QE 841 . U62 ISITY OF KANSAS miscellaneous OT ^ M OF NATURAL HISTORY publication ^ ' • ^ No. 78 Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part III. Fossil Birds By John F. Neas and Marion Anne Jenkinson UNIVERSITY OF KANSAS LAWRENCE 1986 February 5, 1986 5?r A, .1-^ UNIVERSITY OF KANSAS PUBLICATIONS MUSEUM OF NATURAL HISTORY The University of Kansas Publications, Museum of Natural History, beginning with volume 1 in 1946, was discontinued with volume 20 in 1971. Shorter research papers formerly published in the above series are now published as Occasional Papers, Museum of Natural History. The Miscellaneous Publications, Museum of Natural History, began with number 1 in 1946. Longer research papers are published in that series. Monographs of the Museum of Natural History were initiated in 1970. All manuscripts are subjected to critical review by intra- and extramural specialists; final acceptance is at the discretion of the Director. Institutional libraries interested in exchanging publications may obtain the Occasional Papers and Miscellaneous Publications by addressing the Exchange Librarian, University of Kansas Library, Lawrence, Kansas 66045. Individuals may purchase separate numbers of all series. Prices for all publications of the Museum may be obtained from the Publications Secretary, Museum of Natural History, University of Kansas, Lawrence, Kansas 66045. HARVARD UNIVERSITY Library of the Museum of Comparative Zoology The University of Kansas Museum of Natural History Miscellaneous Publication No. 78 February 5, 1986 Type and Figured Specimens of Fossil Vertebrates in the Collection of the University of Kansas Museum of Natural History Part III. Fossil Birds By John F. Neas and Marion Anne Jenkinson Museum of Natural History The University of Kansas Lawrence, Kansas 66045 U.S.A. The University of Kansas Lawrence 1986 University of Kansas Publications, Museum of Natural History Editor: Robert M. Mengel Managing Editor: Joseph T. Collins Miscellaneous Publication No. 78 Pp. 1-14 Published February 5, 1986 MCZ LIBRARY APR 8 1986 HARVARD UNIVERSITY Museum of Natural History The University of Kansas Lawrence, Kansas 66045 U.S.A. ^^ Printed By University of Kansas Printing Service Lawrence, Kansas CONTENTS INTRODUCTION I SHORT HISTORY OF THE AVIAN FOSSIL COLLECTION 1 ACKNOWLEDGMENTS 2 SUMMARY OF CLASSIFICATION 2 CATALOGUE 4 Class Aves 4 Subclass Ornithurae 4 Infraclass Odontoholcae 4 Order Hesperornithiformes 4 Infraclass Neornithes 5 Order Podicipediformes 5 Order Anseriformes 6 Order Galliformes 7 Order Ralliformes 7 Order Ichthyornithiformes 8 Order Charadriiformes 9 Order Columbiformes 10 Order Cuculiformes 10 Order Strigiformes 10 OrderPiciformes 11 Order Passeriformes 11 Vestigia Avium - 12 LITERATURE CITED 12 APPENDIX: GENERA AND SPECIES LISTED 14 INTRODUCTION The present work is part three of a four part series designed to list type and figured specimens of fossil vertebrates in the Uni- versity of Kansas Museum of Natural His- tory (KUVP), as recommended by the International Code of Zoological Nomen- clature. The classification employed follows that of Martin (1983) for higher taxonomic cate- gories. Unless otherwise stated, ordinal to specific nomenclature is that of Brodkorb (1963, 1964, 1967, 1971, 1978). Each spec- imen in the catalogue is provided with the following information: current catalogue number and any previous number used in publication; author and year of publication; pagination of type description; figure(s); preserved anatomical parts of the specimen; geologic stage, formation, member (see Zeller 1968), land mammal age; locality and collector; any pertinent remarks. For each species the holotype, if any, is listed first, followed by paratypes and figured spec- imens. In accordance with what we under- stand will be recommended in the Third Edition of the International Code of Zoo- logical Nomenclature (probably to be pub- lished in 1985), we include as paratypes all remaining specimens from a type series from which a holotype was designated. Herein we use quotation marks around the word "paratype" if the author(s) did not use that word when referring to the specimen in question. SHORT HISTORY OF THE AVIAN FOSSIL COLLECTION Several factors seem particularly impor- tant in the early development of avian pale- ontology at the University of Kansas. One is the presence of the Niobrara Cretaceous chalk beds in western Kansas. These sedi- ments have been actively collected by pale- ontologists since about 1870, the year in which Professor O. C. Marsh of Yale Uni- versity discovered the first North American Mesozoic bird remains. Subsequent collect- ing by Benjamin Mudge, a geology profes- sor at Kansas State University, Manhattan, produced the first known fossil birds with teeth, which Marsh announced in 1873. Most specimens of Hesperornis, Ich- thyornis, and Baptornis known to science come from the Niobrara Chalk of Kansas and many were collected in the late 1800's and early 1900's by Mudge, Samuel Wen- dell Williston, Handel Tong Martin, and Charles and George Sternberg. Of these, only Martin added much avian fossil mate- rial to the collections at the University of Kansas; one specimen collected by him now proves to be different from other recognized species of toothed birds. Martin also col- lected fossils of Phorusrhacos in Patagonia (although only some of this material re- mained in KU's collection) and the type specimen of Griis nannodes from Edson Quarry, Sherman County, Kansas. Similarly important was the association with the museum of Charles Dean Bunker, between 1895 and 1901 and again from 1905 until his retirement in 1942. Bunker inspired an entire generation of KU students of natu- ral history (including William H. Burt, The- odore Downs, David H. Dunkle, E. Raymond Hall, Claude W. Hibbard, John E. Hill, Remington Kellogg, R. A. Sfirton, Alexander Wetmore, and Theodore E. White); many of these became leaders in their fields. Bunker oversaw the collections of both Recent birds and mammals; he also spent a summer in the field with Charles Sternberg. Especially important is the fact that he recognized the value of skeletal material of Recent birds long before most other curators did. His efforts at collecting and preparing skeletal specimens, and those of the students he inspired, resulted in the establishment at KU of one of the world's great collections of avian skeletal specimens. Finally, the fossil avian collection was greatly expanded during the 1930's and 1940's as a result of C. W. Hibbard's development of techniques for collecting microfossils. More than 1,000 fossil avian specimens have been added to the museum collection as a result of numerous field parties conducted by Hibbard at several important Pliocene (Rexroad and Keefe Canyon) and Pleistocene (Borchers. Jones, Sunbrite, Kentuck, Rezabek, and Cudahy) local faunas. Other field parties led by Hib- bard collected in Oklahoma and Nebraska. With the exception of specimens described by Wetmore and others, which include six MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY holotypes, much of the bird material col- lected by Hibbard has been studied only to a minor extent. A hiatus in fossil bird collecting fol- lowed Hibbard' s departure in 1946 and, with few exceptions, continued unbroken for more than two decades. Notable among the exceptions is a collection of many hundreds of bird fossils discovered with bat, insec- tivore, reptile, and amphibian remains that were obtained by James W. Bee in 1957 from three Pleistocene Puerto Rican cave faunas. Also, a small assemblage of late Pleistocene birds was collected from Doni- phan County, Kansas, by Sudi Einsohn in 1969. The appointment of Larry D. Martin as curator in 1972 ushered in an era of revived interest in fossil birds at KU. The museum acquired important specimens of Cretaceous toothed birds discovered by Marion Bonner and his sons Orville and Chuck. These specimens include a partial, juvenile skel- eton of Baptornis advenus collected by Or- ville in 1962, a complete skull of Hesperor- nis discovered by Marion and Chuck in 1981, and a partial skeleton of Parahesper- omis collected by Orville, in the same year. During the same period, J. D. Stewart col- lected seven specimens of Ichthyornis from the Niobrara Cretaceous and more than 100 bird fossils from two new Pleistocene avi- faunas in Kansas (Trap Shoot and Hill City local faunas). Excavations begun in 1975 at Natural Trap Cave (Wyoming) by L. D. Martin and B. Miles Gilbert have yielded a number of Pleistocene and Holocene birds. The museum also houses more than 300 Miocene specimens of birds from the Pungo River Formation of North Carolina collected by Jerry Case, and from the Big Sandy Formation of Arizona collected by Dan Adams around 1975. The Divisions of Pale- ontology and Ornithology have jointly devel- oped a collection of casts of many fossil avian holotypes from around the world. The entire collection of fossil birds has now been entered onto the computer and inventoried, through the efforts of the two authors, plus J. D. Stewart and Kenneth Whetstone. As a result, the 12 holotypic and 17 paratypic specimens (plus the 30 figured specimens which are not holotypes or para- types) have all been located. All of the holotypes and paratypes are valid; 3 of the holotypes and 6 paratypes of another species represent the type-species of their genera. ACKNOWLEDGMENTS We express our gratitude to L. D. Martin for sharing his vast knowledge of the collec- tion with us, to H.-P. Schultze for his assistance with the manuscript, and to Or- ville Bonner, Pierce Brodkorb, Darrel Frost, Robert M. Mengel, J. D. Stewart, Robert W. Wilson, and Glen E. Woolfenden, for their assistance with various matters. Hilde- garde Howard read the manuscript and made valuable suggestions for its improvement. The compilation of the catalogue was supported by the National Science Founda- tion, Grant no. DEB-81 15956. SUMMARY OF CLASSIFICATION Class Aves Subclass Ornithurae Infraclass Odontoholcae Order Hesperornithiformes Family Hesperornithidae Family Baptornithidae Infraclass Neornithes Order Podicipediformes Family Podicipedidae Order Anseriformes Suborder Anseres Family Anatidae Subfamily Anatinae Subfamily Aythyinae Order Galliformes Family Cracidae Family Phasianidae Subfamily Odontophorinae Order Ralliformes Suborder Ralli Family Rallidae Subfamily Rallinae Subfamily Gallinulinae Subfamily Fulicinae Suborder Grues Family Gruidae Subfamily Gruinae Order Ichthyornithiformes Family Ichthyornithidae TYPE AND FIGURED FOSSIL VERTEBRATES AT UNIVERSITY OF KANSAS Order Charadriiformes Suborder Charadrii Family Scolopacidae Subfamily Scolopacinae Family Recurvirostridae Subfamily Presbyornithinae Family Burhinidae Order Columbiformes Suborder Columbae Family Columbidae Subfamily Columbinae Order Cuculiformes Suborder Cuculi Family Cuculidae Order Strigiformes Family Protostrigidae Order Piciformes Suborder Galbulae Family Primobucconidae Order Passeriformes Suborder Passeres Family Icteridae Family Emberizidae Vestigia Avium MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY CATALOGUE Class Aves Linnaeus 1758 Subclass Ornithurae Haeckel 1866 Infraclass Odontoholcae (Stejneger 1885) Order Hesperornithiformes (Fiirbringer 1888) Family Hesperomithidae Marsh 1872 Parahesperomis Martin 1984 Parahesperomis alexi Martin 1984 (Type-species) KUVP 2287 Parahesperomis alexi— Uorim 1984: pp. 143-145; figs. IB-C, 2B, 2E, 2G, 3A, 3C, 4A; tarsometatarsus, lacrimal, coracoid, dorsal and lateral views of restored skull, lateral view of restored skeleton. Martin (1983: p. 317) cited KUVP 2287 as the holotype of Parahesperomis alexi, but it is doubtful that his discussion in that publication can be considered as a diagnosis. We think Martin's 1984 publication should be considered as establishing the name Para- hesperomis alexi. KUVP 2287 is a nearly complete skeleton, includ- ing tarsal scutes and feather impressions. Parts of KUVP 2287 have been figured in a number of publications, under several names: Hesperornis—^iWision 1896: pi. II (showing tarsometatarsus and scute impressions). This same figure appeared in Williston 1898 as pi. VIII. Hesperornis gracilis— Lucas 1903: p. 552; fig. 1-2; quadrate and pterygoid. As Gingerich (1976, p. 27) indicated, the left pterygoid in Lucas's figure 2 is erroneously labelled as a right pterygoid. Lucas established the genus Hargeria for Yale Peabody Museum specimen 1473, the holotype of Hesperornis gracilis. He assumed, however, that KUVP 2287 was conspecific with YPM 1473 and used the characters of the former to distinguish the genus. Hesperornis— Gregory 1951: fig. 2A-C; a restoration of the lower jaw. This same figure, labelled Hesperornis gracilis Marsh, appeared in Gregory 1952 as fig. 7A-C. Hargeria (Hesperornis) gracilis-Schmidt and Keil 1958: fig. 106, 167, 235, 236; cross-sections of teeth. These same illustrations appeared in Schmidt and Keil 1971, as figures 113, 240, 333, 334. Hesperornis regah s—Sv/inton 1975: fig. 15 ( = Gregory 1951, fig. 2A- C). Swinton thought this might possibly be a mosasaur jaw. Hesperornis— G'mgench 1976: fig. 2a-b, 3a-b; ventral view of the skull, quadrate, and pterygoid. Hesperornithid— Martin, Stewart, and Whetstone 1980: tig. 2A, 2D; stereophotographs of complete teeth. Parahesperomis a/ev/— Martin 1983: fig. 9.6B, 9. 73; restorations of the complete skeleton and skull. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; probably west of Hill City, Graham County, Kansas; collected by H. T Martin in 1894. Family Baptornithidae American Ornithologists' Union 1910 (placed in Podicipediformes by Brodkorb, 1963) Baptornis Marsh 1877 Baptornis advenus Marsh 1 877 TYPE AND FIGURP:D FOSSIL VERTEBRATES AT UNIVERSITY OF KANSAS KUVP 2290 Baptornis advenus— Lucas 1903: fig. 6-8; coracoid, partial scapula, radius, ulna, humerus, and patella. The posterior (sternal) end of right coracoid figured by Lucas (1903, fig. 6) appears incorrectly rotated 180°. Compare Martin and Tate, 1976 (fig. 9c, 10b). Baptornis advenus—Martm and Tate 1976: fig. 3d-f, 5b, 9b (based partially on this specimen), 9c-d, 10b, llb-d, 12a-b, 13b, 13d, 16c, 16e, 19 (a composite restoration, only the wing and shoulder girdle being based on this specimen); cervical and thoracic vertebrae, a scapula, coracoid, humerus, radius, ulna, proximal and distal tar- sometatarsi, and synsacrum. Baptornis advenus—Mari'in 1983: fig. 9.6C (modified from Martin and Tate 1976, fig. 19). KUVP 2290 also consists of additional vertebrae, fragments of ribs and pelvis, both femora, both tibiotarsi, and the proximal end of a fibula, none of which has been figured. Upper Cretaceous, Santonian or Campanian, Niobrara Formation, Smoky Hill Chalk Member; ?Logan County, Kansas. KUVP 16112 Baptornis advenus— Martin and Tate 1976: fig. 19; a composite restora- tion, the tip of the bill being based on this specimen. Baptornis advenus— Martin and Bonner 1977: fig. lA, IC, IF (incor- rectly stated as KUVP 16122) and II; premaxillary fragment, right femur, distal end of left tibia without tarsals, and left metatarsal without tarsals. KUVP 16112, an immature specimen, also includes vertebrae, synsacrum, the left femur, the right tibia, the right metatarsal, phalanges, and other fragments, none of which has been figured. Upper Cretaceous, Upper Santonian or Lower Campanian, Niobrara Formation, Smoky Hill Chalk Member; Willow Canyon, 10 miles south of Russell Springs, SE'A. NW'A, Sec. 13. T15S. R35W, Logan County, Kansas; collected by O. Bonner in 1962. Infraclass Neornithes (Gadow 1893) Order Podicipediformes (Fiirbringer 1 Family Podicipedidae (Bonaparte 1831) Colymbus Linnaeus 1758 see Podiceps Latham 1787 Podiceps Latham 1787 Podiceps caspicus (Hablizl 1783) KUVP 5676 Colymbus caspicus— Downs 1954: fig. la; complete left carpometacar- pus. Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mam- mal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Order Anseriformes (Wagler 1831) Suborder Anseres Wagler 1831 Family Anatidae Vigors 1825 Subfamily Anatinae (Vigors 1825) Anas Linnaeus 1758 Anas acuta Linnaeus 1766 KUVP 5641 Anas acuta— Downs 1954: fig. lb, right figure; anterior (furcular) end of left coracoid. KUVP 5644 Anas acuta— Downs 1954: fig. lb, left figure; anterior (furcular) end of left coracoid. 5641, 5644: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. Anas bunkeri (Wetmore 1944) KUVP 3982 Nettion bunkeri— Wetmore 1944: pp. 92-94; fig. 1-3; holotype, right carpometacarpus without metacarpal III (called metacarpal IV by Wetmore). Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age), Rexroad local fauna; NW'A, SW'A, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Anas clypeata Linnaeus 1758 KUVP 5639 Anas clypeata— Downs 1954: fig. Ig; complete left coracoid. Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mam- mal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. Anas sp. KUVP 5649 Anas (teal)— Downs 1954: fig. Id; right carpometacarpus missing meta- carpal III. KUVP 5679 Anas (teal)— Downs 1954: fig. Ic; anterior (furcular) end of right coracoid. KUVP 5683B See KUVP 25464 KUVP 5683C See KUVP 25465 KUVP 25464 Anas (teal)— Downs 1954: fig. le; figured under former no. 5683B; proximal end of left tibiotarsus. KUVP 25465 Anas (teal)— Downs 1954: fig. If; figured under former no. 5683C; distal end of left tibiotarsus. 5649, 5679, 25464, 25465: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. Nettion Kaup 1829 Nettion bunkeri Wetmore 1944 see Anas bunkeri (Wetmore 1944) Subfamily Aythyinae (Delacour and Mayr 1945) Aythya Boie 1822 Aythya sp. KUVP 5652 /lv//7V«— Downs 1954: fig. KUVP 6052 Avthya—Downs 1954: fig. KUVP 6057 Aythya —Downs 1954: fig. li; proximal fragment of right humerus. Ih; complete left coracoid. Ij; distal left humerus. TYPE AND FIGURED FOSSIL VERTEBRATES AT UNIVERSITY OF KANSAS KUVP 9901 KUVP 3981 KUVP 3997 KUVP 3865 KUVP 3867 KUVP 3869 KUVP 3870 KUVP 3871 KUVP 3872 KUVP 3866 KUVP 3868 5652, 6052, 6057: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by J. Tihen and party in 1940 (except 5652 which was collected by C. W. Hibbard and party in 1939). Order Galliformes (Temminck 1820) Family Cracidae Vigors 1825 Cracid, gen. et sp. indet. Cracid, genus and species indeterminate— Martin and Mengel 1984: p. 176; fig. IG-K; distal right tibiotarsus. Tertiary, early Miocene, Pawnee Creek Formation (Hemingfordian land mammal age), Martin Canyon local fauna; "Quarry A," NE'A, Sec. 27, TUN, R53W, Logan County, Colorado; collected by R. W. Wilson and party. Family Phasianidae Vigors 1825 Subfamily Odontophorinae Gould 1844 Colinus GoWuss 1820 Colinus hibbardi Wetmore 1944 Colinus hibbardi— Wetmore 1944: pp. 96-98; fig. 4-5; holotype, right tarsometatarsus missing trochleae for digits II and IV. Colinus hibbardi— Wetmore 1944: fig. 6-8; "paratype," distal right humerus. 3981, 3997: Tertiary, Pliocene, Rexroad Formation (Blancan land mam- mal age), Rexroad local fauna; NW'A, SWA, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Order Ralliformes (Reichenbach 1852) Suborder Ralli (Reichenbach 1852) Family Rallidae Vigors 1825 Subfamily Rallinae (Vigors 1825) RaUus Linnaeus 1758 Rallus prenticei Wetmore 1944 RaUus prenticei— Wetmore 1944: pp. 99-103 plete right humerus. RaUus prenticei— Wetmore 1944: p. 101; fig plete right coracoid. RaUus prenticei— Wetmore 1944: p. sus, distal end. RaUus prenticei— Wetmore 1944: p. distal end. RaUus prenticei— Wetmore 1944: p missing head. RaUus prenticei— Wetmore 1944: p fig. 9-12; holotype, com- 13-14; "paratype," com- 01; "paratype," right tarsometatar- 102; "paratype," right tibiotarsus, 102; "paratype," right humerus, 102; fig. 15-19; "paratype," left tibiotarsus without proximal end. 3865, 3867, 3869-3872: Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age), Rexroad local fauna; NE'A, NW'A, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1936 and (for specimen 3867) in 1937. RaUus prenticei— Wetmore 1944: p. 100; "paratype," left humerus, distal two-thirds. RaUus prenticei— Wetmore 1944: p. 101; "paratype," left coracoid, without sternal end. MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY 3866, 3868: Tertiary, Pliocene, Rexroad Formation (Blancan land mam- mal age), Rexroad local fauna; NW'A, SWA, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Subfamily Gallinulinae Gray 1840 Gallinula Brisson 1760 Gallinula kansarum Brodkorb 1967 KUVP 3994 Gallinula kansarum— Brodkorb 1967: p. 125 (footnote); holotype, distal part of left humerus. Brodkorb notes Wetmore's (1944, p. 103) referral of this specimen to Fulica americana Gmelin. The holotype is first figured by Feduccia (1968; fig. 3, two figures on right). On page 448 Feduccia erroneously refers to this figure as "fig. 1." KUVP 3988 Gallinula kansarum— Brodkorb 1967: p. 125 (footnote); "paratype," right ulna, without olecranon, "referred only tentatively" to this species by Brodkorb. 3994, 3988: Tertiary, Pliocene, Rexroad Formation (Blancan land mam- mal age), Rexroad local fauna; NW'A, SWA, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Subfamily Fulicinae (Nitzsch 1820) Fulica Linnaeus 1758 Fulica americana Gmelin 1789 see Gallinula kansarum Brodkorb 1967 Suborder Grues Bonaparte 1854 Family Gruidae Vigors 1825 Subfamily Gruinae (Vigors 1825) Grus Pallas 1766 Grus nannodes Wetmore and Martin 1930 KUVP 3757 Grus nannodes— WetmorQ and Martin 1930: pp. 62-63; fig. 23-25; holotype, left carpometacarpus without proximal end. The figure of the carpometacarpus shows an intact metacarpal III (which Wetmore and Martin called metacarpal IV). It is now broken and much of the shaft is missing. According to Wetmore and Martin, part of the extreme distal end of the "second metacarpal" ( = metacarpal I) is present, but this is not apparent to us, either in their figures or in the existing specimen. Tertiary, late Miocene, Ogallala Formation (Hemphillian land mammal age); Edson Quarry, SW'A, Sec. 25, TIOS, R38W, Sherman County, Kansas; collected by H. T. Martin in 1924. Order Ichthyornithiformes (Marsh 1873) Family Ichthyornithidae (Marsh 1873) Ichthyornis (Marsh 1872) KUVP 2294 Ichthyornis sp. —Martin and Stewart 1982: fig. IB; presacral vertebra. Upper Cretaceous, Upper Santonian or Lower Campanian, Niobrara Formation, Smoky Hill Chalk Member; 10 miles south of Russell Springs, Willow Canyon, Sec. 11, 12, 13, T15S, R35W, Logan County, Kansas. TYPE AND FKiURED FOSSIL VERTEBRATES AT UNIVERSll Y OF KANSAS Order Charadriiformes (Huxley 1867) Suborder Charadrii (Huxley 1867) Family Scolopacidae Vigors 1825 Subfamily Scolopacinae (Vigors 1825) Bartramia Lesson 1831 Bartramia longicauda (Bechstein 1831) KUVP 5653 Bartramia longicauda—Dowm 1954: fig. Im, right figure; proximal end of right humerus. KUVP 5654 Bartramia loni^icauda—Downs 1954: fig. In: erroneously published as 5659; right humerus without distal end. KUVP 5655 Bartramia longicauda— Downs 1954: fig. lo; distal end of right humerus. KUVP 5659 Bartramia longicauda— Downs 1954: fig. Im, left figure; erroneously published as no. 5654; proximal end of left humerus. KUVP 6394 Bartramia longicauda— Downs 1954: fig. lk-1; complete left tar- sometatarsus. 5653-5655, 5659, 6394: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939 (except for specimens 5654 and 6394, which were collected in 1941). Erolia Vieillot 1816 see Calidris Merrem 1804 Calidris Merrem 1 804 KUVP 6064 £rc»//a— Downs 1954: fig. 5a; complete left tarsometatarsus. Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mam- mal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by J. Tihen and party in 1940. In 1973, the genus Erolia was merged with Calidris (American Ornithologists' Union 1973, p. 415). Family Recurvirostridae Bonaparte 1831 Subfamily Presbyornithinae (Wetmore 1926) Coltonia Hardy 1959 Coltonia recurvirostra Hardy 1959 (Type-species) KUVP 10105 Coltonia recurvirostra— Yiairdy 1959: pp 106-108; fig. 1; holotype, associated distal left humerus, left radius and ulna, radiale, ulnare, and proximal left carpometacarpus. Tertiary, Eocene, Colton Formation (Wasatchian land mammal age), mouth of Ephraim Canyon, Wasatch Plateau, Sanpete County, Utah; collected by K. E. Boker in 1953. This enigmatic group of birds has had an uncertain status ever since Wetmore (1926) first described Presbyomis and allocated it to a new family (Presbyornithidae). According to Olson and Feduccia (1980, p. 22) "until more detailed comparisons are made of the osteology of Presbyor- nis, firm recommendations as to the taxonomic status of the Presbyor- nithidae cannot be made. ... we feel that Presbyomis probably would have to be considered as still having had a charadriiform grade of morphology." 10 KUVP 6822 KUVP 5650 KUVP 3995 KUVP 25629 KUVP 25630 KUVP 25631 KUVP 16601 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Family Burhinidae Mathews 1913 Burhinus Illiger 1811 Burhinus aquilonahs Feduccia 1980 Burhinus aquilonaris— Feduccia 1980: pp. 115-117; fig. 1; holotype, complete left humerus. Bearing same number and data are associated paratypes: left radius and broken left ulna, fragmentary left ilium, and rib fragments. Quaternary, Pleistocene, Sanborn Formation (Rancholabrean land mam- mal age); NE'A, Sec. 3, T3S, R27W, Decatur County, Kansas; collected by C. W. Hibbard in 1943. Order Columbiformes (Latham 1790) Suborder Columbae Latham 1790 Family Columbidae (Illiger 1811) Subfamily Columbinae (Illiger 1811) Zenaidura Bonaparte 1855 Zenaidura macroura (Linnaeus 1758) Zenaidura macroura— Downs 1954: fig. 5b; proximal end of right carpometacarpus. Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mam- mal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939. Zenaidura prior Brodkorb 1969 Zenaidura /7nor— Brodkorb 1969: pp. 174-175; fig. 1; holotype, left humerus without distal end. Tertiary, Pliocene, Rexroad Formation (Blancan land mammal age), Rexroad local fauna; NW'A, SWA, Sec. 22, T33S, R29W, Meade County, Kansas; collected by C. W. Hibbard and party in 1937. Order Cuculiformes (Wagler 1830) Suborder Cuculi Wagler 1830 Family Cuculidae Vigors 1825 Cursoricoccyx Martin and Mengel 1984 Cursoricoccyx geraldinae Martin and Mengel 1984 (Type-species) Cursoricoccyx geraldinae— Mart'm and Mengel 1984: pp. 172-174; fig. 2B-F; holotype, proximal right tarsometatarsus. Cursoricoccyx geraldinae— Martm and Mengel 1984: fig. 2G-J; "para- type," anterior (furcular) end of right coracoid. Cursoricoccyx geraldinae— Man'm and Mengel 1984: fig. lA-F; "para- type," left carpometacarpus. 25629-25631: Tertiary, Miocene, Pawnee Creek Formation (Heming- fordian land mammal age), Martin Canyon local fauna; "Quarry A," NEV4, Sec. 27, TUN, R53W, Logan County, Colorado; collected by R. W. Wilson and party. Order Strigiformes (Wagler 1 830) Family Protostrigidae Wetmore 1933 fo^mjc Brodkorb 1971 Eostrix martinellii Martin and Black 1972 Eostrix martinellii-Marnn and Black 1972: pp. 887-888; fig. lA-C; holotype, distal end of left tarsometatarsus. Tertiary, Eocene, Wind River Formation, Lysite Member (Wasatchian TYPE AND FIGURED FOSSIL VERTEBRATES AT UNIVERSITY OF KANSAS 1 land mammal age); NW'A, Sec. 22, T39N, R90W; Cottonwood Creek, Fremont County, Wyoming; collected by J. Martinelli in 1970. Er- roneously published as NE'A, Sec. 22, T90W. Order Piciformes (Meyer and Wolf 1810) Suborder Galbulae Fiirbringer 1888 Family Primobucconidae Feduccia and Martin 1976 Uintornis Marsh 1872 Uintornis marionae Feduccia and Martin 1976 KUVP 26906 Uintornis marionae— Feduccia and Martin 1976: p. 108; fig. 5a-d; holotype, distal end of left tarsometatarsus missing trochlea for digit IV. This specimen is incorrectly called a right tarsometatarsus in the holotype designation but correctly referred to in the figure legend as a left tarsometatarsus. Tertiary, Eocene, Bridger Formation (Bridgerian land mammal age); Sage Creek, Sweetwater County, Wyoming. Collected by H. H. Lane and H. T. Martin in 1927. Uintornis was originally allocated to the Picidae by Marsh (1872) and later was placed in the Cuculiformes by Cracraft and Morony (1969) and Brodkorb (1970, 1971). Order Passeriformes (Linnaeus 1758) Suborder Passeres Linnaeus 1766 Family Icteridae (Vigors 1825) Agelaius Vieillot 1816 KUVP 5648 lAgelaius— Downs 1954: fig. 5c; complete right carpometacarpus. For locality description, see under KUVP 6067, Icteridae. Molothrus Swainson 1832 KUVP 5669 ? Molothrus— Downs 1954: fig. 5d; right carpometacarpus missing meta- carpal III. For locality description, see under KUVP 6067, Icteridae. KUVP 6062 1 Molothrus— Downs 1954: fig. 5f; proximal end of left humerus. For locality description, see under KUVP 6067, Icteridae. Icteridae, gen. et sp. indet. KUVP 6067 Icteridae— Downs 1954: fig. 5e; proximal end of left ulna. 5648, 5669, 6062, 6067: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S. R27W, Meade County, Kansas; 5648 and 5669 collected by C. W. Hibbard and party in 1939; 6062 and 6067 collected by J. Tihen and party in 1940. KUVP 5647 KUVP 5647A KUVP 5647B KUVP 25467 Family Emberizidae Vigors 1831 Calamospiza Bonaparte 1838 Calamospiza melanocorys Stejneger 1885 Calamospiza melanocorys— Downs 1954: fig. 5g-h; formerly no. 5647 A. published erroneously as no. 5674; rostrum. For description of locality, see under KUVP 6068, Calcarius. See KUVP 5647 See KUVP 25467 Calamospiza melanocorys— Downs 1954: fig. 5k-l; formerly no. 5647B, published erroneously as no. 5674B; rostrum. For description of locality, see under KUVP 6068, Calcarius. 12 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY Calcarius Bechstein 1802 KUVP 6068 Ca/canM^— Downs 1954: fig. 5i-j; rostrum. 5647, 6068, 25467: Quaternary, Pleistocene, Vanhem Formation (Rancholabrean land mammal age), Jones local fauna; Sec. 8, T33S, R27W, Meade County, Kansas; collected by C. W. Hibbard and party in 1939 (except 6068 which was collected by J. Tihen and party in 1940). Pedinorhis Olson and McKitrick 1981 Pedinorhis stirpsarcana Olson and McKitrick 1981 (Type-species) KUVP 37051 Pedinorhis stirpsarcana— Ohon and McKitrick 1981: p. 280; paratype, rostrum. KUVP 37052 Pedinorhis stirpsarcana— Olson and McKitrick 1981: p. 280; paratype, rostrum. KUVP 37053 Pedinorhis stirpsarcana— Olson and McKitrick 1981: p. 280; paratype, rostrum. KUVP 37054 Pedinorhis stirpsarcana— Olson and McKitrick 1981: p. 280; fig. ID-F; paratype, rostrum. KUVP 37055 Pedinorhis stirpsarcana— Olson and McKitrick 1981: p. 280; paratype, rostrum. KUVP 37056 Pedinorhis stirpsarcana— Olson and McKitrick 1981: p. 280; paratype, mandible. 37051-37056: Quaternary, Late Pleistocene; Cuevo del Perro, Ciales Valley, about 2 kilometers south of Barrio de Barahona. Municipio de Morovis, 18°20'57"N, 66°26'47"W, Puerto Rico; collected by J. W. Bee in 1957. Vestigia Avium KUVP 65696 Bird Track— Snow 1887: text fig. on p. 4. Bird Track— Williston 1898: fig. 2 ( = Snow 1887, text fig. on p. 4). Upper Cretaceous, Cenomanian, Dakota Formation; Thompson's Creek, Ellsworth County, Kansas; collected by E. P. West in 1885. Williston (1898, p. 53) agreed with Snow (1887, p. 4) that the track is "in all probability that of a bird. ' ' Currie ( 198 1 , p. 264) refers the track to Irenesauripus, a saurischian track. Neas compared KUVP 65696 with a published illustration of a track of Irenesauripus and thinks that the two show little similarity. The Kansas University specimen indicates the presence of a hallux and the divarication of the toes is small— less than 90°. LITERATURE CITED American Ornithologists' Union. 1973. Thirty- second supplement to the American Or- nithologists" Union Check-list of North Amer- ican birds (Check-list Committee, E. Eisenmann, Chr.). Auk 90: 411-419. Brodkorb, p. 1963. Catalogue of fossil birds: Part 1 (Archaeopterygiformes through Ardeiformes). Bull. Florida State Mus. 7:179-293. Brodkorb, P. 1964. Catalogue of fossil birds: Part 2 (Anseriformes through Galliformes). Bull. Flor- ida State Mus. 8:195-335. Brodkorb, P. 1967. Catalogue of fossil birds: Part 3 (Ralliformes, Ichthyornithiformes, Charadri- iformes). Bull. Florida State Mus. 2:99-220. Brodkorb, P. 1969. An ancestral Mourning Dove from Rexroad, Kansas. Quart. J. Florida Acad. Sci. 31:173-176. Brodkorb, P. 1970. The paleospecies of woodpeckers. Quart. J. Florida Acad. Sci. 33:132-136. Brodkorb, P. 1971. Catalogue of fossil birds: Part 4 (Columbiformes through Piciformes). Bull. Florida State Mus. 15:163-266. Brodkorb, P. 1978. Catalogue of fossil birds: Part 5 (Passeriformes). Bull. Florida State Mus. 23:139-228. Cracraft, J., and J. J. MoRONY, Jr. 1969. A new Pliocene woodpecker, with comments on the fossil Picidae. Amer. Mus. Novitates, 2400:1-8. TYPE AND FIGURED FOSSIL VERTEBRATES AT UNIVERSITY OF KANSAS 13 CuRRiE. R J. 1981. Bird footprints from the Gething Formation (Aptian, Lower Cretaceous) of northeastern British Columbia, Canada. J. Vert. Paleont. 1:257-264. Downs, T. 1954. Pleistocene birds from the Jones Fauna of Kansas. Condor 56:207-221. Feduccia, a. 1968. The Pliocene rails of North America. Auk 85:441-453. Feduccia, A. 1980. A thick-knee (Aves: Burhinidae) from the Pleistocene of North America, and its bearing on Ice Age climate. In: Papers honoring Hildegarde Howard. Sci. Natur. Hist. Mus. Los Angeles Co. 330:115-118. Feduccia, A., and L. D. Martin. 1976. The Eocene zygodactyl birds of North America (Aves: Piciformes). In: Collected papers in avian pale- ontology honoring the 90th birthday of Alex- ander Wetmore. Smiths. Contr. to Paleobiol. No. 27: 101-110. GiNGERiCH, P. D. 1976. Evolutionary significance of the Mesozoic toothed birds. In: Collected papers in avian paleontology honoring the 90th birthday of Alexander Wetmore. Smiths. Contr. to Pa- leobiol. No. 27:23-33. Gregory, J. T. I95I. Convergent evolution: the jaws of Hesperornis and the mosasaurs. Evolution 5:345-354. Gregory, J. T. 1952. The jaws of the Cretaceous toothed birds, Ichlhyornis and Hesperornis. Condor 54:73-88. Hardy, J. W. 1959. A previously undescribed recur- virostrid from the Eocene of Utah. Auk 76:106-108. Lucas, F. A. 1903. Notes on the osteology and relationship of the fossil birds of the genera Hesperornis, Hargeria, Baptornis, and Di- atryma. Proc. U.S. Nat. Mus. 26:545-556. Marsh, O. C. 1872. Notice of some new Tertiary and Post-Tertiary birds. Amer. J. Sci. 4:256-262. Martin, L. D. 1983. The origin and early radiation of birds. In: Perspectives in ornithology. Essays presented for the centennial of the American- Ornithologists' Union. Chapter 9:291-338. Cambridge Univ. Press. Martin, L. D. 1984. A new hesperornithid and the relationships of the Mesozoic birds. Trans. Kan- sas Acad. Sci. 87:141-150. Martin, L. D., and C. C. Black. 1972. A new owl from the Eocene of Wyoming. Auk 89:887-888. Martin, L. D., and O. Bonner, 1977. An immature specimen of Baptornis advenus from the Cre- taceous of Kansas. Auk 94:787-789. Martin, L. D., and R. M. Mengel. 1984. A new cuckoo and a chachalaca from the early Miocene of Colorado. In: Papers in vertebrate paleontol- ogy honoring Robert Warren Wilson. Carnegie Mus. Natur. Hist. Special Publ. No. 9:171-177. Martin, L. D., and J. D. Stewart. 1982. An ichthyornithiform bird from the Campanian of Canada. Canadian J. Earth Sci. 19:324-327. Martin, L. D., J. D. Stewart, and K. N. Whet- stone. 1980. The origin of birds: structure of the tarsus and teeth. Auk 97:86-93. Martin, L. D., and J. Tate, Jr. 1976. The skeleton of Baptornis advenus (Aves: Hcsperornithifor- mes). In: Collected papers in avian paleontology honoring the 90th birthday of Alexander Wet- more. Smiths. Contrib. Paleobiol. No. 27:35-66. Olson, S. L., and A. Feduccia. 1980. Preshyornis and the origin of the Anseriformes (Aves: Cha- radriomorphae). Smiths. Contrib. Zool. No. 323. 24 pp. Olson, S. L., and M. C. McKitrick. 1981. A new genus and species of emberizine finch from Pleistocene cave deposits in Puerto Rico (Aves: Passeriformes). J. Vert. Paleont. 1:279-283. Schmidt, W. J., and A. Keil. 1958. Die gesunden und die erkrankten Zahngewebe des Menschen und der Wirbeltiere im Polarisationsmikroskop. 386 pp. Carl Hanser Verlag, Miinchen. Schmidt, W. J., and A. Keil. 1971. Polarizing microscopy of dental tissues. 603 pp. Pergamon Press, Oxford and New York. Snow, F. H. 1887. On the discovery of a fossil bird track in the Dakota Sandstone. Trans. Kansas Acad. Sci. 10:3-6. Swinton, W. E. 1975. Fossil birds. British Mus. (Nat. Hist.) Publ. 397, 3rd ed., 81 pp. Wetmore, A. 1926. Fossil birds from the Green River deposits of eastern Utah. Ann. Carnegie Mus. Natur. Hist. 16:391-402. Wetmore, A. 1944. Remains of birds from the Rex- road fauna of the upper Pliocene of Kansas. Univ. Kansas Sci. Bull. 30 (pt. 1, no. 9):89-105. Wetmore, A., and H. T Martin. 1930. A fossil crane from the Pliocene of Kansas. Condor 32:62-63. Williston, S. W. 1896. On the dermal covering of Hesperornis. Kansas Univ. Quart. 5:53-54. Williston, S. W. 1898. Birds. Paleontology, Univ. Kansas Geol. Surv., Vol. 4, pp. 43-53. Zeller, D. E. (ed.). 1968. The stratigraphic succes- sion in Kansas. Kansas Geol. Surv. Bull. 189:1-81. 14 MISCELLANEOUS PUBLICATION MUSEUM OF NATURAL HISTORY APPENDIX GENERA AND SPECIES LISTED Agelaius 11 sp 11 Anas 6 acuta 6 bunkeri 6 clypeata 6 sp 6 Aythya 6 sp 6 Baptornis 1.4 advenus 2, 4, 5 Bartramia 9 longicauda 9 Bird Track 12 Burhinus 10 aquilonaris 10 Calamospiza 11 melanocorys 11 Calcarius 11,12 sp 12 Calidris 9 sp 9 Colinus 7 hibbardi 7 Coltonia 9 recurvirostra 9 Colymbus 5 caspicus 5 Cracid 7 Cursoricoccyx 10 geraldinae 10 Eostri.x 10 martinellii 10 Erolia 9 sp 9 Fulica 8 amehcana 8 Gallinula 8 kansarum 8 Grus 8 nannodes L 8 Hargeria (Hesperornis) 4 gracilis 4 Hesperornis 1. 2, 4 gracilis 4 regalis 4 sp 4 Hesperornithid 4 Icteridae 1 1 Ichthyornis 1, 2, 8 sp 8 Irenesauripus 12 Molothrus 11 sp 11 Nettion 6 bunkeri 6 Parahesperornis 2, 4 alexi 4 Pedinorhis 12 stirpsarcana 12 Phorusrhacos 1 Podiceps 5 caspicus 5 Presbyornis 9 Rallus 7 prenticei 7 Uintornis 11 marionae 1 1 Zenaidura 10 macroura 10 prior 10 Harvard MCZ Librai 3 2044 066 317 587 Date Due RECENT MISCELLANEOUS PUBLICATIONS UNIVERSITY OF KANSAS MUSEUM OF NATURAL HISTORY 62. A demographic study of the ringneck snake {Diadophis punctatus) in Kansas. By Henry S. Fitch. Pp. 1-53, 19 figures in text. April 3, 1975. Paper bound. 65. The biology of an equatorial herpetofauna in Amazonian Ecuador. By William E. Duellman. Pp. 1-352, 198 figures in text. August 30, 1978. Paper bound. 66. Leptodactylid frogs of the genus Eleutherodactylus from the Andes of southern Ecuador. By John D. Lynch. Pp. 1-62, 23 figures in text. February 28, 1979. Paper bound. 67. An ecogeographic analysis of the herpetofauna of the Yucatan Peninsula. By Julian C. Lee. Pp. 1-75, 27 plates, 22 figures in text. February 29, 1980. Paper bound. 68. Internal oral features of larvae from eight anuran families: Functional, systematic, evolutionary and ecological considerations. By Richard Wassersug. Pp. 1-146, 37 figures in text. June 24, 1980. Paper bound. 69. The Eleutherodactylus of the Amazonian slopes of the Ecuadorian Andes (Anura: Leptodactylidae). By John D. Lynch and William E. Duellman. Pp. 1-86, 8 figures in text. August 29, 1980. Paper bound. 70. Sexual size differences in reptiles. By Henry S. Fitch. Pp. 1-72, 9 figures in text. February 27, 1981. Paper bound. 71. Late Pleistocene herpetofaunas from Puerto Rico. By Gregory Pregill. Pp. 1-72, 26 figures in text. May 8, 1981. Paper bound. 72. Leptodactylid frogs of the genus Eleutherodactylus in the Andes of northern Ecuador and adjacent Colombia. By John D. Lynch. Pp. 1-46, 22 figures in text. July 8, 1981. Paper bound. 73. Type and figured specimens of fossil vertebrates in the collection of the University of Kansas Museum of Natural History. Part L Fossil fishes. By H.-P. Schultze, J. D. Stewart, A. M. Neuner and R. W. Coldiron. Pp. 1-53. October 6, 1982. Paper bound. 74. Relationships of pocket gophers of the genus Geomys from the central and northern Great Plains. By Lawrence R. Heaney and Robert M. Timm. Pp. 1-59, 19 figures in text. June 1, 1983. Paper bound. 75. The taxonomy and phylogenetic relationships of the hylid frog genus Stefania. By William E. Duellman and Marinus S. Hoogmoed. Pp. 1-39, 30 figures in text. March 1, 1984. Paper bound. 76. Variation in clutch and litter size in New World reptiles. By Henry S. Fitch. Pp. 1-76, 15 figures in text. May 24, 1985. Paper bound. 77. Type and figured specimens of fossil vertebrates in the collection of the University of Kansas Museum of Natural History. Part IL Fossil amphibians and reptiles. By H.-P. Schultze, L. Hunt, J. Chorn and A. M. Neuner. Pp. 1-66. December 3, 1985. Paper bound.