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Preliminary 
Amphibian and Reptile Survey 

of the 
Lewis and Clark National Forest: 1994 



A Report to: 

USDA Forest Service 

Lewis and Clark National Forest 

1101 15th Street N. 

Box 869 

Great Falls, MT 59403 



Submitted by 
JAMES D.REICHEL 

March 1995 

Montana Natural Heritage Program 

1515 East Sixth Avenue 

P.O. Box 201800 

Helena, MT 59620-1800 



1995 Montana Natural Heritage Program 



This document should be cited as follows: 

Reichel, J. D. 1 995. Preliminary amphibian and reptile survey of the Lewis and Clark National Forest: 1 994. 
Montana Natural Heritage Program. Helena, MT. 92 pp. 



ABSTRACT 

A total of 40 surveys and several additional sightings were made in the Lewis and Clark National 
Forest (L&CNF) between May and September 1994. Mostwere surveys of ponds, lakes, seeps, 
streams or other wetlands, made by 1 or 2 individuals. Each survey took 1 0-200 person-minutes and 
consisted of a thorough search of the wetland perimeter and netting of near shore aquatic habitats for 
adults, eggs, larvae, and tadpoles. Stream sampling was done by hand and dipnet. Seeps were 
checked by rolling over rocks and logs in and near wet areas. In addition to surveys, sightings were 
made from road kills, vocal identifications or fortuitous sightings by other reliable individuals. 

Localized areas across the entire forest were covered in the survey, with specific locations 
determined largely by priorities set by Mike Enk, Fisheries Biologist for the Lewis and Clark National 
Forest. 

Six amphibians are reported from the L&CNF: Long-toed Salamander {Ambystoma 
macrodactylum). Tailed Frog (Ascaphus truei). Western Toad (Bufo boreas). Northern Chorus Frog 
(Pseudacris triseriata). Northern Leopard Frog (Ranapipiens), and Spotted Frog (Ranapretiosa) . 
The Spotted Frog was the most widespread amphibian throughout the forest. The Tailed Frog and 
Long-toed Salamander were found in a few locations on the Rocky Mountain Ranger District 
(RMRD). The Western Toad was found in very small numbers on the RMRD in 1994. Historically it 
has also been reported in the Highwood, Little Belt, and Crazy Mountains; however it was not found 
during our surveys. Given the apparent region-wide declines in this species, all sightings should be 
reported. The Northern Leopard Frog was reported historically from a single site on the RMRD and 
several sites in the Highwood Mountains on the L&CNF; none were found on the L&CNF in this 
survey. It also has been reported historically from numerous prairie ponds, outside and at lower 
elevations than L&CNF lands. Northern Leopard Frogs are nearly extirpated from western Montana, 
and anecdotal evidence indicates a decline elsewhere in Montana (except the southeast corner); all 
sightings should be reported. The Northern Chorus Frog was common in prairie ponds, primarily 
outside and lower in elevation than L&CNF lands. Four other prairie-inhabiting amphibians have been 
recorded in the area, though in some cases well away from L&CNF lands; these include the Tiger 
Salamander (Ambystoma tigrimim). Great Plains Toad (Bufo cognatus), Woodhouseis Toad (Bufo 
woodhousii), and Plains Spadefoot (Scaphiopus bombifrons). 

Eleven reptiles have been reported from near the L&CNF, but only two have been definitely 
reported from on the forest: the Western Terrestrial Garter Snake (Thamnophis elegans) and Common 
Garter Snake (Thamnophis sirtalis). Both were found in the RMRD and Little Belt Mountains. The 
Western Terrestrial Garter Snake also was found in the Big Snowy Mountains and there is a specimen 
record of the Common Garter Snake from the Highwood Mountains. The Rubber Boa (Charina 
bottae) and Racer (Coluber constrictor) have been recorded on the Helena NF just to the south and 
west of the L&CNF and therefore are likely to be found on the L&CNF as well. The Spiny Softshell 
(Trionyx spinifera) is present in large rivers at lower elevations; records include the Musselshell River 
near Shawmut and perhaps Canyon Ferry Reservoir This species could also occur on the Smith or 
Judith Rivers. The following reptiles have been recorded in the area and may eventually be found on 



in 



lower elevation L&CNF lands: Painted Turtle {Chrysemys picta). Short-horned Lizard (Phrynosoma 
doiiglasi). Western Hognose Snake (Heterodon nasiciis). Gopher Snake (Pituophis catenifer). Plains 
Garter Snake {Thamnophis radix), and Western Rattlesnake (Crotahis viridis). 



n 



TABLE OF CONTENTS 

ABSTRACT i 

ACKNOWLEDGMENTS vii 

INTRODUCTION 1 

METHODS AND MATERIALS 2 

RESULTS AND DISCUSSION 3 

Species known to be present on the Lewis and Clark National Forest 6 

Long-toed Salamander (Ambystoma macrodactylum) 6 

TailedV vog (Ascaphus tniei) 8 

WestevnToad (Bufo boreas) 10 

Western ChovusVvog(Pseiidacristriseriata) 12 

Northern Leopard Frog (i?a«a/»zp/e«5) 14 

SpottedVvog (Ranapretiosa) 16 

Western Terrestrial Garter Snake (Thamnophis elegans) 18 

Common Garter Snake (Thamnophis sirtalis) 20 

Species Potentially Present on the Lewis and Clark National Forest 22 

Tiger Salamander (Ambystoma tigrinum) 22 

Great^lamsToad (Biifo cognatus) 24 

Woodhouseis Toad (Bufowoodhousii) 26 

Plains Spadefoot (Scaphiopus [=SpeaJ bombifrons) 28 

^amtedTurtle (Chrysemys picta) 30 

Spiny Softshell {Trionyx spiniferus) (=Apalone spinifera) 32 

Sl\ort-\iomed\Azard {Phrynosomadouglasi) 34 

^uhher ^oa{Charmabottae) 36 

^acer {Coluber constrictor) 38 

Western Hognose Snake (Heterodon nasicus) 40 

Gopher Snake {Pituophis catenifer [=melanoleuciisJ) 42 

Vlams Garter Snake {Thamnophis radix) 44 

Western Rattlesnake (Crototev/ntZ/.s') 46 

Rocky Mountain Division Information 47 

Rocky Mountain Division 47 

Jefferson Division Information 47 

Highwood Mountains 47 

Little Belt Mountains 48 

Castle Mountains 48 

Crazy Mountains 48 

Big Snowy Mountains 48 

Little Snowy Mountains 49 



RECOMMENDATIONS 50 

BroLIOGRAPHY 51 

Appendix 1 . Data Sheets used for Reptiles and Amphibian Surveys and Observations 65 

Appendix2. Sites surveyed during 1994 amphibian and reptile surveys 67 

Appendix 3 . Amphibians and reptiles observed during surveys of the Lewis and Clark National 

Forest in 1993-94 71 

Appendix 4. Amphibians and reptiles reported from in and around the Lewis and Clark National 

Forest 74 



M 



ACKNOWLEDGMENTS 

We thank Mike Enk for his help throughout the study. Additional help, location of possible survey 
sites, information on herp observations, and other support was provided by Seth Diamond, Portia 
Jelinek, DavePilliod, Phillip Whitechey, Tom Whitford, and other Forest Service personnel. D. D. 
Dover, C. Jones, K. Jurist, and S. Thweatt assisted with element occurrence and map preparation. 
Financial support for the proj ect came from the Lewis and Clark National Forest (U.S. Forest Service, 
Northern Region) and the Montana Natural Heritage Program (Montana State Library, Natural 
Resources Information System and The Nature Conservancy). 

Museum records were received from: American Museum of Natural History, Academy of Natural 
Science, Brigham Young University, California Academy of Science, Carnegie Museum, University of 
Puget Sound Museum, Field Museum of Natural History, Glacier National Park Museum, Illinois 
Natural History Survey, University of Kansas, Los Angeles County Museum, Louisiana State University 
Museum of Zoology, Museum of Comparative Zoology - Harvard, Milwaukee Public Museum, 
Montana State University Museum, Michigan State University Museum, North Carolina State Museum 
of Natural History, Northern Louisiana University Museum, University of Colorado Museum, University 
of Georgia Museum of Natural History, University of Idaho Museum, University of Michigan Museum, 
University of South Dakota, United States National Museum of Natural History, University of Texas - 
Arlington, University of Texas - El Paso, and Peabody Museum - Yale. Most museum data were 
received with the help of Dr Charles Peterson, Idaho State University, Pocatello. 



vn 



INTRODUCTION 

Many amphibians are apparently declining in the western U. S. and world-wide (Corn and Fogelman 
1984, Phillips 1990, Yoffe 1992). Acid rain, ozone depletion, pollution by toxic chemicals and heavy 
metals, predation and/or competition by exotic species, habitat alteration, climate change, disease, 
immune system problems, and some combination of these factors have all been suggested as possible 
causes (Com and Fogelman 1984,Phimps 1990, Yoffe 1992). 

Bass have been introduced into waters on or near the Lewis and Clark National Forest (L&CNF) 
and have been implicated in declines of native amphibian populations in some areas. Past forestry 
practices and large scale logging continue to be detrimental to resident herpetofauna (Bury etal. 1991). 
The Tailed Frog (Ascaphiis tniei), present on the L&CNF, is thought to be one of the most sensitive 
indicators of stream-side and aquatic community health in forested landscapes (R. B. Bury, pers. 
comm.). Preliminary data indicate the Northern Leopard Frog {Ranapipiens) has disappeared over 
much of its former range in western Montana and is declining in at least some areas of eastern Montana. 
The US Fish and Wildlife Service now lists the Western Toad {Biifo boreas) as a Candidate (C-2) 
species in Colorado, Wyoming and New Mexico. Apparent declines have recently been reported in 
northern Idaho (C. Peterson pers. comm.), northwest Montana (Werner and Reichel 1 994), 
Yellowstone National Park (Peterson et al 1 992), Wyoming, and Colorado (Carey 1 993). 
The U. S. Fish and Wildlife Service lists two Montana amphibians and two reptiles as candidate 
species: the Spotted Frog (C2) {Ranapretiosa), Tailed Frog (C2), Short-horned Lizard {Phrynosoma 
doiiglasi) andNorthern ^a.gQhnx^\\lAzax6i{Sceloporns graciosus graciosus). The U.S. Forest 
Service Region 1 lists the Coeur diAlene Salamander {Plethodon idahoensis) as iSensitivei and is 
considering adding several other amphibians. The Montana Natural Heritage Program and the Montana 
Department of Fish, Wildlife and Parks list 6 amphibians [Coeur diAlene Salamander, Idaho Giant 
Salamander (Dicamptodon aterrimus). Tailed Frog, Canadian Toad (Biifo hemiophrys). Spotted 
Frog, Wood Frog (Rana sylvatica)] and 7 reptiles [Snapping Turtle (Chelydra serpentina). Spiny 
Softshell {Apalone spiniferd). Short-homed Lizard, Sagebmsh Lizard, Westem Hognose Snake 
(Heterodon nasiciis). Smooth Green Snake (Opheodrys vernalis). Milk Snake (Lampropeltis 
triangulum)] as species of special concem in the state. The Northern Leopard Frog and Westem 
Toad (Biifo boreas) are being considered for addition to the list. Eight of these species, the Tailed 
Frog, Western Toad, Spotted Frog, Northem Leopard Frog, Spiny Softshell, Short-homed Lizard, 
Western Hognose Snake, and Milk Snake occur or potentially occur on the L&CNF. 



METHODS AND MATERIALS 

Historic locations of amphibians and reptiles were recorded from literature (see Bibliography) and 
museum specimen records. Records were received from over 20 maj or museums which have 
computerized their collections in North America (see Acknowledgments). Locations derived from these 
sources have been entered into a database and digitized. Records from the Museum of Vertebrate 
Zoology, Berkeley, California, have yet to be received. 

Survey sites were chosen based on 4 criteria: 1) high priority sites as determined by the L&CNF; 
2) location of streams, seeps and wetlands on topographic maps; 3) accessibility of the wetlands by 
roads or hiking trails; and 4) conversations with district biologists regarding stream-seep-wetland 
locations and past Forest Service surveys. Based on the above, 3-6 sites were chosen daily for 
surveys. Ten minutes to 2 hours were spent at each site, depending upon the size of the area and what 
was found. Initially, the entire shoreline, or a maj or part thereof, was searched by walking slowly along 
the edge and up into the surrounding vegetation, including rolling over rocks and logs. At regular 
intervals, the aquatic habitat was sampled for tadpoles or larvae using dipnets. If the initial sampling 
showed amphibian/reptile species present, further effort was expended in order to get some idea of 
abundance and distribution. 

An attempt was made to capture at least the first few individuals of a species seen at a survey site. 
The species name was recorded along with developmental stage and sex (if possible); the animals were 
then released. Representative samples of the more common species in an area were preserved for 
permanent museum records and are deposited at the Idaho State University Museum. Water 
temperature, air temperature, and a general description of the area were recorded. Standard data 
sheets used during this proj ect are given in Appendix 1 ; the amphibian survey data sheet was developed 
by U. S. Fish and Wildlife Service and is used extensively by a variety of researchers in the western U. S. 
Much site-specific data was gathered during these surveys; not all data has been analyzed or is 
presented in this report, but is available from the Montana Natural Heritage Program. 



RESULTS AND DISCUSSION 

A total of 40 sites were surveyed of which 22 had one or more amphibian or reptile species present 
(Figure 1 , Appendices 2 and 3). Although no species were found at 1 8 sites, their absence may have 
been due to the time of day, weather conditions, or other factors at the time of sampling. With three 
exceptions, all of the sites were on L&CNF land. 

In addition to the 40 surveys, there were a number of sightings (i.e. road kills, chance observations) 
for which data are available and the sightings considered reliable. Species location data from surveys, 
chance encounters, and historic records (from the literature and museum specimens) are listed in 
Appendix 4. Distribution maps were created using survey and sighting data and historical records; inset 
statewide maps for each species are based on sight and specimen records, both recent and historic. 

No previous publications or reports on reptiles or amphibians concentrate on the L&CNF area. 
There is a publication on the Tiber Reservoir area to the north (Mosimann and Rabb 1952). Based 
museum specimens, publications, surveys and incidental observations, six amphibians and two reptiles 
have been reported on the L&CNF; an additional four amphibians and nine reptiles may eventually be 
found to occur there. Six amphibian and two reptile species were actually observed during the study. 
The following results are presented as a species summary for the Forest as a whole, followed by 
specific information on each division (and mountain range within the Jefferson Division). 

In the following species accounts, the section on i Similar SpeciesT covers species only which are 
known or suspected to occur in Montana; outside Montana other confusing species may occur which 
are not covered in this report. Photos of all Montana amphibians and reptiles may be found in Reichel 
and Flath (1995). 



Herp Survey Locations on or near the Lewis & Clark National Forest 




Survev locstlooi itvrn the Moritana NaturEl Merttage Pmgrsm, 2f2SfBB 



atiMakmfinJhiUi.tantti 



OcDurrences of Am by stoma macrodactylum on or near the Lewis & Clark National Forest 




^' Pra'1993 data 

V Musflum specimeriB 



Species locatlorKa from the Montana NaturBl Heritage P^ogramx 2i29i9b 



MMimmHtty.env 



Species known to be present on the Lewis and Clark National Forest 

Long-toed Salamander (Ambystoma macrodactyhim) 

Description : Adults are dark gray to black with an irregular (and sometimes broken) green to yellow 
stripe down the middle of the back. Adult snout-vent length varies from 2 to 3 .25 " . All 
salamanders have smooth moist skin without scales. 

Eggs and Larvae: Egg masses are typically laid in small clusters of 5-100 eggs but may be laid 
singly (Nussbaum etal. 1983). Within the clear gelatinous eggs, the embryos are somewhat light- 
colored, while frog and toad embryos are dark (except in Tailed Frogs). Larval Long-toed 
Salamanders are typically brown- or gray-colored, are found in ponds, have three external gills, and 
are relatively small (<1 .75" snout-vent) and slender They are distinguished from Tiger Salamander 
larvae by the 9-13 gill rakers on the inside of the 3rd gill arch (17-22 rakers on the Tiger 
Salamander); they are also smaller and lack the large head and mouth. 

Similar species: Adult Long-toed Salamanders can be distinguished from Coeur dIAlene Salamanders 
by the longest toe on the hind foot which is longer than the sole and a yellow throat patch. Long- 
toed Salamanders lack a groove running vertically from nostril to mouth. 

Habitat and Habits: Long-toed Salamanders are found in a wide variety of habitats from sagebrush to 
nearly alpine. They breed in ponds or lakes (very rarely in slow moving streams), usually those 
without fish present. Adults go to the breeding ponds immediately after snow-melt and are usually 
the earliest breeding amphibians in western Montana. In the Pacific Northwest, eggs hatch in 3 -6 
weeks and metamorphosis occurs after 2-14 months (Nussbaum etal. 1983, Leonard etal. 
1 993). Long-toed Salamanders were found in only two locations on the L&CNF. The earliest 
observation was of three egg masses nearly ready to hatch and about 500 recently-hatched larvae in 
a marshy, backwater pond off Wood Creek on 27 May 1 994. A mid-sized larva was also seen in a 
beaverpondinWagnerBasinon5 July 1994. Individuals were found only in the Rocky Mountain 
Ranger District (RMRD) from 4600 - 5720 ft. elevation. They occurred with the Spotted Frog in 
the Wood Creek pond. 

Surveying: Larvae can readily be seen in ponds during the day and sampled with a dipnet; egg masses 
may beharderto see. During the breeding season, adults may alsobe seen in the water, particularly 
during night surveys. During the rest of the spring, summer and fall, adults may occasionally be 
found in and under logs on the forest floor Metamorphosed individuals are active at night, 
particularly when it is warm and rainy; they may be captured at this time by either night searches or 
pitfall traps. 

Status: The Long-toed Salamander is the most common salamander in western Montana. Preliminary 
indications are that the Long-toed Salamander is uncommon and locally distributed in the RMRD of 
the L&CNF, the eastern edge of its range. 

MontanaNatural Heritage Program rank: G5 S5. 



Occurrences of Ascaphys truet on or near tha Lewis St Clark NatJonal Forest 



*- 





* 1S93-9* data 
^ Pre- 1393 date 



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$pE!i?h^ |q4;:aTiiDrk^ iKtm ih« M<MiTflne NdTunal H^Hiag^ PrE^gram, 2il3l^5 



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Tailed Frog (Ascaphus truei) 

Description : Adults are gray or brown with gray, brown, or occasionally yellow blotches; the skin has a 
distinctly bumpy texture. The adult has a snout-vent length of 1 .5-2" and lacks a tympanum. The 
outer toe of the hind foot is broader than the other toes. The male has a bulbous itailT which acts as 
a penis. 

Eggs and Larvae: Approximately 50 eggs are laid in rosary-like strings attached to the underside 
of rocks. The tadpole (up to 2" long) is unique in that it has a large mouth modified into a sucker; 
the color is quite variable. 

Similar species : No other frog or toad has the outer toe of the hind foot broader than the other toes; all 
other frogs and toads have a tympanum behind each eye. 

Habitat and Habits: Tailed Frogs are found in and along small, swift, cold mountain streams. In the 
L&CNF, they have been found only on the RMRD in four locations (Appendix 3, 4). The 
elevations range from 4900 feet in Falls Creek to about 6000 feet in a small stream on the Mount 
Wright trail. This elevation range will likely increase with additional sightings from the area. In the 
Cascade Mountains of Washington and Oregon, the Tailed Frog appears to be very sensitive to 
siltation and frequently disappears in and downstream from clearcuts and water diversions (Bury, 
pers. comm.). Preliminary findings do not indicate that this is the case in Montana. Eggs are laid 
during the late summer and take approximately 4 weeks to hatch. Tadpoles take 1 -4 years to 
metamorphose, depending on water temperature (Nussbaum et al 1983; Metter 1 967). Sexual 
maturity in Montana is attained at ages 6-7, (Daugherty and Sheldon 1982) which is the latest age 
for sexual maturity of any North American amphibian. 

Surveying: Tadpoles are frequently found while electro-shocking fish. They may also be found by 
turning over rocks in rapid water with a net held just downstream. Adults are best found by walking 
up streams starting 30-60 minutes after dark. 

Status: The Tailed Frogs in this report are the northeastern-most records for the species. Tailed Frogs 
are known from a few other locations east of the Continental Divide in the Beaverhead National 
Forest. While Tailed Frogs should be considered a species with a very localized distribution on the 
RMRD of the L&CNF, it may be more common and widespread in suitable habitat than is currently 
known. It is common and widespread in western Montana. Currently it is a USFWS Candidate 
species (C-2). 

Montana Natural Heritage Program rank: G5 S3 S4. 



Occurrences of Buio biireaa on or near the Lewis Sl Clark National Forest 



* 193334 data 
Pte-1993 data 
"MT MusftLjm fipsclinans 



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Species Ideations from The Montana Natural Hsritag^a F'rDgrBm. 2/3S/9Si 



■tLatAn ipfH q ? -cdi p 



Western Toad {Bufo boreas) 

Description: Adults are colored with a gray, brown, or olive-green mottling and a prominent white or 
yellowish line down the center of the back; very young transformed toads typically lack the dorsal 
line, and the warts are often red-brown in color. The pupils are horizontal. The adult has a body 
length of 2.5-5 ". There are no cranial crests and the skin is relatively dry with many warts and 
glands present. 

Eggs and Larvae: Eggs are laid in long, clear, double strings, and each has a black embryo. 
Tadpoles are typically j et black, while all the Montana frog species tadpoles are green or bronze 
(except for some Tailed Frogs). 

Similar species: Other Montana toads have cranial crests between their eyes. The Plains Spadefoot 
has one tubercle on the sole of the hind feet, a vertical pupil, and smoother skin. NOTE: It is very 
difficult to distinguish among the four Montana toad species in recently-transformed toadlets. 

Habitat and Habits: Adults are largely terrestrial and found in a variety of habitats from valley bottoms 
to high elevations; they breed in lakes, ponds, and slow streams with a preference for shallow areas 
with mud bottoms. Breeding and egg laying in Montana usually takes place 1 -3 months after snow- 
melt, from April at lower elevations to July at higher sites. Western Toads were reported from only 
two locations on the L&CNF in 1994. We found eggs in a beaver pond on a backwater of the 
Teton River on 26 May 1994; one clutch was about half developed the other two recently laid. 
Mike Enk reported a single adult along East Fork Woods Creek. Small tadpoles were seen on 2 1 
June 93 at Pine Butte Swamp Preserve. Tadpoles are typically 2-3 months old at metamorphosis in 
Montana, depending on water temperature (Black 1970). Following metamorphosis, hundreds of 
small toads, many with the tails still present, can be found on the shores of breeding ponds. 

Surveying: Tadpoles are easily seen in ponds during the day and can be sampled with a dipnet. During 
the breeding season, adults may be seen in the water but at other times are found in more terrestrial 
habitats. 

Status: Tadpoles and eggs of the Western Toad were observed at only one site during the 1994 survey 
in the L&CNF; adults were seen at only a single additional site. None were seen in the Highwood, 
Little Belt, or Crazy Mountains, although historic records exist from all three ranges (Appendix 3,4). 
The rarity of this species on the RMRD and lack of recent sightings in the Jefferson Division is of 
concern. The U. S. Fish and Wildlife Service now lists this species as a Candidate (C-2) species in 
Colorado, Wyoming, and New Mexico. Apparent declines have recently been reported in northern 
Idaho (C. Peterson pers. comm.), northwest Montana (Werner and Reichel 1994), Yellowstone 
National Park (Peterson etal 1992), Wyoming, and Colorado (Carey 1993). We would 
recommend that all sightings of this species be reported and that a monitoring program be set up. 

Montana Natural Heritage Program rank: G4 S4. 



10 



Occurrences of Pseudacris trieerjala on qt near the Lewis & Clark National Forest 

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• 199394 data 

^ Pra^lSSa {lata i^ 

j^ Musaum GpaclirMns 

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SpKJU lociptlons fr^iTt tHp ManTa^a NatLihai Harltagft Pvogfatn, 2tlQI^B 



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Western Chorus Frog (Pseudacris triseriata) 

Description: Adults are very small (0.75-1.5") and have tiny, almost unnoticeable toe pads. They have 
a dark line extending from the snout through the eye to the groin. Basic coloration is quite variable 
with the background color being green, brown, gray, orreddish. Typically 3-5 dark longitudinal 
stripes are present on the head and back which may be broken up into spots on some individuals. 
Eggs and Tadpoles: Eggs are laid in small clusters of 1 0- 1 00, usually less than 1 " across and 
attached to submerged vegetation (Wheeler and Wheeler 1 966, Baxter and Stone 1985). 
Individual eggs are about 1 mm in diameter. Tadpoles are brown/bronze and the eyes are located 
on the sides of the head. 

Similar species: Pacific Chorus Frogs {Pseudacris regilla) have obvious toe pads and an eye stripe 
ending at the shoulder Recently metamorphosed Ranid frogs could be confused with this species 
but the coloration differs and the tiny toe pads are lacking. 

Habitat and Habits: Western Chorus Frogs are regularly found in the water only during the breeding 
period in spring. Their presence is obvious during this time due to their call which is given frequently 
at night and sporadically throughout the day. Calls were heard on or near the L&CNF from mid- 
May through early June while doing surveys; however, the precise beginning and end of calling in the 
area is unknown. Following breeding, these frogs move into adjacent uplands and are rarely seen. 
In eastern Montana they breed in temporary ponds and small lakes surrounded by prairie; in some 
locations in Montana they are also found in open forested habitats. Eggs hatch in about 2 weeks 
and tadpoles are about 2 months old at metamorphosis (Wheeler and Wheeler 1 966, Nussbaum et 
al. 1983). 

Surveying: Adults are easily surveyed for, using their calls for identification during the breeding season 
in the spring and early summer During the breeding season, adults may also be seen in the water, 
but their small size and habit of freezing or diving when disturbed makes observation difficult; night 
surveys may be more productive. Egg masses are difficult to find. Tadpoles may be seen in ponds 
during the day and can be sampled with a dipnet. 

Status: Common throughout the prairies of eastern Montana. Probably common at the lower edges of 
the L&CNF, especially where open forest or prairie occurs; perhaps uncommon and local within 
forested habitat at higher elevations. Mike Enk reported two locations with Western Chorus Frogs 
on the L&CNF. These included the headwaters of Arrow Creek in the Highwood Mountains and a 
seep area near Whiterock Creek in the RMRD; an additional historic record exists from lower 
Highwood Creek (Appendix 4). Numerous others were heard calling just outside the forest 
boundary. 

Montana Natural Heritage Program rank: G5 S5. 



12 



Occurrfincss of Rana pipjens on or near the Lewis & Clark National Forest 

Q 1 




• 1993-94 dnti 

^ Pr&-1993 data ^' 

V Museum specimens 



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Spflciu locBtJons from th« Montana Natural Heritage Fnogram, 2JlSfS5 



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Northern Leopard Frog (Ranapipiens) 

Description: Adults are brown or green with large, dark spots surrounded by light-colored halos on the 
sides and back. The dorso-lateral folds (ridges along the sides of the back) are usually lighter in 
color that the surrounding background. The under-side is typically white, but may be cream- 
colored or yellowish. The adult has a body length of 2-5". Newly transformed froglets may lack 
spots and are about 1 " in length (Leonard etal. 1993). 

Eggs and Tadpoles: Eggs are laid in 2-5 " globular masses composed of hundreds to thousands of 
eggs (Hammerson 1982a, Nussbaume^a/. 1983). The tadpoles are brown to dark brown on top 
with some metallic flecking, whereas the underside is often nearly transparent (Nussbaum etal. 
1983). Total length of tadpoles may reach more than 3"; the eyes are located on top of the head. 

Similar species: None, although some newly-transformed froglets may lack spots, which makes them 
extremely difficult to distinguish from Spotted and Wood Frogs. 

Habitat and Habits : Northern Leopard Frogs are found in or near water in non-forested habitats. 
Vegetation is typically dense, as in a cattail marsh or dense sedge-meadow. Breeding takes place in 
lakes, ponds (temporary and permanent), springs, and occasionally backwaters or beaver ponds in 
streams. In Colorado, eggs hatch in 4-1 5 days and tadpoles take 8-15 weeks to metamorphose, 
depending on water temperature (Hammerson 1982a). 

Surveying: Both adults, tadpoles, and eggs are easily seen in and along the water during the day and 
can be sampled with a dipnet; adults may also be captured by hand. At very low densities adults 
may be difficult to find and may be detected using a call recorder Tadpoles are difficult to tell from 
those of the Spotted Frog in areas where the two species may overlap. 

Status: Historically, the Northern Leopard Frog was widespread in Montana but it now appears to be 
extinct throughout much of the western part of the state. It is still common and widespread in the 
southeastern comer of the state, but its status is uncertain in central and northeastern Montana. It 
appears that only localized populations are present on the western edge of the plains. A single 
historic location is known from on the RMRD of the L&CNF; that museum specimen was collected 
in 1958 from the i Sun River, 5500feet.i Although it is impossible to pinpoint the exact locality for 
this record, wetlands in the immediate area should be surveyed. Specimens were collected in upper 
Highwood Creek in the Highwood Mountains in 1 962; although searches were conducted in that 
area during this study, no Northern Leopard Frogs were found. Several other historic records exist 
from near L&CNF lands just to the south and east of the Little Belt Mountains. More recent 
records include: 1) a single individual seen during a two-days period at Pine Butte Swamp in 1993; 
2) adults seen along Belt Creek north of L&CNF lands in 1994; and 3) adults seen along the 
Dearborn River in 1 993 . Due to its significant decline and lack of current reports from the L&CNF, 
all sightings of this species should be documented. 

Northern Leopard Frogs are now absent from many other areas in North America where they 
were common a few decades ago. Widespread extinctions are known from Alberta (Koonz 1 993), 
Wyoming, Colorado (Hammerson 1982b, Com andFogelman 1984), Idaho (Groves and Peterson 
1992), Washington, and Oregon (Leonard etal. 1994). Bullfrog and fish introductions, acid rain, 
ozone depletion, immune system suppression, and iPostmetamorphic Death Syndromei have all 
been suggested as causes for frog extirpations in other areas (Corn and Fogelman 1 984, 
Hammerson 1982b, Carey 1993, Leonard e^ a/. 1993). 

Montana Natural Heritage Program rank: G4 S4. 



14 



Occurrences of Rana pretiosa on or near the Lewis Si Clark National Forest 



• 1933-34 data 
^ Mussum spsclmsns 





"\j^ 




k 



\ 




Sp»cl«s locations from thi« Montana Natural Harltaga Pro<grann, 2/19/95 



■t I ■ b' [TIBf^.lc ID jC [TV 



Spotted Frog (Ranapretiosd) 

Description: The adult has a snout-vent length of 2-4" . Adults are dark to light brown, gray, or olive 
green with dark spots (frequently with lighter centers) found on the back, sides and legs. The 
number and pattern of spotting is quite variable. The back and sides are often covered with small 
bumps. The underside of the legs is bright red, salmon, or orange; this bright color may extend up 
to the chin or be replaced by a light, mottled gray on the chin, chest, and/or belly. In younger 
subadults, bright leg color is often lacking and instead a light, lemon-colored wash is present. In 
these subadults there is often a dark mask present, with a light jaw stripe extending to the shoulder; 
both the mask and jaw stripe may be less obvious in larger, older animals. 
Eggs and Tadpoles: Eggs are laid in large, globular masses of 1 50-500 at the surface of the 
water. The tadpoles are dark green to brown on top with some gold flecking whereas the underside 
has an iridescent bronze or silver color. Total length of tadpoles may reach 3 " ; the eyes are located 
on top of the head. 

Similar species: The bright-colored pigment on the undersides of the adultis legs distinguish this species 
from all other frogs in Montana. Younger individuals, without colored legs, may usually be 
distinguished from other frogs by a combination of 1) dorsal spots usually present but not 
surrounded by light-colored halos; 2) dorsolateral folds present; 3) toes without pads at the tips; 
and 4) a pale gray, (rather than white) belly. 

Habitat and Habits: Spotted Frogs are regularly found at the waterls edge in openings within forest 
habitats. Wetlands in or near treeline are also used, but populations are uncommon in the large, 
open intermountain valleys. The Spotted Frog was commonly found on both divisions of the 
L&CNF from just above the prairie edge at 4860 ft. to over 7400 ft elevations near timberline. 
Individuals were found in every type of wetland habitat, although numbers varied widely from one to 
25 or more per site. Breeding takes place in lakes, ponds (temporary and permanent), springs, and 
occasionally backwaters or beaver ponds in streams. All the egg masses in a particular pond are 
often found in the same location at the margin of the pond; therefore, the eggs are susceptible to 
drying up if pond levels recede substantially before hatching. Both eggs and recently hatched- 
tadpoles were found 27 May 94 at a pond near Wood Creek. Tadpoles were seen from the 
earliest survey on 27 May 94 to the latest survey on 9 July 94; at that date they seemed a few 
weeks from metamorphosis. Eggs hatch in 2-3 weeks and tadpoles take 2-14 months to meta- 
morphose, depending on water temperature (Nussbaum etal 1983, Turner 1958). Young and adult 
frogs often disperse into marsh and forest habitats but are not usually found far from open water 

Surveying: Adults, tadpoles, and eggs are easily seen in and along the water during the day and can be 
sampled with a dipnet; adults may also be captured by hand. Many adults may leave the breeding 
ponds following egg laying and move to nearby feeding areas for the summer Tadpoles are difficult 
to distinguish from those of the Northern Leopard Frog in areas where the two species overlap. 

Status: The most common frog on the L&CNF and in western Montana. It was observed on the 
RMRD, and the Little Belt, Crazy, and Castle Mountains of the Jefferson Division, however few 
breeding location are known at this time and, if found, should be reported. It was not seen in the 
Highwood or Snowy Mountains; any sightings from those mountain ranges should be reported. The 
species is currently a U. S. Fish and Wildlife Service Category 2 Candidate species in Montana; 
elsewhere in its range it is listed as a C-1, with Threatened/Endangered status warranted but 
precluded by work on higher priority species (U.S. Fish and Wildlife Service 1993). Significant 
declines are known from the southern end of the range (Nevada, southern Idaho, Utah). While 
significant declines are also apparent in coastal Washington (McAllister et al. 1 993), Oregon, and 
California, recent (as yet unpublished) research indicates that those populations are actually a 
different species. 

Montana Natural Heritage Program rank: G4 S4. 

16 



Occurrences of Thamnophia elegans on or near the Lewis & Clark National Forest 



1) 




# 



"A,^-^ 



«• 




* 1993-34 date 
O PiB.1^Sl3 data 
V Museum specimens 




* 



1 



* 






7 








'^sJ- 







^* 



* 





t 



T 



Sfhscias locitjons from ttis M'Dntar>B Natural HBritaQie Progrsm, 2i2SiS^ 



atninmptMiB.einf 



Western Terrestrial Garter Snake {Thamnophis elegans) 

Description: Adult Western Terrestrial (or Wandering) Garter Snakes are smaller inbody size than the 
Common Garter Snake, their length varying from 1 8-43 " . Three yellow longitudinal stripes are 
present (one dorsal, two lateral), but the dorsal stripe is much narrower than that of the Common 
Garter Snake. A distinctive feature of the Western Terrestrial Garter Snake is a series of alternating 
black spots which run the length of the body between, and somewhat on, the yellow stripes. The 
background color between the stripes tends to be more gray compared to the dark brown found in 
the Common Garter Snake. The ventral surface has a series of dark black/brown blotches which 
may cover most of the surface. The dorsal scales are keeled and there are normally 8 upper labial 
scales. 
Young: The coloration of young snakes is similar to that of the adults; young are live-bom. 

Similar species: See Common and Plains Garter Snakes. 

Habitat and Habits: The habitat and habits of the Western Terrestrial Garter Snake are similar to the 
Common Garter Snake, i.e., they are found in most habitats but are particularly common around 
wetlands. On the L&CNF the species was found between 5000 ft and 6000 ft but probably 
occurs much lower and higher Females give birth to 4- 1 9 young during the summer (Stebbins 
1985). 

Surveying: Timed sight surveys may be conducted around wetlands and riparian feeding areas or at 
denning areas where higher concentrations of garter snakes occur; clear mornings are the best 
survey times. Much distributional information may come from recording incidental sightings. More 
intensive research may be done using funnel traps in combination with drift fences. More intensive 
research and survey projects may use mark-recapture or radiotelemetry techniques. 

Status: Westem Terrestrial Garter Snakes were found in the RMRD, Little Belt Mountains, and Big 
Snowy Mountains of the L&CNF. There is also a historic record of them from the Highwood 
Mountains in additional to historic records from all ranges where they were found in 1994. Given 
the small number of records from throughout the area, all records should be documented until the 
distribution is better understood; of particular interest would be documentation of denning sites. 
Sightings oi elegans from the Castle, Crazy, and Little Snowy Mountains should be documented to 
confirm their presence in those ranges. 

Montana Natural Heritage Program Rank: G5 S5. 



Occurrences of Thamnophis sirtalis on or near the Lewis & Clerk National Forest 



^' Pro- 1993 datB 




Specie? iDSBtiors from \h9 Monlsne NetuFsl Herrtsge ProBrem, Zf2@fB^ 



«iHMlfl|MAC^S,C>Iip 



Common Garter Snake (Thamnophis sir talis) 

Description: The Common Garter Snake consists of two color phases in western Montana, both 
ranging from 18-52" inlength. Both phases have three yellow longitudinal stripes: one located 
dorsally and one on each side. Between the yellow stripes is a black stripe broken with red spots in 
one color phase but lacking red in the other Ventral coloration varies from yellow to bluish, and 
some individuals of the red-sided color phase have small black spots on the edge of the ventral 
scales. The dorsal scales are keeled, and normally there are 7 upper labial scales. 
Young: The coloration of young snakes is similar to that of the adults; young are live-bom. 

Similar species: The Western Terrestrial Garter Snake has black spots overlapping the dorsal yellow 
stripe; the background color between stripes tends to be brownish. The Plains Garter Snake has the 
side yellow stripe on the 3rd and 4th scale rows above the belly scales and the dorsal stripe is often 
orange or red. 

Habitat and Habits: Garter snakes are found in all forest habitats but are more common at lower 

elevations around marsh-bog-pond situations, where they prey on young fish, frogs, toads, mice and 
invertebrates. They are sometimes confused with water snakes because of their frequent aquatic 
exploits, but there are no true water snakes in Montana. Typical of most garter snakes, they emit a 
noxious secretion when handled and can be aggressive when disturbed. The Common Garter was 
found between 4 1 00 ft to 5 540 ft on or near the L&CNF, but they are likely to occur at lower and 
much higher elevations. Garter snakes eat a variety of vertebrates and invertebrates, with the 
Common Garter Snake concentrating more on amphibians than the Western Terrestrial Garter 
Snake. The Common Garter Snake is a live-bearer giving birth to 12- 1 8 young during the summer 
in Colorado (Hammerson 1982a). 

Surveying: Timed-sight surveys may be conducted around wetlands and riparian feeding areas or at 
denning areas where higher concentrations of garter snakes occur; clear mornings are the best 
survey times. Much distributional information may come from recording incidental sightings. More 
intensive research may be done using funnel traps in combination with drift fences. More intensive 
research and survey projects may use mark-recapture or radiotelemetry techniques. 

Status: Common Garter Snakes were found in the RMRD and Little Belt Mountains of the L&CNF. 
There is also a historic record of them from the Highwood Mountains. Given the small number of 
records from throughout the area, all records should be documented until the distribution is better 
understood; of particular interest would be documentation of denning sites. Sightings ofsirtalis 
from the Castle, Crazy, and Snowy Mountains should be documented to confirm their presence in 
those ranges. Only the red-sided color phase was observed in the present survey, however the 
color phase lacking red spots should be watched for. The relative abundance of this species in this 
area compared to the Western Terrestrial Garter Snake i s not yet clear; in northwestern Montana 
the Common Garter Snake is currently much less abundant. 

MontanaNatural Heritage Program Rank: G5 S5. 



20 



Occurrences of Ambvstoma tigrinum on or near the Lewis & Clerk Natforval Forest 

'W 'n:^ ^ 




• 1933^94 data 

■^ Prfl-1993 data 

V Museum ^p$cirrar9£ 



Species Eocations from ttia Mootena Natural Merrta^e ProgrsfTt, 2^28/95 



titowhi Bpiyie 3 , em p 



Species Potentially Present on the Lewis and Clark National Forest 

Tiger Salamander (Ambystoma tigrinum) 

Description: Adults have a smooth moist skin without scales and the color pattern is highly variable; 
usually the background color is dark, with lighter blotches of yellow, tan or green. The adult is large 
and heavy-bodied with a snout-vent length of 3 -6" . 

Eggs and Larvae: Egg masses are typically laid in small clusters of 5-120, but may be laid singly 
(Nussbaume^a/. 1983, Leonard e^ a/. 1993). They are usually attached to vegetation and placed 
2"- 1 0" below the surface of the water (Hammerson 1 982a). Larval Tiger Salamanders are typically 
pale green or brown-colored, though some are nearly white in bentonite clay ponds. They are 
found in lakes and ponds, have external gills, and are relatively large (0.75-4" snout-vent) and 
heavy-bodied. 

Similar species: Adult Tiger Salamanders have two prominent tubercles on the bottom of each hind foot 
which Idaho Giant Salamanders lack. Idaho Giant Salamanders also have a more marbled color 
pattern and a very large head. 

Habitat and Habits: Tiger Salamanders in Montana are primarily associated with prairie or agricultural 
habitats in eastern Montana. They breed in ponds or lakes, usually those without fish present. In 
arid areas they may also be found in springs, intermittent streams, and stock ponds. In Blue Lake, 
Madison County, Montana, eggs are laid from early June to mid- August, hatched in about 2 weeks, 
and metamorphosed after more than a year (Micken 1 968, 1971). In Colorado and Wyoming egg 
laying takes place from mid-March to mid- August (Hammerson 1 982a, Baxter and Stone 1985). 
Eggs hatch in 2-5 weeks in Colorado and metamorphosis occurs after 2-24 months (Hammerson 
1 982a). Following breeding, adults may remain in the pond or may move to upland areas and live 
in burrows of their own or in those of other animals. In some locations, such as Blue Lake in 
Madison County, larval salamanders never transform, but rather become sexually mature and breed 
while still retaining external gills. This process is referred to as neoteny and these salamanders are 
called iaxolotlsi or iwater dogs.i 

Surveying: Larvae and eggs may be seen in ponds during the day and may be sampled with a dipnet. 
In areas where larvae transform, migrations of hundreds or thousands of newly transformed adults 
are occasionally seen in mid to late summer or in early fall. During the breeding season, adults are 
often seen in the water or moving to or away from it. Pitfall and minnow traps may be used at this 
time to capture adults. The rest of the summer adults are difficult to find; the best techniques may be 
pitfall traps or driving roads on warm rainy nights. 

Status: The most common salamander in eastern Montana. They have not yet been found on the 
L&CNF, but there are records of larvae from ponds just south of the Castle Mountains (north of 
Lennep) and from Pine Butte Swamp Preserve, just east of the RMRD. They should be surveyed 
for in low elevation ponds and lakes, particularly those without fish and within grassland habitats. 
Any located on the L&CNF should be documented. 

Montana Natural Heritage Program rank: G5 S5 . 



22 



Occurrences of Bufo cognatus on or near the Lewis & Clark National Forest 




• 19S3 84 date 

^ Pra-1993 data ^^ 

"V" Museum spicimens 



Spflcifts kttitions from this Montana Natural Haritaga PfAgrdm. lf2&fSS 



■ri isJ mapd^ c 9 .m^ 



Great Plains Toad {Biifo cognatiis) 

Description: Adults have dry skin with small warts. The coloration is dominated by a number of large, 
dark, somewhat symmetrical spots surrounded by light edges on the back. The dorsal background 
color is gray, light brown or olive green. The Great Plains Toad has converging V-shaped cranial 
crests between the eyes and post-orbital crests connecting to them at a right angle behind the eyes; 
the post-orbital crests typically touch the parotoid glands. The pupils are horizontal. The adult has 
two black tubercles on the hind feet and a body length of 2-3 . 5 " . 
Eggs and Tadpoles: Similar to the Western Toad. 

Similar species: Other Montana toads lack the somewhat symmetrical spotted pattern on the back. 
NOTE: It is very difficult to distinguish among the four Montana toad species in recently 
transformed toadlets. 

Habitat and Habits: Adults may favor higher elevation grasslands than Woodhousels Toad which favors 
floodplains (Bragg 1940, Timkin and Dunlap 1965, Black 1970). They have also been found in 
agricultural areas and open Ponderosa pine savannahs in southeastern Montana (Black 1 970). 
They are most active at night, spend much of the year underground, and emerge in response to 
warm rains (Hammerson 1 982a). They normally breed in temporary ponds resulting from heavy 
rains or irrigation runoff or reservoirs with much fluctuation (Bragg 1 940, Hammerson 1 982a). In 
Montana they apparently breed from May to July (Black 1970). Females lay strings of eggs which 
hatch after 2-3 days (Hammerson 1982a). Youngtypically metamorphose after about 1.5 months, 
although metamorphosis has been reported in as little as 17 days (Hahn 1968, Hammerson 1982a). 

Surveying: Adults may be found by listening for their loud calls on warm (>60° F) nights following 
heavy rains (Hammerson 1982a). iRoad huntingi on warm nights may also be effective. Eggs and 
tadpoles are seen in ponds during the day and can be sampled with a dipnet; however, identification 
of toad eggs and tadpoles is difficult or impossible in the field. 

Status: Notyet found on the L&CNF, but known from the prairie region about 70 miles north of the 
Highwood Mountains. Occurs in localized areas in eastern Montana, with large gaps in its known 
range. Geographic and habitat relationships with other toads in Montana are not well known. The 
Great Plains Toad should be watched for at low elevations in prairie or shrub-steppe habitat on the 
L&CNF, especially on the Jefferson Division. Any located on the L&CNF should be well 
documented with a description indicating how the species was differentiated. 

Montana Natural Heritage Program rank: G5 S4. 



24 



Occurrences of Byfo woodhousii on or near the Lewis & Clark National Forest 




• 13^5-94 data 

^ Prih1993 daid 

"V Mufi9jm specimens 



Sjjscies locstdDna fram tlia Montana Natufal MflritigB P^grern, 2/2S/9E 



«lBD^D0fc'le1 1 



Woodhouseis Toad {Bufowoodhousii) 

Description: Adults have dry skin with small warts, and are gray, brown, or olive-green with paler 
mottling or spots. Aprominent white or yellowish line runs down the center of the back; very young 
transformed toads typically lack the dorsal line, and the warts are often red-brown in color 
Woodhouseis Toad has parallel cranial crests between the eyes and post-orbital crests connecting 
to them at a right angle behind the eyes; the post-orbital crests typically touch the parotoid glands. 
If a lump-like boss is present on the snout, it does not extend back between the eyes. The pupils 
are horizontal. The adult has two black tubercles on the hind feet and a body length of 2.5-4" . 
Eggs and Tadpoles: Similar to those of the Western Toad. 

Similar species: Western Toad lacks cranial crests. Great Plains Toad has large, white-bordered, dark, 
dorsal blotches. The Canadian Toad has a lump between the eyes; frequently the parotoid gland is 
separated from the post-orbital crest which may be broken or absent. NOTE: It is very difficult to 
distinguish among the four Montana toad species in recently transformed toadlets. 

Habitat and Habits: Adults are partially terrestrial but often found near water They are usually found in 
irrigated agricultural areas and flood plains, rather than the more upland areas used by Great Plains 
Toads (Bragg 1940, Timkin andDunlap 1965, Black 1970). They are most active at night, 
although they may at times ben found feeding during the day (Hammerson 1 982a). They typically 
breed in permanent lakes, ponds, reservoirs, and slow streams, with a preference for shallow areas 
with mud bottoms (Black 1970, Hammerson 1982a, Baxter and Stone 1985). Breeding and egg 
laying is spread out over the spring and early summer, with known dates from Montana ranging from 
4 May to 1 July (Black 1970). 

Surveying: Adults may easily be found by using their loud calls for identification on warm (>54° F) 

nights; calling peaks during the first few hours after sunset (Hammerson 1 982a). iRoad huntingi on 
warm nights may also be effective. Eggs and tadpoles are seen in ponds during the day and can be 
sampled with a dipnet; however, identification of toad eggs and tadpoles is difficult or impossible in 
the field. 

Status: Not known from theL&CNF; the nearest known record, collected in 1918, is from about 30 
miles southeast of the Little Snowy Mountains, 1 mile west of Kline. Woodhouseis Toad is 
relatively common in southeastern Montana, however, its status elsewhere in the state is unclear 
Geographic and habitat relationships with other toads in Montana are not well known. It should be 
watched for at low elevations in prairie or shrub-steppe habitat on the L&CNF, especially in the 
Jefferson Division; it could occur along the Smith River. Any located on the L&CNF should be well 
documented with a description indicating how the species was differentiated. 

Montana Natural Heritage Program rank: G5 S4. 



26 



Occurrences of Scaphiopus bombifrons on or near the Lewis & Clark National Forest 




• 1993 94 data 
■ ' Fhre-1993 date 



Spaelas liicptlcwia frafn Ihe MpnTsne Natural Hefiteigie Ptosrafn, 2/23/96 



nlak/maiM^lel Bxnp 



Plains Spadefoot {Scaphiopus [=Spea] bombifrons) 

Description: Adults are colored gray or brown with darker mottling on the back and a white belly. 
Some individuals have indistinct longitudinal streaking. The pupils of the Plains Spadefoot are 
vertically elliptical and there is a high, hard lump between the eyes. Its skin is less warty than true 
toads. The adult has a single tubercle on the hind feet and has a body length of less than 2.5". 
Eggs and Tadpoles: Oval egg masses of 1 0-250 eggs are attached to underwater plants or 
debris. Tadpoles are mottled sooty and olive-yellow above and paler below with gold metallic 
flecking over all; iris is gold. 

Similar species: Other Montana frogs and toads have round or horizontally elliptical pupils. 

Habitat and Habits: Adults are found in grassland and sagebrush areas, particularly in areas with sandy 
or loose soil (Wheeler and Wheeler 1966, Hammerson 1982a, Baxter and Stone 1985). Except 
during breeding, they are seldom found in the water They are primarily nocturnal and emerge 
from their burrows only following heavy rains. They breed in shallow temporary pools usually 
following heavy spring or summer rains (Hammerson 1 982a). Males call loudly, with groups being 
heard for up to a mile. Eggs hatch after 2-3 days and tadpoles transform in 6- 1 weeks (Wheeler 
and Wheeler 1966, Hammerson 1982a). 

Surveying: Adults may be easily found by using their calls for identification when breeding at night or 
by iroad huntingi on warm, rainy nights. Calling normally takes place only when the temperature 
is >50° F (Hammerson 1982). Tadpoles are seen in ponds during the day and can be sampled 
with a dipnet. Surveying is complicated by the long time periods which this species spends 
underground, especially during droughts. 

Status: The Plains Spadefoot is not known from L&CNF lands; the nearest records are from 
Cascade County about 20-30 miles west of the Highwood Mountains, WNW of the Little Belt 
Mountains, and in Great Falls. Locally common in eastern Montana; there are large gaps in the 
known range. It should be watched for at low elevations in prairie or shrub-steppe habitat on the 
L&CNF, especially the Jefferson Division. Any located on the L&CNF should be well- 
documented. 

Montana Natural Heritage Program rank: G5 S4? 



28 



Occurrences of Chrysemys picta on or near tha Lewis 8l Clark National Forest 




* 1993^94 data 
C> Pra-1993 data 
V* Museum specimens 





T 




Species k»c«tl9ris from the Mcnitene Natural MsritPfe Prpsrpnip 2f2BfB^B 



^i1 B ■ ^mBpa^'tc 2 3 . cmp 



Painted Turtle (Chrysemys picta) 

Description: Adult Painted Turtles have a relatively flat dorsal shell, or carapace, the length of which 
may reach 9" in females and 7" in males. The background color of the shell may be dark brown, 
olive, or black. A series of short, irregular yellow lines are often scattered across the shell, and a 
red and black border forms the outer edge. The ventral shell, or plastron, is red with a centrally- 
located yellow and black blotch with edges flaring out along the border of the scutes. The edge of 
the plastron also has a series of black and yellow blotches. The head, neck, and legs are marked 
with yellow lines and a red spot appears behind the eye. Very dark colored individuals are 
occasionally found. Males are distinguished by longer front claws and longer tails with the anus 
posterior to the margin of the carapace (Ernst et al. 1 994). 

Eggs and Young: The elliptical, white, soft-shelled eggs are about 28-3 5 mm in length and 1 6-23 
mm in width (Ernst et al. 1 994). They typically number 6-23 per clutch. Coloration of young 
Painted Turtles is more vibrant and the shell is not quite as flattened as adults. 

Similar Species: None. 

Habitat and Habits: Painted Turtles are active during the day and are rarely seen far from ponds, lakes, 
or the slow-moving water of streams. Adults are primarily herbivorous, feeding on a variety of 
aquatic plants, but will also scavenge on animal remains. Eggs are usually laid within 1 0-20 feet of 
the waterls edge, although some individuals will travel up to 600 m seeking a suitable site. During 
egg-laying, the female excavates a hole with her hind feet and deposits the eggs, which are then 
covered by several inches of dirt. Predation on turtle eggs by raccoons, skunks, etc. is common, 
and shell fragments are evidence of such activity. Female Painted Turtles may lay more than one 
clutch of eggs each summer. Young borne of late egg depositions overwinter in the nest and do not 
emerge until the following spring (Ernst etal 1 994). Once females lay their eggs, they return to the 
pond, where they can often be seen basking on logs or rocks along with juveniles and males. 
Painted Turtles are sexually mature at 3-5 years of age and may live to be 30 years or older (Ernst 
etal. 1994). 

Surveying: Although various turtle traps can be used for surveys, visual identification is suitable for 
presence/absence studies since the three turtle species in Montana are easily distinguished. Basking 
peaks at different times during the day, depending on season and location; in the northern states and 
Canada it generally peaks in the morning. Surveys should be done on sunny days with a pair of 
binoculars. During cold or cloudy weather, turtles tend to remain underwater for long periods and 
can be missed on a walk-through survey. 

Status: Painted Turtles are locally quite common in Montana at lower elevations. They were not found 
on the L&CNF, but were seen only a few miles east of the RMRD. There has been some concern 
about Painted Turtle populations nationally, and whether declines have occurred in Montana is 
unknown. It should be watched for particularly at lower elevations in ponds in open forest, prairie 
or shrub-steppe habitat on the L&CNF. Any animals located should be documented. 

Montana Natural Heritage Program Rank: G5 S5 . 



Occurrenoea of Trjonyx spintf^rus on or near the Lewis & Clark National Forest 




■ 1933-54 data 
Pre- 1993 dBfU 

>i^ Mijuuin spaclmflns 



Spfi<:J^fi l4?Btl0nS twrr Itia M^intar? Natural Herrt^te Prflgrpjiip 2/2S/9S 



«rlBuAii«fiB^?9',i;niil 



Spiny Softshell (Trionyx spiniferiis) {=Apalone spinifera) 

Description: Spiny Soft-shells have flexible, leathery shells. The carapace is olive-gray, marked with 
dark spots. The plastron is white or light cream-colored. Female carapace length is up to 1 8 
inches or more, whereas males are typically 6-8 inches. The nostrils are terminal, allowing this turtle 
to remain entirely beneath the surface and take air through its isnorkel.T 

Eggs and Young: The nest is a flask-shaped excavation containing 4-39 (typically 12-18) hard- 
shelled, spherical, white eggs. The individual eggs range in size from 24-32 mm in diameter and 
average about 28 mm. Hatchlings resemble adults and are 30-40 mm in shell length (Ernst etal. 
1994). 

Similar Species: None. 

Habits and Habitat: Spiny Softshells are active during the day. This highly aquatic turtle is found in 
rivers or their connecting backwaters with muddy or sandy bottoms. Unlike other Montana turtles, 
they do not move overland from one water body to another. Mud and sand banks and bars are 
used for both basking and nesting. Hibernation takes place beneath the water, usually beneath 5-10 
cm of bottom substrate (Ernst et al. 1 994). The retracted head and neck combines with the profile 
of the shell to produce a wedge shape, which allows this turtle to escape by literally diving into the 
bottom mud. If necessary, additional strokes of the legs will completely bury it in the substrate, 
hidden from view. Food items include fish, crayfish, frogs, toads, aquatic insects, and carrion. 
Spiny Soft-shells have a surprisingly long, agile neck and can inflict a painful bite. They can be 
safely handled by grasping the shell on each side between the front and rear legs with the head 
pointing away from the captor. 

Surveying: Although various turtle traps can be used for surveys, visual identification is suitable since the 
three turtle species in Montana are easily distinguished . Apair of binoculars is helpful and surveys 
should be done on warm sunny days; basking seldom takes place before 10:00 a.m. (Ernst etal. 
1 994). During cold or cloudy weather, turtles tend to remain underwater for long periods and can 
be missed on a walk-through survey. Care should be taken to watch for the snorkel -like nostrils 
projecting just above the surface of the water. 

Status and Distribution: Found mainly in the Yellowstone and Missouri Rivers and their maj or 

tributaries. These populations may be separated from each other and are believed to be disjunct 
form the population in South Dakota (Ernst et al. 1 994); they have not been reported from North 
Dakota (Wheeler and Wheeler 1 966). The Missouri River population is known from the tail of Fort 
Peck Reservoir upstream to the first dam above, and from most of the Mussellshell River; their 
presence on other tributaries is presently unknown. They have been reported from the Mussellshell 
River to just southeast of the Little Belt Mountains and should be watched for in the Mussellshell, 
Judith, and Smith Rivers on the L&CNF. Any located on the L&CNF should be documented and 
reported. Considered a Species of Special Concern in Montana. 

Montana Natural Heritage Program Rank: G5 S3. Species of Special Concern. 



32 



Occ:urfences of Phrynosoma douglasi on or near the Lewis Et Clark National Forest 



^ 



«- 



\ 



• 1$a3'S>4 data 
'^ Pre- 1993 data 



1 1 






L 






<^ 





iffi^iij 




Specffis loc4tfon» fivm the Montana Natural Hsrltage Program, 2/26/9^5 



ilt^r.lmapiflt27jc!m 



Short-horned Lizard (Phrynosoma douglasi) 

Description: The Short-homed lizard has a broad, somewhat flatted body and relatively short limbs and 
tail. It is generally tan to gray with dark and light spots and blotches; the belly is white. There is a 
distinctive line of pointed scales along each side and the head has short, blunt ihornsT pointing 
backward. Adult lizards range from 1 . 7 - 5 . 5 " in length. 
Young: Young are live-bom and resemble small adults. 

Similar species: None. 

Habitat and Habits: The Short-horned lizard is found in a variety of habitats, including dry open forests, 
grasslands, and sagebrush; the soil is usually loose or sandy. In firmer soil situations it may use the 
burrows of other animals. It is active during the day, typically with the peak of activity in mid-late 
moming. A Short-horned Lizard may squirt blood from its eyes when disturbed. Little is known 
about reproduction in this part of the range; young are bom in late summer Ants are the primary 
food of the species. 

Surveying: They may be surveyed for by slowly walking through appropriate habitat and carefully 
watching for them; look carefully near ant mounds; this technique has low success with Short- 
homed Lizards however As with many lizards and snakes, they are easily missed. Carefully 
documented incidental observations may provide the best clues to their distribution. They may be 
also taken in pitfall or funnel traps in combination with drift fences. 

Status: Widely distributed (but apparently localized) in eastem Montana. This species may be 

vulnerable to collecting for the pet trade and agricultural conversion of native habitats. The Short- 
homed Lizard subspecies found in Montana {P. d. brevirostrd) is currently a U. S. Fish and Wildlife 
Service Category 2 Candidate species (U. S. Fish and Wildlife Service 1 994). In the vicinity of the 
L&CNF it has been found at breaks west of Ulm, Egg Mountain on Nature Conservancy land, 
north of the Highwood Mountains, and near Harlowton southeast of the Little Belt Mountains. It 
should be watched for in open pine, prairie, or shrub-steppe habitat with loose or sandy soils at 
lower elevations on the L&CNF; any sightings should be documented. 

MontanaNatural Heritage Program Rank: G5 S4. 



34 



Occurrences of Charina bottae on or near tha Law is & Clark National Forest 





* 



* 



1993-94 data 
Pre- 1993 dPtfl 



1 



J^ 



r" 




Specie^ iDcatifjTis ffdm tha ManTana Nature] N^ritaga PrDgrsm, Zf^^fBB 



ii^^«fiiAt34ciV 



Rubber Boa (Charina bottae) 

Description: The Rubber Boa looks and feels like mbber, hence its name. It is a small snake (14-33" 
length), stout, and uniformly-colored either brown or green on the dorsal side. The ventral surface 
is cream to tan in color. The scales are small and smooth, except for those on the head which are 
enlarged. The tail is short and blunt and the eyes are very small. It is a very slow moving snake 
which can easily be caught if detected. 

Young: Rubber Boas are born alive and young are more tan (or even pinkish) than the adults on 
both the dorsal and ventral surfaces. 

Similar species: The Racer is much quicker and more active, has larger eyes, and a thin, tapered (not 
blunt) tail. 

Habitat and Habits: The Rubber Boa is a secretive, slow-moving, docile snake, usually found under 
logs and rocks in either moist or dry forest habitats, but rarely in marsh or bog situations. Denning 
locations are typically in areas with fractured rock; recent data indicates it only moves short 
distances from its den (Peterson pers. comm). Occasionally this snake is seen sunning itself on a 
road, trail, or open area, but it is primarily nocturnal. Feeding is primarily on small mice, but also on 
shrews, salamanders, snakes and lizards. Two to eight young are bom alive in late summer or early 
fall. 

Surveying: There are no practical methods for surveying other than systematic searches of a given area 
rolling over rocks, logs, etc. Driving roads at night, particularly after a rain when the temperature is 
> 1 0° C, may be more effective, especially on roads which follow a stream. Previous sightings are 
of value in locating general areas of activity and denning sites. Funnel traps may be effective. 

Status: Sightings of Rubber Boas are infrequent, but they are widely distributed and probably common 
throughout western Montana. They were not found during this survey, nor are there historic records 
of their presence in the L&CNF. However, they probably do occur throughout the RMRD of the 
L&CNF at low to mid-elevations. They should be watched for on the L&CNF; any sightings 
should be documented. Of particular interest would be any reports of this species from the 
Jefferson Division or documentation of any denning sites located. 

Montana Natural Heritage Program Rank: G5 S4. 



36 



Occurrences of Coluber constrictor on or near the Lewis & Clark National Forest 




Species locaticms from the Mortana Natural Herrtage Prograni, 2/28/95 



■tinnAn^H^St .cmp 



Racer {Coluber constrictor) 

Description: A slender, but moderately long snake, the Racer ranges from 20-65 inches in length. Adult 
coloration is uniform across the dorsal side but it can vary from a greenish-gray to brown or blue. 
The ventral side is whitish to pale yellow, the latter color extending onto the upper lip scales and 
nasal region. The eyes are relatively large. The scales are smooth and the nostril is bordered by 
two scales. 

Young: Snakes (up to about 20") have a much different coloration than the adults consisting of a 
series of dorsal brown blotches edged with black which run the length of the animal; a row of 
blotches is also found on each side of the animal extending onto the ventral side. 

Similar species: Young Gopher Snakes may be distinguished by the keeled rather than smooth scales of 
the young Racer Young Western Hognose Snakes have an upturned nose. Smooth Green Snakes 
are smaller and colored bright grass-green and whitish below; their nostrils are centered in single 
scales. Also see Rubber Boa. 

Habitat and Habits: The Racer is associated with more open habitats either in shortgrass, shrub-steppe, 
or forested areas (Hammerson 1982a, Baxter and Stone 1985). It is often found near water and 
rocks. The Racer is an extremely fast and agile snake. A clutch of perhaps 3-7 eggs is laid in the 
summer (Stebbins 1 985). It preys on insects and small vertebrates such as mice and frogs. 

Surveying: They may be surveyed for by slowly walking through appropriate habitat on warm, sunny 
days and carefully watching for them; this technique is moderately effective for the Racer However, 
as with many lizards and snakes, they may easily be missed. Carefully documented incidental 
observations may provide the best clues to their distribution. They may be also taken in funnel traps 
with drift fences. Mark-recapture methods offer the best opportunity for determining population 
status. 

Status: The Racer was not seen in this survey nor are there historic records from the L&CNF. 

However they are known just to the south of the RMRD and from about 25 miles west of the Little 
Belt Mountains. They probably do occur on the L&CNF at low to mid-elevations; any sightings 
should be documented. Of particular interest would be documentation of any denning sites located. 

Montana Natural Heritage Program Rank: G5 S5. 



38 



1 




• 1993-94 dMx 

V" MuMurrv specimieris 



1 




X^ 




Occurrences of Heterodon nasicuB on or near the Lewis & Clark National Forest 



k 





Speelas locatk^nt trtym tha Montana Natural Hsrltags Program, 2i2B}QB 



EtJ uMivaJk 3 2 jDirv 



Western Hognose Snake (Heterodon nasicus) 

Description: The Western Hognose Snake is a mid-sized, heavy-bodied snake reaching 32" . The dorsal 
ground color is yellowish- to grayish-brown, with 3 rows of darker brown to black blotches run 
down the back. The belly is dark gray to black, sometimes checkered. Its nose has a keel on the 
top and is upturned. 

Eggs andYoimg: Eggs are white and elliptical, with thin, papery shells; length averages 32.5 mm 
(26-38 mm) and width 18 mm (14-23 mm). Young are 139-197 mm at hatching and are similar in 
color and pattern to adults (Piatt 1969). 

Similar Species: No other Montana snake has a keeled nose. Coloration is similar to both the Gopher 
Snake, Western Rattlesnake and juvenile Racer 

Habits and Habitat: The Western Hognose Snake is found on the plains of eastern Montana. It seems 
to prefer arid areas, farmlands and floodplains, and particularly areas of gravelly or sandy, loose 
soil. The keeled, or shovel -like, nose is thought to help it to dig down to its food, which it finds by 
smell. Apparently toads are its preferred food, though frogs, insects, and other small animals are 
also eaten (Piatt 1 969). It is active primarily during the daylight hours. Little is known of 
reproduction in Montana. In Kansas, Western Hognose Snakes typically lay clutches of 7- 1 5 eggs 
(Piatt 1 969). It is likely that a female will only breed every other year in Montana. 

The Hognose is famous for its behavior in the face of a threat. At first it will puff up its neck, as 
does a cobra, and hiss and strike at its enemy. However, this is all a bluff and very rarely will it 
actually bite. If this threatening strategy does not work, it will pretend to die. It appears to go into 
convulsions, writhing on the ground, sticking its mouth in the dirt, and eventually rolling on its back 
and going into a trance that makes it appear to be dead. If turned right-side-up, it will roll back 
over and continue its deception. If left alone for a few minutes, it will right itself and continue on its 
way. The initial aggressive display and basic rattlesnake-like coloration cause many to be killed 
needlessly by people who mistakenly believe it to be venomous. 

Surveying: They may be surveyed for by slowly walking through appropriate habitat and carefully 
watching for them; timing surveys for warm sunny days enhances sampling success. However, as 
with many lizards and snakes, they may easily be missed. Carefully documented incidental 
observations may provide the best clues to their distribution. They may be also taken in pitfall or 
funnel traps with drift fences. Mark-recapture methods offer the best opportunity for determining 
population status. 

Status: They were not found during this survey, nor are there historic records of their presence in the 
L&CNF. However, they may occur on the L&CNF at low to mid-elevations, particularly in sandy 
or loose soil areas. The nearest location to the L&CNF is from central Teton County about 30 
miles east of the RMRD. They should be watched for on the L&CNF; any sightings should be 
documented. We have relatively few reports of the Western Hognose Snake from Montana. It is 
collected for the pet trade, and populations may be vulnerable to commercial collectors. 
Additionally, since toads are its preferred food, any decline in toad populations would be expected 
to negatively impact Western Hognose Snakes. Of particular interest would be documentation of 
any denning sites located. 

MontanaNatural Heritage Program Rank: G5 S3? Species of Special Concern. 



40 



Occurrences of Pituophia catenifer on or near the Lewii^ & Clark National Forest 

■■.'"vx; 




• 1993-94 data 
^ Pr*-1S93 data 
V^ MuseLiin spa&iiria.na 



SpQCise Eflcationg, Trqm ttie Momtera Natural Herrtage ProBrarrir 2j'2S/95 



B tlB«ATiviafc3&. cmp 



Gopher Snake (Pituophis catenifer [=melanoleiicus]) 

Description: Montanals largest snake, the adult Gopher Snake (also called Bullsnake or Pine Snake) 
can reach a total length of 7 feet, but most specimens seen in western Montana range between 3-5 
feet. It is readily recognized by a series of large black to brown blotches which run down the back, 
and another series along the sides. The blotches, which are set on a yellow background, become 
more widely spaced and darker towards the tail . The dorsal scales are keeled. There is usually a 
black band on the head located in front of and extending below the eyes. The ventral coloration is 
yellow to white, often spotted with black, and the anal plate is undivided. 

Eggs and Young: Gopher Snakes lay between 2-24 eggs during the summer months (Hammerson 
1 982a), and the young resemble the adults in coloration. 

Similar species: Young Racers have a black border on dark blotches and the scales are not keeled. 
Young Western Hognose Snakes have an upturned nose. Western Rattlesnakes have a rattle on 
their tail and triangular shaped heads. 

Habitat and Habits: Gopher Snakes are associated with dry, arid habitats including grassland, shrub- 
steppe, and open pine forest. They feed on rodents, rabbits and ground dwelling birds, and to a 
lesser extent on frogs, toads, etc., found around stock ponds and other wetlands. They have a habit 
of hissing and vibrating the tail when alarmed, often sounding like rattlesnakes. They occasionally 
climb trees, hence the common name iPine Snake.T 

Surveying: Walk-through surveys, done on a regular basis in warm, sunny weather probably give the 
best results without resorting to trapping techniques. They are most easily found near dens in the 
spring and fall. Funnel trapping is effective and they may occasionally be found by night driving 
during the mid-summer Data can be enhanced by mark-recapture techniques. 

Status: The Gopher Snake was not seen in this survey nor are there historic records from the L&CNF. 
However they are known from lower elevation areas on the plains adj acent to the forest. They 
probably do occur on the L&CNF at low to mid-elevations; any sightings should be documented. 
Of particular interest would be documentation of any denning sites located. 

Montana Natural Heritage Program Rank: G5 S5 . 



42 



OcGurrances of Thamnophis radix on or near the Lewis Si Clark National Forest 



• 1993-94 deu 
O Pr«-1S93 data 

'V Museum spscimens 




Speclas locations Itinti the Montana Natural Heritage Program, 2/2Q/9& 



«[teiuniapiAe3T.cn)p 



Plains Garter Snake {Thamnophis radix) 

Description: The Plains Garter Snake ranges in size from 16-42" in length and has a dorsal background 
color of olive, brown, or black. It has a prominent orange or yellow dorsal stripe and a greenish- 
yellow stripe on each side located on the 3rd and 4th scale rows above the belly scales. It typically 
has black vertical bars on the upper lips. 
Young: Young resemble adults. 

Similar species: The other garter snakes found in Montana have the lateral yellow lines on the 2nd and 
3rd scale rows above the belly scales. 

Habitat and Habits: The habitat and habits of the Western Terrestrial Garter Snake are similar to those 
of the Common Garter Snake. 

Surveying: Timed sight surveys may be conducted around wetlands and riparian feeding areas or at 
denning areas where higher concentrations of garter snakes occur; clear mornings may be the best 
survey times. Much distributional information may come from recording incidental sightings. More 
intensive research may be done using funnel traps in combination with drift fences. More intensive 
research and survey projects may use mark-recapture or radiotelemetry techniques. 

Status: Found over much of eastern Montana. Its status is unclear due to confusion in the identification 
of the 3 garter snakes which occur there. In the vicinity of the L&CNF it has been found near 
Great Falls and north of the Highwood Mountains. It should be watched for in prairie or shrub- 
steppe areas at lower elevation of the L&CNF and any sightings should be well documented with a 
description written at the time of observation including how radix was distinguished from the other 
garter snakes. Of particular interest would be documentation of any denning sites located. 

MontanaNatural Heritage Program Rank: G5 S4. 



44 



Occurrences of Crotalus viridis on or near the Lewis & Clark National Forest 



• 1993-94 data 

V~ Museum ^ecimens 




S|j6[^i«5 Igcatiana from Ihe Mttntana Katural Heritage Pirogrem, 2i23i§S 



■tlH'nMpkAaSS.cnnp 



Western Rattlesnake {Crotalus viridis) 

Description: Rattlesnakes have a heat-sensing pit located between the nostril and the eye. The fangs 
are hollow and hinged, allowing them to be folded back against the roof of the mouth. The head is 
triangular in shape and blunt-nosed. The eyes are slightly elevated. There are several white lines 
which run along the side of the head. Adult Western Rattlesnakes have a narrow neck but a stout 
body with total length ranging from 1 5-60 inches. The dorsal background color varies from pale 
green to brown with a series of brown or black blotches edged with a dark and then light line 
extending the length of the body. The blotches often merge into rings on the tail. There are also 
blotches on the sides of the body. The ventral side is pale yellow to white and without blotches. 
The scales are keeled. The tail ends in a rattle which helps to warn potential predators of the 
snakels presence. The young have the same color pattern, but are brighter in color than adults. 

Similar species: No other snake in Montana has rattles, but see Racer, Gopher Snake and Western 
Hognose Snake which may have similar color patterns. 

Habitat and Habits: The Western Rattlesnake is an inhabitant of more open and arid country but it is 
also found in Ponderosa pine stands or mixed grass-coniferous forests. It is more likely to be 
encountered on south-facing slopes and areas of rock outcrops. It is feared and often needlessly 
killed due to its poisonous bite. Rattlesnakes may den in large numbers, moving up to 7 miles out 
from the dens during the summer (Peterson, pers. comm.); den sites are most common in south- 
facing talus slopes. In Wyoming, it is found at elevations of over 8500 feet (Baxter and Stone 
1985). Rattlesnakes prey on a variety of animals including mice, ground squirrels, rabbits, 
amphibians, and other snakes. Females give birth to 4-21 young in Colorado during the summer 
(Hammerson 1982a). 

Surveying: Walk-through surveys on warm sunny days is probably the best method for determining 
presence/absence; easiest to find near den sites in spring and fall. Funnel traps and night driving are 
both effective techniques. Mark-recapture methods can be used to determine more precise 
numbers. 

Status: The Western Rattlesnake was not been found in the L&CNF but is known from lower elevation 
areas to the north, south, and east. It would most likely be encountered at lower elevations in open 
habitats on the Jefferson Division. The habit of denning at traditional sites in large numbers makes 
rattlesnakes vulnerable to commercial collecting or simply killing by fearful people. Observations of 
Western Rattlesnakes should be reported to document the presence of this species on the L&CNF; 
of particular interest would be documentation of any denning sites located. 

MontanaNatural Heritage Program Rank: G5 S4. 



46 



Rocky Mountain Division Information 

Rocky Mountain Division: Two amphibian species were only found on the Rocky Mountain Division 
of the L&CNF : the Long-toed Salamander and Tailed Frog. Both species appeared to be more 
localized than in areas to the west of the Divide. Additional surveys should be undertaken to determine 
how widespread these species are; all incidental observations should be recorded. The Rubber Boa is 
likely to be present due to its proximity to known localities to the south and west; whether it will also be 
found in the Jefferson Division is unknown. 

Other species found during surveys or for which historic locations are known from on the RMRD 
include the: Western Toad, Western Chorus Frog, Northern Leopard Frog, Spotted Frog, Western 
Terrestrial Garter Snake, and Common Garter Snake. All of these species are also present on the 
Jefferson Division. The Western Toad was only found breeding at a single location on the forest; this 
may be a function of the small amount of sampling or the apparent decline in much of western Montana. 
The breeding location in the backwater^eaver pond on the Teton River at 7 Lazy P Ranch should be 
considered for monitoring to see if Western Toads continue to breed there and are successfully reaching 
metamorphosis. Western Chorus Frogs, though common at lower elevations outside the L&CNF, 
apparently are very uncommon and localized within the forest. No Northern Leopard Frogs were seen 
on surveys. However a museum specimen exists from the iSun River, 5500 feeti; this location 
information is poor because at 5500 feet the Sun River is split into the maj or forks. While doing 
Harlequin surveys. Northern Leopard Frogs should be watched for along the Sun River Substantial 
populations of the Spotted Frog were found throughout the RMRD; it was more commonly 
encountered than any other amphibian or reptile, though ironically it is one of the two C-2 Candidate 
species present on the L&CNF. An ideal marsh-pond area below Wood Lake on Wood Creek might 
be considered for long-term monitoring of Spotted Frogs and Long-toed Salamanders; the area to be 
monitored would have to be carefully delimited, however, since the area is large. Concerned citizens 
might be willing to participate in long-term surveys. 

Given the low numbers of locations for any amphibians or reptiles on the RMRD, all sightings of any 
species should be recorded. A possible exception would be for Spotted Frogs; however, breeding 
locations found should be recorded even for this species until more are known and mapped. Of 
particular interest would be records of the following species which have not yet been recorded on the 
District: Tiger Salamander, Great Plains Toad, Plains Spadefoot, Painted Turtle, Short-homed Lizard, 
Racer, Western Hognose Snake, Gopher Snake, Plains Garter Snake, and Western Rattlesnake. All of 
these potentially present species are most likely to be seen at low elevations in open habitat. 

Jefferson Division Information 

Highwood Mountains : This range is the only Jefferson Division area in which Western Chorus Frogs 
are known; they were found by Mike Enk in 1 994 and had been collected in the 1 960s north of the 
1 994 location. The Western Chorus Frog is the only herp recently recorded from the Highwoods. The 



47 



Western Toad, Northern Leopard Frog, Western Terrestrial Garter Snake, and Common Garter Snake 
were all collected along Highwood Creek or Arrow Creek in the 1 960s but were not relocated during 
our surveys. Perhaps the best chance of finding the Great Plains Toad on the L&CNF occurs in the 
Highwoods, given its known site to the north. Given the meager information available from this range 
and at least two species (known to be declining) recorded from historic records but not relocated, all 
sightings of amphibians and reptiles are of interest from this range and should be recorded. Baseline 
distribution information, particularly for species not yet recorded and breeding sites for known species, 
is necessary before monitoring sites are chosen. 

Little Belt Mountains: Spotted Frogs appeared to be common and widespread in this range. 
Western Toads have been collected historically, but were not located during our surveys; any sightings 
of this declining amphibian should be recorded. The Western Terrestrial Garter Snake and Common 
Garter Snake were the only other herps found during our surveys. Perhaps the best chance of finding 
the Spiny Softshell on the L&CNF occurs in the Little Belts in the Smith, Judith, or Mussellshell Rivers. 
Given the meager information available from this range and at least one species known from historic 
records but not relocated, all sightings of amphibians (except perhaps Spotted Frogs) and reptiles are of 
interest and should be recorded. Even for Spotted Frogs any breeding locations found should be 
recorded. Such baseline distribution information is necessary before monitoring sites are chosen. 

Castle Mountains: The Spotted Frog is the only herp known from this range; there is one historic 
specimen record and the species was also found during our 1 994 surveys in beaver ponds in the West 
Fork of Checkerboard Creek. Tiger Salamander larva were found at the base of the range just north of 
Lennep off USFS Lands. Given the meager information available from this range all sightings of 
amphibians and reptiles are of interest and should be recorded. Baseline distribution information, 
particularly for species not yet recorded and breeding sites for known species, is necessary before 
monitoring sites are chosen. 

Crazy Mountains: The Spotted Frog was the only herp found during surveys in 1994 and was 
located in Forest Lake. Western Toads have been collected historically in the Crazy Mountains, but 
were not located during our surveys; any sightings of this declining amphibian should be recorded. 
Given the meager information available from this range all sightings of amphibians and reptiles are of 
interest and should be recorded. Baseline distribution information, particularly for species not yet 
recorded and breeding sites for known species, is necessary before monitoring sites are chosen. 

Big Snowy Mountains: The Western Terrestrial Garter Snake is the only herp known from this range; 
there are three historic specimen records and this species was found during our 1 994 surveys on the 
shore of Crystal Lake. Given the meager information available from this range, all sightings of 
amphibians and reptiles are of interest and should be recorded. Baseline distribution information, 
particularly for species not yet recorded and breeding sites for known species, is necessary before 
monitoring sites are chosen. 



48 



Little Snowy Mountains: No herps are known from this range; there are no historic specimen 
records and we did not survey here in 1994. Given the total lack of information available from this 
range, all sightings of amphibians and reptiles are of interest and should be recorded. This baseline 
distribution information, particularly for species not yet recorded and breeding sites for known species, 
is necessary before monitoring sites are chosen. 



49 



RECOMMENDATIONS 

1) All incidental sightings of amphibians and reptiles from the L&CNF should be recorded and 
forwarded to the Natural Heritage Program. The single exception being that for the Spotted Frog on 
the RMRD and Little Belt Mountains, only breeding locations are necessary to record. Ahalf-day 
training session for biologists (including seasonal employees) and other interested field people in May 
would raise awareness of this data need and provide the training needed for accurate identification of 
animals observed. Certainly all Tailed Frogs (larvae and adults) found during fisheries surveys should be 
recorded; this is the most efficient way to get data on this species. The Tailed Frog is now a USFWS 
C-2 Candidate. 

2) Due to the time constraints and the large area covered in the 1 994 survey, it should not be regarded 
as a definitive index of all the herptiles or their distribution on the L&CNF. The secretive habits of many 
amphibians and reptiles, and our lack of knowledge regarding their reproductive behavior makes it 
difficult to assess their overall status. We recommend that additional surveys be conducted, 
concentrating on: A) potential Western Toad and Northern Leopard Frog breeding sites; B) low- 
elevation, xeric habitats (including wetlands within this matrix) for reptiles and plains-dwelling 
amphibians; C) re-surveying historic amphibian sites; and D) gathering additional distribution information 
from the isolated ranges on the Jefferson Division. 

3) When more breeding locations for amphibians are known, long-term monitoring of typical marsh- 
pond habitats should be set up at several sites in each Division in order to evaluate relative numbers and 
breeding success of the more common species: Long-toed Salamander, Spotted Frog, Western Toad, 
Western Chorus Frog, Western Terrestrial Garter Snake, and Common Garter Snake. Particular 
attention needs to be given to any Western Toad and Northern Leopard Frog breeding sites found. 

4) Life history and ecology of the amphibians in Montana is poorly known for most species. Long- 
term monitoring will give us information on timing of and habitat requirements needed for successful 
breeding. 

5) Sightings on L&CNF lands of the Tiger Salamander, Great Plains Toad, Woodhousels Toad, Plains 
Spadefoot, Painted Turtle, Spiny Softshell, Short-horned Lizard, Rubber Boa, Racer, Western Hognose 
Snake, Gopher Snake, Plains Garter Snake, and Western Rattlesnake would represent first-time 
occurrences and range extensions, thus it is important to document and record such data. Preferably 
either photos should be taken or, if appropriate, a specimen collected; at the very least, a description 
should be written at the time of the observation. 



50 



BIBLIOGRAPHY 

Anderson, M. E. 1977. Aspects of the ecology of the two sympatric species of Thamnophis and heavy 

metal accumulation within the species. M.S. Thesis, University of Montana, Missoula. 147 pp. 
Baxter, G.T. and M.D. Stone. 1985. Amphibians and reptiles of Wyoming. Wyoming Game and Fish 

Department. Cheyenne, Wyoming. 137 pp. 
Belhs,E.D. 1962. Theinfluenceofhumidity on wood frog activity. Amer Midi. Nat. 68(1):130 148. 
Bergeron, D. No date. Terrestrial wildlife survey. Coal Creek Mine Area, Montana, 1977 1978. West. 

Tech. & Eng., Inc., Helena. 
Bernard, S. R. and K. F. Brown. 1977. Distribution of mammals, reptiles, and amphibians by BLM 

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59 



APPENDIX 1. 

DATA SHEETS USED FOR 

AMPHIBIAN AND REPTILE 

SURVEYS AND OBSERVATIONS 



60 



AMPHIBIAN SURVEY DATA SHEET: INSTRUCTIONS This data sheet is designed to fecilitate quick recording of data from field surveys of amphibians and their habitats. 
It appears complex and intimidatipg, but actually can be completed in a short amount of time after a minimum amount of training. Many variables require only the correct choice 
to be circled, and the remaining variables are numerical and easy to determine. The data sheet is divided into four sections, divided by double lines. Each section describes 
a cohesive set of variables. In addition the back of the sheet includes a grid for a rough sketch of the site and space for additional comments. The map is optional, but the futuie 
value of the data is enhanced if it is supplied. 



SECTION 1 - LOCALITY These data are 

essential. Many amphibian surveys have been 

hankered by the inability to relocate exact 

locations in the historical record. Some of this 

information can be completed in the office ajler 

the survey. 

DATE: Use the format DD-MMM-VY (e.g., 
05-APR-92). 

BEGIN TIME: List the time surv^ of habitat 
for amphibians began in 24 hour format. 

END TIME: List the time the surv^ ended in 
24 hour format. (The total time (END 
TIME - BEGIN TIME) should reflect only 
the amount of time spent searching for 
amphibians. Total time plus number of 
observers mey be used to assess relative 
abundance) 

OBSERVERS: List names or initials of all 
persons involved in searching. 

LOCALITY: Describe the specific geographic 
location of the site. Use air distance in two 
directions (e.g., 5km N and 7.5 km W) of a 
map landmark that likely will not change 
(distance from a large town or city is not all 
that helpful). 

STATE: Use the 2-leltcr abbreviation. 

COUNTY: 

MAP NAME: List the name of the U.S.O.S. 
quadrangle or other map used to locate the 
site. 

OWNER: List the public land manager (e.g., 
Roosevelt Nat. Forest or Rocky Mu» NP), or 
name of the owner if the site is on private 
land (listing the owner's name will make it 
clear that you did not tre^ass to survey the 
site). 

ELEVATION: Circle the scale used; meters 
are preferred. 

T: township R: range S: section 

SECTION DESCRIPTION: Describe the 
location of the site within the section (e.g., 
SE !4 orNE M ofSE U) 

UTM ZONE, NORTHING, EASTING: 
Universal Transverse Mercator coordinates 



are preferred over longitude and latitude. 
The UTM 2»ne is listed on newer 
topographic maps. If you are using a map 
without the UTM grid, substitute latitude for 
Northing and longitude for Easting. 

SECTION 2 - SPECIES DATA Ust all 

amphibian species observed. If garter snakes 

are seen, list them here also. 

SPECIES: Use the scientific name. 
Convenient shorthand is to use a 4-lettcr 
code made up of the first 2 letters of the 
genus and species (e.g., Rana sylvatica 
would be RASY). 

ADULTS/JUVENILES: Indicate presence with 
a check, but numbers seen are more valuable 
data 

CALLING?: Circle Y if frogs are vocalizing 
in a breeding chorus, of if a breedii^ 
aggregation of species that don't call (e.g., 
Bufo boreas) \s observed. 

TADPOLES/LARVAE: Same as for 
adults/juveniles 

EGG MASSES: Same as above. Numbers of 
egg masses are especially valuable data. If 
possible, describe the developmental stage of 
eggs in the space for additional notes on the 
back of the form. 

METHOD: Circle how observations were 
made: VISUAL/AURAL ID - species 
identified without pickii^ it up, either by 
sight or by recognition of the breeding call; 
HAND COLLECTED - animal was picked 
up and identified in the field (higher 
confidencethan visual id); DIP NET/SEINE 
- the usual method of collection for larvae; 
TRAPPED - minncw-type traps are also 
used for larvae; VOUCHER 

COLLECTED? - circle yes or no (voucher 
specimens are recommended for every site, 
especially if identification is uncertain and 
for larvae). Indicate voucher status in 
addition to method used. 

FISH PRESENT?: If yes, list species if you 



can. Circle the question marks if you are 
not certain, but suspect that fish are present. 
ENTIRE STTE SEARCHED?: If no, list 
either the meters of shoreline or the area 
(m^ of habitat (e.g., amount of wet 
meadow) searched. 

SECTION 3 - PHYSICAL AND 
CHEMICAL DATA Wuer chemistry data are 
difficult to collect accurately without thorough 
planning and quality equipment; these data are 
optional. Vkather data are important for 
determining the quality of the observations 
(e.g., was an absence of amphibians due to 
observations made durir^ a blizzard?) 
WEATHER, WIND: Indicate atmospheric 

conditions 
AIR TEMPERATURE: Take at chest height 

in shade. The Celsius scale is preferred. 
WATER TEMPERATURE: Take 1 meter 

from margin and at 2 cm depth, or where 

egg masses are observed. 
COLOR: This is a qualitative assessment of 

whether the water clear or tea-coloied from 

organic (humic) acids. 
TURBIDTTK This is a qualitative assessment 

of whether the water clear or clouded from 

suspended particulate matter. 

SECTION 4 - HABFTAT DESCRIPTION 

These data are important for developing 

hypotheses to explain chaises in abundance of 

amphibians. This section needs to be filled out 

only once for each site (a reasonable 

amphibian survey should iitclude at least 2-3 

visits to each site in one season). 

ORIGIN: Decide whether the lake is a natural 

geologic formation or man-made. Bodies of 

water enlarged by a dam are problematic. 

List them as man-made, but add an 

explanation in the space for additional notes 

on the back of the form. 

DRAINAGE: Circle whether the site has 

permanent drainage, no drainage, or 



occasional drainage. Determining the potential for 
occasional drainage requires judgement. Look for 
clues in the topography and vegetation. 

DESCRIPTION: Decide how best to describe the site. 
If there is evidence of past or present beaver activity, 
circle one of these choices in addition to your choice. 

LENGTH, WIDTH: Record the maximum length and 
width of lakes and ponds. For streams, record the 
length and average width of the reach searched. 

MAXIMUM DEPTH: Most times, you will not have 
access to a boat, so estimate depth (deep lakes are 
usually not important to amphibians). 

STRE^AM ORDER: This is an index of stream size, and 
you will need a topographic map to determine it. First- 
order streams have no tributaries, second-oider streams 
are formed by the confluence of two I'-order streams, 
third-order streams are formed by the confluence of 
two 2"'-order streams, and so on. 

PRIMARY SUBSTRATE: Circle the type that covers 
the majority of the bottom of the site. 

EMERGEKT VEGETATION: Circle the percentage of 
the margin of the site with emergent vegetation present, 
and list the dominant species. If you are botanically- 
disad>«uitaged, list the categories of the dominant 
species (e.g., cattail, sedges, etc.). 

NORTH SHORELINE CHARACTERS: Describe the 
north shore of a lake or pond in terms of shallow water 
and emergent vegetation. This is important in 
evaluating quality of breeding habitat in some mountain 
locations. 

FOREST CHARACTERS: List the closest distance 
between the water and the surrounding forest, and list 
the most common tree species. Leave these fields 
blank if there is no forest. Describe other surrounding 
habitat types in the notes section on the back of the 
form. 



AMPHIBIAN SURVEY DATA SHEET - us HSH^WILDUFE service, 4612MeMU(WYAVE, ft. COUJNS,CO»OBJ6-9400 

leirel* eholc* lor (hadwl varitbiM; •upply v»Iu« (or othort) (ver. 2/7/92) 



DATE 



BEGIN 
TIME 



END 
TIME 



OBSERVERS 



LOCALITY 



STATE 



COUNTY 



MAP 

NAME 



OWNER 



ELEVATION 
(eirdo lealo) 



M 
FT 



SECTION 
DESCRIPTION 



UTM 

ZONE 



NORTHING 
(orLAT) 



EASTING 
lorLON) 



AMPHIBIAN AND/OR GARTER SNAKE SPEOES PRESENT 
nNOICATE NUMBERS IN CATEGORIES IF POSSIBUE) 



Cna£ METHOD AND HdCATE V 
VOUCHER SPEOMOI WAS COLLECTED 



SPEDES 



ADULTS/JUVENILES CALLWG7 TADPOLES/LARVAE 



EGG MASSES 



METHOD- 



Y N 



VISUAUAURAL ID DIP NET/SEINE 
HAND COLLECTED TRAPPED 
VOUCHER COLLECTED? YES NO 



Y N 



VISUAUAURAL ID DIP NET/SQNE 
HAND COLLECTED TRAPPED 
VOUCHER COLLECTED? YES NO 



Y N 



VISUAUAURAL ID DIP NET/SQNE 
HAND COLUCTED TRAPPED 
VOUCHER COLLECTED? YES NO 



Y N 



VISUAUAURAL ID DIP NET/SEINE 
HAND COLLECTED TRAPPED 
VOUCHER COLLECTED? YES NO 



VISUAUAURAL ID DIP NET/SEINE 
HAND COLLECTED TRAPPED 
VOUCHER COLLECTED? YES NO 



RSHPRESENn 



YES 



??? NO 



nsH 

SPEOES: 



ENTIRE SITE 
SEARCHED? 



YES 



NO 



IF NO. INDICATE 
AREA 



METERS OF SHOREUNE 
M* OF HABITAT 



PHYSICAL AND CHEMICAL ENVIRONMENT (CHEMISTRY VAAIABUS OPTIONAL - USE EXTRA SPACES FOR ADDITIONAL MEASUREMENTS! 



WEATHER: 



CLEAR 



OVERCAST 



RAIN 



SNOW 



WIND 



CALM 



UGHT 



STRONG 



AIR TEkff> 
(circlo *eala) 



WATER TEMP 
Idrdt tcalo) 



•C 
•F 



coLoa 



CLEAR 



STAINED 



TURBIDITY: 



CLEAR 



aOUDY 



pH 



ANC 



SITE DESCRIPTIONS ■ {SKETCH SITE AND PUT ADDITIONAL COMMENTS ON BACK OF SHEET) 
OMIT THIS SECTION IF DATA HAVE BEEN COLLECTED ON A PREVIOUS VISIT 



ORIGIN: 



NATURAL 



MAN-MADE 



DRAINAGE. 



PERMANENT 



OCCASIONAL 



NONE 



DESCRIPTION: 



PERMANENT 
LAKOPOND 



TEMPORARY 
UKEff>OND 



MARSH/BOG 



STREAM 



SPRING/SEEP 



ACTIVE 
BEAVER POND 



INACTIVE 
BEAVER POND 



SITE 
LENGTH (Ml 



SITE 
WIDTH (M) 



MAXIMUM DEPTH: 



< 1 M 



1 -2M 



> 2M 



STREAM ORDER 



S + 



PRIMARY SUBSTRATE: 



SILT/MUD 



SAND/GRAVEL 



COBBLE 



BOULOERmEOROCK 



OTHER 



% OF POND LAKE MARGIN WITH EMERGENT VEGETATION: 



1 -25 



25-50 



>50 



EMERGENT VEGETATION SPEDES 
lUST IN ORDER OF ABUNDANCE) 



NORTH SHOREUNE CHARACTERS: 



SHAaOWS 
PRESENT 



SHALLOWS 
ABSENT 



EMERGENT VEG 
PRESENT 



EMERGENT VEG 
ABSENT 



DISTANCE (M) TO FOREST EDGE 



FOREST TREE 
SPECIES: 



Gap/USFS Habitat 



USFWS Habii-ai- 



ROUGH SKETCH OF SITE GRID SPACING IS METERS BETWEEN LINES 



^ 












>..>■••■■>■■■« .•(•••■I 



























































































































































ADDITIONAL NOTES: 



Miscellaneous Observation Form 
Montana Natural Heritage Program 
1515 E 6th Ave 
PO Box 201800 
Helena, MT 59620-1800 



Observer_ 
Address 



Phone No. 




INSTRUCTIONS" Please use this sheet to submit sight, call, or specimen records of any Montana amphibian or reptile species. Use a separate line for each species and site. On 
the back of the sheet include any additional comments or supporting information. Please provide as specific location information as possible, particularly for the following species 
of special concern: Coeur d'Alene Salamander, Idaho Giant Salamander, Tailed Frog, Canadian Toad, Wood Frog, Snapping Turtle, Spiny Softshell, Short-homed Lizard, 
Sagebrush Lizard, Western Hognose Snake, and Smooth Green Snake. Documentation is required for Idaho Giant Salamander and Wood Frog (photo, through description, 
verification by experienced observer, etc.). An identification guide is available in the May/June 1995 issue of Montana Outdoors (reprints available at the MT Nat. Heritage Prog). 



Species 


Location 


County 


Township Range Section 
orUTM 


Date 
Mo/DayA^r 


Time 


# 
Adults 


-- D -~o 

# 

Larvae 


Example: Leopard Frog 


McNab Pond 


Carter 


T01NR59ESect 19NE 


5/20/94 


8:30a 


5 


200 


Example: Milk Snake 


3.4 mi W, 1.2 mi N of Harlowton 


Wheatland 


5145200 N, 584700 E 


8/15/94 


ll:15p 


1 




1. 
















2. 
















3. 
















4. 
















5. 
















6. 
















7. 
















8. 
















9. 
















10. 
















11. 

















Coniinciils: Include method of observation, measurements, documentation for species of special concern, disposition of specimens, weather, etc. Numbers correspond to those on 
the otiicr side of this sheet. Use additional space or sheets if necessary. 



Example: Sunny wann day, about 75°. Adults (3 seen; 2 heard calling only) at margin of ponds in cattails. Very small tadpoles seen; 1 egg mass still present. 



Example: Found dead in the road in sagebrush flat near rinirocks; 24" long; Colored with bands of yellow / black / red / black / yellow...; deposited m MSU Museum 



5. 



6. 



7. 



8. 



10. 



II. 



APPENDIX 2. 

SITES SURVEYED DURING 1993-94 

AMPHIBIAN AND REPTILE SURVEYS 



66 



Appendix 2. Sites surveyed during 1994 amphibian and reptile surveys 



Site 



Location 



Elevation 



Date 



Start Time 



LEWIS AND CLARK NATIONAL FOREST 
Rocky Mountain District 

Bean Lake* 
Beaver Basin 
Beaver Crk pond 
Blacktail Gulch* 
Falls Creek 
Kiyo Crag Lake & 
Jones Crk, . 7 mi 
Little Willow Creek, 
Lubec Lake, NE i^ 



T18N 

T20N 

T21N 

T21N 

T17N 

ponds T29N 

W of on FS144* T25N 

head of T20N 

T30N 



Palookaville beaver ponds 



T30N 



R07W 
R0 9W 
R0 9W 
R0 9W 
R07W 
R12W 
R0 9W 
R0 9W 
R13W 
R12W 



S13&24 

SO 4 NE4 

S15 

S33 SE4 

S03 S2 

S02&01 

S15 

S14 

SIO NE4 

S2 5 NE4 



Pond .4 mi past Cave Mtn trnoff* T25N R09W S26 NE4 



Teton Pass, 1.5 road mi N of* T25N R09W S06 

Teton River, 7 Lazy P. Ranch T25N R09W S26 

Wagner Basin T22N R09W S36 

Willow Creek ponds T20N R09W S23 

Wood Creek* T20N RlOW S22 

Wood Creek pond T20N RlOW S26 

Wood Creek, pond below Wood Lk T20N RlOW S36 



SE4 
SE4SE4 



4555 
6000 
4860 
4800 
4900 
6208 
5200 
5740 
5040 
5360 
5150 
6000 
5100 
4600 
5540 
5480 
5490 
5720 



04 


Jul 


94 


1615 


27 


May 


94 


1420 


27 


May 


94 


1510 


05 


Jul 


94 


1550 


05 


Jul 


94 


1030 


06 


Jul 


94 


1035 


26 


May 


94 


1630 


27 


may 


94 


1350 


25 


May 


94 


1800 


06 


Jul 


94 


1455 


26 


May 


94 


1825 


26 


May 


94 


1730 


26 


May 


94 


1515 


05 


Jul 


94 


1330 


27 


May 


94 


1245 


27 


May 


94 


0945 


27 


May 


94 


1025 


27 


May 


94 


1130 



Sites with no herps found during survey 



Appendix 2. (cont.) Sites surveyed during 1994 amphibian and reptile surveys 



Site 



Location 



Elevation 



Date 



Start Time 



JEFFERSON DIVISION 

Big Snowy Mountains 

Crystal Lake 

Crazy Mountains 

Cottonwood Creek, W Fork* 
Forest Lake 



T12N R17E S13 



T06N RIOE SIO 
T06N RIOE S26&35 



Castle Mountains 

Blackhawk Cabins meadows* T09N ROSE S36 

Checkerboard Crk, E Fork hdwtr* T09N ROSE S25&36 

Checkerboard Creek, W Fork T09N ROSE S24 

Lennep, pond . S mi N TOSN R09E S25 

Thorsen's Pond* T09N R09E S20 NW4 

Highwood Mountains 

Briggs Creek seep* T20N R09E S2S NE4NW4 

Highwood Crk, nr Cow Camp* T20N R09E S29 NW4 

Highwood Creek, middle* T20N R09E S33 center 

Highwood Creek, beaver complex* T19N R09E S09 NE4 

Highwood Creek, hdwtrs seeps* T19N R09E S15 NW4 



6000 



6100 
6490 



6S50 
6700 
6200 
5350 
6560 



4520 
4440 
46S0 
5020 
5320 



07 


Jul 


94 


1400 


OS 


Jul 


94 


0950 


07 


Jul 


94 


1S35 


OS 


Jul 


94 


1345 


OS 


Jul 


94 


1410 


OS 


Jul 


94 


1530 


OS 


Jul 


94 


1205 


OS 


Jul 


94 


1310 


2S 


May 


94 


1300 


2S 


May 


94 


1330 


2S 


May 


94 


1405 


2S 


May 


94 


1430 


2S 


May 


94 


1525 



Sites with no herps found during survey 



Appendix 2. (cont.) Sites surveyed during 1994 amphibian and reptile surveys 



Site 



Location 



Elevation 



Date 



Start Time 



JEFFERSON DIVISION (continued) 
Little Belt Mountains 

Bear Park, Basin Crk hdwtrs* 

Belt Crk Info Cntr, pond .5 mi 

Clyde Park 

Crater Lake & ponds 

Harley Park* 

Onion Park 

Russian Flat pond 

Sheep Creek complex* 

Yogo Creek, upper 



TUN 


RIOE 


S34 




6300 


SIT14N 


ROSE 


S06 


NW4NW4 


5030 


TUN 


RIOE 


S28 


N2 


6V00 


TUN 


ROVE 


S25 


SE4SW4 


5880 


T14N 


ROVE 


S33 




V350 


T13N 


ROVE 


S4&5 




V400 


TUN 


RIOE 


S12 


SW4SW4 


6330 


T12N 


ROVE 


S36 


N4 


5900 


T13N 


RIOE 


S5 




6200 



09 


Jul 


94 


1220 


28 


May 


94 


1830 


09 


Jul 


94 


1105 


29 


May 


94 


1325 


08 


Jul 


94 


1930 


09 


Jul 


94 


0815 


09 


Jul 


94 


1330 


29 


May 


94 


1035 


09 


Jul 


94 


1520 



Sites with no herps found during survey 



APPENDIX 3. 

AMPHIBIANS AND REPTILES 

OBSERVED DURING SURVEYS OF THE 

LEWIS AND CLARK NATIONAL FOREST 

IN 1994 



70 



Appendix 3. Amphibians and reptiles observed during surveys of the Lewis and Clark National 
Forest in 1993-94 

1 

Site Person Total number of adults/juv of each species observed 

Hrs:min AMMA AMTI ASTR BUBO PSTR RAPR THSI THEL 

Rocky Mountain District 

Beaver Basin 0:20 5* 

Beaver Crk pond 0:50 8 

Falls Creek 1:30 * 

Kiyo Crag Lake/pond 3:20 25* 

Little Willow Creek 0:30 6 1 

Lubec Lake, NE i^ 1 : 1 5 3 

Palookaville ponds 1:20 13* 

Teton River, 7 Lazy PI: 10 * 

Wagner Basin 3:10 * 

Willow Creek ponds 1:20 21 1 

Wood Creek pond 1:10 8* 

Wood Ck, belw Wood Lk0:40 * 18 



* 



^AMMA=Amby stoma macrodactylum; AMTI=Ambystoma tigrinum; ASTR=Ascaphus truei; BUBO=Bufo 
boreas; PSRE= Pseudacris regilla; RAPR=Rana pretiosa; THSI=Thamnophis sirtalis; 
THEL=Thamnophis elegans. 



*denotes site with breeding, i.e. tadpoles, larvae, or eggs present 



Appendix 3. (cont.) Amphibians and reptiles observed during surveys of the Lewis and Clark 
National Forest in 1993-94 



Site 



Person Total number of adults/juv of each species observed 
Hrs:min AMMA AMTI ASTR BUBO PSTR RAPR THSI THEL 



JEFFERSON DIVISION 

Big Snowy Mountains 

Crystal Lake 1:35 

Crazy Mountains 

Forest Lake 1:35 

Castle Mountains 

Checkerboard Ck, W Fk0:20 
Lennep, pond .8 mi N 0:11 



Little Belt Mountains 

Belt Crk Info Cntr 
Clyde Park 
Crater Lake & ponds 
Onion Park 
Russian Flat pond 
Yogo Creek, upper 






20 





52 


1 


00 





57 





15 


1 


00 



3 

1* 

1* 

2 

1* 

1 



^AMMA=Amby stoma macrodactylum; AMTI=Ambystoma tigrinum; ASTR=Ascaphus truei; BUBO=Bufo 
boreas; PSRE= Pseudacris regilla; RAPR=Rana pretiosa; THSI=Thamnophis sirtalis; 
THEL=Thamnophis elegans. 



*denotes site with breeding, i.e. tadpoles, larvae, or eggs present 



APPENDIX 4. 

AMPHIBIANS AND REPTILES 

REPORTED FROM IN AND AROUND THE 

LEWIS AND CLARK NATIONAL FOREST 



73 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



LONG TOED SALAMANDER 

Granite .5 to 5 mil / / No Observation 
Kitchen Creek, near Missoula 

Jefferson .5 to 5 mil 4/28/1962 No Museum Specimen 
2 mi. S. of East Helena on branch McClellen Creek 

Jefferson .5 to 5 mil 4/28/1962 No Museum Specimen 
S. of East Helena on Al Palmer Ranch 

Jefferson .5 to 5 mil 4/28/1962 Yes Museum Specimen 
2 mi. s. of East Helena on branch McClellen Creek 

Jefferson .5 to 5 mil 8/25/1959 No Museum Specimen 
Horse trough off McClellen Creek 

Jefferson .5 to 5 mil 7/ /1 972 No Museum Specimen 
Near Clancy 

Lewis & Clark <.5mile. 5/27/1994 Yes Observation 
Wood Creek, just below Wood Lake, 5700 ft. 

Lewis & Clark <.5mile. 7/5/1994 Yes Museum Specimen 
Wagner Basin, 4600 ft. 

Lewis & Clark .5 to 5 mil 7/13/1983 No Museum Specimen 
Upper Grizzly Gulch 

Missoula .5 to 5 mil 4/24/1950 No Observation 
Overflow of Union Creek, near McNamara, ca. 3500 ft. 

Missoula <.5mile. 7/1/1993 No Observation 
S of NWl/4 of section 29, pond E of Boles PL 

Missoula <.5mile. 7/1/1993 No Observation 
NW of NE of section 32, lily pad pond. 

TIGER SALAMANDER 

Cascade 5 to 10 mil 1 1/ /1992 No Observation 
Malmstrom AFB near Great F alls 

Cascade .5 to 5 mil /1 8/1 942 No Museum Specimen 
5 miles S. of Great Falls 

Chouteau > 10 miles. // No Museum Specimen 
Fort Benton 

Chouteau 5 to 10 mil 9/ /1 967 No Observation 
Teton River near Fort Benton 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



TIGER SALAMANDER (contmued) 

Glacier > 10 miles. // No Museum Specimen 
Blackfoot Indian Reservation, 1 6 miles S W of Cut Bank 

Glacier .5 to 5 mil // No Observation 
4 mi NE of Marias Pass summit 

Meagher <.5mile. 7/8/1994 Yes Observation 
Pond 0.8 mi. N. of Lennep 

Teton <.5mile. 6/22/1993 No Observation 

S.E.Pme Butte 

Unknown .5 to 5 mil /1 8/1 917 No Museum Specimen 
VoltaDam, 13.5 miles below GreatFalls 

Unknown > 10 miles. 9/10/1952 No Museum Specimen 
Lothair 

Wheatland .5 to 5 mil 7/ /1 904 No Museum Specimen 
Winnecook Ranch: Stock reservoir 

Wheatland <.5mile. / /1 994 No Observation 
Chief Joseph Park Pond, Harlowton 

TAILED FROG 

Flathead .5 to 5 mil 7/11/1934 No Museum Specimen 
Glacier National Park 3 . 5 miles above mouth of Muir creek 

Flathead > 10 miles. 7/24/1934 No Museum Specimen 
Glacier National Park, Coal Creek tributaries 

Flathead .5 to 5 mil 6/28/1934 No Museum Specimen 
Glacier National Park, 8 mi above mouth of Ole Creek 

Flathead .5 to 5 mil 7/6/1934 No Museum Specimen 
Glacier National Park, 6 mi above mouth of Park Creek 

Flathead 5 to 10 mil // No Observation 
Coal Creek 

Flathead 5 to 10 mil // No Observation 
Ole Creek 

Flathead 5 to 10 mil // No Observation 
Park Creek 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



TAILED FROG (contmued) 

Flathead .5 to 5 mil / /1 969 No Observation 
Soldier and Clark Creeks 

Flathead .5 to 5 mil / /1 969 No Observation 
Bear and Skyland Creeks 

Flathead .5 to 5 mil // No Museum Specimen 
Midvale Creek, near Glacier National Park 

Granite 5 to 10 mil 7/24/1958 No Museum Specimen 
Ranch Creek, ca. 1 1 mi. S. of I 90 on Rock Creek Rd. 

Granite .5 to 5 mil 7/24/1958 No Museum Specimen 
Ranch Creek (tributary of Rock Creek) 

Lewis & Clark 5 to 10 mil // No Observation 
Tributary of Copper Creek, tributary of the Blackfoot River 

Lewis & Clark <.5mile. 7/5/1994 Yes Museum Specimen 
Falls Creek, crossing down to falls. 

Missoula .5 to 5 mil 5/4/1947 No Observation 
Spring on Gold Creek, near Gold Creek Ranger Station 

Missoula 5 to 10 mil 7/22/1950 No Museum Specimen 
Placid Creek, ca. 4 mi. SW of Seeley Lake 

Missoula .5 to 5 mil / /1 969 No Observation 
Owl Creek 

Pondera < .5 mile. 10/7/1994 Yes Observation 
E Fork Woods Creek; also sec. 16, Whiterock Creek 

Powell 5 to 10 mil // No Observation 

Morrell Creek, tributary of Clearwater River 

Teton <.5mile. 7/8/1994 Yes Observation 

On Mt. Wright hiking trail at small stream that trail crosses. 

WESTERN TOAD 

Broadwater > 10 miles. / /1 966 Yes Observation 
Near Canyon Ferry Reservoir, small spring in open ponderosa pine. 

Carbon .5 to 5 mil / /1 966 No Observation 

Kmgs Hill, 7000 ft. 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



WESTERN TOAD (contmued) 

Cascade .5 to 5 mil // No Museum Specimen 
Little Belt Mountams, 1 .2 miles N. (Hwy 89) of Kings Hill Pass, 7200 ft. 

Chouteau .5 to 5 mil 6/24/1962 Yes Museum Specimen 
Highwood Creek near Arrow Creek Divide 

Chouteau .5 to 5 mil 6/24/1962 No Museum Specimen 
Highwood Creek near Arrow Creek Divide 

Chouteau .5 to 5 mil 6/24/1962 Yes Museum Specimen 
Upper Highwood Creek near pass to Arrow Creek 

Chouteau .5 to 5 mil 6/24/1962 Yes Museum Specimen 
Upper Highwood Creek near pass to Arrow Creek 

Flathead >10miles. 7/6/1934 No Museum Specimen 
Park Creek, Glacier National Park. 

Flathead .5 to 5 mil 7/23/1949 No Observation 
South Fork River and Addition Creek 

Granite .5 to 5 mil 7/11/1977 Yes Museum Specimen 
1/4 mi W of Bearmouth, rest area off I 90. 

Lewis & Clark .5 to 5 mil 7/12/1958 No Museum Specimen 
BearLake, 7000 feet 

Meagher > 10 miles. 8/4/1899 No Museum Specimen 
Deep Creek Canyon, Big Belt Mountains 

Meagher 5 to 1 mil 9/5/1918 No Museum Specimen 
6 mi SW of Lennep. 

Meagher > 10 miles. 8/23/1919 No Museum Specimen 
Fort Logan, Camas Creek (4 mi S). 

Meagher .5 to 5 mil 8/20/1951 No Museum Specimen 
Sheep Creek near Jumping Creek Campgrounds 

Meagher .5 to 5 mil 8/24/1951 Yes Museum Specimen 
Adams Ranch, Sheep Creek 

Meagher .5 to 5 mil 8/ 6/1958 No Museum Specimen 
Lake Creek 



Natural Heritage Program 03/04/1 995 

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County Precision Date Breed Data Type 



WESTERN TOAD (contmued) 

Missoula .5 to 5 mil 5/2/1948 No Observation 
Morrel Creek near Seeley Lake; Archibald Creek near Seeley Lake, 4000 ft. 

Missoula .5 to 5 mil / /1 966 No Observation 
Rainy Lake 

Missoula <.5mile. 7/1/1993 No Observation 
S of NWl/4 of section 29, Pond E of Boles Point. 

Phillips .5 to 5 mil 8/10/1972 No Museum Specimen 
Slippery Ann Station 

Pondera <.5mile. 7/13/1994 No Observation 
Damp area along E Fork Woods Creek. 

Teton <.5mile. 5/26/1994 Yes Observation 

7 Lazy P Ranch, Lewis and Clark NF, 5 100 ft. 

Teton <.5mile. 6/21/1993 Yes Observation 

NW of NW section 13, Pine Butte Swamp Preserve. 

Wheatland > 10 miles. / /1 966 No Observation 
Grassy spring in the short grass prarie W. of Harlowton 



GREAT PLAINS TOAD 

Toole .5 to 5 mil 7/20/1950 Yes Observation 

cattle pond on the prairie apprx. 1 mi S of camp ( 1 mi S of Galata) 

WOODHOUSEiS TOAD 

Fergus .5 to 5 mil 7/24/1919 No Museum Specimen 
On Missouri River 5 miles north of Wilder 

Fergus .5 to 5 mil 8/9/1991 No Museum Specimen 
Missouri River at Fred Robinson Bridge. 

Mussellshell . 5 to 5 mil 8/7/1918 No Museum Specimen 
1 mile west of Kline 

WESTERN CHORUS FROG 

Cascade <. 5 mile. 5/28/1994 No CallHeardOnly 
On Hwy 89 go 5 mi. W. of Hwy 200 junction to pond on N. side of road 



Natural Heritage Program 03/04/1 995 

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County Precision Date Breed Data Type 



WESTERN CHORUS FROG (contmued) 

Cascade 5 to 10 mil 6/7/1993 No Observation 
Chestnut Valley Sand hills 

Chouteau 5 to 10 mil 5/ /1 967 No Observation 
Near Fort Benton 

Chouteau .5 to 5 mil 7/22/1962 Yes Museum Specimen 
Swan Ranch E. of Highwood 

Fergus .5 to 5 mil 7/26/1991 No Museum Specimen 
5 mi. E. of Roy 

Glacier < .5 mile. 5/25/1 994 No Call Heard Only 
Both ponds at intersection, T30NR11W S16 

Judith Basm <.5mile. 5/27/1994 No Call Heard Only 
T18NR10ES20 

JudithBasin <.5mile. / /1 994 No Observation 
AlsoT19NR09ESec.22,offFSRD122. 

JudithBasin <.5mile. 5/28/1994 No Museum Specimen 
5 mi. N., 3 mi. W. of Geyser, 4370 ft. 

Lewis& Clark <. 5 mile. 5/27/1994 No CaU Heard Only 
Anderson Lake 

Liberty <.5mile. 7/7/1994 Yes Museum Specimen 
Moffat Bridge on Marias River, ca. 6 mi. ESE of Tiber Dam 

Petroleum 5 to 10 mil 8/ /1 968 No Museum Specimen 
Near War Horse Lake 

Pondera <.5mile. 5/26/1994 No Museum Specimen 
T28NR8WS5 

Pondera <.5mile. 7/13/1994 No Observation 
Spring seep area T29NR12W S9. 

Teton <.5mile. 6/21/1993 No Observation 

NEl/4 of NEl/4 S34, near the Teton River, Pine Butte Swamp Preserve. 

Teton <.5mile. 6/21/1993 No Observation 

NW of NW sec 13 Pine Butte Swamp Preserve. 



Natural Heritage Program 03/04/1 995 

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County Precision Date Breed Data Type 



WESTERN CHORUS FROG (contmued) 

Teton <.5mile. 6/22/1993 No Observation 

SE of SE of S2, Pine Butte Swamp Preserve. 

Teton <.5mile. 6/22/1993 No Observation 

Bellview Rd Pond 

Teton <.5mile. 6/22/1993 No Observation 

NEl/4 of Sect 17, Pine Butte Swamp Preserve. 

Teton <.5mile. 6/22/1993 No Observation 

SEPme Butte 

Toole > 10 miles. 7/ /1 950 No Observation 

in the area near camp ( 1 mi S of Galata) 

Wheatland > 10 miles. 8/29/1918 No Museum Specimen 
5 miles SW of Oka 

PLAINS SPADEFOOT 

Cascade .5 to 5 mil / /1 934 No Museum Specimen 
4.5 miles W. of Great Falls 

Cascade .5 to 5 mil 8/31/1948 No Museum Specimen 
3 mi. S. of Cascade, Missouri River 

Cascade .5 to 5 mil 9/ /1 962 No Museum Specimen 
NE of Great FaUs 

Cascade <. 5 miles / /1 992 Yes Observation 
Great Falls 

Toole .5 to 5 mil 7/21/1950 No Observation 

dry flood plain at camp (10 mi S of Galata) 

NORTHERN LEOPARD FROG 

Broadwater <.5mile. 5/10/1993 No Observation 
Deepdale FAS. Directly E. across channel from Deepdale BE nest 

Cascade <. 5 miles 6/ /1 994 No Observation 
cutoff slough along Belt Creek, T 1 8N R6E Sec 1 2 

Lweis and Clark <. 5 miles / /1993 No Observation 
small riparian ponds along Dearborn River, T 1 8N R7 W Sec. 25 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



NORTHERN LEOPARD FROG (contmued) 

Chouteau .5 to 5 mil 6/24/1962 No Museum Specimen 
Upper Highland Creek near Arrow Creek Divide. 

Chouteau .5 to 5 mil 6/24/1962 No Museum Specimen 
Upper extreme of Highwood Creek, near mouth of Beaver Creek 

Chouteau .5 to 5 mil 8/31/1948 No Museum Specimen 
3 mi. S. of Cascade, Missouri River 

Fergus .5 to 5 mil 8/20/1918 No Museum Specimen 
8 miles W. of Lewistown, on Beaver Creek 

Fergus .5 to 5 mil 8/19/1918 No Museum Specimen 
7 miles NE of Lewistown, Judith Mountains 

Fergus .5 to 5 mil 7/28/1919 No Museum Specimen 
7 miles NE of Hilger 

Fergus .5 to 5 mil 8/4/1919 No Museum Specimen 

5 miles NW of Hilger, Moccassin Mountains 

Fergus <.5mile. 8/28/1994 No Observation 
T13NR22ES23 

Fergus 5 to 10 mil 8/13/1948 No Museum Specimen 
Beaver Creek, S. of Lewistown 

Lewis & Clark 5 to 10 mil 7/1 8/1 958 No Museum Specimen 
Sun River, 5500 feet 

Liberty 5 to 10 mil 6/13/1950 No Observation 

6 miles east of reservoir area at Turner Memorial Park 

Meagher .5 to 5 mil 8/30/1918 No Museum Specimen 
Martinsdale, S. fork of Missellshell River 

Meagher .5 to 5 mil 10/21/1950 No Museum Specimen 
S. Fork Mussellshell near Lennep 

Powell 5to 10 mil 7/3/1973 No Museum Specimen 
North Fork of Blackfoot River 

Teton <.5mile. 7/21/1993 No Observation 

NE of NE of sect 7, Pine Butte Swamp Preserve. 

Teton .5 to 5 mil 8/27/1949 No Museum Specimen 

5 mi. E. of Choteau. 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



NORTHERN LEOPARD FROG (contmued) 

Wheatland .5 to 5 mil /1 8/1 942 No Museum Specimen 
Musselshell River at Shawmut 

Wheatland . 5 to 5 mil 8/28/1 9 1 8 No Museum Specimen 
5 miles SW of Oka 

Wheatland . 5 to 5 mil 8/28/1 9 1 8 No Museum Specimen 
3 miles N. of Judith Gap 

SPOTTED FROG 

Cascade .5 to 5 mil // No Museum Specimen 
Little Belt Mountains, 1.2 miles N. of Kings Hill Pass 

Cascade .5 to 5 mil 8/8/1944 No Museum Specimen 
12 miles S. of Neihart 

Cascade .5 to 5 mil 6/27/1973 No Museum Specimen 
North of Kings Hill Campground, Hwy. 89, Little Belt Mountains 

Cascade <.5mile. 5/28/1994 No Museum Specimen 
Pond 0.5 mi N. of Belt Creek Info Center on E. side of Hwy. 89 

Flathead .5 to 5 mil 7/25/1934 No Museum Specimen 
Galcier National Park, Marthais Basin Lake 

Glacier <.5mile. 5/25/1994 No Observation 
NE 1/4 of Lubec Lake, 5040 ft. 

Glacier <.5mile. 7/6/1994 Yes Observation 
Palookaville Beaver Ponds 

Granite .5 to 5 mil 7/11/1977 No Museum Specimen 
0.25 mile W. of Bear Mouth, at rest area off Hwy. 90. 

Jefferson .5 to 5 mil 7/8/1944 No Museum Specimen 
5 miles W. of Bemice 

JudithBasin <.5mile. 5/20/1994 No Observation 
Along tributary to Dry Pole Gulch. 

JudithBasin <.5mile. 7/9/1994 Yes Observation 
Clyde Park, Little Belt Mountains, 6700 ft. 

JudithBasin <.5mile. 7/9/1994 No Observation 
Upper Yogo Creek, Little Belt Mountains, 6200 ft. 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



SPOTTED FROG (contmued) 

Judith Basin <.5mile. 7/9/1994 Yes Observation 
Pond in Russian Flat, Little Belt Mountains 

JudithBasin .5 to 5 mil 7/12/1991 Yes Museum Specimen 
Clyde Park Pond, Litle Belt Mountains 

Lewis & Clark .5 to 5 mil // No Observation 
2.3 miles W. of Flasher Pass tributary of Blackfoot River 

Lewis & Clark <.5mile. 5/27/1994 Yes Museum Specimen 
Wood Creek, just below Wood Lake 

Lewis & Clark <.5mile. 5/27/1994 Yes Observation 
Willow Creek Ponds, 5540 ft., also Sec. 14 

Lewis & Clark <.5mile. 5/27/1994 No Observation 
Pond above Beaver Creek, S. of Green Timber Basin. 

Lewis & Clark < .5 mile. 9/ 3/1 994 No Observation 
Prickley Pear Creek just W. of Stansfield Lake. 

Lewis & Clark <.5mile. 5/27/1994 No Observation 
Head of Little Willow Creek. 

Lewis & Clark <.5mile. 5/27/1994 Yes Observation 
Beaver Basin (road sign) 

Lewis & Clark <.5mile. 5/27/1994 Yes Museum Specimen 
Wood Creek 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
Pond, ca. 400 m from the main stack of East Helena Smelter 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
Ca. 1.6 km SSE from the East Helena Smelter in springs 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
McClellan Creek Quarry Lake 

Lewis & Clark <.5mile. 5/22/1994 No Museum Specimen 
7 Up Pete Proposed Gold Mine area on HWY 200, ca. 10 mi. E. of Lincoln. 

Meagher . 5 to 5 mil 9/3/1918 No Museum Specimen 
Martinsdale, S. Fork of Mussellshell River 

Meagher .5 to 5 mil 6/ 1/1978 No Museum Specimen 
Dry Fork of Musselshell River 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



SPOTTED FROG (contmued) 

Meagher <.5mile. 5/29/1994 Yes Museum Specimen 
Crater Lake and ponds above, 5880 ft. 

Meagher <.5mile. 7/7/1994 No Museum Specimen 
Forest Lake, Crazy Mountains, also Sec. 35, 6490 ft. 

Meagher <.5mile. 7/8/1994 No Observation 
W. fork Checkerboard Creek, Castle Mountains, 6200 ft. 

Meagher <.5mile. 7/9/1994 No Observation 
Onion Park, Little Belt Mountains, also sec. 5. 

Meagher .5 to 5 mil 8/ 6/1958 No Museum Specimen 
Lake Creek 

Meagher .5 to 5 mil 5/13/1950 No Museum Specimen 
Near Ringling Hot Well 

Meagher .5 to 5 mil 8/ 6/1958 No Museum Specimen 
Lake Creek 

Missoula .5 to 5 mil 5/18/1948 No Observation 
Overflow of Union Creek, near McNamara 

Missoula <.5mile. 7/2/1993 No Observation 
T16NR15WS25 

Pondera <.5mile. 7/6/1994 Yes Observation 
Kiyo CragLake andPond, and sm. pond on RD 9218 

Powell .5 to 5 mil 7/21/1891 No Museum Specimen 
Elliston, near Little Blackfoot River 

PAINTED TURTLE 

Broadwater <.5mile. 5/ /1 993 No Observation 
Canyon Ferry WMA east side. 

Broadwater 5 to 10 mil // No Specimen Reported 
See map in Black 1 970 

Lewis & Clark <.5mile. 6/9/1994 No Observation 
W. of Augusta along Benchmark Rd. W. of Nilan Reservoir, next to 2 ponds 

Powell .5 to 5 mil 9/7/1994 No Observation 
Lahrity Lake in Ovando Valley 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



PAINTED TURTLE (cont.) 

Powell .5 to 5 mil 9/ 8/1 994 No Observation 
Evans Lake 

Powell 5 to 1 mil / / No Specimen Reported 
See map in Black 1 970 

Teton 5 to 1 mil / / No Specimen Reported 

See map in Black, 1 970 

SPINY SOFTSHELL 

Broadwater 5 to 10 mil // No Specimen Reported 
Canyon Ferry Reservoir (may be erroreous) 

Wheatland .5 to 5 mil /1 7/1 942 No Museum Specimen 
Musselshell River, near Shawmut 

Wheatland .5 to 5 mil // No Museum Specimen 
Musselshell River near Shawmut. 

SHORT HORNED LIZARD 

Chouteau > 10 miles. // No Museum Specimen 
Near Fort Benton 

Chouteau 5-10 miles spring 1985 No Observation 
Cascade Hutterite Colony west of Ulm in breaks, 3500 ft. 

Golden Valley > 10 miles. 8/ /1 894 No Museum Specimen 
Painted Robe Creek 

Toole > 10 miles. / /1 950 No Observation 

tops of ridges between coulees near camp (10 mi S of Galata) 

Wheatland .5 to 5 mil /29/1933 No Museum Specimen 
Harlowton 

SAGEBRUSH LIZARD 

Mussellshell 5 to 10 mil 8/3/1908 No Museum Specimen 
Roundup, 8 mi S of 

RUBBER BOA 

Broadwater 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark .5 to 5 mil 3/ /1 949 No Museum Specimen 
Worth Ranch, Canyon Creek 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



RACER 

Flathead 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Toole .5 to 5 mil 7/26/1950 No Observation 

in a couleejustN of camp (10 mi S of Galata) 

WESTERN HOGNOSE SNAKE 

Mussellshell 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Teton 5 to 1 mil / / No Specimen Reported 

See map in Davis 1 963 

GOPHER SNAKE 

Broadwater <.5mile. 5/21/1994 No Observation 
Hwy. 12 between Helena and Townsend 

Cascade 5 to 10 mil 7/6/1993 No Observation 
Chestnut Valley Sandhills 

Cascade 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Cascade <. 5 miles 6/ /1 994 No Observation 
near Belt Creek, T 1 8N R6E Sec 1 2 

Chouteau .5 to 5 mil // No Museum Specimen 
Fort Benton 

Lewis&Clark .5 to 5 mil 10/15/1982 No Museum Specimen 
Junction Sheep and Little Prickly Pear Creeks 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



GOPHER SNAKE (contmued) 

Toole > 10 miles. / /1 950 No Observation 

9 from river valley, 1 from coulees, 2 from Prairie (approx. 1 mi S of Ga) 

Wheatland .5 to 5 mil / /1 932 No Museum Specimen 
Harlowton 

Wheatland .5 to 5 mil // No Museum Specimen 
20 miles E. of Harlowton 

WESTERN TERRESTRIAL GARTER SNAKE 

Broadwater .5 to 5 mil 9/ /1 962 No Museum Specimen 
W. of Winston 

Cascade <.5mile. 5/28/1994 No Observation 
Pond 0.5 mi N. of Belt Creek Info Center on E. side of Hwy. 89 

Cascade .5 to 5 mil 10/15/1992 No Museum Specimen 
5 mi. SE of Fairfield 

Chouteau .5 to 5 mil // No Museum Specimen 
Fort Benton 

Chouteau .5 to 5 mil 7/21/1962 No Museum Specimen 
Shankin Creek, Highwood Mountains 

Chouteau .5 to 5 mil 5/ /1 963 No Museum Specimen 
N. Fork Highwood Creek 

Fergus .5 to 5 mil 8/23/1954 No Museum Specimen 
22 miles S., 12 miles E. of Lewiston 

Fergus .5 to 5 mil 7/2/1961 No Museum Specimen 
3milesW., 18milesS. of Lewiston, Big Snowy Mts. 

Fergus .5 to 5 mil 8/15/1918 No Museum Specimen 
Heath, 1 5 mi S of. Big Snowy Mountains 

Fergus 5 to 10 mil 8/4/1919 No Museum Specimen 
Hilger, 5 mi NW of. Mocassin Mountains 

Fergus 5 to 10 mil 7/27/1919 No Museum Specimen 
Hilger, 7 mi NE of 

Fergus <.5mile. 7/7/1994 No Observation 
Crystal Lake, Big Snowy Mountains, 6000 ft. 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



WESTERN TERRESTRIAL GARTER SNAKE (contmued) 

Gallatin 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Judith B asin . 5 to 5 mil 8/7/1919 No Museum Specimen 
Stanford, 20 SW of. Little Belt Mtns, on Dry Wolf Creek 

JudithBasin .5 to 5 mil 8/13/1919 No Museum Specimen 
Geyser, Otter Creek, (10 mi SW) 

Lewis & Clark .5 to 5 mil // No Museum Specimen 
Sun River at U. S. Hy 287, N of Augusta 

Lewis& Clark >10miles. 7/20/1891 No Museum Specimen 
Helena, near, Mc Clellan Creek 

Lewis & Clark > 10 miles. 7/18/1958 No Museum Specimen 
Sun River, elevation 5500 ft. 

Lewis & Clark <.5mile. 5/27/1994 No Observation 
Head of Little Willow Creek 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
Pond, ca. 400 m from the main stack of East Helena Smelter 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
Where Prickly Pear Creek flows along slag pile at East Helena Smelter 

Lewis & Clark <.5mile. 7/27/1975 No Observation 
McClellan Creek Quarry Lake 

Liberty > 10 miles. II /1950 No Observation 
Liberty County about 6 mi. W of camp (approx. 10 mi. S of Galata) 

Meagher . 5 to 5 mil 9/4/1918 No Museum Specimen 
Grovedale (Groveland), 3 mi W of 

Meagher .5 to 5 mil 8/30/1918 No Museum Specimen 
Martinsdale, S Fork of Musselshell River 

Meagher 5 to 10 mil 8/22/1919 No Museum Specimen 
Fort Logan, Camas Creek, (4 mi S) 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



WESTERN TERRESTRIAL GARTER SNAKE (contmued) 

Meagher . 5 to 5 mil 8/31/1918 No Museum Specimen 
Martinsdale, S fork of Musselshell River 

Missoula <.5mile. 7/1/1993 No Observation 
Grassy hillside, 100m from riparian area Clearwater River 

Missoula <.5mile. 9/ /1 950 No Specimen Reported 
Upper Holland L ake 

Missoula <.5mile. 9/ /1 950 No Specimen Reported 
Gyp Mountain 

Powell > 10 miles. 7/31/1967 No Museum Specimen 
North Fork of the Blackfoot River 

Powell > 10 miles. 7/3/1973 No Museum Specimen 
North Fork of the Blackfoot River 

Powell <.5mile. 6/ /1 950 No Specimen Reported 
Cottonwood Creek 

Teton .5 to 5 mil 10/16/1982 No Museum Specimen 

10 mi. W. of Chouteau 

PLAINS GARTER SNAKE 

Cascade .5 to 5 mil 8/31/1894 No Museum Specimen 
Great Falls 

Chouteau .5 to 5 mil // No Museum Specimen 
Fort Benton 

Toole .5 to 5 mil 7/16/1950 No Observation 

margin of cattle pond 1 .5 mi S of camp(10 mi S Of Galata 

COMMON GARTER SNAKE 

Carbon .5 to 5 mil // No Museum Specimen 

Fort Benton 

Cascade .5 to 5 mil 9/6/1994 No Observation 
Schrammeck L ake 

Chouteau .5 to 5 mil 6/24/1962 No Museum Specimen 
Upper Highwood Creek near pass to Arrow Creek 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



COMMON GARTER SNAKE (contmued) 

Fergus .5 to 5 mil 8/16/1918 No Museum Specimen 
1 mile SE of Lewistown 

Golden Valley . 5 to 5 mil 1 0/10/1 994 No Observation 
Near Big Coulee Creek 

JudithBasin <.5mile. 5/20/1994 No Observation 
Along tributary to Dry Pole Gulch. 

Lewis & Clark <.5mile. 5/27/1994 No Observation 
Willow Creek Ponds, 5540 ft. 

Lewis& Clark <. 5 mile. 5/5/1994 No Observation 
Smith Creek 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Missoula <.5mile. 7/2/1993 No Observation 
NWl/4of SEl/4 Sec25, lOmfrom water 

Teton <.5mile. 7/5/1993 No Observation 

Durr Place, outside tack room. Pine Butte Swamp Preserve 

WESTERN RATTLESNAKE 

Cascade 5 to 10 mil 9/ 1/1894 No Museum Specimen 
Great Falls 

Cascade .5 to 5 mil /10/1937 No Museum Specimen 
7 miles NW of Cascade 

Cascade .5 to 5 mil 7/29/1972 No Museum Specimen 
3 mi. N. Manchester 

Cascade .5 to 5 mil / /1 927 No Museum Specimen 
Cascade 

Chouteau .5 to 5 mil 10/ /1 937 No Museum Specimen 
30 miles east of Brady, 2 miles south of Russell 

Chouteau .5 to 5 mil // No Museum Specimen 
Ft. Benton 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



WESTERN RATTLESNAKE (contmued) 

Fergus .5 to 5 mil 7/24/1919 No Museum Specimen 
5 miles north of Wilder on Missouri River 

Gallatin 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Lewis & Clark .5 to 5 mil 7/28/1 949 No Museum Specimen 
Wirth Ranch 

Liberty <.5mile. 7/6/1994 No Observation 
5.3 mi. E. of Tiber Dam on Prospect Road, 3030ft., also sec. 10. 

Mussellshell 5 to 10 mil // No Specimen Reported 
See map in Davis 1 963 

Toole > 10 miles. 7/24/1950 No Observation 

Marias river near camp (10 mi south of Galata) 

Wheatland .5 to 5 mil // No Museum Specimen 
Harlowton 

Wheatland .5 to 5 mil // No Museum Specimen 
2 miles S. of Hedgesville, 20 miles E. of Harlowton 

Wheatland > 10 miles. 8/12/1903 No Museum Specimen 
vie of Harlowton 

Wheatland > 10 miles. 4/16/1967 No Museum Specimen 
vie Harlowton 

Wheatland > 10 miles. 5/11/1984 No Museum Specimen 
vie of Harlowton 

Wheatland <.5mile. / /1 989 No Observation 
Behind USPS office, Harlowton 

Wheatland .5 to 5 mil /24/1 932 No Museum Specimen 
8 miles S. of Harlowton 

Wheatland .5 to 5 mil /29/1932 No Museum Specimen 
8 miles S. of Harlowton 

Wheatland .5 to 5 mil /22/1933 No Museum Specimen 
2 miles S Hedgesville 



Natural Heritage Program 03/04/1 995 

Montana Animal Atlas (Herpetile) Species Report 

County Precision Date Breed Data Type 



WESTERN RATTLESNAKE (contmued) 

Wheatland .5 to 5 mil / /1 935 No Museum Specimen 
8 miles S. of Harlowton 

Wheatland .5 to 5 mil / /1 932 No Museum Specimen 
2 miles S Hedgeville 

Wheatland .5 to 5 mil /24/1926 No Museum Specimen 
8 miles S. of Harlowton 

Yellowstone 5 to 1 mil / / No Specimen Reported 
See map in Davis 1 963