b,2-4 (after pi. 6, figs. l,la-b, 2-4 in Williston 1894). Desmatochelys Lowii Williston (error)— Wieland 1900: text-fig. 13 (after pi. 4 in Williston 1894). Desmatochelys lowi Williston— Hay 1908: text-figs. 238-243 (element in fig. 242 not relocated). Desmatochelys lowii (error)— Williston 1914: text-fig. 122 (after text- figs. 238,239 in Hay 1908). Desmatochelys lowi Williston— Miiller 1968: text-fig. 82a, b (after text- figs. 238',239 in Hay 1908). Desmatochelys lowi Williston— Zangerl and Sloan 1960: text-fig. 4, sketch of skull made by T. H. Eaton. Desmatochelys lowi Williston— Sukhanov 1964: text-fig. 425a and b (after text-figs. 238 and 239 in Hay 1908). Desmatochelys lowii (error)— Mlynarski 1976: text-fig. 50 (after text-fig. 82a,b in Muller 1968). Desmatochelys lowi— Gaffney 1979: text-fig. 207 (after text-fig. 4 in Zangerl and Sloan 1960); text-fig. 208 (after pi. 4 in Williston 1894). Porthochelys (error)— Bennett 1984: text-fig. 83A, skull and lower jaw. Holotype; complete skull with lower jaw, metacarpals and other pha- langeal elements, pygal and marginalia, fragments of some carapacial elements, right humerus, coracoid, radius and ulna, scapula-precoracoid, right pelvic bones, femur, and caudal vertebrae. Upper Cretaceous, Turonian, Greenhorn Formation, Pfeifer Shale or 28 KUVP 23488 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Jetmore Chalk Member; near Fairbury, Jefferson County, Nebraska; collected by B. Schrantz and N. V. Eaton in 1893. Superfamily Testudinoidea Baur 1893 Family Chelydridae Swainson 1839 Emarginachelys Whetstone 1978 Emarginachelys cretacea Whetstone 1978 (Type-species) Emarginachelys cretacea n. sp. — Whetstone 1978: p. 544, text-figs. 5-8,10-19; holotype; carapace, plastron, skull, missing lower jaw, right stapes, fragmentary hyoid, portions of the right and left forelimbs and girdles and left hind limb and girdle, right ilium, several cervical and caudal vertebrae. Upper Cretaceous, Maestrichtian, Hell Creek Formation; SW'A, NW'/4, Sec. 35, T21N, R37E, Garfield County, Montana; collected by D. and S. Rasmussen and J. Chorn in the summer of 1971. Family Emydidae Gray 1825 Subfamily Emydinae Gray 1825 Chrysemys Gray 1844 Chrysemys hilli (Cope 1878) KUVP 7676 Chr\senn's limnodxtes, new species— Galbreath 1948: p. 269; pi. 1, figs. A,B. Chrysemys limnodytes i = P. hilli)— Adler 1968: text-figs. ID, 2. Holotype; fragmental anterior portion of carapace, left side of plastron, and several marginalia. Tertiary, Late Miocene, Laverne Formation, Marl beds; SWVa, Sec. 15, T4N, R25E, Beaver County, Oklahoma; collected by a University of Kansas field party in the summer of 1946. Adler ( 1968) considered Chrysemys limnodytes Galbreath to be a synonym of Pseudemys hilli (Cope). He also stated that Galbreath's original reconstruction of this fossil is incorrect in several respects. McDowell (1964) contended that Pseudemys and Chrysemys should be combined into a single genus,Chrysemys. Zug (1966) and Ernst and Barbour (1972) agreed with McDowell's conclusions. Emydoidea Gray 1870 Emydoidea blandingi (Holbrook 1838) KUVP 6478 Emys twentei sp. nov.— Taylor 1943: p. 250, pi. 20; holotype, posterior three-fourths of the carapace. Quaternary, Odee Formation, Pleistocene high terrace sands; north side of the Cimarron River, 9 miles south of State Lake, probably Sec. 33, T34S, R29W, XI Ranch, Meade County, Kansas; collected by Jack Twente in the summer of 1942. Schultz (1965) and Gehlbach (1965), both referred to this taxon as Emydoidea twentei Taylor. Preston and McCoy (1971), with additional fossil Emydoidea and Emys material at hand, concluded that Emys twentei Taylor should be referred to Emydoidea blandingi (Holbrook). Family Testudinidae Gray 1825 Hesperotestudo Williams 1950 Hesperotestudo riggsi (Hibbard 1944) Williams (1950) synonymized Testudo riggsi Hibbard into the genus Gopherus, as G. riggsi (Hibbard). Oelrich (1957), considering T. riggsi TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 29 Hibbard to be a synonym of T. turgida, placed both in the genus Geochekme (subgenus Hesperotestudo), and stated that Geochekme riggsi (Hibbard) represents a phyletic intermediate between G. turgida (Cope) and G. incisa (Hay). Bramble in an unpublished work cited by Preston (1979) argued that Geochelone does not occur in the Western hemisphere. Therefore Preston used Hesperotestudo as the generic name for New World "Geochelone. " KUVP 6789 Testudo riggsi sp. nov.— Hibbard 1944: p. 72, text-figs. 1.2. Testiido turgida Cope— Oelrich 1957: text-figs. 2,3. Geochelone riggsi (Hibbard)— Auffenberg 1963: text-fig. 31 (after text- figs. 2,3 in Oelrich 1957), erroneously listed as a paratype. Holotype; a nearly complete carapace and plastron. KUVP 6790 Testudo riggsi sp. nov.— Hibbard 1944: p. 72; paratype; complete plastron and nearly complete carapace. KUVP 6789, 6790 Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); Sec. 35, T34S, R31W, XI Ranch, south side of Cimarron River, Seward County, Kansas; collected by C. W. Hibbard and T. McLaughlin in June, 1943. Hesperotestudo gilberti (Hay 1899) KUVP 1245 Xerobates (?) undata Cope-Gilbert 1898: text-figs. 1-4. Testudo gilbertii— Hay 1899: p. 349, no figures. Testudo gilberti Hay— Hay 1908: pi. 80, figs. 2-5. Holotype; skull with lower jaw, pectoral girdle, pelvic girdle, limbs, cervical vertebrae. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); Phillips County, Kansas; collected by E. P. West and T. R. Overton. Gilbert (1898) tentatively identified this material as Xerobates undata Cope. Hay (1899) placed it in the genus Testudo as a new species, based on the figures provided by Gilbert. Auffenberg (1963, 1974) placed it in the genus Geochelone (subgenijs Hesperotestudo), and suggested that it is probably a synonym of Geochelone osborniana (Hay). Hesperotestudo campester (Hay 1903) KUVP 9399 Testudo rexroadensis sp. nov.— Oelrich 1952: p. 301, text-figs. 2,3; paratype; fragments of a carapace and a nearly complete plastron. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); W'/:, SW'A, Sec. 22, T33S, R29W, Meade County, Kansas; collected by University of Kansas field party in 1940. Auffenberg (1963, 1974) considered Testudo rexroadensis Oelrich to be a member of the genus Geochelone (subgenus Hesperotestudo) and a synonym of Geochelone campester (Hay), closely related to G. orthopygia (Cope). Hesperotestudo orthopygia (Cope 1878) KUVP 1262 Caryoderma snovianum Cope, sp. nov.— Cope 1886: p. 1045. Caryoderma snovianum Cope — Cope 1889: pi. 32, figs. 1-17. The specimens reproduced in figs. 8,15-15a, and 16-16a, have not been relocated. Holotype; various dermal ossicles and a single phalangeal element figured by Cope (1889). Material not figured includes more dermal ossicles, fragmentary limb bones, phalangeal, and carapacial elements. Tertiary, Late Miocene, Ogallala Formarion (Hemphillian land mammal age); Decatur County, Kansas; collected by C. H. Sternberg in 1878. Cope described this material in 1886 as, ". . . a portion of the dermal 30 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY skeleton of an armadillo, probably in the family Glyptodontidae ..." Later (1889) he figured the material. In 1898E. Williston correctly identified these bones as the carapacial osteoderms of a large tortoise and provisionally placed the material in the genus Xerobates, "... specifi- cally probably identical with Testudo iindata Cope." Hay (1902) listed this species as Testudo snoviana (Cope), but later placed Caryoderma snovianum Cope in the synonymy of Testudo orthopygia (Cope). Williams (1950) placed T orthopygia (Cope) in the genus Geochelone Fitzinger. KUVP 16340 Testudo— 'WxWxdLms 1975: text-fig. 20; nearly complete carapace and plastron, pelvis, and limb bones. Tertiary, Late Miocene, Ogallala Formation (Hemphillian land mammal age); road cut north of Bluff Creek, Beckerdite Ranch, near center of Sec. 8, T31S, R22W. Clark County, Kansas; found by L. R. Chilson and collected by O. T. Bonner in 1969. Subclass Sauropsida Huxley 1864 Superorder Lepidosauria Haeckel 1866 Order "Eosuchia" Broom 1914 Family Petrolacosauridae Peabody 1952 Petrolacosaurus Lane 1945 Petrolacosaurus kansensis Lane 1945 (Type-species) KUVP 1424 Petrolacosaurus kansensis, gen. et sp. nov.— Lane 1945A: p. 381, holotype; text-fig. 1, nearly complete hind limb lacking a portion of the distal articulation of the femur and the proximal half of the tibia and fibula. Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 8, adult right femur; text-fig. 9, right pes of adult. Petrolacosaurus Peabody (error)— Huene 1956: text-fig. 460D (after text- fig. 9 in Peabody 1952). Petrolacosaurus kansensis Lane — Miiller 1968: text-fig. 170B (after text- fig. 460D in Huene 1956). Petrolacosaurus kansensis Lane — Reisz 1981: text-fig. 24A, reconstruc- tion of left pes, based primarily on KUVP 1424 and 9951b. KUVP 1423 Podargosaurus hibbardi. gen. et sp. nov.— Lane 1945A: p. 385; holotype; text-fig. 3, nearly complete right forelimb (erroneously listed as the left forelimb). Associated type material includes the crural, tarsal, and pedal portions of a hind limb; a second forelimb, several dorsal vertebrae; thirty or more caudal vertebrae and several ribs. Petrolacosaurus kansensis Lane — Peabody 1952: text-fig. 6C, subadult right forelimb. Peabody (1952) placed this type in the synonymy of Petrolacosaurus kansensis Lane. He also stated that the limb bones figured by Lane ( 1945 A) as the type of Podargosaurus hibbardi cannot be associated with the remaining portions of the skeleton mentioned above. Petrolacosaurus Peabody (error)— Huene 1956: text-fig. 460C (after text- fig. 6C in Peabody 1952; image reversed). Petrolacosaurus kansensis Lane — Miiller 1968: text-fig. 170A (after text- fig. 460C in Huene 1956; image reversed). Petrolacosaurus kansensis Lane — Reisz 1981: text-fig. 21A,B, recon- struction of right manus. based primarily on KUVP 1423, 8355, and 9957b. KUVP 8351 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 1A,B, skull and TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 31 associated cervical vertebrae; pi. lA, immature skull and cervical vertebrae, with associated cupule ot pteridosperm; pi. 2A, immature skull and cervical vertebrae (photograph of specimen represented in text-fig. 1A,B). Petrolacosaurus Peabody (error)— Huene 1956: text-fig. 460B (after text- fig. lA in Peabody 1952). KUVP 1427 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 2, post-cranial skeleton of young adult; text-fig. 3A, cervical vertebrae (details of proatlas from KUVP 8351); text-fig. 3C, fifteenth caudal vertebra; text-fig. 11, reconstruction of skull and skeleton based primarily on KUVP 1427; pi. 2B, forelimb, with associated cone of conifer, Walchia. Petrolacosaurus Peabody (error)— Huene 1956: text-fig. 460A (after text- fig. 1 1 in Peabody 1952). Petrolacosaurus kansensis Lane— Tatarinov 1964B: text-fig. 286 (after text-fig. 11 in Peabody 1952). Petrolacosaurus kansensis Lane— Miiller 1968: text-fig. 169 (after text- fig. 11 in Peabody 1952). KUVP 9954 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 3B; pi. IB, three anterior dorsal vertebrae. Only the anterior (left) most vertebra is figured. Cited by Peabody 1952 as "loaned from Nebraska Geological Survey (no number)." KUVP 1428 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 4, subadult skeleton with fragmentary skull. KUVP 1426 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 6A, immature right hind limb, lacking femur. KUVP 1429 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 6B, immature left forelimb. KUVP 8355 Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 7, right carpus of young adult; pi. IC, carpus (photograph of specimen represented in fig. 7). Petrolacosaurus— CdLvroW 1977: text-fig. IIB (after text-fig. 21 A, re- versed, in Reisz 1981). Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 21 A, B, recon- struction of right manus, based primarily on KUVP 8355, 1423 and 9957b. KUVP 33607 Petrolacosaurus kansensis— CarroW 1975: text-fig. 13 (after text-fig. 2, central figures in Reisz 1977). Petrolacosaurus kansensis— Reisz 1977: text-fig. 2, lower left-hand figure, reconstruction of occipital view of skull, based mainly on KUVP 33607 and 9951; text-fig. 2, central figures, reconstruction of dorsal and palatal views of skull, based mainly on KUVP 33607, 33606, and 9952; text-fig. 2, top figure, based mainly on KUVP 33607 and 9952. Petrolacosaurus— McFarlmd, et al. 1979: fig. 13-8A (redrawn from fig. 2, top figure in Reisz 1977). Petrolacosaurus— Kuhn-Schnyder 1980: fig. 9.8C (redrawn from fig. 2, top figure in Reisz 1977). Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 1 (reconstructed skull only), based primarily on KUVP 33607 and 9952; text-fig. 2 A, top figure (after text-fig. 2, top figure, in Reisz 1977); text-fig. 2B, left mandible, based upon KUVP 33607 and 9952; text-fig. 3, (after text-fig. 2, central figures in Reisz 1977); text-fig. 4 (after text-fig. 2, 32 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY lower left-hand figure in Reisz 1977); text-figs. 12,13, mature skull and lower jaws with articulated cervical vertebrae and isolated cleithrum; text-fig. 14A, left proatlas; text-fig. 14E. atlas-axis com- plex, based on KUVP 33607 and 33604. KUVP 33606 Petrolacosaunis kansensis—CdiXxoW 1975: text-fig. 13 (after text-fig. 2, central figures in Reisz 1977). Petrolacosaurus kansensis— Reisz 1977: text-fig. 2, central figures based primarily upon KUVP 33606, 33607, and 9952. Petrolacosaaunis kansensis— Reisz 1981: text-fig. 3 (after fig. 2, central figures in Reisz 1977); text-fig. 11, mature skull and left lower jaw; text-fig. 22A,B. left femur, based mainly on KUVP 33606 and 9951; text-fig. 24C, right astralagus. KUVP 9951 Petrolacosaunis kansensis— CarroW 1975: text-fig. 13 (after text-fig. 2, central figures in Reisz 1977). Petrolacosaurus kansensis— Reisz 1977; text-fig. 2, lower left hand figure, reconstruction of occipital view of skull, based mainly on KUVP 9951 and 33607. Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 6, mature skull and caudal vertebrae; text-fig. 7, braincase; text-fig. 14G, third cervical vertebra, based on KUVP 9951 and 9956; text-fig. 14J, sixth cervical vertebra, based on KUVP 9951 and 9956; text-fig. 14H,I, fourth and fifth cervical vertebrae; text-fig. 15A-C, dorsal vertebrae 7,8,13,14; text-fig. 15D, vertebrae 17, 18; text-fig. 16A, atlantal rib; 16B axial rib, text-fig. 16G, rib 15. KUVP 9952 Petrolacosaurus kansensis— CarroW 1975: text fig. 13 (after text-fig. 2, central figures in Reisz 1977). Petrolacosaurus kansensis— Reisz 1977: text-fig. 2 (reconstructed skull only), based upon KUVP 9952 and 33607. Petrolacosaurus— Kuhr\-Schnyder 1980: text-fig. 98C (redrawn from text-fig. 2, top figure in Reisz 1977). Petrolacosaurus kansensis Lane— Reisz 1981 : text-fig. 2 A (after fig. 2 top figure in Reisz 1977); text-fig. 2B, left mandible, based upon KUVP 9952 and 33607; text-fig. 8, skull; text-fig. 3 (after text-fig. 2, central figures in Reisz 1977). KUVP 33608 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 5A,B, immature skull and lower jaws. Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 10, skull and lower jaws of young adult. KUVP 33602 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 9A, left maxilla; 9B, cross section. KUVP 9959a Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 9C, quadrate and quadratojugal (erroneously labeled left parietal). KUVP 33603 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 9D, left parietal (erroneously labeled quadrate and quadratojugal). KUVP 33604 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 14B, atlantal intercentrum; text-fig. 14C, left atlantal neural arch; text-fig. 14D, atlantal centrum and fused axial intercentrum; text-fig. 14E, atlas-axis complex, based on KUVP 33604 and 33607; text-fig. 14F, axis; text- fig. 16A,B, atlantal and axial ribs, based primarily on KUVP 33604 and 9951. KUVP 9956 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 14G, third cer- vical vertebra, based on KUVP 9956 and 9951; text-fig. 14J, sixth cervical vertebra, based on KUVP 9956 and 9951; text-fig. 16C, fifth TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 33 rib; text-fig. 16D, sixth (cervical) rib; text-fig. 16E, seventh (tlrst dorsal) rib; text-tig. 16F. eighth rib; text-fig. I6H. anterior dorsal ribs 9-11. KUVP 33606b Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 15E, dorsal ver- tebrae 25 and 26; text-fig. 15G, second sacral vertebra and rib; text- fig. 161, posterior dorsal ribs 23 and 25; text-fig. 16J, first sacral ribs and vertebra. KUVP 33605 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 15F, first sacral neural arch and ribs. KUVP 33606c Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 15H, L anterior caudal vertebrae and associated ribs; text-fig. 18B. right iliac blade. KUVP 9957a Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 17A, left scapulo- coracoid in articulation with limb; text-fig. 19A-D, right humerus, based mainly on KUVP 9957 and 33606a; text-fig. 20A, left radius; text-fig. 20B, left ulna (originally published as KUVP 9957). KUVP 9959b Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 17B, immature interclavicle. KUVP 9958 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 17C, immature left clavicle. KUVP 33609 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 17D, right scapulo-coracoid. KUVP 9961 Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 18A. left pelvis, slightly crushed. KUVP 9951c Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 18C, left pelvis, slightly crushed; 18C originally published as KUVP 9951. KUVP 33606a Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 19A-D, right humerus, based primarily on KUVP 33606a and 9957. KUVP 9957b Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 21B. reconstruc- tion of right manus, based primarily on KUVP 9957b, 1423, and 8355. KUVP 995 Id Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 22A,B, recon- struction of left femur, based mainly on KUVP 995 Id (originally published as KUVP 9950) and 33606. KUVP 9951b Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 23A-C, left tibia; text-tig. 23D, left fibula; text-fig. 24A, reconstruction of left pes, based mainly on KUVP 9951b and 1424; text-fig. 24D, metatarsals. KUVP 9951a Petrolacosaurus kansensis Lane— Reisz 1981: text-fig. 24B, left tarsus. KUVP 1423-4, 1426-9, 8351, 8355, 8357, 9951-2, 9954, 9956-9, 9961, 33602-9 Upper Pennsylvanian, Missourian, Stanton Formation, Rock Lake Member; six miles northwest of Garnett, Sec. 32, T19S, R19E, Anderson County, Kansas. KUVP 1 1473 Order Squamata Oppel 1811 Suborder Sauria Macartney 1802 Infraorder Iguania Latreille 1825 Family Iguanidae Gray 1827 Leiocephalus Gray 1827 Leiocephalus etheridgei Pregill 1981 Leiocephalus etheridgei, new species— Pregill 1981: p. 35, paratypes; a single right, and six left dentaries. Quaternary, Late Pleistocene; Cuevo del Perro, Ciales Valley, about 2 kilometers south of Barrio de Barahona, Municipio de Morovis, 18°20'57"N, 66°26'47"W, Puerto Rico; collected by J. W. Bee in 1957. 34 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Leiocephalus partitiis Pregill 1981 KUVP 64695 Leiocephalus partitus, new species— Pregill 1981: p. 39; paratype; a single right dentary. This specimen originally bore the same number as L. ethehdgei. Quaternary, Late Pleistocene; Cuevo del Perro, Ciales Valley, about 2 kilometers south of Barrio de Barahona, Municipio de Morovis, 18°20'57"N, 66°26'47"W, Puerto Rico; collected by J. W.« Bee in 1957. Phrynosoma Wiegmann 1828 Phrynosoma adinognathus Rickart 1976 KUVP 25258 Phrynosoma adinognathus sp. nov.— Rickart 1976: p. 65; text-fig. lb; holotype; proximal two-thirds of a right dentary. KUVP 25259 Phrynosoma adinognathus sp. nov.— Rickart 1976: p. 65; paratype; proximal half of left dentary. KUVP 25260 Phrynosoma adinognathus sp. nov.— Rickart 1976: p. 65; text-fig. Ic; paratype; distal half of right dentary. KUVP 25261 Phrynosoma adinognathus sp. nov.— Rickart 1976: text-fig. la; referred material, a nearly complete frontal lacking posterolateral arms. KUVP 25258-61 Quaternary, Pleistocene, Crooked Creek Formation, Borchers local fauna; east of State Lake on east side of Crooked Creek, NW'A, NE'A, Sec. 21, T33S, R28W, Meade County, Kansas; collected by C. W. Hibbard in 1939 and 1940. Phrynosoma cf. cornutum (Harlan 1825) KUVP 5099 Eumecoides hibbardi, n. sp.— Taylor 1941A: p. 173, text-fig. 5A-D; holotype; a nearly complete right dentary. Phrynosoma cornutum— Estes 1983: text-fig. 8G (after text-fig. 5A-D in Taylor 1941 A). KUVP 5115 Eumecoides mylocoehis, n. sp.— Taylor 1941A: p. 174, text-fig. 6A-D, holotype; left dentary, missing posterior portion. KUVP 5099, 51 15 Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Taylor (1941 A) originally referred this material to the family Scincidae. Etheridge (1960), on the basis of additional material, considered both Eumecoides hibbardi Taylor and E. mylocoehts Taylor to be conspecific with, though slightly different from, Phrynosoma cornutum (Harlan). Infraorder Scincomorpha Camp 1923 Family Scincidae Gray 1825 Eumeces Wiegmann 1834 Eumeces striatulatus Taylor 1941 KUVP 5079 Eumeces striatulatus, n. sp.— Taylor 1941A: p. 171, text-figs. 3A-D; holotype; fragmentary right dentary. Eumeces striatulatus— Esles 1983: text-fig. 27E (after text-fig. 3A-D in Taylor 1941 A). KUVP 65694 Eumeces striatulatus, n. sp.— Taylor 1941A: text-figs. 4A,B; referred material, middle portion of right dentary. KUVP 5079, 65694 Tertiary, Pliocene. Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade, NE'/4, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Family Teiidae Gray 1827 Cnemidophorus Wagler 1839 Cnemidophorus bilobatus Taylor 1941 TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 35 KUVP 5084 Cnemidophorus bilobatus, n. sp.— Taylor 1941A: p. 167, text-fig. lA-C; holotype; poilion of left dentary, containing eight teeth. Cnemidophorus bilobatus — Estes 1983: text-fig. 2 ID (after text-fig. lA-C in Taylor 1941 A). KUVP 5081 Cnemidophorus bilobatus, n. sp.— Taylor 1941A: text-fig. 2B,C; referred material, fragment of anterior part of right maxilla. KUVP 5091 Cnemidophorus bilobatus, n. sp.— Taylor 1941 A: text-fig. 2A; referred material, anterior tip of left dentary. KUVP 5084, 5081, 5091 Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age); about 16 miles southwest of Meade, NE'A, SWA, Sec. 22, T33S, R29E, Meade County, Kansas; collected by C. W. Hibbard and party in 1938. Infraorder Anguimorpha Fiirbringer 1900 Superfamily Diploglossa ( = Anguinoidae) Cope 1864 Family Anguidae Gray 1825 Subfamily Glyptosaurinae McDowell and Bogert 1954 Helodermoides Douglass 1903 Helodermoides tuberculatus Douglass 1902 KUVP 10698 Helodermoides tuberculatus Douglass— Sullivan 1979: text-fig. 10i,j; nearly complete left frontal and posterior part of left prefrontal. Peltosaurus granulosus— Estes 1983: text-fig. 37 A (after text-figs. 76, 73, 78 in Gilmore 1928). Tertiary, Oligocene, White River Formation (Orellan land mammal age), Cedar Creek Member; East Pawnee Butte, Sec. 22, TION, R59W, Weld County, Colorado; collected by R. W. Wilson on July 11, 1956. Peltosaurus Cope 1873 Peltosaurus granulosus Cope 1873 KUVP 1280 Peltosaurus granulosus Cope— Gilmore 1926: pi. 15, figs. 1-3; skull and dermal scutes. Peltosaurus granulosus Cope — Gilmore 1928: text-figs. 73, 75-78; skull. Peltosaurus granulosus Cope— Piveteau 1955: text-fig. IIB on p. 624 (after text-fig. 75 in Gilmore 1928). Peltosaurus granulosus — Estes 1983: text-fig. 37A (after text-figs. 76, 73, 78 in Gilmore 1928). Tertiary, Oligocene, Brule Formation (Orellan land mammal age), Orella Member; Hat Creek Basin, 1.5 miles west of Anderson Ranch, Sioux County, Nebraska; collected by E. R. Hall in 1922. Gilmore listed this specimen as KUVP 1208 in the figure legends of his 1926 paper (a number currently occupied by the marine turtle Protostega gigas), and variably as KUVP 1208 and 1280 in his 1928 paper. He also listed KUVP 1280 as referred material and a fragmentary humerus from this same specimen as KUVP 1208 in this paper. The number KUVP 1280 is the correct citation for all of this material. KUVP 7654 Peltosaurus granulosus Cope— Meszoely 1970: text-fig. 16D; skull roof and other skull elements. Erroneously cited as KUVP 620 (this is the field number). Tertiary, Oligocene, White River Formation (Orellan land mammal age); WV:, Sec. 7, TUN, R53W. Logan County, Colorado: collected by R. R. Camp on July 31, 1946. Family Anguidae incertae sedis Odaxosaurus Gilmore 1928 36 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Odaxosaunis pawneensis (Gilmore 1928) KUVP 1281 IXestops pawneensis, new species— Gilmore 1928: p. 150; holotype; median section of an articulated skull with the median part of the attached right ramus. Tertiary, Oligocene, White River Formation (Orellan land mammal age); Clyde Ward Ranch, Sec. 28, TUN, R53W, 30 miles north of Sterling, Logan County, Colorado; collected by C. J. Hesse in 1925. Due to the fragmentary nature of this specimen, Gilmore could only tentatively place it in the genus Xestops Cope. Meszoely (1970) erected the new genus, Pancelosaurus as a replacement name for Peltosaurusl pi ger GWvaovQ 1928, P. jepseni Gilmore 1942, and Odaxosaurus obliquus Gilmore 1928, all of which were conspecific, but generically distinct from Peltosaurus Cope, in his opinion. He chose Peltosaurusl piger as the type species of this new genus since it had page priority over Odaxosaurus obliquus in Gilmore's (1928) paper. However, since the genus Odax- osaurus Gilmore (1928) already existed as a replacement name, Mes- zoely's treatment, in effect, synonomized his replacement name Pancelosaurus. Meszoely concluded that IXestops pawneensis Gilmore (1928) should be called Pancelosaurus pawneensis (Gilmore), but the proper designation should be Odaxosaurus pawneensis (Gilmore 1928). Our placement of Odaxosaurus in the Anguidae, incertae sedis, follows Sullivan's (1979) conclusions. KUVP 9734 Odaxosaurus cf. piger (Gilmore)— Sullivan 1981: text-fig. 4.2, anterior portion of frontal complex. Tertiary, Middle Paleocene, Nacimiento Formation (Torrejonian land mammal age); NW'A, Sec. 14, T27N, RllW, San Juan County, New Mexico; collected by R. W. Wilson field party in 1950. Superfamily Platynota Baur 1890 Family Helodermatidae Bonaparte 1840 Heloderma Wiegmann 1829 Heloderma matthewi Gilmore 1928 KUVP 7652 Heloderma matthewi Gilmore— Yatkola 1976: text-fig. IC-E; complete right maxilla. Specimen probably lost. Tertiary, Middle Oligocene, White River Formation (Orellan land mam- mal age); center of WV2. Sec. 7, Tl IN, R53W, Logan County, Colorado; collected by R. R. Camp on August 1, 1946. Family Mosasauridae Gervais 1853 Subfamily Mosasaurinae Williston 1897 Clidastes Cope 1868 Clidastes propython Cope 1 869 KUVP 1022 Clidastes velox Marsh— Williston and Case 1892: pi. 2, left front paddle (image reversed); pi. 3, left hind paddle (now missing the fibula). Clidastes velox Marsh— Williston 1893B: pi. 3. figs. 1-3; restoration of complete skeleton based primarily on KUVP 1022. Williston stated that the palatals and vomers were so distorted in KUVP 1022, that he had to refer to a specimen of C. tortor Cope ( = C. propython Cope), for these cranial elements. Clidastes ve/ojc— Williston 1895 A: pi. 17, fig. 1 (after pi. 3, fig. 2 in Williston 1893B). Clidastes velox Marsh— Willi.ston 1898B: pis. 10-12, reconstruction of skull, based primarily on KUVP 1022; pi. 33 (after pi. 2 in Williston TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 37 and Case 1892); pi. 34 (after pi. 3 in Williston and Case 1892, but with the pelvic girdle elements rearranged); pi. 27, ?figs. 3,4, radii, ?figs. 3A, 4A, ulnae; ?pl. 31, fig. 6, coracoid and scapula; pi. 60, fig. 6, right quadrate, fig. 7. left quadrate (these specimens have not been relocated); pi. 62, fig. 3, nearly complete skeleton (image reversed); pi. 72 top figure, restoration of complete skeleton. Clidastes—WiWiston 1914: text-fig. 72, top figure (redrawn from pi. 10 in Williston 1898B); text-fig. 74 (redrawn from pi. 2 in Williston and Case 1892, image reversed). Clidastes—LuW 1917: text-fig. 69 (after Williston 1914, text-fig. 72, top figure). Clidastes velox Marsh— Zittel 1918: text-fig. 359A (after pi. 10 in Williston 1898). Tylosaimis proriger Cope (error)— Zittel 1918: text-fig. 359B (redrawn from pi. 11 in Williston 1898B); erroneously labled Tylosaimis proriger. Clidastes— ^'xWx^ion 1925: text-fig. 54, top figure (after text-fig. 72, top figure in Williston 1914); text-fig. 146 (after text-fig. 74 in Williston 1914). Tylosaimis (error)— Romer 1933: text-fig. 142A (redrawn from pi. 11 in Williston 1898B). Second edition 1945 text-fig. 169A; third edition 1966 text-fig. 204 A; erroneously labled Tylosaimis. Clidastes-Romer 1933: text-fig. 142B (after Williston 1914, text-fig. 72). Second edition 1945 text-fig. 169B; third edition 1966 text-fig. 204B. Clidastes velo.x— Lane 1946: text-fig. 7 (redrawn from pi. 60, fig. 7 in Williston 1898B). Specimen not relocated. Tylosaimis (error)— Romer 1956: text-fig. 65A (redrawn from pi. 11 in Williston 1898B); erroneously labeled Tylosaimis. Clidastes— Romer 1956: text-fig. 65C (after pi. 3 in Williston and Case 1892); text-fig. 182f, pectoral elements (after pi. 2 in Williston and Case 1892). Clidastes velox Marsh-Callison 1967: text-figs. 1, 9 (after pi. 10 in Williston 1898B). Clidastes— Peyer 1968: text-fig. 102 (taken from Romer 1945, text-fig. 169). Clidastes— Bennett 1984: text-fig. 85B (redrawn from pi. 72, top figure, Williston 1898B). Nearly complete skeleton. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; Butte Creek, Logan County, Kansas; collected by S. W. Williston in the summer of 1891. KUVP 1026 Clidastes Westii, n. sp. -Williston and Case 1892: p. 29-30, pi. 4; nearly complete right front paddle; pi. 5, right hind paddle, missing the terminal phalangeal elements; pi. 6, eighteenth dorsal vertebra. Holotype. Clidastes westii Williston (error)-Williston 1898B: pi. 23. top figure, complete left mandible; pi. 27, fig. 1, radius; fig. lA, ulna; pi. 35 (after pi. 4 in Williston and Case 1892); pi. 36 (after pi. 5 in Williston and Case 1892); pi. 39, top figures, distal ends of humerus; pi. 53 (after pi. 6 in Williston and Case 1892), listed as the thirty-eighth vertebra; pi. 60, fig. 4, right quadrate, not relocated. Clidastes westii (error)-Camp 1923: text-fig. 22 (after Williston 1898B. 38 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY pi. 53, fig. 2). Clidastes—V^'iWiston 1925: text-fig. 158B (redrawn from pi. 4 in Williston and Case 1892). Lower jaw, skull parts, 84 vertebrae, ribs, front and hind paddle. Upper Cretaceous. Campanian, Pierre Shale Formation; near McAllister, Logan County, Kansas; collected by C. H. Sternberg. Williston and Case (1892) stated the type material of C. wesri came from, "... the uppermost of the Niobrara beds, in the vicinity of the old town of Sheridan" (now McAllister), in Logan County, Kansas. KUVP 1000 Clidastes tortor Cope— Williston 1898B: pi. 23, lower figure, complete right mandible; pi. 27, fig. 2, radius; fig. 2a, ulna, not relocated; pi. 28, fig. 6, mandibular tooth (erroneously listed as Platecarpus coryphaeus); pi. 29, fig. 2. posterior cervical vertebra; pi. 37. right scapula and coracoid; pi. 38, proximal half of left front paddle; pi. 39, right-hand figure, pelvic girdle (now missing distal end of pubis); pi. 54, fig. 4, pygal vertebra, fig. 5, anterior caudal vertebrae. Clidastes tortor Cope— Yakovlew 1901: text-fig. 1 (after pi. 23 in Williston 1898B). Clidastes tortor Cope— Williston 1903 A: text-fig. 4 (after pi. 23 in Williston 1898B). Clidastes tortor Cope— Yakovlew 1906: text-fig. 1 (after pi. 23 in Williston 1898B). Clidastes-^xWxsXon 1914: text-fig. 71 (after pi. 23 in Williston 1898B). Clidastes-V^msion 1925: text-fig. 58 (after pi. 23 in Williston 1898B); text-fig. 86A (redrawn from pi. 29, fig. 2 in Williston 1898B). Clidastes— KovaQv 1933: text-fig. 142D (after Williston 1914, text-fig. 71). Second edition 1945 text-fig. 169D; third edition 1966 text-fig. 204D. Clidastes— Komer 1956: text-fig. 108C (after pi. 23 in Williston 1898B). Skull, lower jaws, quadrate, vertebral column, left scapula and coracoid. pelvic girdle, parts of front paddle, and ribs. Upper Cretaceous. Senonian, Niobrara Chalk Formation, Smoky Hill Chalk Member; Smoky Hill River, Kansas; collected by E. P. West in 1889. KUVP 1108 Clidastes \Wav-Williston 1898B: pi. 28. figs. 1-4. Clidastes /error— Williston 1898B: pi. 61. fig. 4; right quadrate. KUVP 14278 Clidastes /orror— Williston 1898B: pi. 61. fig. 4; left quadrate. KUVP? Clidastes ror/or— Williston 1898B: pi. 42, figs. 5,6, posterior cervical vertebrae. Specimen not relocated. KUVP ? Clidastes ror/or— Williston 1898B: pi. 47, left-hand figure, frontal bone. Specimen not relocated. KUVP ? Clidastes rorror— Williston 1898B: pi. 64, fig. 1; left quadrate. Specimen not relocated. KUVP 1108, 14278, ? Upper Cretaceous, Senonian. Niobrara Formation. Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Russell (1967) considered Clidastes velox Marsh, C. westi Williston and Case, and C tortor Cope, to be conspecific with C. propython Cope. Clidastes sp. KUVP 66330 Clidastes sp. -Williston 1898B: pi. 57, fig. 3, tibia. KUVP 69552 Clidastes pumilus Marsh— Abel 1911: text-fig. 107, front paddle (drawing from a composite plaster cast, humerus reversed). Clidastes pumilus Marsh— Abel 1919: text-fig. 529 (from Abel 191 1. text- TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 39 fig. 107), front paddle. Clidastes pumilus Marsh— Abel 1927: text-fig. 292 (from Abel 1911, text- fig. 107), front paddle. Upper Cretaceous; western Kansas. KUVP 14348 CZ/V/a^r^'.v— Callison 1967: text-fig. 3. occipital fragment of skull; text-fig. 5, skull roof; text-fig. 8-3, 4b, right stapes. Specimens not relocated. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Mosasaurus Conybeare 1822 Mosasaurus ivoensis Persson 1963 KUVP 1024 Clidastes stenops Cope— WiWislon 1902C: pi. 12, lower figure, portion of front paddle, missing several phalangeal elements. Mosasaurus /vo^n^/^— Russell 1967: text-fig. 79, anterior portion of muzzle. Partial skull, ribs, pelvic girdle, front and hind paddle. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, western Kansas; collected by C. H. Sternberg. Russell (1967) referred this material, originally identified by Williston (1902C) as C. stenops Cope, to M. ivoensis Persson. Mosasaurus missouriensis (Harlan 1834) KUVP 1034 Mosasaurus horridus—'W\\\\?,iox\ 1895A: p. 166, holotype; "nearly the entire skull, the larger part of a front paddle, and about forty vertebrae"; pis. 14, 15, skull; pi. 16, fig. 1, left front paddle; fig. 2, right quadrate. Mosasaurus horridus Williston— Williston 1898B: pis. 19-21, skull; pi. 32 (after pi. 16, figs. 1,2 in Williston 1895A). Mosasaurus horridus Williston— Shor 1971: plate on p. 49 (after pi. 14 in Williston 1895A). Nearly complete skull, eight vertebrae, portions of the left front paddle. Upper Cretaceous, Campanian, Pierre Shale Formation; Cheyenne River, Custer County, South Dakota; collected by S. W. Williston and the KU Geological Expedition in the summer of 1894. Russell (1967) considered M. horridus Williston indistinguishable from, and a junior synonym of, M. missouriensis (Harlan). Subfamily Plioplatecarpinae (Dollo 1894) Platecarpus Cope 1869 Platecarpus coryphaeus (Cope 1872) KUVP ? Platecarpus caryphaeus (error) Cope— Williston 1897E: pi. 20, lower figure, restoration of skull. Specimen not relocated. Composite? Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP? Platecarpus con'/?/?a^w5— Williston 1898B: pi. 13 (after pi. 20 in Williston 1897E); pis. 14, 15, reconstruction of skull, probably a composite of several specimens; pi. 22, left mandible; pi. 24, fig. 4, postorbitofrontal; pi. 25, figs. 1,2, left maxilla; fig. 3, prefrontal; figs. 4,5, transverse bones; pi. 28, mandibular tooth; pi. 29, fig. 4, anterior end of vomer; fig. 5, indeterminate bones; pi. 52, fig. 3, right lateral piece of atlas; pi. 54, fig. 3, pygal vertebra; pi. 56, figs. 1,2,4, femora; pi. 57, fig. 5, fibula; fig. 6, metatarsal; fig. 8-10 in center, carpals; pi. 58, fig. 3, radius; fig. 6, ulna; fig. 7, radial carpal; fig. 8, 40 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 66337 KUVP 1001 KUVP? KUVP ? KUVP 66336 KUVP 1007 ulnar carpal; figs. 9,10, distal carpals; pi. 63, fig. 1, parietal; fig. 2, petrosals; fig. 5, odontoid; fig. 6, atlantar intercentrum; figs. 7,8, lateral pieces of atlas; fig. 11, articuloangular; pi. 72, middle figure, restoration of complete skeleton, a composite based upon several specimens (mostly KUVP 1007). None of these specimens have been relocated. Platecarpus—^illiston 1914: text-fig. 72, middle figure (redrawn from pi. 15 in Williston 1898B). Specimen not relocated. Mosasaur— Moodie 1923; pi. 40, magnified cross-section of diseased radius; pi. 48, figs. A, B, diseased radius; figs. C, D, diseased dorsal vertebrae. Specimens not relocated. Platecarpus—V^iWiston 1925: text-fig. 54, middle figure (after pi. 72, middle figure in Williston 1914). Platecarpus—Romer 1933: text-fig. 142C (after Williston 1914, text-fig. 72). Second edition 1945 text-fig. 169C; third edition 1966 text-fig. 204C. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Platecarpiis coryphaeus— "^\\\\?.ior\ 1898B: pi. 24, fig. 1, right pterygoid. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Platecarpus coryphaeus— WilUslon 1898B: pi. 24, fig. 5; pi. 63, fig. 3, left jugal. Platecarpus coryphaeus— WxWiston 1899A: pi. 12, cartilaginous tracheal rings erroneously described as a ''nuchal fringe," and cervical ribs. Nearly complete skeleton. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by S. W. Williston and party in the summer of 1898. Platecarpus coryphaeus— WiWislon 1898b: pi. 24, figs. 2,3, premaxilla. Specimen not relocated, if not part of KUVP 1007. Platecarpus coryphaeus— Williston 1898B: pi. 26, fig. 1, parietal; figs. 2,3, frontal. Specimens not relocated, if not part of KUVP 1007. Platecarpus cotyphaeus-Willision 1898B: pi. 26, fig. 4; pi. 63, fig. 4, left prosquamosal. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Platecarpus coryphaeus— 'Williston 1898B: pi. 27, top figure; rear view of complete skull; pi. 29, fig. 3, sclerotic plates; ? pi. 31, tigs. 4,5, pygal vertebrae; ? pi. 52, fig. 2, ulna; pi. 63, fig. 9, coronoid; pi. 72, middle figure, restoration of complete skeleton, a composite based upon several specimens. Platecarpus coryphaeus Cope— Zittel 1902: text-fig. 244 (after pi. 27, top figure in Williston 1898B). Second English edition 1932: text-fig. 435. Platecarpus coryphaeus Cope— Zittel 1902: text-fig. 250 (after pi. 72, middle figure in Williston 1898B). Second English edition 1932: text- fig. 442. Platecarpus— 'Williston 1914: Williston 1898B). Platecarpus coryphaeus Cope- pi. 27, top figure in Williston 1898B). Platecarpus— V^illiston 1925: text-fig. 57 (after pi. 27, top figure, in text-fig. 73 (after pi. 27, top figure in -Zittel 1918: text-fig. 360 (redrawn from TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 41 Williston 1898B). Platecarpus—W WMams 1975: Cover picture. Platecarpus— Bennett 1984: text-fig. 85C (redrawn from pi. 72, middle figure, Williston 1898B). Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, western Kansas; collected by E. P. West in 1890. KUVP 66335 Platecarpus coryphaeus— V/iWiston 1898B: pi. 28, fig. 7, mandibular tooth. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1131 Platecarpus coryphaeus— WiUiston 1898B: pi. 28, fig. 8,8a,8b, radius; pi. 41, fig. 2, pubis; fig. 2a, ischium; pi. 45, fig. l,la,b, radius. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by S. W. Williston on June 3, 1891. KUVP 66333 Platecarpus coryphaeus— WiUiston 1898B: pi. 31. fig. 7. Specimen not identified in plate explanation. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP ? Platecarpus coryphaeus— WiWiston 1898B: pi. 42, figs. 3,4, median dorsal vertebrae. Specimens not relocated. KUVP 1011 Platecarpus coryphaeus— WiUiston 1898B: pi. 46, figs. 1-4; pi. 52, fig. 1, left humerus; pi. 58, figs. 1, 4, radii, not relocated. Scapula, humerus, radius. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1042 Platecarpus coryphaeus— WiUiston 1898B: pi. 51, fig. 1, right humerus; fig. 2, left coracoid; fig. 3, right scapula; fig. 4, left humerus; fig. 5, carpal; fig. 6, radius; fig. 7, cervical hypapophysis (not numbered on photograph); figs. 8,9, cervical ribs; figs. 10-12, thoracic ribs; fig. 13, ossified cartilage. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; Smoky Hill River, either Logan or Gove Counties, Kansas; collected by E. P. West. KUVP 1041 Platecarpus coryphaeus— WiUiston 1898B: pi. 54, fig. 2, pygal vertebra. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, western Kansas; collected by E. P. West. KUVP 1171 Platecarpus coryphaeus— WiUiston 1898B: pi. 57, fig. 4, tibia. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown but probably near Elkader, Gove County, Kansas; collected by E. E. Slosson on June 2, 1891. KUVP 1172 Platecarpus coryphaeus-WiUiston 1898B: pi. 56, figs. 3,5. Mosasaur— Moodie 1917: fig. 18. Platecarpus coryphaeus — Moodie 1918: fig. 15. Mosasaur— Moodie 1923: pi. 34 (after fig. 18, in Moodie 1917). Exostosial radius, carpus, and part of the metacarpus. Specimen not relocated. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by S. W. Williston on June 2, 1891. KUVP 66332 Platecarpus coryphaeus-WiUiston 1898B: pi. 57, figs. 8-10 (lower left- 42 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY hand corner), tarsals (specimen in fig. 10 not relocated). Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1 167? Platecarpus coryphaeus— ^WWsion 1898B: pi. 57, fig. 7, fifth metatarsal. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by S. W. Williston on June 27, 1891. KUVP 1132 Platecarpus coryphaeus— V^WWs.ion 1898B: pi. 58, fig. 2. radius; fig. 5, ulna. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by E. C. Case on June 30, 1891. KUVP 14285 Platecarpus coryphaeus— "^WW^lon 1898B: pi. 60, fig. 3, left quadrate. Platecarpus cor\phaeus—Lane 1946: text-fig. 6 (redrawn from pi. 60, fig. 3 in Wiiliston 1898). Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1178 Platecarpus coryphaeus— WiWiston 1898B: pi. 61, fig. 3, right scapula. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1194 Platecarpus coryphaeus— WiWiston 1898B: pi. 63, fig. 10, surangular. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by S. W. Williston in June of 1891. KUVP 1004 Platecarpus coryphaeus Cope— Williston 1902C: pi. 12, upper figure, nearly complete hind paddle. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; South Fork of Solomon River, 3 miles northwest of Hill City, Sec. 17, T8S, R23W, Graham County, Kansas; collected by S. W. Williston in 1895. KUVP ? Platecarpus coryphaeus — Sternberg 1909: text-fig. 10, skull; no longer in collection. Platecarpus coryphaeus— Abel 1927: text-fig. 295 (after Sternberg 1909 text-fig. 10), skull. Upper Cretaceous, Niobrara Formation, Smoky Hill Chalk Member; western Kansas. KUVP 1081 Platecarpus coryphaeus Cope — Callison 1967: text-fig. 7, left quadrate, four vertebrae, rib fragments. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Wallace County, Kansas; collected by G. P. Cooper. Platecarpus ictericus (Cope 1871) KUVP 1142 Platecarpus /cr^r/cM5— Williston 1898B: pi. 43, scapula and coracoid (coracoid is now missing some pieces). KUVP ? Platecarpus ictericus— WiWhton 1898B: pi. 44, front paddle. KUVP 1142, ? Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, western Kansas; collected by S. W. Williston. Platecarpus sp. KUVP ? Platecarpus sp.— Williston 1895A: pi. 17, fig. 3, reconstruction of skull, probably a composite of several specimens. KUVP 66329 Platecarpus sp.— Williston 1898B: pi. 45, fig. 2,2A,B, tibia (?). TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 43 Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP ? Platecarpus sp.— Williston 1898B: pi. 57, figs. 1,2. radii. Specimens not relocated. KUVP 1051 Platecarpus— Mood'xc 1923: pi. 41a, lower figure, pi. 41c, right-hand specimen, diseased metatarsal; pi. 41a, top figure (specimen not relocated); pi. 41b, diseased terminal phalanges; pi. 41c, upper right- hand figure, diseased terminal phalanx, upper left-hand and center figures (specimens not relocated). Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, western Kansas; collected by E. P. West. Prognathodon Dollo 1889 Prognathodon overtoni (Williston 1897) KUVP 950 Undescribed genus Ov^rro/j/— Williston 1895A: p. 169, no figures; holotype. Brachysaunis overtoni Williston— Williston I897C: p. 96; pi. 8, fig. l,ia, mandibular tooth; fig. 2, fragmentary frontal; fig. 3, quadrate; fig. 4,4a, posterior cervical vertebra; fig. 5, medial caudal vertebra, not relocated; fig. 6, humerus; tig. 7, paddle bone. Brachysaurus overtonii (error) Williston— Williston 1898B: pi. 22, top figure, nearly complete left lower mandible and fragments of the anterior portion of right upper maxilla (tip of mandible is now missing); pi. 30 (after pi. 8, figs. 1-7 in Williston 1897C). Plate 40, figs. 10,11, indeterminate bones; pi. 62, fig. 1. humerus; fig. 2, coracoid. Fragmentary skull, upper and lower jaw, quadrate, ribs, 16 vertebrae, humeri, coracoid, and two paddle bones. Upper Cretaceous, Campanian, Pierre Shale Formation; Cheyenne River, Custer County, South Dakota; collected by T. R. Overton with the KU Geological Expedition in the summer of 1894. Williston (1895 A) recognized this material as probably generically dis- tinct from Mosasaurus, but declined to refer it to any known genus. He did, however, erect the name overtoni at that time. In 1897C he tentatively proposed the generic name, Brachysaurus. Schmidt (1927) stated that the name Brachysaurus is preoccupied by the iguanid genus Brachysaurus Hallowell and proposed a replacement name, Ancylocentrum. Russell (1967) comxdtxtd Ancylocentrum Schmidt ( = Brachysaurus Williston) to be conspecific with Prognathodon Dollo. Subfamily Tylosaurinae Williston 1897 Tylosaurus Marsh 1872 Tylosaurus proriger (Cope 1869) KUVP 1075 Liodon Owen (Tylosaurus Marsh)— Snow 1877: plate on p. 55, impres- sion of dermal scales; text-fig. on p. 57, oudine drawing of right front paddle. Tylosaurus— WiWiston 1897D: text-fig. 1, outline drawing of front paddle; pi. 9, front paddle and ribs, in matrix; pi. 10, hind paddle bones, in matrix; pi. 1 1 , left pelvis, in matrix; pi. 12, dermal scale impressions. Tylosaurus proriger— ^iWiston 1898B: pi. 48, left-hand figure (after fig. 1 in Williston 1897D); right-hand figure (after pi. 9 in Williston 1897D); pi. 49 (after pi. 1 1 in Williston 1897D); pi. 50 (after pi. 10 in Williston 1897D); pi. 68, entire specimen, in matrix; pi. 69 (after pi. 44 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY on p. 55 in Snow 1877); pi. 70 (after pi. 12 in Williston 1897D), image is reversed. T\losaurus—W iWiston 1914: text-fig. 75 (after pi. 9 in Williston 1897D); fig. 78 (after pi. 12 in Williston 1897D). rv/o^flwn/^— Williston 1925: text-fig. 148 (after text-fig. 75 in Williston 1914). Txlosaurus proriger—Shor 1971: plate on p. 48 (after pi. 69 in Williston 1898B). Two front paddles, ribs, pelvic girdle. 20 vertebrae, impression of dermal scales. Upper Cretaceous. Senonian. Niobrara Formation. Smoky Hill Chalk Member; Hackberry Creek. Gove County. Kansas; collected by F. H. Snow and L. L. Dyche in 1877. KUVP ? Txlosaurus proriger Cope— Williston 1898B: pis. 16-18, reconstruction of skull (pi. 16 after pi. 20. top figure in Williston 1897E); pi. 72 (after pi. 13. fig. 3, in Williston 1897G) reconstruction of skeleton. Reconstruction of skull and skeleton from several specimens. Txlosaurus proriger Cope— Zittel 1918: text-fig. 359C (redrawn from pi. 18 in Williston 1898B). Txlosaurus— Williston 1914: pi. 72. lower figure (redrawn from pi. 17 in Williston 1898B). Tylosaurus proriger— Abel 1919: text-fig. 531 (redrawn from pi. 18 in Williston 1898B). Txlosaurus— Williston 1925: text-fig. 54. lower figure (after text-fig. 72, lower figure in Williston 1914). Upper Cretaceous, Senonian, Niobrara Formation. Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 66334 Txlosaurus proriger— Williston 1898B: pi. 29, fig. 1. mandibular tooth. rv/o5a//n/5— Williston 1914: text-fig. 70, image reversed (after pi. 29, fig. 1 in Williston 1898B). Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1195 Txlosaurus proriger-W illiston 1898B: pi. 40, fig. 1, femur; fig. 3, right ischium; fig. 4, left ischium; also includes thirty-one vertebrae and a skull fragment. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by E. C. Case on June 30, 1891. KUVP ? Txlosaurus proriger— Williston 1898B: pi. 40, fig. 2, ilium; fig. 6, radius; fig. 7, metacarpal. Specimens not relocated. KUVP 1017 Tylosaurus proriger-Williston 1898B: pi. 41, fig. la, left ilium; fig. lb, pubis; fig. Ic, ischium. Pelvic girdles. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP 1029 Txlosaurus proriger-Williston 1898B: pi. 40, fig. 5, femur; fig. 8, ulna; fig. 9 radius. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; Smoky Hill River, either Logan or Gove Counties, Kansas; collected by E. P. West in 1890. KUVP 1016 Txlosaurus proriger-Williston 1898B: pi. 55, fig. 1, radius; fig. 2, humerus; fig. 3, coracoid; fig. 4, cartilage. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 45 Member; exact locality and collector unknown, western Kansas. KUVP 1020 Tylosaurus proriger—WiWision 1898B: pi. 60, fig. 1, right quadrate; fig. 2, left quadrate. Txlosaurus proriger— Lane 1946: text-fig. 5 (redrawn from pi. 60, fig. 2 in Williston 1898B). Cranial fragments, quadrates, paddle elements. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; Smoky Hill River, either Logan or Gove Counties, Kansas; collected by E. P. West in 1890. KUVP 1 189 Txlosaurus proriger— WiWiston 1898B: pi. 64, t~ig. 2, left quadrate; fig. 3, anterior caudal vertebrae in matrix (fig. reversed); material in pi. 64 erroneously listed as Platecarpus coryphaeus; pi. 65, posterior tho- racic and anterior dorsolumbar vertebrae and ribs, in matrix. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown. Gove County, Kan.sas; collected by S. W. Williston on June 1, 1891. KUVP 1013 Tylosaurus proriger—'^WWsion 1898B: pi. 66, posterior caudal vertebrae; pi. 67, terminal caudal vertebrae. Twenty-eight vertebrae. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. Tylosaurus indet. fylosaurus dyspelor (Cope 1871) nomen dubium KUVP 1014 Tylosaurus dyspelor— Wilhston 1898B: pi. 41, figs. 4,5, posterior cer- vical vertebra. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. KUVP ? Tylosaurus dyspelor—V^ iWiston 1898B: pi. 42, figs. 1,2, pygal vertebra; pi. 59, fig. 1, humerus; fig. 2, radius; fig. 6, phalangeal element. Specimens not relocated. KUVP 1117 Tylosaurus dyspelor— WiWiston 1898B: pi. 59, fig. 3, ulna; figs. 4-5, 7-8, metacarpals and phalangeal elements. Pieces of skull, 44 vertebrae, bones of front and hind paddles. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by E. P. West. KUVP 1032 Tylosaurus dyspelor— WWhston 1898B: pi. 61, figs. 1 (image reversed), 2, left quadrate. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; mouth of 12 Mile Creek, on the Smoky Hill River, Logan County, Kansas; collected by E. P. West. Russell (1967) declined to refer Tylosaurus dyspelor to either of the two species of Niobrara Tylosaurus, nepaeolicus or proriger. Instead, he regarded the name dyspelor as a nomen vanum, and also stated that the holotype is probably a junior synonym of T proriger (Cope). Tylosaurus sp. KUVP ? Tylosaurus sp.— Williston I895A: pi. 17, fig. 2, reconstruction of skull, probably a composite of several specimens. KUVP 1015 Tylosaurus sp.— Williston 1898B: pi. 54, fig. 1, pygal vertebrae. Upper Cretaceous, Campanian, Pierre Shale Formation, Sharon Springs Member?; exact locality unknown, Wallace County, Kansas; collected by S. W. Williston in 1891. 46 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Mosasauridae indet. KUVP 6176 Mosasaur— Martin and Stewart 1977: text-fig. lA, four dorsal vertebrae and some unidentified fragments. Upper Cretaceous, Turonian, Carlile Shale Formation, Blue Hill Shale Member; near Medicine Peak, Osborn County, Kansas; collected by A. Graffham. KUVP 27032 Mosasaur— Martin and Stewart 1977: text-fig. IB, two daudal vertebrae. Upper Cretaceous, Turonian, Carlile Shale Formation, Blue Hill Shale Member; NW'A, Sec. 19, TllS, R17W, Ellis County, Kansas; collected by J. D. Stewart in 1973. KUVP 25869 Mosasaur— Martin and Stewart 1977: text-fig. IC, posterior tip of right maxilla. Upper Cretaceous, Turonian, Carlile Shale Formation, Fairport Chalk Member; EV:, Sec. 19, TllS. R17W, Ellis County, Kansas; collected by J. D. Stewart in 1975. p. 417; text-fig. 1; holotype; p. 417; text-fig. 2; paratype; Suborder Serpentes Linnaeus 1758 Family Boidae Bonaparte 1831 Geringophis Holman 1976 Gehngophis vetus Holman 1982 KUVP 49126 Geringophis vetus sp. nov.— Holman 1982: p. 490, text-fig. 1; holotype; trunk vertebra. Tertiary, Oligocene, Brule Formation (Late Orellan land mammal age); NE'A, Sec. 8, T33N, R53W, Toadstool Park, Sioux County, Nebraska; collected by L. Martin and O. Bonner field party in 1970's. Infraorder Henophidia incertae sedis Helagras Cope 1883 Helagras orellanensis Holman 1983 KUVP 49127 Helagras orellanensis n. sp.— Holman 1983: trunk vertebra. KUVP 49128 Helagras orellanensis n. sp.— Holman 1983: two fused vertebrae of juvenile. Tertiary, Middle Oligocene, Brule Formation (Orellan land mammal age); Toadstool Park area near Orella, Sioux County, Nebraska, anthill collec- tions; collected by L. Martin, O. Bonner field party in 1970's. Suborder Amphisbaenia Cope 1864 Family Amphisbaenidae Gray 1825 Hyporhina Baur 1893 Hyporhina galbreathi Taylor 1951 KUVP 8221 Hyporhina galbreathi sp. nov.— Taylor 1951: p. 532, text-fig. 2A-C; pi. 58, figs. 6-8; holotype; a nearly complete skull, missing quadrate, part of squamosal and lower jaws. Hyporhina galbreathi— Esles 1983: text-fig. 49A (after text-fig. 2A-C in Taylor 1951). KUVP 8219 Hyporhina galbreathi sp. nov.— Taylor 1951: text-fig. 3A,B; pi. 59, fig. 5; referred material, fragmentary skull with cast of braincase, lacking jaws. KUVP 8222 Hyporhina galbreathi sp. nov.— Taylor 1951: pi. 59. fig. 4; referred material, fragmentary skull, missing lower jaws. KUVP 8219, 8221, 8222 Tertiary, Middle Oligocene, White River Formation (Orellan land mammal age); Cedar Creek Member; Clyde Ward Ranch, SE'A, TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 47 SWV4, Sec. 12, TUN, R54W, 35 miles northwest of Sterling, Logan County, Colorado; collected by E. C. Galbreath in 1948. Rhineura Cope 1861 Rhineura amblyceps Taylor 1951 KUVP 7649 Rhineura amblyceps sp. nov.— Taylor 1951: p. 543; text-fig. 5A-C; pi. 61, figs. 1-3; holotype; basal portion of skull, missing jaws. Er- roneously listed by Taylor in fig. 5 as KUVP 7469. KUVP 7650 Rhineura amblyceps sp. nov.— Taylor 1951: text-fig. 6A-C; pi. 62, figs. 1-3; referred material, fragmentary braincase, missing jaws. KUVP 7649-50 Tertiary, Middle Oligocene, White River Formation (Orellan land mam- mal age). Cedar Creek Member, Oreodon beds; Clyde Ward Ranch, W'/2, Sec. 7, TUN, R53W, about 35 miles northwest of Sterling, Logan County, Colorado; collected by R. R. Camp (C. W. Hibbard field party) on July 29 and July 31, 1946, respectively. KUVP 9041 Rhineura amblyceps sp. nov.— Taylor 1951: pi. 59, fig. 1; pi. 61, figs. 4,5; fragmentary skull, lacking jaws. Tertiary, Middle Oligocene, White River Formation (Orellan land mam- mal age). Cedar Creek Member; Clyde Ward Ranch, WV2, Sec. 7, Tl IN, R53W; about 35 miles northwest of Sterling, Logan County, Colorado; collected by E. C. Galbreath in 1948. Rhineura hatcheri Baur 1893 KUVP 8220 Rhineura hatcheri Baur— Taylor 1951: pi. 65, figs. 1-3; nearly complete skull, lacking jaws. Tertiary, Middle Oligocene (Orellan land mammal age); SE'A, Sec. 21, TUN, R53W, Logan County, Colorado; collected by R. W. Wilson (E. C. Galbreath Field Party) in 1948. KUVP 8960 Rhineura hatcheri Baur— Taylor 1951: pi. 66, figs. 1-3; nearly complete skull with jaws. Tertiary, Middle Oligocene, White River Formation (Orellan land mam- mal age). Cedar Creek Member; Clyde Ward Ranch, NE'A, SE'A, Sec. 3, TUN, R54W, Logan County, Colorado; collected by E. C. Galbreath in 1949. Rhineura wilsoni Taylor 1951 KUVP 7651 Rhineura wilsoni sp. nov.— Taylor 1951: p. 548, text-fig. 7A-C; pi. 63, figs. 1-3; holotype; nearly complete skull, lacking squamosal, quad- rate, columella, extra-columella, and lower jaws. Erroneously listed in text-fig. 7 and pi. 63 as KUVP 7649. Rhineura wilsoni— Esies 1983: text-fig. 50C (after text-fig. 7A-C in Taylor 1951). Tertiary, Middle Oligocene, White River Formation (Orellan land mam- mal age). Cedar Creek Member;" Clyde Ward Ranch, NW'A, Sec. 7, TUN, R53W, about 35 miles northwest of Sterling, Logan County, Colorado; collected by R. R. Camp (C. W. Hibbard field party) on July 31. 1946. Superorder Archosauria Cope 1869 Order Thecodontia Owen 1859 Suborder Pseudosuchia Zittel 1887-1 Infraorder Parasuchia Huxley 1875 Family Parasuchidae Lydekker 1885 Angistorhinus Mehl 1913 Angistorhinus aeolamnis Eaton 1965 890 48 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY KUVP 1 1659 Angistorhinus aeolamnis Eaton n. sp. — Eaton 1965: p. 4, text-figs. 1 A-C, 2A,B; holotype; a nearly complete skull lacking lower jaws. Angistorhinus aeolamnis— Charig et al. 1976: text-fig. 8C. palatal view of skull (redrawn from Eaton 1965). Upper Triassic, Chugwater Formation, Popo Agie Member; 23 miles northwest of Lander via U.S. Highway 287 and 1 1 miles west of Highway 287 on Sage Creek Road, SE'A, Sec. 14, TIN, R3W, Fremont County, Wyoming; collected by University of Kansas tleld party during the summer of 1960. KUVP 11660 Angistorhinus sp.— Eaton 1965: text-fig. 2C; fragment of the interorbital skull roof. Collected at the type locality of A. aeolamnis Eaton by R. D. Platz during the summer of 1960. Order Crocodylia Gmelin 1788 Suborder Mesosuchia Huxley 1875 Family Goniopholididae Cope 1875 Goniopholis Owen 1841 {= Hyposaurus Owen 1849) Goniopholis sp. KUVP 66331 Crocodile-Williston 1894: pi. 1, fig. 5. single vertebra. Crocodile— Williston 1898A: text-fig. 4 (after pi. 1, fig. 5 in Williston 1894). Williston tentatively placed this vertebra in the crocodilian genus Hypo- saurus Cope. This specimen was originally cataloged as KUVP 829. Lower Cretaceous, Albian, Kiowa Formation; Clark County, Kansas. Order Pterosauria Kaup 1834 Suborder Pterodactyloidea Plieninger 1901 Family Pteranodontidae Marsh 1876 Pteranodon Marsh 1876 Pteranodon ingens (Marsh 1872) KUVP 929 Pteranodon sp. — Williston 1892: text-fig. 1, prepubic bones. Upper Cretaceous. Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by E. P. West. KUVP 2212 Pteranodon sp. — Williston 1892: pi. 1. nearly complete skull, lacking medial portion of lower jaw. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, but probably on Smoky Hill River, near Elkader, Logan County, Kansas; collected by E. C. Case and S. W. Williston with the Kansas University Geological Expedition during the summer of 1891. KUVP ? Ornithostoma ingens— 'WWW^Xon 1893A: text-fig. on p. 80, composite reconstruction of pelvic girdle and left hind limb, too generalized to identify specimens. Ornithostoma ingens M.— Williston 1897A: text-fig. on p. 49 (from fig. on p. 80 in Williston 1893A). No locality data. KUVP 974 Ornithostoma — Williston 1895B: pi. 1. fig. 1, reconstruction of posterior half of mandible. Upper Cretaceous, Santonian or Campanian. Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by H. T. Martin. TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 49 KUVP 976 Ornithostoma—V^iWiston 1896: pi. 1, figs. 1.2, posterior half of skull. Pteranodon sp.— Huene 1914: text-fig. 8, posterior half of skull. Upper Cretaceous, Santonian or Campanian. Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by C. H. Sternberg. KUVP 966 Ornirhostoma species— WiWiston 1897A: text-fig. on p. 42, sternum. KUVP 2212 Ornirhostoma ingens M.— Williston 1897 A: pi. 2, restoration of complete skeleton based on several specimens (skull is based on KUVP 2212). KUVP 966, 2212 Cretaceous, Cenomanian or Santonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Trego County, Kansas; collected by H. T. Martin. KUVP 1354 Order Saurischia Seeley 1888 Suborder Sauropodomorpha Huene 1932 Infraorder Sauropoda Marsh 1878 Family Camarasauridae Cope 1877 Subfamily Camarasaurinae Nopcsa 1928 Camarasaurus Cope 1877 ( = Morosaurus Marsh 1878) Camarasaurus sp.? Morosaurus grandis—V\.QrnbQvg 1909 and 1931: text-fig. 20 (after pi. ll'in Williston 1902B). Trinacromerum osborni— 'WiWi^ion 1914: text-fig. 32 (after pi. 11 in Williston 1902B); text-fig. 35, restoration of pectoral girdle; text-fig. 36, restoration of pelvic girdle; text-fig. 38B,C (redrawn from pi. 20 in Williston 1903A); text-fig. 41 (redrawn from pi. 3, fig. 2 in Williston 1903A). Trinacromerum Osborni Williston (error)— Zittel 1918: text-fig. 400 (redrawn from text-fig. 36 in Williston 1914). Trinacromerum osborni— ^\\\'\?Xon 1925: text-fig. 48C (after text-fig. 41 in Williston 1914); text-tig. 80A (redrawn from pi. 22, fig. 5 in Williston 1903 A, image reversed): text-fig. 80O (redrawn from pi. 1 1 in Williston 1903A); text-fig. 102 (redrawn from fig. 35 in Williston 1914); text-fig 125 (redrawn from text-tig. 36 in Williston 1914); text- fig. 159B,C (after text-fig. 38B.C in Williston 1914); text-fig. 174 (after pi. 11 in Williston 1902B). Trinacromerum osborni Williston— Zittel 1932: text-tig. 404 (redrawn from text-fig. 36 in Williston 1914). Trinacromerum osborni— Lane 1946: text-fig. 3; complete skeleton. Trinacromerum Cragin— Huene 1956: text-fig. 459E (redrawn from text- fig. 35 in Williston 1914)— also figured are a skull and front paddle; these are probably based on University of Kansas material but 54 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY comparison is difficult. Trinacromerum—Romer 1956: text-fig. 120O (redrawn from text-fig. 80 in Williston 1925); text-fig. 148C (redrawn from text-fig. 102 in Williston 1925); text-fig. 159B (redrawn from text-fig. 125 in Williston 1925); text-fig. 186C (redrawn from text-fig. 159C in Williston 1925, image reversed); text-fig. 194B (redrawn from text- fig. 159B in Williston 1925, image reversed). Dolichorhynchops osborni Williston— Welles 1962: text-fig. 13A. left humerus; text-fig. 13B, pectoral girdle; text-fig. 13C, left femur; text- fig. 13D, pelvic girdle (all figures redrawn from Williston 1903A); text-fig. 23 (a reduction of text-fig. 13 in Welles 1962). Dolichorhynochops (error)— Bennett 1984: text-fig. 87 A, drawing of complete skeleton. Upper Cretaceous, Santonian or Campanian. Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Logan County, Kansas; collected by C. H. Sternberg in 1901. Dolichorhynchops willistoni (Riggs 1944) KUVP 5070 Thnacromeritm wilUsioni sp. nov.— Riggs 1944: p. 77, holotype; text-fig. 1, skull; text-fig. 2, atlanto-axial complex; text-fig. 3, scapula; text- figs. 4,5, fragmentary clavicles and interclavicle; text-fig. 6, right coracoid; text-fig. 7, fragmentary pelvic girdle; text-fig. 8, fragmen- tary pelvic bones (pubis and ischium); other type material includes approximately fifty vertebrae. Upper Cretaceous, Cenomanian or Turonian, Greenhorn Formation, Hartland Shale Member; road cut on U.S. Hwy. 81, south of Concordia, Sec. 16, T6S, R3W, Cloud County, Kansas; collected by road crew in December of 1936. Welles (1962) considered the genus Trinacromemm Cragin to be based upon non-diagnostic material and consequently transferred T. willistoni Riggs to the genus Dolichorhynchops. Dolichorhynchops indet. KUVP 1325 Trinacromemm anonymum n. sp. — Williston 1903A: p. 66. holotype; text-fig. 9, episternum; pi. 28, ten associated cervical vertebrae, propodial, parts of illim and coracoid. Unfigured portion of holotype includes 16 vertebrae, 2 phalanges and fragments. Upper Cretaceous, Turonian, Benton Limestone Formation; near Paradise Creek, NW'A, Sec. 20, TllS, R14W. Russell County, Kansas; collected by B. F. Mudge in 1873. Welles (1962) considered this material non-diagnostic and the name a nomen "vanum," thus representing an indeterminate form within the genus Dolichorhynchops. Polycotylidae indet. Elasmosaiirus sternbergi Williston 1906, nomen dubium KUVP 1312 Elasmosaurus sternbergi n. sp.— Williston 1906: p. 232, no figures; holotype; two complete dorsal vertebrae and additional fragments. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Gove County, Kansas; collected by C. H. Sternberg in 1895. The KUVP catalogue lists this holotype as KUVP 1302. however. Welles (1952) found that material numbered as KUVP 1302 consisted of seven vertebrae and, proportionally, were very different from the measurements TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 55 provided by Williston (1906). See KUVP 1302 account for more informa- tion. The correct number for this material is KUVP 1312. Based on the measurements in Williston (1906), Welles concluded that this material is pliosaurian not elasmosaurian and of indeterminate affinity. Plesiosaurus gulo Cope 1872, nomen dubium KUVP 1329 Plesiosaurus gulo— Cope 1872: p. 128, no figures; holotype; "eleven cervical, thirteen dorsal, and seven or eight other vertebrae, with portions of the scapular and pelvic arch, and ribs." Plesiosaurus gulo Cope — Williston 1903A: text-fig. 7, ventral extremity of scapula. Lower Cretaceous, Kiowa Formation?; collected by J. Savage. Cope reported that this material is from the Niobrara or Pierre Formation, near Sheridan (now McAllister), Logan County, Kansas. The type of preservation strongly indicates that this is incorrect. P. gulo is not from the Niobrara and probably not from the Pierre Shale. Kiowa Formation is likely but this formation does not occur in Logan County. Welles (1952) stated that the figure in Williston (1903A), if it is a scapular fragment, may be pliosaurian, whereas if it is the distal border of the coracoid, it is probably elasmosaurian. In any case, the material is not diagnostic and P. gulo Cope is a nomen "vanum. " Specimens catalogued as KUVP 1329 are mostly poorly preserved and probably include several taxa. This material consists of more than 40 vertebral centra and other fragments, including the piece of "scapula" figured by Williston (1903A). It is not possible to separate the type from other material with certainty. Plesiosaurus gouldi Williston 1897, nomen dubium KUVP ? Plesiosaurus Gouldii n. sp. (error)— Williston 1897B: p. 57, no figures; holotype; several nearly complete dorsal vertebrae. Specimens not relocated. Plesiosaurus gouldii (error)— Williston 1903A: pi. 27, figs. 2,3, dorsal vertebra. Specimen not relocated. Confusion exists as to what constitutes the holotype of P. gouldi and consequently where this species should be placed within the Plesiosauria. The type consists of several nearly complete dorsal vertebrae. According to Welles (1952, p. 1-6) measurements of one of these vertebrae (Williston 1897B, p. 57; 1903A, p. 13) indicate it is an elasmosaur. However, the description published by Williston in these same papers appears to match the specimen figured and labeled by him as the holotype (pi. 27, figs. 2,3). Measurements taken from this figure indicate this material is from a pliosaur (Welles 1952, p. 1 16). Because Welles (1952) believed (based on Williston's published measurements) that the holotype of P. gouldi is from an elasmosaur, he did not consider the vertebra figured by Williston (1903A) as part of the holotype; P. gouldi is therefore Dolicho- rhynchopidae indet. Lower Cretaceous, Albian. Kiowa Formation; exact locality unknown, Clark County, Kansas; collected by C. N. Gould. Polycotylus latipinnis Cope 1869, nomen dubium KUVP 5916 Polycotylus latipinnis Cope— Williston 1903 A: pi. 21 , right pelvic paddle (published as pectoral paddle). Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown. Smoky Hill River, east of 56 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Fort Wallace, Wallace County. Kansas; collected by G. R. AUman. Pliosauroidea indet. KUVP ? Cimoliosaums sp. (error)— Williston 1894: pi. 1, fig. 3, propodial. Specimen not relocated. Probably pliosaurian (Welles 1952, p. 116). Lower Cretaceous, Albian, Kiowa Formation; vicinity of Upper Bluff and Sand Creeks, near Ashland, Clark County, Kansas; collected by C. N. Gould and S. W. Williston in 1894. KUVP ? Plesiosaurus mudgei—Cvagin 1894: pi. 1, fig. 4, fragmentary propodial (femur?), tentatively referred to P. mudgei by Cragin; propodial element is probably pliosaurian. Lower Cretaceous, Albian, Kiowa Formation (cited as Fredericksburg); Blue Cut Hill, southeastern Kiowa County, Kansas; collected by A. L. Diamond. KUVP 7545 Plesiosaurus mudgei Cragin— Williston 1903A: text-fig. 6, portion of the right scapula. Lower Cretaceous, Albian, Kiowa Formation; exact locality unknown, Clark County, Kansas; collected by C. N. Gould. Welles (1952) stated that this scapula is probably pliosaurian. Plesiosauria indet. "young plesiosaurs"— Williston 1903A: pi. 23, fig. 4, juvenile propodial. ""Polycotylus and Elasmosaurus"'—Moodie 1911: pi. 1, fourth specimen from right-hand side. "young plesiosaurs"— Williston 1903A: pi. 23, fig. 3, juvenile propodial. Upper Cretaceous, Senonian, Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, western Kansas. "Polycotylus and Elasmosaurus^^—y[oo6\Q 191 1: pi. 1, first specimen on left-hand side; juvenile propodial (humerus); material also includes five carpal elements. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; exact locality unknown, Wallace County, Kansas; collected by Kansas University Geological Expedition during the summer of 1895. KUVP? "plesiosaurian propodial"— Moodie 1911: text-figs. 1,2,3; pi. 1 right- hand specimen, embryonic propodial. Specimen not relocated. plesiosaurian— Moodie 1916: text-fig. 5, embryonic plesiosaurian pro- podial (from text-fig. 1 in Moodie 1911). Cretaceous. Niobrara Formation, Smoky Hill Chalk Member; exact locality and collector unknown, probably western Kansas. KUVP? plesiosaur— Moodie 1911: text-figs. 4,5, juvenile propodial; figs. 8,9, mature propodial. Specimens not relocated. plesiosaurian {Polycotylus)—'M.ood\e 1916: text-fig. 3, propodial (re- drawn from text-fig. 4 in Moodie 191 1). Probably Cretaceous of western Kansas; no other data. KUVP ? Polycotylus—Moodxe 1911: text-fig. 6, nearly mature propodial; fig. 7, mature propodial. Specimens not relocated. Probably Cretaceous of western Kansas; no other data. KUVP? ^'Polycotylus and Elasmosaurus" —Moo6\e 1911: pi. 1, second, third, fifth and sixth specimens from the left, propodials. Specimens not relocated. Probably Cretaceous of western Kansas; no other data. KUVP ? plesiosaurian— Moodie 1916: text-fig. 2, distal end of embryonic pro- KUVP 1314 KUVP 1315 KUVP 1314-5 KUVP 1323 TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES podial. Specimen not relocated. Probably Cretaceous of western Kansas; no other data. 57 Subclass Theropsida Seeley 1896 Order Pelycosauria Cope 1878 Suborder Sphenacodontia Marsh 1878 Family Sphenacodontidae Marsh 1878 Virausmosaurus Fox 1962 Tfirausmosaurus serratidens Fox 1962 (Type-species) KUVP 11120 Tlirausmosaurus serratidens new genus and new species— Fox 1962: p. 302; holotype; text-fig. 4, fragmentary left dentary. KUVP 11121 TJnaiismosaiirus serratidens new genus and new species— Fox 1962: text- fig. 5, fragmentary left (?) maxilla (referred specimen). Specimen not relocated. KUVP 1 1 122 Tlirausmosaurus serratidens new genus and new species— Fox 1962: text- fig. 6, fragmentary left dentary (referred specimen). Specimen not relocated. KUVP 11120-22 Lower Permian, Wichita Group, Wellington Formation, fissure fills within the Ordovician Arbuckle Limestone; Dolese Brothers limestone quarry at Richards Spur, just west of U.S. Hwys. 62 and 281, 10.5 miles south of Apache, 6 miles north of Fort Sill, SWV2, Sec. 31, T4N, RllW, Comanche County, Oklahoma. Suborder Edaphosauria Cope 1882 Family Nitosauridae Romer and Price 1940 Delorhynchus Fox 1962 Delorhynchus priscus Fox 1962 (Type-species) KUVP 11117 Delorhynchus priscus new genus and new species— Fox 1962: p. 299; holotype; text-fig. 1, fragmentary left maxilla. KUVP 11118 Delorhynchus priscus new genus and new species— Fox 1962: text-fig. 2, fragmentary right maxilla (referred specimen). KUVP 11 1 19 Delorhynchus priscus new genus and new species— Fox 1962: text-fig. 3, fragmentary left maxilla (referred specimen). KUVP 11117-9 Lower Permian, Wichita Group, Wellington Formation, fissure fills within the Ordovician Arbuckle Limestone; Dolese Brothers limestone quarry at Richards Spur, just west of U.S. Hwys. 62 and 281, 10.5 miles south of Apache, 6 miles north of Fort Sill, SW'/:, Sec. 31, T4N, RllW, Comanche County, Oklahoma. Family Edaphosauridae Cope 1882 Edaphosaurus Cope 1882 Edaphosaurus sp. KUVP 1425 Petrolacosaurus— Lane 1945: text-fig. 2, a nearly complete pelvis. Petrolacosaurus kansensis Lane— Peabody 1952: text-fig. 5, left half of pelvis. Listed by Peabody as the topotype of P. kansensis Lane. Reisz et al. (1982), stated that KUVP 1425 was mistakenly assigned to Petrolacosaurus by Peabody in his 1952 paper, but subsequently referred to Edaphosaurus ecordi in Peabody 's 1957 publication. Collected by H. H. Lane in 1931 or 1932. KUVP 59037 Undescribed edaphosaurian pelycosaur— Reisz et al. 1982: text-fig. 4, articulated presacral vertebral column, partial, skull and humerus. 58 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Pelycosaur— Bennett 1984: text-fig. 62. Reisz et al. stated that this material probably belongs to the genus Edaphosaurus. KUVP 1425, 59037 Upper Pennsylvanian. Missourian, Stanton Formation, Rock Lake Member; six miles northwest of Garnett, Sec. 32, T19S, R19E, Anderson County, Kansas; collected by F. E. Peabody in 1954. Pelycosauria incertae sedis Xyrospondylus Reisz et al. 1982 Xyrospondylus ecordi (Peabody 1957) (Type-species) KUVP 9963 Edaphosaurus ecordi, n. sp.— Peabody 1957: p. 947; holotype; text-fig. 1, a complete posterior cervical vertebra. Xyrospondylus ecordi n. gen. — Reisz et al. 1982; text-fig. 3, holotype. Upper Pennsylvanian, Missourian, Stanton Formation, Rock Lake Mem- ber; six miles northwest of Garnett, Sec. 32, T19S, R19E, Anderson County, Kansas. Reptilia incertae sedis Apatomerus Williston 1903 Apatomerus mirus Williston 1903 (Type-species) KUVP 1198 Crocodile-Williston 1894A: pi. 1, fig. 4, femur. Crocodile— Williston 1898A: text-fig. 3 (after pi. 1, fig. 4 in Williston 1894). Apatomerus mirus, gen. et sp. nov.— Williston 1903B: p. 160; holotype; femur. Lower Cretaceous, Albian, Kiowa Formation; vicinity of Bluff and Sand Creeks, near Ashland, Clark County, Kansas; collected by C. N. Gould in 1893. Williston questionably identified this femur as belonging to a crocodile, however, in a later paper Williston (1903B) identified this femur as representing a new type of pterosaur. He erected the genus and species Apatomerus mirus based on this specimen. Wellnhofer (1978) stated that this material resembles Pteranodon in overall shape, but Apatomerus is distinctly different in being larger in size and thicker boned than this genus. He was unable to determine its affinities. We doubt the assignment to pterosaurs as a whole. 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Sci. 17:195-199. WILLISTON, S. W. 1902A. On the hind limb of Pro- tostega. Amer. J. Sci. 13:276-278. WiLLiSTON. S. W. 1902B. Restoration of Doliclw- rhynchops osborni, a new Cretaceous plesiosaur. Kanas Univ. Sci. Bull. 1:241-244. WiLLiSTON, S. W. 1902C. Notes on some new or little known extinct reptiles. Kansas Univ. Sci. Bull. 247-254. WiLLiSTON, S. W. 1903A. North American plesio- saurs. Pt. 1. Field Columbia. Mus. Publ. No. 73, Geol. Ser. 2:1-77. WiLLiSTON, S. W. 1903B. On the osteology of A'vc- tosaurus (Nyctodactylus) with notes on Ameri- can Pterosaurs. Field Columbia Mus. Publ. 78, Geol. Ser. 2:125-163. WiLLiSTON, S. W. 1904. The temporal arches of the Reptilia. Biol. Bull. 7:175-192. WiLLiSTON, S. W. 1906. North American plesiosaurs: Elasmosaurus, Cimoliasaunis, and Polycotylus. Amer. J. Sci. 21:221-236. WiLLiSTON, S. W. 1908. North American Plesiosaur. Trinacromerum. J. Geol. 16:715-736. WiLLiSTON, S. W. 1910A. Cacops. Desmospondyius: New genera of Permian vertebrates. Bull. Geol. Soc. Amer. 21:249-284. WiLLiSTON, S. W. 1910B. New Permian reptiles; rhachitomous vertebrae. J. Geol. 18:590-594. WiLLiSTON, S. W. 1914. Water Reptiles of the past and present, i-vii + 251 pp. Univ. Chicago Press, Chicago. WiLLiSTON. S. W. 1916. Synopsis of some American Permo-carboniferous tetrapoda. Contrib. Walker Mus. 1:225-226. WiLLiSTON. S. W. 1925. The osteology of the reptiles. 1st ed. Harvard Univ. Press. Cambridge. 300 pp. WiLLiSTON. S. W.. andE. C. Case. 1892. Kansas Mo- sasaurs, Part I. Kansas Univ. Quart. 1:15-32. WiLLiSTON. S. W.. and R. L. Moodie. 1913. New plesiosaurian genus from the Niobrara Cre- taceous of Nebraska (Abstract). Bull. Geol. Soc. America 24:120-121. WiLLiSTON, S. W., AND R. L. MOODIE. 1917. Og- modius martinii, a new plesiosaur from the Cretaceous of Kansas. Kansas Univ. Sci. Bull. 10:61-73. Yakovlew, N. 1901. Restes d'un mosasaurien trouve dans le Cretace superieur du Sud de la Russie. Bull. Com. Geol. (U.S.S.R.). St. Petersbourg 20:507-520. (Russ. with French summary). Yakovlew, N. 1906. Notes sur les mosasauriens. Bull. Com. Geol. (U.S.S.R.), St. Petersbourg 24:135-152. (Russ. with French summary). Yatkola, D. a. 1976. Fossil Heloderma (Reptilia: Helodermatidae). Occ. Pap. Mus. Natur. Hist. Univ. Kansas 51:1-14. Zangerl, R. 1953A. The vertebrate fauna of the Selma Formation of Alabama. Pt. 3. The turtles of the family Protostegidae. Fieldiana, Geol. Mem. 3:61-133. Zangerl, R. 1953B. The vertebrate fauna of the Selma Formation of Alabama. Pt. 4. The turtles of the family Toxochelyidae. Fieldiana, Geol. Mem. 3:135-277. Zangerl, R., and R. E. Sloan. 1960. A new spec- imen of Desmatochelys lowi Williston. A primi- tive cheloniid sea turtle from the Cretaceous of South Dakota. Fieldiana Geol. 14: No. 2:7-40. Zeller, D. E. (ed.). 1968. The stratigraphic succes- sion in Kansas. Kansas Geol. Surv. Bull. 189:1-81. ZiTTEL, K. A. 1902. Text-book of Palaeontology (Trans, a. ed. by R. Eastman) 2:i-viii, 283 pp. Macmillan. New York. ZiTTEL. K. A. 1918. Grundzuge der Paliiontologie (Palaozoologie). Vol. 2. Vertebrata. 3rd ed. iv-l-675 pp. Oldenbourg. Miinchen. ZiTTEL, K. A. 1932. Text-book of Paleontology (Sec- ond English ed., revised with additions by A. S. Woodward) 2:xvii + 464 pp. Macmillan, Lon- don. ZuG, G. R. 1966. The penial morphology and the relationships of Cryptodiran turtles. Occ. Pap. Mus. Zool. Univ. Michigan 647:1-24. 64 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY APPENDIX GENERA AND SPECIES LISTED Acroplous vorcLX Actiobales peabodyi Allosaurus sp. Ambystoma kansense kansensis Anchylorana dubita moorei robustocondyla Ancylocentrum Andrias matthewi Angistorhinus aeolanmis sp. Anura indet. Apatomerus niirus Apatosaurus lonisae Arkanserpeton arcuatum Atlantosaurus sp.? Brachysaurus overtoni overt onii Bufo arenarius hibbardi sp. Camarasaurus sp.? Captorhinus aguti Cardiocephalus cf. Sternberg! peabodyi sp. Caryoderma snovianum Chelosphargis advena Chr\sem\s hilli ' limnodytes Cimoliasaurus snovii snowii PAGE 3,6 Cimoliosaurus 6 n. sp. 7 snowii 7 sp. 49 Clidastes 49 p ropy than 16,19 pumilus 18,20 sp. 16,20 stenops tortor 12.14 1 "1 velox 12 westi westii 13 43 Cnemidophorus bilobatits 16 16 Cricotillus brachydens 47 crocodile 47,48 48 Crossotelos annulatus 14,15,16 Ctenochelys 58 procax 58 sp. stenopora 49 49 stenoporus 6 Delorhynchus 6,7 priscus 49 Desmatochelys 49 lowi lowii 43 43 43 Diplocaulus Dolichorhynchops osborni 10 willistoni 11,12 11,12 Edaphosaurus 12 ecordi sp. 49 undescribed 49 elasmosaur 20,21,22 g. et sp. indet. 21,22,23 Elasmosaurus ischiadicus 8 snowi 8 9 sternbergi Emarginachelys cretacea 3,29,30 Emydoidea 23 twentei 24 blundingi 28 Emys 28 twentei 28 Eoscopus lockardi 51 52 52 51 56 36.37,38,39 36.38 38,39 38 39 36.38 36.37,38 38 37 34 34.35 5 5 48,58 5 5 26 27 26 26,27 26,27 57 57 27 27 27 5 53,54 53,54 54 57,58 57,58 57 57 52 51.52.56 51 51.52 52.54 28 28 28 28 28 28 7 7 TYPE AND FIGURED FOSSIL AMPHIBIANS AND REPTILES 55 Eryops megacephalus Willi stoni Eumeces striatulatus Eumecoides hibbardi mylocoelus Euryodus bonneri peabodyi primus sp. Geochelone campester incisa orthopygia osborniana riggsi turgida Geringophis vetus Goniopholis sp. Gopherus riggsi Helagras orellanensis Heloderma matthewi Helodermoides tuberculatus Hesperoherpelon garnettense Hesperotestudo campester gilberti orthopygia riggsi Hyporhina galbreathi Hyposaurus labyrinthodont unidentified Lanebatrachus martini Leiocephalus etheridgei partitus Leurospondylus ultimas Liodon Mastodonsaurus sp. indet. PAGE PAGE 6 Morosaurus 49 6 grandis 49 6 mosasaur 40,46 34 !\4osasaurus 39,43 34 horridus 39 ivoensis 39 34 missouriensis 39 34 8 Neoscaphiopus 9 noblei 10 8 9 Odaxosaurus 35 8 cf. piger 36 obliquus 36 29.30 pawneensis 36 29 Ogallalabatrachus 29 horarium 19,20 3,29 Ogmodirus 51 29 ischiadicus 51 29 martini 50,51 29 martinii 50 46 Onychopus 46 gigas 20 48 Ornithostoma 48.49 48 ingens 48.49 28 species 49 28 Pancelosaurus 36 46 pawneensis 36 46 Peltosaurus 35 36 granulosus 35 36 jepseni 36 35 piger 36 35 Pentaceratops 50 7,8 cf. sternbergi 50 8 cf. sternbergii 50 28,29 Petrolacosaurus 2,30,31,57 29 kansensis 30,31,32,33,57 29 Phrynosoma 34 29 adinognathus 34 28 cornutum 34 46 cf. cornutum 34 46 Platecarpus 39.40,41,43 48 caryphaeus 39 coryphaeus 39,40,41,42 ictericus 42 9 sp. 42,43 Pleuristion 20 19,20 brachycoelus 20,21 33 plesiosaur (plesiosaurian) 56 33 Plesiosaurus 52,53 34 gouldi 52,55 gouldii 55 51 gulo 2,55 43 mudgei 52,56 Plioambystoma 19 7 kansasenese 16,18,19 7 kansense 16,17,20 kansensis 16,17,18,19,20 66 Podargosaurus hibbardi Polycotylus ischiadicus latipinnis Porthochelys laticeps Prognathodon overtoni Protostega advena gigas Pseudemvs hilli ' Pteranodon ingens sp. Rana dubita ephippium fayeae meadensis moorei panissima rexroadensis robuslocondyla sp. valida Ranidae indet. Rhineiira amblyceps hatcheri wilsoni Sauropodus dinosaur Scaphiopus antiquus diversus pliobatrachus studeri Silvisaurus condraxi Styxosaurus snovii snowii MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY 30 51,56 51 51,55 24,25,27 24 43 43 23 24 23 48.58 48 48.49 14.15,16 12 12 12 12 12 13 13 13 13 13 14 47 47 47 47 49 10 10 10 7,10 10 50 50 52 52 Testudo gilberti gilbertii orthopygia rexroadensis riggsi snoviana turgido iindata Tlialassiosaurus ischiadicus Tfirausmosaurus serratidens Toxochelys brachyrhina brachyrhinus browni latiremis procax serrifer sp. indet. slenopora stenoporus Trimerorhachis leptorhynchus Trinacromerum anonymum osborni willistoni Tylosaurus dyspelor nepaeolicus proriger sp. Xerobates undala Xestops pawneensis Xyrospondylus ecordi young plesiosaurs PAGE 29,30 29 29 30 29 28,29 30 29 30 51 51 57 57 24,25 25 24,25 25.26 24.25,26 26 26 27 26 26 5 53,54 2,54 53 54 37,43,44 45 45 43.44,45 45 30 29 36 36 58 58 56 828 RECENT MISCELLANEOUS PUBLICATIONS UNIVERSITY OF KANSAS MUSEUM OF NATURAL HISTORY 61. Reproductive strategies in a tropical anuran community. By Martha L. Crump. Pp. 1-68, 13 figures in text. November 15, 1974. Paper bound. 62. A demographic study of the ringneck snake (Diadophis punctatus) in Kansas. By Henry S. Fitch. Pp. 1-53, 19 figures in text. April 3, 1975. Paper bound. 65. The biology of an equatorial herpetofauna in Amazonian Ecuador. By William E. Duellman. Pp. 1-352, 198 figures in text. August 30, 1978. Paper bound. 66. Leptodactylid frogs of the genus Eleutherodactylus from the Andes of southern Ecuador. By John D. Lynch. Pp. 1-62, 23 figures in text. February 28, 1979. Paper bound. 67. An ecogeographic analysis of the herpetofauna of the Yucatan Peninsula. By Julian C. Lee. Fp. 1-75, 27 plates, 22 figures in text. February 29, 1980. Paper bound. 68. Internal oral features of larvae from eight anuran fam-ilies: Functional, systematic, evolutionary and ecological considerations. By Richard Wassersug. Pp. 1-146, 37 figures in text. June 24, 1980. Paper bound. 69. The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). By John D. Lynch and William E. Duellman. Pp. 1-86, 8 figures in text. August 29, 1980. Paper bound. 70. Sexual size differences in reptiles. By Henry S. Fitch. Pp. 1-72, 9 figures in text. February 27, 1981. Paper bound. 71. Late Pleistocene herpetofaunas from Puerto Rico. By Gregory Pregill. Pp. 1-72, 26 figures in text. May 8, 1981. Paper bound. 72. Leptodactylid frogs of the genus Eleutherodactylus in the Andes of northern Ecuador and adjacent Colombia. By John D. Lynch. Pp. 1-46, 22 figures in text. July 8, 1981. Paper bound. 73. Type and figured specimens of fossil vertebrates in the collection of the University of Kansas Museum of Natural History. Part I. Fossil fishes. By H.-P. Schultze, J. D. Stewart, A. M. Neuner and R. W. Coldiron. Pp. 1-53. October 6, 1982. Paper bound. 74. Relationships of pocket gophers of the genus Geomys from the central and northern Great Plains. By Lawrence R. Heaney and Robert M. Timm. Pp. 1-59, 19 figures in text. June 1, 1983. Paper bound. 75. The taxonomy and phylogenetic relationships of the hylid frog genus Stefania. By William E. Duellman and Marinus S. Hoogmoed. Pp. 1-39. March 1, 1984. Paper bound. 76. Variation in clutch and litter size in New World reptiles. By Henry S. Fitch. Pp. 1-76. May 24, 1985. Paper bound.