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UNIVERSITY OF ILLINOIS
Volume III
Issued by Agronomy Department, University of Illinois
During the Period September 5^ 1955^ to June l8, 1956
AGITI^
ACES LIBRARY
UNIVERSITY OF ILLINOIS
1101 S. GOODWIN AVE.
URBANA.IL 61801
Digitized by the Internet Archive
in 2011 with funding from
University of Illinois Urbana-Champaign
http://www.archive.org/details/agronomyfacts03univ
TABLE OF CONTENTS
Miscellaneous
Pollination and Fertilization M-13
The Function of CO in Crop Production K-l4
Radiation and Plant Breeding K-15
The Yellow Dwarf (Red Leaf) Disease of Oats M-I6
Corn
Corn Hybrids for Specialized Farm and Market Uses C-7
Tomorrow's Hybrid Corn C-8
Temperature and Other Interrelated Factors in Drought Damage
to the Corn Plant C-9
How to Estimate Hail Losses to Corn C-10
Forage Crops
Identifying Common Legume Seedlings F-IT
Selecting Alfalfa Varieties F-I8
How Hybrid Sorghums Were Developed and Are Being Produced F-I9
Sudangrass in Illinois F-20
Orchardgrass and Its Management F-21
Small Grains
Winter Barley in Illinois G-I3
Winter Wheat G-lU
"Blast" in Oats G-I5
Winter Rye in Illinois^ G-I6
Soybeans
Root and Stem Rot of Soybeans E-5
Estimating Hail Losses to Soybeans S-6
Soil Fertility and Testing
Chemistry of Organic Nitrogen in Soils SF-37
1955 Wheat Yields--Illinois Soil Experiment Fields SF-38
Earthworms SF-39
Band Application of Fertilizers in Illinois — Part 1 SF-40
Band Application of Fertilizers in Illinois — Part 2 SF-4l
Progress Report on a Green-Manuring Project SF-U2
Composts SF-i|-3
The Nature of Exchangeable Calcium and Magnesium and Their
Relation to Soil Acidity and Lime Requirement SF-^ij-
-2-
Soil Management and Conservation
Objectives of Crop Rotations — Introduction and Erosion Control . . .SM-12
Effect of Crop Rotations on Soil Physical Condition SM-I3
Crop Rotations and Insects 5M-1^
An Analysis of the Nitrogen Status of the Agronomy South
Farm Rotations SM-I5
Economic Ohjectives of Crop Rotations £M-l6
Continuous Corn SM-IT
Soil Properties
The Productivity of Some Important Southern Illinois Soils SP-9
Corn Root Distribution in Fertilized and Unfertilized Flanagan
Silt Loam SP-10
The Productivity of Dark, Till-Derived Soils in Northeastern
Illinois SP-11
Bottomland Soils of Illinois SP-12
Organic Soils in Illinois SP-I3
Fragipans in Illinois Soils SP-1^
Basis for Separating and Classifying Soils SP-I5
Weed Control
Controlling Wild Garlic and Wild Onion W-6
Reaction of Various Weeds and Brush to 2,4-1 and 2,k,'^-T V-J
J.-/P: sc
Aug. 1955
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
MISCELLANEOUS
AGRONOMY FACTS
M-13
POLLINATION AND FERTILIZATION
The pollen that we see flying in fields
of corn in July and August plays an im-
portant pai-t in the production of the
corn crop. These tiny grains are essen-
tial for the formation of the kernel;
■without them the cob vciold he hare and
useless. Pollination is essential not
only in producing corn, hut in producing
all other crops. Without this process,
almost all of the seed plants would
disappear, leaving only those that repro-
duce by cuttings or other special mechan-
isms.
Pollen is produced by the male flowers
of the corn tassel. The female flowers
are borne on the ear. Plants of this
sort, in which the flowers are carried
separately, are said to have imperfect
flowers. Most plants have perfect
flowers; that is, the male and female
parts are carried within a single flower,
as in wheat, oats and soybeans. Plants
that have perfect flowers are often
self -fertilized, while those that have
imperfect flowers are usually cross-
fertilized. In many species with per-
fect flowers, stray pollen is usually
excluded because pollination occurs be-
fore the flower actually opens. This
makes it virtually certain that self-
pollination will take place.
At the time of sexual maturity, matiure
pollen is released from anthers in large
amounts. It has been estimated that a
single corn tassel will produce as much
as 25, 000, ceo pollen grains. This esti-
mate is probably high, but even the most
conservative estimate would be that each
silk produces at least 9^000 pollen
grains ,
Mature pollen is transferred from the
anther to a receptive surface, called a
stigma, by wind, gravity, or insects.
Upon alighting, the pollen grain germi-
nates, and the pollen tube grows down
the style (silk in corn) until it reaches
the ovary.
Fertilization in Typical Grass
Species (Schematic)
,Silk (stigma)
Polar
Nuclei
Germinated
Pollen I
Pollen Tube
Sperm
In corn this process usually takes 2k-
to 26 hours. Two sperm move down the
pollen tube and enter the ovary. One of
these sperm unites with the egg and the
other unites with two polar nuclei. The
egg and sperm union produces the embryo
(the new plant), and the polar nuclei-
sperm \inion produces the endosperm.
This process is called fertilization.
Cell division proceeds in these tissues
\intil the mature seed is produced.
Endosperm in corn consists of the aluerone
layer and the soft and horny starch of
the kernel. In soybeans the endosperm
is microscopic, and the food for the new
embryo is stored in stinctiires called
cotyledons. Cotyledons are a part of the
embryo itself, and in young soybean seed-
lings they look much like two thickened
leaves ,
Envirorment plays an important part in
the success of pollination. For ex-
ample, high humidity quickly bursts pol-
len of red clover. The higher seed set
of this species in dry climates is doubt-
less due to the lower humidity. High
temperatures usually reduce the time
that pollen will live. Species vary in
the length of time pollen remains via-
able. Barley pollen is particularly
short-lived if removed from the flower.
In contrast, wheat pollen can be col-
lected and stored at room temperatures
for as long as six hovirs and still ef-
fect fertilization. Corn pollen remains
viable for a shorter period, possibly no
longer than two or three hours under
ordinary field conditions. Viability is
lost much more quickly when temperatures
are high than when they are low.
Under Illinois conditions, pollen via-
bility rarely limits corn production.
In 100° temperatures, pollen remains via-
ble for only a short time. During early
morning hours when temperatures are
lower, enough pollen is usually shed to
insure a satisfactory seed set. How-
ever, extremely high planting rates or
drouth often causes late silking and many
ears may not be adequately pollinated.
The pollination habits of a crop species
largely determine the methods that plant
breeders may use in improving that crop.
In corn, the male and female flowers are
widely separated, and hybrids may be
produced easily and inexpensively. Thus
their large-scale use is practical.
Oats, wheat, and soybeans, however, can-
not be easily hybridized because the
flowers are perfect and tiny. Hybrids
of these crop plants will therefore prob-
ably never be ccmmercially available.
D. E. Alexander
10-17-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
MISCELLANEOUS
AGRONOMY FACTS
M-14
THE FUNCTION OF CO2 IN CROP PRODUCTION
Carbon constitutes kO-k-^ percent of the
dry weight of most crop plants. Wo
other element contributes as many atoms
to the formation of a plant. Carbon mol-
ecules linked one to another form the
basic skeleton of the carbohydrates^
fats^ proteins, and other substances
that make up a plant. The oxidation or
"burning" of the reduced carbon in fat
and sugar provides the energy for plant
growth. Obviously, an adequate supply
of carbon is imperative for good crop
production.
The basic supply of carbon for plants is
in the form of a gas, carbon dioxide, or
CO2. Carbon in this form is at its low-
est energy level — completely oxidized
or burned--and is very stable. The nor-
mal CO2 concentration of the atmosphere
is about three parts in 10,000 parts of
air, or O.O3 percent. Except in the vi-
cinity of the soil or photosynthesizing
plants, this figure is relatively con-
stant. The constancy is maintained by a
vast reservoir of CO2 dissolved in the
oceans, both
as the gas and as carbon-
ates (the reaction product of CO2 and
H2O). Air passing over the oceans gains
or loses CO2 until the normal concentra-
tion is re-established, and large m.asses
of air continually move from oceans to
land and off to the oceans again.
The plant absorbs nearly all of the CO2
it requires frcm the atm.osphere, the gas
diffusing in through small pores in the
leaves. Some CO2 is absorbed by the
roots and is translocated to the leaves
in the upward-moving sap, but the amount
so supplied is believed to be of minor
importance (about 5 percent at most).
Plant leaves are amazingly efficient in
absorbing CO2. They can remove as much
as 50 percent of the CO2 from a layer of
air which is rapidly passed over photo-
synthesizing corn leaves . If it were
not for this high absorption efficiency,
plants would make very poor gro-vrth, since
the concentration of CO2 in the air is
so very small.
The CO2 that is absorbed by the leaf is
transformed to sugars in the chloro-
plasts, which are small, more or less
rounded bodies to be found in the cells
of all green tissue. The chlorophyll of
the chloroplasts absorbs light (prin-
cipally the red and blue wave lengths,
leaving the familiar green color) and
uses the light energy to split off hydro-
gen from water. The hydrogen thus
gained is used to "reduce" carbon diox-
ide to a higher energy state — the state
in which it occurs in sugars and starch.
This reduction to a higher energy state
is an extremely ccmplicated biochemical
process, but it can be summarized as
follows :
1. The CO2 absorbed by the chloroplasts
is added by an enzyme to aphosphory-
lated 5-carbon sugar. This forms an
unstable 6-carbon compound, which
immediately splits to give two phos-
phorylated S-cs^^'bcn acids.
2. The hydrogen produced by the split-
ting of water is added to the 3-carbon
acids, which then condense to give a
stable 6-carbon sugar, a phosphory-
lated glucose. The phosphorylated
glucose is a key compound that can
be readily transformed to starch,
cane sugar, fat, amino acids, etc.
The figure on the back schematica3J.y
depicts the photosynthetic process.
Greenhouse and laboratory experiments
have shown that the concentration of CO2
in the atmosphere is far frcm optimal
for photosynthesis. By increasing the
CO2 concentration about 2.5 times, the
photosynthetic rate of plants in bright
sunlight can be doubled. There is some
question, however, about the effect of
still higher concentrations of COg.
While short-time photosynthesis con-
tinues to increase, the higher concentra-
tions of CO2 have a toxic effect that
reduces grovth.
Many investigators have grown crop plants
to maturityin C02-enriched air in green-
houses, and with few exceptions they re-
port yield increases of 20 to 200 percent.
Exceptions apparently occur when the CO2
concentration is raised too high. There
is no accurate evaluation of just what
concentration of CO2 would produce maxi-
mum photosynthesis without injuring the
plant, hut it appears that a 50-100 per-
cent increase in concentration would he
nearly optimal, depending on the species
concerned.
Field studies on the effect of CO2 en-
richment have also been carried out,
and for the most part these results also
shew yield increases. Field studies
are much more difficult to make than
greenhouse studies, "because the gas is
not contained and winds will carry it
away. However, CO2 is a heavy gas, and
if released near the soil it will tend to
"hang" in the vegetation in high concen-
tration if there is not excessive turbu-
lence due to winds. Even though marked
yield increases of such crops as sugar
beets have been obtained by CO2 enrich-
ment of the air, such "fertilization" is
not commercially practical because of the
excessive cost of supplying the gas.
Actually it appears that sizable yield
increases could be obtained if one
could only maintain the normal concen-
tration of CO2 around the crop during
the daylight hours. As it is, the CO2
concentration in a field of rapidly
growing corn during the daylight hours
will average about 25 percent less than
normal. Measurements made at Ames,
Iowa, show that even at 5OO feet above
the cornfield the concentration will be
reduced 10 percent. Such depletions
occur on windy as well as still days
light
-3-
and leave no doubt that the crop can ah-
sorh CO2 more rapidly than it can be
brought down from the upper atmosphere.
The upper atmosphere is not the only
source of CO2 for the plant ^ however.
The respiration of microorganisms and
plant roots produces large amounts of CO 2
in the soil^ which diffuses up into the
atmosphere about the leaves of the crop.
A fertile^ warm, moist soil, well sup-
plied with organic matter, will give off
as much as 3OO pounds of CO2 per acre
during the daylight hours. Inasmuch as
a very rapidly growing acre of corn will
absorb about 400 pounds of CO2 in the
same period, it can be seen that the soil
can do much to maintain favorable concen-
trations of CO2 about the leaves. If it
were not for this evolution of CO2 from
the soil, the concentration of the gas in
the cornfield during the day would un-
doubtedly be reduced more than the 25
percent below normal previously quoted.
The point bears emphasizing, however,
that, if the soil is to be effective in
this respect, it must be fertile, moist.
and warm, with a high organic matter con-
tent and a structure conducive to rapid
gas exchange. In short, production of CO2
by the soil will be maximal in those
soils that have long been recognized as
producing excellent crops. But just
what share of the yield is to be at-
tributed to increased COo concentration
about the leaves? Experimental evidence
does not permit this question to be
answered with exactitude, although an ex-
periment conducted in Sweden with sugar
beets 30 years ago suggests that it
might be appreciable.
In this experiment, manure, instead of
being worked into the soil, was allowed
to ferment in troughs between the beet
rows. This fermentation prevented the
roots from obtaining the nutritive ele-
ments, but the air around the leaves was
enriched by the escaping C02. A 19 per-
cent increase in yield over controls was
obtained by this treatment. Such results
suggest that an appreciable part of record
corn yields may be due to increased CO2
coming from the heavily manured soils
that seem to be an invariable part of
such yields.
J. B. Hanson
1-16-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
M-15
RADIATION AND PLANT BREEDING
In 1927 and 1928 Dr. H. J. Muller, now
of Indiana University, and the late Dr.
L. J. Stadler, of the University of
Missouri, reported that X-ray and ultra-
violet radiation "brought about heritable
changes (mutations) in living organisms.
(Doctor Muller was awarded the Eobel
prize in Medicine in 19^6 for this work. )
Since the rate of mutation was thousands
of times greater in irradiated material
than in untreated material and, further,
since some of the mutants appeared to
have qualities that plant breeders were
looking for, it was immediately suggested
that plant breeders use this new tech-
nique.
At that time corn breeders were perfect-
ing hybrid corn, and other breeders were
convinced that the new tool was not well
adapted to their programs. Most breeders
and geneticists were impressed with
the fact that the vast majority of the
mutations were deleterious, i.e., the
plant that possessed the new character
yielded lower or was otherwise less de-
sirable than already existing strains.
Hence the use of irradiation in breeding
was largely neglected.
If we examine critically the changes
brought about in the hereditary material
of an irradiated individual, we can make
some interesting observations. Muller
and Stadler found that these changes
could be roughly placed into three
classes:
1. Intra-genic changes, i.e., changes
within the gene itself that are simi-
larto those found in nature. These
changes behaved in a Mendelian fash-
ion. No differences in viability of
pollen or egg were evident and normal
segregation occurred in the F2. Un-
fortunately, these changes are the
least common ones that are induced
by radiation.
2. Deletion, i.e., destruction of a
small part of the chromosome itself.
Critical studies by Stadler revealed
that many of these changes were so
small as to be undetactable when he
looked at the chromosome under a
microscope, but genetic studies
showed that a number of genes had
been destroyed.
3. Structural rearrangement of chromo-
somes. Many of the mutations in-
volved the translocation of part of
one chromosome to another chromosome,
or the inversion, or change in gene
order, of a single chromosome. These
changes produced varying degrees of
sterility, either on the female or
male side, or on both.
Swedish plant breeders, however, started
irradiation breeding in barley and other
cereal crops and were able to show that
mutants could be produced that matured
earlier and had stiffer and shorter
straw, greater resistance to certain
diseases and, in a few cases, higher
yield. However, these useful mutants
were very rare. Swedish workers esti-
mate that only one in a thousand of
these mutations may be of value in a
breeding program.
An increase in radiation breeding oc-
curred after World War II in the U.S.,
largely because of the financial support
of the Atomic Energy Commission. Radia-
tion breeding in corn, wheat, oats, soy-
beans, rye, peanuts, cotton, and several
other field crop species is currently
under way in agricultural experiment
stations throughout this country. It is
still too early to anticipate the even-
tual outcome of all this work, but the
conclusions on the following page appear
to the justified as of now.
2.
It is improbable that the mutants
produced in a strain by irradiation
will be used directly by farmers.
It is more probable that the new
quality will be incorporated into
new varieties by conventional plant
breeding methods and that these va-
rieties will be released only if
they possess advantages over exist-
ing varieties.
The use of radiation does not appear
to be a profitable venture for plant
breeders unless a desired quality
does not exist or cannot be found in
the species under improvement.
3. No drastically new and completely
superior varieties can be expected
through irradiation treatment alone.
Since the discovery of the mutational
effect of ionizing radiation in I926,
millions of individual seeds,, plants,
and pollen grains have been treated
with X-rays, neutrons, and ultra-
violet rays. Some heritable changes
have been wrought. All of them are
more or less similar to the changes
that occur at rarer intervals without
man's intervention, and most of them
are deleterious to the individual
that inherits them.
D. E. Alexander
3-26-56
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
MISCELLANEOUS
AGRONOMY FACTS
M-16
THE YELLOW DWARF (RED LEAF) DISEASE OF OATS
The red leaf (yellov dvarf ) disease of
oats probably has been present in the
U.S. for many years. Excellent descrip-
tions of what appear to be the eame
disease were published as far back as
1898. Since 19^5:) however^ the disease
has become increasingly more prevalent.
The reason for this increase is not
known,, but it does not appear to be as-
sociated with the continual change-over
in oat varieties.
Red leaf is caused by a virus that can
be transmitted by at least five species
of aphids commonly infesting small grain
and grasses. The virus cannot be trans-
mitted mechanically, in the seed, or
through the soil.
The virus has been sho-vm to be identical
with the one causing yellow dwarf dis-
ease of small gi-ain in California. The
name "yellow dwarf" was selected by Cali-
fornia workers on the basis of outstand-
ing symptoms produced by the virus on
barley and wheat.
The disease usually appears first on the
edges of a field and in "spots" or cir-
cular areas varying from a few feet to
30 or more feet in diameter. Sometimes
these areas may overlap, and in years
when aphids are very abimdant a field
may be uniformly affected. Farmers fre-
quently have failed to associate earlier
aphid infestations with the disease,
since the first symptoms usually appear
in about ik days. Sometimes nearly all
aphids have disappeared by the time the
first symptoms can be seen.
The first symptom of the disease in oats
is the appearance, usually near the leaf
tip, of faint yellowish-green blotches
that can best be seen by holding the
leaf blade up to the light. V/hen first
formed, the blotches are somewhat vari-
able in size and shape and are usually
less than a few centimeters in size. The
blotches enlarge rather rapidly, merge,
and turn various shades of red, broim,
and yellow- orange. Cool temperatures
(70° F. and lower) favor the appearance
of red pigments in the affected leaves,
whereas temperatures of 75° F. and above
suppress their appearance. At the same
time, the yellowish-green blotches con-
tinue to appear on successively lower
portions of the leaf in advance of the
changes in color. The affected portions
of the leaf often die rapidly.
In addition, a rather characteristic in-
ward curling of affected leaves frequently
occurs. Symptoms generally appear on
the oldest leaves first and then succes-
sively involve the younger leaves. Oc-
casionally the youngest leaves will show
a longitudinal striping resulting from a
yellow-green color in the interveinal
areas and a darker green in the tissue
over the veins. The root system is as
severely stunted as the tops of the
plants.
Blasting of the florets is the most seri-
ous aspect of the disease. It may vary
from only a few blasted florets to com-
plete failure of the plant to head. A
shriveling and lower test weight of ker-
nels also may occur. The severity of
the disease depends on the variety of
oat infected, the age of the plants at
the time they become infected^ and the
strain of virus involved.
Winged aphids moving into small grain
fields in the spring from various grasses
are believed to be responsible for the
initial spread. Each adult aphid is cap-
able of producing daily from 10 to 20
young, which upon maturity (one to two
weeks) also begin to produce. These
later aphids are usually vingless and
move about by crawling from plant to
plant. When food conditions become un-
favorable, winged forms develop which
fly to other fields of small grain and
grasses.
In order to transmit the virus to healthy
plants, the aphid first must acquire it
by feeding on diseased plants. However,
once the aphid has acquired the virus, it
apparently is able to transmit it for
the rest of its life.
Virus -free aphids --less than a hundred
per plant --cause very little damage to
small grain. The greenbug is an excep-
tion, since it secretes a toxin. A sin-
gle viruliferous apple -grain aphid is
able to transmit the virus to a healthy
oat plant in as short a time as four
hours.
For an aphid-transmitted virus to cause
serious loss in an annual crop raised
from seed, the virus must spread rapidly.
Rapid spreading can occur only if the
crop is easily infected, the source of
the virus is readily available, and the
vector is very numerous and active. For-
tunately, all of these conditions do not
usually occur at the same time, and losses
therefore vary greatly from year to year
and from locality to locality.
No oat variety tested has been found to
possess satisfactory resistance to red
leaf disease. Therefore, an intensive
effort is being made to locate sources
of resistance in oats.
Early planting is the only practice that
can be recommended at present. This rec-
ommendation is based on the fact that
large plants are better able to "toler-
ate" the disease than smaller ones.
R. M. Takeshita
5-li^-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
C-7
CORN HYBRIDS FOR SPECIALIZED FAR/V^ AND MARKET USES
Corn is an extremely versatile crop.
Consequently, breeders are able to se-
lect types for particular uses. They
may work with dent, sweet, pop, flint,
floury, or waxy corn. These types, how-
ever, may be modified greatly by breed-
ing and selection.
Livestock feed. From 75 to 90 percent
of the corn crop in the United States is
fed to livestock. In contrast, most of
the corn crop in many other areas of the
world is used for human food. Most live-
stock feeders in the United States pre-
fer a dent grain that is yellow, soft,
and high in quality and quantity of pro-
tein. Feeders in many other covintries
prefer a white flint corn.
Corn has certain limitations for feed-
ing. It is low in quality and quantity
of protein and is relatively low in
vitamins. Seme feeders also ccmplain of
poor palatability and reduced gains from
certain hybrids. Many feeders grind the
harder types.
Corn breeders need to develop special
hybrids with high tonnage and better
quality of silage and green feed.
Multiple -eared, heavy- tillering strains
may be useful for this purpose. Consid-
eration must be given to yield and per-
centage of dry matter, proportion of
ears to stalks and leaves, and percent-
age 1 of crude fiber and protein. These
studies should be supported by digesti-
bility experiments and by feeding trials.
Milling industry. Wet millers desire
strains that are high in starch, oil,
and protein. Since corn contains about
70 percent starch (dry basis), a varia-
tion of one or two percent is important
in large-scale operation. The industry
claims that flinty types do not steep
and process so well as softer types.
Most dry millers prefer a kernel that is
semihard and vitreous and that does not
have too much soft starch on either the
tip or the dent end. They want neither
the shoe -peg type nor small, roiond
kernels. The dent must not be too deep
or the hull too rough. Kernels of this
type make chaffy or chalky products
which are fit only to put into feed.
The cob color is of little consequence.
An adequate supply of white corn is
another requirement of dry millers. Con-
s\jmers claim a distinct preference for
the flavor and taste of meal made from
white corn. The finished-product demand
is for either pure white or pure yellow.
Protein content and quality. Protein is
an expensive but necessary constituent
of food and feed. The University of
Illinois gave the first evidence that
protein and oil content in corn could be
greatly increased or lowered by breed-
ing. After 50 generations of selection,
the average protein content was 19.5
percent for Illinois High Protein and
4.9 percent for Illinois Lev Protein.
These open-pollinated strains yield only
about 50 percent as much grain as adapted
hybrids. Fortunately, the high pro-
tein trait can be transferred to stand-
ard inbred lines by breeding procedvires.
Quality of protein is fully as inrportant
as quantity of protein. The corn kernel
contains two main types of protein.
That found in the endosperm is primarily
zein. Zein is deficient in tryptophane
and lysine, which are essential for ani-
mal nutrition. The other type of pro-
tein, fo\ind in both endosperm and germ,
contains both tryptophane and lysine and
is biologically balanced. The corn
breeder would like to increase the per-
centage of these amino acids in the en-
dosperm protein. The alternative is to
add them to the diet from other sources.
Oil for industry and high-energy feed.
Corn oil, a valuable by-product of the
starch industry, is high in energy value
for livestock feeding. Most of ■'the oil
is in the germ of the kernel. Germ pro-
tein contains tryptophane and lysine,
is biologically balanced, and is prob-
ably more valuable for livestock feeding
than endosperm protein. High- oil hy-
brids having a high proportion of germ
to endosperm should therefore benefit
both the starch industry and livestock
feeders.
After 50 generations of selection at the
University of Illinois, the average oil
content of Illinois High Oil vas 15.^
percent compared with 1.0 percent for
Illinois Lov Oil. Unfortunately, these
open-pollinated high-oil strains are low
yielding. High oil, however, was trans-
ferred to standard inbred lines at the
Illinois ' station by crossing, followed
by back-crossing, selection, and self-
fertilization. Selection for high oil
was accomplished by selecting ears bear-
ing kernels with large germs.
Breeding programs have been inaugurated
by several Com Belt agricultural exper-
iment stations and private hybrid seed
corn companies to develop hybrids with
high -oil or high -protein content. Seme
of these hybrids appear to be very prom-
ising. For example. 111. 6063 produced
lij-,0 percent more protein, 32.8 percent
more oil, and was 8.5 percent superior
in grain yield to U. S. I3, a standard
hybrid. However, these Illinois experi-
mental combinations are not yet in com-
mercial use.
Zein for special fabrics. Protein con-
tent of corn grain may be increased by
breeding and by high applications of
nitrogen fertilizers. Most of the in-
crease of endosperm protein is zein,
whith is not high in nutritive value be-
cause it is poorly balanced among its
constituent amino acids.
Zein is obtained from the gluten in the
corn wet-milling process. The Northern
Utilization Research Branch of the U. S.
Department of Agriculture, Peoria, Illi-
nois, studied dispersion of zein in
strong alkali, "spinning" it into a fi-
ber- and stretching and curing to give
the fiber added strength.
This fiber, which is available on the
market under the trade name "Vicarai'! is
used mostly in blends with wool for such
garments as socks, swimming suits, and
sweaters. Possible increased use of
zein for special fabrics has created
seme interest in breeding comfbr-higlier
zein content,
Amylose for plastics, cellophane, and
films, Amylose is a linear-type molecule
which can be made into thin, transparent
films resembling cellophane. Acceptable
films require amylose of 80 percent
purity. Ordinary com starch contains
about 27 percent amylose. Samples, how-
ever, have been found in which the starch
was 62 percent amylose. Consequently,
it appears that it may be possible
eventually to obtain dent corn with a
high enough amylose content for the prac-
tical production of plastics, cellophane,
and films.
Vitamin A (B-darotene), An association
has been foijnd between the yellow pig-
ment and vitamin A in corn. There is a
direct quantitative relation between
vltemln A and the number of genes for
yellow pigment in the endosperm. In
fact, the yellow endosperm genes act in
an arithmetic, cimiulative manner, each
gene adding 2,5 units of vitamin A per
gram of grain. Feeding tests have dem-
onstrated that yellow corn is better
than white corn for hogs on drylot feed-
ing.
Niacin (nicotinic acid). Fortunately,
wide differences in niacin concentra-
tions have been found among various
strains of corn. Hybrids tend to rank
between their parents, and the seed par-
ent generally exercises more influence
than the pollen parent. Dent kernels
are lowest in niacin content, waxy ker-
nels intermediate, and sugary " kernels
highest. Adequate niacin in the diet
eliminates certain malnutritional dis-
turbances.
R. W, Jugenheimer
9-5-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
C-8
TOMORROW'S HYBRID CORN
It is becoming more and more difficult
to develop hybrids that are definitely-
superior in all characteristics to the
better ones nov available in the Corn
Belt. For this reason, the Corn Belt
hybrids of I965 may not be radically
different from those of today. However,
it should be possible to greatly im-
prove the hybrids that are adapted to
other areas of the United States and many
other sections of the world.
Hybrids for special purposes and uses
should soon be available. Producers and
industry \d.ll be able to choose between
better dent, flint, sweet, pop, waxy or
possibly floury types.
Livestock feeders in the United States
prefer a yellow grain that is soft and
high in quality and quantity of protein.
Multiple -eared, heavy -tillering strains
will be useful for silage and green
feed.
Industry will eventually be able to ob-
tain more suitable types of corn. Dry
millers prefer white kernels with smooth
dent. Wet millers and livestock feeders
will welcome high- oil hybrids. Waxy
corn is available for food and glue.
High-zein corn can be used for special
fabrics, and a high-amylose corn would hybrids are better husk cover, better
be valuable
and films.
Hybrids differ greatly in drouth re-
sistance. The leaves of some strains
remain green, while others are badly in-
jured by heat. Some hybrids set seed
satisfactorily under conditions of high
temperature and low humidity, while
others shed little pollen for only short
periods.
Excellent standability has contributed
greatly to the popvilarity of hybrids in
the Corn Belt, This desirable trait
needs more emphasis in. many other places.
Lodging lowers quality and yield and
makes harvesting more difficult. Varia-
tions in standability between hybrids
are caused by differences in stalk struc-
ture, root system, ear height, soil
fertility, plant population, and resist-
ance to insects and diseases.
For hand harvesting, farmers want single-
eared strains with the ear borne at a
convenient height. It may be possible to
harvest future hybrids with a mechanical
picker more easily and satisfactorily
than the present types can be harvested.
Hybrids with shorter plants may also be
better adapted for field shelling and
ccm.bine harvesting.
Other traits that may be added to future
for plastics, cellophane, grain quality, higher shelling percent-
age, and resistance to chemical weed
Yields of grain, silage, and fodder will
gradually edge upward because hybrids
will be better able to resist hazards.
Effective disease inoculation and insect
infestation techniques will result in
hybrids that have greater resistance to
diseases and insects. In general, flint
corn germinates better and the seedings
grew more vigorously than dent in the
cooler climates.
sprays ,
By 1965 the use of male sterility and
pollen restorers will probably eliminate
much of the detasseling now required in
producing hybrid seed. This development
should lower production costs and result
in a better product,
R, ¥, Jugenheimer
10-10-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
TEMPERATURE AND OTHER INTERRELATED FACTORS IN
DROUGHT DAMAGE TO THE CORN PLANT
C-9
The question, "Is it high temperature or
lack of moisture that is damaging the
corn plant?" commonly asked in a hot, dry
year is exceedingly hard to answer. The
reason for this is that the two variahles
are so intimately interrelated that it
is impossihle to separate them under
normal field conditions.
Some of the factors involved in "drought
damage" of the corn plant under field
conditions in addition to temperature
are listed below and show the complexity
of the problem.
1. Metabolic status of the plant
2. Soil moisture
3. Atmospheric moisture or relative
humidity
h. Physiological moisture
5. Variety of the plant
6. State of development of the plant
7. Part of the plant
Since under field conditions all of
these factors may be in operation at the
same time, it is extremely hard to dis-
cuss the problem in general. This arti-
cle will therefore emphasize temperature
and try to relate the other factors to it .
The temperature factor. The following
method is commonly used to determine the
temperature that is required to kill
plant tissue and cells: A leaf, branch,
or section of tissue is cut off and im-
mersed in water maintained at a constant
temperattire . The material is removed
after an appropriate time interval, and
staining techniques are used to deter-
mine whether the cells have been killed.
By varying the temperature and the time
of Immersion, and determining the per-
centage of cells killed, it is possible
to determine the "thermal death point."
This term which is used to express the
results is based on three components:
tim£, temperature, and percentage of
cells killed.
The main advantage of this method is
that it separates temperature effects
from many of the environmental factors.
As an illustration, it was found that
100 percent of the cells of range grass
roots were killed when immersed in water
at 162 F. In contrast where the roots
were placed in water which was slowly
warmed over a 150-minute period, the
lethal temperature was 126 F. This
method, or modifications of it, has been
used by many investigators to establish
a critical temperature range that varies
from 113 to l40 F. for many different
plant species. The literature shows
that the corn plant falls within this
temperature range.
In visualising hew high temperatures
kill a plant cell, it should be kept in
mind that all the vital metabolic pro-
cesses are carried out by proteinaceous
compounds called enzymes. Heat, or de-
hydration by heat, can inactivate the en-
zymes by coagulation in much the same
mamier as egg protein is coagulated by
frying or boiling.
Leaves in general, and corn leaves are
no exception, tend to maintain them-
selves at the same temperature as the
air surrounding them. Using over a thou-
sand separate measurementF, experimenters
at the Kansas Experiment Station found
the average temperature of tiirgid leaves
to be 87.2 F. compared to an average of
87 F. for air temperature. Wilted corn
leaves measured from 3 to 8 F. higher.
This indicates that transpiration does
not have a major role in regulating leaf
temperature. The leaf then must maintain
its temperature by a) reflecting a por-
tion of the light and b) radiating heat.
The latter is accomplished in mu'h the
same way as the "leaf -like" finned por-
tion of an air-cooled motor dissipates
heat.
Temperature and metabolism. Plants have
two major metabolic systems that are
"geared together," a) photosynthesis
which synthesizes the carbohydrates
vhich are \ised for "building blocks" and
for energy and b) respiration which oxi-
dizes (burns ) carbohydrates to provide
energy for maintenance and growth^ and
to supply "building blocks" of different
types needed in plant growth . At a tem-
perature of 68 F. the photosynthetic
process exceeds the respiratory process
by a considerable margin. This supplies
an adequate amoiont of carbohydrates for
the respiratory process and also pro-
vides the "building blocks" needed for
grovth. As temperatui-e is increased, a
compensation point is reached where in-
put of carbohydrates just balances the
consumption by respiration, while at
higher temperatures (96 F. ) the respir-
atory utilization exceeds that provided
by the photosynthetic process. Prolonged
exposiu:e of the plant to high temper-
atures would therefore markedly curtail
plant growth.
It has been observed that a wilted leaf
absorbs but one-third the amount of CO2
taken up by a turgid leaf. Naturally,
this would reduce the photosynthetic pro-
duction of carbohydrates by the same
amount. This emphasizes the interrela-
tion of moisture and temperature.
Although the information is meager,
there seems to be seme relationship be-
tween the levels of carbohydrates, pro-
teins, and colloids in the plant cells
and their resistance to dehydration. It
has been pointed out that dehydration of
the proteinaceous material of cells can
result in their death. These levels of
cellular constituents are controlled by
the metabolism of the plant.
Temperature and moisture. Three major
factors involved in maintaining an ade-
quate physiological moisture level in
the plant are: soil moisture, water-
conducting system of the plant, and rate
of transpiration of water loss from the
plant. Soil moisture is the major reserve
of water supply and must be adequate to
meet the demands made by the plant for
maintaining its physiological moisture
level and transpirational losses. Since
water is conducted frcm the soil by the
xylem system of the plant, this "piping
system" must develop rapidly enough and
be large enough to supply the demands of
all parts of the plant. It has long been
known that certain varieties of corn are
more susceptible to top leaf blasting
than others. Recently it has been shcwnl/
that the susceptible varieties were much
slower in developing xylem vessels in the
leaves than the non-blasting varieties.
Consequently, in high transpirational
periods (hot, dry windy weather) the
leaves of the plant with inadequately
developed "piping systems" were severely
desiccated and subsequently died. This
desiccation could occur even with ade-
quate soil moisture since the failure is
in the water trajisport system.
The rate of transpiration from leaves
depends largely on temperature, relative
humidity, and air movement (wind cur-
rents). The relationship between tem-
perat-ure and rela.tive humidity is siio-rfn
by the fcllov^ing: air at 50 , 68 , and
100 F. must contain, respectively; 0.3^
0.6, and 3»5 oimces of water per cubic
yard to achieve 100 percent relative hu-
midity at each temperature. Air at
50 F. and a relative humidity of 80 per-
cent if heated to 68 F. would drop to
40 percent relative hvunidity and at
100 F, would be only 0.6 percent satu-
rated. The loss of water from leaves is
controlled by the gradient between the
relative humidity of the stcmata of the
leaves (assuming 100 percent R.H.) and
that of the air. Rapid air movement
tends to keep this gradient at amaximum.
Temperature and part of the plant. Some
parts of the plant are more susceptible
to heat and desiccation damage than
other parts. For example, pollen and
silk seem to be most sensitive. In
field trials2/ representing some 7^000
pollinations, a good correlation was ob-
tained between high temperatures and
failure to set seed. At 75 F« 'the per-
centage of ovules setting seed was 65,
while at IO5-IIO F, only eight percent
seed set was obtained. Desiccation of
pollen and silks rather than lethal tem-
perature effects was considered the pri-
mary cause of the damage.
1/ Private communication from Dr. L. A.
Tatum, Kansas State College, Manhattan,
Kansas.
2/ lonnquist, J. A. and Jugenheimer, R.W.
Jour. Amer. Soc. Agron. 35:923. 19^3*
R. H. Hageman
2-6-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
C-10
HOW TO ESTIMATE HAIL LOSSES TO CORN
Hail insurance on growing crops in this
country increased gradually from 87
million dollars in 193^ "bo over 1 3/^
"billion in 1953^ or 20 fold. Payments
to farmers for losses ranged from a low
of 1.5 percent of the insured value to
a high of 3-1 percent, with an average
of 2.3 percent over the 20-year period.
The amount of damage inflicted by hail
is hard to estimate. The stage of the
crop when the storm strikes and the se-
verity of the injury are the two main
factors that need to be considered in
appraising losses. Without data from
experiments, an estimate of the loss re-
sulting from a hailstorm might be no
better than a wild guess. Fortunately,
field trials have been conducted by
agricultural experiment stations in
Illinois (6 years), Iowa (7 years),
Nebraska (9 years). South Dakota (2
years), and West Virginia (l year), in
addition to some extensive tests by in-
surance companies. The results of these
experiments agree closely and make it
possible to assess the damaging effect
of a hailstorm on a crop of corn rather
accurately.
this stage, the plant will therefore
produce no grain. Removing all exposed
blades when the plants are younger does
little harm because defoliation at that
time takes off only a little leaf surface
and the plant produces new leaves as the
stem pushes upwards inside the whorl.
The later the blades are removed, the
greater the percentage of leaf surface
destroyed. Thus, grain yield goes down
as blade removal is delayed, and this
continues until the tasseling stage.
After tasseling, however, grain yield
goes up as blade removal is delayed.
This relation between grain yield and
stage of plant development at the time
blades were removed was borne out in t"ne
experiments in all the states. So if a
hailstorm occurs, carefully note the
stage of development your crop is in when
the hail strikes it.
Degree of injury is also important in es-
timating damage from hail. Any injury
to the corn plant will usually decrease
grain yield, because Nature does not pro-
vide the corn plant with enough leaf sur-
face to permit part of it to be sacrificed
without affecting the yield.
Sometimes farmers are not fully satis-
fied with the appraisal of injury to
their hail-damaged crops. They ask how
the losses are estimated. This brief
discussion is presented to explain the
factors that need to be considered in
arriving at an estim.ate.
Stage of crop development has an im-
portant bearing on the losses from hail
injury. Corn plants in the tassel and
ear-shoot-emerging stage are most sub-
ject to injury so far as grain pro-
duction is concerned. No grain has been
produced before the injury, and no new
blades can be produced afterwards. If
all the blades are removed by hail at
Experimental results show that grain
yields are reduced in direct proportion
to the amount of leaf area that is re-
moved. There is a tendency, however, es-
pecially when only small percentages of
leaves are removed, for the yield reduc-
tion to be somewhat less than the amoimt
of leaf surface that is lost. This sug-
gests that the efficiency of the uninjured
leaf surface is stepped up after some of
the leaves have been removed, possibly
because the remaining leaves get more
light.
As soon as possible after the storm sub-
sides, get as careful an estimate as you
can of the amount of blade surface re-
moved from the plants by the hailstorm.
Blade shredding, midrib breaking, and
stalk and ear bruising are other forms
of injury caused by hail. Tests show that
as long as any part of the blade rem.ains
attached to the plant it is capable of
contributing to grain yield. In lowa^
when all the blades were severely shredded,
yield of grain was 37 percent of normal
even when the shredding was done at the
beginning of tasseling. Severe shredding
earlier and later caused progressively
less damage.
Midrib breaking did not do much harm.
With every midrib broken at the most
critical time, namely, tasseling time,
yield was 80 percent of normal.
Stalk bruising decreased yields about 10
percent beyond that caused by blade
shredding. Ear bruising did little harm
to yield, but when it occurred at the
milk stage the market quality of the
grain was reduced somewhat because of
the damage to kernels .
Believe it or not, under some conditions
hail injury may actually increase grain
yields. This happened in Iowa during
the dry year of 1930- Cutting out blades
reduced transpiration, and the moisture
thus conserved was more beneficial to the
plant than the leaf removal was harmful.
Experimental data have taken much of the
"guess" out of estimating losses to corn
from hail injury. Yet it is still neces-
sary to weigh the significance of the
many factors that have a bearing on the
outc ome .
George H. Dungan
5-28-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
IDENTIFYING COMMON LEGUME SEEDLINGS
F-17
It is often necessary to identify cer-
tain legumes before they have flowered
or after they have been closely grazed.
Many times it is necessary to identify
Cot yle denary leaves - the seed leaves of
the embryo -which act as storage organs
in seeds of plants.
Leaflet - one of the divisions of a com-
pound leaf; e.g., the red clover leaf
has three leaflets.
Petiole - the stalk of a leaf.
Pubescent - having fine, soft hairs.
In the following key to identifying
legume seedlings, the cotyledonary leaves
are considered to be the first and second
leaves. Most of the common legvmes
exhibit epigean emergence; i.e., the
cotyledons emerge aboveground. The pea
and vetch, notable exceptions, exhibit
hypogean emergence; i.e., the cotyledons
remain imderground during germination
and emergence.
leg\minous plants in the seedling stage.
The follo-vrLng definitions and distin-
guishing featiires should be helpful in
identifying some ccmnon legume seedlings.
Serrate - having sharp teeth.
Trifoliolate - having three leaflets.
Unifoliolate - having one leaflet.
Variegation - the barring (water marks)
on leaves, seen in nearly all American
strains of red clover.
Vein (nerve, rib) - nerve or rib in
leaves, bracts, scales, sepals, etc.
This key is greatly simplified. It
should be remembered that many weed seed-
lings have characteristics similar to
those of legume seedlings. Further,
there are many variations within the
different species presented in the key;
e.g., most European strains of red clover
are not pubescent. However, for practi-
cal field use, the key will help to
identify some common legume species in
the seedling stage.
Key to the Seedlings of Some Common Legume Species
-la. Third leaf imifoliolate (fourth leaf trif oliolate )
_ 2a, Petiolar branches of vmequal length
3a • Leaflets one -third serrate - alfalfa
yo. Leaflets completely serrate - sveet clover
-213. Petiolar "branches of equal length
_3a. Vegetative parts pubescent
Ua. Variegation present - red clover
4b, No variegation present - crimson clover
3b. Vegetative parts not pubescent
ka. No variegation present - alsike clover
I hh. Variegation present
. 5a- • Giant form - Ladino clover
I 5^3, Snail form - common vhite clover
lb. Third leaf trif oliolate or both third and fourth leaves
unif oliolate
_2a. Third leaf trif oliolate, veins not prominent - birdsfoot
trefoil
_2b. Both third and fovirth leaves unif oliolate, veins
prominent - common, Korean, and sericean lespedeza
A, W, Burger
10-31-55
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SELECTING ALFALFA VARIETIES
F-18
The main point to consider in selecting
an alfalfa variety is the time you ex-
pect the alfalfa to stand before plowing
it down. If you plan to use the stand
several years for hay^ plant seed of a
winter-hardjj wilt-resistant variety like
Ranger or Buffalo. If you plan to use
it only one or two years for hay^ you
can use a winter-hardy^ wilt-susceptible
variety like Atlantic or Du Puits. In
either case, use certified seed.
Bacterial wilt does not reduce alfalfa
yield until about the third year. Be-
cause certain wilt -susceptible vari-
eties, such as Atlantic and Du Puits, are
as productive as Ranger and Buffalo dur-
ing the first year or two, there is no
advantage in usinga wilt-resistant vari-
ety in short rotations.
Several varieties of alfalfa have been
developed in the United States. There
is a good seed supply of most varieties.
Following are descriptions of several
varieties and status of seed supplies.
Ranger, which is resistant to bacterial
wilt, was developed at the Nebraska Ex-
periment Station by intercrossing se-
lected strains of Cossack, Ladak, and
Turkistan. Ranger is a good forage pro-
ducer and is as winter-hardy as the
hardy common alfalfas. The flower color
is variegated. Ranger is recommended
for the northern two-thirds of Illinois.
Seed supply is adequate.
Buffalo, also resistant to bacterial
wilt, was developed by the Kansas Experi-
ment Station out of Kansas Common. It
is a good forage producer and is only
slightly less winter-hardy than Ranger.
Flower color is purple. Buffalo is rec-
ommended in the southern two-thirds of
Illinois. Seed supply is adequate.
Atlantic, a high-yielding variety devel-
oped by the New Jersey Experiment Sta-
tion, is not resistant to bacterial
wilt. It was developed especially for
the eastern states, where bacterial wilt
is not serious. It is about as winter-
hardy as Buffalo. Flower color is varie-
gated. Atlantic is recommended throughout
Illinois for short rotations. Seed sup-
ply is adequate.
Du Puits is a variety developed in France
that has yielded exceptionally well in
tests in Illinois and several other
states. Lu Puits is not wilt-resistant.
It is about as winter-hardy as Buffalo.
Du Puits is recommended for the southern
two-thirds of Illinois in short rota-
tions. Seed supply is limited.
Vernal is a variegated variety developed
at the Wisconsin E^cperiment Station. It
is very cold-resistant and highly resist-
ant to bacterial wilt. It is not so
susceptible to leaf and stem diseases as
Ranger. This variety has not been
tested in Illinois long enough to deter-
mine its value in relation to the recom-
mended varieties. It is not reccmmended
in Illinois at the present time. Seed
supply is limited.
Narragansett, a high-yielding variety
developed by the Rhode Island Experiment
Station, is not resistant to bacterial
wilt. It was developed for use in the
eastern United States north of the area
where Atlantic is adapted. Flower color
is variegated. Atlantic is preferred to
Narragansett in Illinois at the present
time. Seed supply is limited, but it
should be adequate to meet the demand.
Nomad has a high proportion of creeping
plants that will root at stem nodes. It
is from an old field in Oregon found to
have this type of plant. Nomad is sus-
ceptible to bacterial wilt, and it has
not been tested long enough to determine
its adaptability. In most tests it has
not appeared to be so vigorous as other
varieties. Because of its creeping habit
of growth, it may be useful in pastures.
It is not recommended in Illinois at the
present time. A limited amount of seed
is available ccmmercially.
Rhizcma is a broad-crowned type of al-
falfa developed at the British Columbia
Experiment Sta.tion. It does not root at
the nodes and thus is not a true creeping
alfalfa. Rhizoma is a variegated^ very
winter -hardy variety that is not resist-
ant to bacterial wilt. It beccmes dor-
mant very early in the fall and begins
growth very late in the spring. For this
reason it is not recommended in Illinois.
Certified seed is available for all the
varieties recommended for use in Illi-
nois^ and it should be used in preference
to uncertified seed.
Certified seed may be produced outside
the region of adaptation, principally in
California. For seed to be certified
under such conditions, the seed fields
must be established from seed produced
in the region of adaptation.
Seed fields can remain down only six
years; therefore certified seed of
winter-hardy varieties that is produced
in California is only one generation re-
moved from plants that grew in the re-
gion of adaptation. Also, in fields
growing certified seed, precautions must
be taken to prevent the growth of volun-
teer seedlings. Winter-hardiness studies
have shown that, when these precautions
are taken, there is only slight loss of
winter-hardiness. It is only when these
varieties are grown for two or more
generations outside the region of adap-
tation that there is serious loss of
winter-hardiness .
J. A. Jackobs
12-26-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
F-19
HOW HYBRID SORGHUMS WERE DEVELOPED AND ARE BEING PRODUCED
Varieties of grain sorghums like Martin,
Midland, Westland, RedlDine 60, and Com-
"bine Kafir 60 and varieties of forage
sorghums like Atlas, Leoti Red, and Kan-
sas Orange are pure lines- -just as are
varieties of soybeans or oats. Hybrids
of sorghums are first-generation single
or three-way crosses between certain of
these pure lines.
It has been known for a long time that.
Just as in corn, hybrid vigor will result
from crossing certain varieties of sor-
ghum. However, in the past, controlling
pollination has been a major problem.
Since the anthers, or the pollen- shedding
organs, of the sorghums are located in
the same floral envelope as the female
parts, it is impossible to use the prin-
ciple of detasseling to control pollina-
tion, as is done with corn.
Work to devise methods for producing hy-
brid sorghum seed in quantity has been
under way at the Texas Agricultural Ex-
periment Station, in cooperation with
the United. States Department of Agricul-
ture, for more than 20 years
In 1929
J. C. Stephens of the Texas Station dis-
covered an antherless character in Sudan-
grass, and in 1935 he discovered a
better male -sterile in Texas Blackhull
Kafir. In 19^13 Glen H Kuykendall dis-
covered a still better male-sterile in a
field of the Day Milo variety on his
father's farm in Cookeville, Tennessee,
and in I95O J. C. Stephens and H. F.
Holland foixnd cytoplasmic male-sterility
in progeny of crosses between Milo and
Kafir. The cytoplasmic type of male-
sterility is utilized in single crosses
and has advantages over the tln-ee-way
cross which is used with the Day l^'ge of
male -sterility.
Sorghum plants with male -sterility do
not shed pollen, and the flowers are
readily fertilized by pollen carried by
the wind from nonnal plants . With this
type of sterility, it is a simple proc-
ess to produce and maintain both sterile
and fertilized plants in crossing fields
for producing hybrid seed at a reason-
able cost. To make it clear how hybrid
seed is and will be produced, it might
be well to follow the method of produc-
tion tlirough all the steps from mainte-
nance of parental stocks to production
in farmers' fields. The necessary steps
for maintaining stock and producing hy-
brid seed are shown in the following
diagrams, originally prepared by J. C.
Stephens .
Diagrams Showing the Method of Maintaining Seed of a Cytoplasmic Male-Sx.erile
Seed Parent and of Producing Hybrid Sorghum Seed
I. The maintenance and increase of a
male-sterile line is illustrated.
Male -sterile strain A and normal
strain A are identical except
that the male -sterile strain does
not have anthers that shed pol-
len. The chromosome complement
of both strains is the same, since
the male-sterile strain is main-
tained by backcrossing to the
normal A strain.
I MAINTENANCE AND INCREASE OF
MALE-STERILE STRAIN
MALE-STERILE A
COUNTERPART
NORMAL STRAIN A
MALE-STERILE
PRODUCING
/^
WIND-BLOWN
/r\
^
^
POLLEN
X
ISOLATED FIELD
II. Hybrid sorghum seed will be pro-
duced in a second crossing block
in which the seed parent rows
will be male-sterile strain A
ard the pollen parent will be
normal strain B. Lines chosen
as male parents must restore
normal fertility in the succeed-
ing crop, give a good hybrid of
combine height, and be good pol-
len producers. The hybrid seed
from the male-sterile female
rows is harvested and used for
commercial production.
I[ SEED-GROWER CROSSING BLOCK
MALE-STERILE
STRAIN A
\
NORMAL STRAIN B
MALE-FERTILE
RESTORING
WIND-BLOWN
POLLEN
X
ISOLATED FIELD
TO PRODUCE
III. The seed produced in the second
crossing block will be planted
by sorghum growers for commer-
cial production. This is the
single-cross A x B sorghum. All
plants are completely male and
female fertile. With these hy-
brids it is necessary to pur-
chase new seed each year, just
as is true of hybrid corn.
m SINGLE CROSS (A X B) HYBRID SORGHUM
NORMAL CROP ON FARM
At present work in the main
sorghum states is devoted to
crossing many lines and testing
to find the most suitable hy-
brid combinations. It will prob-
ably be several years before seed
stocks can be increased, the
various hybrids can be tested for
adaptability, and seed of "proved"
hybrids is made available for
commercial production to any ex-
tent.
It should be emphasized that not all sor-
ghum hybrids can be expected to be supe-
rior, but yield increases of 30 to 50
percent above those of present commercial
varieties have been reported in states
where these hybrids have been evaluated.
Testing is needed and is being carried
out in Illinois to determine the per-
formance of hybrid sorghums at various
locations in the state.
C. N. Hittle
i+-23-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SUDANGRASS IN ILLINOIS
Sudangrass is ideal for use as a summer
supplemental pasture in Illinois. It
may also be harvested for hay or silage.
This forage grass, which is a sorghum
and an annual, is very leafy, tillers
profusely, and has great capacity for
regrowth after cutting or grazing. It
becomes somewhat dormant under condi-
tions of severe drought, but resumes
growth when rains come during mid- or
late summer.
Sudangrass for Pasture
Sudangrass should not be pastured until
the crop is l8 to 2h inches tall. At
this stage the plants will contain
roughly from 10 to 13 percent protein,
and both common and sweet types will be
readily eaten. The possibility of poi-
soning from hydrocyanic (prussic) acid
is increased if Sudan is pastured at
shorter heights. The grazing management
should consist of rotational grazing
with heavy stocking so that the growth
may be grazed down rather quickly. The
crop should then be allowed to grow
until there is time for l8 to 2^+ inches
of regrowth before it is grazed again.
The place of Sudan in a balanced pasture
program is for late summer grazing in
conjunction with cool-season perennial
pasture crops for spring and fall usage.
When Sudangrass is to be used for pas-
ture, it may be planted with soybeans.
This combination furnishes considerably
more roughage than either crop alone and
is ready for use in midsummer when other
pasturage is short. Soybeans also pro-
vide added protein to the forage and,
since they are not injured by chinch
bugs, they aid in reducing the chinch
bug damage on the Sudangrass.
Other Uses
Although Sudangrass is difficult to cure,
it may be harvested for hay. The best
quality hay is obtained if cut and prop-
erly cured when the very first heads
begin to appear. Yields of up to three
tons of good-quality hay may be expected
from the finer-stemmed varieties. The
coarseness of some of the new varieties
makes them unsuitable for hay.
Sudan may also be ensiled and, when
properly stored, makes good-quality si-
lage. The yield for silage would be the
best when the crop is heading to soft
dough in stage. For use as hay or si-
lage, it should be harvested before leaf
drying occurs.
Frussic Acid Poisoning
Sudangrass contains a glucoside called
dhurrin, which releases hydrocyanic acid
when hydrolyzed in the ruminants. This
may cause HOW (or prussic acid) poison-
ing. Quantities of prussic acid large
enough to cause sickness are usually
fatal, and a poisoned animal may die in
a matter of minutes . However, a remedy
that has sometimes proved effective is
the intravenous injection of sodium
thiosulfate. Symptoms of poisoning are
depression, paralysis, stupor, and dif-
ficult breathing.
Shortly after animals are first turned
in to a Sudan field, they should be ob-
served closely for symptoms of prussic
acid poisoning. Sudangrass whose growth
has been slowed by drought or partially
killed by frost may be dangerous to
graze, since the cattle will graze the
young, tender shoots that are much high-
er in dhurrin. Grain and silage sorghioms
are usually much higher in the glucoside
than is Sudangrass, and if they are al-
lowed to cross with Sudan the resulting
hybrids will usually be higher in poten-
tial HON producers than Sudan.
The danger of prussic acid poisoning can
be largely eliminated by:
1. Using only certified seed or seed
that is known to be pure Sudan.
2. Letting the crop grow at least l8
inches before grazing.
3- Not feeding excessively hungry
cattle.
h. Not grazing frosted or drought-
stunted crops.
Diseases
Probably the most severe disease of
Sudangrass in Illinois is leaf blight,
caused by the fungus Helminthosporium
turcictim. Leaf blight lesions appear
first as water-soaked areas. Drying out
occurs as the lesions spread to elon-
gated, irregular areas. Entire leaf
blades may be and frequently are killed.
Bacterial leaf diseases may also cause
considerable damage. Best control of
the leaf diseases may be obtained by the
use of resistant varieties. Such vari-
eties as Piper, Greenleaf , and _ Lahoma
showed a relatively high degree of dis-
ease resistance in Illinois in 195^ and
1955.
Insects
The chinch bug is more harmful to Sudan
than any other insect; and in years when
chinch bug infestations are severe,
stands of Sudan may be practically elim-
inated. None of the varieties are com-
pletely resistant to this pest, but the
sweet types possess more resistance than
do the common varieties.
Varieties
There are two main classes of Sudan-
grass: those which have sweet and juicy
stalks and those which are non-sweet or
relatively dry-stalked. The mid-rib of
the leaf of sweet types is cloudy ap-
pearing, and the pith in the stalk is
almost completely juicy. The palata-
bility of the sweet types is high, even
at later stages of growth, and they are
generally eaten more readily when pas-
tured than the non-sweet types. The
dry-stalked varieties cure more readily
when cut for hay and are usually higher
in yield than the sweet types.
Sweet Sudangrass was developed in Texas
from a cross of Sudangrass and Leoti
sorghum. Its performance in Illinois
has been good, and excellent quality
certified seed has been available from
Oklahoma, Texas and California. _The
damage caused by leaf diseases on Sweet
varies considerably, depending upon the
location and the year. Sweet is a juicy-
stalked variety, quite early in maturity,
and has reddish-brown colored seed.
Piper was developed by the Wisconsin
Experiment Station and released in 1950'
It is generally more vigorous in the
northern states than other varieties, is
mostly dry-stalked, has a lower level of
hydrocyanic acid potential and increased
resistance to leaf blight and anthrac-
nose. The strain is not homozygous, be-
ing somewhat variable in seed and foli-
age color. In variety trials conducted
in Illinois during the past two years.
Piper has consistently outyielded all
other varieties and has been damaged
only slightly by leaf diseases.
Common, a dry-stalked type, was devel-
oped from many of the early introduc-
tions. In Illinois Common yields well,
but because of its susceptibility to
leaf diseases and extreme early maturity,
it is not recommended when seed of other
varieties can be obtained.
Greenleaf is a new juicy-stalked variety
of Sudangrass released by the Kansas
Experiment Station in 1953- It has a
low HCW potential, and the seed is loni-
formly mahogany-colored. Preliminary
results in Illinois show that it is
exceptional in resistance to leaf dis-
eases and degree of leafiness, but is
somewhat lower yielding than many other
varieties . Further testing of Greenleaf
is necessary before its adaptability in
Illinois can be determined.
Lahoma is a sweet Sudan and in tests in
Oklahoma, where it was developed, it has
shown uniformity, good leafiness, palata-
bility, and greater resistance to leaf
diseases and later maturity than other
sweet Sudans. It is rather tolerant to
chinch bug attack. At Urbana and Browns-
town, Illinois, the yields of Lahoma have
been relatively low, but at Carbondale
it has performed very well. Lahoma also
needs further evaluation in this state
before its performance can be deter-
mined.
Carl W. Hittle
5-7-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
F-21
ORCHARDGRASS AND ITS MANAGEMENT
Orchardgrass is a long-lived perennial
and a distinctly bunch-type grass with
folded leaf hlades and compressed
sheaths. Because it does not produce
stolons or linderground rhizomes^ it forms
an open sod. This characteristic makes
it a good companion crop for pasture and
hay when seeded with such legumes as al-
falfa^ Ladino clover, red clover, and
lespedeza.
Orchardgrass owes its common name to its
shade tolerance and its consequent use
in orchards and other shaded areas. In
many other English-speaking countries,
it is commonly known as cocksfoot, a
name that aptly describes its distinctive
stiffs finger-like panicles upon which
spikelets are "borne in dense clusters or
glomule s .
Characteristics. Although less i-7inter
hardy than some of the other pasture -
type grasses, orchardgrass starts to
grow early in the spring and grows very
rapidly. It matures for hay or seed about
two weeks earlier than smooth bromegrass
and about three weeks earlier than tim-
othy. It recovers rapidly after mowing
or grazing and produces large quantities
of leafy pasturage or aftermath. It may
produce less first-crop hay or silage
than some of the other pasture -type
grasses, but because it continues to grow
during the summer, its total production
of hay is equal to or higher than that
of many other grasses. Because it will
grow during the drier, warmer part of
the summer and recovers rapidly follow-
ing defoliation, its seasonal yield is
also more uniformly distributed.
Regions of adaptation. Orchardgrass is
grown most extensively and is most im-
portant in the southern half of the so-
called timothy-bluegrass belt, extending
from southern New York to southern Vir-
ginia and westward through Kentucky, Ten-
nessee, southern Ohio, southern Indiana,
southern Illinois, Missouri, and eastern
Kansas. However, it can be grown to
advantage in the northern part of Indiana
and Illinois as well.
Orchardgrass is better adapted to and
more productive in the southern range of
the timothy growing belt than smooth
bromegrass or timothy because it will
tolerate more heat and drought. It will
survive and grow better on thinner and
less fertile soils than timothy and
especially smooth bromegrass. However,
orchardgrass is not well adapted to tight,
poorly drained soils, especially if they
tend to be cold. It responds well to
high levels of soil treatment, particu-
larly nitrogen supplied either from a
chemical source or by a legume seeded in
a grass -legume mixture.
Uses of orchardgrass. Orchardgrass is
used primarily for pasture, but it can be
used for hay and silage as well. It is
recommended for use in permanent pasture
mixtures along with annual lespedeza,
white and Ladino clover, and other
grasses. It is frequently substituted
for timothy in mixtures with red clover
or alfalfa for hay; however, it may ma-
ture somewhat earlier than the legume^
and the legume may have to be harvested
sooner than normally to prevent the
orchardgrass from becoming too mature.
Because of its vigor and productiveness,
orchardgrass lends itself well to use in
pasture renovation where short-term pas-
tures are fitted into a forage program.
Seedling stands are easily established,and
excellent yields of forage can be ob-
tained the first crop year.
Management . Although generally con-
sidered to be tolerant to grazing, orchard-
grass does not persist \inder continuous
close grazing. To best utilize the
forage^ it is therefore necessary to fol-
low a program of rotational grazing.
To maintain a high-quality orchardgrass
pasture;, remove the early spring gro\vrth
by grazing or clipping before it reaches
the full head stage. Then rotationally
graze the regroirth so that an area vill
be grazed for about one week and allowed
to rest from three to four weeks. To
control weeds and permit uniform re-
groirth; clip the grazed area to a height
of about four inches with a field mower
following removal of cattle. Undergraz-
ing or delaying grazing not only reduces
palatability and feeding value ^ but may
also weaken the legume stand in a mixture
as a result of excessive competition from
the grass.
Orchardgrass makes excellent hayandpro-
duces high yields if cut in the earlier
stages of maturity, preferably at the
early head stage. After this stage it
m.atures rapidly and becomes woody and
unpalatable. This early maturity is of-
ten a disadvantage at the first cutting
when orchardgrass is seeded in a mixture
with alfalfa.
Orchardgrass makes excellent silage. If
it matures before it can be grazed^, it
can therefore be ensiled. The silage
can be used to supplement late summer
pasture or as winter roughage. Often
the entire first crop is removed for
silage. Orchardgrass should be ensiled
at about the early head stage to insure
high yields of good-quality silage.
Seedling establishment. The best time
to seed orchardgrass is usually in the
early spring , although late summer or
fall seedings are successful if made
early enough to allow the seedlings to
become established before mnter. Or-
chardgrass seedlings are less winter
hardy than many of the common pasture
grasses, and fall seedings are more sub-
ject to winterkilling, particularly in
the northern part of the state. Spring
seedings should be made early enough to
permit the seedlings to become estab-
lished before weeds become a problem.
In general, good stands of orchardgrass
can be obtained in mixtures seeded at
four to eight pounds per acre. In pure
grass stands, the rate should be in-
creased to 10 or 12 pounds or more, de-
pending on the germination percentage of
the seed. Either broadcast or band
seeding m.ethods can be used, but band
seeding requires less seed. For best
results, soil fertility should be kept
as high as possible. Orchardgrass is
not seriously troubled with insect pests,
but such diseases as anthracnose, leaf
stripe, leaf rust, and scald do reduce
the quality of the forage and may reduce
the vigor of the stand.
Varieties of orchardgrass. Recently
Potomac , a new variety, has been released
for certified seed increase. It has some
resistance to leaf rust and shows some
superiority over the common strains in
seedling vigor and leafiness.
Earl C.
Spurrier
6-U-56
JNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
G-13
WINTER BARLEY IN ILLINCIS
Winter barley is not a new crop in Illi-
nois, but there is a demand at present
for a small grain crop to be planted on
areas taken out of wheat production by-
acreage allotments. Winter barley might
well substitute for wheat on these areas
in the southern half of Illinois.
Winter barley has several advantages
over spring -planted barley. \Tiien ' it
survives the -vrinter it yields more and
its quality is better than the yield and
quality of the spring types. Winter
types normally mature earlier and thus
escape the higher temperatures and some
diseases that are serious hazards to the
spring crop.
Winter barley makes excellent fall pas-
ture and is a good companion crop for
legumes and grasses. The grain makes ex-
cellent livestock feed and is considered
nearly equal to corn in feeding value
when fed to poultry and certain types of
livestock.
U. S. Route 3*5, or a line from Hannibal,
Missouri, through Decatur, Illinois, is
usually considered about the northern
limit for winter barley. But the right
variety, planted early under favorable
conditions, has a good chance to sur-
vive somewhat farther north. \-Jhile it
is scme\^hat hazardous to grow imiter
barley north of U. S. Route 36, if it
winterkills you can still plant a spring
crop •VTith little extra seedbed prepara-
tion. Your only loss would be the cost
of seed plus planting charges.
Cultural practices. Winter barley is
not so winter hardy as winter wheat or
winter rye, but it is hardier than win-
ter oats. Early seeding is recommended
to give the crop a chance to become es-
tablished before cold weather prevents
further growth.
Seed in time to insure a good root
growth and at least four inches of top
growth before the crop becomes winter
dormant. The right date is about 10
days to two weeks ahead of normal wheat
seeding, that is, from September 15 to
October 15, depending on how far south
you are. Seed with a grain drill at the
rate of eight pecks an acre on a well-
prepared seedbed.
Barley responds well to fertilizers, and
fertile soils mean good root growth and
more winter hardiness. A high nitrogen
content in soils may cause barley to
lodge. The straw of all barleys will
break rather quickly once they are ma-
txore. For this reason it is important
not to delay harvesting.
Varieties. Besides yield, winter hardi-
ness and straw strength are the most im-
portant characteristics to consider in
choosing a variety of barley. At pres-
ent the fo.iJLowing varieties are being
recommended: Kearney, Reno, and Ken-
tucky #1 for the area between U. S. Route
36 and U. S. Route 50 (which is frcm St.
Louis east to Vincennes, Indiana) and
these three plus Kenbar and Mo, B-UOO
for the axea south of U, S, Route 50*
Kearney is one of the most winter hardy
varieties, is medium early, and has a
medium stiff straw. Reno has a slightly
stiffer straw than Kearney but is not
so winter hardy. Kentucky #1^ is vlliter-
hardy, has a rather weak straw, and is
later than either Kearney or Reno.
Kenbar is the earliest and the shortest
of the varieties recommended. It is not
quite so hardy as Reno but it has a much
better straw. It is more resistant to
smut, mildew, and scald than Reno,
Kearney, or Kentucl^y /)-l.
Mo. B-UOO is a high-yielding variety early as Kenbar nor does it have so good
that is resistant to loose smut, mildew, a straw,
and splot blotch. Because it grows rap-
idly and vigorously in the fall it makes The 1955 yields of the recommended vari-
excellent fall pastxire. It is not so eties are shown in the following table:
Per acre
yield
Variety
Urbana
Brownstown
Dahl gren
Carbondale
Average
bu.
bu.
bu.
bu.
bu.
Kearney
55.7
39.9
24,3
25.2
36.3
Reno
68.6
52,0
26.0
32.9
it4.9
Kentucky #1
5^.0
39.8
19.6
30.7
36.0
Kenbar
2k, Q*
60,9
26,9
18,8^^
32.8
Mo, B-1+00
62.4
• • • •
, , , ,
31.1
U6,7
*Poor fall stands due to low germination reduced yields at these locations.
R, 0. Weibel
9-12-55
AGRONOMY FACTS
G-14
WINTER WHEAT
Winter wheat ranks third as a cash grain
crop in Illinois. The agricult-ure of
southern Illinois is strongly supported
hy this crop. The climate and soils of
this area are veil suited to the produc-
tion of high-quality soft red winter
wheat. On the other hand, hard red win-
ter wheats produced in Illinois, espe-
cially in the southern half of the state,
are usually too low in protein to produce
a satisfactory bread flour. And because
of the characteristic of the protein,
the flour is not suitable for cakes or
pastry.
Reccmmended Varieties
Soft
Saline
Knox
Seneca
Hard
Pawnee
We star
Pone a
Acceptable Varieties
Soft
Royal
Vigo
Butler
Hard
Triumph
Growing both soft and hard wheats in the
same area has caused a considerable
amount of concern. Because local eleva-
tor personnel cannot always distinguish
between them, they are often mixed in
handling or in shipment. Thus they are
graded mixed wheat when they reach the
inspection points. Mixed wheat is below
the straight grades on the market. To
protect himself, the local buyer, if in
doubt, must class the wheat mixed.
This problem will continue as long as
there is a price differential in favor
of hard wheat and as long as we have no
quick way to determine protein at the
local buying points. We can help con-
siderably by recommending that growers
know the variety they are growing. All
persons selling seed should stress the
name of the variety and sell only those
that are reccmmended.
Variety Descriptions. All of the varie-
ties listed below have produced good
yields in Illinois. All have acceptable
grain quality for their class.
Mew Varieties Being Increased
Soft
Dual
Vermillion
Of the recommended soft varieties. Saline
is the tallest and also the latest.
Knox has the shortest straw and is the
earliest, Seneca is intermediate. Pawnee
and Ponca, hard wheats, are very similar
in maturity and straw strength. Both
are a few inches shorter, a little ear-
lier, and have slightly stiffer straw
than We star.
The grain of Ponca will not bleach out
in the field so readily as Pawnee, and
hence its milling quality is better.
Westar is more resistant to mosaic than
Pawnee or Ponca, but all three may be-
come heavily infested when conditions
are favorable.
Other characteristics of the reccmmended
varieties are given in the tables on
the back.
Table 1. --Characteristics of Varieties of VJheat Reccmmended for Illinois
Area of
Lodg-
Winter
"■"Rela-
CI.
state vhere Test
ing
hardi-
tive
Head
Grain
Variety
Ko,
adapted* weight
res.
ness
height
type
texture
Saline
Knox
1267^^
12798
All
C &
S
Med.
Med.
Excel.
Excel.
Excel.
I-med.
Tall
Short
Bearded
Smooth
Soft
Soft
Seneca
Royal
12529
12558
S
C &
S
Low
High
Excel.
Med.
Fair
Excel.
Med,
Tall
Smooth
Bearded
Soft
Soft
Vigo
Butler
12220
12527
c &
s
S
Med.
Med.
Med.
Excel.
I-med.
I-med.
Tall
Tall
Smooth
Bearded
Soft
Soft
Dual
Vermillion
13083
I27I+8
c &
All
S
Lew
Med.
Excel.
Excel.
Excel.
Excel.
Med.
Med.
Smooth
Smooth
Soft
Soft
Pavnee
Ponca
11669
12128
N &
W &
c
c
Med.
Med.
Med.
Med.
Excel.
I-med,
Short
Short
Bearded
Bearded
Hard
Hard
We star
Triumph
12110
12132
N &
c
c
Med.
Med.
Med.
Med.
Excel.
I-med,
Med,
Short
Bearded
Bearded
Hard
Hard
* W = North;
C = Central; S =
South.
1
Tahle 2. --Reaction of Wheat Varieties to Diseases Common in Illinois
Leaf
Stem
Loose
Covered
Sep-
Hessian
Variety
Mosaic
rust
rust
smut
smut
Mildew
toria
Scab
fly
Saline
R
I
I
S
S
S
S
S
s
Knox
R
R
S
S
S
S
S
S
s
Seneca
R
S
S
I
S
S
s
S
s
Royal
R
S
I
S
S
S
s
S
s
Vigo
R
I
S
I
S
S
s
S
s
Butler
R
S
s
I
I
&'
s
S
s
Dual
R
R
S
S
S
I
s
S
R
Vermillion
R
R
s
s
S
S
s
S
s
Pawnee
S
I
R
R
R
S
s
s
S
Ponca
S
I
R
R
R
s
s
S
R
Westar
I
I
S
I
S
S
s
s
s
Tri\iQiph
S
S
S
I
S
s
s
s
s
R = Resistant; S = Susceptible; I = Intermediate,
R. 0. Weibel
9-19-55
AijRONOMY rMciS
"BLAST" IN OATS
G-15
"Blast" is a term that is applied to a
type of sterility in oats. It is evi-
denced by white, empty glumes in the
lower branches at the base of the pani-
cle about the time the oats are in full
head. The condition is also called
blight, blindness, and white ear.
Blast is not a disease; it results from
inability of some of the spikelets to
develop completely. Anything that in-
terferes with the physiological processes
during development of the plant, partic-
ularly the panicle, may cause blast.
Late seeding, lack of moisture, nutrient
deficiencies, insect attacks, and dis-
ease, either singly or in combination,
will increase the condition. Tillers
are affected more than main stems.
The base of the panicle is where blast
is most prevalent. The sequence of de-
velopment in the oat panicle makes it
easy to imderstand why. An oat panicle
is composed of many branches, each of
which ends in a many-flowered spikelet
in which usually only two flowers pro-
duce seed. The main axis of the panicle
terminates in a spikelet. Beneath the
tip spikelet, and placed alternately on
the main stem of the panicle, are five to
six groups of spikelet -bearing branches.
The number of spikelet -bearing branches
increases from the tip of the panicle
downi-ra-rd.
The first structure ■ that develops from
the main axis of the panicle is a branch
primordium (a branch of the first order),
which is the beginning of the group of
branches at each node of the panicle.
From the first -order branches, branches
of the second order are formed; from the
second order, the third order; and so on.
At the nodes, especially the basal group,
branches of the fifth and sixth orders
may be found.
The oat panicle starts to develop from
the tip spikelet and proceeds to the
base. At any node, development begins
with the first -order branch, followed in
sequence by the second, third, etc. Con-
sequently the panicle is oldest at the
tip and yo\ingest at the base, and at any
node the first -order branches are oldest
and the fifth- or sixth-order branches
are youngest.
It takes some time for a panicle to de-
velop. At Urbana a panicle of Clinton
requires 15 to 18 days, or from about
May 10 to 25 or 28, to develop fully.
Heading occurs about 15 days later.
Adverse changes during panicle develop-
ment, especially the first half, will
affect the youngest parts more than the
oldest. The youngest parts are more
susceptible because they are farthest
from producing seed. We can thus expect
to find more blast in the basal groups
of branches because that is where the
largest number of high-order branches
are found.
Varieties differ' in amount of blast even
though their environment and maturity
are similar. This fact would indicate
that plant breeders can produce varie-
ties that have a low percentage of blast
and hence higher yields.
0. T. Bonnett
9-26-55
AGRONOMY
G-16
WINTER RYE IN ILLINOIS
The acreage of -winter rye has climbed
steadily since 1951/ according to the
Illinois Crop Reporting Service, The
crop harvested in 1955 vas the largest
since 1919 . Perhaps we should take a
closer look at this age-old crop that is
making a ccmetack in Illinois. Why the
increased interest in rye?
1, Wheat and corn acreage allotments
have resulted in extra land.
2, More rye is being used for pasture
and forage.
3, Rye is being inter seeded as a winter
cover and as a green maniire crop in
corn and soybean fields both before
and after harvest.
Characteristics . Winter rye is the har-
diest of the cereals — "the roughest,
toughest of them all." It will grow on
poorer soil, in drier soil, and in colder
weather than our other grain crops. It
will produce in soil that is sandy or low
in fertility where other cereals will do
little or nothing. On good soils rye is
capable of making high yields, although
not so high as wheat.
Two-year comparisons of average acre
yields from several rye varieties with
average yields frcm wheat varieties grown
in the same field showed the following
res\ilts: Urbana, central Illinois - wheat
k8 bu. and rye 39 bu. DeKalb, northern
Illinois - wheat 3^ bu. and rye 30 bu.
Although a little spring rye is grown in
the western United States, the Illinois
rye crop is all of the winter type and
therefore is fall seeded. Rye differs
from the other common small grains in
being almost completely cross ferti-
lized. That is, the flowers on one plant
are fertilized not by their own pollen,
but by pollen from other rye plants.
Culture . Rye ^d-ll respond to good cul-
tural practices and to fertilizers, but
the return for fertilization is generally
greater on wheat and other grains. A
seeding rate of 5 "to 6 pecks per acre
is thought best for Illinois. The crop
can be sown frcm August to Kovember, Al-
though a good seed bed is desirable, rye
has the ability to germinate and grow
under poor seeding conditions.
Uses. Rye may be used as a cash grain,
as a grain feed, as pasture, or as a
green manure crop.
As a cash grain the crop generally goes
into the following trade channels: dis-
tilling, dry milling, feed, export, and
seed. The distilling industry uses rye
for whiskey and alcohol. The dry millers
make rye flour, which is generally blended
with wheat flour to make the kind of rye
bread Americans prefer. The feed indus-
try also blends rye with other grains.
Although rye may be used as a feed crop
on the farm, it is less valuable than
corn, wheat, or barley. Alone, it is
somewhat unpalatable and is considered
heavy and sticky. For best results, it
should be mixed with other grains.
Rye makes excellent fall and spring pas-
ture. It is superior to other small
grains in Illinois because it will grow
later in the fall and start growth ear-
lier and more rapidly in the spring.
The protein content of young rye plants
may run 30 percent or higher in the
spring on fertile soils.
Use of rye as a green manure is rela-
tively new in the Corn Belt. At present
the agriciiltural experiment stations are
doing a great deal of research on the
various aspects of this potentially im-
portant cultural practice. In addition
to supplying organic matter^ rye serves
as a winter cover or guard against ero-
sion and it may also be pastured. If
corn is to follow rye, the land should
be plowed in late April or early May,
and nitrogen should be added.
The success of rye as a green manure will
depend to a great extent en soil mois-
ture. In late, wet springs, the crop
will mature before it can be plowed, and
in dry springs it will remove large
amotints of water from the soil profile
before corn is planted.
Results from Winter Rye Variety Trials at Urbana and DeKalb, Illinois,
195^ and 1955
Adams
Balbo
Caribou
Emerald
Imperial
Pierre
Tetra
Petkus
Yield
Test
weight
Height
Plant
erect-
ness
Head-
Urbana DeKalb
ing
Variety
195^ 1955 Av. 195'J- 1955 Av.
date
bu, bu, bu,
in. 7 ^3.8 i^2.8
37.9 ^^.9 ^1.^
bu. bu, bu,
22,0 3^.5 28.3
29. i^ 30.6 30.0
37.3 38.1^ 37.9
29,1 i+1,9 35.5
3>h,S Uo.8 37.7
37.0 36.9 37.0
I • • • • • • •
33.2 3G.k 34.8
3i^.3 36.6 35.5
30,7 32.8 31.8
lbs.
in.
perct.
51.7
57
^5
May 9
51.5
55
57
" 3
51.2
50 •
• •
" 13
51.0
56
hi
" 9
51.0
57
55
" 10
51.8
5^
51
" 8
• • • •
13.i+
• • • t
50.1
53
75
16
Balbo is believed to be the first choice
for pasture because it will grow earlier
in the spring. In extreme northern Illi-
nois, however, it may siiffer from winter-
killing and Pierre or Emerald might be
better. Tetra Petkus has larger kernels
and wider leaves, but it has not out-
yielded Balbo in either grain or forage.
J. W.
Pendleton
10-3-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
m
S-5
ROOT AND STEM ROT OF SOYBEANS
Root rots occur on soybeans in Illinois
from time to time^, depending largely on
weather conditions. They are likely to
he noticeable in wet seasons^ especially
in low spots in fields where drainage is
poor. One of these root rots is caused
hy Rhizoctonia, a fungus that is present
in most soils where crop plants are
grown. The fungus usually attacks young
plants when the soil is abnormally wet^
causing a reddish-brown decay of the
outer layer of the main root and basal
stem. Much of the secondary root system
is destroyed^ and the plants wilt and
die. Dead plants occtu: typically in
areas four to 10 feet in diameter, usually
distributed irregularly over a field.
In most seasons Rhizoctonia root rot is
of little economic importance.
During the 1955 season, xa. root and
stem rot new to Illinois was discovered
in five or six fields in northern and
central Illinois. This disease affects
plants of all ages. Seedlings that have
just emerged may shrivel and die, leaving
gaps ranging from a few inches to several
feet in the rows. Older plants wilt and
ic"y 'U-P, or they may be severely stunted
and perhaps wilt only slightly at midday.
When such plants are dug or pulled, they
show a badly decayed root system. The
disease is, however, not confined to the
roots; often the brown decay is noted on
the stem several inches above the soil
line.
Although this root and stem rot is espe-
cially damaging in poorly drained areas
of the field, it sometimes occurs on
higher ground. The disease is not wide-
spread or serious in Illinois at present,
but it should be watched closely because
of its potential threat to soybean pro-
duction.
This same root rot has "been present in
northwestern Ohio since 1951« It has be-
come prevalent and destructive in the
clay soils of the old lakebed region.
The disease is caused by a fungus, iden-
tified by Ohio Experiment Station pathol-
ogists as a Phytophthora. Sometimes
it is impossible to differentiate the
Phytophthora root rot from the one
caused by Rhizoctonia. However, the Rhi-
zoctonia root rot lesion usually has a
reddish brown color, while that of the
Phytophthora root rot is brown. Also,
the Phytophthora root rot seems to per-
sist throughout most of the growing sea-
son, while the Rhizoctonia root rot
usually disappears before mid-July.
Work at the Ohio Station indicates that
the varieties Illini, Monroe, and Black-
hawk are resistant to Phytophthora,
while Hawkeye, Lincoln, and Harosoy are
susceptible. Seed treatment has no
value in preventing seedling blight. It
is recommended that rotation mth other
crops be practiced on land where the
root rot has appeared.
Investigations on Phytophthora root and
stem rot are under way at the Illinois
Experiment Station, and more information
on this disease will be made available
as controls are developed or resistant
varieties are released.
U. S.
D. W. Chamberlain
Plant Pathologist
Dept. of Agriculture
5-21-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
S-6
ESTIMATED HAIL LOSSES TO SOYBEANS
Because soybeans grow throughout the
cropping season, they are subject to in-
jury by hailstorms. Fully two-thirds of
the hailstorms that occur in the Corn
Belt come between May and September,
when soybeans are in the field.
Insurance to protect the farmer against
loss of his crop by hail is becoming more
common each year. Insurance companies
have conducted and are conducting experi-
ments designed to get information on how
to evaluate losses to soybeans from hail
injury. Experiment stations, too, have
studied the problem, and the insurance
companies have supported some of this
work. Illinois has been conducting such
tests for six years, Iowa for six years,
and North Carolina for one year.
Results at the Iowa Station showed that
when all leaves were removed at the blos-
soming stage the yield of beans was l8
percent of norm.al, and when the stems
were broken in addition to 100 percent
leaf removal the yield was l6 percent of
normal (see Stage 7 in diagram). Leaf
removal earlier and later than this
critical period caused progressively
less reduction in yield.
Recoverability is highest when plants
are about three weeks old and after
three or four trifoliate leaves have un-
rolled. Although leaf removal at this
stage causes some reduction in yield,
the drop-off in production is less than
before or after this stage. In the dia-
gram this stage is shown as 2.
Effect of stage of development on recov-
ery from injury. The soybean plant pos-
sesses great ability to recover from
injury, particularly at certain stages
of development. Knowing the stage at
which the damage occurred is therefore
just as important in estimating losses
as knowing the extent of the injury.
Unless the plants are broken off at the
ground or are otherwise destroyed by
beating of the hail, they will yield
some grain. That is to say, there is no
stage when injury will completely elimi-
nate all yield.
The stage of lowest recoverability is
just at the end of blossoming. At this
time the pods on the lower branches are
nearly full length and beans are devel-
oping in them. Tlie pods on these branches
are far enough along to yield some grain,
but pods in the top part of the plant
are just starting to form. No new blos-
soms or leaves form afterward, and removal
of the leaves robs the plants of the
photosynthetic area when it is needed
most.
Extent of injury. In general, yield is
reduced in direct proportion to the per-
centage of leaf surface rem.oved. Bruis-
ing of the stem also affects yield.
Tests at Iowa showed that breaking of
stems lowered yield below that caused by
leaf removal. This decrease amounted to
only 15 percent at the first-bloom (2)
stage, but increased to over 50 percent
at the critical (7) stage.
A severe storm that kills plants out-
right will reduce the stand and conse-
quently the yield, but the reduction v;ill
vary with stage of growth. If it occurs
during the early (l to 3) stages, as
many as 50 percent of the plants can be
destroyed without any great reduction in
yield. But at Stage 6 or later, a re-
duction in stand will mean a marked de-
crease in yield. By the end of the
blooming period, the plants have lost
their capacity to spread out and take up
the extra space.
Other effects
Loss of leaw
the critical (7) stage delays matui'
the plants, but such injury aft^.
stage seems to hasten maturity. Actu-
ally, however, maturity only appears to
be speeded up. The crop comes only from
the most advanced pods because the leaf
area is not sufficient to provide reserves
for the later formed pods.
large leaf losses reduce seed size and
also decrease the oil content of the
seed if the loss occurs while the grain
is filling.
Estimating losses. It is hard to deter-
mine the percentage of leaves removed by
a hailstorm. It is possible to tell the
stage of plant development by carefully
observing the plants immediately after a
storm. But figuring out how many leaves
remain in relation to the number before
the storm is like probing in the dark.
Because of this difficulty, Dr. James C.
Weill, who did his Ph. D. thesis on the
effects of artificial hail on soybeans
at the University of Illinois, suggested
the possibility of determining the de-
gree of injury by counting the terminal
buds that had been knocked off. In two-
year tests (1950-51) Weill blew cracked
ice through a three-inch rubber tube onto
soybean plants at various stages of de-
velopment. In this way he inflicted in-
juries similar to those caused by natural
hail, removing different percentages of
leaves and at the same time damaging
stems and removing terminal buds .
Weill counted the leaves before treat-
ment and then counted those left on the
plants after treatment. He also coimted
the plants from which the topmost or
terminal bud had been removed. In both
years in which he made these tests he
found a very high correlation between
the percentage of leaves lost and the
percentage of terminal buds lost.
Because in these tests the correlation
between loss of leaves and loss of ter-
minal buds was so close and because the
percentage of terminal buds that are
lost can be easily and accurately deter-
mined after a hailstorm, Weill suggests
that the severity of injury be estimated
by counting the number of plants in a
himdred from which the terminal buds have
been rem^oved.
G. H. Dungan
6-18-56
100
+3
a
Q)
CO
C)
-P
U
C
(U
^^
PnrH
yu
C
-H
Ti
(i)
Ti
u
M
:3
fl)
"-n
•H
r,
>^
•H
a
i^
D
•H
01
<+-l
!h
C)
0
20
lOC;^ leaf remova-l
^. + stem breakrage
3 ^ 567
Stages in plant development
10
Diagram of grain yield reduction as a result of f-: . _ .
hail damage at 10 stages of soybean plant development.
"erities of
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
AND -^^Z
TESTING -^^
■■■' S
SF-37
CHEMISTRY OF ORGANIC NITROGEN IN SOILS
Nitrogen is different frcm the other nu-
trient elements in the soil because it
occurs almost entirely in organic ccm-
"bination. Only a very small fraction of
the nitrogen- -usually frcm 1 to 3 per-
cent --is in the inorganic forms at any
one time.
During the year nitrogen is released
slowly from the organic matter to the
mineral forms ^ which are the available
forms of nitrogen in soils (see SF-4).
The amount that is released depends on
the kind and amount of nitrogen in the
organic matter^ on climatic conditions;,
and on the physical condition of the
soil.
Several methods are used to estimate the
quantity of nitrogen converted to the
available forms during the growing sea-
son, but all of these methods have cer-
tain limitations (see SF-I7). If we
knew more about the nature of organic
nitrogen in soils, we could overcome
seme of the uncertainties involved in
estimating nitrogen availability and in
making fertilizer recommendations.
Unfortunately, we know very little about
the nature of soil organic nitrogen or
about how soils differ in their contents
of organic nitrogen compounds. Do fer-
tile soils contain nitrogen compounds
that are not found in infertile soils?
Are some nitrogen compounds in humus
easily mineralized while others resist
attack by soil microbes? Are well-
fertilized and manured soils different
from infertile soils in their contents of
certain kinds of organic nitrogen com-
pounds?
To answer these questions the Department
of Agronomy has a research project under
way that is concerned with the chemistry
of humus nitrogen. As indicated above,
the purpose of this study is to find out
what forms of organic nitrogen the soils
contain and to determine how cropping
and management practices affect these
forms.
Recently a study was made of the protein
nature of soil humus. The building
units of the proteins--the amino acids--
were isolated frcm several soils and
their amounts were determined. Briefly,
the results showed that soils differed
in the quantity and quality of their pro-
teins. In seme soils as much as half of
the nitrogen was in the form of pro-
teins; in others less than a third of
the nitrogen was proteinaceous. Also,
the amino acid composition of the pro-
tein material in one soil was quite
different frcm that in another. This
difference suggests that the ability of
humus to supply nitrogen to the plant
may depend considerably upon the nature
of its proteins.
The famous Morrow Plots at the Univer-
sity of Illinois were used to determine
the effects of some long-time rotations
on the distribution of amino acids in
soils. The results of this study were
very interesting. It was foimd that the
proteins in the soils from the untreated
plots (such as the i^ntreated continuous-
corn plot) were low in the kinds of ami-
no acids that would be expected to be
ready sources of nitrogen--for example,
mono -amino acids like glycine, alanine,
valine, and leucine, and the amino acid
amides, asparagine and glutamine. The
proteins in the soil frcm the corn-oats-
clover rotation plot that had been
manured, limed, and supplied with phos-
phate were high in these amino acids.
These results show that when soils are
heavily cropped they lose heavily in cer-
tain kinds of amino acids. Hence, not
only does an intensive system of farming
deplete the soil of proteins, but the
protein material that remains is of low
quality.
Needless to say, more research needs to
be dene before the full practical signif-
icance of these findings can be deter-
mined. But the results obtained thus
far have served to emphasize the desir-
ability of using management practices
that will furnish a continuous supply of
actively decomposing organic matter to
the soil. The soil organisms use plant
residues and manures as sources of food.
During growth and reproduction they syn-
thesize body proteins. It is these pro-
teins that later become good sources of
mineralizable nitrogen.
Research work on the chemistry of organ-
ic nitrogen in soils is continuing. In
the future we can look forward to unrav-
eling seme of the mysteries surrounding
the availability of hiimus nitrogen in
soils.
F. J, Stevenson
10/31/55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AND
TESTING
AGRONOMY FACTS
SF-38
1955 WHEAT YIELDS - ILLINOIS SOIL EXPERIMENT FIELDS
Cropping 123^56789
Location system 0 M ML MLF 0 R RL RIP RIPK
Dark-colored soils
bu. bu. bu. bu. bu. bu. bu. bu. bu.
Mercer C-C-O-W kk h2. 39 ^0 U6
Macoupin C-B-W-H 26 "^h 39 38 31 32 37 ^0 ij-5
Hancock C-B-W-H 28 36 !+3 i+3 29 2h 28 32 39
Adams C-B-O-W I8 29 30 33 37
Lee C-O-H-W lj-2 1+8 5I 50 4l hG hh 52 1+9
Logan C-C-O-H 1+1 1+7 1+1 1+3 39
Will C-B-C-0-W-H 20 26 29 3!+ I7 I5 20 3I+ 36
Henry C-O-W-H 31 36 hh 51 1+9
St. Clair C-B-W-H 11 25 33 36 I3 12 28 35 37
Putnam C-C-O-W 17 37 38 1+3
Woodford C-C-O-W 37 39 37 39 ^2
Ogle C-O-W-H j_L.ij.iZ. ♦• 18213137 37
25 36 39 5o 28 32 35 1+0 5l
Field
Ale do
Carlinville
Carthage
Clayton
Dixon
Hartsburg
Joliet
Kevanee
Lebanon
McWabb
Minonk
Mt. Morris
Average
Lip;ht- colored soils
Brownstown
Fayette
C-B-W-H
• •
• •
• •
• •
• •
1
33
^5
51
Enfield
White
C-O-W-H
1+
11
26
30
1+
1+
15
20
33
Eving
Franklin
C-B-W-H
0
1+
23
31
2
1+
17
20
27
Oblong
Crawford
C-B-W-H
13
27
36
1+1
0
9
26
1+0
ho
Raleigh
Saline
C-O-H-W
6
13
27
26
15
12
17
25
32
Toledo
Cumberland
C-B-W-H
9
22
52
56
10
11
21+
36
39
Average
6
15
33
37
7
7
22
31
37
Cropping System Symbols:
Soil Treatment Symbols:
C = Corn, B = Soybeans, W = Wheat, 0 = Oats, H = Hay
(legumes and mixed grasses).
0 = Untreated land, M = Manure returned equal to crops
removed,R = Crop residues, L = Limestone, P = Rock phos-
phate, K = Muriate potash.
(Continued en back)
Average whea.t yields in Illinois for
1955 were the highest on record, 31»5
"bushels. Nevertheless yields varied
widely from location to location and
with different soil productivity and
management pi-actices. Yields from the
Illinois experiment fields (see tahle)
illustrate those variations, as they rep-
resent a cross section of the produc-
tive capacity of Illinois soils.
On untreated dark-colored soils yields
ranged from I3 to kh bushels an acre.
On fully treated land the range was 32
to 52 bushels. The response to treat-
ment varied from none to 26 bushels.
Similar variations occurred on the light -
colored soils of the state.
This kind of information should alert
agricultural workers and farmers to the
danger of making prodictions regarding
the response that can be expected from
a given fertilizer or management prac-
tice in any one year. The data also
show the advantages of keeping a soil
adequately supplied with plant food and
ready for any favoi'able growing condi-
tions that come along, such as those of
1955.
The yields given in the table were ob-
tained after bulk application of such
fertilizing materials as manure, crop
residues, limestone, rock phosphate, and
muriate of potash (on individual plots)
and under the cropping system described.
In these experiments no attempt is m.ade
to fertilize the individual crop. Mate-
rials have been applied in quantities
adequate to determine the needs of the
various soils and to supply all crops in
the system under a wide range of seasonal
conditions.
On many of the fields additional tests
have been made with complete mixed ferti-
lizer, various phosphate carriers, nitro-
gen, and potash. In seme cases yields
have been increased by annual direct ap-
plications, and in other cases they have
not.
A detailed explanation of the treatments,
cropping systems, and soil types is given
in Illinois Bulletin 516, "Effect of
Soil Treatment on Soil Productivity. "
This publication is supplemented by a
mimeograph pamphlet that brings data on
each field up to date for each year.
These publications are available from
the Agronomy Department, University of
Illinois, Urbana.
A. L. Lang
ll-li|-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SF-39
EARTHWORMS
In 1777 Gilbert White published a paper
in which he said: "Worms seem to be the
great promoters of vegetation^ which
would proceed but lamely without them,
by boring, perforating, and loosening
the soil, and rendering it pervious to
rains and the fibers of plants, by draw-
ing straws and stalks of leaves and
twigs into it; and, most of all, by
throwing up such infinite numbers of
lumps of earth called worm-casts, which,
being their excrement, is a fine manure
for grain and grass. . .the earth with-
out worms would become cold, hard bound,
and void of fermentation, and conse-
quently sterile."
Since 1777 much information concerning
the relation between the earthworm
population and soil conditions has been
collected. In spite of this knowledge
concerning the activities of earthworms
and the conditions under which they flour-
ish, it still is not clear whether they
affect the productivity of soils materi-
ally.
Certain facts are, however, recognized
and a few of them are worth enumerating:
1. Earthworms occur in fairly large num-
bers in many farm soils. Observa-
tions in the United States and in
other countries indicate that under
favorable conditions the number may
exceed a million an acre, with a to-
tal live weight of more than l/h
ton.
2. The worms are most abundant in soils
that are high in organic matter.
Consequently they may be found most
extensively in forest, pasture, and
heavily manured soils.
3. Cultivated soils are less favorable
for earthworm survival than are
areas that are covered with sod
crops. This difference is explained
in part by their sensitivity to sud-
den changes in the soil temperature
when the late-fall freezes occur.
They apparently need the insulating
protection that vegetation cover
gives the soil.
k. Earthworms appear to need a contin-
uous supply of calcium, which they
excrete from their digestive tract.
Their numbers therefore decrease as
the pH of the soil goes down. Dif-
ferent species also differ in their
tolerance to soil acidity.
5. In virgin soils and in pastures the
worms drag into their burrows leaves
and other dead surface litter. Mi-
croorganisms then attack the litter
and convert it into the so-called
humus fraction of xhe soil.
6. It is believed that earthworm activ-
ity improves soil aeration. The
channels the earthworms make help to
improve air and water movement.
These channels may also serve as
passageways for plant roots. Since
the worms that burrow deeply in the
soil may carry lime and plant nutri-
ent elements into their channels in
the subsoil, the occurrence of roots
in subsoils, which ordinarily would
be unfavorable for them, may be due
to earthworm activity.
Correspondence wwith farmers indicates
concern in seme areas about the effect
of use of anhydrous ammonia on the earth-
worm population. Although no experi-
mental data are available on this subject,
the lack of proof of the value of earth-
worms in soil productivity suggests that
arguments for and against the use of
anhydrous ammonia should be based on fac-
tors other than its effect on the earth-
worm population.
0. H. Sears
11-21-55
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SF-40
BAND APPLICATION OF FERTILIZERS IN ILLINOIS - PART I
Banded fertilizer may "be defined as any
fertilizer that is banded or drilled
with seed. Banding or drilling ferti-
lizer has been a standard practice in
the East and South for many years. In-
terestingly enough, fertilizer so applied
is generally called starter fertilizer
in the Midwest. An eastern or southern
farmer merely says that he is fertilizing
his crop.
other good management practices are fol-
lowed.
On the other hand, there are alternative
management and fertilization practices
through which high yields can be secured
without the use of banded fertilizers.
The farmer's choice of the alternatives
is not solely an agronomic problem--it
is a joint agronomic -economic problem.
Banded fertilizers may serve a number of
useful functions. It is perhaps lonfor-
tunate, therefore, that the term starter
fertilizer was coined. Under certain
conditions, banded fertilizers promote
rapid, early plant growth and vigor.
The seedlings, therefore, are better
able to overcome such early hazards as
grape colaspis, a root-feeding insect,
and adverse weather. Also, by promoting
early, liniform, vigorous growth in row
crops, banding makes it possible to do a
more thorough job of weed control.
These functions help to get the better
stands that are essential for top yields.
On moderately phosphorus- and potassium-
deficient soils, banded fertilizers may
promote deeper and more extensive root-
ing. In wheat, this reduces winter heav-
ing and assures better winter survival.
Banding may also lead to more complete
fertility exploitation by all crops.
Critics of banded or starter fertilizers
often cite this argument to discredit
band or row applications, saying that
banded fertilizers make it possible to
more thoroughly mine or deplete soils.
This may be true where small amounts of
low-analysis fertilizer are applied, but
it is not true where adequate amounts of
high-analysis fertilizer are applied for
each crop. In fact, it is possible not
only to maintain the mineral fertility
of productive soils, but also to main-
tain very satisfactory yields on infer-
tile soils by applying adequate amounts
of high analysis fertilizer provided
In Illinois the fields are frequently
large and the rows long. Application of
banded fertilizers, particularly in ade-
quate amounts, requires frequent filling
of hoppers, adds to the labor and re-
duces the acreages planted per day as
much as 25 percent. Many farmers are
sold on the value of banded fertilizers,
others question their need, and occasion-
ally seme even say that yields have been
reduced. Virtually all complain of the
added labor and weight-lifting.
In most cases starter effects induced by
banding fertilizer are primarily a re-
flection of seedling response to highly
available soluble phosphorus. So let us
examine the chemical aspects of applying
soluble phosphates to soils. This will
help to clarify the reasoning that gave
rise to the practice of banding or drill-
ing phosphatic fertilizers or mixed fer-
tilizers containing available phosphorus.
When soluble phosphorus is applied to
strongly acidic soils or soils that con-
tain excess lime, phosphorus fixation
takes place. Soil chemists at first be-
lieved that phosphorus fixed by strongly
acidic soils was permanently converted
into unavailable forms. The original
thinking behind placement of phosphorus -
containing fertilizers, therefore, was
to minimize soil-fertilizer contact.
Concentrating the fertilizer close to
the seed made it possible for the plants
to take up most of the phosphorus they
needed before fixation occurred. The
-2-
economic loss of phosphorus through fix-
ation was believed to be reduced.
A "better understanding of the chemistry
of soil phosphorus has developed in re-
cent years. Soil chemists have found
that the phosphorus fixed by acidic soils
is not permanently lost. The fixed phos-
phorus is now merely considered "diffi-
cultly available" to plants. It has been
learned that the use of sufficient lime
makes it possible for plants to use the
phosphorus that acidity had made physio-
logically unavailable. Strongly acidic
soils already containing large amounts
of difficultly available phosphorus will
not respond to phosphorus for several
years when first limed to neutrality.
This does not mean that lime can be sub-
stituted permanently for phosphorus on
such soils. It means that lime makes it
possible to reclaim much of the phos-
phorus previously unavailable to the
plant .
The fixation of phosphorus to difficultly
available forms in calcareous (shelly)
soils is a more serious matter. It is
sometimes claimed that rock phosphate is
produced. Chemically this is an untenable
hypothesis. On calcareous soils the
fixation to difficultly available forms
is caused by excess lime. There is no
practical way to remove the lime. For-
tunately the reversion to difficultly
available forms is slow, usually taking
a full growing season. Annual applica-
tions of phosphate-containing fertilizers
are preferable on calcareous soils.
lizer is applied just ahead of seeding.
Top-dressing of permanent meadows in the
spring is preferable. Banding or drill-
ing is best, however, and leads to the
most efficient use of soluble phosphates,
particularly where small applications
are made.
A second problem in the use of mixed fer-
tilizers containing phosphorus is posed
by the immobile nature of soil phospho-
rus. For all practical purposes plant
roots must forage for phosphorus. The
degree to which seedlings are stimulated
into faster growth depends, therefore,
on the extent of the soil phosphorus de-
ficiency. An estimate of the early
growth stimulation of various crops is
illustrated in Figure 1. It is evident
that on very phosphorus-deficient soils
the starter effects can be quite large.
Early plant stimulation may or may not
be reflected in final yields. On moder-
ately acid soils testing medium to high
in available phosphorus, the growth stim-
ulation effects, except for wheat and
clovers, are apt to be small or even ab-
sent.
Potassium is likewise an immobile nutri-
ent for which plant roots must forage.
The stimulating effects of potash in
mixed fertilizers will vary with the
level of soil potassium availability,
much as is the case for phosphorus. On
most soils, however, early growth stimu-
lation is essentially a phosphorus ef-
fect because the phosphorus requirements
of seedlings are usually very high.
To avoid excessive fixation, the ferti-
lizer may be banded or drilled with the
seed. Broadcasting is also satisfactory
if excessive mixing (disking) is avoided
and if the phosphate-containiDg ferti-
The production of radioactive phosphorus
by the Atomic Energy Commission at Oak
Ridge, Tennessee, has made it possible,
through the use of tagged radioactive
phosphorus, to trace the proportion of
Table 1. --Percent of Phosphorus in Corn Secured From Banded
Fertilizer During the Growing Season
Available phosphorus
Percent
of
phosphorus
m
plant from fertilizer
in soil
30 days
60 days
90 days
110 days
Low
High
58
26
36
21
28
17
23
15
-3-
the phosphorus absorhed hy plants from
fertilizers throughout the growing sea-
son. Results of such an experiment con-
ducted with corn in North Carolina are
given in Table 1.
These data indicate that during the
first 30 days of growth corn absorbed
slightly more than twice as much phospho-
rus from band applications on soils that
were low or deficient in available phos-
phorus as from the same applications on
soils that were high in available phos-
phorus. In other words, the phosphorus
in banded fertilizers can be expected to
be more effective in promoting rapid,
early growth with increasing soil phos-
phorus deficiencies.
The data also show that, regardless of
the level of available soil phosphorus,
less and less phosphorus is derived from
the fertilizer as growth progresses and
the root system develops. After 110
days, on soils that were high and low in
available phosphorus, only 23 and 15 per-
cent, respectively, of the phosphorus
needs of the crop came from the applied
fertilizer. The remainder of the phos-
phorus that was absorbed--some 77 sjid 85
percent--came from the soil. This ex-
plains why, on moderately fertile soils,
early growth stimulation of corn by
banded fertilizers often is not reflected
in the final yield. This being the case,
benefits other than yield increases must
be assigned to banded fertilizers for
some soils.
Fig. 1. — Estimated Relative Early Growth Stimulation Induced
by Soluble Phosphorus Applied in Bands to Soils of
Different Soil Phosphorus Test Values
a
o
•H
-p
H
I
•H
-P
CO
o
u
o
a
(U
0)
CD
i)
>
•H
1^
H
(U
Ti
V
-p
-P
to
\^^^^M wheat and Clovers
Corn
Soybeans and Oats
i
b^
t^
^
i:^
El
Low Slight Medium High
Illinois Soil Phosphorus Test Values
E. H. Tyner
12-5-55
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
^oiL PERTitmr
AND ,^
TESTING , -
AGRONOMY FACTS
SF-41
BAND APPLICATION OF FERTILIZERS IN ILLINOIS - PART 2
The preceding Fact Sheet was devoted pri-
marily to some of the fundamental aspects
of hand fertilization. This sheet will
deal with the more practical aspects of
the problem- -amounts^ fertilizer ratios
in relation to soil test values^ etc.
Except on calcareous (shelly) soils ^
banding fertilizers with each row crop
is not urgent on soils testing at least
M- in available phosphorus or above 170
pounds in available potassiiim. On such
soils the primary benefits from banded
fertilizers, if any, are likely to be
secured only with wheat and legumes (see
Fig. 1, Fact Sheet SF-ij-O).
Under these conditions applying mainte-
nance amounts of phosphorus and potassium
once in the rotation will maintain enough
mineral fertility for high crop yields,
assuming adequate nitrogen fertility.
Moreover, these amounts can be applied
with bulk distribution equipment during
slack periods. One exception might be
where grape colaspis must be controlled,
but it can also be controlled with pes-
ticide sprays. Farmers who have serious
grass weed problems or who are shooting
for extraordinary yields may still prefer
to apply seme banded fertilizer in order
to insure better stands.
Banded fertilizers applied to each crop,
however, have a definite place where de-
pleted farms are operated by tenants with
uncertain tenure, or where capital is
not sufficient for build-up applications
or where it is needed more in other
phases of the farm business than for
rapid build-\ip of basic soil fertility.
In general, on very deficient soils one
cannot apply enough fertilizer to get
maximum yields by hill-dropping or band-
ing in the row. Even so, it is possible
to get good yield increases with mixed
fertilizers alone, provided the analysis
supplies reasonable amounts of the most
needed nutrients and provided nitrogen
fertility is adequate. Table 1 lists
first-year phosphorus and potassivmi re-
quirements for moderately high yields at
various soil test values.
Table 1. --Phosphorus and Potassium Re-
quired the First Year at Vari-
ous Soil Test Values
Phosphorus
Potassium
Test value
(P2O5)
(K2O)
lb. /A
lb. /A
L-
60
L
5^
L+
kQ
S-
k2
S
36
s+
30
Uo
• •
120
80
• •
70
120
• •
1|0
170-200
• •
30
The above requirements are stated in
terms of phosphorus (P2O5) and potassium
(K2O) needed per acre. How can we use
such information in determining the prop-
er fertilizer analysis to buy: Let us
assume that the soil test indicates
phosphorus availability as L- and potas-
sium availability as 120 pounds. On the
basis of this information, we would need
to apply 60 poimds of P2O5 and ^0 poionds
of K2O per acre. The P205:K20 ratio needed
in a mixed fertilizer would therefore
be 1.5 to 1.0. It would be difficult
to find a mixed commercial fertilizer
with this ratio. Therefore, we will
choose a fertilizer with a 2 to 1
phosphorus-potassiijm ratio, such as 0-20-
10. If we were to apply 3OO pounds of
this analysis per acre, it would supply
the 60 poimds of P2O5, but only 30 pounds
rather than to pounds of K2O. Even so, we
could still obtain satisfactory yields.
Can this amount of fertilizer be hill-
dropped or banded for row crops? Large
amounts of fertilizer close to the seed
can delay or even prevent germination.
The reason is that the "salt action" in-
hibits moisture uptake by the seed or the
roots just emerging frcm the seed. In
general practice, it is only the nitrogen
and potassium in mixed fertilizers that
are "hot" and that interfere with germi-
nation.
A good rule-of -thumb is not to hill-drop
or band more than 30 pounds of "hot"
solubles (nitrogen (n) plus potassium.
(K20))per acre. It is apparent that the
safe amount will vary with the analysis,
e.g., 3-12-12 (200 pounds), 10-10-10
(150 pounds), 3-9-27 (100 pounds). The
30-pound rule is for dry to average sea-
sons. In wet seasons higher rates may
not be injurious. According to this
rule, the 3C0-pound application of 0-20-
10 could be made provided the fertilizer
attachment would deliver this amount.
If not, 100 pounds could be banded and
the rest broadcast and either plowed
Tinder or disked in.
This explanation shows why it is diffi-
cult to prescribe certain fertilizer ra-
tios for use on crops grown on soils
that may vary considerably in basic fer-
tility. Moreover, a certain analysis
may be satisfactory to apply for a few
years, but carry-over effects may cause
the ratio and the amount to change. Pe-
riodic soil testing will help to deter-
mine the most practical and profitable
ratios and rates.
How necessary is nitrogen in mixed banded
fertilizers? This is a question on which
agronomists do not agree. Experimental
evidence indicates that nitrogen in-
creases the uptake of phosphorus. It
may therefore be argued that nitrogen in
banded fertilizers serves a very useful
purpose. This may be true on soils that
are highly deficient in available phos-
phorus, but it is not so important on
soils that are moderately supplied with
available phosphorus. More will be said
on this point later.
In general, where experience indicates a
substantial response to nitrogen, large
amounts of nitrogen must be applied. It
is usually best to apply a nitrogen ma-
terial, e.g., 20-0-0, 33-0-0, 82-0-0.
This does not mean that 12-12-12, etc.,
are not satisfactory sotirces of nitrogen.
Such analyses, if used in amounts ade-
quate to supply the required nitrogen,
often supply more phosphorus and potas-
sium than can be used efficiently. But
the excess phosphorus and potassium are
not lost through leaching and can be used
later.
Applying nitrogen in large amounts in
mixed fertilizers can, therefore, lead
to inefficient use of phosphorus or po-
tassium if the soil already tests moder-
ately high in these elements. Where high
nitrogen mixed fertilizers are used, the
carry-over effects of excess phosphorus
and potassium should be carefully ana-
lyzed before subsequent fertilizer appli-
cations are made.
In general, conditions preceding planting
and level of available soil phosphorus
should determine the advisability of ap-
plying amounts of nitrogen in mixed fer-
tilizers, such as 3-12-12, Normally the
decay of soil organic matter will take
care of the small nitrogen requirements
of seedlings. If the weather is abnor-
mally cool with frequent rains before
corn is planted, it might be advisable to
use a mixed fertilizer containing seme
nitrogen, particularly on phosphorus defi-
cient soils. On the other hand, if tem-
peratures are normal and moisture is
average, it is questionable whether ni-
trogen will be needed in starter ferti-
lizers even to increase phosphorus
uptake. Nitrogen in mixed fertilizers
may, however, be beneficial in most sea-
sons on weedy fields where a considerable
amount of trash, particularly straw, has
been plowed under.
Is nitrogen necessary for establishing
stands of winter wheat. If wheat follows
soybeans and is seeded just after the
fly-free date, the value of applying ni-
trogen in mixed fertilizer to stimulate
fall growth is questionable. On the other
hand, if seeding is delayed because of
wet weather, seme nitrogen- -perhaps not
more than 3 to 5 pounds per acre --might
be desirable. Applying nitrogen in the
fall on clay-pan soils to avoid the need
for spring applications is a different
matter. See Fact Sheet SF-28.
E. H. Tyner
12-12-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SF-42
PROGRESS REPORT ON A GREEN-MANURING PROJECT
In 1953 3.n experiment was started to de-
termine the value of northern and south-
ern alfalfa varieties, red clover, ladino
clover, sweet clover and lespedeza as
green-manure crops. A comparison of the
green-manure crops with chemical nitro-
gen was also included.
The Series 1102 plots on the Agronomy
South Farm, where the various green-manure
crops were seeded in oats in 1953^ had
the following crop history:
19^8 Corn followed hy an application of
1^ tons of manure an acre.
1949 Corn
1950 Soybeans harvested for seed
1951 Oats in which clover was seeded but
failed
1952 l-Jheat (straw left on land)
1953 Before oats were seeded, three tons
of 200-mesh limestone, 5OO pounds
of 0-20-0, and 110 pounds of O-O-5O
were applied
The Series W-6C0 plots had been in a
rotation of corn, corn, oats, and clover
for 50 years. The clover failed in 1953
and oats were seeded again in 195^ as a
companion crop for the green-manure
crops .
In each series the green-manure crops
were plowed under about a yea^ after
seeding. To some plots where a green
manure had not been seeded (NHi^)2S0i^. was
applied in 195^ ^nd plowed under at once.
KHPIO3 was used in the same way on certain
plots in 1955. Corn was planted by hand
at the rate of four kernels per hill. A
nearly perfect stand was sec\ired in each
year.
The yields of No. 2 corn on these plots
for the years 195^ snd 1955 are given
below.
CORK YIELDS FOLLOWING GREEN-MANURE CROPS
AND WITH NITROGEN FERTILIZERS
Acre
yields
Green-man\ire crops
195^
1955
bu.
bu.
Alfalfa
African
119
102
Chilean
120
111
Indian
123
103
New Mexico
120
108
Northern Common
^\?^
109
Ranger
119
110
Clover
Ladino
119
103
Medium red
124
103
Lespedeza (Korean)
122
103
Sweet clover
White blossom
117
109
Yellow blossom
116
109
AVERAGE
120
103
None
119
103
None plus nitrogen
60 poiinds/acre
m
97
None plus nitrogen
120 pounds/acre
—
99
In each year a randomized block design
■vriLth four replications was used. Conse-
quently it was possible to make a sta-
tistical analysis of the data.
Although seme differences in corn yields
were associated with the previous green-
manure crops and with nitrogen fertilizer
these differences were not statistically
significant. On the basis of this
Information, it cante concluded only that
neither nitrogen fertilizer nor legumi-
nous green-manure crops had a measiirahle
effect on yields of corn in these plots
in 195^ and 1955* This information
should not be interpreted to mean that in
other years or on other soils in the same
years the use of leguminous green-manure
crops or nitrogen fertilizers would not
or did not affect corn yields favorably.
The question then arises why increased
corn yields did not occur. While it is
not possible to say with authority why
these results were obtained, some possi-
ble ejcplanations can be presented: Obvi-
ously these soils were furnishing a
larger amount of available nitrogen than
is usually the case. Again the question
why arises naturally.
In the years 1953, 195^ and 1955;, there
were no periods of excessive rainfall.
In fact, a moisture deficiency occurred
for several extended periods in each of
these years. As a result, it seems rea-
sonable to postulate that the soils, even
without green m.anure, fiirnished suffi-
cient available nitrogen for maximum
yields under the seasonal conditions
that prevailed. This large amoiuit of
available nitrogen was the result of:
1. Moisture and temperature favorable
for nitrate formation.
2. IJot enough soil moisture to cause
leaching losses of nitrate nitrogen
or denitrification.
In other areas on the Agronomy Farm there
was evidence that the availability of
soil forms of nitrogen was high in 195^
and 1955* Consequently the response to
added nitrogen was less than many people
expected.
These data should not be construed to
mean that leguminous green-manixre crops
and chemical nitrogen are not important
in the nitrogen economy of the land.
Rather, they indicate that seasonal con-
ditions must be considered in evaluating
the worth of any soil-improving practice
in a particular year.
J, A. Jackobs
0. H, Sears
1-9-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SF-43
COMPOSTS
Composts have iDeen used in agricultiire
for centuries. Roman farmers prepared
composts from the strav and chaff of
grain crops and from the leaves of for-
est trees. Although the farmers of an-
cient times did not understand all of
the principles involved, they were famil-
iar vith the kinds of ingredients needed
for successful composting.
One kind of composting, knovn as sheet
composting, has been used in Illinois
for more than half a century. It con-
sists of plowing under such materials as
straw, manure, and green manure crops
grown for soil improvement purposes.
Composts have "been used extensively in
the United States for mulching shrubbery
where it is not desirable to incorporate
the organic residues in the soil. They
have also been used to some extent in
vegetable crops and flower gardens to
conserve moisture and to prevent the
soil structure from being destroyed by
beating rains.
be used, but grease and meat scraps will
develop offensive odors and attract rats
and flies.
During composting a considerable part of
the materials decay and part of them go
into the air as gases. The remainder
is similar to well -rotted stable manure.
A good manure from composts must go
through this decomposition process.
Bacteria and molds are largely responsi-
ble for decomposition. Although they de-
rive energy from the refuse which they
decompose, they also need other sub-
stances for growth. One of the most es-
sential is nitrogen. Because most of
the substances which go into a compost
heap are low in this element, it is nec-
essary to furnish additional nitrogen if
a rapid rate of decay is to be obtained.
In the process of decomposition, acids
are produced. Some nitrogen fertilizers
also form acids. Consequently, it is ad-
vantageous to add limestone.
One advantage of decomposing organic ma,t-
ter is lost when composts are prepared
in a compost heap. One reason for in-
corporating organic m.atter in the soil
is to improve soil structure or tilth.
As the organic materials decompose, the
microbes produce substances that in-
crease the aggregation or crumb struc-
ture of the soil. Although soil tilth
is not improved solely by microbial
activity, it is recognized generally
that microbes have an important function
in keeping soils in good physical condi-
tion.
Many materials can be used in the com-
post heap, including leaves from trees
and shrubs, weeds, la^-m clippings, and
garden residues. Even table scraps can
A satisfactory compost may
by using the following:
be prepared
Organic residues (dry weight) 100 pounds
Ammonium sulfate 5 pounds
Limestone 5 pounds
Five hundred pounds of green plant resi-
dues will contain about 100 pounds of
dry material. A bushel of leaves weighs
about 5 to 8 pounds, depending upon con-
dition and packing.
To build a compost heap, place about 1/5
to 1/6 of the leaves, grass clippings,
or garden residues in a layer 10 to 12
inches thick. Moisten this layer and
spread I/5 to 1/6 of the mineral mixture
over it. Alternate the layers of residues
and minerals until the stack is complete.
Make the top of the stack concave to
catch and hold rain. If the material is
kept moist, rotting will proceed rapidly.
One ton of dry material will produce
about 2 1/2 tons of moist artificial ma-
nure.
If the heap is made in early or even
late summer, the compost will "be avail-
able for early spring use. However, if
the materials are not composted until
late fall, when the temperatures are low,
decay will be slow, and a sufficient de-
gree c£ decomposition may not be obtained
by early spring.
Although it is not necessary to add phos-
phates to decompose the residues, 2
pounds of superphosphate added to the
mineral mixture will increase the value
of the manure, particularly if it is to
be used on soils that are low in avail-
able phosphorus.
Even though artificial manure in the
stack does not have a noticeable odor
during decomposition, when spread it has
an odor resembling that of natural ma-
nures.
Prepared mineral niixtures are available
in some localities. More recently a num-
ber of commercial products which are
said to contain the microbes needed for
decomposition have been sold. Investi-
gations in California and Florida indi-
cate that there is little need to apply
microbial concoctions. The same results
may be secured by adding small amovmts
of fertile soil to the coiirpost.
0. H. Sears
i<-2-56
.^IIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
AND
TESTING
m
>
■
SF-44
THE NATURE OF EXCHANGEABLE CALCIUM AND MAGNESIUM AND
THEIR RELATION TO SOIL ACIDITY AND LIME REQUIREMENT
In moist soils calcium and magnesium are
present mainly in exchangeable form.
This form is attached by valence bonds
to the surfaces of the soil clay and or-
ganic matter particles. Exchangeable
potassium also occurs on these siirfaces,
although usually only in small amounts
in relation to calcium and magnesium.
The exchangeable form is usually the
principal available form of each of these
soil nutrients --the form the plant de-
pends on for grovth. ^^
However^ calcium and magnesium play an-
other role in soil fertility. They
control the acidity or pH of the soil.
If the clay and organic matter particles
are covered mostly with calcium and mag-
nesium;, the soil will be nearly neutral
or "sweet." But whenever a cation like
the calcium or magnesium ion is lost by
leaching or removed by plant roots, hy-
drogen, the "acid" ion takes its place
and the soil becomes more acid. Liming
is merely the process of replacing the
exchangeable calcium and magnesium lost
by leaching in order to bring the soil
back to a pH favorable for plant growth.
In the claypan soils of southern Illi-
nois, the native soil is usually highly
acid. The calcium and sometimes the mag-
nesiiim are so low that liming is needed
for two purposes: First, liming adds
calcium and magnesi'um and thus overcomes
deficiencies to the extent that these
nutrients are supplied in available form.
Second, proper liming corrects the acid-
ity sufficiently to make the soil reac-
tion (pH) favorable for plant growth.
In one sense the two fimctions of liming
are completely independent of each other.
In fact, except in southern Illinois ,
most of the soils in the state already
contain so much exchangeable calcium and
magnesi'um that they provide plenty of
nutrient for plant growth, yet they are
often so low in these elements in rela-
tion to the acid ion that the soil can
be harmfully acid to some crops. Except
in these claypan soils, therefore, the
chief purpose of liming is usually to
change the soil reaction.
One effect of changing the pH upward is
that it makes it possible for plants to
feed more efficiently on the available
phosphorus in the soil. Another is that
many legumes nodulate only at the higher
pH levels. (A low pH means a more acid
reaction. ) Organic matter appears to
decompose more readily at the higher pH
levels.
Except in soils that are very low in
clay and organic matter, a favorable pH
level is practically a sure sign that
calcium will not be deficient. In such
soils a high pH could theoretically be
due mostly to potassium and magnesium.
But such situations have never been found
in Illinois.
The pH of a soil is not determined by
how much or how little exchangeable cal-
cium and magnesium it contains, but
rather by the proportion of these basic
ions to the acid hydrogen ions. The to-
tal amount of exchangeable cations Ca++,
iy[g++^ and H+ held on the clay surfaces
is called the base exchange capacity and
is measured in terms of milligram-
equivalents per 100 grams of soil (m.e.
per 100 grams).
One m.e. per 100 grams is about the
amount of calcium and magnesium contained
in 1,000 pounds of high-grade lime-
stone. If a soil contains on its clay
and organic matter surfaces 8 m.e. of
bases and 2 m.e. of acid ion, or a total
of 10 m.e., it will be in the sweet
range because it is 80 percent saturated
with exchangeable bases and only 20 per-
cent saturated with the acid ion. If it
contains l6 m.e. of bases and ^4- m. e. of
acid^ it will have a total of 20 m.e.,
but the percent of saturation and also
the pH will be the same as before. So
the degree of acidity or pH depends not
on the actual amount of acid that is
present; but rather on the proportion of
acid to base.
Lime requirement is another matter. Sup-
pose the first soil has 5 m.e. of bases
and 5 m.e. of acid, for a total of 10
m.e. It is then 50 percent saturated
with bases and is in the unfavorable
range of acidity. A more favorable range
is 80 to 90 percent saturation with
bases. This means that from 3 "to ^ m.e.
of hydrogen must be neutralized. This is
equal to 3^000 to ^,000 pounds of pure
limestone. Hence the tentative lime re-
quirement is 3^000 to ^,000 pounds.
Wow suppose a 20 m.e. soil is also only
50 percent saturated and it is desirable
to increase the saturation to around 80
to 90 percent. Fifty percent of 20 m.e.
is 10 m.e. of the bases present. Eighty
to 90 percent saturation is 16 to I8
m.e. of bases needed. If 10 m.e. are
present and I6 to I8 are needed, then 6
to 8 m.e. or 6,000 to 8,000 pounds of
limestone is the tentative lime require-
ment.
Silt loam and clay soils have higher
base requirements for the same pH value
than do the lighter soils. Sands gener-
ally have the lowest.
Theoretically, in order to estimate a
practical lime requirement, one should
first know the magnitude of the base-
exchange capacity and the sum of the mil-
liequivalents of the bases present. This
would require a quantitative estimate of
the exchangeable calcium, magnesium, and
hydrogen (acid ion).
But that is not all one would have to
know. Knowing that, theoretically, it
would take 6,000 pounds of pure lime-
stone to sufficiently neutralize the
soil is only one item. Next one must
consider the neutralizing value of the
limestone. If it is only 90 percent C.C.E.
(calcium carbonate equivalent ), the orig-
inal 6,000-pound calculation must be
increased to compensate for the 10 per-
cent impurities. This increases the
requirement to 6^666 pounds.
However, this is still not the answer.
The fineness of the limestone must be
rated according to its rate of neutrali-
zation, i.e., how much acid it can neu-
tralize within a relatively short time.
For years, until a government agency took
over limestone inspection, such ratings
were made on all Illinois limestones and
an effectiveness score was given to the
stone. If the effectiveness score is low,
more limestone is needed. Wow suppose
the limestone has a score of 85 percent
effectiveness over the first three years.
This was about the average score for Il-
linois quarries at the time effectiveness
inspection was discontinued. Applying this
correction to the above 6,666 pounds gives
7,840 pounds as the corrected requirement.
But again this is not the whole story.
Many, if not most, soils have acid sub-
soils. But the acidity of the subsur-
face varies; in the claypan soils it is
higher than in the dark-colored silt
loams. With deeper plowing this acid
soil gets mixed with the top soil, in-
creasing the acidity. Also the transfer
of bases downward increases when sub-
soils are more acid. These things would
all affect the need for limestone over a
period of years.
And, last, there is the matter of quick-
ness of leaching. A sandy soil with a
sufficiently low exchange capacity can
become acid through the loss of only one
-3-
milliequivalent of calcium. And more
water moving through such a soil hastens
leaching. But heavy soils change in pH
much more slowly because their capacity
is higher and less water goes out the
tile. The limestone reconmiendation must
include recognition of this point.
So estimating a practical lime require-
ment is not merely a matter of knowing
the amounts of exchangeable bases and acid
present^ although this is the soundest
starting point for estimating the theoret-
ical requirement. Other considerations
are the effectiveness of the limestone
(fineness)^ purity of the limestone,
acidity of the subsurface, and rate of
leaching of the exchangeable bases. When
all of these things are considered, the
final recommendation may appear to have
little relation to the theoretical re-
quirement .
For example, some sands in Illinois have
a base -exchange capacity of 3 milli-
equivalents per 100 grams. This means
that they can hold calcium and magnesium
equivalent to not much over 2., 500 pounds
of a fine 100 percent C.C.E. stone. Yet
when they are acid, 3 "to 4 tons of ordi-
nary limestone are commonly added after
considering screen score, C.C.E. , and
especially the high rate of leaching on
sandy soils. On the other hand, a soil
like Muscatine silt loam with a base-
exchange capacity of 20 m.e. per 100 grams
will, at the same pH, have a much higher
theoretical requirement, but the practical
requirement may be little higher than the
practical requirement for the sand when
all the other factors are considered.
It is therefore not "fool proof" to base
the lime requirement on measurement of
the exchangeable bases and the exchange-
able hydrogen (acid) alone.
The lime requirement recommendations
based on pH or thiocyanate readings have
resulted from practical experience over
a period of 30 or more years. VJhile a
thorough study of Illinois soil types
and of their base-exchange capacities and
pH — degree of saturation relationships
was made a couple of decades ago, it has
not yet appeared practical to apply this
knowledge to refining the lime require-
ment recommendations made in our county
laboratories.
The Missouri "Lime Meter" method repre-
sents one approach to this problem, but
when corrected for these other factors
it correlates too closely with our recom-
mendations based on pH or thiocyanate to
make it practical to use.
The pH and thiocyanate methods, while
they do not involve a determination of
the bases and acid, do give a measure of
degree of saturation with bases and can
be Interpreted in terms of lime require -
m.ent when the base -exchange capacities
of the soil types are known.
In general, the high-capacity soils occur
in the northern two-thirds of Illinois,
and their capacities and pH vs. degree
of saturation relationships have been
thoroughly studied and found sufficiently
similar to make it practical to use pH or
thiocyanate as a measure of the lime
requirement.
On the other hand, the soils of southern
Illinois, while lower in base-exchange
capacity^ have much more acid subsoils,
which would tend to increase the rate at
which a limed soil becomes acid and thus
require more than the theoretical amount
of lime.
Measuring the theoretical amount is there-
fore not the whole answer. The higher
cost of the required tests leads to a
tendency to run only one or two samples,
whereas in the Illinois plan 11 samples
in a UO-acre field are analyzed. Given
a choice, the 11 samples run by the pH or
thiocyanate test are generally preferable
and may be more accurate than a single
composite of a whole field run by a the-
oretically better method. In fact, some
results reported to us of interpretations
-1^-
based on theoretically better determina-
tions indicate that the interpretations
must have oeen made "by persons mth so
little knovledge of soil chenistry that
the recommendations -were unsoimd and
vorthless.
Years ago this station put out a very
siniple titration-indicator method that
involved using several samples of the
same soil and shaking each one •'.■ri.th a
salt containing a different amount of a
base. This procedixre measured the theo-
retical lime requirement directly^ re-
gardless of variations in base-exchange
capacity, and is easier to run than
methods involving the total of each ion
involved.
Roger H. Bray
6-11-56
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
AGRONOMY FACTS
MANAGEM
CONSERVATIO.
SM-12
OBJECTIVES OF CROP ROTATIONS - INTRODUCTION AND EROSION CONTROL
This is the first in a series of five
Agroncmy Fact sheets devoted to the "Ob-
jectives of Crop Rotations." This one
is concerned with control of soil ero-
sion. The second one ■will discuss the
effects of crop rotations on soil physi-
cal properties^ such as absorption of
rainfall;, internal drainage^ and compac-
tion. Nvunber 3 will explain how crop
rotations may help to control insect
pests and crop diseases. Number h -vrill
take up the problem of plant nutrients
with particular emphasis on nitrogen.
The fifth and last of this series will
discuss the economic factors to be con-
sidered in deciding what crop rotation
to follow.
Crop Rotations and Erosion Control
The effectiveness of a crop rotation in
controlling soil erosion depends upon
the type of growth^ the amount of growth
and the time of growth of the crops in-
cluded in the rotation and the propor-
tion of the time that the crops with
different characteristics are on the
land.
Type of Growth
Grasses and legumes are erosion resist-
ing crops. If a good stand of such
crops occupies the land^ erosion losses
are negligible even on relatively steep
slopes. At the other extreme is bare
soil which is subject to serious erosion
even on gentle slopes. Inter-tilled
crops, like corn and soybeans, furnish
some protection against erosion and are
therefore better for the soil than no
crop. Small grains are intermediate be-
tween grasses and legumes and inter-
tilled crops in holding down erosion.
Amount of Growth
In the case of any crop the amount of
top growth determines to a considerable
extent its effectiveness in controlling
erosion. The more completely the soil
is covered by a crop the less it is
exposed to the beating action of rain-
drops which break up the soil crumbs or
graniiles. The character and extent of
the root system of a crop are also impor-
tant. For example, in August a crop of
soybeans ma,y cover the ground as com-
pletely as a crop of alfalfa and still
the soil may be subject to much greater
erosion losses . This is due to the fact
that the bean ground is in a looser con-
dition than the alfalfa ground. The
bean roots are also less extensive and
are usually localized in rows.
Abundant root growth of crops improves
the physical condition of soils so that
water is more readily absorbed and the
soil is less damaged by the beating
action of raindrops. A soil full of
roots also resists the cutting action of
water concentrated into small streams.
Time of Growth
The amount of erosion that m.ay be attri-
buted to a particular crop depends
largely on the time of its groirth. In
Illinois much of the corn is planted in
May, when the amount and intensity of
rainfall is relatively high. Since the
soil is loose from tillage and the soil
gets no protection from, the crop^ erosion
losses are usually high. In June the
corn is in the active growing stage, the
soil is cultivated and the amount and
the intensity of rainfall are relatively
high. In this month also erosion losses
are high. From kO to 60 percent of the
annual soil losses from corn land on the
Agronomy South Farm occurred in June.
Expected Soil Losses
From available data it is possible ^- -r-
timate the soil loss by erosio:. -, u
will occur on different slopes, from
different kinds of soils under a particu-
lar rotation. If the sril loss is
greater than the estir. ermissable
loss, then more sc;^ .;-- be included
in the rotation. .' in Illinois
-2-
farmers are interested in growing corn
and soybeans. To figure out a crop rota-
tion that will satisfactorily contx'ol
erosion just enough sod crops are included
to keep the soil loss dovn to a per-
missahle amount. Permissable loss is
the amount in tons per acre which it is
estimated can "be lost annually from a
particular soil and still maintain the
land in continued productivity. (The
table appearing at the end of this re-
port shows how this works . )
Supporting Practices
By using supporting practices a rotation
may he followed that would permit too
high soil loss without the supporting
practices. It works like thisl A
farmier selects the crops he wants to
grow and the amount of a particular crop^
like corn, he wants to include in the ro-
tation. This is checked for the condi-
tion of his land, the percent .of slope,
the length of slope, the kind of soil
and its condition, particularly with
reference to past erosion damage. If
the erosion losses are higher than per-
mis sable under the conditions, the use
of supporting practices, such as contour-
ing, strip cropping, and terracing may
reduce the soil loss enough so the de-
sired rotation can be followed.
Expected Soil Losses from Selected Rotations on Four
Soil Types on a 200-foot Contoured Slope in Illinois
a
:/
Rota.tion— '
CCCG*
CCCGM
CCGM
CGM
CGMM
CGMMI#I
Permis sable
Loss
Flanagan
2i Slope
Siv-ygert
5i
Clinton
epjo
Grant sbur
5fo
Tons /A
Tons/A
6.2
Tons/A
14.6
Tons/A
1.9
7.8
1.2
4.2
9.8
5.3
1.0
3.4
2.0
7.8
4.5
4.2
.6
2.4
.k
1.2
2.8
1.5
.2
0.8
1.8
1.0
4.5
1.5
3.0
2.5
(Losses below heavy line are small enough to permit use of
rotation indicated. )
a/ Calculations are based on expected losses on soils with 8 or
more inches of siirface soil remaining. Average management is
assumed in preparing the table. With poor management the losses
should be multiplied by 1.3. For high-level management of crops
and soil multiply by 0.7.
b/ C - corn; G - small grain; M - meadow. *G - small grain with
catch crop to plow down.
C. A. Van Doren & R. S. Stauffer
1-30-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
MANAGEMENT &
CONSERVATION
EFFECT OF CROP ROTATIONS ON SOIL PHYSICAL CONDITION
SM-13
The physical condition of soils is de-
termined largely by their texture^ state
of compaction, and degree of aggregation.
Cultural practices have little effect on
soil texture, hut such management vari-
ables as organic matter management, sur-
face cover, and tillage will change soil
compaction and aggregation and thus help
to determine physical condition.
Organic matter management . The degree
of aggregation is closely related to the
amount of readily decomposable organic
matter in the soil. For this reason,
cropping systems that add large amounts
of readily decomposable organic matter
to soils are the systems that promote
good aggregation and thus improve physi-
cal condition. The effectiveness of
residues in increasing organic matter in
the soil, and thus improving soil aggre-
gation, depends on the amount of resi-
dues, their ability to decompose, and
the thoroughness with which they are
mixed with the soil.
If other conditions are constant, crop
rotations that add the greatest amount
of readily decomposable crop residues to
the soil will be most effective in cre-
ating good physical condition.
Surface cover. The amount of protection
provided by the crops grown in the
rotation also affects soil physical con-
dition. Cropping systems that provide
a protective vegetative cover, either
living or dead, for the soil during the
greatest part of the year help to lessen
the impact of raindrops. Heavy rains
destroy aggregation in the surface soil
and often cause serious crusting. Air
and water cannot then move freely into
the soil, and seedlings may not be able
to emerge.
Tillage . Tilling tends to cause soil
physical condition to deteriorate, par-
ticularly when the soil is wet. Crop
rotations differ greatly in the amount
of tillage they require. Intertilled
crops have a greater structure -depleting
effect than others.
Sod crops provide readily decomposable
organic matter and vegetative cover and
require fewer tillage operations than
intertilled crops. It is thus easy to
understand how the sod crops help to im-
prove soil structure. Possibly some of
the new techniques now being used to
produce intertilled crops may eventually
reduce their structure -depleting effect
enough to make it necessary to reevaluate
their usefulness in crop rotations for
improving soil physical structure .
M. B. Russell
2-13-56
JNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
MANAGEMENT &
CONSERVATION
SM-14
CROP ROTATIONS AND INSECTS
Cultural practices are essential in com-
bating attacks of many species of in-
sects. Crop rotations are a cultural
practice that reduces the number of in-
sects farmers have to combat and thus
cuts down the amount of insecticide they
have to use. Insecticides are only a
supplementary means of controlling in-
sects. If populations were not reduced
by rotations and other cultural prac-
tices^ as well as by the weather, the
insecticide bill on each farm would be
astounding.
Northern corn rootworm is a good example
of how insects can be controlled by cor-
rect crop rotation. This pest, which
has a life cycle of one year, presents a
serious problem where corn is grown in
the same field for more than two years
in succession. The rootworm overwinters
in the egg stage, and the small larvae
hatch in the spring and stay in the soil
waiting to feed on the corn roots. The
damage they do may cause the corn to
lodge seriously.
Correct rotations will adequately con-
trol this pest. Growing other crops in
the rotation for two years after corn
reduces the niitnber of larvae and emerg-
ing adults, providing adequate control
without use of an insecticide. A two-
year break is necessary, however, be-
cause rootworms will survive for one
year on some crops besides corn. ^-Jheat
jointworms can also be partly controlled
by rotating crops.
White grubs are another pest that can be
controlled by rotations, but not so eas-
ily as the northern corn rootworm. Be-
cause grubs have a three -year life cycle,
rotation plans must be made two years
before the damage is expected. In the
first year of the cycle, June beetles,
the adults of white grubs, lay their eggs
in the spring. The grubs hatch, feed,
and overwinter. The next summer, during
the second year of the cycle, they feed
heavily, severely damaging the crop
roots. During the third year, they feed
during May, pupate, and remain in the
soil as adults the following winter,
emerging in the spring to lay eggs.
To avoid damage from grubs during the
year of heavy feeding, it is necessary
to have had some crop on the land that
was not suitable to the adults for lay-
ing eggs during the previous season (the
first year of the cycle). Such crops
are clover, alfalfa, and corn. At one
time soybeans were included in the list
of resistant crops, but there is now one
species of grub that will lay eggs in
this crop.
A carefully planned rotation will also
reduce some species of wireworms and
cutworms. If length of life cycle and
other facts about the insect are known,
then a crop rotation can be planned that
should reduce its attacks on the crop.
H, B. Petty
2-20-56
(Over )
SM-14
CROP ROTATIONS AND DISEASE
It is obvious that continuous cropping
perpetuates and increases the pathogenic
organisms in the soil. Good examples of
this process in Illinois are the in-
crease in the amount of take-all and of
bunt through soil infestation in fields
where wheat has followed wheat for two
successive years. Under such situations
rotating crops would constitute a good
method of control. Crop rotation may^
however, be recommended as a means of
controlling disease without considering
whether the pathogen or pathogens would
be effectively reduced.
The first question to consider in rec-
ommending crop rotation as a means of
controlling a specific disease is to de-
termine whether the pathogen that is re-
sponsible for the disease is a true soil
inhabitant or whether it is a soil in-
vader of short duration. Some organisms
are capable of living in the soil for 10
to 20 years without coming into contact
with their natural host. Some examples
are species of Fusaria and damping-off
and root-rotting fungi like Pythium.
Crop rotation could obviously be of lit-
tle value in controlling diseases caused
by this group because it would take too
long to eliminate them.
may be at least partly eliminated by ro-
tating crops. Most of these fungi, how-
ever, are parasites of vegetable crops.
Some of the more common ones are the or-
ganisms causing bean anthracnose, black-
leg of cabbage, and bacterial blight of
common beans.
Some work done in Illinois to determine
the persistence of the soil-borne wheat
mosaic virus showed that growing nonsus-
ceptible crops for at least four years
reduced the amount of virus but did not
eliminate it. Therefore crop rotation
cannot be recommended as an adequate con-
trol for this disease, in which the virus
is known to persist in the soil for a
long time.
In most cases rotation is of little value
as the sole means of controlling soil-
borne diseases in agronomic crops. On
the other hand, crop rotation together
with other types of control could be
very effective. Often the two combined
will keep the disease-producing organisms
from building up or hold them to a mini-
mum so that they will cause little loss
in yield. If they accomplished no other
purpose, this result alone would make
crop rotation worth while.
On the other hand, those fungi that can
live for no more than two to four years
in the soil without their natural host
Wayne M. Bever
Department of Plant Pathology
2-20-56
( Over )
UNIVERSITY OF ILLINOIS ■ COLLEGE OF AGRICULTURE
AGRONOMY FACTS
MANAGEMENT &
CONSERVATION
SM-15
AN ANALYSIS OF THE NITROGEN STATUS OF THE AGRONOMY SOUTH FARM ROTATIONS
Numerous experiments ha"ve shoi-m that sat-
isfactory^ although not necessarily maxi-
mum, grain yields can be maintained by
using rotations where the legume -grain
crop ratio is 1 to 3 or 1 to ^4-. The
marked improvement in soil physical con-
dition (tilth) caused by the legume mead-
ow and the noticeable deterioration in
physical condition after several years
of tillage have tended, however, to ob-
scure the primary fiinction of legumes in
rotations, namely, to restore and main-
tain nitrogen. It is only when nitrogen
is applied to grain crops several years
removed from the legume that it becomes
evident that the decline in crop yields
which necessitates a return to legumes
is primarily a reflection of lower ni-
trogen fertility rather than of tilth
deterioration.
The transitory nature of nitrogen fer-
tility in rotations where legumes are the
primary source of nitrogen is illustrated
in Figure 1. The corn, corn, oat, wheat
rotation was planned to determine the
effects of catch-legxanes sown in small
grains en subsequent grain yields . There
were check plots where legumes were not
seeded and plots where legumes were
seeded. Thus it is possible to evaluate
the yield effects of seeding the catch
legumes.
Figure 1 shows that the catch legume in-
creased yields of all grains. The yield
effect of the legume, however, expressed
as percent of increase in yield, was al-
ways greatest the first year. A rather
sharp break in the residual yield effect
of the legume is evident in the second
year. The yield effect of the legume
continues to decline thereafter, but at
a lesser rate.
The data in Figure 1 for the yield ef-
fects of commercial nitrogen indicate
that commercial nitrogen is equally as
50
0)
o
c
0)
30"
so-
lo
-^
For
nitrogen
^
For catih legume
Corn
Corn
Oats
Wheat
Fig, 1, Percent of Increase In grain yields
for planting a catch legume and using nitrogen
only where yields of plots receiving no nitro-
gen or legume seedlngs are the standard for
comparison. For nitrogen graph line only:
first corn, 100 lb. N; second corn, no N; oats,
20 lb, N; wheat, 20 lb. N. (Data from Illinois
Agronomy South Farm)
satisfactory as legume nitrogen in so far
as first-year corn yields are concerned.
The residual effects of commercial nitro-
gen applied to first-year corn on second-
year corn yields were insignificant. In
this respect legume nitrogen was superior
to commercial nitrogen. It is quite ap-
parent, however, that the leg-ume system
did not supply enough available nitrogen
for maximum small grain yields, because
oat and wheat yields were higher when
nitrogen was applied.
The preceding discussion presented
evidence of the transitory nature of
nitrogen fertility. Figure 2 gives a
generalized concept of the nitrogen fer-
tility pattern one might expect in rota-
tions where the ratio of non-legume to
legume crops is 3 to 1. The pattern is
carried through nine years, or two full
rotation periods, to show the saw-toothed
cyclic nature of nitrogen fertility in-
duced into rotations by periodic legume
growth.
-2-
4J
\
\
/
■H
H
-p
\
\l
/
0)
c
\
\
/
bO
O
U
■\^
X
/
B
Years
1234567 89
Non-legume crops Non-leguKe crops
Legume meadow Legume meadow
Fig, 2, A generalized nitrogen fertility pat-
tern for rotations where the grain to legume
ratio Is 3 to 1.
Two areas in the nitrogen fertility pat-
tern are of interest : the peak and the
floor. In terms of minimum rotational
nitrogen availability^ the floor can be
expected to vary with the organic matter
content of soils ^ the activity of decay
of this organic matter, and the residual
legume nitrogen. If the legume is not
sown or if it fails, there can be no
peak. The floor of minimal nitrogen
availability would then appearto broaden
and gradually sink, since nitrogen avail-
ability at this point depends primarily
on the decay rate of an ever-decreasing
soil organic matter supply.
If the stand is pure, the height to which
nitrogen availability or fertility rises
following a legume would appear to be
determined primarily by the amount of
nitrogen the legume contributes. Lower
peaks, and frequently double peaks, with
the higher of the two peaks in the second
year, are possible where mixed legume -
grass sods or legumes mixed with a con-
siderable amount of straw are turned
under.
Figure 3 shows examples of multiple ni-
trogen availability peaks of this nature.
Thus the generalized nitrogen fertility
pattern illustrated in Figure 2 can vary
with different rotations. Moreover, the
peak nitrogen fertility associated with
a specific rotation may not represent
the optimum nitrogen fertility needed
for maximxim grain yields.
3_.0 _ Critical leaf nitrogen
„ "content {2.3fa N)
2.8
2.6
2.4
2.2
2.0
1.8
1.6
n
2345
Rotation No.
6 7 8 9 10
[See Table 1)
Fig, 3. Nitrogen adequacy peak measure by leaf
analysis for various rotations, Illinois Agron-
omy South Farm.
Leaf analysis studies with corn have in-
dicated that a leaf nitrogen content of
2.9 percent gives maximimi corn yields.
This is the critical leaf nitrogen per-
centage for corn. By means of leaf analy-
sis it has been possible, in terms of
leaf nitrogen contents, to define the ni-
trogen fertility peaks associated with
various rotations at the Agronomy South
Farm, Ten rotations and associated corn
leaf nitrogen contents are listed in
Table 1. The results are presented
graphically in Figure 3«
Table 1. Rotations and associated corn leaf
nitrogen contents at the University of Illinois
Agronomy South Farm*
Rotation
number Rotation
1 Corn, soybeans
2 Corn, oats
3 Corn, oats**
4 Corn, oats, wheat**
5 Corn, corn, wheat**
6 Corn, oats, wheat, timothy
7 Corn, oats, wheat, alfalfa
8 Corn, corn, oats**, wheat**
9 Corn, corn, soybeans, wheat**
10 Corn, soybeans, oats, wheat**
Percent
leaf nitrogen
2.42
2.17
2.67
2.67
1.80 & 1.70
2.14
2.85
1.82 &
2.07
1.94 & 2.07
2.05
*A11 plots Included In rotation experiment
have been limed and phosphated. Soil test
Indicates adequate potash,
**Catch legumes seeded.
-3-
Rotation 7? which is a corn, oat, wheat,
alfalfa sequence, was the only rotation
in which soil nitrogen fertility, as in-
dicated by leaf analysis, approached
that considered optjjnum for first-year
corn. Rotations 2, h. 5^ 6, 8, 9, and
10, judged by the same standard, gave
soil nitrogen fertility peaks that were
substantially suboptimum for maximum
corn yields.
The leaf nitrogen content observed for
rotation 1, a corn-soybean sequence, is
of considerable interest. Most workers
do not consider that harvested soybean
residues contribute m.uch nitrogen to the
soil. Yet, except in rotations 3 and J,
the leaf nitrogen content of corn fol-
lowing soybeans exceeds that for other
rotations.
The microbial demands for nitrogen set
up by corn residues probably have no ef-
fect on yields of nodulated soybeans.
Thus soil nitrogen perhaps contributes
toward the decay of most of the corn
residues diiring the season in which soy-
beans occupy the land. ' Corn following
soybeans would then appear to grow in a
soil environment that is virtually free
of microbial competition for nitrogen.
Thus the greater part of the available
nitrogen is released from soil organic
matter. This nitrogen plus that re-
leased from partly decayed corn and de-
caying soybean residues probably makes
the soil nitrogen fertility status higher
than one might expect. It should be ob-
vious, however, that this practice can
not assure above-average corn yields in-
definitely because under this highly
nitrogen-deficient rotation the soil or-
ganic matter will eventually be drastic-
ally reduced, leading to progressively
lower corn yields.
The effect of the kind of residues that
precede corn on net soil nitrogen avail-
ability can be evaluated by comparing
rotations 1 and 2. Where soybeans pre-
cede corn, microbial nitrogen demands
are probably at a minimum for the rea-
sons previously given. Where corn
follows oats (rotation 2), microbial com-
petition for available nitrogen appar-
ently persists during most of the growing
season. This is reflected in lower leaf
nitrogen contents and lower corn yields
for the corn-oat sequence. Planting a
catch legume in the oats (rotation 3 )
appears to greatly increase the net ni-
trogen availability, and this increase
is reflected in a higher leaf nitrogen
content.
Multiple nitrogen fertility peaks, with
maximiim nitrogen fertility in the second
year, are not uncommon for some rota-
tions. This is true for rotations 8 and
9. It would appear that the nitrogen
contributed by the catch legume that
preceded corn was not sufficient to over-
come all of the microbial nitrogen de-
mands occasioned by the decay of the
wheat straw. Thus first-year corn is
produced in a less favorable soil nitro-
gen availability environment.
The initial decay processes that reduce
first -year nitrogen availability, how-
ever, appear to be more or less complete
prior to August of the second year. The
subsequent release of available nitrogen
previously withheld during decay then
gives a higher second-year nitrogen fer-
tility pattern, as the data for corn
leaf composition show.
The nitrogen fertility pattern illus-
trated in Figure 2 might be considered
to represent a normal nitrogen fertility
pattern. The data in Figure 3^ however,
indicate that rotations can alter the
normal pattern, i.e., multiple nitrogen
peaks. Moreover, it is apparent that
the nitrogen fertility peaks achieved
by some rotations do not constitute ni-
trogen adequacy in so far as maxim'um
yields are concerned. Supplementary
nitrogen fertilization throughout such
rotations is needed to obtain maximum
grain yields.
Edward H, Tyner
2-27-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
MANAGEMENT &
CONSERVATION
AGRONOMY FACTS
SM-16
ECONOMIC OBJECTIVES OF CROP ROTATIONS
Four previous Agronomy Fact sheets have
discussed the various agronomic objec-
tives of crop rotations: controlling
erosion;, maintaining desirahle soil
physical properties^ controlling insects
and diseases, and supplying plant nutri-
ents to crops. Attaining any one of
these four objectives to the fullest ex-
tent is likely to mean sacrificing the
complete fulfillment of one or more of
the others. They must therefore he bal-
anced when a rotation is selected and
put into effect in the farm business.
An economic analysis of this problem
should consider the "best" use of labor
and capital, as well as the "best" use
of the land. In short, the economic ob-
jective of crop rotations should be to
select the rotation that will give the
maximum profit for the total farm busi-
ness over a period of years.
Economic Principle Involved. One crop
can be substituted for another to vary-
ing degrees in rotations. For example,
going from a C-C-0 (clover catch crop)
rotation to a C-C-O-M rotation means
that meadow is being substituted for
part of the corn and oats. In terms of
total production of each crop (per acre
yield times number of acres), the result
of substituting one crop for another in
a rotation is not always the same. For
example, the number of bushels of corn
sacrificed for each ton of legume hay
gained is likely to be less in shifting
from, say, a C-C-O-M to a C-C-0-M-M ro-
tation than in shifting from a C-O-M-M
to a C-0-M-M-M. This difference is due
to the different effects of crop se-
quences on yields as the proportion of
land in each crop changes.
The economic principle involved in se-
lecting a rotation is that, in order to
maximize profits, one crop (for example,
meadow) is substituted for another (for
example, corn) until the returns sacri-
ficed by decreasing corn production are
exactly balanced by the gain in returns
from increasing meadow.
Rotation experiments frequently show a
"complementary" effect on corn when leg-
\imes are added to the rotation in small
amoionts. That is, total corn production
(per acre yield times number of acres )
increases, up to a point, as legumes oc-
cupy a larger percent of the rotation.
If legumes and corn are complementary,
adding legumes to the rotation will be
profitable even if the roughage that is
produced is not sold or used by live-
stock. In m.any commercial fertilizer
programs, however, this complementary re-
lationship disappears, and corn and leg-
umes compete throughout a wider range
of rotations. This means that the farmer
has a larger ntmiber of choices, and
thus his decisions regarding cropping
systems become more difficult,
Basic Data Needed for Choosing Rotation.
The economic principle can be applied
by using expected yields from each alter-
native rotation. The expected yields
summarize, in a sense, the combined ef-
fect of the relationships described in
the four previous Agronomy Fact sheets
on rotation objectives. Since the prob-
lem of selecting a rotation cannot be
divorced from that of applying fertilizer
to individual crops within the rotation,
alternative fertilizer programs and
their expected effects on yield must be
considered in combination with each ro-
tation.
Except where a cash-grain system is to
be followed, the livestock system must
also be considered. Roughage -consimiing
livestock may be desirable to use labor
during slack seasons. Livestock may in
turn require a higher percent of leg-
umes in the rotation than would be dic-
tated by a simple cost-returns analysis
of the cropping system by itself.
For example, a cost-returns analysis of
the cropping system made independently
of the livestock system might show a
catch-crop rotation to "be more profit-
atile than a stand-over rotation. If,
however, the farm is small and the farmer
wishes to increase his vol'ume of busi-
ness by using roughage-consuming live-
stock, the stand-over rotation will be
more appropriate.
The place of manure and its effect on
yields also needs to be taken into ac-
count. And the effect of substituting
supporting practices for meadow to sat-
isfy soil conservation objectives must
be estimated.
Comparing Relative Profits of Rotations.
Using the basic data outlined above, the
farmer can make a "budget" of estimated
costs and returns for each alternative
rotation, including other parts of the
farm business that are related to the
rotation. This comparison of costs and
returns can be simplified by considering
only those costs that differ among the
alternative plans. For example, "such
fixed costs as taxes, interest on land,
overhead on machinery, etc . , can be
omitted because they stay the same re-
gardless of rotation. Operator and fam-
ily labor can also sometimes be omitted.
Since all costs are not included, the
resulting figures should not be confused
with "profits . " This budget is simply a
tool to help compare the effects of dif-
ferent rotations; it is not a measure of
profitability of the farm business.
Legume Nitrogen vs. Commercial Nitrogen.
An added advantage in selecting a rota-
tion as part of the total farm business
is that there is less need for "internal"
accounting. We need not, for example,
attempt to calculate a cost for produc-
ing nitrogen from legumes. So long as
the yield estimates for each rotation
adequately reflect the effect of the
crop sequence, and varying levels of ni-
trogen are considered in combination
with the rotations, the value of the leg-
ume will show up in subsequent yield
increases and, in a livestock system, in
increased livestock production. In-
terest should focus on the comparative
returns from the alternative total farm
plans and not on the value of " goods- in -
process. "
Computing Met Returns for a Period of
Years. Adoption of a rotation implies
that the farm operator expects to re-
ceive income and to inctir expenses over
a period of years. Alternative cropping
systems may give him a choice in the way
his income and expenses are spread over
this period. For example, some cropping
systems may require heavy initial ex-
penses for fertilizer, while others may
require smaller initial expense but
cover a longer time. The time at which
the crop sequence will affect yields also
differs among rotations. The farmer must
consider these differences in developing
a budget that is designed to help him
pick a rotation.
Budgets of returns and expenses over a
period of years can be compared more ac-
curately by using a discounting proce-
dure. This will give the net returns
over a period of time from each plan in
terms of its present value. For in-
stance, using an annual discount rate of
5 percent, we find that an income of
$100 three years from now is worth only
$86 today. The same procedure must be
applied to costs. Discounting is espe-
cially important in comparing alterna-
tive plans that have widely different
timing of expenses of income.
Comparative Risks. Some farmers prefer
a lower, more stable income to one that
is higher but varies more from year to
year. Since weather and price fluctua-
tions do not affect all crops in the same
way, rotations differ in variability of
returns as well as in average level of
returns. In the final selection of a
rotation, these differences in risks
need to be balanced against the compara-
tive returns from the alternative rota-
tions.
Earl R. Swans on
3-5-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
MANAGEMENT &
CONSERVATION
CONTINUOUS CORN
SM-17
How often should corn be grown on (pro-
ductive) corn-belt soil? This much-
discussed question has been studied
experimentally for 80 years on the Morrow
plots on the campus of the University of
Illinois. Many other tests have been
made on experiment fields and on farmers'
fields throughout the state.
At the Momence soil experiment field in
Kankakee county;, corn was grown continu-
ously for 1^ years, beginning in 1902.
Average yields were 6.5 bushels an acre
on untreated land; 4ij-.3 bushels with
limestone, phosphate, and potash (LPK);
and 50«6 bushels with limestone, phos-
phate, potash, and nitrogen (lIPKW).
Fertilizers were applied at the following
acre rates each year: phosphorus as
bone meal, 200 pounds; potassium as mu-
riate of potash, 150 poimds; and nitro-
gen as dried blood (l2foNO), 8OO pounds.
The soil was peaty loam.
On the Davenport plots at Urbana, a con-
tinuous corn experiment was operated for
13 years, beginning in I9OI. Fertilizers
were used in the same amounts as described
above in the tests at Momence. Average
yields were as follows: LPK, 50.1;
and LPKN, 6l.h. During these same years
(1901, 1913) J a, rotation of corn, oats,
clover, and wheat catch crop was also
used on the Davenport plots. Average
yields were 57«9 bushels with no soil
treatment and 78.7 bushels with LPK. No
nitrogen fertilizer was used on the rota-
tion plots. The soil on this area was
Flanagan silt loam.
A continuous corn culture study was
started on the Dixon soil experiment
field in 1932. To maintain a high test
level, soil treatments have included
lime, superphosphate, and potash. Since
19^2, 600 pounds of ammonium sulphate an
acre have been plowed under each year,
and 130 pounds of 3-12-12 have been used
as a starter. Rye has been seeded in
the fall as a cover crop since 1951-
Twelve-year average corn yields 0-9^^-55)
for these plots and for corn under
rotation at Dixon and at the Morrow
plots are reported in Table 1. Growing
costs are indicated in terms of bushels
of corn (calculated at $1.25 a bushel).
These costs were subtracted from yield
to give "take-home" corn per acre .
During recent years hybrid corn, cheaper
nitrogen fertilizers, and corn price sup-
ports have caused renewed interest in
continuous corn. At the Lebanon experi-
ment field on Jarvis-LeClaire silt loam
and at the Newton experiment field on
Cisne silt loam, tests with continuous
corn were established in 1951 on land
that had been in rotation with soil
treatment for many years. Treatments
with continuous corn include the use of
non-legume cover crops and nitrogen at
the rate of 1^0 pounds each year. Yields
were low as a result of severe drought
in 195^ and moderate drought in 1955 •
Table 2 gives average corn yields for
the four years 1952-1955 with continuous
corn and with corn in a corn, beans,
wheat, hay rotation.
At the Urbana Mumford (M-9) plots an
area of land has been in continuous corn
culture since 1935* Average yield on
the untreated areas of these plots from
19^0 to 1952 was 6h bushels an acre.
Table 3 lists treatments and yields for
the past three years (1953-1955 )• This
soil is a Drummer clay loam that has
been adequately limed and phosphated.
A. L. Lang
L. B. Miller
C. H. Farnham
P. E. Johnson
D. L. Mulvaney
3-26-56
Table 1. Continuous Corn Versus Rotation Corn at Dixon and at Morro-w Plots
12-yr. Averages^ 19ij-i4-1955
Costs in
bushels
Ratio-/ of
of corn per acre
"Take-
Soil
Acre
Soil
Total /
costs-
home"
yield to
Rotation
treatment
yield
treatment
corn
"Take -home"
Dixon Soil
Experiment Field - Tama
Silt Loam
bu.
bu.
bu.
bu.
Continuous corn
LPKN
3-12-
12
75.9
18.0
58.2
17.7
4.29
Continuous corn
L
^9.2
1.0
38.6
10.6
4.64
C-O-Cl-W
LPK
95.1
5.0
i^7.0
48.1
1.98
C-O-Cl-W
0
58.5
-
38.5
20.0
2.92
C-0-W( legume)
LPK
80.2
6.0
k6.6
33.6
2.39
C-O-W
LPK
^6.0
k.o
i^l.3
4.7
9.79
Morrow
Plots - Flanagan Silt
Loam
Continuous cox-n
0
22.0
M
39.0
-17.0
X
Continuous corn
MIP
64.0
5.0
41^.0
20.0
3.20
C-O-Cl
0
68.0
39.4
28.6
2.38
C-O-Cl
I/ELP
108.0
5.0
48.2
59.8
1.81
1/ Includes fertilizer costs, harvesting, and marketing @ 125z!/bu. plus all other
costs, based on Detailed Cost Report for Central Illinois. (A. E. Mimeo 2969)
2/ This ratio indicates how many bushels of corn must be harvested for each bushel
of net or profit. It also shows hov many bushels must go to market or be added
to surplus for the producer to maice the same net profit.
Table 2. Continuous Corn vs. Rotation Corn at Newton and
Lebanon, 1952-1955
Soil
Average corn yields
Cropping
treatment
Neirton
Lebanon
bu.
bu.
Corn continuous
LPK
35.7
61.0
Corn continuous and cover crop
LPK
30.1
54.3
Corn continuous
LPKN
35.1
63.0
Corn continuous and cover crop
LPKN
37.9
64.3
Corn, beans, wheat, hay
LPK
45.9
83.4
Table 3* Continuous Corn With and
Without Rye,
Mumford Plots,
M-9, Urbana
Year K Rye K
K Rye nV
10
-lo-ioV
Nl/
lo-io-ioi/
'17
bu. bu.
bu.
bu.
bu.
1953 57 66
1954 58 65
69
69
79
72
74
68
1955 59 71
87
75
88
90
Average 58 67
79
77
1/ Treatment 5OO lb. /A 10-10-10, N - 100 lb. N alone or 50 lb.
with 10-10-10.
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SP-9
THE PRODUCTIVITY OF SOME IMPORTANT SOUTHERN ILLINOIS SOILS
In the last few years a considerable
amo-unt of interest has centered on in-
creasing the yield possibilities of seme
of the southern Illinois soils. This
interest has arisen as a result of the
relatively higher prices received for
central and northern Illinois soils and
some rather favorable responses to soil
management on experimental fields in the
southern area.
Productivity is the ability of a soil to
produce crops or other plants under vari-
ous management practices and weather con-
ditions. Productivity may be measured
in per acre yields of certain crops or
in terms of an index of combined yields
of grain crops_, forage^ or timber (see
AG-141+3 ) .
The soils included in this report occur
generally south of a line from Calhoion
county, Illinois, on the vest to Clark
county on the east. On the soil associ-
ation map (see AG-l4^3) they occur in
Areas M, N, 0, and P. These soils have
generally developed in less than five
feet of loess lying on a weathered gla-
cial till of Illinoian age (SP-l). In
most cases they are leached of free car-
bonates, and their degree of base satu-
ration is low (SM-7).
Soils in Areas M, N, and P were gener-
ally developed under a native prairie
grass vegetation and are somewhat better
supplied with organic matter and with
the desirable base nutrients (Ca, Mg, K)
than are the soils of Area 0. The soils
Table 1. --Average Per Acre Yields of Hybrid Corn, Soybeans,
and VJheat Obtained by Farmers on Southern Illinois Soils
Under Medium and Moderately High Management Levels*
Hybrid
corn,
Soyb
2ans,
.Jheat,
Soil
Soil type
management**
manag
iiaent**
management**
assn.
or
Mod.
Mod.
Mod.
area
associated types
Med.
high
Med.
hiph
Med.
high
bu.
bu.
bu.
bu.
bu.
bu.
M
Herrick silt loam and
Virden silty clay loam
61
68
#
#
#
M-
tr
M
Herrick silt loam
5h
59
25
28
30
32
N
Ccwden silt loam and
Oconee silt loam
1+8
62
JL
.J.
25
27
N
Cowden silt loam
hi
56
#
7r
2k
26
P
Cisne silt loam
^5
53
22
30
21
21+
0
Bluford silt loam and
Ava silt loam
1+6
57
ir
#
19
20
*About 95 percent of data represent the I9I+O-5O decade.
**The standard error of estimate of these figures, when considered as 10-year
average yields, is calculated to be about + 5 percent.
//Insufficient data reported.
-2-
of Area 0 were generally developed under
forest vegetation and are usually quite
acid (pH 5»0) iJ^ their untreated state.
The yields presented in Table 1 are
hased on data reported by farmers who
have been cooperating with the Agronomy
Department in a state-wide project aimed
at evaluating the productivity of some
major Illinois soils. The calculated
yields^ based on farm-reported data^ are
tiiven for two levels of management. For
convenience, these management levels are
referred to as medium and moderately
high and are expressed in terms of pounds
of nitrogen, phosphate, and potash used
and the interval of time between legumes
and the crop under study.
For hybrid corn and soybeans, yields
were calculated by assuming 6.5 inches
of total rainfall and an average maximum
temperature of 90 for the months of
July and August. These figures are aver-
ages for the period I925-5O reported by
the weather stations in the area.
A medium level of management for esti-
mating hybrid corn yields consisted of
50 pounds of N per acre in the current
and the previous year, contributed from
both legume and nonlegume sources; 20
pounds each of equivalent PpOc and K„0
per acre, applied or estimated as resid-
ual from previous applications (30 pounds
for Bluford-Ava soils); and a legume or
legume -grass mijcture two years before
the corn crop.
Comparable figures for a moderately high
level of management were 100 pounds of W
per acre, ^l-O pounds of PpO^ (60 pounds
for Bluford-Ava soils), 60 pounds of K 0
{kO pounds for Herrick soils), and the
equivalent of an alfalfa — red clover mix-
ture immediately preceding the corn
crop.
For soybeans and wheat the comparable
figures for medium and moderately high
management are approximately one -half of
those given for corn.
Table 2 lists yields from similar crops
gro-ji/n on University of Illinois soil ex-
periment fields having the same soils as
those included in Table 1 or closely
associated soils. Huey silt loam listed
in Table 2 includes many of the so-
called slick spots in southern Illinois
(AG-li|i4-3 and SM-8).
The check-plot yields show the productiv-
ity of these soils in an untreated condi-
tion, and particularly the influence of
so-called slick spot soils. The combina-
tion of slick spots and low organic mat-
ter content in the soil surface may be
partly responsible for the low soybean
yields at the Sparta field. Good germi-
nation and early growth are frequently
poor because of "crusting" after rain.
The yields on the plots having full
treatment show how these soils respond
to adequate fertilization and appropri-
ate rotations. These yields are higher
than those reported for a moderately high
management level in Table 1. Farmers
whose long-time yields are similar to
those given in Table 1 may wisely examine
their crop production practices not only
with regard to soil management, but also
weed and disease control, choice of hy-
brids or varieties, planting rates, and
harvesting procedures. The adoption of
appropriate improved management prac-
tices should bring their soil up to such
a level that they could, within a reason-
able time, expect to obtain yields similar
to those shown in Table 2.
-3-
Table 2. --Average Per Acre Yields of Hybrid Corri;, Soybeans, and
Wheat Obtained on University Soil Experiment Fields,
19i|0-195^-^
Soil
Soil type
Hy.
corn.
Soyb
eans,
Wheat,
assn.
or
treatment
treatment
treatment
Experiment
area
associated types
None
Full**
None
Full**
None
Full-J«^
fields***
bu.
bu.
bu.
bu.
bu.
bu.
Clayton
M
Dominantly Herrick
Carlinville
silt loam
h9
90
23
32
1?
32
Lebanon
P
Cisne silt loam
23
73
11
21
5
27
Oblong
P
Cisne silt loam and
Hoyleton silt loam
18
62
8
21
2
25
Ewing
P
Cisne silt loam and
Huey silt loam
9
58
9
21
1
22
Nevton
0
Wynoose silt loam
and Huey silt loam
6
52
2
12
3
26
Sparta
*See 111. Bui. 516, Effect of Soil Treatment on Soil Productivity, for history of
fields and yields prior to 19^2.
**Full treatment represents an average of yields obtained on plots treated with
manure, limestone, and phosphate (MIP) and with residues, limestone, phosphate,
and potash (RIPK).
***See SF-21 for rotations used. A rotation of corn-soybeans-wheat-hay is used on
most of the fields.
R. H. Rust
11-28-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SP-10
CORN ROOT DISTRIBUTION IN FERTILIZED AND UNFERTILIZED
FLANAGAN SILT LOAM
Comparison of corn root development in
fertilized and unfertilized Flanagan
silt loam at Urbanaj Illinois^ during
the moderately dry season of 195^ shoved
greater root penetration and root growth
and also higher corn yield on the ferti-
lized than on the unfertilized plot.
Over a long period^ the fertilized plot
had received residue (stover^ straw, leg-
umes), lime, and rock phosphate, whereas
the unfertilized plot had received only
crop residues.
Soil tests indicated that, except for ni-
trogen, the greatest difference in fer-
tility was in the sizrface soil, where
the available phosphorus was high on the
fertilized plot and low on the unferti-
lized plot. There was very little dif-
ference in available phosphorus below
the surface soil, and practically no dif-
ference in acidity or in available po-
tassium throughout the soil profiles
from the two plots.
Although Flanagan is a dark-colored, per-
meable, naturally fertile soil, crop
yields invariably decrease with con-
tinued farming unless a good soil manage-
ment program is followed.
Corn yield was 79 bushels an acre on the
fertilized plot and 66 bushels on the
unfertilized plot. Total root weight on
the unfertilized (r) plot was 1,398
pounds an acre, and the roots penetrated
to about hQ inches (see illustration on
back). To this depth this soil is capa-
ble of storing about 10.5 acre-inches of
available water.
On the fertilized (RIP) plot, total root
weight was 1, &^6 poiinds an acre, and the
roots penetrated to about 60 inches.
VJith this larger rooting volume, the
soil had a greater available soil mois-
ture storage capacity (l2.8 acre-inches)
and a greater supply of nutrients for the
crop to draw upon.
Calculated acre weights of corn roots by
soil horizons are given in the table on
page 2. Soil horizons are marked on the
left of the photographs on page 3'
Roots were sampled with a soil-core sam-
pling machine that took four-inch diam-
eter cores to a depth of 72 inches.
Core samples were taken in five concen-
tric rings each four inches wide aroiind
corn hills. In this way
were obtained up to
corn hill, or halfway
since the corn was
inches. Each of the five vertical sec-
tions of roots in the photographs is
from one four-inch diameter core 72
inches long at distances from the corn
hill specified at the top of the root
panels .
For comparison of corn root development
in fertilized and unfertilized Cisne
silt loam during the moderately dry sea-
son of 1952, see Agronomy Facts SM-5.
J, B, Fehrenbacher
12-19-55
root samples
20 inches from the
to the next hill,
checked i^-0 by kO
-2-
Calculated
Calculated
Percent
Soil
root weights
root weights
of total
horizon
Depth
per acre
per acre-inch
roots
in.
lb.
lb.
perct.
R Plot
^1
0-li+
kkk
32
31.8
^
11^-18
11.5
36
10.1+
\
18-23
168
3i^
12.0
\
23-39
1^58
29
32.7
^
39-i^3
85
21
6.1
^1
^3-5^^
77
7
5.5
\
5U-72
21
1
1.5
Total
1,398
--
100.0
RIP Plot
h
0-16
769
1^8
1+1.7
s
16-20
li^3
36
7.7
\
20-2I4-
142
36
7.7
\
2^4-^0
435
27
23.6
^3
ko-k6
165
27
8.9
^1
h6-^l
138
13
7.5
^1
57-72
3h
k
2.9
Total
l,eh6
--
100.0
CORN ROOTS IN FLANAGAN SILT LOAM-R PLOT
CORN ROOTS IN FLANAGAN SILT LOAM-RLP PLOT
INCHES FROM HILL
0 4 9 12 16 20
SURFACE Ai
SliBSURFACE_ Aj^
Bi
SUBSOIL Ba JV'
|te4
2>:
SUBSTRATA b:> .
SURFACE A I
SUBSURFACE
SUBSOIL
B3
SUBSTRATA
INCHES FROM HILL
0 4 B 12 16 20
if
I FT
2 FT
3 FT
4 FT
5 FT
6 FT
Corn roots in Flanagan silt loam from a fertilized (RLP) plot on the left and
an unfertilized (r) plot on the right. In both photographs each of the five
vertical sections of the root panels is from one soil core k inches in diameter
and 72 inches long. Distance of each vertical section from the corn hill is
indicated at the top of the root panel.
jNIVERSin >-yr ILLINOIS ■ COLLEGE OF AGRICULTURE
AGRONOMY FACTS
I
SP-11
THE PRODUCTIVITY OF DARK, TILL-DERIVED SOILS IN NORTHEASTERN ILLINOIS
Soils in the northeastern one -fifth of
Illinois are derived primarily from cal-
careous glacial till of Wisconsin age.
Although most of these soils are dark
colored and appear similar to the casual
observer^ they differ widely in produc-
tivity (Table l).
These differences in productivity of the
various soils are related to the texture
of the subsoil and underlying glacial
till (SP-7). The underlying parent ma-
terial of Clarence -Rove soils contains
so much clay that moisture movement (data
in Soil Sci. Soc. Am. Proc.^ Vol. l^i:
p. 51-55^ 1950^ and Agricultural Engi-
neering^ Vol. 30: p. 38if-386;, 19k9) and
root penetration (unpublished data of J. B.
Fehrenbacher ) are restricted and crop
yields are rather low^ even under a
moderately high level of management.
In contrast^ such soils as Saybrook^ Lis-
bon^ and Drummer^ which are derived from
permeable loam till^ produce consider-
ably higher crop yields under comparable
management. Such management problems as
drainage and erosion are also much more
difficult on Clarence-Rowe soils than on
Saybrook, Lisbon, and Drummer. The phys-
ical properties, productivity, and man-
agement problems of Swygert-Bryce and
Elliott -Ashkum soils are intermediate
between those of the preceding two soil
associations.
Table 1. --Average per acre yields* of corn, soybeans, and oats obtained
by farmers on certain northeastern Illinois soils under medium
and moderately high levels of management
Soil
Texture of
underlying
Corn
Soybe
Me-
2 ans
Mod.
Oat
s
associ-
Me-
Mod.
Me-
Mod.
ation
glacial
dium
high
dium
high
dium
high
area**
till
Soil series**
mgt.
mgt.
mgt.
mgt.
mgt.
bu.
mgt.
bu.
bu.
bu.
bu.
bu.
C
Loam
Saybrook, Lisbon,
and Drummer
70
79
28
33
56
65
E
Silty clay
loam
Elliott and Ashkum
62
66
27
30
50
53
G
Silty clay
Swygert and Bryce
56
6k
25
26
kk
51
G
Clay
Clarence and Rowe
53
61
21
27
36
k9
*About 95 percent of the yield data represent the decade from 19^0 to 1950. The
standard error of estimate of these figures, when considered as 10-year average
yields, is calculated to be approximately + 5 percent.
**Described in Illinois Agricultural Experiment Station publication AG-1^^3^ en-
titled "Illinois Soil Type Descriptions."
The crop yields in Table 1, which are
based upon detailed records kept by
farmers, are given for two levels of
management. These two management levels,
designated 'inedium" and "moderately high,"
are defined in terms of pounds of nitrogen.
phosphate, and potash used and the in-
terval of time between legumes and the
crop under study.
For corn and soybeans, yields were cal-
culated by assiffiiing 6.1 inches of total
rainfall and an average maximum tempera-
ture of 86 F, for the months of July and
August. These figures are averages for
the period 1925-50 reported "by the
weather stations in the area.
A medium level of management for estimat-
ing corn yields consisted of 50 pounds
of N per acre in the current and previ-
ous year^ contributed from both legume
and nonlegiime sources; 20 pounds each of
equivalent P2O5 and K2O per acrO;, applied
or estimated as residual from previous
applications; and a legume or legume -
grass mixture two years before the corn
crop.
Comparable figures for a moderately high
level of management were 100 pounds of K
per acre_, kO pounds each of equivalent
P2O5 and KgO per acre^ and the equiva-
lent of a legume mixture^ such as alfalfa-
red clover^ immediately preceding the
corn crop.
For soybeans and oats the corresponding
requirements for medium and moderately
high management were approximately 70
percent (ranging from ^+0 to 100 percent)
of those given for corn.
Crop yields on the Joliet experiment
field, which is located on Elliott silt
loam and Ashkum silty clay loam, are
given in Table 2. These data indicate
that both limestone and phosphate are
needed on these soils if near -maximum
crop yields are desired. Comparison of
yields obtained with RIPK treatment on
the Joliet experiment field (Table 2)
with the yields in Table 1 for Elliott
and Ashkum soils indicates that farm
yields under the moderately high level of
management are approximately 80 to 90
percent of the yields with RLPK treat-
ment. These results indicate that with
improved management practices farmers
may obtain higher crop yields on Elliott -
Ashkum and probably other till-derived
soils in northeastern Illinois.
Table 2. --Average per acre yields of corn, soybeans, and oats obtained
with various soil treatments* on Elliott silt loam and Ashkum
silty clay loam at the Joliet Experiment Field, 19^^-1955
Yields with various treatments*
Crop
**
RL
RIP
RLPK
First- year corn (after alfalfa — red clover)
Second- year corn (after soybeans)
Soybeans
Oats
bu.
ko
32
23
51
bu.
53
hi
25
52
bu.
77
60
29
62
bu.
8lf
72
32
58
^Symbols for the various soil treatments are: 0 = no treatment; R = crop residues;
L — limestone; P = rock phosphate; K = muriate of potash. Refer to Illinois Agri-
cultural Experiment Station Bulletin 5I6 for information on soil treatment and
cropping prior to 19^2.
**The cropping system followed is corn, soybeans, corn, oats (legume catch crop),
wheat, alfalfa — red clover.
R. T. Odell
1-2-1956
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
SP-12
BOTTOMLAND SOILS OF ILLINOIS
Because interest in the management of
bottomland soils has heen increasing,
the following brief discussion and the
key on the opposite page are given to
help identify such soils and group them
J according to their management needs and
adaptation.
' All of the soils in the very poorly
drained column in the key need drainage.
Bonnie_, Jacob, Fordyce, Karnak, Darwin,
Turtle Creek, Muskrat, and Wabash are so
slowly permeable to water that drainage
by tile is usually not practical. In
such soils open ditches are recommended
and are usually used. Jacob is so diffi-
cult to drain that it is best used for
timber or pasture. Some of the others,
like Karnak, Darwin, Muskrat, and VJabash,
sometimes occur in sloughs where very
poor outlets make drainage impractical.
Such areas are best used for production
of water-loving species of trees or for
wildlife.
Most large areas of any of the very
poorly to poorly drained soils except
Romeo need some dredging to get proper
outlets for either tile or open-ditch
drainage systems. Some of the soils
having sandy substrata, like Wewart,
Riley, Gorham, and Ambraw, may present
difficult problems of ditch-bank main-
tenance if ditches must be cut through
them into the sandy, underlying mate-
rials.
The soils in the moderately well to well
drained column do not need drainage.
Some of the soils in the imperfectly
drained column, like Belknap, Dupo, VJake-
land, Coffeen, Tice, and Gorham, need
drainage for best crop growth. Of this
group, Bellaiap is least responsive to
tile drainage, and in it open ditches
are usually recommended. The others usu-
ally have low enough water tables and
are permeable enough to keep drainage
from becoming much of a problem.
Surface soil texture and also organic
matter content, of which color is a
fairly good indicator, determine work-
ability to a large extent. So far as
possible in the key, surface texture is
arranged within pH groups, with the
coarser textures at the top and the
finer textures at the bottom. On the
fine-textured soils (silty clay lo.am or
finer), plowing in the fall and working
only when moisture conditions are favor-
able will help to prevent compaction and
cloddiness.
Assuming adequate drainage, the light -
colored soils have the highest nitrogen
needs, and the dark-colored soils the
lowest nitrogen needs for satisfactory
crop growth. The strongly to moderately
acid (pH<5.8) light-colored soils have
the highest limestone, phosphate, and
potash needs. Superphosphate and mixed
fertilizers should be used for specific
crops on those soils that are alkaline
(pH>7.5)» Limestone should not be used
on the soils that have high pH values.
Most of the soils having pH values with-
in the range of 5«8 to 7*5 are mediiun
to high in available phosphorus and
available potassium. However, the soils
in this group having pH values near ^.Q
will need some limestone, phosphorus,
and potassium to produce high yields of
most crops, particularly legimies.
Two of these bottomland soils, Burnside
and Romeo, are not well suited for gen-
eral crops because they have only shal-
low soil profiles over bedrock. Pasture,
or in some cases timber, is the best use
for these two soils. Perks, because of
its very sandy texture, is usually
drouthy and not well suited for simmer
crops like corn and soybeans.
Lack of space makes it possible to show
only the major characteristics that are
useful in identifying and classifying
the bottomland soils in the key.
, A
KEY TO BOTTOMIAND SOILS OF ILLINOIS
/
(
Colorft/
sur-
face
soil
of
pro-
file
Inches
of
sur-
face
Texture
Soil seriesii/ grouped according
to natural drainage;^/ of profile
Surface soil
Below surface
Very poor
to Door
Imperfect
ModerateOj
well to well
Light
<5.8
12-36
silt loam
sandstone
Burns ide
Btirnside
>8e/
silt loam
silt loam
BeUcnap
Sharon
<8
silt loam
silt loam
Bonnie
silty clay loam
silty clay loam
Piopolis
silty clay to clay
silty clay to clay
Jacob
5.8
to 7.5
12-36
fine sandy loam
sand
Landes 1
15-ij-o
silt loam
silty clay loam
to silty clay
Dupo
Dupo
Arenzville
>8
silt loam
silt loam
Wake land
Haymond
1
<8
silt loam
silt loam
Birds
silty clay to clay
For dye e
silty clay loam
silty clay loam
Petroli^
silty clay to clay
silty clay to clay
KarnakL/
>7.5
>8
sand
sand
Perks
silt loam
silty loam to
siltv clay loam
Jules
Jules
Moder-
ately
dark
5.8
to 7.5
30-ito
silt loam
sand
Newart
clay loam to
silty clay loam
sand
Newart
8-30
silty clay loam
sand
Riley
>8
silt loam
silt loam
Coffeeni/
Coffeeni/
Kempers/
silty clay loam
silty clay loam
Beaucoup
Tice
Allison
gravelly clay loam.
gravelly clay loam
Beaucoup
, /
clay loam to
silty clay loam
clay loam to
silty clay loam
Ambraw^/
GorhamS/
silty clay to clay
silty clay
to clay
Darwin
Turtle Creek
1
Dark
5.8
to 7.5
12-30
silt loam
silty clay loam
Radford
>8
loam
sandy clay
Muskrat
sandy loam to
clay loam
Otter
Huntsville
silt loam
silt loam
Huntsville
gravelly clay loam
gravel
Huntsville
silty clay loam
silty clay loam
Sawmill
clay loam
silty clay loam
Sawmill
silxy clay to clay
silty clay to clay
VJabash
>7.5
>8
loam to silt loam
loam to silt loam
Millington
DuPage
DuPage
2-10
silt loam
limestone
Romeo
Romeo
i
a/ Light colors have values of k or more on Munsel soil color charts (dark gray to hrovn or;
lighter). Moderately dark color values are usually 3 (very dark gray to dark hrown).
Dark color values are usually 2 ("black to very dark "brcvn).
h/ pH refers to reaction: pH < 5.8 is strongly to moderately acid; pH 5.8 to 7.5 is
slightly acid to neutral; pH >7.5 is plkaline (usually calcareous),
c/ Soil series name plus surface soil texture equals soil type name,
d/ For an explanation of natural soil drainage classes, see Agronomy Facts SP-3.
e/ The symbol > 8 means more than 8. The symbol <C 8 means less that 8.
f/ The pH of Karnak may be as low as 5«0«
g/ Tentative series (not yet correlated),
h/ Ambrav and Gorham are sandy below Uo inches.
J. B. Fehrenbacher
1-23-5^
UNIVERSITY OF ILLINOIS
AGRONOMY FACTS
.-.ULTURE
ORGANIC SOILS IN ILLINOIS
SP-13
Organic soils are soils that contain
more than ahout 25 to 30 percent of or-
ganic matter. They occur in moist to
wet locations where organic materia.1--
primarily plant remains --accimiulated
faster than hirnius decomposed or decayed.
to the botanical composition of the plant
remains and to the nature of accumula-
tion. These in turn are responsible for
differences in texture ^ color ^ shrinkage,
etc.; and determine the uses to which
the peat m.aterials may be put.
Organic soils are of two kinds : peat
soils are those in which the plant re-
mains are sufficiently well preserved to
permit the plant forms to be identified.
Muck soils are those in which the plant
remains are so thoroughly decayed that
the plant parts cannot be recognized.
In most areas peat probably formed be-
fore muck. Some muck may possibly have
formed without first going through the
long period of thick peat accumulation.
The development of muck in this way, how-
ever, would have required short but reg-
ular wet periods for organic matter to
acciomulate, alternating with drier peri-
ods for it to decompose. This alternat-
ing wet and dry cycle would have had to
continue throughout the entire period of
muck development.
About one-foiirth to one -third of the
total area of organic soils in Illinois
is peat, and the remainder is muck.
Little or no true woody peat exists, al-
though shrubs and trees contributed part
of the plant rem.ains in som^e areas, such
as the area at Manito in Mason county.
Also, there is very little true Sphagnum
moss peat in this state. This is the
strongly acid type of peat so well kno'i-m
in Canada and northern United States.
Only two small areas, located in Lake
coiinty, are known in Illinois.
Peat materials in Illinois are chiefly
of two kinds: fibrous (reed-sedge vari-
ety) and sedimentary or colloidal. Dif-
ferences between these two peats are due
Fibrous peat is the most commonly kno'^-m
peat in Illinois. It is formed in shal-
low water from marsh plants like sedges,
reeds, certain grasses, and rushes. These
are upright -growing plants that live
where the watertable remains permanently
at or near the surface. A few of the
more common mosses, particularly some
s ■ :lytrichum and Hypnum, also
hc_kCL. ^o i^iiii the fibrous peat, but not
to the same extent as the m.arsh plants.
Fibrous peat in Illinois is . . ji or
felted, stringy mass that resembles
firmly compressed, half-rc"^""- -traw. It
is usually bro^^m in color _-_ . .ut neu-
tral in rea.ction it .3
large amounts of snaj._L .^nells oi- c-ne_l
fragments. The shells and shell frag-
ments scm.etimes give a grayish cast to
the : --"' ' mass and also provide an ex-
cess 1 /:iount of calcium, carbonate.
Freshly exposed fibrous peat usually
gives off hydrogen sulfide gas, which
has a very distinctive odor.
Fibrous peat is low in ash, usuaJ.j.y con-
taining: less than about 8 to 10 percent.
It i in organic matter, averaging
f • art between 60 and 70 per-
-atively high in nitrogen,
: ween 2 and k percent. It
has about as much total phosphorus as
the surface .l;:vr:v of an average brovm
silt loam pra ' il, such as Saybrook
or Elliott. It contains less total po-
tassium and somewhat less total magnesivim
than the surface of Saybrook or Elliott.
It is- lered high in cellulose, hemi-
cell^^^.^^,. and lignin.
Sedimentary peat is foi-med in small
lakes where the water is at least a foot
or two deep. It is composed primarily
of the remains of aquatic plants^ such
as water lilies, pondweeds, and stone -
worts, and free-floating plants, such as
algae, duckweeds, and diatoms. The re-
mains of such plants tend to disinte-
grate rather thoroughly- -except diatoms,
which are already very small --and upon
settling to the lake bottom form a finely
divided, incoherent, structureless ooze.
The sedimentary type of peat is mostly
gray in color and calcareous--i.e. , high
in lime--and will effervesce with dilute
hydrochloric acid. Compared with fi-
brous peat, it is high in ash, averaging
between kO and 50 percent. It contains
less organic matter than fibrous peat
(about 30 percent) and considerably less
nitrogen (between 1 and 2 percent ) . It
is also relatively low in cellulose, hemi-
cellulose, and lignin.
Sedimentary peat is soft and smooth when
wet, shrinks greatly upon drying, and
dries to a fine, powdery dust. This
dust is easily stirred up by tillage and
by wind, and the diatoms, which have si-
licified cell walls, cause itching when
the soil contacts the skin.
In Illinois most sedimentary peats are
covered by a layer of fibrous peat.
After drainage and cultivation, however,
this layer of fibrous peat decays rapidly
and soon becomes muck. Most mucks are
black and usually contain some added
mineral matter. Although muck soils are
harder to form than the mineral soils,
they are more stable than the peats and
usually need less specialized fertilizer
treatment and management.
More than 90 percent of the organic
soils in Illinois are in the northeast-
ern one -fifth of the state or in the re-
gion lying north, northeast, and east of
McLean county. The few remaining im-
portant areas are in V/hiteside, Henry,
Bureau, and Mason counties, although
other small spots occur in various other
parts of the state.
Individual areas of organic soils vary
in size from small spots of less than
one acre up to more than 1,000 acres.
The combined area of such soils in Illi-
nois totals about 25O square miles. Al-
though this area is small compared with
the total area of the state (about l/2
percent), the organic soils are extremely
important on the individual farms where
they occur. They often differ radi-
cally from the mineral soils with which
they are associated in drainage and fer-
tilizer requirements, workability, and
adaptation to crops.
Three series of organic soils --Houghton,
Lena, and Edwards - -have been established
in Illinois to date. Houghton peat (No.
97 ) is fibrous peat that is about neu-
tral in reaction, and Houghton muck (No.
103) is muck that is approximately neu-
tral in reaction and that decomposed pri-
marily from fibrous peat. Lena peat
(No. 32^) is primarily calcareous fi-
brous peat or fibrous peat that is highly
charged with snail shell fragments, and
Lena muck (No. 210 ) is calcareous muck
formed from both calcareous or shelly
fibrous peat and calcareous sedimentary
peat. Edwards muck (No. 312) is neutral
to calcareous muck between 12 and 36 inches
thick on marl. It is decomposed from
either fibrous or sedimentary peat. No
areas of sedimentary peat that are -cov-
ered by fibrous peat have been mapped.
A few areas of peat and muck consisting
of shallow to mineral material of sand,
silt, and clay textures are known, but
to date they have not been described and
designated as separate soil series.
H. L. Wascher
3-12-56
UNIVERSirt
AGRONOMY FACTS
SP-14
FRAGIPANS IN ILLINOIS SOILS
The term fragipan was formed by combin-
ing paxt of the Latin word fragilis
(fragi ), meaning brittle, with the word
pan which, in reference to soils, means
a horizon or layer that is strongly com-
pacted and dense, indurated, or very
high in clay.
Fragipans differ from claypans in usually
being relatively low in clay, but having
a high silt and/ or sand content. Fragi-
pans that are high in silt have often
been called "siltpans." These veiy slowly
permeable, dense horizons that are ex-
tremely hard when dry usually occur in
soils in the lower part of the subsoil.
In Illinois fragipans are most common in
seme of the upland soils in the southern
part of the state. In this area upland
soils that developed from moderately
thick and thin loess usually nave "pan"
horizons. On the flats, claypans are
found; and on the moderately rolling,
but not steep, better drained areas,
soils with fragipans are common. The
Grantsburg, Hosmer, and Ava soils of
southern Illinois all have fragipan hori-
zons of varying degrees of development
in the lower part of their subsoils. In
the tlilck loess areas bordering the Mis-
sissippi and Wabash river valleys,
weathering or soil development has not
progressed far enough for fragipans to
have formed. However, there is little
doubt but that soil development in these
thick loess areas is in the direction of
fragipan formation.
A| -Dark grayish -brown silt loam, crumb structure.
A2 -Brownish -yellow s: I t loom, ploty structure.
Yellowish-brown silty clay loam, granulor to
' fine subongulor blocky structure.
Bj -Yellowish-brown silty day loom, subongulor blocky structure.
_ Yellowish- brown silty clay loam, subongulor blocky
5roy '-°i'"' "nodyigj heovily cooted with gray silt.
B _ Yellowish-brown silty cloy loom mottled with groy,
priimotlc Structure breoking to blocky oggregotes.
Yellowish-brown silt ipom mottled with groy , extremely
Frogipon — lorge prismatic polygonal aggregates or mony sided
blocks surrounded by very gray streaks of cloy and silt.
The usual morphology of southern Illi-
nois soils with fragipan horizons is
shown in the diagram on the opposite
page. The major type of structure in
each horizon is shown in the diagram and
is also indicated along with color and
texture in the description at the right.
The upper part of the profile above the
gray layer has uniform colors of a well
drained soil} hut because of the mottled
colors in and below the gray layer, the
entire profile is considered to be only
moderately well drained. Fragipan hori-
zons are mixed yellowish brown and gray.
\Then dry they are very hard and brittle,
but upon thorough wetting they slake
down to a noncohesive or only slightly
plastic mass. They seem, therefore, to
be reversibly cemented by some agent.
\lhether the cementing agent is chemical
or whether it is small amounts of cohe-
sive clay between closely packed silt
particles is still a question.
such as Grantsburg and Hosmer, water
moves readily through the upper part of
the profile above the gray layer. Be-
cause of the very slow permeability of
the fragipan, the water often moves lat-
erally in the gray layer above the pan.
During late winter and early spring the
upper part of the profile often is satu-
rated with water, whereas the fragipan
may be only moist. The moisture stor-
age capacity of the fragipans of south-
ern Illinois, in the range available to
plants, is reasonably good, but the in-
ability of roots to penetrate the pan
means that plants are usually deprived
of most of this water.
In Illinois soils fragipans occur at -vari-
ous depths, depending on soil type,
slope, and amovmt of erosion. In most
uneroded areas the top of the pan is be-
low 2^ or 3 feet. The lower boundary is
often indefinite, but the pan is commonly
2 to 3 feet thick.
Fragipan horizons are not entirely lack-
ing in structure, although the structural
aggregates are usually very large, as
shown in the diagram on the opposite
page. The large aggregates are separated
by gray or almost white streaks that are
composed largely of clay. The sides of
the large aggregates next to the gray
streaks are often covered with a black
substance. The vertical dimensions of
these large aggregates, bounded by the
gray streaks, are usually greater than
the horizontal widths so that in excava-
tions with big machinery, such as is
used in road construction or strip min-
ing, the aggregates break out as large
prisms.
Fragipans are very slowly permeable to
water and restrict root penetrations
largely to the gray streaks. In soils
At present there is no proved and prac-
tical means of correcting the adverse
conditions in fragipans. Deep tillage
or mechanical breaking up of the pan re-
quires tremendous power, and the benefi-
cial effects of such treatment, if any,
are unknown. Chemical treatment has not
been tried to any great extent to date.
Fragipans usually have low pH values,
ranging from k.'^ to about 5»5» Base
saturation and available phosphorus are
low, but available potassium is generally
moderate to high. The poor chemical prop-
erties appear to be easier to correct
than the adverse physical properties.
Controlling erosion on soils with fragi-
pan horizons is doubly important. Ero-
sion not only removes valuable topsoil,
but also reduces the depth to the fragi-
pan and thus reduces the rooting depth
of plants.
J. B. Fehrenbacher
lf-9-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICl '"
AGRONOMY FACTS
LAllii.
SP-15
BASIS FOR SEPARATING AND CLASSIFYING SOILS
Soils are characterized and classified
primarily for two reasons: (a) to under-
stand hov they differ and why they differ
and (h) to develop^ as a result of laho-
ratory, greenhouse and field research^
systems of management suitable for dif-
ferent kinds of soils.
The identification and separation of
soil types are hased upon the important
characteristics of the profile (see SP-2)^
especially the following:
1. Color of the various soil horizons
2. Wumher, thickness^ and arrangement
of the horizons
3. Texture of the horizons
k. Structure of the horizons
5. Chemical characteristics of the ho-
rizons^ such as total cation exchange
capacity, kind and amount of ex-
changeable ions, percentage of base
saturation, pH^etc.
6. Mineralogical composition of the ho-
rizons, with special emphasis on the
clay mineral fraction.
Color is one of the most easily observed
soil characteristics. In Illinois, as
elsewhere, color tends to indicate two
entirely different properties: (a) or-
ganic carbon accumulation and (b) degree
of oxidation and diffusion of iron and
manganese compounds.
Organic carbon is black and its accumula-
tion is responsible for the very dark
gray to dark brown to black soil colors,
particularly of the upper or surface ho-
rizons. The acc^umulation of organic
carbon (organic matter) tends to be
greater under grass vegetation than
under forest, other things being equal
(see SP-1 and 2), and greater under
anaerobic (wet) conditions than londer
aerobic (dry). Thus the greatest accumu-
lation of organic carbon and resulting
blackness of the surface or "A" horizon
occurs under grass vegetation in wet
places. Oxides of manganese are also
dark brown to black, particularly in the
hydrated state, but in Illinois they are
never present in sufficient quantity to
produce more than a few dark concretions
or dark splotches on some of the soil
aggregates.
Oxides of iron, primarily in the ferric
state, vary from yellowish brown to red-
dish brown. Under aerobic (well drained)
conditions these compounds are diffused
throughout the soil mass (see SP-3).
They coat so many of the individual soil
particles that the color is uniform
where it is not obscured by organic
matter. Under anaerobic (poorly drained)
conditions the iron compounds are in the
reduced state and are more generally
concentrated into concretions. It is
believed that, as some soils age, mole-
cules of water are lost from the iron
compoiHids and the color gradually changes
from yellowish brown to reddish brown.
The number, thickness, and arrangement
of soil horizons, discussed at some
length in SF-2, are also rather easily
observed characteristics that are used
in identifying soil units. Unweathered
parent material is thought of as having
one soil horizon, i.e., a "C" horizon.
As weathering progresses, horizons de-
velop one by one. In some areas in
Illinois as many as six horizons have de-
veloped, each having features that tend
to distinguish it from adjacent horizons.
The soils known to be most productive
for agricultural purposes under the cli-
matic conditions prevailing in Illinois
are those having a few mediijun-textiired
(loam, silt loam, silty clay loam) hori-
zons and a thick, dark surface.
Texture is another rather easily ob-
served soil characteristic. It often
varies markedly from one horizon to an-
other. In the field it is determined "by
rubbing some soil between the thumb and
fingers (see 111. Cir. 758)^ ^^"t experi-
ence is needed in texturing standard
samples before close correlations can be
made. Laboratory analyses are sometimes
needed for final comparison.
Textiore is a function of relative parti-
cle size and therefore is an indicator
of permeability. Coarse materials^ such
as gravels and sands, have large pores
through which water moves freely. As
the particles become smaller and smaller,
a point is eventually reached in very
fine clay where the pores are so small
that moisture and air movement are often
seriously restricted.
The kind and arrangement of structiiral
aggregates in the various soil horizons
are also useful in characterizing soils.
The size, shape, and arrangement of
structural aggregates give some indica-
tion of the moisture -absorptive capacity
of a horizon, as well as some indication
of its permeability. Loosely packed
granular to rounded aggregates absorb
water more readily and permit easier air
and water movement than tightly packed
angular to square or platy aggregates.
But any form of aggregation tends to pro-
duce cleavage planes or channels that
permit freer water movement and root
penetration than would otherwise be pos-
sible.
Soil acidity, including the presence or
absence of highly calcareous material,
is used as a criterion in separating
certain soil types. Tests with dilute
hydrccMcric acid are used to detect areas
of high-lime (calcareous) soils. pH
tests indicate the relative acidity or
alkalinity of a soil, its probable re-
sponse to liming materials, and its ap-
proximate base saturation. A soil or
any one of its horizons having a pH of
5.0 or less is considered strongly acid.
It is also likely to be relatively low
in exchangeable bases, such as calcium,
magnesiiom, and potassium. Soils having
a pH of about 6.0 are considered slightly
acid, and many of them will be about 70
to 80 percent saturated with bases.
Neutral soils with a pH of 7-0 are in
general 90 to 100 percent saturated with
bases, whereas soils of pH 8.0 usually
contain free basic salts, primarily
calcium carbonate and, less frequently,
salts of magnesium and sodium.
The kind and amount of clay minerals are
important in classification as well as
in the use and management of many soils
(see SP-8). Clays impart sticky or
plastic properties to soils, and any in-
crease in clay above a certain point
increases tillage and drainage difficul-
ties. On the other hand, any decrease
in clay content below a certain point,
especially of montmorillonitic clay, re-
duces the ability of a soil to hold
nutrients in a readily available form
and release them to plants.
The accurate characterization of soil
units consolidates past experience with
those units and indicates the probable
future behavior of the soils under similar
conditions. Classifying soils through
accurately defined characteristics is not
only an attempt to better understand and
interpret nature, but a means of preserv-
ing in an orderly manner the facts known
about the soils that produce our food.
H. L. Wascher and R. T. Odell
1^-16-56
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
CONTROLLING WILD GARLIC AND WILD ONION
W-6
It has been estimated that wild garlic
and wild onion cost Illinois farmers
well over a half -million dollars yearly.
This loss is mainly through dockage in
wheat and does not include loss of milk
and meat products due to undesirable
flavor caused by these weeds.
Wild garlic (Allium vineale) is much
more common than wild onion. VJild onion
(Allium canadense ) is found in the same
areas as wild garlic but usually pre-
sents a small problem compared with its
neighbor.
How to Tell the Plants Apart
Wild garlic
Wild onion
leaves
Form
Base
Cylindrical
Sometimes above-
ground on stems
Bulbs (undergrovind)
Number Clusters at base
of each plant
Covering Fibrous
Kind Hard- shelled (brown)
and soft-shelled
Flavor and odor
Strong
Both of these weeds begin to grow in the
fall from the old plants as well as from
the bulblets. Fall growth starts any
time from September to November, depend-
ing on fall rains. Both weeds seemingly
"grow under the snow" during the winter.
Wild garlic begins to form undergro\and
bulblets in March. In May the aerial
bulblets begin to form on both wild gar-
lic and wild onion.
Both plants have matured by small grain
harvest, and the aerial biolblets are
harvested with the small grain. By mid-
July they become dormant, and plowing
or disking during the summer has little
effect on them. The old plants and the
new bulblets are ready to start their
life cycle again with the fall rains of
September and October,
Both wild garlic and wild onion can be
controlled and eliminated by growing
crops that can be plowed in either late
fall or early spring. If such crops can
be combined with one that can be culti-
vated during the growing season, such
Flat
At ground level, rising
out of the bulb
One at base of each
plant
Ketlike
All soft-shelled
Moderate
as corn or soybeans, both weeds can be
eliminated in three to four years.
Plowing in late fall or early spring
is particularly effective because it
smothers the plants that have germinated
and fall plowing usually prevents the
underground garlic bulblets from form-
ing. The cultivated crop then destroys
any seedlings that may appear with rains
during the growing season. Three or
four years of this program will practi-
cally exhaust all seeds or bulblets that
are in the soil,
2,^-D is also effective in destroying
wild garlic. Experiments conducted from
19^8 to 1950 by the Agronomy Department
showed that 1 l/2 to 2 pounds of 2,U-D
ester in late fall destroyed both the old
plants and the newly germinated seed-
lings. The same rates in early April
were only slightly less effective than
the late fall spraying.
Unfortunately, winter wheat will not
tolerate these rates of 2,i|— D. Only
pastures or stubblefields can be sprayed
with these amounts.
The folloving table on wheat yields in-
dicates that winter wheat will not
tolerate a rate of 2,k-'D that will effec-
tively control aerial bulblets without
seriously reducing yields.
Percent
Rate of
Yield of
control of
2,l^-D ester/A
wheat aerial bulhlets
Fall Applica
ticn, November 22, 1948
Check
32.0
0
1/8
28.0
20
lA
31.1
40
1/2
25.3
65
1
23.5
100
1 1/2
20.9
ICO
Spring Appli
cation, April, 5^
1949
Check
21.4
0
1/k
18.8
25
1/2
19.1
50
1
16.4
70
2
lo.o
eo
Spring Application, April 29,
, 1949
Check
16.8
0
l/,3
16.3
0
2/3
14.4
10
11/3
12.0
30
2 2/3
13.0
40
Applications made about the first week
in April at I/2 pound of 2,4-D ester per
acre have had very little effect on
wheat yields but have reduced aerial
bulblet formation an average of 50 per-
cent. In addition, the remaining plants
have been so deformed that the combine
has picked up very few of them. Appli-
cations in late April have not been ef-
fective in controlling garlic in winter
wheat .
The best way to control both wild garlic
and wild onion is to change the cropping
sequence so that ctiltivated crops are
grown continuously for three to four
years. If this program cannot be car-
ried out, then applications of l/2 pc\and
of 2,4-D ester applied to winter wheat
about the first week in April will ma-
terially reduce aerial biilblet formation
and will gradually thin out stands of
wild garlic. One and a half to 2 pounds
of 2,4-D acid in the ester form is high-
ly effective in eliminating garlic either
in pastures without legumes or in stub-
blefields. The application can be made
either in late fall or in early spring.
F. W. Slife
10-24-55
UNIVERSITY OF ILLINOIS • COLLEGE OF AGRICULTURE
AGRONOMY FACTS
W-7
REACTION OF VARIOUS V/EEDS AND BRUSH TO 2,4-D AND 2,4,5-T
Many veeds and woody plants are not
killed when sprayed with 2^i4--D and
2,k,'^-T, while others are easily con-
trolled. Following is a listing of
various weeds and woody plants and their
reactions to 2,U-Dand 2,h,^-1! herbicides.
Annuals
Weeds and Their Reaction to 2,4-E
Susceptible Perennials & Biennials - Susceptible (Cont.)
Beggar-ticks
Bitter wintercress
Black medic
Carpet weed
Cocklebur
False flax
Flower-of -the -hour
Hemp
Hemp -nettle
Henbit
Jewelweed
Kochia
Lambsquarters
Marsh elder
Morning glory, annual
Mustards
Peppergrasses
Pigweeds
Plantain, annual
Prostrate verbain
Radish, wild
Ragweeds
Rape, annual
Rough cinquefoil
Sow thistle, annual
Stinkweed
Sunflower
Vetch
Yellow star thistle
Annuals - less Susceptible
(plants may recover under seme conditions,
Bedstraw
Buckwheat, wild
Chickweed
Dodder
Dog-fennel
Fleabane
Goosefoot
Jimsonweed
Khotweed
Lettuce, wild
Mallow, roundleaved
Purslane
Russian thistle
Shepherd's purse
Smartweeds
Speedwells
Velvet weed
Annuals - Not Susceptible
Annual grasses
Black nightshade
Buffalo bur
Catchfly
Corn cockle
Cow cockle
Wild cucumber
Wood sorrel
Perennials and Biennials - Susceptible
(Frequently killed by one application)
Artichoke
Broadleaf plantain
Buckhorn
Bull thistle
Burdock
Catnip
Chicory
Cinquefoils
Coneflowers
Creeping charley
Dandelion
Dragonhead
Evening primrose
False ragweed
Figwort
Four-o-clock
Gummweed
Heal-all
Hedge bindweed
Hedge nettle
Hoary alyssum
Horsetail
Licorice, wild
Moonseed
Kettle, stinging
Poppy, mallow
Roadside thistle
Rosin weed
Skelton weed
Slender rush
Verbains
Water hemlock
VJild parsnip
Perennials & Biennials - Less Susceptible
(Tops are killed but regrowth may occur)
Bouncing Bett
Buttercups
Canda thistle
Carrot, wild
Docks
Dogbane
Field bindweed
Goatsbeard
Goldenrod
Lettuce, blue
Poke weed
Poverty weed
Sorrel, red
Teasel
Yarrow
Perennials & Biennials - Wot Susceptible
Asters
Bittersweet
Bracken
Catchfly
Cattail
Chickweed, mouse-ear
Climbing milkweed
Ferns
Foxglove
Goatweed
Ground cherry
Hoary verbain
Milkweeds
Mullen, ccmmon
Nettles, horse
Ox-eye daisy
Russian knapweed
Smartweed, swamp
Sorrel, yellow
Spurges
Strawberry, wild
Tansy rag\rort
Tick-trefoils
Toadflax
Violets
White cockle
Woody Plants
Woody Plants - Susceptible (Cont.
A mixture of 2,U-D and 2,k,'^-1 or
straight 2.,h,^-']l is more effective than
2,4-D alone on most of these plants.
Vfoody Plants - Susceptible
(Current growth killed but retreatment
may be required. )
Alder
Apple ^ crab
Aspen
Barberry
Birch^ black
Blackberry
Boxelder
Buckbrush
Cherry, wild
Cottonwood
Currant
Dogwood
Elderberry
Elm
Gooseberry
Grape, wild
Hackberry
Hazelnut
Honeysuckle
Locust, black
Mulberry
Osage orange
Persimmon
Plum, wild
Poison ivy
Sassafras
Sumac
Tamarisk
Trumpet vine
Virginia creeper
Walnut, black
Willows
Woody Plants - Not Susceptible
Ash
Basswood
Bittersweet
Lead plant
Locust, honey
Oak
Raspberries
Red cedar
Rose, wild
Earl C. Spurrier
U-30-56
8/4/2010
Z
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