UNIVERSITY OF ILLINOIS LIBRARY at URBANA-CHAMPAIGN M BIOLOGY APR 91992 Zoology NEW SERIES, NO. 53 World Catalog of the Genera of Pselaphidae (Coleoptera) Alfred F. Newton, Jr. Donald S. Chandler vo\ April 28, 1989 Publication 1400 JUHU19&9 PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY Information for Contributors to Fieldiana General: Fieldiana is primarily a journal for Field Museum staff members and research associates, although manuscripts from nonaffiliated authors may be considered as space permits. The Journal carries a page charge of $65 per printed page or fraction thereof. Contributions from staff, research associates, and invited authors will be considered for publication regardless of ability to pay page charges, but the full charge is mandatory for nonaffiliated authors of unsolicited manuscripts. Payment of at least 50% of page charges qualifies a paper for expedited processing, which reduces the publication time. 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Journal of Ecology, 51: 567-601. Langdon, E. J. M. 1979. Yage among the Siona: Cultural patterns in visions, pp. 63-80. In Browman, D. L., and R. A. Schwarz, eds., Spirits, Shamans, and Stars. Mouton Publishers. The Hague, Netherlands. Murra, J. 1946. The historic tribes of Ecuador, pp. 785-821. In Steward, J. H., ed., Handbook of South American Indians. Vol. 2, The Andean Civilizations. Bulletin 143, Bureau of American Ethnology, Smithsonian Institution, Washington, D.C. Stolze. R. G. 1981. Ferns and fern allies of Guatemala. Part II. Polypodiaceae. Fieldiana: Botanv, n.s., 6: 1- 522. Illustrations: Illustrations are referred to in the text as "figures" (not as "plates"). Figures must be accompanied by some indication of scale, normally a reference bar. Statements in figure captions alone, such as "x 0.8." are not acceptable. Captions should be typed double-spaced and consecutively. See recent issues of Fieldiana for details of style. 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Each author will normally receive a copy of the edited manuscript on which deletions, additions, and changes can be made and queries answered. Only one set of page proofs will be sent. All desired corrections of type must be made on the single set of page proofs. Changes in page proofs (as opposed to corrections) are very expensive. Author-generated changes in page proofs can only be made if the author agrees in advance to pay for them. THIS PUBLICATION IS PRINTED ON ACID-FREE PAPER. FIELDIANA Zoology NEW SERIES, NO. 53 World Catalog of the Genera of Pselaphidae (Coleoptera) Alfred F. Newton, Jr. Division of Insects Field Museum of Natural History Chicago, Illinois 60605-2496 Donald S. Chandler Department of Entomology University of New Hampshire Durham, New Hampshire 03824 Accepted May 1, 1987 April 28, 1989 Publication 1400 PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY © 1989 Field Museum of Natural History Library of Congress Catalog Card Number: 88-82531 ISSN 0015-0754 PRINTED IN THE UNITED STATES OF AMERICA Table of Contents Abstract 1 Introduction 1 Methods 3 Acknowledgments 4 Discussion of Changes and Problems .... 4 Family-Group Name Changes 5 New Names for Newly Discovered Junior Homonyms 5 Newly Discovered Junior Homonyms with Available Names 5 Validation of Selected Nomina Nuda 5 Remaining Nomina Nuda 5 Type Species Designations 6 Miscellaneous Changes 7 Problems Yet to be Resolved 8 Genera Transferred from or to Pselaphidae 8 SUPRAGENERIC CLASSIFICATION 9 List of Family-Group Names 10 Catalog of Genera 13 Abbreviations 14 Recent Taxa 14 Faroninae 14 Bythinoplectini 14 Dimerini 17 Faronini 18 Mayetiini 19 Euplectinae 19 Euplectomorphi 19 Euplectini 19 Pteracmini 27 Raffrayiini 28 Trogastrini 29 Jubomorphi 31 Jubini 31 Batrisinae 31 Amauropsini 31 Batrisini 32 Metopiini 41 Goniacerinae 41 Brachyglutomorphi 41 Arnylliini 41 Brachyglutini 41 Bythinomorphi 47 Bythinini 47 Goniacerini 49 Pygoxyini 50 Tanypleurini 50 Trichonychini 52 Valdini 52 Proteromorphi 52 Imirini 52 Proterini 52 Tychomorphi 53 Speleobamini 53 Tychini 53 Pselaphinae 54 Arhytodimorphi 54 Arhytodini 54 Ctenistomorphi 54 Attapseniini 54 Ceophyllini 54 Chalcoplectini 54 Ctenistini 54 Odontalgini 56 Pachygastrodini 56 Petanopini 56 Schistodactylini 56 Somatipionini 56 Tmesiphorini 56 Tyrini 57 Cyathigerimorphi 61 Barrosellini 61 Cyathigerini 61 Hybocephalini 61 Machadoini 62 Pselaphomorphi 62 Phalepsini 62 Pselaphini 62 Clavigerinae 63 Adranini 63 Apoderigerini 63 Clavigerini 63 Clavigerodini 63 Dimerometopini 66 Disarthricerini 66 Hoplitoxenini 66 Lunillini 67 Mastigerini 67 Miroclavigerini 67 Neoceratopsini 67 Neocerini 67 Radamini 67 Thysdariini 67 Tiracerini 68 Extinct Taxa 68 Taxa Based on Larvae Only 69 Nomen Dubium 69 Literature Cited 69 Taxonomic Index 85 111 List of Illustrations List of Tables 1. Names and boundaries of zoogeographic 1. New names for newly discovered junior regions used in generic catalog 14 homonyms 6 2. Newly discovered junior homonyms with available names 6 3. Earlier names for which no type species have apparently been designated 7 World Catalog of the Genera of Pselaphidae (Coleoptera) Abstract All names that have been proposed above the species level in the family Pselaphidae are cata- loged, with a full bibliography of the original de- scriptions and a full taxonomic index. The ap- proximate world distribution and number of species of each currently recognized genus are giv- en, along with the present status, type species, and mode of type species designation for all genus- group names. Changes and problems at the generic level are summarized in a separate Discussion section. New names are proposed for 18 newly discovered ju- nior homonyms representing currently recognized pselaphid genera; six additional homonyms have available names. Four unavailable names in cur- rent use are validated by designation of a type species for each; 1 3 additional unavailable names, all of Karaman and most recently cited as syn- onyms, are listed but not validated. Type species are designated for the first time for 2 1 older names. Previously unnoticed type species designations make Fustigerodes a senior objective synonym of Commatoceropsis. and Ramecia a senior objective synonym of Liniolis; Novoclaviger is available for Fustigerodes of recent usage, and a new genus Ra- cemia is proposed for Ramecia of recent usage. Rhytus Westwood (not preoccupied) replaces Ar- hytodes Reitter. Problems with previous type species designations for Da/modes, Dahnoplectus, and Fustiger are reviewed. Unresolved problems involving the forgotten name Alytus, previously overlooked type species designations for Batrisus, and the identity of the type species of Rybaxis are discussed. A list of genera recently transferred to or from Pselaphidae is also given. The genera are placed into higher taxa according to the authors' estimate of a currently accepted world classification, based on that of Jeannel. Problems with this classification, including the resolution of conflicting regional classifications, are discussed, but only one new change is made: Aux- enocerini Jeannel is synonymized under Jubini Raffray. Application of the principle of priority to family-group names requires the following changes in recent usage: Bythininae to Goniacerinae. Fus- tigerini to Clavigerodini, Octomicrini to Dimerini, Pyxidicerini to Bythinoplectini, and Zethopsina to Bythinoplectina. Citations for family-group names are given in a separate list which serves as a con- spectus of the suprageneric classification. Since the last world catalog of Recent Psela- phidae (Raffray, 1 9 1 1 ), the family has grown from 3,300 species in 468 genera to about 8,377 species in 1,091 genera, placed in two divisions, six subfamilies, 10 sections, 57 tribes, and 40 sub- tribes. An additional 16 genera and 36 species of extinct taxa, and two genera with two species based on larvae, are listed. Introduction The family Pselaphidae, with about 8,400 de- scribed species, is among the ten largest beetle families, but it is easily the least well known of these families. This is undoubtedly due in large measure to the small size of the beetles (usually 1 to 3 mm) and their reclusive habits. Despite a general scarcity of specimens in most insect col- lections, the group has attracted the attention of several very productive taxonomists, perhaps due to the intriguing variation in external structure which often includes striking cuticular modifica- tions of the males. Use of modern collecting meth- ods (Berlese/Tullgren funnels. Winkler/Moczarski extractors, blacklight and flight-intercept traps) is NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE now providing a wealth of material, and interest in the family is higher than ever among system- atists as well as other biologists. An understanding of the present state of knowledge of the classifi- cation and distribution of the family is greatly hampered, however, by the lack of any worldwide summary since the last catalog of the family (Raf- fray, 1911). The present work attempts to provide such a summary above the species level. Knowledge of the natural history of pselaphids is usually, at best, restricted to the habitat in which the adults are collected. Most species are found in leaf and wood litter of the forest floor (e.g., Park, 1947a; Tanokuchi & Itoh, 1983; Chandler, 1987a), usually in low numbers of individuals. Species richness in this habitat may be very high, however, with 30 or more species often turning up in litter samples from small plots of warm temperate and tropical forests (pers. obs.). Species can also be discovered in wetland, grassland, desert, beach, cave, and even arboreal habitats, wherever debris, moss, or root mats maintain a zone of high hu- midity. A number of species scattered through five of the subfamilies are inquilines in the nests of social insects, while all species of the sixth subfam- ily, the Clavigerinae, are thought to be obligate associates of ants or termites (Kistner, 1982). No complete life histories have yet been worked out. The immature stages are very poorly known, with larvae of fewer than a dozen genera and pupae of three genera described (see especially Kemner, 1927; B6ving& Craighead, 1931; Besuchet, 1952, 1956b; Kasule, 1966; De Marzo & Vit, 1982; De Marzo, 1986; Newton, in press). Both larvae and adults are assumed to be predatory on small ar- thropods and worms, based on limited observa- tions (Park, 1947a; Engelmann, 1956; Reichle, 1969; De Marzo, 1986) and on general mouthpart structure and gut contents. Members of the family are of no known com- mercial importance, but the high species richness and abundance in certain microhabitats suggests that pselaphids may be ecologically important (Park, 1947a; Chandler, 1987a). This diversity, in combination with a wealth of taxonomic charac- ters that should facilitate phylogenetic analysis, also suggests a high potential for use of pselaphids in various evolutionary studies such as local and large-scale biogeography and integration of guests into social insect systems. Pselaphids have been maintained as a distinct family of stable limits since the early part of the 19th century. The family is placed in the super- family Staphylinoidea, adjacent to the even larger family Staphylinidae (32,000+ named species), most members of which resemble pselaphids in having short elytra. Pselaphids differ from most staphylinids in having a reduced tarsal formula (all three- or two-segmented), somewhat clubbed antennae, and a distinctive body form with a rel- atively rigid abdomen. They also have a unique system of distinctive cuticular foveae, highly mod- ified female genitalia, and other adult and larval characters which imply that the family is mono- phyletic. Pselaphids are generally considered to be derived from staphylinids, with the main source of argument being to which subgroup of Staphy- linidae they are most closely related. It is likely that, when a more phylogenetically defensible family classification of Staphylinoidea is adopted, Pselaphidae will be combined with at least part of the present family Staphylinidae, as already done by Naomi (1985). For general discussions on the relationship between these two families and the difficulties of distinguishing them, see Casey ( 1 894), Raffray (1908a), Scheerpeltz (1926), Park (1942), Jeannel (1950), Kasule (1966), Naomi (1985), and Thayer (1987). The anatomy of the Pselaphidae is discussed by Raffray (1908a), Park (1942), and Jeannel (1950). There have been two periods of explosive growth in the number of described species, genera, and higher taxa of Pselaphidae. The first period ex- tended from the late 1 870s to about 1918, and was largely the result of the work of three men: Achille Raffray, Edmund Reitter, and Ludwig Schaufuss. All three produced numerous descriptive taxo- nomic papers dealing primarily with faunal col- lections from various parts of the world, and each also produced one or more attempts at proposing a higher classification of the family. Two large synthetic works by Raffray on the world fauna of the family, in the Genera Insectorum (1908a) and Coleopterorum Catalogus (1911) series, summa- rize the results of this first active period. The fam- ily grew from 450 species in 44 genera (Gemmin- ger & Harold, 1868) prior to this period, to 3,300 species in 468 genera and 1 9 tribes (Raffray, 1911). The second very active period ran from about 1942 to 1964 and was dominated by two men: Orlando Park and the remarkably prolific Rene Jeannel. This period is also characterized primar- ily by the descriptive taxonomy of faunas, with each author confining his work to one faunal re- gion for long periods of time. Park worked initially on the Neotropical fauna, and in his later years on the Nearctic and portions of the Pacific faunas. Jeannel initially worked on the western European FIELDIANA: ZOOLOGY fauna, following with extensive work on all por- tions of the African fauna. Later in his career he touched on the Japanese, Chilean, and to a limited extent the Oriental faunas. The major workers following this second period have generally continued a regional approach but have increasingly emphasized thorough revision- ary studies of more restricted taxonomic groups. These include C. Besuchet (Palearctic), N. Leleup (African), I. Lobl (Oriental), and R. O. Schuster and A. A. Grigarick (Nearctic). G. Coulon (1986) has completed the first worldwide revision of the genera and higher taxa of a sizeable group (Faron- inae excluding Faronini). Grigarick and Schuster (1980) and Coulon (1986) have presented the first explicit phylogenetic analyses of certain groups in the family. Although the number of species, genera, and higher taxa of pselaphids has more than doubled since Raffray's (1911) catalog, no attempt has been made previously to summarize this growth on a worldwide basis. Such a summary is made partic- ularly necessary, but difficult, by the regional ap- proach taken by most specialists on the family. At the present time, regional diagnoses of faunas have been produced for France (Jeannel, 1950), central Europe (Besuchet, 1974a), northern Africa (Jean- nel, 1956a), central Africa (Jeannel, 1959d), South Africa (Jeannel, 1964c), Madagascar (Jeannel, 1 960b), Sri Lanka (Jeannel, 1961), Japan (Jeannel, 1958b), the Neotropical region (Park, 1942, 1952a- b), and Chile (Jeannel, 1962). To a certain extent, new generic and even higher-level concepts have been developed independently for these better- studied parts of the New and Old World faunas, while Southeast Asia and the Australian and Pa- cific regions have been largely neglected since the earlier active period. The primary purpose of this catalog is to sum- marize the present classification of Pselaphidae at the generic and higher levels. Only changes re- quired by the integration of somewhat conflicting regional treatments are made here. We have also attempted to resolve a number of nomenclatural matters including the determination of type species designations and priority of family-group names. Finally, we include an estimate of the known dis- tribution and number of described species of each genus. We do not necessarily endorse this classi- fication or intend to stabilize it with the publica- tion of this catalog. Rather, we hope this summary will facilitate and perhaps stimulate broad-based studies on the classification, phylogeny, and bio- geography of this interesting and diverse family. Methods This catalog was undertaken in part to deter- mine the feasibility of using a microcomputer da- tabase for constructing generic checklists for the families of Staphylinoidea more easily than could be done on paper or with a word processor. The basic procedure was to ( 1 ) set up a suitable data- base (a commercially available database program, dBASE II®, was used on Morrow® and Norths- tar® CP/M® microcomputers); (2) begin with the last world catalog of the family (Raffray, 1911), entering all generic names and relevant data in- cluding distribution and number of species for each recognized genus; and (3) go through the Zoolog- ical Record in sequence from 191 1 to 1986, adding new generic names, changing synonymies, and modifying the placement, distribution, and species number of each genus as needed. A separate da- tabase was set up, using the same procedures, for the family-group names. The Literature Cited section was set up as a third database. The full article titles, as well as journal or book titles, were entered without shortening or abbreviations. Care was taken to establish as spe- cifically as possible the correct publication date of all citations, principally by searching for a precise date on the flyleaf of books, on journal issue cov- ers, or at the top or bottom of pages. Accurate publishing dates for several papers could not be established, and in these cases the "year" of the volume is assumed to be correct. The sequence of articles by an author in the Literature Cited section is based as closely as possible on the actual pub- lication dates. All of the descriptive taxonomic literature on genera and higher taxa was examined to check dates, original spellings, page citations, and type species designations, if any. We have made no attempt to list all misspellings, lapsus calami, or printer's errors, but have cited those that are used consistently in one or more significant works on Pselaphidae and might therefore be confusing. With a few significant exceptions, misidentifications are not cited. Each type species is cited in its original combination, with the present senior synonym (if any) in parentheses in its own original combina- tion. Standard nomenclators such as the Nomenclator Zoologicus (Neave, 1939, 1940, 1950; Edwards & Hopwood, 1966; Edwards & Vevers, 1975), the Nomenclator Animalium Generum et Subgenerum (Schulze et al., 1926-1954), and the Zoological Record through 1 986 were checked for cases of NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE junior homonymy of names used in Pselaphidae. Following Blackwelder's (1952) guidelines, we sought subsequent designations of type species in a number of general works known to provide many such designations, including Latreille (1810), Westwood (1838), Shuckard (1839), Thomson (1859), and Lucas (1920). We followed Black- welder's (1952) interpretation of Lucas's (1920, p. v) introductory comments, and accepted a single species name or the first of two or more names listed for a genus as a designation of that species as type species for the genus, unless there was a query, or the first two names were connected by "et". Helpful works for locating the earliest use of family-group names included Lucas (1920), Han- dlirsch (1925), Watt (1975), and Spahr (1981), in addition to the catalogs of Raffray (1908a, 1911) and other major works on Pselaphidae. The catalog was cross-checked and augmented in several ways. Reports from the databases sorted by various criteria such as author- year-page, name, or taxonomic placement were used as a means of error-checking. Corrected and appropriately for- matted reports were directly incorporated into the final manuscript, which was edited and indexed with a word processor (WordPerfect® on a Tan- dy® MS-DOS® microcomputer). Because the pro- cedure for constructing the catalog subsequent to Raffray (1911) did not include keeping track of individual species, and because the Zoological Re- cord often has not specified the fate of species in genera whose limits have been modified, the dis- tributions and species numbers for large and fre- quently modified genera may be inaccurate. To help correct these and any other errors, a draft of the catalog was sent to most currently active pse- laphid specialists for checking and comments (see Acknowledgments). Virtually all of their suggested corrections and changes were adopted. We have attempted to follow the current Inter- national Code of Zoological Nomenclature (ICZN, 1985) in all nomenclatural matters, including the priority of family-group names, with the three ex- ceptions noted below under Problems Yet to be Resolved. The content and format of the catalog conforms in general to unpublished guidelines being developed by the Cataloging Committee of the Systematic Entomology Laboratory, United States Department of Agriculture, Washington, D.C. (R. W. Hodges, Chairman). The senior author was primarily responsible for constructing the databases, entering the data (ex- cept type species and bibliography), checking for homonyms, and producing the final output; the junior author, for producing the bibliography, checking type species designations, and placing taxa of uncertain position. We have both been involved to some extent in all these activities and bear joint responsibility for any errors or omissions. We intend to keep active and up-to-date copies of the databases on which this catalog is based on disk, and would therefore appreciate being in- formed of any corrections or additions. We will consider publishing a revised version if and when this seems warranted. Acknowledgments We thank several individuals for sending copies of papers that were not available at the Harvard University libraries: R. Baranowski, Lund; E. C. Becker, Ottawa; C. Besuchet, Geneva; E. R. Hoe- beke, Ithaca; Z. Kaszab, Budapest; I. Lobl, Ge- neva; S. Miller, Cambridge; and K. Sawada, Koy- oen (Japan). F. M. Carpenter and S. Regen of Harvard University were very helpful with certain obscure references. We also thank C. P. Ciali, Ex- tension Consultant, and C. A. Penniman, Bio- metrics, both University of New Hampshire, for their invaluable help in assisting with transfer of files between two disparate microcomputers. Last- ly, we extend special thanks to all those who agreed to review the manuscript for their constructive comments and corrections: C. Besuchet, Museum d'Histoire Naturelle, Geneva; J. F. Burger, Uni- versity of New Hampshire, Durham; G. Coulon, Institut Royal des Sciences Naturelles, Brussels; N. Leleup, Musee Royal de l'Afrique Centrale, Tervuren; I. Lobl, Museum d'Histoire Naturelle, Geneva; R. M. Reeves, University of New Hamp- shire, Durham; R. O. Schuster, University of Cal- ifornia at Davis; M. K. Thayer, Field Museum of Natural History, Chicago; and J. A. Wagner, Ken- dall College, Evanston. Dr. Besuchet's comments were particularly extensive, and he has prepared a publication (Besuchet, 1 986) for the express pur- pose of establishing numerous synonymies and other changes for inclusion in this catalog. Com- ments from two anonymous reviewers also sig- nificantly improved the manuscript. Discussion of Changes and Problems In the course of preparing this catalog, we have made a number of discoveries that require changes FIELDIANA: ZOOLOGY in the application of certain names at both the family-group and genus-group levels. Some of these discoveries have been independently made and adopted by G. Coulon in his doctoral thesis (Cou- lon, 1 986), parts of which are to be published soon. [Note: the thesis itself is not considered a publi- cation, so our checklist does not include the many new genera and generic synonymies proposed by Coulon; we have, however, followed his tribal placement of existing genera.] Besuchet ( 1 986) has also recognized and corrected some of the prob- lems. The changes are summarized here. Family-Group Name Changes Articerini of Schaufuss, 1872, and Jeannel, 1954, is based on a misidentified type genus, Articerus of Hope, 1845 (not Dalman, 1826). Besuchet (1986) has reviewed this problem and corrected it by es- tablishing a new genus, Tiracerus, for Articerus of Hope, and the new tribe Tiracerini for Articerini of authors. Auxenocerini Jeannel, 1962 = Jubini Raffray, 1904, NEW SYNONYMY. The Chilean tribe Auxenocerini clearly represents the same taxon as the Neotropical tribe Jubini, which Jeannel ap- parently overlooked. The two tribes are here com- bined and Auxenocerini synonymized under Ju- bini (see also Coulon, 1986). Bythinoplectina Schaufuss, 1890, has priority over Zethopsina Jeannel, 1952. Bythinoplectini Schaufuss, 1890, has priority over Pyxidicerini Raffray, 1904. Clavigerodini Schaufuss, 1 882, has priority over Fustigerini Jeannel, 1949. Dimerini Raffray, 1908, has priority over Oc- tomicrini Jeannel, 1952. The latter name was adopted by Jeannel ( 1 956e) when he synonymized Dimerus under Octomicrus, but in our view Oc- tomicrini has not become sufficiently established to justify suppression of Dimerini under the pro- visions of Article 40b of the Zoological Code (ICZN, 1985; see also Coulon, 1986). Goniacerinae Reitter, 1882 (1872), has priority over Bythininae Raffray, 1890. This subfamily, when first recognized in its present sense by Jean- nel (1955b), was called Bryaxinae, a name based on the junior homonym Bryaxis Leach (now Re- jected Name No. 1953). Later, Jeannel (1959d) correctly rejected the name Bryaxinae, replacing it with Bythininae under the mistaken understand- ing that this name was as old as any other available name. Goniacerides Reitter actually takes prece- dence from 1872, since it was apparently intended to replace Goniastini Schaufuss, 1872, when Gon- iastes was synonymized (temporarily) under Gon- iacerus. New Names for Newly Discovered Junior Homonyms We have found 24 cases of previously unrec- ognized junior homonyms in Pselaphidae. Eigh- teen of these names are currently in use as the valid names for genera and have no available syn- onyms to replace them, so we propose new names for these here (table 1). All of the new names are whole or reduced anagrams of the names they re- place, and each (except Pedisinops, masculine) has the same gender as the replaced name. Newly Discovered Junior Homonyms with Available Names Six junior homonyms are currently treated as subjective synonyms of available generic names, so we do not propose new names for them (table 2). Validation of Selected Nomina Nuda A number of names proposed since 1930 meet the requirements of availability except for the mandatory designation of type species (ICZN, 1985, art. 13b). Those names subsequently vali- dated by designation of a type species are listed in the generic catalog, with the original citation of each indicated as a nomen nudum. Four names generally listed as valid names for a genus and three subgenera are validated in the generic catalog by our designation of a type species for each. The new taxa are: Decarfuss NEW SUBGENUS of Decarthron (validation of Park, 1942:194) Paradrasinus NEW SUBGENUS of Drasinus (validation of Park, 1942:182) Raphitreodes NEW GENUS (validation of Jeannel, 1958b: 119) Sinistrocedius NEW SUBGENUS of Cedius (validation of Park, 1949b:336) Remaining Nomina Nuda The following nomina nuda have never been validated by designation of a type species. Because NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE Table 1. New names for newly discovered junior homonyms. New name For Not Anama Apotectus Asanis Batrischema Batrixenus Bythiotes Cerylambus Dalminia Humbertella Hyobontus Merinia Mnia Nondulia Pedisinops Plectoprotus Rhamophorus Stethotaphrus Sufifer Amana Raffray, 1890 Autoplectus Raffray, 1883 Anasis Raffray, 1891 Batrisoschema Jeannel, 1960 Batrisoxenus Dajoz, 1982 Bythonesiotes Jeannel, 1958 A mblycerus Raffray, 1895 Damilina Jeannel, 1 964 Humbertiella Jeannel, 1960 Hybonotus Jeannel, 1953 /mer/na Raffray, 1897 Mina Raffray, 1890 Nodulina Raffray, 1904 Pedinopsis Raffray, 1 890 Protoplectus Leleup, 1974 Harmophorus Schaufuss, 1 886 Taphrostethus Schaufuss, 1882 Fusifer Raffray, 1900 Walker, 1855 Balsamo-Crivelli, 1843 Castelnau, 1868 Reitter, 1884 Leleup, 1971 Jeannel, 1956 Thunberg, 1815 Horny, 1961 Saussure, 1869 Klug, 1803; Dejean, 1834 Ragonot, 1891 Walker, 1858 Rhumbler, 1895 Cotteau, 1863 Raffray, 1898 Motschulsky, 1851 Fieber, 1844 Dendy, 1896 they have been listed most recently as synonyms (Besuchet, 1974a), we do not validate them here: Bajulobythus Karaman, 1954b:86 (as subgenus of Areopagus) Bajulobythus Karaman, 1957:171 (as subgenus of Balcanobythus) Balcanella Karaman, 1967:60 (as genus, Brach- yglutini; not Balcanella S. Karaman, 1933) Bythinoides Karaman, 1957:167 (as subgenus of Bythinus) Clavidius Karaman, 1957:168 (as subgenus of Areopagus) Dalmatobythus Karaman, 1957:173 (as subge- nus of Argiobythus) Erichobythus Karaman, 1954b:81 (as subgenus of Areopagus) Erichobythus Karaman, 1957:169 (as subgenus of Bythinites) Jugobythus Karaman, 1953:92 (as subgenus of Areopagus) Mesoplectus Karaman, 1962a:61 (as genus, Eu- plectini) Nodibythus Karaman, 1 957: 1 67 (as subgenus of Bythinus) Orientobythus Karaman, 1 957: 1 68 (as subgenus of Areopagus) Plectophleoides Karaman, 1962b: 146 (as sub- genus of Plectophloeus) The following name has never been described and has been ignored by later authors: Eupsinus Motschulsky, 1856a:7 (as genus, in- cluding two species from Louisiana, U.S.A., that are also no mina nuda) Type Species Designations In addition to the names described since 1930 which have never been validated by designation of a type species (see above), we have found 21 earlier names for which no type species have ap- parently been designated (table 3). We have des- ignated a type species for each of these names in the generic catalog. Table 2. Newly discovered junior homonyms with available names. Available name For Not Sognorus Reitter Globosulus Jeannel Jubus Schaufuss Sogaella Jeannel Octomicrellus Jeannel Gerallus Sharp Agnesia Lokay, 1907 Globulites Jeannel, 1954 Gamba Schaufuss, 1872 Leiochrotina Jeannel, 1959 Leptomicrus Jeannel, 1951 Zosimus Schaufuss, 1886 Koninck, 1883; Michaelsen, 1898 de Haan, 1825 Liais, 1872 Westwood, 1883 Fauvel, 1878 Leach, 1825 FIELDIANA: ZOOLOGY Miscellaneous Changes Dalmodes was originally proposed (Reit- ter, 1 882c, p. 2 10) without species; two new species, D. batrisoides and D. rybaxides, were subsequently added (Reiner, 1883d, pp. 382-383), and D. ry- baxides later fixed as type species of the genus (Lucas, 1920, p. 225). Raffray (1890a, p. 102) cre- ated a new monobasic genus Dalmoplectus for Dalmodes "rybaxoides" (= rybaxides) Reitter, but in his discussion of Dalmodes in the same paper (Raffray, 1 890a, p. 1 30), he referred to his removal of Dalmodes batrisoides Reitter to form Dalmo- plectus1. Raffray (1890a, pp. 102, 130) clearly in- tended to remove only one of these species to Dal- moplectus, and the following facts suggest that he intended to base Dalmoplectus on Dalmodes ba- trisoides rather than D. rybaxides: his generic key (Raffray, 1 890a, p. 96) mentions for Dalmoplectus characters of Dalmodes batrisoides rather than D. rybaxides (such as a trifoveate pronotum; see Reit- ter, 1883d, p. 383); he labels a subsequent figure as Dalmoplectus batrisoides (Raffray, 1 890b, p. 2 1 8, fig. 27), and this figure differs grossly from Reitter's (1885c, p. 339, fig. 29) figure of Dalmodes rybax- ides: and, although he mentioned D. rybaxides one later time under the genus Dalmoplectus (Raffray, 1898c, p. 227), all subsequent works of Raffray (e.g., 1904a-b. 1908a, 1911) refer Dalmodes ba- trisoides to Dalmoplectus and D. rybaxides to Dal- modes, without comment. We conclude from this available evidence that Raffray's (1890a, p. 102) mention of Dalmodes rybaxides as the sole species of Dalmoplectus is a lapsus calami for Dalmodes batrisoides, which is mentioned as the sole species of Dalmoplectus on page 1 30 and in most of his later publications. This interpretation preserves the most general usage and placement of both Dal- modes and Dalmoplectus. e.g., by Lucas (1920) and Park (1942). Fustiger was first described by LeConte (1866, p. 108, published in May) for two named species (Articerus brazi liens is West wood and syriacus Saulcy) plus an unnamed species from Tennessee. The latter species was shortly named by Brendel ( 1 866. p. 1 90, published in June) as Fustiger fuch- sii. Lucas (1920, p. 296) and later authors have cited F. fuchsii as type species of Fustiger, and Brendel as author of the genus. LeConte is clearly the author of the genus, and the designation of F. fuchsii (not originally included by name) as type species is not valid. We here designate Articerus braziliensis Westwood as type species of Fustiger, leaving the current concept of the genus un- changed. Table 3. Earlier names for which no type species have apparently been designated. Abryxis Raffray Anabaxis Raffray Anasopsis Raffray Atherocolpus Raffray Atychodea Reitter Batrisomorpha Raffray Brabaxis Raffray Ctenicellus Raffray Ctenisophus Raffray Dacnolitlus Raffray Gamba Schaufuss Glapharsenus Raffray Hypochraeus Raffray Maya Blattny Neophamisus Reitter Nisa Casey Plagiophorus Motschulsky Stenosagola Broun Tychotyrus Broun Vidamus Raffray Zosimus Schaufuss Fustigerodes Reitter was described (Reitter, 1884a) for an unnamed species and Fustiger mad- agascariensis Raffray, which became the type species by monotypy. Raffray (1890a, p. 167) as- signed a name to the unnamed species of Reitter (1884a) and considered this to be the only species in the genus Fustigerodes; he based a new genus, Commatoceropsis, on Fustiger madagascariensis Raffray. Raffray's concepts of Fustigerodes and Commatoceropsis have been accepted by subse- quent authors. The original type fixation of Fus- tigerodes by Reitter results in Fustigerodes being a senior objective synonym of Commatoceropsis, replacing the latter name. The name Novoclaviger Wasmann is available for Fustigerodes of Raffray and subsequent authors (not Reitter). Ramecia Casey (1894, p. 450) was restricted in concept by Grigarick and Schuster (1980), who recognized the genus in the sense established by Bowman's ( 1 934, p. 1 44) designation of Euplectus arcuatus LeConte as type species. The previously overlooked prior fixation of Euplectus crinitus Brendel as type species of Ramecia (Lucas, 1 920, p. 564) is unfortunate, since this species was re- moved from Ramecia by Grigarick and Schuster (1980, p. 34) to form their new monobasic genus Liniolis. The name Ramecia Casey is thus a senior objective synonym of Liniolis, and Ramecia of Grigarick and Schuster ( 1 980, p. 1 8) is left without a name since there are no available synonyms. We therefore propose the new name Racemia NEW GENUS (type species: Euplectus arcuatus Le- Conte, designated here) for Ramecia of Grigarick and Schuster (1980). Rhytus Westwood was replaced by Arhytodes Reitter ( 1 882c) in the belief that Westwood's name was preoccupied by Rytus King. According to the current rules of homonymy (ICZN, 1 985, art. 56b), the single-letter difference between Rytus and Rhy- tus is sufficient to prevent homonymy between NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE them. Thus Rhytus is not preoccupied and should be used instead of Arhvtodes. Problems Yet to be Resolved Alytus and Batrisodes— Hampe (1863), on describing a new species Trichonyx adnexus, pro- posed a new subgeneric name Alytus for the species. This species is now included in the genus Batri- sodes Reitter (1882a). The name Alytus Hampe has been virtually forgotten in the pselaphid lit- erature and overlooked by standard zoological no- menclators. A junior homonym, Alytus Jacoby, 1887, is the currently accepted name for a genus of Chrysomelidae. Dr. C. Besuchet (pers. comm.) submitted a proposal for suppression of Alytus Hampe to the International Commission on Zoo- logical Nomenclature in August 1976, to conserve existing usage of both Batrisodes and Alytus Ja- coby. Apparently no formal action has been taken by the Commission on this proposal. We continue existing usage of Batrisodes here (see also next paragraphs). Batrisus and Batrisodes— Batrisus was pro- posed by Aube (1833, p. 45) to contain eight species, with no type species being designated. Subsequently, Westwood (1838, p. 21) designated an originally included species, Bryaxis nigriventris Denny (junior subjective synonym of Pselaphus venustus Reichenbach, synonymized by Aube, 1833) as type species. Shuckard(1839, p. 145) and Thomson (1859, p. 53) also cited this species (as Pselaphus venustus) as type species of Batrisus. Reitter (1882a, p. 134; 1882c, p. 205) described Batrisodes as a subgenus of Batrisus, initially with- out included species. Five species were subse- quently added to this subgenus (Reitter, 1882d), including B. venustus, but without designation of a type species; the subgenus Batrisus was restricted to B. formicarius Aube. These subgenera were soon elevated to generic rank and have been used in Reiner's sense by all subsequent workers. Lucas (1920, p. 135) fixed the type species of Batrisodes as Batrisus delaporti Aube, while Lucas (1920, p. 136) and all later authors have cited Batrisus for- micarius as type species of Batrisus. A century of Holarctic literature on the 160 species now included in Batrisodes and Batrisus is based on the current type species concepts of these two genera. Recognition of Westwood's fixation of Bryaxis nigriventris as type species of Batrisus would make Batrisus a senior subjective synonym of Batrisodes, replacing the latter name, and would leave Batrisus of Reitter and all later authors with- out a name. It may thus be best to petition the International Commission on Zoological Nomen- clature to set aside previous type species desig- nations for Batrisus and designate Batrisus for- micarius Aube as type species, which would conserve existing usage of both Batrisus and Ba- trisodes. Existing usage of each genus is followed here. Rybaxis Saulcy— Problems with the identifi- cation of the type species of this genus, Bryaxis sanguinea (attributed to various authors), have been reviewed by Besuchet (1969) in a proposal to the International Commission on Zoological Nomenclature. Apparently, no formal action has been taken on this proposal, although Mayr (1971) has published a brief comment. It should be noted that the designation of "sanguinea Leach 1817" by Bowman (1934, p. 144) predates the designa- tion of ' " sanguinea Fabricius" by Jeannel (1950, p. 280) as type species of Rybaxis, but this does not alter the confusing situation. Existing usage of Rybaxis is followed here. Genera Transferred from or to Pselaphidae The family limits of Pselaphidae have been quite stable for over a century and a half. However, the close relationship and similar habitus of the family to Staphylinidae has resulted in some confusion on the placement of a few genera. The presently accepted placement of these genera is given here. Genera described as Pselaphidae, now in Sta- phylinidae, subfamily Euaesthetinae (Fauvel, 1 902; Puthz, 1974): Edaphus Motschulsky = Microphthartus Blattny = Rhenanus Wiisthoff = Tetrameres Schaufuss = Tetratarsus Schaufuss Tamotus Schaufuss Genus described as Staphylinidae, now in Pse- laphidae (transferred by Park, 1947b, 1951): Mayetia Mulsant & Rey Genus of Pselaphidae sometimes erroneously placed in Staphylinidae (by Bernhauer, 1 908, etc.): Dimerus Fiori (= Octomicrus Schaufuss) FIELDIANA: ZOOLOGY Suprageneric Classification The first formal subdivision of the family was made by Leach (1815) who recognized two groups, Pselaphides and Clavigerides. These two groups, usually treated as subfamilies, were divided into a steadily increasing number of tribes during the first period of active work on the family in the late 19th and early 20th centuries (especially by LeConte, 1861; Schaufuss, 1872, 1890; Reitter, 1882c; Raffray, 1890a, 1904a-b). Raffray (1890a) introduced two informal groups, or divisions, of the tribes of Pselaphinae. By the end of this period (Raffray, 1911), 18 tribes of Pselaphinae had been recognized, while the Clavigerinae remained un- divided. During the second period of rapid growth, the higher classification of the family was extensively modified by Jeannel (1949a), who introduced another informal category between subfamily and tribe, the section. Later, Jeannel (1955b) revised the classification again, expanding the number of subfamilies to six and elevating Raffray's two di- visions to rank above subfamily. The number of tribes grew to over 50 during this period, and sub- tribes were introduced (see especially Jeannel, 1949a, 1950, 1954a, 1955b, 1959d, 1962). Jeannefs system, intended to be worldwide but based primarily on his work on the pselaphid fau- nas of the Old World (especially Africa and Eu- rope), was adopted to some extent by Park (1951) for the New World. However, there was incom- plete recognition and utilization by the two au- thors of the higher taxa proposed by each other, probably due to their work on such disjunct fau- nas. The resulting two classifications are not well integrated, especially in the Euplectinae and Tan- ypleurini, where the definitions of the tribes and subtribes at the world level are inconsistent and appear to be phylogenetically unsound. Grigarick and Schuster (1980) reexamined the North and Central American Euplectinae and proposed yet another arrangement, based on the first attempt at a suprageneric phylogenetic analysis in the fam- ily, but they did not propose a formal classification or deal specifically with the existing ones of Jean- nel and Park. Coulon's (1986) review of the clas- sification of Faroninae provides the only up-to- date worldwide summary for a sizeable group of pselaphids. The classification adopted here is essentially that of Jeannel (1955b), which is the most recent su- prageneric classification that was intended to be complete and worldwide down to the tribal level. We have modified Jeannel's system primarily by incorporating Park's (1951) taxa which were over- looked by Jeannel; adding taxa and making other changes published subsequent to Jeannel (1955b); and attempting to resolve conflicts between dif- fering regional classifications, especially those used by Park and coworkers in the New World and those of Jeannel and other Europeans in the Old World and Chile. Many genera, mostly from the poorly studied Oriental, Australian, and Pacific regions, were originally placed in tribes that have now been divided into several tribes, or subdivid- ed into a system of subtribes. We have placed these genera in the present system of tribes and subtribes primarily based on information given in their orig- inal descriptions. We emphasize that the classification used here is presented as the closest we can come to a "pres- ently accepted" classification that is consistent worldwide. We do not necessarily endorse it. In- deed, our experience in attempting to place taxa of uncertain position suggests that there are prob- lems at three different levels with the present clas- sification. First, several of the published charac- terizations of suprageneric taxa, as well as the identification keys derived from these character- izations, are inaccurate and therefore of dubious value. Second, the higher classification is some- what inflated relative to the tribal and subfamily concepts in use elsewhere in Staphylinoidea, and probably should be rendered more consistent with the other families. Third, no attempt has yet been made to analyze most pselaphid groups phyloge- netically. A thorough cladistic analysis of the higher taxa is badly needed, and would probably lead to very substantial changes in the classification. The following synonymical list of family-group taxa includes all names of which we are aware. Since rules on priority of family-group names have been standardized only in the past quarter century, and most older catalogs pay little if any attention to this subject, there are very likely to be some omissions in this list. Many names validated by Schaufuss (1872, 1890) and Reitter (1882c, 1909) were unknown to or ignored by Raffray (1890a and later) and Jeannel (1949a and later), who in- dependently proposed names for the same taxa. In many cases the type genus is the same and only the author and citation need be changed, but in four cases (see Discussion, above) the prior use of a name based on a different type genus requires changing the established name. In these cases, the NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE more widely used junior name is given in paren- theses after the prior name adopted here. The hierarchy of categories used in Pselaphidae, with the present number of taxa of each rank, may be diagrammed thus: Family Division [2] Subfamily [6] Section [10, in 3 subfamilies] Tribe [57] Subtribe [40] List of Family-Group Names In this list, family-group names we accept here are boldfaced and in upper-case letters. Those of the four higher categories are labeled and flush left, those of tribes, indented once, and of subtribes, indented twice under the appropriate tribe. The subfamilies are listed in the generally accepted se- quence of Jeannel (1955b) (presumed to be from phylogenetically more primitive to more derived), but sections, tribes, and subtribes are each listed in alphabetical order within the next higher cat- egory. Citations are triple-indented and give the orig- inal spellings. The citations given here are for no- menclatural purposes only, in contrast to the fre- quent use of similar citations in the pselaphid literature to indicate taxonomic concepts. Cita- tions are given only for the first valid use of a family-group name based on a given type genus; for independent proposals of names based on the same genus; and for a few subsequent uses of names using different orthographies or suffixes, or when it is not clear whether the subsequent use is based on an earlier name or independently proposed. According to the current code (ICZN, 1985), all family-group names based on a given type genus are coordinate, that is, take the author and date of the first valid name based on that genus re- gardless of rank. The citations for each such group of coordinate names are given here only under the highest-ranked taxon of the group (e.g., under Eu- plectinae, not repeated under Euplectini or Eu- plectina). Each boldfaced name should be given the author and date of the first (oldest) citation of a valid name shown here (i.e., one that is not in- dicated as based on an unavailable generic name). Family PSELAPHIDAE Pselaphii Latreille, 1802:239 Pselaphides Leach, 1815:116 Pselaphidea Leach, 1817:80 [sic] Pselaphidae Fleming, 1821:51 Division BRACHYSCELIA Brachyscelidae Raffray, 1890a:82 (group between subfamily and tribe) Brachyscelia Park, 1942:34 Brachyscelides Jeannel, 1955b:5 (revised status, group above subfamily) Subfamily FARONINAE Faronides Reitter, 1882c: 199 Faronina Sharp, 1887:44 Faronini Raffray, 1890a:84 BYTHINOPLECTINI (= PYXIDICERINI) Bythinoplectini Schaufuss, 1890:plate BYTHINOPLECTINA (= ZETHOPSINA) Zethini Schaufuss, 1890:plate (based on preoccupied generic name) Zethopsina Jeannel, 1952a:51 PYXIDICERINA Pyxidicerini Raffray, 1904a:500 DIMERINI (= OCTOMICRINI) Dimerini Raffray, 1908a:412 (published February 1908) Dimerini Bernhauer, 1908:327 (published March 1908) Octomicrini Jeannel, 1952a:43 FARONINI MAYETIINI Mayetinae Scheerpeltz, 1933:1 139 Mayetini Park, 1947b: 124 Subfamily EUPLECTINAE Euplectini LeConte, 1861:57 Euplectini Reitter, 1882c: 194 Euplectini Raffray, 1890a:91 Euplectidae Schaufuss, 1890:plate Section EUPLECTOMORPHI [of EUPLECTIN- AE] Euplectomorphi Jeannel, 1 949a:44 EUPLECTINI ACETALIINA Acetaliini Jeannel, 1958b:81 BIBLOPLECTINA Bibloplectina Jeannel, 1959d:110 BIBLOPORELLINA Bibloporellina Jeannel, 1952a:92 BIBLOPORINA Bibloporini Park, 1951:64 CHRESTOMERINA Chrestomerina Jeannel, 1962:344 10 FIELDIANA: ZOOLOGY EUPLECTINA PANAPHANTINA Panaphantina Jeannel, 1950:76 RHINOSCEPSINA Rhinoscepsii Bowman, 1934:8 TRIMIINA Trimii Bowman, 1934:8 Trimiina Jeannel, 1950:139 TRIMIODYTINA Trimiodytina Jeannel, 1964c:39 PTERACMINI Pteracmini Jeannel, 1962:347 RAFFRAYIINI Raffrayina Jeannel, 1949a:76 Ranavalini Jeannel, 1954a: 183 TROGASTRINI Trogastrini Jeannel, 1949a:75 MITRAMETOPINA Mitracephalini Park, 1951:64 (based on preoccupied generic name) Mitrametopina Park, 1952c:87 (new name for Mitracephalini Park) PHTEGNOMINA Phtegnomini Park, 1951:64 RHEXIINA Rhexini Park, 1951:63 TRISIGNINA Trisignina Park & Schuster, 1955:1 TROGASTRINA Section JUBOMORPHI [of EUPLECTINAE] Jubomorphi Park, 1951:59 JUBINI (= AUXENOCERINI) Jubini Raffray, 1904a:507 Jubinini Raffray, 1908a:25 Auxenocerini Jeannel, 1962:319 (NEW SYNONYMY) Subfamily BATRISINAE Batrisini Reitter, 1882c: 187 Batrisini Raffray, 1890a: 108 Batrisomorphi Jeannel, 1949a: 113 AMAUROPSINI Amauropsini Jeannel, 1948a:l BATRISINI AMBICOCERINA Ambicocerina Leleup, 1970a:309 BATRISINA Oropygiina Jeannel, 1949a: 114 Trabisina Jeannel, 1 949a: 1 1 6 LEUPELIINA Leupeliina Jeannel, 1954e:106 STILIPALPINA Stilipalpina Jeannel, 1954e: 118 METOPIINI Metopiini Raffray, 1904b: 106 Metopiasini Jeannel, 1949a:42 Subfamily GONIACERINAE (= BRYAXINAE, BYTHININAE) Goniastini Schaufuss, 1872:245 (rejected, ICZN, 1985, art. 40b) Goniacerides Reitter, 1882c: 188 Goniacerina Sharp, 1887:21 Goniacerini Raffray, 1 890a: 1 3 1 Section BRACHYGLUTOMORPHI [of GONI- ACERINAE] Bryaximorphi Jeannel, 1949a:86 (based on rejected generic name) Brachyglutomorphi Jeannel, 1959d:510 (new name for Bryaximorphi Jeannel) ARNYLLIINI Arnylliini Jeannel, 1952b:100 BRACHYGLUTINI (= BRYAXINI) Bryaxes LeConte, 1861:57 (based on re- jected generic name) Bryaxini Reitter, 1882c: 188 (based on re- jected generic name) Bryaxini Raffray, 1890a:l 17 (based on re- jected generic name) Brachyglutini Raffray, 1904b: 108 (new name for Bryaxini of authors) BARADINA Baradiini Park, 1951:62 BRACHYGLUTINA Reichenbachiina Jakobson, 1910:577 DECARTHRONINA Decarthronini Park, 1951:61 EUPSENIINA Eupseniini Park, 1951:61 HALORABYXINA Halorabyxina Leleup, 1969a: 138 PSELAPTINA Pselaptina Park, 1976:48 Section BYTHINOMORPHI [of GONIACERI- NAE] Bythinomorphi Jeannel, 1949a:79 BYTHININI Bythinini Raffray, 1 890a: 1 26 BYTHININA MACHAERITINA Machaeritina Jeannel, 1950:168 XENOBYTHINA Xenobythina Jeannel, 1950:201 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 11 GONIACERINI Listriophorini Schaufuss, 1890:plate Simini Schaufuss, 1890:plate (based on preoccupied generic name) PYGOXYINI Pygoxyini Reitter, 1909:202 TANYPLEURINI Tanypleurini Jeannel, 1949a:79 DALMODINA Dalmodiini Park, 1951:61 GLOBINA Globina Jeannel, 1959d:471 INIOCYPHINA Iniocyphini Park, 1951:60 TANYPLEURINA TRICHONYCHINI Trichonyides Reitter, 1882c: 198 Trichonycina Sharp, 1887:40 Trichonyni Raffray, 1890a: 102 Trichonychini Ganglbauer, 1895:798 VALDINI Valdiini Park, 1953a:261 Section PROTEROMORPHI [of GONIACERI- NAE] Proteromorphi Jeannel, 1959d:492 IMIRINI Mirini Raffray, 1917:1 10 (based on preoc- cupied generic name) Imirini Jeannel, 1949a:41 (new name for Mirini Raffray) PROTERINI Proterini Jeannel, 1949a:41 Section TYCHOMORPHI [of GONIACERI- NAE] Tychomorphi Jeannel, 1959d:461 SPELEOBAMINI Speleobamini Park, 1951:51 TYCHINI Tychini Raffray, 1904b:254 Division MACROSCELIA Macroscelidae Raffray, 1890a:83 (group between subfamily and tribe) Macroscelia Park, 1942:34 Macroscelides Jeannel, 1955b:6 (revised status, group above subfamily) Subfamily PSELAPHINAE Section ARHYTODIMORPHI [of PSELAPHIN- AE] Arytodimorphi Jeannel, 1955b:9 ARHYTODINI Arhytodini Raffray, 1890a: 162 Holozodini Raffray, 1900a:518 Section CTENISTOMORPHI [of PSELAPHIN- AE] Ctenistomorphi Jeannel, 1949a: 177 ATTAPSENIINI Attapseniini Bruch, 1933:26 CEOPHYLLINI Ceophyllini Park, 1951:67 CHALCOPLECTINI Chalcoplectini Oke, 1925:13 CTENISTINI Ctenistini Reitter, 1882c: 183 Chenniides Reitter, 1882c: 184 Ctenistini Raffray, 1 890a: 1 40 Chenniina Jakobson, 1910:585 ODONTALGINI Odontalgini Jeannel, 1949a: 177 PACHYGASTRODINI Pachygastrodini Leleup, 1969b:282 PETANOPINI Petanopini Jeannel, 1954c:102 SCHISTODACTYLINI Schistodactylini Raffray, 1890a: 162 SOMATIPIONINI Somatipionini Jeannel, 1949a:208 TMESIPHORINI Tmesiphorini Jeannel, 1949a:202 TYRINI Tyrides Reitter, 1882c: 185 Tyrina Sharp, 1887:3 Tyrini Raffray, 1890a: 144 CENTROPHTHALMINA Centrophthalmina Jeannel, 1949a:209 HAMOTINA Hamotini Park, 1951:67 JANUSCULINA Janusculina Cerruti, 1970:123 TYRINA Section CYATHIGERIMORPHI [of PSELA- PHINAE] Cyathigerimorphi Jeannel, 1951c: 100 BARROSELLINI Barrosellini Leleup, 1973a:81 CYATHIGERINI Cyathigerini Schaufuss, 1872:245 Cyathigerini Raffray, 1890a: 133 HYBOCEPHALINI Hybocephalini Raffray, 1890a: 134 Mestogastrina Jakobson, 1910:585 12 FIELDIANA: ZOOLOGY MACHADOINI Machadoini Jeannel, 1951c: 105 Section PSELAPHOMORPHI [of PSELAPHIN- AE] Pselaphomorphi Jeannel, 1949a: 162 PHALEPSINI Phalepsini Jeannel, 1949a:208 PSELAPHINI Subfamily CLAVIGERINAE Clavigerides Leach, 1815:117 Clavigeridae Hope. 1836:139 ADRANINI Adranini Schaufuss, 1872:245 Adraniini Park, 1951:58 APODERIGERINI Apoderigerini Jeannel, 1954a:310 CLAVIGERINI CLAVIGERODINI (= FUSTIGERINI) Clavigerodini Schaufuss, 1882a:205 Commatocerini Schaufuss, 1882b:349 Clavigeropsini Schaufuss, 1890:plate Fustigerini Jeannel, 1949a:31 DIMEROMETOPINI Dimerometopini Celis, 1970:244 DISARTHRICERINI Diarthricerini Jeannel, 1949a:29 [sic] HOPLITOXENINI Hoplitoxenini Celis, 1969b:418 LUNILLINI Lunillini Celis, 1969b:416 MASTIGERINI Mastigerini Jeannel, 1954a:291 MIROCLAVIGERINI Miroclavigerini Jeannel, 1949a:29 NEOCERATOPSINI Neoceratopsini Celis, 1970:260 NEOCERINI Neocerini Jeannel, 1954a:316 NEOCERINA THEOCERINA Theocerini Jeannel, 1954a:314 RADAMINI Radamini Jeannel, 1954a:319 THYSDARIINI Thysdariini Jeannel, 1954a:332 TIRACERINI Articerini of Schaufuss, 1 872:245 (based on misidentified type genus) Articerini of Jeannel, 1954a:291 (based on misidentified type genus) Tiracerini Besuchet, 1986:263 Catalog of Genera All names of the genus-group that have been used in Pselaphidae through 1 986 are listed in this catalog, with their present status and placement given as near as we can determine. Subsequent generic names or changes that have come to our attention up to the time of final editing (March 1988) are also included. Family-group taxa are listed in the sequence giv- en in the previous section, and genera are listed in alphabetical order within the next higher cat- egory. Names currently recognized as subgenera, synonyms, nomina nuda (if subsequently validat- ed), emendations, and misspellings (incorrect orig- inal or subsequent spellings) are all listed together in chronological order under the currently valid generic name, with the status of each name indi- cated if not a synonym. Taxa described from fos- sils, those based on larvae, and nomina dubia are listed separately at the end of the catalog of Recent adults. In cases of multiple publication of the same name for the same taxon by the same author, mul- tiple citations for the name are given in chrono- logical order. The type species of each genus-group name is given in its original combination, with the method of designation indicated. If the type species is cur- rently a junior synonym, the senior synonym is given in parentheses in its own original combi- nation. The distribution and number of species given for each genus are intended to be approximately correct as of the end of 1986, but both may be inaccurate in larger genera whose limits have been changed often. In general the country or countries and/or major islands of known occurrence are giv- en, but for brevity more or less standard terms for larger zoogeographic regions (modified from Dar- lington, 1957, chap. 7) have usually been used when a genus is known from two or more widely separated countries in such a region. These terms and their definitions as used here are (see also fig. 1): Afrotropical (= Ethiopian): Africa south of the Sahara Desert, but excluding the Republic of South Africa, and including the southern half of the Arabian Peninsula and Mada- gascar (use of "Afrotropical" avoids con- fusion of "Ethiopian" with "from Ethio- pia"; see Crosskey & White, 1977). Holarctic: Nearctic plus Palearctic regions. Nearctic: North America north of Mexico. Neotropical: Mexico, Central and South Amer- NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 13 Fig. 1 . Names and boundaries of zoogeographic regions used in generic catalog. Country and/or island names are given for distributions outside of these regions and for restricted distributions within the regions. ica, and the West Indies, but excluding Chile and the southern half of Argentina. Oriental: India, the Himalayan region, southern China and Taiwan to Indonesia, excluding New Guinea. Palearctic: Europe, Asia excluding the Oriental region, and Africa north of the Sahara Des- ert. (Note: The terms Afrotropical and Neotropical as used here differ from conventional usage in the exclusion of the southern temperate region of each as South Africa and Chile/southern Argentina, re- spectively. As is the case for many groups of in- sects, the pselaphid fauna of each of these tem- perate areas differs significantly from that of the adjacent tropical area to the north. Note too that northern Mexico is included in the Neotropical rather than Nearctic region.) Abbreviations E eastern N northern S southern w western c central des. designated orig. des. preocc. subseq. mon. Recent Taxa original designation preoccupied subsequent monotypy Family PSELAPHIDAE Division BRACHYSCELIA Subfamily FARONINAE Tribe BYTHINOPLECTINI Subtribe BYTHINOPLECTINA Acrodimerus Jeannel, 1954d:162 Type species: Acrodimerus alticola Jeannel (orig. des.) Kenya, Tanzania [2 spp.] Afrozethopsus Jeannel, 1959a: 172 Type species: Afrozethopsus cantaloubei Jean- nel (orig. des.) Cameroon [1 sp.] Anomozethus Jeannel, 1951c:25 Type species: Anomozethus caecus Jeannel (orig. des.) Angola, Zaire [2 spp.] Anozethopsis Jeannel, 1956f:6 Type species: Anozethopsis perinetanus Jean- nel (orig. des.) Madagascar [2 spp.] 14 FIELDIANA: ZOOLOGY Apozethopsus Jeannel, 1954a: 159 Type species: Apozethopsus vadoni Jeannel (orig. des.) Madagascar [3 spp.] Archaeozethus Leleup, 1973a:44 Type species: Archaeozethus mirabilis Leleup (orig. des.) Ivory Coast, Zaire [2 spp.] Aulacozethus Leleup, 1977b:693 Type species: Aulacozethus grossepunctatus Leleup (orig. des.) Ghana, Ivory Coast [1 sp.] Basilewskyozethus Leleup, 1977a:76 Type species: Basilewskyozethus antennalis Leleup (orig. des.) Madagascar [1 sp.] Besucheteidos Comellini, 1985:748 Type species: Besucheteidos guai mi Comellini (orig. des.) Neotropical [13 spp.] Bolbozethus Coulon, 1982b: 145 Type species: Bolbozethus boosteni Coulon (orig. des.) Sri Lanka, Vietnam [2 spp.] Bythinoplectoides Comellini, 1985:739 Type species: Bythinoplectoides gigas Com- ellini (orig. des.) Mexico [1 sp.] Bythinoplectus Reitter, 1882c:209; 1883a:37 Type species: Bythinoplectus foxeatus Reitter (subseq. mon. Reitter, 1883a:37) Neotropical, SW Nearctic, Galapagos Is- lands [34 spp.] Cephalozethus Jeannel, 1955c:300 Type species: Cephalozethus celisi Jeannel (orig. des.) Uganda, Zaire [1 sp.] Couloniella Besuchet, 1983:509 Type species: Couloniella mirabilis Besuchet (orig. des.) India [1 sp.] Dichocoryna Comellini, 1985:742 Type species: Dichocoryna longipalpis Com- ellini (orig. des.) Costa Rica [1 sp.] Dichozethinus Jeannel, 1956d:363 Type species: Dichozethinus franzi Jeannel (orig. des.) Zaire [1 sp.] Dimorphozethus Jeannel, 1952a:62 Type species: Dimorphozethus leleupi Jeannel (orig. des.) Burundi, Zaire [6 spp.] Diplomelinus Jeannel, 1957b:27 Type species: Diplomelinus machadoi Jeannel (orig. des.) Angola [1 sp.] Echinozethus Jeannel, 1954f:76 Type species: Zethopsus furcifer Raffray (orig. des.) Sri Lanka [1 sp.] Euplectomorphus Motschulsky, 1863:424 Type species: Euplectomorphus pygmaeus Motschulsky (monotypy) Sri Lanka [1 sp.] Zethinellus Jeannel, 1 949c: 3 1 Type species: Zethopsus nitidulus Reitter (orig. des.) Glyphozethopsis Jeannel, 1 954e: 1 3 Type species: Glyphozethopsis sulcipennis Jeannel (orig. des.) Zaire [1 sp.] Glyphozethus Jeannel, 1953b:48 Type species: Glyphozethus elongatus Jeannel (orig. des.) Zaire [1 sp.] Goniozethus Jeannel, 1959c: 19 Type species: Goniozethus saegeri Jeannel (orig. des.) Zaire [1 sp.] Hendecameros Comellini, 1985:754 Type species: Hendecameros costaricense Comellini (orig. des.) Neotropical [9 spp.] Heptaleptus Jeannel, 1952a:85 Type species: Heptaleptus punctaticeps Jean- nel (orig. des.) Burundi, Zaire [3 spp.] Jeannelia Raffray, 1 9 1 3a: 1 1 Type species: Jeannelia microphthalma Raf- fray (monotypy) Ivory Coast, Kenya [3 spp.] Leleupiozethus Coulon, 1979:395 Type species: Leleupiozethus mahnerti Cou- lon (orig. des.) Kenya [1 sp.] Leptozethodes Jeannel, 1960c:79 Type species: Leptozethodes leleupi Jeannel (orig. des.) Ivory Coast, Zaire [3 spp.] Loebliozethus Coulon, 1983:240 Type species: Loebliozethus cavaticeps Cou- lon (orig. des.) Ivory Coast [2 spp.] Mecynozethus Jeannel, 1952a:75 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 15 Type species: Mecynozethus leleupi Jeannel (orig. des.) Burundi, Ruanda, Zaire [12 spp.] Metazethopsis Jeannel, 1960b:68 Type species: Metazethopsis terricola Jeannel (orig. des.) Madagascar [1 sp.] Microzethinus Jeannel, 1954a: 165 Type species: Microzethinus endogaeus Jean- nel (orig. des.) Madagascar [1 sp.] Microzethopsis Jeannel, 1960b:64 Type species: Microzethopsis curtus Jeannel (orig. des.) Madagascar [2 spp.] Microzethus Jeannel, 1959d:38 Type species: Microzethus laevis Jeannel (orig. des.) Zaire [1 sp.] Neozethopsus Jeannel, 1954f:77 Type species: Pyxidicerus triophthalmus Schaufuss (orig. des.) E Oriental, New Guinea [6 spp.] Nesiotozethus Jeannel, 1 954a: 1 64 Type species: Nesiotozethus dimorphus Jean- nel (orig. des.) Madagascar [3 spp.] Octomeros Comellini, 1985:714 Type species: Octomeros sulcatum Comellini (orig. des.) Costa Rica, Brazil [2 spp.] Octozethodes Jeannel, 1953b:59 Type species: Octozethodes angulifrons Jean- nel (orig. des.) Zaire [1 sp.] Octozethus Jeannel, 195 lc:23 Type species: Octozethus angolensis Jeannel (orig. des.) Afrotropical [22 spp.] Octozethinus Jeannel, 1952a:69 Type species: Octozethinus urundianus Jean- nel (orig. des.) Opisthosphaera Jeannel, 1952a:67 Type species: Opisthosphaera gracilicornis Jeannel (orig. des.) Burundi, Zaire [6 spp.] Oxyzethodes Jeannel, 1954e:79 Type species: Oxyzethodes acutipalpus Jean- nel (orig. des.) Zaire [1 sp.] Oxyzethus Jeannel, 1953b:46 Type species: Oxyzethus rostratus Jeannel (orig. des.) Zaire [1 sp.] Pachyzethopsus Jeannel, 1954f:67 Type species: Zethopsus procerus Raffray (orig. des.) Malaya [1 sp.] Parazethopsus Jeannel, 1 952d: 16 Type species: Parazethopsus wittei Jeannel (orig. des.) Afrotropical [2 spp.] Petalozethopsis Jeannel, 1960a:6 Type species: Petalozethopsis andriai Jeannel (orig. des.) Madagascar [1 sp.] Poeciloceras Comellini, 1985:712 Type species: Poeciloceras portentosum Com- ellini (orig. des.) Paraguay [1 sp.] Proboscites Jeannel, 1949c:30 Type species: Proboscites ghesquierei Jeannel (orig. des.) Afrotropical [5 spp.] Protozethopsus Jeannel, 1954a: 162 Type species: Protozethopsus integricornis Jeannel (orig. des.) Madagascar, Ivory Coast [8 spp.] Protozethus Jeannel, 1954f:62 (misspelling) Pyxidion Comellini, 1985:744 Type species: Pyxidion acinaces Comellini (orig. des.) Neotropical [3 spp.] Schizocoryna Comellini, 1985:740 Type species: Schizocoryna wagneri Comel- lini (orig. des.) Panama [1 sp.] Selenozethus Jeannel, 1955a:14 Type species: Selenozethus brevipennis Jean- nel (orig. des.) Uganda [1 sp.] Selinozethus Jeannel, 1954f:65 (nomen nudum, no species included) Trizethopsis Dajoz, 1982:482 Type species: Trizethopsis dent at us Dajoz (orig. des.) Madagascar [1 sp.] Typhloproboscites Jeannel, 1952a:64 Type species: Typhloproboscites leleupi Jean- nel (orig. des.) Afrotropical [4 spp.] Typhlozethus Jeannel, 195 lc:23 Type species: Octozethus machadoi Jeannel (orig. des.) Angola, Zaire [3 spp.] Urozethidius Jeannel, 1959d:91 16 FIELDIANA: ZOOLOGY Type species: Urozethidius pygidialis Jeannel (orig. des.) Ivory Coast, Tanzania [2 spp.] Zethiniotus Jeannel, 1 964a: 1 8 Type species: Zethiniotus spinifrons Jeannel (orig. des.) Zimbabwe [1 sp.] Zethinomorphus Jeannel, 1960b:66 Type species: Zethinomorphus caecus Jeannel (orig. des.) Madagascar [1 sp.] Zethinus Raffray, 1908b:7 Type species: Zethinus severini Raffray (monotypy) Afrotropical [5 spp.] Zethopsimorphus Jeannel, 1954f:73 Type species: Zethopsus crassus Jeannel (orig. des.) Zaire [1 sp.] Zethopsinidius Jeannel, 1954f:79 Type species: Zethopsus decipiens Raffray (orig. des.) Afrotropical [4 spp.] Zethopsinus Jeannel, 1949a:22 Type species: Zethopsus major Raffray (orig. des.) Afrotropical [12 spp.] Zethopsiola Jeannel, 1954f:86 Type species: Zethopsinus urundianus Jean- nel (orig. des.) Afrotropical [63 spp.] Zethopsiolites Jeannel, 1959d:85 Type species: Zethopsiolites fallaciosus Jean- nel (orig. des.) Tanzania [1 sp.] Zethopsoides Jeannel, 1953b:40 Type species: Zethopsoides rugosus Jeannel (orig. des.) Ivory Coast, Zaire [1 sp.] Zethopsus Reitter, 1880b:85 (new name for Ze- thus Schaufuss) Oriental, Japan [21 spp.] Zethus Schaufuss, 1872:246 (preocc. not Fa- bricius, 1805; Pander, 1830) Type species: Zethus opacus Schaufuss (sub- seq. mon. Schaufuss, 1877a: 12) Tribe BYTHINOPLECTINI Subtribe PYXIDICERINA Cerennea Raffray, 1913b: 125 Type species: Cerennea brevipennis Raffray (monotypy) Seychelles [1 sp.] Cerennia Jeannel, 1949a:21 (misspelling) Hughia Raffray, 191 3b: 122 Type species: Hughia phantasma Raffray (des. Lucas, 1920:338) Seychelles [2 spp.] Scottia Jeannel, 1949a:21; 1952a:51 (apparent error for Hughia) Orlandia Comellini, 1985:758 Type species: Orlandia parki Comellini (orig. des.) Mexico [1 sp.] Parapyxidicerus Sawada, 1964:11 Type species: Parapyxidicerus carinatus Sa- wada (orig. des.) Japan [1 sp.] Pyxidicerinus Jeannel, 1952b:72 Type species: Pyxidicerinus barbieri Jeannel (orig. des.) Vietnam [1 sp.] Pyxidicerus Motschulsky, 1863:422 Type species: Pyxidicerus castaneus Mot- schulsky (monotypy) Oriental, New Guinea [9 spp.] Tribe DIMERINI Barroeuplectoides Park, 1 942: 1 0 1 Type species: Barroeuplectoides zeteki Park (orig. des.) Panama [1 sp.] Ocabaraja Schuster & Grigarick, 1980:39 Type species: Ocabaraja romingeri Schuster & Grigarick (orig. des.) Dominican Republic [1 sp.] Ocabajara Schuster & Grigarick, 1980:39 (mis- spelling, first revised here) Ocarabaja Schuster & Grigarick, 1980:39 (mis- spelling, first revised here) Octomicrellus Jeannel, 1 949a: 1 8 Type species: Octomicrus angustatus Raffray (orig. des.) Afrotropical [23 spp.] Leptomicrus Jeannel, 1951c: 13 (preocc, not Fauvel, 1878) Type species: Leptomicrus rostratus Jeannel (orig. des.) Octomicrites Jeannel, 1956e:94 Type species: Octomicrus bifrons Jeannel (orig. des.) Vietnam [1 sp.] Octomicrus Schaufuss, 1877a: 14 Type species: Octomicrus longulus Schaufuss (monotypy) Afrotropical, Oriental, W Palearctic [ 1 8 spp.] Singhala Raffray, 1898c:213, 255 (preocc, not Blanchard, 1851) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 17 Type species: Octomicrus punctipennis Raf- fray (monotypy) Shingala Raffray, 1898c:255 (misspelling, first reviser Raffray, 1904a:577) Dimerus Fiori, 1899:103 (subgenus) Type species: Dimerus staphylinoides Fiori (monotypy) Diastictulus Raffray, 1904a:577 (new name for Singhala Raffray) Phthartomerus Sawada, 1956:102 Type species: Phthartomerus nanus Sawada (orig. des.) Japan [1 sp.] Tuberoplectus Park, 1952c:96 Type species: Tuberoplectus plaumanni Park (orig. des.) Brazil, Mexico [2 spp.] Tribe FARONINI Delenda Croissandeau, 1891:152 Type species: Delenda carthago Croissandeau (monotypy) Turkey [1 sp.] Eusonoma Reitter, 1893:172 Type species: Eusonoma frivaldszkyi Reitter (monotypy) Exeirarthra Broun, 1893a: 1054 Type species: Exeirarthra enigma Broun (monotypy) New Zealand [5 spp.] Faronidiellus Jeannel, 1964c:37 Type species: Faronidius monilis Raffray (orig. des.) South Africa [1 sp.] Faronidius Casey, 1887b:381 Type species: Faronidius africanus Casey (monotypy) South Africa [5 spp.] Faronites Jeannel, 1954a:153 Type species: Faronites vadoni Jeannel (orig. des.) Madagascar [11 spp.] Faronitopsis Jeannel, 1960b:54 Type species: Faronites convexicollis Jeannel (orig. des.) Madagascar [1 sp.] Faronus Aube, 1844:157 Type species: Faronus lafertei Aube (mono- typy) W Palearctic [35 spp.] Faronellus Jeannel, 1956a:13 Type species: Faronus nicaeensis Saulcy (orig. des.) Golasa Raffray, 1904a:494 Type species: Sagola microcephala Reitter (monotypy) Chile, S Argentina [5 spp.] Golasidius Jeannel, 1 962:3 1 1 Type species: Golasidius microps Jeannel (orig. des.) Chile [2 spp.] Golasina Jeannel, 1962:311 Type species: Golasina robusta Jeannel (orig. des.) Chile [4 spp.] Golasites Jeannel, 1962:310 Type species: Golasites longicornis Jeannel (orig. des.) Chile [1 sp.] Megarafonus Casey, 1897:550 Type species: Megarafonus ventralis Casey (monotypy) W Nearctic, Mexico [5 spp.] Nanorafonus Schuster & Marsh, 1958b: 192 (subgenus) Type species: Megarafonus parvus Schuster & Marsh (orig. des.) Nafonus Schuster & Marsh, 1958b: 194 (subge- nus) Type species: Megarafonus fundus Park (orig. des.) Parafaronus Jeannel, 1954a: 156 Type species: Parafaronus pauliani Jeannel (orig. des.) Madagascar [3 spp.] Prosagola Raffray, 1904a:494 Type species: Sagola elfridae Reitter (mono- typy) Chile [3 spp.] Sagola Sharp, 1874c:506 Type species: Sagola misella Sharp (des. Oke, 1928:5) New Zealand, Australia [139 spp.] Salagosa Raffray, 1904a:494 Type species: Sagola brevipennis Reitter (monotypy) Chile [6 spp.] Sonoma Casey, 1886:195 Type species: Sonoma corticina Casey (des. Lucas, 1920:597) Nearctic [25 spp.] Rafonus Casey, 1894:441 Type species: Faronus tolulae LeConte (monotypy) Stenosagola Broun, 1921a:504 Type species: Stenosagola planiocula Broun (des. here) New Zealand [6 spp.] 18 FIELDIANA: ZOOLOGY Tribe MAYETIINI Mayetia Mulsant & Rey, 1875:87; 1876:9 Type species: Mayetia sphaerifera Mulsant & Rey (monotypy) W Palearctic, Nearctic, Gabon [127 spp.] Promayetia Coiffait, 1962:249 (subgenus) Type species: Mayetia minima Coiffait (orig. des.) Metamayetia Coiffait, 1962:249 (subgenus) Type species: Mayetia nevesi Jarrige (orig. des.) Typhloleptodes Jeannel, 1953b:67 Type species: Typhloleptodes filiformis Jean- nel (orig. des.) Zaire [1 sp.] Typhloleptus Jeannel, 1 95 1 c: 1 5 Type species: Typhloleptus machadoi Jeannel (orig. des.) Afrotropical [12 spp.] Subfamily EUPLECTINAE Section EUPLECTOMORPHI Tribe EUPLECTINI Subtribe ACETALIINA Acetalius Sharp, 1883a:322 Type species: Acetalius dubius Sharp (mono- typy) Japan [2 spp.] Philoscotus Sawada, 1957:23 Type species: Philoscotus longulus Sawada (orig. des.) Japan [4 spp.] Typhlacetalius Jeannel, 1 959b: 1 7 Type species: Typhlacetalius coiffaiti Jeannel (orig. des.) Tribe EUPLECTINI Subtribe BIBLOPLECTINA Aminosimus Raffray, 1898c:267 Type species: Aminosimus madagascariensis Raffray (monotypy) Madagascar [12 spp.] Plectasymus Jeannel, 1960b:80 (subgenus) Type species: Aminosimus caudatus Jeannel (orig. des.) Anotimus Dajoz, 1982:484 Type species: Anotimus elongatus Dajoz (orig. des.) Madagascar [1 sp.] Apheloplectus Raffray, 1913b:126 Type species: Apheloplectus longicollis Raf- fray (monotypy) Seychelles [1 sp.] Asymoplectidius Jeannel, 1956f: 1 1 Type species: Asymoplectidius imerinae Jean- nel (orig. des.) Madagascar [1 sp.] Asymoplectus Raffray, 1897b:55 Type species: Euplectus discicollis Raffray (des. Lucas, 1920:124) Afrotropical, South Africa, Oriental [54 spp.] Asymoplectodes Jeannel, 1960b:75 (subgenus) Type species: Asymoplectus nigripennis Jean- nel (orig. des.) Bibloplectinus Jeannel, 1949a:50 Type species: Bibloplectus variabilis Raffray (orig. des.) Afrotropical, South Africa [16 spp.] Bibloplectopsis Jeannel, 1962:343 Type species: Bibloplectopsis chilensis Jeannel (orig. des.) Chile [1 sp.] Bibloplectus Reitter, 1881:529 Type species: Pselaphus ambiguus Reichen- bach (des. Lucas, 1920:137) Holarctic, Mexico, Oriental, Afrotropi- cal [42 spp.] Bibloplectodes Jeannel, 1949a:49 (subgenus) Type species: Euplectus ruficeps LeConte (orig. des.) Minitiola Karaman, 1969:53 Type species: Minitiola eubeae Karaman (monotypy) Eleusomatus Raffray, 1904a:548 Type species: Euplectus allocephalus Broun (des. Lucas, 1920:259) New Zealand [6 spp.] Eleusotamus Raffray, 1904a:548 (misspelling, first reviser Raffray, 1 908a: 1 12) Euphiliops Jeannel, 1 959d: 1 1 9 Type species: Philiopsis humicola Jeannel (orig. des.) Afrotropical [31 spp.] Euplectodina Raffray, 1898c:228 Type species: Euplectus hipposideros Schau- fuss (monotypy) Oriental [1 sp.] Haploplectus Jeannel, 1 95 1 f:25 Type species: Haploplectus b if oveat us Jeannel (orig. des.) Zaire, Angola, South Africa [2 spp.] Humbertella NEW NAME (for Humbertiella Jeannel) Madagascar [1 sp.] Humbertiella Jeannel, 1960a:7 (preocc, not Saussure, 1869) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 19 Type species: Humbertiella caeca Jeannel (orig. des.) Hypoplectus Raffray, 1913a:24 Type species: Hypoplectus abdominalis Raf- fray (monotypy) Afrotropical [16 spp.] Hypoplectinus Jeannel, 1952a: 122 (subgenus) Type species: Pseudoplectus crassipes Jeannel (orig. des.) Leptophiliops Jeannel, 1 952a:l 14 Type species: Leptophiliops alticola Jeannel (orig. des.) Zaire [2 spp.] Limoniates Raffray, 1898c:231 Type species: Limoniates australis Raffray (monotypy) Australia [6 spp.] Methorius Raffray, 1904a:537 Type species: Periplectus bicolor Raffray (monotypy) Zimbabwe, Vietnam [2 spp.] Microplectus Raffray, 1898c:264 Type species: Microplectus exiguus Raffray (monotypy) Singapore, New Guinea [5 spp.] Neoplectidius Jeannel, 1960b:84 Type species: Neoplectidius convexifrons Jeannel (orig. des.) Madagascar [1 sp.] Neothesiastes Jeannel, 1960b:73 Type species: Neothesiastes scotti Jeannel (orig. des.) Madagascar [2 spp.] Omotimellus Jeannel, 1952a: 116 Type species: Omotimellus exilis Jeannel (orig. des.) Zaire [2 spp.] Omotimiotes Jeannel, 1954a: 170 Type species: Omotimiotes caecus Jeannel (orig. des.) Madagascar [1 sp.] Omotimus Raffray, 1904a:534 Type species: Trimiodytes sulcifrons Raffray (monotypy) South Africa [1 sp.] Pachyeuplectus Jeannel, 1 954a: 1 79 Type species: Pachyeuplectus vadoni Jeannel (orig. des.) Madagascar [1 sp.] Paraphiliopsis Jeannel, 1959d:120; 1960b:70 Type species: Philiopsis unicolor Raffray (orig. des.) Madagascar [5 spp.] Patreus Broun, 1904:47 Type species: Patreus lewisi Broun (mono- typy) New Zealand [1 sp.] Periplectus Raffray, 1887:55 Type species: Periplectus nigripennis Raffray (monotypy) Afrotropical [14 spp.] Philiopsis Raffray, 1893a:471 Type species: Philiopsis exigua Raffray (monotypy) Oriental, Japan, New Guinea, ?Zaire [7 spp.] Placodium Broun, 1893a: 1431 Type species: Placodium zenarthrum Broun (monotypy) New Zealand, New Caledonia [2 spp.] Plectiastes Jeannel, 1960a:9 Type species: Plectiastes lepidus Jeannel (orig. des.) Madagascar [1 sp.] Plectodytes Jeannel, 1956f:8 Type species: Omotimiotes endogaeus Jean- nel (orig. des.) Madagascar [2 spp.] Prodalma Raffray, 1897b:51 Type species: Prodalma capensis Raffray (monotypy) South Africa [1 sp.] Pseudomotimus Jeannel, 1952d:28 Type species: Pseudomotimus alutaceus Jean- nel (orig. des.) Zaire [2 spp.] Pseudoplectus Reitter, 1881:531 Type species: Euplectus perplexus Duval (monotypy) W Palearctic [1 sp.] Rhynchoplectus Jeannel, 1959d: 1 35 Type species: Microplectus minutus Jeannel (orig. des.) Zaire [1 sp.] Tiomomus Jeannel, 1962:341 Type species: Tiomomus rostratus Jeannel (orig. des.) Chile, S Argentina [2 spp.] Trimiophanes Jeannel, 1954a: 168 Type species: Trimiophanes globulicornis Jeannel (orig. des.) Madagascar [16 spp.] Vidamodes Broun, 1921b:606 Type species: Vidamodes furvus Broun (monotypy) New Zealand [1 sp.] 20 FIELDIANA: ZOOLOGY Tribe EUPLECTINI Subtribe BIBLOPORELLINA Aphilia Reitter, 1882c:210; 1883c:297 Type species: Aphilia femorata Reitter (sub- seq. mon. Reitter, 1883c:297) Oriental [4 spp.] Apoterus Raffray, 1894b:21 1 Type species: Apoterus laevis Raffray (mono- typy) Singapore, New Guinea [2 spp.] Bibloporellus Jeannel, 1949a:47 Type species: Bibloporellus microphthalmus Jeannel (orig. des.) Afrotropical [18 spp.] Chaetorhopalus RafTray, 1887:48 Type species: Chaetorhopalus unicolor Raf- fray (monotypy) Afrotropical, Oriental [11 spp.] Chaetorrhopalus Jeannel, 1949a:46 (misspell- ing) EupinionOke, 1928:14 Type species: Eupinion crassipes Oke (orig. des.) Australia [1 sp.] Narcissiella Jeannel, 1953b:82 Type species: Narcissiella elegantula Jeannel (orig. des.) Zaire, Kenya [2 spp.] Omobathus Raffray, 19 13b: 127 Type species: Omobathus elegans Raffray (monotypy) Seychelles [1 sp.] Opisthosemus Jeannel, 1952a:97 Type species: Opisthosemus leptus Jeannel (orig. des.) Zaire [1 sp.] Pseudozibus Jeannel, 1956d:365 Type species: Pseudozibus simplicipennis Jeannel (orig. des.) Zaire, Madagascar, Italy, Japan [5 spp.] Aphiliopsis Besuchet, 1956a:369 Type species: Aphiliops crassipes Raffray (orig. des.) Typhlokahusia Jeannel, 1953b:80 Type species: Typhlokahusia ziboides Jeannel (orig. des.) Zaire [1 sp.] Tribe EUPLECTINI Subtribe BIBLOPORINA Abdiunguis Park & Wagner, 1962:10 Type species: Abdiunguis fenderi Park & Wag- ner (orig. des.) W Nearctic (Oregon) [1 sp.] Biblomelba Park, 1952c: 106 Type species: Biblomelba profunda Park (orig. des.) Mexico [1 sp.] Biblomimus RafTray, 1904a:545 Type species: Biblomimus minutus Raffray (monotypy) Mexico, Honduras, West Indies [4 spp.] Bibloporus Thomson, 1859:53 Type species: Euplectus bicolor Denny (orig. des.) Palearctic, Zaire, E Nearctic [8 spp.] Faliscus Casey, 1 884: 1 20 Type species: Faliscus bicanalis Casey (mono- typy) Bibloporites Jeannel, 1950:80 (subgenus) Type species: Bibloporus mayeti Guillebeau (orig. des.) Synbibloporus Sawada, 1956:101 (subgenus) Type species: Bibloporus japonicus Sawada (orig. des.) Biblomorphus Lobl, 1961:99 (subgenus) Type species: Bibloporus variicolor Reitter (orig. des.) Dalma Sharp, 1874c:504 Type species: Dalma pubescens Sharp (mono- typy) New Zealand [4 spp.] Euglyptus Broun, 1893a: 141 1 Type species: Euglyptus elegans Broun (monotypy) New Zealand [8 spp.] Microtyrus Broun, 1893a: 141 3 Type species: Microtyrus punctatus Broun (monotypy) Euplectopsis Raffray, 1890a: 101 Type species: Trichonyx microcephalus Reit- ter (monotypy) New Zealand [40 spp.] Reitteronymus Raffray, 1 890a: 107 Type species: Trichonyx longicollis Reitter (monotypy) Eutyphlus LeConte, 1880:185 Type species: Eutyphlus similis LeConte (monotypy) E Nearctic [5 spp.] Nicotheus Casey, 1 884: 1 2 1 Type species: Nicotheus tibialis Casey (mono- typy) Nicothaeus Raffray, 1890a: 102 (misspelling) Planityphlus Park, 1956:62 (subgenus) Type species: Eutyphlus thoracicus Park (orig. des.) Hatchia Park & Wagner, 1962:9 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 21 Type species: Hatchia carinata Park & Wag- ner (orig. des.) W Nearctic (Washington) [ 1 sp.] Macroplectus Raffray, 1898c:255 Type species: Macroplectus calcaratus Raffray (des. Lucas, 1920:389) Australia [7 spp.] Mexigaster Park, 1952c: 110 Type species: Mexigaster boneti Park (orig. des.) Mexico [1 sp.] Mexiplectus Park, 1943:174 Type species: Mexiplectus emersoni Park (orig. des.) Mexico [2 spp.] Plectomorphus Raffray, 1898c:265 Type species: Euplectus spinifer Broun (monotypy) New Zealand [ 1 4 spp.] Plectostenus Lea, 1910:146 Type species: Plectostenus gracilicornis Lea (monotypy) Australia [2 spp.] Plectusodes Raffray, 1898c:261 Type species: Plectusodes breviceps Raffray (monotypy) Australia [3 spp.] Protoplectus Raffray, 1 898c:260 (not Leleup, 1 974) Type species: Protoplectus pubescens Raffray (monotypy) Australia [1 sp.] Trychioplectus Raffray, 1898c:272 Type species: Trychioplectus geminatus Raf- fray (monotypy) Australia [1 sp.] Whitea Hutton, 1 904: 1 79 (new name for Brounia Raffray) (published by May 1 904) New Zealand [1 sp.] Brounia Raffray, 1898c:266 (preocc, not Sharp, 1878) Type species: Euplectus laevifrons Broun (monotypy) Brouniella Raffray, 1904a: 5 84 (unnecessary new name for Brounia Raffray) (published September 1904) Zelandius Raffray, 1898c:272 Type species: Euplectus obscurus Broun (monotypy) New Zealand [17 spp.] Tribe EUPLECTINI Subtribe CHRESTOMERINA Chrestomera Jeannel, 1962:344 Type species: Chrestomera longicollis Jeannel (orig. des.) Chile [5 spp.] Tribe EUPLECTINI Subtribe EUPLECTINA Acolonia Casey, 1894:454 Type species: Euplectus cavicollis LeConte (monotypy) SE Nearctic (Florida) [ 1 sp.] Adrogaster Raffray, 1890a: 100 Type species: Adrogaster longipennis Raffray (subseq. mon. Raffray, 1890b: 195) Brazil [1 sp.] Afroplectus Jeannel, 1952a: 130 Type species: Euplectus africanus Raffray (orig. des.) Afrotropical, South Africa [23 1 spp.] Afroplectidius Jeannel, 1952e:200 (subgenus) Type species: Euplectus raffrayi Jeannel (orig. des.) Afroplectodes Jeannel, 1952e:202 (subgenus) Type species: Euplectus montanus Raffray (orig. des.) Afroplectaulax Jeannel, 1959e:202 (subgenus) Type species: Afroplectus remyi Jeannel (orig. des.) Aloxomidus Raffray, 1903a:34 Type species: Aloxomidus variolosus Raffray (monotypy) New Guinea [ 1 sp.] Anomoplectus Jeannel, 1 95 1 e: 1 29 Type species: Euplectus micropalpus Jeannel (monotypy) Zaire [1 sp.] Echinoplectus Jeannel, 1 960c: 1 04 Type species: Echinoplectus joannae Jeannel (orig.des.) Aulaxus Raffray, 1898c:225 Type species: Aulaxus rugicollis Raffray (des. Lucas, 1920:128) Australia [2 spp.] Bothriocephalotes Leleup, 1978b:668 Type species: Bothriocephalotes graft Leleup (orig. des.) Ivory Coast [2 spp.] Calarus Raffray, 1903a:32 Type species: Calarus puncticeps Raffray (des. Lucas, 1920:156) New Guinea [3 spp.] Coptoplectus Leleup, 1974a: 183 22 FIELDIANA: ZOOLOGY Type species: Coptoplectus bifalcatus Leleup (orig. des.) Ethiopia [1 sp.] Deroplectus Raffray, 1898c:271 Type species: Euplectus excisus King (mono- typy) Australia [1 sp.] Diarrogus Raffray, 1903a:36 Type species: Diarrogus pubescens Raffray (monotypy) New Guinea [1 sp.] Dissemoplectus Jeannel. 1 952a: 1 6 1 Type species: Euplectus tuberculiceps Raffray (orig. des.) South Africa [1 sp.] Epithematus Raffray, 1903a:39 Type species: Epithematus nitidus Raffray (monotypy) New Guinea [1 sp.] Euplectamecia Park, 1952c: 102 Type species: Euplectamecia hidalgensis Park (orig. des.) Mexico [1 sp.] Euplecterga Park & Wagner, 1962:15 Type species: Oropodes impressicollis Park & Wagner (orig. des.) W Nearctic [3 spp.] Euplectus Leach, 1817:82 Type species: Euplectus reichenbachii Leach (= Pselaphus nanus Reichenbach) (monotypy) Holarctic, Neotropical, Oriental, Afrotropi- cal, ?New Zealand, ?Australia [117 spp.] Diplectellus Reitter, 1909:207 (subgenus) Type species: Euplectus afer Reitter (des. Jeannel, 1950:100) Euplectellus Reitter, 1909:207 (subgenus) Type species: Euplectus huemmleri Reitter (monotypy) Endoplectus Raffray. 1 9 1 0b:254 Type species: Euplectus doderoi Reitter (monotypy) Pareuplectus Jeannel, 1952e:211 (subgenus) Type species: Euplectus delamarei Jeannel (orig. des.) Euplectoides Jeannel, 1 954b: 1 06 Type species: Euplectoides pearcei Jeannel (orig. des.) Diplectidius Jeannel, 1956a:31 (subgenus) Type species: Euplectus theryi Guillebeau (orig. des.) Euplectinus Jeannel, 1 956a:3 1 Type species: Euplectus afer Reitter (orig. des.) Fijiastes Park, 195 2b: 11 Type species: Fijiastes zimmermani Park (orig. des.) Fiji [1 sp.] Gabata Raffray, 1898d:387 Type species: Gabata semipunctata Raffray (monotypy) South Africa, New Zealand [5 spp.] Leptoplectus Casey, 1908:266 Type species: Euplectus pertenuis Casey (des. Lucas. 1920:371) E Nearctic, Palearctic, Oriental, New Guinea [8 spp.] Grammoplectus Jeannel, 1 950: 1 00 Type species: Euplectus spinolae Aube (orig. des.) Archeuplectus Jeannel, 1 952a: 1 30 Type species: Euplectus annamita Raffray (orig. des.) Lioplectus Raffray, 1898c:260 Type species: Lioplectus nitidus Raffray (monotypy) Argentina [7 spp.] Meliceria Raffray, 1898c:254 Type species: Euplectus acanthifer Reitter (monotypy) W Palearctic [5 spp.] Cyrtoplectus Normand, 1904:200 (subgenus) Type species: Euplectus sulciventris Guille- beau (monotypy) Metaplectodes Jeannel, 1960c:93 Type species: Metaplectodes leleupi Jeannel (orig. des.) Zaire [1 sp.] Nesiotoplectus Jeannel. 1954a: 175 Type species: Euplectus megacephalus Raf- fray (orig. des.) Madagascar, Comoros [15 spp.] Oropodes Casey, 1894:453 Type species: Oropodes orbiceps Casey (monotypy) W Nearctic [6 spp.] Panaramecia Park, 1942:94 Type species: Panaramecia williamsi Park (orig. des.) Panama [1 sp.] Plectophloeus Reitter, 1 89 1 a: 1 23 Type species: Euplectus erichsoni Aube (des. Jeannel, 1950:132) W Palearctic [9 spp.] Plectophleus Karaman, 1 962b: 1 45 (misspelling) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 23 Plectoprotus NEW NAME (for Protoplectus Le- leup) Zaire [1 sp.] Protoplectus Leleup, 1974a: 186 (preocc, not Raffray, 1898) Type species: Protoplectus lejeunei Leleup (orig. des.) Pseudactium Casey, 1908:271 Type species: Pseudactium carolinae Casey (orig. des.) E Nearctic [3 spp.] Pycnoplectus Casey, 1897:552 Type species: Euplectus linearis LeConte (des. Wagner, 1975:158) Nearctic [1 1 spp.] Racemia NEW GENUS (for Ramecia of Grigarick & Schuster, 1980, not Casey, 1894; see Dis- cussion) Type species: Euplectus arcuatus LeConte (des. here) E Nearctic [5 spp.] Ramecia Casey, 1894:450 (see Discussion) Type species: Euplectus crinitus Brendel (des. Lucas, 1920:564) E Nearctic [1 sp.] Liniolis Grigarick & Schuster, 1980:34 Type species: Euplectus crinitus Brendel (orig. des.) Saxet Grigarick & Schuster, 1980:37 Type species: Euplectus decoris Casey (orig. des.) SW Nearctic [3 spp.] Scotoplectus Reitter, 1879:44 Type species: Scotoplectus capellae Reitter (monotypy) W Palearctic [6 spp.] Moczarskia Blattny & Blattny, 1914:93 Type species: Scotoplectus winkleri Blattny & Blattny (monotypy) Seleneuplectus Jeannel, 1955a:20 Type species: Seleneuplectus scutellatus Jean- nel (orig. des.) Uganda [1 sp.] Stenoplectus Raffray, 1898c:243 Type species: Stenoplectus sternalis Raffray (monotypy) Australia [1 sp.] Thesiastes Casey, 1894:457 Type species: Euplectus debilis LeConte (des. Bowman, 1934:144) Nearctic, Neotropical, Chile, Oriental, New Guinea, Tanzania [19 spp.] Trigonoplectus Bowman, 1934:37 24 Type species: Trigonoplectus minutus Bow- man (orig. des.) E Nearctic [2 spp.] Trimioplectus Brendel, 1891:50 Type species: Trimioplectus obsoletus Brendel (des. Lucas, 1920:655) E Nearctic [2 spp.] Tribe EUPLECTINI Subtribe PANAPHANTINA Acotreba Reitter, 1882c:209; 1883b:52 Type species: Acotreba simoni Reitter (sub- seq. mon. Reitter, 1883b:52) Chile [1 sp.] Panaphantus Kiesen wetter, 1858:48 Type species: Panaphantus atomus Kiesen- wetter (monotypy) W Palearctic, Zaire [2 spp.] Thesiectus Park, 1960a:8 Type species: Thesiectus probus Park (orig. des.) Jamaica [1 sp.] Thesium Casey, 1884:117 Type species: Euplectus cavifrons LeConte (des. Bowman, 1934:144) E Nearctic, Neotropical [18 spp.] Apothinus Sharp, 1887:35 Type species: Apothinus impressifrons Sharp (des. Lucas, 1920:113) Verabarolus Park, 1942:104 Type species: Verabarolus subdendrus Park (orig. des.) Panama [1 sp.] Tribe EUPLECTINI Subtribe RHINOSCEPSINA Morius Casey, 1894:445 Type species: Morius occidens Casey (mono- typy) W Nearctic (California) [1 sp.] Rhinoscepsis LeConte, 1878:382 Type species: Rhinoscepsis bistriatus LeConte (monotypy) Nearctic, Neotropical [11 spp.] Rhinosceptis Reitter, 1882c:209 (misspelling) Rhynosceptis Raffray, 1890a:97 (misspelling) Rhynoscepsis Raffray, 1898c:223 (misspelling) Rafrhisis Park, 1945a:283 (subgenus) Type species: Rhinoscepsis orbis Park (mono- typy) FIELDIANA: ZOOLOGY Tribe EUPLECTINI Subtribe TRIMIINA Actiastes Casey, 1897:561 Type species: Trimium foveicolle LeConte (orig. des.) Nearctic [8 spp.] Actiades Lucas, 1920:72 (misspelling) Actionoma Raffray, 1898c:235 Type species: Actionoma obesum Raffray (monotypy) Mexico [1 sp.] Actinoma Park, 1942:108 (misspelling) Actium Casey, 1886:201 Type species: Trimium californicum LeConte (des. Brendel, 1891:40) Nearctic, Mexico, Central America, West In- dies [38 spp.] Proplectus Raffray, 1 890a: 107 Type species: Proplectus decipiens Raffray (subseq. mon. Raffray, 1890b: 197) Actizona Chandler, 1985:112 Type species: Actizona chuskae Chandler (orig. des.) Nearctic [2 spp.] Allobrox Fletcher, 1928:205 Type species: Allobrox dampfi Fletcher (monotypy) SW Nearctic, Mexico [3 spp.] Cutrimia Park, 1945b:336 Type species: Cupila mexicana Park (orig. des.) Allomelba Park, 1954a: 16 Type species: Allomelba antennata Park (orig. des.) West Indies [1 sp.] Alloplectus Broun, 191 1:697 Type species: Euplectus claviger Broun (orig. des.) New Zealand [3 spp.] Allotrimium Park, 1943:187 Type species: Allotrimium michoacanensis Park (orig. des.) SW Nearctic, Mexico [3 spp.] Amudrocerus Raffray, 1894b:218 Type species: Amudrocerus grandiceps Raf- fray (monotypy) Singapore [1 sp.] Aphiliops Reitter, 1 884c:208 (new name for Philus Saulcy) France (Corsica) [1 sp.] Philus Saulcy, 1874:40 bis (preocc, not Saun- ders, 1853) Type species: Philus aubei Reitter (des. Reit- ter, 1881:518) Basolum Casey, 1897:571 Type species: Trimium impunctatum Brendel (orig. des.) E Nearctic [2 spp.] Cupila Casey, 1897:561 Type species: Trimium clavicorne Maklin (orig. des.) W Nearctic [3 spp.] Dalmomelba Park, 1954a:6 Type species: Dalmomelba cazieri Park (orig. des.) Bahamas [1 sp.] Dalmoplectus Raffray, 1 890a: 102 (see Discussion) Type species: Dalmodes batrisoides Reitter (monotypy) (see Discussion) Brazil [1 sp.] Dalmosanus Park, 1952c: 126 Type species: Dalmosanus inoculus Park (orig. des.) Mexico [2 spp.] Triangusella Park, 1952c: 132 Type species: Triangusella acuta Park (orig. des.) Dalmosella Casey, 1897:570 Type species: Dalmosella tenuis Casey (orig. des.) E Nearctic [ 1 sp.] Haasellia Wagner, 1984:282 (new name for Haa- siella Park) Puerto Rico [1 sp.] Haasiella Park, 1976:35 (preocc, not Pocock, 1901; Kieffer, 1913; Lindholm, 1925; Wirth, 1973) Type species: Haasiella medicina Park (orig. des.) Hanfordia Park, 1 960a: 1 0 Type species: Hanfordia absoluta Park (orig. des.) Jamaica [1 sp.] Hispanisella Park, 1976:32 Type species: Hispanisella haitiana Park (orig. des.) Haiti [2 spp.] Latomelba Park, 1955:111 Type species: Latomelba quadrisicca Park (orig. des.) Jamaica [2 spp.] Lemelba Park, 1953b:l Type species: Lemelba davisi Park (orig. des.) SE Nearctic, West Indies [3 spp.] Malleoceps Park, 1954b:l NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 25 Type species: Malleoceps darlingtoni Park (orig. des.) West Indies [2 spp.] Melba Casey, 1897:565 Type species: Trimium thoracicum Brendel (des. Bowman, 1934:144) Nearctic, Neotropical [25 spp.] Rameloidea Park, 1942:121 (subgenus) Type species: Melba temporalis Raffray (monotypy) Vertelba Park, 1942:120 (subgenus) Type species: Melba caviceps Raffray (mono- typy) Frontelba Park, 1942:120 (nomen nudum, no type species designated) Asymmelba Park, 1942:121 {nomen nudum, no type species designated) Asymmelba Park, 1952c: 140 (subgenus) Type species: Melba clavata Raffray (orig. des.) Frontelba Park, 1952c: 140 (subgenus) Type species: Melba frontalis Raffray (orig. des.) Cismelba Park, 1960a: 16 (subgenus) Type species: Melba trifoveata Park (orig. des.) Melbamima Raffray, 1909:17 Type species: Melbamina clavicornis Raffray (monotypy) Brazil [1 sp.] Minibi Grigarick & Schuster, 1 980:40 Type species: Melba zonula Park (orig. des.) Bahamas [1 sp.] Neactium Jeannel, 1962:339 Type species: Pseudoplectus trimiiformis Reitter (orig. des.) Chile [2 spp.] Neodalmus Raffray, 1891a:304 Type species: Neodalmus carinatus Raffray (monotypy) Venezuela [1 sp.] Paractium Jeannel, 1962:334 Type species: Paractium kuscheli Jeannel (orig. des.) Chile, S Argentina [7 spp.] Perimelba Park, 1952c: 141 Type species: Melba granulosa Park (orig. des.) Neotropical [7 spp.] Perimelba Park, 1943:183 {nomen nudum, no type species designated) Pilactium Grigarick & Schuster, 1970:37 Type species: Pilactium summersi Grigarick & Schuster (orig. des.) W Nearctic [2 spp.] Prophilus Raffray, 1890a: 100 Type species: Prophilus minutus Raffray (sub- seq. mon. Raffray, 1890b: 195) Singapore [2 spp.] Pseudotrimium Raffray, 1898c:230 Type species: Pseudotrimium microcephalum Raffray (monotypy) Mexico [1 sp.] Pygmactium Schuster & Grigarick, 1968:1 12 Type species: Pygmactium steevesi Schuster & Grigarick (orig. des.) E Nearctic [2 spp.] Quadrelba Park, 1952c: 140 Type species: Trimiopsis parmata Reitter (orig. des.) Neotropical [3 spp.] Quadrelba Park, 1942:120 {nomen nudum, no type species designated) Ramelbida Park, 1942:1 12 Type species: Melba quadrifoveata Raffray (orig. des.) Virgin Islands [1 sp.] Sandersonella Park, 1976:38 Type species: Sandersonella transversa Park (orig. des.) Haiti [1 sp.] Saulcyella Reitter, 1901:229 Type species: Euplectus schmidti Maerkel (monotypy) W Palearctic [1 sp.] Simplona Casey, 1897:561 Type species: Simplona arizonica Casey (orig. des.) SW Nearctic (Arizona) [2 spp.] Tomoplectus Raffray, 1898c:266 Type species: Tomoplectus cordicollis Raffray (monotypy) Mexico, SW Nearctic (Arizona) [3 spp.] Trimioarcus Park, 1952c: 130 Type species: Trimioarcus incisurus Park (orig. des.) Mexico, SW Nearctic (Arizona) [2 spp.] Trimiodina Raffray, 1898c:231 Type species: Trimium concolor Sharp (monotypy) Guatemala [1 sp.] Trimiomelba Casey, 1897:563 Type species: Euplectus dubius LeConte (des. Lucas, 1920:655) E Nearctic [3 spp.] Trimiomorphus Raffray, 1 890a: 1 00 Type species: Trimiomorphus elongatus Raf- fray (subseq. mon. Raffray, 1 890b: 1 94) Singapore [1 sp.] 26 FIELDIANA: ZOOLOGY Trimiopsis Reitter, 1882a: 149 Type species: Trimiopsis claviceps Reitter (monotypy) Neotropical [4 spp.] Trimiosella Raffray, 1898c:236 Type species: Trimiopsis anguina Reitter (monotypy) Virgin Islands [1 sp.] Trimiovillus Park, 19 54a: 11 Type species: Trimiovillus bahamicus Park (orig. des.) West Indies [2 spp.] Trimium Aube, 1833:44 Type species: Pselaphus brevicornis Reichen- bach (monotypy) W Palearctic [25 spp.] Aphanogramme Dodero, 1919:242 (subgenus) Type species: Trimium asturicum Dodero (monotypy) ZibusSaulcy, 1874:40 Type species: Euplectus leiocephalus Aube (des. Lucas, 1920:680) W Palearctic [1 sp.] Zolium Casey, 1897:562 Type species: Trimiopsis eggersi Reitter (orig. des.) Virgin Islands [1 sp.] Zonaira Grigarick & Schuster, 1980:43 Type species: Zonaira trilinea Grigarick & Schuster (orig. des.) SW Nearctic (Arizona) [2 spp.] Tribe EUPLECTINI Subtribe TRIMIODYTINA Anoplectus Raffray, 1897b:59 Type species: Anoplectus niger Raffray (monotypy) South Africa [2 spp.] Trimiodytes Raffray, 1897b:52 Type species: Trimiodytes palustris Raffray (monotypy) South Africa [10 spp.] Tribe EUPLECTINI, incertae sedis Dalmisus Sharp, 1886:381 Type species: Dalmisus batrisodes Sharp (monotypy) New Zealand [1 sp.] Epiplectus Raffray, 1898c:264 Type species: Philiopsis cavifrons Raffray (monotypy) Singapore, Burma [3 spp.] Euplectina Raffray, 1894b:216 Type species: Euplectina nigripennis Raffray (des. Lucas, 1920:284) Singapore [2 spp.] Glastus Raffray, 1898c:254 Type species: Euplectus cicatricosus Raffray (monotypy) Singapore [1 sp.] Kenocoelus Broun, 191 1:700 Type species: Kenocoelus dimorphus Broun (monotypy) New Zealand [1 sp.] Mirellus Raffray, 1894b:204 Type species: Mirellus sulcicollis Raffray (monotypy) Singapore [1 sp.] Pareuplectops Jeannel, 1957a:7 Type species: Phthartomicrus pubescens Raf- fray (orig. des.) Oriental [2 spp.] Philotrimium Blattny, 1925:186 Type species: Trimium abdominale Mot- schulsky (monotypy) Burma [1 sp.] Sagolonus Raffray, 1898c:265 Type species: Euplectus patronus Broun (monotypy) New Zealand [3 spp.] Sampsa Raffray, 1898c:254 Type species: Sampsa longipennis Raffray (monotypy) Sumatra [1 sp.] Trimicerus Motschulsky, 1855:16 Type species: Trimicerus pacificus Motschul- sky (des. Lucas, 1920:654) Panama [2 spp.] Vidamus Raffray, 1898c:252 Type species: Euplectus incertus Reitter (des. here) New Zealand [20 spp.] Tribe PTERACMINI Andiplectops Jeannel, 1962:358 Type species: Andiplectops temporalis Jeannel (orig. des.) Chile [2 spp.] Araucaniotes Jeannel, 1962:371 Type species: Araucaniotes parallelus Jeannel (orig. des.) Chile [1 sp.] Caligrua Jeannel, 1962:352 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 27 Type species: Caligma thoracica Jeannel (orig. des.) Chile, S Argentina [3 spp.] Caligruacmes Jeannel, 1963:359 Type species: Caligruacmes convexicollis Jeannel (orig. des.) Chile [1 sp.] Daliacmes Jeannel, 1962:354 Type species: Daliacmes insularis Jeannel (orig. des.) Chile [5 spp.] Etopias Jeannel, 1962:357 Type species: Etopias longulus Jeannel (orig. des.) Chile [1 sp.] Euplectops Reitter, 1882c:210 Type species: Euplectus odewahni King (des. Raffray, 1900b: 132) Australia [15 spp.] Nothoplectus Jeannel, 1963:360 Type species: Nothoplectus depressus Jeannel (orig. des.) Chile [1 sp.] Paraplectus Raffray, 1898c:269 Type species: Paraplectus punctulatus Raffray (des. Lucas, 1920:482) Australia [4 spp.] Parapteracmes Jeannel, 1962:361 Type species: Parapteracmes kuscheli Jeannel (orig. des.) Chile [5 spp.] Pteracmes Raffray, 1890a: 102 Type species: Pteracmes schaufussi Raffray (subseq. mon. Raffray, 1890b: 196) Chile, S Argentina [5 spp.] Pteracmidius Jeannel, 1962:365 Type species: Pteracmidius frontalis Jeannel (orig. des.) Chile [6 spp.] Pteroplectus Raffray, 1898c:259 Type species: Euplectus grandicornis Schau- fuss (monotypy) Chile [2 spp.] Trepacmes Jeannel, 1962:360 Type species: Trepacmes germaini Jeannel (orig. des.) Chile [1 sp.] Tribe RAFFRAYHNI Ambalavoa Dajoz, 1982:487 Type species: Ambalavoa antennalis Dajoz (orig. des.) Madagascar [1 sp.] Ankavena Jeannel, 1954a:192 Type species: Ankavena depressa Jeannel (orig. des.) Madagascar [2 spp.] Apotectus NEW NAME (for Autoplectus Raffray) Madagascar, Angola [7 spp.] Autoplectus Raffray, 1 883:248 (preocc, not Bal- samo-Crivelli, 1843) Type species: Autoplectus torticornis Raffray (monotypy) Badensia Jeannel, 1 954a: 1 93 Type species: Badensia pauliani Jeannel (orig. des.) Madagascar [13 spp.] Dalmina Raffray, 1887:46 Type species: Dalmina globulicornis Raffray (monotypy) South Africa, Zimbabwe [20 spp.] Dalminiola Jeannel, 1964c:97 (subgenus) Type species: Dalmina nigrita Jeannel (monotypy) Dalminella Jeannel, 1964c:75 Type species: Dalminella parvula Jeannel (orig. des.) South Africa [1 sp.] Dalminia NEW NAME (for Damilina Jeannel) South Africa [6 spp.] Damilina Jeannel, 1964c:69 (preocc, not Hor- ny, 1961) Type species: Damilina leleupi Jeannel (orig. des.) Dalminiastes Jeannel, 1962:373 Type species: Dalminiastes semirugosus Jean- nel (orig. des.) Chile, S Argentina [5 spp.] Dalminiomus Jeannel, 1962:378 Type species: Dalminiomus araucanus Jean- nel (orig. des.) Chile [1 sp.] Damilinidius Jeannel, 1964c:74 Type species: Damilinidius capensis Jeannel (orig. des.) South Africa [1 sp.] Epullea Jeannel, 1964c:66 Type species: Epullea capensis Jeannel (orig. des.) South Africa [2 spp.] Faronoma Raffray, 1893b:259 Type species: Rybaxis cavangula Reitter (monotypy) Chile [1 sp.] Frutillariotes Jeannel, 1962:381 Type species: Frutillariotes kuscheli Jeannel (orig. des.) Chile [1 sp.] 28 FIELDIANA: ZOOLOGY Imeriniella Jeannel, 1960a: 13 Type species: Ranavala subterranea Jeannel (orig. des.) Madagascar [3 spp.] Imerinella Jeannel, 1960b: 110 (objective syn- onym of Imeriniella Jeannel) Type species: Ranavala subterranea Jeannel (orig. des.) Macroraffrayia Jeannel, 1955b:69 Type species: Raffrayia longula Rattray (orig. des.) South Africa [2 spp.] Mildana Jeannel, 1964b:9 Type species: Mildana castrii Jeannel (orig. des.) Chile [1 sp.] Neoraffrayia Jeannel, 1955b:90 Type species: Raffrayia nasuta RafFray (orig. des.) South Africa [22 spp.] Parepullea Jeannel, 1964c:69 Type species: Dalmina elegans Raffray (orig. des.) South Africa [1 sp.] Platyplectus Jeannel, 1962:377 Type species: Platyplectus chilotus Jeannel (orig. des.) Chile [2 spp.] Pseudeuplectus Enderlein, 1909:383 Type species: Pseudeuplectus antarcticus En- derlein (monotypy) Crozet Islands [1 sp.] Raffrayia Reitter, 1882c:21 1 Type species: Trichonyx antennatus Raffray (orig. des.) Afrotropical, South Africa [37 spp.] Raffrayidius Jeannel, 1955b:88 Type species: Raffrayidius rhinoceros Jeannel (orig. des.) South Africa [14 spp.] Raffrayites Jeannel, 1955b:99 Type species: Raffrayites braunsi Jeannel (orig. des.) South Africa [11 spp.] Raffrayitidia Jeannel, 1964c:l 19 (subgenus) Type species: Raffrayites pubescens Jeannel (orig. des.) Raffrayola Jeannel, 1955b:70 Type species: Raffrayia bicolor Raffray (orig. des.) South Africa [15 spp.] Ranavala Raffray, 1898c:224 Type species: Autoplectus integricollis Raffray (monotypy) Madagascar [21 spp.] Masoala Jeannel, 1954a: 190 (subgenus) Type species: Masoala curta Jeannel (orig. des.) Fanovana Jeannel, 1960b: 100 (subgenus) Type species: Ranavala longifrons Jeannel (orig. des.) Ranavalidius Jeannel, 1959e:204 Type species: Ranavalidius nasutus Jeannel (orig. des.) Madagascar [3 spp.] Ranavalodes Jeannel, 1960b:99 Type species: Ranavalodes caecus Jeannel (orig. des.) Madagascar [2 spp.] Typhloraffrayia Jeannel, 1955b:89 Type species: Raffrayia abdominalis Raffray (orig. des.) South Africa [ 1 sp.] Xenogyna Raffray, 1897b:60 Type species: Xenogyna heterocera Raffray (monotypy) South Africa [6 spp.] Xenogynidia Jeannel, 1964c:80 Type species: Xenogynidia laticollis Jeannel (orig. des.) South Africa [1 sp.] Tribe TROGASTRINI Subtribe MITRAMETOPINA Mitrametopus Raffray, 1911:50 (new name for Mitracephala Raffray) Brazil [1 sp.] Mitracephala Raffray, 1890a: 107 (preocc, not Thomson, 1859) Type species: Mitracephala longipennis Raf- fray (subseq. mon. Raffray, 1 890b: 1 98) Tribe TROGASTRINI Subtribe PHTEGNOMINA Phtegnomus Raffray, 1890a: 107 Type species: Phtegnomus oberthuri Raffray (subseq. mon. Raffray, 1890b: 198) Brazil, Venezuela [3 spp.] Tribe TROGASTRINI Subtribe RHEXIINA Rhexius LeConte, 1849:102 Type species: Rhexius insculptus LeConte (monotypy) Nearctic, Neotropical [19 spp.] Rafrhexius Park, 1952c:75 (subgenus) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 29 Type species: Rhexius elegans Raffray (orig. des.) Rhexinexus Park, 1952c:85 (subgenus) Type species: Rhexius giganticus Park (orig. des.) Tribe TROGASTRINI Subtribe TRISIGNINA Trisignis Park & Schuster, 1955:1 Type species: Trisignis marshi Park & Schus- ter (orig. des.) W Nearctic (California) [2 spp.] Tribe TROGASTRINI Subtribe TROGASTRINA Aboeurhexius Park, 1952c:62 Type species: Aboeurhexius crenulatus Park (orig. des.) Mexico [1 sp.] Adalmus Reitter, 1882c:210 Type species: Adalmus velutinus Reitter (sub- seq. mon. Reitter, 1885c:338) New Zealand [1 sp.] Anarmodius Raffray, 1 890a: 1 06 Type species: Trichonyx gibbus Schaufuss (monotypy) Brazil, Venezuela [2 spp.] Aporhexius Raffray, 1904a:593 Type species: Aporhexius pubescens Raffray (monotypy) Bolivia [1 sp.] Bontomtes Grigarick & Schuster, 1980:24 Type species: Bontomtes riparie Grigarick & Schuster (orig. des.) W Nearctic (Oregon) [1 sp.] Conoplectus Brendel , 1888:301 Type species: Euplectus canaliculars Le- Conte (monotypy) E Nearctic, Mexico [13 spp.] Prorhexius Raffray, 1890a: 107 Type species: Prorhexius sylvaticus Raffray (= Euplectus canaliculars LeConte) (sub- seq. mon. Raffray, 1890b: 197) Hexirhexius Grigarick & Schuster, 1980:20 Type species: Euplectus canaliculars Le- Conte (orig. des.) Euboarhexius Grigarick & Schuster, 1966:31 Type species: Euboarhexius sinus Grigarick & Schuster (orig. des.) Nearctic [4 spp.] Eurhexius Sharp, 1887:41 Type species: Eurhexius vestitus Sharp (des. Park, 1952c:74) Neotropical [30 spp.] Fletcherexius Park, 1942:72 Type species: Eurhexius macrodactylus Fletcher (orig. des.) Mexico [1 sp.] Foveoscapha Park & Wagner, 1962:13 Type species: Foveoscapha terracola Park & Wagner (orig. des.) W Nearctic (Washington) [1 sp.] Mesoplatus Raffray, 1890a: 106 Type species: Batrisus barbatus King (mono- typy) Australia, New Guinea [8 spp.] Neosampa Broun, 192 la:5 1 8 Type species: Neosampa granulata Broun (monotypy) New Zealand [1 sp.] Oropus Casey, 1886:196 Type species: Oropus convexus Casey (orig. des.) W Nearctic [34 spp.] Rhexidius Casey, 1887a:478 Type species: Rhexidius granulosus Casey (monotypy) Nearctic, Mexico [13 spp.] Rhexinia Raffray, 1890a: 106 Type species: Rhexinia angulata Raffray (sub- seq. mon. Raffray, 1890b: 196) Neotropical [3 spp.] Rhexiola Park, 1952c:57 Type species: Rhexinia mexicana Park (orig. des.) Mexico [2 spp.] Tetrascapha Schuster & Marsh, 1957:149 Type species: Tetrascapha dasycerca Schuster & Marsh (orig. des.) W Nearctic [3 spp.] Trogaster Sharp, 1874b:83 Type species: Trogaster aberrans Sharp (monotypy) Corsica, Italy [10 spp.] Heteronyx Saulcy, 1 874: 1 28 (preocc, not Guer- in-Meneville, 1838) Type species: Heteronyx aberrans Saulcy (des. Jeannel, 1950:147) Speleoplectus Minozzi, 1917:168 Type species: Plectophloeus fiorii Minozzi (monotypy) Trogasteropsis Dodero, 1918:108; 1919:176 Type species: Trogasteropsis anophthalmus Dodero (monotypy) Spain, Sardinia [5 spp.] 30 FIELDIANA: ZOOLOGY Xerhius Raffray, 1891a:305 Type species: Xerhius cordicollis Raffray (monotypy) Venezuela [1 sp.] Xherius Raffray, 1904a:587 (misspelling) Section JUBOMORPHI [of EUPLECTIN AE] Tribe JUBINI Arctophysis Reitter, 1882c:21 1 Type species: Arctophysis gigantea Reitter (subseq. mon. Reitter, 1883d:384) Colombia [1 sp.] Auxenocerus Jeannel, 1962:324 Type species: Auxenocerus bispina Jeannel (orig. des.) Chile [8 spp.] Balega Reitter, 1882c:211; 1883a:43 Type species: Balega elegans Reitter (subseq. mon. Reitter, 1883a:43) Mexico, West Indies [4 spp.] Barrojuba Park, 1942:59 Type species: Barrojuba albertae Park (orig. des.) Panama [6 spp.] Canthoderus Motschulsky, 1855:15 Type species: Canthoderus villosus Motschul- sky (monotypy) Neotropical [6 spp.] Stratus Schaufuss, 1872:246 Type species: Stratus ursinus Schaufuss (sub- seq. mon. Schaufuss, 1877b:452) Endytocera Sharp, 1887:45 Type species: Endytocera vestita Sharp (orig. des.) Panama [2 spp.] Germainites Jeannel, 1962:330 Type species: Germainites crassus Jeannel (orig. des.) Chile [1 sp.] Jubomorphus Raffray, 1891a:299 Type species: Jubomorphus simoni Raffray (monotypy) Venezuela [1 sp.] Jubus Schaufuss, 1872:247 Type species: Jubus spinicollis Schaufuss (des. Lucas, 1920:355) Neotropical [54 spp.] Gamba Schaufuss, 1872:247 (not Liais, 1872) Type species: Gamba brucki Schaufuss (des. here) Gasola Reitter, 1882a: 150 Type species: Gasola simoni Reitter (mono- typy) Duciola Reitter, 1882c:21 1 Type species: Duciola tetratoma Reitter (sub- seq. mon. Reitter, 1883d:384) Kuscheliotes Jeannel, 1962:321 Type species: Kuscheliotes rugosus Jeannel (orig. des.) Chile [5 spp.] Macta Raffray, 1890a:87 Type species: Macta constricta Raffray (sub- seq. mon. Raffray, 1890b: 193) Brazil [1 sp.] Phamisus Aube, 1844:94 Type species: Phamisus reichenbachii Aube (monotypy) Neotropical [4 spp.] Pselaphomorphus Motschulsky, 1855:15 Type species: Pselaphomorphus sculpturatus Motschulsky (monotypy) Neotropical [5 spp.] Sebaga Raffray, 1891a:300 Type species: Sebaga centralis Raffray (monotypy) Neotropical [10 spp.] Subfamily BATRISINAE Tribe AMAUROPSINI Amaurops Fairmaire, 1851:527; 1852:74 Type species: Amaurops aubei Fairmaire (monotypy) W Palearctic [9 spp.] Amauropus Reitter, 1918:202 Type species: Amauropus lobipes Reitter (monotypy) Albania [1 sp.] Arianops Brendel, 1893:278 (new name for Anops Brendel) E Nearctic [31 spp.] Anops Brendel, 1891:80 (preocc, not Oken, 1815; Bell, 1833; Boisduval, 1836; d'Or- bigny, 1836) Type species: Anops amplyoponica Brendel (monotypy) Eusanops Brendel, 1893:278 (published in syn- onymy with Arianops; unavailable, never validated) Arispeleops Park, 1951:41 Type species: Arianops caver nensis Park (orig. des.) Bergrothia Reitter, 1 884c:207 (new name {or Ami- crops Saulcy) Caucasus area [4 spp.] Amicrops Saulcy, 1879:467 (preocc, not Mon- trouzier, 1858) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 31 Type species: Amaurops saulcyi Reitter (des. Lucas, 1920:137) BergrothiellaReitter, 1897:241 (unnecessary new name for Bergrothia Reitter) Orientamaurops Karaman, 1961:158 Type species: Amaurops syriaca Reitter (monotypy) Lebanon [1 sp.] Paramaurops Jeannel, 1948a:7 Type species: Amaurops exaratus Baudi (orig. des.) S Europe [40 spp.] Amauropidius Jeannel, 1 948a: 1 5 Type species: Amaurops gallicus Delarouzee (orig. des.) Protamaurops Miiller, 1944:84 Type species: Amaurops macrophthalma Miiller (orig. des.) Balkan Peninsula, Turkey [5 spp.] Schweigeria L6b\, 1967:1 Type species: Schweigeria uludagensis Lobl (orig. des.) Pseudamaurops Jeannel, 1948a:7 Type species: Bergrothiella albanica Apfel- beck (orig. des.) Albania [2 spp.] Seracamaurops Winkler, 1925:147 Type species: Amaurops frieseni Winkler (orig. des.) Yugoslavia [7 spp.] Troglamaurops Ganglbauer, 1903:178 Type species: Amaurops leptoderina Reitter (monotypy) Yugoslavia [2 spp.] Tropidamaurops Jeannel, 1 948a: 1 3 Type species: Amaurops carinatus Baudi (orig. des.) Italy [1 sp.] Zoufalia Reitter, 1918:201 Type species: Amaurops corcyrea Reitter (des. Jeannel, 1948a:5) Greece [5 spp.] Tribe BATRISINI Subtribe AMBICOCERINA Ambicocerodes Leleup, 1970a:334 Type species: Ambicocerodes basilewskyi Le- leup (orig. des.) Afrotropical [4 spp.] Ambicoceropsis Leleup, 1971a:332 Type species: Ambicoceropsis vandeli Leleup (orig. des.) Gabon [1 sp.] Ambicocerus Leleup, 1970a:314 Type species: Ambicocerus carinaticollis Le- leup (orig. des.) Afrotropical [9 spp.] Pseudambicocerus Leleup, 1970a:337 Type species: Pseudambicocerus regressus Le- leup (orig. des.) Gabon [1 sp.] Tribe BATRISINI Subtribe BATRISINA Acanthanops Jeannel, 1952a:242 Type species: Acanthanops bambuseti Jeannel (orig. des.) Zaire [1 sp.] Acanthicomus Jeannel, 1949c:273 Type species: Acanthicomus leptus Jeannel (orig. des.) Afrotropical [6 spp.] Basilewskyola Jeannel, 1959d:421 Type species: Basilewskyola spinicollis Jean- nel (orig. des.) Acanthocliarthrus Leleup, 1970b:36 Type species: Acanthocliarthrus troglobioides Leleup (orig. des.) Gabon [1 sp.] Adiastulus Raffray, 1904b:40 Type species: Batrisus ophthalmicus Raffray (monotypy) Afrotropical [5 spp.] Anama NEW NAME (for Amana Raffray) Singapore, Sumatra [2 spp.] Amana Raffray, 1890a:l 13 (preocc, not Walk- er, 1855) Type species: Amana crassicornis Raffray (subseq. mon. Raffray, 1890b:203) Ancistromus Jeannel, 1957a: 13 Type species: Coryphomus longulus Jeannel (orig. des.) Vietnam [1 sp.] Apobatrisus Raffray, 1897a:235 Type species: Batrisus gabonicus Raffray (monotypy) Uganda [1 sp.] Apocliarthrus Jeannel, 1959d:310 Type species: Hemicliarthrus spinicollis Jean- nel (orig. des.) Afrotropical [5 spp.] Apotrabisus Jeannel, 1959d:229 Type species: Trabisus brachypterus Jeannel (orig. des.) Uganda [1 sp.] 32 FIELDIANA: ZOOLOGY Arthmius LeConte, 1849:91 Type species: Arthmius globicollis LeConte (monotypy) Nearctic, Neotropical, Chile [109 spp.] Arthromelodes Jeannel, 1954a:247 Type species: Arthromelodes cartel Jeannel (orig. des.) Vietnam, Japan [2 spp.] Arthromelus Jeannel, 1 949a: 1 49 Type species: Batrisus caudatus Raffray (orig. des.) Afrotropical, S. Africa, Oriental [110 spp.] Scaiomelus Jeannel, 195 ld:50 (subgenus) Type species: Arthromelus bredoi jeannel (orig. des.) Euthiomelus Jeannel, 1957b:73 (subgenus) Type species: Arthromelus dundoanus Jeannel (orig. des.) Atheropterus Raffray, 1882:77 Type species: Atheropterus longipalpis Raffray (monotypy) Afrotropical, South Africa [107 spp.] Trabisus Raffray, 1 890a: 1 1 4 Type species: Batrisus zanzibaricus Raffray (des. Jeannel, 1949a: 11 7) Probatrisus Raffray, 1890a:l 15 (subgenus) Type species: Batrisus sulcatus Raffray (monotypy) Atheropterodes Jeannel, 1 949c: 1 1 4 Type species: Atheropterodes devroeyi Jeannel (orig. des.) Trabisotus Jeannel, 1 949c: 127 Type species: Trabisotus brevipilis Jeannel (orig. des.) Trabisellus Jeannel, 1952a: 194 (subgenus) Type species: Protrabisus abdominalis Jean- nel (orig. des.) Trabisidius Jeannel, 1952a: 194 (subgenus) Type species: Trabisus planifrons Jeannel (orig. des.) Trabisorites Jeannel, 1952a: 194 (subgenus) Type species: Trabisus alticola Jeannel (orig. des.) Trablsonoma Jeannel, 1955d:205 (subgenus) Type species: Trablsonoma gedyei Jeannel (orig. des.) Trabisomorphus Jeannel, 1959d:229 (subgenus) Type species: Batrisus hydropicus Raffray (orig. des.) Trabisinus Jeannel, 1959d:229 (subgenus) Type species: Hypobatrisus celisi Jeannel (orig. des.) Echinotrabisus Jeannel, 1959d:229 (subgenus) Type species: Trabisus echinatus Jeannel (orig. des.) Paratrabisus Jeannel, 1959d:229 (subgenus) Type species: Trabisus trigonoderus Jeannel (orig. des.) Basitrodes Jeannel, 1958b:26 Type species: Batrisus vest It us Sharp (orig. des.) Japan, USSR [10 spp.] Batoctenus Sharp, 1887:18 Type species: Batoctenus puncticollis Sharp (des. Lucas, 1920:135) Neotropical [5 spp.] Batoxyla Raffray, 1897b:83 Type species: Batoxyla punctata Raffray (monotypy) Afrotropical [7 spp.] Batriasymmodes Park, 1951:11 Type species: Batrisus monstrosus LeConte (orig. des.) E Nearctic [14 spp.] Extollodes Park, 1965:185 (subgenus) Type species: Batriasymmodes wagneri Park (orig. des.) Speleodes Park, 1965:186 (subgenus) Type species: Batrisodes troglodytes Park (orig. des.) Batribolbus Raffray, 1904b:60 Type species: Eubatrisus palpator Raffray (des. Lucas, 1920:135) Sri Lanka [3 spp.] Batricrator Jeannel, 1957a: 1 1 Type species: Batrisus rajah Raffray (orig. des.) Oriental [3 spp.] Batrictenistes Lobl, 1976:121 Type species: Batrictenistes decorata Lobl (orig. des.) Java, Malaya [3 spp.] Batrifigia Park, 1952b:33 Type species: Batrifigia sulcata Park (orig. des.) Fiji [4 spp.] Batrinanda Park, 1952b:50 Type species: Batrinanda foveata Park (orig. des.) Fiji [1 sp.] Batriplica Raffray, 1904b:3 Type species: Batrisus dohrni Schaufuss (des. Lucas, 1920:135) Oriental [3 spp.] Batrisaulax Jeannel, 1959d:352 Type species: Syrbatus foveiventrts Raffray (orig. des.) Afrotropical [9 spp.] Batriscenaulax Jeannel, 1958b:57 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 33 Type species: Batrisus modestus Sharp (orig. des.) Japan [4 spp.] Batriscenellus Jeannel, 1958b:60 Type species: Batrisus fragilis Sharp (orig. des.) Japan [4 spp.] Scaioscenellus Jeannel, 1958b:60 (subgenus) Type species: Batrisus similis Sharp (orig. des.) Batrisceninus Jeannel, 1952b:93 Type species: Batrisocenus serraticornis Raf- fray (orig. des.) New Guinea [5 spp.] Batrisceniola Jeannel, 1958b:65 Type species: Batrisus dissimilis Sharp (orig. des.) Japan [1 sp.] Batriscenites Jeannel, 1952b:95 Type species: Batrisus morulus Reitter (orig. des.) Java [1 sp.] Batriscenodes Jeannel, 1952b:93 Type species: Batrisocenus siamensis Raffray (orig. des.) Oriental [5 spp.] Batrischema NEW NAME (for Batrisoschema Jeannel) Madagascar [1 sp.] Batrisoschema Jeannel, 1960b: 123 (preocc, not Reitter, 1883) Type species: Batrisoschema barbatum Jean- nel (orig. des.) Batrisiella Raffray, 1904b:59 Type species: Eubatrisus caviventris Raffray (monotypy) Oriental, Japan, Korea [15 spp.] Arthromelinus Jeannel, 1952b:98 Type species: Batrisus angulatus Raffray (orig. des.) Batrisinus Raffray, 1893a:476 Type species: Batrisus bouchardi Raffray (monotypy) Sumatra [4 spp.] Batrisiotes Jeannel, 1951c:90 Type species: Batrisiotes clavigeroides Jeannel (orig. des.) Afrotropical [3 spp.] Batrisocenus Raffray, 1903a:48 Type species: Batrisocenus bironis Raffray (des. Jeannel, 1949a: 149) Oriental, Japan, New Guinea, Australia [92 spp.] Batrisochaetus Jeannel, 1959d:349 Type species: Batristellus scopsi Jeannel (orig. des.) Afrotropical [7 spp.] Batrisochorus Jeannel, 1 949a: 1 46 Type species: Batrisus gracilicornis Raffray (orig. des.) Afrotropical [16 spp.] Batristellus Jeannel, 1 949c: 1 54 (subgenus) Type species: Batristellus burgeoni Jeannel (orig. des.) Batrisoconnus Leleup, 1975b:235 Type species: Batrisoconnus longiceps Leleup (orig. des.) Afrotropical [2 spp.] Batrisodella Jeannel, 1954a:243 Type species: Batrisodella puncticollis Jeannel (orig. des.) Madagascar [2 spp.] Batrisodellus Jeannel, 1958b:37 Type species: Batrisodes nipponensis Raffray (orig. des.) Japan [1 sp.] Batrisodema Raffray, 1 890a: 1 1 3 Type species: Batrisodema tuberculata Raf- fray (subseq. mon. Raffray, 1890b:200) Singapore [1 sp.] Batrisodes Reitter, 1882a: 134; 1882c:205 (see Discussion) Type species: Batrisus delaporti Aube (des. Lucas, 1920:135) Holarctic [154 spp.] Alytus Hampe, 1863:286 (see Discussion) Type species: Trichonyx adnexus Hampe (monotypy) Batrisodinus Jeannel, 1950:351 Type species: Batrisus oculatus Aube (orig. des.) Babnormodes Park, 1951:12 (subgenus) Type species: Pselaphus riparius Say (orig. des.) Pubimodes Park, 1951:12 (subgenus) Type species: Batrisus nigricans LeConte (orig. des.) Excavodes Park, 1951:12 (subgenus) Type species: Batrisus frontalis LeConte (orig. des.) Elytrodes Park, 1951:13 (not Fischer- Wal- dheim, 1829, which is a nomen nudum) (subgenus) Type species: Batrisus ionae LeConte (orig. des.) Declivodes Park, 1951:13 (subgenus) Type species: Batrisus bistriatus LeConte (orig. des.) 34 FIELDIANA: ZOOLOGY Spifemodes Park, 1953c:315 (subgenus) Type species: Batrisus schaumii Aube (orig. des.) Empinodes Park, 1953c:315 (subgenus) Type species: Batrisus albionicus Aube (orig. des.) Batrisodiola Jeannel, 1960d:414 Type species: Batrisodiola transversicollis Jeannel (orig. des.) NE India [1 sp.] Batrisodites Jeannel, 1954a:244 Type species: Batrisodites laeviceps Jeannel (orig. des.) Madagascar [19 spp.] Batrisolius Jeannel, 1956f:25 Type species: Batrisodites laevicollis Jeannel (orig. des.) Batrisomalus Raffray, 1 904b:60 Type species: Batrisus microphthalmus Raf- fray (des. Lucas, 1920:135) India, Sri Lanka [4 spp.] Batrisomellus Leleup, 1976c:446 Type species: Batrisomellus parvulus Leleup (orig. des.) Gabon [1 sp.] Batrisomicrus Jeannel, 1954e:29 Type species: Batrisomicrus acutispina Jean- nel (orig. des.) Zaire [1 sp.] Batrisomina Raffray, 1903b:316 Type species: Batrisomina strigicollis Raffray (monotypy) Madagascar [11 spp.] Batrisopachys Jeannel, 1960d:414 Type species: Batrisopachys tibialis Jeannel (orig. des.) N India [1 sp.] Batrisophyma Raffray, 1 904b:90 Type species: Batrisus granosum Raffray (monotypy) E Oriental [1 sp.] Batrisoplatus Raffray. 1894b:226 Type species: Batrisoplatus rugosulus Raffray (des. Lucas, 1920:135) Malaya [1 sp.] Batrisoplisus Raffray, 1908a: 180 Type species: Batrisus antennatus Weise (monotypy) Japan [5 spp.] Batrisoschema Reitter, 1884b:399 (not Jeannel, 1960) Type species: Batrisoschema lateridentata Reitter (monotypy) Oriental, Japan [5 spp.] Batrisoxenus Leleup, 1971c: 186 (not Dajoz, 1982) Type species: Batrisoxenus kaszabi Leleup (orig. des.) Congo-Brazzaville [1 sp.] Batristerus Jeannel, 1949a: 133 Type species: Batrisodes tricarinatus Raffray (orig. des.) Oriental, New Guinea, Australia [68 spp.] Batristhenes Jeannel, 1960d:422 Type species: Batristhenes clypeatus Jeannel (orig. des.) India [2 spp.] Batristidius Jeannel, 1949c: 169 Type species: Batristidius fossulatus Jeannel (orig. des.) Afrotropical [4 spp.] Batristilbus Raffray, 1909:22 Type species: Batrisus politus Sharp (mono- typy) Japan [2 spp.] Batristites Jeannel, 1949a: 133 Type species: Batrisodes articularis Raffray (orig. des.) Afrotropical [4 spp.] Paracliarthrus Jeannel, 1 949a: 1 44 Type species: Cliarthrus palmarum Raffray (orig. des.) Batristogenius Jeannel, 1959d:388 Type species: Batristogenius contractus Jean- nel (orig. des.) Tanzania [1 sp.] Batrisus Aube, 1833:45 (see Discussion) Type species: (see Discussion) Palearctic [4 spp.] Batrivitis Park, 1952b:42 Type species: Batrivitis facialis Park (orig. des.) Fiji [2 spp.] Batrixenus NEW NAME (for Batrisoxenus Dajoz) Madagascar [1 sp.] Batrisoxenus Dajoz, 1982:495 (preocc, not Le- leup, 1971) Type species: Batrisoxenus gracilis Dajoz (orig. des.) Besuchetidia Leleup, 1981:23 Type species: Besuchetidia antennalis Leleup (orig. des.) Ivory Coast [1 sp.] Borneana Schaufuss, 1882c: 140 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 35 Type species: Borneana biformis Schaufuss (monotypy) Borneo [1 sp.] Bothriotelus Jeannel, 195 1 f:46 Type species: Bothriotelus leleupi Jeannel (orig. des.) Afrotropical [2 spp.] Camptomodes Jeannel, 1959d:407 Type species: Coryphomus carinatus Jeannel (orig. des.) Ivory Coast [1 sp.] Catoxyomus Jeannel, 1949c: 145 Type species: Catoxyomus crenulatus Jeannel (orig. des.) Zaire [3 spp.] Celisia Jeannel, 1955a:29 Type species: Celisia longula Jeannel (orig. des.) Afrotropical [7 spp.] Ceroderma Raffray, 1890a: 112 Type species: Ceroderma asperata Raffray (subseq. mon. Raffray, 1890b: 199) Malaya [1 sp.] Clarkeorites Leleup, 1974a: 189 Type species: Clarkeorites abyssinicus Leleup (orig. des.) Afrotropical [1 sp.] Cliarthrinidius Jeannel, 1957b:51 Type species: Cliarthrinidius carvalhoi Jean- nel (orig. des.) Angola [1 sp.] Cliarthrinus Jeannel, 1 949a: 1 38 Type species: Cliarthrus elaphoides Jeannel & Paulian (orig. des.) Afrotropical [14 spp.] Cliarthrodes Jeannel, 1949a: 146 Type species: Batrisus spinicollis Raffray (orig. des.) Afrotropical [7 spp.] Cliarthromorphus Jeannel, 1 9 5 1 e: 1 34 Type species: Cliarthromorphus acuminatus Jeannel (orig. des.) Afrotropical [2 spp.] Cliarthrus Raffray, 1877:290 Type species: Cliarthrus bicolor Raffray (monotypy) Afrotropical, South Africa [3 spp.] Connodontinus Jeannel, 1 953b: 171 Type species: Connodontus vanboveni Jeannel (orig. des.) Zaire [2 spp.] Connodontus Raffray, 1882:52 Type species: Connodontus acuminatus Raf- fray (monotypy) Afrotropical [8 spp.] Connodontites Jeannel, 1959d:284 (subgenus) Type species: Connodontus scutellatus Jean- nel (monotypy) Conogastridius Jeannel, 1949c: 164 Type species: Conogastridius microcephalus Jeannel (orig. des.) Afrotropical [2 spp.] Conopygidia Jeannel, 1949c: 109 Type species: Conopygidia saegeri Jeannel (orig. des.) Zaire [1 sp.] Conuridius Jeannel, 1949c: 166 Type species: Conuridius bequaerti Jeannel (orig. des.) Zaire [4 spp.] Coryphomodes Jeannel, 1960d:416 Type species: Coryphomodes cristatus Jeannel (orig. des.) Oriental, Japan [7 spp.] Coryphomodes Jeannel, 1959d:408 (nomen nu- dum, no described species included) Coryphomoides Lobl, 1982:205 Type species: Coryphomoides osellai Lobl (orig. des.) Andaman Islands [1 sp.] Coryphomus Jeannel, 1949a: 133 Type species: Batrisodes gladiator Raffray (orig. des.) Afrotropical [37 spp.] Camptomites Jeannel, 1949b: 122 (subgenus) Type species: Camptomites termitophilus Jeannel (orig. des.) Camptomidius Jeannel, 1959d:407 (subgenus) Type species: Coryphomus basiplicatus Jean- nel (orig. des.) Coryphomellus Jeannel, 1959d:407 (subgenus) Type species: Camptomites acanthurus Jean- nel (orig. des.) Cratna Raffray, 1890a:113 Type species: Cratna torticornis Raffray (sub- seq. mon. Raffray, 1890b:202) Oriental [13 spp.] Cratnodes Jeannel, 1961:433 Type species: Cratnodes lewisi Jeannel (orig. des.) Sri Lanka [1 sp.] Cylindroma Raffray, 1897a:232 Type species: Cylindroma excavata Raffray (monotypy) Sumatra [1 sp.] Diaposis Raffray, 1904b:38 36 FIELDIANA: ZOOLOGY Type species: Batrisus carinicollis Raffray (monotypy) Sumatra [1 sp.] Diaugis Raffray, 1904b: 14 Type species: Diaugis granulosa Raffray (des. Lucas, 1920:236) E Oriental [3 spp.] Ectotrabisus Leleup, 1979:445 Type species: Probatrisus luberculaius Jean- nel (monotypy) Zaire, Ivory Coast [3 spp.] Eleodimerina Jeannel, 1960c: 139 Type species: Eleodimerina joannae Jeannel (orig. des.) Zaire [1 sp.] Eleodimerodes Leleup, 1981:47 Type species: Eleodimerodes antennalis Le- leup (orig. des.) Ivory Coast [1 sp.] Eleodimerus Jeannel, 1 949a: 1 46 Type species: Batrisus deformis Reitter (orig. des.) Afrotropical [101 spp.] Eubatrisus Raffray. 1 890a: 1 14 Type species: Batrisus pubifer Reitter (mono- typy) Borneo [1 sp.] Euphalepsus Reitter. 1883d:378 Type species: Euphalepsus globipennis Reitter (des. Park, 1942:258) Neotropical [24 spp.] Euphalespus Raffray, 1887:39 (misspelling) Exallidius Jeannel. 1957b:49 Type species: Exallidius dentaticornis Jeannel (orig. des.) Angola [1 sp.] Exallomorpha Jeannel, 1949c: 144 Type species: Exallomorpha leleupi Jeannel (orig. des.) Zaire [2 spp.] Exallus Raffray. 1 904b: 1 8 Type species: Batrisus semiopacus Raffray (monotypy) Afrotropical [7 spp.] Exechophyes Jeannel, 1 949c: 1 73 Type species: Exechophyes ghesquierei Jean- nel (orig. des.) Zaire [2 spp.] Exedrus Raffray. 1903a:47 Type species: Exedrus caviceps Raffray (monotypy) Singapore, New Guinea [2 spp.] Franzorella Leleup. 1977a:82 Type species: Franzorella trifossulata Leleup (orig. des.) Madagascar [1 sp.] Hemicliarthrus Jeannel, 1951 f:47 Type species: Hemicliarthrus puncticeps Jean- nel (orig. des.) Afrotropical [3 spp.] Arthriclius Jeannel, 1953b:207 Type species: Arthriclius parvicollis Jeannel (orig. des.) Hingstoniella Jeannel, 1960d:410 Type species: Hingstoniella lata Jeannel (orig. des.) Sikkim, Tibet [1 sp.] Hulstaertites Jeannel, 1949c: 163 Type species: Hulstaertites punctatus Jeannel (orig. des.) Zaire [1 sp.] Hyobontus NEW NAME (for Hybonotus Jeannel) Zaire [1 sp.] Hybonotus Jeannel, 1953b:245 (preocc, not Klug, 1803; Dejean, 1834) Type species: Hybonotus leleupi Jeannel (orig. des.) Hypochraeus Raffray, 1 904b: 1 3 Type species: Batrisoschema humeralis Raf- fray (des. here) Singapore, Sumatra [3 spp.] Iteticus Raffray, 1904b: 19 Type species: Batrisus princeps Reitter (des. Park, 1942:248) Neotropical, Chile [9 spp.] Jochmansiella Leleup, 1976e:307 Type species: Jochmansiella suzannae Leleup (orig. des.) Madagascar [1 sp.] Kigatrodes Jeannel, 1958b:39 Type species: Batrisus solitarius Sharp (orig. des.) Japan [2 spp.] Korovodes Park, 1952b:48 Type species: Korovodes femoralis Park (orig. des.) Fiji [1 sp.] Leleupia Jeannel, 1949c: 137 Type species: Leleupia globicephala Jeannel (orig. des.) Zaire, South Africa [4 spp.] Paraleleupia Jeannel. 1949c: 137 (subgenus) Type species: Leleupia laevipennis Jeannel (orig. des.) Leleupiastes Jeannel, 1960c: 108 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 37 Type species: Leleupiastes trigonocephalus Jeannel (orig. des.) Afrotropical [2 spp.] Leptobatrisus Jeannel, 195 lc:91 Type species: Leptobatrisus longulus Jeannel (orig. des.) Afrotropical [11 spp.] Strongylomus Jeannel, 1952a:240 (subgenus) Type species: Strongylomus punctaticollis Jeannel (orig. des.) Loebliella Leleup, 1981:27 Type species: Loebliella ambicoceroides Le- leup (orig. des.) Ivory Coast [1 sp.] Lukwangulorites Leleup, 1 976f:8 14 Type species: Lukwangulorites alticola Leleup (orig. des.) Tanzania [1 sp.] Macrodelphus Leleup, 1977a:84 Type species: Batrisodites laevissimus Jeannel (orig. des.) Madagascar [1 sp.] Madrasorites Jeannel, 1960d:416 Type species: Madrasorites bifasciculatus Jeannel (monotypy) India [1 sp.] Manniconnus Park, 1949a:253 Type species: Connodontus manni Park (orig. des.) Afrotropical [5 spp.] Megabatrus Lobl, 1979:193 Type species: Megabatrus caviceps Lobl (orig. des.) China (Fukien) [1 sp.] Microbatrisodes Jeannel, 1949b: 120 Type species: Microbatrisodes arboricola Jeannel (orig. des.) Ivory Coast [1 sp.] Microbatrisus Jeannel, 1949c: 135 (misspelling) Mnia NEW NAME (for Mina Raffray) Oriental [31 spp.] Mina Raffray, 1890a:l 12 (preocc, not Walker, 1858) Type species: Mina elegans Raffray (subseq. mon. Raffray, 1890b:200) Nenemeca Raffray, 1904b: 3 7 Type species: Batrisodema orbata Raffray (des. Lucas, 1920:434) Oriental [2 spp.] Neotrabisus Jeannel, 1 949b: 1 1 5 Type species: Neotrabisus a no m mat is Jeannel (orig. des.) Afrotropical [31 spp.] Hypotrabisus Jeannel, 1951 f:44 Type species: Protrabisus hirticollis Jeannel (orig. des.) Nesiotomina Jeannel, 1 96 1 :43 1 Type species: Batrisus spinicollis Motschulsky (orig. des.) Sri Lanka [2 spp.] Odonticoscapus Jeannel, 1956c: 123 Type species: Odonticoscapus celisi Jeannel (orig. des.) Zaire [1 sp.] Odontoconnus Jeannel, 1959d:287 Type species: Connodontus rugosiceps Jean- nel (orig. des.) Afrotropical [3 spp.] Ophelius Raffray, 1904b:36 Type species: Ophelius simplex Raffray (monotypy) Sumatra [1 sp.] Orropygia Raffray, 1910a:419 Type species: Orropygia myrmecophila Raf- fray (monotypy) South Africa [1 sp.] Oropygia Raffray, 1911:52 (misspelling) Oxarthrius Reitter, 1883d:378 Type species: Batrisus bispinosus Reitter (des. Lucas, 1920:469) Neotropical [12 spp.] Baroxarthrius Park, 1942:244 (subgenus) Type species: Oxarthrius escharus Park (monotypy) Oxyomera Raffray, 1 890a: 1 1 3 Type species: Oxyomera denticollis Raffray (subseq. mon. Raffray, 1890b:203) Singapore, Sumatra [2 spp.] Oxyomites Jeannel, 1954e:22 Type species: Oxyomites leleupi Jeannel (orig. des.) Zaire [1 sp.] Pachypygidia Jeannel, 1 9 5 1 c: 7 1 Type species: Pachypygidia alifera Jeannel (orig. des.) Angola [1 sp.] Pachytrabisus Jeannel, 1949c: 129 Type species: Pachytrabisus pubescens Jean- nel (orig. des.) Afrotropical [19 spp.] Panaphysis Reitter, 1882b: 185 Type species: Panaphysis koppi Reitter (monotypy) Afrotropical [6 spp.] Histricoidius Jeannel, 1 952a:2 1 0 Type species: Histricoidius leleupi Jeannel (orig. des.) Pantosiella Jeannel, 1958a:346 38 FIELDIANA: ZOOLOGY Type species: Pantosiella ophthalmicus Jean- nel (orig. des.) Zaire [1 sp.] Parabatrisus Jeannel, 1 949c: 1 68 Type species: Parabatrisus sulcatus Jeannel (orig. des.) Afrotropical, Aden [23 spp.] Allobatrisus Leleup, 1976a:86 Type species: Allobatrisus incisicollis Leleup (orig. des.) Passosiella Leleup. 1975b:231 Type species: Passosiella carvalhoi Leleup (orig. des.) Angola [1 sp.] Petaloscapus Jeannel, 1958b:33 Type species: Batrisus ornatus Sharp (orig. des.) Japan [4 spp.] Phalespoides Raffray. 1890a: 1 16 (justified emen- dation of Phalespoides, incorrect original spelling) Type species: Euphalepsus punctatissimus Reitter(des. Lucas, 1920:498) Brazil [4 spp.] Phalepsoides Raffray. 1904b: 104 (unjustified emendation of Phalespoides Raffray) Physomerinus Jeannel, 1952b:96 Type species: Batrisus septemfoveolatus Schaufuss (orig. des.) Oriental, Japan [9 spp.] Plagiophorus Motschulsky, 1851:496 Type species: Plagiophorus paradoxus Mot- schulsky (des. here) "Indes oriental" [2 spp.] Plocamarthrus Jeannel, 1960d:423 Type species: Plocamarthrus championi Jean- nel (orig. des.) India [4 spp.] Procheilophorus Leleup, 1981:39 Type species: Procheilophorus amplilabris Le- leup (orig. des.) Ivory Coast [1 sp.] Pseudobatriscenus Jeannel, 1958b:66 Type species: Batrisocenus dilalatus Raffray (orig. des.) ?Yunnan or Japan (doubtful loc.) [1 sp.] Pseudobatrisus Raffray, 191 3a:34 Type species: Pseudobatrisus rhinoceros Raf- fray (monotypy) Zimbabwe, Kenya [2 spp.] Pseudocliarthrus Jeannel, 1953b:205 Type species: Pseudocliarthrus laeviceps Jean- nel (orig. des.) Zaire [4 spp.] Pseudoconnus Leleup, 1971 c:200 Type species: Pseudoconnus kistneri Leleup (orig. des.) Uganda [1 sp.] Pseudoctenistes Leleup, 1975a: 182 Type species: Pseudoctenistes pertusus Leleup (orig. des.) Gabon [1 sp.] Pseudotrabisoides Jeannel, 1 953b: 1 89 Type species: Trabisoides cylindricornis Jean- nel (orig. des.) Zaire [1 sp.] Ruacorites Jeannel, 1949c: 140 Type species: Ruacorites wittei Jeannel (orig. des.) Zaire [4 spp.] Ryxabis Westwood, 1870:131 Type species: Ryxabis anthicoides Westwood (monotypy) Singapore [1 sp.] Sathytes Westwood, 1870:128 Type species: Sathytes punctiger Westwood (monotypy) Oriental [18 spp.] Batoxylina Jeannel, 1957a:8 Type species: Batoxylina clavalis Jeannel (orig. des.) Seydelites Jeannel. 195 1 f:75 Type species: Seydelites spiniventris Jeannel (orig. des.) Afrotropical [7 spp.] Siteromina Lobl, 1979:189 Type species: Siteromina dayak Lobl (orig. des.) Borneo [1 sp.] Speobatrisodes Jeannel, 1958b:41 Type species: Speobatrisodes punctaticeps Jeannel (orig. des.) Japan [2 spp.] Stenocliarthrus Leleup, 1970b:34 Type species: Arthriclius antennalis Jeannel (orig. des.) Afrotropical [2 spp.] Sulcifigia Park, 1952b:46 Type species: Sulcifigia bishopae Park (orig. des.) Fiji [1 sp.] Syrbatomorphus Jeannel, 1960c: 126 Type species: Syrbatus rostratus Jeannel (orig. des.) Afrotropical [2 spp.] Syrbatus Reitter, 1882c:205 Type species: Batrisus clypeatus Reitter (des. Jeannel, 1949a: 128) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 39 S Neotropical, Afrotropical, South Afri- ca [HOspp.] Syrbatidius Jeannel, 1952a:212 (subgenus) Type species: Syrbatus collarti Jeannel (orig. des.) Syrbatodes Jeannel, 1 959d:3 1 8 Type species: Syrbatus leleupi Jeannel (orig. des.) Syrmocerus Raffray, 1898b:459 Type species: Batrisus cervus Raffray (des. Lu- cas, 1920:623) Brazil [5 spp.] Texamaurops Barr & Steeves, 1963:1 17 Type species: Texamaurops reddelli Barr & Steeves (orig. des.) S Nearctic (Texas) [1 sp.] Trabisodema Jeannel, 1949c: 128 Type species: Trabisodema wittei Jeannel (orig. des.) Zaire [1 sp.] Trabisoides Jeannel, 1 949c: 1 29 Type species: Trabisoides burgeoni Jeannel (orig. des.) Zaire [4 spp.] Trabisostenus Jeannel, 1 949c: 1 1 8 Type species: Trabisostenus schoutedeni Jean- nel (orig. des.) Afrotropical [8 spp.] Trabisoxenus Leleup, 197 lb: 155 Type species: Trabisoxenus debeckeri Leleup (orig. des.) South Africa, Afrotropical [4 spp.] Tribasodema Jeannel, 1961:429 Type species: Batrisus armatus Raffray (orig. des.) Sri Lanka [1 sp.] Tribasodes Jeannel, 1958b:44 Type species: Batrisus longicornis Sharp (orig. des.) Japan [3 spp.] Tribasodites Jeannel, 1960d:411 Type species: Tribasodites antennalis Jeannel (orig. des.) N India, Japan [3 spp.] Trichonomorphus Raffray, 1 890a: 1 1 3 Type species: Trichonomorphus ursinus Raf- fray (subseq. mon. Raffray, 1890b:201) Malaya, Sumatra [2 spp.] Trisinarthrus Jeannel, 1960d:420 Type species: Trisinarthrus depressifrons Jeannel (orig. des.) NE India [1 sp.] Trisiniotus Jeannel, 1960d:420 Type species: Trisiniotus nodicornis Jeannel (orig. des.) India [1 sp.] Typhlobatrisus Jeannel, 1953b:254 Type species: Typhlobatrisus globulicollis Jeannel (orig. des.) Zaire [2 spp.] Typhlorites Jeannel, 1952a:238 Type species: Typhlorites leleupi Jeannel (orig. des.) Zaire [1 sp.] Xenadiastus Jeannel, 1949c: 175 Type species: Xenadiastus vanboveni Jeannel (orig. des.) Zaire [1 sp.] Xenobasilewskyia Leleup, 1976f:796 Type species: Xenobasilewskyia myrmecobia Leleup (orig. des.) Tanzania [1 sp.] Xenobatrisus Wasmann, 1918b:pl.2 Type species: Xenobatrisus myrmicariae Wasmann (monotypy) Zaire, Cameroon [1 sp.] Xenocliarthrus Jeannel, 1956c:37 Type species: Xenocliarthrus myrmicariae Jeannel (orig. des.) Zaire [1 sp.] Xenoconurus Jeannel, 1 960c: 1 1 7 Type species: Xenoconurus leleupi Jeannel (orig. des.) Zaire [2 spp.] Xenomachadoella Leleup, 1975b:239 Type species: Xenomachadoella jeanneli Le- leup (orig. des.) Angola [1 sp.] Xenopygia Jeannel, 1949c: 106 Type species: Xenopygia raignieri Jeannel (orig. des.) Zaire [3 spp.] Xenopygiella Jeannel, 1953b: 174 Type species: Xenopygiella myrmidon Jean- nel (orig. des.) Zaire [1 sp.] Tribe BATRISINI Subtribe LEUPELIINA Leupelia Jeannel, 1954e:106 Type species: Leupelia longicornis Jeannel (orig. des.) Zaire [1 sp.] 40 FIELDIANA: ZOOLOGY Tribe BATRISINI Subtribe STILIPALPINA Codonoderus Jeannel, 1956d:367 Type species: Codonoderus bicolor Jeannel (orig. des.) Zaire [1 sp.] Pelulea Jeannel, 1 954e: 1 1 9 Type species: Pelulea grandiceps Jeannel (orig. des.) Zaire [2 spp.] Peluleotes Jeannel, 1 954e: 1 20 Type species: Peluleotes trichosus Jeannel (orig. des.) Zaire [2 spp.] Stilipalpus Jeannel, 1 9 5 1 f : 7 3 Type species: Stilipalpus clavipes Jeannel (orig. des.) Zaire [2 spp.] Tribe BATRISINI, incertae sedis Batrisopsis Raffray, 1894b:264 Type species: Batrisopsis myrmecophila Raf- fray (monotypy) Malaya, Sumatra [1 sp.] Harmophorus Motschulsky, 185 1 :490 (not Schau- fuss, 1886) Type species: Harmophorus gibbioides Mot- schulsky (monotypy) "Indes oriental" (Burma?) [1 sp.] Emanuelia Blattny, 1925:204 (unnecessary new name for Harmophorus Motschulsky) Podus Raffray, 1882:52 Type species: Stictus femoralis Raffray (monotypy) New Guinea [1 sp.] Stictus Raffray, 1882:49 Type species: Stictus denticollis Raffray (des. Lucas, 1920:614) New Guinea [3 spp.] Trisinus Raffray, 1894b:265 Type species: Trisinus cratocerus Raffray (des. Lucas, 1920:657) Singapore, Sumatra [2 spp.] Xenolejeunea Leleup & Celis, 1 969a:48 Type species: Xenolejeunea wilfridi Leleup & Celis (monotypy) Zaire [1 sp.] Tribe METOPIINI Barrometopia Park, 1942:210 Type species: Barrometopia quasimoda Park (orig. des.) Panama [1 sp.] Bibrax Fletcher, 1927:149 Type species: Bibrax bradleyi Fletcher (orig. des.) Panama [1 sp.] Metopias Gory, 1832: unnumbered page Type species: Metopias curculionoides Gory (monotypy) Neotropical [4 spp.] Marnax LaPorte, 1835:137 (unnecessary new name for Metopias Gory) Metopiellus Raffray, 1908a: 188 Type species: Metopias hirtus Reitter (des. Park, 1942:205) Brazil, N Argentina [3 spp.] Metopiosoma Raffray, 1908a: 188 Type species: Metopias pacificus Westwood (monotypy) Brazil, N Argentina [2 spp.] Metopioxys Reitter, 1885c:334 Type species: Metopioxys gladiator Reitter (des. Lucas, 1920:410) S Neotropical [28 spp.] Subfamily GONIACERINAE Section BRACHYGLUTOMORPHI Tribe ARNYLLIINI Arnyllium Reitter, 1884b:391 Type species: Arnyllium pectinatum Reitter (des. Jeannel, 1952b: 101) Oriental [1 1 spp.] Tribe BRACHYGLUTINI Subtribe BARADINA Barada Raffray, 1891a:314 Type species: Barada mucronata Raffray (monotypy) Venezuela [1 sp.] Tribe BRACHYGLUTINI Subtribe BRACHYGLUTINA Acamaldes Reitter, 1882a: 140 Type species: Acamaldes bythinoides Reitter (subseq. mon. Reitter, 1882b: 191) Afrotropical [5 spp.] Achilia Reitter, 1890:212 (new name for Byraxis Raffray) Chile, S Argentina [58 spp.] Byraxis Raffray, 1890a: 123 (preocc, not Reit- ter, 1880) Type species: Bryaxis valdiviensis Reitter (= Achilia blanchardi Raffray) (des. Jean- nel, 1962:396) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 41 Achillia Raffray, 1 904b: 1 1 3 (misspelling) Clermontodes Jeannel, 1 950:3 1 7 Type species: Clermontodes insularis Jeannel (orig. des.) Achillidia Jeannel, 1962:391 Type species: Bryaxis bituberculata Reitter (orig. des.) Chile [1 sp.] Achilliotes Jeannel, 1962:445 Type species: Achilla brevicornis Raffray (orig. des.) Chile [1 sp.] Anabaxis Raffray, 1908a:252 Type species: Bryaxis lunatlca King (des. here) Australia, New Zealand [4 spp.] Anarmoxys Raffray, 1 900b: 1 66 Type species: Anarmoxys simpllclfrons Raf- fray (monotypy) Australia [1 sp.] Anasopsis Raffray, 1904b: 179 Type species: Anasls savesi Raffray (des. here) New Caledonia, Loyalty Islands [4 spp.] Asanis NEW NAME (for Anasls Raffray) Oriental [3 spp.] Anasls Raffray, 1891b:492 (preocc, not Castel- nau, 1868) Type species: Bryaxis laevicollis Raffray (monotypy) Barbiera Jeannel, 1952b:88 Type species: Barbiera frontalis Jeannel (orig. des.) Vietnam, Korea [2 spp.] Baxyridius Jeannel, 1954a:210 Type species: Baxyridius longifrons Jeannel (orig. des.) Madagascar [2 spp.] Baxyris Jeannel, 1949c:74 Type species: Baxyris burgeoni Jeannel (orig. des.) Afrotropical [26 spp.] Brachygluta Thomson, 1859:54 Type species: Pselaphus fossulatus Reichen- bach (orig. des.) Holarctic, Afrotropical, Burma [93 spp.] Nisa Casey, 1886:182 Type species: Bryaxis cavicornls Brendel (des. here) Brachyglutina Jeannel, 1950:286 Type species: Bryaxis globullcollis Mulsant & Rey (orig. des.) Dicrobiella Jeannel, 1950:315 (subgenus) Type species: Bryaxis dentiventris Saulcy (orig. des.) Neobrachygluta Kaszab & Szekessy, 1953:475 Type species: Brachygluta dudichl Kaszab & Szekessy (= Pselaphus fossulatus Rei- chenbach) (orig. des.) Brachyglutodes Jeannel, 1956a:97 Type species: Bryaxis leprieuri Saulcy (orig. des.) Dlcrobionoma Jeannel, 1956a:97 (subgenus) Type species: Bryaxis caligata Saulcy (orig. des.) Braxyda Raffray, 1904b: 145 Type species: Braxyda ha mat a Raffray (des. Lucas, 1920:149) Bolivia [2 spp.] Braxida Jeannel, 1962:388 (misspelling) Briara Reitter, 1882c:207 (new name for Gona- tocerus Schaufuss) Australia [5 spp.] Gonatocerus Schaufuss, 1880:506 (preocc, not Nees, 1834) Type species: Gonatocerus communis Schau- fuss (= Bryaxis basalis King) (mono- typy) Briaraxis Brendel, 1894:158 Type species: Briaraxis depressa Brendel (monotypy) SE Nearctic (Dry Tortugas) [1 sp.] Bryaxella Raffray, 1903b:319 Type species: Bryaxella spectralls Raffray (monotypy) Madagascar [1 sp.] Bryaxina Raffray, 1904b: 140 Type species: Bryaxis fraudatrix Schaufuss (des. Park, 1942:159) Brazil [10 spp.] Bryaxinella Jeannel, 1962:394 Type species: Bryaxinella nodlcornis Jeannel (orig. des.) Argentina (San Juan) [1 sp.] Bunoderus Raffray, 1 904b: 1 5 1 Type species: Bunoderus carinicollis Raffray (des. Lucas, 1920:152) Neotropical [18 spp.] Byraxorites Jeannel, 1962:389 Type species: Bryaxorites alticola Jeannel (orig. des.) Chile [1 sp.] Dranisaxa Park, 1945b:391 Type species: Dranisaxa gloydi Park (orig. des.) Mexico [1 sp.] Drasinus Raffray, 1904b: 148 Type species: Drasinus binodulus Raffray (monotypy) Neotropical [4 spp.] 42 FIELDIANA: ZOOLOGY Paradrasinus NEW SUBGENUS (validation of Paradrasinus Park) (subgenus) Type species: Drasinus hirsutus Fletcher (des. here) Paradrasinus Park, 1942:182 {nomen nudum, no type species designated) Ectopocerus Raffray, 1904b: 149 Type species: Decarthron verticicornis Reitter (monotypy) Chile [1 sp.] Ephymata Raffray, 1904b: 149 Type species: Bryaxis mucronata Raffray (monotypy) Singapore [1 sp.] Ergasteriocerus Leleup, 1973a:77 Type species: Ergasteriocerus spectabilis Le- leup (orig. des.) Ghana [1 sp.] Eupinella Raffray, 1908a:212 Type species: Eupinella dentiventris Raffray (monotypy) Australia [1 sp.] Eupinogitus Broun, 192 la:5 1 7 Type species: Eupinogitus sulcipennis Broun (monotypy) New Zealand [1 sp.] Eupinopsis Raffray, 1897a:261 Type species: Gerallus perforatus Schaufuss (monotypy) Australia [3 spp.] Fagniezia Jeannel, 1950:328 Type species: Pselaphus impressus Panzer (orig. des.) W Palearctic [2 spp.] Ghesquierites Jeannel, 1949c: 101 Type species: Ghesquierites spinicornis Jean- nel (orig. des.) Afrotropical [5 spp.] Leiochrotella Jeannel, 1954a:231 Type species: Leiochrotella vadoni Jeannel (orig. des.) Madagascar [3 spp.] Madabaxyris Dajoz, 1982:501 Type species: Madabaxyris difformis Dajoz (orig. des.) Madagascar [1 sp.] Malleecola Oke, 1926:281; 1928:15 Type species: Malleecola myrmecophila Oke (monotypy; orig. des.) Australia [1 sp.] Metaxis Motschulsky, 1851:490 Type species: Metaxis corpulentus Motschul- sky (des. Lucas, 1920:409) "Indes oriental," Sri Lanka, Panama [4 spp.] Nisaxis Casey, 1886:183 Type species: Bryaxis tomentosa Aube (monotypy) Nearctic [4 spp.] Noduliceps Jeannel, 1958b: 104 Type species: Bryaxis latifrons Sharp (orig. des.) Japan [1 sp.] Nondulia NEW NAME (for Nodulina Raffray) Surinam, Brazil [1 sp.] Nodulina Raffray, 1904b: 153 (preocc, not Rhumbler, 1895) Type species: Bryaxis convexa Schaufuss (monotypy) Panabachia Park, 1942:153 Type species: Bryaxis vulnerata Sharp (orig. des.) Neotropical [2 spp.] Pedisinops NEW NAME (for Pedinopsis Raffray) New Guinea [1 sp.] Pedinopsis Raffray, 1890a: 102 (preocc, not Cotteau, 1863) Type species: Euplectus major Raffray (monotypy) Physoplectus Reitter, 1882c:210 Type species: Euplectus armipes Raffray (sub- seq. mon. Reitter, 1885c:338) New Guinea, New Caledonia [2 spp.] Plectrobythus Leleup & Celis, 1969b: 79 Type species: Plectrobythus antennatus Le- leup & Celis (monotypy) Ivory Coast [1 sp.] Pseudachillia Jeannel, 1963:367 Type species: Pseudachillia bicolor Jeannel (orig. des.) Chile [2 spp.] Pseudocamaldes Jeannel, 1955a:36 Type species: Pseudocamaldes Jletcheri Jean- nel (orig. des.) Uganda [1 sp.] Rabyxis Raffray, 1 890a: 1 24 Type species: Bryaxis striolata Raffray (sub- seq. mon. Raffray, 1890b:205) Madagascar [26 spp.] Pseudobaxyris Jeannel, 1 960b: 1 29 (subgenus) Type species: Bryaxis labiata Raffray (orig. des.) Raxybis Raffray, 1908a:222; 1908d:69 Type species: Raxybis nodosa Raffray (des. Lucas, 1920:565) Argentina (Buenos Aires) [2 spp.] Reichenbachella Jeannel, 1949c:80 Type species: Bryaxis punctulata Raffray (orig. des.) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 43 Afrotropical, South Africa, Oriental, Paki- stan [19 spp.] Reichenbachia Leach, 1826:451 Type species: Bryaxis juncorum Leach (des. Bowman, 1934:144) Worldwide (except Chile) [340 spp.] Dierobia Thomson, 1859:54 Type species: Bryaxis juncorum Leach (orig. des.) Dierobia Reitter, 1881:474 (misspelling of Di- erobia) Reichenbachius Casey, 1906:359 (unnecessary new name for Reichenbachia Leach) Rougemontiella Leleup, 1974a:202 Type species: Rougemontiella cavaticeps Le- leup (orig. des.) Ethiopia [1 sp.] Rybaxidia Jeannel, 1962:392 Type species: Rybaxidia kuscheli Jeannel (orig. des.) Chile [2 spp.] Rybaxis Saulcy, 1876:96 Type species: Bryaxis sanguinea Leach (des. Bowman, 1934:144; see Discussion) Holarctic, Afrotropical, South Africa, Ori- ental, Australia, Fiji [88 spp.] Bryaxis Leach, 1817:85 (Rejected Name No. 1953, ICZN, 1969:133) Simkinion Park & Pearce, 1962:251 Type species: Simkinion prelaticum Park & Pearce (orig. des.) New Zealand [2 spp.] Speobaxyris Jeannel, 1949c:74 Type species: Baxyris troglobius Jeannel (orig. des.) Zaire [3 spp.] Startes Broun, 1886:829 Type species: Startes sculpturata Broun (monotypy) New Zealand [2 spp.] Strombopsis Raffray, 1904b: 133 Type species: Strombopsis breviventris Raf- fray (monotypy) Brazil [1 sp.] Tremissus Besuchet, 1982:317 Type species: Tremissus inexspectatus Be- suchet (orig. des.) Turkey, Israel [2 spp.] Tribatus Motschulsky, 1851:484 Type species: Batrisus thoracicus Motschul- sky (monotypy) Palearctic [5 spp.] Abatrisops Reitter, 1881:518 Type species: Batrisus thoracicus Motschul- sky (monotypy) Trissemus Jeannel, 1949a:95 Type species: Bryaxis antennata Aube (orig. des.) Afrotropical, South Africa, Palearctic, Ori- ental [130 spp.] Corynecerus Jeannel, 1 949a: 111 (subgenus) Type species: Reichenbachia nigricornis Raf- fray (orig. des.) Trissemodes Jeannel, 1949c:84 (subgenus) Type species: Trissemus burgeoni Jeannel (orig. des.) Trissemidius Jeannel, 1952a: 182 (subgenus) Type species: Trissemus apterus Jeannel (orig. des.) Apotrissemus Jeannel, 1954a:211 (subgenus) Type species: Trissemus cavifrons Jeannel (orig. des.) Trissemites Jeannel, 1959d:529 (subgenus) Type species: Bryaxis abyssinica Raffray (orig. des.) Trissemellus Jeannel, 1959d:529 (subgenus) Type species: Reichenbachia obtusa Raffray (orig. des.) Trissemosus Jeannel, 1959d:529 (subgenus) Type species: Bryaxis rupestris Raffray (orig. des.) Xiphobythus Jeannel, 1951c: 5 9 Type species: Xiphobythus machadoi Jeannel (orig. des.) Afrotropical [3 spp.] Tribe BRACHYGLUTINI Subtribe DECARTHRONINA Decarthron Brendel, 1865:30 Type species: Bryaxis formiceti LeConte (des. Bowman, 1934:144) Nearctic, Neotropical [110 spp.] Decarfuss NEW SUBGENUS (validation of De- carfuss Park) (subgenus) Type species: Bryaxis monoceros Schaufuss (des. here) Decarfuss Park, 1942:194 (nomen nudum, no type species designated) Euteleia Raffray, 1 904b: 1 83 Type species: Bryaxis recens Schaufuss (des. Park, 1942:201) Neotropical [3 spp.] Raffrayolus Jakobson, 1914:528 (new name for Itamus Raffray) Brazil [1 sp.] 44 FIELDIANA: ZOOLOGY Itamus Raffray, 1904b: 196 (preocc, not Schmidt-Gobel, 1846; Loew, 1849; Foerster, 1868) Type species: Itamus laticeps Raffray (mono- typy) Tribe BRACHYGLUTINI Subtribe EUPSENIINA Eupsenina Raffray, 1909:35 Type species: Eupsenina fracticornis Raffray (monotypy) Neotropical [3 spp.] Eupsenius LeConte. 1849:90 Type species: Eupsenius glaber LeConte (monotypy) E Nearctic, Neotropical [ 1 1 spp.] Euplenius Motschulsky, 1855: 16 (apparent mis- spelling of Eupsenius) Euprenius Raffray, 1897a:262 (misspelling) Tribe BRACHYGLUTINI Subtribe HALORABYXINA Halorabyxis Jeannel, 1954a:338 Type species: Halorabyxis vinsoni Jeannel (orig. des.) Mauritius, Reunion, Somalia, Society Is- lands [4 spp.] Tribe BRACHYGLUTINI Subtribe PSELAPTINA Anasidius Jeannel, 1952c: 139 Type species: Anasidius laevicollis Jeannel (orig. des.) Java [1 sp.] Anchylarthron Brendel, 1887:208 Type species: Decarthron cornutum Brendel (monotypy) E Nearctic, Mexico [4 spp.] Verticinotus Brendel, 1 888:300 (unnecessary new name for Anchylarthron Brendel) Baraxina Raffray. 1896a:301 Type species: Baraxina francoisi Raffray (monotypy) New Caledonia [1 sp.] Berdura Reitter, 1882c:206; 1883a:35 Type species: Berdura excisula Reitter (sub- seq. mon. Reitter, 1883a:36) Panama, West Indies [3 spp.] Berlara Reitter, 1882c:206; 1883c:286 Type species: Berlara crassipalpis Reitter (subseq. mon. Reitter, 1883c:287) Java [1 sp.] Br yaxonoma Raffray, 1898d:401 Type species: Bryaxonoma filiceum Raffray (monotypy) South Africa [2 spp.] Bythinogaster Schaufuss, 1887:111 Type species: Bythinogaster simplex Schau- fuss (monotypy) West Indies [5 spp.] Caligocara Park, 1945b:397 Type species: Caligocara carinifera Park (orig. des.) Mexico [2 spp.] Cryptorhinula Schaufuss, 1887:150, 163 Type species: Bryaxis nodifera Schaufuss (monotypy) Neotropical [7 spp.] Cryphorhinula Schaufuss, 1887:150 (misspell- ing, first reviser Raffray, 1 890a: 1 24) Dicrobiotus Jeannel, 1953b: 160 Type species: Dicrobiotus glaber Jeannel (orig. des.) Angola, Zaire [1 sp.] Eremomus Raffray, 1904b: 177 Type species: Eremomus obesus Raffray (des. Lucas, 1920:272) Bolivia, Brazil [2 spp.] Eupifigia Park, 1952b: 18 Type species: Eupifigia zimmermani Park (orig. des.) Fiji [6 spp.] Eupines King, 1866:309 Type species: Bryaxis clavatula King (des. Jeannel, 1952b:84) Oriental, New Guinea, Australia, New Zea- land, New Caledonia [130 spp.] Byraxis Reitter, 1880a: 166 (subgenus) Type species: Byraxis monstrosa Reitter (monotypy) Patranus Raffray, 1890a: 123 Type species: Tychus politus Schaufuss (= Bryaxis globulifer Schaufuss) (mono- typy) Brabaxis Raffray. 1890a: 125 Type species: Bryaxis siamensis Schaufuss (= Bryaxis sphaerica Motschulsky) (des. here) Eupinidius Jeannel, 1952b:82 Type species: Eupinidius barbieri Jeannel (orig. des.) Vietnam [1 sp.] Eupinoda Raffray, 1900b: 168 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 45 Type species: Eupinoda leana Raffray (des. Lucas, 1920:284) Australia [4 spp.] Eupinolus Oke, 1928:11 Type species: Eupinolus lucifugus Oke (orig. des.) Australia [4 spp.] Eutrichites LeConte, 1880:184 Type species: Eutrichites zimmermanni LeConte (monotypy) E Nearctic [1 sp.] Gastrobothrus Broun, 1882:287; 1886:758 Type species: Bryaxis abdominalis Broun (monotypy) New Zealand [3 spp.] Physa Raffray, 1890a: 122 (preocc, not Drapar- naud, 1801) Type species: Bryaxis inflata Sharp (mono- typy) Physobryaxis Hetschko, 1913:182 (new name for Physa Raffray) Achiraffraya Navas, 1925:29 (unnecessary new name for Physa Raffray) Leiochrotidius Jeannel, 1 960b: 1 6 1 Type species: Leiochrotidius brunneus Jean- nel (orig. des.) Madagascar [1 sp.] Leptachillia Jeannel, 1962:446 Type species: Leptachillia laevissima Jeannel (orig. des.) Chile [1 sp.] Leptorrachis Jeannel, 1961:445 Type species: Leptorrachis nigra Jeannel (orig. des.) Sri Lanka [1 sp.] Mitona Raffray, 1904b: 129 Type species: Mitona quadraticeps Raffray (des. Lucas, 1920:420) Neotropical [5 spp.] Pselaptus LeConte, 1880:184 Type species: Pselaptus belfragei LeConte (monotypy) Neotropical, SW Nearctic [10 spp.] Scalenarthrus LeConte, 1880:185 Type species: Scalenarthrus hornii LeConte (monotypy) SW Nearctic, Neotropical [26 spp.] Cylindrembolus Schaufuss, 1887:103 Type species: Cylindrembolus marginalis Schaufuss (monotypy) Abryxis Raffray, 1890a: 124 Type species: Bryaxis denticornis Schaufuss (des. here) Silillicus Lucas, 1 920:504 (new name for Phoberus Raffray) Bolivia [1 sp.] Phoberus Raffray, 1904b: 176 (preocc, not MacLeay, 1819;Kirsch, 1873; Milne-Ed- wards, 1881) Type species: Phoberus armatus Raffray (monotypy) Xenobryaxis Jeannel, 1954a:234 Type species: Xenobryaxis pauliani Jeannel (orig. des.) Madagascar [1 sp.] Xybarida Raffray, 1897a:260 Type species: Xybarida clavata Raffray (monotypy) Neotropical [4 spp.] Xybaris Reitter, 1882a: 140 Type species: Xybaris spiniceps Reitter (des. Lucas, 1920:674) Neotropical [8 spp.] Tribe BRACHYGLUTINI, incertae sedis Arachis Raffray, 1890a: 121 Type species: Batrisomorpha crassicornis Raf- fray (monotypy) New Guinea [1 sp.] Atenisodus Raffray, 1904b: 121 Type species: Comatopselaphus longicornis Raffray (des. Lucas, 1920:125) E Oriental [3 spp.] Batraxis Reitter, 1881:464 Type species: Batraxis hampei Reitter (mono- typy) Greece, Oriental, New Guinea, Australia, Seychelles [43 spp.] Batrisomorpha Raffray, 1882:38 Type species: Bryaxis armitagei King (des. here) Batraxys Raffray, 1900b: 147 (misspelling) Raffrayella Blattny, 1925:204 Type species: Raffrayella raffrayana Blattny (monotypy) Berlaraxis Jeannel, 1957a: 19 Type species: Berlaraxis coomani Jeannel (orig. des.) Vietnam [2 spp.] Comatopselaphus Schaufuss, 1882b:368 Type species: Comatopselaphus opacicollis Schaufuss (monotypy) E Oriental, New Guinea [7 spp.] Diroptrus Motschulsky, 1858:28 Type species: Diroptrus ceylonicus Motschul- sky (monotypy) Sri Lanka, India [4 spp.] 46 FIELDIANA: ZOOLOGY Obricala Raffray, 1890a: 121 Type species: Batrisomorpha foveicollis Raf- fray (monotypy) Java [1 sp.] Tetrasemus Jeannel, 1962:386 Type species: Tetrasemus delamarei Jeannel (orig. des.) Argentina (Tucuman) [1 sp.] Section BYTHINOMORPHI [of GONIACERI- NAE] Tribe BYTHININI Subtribe BYTHININA Bryaxis, Kugelann, 1794:580 Type species: Pselaphus bulbifer Reichenbach (des. ICZN, 1969:133) Palearctic [243 spp.] Areopagus Leach, 1 8 1 7:83 (Rejected Name No. 1954, ICZN, 1969:133) Kunzea Leach, 1826:448 Type species: Kunzea nigriceps Leach (mono- typy) Picrogyne Gozis, 1 886: 1 6 Type species: Bythinus pandellei Saulcy (orig. des.) Bythobletus Reitter, 1909:214 Type species: Areopagus puncticollis Denny (des. Jeannel, 1950:232) Megalobythus Jeannel, 1922:232 Type species: Megalobythus goliath Jeannel (monotypy) Embolobythus Jeannel, 1950:232 Type species: Bythinus curtisi Leach (orig. des.) Bythinites Jeannel, 1950:232 Type species: Bythinus latebrosus Reitter (orig. des.) Bythinidius Jeannel, 1950:232 Type species: Bythinus crassicornis of authors (= Bythinus glabricollis Schmidt-G6b- el, teste C. Besuchet) (orig. des.) Pyreneobythus Jeannel, 1950:232 Type species: Bythinus pandellei Saulcy (orig. des.) Bythinomorphus Jeannel, 1950:275 Type species: Bythinus fagei Jeannel (orig. des.) Parabythinus Jeannel, 1950:220 Type species: Bythinus pyrenaeus Saulcy (orig. des.) Balcanobythus Karaman, 1954b:83 (nomen nu- dum, no type species designated) Argiobythus Karaman, 1957:173 Type species: Bythinus peloponnesius Reitter (orig. des.) Balcanobythus Karaman, 1957:170 Type species: Bythinus lokayi Machulka (orig. des.) Illyrobythus Karaman, 1957:172 Type species: Bythinus longulus Kiesenwetter (monotypy) Plattibythus Karaman, 1957:173 Type species: Bythinus verrucipalpus Apfel- beck (orig. des.) Parabythus Karaman, 1957:174 (misspelling of Parabythinus) Iberobythus Franz, 1958:123 Type species: Bryaxis crotchi Sharp (orig. des.) Arcobythus Jeannel, 1958b:85 (subgenus) Type species: Bryaxis frontalis Jeannel (orig. des.) Bythiniama Jeannel, 1958b:85 (subgenus) Type species: Bryaxis japonieus Sharp (orig. des.) Bythinopsidius Meggiolaro, 1 960:60 Type species: Bythinus pentagonoceras Stolz (monotypy) Bythinus Leach, 1817:82 Type species: Pselaphus securiger Reichen- bach (des. Westwood, 1838:21) W Palearctic [25 spp.] Bolbobythus Raffray, 1 904b:289 (Rejected Name No. 1955, ICZN, 1969:133) Glyphobythus Raffray, 1904b:274 Type species: Bythinus maritimus Reitter (des. Jeannel, 1950:206) France, Italy [8 spp.] Glyphobythoides Jeannel, 1950:206 Type species: Bythinus gracilipes Saint-Claire Deville (orig. des.) Linderia Saulcy, 1863:81 Type species: Machaerites mariae Jacquelin du Val (monotypy) France, Spain [12 spp.] Macrobythus Raffray, 1908a:280 Type species: Machaerites armatus Schaufuss (des. Jeannel, 1950:190) Lophobythus Raffray, 1908a:287 Type species: Machaerites cristatus Saulcy (des. Jeannel, 1950:199) Machaerodes Brendel, 1890:252 Type species: Machaerodes carinatus Brendel (des. Lucas, 1920:386) E Nearctic [1 sp.] Typhlobythus Jeannel, 1926:285 Type species: Machaerites breuili Jeannel (orig. des.) Spain [1 sp.] NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 47 Tribe BYTHININI Subtribe MACHAERITINA Antrobythus Besuchet, 1985:511 Type species: Antrobythus lecLrci Besuchet (orig. des.) France [1 sp.] Bathybythus Besuchet, 1974b:41 Type species: Bathybythus bleyi Besuchet (orig. des.) Spain [1 sp.] Bythoxenites Jeannel, 1958b:83 Type species: Machaerites japonica Yoshida & Nomura (orig. des.) Japan [1 sp.] Bythoxenus Motschulsky, 1859:132 Type species: Bythoxenus subterraneus Mot- schulsky (monotypy) Yugoslavia [1 sp.] Facetus Schaufuss, 1872:246 Type species: Machaerites plicatulus Schau- fuss (subseq. mon. Schaufuss, 1877b: 454) Ceratobythus Normand, 1932:15 Type species: Bythoxenus monoceros Nor- mand (monotypy) Algeria [1 sp.] Decatocerus Saulcy, 1870:89 Type species: Decatocerus alhambrae Saulcy (monotypy) S Europe [6 spp.] Leptobythus Jeannel, 1955e:27 Type species: Leptobythus palaui Jeannel (orig. des.) Balearic Islands (Majorca) [1 sp.] Machaerites Miller, 1855:509 Type species: Machaerites spelaeus Miller (monotypy) Yugoslavia [2 spp.] Prionobythus Jeannel, 1921:521 Type species: Prionobythus bolivar i Jeannel (orig. des.) France, Spain [3 spp.] Spelaeobythus Lobl, 1965:589 Type species: Spelaeobythus regulis Lobl (orig. des.) Greece [1 sp.] Speleochus Park, 1951:46 Type species: Machaerites stygicus Park (orig. des.) SE Nearctic (Alabama) [3 spp.] Subterrochus Park, 1960b:91 Type species: Machaerites ferus Park (orig. des.) SE Nearctic (Alabama) [3 spp.] Tychobythinus Ganglbauer, 1896:170 Type species: Bythinus ononis Ganglbauer (des. Besuchet, 1962:348) Holarctic [86 spp.] Trichobythus Dodero, 1900:408 Type species: Bythinus anophthalmus Dodero (monotypy) Eccoptobythus Sainte-Claire Deville, 1901:271 Type species: Bythinus paradoxus Sainte- Claire Deville (monotypy) Tychobythus Raffray, 1904b:255 (misspelling of Tychobythinus) Bythinopsis Raffray, 1908a:282 Type species: Bythinus myrmido Reitter (des. Bowman, 1934:144) Apobythus Raffray, 1908a:285 Type species: Bythinus gladiator Reitter (des. Lucas, 1920:112) Trogobythus Dodero, 1919:210 Type species: Bythinus gracilicornis Raffray (des. Jeannel, 1950:171) Acanthobythus Normand, 1930:163 Type species: Bythoxenus araneipes Nor- mand (monotypy) Collartia Jeannel, 1948b:33 Type species: Collartia belgica Jeannel (monotypy) A maurobythus Jeannel, 1950:180 Type species: Bythinus revelierei Reitter (orig. des.) Crenobythus Karaman, 1954a: 183 Type species: Bythinus neumanni Miiller (monotypy) Microbythus Karaman, 1954a: 190 Type species: Bythinus apfelbecki Ganglbauer (monotypy) Odontobythus Karaman, 1954a: 185 Type species: Odontobythus ognjevae Kara- man (monotypy) Anopsibythus Jeannel, 1956a:60 Type species: Machaerites theryi Guillebeau (orig. des.) Chiasmatobythus Jeannel, 1956a:67 Type species: Bythinus algiricus Raffray (orig. des.) Globobythus Karaman, 1957:176 Type species: Bythinus latifrons Miiller (orig. des.) Globobythus Karaman, 1954a: 180 (nomen nu- dum, no type species designated) Tribe BYTHININI Subtribe XENOBYTHINA Pselaptrichus Brendel, 1889:194 48 FIELDIANA: ZOOLOGY Type species: Pselaptrichus tuberculipalpus Brendel (monotypy) W Nearctic [37 spp.] Vestitrichus Park, 1953a:259 (subgenus) Type species: Pselaptrichus curiosus Park (orig. des.) Xenobythus Peyerimhoff, 1901:203 Type species: Bythinus serullazi Peyerimhoff (monotypy) France [1 sp.] Tribe GONIACERINI Adrocerus Raffray, 1890a: 133 Type species: Adrocerus cavicornis Raffray (subseq. mon. Raffray, 1890b:208) Brazil [1 sp.] Basilewskydiella Leleup, 1978a:322 Type species: Basilewskydiella jeanneli Le- leup (orig. des.) Zaire [1 sp.] Bredoella Jeannel, 1949c:71 Type species: Bredoella gracilis Jeannel (orig. des.) Zaire [1 sp.] Debeckeria Leleup, 1976f:828 Type species: Debeckeria morogoroensis Le- leup (orig. des.) Tanzania [1 sp.] Goniaceroides Raffray, 1918:483 Type species: Goniaceroides umbilicatus Raf- fray (monotypy) Paraguay [1 sp.] Goniacerus Motschulsky, 1855:17 Type species: Goniacerus gibbus Motschulsky (monotypy) Neotropical [4 spp.] Metopioides Schaufuss, 1872:245 Type species: Metopioides setifer Schaufuss (subseq. mon. Schaufuss, 1877b:451) Goniastes Westwood, 1870:125 Type species: Goniastes sulcifrons Westwood (monotypy) Neotropical [8 spp.] Heptameron Comellini, 1979:687 Type species: Heptameron reticulatum Com- ellini (orig. des.) Costa Rica [1 sp.] Ipsimodes Jeannel, 1 9 5 1 f : 7 1 Type species: Ipsimodes joannae Jeannel (orig. des.) Afrotropical [3 spp.] Ipsimus Reitter, 1885b:228 (new name for Simus Raffray) Eritrea, Zaire [2 spp.] Simus Raffray, 1882:6 (preocc, not Bonaparte, 1838; Hodgson, 1841) Type species: Simus fracticornis Raffray (monotypy) Kistneriella Leleup, 197 lb: 161 Type species: Kistneriella termitobia Leleup (orig. des.) South Africa [1 sp.] Leleupites Jeannel, 1949c:69 Type species: Leleupites termitophilus Jeannel (orig. des.) Zaire [1 sp.] Listriophorus Schaufuss, 1872:245 Type species: Listriophorus felix Schaufuss (subseq. mon. Schaufuss, 1874:289) Neotropical [8 spp.] Microgmocerus Leleup, 1974a: 198 Type species: Microgmocerus grassei Leleup (orig. des.) Gabon [1 sp.] Notogmocerus Leleup, 1978a:335 Type species: Ogmocerus rugosus Raffray (monotypy) South Africa [1 sp.] Ogmoceridius Leleup, 1982:200 Type species: Ogmoceridius usambarensis Le- leup (orig. des.) Tanzania [1 sp.] Ogmocerodes Jeannel, 1949c:65 Type species: Ogmocerus agymsibanus Raf- fray (orig. des.) Afrotropical [13 spp.] Ogmoceropsis Jeannel, 1960c: 156 Type species: Ogmocerodes leleupi Jeannel (orig. des.) Afrotropical [1 sp.] Ogmocerus Raffray, 1882:7 Type species: Ogmocerus giganteus Raffray (monotypy) Afrotropical, South Africa [ 1 2 spp.] Paragoniastes Comellini, 1979:681 Type species: Paragoniastes raffrayi Comel- lini (orig. des.) Brazil [4 spp.] Parasimus Jeannel, 1949b: 114 Type species: Parasimus pachycerus Jeannel (monotypy) Afrotropical [4 spp.] Vadonites Jeannel, 1953a:77 Type species: Vadonites camerunensis Jean- nel (orig. des.) Cameroon, Ivory Coast [1 sp.] Xenogmocerodes Leleup, 1978a:335 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 49 Type species: Ogmocerus raffrayanus Brauns (monotypy) Zimbabwe [1 sp.] Tribe PYGOXYINI Pygoxyon Reitter, 1880c:508 Type species: Pygoxyon scydmaeniforme Reitter (monotypy) W Palearctic [ 1 7 spp.] Tribe TANYPLEURINI Subtribe DALMODINA Dalmodes Reitter, 1882c:210 (see Discussion) Type species: Dalmodes rybaxides Reitter (des. Lucas, 1920:225) Neotropical [10 spp.] Batrisobryaxis Schaufuss, 1887:148 Type species: Batribraxis labialis Schaufuss (monotypy) Dalmonexus Park, 1942:272 Type species: Dalmonexus seeversi Park (monotypy) Panama, Tobago [2 spp.] Euharmophola Park, 1960a:23 Type species: Euharmophola carinata Park (orig. des.) Jamaica [1 sp.] Harmomima Raffray, 1904b:269 Type species: Harmomima grandiceps Raf- fray (des. Lucas, 1920:316) Bolivia [2 spp.] Harmophola Raffray, 1896b: 130 Type species: Harmophola clavata Raffray (monotypy) Brazil [1 sp.] Insulomodes Park, 1976:13 Type species: Insulomodes tobagoensis Park (orig. des.) Tobago [2 spp.] Rhamophorus NEW NAME (for Harmophorus Schaufuss) Brazil [1 sp.] Harmophorus Schaufuss, 1 886:264 (preocc, not Motschulsky, 1851) Type species: Harmophorus manticoroides Schaufuss (monotypy) Tribe TANYPLEURINI Subtribe GLOBINA Globa Raffray, 1887:37 Type species: Globa longipes Raffray (mono- typy) Neotropical [3 spp.] Globosulus Jeannel, 1949c: 5 7 Type species: Globosulus cornutus Jeannel (orig. des.) Afrotropical [50 spp.] Globulites Jeannel, 1954e:96 (preocc, not de Haan, 1825) Type species: Globulites sulcifrons Jeannel (orig. des.) Tribe TANYPLEURINI Subtribe INIOCYPHINA Iniocyphus Raffray, 1912:435 Type species: Iniocyphus jheringi Raffray (monotypy) Brazil [1 sp.] Tribe TANYPLEURINI Subtribe TANYPLEURINA Acrocomus Raffray, 1894a:460 Type species: Acrocomus cribratus Raffray (monotypy) Sri Lanka [1 sp.] Anoplobraxis Park, 1942:277 Type species: Anoplobraxis guianensis Park (monotypy) Guyana [1 sp.] Apoplectus Raffray, 1890a: 102 Type species: Euplectus crassus Raffray (monotypy) New Guinea, Seychelles, Philippines [7 spp.] Glapharsenus Raffray, 1903a:86 Type species: Glapharsenus asperatus Raffray (des. here) Auchenotropidius Jeannel, 1 952a: 1 75 Type species: Auchenotropidius tuberculifer Jeannel (orig. des.) Zaire [1 sp.] Auchenotropis Raffray, 1913a:49 Type species: Auchenotropis dentimana Raf- fray (monotypy) Afrotropical [33 spp.] Batriphysis Park, 1942:270 Type species: Batriphysis epicranis Park (monotypy) Panama [1 sp.] Batrybraxis Reitter, 1882a: 141 Type species: Batrybraxis fortis Reitter (des. Lucas, 1920:136) Neotropical [7 spp.] Batribraxis Schaufuss, 1887:147 (misspelling) Buris Fletcher, 1928:227 Type species: Buris brunneus Fletcher (orig. des.) Mexico, Venezuela, West Indies [4 spp.] 50 FIELDIANA: ZOOLOGY Bythinophysis Raffray, 1908a:266 Type species: Bythinophysis punctipennis Raf- fray (monotypy) French Guiana, Guadeloupe [3 spp.] Bythiotes NEW NAME (for Bythonesiotes Jean- nel, 1958) Japan [2 spp.] Bythonesiotes Jeannel, 1958b:75 (preocc, not Jeannel, 1956) Type species: Bythonesiotes coiffaiti Jeannel (orig. des.) Bythonesiotes Jeannel, 1956b:280 (not Jeannel, 1958) Type species: Bythonesiotes vinsoni Jeannel (orig. des.) Mauritius [2 spp.] Capnites Raffray, 1898c:245 Type species: Capnites angustus Raffray (monotypy) Madagascar [1 sp.] Dalmoburis Park, 1942:266 Type species: Dalmoburis petrunfcevitchii Park (orig. des.) Panama [1 sp.] Dalmomima Raffray, 1908a:270 Type species: Batrybraxis caviceps Raffray (monotypy) Brazil [1 sp.] Dalmophysis Raffray, 1897a:267 Type species: Dalmophysis cylindrica Raffray (monotypy) Mexico [1 sp.] Gnesion Raffray, 1900b: 192 Type species: Gnesion rufulum Raffray (monotypy) Australia [1 sp.] Leleupiella Jeannel, 1 952a: 1 74 Type species: Leleupiella hirsuta Jeannel (orig. des.) Zaire, Zimbabwe [1 sp.] Machulkaia I obi. 1964a:l Type species: Machulkaia mirabilis Lobl (orig. des.) E China [1 sp.] Maya Blattny, 1925:209 Type species: Tychus dilatatus Motschulsky (des. here) Burma, Sri Lanka [2 spp.] Morana Sharp, 1 874a: 1 1 7 Type species: Morana discedens Sharp (monotypy) Japan [1 sp.] Nedarassus Raffray, 1895a:44 Type species: Nedarassus punctatus Raffray (monotypy) Malaya [1 sp.] Nipponobythus Jeannel, 1958b:77 Type species: Nipponobythus syrbatoides Jeannel (orig. des.) Japan, China [1 1 spp.] Phybytharsis Park, 1942:274 Type species: Phybytharsis gambosis Park (monotypy) Panama [1 sp.] Prosthecarthron Raffray, 19 14a: 2 Type species: Prosthecarthron sauteri Raffray (monotypy) Taiwan [1 sp.] Rugegius Jeannel, 1952a: 173 Type species: Rugegius leleupi Jeannel (orig. des.) Zaire [1 sp.] Sogaella Jeannel, 1960b: 125 Type species: Sogaella tanypleuroides Jeannel (orig. des.) Madagascar [5 spp.] Leiochrotina Jeannel, 1959e:208 (preocc, not Westwood, 1883) Type species: Leiochrotina nodulicornis Jean- nel (orig. des.) Sunorfa Raffray, 1882:28 Type species: Sunorfa capitata Raffray (monotypy) Oriental, New Guinea, Fiji, Santa Cruz Is- lands, Seychelles [25 spp.] Bythinomorpha Schaufuss, 1887:109 Type species: Bythinomorpha exsculpta Schaufuss (monotypy) Sunorfoides Raffray, 1 9 1 3b: 1 34 (subgenus) Type species: Sunorfa nigripennis Raffray (des. Lucas, 1920:620) Takaorites Jeannel, 1958b:73 Type species: Takaorites torticornis Jeannel (orig. des.) Japan [1 sp.] Tanypleurus Raffray, 1 890a: 1 30 Type species: Tanypleurus malaianus Raffray (subseq. mon. Raffray, 1890b:206) Malaya, Sumatra [5 spp.] Trichopnites Dajoz, 1982:500 Type species: Trichopnites pubescens Dajoz (orig. des.) Madagascar [1 sp.] Triomicrus Sharp, 1883a:325 Type species: Bryaxis protervus Sharp (des. Lucas, 1920:655) China, Japan, Seychelles [7 spp.] NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 51 Trimiomicrus Raffray, 1911:110 (misspelling of Triomicrus) Tribe TRICHONYCHINI Amauronyx Reitter, 1881:519 Type species: Euplectus maerkelii Aube (des. Jeannel, 1950:160) W Palearctic [24 spp.] Berberonyx Jeannel, 1956a:52 Type species: Trichonyx lapidicola Raffray (orig. des.) Namunia Reitter, 1882c:21 1 Type species: Namunia myrmecophila Reitter (subseq. mon. Reitter, 1885a:81) Turkey, Greece [3 spp.] Trichonyx Chaudoir, 1845:164 Type species: Pselaphus sulcicollis Reichen- bach (monotypy) W Palearctic, E Nearctic [2 spp.] Tribe VALDINI Valda Casey, 1894:493 Type species: Valda frontalis Casey (mono- typy) W Nearctic (California) [1 sp.] Section PROTEROMORPHI [of GONIACERI- NAE] Tribe IMIRINI 1 minis Reitter, 1885b:228 (new name for Mirus Saulcy) France, Spain, Corsica [3 spp.] Mirus Saulcy, 1877:169 (preocc, not Albers, 1850) Type species: Mirus permirus Saulcy (mono- typy) Tribe PROTERINI Aulacobythus Leleup, 1986:533 Type species: Ectoparyphodes canaliculars Jeannel (orig. des.) Ivory Coast, Zaire [4 spp.] Breguetiella Leleup, 1973a:68 Type species: Breguetiella miserezi Leleup (orig. des.) Gabon [1 sp.] Bythinoderes Reitter, 1884b:407 Type species: Bythinoderes grabowskyi Reit- ter (monotypy) Borneo, Sumatra [2 spp.] Bythinophanax Reitter, 1884b:405 Type species: Bythinophanax bicornis Reitter (des. Lucas, 1920:153) Borneo, Sumatra [4 spp.] Cephalaxis Jeannel, 1960d:433 Type species: Cephalaxis championi Jeannel (orig. des.) India [1 sp.] Craspedopterus Jeannel, 1949c:51 Type species: Craspedopterus ghesquierei Jeannel (orig. des.) Ivory Coast, Zaire [4 spp.] Delamarea Jeannel, 1949b: 108 Type species: Delamarea laticeps Jeannel (monotypy) Afrotropical [4 spp.] Ectoparyphidius Jeannel, 1964a:33 Type species: Ectoparyphidius globosus Jean- nel (orig. des.) Zambia [1 sp.] Ectoparyphodes Jeannel, 1952a: 168 Type species: Ectoparyphodes incisifrons Jeannel (orig. des.) Afrotropical [ 1 1 spp.] Ectoparyphus Jeannel, 1949b:l 10 Type species: Ectoparyphus pauliani Jeannel (orig. des.) Afrotropical [10 spp.] Elaphobythus Jeannel, 1952a: 170 Type species: Elaphobythus leleupi Jeannel (orig. des.) Afrotropical [11 spp.] Goniomellus Jeannel, 1949c: 5 2 Type species: Goniomellus vrijdaghi Jeannel (orig. des.) Afrotropical [6 spp.] Hypoparyphantus Jeannel, 1953b: 134 Type species: Hypoparyphantus bucculentus Jeannel (orig. des.) Zaire [1 sp.] Imtempus Reitter, 1882c:209 Type species: Imtempus punctatissimus Reit- ter (subseq. mon. Reitter, 1885c:337) Philippines [1 sp.] Intempus Raffray, 1890a:99 (misspelling) Loebliobythus Leleup, 1986:550 Type species: Loebliobythus fimbriaticollis Leleup (orig. des.) Ivory Coast, Zaire [2 spp.] Mechanicus Schaufuss, 1887:158 Type species: Mechanicus chlamydophorus Schaufuss (monotypy) Oriental, New Guinea [5 spp.] Mimoplectus Raffray, 1918:490 52 FIELDIANA: ZOOLOGY Type species: Mimoplectus orientalis Raffray (monotypy) Laos [1 sp.] Neodeuterus Schaufuss, 1887:151 Type species: Neodeuterus admirandus Schaufuss (des. Lucas, 1920:435) Sumatra [2 spp.] Oxygastrobythus Leleup. 1986:567 Type species: Oxygastobythus loebli Leleup (orig. des.) Ivory Coast [1 sp.] Paryphantellus Jeannel, 195 lc:50 Type species: Paryphantellus latipennis Jean- nel (orig. des.) Afrotropical [9 spp.] Paryphopterus Jeannel. 1952d:36 Type species: Paryphopterus brevicollis Jean- nel (orig. des.) Ivory Coast, Zaire [2 spp.] Phthartomicrus Schaufuss, 1887:156 Type species: Phthartomicrus externus Schau- fuss (monotypy) Sumatra [1 sp.] Platycerobythus Jeannel, 1959d:500 Type species: Platycerobythus lamellifer Jean- nel (orig. des.) Tanzania [1 sp.] Proterus Raffray, 1897a:231 Type species: Proterus punctatus Raffray (monotypy) Sumatra [1 sp.] Pseudoterus Raffray, 1893a:469 Type species: Pseudoterus crassicornis Raffray (monotypy) Sumatra [1 sp.] Salpictomelus Jeannel, 1960d:434 Type species: Salpictomelus pachys Jeannel (orig. des.) India [1 sp.] Ulugurubythus Leleup. 1973b:263 Type species: Elaphobythus rugosicollis Jean- nel (orig. des.) Tanzania [1 sp.] Section TYCHOMORPHI [of GONIACERI- NAE] Tribe SPELEOBAMINI Prespelea Park, 1953a:251 Type species: Prespelea quirsfeldi Park (orig. des.) SE Nearctic (Tennessee) [2 spp.] Fusjuguma Park, 1956:55 (subgenus) Type species: Prespelea copelandi Park (orig. des.) Speleobama Park, 1951:52 Type species: Speleobama vana Park (orig. des.) SE Nearctic [1 sp.] Tribe TYCHINI Atychodea Reitter, 1884b:412 Type species: Atychodea simoniana Reitter (des. here) Borneo [4 spp.] Cylindrarctus Schaufuss, 1887:91 Type species: Cylindrarctus americanus Schaufuss (monotypy) Nearctic [5 spp.] Hesperotychus Schuster & Marsh, 1958a: 126 Type species: Hesperotychus adustus Schuster & Marsh (orig. des.) W Nearctic [13 spp.] Hylotychus Grigarick & Schuster, 1 962: 1 70 Type species: Hylotychus dentatus Grigarick & Schuster (orig. des.) W Nearctic [13 spp.] Lucifotychus Park & Wagner, 1962:22 Type species: Lucifotychus impellus Park & Wagner (orig. des.) Nearctic [12 spp.] Custotychus Park & Wagner, 1962:23 (subge- nus) Type species: Tychus daggyi Park (orig. des.) Paratychus Besuchet, 1960:24 Type species: Tychus appendiculatus Sahlberg (orig. des.) Lebanon, Israel [3 spp.] Pselaphotychus Khnzorian, 1957:159 Type species: Pselaphotychus onobrychidis Khnzorian (orig. des.) USSR (Armenia) [1 sp.] Tychomorphus Jeannel, 1950:342 Type species: Tychus tuberculatus Aube (orig. des.) W Palearctic [18 spp.] Tychus Leach, 1817:84 Type species: Pselaphus niger Reichenbach (monotypy) Palearctic [70 spp.] Tychoides Karaman, 1955:124 Type species: Tychus hirtulus Reitter (orig. des.) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 53 Division MACROSCELIA Subfamily PSELAPHINAE Section ARHYTODIMORPHI Tribe ARHYTODINI Caccoplectinus Chandler & Wolda, 1986:521 Type species: Caccoplectinus afoveatus Chan- dler & Wolda (orig. des.) French Guiana [1 sp.] Caccoplectus Sharp, 1887:22 Type species: Caccoplectus celatus Sharp (monotypy) SW Nearctic, Neotropical [36 spp.] Eichiella Leleup, 1976e:311 Type species: Eichiella demeyeri Leleup (orig. des.) Madagascar [2 spp.] Holozodinus Jeannel, 1949c:216 Type species: Holozodinus schoutedeni Jean- nel (orig. des.) Zaire [1 sp.] Holozodoides Dajoz, 1982:503 Type species: Holozodoides peyrierasi Dajoz (orig. des.) Madagascar [1 sp.] Holozodus Fairmaire, 1898b:346 (new name for Hologlyptus Fairmaire) Madagascar [4 spp.] Hologlyptus Fairmaire, 1898a:338 (preocc, not LeConte, 1866; Pomel, 1883) Type species: Hologlyptus raffrayi Fairmaire (monotypy) Pasiglyptus Berg, 1899:79 (unnecessary new name for Hologlyptus Fairmaire) Rhytus Westwood, 1870:126 (not preoccupied by Rytus King, 1866; see Discussion) Type species: Rhytus vestitus Westwood (monotypy) S Neotropical [ 1 1 spp.] Arhytodes Reitter, 1882c:209 (new name for Rhytus Westwood) Tetraglyptinus Jeannel, 1960b: 167 Type species: Tetraglyptinus longifrons Jean- nel (orig. des.) Madagascar [1 sp.] Tetraglyptus Jeannel, 1956f:46 Type species: Tetraglyptus pauliani Jeannel (orig. des.) Madagascar, Comoros [4 spp.] Section CTENISTOMORPHI [of PSELAPHIN- AE] Tribe ATTAPSENHNI Attapsenius Bruch, 1933:27 Type species: Attapsenius chernosvitovi Bruch (orig. des.) N Argentina, Brazil [2 spp.] Tribe CEOPHYLLINI Ceophyllus LeConte, 1849:73 Type species: Ceophyllus monilis LeConte (monotypy) E Nearctic [1 sp.] Tribe CHALCOPLECTINI Chalcoplectus Oke, 1925:13 Type species: Chalcoplectus depressus Oke (monotypy) Australia [1 sp.] Tribe CTENISTINI AtinusHorn, 1868:127 Type species: Ctenistes monilicornis Brendel (monotypy) E Nearctic [2 spp.] Biotus Casey, 1887a:456 Type species: Biotus formicarius Casey (monotypy) SW Nearctic [1 sp.] Centrotoma Heyden, 1849:182 Type species: Centrotoma lucifuga Heyden (monotypy) Palearctic, India [9 spp.] Chenniopsis Raffray, 1904b:338 Type species: Chenniopsis madecassa Raffray (monotypy) Madagascar [1 sp.] Chennium Latreille, 1807:77 Type species: Chennium bituberculatum La- treille (monotypy) W Palearctic [1 1 spp.] Ctenicellus Raffray, 1904b:344 Type species: Ctenicellus major Raffray (des. here) Australia [2 spp.] Ctenisis Raffray, 1890a: 143 Type species: Desimia dispar Sharp (des. Raf- fray, 1897a:271) Neotropical, SW Nearctic [8 spp.] Ctenisodes Raffray, 1897a:274 Type species: Ctenisodes laticeps Raffray (monotypy) Mexico, Cuba [2 spp.] Ctenisomimus Raffray, 1904b:341 Type species: Sognorus oneili Raffray (mono- typy) South Africa, Angola [2 spp.] 54 FIELDIANA: ZOOLOGY Ctenisomorphus Raffray, 1890a: 142 Type species: Ctenistes major Raffray (mono- typy) Ethiopia, W Palearctic [4 spp.] Ctenisophanes Jeannel, 1954a:259 Type species: Centrophthalmus angustior Fairmaire (orig. des.) Madagascar [4 spp.] Ctenisophus Raffray, 1900b: 208 Type species: Ctenisophus morosus Raffray (des. here) Oriental, New Guinea, Australia [2 1 spp.] Ctenisoschema Jeannel, 1956a: 179 Type species: Ctenisomorphus garamanticus Peyerimhoff (orig. des.) Algeria [1 sp.] Ctenistes Reichenbach, 1 8 1 6:75 Type species: Ctenistes palpalis Reichenbach (monotypy) Palearctic, Oriental, Afrotropical, South Af- rica [30 spp.] Diony.x LePeletier & Serville, 1825:221 Type species: Dionyx dejeanii LePeletier & Serville (monotypy) Tecnesis Peyerimhoff, 1925:59 (subgenus) Type species: Ctenistes vaulogeri Pic (mono- typy) Leptoctenistes Jeannel, 1956a: 167 (subgenus) Type species: Ctenistes wittmeri Jeannel (orig. des.) Tectenis Jeannel, 1 959d:6 1 7 (misspelling of Tecnesis) Tecnesites Jeannel, 1961:451 (subgenus) Type species: Ctenistes ceylanicus Jeannel (orig. des.) Ctenistidius Jeannel, 1957a:27 Type species: Ctenistidius coomani Jeannel (orig. des.) Oriental, Afghanistan [3 spp.] Ctenistodites Jeannel, 1957b:105 Type species: Ctenistodites pachycerus Jean- nel (orig. des.) Angola [1 sp.] Desimia Reitter, 1 88 1 :457 (new name for Tetrads Sharp) W Palearctic, Afrotropical [23 spp.] Tetracis Sharp, 1874b:79 (preocc, not Guenee, 1857) Type species: Tetracis complex Sharp (mono- typy) Desmia Leng, 1920:132 (misspelling) Desimiella Jeannel, 1949a:200 (subgenus) Type species: Desimia subcalva Reitter (orig. des.) Xenodesimia Jeannel, 1959d:632 (subgenus) Type species: Desimia rugosiventris Jeannel (orig. des.) Edocranes Reitter, 1 885b:228 (new name for Nar- codes King) Australia [8 spp.] Narcodes King, 1863:38 (preocc, not Agassiz, 1845) Type species: Narcodes varia King (des. Lu- cas, 1920:431) Enoptostomus Schaum, 1864:528 Type species: Enoptostomus wollastoni Schaum (monotypy) Palearctic, Afrotropical, South Africa, Bur- ma, Brazil [23 spp.] Epicaris Reitter, 1882c:202 Type species: Tetracis ventralis Raffray (monotypy) Afrotropical, South Africa [2 spp.] Taphrophorus Schaufuss, 1882b:350 Type species: Taphrophorus doriae Schaufuss (des. Jeannel, 1949a: 192) Gnorosus Raffray, 1908a:335 Type species: Sognorus croissandeaui Reitter (des. Lucas, 1920:306) W Palearctic [2 spp.] Hynneophorus Leleup, 1972:171 Type species: Hynneophorus franzi Leleup (orig. des.) Madagascar [1 sp.] Laphidioderomimus Leleup, 1972:167 Type species: Laphidioderomimus madecas- sus Leleup (orig. des.) Madagascar [1 sp.] Laphidioderus Raffray, 1887:20 Type species: Laphidioderus capensis Raffray (monotypy) South Africa [4 spp.] Metactenistes Jeannel, 1 956a: 1 74 Type species: Ctenisomorphus alternans Raf- fray (orig. des.) Sumatra [1 sp.] Parastectenis Jeannel, 1957b: 106 Type species: Parastectenis spinosus Jeannel (orig. des.) Angola [1 sp.] Pilopius Casey, 1897:617 Type species: Pilopius lacustris Casey (des. Bowman, 1934:144) Nearctic, Neotropical, Japan, Korea [17 spp.] Poroderopsis Jeannel, 1960d:448 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 55 Type species: Poroderopsis championi Jeannel (orig. des.) India [2 spp.] Poroderus Sharp, 1883a:294 Type species: Ctenistes armatus Sharp (des. Lucas, 1920:531) Oriental, Japan [12 spp.] Sognorus Reitter, 1881:458 Type species: Ctenistes calcaratus Baudi (des. Raffray, 1908a: 341) Palearctic, Oriental, Afrotropical [1 1 spp.] Agnesia Lokay, 1907:87 (preocc, not Koninck, 1883; Michaelsen, 1898) Type species: Agnesia cilicica Lokay (mono- typy) Stectenis Raffray, 1908a:339 Type species: Ctenistes simonis Reitter (monotypy) Indiella Blattny, 1925:215 Type species: Ctenistes birmanensis Mot- schulsky (monotypy) Sognorites Jeannel, 1949c:227 Type species: Sognorites saegeri Jeannel (orig. des.) Gonorsus Jeannel, 1 958b: 1 1 6 Type species: Ctenistes breviceps Sharp (orig. des.) Stectenidius Jeannel, 1957b: 107 Type species: Stectenidius latitarsis Jeannel (orig. des.) Angola [1 sp.] Tribe ODONTALGINI Algodontodes Jeannel, 1960b: 175 Type species: Algodontodes acutipalpus Jean- nel (orig. des.) Madagascar [1 sp.] Algodontus Jeannel, 1955b:144 Type species: Odontalgus costatus Raffray (orig. des.) South Africa [5 spp.] Madontalgus Dajoz, 1982:506 Type species: Madontalgus sculpturatus Da- joz (orig. des.) Madagascar [1 sp.] Odontalgus Raffray. 1877:286 Type species: Odontalgus tuberculatus Raf- fray (des. Jeannel, 1949a: 178) Afrotropical, South Africa, Oriental, Ko- rea [49 spp.] Herminiella Blattny, 1925:211 Type species: Ctenistes costulatus Motschul- sky (monotypy) Tribe PACHYGASTRODINI Pachygastrodes Leleup, 1969b:284 Type species: Pachygastrodes minimipennis Leleup (monotypy) Madagascar [1 sp.] Pachygastrodirius Leleup, 1976d:268 Type species: Pachygastrodirius globulicollis Leleup (orig. des.) Madagascar [2 spp.] Tribe PETANOPINI Daveyia Lea, 1912b:45 Type species: Daveyia mira Lea (monotypy) Australia [1 sp.] Petanops Jeannel, 1954c:99 Type species: Petanops armstrongi Jeannel (monotypy) Australia [1 sp.] Tribe SCHISTODACTYLINI Leanymus Raffray, 1900b:219 Type species: Leanymus palpalis Raffray (monotypy) Australia [2 spp.] Schistodactylus Raffray, 1883:243 Type species: Schistodactylus phantasma Raf- fray (monotypy) Australia [3 spp.] Tribe SOMATIPIONINI Somatipion Schaufuss, 1877b:457 Type species: Somatipion globulifer Schau- fuss (= Faronus punctatus King) (monotypy) Australia [1 sp.] Tribe TMESIPHORINI Afrotyrus Jeannel, 1949c:256 Type species: Tyrus glabratus Jeannel (orig. des.) Zaire, Uganda [4 spp.] Tmesiphorellus Jeannel, 1 954d: 1 69 Type species: Tmesiphorellus leleupi Jeannel (orig. des.) Aphanethrix Raffray, 1908c:209 Type species: Aphanethrix andrewesi Raffray (monotypy) India [1 sp.] Brinckidiella Leleup, 19 74b: 504 Type species: Brinckidiella abnormipalpa Le- leup (orig. des.) South Africa [1 sp.] 56 FIELDIANA: ZOOLOGY Dacnotillus Raffray, 1908a:374 Type species: Tmesiphorus pubescens Raffray (des. here) E Oriental [7 spp.] Elaphidipalpus Jeannel, 1964c: 197 Type species: Elaphidipalpus leleupi Jeannel (orig. des.) South Africa [1 sp.] Gasterotropis Raffray, 1914b:463 Type species: Gasterotropis poweri Raffray (monotypy) South Africa [1 sp.] Itombworites Jeannel, 1 960c: 1 70 Type species: Itombworites leleupi Jeannel (orig. des.) Zaire [1 sp.] Machadoites Jeannel, 1 95 lc: 1 2 1 Type species: Machadoites termitophilus Jeannel (orig. des.) Angola [1 sp.] Neosintectes Jeannel, 1964c: 195 Type species: Neosintectes brevipennis Jean- nel (orig. des.) South Africa [2 spp.] Pselaphocerodes Jeannel. 1955b: 1 65 Type species: Pselaphocerus nodicornis Raf- fray (orig. des.) South Africa [1 sp.] Pselaphocerus Raffray. 1887:28 Type species: Pselaphocerus peringueyi Raf- fray (des. Lucas, 1920:544) South Africa [10 spp.] Raphitreodes NEW GENUS (validation of Ra- phitreodes Jeannel) Type species: Raphitreus dentimanus Raffray (des. here) Oriental [2 spp.] Raphitreodes Jeannel, 1958b: 119 (nomen nu- dum, no type species designated) Raphitreus Sharp, 1883a:298 Type species: Tmesiphorus speratus Sharp (monotypy) Japan [1 sp.] Riphaterus Jeannel, 1964c: 199 Type species: Riphaterus galeatus Jeannel (orig. des.) South Africa [1 sp.] Smilestethus Cerruti, 1955:96 Type species: Smilestethus patrizii Cerruti (monotypy) Kenya [1 sp.] Syntectodes Reitter, 1882c:202 Type species: Syntectodes diversipalpus Reit- ter (subseq. mon. Reitter, 1885c:333) Sri Lanka [2 spp.] Sintectodes Reitter, 1885c:333 (misspelling) Tapiloites Jeannel, 1958a:352 Type species: Tapiloites pantosi Jeannel (orig. des.) Zaire [1 sp.] Tmesiphorinus Leleup, 197 lb: 169 Type species: Tmesiphorites longipes Jeannel (orig. des.) South Africa [1 sp.] Tmesiphorites Jeannel, 1953b:299 Type species: Tmesiphorites grossepunctatus Jeannel (orig. des.) Afrotropical, South Africa [1 1 spp.] Tmesiphoromimus Lobl, 1964b: 1 Type species: Tmesiphoromimus samsinaki Lobl (orig. des.) S China [1 sp.] Tmesiphorus LeConte, 1849:75 Type species: Pselaphus carinatus Say (des. Lucas, 1920:643) Holarctic, Afrotropical, South Africa, Ori- ental, New Guinea, Australia [60 spp.] Glyptosoma Motschulsky, 1851:480 Type species: Glyptosoma paulinae Mot- schulsky (monotypy) Sintectes Westwood, 1 870: 1 29 Type species: Sintectes carinatus Westwood (= Tmesiphorus westwoodi Raffray) (monotypy) Syntectes Jeannel, 1949a:203 (misspelling) Tropeogaster Jeannel, 1949a:207 Type species: Tmesiphorus denticornis Raf- fray (orig. des.) Afrotropical [4 spp.] Xenotmesiphorus Leleup, 1974b:500 Type species: Xenotmesiphorus messoris Le- leup (orig. des.) South Africa [1 sp.] Tribe TYRINI Subtribe CENTROPHTHALMINA Camaldites Jeannel, 1949c:251 Type species: Camaldites gracilis Jeannel (orig. des.) Zaire [1 sp.] Camaldosis Jeannel, 1 95 1 f:69 Type species: Camaldosis crassicornis Jeannel (orig. des.) Zaire [1 sp.] Centrophthalmina Raffray, 1908a:353, 365 Type species: Centrophthalmus laevis Raffray (monotypy) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 57 E Oriental [2 spp.] Centrophtalmina Raffray, 1908a:365 (misspell- ing, first reviser Raffray, 1911:154) Centrophthalmosis Raffray, 1904b:376 Type species: Centrophthalmosis reitteri Raf- fray (des. Lucas, 1920:170) Afrotropical, South Africa, Egypt, Israel [27 spp.] Centrophthalmus Schmidt-Gobel, 1838:7 Type species: Centrophthalmus paria Schmidt- Gobel (monotypy) Canary Islands, N Africa, Afrotropical, South Africa, Oriental, New Guinea [90 spp.] Camaldus Fairmaire, 1863:637 Type species: Camaldus villosulus Fairmaire (monotypy) Enantius Schaufuss, 1877a: 18 Type species: Enantius punctipennis Schau- fuss (monotypy) Oriental [7 spp.] Tribe TYRINI Subtribe HAMOTINA Apharus Reitter, 1882a: 129 Type species: Apharus muelleri Reitter (monotypy) Neotropical [6 spp.] Aploderina Raffray, 1904b:386 Type species: Aploderina sulcicornis Raffray (monotypy) Bolivia [1 sp.] Cercoceroides Raffray, 1896c: 137 Type species: Cercocerus germaini Raffray (des. Lucas, 1920:173) Neotropical [7 spp.] Cercoceropsis Raffray, 1904b:387 Type species: Cercoceropsis longipes Raffray (monotypy) Brazil, Paraguay [2 spp.] Cercocerulus Raffray, 1905:411 Type species: Phalepsus hirsutus Schaufuss (monotypy) Brazil [1 sp.] Circocerus Motschulsky, 1855:16 Type species: Circocerus perplexus Motschul- sky (monotypy) Nearctic, Neotropical [2 spp.] Cercocerus LeConte, 1861:57 (misspelling) Upoluna Schaufuss, 1886:283 Type species: Upoluna flavitarsis Schaufuss (= Circocerus batrisioides Motschul- sky) (monotypy) Hamotidius Jeannel, 1962:476 Type species: Hamotidius hirtellus Jeannel (orig. des.) Chile [1 sp.] Hamotocellus Raffray, 1912:444 Type species: Hamotocellus hirsutus Raffray (monotypy) Brazil [2 spp.] Hamotulus Schaufuss, 1887:108 Type species: Bryaxis chamaeleon Schaufuss (monotypy) Australia, New Zealand [ 1 2 spp.] Tychotyrus Broun, 1893b: 168 Type species: Tychotyrus sternalis Broun (des. here) Hamotus Aube, 1844:91 Type species: Hamotus lateritius Aube (des. Lucas, 1920:314) Nearctic, Neotropical, Chile [94 spp.] Hamotoides Schaufuss, 1888:298 (subgenus) Type species: Hamotus tritomus Reitter (des. Jeannel, 1962:473) Mothaus Jeannel, 1962:477 Type species: Mothaus delamarei Jeannel (orig. des.) Argentina (Tucuman) [ 1 sp.] Phamisulus Reitter, 1888:234 Type species: Phamisulus hetschkoi Reitter (monotypy) Brazil [1 sp.] Neophamisus Reitter, 1888:234 Type species: Phamisus horroris Schaufuss (des. here) Pselaphocompsus Raffray, 1908a:402; 1912:447 Type species: Pselaphocompsus punctatus Raffray (monotypy) Brazil [1 sp.] Pseudohamorus Raffray, 1890a: 154 Type species: Hamotus conjunctus Reitter (des. Park, 1942:337) Neotropical [5 spp.] Saltisedes Kubota, 1944:8 Type species: Saltisedes brunneus Kubota (orig. des.) Japan [1 sp.] Stethotaphrus NEW NAME (for Taphrostethus Schaufuss) E Oriental, New Guinea [4 spp.] Taphrostethus Schaufuss, 1882d:151 (preocc, not Fieber, 1844) Type species: Epicaris hamotoides Schaufuss (monotypy) 58 FIELDIANA: ZOOLOGY Tribe TYRINI Subtribe JANUSCULINA Janusculus Cerruti, 1 970: 1 1 7 Type species: Janusculus leleupi Cerruti (monotypy) Eritrea [1 sp.] Tribe TYRINI Subtribe TYRINA Abascantodes Strand, 1 928:2 (new name for Abas- cantus Schaufuss) Australia [1 sp.] Abascantus Schaufuss, 1886:258 (preocc, not Stal, 1864) Type species: Abascantus sannio Schaufuss (monotypy) Abascantis Strand, 1928:2 (misspelling) Acylobythus Jeannel, 1 960b: 1 78 Type species: Acylobythus sulcatus Jeannel (orig. des.) Madagascar [2 spp.] Acylopselaphus Raffray, 1883:237 Type species: Acylopselaphus mariae Raffray (monotypy) Madagascar [11 spp.] Acylotyrus Jeannel, 1954a:278 Type species: Centrophthalmus communis Schaufuss (orig. des.) Madagascar [13 spp.] Leptotyrus Jeannel, 1954a:283 Type species: Leptotyrus vadoni Jeannel (orig. des.) Agatyrus Broun, 1917:382 Type species: Agatyrus fulvihirtus Broun (monotypy) New Zealand [1 sp.] Anagonus Fauvel, 1903:284 Type species: Anagonus fracticornis Fauvel (monotypy) New Caledonia [1 sp.] Ancystrocerus Raffray, 1893a:500 Type species: Ancystrocerus sumatrensis Raf- fray (monotypy) Malaysia, Sumatra [9 spp.] Anitra Casey, 1 894:499 Type species: Anitra glaberula Casey (mono- typy) SW Nearctic (Arizona) [ 1 sp.] Cedius LeConte, 1849:74 Type species: Cedius spinosus LeConte (des. Lucas, 1920:169) E Nearctic [4 spp.] Sinistrocedius NEW SUBGENUS (validation of Sinistrocedius Park) (subgenus) Type species: Cedius ziegleri LeConte (des. here) Sinistrocedius Park, 1949b:336 (nomen nudum, no type species designated) Chasoke Chandler, 1987b:32 Type species: Chasoke victoriae Chandler (orig. des.) Australia [1 sp.] Collacerothorax Lea, 1912a:450 Type species: Collacerothorax sculpt iceps Lea (monotypy) Australia [2 spp.] Ctenotillus Raffray, 1897a:277 Type species: Ctenotillus costatus Raffray (monotypy) Sumatra [1 sp.] Durbos Sharp, 1874c:495 Type species: Durbos priscus Sharp (mono- typy) Australia, Taiwan [3 spp.] Ephimia Reitter, 1882c:203; 1883a:34 Type species: Ephimia simoni Reitter (sub- seq. mon. Reitter, 1883a:34) Panama, West Indies [5 spp.] Ephymia Reitter, 1883a:34 (misspelling) Juxtahamotopsis Park, 1942:303 Type species: Juxtahamotopsis bardeni Park (monotypy) Eudranes Sharp, 1892:241 Type species: Eudranes carinatus Sharp (monotypy) Australia [1 sp.] Eulasinus Sharp, 1892:240 Type species: Eulasinus walkeri Sharp (monotypy) China (Chekiang) [1 sp.] Franziotus Leleup, 1972:175 Type species: Franziotus hirsutus Leleup (orig. des.) Madagascar [3 spp.] Gerallus Sharp, 1874c:493 Type species: Gerallus nanus Sharp (mono- typy) Australia [8 spp.] Zosimus Schaufuss, 1886:293 (preocc, not Leach, 1825) Type species: Zosimus longipes Schaufuss (des. here) Zozimus Raffray, 1890a: 160 (misspelling) Hamotopsis Raffray, 1 900b: 2 1 7 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 59 Type species: Hamotopsis australasiae Raf- fray (monotypy) Australia [4 spp.] Horniella Raffray, 1905:434 (new name for Hor- nia Raffray) Sri Lanka [1 sp.] Hornia Raffray, 1901b:29 (preocc., not Riley, 1878) Type species: Hornia hirtella Raffray (mono- typy) Labomimus Sharp, 1883a:300 Type species: Labomimus reitteri Sharp (monotypy) Japan, W Oriental [5 spp.] Lasinus Sharp, 1874a: 106 Type species: Lasinus spinosus Sharp (mono- typy) Japan, Oriental [5 spp.] Lethenomus Raffray, 1895b:396 Type species: Aplodea villosa Schaufuss (monotypy) Chile, S Argentina [3 spp.] Marellus Motschulsky, 1851:483 Type species: Marellus aegyptiacus Motschul- sky (monotypy) Afrotropical, South Africa, N Africa [ 1 1 spp.] Mipseltyrus Park, 1953a:262 Type species: Mipseltyrus nicolayi Park (orig. des.) Nearctic [4 spp.] Neopalimbolus Oke, 1926:285; 1928:22 Type species: Neopalimbolus goudiei Oke (monotypy) Australia [1 sp.] Neotyrus Raffray, 1895b:396 Type species: Aplodea gibbicollis Schaufuss (monotypy) Neotropical [5 spp.] Nesiotyrodes Jeannel, 1954a:285 Type species: Nesiotyrodes perrieri Jeannel (orig. des.) Madagascar [1 sp.] Palimbolus Raffray, 1890a: 159 Type species: Tyrus mirandus Sharp (mono- typy) Australia [23 spp.] DidimoproraRafTray, 1890a: 159 Type species: Tyrus victoriae King (mono- typy) Didymoprora Raffray, 1905:428 (misspelling) Palodea Jeannel, 1962:450 Type species: Aplodea hirta Reitter (orig. des.) Brazil [3 spp.] Phormiobius Broun, 1917:381 Type species: Phormiobius halli Broun (monotypy) New Zealand [1 sp.] Plesiotyrus Broun, 1914:94 Type species: Tyrus crassipes Broun (orig. des.) New Zealand [1 sp.] Pselaphodes Westwood, 1870:129 Type species: Pselaphodes villosus Westwood (monotypy) Oriental [10 spp.] Atherocolpus Raffray, 1882:15 Type species: Pselaphodes heterocerus Raffray (des. here) Pseudotychus Raffray, 1 897b: 1 14 Type species: Pseudotychus nigerrimus Raf- fray (monotypy) South Africa [1 sp.] RytusKing, 1866:302 Type species: Rytus punctatus King (= Tyrus subulatus King) (orig. des.) Australia, New Guinea [9 spp.] Schaufussia Raffray, 1883:238 Type species: Bryaxis brevis Schaufuss (= Ty- rus formosus King) (monotypy) Australia [2 spp.] Spilorhombus Raffray, 1900b:225 Type species: Spilorhombus hirtus Raffray (monotypy) Australia [1 sp.] Subulipalpus Schaufuss, 1877a:23 Type species: Subulipalpus spinicoxis Schau- fuss (monotypy) Oriental [2 spp.] Tasmanityrus Chandler, 1987b:20 Type species: Tyromorphus auricomus Lea (orig. des.) Australia [7 spp.] Termitotyrus Borgmeier, 1954:209 Type species: Termitotyrus palpalis Borg- meier (orig. des.) Brazil [1 sp.] Tyrodes Raffray, 1908a:379 Type species: Tyrus histrio Schaufuss (des. Lucas, 1920:663) Oriental [5 spp.] Tyrodites Jeannel, 1960d:455 Type species: Tyrodites championi Jeannel (orig. des.) Tyrogatunus Park, 1942:338 Type species: Tyrogatunus zeteki Park (monotypy) Panama [1 sp.] Tyrogetus Broun, 1893b: 165 60 FIELDIANA: ZOOLOGY Type species: Tyrogetus optandus Broun (monotypy) Australia, New Zealand [5 spp.] Tyromorphus Raffray, 1883:240 Type species: Tyromorphus nitidus Raffray (monotypy) Australia, New Guinea [18 spp.] Tyropsis Saulcy, 1874:80 Type species: Tyropsis chevrolati Saulcy (= Pselaphus castaneus Blanchard) (monotypy) Chile, S Argentina [20 spp.] Aplodea Reitter, 1882c:203; 1883b:47 Type species: Aplodea palpalis Reitter (orig. des.) Tyrus Aube, 1833:15 Type species: Pselaphus mucronatus Panzer (monotypy) Holarctic, Oriental [16 spp.] Pytna Casey, 1887a:457 Type species: Pytna corticina Casey (mono- typy) Pseudotyrus Raffray, 1890a: 153 Type species: Pseudotyrus corticalis Raffray (subseq. mon. Raffray, 1890b:215) Novissimus Blattny, 1925:217 Type species: Novissimus helferi Blattny (monotypy) Vadoniotus Jeannel, 1954a:287 Type species: Vadoniotus filipalpis Jeannel (orig. des.) Madagascar [1 sp.] Zeatyrus Sharp, 1881:48 Type species: Zeatyrus lawsoni Sharp (mono- typy) New Zealand [1 sp.] Parmipalpus Broun, 1881:662 Type species: Parmipalpus montivagus Broun (monotypy) Section CYATHIGERIMORPHI [of PSELA- PHINAE] Tribe BARROSELLINI Barrosellus Jeannel, 1951c: 107 Type species: Barrosellus sphaeroides Jeannel (orig. des.) Afrotropical [6 spp.] Tribe CYATHIGERINI Cyathiger King, 1865:174 Type species: Cyathiger punctatus King (monotypy) Australia, Malaysia, Singapore [13 spp.] Cyathigerodes Jeannel, 1 95 lc: 1 10 Type species: Cyathigerodes machadoi Jean- nel (orig. des.) Afrotropical [32 spp.] Denicyathiger Jeannel, 1951c: 109 Type species: Cyathiger bironis Raffray (orig. des.) New Guinea [1 sp.] Manuleiger Jeannel, 1961:449 Type species: Manuleiger remyi Jeannel (orig. des.) Sri Lanka [1 sp.] Paracyathiger Jeannel, 1 95 1 c: 1 09 Type species: Cyathiger heterocerus Raffray (orig. des.) Oriental, Japan, New Guinea [42 spp.] Tribe HYBOCEPHALINI Apharina Reitter, 1882c:209; 1883c:295 Type species: Apharina simonis Reitter (des. Jeannel, 1949c:210) Oriental, Afrotropical [17 spp.] Apharinodes Raffray, 1 890a: 1 36 Type species: Apharinodes squamosa Raffray (subseq. mon. Raffray, 1890b:210) Singapore [2 spp.] Filigerinus Jeannel, 1949a: 176 Type species: Filiger vestitus Raffray (orig. des.) Afrotropical, Sumatra [7 spp.] Filigerodes Jeannel, 1949c:207 Type species: Filigerodes verrucosus Jeannel (orig. des.) Sudan, Zaire [3 spp.] Hybocephalodes Raffray, 1908a:322 Type species: Hybocephalus dentiventris Reit- ter (des. Lucas, 1920:339) Borneo [3 spp.] Hybocephalus Motschulsky, 1851:482 Type species: Hybocephalus minimus Mot- schulsky (des. Lucas, 1920:339) Oriental [2 spp.] Mecochelia Motschulsky, 1851:480 Type species: Mecochelia impressa Motschul- sky (des. Lucas, 1920:394) "Indes oriental" [2 spp.] Mestogaster Schmidt-Gobel, 1838:9 Type species: Mestogaster crassicornis Schmidt-Gobel (monotypy) Oriental, Afrotropical [8 spp.] Metaxoides Schaufuss, 1 877a: 1 3 Type species: Metaxoides bruchiformis Schaufuss (monotypy) Mestogastridius Jeannel, 1952d:54 (subgenus) NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 61 Type species: Mestogastridius punctaticeps Jeannel (orig. des.) Pseudapharina Raffray, 1 890a: 1 36 Type species: Filiger primus Reitter (mono- typy) Borneo [1 sp.] Stipesa Sharp, 1874a: 108 Type species: Stipesa rudis Sharp (monotypy) Afrotropical, Oriental, Japan [22 spp.] Filiger Schaufuss, 1 877a: 1 7 Type species: Filiger ampliventris Schaufuss (des. Lucas, 1920:295) Tribe MACHADOINI Machadous Jeannel, 1951c: 105 Type species: Machadous punctatus Jeannel (orig. des.) Afrotropical [15 spp.] Section PSELAPHOMORPHI [of PSELAPHIN- AE] Tribe PHALEPSINI Phalepsus Westwood, 1870:131 Type species: Phalepsus subglobosus West- wood (monotypy) Neotropical [10 spp.] Phalespus Raffray, 1890a: 162 (misspelling) Pseudophanias Raffray, 1890a: 161 Type species: Pseudophanias malaianus Raf- fray (subseq. mon. Raffray, 1890b:214) Malaysia, Sumatra [10 spp.] Tribe PSELAPHINI Acmoeonotus Motschulsky, 1851:483 Type species: Acmoeonotus motschoulskyi Motschulsky (monotypy) Burma [1 sp.] Acmaeonotus Gemminger & Harold, 1868:686 (misspelling) Af ropselaphus Jeannel , 1950:384 Type species: Pselaphus algericus Raffray (orig. des.) Canary Islands, NW Africa [9 spp.] Callithoracoides Strand, 1928:2 (new name for Callithorax Motschulsky) Bengal [1 sp.] Callithorax Motschulsky, 185 1:482 (preocc, not Agassiz, 1846) Type species: Callithorax subtilis Motschul- sky (monotypy) Curculionellus Westwood, 1870:127 Type species: Curculionellus angulicollis Westwood (= Pselaphus punctatus King) (des. Jeannel, 1951a:6) E Oriental, New Guinea, Australia [ 1 2 spp.] Dicentrius Reitter, 1882c:208 Type species: Pselaphus merklii Reitter (monotypy) Palearctic, Afrotropical, China, New Zea- land [38 spp.] Pselaphogenius Reitter, 1910:155 Type species: Pselaphus quadricostatus Reit- ter (des. Jeannel, 1950:384) Pselaphodinus Jeannel, 1950:389 Type species: Pselaphus longipalpis Kiesen- wetter (orig. des.) Geopselaphus Jeannel, 1956a: 152 Type species: Pselaphus sencieri Coquerel (orig. des.) NW Africa, Spain, Balearic Islands [23 spp.] Margaris Schaufuss, 1877b:453 Type species: Margaris imperialis Schaufuss (monotypy) Australia [1 sp.] Mentraphus Sharp, 1883b:225 Type species: Mentraphus pselaphodes Sharp (monotypy) Afrotropical, C Palearctic, Oriental [6 spp.] Psilocephalus Raffray, 1877:284 (preocc, not Swainson, 1839; Salter, 1866) Type species: Psilocephalus formicetorum Raf- fray (monotypy) Apenicillus Reitter, 1885b:228 (new name for Psilocephalus Raffray) Apenicellus Jeannel, 1949c: 197 (misspelling) Neopselaphus Jeannel, 1951a:8 Type species: Pselaphus bizonatus Schaufuss (orig. des.) Neotropical, S Nearctic (Texas) [8 spp.] Pselaphaulax Reitter, 1909:218 Type species: Pselaphus dresdensis Herbst (monotypy) Afrotropical, South Africa, Palearctic, Ori- ental, Australia [6 1 spp.] Neopselaphaulax Jeannel, 1959d:593 (subge- nus) Type species: Pselaphus bifossulus Raffray (orig. des.) Pselaphellus Raffray, 1908a:307; 1908d:77 Type species: Pselaphus opacus Schaufuss (des. Lucas, 1920:544) Neotropical [8 spp.] Pselaphidius Jeannel, 195 la:9 Type species: Pselaphus filipalpis Reitter (orig. des.) Afrotropical, Palearctic, Oriental [15 spp.] 62 FIELDIANA: ZOOLOGY Pselaphischnus Raffray, 1897b:99 Type species: Pselaphischnus squamosus Raf- fray (monotypy) South Africa [1 sp.] Pselaphophus Raffray, 1890a: 139 Type species: Bryaxis atriventris Westwood (des. Lucas, 1920:544) Australia, New Guinea [5 spp.] Pselaphoptrus Reitter, 189 lb: 139 Type species: Pselaphoptrus kubischteki Reit- ter (monotypy) C Palearctic (Turkestan) [2 spp.] Faradayus Reitter, 1909:217 Type species: Pselaphus banghaasi Reitter (des. Jeannel, 195 la:6) Pselaphorites Jeannel, 1952a:260 Type species: Pselaphorites carinatus Jeannel (orig. des.) Afrotropical [18 spp.] Pselaphostomus Reitter. 1909:218 Type species: Pselaphus stussineri Saulcy (orig. des.) W Palearctic [21 spp.] Pselaphopsis Jeannel, 1950:389 Type species: Pselaphus kiesenwetteri Reitter (orig. des.) Pselaphotheseus Park. 1964:391 Type species: Pselaphotheseus hippolytae Park (orig. des.) New Zealand (Campbell Island) [1 sp.] Pselaphotrichus Besuchet, 1986:259 (new name for Trichopselaphus Jeannel) Afrotropical, India [6 spp.] Trichopselaphus Jeannel, 1 949c: 187 (preocc, not Chaudoir, 1843) Type species: Pselaphus gibbicollis Raffray (orig. des.) Pselaphoxys Raffray, 1890a: 139 Type species: Pselaphus delicatulus Raffray (monotypy) Afrotropical, South Africa [5 spp.] Pselaphus Herbst 1792:106 Type species: Pselaphus heisei Herbst (des. Shuckard, 1839:147; designations of Latreille, 1810:432, and Westwood, 1838:21, not valid, species not origi- nally included) Worldwide (except Neotropical, South Africa, Australia) [59 spp.] Stenosthetus Grimih& Pidgeon, 1832:300 (pub- lished in synonymy with Pselaphus: un- available, never validated) (see Black- welder, 1952:359) Pselaphinus Jeannel. 1 95 1 a: 1 0 (subgenus) Type species: Pselaphus pilipalpis Reitter (orig. des.) Tyraphus Sharp, 1874c:489 Type species: Tyraphus planus Sharp (des. Lucas, 1920:663) Japan, Oriental, Australia [19 spp.] Subfamily CLAVIGERINAE Tribe ADRANINI Adranes LeConte, 1849:83 Type species: Adranes coecus LeConte (monotypy) Nearctic [6 spp.] Tribe APODERIGERINI Ambrosiger Silvestri, 1926:261 Type species: Ambrosiger gravis Silvestri (orig. des.) China (Macao) [1 sp.] Apoderiger Wasmann, 1897:263 Type species: Apoderiger cervinus Wasmann (monotypy) Madagascar [2 spp.] Micrapoderiger Jeannel, 1960b:202 Type species: Micrapoderiger minutissimus Jeannel (orig. des.) Madagascar [1 sp.] Trymalius Fairmaire, 1898b:345 Type species: Trymalius foveicollis Fairmaire (monotypy) Madagascar [1 sp.] Tribe CLAVIGERINI Claviger Preyssler, 1790:68 Type species: Claviger testaceus Preyssler (monotypy) W Palearctic [42 spp.] Clavifer LaPorte. 1835:137 Type species: Claviger longicornis Muller (monotypy) Tribe CLAVIGERODINI Amphironchus Peyerimhoff 1939:347 Type species: Amphironchus alfierii Peyer- imhoff (monotypy) Egypt [1 sp.] Anaclasiger Raffray, 1890a: 165 Type species: Anaclasiger sinuaticollis Raffray (subseq. mon. Raffray, 1890b:216) Singapore [1 sp.] Ankarahitra Jeannel, 1954a:308 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 63 Type species: Ankarahitra perrieri Jeannel (orig. des.) Madagascar [1 sp.] Archiclaviger Heller, 1936:66 Type species: Archiclaviger overbecki Heller (monotypy) Java [1 sp.] Articerodes Raffray, 1890a: 167 Type species: Articerus syriacus Saulcy (monotypy) C Palearctic, Zaire, South Africa, ?Suma- tra [4 spp.] Pararticerus Jeannel, 1 955b: 1 79 Type species: Articerodes latus Raffray (orig. des.) Articeronomus Raffray, 1898a:268 Type species: Articeronomus nitidus Raffray (monotypy) Madagascar [1 sp.] Articeropsis Wasmann, 1893b:257 Type species: Articeropsis sikorae Wasmann (monotypy) Madagascar, Sri Lanka [4 spp.] Articerus Dalman, 1826:394 Type species: Articerus armatus Dalman (monotypy) Oriental [2+ undescribed spp.; see Besuch- et, 1986:262] Bironia Raffray, 1903a:99 Type species: Bironia cavernosa Raffray (monotypy) New Guinea [1 sp.] Cerylambus NEW NAME (for Amblycerus Raf- fray) Singapore [1 sp.] Amblycerus Raffray, 1895a:79 (preocc, not Thunberg, 1815) Type species: Amblycerus reticulatus Raffray (monotypy) Clavigerodes Raffray, 1877:279 Type species: Clavigerodes abyssinicus Raf- fray (monotypy) Afrotropical [4 spp.] Clavigeropsis Raffray, 1882:3 Type species: Clavigeropsis formicarius Raf- fray (monotypy) Zaire, Ethiopia, Australia [3 spp.] Colletocerus Celis, 1971:189 Type species: Colletocerus bouilloni Celis (orig. des.) Zaire [1 sp.] Commatocerodes Peringuey, 1888:85 Type species: Commatocerodes raffray i Per- inguey (monotypy) South Africa [1 sp.] Corynotopsis Jeannel, 195 lb:228 Type species: Corynotopsis scotti Jeannel (orig. des.) Ethiopia [1 sp.] Corynotus Jeannel, 1951c:27 Type species: Corynotus sulcatus Jeannel (orig. des.) Afrotropical [4 spp.] Dejaegeria Jeannel, 1964c:212 (justified emen- dation ofDejageria, incorrect original spell- ing) Type species: Dejaegeria joannae Jeannel (orig. des.) South Africa [1 sp.] Diartiger Sharp, 1883a:329 Type species: Diartiger fossulatus Sharp (des. Lucas, 1920:235) Japan [4 spp.] Microdiartiger Sawada, 1 964: 1 2 Type species: Microdiartiger japonicus Sa- wada (orig. des.) Coiffaitius Karaman, 1969:50 Type species: Coiffaitius ispartae Karaman (monotypy) Elasmatus Raffray, 1903a:97 Type species: Elasmatus lamellipes Raffray (monotypy) New Guinea [1 sp.] Fossiger Wasmann, 1918a:405 Type species: Fossiger assmuthi Wasmann (monotypy) India [1 sp.] Fustiger LeConte, 1866:109 Type species: Articerus braziliensis Westwood (des. here; see Discussion) Nearctic, Neotropical, Afrotropical, C Pale- arctic, India, Fiji [53 spp.] Commatocerus Raffray, 1882:1 Type species: Commatocerus elegantulus Raf- fray (monotypy) Pseudfustiger Reitter, 1 884a: 1 68 Type species: Articerus stricticornis Reitter (orig. des.) Pseud of ustiger Raffray, 1 890a: 1 66 (misspelling) Afrofustiger Jeannel, 1 952d: 1 8 Type species: Afrofustiger pleuralis Jeannel (orig. des.) Corynotellus Jeannel, 1 960c: 1 75 Type species: Corynotellus tubifer Jeannel (= Afrofustiger pleuralis Jeannel) (orig. des.) 64 FIELDIANA: ZOOLOGY Fustigerillus Wasmann, 1918a:400 Type species: Fustigerillus fusicornis Was- mann (monotypy) India [1 sp.] Fustigerinus Wasmann, 1912:105 Type species: Fustigerinus kohli Wasmann (des. Lucas, 1920:296) Zaire, Guinea, "Hungary" [8 spp.] Neocommatocerus Jeannel, 1949c:37 Type species: Fustigerinus nasicornis Rei- chensperger (orig. des.) Fustigerodes Reitter, 1884a: 168 (new sense; see Discussion) Type species: Fustiger madagascariensis Raf- fray (monotypy) Madagascar, Bolivia [3 spp.] Commatoceropsis Raffray, 1890a: 167 (revised status; see Discussion) Type species: Fustiger madagascariensis Raf- fray (monotypy) Commatocerinus Wasmann, 1897:260 Type species: Commatocerinus ranavalonae Wasmann (monotypy) Fustigeromimus Dajoz, 1982:515 Type species: Fustigeromimus pubescens Da- joz (orig. des.) Madagascar [1 sp.] Fustigeropsis Raffray, 1890a: 167 Type species: Commatocerus peringueyi Raf- fray (monotypy) South Africa [2 spp.] Gericlavodes Jeannel, 1960c: 178 Type species: Clavigerodes breviceps Jeannel (orig. des.) Afrotropical [2 spp.] Gomyia Celis, 1974b:772 Type species: Gomyia monilicornis Celis (orig. des.) Reunion [1 sp.] Hexamerodes Jeannel, 1959d:709 Type species: Hexamerodes leleupi Jeannel (orig. des.) Tanzania [1 sp.] Ischyroceros Reichensperger, 1915a:8 Type species: Ischyroceros mirus Reichen- sperger (monotypy) South Africa [1 sp.] Kaisia Mann, 1920:67 Type species: Kaisia oceanica Mann (orig. des.) Fiji [1 sp.] Leptocorynotus Coulon, 1982a:55 Type species: Pseudoclavigerodesjeanneli Celis (orig. des.) Gabon [1 sp.] Marofusiger Dajoz, 1982:517 Type species: Marofusiger grandis Dajoz (orig. des.) Madagascar [1 sp.] Mesoleptochir Jeannel, 1959c:66 Type species: Mesoleptochir saegeri Jeannel (orig. des.) Afrotropical [2 spp.] Microfustigerinus Jeannel, 1 960c: 1 74 Type species: Microfustigerinus nitens Jean- nel (orig. des.) Zaire [1 sp.] Nadarimanu Mann, 1920:65 Type species: Nadarimanu alewa Mann (orig. des.) Fiji [1 sp.] Nearticerodes Jeannel, 1954a:299 Type species: Commatocerinus laevis Raffray (orig. des.) Madagascar [1 sp.] Neocorynotus Jeannel, 1960b: 198 Type species: Neocorynotus pusillus Jeannel (orig. des.) Madagascar [1 sp.] Neofustiger Bruch, 1929:157 Type species: Neofustiger cochlearis Bruch (orig. des.) Argentina [1 sp.] Neofustigerinus Jeannel, 1 960b: 1 94 Type species: Neofustigerinus lativentris Jean- nel (orig. des.) Madagascar [2 spp.] Neoradamopsis Celis, 1969a:218 Type species: Neoradamopsis coiffaiti Celis (orig. des.) Gabon, Ghana [2 spp.] Novoclaviger Wasmann, 1 894:2 1 4 (revised status; see Discussion) Type species: Novoclaviger wroughtoni Was- mann (monotypy) South Africa, Ghana [8 spp.] Fustigerodes of Raffray, 1890a: 167 (misidenti- fied, not Reitter, 1884) Novofustiger Wasmann, 1 893a: 1 06 Type species: Novofustiger raffrayi Wasmann (monotypy) Madagascar [1 sp.] Palaeoclaviger Reichensperger, 1930:379 Type species: Palaeoclaviger uelensis Rei- chensperger (orig. des.) Zaire [2 spp.] Paliger Wasmann, 1918a:402 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 65 Type species: Paliger latirostris Wasmann (monotypy) India [1 sp.] Paussiger Wasmann, 1893b:257 Type species: Paussiger limicornis Wasmann (monotypy) Madagascar [1 sp.] Platycerodes Jeannel, 1 960b: 1 93 Type species: Platycerodes trichosus Jeannel (orig. des.) Madagascar [1 sp.] Pseudacerus Raffray, 1895a:80 Type species: Pseudacerus furcatus Raffray (monotypy) Singapore [1 sp.] Pseudoclavigerodes Reichensperger, 1 9 1 5b: 1 24 Type species: Pseudoclavigerodes kohli Rei- chensperger (monotypy) Afrotropical [2 spp.] Articerodites Jeannel, 1 960c: 1 76 Type species: Articerodites ellipticus Jeannel (orig. des.) Radamopsis Reichensperger, 19 15b: 126 Type species: Radamopsis bickmanni Rei- chensperger (monotypy) Afrotropical [7 spp.] Paraclavigerodes Celis, 1 974a: 1 5 Type species: Articerodes vanboveni Jeannel (orig. des.) Rhynchoclaviger Wasmann, 1891:4 Type species: Rhynchoclaviger cremastogas- tris Wasmann (monotypy) Madagascar [1 sp.] Rynchoclaviger Raffray, 1911:175 (misspelling) Stenofustigerinus Celis, 1970:241 Type species: Stenofustigerinus franzi Celis (orig. des.) Madagascar [1 sp.] Sufifer NEW NAME (for Fusifer Raffray) Madagascar [2 spp.] Fusifer Raffray, 1900a:524 (preocc, not Dendy, 1896) Type species: Commatoceropsis perrieri Fair- maire (monotypy) Syrraphesina Raffray, 1 903a: 1 00 Type species: Syrraphesina pliciventris Raf- fray (monotypy) New Guinea [1 sp.] Triartiger Kubota, 1944:10 Type species: Triartiger urceus Kubota (orig. des.) Taiwan [1 sp.] Triceratomerus Jeannel, 1 960b: 1 95 Type species: Triceratomerus sparsepunctatus Jeannel (orig. des.) Madagascar [1 sp.] Villofustiger Leleup & Celis, 1968:409 Type species: Villofustiger gibbiceps Leleup & Celis (orig. des.) South Africa [1 sp.] Xenalluaudia Raffray, 1913a: 5 5 Type species: Xenalluaudia crematogastris Raffray (orig. des.) Kenya [2 spp.] Xenobesuchetia Leleup, 1976b:307 Type species: Xenobesuchetia perreti Leleup (orig. des.) Kenya [1 sp.] Xenocelisia Coulon, 1982a:63 Type species: Radamopsis vihambai Celis (orig. des.) Afrotropical [6 spp.] Xenoleleupia Jeannel, 1952a:90 Type species: Xenoleleupia nodicornis Jeannel (orig. des.) Zaire, Tanzania [3 spp.] Xenomussardia Celis, 1975:655 Type species: Xenomussardia besucheti Celis (orig. des.) Zimbabwe [1 sp.] Tribe DIMEROMETOPINI Dimerometopus Celis, 1970:245 Type species: Hadrophorus bicephalus Jean- nel (orig. des.) Madagascar [1 sp.] Tribe DISARTHRICERINI Disarthricerus Raffray, 1895a:78 Type species: Disarthricerus integer Raffray (monotypy) Singapore, Borneo [2 spp.] Diarthricerus Jeannel, 1949a:29 (misspelling) Tribe HOPLITOXENINI Burgeonilla Reichensperger, 1930:381 Type species: Burgeonilla polygona Reichen- sperger (orig. des.) Zaire, Cameroon [1 sp.] Hadrophorus Fairmaire, 1898b:342 Type species: Hadrophorus humerosus Fair- maire (monotypy) Madagascar [1 sp.] Xenofranzia Celis, 1969b:422 66 FIELDIANA: ZOOLOGY Type species: Xenofranzia spinipennis Celis (monotypy) Hoplitoxenus Jeannel, 1 960c: 1 73 Type species: Hoplitoxenus joannae Jeannel (orig. des.) Zaire [1 sp.] Monodiger Paulian, 1948:143 Type species: Monodiger sulcatus Paulian (orig. des.) Ivory Coast [1 sp.] Tribe LUNILLINI Lunilla Jeannel, 1957b: 110 Type species: Lunilla saprinoides Jeannel (orig. des.) Angola [1 sp.] Tribe MASTIGERINI Mastiger Motschulsky, 1851:501 Type species: Mastiger abruptus Motschulsky (monotypy) Oriental [2 spp.] Tribe MIROCLAVIGERINI Miroclaviger Wasmann, 1893a: 108 Type species: Miroclaviger cervicornis Was- mann (monotypy) Madagascar [5 spp.] Tribe NEOCERATOPSINI Neoceratopsis Jeannel, 1956f:50 Type species: Neoceratopsis pauliani Jeannel (orig. des.) Madagascar [1 sp.] Tribe NEOCERINI Subtribe NEOCERINA Neocerus Wasmann, 1893a: 105 Type species: Neocerus compressicornis Was- mann (monotypy) Madagascar [1 sp.] Tribe NEOCERINI Subtribe THEOCERINA Antalaha Jeannel, 1954a:318 Type species: Antalaha vadoni Jeannel (orig. des.) Madagascar [5 spp.] Antahala Dajoz, 1982:521 (misspelling) Theocerus Raffray, 1897a:280 Type species: Theocerus crenulatus Raffray (monotypy) Madagascar [2 spp.] Tribe RADAMINI Madara Dajoz, 1982:520 Type species: Madara myrmecophila Dajoz (orig. des.) Madagascar [1 sp.] Merinia NEW NAME (for Imerina Raffray) Madagascar [2 spp.] Imerina Raffray, 1897a:281 (preocc, not Ra- gonot, 1891) Type species: Imerina breviceps Raffray (des. Jeannel, 1954a:328) Pseudoradama Dajoz, 1982:519 Type species: Pseudoradama marojejensis Dajoz (orig. des.) Madagascar [1 sp.] Radama Raffray, 1883:230 Type species: Radama inflatus Raffray (des. Lucas, 1920:564) Madagascar [4 spp.] Radamellus Raffray, 1905:456 Type species: Radamides minutus Raffray (des. Jeannel, 1954a:324) Madagascar [3 spp.] Radamides Wasmann, 1897:261 Type species: Radamides trifoveolatus Was- mann (monotypy) Madagascar, South Africa [4 spp.] Radamira Reichensperger, 191 5a: 5 Type species: Radamira traegaordhi Reichen- sperger (monotypy) South Africa [2 spp.] Semiclaviger Wasmann, 1 893a: 1 02 Type species: Semiclaviger sikorae Wasmann (monotypy) Madagascar [1 sp.] Tribe THYSDARIINI Arnoldiella Brauns, 1 925: 1 1 7 Type species: Arnoldiella tibialis Brauns (monotypy) Zimbabwe [1 sp.] Braunsiella Raffray, 1901a:201 Type species: Braunsiella pubiventris Raffray (monotypy) South Africa [1 sp.] Eurycheiles Jeannel, 1 95 lc:28 Type species: Eurycheiles machadoi Jeannel (orig. des.) Angola, Cameroon, Zaire [1 sp.] NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 67 Soalala Dajoz, 1982:512 Type species: Soalala spinosa Dajoz (orig. des.) Madagascar [1 sp.] Thysdariopsis Jeannel, 1960b:211 Type species: Thysdarius gracilis Raffray (orig. des.) Madagascar [1 sp.] Thysdarius Fairmaire, 1 904: 1 1 7 (new name for Thysdrus Fairmaire) Madagascar [2 spp.] Thysdrus Fairmaire, 1898b:344 (preocc, not Stal, 1874) Type species: Thysdrus perrieri Fairmaire (monotypy) Trichomatosus Celis, 1970:268 Type species: Trichomatosus pusillus Celis (orig. des.) Madagascar [1 sp.] Tribe TIRACERINI Tiracerus Besuchet, 1986:262 Type species: Articerus curvicornis Westwood (orig. des.) Australia, Lord Howe Island [49 spp.] Articerus of Hope, 1845:106 (misidentified, not Dalman, 1826; see Discussion) Articeros Gemminger & Harold, 1 868:699 (mis- spelling) Extinct Taxa Most of these taxa have not been described ad- equately by present standards, and most were not placed to tribe by their authors. Tribal placement here is therefore tentative and is based on infor- mation given in the original descriptions. Al- though all of the species are reputedly extinct, some (especially those from copal deposits) may rep- resent living species (Burleigh & Whalley, 1983). See Spahr (1981) for a complete catalog of the species. Subfamily FARONINAE Tribe FARONINI Faronus Aube [Recent genus] Baltic Amber (middle Tertiary) [2 spp.] Nugaculus Schaufuss, 1890:147 Type species: Nugaculus calcitrans Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Subfamily EUPLECTINAE Tribe EUPLECTINI Euplectus Leach [Recent genus] Baltic Amber (middle Tertiary) [4 spp.] Eupsinoides Motschulsky, 1856b:26 Type species: Eupsinoides glabrellus Mot- schulsky (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Euspinoides Handlirsch, 1907:732 (misspelling) Hetereuplectus Schaufuss, 1890:156 Type species: Hetereuplectus retrorsus Schau- fuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Monyx Schaufuss, 1890:129 Type species: Monyx spiculatus Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Nugator Schaufuss, 1890:149 Type species: Nugator stricticollis Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Subfamily BATRISINAE Tribe BATRISINI Batrisus Aube [Recent genus] Baltic Amber (middle Tertiary) [2 spp.] Pantobatrisus Schaufuss, 1890:145 Type species: Pantobatrisus cursor Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Temnodera Hope, 1837:52 Type species: Temnodera testacea Hope (monotypy) Copal, ?India (probably Recent) [1 sp.] Tribe METOPHNI Hagnometopias Schaufuss, 1890:133 Type species: Hagnometopias pater Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Subfamily GONIACERINAE Tribe BRACHYGLUTINI Barybryaxis Schaufuss, 1890:121 Type species: Barybryaxis lata Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Parabryaxis Handlirsch, 1907:732 (misspelling) Eupines King [Recent genus] Kauri Resin, Australia (late Tertiary) [ 1 sp.] Rybaxis Saulcy [Recent genus] Baltic Amber (middle Tertiary) [3 spp.] 68 FIELDIANA: ZOOLOGY Tribe BYTHININI Bythinus Leach [Recent genus] Baltic Amber (middle Tertiary) [4 spp.] Type species: Greys conciliator Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Tribe TYCHINI Tychus Leach [Recent genus] Baltic Amber (middle Tertiary) [2 spp.] Subfamily PSELAPHINAE Tribe CTENISTINI Ctenistodes Schaufuss, 1890:141 Type species: Ctenistodes claviger Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Tribe TMESIPHORINI Tmesiphoroides Motschulsky, 1856b:26 Type species: Tmesiphoroides cariniger Mot- schulsky (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Tribe TYRINI Cymbalizon Schaufuss, 1890:137 Type species: Cymbalizon tyroides Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Dantiscanus Schaufuss, 1890:142 Type species: Dantiscanus costalis Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Deuterotyrus Schaufuss, 1890:131 Type species: Deuterotyrus redivivus Schau- fuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Pammiges Schaufuss, 1890:144 Type species: Pammiges spectrum Schaufuss (monotypy) Baltic Amber (middle Tertiary) [1 sp.] Tyrus Aube [Recent genus] Baltic Amber (middle Tertiary) [1 sp.] Subfamily CLAVIGERINAE Tribe CLAVIGERODINI Articerus Dalman, 1826:394 [also Recent genus; see Besuchet, 1986:261] Copal, ?Oriental (probably Recent) [1 sp.] Family PSELAPHIDAE, incertae sedis Greys Schaufuss, 1 890: 1 1 3 Taxa Based on Larvae Only Ceroncinus Silvestri, 1920:311 Type species: Ceroncinus setosus Silvestri (orig. des.) 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Transactions of the Entomological Society of London: 125-132. Winkler, A. 1925. Ergebnisse von Exkursionen auf Blindkafer in der sudlichen Herzegowina. Koleopter- ologische Rundschau, 11: 137-148. philen und Termitophilen Arthropoden. Mit Angabe NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 83 Taxonomic Index This index includes all generic and higher-level names listed in the text. Suprageneric names in current use are given in all UPPER-CASE letters; synonyms and alternate spellings are in upper- and lower- case letters, in italics. Generic names include author, and are boldfaced for currently recognized genera and given in italics for subgenera, synonyms, and invalid names. When more than one page reference is given, the page number of the main entry in the catalog is boldfaced. Abascantis Strand 59 Abascantodes Strand 59 Abascantus Schaufuss 59 Abatrisops Reitter 44 Abdiunguis Park & Wagner 2 1 Aboeurhexius Park 30 Abryxis Raffray 7. 46 Acamaldes Reitter 4 1 Acanthanops Jeannel 32 Acanthicomus Jeannel 32 Acanthobythus Normand 48 Acanthocliarthrus Leleup 32 ACETALIINA 10, 19 Acetaliini 10 Acetalius Sharp 19 Achilia Reitter 4 1 Achillia Raffray 42 Achillidia Jeannel 42 Achilliotes Jeannel 42 Achiraffraya Navas 46 Acmaeonotus Gemminger & Har- old 62 Acmoeonotus Motschulsky 62 Acolonia Casey 22 Acotreba Reitter 24 Acrocomus Raffray 50 Acrodimerus Jeannel 1 4 Actiades Lucas 25 Actiastes Casey 25 Actinoma Park 25 Actionoma Raffray 25 Actium Casey 25 Actizona Chandler 25 Acylobythus Jeannel 59 Acylopselaphus Raffray 59 Acylotyrus Jeannel 59 Adalmus Reitter 30 Adiastulus Raffray 32 Adranes LeConte 63 Adraniini 13 ADRANINI 13,63 Adrocerus Raffray 49 Adrogaster Raffray 22 Afrofustiger Jeannel 64 Afroplectaulax Jeannel 22 Afroplectidius Jeannel 22 Afroplectodes Jeannel 22 Afroplectus Jeannel 22 Afropselaphus Jeannel 62 Afrotyrus Jeannel 56 Afrozethopsus Jeannel 1 4 Agatyrus Broun 59 Agnesia Lokay 6, 56 Algodontodes Jeannel 56 Algodontus Jeannel 56 Allobatrisus Leleup 39 Allobrox Fletcher 25 Allomelba Park 25 Alloplectus Broun 25 Allotrimium Park 25 Aloxomidus Raffray 22 Alytus Hampe 8, 34 Alytus Jacoby 8 Amana Raffray 6, 32 A maurobythus Jeannel 48 Amauronyx Reitter 52 Amauropidius Jeannel 32 Amaurops Fairmaire 3 1 AMAUROPSINI 11,31 Amauropus Reitter 3 1 Ambalavoa Dajoz 28 AMBICOCERINA 11,32 Ambicocerodes Leleup 32 Ambicoceropsis Leleup 32 Ambicocerus Leleup 32 Amblycerus Raffray 6, 64 Ambrosiger Silvestri 63 Amicrops Saulcy 3 1 Aminosimus Raffray 1 9 Amphironchus PeyerimhofF 63 Amudrocerus Raffray 25 Anabaxis Raffray 7, 42 Anaclasiger Raffray 63 Anagonus Fauvel 59 Anama NEW NAME 6, 32 Anarmodius Raffray 30 Anarmoxys Raffray 42 Anasidius Jeannel 45 Anasis Raffray 6, 42 Anasopsis Raffray 7, 42 Anchylarthron Brendel 45 Ancistromus Jeannel 32 Ancystrocerus Raffray 59 Andiplectops Jeannel 27 Anitra Casey 59 Ankarahitra Jeannel 63 Ankavena Jeannel 28 Anomoplectus Jeannel 22 Anomozethus Jeannel 1 4 Anoplectus Raffray 27 Anoplobraxis Park 50 Anops Brendel 31 Anopsibythus Jeannel 48 Anotimus Dajoz 19 Anozethopsis Jeannel 1 4 Antahala Dajoz 67 Antalaha Jeannel 67 Antrobythus Besuchet 48 Apenicellus Jeannel 62 Apenicillus Reitter 62 Aphanethrix Raffray 56 Aphanogramme Dodero 27 Apharina Reitter 6 1 Apharinodes Raffray 6 1 Apharus Reitter 58 Apheloplectus Raffray 1 9 Aphilia Reitter 21 Aphiliops Reitter 25 Aphiliopsis Besuchet 2 1 Aplodea Reitter 6 1 Aploderina Raffray 58 Apobatrisus Raffray 32 Apobythus Raffray 48 Apocliarthrus Jeannel 32 Apoderiger Wasmann 63 APODERIGERINI 13,63 Apoplectus Raffray 50 Aporhexius Raffray 30 Apotectus NEW NAME 6, 28 Apoterus Raffray 2 1 Apothinus Sharp 24 Apotrabisus Jeannel 32 Apotrissemns Jeannel 44 Apozethopsus Jeannel 1 5 Arachis Raffray 46 Araucaniotes Jeannel 27 Archaeozethus Leleup 1 5 Archeuplectus Jeannel 23 Archiclaviger Heller 64 Arcobythus Jeannel 47 Areopagus Leach 6, 47 Arctophysis Reitter 3 1 Argiobythus Karaman 6, 47 Arhvtodes Reitter 7, 54 ARHYTODIMORPHI 12, 54 ARHYTODINI 12,54 Arianops Brendel 31 Arispeleops Park 31 Arnoldiella Brauns 67 ARNYI.LIINI 11.41 Arnyllium Reitter 41 Arthmius LeConte 33 Arthriclius Jeannel 37 Arthromelinus Jeannel 34 Arthromelodes Jeannel 33 Arthromelus Jeannel 33 Articerini 5, 13 Articerodes Raffray 64 Articerodites Jeannel 66 Articeronomus Raffray 64 Articeropsis Wasmann 64 Articeros Gemminger & Harold 68 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 85 Articerus Dalman 5, 64, 69 Articerus of Hope 5, 68 Arytodimorphi 12 Asanis NEW NAME 6, 42 Asymmelba Park 26 Asymoplectidius Jeannel 1 9 Asymoplectodes Jeannel 1 9 Asymoplectus Raffray 1 9 Atenisodus Raffray 46 Atherocolpus Raffray 7, 60 Atheropterodes Jeannel 33 Atheropterus Raffray 33 Atinus Horn 54 ATTAPSENIINI 12, 54 Attapsenius Bruch 54 Atychodea Reitter 7, 53 Auchenotropidius Jeannel 50 Auchenotropis Raffray 50 Aulacobythus Leleup 52 Aulacozethus Leleup 1 5 Aulaxus Raffray 22 Autoplectus Raffray 6, 28 Auxenocerini 5, 11 Auxenocerus Jeannel 3 1 Babnormodes Park 34 Badensia Jeannel 28 Bajulobythus Karaman 6 Balcanella Karaman 6 Balcanobythus Karaman 6, 47 Balega Reitter 3 1 Barada Raffray 41 Baradiini 1 1 BARADINA 11,41 Baraxina Raffray 45 Barbiera Jeannel 42 Baroxarthrius Park 38 Barroeuplectoides Park 1 7 Barm, juba Park 3 1 Barrometopia Park 4 1 BARROSELLINI 12,61 Barrosellus Jeannel 6 1 Barybryaxis Schaufuss 68 Basilewskydiella Leleup 49 Basilewskyola Jeannel 32 Basilewskyozethus Leleup 1 5 Basitrodes Jeannel 33 Basolum Casey 25 Bathybythus Besuchet 48 Batoctenus Sharp 33 Batoxyla Raffray 33 Batoxylina Jeannel 39 Batraxis Reitter 46 Batraxys Raffray 46 Batriasymmodes Park 33 Batribolbus Raffray 33 Batribraxis Schaufuss 50 Batricrator Jeannel 33 Batrictenistes Lobl 33 Batrifigia Park 33 Batrinanda Park 33 Batriphysis Park 50 Batriplica Raffray 33 Batrisaulax Jeannel 33 Batriscenaulax Jeannel 34 Batriscenellus Jeannel 34 Batrisceninus Jeannel 34 Batrisceniola Jeannel 34 Batriscenites Jeannel 34 Batriscenodes Jeannel 34 Batrischema NEW NAME 6, 34 Batrisiella Raffray 34 BATRISINA 11,32 BATRISINAE 11,31, 68 BATRISINI 11,32,68-69 Batrisinus Raffray 34 Batrisiotes Jeannel 34 Batrisobryaxis Schaufuss 50 Batrisocenus Raffray 34, 69 Batrisochaetus Jeannel 34 Batrisochorus Jeannel 34 Batrisoconnus Leleup 34 Batrisodella Jeannel 34 Batrisodellus Jeannel 34 Batrisodema Raffray 34 Batrisodes Reitter 8, 34 Batrisodinus Jeannel 34 Batrisodiola Jeannel 35 Batrisodites Jeannel 35 Batrisolius Jeannel 35 Batrisomalus Raffray 35 Batrisomellus Leleup 35 Batrisomicrus Jeannel 35 Batrisomina Raffray 35 Batrisomorpha Raffray 7, 46 Batrisomorphi 1 1 Batrisopachys Jeannel 35 Batrisophyma Raffray 35 Batrisoplatus Raffray 35 Batrisoplisus Raffray 35 Batrisopsis Raffray 41 Batrisoschema Jeannel 6, 34 Batrisoschema Reitter 35 Batrisoxenus Dajoz 6, 35 Batrisoxenus Leleup 35 Batristellus Jeannel 34 Batristerus Jeannel 35 Batristhenes Jeannel 35 Batristidius Jeannel 35 Batristilbus Raffray 35 Batristites Jeannel 35 Batristogenius Jeannel 35 Batrisus Aube 8, 35, 68 Batrivitis Park 35 Batrixenus NEW NAME 6, 35 Batrybraxis Reitter 50 Baxyridius Jeannel 42 Baxyris Jeannel 42 Berber onyx Jeannel 52 Berdura Reitter 45 Bergrothia Reitter 3 1 Bergrothiella Reitter 32 Berlara Reitter 45 Berlaraxis Jeannel 46 Besucheteidos Comellini 1 5 Besuchetidia Leleup 35 Biblomelba Park 2 1 Biblomimus Raffray 2 1 Biblomorphus Lobl 2 1 BIBLOPLECTINA 10, 19 Bibloplectinus Jeannel 1 9 Bibloplectodes Jeannel 1 9 Bibloplectopsis Jeannel 1 9 Bibloplectus Reitter 19 BIBLOPORELLINA 10, 21 Bibloporellus Jeannel 2 1 BIBLOPORINA 10, 21 Bibloporini 10 Bibloporites Jeannel 2 1 Bibloporus Thomson 2 1 Bibrax Fletcher 4 1 Biotus Casey 54 Bironia Raffray 64 Bolbobythus Raffray 47 Bolbozethus Coulon 1 5 Bontomtes Grigarick & Schuster 30 Borneana Schaufuss 36 Bothriocephalotes Leleup 22 Bothriotelus Jeannel 36 Brabaxis Raffray 7, 45 Brachygluta Thomson 42 BRACHYGLUTINA 11, 41 Brachyglutina Jeannel 42 BRACHYGLUTINI 6, 11, 41, 68 Brachyglutodes Jeannel 42 BRACHYGLUTOMORPHI 1 1 , 41 BRACHYSCELIA 10, 14 Brachyscelidae 10 Brachyscelides 10 Braunsiella Raffray 67 Braxida Jeannel 42 Braxyda Raffray 42 Bredoella Jeannel 49 Breguetiella Leleup 52 Briara Reitter 42 Briaraxis Brendel 42 Brinckidiella Leleup 56 Brounia Raffray 22 Brouniella Raffray 22 Bryaxella Raffray 42 Bryaxes 1 1 Bryaximorphi 1 1 Bryaxina Raffray 42 Bryaxinae 5 Bryaxinella Jeannel 42 Bryaxini 1 1 Bryaxis Kugelann 47 Bryaxis Leach 5, 44 Bryaxonoma Raffray 45 Bunoderus Raffray 42 Burgeonilla Reichensperger 66 Buris Fletcher 50 Byraxis Raffray 4 1 Byraxis Reitter 45 Byraxorites Jeannel 42 Bythiniama Jeannel 47 Bythinidius Jeannel 47 BYTHININA 11,47 Bvthininae 5 BYTHININI 11,47, 69 Bythinites Jeannel 6, 47 Bythinoderes Reitter 52 Bythinogaster Schaufuss 45 Bythinoides Karaman 6 Bythinomorpha Schaufuss 5 1 BYTHINOMORPHI 1 1, 47 Bythinomorphus Jeannel 47 86 FIELDIANA: ZOOLOGY Bythinophanax Reitter 52 B vthinophvsis RafFray 5 1 BYTHINOPLECTINA 5, 10, 14 BYTHINOPLECTINI 5, 10, 14 Bythinoplectoides Comellini 1 5 Bythinoplectus Reitter 1 5 Bythinopsidius Meggiolaro 47 Bythinopsis RafFray 48 Bythinus Leach 6, 47, 69 Bythiotes NEW NAME 6, 51 Bythobletus Reitter 47 Bythonesiotes Jeannel 1956 51 Bythonesiotes Jeannel 1958 6, 51 Bythoxenites Jeannel 48 Bythoxenus Motschulsky 48 Caccoplectinus Chandler & Wolda 54 Caccoplectus Sharp 54 ( alar us RafFray 22 Caligocara Park 45 Caligrua Jeannel 27 Caligruacmes Jeannel 28 Callithoracoides Strand 62 Callithorax Motschulsky 62 ( a ma Id it os Jeannel 57 Camaldosis Jeannel 57 Camaldus Fairmaire 58 Camptomidius Jeannel 36 Camptomites Jeannel 36 Camptomodes Jeannel 36 Canthoderus Motschulsky 3 1 Capnites RafFray 5 1 Catoxyomus Jeannel 36 Cedius LeConte 5, 59 Celisia Jeannel 36 Centrophtalmina RafFray 58 CENTROPHTHALMINA 12, 57 Centrophthalmina RafFray 57 ( tiitniph thai mosis RafFray 58 Centrophthalmus Schmidt-Gobel 58 ( \ii trot oma Heyden 54 CEOPHYLLINI 12,54 Ceophyllus LeConte 54 Cephalaxis Jeannel 52 Cephalozethus Jeannel 1 5 Ceratobythus Normand 48 Cercoceroides RafFray 58 Cercoceropsis RafFray 58 Cercocerulus RafFray 58 Cercocerus LeConte 58 Cerennea RafFray 1 7 Cerennia Jeannel 1 7 Ceroderma RafFray 36 Ceroncinus Silvestri 69 Cerylambus NEW NAME 6, 64 Chaetorhopalus RafFray 2 1 Chaetorrhopalus Jeannel 2 1 CHALCOPLECTINI 12, 54 Chalcoplectus Oke 54 Chasoke Chandler 59 Chenniides 12 Chenniina 12 Chenniopsis RafFray 54 Chennium Latreille 54 Chiasmatobythus Jeannel 48 ( h res to mora Jeannel 22 CHRESTOMERINA 10, 22 Circocerus Motschulsky 58 Cismelba Park 26 Clarkeorites Leleup 36 Clavidius Karaman 6 Clavifer LaPorte 63 Claviger Preyssler 63 Clavigeridae 13 Clavigerides 9, 13 CLAVIGERINAE 9, 13, 63, 69 CLAVIGERINI 13,63 Clavigerodes RafFray 64 CLAVIGERODINI 5, 13, 63, 69 Clavigeropsini 13 Clavigeropsis RafFray 64 Clermontodes Jeannel 42 Cliarthrinidius Jeannel 36 ( liarthriinis Jeannel 36 Cliarthrodes Jeannel 36 Cliarthromorphus Jeannel 36 Cliarthrus RafFray 36 Codonoderus Jeannel 4 1 Coiffaitius Karaman 64 Collacerothorax Lea 59 Collartia Jeannel 48 Colletocerus Celis 64 Comatopselaphus Schaufuss 46 Commatocerini 13 Commatocerinus Wasmann 65 Commatocerodes Peringuey 64 Commatoceropsis RafFray 7, 65 Commatocems RafFray 64 Connodontinus Jeannel 36 Connodontites Jeannel 36 Connodontus RafFray 36 Conogastridius Jeannel 36 Conoplectus Brendel 30 Conopygidia Jeannel 36 Conuridius Jeannel 36 Coptoplectus Leleup 22 Corynecerus Jeannel 44 Corynotellus Jeannel 64 Corynotopsis Jeannel 64 Corynotus Jeannel 64 Coryphomellus Jeannel 36 Coryphomodes Jeannel 1960 36 Coryphomodes Jeannel 1959 36 Coryphomoides Lobl 36 Coryphomus Jeannel 36 Couloniella Besuchet 1 5 Craspedopterus Jeannel 52 Cratna RafFray 36 Cratnodes Jeannel 36 Crenobythus Karaman 48 Cryphorhinula SchauFuss 45 Cryptorhinula Schaufuss 45 Ctenicellus RafFray 7, 54 Ctenisis RafFray 54 Ctenisodes RafFray 54 Ctenisomimus RafFray 54 ( ten isomorphic RafFray 55 Ctenisophanes Jeannel 55 ( ten i soph us RafFray 7, 55 Ctenisoschema Jeannel 55 Ctenistes Reichenbach 55 Ctenistidius Jeannel 55 CTENISTINI 12, 54, 69 Ctenistodes Schaufuss 69 ( tenistodites Jeannel 55 CTENISTOMORPHI 12,54 ( tenotillus RafFray 59 Cupila Casey 25 Curculionellus Westwood 62 Custotychus Park & Wagner 53 Cutrimia Park 25 Cyathiger King 6 1 CYATHIGERIMORPHI 12,61 CYATHIGERINI 12,61 Cyathigerodes Jeannel 6 1 Cylindrarctus Schaufuss 53 Cylindrembolus Schaufuss 46 Cylindroma RafFray 36 Cymbalizon Schaufuss 69 Cyrtoplectus Normand 23 Dacnotillus RafFray 7, 57 Daliacmes Jeannel 28 Dalma Sharp 2 1 Dalmatobythus Karaman 6 Dalmina RafFray 28 Da I mind la Jeannel 28 Dalminia NEW NAME 6, 28 Dalminiastes Jeannel 28 Dalminiola Jeannel 28 Dalminiomus Jeannel 28 Dalmisus Sharp 27 Dalmoburis Park 5 1 Dalmodes Reitter 7, 50 Dalmodiini 12 DALMODINA 12,50 Dalmomelba Park 25 Dalmomima RafFray 5 1 Dalmonexus Park 50 Da lmoph y sis RafFray 5 1 Dalmoplectus RafFray 7, 25 Dalmosanus Park 25 Dal mose 11a Casey 25 Damilina Jeannel 6, 28 Damilinidius Jeannel 28 Dantiscanus Schaufuss 69 Daveyia Lea 56 Debeckeria Leleup 49 Decarfuss NEW SUBGENUS 5, 44 Decarfuss Park 44 Decart h ron Brendel 5, 44 DECARTHRONINA 1 1, 44 Decarthronini 1 1 Decatocerus Saulcy 48 Declivodes Park 34 Dejaegeria Jeannel 64 Delamarea Jeannel 52 Delenda Croissandeau 1 8 Denicyathiger Jeannel 6 1 Deroplectus RafFray 23 Desimia Reitter 55 Desimiella Jeannel 55 Desmia Leng 55 Deuterotyrus Schaufuss 69 Diaposis RafFray 37 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 87 Diarrogus Raffray 23 Diarthricerini 13 Diarthricerus Jeannel 66 Diartiger Sharp 64 Diastictulus Raffray 1 8 Diaugis Raffray 37 Dicentrius Reitter 62 Dichocoryna Comellini 1 5 Dichozethinus Jeannel 1 5 Dicrobia Reitter 44 Dicrobiella Jeannel 42 Dicrobionoma Jeannel 42 Dicrobiotus Jeannel 45 Didimoprora Raffray 60 Didymoprora Raffray 60 Dierobia Thomson 44 DIMERINI 5, 10, 17 DIMEROMETOPINI 13, 66 Dimerometopus Celis 66 Dimerus Fiori 5, 8, 18 Dimorphozethus Jeannel 1 5 Dionyx LePeletier & Serville 55 Diplectellus Reitter 23 Diplectidius Jeannel 23 Diplomelinus Jeannel 1 5 Diroptrus Motschulsky 46 DISARTHRICERINI 13,66 Disarthricerus Raffray 66 Dissemoplectus Jeannel 23 Dranisaxa Park 42 Drasinus Raffray 5, 42 Duciola Reitter 3 1 Durbos Sharp 59 Eccoptobythus Sainte-Claire De- ville 48 Echinoplectus Jeannel 22 Echinotrabisus Jeannel 33 Echinozethus Jeannel 1 5 Ectoparyphidius Jeannel 52 Ectoparyphodes Jeannel 52 Ectoparyphus Jeannel 52 Ectopocerus Raffray 43 Ectotrabisus Leleup 37 Edaphus Motschulsky 8 Edocranes Reitter 55 Eichiella Leleup 54 Elaphidipalpus Jeannel 57 Elaphobythus Jeannel 52 Elasmatus Raffray 64 Eleodimerina Jeannel 37 Eleodimerodes Leleup 37 Eleodimerus Jeannel 37 Eleusomatus Raffray 1 9 Eleusotamus Raffray 1 9 Elytrodes Park 34 Emanuelia Blattny 41 Embolobythus Jeannel 47 Empinodes Park 35 Enantius Schaufuss 58 Endoplectus Raffray 23 Endytocera Sharp 3 1 Enoptostomus Schaum 55 Ephimia Reitter 59 Ephymata Raffray 43 Ephymia Reitter 59 Epicaris Reitter 55 Epiplectus Raffray 27 Epithematus Raffray 23 Epullea Jeannel 28 Ere mom us Raffray 45 Ergasteriocerus Leleup 43 Erichobythus Karaman 6 Etopias Jeannel 28 Euaesthetinae 8 Eubatrisus Raffray 37 Euboarhexius Grigarick & Schuster 30 Euceroncinus Kemner 69 Eudranes Sharp 59 Euglyptus Broun 2 1 Euharmophola Park 50 Eulasinus Sharp 59 Euphalepsus Reitter 37 Euphalespus Raffray 37 Euphiliops Jeannel 1 9 Eupifigia Park 45 Eupinella Raffray 43 Eupines King 45, 68 Eupinidius Jeannel 45 Eupinion Oke 2 1 Eupinoda Raffray 45 Eupinogitus Broun 43 Eupinolus Oke 46 Eupinopsis Raffray 43 Euplectamecia Park 23 Euplectellus Reitter 23 Euplecterga Park & Wagner 23 Euplectidae 10 EUPLECTINA 11,22 Euplectina Raffray 27 EUPLECTINAE 9-10, 19, 68 EUPLECTINI 6, 10, 19, 68 Euplectinus Jeannel 23 Euplectodina Raffray 1 9 Euplectoides Jeannel 23 EUPLECTOMORPHI 10, 19 Euplectomorphus Motschulsky 1 5 Euplectops Reitter 28 Euplectopsis Raffray 2 1 Euplectus Leach 23, 68 Euplenius Motschulsky 45 Euprenius Raffray 45 EUPSENIINA 11,45 Eupseniini 1 1 Eupsenina Raffray 45 Eupsenius LeConte 45 Eupsinoides Motschulsky 68 Eupsinus Motschulsky 6 Eurhexius Sharp 30 Eurycheiles Jeannel 67 Eusanops Brendel 3 1 Eusonoma Reitter 1 8 Euspinoides Handlirsch 68 Euteleia Raffray 44 Euthiomelus Jeannel 33 Eutrichites LeConte 46 Eut yphlus LeConte 2 1 Exallidius Jeannel 37 Exallomorpha Jeannel 37 Exallus Raffray 37 Excavodes Park 34 Exechophyes Jeannel 37 Exedrus Raffray 37 Exeirarthra Broun 18 Extol lodes Park 33 Facetus Schaufuss 48 Fagniezia Jeannel 43 Faliscus Casey 2 1 Fanovana Jeannel 29 Faradayus Reitter 63 Faronellus Jeannel 1 8 Faronides 10 Faronidiellus Jeannel 1 8 Faronidius Casey 1 8 Faronina 10 FARONINAE 3, 10, 14, 68 FARONINI 10, 18, 68 Faronites Jeannel 1 8 Faronitopsis Jeannel 1 8 Faro no ma Raffray 28 Faronus Aube 18, 68 Fijiastes Park 23 Filiger Schaufuss 62 Filigerinus Jeannel 6 1 Filigerodes Jeannel 6 1 Fletcherexius Park 30 Fossiger Wasmann 64 Foveoscapha Park & Wagner 30 Franziotus Leleup 59 Franzorella Leleup 37 Frontelba Park 26 Frutillariotes Jeannel 28 Fusifer Raffray 6, 66 Fusjuguma Park 53 Fustiger LeConte 7, 64 Fustigerillus Wasmann 65 Fustigerini 5, 13 Fustigerinus Wasmann 65 Fustigerodes of Raffray 7, 65 Fustigerodes Reitter 7, 65 Fustigeromimus Dajoz 65 Fustigeropsis Raffray 65 Gabata Raffray 23 Gamba Schaufuss 6-7, 31 Gasola Reitter 3 1 Gasterotropis Raffray 57 Gastrobothrus Broun 46 Geopselaphus Jeannel 62 Gerallus Sharp 6, 59 Gericlavodes Jeannel 65 G ermai nites Jeannel 3 1 Ghesquierites Jeannel 43 Glapharsenus Raffray 7, 50 Glastus Raffray 27 Globa Raffray 50 GLOBINA 12,50 Globobythus Karaman 48 Globosulus Jeannel 6, 50 Globulites Jeannel 6, 50 Glyphobythoides Jeannel 47 Glyphobythus Raffray 47 Glyphozethopsis Jeannel 1 5 88 FIELDIANA: ZOOLOGY Glyphozethus Jeannel 1 5 Glyptosoma Motschulsky 57 Gnesion Raffray 5 1 Gnorosus Raffray 55 Golasa Raffray 1 8 Golasidius Jeannel 1 8 Golasina Jeannel 1 8 Golasites Jeannel 1 8 Gomyia Celis 65 Gonatocerus Schaufuss 42 Goniacerides 5, 11 Goniacerina 1 1 GONIACERINAE 5, 1 1, 41, 68 GONIACERINI 12,49 Goniaceroides Raffray 49 Goniacerus Motschulsky 5, 49 Goniastes Westwood 5. 49 Goniastini 5, 11 Goniomellus Jeannel 52 Goniozethus Jeannel 1 5 Gonorsus Jeannel 56 Grammoplectus Jeannel 23 Greys Schaufuss 69 Haasellia Wagner 25 Haasiella Park 25 Hadrophorus Fairmaire 66 Hagnometopias Schaufuss 68 HALORABYXINA 1 1 , 45 Halorabyxis Jeannel 45 Hamotidius Jeannel 58 HAMOTINA 12,58 Hamotini 12 Hamotocellus Raffray 58 Harriot oides Schaufuss 58 Hamotopsis Raffray 59 Hamotulus Schaufuss 58 Hamotus Aube 58 Hanfordia Park 25 Haploplectus Jeannel 1 9 Harmomima Raffray 50 Harmophola Raffray 50 Harmophorus Motschulsky 4 1 Harmophorus Schaufuss 6, 50 Hatchia Park & Wagner 2 1 Hemicliarthrus Jeannel 37 Hendecameros Comellini 1 5 Heptaleptus Jeannel 1 5 Heptameron Comellini 49 Herminiella Blattny 56 Hesperorychus Schuster & Marsh 53 Hetereuplectus Schaufuss 68 Heteronyx Saulcy 30 Hexamerodes Jeannel 65 Hexirhexius Grigarick & Schuster 30 Hingstoniella Jeannel 37 Hispanisella Park 25 Histricoidius Jeannel 38 Hologlyptus Fairmaire 54 Holozodini 12 Holozodinus Jeannel 54 Holozodoides Dajoz 54 Holozodus Fairmaire 54 HOPLITOXENINI 13,66 Hoplitoxenus Jeannel 67 Hornia Raffray 60 Horniella Raffray 60 Hughia Raffray 1 7 Hulstaertites Jeannel 37 Humbertella NEW NAME 6, 19 Humbertiella Jeannel 6, 19 HYBOCEPHALINI 12, 61 Hybocephalodes Raffray 6 1 Hybocephalus Motschulsky 6 1 Hybonotus Jeannel 6, 37 Hylorychus Grigarick & Schuster 53 Hynneophorus Leleup 55 Hyobontus NEW NAME 6, 37 Hypochraeus Raffray 7, 37 Hypoparyphantus Jeannel 52 Hypoplectinus Jeannel 20 Hypoplectus Raffray 20 Hypotrabisus Jeannel 38 Iberobythus Franz 47 Illyrobythus Karaman 47 Imerina Raffray 6, 67 Imerinella Jeannel 29 Imeriniella Jeannel 29 IMIRINI 12,52 Imirus Reitter 52 Imtempus Reitter 52 Indiella Blattny 56 INIOCYPHINA 12,50 Iniocyphini 12 Iniocyphus Raffray 50 Insulomodes Park 50 Intempus Raffray 52 Ipsimodes Jeannel 49 Ipsimus Reitter 49 Ischyroceros Reichensperger 65 Itamus Raffray 45 Iteticus Raffray 37 Itombworites Jeannel 57 JANUSCULINA 12, 59 Janusculus Cerruti 59 Jeannelia Raffray 15 Jochmansiella Leleup 37 JUBINI 5, 11,31 Jubinini 1 1 JUBOMORPHI 11,31 Jubomorphus Raffray 3 1 Jubus Schaufuss 6, 31 Jugobythus Karaman 6 Juxtahamotopsis Park 59 Kaisia Mann 65 Kenocoelus Broun 27 Kigatrodes Jeannel 37 Kistneriella Leleup 49 Korovodes Park 37 Kunzea Leach 47 Kuscheliotes Jeannel 3 1 La bom im us Sharp 60 Laphidioderomimus Leleup 55 Laphidioderus Raffray 55 Lasinus Sharp 60 Latomelba Park 25 Leanymus Raffray 56 Leiochrotella Jeannel 43 Leiochrotidius Jeannel 46 Leiochrotina Jeannel 6, 51 Leleupia Jeannel 37 Leleupiastes Jeannel 37 Leleupiella Jeannel 5 1 Leleupiozethus Coulon 1 5 Leleupites Jeannel 49 Lemelba Park 25 Leptachillia Jeannel 46 Leptobatrisus Jeannel 38 Leptobythus Jeannel 48 Leptocorynotus Coulon 65 Leptoctenistes Jeannel 55 Leptomicrus Jeannel 6,17 Leptophiliops Jeannel 20 Leptoplectus Casey 23 Leptorrachis Jeannel 46 Leptotyrus Jeannel 59 Leptozethodes Jeannel 1 5 Lethenomus Raffray 60 Leupelia Jeannel 40 LEUPELIINA 11,40 Limoniates Raffray 20 Linderia Saulcy 47 Liniolis Grigarick & Schuster 7, 24 Lioplectus Raffray 23 Listriophorini 12 Listriophorus Schaufuss 49 Loebliella Leleup 38 Loebliobythus Leleup 52 Loebliozethus Coulon 1 5 Lophobythus Raffray 47 Lucifotychus Park & Wagner 53 Lukwangulorites Leleup 38 Lunilla Jeannel 67 LUNILLINI 13,67 MACHADOINI 13,62 Machadoites Jeannel 57 Machadous Jeannel 62 Machaerites Miller 48 MACHAERITINA 11,48 Machaerodes Brendel 47 Machulkaia I .obi 5 1 Macrobythus Raffray 47 Macrodelphus Leleup 38 Macroplectus Raffray 22 Macroraffravia Jeannel 29 MACROSCELIA 12,54 Macroscelidae 12 Macroscelides 12 Macta Raffray 3 1 Madabaxyris Dajoz 43 Madara Dajoz 67 Madontalgus Dajoz 56 Madrasorites Jeannel 38 Malleecola Oke 43 Malleoceps Park 25 Manniconnus Park 38 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 89 Manuleiger Jeannel 6 1 Marellus Motschulsky 60 Margaris Schaufuss 62 Marnax LaPorte 41 Marofusiger Dajoz 65 Masoala Jeannel 29 Mas tiger Motschulsky 67 MASTIGERINI 13,67 Maya Blattny 7, 51 Mayetia Mulsant & Rey 8, 19 MAYETIINI 10, 19 Mayetinae 10 Mayetini 10 Mechanicus Schaufuss 52 Mecochelia Motschulsky 6 1 Mecynozethus Jeannel 1 5 Megabatrus Lobl 38 Megalobythus Jeannel 47 Megarafonus Casey 1 8 Melba Casey 26 Melbamima Raffray 26 Meliceria Raffray 23 Mentraphus Sharp 62 Merinia NEW NAME 6, 67 Mesoleptochir Jeannel 65 Mesoplatus Raffray 30 Mesoplectus Karaman 6 Mestogaster Schmidt-Gobel 6 1 Mestogastridius Jeannel 6 1 Mestogastrina 12 Metactenistes Jeannel 55 Metamayetia Coiffait 1 9 Metaplectodes Jeannel 23 Metaxis Motschulsky 43 Metaxoides Schaufuss 6 1 Metazethopsis Jeannel 16 Methorius Raffray 20 Metopias Gory 41 Metopiasini 1 1 Metopiellus Raffray 4 1 METOPIINI 11,41,68 Metopioides Schaufuss 49 Metopiosoma Raffray 4 1 Metopioxys Reitter 4 1 Mexigaster Park 22 Mexiplectus Park 22 Micrapoderiger Jeannel 63 Microbatrisodes Jeannel 38 Microbatrisus Jeannel 38 Microbythus Karaman 48 Microdiartiger Sawada 64 Microfustigerinus Jeannel 65 Microgmocerus Leleup 49 Microphthartus Blattny 8 Microplectus Raffray 20 Microtyrus Broun 2 1 Microzethinus Jeannel 1 6 Microzethopsis Jeannel 1 6 Microzethus Jeannel 1 6 Mildana Jeannel 29 Mimoplectus Raffray 52 Mina Raffray 6, 38 Minibi Grigarick & Schuster 26 Minitiola Karaman 1 9 Mipseltyrus Park 60 Mirellus Raffray 27 Mirini 12 Miroclaviger Wasmann 67 MIROCLAVIGERINI 13, 67 Mirus Saulcy 52 Mitona Raffray 46 Mitracephala Raffray 29 Mitracephalini 1 1 MITRAMETOPINA 1 1, 29 Mitrametopus Raffray 29 Mnia NEW NAME 6, 38 Moczarskia Blattny & Blattny 24 Monodiger Paulian 67 Monyx Schaufuss 68 Morana Sharp 5 1 Morius Casey 24 Mothaus Jeannel 58 Nadarimanu Mann 65 Nafonus Schuster & Marsh 1 8 Namunia Reitter 52 Nanorafonns Schuster & Marsh 1 8 Narcissiella Jeannel 2 1 Narcodes King 55 Neactium Jeannel 26 Nearticerodes Jeannel 65 Nedarassus Raffray 5 1 Nenemeca Raffray 38 Neobrachygluta Kaszab & Szekessy 42 NEOCERATOPSINI 13,67 Neoceratopsis Jeannel 67 NEOCERINA 13,67 NEOCERINI 13,67 Neocerus Wasmann 67 Neocommatocerus Jeannel 65 Neocorynotus Jeannel 65 Neodalmus Raffray 26 Neodeuterus Schaufuss 53 Neofustiger Bruch 65 Neofustigerinus Jeannel 65 Neopalimbolus Oke 60 Neophamisus Reitter 7, 58 Neoplectidius Jeannel 20 Neopselaphaulax Jeannel 62 Neopselaphus Jeannel 62 Neoradamopsis Celis 65 NeorafTrayia Jeannel 29 Neosampa Broun 30 Neosintectes Jeannel 57 Neothesiastes Jeannel 20 Neotrabisus Jeannel 38 Neotyrus Raffray 60 Neozethopsus Jeannel 1 6 Nesiotomina Jeannel 38 Nesiotoplectus Jeannel 23 Nesiotozethus Jeannel 16 Nesiotyrodes Jeannel 60 Nicothaeus Raffray 2 1 Nicotheus Casey 2 1 Nipponobythus Jeannel 5 1 Nisa Casey 7, 42 Nisaxis Casey 43 Nodibythus Karaman 6 Noduliceps Jeannel 43 Nodulina Raffray 6, 43 Nondulia NEW NAME 6, 43 Nothoplectus Jeannel 28 Notogmocerus Leleup 49 Novissimus Blattny 6 1 Novoclaviger Wasmann 7, 65 Novofustiger Wasmann 65 Nugaculus Schaufuss 68 Nugator Schaufuss 68 Obricala Raffray 47 Ocabajara Schuster & Grigarick 1 7 Ocabaraja Schuster & Grigarick 1 7 Ocarabaja Schuster & Grigarick 1 7 Octomeros Comellini 1 6 Octomicrellus Jeannel 6, 17 Octomicrini 5, 10 Octomicrites Jeannel 1 7 Octomicrus Schaufuss 5, 8, 17 Octozethinus Jeannel 1 6 Octozethodes Jeannel 1 6 Octozethus Jeannel 1 6 ODONTALGINI 12, 56 Odontalgus Raffray 56 Odonticoscapus Jeannel 38 Odontobythus Karaman 48 Odontoconnus Jeannel 38 Ogmoceridius Leleup 49 Ogmocerodes Jeannel 49 Ogmoceropsis Jeannel 49 Ogmocerus Raffray 49 Omobathus Raffray 2 1 Omotimellus Jeannel 20 Omotimiotes Jeannel 20 Omotimus Raffray 20 Ophelius Raffray 38 Opisthosemus Jeannel 2 1 Opisthosphaera Jeannel 1 6 Orientamaurops Karaman 32 Orientobythus Karaman 6 Orlandia Comellini 1 7 Oropodes Casey 23 Oropus Casey 30 Oropygia Raffray 38 Oropygiina 1 1 Orropygia Raffray 38 Oxarthrius Reitter 38 Oxygastrobythus Leleup 53 Oxyomera Raffray 38 Oxyomites Jeannel 38 Oxyzethodes Jeannel 1 6 Oxyzethus Jeannel 1 6 Pachyeuplectus Jeannel 20 Pachygastrodes Leleup 56 PACHYGASTRODINI 12, 56 Pachygastrodirius Leleup 56 Pachypygidia Jeannel 38 Pachytrabisus Jeannel 38 Pachyzethopsus Jeannel 1 6 Palaeoclaviger Reichensperger 65 Paliger Wasmann 65 Palimbolus Raffray 60 Palodea Jeannel 60 Pammiges Schaufuss 69 90 FIELDIANA: ZOOLOGY Panabachia Park 43 PANAPHANTINA 1 1, 24 Panaphantus Kiesenwetter 24 Panaphysis Reitter 38 Panaramecia Park 23 Pantobatrisus Schaufuss 68 Pantosiella Jeannel 38 Parabatrisus Jeannel 39 Parabryaxis Handlirsch 68 Parabythinns Jeannel 47 Parabythus Karaman 47 Paraclavigerodes Celis 66 Paracliarthrus Jeannel 35 Paractium Jeannel 26 Paracyathiger Jeannel 6 1 Paradrasinus NEW SUBGENUS 5, 43 Paradrasinus Park 43 Parafaronus Jeannel 1 8 Paragoniastes Comellini 49 Paraleleupia Jeannel 37 Paramaurops Jeannel 32 Paraphiliopsis Jeannel 20 Paraplectus Raffray 28 Parapteracmes Jeannel 28 Parapyxidicerus Sawada 1 7 Pararticerus Jeannel 64 Parasimus Jeannel 49 Parastectenis Jeannel 55 Paratrabisus Jeannel 33 Paratychus Besuchet 53 Parazethopsus Jeannel 1 6 Parepullea Jeannel 29 Pareuplectops Jeannel 27 Pareuplectus Jeannel 23 Parmipalpus Broun 6 1 Paryphantellus Jeannel 53 Paryphopterus Jeannel 53 Pasiglyptus Berg 54 Passosiella Leleup 39 Patranus Raffray 45 Patreus Broun 20 Paussiger Wasmann 66 Pedinopsis Raffray 6, 43 Pedisinops NEW NAME 5-6, 43 Pelulea Jeannel 4 1 Peluleotes Jeannel 4 1 Perimelba Park 1952 26 Perimelba Park 1943 26 Periplectus Raffray 20 Petaloscapus Jeannel 39 Petalozethopsis Jeannel 1 6 PETANOPINI 1 2, 56 Petanops Jeannel 56 PHALEPSINI 13,62 Phalepsoides Raffray 39 Phalepsus Westwood 62 Phalespoides Raffray 39 Phalespus Raffray 62 Phamisulus Reitter 58 Phamisus Aube 3 1 Philiopsis Raffray 20 Philoscotus Sawada 1 9 Philotrimium Blattny 27 Philus Saulcy 25 Phoberus Raffray 46 Phormiobius Broun 60 PHTEGNOMINA 11,29 Phtegnomini 1 1 Phtegnomus Raffray 29 Phthartomerus Sawada 1 8 Phthartomicrus Schaufuss 53 Phybytharsis Park 5 1 Physa Raffray 46 Physobryaxis Hetschko 46 Physomerinus Jeannel 39 Physoplectus Reitter 43 Picrogyne Gozis 47 Pilactium Grigarick & Schuster 26 Pilopius Casey 55 Placodium Broun 20 Plagiophorus Motschulsky 7, 39 Planityphlus Park 2 1 Plattibythus Karaman 47 Platycerobythus Jeannel 53 Platycerodes Jeannel 66 Platyplectus Jeannel 29 Plectasymus Jeannel 19 Plectiastes Jeannel 20 Plectodytes Jeannel 20 Plectomorphus Raffray 22 Plectophleoides Karaman 6 Plectophleus Karaman 23 Plectophloeus Reitter 6, 23 Plectoprotus NEW NAME 6, 24 Plectostenus Lea 22 Plectrobythus Leleup & Celis 43 Plectusodes Raffray 22 Plesiotyrus Broun 60 Plocamarthrus Jeannel 39 Podus Raffray 4 1 Poeciloceras Comellini 16 Poroderopsis Jeannel 55 Poroderus Sharp 56 Prespelea Park 53 Prionobythus Jeannel 48 Probatrisus Raffray 33 Proboscites Jeannel 1 6 Procheilophorus Leleup 39 Prodalma Raffray 20 Promayetia Coiffait 1 9 Prophilus Raffray 26 Proplectus Raffray 25 Prorhexius Raffray 30 Prosagola Raffray 18 Prosthecarthron Raffray 5 1 Protamaurops Miiller 32 PROTERINI 12,52 PROTEROMORPHI 12, 52 Proterus Raffray 53 Protoplectus Leleup 6, 24 Protoplectus Raffray 22 Protozethopsus Jeannel 1 6 Protozethus Jeannel 1 6 Pselaphaulax Reitter 62 I'm- la phi- 1 lus Raffray 62 PSELAPHIDAE 10. 14 Pselaphidea 10 Pselaphides 9, 10 Pselaphidius Jeannel 62 Pselaphii 10 PSELAPHINAE 9, 12,54 PSELAPHINI 13,62 Pselaphinus Jeannel 63 Pselaphischnus Raffray 63 Pselaphocerodes Jeannel 57 Pselaphocerus Raffray 57 Pselaphocompsus Raffray 58 Pselaphodes Westwood 60 Pselaphodinus Jeannel 62 Pselaphogenius Reitter 62 PSELAPHOMORPHI 13,62 Pselaphomorphus Motschulsky 3 1 Pselaphophus Raffray 63 Pselaphopsis Jeannel 63 Pselaphoptrus Reitter 63 Pselaphorites Jeannel 63 Pselaphostomus Reitter 63 Pselaphotheseus Park 63 I Nc la phot rich us Besuchet 63 Pselaphotychus Khnzorian 53 Pselaphoxys Raffray 63 Pselaphus Herbst 63 PSELAPTINA 1 1 , 45 Pselaptrichus Brendel 48 Pselaptus LeConte 46 Pseudacerus Raffray 66 Pseudachillia Jeannel 43 Pseudactium Casey 24 Pseudamaurops Jeannel 32 Pseudambicocerus Leleup 32 Pseudapharina Raffray 62 Pseudeuplectus Enderlein 29 Pseudfustiger Reitter 64 Pseudobatriscenus Jeannel 39 Pseudobatrisus Raffray 39 Pseudobaxyris Jeannel 43 Pseudobryaxis Motschulsky 69 Pseudocamaldes Jeannel 43 Pseudoclavigerodes Reichensperger 66 Pseudocliarthrus Jeannel 39 Pseudoconnus Leleup 39 Pseudoctenistes Leleup 39 Pseudofustiger Raffray 64 Pseudohamotus Raffray 58 Pseudomotimus Jeannel 20 Pseudophanias Raffray 62 Pseudoplectus Reitter 20 Pseudoradama Dajoz 67 Pseudoterus Raffray 53 Pseudotrabisoides Jeannel 39 Pseudotrimium Raffray 26 Pseudotychus Raffray 60 Pseudotyrus Raffray 6 1 Pseudozibus Jeannel 2 1 Psilocephalus Raffray 62 Pteracmes Raffray 28 Pteracmidius Jeannel 28 PTERACMINI 11, 27 Pteroplectus Raffray 28 Pubimodes Park 34 Pycnoplectus Casey 24 Pygmactium Schuster & Grigarick 26 PYGOXYINI 12,50 Pygoxyon Reitter 50 Pyreneobythus Jeannel 47 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 91 Pvtna Casey 6 1 PYXIDICERINA 10, 17 Pyxidicerini 5, 10 Pyxidicerinus Jeannel 1 7 Pyxidicerus Motschulsky 17 Pyxidion Comellini 16 Quadrelba Park 1952 26 Quadrelba Park 1942 26 Rabyxis Raffray 43 Racemia NEW GENUS 7, 24 Radama Raffray 67 Kudu melius Raffray 67 Kudu m ides Wasmann 67 RADAMINI 13,67 Radamira Reichensperger 67 Kuclumopsis Reichensperger 66 Raffrayella Blattny 46 Kuffruyiu Reitter 29 Raffrayidius Jeannel 29 RAFFRAYIINI 11, 28 Raffrayina 1 1 Kuf fruyites Jeannel 29 Raffrayitidia Jeannel 29 Raffrayola Jeannel 29 Raffrayolus Jacobson 44 Rafonus Casey 1 8 Rafrhexius Park 29 Rafrhisis Park 24 Ramecia Casey 7, 24 Ramelbida Park 26 Rameloidea Park 26 Ranavala Raffray 29 Ranavalidius Jeannel 29 Ranavalini 1 1 Ranavalodes Jeannel 29 Raphitreodes Jeannel 57 Raphitreodes NEW GENUS 5, 57 Kap bit reus Sharp 57 Raxybis Raffray 43 Reichenbachella Jeannel 43 Reichenbachia Leach 44 Reichenbachiina 1 1 Reichenbachius Casey 44 Reitteronymus Raffray 2 1 Rhamophorus NEW NAME 6, 50 Rhenanus Wusthoff 8 Rhexidius Casey 30 RHEXIINA 11,29 Rhexinexus Park 30 Rhexini 1 1 Rhexinia Raffray 30 Rhexiola Park 30 Rhexius LeConte 29 Rhinoscepsii 1 1 RHINOSCEPSINA 11, 24 Rhinoscepsis LeConte 24 Rhinosceptis Reitter 24 Rhynchoclaviger Wasmann 66 Rhynchoplectus Jeannel 20 Rhynoscepsis Raffray 24 Rhynosceptis Raffray 24 Rhytus Westwood 7, 54 Riphaterus Jeannel 57 Rougemontiella Leleup 44 Ruacorites Jeannel 39 Rugegius Jeannel 5 1 Rybaxidia Jeannel 44 Rybaxis Saulcy 8, 44, 68 Rynchoclaviger Raffray 66 Rytus King 7, 60 Ryxabis Westwood 39 Sagola Sharp 1 8 Sagolonus Raffray 27 Salagosa Raffray 1 8 Salpictomelus Jeannel 53 Saltisedes Kubota 58 Sunipsu Raffray 27 Sandersonella Park 26 Sathytes Westwood 39 Saulcyella Reitter 26 Saxet Grigarick & Schuster 24 Scaiomelus Jeannel 33 Scaioscenellus Jeannel 34 Scalenarthrus LeConte 46 Schaufussia Raffray 60 SCHISTODACTYLINI 12, 56 Schistodactylus Raffray 56 Schizocoryna Comellini 1 6 Schweigeria Lobl 32 Scotoplectus Reitter 24 Scottia Jeannel 1 7 Sebaga Raffray 3 1 Seleneuplectus Jeannel 24 Selenozethus Jeannel 1 6 Selinozethus Jeannel 1 6 Semiclaviger Wasmann 67 Seracamaurops Winkler 32 Seydelites Jeannel 39 Shingala Raffray 18 Silillicus Lucas 46 Simini 12 Simkinion Park & Pearce 44 Simplona Casey 26 Simus Raffray 49 Singhala Raffray 1 7 Sinistrocedius NEW SUBGENUS 5,59 Sinistrocedius Park 59 Sintectes Westwood 57 Sintectodes Reitter 57 Siteromina Lobl 39 Smilestethus Cerruti 57 Sou lulu Dajoz 68 Sogaella Jeannel 6, 51 Sognorites Jeannel 56 Sognorus Reitter 6, 56 Somatipion Schaufuss 56 SOMATIPIONINI 12, 56 Sonoma Casey 1 8 Spelaeobythus Lobl 48 Speleobama Park 53 SPELEOBAMINI 12,53 Speleochus Park 48 Speleodes Park 33 Speleoplectus Minozzi 30 Speobatrisodes Jeannel 39 Speobaxyris Jeannel 44 Spifemodes Park 35 Spilorhombus Raffray 60 STAPHYLINIDAE 2, 8 STAPHYLINOIDEA 2 Startes Broun 44 Stectenidius Jeannel 56 Stectenis Raffray 56 Stenocliarthrus Leleup 39 Stenofustigerinus Celis 66 Stenoplectus Raffray 24 Stenosagola Broun 7, 18 Stenosthetus Griffith & Pidgeon 63 Stethotaphrus NEW NAME 6, 58 Stictus Raffray 4 1 STILIPALPINA 11,41 Stilipalpus Jeannel 4 1 Stipesa Sharp 62 Stratus Schaufuss 3 1 Strombopsis Raffray 44 Strongylomus Jeannel 38 Subterrochus Park 48 Subulipalpus Schaufuss 60 Sufifer NEW NAME 6, 66 Sulcifigia Park 39 Sunorfa Raffray 5 1 Sunorfoides Raffray 5 1 Synbibloporus Sawada 2 1 Syntectes Jeannel 57 Syntectodes Reitter 57 Syrbatidius Jeannel 40 Syrbatodes Jeannel 40 Syrbatomorphus Jeannel 39 Syrbatus Reitter 39 Syrmocerus Raffray 40 Syrraphesina Raffray 66 Takaorites Jeannel 5 1 Tamotus Schaufuss 8 TANYPLEURINA 12, 50 TANYPLEURINI 9, 12, 50 Tanypleurus Raffray 5 1 Taphrophorus Schaufuss 55 Taphrostethus Schaufuss 6, 58 Tapiloites Jeannel 57 Tasmanityrus Chandler 60 Tecnesis Peyerimhoff 55 Tecnesites Jeannel 55 Tectenis Jeannel 55 Temnodera Hope 68 Termitotyrus Borgmeier 60 Tetrads Sharp 55 Tetraglyptinus Jeannel 54 Tetraglyptus Jeannel 54 Tetrameres Schaufuss 8 Tetrascapha Schuster & Marsh 30 Tetrasemus Jeannel 47 Tetratarsus Schaufuss 8 Texamaurops Barr & Steeves 40 THEOCERINA 1 3, 67 Theocerini 13 Theocerus Raffray 67 Thesiastes Casey 24 Thesiectus Park 24 Thesium Casey 24 92 FIELDIANA: ZOOLOGY THYSDARIINI 13,67 Thysdariopsis Jeannel 68 Thysdarius Fairmaire 68 Thysdrus Fairmaire 68 Tiomomus Jeannel 20 TIRACERINI 5, 13,68 Tiracerus Besuchet 5, 68 Tmesiphorellus Jeannel 56 TMESIPHORINI 12, 56, 69 Tmesiphorinus Leleup 57 Tmesiphorites Jeannel 57 Tmesiphoroides Motschulsky 69 Tmesiphoromimus Lobl 57 Tmesiphorus LeConte 57 Tomoplectus Raffray 26 Trabisellus Jeannel 33 Trabisidius Jeannel 33 Trabisina 1 1 Trabisinus Jeannel 33 Trabisodema Jeannel 40 Trabisoides Jeannel 40 Trabisomorphus Jeannel 33 Trabisonoma Jeannel 33 Trabisorites Jeannel 33 Trabisostenus Jeannel 40 Trabisotus Jeannel 33 Trabisoxenus Leleup 40 Trabisus Raffray 33 Tremissus Besuchet 44 Trepacmes Jeannel 28 Triangusella Park 25 Triartiger Kubota 66 Tribasodema Jeannel 40 Tribasodes Jeannel 40 Tribasodites Jeannel 40 Tribatus Motschulsky 44 Triceratomerus Jeannel 66 Trichobythus Dodero 48 Trichomatosus Celis 68 Trichonomorphus Raffray 40 TRICHONYCHINI 12, 52 Trichonycina 12 Trichonyides 12 Trichonyni 12 Trichonyx Chaudoir 52 Trichopnites Dajoz 5 1 Trichopselaphus Jeannel 63 Trigonoplectus Bowman 24 Trimicerus Motschulsky 27 Trimii 1 1 TRIMIINA 11, 25 Trimioarcus Park 26 Trimiodina Raffray 26 Trimiodytes Raffray 27 TRIMIODYTINA 11, 27 Trimiomelba Casey 26 Trimiomicrus Raffray 52 Trimiomorphus Raffray 26 Trimiophanes Jeannel 20 Trimioplectus Brendel 24 Trimiopsis Reitter 27 Trimiosella Raffray 27 Trimiovillus Park 27 I rimium Aube 27 Triomicrus Sharp 5 1 TRISIGNINA 11,30 Trisignis Park & Schuster 30 I risinarthrus Jeannel 40 Trisiniotus Jeannel 40 Trisinus Raffray 4 1 Trissemellus Jeannel 44 Trissemidius Jeannel 44 Trissemites Jeannel 44 Trissemodes Jeannel 44 Trissemosus Jeannel 44 Trissemus Jeannel 44 Trizethopsis Dajoz 1 6 Trogaster Sharp 30 Trogasteropsis Dodero 30 TROGASTRINA 11,30 TROGASTRINI 11,29 Troglamaurops Ganglbauer 32 Trogobythus Dodero 48 Tropeogaster Jeannel 57 Tropidamaurops Jeannel 32 Trychioplectus Raffray 22 Trymalius Fairmaire 63 Tuberoplectus Park 1 8 TYCHINI 12,53,69 Tychobythinus Ganglbauer 48 Tychobythus Raffray 48 Tychoides Karaman 53 TYCHOMORPHI 12, 53 Tychomorphus Jeannel 53 Tychotyrus Broun 7, 58 Tychus Leach 53, 69 Typhlacetalius Jeannel 19 Typhlobatrisus Jeannel 40 Typhlobythus Jeannel 47 Typhlokahusia Jeannel 2 1 Typhloleptodes Jeannel 1 9 Typhloleptus Jeannel 1 9 Typhloproboscites Jeannel 1 6 Typhloraffrayia Jeannel 29 Typhlorites Jeannel 40 Typhlozethus Jeannel 1 6 Tyraphus Sharp 63 Ty rides 12 TYRINA 12, 59 TYRINI 12,57,69 Tyrodes Raffray 60 Tyrodites Jeannel 60 Tyrogatunus Park 60 Tyrogetus Broun 60 Tyromorphus Raffray 6 1 Tyropsis Saulcy 6 1 Tyrus Aube 61, 69 Ulugurubythus Leleup 53 Upoluna Schaufuss 58 Urozethidius Jeannel 1 6 Vadoniotus Jeannel 6 1 Vadonites Jeannel 49 Valda Casey 52 Valdiini 12 VALDINI 1 2, 52 Verabarolus Park 24 Vertelba Park 26 Vert ici not us Brendel 45 Vestitrichus Park 49 Vidamodes Broun 20 Vidamus Raffray 7, 27 Villofustiger Leleup & Celis 66 Whitea Hutton 22 Xenadiastus Jeannel 40 Xenalluaudia Raffray 66 Xenobasilewskyia Leleup 40 Xenobatrisus Wasmann 40 Xenobesuchetia Leleup 66 Xenobryaxis Jeannel 46 XENOBYTHINA 11,48 Xenobythus Peyerimhoff49 Xenocelisia Coulon 66 Xenocliarthrus Jeannel 40 Xenoconurus Jeannel 40 Xenodesimia Jeannel 55 Xenofranzia Celis 66 Xenogmocerodes Leleup 49 Xenogyna Raffray 29 Xenogynidia Jeannel 29 Xenolejeunea Leleup & Celis 4 1 Xenoleleupia Jeannel 66 Xenomachadoella Leleup 40 Xenomussardia Celis 66 Xenopygia Jeannel 40 Xenopygiella Jeannel 40 Xenotmesiphorus Leleup 57 Xerhius Raffray 3 1 Xherius Raffray 3 1 Xiphobythus Jeannel 44 Xybarida Raffray 46 Xybaris Reitter 46 Zeatyrus Sharp 6 1 Zelandius Raffray 22 Zethinellus Jeannel 1 5 Zethini 10 Zethiniotus Jeannel 1 7 Zethinomorphus Jeannel 1 7 Zethinus Raffray 1 7 Zethopsimorphus Jeannel 1 7 Zethopsina 5, 10 Zethopsinidius Jeannel 1 7 Zethopsinus Jeannel 1 7 Zethopsiola Jeannel 1 7 Zethopsiolites Jeannel 1 7 Zethopsoides Jeannel 1 7 Zethopsus Reitter 1 7 Zethus Schaufuss 1 7 Zibus Saulcy 27 Zolium Casey 27 Zonaira Grigarick & Schuster 27 Zosimus Schaufuss 6, 7, 59 Zoufalia Reitter 32 Zozimus Raffray 59 NEWTON & CHANDLER: CATALOG OF PSELAPHIDAE 93 Field Museum of Natural History Roosevelt Road at Lake Shore Drive Chicago, Illinois 60605-2496 Telephone: (312) 922-9410 UNIVERSITY OF ILLINOIS URBANA 590 5FIN.S C001 FIELOIANA : ZOOLOGY. $ NEW SERIES SCHGO 40-54 1988-89 3 0112 009378735