1DJANA
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no. 102
Zoology
NEW SERIES, NO. 102
Atlas of New World Marsupials
Barbara E. Brown
January 30, 2004
Publication 1527
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FIELDIANA
Zoology
NEW SERIES, NO. 102
Atlas of New World Marsupials
Barbara E. Brown
Division of Mammals
Department of Zoology
Field Museum of Natural History
1400 South Lake Shore Drive
Chicago, Illinois 60605-2496
U.S.A.
Accepted October 23, 2002 BIOLOGY LIBRARY
Published January 30, 2004 BURRILL HALL
Publication 1527 A __ ,, -
SEP 1 2004
PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY
2004 Field Museum of Natural History
ISSN 0015-0754
PRINTED IN THE UNITED STATES OF AMERICA
Table of Contents
Introduction 1
Acknowledgments 2
The synonymies follow:
I. Order Didelphimorphia 4
Family Didelphidae Gray, 1821 4
Subfamily Caluromyinae Kirsch, 1977 4
Genus Caluromys J. A. Allen, 1900 4
Caluromys derbianus (Waterhouse, 1 84 1 ) 4
Caluromys lanatus (Olfers, 1818) 8
Caluromys philander (Linnaeus, 1758) 12
Genus Caluromysiops, Sanborn, 1951 14
Caluromysiops irrupta Sanborn, 1951 14
Genus Glironia Thomas, 1912 14
Glironia venusta Thomas, 1912 14
Subfamily Didelphinae Gray, 1821 16
Genus Chironectes Illiger, 1811 16
Chironectes minimus (Zimmermann, 1780) 17
Genus Didelphis Linnaeus, 1758 22
Didelphis albiventris Lund, 1841 22
Didelphis aurita Wied-Neuwied, 1826 30
Didelphis marsupialis Linnaeus, 1758 31
Didelphis virginiana Kerr, 1792 40
Genus Gracilinanus Gardner and Creighton, 1989 41
Gracilinanus aceramarcae (Tate, 1 93 1 ) 41
Gracilinanus agilis (Burmeister, 1854) 41
Gracilinanus dryas (Thomas, 1898) 44
Gracilinanus emiliae (Thomas, 1909) 46
Gracilinanus marica (Thomas, 1898) 47
Gracilinanus microtarsus (Wagner, 1842) 48
Gracilinanus perijae Hershkovitz, 1 992 49
Genus Hyladelphys (Voss, 2001) 49
Hyladelphys kalinowskii (Hershkovitz, 1992) 49
Genus Lestodelphys Tate, 1934 49
Lestodelphys halli (Thomas, 1921) 50
Genus Lutreolina Thomas, 1910 50
Lutreolina crassicaudata (Desmarest, 1804) 51
Genus Marmosa Gray, 1821 54
Marmosa andersoni Pine, 1972 56
Marmosa canescens (J. A. Allen, 1893) 58
Marmosa lepida (Thomas, 1888) 59
Marmosa mexicana Merriam, 1897 59
Marmosa murina (Linnaeus, 1758) 60
Marmosa robinsoni Bangs, 1898 65
Marmosa rubra Tate, 1931 68
Marmosa tyleriana Tate, 1931 68
Marmosa xerophila Handley and Gordon, 1979 69
Genus Marmosops Matschie, 1916 69
Marmosops bishopi (Pine, 1981) 69
Marmosops cracens Handley and Gordon, 1979 70
Marmosops dorothea (Thomas, 1911) 70
Marmosops fuscatus (Thomas, 1896) 71
Marmosops handleyi Pine, 1981 74
Marmosops impavidus (Tschudi, 1844) 74
Marmosops incanus (Lund, 1840) 77
Marmosops invictus (Goldman, 1912) 79
Marmosops juninensis (Tate. 1 93 1 ) 80
Marmosops neblina (Gardner, 1990) 80
Marmosops noctivagus (Tschudi, 1845) 80
Marmosops panidens (Tate, 1 93 1 ) 83
Marmosops paulensis (Tate, 1 93 1 ) 85
Marmosops pinheiroi (Pine, 1981) 85
Genus Metachirus Burmeister, 1854 86
Metachirus nudicaudatus (E. Geoffrey, 1 803) 87
Genus Micoureus Lesson, 1842 92
Micoureus alstoni (J. A. Allen, 1900) 93
Micoureus constantiae (Thomas, 1904) 93
Micoureus demerarae (Thomas, 1905) 95
Micoureus regina (Thomas. 1898) 100
Genus Monodelphis Burnett, 1830 [1829] 101
Monodelphis adusta (Thomas, 1897) 102
Monodelphis americana (Mullen 1776) 102
Monodelphis brevicaudata (Erxleben, 1777) 104
Monodelphis brevicaudis Olfers. 1818 107
Monodelphis dimidiata (Wagner, 1847) 107
Monodelphis domestica (Wagner, 1 842) 110
Monodelphis emiliae (Thomas, 1912) Ill
Monodelphis glirina (Wagner, 1 842) Ill
Monodelphis iheringi (Thomas, 1 888) 113
Monodelphis kunsi Pine, 1975 114
Monodelphis maraxina Thomas, 1 923 114
Monodelphis osgoodi Doutt, 1 938 114
Monodelphis palliolata (Osgood, 1914) 116
Monodelphis rubida (Thomas, 1 899) 117
Monodelphis scalops (Thomas, 1 888) 118
Monodelphis sorex (Hensel, 1 872) 118
Monodelphis theresa Thomas, 1921 120
Monodelphis unistriata (Wagner, 1842) 120
Genus Philander Brisson, 1 762 121
Philander andersoni (Osgood, 1913) 1 22
Philander frenata (Olfers, 1818) 124
Philander mcilhennyi Gardner and Patton, 1 972 127
Philander opossum (Linnaeus, 1 758) 1 29
Genus Thylamys Gray, 1843 137
Thylamys elegans (Waterhouse, 1839) 137
Thylamys macrura (Olfers, 1818) 139
Thylamys pallidior (Thomas, 1902) 140
Thylamys pusilla (Desmarest, 1 804) 143
Thylamys velutinus (Wagner, 1842) 145
Thylamys venustus (Thomas, 1902) 145
II. Order Paucituberculata 147
Family Caenolestidae Trouessart, 1898 147
Genus Caenolestes Thomas, 1895 147
Caenolestes caniventer Anthony, 192 1 147
Caenolestes condorensis Albuja and Patterson, 1996 148
Caenolestes convelatus Anthony, 1924 148
Caenolestes fuliginosus (Tomes, 1863) 148
Genus Lestoros Oehser, 1 934 151
Lestoros inca (Thomas, 1917) 151
Genus Rhyncholestes Osgood, 1924 152
Rhyncholestes raphanurus Osgood, 1924 152
III. Order Microbiotheria 153
Family Microbiotheriidae Ameghino, 1887 153
Genus Dromiciops Thomas, 1894 153
Dromiciops gliroides Thomas, 1894 153
Literature Cited 156
Gazetteer of Collecting Localities 176
List of Maps
1 . Distribution of the genus Caluromys in Central America 4
2. Distribution of the genus Caluromys in southern Central America 5
3. Distribution of Caluromys derbianus in Central America and
South America 7
4. Distribution of Caluromys lanatus 11
5. Distribution of Caluromys philander 15
6. Distribution of Caluromysiops irrupta 16
7. Distribution of Glironia venusta 17
8. Distribution of Chironectes minimus in Central America 20
9. Distribution of Chironectes minimus in South America 21
10. Distribution of the genus Didelphis in Central America 23
1 1 . Distribution of the genus Didelphis in southern Central and South America ... 24
12. Distribution of Didelphis albiventris in South America 28
1 3. Distribution of Didelphis aurita in South America 32
14. Distribution of Didelphis marsupialis in Central America 36
15. Distribution of Didelphis marsupialis in South America 37
1 6. Distribution of the genus Gracilinanus 42
1 7. Distribution of Gracilinanus aceramarcae 43
1 8. Distribution of Gracilinanus agilis 45
19. Distribution of Gracilinanus dryas 46
20. Distribution of Gracilinanus emiliae 47
2 1 . Distribution of Gracilinanus marica 48
22. Distribution of Gracilinanus microtarsus 50
23. Distribution of Gracilinanus perijae 51
24. Distribution of Hyladelphys kalinowskii 52
25. Distribution of Lestodelphys halli 53
26. Distribution of Lutreolina crassicaudata 55
27. Distribution of the genus Marmosa in Central America 56
28. Distribution of the genus Marmosa in South America 57
29. Distribution of Marmosa andersoni 58
30. Distribution of Marmosa canescens 59
3 1 . Distribution of Marmosa lepida 60
32. Distribution of Marmosa mexicana 61
33. Distribution of Marmosa murina 66
34. Distribution of Marmosa robinsoni 69
35. Distribution of Marmosa rubra 70
36. Distribution of Marmosa tyleriana 71
37. Distribution of Marmosa xerophila 72
38. Distribution of the genus Marmosops 73
39. Distribution of Marmosops bishopi 74
40. Distribution of Marmosops cracens 75
41. Distribution of Marmosops dorothea 76
42. Distribution of Marmosops fuscatus 77
43. Distribution of Marmosops handleyi 78
44. Distribution of Marmosops impavidus 79
45. Distribution of Marmosops incanus 80
46. Distribution of Marmosops invictus 81
47. Distribution of Marmosops juninensis 82
48. Distribution of Marmosops nebllna 83
49. Distribution of Marmosops noctivagus 84
50. Distribution of Marmosops parvidens 85
5 1 . Distribution of Marmosops paulensis 86
52. Distribution of Marmosops pinheiroi 87
53. Distribution of Metachirus nudicaudatus 90
54. Distribution of the genus Micoureus 94
55. Distribution of Micoureus alstoni 95
56. Distribution of Micoureus constantiae 96
57. Distribution of Micoureus demerarae 99
58. Distribution of Micoureus regina 101
59. Distribution of the genus Monodelphis 103
60. Distribution of Monodelphis adusta 1 04
61. Distribution of Monodelphis americana 105
62. Distribution of Monodelphis brevicaudata 1 08
63. Distribution of Monodelphis brevicaudis 1 09
64. Distribution of Monodelphis dimidiata 110
65. Distribution of Monodelphis domestica 112
66. Distribution of Monodelphis emiliae 113
67. Distribution of Monodelphis iheringi 114
68. Distribution of Monodelphis kunsi 115
69. Distribution of Monodelphis maraxina 116
70. Distribution of Monodelphis osgoodi 117
7 1 . Distribution of Monodelphis palliolata 118
72. Distribution of Monodelphis rubida 119
73. Distribution of Monodelphis scalops 120
74. Distribution of Monodelphis sorex 121
75. Distribution of Monodelphis theresa 122
76. Distribution of Monodelphis unistriata 123
77. Distribution of the genus Philander in Central America 124
78. Distribution of the genus Philander in South America 125
79. Distribution of Philander andersoni 126
80. Distribution of Philander frenata 128
8 1 . Distribution of Philander mcilhennyi 1 29
82. Distribution of Philander opossum in Central America 134
83. Distribution of Philander opossum in South America 135
84.
85.
86.
87.
89.
90.
91.
92.
93.
94.
95.
96.
97.
98.
Distribution
Distribution
Distribution
Distribution
88. Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
Distribution
of the genus Thylamys 138
of Thylamys elegans 139
of Thylamys macrura 140
of Thylamys pallidior 142
of Thylamys pusilla 144
of Thylamys velutinus 146
of Thylamys venustus 148
of the genus Caenolestes 1 49
of Caenolestes caniventer 150
of Caenolestes condorensis 151
of Caenolestes convelatus 152
of Caenolestes fuliginosus 153
of Lestoros inca 1 54
of Rhyncholestes raphanurus 155
of Dromiciops gliroides 156
VII
Atlas of New World Marsupials
Barbara E. Brown
Introduction
Recent Neotropical marsupials now include
three orders and at least 21 genera and 83 species.
New species are still being discovered, and most
groups stand in need of taxonomic revisions.
New World marsupials range from Canada to
Argentina and from the Atlantic to the Pacific
Ocean. They inhabit every major biome, and a
wide range of habitat types within them. Never-
theless, their distributions are not well document-
ed, and detailed taxonomic history and locality
data have not previously been available in one
publication. This atlas is intended fill that gap.
The work was started in 1992 under the direc-
tion of Philip Hershkovitz. Its modest goal was to
determine whether Didelphis marsupialis and D.
albiventris were anywhere sympatric. In Brazil,
we found that even when the same localities were
given for the two species, their elevational ranges
differed, and, in the few cases where they were
the same, the habitat was severely disturbed. Un-
der Hershkovitz's aegis, the work grew to include
the distribution of all the marsupials of South
America, and then Central America and Mexico.
Maps of locality records were also made only to
the species level. Range maps can be found in the
following publications: Emmons and Feer (1990),
Streilein (1982a), and Ojeda and Giannoni (1997,
draft copy). However, unpublished works by
Hershkovitz (1997) and Patterson and Albuja
were available to me in manuscript form and are
included in pertinent accounts. Because Didelphis
virginiana occurs from southern Canada south to
northwestern Costa Rica, and because there are so
many locality records, I decided not to include a
range map of them. For a full account of the ge-
nus Didelphis in North and Central America, see
Gardner (1973). The cutoff date for included ci-
tations is 31 December 2001.
The nomenclature and taxonomy of Gardner
(1993) were followed except in the following in-
stances. Hershkovitz recognized Monodelphis
brevicaudis as a distinct species (see remarks un-
der synonymy), whereas Gardner omitted this
species completely. Voss et al. (2001) recognized
Monodelphis glirina and Monodelphis palliolata
as distinct species, and Gardner listed them under
M. hrevicaudata. Gardner recognized five species
of Thylamys. Subsequent accounts list six; how-
ever, the specific taxonomy of this genus is un-
settled (see Flores et al., 2000, and Mares and
Braun, 2000). Since 1993, one new genus, Hyla-
delphys, and 1 1 new species have been described;
they are Caenolestes condorensis, Hyladelphys
kalinowskii, Marmosops bishopi, Marmosops neb-
Una, Marmosops paulensis, Marmosops pinheiroi,
Monodelphis glirina, Monodelphis palliolata,
Philander frenata, Philander mcilhennyi, and
Thylamys venustus. Under Metachirus, Hershko-
vitz had Didelphis nudicaudatus E. Geoffroy
(1803) (by subsequent designation [Thomas,
1888]) as type species, not Didelphis myosuros
Temminck (1824-1827) in Gardner (see synony-
my for Metachirus); Grubb (2001:41) proposed
that Geoffroy (1803) be placed on the Official List
of the Works Approved as Available for Zoolog-
ical Nomenclature.
Names listed here were excluded by Gardner as
nomena nuda. The following differences between
Hershkovitz's unpublished notes and personal
communications and Gardner (1993) are also not-
ed. Under Lutreolina, Hershkovitz listed Didel-
phis macroura Desmoulins, 1824:492 (= Didel-
phis macroura Illiger, 1815:107), nomen nudum,
and Didelphis mustelina Waterhouse, 1846:497-
499 (nomen nudum) as synonyms; Gardner did
not list them. Under Marmosa, Hershkovitz listed
Marmosa phaea, J. A. Allen, 1912a: 17 (part, not
Thomas, misidentification), as a synonym of M.
robinsoni, and Marmosa murina zeledoni J. A.
Allen, 1912a:73 (not Goldman, misidentification)
as a synonym of M. robinsoni, while Gardner did
not list them. Hershkovitz assigned Caluromys
(Micoureus) antioquiae Matschie, 1917:28, to C.
derbianus, Gardner to C. lanatus. Under Caluro-
mys philander, Hershkovitz listed V. [sic =
D(idelphis)] flavescens Brongniart, 1792:115,
FIELDIANA: ZOOLOGY, N.S., NO. 102, JANUARY 30, 2004, PP. 1-308
whereas Gardner excluded it as a nomen nudum;
Hershkovitz listed Didelphys macrura Pelzeln,
1883:111, while Gardner did not; Hershkovitz
listed D[idelphys] longicaudatci Pelzeln, 1883:
111, and [Didelphys {Philander)] longicauda
Trouessart, 1898:1237, but Gardner excluded the
former as a nomen nudum and the latter as a lap-
sus.
Locality records have been gathered from pub-
lications and from specimens in certain museums.
The following sources for the coordinates of each
locality are Geonet; the United States Board on
Geographic Names (1955-1983); the series of or-
nithological gazetteers published by Paynter
(1982-1994), Paynter and Tray lor (1981-1991),
and Stephens and Traylor (1983-1985); Hanson
(1945); Gardner (1973); Willet (1987); Handley
(1976); published articles on species containing
coordinates of collected localities; Hershkovitz
(1977); various cited records; and the working re-
cords of Hershkovitz compiled over a span of 60
years. The latter are part of the Field Museum of
Natural History Division of Mammals archives.
Authors cited in the synonymies are listed in
the bibliography. Also listed in the synonymies
are additional previously unpublished locality re-
cords for all species except those of Gracilinamis,
Marmosa, Marmosops, Micoureus, Monodelphis,
and Thylamys.
On the maps, solid circles denote the localities
for which there are published records, and open
circles denote the additional (unpublished) re-
cords.
Under each locality name in the Gazetteer, geo-
graphic coordinates and elevations, where known,
are given.
The following abbreviations are used:
AMNH American Museum of Natural History,
New York
BMNH Natural History Museum, London
EPN Ciencias Biologicas, Escuela Politec-
nica Nacional, Quito
FMNH Field Museum of Natural History, Chi-
cago
KU University of Kansas, Natural History
Museum, Lawrence
LACM Los Angeles County Museum, Los
Angeles
LSUMZ Louisiana State University Museum of
Zoology. Baton Rouge
MCZ Museum of Comparative Zoology,
Harvard University, Cambridge
MMNH Bell Museum of Natural History, Uni-
versity of Minnesota, St. Paul
MMP Museo Municipal de Ciencias Natura-
les "Lorenzo Scaglia" de Mar del Pla-
ta, Mar del Plata
MNRJ Museu Nacional, Rio de Janeiro
MPEG Museu Paraense Emilio Goeldi, Belem
MVZ University of California, Museum of
Vertebrate Zoology, Berkeley
MZUSP Museu de Zoologia de Universidad de
Sao Paulo, Sao Paulo
RMNH Rijksmuseum van Naturlijke Historic
Leiden
UMMZ University of Michigan, Museum of
Zoology, Ann Arbor
USNM United States National Museum,
Washington, D.C.
UV Departmento de Zoologia, Universidad
del Valle, Cali
ZFMK Zoologisches Forschungsinstitut und
Museum Alexander Koenig, Bonn
ZMB Zoologisches Museum der Humboldt-
Universitat, Berlin
Acknowledgments
I wish to thank the following persons and in-
stitutions for lists of specimens in their marsupial
collections: Dr. Robert M. Timm, University of
Kansas; Dr. James L. Patton, Museum of Verte-
brate Zoology, University of California, Berkeley;
Dr. Charles O. Handley, Jr., United States Nation-
al Museum; Dr. Mark S. Hafner, Museum of Nat-
ural Science; Louisiana State University Museum;
Dr. Robert S. Voss, American Museum of Natural
History; and Dr. Philip Myers of the University
of Michigan. The assistance of Eunice Hoshizaki
for typing the manuscript and helping in editing
and proofreading and C. Victoria Ramos for ini-
tially generating the maps using ArcView was
deeply appreciated. Lucia Luna provided me with
exceptional help in research and editing the man-
uscript and helped eliminate many editorial short-
comings. Steven Foldi generated new maps for
the final revision and provided further typing and
proofreading. I found truly helpful the comments
and corrections of Dr. Alfred L. Gardner of the
Smithsonian Institution and Dr. Robert M. Timm
of the University of Kansas. The insight, encour-
agement, editorial help, and reading of this work
by Dr. Jack Fooden and especially Dr. Bruce D.
Patterson of the Field Museum were invaluable. I
am most grateful for the aid, advice, and encour-
agement of the late Philip Hershkovitz, without
whose prodding this project would never have
been started or completed.
FIELDIANA: ZOOLOGY
Lutreolina crassicaudata Thomas
BROWN: ATLAS OF NEW WORLD MARSUPIALS
L Order Didelphimorphia
Family Didelphidae Gray, 1821
Subfamily Caluromyinae Kirsch, 1977
Genus Caluromys J. A. Allen, 1900
Sarigua Muirhead, 1819:429 part, S. cayopollin list-
ed among included species.
Asagis and Notagogus Gloger, 1842:82 type by
monotypy the Cayopollin - Caluromys philander.
Notagogus Gloger, 1842:82 name preoccupied by
Notagogus Agassiz, 1833, an extinct genus of fish
of the family Macrosemiidae.
Philander: Burmeister, 1856:74 (not Brisson, 1762)
included species: Philander cayopollin Burmeister
(not Schreber) [= Caluromys philander Linnaeus],
Philander dichura [= Caluromys philander Lin-
naeus]; generic name a homonym of Philander Tie-
demann, 1808, the four-eyed pouched opossum.
Thomas, 1888:336 subgenus of Didelphys; type
species, D. philander Linnaeus. J. A. Allen, 1911:
246 attributed to Brisson, a nonbinomialist, with
type Didelphis philander; Caluromys J. A. Allen in
synonymy. Thomas, 1920a: 195, footnote type
species, Philander philander, by tautonomy. Hop-
wood, 1947:535 Philander Brisson rejected, ge-
neric name attributed to Gronovius, 1763, a non-
binomialist. Gardner, 1993:15 listed.
Caluromys J. A. Allen, 1900b: 189. Gardner, 1993:
15 listed.
Mallodelphys Thomas, 1920a: 195, footnote replace-
ment name for Micoureus Lesson, 1 842 sensu Mat-
schie, 1916, type, Didelphis lanigera Desmarest [ =
Caluromys lanatus Olfers]. Gardner, 1993:15 list-
ed.
Mallodelphis Miranda Ribeiro, 1936:353 unjustified
emendation of Mallodelphys Thomas; included
species: Mallodelphis lanigera.
Type Species Didelphis philander Linnaeus,
by original designation.
Distribution (Figs. 1 and 2) Forested tropi-
cal zones of the Americas from southern Mexico
through Central America and South America to
Paraguay and Rio Grande do Sul, Brazil, on the
south and east; the Pacific coast of Ecuador and
Colombia to the west of the Andes; and the is-
lands of Trinidad and Margarita.
Caluromys derbianus (Waterhouse, 1841)
Didelphvs derbianus (Waterhouse, 1841:97). Water-
house, 1846:493. Alston, 1882:199 distribution
(Nicaragua to Peru). Gardner, 1983:15 listed.
[Didelphys] derbianus: J. A. Allen, 1904a:57 (in
text) COLOMBIA: Valle del Cauca (Rio Cau-
quita, Rio Cauca valley).
Caluromys derbianus: J. A. Allen, 1900a: 189 clas-
sification. Loftin, 1965:83 PANAMA: Veraguas
Fig. 1 . Distribution of the genus Caluromys in Central America: southern Mexico through Panama. See Figure
2 for distribution in South America.
FIELDIANA: ZOOLOGY
20 10
Fig. 2. Distribution of the genus Caluromys in southern Central America and South America: from Belize,
including islands of Trinidad and Margarita, south to southern Paraguay and southern Brazil. See Figure 1 for
distribution in Central America.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
(Santiago). Handley, 1966: 755 PANAMA: Bo-
cas del Torn (Cayo de Agua; Isla Bastimentos);
Chiriqui (Divala; Isla Brava); Veraguas (Isla Ce-
baco; Santa Fe); Los Santos (Guanico); Panama
(Panama Viejo); Herrera (Parita). Reig et al., 1977:
212 COSTA RICA: Cartago (Pacuare, Rio Pac-
uare). Steiner, 1981:506 PANAMA: Colon/Pan-
ama (Isla Barro Colorado [= Barro Colorado]).
Timm, 1989:15 COSTA RICA: Heredia (Finca
La Selva). Albuja, 1992:124 ECUADOR: Man-
abi (Cerro Pata de Pajaro); Los Rfos (Jauneche).
Emmons and Albuja, 1992:120 ECUADOR:
Guayas-Manabi (Parque Nacional Machalilla).
[Didelphis (Micoureus)] derbianus: Matschie, 1916:
269 taxonomy. Matschie, 1917:292 synonyms;
pictus Thomas, pyrrhus Thomas.
Ph[ilander] l[aniger] derbianus: Thomas, 1913b:
358 holotype in Liverpool Museum; COLOM-
BIA: Valle del Cauca (Rio Cauca Valley).
Caluromys derbianus derbianus: Cabrera, 1958:2
classification. Hall and Kelson, 1959:18 COSTA
RICA: Tabernilla [Tabernilla is in Panama, Pana-
ma].
[Didelphis (Philander)] derbyana: Trouessart, 1898:
1238 incorrect subsequent spelling of derbiana,
in synonymy of [Didelphis (Philander)] lanigera.
Caluromys (Mallodelphis) derbiana: Gilmore, 1941:
318 distribution.
Didelphys lanigera: Thomas, 1888b:339 PANA-
MA: Panama (Chepo); COLOMBIA: Antioquia
(Medellfn); ECUADOR: Guayas [now in Manabi]
(Balzar Mountains [= Montanas de Balzar]).
Philander laniger: Pocock, 1913:1094 COLOM-
BIA: Valle del Cauca (Buenaventura).
Philander laniger guayanus Thomas, 1899b:286
ECUADOR: Guayas [now in Manabi] (type local-
ity, Balzar Mountains [= Montanas de Balzar]; ho-
lotype, male, skin and skull, BMNH 80.5.6.87; col-
lected by Illingworth). Lonnberg, 1921:69 EC-
UADOR: Pichincha (Rio Mindo). Gardner, 1993:
15 listed.
Caluromys laniger guayanus: J. A. Allen, 1900a:
1 89 taxonomy.
P[hilander] l[aniger] guayanus: Thomas, 1913b:
358 ECUADOR: (western).
[Didelphis (Micoureus)] guayanus: Matschie, 1916:
269 taxonomy.
Philander centralis Hollister, 1914: 103 COSTA
RICA: Limon (type locality, Talamanca; holotype,
female, USNM 12218/38522, collected by W. M.
Gabb). Harris, 1943:7 COSTA RICA: Limon
(Talamanca [= Sipurio]). Gardner, 1993:15 listed.
Caluromys laniger centralis: Harris, 1943:6 COS-
TA RICA: Limon (Sipurio, 15 mi from Caribbean
coast).
Caluromys derbianus centralis: Hall and Kelson,
1959:17 COSTA RICA: Cartago (type locality, 5
km SE of Turrialba, 1950 ft [= 610 m], KU 26923).
[Didelphis (Micoureus)] canus Matschie, 1917:284
NICARAGUA (type locality). Gardner, 1993:15
listed.
[Didelphis (Micoureus)] antioquiae Matschie, 1917:
286 COLOMBIA: Antioquia (type locality, Ca-
ceres, Rio Cauca; holotype, juvenile female, ZMB
5538/26753; collected by Grosskopf).
Philander laniger pallidas Thomas, 1899b:286
PANAMA: Chiriqui (type locality, Bugaba; holo-
type, male, BMNH, collected 12 October 1895 by
H. J. Watson). Hall and Kelson, 1959:18 NICA-
RAGUA; COSTA RICA: San Jose (Fuentes); Pun-
tarenas (Puerto Cortez); Cartago (Irazu); PANA-
MA: Chiriqui (Boqueron; Bugaba). Gardner, 1993:
15 listed.
Caluromys laniger pallidus: J. A. Allen, 1910:92
NICARAGUA: Matagalpa (Matagalpa).
Caluromys derbianus pallidus: Goodwin, 1946:285
COSTA RICA: San Jose (Fuentes); Cartago (Ira-
zii); Puntarenas (Boruca; Puerto Cortez); PANA-
MA: Chiriqui (Boqueron). Hall and Kelson, 1959:
18 NICARAGUA: Matagalpa (Maltagalpa).
Caluromys laniger pyrrhus Thomas, 190 lb: 196
COLOMBIA: Valle del Cauca (type locality, Rio
Oscuro, near Cali, 1000 m; holotype, male, skin
and skull, BMNH 99.9.6.50; collected June 1898
by J. H. Batty, Parish et al.); ECUADOR: Esmer-
aldas (San Javier, Rio Cachabf). Trouessart, 1910:
A30 ECUADOR: Pichincha (Puente de Ayuri-
quin, on road from Quito to Santo Domingo, 700
m). Gardner, 1993:15 listed.
[Didelphis (Micoureus)] pyrrhus: Matschie, 1916:
269 taxonomy.
Philander laniger senex Thomas, 1913c:573 EC-
UADOR: Pichincha (type locality, Mindo, 4200 ft
[= 1300 m]; holotype, female, skin and skull,
BMNH 13.10.24.75; collected 25 June 1913 by
Gilbert Hammond). Gardner, 1993:15 listed.
[Didelphis (Micoureus)] senex: Matschie, 1916:269
taxonomy.
Philander laniger pictus Thomas, 1913b:360 CO-
LOMBIA: Caldas [now Risaralda] (type locality,
Apia, Rfo Apia, Rio Cauca valley, 5000 ft [= 1560
m]; holotype, female, skin and skull, BMNH
10.7.16.5; collected 10 November 1909 by M. G.
Palmer); ECUADOR: Esmeraldas (San Javier).
Lonnberg, 1921:68 ECUADOR: Pichincha (Pae-
to, below Gualea, 3000 ft [= 930 m]). Gardner,
1993:15 listed.
[Didelphis (Micoureus)] pictus: Matschie, 1916:
269 taxonomy.
[Didelphis (Micoureus)] pulcher Matschie, 1917:
281 (type locality unknown; holotype, female,
skin and skull, ZMB 20467/8; purchased alive from
dealer Rath in Hamburg and died 17 February 1915
in Berlin Zoological Gardens). Gardner, 1993:15
listed.
Philander laniger aztecus Thomas, 1913b:359
MEXICO: Veracruz-Llave (type locality, San Juan
de la Punta; holotype, female, BMNH 94.12.18.28,
received from the Museo Nacional, Mexico); Ta-
basco (Teapa). Gardner, 1993:15 listed.
Caluromys derbianus aztecus: Hall and Kelson, 1959:
17 MEXICO: Veracruz-Llave (Jesus Carranza;
Teapa; San Juan de la Punta).
Philander laniger nauticus Thomas, 1913b:359
PANAMA: Veraguas (type locality, Isla Goberna-
dora; holotype, male, BMNH 03.3.1.124, collected
by J. H. Batty). Goldman, 1920:55 PANAMA:
Chiriqui (Isla Brava; Isla Parida); Veraguas (type
locality, Isla Cebaco). Gardner, 1993:15 listed.
Philander laniger fervidus Thomas, 191 3b:359
Type locality, GUATEMALA (holotype probably
from lowlands of east-central Guatemala or north-
F1ELDIANA: ZOOLOGY
30
20'
10
Caluromys derbianus
Fig. 3. Distribution of Caluromys derbianus: from southern Mexico through Panama in Central America and
from western Colombia and western Ecuador in South America, o = localities from unpublished records; = localities
given in published records.
ern Honduras according to Goodwin, 1942:1 14; ho-
lotype, female, BMNH 1906.10.22.1, collected by
J. Rodriguez). Gardner, 1993:15 listed.
Caluromys derbianus fervidus: Goodwin, 1942:1 14
HONDURAS: Yoro (Catacamas; Chamelecon; lla-
ma). Hershkovitz, 1951:552 BELIZE: Cayo (Ka-
tes Lagoon). Hall and Kelson, 1952:325 HON-
DURAS: Cortes (San Pedro Sula).
Holotype Originally in Lord Derby's Muse-
um but now in the Liverpool Museum (cf. Thom-
as, 1913b:358).
Type Locality Fixed by Cabrera, 1957:2, as
Valle del Cauca, Colombia (type locality fixation
incorrectly attributed to J. A. Allen, 1904a:57, in
text).
Distribution (Fig. 3) Forests of Mexico
through Central America to western Colombia
and western Ecuador.
Additional Records MEXICO: Veracruz-
Llave; Motzorongo (1935, USNM); San Andres
Tuxtla (Dalquest, 1948, KU); Tabasco; Balancan
(Lay, 1960, LSUMZ); La Venta (Weber, 1943,
USNM); Campeche; Escarcega, near (Clifton,
1963, KU); Chiapas; Palenque (Kuns, 1949,
FMNH); BELIZE: Cayo; Baking Pot (Disney,
1964, FMNH); HONDURAS: Yoro; Agua Color-
ada (Stadelman, 1939, FMNH); NICARAGUA:
Matagalpa; Muy Muy (Alcorn, 1956, KU); Es-
quipulas (Choate, 1968, KU; Jones, Jr., 1968, KU;
Smith, 1968, KU); Chinandega; Hacienda Bella-
vista, Volcan Casita (Lawlor, 1966, KU; Turner,
1966, KU); Hacienda San Isidro (Genoways,
1967, KU); Boaco; Boaco, near Santa Rosa (Gen-
oways, 1967, KU; Kirsch, 1987, KU; Phillips,
1967, KU; Smith, 1967, KU); Los Cocos, 14 km
S of Boaco (Choate, 1968, KU; Kirsch 1968, KU;
Phillips, 1968, KU); Tuestepe [= Teustepe] (Phil-
lips, 1968, KU); Managua; Finca America (Bosh-
ell, 1953, USNM); Managua (Alcorn, 1956, KU;
Kellers, USNM); Sabana Grande (Choate, 1968,
KU; Genoways, 1967, 1968, KU; Jones, Jr., 1964,
KU; Lawler, 1964, KU; McGee, 1968, KU;
Schlitter, 1967, KU; Smith, 1964, KU); Masaya;
Masaya (Alcorn, 1956, KU); Zelaya; El Recreo
(Schlitter, 1966, KU); Choluteca; Diriamba (Gen-
oways, 1967, KU; Kirsch, 1968, KU; Phillips,
1967, KU); Granada; Finca Santa Cecilia (For-
man, 1966, KU; Lawlor, 1966, KU; Turner, 1966,
KU); La Calera, Nandaime (Clark, 1964, KU;
Clark, 1964, USNM); Mecatepe, Nandaime
(Clark, 1964, KU; Clark, 1964, USNM); Rivas;
BROWN: ATLAS OF NEW WORLD MARSUPIALS
Rivas (Jones, Jr., 1964, KU); Finca Amayo, 13 km
E of Rivas (Forman, 1966, KU; Lawler, 1966,
KU; Turner. 1966, KU); Sapoa (Jones. Jr.. 1964,
KU); COSTA RICA: Cartago; San Juan (Zele-
don. 1978. USNM); Guanacaste; Finca Jimenez.
0.5 mi E. not located (Hooper. 1996. UMMZ):
Puntarenas; Dominical. 0.5 mi N (Musser. 1962.
UMMZ); San Jose: Monterrey. 22 km S of San
Jose (Willie. 1947, KU): Santa Ana, 2 km N
(Gardner. 1967. LSUMZ): Finca Lornessa. 2 km
NW of Santa Ana (McPherson. 1970. LSUMZ);
San Ignacio (1931. USNM); Caspirola (McPher-
son. 1966, LSUMZ); Limdn; Fortuna. Pandora
(Setzer. 1949. USNM): Finca La Lola, 50 m, not
located (Huckaby, 1964. LSUMZ); PANAMA:
Bocas del Toro; Sibube (Handley. 1963. USNM):
Rio Changuinola (Barrett, 1962, USNM): Chan-
guinola (Handley. 1960. USNM); Boca del Drago
(Handley. 1960. USNM): La Gruta. Isla Colon
(Greenwell. 1988. USNM); Long Bay. Isla Bas-
timentos (Handley. 1963. USNM); Almirante
(Handley. 1960. USNM); Salt Creek, Isla Basti-
mentos (Handley, 1963. USNM; Jacobs. 1987.
USNM); Old Point. Isla Bastimentos (Handley.
1963. USNM); Cedar Creek, Isla Bastimentos
(Handley, 1963, USNM): Bocatorito, Isla San
Cristobal (Greenwell. 1989. USNM): Cuan Creek
(Greenwell. 1989. USNM); Punta Alegre. Penin-
sula Valiente (Greenwell. 1990. USNM); Sum-
wood Channel, Isla Popa (Greenwell. 1991.
USNM); Isla Escudo de Veraguas (Greenwell.
1990. USNM; Handley, 1991. USNM; Nelson.
1991. USNM); Bisira (Handley. 1993. USNM);
Cerro Chucanti. not located (Wetmore. 1950.
USNM); Chiriqui; Progreso (Greenwell. 1966.
USNM); Guabala (Tyson, 1961, USNM); Los
Santos; Las Palmitas (Handley. 1962, USNM);
Colon; Fort Sherman. Camp Pina (1955. 1956.
USNM; Fleming. 1966. USNM); France Field
(1956. USNM); Fort Gulick (Keenen. 1961.
USNM); Fort Davis (Keenen. 1953, USNM; Setz-
er, 1953. USNM); Frijoles. Bahia Peninsula (Pine.
1972, USNM); Las Cascadas (Enders. 1932.
USNM); Panama; Tabernilla (Goldman. 1911.
USNM): Gamboa (Koford. 1956. USNM); Sum-
mit (Keenen. 1955. USNM); Chiva-Chiva (Kee-
nen. 1952. USNM); Fort Clayton (Keenen. 1955.
USNM); Albrook (Keenen. 1954, USNM); Cu-
rundu (Setzer. 1953. USNM): Fort Kobbe (1954.
USNM: Keenen. 1955. USNM); Panama City
(Russell. 1967. USNM); Arraijan (Mendez. 1959.
USNM); La Chorera (Zetek. USNM); La Men-
doxa. not located (Handley. 1958, USNM); 5a/;
Bias; Armila. Quebrada Venado (Handley. 1963.
USNM); Darien; Cana (Goldman. 1912, USNM);
El Real (Handley. 1964. USNM); Tacarcuna
(Handley. 1959. USNM); Cerro Tacarcuna (Han-
dley. 1964, USNM); Rio Pucro, head, Cerro Mali
(Handley. 1964. USNM); Jaque. Rio Imamado
(Handley. 1966. USNM): COLOMBIA: Turreke.
not located (Buckow. USNM); Choco; Unguia
(Hershkovitz, 1950. FMNH); Cordoba; Rio Nam
(Hershkovitz, 1949. FMNH): Valle del Cauea;
Virology Field Camp. Rio Raposo (Duran. 1962.
USNM"); ECUADOR: hnbabura; Hacienda Par-
amba (Mena. 1941. FMNH); Bolivar; Balzapam-
ba (Clarke-Macintyre. 1938, MVZ).
Caluromys ktnatus (Olfers, 1818)
Didelphis Cayopollin et Didelphis Dorsigera Moreau
Saint-Mery (not authors). 1801, in Azara:275
misidentification of "Micoure second ou Micoure
Laineux."
Lanoso. Azara. 1802:221.
Didelphis cayopollin: Fischer (not "Der Kayopollin"
of Schreber). 1813:580 misidentification of mi-
coure second of Moreau-Saint-Mery in Azara
(1801:275).
[Didelphis] lanata Illiger. 1815:107 nomen nudum.
D[idelphys] lanata: Olfers. 1818:206. Gardner. 1993:
15 listed, synonymy.
P[= Caluromys] Lanata: Hershkovitz. 1951:552
comparison; laniger Desmarest. a synonym.
Caluromys lanatus: Yunis et al.. 1972:265 CO-
LOMBIA: Mela (Rio Ocoa. near, on the Villavi-
cencio-Acacias Road). Reig et al.. 1977:195. 212
PERU: Ucayali (Balta. Rio Curanja); Loreto (Yar-
inacocha [now in Ucayali]). Terborgh et al.. 1984:
1 PERU: Madre de Dios (Cocha Cashu Biologi-
cal Station. Manu National Park [= Parque Na-
cional del Manu]). Gardner. 1988:695 VENE-
ZUELA: Amazonas (Base Camp. Cerro de La
Neblina). Emmons et al.. 1994:144 PERU: Ma-
dre de Dios (Explorer's Inn Reserve). Emmons et
al.. 1994:146 PERU: Madre de Dios (Puesto San
Antonio). Anderson. 1997:140 BOLIVIA: La Paz
(Alto Rio Madidi). Patton et al.. 2000:85 BRA-
ZIL: Acre (Igarape Porongaba. right bank Rio Ju-
rua; Igarape Porongaba. opposite, left bank Rio Ju-
rua; Flora [= Fazenda Santa Fe]: Nova Vida. right
bank Rio Jurua: Sobral): Amazonas (Sacado: Al-
tamira. right bank Rio Jurua: Altamira. opposite,
left bank Rio Jurua; Barro Vermelho. left bank Rio
Jurua: Ilha Paxiuba. right bank Rio Jurua; Colo-
cacao Vira-Volta. left bank Rio Jurua on Igarape
Arabidi).
Caluromys lanatus lanatus: Hershkovitz. 1959:338.
345 taxonomy. Avila Pires. 1968:163 BRAZIL:
Minas Gerais (Barra do Rio Paraopeba). Crespo.
1974:3 BOLIVIA: Soma Cruz (Buena Vista);
PARAGUAY: Caazapd (Caazapa). Handley. 1976:
5 VENEZUELA: Apure (Nulita; Selvas de San
Camilo. 29 km SSW of Santo Domingo): Zulia (El
Rosario): Trujillo (Valera); Amazonas (San Juan de
FIELDIANA: ZOOLOGY
Manapiare; Puerto Ayacucho; Boca Mavaca; Es-
meralda; Belen; Acanana). Massoia et al., 1987:
1 16 ARGENTINA: Misiones (Rio Urugua-i). An-
derson et al., 1993:16 BOLIVIA: Santa Cruz
(Buena Vista; Santa Cruz de la Sierra). Avila Pires,
1994:369 BRAZIL: Rio Grande do Sul (Rio
Cahy [= Cail). Anderson, 1997:1 16 BOLIVIA:
Beni (San Joaquin; Piedras Blancas; Estancia Yu-
tiole, 20 km S of San Joaquin; Los Tajibos, 40 km
E of San Borja); Pando (Isla Gargantua); Santa
Cruz (Buena Vista; Santa Cruz de la Sierra).
Didelphis lanigera Desmarest, 1820:258 name
based on the micoure second ou micoure laineux
of Azara. Jentink, 1887b:302 "Demerara" [=
Peru]. Jentink, 1888:221 PERU: ("Demerara" [=
Peru]). Rennger, 1830:225 PARAGUAY. Gard-
ner, 1993:15 listed.
Didelphys lanigera: Waterhouse, 1841:98. Pelzeln,
1883:113 BRAZIL: Mato Grosso (Caicara). Jen-
tink, 1887a:225 PERU: Loreto (Samiria). Thom-
as, 1888b:339 PERU: Loreto (Chamicuros); EC-
UADOR: Pastaza (Sarayacu).
D[idelphys] lanigera: Wagner, 1855:233 BRAZIL:
Mato Grosso (Caicara).
Micoureus lanigera: Lesson, 1842:136 classifica-
tion.
Philander lanigera: Burmeister, 1856:88. Bertoni,
1914:69 PARAGUAY: Alto Parana (Puerto Ber-
toni, upper Rio Parana). Bertoni, 1939:7 PARA-
GUAY: Alto Parana (Puerto Bertoni).
[Cuica] lanigera: Liais, 1872:330 classification.
Philander laniger: Winge, 1893:6, 33 incorrect sub-
sequent spelling of lanigera BRAZIL: Minas
Gerais (Lagoa Santa, recent and fossil). Miranda
Ribeiro, 1914:48 BRAZIL: Mato Grosso (Porto
Espiridao; Salto da Felicidade, Rio Sepotuba). Mi-
randa Ribeiro, 1936:356 BRAZIL: Mato Grosso
(Rio Jauru, upper Rio Paraguay). Vieira, 1955:
347 BRAZIL: Mato Grosso (Rio Jauru, upper
Rio Paraguay).
Caluromys laniger: J. A. Allen, 1900a: 189 incorrect
subsequent spelling of lanigera taxonomy.
Micoureus laniger: Matschie, 1916:269 incorrect
subsequent spelling of lanigera PARAGUAY:
(type locality, Caazapd, Asuncion [now in Cen-
tral]).
Caluromys {Mallodelphis) laniger: Gilmore, 1941:
317 incorrect subsequent spelling of lanigera
taxonomy.
Philander laniger laniger: Cabrera, 1919:33 incor-
rect subsequent spelling of lanigera Vieira, 1955:
346 BRAZIL: Sao Paulo (Araraquara).
Caluromys laniger laniger: Vieira, 1949:349 incor-
rect subsequent spelling of lanigera BRAZIL:
Sao Paulo (Araraquara).
Didelphys ochropus Wagner, 1842:359 type locality,
Barra. Wagner, 1850:141 BRAZIL: Amazonas
(type locality, Barra). Wagner, 1847:148 type lo-
cality clarified as BRAZIL: Amazonas, Barra do
Rio Negro. Wagner, 1855:236 BRAZIL: Amazon-
as (type locality, Barra do Rio Negro; holotype,
male, in Vienna Museum, collected 7 November
1825 by Johann Natterer). Pelzeln, 1883:113
BRAZIL: Amazonas (Barra do Rio Negro). Gard-
ner, 1993:15 listed.
Micoureus ochropus: Matschie, 1916:269 taxono-
my.
[Caluromys laniger] ochropus: Thomas, 1901b: 196.
Tate, 1939:163 GUYANA.
Mallodelphis lanigera ochropus: Miranda Ribeiro,
1936:355 BRAZIL: Amazonas (Barra do Rio Ne-
gro).
Caluromys laniger ochropus: Vieira, 1948:272
BRAZIL: Amazonas (Santa Cruz, Rio Eini, tribu-
tary of Rio Jurua; Rio Eini, Rio Jurua; Igarape
Grande, Rio Jurua).
Caluromys lanatus ochropus: Cabrera, 1958:3 (part,
not Wagner, 1842) vitalina, in synonymy only.
Baker, 1974:135 ECUADOR: Napo (Santa Cecil-
ia). Reig et al., 1977:201 PERU: Loreto (Yari-
nacocha [now in Ucayali]). Perez-Hernandez,
1989:375 VENEZUELA: Amazonas (Acanana;
Cacuri; Carlo Cotua, Boca; Parque Nacional Ya-
pacana, Cano; Cano Yagua, Cerro Cucurito; Rio
Cunucunuma; La Esmeralda; Raudal Cerguera; San
Carlos de Rio Negro; Campamanto, La Neblina).
D[idelphys] ornata Tschudi, 1844a: 14, 146, 151.
Tschudi, 1844b:250 PERU: (type locality, tropi-
cal forests of central Peru, between 10 and 20S,
i.e., eastern slope of Andes, in Pasco and Junin;
holotype not known to be in existence). Gardner,
1993:15 listed.
Didelphys ornata: Waterhouse, 1846:525. Tomes,
1860:268 ECUADOR.
Micoureus ornatus: Matschie, 1916:269 taxonomy.
Caluromys derbianus ornatus: J. A. Allen, 1900a:
189 taxonomy.
Ph[ilander] l[anatus] ornatus: Thomas, 1913b:358
classification.
Philander laniger ornatus: Thomas, 1920c:247
PERU: Cusco (Rio Comberciato, 2000 ft [= 625
m]). Thomas, 1927a:373 PERU: San Martin (Yu-
rac Yacu). Thomas, 1928a:265 PERU: Junin
(Chicosa [now in Ucayali], 454 m); Loreto (Masi-
sea [now in Ucayali]). Sanborn, 1949:277 PERU:
Loreto (Yarinacocha [now in Ucayali]; Agua Cal-
iente [now in Hudnuco]).
Caluromys laniger ornatus: Sanborn, 1 95 lb:3
PERU: Cusco (Hacienda Cadena).
Caluromys lanatus ornatus: Cabrera, 1958:3 clas-
sification. Baker, 1974:135 ECUADOR: Sucum-
bios (Santa Cecilia).
D[idelphys] bicolor Schinz, 1844:503, footnote no-
men nudum attributed to Tschudi as ms. name.
Philander cicur Bangs, 1898b: 161 COLOMBIA:
Magdelena (type locality, Santa Marta). Bangs,
1900:90 COLOMBIA: Magdalena (type locality,
northern slope of Sierra Nevada de Santa Marta,
above Pueblo Viejo [= Puebloviejo], 4000 ft [ =
1250 m]; holotype, female, skin and skull, MCZ
8114, collected 27 March 1898 by W W Brown;
Palomino, near Santa Marta). Gardner, 1993:15
listed.
Caluromys cicur: J. A. Allen, 1900a: 189. J. A. Allen,
1904d:4 18 COLOMBIA: Magdalena (Minca).
Micoureus cicur: Matschie, 1916:269 taxonomy.
Ph[ilander] l[aniger] cicur: Thomas, 1913b:358
distribution.
Philander laniger cicur: J. A. Allen, 1916a:202 CO-
LOMBIA: Cundinamarca (Tolima, near Bogota).
Anthony, 1923:4 COLOMBIA: Cundinamarca
BROWN: ATLAS OF NEW WORLD MARSUPIALS
(Fusagasuga; Cordillera del Rusio; Vioxa [= Vi-
ola]).
Philander laniger cerca de cicur: Mendez, 1953:82
VENEZUELA: Zulia (Kunana, Sierra de Perija).
Philander laniger cicus: Cabrera, 1958:2 incorrect
subsequent spelling for cicur in synonymy of Cal-
uromys lanatus cicur.
Caluromys lanatus cicur: Cabrera, 1958:2 classifi-
cation; meridensis in synonymy. Mendez, 1953:
82 VENEZUELA: Zulia (Kunana). Perez-Her-
nandez, 1989:377 VENEZUELA: Tdchira (Rio
Burgua; La Frfa, Colonia Agropecuaria del IAN);
Zulia (Kunana; Mision del Tocuco [= Tukuko]);
Barinas (La Lengueta); Apure (Nula); Merida
(Zea).
Philander laniger jivaro Thomas, 1913b:360 EC-
UADOR: Pastaza (type locality, Sarayacu. Rio Bo-
bonaza; holotype, male, skin and skull, BMNH
80.5.6.75; collected by Clarence Buckley; Canelos,
Rfo Bobonaza). Gardner, 1993:15 listed.
Micoureus juninensis Matschie, 1917:283 PERU:
Junin (type locality, La Merced, 1000 m; holotype,
female, skin and skull, ZMB 26754/55; collected
May 1908 by G. O. Schunke). Gardner, 1993:15
listed.
Micoureus meridensis Matschie, 1917:285 VENE-
ZUELA: Merida (type locality, "Montana de la Si-
erra bei Merida . . . ," 2500 m; holotype, female,
skin and skull, ZMB 26759/26760; collected 28
June 1903 by Sr. Bricefio). Gardner, 1993:15 list-
ed.
Micoureus bartletti Matschie, 1917:288 PERU: Lor-
eto (type locality, Rio Chamicuros, a tributary of
the Maranon between the Ribs Huallaga and Uca-
yali; holotype, juvenile female, skin and skull,
ZMB 4106/26758; collected by E. Bartlett). Gard-
ner, 1993:15 listed.
Micoureus cahyensis Matschie, 1917:288 BRAZIL:
Rio Grande do Sul (type locality, Rio Cahy [ =
Caf]; holotype, juvenile male, skin and skull, ZMB
26756/57; collected by Ehrenreich). Avila Pires,
1987:123 BRAZIL: Rio Grande do Sul (Rio
Cahy [= Caf]). Gardner, 1993:15 listed.
Philander calmensis: Vieira, 1955:347 incorrect
subsequent spelling of cahyensis Matschie, 1917.
[Micoureus) nattereri MaiscYnz, 1917:291 BRAZIL:
Mato Grosso (type locality, Caicara; name based
on a female in the Vienna museum collected 7 No-
vember 1825 by Johann Natterer). Gardner, 1993:
15 listed.
Mallodelphis laniger nattereri: Miranda Ribeiro,
1936:356 BRAZIL: Mato Grosso (Rio Jauru, up-
per Rio Paraguay; Caceres).
Philander laniger nattereri: Vieira, 1955:347 BRA-
ZIL: Mato Grosso (Rio Jauru [erroneously cited as
type locality]).
[^Mallodelphis lanigera hemiura Miranda Ribeiro,
1936:355 [misnumbered 455] (type locality un-
known; holotype, a skin and incomplete skull, Mu-
seu Nacional, Rio de Janeiro).
Mallodelphis lanigera modesta Miranda Ribeiro,
1936:356 BRAZIL: (type locality said to be in
Mato Grosso do Sul, probably Pantanal; holotype,
female, skin and skull, MNRJ; received from the
Zoological Gardens in Rio de Janeiro). Gardner,
1993:15 listed.
Mallodelphis lanigera vitalina Miranda Ribeiro,
1936:355 [misnumbered 455] BRAZIL: Minos
Gerais (type locality, Barra do Rio Paraopeba, Rio
Sao Francisco basin; holotype, female, skin and
skull, MNRJ, collected by Vital Rodriguez de Sou-
za). Gardner, 1993:15 listed.
Caluromys lanatus vitalina: Avila Pires, 1968:163
BRAZIL: Minas Gerais (Barra do Rio Paraopeba).
Holotype In alcohol, Museo de Ciencias Na-
turales, Madrid, no. 528; received 22 July 1789;
collected by Felix Azara (cf. Cabrera, 1916:1).
Type Locality Paraguay; Caazapa southeast
of Asuncion and Santa Maria, as restricted by
Matschie (1916:269).
Distribution (Fig. 4) Forests of northern and
central Colombia, northwestern and southern Ve-
nezuela, eastern Ecuador, eastern Peru, eastern
Bolivia, western and southern Brazil, southern
Paraguay, and Misiones in Argentina.
Additional Records COLOMBIA: Cesar;
Colonia Agricola de Caracolicito (Hershkovitz,
1942, USNM); Magdalena; Hacienda Cincinati,
Santa Marta (Carriker, 1941, USNM); Pueblo Bel-
lo (Hershkovitz, 1942, USNM); La Guajira; Mar-
imonda (= Las Marimondas) (Hershkovitz, 1943,
USNM); Sierra Negra near Villanueva (Hershko-
vitz, 1943, USNM); Cundinamarca; Sasaima
(Adolfo, 1982, USNM; Niceforo Maria, 1940,
USNM); Norte de Santander; Rio Zulia (Niceforo
Maria, FMNH); Arboledas, W of Cucuta (Nice-
foro Maria, 1948, FMNH); Toledo (Niceforo Ma-
ria, 1938, USNM); Sarcuta, not located (Niceforo
Maria, 1950, FMNH); Santander; San Gil (Nice-
foro Marfa, 1948, FMNH); Boyacd; Muzo
(Hershkovitz, 1952, FMNH); Meta; San Juan de
Arama (von Sneidern, 1957, FMNH); Parque de
La Macarena, Rfo Guapaya (von Sneidern, 1957,
FMNH); Huila; Villavieja (Hendrickson, 1950,
MVZ; Miller, 1949, MVZ; Pearson, 1950, MVZ);
San Isidro, Valle de Suaza (Del Aguila, 1971,
USNM); Naranjal, Valle de Suaza (Del Aguila,
1971, USNM); Pitalito (Hershkovitz, 1951,
FMNH); Tolima; Natagaima, left bank Rio Mag-
dalena (Niceforo Maria, 1926, USNM); ECUA-
DOR: Napo; Lago Agrio, left bank Rfo Aguarico
(Duellman and Fritts, 1969, KU); Marian (Timm,
1983, 1984, FMNH); Limoncocha (Emmons,
1977, USNM); Pastaza; Rio Pindo Yacu (Olalla,
1934, FMNH); Montalvo, Rfo Bobonaza (Olalla,
1932, FMNH); PERU: Amazonas; La Poza, Rfo
Santiago (Patton, 1979, MVZ); Bagua (= Bagua
Chica) (Graham, LSUMZ); Aguaruna Village,
Rfo Cenepa, near Huampami, not located (Patton,
1977, MVZ); Loreto; Santa Rita (Kalinowski,
10
FIELDIANA: ZOOLOGY
Fig. 4. Distribution of Caluromys lanatus: Venezuela and Colombia south through Ecuador, Peru, Bolivia, and
Paraguay; scattered throughout Brazil. = localities from unpublished records; = localities given in published
records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
11
1956, FMNH): Biological Station Pithecia
(Hershkovitz, 1980. FMNH): Santa Elena (Kali-
nowski. 1950. FMNH): Santa Maria. Mariscal Ra-
mos Castilla, not located (Soini. 1973. FMNH);
San Martin: Tarapoto. 28 km NE (Sanchez. 1983.
LSUMZ): Hudnuco; Ganso Azul (Sanborn. 1946.
FMNH): Plantacion San Huberto. Tingo Maria
(Nybukken. 1960. MVZ): Hacienda Buena Vista
(Heller. 1922. FMNH): Pasco; Oxapampa (Tuttle.
1964. USNM): Jum'n; Chanchamayo (Schunke.
FMNH: Schunke. 1986. USNM): San Ramon
(Anderson. 1914. FMNH): Cusco: Hacienda Villa
Carmen (Kalinowski. 1954. FMNH): Quincemil
(Kalinowski. 1953. FMNH); Madre de Dios: Al-
bergue Cusco Amazonico. 12 km E of Puerto
Maldonado (Warner. 1984. MVZ): Reserva Cusco
Amazonico. 14 km E of Puerto Maldonado
(Timm. 1990. KU): Aguas Calientes, 1 km below
Shintuya (Myers. 1984. UMMZ): VENEZUELA:
Merida; Montana de Edrama. Sierra de Merida
(Gabaldon. 1903. FMNH): Distrito Federal; El
Valle. Caracas (Barnes. 1939. LSUMZ): Amazon-
as; Tamatama (Brevington. 1967. USNM): Rio
Mawarinuma. Cerro de La Neblina (Gardner.
1984, USNM): BRAZIL: Acre; Sena Madureira
(1976. USNM): PARAGUAY: Canendiyu; Curu-
guaty. 13.3 km N (Myers. 1979. MZUM).
Caluromys philander (Linnaeus, 1758)
Tai-ibi Brasiliensibus: Marcgrave. 1648:223.
Mus africanus Kayopolin dictus mas.. Seba. 1734:49.
Tlaqitazin s. tai-ibi Brasiliensibus Seba. 1734:57.
Philander africanus: Brisson. 1762:212 description
based primarily on Mus africanus Kayopolin dictus
of Seba: work rejected for nomenclatural purposes
(International Commission on Zoological Nomen-
clature. 1998:64).
Philander africanus Schreber. 1777:541 der Kayo-
pollin: name ex Brisson. 1762.
Cayopollin: Buffon. 1763:350. not the coyopollin of
Hernandez (1651. Nov. Hisp.. p. 10. a Marmosa).
Didelphis Cajopolin Mullen 1776:35 name based
primarily on the caxopollin of Buffon. Gardner.
1993:15 listed.
Didelphys cayopollin Schreber. 1777:544 (der Koy-
oppolin). 589 Guianas ["Mexico" based on the
coyopollin of Hernandez, a Marmosa. but Buffon "s
caxopollin originated in the Guianas: see E. Geof-
frey. 1803. below]. Gardner. 1993:15 listed.
[Didelphis] Cayopollin: Gmelin. 1788:106 the cay-
opollin of Button.
Didelphis Caxopollin: Shaw. 1800:478 "Mexican"
opossum, ex Buffon.
Didelphis cayopollin: E. Geoffroy. 1803:142
FRENCH GUIANA: male specimens, basis of Buf-
fon 's (1763:350) "Cayopollin." and female speci-
men in spirits, from the Dutch collections [= of
Seba?].
S[arigua] caxopollin: Muirhead. 1819:429 part, new
combination.
Philander Caxopollin: Burmeister. 1856:76. pi. 14
(animal) part. BRAZIL: GUIANAS.
[Sarigua] caxopollin. Muirhead. 1919:429.
[Didelphis] Philander Linnaeus. 1758:54. Gardner.
1983:15 listed, synonymy.
Didelphis philander: Latreille. 1803:304 philander
de Surinam. Temminck. 1827:43 SURINAME:
FRENCH GUIANA. Schomburgk. 1840:344
GUYANA. Jentink. 1887b:301 BRAZIL. Jentink.
1888b:221 SURINAME.
Didelphis (Philander) philander: J. A. Allen and
Chapman. 1893:203 TRINIDAD AND TOBA-
GO.
Didelphxs philander: Schreber. 1777:541. Goldfuss.
1809:215 SURINAME. Waterhouse. 1841:102
SURINAME: GUYANA. Cabanis and Schom-
burgk. 1848:777 GUYANA. Wagner. 1850:130
BRAZIL: Amazonas (Borba. Rio Madeira): Rio
Branco [now Roraima] (Rio Branco). Pelzeln.
1883:111 BRAZIL: Amazonas (Borba): Rio
Branco [now Roraima] (Forte do Rio Branco).
Thomas. 1888b:337 GUYANA: SURINAME:
BRAZIL: Amazonas (Manaus). Thomas. 1893:
10 TRINIDAD AND TOBAGO.
D[idelphys] Philander: Wagner. 1842:358.
Didelphxs (Philander) philander: Menegaux. 1902:
496 FRENCH GUIANA: (Saint Georges).
Gamba philander: Liais. 1872:330 BRAZIL.
Metachirus philander: Ihering. 1894:1 1 BRAZIL:
Sao Paulo.
Caluromys philander: J. A. Allen. 1900a: 189 part.
GUIANAS: BRAZIL (northeastern): part. VENE-
ZUELA. Robinson and Lyon. 1901:138 VENE-
ZUELA: Distrito Federal (San Julian). G. M. Al-
len. 1902:92 VENEZUELA: Nueva Esparto (Isla
Margarita). Thomas. 19 10b: 189 GUYANA: Pom-
eroon-Supenaam (Supenaam River). Thomas.
191 lb: 143 SURINAME: (type locality): holotype
history. Miranda Ribeiro. 1935:39 BRAZIL: Rio
de Janeiro (Teresdpolis). Miranda Ribeiro. 1936:
358 BRAZIL: Para (Para: Rio Tocantins): Mato
Grosso (Salto Utiarity: Rio Papagaio [= Sauerui-
na]: Tapirapuan [= Tapirapua]): Maranhao: Rio de
Janeiro (Teresdpolis): Santa Catarina. Davis.
1945a: 1 19 BRAZIL: Rio de Janeiro (Teresdpol-
is). Davis. 1945b:280 BRAZIL. Davis. 1947:2
BRAZIL: Rio de Janeiro (Teresdpolis). Anduze.
1956-57:8 VENEZUELA: Bolivar (Suapure. left
bank Rio Caura: Valle del Caura: Ciudad Guyana):
Distrito Federal (La Guaria). Carvalho. 1960:
122 BRAZIL: Para (Conceicao do Araguaia).
Handley. 1976:5 VENEZUELA: Amazonas (Be-
len: Boca Mavaca: Cario Culebra: Puerto Ayacu-
cho: San Juan de Manapiure: Tamatama): Sucre
(Cumana: Manacal): Miranda (Curupao); Distrito
Federal-Miranda (Instituto Venezolano de Investi-
gaciones Cientfficas [= IVIC]: Los Venados): Fal-
con (Mirimire: Urama): Carabobo (Montalban):
Bolivar (Los Patos): Monagas (San Agustih). Reig
et al.. 1977:212 VENEZUELA: Miranda (Baru-
ta): Guarico (Dos Caminos): Aragua (La Horqueta.
Tiara: Rancho Grande): Distrito Federal (Los Ven-
ados): TRINIDAD AND TOBAGO: Simla (North-
12
FIELDIANA: ZOOLOGY
em Range, not located). Husson, 1978:9 SURI-
NAME: Nickehe (Sipaliwini Airstrip); Commewi-
jne (Marienburg; Clevia; Nieuw Amsterdam;
Combe); Paramaribo (Culturutuin, Paramaribo;
Ma Retrate; Pan van Wanica). Alkins, 1979:175
TRINIDAD AND TOBAGO. Eisenberg and
O'Connell, 1979:187 VENEZUELA: Miranda-
Guatopo (Parque Nacional Guatopo). Charles-
Dominique et al., 1981:355 FRENCH GUI-
ANA: (Cabassou). Da Fonseca and Kierulff,
1988:109 BRAZIL: Minas Gerais (Rio Doce
State Park [= Parque Estadual do Rio Doce]).
Stallings, 1989:176 BRAZIL: Minas Gerais
(Rio Doce State Park [= Parque Estadual do Rio
Doce]). Julien-Laferriere and Atramentowiez,
1990:405 FRENCH GUIANA: (Cabassou, Piste
de Saint-Elie). Gardner, 1993:15 TRINIDAD
AND TOBAGO: Tobago (Tobago). Julien-Laferri-
ere, 1997:251 FRENCH GUIANA: Piste de
Saint-Elie Ecerex Station. Pires et al., 1999:41
BRAZIL; Rio de Janeiro (Poco das Antas, Reserva
Biologica).
Caluromys philander philander: Vieira, 1953:220
BRAZIL: Alagoas (Mangabeiras). Avila Pires,
1958:2 BRAZIL: Para (Utinga, Belem de Para).
Cabrera, 1958:4 venezuelae in synonymy only.
Carvalho, 1962:284 BRAZIL: Amapd (lower Rio
Araguari, Macapa; Mazagao, Rio Maraca; Vila Vel-
ha do Cassipore, Oiapaque). Pereira and Machado
filho, 1968:283 BRAZIL: Amazonas (km 46, Es-
trada AM-1). Pine, 1973b:54 BRAZIL: Para (Be-
lem). Husson, 1978:9 SURINAME: Paramaribo
(Pad van Wanica). Patterson, 1992:8 BRAZIL:
Para (Cameta); Amazonas (Itacoatiara).
Calurosmys philander philander: Avila Pires, 1964:
8, 1 1 incorrect subsequent spelling of Caluromys
philander philander. Vieira, 1953:220 BRAZIL:
Amazonas (km 50, Manaus-Itacoatiara RR).
Philander philander: Miranda Ribeiro, 1914:48
BRAZIL: Mato Grosso (Salto Utiarity; Rio Papa-
gaio [= Saueruina]; Tapirapoan [= Tapirapua], Rio
Sepotuba). Sanderson, 1949:788 SURINAME.
[Philander philander] philander: Cabrera, 1919:34
classification.
Philander philander philander: Vieira, 1955:347
BRAZIL: Amazonas (Lago do Baptista); Para (Ca-
meta; Caxiricatuba; Foz do Curua); Maranhao
(Miritiba); Alagoas (Mangabeiras); Mato Grosso
(Rio Papagaio; Tapirapoan [= Tapirapua], Rio Se-
potuba); Bahia; Espirito Santo; Rio de Janeiro.
Carvalho, 1960:122 BRAZIL: Para (Conceicao
do Araguaia).
V[sic = D(idelphis)] Flavescens Brongniart, 1792:
115 FRENCH GUIANA: (type locality, Cay-
enne).
Didelphys dichura Wagner, 1842:358 BRAZIL: Sao
Paulo (type locality, Ipanema; holotype, male, skin,
Vienna Museum, collected 14 August 1819 by Jo-
hann Natterer). Waterhouse, 1846:492 BRAZIL.
Gardner, 1993:15 listed.
Didelphys dichrura: Wagner, 1850:134 incorrect
subsequent spelling of dichura Wagner, 1842
BRAZIL: Sao Paulo (Ipanema). Pelzeln, 1883:
1 12 BRAZIL: Sao Paulo (Ipanema; Itarare).
Philander dichura: Burmeister, 1856:76 BRAZIL:
Bahia.
D[idelphis] dichura: Matschie, 1916:263 BRAZIL:
Bahia; skull no. 25306 in Berlin Zoological Mu-
seum erroneously indicated as holotype.
[Caluromys] dichura: Matschie, 1916:269 BRA-
ZIL: Sao Paulo (type locality, Ipanema).
[Philander philander] dichrura: Cabrera, 1919:34
incorrect subsequent spelling of dichura Wagner,
1 842:358 classification.
Philander philander dichrura: Pohle, 1927:241 in-
correct subsequent spelling of dichura Wagner,
1842 BRAZIL: Para (Santa Gertrudes, Araras);
Sao Paulo (Jacarehy [= Jacarei]); holotype cor-
rectly indicated as in Vienna Museum, not Berlin
(cf. Matschie, 1916:263).
Caluromys philander dichrurus: Vieira, 1949:345
incorrect subsequent spelling of dichura Wagner,
1842:358 BRAZIL: Sao Paulo. Vieira, 1955:
347 BRAZIL: Sao Paulo (Sao Sebastiao; Piquete;
Tambi; Monte Alegre [= Monte Alegre do Sul]).
Avila Pires and Gouvea, 1977:10 BRAZIL: Rio
de Janeiro Minas Gerais (Almoxarifado, Parque
Nacional do Itatiaia); Sao Paulo (Ipanema).
Didelphis affinis Wagner, 1842:358 BRAZIL: Mato
Grosso (type locality, Rio Jurare; holotype, female,
skin, Vienna Museum; collected 14 March 1829 by
Johann Natterer). Wagner, 1850:136 BRAZIL:
Mato Grosso. Pelzeln, 1883:1 1 1 BRAZIL: Mato
Grosso (Rio Jurare). Gardner, 1993:15 listed.
Caluromys affinis: J. A. Allen, 1900a: 189 classifi-
cation.
[Philander philander] affinis: Cabrera, 1919:34
classification.
Caluromys philander affinis: Cabrera, 1958:3 clas-
sification.
Philander marsupialis: Gray, 1843:101 BRAZIL.
Didelphys macrura Pelzeln, 1883:1 1 1 BRAZIL:
Amazonas (Borba), Natterer manuscript name in
synonymy of Didelphys philander; nomen nudum.
D[idelphys] longicaudata Pelzeln, 1883:111 Natter-
er manuscript name in synonymy of Didelphys phi-
lander; nomen nudum.
[Didelphys (Philander)] longicauda Trouessart, 1898:
1237 incorrect subsequent spelling for longicau-
data Pelzeln (nomen nudum), in synonymy of P.
philander.
Didelphis {Philander) trinitatis Thomas, 1894a:438
TRINIDAD AND TOBAGO: Port-of-Spain (type
locality, Botanic Garden; holotype, female in al-
cohol, BMNH, collected by J. H. Hart).
Philander trinitatis: J. A. Allen and Chapman, 1897:
26 TRINIDAD AND TOBAGO: St. George
(Caura). Rendall, 1897:342 TRINIDAD AND
TOBAGO.
Caluromys trinitatis: J. A. Allen, 1900a: 189. Vesey-
Fitzgerald, 1936:6 TRINIDAD AND TOBAGO:
western end of northern range. Beebe, 1952:181
TRINIDAD AND TOBAGO (Aruma Valley, not
located). Handley, in Torrealba, 1958:97 VENE-
ZUELA: Guarico (Municipio Parapara).
Didelphys trinitatis: Elliot, 1907:6 TRINIDAD
AND TOBAGO: St George (Caura).
Caluromys trinitatis venezuelae Thomas, 1903:493
VENEZUELA: Sucre (type locality, Ypure, near
Cumana, 2350 ft [= 700 m]; holotype, old female,
skin and skull, BMNH 0.5.1.57, collected 8 March
1899 by E. Andre). Gardner, 1993:15 listed.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
13
Caluromys trinitatis leucurus Thomas, 1904b:36
VENEZUELA: Bolivar (type locality, Maripa, Rio
Caura. lower Rio Orinoco: holotype. male, skin and
skull. BMNH 4.5.7.42. collected 26 October 1903
by S. M. Klages). J. A. Allen. 1904d:337 VE-
NEZUELA: Bolivar (Ciudad Bolfvar). Gardner.
1993:15 listed.
Holotype Name based on the tlaquazin or
tai-ibi brasiliensibus of Seba; the type became
part of the Lidth de Jeude collection subsequently
acquired by the British Museum. It is now num-
bered 414 (Thomas, 1892, Proc. Zool. Soc. Lon-
don, 1892:314).
Type Locality Suriname.
Distribution (Fig. 5) Venezuela, Trinidad
and Tobago. Guyana, Suriname. French Guiana,
and Brazil.
Additional Records VENEZUELA: Falcon;
La Pastora (Smithsonian Expedition, 1967.
USNM): Guarico; Municipio Parapara. Rosco
(Torrealba, 1956. USNM); San Juan de los Mor-
ros (Torrealba, 1957, FMNH); Camp Rangel, not
located (de la Barena. 1959, USNM); TRINIDAD
AND TOBAGO: St. Andrew; Mt. Harris (Wonder.
1947, FMNH); GUYANA: Demerara-Mahaica;
Hyde Park (Warren, FMNH); Cuyuni-Mazaruni;
Essequibo, Oko Mts. (Blake, 1937, FMNH); SU-
RINAME: Saramacca; La Poule (Hershkovitz.
1962, FMNH); BRAZIL: Para; Braganca (Belem
Virus Laboratory, 1968. USNM); Santarem. Mo-
jui dos Campos (1975, USNM); Tapaiuna, right
bank Rio Tapajos (Olalla, 1959, FMNH); Alta-
mira, left bank Rio Xingu (Carleton, 1986.
USNM; Santos, 1976, USNM); Fordlandia. right
bank Rio Tapajos (Olalla, 1958, 1959. FMNH);
Maraba (Peterson, 1975, USNM); Maranhao;
Carolina (Snethlage, 1925, FMNH); Mato Gros-
so; Serra do Roncador, 264 km N of Xavantina
[= Chavantina] (Pine, 1968, USNM); Amapd;
Serra do Navio (1968, USNM); Minas Gerais;
Passos (1946, USNM).
Genus Caluromysiops Sanborn, 1951
Caluromysiops Sanborn. 1951a:473. Gardner. 1993:
15 listed.
Type Species Caluromysiops irrupta Sanborn.
Distribution Southeastern Peru and western
Brazil. Simonetta's (1979) record from southeast-
ern Colombia, cited by Cuervo Diaz et al. (1986)
and de Vivo and Gomes (1989), is probably based
on a transported captive.
Caluromysiops irrupta Sanborn, 1951
Caluromysiops irrupta Sanborn. 1951a:473. Sanborn.
1951b:3 PERU: Cusco (Quincemil). Grimwood.
1968:1 PERU: Madre de Dios (Rio Alto Madre
de Dios. Parque Nacional del Manu). Simonnetta.
1979:247 COLOMBIA: Amazonas (Leticia). Ter-
borgh et al.. 1984:1 PERU: Madre de Dios (Co-
cha Cashu Biological Station. Parque Nacional del
Manu). Cuervo Diaz et al., 1986:472 COLOM-
BIA: Amazonas (Leticia). De Vivo and Gomez.
1989:310 COLOMBIA: Amazonas (Leticia):
PERU: Madre de Dios (Cocha Cashu Biological
Station. Parque Nacional del Manu: Itahuania):
Cusco (Quincemil. Quispicanchis Province); BRA-
ZIL: Rondonia (Upper Rio Jani). Gardner. 1993:
15 listed.
Holotype Male, skin and skull, FMNH
68336; collected 11 July 1950 by Celestino Kal-
inowski.
Type Locality Quincemil, Quispicanchis
Province, Cusco, Peru; 680 m.
Distribution (Fig. 6) Amazonas in Colombia,
western Brazil, and southeastern Peru.
Genus Glironia Thomas, 1912
Glironia Thomas. 1912b:239. Reig. 1955:127 re-
ferred to subfamily Microbiotheriinae. Gardner.
1993:16 listed.
Type Species Glironia venusta Thomas, by
original designation.
Distribution Ecuador, Peru. Bolivia, and the
Amazon basin of Brazil.
Glironia venusta Thomas, 1912
Glironia venusta Thomas. 1912b:240. Thomas.
1912c:47 BOLIVIA: La PazCochabamba
(Yungas region). Cuervo-Dias et al., 1986:472
COLOMBIA: Putumavo [now Amazonica]. Da Sil-
va and Langguth, 1989:873 BRAZIL: Para (Rio
Mapuera). Emmons. 1990:72-73 BOLIVIA: La
Paz (Rio Machariapo). Pacheco et al., 1993:5
PERU: Madre de Dios (Cocha Cashu). Tarifa and
Anderson. 1997:1 12 PERU: Madre de Dios
(Parque Nacional Manu); BOLIVIA: La Paz (Val-
ley de Machariapo, Rio Machiriapo. Chaquinayo):
Santa Cruz (6 mi E on road from San Javier to El
Puente; Meseta de Huanchaca. Parque Nacional
Noel Kempff Mercado). Anderson. 1997:140
BOLIVIA: La Paz (Rio Machariapo: "Yungas"):
Santa Cruz (San Javier, 12 km S).
Glironia venusta: Cabrera. 1958:5 part.
Glironia aequatorialis Anthony. 1926:1 ECUA-
DOR: Napo (type locality, outlets of Lagarto Co-
cha, on the Rio Aguarico above its confluence with
14
FIELDIANA: ZOOLOGY
Caluromys philander
1500 km
lliLl
90
80
70
60
50
40
30
20"
10
Fig. 5. Distribution of Caluromys philander: Venezuela, Trinidad and Tobago, east through Guyana, Suriname,
French Guiana, and south through Brazil. = localities from unpublished records; = localities given in published
records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
15
Caluromysiops irrupta
1500 km
30
Fig. 6. Distribution of Caluromysiops irrupta: southeastern Colombia, western Brazil, and southeastern Peru.
o = localities from unpublished records; = localities given in published records.
the Rio Napo, Napo; holotype, female, skin and
skull, AMNH 71393, collected 15 January 1926 by
Olalla and Sons). Marshall, 1978:1 ECUADOR:
Pastaza (Montalvo, Rio Bobonaza); Pasco (Pozu-
zo). Gardner, 1993:16 listed.
Glironia criniger Anthony, 1926:2 PERU: Loreto
(type locality, junction of Rio Curaray and Napo;
holotype, female, skin and skull, AMNH 71394,
collected 5 December 1925 by Olalla and Sons).
Gardner, 1993:16 listed.
Holotype Male, skin and skull, BMNH
12.1.15.7; collected April 1908 by L. Egg.
Type Locality Pozuzo, Pasco, Peru, 800 m.
Distribution (Fig. 7) Ecuador, Peru, Bolivia,
and the Amazon basin of Brazil.
Additional Records PERU: Loreto; Quebra-
da Vanilla (= Vainilla), 10 km SSW mouth of Rio
Napo on east bank (Maxwell, 1983, LSUMZ);
BOLIVIA: Tarija; Fabrica de Papel, N Cuyam-
buyo, not located (Creighton, 1980, MVZ); Ha-
cienda Conzulman, N Cuyambuyo, not located
(Creighton, 1980, MVZ).
Subfamily Didelphinae Gray, 1821
Genus Chironectes Illiger, 1811
Chironectes Illiger, 1811:76. Gardner, 1993:16 list-
ed.
Memina Fischer, 1813:579 type species, Lutra
Meminna [sic] Boddaert [= Chironectes minimus
Zimmerman]. Gardner, 1993:16 listed.
Sarigua Muirhead, 1819:429 (part) S. memmima
listed among included species.
Cheironectes Gray, 1821:308 emendation of Chi-
ronectes Illiger.
Cheronectis Fleming, 1822:212 name in text, lapsus
for Chironectes Illiger, with D. palmata only spe-
cies mentioned.
Gamba Liais, 1872:329 included species: Gamba
palmata, G. aurita brasiliensis Liais [= Didelphis
marsupialis azarae Temminck], Gamba aurita vir-
giniana [= Didelphis marsupialis virginiana Kerr],
Gamba opossum [ Philander opossum Linnaeus],
Gamba philander [= Caluromys philander Linnae-
us]. Hershkovitz, 1949:12 type species Gamba
palmata Liais [= Chironectes minimus Zimmer-
man], by designation. Gardner. 1993:16 listed.
Chironeytes Goeldi and Hagmann, 1904:100 incor-
16
FIELDIANA: ZOOLOGY
Fig. 7. Distribution of Glironia venusta: Ecuador, Peru, northern Bolivia, and the Amazon basin of Brazil, o
localities from unpublished records; = localities given in published records.
rect subsequent spelling of Chironectes Illiger,
1811, in combination with palmata Cuvier.
Type Species Lutra minima Zimmermann by
monotypy.
Distribution Neotropical forested parts of
Central and South America from southern Mexico
to northeast Argentina.
Chironectes minimus (Zimmermann, 1780)
Saricovienne Buffon, 1755, Hist. Nat., 319 descrip-
tion of the "La Plata" saricovienne of Thevet,
1558, Les singularites de la France antarctique au-
trement nombree Amerique . . . , pp. 107, 108.
Petite loutre d'eau douce de Cayenne Buffon, 1776:
159, pi. 22 FRENCH GUIANA.
Small Guiana otter Smellie, 1780:237, pi. 69.
Latra minima Zimmermann, 1780:317 incorrect
subsequent spelling of generic name. Gardner,
1983:16 listed.
Lutra minima: Cabrera, 1958:43 incorrect subse-
quent spelling of generic name of Latra minima in
synonymy of Chironectes minimus.
Must[ela]. (Lutra) minima: Wolf, 1796:289 classi-
fication.
Chironectes minimus: Lydekker, 1887:329. Thomas,
1888b:368 COLOMBIA: Antioquia (Medellfn);
ECUADOR; GUYANA; BRAZIL: Para; Rio de
Janeiro; Rio Grande do Sul (San Lorenzo [= Sao
Lourenco do Sul]). De Winton, 1896:513 EC-
UADOR: Pichincha (Nanegal, west of Quito; trib-
utary of the Rio Guaillabamba, north of Quito). J.
A. Allen, 1900c:219 PERU: Puno (Juliaca [error
for Inca Mines, Rio Inambari]). Bangs, 1900:90
COLOMBIA: Magdalena (Santa Marta). Thomas,
1900a:549 BRAZIL: Parana (Palmeira). Thom-
as, 1 90 lc:536 BRAZIL: Minas Gerais (Rio Jor-
dao). J. A. Allen, 1912a:74 COLOMBIA: Valle
del Cauca (Juanchito; Palmira). Osgood, 1914b:
150 PERU: San Martin (Moyobamba). Miiller,
1914:8 BRAZIL: Para (Peixe-boi, Belem). Ber-
toni, 1914:68 PARAGUAY: (Yaguarasapa). J. A.
Allen, 1916a:202 COLOMBIA: Valle del Cauca
(Juanchito; Palmira; Barbacoas [now in Narino];
Bagado [now in Choco]; Rio Frio). Lonnberg,
1921:67 ECUADOR: Pichincha (Gualea, 3000 ft
[= 930 m]). Bertoni, 1923:51 PARAGUAY: Cen-
tral (Bahia de Asuncion); Itapud (Pirayii-hi). Mi-
randa Ribeiro, 1935:39 BRAZIL: Rio de Janeiro
(Teresopolis). Miranda Ribeiro, 1936:390 BRA-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
17
ZIL: Santa Catarina (Rio Novo do Itapocii): Rio
de Janeiro (Jacarepagua: Colonia Alpina. Tereso-
polis: Novo Friburgo): Para (Ilha das On^as. Be-
lem). Bertoni. 1939:6 PARAGUAY: Central
(Asuncion): Aha Parana (Puerto Bertoni): AR-
GENTINA: Misiones (Iguasu). Vieira. 1949: 347
BRAZIL: Minas Gerais; Goids: Rio de Janeiro:
Para. Crespo. 1950:5 ARGENTINA: Misiones
(San Javier. Rio Paraguay; Rio Parana-Guazu. 15
km from the banks of the Rio Parana). Sanborn.
1951b:3 PERU: Cusco (Hacienda Cadena): San
Martin (Movobamba): Hudnuco (Pozuzo [now in
Pasco]). Sanborn. 1953:4 PERU: Puna (Yalle
Grande. Sandia. not located). Kuhlhorn. 1953a:7
BRAZIL: Mato Grosso (Rio Pardo). Kuhlhorn.
1953b:121 BRAZIL: Mato Grosso (Ribeirao
Pedra [= Ribeirao das Pedras] near Rancho Pedra.
Rio Pardo). Mondolfi and Medina. 1958:144 VE-
NEZUELA: Aragua (Rio Cumboto: Rio Ocumare:
Parque Nacional de Rancho Grande: El Consejo:
Rio Periquito): Carabobo (Aguacatacal): Guarico
(Rio Guarico). Reig. 1958:262 (footnote) AR-
GENTINA: Entre Rfos (fossil). Lyne. 1959:8-1
PERU: Junm (Chanchamayo): BRAZIL: Rio
Grande do Sid (Sao Lourenco do Sul). Guallart.
1962:157 PERU: Amazonas (upper Rio Mara-
non). Schaldach. 1965:130 MEXICO: Oaxaca
(north of Matias Romero. Rio Sarabia. 1800 m).
Ruschi. 1965:3 BRAZIL: Espirito Santo (Santa
Teresa). Grimwood. 1968a:9 PERU: San Martin
(Movobamba): Puno (Carabaya): Hudnuco (Rio
Pachitea: Tingo Maria): Juni'n (Tarma): Ucayali
(Pucallpa: Rio Curanja): Madre de Dios-Cusco
(Parque Nacional del Manu): Madre do Dios
(Manu): Cusco (Quispicanchi). Yunis et al.. 1972:
265 COLOMBIA: Meta (Rio Ocoa. Villavicen-
cio- Acacias Road). Handley. 1976:9 YENEZUE-
LA: Merida (Santa Rosa): Yaracuy (Minas de
Aroa): Monagas (San Agusti'n). Reig et al.. 1977:
212 VENEZUELA: Aragua (Rio Choroni). Alk-
ins. 1979:1 TRINIDAD AND TOBAGO, de Frei-
tas et al.. 1979:613 BR.AZIL: Rio Grande do Sul.
Eisenberg et al.. 1979:187 VENEZUELA: Mi-
randa-Guarico (Parque Nacional Guatopo). Tvn-
dale-Biscoe. 1980:712 COLOMBIA: Volte 'del
Cauca (Cali). Azevedo et al.. 1982:95 BRAZIL:
Santa Catarina (Corupa: Blumenau: Camboriu:
Brusque). Wilson. 1983:443 COSTA RICA: Pun-
tarenas (Rincon. Corcovado National Park). Ter-
borgh et al.. 1984:1 PERU: Madre de Dios (Co-
cha Cashu Biological Station. Parque Nacional del
Manu). Alho. Pereira. and Paula. 1986:446 BRA-
ZIL: Distrito Federal (Parque Nacional de Brasi-
lia). Gardner. 1988: 700 VENEZUELA: Amazon-
as (Base Camp [= Campamento], Cerro de La
Neblina). Timm et al.. 1989:16 COSTA RICA:
Heredia (Puerto Viejo). Albuja. 1992:126 EC-
UADOR: Los Rfos (Jauneche). Pacheco et al..
1993:5 PERU: Cusco (Hacienda Villa Carmen:
Hacienda Erika). Yensen et al.. 1994:406
BOLIVIA: La Paz (Apolo). Raseot and Albuja.
1994: 176 ECUADOR: Pastaza (Mera). Heinen
and Chebez. 1 99^: 19 ARGENTINA: Misiones
(Parque Nacional Isuazui. Voss et al.. 2001:20
FRENCH GUIANA (Paracou).
Cheironectes minimus: Jentink. 1887:302 incorrect
subsequent spelling of Chironectes Illiger. 1811
SURINAME.
[Lutra] menina: Boddaert. 1784:168 incorrect sub-
sequent spelling of minima Zimmerman, 1780.
Did[elphis] memina: Cuvier. 1798:125 incorrect
subsequent spelling of Lutra menina Boddaert.
178-1 FRENCH GUIANA (type locality).
Didelphis memina: E. Geoffroy. 1803:144 incorrect
subsequent spelling of menina Boddaert. 178-1
FRENCH GUIANA: two specimens in Paris Mu-
seum.
Lutra memina: Desmarest. 1816:546 incorrect subse-
quent spelling of menina Boddaert. 1784 palmata
E. Geoffroy a synonym.
Chironectes memina: Cope. 1889:130 incorrect sub-
sequent spelling of menina Boddaert. 1784 BRA-
ZIL: Mato Grosso (Chapada).
Lutra memia: Desmarest. 1803:147 incorrect sub-
sequent spelling of menina Boddaert. 178-1 GUY-
ANA.
Lutra memmina: Desmarest. 1804:507 incorrect
subsequent spelling of menina Boddaert. 1784.
S[arigua] memmima: Muirhead. 1819:429 new
combination.
Chironectes menima: Pohle. 1927:243 (in text) in-
correct subsequent spelling of menina Boddaert.
178-4 VENEZUELA: Distrito Federal (Caracas):
PERU: Cusco (Marcapata).
Chironectes minima minima: Krumbiegel. 1940:66
VENEZUELA: Aragua (Choroni: Maracay); CO-
LOMBIA: Cundinamarca (La Vega. E of Bogota).
Chironectes minimus minimus: Avila Pires. 1958:5
BRAZIL: Para (Ilha das Oncas. Belem). Goodwin
and Greenhall. 1961:201 TRINIDAD AND TO-
BAGO. Carvalho. 1965:251 BRAZIL: Sao Paulo
(Boraceia). Carvalho and Tocheton. 1969:217
BRAZIL: Para (Utinga). Baker. 1974:135 EC-
UADOR: Sapo (Santa Cecilia). Crespo. 1974:-!
ARGENTINA: Misiones (Pinalitos: Puerto Pirav:
Rio Urugua-i). Massoia. 1976:30 ARGENTINA:
Misiones (Department of General Belgrano. not lo-
cated: Iguazii [Iguasii]: Cuarto Rio Victoria. Rt. 14.
km 265. not located). Massoia et al.. 1987:111
ARGENTINA: Misiones (Rio Urugua-i: San Javier:
Arrovo Garupa: Arrovo Piray Guazii: Arroyo Tac-
uara.' Cerro Azul). Gardner. 1988:700 VENE-
ZUELA: Amazonas (Base Camp [= Campamento].
Cerro de La Neblina). Perez-Hernandez. 1989:
37-1 VENEZUELA: Amazonas (Rio Cunucunu-
ma: Campamento La Neblina: Parla Chonto. not
located): Bolivar (Santa Maria del Erebato). An-
derson et al.. 1993: 19 BOLIVIA: Santa Cruz (Rio
Pitisama. Parque Nacional de Amboro). Anderson
1997:141 BOLIVIA: Santa Cruz (Rio Pitisama.
Parque Nacional de Amboro): Beni (Campamento
6 de Agosto): La Paz (Apolo: La Reserva).
M[ustela] (Lutra) paraguensis Kerr. 1792:172 PAR-
AGUAY: (type locality. Rio de la Plata" [i.e.. the
region comprising northeastern .Argentina and bor-
dering parts of Uruguay and Paraguay with type
locality, the latter by patrinomy]): name based on
the saricovienne of Smellie"s translation of Buffon.
Gardner. 1993:16 listed.
M[ustela] (Lutra) guianensis Kerr. 1792:174
FRENCH GUIANA: (type locality. Cayenne).
Gardner. 1 993: 1 6 listed.'
18
FIELDIANA: ZOOLOGY
L[utra] gujanensis Link, 1795:84 FRENCH GUI-
ANA: (type locality, by patrinomy; holotype, the
petite loutre d'eau douce de Cayenne of Buffon,
by implication). Gardner, 1993:16 listed.
Lutra saricovienna Shaw, 1800:447 FRENCH GUI-
ANA: (type locality, Cayenne). Gardner, 1993:16
listed.
[Mustela] cayennensis Turton, 1802:58 FRENCH
GUIANA: (type locality, Cayenne); Cayenne otter,
name based on the petite loutre d'eau douce de
Cayenne of Buffon. Gardner, 1993:16 listed.
Didelphis palmata Daudin in Lacepede, 1 799: 1 52
name based on the petite loutre d'eau douce de
Cayenne of Buffon.
Didelphys palmata: Burmeister, 1854:133 BRAZIL:
Rio de Janeiro (Novo Friburgo).
Chironectes palmata: Gray, 1827:135 caption for
colored plate.
Gamba palmata: Liais, 1872:329 BRAZIL.
Cheironectes Palmata: Burnett, 1830:351 incorrect
subsequent spelling of Chironectes Illiger, 1811
classification.
Chironeyctes palmata: Goeldi and Hagmann, 1904:
100 incorrect subsequent spelling of Chironectes
Illiger, 1811 BRAZIL: Para (Ilha do Mosqueiro,
near Belem).
Chironectes palmatus: Hensel, 1873:121 BRAZIL
Rio Grande do Sul. Ihering, 1893:101 BRAZIL
Rio Grande do Sul (Costa da Serra, not located
Serra do Herval [Erval]). Ihering, 1894:12 BRA-
ZIL: Sao Paulo. Goeldi, 1894:465 BRAZIL: Rio
de Janeiro (Serra dos Orgaos). Schirch, 1932:84
BRAZIL: Rio de Janeiro (Teresopolis).
Mustela lutris Daudin in Lacepede, 1799:164 name
based on the saricovienne of Buffon.
Chironectes variegatus Illiger, 1815:107 substitute
name for Lutra minima Zimmermann. Olfers, 1818:
206 substitute name for Lutra minima Zimmer-
mann. Burmeister, 1856:74 BRAZIL: Rio de Ja-
neiro (Novo Friburgo). Sclater, 1872:702 CO-
LOMBIA: Antioquia (Medellfn). Pelzeln, 1883:
118 BRAZIL: Sao Paulo (Ipanema). Winge,
1893:850 BRAZIL: Minas Gerais (Lagoa Santa).
Gardner, 1993:16 listed.
Ch[ironectes] variegates: Cabanis and Schomburgk,
1848:778 GUYANA.
Chironectes yapock Desmarest, 1820:261 replace-
ment name for "memina" [= minima] Zimmer-
mann. Gray, 1827:35 FRENCH GUIANA: (banks
of Yapock [= Oyapock] River). Ogilby, 1836:56
BRAZIL. Gardner, 1993:16 listed.
Didelphys Yapock: Waterhouse, 1841:112.
Chironectes Langsdorffii Boitard, 1845:288 BRA-
ZIL: Rio de Janeiro (type locality, near Rio de Ja-
neiro). Gardner, 1993:16 listed.
Chironectes panamensis Goldman, 1914:1 PANA-
MA: Darien (type locality, Santa Cruz de Cana,
upper Rio Tuyra, 606 m; holotype, male, skin and
skull, USNM 179164, collected 23 March 1912 by
E. A. Goldman). Zelek, 1 930:470 PANAMA. En-
ders, 1935:413 PANAMA: Colon/Panama (Barro
Colorado). Dickey, 1928:15 EL SALVADOR: La
Libertad (Hacienda Zapotitan); Handley, 1966:
755 PANAMA: Chiriqui (Rio Colorado); Los
Santos (Guanico; Cerro Hoya); Colon/Panama
(Isla Barro Colorado [= Barro Colorado]); Panama
(Cerro Azul); San Bias (Mandinga; Armila); Da-
rien (Tacarcuna; Boca de Rio Paya; Rio Jaque,
Jaque; Guayabo); Veraguas (Rio Bubi, Rio Viejo).
Baker, 1974:135 ECUADOR: Esmeraldas (Quin-
inde). Perez-Hernandez, 1989:374 VENEZUE-
LA: Merida (La Mucuy); Portuguesa (Colonia
Agricola Turen); Aragua (Cuyagua; Estacion Biol-
ogica de Rancho Grande; Rio Tuy); Guarico (Re-
presa de Guanapito); Miranda (Rio Birongo; La
Toma Capaya; Casupito, Parque Nacional Guato-
po); Sucre (Playa Colorado); Monagas (Caripe).
Gardner, 1993:26 listed.
Chironectes menima bresslaui Pohle, 1927:242 in-
correct subsequent spelling BRAZIL: Rio de Ja-
neiro (type locality, Teresopolis); Sao Paulo (Pir-
acicaba); (holotype, female, skin and skull, ZMB
35441). Gardner, 1993:16 listed.
Chironectes minima bresslaui: Krumbiegel, 1940:
67 BRAZIL: Rio de Janeiro (Teresopolis); Sao
Paulo (Piracicaba).
Chironectes argyrodytes Dickey, 1928:15 EL SAL-
VADOR: La Libertad (type locality, Hacienda Za-
potitan, Rio Sucio; holotype, male, skin and skull,
no. 12986, Donald R. Dickey Collection, MVZ,
collected 20 June 1927 by R. A. Stirton). Goodwin,
1942:112 HONDURAS: Gracias (Las Flores
Gracias); Comayagua (Minas de Oro); Francisco
Morazdn (Tegucigalpa); EL SALVADOR: La Lib-
ertad (Hacienda Zapotitan). Ramirez-Pulido et al.,
1982:13 MEXICO: Tabasco (Golfo de Campe-
che); Chiapas (Pacific coast); Oaxaca (Matfas
Romero). Gardner, 1993:16 listed.
Chironectes minima argyrodytes: Krumbiegel, 1940:
68 classification.
Chironectes minimus argyrodytes: Lay, 1962:374
MEXICO: Tabasco (Rio Teapa).
Type None in existence. Name based on Buf-
fon's petite loutre d'eau douce de Cayenne.
Type Locality Cayenne, French Guiana.
Distribution (Figs. 8 and 9) The originally
forested parts of the Neotropical region from Oa-
xaca and Tabasco, Mexico, south through Central
America and Panama, Colombia, Ecuador, Peru,
Venezuela, and the Guianas, Bolivia, the Orino-
coan and Amazonian regions to Rio Grande do
Sul, Brazil, Paraguay, and northeastern Argentina.
Additional Records MEXICO: Tabasco;
Teapa, 2 mi NW (Armstrong, 1961, LSUMZ; Lay,
1960, LSUMZ); Chiapas; Pichucalco (Smith,
1965, KU); HONDURAS: Lempira; Siguatepeque
(Edwards, 1935, USNM); EL SALVADOR: La
Libertad; Zapotitan (Stirton, 1927, MVZ); GUA-
TEMALA: Izabal; Rio Las Escobas, 3 mi WSW
Puerto Santo Tomas (Campbell, 1981, KU); Finca
Rio Frio (McLean, 1984, USNM); NICARA-
GUA: Nueva Segovia; Jalapa (Phillips, 1967, KU;
Smith, 1967, KU); Boaco; Santa Rosa, 17 km N,
15 km E of Boaco (Genoways, 1967, KU; Jones,
1967, 1968, KU); Teustepe (Kirsch, 1968, KU);
BROWN: ATLAS OF NEW WORLD MARSUPIALS
19
Fig. 8. Distribution of Chironectes miniums in Central America: Oaxaca and Tabasco, Mexico, south through
Panama. See Figure 9 for distribution in South America, o = localities from unpublished records; = localities given
in published records.
Los Cocos (Choate, 1968, KU; Genoways, 1968,
KU; Smith, 1968, KU); Zelaya; El Recreo, S side
Rio Mico (Jones, 1968, KU); Cara de Mono
(Schlitter, 1967, KU); Matagalpa; San Ramon,
not located (Alcorn, 1956, KU); COSTA RICA:
San Jose; San Jose (Zeledon, 1866, USNM);
Guanacaste; Finca Pacffica, 3 mi N of Canas
(Janzen, 1966, UMMZ); Cartago; Agua Caliente
(Torres, 1948, KU); Rio Reventazon, 5 km SE of
Turrialba (Westman, 1947, KU); Linton; Cariari
(Gardner, 1967, LSUMZ); Alajuela; San Carlos
(Smith, 1934, FMNH); PANAMA: Chiriqui; Fin-
ca Santa Clara, El Volcan (Enders, 1971, 1972,
USNM); Escopeta Camp (Pine, 1980, USNM);
Veraguas; Rio Bubi, near Sona (Curtis, 1951,
USNM); Cornes, Rio Viego (Curtis, 1950,
USNM); Isla Cebaco (Tyson, 1965, USNM); Co-
lon; Mohinga Valley, 3 mi S of Fort Sherman
(Setzer, 1953, USNM); Panama; Pedro Miguel
(Keenen, 1953. 1954, USNM); Chepa (Gorgas
Memorial Laboratory, USNM); Colon/Panama;
Nuevo San Juan, not located (Clark, 1951,
USNM); San Bias; Rio Pito (Handley, 1963,
USNM); Quebrada Venado (Handley, 1963,
USNM); Quebrada Repressa. Puerto Obaldia
(Handley, 1963, USNM); Darien; Cana (Gold-
man, 1912, USNM); Rfo Imamado, Jaque (Green-
well, 1966, USNM); Cerro Mali (Handley, 1964,
USNM); Bocas del Tow; Rio Changuinola (Bar-
rett, 1962, USNM; Handley, 1963, USNM);
Changuinola (Handley, 1960, USNM); Sibube
(Handley, 1963, USNM); COLOMBIA: Cordoba;
Rfo Nam, upper Rio Sinu (Hershkovitz, 1949,
FMNH); Socorre, upper Rfo Sinu (Hershkovitz,
1949, FMNH); Choco; Rfo Baudo, Rfo Sando
(von Sneidern, 1958, FMNH); Rfo Docampado
(von Sneidern, 1958, FMNH); Valle del Cauca;
Zabaletas (von Sneidern, 1956, FMNH); Cauca;
La Boca Rfo Saija [= Rfo Saija] (von Sneidern,
1958, FMNH); El Papayo (von Sneidern, 1958,
FMNH); Sabanetas (von Sneidern, 1958, FMNH);
Huila; San Calixto, Valle de Suaza (Hershkovitz,
1943, USNM); Cesar; Colonia de Agrfcola de
Caracolicito (de Santa Marta) (Hershkovitz, 1943,
USNM); La Guajira; Villanueva (Hershkovitz,
1943, USNM); Boyacd; Rio Cobaria (von Snei-
dern, 1959, FMNH); Meta; Villavicencio, near
(Medem, 1953, FMNH); Los Micos, San Juan de
Arama (von Sneidern, 1957, FMNH); ECUA-
DOR: Manabi; San Jose (Gomez, 1942, FMNH);
20
FIELDIANA: ZOOLOGY
Fig. 9. Distribution of Chironectes minimus in South America: Honduras south to Colombia, Ecuador, Peru,
Bolivia, Paraguay, northeastern Argentina, and southern Brazil; Venezuela, the Guianas, and northeastern Brazil. See
Figure 8 for distribution in Central America, o = localities from unpublished records; = localities given in published
records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
21
Pichincha; Mindo (Voss, 1980, MZUM); El Oro;
Puente de Moromoro (Gardner, 1976, USNM);
PERU: Amazonas; Rio Cenepa, N of Huampami
(Patton, 1977, MVZ); Loreto; Centro Union, Rio
Aucayo, not located (Soini, 1972, FMNH); Uca-
yali; Rio Alto Purus (Agnew and Adams, 1962,
LSUMZ); Balta. Rio Curanja (Gardner, 1966,
LSUMZ); Madre de Dios; Shintuya, right bank
Rio Alto Madre de Dios (Willard, 1980, FMNH);
Cusco; Huajyumbe (Kalinowski, 1953, FMNH);
Quincemil (Kalinowski, 1953, FMNH); VENE-
ZUELA: Aragua; Ocumare de la Costa (Handley.
1960, USNM); Amazonas; Puerto Ayacucho
(Smithsonian Venezuelan Project, 1967, USNM);
GUYANA: Barima-Wiani; Warapaco Mission,
Warapoko Creek, Wiani Creek [= Waropoko Mis-
sion, Waropoko Creek, Wiani Creek] (Evans,
1953, USNM); BRAZIL: Acre; Rio Jurua. upper
(Hill, FMNH); Para; Barcarena, 23 km SW of
Belem (Pine, 1968, USNM); Cameta, 25 m, left
bank Rio Tocantins (Olalla, 1933, FMNH); Sao
Paulo; Rocha at ca. 50 m, ca. 40 km NNW of
Juquia (Olalla, 1961, FMNH); PARAGUAY: La
Cordillera; Salto de Pirareta, 10 km S of Piribe-
buy, not located (Myers, 1973, UMMZ); Para-
guari; Piribebuy (Myers, 1976, UMMZ); Parque
Nacional Ybycuf (Creighton, 1979, UMMZ; Dob-
son, 1979, UMMZ; Myers, 1979, MZUM); Itap-
ud; Puerto San Rafael [= San Rafael], 2 km
NNW (Myers, 1978, UMMZ); Canendiyu; Curu-
guaty, 6.3 km N (Myers, 1976, UMMZ).
Genus Didelphis Linnaeus, 1758
Didelphis Linnaeus, 1758:54 included species, mar-
supialis, philander [Caluromys philander], opos-
sum [= Philander], murina [ = Marmosa marmo-
sa], dorsigera [= Marmosa murina]. Gardner.
1993:16 listed.
Didelphys Schreber, 1778:532 classification: name
and emendation of Didelphis Linnaeus.
Opossum Schmid. 1818:115 included species: mar-
supialis Linnaeus [here designated type species],
virginiana [not Kerr; = Philander opossum], mu-
rina [= Marmosa murina].
Sarigua Muirhead, 1819:429 part, included species
Didelphis marsupialis and cancrivora, Linnaeus
and S. [sarigua] opossum (part).
Didelphus I. Geoffroy, 1831:17 (atlas) incorrect
subsequent spelling of Didelphis Linnaeus, 1758.
Gamba Liais, 1872:329 type species, Gamba aurita
var. brasiliensis Liais [= Didelphis aurita Wied-
Neuwied] by monotypy.
Dimerodon Ameghino, 1 889(6):479 type species D.
mutilatus Ameghino, by monotypy.
Leucodidelphis Ihering, 1914:356 type species, D.
paraguayensis Oken 1816 [= Didelphis albiventris
Lund] by original designation.
Leucodidelphys: Krumbiegel, 1941a:34 incorrect
subsequent spelling of Leucodidelphis Ihering,
1914 subgenus of Didelphys; type species, Leu-
codidelphys paraguayensis Oken [= Didelphis al-
biventris Lund], by original designation.
Leucodidelphis: Krumbiegel, 1941a:34.
Leucodelphis Cabrera, 1958:41 incorrect subsequent
spelling of Leucodidelphis Ihering. 1914 in syn-
onymy of Didelphis Linnaeus.
Type Species Didelphis marsupialis Linnae-
us, by subsequent designation (Thomas, 1911).
Distribution (Figs. 10 and 11) From south-
ern Canada south through Central America, and
South America into northern half of Argentina.
Didelphis albiventris Lund, 1841
Carigueya brasiliensis: Marcgrave, 1648:222.
Tai-ibi brasiliensibus: Marcgrave. 1648:223.
Micoure premier ou Micoure proprement dit: Mo-
reau-Saint-Mery in Azara, 1801:244.
Didelphis Virginiana, Desmarest (part, not Kerr).
1817:420 Azara's micoure premier in synonymy.
Didelphis paraguayensis Froriep, 1822:273 nomen
nudum in notice of specimens received from Brazil
by the Vienna Museum; not an available name.
Didelphis paraguayensis J. A. Allen, 1902:267
PARAGUAY: Central (type locality, Asuncion);
URUGUAY; BRAZIL: Minas Gerais (Lagoa San-
ta: Rio Jordao): Rio Grande do Sul; ARGENTINA:
Tucumdn; Corrientes (Goya); new name estab-
lished by use of previously unavailable name (In-
ternational Code of Zoological Nomenclature,
1999:99). Thomas. 1903:243 BRAZIL: Mato
Grosso (Santa Anna de Chapada [= Chapada dos
Guimaraes]). Thomas. 1 9 10a:247 ARGENTINA:
Buenos Aires (Los Inglesas). Ihering, 1914:344
BRAZIL: Sao Paulo (Ituverava; Rincao; Franca);
Bahia (Villa Nova): Ceard. L. E. Miller, 1916:
589 BRAZIL. Cabrera, 1917:10 ECUADOR:
Pichincha (Quito). Thomas, 1926b:327 BOLIV-
IA: Tarija (Carlazo). Thomas, 1926c:608 AR-
GENTINA: Tucumdn (Cerro del Campo; Burruy-
acu, 800 m). Sanborn, 1929:150 URUGUAY. De-
vincenzi, 1935:88 URUGUAY Bertoni, 1939:6
PARAGUAY Gilmore, 1941:316 distribution.
Krumbiegel. 1941a:32, 37 distribution. Vieira,
1949:343 BRAZIL: Sao Paulo; Rio Grande do
Sul; Mato Grosso; Maranhao; Bahia; Goids. Vi-
eira, 1951:124 BRAZIL: Mato Grosso (Rio das
Mortes; Sao Domingos). Kuhlhorn, 1953a:5
BRAZIL: Mato Grosso (Alto Parana [now in Mato
Grosso do Sul]: (Rio Ivinheima now in Mato Gros-
so do Sul). Kuhlhorn, 1953b: 120 BRAZIL: Mato
Grosso do Sul (Rio Ivinheima). Kuhlhorn, 1955:
162 BRAZIL: Mato Grosso. Ruschi, 1965:1
PARAGUAY: Central (Asuncion). Steyer and Pau-
lette Vanrell, 1970:434 BRAZIL: Rio Grande do
Sul (Rio Taquari, valley of).
D[idelphis] paraguayensis: Miranda Ribeiro, 1936:
22
FIELDIANA: ZOOLOGY
Fig. 10. Distribution of Didelphis in northern Central America: northern Mexico south through Central America
to Colombia. See Figure 1 1 for distribution in South America (North American distribution not shown).
399 BRAZIL: Mato Grosso do Sul (Coxim);
Mato Grosso (Caceras); Golds (Tocantins, not lo-
cated).
Didelphis paraguayensis paraguayensis: J. A. Allen,
1 9 16b:562 PARAGUAY: Presidente Hayes (Rio
Negro); Central (Trinidad); BRAZIL: Mato Grosso
(Utiarity). Yepes, 1936:696 ARGENTINA: Men-
doza (La Paz; San Rafael; Rivadavia; San Carlos);
Cordoba (La Paz); San Luis (Rincon); dennleri
Marelli, in synonymy. Moojen, 1943:2 BRAZIL:
Pernambuco (Pocao).
Didelphys paraguayensis: Krieg, 1924:637 AR-
GENTINA: Sante Fe. Krumbiegel, 1941a:37
BRAZIL: Mato Grosso do Sul (Rio Ivinheima); Rio
Grande do Sul (Passo Fundo); PARAGUAY: Cha-
co (San Luis de la Sierra, not loaded; Centurion
[now in Concepcion]; Mennoniten Kolonien [=
Colonia Mennonita, now in Boqueron]; Nueva Ger-
mania [now in San Pedro]); BOLIVIA: Santa Cruz
(San Jose, Rio San Miguel); ARGENTINA: Mi-
siones; Formosa (La Urbana, Rio Pilcomayo); San-
ta Fe (Estancia La Germania); Santa Fei 'Santiago
del Estero (Estancia la Geraldina).
Did[elphys]. paraguayensis paraguayensis: Krumbie-
gel, 1941b:41 distribution.
[Leucodidelphys] paraguayensis paraguayensis:
Krumbiegel, 1941a:34 classification.
Didelphis Azarae Rengger (not Temminck, 1824),
1830:223 PARAGUAY Waterhouse, 1839:93
URUGUAY: Maldonado (Maldonado). Burmeister,
1856:61 PARAGUAY; URUGUAY: Montevideo
(Montevideo); BRAZIL: Sao Paulo southward.
Thomas, 1902b:245 ARGENTINA: Cordoba
(Cruz del Eje).
Didelphis azarae: Jentink (not Temminck, 1824),
1887:300 ECUADOR: Pichincha (Quito). Jen-
tink, 1888:219 part, BRAZIL; ECUADOR: Pi-
chincha (Quito); ARGENTINA: Buenos Aires
(Buenos Aires). Thomas, 1901c:536 BRAZIL:
Minas Gerais (Rio Jordao). Yepes, 1944:60 AR-
GENTINA: Formosa (Las Lomitas); Santa Fe
(Malabrigo; Villa Ana); Buenos Aires (25 de
Mayo); Salta (Zuviria; Lumbrera; Cayafate [= Ca-
fayate]; El Quebrachal; Dragones; Tabacal; Agu-
aray); Jujuy (Arenal; Rib Blanco); Mendoza (Ri-
vadavia; San Carlos; San Rafael). Reig, 1964:
207 ARGENTINA: Buenos Aires (Arroyo Brus-
quitas; Arroyo Chapadamalal; Baliza Caviu; Mar
del Plata [= Ciudade]; Mar del Sur; Miramar).
Contreras, 1973:215 ARGENTINA: Buenos Ai-
res (Laguna Chasico). Mendez, 1977:111 CO-
LOMBIA: Valle del Cauca (Finca Hollanda, near
Paramo de Chinche [Hershkovitz, pers. comm., as-
signs this specimen to albiventris]).
D[idelphis] Azarae: Cabrera (not Temminck, 1824),
1913:145 ECUADOR.
Didelphus azarae, I. Geoffroy (not Temminck, 1 824),
1831:17 (atlas) incorrect subsequent spelling of
Didelphis Linnaeus, 1758 BRAZIL.
Didelphys Azarae: Waterhouse (not Temminck,
1824), 1841:83 incorrect subsequent spelling of
Didelphis Linnaeus, 1758 BRAZIL; URUGUAY:
Maldonado (Maldonado [Darwin collector]). Bur-
meister, 1861:412 ARGENTINA: Tucumdn;
BROWN: ATLAS OF NEW WORLD MARSUPIALS
23
Fig. 1 1 . Distribution of Didelphis in southern Central and South America: throughout South America, excluding
Chile and southern Argentina. See Figure 10 for distribution in northern Central America (North American distribution
not shown).
24
FIELDIANA: ZOOLOGY
URUGUAY: [= Banda Oriental (Oriente)]. Hensel,
1873:111 BRAZIL: Rio Grande do Sul; azarae
authors, not Temminck [= D. marsupialis]. Bur-
meister, 1 879: 1 89 ARGENTINA. Ihering, 1892:
98 BRAZIL: Rio Grande do Sul.
D[idelph\s\ azarae: Wagner (not Temminck, 1824),
1843b": 3 8 part. PARAGUAY: Gran Chaco; AR-
GENTINA: Entre Rios; URUGUAY: (Banda Ori-
ental [Oriente], not located). Tschudi, 1844a: 143
PERU: part. Tschudi, 1844b: 250 PERU: part.
Didelphys azarae: Waterhouse (not Temminck, 1 824),
1846:470 BRAZIL; PARAGUAY; BOLIVIA;
URUGUAY: Maldonado (Maldonado; "El Cor-
te"); COLOMBIA: Cundinamarca (Santa Fe [now
Santafe] de Bogota). Tomes, 1860:268 ECUA-
DOR: Azuay (Cuenca). Hensel, 1867:290 BRA-
ZIL: Rio Grande do Sul. Aplin, 1894:314 URU-
GUAY. Thomas, 1900a:549 BRAZIL: Parana
(Palmeira).
Didelphys marsupialis var. azarae: Thomas (not Tem-
minck, 1824), 1888b:328 azarae Burmeister, not
Temminck [= marsupialis]; COLOMBIA: Cundi-
namarca (Santa Fe [now Santafe] de Bogota); BO-
LIVIA; BRAZIL: Rio Grande do Sul (Taquara);
CHILE (purchased specimen, probably not origi-
nating in Chile). Lyne, 1959:84 ECUADOR: Pi-
chincha (Cayambe).
Didelphys marsupialis azarae: Cope (not Temminck,
1824), 1889:129 BRAZIL: Rio Grande do Sul
(Sao Joao do Monte Negro).
Didelphys marsupialis var. D. azarae: Figueira (not
Temminck, 1824), 1894:8 (separate) URUGUAY.
Didelphis azarae azarae: Cabrera (not Temminck,
1824), 1958:41 classification; synonyms, albiven-
tris, poecilotis, poecilonota, leucotis, lechii, para-
guayensis, bonariensis, dennleri. Fornes and Mas-
soia, 1965:101 ARGENTINA: Buenos Aires
(Miramar).
Didelphys azarai: Ringuelet, 1954:295 incorrect
subsequent spelling of Didelphis azarae Rengger,
1830 ARGENTINA: Cordoba (Barrancas del Rio
Primero [Post Pampean, lower Quaternary]).
Didelphis albiventris Lund, 1841:106. Lund, 1842:
133, 199 BRAZIL: Minus Gerais (Lagoa Santa).
Ihering, 1914:345 BRAZIL: Sao Paulo (Ituvera-
va; Rincao; Franca); Bahia (Villa Nova); Ceard;
synonyms, tai-ibi brasiliensibus Marcgrave, 1648;
poecilotis Wagner, 1842; paraguayensis Allen,
1902. Ruschi, 1965:191 PARAGUAY: Central
(Asuncion). Ximenez and Langguth, 1971:48
URUGUAY: Rio Negro (Rio Negro). Wetzel and
Lovett, 1974:205 PARAGUAY: Boqueron (Es-
tancia Juan de Zalazar); Chaco (Fortin Juan de Zal-
azar [now in Presidente Hayes). Mendez, 1977:
111 COLOMBIA: Valle del Cauca (Finca Hol-
landa). Handley, 1976:5 VENEZUELA: Tdchira
(Buena Vista); Merida (Paramito; Tabay). Reig et
al., 1977:211 PERU: Hudnuco (Cordillera Car-
pish); VENEZUELA: Merida (Laguna Negra; Tim-
otes); ARGENTINA: Buenos Aires (Punta Lara).
Olrog, 1979:10 ARGENTINA: Jujuv (Cerro Cal-
ilegua). Genoways et al., 1981:319 SURINAME:
Brokopondo (Brownsberg Nature Park). Tyndale-
Biscoe, 1980:712 COLOMBIA: Valle del Cauca
(Pichinde; Pilimbala, not located). Mares et al.,
1981:167 ARGENTINA: Salta (El Breal [= Ri-
vadavia]). Cajal, 1981:27 ARGENTINA: Tucu-
mdn (Aguas Chiquitas). Varejao and Valle, 1982:
37 BRAZIL: Minas Gerais (Rio Acima; Aragu-
ari; Bambui; Belo Horizonte; Bonfinopolis de Mi-
nas; Cachoeira do Campo; Caete (= Rocas Novas);
Carmiopolis de Minas; Douradoquara; Fazenda
Cascata; Florestal; Itabira; Jose de Melo; Lagoa
Santa; Lassance; Luz; Machado; Mateus Leme;
Mato Verde; Matozinhos; Monte Bello; Novo
Lima; Para de Minas; Paraopeba; Passos; Patos de
Minas; Pedro Leopoldo; Pirapora; Pompeu; Prados;
Presidente Juscelino; Presidente de Morais; Sahara;
Salinas, Santa Luzia; Sao Gon^alo Rio Abaixo
[Ciudade]; Sao Roque de Minas; Tupaciguara; Ser-
ra do Caraca [= Santa Barbara]); Goids (Anapolis);
Distrito Federal (Brasilia). Yonenaga-Yasuda,
1982:72 BRAZIL: Sao Paulo (Americana; Nova
Odessa; Santa Maria da Serra). Crespo, 1982a:59
ARGENTINA: Misiones (Parque Nacional Iguazii).
Seluja et al., 1984:702 URUGUAY: Canelones
(Banado de Carrasco; Las Piedras); Rocha (La
Tuna); Tacuarembo (Rio Negro); Maldonado (Pir-
apolis). Fonseca and Redford, 1984:518 BRA-
ZIL: Distrito Federal (Reserva Ecologica Biologi-
ca do Instituto de Geografia e Estatfstica [IBGE]).
Cerqueira, 1984:96 BRAZIL: Pernambuco (Bom
Conselho; Triunfo; Exu; Garanhuns; Pedra; Pes-
queira); Paraiba (Princesa Isabel); Ceard (Crato);
Bahia (Feira de Santana; Jabaquara; Jequie; Ju-
azeiro; Palmeiras; Seabra; Serrinha); Alagoas (Pal-
meira dos Indios; Quebrangulo; Vicosa). Cerqueira,
1985:135 BRAZIL: Bahia (Castro Alves); Ala-
goas (Vicosa). Alho et al., 1986:451 BRAZIL:
Distrito Federal (Parque Nacional de Brasilia). Ma-
res et al., 1986:294 BRAZIL: Distrito Federal
(Brasilia). Massoia et al., 1987:116 ARGENTI-
NA: Misiones (Rio Urugua-i). Nogueira, 1988:
233 BRAZIL: Minas Gerais (Belo Horizonte).
Gardner, 1988:690 VENEZUELA: Amazonas
(Camp II, Cerro de La Neblina; Camp VII, Cerro
de La Neblina). Gardner, 1 990:4 1 2 VENEZUE-
LA: Bolivar (Auyan-tepuf); Amazonas (Camp II,
Neblina; Camp VII, Neblina). Gardner, 1993:16
listed. Heinonen and Bosso, 1994:55 ARGEN-
TINA: Jujuy (Parque Nacional Calilegua). Rageot
and Albuja, 1994:172 ECUADOR: Pastaza
(Mera). Mares and Ernest, 1995:775 BRAZIL:
Distrito Federal (Reserva Ecologica Biologica do
Instituto de Geografia e Estatfstica [= IBGE)). Ma-
res et al. 1996:101 ARGENTINA: Tucumdn
(Aconquija; Concepcion; Cerro del Campo; El
Cadillal Dike, 25 km NW of San Miguel de Tu-
cuman; El Cajon; 4 km W of junction 338 and road
to Horco Molle, on road to San Javier; Biological
Reserve at Horco Molle; Los Romanos; Mala
Mala; Escuela Normal, Monteros; Piedra Tendida;
San Fernando; San Pedro de Colalao; km 42, on
highway 364, south of San Pedro de Colalao;
Aguas Chiquitas; Pozo Hondo; Dique Escaba; San-
ta Rosa de Leales; Tafi del Valle). Capllonch et al.,
1997:53 ARGENTINA: Tucumdn (Parque Biol-
ogico Sierra de San Javier). Mares et al., 1997:
100 ARGENTINA: Catamarca (Choya; Andal-
gala). Morando and Polop, 1997:130 ARGEN-
TINA: Cordoba. Catzeflis et al., 1997:231
FRENCH GUIANA (Sinnamary River; Crique
BROWN: ATLAS OF NEW WORLD MARSUPIALS
25
Tigre). Mares and Braun, 2000:30 ARGENTINA:
Catamarca (Andalgala; Choya; La Merced); Men-
doza (Tunuyan, 27 km S, 96 km N); Misiones (Ar-
istobulo de Valle, 10 km W on Rio Cunapiru); Sal-
ta (Aguaray; Cafayate; Dragones; El Quebrachal;
Lumbrera; Parque Nacional Baritu; Parque Nacion-
al El Rey; Santa Victoria, El Breal, 6 km SW of;
Tabacal); San Luis (San Francisco del Monte de
Oro); Santiago del Estero (Santo Domingo); Tu-
cumdn (Aconquija, Concepcion; Cerro del Campo;
El Cadillal Dike; El Cajon; Biological Reserve at
Horco Molle; junction 338 road to Horco Molle, 4
km W; Los Romanos; Mala Mala; Escuela Normal,
Monteros; Piedra Tendida; San Fernando; km 42,
highway 64, S of San Pedro de Colalao; San Pedro
de Colalao); Buenos Aires (Arroyo Brusquitas; Ar-
royo Brusquitas-Miramar; Arroyo Chapadmalal;
Azul, 4 km NE; Azul Benson Farm, not located;
Baliza Caniii; Laguna de Los Padres; La Plata,
Punta Lara; Los Yngleses; Mar del Plata, S of;
Ciudad de Mar del Plata; Mar del Sur; Miramar;
Punta Lara, Ribera; Reserva Natural Otamendi; 25
de Mayo); Chaco (Parque Nacional Chaco; Puerto
Bermejo, 10 km S); Cordoba (Cruz de Eje; La
Paz); Corrientes (Centro Argentino de Primates, 20
km SE, not located; Parque Nacional Mburucuya);
Entre Rios (Parque Nacional Diamante; Parque Na-
cional El Palmar); Formosa (El Colorado; Laguna
Blanca; Las Lomitas; Parque Nacional Rio Pilco-
mayo; Reserva Natural Formosa); Jujuy (Arenal;
Calilegua; Parque Nacional Calilegua; Rib Blanco);
La Rioja (Villa Castelli); Mendoza (Mendoza; Na-
cufian; Reserva Ecologica de Nacunan; Rivadavia;
San Carlos; San Rafael); Misiones (Arroyo Vina
[brazo del Arroyo Garupa]; El Dorado; Parque Na-
cional Iguazii; Puerto Schwelm, not located); San
Luis (Rincon); Santa Fe (Malabrigo; Villa Ana);
Tucumdn (Aguas Chiquitas; Cerro del Campo, Bur-
rucayii; Cerro Medici; Cerro San Javier; Dique Es-
caba; Parque Biologico Sierra de San Javier, access
road; Pozo Hondo; Reserva Experimental de Flora
y Fauna de Horco Molle; Rio Las Cafias, Horco
Molle; Santa Rosa de Leales; Taff del Valle; junc-
tion at 338 and road to Horco Molle). Anderson,
1997:143 BOLIVIA: Chuquisaca (Tarabuco, 12
km N, 11 km E; Porvenir); Potosi (Rio Chancha-
mayo); Santa Cruz (San Ignacio de Velasco; Com-
arapa, 6 km S, 10 km E; Comarapa, 15 km W);
Tarija (Tapecua; Rancho Tambo, 61 km E; Carla-
zo). Voss et al., 2001:23 FRENCH GUIANA:
Piste St. Elie [= Piste de Saint-Elie Ecerex Sta-
tion]; Petit Saut).
D[idelphvs] albiventris: Wagner, 1843:38. Burmeister,
1856:62 BRAZIL.
Didelphys albiventris: Waterhouse, 1846:472 BRA-
ZIL. Burmeister, 1854:132 BRAZIL: Minas Ger-
ais (Lagoa Santa). Ihering, 1894:10 BRAZIL:
Sao Paulo.
Didelphys marsupial is var. albiventris: Winge, 1893:
3, 7, 46 BRAZIL: Minas Gerais (Lagoa Santa,
Recent, and Pleistocene).
Didelphys poecilotis Wagner, 1842:358 BRAZIL:
Mato Grosso (type locality, "Angaba" [= Cuiaba];
holotype. female, Vienna Museum, collected 19
July 1824 by Johann Natterer). Gardner, 1993: 16
listed.
D[idelphys] poecilotis: Wagner, 1855:219. Pelzeln,
1883: i09 BRAZIL: Mato Grosso (Cuyaba [ =
Cuiaba]).
Didelphys paeci lotus: Wagner, 1843:43 incorrect
subsequent spelling of poecilotis Wagner, 1942.
Waterhouse, 1846:524 BRAZIL.
D[idelphys] poecilonota: Schinz, 1844:504 incor-
rect subsequent spelling of poecilotus Wagner.
Didelphys leucotis Wagner, 1850:127 name based
on Azara's micoure premier and azarae of authors,
not Temminck; type locality, by implication, Para-
guay and Uruguay. Gardner, 1993:16 listed.
D[idelphys] leucotis: Wagner, 1855:223 PARA-
GUAY; URUGUAY: (Rio de La Plata, not located).
Didelphis aurita: Alston (not Wied-Neuwied, 1826),
1882:197 part, COLOMBIA: Cundinamarca
(Santa Fe [now Santafe] de Bogota).
Didelphis marsupialis aurita: Yepes, 1936:34 AR-
GENTINA: La Rioja (Villa Castelli).
Didelphys Lechei Ihering, 1892:98 BRAZIL: Rio
Grande do Sul (type locality; holotype, a gray
phase individual or variety of "Didelphys Aza-
rae"). Gardner, 1993:16 listed.
Didelphys lechii: Cabrera, 1958:41 incorrect subse-
quent spelling in synonymy of "Didelphis azarae
azarae."
Didelphis lechi: Avila Pires, 1994:371 incorrect
subsequent spelling of Lechei Ihering, 1892
BRAZIL: Rio Grande do Sul (Taquara).
Didelphis albiventris pernigra J. A. Allen, 1900b:
191 PERU: Puno (type locality, "Juliaca" [ =
Inca Mines, Rio Inambari], 7000 ft [= 2180 m];
holotype, female, skin and skull, AMNH 16071;
collected 12 February 1900 by H. H. Keays). J. A.
Allen, 1900c:219 "Juliaca" corrected to Inca
Mines. J. A. Allen, 1901a:41 PERU: Puno (Inca
Mines). Thomas, 1901b: 191 PERU: Cusco (Mar-
capata); Puno (Limbani). Truessart, 1910:A26
ECUADOR: Pichincha (Gualea). Perez-Hernan-
dez, 1989:372, 384, 385 VENEZUELA: Amazon-
as (Cerro Duida, 1400 m) Merida (La Mucuy; La-
guna Negra, Paramo de Mucubaji; Las Porqueras);
Tdchira (Betania; Cerro Las Copos, La Revancha;
Paramo Zumbader). Gardner, 1993:16 listed.
Anderson, 1997: 144 BOLIVIA: Cochabamba
(Charuplaya; Yungas; Colomi, 5 km N; Cochabam-
ba; Vinto; Tapacari; Jamachuma; Rodeo, 7.5 km
SE); La Paz (Ixiamas; Saynani; Nequejahuira; Rio
Aceromarca; Pitiguaya; Zongo Valley, "Centro
Hidroelectrica").
Didelphis paraguayensis pernigra: J. A. Allen, 1902:
271, 279 PERU: Lima (Callao [although cited,
this specimen, given the locality, must be marsu-
pialis]; Surco; San Mateo); Junin (Huacapistona;
Perene); Puno (Inca Mines; Limbani); Cajamarca
(Carao); BOLIVIA: La Paz. Trouessart, 1910:
A29 ECUADOR: Pichincha (Tumbaco, 2390 m).
Thomas, 1920c: 247 PERU: Cusco (Chospyoc,
10,000 ft [= 3125 m]; Torontoy; Machu Picchu [=
Macchu Picchu]). Thomas, 1926c:318 PERU: Ju-
nin (Yana Mayo, Rio Tarma, 8500 ft [= 2650 m]).
Thomas, 1927a:372 PERU: Cajamarca (Celen-
din). Gardner, 1993:16 listed.
Did[elphis] paraguayensis pernigra: Krumbiegel,
1941a:41, 42 distribution.
Didelphis paraguayensis andina J. A. Allen, 1902:
26
FIELDIANA: ZOOLOGY
272, 279 ECUADOR: Azuay (type locality,
Cuenca; holotype, male, BMNH 99.12.7.6; collect-
ed 2 June 1899 by P. O. Simons); Pichincha (Quito;
Cayambe); Canar (Canar); hnbabura (Ibarra); Loja
(Loja); Chimborazo (Riobamba). Trouessart, 1910:
A28 ECUADOR: Pichincha (Gualea). J. A. Al-
len, 191 2a:74 COLOMBIA: Cauca (Munchique,
2100-2400 m). Lonnberg, 1913:35 ECUADOR:
Pichincha (Quito; Nanegal). Osgood, 1914b: 148
PERU: Cajamarca (Hacienda Limon, near Balsas);
Amazonas (Rio Utcubamba, 15 mi above Chacha-
poyas). J. A. Allen, 1916a:200 COLOMBIA: Cau-
ca (Munchique); Antioquia (Paramillo). J. A. Allen,
1916d:l 16 ECUADOR: Pichincha (Quito; El
Corazon). Gardner, 1993:16 listed.
Did[elphis] paraguayensis andina: Krumbiegel,
1941a:41, 42 distribution.
Didelphis marsupialis: Stone (not Linnaeus, 1758),
1914:17 ECUADOR: Pichincha (Volcan Cay-
ambe [= Cerro Cayambe]).
[Didelphis] leucoprymnus Matschie, 1916:268 no-
men nudum; BRAZIL: Rio de Janeiro (Cabo Frio).
Didelphis paraguayensis meridensis J. A. Allen,
1902:274 VENEZUELA: Merida (type locality,
Merida; holotype, juvenile male, skin and skull,
BMNH 97.7.1.14); COLOMBIA: Cundinamarca
(Bogota). Osgood, 1912:39 VENEZUELA: Me-
rida (Paramo de Tama, 6000-7000 ft [= 1875-
2180 m] [now in Tdchira]). J. A. Allen, 1916a:
200 COLOMBIA: Cundinamarca (Fusagasuga;
El Roble). Anthony, 1923:4 COLOMBIA: Cun-
dinamarca (Choachi; Bogota). Gardner, 1993:16
listed.
Did[elphis] paraguayensis meridensis: Krumbiegel,
1941a:42 distribution.
Didelphis paraguayensis bonariensis Marelli, 1930:2
[work not seen]." Marelli, 1932:69 ARGENTINA:
(type locality, Provinces of Buenos Aires and Santa
Fe; [holotype probably a menagerie specimen the
remnants of which may be preserved in the Buenos
Aires Museum]). Gardner, 1993:16 listed.
Didelphis paraguayensis Dennleri Marelli, 1930:2
[work not seen]. Marelli, 1932:69 ARGENTINA:
(type locality, Buenos Aires Province [holotype
probably a menagerie specimen, the remains of
which may be preserved in the Buenos Aires Mu-
seum]).
Didelphis albiventris imperfectus Mondolfi and Perez-
Hernandez, 1984:407 VENEZUELA: Bolivar
(type locality, km 125, El Dorado-Santa Elena
Highway; holotype, adult female, skin and skull,
MHNLS 175). Perez-Hernandez, 1989:372-382
VENEZUELA: Bolivar (km 124, 125, 126, El Do-
rado-Santa Elena Highway; km 70-88, [Carretera]
El Dorado-Santa Elena Highway; La Tigre; Torono
Tepui [= Torono-tepui], Chimanta; Salto Ichum;
Rio Marajano, Meseta de Jaua; La Laja; El Pauji;
Monumento al Soldado Pionero, Gran Sabana);
Amazonas (Cano Culebra, Cerro Duida; Cumbre
del Cerro, Marahuaca, 2550 m; Alto Rio Atacavi;
Cerro de La Neblina). Gardner, 1993:16 listed.
Didelphis sp.: Handley, 1976:9 VENEZUELA: Bo-
livar (Icabani; km 125, 85 km SSE of El Dorado);
Amazonas (Boca Mavaca; Cabecera del Cano Cu-
lebra, Cerro Duida; Cano Culebra, Cerro Duida;
Puerto Ayacucho, 20-30 km S).
Holotype Presumably in the Zoological Mu-
seum of Copenhagen; collected by Peter Wilhelm
Lund.
Type Locality Rio das Velhas, Lagoa Santa,
Minas Gerais, Brazil.
Distribution (Fig. 12) Subtropical and tem-
perate zones of Colombia, Ecuador, Peru, Vene-
zuela, and Suriname; the highlands of Guyana; the
northern half of Brazil and Brazil east of the Rio
Tocantins; Bolivia; Uruguay; Paraguay; and the
northern half of Argentina.
Additional Records COLOMBIA: Valle del
Cauca; San Antonio, 4 km NW (Miller, 1958,
MVZ); Cauca; Charguayaco (von Sneidern, 1958,
AMNH); Antioquia; Valdivia (Hershkovitz, 1950,
FMNH); Sonson (Hershkovitz, 1950, FMNH);
Huila; San Antonio (Hershkovitz, 1951, FMNH);
Rio Ovejera (Hershkovitz, 1951, FMNH); Aguas
Claras, Rio Sauza (Hershkovitz, 1951, FMNH);
Cundinamarca; Guasca, Rio Balcones (Hershko-
vitz, 1952, FMNH); Boqueron de San Francisco
(Hershkovitz, 1952, FMNH; Niceforo Maria,
1922, AMNH); San Cristobal, suburb of Bogota
(Hershkovitz, 1952, FMNH); Meta; Villavicencio
(Dillon, 1942, AMNH; Gilmore, 1940, AMNH);
La Guajira; Villanueva (Hershkovitz, 1942, 1943,
USNM); Norte de Santander; Pamplona (von
Sneidern, 1959, FMNH); ECUADOR: El Oro;
Santa Rosa (Tate, 1922, AMNH); El Chiral (An-
thony, 1920, AMNH); Taraguacocha, Cordillera
de Chilla (Anthony, 1920, AMNH); Canar; Chi-
cal (Tate, 1922, AMNH); San Antonio, 1 hr N of
Tambo (Tate, 1922, AMNH); Carchi; Montufar,
San Gabriel (Hershkovitz, 1934, UMMZ); Pichin-
cha; Mindo (Olalla, 1960, FMNH); Guaillabam-
ba, Riobamba (Simons, 1899, AMNH); Canon del
Rio Pita (Tate, 1923, AMNH); Tambillo, 3 km S
(Timm, 1983, FMNH); Las Maquinas, Santo Do-
mingo Trail (Tate, 1923, AMNH); Santa Rosa, not
located (Tate, 1922, AMNH); Tunguarahua; Ba-
nos, Oriente (Clark-Macintyre, 1938, MVZ, 1938,
FMNH); Palmera, E of Banos (Tate, 1924,
AMNH); Chimborazo; San Antonio (Tate, 1922,
AMNH); Azuay; Molleturo (Tate, 1922, AMNH);
Bestion (Anthony, 1921, AMNH); Zamora-Chin-
chipe; Sabinilla, Rio Zamora, near mouth Rio Sa-
banilla (Anthony, 1920, AMNH); Napo; Papallac-
ta (Voss, 1980, UMMZ); Marian (Timm, 1983,
FMNH); PERU: Piura; Huancabamba, Can-
chaque (Kalinowski, 1954, FMNH); Cajamarca;
Chaupe (Watkins, 1923, AMNH); Hacienda Tau-
lis (Koepcke, 1952, MVZ); Cajamarca (Simons,
1899, USNM); La Libertad; Pagash, not located
(1980, MVZ); Lima; Lima (Ramirez, 1955, MVZ);
BROWN: ATLAS OF NEW WORLD MARSUPIALS
27
Fig. 12. Distribution of Didelphis albiventris in South America: absent only from Amazon basin of Brazil, Chile,
and southern Argentina. This species in not known to occur in Central America, o = localities from unpublished
records; = localities given in published records.
28
FIELDIANA: ZOOLOGY
Amazonas; Corosha (Martinez, 1970, MVZ); Lor-
eto; Yanayaquilla, Rib Samiria (Bluntschli, 1912,
AMNH); Rio Samiria (Bluntschli, 1912, AMNH);
Hudnuco; Hacienda Buena Vista, Rio Chinchao
(Heller, 1922, FMNH); Carretera Central, E slope
Cordillera Carpish (Gardner, 1968, LSUMZ);
Huanuco (Zimmer, 1922, FMNH); Ambo (Heller,
1922, FMNH); Pasco; San Juan (Perez, 1964,
FMNH); Junin; Chanchamayo (Schunke, 1924,
USNM; 1948, FMNH); La Merced (Schunke,
1924, USNM); Utcuyacu (Watkins, 1919,
AMNH); Matichaca, near Puente Malalma
(O'Neill, 1972, LSUMZ); Tarma, 17 km below
(Gochfeld, 1972, AMNH); Ayacucho; Ayacucho
(Church, 1864, USNM; Thomas, USNM); Cusco;
Occobamba Valley [= Rio Ocobamba] (Heller,
1915, USNM); Ocobamba, below La Raya Pass
(1974, MVZ); Ocobamba (Pass) (Heller, 1915,
USNM); Ollantaytambo (Heller, 1915, USNM);
Limacpunco (Kalinowski, 1953, FMNH); Cusco
(Ceballos, 1954, FMNH); Apurimac; Uripa (Kal-
inowski, 1953, FMNH); Hacienda La Palmira
(Kalinowski, 1953, FMNH); Hacienda La Lagu-
na, near Andahuaylas (Kalinowski, 1953,
FMNH); Quebrada Matara (Sanborn, 1941,
FMNH); Puno; Oconeque (Koford, 1951, MVZ;
Sanborn, 1942, FMNH); VENEZUELA: Trujillo;
Trujillo, 15 km E (Kirsch, 1969, KU); Merida;
Hacienda La Carbonera, Paramo Tambor (Osgood
and Conover, 1920, FMNH); Rio Mucujun (Os-
good and Conover, 1920); Sucre; Cumanacoa
(Tate, 1925, AMNH); Bolivar; San Ignacio de Yu-
ran [= Yuruan], Rio Yuruan (Castellanos, 1986,
USNM); Chimanta-tepui (Steyermark and Wur-
dack, 1955, FMNH); Cerro Roraima (Tate and
Carter, 1927, 1928, AMNH); Arabopo (Tate and
Carter, 1928, AMNH); Towashing [= to wash-
ing?] Stream (Tate and Carter, 1927, AMNH);
Glycon Ridge (Tate and Carter, 1927, AMNH);
Amazonas; Central Camp, Cumbre no. 1 ; Cumbre
no. 13, Despiladero, Cerro Duida (Olalla Broth-
ers, 1929, AMNH); Cumbre no. 15, Cerro Duida
(Olalla Brothers, 1929, AMNH); Agiiita (Olalla
Brothers, 1929, AMNH); Laterite Valley (Olalla
Brothers, 1929, AMNH); BRAZIL: Mato Grosso
do Sul; Corumba (Hibbs, 1965, USNM; Kuns,
1965, USNM); Campo Grande, 13 km N (Hibbs,
1965, USNM; Kuns, 1965, USNM); Maracaju
(Gilmore, 1937, AMNH); Dourados, 25 km S
(Hibbs, 1965, USNM; Kuns, 1965, USNM); Mar-
anhdo; Sao Joao dos Patos (Cardosa, AMNH);
Piaui; localities along lower Rio Parnahyba, not
precisely located (Kaempfer, 1926, AMNH); Ba-
hia; Macaco Secco, near Serra Andarai [= An-
darahy] (Becker, 1913, FMNH); Sao Paulo; Cru-
zeiro (Robert, 1900, AMNH); Piquete (Robert,
1901, AMNH); Alto Parana, not located (Kuhl-
horn, 1953, FMNH); Rio Grande do Sul; Paso
Duraznero, Ayo da Mina [= Arroyo de la Mina]
(Ximenez, 1970, AMNH); Ceard; Juan, near Ig-
uatu (Becker, 1913, FMNH); BOLIVIA: La Paz;
Cochabamba; Incachaca (Miller and Boyle, 1915,
AMNH); Santa Cruz; PARAGUAY: Boqueron;
Fortin Guachalla (Willim, 1945, FMNH); Presi-
dente Hayes; Fortin General Genes [= General
Genes], 1.6 km NE of Asuncion (Myers, 1972,
MVZ); Villa Hayes, 24 km NW (Creighton, 1979,
UMMZ; Myers, MVZ); Central; Recoleta, Asun-
cion (Myers, 1977, AMNH; Myers, 1977, 1979,
UMMZ); Paraguari; Sapucay [= Sapucaf] (Fos-
ter, 1901, 1902, 1903, USNM); Parque Nacional
Ybycuf (Creighton, 1979, UMMZ; Dobson, 1979,
LSUMZ, UMMZ; Myers, 1979, LSUMZ, UMMZ);
Saltos de Pirareta, not located (Creighton, 1979,
UMMZ); Misiones; San Ignacio, 20 km W
(Hibbs, 1965, USNM); Ayolas, 5 km ENE (My-
ers, 1977, AMNH; Myers, 1977, UMMZ); Amam-
bay; Cerro Cora (Myers, 1977, UMMZ); Canen-
diyu; Curuguaty (Myers, 1979, UMMZ); Itapua;
Puerto San Rafael [= San Rafael] (Creighton,
1979, UMMZ); Encarnacion (Creighton, 1979,
UMMZ; Talbot, 1976, UMMZ); Caaguazii: Es-
tancia San Ignacio, 24 km NNW of Carayao (My-
ers, 1979, UMMZ); Coronel Oviedo (Myers,
1976, UMMZ); Colonia Sommerfield no. 11
(Wharton, 1950, USNM); Guaira; Villarica (Gil-
more, 1940, AMNH; Schade, 1923, AMNH); La
Cordillera; Altos, 20 km N (Myers, 1973, MVZ);
Tobati (Myers, 1979, UMMZ); URUGUAY: Ar-
tigas; Belen del Dragon, 6 km NNW (Langguth,
1963, AMNH); Rio Negro; Arroyo Negro, 15 km
S of Paysandu (Tuttle, 1963, AMNH; Ximinez,
1963, AMNH); Soriano; Estancia Concordia, 15
mi SW of Dolores (Sanborn, 1926, 1927, FMNH);
Cardona, 3 km E (Tuttle, 1963, AMNH); Colonia;
La Lata, 10 km S (La Lata is 40 km N of Rosa-
rio), (Sanborn, 1927, FMNH); Durazno; San Jor-
ge, Rio Negro, 15 km NNW (Barlow, 1963,
AMNH); Cerro Largo; Las Manas (Anderson,
1963, AMNH); Estancia Melo, 6 km SE (Barlow,
1963, AMNH; Koopman, 1963, AMNH; Lang-
guth, 1963, AMNH; Tuttle, 1963, AMNH; Xim-
enez, 1963, AMNH); Trienta y Tres; Boca del Rib
Tacuari (Barlow, 1963, AMNH; Tuttle, 1963,
AMNH; Ximenez, 1963, AMNH); Quebrada de
los Cuervos, 43 km N of Trienta y Tres (Sanborn,
1926, FMNH); ARGENTINA: Jujuy; Yuto (Con-
tino, 1957, 1958, 1960, AMNH); Salta; Rosario
BROWN: ATLAS OF NEW WORLD MARSUPIALS
29
de Lerma (Miller and Boyle, 1916, AMNH); Tu-
cumdn; Sarmiento [= Los Sarmientos] (Miller
and Boyle, 1916. AMNH); Corrientes: Itati
(Nachman, 1989, UMMZ): Corrientes, 20 km SE
(Thorington, Jr., 1980. USNM); Misiones; Foz do
Igua^u. 14 km NE, Itaipu Binacional, Refugio
Bela Vista (Mustrangi, 1993, MVZ); Misiones,
ARGENTINA Parana, BRAZIL; San Antonio
(Myers, 1976. UMMZ).
Didelphis aurita Wied-Neuwied, 1826
Didelphis azarae Temminck, 1824:30 (type locality,
none given, here restricted to eastern Sao Paulo:
holotype name designated: description based on
some juveniles in the Paris Museum, adults and
juveniles in the Vienna Museum and Frankfurt Mu-
seum. Gray. 1843:100 part. BRAZIL. Bertoni,
1923:51 PARAGUAY. Gardner. 1993:13 listed.
D[idelphis] azarae: Wagner. 1843b:38 part, descrip-
tion, part, synonymy. Wagner. 1850:126 a black-
eared opossum, not Azara's white-eyed micoure
premier; BRAZIL. Wagner. 1855:223 aurita
Wied-Neuwied. a synonym. Burmeister, 1856:6.
Thomas. 1888:328.
D[idelphys] aurita Wied Neuwied. 1826:395 BRA-
ZIL: Bahia (Villa Vicosa [= Nova Vicosa], Rio
Perufpe). Miranda-Ribero. 1936:336 BRAZIL:
Santa Catarina (Humboldt): Rio de Janeiro (Ther-
ezopolis).
Didelphys aurita: Burmeister. 1854:130 BRAZIL.
Burmeister. 1856:64 BRAZIL: Rio de Janeiro;
azarae Temminck. a synonym. Pelzeln. 1883:
109 BRAZIL: Sao Paulo (Rio Ipanema). Ihering.
1893:99 BRAZIL: Rio Grande do Sul. Ihering.
1894:10 BRAZIL: Sao Paulo.
Didelphis aurita: Lund. 1839:233 BRAZIL: Minas
Gerais (Rio das Velhas). Hensel. 1869:290 BRA-
ZIL: Rio Grande do Sul. Goeldi. 1894:457 BRA-
ZIL: Rio de Janeiro (Serra dos Orgaos). Ihering.
1914:343 part, synonyms; koseritzi Ihering. nigra
Ihering; BRAZIL: Rio Grande do Sul. Miranda Ri-
beiro. 1936:336 BRAZIL: Santa Catarina. Vi-
eira. 1949:342 BRAZIL: Sao Paulo; Espirito
Santo; Minas Gerais; Rio de Janeiro: Rio Grande
do Sul. Avila Pires. 1965:4 BRAZIL: Bahia (type
locality. Villa Vicosa [= Maroba = Nova Vicosa],
Rio Perufpe). lectotype. by selection, male, skin
mounted with skull. AMNH 836. Cerqueira. 1985:
141 BRAZIL: Alagoas (Sao Miguel dos Cam-
pos): Bahia (Bahia do Todos Santos [= Reconca-
vo]). Pires et al.. 1999:41 BRAZIL: Rio de Ja-
neiro (Poco das Antas Biological Reserve). Patton
et al.. 2000:47 BRAZIL: Bahia (Fazenda Beijo
Grande. 12 km S. 1 km E of Itabuna): Espirito
Santo (Mata da Caixa D' Agua). 1 .7 km W of Santa
Tereza): Parana (Fazenda Guaricana. Guaratuba.
Morretes): Sao Paulo (Fazenda Intervales, Base
Saibadela. Sete Barros). Mares and Braun, 2000:
32 ARGENTINA: Misiones (junction of High-
way 14 and 2 de Mayo Road, not located: junction
of Highway 21 and Arroyo Oveja Negra. 2 km W
of Parque Nacional Mocona. not located; Deseado;
Parque Nacional Iguazii: Puerto Iguazii; Puerto
Schwelm. not located; Reserva Natural Estricta San
Antonio: Rio Iguazii).
Didelphys marsupialis aurita: Cope, 1 889: 1 29 (un-
certain locality).
Didelpfns marsupialis aurita: J. A. Allen, 1902:
265,278 BRAZIL: Rio de Janeiro ("Rio Rodeio,
Serra do Mar. near Rio de Janeiro"); Sao Paulo
(Cruzeiro: Piquete); Rio Grande do Sul (Taquara).
Yepes. 1936:34 BRAZIL: Rio Grande do Sul;
Sao Paulo; ARGENTINA: La Rioja (Villa Castelli)
Yepes. 1938:40 BRAZIL: Rio Grande do Sul;
PARAGUAY ARGENTINA: La Rioja (Villa Cas-
telli). Travassos. 1945:494 BRAZIL: Espirito
Santo (Rio Itauna). Carvalho. 1965:251 BRA-
ZIL: Sao Paulo (Boraceia. Estacao Biologica de
Boraceia). Crespo, 1974:2 ARGENTINA; Mi-
siones (Deseado: Rio Iguazii). Tyndale-Biscoe and
Mackenzie. 1976:260 BRAZIL: Rio de Janeiro
(Teresopolis; Jardim Botanico). Crespo. 1982a:
59 ARGENTINA: Misiones (Parque Nacional
Iguazii). Massoia et al.. 1987:1 16 ARGENTINA:
Misiones (Rio Urugua-i'). Fernandez et al., 1988:
221 BRAZIL; Rio de Janeiro (Ilha Grande; Ita-
curuca; Jaguanum).
Didelphis marsupialis L. subsp. Aurita: Luederwaldt.
1929:27 BRAZIL: Scio Paulo (Ilha de Sao Sebas-
tiao). Kumbiegel, 1941a:42 part, BRAZIL: Sao
Paulo (Serrinha; Serra do Bananal: Juquia: Serra
do Mar: Mato Grosso (Rio Ivinheima [now in Mato
Grosso do Sul]): ARGENTINA: Misiones; PAR-
AGUAY: San Pedro (Nueva Germania).
Didelphis marsupialis van rvpica: Thomas. 1888b:
323 part. BRAZIL: Rio de Janeiro.
Gamba aurita var. Brasiliensis Lias. 1872:329 new
name combination for aurita Wied-Neuwied. Gard-
ner, 1993:16 listed.
Didelphys koseritzi Ihering. 1893:99 BRAZIL: Rio
Grande do Sul (type locality: holotype presumably
in the Sao Paulo Museum, presented by Sefior Th.
Bischoff). Ihering, 1894:10 BRAZIL: Sao Paulo.
Gardner. 1 993: 1 6 listed.
[Didelphys (Marmosa)] koseritti: Trouessart, 1898:
1 240 incorrect spelling of koseritzi Ihering. 1893;
classification.
Didelphis koseritzi: Avila Pires. 1994:371 BRAZIL:
Rio Grande do Sul (Taquara).
Didelphxs cancrivora (not Gmelin. 1788): Hensel:
1873:114 BRAZIL: Rio Grande do Sul. Pelzeln,
1883:110 BRAZIL: Rio de Janeiro (Sapitibi [ =
Sepetiba]: Registo do Sai). Winge. 1893:8,49
BRAZIL: Minas Gerais (Lagoa Santa. Recent and
Pleistocene). Bertoni. 1914:68 PARAGUAY. Ber-
toni, 1939:6 PARAGUAY
Didelphis marsupialis: Thomas, 1902d:64 BRA-
ZIL: Parana (Roca Nova). Davis, 1945a: 119
BRAZIL: Rio de Janeiro (Teresopolis). Davis,
1945b:280 BRAZIL. Davis. 1947:2 BRAZIL.
Vieira, 1953:219 BRAZIL: Alagoas (Sao Miguel
dos Campos; Mangabeiras). Varejao and Valle,
1982:42. 43 BRAZIL: Bahia (Ilheus; Nova Vi-
cosa): Minas Gerais (Abre Campo; Alem Paraiba;
Alfenas; Araponga; Barao de Cocais; Barra Longa:
Bela Vista de Minas; Cachoeira do Campo; Caete.
Cidade; Campanha; Canaa; Carmo de Cachoeira;
30
FIELDIANA: ZOOLOGY
Fazenda da Cascata; Guaraciaba; Itabira; Itabirito;
Jequitinhonha; Juiz de Fora; Lambari; Leopoldina;
Manhuacu; Monte Bello; Ouro Preto; Parque For-
estal Estadual do Rio Doce; Parque Nacional de
Caparao; Pecanha; Pogos de Caldas; Ponte Nova;
Prados; Sabinopolis; Santa Maria de Itabira; Santa
Rita do Sapucaf; Sao Domingos do Prata; Sao Joao
de Jacutinga; Teofilo Otoni; Uba; Viscosa; Viscon-
de do Rio Branco; Volta Grande); Rio de Janeiro
(Nova Friburgo; Parque Nacional de Itatiaia; Rio
de Janeiro; Teresopolis); Espirito Santo (Nova Ve-
necia; Ponta da Fruta; Santa Teresa; Sao Francisco;
Venda Nova; Vitoria). Yonenaga Yasuda, 1982:
72 BRAZIL: Sao Paulo (Boraceia, Sao Paulo).
Santori et al., 1996:307 BRAZIL: Rio de Janeiro
(Barra de Marica). Caceres and Montero-Filho,
1998:169 BRAZIL: Parana (Curitiba).
Didelphis marsupialis marsupialis: Avila Pires and
Gouvea, 1977: 10 BRAZIL: Rio de Janeiro (Mon-
te Serrat [= Pico das Agulhas Negras]).
Didelphis aurita longipilis Miranda Ribeiro, 1935:
35 BRAZIL: Rio de Janeiro (type locality, Co-
lonia Alpina, Teresopolis; cotypes, 2 males, MNRJ,
collected by Senor Reinisch). Avila Pires, 1968:
169 BRAZIL: Rio de Janeiro (Colonia Alpina).
Gardner, 1993:16 listed.
D[idelphis] longipilis: Miranda Ribeiro, 1936:338,
footnote 1 BRAZIL: Rio de Janeiro (type local-
ity, Teresopolis; cotypes MNRJ M6, M7).
Did[elphis aurita] melanoides Miranda Ribeiro, 1935:
36 BRAZIL: Miranda Ribeiro, 1935:35 BRA-
ZIL: Rio de Janeiro (type locality, Teresopolis; ho-
lotypes or cotypes in MNRJ). Gardner, 1993:16
listed.
Holotype Female, skin and skull, presumably
in the Wied-Neuwied Museum [cf. holotype of
azarae Temminck].
Type Locality Villa Vicosa [= Nova Vic;o-
sa], Rio Peruipe [= Rio Peruhybe], Bahia, Brazil.
Distribution (Fig. 13) Atlantic forest of
southeastern Brazil, southeastern Paraguay and
northeastern Argentina.
Additional Records BRAZIL: Bahia; Uru-
cutuca (1944, 1945, KU; Rockefeller Institute,
1944, FMNH); Rio do Braco (1944, KU); Fazen-
da Almada (Rockefeller Institute, 1944, FMNH);
Fazenda Pirataquisse (1944, KU); Sao Paulo;
Guaratuba (de Souza Lopez, 1972, USNM); Mor-
retenho (Olalla, 1961, FMNH); Ribeirao Fundo
(Ollala, 1961, FMNH); Rocha (Ollala, 1960,
FMNH); Quebrado Penteado (Ollala, 1961,
FMNH); Primeiro Morro (Ollala, 1960, FMNH);
Estacao Biologica de Boraceia (Mustrangi, 1994,
MVZ); Santa Catarina; Joinville [= Joenvile]
(Schulter and Mass, FMNH); Rio de Janeiro; Pa-
raiba do Sul (Gilmore, 1937, AMNH); Guanabara,
Baia de (Gilmore, 1937, AMNH); Gavea, Rio de
Janeiro (Gilmore, 1937, AMNH); Tijuca (Gil-
more, 1937, AMNH); Trapicheiro (Whitman,
1938, AMNH).
Didelphis marsupialis Linnaeus, 1758
[Didelphis] marsupialis Linnaeus, 1758:54. Gardner,
1993:16 listed.
0[possum]. Marsupialis: Schmid, 1818:116 classi-
fication.
S[arigua] marsupialis: Muirhead, 1819:429 part,
new combination.
Didelphis marsupialis: Rendall, 1897:342 TRINI-
DAD AND TOBAGO: Tobago (Tobago); St. Vin-
cent. J. A. Allen, 1900a:185. J. A. Allen, 1901d:
166, 182, 183 part (type locality restricted to
Valley of Mexico on basis of the tlacuatzin of
Hernandez). J. A. Allen, 1901e:92 name now re-
moved from the Virginian opossum and restricted
to the Tlacuatzin or Mexican opossum of Hernan-
dez, 1651. Thomas, 190 la: 153 GUYANA: Upper
Takutu-Upper Essequibo (Kanuku Mts.); name
based on the philander of Seba (not Linnaeus); type
locality, "Guiana." J. A. Allen, 1902:257, 276
name now regarded as based on philander maxi-
mum orientalis Seba; type locality "Guiana"; dis-
tribution; Guiana, eastern Venezuela, northern
BRAZIL. Menegaux, 1902:495 FRENCH GUI-
ANA: (Manicore, not located). J. A. Allen, 1904d:
337 VENEZUELA: Bolivar (Ciudad Bolivar).
Osgood, 1910:39 VENEZUELA: Carabobo (Lake
[= Lago] Valencia). Thomas, 19 10b: 188 GUY-
ANA: Pomeroon-Supenaam (Supenaam River).
Thomas, 1911:143 SURINAME: (type locality);
Linnaean name based primarily on reference to
Seba. Osgood, 1914b: 147 PERU: San Martin
(Moyobamba); Loreto (Yurimaguas). Ihering,
1914:342 distribution. Pohle, 1927:244 BRA-
ZIL: Pernambuco (Dois Irmaos; Fundao); Paraiba
(Penha). Thomas, 1928a:264 PERU: Loreto (Sar-
ayacu). Gilmore, 1941:316 distribution. Harris,
1943:1 COSTA RICA: Cartago (El Muneco).
Tate, 1947:65 VENEZUELA: Aragua (Rancho
Grande, 1100 m). Sanderson, 1949:786 SURI-
NAME: Paramaribo. Vieira, 1952:31 BRAZIL:
Acre (Iquiri). Sanborn, 1953:4 PERU: Puno (San
Juan, Sandia, not located). Veiria, 1953:219
BRAZIL: Alagoas (Magabeira). Kuhlhorn, 1955:
163 BRAZIL: Mato Grosso. Guallart, 1962:
157 PERU: Amazonas (upper Rio Maranon). Oli-
vares, 1962:307 COLOMBIA: Meta (Rio Guaya-
bero). Pirlot, 1963:175, 178, 180 VENEZUELA:
Zulia (Kasmera, Rio Yasa). Avila Pires, 1964:12
BRAZIL: Amazonas (km 50, Manaus-Itacoatiara).
Alvarez and Ferrusquia, 1967:107 MEXICO:
Aguas Calientes (Aguascalientes [= Aguas Calien-
tes] [Pleistocene]). Handley, 1976:9 VENEZUE-
LA: Merida (Tabay); Barinas (Altamira); Miranda
(Alto No Leon [= No Leon, now in Distrito Fed-
eral], 31 km WSW; Caracas [now in Distrito Fed-
eral]; Rio Chico; Instituto Venezolano de Investi-
gaciones Cientificas [= I VIC]; Parque Nacional
Guatopo [now in Miranda-Guarico]; San Andres);
Amazonas (Belen; Boca Mavaca; Capibara; Puerto
BROWN: ATLAS OF NEW WORLD MARSUPIALS
31
90 80 70 60 50 40 30 20 10
Fig. 13. Distribution of Didelphis aurita: southeastern Brazil, southeastern Paraguay, and northern Argentina.
32
FIELDIANA: ZOOLOGY
Ayacucho; San Juan; Tamatama); Tdchira (Las
Mesas); Distrito Federal (Boca de Tigre Valley;
Caracas; Hacienda Carapiche; Los Venados); Tru-
jillo (La Ceiba; Valera); Falcon (Mirimire; Boca de
Yaracuy; Rio Socopito); Lara (Caserio Boro); Zulia
(Cerro Azul; El Rosario; Kasmera); Apure (Rio
Cinaruco; Hato Cariben; Nulita); Anzodtegui (Clar-
ines); Sucre (Cumana; Ensenada Cuaranta; Mana-
cal); Guarico (Estacion Biologica de los Llanos);
Bolivar (Hato La Florida; Icabaru, 23-45 km NE;
Hato San Jose; km 125, 85 km SSE of El Dorado;
Rio Supamo); Monagas (Hato Mata de Bejuco; San
Agustin); Carabobo (Montalban); Aragua (Esta-
cion Biologica de Rancho Grande); Yaracuy (Mi-
nas de Aroa); Distrito Federal-Miranda (Pico Avi-
la); Carabobo- Falcon-Yaracuy (Urama). Mendez,
1977:166 COLOMBIA: Valle del Cauca (Buga;
Lago Calima; La Maria; Pichinde); Choco (Rio
Curiche). Reig et al., 1977:211 VENEZUELA:
Miranda (Carrizal); Merida (Bejuquero); Aragua
(Bahia de Cata; Rancho Grande); PERU: Ucayali
(Rio Curanjo, Balta). Husson, 1978:32 SURI-
NAME: Marowijne (Bigisanti); Sipaliwini (Oele-
marie Airstrip; Lucie River); Commewijne ("Cle-
via"; Nieuw Amsterdam); Paramaribo (Cultuur-
tuin; Paramaribo); Saramacca (Groningen); Nickeri
(Sipaliwini Airstrip; Wageningen); Coronie (Tot-
ness). Eisenberg et al., 1979:187 VENEZUELA:
Guarico (Fundo Pecuario Masaguaral); Miranda-
Guarico (Parque Nacional Guatopo). O'Connell,
1979:73 VENEZUELA: Guarico (Fundo Pecu-
ario Masaguaral); Miranda-Guarico (Parque Na-
cional Guatopo). Tyndale-Biscoe, 1980:711-712
COLOMBIA: Valle del Cauca (Buenaventura;
Cali; Pichinde); Meta (Finca El Buque). Terborgh
et al., 1984:1-29 PERU: Madre de Dios (Cocha
Cashu Biological Station, Parque Nacional del
Manu). Cerqueira, 1985:140 ECUADOR: Guay-
as (Ancon); PERU: Lambayeque (Eten; Lamba-
yeque); La Libertad (Pacasmayo); Piura (Piura).
Gardner, 1988:100 VENEZUELA: Amazonas
(Base Camp, Cerro de La Neblina) George et al.,
1988:39 BRAZIL: Para (Parque Nacional da
Amazonia). Perez-Hernandez, 1989:381-386
VENEZUELA: Yaracuy (Agua Negra; Norte de
Yaritagua; Sierra de Aroa); Zulia (Alguacil; Chi-
rulf); Distrito Federal (Alta Florida; Caracas; Car-
retera El Limon; Cerro El Avila; Urb. El Marques;
Corozal, Parque Nacional "El Avila"; Parque Na-
cional "El Avila"; El Eden, Parque Nacional "El
Avila"; El Encantado, Hacienda El Limon; La
Conchita, Hacienda El Limon; Los Chorros; Los
Venados, Parque Nacional El Avila; Zona Guaca-
tal. Hacienda El Limon; Suapo, Hacienda El Li-
mon); Miranda (Altos de Pipe, Instituto Venezo-
lano de Investigaciones Cientificas [I VIC]; Baruta;
Carrizales; La Cienega; La Providencia; La Toma
Capaya; La Union; El Hatillo; Los Guayabitos; Los
Teques; Turgua; Cortada del Guayabo); Carabobo
(Bahfa de Patanemo; Camino de los Espafioles; Los
Dos Bocas; Las Quiguas; Los Campamentos; Punto
Cabito; Valle Pascua); Merida (Bejuquero; Carlo
Tigre; Hacienda San Pedro, Santa Cruz de Mora
[= Santa Cruz]; Mina Carbon; Zea); Guarico (Car-
retera Calabozo-San Fernando de Apure; Dos Cam-
inos; El Socorro; Estacion Biologica de los Llanos,
Calabozo; Morichal Villasmil, Calabozo); Aragua
(Centra de Investigaciones de Fauna Silvestre; Es-
tacion de Piscicultura; Hacienda La Esperanza);
Portuguesa (Colonia Agricola Turen); Delta Ama-
curo (Giiiniquina; Los Guires); Cojedes (Carretera
San Carlos-Manrique; Hato El Tirado; Itabana al
Sur de las Vegas; Los Chupones); Anzodtegui (Pe-
kin Abajo; Valle Guanape). Timm et al., 1989:16
COSTA RICA: Heredia (Parque Nacional Braulio
Carrillo). Julien-Laferriere and Atramenwiez, 1990:
405 FRENCH GUIANA (Cabassou; Piste de
Saint Elie). Emmons, 1990:73 BOLIVIA: La Paz
(Rio Machariapo). Albuja, 1992:124 ECUADOR:
Guayas (Cerro Blanco); Los Rios (Jauneche). Em-
mons and Albuja, 1992:123 ECUADOR: Guay-
as-Manabi (Parque Nacional Machalilla). Pacheco
et al., 1993:5 PERU: Cusco (Hacienda Erika).
Emmons and Romo, 1994:140-143 PERU: Ma-
dre de Dios (Ccolpa de Guacamayas). Emmons et
al., 1994:146-149 PERU: Madre de Dios
("Aguas Claras" Camp; Puesto San Antonio; Re-
fugio Juliaca). Catzeflis, 1997:231 FRENCH
GUIANA (Sinnamary River; Crique Tigre). Adler
et al., 1997:361 COLOMBIA: Cordoba (El Con-
tento). Patton et al., 2000:47 COSTA RICA: Pun-
tarenas (Wigdale House; Monteverde); PANAMA:
Bocas del Toro (Isla Popa, 1 km E of Sumwood
Channel; Punta Alegre, Peninsula Valiente); GUY-
ANA: East Berbice-Corentyne (Mapenna Creek,
ca. 6 km from Corentyne River); Barima-Waina
(Baramita); ECUADOR: Napo (Parque Nacional
Yasuni, 20 km S of Pompeya Sur); BRAZIL: Para
(Floresta Nacional Tapirape-Aquiri, Municipio de
Maraba); Amazonas (Ilhazinha, left bank Rio Jurua
on Igarape Arabidi, affluent of Parana Breu; Ser-
ingal Condor, left bank Rio Jurua; Barro Vermelho,
left bank Rio Jurua; Estrada Picarreira, right bank
Rio Cauaburi, Parque Nacional do Pico da Neblina;
right bank Rio Jau above mouth; Tambor, left bank
Rio Jau; Lago Meduinim, left bank Rio Negro; Boa
Esperanca, right bank Rio Jurua; Jainu, right bank
Rio Jurua; Ilha Paxiuba, right bank Rib Jurua; Lago
Vai-Quem-Quer, right bank Rib Jurua; Penedo; Al-
tamira); Acre (Igarape Porongaba, opposite, left
bank Rio Jurua; Igarape Porongaba, right bank Rio
Jurua; Nova Vida; Sobral). Voss et al., 2001:23
FRENCH GUIANA: (Piste St. Elie [= Piste St.
Elie Ecerex Station]; Petit Saut; Paracou) SURI-
NAME: Brokopondo (Brownsberg Nature Park);
Coronie (Totness); Saramacca (Bigi Poika).
Didelphys marsupialis: Goldfuss, 1809:215 classi-
fication. Thomas, 1893:10 TRINIDAD AND TO-
BAGO. Goeldi and Hagmann, 1 894:99 BRAZIL:
Para. Krumbiegel, 1941a:32, 42 distribution.
Sanderson, 1949:775, 786 SURINAME.
Didelphis marsupialis marsupialis: J. A. Allen, 191 1:
247 VENEZUELA: Bolivar (Peru Mines, El Cal-
lao). J. A. Allen, 1916b:562 BRAZIL: Amazonas
(lower Rio Solimoes, not located). Anthony, 1921:
267 GUYANA: Essequibo (Kartabo [now in Cu-
yuni-Mazaruni]). Tate, 1939:160 part, VENE-
ZUELA: Bolivar (Cerro Auyan-tepuf [= Auyan-te-
puf]); Amazonas (Cerro Duida); GUYANA: Cuyu-
ni-Mazaruni (Mt. Roraima). Borrero and Hernan-
dez-Camacho, 1957:213 COLOMBIA: Santander
(San Vicente de Chucuri). Vieira, 1957:125
BROWN: ATLAS OF NEW WORLD MARSUPIALS
33
BRAZIL: Maranhao (Barra do Corda, Rio Mear-
im). Carvalho, 1961:5 BRAZIL: Roraima (Para-
na do Pocao, Caracaraf; Pocao, Boa Vista). Car-
valho, 1962:283 BRAZIL: Amapd (Vila Velha do
Cassipore, Oiapoque; Rio Maruanum; Macapa;
Cachoeira do Pancada [= Prosperidade], Rio Ma-
raca, Mazagao). Butterworth and Starret, 1964:3
VENEZUELA: Monagas (Maturfn, 32 km SE).
Anderson, 1997:145 BOLIVIA: Beni (Guayara-
merin; Riberalta; Rio Itenez, Costa Marquez, op-
posite [BRAZIL]; Rio Itenez, S of and 4 km SW
of Costa Marquez; Horquilla, Rio Machupo, 15 km
above; Versalles; Puerto Siles; Azunta; Camino
Vilches; San Joaquin; Aserradero Monte de San
Pablo; San Pablo Village; Exaltacion, 8 km N; Lar-
angeiras, 20 km W; Bahia de los Casara; Barran-
quita; Piedras Blancas; Yutiole; Rib Mamore, W
side; Los Totumu; Rio Tijamuchi, 5 km above
mouth; Rurrenabaque; Rio Ibare, mouth; Rio Ibare,
27 km above mouth; Ibiato; El Trapiche, 2 km E;
Campamento El Trapiche; Reseva Biologica del
Beni; San Ignacio de Moxos; Casarabe; La Em-
bocada; San Antonio, 10 km E; Camiaco; Rib Ma-
more 5 km NW mouth Rio Grande); Cochabamba
(Campamento Yuqui; Todos Santos; Villa Tunari, 2
km E; Hotel Las Pozas, Villa Tunari; Yungas; Villa
Tunari, 3 km SW; Chapare); La Paz (Rio Beni;
Mapiri; Puerto Linares; Guanay, Rio Challana, 5
km SE; Palos Blancos; Entre Ribs, 2 km W, not
located; La Reserva; Caranavi; El Vertego, Zongo
Valley; Chijchipani; Chijchijpa; Yuncas; Yolosito;
Astillero; Pitiguaya; Chulumani); Pando (Cobija,
10 km S; Rfo Nareuda; Manuripi Health National
Wildlife Reserve; Isla Gargantiia); Santa Cruz (Los
Palmares; Totaitu; Rio Ichilo, 54 km S mouth Rfo
Chapare; Zapoco; Yapacani; Santa Rosa, 7 km N;
San Miguel Rincon; Tocomechi; Buenavista, 7 km
N and 17 km W; Warnes; Santa Rosita; San Rafael
de Amboro; Santa Cruz de la Sierra; km 5 on road
to Cochabamba).
D[idelphis] m[arsupialis] marsupialis: Miranda Ri-
beiro, 1936:333 BRAZIL: Maranhao; Para. Car-
valho, 1961:5 BRAZIL: Rio Branco [now Rorai-
ma] (Po^ao; Boa Vista; Parana do Poao, Caracar-
af).
Didelphis m. marsupialis: Carvalho, 1961:5 BRA-
ZIL: Roraima (Pocao, Boa Vista; Parana do Pocao,
Caracaraf).
Didelphis marsuapialis marsuapialis: Avila Pires,
1964:8, 12 incorrect subsequent spelling for Di-
delphis marsupialis marsupialis Allen, 1911
BRAZIL: Amazonas (km 50, Manaus-Itacoatira
RR).
0\possum] marsupialis: Schmid, 1818:16 classifi-
cation.
Didelphis karkinophaga Zimmerman, 1780:226
FRENCH GUIANA: (type locality, Cayenne);
name based on Buffon's crabier. J. A. Allen and
Chapman, 1897:23 TRINIDAD AND TOBAGO:
Victoria (Princestown); Caroni (Caparo); St.
George (Caura); DOMINICA. Bangs, 1900:89
COLOMBIA: La Guajira (Pueblo Viejo, Santa
Marta). J. A. Allen, 190 Id: 186 TRINIDAD AND
TOBAGO. Robinson and Lyon, 1901:137 VE-
NEZUELA: Disthto Federal (La Guaira). Gardner,
1993:17 listed.
[Didelphis] Carcinophaga Boddaert, 1784:77 the
crabier of Buffon.
Did[elphis] carcinophaga: Cuvier, 1798:124.
[Didelphis] cancrivora Gmelin, 1788:108 FRENCH
GUIANA: (type locality, Cayenne); name based on
the crabier of Buffon. Gardner, 1993:17 listed.
Didelphis cancrivora: Kerr, 1792:195 Cayenne
opossum. Jentink, 1887:300 SURINAME: (spec-
imen figured by Temminck). Jentink, 1888:220
SURINAME.
Didelphis Cancrivorus: Shaw, 1800:476 incorrect
ending of specific name. E. Geoffroy, 1803:139
FRENCH GUIANA: (Cayenne).
Didelphys cancrivora: Waterhouse, 1841:85 BRA-
ZIL; GUYANA. Waterhouse, 1 846:473 northern
parts of South America. Tomes, 1860:268 EC-
UADOR. Burmeister, 1854:129 BRAZIL.
D[idelphvs] cancrivora: Cabanis and Schomburgk,
1848:777 GUYANA. Wagner, 1843:41 GUYA-
NA; BRAZIL.
D[idelphis] azarae: Tschudi, 1844:143 PERU: part,
tropical zone only.
Didelphis azarae: Jentink (not Temminck, 1824),
1888:219 SURINAME.
Didelphys aurita: Pelzeln, 1883: 109 BRAZIL:
Amazonas (Borba, Rfo Madeira); Mato Grosso
(Matogrosso).
Didelphis aurita: Ihering, 1904:425 BRAZIL: Ama-
zonas (Rfo Jurua).
Didelphis marsupialis aurita: Miiller (not Wied-Neu-
weid), 1914:8 BRAZIL: Para (Peixe-boi, Be-
lem). Krumbiegel, 1941:42 BRAZIL: Para (Ilha
Mexiana). Anderson et al., 1993:19 BOLIVIA:
Santa Cruz (Buena Vista; Buena Vista, near; Co-
chabamba; San Miguel Rincon; San Rafael de Am-
boro; Santa Cruz de la Sierra; Santa Rosita; Toco-
mechi; Warnes; Rfo Yapacanf).
Didelphys marsupialis van aurita: Menegaux (not
Wied-Neuweid), 1902:495 FRENCH GUIANA:
(Ouanary River).
Didelphys marsupialis L. subsp. Aurita: Krumbiegel,
1941a:42 part, BRAZIL: Para (Ipitinga; Fazenda
Nazareth, Ilha Mexiana).
Didelphys marsupialis van typica: Thomas (not Lin-
naeus, 1758), 1888b:323 part, TRINIDAD AND
TOBAGO; BRAZIL: Para (Santarem); COLOM-
BIA: Antioquia (Concordia); VENEZUELA; GUY-
ANA: West Pomeroon-Supenaam Coast (Better
Hope); PERU: San Martin (Moyobamba). Krum-
biegel, 1941a:42 part, VENEZUELA: Aragua
(Maracay). Gardner, 1993:17 listed.
[Didelphys (Marmosa)] koseriti: Trouessart, 1898:
1240 incorrect subsequent spelling of koseritzi
Ihering, 1893 classification.
Didelphis karkinophaga caucae J. A. Allen, 1900b:
192 COLOMBIA: Valle del Cauca (type locality,
Cali; holotype, female, skin and skull, AMNH
14192; collected June 1898 by J. H. Batty). Gard-
ner, 1993:17 listed.
Didelphis marsupialis caucae: J. A. Allen, 1902:
261 COLOMBIA: Valle del Cauca (Cali). J. A.
Allen, 1 91 6a:200 COLOMBIA: Antioquia (Puerto
Valdivia; La Frijolera; Barro Blanco; Dabeiba);
Valle del Cauca (Rfo Frio); Quindio (Salento);
Huila (San Agustfn; Andalucfa; Honda [now in To-
lima]). Anthony, 1923:4 COLOMBIA: Antioquia
34
FIELDIANA: ZOOLOGY
(El Poblado, near Medellfn). Gardner, 1973:69-71
(see for further information on localities in Mexico
and Central America) MEXICO: Tamaulipas
(Ejido Santa Isabel); San Luis Potosi (El Salto, Rib
Naranjo; El Naranjo [= Naranjos]; Rib Axtla; Xil-
itla; Tamazunchale; "Rancho Sabinal"); Hidalgo
(San Felipe Orizatlan); Veracruz-Llave (El Higo;
Hacienda Tamiahua, Cabo Rojo; Tuxpan; Tihu-
atlan; Papantla; Nautla; San Marcos; Tlapacoyan;
Las Minas; Jalapa; Jico; Mirador; Veracruz; Boca
del Rib; Fortm; Rio Metlac, 1 km W of Fortfn;
Potrero Viejo; Orizaba; San Juan de la Punta; Rio
Blanco. 20 km W of Piedras Negras; San Andres
Tuxtla: Catemaco; Lago Catemaco; Agua Dulce:
Jesus Carranza [= Santa Lucrecia]); Puebla (Met-
laltoyuca); Oaxaca (Yetla [= San Mateo Yetla];
Santo Domingo [= Santo Domingo Petapa]; Cerro
El Baiil, Rancho Carlos Minne; Tapanatepec [=
San Pedro Tapanatepec]); Tabasco (Alvaro Obre-
gon [= Frontera]; La Venta; Cardenas; Villa Her-
mosa; Mascuspana; Teapa; Rio Puyacatengo, 15
km E of Teapa); Chiapas (Palenque; Solusuchiapa;
Soyalo; El Real; Tuxtla; Chiapa de Corzo; Chiapa;
Comitan, 20 mi W; Zapaluta [= La Trinitaria], 18
mi E; Finca Ocuilapa; Risa de Oro; Finca Prusia;
Pa vol; Finca La Esperanza, 45 km N of Huixtla;
Finca Germania; Rib Huixtla, ca. 13 km N of Huix-
tla); Campeche (Apazote; Escarcega); Yucatan
(Chichen Itza); BELIZE: Cayo (Central Farm);
GUATEMALA: Peten (Toocoq); Aha Verapaz
(Chinaja; Chimoxan); Suchitepequez (Finca Sela-
che); Santa Rosa (Astillero); HONDURAS: Gra-
cias a Dios (Patuca; "Patuca River"); Atlantida
(La Ceiba); Cortes (San Pedro Sula); Santa Bar-
bara (Santa Barbara; Las Ventanas, Lago Yojoa);
Copdn (Copan); Lempira (Las Flores, Gracias);
Ocotepeque (Las Ventanas, near Belen Gualcho;
Nueva Octopeque); Francisco Morazdn (El Zapote:
Tegucigalpa); EL SALVADOR: Chalatenango
(San Jose del Sacare); Ahuachapdn (Barra de San-
tiago); Cuscatldn (Colima); San Miguel (Cerro Ca-
caguatique, N slope; Volcan de San Miguel; Lago
Olomega); Morazdn (Divisadero; Carolina Mine, 4
km W of Divisadero); Libertad (La Libertad); Son-
sonate (Hacienda Chilata); Usulutdn (Puerto del
Triunfo); NICARAGUA: Nueva Segovia (Jalapa);
Chinandega (Hacienda San Isidro; San Antonio);
Jinotega (Yali; Hacienda La Trampa, not located);
Matagalpa ("Lavala [= Savala]"; Santa Maria de
Ostuma; Finca Tepeyac); Zelaya ("Rib Coco"; Bo-
nanza; Kurinwas River; Rio Mico, S side El Re-
creo; El Recreo; Rib Escondido, 50 mi from Blue-
fields); Managua (Managua; Hacienda Azacualpa);
Boaco (Santa Rosa); Granada (Hacienda Mecate-
pe; La Calera, not located); Choluteca (Diriamba);
Chontales (Villa Somoza); Rio San Juan (La Es-
peranza; Toro Rapids; Greytown [= San Juan del
Norte]); COSTA RICA: Guanacaste (La Irma, Rfo
Abangares); Alajuela (Alajuela); Puntarenas (Mon-
teverde; San Geronimo Pirris; Finca Ligia, Parrita;
Pozo Azul; Palmar [= Palmar Sur]; Puerto Golfeto;
Camp Seattle, Osa Peninsula); San Jose (Finca Lor-
nessa; San Jose; San Pedro, Montes de Oca; Hatil-
lo; San Isidro del General [= Urefia]); Cartago
(Santa Teresa Peralta; Turrialba; Cartago); Limon
(Cariari; Pandora; "Talamanca"); PANAMA: Chi-
riqui (Cerro Punta; Palo Santo; Boquete; Concep-
tion; Boqueron; Progreso; Bambito; Tale; Guaba-
la); Bocas del Toro (Nievecita Farm: Boca del Dra-
go; Almirante); Veraguas (Santa Fe, Rio Santa Ma-
ria; Isla Coiba); Colon (Fort Sherman; Mojinga
Swamp; Camp Pina; Rio Indio; Gatun; Fort Davis;
Porto Bello; Colon); Code (El Valle); Los Santos
(Guanico; Cerro Hoya); Colon/Panama (Lion Hill;
Barro Colorado [= Barro Colorado Island]; Rio
Chagres); Panama (Juan Mina Station, Rib Cha-
gres; Rio Mandinga; Madden Road; Empire; Red
Tank; Fort Clayton; Curundu; Corozal; Ancon;
Quarry Heights; Fort Kobbe; Cerro Azul [= La
Zumbadora]; San Miguel Island [= Isla del Rey]);
San Bias (Mandinga; Armila); Darien (Rio Chu-
cunaque; El Real; Tacarcuna; Capeti; Cituro; Boca
de Rio Paya; Cerro Pirre; Cana; Jaque). Baker,
1974:135 ECUADOR: Esmeraldas (Cupa; Es-
meraldas; Quininde; Rib Verde); Guavas (Mira-
dor). Reig et al., 1977:21 1 COSTA RICA: San
Jose (Santa Ana). Wilson, 1983:443 COSTA
RICA: Puntarenas (Monteverde Forest Reserve;
Rincon; San Vito de Java); Heredia (Puerto Viejo
de Sarapaquf).
Didelphis karkinophaga colombica J. A. Allen.
1900b: 193 COLOMBIA: Magdalena (type local-
ity, Bonda, Sierra Nevada de Santa Marta; holo-
type, male, skin and skull, AMNH 15453, collected
19 April 1899 by H. H. Smith). Osgood, 1912:39
VENEZUELA: Zulia (El Panorama, Rfo Aurare).
Gardner, 1993:17 listed.
Didelphis marsupialis colombica: J. A. Allen, 1902:
260, 276 COLOMBIA: Cundinamarca (near Bo-
gota); Magdalena (Santa Marta). J. A. Allen,
1904d:4 18 COLOMBIA: Magdalena (Bonda;
Valparaiso).
Didelphis marsupialis cerca de colombica: Mendez,
1951:205 VENEZUELA: Miranda (El Hafillo,
La Union; Turgua).
Didelphis marsupialis tabascensis J. A. Allen, 190 Id:
1 73 MEXICO: Tabasco (type locality, Teapa; ho-
lotype, USNM 100512). W. B. Davis, 1944:374
MEXICO: Veracruz-Llave (Boca del Rio). Gardner,
1993:17 listed.
Didelphis marsupialis richmondi J. A. Allen, 190 Id:
175 NICARAGUA: Rio San Juan (type locality,
Greytown; holotype, adult female, USNM 33136/
45140). Harris, 1943:7 NICARAGUA: Rio San
Juan (Greytown); COSTA RICA: Cartago (El Mu-
neco). Gardner, 1993:17 listed.
Didelphis marsupialis insularis Rendall, 1897:342
TRINIDAD AND TOBAGO. J. A. Allen, 1902:
259, 276 TRINIDAD AND TOBAGO: Caroni
(type locality, Caparo, holotype, male, skin and
skull, AMNH 7740/6124; collected 19 March 1894
by Frank M. Chapman); DOMINICA; GRENADA;
ST. VINCENT. G. M. Allen, 191 1:193 TRINI-
DAD AND TOBAGO; GRENADA (import); ST
VINCENT (import). Hollister, 1912:93 MARTI-
NIQUE. Vesey-Fitzgerald, 1936:6 TRINIDAD
AND TOBAGO: (northern range). Beebe, 1952:
181 TRINIDAD AND TOBAGO: (Aruma Val-
ley, not located). Gardner, 1993:17 listed.
Didelphis marsupialis etensis J. A. Allen, 1902:262,
276 PERU: Piura (now in Lambaxaque) (type lo-
cality, Eten, 50 ft [= 15 m]); ECUADOR: Guayas
BROWN: ATLAS OF NEW WORLD MARSUPIALS
35
Fig. 14. Distribution of Didelphis marsupialis in Central America: Mexico south through Panama. See Figure 15
for distribution in South America, o = localities from unpublished records; = localities given in published records.
(Puna; Guayaquil); Esmeraldas (San Javier; Pam-
bilar): PANAMA: Chiriqui (Boquerdn: Boquete):
Panama (Isla San Miguel [= Isla del Rey]). J. A.
Allen, 1912a:74 COLOMBIA: Caldas (San Jose).
Osgood. 1914b: 148 PERU: La Libertad (Meno-
cucho: Hacienda Llagueda). J. A. Allen. 1916d:
116 ECUADOR: Esmeraldas; Guayas (Puna [=
Isla Puna]; Daule); Loja (now in El Oro) (Zaruma).
J. A. Allen. 1 9 16a:200 COLOMBIA: Caldas
(San Jose); Narino (Barbacoas): Owed (Bagado).
Thomas. 1920c: 247 PERU: Cusco (Santa Ana).
Thomas. 1927a:372 PERU: La Libertad (Pacas-
mayo); Amazonas (San Nicolas. 1364 m). Gardner.
1993:17 listed.
Didelphis marsupialis battyi Thomas. 1902e:137
PANAMA: (type locality. Coiba Island [= Isla Coi-
ba], Veraguas; holotype. old female. BMNH orig-
inal number 106). Gardner. 1993:17 listed.
Did[elphis] mesamericana mesamericana J. A. Allen.
1902:251 (part, not available from Oken. 1816).
Davis. 1944:374 part. MEXICO: Puebla (Rio
Otlati). Nuevo Leon (Rfo Ramos). Gardner. 1993:
17 listed.
Didelphis marsupialis particeps Goldman. 1917:
107 PANAMA: (type locality. San Miguel Island
[= Isla San Miguel]. Golfo de Panama. Panama;
holotype. adult female. Banas Collection. MCZ
8439). Gardner. 1993:17 listed.
Didelphis [sp.]: Guallan. 1962:157 PERU: Amazon-
as (upper Rio Maraiion).
Holotype None in existence, name based on
Seba's (1734:64) philander maximus orientalis,
Oldfield Thomas, 1911, by subsequent designa-
tion.
Type Locality SURINAME.
Distribution (Figs. 14 and 15) Tropical and
subtropical zones of Mexico from Tamaulipas
south through Central America into Panama, Co-
lombia. Ecuador, Peru, Venezuela, the Guianas,
Bolivia. Brazil, and Paraguay.
Additional Records MEXICO: Tarnaulipas;
Llera, Santa Isabel (Schaldach and Grissi. 1954,
KU); San Luis Potosi; Ebano (Dalquest, 1950,
LSUMZ); Bledos (Dalquest, 1950, LSUMZ); Ha-
cienda Capulin (Dalquest. 1950, LSUMZ); Axtla
(Dalquest, 1946, KU); La Joya. near Xilitla
(Shaw, 1947. LSUMZ); Veracruz-Have; Potrero
(Lee, 1960. KU); Tenochtitlan (Lackey, 1958,
UMMZ); Oaxaca; Acatlan (Miller, 1976,
UMMZ); Mitla, 25 mi S of Oaxaca (Eyster, 1968,
LSUMZ); Chiapas; Palenque. Ruinas de (Kilgore,
1965. KU); Pichucalco (Smith, 1965, KU); Cam-
peche; Isla del Carmen (Stanley, 1962, KU); BE-
LIZE: Cayo; Mt. Pine Ridge, not located (Murie,
1931. UMMZ); GUATEMALA: Peten; Uaxactun
(Murie. 1931, UMMZ): Tikal (Cantrall, 1956,
UMMZ): La Libertad (Jones, Jr., 1960, KU); San-
36
FIELDIANA: ZOOLOGY
| - II Didelphis marsupialis
10
20
30
40
50
90
80
70
60
50
40
30
20
10
Fig. 15. Distribution of Didelphis marsupialis in South America: throughout South America and Trinidad and
Tobago, but currently unknown in Chile, Argentina, and Uruguay. See Figure 14 for distribution in Central America.
o - localities from unpublished records; = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
37
ta Rosa; Finca El Progresso (de la Torre, 1951,
UMMZ): Suchitepequez; Salache (Anthony, 1924,
AMNH): EL SALVADOR: Morazdn; Monte
Cristo Mine, not located (Hildebrand, 1941,
MVZ); NICARAGUA: Managua; Lago Jiloa
(Kuhn, 1958, AMNH); Matagalpa; Hacienda Te-
peyac. near Matagalpa (Clark, 1914. KU); Boaco;
Boaco (Schlitter. 1967, KU; COSTA RICA: Pun-
(arenas; Jaco, 10 km N (Hafner, 1984. LSUMZ);
San Jose; Universidad [de Costa Rica], San Jose
(McPherson. 1966, LSU); Cartago; Santa Rosa
(Hafner, 1987, LSUMZ); Alajuela; Palmira de
Zarcero (Smith, 1936, FMNH); PANAMA: Chi-
riqui; Nicaragua Boquete, not located (Gaige,
1923, UMMZ): Panama; Balboa (Godfrey. 1948,
UMMZ); Tapia (1923, FMNH); Darien; Yaviza
(Cavalier, 1967. MVZ); La Laguna (1963,
UMMZ); COLOMBIA: Owed; Ungufa (Hersh-
kovitz, 1950. FMNH); COLOMBIA: Valle del
Cauca; Riolima. near San Antonio. W of Cali
(Batty. 1898. AMNH); San Jose (Richardson,
1910, AMNH): Rio Raposo (Thornton, 1962,
USNM); Caldas [= Dagua] (Richardson. 1910,
AMNH); Las Lomitas [= Lomitas] (Richardson,
1911, AMNH); Rio Bariatou (= Barratow), near
Cali (Batty, 1898, AMNH); Guengue (Richard-
son. 1911, AMNH); Cauca; Cerro Munichique
(von Sneidern, 1957, FMNH); Rio Saija (von
Sneidern, 1958, FMNH); Charguayaco (von Snei-
dern, 1956, FMNH); Sabanetas (von Sneidern,
1957, 1958, FMNH); Chisqufo (von Sneidern,
1957, FMNH); Totoro (von Sneidern, 1957,
FMNH); Popoyan (von Sneidern, 1957, FMNH);
Purace (Carriker Jr., 1956, USNM); Cauca Seca
(= Rio Cauca). not precisely located (Batty. 1898.
AMNH); Atldntico; La Pena (Hershkovitz, 1952.
FMNH); Cordoba; Socorre (Hershkovitz, 1949,
FMNH); Antioquia; Turbo (Hershkovitz, 1950,
FMNH); Zaragoza (Peterson, 1970, 1971.
USNM); La Cabana (Hershkovitz, 1950, FMNH);
Valdivia (Hershkovitz, 1950, FMNH); Urrao
(Hershkovitz. 1951, FMNH); Club Campestre, 1
mi S of El Poblado (Van Deusen, 1952, AMNH):
Sonson (Niceforo Maria, 1922, AMNH; 1923,
USNM); Huila; Villavieja (Miller, 1949, MVZ);
Naranjal, Valle de Suaza (Del Aguila, 1971,
USNM); Guapaton, Valle de Suaza (Del Aguila,
1971. USNM): San Isidro, Valle de Suaza (Del
Aguila, 1971. USNM): Picuma, Valle de Suaza
(Del Aguila. 1971. USNM); Marmato, Valle de
Suaza (Del Aguila. 1971. USNM); Avispero, Val-
le de Suaza (Del Aguila. 1971. USNM): El Alto,
Valle de Suaza (Del Aguila. 1971, USNM); Las
Quemadas, Valle de Suaza (Del Aguila, 1971,
USNM); Belgica, Valle de Suaza (Del Aguila,
1971, USNM); San Calixto, Valle de Suaza (Del
Aguila, 1971, USNM); Valle de Sauza (Del Agui-
la, 1971, USNM); San Adolfo (Hershkovitz,
1951, FMNH); Putumayo; Puerto Asis (Kirsch,
1969, KU); Rio Mecaya, Rio Caqueta (Hershko-
vitz, 1952, FMNH); Caqueta; Tres Troncas
(Hershkovitz, 1952, FMNH); Magdalena; Pueblo
Bello, Valledupar (Hershkovitz, 1942, USNM); El
Salado (Hershkovitz, 1942, USNM); Cesar; San
Sebastian (Hershkovitz, 1950, FMNH); Colonia
Agncola de Caracolicito (Hershkovitz. 1942,
USNM); Rio Guaimaral (Hershkovitz, 1942,
USNM); El Orinoco, Rio Cesar (Hershkovitz,
1943, USNM); Bolivar; San Juan Nepomuceno
(Hershkovitz, 1949, FMNH); Cundinamarca; San
Juan de Rioseco (Niceforo Maria, 1924, AMNH);
Meta; Guaicaramo [= Guaycarama] (Niceforo
Maria, 1926, AMNH): Finca El Capricho, Villa-
vicencio (Kirsch. 1969, KU); Villavicencio (Gil-
more, 1939, AMNH; Moreno, 1946, MVZ; Ni-
ceforo Maria, 1925, AMNH); Serrania de Macar-
ena [= La Macarena] (Dillon, 1942, AMNH); Ca-
ney [= Restrepo] (Boshell, 1939, AMNH; Wellan,
1937, AMNH); La Guajira; Honda Bay [= Bahia
Honda] (Hosves, 1913, AMNH); Villanueva
(Hershkovitz, 1942, USNM); Las Marimondas
(Hershkovitz, 1942, 1943, USNM); Norte de San-
tander; Rio Tarra (Hershkovitz, 1943, USNM);
Guamalito (Hershkovitz, 1943, USNM); Arauca;
Rio Arauca (von Sneidern, 1959, FMNH); EC-
UADOR: Manabi; Bahia de Caraquez (Tate,
1923, AMNH); Guayas; Cordillera de Colconche,
near Manglaralto (Hershkovitz, 1934, UMMZ);
Cerro Manglaralto (Anthony and Tate, 1923,
AMNH); Bucay (Tate, 1921, AMNH); Ventura
(Tate, 1922, AMNH); El Oro; Santa Rosa (Tate,
1921, AMNH); Portovelo (Anthony, 1920, AMNH);
Rio Pindo (Anthony, 1920, AMNH); Salinas,
(Anthony, 1920, AMNH); Loja; Alamor (An-
thony, 1920, AMNH; Tate, 1921, AMNH); San
Bartolo (Tate, 1921, AMNH); Guainche (Tate,
1921, AMNH); Limona, not precisely located
(Tate, 1921, AMNH); Los Rios; Limon (Tate,
1923, AMNH); Vinces (Tate, 1922, AMNH); Ha-
cienda Santa Teresi (McLean, 1974, USNM);
Chimborazo; Pallatanga (Tate, 1922, AMNH);
Puente de Chimbo (Tate, 1922, AMNH); Rio
Chimbo. Coto (Tate, 1922, AMNH); San Antonio
(Tate. 1922, AMNH); Zamora-Chinchipe; Zamora
(Anthony, 1920, AMNH); Napo; Santa Cecilia
(Lynch, 1967, KU); Limoncocha (Emmons, 1974,
USNM); San Jose Nuevo [= San Jose] (Olalla,
1924, AMNH); Rio Napo, not precisely located
38
FIELDIANA: ZOOLOGY
(von Baumann/Roosevelt, 1936, AMNH); Pasta-
za; Sarayacu (Tate, 1924, AMNH); Montalvo
(Olalla, 1932, FMNH); Copataza (Tate, 1924,
AMNH); PERU: Tumbes; Tumbes (Steere, 1873,
UMMZ); Piura; "Batan," on Zapalache-Carmen
Trail (Barkley, 1980, LSUMZ); Palambla (Wat-
kins, 1922, AMNH); Lambayeque; Motupe
(Smith, 1981, LSUMZ); Lima; Yangas (Pearson,
1968, MVZ); Cajamarca; Huarandosa, Rio Chin-
chipe (Watkins, 1923, AMNH); Amazonas; Rio
Kagka, headwaters (Patton, 1978, MVZ); La
Poza, Rio Santiago (Patton, 1979, 1981, MVZ);
Luya (Patterson, 1987, FMNH); Huampami (Agu-
aruna Village, Rio Cenepa) (Patton, 1977, MVZ);
Tseasim (Aguaruna Village, Rio Huampami), not
precisely located (Patton, 1977, MVZ); Loreto;
Rio Curaray, boca (Olalla, 1925, AMNH); Que-
brada Oran (Chupasko, 1984, LSUMZ); Rio
Napo, lower (Bassler, 1928, AMNH); Santa Rita
(Kalinowski, 1956, FMNH); Iquitos (Bassler,
1927, AMNH); Quistococha (Hershkovitz, 1980,
FMNH); Nauta, left bank Rio Samiria (McGrath,
1980, KU); Santa Elena (Kalinowski, 1956,
FMNH); Yanayaquillo, Rio Samiria (Bluntschli,
1912, AMNH); Rio Samiria (Bluntschli, 1912,
AMNH); San Lorenzo (Kalinowski, 1957,
FMNH); Biological Station "Pithecia" (Hersh-
kovitz, 1980, FMNH); Hudnuco; Tingo Maria
(Heller, 1922, FMNH); Hacienda Exito (Heller,
1922, FMNH); Hacienda Porvenir, 4 mi above
Hacienda Exito (Heller, 1922, FMNH); Hacienda
Buena Vista (Heller, 1922, FMNH); Pasco; Po-
zuzo, Oxapampa (Heller, 1923, FMNH); Nevati
Mission (Buck, 1964, AMNH; Meyers, 1964,
AMNH; Tuttle, 1964, AMNH); San Pablo (Tuttle,
1964, AMNH); Junin; Quinisa, Chanchamayo
(Schunke, 1924, USNM); San Ramon (Anderson,
1914, FMNH); Utcuyacu (Watkins, 1919,
AMNH); Ayacucho; Luisiana [= Hacienda Luisi-
ana] (Gardner, 1971, LSUMZ; Koford, 1963,
AMNH); Ucayali; Yarinacocha (Schunke, 1945,
FMNH); Pucallpa (Hinsch, 1971, 1972, USNM);
Balta (Gardner, 1966, 1968, LSUMZ; O'Neill,
1971, LSUMZ); Lagarto (Olalla, 1928, AMNH);
Santa Rosa, Alto Ucayali (Olalla, 1927, AMNH);
Cusco; Kiteni (Thomas, 1974, LSUMZ); Madre
de Dios; Altamira (Kalinowski, 1962, FMNH);
Puerto Maldonado, Reserva Cuzco Amazonico
(Alegria, MVZ; Hooper, Jr., 1990, KU; Schmidt,
1989, KU; Timm, 1990, KU; Warner, 1983, MVZ;
Woodman, 1989, KU); Tambopata (Hibbs, 1965,
USNM); Puno; Challoma, 3 mi below San Juan
on Rio Tambopata (Heller, 1950, 1951, 1952,
FMNH); VENEZUELA: Zulia; Rio Anaure [ =
Aurare] (Osgood and Jewett, Jr., 1911, FMNH)
Machiques, Kasmera (Kirsch, 1969, KU); Meri
da; Merida (Osgood and Conover, 1920, FMNH
Gabaldon, 1904, AMNH); San Jacinto (Briceno
1903, AMNH; Osgood and Conover, 1920
FMNH); La Vega del Rio Chama (Gabaldon
1907, AMNH); Sierra Nevada de Merida (Osgood
and Conover, 1920, FMNH); Tdchira; La Grita
(Gochfeld, 1974, AMNH); Falcon; La Pastora
(Smithsonian Venezuelan Project, 1967, USNM);
Lara; El Tocuyo (Smithsonian Venezuelan Pro-
ject, 1968, USNM); Trujillo; Motatan (Kirsch,
1969, KU); Trujillo, 15 km E of (Kirsch, 1969,
KU); Apure; Puerto Paez (Smithsonian Venezue-
lan Project, 1966, USNM); Aragua; Ocumare de
la Costa (Handley, Jr., 1960, USNM); Camp Ran-
gel, not located (De La Barera, USNM); Miranda;
Nucleo El Laurel, not located (Kirsch, 1969, KU);
Sucre; Cuchivano (Tate and Clement, 1925,
AMNH; Tate, 1925, AMNH); Latal (Tate, 1925,
AMNH); Neveri (Tate, 1925, AMNH); Monagas;
Cocollar (Tate, 1925, AMNH); Caicara de Matu-
rin [= Caicara] (Smith, Jr., USNM); Bolivar; Cai-
cara (Cherrie, 1907, AMNH); Kanavayen (Mc-
Guire, 1952, AMNH); Chimanta Massif, Lower
Cumbre Camp, Chimanta-tepui, Rio Tiraca (Stey-
ermark and Wurdach, 1955, FMNH); Amazonas;
Cano Leon, Cerro Duida (Olalla, 1928, AMNH);
Chorrera de Vegas (Olalla Brothers, 1929,
AMNH); Middle Camp, Cerro Duida (Tate, 1928,
AMNH); Laterite Valley, Cerro Duida (Olalla,
1929, AMNH); Sabana Grande, Cerro Duida
(Olalla, 1929, AMNH); Cumbre no. 1, Cerro Dui-
da (Olalla, 1929, AMNH); Cumbre no. 15, Cerro
Duida (Olalla, 1929, AMNH); Desfiladero, Cum-
bre no. 13 (Olalla, 1929, AMNH); Esmeralda
(Olalla, 1928, AMNH); Rio Ocamo [= Rio Ocor-
no], boca, Rio Orinoco (Olalla, 1929, AMNH);
San Geronimo, Cerro San Miguel, not located
(Olmstead, 1938, AMNH; Roe, 1938, 1939,
AMNH); Rio Casiquiare (Olalla, 1929, AMNH);
La Fortuna, Cerro San Miguel, not located (Roe,
1938, 1939, AMNH); San Miguel, Palenque, not
located (Roe, 1938, AMNH); TRINIDAD AND
TOBAGO: St. George; Port-of-Spain (AMNH);
San Rafael (Wonder, 1942, FMNH); Carom;
Brickfield (Wonder, 1947, FMNH); St. David;
Grande Riviere, near Tree Station (Downs, 1956,
AMNH); Heights of Oropuche [= El Cerro del
Oropuche] (Carriker, 1910, AMNH); St. Andrew;
Cumaca [= Cumacca] (1967, AMNH); Mt. Harris
(Wonder, 1947, FMNH); Matura (Downs, 1956,
AMNH); Turure Forest (1967, AMNH); Rio San-
gre Grande (Downs, 1956, AMNH); Maingot Es-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
39
tates, 5 mi from Sangre; Nariva; Bush Bush, Na-
riva Swamp (Downs, 1960, AMNH); GUYANA:
Cuyuni-Mazaruni; Oko Mts. (Blake, 1937,
FMNH); Kalacoon [= Kalakun] (Beebe, 1916,
AMNH); Potaro-Siparuni; Tumatumari (Miller,
1913, AMNH); Upper Takutu-Upper Essequibo;
Pakani (Mortensen, 1975, USNM); Dadanawa, 15
mi E (Brock, 1964, USNM); Demerara-Mahaica;
Rio Demerara (Dahlgren, 1922; Warren, 1924,
FMNH); Buxton, 1 mi E (Blake, 1937, FMNH);
SURINAME: Sipaliwini; Tapanahoni, Paloemeu;
Wilhelmina Mountains, West River (Beatty, 1961,
FMNH); Nickerie; Kayser Gebirgte Airstrip
(Beatty, 1960, FMNH); Saramacca; La Poule, left
bank Saramacca River (Hershkovitz, 1962,
FMNH); Para; Lelydorplan [= Lelydorp] (Hersh-
kovitz, 1961, 1962, FMNH); Brokopondo; Loksie
Hattie, Saramacca River (Hershkovitz, 1961,
FMNH); BRAZIL: Manuinhos, not located (Ven-
ancio, 1934, USNM); Roraima; Flexal [= Fre-
chal], Rio Surumu (Tate and Carter, 1927.
AMNH); Serra da Lua, near Boa Vista (Anderson
and Becker, 1913, FMNH); Conceicao (Anderson,
1912, FMNH); Amazonas; Uaupes [= Sao Gabri-
el] (Olalla Brothers, 1928, AMNH; Olalla, 1929,
AMNH); Yucali [= Iucaf] (Olalla Brothers, 1929,
AMNH); Santa Isabel [= Tapurucuara] (Holt,
1930, USNM); Miripinima [= Mirapinima] (Olal-
la Brothers, 1929, AMNH); Parintins [= Villa
Bella Imperatriz = Bella Imperatriz], Boca Rio
Andira (Olalla Brothers, 1930, 1931, AMNH);
Cacau Pirera [= Cacao Pereira] (Olalla Brothers,
1929, 1930, AMNH); Manaus (Becker, 1913,
FMNH); Itacoatiara (Becker, 1915, FMNH); Ro-
sarinho [= Rozarinho] (Olalla Brothers, 1930.
AMNH); Auara Igarape (Olalla Brothers, 1930,
AMNH); Acre; Sena Madureira (1976, USNM);
Amapd; Calcoene (USNM); Serra do Navio
(1966, 1967, 1968, USNM; Belem Virus Labo-
ratory, 1966, USNM; de Pinheiro, 1968, USNM);
Para; Ilha de Marajo (Gilmore, 1936, AMNH:
Kumm, 1936, AMNH); Estrada Ferro de Bragan-
ca (Snethlage, 1911, FMNH); Belem (Belem Vi-
rus Laboratory, 1967, USNM; Pine, 1968,
USNM; Pine, 1973); Villarinho do Monte, Rio
Xingu (Olalla Brothers, 1931, AMNH); Capim
(Belem Virus Laboratory, 1960, AMNH); Porto
de Moz, Rio Xingu (Olalla Brothers, 1931,
AMNH); Ilha do Taiuna. Rio Tocantins (Olalla
Brothers, 1931, AMNH): Igarape Bravo, Rio Ta-
pajos (Olalla Brothers, 1931, AMNH); Faro (Olal-
la Brothers, 1931, AMNH); Sao Jose, near Faro
(Olalla, 1931, AMNH); Cameta, Rio Tocantins
(Olalla Brothers, 1931, AMNH); Igarape Amorin
[= Limontuba], Rio Tapajos (Olalla Brothers,
1931, AMNH); Limoal, near Igarape Amorin, not
located (Olalla Brothers, 1931, AMNH); Moca-
juba, Rio Tocantins (Olalla Brothers, 1931,
AMNH); Piquiatuba, Rio Tapajos (Olalla Broth-
ers, 1931, AMNH); Baiao, Rio Tocantins (Olalla
Brothers, 1931, AMNH); Caxiricatuba, Rio Ta-
pajos (Olalla Brothers, 1931, AMNH); Tauari [ =
Tauary], Rio Tapajos (Olalla Brothers, 1931,
AMNH); Altamira (Emmons, 1986, USNM); For-
dlandia (Gilmore, 1938, AMNH); Itaituba (1976,
USNM; Pinheiro, 1972, USNM); Inajatuba, lower
Rio Tapajos, not located (Olalla Brothers, 1931,
AMNH); Mato Grosso; Aripuana (1974, USNM);
Caceres (Hibbs, 1965, USNM; Kuns, 1965,
USNM); Narinja, not located (AMNH); Maran-
hao; Sao Luis [= Maranhao] (Kaempfer, AMNH);
Distrito Federal; Cruzeiro, near Brasilia (Robert,
1900, AMNH); BOLIVIA: Beni; Rio Beni (An-
derson, 1985, AMNH); Rio Mamore, 5 km NW
of Rio Grande (Anez, 1965, AMNH); Vaca Diez
(Hibbs, 1965, USNM); Estancia Yutcole, 20 km
S of San Joaquin (Anez, 1965, AMNH); Estancia
La Cabana, 1 km E of La Embocada (Creighton.
1979, UMMZ); Santa Cruz; Cerro Amboro, Rio
Pitisama [= Pitasama] (Riddle, 1985, AMNH);
PARAGUAY: Caaguazu; Colonia Sommerfield
no. 11 (Wharton, 1950, USNM); Itapua; El Tirol
(Jones, 1978, MVZ).
Didelphis virginiana Kerr, 1792
Didelphis virginiana Kerr. 1792. In Linnaeus, Animal
Kingdom, p. 193. Synonymy omitted. Gardner,
1993:17 listed.
Type Based on the "Virginian Opossum" of
Pennant, 1781:301.
Type Locality Virginia.
Distribution (not mapped) From British Co-
lumbia to southern Ontario, Canada, into the Unit-
ed States, and introduced into California, south
into Baja California del Norte; Mexico and Cen-
tral America into northern Costa Rica.
Remarks See Gardner, 1973 and 1993; Reig
et al., 1977, for synonymy.
Regarded as extralimital Gardner (1973) list-
ed the following subspecies, Didelphis virginiana
pigra, Bangs, 1898; Didelphis virginiana califor-
nica Bennett, 1833; Didelphis virginiana yucata-
nensis Allen, 1901e.
40
FIELDIANA: ZOOLOGY
Genus Gracilinanus Gardner and Creighton,
1989
Marmosa microtarsus Group: Tate, 1933:8, 10, 34,
43, 47, 49, 185 part, the microtarsus section only.
Creighton, 1984:51, 98 species.
Thylamys: Kirsch and Calaby, 1977:14 subgenus of
Marmosa, part, the species of Tate's microtarsus
section only. Reig et al., 1987:7 part, the species
of Tate's microtarsus section; distribution.
Gracilinanus Gardner and Creighton, 1989:4. Gard-
ner, 1993:17 listed.
Type Species Didelphys microtarsus Wagner,
1842, by original designation.
Distribution (Fig. 16) Forested or wooded
areas of South America from the Caribbean coast
of Colombia and Venezuela in the north to the
Delta del Rio Parana in the south.
Gracilinanus aceramarcae (Tate, 1931)
Marmosa aceramarcae Tate, 1931:12. Tate, 1933:
202 BOLIVIA: holotype only. Goodwin, 1953:
226.
Gracilinanus aceramarcae: Gardner and Creighton,
1989:5. Gardner, 1993:17 listed.
Holotype Young adult female, skin and skull,
AMNH 72568; collected 26 May 1926 by G.H.H.
Tate.
Type Locality Rio "Aceramarca" [= Rio
Aceromarca], Rio Unduavi, Yungas, La Paz, Bo-
livia, 10800 ft (= 3370 m).
Distribution (Fig. 17) Known only from east
side of Lago Titicaca and Bolivia.
Gracilinanus agilis (Burmeister, 1854)
Didelphis murina: Lund (not Linnaeus, 1758), 1839:
233 BRAZIL: Minas Gerais (Rio das Velhas).
Lund, 1841: 51, 52, 106, 107, 134, 237, 265, 293
BRAZIL: Minas Gerais (Lagoa Santa). Lund,
1842:135 BRAZIL: Minas Gerais (Lagoa Santa).
Didelphis pusilla: Lund (not Desmarest, 1804), 1839:
233 BRAZIL: Minas Gerais (Rio das Velhas).
Lund, 1841:106, 134, 237, 265, 293 BRAZIL:
Minas Gerais (Lagoa Santa). Lund, 1842:133,
199 BRAZIL: Minas Gerais (Lagoa Santa).
M[armosa] pusilla: Thomas (not Desmarest, 1804),
1900a:549 part, BRAZIL: Minas Gerais (Lagoa
Santa); agilis Burmeister a synonym.
Grymaeomys pusillus: Winge (not Desmarest, 1804),
1893:27 BRAZIL: Minas Gerais (Lagoa Santa).
Didelphis elegans: Lund (not Waterhouse, 1839),
1840:312, 315 BRAZIL: Minas Gerais; murina
Lund (not Linnaeus), a synonym. Lund, 1842:133,
135, 199 BRAZIL: Minas Gerais (Lagoa Santa).
Marmosa elegans: Miranda Ribeiro (not Waterhouse,
1839), 1914 Annexo, 5:48 BRAZIL: Mato Gros-
so (Salto do Sepotuba).
Didelphys [(Grymaeomys)] agilis Burmeister, 1854:
139- BRAZIL type description.
Grymaeomys agilis: Burmeister, 1856:82 BRAZIL:
Minas Gerais (Lagoa Santa).
[Marmosa] agilis: Trouessart, 1898:1241 classifica-
tion. Krumbiegel, 1941c:7 BOLIVIA: Tarija
(Villa Montes).
Didelphis agilis: Mohr, 1941:226 type history.
Marmosa agilis: Alho et al., 1986:447, 451, 452
BRAZIL: Distrito Federal (Parque Nacional de
Brasilia; Fazenda Agua Limpa). Mares et al., 1986:
294 BRAZIL: Distrito Federal (Fazenda Agua
Limpa, Universidade de Brasilia; Reserva Ecolo-
gica Biologica do Instituto de Geografia e Estatfs-
tica [IBGE]). Redford and da Fonseca, 1986:129
BRAZIL: distribution. Nitikman and Mares, 1987:
83, 84, 86, 87, 90, 91 BRAZIL: Distrito Federal
(Fazenda Agua Limpa, 1000 m). Da Fonseca and
Kierulff, 1988:99 BRAZIL: Minas Gerais (Fa-
zenda Montes Claros; Fazenda Esmeralda; Rio
Doce State Park). Mares et al., 1989:1 1, 12 BRA-
ZIL: Distrito Federal (15-25 km S of Brasilia);
Golds (22 km NE of Cristalina); Mato Grosso
(108-115 kmS of Pocone).
Gracilinanus agilis: Gardner and Creighton, 1989:
5 synonyms (heatrix Thomas, muscula Shamel,
formosa Shamel, chacoensis Tate, buenavistae
Tate, peruana Tate, unduaviensis Tate, blase ri Mi-
randa-Ribeiro, rondoni Miranda-Ribeiro). Mares et
al., 1989:11 BRAZIL: Mato Grosso (Pocone);
Golds (Cristalina); Distrito Federal (Brasilia; Re-
serva Ecologica Biologica do Instituto de Geografia
e Estatistica [IBGE]). Patterson, 1992:7 BOLIV-
IA: Beni (Reyes, Rio Beni). Hershkovitz, 1992: 53-
56 COLOMBIA: Choco (Ungufa); PERU: Ama-
zonas (Bella Vista); BOLIVIA: (Lago Balzon, not
located); Ouro (Cerro Sajama); Beni (San Joaquin);
Santa Cruz (Rio Palometillas; Santiago; Puerto
Suarez); Tarija (Villa Montes; Tablada); BRAZIL:
Mato Grosso (Pocone); Minas Gerais (Brumado;
Lagoa Santa); Maranhdo (Cocos; Alto Rio Parna-
hyba); Golds (Cana Brava; Cristalina); Distrito
Federal (Fazenda Agua Limpa; Brasilia, Fundacao
Zoologica Botanica; Parque Nacional de Brasilia
Brasilia; Reserva Ecologia); Ceard (Ibiapaba; Jua)
URUGUAY: Rio Negro (Bopicua); PARAGUAY:
Concepcion (Concepcion); Guaird (Villa Rica; Ita-
pe Camp); Central (Colonia Nueva Italia); AR-
GENTINA: Buenos Aires (Ingeniero Juarez [= In-
geniero Guillermo-N. Juarez, now in Formosa];
Isla Roble; Zarate; Campana); Formosa (Pozo del
Tigre; Estancia Linda Vista [= Riacho Pilaga]);
Chaco (Las Palmas); Entre Ri'os (Brazo Largo); Mi-
siones (Puerto Libertad-Iguazu, Rio Urugua-i"). Pat-
terson, 1992:1 BOLIVIA: Beni (Reyes). Pacheco
et al., 1993:1 PERU: Madre de Dios (Pakitza).
Anderson et al., 1993:16 BOLIVIA: Santa Cruz
(Buena Vista). Gardner, 1993:17 listed. Mares
and Braun, 2000:32 ARGENTINA: Buenos Aires
(Campana; Isla Roble, Delta de Parana; Reserva
Natural Estricta Otamendi; Zarate, Arroyo Nacu-
rutii); Chaco (Las Palmas; Parque Nacional Cha-
co); Corrientes (near Laguna Gonzalez, Santa
BROWN: ATLAS OF NEW WORLD MARSUPIALS
41
80 70 60 50 40 30 20 10
Fig. 16. Distribution of the genus Gracilinanus: northern coast of Colombia and Venezuela, south to the Delta
del Rib Parana.
42
FIELDIANA: ZOOLOGY
Gracilinanus aceramarcae
500 1000 1500 km
20
80 70 60" W~~ 4G* 30
Fig. 17. Distribution of Gracilinanus aceramarcae: known only from Bolivia. = localities given in published records.
Ana); Entre Rios (Brazo Largo, Delta de Parana);
Formosa (Ingeniero Juarez; Pozo del Tigre; Riacho
Pilaga; Estancia Linda Vista); Misiones (Puerto
Libertad-Iguazu; Rio Urugua-f; Reserva Natural
Estricta San Antonio).
Marmosa agilis agilis: Tate, 1933:194 BRAZIL:
Minas Gerais (Lagoa Santa; Brumado); PARA-
GUAY: Chaco ("V. Clarrica" [probably Villa Rica
in Guaird]); Concepcion (Concepcion); Paraguari
(Sapucay [= SapacafJ); Guaird (Itape). Vieira,
1955:353 (part) BRAZIL: Minas Gerais (Lagoa
Santa; Brumado). Avila Pires, 1968:164 BRA-
ZIL: Goids (Sao Bento). Funayama and Pereira
Barretto, 1971:263 BRAZIL.
Marmosa beatrix Thomas, 19 10c:502 BRAZIL:
Ceard (type locality, Ipu; holotype, female, skin
and skull, BMNH 11.4.23.24; collected 15 June
1910 by Emilia Snethlage). Gardner and Creighton,
1989:5 BRAZIL: Ceard (Ipu). Hershkovitz,
1992:54 BRAZIL: Ceard (Ipu).
[Grymaeomys] beatrix: Matschie, 1916:270 classi-
fication.
Marmosa agilis beatrix: Tate, 1933:196 BRAZIL:
Ceard (Ipu; Jua, near Iguatu; Serra Ibiapaba).
Marmosa marica: Thomas (not Thomas, 1898),
1927b:608 PERU: Hudnuco (Tingo Maria, 606
m; male and female of which the male [Coll. Hen-
dee, no. 1063] became holotype of Marmosa agilis
peruana Tate, 1 93 1 : 1 1 ).
Marmosa musculo Shamel, 1930a:83 ARGENTI-
NA: Formosa (type locality, Riacho Pilaga [= Es-
tancia Linda Vista], 10 mi NW of km 182; holo-
type, male, skin and skull, USNM 236330; col-
lected 9 August 1920 by Alexander Wetmore;
name preoccupied by Didelphis (Marmosa) mus-
culo Cabanis, 1848 [= Marmosa murina Linnae-
us]).
Marmosa formosa Shamel, 1930b:311 replacement
name for Marmosa musculo Shamel, preoccupied.
Tate, 1933:232 type only.
Marmosa velutina formosa: Cabrera, 1958:33. Olrog,
1959:409 ARGENTINA: Formosa (Ingeniero
Juarez [= Ingeniero Guillermo N. Juarez]).
Marmosa agilis buenavistae Tate, 1931:10 BOLIV-
IA: Santa Cruz (type locality, Buena Vista; holo-
type, female, skin and skull, BMNH 26.12.4.91,
collected 7 March 1916 by J. Steinbach). Tate,
1933:197 BOLIVIA: Lago Balzon, not located;
Santa Cruz (Buena Vista; Rio Palometillas; Puerto
Suarez, Sara Province). Avila Pires, 1968:167
BRAZIL: Mato Grosso (Salto do Sepotuba). Gard-
ner and Creighton, 1989:5 BOLIVIA: Santa Cruz
(Buena Vista). Hershkovitz, 1992:53 BOLIVIA:
Santa Cruz (Buena Vista). Anderson, 1997:147
BROWN: ATLAS OF NEW WORLD MARSUPIALS
43
BOLIVIA: Beni (Pampa de Meio: Rio Baures.
mouth; San Joaquin: Puerto Caballo: km 35 NW
Yucumo); Cochabamba (Comarapa. 25 km W);
Pando (Independencia): Santa Cruz (Estancia Ca-
chuela Esperanza: Montero, 3 km SE. 1 km N of
Villa Copacabana: Buena Vista: Rio Palomotitlas.
Buenavista: Las Lomitas: Ingeniero Moro. 7 km E.
3 km N; Zanja Honda, 3 km N of Hacienda El
Peh'cano; Santiago de Chiquitas: Santiago: Puerto
Suarez).
Marmosa agilis chacoensis Tate, 1931:10 PARA-
GUAY: Paraguari (type locality. Sapucay [= Sap-
ucaf]: holotype. male, skin and skull. BMNH
4.1.5.48. collected 11 September 1903 by W. Fos-
ter). ARGENTINA: Chaco (Las Palmas): BOLIV-
IA: Tarija (Villa Montes. incorrectly cited as from
Argentina). Tate. 1933:196 PARAGUAY: Para-
guari (Sapucay [= Sapucaf]): Guaird (Itape Camp.
near Sapucay [= Sapucaf]): ARGENTINA: Chaco
(Las Palmas). Massoia and Fornes. 1972:517 AR-
GENTINA: Buenos Aires; Formosa: Chaco; Mi-
siones; Entre Rios. Wainberg et al.. 1979:33 AR-
GENTINA: Entre Rios (Delta del Parana). Gonza-
lez. 1985:1 URUGUAY: Rio Negro (Rio Uru-
guay, near Fray Bentos-Puerto Ungue: Bopicua.
Rio Uruguay: Arroyo Bopicua). Gardner and
Creighton. 1989:5 PARAGUAY: Paraguari (Sap-
ucay" [= Sapucaf]). Hershkovitz, 1992:54 PAR-
AGUAY: Paraguari (Sapucay [= Sapucaf]). Flores
et al. 2000:322 ARGENTINA: Jujuy (Ingenio La
Esperanza; Santa Barbara): Salta (Ingenio San
Martin de Tabacal. Lote "Milagros"); PARA-
GUAY: Paraguari (Supacay [= Sapucaf]). Ander-
son, 1997:148 BOLIVIA: Chuquisaca (Rio Li-
mon); Tarija (Villa Montes, upper Rio Pilcomayo:
Tablada).
Marmosa agilis peruana Tate. 1931:11 PERU:
Hudnuco (type locality. Tingo Maria, 606 m; ho-
lotype "young adult" male, skin and skull, BMNH
27.11.1.268: collected 20 January 1927 by R. W.
Hendee). Tate. 1933:198 PERU: Hudnuco (Tingo
Maria. Rio Huallaga); Amazonas (Bellavista. Rio
Marafion valley, 303 m, "probably but not certainly
distinct"). Gardner and Creighton. 1989:5 PERU:
Hudnuco (Tingo Maria). Hershkovitz. 1992:53
PERU: Hudnuco (Tingo Maria).
Marmosa agilis peruania: Tate, 1933:194 incorrect
subsequent spelling.
Marmosa agilia peruania: Cabrera. 1958:28 incor-
rect subsequent spelling.
Marmosa unduaviensis Tate. 1931:1 1 BOLIVIA: La
Paz (type locality, Pitiguaya [= La Florida], Rio
Unduavi, 5800 ft'[= 1800 m]: holotype male, skin
and skull. AMNH 72563: collected 12 May 1926
by G.H.H. Tate). Tate. 1933:201 (skull) BOLIV-
IA: La Paz (Pitiguaya [= La Florida]). Gardner and
Creighton. 1989:5 BOLIVIA: La Paz (Pitiguaya
[= La Florida]. Rio Unduavi. Yungas). Hershko-
vitz. 1992:53 BOLIVIA: La Paz (Pitiguaya [= La
Florida]).
Marmosa agilis unduaviensis; Anderson. 1997:148
BOLIVIA: Beni (Reyes): La Paz (Pitiguaya [= La
Florida]): Oruro (Mount Sajama).
Marmosa blaseri Miranda Ribeiro, 1936:361, 373
BRAZIL: Goids (type locality. Sao Bento; holotype
by monotypy. skin and skull. MNRJ 1250. pur-
chased 1929 from Sr. Jose Blaser). Moojen in Mi-
randa Ribeiro, 1955:393 listed. Vieira, 1955:
352 BRAZIL: Goids (Cana Brava, Nova Roma).
Avila Pires. 1968:164 erroneous attribution of in-
valid lectotype designation to Moojen in Miranda
Ribeiro. 1955:393. Gardner and Creighton. 1985:
5 BRAZIL: Goids (Sao Bento).
Thylamys rondoni Miranda Ribeiro. 1936:385. 387
BRAZIL: Mato Grosso (type localities, Sal to de
Sepotuba and Sao Joao de Serra do Norte: types in
alcohol, skulls out, MNRJ).
Marmosa rondoni: Vieira. 1955:352 listed.
Holotype Young adult, sex undetermined,
skin mounted, Zoologische Museum, Halle, no.
147, collected by H. Burmeister.
Type Locality Rio das Velhas, Lagoa Santa.
Minas Gerais, BRAZIL. 760 m.
Distribution (Fig. 18) From the Colombian-
Panamanian border into Peru. Bolivia. Brazil, Par-
aguay. Uruguay, northern Argentina, and the Rio
Parana basin of Argentina.
Gracilinanus dryas (Thomas, 1898)
Marmosa drxas Thomas. 1898a:456. Tate. 1933:
203 VENEZUELA: Merida (Selva Culata. 4000
m: Montanas Uchisera. 3000 m; Montes de la Si-
erra [= Sierra Montanas]). Handley. 1976:6 VE-
NEZUELA: Merida (Tabay); Tdchira (Buena Vis-
ta); Trujillo (Hacienda Misisf). Handley and Gor-
don. 1979:67 COLOMBIA: Cundinamarca (Bo-
queron de San Francisco. Bogota): VENEZUELA:
Merida (Merida: La Montana; La Mucuy; Tabay;
Culata; Montes de la Serra [= Sierra Montanas];
Uchisera. Cordillera de Merida); Tdchira (Buena
Vista): Trujillo (Hacienda Misisf). Aagaard. 1982:
1 VENEZUELA: Merida (Parque Nacional de Si-
erra Nevada). Perez-Hernandez. 1989:369 VE-
NEZUELA: Tdchira (Betania. Distrito Junfn); Cer-
ro Las Copas. La Revancha; Buena Vista. 41 km
SW of San Cristobal: Trujillo (13-15 km E of Tru-
jillo): altitudinal range 2210-4000 m.
[Grymaeomys] dryas: Matschie. 1916:270 listed.
Gracilinanus drxas: Gardner and Creighton. 1989:
6 VENEZUELA: Merida (La Culata). Hershko-
vitz. 1992:52 VENEZUELA: Merida (Uchisera.
Cordillera de Merida: La Culata; Tabay: La Mucuy;
Cafetales de Merida: Cafetal de Milla; Montes de
la Serra [= Sierra Montanas]); Trujillo (Trujillo:
Misisf): Tdchira (Buena Vista: Betania). Gardner.
1993:17 listed.
Holotype Old adult male, skin and skull.
BMNH 98.5.15.2; collected 14 December 1896
by Salamon Bricefio.
Type Locality Culata. Sierra de Merida, Me-
rida. Venezuela. 4000 m.
Distribution (Fig. 19) Northern Cordillera
44
FIELDIANA: ZOOLOGY
30 20
Fig. 18. Distribution of Gracilinanus agilis: south from Colombia-Panama border to the Rio Parana basin of
Argentina. = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
45
Fig. 19. Distribution of Gracilinanus dryas: from Colombia to northwestern Venezuela. = localities given in
published records.
Oriental of Colombia north into the Sierra de Me-
rida of northwestern Venezuela.
Gracilinanus emiliae (Thomas, 1909)
[l]Didelphxs pusilla: Thomas (part, not Desmarest.
1804), "l888b:349 part. BRAZIL: Para (Santa-
rem).
Marmosa emiliae Thomas, 1909:379. Tate. 1933:
189 holotype only. Avila Pires, 1964:8. 11
BRAZIL: Amazonas (km 50. Manaus-Itacoatiara
RR). Husson, 1978:23 SURINAME: Marowijne
(Langamankondre).
[Marmosa] Emiliae: Cabrera. 1913:14 classifica-
tion.
[Grymaeomys] emiliae: Matschie, 1916:270 listed.
Gracilinanus emiliae: Gardner and Creighton. 1989:
6. Hershkovitz. 1992:53-54 BRAZIL: Amazonas
(Igarape Grande. Rio Jurua. W bank; Codajas; km
50, Manaus-Itacoatiara RR); Ceard (Crato); Para
(Para [ = Belem]; Santarem). Patterson, 1992:8
BRAZIL: Amazonas (Codajas; Igarape Grande).
Gardner. 1993:17 listed. Voss et al.. 2001:25
BRAZIL: Paid (Belem: Capim); COLOMBIA:
Meta (Los Micos): FRENCH GUIANA (Paracou);
GUYANA: Upper Takeiii-Upper Essequibo (Dad-
anawa); SURINAME: Marowijne (Langamankon-
dre); VENEZUELA: Monagas (Maturin. 47 km
SE).
Marmosa agricolai Moojen. 1943:2 BRAZIL:
Ceard (type locality. Crato; holotype. male, skin
and skull. MNRJ no. 1495: collected 26 August
1936 by A. Leitao de Carvalho). Voss et al., 2001:
29 possibly a valid species, status of name as a
synonym uncertain.
Marmosa microtarsus microtarsus Tate (part, not
Wagner), 1933:192 BRAZIL: Pard (Santarem);
comment on BMNH specimen recorded as Didel-
phys pusilla by Thomas.
Gracilinanus longicaudus Hershkovitz. 1992:38
COLOMBIA: Meta (San Juan de Arama). CO-
LOMBIA: Meta (type locality. Los Micos, San
Juan de Arama; holotype, adult male, skin and
skull, FMNH 87924: collected 29 April 1957 by
Kjell von Sneidern).
Marmosa lepida: Hershkovitz (not Thomas, 1888).
1992:54 BRAZIL: Amazonas (Codajas. Igarape
Grande). Patterson, 1992:8 BRAZIL: Amazonas
(Codajas. Rio Solimoes; Igarape Grande, upper Rio
Jurua).
Holotype Subadult male, skin and skull.
BMNH 9.3.9.10; collected 13 February 1909 by
Emilia Snethlage.
46
FIELDIANA: ZOOLOGY
Fig. 20. Distribution of Gracilinanus emiliae: Colombia, Venezuela, the Guianas, and eastern Brazil. = localities
given in published records.
Type Locality "Para (Belem)," BRAZIL.
Distribution (Fig. 20) Colombia, northeast-
ern Venezuela, the Guianas, and northern Brazil.
Gracilinanus marica (Thomas, 1898)
M[armosa]. pusilla: Thomas (not Desmarest, 1804),
1896:314 VENEZUELA: Merida (Sierra de Me-
rida).
Mannosa pusilla: Elliot (not Desmarest, 1804), 1907:
4 VENEZUELA.
Marmosa marica Thomas, 1898a:455. Elliot, 1907:
4 VENEZUELA. Tate, 1933:200 VENEZUE-
LA: Merida (Rfo Albarregas; Cafetal de Milla; Lla-
no de Merida; Cafetal de Merida [= Cafetales de
Merida]; La Pedregosa); Sucre-Monagas (Cerro
Turumiquire). Handley, 1976:7 VENEZUELA:
Distrito Federal (Los Venados; Pico Avila); Falcon
(Urama); Monagas (Hato Mata de Bejuco). Hand-
ley and Gordon, 1979:68 (part) COLOMBIA:
Cundinamarca (La Selva); La Guajira (Marimonda
[= Las Marimondas]); VENEZUELA: Distrito
Federal (Los Venados: Pico Avila); Falcon (Ura-
ma); Sucre-Monagas (Cerro Turumiquire); Mona-
gas (Hato Mata de Bejuco); Merida (Cafetal de
Chama; Cafetal de Merida [= Cafetales de Meri-
da]; Llano de Merida; Cafetal de Milla; Merida;
Rio Albarregas; La Pedregosa). Perez-Hernandez,
1989:369 VENEZUELA: Distrito Federal (EI
Junquito; Carretera El Junquito; Carretera Caracas,
La Colonia Tovar, km 40; Sabaneta; Carretera El
Limon; El Encantado, Hacienda El Limon; Suapo,
Hacienda El Limon, La Conchita, Hacienda El Li-
mon; Zona Quacatal, Hacienda El Limon; Aragua
(Estacion Biologica de Rancho Grande, Rancho
Grande; La Quinta, Rancho Grande; Palo de Vaca,
Rancho Grande; Pica Periquito, El Portachuelo,
Rancho Grande; Don Ramon, Rancho Grande; Pan-
toja, Carretera Maracay Ocumare; Guacamaya,
Rancho Grande; Guamitas, Rancho Grande).
[Grymaeomys] marica: Matschie, 1916:270 VE-
NEZUELA: Merida (Rio Albarragas).
Gracilinanus marica: Gardner and Creighton, 1989:
6 VENEZUELA: Merida (Rfo Albarregas).
Hershkovitz, 1992:51-52 COLOMBIA: Cundi-
namarca (La Selva); La Guajira (Marimonda [=
Las Marimondas]); VENEZUELA: Merida (Llano
de Merida; Pedregosa; Cafetos de Merida [= Caf-
etales de Merida]; Merida; Rfo Albarregas; Cafetal
de Milla); Falcon (Urama); Aragua (Rancho
Grande [= Estacion Biologica de Rancho Grande]);
Distrito Federal (Los Venados; Pico Avila; Haci-
enda El Limon; El Junquito); Sucre-Monagas (Cer-
ro Turumiquire); Monagas (Hato Mata de Bejuco).
Gardner, 1993:17 listed.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
47
Fig. 2 1 . Distribution of Gracilinanus marica: Colombia and northern Venezuela. = localities given in published
records.
Holotype Young adult male, skin and skull.
BMNH 98.5.15.1; collected 24 January 1897 by
Salamon Briceno.
Type Locality Rio Albarregas. Merida. Ve-
nezuela, 1630 m.
Distribution (Fig. 21) Northern Venezuela
and northern Colombia.
Gracilinanus microtarsus (Wagner, 1842)
Didelphys microtarsus Wagner. 1842:359. Wagner.
1843:43. Wagner. 1850:147 BRAZIL: Sao Paulo
(type locality. Ipanema [= Ypanema]). Pelzeln.
1883:1 14 part. BRAZIL: Sao Paulo (Ipanema [ =
Ypanema]).
Grymaeomxs microtarsus: Winge. 1893:24 BRA-
ZIL.
[Didelphys (Marmosa)} microtarsus: Trouessart.
1898:1239 as a synonym of Marmosa murina.
Marmosa microtarsus: Thomas. 1900a:549 BRA-
ZIL: Parana (Palmeira). Miranda Ribeiro, 1935:
38 BRAZIL.: Rio de Janeiro (Teresopolis). Davis.
1947:3 BRAZIL: Rio de Janeiro (Teresopolis).
Wainberg et al.. 1979:33 BRAZIL. Fonseca and
Kierulff. 1988:108. 116 BRAZIL: Minas Gerais
(Parque Estadual do Rio Doce). Stallings. 1988:
159 BRAZIL: Minas Gerais (Parque Estadual do
Rio Doce). De Azevedo et al., 1982:93 BRAZIL:
Santa Catarina (Seminario Cora^ao de Jesus. Co-
rupa: Blumenau).
Gracilinanus microtarsus: Gardner and Creighton.
1989:6. Gardner and Creighton. 1989:4 BRAZIL:
Sao Paulo (Ipanema [= Ypanema]). Hershkovitz.
1992:54-55 BRAZIL: Rio de Janeiro (Porto
Real; Teresopolis: Rio de Janeiro; Ilha Grande):
Sao Paulo (Ituverava: Franca; Jabuticabal; Piraci-
caba; Perus; Ipanema [= Bacaetava = Ypanema];
Sao Sebastiao; Ilha da Vitoria: Primeiro Morro:
Boa Vista. Rio Ipiranga); Parana (Palmeira): Santa
Catarina (Joinville [= Join vile]; Humboldt; Rio
Itapocu: Blumenau: Ararangua): Rio Grande do Sul
(Taquara: Sao Lourenco). Gardner. 1993:17 list-
ed.
Marmosa microtarsus microtarsus: Tate, 1933:190
BRAZIL: Sao Paulo (Sao Paulo: Ipanema [= Ypa-
nema]); Rio de Janeiro (Porto Real: Rio de Janei-
ro): Parana (Palmeira): Santa Catarina (Joinville
[= Joinvile], near Rio Itapocu: Blumenau: Arar-
angua). Vieira. 1949(1950):352 BRAZIL: Sao
Paulo (Piracicaba: Perus: Ipanema; Ituverava); Rio
de Janeiro (Ilha Grande). Vieira. 1955:350 BRA-
ZIL: Sao Paulo (Ituverava. Piracicaba; Perus. Ipi-
ranga); Rio de Janeiro (Ilha Grande; Teresopolis);
48
FIELDIANA: ZOOLOGY
Parana (Palmeira); Santa Catarina (Ararangua;
Blumenau; Joinville [= Joinvile]). Avila Pires,
1968:165 BRAZIL: Santa Catarina (Humboldt).
Funayama and Pereira Barretto, 1971:263 BRA-
ZIL: Sao Paulo (Franca).
Grymaeomys agilis: Hensel (not Burmeister, 1854),
1872:123 BRAZIL: Rio Grande do Sul (Island in
Rio Guahyba [= Una Guaiba], near Porto Alegre).
Marmosa agilis: Davis (not Burmeister), 1945:119
BRAZIL: Rio de Janeiro (Fazenda Boa Fe, Tere-
sopolis). Da Fonseca and Kierulff, 1988:108, 112,
128, 137 BRAZIL: Minas Gerais (Parque Estad-
ual do Rio Doce).
Marmosa agilis agilis: Vieira (part, not Burmeister),
1949(1950):351 BRAZIL: Sao Paulo (Jabutica-
bal; Ilha Victoria [= Ilha da Vitoria]). Vieira, 1955:
353 BRAZIL: Sao Paulo (Jabuticabal; Ilha Vic-
toria [= Ilha da Vitoria]; Ilha de Sao Sebastiao).
Didelphvs pusilla: Thomas (part, not Desmarest,
1804), 1888b:348 part, not synonymy; BRAZIL:
Rio de Janeiro; Santa Catarina; Rio Grande do Sul
(Taquara; Sao Lourenco).
M[armosa] pusilla: Thomas (not Desmarest, 1804),
1900a:549 BRAZIL: Parana (Palmeira).
[^Marmosa pusilla Shufelt (not Desmarest, 1804),
1926:563, 574 BRAZIL: Rio Grande do Sul (Sao
Lourenco).
Micoureus pusillus: Goeldi (not Desmarest, 1804),
1894:462 incorrect subsequent spelling BRA-
ZIL: Rio de Janeiro (Serra dos Orgaos). Ihering,
1894:1 1 BRAZIL: Sao Paulo.
[?\Marmosa herhardti Miranda Ribeiro, 1936:361,
382 BRAZIL: Santa Catarina (type locality,
Humboldt; holotype, skin [skull?] in MNRJ, pur-
chased from W. Erhardt). Avila Pires, 1968:165
BRAZIL: Santa Catarina (type locality Humboldt;
holotype, adult female, skin and skull [MNRJ no.
1226] collected 23 December 1915). Gardner and
Creighton, 1989:6 BRAZIL: Santa Catarina
(Humboldt).
Marmosa microtarsus guahybae Tate, 1931:10
BRAZIL: Rio Grande do Sul (type locality, Ilha de
Guahyba [= Ilha Guaiba], near Porto Alegre, ZMB
4306; collected by Reinhold Hensel). Tate, 1933:
192 BRAZIL: Rio Grande do Sul (Ilha Guahyba
[= Ilha Guaiba]; Taquara; Sao Lourenco). Gardner
and Creighton, 1989:6 BRAZIL: Rio Grande do
Sul (Isla Guahyba [= Ilha Guaiba]). Avila Pires,
1994:369 BRAZIL: Rio Grande do Sul (Ilha do
Guahyba [= Ilha Guaiba]).
Type Series Eight specimens in Vienna Mu-
seum; collected January, March, June, and Octo-
ber between 1819 and 1821 by Johann Natterer
(cf. Pelzeln, 1883, p. 114). Tate (1933, table 1, p.
191) designated the adult male "Vienna 48A" as
"type," i.e., lectotype.
Type Locality Ypanema [= Ipanema], be-
tween Sao Paulo and Sorocaba, Sao Paulo, Brazil.
Distribution (Fig. 22) Atlantic coastal forests
from Minas Gerais to Rio Grande do Sul in south-
eastern Brazil.
Gracilinanus perijae Hershkovitz, 1992
Marmosa marica: Handley and Gordon (part, not
Thomas, 1898), 1979:68 COLOMBIA: La Gua-
jira (Marimonda [= Las Marimondas], Serrania de
Perija).
Gracilinanus perijae Hershkovitz, 1992:41 CO-
LOMBIA: La Guajira (Las Marimondas).
Holotype Adult male, skin and skull, USNM
no. 280881; collected 10 March 1943 by Philip
Hershkovitz, original no. 1826.
Type Locality Las Marimondas, the site of a
former coffee plantation, western slope Serrania
de Perija, Cordillera Oriental de los Andes, north
of and above Fonseca at the time in the Depart-
ment of Magdalena, now incorporated into the
Department of La Guajira, 1450 m, Colombia.
Distribution (Fig. 23) Known only from type
locality on western slope Serrania de Perija, La
Guajira, Colombia.
Genus Hyladelphys Voss, 2001
Hyladelphys kalinowskii (Hershkovitz, 1992)
Gracilinanus kalinowskii Hershkovitz, 1992:37
PERU: Junin (Chanchamayo); Cusco (Hacienda
Cadena).
Hyladelphys kalinowskii: Voss et al., 2001:32
FRENCH GUIANA (Paracou); GUYANA: East
Berbice-Corentyne (New River Falls); Peru: Cusco
(Hacienda Cadena); PERU: Junin (Chanchamayo);
Loreto (Nuevo San Juan on Rio Galvez).
Holotype Adult female, skin and skull,
FMNH 89991; collected 3 July 1958 by Celestino
Kalinowski; original no. 457.
Type Locality Hacienda Cadena, Marcapata,
Cusco, Peru, 890 m.
Distribution (Fig. 24) Known only from for-
ested slopes of southeastern Peru in the Depart-
ment of Junin into that of Cusco, eastern Guyana
and northern French Guiana.
Genus Lestodelphys Tate, 1934
Notodelphys Thomas, 192 lb: 137 type species, No-
todelphys halli Thomas, 1921b, by original desig-
nation; generic name preoccupied by Notodelphys
Allman, 1847, a genus of copepod, and Notodel-
phys Lichtenstein and Weinland, 1854, a batrachi-
an. Gardner, 1993:17 listed.
Lestodelphys Tate, 1934:154 replacement for Noto-
delphys Thomas, preoccupied. Gardner, 1993:17
listed.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
49
40
Fig. 22. Distribution of Gracilinanus microtarsus: coastal forests of southeastern Brazil. = localities given in
published records.
Notodelphis Cabrera, 1958:11 incorrect subsequent
spelling of Notodelphys Thomas, 1921b.
Lestodelphis Cabrera, 1958:11 incorrect subsequent
spelling of Lestodelphys Tate, 1937.
Type Species Notodelphys halli Thomas,
1921b.
Distribution Western and southern Argenti-
na.
Lestodelphys halli (Thomas, 1921)
Notodelphys halli Thomas, 192 lb: 137. Thomas,
1929:45 ARGENTINA: Chubut (Pico Salaman-
ca, 100); Santa Cruz (type locality, Estancia Mach-
ijada [= Estancia La Machijada], near Puerto De-
seado). Reig. 1959:57 ARGENTINA: Chubut
(Estancia Los Manantiales, Languineo; Valle de
Rio Chico; Sarmiento. Crespo, 1974:5 ARGEN-
TINA: Chubut (Pico Salamanca). Gardner, 1993:
17 listed.
Lestodelphys halli: Tate. 1934:154 type species.
Reig, 1959:58 ARGENTINA: Chubut (Estancia
Los Manantiales, Languineo). Birney et al., 1996:
150 ARGENTINA: Chubut (Pampa de Agnia, 30
km NW). Mares and Braun, 2000:33 ARGEN-
TINA: Chubut (Estancia Los Manantiales near
Languineo; Pampa de Agnia, 30 km NW; Pico Sal-
amanca); La Pampa (Lihue Calel; Parque Nacional
Lihue Calel); Mendoza (Chacras de Coria); Neu-
quen (Estancia Tehuel Malal); Rio Negro (Cerro
Leones; Clemente Onelli; Comallo, 8-10 km
WSW; Los Menucos, 15 km SE); Santa Cruz
(Cabo Tres Puntas; Estancia La Madrugada; Meseta
El Pedrero).
Holotype Male, skin and skull, BMNH
21.6.7.19; collected by T. H. Hall.
Type Locality Originally stated to be Cabo
Tres Puntas, SE Patagonia, 42S, Santa Cruz, but
later (Thomas, 1929:45) emended to Estancia La
Madrugada, not far from Puerto Deseado, Santa
Cruz, Argentina.
Distribution (Fig. 25) Argentina from Men-
doza south to Santa Cruz.
Genus Lutreolina Thomas, 1910
Sarigua Muirhead, 1819:429 part, S. crassicaudata
listed among included species.
50
FIELDIANA: ZOOLOGY
Gracilinanus perijae
1500 km
30
Fig. 23. Distribution of Gracilinanus perijae: known only from type locality in northeastern Colombia. =
localities given in published records.
Lutreolina Thomas, 1910a:247 included species:
crassicaudata Desmarest. Cabrera, 1919:44 -char-
acters; synonymy. Thomas, 1923b:583 review.
Miranda Ribeiro, 1936:396 characters. Krumbie-
gel, 194 lb: 189 taxonomic review. Reig, 1958:
261 comparisons. Ximenez, 1967:1 review.
Gardner, 1993:18 listed, synonymy.
Peramys: Matschie (not Lesson), 1916:259, 266,
269 subgenus of Didelphis Linnaeus; included
species crassicaudata Desmarest ("Typus") and
turneri Gunther.
Type Species by Monotypy Didelphis cras-
sicaudata Desmarest.
Distribution Southern Brazil, eastern Boliv-
ia, Uruguay, Paraguay, northern Argentina, Guy-
ana, Venezuela, and eastern Colombia.
Remarks The disjunct distribution of Lutreo-
lina in Colombia, Venezuela, and northern Guy-
ana, and Bolivia, southeastern Brazil, Paraguay,
Uruguay, and northern Argentina, is unique.
Lutreolina crassicaudata (Desmarest, 1804)
Micoure troisieme ou micoure a queue grosse Azara,
1801:284.
Coligrueso Azara, 1802:229.
Didelphis crassicaudata Desmarest, 1804:19. Ren-
nger, 1830:226 PARAGUAY. Waterhouse, 1839:
94 URUGUAY: Maldonado (Maldonado). Gard-
ner, 1993:18 listed.
Sarigua crassicaudata: Muirhead, 1819:429 new
combination.
D[idelphis]. Crassicaudata: Fischer, 1829:267.
Didelphys crassicaudata: Waterhouse, 1841:100
PARAGUAY; URUGUAY; ARGENTINA: Buenos
Aires. Waterhouse, 1 846:497 URUGUAY; BRA-
ZIL mustelina E. Geoffroy a synonym. Thomas,
1888b:334 GUYANA: West PomeroonSupe-
naam Coast (Better Hope); turneri Gunther, in syn-
onymy; BRAZIL: Rio Grande do Sul (San Louren-
50 do Sul [= Sao Lourenco do Sul]); URUGUAY:
Maldonado (Maldonado); ARGENTINA: Buenos
Aires (La Plata; Belgrano; Mariano Moreno).
Winge, 1893:7 BRAZIL: Minus Gerais (Lagoa
Santa, Recent and Pleistocene). Figueira, 1894:8
URUGUAY. Aplin, 1894:315 URUGUAY.
D[idelphvs] crassicaudata: Cabanis and Schomburgk,
1848:777 GUYANA.
Peramys crassicaudata: Lesson, 1842:187 classifi-
cation. Matschie, 1916:269 classification.
[Cuica] crassicaudata: Liais, 1872:330 BRAZIL.
Lutreolina crassicaudata: Thomas, 1910a:247 AR-
GENTINA: Buenos Aires (Los Ingleses). Thomas,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
51
Hyladelphys kalinowski
500 1000 150:k~:
Fig. 24. Distribution of Hyladelphys kalinowskii: Peru. Guyana, and French Guiana.
published records.
30*
localities given in
191 7a: 100 ARGENTINA: Buenos Aires (Isla
Ella, Parana delta). Thomas. 1923b:583 ARGEN-
TINA: Cordoba (Noetinger). Devincenzi. 1935:
88 distribution. BertonC 1939:6 PARAGUAY.
Krumbiegel. 194 lb: 190 ARGENTINA: Formosa
(San Jose); PARAGUAY: Caaguazii (Cerro Pila-
do): BRAZIL: Mato Grosso do Sul (Rio Ivinhei-
ma). Kuhlhorn, 1955:163 BRAZIL: Mato Gros-
so. Reig. 1964:207 ARGENTINA: Buenos Aires
(Arroyo Corrientes; Arroyo Brusquitas. General
Pueyrredon; Mar del Sur). Avila Pires. 1968:168
BRAZIL: Sao Paulo (Guariba). Ayala et al., 1973:
374 COLOMBIA: Arauca (Puerto Gaitan). Ol-
rog. 1976:6 ARGENTINA: Jujuy (Abra de Ca-
nos: Cerro [= Sierra] Santa Barbara): Tucumdn
(Raco). Handley. 1976:8 VENEZUELA: Bolivar
(Hato San Jose). Reig et al.. 1977:212 ARGEN-
TINA: Buenos Aires (Banado de San Jorge, Parque
Pereyra lraola). Avila Pires. 1977:7 BRAZIL: Rio
de Janeiro (Pico das Agulhas Negras. Maromba).
Marshall. 1978:2 ARGENTINA: Buenos Aires
(Chapadamal Formation, not located: Monte Her-
moso Formation, not located: Rio Lujan); Cata-
marca (Corral Quemado Formation). Olros. 1979:
10 ARGENTINA: Jujuy (Calilegua Mountain [ =
Serrania de Calilegua]). Mares et al.. 1981:166
ARGENTINA: Salta (Agua Blanca: Oran). Crespo.
1982:1 ARGENTINA: Entre Rios (Parque Na-
cional El Palmar). Cajal. 1981:27 ARGENTINA:
Tucumdn (Aguas Chiquitas). De Azevedo et al.,
1982:94 BRAZIL: Santa Catarina (Armacao do
Sul: Botuvera). Lemke, 1982:226 COLOMBIA:
Meta (Villavicencio): Vichada (Cano Avispa). Se-
luja. 1984:702 URUGUAY: Rocha (Pinamar. not
located): Canelones (Banado de Carrasco: Las Pie-
dras); Rocha (La Tuna): Rio Negro (Rio Negro):
Maldonado (Piriapolis). Crespo Diaz et al.. 1986:
472 COLOMBIA: Meta (Sabanas de Vichada y
Meta. not located). Avila Pires. 1987:123 BRA-
ZIL: Rio Grande do Sul (Sao Lourenco [= Sao
Lourenco do Sul]). Boker Bentti. 1988:111 VE-
NEZUELA: Bolivar (Hato San Jose: San Ignacio
de Yuruanf): Anzoategui (Paso Bajito); COLOM-
BIA: Meta (Villavicencio); Vichada (Cano Avispa;
Comisaria del Vichada): Arauca (Puerto Gaitan);
ARGENTINA: Salta (Agua Blanca). Perez-Her-
nandez. 1989:374 VENEZUELA. Anzoategui
(Paso Bajito); Bolivar (San Ignacio de Yuruanf):
Monagas (Maturin): GUYANA: West Pomeroon-
Supinaam Coast (Better Hope) (the name Lutreo-
Hna crassicaudata turneri used). Heinonen and
Bosso. 1994:55 ARGENTINA: Jujuy (Parque
Nacional Calilegua. Caimancito). Mares et al..
1996:105 ARGENTINA: Tucumdn (Aconquija:
El Cadillal. Aguas Chiquitas. Sierras de Medina;
Piedras Coloradas; Quebrada de Lules; Raco; Cerro
52
FIELDIANA: ZOOLOGY
20
30
40
50
! \
^
h(\
Lestodelphys halli
500
1000
1500 km
90
80 u
70
60 D
50"
40
30
20"
Fig. 25. Distribution of Lestodelphys halli: confined to western and southern Argentina.
published records.
10
localities given in
[= Sierra] San Javier; San Miguel de Tucuman, 25
km NW; San Pablo; El Indio, Ruta 307. Capllonch
et al., 1997:53 ARGENTINA: Tucuman (Parque
Biological Sierra de San Javier). Morando and Po-
lop, 1997:31 ARGENTINA: Cordova. Mares and
Braun, 2000:33 ARGENTINA: Buenos Aires
(Arroyo Brusquitas; Arroyo Corrientes; Arroyo La
Matanza, not located; Bafiado de San Jorge, Parque
Pereyra Iraola, delta de Parana; Los Yngleses; Mar
del Sur; Miramar; Reserva Natural Estricta Ota-
mendi); Chaco (Parque Nacional Chaco); Corri-
entes (Centra Argentino de Primates, SE Corrien-
tes, not located; Estancia Ita Caabo, Mercedes;
Parque Nacional Mburucuya); Formosa (El Colo-
rado; Laguna Blanca; Parque Nacional Rio Pilco-
mayo); Jujuy (Abra de Canas; Cerro Calilegua;
Cerro Santa Barbara; Parque Nacional Calilegua);
Misiones (Parque Nacional Iguazu); Salta (Aguas
Blancas, 24 km NW; Parque Nacional El Rey; Alto
Rio Bermejo); Santa Fe (Malabrigo); Tucuman
(Aconquija; Sierras de Medina; Aguas Chiquitas;
El Cadillal; Piedras Coloradas; Quebrada de Lules;
Raco, Sierra San Javier; San Miguel de Tucuman,
25 km NW; San Pablo; Aguas Chiquitas; Cerro San
Javier; El Indio, Ruta 307; Horco Molle; Raco; Tafi
Viejo).
L[utreolina] c\rassicaudata\. Thomas, 1923b:583,
584 PARAGUAY: Central (Asuncion); Presiden-
te Hayes (Chaco).
[Lutreolina] crassicaudata: Krumbiegel, 1941b:
190 PARAGUAY: San Pedro (Estancia La Ger-
mania); Central (Rib Pilcomayo, boca); Caaguazii
(Cerro Pilado); ARGENTINA: Santa Fe (Galvez);
BRAZIL: Mato Grosso do Sul (Rio Ivinheima);
BOLIVIA: Santa Cruz (San Jose).
Micoureus crassicaudatus: Gervais, 1855:287.
Didelphys crassicaudatus: Burmeister, 1856:88
mustelinus Geoffroy manuscript name in synony-
my. Burmeister, 1879:190 distribution.
[Didelphis (Metachirus)] crassicaudatus: Hensel,
1872:121 BRAZIL: Rio Grande do Sul. Ihering,
1892:100 BRAZIL: Rio Grande do Sul. Bertoni,
1923:51 PARAGUAY: Central (Bahia de Asun-
cion). Krieg, 1924:637 ARGENTINA: Santa Fe.
Didelphis crassicaudis: Illiger, 1815, 1804-1811:
107 incorrect subsequent spelling of crassicau-
data.
Lutreolina crassicaudus: Hildebrand, 1961:244-
249 incorrect subsequent spelling. Olfers, 1818:
206.
Lutreolina crassicaudata crassicaudata: Miranda Ri-
beiro, 1936:400 PARAGUAY: Presidente Hayes
(Chaco [= Gran Chaco]). Vieira, 1949:355 BRA-
ZIL: Sao Paulo; travassosi Miranda Ribeiro in syn-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
53
onymy. Vieira. 1955:348 BRAZIL: Sao Paulo
(Ituverava; Franca; Ribeirao Preto; Ribeirao Boni-
to; Bauni); Rio de Janeiro (Angra dos Reis). Kan-
tis, 1963:54 ARGENTINA: Chaco (Rio de Oro).
Anderson, 1997:149 BOLIVIA: Beni (San Joa-
quin: San Pedro, 10 km W; San Javier, 23 km W);
tarija (Cuyambuyo).
Didelphis macroura Desmoulins, 1824:492 name
based on the micoure a queue grosse of Azara.
Didelphis mustelina Waterhouse. 1846:497-499 no-
men nudum; name attributed to E. Geoffroy, re-
portedly based on specimen in the "Paris Muse-
um"; cited as a synonym of Didelphis crassicau-
data.
Didelphxs Turned Gunther. 1879:108. Gardner, 1993:
18 listed.
[Peramys] turneri: Matschie, 1916:269.
L[utreolina] c[rassicaudata] turneri: Thomas, 1923b:
583, 584 GUYANA: Demerara (type locality.
Better Hope [now in West Pomeroon-Supenaam
Coast]: holotype. male, skin and skull. BMNH
79.5.1.3; collected by W. Y. Turner).
D[idelphis] Ferruginea Larrahaga, 1923:346 name
based on the coligrueso of Azara. Gardner, 1993:
18 listed.
L[utreolina] c[rassicaudata] paranalis Thomas, 1923b:
584 ARGENTINA: Santa Fe (type locality, Las
Rosas; holotype, female, skin and skull. BMNH
17.5.2.22; collected 6 November 1916 by Robin
Kemp); Corrientes (Goya): Buenos Aires (Isla Ella.
Parana Delta); URUGUAY: Montevideo (Colon;
Montevideo). Ximenez. 1967:2 URUGUAY: Ca-
nelones (Banada Tropa Vieja; Carrasco; Arroyo Las
Brujas); Rocha (Camino del Indio); Artigas (San
Gregorio); Montevideo (Cerro; Parque Lecoq):
BRAZIL: Rio Grande do Sul (San Lorenzo [= Sao
Lourenco do Sul]); Sao Paulo (Ribeirao Preto; Ri-
beirao Bonito: Pirassununga; Franca); Mato Grosso
(Maracajii [now in Mato Grosso do Sul]): ARGEN-
TINA: Buenos Aires (Quiroga; Gonzalez Catan; Ar-
royo Pereyra; Mar del Sur, Miramar; Rio Pasaje Ta-
lavera; Zdrate; Punta Lara; Hurlingham); lutrilla a
synonym. Gardner, 1993:18 listed.
Lutreolina crassicaudata paranalis: Sanborn. 1929:
151 URUGUAY: Colonia (Colonia Suiza). De-
vincenzi. 1935:90 URUGUAY. Yepes. 1945:60
ARGENTINA: Santa Fe (Malabrigo). Cabrera.
1958:40. Massoia and Fornes. 1964:29 ARGEN-
TINA: Buenos Aires (delta del Parana). Fornes and
Massoia, 1965:102 ARGENTINA: Buenos Aires
(Miramar).
[Lutreolina crassicaudata] paranalis: Krumbiegel.
1941b: 192 ARGENTINA: Santa Fe (Galvez).
L[utreolina] c[rassicaudata] lutrilla Thomas, 1923b:
585 BRAZIL: Rio Grande do Sul (type locality,
Sao Lourenco do Sul; holotype, male, skin and
skull, BMNH 85.6.26.26; collected by H. von Iher-
ing); URUGUAY: Maldonado (Maldonado). San-
born. 1929:151 URUGUAY: Lavalleja (Arroyo
Polanco, Minas). Gardner, 1993:18 listed.
Lutreolina crassicaudata lutrilla: Cabrera, 1958:39.
Avila Pires, 1994:370 BRAZIL: Rio Grande do
Sul (Sao Lourenco do Sul).
L[utreolina] c[rassicaudatus] bonaria Thomas,
1923b:585 ARGENTINA: Buenos Aires (type lo-
cality, Los Ingleses, Ajo; holotype, male, skin and
skull, BMNH 20.2.7.44; collected 20 July 1919 by
H. E. Box); range from Belgrano to Cape San An-
tonio.
Lutreolina crassicaudata bonaria: Miranda Ribeiro,
1936:401.
Lutreolina c[rassicuadata] travossosi Miranda Ribei-
ro, 1936:402 BRAZIL: Sao Paulo (Guariba, E of
Sao Paulo); (holotype, skin and skull, MNRJ, pre-
sented by Dr. Lauro Travassos). Gardner, 1993:
18 listed.
Holotype None in existence; name based
solely on the micoure a queue grosse of Azara.
Type Locality Paraguay; restricted to Asun-
cion by Cabrera (1958, Rev. Mus. Argentino
Cienc. Nat. "Bernardino Rivadavia," 4:39).
Distribution (Fig. 26) Two populations in
South America: one in eastern Colombia, Vene-
zuela, and western Guyana, the other in southern
Brazil from Rio de Janeiro south into Rio Grande
do Sul, west into southern Mato Grosso through
Uruguay, Paraguay, eastern Bolivia, and northern
Argentina.
Additional Records VENEZUELA: Boli-
var; La Paragua, 20 km W (Smithsonian Expe-
dition, 1967, USNM); PARAGUAY: Central;
Luque (Myers, 1976, UMMZ); Colonia Nueva
Italia (Willem, 1944, FMNH); Presidente Hayes;
Rio Aguaray-Guazu (Creighton, 1979, UMMZ);
Caaguazii; Estancia San Ignacio, 24 km NNW of
Carayao (Dobson, 1979, UMMZ); Conception;
Concepcion (Creighton, 1979, UMMZ); Canen-
diyu; Curuguaty (Creighton, 1978, UMMZ; Dob-
son, 1979, UMMZ; Myers, 1976, UMMZ); La
Cordillera; Tobati (Creighton, 1979, UMMZ;
Myers, 1978, UMMZ; Voss, 1978, UMMZ); Par-
aguari; Parque Nacional Ybycuf (Myers, 1979,
UMMZ); Neembucu; Pilar (Creighton, 1979,
UMMZ); Misiones; Ayolas (Creighton, 1979,
UMMZ); URUGUAY: Lavalleja; Polanco, Minas
(Sanborn, 1926, FMNH); ARGENTINA: Chaco;
Palm Swamp, 12 km ESE of Resistencia (Sage,
1982, MVZ); Santa Fe; Santa Fe (Nachman,
1989. UMMZ); Buenos Aires; Buenos Aires Zoo-
logical Gardens (Onelli, 1909, USNM); Urdam-
pilleta [= La Torrecita] (Osgood, 1923, FMNH);
Corrientes; Corrientes, left bank Rio Parana
(Thorington, 1978, USNM); Entre Rios; Las Cue-
vas (Nachman, 1989, UMMZ); Misiones; Santa
Teresa, 30 km W of San Ignacio (Kuns, 1965,
USNM).
Genus Marmosa Gray, 1821
Opossum Schmid, 1818:115 included species O.
marsupialis, O. virginianus, O. murinus.
54
FIELDIANA: ZOOLOGY
Fig. 26. Distribution of disjunct populations of Lutreolina crassicaudata in South America: one population in
Colombia, Venezuela, and Guyana, the other from Bolivia south into Argentina and east into Paraguay, Uruguay, and
southern Brazil, o = localities from unpublished records; - localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
55
110
100
Fig. 27. Distribution of the genus Marmoset in Central America: Mexico south to Panama. See Figure 28 for
distribution in South America.
Sahgua, Muirhead. 1819:429 part. S. murina listed
among included species.
Marmosa Gray, 1821:308. Thomas. 1895a: 190 no-
menclature; Micoureus Lesson and Asagis Gloger
listed as synonyms. Matschie, 1916:260 taxono-
my. Tate, 1933:20 taxonomic revision. Gardner,
1993:18 listed.
Grymaeomys Burmeister, 1854:viii, 138 subgenus of
Didelphys; included species, murina, agilis, pusil-
la, tristriata [= Monodelphis americana Midler] ,
brachyura [ Monodelphis brevicaudata Erxle-
ben], velutina. Burmeister, 1856:77 taxonomy.
Thomas, 1888b:340 type species Didelphis mu-
rina Linnaeus by subsequent designation. Mat-
schie, 1916:260 designation of pusilla Desmarest
as type species (p. 270) invalid. Gardner, 1993:
18 listed.
Cuica Liais, 1872:329 included species, myosuros
[= Metachirus myosuros Temminck], murina, pus-
illa, cineria, lanigera [= Philander lanata Olfers],
crassicaudata [= Lutreolina crassicaudata Des-
marest], tricolor [= Monodelphis touan Shaw],
tristriata [= Monodelphis americana Midler].
Hershkovitz, 1949:12 type species. Cuica murina
Liais [= Didelphis murina Linnaeus]. Gardner,
1993:18 listed.
Quica Cabrera, 1958:12 Incorrect subsequent spell-
ing of Cuica Liais, 1 872, in synonymy of Marmosa
Gray.
Asagis Gloger, 1842:82 type species Didelphis mu-
rina Linnaeus by subsequent designation (Thomas,
1888b:340). Gardner, 1993:18 listed.
Notagogus Gloger, 1842:82 type species Didelphis
murina Linnaeus, by subsequent designation
(Thomas, 1888b:340). Gardner, 1993:18 listed.
Stegomarmosa Pine. 1972:279 type species Mar-
mosa (Stegomarmosa) andersoni Pine. Gardner,
1993:18 listed.
Type Species Didelphis marina Gray, 1821
incorrect subsequent spelling of Didelphis murina
Linnaeus, 1758.
Distribution (Figs. 27 and 28) Neotropical re-
gion from northern Mexico to Bolivia and Brazil.
Marmosa andersoni Pine, 1972
Marmosa andersoni Pine, 1972:279 PERU: Cusco
(Hacienda Villa Carmen, Cosnipata). Gardner,
1983:18 listed, synonymy.
Holotype Adult male, skin and skull, FMNH
84252, collected 21 September 1954 by C. Kali-
nowski.
Type Locality Hacienda Villa Carmen, Cos-
nipata, Cusco, Peru, 600 m.
Distribution (Fig. 29) Known only from type
locality.
Comments Type species of Stegomarmosa.
56
FIELDIANA: ZOOLOGY
90
80 70 60 50 40 30 20 10
Fig. 28. Distribution of the genus Marmosa in South America: throughout northern half of continent, south
through Bolivia and Brazil. See Figure 27 for distribution in Central America.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
57
Fig. 29. Distribution of Marmosa andersoni: known only from type locality in Department of Cusco. Peru.
localities given in published records.
Marmosa canescens (J. A. Allen, 1893)
Mannosa canescens canescens J. A. Allen. 1893:
235 MEXICO: Oaxaca (type locality. Santo Do-
mingo de Guzman. Isthmus of Tehuantepec: holo-
type^ adult male. AMNH 3111/2433: collected 30
April by A. C. Buller). Tate. 1933:141 MEXICO:
Nayarit (Tepic): Oaxaca (Santo Domingo de Guz-
man: Puerto Angel: Tehuantepec: Tlapacingo):
Guerrero (Tlapa. Rfo Xochihuehuellen: Acapulco:
Sochi, not located): Puebla (Amolac): Michoacdn
(Los Reyes): Colima (Hacienda Magdalena): Nay-
arit (Tepic): Yucatan (Yaxcaba). Hall and Villa.
1950:164 MEXICO: Michoacdn (Los Reyes).
Hershkovitz. 1951:551 MEXICO: Yucatan (Chi-
chen Itza: Yaxcaba). Hooper. 1955:7 MEXICO:
Sinaloa (San Bias): Puebla (Tehuacan). Gardner.
1993:18 listed. Ramirez-Pulido and Castro-Cam-
pillo. 1994:18 MEXICO: Morelos: Oaxaca;
Guerrero.
Marmosa canescens oaxacae Merriam. 1897:43
MEXICO: Oaxaca (type locality. Oaxaca: holo-
type. adult male. USNM 68240: collected 14 Au-
gust 1894 by E. W. Nelson and E. A. Goldman).
Tate. 1933:143 MEXICO: Oaxaca (Oaxaca).
Gardner. 1993:18 listed.
Marmosa canescens sinaloae J. A. Allen. 1898:143
MEXICO: Nayarit (type locality. Tatemales: holo-
type. adult male. BMNH 98.3.2.161. collected 14
May 1897 by P. O. Simons). Tate, 1933:142-143
MEXICO: Sinaloa (Tatamales. not located: Culia-
can: Mazatlan: Esquinapa: Rosario: Rancho Palo
Amarillo. Amatlan. not located): Durango (Ventan-
as, not located). Gardner. 1993:18 listed.
Marmosa canescens insularis Merriam. 1898:14
MEXICO: Sinaloa (type locality. Maria Madre Is-
land [= Isla Marfa Madre]. Tres Marias Islands [ =
Isla Tres Marias]: holotype. young adult male.
United States Biological Survey 89215. collected
16 May 1897). Tate/l933: 144 MEXICO: Sinaloa
(Isla Maria Madre: Islas Tres Marias). Ramirez-
Pulido and Castro-Campillo. 1994:18 MEXICO:
Nayarit. Gardner. 1993:18 listed.
Marmosa canescens gaumeri Osgood. 1913a: 175
MEXICO: Yucatan (type locality. Yaxcaba. Chi-
chen Itza: holotype. FMNH 19995. collected No-
vember 1912 by G. F. Gaumer). Gardner. 1993:
18 listed.
Holotype Adult male. AMNH 3111/2433,
collected 30 April 1890 by A. C. Buller.
Type Locality Santo Domingo de Guzman.
Isthmus of Tehuantepec. Oaxaca. Mexico.
Distribution (Fig. 30) In Mexico from southern
Sonora to Oaxaca. Yucatan, and Islas Tres Marias.
58
FIELDIANA: ZOOLOGY
110 100 90 80
Fig. 30. Distribution of Marmosa canescens: Mexico. = localities given in published records.
Marmosa lepida (Thomas, 1888)
Didelphys pusilla: Tomes (not Desmarest, 1804),
1858:548 ECUADOR: Marona-Santiago (Guala-
quiza).
Didelphys (micoureus) lepida Thomas, 1888a: 158
PERU: Thomas, 1888b:347 PERU: Loreto (Santa
Cruz, Rfo Huallaga); BOLIVIA. Gardner, 1983:
18 listed.
[Marmosa] lepida: Trouessart, 1898:1240 classifi-
cation.
M[armosa] lepida: Thomas, 1927d:374 BOLIVIA:
Santa Cruz (Buenavista).
Marmosa lepida: Cabrera, 1958:18 classification;
grandis Tate, a synonym. Husson, 1978:16-17
SURINAME: Paramaribo (Paramaribo); Brokopon-
do (Gansee); Marowijne (Wiawia Bank); Sipaliwini
(Nassau Mountains). Anderson et al., 1993:16 BO-
LIVIA: Santa Cruz (Buenavista). Anderson, 1997:
150 BOLIVIA: Santa Cruz (Buenavista).
Marmosa lepida lepida: Tate, 1933:205 PERU: Lor-
eto (Santa Cruz, Rio Huallaga); Ucayali (Lagarto,
Rio Ucayali); Hudnuco (Puerto Marquez, Rio
Pachitea); BOLIVIA: Santa Cruz (Buenavista);
ECUADOR: Pastaza (Sarayacu, Rio Bobonaza);
SURINAME: ("San Alouatta" [sic], not located);
Paramaribo (Paramaribo). Lemke et al., 1982:
226 COLOMBIA: Meta (Cano Entrada, Sierra de
La Macarena); Amazonas (Rio Amacayacu [= Rio
Amaca-Yacu]).
Marmosa lepida grandis Tate, 1931:12 BOLIVIA:
Santa Cruz (type locality, Buenavista; holotype, fe-
male, skin and skull, BMNH 26.12.4.94, collected
25 January 1926 by J. Steinbach). Tate, 1933:
207 BOLIVIA: Santa Cruz (Buenavista). Gard-
ner, 1993:18 listed.
Holotype Female, skin and skull, BMNH
69.3.31.4; collected by E. Bartlett.
Type Locality Santa Cruz, near Yurimaguas,
lower Rio Huallaga, Loreto, Peru.
Distribution (Fig. 31) Colombia, Ecuador,
Peru, Suriname, and Santa Cruz in Bolivia.
Marmosa mexicana Merriam, 1897
Marmosa murina mexicana Merriam, 1897:44
MEXICO: Oaxaca (Juquila). Gardner, 1993:18
listed. Ramfrez-Pulido and Castro-Campillo, 1944:
18 MEXICO: Oaxaca.
Marmosa mexicana mexicana: Tate, 1933:133
MEXICO: Chiapas (Chicharras; Huehuetan); Oa-
xaca (Juquila); Veracruz-Llave (Tuxpan; Jalapa;
Achotal; Texolo; San Andres Tuxtla; Mirador; Pasa
Nueva); GUATEMALA: Baja Verapaz (San Jeron-
imo, E of Salama; Chilasco, on tributary of Rfo
Palochic); Aha Verapaz (Cajabon; Coban; San
Cristobal); HONDURAS: (Rfo Umulla, not locat-
ed); NICARAGUA: Matagalpa; (Matagalpa); Ri-
vets (San Emilio, Lago Nicaragua; Lago Nicara-
gua); Nueva Segovia (Ocotal); Managua (Mana-
gua); Zelaya (Rfo Coco); Boaco (Bijagua); Jinote-
ga (San Rafael del Norte); Esteli (Chinandega);
BROWN: ATLAS OF NEW WORLD MARSUPIALS
59
Fig. 31. Distribution of Marmosa lepida: central Colombia south to Bolivia: also in Suriname. = localities
given in published records.
Zela\a (Corn Islands [= Islas del Maiz]). Goodwin.
1954:3 MEXICO: Tamaulipas (Aserradero del
Infernillo). Ramfrez-Pulido and Castro-Campillo.
1994:18 MEXICO: Chiapas; Oaxaca.
Marmosa mexicana savannarum Goldman. 1917:
108 PANAMA: Chiriqui (type locality. Boquer-
6n, holotype, subadult male. AMNH 18915. col-
lected 31 October 1901 by J. H. Batty). Goldman.
1917:108 PANAMA: Chiriqui (Boqueron). Tate.
1933:134 PANAMA: Chiriqui (Bogava [= Bug-
aba]; Colon and Code. W of. not located). Bole.
1937:145 PANAMA: Veraguas (Mariato). Gard-
ner. 1993:18 listed.
Marmosa mexicana zeledoni Goldman. 1917:238
COSTA RICA: Cartago (type locality. Navarro:
holotype. young adult male, USNM 12885-37133,
collected 1 March 1878 by Juan Cooper for Jose
C. Zeledon). Tate. 1933:136 COSTA RICA: Car-
tago (Navarro); Puntarenas (Boruca); Limon
(Jimenez, Rio Jimenez; Rio Pacuare); NICARA-
GUA: Zelaya (Rio Tuma. E of Matagalpa; Blue-
fields). Goodwin. 1946:287 COSTA RICA: Pun-
tarenas (Boruca; Agua Buena): Limon (Rio Pacu-
are). Gardner, 1993:18 listed.
Marmosa mexicana mayensis Osgood. 1913a: 176
MEXICO: Yucatan (type locality, Izamal; holotype,
young adult male. FMNH 19994, collected March
1913 by G. F. Gaumer). Tate, 1933:135 MEXI-
CO: Yucatan (Chichen Itza: Izamal). Ramfrez-Pul-
ido and Castro-Campillo, 1994:18 MEXICO: Yu-
catan. Gardner. 1993:18 listed.
Holotype Young male, USNM 71526, col-
lected 28 February 1895 by E. W. Nelson and E.
A. Goldman.
Type Locality Juquila, Oaxaca, Mexico, at
1500 m.
Distribution (Fig. 32) Tamaulipas, Mexico,
to W Panama in Central America.
Marmosa murina (Linnaeus, 1758)
Een soort van een Bos-rat, Merian. 1719:66.
Mus sylvestris americanus Seba, 1734:48, 49, 50
(marmosa of the Brasilians).
Glis sylvestris Americanus Seba. 1735:90.
Marmose Buffon. 1763:335.
Merian opossum Pennant. 1771:210.
Murine opossum Pennant, 1793:23 GUIANA.
[Didelphis] murina Linnaeus. 1758:55 name based
on Seba (1734. figs. 1. 2. 3. 6 [not "12. 36" as
60
FIELDIANA: ZOOLOGY
Fig. 32. Distribution of Marmosa mexicana: from Tmaulipas, Mexico, south through western Panama.
localities given in published records.
given by Linnaeus and authors]. Gardner, 1993:
18 listed.
Did[elphis] murina: Cuvier, 1798:125 la marmose.
Didelphis murina: Latreille, 1803:304 marmose.
Desmarest, 1816:427 FRENCH GUIANA: (Cay-
enne). Temminck, 1825:50 FRENCH GUIANA.
Jentink, 1887:302 SURINAME. Waterhouse,
1841:105 "GUIANA." Waterhouse, 1846:508
part, "GUIANA." Thomas, 1887:153 GUYANA:
Demerara-Mahaica (Pen Hope). Jentink, 1888:
221 part, SURINAME; part, PERU: Loreto (Rio
Javarf [= Rio Yavarf]). Thomas, 1888b:343 part,
SURINAME: Lidth de Jeude collection in the Brit-
ish Museum; GUYANA: Demerara-Mahaica
(Plantation Hope); BRAZIL: Para; part, synonyms,
dorsigera Linnaeus, guianensis Kerr, musculo Ca-
banis. Thomas, 1892:314 identification of Mus
sylvestris americanus Seba with specimens of the
Lidth de Jeude collection in the British Museum.
Didelphis Murina: Shaw, 1800:484 murine opos-
sum.
Didelphis marina: E. Geoffroy, 1803:143 incorrect
subsequent spelling South America.
Sarigua murina: Muirhead, 1819:429 new combi-
nation.
[l]D[idelphys] murina: Wied-Neuwied (not Linnaeus,
1758), 1*826:41 1 BRAZIL: Espirito Santo-Ba-
hia-Minas Gerais (Rio Mucuri).
D[idelphys] murina: Tschudi, 1844:14, 146, 151
PERU.
['l]Didelphys murina: Burmeister, 1854:138 charac-
ters based primarily on descriptions by Wied-Neu-
wied, 1826.
Grymaeomys murina: Burmeister, 1856:80 muscu-
lus Cabanis, a synonym.
[Cuica] murina: Liais, 1872:330 classification.
Marmosa murina: Thomas, 1895:58 classification.
J. A. Allen, 1904d:337 VENEZUELA: Bolivar
(Rio Suapure). Thomas, 191 lb: 144 SURINAME
type locality; "cotypes" in BMNH. Tate, 1933
106 PERU: Cusco (Marcapata). Pine et al., 1970
668 BRAZIL: Mato Grosso (Xavantina [= Chav-
antina]). Handley, 1976:7 VENEZUELA: Zulia
(El Rosario); Yaracuy (Urama); Amazonas (San
Juan; Belen; Tamatama; Capibara; Boca Mavaca);
Monagas (San Agustfn); Bolivar (Los Patos; Rib
Supamo; Hato San Jose; El Manaco; km 125, 85
km SSE of El Dorado; Icabaru). Reig et al., 1977:
211 PERU: Ucayali (Balta); VENEZUELA: Bo-
livar (Santa Elena de Uairen). Avila Pires and Gou-
vea, 1977:9 BRAZIL: Rio de Janeiro (Campo
Bello, Itatiaya [= Pico das Agulhas Negras]). Hus-
son, 1978:19-20 SURINAME: Nickerie (Vier
Gebroeders Mountain); Sipaliwini (Bitagron;
Adampada Kreek; Raleigh Falls); Paramaribo
(Paramaribo; Cultuurtuin; Morgenstond); Com-
mewijne ("Clevia"; Peperpot); Para (Zanderij);
Saramacca ("Jarikaba"); Wanica (Lelydorp); Ma-
rowijne (Moengo). Eisenberg et al., 1979:191
VENEZUELA: Miranda-Guarico (Parque Nacion-
al Guatopo). Kock, 1986:269 SURINAME. Pat-
terson, 1992:7 BOLIVIA: Beni (Puerto Salinas);
BROWN: ATLAS OF NEW WORLD MARSUPIALS
61
BRAZIL: Amazonas (Itacoatiara; Codajas; Joao
Pessoa; Igarape Grande); Para (Casa Nova: Iro^an-
ga; Itapoama). Rageot and Albuja. 1994:176 EC-
UADOR: Pastaza (Mera). Anderson. 1997:151
BOLIVIA: Beni (Puerto Salinas; Centinela. La Em-
bocada); Cochabamba (Villa Tunari); La Paz (Moi-
re [= Moira Camp, alto Rio Madidi]; Chijchipani):
Santa Cruz (Las Cruces). Patton et al., 2000:51
BRAZIL: Amazonas (Altamira, right bank Rio
Jurua; Barro Vermelho, left bank Rio Jurua: Ilha
Paxiuba. right bank Rio Jurua). Voss et al.. 2001:
41 FRENCH GUIANA (Arataye; Cayenne: Par-
acou: Les Nouragues; Piste Saint Elie; Saul): GUY-
ANA: Demerara-Mahica (Demerara River. 29 mi
above Georgetown): Cuyuni-Mazaruni (Kartabo):
West Demerara-Essiquibo Islands ("Caraiben-
Niederlassung Arrai am obern Pomeroon") [holo-
type of musculo]; SURINAME: Brokopondo (Fin-
isanti); Para (Lelydorpplan: Zanderij); Sipaliwini
(Oelemarie; Raleigh Falls): VENEZUELA: Ama-
zonas (Mt. [= Cerro] Duida); Bolivar (Ciudad Bo-
livar: Arabupii).
[Marmosa] murina: Trouessart. 1898:1239 part.
classification.
M[armosa] murina: Miranda Ribeiro. 1905:189
BRAZIL: Rio de Janeiro (Monte Serrat. Itatiaya).
Marmosa murina murina: Tate. 1933:92 BRAZIL:
Para (Igarape Assu [= Igarape Acu], [holotype of
parata]; Caldeirao, Marajo [= Ilha de Marajo]):
Maranhdo (Parnaiba [= Parnahyba, now in Piauf];
Cocos, Codo: Tury-assu); Pernambuco (Dois Ir-
maos, Recife); Paraiba (Penha): GUYANA: Esse-
quibo (Hoolie Island. Essequibo River, not locat-
ed): SURINAME. Vieira, 1953:220 BRAZIL: Al-
agoas (Mangabeiras [= Mangabeira]). Avila Pires.
1958:3 BRAZIL: Para (Utinga. Belem de Para).
Carvalho, 1962:285, 296 BRAZIL: Amapd (Rio
Maruanum. Macapa: Rio Amapari. Macapa). Avila
Pires, 1968:166 BRAZIL: Rio de Janeiro (Monte
Serrat [= Pico das Agulhas Negras], Itatiaia). Pine.
1973b:55 BRAZIL: Para (Belem). da Fonseca
and Redford. 1984:518 BRAZIL: Distrito Fed-
eral (Reserva Ecologica Biologica do Instituto de
Geografia e Estatfstica [IBGE]).
Marmosa murina murina: Cabrera (part, not Linnae-
us). 1958:20 moeirae in synonymy only.
P[hilander] murinus: Tiedemann. 1808:427 classi-
fication.
0[possum] murinus: Schmid. 1818:117 marmose of
Buffon.
Philander murinus: Gray, 1843:101 tropical Amer-
ica.
Micoureus murinus: Lesson, 1842:186 classifica-
tion.
[Didelphis] dorsigera Linnaeus, 1758:55 "Ameri-
ca" (type locality): name based on the mus sylves-
tris americanus of Seba, "Merian Surin[ame]"
Bos-rat. Gardner, 1993:18 listed.
Didelphis Dorsigera: Shaw. 1 800:485 Merian opos-
sum.
Didelphis dorsigera: Temminck, 1825:48 compari-
son with D. caxopollin. Schomburgk. 1840:345
GUYANA. Jentink, 1887:302 SURINAME. Jen-
tink, 1888:221 SURINAME.
Didelphys dorsigera: Waterhouse, 1841:104 Meri-
an 's opossum. Burmeister. 1879:192 part, distri-
bution.
D[idelphxs] dorsigera: Wagner. 1843:48. Cabanis and
Schomburgk, 1848:777 GUYANA.
Micoureus dorsigera: Lesson, 1842:186 classifica-
tion.
Philander dorsigera: Gray, 1843:101 specimens
without locality listed.
Marmosa dorsigera: Thomas, 191 lb: 144 SURI-
NAME: (type locality); synonyms of murina Lin-
naeus; "type" in BMNH (Lidth de Jeude collec-
tion).
[Monodelphis] dorsigens: Burnett, 1830:351 incor-
rect subsequent spelling classification.
Grymaeomys dorsiger: Burmeister, 1856:80 incor-
rect subsequent spelling part, distribution.
Didelphis guianensis Kerr. 1792:194 SURINAME:
name based on the Guiana opossum of Pennant.
Gardner. 1993:18 listed.
Marmosa murina guianensis, Carvalho, 1962:285,
296 BRAZIL: Amapd (Macapa).
Didelphys macrotarsus Wagner. 1842:359 BRAZIL:
(type locality, Rio Madeira, Amazonas; holotype,
male, in the Vienna Museum, no. 195, collected by
Johann Natterer: not Didelphys macrotarsos Schre-
ber. 1778 [= Tarsius sxrichta Linnaeus]). Wagner,
1850:145 BRAZIL: Amazonas (type locality, Rio
Madeira, not located). Pelzeln. 1883:114 BRA-
ZIL: Amazonas (Rio Madeira, not located). Gard-
ner, 1993:18 listed.
Thylamys macrotarsus: Miranda Ribeiro. 1936:386
characters of type specimen.
[Marmosa] macrotarsus: Trouessart. 1898:1239 list-
ed in synonymy of Marmosa murina Linnaeus.
D[idelph\s]. musculus Cabanis, in Schomburgk,
1848:778 GUYANA: Essequibo (type locality,
Caribe colony of Arrai, upper Pomeroon River
[now West Demerara Essequibo Islands]; holo-
type. male, skin and skull, collected by Richard
Schomburgk). Matschie. 1916:264, 272 holotype
in Berlin museum, skin no. 2331, skull no. 24797.
Gardner. 1993:18 listed.
[Marmosa] musculo: Trouessart, 1898:1239 listed in
synonymy of Marmosa murina Linnaeus.
Marmosa murina musculo: Tate, 1933:96 GUYA-
NA: Pomeroon-Supenaam (Supenaam River, trib-
utary of lower Essequibo); Demerara-Mahaica
(Dunoon: Pen Hope, 120 [sic = 20] mi E of
Georgetown; Hyama Creek. Demerara River; Hyde
Park. Demerara River); Essequibo Islands-West
Demerara (Clarkesdale, Demerara River); Upper
Demarara-Berbice (Comackka, 80 mi up the De-
merara River); Cuyuni-Mazaruni (Kartabo, Esse-
quibo River: Bartica Grove. Essequibo River); Es-
sequibo Islands-West Demerara (Bonasica, Esse-
quibo River); Upper Takutu-Upper Essequibo
(Moon Mountains; Kanuku Mountains, N of); SU-
RINAME: Marowijne (Maroni); Brokopondo (Ber-
gendal): FRENCH GUIANA: (Cayenne); holotype,
a juvenile male; chloe Thomas, a synonym.
Grymaeomxs impavida: Burmeister (? not Tschudi,
1844), 1856:81 BRAZIL: precise locality un-
known; macrotarsus Wagner and murina Burmeis-
ter 1854 regarded as synonyms.
Didelphys Materhousii Tomes. 1860:58 part. Thom-
as, 1 92 la:522 ECUADOR: Morona-Santiago
62
FIELDIANA: ZOOLOGY
(type locality, Gualaquiza; holotype skin lost, skull
only in BMNH 7.1.1.215, collected by Fraser; an-
imal pouched female in spirits figured by
Tomes, 1860, pi. 76, as type, is a member of Mar-
moset cinerea group originally figured by Water-
house, 1846:505). Gardner, 1993:18 listed.
D[idelphys] waterhousei: Thomas, 1888b:342 in
synonymy of Didelphys cinerea Temminck.
[Marmosa] Waterhousei: Trouessart, 1898:1238
listed in synonymy of Marmosa cinerea.
[^Marmosa waterhousei: Lonnberg, 1921:69 EC-
UADOR: Napo (below Baeza, 5000 ft [= 1560
m]).
M[armosa] Waterhousei: Thomas, 1927a:374 mar-
anii Thomas, a synonym.
Marmosa murina waterhousei: Tate, 1933:103 part,
ECUADOR: Tungurahua (Mirador); Morona-San-
tiago (Gualaquiza, Rio Zamora); Zamora-Chinchi-
pe (Zamora); Pastaza (Rio Curaray, mouth, not lo-
cated); bombascarae Anthony, a synonym. Han-
dley and Gordon, 1979:70 VENEZUELA: Zulia
(El Rosario).
[Caluromys] waterhousei: Matschie, 1916:269 list-
ed.
Remarks The type of waterhousei was first
recorded by Tomes in 1859 with a note from the
collector, Mr. Fraser, that it had "five young in
her pouch, each 3 inches long." When naming the
animal in 1860, Tomes (p. 58) described the un-
derparts as "brownish-buff, with a stripe of yel-
lowish-white along the centre of throat and
breast." On page 59 (op. cit.), Tomes repeats that
"on all under parts the hairs are unicolour, of a
pale buff, palest on the mesial line, and on the
throat and breast taking the form of a well defined
streak of pale yellow." He then adds (p. 60) that
the type "unquestionably possessed a complete
pouch, as might be seen from an examination of
the skin preserved in spirit; and Mr. Fraser's note
accompanying the specimen." The animal figured
in the colored plate (op. cit., pi. 76) with chops
and sides of ventrum creamy-buff is not the one
characterized in the text.
In reporting on a second collection from Ec-
uador, made by Mr. Fraser, Tomes mentions a sec-
ond, somewhat smaller specimen of waterhousei
in which the hair of "the region of the pouch and
pubis [is] of a brownish-yellow colour." Finally,
on page 303 (1860), the author offers a figure of
the skull of the type of waterhousei and adds that
the "type possesses a complete pouch, in which,
according to that gentleman's [Fraser's] note, were
several young ones. When it came into my hands
this part contained cotton-wool, and was about the
size of a large hazel-nut."
The type skull is undoubtedly that of a Mar-
mosa. It is impossible, however, to reconcile a
Marmosa skull with a pouched skin. Furthermore,
no known didelphid as small as Marmosa pos-
sesses a marsupium. Although Tomes very defi-
nitely mentions the presence of a pouch, he does
not refer to it in his formal description of the un-
derparts or even notice a reddish mammary region
that covers the ventrum of any breeding female.
These considerations forced Hershkovitz (pers.
comm.) to assume that Tomes may have erred in
his judgment. It was already shown by Thomas
(1921 [9], 7:522) that "Tomes allowed a figure of
Waterhouse's specimen [of Marmosa cinerea,
1846, Nat. Hist. Mamm., 1:505] to be published
as his D. waterhousei when at the same time he
was describing under that name the skin brought
home by Frazer from Ecuador." Thomas then not-
ed that the skull of the type of waterhousei was
in the British Museum but that the skin had dis-
appeared. From the foregoing, it seems advisable
to restrict the name Didelphys waterhousei Tomes
to the type skull and leave the identification of the
skin to those who may find it.
Marmosa bombascarae Anthony, 1922:5 ECUA-
DOR: Zamora-Chinchipe (type locality, Zamora, at
junction of Rio Bombascaro and Zamora; holotype,
female, skin and skull, AMNH 47186, collected 24
November 1920 by H. E. Anthony). Gardner, 1993:
18 listed.
Marmosa maranii Thomas, 1924c:537 PERU: Lor-
eto (type locality, San Lorenzo, Rio Maranon,
above mouth of Rio Huallaga, 500 ft [= 150 m];
holotype, male, skin and skull, BMNH 24.2.22.65,
collected by Latham Rutter). Thomas, 1927:374
a synonym of waterhousei. Gardner, 1993:18 list-
ed.
Marmosa murina maranii: Tate, 1933:102 PERU:
Loreto (San Lorenzo, Rib Maranon; Orosa, oppo-
site mouth of Rio Napo; Pebas, Rio Maranon: Rio
Curaray, mouth); " 'co-type' named by Thomas ap-
pears from the characters of the skin to be a dark
noctivaga. It has no skull."
Marmosa quichua Thomas, 1899:43. Thomas, 1920:
248 PERU: Cusco (type locality, Ocobamba,
9000 ft [= 2800 m]; holotype, female, skin and
skull, BMNH 98.1 1.6.18, collected 2 October 1897
by Otto Garlepp). Thomas, 1928a:265 PERU:
Loreto (Chicosa [now in Ucayali]; Cerro Azul;
Contamana, Rio Ucayali). Tate, 1933: 106 PERU:
San Martin (Moyobamba; Yurac Yacu); Loreto
(Contamana, Rio Ucayali; Cerro Azul; Chicosa
[now in Ucayali]); Hudnuco (Tingo Maria; Rfo
Chinchao); Pasco (Pozuzo); Cusco (Ocobamba;
Marcapata. Rfo Inambari); Junin (San Ramon, near
Chanchamayo; Chanchamayo); musicola Osgood, a
synonym. Ceballos, 1959:266 PERU: Cusco (Ha-
cienda Cadena, Camanti). Gardner, 1993:18 list-
ed.
Marmosa quichua: Cabrera (not Thomas), 1917:10
PERU: Loreto (Rfo Napo).
[Marmosa] quichua: Cabrera (not Thomas), 1911:
BROWN: ATLAS OF NEW WORLD MARSUPIALS
63
12 PERU: Loreto (Rfo Napo); mitis Cabrera, a
synonym.
M[armosa] mitis: Cabrera (not Bangs), 1912:146
PERU: Loreto (Rio Napo).
Marmosa musicola Osgood, 1913b:95 PERU: San
Martin (type locality, Moyobamba; holotype, old
adult female, skin and skull, FMNH 19354, col-
lected 30 July 1912 by W. H. Osgood and M. P.
Anderson). Osgood, 1914b: 150 PERU: San Mar-
tin (Moyobamba). Thomas, 1927a:374 PERU:
San Martin (Yurac Yacu); may be a race of M.
quichua. Gardner, 1933:18 listed.
[Caluromys] musicola: Matschie, 1916:269 classi-
fication.
Marmosa quichua musicola: Thomas, 1927b:608
PERU: Hudnuco (Chinchavito, 3000 ft [= 930 m];
Tingo Maria, 2000 ft [= 625 m]).
Marmosa Klagesi J. A. Allen, 1900a: 198 VENE-
ZUELA: Bolivar (type locality, Ciudad Bolivar;
holotype, male, skin and skull, AMNH 16121; col-
lected 26 October 1899 by S. M. Klages). Thomas,
190 la: 154 GUYANA: Upper Takutu-Upper Es-
sequibo (Kanuku Mts.). J. A. Allen, 1904d:337
VENEZUELA: Bolivar (Ciudad Bolivar; La Union,
Rio Caura). Gardner, 1993:18 listed.
Marmosa murina klagesi: Tate, 1933:98 VENE-
ZUELA: Amazonas (Rio Munduapo, near San Fer-
nando de Atabapo; Bolivar (Ciudad Bolivar; Sua-
pure, Rio Caura; La Union, Rfo Caura). Perez-Her-
nandez, 1989:367 VENEZUELA: Bolivar (Zona
de la Represa del Guri, Rio Caronf; Campamento,
Represa del Guri, Rfo Caronf).
[Marmosops] Klagesi: Matschie, 1916:98 classifi-
cation.
Marmosa chloe Thomas, 1907:167 GUYANA: De-
merara-Mahaica (type locality, Demerara River, 29
mi above Georgetown; holotype, male, skin and
skull, BMNH 7.6.20.16, collected 6 December
1906 by S. B. Warren). Thomas, 1 9 1 0b: 1 89
GUYANA: Pomeroon-Supenaam (Supenaam Riv-
er). Anthony, 1921:267 GUYANA: Cuyuni-Ma-
zaruni (Kartabo). Gardner, 1993:18 listed.
[Marmosops] chloe: Matschie, 1916:271 classifica-
tion.
Marmosa parata Thomas, 1911 a:5 1 7 BRAZIL:
Parti (type locality, Igarape-Assu [= Igarape Acu],
50 m; holotype, female, skin and skull, BMNH
4.7.4.100, collected 12 May 1904 by A. Robert).
Thomas, 1920b:283 BRAZIL: Para (Villa Bra-
ga); Ceard. Gardner, 1993:18 listed.
Marmosa murina parata: Pohle, 1927:241 BRA-
ZIL: Pernambuco (Penha; Dois Irmaos).
Marmosa tobagi Thomas, 191 la:5 15 TRINIDAD
AND TOBAGO: (Richmond); (type locality, Wa-
terloo; holotype, male, skin and skull, BMNH
97.6.7.53, collected 1 1 March 1897 by Perry Ren-
dall). Clark, 1914:69 TRINIDAD AND TOBA-
GO. Gardner, 1993:18 listed.
Marmosa murina tobagi, Goodwin, 1961:9 TRINI-
DAD AND TOBAGO: (Waterloo, north side of is-
land inland from Castara); regarded as specifically
distinct from M. mitis.
Marmosa madeirensis Cabrera, 1913:12 new name
for macrotarsus Wagner, preoccupied. Gardner,
1993:18 listed.
Marmosa murina madeirensis: Tate, 1933:100
BRAZIL: Amazonas (type locality, Rio Madeira;
"co-type," in Vienna Museum, collected by Johann
Natterer). (According to Tate [1933, AMNH], the
type is an adult male, no. 195, formerly mounted,
now preserved as a skin and skull); (Rosarinho,
Lago Miguel, Rio Madeira; Rio Yavarf [= Rio Ja-
varf]; Manacapuni, near mouth of Rio Negro; Villa
Bella Imperatriz [= Parintins], Rio Andira); Para
(Santarem, Rio Tapajos; Villa Braga, Rio Tapajos).
Marmosa murina roraimae Tate, 1931:4 VENE-
ZUELA: Bolivar (type locality, Arabopo [= Ara-
bupii]; foot of Mt. [= Cerro] Roraima, 4200 ft [ =
1300 m]; holotype, male, skin and skull, AMNH
75703, collected 25 December 1927 by G.H.H.
Tate and T. D. Carter). Tate, 1933:98 VENEZUE-
LA: Bolivar (Paulo. Rfo Cuquenam [= Rfo Cu-
quenan]; Mt. [= Cerro] Roraima; Arabupii, Rfo Ar-
abupii, Mt. [Cerro] Roraima). Tate, 1939:164 VE-
NEZUELA: Bolivar (Cerro Roraima. 1400 m; Au-
yan-tepuf, 460 m, 1 100 m, 1850 m). Gardner, 1993:
18 listed.
Marmosa murina duida Tate, 1931:5 VENEZUE-
LA: Amazonas (type locality. Middle Camp, foot
of Cerro Duida, 8 mi N of Esmeralda, upper Rfo
Orinoco, 350 ft [= 110 m]; holotype, male, skin
and skull, AMNH 76984, collected 3 February
1929 by Olalla Bros.). Tate, 1933:99 VENEZUE-
LA: Amazonas (foot of Cerro Duida; Rfo Ocamo,
upper Rfo Orinoco; La Laja. Esmeralda, upper Rfo
Orinoco). Tate, 1939:164 VENEZUELA: Ama-
zonas (foot of Cerro Duida, 100 m). Gardner, 1993:
18 listed. Perez-Hernandez, 1989:366 VENE-
ZUELA: Amazonas (Mavaca, Alto Rfo Orinoco).
Marmosa ruatanica mimetra Tate (part, not Thomas,
1921). 1933:126 part, ECUADOR: Napo (Bae-
za).
Marmosa meridionalis Miranda Ribeiro, 1936:361,
371 BRAZIL: Mato Grosso (type locality, Para-
tudal [= Paratubal], not located; holotype in alco-
hol, MNRJ, collected by the Roosevelt-Rondon
Commission to Mato Grosso). Gardner, 1993:18
listed.
Marmosa murina meridionalis: Cabrera, 1958:19.
Avila Pires, 1968:165 BRAZIL: Mato Grosso
(Paratudal [= Paratubal], not located).
Marmosa moreirae Miranda Ribeiro, 1936:361.
380 BRAZIL: Rio de Janeiro (type locality Ita-
tiaya; holotype, male, skin, MNRJ, collected by Sr.
Carlos Moreira). Gardner, 1993:18 listed.
Didelphys [?sp.] Tomes, 1859:548 ECUADOR: Mo-
rona-Santiago (Gualaquiza).
Types Name based on animals 1, 2, 3, 6 (in-
correctly cited as "12, 36") figured and described
by Seba (1734:48, 49). The specimens in the
Lidth de Jeude collection, BMNH 67.4.12.541-
542, are regarded as being part of the Seba col-
lection and were designated "cotypes" by Thom-
as, 1892:314. These specimens might be labeled
syntypes instead, although the evidence that they
actually figured in Seba's description is specula-
tive. Lectotype designated by Husson 1978:22.
64
FIELDIANA: ZOOLOGY
Type Locality "Asia, America," restricted
to Suriname by Thomas, 1892:314.
Distribution (Fig. 33) Colombia, Venezuela,
Guyana, French Guiana, Suriname, Trinidad and
Tobago, Ecuador, Peru, Brazil, and Bolivia.
Marmosa robinsoni Bangs, 1898
Didelphys murina: Thomas, 1882a:l 1 1 PERU: Ca-
jamarca (Tambillo, 5800 ft [= 1800 m]); Tumbez
(Tumbez). Thomas, 1888b: 343 part, PERU: Tum-
bez (Tumbez); Cajamarca (Tambillo).
Didelphys murina: Thomas (part, not Linnaeus,
1758), 1888b:343 part, COLOMBIA: Antioquia
(Medellin); part ECUADOR: Guayas (Montanas
de Balzar). Robinson, 1896:651 VENEZUELA:
Nueva Esparta (Isla Margarita).
Didelphis (Micoureus) murina, J. A. Allen and Chap-
man, 1893:230 TRINIDAD AND TOBAGO. J.
A. Allen and Chapman, 1897:27 TRINIDAD
AND TOBAGO: St. George (Caura); Caroni (Ca-
paro).
Marmosa murina: Rendall, 1897:342 TRINIDAD
AND TOBAGO.
Marmosa murina: Pirlot (not Linnaeus, 1758), 1963:
175, 178, 179 VENEZUELA: Zulia (Tukuko [ =
El Tukuko]). Tamsitt and Valdivieso (not Linnae-
us), 1964:327 COLOMBIA: Tolima (Mariquita).
Marmosa robinsoni Bangs, 1898a:95. G. M. Allen,
1902:92 VENEZUELA: Nueva Esparta (Marga-
rita [= Isla Margarita]). Handley, 1966:755 PAN-
AMA: Panama (Isla del Rey). Reig, 1968:185
VENEZUELA: Apure (Hato Acapulco). Fleming,
1972:619 PANAMA: Colon (Fort Sherman Mil-
itary Reservation); Panama (Rodman Naval Am-
munition Depot). Handley, 1976:7 VENEZUE-
LA: Zulia (Cerro Azul; Novito); Falcon (Cerro
Santa Ana, Peninsula de Paraguana; Mirimire-La
Pastora; Cerro Socopo; Rfo Socopito); Lara (Cas-
erfo Boro; La Concordia); Trujillo (La Ceiba; Va-
lera); Barinas (Altamira); Carabobo-Falcon-Yara-
cuy (Urama); Carabobo (Montalban); Miranda
(San Andres); Guarico (Hato Las Palmitas; Esta-
cion Biologica de Los Llanos); Nueva Esparta (Isla
Margarita); Sucre (Cumana); Monagas (San Agus-
tin; Hato Mata de Bejuco). Reig et al., 1977:21 1
VENEZUELA: Apure (Mato Salado); Miranda
(Curupao); Guarico (Los Dos Caminos; Estacion
Biologica de Los Llanos). Eisenberg et al., 1979:
191 VENEZUELA: Guarico (Fundo Pecuario
Masaguaral). O'Connell, 1979:717 VENEZUE-
LA: Guarico (Fundo Pecuario Masaguaral). Han-
dley and Gordon, 1979:68 VENEZUELA: Nueva
Esparta (Cerro Matasiete, Isla Margarita). Burton
et al., 1987:103 BELIZE: Toledo (Forestry
Camp). Perez-Hernandez, 1989:368 VENEZUE-
LA: Distrito Federal (Los Canales de Naiguata;
Parque Nacional El Avila); Aragua (Carretera Vieja
de Cata); Barinas (Barinitas; Rio Barragan, cerra
de la desmbocadura, Barinitas; La Erica, 20 km SW
of Barinas); Carabobo (Valencia; Punta Cabito,
Lago de Valencia; Pira Pira); Falcon (Mirimire, 5
km N and 13 km E); Miranda (El Hatillo; La
Union, EI Hatillo; La Providencia, Turgua; Los
Guayabitos; Baruta; La Cienega, Baruta; Cortada
del Guayabo; Estanaque de Curupao, N of Guar-
enas); Monagas (Cerro Papelon); Nueva Esparta
(Serrania de Copey, Margarita; La Sierra, Marga-
rita; Cerro Mastasiete; Fuenteydueno [al Este de la
Isla]; Tacarigua; Hacienda Ochenta); Yaracuy
(Agua Negra; Yaritagua, 30 km N); Apure (La Trin-
idad, Hato La Trinidad de Arauca; "Mato Salada,"
Hato Acapulco): Cojedes (Hato Nuevo; Hato El
Tirado): Guarico (Represa de Guanapito, Altagra-
cia de Orituco; Rio Memo, cerca de Altagracia;
Hato La Palmita, San Francisco de Tiznados; Ha-
cienda "La Palmita"; Estacion Biologica de Los
Llanos, Calabozo; Mondial Villasmil, Calabozo;
Santa Rita, near Rio Manapire; Carlo Bartolo, not
located); Gardner, 1993:18 listed. Adler et al.,
1997:361 COLOMBIA: Cordoba (El Contento).
Lopez-Fuster et al., 2000:836-837 VENEZUE-
LA: Aragua (Carretera Vieja de Cata); Carabobo
(Aguirre, 6 km SSE of Montalban; Cano Alparga-
ton; Embalse Rio Moron; Fundo Caricacho, Ura-
ma; La Batea, 5 km SW of Moron; Montalban,
Punta Cabito, Lago de Valencia; Urama, 19 km
NW); Distrito Federal (Los Canales de Naiguata;
Parque Nacional El Avila); Miranda (Curupao);
Yaracuy (Agua Negra); Monagas (Campamento
M.A.R.N.R., Rio Guarapiche; Caripe); Nueva Es-
parta (Cerro Matasiete; Punta Arena, Peninsula
Macanao; Serranfa de Copey); Sucre (Cariaco,
Chacopata; Guaraunos, Finca Vuelta Larga; Haci-
enda Quetepe, 16 km E of Cumana; Laguna de
Chacopata; Serrania del Turumiquire); Anzodtegui
(Morichal Largo; Rfo Caris, S of El Tigre; Rio Or-
itupano); Monagas (Campamento El Merey, 45 km
S of Temblador; Uverito, 35 km S of Temblador);
Apure (Hato Acapulco; Hato El Frio; La Trinidad
de Arauca); Cojedes (Hato El Tirado; Hato Nuevo;
Hato Pinedo); Guarico (Estacion Biologica de Cal-
abozo; Hato Flores Moradas; Hato La Fe, Coro-
zopando; Hato Las Palmitas); Portuguesa (Estero
de Chirigiiare); Barinas (Reserva Forestal de Ti-
coporo); Falcon (Cerro de Santa Ana, Peninsula de
Paraguana; Coro, 49 km N, 32 km W; Monte Cano,
Pueblo Nuevo).
Marmosa mitis robinsoni: Tate, 1933:118 VENE-
ZUELA: Nueva Esparta (Isla Margarita); Sucre
(Cuchivano, near Cumanacoa; Ypure; Campo Ale-
gre); Monagas (San Antonio de Maturin).
Marmosa robinsoni robinsoni: Cabrera, 1958:24.
Handley and Gordon, 1979:70 VENEZUELA:
Nueva Esparta (Isla Margarita; San Antonio); Su-
cre (Cuchivano).
Marmosa mitis Bangs, 1898b: 162 COLOMBIA: La
Guajira (type locality, Pueblo Viejo, northern slope
Sierra Nevada de Santa Marta; holotype, male, skin
and skull, MCZ 8123, collected 25 March 1898 by
W. W. Brown). Bangs, 1900:90 COLOMBIA:
Magdalena (Palomino, Sierra Nevada de Santa
Marta; La Conception, Sierra Nevada de Santa
Marta); La Guajira (Pueblo Viejo; San Miguel, Si-
erra Nevada de Santa Marta). J. A. Allen, 1904c:
337 VENEZUELA: Bolivar (Ciudad Bolivar). J.
A. Allen, 1904d:4 17 COLOMBIA: Magdalena
(Bonda, Santa Marta; Mamatoco, Santa Marta; Ta-
ganga, Santa Marta; Minca, Santa Marta). J. A. Al-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
65
90 80" 70 60 50 40 30 20 ' 10
Fig. 33. Distribution of Marmosa murina: from northernmost South America south to central Bolivia and southern
Brazil. = localities given in published records.
66
FIELDIANA: ZOOLOGY
len, 1916a:201 COLOMBIA: Cundinamarca (El
Triumfo [= El Triunfo], 600 ft [= 180 m]); Mag-
dalena (Rib Magdalena). Thrasher et al., 1971:
526 COLOMBIA: Atldntico (Barranquilla). Gard-
ner, 1993:19 listed.
Marmosa raids Lonnberg (not Bangs, 1898), 1921:
69 ECUADOR: Pichincha (Nanegal, 5000 ft [=
1560 m]).
Marmosa mitis mitis: Tate, 1933:115 COLOMBIA:
Magdalena (Pueblo Viejo; Palomino; La Concep-
cion; Santa Marta; Rio Don Diego, 45 mi E of San-
ta Marta; Bonda; Cincinati; Taganga; Minca; Ma-
matoco, Rio Manzanares); La Guijira (San Mi-
guel); Tolima (Honda, Rio Magdalena); VENE-
ZUELA: Merida (Milla, Sierra de Merida; Chama,
Sierra de Merida; Montes de Pedregoso [= Que-
brada La Pedregosa], Sierra de Merida).
Marmosa robinsoni mitis: Tate, 1933: 118 VENE-
ZUELA: Monagas (San Antonio de Maturin); Nue-
va Esparta (Isla Margarita); Sucre (Cuchivano; Yp-
ure). Handley and Gordon, 1979:70 COLOM-
BIA: Magdalena (Taganga; Mamatoco; Bondo; La
Concepcion; Minca; Palomino; Pueblo Viejo); La
Guajira (Sierra Negra, E of Villanueva; San Mi-
guel; Villanueva; Las Marimondas); VENEZUE-
LA: Zulia (Cerro Azul; Novito); Merida (Merida;
Cafetos de Milla [= Cafetal de Milla]); Falcon
(Cerro Santa Ana, Peninsula de Paraguana).
Marmosa rapposa: Trouessart (not Thomas, 1899),
1910:A30 ECUADOR: Pichincha (Auca, on road
to Nanegal).
Marmosa simonsi Thomas, 1899b:287 ECUADOR:
(type locality, Puna Island [= Isla Puna], Golfo de
Guayaquil, Guayas; holotype, female, skin and
skull, BMNH 99.8.1.20; collected 3 November
1898 by Perry O. Simons). ECUADOR: Guayas
(Puna [= Isla Puna]; Guayaquil; Chongoncito, near
Guayaquil; Cerro Bajo Verde, near Guayaquil); Los
Rios (Vinces); Manabi (Rib Briceno; Bahia de Ca-
raquez); El Oro (Los Pozos, near Arenillas); Loja
(Valle de Casanga); PERU: Tumbez (near Tumbez);
Cajamarca (Tambillo). Tate, 1933:121 ECUA-
DOR: Guayas (Puna [= Isla Puna]; Hacienda San
Ramon, Puna [= Isla Puna]; Guayaquil; Chongon-
cito; Cerro Bajo Verde); Los Rios (Vinces); Manabi
(Rio Briceno; Bahia de Caraquez); El Oro (Los Po-
zos); Loja (Valley of Casanga [= Valle de Casan-
ga]); PERU: Tumbez (near Tumbez); Cajamarca
(Tambillo). Gardner, 1993:19 listed.
[Caluromys] simonsi: Matschie, 1916:269 classifi-
cation.
Marmosa mitis simonsi: Hershkovitz, 1951:552
classification.
Marmosa robinsoni simonsi: Cabrera, 1958:25 clas-
sification.
Marmosa chapmani J. A. Allen, 1900b: 197 TRIN-
IDAD AND TOBAGO: St. George (type locality,
Caura, Northern range, 500 ft [= 150 m]; holotype,
male skin and skull, AMNH 7666/6052, collected
7 April 1894 by Frank M. Chapman). Elliot, 1907:
4 TRINIDAD AND TOBAGO: St. George (Cau-
ra); Victoria (Princes Town). G. M. Allen, 1911:
194 GRENADA; grenadae Thomas, a synonym.
Tate, 1933:119 TRINIDAD AND TOBAGO: St.
George (Caura; Savanna Grande); Caroni (Capa-
ro); Victoria (Princes Town); St. David (Heights of
Ourepouche [= El Cerro del Oropuche]); GRE-
NADA; TRINIDAD AND TOBAGO; synonyms,
grenadae, tobagi, nesaea. Vesey-Fitzgerald, 1936:
6 TRINIDAD AND TOBAGO: St. George (Ari-
po Valley; Botanic Gardens, Port-of-Spain). Good-
win, 1961:8 GRENADA: (Grand Etang; St.
George); TRINIDAD AND TOBAGO: (Waterloo).
Gardner, 1 993: 1 8 listed.
Marmosa mitis chapmani: Hershkovitz, 1951:552.
Beebe, 1952:181 TRINIDAD AND TOBAGO:
Arima (Arima Valley). Goodwin, 1961:4 TRINI-
DAD AND TOBAGO: St. George (Caura); Caroni
(Caparo); Victoria (Princes Town); St. David
(Heights of Orepouche [= El Cerro del Oropuche];
St. Andrew (Sangre Grande).
Marmosa robinsoni chapmani: Reig et al., 1977:
211 TRINIDAD AND TOBAGO: St. Andrew
(Turure Forest).
Marmosa fulviventer Bangs, 1901:632 PANAMA:
(type locality, San Miguel Island [ = Isla San Mi-
guel], Gulf of Panama, Panama, Pacific side; ho-
lotype, male, skin and skull, MCZ 8435; collected
28 April 1900 by W. W. Brown Jr.). Gardner, 1993:
18 listed.
Marmosa mitis fulviventer: Tate, 1933:117 PANA-
MA: Panama (San Miguel Island [= Isla San Mi-
guel]; Saboga Island [= Isla Saboga]).
Marmosa grenadae Thomas, 191 la:5 14 GRENA-
DA: (type locality, Annandale; holotype, male, skin
and skull, BMNH 87.6.30.5, collected 1886 by the
Eclipse Expedition). Gardner, 1993:18 listed.
Marmosa mitis grenadae: Goodwin, 1961:8 GRE-
NADA: (Grand Etang; St. George's).
Marmosa nesaea Thomas, 191 la:5 15 TRINIDAD
AND TOBAGO: St. George (type locality, Savan-
na Grande); holotype, male, skin and skull, BMNH
97.6.7.24, collected 1 1 January 1897 by Perry Ren-
dall). Gardner, 1993:19 listed.
Marmosa ruatanica Goldman, 1911:237 HONDU-
RAS: (type locality, Roatan Island [= Ruatan Is-
land], one of the islands of Islas de la Bahia, off
north coast of Honduras; holotype, adult male, skin
and skull, USNM 7785/37700, collected 4 Febru-
ary 1864 by J. Akhurst). Gardner, 1993:19 listed.
Marmosa ruatanica ruatanica: Tate, 1933:124
HONDURAS: (Ruatan Island).
Marmosa mitis casta Thomas, 191 la:5 1 6 VENE-
ZUELA: Carabobo (type locality, San Esteban; ho-
lotype, male, skin and skull, BMNH 11.5.25.184,
collected 6 February 1911 by S. M. Klages). J. A.
Allen, 1911:246 VENEZUELA: Carabobo (San
Esteban); Lara (Anzoategui). Tate, 1933:1 16 VE-
NEZUELA: Carabobo (El Trompillo; Puerto Ca-
bello; San Esteban); Lara (Anzoategui); COLOM-
BIA: Norte de Santander (Valera, 10 mi N of Cu-
cuta); pallidiventris Osgood, a synonym. Yepes,
1945:61 VENEZUELA: Yaracuy (San Felipe);
palliviventris Osgood, a synonym.
Marmosa robinsoni casta: Handley and Gordon,
1979:70 VENEZUELA: Falcon (La Pastora; Mir-
imire; Cerro Socopo; Rio Socopito); Lara (Caserio
Boro; La Concordia; Rib Tocuyo); Trujillo (La Cei-
ba; Valera); Barinas (Altamira); Carabobo (Urama;
San Estaban; Montalban); Aragua (Ocumare [ =
Ocumare de la Costa]); Miranda (Curupao; San
BROWN: ATLAS OF NEW WORLD MARSUPIALS
67
Andres); Nueva Esparto (Isla Margarita). Gardner,
1993:18 listed.
Marmosa mitis pallidiventris Osgood, 1912:39 CO-
LOMBIA: Norte de Santander (type locality, El
Guayabal [= Guayabal], 10 mi N of Cucuta: ho-
lotype, female, skin and skull, FMNH 18692. col-
lected 14 March 191 1 by W. H. Osgood and S. G.
Jewett). Gardner. 1993:18 listed.
Marmosa phaea: J. A. Allen (part, not Thomas,
1 899), 1 9 1 2a:73 misidentification.
Marmosa murina zeledoni: J. A. Allen (not Goldman,
1911), 191 2a:73 misidentification.
Marmosa isthmica Goldman, 1912:1 PANAMA:
(type locality, Rio Indio, near Gatiin, Colon; holo-
type, male, skin and skull, USNM no. 170969. col-
lected 16 February 1911 by Edward A. Goldman).
J. A. Allen, 191 6a:200 COLOMBIA: Valle del
Cauca (San Jose, coast; Las Lomitas [= Lomitas]).
Anthony, 1916:363 PANAMA. Enders. 1935:
385 PANAMA: Colon/Panama (Barro Colorado
Island [= Barro Colorado]). Gardner, 1993:18
listed.
Marmosa mexicana isthmica: Goldman, 1917:109.
Enders, 1930:282 PANAMA: Colon/Panama
(Barro Colorado).
Marmosa ruatanica isthmica: Tate, 1933:125 PAN-
AMA: Chiriqui (Boquete); Colon (Gatun, Rio In-
dio; Lago Gatiin); Colon/Panama (Isla Barro Col-
orado); Darien (El Real de Santa Maria. Rio Tuyra;
Tapaliza. near Cerro Tacarcuna; Cerro Tacarcuna;
Santa Cruz de Cana [= Cana]); Panama (Maxon
Ranch, Rio Trinidad); COLOMBIA: Antiquia (Me-
dellin); Choco (Condoto); Cauca (San Jose, coast);
Valle del Cauca (Los Lomitas [= Lomitas]). Tate
(part, not Goldman), 1933:125 COLOMBIA:
(Tusma, Rio Magdalena, not located).
Marmosa mitis isthmica: Hershkovitz. 1951:552
classification.
Marmosa robinsoni isthmica: Cabrera, 1958:24
classification.
Marmosa mimetra Thomas, 1921a:521 ECUADOR:
Pichincha (type locality, Santo Domingo de los Co-
lorados, 1600 ft [= 500 m]; holotype, male, skin
and skull. BMNH 15.1.1.54, collected 19 July 1914
by Gilbert Hammond). Gardner, 1993:18 listed.
Marmosa ruatanica mimetra: Tate, 1933:126 EC-
UADOR: Pichincha (Santo Domingo de los Color-
ados; Baeza; Nanegal: Mindo; Gualea); Guayas
(Balzar: Cerro Manglaralto); Los Rids (Vinces):
Chimborazo (Puente de Chimbo); Imbabura (Par-
amba; Chota); Esmeraldas (San Javier; Caronde-
let); Manabi (El Destino, Rio Cuaque [= Rio
Coaque]; Rio Briceno; Bahia de Caraquez); El Oro
(Santa Rosa).
Marmosa mitis mimetra: Hershkovitz, 1951:552
classification.
Marmosa murina waterhousei: Tate (part, not Tomes,
1860), 1933:103 COLOMBIA: Cundinamarca
(Paime, '"Rio Meta" [Rio Minero]; Bogota; Fusa-
gasuga. W of Bogota); VENEZUELA: Merida (La
Azulita, S of Lake Maracaibo).
Marmosa mitis luridavolta Goodwin, 1961:5 TRIN-
IDAD AND TOBAGO: (type locality, Speyside, 2
mi inland from Tyrrel's Bay; holotype, male, skin
and skull, AMNH 184848; collected 19 March
1960 by Franklin Thurab). Gardner, 1993:18 list-
ed.
Holotype Male, skin and skull, MCZ 7749;
collected 12 July 1895 by Lt. Wirt Robinson.
Type Locality Margarita Island [= Isla Mar-
garita], Nueva Esparta, Venezuela.
Distribution (Fig. 34) Gulf coast of Belize.
Isla Ruatan in Honduras, Panama south through
northern and western Colombia, Ecuador, Peru,
northern Venezuela, Trinidad and Tobago, and
Grenada.
Marmosa rubra Tate, 1931
Marmosa rubra Tate. 1931:6 PERU: Loreto. Tate,
1933:105 PERU: Loreto (Rio Curaray. mouth);
ECUADOR: Napo (below San Jose Nuevo. near
Mt. [= Volcan] Sumaco). Gardner. 1993:19 list-
ed. Pacheco et al.. 1993:1 PERU: Madre de Dios
(Pakitza): Cusco (Hacienda Villa Carmen). Mus-
trangi and Patton. 1997:22 PERU: Amazonas
(Huampami. Rio Cenepa).
Holotype Female, skin and skull, AMNH
71973, collected 7 December 1925 by Olalla
Brothers.
Type Locality Mouth of Rio Curaray, Lore-
to, Peru.
Distribution (Fig. 35) Northeastern Ecuador
and Peru.
Marmosa tyleriana Tate, 1931
Marmosa tyleriana Tate, 1931:6. Tate. 1933:105
VENEZUELA: Amazonas (Cerro Duida plateau).
Tate. 1939:164 VENEZUELA: Amazonas (Cerro
Duida, 4800-6900 ft [= 1500-2150 m]). Perez-
Hernandez. 1989:369 VENEZUELA: Bolivar
(Rio Marajano. Meseta de Jaua). Gardner, 1993:
19 listed.
Marmosa tyleriana phelpsi Tate, 1939:164 VENE-
ZUELA: Bolivar (type locality. Auyan-tepui pla-
teau. 1850 m: holotype. male, skin and skull.
AMNH 130502. collected by G.H.H. Tate). Gard-
ner. 1993:19 listed.
Holotype Female, skin and skull, AMNH
76983. collected 6 February 1929 by Olalla and
Hermanos.
Type Locality Central Camp, Cerro Duida
plateau, upper Rio Orinoco, Amazonas, Venezue-
la, altitude 4800 ft (= 1500 m).
Distribution (Fig. 36) In Venezuela, the Cer-
ro Duida plateau from 1500 m to 2200 m altitude
in the state of Amazonas and the state of Bolivar.
68
FIELDIANA: ZOOLOGY
Fig. 34. Distribution of Marmosa robinsoni in Central and South America: Belize and Isla Ruatan in Honduras;
Panama south through Colombia, Ecuador, and Peru; also northern Venezuela, Trinidad, Tobago, and Grenada. =
localities given in published records.
Marmosa xerophila Handley and Gordon,
1979
Marmosa xerophila Handley and Gordon, 1979:68
COLOMBIA: La Guajira (type locality, La Isla;
near Cojoro, 34-37 km NNE Paraguaipoa in Zulia,
Venezuela, 5-15 m); VENEZUELA: Falcon (Ca-
patarida; Pueblo Nuevo, Peninsula de Paraguana,
13-90 m). Gardner, 1993:19 listed.
Holotype Adult male, skin and skull, USNM
443819, collected 28 June 1968 by N. E. Peterson,
F. B. Brown, Jr., and J. O. Matson.
Type Locality La Isla, near Cojoro, 37 km
NNE of Paraguaipoa, La Guajira, Colombia, ele-
vation 5-15 m.
Distribution (Fig. 37) Northeastern Colom-
bia and northwestern Venezuela.
Genus Marmosops Matschie, 1916
Marmosops Matschie, 1916:262, 267. 270 Didel-
phis incana type by original designation. Reig,
1958:274 (footnote), recognized as a valid subge-
nus equivalent to the Marmosa noctivaga group of
Tate. Gardner, 1993:19 listed.
Type Species Didelphis incana Lund, 1838:
237, by original designation.
Distribution (Fig. 38) Neotropical region
from Panama south to Parana, Brazil.
Comments Previously in Marmosa {sensu
lato; see Gardner & Creighton, 1989, p. 3).
Marmosops bishopi (Pine, 1981)
Marmosa parvidens bishopi Pine, 1981:63 BRA-
ZIL: Mato Grosso (Xavantina [= Chavantina]),
264 km N, Serra do Roncador. Gardner, 1993:20
listed.
Marmosa bishopi: Voss et al., 2001:48 BRAZIL:
Mato Grosso (Xavantina [= Chavantina]); BOLIV-
IA: La Paz (provisionally referred to M. bishopi);
PERU: Huanuco (Montealegre) (provisionally re-
ferred to M. bishopi).
Holotype Adult female, skin and skull,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
69
80 70 60" ~5Q S 40^ 30
Fig. 35. Distribution of Marmosa rubra: eastern Ecuador and Peru. = localities given in published records.
USNM 393535, collected by R. H. Pine, field
number 5157.
Type Locality Xavantina [= Chavantina],
264 km N; Serra do Roncador, Mato Grosso, Bra-
zil, altitude ca. 1750 ft.
Distribution (Fig. 39) Known only from
Mato Grosso, Brazil, Huanuco, Peru, and La Paz,
Bolivia.
Marmosops cracens Handley and Gordon,
1979
Marmosops cracens Handley and Gordon, 1979:68
VENEZUELA: Falcon (La Pastora, 150 m). Gard-
ner, 1993:19 listed.
Marmosops cracens: Perez-Hernandez, 1989:365
VENEZUELA: Falcon (Mirimire, 5 km N and 13
km E, near La Pastora), 170 m.
Type Locality La Pastora, 14 km ENE Mir-
imire, 125-170 m, Falcon, Venezuela.
Holotype Young adult male, skin and skull,
USNM 418503, collected 11 November 1967 by
E. Peterson, F. P. Brown, Jr., and J. O. Matson.
Distribution (Fig. 40)-
con in Venezuela.
-Known only from Fal-
Marmosops dorothea (Thomas, 1911)
Marmosa dorothea Thomas, 191 la:516 BOLIVIA:
La Paz (Rio Solocame, Rio Bopi; Mapiri). Gardner,
1993:19 listed.
[Marmosops] dorothea: Matschie, 1916:271 classi-
fication.
Marmosops dorothea: Mustrangi and Patton, 1997:
21 BOLIVIA: Santa Cruz (Santa Cruz, 15 km S
of Tita). Anderson, 1997:151 BOLIVIA: La Paz
(Serrania Bella Vista; La Reserva; Chijchijpa; Rio
Solocame; Pitiguaya); Santa Cruz (Ascencion, 6
km S; San Ramon, 10 km N; Palmarito, Rib San
Julian; San Miguel Rincon; Cerro Hosana; Buena-
vista; Buena Vista, 4 km SW; Santa Rosita; Esta-
cion Pailon, 3.5 km W; Cerro Amboro, Rib Piti-
sama; El Palmar Road, 7 km SE of Santa Cruz;
Ayacucho; Santa Cruz, 15 km S; Las Lomitas; San-
ta Cruz, 27 km SE; Ingeniero Mora, 1 km E and 3
km N; Hacienda Pelicano; Tita).
Marmosa noctivaga dorothea: Tate, 1933:160 BO-
LIVIA: La Paz (Rio Solocame, Rib Bopi; Mapiri,
near Rio Beni, 2000 ft [= 625 m]).
Marmosa yungasensis Tate, 1931:7 BOLIVIA: La
70
FIELDIANA: ZOOLOGY
Fig. 36. Distribution of Marmosa tyleriana: states of Amazonas and Bolivar in Venezuela. = localities given
in published records.
Paz (type locality, Pitiguaya [= La Florida], Rio
Unduavi, Yungas; holotype, female, skin and skull,
AMNH 72558, collected 10 May 1926 by G.H.H.
Tate). Tate, 1933:160 BOLIVIA: La Paz (Piti-
guaya [- La Florida]). Gardner, 1993:19 listed.
Marmosa ocellata Tate, 1931:7 BOLIVIA: Santa
Cruz (type locality, Buenavista, 500 m; holotype,
old adult male, skin and skull, BMNH 26.1.5.25,
collected 6 August 1924 by J. Steinbach). Tate,
1933:162 BOLIVIA: Santa Cruz (Buenavista).
Gardner, 1993:19 listed.
Marmosa dorothea ocellata: Anderson et al., 1993:
17 BOLIVIA: Santa Cruz (San Miguel Rincon;
Buenavista; Santa Rosita; Camino El Palmar, 7 km
SW of Santa Cruz; Ayacucho; Cerro Hosana).
Holotype Female, skin and skull, BMNH
1.6.7.79, collected 26 January 1901 by P. O. Si-
mons.
Type Locality "Rio Solocame," 16S, 67W,
La Paz, Bolivia, elevation 2300 m.
Distribution (Fig. 41) In Bolivia in the states
of La Paz and Santa Cruz.
Marmosops fuscatus (Thomas, 1896)
Marmosa fuscata Thomas, 1896:313. J. A. Allen,
1911:246 VENEZUELA: Carabobo (La Cumbre
de Valencia, 5000 ft [= 1560 m]). Tate, 1933:
172 VENEZUELA: Merida (La Azulita; Rio Al-
barregas; Merida); Carabobo (La Cumbre de Va-
lencia); COLOMBIA: Cundinamarca (Fusagasu-
ga); Quindio (El Roble); perfusca Thomas, a syn-
onym. Tate, 1947:65 VENEZUELA: Aragua
(Rancho Grande). Handley, 1 976:6 VENEZUE-
LA: Trujillo (Hacienda Misisi'); Carabobo (La
Copa); Aragua (Estacion Biologica de Rancho
Grande); Distrito Federal (Los Venados; Alto No
Leon [= No Leon]); Distrito Federal-Miranda
(Pico Avila); Miranda (Curapao; Instituto Venezo-
lano de Investigaciones Cientificas, 15 km SW of
Caracas); Monagas (San Agustin). Reig et al., 1977:
201 VENEZUELA: Aragua (Rancho Grande); Mi-
randa (Altos de Pipe, Instituto Venezolano de In-
vestigaciones Cientificas [IVIC]). Handley and
Gordon, 1979:67 VENEZUELA: Merida (La
Azulita; Merida; Rio Albarregas; Cafetos de Milla
[= Cafetal de Milla]); Trujillo (Hacienda MisisQ;
Carabobo (La Cumbre de Valencia; La Copa); Ar-
agua (Estacion Biologica de Rancho Grande); Dis-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
71
Fig. 37. Distribution of Mannosa xerophila: northeastern Colombia and northwestern Venezuela. = localities
given in published records.
trito Federal (Los Venados; El Junquito; Alto No
Leon [= No Leon]); Distrito Federal-Miranda
(Pico Avila); Miranda (Curupao; Institute Vene-
zolano de Investigaciones Cientificas [IVIC]. 15
km SW Caracas); Sucre (Cerro Papelon); Monagas
(San Agustin; Santa Ines); TRINIDAD AND TO-
BAGO: St. Andrew (Cumaca [= Cumacca]; Sangre
Grande, Rib Grande Forest); St. David (St. Agus-
tine); Caroni (Caparo). O'Connell, 1979:77
VENEZUELA: Miranda-Guarico (Parque Nacional
Guatopo). Eisenberg et al.. 1979:191 VENEZUE-
LA: Miranda-Guarico (Parque Nacional Guatopo).
Perez-Hernandez, 1 989:366 VENEZUELA Dis-
trito Federal (Parque Nacional *'E1 Avila"; El
Eden, Parque Nacional "El Avila"; Cerro El Avila:
Corozal, Parque Nacional El Avila; El Junquito:
Carretera El Junquito: Carretera Caracas, La Co-
lonia Tovar, km 40; Sabaneta, cerca de El Junqui-
to); Aragua (Estacion Biologica de Rancho Grande.
Parque Nac. "Henry Pittier": Rancho Grande [ =
Estacion Biologica de Rancho Grande]: La Quinta.
Rancho Grande: Palo de Vaca, Rancho Grande;
Pica Periquito, El Portachuelo, Rancho Grande;
Don Ramon, Rancho Grande; Guamitas. Rancho
Grande; Pantoja, Carretera Maracay-Ocumare;
Guacamaya, Rancho Grande); Merida (San Euse-
bio, Carbonera); Miranda (Los Teques; Altos de
Pipe, IVIC; Carrizales via Acuario "Augustin Cod-
azzi"). Gardner. 1993:19 listed.
[Mannosops] fuscata: Matschie, 1916:270 listed.
Thylamys carri J. A. Allen and Chapman. 1897:27
TRINIDAD AND TOBAGO: Caroni (type locali-
ty, Caparo; holotype. male, skin and skull, AMNH
7314/5922, collected 20 March 1894 by Frank M.
Chapman). Gardner. 1993:19 listed.
[Tylomys] Carri: Trouessart. 1897:520 incorrect
subsequent spelling for Thylamys carri, classified
as Rodentia.
[Mannosa] Carri: Trouessart, 1898:1240 classifica-
tion.
Mannosa carri: Tate, 1933:170 TRINIDAD AND
TOBAGO: Caroni (Caparo). Vesey-Fitzgerald.
1936:6 TRINIDAD AND TOBAGO: St. George
(Maracas Valley).
Mannosa fuscata carri: Goodwin, 1961:14 TRINI-
DAD AND TOBAGO: Caroni (Caparo): St. David
(St. Agustine): St. Andrew (Sangre Grande: Cu-
maca [Cumacca]); Arima (Arima).
Mannosa perfusca Thomas. 1924a:236 COLOM-
BIA: Cundinamarca (type locality. Fusagasuga;
holotype. male, skin and skull, BMNH 23.1 1.13.18,
collected September 1923 by Niceforo Maria).
Tate, 1933:173 and footnote type a female, not
72
FIELDIANA: ZOOLOGY
20 10 0"
Fig. 38. Distribution of the genus Marmosops: Panama in Central America, throughout northern South America,
south through Bolivia and Parana, Brazil. = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
73
20 -
Fig. 39. Distribution of Mannosops bishopi: known only from Brazil. Bolivia (not shown), and Peru. = localities
given in published records.
male; regarded as a synonym of fuscata Thomas.
Gardner. 1993:19 listed.
[?]Mannosa caucae caucae: Tate (part, not Thomas).
1933:178 COLOMBIA: Cundinamarca (Bogata;
Fusagasuga, near Bogota).
Type Locality Valdivia, 9 km S, Antioquia,
Colombia, at 1400 m.
Distribution (Fig. 43) Known only from
Cordillera Central in Antioquia, Colombia.
Holotype Female, skin and skull, BMNH
96.1 1.1.6; collected 6 April 1896 by Salamon Bri-
ceno Gabaldon.
Type Locality Rio Albarregas, Merida, Ve-
nezuela, 1630 m altitude.
Distribution (Fig. 42) The eastern Andes of
Colombia, northern Venezuela, and Trinidad and
Tobago.
Marmosops handleyi Pine, 1981
Mannosops handleyi Pine, 1981:67 COLOMBIA:
Antioquia (Valdivia, 1400 m). Gardner. 1983:19
listed, synonymy.
Holotype Adult female, skin, skull and skel-
eton, FMNH 69838, collected 16 June 1950 by
Philip Hershkovitz.
Marmosops impavidus (Tschudi, 1844)
D[idelphys] impavida Tschudi. 1844:14. 149. 151
PERU: (type locality, '"mitteren und tiefen Waldre-
gion." not over 3500 ft [= 1090 m] elevation, and
between 10 and 12S). Thomas. 1927:607 type
lost; "had best be considered as a synonym of noc-
tivaga." Gardner. 1983:19 listed.
Didelphys impavida: Waterhouse. 1846:526. Mat-
schie. 1916:263. 272 holotype said to be in Berlin
Zoological Museum, mounted skin no. 3375. skull
no. 24771.
[Marmosa] impavida: Trouessart. 1898:1239 listed
in synonymy of Marmosa murina.
M[armosa] impavida: Cabrera. 1912:146 ECUA-
DOR: Napo (Rio Coca).
Marmosa impavida: Osgood. 1914b: 149 PERU:
Lorero (Yurimaguas). Osgood. 1915:188 PERU:
Junin (specimens from San Ramon. Rio Chancha-
mayo. regarded as topotypical). Thomas. 1920c:
248 PERU: Cusco (Torontoy. 8000 ft [= 2500
74
FIELDIANA: ZOOLOGY
80 70 60 50 W 30
Fig. 40. Distribution of Marmosops cracens: northern Venezuela. = localities given in published records.
m]). Handley, 1966:755 PANAMA: Darien (Cer-
ro Pirre, 5300 ft [= 1650 m]; Loma Cana). Hand-
ley, 1976:7 VENEZUELA: Tdchira (Buena Vis-
ta). Handley and Gordon, 1979:68 COLOMBIA:
Valle del Cauca (Rio Cauquita, near Cali, 1000 m);
La Guajira (Sierra Negra, near Villanueva); VEN-
EZUELA: Tdchira (Buena Vista, near Paramo de
Tama, 2380-2415 m). Perez-Hernandez, 1989:
366 VENEZUELA: Tachira (Betania, Junin; Cer-
ro Las Copas, La Revancha; San Cristobal [= Bue-
na Vista], 35 km S and 22 km W); Amazonas (Cer-
ro La Neblina). Anderson et al., 1993:16 BOLIV-
IA: Santa Cruz (Caranda).
Marmosa impavida impavida: Cabrera, 1958:16
PERU: Junin (Pueblo Nuevo [= Montana Vitoc],
Cerca de Chanchamayo) ucayaliensis Tate, a syn-
onym.
Marmosops impavidus: Pacheco et al., 1993 PERU:
Cusco (above San Pedro). Anderson, 1997:153
BOLIVIA: Beni (Embocada); Cochabamba (Inca-
chaca); La Paz (Caranavi, 20 km NNE); Pando (In-
dependencia); Santa Cruz (Caranda). Patton et al.,
2000:53 BRAZIL: Amazonas (Barro Vermelho,
left bank Rio Jurua; Seringal Condor, left bank Rio
Jurua; Altamira, right bank Rio Jurua; Coloca?ao
Vira-Volta, left bank Rio Jurua); Acre (Igarape Po-
rongaba, right bank Rio Jurua); BOLIVIA: Santa
Cruz (Tita).
Marmosa caucae Thomas, 1900c:221 COLOMBIA:
Valle del Cauca (type locality, Rio Cauquita, Cali,
1000 m; holotype, male, skin and skull, BMNH
99.9.6.51; collected August 1897 by J. M. Batty).
Gardner, 1993:19 listed.
Thylamys caucae: J. A. Allen, 1912a:73 COLOM-
BIA: Valle del Cauca (San Antonio, 7000 ft [ =
2180 m]). J. A. Allen, 1 9 16a:201 COLOMBIA:
Valle del Cauca (San Antonio); Narino (Ricaurte,
5000 ft [= 1560 m]); Quindio (near Salento); Cun-
dinamarca (El Roble).
[Marmosops] caucae: Matschie, 1916:270 listed.
Marmosa caucae caucae: Tate, 1933:178 COLOM-
BIA: Narino (Ricaurte, 5000 ft [= 1560 m]); Valle
del Cauca (San Antonio, Rio Cauca, near Cali,
7000 ft [= 2180 m]; Rio Cauquita, near Cali);
Quindio (western Quindio, not located, 9000 ft [=
2800 m]).
[l]Marmosa caucae caucae: Tate (part, not Thomas,
1900), 1933:178 COLOMBIA: Cundinamarca
(Mambita, upper Rio Meta, eastern Cundinamarca:
Medina, upper Rio Meta, eastern Cundinamarca;
Fusagasuga; Bogota); Meta (Guaicaramo, upper
Rio Meta); ECUADOR: Napo (Baeza; Palmera,
Rio Pastaza, not located; San Antonio, Rio Ulva;
San Jose de Sumaco [= San Jose Nuevo]).
Marmosa impavida caucae: Cabrera, 1958:16 clas-
sification.
Marmosa sobrina Thomas, 1913c:573 ECUADOR:
Pichincha (type locality, Mindo, NW of Quito,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
75
Marmosops dorothea
Fig. 41. Distribution of Marmosops dorothea: west of Rio Beni in La Paz and east of Santa Cruz. Bolivia.
localities given in published records.
4200 ft [= 1300 m]: holotvpe. male, skin and skull.
BMNH 13.10.24.70. collected 3 July 1913 by Gil-
bert Hammond). Gardner. 1993:19 listed.
[Marmosops] sobrina: Matschie. 1916:270 classifi-
cation.
Marmosa caucae sobrina: Tate. 1933:179 ECUA-
DOR: Pichincha (Mindo: Santo Domingo de los
Colorados): Chimborazo (Rio Chanchan: Puente de
Chimbo: Pallatanga): Guayas (Ventura).
Marmosa madescens Osgood. 1913b:94 PERU:
Amazonas (tvpe localitv. Tambo Ventija [= Yentil-
la]. 9000 ft [= 2800 m]; holotvpe. male, skin and
skull. FMNH 19689. collected 15 June 1912 by W.
H. Osgood and M. P. Anderson). Osgood. 1914b:
149 PERU: San Martin (Tambo Almirante. near
Uchco [= Uscho]): Amazonas (Tambo Ventija [ =
Ventilla]): San Martin (Puca Tambo: Moyobamba).
Thomas. 1927a:375 PERU: Amazonas (San Ni-
colas [= Mendoza]. 4500 ft [= 1400 m]). Thomas.
1928a:265 PERU: Loreto (Contamana. Rio Uca-
vali: San Jeronimo. Rio Ucayali). Gardner. 1993:
19 listed.
[Marmosops] madescens: Matschie. 1916:270 clas-
sification.
Marmosa caucae madescens: Tate. 1933:181
PERU: Cajamarca (Taulis. west of Cajamarca.
9000 ft [= 2800 m]): San Martin (Moyobamba:
Puca Tambo: Yurac Yacu: Uchco [= Uscho]. 5000
ft [= 1560 m]): Amazonas (Tambo Ventija [= Ven-
tilla]).
Marmosa impavida madescens: Cabrera. 1958:16
classification.
Marmosa oroensis Anthony. 1922:3 ECUADOR: El
Oro (type locality. Portovelo. 2000 ft [= 625 m]:
holotvpe. male, skin and skull. AMNH 47180. col-
lected 2 September 1920 bv H. E. Anthonv). Gard-
ner. 1993:19 listed.
Marmosa caucae oroensis: Tate. 1933:180 ECUA-
DOR: El Oro (Portovelo: Alamor [now in Loja]:
Guainche. Alamor [now in Loja]: Salvias; Tunama.
near Alamor [now in Loja]): Loja (Celica. 6900 ft
[= 2150 m]): celicae Anthony, a synonym.
Marmosa impavida oroensis: Cabrera. 1958:17
classification.
Marmosa celicae Anthony. 1922:4 ECUADOR:
Loja (type locality. Celica. 6900 ft [= 2150 m]:
holotype. male, skin and skull. AMNH 47128. col-
lected 28 September 1920 by H. E. Anthony).
Gardner. 1 993 : 1 9 listed.
Marmosa caucae ucayaliensis Tate. 1931:9 PERU:
Ucayali (type locality. Lagarto. Rio Ucayali. 25 m
below mouth of Rio Tambo: holotvpe. male, skin
and skull. AMNH 78952. collected 20 March 1928
by Olalla and Sons): PERU: Ucayali (Lagarto. Rio
Ucayali): Loreto (San Jeronimo. Rio Ucayali: Con-
tamana. Rio Ucavali: Leguia. Rio Pachitea. not lo-
76
FIELDIANA: ZOOLOGY
20-'
Fig. 42. Distribution of Marmosops fuscatus: west-central Colombia, north and east through northern Venezuela,
Trinidad and Tobago. = localities given in published records.
cated; Rio Curaray, mouth at Rio Napo). Gardner,
1993:19 listed.
Marmosa caucae albiventris Tate, 1931:9 PERU:
Cusco (type locality, Torontoy, 8000 ft [= 2500 m];
holotype, male, skin and skull, USNM 194378, col-
lected 30 April 1915 by E. Heller). Tate, 1933:
182 PERU: Cusco (Torontoy). Gardner, 1993:
19 listed.
Marmosa impavida albiventris: Cabrera, 1958:16
classification.
Marmosa albiventris: Mustrangi and Patton, 1997:
22 PERU: Cusco (Amaybamba, 2 km NE and 3
km E, Lucumayo Valley; Paucartambo, 72 km NE).
[?]Marmosa fuscata Tate (?part, not Thomas, 1896),
1933:172 COLOMBIA: Cundinamarca (El Ro-
ble).
Holotype None designated. Cotype is said to
be in Berlin Zoological Museum, mounted skin
no. 3375, skull no. 24771.
Type Locality "Der mittleren und tiefen
Waldregion," not over 1061 m and between 10
and 12S, interpreted by Cabrera, 1958:16, as
"Montana de Vitoc, circa de Chanchamayo, Ju-
nin, Peru."
Distribution (Fig. 44) Mountains of western
Panama to Colombia, Ecuador, Peru, and Bolivia,
and western Brazil, with an isolated population in
Tachira and Amazonas, Venezuela.
Marmosops incanus (Lund, 1840) [preprint of
Lund, 1841]
Didelphis incana Lund, 1839:233 nomen nudum;
Lund, 1841:236-238, 265, 293. Lund, 1842:133,
199 BRAZIL: Minas Gerais (Lagoa Santa). Wa-
terhouse, 1846:503. Gardner, 1993:19 listed.
D[idelphvs] incana: Wagner, 1845:40 description ex
Lund. Thomas, 1 894b: 1 84 BRAZIL: Minas Ger-
ais; Rio de Janeiro; not a synonym of Didelphys
grisea Desmarest.
M[armosa] incana: Thomas, 1896:313 (footnote)
Didelphys grisea Thomas (not Desmarest) of Cat.
Mus. B. M. p. 349.
Marmosa incana: Miranda Ribeiro, 1935:37 BRA-
ZIL: Rio de Janeiro (Teresopolis). Cabrera, 1958:
17 classification; synonyms, bahiensis Tate, pau-
lensis Tate. Avila Pires and Gouvea, 1977:9
BRAZIL: Minas Gerais (Lagoa Santa); Rio de Ja-
neiro (Monte Serrat). da Fonseca and Kierluff,
1988:114 BRAZIL: Minas Gerais (Fazenda
BROWN: ATLAS OF NEW WORLD MARSUPIALS
77
Marmosops handleyi
1500 km
30
Fig. 43. Distribution of Marmosops handleyi: known only from Antioquia, Colombia. = locality given in
published records.
Montes Claros: Fazenda Esmeralda; Rio Doce State
Park [= Parque Estadual do Rio Doce]). Stallings,
1988:174 BRAZIL: Minas Gerais (Rio Doce
State Park [= Parque Estadual do Rio Doce]).
Tribe, 1990:566-567 BRAZIL: Minas Gerais
(Alem Parafba); Bahia (Lamarao); Rio de Janeiro
(Teresopolis).
Marmosops incana: Matschie, 1916:271 BRAZIL:
Minas Gerais (Lagoa Santa).
Marmosops incanus: Mustrangi and Patton, 1997:
19 BRAZIL: Bahia (Lamarao; Serrinha; Sao
Goncalo dos Campos, 30 km SW of Feira de San-
tana; Seabra; Jequie; Jaguaquara; Itabuna, 25 km
W of Ilheus; Ilheus; Porto Seguro); Minas Gerais
(Mocambinho, Manga, Rio Sao Francisco; Fabiao,
Itacarambi, Rio Peruacu; Parque Florestal Estadual
de Jafba, Jaiba; Grota Grande, Turmalina; Parque
Nacional de Serra do Cipo, Jaboticatubas, in south-
ern Serra do Espinhaco; Conceicao do Mato Den-
tro, Serra do Espinhaco; Lagoa Santa; Esta^ao de
Pesquisas de Peti; Sao Goncalo do Rio Abaixo;
Parque Estadual do Rio Doce, Marlieria; Estacao
Biologica de Caratinga, Fazenda Montes Claros,
Caratinga; Parque Nacional de Caparao, Serra do
Caparao; Fervedouro; Alem Parafba; Volta Grande;
Juiz de Fora; Porto Novo; Simao Pereira); Espirito
Santo (Reserva Florestal Companhia Vale do Rio
Doce, Linhares; Estacao Biologica de Santa Lucia,
Santa Tereza; Reserva Biologica Nova Lombardia,
Santa Tereza; Rio Sao Jose; Aracruz Florestal, Ar-
acruz; Campinho); Rio de Janeiro (Teresopolis,
Serra dos Orgaos; Mendes; Sao Joao Marcos;
Parque Estadual da Pedra Branca; Floresta da Ti-
juca and Silvestre, Rio de Janeiro; Reserva Biolo-
gica de Tingua, Nova Iguagu; Ibicuf, Mangaratiba;
Angra dos Reis; Ilha Grande, Angra do Reis;
Parque Nacional do Itatiaia, Itatiaia, Serra da Man-
tiqueira); Sao Paulo (Estagao Biologica de Bora-
ceia, Salesopolis, Serra do Mar; Sao Sebastiao; Ilha
de Sao Sebastiao; Caucaia do Alto, Cotia; Sao Pau-
lo; Fazenda Intervales [base do Carmo]; Capao Bo-
nito, Serra de Paranapiacaba).
Marmosa incana incana: Tate, 1933:164 BRAZIL:
Minas Gerais (Lagoa Santa; Mandi, Lagoa Santa).
Grymaeomys incanus Burmeister, 1856:83.
M[armosa] incanus: Thomas, 1894b: 186 (in text)
classification.
Grymaeomys scapulatus Burmeister, 1856:79 BRA-
ZIL: Minas Gerais (type locality, Lagoa Santa; ho-
lotype, male, skin and skull, Berlin Museum, skin
no. 2330, skull no. 26000; collected by Ignatz von
Olfers). Gardner, 1993:19 listed. Mustrangi,
1994:252 BRAZIL: Sao Paulo (Intervales Bio-
logical Reserve).
Marmosa scapulatus: Tate, 1933:167 BRAZIL: Mi-
nas Gerais (holotype).
78
FIELDIANA: ZOOLOGY
500 1000 1500 km
30
Fig. 44. Distribution of Marmosops impavidus: in South America from Colombia south through Ecuador, Peru,
Bolivia, and northwestern Brazil, with an isolated population in Venezuela. = localities given in published records.
[Marmosops] scapulata: Matschie, 1916:271 listed.
[Marmosa] scapanus: Trouessart, 1898:1238 incor-
rect subsequent spelling- lapsus for scapulatus in
synonymy of Marmosa cinerea.
Didelphys cinerea: Thomas (part, not Temminck,
1824), 1888b:342 part, scapulata in synonymy
only.
Didelphys grisea: Lydekker (not Desmarest, 1827),
1887:281. Thomas, 1888b:349 part, BRAZIL:
Rio de Janeiro.
Grymaeomys griseus: Winge (not Desmarest, 1827),
1893:6, 12 BRAZIL: Minas Gerais (Lagoa San-
ta).
Micoureus griseus: Goeldi (not Desmarest, 1827),
1894:461 BRAZIL: Rio de Janeiro (Serra dos Or-
gaos).
Marmosa incana bahiensis Tate, 1931:8 BRAZIL:
Bahia (type locality, Lamarao; holotype, male, skin
and skull, BMNH 3.9.5.137, collected 1903 by A.
Robert). Tate, 1933:165 BRAZIL: Bahia (Lamar-
ao). Gardner, 1993:19 listed.
Distribution (Fig. 45) Eastern Brazil from
Bahia south to Parana.
Marmosops invictus (Goldman, 1912)
Marmosa invicta Goldman, 1912:3. Tate, 1933:171
PANAMA: Darien (Cana). Handley, 1966:755
PANAMA: Darien (Tacarcuna Laguna). Handley
and Gordon, 1979:68 PANAMA: Bocas del Toro
(Cylindro, 1220+ m); Panama (Cerro Azul, 640
m); Darien (Cerro Tacarcuna, 1464 m; La Laguna,
975 m; Casita, 458 m; Cana, 610 m). Pine, 1981:
66 PANAMA: Panama (Cerro Azul); Darien
(Cerro Tacarcuna, 4800 ft [= 1500 m]; Tacarcuna
Laguna Camp, 4000 ft [= 1250 m]; Tacarcuna Ca-
sita Camp, 2700 ft [= 840 m]; Cana). Gardner,
1993:19 listed.
[Marmosops] invicta: Matschie, 1916:270 PANA-
MA: Darien (Cana).
Holotype Two cotypes, males, no. 223 (in
spirits); male skin no. 224 and skull in Copen-
hagen Museum, collected by P. W. Lund.
Type Locality Lagoa Santa, Minas Gerais,
Brazil.
Holotype Male, skin and skull, USNM
178708; collected 14 March 1912 by E. A. Gold-
man.
Type Locality Cana, Darien, Panama, alti-
tude 2000 ft (= 625 m).
BROWN: ATLAS OF NEW WORLD MARSUPIALS
79
Fig. 45. Distribution of Marmosops incanus: eastern Brazil from Bahia south to Parana.
published records.
10
localities given in
Distribution (Fig. 46) Panama.
Marmosops juninensis (Tate, 1931)
Marmosa juninensis Tate, 1931:13 PERU: Junin
(Utcuyacu, between Tarma and Chanchamayo).
Tate, 1933:207 PERU: Junin (Utcuyacu). Gard-
ner, 1993:20 listed.
Marmosa parvidens juninensis: Pine, 1981:65
PERU: Junin (Utcuyacu).
Marmosops juninensis: Voss et al., 2001: 47 PERU:
Junin (Utcuyacu, "4800 ft near Tarma"; Tarma).
Holotype Female, skin and skull, AMNH
63864, collected 15 November 1919 by H. Wat-
kins.
Type Locality Utcuyacu, between Tarma and
Chanchamayo, altitude 4800 ft (= 1500 m).
Distribution (Fig. 47) Known only from Ut-
cuyacu in Junin, Peru.
(Camp II, Camp VI, Camp VII, Camp XI, Cerro
de La Neblina); Cerro da La Neblina.
Marmosops impavidus neblina: Gardner, 1990:414
VENEZUELA: Amazonas (Cerro de La Neblina).
Marmosops neblina: Mustrangi and Patton, 1997:
56 VENEZUELA: Amazonas (Cerro de La Neb-
lina, Camp VII). Patton et al., 2000:53 BRAZIL:
Acre (Igarape Parongaba; Nova Vida; Ocidente);
Amazonas (Igarape Nova Empresa, left bank Rio
Jurua; Altamira, right bank Rio Jurua; Altamira,
opposite, left bank Rio Jurua; Jainu, right bank Rio
Jurua); VENEZUELA: Amazonas (Cerro de La
Neblina, Camp VII).
Holotype Adult male, AMNH 560735; col-
lected 5 February 1985 by Alfred L. Gardner.
Type Locality Camp VII, Cerro de La Neb-
lina, Amazonas, Venezuela, altitude 1800 m.
Distribution (Fig. 48) Cerro de La Neblina
in Amazonas, Venezuela and Rio Jurua region,
Amazonas, Brazil.
Marmosops neblina (Gardner, 1990)
M[armosa] impavida neblina Gardner, 1988:698
name cited as in press; VENEZUELA: Amazonas
Marmosops noctivagus (Tschudi, 1845)
D[idelph\s] noctivaga Tschudi, 1845:148. Gardner,
1993:20 listed.
80
FIELDIANA: ZOOLOGY
110 100
Fig. 46. Distribution of Marmosops invictus: Panama in Central America.
records.
localities given in published
Didelphys noctivaga: Waterhouse, 1846:527.
[Marmosa] noctivaga: Trouessart, 1898:1238 listed
in synonymy of Marmosa cinerea. Cabrera, 1913:
12 classification.
M[annosa] noctivaga: Thomas, 1920c:248 type ex-
amined in Neuchatel Museum; impavida Tschudi
regarded a synonym.
Marmosa noctivaga: Thomas, 1926e:616 PERU:
Amazonas (Tambo Carrizal). Thomas, 1927b:606
PERU: Hudnuco (Chinchavito, 3000 ft [= 930 m];
Tingo Marfa, 2000 ft [= 625 m]); type examined;
type of impavida lost, probably = noctivaga.
Thomas, 1928b:294 part, PERU: Loreto (Pebas).
Terborgh et al., 1984:27 PERU: Madre de Dios
(Cocha Cashu Biological Station, Parque Nacional
del Manu). Rageot and Albuja, 1994:176 EC-
UADOR: Pastaza (Mera).
[Marmosops] noctivaga: Matschie, 1916:270 listed.
Marmosops noctivaga: Emmons, 1990:72 BOLIV-
IA: La Paz (Ixiamas). Emmons and Romo, 1994:
140 PERU: Madre de Dios (Ccolpa de Guaca-
mayas). Emmons, Ascorra and Romo, 1994:146
PERU: Madre de Dios ("Aguas Claras" Camp).
Emmons, Barkley and Romo, 1994:144 PERU:
Madre de Dios (Explorer's Inn Reserve).
Marmosa noctivaga noctivaga: Tate, 1933:152
PERU: Juni'n (Vitoc; Chanchamayo; San Ramon,
Chanchamayo; Rio Cayumba, not located); Hudn-
uco (Tingo Maria; Chinchavito; Rio Chinchao; Po-
zuzo [now in Pasco]); Loreto (Pebas); ECUADOR:
Tungurahua (Mirador, Rio Napo); Pastaza (Cane-
los; Rio Copataza); Morona-Santiago (Gualaqui-
za). Cabrera, 1958:22 classification.
Marmosops noctivagus: Pacheco et al., 1993:5
PERU: Cusco (Hacienda Villa Carmen; Pilcopata;
Albergue Bosque de las Nubes at Puerte Union);
Madre de Dios (Pakitza). Mustrangi and Patton,
1997:21 BRAZIL: Amazonas (Penedo. right bank
Rio Jurua). Patton et al., 2000:53 PERU: Cusco
(Amaybamba, 2 km NE, Lucumayo Valley; Pau-
cartambo); BRAZIL: Amazonas (Macaco, left bank
Rio Jau; Tambor, left bank Rio Jau; Penedo, right
bank Rio Jurua; Seringal Condor, left bank Rio Ju-
rua; Lago Vai-Quem-Quer, right bank Rio Jurua);
Acre (Igarape Porongaba, right bank Rio Jurua;
Igarape Porongaba, opposite, left bank Rio Jurua;
Nova Vida, right bank Rio Jurua); BOLIVIA: La
Paz (Chijchijpa).
Thylamys keaysi J. A. Allen, 1900b: 198 PERU. J.
A. Allen, 1900c:219 PERU: Puno (type locality,
Inca Mines, not Juliaca; holotype, male, skin and
skull, AMNH 16068, collected 8 January 1900 by
H. H. Keays). J. A. Allen, 1901a:42 PERU: Puno
(Inca Mines, Rio Inambari). Gardner, 1993:20
listed.
[Marmosa] Keaysi: Trouessart, 1904:856 classifica-
tion.
Marmosa noctivaga keaysi: Tate, 1933:158 PERU:
Puno (Inca Mines); Cusco (Marcapata, Rio Inam-
bari; Machu Picchu); BOLIVIA: La Paz (Ticun-
huaya, Rio Tipuani, 4800 ft [= 1500 m]); Cocha-
bamba (Incachaca). Sanborn, 1951b:2 PERU:
BROWN: ATLAS OF NEW WORLD MARSUPIALS
81
Marmosops junenensis
500 1000 1500 km
Fig. 47. Distribution of Marmosops juninensis: known only from Junin. Peru. = locality given in published
records.
Cusco (Hacienda Cadena. Quincemil). Anderson et
ah. 1993:17 BOLIVIA: Santa Cruz (San Rafael
de Amboro). Anderson. 1997:154 BOLIVIA:
Beni (Embocada): Chuquisaca (Rio Limon): Co-
chabamba (Yungas: Tablas Monte. 9.5 km NE: Ta-
blas Monte. 4.4 km N): La Paz (Moira Camp: Alto
Rio Madidi: Ixiamas: Mapiri: Ticunhuaya: Puerto
Linares: Chijchijpa): Pando (Puerto Camacho. 15
km NW: Palmira: Santa Rosa: Santa Rosa. 8 km
S\V): Santa Cruz (Perseveranciai.
Marmosa polita Cabrera. 1912:10 ECUADOR:
Napo (type locality. Rib Coca, near mouth at Rio
Napo: holotype. male, skin and skull. Museo de
Ciencias Naturales. Madrid, no. 797. collected by
Marcos Jimenez de la Espada). Cabrera. 1917:10
ECUADOR: Sapo (Rio Coca). Gardner. 1993:20
listed.
Marmosa noctivaga polita: Tate. 1933:155 ECUA-
DOR: Xapo (San Jose de Sumaco [= San Jose
Nuevo]. ca. 3500 ft [= 1090 m]): PERU: Loreto
(Rib Curaray. mouth: Orosa. mouth of Rib Napo):
BR.AZIL: Amazonas (Santa Rita. Tabatinga: Igar-
ape Cacao Pereira [= Cacau Pirera]. Rio Negro).
Baker. 1974:135 ECUADOR. \apo (Limon^Co-
cha [= Limoncocha]).
Marmosa purui Miller. 1913:31 BRAZIL: Amazon-
as (type locality. Iutanaa [= "Hyutanaham" =
Huitanaa]: holotype. male, in alcohol, USNM
105543: collected' March 1901 by J. B. Steere).
[Thylamys] purui: Matschie. 1916:271 listed.
Marmosa caucae purui: Tate. 1933:18-1 BRAZIL:
Amazonas ("Hyutanaham" [= Huitanaa]. upper
Rio Purus: Rosarinho. Lago Miguel. Rio Madeira).
Marmosa impavida purui: Cabrera. 1958:17 classi-
fication.
Marmosa impavida neglecta Osgood. 1915:187
PERU: Loreto (type locality. Yurimaguas. Rib
Huallaga. 600 ft [= 180 m]: holotype. female, skin
and skull. FMNH 19636. collected 28 September
1912 bv M. R Anderson). Gardner. 1993:20 list-
ed.
M[armosa] noctivaga neglecta: Thomas. 1924c:
358 classification.
Marmosa noctivaga neglecta: Tate. 1933:159
PERU: Loreto (Yurimaguas): Hudnuco (Tingo Ma-
ria): "regarded as a red phase of the female sex of
noctivaga."'
Marmosa collega Thomas. 1920b:281 BRAZIL:
Para (type locality. Villa Braga. Rib Tapajos: ho-
lotype. male, skin and skull, BMNH 20.7.14.40:
collected 7 June 1916 by Emilie Snethlage). Gard-
ner. 1993:20 listed.
Marmosa noctivaga collega: Tate. 1933:157 BRA-
82
FIELDIANA: ZOOLOGY
20
80 70 60 50* 3IT 30
Fig. 48. Distribution of Marmosops neblina: Rio Jurua region of Amazonas, Brazil, and southern Venezuela.
= localities given in published records.
ZIL: Para (Villa Braga, Rio Tapajos). Avila Pires,
1968:166 BRAZIL: Mato Grosso (Aripuana).
Marmosa leucastra Thomas, 1927b:607 PERU:
Amazonas (type locality, Tambo Carrizal, 40 mi S
of Chachapoyas, 9000 ft [ = 2800 m]; Chachapoy-
as). Tate, 1933:161 PERU: Amazonas (Tambo
Carrizal; Chachapoyas; near Perico, Rio Maranon
Valley, not located; may be Perico in Cajamarca).
Gardner, 1993:20 listed.
Marmosa stollei Miranda Ribeiro, 1936:261, 272
BRAZIL: Amazonas (type locality, Rio Aripuanan
[= Rio Aripuana], lower Rio Madeira; holotype,
female taken from stomach of snake [Cenchris],
collected by E. Stolle). Gardner, 1993:20 listed.
Marmosa noctivaga lugenda Thomas, 1927a:373
PERU: San Martin (type locality, Yurac Yacu; ho-
lotype, male, skin and skull, BMNH 27.1.1.178,
collected 2 July 1926 by R. W. Hendee). Tate,
1933:157 PERU: San Martin (Yurac Yacu); EC-
UADOR: Morona-Santiago (Guayabo, near Guala-
quiza); Pastaza (Mera, Rio Pastaza). Gardner,
1993:20 listed.
Holotype Not known to be in existence; ac-
cording to Tate (1933, Bull. AMNH 66:153),
"Berlin 3375, a rather young female (?) with
small teeth, marked as collected by Tschudi, is to
be considered a co-type. None are to be found in
Neuchatel. No others elsewhere." However, ac-
cording to Matschie (1916:263, 272), the Berlin
specimen is type of Didelphys impavida Tschudi.
Type Locality Forest region between 1 and
12S and not above 1061 m; restricted to the Mon-
tana de Vitoc, near Chanchamayo, Junin, by Tate
(1933, pp. 153, 155) on the basis of Tschudi's
account of the species (cf. Reisenskizzen, 2:248-
249). Osgood (1915, p. 188) regarded as "abso-
lutely typical" a specimen from San Ramon, Rio
Chanchamayo. This locality is in the same region
as the Montana de Vitoc, specifically mentioned
by Tschudi in the Reisenskizzen.
Distribution (Fig. 49) Ecuador, Peru, Boliv-
ia, and Amazonian Brazil.
Marmosops parvidens (Tate, 1931)
Marmosa parvidens Tate, 1931:13. Tate, 1933:208
type only. Pine, 1973:55 BRAZIL: Para (Belem).
Handley, 1976:6 VENEZUELA: Carabobo-Fal-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
83
Fig. 49. Distribution of Mannosops noctivagus: in South America from Ecuador south through Peru, Bolivia,
and Amazonian Brazil. = localities given in published records.
con-Yaracuy (Urama); Amazonas (Capibara. Brazo
Casiquiare); Bolivar (km 125, 85 km SSE of El
Dorado). Handley and Gordon. 1979:68 VENE-
ZUELA: Carabobo-Falcon-Yaracuy (Urama):
Amazonas (Capibara. Brazo Casiquiare): GUYA-
NA: Demerara-Mahaica (Hyde Park). PERU: dis-
co (Hacienda Villa Carmen). Gardner. 1993:20
listed.
Mannosops pan'idens: Pacheco et al.. 1993:5
PERU: Madre de Dios (Pakitza). Mustrangi and
Patton. 1997:21 BOLIVIA: La Paz (La Reserva);
BRAZIL: Amazonas (Barro Vermelho. left bank
Rio Jurua; Ilhazinha, left bank Rio Jurua): Paid
(Altamira. 52 km SSW. E bank Rio Xingu): GUY-
ANA: Upper Takutu-Upper Essequibo (Karanam-
bo. Rupununi River). Patton et al.. 2000:53
GUYANA: Upper Takutu-Upper Essequibo (Kar-
anambo. Rupununi River, ca. 45 km N of Kanuku
Mtns.); BRAZIL: Para (Altamira. 52 km SSE, E
bank Rio Xingu): Amazonas (Lago Meduinim. left
bank Rio Negro; Barro Vermelho. left bank Rio
Jurua: Ilha Paxiuba. right bank Rio Jurua: Colo-
cacao Vira-Volta, left bank Rio Jurua; Lago Vai-
Quem-Quer. right bank Rio Jurua; Ilhazinha. left
bank Rio Jurua: BOLIVIA: La Paz (La Reserva).
Voss et al.. 2001:46 BRAZIL: Amazonas (Boca
Rio Piratucu; 80 km N of Manaus); Para (Ilha do
Taiuna, lower Rio Tocantins); FRENCH GUIANA
(Paracou: Arataye); GUYANA: Demerara-Mahai-
ca (Hyde Park): Upper Takutu-Upper Essequibo
(Karamambo).
Marmosa pan-idens panidens: Pine. 1981:60 CO-
LOMBIA: Antioquia (Villa Arteaga, Uraba; Val-
divia: Quebrada Valdiva: La Cabana); Caldas
(Samana): Huila (Rio Aguas Claras, Acevedo);
Boyacd (Muzo): PERU: Cusco (Hacienda Villa
Carmen. Cosnipata): VENEZUELA: Carabobo-
Falcon-Yaracuy (Urama): Amazonas (Capibara,
Brazo Casiquiare): SURINAME: Nickerie (Kais-
erberg Airstrip. Zuid River): Brokopondo (Finisan-
ti): GUYANA: East Demerara (Hyde Park. De-
merara River).
M[armosa] panidentata: Tate, 1933:44 incorrect
subsequent spelling for panidens.
Marmosa panidens woodalli Pine, 1981:62 BRA-
ZIL: Para (type locality, Utinga, Belem: holotype,
adult female, skin and skull. USNM 393532, col-
lected 14 June 1968 by R. H. Pine). Gardner. 1993:
20 listed.
Holotype Skin and skull. FMNH 18545, col-
lected 8 September 1906 by S. B. Warren.
Type Locality Hyde Park, 30 mi up the De-
merara River, Demerara. Guyana, altitude 2 m.
84
FIELDIANA: ZOOLOGY
Fig. 50. Distribution of Marmosops parvidens: throughout northern South America south through Bolivia, with
scattered populations in Brazil. = localities given in published records.
Distribution (Fig. 50) Colombia south to Bo-
livia, Venezuela, Guyana, Suriname, French Gui-
ana, and scattered populations in Amazonas and
Para, Brazil.
Type Locality Teresopolis, in the Serra dos
Orgaos, Rio de Janeiro, Brazil.
Distribution (Fig. 51) Coastal mountains of
southeastern Brazil above 800 m at approximately
20-25S.
Marmosops paulensis (Tate, 1931)
Marmoset incana paulensis Tate, 1931:8 BRAZIL:
Rio de Janeiro (Teresopolis); Sao Paulo. Tate,
1933:166 BRAZIL: Rio de Janeiro (Teresopolis;
Rio de Janeiro); Sao Paulo (Sumidouro, near Rio
de Janeiro); Parana (Roca Nova). Gardner, 1993:
19 listed.
Marmosops paulensis: Mustrangi and Patton, 1997:
21,46 BRAZIL: Minus Gerais (Parque Nacional
Caparao, Serra do Caparao; Itanhandu; Passa Qua-
tro); Rio de Janeiro (Bonsucesso, Serra dos Or-
gaos; Teresopolis, Serra dos Orgaos); Sao Paulo
(Estacao Biologica de Boraceia, Serra do Mar; Fa-
zenda Intervales, Serra de Paranapiacaba); Parana
(Roca Nova).
Holotype Adult male, skin and skull, FMNH
26576, collected 8 July 1926 by C. C. Sanborn.
Marmosops pinheiroi (Pine, 1981)
Marmosops parvidens: Tate, 1939 VENEZUELA:
Bolivar (Auyan-tepuO- Handley and Gorden,
1979 VENEZUELA: Bolivar (Auyan-tepuO;
BRAZIL: Amapa (Serra do Navio).
Marmosops parvidens pinheiroi Pine, 1981:61, 63
BRAZIL: Amapa (Rio Amapari, Serra do Navio);
VENEZUELA: Bolivar (km 121, km 125, 85 km
SSE of El Dorado; Auyan-tepuO. Gardner, 1993:
20 listed.
Marmosops pinheiroi: Voss, 2001:49 BRAZIL:
Amapa (Serra do Navio); Para (Belem; Utinga);
FRENCH GUIANA (Paracou); GUYANA: Potaro-
Siparuni (Iwokrama Reserve); VENEZUELA: Bo-
livar (Auyan-tepui; Churi-tepui; km 121, 85 km SE
of El Dorado).
Holotype Adult female, skin and skull,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
85
20-
30
-10
50
Marmosops paulensis
500 1000 1500 km
90 80 70 60 50 40 30 20 10
Fig. 51. Distribution of Marmosops paulensis: southeastern Brazil. = localities given in published records.
USNM 461459, collected 8 May 1969 by the In-
stitute Evandro Chagas.
Type Locality Rio Amapari, Serra do Navio.
Brazil.
Distribution (Fig. 52) Eastern Venezuela.
Guyana, northern French Guiana, and Amapa,
Brazil.
Genus Metachirus Burmeister, 1854
Metachirus Burmeister, 1854:vii, 135 Subgenus of
Didelphys [sic = Didelphis Linnaeus]: included
species, myosurus [sic] Temminck [not myosuros
Temminck = Didelphis nudicaudatus E. Geoffroy
as a synonym], quica [= Philander opossum quica
Temminck], cinerea [= Micoureus demerarae
Thomas], incana [= Marmosops incana Lund],
murina [= Marmosa murina], agilis [= Gracili-
nanus agilis], pusilla [= Thylamys pusilla], brach-
yura [= Monodelphis brachyura], tristriata [-
Monodelphis americana], velutinus [= Thylamys
velutinus]. Burmeister. 1856:68 included species,
opossum [- Philander opossum], myosurus [sic]
not Temminck [nudicaudatus E. Geoffroy a syno-
nym], quica Temminck [= Philander opossum qui-
ca Temminck]. Thomas, 1888b:329 part, subge-
nus of Didelphys; included species, D. nudicau-
datus E. Geoffroy, designated type, crassicaudatus
Desmarest [= Lutreolina crassicaudata], opossum
Linnaeus [= Philander opossum opossum]. Mat-
schie, 1916:260. 267. 268 taxonomy; type, myo-
surus [sic] Temminck (not Temminck). Cabrera,
1919:45 classification; type, "Didelphis myosu-
rus Temminck [not Temminck], = D. nudicaudatus
Geoffroy." Miranda Ribeiro. 1936:320-321, 343
taxonomic history. Krumbiegel, 194 lb: 194 re-
view. Gardner. 1993:20 species (nudicaudatus
Desmarest. 1817); synonymy; distribution.
Philander Pine (not Tiedemann, 1808), 1973a: 398
regarded as correct generic name for four-eyed
pouchless opossum "'presently known as Metachi-
rus nudicaudatus." Hall, 1981:317 name adopted
after Pine, 1973a.
Type Species Didelphis nudicaudatus E.
Geoffroy (by subsequent designation [Thomas,
1888, p. 329]). Didelphis myosuros Temminck,
1824-1827, given as type species by Gardner
(1993, p. 20) is the four-eyed pouched opossum,
Philander opossum.
Distribution Nicaragua south to northern Ar-
gentina.
86
FIELDIANA: ZOOLOGY
Fig. 52. Distribution of Marmosops pinheiroi: eastern Venezuela, Guyana. French Guiana, and Amapa, Brazil.
localities given in published records.
Metachirus nudicaudatus (E. Geoffroy, 1803)
Didelphis nudicaudatus E. Geoffroy, 1803:142. Des-
marest, 1817:424 FRENCH GUIANA: (Cay-
enne). Carvalho, 1969:217 BRAZIL: Para (Be-
lem; km 87-94, Belem-Brasflia; Sao Miguel do
Guama [= Guama]).
Didelphys nudicaudatus: Desmarest, 1827:390 part,
myosuros (not Temminck) a synonym. Jentinck,
1888:220 part, BRAZIL: "syntype a" (adult,
mounted), "syntype b" (adult male, mounted), "c"
(adult male, mounted). Waterhouse, 1841:94
BRAZIL: Bahia (Bahia). Waterhouse, 1846:482
part, BRAZIL; FRENCH GUIANA; not Didelphis
myosuros in synonymy. Thomas, 1888b:332 part.
BRAZIL: Bahia (Bahia). Lydekker, 1896:201
part, distribution.
Metachirus nudicaudatus: J. A. Allen, 1897b: 119
PERU: San Martin (type locality, Guayabamba [=
Santa Rosa de Huayabamba], 5500 ft [= 1700 m];
holotype, female, skin and skull, AMNH no.
11815/10107, collected 30 August 1894 by O. T.
Baron). Bangs, 1900:87 COLOMBIA: Magdale-
na (Palomino, Sierra Nevada de Santa Marta).
Thomas, 1901a: 153 GUYANA: Upper Takutu-
Upper Essequibo (Kanuku Mts., 600 ft [= 180 m]).
J. A. Allen, 1904c:337 VENEZUELA: Bolivar
(Maripa). Ihering, 1906:1414 BRAZIL: Mato
Grosso (Rio Jauru). Thomas, 1910b: 188 GUYA-
NA: Pomeroon-Supenaam (Supenaam River). Mi-
randa Ribeiro, 1914:48 BRAZIL: Rondonia
(Mattas do Piroculuina [= Piroculuina]); Mato
Grosso (Chapadao [= Chapada] dos Parecis). J. A.
Allen, 1 91 6b:562 BRAZIL: Mato Grosso (Tapi-
rapoan [= Tapirapua], Rio Sepotuba). Pohle, 1927:
242 BRAZIL: Pemambuco (Dois Irmaos); Rio de
Janeiro (Teresopolis; Barreira). Thomas. 1928b:
294 PERU: Loreto (Pebas, "like specimens from
Para and Pernambuco"). Miranda Ribeiro, 1935:
37 BRAZIL: Rio de Janeiro (Teresopolis). Tate,
1939:162 VENEZUELA: Amazonas (foot of Cer-
ro Duida, 300 ft [= 90 m]); Bolivar (Auyan-tepuf,
1500-3500 ft [= 470-1090 m]); GUYANA: East
Demerara (Kartabo [now in Mazaruni-Potaro]).
Miranda Ribeiro. 1940:35 BRAZIL: Mato Gros-
so (Rio Jauru). Vieira. 1955:345 BRAZIL: Mato
Grosso (Rio Jauru). Olivares, 1962:307 COLOM-
BIA: Meta (Rio Guayabero). Pirlot, 1963:175, 178,
180 VENEZUELA: Zulia (Tukuko; Kasmera, Rfo
Yasa). Pine et al., 1970:668 BRAZIL: Mato
Grosso (Xavantina [= Chavantina], 264 km N of
Serra do Roncador). Yunis et al., 1973:369 CO-
LOMBIA: Arauca (Puerto Gaitan). Reig et al.,
1977:197 part, PERU: Ayacucho (Hacienda Lu-
isiana. Rio Apurimac; San Jose, Rfo Santa Rosa);
Ucayali (Balta); part, VENEZUELA: Trujillo (Mo-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
87
tatan); Merida (Bejuquero. Zea). Redford and Fon-
seca, 1986:129. 132 BRAZIL: Distrito Federal
(cerrado region, gallery forest dependent). Fonseca
and Kierulff. 1988:1 16 BRAZIL: Minas Gerais
(Fazenda Esmeralda: Fazenda Montes Claros: Rio
Doce State Park [= Parque Estadual do Rio Doce]).
Stallings. 1988:174. 187. 194 BRAZIL: Minas
Gerais (Rio Doce State Park [= Parque Estadual
do Rio Doce]). Gardner. 1988: 690 VENEZUE-
LA: Amazonas (Camp II. Cerro de La Neblina:
Camp VII. Cerro de La Neblina). Husband et al..
1992:298 BRAZIL: Sergipe (Crasto). Pacheco et
al.. 1993:5 PERU: Madre de Dios (Pakitza: Rio
Manu. lower: Cocha Cashu). Emmons and Romo.
1994: 140 PERU: Madre de Dios (Ccolpa de Gua-
camayos). Emmons et al.. 1994:14-1 PERU: Ma-
dre de Dios (Explorer's Inn Reserve). Rageot and
Albuja. 1994:176 ECUADOR: Pastaza (Mera).
Sarton et al.. 1996:307 BRAZIL: Rio de Janeiro
(Barra de Marico). Patton et al.. 2000:63 BRA-
ZIL: Acre (Igarape Porongaba. right bank Rio Ju-
rua; Igarape Porongaba. left bank Rio Jurua: Flora
[= Fazenda Santa Fe]. left bank Rio Jurua: Oci-
dente. opposite, left bank Rio Jurua: Nova Vida:
Sobral): Espirito Santo (Estacao Biologica de Santa
Lucia: Aracruz. Santa Cruz): Sao Paulo (Fazenda
Intervales. Base Saibadela. Sete Barros): Rio de Ja-
neiro (Ciadade Mineradoras Brasileiras Reunidas.
Ibicui. Mangaratiba): Bahia (Fazenda Beijo
Grande. 12 km S. 1.1 km E of Itabuna): Amazonas
(Barro Vermelho. left bank Rio Jurua: Lago Vai-
Quem-Quer. right bank Rio Jurua: Colocacao Vira-
Volta. left bank Rio Jurua: Alto Rio Urucu: Tefe:
Seringal Condor, left bank Rio Jurua: Rio Jaii; Ma-
caco, left bank Rio Jau: Tambor. left bank Rio Jaii:
above mouth, right bank Rio Jaii: Lago Meduinfm.
left bank Rio Negro: Penedo: Altamira: Ilha Pax-
iuba; Para (Floresta Nacional Tapirape-Aquiri.
Municipio de Maraba). PERU: Amazonas (Huam-
pami. Rio Cenepa). FRENCH GUIANA: (Sinna-
mary River). Mares and Braun. 2000:34 AR-
GENTINA: Misiones (Parque Nacional Iguazii: Rio
Urugua-f). Voss et al.. 2001:51 FRENCH GUI-
ANA (Paracou: Cayenne).
M[etachirus] nudicaudatus: J. A. Allen. 1900b: 197
(in text).
[Metachirus] nudicaudatus: Krumbiegel. 1941b:
198 BRAZIL: Paid (Ipitinga. Rio Acara).
Didelphis nudicaudata: Lesson. 1827:212 incorrect
subsequent spelling classification.
Didelphys nudicaudata: Thomas. 1 882a: 111 incor-
rect subsequent spelling PERU: Amazonas (Chir-
imoto). Thomas. 1888b:332 part. PERU: Ama-
zonas (Chirimoto).
Metachirus nudicaudata: Bertoni. 1914:69 incorrect
subsequent spelling PARAGUAY: Alto Parana
(Puerto Bertoni).
Lutreolina nudicaudata: Bertoni. 1939:6 incorrect
subsequent spelling PARAGUAY: Alto Parana
(Puerto Bertoni).
Philander nudicaudus: Gray. 1843:100 incorrect
subsequent spelling part, not synonymy. Miles et
al.. 1981a:331 BRAZIL: Para (Belem). Guerrero.
1985:40 VENEZUELA.
M[etachirus nudicaudatus] nudicaudatus: Cabrera.
1 9 1 9:46 classification.
Metachirus nudicaudatus nudicaudatus: Anthony.
1 92 lb:268 GUYANA: East Demerara (Kartabo
[now in Mazaruni-Potaro]). Miranda Ribeiro. 1936:
344, 345 part. GUIANAS: BRAZIL: Para (speci-
mens collected by E. Snethlage). Krumbiegel.
194 lb: 198 BRAZIL: Para (Ipitinga). Anduze.
1956:14 VENEZUELA: Territorio Federal Delta
Amacuro. Sanderson. 1949:787 SURINAME:
Para (Zanderij Airfield). Husson, 1973:6 SURI-
NAME. Husson. 1978:28 SURINAME: Para (Re-
publiek: Zanderij: Zanderij Airfield); Commewijne
(Coropina Kreek). Perez-Hernandez. 1989:371
VENEZUELA: Amazonas (Alto Rio Cacuri. Alto
Rio Ventuari; Tamatama: San Felipe [= Raudalito],
Rio Sipapo. not located: Cano Yureba. Salto Yureba:
Capibara; Campamento La Neblina [= Cerro La
Neblina]): Bolivar (Campamento): Delta Amacuro
Territorio Federal (Los Giiires: Tobesobe).
Metachirus n[udicaudatus] nudicaudatus: Carvalho
and Toccheton (not E. Geoffroy). 1969:217 BRA-
ZIL: Para (Guama: km 87-94 Belem-Brasflia).
Metachirus nudicaudatus nudicaudatus: Cabrera.
1919:46 incorrect subsequent spelling classifi-
cation.
D[idelph\s]s mxosuros: Wied-Neuwied (not Tem-
minck". 1824). 1826:400 BRAZIL: Bahia ("Co-
mechatiba" [= Comoxatiba = Ponta Cumuruxati-
ba]).
D[idelphis] mxosuros: Wagner (not Temminck. 1824).
1843:43 BRAZIL: part (description ex Wied-
Neuwied. 1826). Pelzeln. 1883:10 BRAZIL: Sao
Paulo (Ipanema): Rio de Janeiro (Registo do Sai).
M[etachirus] n[udicaudatus] mxosuros: Thomas (not
Temminck. 1824). 1923c:606 classification.
Didelphys mxosurus Burmeister (not Temminck),
1854:135 incorrect subsequent spelling BRA-
ZIL.
Metachirus mxosurus: Burmeister (not Temminck.
1824). 1856:69 incorrect subsequent spelling
BRAZIL: (specimen believed to be from Para).
Ihering. 1894:1 1 BRAZIL: Sao Paulo.
Didelphis mxosurus: Husson (not Temminck. 1824).
1978:29 incorrect subsequent spelling 2 "syn-
types" in Leiden Museum, skulls broken.
Metachirus nudicaudatus mxosurus: Pohle (not Tem-
minck. 1824), 1927:242 incorrect subsequent
spelling BRAZIL: Rio de Janeiro (Rio de Janei-
ro: Barreira): Sao Paulo ("type" locality restricted
to Ipanema: specimen designated "lectotype." skin
and skull. Vienna Museum. 1 16a. collected 25 Jan-
uary 1822 by Johann Natterer). Bresslau. 1927:
190 BRAZIL: Rio de Janeiro. Miranda Ribeiro.
1936:345. 351 BRAZIL: Bahia ("type" locality
restricted to Bahia): Espirito Santo (Lagoa de Ju-
parana): Rio de Janeiro (type locality. Realango.
Serra da Piraqua: Angra dos Reis: holotype. male,
skin and skeleton. MNRJ 1218). Vieira. 1949:
358 BRAZIL: Sao Paulo; Santa Catarina. Vieira.
1955:346 part. BRAZIL: Espirito Santo (Lagoa
de Juparana): Rio de Janeiro: Sao Paulo (Ipanema;
Bauni: Lins): Santa Catarina (Ararangua). Ca-
brera. 1958:38 personatus Miranda Ribeiro a
synonym. Carvalho. 1965:251 BRAZIL: Sao
Paulo (Boraceia). Ruschi. 1965:2 BRAZIL: Es-
pirito Santo (Rio Doce: Linhares; Lagoa Juparana).
M[etachirus] n[udicaudatus] mxosurus: Miranda Ri-
88
FIELDIANA: ZOOLOGY
beiro (not Temminck, 1824), 1936:352, footnote
incorrect subsequent spelling BRAZIL: Espirito
Santo (Lagoa de Juparana).
Metachirus tschudii J. A. Allen, 1900b: 195. Goodwin,
1953:227 type history.
M[etachirus] tschudii: J. A. Allen, 1900c:412 (in
text) "of the M. opossum group [!]." Gardner,
1993:20 listed.
Metachirus nudicaudatus tschudii: Osgood, 1914:
149 PERU: San Martin (Moyobamba). Thomas,
1927a:373 PERU: Amazonas (San Nicolas, 4500
ft [= 1400 m]); San Martin (Yurac Yacu, 2500 ft
[= 780 m]). Thomas, 1927b:606 PERU: Hudnu-
co (Chinchavito, 3000 ft [= 930 m]; Tingo Mana,
3000 ft [= 930 m]).
Metachirus nudicaudatus colombianus J. A. Allen,
1900b: 196. J. A. Allen, 1904d:4 18 COLOM-
BIA: Magdalena (type locality, Don Amo, west-
ern slope Sierra Nevada de Santa Marta; holotype,
male, skin and skull, AMNH 15448; collected 20
April 1899 by H. H. Smith). J. A. Allen, 1901c:
412. Allen, 1916a:201 COLOMBIA: Cundina-
marca (Buena Vista, near Bogota). Cabrera (part,
not J. A. Allen), 1958:37 imbutus a synonym.
Goodwin, 1953:227 type history. Hernandez-
Camacho and Borrero, 1957:213 COLOMBIA:
Santander (San Vicente de Chucurf). Perez-Her-
nandez, 1989:371 VENEZUELA: Barinas (Bar-
initas, Rio Barragan, 440 m; La Lengueta; La Er-
ica; Reserva Forestal Ticoporo); Merida (Zea;
Mesa Bolivar; Pueblo Las Virtudes, 17 km from
Caja Seca, Caserio El Amparo); Trujillo (Motatan;
Agua Viva; Valera); Zulia (Alguacil; Kasmera,
Rio Yasa; Novito; Tocuco [= Tukuko]; Puerto Ca-
tatumbo; Machiques). Gardner, 1993:20 listed.
M[etachirus] nudicaudatus colombica: J. A. Allen,
1901:412 incorrect subsequent spelling CO-
LOMBIA: Cundinamarca (near Bogota).
Metachirus nudicaudatus bolivianus J. A. Allen,
1901c:41 1 BOLIVIA: La Paz (type locality, Chu-
lumani, Yungas, 2000 ft [= 625 m], 2000 m; ho-
lotype, female, skin and skull BMNH 1.6.7.73, col-
lected 14 February 1901 by P. O. Simons). Thomas,
1902a: 143 BOLIVIA: Cochabamba (Charuplaya,
1350 m). Krumbiegel, 194 lb: 196 part, BOLIV-
IA: Santa Cruz (Santa Cruz de la Sierra; San Jose).
Cabrera, 1958:37 listed. Cabrera, 1958:37 in-
fuscus Thomas a synonym. Anderson et al., 1993:
18 BOLIVIA: Santa Cruz (Rfo Surutu). Gardner,
1993:20 listed. Anderson, 1997:155 BOLIVIA:
Beni (El Trapiche); Cochabamba (Charuplaya; Vil-
la Tunari; Yungas; Villa Tunari, 12.5 km SW
Parque Nacional Carrasco; San Rafael); La Paz
(Mapiri, Rio Chimate; Guanay, 5 km SE of Rio
Challana; Serrania Bellavista; Alcoche, 4 km NW;
Caranavi, 20 km NNE; La Reserva; Chijchipani;
Yolosito; Chulumani; Irupana); Pando (Rio Nareu-
da); Santa Cruz (Rio Suturu); Cochabamba-Santa
Cruz (Rio Ichilo, 9 km S of mouth of Rio Chapare;
Rio Ichilo, 52 km from mouth of Rio Chapare).
[Metachirus nudicaudatus] bolivianus: Krumbiegel
(part, not J. A. Allen, 1901), 194 lb: 196 ARGEN-
TINA: Formosa (La Urbana [= Estancia Tapiki-
ole], Rio Pilcomayo).
Metachirus nudicaudatus phaerus Thomas, 190 Id:
545 ECUADOR: Esmeraldas (type locality, San
Javier, Rio Cachavi, 20 m; holotype, male, skin and
skull, BMNH 1.3.19.44, collected 19 May 1900 by
G. Fleming). Gardner, 1993:20 listed.
Metachirus nudicaudatus dentaneus Goldman, 1912:
2 PANAMA: Colon (type locality, Gatiin; holo-
type, male, skin and skull, USNM 172732, collect-
ed 12 January 1911 by E. A. Goldman). Enders,
1935:41 1 PANAMA: Colon/Panama (Barro Col-
orado). Goodwin, 1942:1 13 NICARAGUA:
(Chontales). Hall and Kelson, 1959:19 map 10
(Middle American distribution). Gardner, 1993:
20 listed.
Philander nudicaudatus dentaneus: Hall, 1981:18
map 1 1 (Central American range); classification;
distribution.
Metachirus nudicaudatus antioquiae J. A. Allen,
19 16c:83 COLOMBIA: Antioquia (type locality,
La Frijolera; holotype, male, skin and skull,
AMNH 37751, collected 1 January 1915 by L. E.
Miller and H. S. Boyle). Goodwin, 1953:227 type
history. Gardner, 1993:20 listed.
Metachirus nudicaudatus modestus Thomas, 1923c:
606 PARAGUAY: Paraguari (type locality, Sap-
ucay [= Sapucaf]; holotype, female, skin and skull,
BMNH 1903.2.3.38, collected 2 September 1902
by W. Foster). Crespo, 1950:7 ARGENTINA:
Misiones (Rio Urugua-i, 30 km from Puerto Bem-
berg; Puerto Bemberg). Gardner, 1993:20 listed.
Metachirus nudicaudatus infuscus Thomas, 1923c:
606 PERU: Madre de Dios (type locality, Rio In-
ambari; holotype, subadult male, skin and skull,
BMNH 2.7.27.8, collected by G. Ockenden).
Thomas, 1928b:285 PERU: Ucayali (Yarinaco-
cha); Loreto (Contamana; Pebas). Sanborn, 1949:
277 PERU: Ucayali (Yarinacocha); Loreto (Cerro
Azul). Gardner, 1993:20 listed.
Metachirus nudicaudatus imbutus Thomas, 1923c:
605 ECUADOR: Pichincha (type locality, Min-
do, W slope Cordillera Occidental; ca. 4200 ft | =
1300 m]; holotype, male, skin and skull, BMNH
13.10.24.71, collected 12 June 1913 by Gilbert
Hammond). Gardner, 1993:20 listed.
Metachirus nudicaudatus personatus Miranda Ribei-
ro, 1936:344, 351, 352, footnote BRAZIL: Espir-
ito Santo (Lagoa de Juparana); Rio de Janeiro (Re-
gisto do Sai, Natterer specimen in Vienna Muse-
um); Guanabara (Realango, Serra da Piraquara
[now in Rio de Janeiro]). Miranda Ribeiro, 1955:
416 "type" history. Avila Pires, 1968:168
"type" history. Gardner, 1993:20 listed.
Holotype Adult female, skin mounted with
young attached to teats, skull separate, MNHN,
Paris, original catalog no. 302, exhibition hall cat-
alog no. 434; collected by H. Martin, then French
government botanist in Cayenne. Recataloged as
a C G 1990-420 (see Julien-Laferriere, 1994,
Mammalia 58[1]:3).
Type Locality Cayenne, French Guiana.
Distribution (Fig. 53) Honduras to Paraguay
and northern Argentina.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
89
20
10
10'
20
30
40
50^
Metachirus nudicaudatus \
o<P
.CD
>
0)8
'
o o
c*
%
i~. 500 1000 1500 km
90
80 ;
70 r
60
50
40
30 20 10
Fig. 53. Distribution of Metachirus nudicaudatus: from Honduras in Central America south to Misiones. Argen-
tina, and Rio Grande do Sul. Brazil, o = localities from unpublished records: = localities given in published
records.
90
FIELDIANA: ZOOLOGY
Additional Records NICARAGUA: Zelaya;
El Pool, Bluefields (Myers, 1967, MVZ); COSTA
RICA: San Jose; Angostura (Goldman, 1876,
USNM); PANAMA: Bocas del Toro; Changuin-
ola, 7 km SSW (Handley, Jr., 1960, USNM); Pen-
insula Valiente, Punta Alegre (Greenwell, 1990,
USNM); Nun (Greenwell, 1993, USNM); Code;
El Valle, 6 mi E (Enders, 1957, USNM); Colon;
Fort Sherman (1955, USNM); Cristobal (Fleming,
1967, USNM); Camp Pina, Fort Sherman (Keen-
an, 1956, USNM); Fort Davis (1955, USNM);
Bohio Peninsula (Pine, 1972, USNM); Buena Vis-
ta Peninsula (Pine, 1972, USNM); Panama; Bal-
boa (Fleming, 1967, USNM); Fort Kobbe (1955,
USNM); Rodman Naval Ammunition Depot
(Fleming, 1967, UMMZ); Cerro Azul (1955,
USNM); La Zumbadora [= Cerro Azul] (1957,
USNM; Keenan, 1955, 1956, 1957, USNM);
Maxon Ranch (Anthony, 1914, AMNH; Richard-
son, 1914, AMNH); Cerro Campana (Field, 1954,
USNM); San Bias; Mandinga (Handley, Jr., 1957,
USNM); Armila, Quebrada Venado (Handley, Jr.,
1963, USNM); Darien; Tacarcuna (Village) [=
Tacarcuna] (Anthony, 1915, AMNH); Tacarcuna
Village Camp [= Tacarcuna] (Handley, Jr., 1959,
USNM; Handley, Jr., 1963, UMMZ); Paya Camp
(Mendez, 1959, USNM); Cana (Goldman, 1912,
USNM); Rio Jaque (Wetmore, 1947, USNM);
Jaque (Greenwell, 1966, USNM); COLOMBIA:
Choco; Rio Sando, Rio Baudo (von Sneidern,
1958, FMNH); Valle del Cauca; Buenaventura, 6
km N (Emmons, 1981, USNM); Cordoba; Socor-
re, Rio Sinu (Hershkovitz, 1949, FMNH); Antio-
quia; Puri, Rio Puri, Rio Cauca (Hershkovitz,
1950, FMNH); Tamara [= La Tirana], Rio Nechi
(Peterson, 1971, USNM); Caquetd; Montanita,
Florencia, left bank Rio Orteguaza (Hershkovitz,
1951, FMNH); Tres Troncos, La Tagua, Rio Ca-
queta (Hershkovitz, 1952, FMNH); Norte de San-
tander; Rio Tarra, Rio Catatumbo (Hershkovitz,
1943, USNM); San Calixto (Hershkovitz, 1943,
USNM); Boyacd; Rio Covaria [= Cobana] (von
Sneidern, 1959, FMNH); La Argentina, Rio Cu-
bugdn (von Sneidern, 1959, FMNH); Muzo
(Hershkovitz, 1952, FMNH); Cundinamarca;
Mambita, Rio Guavio (Gonzalez, 1922, AMNH);
Medina, eastern slope Andes (Niceforo Maria,
1925, AMNH); Meta; Buena Vista (Chapman,
1913, AMNH); Guaicaramo (Niceforo Maria,
1926, FMNH); Canay (1937, AMNH); Restrepo
(1937, AMNH); Upun [= Upin], Restrepo (1937,
AMNH); Los Micos, San Juan cte Arama, Rio
Giiejar (von Sneidern, 1957, FMNH); Villavicen-
cio (Niceforo Maria, 1939, AMNH); ECUADOR:
Esmeraldas; Rosa Zarate [= Quininde] (Rageot,
1980, USNM); Pinchincha; Pachahal [= Pacha-
tal], below Mindo (Soderstrom, 1913, AMNH);
Manabi; Rio Pescado (Tate, 1922, AMNH);
Guayas; Cerro Manglaralto (Tate, 1923, AMNH);
Napo; Santa Cecilia (Duellman, 1968, KU); Li-
moncocha (Voss, 1980, UMMZ); San Jose Nuevo
[= San Jose Abajo] (Olalla Brothers, 1924,
AMNH); Pastaza; Rio Yana Rumi (Olalla, 1934,
FMNH); Tiguino, 130 km S of Coca (Jacobs,
1990, USNM); Sarayacu, Rio Bobonaza (Tate,
1924, AMNH); Montalvo, Rio Bobonaza (Olalla,
1932, FMNH); Rio Pindo Yacu (Olalla, 1934,
FMNH); Rio Capihuara (Olalla, 1934, FMNH);
Rio Curaray, not precisely located (Olalla Broth-
ers, 1925, AMNH); Zamora-Chinchipe; Zamora
(Anthony, 1920, AMNH); Los Encuentros, 4 km
ENE, not precisely located (Gardner, 1976,
USNM); PERU: Cajamarca; Quebrada Huaran-
dosa, Rio Chinchipe (Watkins, 1923, AMNH);
Amazonas; La Poza, Rio Santiago (Patton, 1979,
MVZ); Pomara (Watkins, 1924, AMNH); Hudn-
uco; Hacienda Santa Elena [= Santa Elena], 35
km NE of Tingo Marfa (Tallman, 1972, LSUMZ);
Hacienda Exito, Rio Cayumba (Heller, 1922,
FMNH); Pozuzo [now in Pasco] (Heller, 1923,
FMNH); Pasco; Oxapampa [= San Juan de Ox-
apampa] (Tuttle, 1964, USNM); Junin; Chancha-
mayo (Schunke, 1904, FMNH); San Ramon (An-
derson, 1914, FMNH); Ayacucho; Huanhuachayo
(Gardner, 1971, LSUMZ); Cusco; Hacienda Eri-
ka, Rio Alto Madre de Dios (Myers, 1984,
UMMZ; Patton, 1984, MVZ); Kiteni, "60 km
from Rosalina on road from Quellabamba" (My-
ers, 1984, UMMZ; Patton, 1984, MVZ; Thomas,
1974, LSUMZ); Uvini [= Yuvini], Rio Cosireni
(Heller, 1915, USNM); Huaijumbe [= Huajyum-
be] (Kalinowski, 1954, FMNH); Quincemil (Kal-
inowski, 1953, FMNH); Madre de Dios; Tambo-
pata (Kuns, 1965, USNM); Puerto Maldonado
(Hibbs, 1965, USNM; Kuns, 1965, USNM); Al-
bergue Cuzco Amazonico, Puerto Maldonado, 12
km E (Warner, 1983, MVZ); Reserva Cuzco
Amazonico (Timm, 1990, KU; Woodman, 1989,
KU); Lago Sandoval, Rio Madre de Dios (Patton,
1979, MVZ); Loreto; Rio Curaray, boca (Olalla
Brothers, 1925, AMNH); Libertad (Cardiff, 1982,
LSUMZ); Santa Luisa, Rio Nanay (Kalinowski,
1956, FMNH); Puerto Indiana, N bank Rio Ma-
ranon (Olalla Brothers, 1926, AMNH); Iquitos,
Rfo Maranon (Bassler, 1924, 1928, AMNH); San
Fernando, left bank Rio Yavari (Kalinowski,
1957, FMNH); Rio Amaya, boca, W bank Rfo
Morona (Kalinowski, 1957, FMNH); Santa Elena,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
91
Rio Samiria (Kalinowski, 1956, 1957, FMNH);
Ucayali; Pucallpa, 57 km W (Hinsch. 1971, 1972.
USNM); Pucallpa, W bank Rio Ucayali (Hinsch.
1971, USNM); Rio Urubamba, boca (Olalla
Brothers, 1927, AMNH); VENEZUELA: Zulia;
Cerro Azul (1955, 1956, AMNH; 1956, USNM);
Maracaiba (Hershkovitz personal notes); Mision
Tukuko (Ayarzaguena, 1986, USNM; Colonnello,
1986, USNM; Voss, 1986, USNM); Merida; La
Azulita (Osgood and Conover, 1920. FMNH);
Barinas; Altamira (Smithsonian- Venezuelan Ex-
pedition. 1967, USNM); La Quinta (Tuttle, In-
quillo, Stromeyer. and Peterson. 1967. USNM);
Bolivar: Cinco Rancho, 38 km NE Icabani (Tut-
tle, 1968. USNM); Amazonas; Fish Creek, Cerro
Duida (Olalla Brothers, 1929, AMNH); Esmeral-
da, Cerro Duida (Olalla Brothers, 1928. AMNH);
GUYANA: Upper Takutu-Upper Essequibo;
Boundary Camp, Itabu Creek. Corentyne River
(Blake, 1938, FMNH); FRENCH GUIANA: Ini-
ni; River Arataye (Emmons. 1984, USNM):
BRAZIL: Amapd; Maraca. Serra do Navio
(Woodall. 1966, 1967. 1970. USNM); Para; Car-
atateua. E bank Rio Caete (Instituto Oswaldo
Cruz. 1968, USNM); Braganca (Instituto Oswaldo
Cruz, 1968. USNM); Igarape-Acu (Hershkovitz
specimen notes from Instituto Oswaldo Cruz);
Utinga (Instituto Oswaldo Cruz. 1968. USNM):
Murucutii, Instituto Agronomico do Norte (Lima.
1923, FMNH); Capim. 8 mi SE of Belem (Insti-
tuto Oswaldo Cruz, 1968); Lazaropolis, Belem.
not located (Hershkovitz, 1996. personal notes);
Mojui dos Campos (Instituto Oswaldo Cruz.
1975. USNM); Rio Mojui (Hershkovitz personal
notes): Ilha do Taiuna, Rio Tocantins (Olalla.
1931, AMNH); Amarim Igarape, Rio Tapajoz
(Olalla, 1931, AMNH); km 84, Santarem-Cuiaba
(Instituto Oswaldo Cruz, 1972; Instituto Evandro
Chavez. 1972, USNM); Baiao. Rio Tocantins
(Olalla. 1931. AMNH); Altamira, W bank Rio
Xingu (de Paula Pinheiro, Instituto Oswaldo Cruz.
1972, USNM); km 43, Altamira-Itaituba (de Paula
Pinheiro, Instituto Oswaldo Cruz, 1972); Fordlan-
dia, E bank Rio Tapajoz (Olalla. 1960. FMNH):
Itaituba (Hershkovitz field notes); km 19. Itaituba-
Jacareacanga [= Jacare-Acanga] (Ramos. Institu-
to Oswaldo Cruz, 1972, USNM: de Paula Pinhei-
ro, Instituto Oswaldo Cruz, 1972, USNM): San-
tarem (Hershkovitz personal notes); Rio Acara
(Hershkovitz personal notes); Jacareacanga [= Ja-
care-Acanga], W bank Rio Tapajos (Ramos. In-
stituto Oswaldo Cruz. 1972. USNM: Hershkovitz
personal notes); Amazonas; Manaus (Malcolm.
1985, USNM); km 46, Manaus-Itacoatiara
(Hershkovitz field notes); Bahia; Rio do Braco
(1944, KU); Fazenda Almada (Rockefeller Insti-
tute, 1945, FMNH); Fazenda Almada, Ilheus (Ser-
vicio do Estudos e Pesquisas sobre a Febra Amar-
eia [SEPSFA], 1945, FMNH); Urucutuca, Ilheus
(Servicio do Estudos e Pesquisas sobre a Febra
Amareia [SEPSFA]; Riberao da Fortuna [Servicio
do Estudos e Pesquisas sobre a Febra Amareia
(SEPSFA)]. 1944. FMNH); Espirito Santo; Frag-
mentos Florestals da Cai, Aracruz Calulore, Ara-
cruz (Mustrangi, 1993, MVZ); Rio de Janeiro;
Gavea (Gilmore, 1937, AMNH); Floresta da Ti-
juca (1944, 1945, KU); Sao Paulo; Rocha (Olalla,
1961, FMNH): Fazenda Capricornio, Ubatuba
(Lara, 1992, MVZ); Varjao, m, 5 km S of Casa
Grande (Souza Lopes, 1969, 1970, USNM);
Guaratuba (Souza Lopes, 1969, 1970, USNM);
Itanhaem (Souza Lopes, USNM); Boa Vista, Rio
Iporanga (Olalla. 1960. FMNH); Mato Grosso;
Aripuana (1974, 1975, USNM); Mato Grosso do
Sul; Urucum de Corumba (Hershkovitz field
notes): Dourados, 12 km N (Kuns, 1965, USNM);
Rondonia; Pista Nova [= Nueva], 8 km N of Por-
to Velho (Kuns. 1965, USNM); BOLIVIA: La
Paz; Rio Chimate, Rio Mapiri (Tate, 1926,
AMNH); Rio Challana, SE Guanay (Creighton,
1978, UMMZ); PARAGUAY: San Pedro; Aca
Poi (Wharton, 1950, USNM); Tacuati, Aca Poi, S
bank Rio Ypani (Wharton, 1950, USNM); Caa-
guazii; Caaguazii. Summerfield Colony no. 1 1
(Wharton, 1953, USNM); Paraguari; Parque Na-
cional Ybycuf (Myers, 1978, UMMZ); Canendi-
yu; Curuguaty (Myers, 1978, UMMZ).
Genus Micoureus Lesson, 1842
Micoureus Lesson. 1842:186 included species, ci-
nereus Temminck. dorsiguera Linnaeus, murina
Linnaeus, tricolor [= Monodelphis tricolor E.
Geoffroy], lanigera [= Philander lanata Olfers],
elegans Waterhouse. californicus [= Didelphis vir-
giniana Kerr], breviceps [= Didelphis virginiana
Kerr]. Thomas. 1888b:340 Didelphis cinerea
Temminck designated type species. Gardner. 1993:
20 listed.
Micoreus Vieira. 1955:350 incorrect subsequent
spelling of Micoureus Lesson in synonymy of Mar-
mosa Gray.
Micoures Reig. 1958:272 (footnote) incorrect sub-
sequent spelling of Micoureus Lesson.
Type Species Didelphis cinerea Temminck,
1824, preoccupied (= Micoureus demerarae,
Thomas, 1905), by subsequent designation
(Thomas. 1888b).
92
FIELDIANA: ZOOLOGY
Distribution (Fig. 54) Neotropical region
from Central America and adjacent Caribbean is-
lands to northern Argentina, Paraguay and Brazil.
Comments Previously included with Marmo-
set (sensu lato, see Gardner and Creighton, 1989:
3).
Micoureus alstoni (J. A. Allen, 1900)
Didelphys cinerea: Alston, 1882: 199 COSTA
RICA.
Didelphys {Micoureus) cinerea: J. A. Allen, 1891:
218 COSTA RICA: (San Jose).
Marmosa cinerea: J. A. Allen, 1897a:43 COSTA
RICA: (type locality; holotype, male, skin and
skull, AMNH 11790/16210).
[l]Marmosa cinerea: Beach, 1939:315 ECUADOR.
Caluromys alstoni J. A. Allen, 1900a: 189. Gardner,
1983:20 listed.
[Marmosa] alstoni: Thomas, 1901e: 145 classifica-
tion.
M[armosa] c[inerea] Alstoni: Thomas, 1905:313
(footnote) classification.
Micoureus alstoni: Reig et al., 1977:212 COSTA
RICA: San Jose (Santa Ana). Timm et al., 1989
COSTA RICA: Heredia (Parque Nacional Braulio
Carrillo; Concepcion de San Rafael). Patton et al.,
2000:68 PANAMA: Bocas del Tow (Boca Torito,
Isla San Cristobal).
Marmosa alstoni alstoni: Tate, 1933:67, 68 NICA-
RAGUA; HONDURAS; COSTA RICA: Limon
(Cuba); COLOMBIA: Antioquia (Jerico); Quindio
(Calarca); Cundinamarca (Bogota). Goodwin,
1942:115 HONDURAS: (Segovia River [= Rio
Segovia]). Goodwin, 1946:286-287 COSTA RICA:
San Jose (San Jose; Piedras Negras; Escazu); Li-
mon (Cubre [= Cuabre]); Cartago (Agua Caliente;
Cartago; Tres Ribs, Pozo Ancho; San Juan).
Marmosa cinerea nicaraguae Thomas, 1905:313
NICARAGUA: Zelaya (type locality, Bluefields;
holotype, adult male, BMNH 5.10.31.5, collected 9
January 1905 by G. Palmer, original no. 5).
Marmosa alstoni nicaraguae: Tate, 1933:69 NIC-
ARAGUA: Zelaya (Bluefields); COSTA RICA: Li-
mon (Siquirres, Rio Pacuare).
Marmosa demerarae meridae: Tate (part, not Tate),
1933:65 COLOMBIA: Cundinamarca ("S. [ =
Santo] Domingo de Bogota"); Boyacd ("Miraflo-
res").
Holotype Male, skin and skull, AMNH
11790/16210; collected 17 August 1893 by
George K. Cherrie.
Type Locality Tres Rfos, Cartago, Costa
Rica.
Distribution (Fig. 55) Central America, ad-
jacent Caribbean islands, and Colombia west of
the crest of the Cordillera Oriental.
Micoureus constantiae (Thomas, 1904)
Didelphis cinerea: Lund, 1841:237 BRAZIL: Minus
Gerais (Lagoa Santa).
Didelphys [(Micoureus)] cinerea: Thomas, 1888b:
342 part, BOLIVIA: Santa Cruz (Santa Cruz de
la Sierra).
Grymaeomys cinerea: Winge, 1893:46 BRAZIL:
Minas Gerais (Lagoa Santa).
Marmosa cinerea: Thomas, 1901c:536 BRAZIL:
Minas Gerais (Rio Jordao). Miranda Ribeiro, 1936:
361 part, BRAZIL: Minas Gerais (Lagoa Santa).
Marmosa constantiae Thomas, 1904c:243 BRAZIL:
(type locality). Gardner, 1993:20 listed. Heinonen
and Bosso, 1994:55 ARGENTINA: Jujuy (Parque
Nacional Calilegua, Caimancito).
[Caluromys] constantiae: Matschie, 1916:270 clas-
sification.
Marmosa constantiae constantiae: Tate, 1933:75
BRAZIL: Mato Grosso (Chapada [= Chapada dos
Guimaraes]); BOLIVIA: Santa Cruz (Buenavista;
Rfo Sara, near Buenavista; Cerro Hosana; Rio Su-
rutii). Vieira, 1955:351 BRAZIL: Mato Grosso
(Chapada [= Chapada dos Guimaraes]; Porto Faia,
not located); Mato Grosso do Sul (Campo Grande).
Micoreus constantiae constantiae: Anderson, 1997:
156 BOLIVIA: Beni (Cascajal, across river; Rio
Itenez, opposite Costa Marques; Villa Monte; El
Porvenir, 5 km N); Cochabamha (Yungas; Tablas
Monte, 9.4 km N; Tablas Monte, 4.4 km N); La
Paz (Campamento Moira, Alto Madidi; Ixiamas, 13
km W; Rio Machariapo; Puerto Linares; Palos
Blancos; La Reserva); Pando (La Cruz). Mares and
Braun, 2000:34 ARGENTINA: Jujuy (Altura de
Yuta, Rio San Francisco; Parque Nacional Calile-
gua); Salta (Oran; Parque Nacional Baritu).
Marmosa budini Thomas, 1920a: 195 ARGENTI-
NA: Jujuy (type locality, Altura de Yuto, 70 km N
of Villa Carolina, Rfo San Francisco, 500 m; ho-
lotype, male, skin and skull, BMNH 20.1.7.134,
collected 3 July 1919 by E. Budin). Gardner, 1993:
20 listed.
[Marmosa] budini: Krumbiegel, 1941c:6 BOLIVIA:
Santa Cruz (San Jose, Rfo San Miguel).
Marmosa constantiae budini: Tate, 1933:76 AR-
GENTINA: (holotype).
Marmosa constanciae budini: Olrog, 1959:404 in-
correct subsequent spelling ARGENTINA: Salta
(Oran |= San Ramon de la Nueva Oran|).
Marmosa constantiae budini: Anderson, 1997:156
BOLIVIA: Chuquisaca (Rfo Limon; Chuyayacu, 2
km E); Santa Cruz (Pajaral; Rfo Mamore, 2 km N
of mouth of Rfo Chapare; Rfo Ichilo, 54 km S of
mouth of Rfo Chapare; Yapacanf; Santa Rosa, 7 km
N; Buena Vista, 25 km W, W bank Rfo Yapacanf;
Rfo Surutu; Cerro Hosana; Buenavista; Warnes;
Santa Cruz de la Sierra; Santa Cruz, 27 km SE;
Chiquitos; Estancia Cuevas; Terebinto; Boyuibe, 53
km E); Tarija (Entre Rfos, 5 km NNW; Caraparf,
3 km WNW). Flores et al., 2000:324 ARGEN-
TINA: Jujuy (Aguas Negras; Aguas Negras, camp-
ing, Parque Nacional Calilegua; Caimancito; Cali-
legua; Mesada de las Colmenas; Rfo San Francisco,
altura de Yuto); Salta (43.7 km NW of junction of
Argentine Highways 50 and 1 8 on road to Isla de
BROWN: ATLAS OF NEW WORLD MARSUPIALS
93
Micoureus
90 80 70" 60 50 40 30 20 10
Fig. 54. Distribution of the genus Micoureus: Belize to southern Panama in Central America and from northern
South America to northern Argentina and southern Bolivia and Brazil.
94
FIELDIANA: ZOOLOGY
Fig. 55. Distribution of Micoureus alstoni: Central America and adjacent Caribbean islands south to western
Colombia. = localities given in published records.
Cafias; Oran; Parque Nacional Baritu, desembocad-
ura arroyo Santelmita).
Marmoset cinerea hudini: Anderson et al., 1993:17
BOLIVIA: Santa Cruz (Rio Surutu; Buenavista;
Warnes; Santa Cruz de la Sierra; Cerro Hosana,
Parque Nacional de Amboro).
Holotype Adult male, BMNH 3.7.7.157, col-
lected 27 August 1902 by A. Robert.
Type Locality Santa Anna de Chapada [=
Chapada dos Guimaraes], Mato Grosso, Brazil.
Distribution (Fig. 56) Bolivia south into
northern Argentina and Brazil.
Micoureus demerarae (Thomas, 1905)
Didelphis cinerea Temminck, 1824:46 BRAZIL: Ba-
hia (type locality, Morro de Arara, Rio Mucuri, near
border of Espfrito Santo [cf. Wied-Neuwied, 1 826:
406]; syntypes several specimens collected by
Prince Maximillian Wied-Neuwied, Johann Natterer
and Sellow [?]. Most, if not all, were originally pre-
served in the Vienna Museum. In 1870, the Ameri-
can Museum of Natural History purchased part of
the Wied-Neuwied collection, including one of the
male syntypes of cinerea. The syntype, a skin only,
is mounted, faded, and in poor condition. Its present
catalog number is 845. A second syntype, in the
Vienna Museum, is numbered 69A [cf. Pohle, 1927:
57; Goodwin, 1953:223]). Jentink, 1888:221
BRAZIL: one of syntypes received from Vienna
Museum. Gardner. 1993:20 listed.
D[idelph\s] cinerea: Wied-Neuwied (not Goldfuss,
1809)," 1826:406 BRAZIL: Bahia (Morro d' Arara
[= Morro de Arara], Rio Mucuri); name combi-
nation preoccupied by Didelphys cinerea Goldfuss
(1809:220 [= Trichosurus vulpecula Kerr]).
Didelphx.s cinerea: Waterhouse, 1841:103 BRAZIL.
Waterhouse, 1846:501 BRAZIL. Burmeister,
1854:137 BRAZIL. Pelzeln, 1883:1 14 BRA-
ZIL: Bahia. Goeldi and Hagmann, 1904:100
BRAZIL: Para.
\Marmosa] cinerea: Trouessart, 1898:1238 part,
classification. Thomas, 1901e: 145 classification.
Marmosa cinerea: Elliot, 1907:4 VENEZUELA.
Miranda Ribeiro, 1936:361 part, BRAZIL: Rio de
Janeiro (Angra dos Reis). Lyne, 1959:84 VE-
NEZUELA: Merida. Pine et al., 1970:668 BRA-
ZIL: Mato Grosso (Xavantina [= Chavantina]).
Handley, 1976:6 VENEZUELA: Falcon (Ura-
ma); Aragua (Estacibn Bioldgica de Rancho
Grande); Miranda (Curupao); Amazonas (Puerto
Ayacucho; San Juan; Belen; Cano Culebra, Cerro
BROWN: ATLAS OF NEW WORLD MARSUPIALS
95
90 80 70 60 50" 40 30 20 10
Fig. 56. Distribution of Micoureus constantiae: Bolivia, northern Argentina, and southern Brazil. = localities
given in published records.
96
FIELDIANA: ZOOLOGY
Duida; Capibara, Brazo Casiquiare); Bolivar (Hato
La Florida; Hato San Jose; km 125, 85 km SSE of
El Dorado). Reig et al., 1977:212 PERU: Ucayali
(Rio Curanja, Balta); VENEZUELA: Aragua (Ran-
cho Grande); Merida (Hacienda San Pedro, Santa
Cruz de Mora [= Santa Cruz]); Amazonas (Esmer-
alda), Cerro Duida); Bolivar (El Guri [= El Man-
teco]; Caicara del Orinoco [= Caicara]). Husson,
1978:14-15 GUYANA: East Demarara-West
Coast Berbice (Coomacka [= Takama]); SURI-
NAME: Coronie (Plantation "Mary's Hope, E of
Totness"); Sipaliwini (Raleigh Falls); Para (On-
verwacht); Marowijne (Langamankondre).
O'Connell, 1979:77 VENEZUELA: Miranda-
Guarico (Parque Nacional Guatopo). Eisenberg et
al., 1979:191 VENEZUELA: Miranda-Guarico
(Parque Nacional Guatopo). Mares et al., 1981:
101 BRAZIL: Bahia (Fazenda Almeida, Ilheus;
Banco da Vitoria, Pirataquisse, Ilheus; Aritagua
Urucutuca, Ilheus; Buerarema da Fortuna [= Buer-
arema Ribeirao da Fortuna, Ilheus]); Pernambuco
(Dois Irmaos). Terborgh et al., 1984:1 PERU:
Madre de Dios (Cocha Cashu Biological Station,
Parque Nacional del Manu). Emmons, 1984:1
PERU: Madre de Dios (Cocha Cashu Biological
Station, Parque Nacional del Manu). Stallings,
1988:175 BRAZIL: Minas Gerais (Rio Doce
State Park [= Parque Estadual do Rio Doce]). da
Fonseca and Kierulff, 1988:108 BRAZIL: Minas
Gerais (Rio Doce State Park [= Parque Estadual
do Rio Doce]). Gardner, 1988:696 VENEZUE-
LA: Amazonas (Base Camp [= Campamento];
Base Camp I, Cerro de La Neblina; Camp V, Cerro
de La Neblina; Camp VII, Cerro de La Neblina;
Camp XI, Cerro de La Neblina).
Marmosa (Micoureus) cinerea: Pohle, 1927:241
BRAZIL: Pernambuco (Dois Irmaos).
[l]Marmosa cinerea: Sanderson, 1949:788 SURI-
NAME.
Marmosa {Micoureus) cinerea subsp.: Pohle, 1927:
241 BRAZIL: Rio de Janeiro.
[Marmosa] cinerea cinerea: Cabrera, 1919:36 listed.
Marmosa cinerea cinerea: Tate, 1933:55 BRAZIL:
Espi'rito Santo-Bahia-Minas Gerais (Rio Mucuri,
probably near Morro de Arara); Pernambuco (Dois
Irmaos); Rio de Janeiro; Para (Marajo [= Ilha de
Marajo]). Vieira, 1955:351 BRAZIL: Para; Per-
nambuco; Espirito Santo (Rio Mucuri; Rio Sao
Jose); Rio de Janeiro (Angra dos Reis). Cabrera,
1958:13 classification. Carvalho, 1958:123
BRAZIL: Para (Gradaus [= Gorotire], Rio Fres-
co). Avila Pires, 1968:165 BRAZIL: Rio de Ja-
neiro (Angra dos Reis). Pine, 1973b:55 BRAZIL:
Para (Belem). Avila Pires and Gouvea, 1977:9
BRAZIL: Rio de Janeiro (Monte Serrat;
Macieiras).
Micoureus cinereus: Lesson, 1842:186. Emmons and
Romo, 1994:140 PERU: Madre de Dios (Cerros
del Tavara). Emmons, 1990:72 BOLIVIA: La Paz
(Ixiamas). Patterson, 1992:7 BRAZIL: Amazonas
(Codajaz); Para (Cameta).
Philander cinereus: Gray, 1843:101 South America.
Grymaeomys cinereus: Burmeister, 1856:78 classi-
fication.
Caluromys cinereus: J. A. Allen, 1900a: 189 classi-
fication.
Didelphis cinereus: Avila Pires, 1965:4 BRAZIL:
Bahia (type locality, Morro d' Arara [= Morro de
Arara], some 30 km W of mouth of Rio Mucuri;
holotype, male, skin, formerly mounted, AMNH
845).
Marmosa cinerea demerarae Thomas, 1905:313
GUYANA (type locality). Anthony, 1921:267
GUYANA: Essequibo (Kartabo |now in Cuvani-
Mazaruni]). Perez-Hernandez, 1989:364 VENE-
ZUELA: Bolivar (Campamento, Zona de la Repre-
sa del Guri, Rio Caroni; Alrededores de Caicara del
Orinoco, Orinoco Medio); Merida (Las Porqueras).
Gardner, 1993:20 listed.
Marmosa demararae demararae: Tate, 1933:62 in-
correct subsequent spelling GUYANA: Upper
Demerara-Berbice (Comaccka [= Takama]); De-
merara-Mahaica (Hyde Park); Cuyuni-Mazaruni
(Kartabo; Bartica Grove; Kalacoon); Potaro-Sipa-
runi (Potaro District); Upper Takutu-Upper Esse-
quibo (Ireng Valley); SURINAME; VENEZUELA:
Sucre (Maturfn [now in Monagas]). Tate, 1939:
164 listed.
Marmosa cineaea demararae: Cabrera, 1958:13 in-
correct subsequent spelling listed.
[Caluromys] demararae: Matschie, 1916:269 incor-
rect subsequent spelling listed.
Marmosa demararae: Tate, 1933:60 description.
Marmosa demararae ssp.: Mendez, 1951:207 incor-
rect subsequent spelling VENEZUELA: Miranda
(La Cienaga [= La Cienega], near Baruta); Miran-
da (Turgua).
Micoureus demerarae: Gardner, 1993:20. Peres et al.,
1999:39 BRAZIL: Rio de Janeiro (Poco das An-
tas Biological Reserve). Patton et al., 2000:68
Acre (Nova Vida, right bank Rio Jurua; opposite
Igarape Porongaba, left bank Rio Jurua, Igarape
Porongaba, right bank Rio Jurua; Sobral). Amazon-
as (above mouth, right bank Rio Jau; Macaco, left
bank Rio Jau; Boa Esperanca, right bank Rio Jurua;
Ilha Paxiuba; Tambor, left bank Rio Jau; Comuni-
dade Colina, right bank Rio Tiquie, Municipio San
Gabriel de Cachoeira; Lago Medufnim, left bank
Rio Negro; Seringal Condor, left bank Rio Jurua;
Penedo; Barro Vermelho, Condor, left bank Rio Ju-
rua; Lago Vai-Quem-Quer; Altamira, right bank
Rio Jurua; Colocacao Vira-Volta, left bank Rio Ju-
rua; Alto Rio Urucu, Tefe; Para (Altamira, 52 km
SSW); VENEZUELA: Amazonas (Camp VII, Cer-
ro de La Nablina); GUYANA: Rupununi (Quarter-
mile Landing, Rupununi River, 5 km S of Anna);
BOLIVIA: Santa Cruz (Flor de Oro, left bank Rio
Itenez, Parque Noel Kempff Mercado); PERU:
Cusco (Tangoshiari, 2 km SW, Rio Pangoreni). Ma-
res and Braun, 2000:34 ARGENTINA: Misiones
(Arroyo Vina |= brazo del Arroyo Garupa]; Parque
Nacional Iguazu). Voss et al., 2001:53 FRENCH
GUIANA (Paracou); GUYANA: Cuyuni-Mazaruni
(Kartabo); Potaro-Siparuni (Surama, 5 km SE;
Iwokrama Reserve); Upper Demerara-Upper Es-
sequibo (Comackka on Demerara River); Upper
Takutu-Upper Essequibo (Ireng Valley; Kuitaro
River, 40 mi E of Dadanawa; Weri More, not lo-
cated; Achamere Wan, not located).
Marmosa domina Thomas, 1920b:280 BRAZIL:
Para (type locality, Villa Braga, Rio Tapajos; ho-
lotype, female, skin and skull, BMNH 20.7.14.39,
collected 19 June 1917 by Emilia Snethlage). Tate,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
97
1933:72 BRAZIL: "Amazonia*: Para (Villa
Braga. Rio Tapajoz): Roraima [formerly Rio Bran-
co] (Serra da Lua. near Boa Vista. Rio Branco):
Amazonas (Lago do Capiranga: Lago do Arara:
Ayapua [= Aipua]. Rio Purus: Villa Bella Imper-
atriz [= Parintins]: Rio Andira. boca). Carvalho.
1962:285. 296 BRAZIL: Amapd (Rio Araguari.
Macapa). Avila Pires. 1964:8. 1 1 BRAZIL: Para
(Villa Braga. Rio Tapajos): Amazonas (Manaus).
Gardner. 1993:20 listed.
Marmosa limae Thomas. 1920b:282 BRAZIL:
Ceard (tvpe locality; holotvpe. female, skin and
skull. BMNH 20.7.14.41: collected 1916 by F.
Lima and lived for some months in the Para Mu-
seum, died 11 March 1916). Gardner. 1993:20
listed.
Micoureus limae: Patton et al.. 2000:67-68 (tenta-
tively to the coastal Mata Atlantica. Brazilian
clade) BRAZIL: Minas Gerais (Parque Estadual
do Rio Preto. 15 km S of Sao Goncalo do Rio
Preto: Parque Estadual do Rio Doce. 13 km E of
Marlieria): Espirito Santo (Fazenda Santa Terezin-
ha. 33 km NE of Linhares): Sao Paulo (Fazenda
Intervales. Capao Bonito).
Marmosa demararae areniticola Tate. 1931:2 VE-
NEZUELA: Bolivar (type locality. Arabupu [= Ar-
abopo]. foot of Cerro* Roraima.' 4200 ft [= 1300
mj: holotvpe. male, skin and skull. AMNH 75691.
collected 29 December 1927 bv G.H.H. Tate and
T. D. Carter). Tate. 1933:63 VENEZUELA: Bo-
livar (Arabupu [= Arabopo], Cerro Roraima). Tate.
1939:164 VENEZUELA: Bolivar (Cerro Rorai-
ma. 1300 m: Auvan-tepui. 460-1850 m). Gardner.
1993:20 listed."
Marmosa cinerea arenticola: Perez-Hernandez. 1989:
364 VENEZUELA: Bolivar (Carretera El Dora-
do Santa Elena).
Marmosa demararae esmeraldae Tate. 1931:2 VE-
NEZUELA: Amazonas (type locality. Esmeralda,
near foot of Cerro Duida. upper Orinoco. 325 ft [=
95 m]: holotvpe. male. AMNH 76964: collected 7
October 1928 by G.H.H. Tate: Casiquiare Canal
and Cerro Duida region to 6200 ft [= 1930 m]).
Tate. 1933:64 VENEZUELA: Amazonas (Esmer-
alda: Cerro Duida: upper Rio Orinoco: Rio Ocamo.
SW of Esmeralda: Casiquiare Canal: Quemapure.
Casiquiare Canal). Tate. 1939:164 VENEZUE-
LA: Amazonas (Cerro Duida. 100-2000 m). Gard-
ner. 1993:20 listed.
[Marmosa demararae] esmeraldae: Krumbieael.
1 94 lc:5 VENEZUELA: Aragua (Maracay).
Marmosa cinerea esmeraldae: Perez-Hernandez.
1989:365 VENEZUELA: Amazonas (La Esmer-
alda: Campamento No. 2. Duida: Isla Cigarron. Rio
Negro: Rio Manapiare. San Juan de Manapiare: Rio
Casiquiare. Capibara: Raudal Cerguera. Rio Au-
tana).
Marmosa cinerea paraguayana Tate. 1931:1 PAR-
AGUAY: Guaird (type locality. Villa Rica [= Vil-
larica]: holotvpe. adult male. BMNH 25.5.1.15.
collected by Schade): BR.AZIL: Minas Gerais (La-
goa Santa): Santa Catarina. Tate. 1933:58 PAR-
AGUAY: Guaird (Villa Rica [= Villarica]): Para-
guari (Sapucay [= Sapucaf]); BRAZIL: Parana
(Rio Jordao): Santa Catarina (Hansa [= Corupa]):
"Rio de Janeiro. Sao Paulo": Sao Paulo (Piquete):
Minas Gerais (Lagoa Santa). Yepes. 1938:42
PARAGUAY: BRAZIL: ARGENTINA: (no speci-
mens). Vieira. 1949:350 BRAZIL: Rio Grande
do Sul. Vieira. 1955:351 BRAZIL: Minas Gerais
(Lagoa Santa): Sao Paulo (Itapetininga: Bauni;
Ubatuba: Tambaii): Parana: Santa Catarina (Join-
ville [= Join vile]: Colonia Hansa [= Corupa]). Ca-
brera. 1958:1-4 classification. Gardner. 1993:20
listed.
[Marmosa cinerea] paraguaxana: Krumbiegel.
1 94 lc:3 PARAGUAY: San Pedro (Nueva Ger-
mania).
Marmosa demararae meridae Tate. 1931:3. 65 VE-
NEZUELA: Merida (type locality. Cafetos de Me-
rida. 1630 m: holotvpe. male, skin and skull.
USNM 137510: collected 14 June 1904 by Sala-
mon Bricefio Gabaldon). Tate. 1933:65. 66 part.
VENEZUELA: Merida (Merida region; Cafetal de
Chama [= Hama]: Cafetal de Llano, not located;
Cafetal Fuente. not located: Cafetal de Merida;
Cafetal de Milla: Platanales de Ios Campos, not lo-
cated): Carabobo (San Esteban); COLOMBIA:
Boyacd (Miraflores): Cundinamarca (Santo Do-
mingo de Bogota): Meta (Villavicencio). Tate.
1947:65 VENEZUELA: Aragua (Rancho Grande.
1100 m). Gardner. 1993:20 listed.
Marmosa cinerea meridae: Perez-Hernandez. 1989:
365 VENEZUELA: Aragua (Estacion Biologica
de Rancho Grande. Parque Nacional "'Henry Pit-
tier": Rancho Grande: La Quinta. Rancho Grande:
Pica Periquito. El Portachuelo. Rancho Grande:
Don Ramon. Rancho Grande: Guamitas. Rancho
Grande); Amazonas (Pantoja. Carretera Maracay-
Ocumare: Guacamaya. Rancho Grande): Merida
(Hacienda San Pedro: Las Porqueros): Miranda
(Estacion Experimental de Rio Negro): Lara (La
Blanquita. Parque Nacional Yacambu).
M[armosa] c[inerea] pfrimeri Miranda Ribeiro. 1936:
360. 365. 366 BRAZIL: Goids (type locality. Pal-
ma: syntypes. male and female, skins and skulls.
MNRJ. presented by Dr. Rudolph Pfrimmer). Gard-
ner. 1993:20 listed.
Marmosa cinerea pfrimeri: Vieira. 1955:351 listed.
M[armosa] c[inerea] travassosi Miranda Ribeiro.
1936:360. 365. 366 BRAZIL: Rio de Janeiro
(type locality. Angra dos Reis: holotvpe. skin and
skull. MNRJ. presented by Dr. Lauro Travassos).
Gardner. 1993:20 listed. '
Holotype Female, skin and skull. BMNH
5.1 1.1.25; collected 29 June 1905 by S. B. Warren.
Type Locality Comaccka [= Takama], 80 mi
above mouth Demerara River. East Demerara-
West Coast Berbice. Guyana.
Distribution (Fig. 57) South America in Co-
lombia. Peru. Venezuela, Guyana, Suriname.
French Guiana. Brazil. Bolivia. Paraguay, and Mi-
siones. Argentina.
Comments Previously known as Marmosa ci-
nerea; cinerea preoccupied by D. cinerea Gold-
fuss, 1812.
98
FIELDIANA: ZOOLOGY
Micoureus demerarae
l... 500 1000 1500 km
1 ' go 5 80* ' 70 5 ' 60 s 50 40 30 20 10
Fig. 57. Distribution of Micoureus demerarae: Colombia, Venezuela, the Guianas, Suriname, Brazil, Peru, Bo-
livia, and Paraguay. = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
99
Micoureus regina (Thomas, 1898)
Didelphys cinerea: Thomas (not Goldfuss, 1812),
1880:403 ECUADOR: Pastaza (Sarayacu). Thom-
as, 1882a:lll PERU: Amazonas (Chirimoto).
Didelphys [{Micoureus)] cinerea: Thomas (not Gold-
fuss, 1812), 1888b:342 part, PERU: (upper Ama-
zonas); ECUADOR: Pastaza (Sarayacu).
Marmosa regina Thomas 1898b:274. Tate, 1933:83
COLOMBIA: (type locality "West Cundinamarca
[Bogota region]"; holotype, male, skin and skull,
BMNH 98.5.15.4). Cuervo Diaz et al., 1986:473
COLOMBIA: Cundinamarca (Bogota). Gardner,
1993:20 listed.
Philander regina: J. A. Allen, 191 1:246 VENE-
ZUELA: Aragua (San Esteban [now in Carabo-
bo\).
[Caluromys] regina: Matschie, 1916:269 classifica-
tion.
Micoureus regina: Pacheco et al., 1993:5 PERU:
Madre de Dios (Pakitza; Altamira; Hacienda Ama-
zonia; Itahuanfa; Cocha Cashu). Emmons, Ascorra,
and Romo, 1994 PERU: Madre de Dios (Refugio
Picoplancha; Explorer's Inn Reserve). Anderson,
1997:157 BOLIVIA: La Paz (Mamas, 13 km
SW; Apolo, 17 km N; Mapiri; Ticunhuaya; Guan-
ay; Guanay, 5 km SE; Alcoche, 4 km NW; Car-
anavi, 20 km NNE; Pitiguaya; Nequejahuira); San-
ta Cruz (Punta Rieles). Patton et al., 2000:68-75
BRAZIL: Acre (Sobral, left bank Rio Jurua; op-
posite Igarape Porongaba, left bank Rio Jurua; Igar-
ape Porongaba, right bank Rio Jurua; Nova Vida);
Amazonas (Seringal Condor, left bank Rio Jurua;
Boa Esperanca; Jainu; Barro Vermelho; Sacado;
Nova Empresa; opposite Altamira; Penedo, right
bank Rio Jurua); PERU; Amazonas (Huampami).
Marmosa rapposa Thomas, 1899c:42 PERU: Cusco
(type locality, Rio Vilcanota, 1500 m; holotype, fe-
male, skin and skull, BMNH 98.11.1.13; collected
December 1897 by Otto Garlepp). Thomas, 1920c:
247 PERU: Cusco (Rio Vilcanota). Tate, 1933:
77 PERU: Cusco (Rio Vilcanota); Puno (Santo
Domingo, Rio Inambari, 6000 ft [= 1875 m]); Ju-
nin (Utcuyacu; Rio Perene, 4800 ft [= 1500 m]).
Gardner, 1993:20 listed.
[Caluromys] rapposa: Matschie, 1916:269 listed.
Marmosa phaea Thomas, 1899c:44 COLOMBIA:
Narino (type locality, San Pablo; holotype, female,
skin and skull, BMNH 98.9.5.2, collected 29 March
1897 by Gustav Hopke). J. A. Allen, 1916a:201
part, COLOMBIA: Narino (Ricaurte); Huila (La
Candela). Lonnberg, 1921:69 ECUADOR: Pi-
chincha (Mindo, 5500 ft [= 1700 m]). Tate, 1933:
84 COLOMBIA: Narino (San Pablo; Ricaurte,
5000 ft [ = 1560 m]); Huila (La Candela, 6000 ft
[= 1875 m]); COLOMBIA: Cundinamarca ("Cha-
carcito, Itoco, near Bogota"); ECUADOR: Pichin-
cha (Ojo Negro, road from Nanegal to Anca; Na-
negal); Loja (Santa Ana; Guainche); El Oro (Pi-
fias); perplexa Anthony, a synonym. Gardner, 1993:
20 listed.
Marmosa germana Thomas, 1904a: 143. Thomas,
1 92 la:22 ECUADOR: Pastaza (type locality,
Sarayacu, Rib Bobonaza, 2500 ft [= 780 m]; ho-
lotype, female, skin and skull, BMNH 80.5.6.77,
collected by Clarence Buckley). Thomas, 1927a:
373 PERU: San Martin (Yurac Yacu; San Anto-
nio, Lamas, SE of Moyobamba). Thomas, 1927b:
606 PERU: Hudnuco (Chinchavito, 3000 ft [=
930 m]). Gardner, 1993:20 listed.
[Caluromys] germana: Matschie, 1916:269 listed.
Marmosa germana germana: Tate, 1933:80 EC-
UADOR: Pastaza (Sarayacu, Rfo Bobonaza, Or-
iente; Canelos, Rio Bobonaza).
Marmosa waterhousei: Osgood (not Tomes, 1860),
1914b:150 PERU: San Martin (Moyobamba);
Loreto (Yurimaguas); synonyms, part, cinerea
Thomas, 1888; Marmosa germana Thomas, 1904,
from Sarayacu, Ecuador.
[Marmosa] waterhousei: Cabrera (part, not Tomes,
1860), 1919:39 germana incorrectly regarded as
a synonym.
Marmosa perplexa Anthony, 1922:3 ECUADOR:
Loja (type locality, Santa Ana trail from Zaruma to
Loja, 3650 ft [= 1 140 m]; holotype, female, skin
and skull, AMNH 47188, collected 21 December
1920 by H. E. Anthony). Gardner, 1993:20 listed.
Marmosa rutteri Thomas, 1924c:536 PERU: Loreto
(type locality, Tushemo near Masisea, Rio Ucayali,
1000 ft [= 300 m]; holotype, male, skin and skull,
BMNH 24.2.22.67, collected 10 September 1923
by L. Rutter). Thomas, 1928a:265 PERU: Loreto
(San Jeronimo, Rio Ucayali; Contamana, Rio Uca-
yali). Thomas, 1928b:294 PERU: Loreto (Pebas).
Gardner, 1993:20 listed.
Marmosa germana rutteri: Tate, 1933:81 PERU:
Ucayali (near Masisea, Rio Ucayali); Loreto (San
Jeronimo, Rfo Ucayali; Contamana, Rio Ucayali;
Sarayacu, Rfo Ucayali; Yurimaguas, Rfo Huallaga;
Pebas, Rfo Maranon; Orosa, boca Rfo Napo; boca
Rfo Curaray; Lagarto Cocha [= Lagartococha], Rfo
Napo); ECUADOR: Napo (Rfo Suno, Rfo Napo;
San Jose Nuevo, Rfo Napo). Sanborn, 1949:277
PERU: Loreto (Yarinacocha [now in Ucayali], Rfo
Ucayali). Baker, 1974:135 ECUADOR: Napo
(Santa Cecilia; Limoncocha).
Marmosa mapiriensis Tate, 1931:3 BOLIVIA: La
Paz (type locality, Ticunhuaya, on road from Sorata
to Guanay, Rfo Tipuani, 4800 ft [= 1500 m]; ho-
lotype, male, skin and skull, AMNH 72555, col-
lected 23 April 1926 by G.H.H. Tate). Tate, 1933:
76 BOLIVIA: La Paz (Ticunhuaya, Rfo Tipuani;
Nequejahuira, Rfo Unduavi, 8000 ft [= 2500 m];
Mapiri, Rfo Mapiri, 2000 ft [= 625 m]; Guanay,
Rfo Mapiri, 1700 ft [= 530 m]); PERU: Puno (San-
to Domingo, Rfo Inambari; Rfo Tavara, Rfo In-
ambari). Sanborn, 1951b:2 PERU: Cusco (Haci-
enda Cadena). Gardner, 1993:20 listed.
Marmosa germana parda Tate, 1931:4 PERU:
Hudnuco (type locality, Huachipa, mouth Rfo Ca-
yumba, upper Rfo Huallaga, 3000 ft [= 930 m];
Chinchavito, Rfo Huallaga; holotype, male, skin
and skull, FMNH 24140, collected 28 September
1922 by J. T. Zimmer). Tate, 1933:82 PERU: San
Martin (San Antonio, Lamas, Rfo Mayo [upper];
Yurac Yacu, Rfo Mayo [upper]; Moyobamba, Rfo
Mayo [upper]); Hudnuco (Huachipa, Rfo Huallaga
[upper]; Rfo Cayumba [upper], Rfo Huallaga [up-
per], 3000 ft [= 930 m]; Rfo Chinchao [upper], Rfo
Huallaga [upper]; Chinchavito, Rfo Huallaga; Po-
zuzo [now in Pasco], Rfo Pachitea [upper]); Junin
(Rio Perene [Tschudi coll.] [upper]). Gardner,
1993:20 listed.
100
FIELDIANA: ZOOLOGY
Fig. 58. Distribution of Micoureus regina: Colombia and northwestern Venezuela south to Ecuador, Peru, Bolivia,
and Brazil. = localities given in published records.
Holotype Male, skin and skull, BMNH
98.5.15.4; collected 1 November 1895 by G. D.
Child.
Type Locality Bogota region, Cundinamar-
ca, Colombia.
Distribution (Fig. 58) Colombia from the
Bogota and Rio Magdalena region north into
northwestern Venezuela; south through Ecuador,
Peru, Bolivia, and western Brazil.
Genus Monodelphis Burnett, 1830 [1829]
Sarigua Muirhead, 1819:429 part, 5. brachyura list-
ed among included species.
Monodelphis Burnett, 1830:351 included species,
dorsigerens [sic] [ = Marmosa dorsiguera Linnae-
us], brachyura. Gardner, 1993:21 listed.
Peramys Lesson, 1 842: 1 87 included species: brack-
yurus (= brachyuros Schreber [type species by
subsequent designation], crassicaudata Desmarest
[= Lutreolina crassicaudata], tristriata Illiger, pus-
ilia Desmarest [= Thylamys pusilla]). Thomas,
1888b:354 subgenus of Didelphys [sic] Linnaeus;
type species designation, "D. brevicaudata" [= D.
brachyuros Schreber]. Matschie, 1916:269 sub-
genus of Didelphis Linnaeus; invalid designation of
crassicaudata Desmarest as type species, earlier
fixation overlooked. Cabrera, 1919:41 type, D.
brachyuros Schreber [= D. brevicaudata Erxle-
ben].
Hemiurus Gervais, 1855:101 name attributed to I.
Geoffroy; included species: tristriata, tricolor,
brachyura, hunteri [Gervais, not Waterhouse = do-
mestica Wagner]; generic name preoccupied by
Hemiurus Rudolphi, 1809, a genus of fluke (Trem-
atoda). Thomas, 1888b:354 synonym of subgenus
Peramys; selected type species, "D. brevicaudata"
[= brachyura]. Matschie, 1916:261 selected type
species, hunteri Gervais [ = Monodelphis domesti-
ca Wagner].
Microdelphys Burmeister, 1856:83 included species,
tristriata, tricolor, brachyura velutina (a Marmo-
sa). Thomas, 1888b:354 synonym of subgenus
Peramys; selected type species, "D. americana"
[ D. tristriata].
Microdidelphys Trouessart, 1898:1242 incorrect
subsequent spelling of Microdelphys Burmeister in
synonymy of Peramys.
Microdelphis Cabrera, 1958:6 incorrect subsequent
spelling of Microdelphys Burmeister.
Monodelphiops Matschie, 1916:261, 267, 272 type
species, M. sorex Hensel, by original designation.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
101
Minuania Cabrera, 1919:43 type species, Didelphis
dimidiata Wagner, by original designation.
Type Species Monodelphis brachyura Burnett
[= Didelphis brachyuros Schreber = Didelphis
brevicaudata Erxleben], by subsequent selection
(Matschie, 1916:271).
Distribution (Fig. 59) Panama south to cen-
tral Argentina.
Monodelphis adusta (Thomas, 1897)
Didelphis brachxura: Roulin (not Schreber), 1843:
23 COLOMBIA.
Peramys adustus Thomas, 1897:219 description.
[Monodelphis] adustus: Matschie, 1916:271 listed.
Monodelphis adusta: Tammitt and Valdivieso, 1964:
325 COLOMBIA: Meta (Villavicencio). Han-
dley, 1966:755 PANAMA: Darien (Tacarcuna:
Cana). Cuervo Diaz et al., 1986:473 COLOM-
BIA: Choco (Serrania del Darien). Pacheco et al..
1993:1 PERU: Cusco (km 152, above San Pedro.
72 km NE Paucartambo); Madre de Dios (Haci-
enda Amazonia). Gardner, 1993:21 listed. Rageot
and Albuja, 1994:177 ECUADOR: Pastaza
(Mera). Perez-Hernandez, 1989:370 VENEZUE-
LA: Tdchira (Pregonero). Perez-Hernandez et al.,
1994:28 VENEZUELA: Merida (Cordillera de
Merida, south of Lago do Maracaibo). Anderson.
1997:158 BOLIVIA: La Paz (La Reserva).
Peramys melanops Goldman, 1912:2 PANAMA:
Darien (type locality, Santa Cruz de Cana, upper
Rio Tuyra; holotype. male, skin and skull, USNM
179609, collected 23 May 1912 by E. A. Gold-
man). Gardner, 1993:21 listed.
[Monodelphis] melanops: Matschie. 1916:271 clas-
sification.
Peramys peruvianas Osgood, 1913b:93 PERU: San
Martin (type locality, Moyobamba; holotype, male
skin and skull, FMNH 19362. collected 21 July
1912 by W. H. Osgood and M. P. Anderson). Os-
good, 1914b: 147 PERU: San Martin (Moyo-
bamba). Thomas, 1920c:248 PERU: Cusco (Oco-
bamba Valley). Gardner, 1993:21 listed.
Monodelphis peruvianus: Matschie. 1916:271 list-
ed.
Monodelphis peruviana peruviana: Ceballos. 1958:
48 incorrect subsequent spelling PERU: Loreto
(Pucallpa [now in Ucayali]).
Monodelphis adusta peruviana: Cabrera. 1958:6 in-
correct subsequent spelling Ceballos. 1959:265
PERU: Loreto (Pucallpa [now in Ucayali]).
Monodelphis kunsi: Anderson, 1997 BOLIVIA: La
Paz (La Reserva).
Holotype Skin and skull, BMNH 97.7.2.1.
Type Locality "W. Cundinamarca, in the
low-lying hot regions," Colombia.
Distribution (Fig. 60) Panama south through
Colombia, Ecuador, Peru, and Bolivia; northwest-
ern Venezuela.
Monodelphis americana (Miiller, 1776)
Mus araneus, figura muris Marcgrave, 1648:229.
Musaraigne de Bresil Buffon. 1767:160.
Sorex americanus Miiller. 1776:36 Gardner, 1993:
21 listed.
Peramys americanus: Ihering, 1894:12 BRAZIL:
iheringi Thomas, probably a synonym. Thomas,
19 12a:89 BRAZIL: Para (Benavides, near Para).
Miranda Ribeiro, 1936:414 BRAZIL: Para.
Monodelphis (Microdelphys) americanus: Pohle,
1927:240 BRAZIL: Pernambuco (Dois Herma-
nos [= Dois Irmaos]).
Didelphys americana: Thomas, 1888b:363 BRA-
ZIL: synonyms, brasiliensis Erxleben. tristriatus II-
liger, trilineata Lund.
[Microdelphys] americana: Matschie, 1916:272
tristriata Illiger a synonym.
Monodelphis americana: Davis, 1945b:280 BRA-
ZIL: Rio de Janeiro (Teresopolis). Davis. 1947:3
BRAZIL: Rio de Janeiro (Teresopolis). Vieira,
1949:358-359 BRAZIL: Sao Paulo (Bauru; Uba-
tuba: Piracicaba): Rio de Janeiro (Ilha Grande); Mi-
nas Gerais (Mariana). Avila Pires. 1968:167
BRAZIL: Goids (Veadeiros). da Fonseca and Ki-
erulff, 1988:117 BRAZIL: Minas Gerais (Fazen-
da Montes Claros). Langguth and Lima, 1988:1
BRAZIL: Paraiba (Joao Pessoa). Mares and
Ernest. 1995:755 BRAZIL: Distrito Federal (Re-
serva Ecologica Biologica do Instituto de Geografia
e Estatistica [IGBE]).
[Sorex] brasiliensis Erxleben, 1777:127 name based
primarily on the Mus araneus of Marcgrave; type
locality. Brazil. Gardner. 1993:21 listed.
Sorex brasiliensis: Schreber. 1777:577.
Sorex Brasiliensis: E. Geoffroy, 1811:172, footnote
1 Didelphis tricolor E. Geoffroy regarded as a
synonym.
Sorex braziliensis: Thomas, 1888b:363 incorrect
subsequent spelling.
[Didelphys] tristriata Illiger. 1815:107. 112 BRA-
ZIL: Pernambuco (type locality [now restricted]);
name based on the 'Sorex brasiliensis" of Marc-
grave. Gardner. 1993:21 listed.
D[idelphys]. tristriata: Olfers, 1818:206 BRAZIL:
Para; Maranhdo; Pernambuco.
Didelphys tristriata: Lichtenstein, 1817:217 Mus
araneus of Marcgrave. Pelzeln. 1883: 116 BRA-
ZIL: Sao Paulo (Ipanema).
Didelphis tristriata: Kuhl. 1820:63 BRAZIL; spec-
imen in Berlin Museum; Sorex surinamensis E.
Geoffroy a possible synonym.
Microdelphys tristriata: Burmeister, 1856:84 BRA-
ZIL: Rio de Janeiro (Novo Friburgo).
[Cuica] tristriata: Liais, 1872:330 classification.
Peramys tristriatus: Lesson, 1842:187. Goeldi, 1894:
463 BRAZIL: Rio de Janeiro (Serra dos Orgaos).
Goeldi and Hagmann, 1904:100 BRAZIL: Para
(vicinity of Para [= Belem]).
Hemiurus tristriatus: Winge, 1893:8, 58 BRAZIL:
Minas Gerais (Lagoa Santa).
H[emiurus] tristriatus: Miranda Ribeiro, 1905:189
BRAZIL: Rio de Janeiro (Monte Serrat. Itatiaya [=
Pico das Agulhas Negras]).
Didelphis trilineata Lund. 1839:233 nomen nudum;
102
FIELDIANA: ZOOLOGY
Monodelphis
/
\ /
C i o" 80 ' 70 "' 60~ 50 40 30 20 10
Fig. 59. Distribution of the genus Monodelphis: southern Panama in Central America and throughout South
America east of the Andes. = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
103
Monodelphis adusta
1500 km
30
Fig. 60. Distribution of Monodelphis adusta in Central and South America: Panama south through Colombia,
Ecuador, and Peru and in northwestern Venezuela. = localities given in published records.
BRAZIL: Minas Gerais (Rio das Velhas). Gardner.
1993:21 listed.
Holotype Not known to be in existence:
name based on ^brasilianische Spitzmause, i.e..
the musaraigne de Bresil of Buffon and the mus
araneus of Marcgrave."
Type Locality Brazil; restricted to Pernam-
buco (cf. Thomas, 191 lb: 123), the general locale
of Marcgrave 's animals.
Distribution (Fig. 61) Eastern Brazil from
Para in the north to Santa Catarina in the south.
Monodelphis brevicaudata (Erxleben, 1777)
Muris silvestris americani femina Seba. 1734:50.
[?]Die surinamische Spitzmaus Schreber, 1777:575
SURINAME.
Touan Buffon. 1789:252 FRENCH GUIANA:
(Cayenne).
[Didelphis] brevicaudata: Erxleben, 1777:80. Gard-
ner. 1993:21 listed.
Didelphys brevicaudata: Thomas, 1888b:356 BRA-
ZIL: Bahia; synonyms, brachyura Schreber, suri-
namensis Gmelin. touan Shaw, touan Bechstein,
tricolor E. Geoffroy. brevicaudis Illiger [?], sebae
Gray, humeri Waterhouse. BRAZIL: Bahia. Thom-
as. 1892:314 type of Seba's Muris silvestris
americani femina in the Lidth de Jeude Collection
in the British Museum.
Monodelphis brevicaudata, Tate. 1923:65 VENE-
ZUELA: Bolivar (Rio Supamo: El Manaco; Ica-
baru): Amazonas (Puerto Ayacucho. Acanana; Es-
meralda: Tamatama; Capibara: Boca Mavaca).
Husson. 1978:12 SURINAME: Brokopondo
(Brokopondo: Brownsberg): Paramaribo (De Mor-
genstond: Paramaribo: Cultuurtuin); Commewijne
("Clevia"): Saramacca ("Jarikaba"); Marowijne
(Langamankondre). Patterson. 1992:6 BRAZIL:
Para (Cameta: Marai). Pacheco et al.. 1993:1
PERU: Madre de Dios (Pakitza). Anderson, 1997:
158 BOLIVIA: La Paz (Guanay: Guanay, 5 km
SE; Alcoche. 4 km NW); Pando (Centro Dieci-
ocho: Independencia: Santa Rosa). Voss et al.,
2001:54 GUYANA: Pomeroon-Supenaam (Su-
penaam River): Cuyuni-Mazaruni (First Falls on
Cuyuni River: Kartabo: Bartica Grove: Kamakusa);
Essequibo Islands-West Demerara (Buck Hall):
Potaro-Siparuni (Anundabaru: Minnehaha Creek:
Potaro) Upper Demerara-Berbice (Dubulay
Ranch): FRENCH GUIANA (Arataye: Cacao;
Cayenne: St. Eugene; Paracou: Sophie; Tamanoir:
104
FIELDIANA: ZOOLOGY
c=>
Monodelphis americana
500 1000 1500 km
90
80
70
60
50"
40"
30
20
10
Fig. 61. Distribution of Monodelphis americana: eastern Brazil from Para south to Santa Catarina. = localities
given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
105
Trois Sauts); SURINAME: Brokopondo (Browns-
berg: Finisanti); Marowijne (Albina. 3 km SW;
Langamankondre; Moengo. 10 km N and 4 km W);
Sipaliwini (Oelemarie: Paloemeu Airstrip: Raleigh
Falls): Nickerie (Avanavero: Kayserberg Airstrip:
King Frederick Williams Falls); Saramacca (La
Poule: Jarikaba near Uitkijk): Paramaribo (Cul-
tuurtuin. near Paramaribo: Plantage De Morgen-
stond near Paramaribo); Commewijne (Plantation
Clevia near Paramaribo): VENEZUELA: Bolivar
(El Dorado, 65 km SSE: El Manteco. 56 km SE:
Reserva Forestal Imataca, not located; Arabupu:
Auyan-tepuf: Caicara: Ciudad Bolivar); Amazonas
(Boca Rio Ocamo: Esmeralda: Mt. [= Cerro] Dui-
da; "Rio Casiquiari": Sierra [= Cerro de La] de
Neblina): BRAZIL: Amapd (Serra do Navio: Serra
do Tumucumaque); Amazonas (Manaus, 80 km N;
Santo Antonio da Amatary); Para (Cachoeira Por-
teira; Faro): type locality emended to Guyana. Kar-
tabo. Cuyuni-Mazaruni.
Pramys brevicaudatus: Cabrera. 1958:8 incorrect
subsequent spelling of Peramys in synonymy of
Monodelphis brevicaudata.
[Peramvs brevicaudatus] brevicaudatus: Cabrera.
1919:42 part, listed.
Monodelphis brevicaudata brevicaudata: Anthony.
1921b:268 GUYANA: Cuyuni-Mazaruni (Karta-
bo). Tate. 1939:166 GUYANA: SURINAME:
(type locality). Vieira. 1955:348 BRAZIL: Ron-
donia (Rio Mamore. not located). Carvalho. 1962:
284 BRAZIL: Amapd (lower Rio Araguari: Rio
Amapari. Macapa: Igarape Rio Branco. upper Rio
Maraca. Mazagao). Cabrera. 1958:7 glirina in
synonymy only.
[Monodelphis brevicaudata] brevicaudata: Avila Pi-
res. 1964:10 (in text) BRAZIL: Para (Castanhal.
Rio Jamunda [= Nhamunda. not located]): Rorai-
ma (Tapajos. not located).
Monodelphis [brevicaudata] brevicaudata: Carvalho.
1960:123 BRAZIL: Amapd (Rio Araguari: Rio
Amapari).
Monodelphis b. brevicaudata: Perez-Hernandez.
1989:370 VENEZUELA: Bolivar (San Martin de
Turumban. Anacoco. Rio Cuyuni; km 88. Carretera
El Dorado-Santa Elena: La Laja. Campamento
MOP km 109. S of El Dorado. Carretera El Do-
rado-Santa Eleana de Uairen: km 124-134. Car-
retera El Dorado-Santa Elena): Amazonas (Acan-
ana, Rio Cunucunuma: Tamatama: San Carlos de
Rio Negro; Rio Negro. Cano Ardabo).
P[eramys] brevicaudatus: Thomas. 1899a: 155 tax-
onomic comparison.
[Monodelphis] brevicaudatus: Matschie. 1916:271
classification. Type locality designated.
[Peramys brevicaudatus] brevicaudatus: Cabrera
part (not Erxleben. 1777). 1919:42 glirina in syn-
onymy only.
[Didelphis] Brevicauda Boddaert. 1784:77 (name ap-
parently proposed independently of brevicaudata
Exleben) name based on the muris silvestris
americani femina of Seba.
Monodelphis brevicauda: Sanderson. 1949:788 SU-
RINAME: Para (Zanderij).
Didelphys brachyuros Schreber. 1777:548. 589
South America: (type locality): name based pri-
marily on the muris silvestris americani femina of
Seba. Gardner. 1993:21 listed.
[Didelphis] brachyura: Gmelin. 1788:108. Wagner.
1 855:25 1 synonymy.
Didelphis Brachyura: Shaw. 1800:479 synonymy.
Didelphis bachyura: E. Geoffroy. 1803:145 char-
acters. Demarest. 1816:429. 430 characters and
synonomy. Desmarest. 1820:260 characters. Des-
marest. 1827:398 characters.
Did[elphys] brachyura [sic]: Cuvier. 1798:125 the
touan of the Guianas.
D[idelphys] brachyura: Olfers. 1818:205 touan
Shaw, a synonym.
Didelphys brachyura: Gervais. 1855:101 included
in genus Hemiurus.
Monodelphis Brachyura: Burnett. 1830:351.
Sarigua brachyura: Muirhead. 1830:420 listed.
Didelphys tricolor vel brachyura: Gray. 1827:33
part. FRENCH GUIANA: (Cayenne).
P[hilander] Brachyurus: Tiedemann. 1808:428 clas-
sification.
Peramys brachyurus: Lesson. 1842:187 classifica-
tion.
Paramys brachyurus: Cabrera. 1958:7 incorrect
subsequent spelling of Peramys in synonymy of
Monodelphis brevicaudata brevicaudata.
Sorex surinamensis Zimmerman. 1780:386. Gardner.
1993:21 listed.
[Sorex] surinamensis: Gmelin. 1788:114 SURI-
NAME.
Viverra Touan: Shaw. 1800:432 name based on
Buffon's touan. from Cayenne. Gardner. 1993:21
listed. Voss et al.. 2001:58 neotype designed:
adult male FMNH 21720 (FRENCH GUIANA:
Cayenne).
Mustela touan. Bechstein. 1800:358. 687 name
based on Buffon's touan.
Mustela Tuan: Bechstein. 1800:359. Gardner. 1993:
21 listed.
[Monodelphis] touan: Matschie. 1916:271 classifi-
cation.
Monodelphis touan touan: Cabrera, 1958:10 classi-
fication.
Monodelphis brevicaudata touan: Avila Pires. 1964:
8, 9 BRAZIL: Amazonas (Manaus; Itacoatiara;
Porto de Moz [now in Para]: Santo Antonio do
Amaterei [= Amateri]): GUYANA: Cuyuni-Maza-
runi (Kartabo): Potaro-Siparuni (Anundabaru [An-
andabaru]; Minnehaha Creek): SURINAME;
FRENCH GUIANA: (Cayenne: Tamanoir): VE-
NEZUELA.
Didelphis tricolor E. Geoffroy. 1803:144 holotype
from Cayenne, in Paris Museum, specimen number
uncertain: cf. Julien Laferriere. 1994. Desmarest,
1820:260 FRENCH GUIANA. Desmarest. 1827:
397 FRENCH GUIANA: (Cayenne). Jentink.
1887b:302 FRENCH GUIANA: (Cayenne):
GUYANA: Demerara [= Georeetown. Demerara-
Mahaica]. Jentink. 1888:222 FRENCH GUI-
ANA; GUYANA: Demerara [= Georgetown. De-
merara-Mahaica]. Gardner, 1993:21 listed.
Didelphys tricolor: Waterhouse, 1841:109 charac-
ters.
[Peramys] tricolor: Lesson. 1842:186 classification.
Microdelphys tricolor: Burmeister. 1856:85 charac-
ters.
106
FIELDIANA: ZOOLOGY
Cuica tricolor: Liais, 1872:330 classification.
D[idelphys] sebae Gray, 1827:190.
[Didelphys] hunteri Waterhouse, 1841:110 (type lo-
cality unknown; holotype, female, in alcohol, orig-
inally in Museum of College of Surgeons, BMNH
89.1.31.1). Gardner, 1993:21 listed.
Didelphys Hunteri: Waterhouse, 1846:524.
D[idelphis] macroura: Schinz, 1844:509 name er-
roneously attributed to Pallas, 1780, in synonymy
of D. tricolor E. Geoffroy, 1803.
Peramys brevicaudatus orinoci Thomas, 1 899a: 1 54
VENEZUELA: Bolivar (type locality, Caicara, Rio
Orinoco; holotype, male, skin and skull, BMNH
98.12.1.22, collected 10 August 1898 by George
K. and Stella M. Cherrie). Gardner, 1993:21 list-
ed.
Monodelphis brevicaudata orinoci: Tate, 1939:166
VENEZUELA: Bolivar (Mt. Auyantepui [= Cerro
Auyan-tepuf]); Amazonas (Cerro Duida; Rib Oca-
mo; Casiquiare [= Brazo Casiquiare]); GUYANA;
dorsalis J. A. Allen, a synonym, BRAZIL: Rorai-
ma (Limao, right bank Rio Cotinga). Anduze,
1956:10 (not Thomas); VENEZUELA: Miranda
(La Florida [now in Distrito Federal]). Reig and
Bianchi, 1969:210 VENEZUELA: Aragua (Ran-
cho Grande).
[Monodelphis] orinoci: Matschie, 1916:271 classi-
fication.
Monodelphis orinoci: Reig et al., 1977:212 VENE-
ZUELA: Guarico (Estacion Biologica los Llanos).
Perez-Hernandez, 1989:371 VENEZUELA: Ara-
gua (Hacienda La Esperanza, 8 km de Magdaleno,
Lago de Valencia); Bolivar (Campamento, Zona de
la Represa del Guri, Rio Caroni); Cojides (Hato
Nuevo); Guarico (Cuartal Zaraza-San Juan de los
Morros; San Juan de los Morros, 50 km from; Hato
La Palmita, San Francisco de Tiznados; Hacienda
"La Palmita," between Ortez and San Francisco de
Tiznados; Estacion Biologica de los Llanos, Cala-
bozo; Morichal Villasmil, Calabozo).
Peramys brevicaudatus dorsalis J. A. Allen, 1904b:
327 VENEZUELA: Bolivar (type locality,
Ciudad Bolivar; holotype, male, skin and skull,
AMNH 16126, collected 19 December 1899 By S.
M. Klages). J. A. Allen, 1904d:337 VENEZUE-
LA: Bolivar (Ciudad Bolivar; Suapure). Gardner,
1993:21 listed.
Paramys brevicaudatus dorsalis: Cabrera, 1958:8
incorrect subsequent spelling of Peramys.
Monodelphis [sp.]: Tate, 1931:248 GUYANA: (An-
undabaru [= Anandabaru, now in Potaro-Siparu-
ni]).
Holotype Name based on the muris silvesths
americani femina of Seba, preserved in spirits
with extracted skull in the Lidth de Jeude Collec-
tion, BMNH 67.4.12.540.
Type Locality Forests of South America; re-
stricted to Suriname by Matschie (1916:271);
emended to GUYANA: Cuyuni-Mazaruni (Kar-
tabo) by Voss et al., 2001:57, who explain that
the pelage color of the holotype is incompatible
with an origin in Suriname.
Distribution (Fig. 62) Venezuela, Suriname,
Guyana, French Guiana, south to Peru, Bolivia,
the Amazon basin of Brazil, and Minas Gerais in
southeastern Brazil.
Monodelphis brevicaudis Olfers, 1818
Micoure cinquieme ou micoure a queue courte Azara,
1801:295.
Col icorto Azara, 1802:258.
[Didelphys] brevicaudis Illiger, 1815:107 nomen
nudum.
D[idelphys] brevicaudis: Olfers, 1818:205 based on
"Micoure a queue Az.I p. 298."
S[arigua] brachyura: Muirhead, 1819:429 new
combination.
Monodelphis brevicaudis: Hershkovitz, 1959:338
wagneri Matschie, a synonym. Wetzel and Lovett,
1974:206 PARAGUAY: Presidente Hayes (Fortin
Juan de Zalazar).
[Monodelphis] wagneri Matschie, 1916:272 new
name for D. brachyura wagneri (Azara's micoure
a queue courte).
Didelphis brachyura: Desmarest (part, not Schreber,
Mil), 1816:429 micoure cinquieme ou micoure
a queue courte Azara in synonymy.
D[idelphis] Tricolor vel brachyura: Gray (part, not E.
Geoffroy, 1803 and Schreber, 1777), 1827:33
PARAGUAY: "Azara, during his stay at Paris, rec-
ognized in the individuals of this species which
form part of the collection of the Museum of Nat-
ural History, his own fifth micoure."
Didelphis tricolor: Desmarest (part, not E. Geoffroy,
1803), 1820:260 Azara's micoure a queue courte
in synonymy. Desmarest, 1827:397 Azara's mi-
coure a queue courte in synonymy.
Peramys brevicaudatus: Bertoni (not Erxleben, 1777),
1914:69 PARAGUAY: Alto Parana (Piraqui, not
located). Bertoni, 1939:6 PARAGUAY.
Peramys brevicaudata: Dennler (not Erxleben, 1777),
1939:232.
Type Azara's description of the micoure a
queue courte is based in two specimens from Par-
aguay; neither of these specimens is known to
have been preserved
Type Locality "Paraguay" as then defined.
Distribution (Fig. 63) Paraguay: exact distri-
bution unknown.
Remarks Hershkovitz (pers. comm.) treated
brevicaudis as a distinct species. This is contro-
versial, and future researchers may find it neces-
sary to reassign this species name.
Monodelphis dimidiata (Wagner, 1847)
D[idelphis] dimidiata Wagner, 1847:151, third foot-
note. Thomas, 1888b:355 BRAZIL: Rio Grande
BROWN: ATLAS OF NEW WORLD MARSUPIALS
107
20 10
Fig. 62. Distribution of Monodelphis brevicaudata: Venezuela east through the Guianas and Suriname. south
through Brazil, southern Peru, and western Bolivia. = localities given in published records.
108
FIELDIANA: ZOOLOGY
20
30
40
50
Monodelphis brevicaudis
1500 km
90 80 70" 60 50 40 30 20 10
Fig. 63. Distribution of Monodelphis brevicaudis: Paraguay. = localities given in published records.
do Sul (Taquara; Sao Lourenco do Sul); URU-
GUAY: Maldonado (Maldonado). Gardner, 1993:
21 listed.
[Monodelphis] dimidiata: Matschie, 1916:272 clas-
sification.
Monodelphis dimidiata: Sanborn, 1929:150 URU-
GUAY: Montevideo (Montevideo). Talace et al.,
1960:149 URUGUAY: Cane tones (40 km N of
Canelones). Fornes and Massoia, 1965:102 AR-
GENTINA: Buenos Aires (Miramar). Avila Pires,
1968:163 BRAZIL: Rio de Janeiro (Campo Bel-
lo). Reig and Bianchi, 1969:1210 ARGENTINA:
Buenos Aires (Arroyo Brusquitas, General Pueyr-
reddn). Reig et al., 1977:212 ARGENTINA:
Buenos Aires (Arroyo Brusquitas). Avila Pires and
Gouvea, 1977:8 BRAZIL: Rio de Janeiro (Cam-
po Bello; Monte Serrat; Maromba; Maromba-
Maua). Pine et al., 1985:196 ARGENTINA: Cor-
doba (Rib Ceballos); Buenos Aires (Balcarce; Bal-
carce, 15 km NW; Estacion Experimental del In-
stituto Nacional de Technologica Agropecuaria
[INTA]; INTA Areas [ditches on Ruta 226, en-
trance gate to INTA; "Hill Plot"; Crovetto Field,
6 km N of Balcarce; Pergamino; Ruta 188, Curva
de Pena]); URUGUAY: Maldonado (Maldonado).
Mares et al., 1997:105 ARGENTINA: Tucumdn
(Concepcion). Morando and Polop, 1997:131
ARGENTINA: Cordoba. Mares and Braun, 2000:
35 ARGENTINA: Buenos Aires (Arroyo Brus-
quitas; Arroyo Chocori; Arroyo del Pescado; Baliza
Caniu; Baliza San Andres; Miramar; Estacion Ex-
perimental del Instituto de Tecnologfa Agropecu-
aria [INTA], 15 km NW Balcarce; INTA areas,
Ruta 226, between km 65 and 75; Ruta 226; "Hill
Plot"; "Crovetto Field"; Estacion Tambo Nuevo y
Ruta 188, km 60, Partido de Pergamino; Pergami-
no; Ruta 188, Curva de Pena; Santa Clara del Mar;
Sierra de La Peregrina); Cordoba (Rio Ceballos;
Yacanto); La Pampa (Caleufu); Misiones (Arroyo
Zaiman; El Cruce, not located); Salta (Tartagal);
Buenos Aires (La Tinta; Sierra de La Tinta).
[Minuania] dimidiata: Cabrera, 1919:44 classifica-
tion.
Minuania dimidiata: Devincenzi, 1935:91 URU-
GUAY.
M\inuania\ dimidiata: Miranda Ribeiro, 1936:419
BRAZIL: Rio Grande do Sul (Taquara do Mundo
Novo [= Taquara]; Sao Lourenco do Sul).
Peramys dimidiatus: Ihering, 1892:100 BRAZIL:
Rio Grande do Sul. Thomas, 1900:349 BRAZIL:
Parana (Palmeira). Bertoni, 1939:6 PARA-
GUAY: Alto Parana (Puerto Bertoni); not the col-
icorto of Azara. Lyne, 1959:84 BRAZIL: Parana
(Palmeira).
Didelphys brachyura: Waterhouse (not Schreber,
1777), 1838:64. Waterhouse, 1839:97 URU-
GUAY: Maldonado (Maldonado). Waterhouse,
1841:111 URUGUAY: Maldonado (Maldonado).
BROWN: ATLAS OF NEW WORLD MARSUPIALS
109
20
30
40
50
Monodelphis dimidiata
--. 500 1000 1500 km
90
80 r
70 r
60
5C ;
40='
30'"
20*
10
:
Fig. 64. Distribution of Monodelphis dimidiata: southeastern Brazil. Uruguay, and northern and central Argentina.
= localities given in published records.
Waterhouse, 1 846:522 part, URUGUAY: Mal-
donado (Maldonado). Burmeister, 1854:141
BRAZIL. Burmeister. 1856:86 part. URUGUAY.
Burmeister, 1879:194 part, URUGUAY.
D[idelph\s] brachyura: Wagner (not Schreber, 1777),
1855:251 part", URUGUAY.
Microdelphxs brachyura: Burmeister. 1856:86. Hen-
sel, 1873:122 BRAZIL: Rio Grande do Sid.
Didelphxs brevicaudata: Figueira (not Schreber).
1894:7 (separate) part, URUGUAY.
Monodelphis fosteri Thomas. 1924b:586 ARGEN-
TINA: La Pampa (type locality, Caleufu. station on
Western Railway: holotype. immature female, skin
and skull. BMNH 24.4.5.1, collected 10 April 1923
and presented by Frank Foster). Pine et al.. 1985:
206 ARGENTINA: La Pampa (Caleufu): URU-
GUAY: Montevideo (Colon). Gardner, 1993:21
listed.
Minuania fosteri: Cabrera and Yepes, 1940:34.
Monodelphis (Minuania) fosteri: Crespo. 1964:62.
124 ARGENTINA: Cordoba (Yacanto), Buenos
Aires (Abra de la Ventana, Sierra de la Ventana:
Tornquist, 550 m); Salta (Tartagal).
Minuania dimidiata itatiayae. Cabrera, 1958:11 cit-
ed in synonymy of Monodelphis dimidiata Wagner.
Holotype Male, skin and skull, BMNH; col-
lected by Charles Darwin.
Type Locality Maldonado, Uruguay.
Distribution (Fig. 64) Uruguay, Paraguay,
southeastern Brazil, and the northern half of Ar-
gentina.
Monodelphis domestica (Wagner, 1842)
Didelphxs domestica Wagner. 1842:359. Wagner.
1848:153 BRAZIL: Mato Grosso (Cuyaba [Cu-
iaba]). Thomas. 1888b:358 BRAZIL: Mato Gros-
so (Cuyaba [Cuiaba]. cotype): Goids. Vieira. 1955:
348 BRAZIL: Bahia (Machada Portela: Feira de
Santana): Goids (Cana Brava; Rio Sao Domingos);
Mato Grosso do Sul (Miranda): Minus Gerais (Bar-
ro Alto Rio Sao Francisco, not located). Gardner,
1993:21 listed.
Microdelphxs domestica: Burmeister. 1856:117
BRAZIL.
Monodelphis domestica: Pine. 1979:497 BOLIVIA:
Beni (San Joaquin: La Granja. Itenez); Chuquisaca
(Tomima): Santa Cruz (San Ignacio de Velasco:
Basilio); BRAZIL: Mato Grosso (Lava Pes. Facao
Road. Sao Luis de Caceres): Mato Grosso do Sul
(Sao Marcus Road. Corumba): Ceard (Quixada;
Santana do Cariri). Mares et al.. 1981:101 BRA-
ZIL: Piaui (Fazenda Olho da Agua); Ceard (Sftio
110
FIELDIANA: ZOOLOGY
Cereado, Jardim, Solonopole; Sftio Volta, Campo
Sales; Sftio Constantino, Crato; Sftio Olaria, Crato;
Sftio Araruna, Missao Velha; Sftio Cinta da Soli-
dade, Missaa Velha; Sftio Emboscada, Missao Vel-
ha; Sftio Lapinha, Missao Velha; Sftio Camara,
Milagres); Bahia (Fazenda Capoeira do Rosario,
Feira; Fazenda Cayumba, Feira; Fazenda Jacii, Fei-
ra; Fazenda Salgado, Feira; Fazenda Cacua, Serrin-
ha; Fazenda Oiteiro, Serrinha; Fazenda Umburana,
Serrinha); Pernambuco (Exu; Fazenda Pinheira,
Exu; Fazenda Batente, Exu; Escola Agricola de
Exu; Sftio Cana Brava de Jerico, Triunfo; Sftio Boa
Esperanca de Jerico, Triunfo; Sftio Paus Preto, Bo-
doco; Sftio Sao Goncalo, Bodoco; Sftio Belem, Bo-
doco; Sftio Macaco de Baixa Verde, Triunfo; Sftio
Sao Mateus, Triunfo; Sftio Novo, Triunfo; Sftio
Borgens, Triunfo; Sftio Corredor do Vento, Triunfo;
Fazenda Caianinha, Pesqueira; Fazenda Sororoca,
Pesqueira; Sftio Xique-Xique, Bodoco; Fazenda
Saco; Sftio Oiti, Triunfo; Sftio Riacho Fundo, Gar-
anhuns); Alagoas (Sftio Cachoeira Grande, Vicosa;
Sftio Lagoinha, Santana do Ipanema; Sftio Capu-
ma, Palmeira dos Indios; Sftio Dormiao, Palmeira
dos Indios; Sftio Riveira, Palmeira dos Indios; Sftio
Sabia, Palmeira dos Indios; Sftio Barracao, Limo-
eira da Anadia). Mares et al., 1989:18 BRAZIL:
Mato Grosso do Sul (Fazenda Nhumirim); Goids
(Cristalina). Anderson et al., 1993:16 BOLIVIA:
Santa Cruz (San Rafael de Amboro; Basilio). An-
derson, 1997:159 BOLIVIA: Beni (San Joaqufn;
La Granja); Chuquisaca (Rio Limon; Tihumayu,
Tomina province; Porvenir; Carandayti, E of); San-
ta Cruz (San Ignacio de Velasco; Rfo Quiser [=
Quizar]; Pailon, 8 km N and 1 km E; Pailon, 7 km
N of San Rafael de Amboro; hunting camp, 4 km
S and 13 km W of San Rafael de Amboro; Santa
Cruz, 27 km S; Chiquitos; Ingeniero Mora, 7 km
E and 3 km N; Basilio; Basilio, 3.4 km S; Santiago
de Chiquitos, 4 km N and 1 km W; Tita; Cerro
Colorado; Estancia Isibobo, 1 km S and 3 km W;
Camiri, near; Boyuibe, 53 km E; Cafeces, Monte
Grande; Santa Cruz de la Sierra, near); Tarija (Ca-
matindi; Estancia Bolivar). Mares and Braun, 2000:
35 ARGENTINA: Formosa (Reserva Natural
Formosa).
Monodelphis domestica domestica: Cabrera, 1958:
8 classification.
Hemiurus domesticus: Winge, 1893:8, 55 BRAZIL:
Minas Gerais (Lagoa Santa).
Peramys domesticus: Thomas, 1910c:502 BRAZIL:
Ceard (Ipti; Serra de Ibiapaba [= Serra Ibiapaba]).
Miranda Ribeiro, 1936:405 BRAZIL: Mato Gros-
so (Cuyaba [= Cuiaba], holotype); Ceard (Ipii);
Goids (Veadeiro [= Veadeiros]). Moojen, 1943:2
BRAZIL: Paraiba (Lagoa do Monteiro); Pernam-
buco (Pocao); Ceard (Crato).
[Monodelphis] domesticus: Matschie, 1916:272. Poh-
le, 1927:240 BRAZIL: Pernambuco (Dois Ir-
maos); Paraiba (Penha).
Hemiurus concolor Gervais, 1855:101. Gardner,
1993:21 listed.
\Monodelphis] concolor: Matschie, 1916:271 clas-
sification.
Hemiurus Hunteri: Gervais (not Waterhouse, 1841),
1855:101 BRAZIL: Goids.
Holotype In the Vienna, Munich, and British
Museums; lectotype, adult female, skin and skull
in Vienna Museum, collected March 1824 by Jo-
hann Natterer.
Type Locality Cuyaba [= Cuiaba], Mato
Grosso, Brazil.
Distribution (Fig. 65) Brazil to eastern and
central Bolivia, and northern Argentina.
Monodelphis emiliae (Thomas, 1912)
Peramys emiliae Thomas, 19 12a: 89. Gardner, 1993:
21 listed.
[Monodelphis] emiliae: Matschie, 1916:271 classi-
fication.
Monodelphis emiliae: Thomas, 1920b:283 BRA-
ZIL: Para (Vila Braga [= Villa Braga], Rio Tapa-
j6z). Pine and Handley, 1984:240 PERU: Loreto
(Quebrada Aucayo); BRAZIL: Para (Cameta; For-
dlandia; km 217, Santarem-Cuiaba Highway; Vila
Braga [= Villa Braga]; Flexal; km 212, Transa-
mazonian Highway between Itaibuba and Jacarea-
Canga). Patterson, 1992:6 BRAZIL: Para (Lago
Tapayuna). Anderson, 1997:160 BOLIVIA: Pan-
do (Centra Dieciocho). Patton et al., 2000:77 BO-
LIVIA: Pando (Centra Dieciocho); BRAZIL: Acre
(Igarape Porongaba, right bank Rfo Jurua; Nova
Vida, right bank Rfo Jurua); Amazonas (Seringal
Condor, left bank Rfo Jurua; Penedo).
P[eramys] tricolor emiliae: Miranda Ribeiro, 1936:
408 BRAZIL: Para (Villa Braga, Rio Tapajdz).
Monodelphis brevicauda emiliae: Carvalho, 1960:
122 BRAZIL: Para (Gradaus [= Gorotire], Rio
Fresco; Riosinho, Rio Fresco).
Monodelphis touan emiliae: Cabrera, 1958:9 clas-
sification.
Monodelphis brevicaudata: Thomas, 1920b:283
BRAZIL: Paid (Tapirinha, near Santarem).
Holotype Male, skin and skull, BMNH
1 1.12.22.16; collected 13 September 191 1 by Em-
ilia Snethlage.
Type Locality Boim, lower Rio Tapajoz,
Para, Brazil.
Distribution (Fig. 66) The Amazon basin of
Brazil, Peru, and northern Bolivia.
Monodelphis glirina (Wagner, 1842)
Didelphys glirina Wagner, 1842:359 BRAZIL: Ron-
donia (type locality Cachoeira do Rfo Grande, Rfo
Mamore).
D[idelphys[ glirina: Wagner. 1855:253.
[Monodelphis] glirina: Matschie, 1916:271 classifi-
cation.
Monodelphis glirina: Voss et al., 2001:54 BRAZIL
(S of the Amazon and W of the Xingu.
Peramys brevicaudatus: Miranda Ribeiro (not Erxe-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
111
90 80 70 60 50" 40 30 20 10
Fig. 65. Distribution of Monodelphis domestica: eastern Brazil west to eastern Bolivia and northern Argentina.
= localities given in published records.
1 12
FIELDIANA: ZOOLOGY
20
^O*
V-
s^r
'
\ -v..
Monodelphis emiliae
"V--.
10'
10'
20
Fig. 66. Distribution of Monodelphis emiliae: the Amazon basin of Brazil west to northeastern Peru. = localities
given in published records.
leben), 1936:406 BRAZIL: Rondonia (Cachoeira
do Rio Grande, Rio Mamore).
[Peramys brevicaudatus] brevicaudatus: Cabrera,
1919:42 glirina in synonymy only.
Monodelphis brevicaudata brevicaudata: Cabrera
(part, not Erxeleben), 1958:7 glirina in synonymy
only.
Holotype Male, skin and skull, Vienna Mu-
seum, collected September 1829 by Johann Nat-
terer.
Type Locality Cachoeira do Rio Grande, Rio
Mamore, Rondonia, Brazil.
Distribution South of the Amazon and west
of the Xingii in Brazil (no distribution map; lo-
cality not located).
Monodelphis iheringi (Thomas, 1888)
Didelphvs (Peramys) Iheringi Thomas, 1888a: 159
BRAZIL: Rio Grande do Sul (type locality, Ta-
quara; holotype, male, in alcohol, BMNH
82.9.30.43; presented by Hermann von Ihering).
Vieira, 1955:349 BRAZIL: Espirito Santo (Chav-
es). Avila Pires, 1987:123 BRAZIL: Rio Grande
do Sul (Taquara). Gardner, 1993:21 listed.
Didelphis iheringi: Thomas, 1888b:364 BRAZIL:
Rio Grande do Sul (Taquara).
Peramys Iheringi: Ihering, 1892:101 BRAZIL: Rio
Grande do Sul (Taquara do Mundo Novo [= Ta-
quara]).
Peramys iheringi: Shufeldt, 1926:563, 573, 574
BRAZIL: Sao Paulo (Serra da Cantareira).
[Microdelphys] iheringi: Matschie, 1916:272 clas-
sification.
Peramys iheringii: Miranda Ribeiro, 1936:415 in-
correct subsequent spelling classification.
Monodelphis iheringii: Vieira. 1953:357 incorrect
subsequent spelling BRAZIL: Santa Catarina;
Sao Paulo; Espirito Santo. Pine, 1977:235 BRA-
ZIL: Sao Paulo (Iguape); Santa Catarina (Joinvil-
le; Colonia Hansa [= Corupa]; Hamonia [= Ibira-
ma]; Ararangua). Mares and Braun, 2000:35 AR-
GENTINA: Misiones (Rib Victoria, not located;
Parque Nacional Iguazii).
Monodelphis americana iheringi: Cabrera, 1958:7.
Holotype Male, in alcohol, BMNH 82.9.30.43;
presented by Hermann von Ihering.
Type Locality Taquara, Rio Grande do Sul,
Brazil.
Distribution (Fig. 67) Southeastern Brazil
and Misiones in Argentina.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
113
20
20
^:
50
Monodelphis iheringi
-- 500 1000 1500 km
90
80
70
60
50
40
30-
20
10 ;
0=
Fig. 67. Distribution of Monodelphis iheringi: southeastern Brazil from state of Sao Paulo south to Rio Grande
do Sul and west to Misiones, Argentina. = localities given in published records.
Monodelphis kunsi Pine, 1975
Monodelphis kunsi Pine. 1975:321 BOLIVIA: Beni
("La Granja"); Tarija (Rio Lipeo). Mares et al..
1989:18 BRAZIL: Disthto Federal (Brasilia. 20
km S). Gardner. 1993:21 listed. Anderson. 1997:
160 BOLIVIA: Tarija (Tapecua; Rio Lipeo).
Holotype Adult male, skin, skull and skele-
ton, USNM 461348; collected July 1971 by Merle
L. Kuns.
Type Locality "La Granja."" Beni. Bolivia.
Distribution (Fig. 68) Known only from four
localities, three in Bolivia and one in Brazil.
Monodelphis maraxina Thomas, 1923
Monodelphis maraxina Thomas, 1923a: 157 BRA-
ZIL: Para (type locality. Caldeirao. Ilha de Marajo:
Soure. Ilha de Marajo). Pine. 1979:497 BRAZIL:
Para (Fazenda Tesa. Soure. Ilha de Marajo; Soure.
Ilha de Marajo). Gardner. 1993:21 listed.
Monodelphis domestica maraxina: Cabrera. 1958:8
classification.
Holotype Female, skin and skull, BMNH
28.8.9.9: collected 21 February 1923 by W. Ehr-
hardt.
Type Locality Caldeirao. Ilha de Marajo,
Para. Brazil.
Distribution (Fig. 69) Known only from the
Ilha de Marajo. mouth of Amazon. Para. Brazil.
Monodelphis osgoodi Doutt, 1938
Peramys peruvianus: Thomas (not Osgood. 1913).
1920:248 PERU: Cusco (Tocopoqueu, Ocam-
bamba Valley).
Monodelphis peruvianus osgoodi Doutt. 1938:100.
Monodelphis adusta osgoodi: Cabrera. 1958:6 clas-
sification.
Monodelphis osgoodi: Gardner. 1993:21 listed. An-
derson. 1997:160 BOLIVIA: Cochabamba (Cho-
re; Tablas Monte. 4.4. km N: Incachaca).
Holotype Adult male. Carnegie Museum
5242, collected 12 September 1921 by Jose Stein-
bach.
114
FIELDIANA: ZOOLOGY
Monodelphis kunsi
" ></
Fig. 68. Distribution of Monodelphis kunsi: known only from four widely scattered localities in northern and
extreme southern Bolivia and Distrito Federal, Brazil. = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
115
Monodelphis maraxina
80 70
Fig. 69. Distribution of Monodelphis maraxina: known only from Una de Marajo, Para, Brazil,
given in published records.
Type Locality Incachaca, Cochabamba, Bo-
livia, 2600 m.
Distribution (Fig. 70) Southeastern Peru and
central Bolivia.
Monodelphis palliolata (Osgood, 1914)
Peramys palliolatus Osgood, 1914a: 135 VENE-
ZUELA: Tdchira (type locality San Juan de Co-
lon). Gardner, 1993:21 listed.
Monodelphis palliolatus: Matschie, 1916:271 clas-
sification. Voss et al., 2001:54 COLOMBIA; VE-
NEZUELA.
Monodelphis (Monodelphis) palliotus: Anduze, 1 956:
10 VENEZUELA: Carabobo (San Esteban).
Monodelphis b. palliolata: Perez-Hernandez, 1989:
370 VENEZUELA: Distrito Federal (Los Cana-
les de Naiguata, Parque Nacional El Avila; Parque
Nacional "El Avila"; El Eden Parque Nacional "El
Avila"; Cerro El Avila; Corozal, Parque Nacional
El Avila; Hacienda El Limon; Carretera El Limon;
El Encantado, Hacienda El Limon; Suapo, Haci-
enda El Limon; La Conchita, Hacienda El Limon;
Zona Guacatal, Hacienda El Limon; Chichirivichi);
Barinas (Barinitas; Rib Barragan, Barinitas; Bar-
inita, 3 km SW); Carabobo (Bahio de Patenemo;
Valle Pascua, Patenemo; Los Campamentos, Pate-
nemo; Camino de los Espanoles, San Esteban; Los
Quigas, San Esteban; Pira Pira); Cojedes (El Can-
delo, Tucuragua); Merida (Zea; Bejuquero, 15 km
N of Tovar; Cano Tigre, Zea; Mina Carbon, Zea;
Hacienda San Pedro, Santa Cruz de Mora); Miran-
da (El Hatillo; La Union, El Hatillo; Targua; La
Providencia, Turgua; Los Guayabitos; Baruta; La
Cienega, Baruta; Cortada del Guayabo; La Toma
Capaya, Rib Marismita); Yaracuy (Agua Negra,
Yaritagua, 30 km N); Zulia (Kasmera, base of La
Serranfa de Perija; Estacion de Kasmera, Rfo Yasa).
Peramys brevicaudatus: Robinson and Lyon (not Erx-
eleben, 1777), 1901:138 VENEZUELA: Distrito
Federal (San Julian).
Monodelphis brevicaudata: Tate (not Erxeleben,
1777), 1923:65 VENEZUELA: Aragua ([Esta-
cion Biologico de] Rancho Grande). Sanderson,
1949:788 SURINAME. Pirlot, 1963:175, 178
VENEZUELA: Zulia (Kasmera, Rio Yasa). Han-
dley, 1976:6 VENEZUELA: Zulia (Kasmera, Rio
Yasa); Falcon (Mirimire); Guarico (Estacion Biol-
ogica de los Llanos); Carabobo-Falcon-Yaracuy
(Urama); Trujillo (Valera; La Ceiba); Barinas (Al-
tamira); Yaracuy (Boca de Yaracuy; Minas de
Aroa); Carabobo (Montalban); Aragua (Estacion
Biologica de Rancho Grande); Distrito Federal
116
FIELDIANA: ZOOLOGY
Monodelphis osgoodi I \
500 1000 1500 km
Fig. 70. Distribution of Monodelphis osgoodi: southeastern Peru and central Bolivia.
published records.
30
localities given in
(Hacienda Carapiche); Miranda (Curupao; Biron-
go; Rio Chico); Miranda-Guarico (Parque Nacion-
al Guatopo); Sucre (Manacal). Reig et al., 1977:
212 VENEZUELA: Zulia (Kasmera, Rio Yasa);
Aragua ([Estacion Biologica dej Rancho Grande);
Merida (Hacienda San Pedro, Santa Cruz del
Moro). Eisenberg et al., 1979:187 VENEZUELA:
Miranda-Guarico (Parque Nacional Guatopo).
[Monodelphis brevicaudata] brevicaudata: Avila-Pi-
res, 1964:10 (in text) VENEZUELA: Aragua
([Estacion Biologica de] Rancho Grande).
Monodelphis brevicaudata orinoci: Anduze (not
Thomas, 1899), 1956:10 VENEZUELA: Distrito
Federal (La Florida). Reig and Bianchi, 1969:
210 VENEZUELA: Aragua ([Estacion Biologica
de] Rancho Grande).
Holotype Male, skin and skull, FMNH
20524, collected 14 November 1913 by M. P. An-
derson.
Type Locality San Juan de Colon, Tachira,
Venezuela, altitude 180 m.
Distribution (Fig. 71) North and west of Rio
Orinoco in northern Venezuela.
Monodelphis rubida (Thomas, 1899)
Didelphvs brevicaudata: Thomas, 1888b:356 part,
BRAZIL: Bahia.
Peramys rubidus Thomas, 1899a: 155. Gardner, 1993:
22 listed.
[Monodelphis] rubidus: Matschie, 1916:271 classi-
fication.
P[eramys] tricolor rubidus: Miranda Ribeiro, 1936:
408 classification.
Monodelphis tricolor rubidus: Vieira, 1955:350 list-
ed.
Monodelphis touan rubidus: Cabrera, 1958:10 clas-
sification.
M[inuania[ goyana Miranda Ribeiro, 1936:419 ap-
parent lapsus for M. umbristriata. Gardner, 1993:
22 listed.
Minuania umbristriata Miranda Ribeiro, 1936:422,
424 BRAZIL: Golds (type locality, Veadeiros;
holotype, male, skin and skull, Museu Nacional,
Rio de Janeiro, purchased from Jose Blaser). Gard-
ner, 1993:22 listed.
Monodelphis umbristriatus: Vieira, 1955:350. Pine,
1976:785 BRAZIL: Goids (Veadeiros); Minas
Gerais (Passos).
Monodelphis americana americana: Cabrera, 1958:
BROWN: ATLAS OF NEW WORLD MARSUPIALS
117
20'
Monodelphis palliolata
1500 km
80" 70 60 5C W 30
Fig. 71. Distribution of Monodelphis palliolata: northern Venezuela. = localities given in published records.
6 part? (not Miiller, 1 776), umbristriata question-
ably included in synonymy.
Holotype Male, skin and skull, BMNH
55.11.26.9.
Type Locality Bahia, Brazil.
Distribution (Fig. 72) In eastern Brazil from
Goias south to Minas Gerais.
Monodelphis scalops (Thomas, 1888)
Didelphys (Peramys) scalops Thomas, 1888a: 158.
Gardner, 1993:22 listed.
Didelphys scalops: Thomas, 1888b:359. Pohle, 1927:
240 BRAZIL: Rio de Janeiro (type locality re-
stricted to Rio de Janeiro; Teresopolis). Leuder-
waldt, 1929:27 BRAZIL: Sao Paulo.
[Monodelphis] scalops: Matschie, 1916:271 classi-
fication.
Monodelphis scallops: Cabrera, 1958:9. Vieira, 1949:
360 BRAZIL: Sao Paulo (Sao Sebastiao; Piraci-
caba; Piquete). Pine, 1978:380 BRAZIL: Sao
Paulo (Piquete; Piracicaba; Sao Sebastiao); Espir-
ito Santo (Caixa D'Agua; Forest Preserve Nova
Lombardia); Rio de Janeiro (Teresopolis). Mares
and Braun, 2000:35 ARGENTINA: Misiones
(Parque Nacional Iguazii).
[Peramys] scalops: Cabrera, 1919:43 classification.
Peramys scalops: Miranda Ribeiro, 1935:38 BRA-
ZIL: Rio de Janeiro (Teresopolis). Miranda Ribei-
ro, 1936:412 BRAZIL: Rio de Janeiro (type lo-
cality, Teresopolis). Davis, 1947:3 BRAZIL: Rio
de Janeiro (Teresopolis). Vieira, 1949:360 BRA-
ZIL: Sao Paulo. Vieira, 1955:349 BRAZIL: Sao
Paulo (Sao Sebastiao; Piracicaba; Piquete); Rio de
Janeiro (Teresopolis).
Holotype Male, skin and skull, BMNH
51.7.21.23 (skin), 51.8.30.10 (skull); purchased
from Lord Derby.
Type Locality Brazil; restricted to Rio de Ja-
neiro by Pohle (1927:240) and further restricted
to Teresopolis by Vieira (1955:349).
Distribution (Fig. 73) Southeastern Brazil
from Espfrito Santo to Santa Catarina and Mi-
siones, Argentina.
Monodelphis sorex (Hensel, 1872)
Microdelphys sorex Hensel, 1872:122. Gardner, 1993:
22 listed.
118
FIELDIANA: ZOOLOGY
90 80 70 60 50
Fig. 72. Distribution of Monodelphis rubida: eastern Brazil, south from Goias to Minas Gerais. = localities
given in published records.
Didelphis sorex: Thomas, 1888b:362 BRAZIL: Rio
Grande do Sul (Taquara; San Lorenzo [= Sao Lou-
renco do Sul]).
Peramys sorex: Ihering, 1892:101 BRAZIL: Rio
Grande do Sul. Bertoni, 1939:6 PARAGUAY:
Alto Parana (Puerto Bertoni).
[Monodelphiops] sorex: Matschie, 1916:261, 267,
272 listed.
Monodelphis sorex: Cabrera, 1958:9 classification.
Mares and Braun, 2000:36 ARGENTINA: Mi-
siones (Parque Nacional Iguazu; Reserva Natural
Estricta San Antonio).
Didelphys {Peramys) Henseli Thomas, 1888a: 159
BRAZIL: Rio Grande do Sul (type locality, Ta-
quara; holotype, female in spirits BMNH
82.9.30.37 collected by H. von Ihering). Vieira,
1955:349 BRAZIL: Rio de Janeiro (Serra de Ma-
cae). Avila Pires, 1994:369 BRAZIL: Rio Grande
do Sul (Taquara). Gardner, 1993:22 listed.
Didelphys henseli: Thomas, 1888b:360 BRAZIL:
Rio Grande do Sul (Taquara); ARGENTINA: Entre
Ri'os (no specimens recorded).
Peramys Henseli: Ihering, 1892:100 BRAZIL: Rio
Grande do Sul.
Peramys henseli: Bertoni, 1914:69 PARAGUAY:
Alto Parana (Puerto Bertoni). Bertoni, 1939:7
PARAGUAY: Alto Parana (Puerto Bertoni).
Peramys henselii: Miranda Ribeiro, 1936:409 incor-
rect subsequent spelling BRAZIL: Rio Grande do
Sul.
[Monodelphis] henseli: Matschie, 1916:271 classi-
fication.
Monodelphis henseli: Cabrera, 1958:9 classification.
Pine, 1979:496 BRAZIL: Minas Gerais (Passos);
Sao Paulo (Casa Grande, Boraceia; Ribeirao da La-
goa); Rio de Janeiro (Campo Bello). Azevedo et
al., 1982:94 BRAZIL: Santa Catarina (Seminario
Coracao de Jesus, Corupa).
[Monodelphis] lundi Matschie. 1916:271 BRAZIL:
Minas Gerais (type locality, Rio das Velhas, Lagoa
Santa); name based on specimens recorded as D.
tricolor by Lund (1841:236; 1842:135). Gardner,
1993:22 listed.
P[eramys (Minuania)] d[imidiata] itatiayae Miranda
Ribeiro, 1936:421 BRAZIL: Rio de Janeiro (type
locality, Campo Bello, Itatiaya; holotype, male,
skin and skull, MZUSP; collected by Senhor Ziken
and presented by Senhor E. May). Gardner, 1993:
22 listed.
Monodelphis tricolor paulensis Vieira, 1949:359
BRAZIL: Sao Paulo (Pirituba; Piquete; Mogi das
Cruzes; Ibiti [= Monte Alegre do Sul]). Vieira,
1955:380 BRAZIL: Sao Paulo (type locality, Pir-
ituba, outside Sao Paulo; Capital; Mogi das Cruzes;
Monte Alegre [ = Monte Alegre do Sul]; Piquete;
Ibiti [ = Monte Alegre do Sul]; holotype, male, skin
BROWN: ATLAS OF NEW WORLD MARSUPIALS
119
90 80 70 60 50 40 30 20 10
Fig. 73. Distribution of Monodelphis scalops: southeastern Brazil from Espirito Santo south to Santa Catarina.
= localities given in published records.
and skull, Departamento de Zoologia, Secretaria de
Agricultura, Sao Paulo, Brazil, no. 2557, collected
13 January 1908 by L. P. Barreto). Gardner, 1993:
22 listed.
Monodelphis touan paulensis: Cabrera, 1958:9 clas-
sification.
Syntypes Male and female, Berlin Zoological
Museum; collected by R. Hensel.
Type Locality Taquara, Rio Grande do Sul,
Brazil.
Distribution (Fig. 74) Southeastern Brazil,
southeastern Paraguay, and northeastern Argenti-
na.
Monodelphis theresa Thomas, 1921
Monodelphis theresa Thomas, 1921d:441 BRAZIL:
Rio de Janeiro. Gardner, 1993:22 listed.
Peramys theresae Miranda Ribeiro, 1935:39 incor-
rect subsequent spelling BRAZIL: Rio de Janeiro
(Teresopolis).
Peramys therezae: Miranda Ribeiro, 1936:417 in-
correct subsequent spelling BRAZIL: Rio de Ja-
neiro (Teresopolis).
Monodelphis americana theresa: Cabrera, 1958:7.
Holotype Skin (from alcohol preserved fe-
male, skull, BMNH 21.8.6.2, collected by Prof. J.
P. Hill).
Type Locality Serra dos Orgaos, Rio de Ja-
neiro, Brazil.
Distribution (Fig. 75) Southeastern Brazil.
Monodelphis unistriata (Wagner, 1842)
Didelphys unistriata Wagner, 1842:360. Pelzeln,
1883:1 16 BRAZIL: Sao Paulo (Itarare). Gardner,
1993:22 listed.
D[idelphys] unistriata: Wagner, 1855:249 charac-
ters.
Microdelphys unistriata: Burmeister, 1856:87.
Peramys unistriata: Ihering, 1894:11.
Monodelphis unistriatus: Vieira, 1949:358.
Holotype Male, in Vienna Museum; collect-
ed 11 March 1821 by Johann Natterer.
Type Locality Itarare, Sao Paulo, Brazil.
120
FIELDIANA: ZOOLOGY
20
30
40
50
Monodelphis sorex
1500 km
90"
80'
70
60
50"
40
30
20
10"
Fig. 74. Distribution of Monodelphis sorex: southeastern Brazil, northeastern Argentina, and southern Paraguay.
= localities given in published records.
Distribution (Fig. 76) Known from type lo-
cality only.
Genus Philander Brisson, 1762
Philander Brisson, 1762:13, 207-214 name pub-
lished in rejected work, subsequently conserved by
the International Commission on Zoological No-
menclature in the Bulletin of Zoological Nomen-
clature (1998:64) and type species designated as
Didelphis opossum Linnaeus. Tiedemann, 1808:
426 included species, P. virginianus Tiedemann
[= D. opossum Linnaeus], P. murinus [= Marmosa
murina], P. brachyurus [= Monodelphis brachyu-
ra\. Gilmore, 1941:316 Metachirops Matschie
antedated; distribution. Hopwood, 1947:533
names from Brisson not valid; Philander Gronov-
ius non-Linnaean and not available (see Bulletin of
Zoological Nomenclature, 51 [2]: 135-138). Hersh-
kovitz, 1949:11 type, P. virginianus Tiedemann
= Didelphis opossum Linnaeus; Philander Brisson
1762, and Philander Gronovius, 1763, not valid.
Pine, 1973a:391 not the four-eyed pouched opos-
sum. Hershkovitz, 1976:295 history, nomencla-
ture. Husson, 1978:27 treated as synonym of Di-
delphis Linnaeus by designation of the Virginia
opossum as lectotype of type species Philander vir-
ginianus Tiedemann. Hershkovitz 1981:943 Phi-
lander valid generic name; type P. virginianus Tie-
demann = Didelphis opossum Linnaeus. Gardner,
1981:447 Philander valid; Husson opinion re-
jected. Jenkins and Kurtson, 1 983 type specimens
in Natural History Museum, London. Gardner,
1993:22 synonymy; species (P. opossum, P. an-
dersoni) distribution.
Sarigua Muirhead, 1819:429 part, 5. opossum (=
Didelphis opossum Linnaeus) listed among includ-
ed species.
Metachirops Matschie, 1916:262, 267, 268 included
species, pallidus Allen, fuscogriseus Allen, grises-
cens Allen, melanurus Thomas, opossum Linnaeus,
canus Osgood, andersoni Osgood, quica Tem-
minck (designated type), frenata Lichtenstein; tax-
onomy. Miranda Ribeiro, 1936:340. Krumbiegel,
194 lb: 199 review. Pine, 1973a:391 valid ge-
neric name for four-eyed pouched opossum; Phi-
lander regarded a synonym of Didelphis.
Holothylax Cabrera, 1919:47 type, Didelphis opos-
sum Linnaeus, by original designation.
Metachirus Burmeister, 1854:135 subgenus of Di-
delphys [= incorrect spelling of Didelphis] Linnae-
us, part, D. quica Temminck only. Burmeister,
1856:68 part, M. opossum Linnaeus only. Thom-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
121
20
30'
40
50
Monodelphis theresa
500 1000 1500 km
90 80 70 60 50 40
Fig. 75. Distribution of Monodelphis theresa: southeastern Brazil.
30 20 10
localities given in published records.
as, 1888b:329 part, Didelphys opossum Linnaeus
only.
Metacheirus Sanderson, 1949:787 incorrect subse-
quent spelling of Metachirus, in combination with
M. opossum.
Type Species Didelphis opossum Linnaeus,
by designation of International Commission on
Zoological Nomenclature (1998:64).
Distribution (Figs. 77 and 78) Tropical and
subtropical forests including second growth, Cen-
tral and South America, from Mexico south to
Misiones and Chaco in Argentina and western
Peru to southeastern Brazil; altitudinal range from
sea level to about 2000 m above.
Philander andersoni (Osgood, 1913)
[l]D[idelphys]. myosuros: Tschudi (not Temminck,
1824), 1844:14, 145, 151 PERU. Tschudi, 1844:
250 PERU.
Metachirus andersoni Osgood, 1913:95. Osgood,
1914:149 PERU: Ucayali ("Yane Yacu" [= Yan-
ayacu, now in Loreto]; Rio Paranapura, 8 mi from
Yurimaguas [now in Loreto]). Sanborn, 1947:
215 type history.
M[etachirops] andersoni: Matschie, 1916:268 clas-
sification.
[Holothylax] andersoni: Cabrera, 1919:47 classifi-
cation.
Philander andersoni: Patton and da Silva, 1997:
101 ECUADOR: Napo (Parque Nacional Yasuni,
Onkone Sur, 38 km S of Pompeya Sur); PERU:
Amazonas (Huampami, Rio Cenepa); Loreto (San
Jacinto, Rio Tigre). Patton et al., 2000:82 EC-
UADOR: Napo (Okone Gare, 38 km SE of Pom-
peya Sur, Parque Nacional Yasuni'); PERU: Ama-
zonas (Huampami, Rio Cenepa); Loreto (San Ja-
cinto, Rio Pastaza); BRAZIL: Amazonas (Comu-
nidade Colina, right bank Rio Tiquie, Municipio
San Gabriel de Cachoeira; Rio Jau, above mouth
right bank; Macaco, left bank Rio Jau).
M[etachirus] opossum andersoni: Thomas, 1923c:
603 classification.
Metachirus opossum andersoni: Thomas, 1928b:
294 PERU: Loreto (Iquitos).
Philander opossum andersoni: Cabrera, 1958:34.
Perez-Hernandez, 1989:373 VENEZUELA: Ama-
zonas (Cano Majagua; Cano Yagua, Cerro Cucuri-
to; Rio Manapiare, San Juan de Manapiare; Ta-
matama; Rio Cunucunuma, Belen; Rio Casiquiare,
Capibara; Campamento La Neblina, Cerro La Neb-
lina; Mavaca, Alto Rio Orinoco; Sierra Parima).
P[hilander] opossum andersoni: Gardner and Patton,
1972:5 PERU: Ayacucho (Huanhuachayo; San
Jose).
122
FIELDIANA: ZOOLOGY
20'
30
40
50'
Monodelphis unistriata
1500 km
7\
90 1
80"
70
60
50
40
30
20
10
Fig. 76. Distribution of Monodelphis unistriata: known only from type locality in state of Sao Paulo, Brazil.
localities given in published records.
P\hilander] o[possum] andersoni: Baker, 1974:135
ECUADOR: Napo (Santa Cecilia).
Philander andersoni andersoni, Hershkovitz, 1997:
61, 85-91 COLOMBIA: Putumayo (Puerto Asis);
Caquetd (Tres Troncos); ECUADOR: Napo (San
Jose, abajo; Puerto Napo, Rio Napo; Santa Cecilia);
Pastaza (Rio Yana Rumi; Sarayacu; Montalvo; Rio
Copataza; Rio Pindo Yacu); PERU: Ucayali ([the
following localities are now in Loreto] Rio Curar-
ay, boca; Lago Miranes, Rio Napo; Rio Mazan,
boca; Santa Luisa; Orosa; Nauta; Santa Elena, Rio
Samiria; Yurimaguas, Rio Huallaga; Yana Yacu [=
Yanayacu], Iquitos; San Jeronimo); Ucayali (Cu-
meria [= Cumaria]); Amazonas (La Poza, Rio San-
tiago; Huampami, 5 mi W, Rio Cenepa); Pasco
(Oxapampa, Inanez); Junin (Chanchamayo; Utcuy-
acu); PERU: Ayacucho (San Jose, Huanhuachayo
[Hershkovitz placed these specimens from Ayacu-
cho under P. andersoni andersoni, but Gardner
placed them under P. opossum]); VENEZUELA:
Amazonas (Acanana, Rio Cunucunuma; Playa del
Rio Base, Cerro Duida; Esmeralda; Merey; Belen;
San Juan; Tamatama; Cano Majagua; Cerro Cucur-
ito; Sierra Parima; Capibara; Mavaca; Campamento
La Neblina).
Metachirus opossum Tate, 1939:161 VENEZUELA:
Amazonas (Playa del Rio Base, Cerro Duida).
Philander opossum: Gardner and Patton, part (not
Linnaeus, 1758), 1972:5 PERU: Ayacucho (San
Jose, Rio Santa Rosa, 1000 m; Huanhuachayo,
1660 m); comparisons. Reig et al., 1977:197
PERU: Ayacucho (San Jose, Rio Santa Rosa; Huan-
huachayo).
Metachirus opossum nigratus Thomas, 1923c:603
PERU: Junin (type locality, Utcuyacu, 1600 m;
Chanchamayo; holotype, male, skin and skull,
BMNH 1900.7.7.62, collected 21 April 1900 by P.
O. Simons). Thomas, 1928:250, footnote PERU:
Junin (Inanez, not located). Gardner and Patton,
1972:5.
Philander opossum canus: Cabrera (part, not Osgood,
1913) 1958:35.
Holotype Male, skin and skull, FMNH
19655; collected 11 September 1912 by M. P. An-
derson.
Type Locality "Yane Yaku" [= Yanayacu],
a chacra or farm on the Paranapura River about
8 mi from Yurimaguas, Ucayali, Peru (now in
Loreto, Peru). The latter is on the Rio Huallaga
at the mouth of the Rio Paranapura (cf. Osgood,
1914, p. 147).
Distribution (Fig. 79) Southern Venezuela,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
123
Fig. 77. Distribution of the genus Philander in Central America: north-central Mexico south through Panama.
See Figure 78 for distribution in South America.
southern Colombia, Ecuador, Peru, and north-
western Brazil.
Additional Records ECUADOR: Napo; Li-
moncocha (Emmons, 1977, USNM); Llunchi, Rio
Napo (Hershkovitz, 1936, UMMZ); Zancudo
(Timm, 1983, FMNH); Pastaza; Intillama (= In-
dillama), Rio Napo (Hershkovitz, 1936, UMMZ);
PERU: Amazonas; Wanuma, Adelbert Mts., not
located (Patton, 1977, MVZ); Loreto; Quebrada
Oran (Barkley, 1984, LSUMZ); Quisto Cocha [=
Quistococha]; Maynas (McGrath, 1980, KU).
Philander frenata (Olfers, 1818)
D[idelphis] frenata Illiger, 1815 (1 804- 1 8 1 1 ): 1 07
nomen nudum.
D[idelphys] frenata Olfers, 1818:204 listed.
[Metachirops] frenata: Matschie, 1916:268 BRA-
ZIL: Bahia (type locality originally given as Brazil,
according to Wagner [1843, footnote 25, p. 44], the
specimen labeled D. frenata in the Berlin Museum
was collected by Herr Koehne in Bahia, presum-
ably in the present Sao Salvador, which may be
taken as the restricted type locality).
Met[achirops] opossum frenatus: Krumbiegel, 1941:
206 (in text) incorrect subsequent spelling.
Philander opossum frenatus: Olfers, 1818:201 in-
correct subsequent spelling.
Philander opossum frenata: Hershkovitz, 1959:338,
342 taxonomy; part, synomyms (superciliaris Ol-
fers, myosuros Temminck). Hershkovitz, 1997:51,
93 BRAZIL: Bahia (Bahia [= Salvador]).
Philander frenata: Patton and da Silva, 1997:97-
101 BRAZIL: Parana; Bahia; Goias; Espirio
Santo (Estacao Biologica Santa Lucia. Santa Tereza
[= Teresa]; Mato do Larva d'Agua, 1.7 km from
Santa Tereza); Minas Gerais (Parque Estadual do
Ibitipoca; Fervedouro); Sao Paulo (Fazenda Inter-
vales, Capao Bonito; Fazenda da Toca, Ilha de Sao
Sebastiao, Ilhabela; Praia do Felix, Uratuba; Esta-
cao Biologica Boraceia; Serra do Japi, 7 km W of
Jundiaf); Rio de Janeiro (Parque Nacional de Ita-
tiaia, Penedo, Municipio de Itatiaia; Marica; Gar-
rafao, Estacion Rio Terezopolis [= Terasopolis],
Maje). Gentile et at, 1997:105 BRAZIL: Rio de
Janeiro (Barra de Marica). Patton et al., 2000:82
BRAZIL: Espirito Santo (Estacao Biologica Santa
Lucia, Santa Tereza); Rio de Janeiro (Garrafao, Es-
tacion Rio Terezopolis [= Teresopolis], Maje); Sao
Paulo (Fazenda Intervales, Capao Bonito); Minas
Gerais (Parque Estadual do Ibitipoca, 30 km N of
Lima Duarte).
D[idelphys] superciliaris Illiger, 1815(1804-1811):
107 nomen nudum. Olfers, 1818:204 type lo-
cality "Sudamerica".
Didelphis myosuros Temminck, 1824:38 BRAZIL:
(type locality); syntypes in the Leiden, Vienna,
Frankfurt, and Maximiliam Zu Wied-Neuwied Mu-
seum; female syntype in the Vienna Natural His-
tory Museum, collected 3 March 1818 by Johann
124
FIELDIANA: ZOOLOGY
Fig. 78. Distribution of the genus Philander in South America: throughout northern South America south to
northeastern Argentina; in Brazil west of the Rio Tocantins-Araguaia, with the exception of Bahia. See Figure 77 for
distribution in Central America.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
125
20
10
1500 km
30
Fig. 79. Distribution of Philander andersoni: southern Venezuela, southern Colombia, Ecuador, Peru, and north-
western Brazil, o = localities from unpublished records; = localities given in published records.
Natterer, designated lectotype by Hershkovitz
(1959:34). Hershkovitz, 1959:343, in synonymy of
Philander opossum frenata Olfers.
Didelphis myosurus: Wagner, 1843:footnote 25, p.
44 incorrect subsequent spelling of myosuros
BRAZIL: Bahia; type of "frenata Lichtenstein"
from Bahia; collected by Kaehne.
Metachirus nudicaudatus myosurus, Miranda Ribeiro,
1936:345 incorrect subsequent spelling of myo-
suros BRAZIL: Bahia.
Didelphis quica Temminck, 1824:36. Desmarest,
1827:387. Jentink, 1887:301 BRAZIL: Tem-
minck collection. Jentink, 1888:200 part, BRA-
ZIL; syntype.
Didelphys quica: Waterhouse, 1841:90 part, BRA-
ZIL. Waterhouse 1 846:480 part, BRAZIL. Bur-
meister, 1854:136 BRAZIL: Rio de Janeiro
(Novo Friburgo). Pelzen, 1883:1 10 BRAZIL: Rio
de Janeiro (Sepetiba); Sao Paulo (Ipanema [ =
Ypanema]).
D[idelphys] quica: Wagner, 1855:225. Giebel, 1859:
227 "der Guica."
Metachirus quica: Burmeister, 1856:70 part, BRA-
ZIL: Rio de Janeiro (Novo Friburgo). Burmeister,
1856:70 (part, not Temminck) BRAZIL: Bahia;
Rio de Janeiro (Novo Friburgo). Hensel, 1872:
120 BRAZIL: Rio Grande do Sul. Goeldi, 1894:
460 Rio de Janeiro (Serra dos Orgaos). J. A. Al-
len, 1900:195 BRAZIL: (type locality, coast re-
gion of Brazil, just south of Rio de Janeiro).
[Holothylax] quica: Cabrera, 1919:48 classification.
Metachirus opossum quica: J. A. Allen, 1916b:562
BRAZIL: Rio de Janeiro (Sapitiba [= Sepetiba],
restricted type locality).
[Metachirops] quica: Matschie: 1916:268 BRAZIL:
Rio de Janeiro (type locality "Sapitibi").
Metachirops quica: Pohle: 1927:243 BRAZIL: Rio
de Janeiro (Barreira; Teresopolis). Schirch, 1932:
85 BRAZIL: Rio de Janeiro (Teresopolis).
Metachiropos opossum quica: Miranda Ribeiro, 1936:
340 BRAZIL: Santa Catarina (Porto Feliz, Rio
Urugai, not located); Rio de Janeiro (Teresopolis).
Vieira, 1949:345 BRAZIL: Sao Paulo; Minas
Gerais; Espirito Santo. Ruschi, 1965:2 BRAZIL:
Espirito Santo (coast).
Metachirus opossum quichua: Krumbiegel, 1941b:
200, 201, 206 incorrect subsequent spelling for
quica Temminck BRAZIL: Sao Paulo (Juquia,
Serra do Mar).
Philander opossum quica: Perodini and Perodini,
1966:28 BRAZIL: Sao Paulo (Cantareira Forest-
al Preserve, Sao Paulo). Carvalho, 1965:251
BRAZIL.: Sao Paulo (Boraceia). Correa Gomez,
1984:369 BRAZIL: Mato Grosso do Sul (Salob-
ra). Da Fonseca and Redford, 1984:517 BRAZIL:
Distrito Federal (Parque Nacional Brasilia). Alho
126
FIELDIANA: ZOOLOGY
et al., 1986:447, 451, 452 BRAZIL; humid galley
forest and valley side wet campo; cerrado [un-
known]). Hershkovitz, 1997:86-95 BRAZIL:
Goids (Aragarcas; Formosa; Anapolis; Trinidade);
Distrito Federal (Parque Nacional de Brasilia; Bra-
silia; Fazenda Agua Limpa); Minas Gerais (Con-
ceicao do Mato Dentro; Fazenda Esmeralda; Fa-
zenda Montes Claros; Lagoa Santa; Quartel do Sac-
ramento; Fazenda da Floresta, Rio Matipo; Boa Es-
peranca, Serra de Caparao; Fazenda Cardosa, Serra
de Caparao; Alem Paraiba; Juiz do Fora; Benfica,
Serra de Itatiaia [see Benfica, Rio de Janeiro]); Es-
pirito Santo (Santa Teresa; Sao Joao de Petropolis;
Vale Verde, Parque Nacional do Caparao; Pedra
Roxa, Parque Nacional de Caparao; Cachoeira
Bonita, Parque Nacional de Caparao; Serra; Cam-
pinho; Vitoria; Vila Velha, Morro de Angoles;
Caixa D'Agua; Valao de Sao Lourenco; Engenhei-
ro Reeve [= Rive]); Rio de Janeiro (Fazenda Boa
Vista; Rio de Janeiro; Fazenda Boa Fe [= Tereso-
polis]; Rodeio, Serra do Mar; Sao Joao Marcos;
Novo Friburgo; Barreira; Serra dos Orgaos; Monte
Serrat [= Pico das Agulhas Negras], Parque Na-
cional de Itatiaya [= Itatiaia]; Maceira, Itatiaia; Se-
petiba; Mangaratiba; Barro Branco; Pedra Branca,
Parati; Bemfica [= Benfica]; Sao Paulo (Serra de
Itatiaia [Collector E. G. Holt, June 1922, gives co-
ordinates as 2141'S, 4326'W in Minas Gerais.
Paynter and Tray lor (1991) place Benfica in Rio de
Janeiro]; Rio de Janeiro (Parque Nacional de Ita-
tiaia); Sao Paulo (Avanhandava, Lajeado; Bora-
ceia; Iporanga, Lajeado; Monte Alegre, Amparo [=
Monte Alegre do Sul]; Sao Paulo; Cotia; Serra da
Cantareira; Butanta; Costao dos Engenhos; Alto da
Serra; Vila Oliveira, Mogi das Cruzes; Salesopolis;
Casa Grande; Ponta da Boraceia; Piquete; Sao Se-
bastiao; Serra de Itatiaia; Ipanema; Juquia); Parana
(Rio Paracaf; Roa Nova); Santa Catarina (Join-
ville; Hansa [= Corupa]).
Didelphys opossum: Goldfuss, 1809:215. Waterhouse,
1841:90 part, BRAZIL. Waterhouse, 1846:485
part, BRAZIL. Thomas, 1888b:329 BRAZIL: Rio
Grande do Sul (Taquara). Winge, 1893:7, 38
BRAZIL: Minas Gerais (Lagoa Santa, Recent and
fossil).
Metachirus opossum: Ihering, 1892:99 BRAZIL:
Rio Grande do Sul. Ihering, 1894:10 BRAZIL:
Sao Paulo. Thomas, 1902d:64 BRAZIL: Parand
(Ro?a Nova, Serra do Mar).
M[etachirus] opossum: Miranda Ribeiro, 1905:189
BRAZIL: Rio de Janeiro (Monte Serrat, Parque
Nacional do Itatiaia [= Pico das Agulhas Negras]).
Philander opossum: Davis, 1945a: 122 BRAZIL:
Rio de Janeiro (Fazenda Boa Fe; Teresopolis).
Mello and Moojen, 1979:29 BRAZIL: (Bom Jar-
din, gallery forest); Distrito Federal (Brasilia, gal-
lery forest; Fazenda Agua Limpa, gallery; Guara,
gallery, not located).
Philander opossum opossum: da Fonseca and Kierulff
(not Linnaeus, 1758), 1989:118, 143 BRAZIL:
Minas Gerais (Fazenda Esmeralda; Fazenda Mon-
tes Claros).
Didelphis nudicaudatus: Desmarest (part, not E.
Geoffrey, 1803), 1827:390 myosuros Temminck
a synonym. Waterhouse, 1841:94. Jentick, 1888:
220 part, syntypes of myosuros Temminck and
opossum of Temminck collection. Thomas, 1 888b:
332 part, myosuros in synonymy.
Metachirus nudicaudatus nudicaudatus: Cabrera
(part, not E. Geoffrey, 1803), 1958:38 "frenata
Lichtenstein (Wagner)" in synonymy.
Didelphis larvata Jentink, 1888:220 Natterer MS
name on label of quica Temminck.
Didelphis larvata in MS name in synonymy. Avila-
Pires and Souvea, 1977:9 BRAZIL: Rio de Ja-
neiro (Monte Serrat [= Pico das Agulhas Negras];
Maceira [= Maceiras = Pico das Agulhas Negras];
Parque Nacional Itatiaia [= Pico das Agulhas Ne-
gras]); Minas Gerais (Benfica).
Holotype Adult female, skin mounted, Berlin
Zoological Museum, collected before 1815 by
Herr Kaehne.
Type Locality "Sudamerica"; restricted to
Bahia, Brazil, by Wagner, 1843:44, footnote 25,
restricted to Sepetiba, Rio de Janeiro, by Pelzen,
1883:110.
Distribution (Fig. 80) Coastal region of east-
ern and southeastern Brazil and from Goias to Pa-
rana.
Additional Records BRAZIL: Rio de Janei-
ro; Floresta da Tijuca (G.I.P., 1945, 1946, KU);
Espirito Santo; Pedra Roxa, Parque Nacional do
Caparao (Hershkovitz, 1992, FMNH); Minas Ger-
ais; Rio Matipo (Fonseca, 1919); San Francisco
de C, not located (Robert, 1960, BMNH); Sao
Paulo; Iguape (de Souza-Lopes, 1976, USNM);
Banagena, not located (1966, 1967, 1968,
MZUSP); Parque Estadual de Ilhabella; Ilha de
San Sabastiao (Mustrangi, 1993, MVZ).
Philander mcilhennyi Gardner and Patton,
1972
Philander mcilhennyi: Gardner and Patton, 1972:2
PERU: Ucayali (Balta, Rio Curanja, upper Rio Pu-
rtis). Reig et al., 1977:197, 212 PERU: Ucayali
(Balta, Rio Ucayali). Hutterer et al., 1995:10
PERU: Huanuco (Panguana, Rfo Pachitea). Patton
et al., 1995:7, 20, 21 BRAZIL: Amazonas (Alto
Rio Urucu). Emmons and Feer, 1997:8 PERU:
Ucayali (Balta); BRAZIL: Acre; Amazonas. Patton
and da Silva, 1997:97, 101 PERU: Ucayali (Bal-
ta; Pucallpa; Pucallpa, 59 km SW of and 59 km W
of); Huanuco (Panguana); Loreto (Jenaro Herrera
Biological Station; Rio Yarapa); BRAZIL: Acre
(Igarape Porongaba, right bank Rfo Jurua; Seringal
Sobral, left bank Rio Jurua); Amazonas (Seringal
Penedo, right bank Rfo Jurua; Seringal Altamira;
right bank Rfo Jurud; Alto Rfo Uriicu; Seringal
Condor, left bank Rfo Jurua). Patton et al., 2000:
83 BRAZIL: Acre (Igarape Porongaba, right bank
Rfo Jurua; Igarape Porongaba, opposite, left bank
Rfo Jurua; Nova Vida, right bank Rfo Jurua; So-
bral, left bank Rfo Jurua); Amazonas (Seringal
BROWN: ATLAS OF NEW WORLD MARSUPIALS
127
20 10
Fig. 80. Distribution of Philander frenata: in Brazil from Parana east to Bahia and south to Santa Catarina. o
localities from unpublished records; = localities given in published records.
128
FIELDIANA: ZOOLOGY
Fig. 81. Distribution of Philander mcilhennyi: from eastern Peru east to Acre and Amazonas in Brazil.
localities from unpublished records; = localities given in published records.
o =
Condor, right bank Rib Jurua; Penedo, right bank
Rio Jurua; Altamira, right bank Rio Jurua; Alto Rio
Urucu, Tefe).
Philander mcilhenni: Engstrom and Gardner, 1988:
232 incorrect subsequent spelling of mcilhennyi.
Philander andersoni mcilhennyi: Hershkovitz, 1997:
87 PERU: Ucayali (Balta, Rio Curanja; Pucallpa,
Rio Ucayali).
Holotype Adult female, skin and skull,
LSUMZ 16395; collected 10 April 1971 by Alfred
L. Gardner, original number 1 1 54 1 .
Type Locality Balta, Rio Curanja, upper Rio
Purus, altitude ca. 300 m.
Distribution (Fig. 81) Eastern Peru in the
Rio Ucayali Basin, southeast Balta and east into
Acre and Amazonas of Brazil.
Philander opossum (Linnaeus, 1758)
Philander, Opassum, sive Carigueja, Brasiliensis;
mas. [sic], Seba, 1734:56 (original spelling is as
indicated).
Philander, Americanus, seu Carigueja, cum catulis.
saccum ventris intrantihus; foemina Seba, 1 734:57
(original spelling is as indicated).
Carigueja, junior, sive Opassum, cum sacco aperto,
Seba, 1 734:57 description.
Sarigue, Buffon and Daubenton, 1763:279 descrip-
tion.
[Didelphis] opossum: Linnaeus, 1758:55.
Didelphis oposum: Brongniart, 1792:115 incorrect
subsequent spelling FRENCH GUIANA: (Cay-
enne).
Didelphis opossum: Cuvier, 1798:1 14 le sarigue. E.
Geoffroy St. Hilaire, 1803:141. Latreille, 1803:
304 sarigue ou opossum. Desmarest, 1820:256.
Temminck, 1825:41. Gray, 1843:101. Jentink,
1887b:301 SURINAME; FRENCH GUIANA:
(Cayenne). Jentink, 1888:220 SURINAME. J. A.
Allen, 1900b: 195 SURINAME: (type locality).
l'?]Didelphis opossum: Thomas, 1888b:329 part,
ECUADOR: Guayas (Guayaquil).
Molucca opossum: Shaw, 1800:476.
Didelphys opossum: Goldfuss, 1809:215. Waterhouse,
1841:90 part, GUYANA; SURINAME. Water-
house 1 846:485 GUYANA. Thomas, 1888b:329,
331 part, GUATEMALA: Esquintla (Vera Paz);
Aha Verapaz (Coban); part, GUYANA: Pomeroon-
Supenaam (Better Hope); SURINAME; ARGEN-
TINA: Chaco; BOLIVIA: Santa Cruz (Santa Cruz
BROWN: ATLAS OF NEW WORLD MARSUPIALS
129
de la Sierra). Lydekker, 1896:200 part, distribu-
tion. Goeldi and Hagmann, 1904:100 BRAZIL:
Para.
D[idelphys\ opossum: Olfers, 1818:204 part, syn-
onymy. Schinz, 1844:501. Tschudi, 1844:14, 144,
151 PERU. Wagner, 1855:226 listed.
S[arigua] opossum: Muirhead, 1819:429 part, new
combination.
Didelphys {Metachirus) opossum: Menegaux, 1902:
496 FRENCH GUIANA: (Ouanary River).
Sarigua opossum: Muirhead, 1830 (1819):429 clas-
sification.
Metachirus opossum: Burmeister, 1856:69. Thomas,
190 la: 153 GUYANA: Upper Takutu-Upper Es-
sequibo (Rupununi River; Kanuku Mts.). Thomas,
1 9 1 0b: 1 88 GUYANA: Pomeroon-Supenaam (Su-
penaam River). Thomas, 191 lb:143 SURINA-
ME: (type locality); Seba sole reference for opos-
sum Linnaeus. J. A. Allen, 1916a:201 COLOM-
BIA: Meta (Villavicencio). Bertoni, 1923:51
PARAGUAY: Central (Bahia de Asuncion). Lyne,
1959:84 VENEZUELA: Merida.
Metacherius opossum, Sanderson, 1949:787 incor-
rect subsequent spelling part, SURINAME.
Gamba opossum: Liais, 1872:329 classification.
[Metachirops] opossum: Matschie, 1916:268 SU-
RINAME: Paramaribo (type locality, Paramaribo).
Metachirops opossum: Bertoni, 1939:6 PARA-
GUAY. Crespo, 1950:6 ARGENTINA: Misiones
(Rio Urugua-i, 39 km from Puerto Bemberg).
[Holothylax opossum] opossum: Cabrera, 1919:48
classification.
Philander opossum: Gilmore, 1941:309 classifica-
tion. Moojen and Avila Pires, 1966:397 BRAZIL:
Cerrado formation (absent). Gardner and Patton,
1972:5 PERU: Ucayali (Balta; Yarinacocha);
Ayacucho (San Jose; Huanhuachayo [Hershkovitz
(1977) identified these specimens as Philander an-
dersoni andersoni]). Pine, 1973b:56 BRAZIL:
Para (Utinga, suburb of Belem). Reig et al., 1977:
197 part, VENEZUELA: Barinas (Guaquitas).
Reig et al., 1977:197, 212 part, PERU: Ucayali
(Yarinacocha; Balta); BRAZIL: Mato Grosso (To-
rixoreu, gallery forest; Baliza, gallery forest; Barra
do Garcas gallery forest; Pocone, gallery forest).
Tyndale-Biscoe, 1980:21 1 part, COLOMBIA:
Meta (Villavicencio); Valle del Cauca (Cali; Bue-
naventura). Tuttle et al., 1981:333 PANAMA:
Colon/Panama (Isla Barro Colorado [= Barro Col-
orado]). Mills et al., 1 98 la:331 BRAZIL: Para
(Belem). Miles et al., 1 98 lb:272 BRAZIL: Para
(Belem). Charles-Dominique et al.. 1981:342
FRENCH GUIANA: (Cayenne). Charles-Domi-
nique, 1983:395-422 FRENCH GUIANA: (Cay-
enne). Guerrero, 1985:41 VENEZUELA. Red-
ford and Da Fonseca, 1986:126, 132 BRAZIL:
cerrado region (gallery forest dependent). Atra-
mentowicz, 1986:123 FRENCH GUIANA: (Cay-
enne). Atramentowicz, 1988:47 FRENCH GUI-
ANA: (Cayenne). Gardner, 1988:698 VENE-
ZUELA: Amazonas (Base Camp I, Cerro de La
Neblina). Mares et al., 1989:19 BRAZIL: Mato
Grosso (Pocone). Pacheco et al., 1993:5 PERU:
Madre de Dios (Pakitza; Cocha Cashu; Aguas Cal-
ientes); Cusco (Hacienda Erika). Julien-Laferriere
and M. Atrementowicz, 1994 FRENCH GUI-
ANA (Cabassou; Piste de Saint Elie). Emmons et
al., 1994:146 PERU: Madre de Dios (Puesto San
Antonio). Emmons et al., 1994:144 PERU: Ma-
dre de Dios (Explorer's Inn Reserve). Emmons and
Remo, 1994:140 PERU: Madre de Dios (Ccolpa
de Guacamayas). Rageot and Albuja, 1994:178
Ecuador: Pastaza (Mera). Hershkovitz, 1997:90
BRAZIL: Sao Paulo (Iporanga, Parque Estudual
Petar); ARGENTINA: Chaco (Rio de Oro, boca);
Misiones (Rio Urugua-i; Fracran, San Pedro). Ma-
res and Braun, 2000:36 ARGENTINA: Chaco
(Parque Nacional Chaco; Puerto Bermejo, 10 km
S; Rio de Oro); Formosa (Laguna Blanca; Parque
Nacional Rio Pilcomayo); Misiones (Fracran,
Parque Nacional Iguazii; Rio Urugua-i). Voss et al.,
2001:63 FRENCH GUIANA (Paracou); SURI-
NAME: Coronie (Totness); Marowijne (Moengo;
Perica); Sipaliwini (Avanavero; Wilhemina Moun-
tains); Nickerie (Sipaliwini Airstrip; Kayserberg
Airstrip); Saramacca (Bigi Poika; La Poule); Com-
mewijne (Clevia; Plantation Clevia); Para (Lely-
dorpplan; Zanderij); Suriname (Powakka).
Metachirus opossum opossum: J. A. Allen, 1911:
246 VENEZUELA: Bolivar (Rio Yuruan).
Metachirops opossum opossum: Krumbiegel, 1941b:
200 part, BRAZIL: Para (Ipitinga, Rio Acara;
Peixe-boi). Husson, 1978:24 SURINAME: Sipal-
iwini (Avanavero Falls, Kabalebo River; upper
Nickerie River); Paramaribo (Paramaribo, various
localities in and near town; near Para River on road
to Domburg, 10 km SE of Paramaribo); Para (Le-
lydorp); Commewijne (near Commewijne River;
Clevia); Marowijne (Albina).^
Philander opossum opossum: Avila-Pires, 1958:4
BRAZIL: Para (Utinga [suburb of Belem], Belem
de Para). Husson, 1973:5 SURINAME. Carvalho,
1961 :5 BRAZIL: Roraima (formerly Rio Branco)
(Parana do Pocao, Caracarai; Pocao, Boa Vista).
Carvalho, 1962:285 BRAZIL: Amapci (Macapa,
Rio Amapari; Mazagao, Rio Maraca; Rio Branco;
Rio Maraca); Para (Sapucajuba; Utinga, suburb of
Belem). Pine, 1973b:56 BRAZIL: Para (Belem;
Utinga, suburb of Belem). Perez-Hernandez, 1989:
373 VENEZUELA: Bolivar (El Palmar; San
Martin de Turumban; Raudal Chalimana); Amazon-
as (Campamento, La Neblina; Rio Casiquiare).
Hershkovitz, 1997:90-92 VENEZUELA: Bolivar
(El Palmar; Maripa; Hato San Jose; San Martin de
Turumban; Rio Yuruan; Raudal Chalimana); GUY-
ANA: Pomeroon-Supenaam (Supenaam River;
Better Hope); Demerara-Mahaica (Georgetown;
Hyde Park); Essequibo Islands-West Demerara
(Buck Hall [= Buckhall]); Upper Takutu-Upper
Essequibo (Rupununi River; Kanuku Mts.). SU-
RINAME: Nickerie (Makerie; Kaiserberg Airstrip);
Sipaliwini (Avanavero Falls; Nickerie River); Bro-
kopondo (Loksie Hattie); Saramacca (La Poule);
Suriname (Culturutuin [= Cultuurtuin, now in Par-
amaribo}; Rijweg [now in Paramaribo]; Lelydorp-
plan [now in Para]; Paramaribo [now in Paramar-
ibo]; Clevia [now in Commewijne]; Para River
[now in Paramaribo]; Lelydorp [now in Wanica]);
Commewijne (Commewijne River); Marowijne
(Albina); FRENCH GUIANA: (Cayenne; Ouanary
River); BRAZIL: Amapd (Serra do Navio; Teresin-
ha; Macapa; Mazagao; Rio Branco, Rio Maraca);
130
FIELDIANA: ZOOLOGY
Roraima (Pocao, Boa Vista, km 192, Ferovia Ama-
pa; Pocao, Boa Vista; Pocao, Caracaraf: Caracaraf,
Rio Mucajai); Amazonas (Rio Xiriviny; Santo Isi-
dore; Ipixuna [= Lago do Ipixuna]; Serra de Par-
intins; Parintins; Santa Clara; Villa Bella Imperatriz
[= Parintins]; Lago do Baptista; Rio Madeira,
boca; Rosarhino; Auara Igarape; "Rio Negro," not
located; Santo Antonio de Uayara, Rio Eiru); Acre
(Seringal Oriente); Para (Fordlandia, Rio Tapajoz;
km 19, Itaituba-Jacareacanga [= Jacare-Acanga];
km 216, Estrada de Santarem-Cuiaba; Canudos;
Igarape Tapereba; Altamira, Rio Xingu; Gradaiis
[= Gorotire]; Cameta; Ilha do Taiuna; Mazagao;
Mocajuba; Baiao; Arumateua; Maraba; km 90, Ro-
dovia, Belem-Brasflia; Ipeau-Apez, suburb of Be-
lem; Sapucajuba, suburb of Belem; Utinga, suburb
of Belem; Marco, suburb of Belem; Murutucu [=
Instituto Agronomico do Norte]; Ananindeua; Ipi-
tinga; Flor do Prado, near Quatipuru; Santa Maria,
Braganca; Peixe-boi; Igarape Acu, Colonia do Pra-
ta; Capim; Sao Miguel do Guama [= Guama]; Ir-
iteria [= Guama]). Patton and da Silva, 1997:102
GUYANA: Potaro Siparuni (Surama, 30 km SE);
Barima-Waini (Santa Cruz; Waikerebi); BRAZIL:
Para (Rio Xingu, E bank, 52 km SSW of Altami-
ra). Patton et al., 2000:81-82 GUYANA: Potaro
Sipalwini (Surama, 30 km NE); Barina-Waini
(Waikerebi); BRAZIL: Amazonas (Ilha das Oncas,
left bank Rio Negro; Lago Meduinfm, left bank Rio
Negro; Sacado, right bank Rfo Jurua; Igarape Nova
Empresa, left bank Rio Jurua); Para (Floresta Na-
cional Tapirape-Aquiri, Municipio da Maraba);
Acre (Igarape Porongaba. right bank Rio Jurua;
Flora [= Fazenda Santa Fe], left bank Rio Jurua;
Ocidente, opposite, right bank Rio Jurua; Nova
Vida, right bank Rio Jurua; Sobral, left bank Rio
Jurua); FRENCH GUIANA (La Trinite Moun-
tains).
Philander o\possum] Opossum: Carvalho and Toc-
cheton, 1969:217 BRAZIL: Para (Utinga, Be-
lem; Sapucajuba, Belem; km 94 Belem-Brasflia).
Didelphis Marsupialis: Shaw (not Linnaeus, 1758),
1 800:476 Molucca opossum.
P[hilander] virginianus Tiedeman, 1808:427 re-
placement name for D. opossum Linnaeus.
Didelphis austro-americana Thomas, 1923:604
name ex Oken, 1816:1 148 a non-Linnaean work:
synonym of D. opossum Linnaeus.
Didelphis quica: Schomburgk, 1840:344 GUYA-
NA. Jentink, 1888:220 SURINAME.
Didelphys quica: Waterhouse, 1841:90 part, GUY-
ANA; SURINAME. Waterhouse, 1 846:480 part.
GUYANA; SURINAME. Alston, 1880:198
MEXICO; GUATEMALA: Suchitepequez (San
Miguel Duenas [= Duenas]); Alta Verapaz (Cob-
an); COSTA RICA: Cartago (Tucurriqui). Pelzeln,
1883:110 BRAZIL: Mato Grosso (Mato Grosso).
D[idelphys] quica: Cabanis and Schomburgk (not
Temminck), 1848:777 GUYANA.
Metachirus opossum quica: Miller, 1916:589 J. A.
Allen, 19 16b:562 BRAZIL: Mato Grosso do Sul
(Urucum). Carvalho, 1957:3 BRAZIL: Acre (Ser-
ingal Oriente, Rio Jurua).
Metachiropos opossum quica: Vieira, 1945:421
BRAZIL: Mato Grosso (Monte Alegre, Palmeiras).
Vieira, 1949:345 BRAZIL: Mato Grosso. Car-
valho, 1958:123 BRAZIL: Para (Gradaiis [= Go-
rotire], Rio Fresca).
Philander opossum quica: Hershkovitz. 1997:86-
95 COLOMBIA: Boyacd (Rfo Cobaria); Meta
(Caney; Restrepo; Villavicencio; Finca El Capri-
cho); PERU: Loreto (Itaya; Iquitos; Apayacu; Sar-
ayacu, Rio Ucayali; San Jeronimo); Ucayali (Pu-
callpa; Lagarto Cocha; Rfo Urubamba; Santa
Rosa); San Martin (Yurac Yacu; Moyobamba; Rio-
ja); Hudnuco (Tingo Maria); Cusco (Hacienda Cad-
ena; San Jeronimo; Quincemil); Mad re de Dios
(Rfo Colorado, boca; Puerto Maldonado); BOLIV-
IA: La Paz (Chulumani); Beni (Guayamarfn; Mer-
redor; Rfo Mamore; Itonama; Puerto Siles, Rfo Ma-
more; Santa Rosa; Barranquita; Arruda; Centinela;
Camino Vilches; Santo Dios; San Joaqufn; Exal-
tation. Rfo Mamore; Palacios; Puerto Caballo; San
Ramon, Rfo Machupo; Monte Chocolatal; Magda-
lena; San Pablo; El Carmen, Rfo Blanco; Rfo Ibare,
boca; Vaca Diez; San Ignacio de Moxos; Camiaco);
Cochabamba (Rfo Ichilo); Cochabamba-Santa
Cruz (Rfo Mamore, 2 km S mouth Rfo Chapare);
Santa Cruz (Ascencion de Guarayos; Mercedes,
Rfo Guapore; Buenavista [= Buena Vista]; Warnes;
Santa Rosita; Tocomechi; Hamacas; El Palmar;
Ayacucho, Ibanez; Santa Cruz de la Sierra [= Santa
Cruz], 18 km SW; San Ramon); PARAGUAY: San
Pedro (Tacuati); Central (Asuncion; Lapango);
Paraguari (Sapucay [= Sapucaf]); BRAZIL: Acre
(Seringal Oriente); Rondonia (Porto Velho); Mato
Grosso (Mato Grosso; Barra do Garcas; Pocon6;
Balisa; Caceres; Monte Alegre, Palmeiras); Mato
Grosso do Sul (Santa Teresa; Urucum; Miranda);
Rio Grande do Sul (Passo Fundo; Taquara).
Metachirus fuscogriseus: J. A. Allen, 1900b: 194 (ho-
lotype, probably female, skin and skull, AMNH
9920/8252; presented by Albert Smith. The skin is
labeled female, skull tag and box are labeled male).
Allen, 1 90 lb:2 13 NICARAGUA: Rio San Juan
(Greytown [= San Juan del Norte]); Zelaya (Blue-
fields).
Metachirops fuscogriseus: Matschie, 1916:268 clas-
sification.
Metachirus opossum fuscogriseus: J. A. Allen, 1911:
247 NICARAGUA: Rio San Juan (Greytown [ =
San Juan del Norte]).
[Holothylax fuscogriseus] fuscogriseus: Cabrera,
1 9 1 9:47 classification.
Metachirops opossum fuscogriseus: Enders, 1935:
410 PANAMA: Colon/Panama (Barro Colora-
do).
Philander opossum fuscogriseus: Alston, 1880: 198
GUATEMALA: Suchitepequez (San Miguael
Duenas). Allen, 1 90 lb:2 13 NICARAGUA: Ze-
laya (Bluefields). Allen, 1910:13 NICARAGUA:
(Rfo Coco, on border of Honduras and Nicaragua);
Chinandega (Volcan de Chinandega); Madriz (San
Juan); Zelaya (Rfo Escondido); Jinotega. Goodwin,
1942:113 NICARAGUA: (type locality, Grey-
town; "found [in New York, 11 January 1895] in
a bunch of bananas in unloading a fruit steamer
from a Central American port, most likely Colon
. . ." [J. A. Allen, 1900b: 194]; "subsequently fixed
as Greytown [= San Juan del Norte]," Nicaragua
[Hershkovitz, 1997:56]); HONDURAS. Hall and
Kelson, 1959:11 distribution in Middle America.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
131
Yates and Froehlich, 1984:512 HONDURAS:
Gracias a Dios (Rfo Platano Biosphere Reserve).
Hershkovitz, 1997:52, 85-87 COLOMBIA: An-
tioquia (Alto Bonito; Bellavista); Caldas (Rio Hon-
do, Samana); Chocd (Bagado; Condoto; Rio Sando,
Rfo Baudo; Rfo Docampado; Novita); Valle del
Cauca (Rfo Frfo; Buenaventura; Buenaventura, 28
km NE; Rfo Raposo; Cali; Rfo Cauca [see Cali));
Cauca (Rfo Saija, boca; Cocal); ECUADOR: Man-
abi(Rio Pescado; Rfo de Oro); Guayas (Guayaquil;
Rfos Chimbo-Coco; Bucay, Rfo Chimbo; Ventura,
Rfo Chanchan; Puente de Chimbo |now in Chim-
borazo]); El Oro (El Chiral; Pifias; Salvias); Loja
(Punta Santa Ana; Cebollal |= Seboyal]). Patton
and da Silva, 1997:102 PANAMA: Bocas del
Toro (Old Point, Isla Bastimentos; Isla Popa, 1 km
SE of Deer Island Channel). Patton et al., 2000:
82 PANAMA: Bocas del Toro (Old Point, Isla
Bastimentos; Isla Popa, 1 km SE of Deer Island
Channel).
Metachirus fuscogriseus pallidas: J. A. Allen, 1901b:
213 MEXICO: Veracruz-Llave (type locality,
Orizaba; holotype, male, skin and skull, USNM
51858, collected 24 January 1894 [not 24 June
1894 as in original description] by E. W. Nelson
and E. A. Goldman); Veracruz-Llave (Motzoron-
go); Oaxaca (Tuxtepec); Puebla (Metlaltoyuca);
Tabasco (Montecristo [= Monte Cristo]; Frontera
[= Alvaro Obregon]; Teapa); Chiapas (Huehuetan;
La Chicharras [= Las Chicharras]).
[Holothylax fuscogriseus] pallidus: Cabrera, 1919:
47 classification.
Metachirops opossum pallidus: Goodwin, 1934:4
GUATEMALA; distribution. Davis, 1994:375
MEXICO: Veracruz-Llave (Boca del Rfo).
Philander opossum pallidus: Allen, 1901b:215-
216 MEXICO: Veracruz-Llave (Orizaba). Good-
win, 1934:5 GUATEMALA: Aha Verapaz (Finca
Sepacuite); San Martin (Finca Carolina). Suchite-
pequez (Finca Cipres). Villa, 1949:497 MEXICO:
Chiapas (Mapastepec). Dalquest, 1953:19 MEX-
ICO: San Luis Potosi (El Salto; Xilitla). Baker,
1951:207 MEXICO: Tamaulipas (Ciudad Victo-
ria, 70 km S; El Carrizo, 2 km W, 757 m). Hall
and Kelson, 1959:11 distribution. Hall and Dal-
quest, 1963:195 MEXICO: Veracruz-Llave (dis-
tribution, near sea level to 1697 m above). Alvarez,
1963:394 MEXICO: Tamaulipas (E slope Sierra
Madre Oriental, not precisely located; La Purfsma,
7 km SW; Rancho Pano Ayucutle, 6 mi N of Go-
mez Farias, 25 mi N of El Mante; El Encino; El
Carrizo).
Metachirus grisescens J. A. Allen, 1 901 b:21 7 CO-
LOMBIA: Valle del Cauca (type locality, "Rio
Cauca"; holotype, female, skin and skull, AMNH
15072, collected June 1888 by J. H. Batty. Good-
win, 1953:227.
[Metachirops] grisescens: Matschie, 1916:268 clas-
sification.
[Holothylax grisescens[ grisescens: Cabrera, 1919:
47 classification.
Metachirus opossum grisescens: J. A. Allen, 1916a:
201 COLOMBIA: Valle del Cauca (Rfo Frfo).
Philander opossum grisescens: Goodwin, 1953:227
classification. Perez- Hernandez (not J. A. Allen),
1989:373 VENEZUELA: Barinas (La Lengueta,
SE Barinas; Guayquitas; Cafio Anaru, "Las Boni-
tas," 22 km NE Guasdalito; Las Bonitas; Ticopo-
ro); Apure (Nulita); Tdchira (La Ponchera); Trujillo
(Motatan; Rio Motatan; El Dividivf); Zulia (Encon-
trados; Alguacil; Churulf; Puerto Catatumbo); Me-
rida (Zea; Bejuquero; Carlo Tigre, Zea; Mina Car-
bon, Zea).
Metachirus opossum melantho Thomas, 1923c:602
COLOMBIA: Chocd (type locality, Condoto, 300
ft [= 90 m]; holotype, young adult male, skin and
skull, BMNH 1914.5.28.30, collected 1 February
1914 by G. F Spurrell).
Metachirus opossum melanurus Thomas, 1 899b:
285 ECUADOR: Imbabura (type locality, Par-
amba [= Hacienda Paramba], Rfo Mira, western
Ecuador, 1100 m; holotype, old male, skin and
skull BMNH 1897.1 1.7.61; collected 1 1 April 1897
by W. F. H. Rosenberg). Lonnberg, 1913:36 EC-
UADOR: Pichincha (Gualea). J. A. Allen, 1912:
73 COLOMBIA: Cauca (Cocal); J. A. Allen,
1916:116, 201 COLOMBIA: Cauca (Cocal);
Chocd (Bagado); Antioquia (Alto Bonito). Cabrera,
1958:35 part.
[Metachirops] melanurus: Matschie, 1916:268 clas-
sification.
[Holothylax opossum] melanurus: Cabrera, 1919:
48 classification.
Metachirops opossum melanurus: Lonnberg, 1921:
68 ECUADOR: Pichincha (Gualea).
Philander opossum melanurus: Cabrera, 1958:35.
Hershkovitz, 1997:51, 86 ECUADOR: Limon,
not located; Esmeraldas (Cachavi, Rfo Cachavf;
Carondelet; San Javier; Esmeraldas; Quininde [=
Rfo Quininde]); Imbabura (Paramba); Pichincha
(Rfo Caonf; Gualea; Mindo).
P[hilander] o[possum] melanurus: Baker, 1974:135
ECUADOR: Esmeraldas (Quininde).
Metachirus canus Osgood, 191 3b:96 PERU: San
Martin (type locality, Moyobamba; holotype, male,
skin and skull, FMNH 19347; collected 4 August
1912 by W. H. Osgood and M. P. Anderson). Os-
good, 1914b: 148 PERU: San Martin (Moyob-
amba).
[Metachirops] canus: Matschie, 1916:268 classifi-
cation.
[Holothylax grisescens] canus: Cabrera, 1919:47
classification.
Metachirus opossum canus: Thomas, 1927a:372
PERU: San Martin (Rioja; Moyobamba; Yurac
Yacu). Thomas, 1927b:606 PERU: Hudnuco
(Tingo Maria, 2000 ft [= 625 m]). Thomas, 1928a:
264 PERU: Ucayali (Cumerfa [= Cumarfa], 1000
ft [= 300 m]; Chicosa, 1000 ft [= 300 m]; San
Jeronimo [now in Loreto], 500 ft [= 150 m]).
M[etachirus] o[possum] canus: Thomas, 1928b:294.
Philander opossum canus: Sanborn, 1949:277
PERU: Ucayali (Yarinacocha). Sanborn, 1951b:2
PERU: Cusco (Hacienda Cadena). Patton and da
Silva, 1997:102 BRAZIL: Acre (Fazenda Santa
Fe [= Flora], left bank Rfo Jurua; Seringal Ociden-
te, right bank Rfo Jurua); Amazonas (Nova Empre-
sa, left bank Rfo Jurua). Anderson, 1997:162 BO-
LIVIA: Beni (Nicolas Suarez Isalnd, Rfo Mamore,
near Guayaramerin; Guayaramerin; Guayaramerin,
5 km S; Riberalta; Buena Hora, 6 km S; Rfo Ma-
more; Puerto Siles; Camino Vilches, San Joaqufn;
132
FIELDIANA: ZOOLOGY
San Pablo; Exaltacion, 8 km N; Boroica; Barran-
quita; Piedras Blancas; Exaltacion; San Ramon;
Magdalena; Palacios, 4 km SE; Puerto Caballo; El
Carmen; Espiritu; Yacuma, 45 km N; Santa Rosa;
Rurrenabaque; Rio Ibare, mouth; Rio Ibare, 24 km
from mouth; Rio Ibary, 27 km from mouth; Cas-
arabe; San Ignacio de Moxos; Rfo Tijamuchi; Cam-
iaco; Alto Beni, not located; Arruda; Centenela;
Mamore; Santo Dios); Cochabamba (Rio Ichilo);
La Paz (Rio Beni); Pando (Agua Dulce; Rio Na-
reuda; Bella Vista; Independencia; Isla Gargantua);
Santa Cruz (Aserradero Moira, km 7 E; Pajaral;
Ascencion de Guarayos; Ascencion, 6 km W; Los
Palmares; Rfo Mamore; Totaitu; Yapacani; San Ra-
mon, 10 km N; Estancia Cachuela Esperanza; Pal-
marito; Santa Rosa, 7 km N; Montero, SE of; San
Miguel Rincon; Tocomechi; Buenavista; Warnes;
Santa Rosita; La Belgica, 2 km S; Hamecas; Co-
toca, 2 km SE; El Palmar road; Santa Cruz de la
Sierra; El Palmar; Ayacucho; Santa Cruz, 15 km S;
Santa Cruz, 15 km SW; Santiago, Chiquitos; Tita).
Patton et al., 2000:81-82 BRAZIL: Acre (Oci-
dente, right bank Rio Jurua; Ocidente, opposite, left
bank Rfo Jurua; Igarape Porongaba, right bank Rfo
Jurua; Flora [= Fazenda Santa Fe], left bank Rfo
Jurua; Nova Vida, rigtht bank Rfo Jurua; Sobral,
left bank Rfo Jurua); Amazonas (Sacado, right bank
Rfo Jurua; Igarape Nova Empresa, left bank Rfo
Jurua).
Metachirus opossum azaricus Thomas, 1923c:604
PARAGUAY: Paraguari (type locality, Sapucay
[= Sapucaf]; holotype, female, skin and skull,
BMNH 1903.2.3.36; collected 8 August 1892 by
W. Foster).
Met[achirops] opossum azaricus: Krumbiegel, 1941b:
203, 206 PARAGUAY: Central (Lapango); BO-
LIVIA: Santa Cruz (San Ramon).
Philander opossum azaricus: Cabrera, 1958:34. Kan-
tis, 1963:54 ARGENTINA: Chaco (Rfo de Oro,
boca); Misiones (San Pedro). Crespo, 1974:3 AR-
GENTINA: Chaco (Rfo de Oro, boca); Misiones
(Fracran, San Pedro).
Metachirus opossum crucialis Thomas, 1923e:604
BOLIVIA: Santa Cruz (type locality, Santa Cruz
de la Sierra; holotype, female, skin [originally
mounted] and skull, BMNH 1847.11.22.15, col-
lected by Thomas Bridges).
Philander opossum subspecies nov.: Perez-Hernan-
dez, 1990:373, 377 VENEZUELA: Monagas
(Cerro Papelon); Delta Amacuro Territorio Federal
(Tobesobe, Guayo; Los Giiires; Guiniquina).
Hershkovitz, 1997:52, 90 VENEZUELA: Trujillo
(Agua Viva).
Philander opossum subspecies: Hershkovitz, 1997:
90 VENEZUELA: Zulia (Puerto Catatumbo; Rfo
de Oro, boca; Alguacil; Churulf; Encontrados);
Delta Amacuro Territorio Federal (Los Giiires; To-
besobe; Guiniquina); Merida (Merida); Barinas
(Ticoporo; Guaquitas; Las Bonitas, Cano Amaru;
La Lengueta); Apure (Nulita; La Blanquita); Tdch-
ira (La Ponchera); Trujillo (Motatan; Rfo Motatan;
Agua Viva; El Dividivf).
Philander\T\ opossum subspecies: Perez-Hernandez,
1989:373 VENEZUELA: Monagas (Cerro Pape-
lon). Hershkovitz, 1997:90 VENEZUELA: Mon-
agas (Cerro Papelon).
Type Name based on Seba's (1734) "Philan-
der Opassum, sive Carigueja, Brasiliensis mas,"
and "Philander, Americanus, sive Carigueja, cum
catulus, sacrum bentris intrantibus foemina^ '; the
female, RMNH 25421, in alcohol, designated lec-
totype by Hershkovitz (1976), is preserved in the
Rijksmuseum van Naturlijke Historie, Leiden.
Type Locality "America," restricted to Su-
riname by J. A. Allen, 1900b: 195, and further re-
stricted to Paramaribo, Suriname, by Matschie,
1916:261.
Distribution (Figs. 82 and 83) From Tamau-
lipas, Mexico, through Central and South America
to Paraguay and northeastern Argentina.
Additional Records MEXICO: Tamaulipas;
El Mante (Schaldach, 1954, KU); Rfo Sabinas, 10
km N of El Encino (Schaldach, 1954, KU);
Ocampo (Clifton, 1965, KU); La Purfsma (Dal-
quest, 1946, KU); San Luis Postosi; El Salto Falls
(Stock, 1963, USNM); Valles (Shaw, 1946,
LSUMZ); Huichihuyan (Dalquest, 1950, LSUMZ);
Veracruz-Llave; Tuxpan (Lee, 1960, KU); Tihu-
atlan (Bodley, 1961, KU; Clifton, 1961, KU); Pa-
pantla (Nelson, 1898, USNM); San Marcos (Dal-
quest, 1947, KU); Teocelo (Dalquest, 1949, KU);
Veracruz-Llave, Perez (Friesser, 1903, USNM);
Fortin (Dalquest, 1946, KU); Rfo Atoyac (Dal-
quest, 1946, KU); Potrero (Dalquest, 1946, KU;
Sumichrast, USNM); Potrero Viejo, 5 km W of
Potrero (Dalquest, 1946, KU); Cordoba, 3 mi SE
of Yongas (Cutter, 1956, KU); Rfo Seco, near
Cordoba (Sumichrast, 1866, USNM); Rfo Blanco,
20 km WNW of Piedras Negras (Dalquest, 1946,
KU); San Andres Tuxtla (Dalquest, 1948, KU;
Parker-Robinson, 1959, UMMZ); Catemaco (Nel-
son, 1891, USNM); La Jimba [= Jimba] (Dal-
quest, 1947, KU); Tenochtitlan (Lackey, 1968,
UMMZ); Achotal (Heller, 1904, FMNH); Jesus
Carranza (Dalquest, 1948, 1949, KU); Chichicaf-
te, not located (Nelson, 1894, USNM); Oaxaca;
Metates, SSW of Sierra Juarez (Campbell, 1981,
KU); Matfas Romero, S of (Straney, 1977, MVZ);
Tapanatepec (Musser, 1962, UMMZ); Tabasco;
Miramar (Woods, 1960, LSUMZ); La Venta (We-
ber, 1943, USNM); Cardenas (Dickerman, 1955,
KU); Villa Hermosa (Duellman, 1958, UMMZ);
Balancan (Lay, 1960, LSUMZ); Macuspana, 19
km N, 5 mi SW (Dickerman, 1955, KU); Chia-
pas; Palenque (Kuns, 1949, FMNH); Tuxtla Gu-
tierrez (Dickerman, 1955, KU); Finca Ocuilapa,
10 km S of Tonala (Huckaby, 1969, UMMZ); El
Talisman [= Talisman] (Smith, 1965, KU); Cam-
peche; Escarcega, 65 km S, 128 km E Clanton,
1963, KU); La Tuxpena [= Chompoton] (Shu-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
133
110
100
Fig. 82. Distribution of Philander opossum in Central America: from Tamaulipas. Mexico, south through Panama.
See Figure 83 for distribution in South America, o = localities from unpublished records; = localities given in
published records.
feldt, 1912, USNM); Yucatan; Tekom (Sanderson.
1940, FMNH); Hacienda de Carmen, not located
(Rice, USNM); GUATEMALA: Peten; Uaxactun
(Murie, 1931, UMMZ); Parque Nacional Tikal
(McLean and Navin, 1983, 1987, USNM); La
Libertad [= Libertad] (Malleis, 1923, USNM);
Alta Verapaz; Chinaja (Jones, Jr., 1960, KU); Iza-
bal; El Estor-Los Dantas (Koford, 1978, MVZ);
Bobos (Schmidt, 1933, FMNH); Los Amates
(Heller and Barber, 1905, FMNH); El Progresso;
San Agustin, Aldea Puerto Golpe (McLean, 1983,
USNM); Esquintla; Finca El Salto, 1 mi E of Es-
quintla (Lawler, 1967, UMMZ); Guanagazapa
(McLean, 1984, USNM); Masagua, Finca Santil-
lana del Esquintla (McLean, 1984, USNM); Con-
cepcion del Mar (Blake, 1934. FMNH); Santa
Rosa; Finca El Progresso (De La Torre, 1951,
UMMZ); Jutiapa; Laguna de (= Lago) Atesca-
tempa (Handley, Jr., 1947, USNM); Baja Verapaz
[probably Alta Verapaz]; Astillero (Bee, 1955,
KU); BELIZE: Gallon Jug, Orange Walk District
(Lancaster, 1956, LSUMZ); Belize (Murie. 1931,
UMMZ); Cayo District; Cayo (Murie, 1931,
UMMZ); Cayo; Mt. Pine Ridge, S of Cayo (Mu-
rie, 1931, UMMZ); Central Farm (Disney, 1964,
USNM); Stann Creek District; Pomona, 4 mi E
(Dennett, Jr., 1956, LSUMZ); HONDURAS: Cor-
tes; San Pedro, near Lago Ticamaya (Schmitt and
Walters. 1923, FMNH); Gracias a Dios; Patuca
(Perry, USNM); Valle; Coray (McLean, 1984,
USNM); Nacaome, Rio de Coray (McLean, 1983,
USNM); EL SALVADOR: Ahuachapdn; Barra de
Santiago Ahuachapan (Stirton, 1927, MVZ); Son-
sonate; Hacienda Chilata (Hildebrand, 1942,
MVZ); Usulutdn; Puerto del Triunfo (Stirton,
1925, 1926, MVZ); San Miguel; Rio San Miguel
(Stirton, 1926, MVZ); Lago Olomega (Marshall
Jr.. 1942. MVZ; Stirton, G. D., 1925, MVZ; Stir-
ton, R. A., 1942, MVZ); Morazdn; Divisadero, 2
mi N (Stirton, 1926, MVZ); NICARAGUA: Nue-
va Segovia; Jalapa (Schlitter, 1967, KU; Geno-
ways, 1967, KU); Jinotega; Hacienda La Trampa,
16 km NE Jinotega (Clark, 1964, KU); Chinan-
dega; Cosigiiina (Smith, 1968, KU); Hacienda
San Isidro, El Realejo. 10 km S of Chinandega
(Clark. 1963. USNM; 1963, KU); Managua; Ma-
nagua (Alcorn, 1956, KU); Finca America, Ma-
nagua (Boshell, 1953, USNM); Sabana Grande
(Choate, 1968, KU; Genoways, 1968, KU; Jones.
Jr.. 1964, 1968. KU; Kirsch. 1968, KU; Lawler,
134
FIELDIANA: ZOOLOGY
>y
90 80
Fig 83 Distribution of Philander opossum in South America: from northern South America south through north-
eastern Argentina, Paraguay, and southern Brazil. See Figure 82 for distribution in Central America, o = localities
from unpublished records; = localities given in published records.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
135
1964, KU; Phillips, 1968, KU); Hacienda Aza-
cualpa, 5 km NW of Villa El Carmen (Clark,
1964, KU; Clark, 1964, USNM); Matagalpa;
Santa Maria de Ostuma (Genoways, 1966, KU;
Schlitter, 1966, KU; Turner, 1966, KU); Hacienda
Tepeyac, near Matagalpa (Clark, 1964, USNM;
Clark, 1964, KU); Esquipulas (Choate, 1968,
KU); Hacienda Tepello, not located (Clark, 1963,
USNM); Boaco; Teustepe (Choate, 1968, KU;
Kirsch, 1968, KU; Phillips, 1968, KU); Cholute-
ca; Diriamba (Choate, 1968, KU; Genoways,
1968, KU); Granada; Mecatepe, 2 km N, 1 1 km
E of Nandaime (Clark, 1964, KU; Clark, 1964,
USNM); La Calera, W of Nandaime (Clark, 1964,
KU; Clark, 1964, USNM); Rivas; Finca Amayo,
SE of Rivas (Clanton, 1963, KU; Jones, Jr., 1964,
KU; Lawler, 1966, KU); Sapoa (Turner, 1966,
KU); Ometepe (Nutting, 1983, USNM); Zelaya;
Bonanza (Clark, 1964, KU; Clark, 1963, USNM);
Rfo Kurinwas [= Rib Curinhuas] (Correra, 1967,
USNM); Rfo Siquia (Allen, 1935, UMMZ); El
Recreo (Choate, 1968, KU; Clanton, 1963,
USNM; Clanton, 1963, KU; Smith, 1968, KU);
Cara de Mond, (Phillips, 1968, KU); Rio San
Juan; La Esperanza (Clark, 1965, KU; Clark,
1965, USNM); COSTA RICA: El Sance Peralta,
not located (Underwood, 1931, USNM); Alajuela;
San Carlos (Smith, 1934, FMNH); Villa Quesada
(Smith, 1934, FMNH); Puntarenas; Monteverde,
Rfo Guacimal (Brown, 1966, UMMZ); Guana-
caste; Hacienda Santa Maria (Smith, 1932,
UMMZ); Finca Jimenez (Hooper, 1966, UMMZ);
San Jose; Monterrey, 22 km S of San Jose (Wille,
1947, KU); Santa Ana (McPherson, 1970,
LSUMZ); San Ignacio (Underwood, 1931,
USNM); Santa Juana, 10 km NE of Quepos (Wil-
le, 1947, KU); Savanillas de Pirns, not located
(Underwood, 1931, FMNH); Heredia; Puerto
Viejo, Rio Sarapiqui (Musser, 1962, UMMZ);
Cartago; Pacuare, Rfo Pacuare (Gardner, 1967,
LSUMZ; Zeladon, 1876, 1884, USNM); Turrialba
(Hall, 1947, KU; Musser, 1962, UMMZ; Torres,
1947, KU); PANAMA: Bocas del Toro; Chan-
guinola (Handley, Jr., 1960, USNM; Hinds, 1960,
USNM); Boca del Drago (Handley, Jr., 1960,
USNM); Isla Colon (Greenwell, 1988, USNM;
Handley, Jr., 1960, USNM); Cayo Nancy (Green-
well, 1988, USNM); Almirante (Handley, Jr.,
1960, USNM); Isla Bastimentos (Handley, Jr.,
1962, USNM; Hinojosa, 1987, USNM; Jacobs,
1987, USNM); Isla San Cristobal (Greenwell,
1989, USNM); Tierra Oscura (Greenwell, 1989,
USNM); Cayo de Agua (Greenwell, 1990,
USNM; Handley, 1962, USNM); Penfnsula Val-
iente, Punta Alegre (Greenwell, 1990, USNM);
Penfnsula Valiente, Quebrada Hido (Greenwell,
1990, USNM); Nun (Greenwell, 1993, USNM;
Handley, Jr., 1993, USNM); Rfo Changena Camp
(Handley, Jr., 1962, USNM; Keenan, 1961,
USNM); Tabori, not located (Greenwell, 1983,
USNM); Chiriqui; El Volcan, 14.5 km NW of
Finca Santa Clara (Enders, 1971, 1972, 1973,
USNM); Progreso (Greenwell, 1966, USNM);
Bugaba (Julatson, 1911, USNM); Puerto Armuel-
les (Greenwell, 1966, USNM); Veraguas; Isla
Gobernadora (Tyson, 1962, USNM); Isla Cebaco
(Tyson, 1965, USNM); Code; El Valle (Enders,
1957, USNM; Keenan, 1954, USNM; Wislocki,
1955, USNM); Colon; Salud (1957, USNM);
Coco Solo (1955, USNM); Camp Pina, Fort Sher-
man (1955, 1956, USNM; Keenan, 1956, USNM);
Cristobal (Fleming, 1966, 1967, USNM); Fort Da-
vis (Beauregard, 1953, USNM; Enders, 1930,
UMMZ; Keenan, 1952, 1955, USNM; Setzer,
1953, USNM); Gatun, Rfo Indio (Goldman, 191 1,
USNM); Isla Puma (1956, USNM); Lago Gatun
(Gardner, 1973, LSUMZ); Bohio Penfnsula (Wil-
son, USNM); Buena Vista Penfnsula (Pine, 1972,
USNM); Gamboa, 5 mi NE (Koford, 1952,
MVZ); Gamboa (Setzer, 1953, USNM); Panama;
Bruja Point (Enders, 1930, USNM); Pina (Keen-
an, 1960, USNM); Juan Mina [Juan Mina Sta-
tion]; Tabernilla (Goldman, 1911, USNM); Rfo
Mandinga (Keenan, 1957, USNM); Madden Road
(Keenan, 1955, USNM); Empire (Enders, 1957
USNM; Goldman, 1912, USNM); Cocoli (1956
USNM); Curundu (Keenan, USNM); Albrook
Field (Field, 1954, USNM; Setzer, 1953, USNM)
Albrook (Keenan, 1954, USNM); Rodman Naval
Ammunition Depot (Fleming, 1967, UMMZ)
Balboa (Fleming, 1966, 1967, USNM); Isla Bue
naventura, not located (Goldman, 1911, USNM)
La Zumbadora [= Cerro Azul] (1956, 1957, 1958
USNM; Handley, Jr., 1957, USNM; Keenan
1956, USNM); Chepo (Field, 1954, USNM); Cer-
ro Campana (Enders, 1955, USNM); Darien
Jaque (Greenwell, 1966, USNM); Los Santos
Cerro Hoya (Handley, Jr., 1962, USNM); CO-
LOMBIA: Valle del Cauca; Alto Rfo Anchicaya
(Simons, 1981, USNM); ECUADOR: Imbabura
Intag, Penaherra (Hershkovitz, 1934, UMMZ)
Carchi; La Concepcion (Hershkovitz, 1934
UMMZ); Los Rios; Vinces (McLean, 1974
USNM); Hacienda Santa Teresi (McLean, 1974
USNM); Puebloviejo (McLean, 1974, USNM)
Lima Pareja (McLean, 1975, USNM); El Papayo
(McLean, 1975, USNM); Manabi; La Papaya, not
located (Mena, 1942, FMNH); El Oro; Moromoro
136
FIELDIANA: ZOOLOGY
(Gardner, 1976, USNM); PERU: Ucayali; Sha-
hufa, Rio Ucayali, opposite Cumaria (Hendee,
1927, AMNH); Amazonas; Chiriaco, 43 km NE
(Graham, 1978, LSUMZ); Madre de Dios; Reser-
va Cuzco Amazonico, 14 km from Puerto Mal-
donado (Hooper, Jr., 1990, KU; Woodman, 1989,
KU); Rio La Torre, boca, S bank Rio Tambopata
(Barkley, 1979, LSUMZ); Lago Sandoval, Rio
Madre de Dios (Patton, 1979, MVZ); VENEZUE-
LA: Zulia; Cerro Azul, near (Keenan, 1956,
USNM; Handley, 1957, 1958, USNM); El Rosa-
rio (Smithsonian- Venezuelan Expedition, 1968,
USNM); Aragua; Rancho Grande (Rood, 1960,
UMMZ); GUYANA: Pomeroon-Supenaam; Pom-
eroon (1933, USNM); FRENCH GUIANA: Cay-
enne; Sinnamarey (Hershkovitz notes); BRAZIL:
Amapd; Rio Amapari, Serra do Navio (Dente,
1964, LSUMZ); Ferrovia Amapa, 192 km Serra
do Navio, not located (Dente, 1964, LSUMZ);
Para; Oriximina (1976, USNM); Mato Grosso;
Limao, Rio Jurua (Hibbs, 1965, USNM); Cuiaba
(Hibbs, 1965, USNM); Mato Grosso do Sul; Cor-
umba, Rio Paraguay (Kuns, 1965, USNM); Dour-
ados (Kuns, 1965, USNM); Districto Federal;
Guara, gallery, not located (Hershkovitz, 1997,
notes). BOLIVIA: Santa Cruz; Cordillera Basilio
(Becerra, 1966, USNM); PARAGUAY: Concep-
cion; Rio Aquidaban, Paso Horqueta (Creighton,
1979, UMMZ); Aca Poi (Wharton, 1950, USNM);
Cordillera; Tobati, 12 km N (Myers, 1973,
MVZ); Central; Luque, 17 km E (Koford, 1972,
MVZ; Myers, 1976, UMMZ); ARGENTINA:
Formosa; Estancia Linda Vista (see also Riacho
Pilaga), NNE Formosa (Creighton, 1980,
UMMZ).
Genus Thylamys Gray, 1843
Sarigua, Muirhead, 1819:429 part, S. pusilla listed
among included species.
Thylamys Gray, 1843:xxiii, 101 type species, Thy-
lamys elegans [= Didelphis elegans Waterhouse]
by monotypy. Thomas, 1888b:340 type species,
Didelphis elegans Waterhouse. Matschie, 1916:
262, 266, 271 taxonomy. Miranda Ribeiro, 1936:
386 taxonomy. Reig, 1958:274 (footnote) rec-
ognized as a valid subgenus equivalent to the ele-
gans and microtarsus groups of Tate.
Type Species Thylamys elegans [Didelphis
elegans Waterhouse, 1839] by monotypy.
Distribution (Fig. 84) Peru south to Chile
west of the Andes, east of the Andes from Bolivia
to Argentina, Paraguay, and scattered locations in
eastern and western Brazil.
Thylamys elegans (Waterhouse, 1839)
1 839:95 hortensis
1844:154 CHILE:
-CHILE. Gardner,
Didelphis elegans Waterhouse,
Reid in synonymy. Bridges,
Aconcagua. Gay, 1847:84-
1993:23 listed.
Didelphys elegans: Waterhouse, 1 84 1 : 1 06 CHILE:
Valparaiso. Baird, 1855:155 CHILE: Valparaiso
(Colchagua [= Cachagua]). Thomas, 1888b:352
part, CHILE: Aconcagua; Valparaiso.
Micoureus elegans: Lesson, 1842:186 classification.
Thylamys elegans: Gray, 1843:101 CHILE: Valpa-
raiso. Gerrard, 1862:140 CHILE.
Marmosa elegans: Thomas, 1902c: 158, 160
CHILE: Valparaiso. Wolffsohn and Porter, 1908:
69 CHILE: Valparaiso (Quillota). Kreig, 1925:
169 CHILE. Reig et al., 1977:21 1 CHILE: Val-
paraiso (Cachagua, 3 km S). Pine et al., 1979:
340 CHILE: Tarapacd (Belen; Chapiquina;
Tignamar; Zapahuira); Maule (Cerro Robles); Val-
paraiso (Penuelas Forest Preserve, 7 km E of Las
Tablas); Santiago (Bocatoma [= Lo Valdes]; Em-
balse Huechun; Las Condes; Rio Colorado, 30 km
E of Guayacan); Atacama (Mina Altamira; Cordil-
leras del Transito near Vallenar); Coquimbo (Ilia-
pel; Los Vilos); Los Lagos (Rio Maule, junction of
Rio Claro).
M[armosa] elegans: Wolffsohn, 1921:512 CHILE.
[Marmosa] elegans: Wolffsohn, 1923:160 CHILE:
Valparaiso (Valparaiso; Quilpue).
[Marmosa (Thylamys) elegans] elegans: Cabrera,
1 9 1 9:40 classification.
Marmosa elegans elegans: Tate, 1933:214 CHILE:
Valparaiso (La Cruz; Limache; Papudo; Longato-
ma; Quilpue; Olmue; Calera [= La Calera], Lagu-
na); Santiago (coast hills, not located; La Cruz; Pu-
ente Alto, Rio Maipo). Osgood, 1943:44 CHILE:
Valparaiso (near Calera [= La Calera]; Longato-
ma; Papudo; Limache; Olmue; Palmilla [= La Pal-
milla]); Santiago (La Cruz); O'Higgins (Palos Que-
mados). Mann, 1950:5 CHILE: Tarapacd.
M[armosa] elegans elegans: Schneider, 1946:68
CHILE: Concepcion.
Marmosa (Thylamys) elegans: Reig et al., 1972:37
CHILE: Valparaiso (Cachagua).
Didelphys soricina R. A. Philippi, 1894:36 CHILE:
Los Lagos (type locality, Valdivia; holotype, very
young, skin mounted with skull in Museo Nacional
de Chile, collected by R. A. Philippi). Gardner,
1993:23 listed.
[Peramys] soricina: Trouessart, 1898:1244 classifi-
cation.
[Dromiciops] soricina: Matschie, 1916:271 classi-
fication.
Dromictops soricina Cabrera: 1958:30 incorrect
subsequent spelling in synonymy of Marmosa ele-
gans soricina Philippi, p. 29.
Didelphys (Paramys) soricina: Cabrera, 1958:29 in-
correct subsequent spelling in synonymy of Mar-
mosa elegans soricina Philippi.
Marmosa elegans soricina: Tate, 1933:216 CHILE:
Araucania (Angol [now in Malleco]).
Marmosa elegans coquimbensis Tate, 1931:14
CHILE: Coquimbo (type locality, Paiguano, 1000
m; holotype, female, skin and skull, FMNH 23302,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
137
Thylamys
Fig. 84. Distribution of the genus Thylamys: west of the Andes in Peru and Chile: southern Bolivia through
Argentina and Paraguay, and east-central Brazil south to the state of Parana.
138
FIELDIANA: ZOOLOGY
20'
30'
40
50
1 j
1
t
1* 1
-) (
\ # 1
hi
Thylamys elegans
500 1000 1500 km
90 80 70" 60 50 40 30 20 10
Fig. 85. Distribution of Thylamys elegans: Chile, west of the Andes. = localities given in published records.
collected 20 June 1923 by C. C. Sanborn). Gardner,
1993:23 listed.
Holotype Male, skin and skull, BMNH
53.8.29.18; collected by Charles Darwin, accord-
ing to Waterhouse in October, but Darwin reached
Valparaiso 11 March 1835 and left on horseback
for Coquimbo 27 April 1835.
Type Locality Valparaiso, Valparaiso, Chile.
Distribution (Fig. 85) Chile.
Thylamys macrura (Olfers, 1818)
Micoure a queue longue Azara, 1801:290 descrip-
tion.
Micoure nain Azara, 1801:304 description.
Colilargo Azara, 1802:251. Azara, 1809:289 char-
acters.
Enano Azara, 1802:262 description.
Micoure sixieme, ou Micoure nain Azara, 1801:304
description.
[Didelphys] macrura Illiger, 1815:107 nomen nu-
dum. Olfers, 1818:205 based on micoure a queue
longue of Azara. Gardner, 1993:23 listed.
Thylamys macrura: Palma, 1995:2 PARAGUAY:
Concepcion (Escuela Agropecuaria).
Thylamys macrurus: Anderson, 1997:163 incorrect
subsequent spelling BOLIVIA: Santa Cruz.
D[idelphis] grisea Desmarest, 1827:393 PARA-
GUAY: Central (type locality, Tapua, NE of Asun-
cion); based on the micoure a longue queue of Aza-
ra. Thomas, 1894b: 184 ARGENTINA: Corrien-
tes (Goya).
[Marmosa] grisea: Trouessart, 1898:1241 classifi-
cation.
Marmosa grisea: Bertoni, 1914:69 PARAGUAY.
Bertoni, 1939:6 PARAGUAY.
[Thylamys] grisea: Matschie, 1916:271 listed.
Grymaeomys griseus: Burmeister, 1856:83 classifi-
cation.
Holotype None designated.
Type Locality "Sudamerica," restricted by
Tate (1933:218) to "Tapoua" (= Tapua), Presi-
dente Hayes, Paraguay.
Distribution (Fig. 86) Paraguay and northern
Argentina.
Comments Based on Azara's (1801:290)
"Micoure a queue longue.''''
BROWN: ATLAS OF NEW WORLD MARSUPIALS
139
Fig. 86. Distribution of Thylamys macrura: Paraguay and northern Argentina. = localities given in published
records.
Thylamys pallidior (Thomas, 1902)
Grymaeomys elegans: Burmeister (not Waterhouse,
1839), 1856:83 part, figure and description of an-
imal from Mendoza.
Didelphys elegans: Burmeister (not Waterhouse,
1839), 1861:412 ARGENTINA: Mendoza. Bur-
meister, 1879:193 part, ARGENTINA: Mendoza.
Marmosa elegans: Thomas (not Waterhouse, 1839),
1902f:230 BOLIVIA: Oruro (type locality Chal-
lapata, 3700 m). Reig et al., 1977:21 1 PERU: Ar-
equipa (Arequipa); Huancavelica (Ticrapo).
[Thylamys] pallidior: Matschie, 1916:271 classifi-
cation.
Thylamys pallidior: Yensen and Tarifa, 1993:51
BOLIVIA: Oruro (Quebrada Kohuiri; Loma Jincho
Asjasta). Palma, 1995:2 BOLIVIA: Chusquisaca
(Camargo); Tarija (Serrania Sama). Mares et al.,
1996:107 ARGENTINA: Tucumdn (El Bracho;
El Cadillal; Las Mesadas; San Pedro de Colalao;
Tafi del Valle; Tapia). Mares et al., 1997: 100 AR-
GENTINA: Catamarca (Andalgala; Rib Andalgala;
Belen; Chumbicha; Minas Capillitas; Highway 62;
Quiros; Otro Cerro). Anderson, 1997: 164 BO-
LIVIA: Chuquisaca (Camargo, 68 km N); Cocha-
bamba (Jamachuma; Rodeo, 7.5 km E); La Paz (La
Paz; Huajchilla; Huajchilla, 4 km SW; Mecapaca;
Caracato; Huaraco); Oruro (Sajama; Quebrada Ko-
huir; Pazna; Challapata; Luca); Potosi (Yuruma);
Santa Cruz (Comarapa, 5 km SE); Tarija (Serrania
Sama; Iscayachi, Rio Tomayapo; Camataqui, 25
km SSE). Flores et al., 2000:327 ARGENTINA:
Catamarca (Belen; Chumbicha; Hualfin; Inmed. N
Andalgala, Rib Andalgala; Minas Capillitas; An-
dalgala, highway E of; Otro Cerro; Quiros); Jujuy
( 1 1 km E of Humahuaca, 2 km E of Pucara on road
to Cianzo; Humapuaca Barcena, 9 km NW; Abra
Pampa; Alfarcito; Casabindo; Cerro Casabindo;
Cuesta del Huron, 29 km W of Cineguillas on road
64; Curques, 24 km N of Susques on road 74; La
Quiaca, 17 km W, 3 km S; Laguna de Pozuelo;
Maimara; Mina Pirquitos, 31 SE, Sierra de Qui-
chagua; Miyuyoc; Salar Cauchari, 31 km N of Cau-
cahri; Sierra de Tilcara); Salta (Cachi, 30 km E);
Tucumdn (Amaicha del Valle, near; Pichao, Sierra
de Quilmes; Tafi del Valle); BOLIVIA: Oruro
(Challapata). Mares and Braun, 2000:36 AR-
GENTINA: Catamarca (Andalgala, N of, Rio An-
dalgala; Belen; Chumbicha; Otro Cerro; Minas
Capillitas; Quiros, Highway 62, 8 km E of junction
of Highways 1 and 62; Gualfin [probably = Hual-
fin]); Mendoza (El Sosneado, 8 km NW; Tre 40,
10.5 km W on road to Lago Diamonte; Malargue,
12.8 km E; Mendoza, 37 km N along Villavicinia
140
FIELDIANA: ZOOLOGY
Highway 4; Nacunan; Pareditas, 35 km S; Puesto
Punta de Agua; Refugio Militar Gral. Alvarado, 3
km W, not located; Salinas del Diamante R.R. Sta-
tion; Salinas del Diamante; Tunuyan, 31-32 km W;
Tunuyan, 50 km W; Uspallata, 7 km S; Valle
Grande Dike on Hwy 73; 25 de Mayo, 402 km W
on Route 150); San Juan (Castano Nuevo; Com-
plejo Astronomico El Leoncito, not located; Estan-
cia Leoncito W of Observatorio Astronomico; Jose
Marti; 17 km ESE; Quebrada de Las Flores; Tud-
cum, "Nacedero"; Villa Nueva, 9 km NW; km 514,
6 km N on Hwy 20); San Luis (Hualtaran; Parque
Provincial Sierra de las Quijadas; Paso del Rey,
Arroyo de la Canada Honda; Quebrada de Lopez,
San Francisco del Monte de Oro; Rio Gomes, 7 km
E of San Francisco del Monte de Oro; Salinas del
Bebedoro, 15 km E, 15 km SE; Varela, 12 km N);
Salta (Cachi, 30 km E); Tucumdn (El Bracho; El
Cadillal; Las Mesadas; San Pedro de Cololao);
Buenos Aires (Laguna Chasico; Vivero Chasico);
Chubut (Punta Norte; Peninsula Valdes, 3 km S;
Dolaron, ca. 200 km W; Dolaron, 280 km W; Isla
de los Pajaros; Istmo Ameghino; Puerto Lobos;
Puerto Piramides); Jujuy (Humahuaca, 1 1 km E, on
road to Cianzo; Abra Pampa; Alfarcitos; Casabin-
do; Laguna de Pozuelos; Maimara); La Pampa
(Parque Nacional Lihue Calel); La Rioja (Cerro [ =
Loma de] Famatina; La Invernada; Pagancillo; Pa-
tquia; Poterillo); Mendoza (Cacheuta; Callao, not
located; Challao; Colonia Alocar Rio Atuel; De-
saguadero; La Paz; Las Heras; Loncovaca; Lujan
de Cuyo; Nacunan; Punta del Agua; Santa Rosa;
Tunuyan; Tupungato); Neuquen (Chos Malal; Col-
Ion Cura; Las Lajas; Parque Nacional Laguna Blan-
ca; Zapala); Rio Negro (Estancia Maria Sofia; Gen-
eral Roca); San Juan (Reserva Natural Estricta El
Leoncito); San Luis (Alto Pencoso; Parque Nacion-
al Sierra de las Quijadas; Villa Valeria); Santiago
del Estero (Esteros; Lago Muyo; La Valle); Tucu-
mdn (Dique de Cadillal; El Bracho; Las Mesadas,
Trancas; Tafi del Valle; Tapia; Tucuman, 90 km
NNW). Solari, 2001 (pers. comm.) PERU: Ancash
(Huaras; Pariacoto; Chasquitambo); Lima (Meta-
cana).
Marmosa pallidior: Tate, 1933:229 BOLIVIA: Oru-
ro (Challapata); La Paz; Tarija (Estacion Yuruma
[now in Potosi]); ARGENTINA: Jujuy (Maimara;
Abra Pampa; Casabindo); Tucumdn (Tafi del Val-
le); Santiago del Estero (La Valle; Esteros); Cata-
marca (Otro Cerro; Chumbicha; Gualfin [= Hual-
fin]); La Rioja (La Invernada; Potrerillo); Mendoza
(Tupungato; Colonia Alocar, Rib Atuel); Neuquen
(Chos-Malal; Collon Cura; Las Lajas); Rio Negro
(General Roca). Yepes, 1936a:699 ARGENTI-
NA: Mendoza.
Marmosa elegans pallidior Thomas, 1902c: 159,
161 BOLIVIA: Oruro (Challapata). Thomas,
191 3a: 143 ARGENTINA: Jujuy (Maimara, 2230
m). Thomas, 1919c: 1 18 ARGENTINA: Catamar-
ca (Chumbicha). Thomas, 1919b: 135 ARGENTI-
NA: Jujuy (Abra Pampa, 3500 m; Casabindo, 4000-
4500 m). Thomas, 1920d:422 ARGENTINA: La
Rioja (La Invernada; Potrerillo; Sierra de Famati-
na). Thomas, 1 92 le:6 17 ARGENTINA: Jujuy
(Alfarcito, 2600 m). Thomas, 1926f: 195 BOLIV-
IA: Potosi (Yuruma). Thomas, 1926g:641 AR-
GENTINA: Neuquen (Chos-Malal, 805 m; Las La-
jas, 640 m; Zapala, 1062 m). Thomas, 1927d:657
ARGENTINA: Neuquen (Zapala). Thomas, 1927c:
202 ARGENTINA: Neuquen (Collon Cura). Ma-
res et al., 1981:165 ARGENTINA: Salta (Que-
brada de San Lorenzo).
M[armosa] e[legans] pallidior: Osgood, 1943:46-
47 taxonomy.
Marmosa pusilla pallidior: Cabrera, 1958:32 clas-
sification. Olrog, 1959:407 ARGENTINA: La
Rioja (Cerro Famatina [= Loma de Famatima],
3200 m); Tucumdn (Dique de Cadillal, 700 m; Las
Mesadas, Trancas, 2000 m); Jujuy (Abra Pampa,
3500 m). Olrog, 1979:9 ARGENTINA: Jujuy
(Cerro Calilegua).
Marmosa {Thylamys) pusilla pallidior: Anderson et
al., 1993:18 BOLIVIA: Santa Cruz (Santa Cruz
de la Sierra [= Santa Cruz]). Collected by P. O.
Simons; PERU: Lima (Surco, near Lima).
Marmosa venusta venusta: Tate, 1933:225 PERU:
Lima (Matucana, Rio Rimac; Surco, Rfo Rimac).
Marmosa bruchi Thomas, 1 92 la:5 19 ARGENTI-
NA: San Luis (type locality, Alto Pencoso, W of
San Luis; holotype, young adult male, skin and
skull, BMNH 21.4.21.8, collected February 1914
by Carlos Bruch). Tate, 1933:231 ARGENTINA:
San Luis (Alto Pencoso); Cordoba (Villa Valeria).
Yepes, 1936b:34 ARGENTINA: La Rioja (Pa-
gancillo; Patqufa); San Luis (Alto Pencoso [para-
type]). Gardner, 1993:23 listed.
Marmosa pusilla bruchi: Cabrera, 1958:32 classifi-
cation. Olrog, 1959:407 ARGENTINA: Tucumdn
(El Bracho, 400 m); Santiago del Estero (Lago
Muyo). Contreras, 1973:215 ARGENTINA: Chu-
but (Laguna Chasico). Crespo, 1974:3 ARGEN-
TINA: Santa Cruz (Isla de los Pajaros); San Luis
(Alto Pencoso). Daciuk, 1974:23 ARGENTINA:
Santa Cruz (Isla de los Pajaros).
Marmosa elegans fenestrae Marelli, 1932:68 AR-
GENTINA: Buenos Aires (type locality, Sierra de
la Ventana; holotype presumably an individual
once living in the La Plata Zoological Garden).
Gardner, 1993:23 listed.
Marmosa pulchella: Cabrera, 1958:32 name com-
bination in synonomy of Marmosa pusilla bruchi
Thomas.
Marmosa tatei Handley, 1957:402 PERU: Ancash
(type locality, Chasquitambo, 70 m; holotype,
male, skin and skull from specimen originally pre-
served in spirits, USNM 302915; collected Decem-
ber 1955 by Jose Maria de la Barrera). Gardner,
1993:23 listed.
Note: Sergio Solari in his master's thesis in prepara-
tion recognizes M. tatei as a separate species [pers.
comm.].
Holotype Young adult male, skin and skull,
BMNH 2.2.2.116, collected 26 October 1901 by
P. O. Simons.
Type Locality Challapata, near Lake Poopo,
Oruro, Bolivia, elevation 3800 m.
Distribution (Fig. 87) Peru, Bolivia, and Ar-
gentina.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
141
Thylamys pallidior
xDs
/..
500 1000 1500 km
90
80
40
30
20
10
Fig. 87. Distribution of Thylamys pallidior: Peru. Bolivia, and Argentina. = localities given in published
records.
142
FIELDIANA: ZOOLOGY
Thylamys pusilla (Desmarest, 1804)
Didelphis pusilla: Desmarest, 1804:19. Gardner,
1993:23 listed.
Didelphis pusila: Desmarest, 1816:430 incorrect
subsequent spelling.
Sarigua pusilla: Muirhead, 1819:429 new combi-
nation.
Didelphys pusilla: Waterhouse, 1841:112 incorrect
subsequent spelling.
Peramys pusilla: Lesson, 1842:187 classification.
[Cuica] pusilla: Liais, 1872:330 classification.
M[armosa] pusilla: Thomas, 1898a:455 PARA-
GUAY.
Marmosa pusilla: Bertoni, 1914:69 PARAGUAY:
Alto Parana (Puerto Bertoni). Tate, 1933:221
ARGENTINA: Corrientes (Goya; Estancia Coropa,
Goya); Entre Rios (La Paz); Salta (40 km NNW of
Tucuman). Yepes, 1938:41 PARAGUAY; AR-
GENTINA: Entre Rios; Salta (90 km NNW of Tu-
cuman) BRAZIL: Rio Grande do Sul. Bertoni,
1939:6 PARAGUAY: Alto Parana (Puerto Ber-
toni; Rio Parana, not located); agilis Burmeister re-
garded as a variety. Reig et al., 1977:201 AR-
GENTINA: Buenos Aires (Vivero Chasico, Chasi-
co). Mares et al., 1981:160 ARGENTINA: Salta
(Tolloche; Cachi [now T. pallidior, see Mares and
Braun, 2000:39]). Crespo, 1928:33 ARGENTI-
NA: Entre Rios (Parque Nacional El Palmar).
Thylamys pusilla: Mares and Braun, 2000:38 AR-
GENTINA: Chaco (El Mangrullo, 20 km N and
NW and 1 1 km NE, not located); Formosa (Ingen-
iero Guillermo N. Juarez, 35 km S, Puesto Divi-
sadero); Salta (Hickman; Piquirenda Viejo, 6 km
W; Tolloche, 5 km S); Santiago del Estero (Pampa
de los Guanacos, 6 km S, 2 km E; Virgen del Valle
picnic area on Highway 64 between Santa Catalina
and La Puerta Chiquita); Corrientes (Estancia Cor-
opa, Goya; Goya); Entre Rios (La Paz); Formosa
(Parque Nacional Rio Pilcomayo). Flores et al.,
2000:328 ARGENTINA: Salta (Tolloche, 5 km S
on Vinalito Road; Piquirenda Viejo, 6 km W; Tu-
cuman, 90 km NNW; Campo Grande, 17 km E of
Santo Domingo; Departamento Capital; Dragones;
Hickman; Los Colorados, 17 km E of Santo Do-
mingo; Puesto Campo Grande, 17 km E of Santo
Domingo); Santigo del Estero (Picada de Olmos,
15 km SE, W of Los Pirpintos; Pampa de los Gua-
nacos, 6 km S, 2 km E; Estancia Guampacha; Es-
teros; Lavalle [= La Valle]; Lago Muyo; Virgen
del Valle picnic area on Highway 64 between Santa
Catalina and La Puerta Chiquita; Robles; San An-
tonio; Villa La Punta); Tucuman (Cerro Medici; El
Bracho; El Cadillal; Estacion Vipos [= Vipos]; Las
Mesadas; San Pedro de Colalao; Tapia).
Marmosa pusilla pusilla: Cabrera, 1958:33 classifi-
cation. Olrog, 1959:408 ARGENTINA: Salta
(Hickmann). Wetzel and Lovett, 1974:203 PAR-
AGUAY: Nueva Asuncion (Teniente Ochoa); Pres-
idente Hayes (Juan de Zalazar [= Fortin Juan de
Zalazarj).
Grymaeomys pusillus: Burmeister, 1856:83 classifi-
cation.
Philander pusillus: Cope, 1889:130 BRAZIL: Mato
Grosso (Chapada [= Chapada dos Guimaraes]).
Thylamys pusillus: Morando and Polop, 1997:131
ARGENTINA: Cordoba. Anderson, 1997:164
BOLIVIA: Chuquisaca (Tarabuco, 12 km N, 1 1 km
W; Padilla, 9 km N; Carandayti, 3.8 km E); Santa
Cruz (Santa Cruz de la Sierra; Puesto Perforacion;
Boyuibe, 53 km E); Tarija (Villa Montes, 8 km S
and 10 km E; Villa Montes, 35 km SE; Entre Rios;
Estancia Bolivar).
Marmosa janetta pulchella Cabrera, 1934:126 AR-
GENTINA: Santiago del Estero (type locality, Ro-
bles; holotype, female, skin and skull, Museo de
La Plata, collected by Jorge Arganarez). Gardner,
1993:23 listed.
[Didelphys] nana Illiger, 1815:107 nomen nudum.
D[idelphys] nana Olfers, in Eschwege, 1818:206
renaming of pusilla Desmarest. Gardner, 1993:23
listed.
Did[elphis] nana: Schinz, 1821:252 PARAGUAY.
[Marmosa] nana: Trouessart, 1898:1241 listed in
synonymy of Marmosa pusilla.
Marmosa marmota Thomas, 1896:footnote, pp. 313
314 name based on Didelphys marmota Oken,
1816 [a nonbinomialist, hence valid only as of
Thomas, 1896; regarded as a senior synonym of
grisea Desmarest]. Thomas, 1902c: 158 ARGEN-
TINA: Corrientes (Goya); grisea Desmarest a syn-
onym. Gardner, 1993:23 listed.
M[amosa] marmota: Thomas, 1912d:410 PARA-
GUAY: Paraguari (Sapucay [= Sapucaf]); Central
(Asuncion).
Marmosa marmota marmota: Tate, 1933:218 PAR-
AGUAY: Paraguari (Sapucay [= Sapucaf]; Cen-
tral (Asuncion); grisea Desmarest a synonym.
Marmosa citella Thomas, 191 2d:409 ARGENTI-
NA: Corrientes (type locality, Goya, 181 m; ho-
lotype, young adult male, skin and skull, BMNH
98.8.19.9, collected 25 July 1895 by R. Perrens).
Gardner, 1993:23 listed.
[Thylamys] citella: Matschie, 1916:271 listed.
Marmosa verax Thomas, 1 92 la:520 PARAGUAY:
Chaco (type locality, Misiones, W of Concepcion,
not located; holotype, female, skin and skull,
BMNH 20.12.18.34, collected 29 July 1920 and
presented by the Marquis de Wavrin). Gardner,
1993:23 listed.
Marmosa marmota verax: Tate, 1933:220 PARA-
GUAY: Chaco (Mision [= Misiones], not located;
Bermalcue, not located).
[Marmosa marmota] verax: Krumbiegel, 1941c:9
PARAGUAY: Boqueron (Esteros, Rio Pilcomayo);
ARGENTINA: Formosa (La Urbana [= Tapikiole],
Rio Pilcomayo).
Marmosa elegans: Bertoni (not Waterhouse), 1923:
51 PARAGUAY. Bertoni, 1939:6 PARAGUAY:
Central (Asuncion).
Marmosa pallidior: Tate, 1933: 229 ARGENTINA:
Salta (Tucuman, 90 km NNW; Estacion Vipos;
Tapia); BOLIVIA: Santa Cruz (Santa Cruz de la
Sierra) [= Santa Cruz]); ARGENTINA: Santiago
de Estero (La Valle; Esteros; Villa Montes, not lo-
cated).
Thylamys pallidior: Mares and Braun, 2000:36 AR-
GENTINA: Salta (Tucuman, 90 km NNW).
Marmosa karimii Petter, 1968:313 BRAZIL: Per-
nambuco (type locality, Exu; holotype, male, skin
and skull, no. 1968-148, Museu National d'Histoire
BROWN: ATLAS OF NEW WORLD MARSUPIALS
143
Thylamys pusilla
90 80
70 60 50 40
Fig. 8
south to
1. Distribution of Thylamys pusilla: eastern and central Brazil, southern Bolivia. Paraguay, and Argentina
Rio Negro. = localities given in published records.
144
FIELDIANA: ZOOLOGY
Naturelle, Paris, collected by Dr. Y. Karimi). Pine
et al., 1970:669 BRAZIL: Mato Grosso (Xavati-
na, 264 km N). Mares et al., 1981:101 BRAZIL:
Pemambuco (Escola Agricola de Exu; Exu; Fazen-
da Guarani).
Holotype Not known to be in existence;
name based on the micoure nain of Azara.
Type Locality Saint-Ignace-Gouazou [= San
Ignacio Misiones]; southern Paraguay.
Distribution (Fig. 88) Central and eastern
Brazil, Paraguay, and southern Bolivia south to
Rio Negro, Argentina.
Thylamys velutinus (Wagner, 1842)
Didelphys velutina Wagner, 1842:360. Burmeister,
1854:142 BRAZIL: Minus Gerais (Lagoa Santa).
Pelzeln, 1883:115 BRAZIL: Sao Paulo (Ipane-
ma). Thomas, 1888b:352 BRAZIL: Sao Paulo.
Gardner, 1993:23 listed.
Macrodelphys velutina: Burmeister, 1856:86 BRA-
ZIL: Minas Gerais (Lagoa Santa).
[Marmosa] velutina: Trouessart, 1898:1241 classi-
fication.
[Thylamys] velutina: Matschie, 1916:271 listed.
Marmosa velutina: Tate, 1933:234 BRAZIL: Sao
Paulo (Ipanema [type]); Minas Gerais (Lagoa San-
ta). Vieira, 1949:351 BRAZIL: Sao Paulo. Vi-
eira, 1949:351 BRAZIL: Sao Paulo. Vieira,
1955:351 BRAZIL: Sao Paulo (Capital, not lo-
cated; Ipanema).
Marmosa velutina velutina: Cabrera, 1958:34 clas-
sification.
Grymaeomys velutinus: Winge, 1893:29.
Peramys velutinus: Ihering, 1894:11 BRAZIL: Sao
Paulo.
Didelphys pimelura Reinhardt, 1851:5 BRAZIL:
Minas Gerais (type locality, Lagoa Santa; holotype,
male, skin mounted with skull in Copenhagen Mu-
seum no. 164). Gardner, 1993:23 listed.
[Marmosa] pimelura: Trouessart, 1898:1242 listed
in synonymy of Marmosa elegans Waterhouse.
Holotype Male, skin and skull, Vienna Zoo-
logical Museum no. 81; collected 10 June 1821
by Johann Natterer.
Type Locality Ipanema, Sao Paulo, Brazil.
Distribution (Fig. 89) Southeastern Brazil.
Thylamys venustus (Thomas, 1902)
Marmosa elegans: Thomas (not Waterhouse, 1839),
1898c:4 BOLIVIA: Tarija (Caiza). Thomas (not
Waterhouse, 1839), 1900b:302) ARGENTINA:
Tucumdn (Tucuman, Dinelli, collector). Thomas,
1902a: 143 BOLIVIA: Cochabamba (Parotani).
Thomas, 1902b:230 Bolivia: Chuquisaca (Sucre).
Thylamys elegans: Heinonen and Bosso, 1994:55 (not
Waterhouse, 1839) ARGENTINA: Jujuy (Duraz-
nillar, Parque Nacional Calilegua). Mares et al.,
1996:106 ARGENTINA: Catamarca (Las Hig-
uerillas, 5 km N); Tucumdn (Aconquija; Burruya-
cu; Cafayate, 45 km S; Cerro del Campo; Cerro de
Taff Viejo; Cerro San Javier; Concepcion; El Cad-
illal; Biological Reserve at Horco Molle; 4 km W
of junction Highway 338 and road to Horco Molle;
road to San Javier; Horco Molle; La Higuera; Lu-
les; Norco, Vipos; Piedra Tendida; San Pablo; km
42, Highway 364 S of San Pedro de Colalao; San
Pedro de Colalao, 1 1 km S; San Miguel de Tucu-
man, 17 km NW; Tucuman; Villa Nougues, Vipos;
Cerro de Raco, Trancas). Mares et al., 1997:100
ARGENTINA: Catamarca (Las Higuerillas). Ca-
pllonch et al., 1997:53 ARGENTINA: Tucumdn
(Cerro de Taff Viejo; Horco Molle; Reserva Biol-
ogica de Horco Molle, cerca de la Residencia; San
Javier; Taff Viejo; Cerro de Raco; Highway 338
and road to Horco Molle [junction] on road to San
Javier; Villa Nougues).
Marmosa elegans venusta Thomas, 1902c: 159, 160
BOLIVIA: Cochabamba (Parotani); Chuquisaca
(Sucre). Osgood, 1916:200 BOLIVIA: Cocha-
bamba (Parotani). Olrog, 1959:405 ARGENTI-
NA: Salta (Aguary; Oran); Jujuy (Yuto). Mares et
al., 1981:165 ARGENTINA: Salta (Aguas Blan-
cas [= Agua Blanca]; Juntas de San Antonio; Tar-
taga). Gardner, 1993:23 listed.
Marmosa (Thylamys) elegans venusta: Anderson et
al., 1993:17 BOLIVIA: Santa Cruz (Vermejo [ =
Bermejo]; Buena Vista; Vallegrande).
[Thylamys] venusta: Matschie, 1916:271 classifica-
tion.
Thylamqs venusta: Cabrera, 1958:30 incorrect sub-
sequent spelling in synonymy of Marmosa elegans
venusta Thomas.
Marmosa venusta venusta: Tate, 1933:225 BOLIV-
IA: Cochabamba (Parotani); Chuquisaca (Sucre);
Santa Cruz ("Vermejo" [= Bermejo]).
Thylamys venustus: Hefnonen and Bosso, 1994:55
ARGENTINA: Jujuy (Duraznillar, Parque Nacional
Calilegua; Aguas Negras, Parque Nacional Calile-
gua; Caimancito, Parque Nacional Calilegua; Mes-
ada de las Colmenas, Parque Nacional Calilegua;
Abra de Cafias, Parque Nacional Calilegua; Cerro
Hermosa, Parque Nacional Calilegua). Flores et al.,
2000:331 ARGENTINA: Salta (Agua Blanca, 27
km W; Angosto del Rfo Pescado; Finca Yakulica,
Angosto del Rfo Pescado; Arroyo Santelmita,
Parque Nacional Baritu; Cerro Chaguar, Parque Na-
cional Baritii); BOLIVIA: Cochabamba (Paratani).
Anderson, 1997:165 BOLIVIA: Chuquisaca (Su-
cre; Horcus; Rfo Limon; Monteagudo, 2 km SW;
Porvenir; Carandayti, 3.8 km E); Cochabamba
(Cochabamba; Vacas; Paratani; Rodeo, 7.5 km SE;
Totora, Tinkusiri, 17 km E); La Paz (Huajchilla;
Mecapaca); Oruro (Oruro; Oruro, 64 km S; Chal-
apata); Santa Cruz (Buenavista; Tambo, 2.5 km N;
Quifie [= Estancia Jahue], 6 km NNE; Quifie, 5
km NE; Vermejo; Quine [= Estancia Laja], 17 km
S; Ariruma, 7 km SE; Tita; Vallegrande, Rfo Cie-
nega, 5.5 km NNE; Vallegrande; Gutierrez; Cerro
Colorado; Cerro Itahuatica; Camiri, 10 km S); Tar-
ija (Camatindi; Camataqui, 40 km SSE; Tapecua;
Rancho Tambo; Tucumilla; Pino; Carlazo; Erquis;
BROWN: ATLAS OF NEW WORLD MARSUPIALS
145
20
30
40
50
Thylamys velutinus
90 80 70 60 50 40 30 20 10
Fig. 89. Distribution of Thylamys velutinus: southeastern Brazil. = localities given in published records.
Entre Ribs, 5 km NNW; Tarija; Tablada; Tarija, 8
km S; Padcaya, 1 1.5 km N and 5.5 km E; Carapari,
3 km WNW; Caiza; Carapari; Yacuiba; Cuyambu-
yo, 8 km N; Cuyambuyo, 4 km N; Cuyambuyo, 3
km SE); La Paz (Palos Blancos, 2 km S, 5 km E).
Mares and Braun, 2000:39 ARGENTINA: Cata-
marca (Las Higuerillas, 5 km S on Highway 9);
Jujuy (Salta, Highway 9, campground on way to El
Carmen; El Simbolar; Laguna La Brea; El Palmar,
Sierra de Santa Barbara, E of; Highway 29, 10 km
W of Tiraxi; Highways 34 and 3, junction of; Cal-
ilegua; Caimancito; Cerro Calilegua; Jujuy; Leon;
Parque Nacional Calilegua; Sunchal, Sierra de San-
ta Barbara; Vala [= Yala]; Villa Carolina; Yuto);
Salta (Juntas de San Antonio; Oran, 15 km S and
15 km W; Agua Blanca, 24 km NW; Salta, 12 km
NW; Tartagal, 27 km E; Arroyo de los Noquez,
Parque Nacional El Rey; Campo Duran, 8 km S, 8
km E; Aguaray; Parque Nacional Baritii; Parque
Nacional El Rey); Tucumdn (Aconquija; Burruya-
cii; Cafayate, 45 km S; Cerro del Campo; Cerro de
Taff Viejo; Cerro San Javier [= Parque Biologica
Sierra de San Javier]; Concepcion; El Cadillal; Bi-
ological Reserve at Horco Molle; Highway 338 and
road to Horco Molle [junction] on road to San Ja-
vier [= Parque Biologica Sierra de San Javier];
Horco Molle; La Higuera; Lules; Norco, Vipos;
Piedra Tendida, Rio Cajon; San Pablo; Highway
354, km 42 S of San Pedro de Colalao; San Miguel
de Tucuman; San Pedro de Colalao, 11 km S; Tu-
cuman; Villa Nougues; Vipos; Cerro de Raco; Cer-
ro Medici; San Javier; Sierra [= Serra] de Tucu-
man; Taff Viejo; Estacion Vipos).
Marmosa elegans Cinderella, Thomas; 1902c: 159,
161 ARGENTINA: Tucumdn (type locality, Tu-
cuman; holotype, female, skin and skull, BMNH
0.7.9.205, collected 1 December 1899 by L. Di-
nelli). Thomas, 1918:193 ARGENTINA: Jujuy
(Leon). Thomas, 1 920a: 1 96 ARGENTINA: Jujuy
(Jujuy; Villa Carolina). Thomas, 1925:582 BO-
LIVIA: Tarija (Carapari). Thomas, 1926c:608
ARGENTINA: Tucumdn (Cerro del Campo, Bur-
ruyacu; Norco, Vipos; Cerro de Taff Viejo). Olrog,
1959:405 ARGENTINA: Tucumdn (Concepcion;
San Javier; Vipos). Gardner, 1993:23 listed.
M[armosa] e[legans] Cinderella: Osgood, 1943:46-
47 taxonomy.
Marmosa venusta Cinderella: Tate, 1933: 226 AR-
GENTINA: Tucumdn (Norco, Vipes; Villa Nouges,
San Pablo; Cerro de Taff Viejo; Concepcion; Tran-
cas; Cerro de Raco; Cerro del Campo, Burruyacii;
Sierra de Tucuman); Jujuy (Jujuy; Caimancito;
Leon); BOLIVIA: Tarija (Carapari).
[Thylamys] Cinderella: Matschie, 1916:271 listed.
Thylamys Cinderella: Flores et al., 2000:325 AR-
GENTINA: Jujuy (4 km W junction of Highways
34 and 3 on Highway 3; Caimancito; junction road
to Puesto Viejo and Highway 34, 4 km E on road
146
FIELDIANA: ZOOLOGY
to Puesto Viejo; Jujuy; Laguna La Brea; Leon; El
Carmen, campground on Highway 9; Oyeros, 3 km
N; Santa Rita, 2 km NW; Santa Rita, 4 km NW;
Villa Carolina, Rio Lavallen; Yuto); Salta (Agu-
aray; Giiemes, Highway 34, km 1383; Playa An-
cha; Rio Mojotoro, bridge of Ferrocarril Belgrano;
San Ramon de la Nueva Oran; Tartagal, Laguna de
Las Catas); Tucumdn (Cerro de Raco; Cerro de Tafi
Viejo; Cerro del Campo; Concepcion; Finca El Ja-
giiel, 4.5 km NW road 304 on way to Requelme;
Norco-Vipos; Raco; Tucuman; Villa Nougues, San
Pablo).
Marmosa elegans sponsoria Thomas, 1921c: 186
ARGENTINA: Jujuy (type locality Sunchal, Sierra
de Santa Barbara, 1200 m; holotype, male, skin and
skull BMNH 21.1.8.85, collected 23 June 1920 by
E. Budfn). Gardner, 1993:23 listed.
Marmosa venusta sponsoria: Tate, 1933:228 AR-
GENTINA: Jujuy (Sunchal, Sierra de Santa Bar-
bara; Higuerilla; near Vala; Calilegua; Leon; Cai-
mancito).
Thylamys sponsoria: Flores et al., 2000:330 AR-
GENTINA: Jujuy (Barcena, 9 km NW; Abra de
Canas; El Monolito, Abra de Canas; Aguas Negras
Arroyo La Horqueta, 3 km SE of Laguna de Yala
Arroyo La Horqueta, 6 km SE of Laguna de Yala
Arroyo Yuto, 13 km SW of Yuto; Caimancito; Cal-
ilegua; Cerro Hermoso; El Simbolar; Higuerilla;
Mesadas de las Colmenas; Mountains W of Yala;
near Yala; Highway 29, 10 km W of Tiraxi; Palma
Sola; Parque Nacional Calilegua; Rio Blanco, 9 km
SW San Antonio; Rio Tesorero; Rio Tiraxi; Santa
Barbara; Sunchal, Sierra de Santa Barbara); Salta
(Potrerillos; Highways 50 and 18, junction, 43.7
km NW on road to Isla Canas; Rio de Las Conchas;
Rio Santa Maria; Vado de Arrazayal); Tucumdn
(Aconquija; Biological Reserve Horco Molle, near
Residencia; Cerro San Javier; Concepcion; Horco
Molle; Las Agiiitas, El Taficillo; Parque Provincial
El Cochuna; Piedra Tendida; Reserva Provincial La
Florida, 7 km from Ibatin on Rio Pueblo Viejo; San
Javier; San Javier, Estacion Biologica R. Schreiter;
San Miguel de Tucuman; Villa Nougues).
Marmosa janetta Thomas, 1926b:327 BOLIVIA:
Tarija (type locality Carlazo, 2300 m; holotype,
male, skin and skull, BMNH 26.1.1.167, collected
26 February 1925 by Emilio Budfn). Tate, 1933:
220 BOLIVIA: Tarija (Carlazo; Pino [= Pinos);
Tallada [= Tablada]). Gardner, 1993:23 listed.
M[armosa] janetta: Handley, 1956:403 BOLIVIA.
Marmosa elegans janetta: Olrog, 1959:406 ARGEN-
TINA: Tucumdn (La Higuera, Trancas); Jujuy (Yuto).
Didelphys dorsigera: Burmeister (not Linnaeus),
1879:192 part, ARGENTINA: northern Jujuy.
Holotype Female, skin and skull, BMNH
2.1.1.120, collected by P. O. Simmons.
Type Locality Parotani, Cochabamba, Boliv-
ia, 2800 m.
Remarks The genus Thylamys is under revi-
sion; some authors (Flores et al., 2000) recognize
Cinderella and sponsoria as distinct species; oth-
ers (Mares and Braun, 2000) do not. Work is on-
going.
Distribution (Fig. 90) Bolivia and northern
Argentina.
II. Order Paucituberculata
Family Caenolestidae Trouessart, 1898
Genus Caenolestes Thomas, 1895
Hyracodon Tomes, 1863:50 name preoccupied by
Hyracodon Leidy, 1 856, a genus of Perissodactyla.
Gardner, 1993:25.
Caenolestes Thomas, 1895:367 substitute name for
Hyracodon Tomes, preoccupied. Gardner, 1993:
25 listed.
Type Species Hyracodon fuliginosus Tomes,
by original designation.
Distribution (Fig. 91) Temperate zone forests
of the Andes of western Venezuela, Colombia, and
Ecuador; elevational range from approximately
2100 to 4000 m (see Albuja and Patterson, 1996).
Caenolestes caniventer Anthony, 1921
Caenolestes caniventer Anthony, 1921a:6. Albuja and
Patterson, 1996:52 ECUADOR: El Oro (El Chi-
ral); Azuay (Molleturo); PERU: Piura (Cerro Chin-
guela; "Batan" on Zapalache-Carmen Trail; Huan-
cabamba). Gardner, 1993:25 listed, synonomy.
Holotype Male, skin and skeleton, AMNH
47174; collected 2 August 1920 by Harold E. An-
thony.
Type Locality El Chiral, Cordillera Occiden-
tal, El Oro, Ecuador, 5350 ft (= 1670 m).
Distribution (Fig. 92) Western Ecuador and
northwestern Peru.
Additional Records (compiled from Patter-
son and Albuja an unpublished monograph on
shrew-opossums). ECUADOR: Pichincha; Ver-
decocha (Anthony, 1923, AMNH); Canon del Rio
Pita (AMNH); San Antonio (Tate, 1922, AMNH);
Santa Rosa above Rio Pita (AMNH); Loja; Ca-
januma, Laguna Compadre (EPN); Azuay; Pin-
angu (Tate, 1922, AMNH; Wickenheiser, 1922,
AMNH); Pauji [= Paujchi] (Tate, 1922, AMNH;
Wickenheiser, 1922, AMNH); Mazan (Barnett,
1984, BMNH); El Oro; Cordillera de Chilla, Tar-
aguacocha (Anthony, 1920, AMNH); Tungura-
hua; San Antonio (AMNH); San Francisco (Tate,
1922, AMNH); ZamoralChinchipe; Zamora (An-
thony, 1923, AMNH).
BROWN: ATLAS OF NEW WORLD MARSUPIALS
147
20
30
40
50
Thylamys venustus
90
80"
70
60'"
50^'
40
30
20
10 L
Fig. 90. Distribution of Thylamys venustus: Bolivia and northern Argentina. = localities given in published
records.
Caenolestes condorensis Albuja and Patterson,
1996
Caenolestes condorensis Albuja and Patterson, 1996:
42 ECUADOR: Morona-Santiago (Achupallas).
Holotype Adult male, skin and skull. FMNH
152134; collected 23 July 1993 by Luis Albuja
and Alfredo Luna.
Type Locality "Achupallas," a place name
coined by members of the Rapid Assessment Pro-
gram expedition for their camp on the upper pla-
teau of the Cordillera del Condor in the Provincia
de Morona-Santiago, Ecuador, altitude 2080 m.
Distribution (Fig. 93) Found only in the Cor-
dillera del Condor, southeastern Ecuador.
Caenolestes convelatus Anthony, 1924
Caenolestes convelatus Anthony, 1924:1. Gardner.
1993:25 listed. Albuja and Patterson, 1996:52
COLOMBIA: Antioquia (Paramo Frontino, Caice-
do; Santa Barbara): ECUDAOR: Imbabura (Haci-
enda La Vega); Pichincha (Gualea. Ilambo Valley;
Galaya [= Saloya] West; Santa Rosa, Mindo; Las
Maquinas; El Hato, Antisana; El Castillo). Kirsch
and Waller, 1979:390 ECUADOR: Pichincha
(Aloag, 1 1 km W).
Holotype Male, skin and skull, AMNH
64462; collected 24 September 1923 by Harold E.
Anthony.
Type Locality Las Maquinas, on trail from
Aloag to Santo Domingo de los Colorados. Cor-
dillera Occidental, Ecuador, 7000 ft (= 2180 m).
Distribution (Fig. 94) Western Colombia and
northwestern Ecuador.
Additional Records (compiled from Patter-
son and Albuja an unpublished monograph on
shrew-opossums) COLOMBIA: Valle del Cau-
ca-Choco; Alto de Galapagos (UV); ECUADOR:
Pichincha; Old Santo Domingo Trail, Rio Saloya,
not located (UMMZ).
Caenolestes fuliginosus (Tomes, 1863)
H[yracodon] fuliginosus Tomes,
1993:25 listed.
1863:51. Gardner.
148
FIELDIANA: ZOOLOGY
Caenolestes
20
1500 km
Fig. 91.
Ecuador.
80" 70 60 50 3 30* 30
Distribution of the genus Caenolestes: extreme western Venezuela south through western Colombia and
C[aenolestes] fuliginosus: Thomas, 1895:367 type
of genus Caenolestes Thomas.
Caenolestes fuliginosus: Stone, 1914:17 ECUA-
DOR: Pichincha (Hacienda Garzon, Cerro Pichin-
cha). Lonnberg, 1921:70, 72 ECUADOR: Imba-
bura (Cerro Cotacachi, 8000 ft [= 2500 m]); Pi-
chincha (road to Nanegal, 6000 ft [= 1875 m]; Cer-
ro Illiniza, 12000 ft [= 3750 m]; La Carolina, 9400
ft [= 2930 m]). Tate, 1931:248 ECUADOR: sub-
paramo zone. Kirsch and Waller, 1979:390 EC-
UADOR: Pichincha (Volcan Pichincha, 3300 m).
Albuja and Patterson, 1996:52 COLOMBIA: An-
tioquia (Paramo Frontino, Paramo Sonson, 8800 ft
[= 2750 m]); Nariho (Pasto); Caldas (Rio Terma-
les; Termales del Ruiz); Valle del Cauca (Paramo
de Barragan); Cauca (Paramo de Guanacas; La-
guna San Rafael, Paramo de Purace); Quindio (Fin-
ca "Bengala," Carretera Salento-Cocora; Finca
Rincon Santo, Municipio Salento); Huila (Rio Ove-
jera; San Antonio; Santa Marta; Rio Magdalena);
Cundinamarca (Guasca, Rio Balcones; Finca El
Soche, Soacha; San Miguel-Sibate, Soacha; San
Cristobal; Bogota); Boyacd (Hacienda La Prima-
vera); Norte de Santander (Paramo de Tama); EC-
UADOR: Pichincha (Chinchin Cocha; Tablahuasi
occidente; Galaya [= Saloya Occidente]; Verde-
cocha; Quebrada de Pichan; Pichan; Inga Pirca;
San Ignacio, Volcan Pichincha; Faldas Iliniza; Fal-
das Occidentales del Pichincha; Volcan Pichincha,
oriente; Paramos of Chimborazo or Pichincha, not
located; Laderas Iliniza); Bolivar (Sinche, 1 1 100 ft
[= 3460 m]); Chimborazo (Cerro Chimborazo);
Napo (Cosanga, Rio Aliso; Cerro Antisana; Rio
Bermejo); VENEZUELA: Tdchira (San Cristobal);
COLOMBIA/VENEZUELA: Norte de Santander
(Paramo de Tama).
[Caenolestes] fuliginosus: Cabrera, 1919:98. Thomas,
1920c:246 ECUADOR: Pichincha (Gualea).
Caenolestes obscurus Thomas, 1895:367 COLOM-
BIA: Cundinamarca (Bogota) (type locality, Estan-
cia La Selva, Bogota; holotype, in spirits, BMNH
96.1.7.1, collected 7 May 1895 by G. D. Child).
Osgood, 1912:38 COLOMBIA: Norte de Santan-
der (Paramo de Tama). Lonnberg, 1921:72 EC-
UADOR: Pichincha (road to Gualea; La Carolina,
3 mi N of Quito, 9400 ft [= 2930 m]). Osgood,
1924:167 COLOMBIA: Norte de Santander
(Paramo de Tama). Niceforo Maria, 1925:47
COLOMBIA. Handley, 1976:9 VENEZUELA:
Tdchira (Buena Vista). Mendez, 1977:91 CO-
LOMBIA: Cauca (Parque de Purace); Cundina-
marca (El Soche). Kirsch and Waller, 1979:390
COLOMBIA: Cauca (Laguna San Rafael, Paramo
de Purace, 3000 m); Cundinamarca (Finca El So-
che, 2630 m). Tyndale-Biscoe, 1980:714 CO-
LOMBIA: Cauca (Paramo de Purace); Cundina-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
149
Fig. 92. Distribution of Caenolestes caniventer: western Ecuador and northwestern Peru, o
unpublished records; = localities given in published records.
localities from
marca (Paramos El Soche); COLOMBIA/VENE-
ZUELA: Norte de Santander Tdchira (Paramo de
Tama). Cuervo Diaz et al., 1986:474 COLOM-
BIA: Cundinamarca (Bogota, cerca de); Narino.
Gardner, 1993:25 listed. Albuja and Patterson,
1996:52 COLOMBIA: Cauca (Laguna de Pura-
ce).
[Caenolestes] obscurus: Cabrera, 1919:99 classifi-
cation.
C[aenolestes] Obscurus: Thomas, 1920c:247.
Caenolestes tatei Anthony, 1923:1 ECUADOR:
Azuay (type locality, Molleturo, 7600 ft [= 2375
m]; holotype, female, skin and skull, AMNH
61860; collected 11 June 1922 by G. H. H. Tate).
Gardner, 1993:25 listed.
Holotype Young individual in spirits, BMNH
7.1.1.191; collected by Fraser.
Type Locality Ecuador; precise locality not
given, but Fraser may have taken the type either
in Pallatanga or in the vicinity of Quito.
Distribution (Fig. 95) Andes of Colombia,
Ecuador, northern Peru and northwestern Vene-
zuela.
Additional Records (compiled from Patter-
son and Albuja, an unpublished manuscript on
shrew-opossums) ECUADOR: Pichincha; Alas-
pungo, Cerro Pichincha (MCZ); Las Maquinas
(AMNH); Santa Rosa, Mindo (Tate, 1923,
AMNH); Rio San Rafael, near Santa Rosa (An-
thony, 1923, AMNH); Santa Rosa (AMNH); An-
tisanilla (Tate, 1923, AMNH; Anthony, 1923,
AMNH); Lloa, Rio Condor Huashana (USNM);
Corazon, forest NW of (Tate, 1923, AMNH;
Anthony, 1923, AMNH); El Chaupi, NW of
(ZFMK); Quebrada Tasinchana (ZFMK); Cerro
Pichincha, Crater (MCZ); Old Santo Domingo
Trail, not located (UMMZ); Canon del Rio Pita
(AMNH); Rio Pita, upper (AMNH); Rio Pita
(MCZ, AMNH); Napo; Papallacta, near (Voss,
1978, AMNH); COLOMBIA: Agua Larga, not lo-
cated (Child, 1895, BMNH); Quindio; La Guneta
(= Laguneta) (AMNH).
Other Additional Records COLOMBIA:
Antioquia; Paramo Frontino (Hershkovitz, 1951,
FMNH); Cauca; Munchique (von Sneidern, 1958,
FMNH); PERU: Piura; Cerro Chinguela, 5 km
NE of Zapalache (Barkley, 1980, LSUMZ), "Ba-
tan," on Zapalache-Carmen Trail (Barkley, 1980,
LSUMZ).
150
FIELDIANA: ZOOLOGY
Caenolestes condorensis I <
500
1000 1
MM ion
\l
j
...._.
__ i
30"
Fig. 93. Distribution of Caenolestes condorensis: found only in Cordillera del Condor in southeastern Ecuador.
= localities from unpublished records; = locality given in published records.
Genus Lestoros Oehser, 1934
Orolestes Thomas, 1917b:3 type species, Orolestes
inca Thomas; generic name preoccupied by Orolestes
MacLachlan, 1895, a genus of dragonfly. Osgood,
1921:11 distribution. Gardner, 1993:25 listed.
Cryptolestes Tate, 1934:154 replacement name for
Orolestes Thomas, preoccupied; type species,
Cryptolestes inca Thomas; generic name preoccu-
pied by Cryptolestes Ganglbauer, a genus of Co-
leoptera. Gardner, 1993:25 listed.
Lestoros Oehser, 1934:240 replacement name for
Cryptolestes Tate, preoccupied; type species, Les-
toros inca Thomas. Gardner, 1993:25 listed.
Type Species Orolestes inca Thomas, by orig-
inal designation.
Distribution Highlands of southern Peru and
northwestern Bolivia.
Lestoros inca (Thomas, 1917)
Orolestes inca Thomas, 1917b:3. Thomas, 1920c:
244 PERU: Cusco (Torontoy; Machu Picchu,
12000-14000 ft [= 3750-4375 m]; Ocobamba
Valley, 9100 ft [= 2840 m]). Gardner, 1983:25
listed, synonomy.
Cryptolestes inca: Tate, 1934:154 type of Crypto-
lestes Tate.
Lestoros inca: Oehser, 1934:240 type of Lestoros
Oehser. Hayman, 1971:194 PERU: Cusco (Ollan-
taytambo). Kirsch and Waller, 1979:390 PERU:
Cusco (Ollantaytambo). Pacheco et al., 1993:4
PERU: Cusco (Tres Cruces; Pillahuata; Quebrada
Morro Legufa; Trocha Ericsson, trail below P.V.
Acjanaco; La Esperanza; Abra Acjanaco). Albuja
and Patterson, 1996:53 PERU: Cusco (Cedro-
bamba Ruins, Machu Picchu; Limacpunco; Pau-
catambo; Quillabamba; Marcapata); Puno (Agu-
alani). Anderson, 1997:167 BOLIVIA: La Paz
(Llamachaque, near Pelechuco).
Holotype Maie, skin and skull, USNM
194401; collected 14 May 1915 by Edmund Heller.
Type Locality Torontoy, Cusco, Peru; 14000
ft (= 4375 m).
Distribution (Fig. 96) Known only from the
temperate zone in Cusco, southeastern Peru, and
La Paz, Bolivia.
Additional Records (compiled from Patter-
son and Albuja, an unpublished monograph on
BROWN: ATLAS OF NEW WORLD MARSUPIALS
151
Fig. 94. Distribution of Caenolestes ccmvelatus: western Colombia and northwestern Ecuador, o
unpublished records; = localities given in published records.
localities from
shrew-opossums) PERU: Cusco; Machu Picchu.
Runcaraccay Ruins (Heller, 1915, USNM); To-
copoqueu, Ocobamba Valley (Heller, 1915,
USNM); Puno; Limbani (Patterson and Albuja).
Other Additional Records PERU: Cusco;
San Luis, km 185 on Ollantaytambo-Quellabamba
Rd. (Thomas, 1974. LSUMZ); Calca, 55 km N
(Patton, 1984, MVZ); Puno; Limbani (Koford.
1951, MVZ; Patton, MVZ).
Genus Rhyncholestes Osgood, 1924
Rhyncholestes Osgood, 1924:170. Gardner.
25 listed.
1993:
Type Species Rhyncholestes raphanurus Os-
good, by original designation.
Distribution Southern Chile and adjacent
Argentina.
Rhyncholestes raphanurus Osgood, 1924
Rhyncholestes raphanurus Osgood. 1924:170. Osgood.
1943:50 CHILE: Los Lagos (Refugio. Volcan
Osorno; boca Rio Inio. Isla de Chiloe [= Isla Chi-
loe]). Pine et al 1979:343 CHILE: Los Lagos
(Quellon. near mouth of Rio Yaldad). Gallardo and
Patterson. 1987:1 1 1 CHILE: Los Lagos (Volcan
Osorno). Patterson and Gallardo. 1987:1 CHILE:
Los Lagos (Maicolpue: Parque Nacional Puyehue;
Valle de La Picada: Refugio. Volcan Osorno: Peulla;
Lago Chapo; Cucao; Puerto Carmen; Rio Inio. Isla
Chiloe: Quellon; Fundo El Venado. Isla Chiloe. not
located). Meserve et al.. 1988:722 CHILE: Los La-
gos (Valle de La Picada. Parque Nacional Vincente
Perez Rosales). Gardner. 1993:25 listed. Bimey et
al.. 1996:1 ARGENTINA: Rio Negro (Puerto
Blest, Parque Nacional Nahuel Huapf). Albuja and
Patterson. 1 996:53 CHILE: Los Lagos (Maicolpue;
Parque Nacional Puyehue; Centro de Ski Antillanca
[= Antillanca Ski Area]; Osomo; Puerto Octay: Val-
le de La Picada. Refugio; Valle de La Picada, La
Picado Alto; Parque Nacional Vincente Perez Ro-
sales. La Picada Forest: Contao; Rio Negro: Rib
Inio); ARGENTINA: Rio Negro (Puerto Blest). Hei-
nonen Fortabat and Chebez. 1997:20 ARGENTI-
NA: Neuquen (Parque Nacional Nahuel Huapf). Ma-
res and Braun. 2000:40 ARGENTINA: Neuquen
(Parque Nacional Nahuel Huapf): Rio Negro (Parque
Nacional Nahuel Huapf; Puerto Blest. Parque Na-
cional Nahuel Huapf).
152
FIELDIANA: ZOOLOGY
Fig. 95. Distribution of Caenolestes fuliginosus: along the Andes from northwestern Venezuela through Colombia,
Ecuador, and northern Peru, o = localities from unpublished records; = localities given in published records.
Holotype Skin and skeleton; FMNH 22422;
collected 12 January 1923 by Wilfred H. Osgood.
Type Locality Mouth of Rio Inio, south end
of Isla de Chiloe [= Isla Chiloe], Chile.
Distribution (Fig. 97) South-central Chile,
including Isla Chiloe and adjacent Argentina.
Additional Records CHILE: Los Lagos Val-
le de La Picada, La Picada Bajo (Patterson and
Albuja, an unpublished monograph, p. 24); Los
Lagos; Lago Chapo, 12.4 km WNW, 19.7 km N
(Meserve, 1986, 1987, FMNH).
III. Order Microbiotheria
Family Microbiotheriidae Ameghino,
1887
Genus Dromiciops Thomas, 1894
Type Species Dromiciops gliroides Thomas
by original designation.
Distribution Chile and adjacent Argentina
from about 36S to near 43S.
Dromiciops gliroides Thomas, 1894
Didelphys elegans Cunningham (not Thylamys ele-
gans Waterhouse, a marmosid), 1871:362.
Didelphys australis F. Philippi (not D. a. Goldfuss,
1 8 1 2, a phalangerid), 1 893a:3 1 8 CHILE: Los La-
gos (type locality near Union, holotype a mounted
skin, skull inside, Museo Nacional, Santiago de
Chile). F. Philippi, 1893b: 33 CHILE: Los Lagos
(near Union). R. A. Philippi, 1894:33 CHILE:
Valdivia [now in Los Lagos]; Llanquihue; Arau-
cania. Wolffsohn and Porter, 1908:69 CHILE:
Araucania (Curacautfn); gliroides Thomas, a syn-
onym.
[Didelphys (Peramys)] australis: Trouessart, 1898:
1244 listed.
Dromiciops australis Thomas, 1919d:212 ARGEN-
TINA: Neuquen (Beatriz, Nahuel Huapf, 800 m);
CHILE: Araucania (Temuco). Osgood, 1943:48
CHILE: Araucania (Villa Portales, Rib Colorado).
Mann, 1955:159 CHILE: Los Lagos (Puerto
Montt; between Valdivia and Puerto Montt, not lo-
cated). Reig et al., 1977:201 CHILE: Los Lagos
(Valdivia). Marshall, 1978c:l, fig. 4 (map distri-
bution). Jimenez and Rageot, 1979:83 CHILE:
Araucania-Biohio (Parque Nacional Nahuelbuta);
Los Lagos Parque Nacional Tolhuaca); Araucania;
BROWN: ATLAS OF NEW WORLD MARSUPIALS
153
Fig. 96. Distribution of Lestoros inca: presently confined to the highlands of southern Peru and La Paz. Bolivia.
c = localities from unpublished records (too close to published localities to be identified separately on map): =
localities given in published records.
(Cerro Nielol: Parque Nacional Villarrica: Los
Parques Nacionales Conguillfo). Marshall. 1982:
35. fig. 12 (map distribution). Pearson and Pearson.
1982M 29 ARGENTINA: Rio Negro (Parque Na-
cional Nahuel Huapf). Meserve et al.. 1988:721
CHILE: Los Lagos (La Picada). Patterson et al..
1989:67 CHILE: Los Lagos (La Picada). Patter-
son et al.. 1990:620 CHILE: Los Lagos (La Pi-
cada). Birney et al.. 1996:153 ARGENTINA: Rio
Negro (Puerto Blest).
Dromiciops australis austral is: Osgood. 1943:48
CHILE: Araucania (Rio Colorado: Victoria: Te-
muco): Los Lagos (Cayetue. Lago Todos Santos
Peulla. Lago Todos Santos: Cerro Nahuelbuta)
Biobio (Lota. SW of Conception): ARGENTINA
Neuquen (Beatriz. Nahuel Huapf). Santos Gollan.
1946:191 ARGENTINA: Neuquen (Huemul. Na-
huel Huapf: Villa Angostura. Nahuel Huapf). Greer.
1965:103 CHILE: Araucania (Angol: Banos Rio
Blanco: Curacautfn: Jauja: Laguna Malleco: Lon-
quimay: Paso Pino Hachado: Rio Ranquil: Selva
Oscura: Troyo): Los Lagos (Cordillera [= Cerro]
de Nahuelbuta): Biobio (Cordillera de los Andes:
Contulmo). Pine et al.. 1979:339 CHILE: Los La-
gos (Lago Paraiso. Parque Nacional Puyehue: An-
tillanca Ski Area [= Centro de Ski]. Parque Na-
cional Puyehue). Tamayo and Frassinetti. 1980:
327 CHILE: Los Lagos (Rio Itata). Pearson.
1 983 :483 ARGENTINA.
D[romiciops] australis australis: Schneider. 1946:
68 CHILE: Biobio (Huepil. N of Rio Laja): Con-
ception (Parque Pedro del Rio).
Dromiciops gliroides Thomas. 1894b: 187 CHILE:
Los Lagos (type locality. Huite. Chiloe Island).
Thomas. 1919d:212 CHILE: Araucania (Temu-
co). Krumbiegel. 1941c: 1 1 CHILE: Valdivia
[now Los Lagos]. Patterson and Rogers, in press:
ms. p. 1 1 ARGENTINA: Neuquen (Lago Lacar:
Parque Nacional Nahuel Huapf): CHILE: Arauca-
nia (Paso Pino Hachado): Biobio (Lota): Los Lagos
(Quellon. near mouth of Rfo Yaldad. Isla Chiloe;
Puerto Carmen. Isla Chiloe: Cucao. Isla Chiloe;
Huite [= Puerto Huite]. Isla Chiloe: Rfo Inio. boca.
Isla Chiloe: Peulla: Seno Reloncavf; Valdivia: Rfo
Itata: Maicolpue: Parque Nacional Puyehue: Valle
de La Picada). Mares and Braun. 2000:41 AR-
GENTINA: Chubut (Parque Nacional Los Alerces:
Parque Nacional Lago Puelo): Neuquen (Beatriz.
Nahuel Huapi: Isla Victoria. Lago Nahuel Huapi:
Parque Nacional Lanfn: Parque Nacional Los Ar-
rayanes: Parque Nacional Nahuel Huapi): Rio Ne-
gro (Pampa Linda. 5 km NW; Parque Nacional Na-
154
FIELDIANA: ZOOLOGY
20
30
40
50
j \
\
j
\ 1
-
s/f
Rhyncholestes raphanurus
500
1000
1500 km
3Lj_
90
80
70
60
50
40"
30
20
10 :j
Fig. 97. Distribution of Rhyncholestes raphanurus: south-central Chile, including Isla Chiloe, and adjacent Ar-
gentina. = localities from unpublished records; = localities given in published records.
huel Huapi; Puerto Blest, Parque Nacional Nahuel
Huapi).
Dromiciops australis gliroides, Osgood, 1943:50
CHILE: Los Lagos (Quellon; Huite). Pine et al.,
1979:340 CHILE: Los Lagos (Quellon, near
mouth of Rio Yaldad, Isla Chiloe). Tamayo and
Frassinetti, 1980:377 CHILE: Los Lagos (Isla
Chiloe) (0-300 m).
Holotype Male, skin and skull, BMNH
92.9.5.3; collected 20 May 1868 by Robert O.
Cunningham on the voyage of H.M.S. Nassau.
Type Locality Huite, near Ancud, northeast-
ern Chiloe Island, Chile.
Distribution (Fig. 98) The rain forests of
central Chile and bordering parts of Argentina
from about 36S to 43S in Chiloe but to 41S on
the mainland, or from the latitude of Concepcion
to the southern border of Isla Chiloe, east to the
lake district of Nahuel Huapi in Neuquen, and Rio
Negro, Argentina. The elevational range extends
from sea level on Isla Chiloe to 1450 m in Mal-
leco Province.
Additional Records CHILE: Araucania;
Thermas de San Luis, 30 km E of Pucon (Creigh-
ton, 1980, UMMZ); Los Lagos; Mafil (Castillo,
1977, MVZ); Catamutun (Solis, 1975, MVZ);
Osorno (Kelt, 1985, FMNH); La Picada Forest,
Parque Nacional Vicente Perez Rosales [= Re-
fugio] (Kelt, 1984, 1986, FMNH); Volcan Osor-
no, La Picada (Gallardo, 1976, MVZ; Sepulveda,
1973, MVZ); Rio Negro (Meserve, 1987,
FMNH); Contao, Palena (Kelt, FMNH); Lago To-
dos los Santos (Hershkovitz, 1996, personal
notes); Yaldad, Cocauque, Isla Chiloe (Meserve
and Lang, 1984, FMNH); ARGENTINA: Neu-
quen; Lago Quillen (Sage, 1982, MVZ); Arroyo
de las Terras Coloradas, Lago Huechulfaquen,
near Cerro del Chico (Sage, 1982, MVZ); Refu-
gio, near Paso Coihue (Patton, 1996, MVZ); Cas-
cada Diana, 16 km WNW of Villa La Angostura
(Pearson, 1985, MVZ); Rio Negro; Refugio Neu-
meyer, SSW of San Carlos de Bariloche (Phillips,
1983, MVZ); Hotel Tunquelen, 25 km E of San
Carlos de Bariloche (Christie, 1982, MVZ); Rio
Castano Overo, W of San Carlos de Bariloche
(Pearson, 1978, 1979, MVZ).
BROWN: ATLAS OF NEW WORLD MARSUPIALS
155
Fig. 98. Distribution of Dromiciops gliroides: south-central Chile and bordering Argentina, west through Isla
Chiloe. o = localities from unpublished records; = localities given in published records.
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FIELDIANA: ZOOLOGY
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BROWN: ATLAS OF NEW WORLD MARSUPIALS
175
Gazetteer
Abra Acjanaco; Cusco, Peru (ca. 1315'S.
7130'W, Pacheco et al., 1993)
Lestoros inca, 3200 m
Abra de Cafias, Parque Nacional Calilegua; Jujuy,
Argentina (2340'S, 6454'W, Flores et al..
2000)
Lutreolina crassicaudata, 1700 m. 1500 m
Thylamys venustus, 1700 m
Abra de La Ventana (see Sierra de la Ventana);
Buenos Aires, Argentina
Abra Pampa; Jujuy. Argentina (2243'S, 6542'W.
Paynter, 1985)
Thylamys pallidior, 3500 m
Abre Campo; Minas Gerais. Brazil (2018'S.
4229'W, Paynter and Traylor. 1991)
Didelphis aurita, 548 m
Acanafia. Rio Cunucunuma; Amazonas, Venezuela
(0313'N, 6558'W, Perez-Hernandez. 1989)
Caluromys lanatus, 145 m
Monodelphis brevicaudata, 145 m
Philander andersoni, 145 m (0339'N. 6566'W)
Aca Poi. 10 km S of Rio Ypane, 1 km N of San
Pedro; Concepcion, Paraguay (2305'S.
5607'W. USBGN)
Metachirus nudicaudatus
Philander opossum (10 km S of Rio Ypane)
Acapulco; Guerrero, Mexico (1651'N, 9955'W.
Geonet)
Marmosa canescens
Acatlan; Oaxaca. Mexico (1832'N. 9637'W,
Geonet)
Didelphis marsupialis
Achamere Wan; Upper Takutu-Upper Essequibo.
Guyana (not located)
Micoureus demerarae
Achotal; Veracruz-Llave, Mexico (1744'N.
9510'W, Geonet)
Marmosa mexicana
Philander opossum
Achupallas; Morona-Santiago. Ecuador (0327'S,
7821'W, Albuja and Patterson, 1996)
Caenolestes eondorensis, 2100 m
Aconquija. Concepcion; Tucuman. Argentina
(2713'S. 6608'W, Mares et al., 1997)
Didelphis albiventris
Lutreolina crassicaudata
Thylamys venustus, 3000 m
Adampada Kreek; Sipaliwini, Suriname (0434'N.
5632'W, Geonet)
Marmosa murine
Agua Blanca (= Aguas Blancas). 24 km NW; Sal-
ta. Argentina (2244'S, 6422'W, Flores et al..
2000)
Lutreolina crassicaudata, ca. 400 m
Thylamys venustus
Agua Buena; Puntarenas, Costa Rica (0845'N,
8256'W, 3 localities within a few km of each
other. Geonet)
Marmosa mexicana
Agua Caliente; Cartago. Costa Rica (1026'N,
8516'W. USBGN)
Micoureus alstoni, 1300 m
Chironectes minimus, 1400 m
Agua Caliente (= Aguas Calientes); Huanuco,
Peru (0850'S. 7441'W, Geonet)
Caluromys lanatus, ca. 150 ft (= 45 m)
Aguacatal; Carabobo. Venezuela (1012'N,
6807'W. Geonet)
Chironectes minimus, 1 100 m
Agua Colorada; Yoro. Honduras (1523'N,
8725'W, Geonet)
Caluromxs derhianus, 3000 ft (= 930 m)
Agua Dulce; Pando. Bolivia (lrOl'S, 6612'W,
Anderson, 1997)
Philander opossum
Agua Dulce; Veracruz-Llave. Mexico (1808'N,
9408'W. Geonet)
Didelphis marsupialis
Agualani; Puno. Peru (1406'S. 6942'W, Geonet)
Lestoros inca, 2840 m. 9200 ft (= 2870 m)
Agua Larga; Colombia (not located)
Caenolestes fuliginosus
Agua Negra; Yaracuy. Venezuela (1004'N,
6909'W. Perez-Hernandez. 1989)
Didelphis marsupialis, 80 m
Marmosa robinsoni, 80 m
Monodelphis palliolata, 80 m
Aguaray: Salta, Argentina (2216'S, 6345'W,
Paynter, 1985)
Didelphis albiventris, 700 m
Thylamys venustus, 565 m
Aguaruna Village. Rio Cenepa. near Huampami
(= Kayamas. Aguaruna Village = Tseasim);
Amazonas. Peru (not located)
Caluromys lanatus
Didelphis marsupialis
Philander andersoni
Aguas Blancas (see Agua Blanca); Salta, Argentina
Aguas Caliente (see Agua Caliente); Huanuco,
Peru
Aguas Calientes, 1 km below Shintuya; Madre de
Dios. Peru (Shintuya = 1241'S. 7115'W, Geo-
net)
Caluromys lanatus
Philander opossum, 450 m (left bank Rio Ma-
dre de Dios)
Aguascalientes; Aguascalientes. Mexico (2153'N,
10212'W. Geonet)
176
FIELD1ANA: ZOOLOGY
Didelphis marsupialis
Aguas Chiquitas; Tucuman, Argentina (2637'S,
6512'W, Mares et al., 1996)
Didelphis albiventris
Lutreolina crassicaudata
Aguas Claras, near San Adolfo, Rio Suaza; Huila,
Colombia (ca. 0137'N, 7559'W, Paynter,
1997)
Didelphis albiventris, 1500 m
"Aguas Claras" Camp; Madre de Dios, Peru
(1257'S, 6854'W, Emmons et al., 1994)
Didelphis marsupialis, ca. 190 m
Marmosops noctivagus, ca. 190 m
Aguas Negras (see Parque Nacional Calilegua);
Jujuy, Argentina
Aguas Negras, Parque Nacional Itatiaia (see Pico
das Agulhas Negras); Rio de Janeiro, Brazil
Agua Viva; Trujillo, Venezuela (0933'N,
7038'W, Geonet)
Metachirus nudicaudatus, 164 m
Philander opossum, 164 m
Aguirre; Carabobo, Venezuela (1014'N,
6814'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Agiiita, Cerro Duida; Amazonas, Venezuela (ca.
0320'N, 6532'W, Paynter, 1982)
Didelphis albiventris, 3250 ft (= 1000 m)
Aipua (= Ayapua), W side Rio Purus; Amazonas,
Brazil (0427'S, 6208'W, Paynter and Tray-
lor, 1991)
Micoureus demerarae
Alajuela; Alajuela, Costa Rica (1001'N, 8413'W,
Geonet)
Didelphis marsupialis
Alamor; Loja, Ecuador (0402'S, 8002'W, Payn-
ter, 1993)
Didelphis marsupialis, 1325 m
Marmosops impavidus, 1325 m
Alaspungo, Cerro Pichincha; Pichincha, Ecuador
(0000\ 7836'W, Paynter, 1993)
Caenolestes fuliginosus
Albergue Bosque de las Nubes, at Puente Union;
Cusco, Peru (ca. 1309'S, 7117'W, Pacheco
et al., 1993)
Marmosops noctivagus, 1700 m
Albergue Cusco (Cusco) Amazonico, Puerto Mal-
donado; Madre de Dios, Peru (Puerto Mal-
donado = 1236'S, 6911'W, Paynter, 1983)
Caluromys lanatus (12 km E of Puerto Mal-
donado)
Metachirus nudicaudatus, 200 m (12 km E of
Puerto Maldonado)
Albina; Marowijne, Suriname (0530'N, 5403'W,
Stephens and Tray lor, 1985)
Monodelphis brevicaudata
Philander opossum
Albrook; Panama, Panama (0858'N, 7934'W,
Geonet)
Caluromys derbianus
Philander opossum
Albrook Field; Panama, Panama (0859'N,
7933'W, Geonet)
Philander opossum (2 mi N of Albrook Field)
Alcoche; La Paz, Bolivia (1542'S, 6740'W,
Paynter, 1992)
Metachirus nudicaudatus
Micoureus regina, 424 m
Monodelphis brevicaudata, 424 m
Alem Paraiba; Minas Gerais, Brazil (2152'S,
4241'W, Paynter and Traylor, 1991)
Didelphis aurita, 140 m (2145'S, 4232'W)
Marmosops incanus, 140 m (left bank Rio Pa-
raiba)
Philander frenata
Alfarcito, 15 km NE of Maimara, 9.7 km E of
Tilcara; Jujuy, Argentina (2335'S, 6521'W,
Geonet)
Thylamys pallidior, 2600 m
Alfenas; Minas Gerais, Brazil (2126'S, 4557'W,
Paynter and Traylor, 1991)
Didelphis aurita, 843 m
Alguacil; Caja Seca, Zulia, Venezuela (0909'N,
7104'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
Alluriquin (see Puente de Ayuriquin); Pichincha,
Ecuador
Almirante; Bocas del Toro, Panama (0918'N,
8224'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Philander opossum
Almoxarifado, Parque Nacional do Itatiaia; Rio de
Janeiro-Minas Gerais, Brazil (Parque
2220'S, 4435'W, Geonet)
Caluromys philander
Aloag, 11 km W; Pichincha, Ecuador (0028'S,
7835'W, Paynter, 1993)
Caenolestes convelatus, 2960 m
Alrededores de Caicara del Orinoco, Orinoco Me-
dio; Bolivar, Venezuela (737'N, 6510'W,
Perez-Hernandez, 1989)
Micoureus demerarae, 50 m
Alta Florida, Caracas; Distrito Federal, Venezuela
(Caracas = 1030'N, 6655'W, Perez-Hernan-
dez, 1989)
Didelphis marsupialis, ca. 980 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
177
Altamira, right bank Rio Jurua: Amazonas
(0635'S, 6854'W. Patton et al., 2000)
Caluromys lanatus
Didelphis marsupialis
Marmosa murina
Marmosops neblina
Metachirus nudicaudatus
Micoureus demerarae
Philander mcilhennyi
Altamira. opposite, left bank Rio Jurua: Amazon-
as. Brazil (0635'S. 6856'W, Patton et al..
2000)
Caluromys lanatus
Marmosops neblina
Micoureus regina
Altamira: Barinas. Venezuela (0850'N. 7031'W,
Paynter. 1982)
Didelphis marsupialis, 794 m
Marmosa robinsoni, 611 m. 697 m
Metachirus nudicaudatus
Monodelphis palliolata, 697-794 m
Altamira, 35 km above Manu: Madre de Dios.
Peru (1213'S. 7055'W. Stephens and Tray-
lor. 1983)
Didelphis marsupialis, 400 m
Micoureus regina
Altamira; Para. Brazil (0312'S. 5212'W. Paynter
and Traylor. 1991)
Caluromys philander (left bank Rio Xingu)
Didelphis marsupialis left bank Rio Xingu
Marmosops parvidens (52 km SSW. E bank Rio
Xingu)
Metachirus nudicaudatus (W bank Rio Xingu)
Micoureus demerarae (52 km SSW. E bank Rio
Xingu)
Philander opossum (E and W bank Rio Xingu)
Alto Beni: Beni. Bolivia (not located)
Philander opossum
Alto Bonito: Antioquia, Colombia (ca. 0705'N.
7630'W, Paynter. 1997)
Philander opossum. 1500 ft (= 460 m) (upper
Rio Sucio, W slope Cordillera Occidental)
Alto da Serra: Sao Paulo. Brazil (2347'S,
4619'W. Paynter and Traylor, 1991)
Philander frenata, 800 m. 1375 m
Alto de Galapagos: Valle del Cauca-Choco. Co-
lombia (0453'N. 7613'W. Patterson and Al-
buja. 1996)
Caenolestes convelatus
Alto No Leon (see No Leon); Distrito Federal.
Venezuela
Alto Parana, southeastern Mato Grosso do Sul/
Parana; Brazil (2305'S. 5222'W. Paynter
and Traylor. 1991)
Didelphis albiventris
Alto Parana: Sao Paulo. Brazil (not located)
Didelphis albiventris
Alto Parnahyba (see Alto Rio Parnaiba); Maran-
ahao. Brazil
Alto Pencoso: San Luis, Argentina (3326'S,
6656'W. Geonet)
Thylamys pallidior
Alto Rio Anchicaya (see Rio Anchicaya. alto);
Valle del Cauca, Colombia
Alto Rio Atacavi, Campamento 2; Amazonas, Ve-
nezuela (0305'N. 6654'W, Perez-Hernan-
dez. 1989)
Didelphis albiventris, 140 m
Alto Rio Bermejo: Salta, Argentina (2652'S,
5823'W. Paynter. 1985)
Lutreolina crassicaudata
Alto Rio Madidi. Rio Itenez; La Paz, Bolivia
(1353'S. 6846'W. Anderson, 1997)
Caluromys lanatus
Mannosops noctivagus
Alto Rio Parnaiba (= Alto Parnahyba); Marana-
hao. Brazil (0646'S, 4300'W, Hershkovitz,
1992)
Gracilinanus agilis, 400-600 m
Alto Rio Urucu (= Rio Urucu); Amazonas, Brazil
(0451'S. 6516'W. Hershkovitz, 1997)
Metachirus nudicaudatus
Micoureus demerarae
Philander mcilhennyi
Altos. 20 km N; La Cordillera. Paraguay
(2514'S. 5714'W. USBGN)
Didelphis albiventris
Altos de Pipe. Instituto Venezolano de Investiga-
ciones Cientificas (IVIC): Miranda. Venezue-
la (1021'N. 6702'W. Perez-Hernandez,
1989)
Didelphis marsupialis
Mannosops fuscatus
Altura de Yuto; Jujuy. Argentina (Yuto =
2338'S. 6428'W. Paynter, 1985)
Micoureus constantiae, 500 m
Alvarenga (see Fazenda da Floresta); Minas Ger-
ais. Brazil
Alvaro Obregon (= Frontera): Tabasco, Mexico
(1835'N. 9239'W. Geonet)
Didelphis marsupialis, 10 ft (= 3 m)
Philander opossum, 3 m
Amaicha del Valle. near: Tucuman. Argentina
(2636'S. 6555'W. Flores et al., 2000)
Thylamys pallidior
Amarim Igarape (see Igarape Amorin): Para. Bra-
zil
178
FIELDIANA: ZOOLOGY
Amaybamba; Cusco, Peru (1304'S, 7208'W, Pat-
ton, pers. comm.)
Marmosops impavidus, 2000 m, 2200 m (2 km
NE and 3 km E of Amaybamba, Lucumayo
valley)
Marmosops noctivagus
Amazonica (formerly Putumayo); southern Co-
lombia on Eucador-Peru border; Colombia
(no specific locality)
Glironia venusta
Ambo; Huanuco, Peru (1007'S, 7610'W, Geo-
net)
Didelphis albiventris, 6700 ft (= 2090 m)
Americana; Sao Paulo, Brazil (2244'S, 4720'W,
Geonet)
Didelphis albiventris
Amolac; Puebla, Mexico (1803'N, 9823'W,
Geonet)
Marmosa canescens
Anandabaru (= Anundabaru); Potaro-Suparuni,
Guyana (Anandabaru Mountains = 0459'N,
5944'W, Geonet)
Monodelphis brevicaudata
Ananindeua, E.E Braganca; Para, Brazil (0122'S,
4823'W, Paynter and Traylor, 1991)
Philander opossum
Anapolis (= Annapolis); Goias, Brazil (1620'S,
4858'W, Paynter and Traylor, 1991)
Didelphis albiventris, 1000 m, 1010 m
Philander frenata, 1000 m
Ancon; Panama, Panama (0857'N, 7933'W,
Geonet)
Didelphis marsupialis
Ancon; Guayas, Ecuador (0220'S, 8052'W,
Paynter, 1993)
Didelphis marsupialis, m
Andalgala; Catamarca, Argentina (2736'S,
6620'W, Mares et al., 1997)
Didelphis albiventris
Thylamys pallidior
Andalgala, Highway E of, at junctions of High-
way 62 and Highway 1; Catamarca, Argen-
tina (2736'S, 6615'W, Flores et al., 2000)
Thylamys pallidior
Andalgala, N of, Rio Andalgala; Catamarca, Ar-
gentina (2734'S, 6616'W, Flores et al.,
2000)
Thylamys pallidior
Andalucia; Huila, Colombia (0154'N, 7540'W,
Geonet)
Didelphis marsupialis, 600-990 m, 7000 ft ( =
2180 m)
Andarahy (see Macaco Secco); Bahia, Brazil
Angaba (see Cuiaba); Mato Grosso, Brazil
Angol; Araucania, Chile (3748'S, 7243'W, Geo-
net)
Dromiciops gliroides, 537 m, 685 m, 1110 m,
1 170 m (N and W of Angol)
Thylamys elegans, 72 m
Angosto del Rio Pescado; Salta, Argentina
(2233'S, 6432'W, Flores et al., 2000)
Thylamys venustus
Angostura; San Jose, Costa Rica (0917'N,
8341'W, Geonet)
Metachirus nudicaudatus
Angra dos Reis; Rio de Janeiro, Brazil (2300'S,
4418'W, Paynter and Traylor, 1991)
Lutreolina crassicaudata, 2 m
Marmosops incanus, sea level
Metachirus nudicaudatus
Micoureus demerarae, 2 m
Annandale; Grenada (1205'N, 6143'W, Geonet)
Marmosa robinsoni
Annapolis (see Anapolis); Goias, Brazil
Anundabaru (see Anandabaru); Potaro-Siparuni,
Guyana
Antillanca Ski Area ('= Centro de Ski), 98 km E
of Osorno, Parque Nacional Puyehue; Los
Lagos, Chile (4045'S, 7209'W, Albuja and
Patterson, 1996, Geonet)
Dromiciops gliroides, 780 m, 970 m
Rhyncholestes raphanurus, 690-820 m
Antisanilla; Pichincha, Ecuador (0023'S,
7817'W, Paynter, 1993)
Caenolestes fuliginosus, 11500 ft (= 3590 m)
Anzoategui; Lara, Venezuela (0936'N, 6954'W,
Paynter, 1982)
Marmosa robinsoni, ca. 1500 m
Apayacu; Loreto, Peru (0319'S, 7207'W, Ste-
phens and Traylor, 1983)
Philander opossum
Apazote; Campeche, Mexico (1854'N, 9020'W,
Geonet)
Didelphis marsupialis
Apia, Rio Apia, Rio Cauca Valley; Risaralda, Co-
lombia (0505'N, 7556'W, Geonet)
Caluromys derbianus, 5000 ft (= 1560 m)
Apolo; La Paz, Bolivia (1446'S, 6827'W, Payn-
ter, 1992)
Chironectes minimus, 1436 m
Micoureus regina
Arabopo (= Arabupii), 8 mi SE of Cerro Roraima;
Bolivar, Venezuela (0506'N, 6044'W, Payn-
ter, 1982)
Didelphis albiventris, 4200 ft (= 1300 m)
Marmosa murina
Micoureus demerarae, 4200 ft (= 1300 m)
Monodelphis brevicaudata
BROWN: ATLAS OF NEW WORLD MARSUPIALS
179
Arabupu (see Arabopo); Bolivar, Venezuela
Aracruz, Santa Cruz; Espfrito Santo, Brazil
(1956'S, 4009'W, Paynter and Traylor,
1991)
Metachirus nudicaudatus
Aracruz Forestal, Aracruz; Espirito Santo, Brazil
(1947'S, 4007'W, Mustrangi and Patton,
1997)
Marmosops incanus, 50 m
Aragarcas; Goias, Brazil (1555'S, 5215'W,
Paynter and Traylor, 1991)
Philander frenata (gallery forest)
Araguari; Minas Gerais, Brazil (1838'S,
4808'W, Paynter and Traylor, 1991)
Didelphis albiventris, 929 m
Araponga; Minas Gerais, Brazil (2040'S,
4232'W, Geonet)
Didelphis aurita, 1060-1212 m
Araraquara; Sao Paulo, Brazil (2147'S, 4810'W,
Paynter and Traylor, 1991)
Caluromys lanatus
Ararangua; Santa Catarina, Brazil, (2856'S,
4929'W, Paynter and Traylor, 1991)
Gracilinanus microtarsus, 13 m
Metachirus nudicaudatus
Monodelphis iheringi
Arataye (see Arataye Crique); French Guiana
Arataye Crique (= Arataye Riviere = Riviere Ar-
ataye); French Guiana (0359'N, 5234'W,
USBGN)
Marmosa murina
Marmosops parvidens (right bank)
Metachirus nudicaudatus
Monodelphis brevicaudata
Arataye Riviere, French Guiana (see Arataye
Crique); French Guiana
Arboledas, W of Ciicuta; Norte de Santander, Co-
lombia (0740'N, 7245'W, Geonet)
Caluromys lanatus
Arenal; Jujuy, Argentina (2420'S, 6421'W,
Paynter, 1985)
Didelphis albiventris, 1000 m
Arequipa; Arequipa, Peru (1623'S, 7132'W,
Geonet)
Thylamys pallidior, 10600 ft (= 3300 m) (18
km E of Arequipa)
Arima; Arima, Trinidad and Tobago (1038'N,
6117'W, Geonet)
Marmosa robinsoni (Arima Valley)
Marmosops fuscatus
Aripo Valley; St. George, Trinidad and Tobago
(Aripo River = 1035'N, 6114'W, Geonet)
Marmosa robinsoni
Aripuana (= Nucleo Pionefro Humboldt); Mato
Grosso, Brazil (0910'S, 6038'W, Paynter
and Traylor, 1991)
Didelphis marsupialis
Marmosops noctivagus, 200 m (right bank, trib-
utary of Rio Madeira)
Metachirus nudicaudatus
Ariruma, 7 km SE; Santa Cruz, Bolivia (1824'S,
6419'W, Anderson, 1997)
Thylamys venustus
Aristobulo de Valle, 19 km W on Rio Cunapirus;
Misiones, Argentina (2707'S, 5455'W,
Paynter, 1985)
Didelphis albiventris
Aritagua Urucutuca; Bahia, Brazil (1443'S,
3906'W, Geonet)
Micoureus demerarae
Armacao do Sul; Santa Catarina, Brazil, (2721'S,
4834'W, USBGN)
Lutreolina crassicaudata
Armila; Quebrada Venado; San Bias, Panama
(0840'N, 7728'W, Geonet)
Caluromys derbianus
Chironectes minimus
Didelphis marsupialis
Metachirus nudicaudatus
Arraijan; Panama, Panama (0857'N, 7939'W,
Geonet)
Caluromys derbianus
Arroyo Bopicua (see Bopicua); Rio Negro, Uru-
guay
Arroyo Brusquitas, General Pueyrredon; Buenos
Aires, Argentina (3815'S, 5746'W, Geonet)
Didelphis albiventris
Lutreolina crassicaudata
Monodelphis dimidiata
Arroyo Chapadmalal, 28 km SW of Mar del Plata;
Buenos Aires, Argentina (3803'S, 5742'W,
USBGN)
Didelphis albiventris
Arroyo Chocori; Buenos Aires, Argentina
(3824'S, 5807'W, USBGN)
Monodelphis dimidiata
Arroyo Corrientes; Buenos Aires, Argentina (ca.
3810'S, 5715'W, Reig, 1964)
Lutreolina crassicaudata
Arroyo de la Mina (see Paso Duraznero); Rio
Grande do Sul, Brazil
Arroyo de las Terras Coloradas, Lago Huechul-
faquen, near Cerro del Chico; Neuquen, Ar-
gentina (Lago Huechulfaquen = 3946'S,
7128'W, USBGN)
Dromiciops gliroides, 950 m
Arroyo de Los Noquez, Parque Nacional El Rey;
180
FIELDIANA: ZOOLOGY
Salta, Argentina (Los Noquez = 2444'S,
6433'W, USBGN)
Thylamys venustus
Arroyo El Pescado; Buenos Aires, Argentina
(3454'S, 5747'W, USBGN)
Monodelphis dimidiata
Arroyo Garupa; Misiones, Argentina (2727'S,
5548'W, USBGN)
Chironectes minimus
Arroyo La Horqueta; Jujuy, Argentina (2407'S,
6527'W, Flores et al., 2000)
Thylamys venustus, 2100 m
Arroyo La Matanza; Buenos Aires, Argentina (not
located, may be Rio Matanza at 3442'S,
5828'W, Paynter, 1985)
Lutreolina crassicaudata
Arroyo Las Brujas; Canelones, Uruguay (ca.
3438'S, 5620'W, Paynter, 1994)
Lutreolina crassicaudata
Arroyo Nacurutu (= Zarate, Arroyo Nacurutu);
Buenos Aires, Argentina (3404'S, 5902'W,
Geonet)
Gracilinanus agilis
Arroyo Negro, 15 km S of Paysandu; Rio Negro,
Uruguay (Paysandu = 3219'S, 5808'W,
Paynter, 1994)
Didelphis albiventris, 48 m
Arroyo Pereyra; Buenos Aires, Argentina
(3451'S, 5808'W, USBGN)
Lutreolina crassicaudata
Arroyo Piray Guazii; Misiones, Argentina (Puerto
Piray = 2627'S, 5442'W, USBGN)
Chironectes minimus
Arroyo Pirayii-i (= Pirayii-hi); Itapua, Paraguay
(2651'S, 5514'W, Geonet)
Chironectes minimus
Arroyo Polanco, 52 km N of Minas; Lavalleja,
Uruguay (3353'S, 5503'W, Geonet)
Lutreolina crassicaudata
Arroyo Santelmita (see Parque Nacional Baritii);
Salta, Argentina
Arroyo Tacuara, Cerro Azul; Misiones, Argentina
(Cerro Azul = 2738'S, 5529'W, Paynter,
1985)
Chironectes minimus, ca. 200 m
Arroyo Yuto; Jujuy, Argentina (2340'S, 6437'W,
Flores et al., 2000)
Thylamys venustus
Arroyo Vina (brazo del Arroyo Garupa); Mi-
siones, Argentina (Arroyo Garupa = 2727'S,
5548'W, Geonet)
Didelphis albiventris
Micoureus demerarae
Arroyo Zaiman; Misiones, Argentina (2727'S,
5554'W, Geonet)
Monodelphis dimidiata
Arruda, 4 km NE of San Joaquin; Beni, Bolivia
(San Joaquin = 1304'S, 6449'W, Hershko-
vitz, 1997)
Philander opossum
Arumateua (= Arumatheua); Para, Brazil
(0354'S, 4941'W, Hershkovitz, 1997)
Philander opossum (left bank Rio Tocantins)
Arumatheua (see Arumateua); Para, Brazil
Aruma Valley; Trinidad and Tobago (not located,
possibly Arima, stream at 1035'N, 6116'W,
Geonet)
Caluromys philander
Didelphis marsupialis
Ascencion, 6 km W; Santa Cruz, Bolivia (1543'S,
6309'W, Anderson, 1997)
Marmosops dorothea
Philander opossum
Ascencion de Guarayos; Santa Cruz, Bolivia
(1457'S, 6124'W, Hershkovitz, 1997)
Philander opossum
Aserradero del Infernillo, 1 1 km W of Gomez
Farias; Tamaulipas, Mexico (Gomez Farias =
2303'N, 9909'W, Geonet)
Marmosa mexicana, 1300 m
Aserradero Moisa, 7 km E; Santa Cruz, Bolivia
(1433'S, 6108'W, Anderson, 1997)
Philander opossum
Aserradero Monte de San Pablo; Beni, Bolivia
(1305'S, 6453'W, Anderson, 1997)
Didelphis marsupialis
Astillero; Alta Verapaz, Guatemala (1546'N,
8934'W, Geonet)
Philander opossum, 25 ft (= 7 m)
Astillero; La Paz, Bolivia (1616'S, 6733'W, An-
derson, 1997)
Didelphis marsupialis
Astillero; Santa Rosa, Guatemala (1351'N,
9021'W, Geonet)
Didelphis marsupialis, 25 ft (= 7 m)
Asuncion; Central, Paraguay (2516'S, 5740'W,
Paynter, 1989)
Caluromys lanatus
Chironectes minimus, 11 m
Didelphis albiventris, 200 m
Lutreolina crassicaudata
Philander opossum
Thylamys pusilla, 11 m
Auara Igarape; Amazonas, Brazil (0422'S,
5843'W, Hershkovitz, 1997)
Didelphis marsupialis
Philander opossum
BROWN: ATLAS OF NEW WORLD MARSUPIALS
181
Auca, on road to Nanegal; Pichincha, Ecuador
(Nanegal = 0007'S, 7846'W, Paynter, 1993)
Marmosa robinsoni
Aurare (see Rio Anaure); Zulia, Venezuela
Auyan-tepui (= Cerro Auyan-tepui); Bolivar, Ve-
nezuela (0555'N, 6232'W, Paynter, 1982)
Didelphis albiventris, 1 100 m, 1850 m, 2500 m
Didelphis marsupialis
Marmosa tyleriana, 1850 m
Marmosops pinheroi, 460 m, 1100 m
Metachirus nudicaudatus, 1500-3500 ft ( =
460-1000 m)
Micoureus demerarae, 460 m, 1 100 m, 1850 m
Monodelphis brevicaudata, 2560 m (see also
Rancho Grande, Auyan-tepui)
Avanavero Falls (see Avanavero Vallen); Sipali-
wini, Suriname
Avanavero Vallen (= Avanavero Falls), Kabalebo
River; Sipaliwini, Suriname (0449'N.
5724'W, Geonet)
Mondelphis brevicaudata
Philander opossum
Avanhandava, Lajeado: Sao Paulo, Brazil
(2057'S, 4846'W, Hershkovitz. 1997)
Philander frenata
Avispero, Valle de Suaza. near Rio Suaza: Huila.
Colombia (Rio Suaza = 0210'N. 7441'W,
USBGN, 1964)
Didelphis marsupialis
Axtla; San Luis Potosi, Mexico (2128'N,
9851'W, Geonet)
Didelphis marsupialis
Ayacucho; Ayacucho, Peru (1309'S, 7413'W,
Geonet)
Didelphis albiventris, 2746 m
Ayacucho, Ibanez; Santa Cruz, Bolivia (1751'S.
6320'W, Hershkovitz, 1997)
Marmosops dorothea
Philander opossum, 500 m
Ayapua (see Aipua); Amazonas, Brazil
Ayolas; Misiones, Paraguay (2724'S, 5654'W,
Paynter, 1989)
Didelphis albiventris (5 km ENE of Ayolas)
Lutreolina crassicaudata
Azul, 4 km NE; Buenos Aires, Argentina
(3647'S, 5951'W, Paynter, 1985)
Didelphis albiventris, 137 m
Azul Benson Farm; Buenos Aires, Argentina (not
located)
Didelphis albiventris
Azunta (see Casa Morava); Beni, Bolivia
B
Bacabena, 23 km SW of Belem (see Belem); Para,
Brazil
Bacaetava (see Ipanema); Sao Paulo, Brazil
Baeza; Napo, Ecuador (0027'S, 7753'W, Payn-
ter, 1993)
Marmosa murina, 5000 ft (= 1560 m) (below
Baeza)
Marmosa robinsoni, 1525 m
Marmosops impavidus, 1525 m
Bagado. Rio Andagueda; Chocd, Colombia
(0525'N, 7624'W, Paynter, 1997)
Chironectes minimus
Didelphis marsupialis, 650 ft (= 200 m)
Philander opossum
Bagua (= Bagua Chica); Amazonas, Peru
(0538'S, 7831'W, Geonet)
Caluromys lanatus, 1350 ft (= 420 m)
Bagua Chica (see Bagua); Amazonas, Peru
Bahia (= Salvador); Bahia, Brazil (1259'S,
3831'W, Paynter and Traylor, 1991)
Metachirus nudicaudatus
Philander frenata
Bahia de Asuncion; Central, Paraguay (2516'S,
5737'W, Geonet)
Chironectes minimus
Lutreolina crassicaudata, 11 m
Philander opossum
Bahia de Caraquez. Rio Briseno [Briceno]; Man-
abi, Ecuador (0036'S, 8025'W, Paynter,
1993)
Didelphis marsupialis, sea level
Marmosa robinsoni, sea level (on Rio Briceno)
Bahia de Cata; Aragua, Venezuela (ca. 1028'N,
6744'W, Paynter, 1982)
Didelphis marsupialis, sea level
Bahia de los Casara; Beni, Bolivia (1313'S,
6221'W. Anderson, 1997)
Didelphis marsupialis
Bahfa de Patanemo; Carabobo, Venezuela
(1026'N, 6800'W, Perez-Hernandez. 1989)
Didelphis marsupialis, 10 m
Monodelphis palliolata, 10 m
Bahia de Todos Santos (see Reconcavo); Bahia,
Brazil
Bahia Honda (= Honda Bay); La Guajira, Colom-
bia (1221'N, 7147'W, Paynter, 1997)
Didelphis marsupialis
Baia de Guanabara (see Guanabara); Rio de Ja-
neiro. Brazil
Baiao. Rio Tocantins; Para. Brazil (0241'S,
4941'W, Paynter and Traylor, 1991)
Didelphis marsupialis, 25 m
Metachirus nudicaudatus
Philander opossum (right bank Rio Tocantins)
Baking Pot; Cayo, Belize (1712'N, 8901'W,
Geonet)
182
FIELDIANA: ZOOLOGY
Caluromys derbianus
Balancan; Tabasco, Mexico (1748'N, 9132'W,
Geonet)
Caluromys derbianus
Philander frenata
Balboa; Panama, Panama (0857'N, 7934'W,
Geonet)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
Balcarce (= "Hill Plot"); Buenos Aires, Argen-
tina (3750'S, 5815'W, Paynter, 1985)
Monodelphis dimidiata, 100 m, 108 m
Balisa; Mato Grosso, Brazil (1615'S, 5226'W,
Hershkovitz, 1997)
Philander opossum (gallery forest)
Baliza Caviu, 12.5 km SSW of Mar del Plata (see
Mar del Plata [Cuidad]); Buenos Aires, Ar-
gentina
Baliza San Andres; Buenos Aires, Argentina (ca.
3815'S, 5735'W, Reig, 1964)
Monodelphis dimidiata
Balta, Rio Curanja; Ucayali, Peru (1006'S,
7114'W, Geonet)
Caluromys lanatus, 300 m
Chironectes minimus, 300 m
Didelphis marsupialis, 300 m
Marmosa murina, 300 m
Metachirus nudicaudatus, 300 m
Philander mcilhennyi, 300 m
Philander opossum, 300 m
Balzapamba; Bolivar, Ecuador (0147'S, 7913'W,
Paynter, 1993)
Caluromys derbianus, 700 m
Balzar; Guayas, Ecuador (0122'S, 7954'W,
Paynter, 1993)
Marmosa robinsoni, 100 m
Balzar Mountains (see Montanas de Balzar);
Manabi, Ecuador
Bambito; Chiriqui, Panama (0850'N, 8237'W,
Geonet)
Didelphis marsupialis
Bambui; Minas Gerais, Brazil (2001'S, 4558'W,
Geonet)
Didelphis albiventris
Bafiada Tropa Vieja, ca. 35 km NE of Montevi-
deo; Canelones, Uruguay (3447'S, 5552'W,
Paynter, 1994)
Lutreolina crassicaudata
Bafiado de Carrasco; Canelones, Uruguay
(3450'S, 5603'W, Paynter, 1994)
Didelphis albiventris, 100 m
Lutreolina crassicaudata, 100 m
Banado de San Jorge, Parque Pereyra Iraola;
Buenos Aires, Argentina (Parque Pereyra Ir-
aola = 3451'S, 5804'W, Paynter, 1985)
Lutreolina crassicaudata, ca. 20 m
Banagena; Sao Paulo, Brazil (not located)
Philander opossum
Banco da Vitoria, Pirataquisse, Ilheus; Bahia, Bra-
zil (Ilheus = 1449'S, 3902'W, Paynter and
Traylor, 1991)
Micoureus demerarae, sea level
Banda Oriente (= Uruguay); Uruguay (no specific
locality)
Didelphis albiventris
Banos, Oriente; Tunguarahua, Ecuador (0124'S,
7824'W, Paynter, 1993)
Didelphis albiventris, 4000 ft (= 1250 m) (Pal-
mera, E of Banos), 5800 ft (= 1800 m)
Banos Rio Blanco; Araucania, Chile (3834'S,
7134'W, Geonet)
Dromiciops gliroides, 980 m
Baramita; Barima-Waina, Guyana (0721'N,
6029'W, Geonet)
Didelphis marsupialis
Barao de Cocais; Minas Gerais, Brazil (1956'S,
4326'W, Geonet)
Didelphis aurita
Barbacoas; Narino, Colombia (0141'N, 7809'W,
Paynter, 1997)
Chironectes minimus
Didelphis marsupialis, 36 m
Barcarena, 23 km SW of Belem; Para, Brazil
(0130'S, 4840'W, Paynter and Traylor,
1991)
Chironectes minimus
Barcena; Jujuy, Argentina (2357'S, 6530'W,
Flores et al., 2000)
Thylamys pallidior
Thylamys venustus
Barinitas, Rio Barragan; Barinas, Venezuela
(0840'N, 7025'W, Perez-Hernandez, 1989)
Marmosa robinsoni, 400 m
Metachirus nudicaudatus, 440 m
Monodelphis palliolata, 440 m (also at 3 km
SW of Barinitas)
Barra (see Manaus); Amazonas, Brazil
Barra de Marica; Rio de Janeiro, Brazil (2256'S,
4250'W, Paynter and Traylor, 1991)
Didelphis aurita
Metachirus nudicaudatus
Ph i lander frenata
Barra de Santiago; Ahuachapan, El Salvador
(1341'N, 9000'W, Geonet)
Didelphis marsupialis
Philander opossum
Barra do Corda, Rio Mearim; Maranhao, Brazil
BROWN: ATLAS OF NEW WORLD MARSUPIALS
183
(0530'S, 4215'W, Paynter and Traylor,
1991)
Didelphis marsupialis
Barra do Garcas; Mato Grosso, Brazil (1553'S,
5215'W, Hershkovitz, 1997)
Philander opossum (gallery forest)
Barra do Rio Negro (see Manaus); Amazonas,
Brazil
Barra do Rio Paraopeba; Minas Gerais, Brazil
(1850'S, 4511'W, Geonet)
Caluromys lanatus
Barra Longa; Minas Gerais, Brazil (2017'S,
4303'W, Geonet)
Didelphis aurita
Barrancas del Rio Primero, left bank (lower Pla-
tense); Cordoba, Argentina (Ciudad Rio Pri-
mero = 3120'S, 6337'W, Paynter, 1985)
Didelphis alhiventris
Barranquilla; Atlantico, Colombia (1059'N,
7448'W, Geonet)
Marmosa robinsoni, ca. 100 m
Barranquita, 10 km SW of San Joaquin; Beni, Bo-
livia (1313'S, 6448'W, Anderson, 1997)
Didelphis marsupialis
Philander opossum
Barreira, Serra dos Orgaos; Rio de Janeiro, Brazil
(2230'S, 4256'W, Paynter and Traylor,
1991)
Metachirus nudicaudatus
Philander frenata (2256'S, 4256'W)
Barro Alto Rio Sao Francisco; Minas Gerais, Bra-
zil (not located)
Monodelphis domestica
Barro Blanco; Antioquia, Colombia (ca. 0615'N,
7523'W, Paynter, 1997)
Didelphis marsupialis, 2160 m
Barro Branco; Rio de Janeiro, Brazil (2223'S,
4430'W, Hershkovitz, 1997)
Philander frenata
Barro Colorado (= Isla Barro Colorado = Barro
Colorado Island); Colon/Panama, Panama
(0919'N, 7950'-7956'W, Geonet)
Caluromys derbianus
Chironectes minimus
Didelphis marsupialis
Marmosa robinsoni
Metachirus nudicaudatus
Philander opossum
Barro Colorado Island (see Barro Colorado); Co-
lon/Panama, Panama
Barro Vermelho, left bank Rio Jurua; Amazonas,
Brazil (0628'S, 6846'W, Mustrangi and Pat-
ton, 1997)
Caluromys lanatus
Didelphis marsupialis
Marmosa murina
Marmosops impavidus
Marmosops parvidens
Metachirus nudicaudatus
Micoureus demerarae
Micoureus regina
Bartica Grove; Cuyuni-Mazaruni, Guyana
(0624'N, 5837'W, Geonet and Times Atlas,
1992)
Marmosa murina
Micoureus demerarae
Monodelphis brevicaudata
Baruta; Miranda, Venezuela (1022'N, 6645'W,
Perez-Hernandez, 1989)
Caluromys philander, ca. 1 000 m
Didelphis marsupialis
Marmosa robinsoni
Monodelphis palliolata
Base de Carmo (see Fazenda Intervales); Sao Pau-
lo, Brazil
Base Camp (see Cerro de La Neblina); Amazonas,
Venezuela
Base Camp La Neblina (see Campamento); Ama-
zonas, Venezuela
Basilio (= Cordillera Basilio); Santa Cruz, Boliv-
ia (1808'S, 6312'W, Anderson, 1997)
Monodelphis domestica
Philander opossum
"Batan," on Zapalache-Carmen Trail; Piura, Peru
(Zapalache = 0509'S, 7927'W, Stephens
and Traylor, 1983)
Caenolestes caniventer, 2250 m
Caenolestes fuliginosus, 2250 m
Didelphis marsupialis, 2250 m
Bauru; Sao Paulo, Brazil (2219'S, 4904'W,
Paynter and Traylor, 1991)
Lutreolina crassicaudata, 499 m
Metachirus nudicaudatus
Micoureus demerarae, 499 m
Monodelphis americana
Beatriz, Nahuel Huapf (also see Parque Nacional
Nahuel Huapi); Neuquen, Argentina
(4051'S, 7139'W, USBGN)
Dromiciops gliroides, 800 m
Bejuquero, N of Zea; Merida, Venezuela
(0823'N, 7147'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 500 m
Metachirus nudicaudatus, 500 m
Monodelphis palliolata, 500 m (15 km N of
Tovar)
Philander opossum, 500 m
Bela Vista de Minas; Minas Gerais, Brazil
(1952'S, 4305'W, Varejao and Valle, 1982)
184
FIELDIANA: ZOOLOGY
Didelphis aurita
Belem (= Belem de Para = Instituto Agronomico
do Norte = Ipeau-Apez = Murucutu = Para
= Utinga); Para, Brazil (0127'S, 4829'W,
Paynter and Traylor, 1991)
Caluromys philander, 10 m
Chironectes minimus, 10 m (Bacabena, 23 km
SW of Belem)
Didelphis marsupialis, 10 m
Gracilinanus emiliae, m
Marmosa murina, 10 m
Marmosa parvidens
Marmosops pinheiroi, 1 m
Metachirus nudicaudatus
Micoureus demerarae, 10 m
Monodelphis americana, 10 m (near Belem)
Philander opossum (Sapucajuba, suburb of Be-
lem; Marco, suburb of Belem; Utinga, suburb
of Belem)
Belem de Para (see Belem); Para, Brazil
Belem Virus Laboratory (see Macapa); Amapa,
Brazil and Para, Brazil
Belen; Catamarca, Argentina (2739'S, 6702'W,
Mares et al., 1997)
Thylamys pallidior
Belen; Tarapaca, Chile (1829'S, 6931'W, Geo-
net)
Thylamys elegans, 10500 ft (= 3280 m)
Belen, Rio Cunucunuma; Amazonas, Venezuela
(0343'-0337'N, 6553'-6546'W, Handley,
1976)
Caluromys lanatus, 150 m
Caluromys philander, 150 m
Didelphis marsupialis, 150 m
Marmosa murina, 150 m
Micoureus demerarae, 150 m
Philander andersoni, 150 m
Belen del Dragon, 6 km NNW; Artigas, Uruguay
(Artigas = 3024'S, 5628'W, Paynter, 1994)
Didelphis albiventris
Belgica, Valle de Suaza; Huila, Colombia (Rio
Suaza = 0302'N, 7441'W, USBGN)
Didelphis marsupialis
Belgrano; Buenos Aires, Argentina (3434'S,
5828'W, Paynter, 1985)
Lutreolina crassicaudata, sea level
Belize City; Belize (1729'N, 8811'W, Geonet)
Philander opossum
Bella Imperatriz (see Parintins); Amazonas, Brazil
Bella Vista; Amazonas, Peru (0530'S, 7840'W,
Hershkovitz, 1992)
Gracilinanus agilis, 300 m (right bank Rio Ma-
rafion)
Bella Vista; Pando, Bolivia (1123'S, 6712'W,
Anderson, 1997)
Philander opossum
Bellavista, upper Rio Porce, Cordillera Central;
Antioquia, Colombia (0633'N, 7518'W,
Paynter, 1997)
Philander opossum, 1 200 m
Belo Horizonte; Minas Gerais, Brazil (1955'S,
4356'W, Paynter and Traylor, 1991)
Didelphis albiventris, 835 m
Bemberg (see Puerto Libertad); Misiones, Argen-
tina
Bemfica (see Benfica); Rio de Janeiro, Brazil
Benavides (see Benevides); Para, Brazil
Benevides ( = Benavides); Para, Brazil (0122'S,
4815'W, Paynter and Traylor, 1991)
Monodelphis americana, 30 m
Benfica (= Bemfica); Rio de Janeiro, Brazil
(2225'S, 4438'W, Paynter and Traylor,
1991)
Philander frenata, 2000 ft (= 625 m) (Serra de
Itatiaia) (Hershkovitz [1997] used Holt's co-
ordinates of 2141'S, 4326'W, in Minas Ger-
ais, but Paynter and Traylor [1991] place this
locality in Rio de Janeiro)
Berg En Dal [Bergendal]; Brokopondo, Suriname
(0509'N, 5504'W, Geonet)
Marmosa murina
Bermalcue; Chaco, Paraguay (not located)
Thylamys pusilla
Bermejo (= Vermejo); Santa Cruz, Bolivia (ca.
1810'S, 6336'W, Paynter, 1992)
Thylamys venustus, 400 m
Bestion; Azuay, Ecuador (0330'S, 7913'W,
Hershkovitz files)
Didelphis albiventris, 3075 m
Betania; Tachira, Venezuela (0727'N, 7226'W,
Perez-Hernandez, 1989)
Didelphis albiventris, 2250 m
Gracilinanus dryas, 2250 m
Marmosops impavidus, 2250 m
Better Hope; West Pomeroon-Supenaam Coast,
Guyana (0723'N, 5831'W, Geonet)
Didelphis marsupialis
Lutreolina crassicaudata
Philander opossum
Biagi Santi (see Bigisanti); Marowijne, Suriname
Bigi Poika; Saramacca, Suriname (0525'N,
5530'W, Geonet)
Didelphis marsupialis
Philander opossum
Bigisanti (= Biagi Santi), Wia-Wia Nature Re-
serve; Marowijne, Suriname (ca. 0555'N,
5430'W, Stephens and Traylor, 1985)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
185
Didelphis marsupialis
Bijagua (= Vijagua); Boaco, Nicaragua (1235'N,
8513'W, USBGN)
Marmosa mexicana
Biological Reserve at Horco Molle (see Horco
Molle); Tucuman. Argentina
Biological Station "Pithecia"; Loreto, Peru
(0510'S, 7430'W, Hershkovitz. 1983)
Caluromys lanatus, 150 m
Didelphis marsupialis, 150 m
Birongo (see Rio Birongo); Miranda. Venezuela
Bisira; Bocas del Toro, Panama (0854'N.
8152'W, Geonet)
Caluromys derbianus
Bitagron. right bank Coppename River: Sipali-
wini. Suriname (0510'N, 5606'W, Geonet)
Marmosa murina
Bledos; San Luis Potosi, Mexico (2151'N,
10107'W, Geonet)
Didelphis marsupialis
Bluefields; Zelaya, Nicaragua (12WN. 8345'W,
Geonet)
Marmosa mexicana
Mieoureus alstoni, ca. sea level (1220'N.
8350'W)
Philander opossum (1220'N. 8350'W)
Blumenau; Santa Catarina, Brazil, (2656'S.
4903'W. Paynter and Traylor. 1991)
Chironectes minimus, 14 m (near Blumenau)
Gracilinanus microtarsus, 14 m. 22 m
Boaco; Boaco, Nicaragua (1228'N. 8540'W.
Geonet)
Caluromys derbianus (near Santa Rosa)
Didelphis marsupialis, 300 m
Boa Esperanca. right bank Rio Jurua; Amazonas.
Brazil (0632'S, 6855'W. Patton et al 2000)
Didelphis marsupialis
Mieoureus demerarae
Mieoureus regina
Boa Esperanca. Serra de Caparao; Minas Gerais,
Brazil (2010'S, 4146'W. Hershkovitz. 1997)
Philander frenata, 3000 ft (= 930 m)
Boa Vista, Pocao, right bank Rio Branco: Rorai-
ma. Brazil (0249'N, 6040'W, Paynter and
Traylor, 1991)
Didelphis marsupialis, 55 m
Philander opossum
Boa Vista. Rio Ipiranga: Sao Paulo. Brazil
(2422'S, 4738'W, Paynter and Traylor,
1991)
Gracilinanus microtarsus (2435'S, 4738'W)
Metachirus nudicaudatus
Boa Vista, Teresopolis (see Teresopolis); Rio de
Janeiro. Brazil
Bobos; Izabal. Guatemala (1521'N, 8849'W,
Geonet)
Philander opossum
Boca Colorado: Madre de Dios. Peru (1230'S,
7040'W. Hershkovitz, 1997)
Philander opossum, 279 m
Boca de Lagarto Cocha, Rio Aguarico at conflu-
ence of Rio Napo: Napo. Ecuador (0039'S.
7516'W, Paynter, 1993)
Glironia venusta, ca. 300 m
Boca del Drago: Bocas del Toro. Panama
(0925'N. 8220'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Philander opossum
Boca del Rio; Veracruz-Llave, Mexico (1906'N,
9606' W, Geonet)
Didelphis marsupialis
Philander opossum, 10 ft (= 3 m) (E coast)
Boca del Rio Tacuari; Trienta y Tres. Uruguay
(3246'S. 5318'W. Paynter. 1994)
Didelphis albiventris
Boca de Rio Paya: Darien. Panama (0755'N,
7731'W. Handley. 1966. Gardner. 1973)
Chironectes minimus, 2000 ft (= 625 m)
Didelphis marsupialis
Boca de Tigre Valley. 5 km NW of Caracas; Dis-
trito Federal. Venezuela (1032'N, 6654'W,
Handley. 1976)
Didelphis marsupialis, 1394 m
Boca de Yaracuy; Yaracuy. Venezuela (1035'N,
6815'W, Handley. 1976)
Didelphis marsupialis, 2 m
Monodelphis palliolata, 2 m
Boca Mavaca. 84 km SSE of Esmeralda; Ama-
zonas. Venezuela (0230'N. 6513'W, Han-
dley. 1976)
Caluromys lanatus, 138 m
Caluromys philander, 138 m
Didelphis albiventris, 138 m
Didelphis marsupialis, 138 m
Marmosa murina, 138 m
Monodelphis brevicaudata, 138 m
Bocatoma. near Lo Valdes; Santiago, Chile
(3355'S, 7013'W, Geonet)
Thylamys elegans
Bocatorito, Isla San Cristobal; Bocas del Toro.
Panama (0914'N, 8216'W, Geonet)
Caluromys derbianus
Mieoureus alstoni
Bodoco; Pernambuco. Brazil (0747'S, 3955'W.
USBGN)
Monodelphis domestica (Si'tio Belem. Si'tio
Paus Preto. Si'tio Sao Goncalo)
186
FIELDIANA: ZOOLOGY
Bogava (see Bugaba); Chiriqui, Panama
Bogota (= Santafe de Bogota); Cundinamarca,
Colombia (ca. 0436'N, 7405'W, Paynter,
1997)
Caenolestes fuliginosus
Didelphis albivenths, 2590 m, 2600 m, 2620
m, 2700 m
Didelphis marsupialis, 2620 m
Marmosa robinsoni, 2590 m
Marmosops fuscatus, 2590 m
Marmosops impavidus
Metachirus nudicaudatus
Micoureus alstoni, 2590 m
Micoureus regina, 2590 m
Bohio Peninsula; Colon, Panama (0912'N,
7950'W, Geonet)
Metachirus nudicaudatus
Philander opossum
Boim; Para, Brazil (0249'S, 5510'W, Paynter
and Traylor, 1991)
Monodelphis emiliae, ca. 50 m
Bom Conselho; Pernambuco, Brazil (0910'S,
3641'W, Geonet)
Didelphis albiventris
Bomfim (see Villa Nova); Bahia, Brazil
Bom Jardin; Brazil (not located, many so named
in many states)
Philander frenata
Bonanza; Zelaya, Nicaragua (1401'N, 8435'W,
Geonet)
Didelphis marsupialis
Philander opossum
Bonasika, Essequibo River; Essequibo Islands-
West Demerara, Guyana (0645'N, 5830'W,
Geonet)
Marmosa murina
Bonda; Magdalena, Colombia (1 114'N, 7408'W,
Paynter, 1997)
Didelphis marsupialis, ca. 50 m
Marmosa robinsoni, ca. 50 m, 150-250 ft ( =
47-78 m)
Bonfinopolis de Minas; Minas Gerais, Brazil
(1632'S, 4600'W, Varejao-Valle, 1982)
Didelphis albiventris
Bonsucesso, Rio de Janeiro, Brazil (2212'S,
4244'W, Mustrangi and Patton, 1997)
Marmosops paulensis, 1000 m
Bopicua, Rio Uruguay; Rib Negro, Uruguay (ca.
3306'S, 5801'W, Paynter, 1994)
Gracilinanus agilis, ca. 10 m (also Arroyo
Bopicua)
Boqueron; Chiriqui, Panama (083<Q'N, 8235'W,
Geonet)
Caluromys derbianus
Didelphis marsupialis
Marmosa mexicana
Boqueron de San Francisco; Cundinamarca, Co-
lombia (San Francisco = 0458'N, 7418'W,
Paynter, 1997)
Didelphis albiventris
Gracilinanus dryas, 2250 m, 3000 m
Boquete; Chiriqui, Panama (0847'N, 8226'W,
Geonet)
Didelphis marsupialis
Marmosa robinsoni, 4000 ft (= 1250 m)
Boraceia (see also Estacao Biologica de Bora-
ceia); Sao Paulo, Brazil (ca. 2239'S,
4554'W, Paynter and Traylor, 1991)
Chironectes minimus, 850 m
Didelphis aurita, ca. 850 m
Metachirus nudicaudatus
Philander frenata (upper Rio Tiete, 2210'S,
4845'W)
Borba; Amazonas, Brazil (0424'S, 5935'W,
Paynter and Traylor, 1991)
Caluromys philander (right bank Rio Madeira)
Didelphis marsupialis, 25 m
Boroica; Beni, Bolivia (1313'S, 6409'W, Ander-
son, 1997)
Philander opossum
Boruca; Puntarenas, Costa Rica (0900'N,
8319'W, Geonet)
Caluromys derbianus
Marmosa mexicana, 1700 ft (= 530 m)
Botanic Garden; Port-of-Spain, Trinidad and To-
bago (1039'N, 6131'W, Geonet)
Caluromys philander
Marmosa robinsoni
Botuvera; Santa Catarina, Brazil (2713'S,
4905'W, Geonet)
Lutreolina crassicaudata
Boundary Camp, Itabu Creek Head; Upper Tak-
utu-Upper Esequibo, Guyana (ca. 0133'N,
5810'W, Stephens and Traylor, 1985, and
Times Atlas, 1992)
Metachirus nudicaudatus
Boyuibe, 53 km E; Santa Cruz, Bolivia (2027'S,
6250'W, Anderson, 1997)
Micoureus constantiae
Monodelphis domestica
Thylamys pusilla
Braganca; Para, Brazil (0103'S, 4646'W, Paynter
and Traylor, 1991)
Caluromys philander
Metachirus nudicaudatus
Brasilia; Distrito Federal, Brazil (1547'S,
4755'W, Paynter and Traylor, 1991)
Didelphis albiventris, ca. 1 100 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
187
Philander frenata (gallery forest)
Brasilia, 20 km S; Distrito Federal. Brazil
(1556'S, 4753'W, Mares et al., 1989)
Gracilinanus agilis, 15-25 km S
Monodelphis kunsi
Brasilia, Fundacao Zoologica Botanico; Distrito
Federal, Brazil (1547'S, 4755'W, Paynter
andTraylor, 1991)
Gracilinanus agilis, 1100 m
Brazo Casiquiare (= Casiquiare Canal, Cerro Dui-
da); Amazonas, Venezuela (0237'N,
6619'W, Handley, 1976)
Micoureus demerarae, up to 6200 ft (= 1930
m)
Monodelphis brevicaudata
Brazo Largo, Delta del Parana, near: Entre Ribs.
Argentina (3347'S, 5836'W, Paynter, 1985)
Gracilinanus agilis, sea level
Brickfield; Caroni. Trinidad and Tobago (1029'N,
6128'W, Geonet)
Didelphis marsupialis
Brokopondo: Brokopondo, Suriname (0504'N,
5458'W. Geonet)
Monodelphis brevicaudata
Brownsberg [Browns Berg]; Brokopondo, Suri-
name (0453'N, 5512'W, Geonet)
Monodelphis brevicaudata, 475 m
Brownsberg [Browns Berg] Nature Park; Broko-
pondo, Suriname (0455'N, 5512'W, Ste-
phens and Traylor, 1985)
Didelphis albiventris
Didelphis marsupialis
Bruja Point; Panama, Panama (0853'N, 7935'W,
Geonet)
Philander opossum
Brumado; Minas Gerais, Brazil (2107'S.
4418'W, Hershkovitz, 1997)
Gracilinanus agilis
Brusque: Santa Catarina, Brazil (2706'S.
4856'W, Paynter and Traylor, 1991)
Chironectes minimus, 21 m
Bucay. Rio Chimbo; Guayas, Ecuador (0210'S.
7906'W, Paynter, 1993)
Didelphis marsupialis, 298 m
Philander opossum, 1000 ft (= 300 m)
Buckhall; Essequibo Islands-West Demerara,
Guyana (0656'N, 5833'W, Geonet)
Monodelphis brevicaudata
Philander opossum, 9 m (Essequibo River)
Buena Hora; Beni, Bolivia (1134'S; 6512'W,
Anderson. 1997)
Philander opossum
Buenaventura; Valle del Cauca, Colombia
(0353'N, 7704'W, Paynter, 1997)
Caluromys derbianus, sea level
Didelphis marsupialis, 10 m, 150 m
Metachirus nudicaudatus (6 km N of Buena-
ventura)
Philander opossum, near sea level; 150 m (28
km NE of Buenaventura)
Buena Vista, near Bogota; Cundinamarca, Colom-
bia (ca. 0420'N, 7340'W)
Metachirus nudicaudatus
Buena Vista; Meta, Colombia (0410'N, 7341'W
Paynter. 1997)
Metachirus nudicaudatus, 1500 m
Buena Vista [Buenavista]; Santa Cruz, Bolivia
(1727'S, 6340'W, Paynter, 1992)
Caluromys lanatus, 400 m
Didelphis marsupialis, 125 m, ca. 400 m (Bue-
na Vista, Ichilo)
Gracilinanus agilis, ca. 400 m
Marmosa lepida, ca. 400 m
Marmosops dorothea, 400 m, 500 m (Buem
Vista. Ichilo)
Micoureus constantiae, 400 m (W bank Ric
Yapacani)
Philander opossum
Thylamys venustus
Buena Vista [Buenavista], near Paramo de Tama
41 km SW of San Cristobal; Tachira, Vene-
zuela (0727'N. 7226'W. Handley, 1976)
Caenolestes fuliginosus, 2390-2400 m
Didelphis albiventris, 1700 m, 2380-2390 m
Gracilinanus dryas, 2405-2410 m (0754'N
7159'W, Hershkovitz, 1997)
Marmosops impavidus, 2350-2430 m (near Pa-
ramo de Tama)
Buena Vista, near; Santa Cruz, Bolivia (1724'S
6344'W, Anderson et al., 1993)
Didelphis marsupialis
Buena Vista Peninsula, near Frijoles; Colon, Pan-
ama (Frijoles = 0910'N, 7948'W, Geonet)
Metachirus nudicaudatus
Philander opossum (1.75 km NNW of Frijoles'
Buenos Aires; Buenos Aires, Argentina (3436'S
5827'W, Paynter, 1985)
Didelphis albiventris, sea level
Buenos Aires Zoological Gardens; Buenos Aires
Argentina (3436'S, 5827'W, Paynter, 1985)
Lutreolina crassicaudata
Buerarema (see Buerarema Ribeirao da Fortuna):
Bahia. Brazil
Buerarama da Fortuna (see Buerarema Ribeirac
da Fortuna); Bahia, Brazil
Buerarema Ribeirao da Fortuna (= Buerarama da
Fortuna = Buerarema); Bahia, Brazil
(1457'S, 3919'W, USBGN)
188
FIELDIANA: ZOOLOGY
Micoureus demerarae, sea level
Buga; Valle del Cauca, Colombia (0354'N,
7617'W, Paynter, 1997)
Didelphis marsupialis, 965 m
Bugaba (= Bogava); Chiriquf, Panama (0829'N,
8237'W, Geonet)
Caluromys derbianus, 2500 m
Marmosa mexicana
Philander opossum
Burruyacii; Tucuman, Argentina (2630'S,
6445'W, Paynter, 1985)
Didelphis albiventris, 800 m
Thylamys venustus, 1300 m
Bush Bush, Nariva Swamp; Nariva, Trinidad and
Tobago (Nariva Swamp = 1025'N, 6104'W,
USBGN)
Didelphis marsupialis
Butanta; Sao Paulo, Brazil (ca. 2334'S, 4643'W,
Hershkovitz, 1997)
Philander frenata
Buxton, 1 mi E; Demerara-Mahaica, Guyana
(0647'N, 5802'W, Geonet)
Didelphis marsupialis
Caazapa; Caazapa, Paraguay (2609'S, 5624'W,
Paynter, 1989)
Caluromys lanatus
Cabassou; French Guiana (0454'N, 5218'W,
USBGN)
Caluromys philander
Didelphis marsupialis
Philander opossum
Cabecera del Cano Culebra, Cerro Duida, 40 km
NNW of Esmeraldas; Amazonas, Venezuela
(0330'N, 6543'W, Handley, 1976)
Didelphis albiventris, 1400 m
Cabo Frio; Rio de Janeiro, Brazil (2253'S,
4201'W, Paynter and Tray lor, 1991)
Didelphis albiventris, sea level
Cabo Tres Puntas; Santa Cruz, Argentina
(4706'S, 6533'W, Paynter, 1985)
Lestodelphys halli
Cacao; French Guiana (0435'N, 5228'W, Ste-
phens and Tray lor, 1985)
Monodelphis brevicaudata
Cacao Pereira (see Cacau Pirera); Amazonas, Bra-
zil
Cacau Pirera (= Cacao Pereira); Amazonas, Bra-
zil (ca. 0308'S, 6005'W, Paynter and Tray-
lor, 1991)
Didelphis marsupialis, ca. 20 m
Marmosops noctivagus (Igarape Cacao Pereira)
Caceres; Antioquia, Colombia (0735'N, 7520'W,
Paynter, 1997)
Caluromys derbianus, 75 m
Caceres (= Sao Luis de Caceres); Mato Grosso,
Brazil (1604'S, 57 41'W, Paynter and Tray-
lor, 1991)
Caluromys lanatus, 152 m
Didelphis albiventris, 1 52 m
Didelphis marsupialis
Philander opossum
Cachabi (= Cachavf), Rio Cachabi; Esmeraldas,
Ecuador (0103'N, 7850'W, Hershkovitz,
1987)
Philander opossum
Cachagua; Valparaiso, Chile (3235'S, 7125'W,
Geonet)
Thylamys elegans, sea level
Cachavf (see Cachabi); Esmeraldas, Ecuador
Cacheuta; Mendoza, Argentina (3306'S,
6907'W, Geonet)
Thylamys pallidior
Cachi, 30 km E; Salta, Argentina (2506'S,
6611'W, Paynter, 1985)
Thylamys pallidior
Cachoeira Bonita, Parque Nacional do Caparao;
Minas Gerais, Brazil (Parque = 2031'-
2040'S, 41 o 40'-4142'W, Paynter and Tray-
lor, 1991)
Philander frenata
Cachoeira de Pancada, Rio Maraca, Mazagao (see
Prosperidade); Amapa, Brazil
Cachoeira do Campo (= Glaura); Minas Gerais,
Brazil (2020'S, 4340'W, Paynter and Tray-
lor, 1991)
Didelphis albiventris, ca. 1 1 00 m
Didelphis aurita, ca. 1 100 m
Cachoeira do Rio Grande, Rib Mamore; Rondon-
ia, Brazil (not located, border of Bolivia and
Brazil)
Monodelphis glirina
Cachoeira Porteira; Para, Brazil (0105'S,
5704'W, Paynter and Traylor, 1991)
Monodelphis brevicaudata
Cacuri, Alto Ventuari; Amazonas, Venezuela
(0449'N, 65 21'W, Perez-Hernandez, 1989)
Caluromys lanatus
Metachirus nudicaudatus
Caete, Cidade (see Cidade Caete); Minas Gerais,
Brazil
Caete, Ro^as Novas; Minas Gerais, Brazil
(1945'S, 4337'W, Varejao-Valle, 1982)
Didelphis albiventris, ca. 1 1 00 m
Cafayate (= Cayafate); Salta, Argentina (2605'S,
6558'W, Paynter, 1992)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
189
Didelphis albiventris, 1600 m
Cafayate, 45 km S; Tucuman, Argentina (2620'S,
6558'W, Mares et al., 1996)
Thylamys venustus
Cafeces, Montegrande, Ibanez; Santa Cruz, Boliv-
ia (Ibanez = ca. 1748'S, 6308'W, Paynter,
1992)
Monodelphis domestica
Cafetal de Chama (= Rio Chama [= Hama]); Me-
rida, Venezuela (ca. 0832'N, 7121'W, Payn-
ter, 1982)
GracUinanus marica, 1600 m
Micoureus demerarae, 1630 m
Cafetal de Llano; Merida, Venezuela (not located)
Micoureus demerarae
Cafetal de Merida (see Cafetales de Merida); Me-
rida, Venezuela
Cafetal de Milla (= Cafetos de Milla); Merida,
Venezuela (0830'N, 7122'W, Hershkovitz,
1992)
GracUinanus dryas
GracUinanus marica, 1600 m
Marmosa robinsoni, 1680 m
Marmosops fuscatus, 1630 m
Micoureus demerarae
Cafetales de Merida (= Cafetal de Merida = Caf-
etos de Merida); Merida, Venezuela
(0836'N, 7108'W, Hershkovitz, 1992)
GracUinanus dryas, 1640 m
GracUinanus marica, 1600-1630 m
Micoureus demerarae, 1630 m
Cafetal Fuente; Merida, Venezuela (not located)
Micoureus demerarae
Cafetos de Merida (see Cafetales de Merida); Me-
rida, Venezuela
Cafetos de Milla (see Cafetal de Milla); Merida,
Venezuela
Cahabon (see Cajabon); Alta Verapaz, Guatemala
Caicara; Bolivar, Venezuela (0737'N, 6610'W,
Paynter, 1982)
Didelphis marsupialis
Micoureus demerarae, 46 m (Caicara del Ori-
noco)
Monodelphis brevicaudata
Caicara; Mato Grosso, Brazil (ca. 1604'S,
5742'W, Paynter and Traylor, 1991)
Caluromys lanatus, ca. 150 m
Caicara (see Caicara de Maturin); Monagas, Ve-
nezuela
Caicara de Maturin (= Caicara); Monagas, Ve-
nezuela (0949'N, 6336'W, Paynter, 1982)
Didelphis marsupialis, 175 m
Caicedo; Antioquia, Colombia (0625'N,
7600'W, Paynter, 1997)
Caenolestes convelatus, 360 m
Caimancito, Parque Nacional Calilegua; Jujuy,
Argentina (2344'S, 6435'W, Paynter, 1985)
Thylamys venustus, 400 m, 550 m, 600 m, 700
m, 6000 ft (= 1875 m)
Micoureus constantiae
Caixa D'Agua; Espirito Santo, Brazil (2038'S,
4055'W, Hershkovitz, 1997)
Monodelphis scalops, 750 m
Philander frenata
Caiza; Tarija, Bolivia (2149'S, 6334'W, Ander-
son, 1997)
Thylamys venustus
Cajabon (= Cahabon), 40 mi NE of Coban; Alta
Verapaz, Guatemala (1534'N, 8949'W,
Geonet)
Marmosa mexicana, 220 m
Cajamarca; Cajamarca, Peru (0710'S, 7831'W,
Geonet)
Didelphis albiventris, 2800 m
Cajanuma, Laguna Compadre; Loja, Ecuador
(0410'S, 7907'W, Patterson and Albuja;
Paynter, 1993, lists Cajanuma, Nudo de, at
0405'S, 7912'W).
Caenolestes caniventer, 2900-3200 m
Calca, 55 km N; Cusco, Peru (1320'S, 7157'W,
Stephens and Traylor, 1983)
Lestoros inca, 3560 m
Calarca; Quindio, Colombia (0431'N, 7538'W,
Paynter, 1997)
Micoureus alstoni, ca. 1500 m
Calcoene; Amapa, Brazil (0230'N, 5057'W,
Paynter, 1997)
Didelphis marsupialis
Caldas (see Dagua); Valle del Cauca, Colombia
Caldeirao (see Ilha de Marajo); Para, Brazil
Calera (see La Calera); Valparaiso, Chile
Caleufu; La Pampa, Argentina (3535'S, 6433'W,
Paynter, 1985)
Monodelphis dimidiata, 260 m
Cali (= Rio Bariatou [Barratow] = Rio Cauca);
Valle del Cauca, Colombia (0327'N,
7631'W, Paynter, 1997)
Caluromys derbianus, 1 000 m (Rio Cauca Val-
ley)
Chironectes minimus, 1000 m
Didelphis marsupialis, 957-1050 m
Philander opossum (upper Rio Cauca Valley)
Calilegua; Jujuy, Argentina (2347'S, 6447'W,
Paynter, 1985)
Didelphis albiventris
Micoureus constantiae
Thylamys venustus, ca. 500 m
190
FIELDIANA: ZOOLOGY
Calilegua Mountain (see Serrania de Calilegua);
Jujuy, Argentina
Callao; Mendoza, Argentina (not located)
Thylamys pallidior
Camargo; Chusquisaca, Bolivia (2039'S,
6513'W, Paynter, 1992)
Thylamys pallidior, 3400 m
Camataqui (= Camataquin); Tarija, Bolivia
(2119'S, 6513'W, Anderson, 1997)
Thylamys pallidior, 11500 ft (= 3590 m)
Thylamys venustus
Camataquin (see Camataqui); Tarija, Bolivia
Camatindi; Tarija, Bolivia (2100'S, 6323'W, An-
derson, 1997)
Monodelphis domestica
Thylamys venustus
Camboriu; Santa Catarina, Brazil, (2701'S,
4838'W, Paynter and Traylor, 1991)
Chironectes minimus, 9 m
Cameta; Para, Brazil (0215'S, 4930'W, Paynter
and Traylor, 1991)
Caluromys philander, 25 m (left bank Rio To-
cantins)
Chironectes minimus, 25 m (left bank Rio To-
cantins)
Didelphis marsupialis, 25 m (Rio Tocantins)
Micoureus demerarae, 25 m (left bank Rio To-
cantins)
Monodelphis brevicaudata, 25 m (left bank Rio
Tocantins)
Monodelphis emiliae (W bank Rio Tocantins,
0214'S, 4930'W)
Philander opossum (left bank Rio Tocantins,
0215'S, 4929'W)
Camiaco, ca. 12 km NNW of Limoqueje; Beni,
Bolivia (1524'S, 6446'W, Herskovitz, 1997)
Didelphis marsupialis
Philander opossum
Camino del Indio, 20 km NE of Castillos; Rocha,
Uruguay (Castillos = 3412'S, 5350'W,
Paynter, 1994)
Lutreolina crassicaudata
Camino de los Espanoles, San Esteban; Carabobo,
Venezuela (San Esteban = 1026'N, 6800'W,
Paynter, 1982)
Didelphis marsupialis, 365 m
Monodelphis palliolata, 365 m
Camino El Palmar, 7 km SW of Santa Cruz; Santa
Cruz, Bolivia (1748'S, 6308'W, Anderson
et al., 1993)
Marmosops dorothea
Camino Vilches, 1 mi E of San Joaquin (see San
Joaquin); Beni, Bolivia
Camiri, near; Santa Cruz, Bolivia (2005'S,
6334'W, Anderson, 1997)
Monodelphis domestica
Camp I, Cerro de La Neblina; Amazonas, Vene-
zeula (0046'N, 6600'W, Geonet)
Micoureus demerarae
Philander opossum
Camp II, 2.5 km NE of Pico Phelps, Cerro de La
Neblina; Amazonas, Venezuela (0046'N,
6600'W, Geonet)
Didelphis albiventris, 2085-2100 m
Marmosops neblina
Metachirus nudicaudatus
Camp V, 2 km NNW of Pico Phelps, Cerro de La
Neblina; Amazonas, Venezuela (0046'N,
6600'W, Geonet)
Micoureus demararae
Camp VI, Cerro de La Neblina; Amazonas, Ve-
nezuela (0046'N, 6600'W, Gardner, 1988)
Marmosops neblina
Camp VII, Cerro de La Neblina; Amazonas, Ve-
nezuela (0046'N, 6600'W, Geonet)
Didelphis albiventris, 1730 m, 2100 m, 3045 m
(2.5 km NE of Pico Phelps [ = Cerro de La
Neblina])
Marmosops neblina, 1800 m
Metachirus nudicaudatus
Micoureus demerarae, 1800 m
Camp XI, Cerro de La Neblina; Amazonas, Ve-
nezuela (0045'N, 6558'W, Gardner, 1988)
Micoureus demerarae
Marmosops neblina
Campamento, Repressa del Guri (= Zona de la
Repressa del Guri), Rio Caroni; Bolivar, Ve-
nezuela (0718'N, 6300'W, Perez-Hernan-
dez, 1989)
Micoureus demerarae, 180 m
Marmosa murina, 180 m
Monodelphis brevicaudata, 180 in
Campamento; Bolfvar, Venezuela (0629'N,
6126'W, Perez-Hernandez, 1989)
Metachirus nudicaudatus, 100 m
Campamento (= Base Camp La Neblina), Cerro
de La Neblina; Amazonas, Venezuela
(0052'N, 6614'W, Perez-Hernandez, 1989)
Caluromys lanatus
Chironectes minimus
Didelphis marsupialis
Metachirus nudicaudatus
Micoureus demerarae
Philander andersoni
Philander opossum
Campamento No. 2, Duida; Amazonas, Venezuela
(0310'N, 6532'W, Perez-Hernandez, 1989)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
Micoureus demerarae, 1000 m
Campamento 6 de Agosto; Beni, Bolivia
(1517'S, 6785'W, Anderson, 1997)
Chironectes minimus
Campamento El Merey, 45 km E of Temblador;
Monagas, Venezuela (0850'N, 6242'W, Lo-
pez-Fuster et al., 2000)
Marmosa robinsoni
Campamento El Trapiche; Beni, Bolivia (1448'S,
6619'W, Anderson, 1997)
Didelphis marsupialis
Campamento M.A.R.N.R., Rio Guarapiere; Mon-
agas, Venezuela (0955'N, 6255'W, Lopez-
Fuster et al., 2000)
Marmosa robinsoni
Campamento Moira, Alto Madidi; La Paz, Bolivia
(1335'S, 6846'W, Anderson, 1997)
Micoureus constantiae
Campamento Yuqui; Cochabamba, Bolivia
(1647'S, 6457'W, Anderson, 1997)
Didelphis marsupialis
Campana; Buenos Aires, Argentina (3513'S,
5854'W, Hershkovitz, 1992)
Gracilinanus agilis
Campanha; Minas Gerais, Brazil (2150'S.
4524'W, Paynter and Traylor, 1991)
Didelphis aurita, 874 m
Campinho; Espirito Santo, Brazil (2007'S,
4018'W, Hershkovitz, 1992)
Marmosops incanus, sea level (2009'S.
4017'W)
Philander frenata, 500 m
Campo Alegre; Sucre, Venezuela (ca. 1010'N,
6345'W, Paynter, 1982)
Marmosa robinsoni
Campo Bello, Itatiaya (see Pico das Agulhas Ne-
gras); Rio de Janeiro, Brazil
Campo Duran, 8 km S, 8 km E; Salta, Argentina
(2214'S, 6342'W, USBGN)
Thylamys venustus
Campo Grande, 13 km N; Mato Grosso do Sul,
Brazil (2027'S, 5437'W, Paynter and Tray-
lor, 1991)
Didelphis albiventris
Micoureus constantiae
Campo Grande, 17 km E of Santo Domingo; Sal-
ta, Argentina (2437'S, 6320'W, Flores et al.,
2000)
Thylamys pusilla
Campo Sales; Ceara, Brazil (0704'S, 4023'W,
USBGN)
Monodelphis domestica (Si'tio Volta)
Camp Pifia, Fort Sherman; Colon, Panama
(0922'N, 7957'W, Geonet)
Didelphis marsupialis
Metachirus nudicaudatus (0916'N, 8000'W)
Philander opossum
Camp Rangel; Aragua, Venezuela (not located)
Didelphis marsupialis
Camp Rangel; Guarico, Venezuela (not located)
Caluromys philander
Camp Seattle, Osa Peninsula; Puntarenas, Costa
Rica (Osa Peninsula = 0834'N, 8331'W,
USBGN)
Didelphis marsupialis
Cana (= Santa Cruz de Cafia); Darien, Panama
(0747'N, 7742'W, Gardner, 1973)
Caluromys derbianus
Chironectes minimus, 606 m, 2000 ft (= 625
m) (upper Rio Tuyra, Rio Tuyra = 0821'N,
7803'W)
Didelphis marsupialis
Marmosa robinsoni, 1800 ft (= 560 m), 600 m
Marmosops invictus, 610 m, 2000 ft (= 625 m)
Metachirus nudicaudatus
Monodelphis adusta, 2000 ft (= 625 m) (upper
Rio Tuyra, Rio Tuyra = 0821'N, 7803'W)
Canaa; Minas Gerais, Brazil (2040'S, 4235'W,
Varejao and Valle, 1982)
Didelphis aurita
Cana Brava; Nova Roma, Goias, Brazil (1340'S,
4807'W, Hershkovitz, 1992)
Gracilinanus agilis
Monodelphis domestica
Canar; Canar, Ecuador (0233'S, 7856'W, Payn-
ter, 1993)
Didelphis albiventris, 2600-3176 m
Canay (see Restrepo); Meta, Colombia
Canelones, 4 km N; Canelones, Uruguay
(3432'S, 5617'W, Paynter, 1994)
Monodelphis dimidiata
Canelos; Pastaza, Ecuador (0135'S, 7745'W,
Paynter, 1993)
Caluromys lanatus, 600 m (Rio Bobonaza)
Marmosops noctivagus, 530 m
Micoureus regina, 530 m (left bank Rio Bo-
bonaza)
Caney, Rib Guatiqufa (see Restrepo); Meta, Co-
lombia
Cano Alpargaton; Carobobo, Venezuela (1028'N,
6815'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Cano Anaru; Barinas, Venezuela (0723'N,
7044'W, Perez-Hernandez, 1989)
Philander opossum
Cano Avispa; Vichada. Colombia (0516'N,
7042'W, Geonet)
Lutreolina crassicaudata, 100 m
192
FIELDIANA: ZOOLOGY
Cano Bartalo; Guarico, Venezeuela (not located)
Marmosa robinsoni
Cano Cotua, Boca, Parque Nacional Yapacana;
Amazonas, Venezuela (0341'N, 6652'W,
Perez-Hernandez, 1989)
Caluromys lanatus
Cano Culebra, Cerro Duida; Amazonas, Venezue-
la (0337'N, 6541'W, Paynter, 1982)
Caluromys philander, 750 m
Didelphis albiventris, 750 m, 1400 m (50 km
NNW of Esmeralda)
Micoureus demerarae, 750 m
Cano Entrada, Sierra de la Macarena; Meta, Co-
lombia (0308'N, 7252'W, Paynter, 1997)
Marmosa lepida, 550 m
Cano Leon, Cerro Duida; Amazonas, Venezuela
(0325'N, 6540'W, Paynter, 1982)
Didelphis marsupialis
Cano Majagua, Rio Ventuari; Amazonas, Vene-
zuela (0520'N, 6540'W, Hershkovitz, 1997)
Philander andersoni
Canon del Rio Pita; Pichincha, Ecuador (ca.
0018'S, 7828'W, Paynter, 1993)
Caenolestes caniventer
Caenolestes fuliginosus
Didelphis albiventris
Cano Tigre, Zea; Merida, Venezuela (0851'N,
7123'W, Geonet)
Didelphis marsupialis, 450 m
Monodelphis palliolata, 450 m
Philander opossum, 450 m
Cano Yagua, Cerro Cucurito; Amazonas, Vene-
zuela (0335'N, 6646'W, Geonet)
Caluromys lanatus, 120 m
Philander andersoni, 120 m
Cano Yureba, Salto Yureba; Amazonas, Venezue-
la (04 1 l'N, 6642'W, Geonet)
Metachirus nudicaudatus
Cantareira (see Serra da Cantareira); Sao Paulo,
Brazil
Cantareira Forestal Preserve (see Serra da Can-
tareira); Sao Paulo, Brazil
Canudos; Para, Brazil (0716'S, 5807'W, Hersh-
kovitz, 1997)
Philander opossum
Capao Bonito (see Fazenda Intervales); Sao Pau-
lo, Brazil
Caparo; Caroni, Trinidad and Tobago (1027'N,
6120'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni
Marmosops fuscatus
Capatarida; Falcon, Venezuela (1110'N,
7037'W, Handley, 1976)
Marmosa xerophila, 40-75 m
Capeti; Darien, Panama (0804'N, 7733'W, Gard-
ner, 1973)
Didelphis marsupialis
Capibara, Brazo Casiquiare, 106 km SW of Es-
meraldas; Amazonas, Venezuela (0237'N,
6619'W, Handley, 1976)
Didelphis marsupialis, ca. 100 m, 130 m
Marmosa murina, 130 m
Marmosops parvidens, 1 30 m
Metachirus nudicaudatus, 130 m
Micoureus demerarae, 130 m
Monodelphis brevicaudata, 130 m (left bank
Brazo Casiquiare)
Philander andersoni, 130 m
Capim (= km 87-94, Belem-Brasilia); Para, Bra-
zil (0140'S, 4747'W, Paynter and Traylor,
1991)
Didelphis marsupialis, 25 m
Gracilinanus emiliae
Metachirus nudicaudatus (8 km SE of Belem
= 0130'S, 4820'W, km 87-94, Belem-Bra-
silia)
Philander opossum (0130'S, 4820'W)
Capital; Sao Paulo, Brazil (not located)
Monodelphis sorex
Thylamys velutinus
Carabaya (province); Puno, Peru (1350'S,
7015'W, Stephens and Traylor, 1983)
Chironectes minimus
Caracaraf, Parana do Pocao, Rio Mucajai (= Pa-
rana do Pocao); Roraima, Brazil (0150'N,
6108'W, Paynter and Traylor, 1991)
Didelphis marsupialis (also at 0249'N,
6040'W)
Philander opossum
Caracas; Distrito Federal, Venezuela (1030'N,
6655'W, Paynter, 1982)
Chironectes minimus, 1 1 10-1 170 m
Didelphis marsupialis, 917-1170 m
Caracato; La Paz, Bolivia (1659'S, 6749'W, An-
derson, 1997)
Thylamys pallidior
Cara de Mono; Zelaya, Nicaragua (1207'N,
8428'W, Geonet)
Chironectes minimus, 50 m
Philander opossum, 50 m
Caraiben-Niederlassungung Arrai am obern Pom-
eroon (see Caribe Colony of Arrai); West De-
merara-Essequibo Islands, Guyana
Caranavi; La Paz, Bolivia (1546'S, 6736'W,
Paynter, 1992)
Didelphis marsupialis
Marmosops impavidus
BROWN: ATLAS OF NEW WORLD MARSUPIALS
193
Metachirus nudicaudalus (20 km NNE of Car-
anavi)
Micoureus regina
Caranda, 2 km S; Santa Cruz, Bolivia (1733'S,
6332'W, Anderson et al., 1993)
Marmosops impavidus
Carandaiti (see Carandaity); Chusquisaca, Bolivia
Carandaity (= Carandaiti), 30 km SE; Chusquis-
aca, Bolivia (2045'S, 6304'W, Paynter,
1992)
Monodelphis domestica
Thylamys pusilla
Thylamys venustus
Carao, Cajamarca [probably Caras in Ancash];
Peru (Caras = 0904'S, 7749'W, Geonet)
Didelphis albiventris, 2200 m
Carapari; Tarija, Bolivia (2149'S, 6346'W, Payn-
ter, 1992)
Micoureus constantiae
Thylamys venustus, 1000 m
Caratateua, E bank Ri'o Caete; Para, Brazil
(0059'S, 4642'W, USBGN)
Metachirus nudicaudatus
Caratinga (= Sao Joao de Jacutinga); Minas Ger-
ais, Brazil (1946'S, 4145'W, Varejao and
Valle, 1982)
Didelphis aurita
Cardenas; Tabasco, Mexico (1759'N, 9322'W,
Geonet)
Didelphis marsupialis
Philander opossum
Cardona, 3 km E; Soriano, Uruguay (3353'S,
5723'W, USBGN)
Didelphis albiventris
Cariaco; Sucre, Venezuela (1050'N, 6343'W,
Lopez-Fuster et al., 2000)
Marmosa robinsoni
Cariari; Limon, Costa Rica (1022'N, 8344'W,
Gardner, 1973)
Chironectes minimus
Didelphis marsupialis
Caribe Colony of Arrai (= Caraiben-Niederlas-
sungung Arrai am obern Pomeroon), upper
Pomeroon River; West Demerara-Essequibo
Islands, Guyana (ca. 0710'N, 5840'W,
Schomburgk, 1848)
Marmosa murina
Caripe; Monagas, Venezuela (1012'N, 6329'W,
Paynter, 1982)
Chironectes minimus, 860 m
Marmosa robinsoni
Carlazo; Tarija, Bolivia (2128'S, 6432'W,
USBGN)
Didelphis albiventris, 2300 m
Thylamys venustus, 2300-2400 m
Carmo da Cachoeira; Minas Gerais, Brazil
(2130'S, 4515'W, Varejao and Valle, 1982)
Didelphis aurita
Carmopolis de Minas; Minas Gerais, Brazil
(2033'S, 4438'W, Geonet)
Didelphis albiventris
Carolina; Maranhao, Brazil (0720'S, 4728'W,
Paynter and Traylor. 1991)
Caluromys philander, 169 m
Carolina Mine, 4 km W of Divisadero; Morazan,
El Salvador (Divisadero = 1336'N,
8804'W, Geonet)
Didelphis marsupialis, 750 ft (= 230 m)
Carondelet; Esmeraldas, Ecuador (0106'N,
7842'W, Paynter, 1993)
Marmosa robinsoni, 70 m
Philander opossum, 60 ft (= 18 m) (0103'N,
7850'W)
Carrasco; Montevideo, Uruguay (3450'S,
5603'W, Paynter, 1994)
Lutreolina crassicaudata
Carretera El Limon; Distrito Federal, Venezuela
(1028'N, 6717'W, Perez-Hernandez, 1989)
Gracilinanus marica
Didelphis marsupialis, 600 m, 1750 m
Monodelphis palliolata, 1 750 m
Carretera Calabozo-San Fernando de Apure, Par-
celamiento; Guarico, Venezuela (0816'N,
6736'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Carretera Caracas, La Colonia Tovar, km 40; Dis-
trito Federal, Venezuela (1028'N, 6705'W,
Perez-Hernandez. 1989:381)
Gracilinanus marica
Marmosops fuscatus
Carretera Central, Cordillera Carpish; Huanuco,
Peru (Cordillera Carpish = 0955'-0918'S,
7614'-7559'W, Stephens and Traylor,
1983)
Didelphis albiventris, 2400 m (E slope Cordil-
lera Carpish)
Carretera El Dorado-Santa Elena; Bolivar, Vene-
zuela (556'N, 6127'W, Perez-Hernandez,
1989)
Micoureus demerarae, km 124 at 1350 m, km
125 at 1100 m, km 126 at 1200 m, km 134
at 1300 m
Carretera El Dorado-Santa Elena de Uairen, 70
km; Bolivar, Venezuela (06WN, 6127'W,
Perez-Hernandez, 1989)
Didelphis albiventris
Carretera El Limon; Distrito Federal, Venezuela
194
FIELDIANA: ZOOLOGY
(1028'N, 6717'W, Perez-Hernandez,
1989)
Monodelphis palliolata, 1750 m
Carretera El Junquito (see El Junquito); Distrito
Federal, Venezuela
Carretera San Carlos-Manrique; Cojedes, Vene-
zuela (0950'N, 6832'W, Perez-Hernandez,
1989)
Didelphis marsupialis
Carretera Vieja de Cata; Apure, Venezuela
(1028'N, 6744'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Carrizal; Miranda, Venezuela (1021'N, 6659'W,
Geonet)
Didelphis marsupialis
Carrizales; Miranda, Venezuela (1021'N,
6702'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 1 100 m
Marmosops fuscatus
Cartago; Cartago, Costa Rica (0952'N, 8355'W,
Geonet)
Didelphis marsupialis, 1432 m
Micoureus alstoni
Casabindo; Jujuy, Argentina (2258'S, 6600'W,
USBGN)
Thylamys pallidior, 4000-4500 m
Casa Grande, Boraceia, 20 km S of Salesopolis;
Sao Paulo, Brazil (Salesopolis = 2332'S,
4551'W, Hershkovitz, 1997)
Monodelphis sorex
Philander frenata
Casa Morava (= Azunta), 18 km NW of San Joa-
quin; Beni, Bolivia (San Joaquin = 1304'S,
6449'W, Geonet)
Didelphis marsupialis
Casa Nova, Rio Arapuins; Para, Brazil (0218'S,
5539'W, Hershkovitz, 1977)
Marmosa murina
Casarabe, 16 km W; Beni, Bolivia (1448'S,
6414'W, Geonet)
Didelphis marsupialis, 230 m
Philander opossum
Cascada Diana, 16 km WNW of Villa La Angos-
tura; Neuquen, Argentina (Villa La Angos-
tura = 4047'S, 7140'W, USBGN)
Dromiciops gliroides
Cascajal; Beni, Bolivia (1213'S, 6513'W, An-
derson, 1997)
Micoureus constantiae
Caserio Boro (= La Concordia); Lara, Venezuela
(0953'-1002'N, 6926'-6947'W, Handley,
1976)
Didelphis marsupialis, 5 1 8-900 m
Marmosa robinsoni, 528-616 m
Casiquiare Canal, Cerro Duida (see Brazo Casi-
quiare); Amazonas, Venezuela
Casita; Darien, Panama (0801'N, 7722'W, Wen-
zel and Tipton, 1966)
Marmosops invictus, 458 m
Caspirola; San Jose, Costa Rica (0941'N,
8417'W, Geonet)
Caluromys derbianus
Castanhal, Rio Jamunda [Nhamunda]; Para, Brazil
(not located)
Monodelphis brevicaudata
Castano Nuevo; San Juan, Argentina (3102'S,
6933'W, Geonet)
Thylamys pallidior, 5040 ft (= 1575 m)
Castro Alves; Bahia, Brazil (1245'S, 3926'W,
Geonet)
Didelphis albiventris
Casupito, Parque Nacional Guatopo; Miranda, Ve-
nezuela (1003'N, 681 l'W, Geonet)
Chironectes minimus, 350 m
Catacamas; Yoro, Honduras (1448'N, 8554'W,
Geonet)
Caluromys derbianus
Catamutun;' Los Lagos, Chile (4007'S, 7307'W,
Geonet)
Dromiciops gliroides
Catemaco; Veracruz-Llave, Mexico (1825'N,
9507'W, Geonet)
Didelphis marsupialis
Philander opossum
Caucaia do Alto, Cotia; Sao Paulo, Brazil
(2341'S, 4702'W, Geonet)
Marmosops incanus, 1000 m
Cauca Seca (= Rio Cauca); Colombia (not pre-
cisely located)
Didelphis marsupialis
Caura; St. George, Trinidad and Tobago (1043'N,
6121'W, Geonet)
Caluromys philander, 600 ft (= 180 m)
Didelphis marsupialis
Marmosa robinsoni, 500 ft (= 150 m), 600 ft
(= 180 m)
Caxiricatuba, right bank Rfo Tapajos; Para, Brazil
(0236'S, 5456'W, Hershkovitz, 1977)
Caluromys philander
Didelphis marsupialis
Cayafate (see Cafayate); Salta, Argentina
Cayambe, foot of Volcan Cayambe; Pichincha,
Ecuador (0003'N, 7808'W, Paynter, 1993)
Didelphis albiventris, 9340 ft (= 2900 m)
Cayenne; French Guiana (0456'N, 5220'W, Ste-
phens and Traylor, 1985)
Caluromys philander
Chironectes minimus, sea level
BROWN: ATLAS OF NEW WORLD MARSUPIALS
195
Didelphis marsupialis, sea level
Marmosa murina
Metachirus nudicaudatus
Monodelphis brevicaudata
Philander opossum
Cayetue. Lago Todos Santos; Los Lagos, Chile
(4114'S, 7217'W. Paynter. 1988)
Dromiciops gliroides, 250 m
Cayo; Cayo District. Belize (ca. 1709'N.
8904'W, Geonet)
Philander opossum
Cayo de Agua: Bocas del Toro. Panama (0909'N.
8202'W, Geonet)
Caluromys derbianus (Punta Norte)
Philander opossum
Cayo Nancy; Bocas del Toro. Panama (0920'N.
8213'W, Geonet)
Philander opossum
Ccolpa de Guacamayos. W bank Rio Tambopata;
Madre de Dios, Peru (1308'S. 6936'W, Em-
mons et al.. 1994)
Didelphis marsupialis
Marmosops noctivagus
Metachirus nudicaudatus
Philander opossum
Cebollal (= Seboyal): Loja. Ecuador (0402'S.
8002'W. Hershkovitz. 1997)
Philander opossum. 3100 ft (= 970 m)
Cedar Creek. Isla Bastimentos: Bocas del Toro.
Panama (0917'N. 8207'W, Geonet)
Caluromys derbianus
Cedrobamba Ruins. Macchu Pichu (see Macchu
Pichu, Urumbamba. Cedrobamba Ruins);
Cusco, Peru
Celendin; Cajamarca. Peru (0652'S. 7809'W.
Geonet)
Didelphis albiventris, 2709 m
Celica; Loja. Ecuador (0407'S, 7659'W. Paynter.
1993)
Marmosops impavidus, 6900 ft (= 2150 m)
Centinela (= Puerto Centinela). 1.5 km E of San
Joaquin; Beni, Bolivia (San Joaqufn =
1304'S, 6449'W. Hershkovitz. 1997)
Marmosa murina
Philander opossum
Central Camp, Cerro Duida (see Middle Camp);
Amazonas, Venezuela
Central Camp, Cumbre #1. Cerro Duida; Ama-
zonas, Venezuela (Cerro Duida = 0325'N.
6540'W. Paynter, 1982)
Didelphis albiventris. 4800 ft (= 1500 m)
Central Farm; Cayo. Belize (17H'N, 89WW.
Geonet)
Didelphis marsupialis
Philander opossum
Centro Argentino de Primates, 20 km SE. Corri-
entes; Corrientes. Argentina (not located)
Didelphis albiventris
Lutreolina crassicaudata
Centro de Investigaciones de Fauna Silvestre, El
Limon; Aragua. Venezuela (1019'N,
6738'W, Perez-Hernandez. 1989)
Didelphis marsupialis
Centro de Ski Antillanca (see Antillanca Ski
Area); Osorno, Chile
Centro Dieciocho: Pando, Bolivia (10 C 36'S.
6647'W. Anderson, 1997)
Monodelphis brevicaudata
Monodelphis emiliae
Centro Union, Rio Aucayo; Loreto, Peru (not lo-
cated)
Chironectes minimus
Centurion; Concepcion, Paraguay (ca. 2215'S.
5735'W, Paynter, 1989)
Didelphis albiventris
Cerro; Montevideo. Uruguay (Montevideo =
3453'S. 56H'W. Paynter. 1994)
Lutreolina crassicaudata
Cerro Amboro. Rio Pitisama (also see Rio Pitis-
ama). 4.5 km N, 1.5 km E: Santa Cruz, Bo-
livia (1744'S, 6339'W, Paynter, 1992)
Didelphis marsupialis, 620 m
Mannosops dorothea
Cerro Antisana. Oriente; Napo, Ecuador (0030'S.
7808'W. Paynter. 1993)
Caenolestes fuliginosus, 4000 m
Cerro Auyan-tepui (see Auyan-tepuf); Bolivar.
Venezuela
Cerro Azul; Loreto, Peru (ca. 0713'S, 7430'W,
Stephens and Traylor. 1983)
Marmosa murina, ca. 500 m
Metachirus nudicaudatus, ca. 2000 ft (= 625
m) (E bank Rio Ucayali, 0714'S, 7434'W)
Cerro Azul (= La Zumbadora); Panama. Panama
(0910'N. 7925'W, Geonet)
Chironectes minimus, 2000 ft (= 625 m)
Didelphis marsupialis
Marmosops invictus, 265-1000 m, 640 m
Metachirus nudicaudatus
Philander opossum
Cerro Azul. near; Zulia. Venezuela (1051'N,
7216'W, Handley. 1976)
Didelphis marsupialis, 75-80 m
Marmosa robinsoni, 75-80 m
Metachirus nudicaudatus
Philander opossum
Cerro Bajo Verde, near Guayaquil; Guayas, Ec-
196
FIELDIANA: ZOOLOGY
uador (Guayaquil = 0210'S, 7950'W, Payn-
ter, 1993)
Marmosa robinsoni
Cerro Blanco; Guayas, Ecuador (0210'S,
8002'W, Paynter, 1993)
Didelphis marsupialis, ca. 100-420 m
Cerro Cacaguatique, N slope; San Miguel, El Sal-
vador (1346'N, 8813'W, Geonet)
Didelphis marsupialis, 3500 ft (= 1090 m),
1090 m, 4500 ft (= 1400 m)
Cerro Calilegua (see Serranfa de Calilegua); Ju-
juy, Argentina
Cerro Campana; Panama, Panama (0843'N,
7954'W, Geonet)
Metachirus nudicaudatus
Philander opossum
Cerro Casabindo; Jujuy, Argentina (2256'S,
6607'W, Flores et al., 2000)
Thylamys pallidior, 4500 m.
Cerro Cayambe (see Volcan Cayambe); Pichin-
cha, Ecuador
Cerro Chaguar, Parque Nacional Baritii; Salta, Ar-
gentina (2235'S, 6437'W, Flores et al.,
2000)
Thylamys venustus
Cerro Chimborazo; Chimborazo, Ecuador
(0128'S, 7848'W, Paynter, 1993)
Caenolestes fuliginosus, 4000 m
Cerro Chinguela, 5 km NE of Zapalache; Piura,
Peru (0507'S, 7923'W, Paynter, 1983)
Caenolestes caniventer, 2900 m
Caenolestes fuliginosus, 2900 m
Cerro Chucanti; Bocas del Toro, Panama (not lo-
cated)
Caluromys derbianus
Cerro Colorado; Santa Cruz, Bolivia (1927'S,
6221'W, Anderson, 1997)
Monodelphis domestica
Thylamys venustus
Cerro Cora; Amambay, Paraguay (2237'S,
5559'W, Geonet)
Didelphis albiventris
Cerro Cotacachi; Imbabura, Ecuador (0022'N,
7820'W, Paynter, 1993)
Caenolestes fuliginosus, 8000 ft (= 2500 m)
Cerro Cucurito; Amazonas, Venezuela (0338'N,
6625'W, Hershkovitz, 1997)
Philander andersoni
Cerro de Famatima (see Loma de Famatima); La
Rioja, Argentina
Cerro de La Neblina (= Cerro La Neblina = La
Neblina); Amazonas, Venezuela (ca.
0050'N, 6604'W, Perez-Hernandez, 1989)
Caluromys lanatus (Base Camp, Rio Mawar-
inuma)
Didelphis albiventris, 2100 m
Didelphis marsupialis
Marmosops impavidus, 2100 m
Marmosops neblina
Metachirus nudicaudatus
Cerro del Campo, 100 km NE of San Miguel de
Tucuman, Burruyacu; Tucuman, Argentina
(San Miguel de Tucuman = 2649'S,
6413'W, Paynter, 1985)
Didelphis albiventris, 800 m
Thylamys venustus, 1300 m, 2400 m
Cerro de Raco, Trancas; Tucuman, Argentina
(2643'S, 6530'W, Mares et al., 1996)
Thylamys venustus
Cerro de Santa Ana, Penisula de Paraguana; Fal-
con, Venezuela (1148'N, 6955'W, Lopez-
Fuster et al., 2000)
Marmosa robinsoni
Cerro de Taff Viejo, Taff Viejo; Tucuman, Argen-
tina (Taff Viejo = 2644'S, 6516'W, Paynter,
1985)
Thylamys venustus, 2400 m
Cerro de Tucuman (see Serra de Tucuman); Tu-
cuman, Argentina
Cerro Duida; Amazonas, Venezuela (0325'N,
6540'W, Paynter, 1982)
Didelphis albiventris, 1400 m, 2240 m
Didelphis marsupialis, 2240 m
Marmosa tyleriana, 4800-6900 ft (= 1500-
2150 m) (plateau Mt. Duida)
Micoureus demerarae, 100-2000 m
Monodelphis brevicaudata, 2240 m
Cerro Duida, base; Amazonas, Venezuela
(0325'N, 6540'W, Paynter, 1982)
Marmosa murina, 100 m
Metachirus nudicaudatus, 300 ft (= 90 m)
Cerro El Avila; Distrito Federal, Venezuela
(1031'N, 6652'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Marmosops fuscatus
Monodelphis palliolata
Cerro El Baiil, Ranch Carlos Minne; Oaxaca,
Mexico (ca. 1633'N, 9410'W, Gardner,
1973)
Didelphis marsupialis
Cerro Hermoso, Parque Nacional Calilegua; Ju-
juy, Argentina (2335'S, 6453'W, Flores et
al., 2000)
Thylamys venustus, 2800 m
Cerro Hosana; Santa Cruz, Bolivia (1727'S,
6340'W, Anderson et al., 1993)
Marmosops dorothea
BROWN: ATLAS OF NEW WORLD MARSUPIALS
197
Micoureus constantiae
Cerro Hoya; Los Santos, Panama (0718'N,
8042'W, Geonet)
Chironectes minimus, 1450 ft (= 450 m)
Didelphis marsupialis
Philander opossum
Cerro Ibiapaba (see Serra Ibiapaba); Ceara, Brazil
Cerro Iliniza (= Mt. Illininiza); Pichincha, Ecua-
dor (0040'S, 7842'W, Paynter, 1993)
Caenolestes fuliginosus, 12000 ft (= 3750 m)
Cerro Itahuatica; Santa Cruz, Bolivia (1948'S,
6331'W, Anderson, 1997)
Thy I amy s venustus
Cerro La Neblina (see Cerro de La Neblina);
Amazonas, Venezuela
Cerro Las Copas, La Revancha; Tachira, Vene-
zuela (0727'N, 7226'W, Perez-Hernandez,
1989)
Gracilinanus dryas, 2200 m
Didelphis albiventris, 2200 m
Marmosops impavidus, 2200 m
Cerro Leones, 16 km ENE of San Carlos de Bar-
iloche (= Cerro Lesnea); Rio Negro, Argen-
tina (San Carlos de Bariloche = 4109'S,
7118'W, Paynter, 1985)
Lestodelphys halli
Cerro Lesnea (see Cerro Leones); Rio Negro, Ar-
gentina
Cerro Mali; Darien, Panama (0807'N, 7714'W,
Geonet)
Chironectes minimus
Cerro Manglaralto; Guayas, Ecuador (0150'S,
8020'W, Paynter, 1993)
Didelphis marsupialis, 375-450 m
Marmosa robinsoni
Metachirus nudicaudatus, 1200-1500 ft (=
375-460 m) (that portion of the Sierra de Co-
lonche east of the town of Manglaralto)
Cerro Matasiete, Isla Margarita; Nueva Esparta,
Venezuela (1 103'N, 6351'W, Perez-Hernan-
dez, 1989)
Marmosa robinsoni, 120-524 m
Cerro Medici; Tucuman, Argentina (2637'S,
6512'W, Flores et al., 2000)
Didelphis albiventris
Thylamys pusilla
Thylamys venustus
Cerro Munchique; Cauca, Colombia (0232'N,
7657'W, Paynter, 1997)
Didelphis marsupialis, 1750 m, 2000 m, 2100
m, 2300 m, 2400 m, 2570 m
Cerro Nahuelbuta (= Sierra Nahuelbuta); Los La-
gos, Chile (3743'S, 7304'W, Geonet)
Dromiciops gliroides, 1440 m
Cerro Nielol; Araucania, Chile (3840'S, 7238'W,
Geonet)
Dromiciops gliroides
Cerro Papelon; Monagas, Venezuela (1001'N,
6354'W, Perez-Hernandez, 1989)
Marmosa robinsoni, 900 m
Philander opossum, 900 m
Cerro Papelon; Sucre, Venezuela (1011'N,
6349'W, Perez-Hernandez, 1989)
Marmosops fuscatus, 900 m
Cerro Pata de Pajaro; Manabi, Ecuador (0002'N,
7958'W, Paynter, 1993)
Caluromys derbianus, 800 m
Cerro Pichincha, crater; Pichincha, Ecuador
(0010'S, 7833'W, Paynter, 1993)
Caenolestes fuliginosus
Cerro Pilado; Caaguazu, Paraguay (2500'S,
5530'W, Krumbiegel, 1941b)
Lutreolina crassicaudata
Cerro Pirre, near head of Rio Limon; Darien, Pan-
ama (0756'N, 7743'W, Geonet)
Didelphis marsupialis
Marmosops impavidus, 5300 ft (= 1650 m)
Cerro Punta; Chiriqm, Panama (0853'N,
8235'W, Geonet)
Didelphis marsupialis
Cerro Robles, 125 km NNW of Santiago; Maule,
Chile (3514'S, 7123'W, Geonet)
Thylamys elegans, 1600 m
Cerro Roraima (= Monte Roraima = Mt. Rorai-
ma) (see also Arabopo); Bolivar, Venezuela
(0512'N, 6044'W, Paynter, 1982)
Didelphis albiventris
Marmosa murina, 1400 m, 4200 ft (= 1300 m)
(foot of Cerro Roraima)
Micoureus demerarae, 4200 ft (= 1300 m)
(foot of Cerro Roraima)
Cerro Sajama (see Sajama Nevado); Oruro, Bo-
livia
Cerro San Javier; Tucuman, Argentina (2647'S,
6524'W, Mares et al., 1996)
Didelphis albiventris
Lutreolina crassicaudata
Thylamys venustus, 1000 m
Cerro Santa Ana, Peninsula de Paraguana; Falcon,
Venezuela (1149'N, 6956'W, Geonet)
Marmosa robinsoni, 500-615 m
Cerro Santa Barbara (= Sierra Santa Barbara); Ju-
juy, Argentina (2407'S, 6429'W, Paynter,
1985)
Lutreolina crassicaudata, 1800 m
Cerros del Tavara; Madre de Dios, Peru (1302'S,
6941'W, Emmons et al., 1994)
Micoureus demerarae, 500 m
198
FIELDIANA: ZOOLOGY
Cerro Socopo; Falcon, Venezuela (1028'N,
7048'W, Handley, 1976)
Marmosa robinsoni, 1257-1280 m
Cerro Tacarcuna; Darien, Panama (0811'N,
7717'W, Geonet)
Caluromys derbianus
Marmosa robinsoni
Marmosops invictus, 1464 m, 4800 ft (= 1500
m) (0810'N, 7718'W)
Cerro Turumiquire; Sucre-Monagas, Venezuela
(1007'N, 6352'W, Paynter, 1982)
Gracilinanus marica, 8500 ft (= 2650 m),
2596 m
Chacarcito, Itoco, near Bogota; Cundinamarca,
Colombia (Bogota = 0436'N, 7405'W,
Paynter, 1997)
Micoureus regina
Chachapoyas; Amazonas, Peru (0613'S,
7751'W, Geonet)
Marmosops noctivagus, 9000 ft (= 2800 m)
Chaco (= Gran Chaco), area between Ribs Pil-
comayo and Paraguay, NW Asuncion; Pres-
idente Hayes, Paraguay (ca. 2521'-2718'S,
5740'-5838'W, Paynter, 1989)
Lutreolina crassicaudata
Chacras de Coria; Mendoza, Argentina (3300'S,
6852'W, Paynter, 1985)
Lestodelphys halli
Challao; Mandoza, Argentina (3251'S, 6856'W,
Paynter, 1985)
Thylamys pallidior
Challapata; Oruro, Bolivia (1854'S, 6647'W,
Paynter, 1992)
Thylamys pallidior, 3700-3800 m
Thylamys venustus
Challoma, Sandia, 3 mi below San Juan on Rio
Tambopata; Puno, Peru (not located)
Didelphis marsupialis, 5000 ft (= 1560 m)
Chama, Sierra de Merida; Merida, Venezuela
(0903'N, 7137'W, Paynter, 1982)
Marmosa robinsoni
Chamelecon (= Chemelacon); Cortes, Honduras
(1526'N, 8801'W, Geonet)
Caluromys derbianus
Chamicuros (see Rio Chamicuros); Loreto, Peru
Chanchamayo (see also Pueblo Nuevo); Junfn,
Peru (ca. 1103'S, 7519'W, Stephens and
Traylor, 1983)
Caluromys lanatus, 1200 m
Chironectes minimus
Didelphis albiventris, 1200 m
Hyladelphys kalinowskii, 1100 m (1110'S,
7520'W, Hershkovitz, 1992)
Marmosa murina (1055'S, 7518'W)
Marmosops noctivagus
Metachirus nudicaudatus, 1 200 m
Philander andersoni, 1 200 m
Changuinola; Bocas del Toro, Panama (0926'N,
8231'W, Geonet)
Caluromys derbianus
Chironectes minimus
Metachirus nudicaudatus (7 km SSW of Chan-
guinola)
Philander opossum
Chapada (see Chapada dos Guimaraes); Mato
Grosso, Brazil
Chapada dos Guimaraes (= Chapada = Santa Ana
[Anna] de Chapada); Mato Grosso, Brazil
(1526'S, 5545'W, Paynter and Traylor,
1991)
Chironectes minimus
Didelphis albiventris, 793 m
Micoureus constantiae
Thylamys pusilla
Chapadao dos Parecis (see Serra dos Parecis);
Mato Grosso, Brazil
Chapadamal Formation; Buenos Aires, Argentina
(not located)
Lutreolina crassicaudata
Chapare; Cochabamba, Bolivia (1711'S,
6549'W, Anderson, 1997
Didelphis marsupialis
Chapiquina; Tarapaca, Chile (1823'S, 6933'W,
Geonet)
Thylamys elegans, 10500 ft (= 3280 m)
Charapaya (see Charuplaya); Cochabamba, Bolivia
Charguayaco; Cauca, Colombia (ca. 0240'N,
7657'W, Paynter, 1997)
Didelphis albiventris, 2000 m
Didelphis marsupialis, 2000-2200 m
Charuplaya (= Charapaya); Cochabamba, Bolivia
(1722'S, 6645'W, Geonet)
Didelphis albiventris
Metachirus nudicaudatus, 1350 m
Chasquitambo; Ancash, Peru (1018'S, 7737'W,
Handley, 1956)
Thylamys pallidior
Chaupe; Cajamarca, Peru (0600'S, 7909'W,
Geonet)
Didelphis albiventris, 6100 ft (= 1900 m)
Chavantina (= Xavantina), 264 km N of Serra do
Roncador; Mato Grosso, Brazil (1251'S,
5146'W, Pineet al., 1970)
Caluromys philander
Marmosa murina
Marmosops bishopi, ca. 1750 ft (= 540 m)
Metachirus nudicaudatus
Micoureus demerarae
BROWN: ATLAS OF NEW WORLD MARSUPIALS
199
Thylamys pusilla (1249'S, 4245'W)
Chaves; Espirito Santo, Brazil (ca. 2006'S,
4032'W, Paynter and Traylor, 1991)
Monodelphis iheringi
Chemelacon (see Chamelecon); Cortes, Honduras
Chepa (Quebrada); Panama, Panama (0902'N,
7945'W, Geonet)
Chironectes minimus
Chepo; Panama, Panama (0910'N, 7906'W,
Geonet)
Caluromys derbianus
Philander opossum
Chiapa, 5 km S; Chiapas, Mexico (1642'N,
9300'W, Geonet)
Didelphis marsupialis
Chiapa de Corzo, 4 mi NE; Chiapas, Mexico
(1642'N, 9300'W, Geonet)
Didelphis marsupialis
Chical; Canar, Ecuador (0224'S, 7858'W, Payn-
ter, 1993)
Didelphis albiventris, 3125 m
Chicharras (see Las Chicharras); Chiapas, Mexico
Chichen Itza; Yucatan, Mexico (2040'N,
8834'W, Geonet)
Didelphis marsupialis
Marmosa canescens
Marmosa mexicana
Chichicafte; Veracruz-Llave, Mexico (not located)
Philander opossum
Chichirivichi; Distrito Federal, Venezuela
(1028'N, 6717'W, Perez-Hernandez, 1989)
Monodelphis palliolata
Chicosa; Ucayali, Peru (1028'S, 7403'W, Geo-
net)
Caluromys lanatus, 454 m, 1500 ft (= 500 m)
Marmosa murina
Philander opossum, 1000 ft (= 300 m)
Chijchijpa, Rio Yarisa; La Paz, Bolivia (1609'S,
6745'W, Mustrangi and Patton, 1997)
Didelphis marsupialis
Marmosops dorothea, 1114m
Marmosops noctivagus, 1114m, 1 224 m
Chijchipani; La Paz, Bolivia (1608'S, 6744'W,
Anderson, 1997)
Didelphis marsupialis
Marmosa murina
Metachirus nudicaudatus
Chilasco; Baja Verapaz, Guatemala (1507'N,
9006'W, Geonet)
Marmosa mexicana, 5000 ft (= 1560 m)
Chimanta-tepuf; Boli'var, Venezuela (0518'N,
6210'W, Paynter, 1982)
Didelphis albiventris, 6300 ft (= 1960 m)
Chimanta Massif, Lower Cumbre Camp, Chiman-
ta-tepuf, Rib Tiraca; Boli'var, Venezuela
(0518'N, 6210'W, Paynter, 1982)
Didelphis marsupialis, 6300 ft (= 1960 m)
Chimoxan; Alta Verapaz, Guatemala (1538'N,
8939'W, Geonet)
Didelphis marsupialis
Chinaja; Alta Verapaz, Guatemala (1601'N,
9013'W, Geonet)
Didelphis marsupialis, 550 ft (= 170 m)
Philander opossum, 550 ft (= 170 m)
Chinandega; Esteli, Nicaragua (1237'N,
8709'W, Geonet)
Marmosa mexicana
Chinchavito; Huanuco, Peru (0929'S, 7555'W,
Stephens and Traylor, 1983)
Marmosa murina, 3000 ft (= 930 m)
Marmosops noctivagus, 3000 ft (= 930 m)
Metachirus nudicaudatus, 3000 ft (= 930 m)
Micoureus regina, 3000 ft (= 930 m)
Chinchin Cocha; Pichincha, Ecuador (0003'N,
7810'W, Patterson and Albuja, Geonet lists
a Chinchin Lima at 0003'N, 7817'W)
Caenolestes fuliginosus, 4000 m
Chiquitos; Santa Cruz, Bolivia (1800'S, 6600'W,
Anderson, 1997)
Micoureus constantiae
Monodelphis domestica
Chiriaco, 43 km NE; Amazonas, Peru (Chiriaco
= 0510'S, 7818'W, Geonet)
Philander opossum, 328 m
Chirimoto; Amazonas, Peru (0631'S, 7724'W,
Stephens and Traylor, 1983)
Metachirus nudicaudatus, 5400 ft (= 1680 m)
(Rio Huambo)
Micoureus regina, 2100 m
Chiruli; Zulia, Venezuela (0909'N, 7104'W,
Perez-Hernandez, 1989)
Didelphis marsupialis
Chisqufo; Cauca, Colombia (0229'N, 7652'W,
Paynter, 1997)
Didelphis marsupialis, 1800 m
Chiva-Chiva; Panama, Panama (0902'N,
7935'W, Geonet)
Caluromys derbianus
Choachf; Cundinamarca, Colombia (0432'N,
7356'W, Paynter, 1997)
Didelphis albiventris, 1970 m
Chompoton (see La Tuxpena); Campeche, Mexico
Chongoncito, near Guayaquil; Guayas, Ecuador
(0214'S, 8005'W, Paynter, 1993)
Marmosa robinsoni, ca. 100 m
Chontales (Department); Nicaragua (Center of
Department = 1205'N, 8510'W, USBGN)
Metachirus nudicaudatus
200
FIELDIANA: ZOOLOGY
Choro (see Choros); Cochabamba, Bolivia
Choros (= Choro); Cochabamba, Bolivia
(1656'S, 6642'W, Anderson, 1997)
Monodelphis osgoodi, 3500 m
Choroni; Aragua, Venezuela (1029'N, 6737'W,
Paynter, 1982)
Chironectes minimus
Chorrera de Vegas; Amazonas, Venezuela (ca.
0323'N, 6536'W, Paynter, 1982)
Didelphis marsupialis, 4700 ft (= 1460 m)
Chos-Malal; Neuquen, Argentina (3723'S,
7016'W, Paynter, 1985)
Thylamys pallidior, 805 m
Chospiyoc (see Chospyoc); Cusco, Peru
Chospyoc (= Chospiyoc); Cusco, Peru (ca.
1316'S, 7221'W, Stephens and Traylor,
1983)
Didelphis albiventris, 10000 ft (= 3125 m)
Chota; Imbabura, Ecuador (0029'N, 7804'W,
Paynter, 1993)
Marmosa robinsoni
Choya, 13 km NNW of Andalgala; Catarnarca,
Argentina (2732'S, 6624'W, Mares et all,
1997)
Didelphis albiventris, 1219 m
Chulumani (= Culumani); La Paz, Bolivia
(1624'S, 6731'W, Hershkovitz, 1997)
Didelphis marsupialis
Metachirus nudicaudatus, 2000 m, 2200 m
Philander opossum
Chumbicha; Catarnarca, Argentina (2852'S,
6614'W, Paynter, 1985)
Thylamys pallidior, 415 m, 457 m, 460 m,
600 m
Churi-Tepui, 85 km SE of El Dorado; Bolivar, Ve-
nezuela (0513'N, 6154'W, Geonet)
Marmosops pinheiroi
Churuli, Caja Seca; Zulia, Venezuela (0909'N,
7104'W, Hershkovitz, 1997)
Philander opossum
Chuyayacu, 2 km E; Chuquisaca, Bolivia
(1943'S, 6351'W, Anderson, 1997)
Micoureus constantiae
Ciadade Mineradoras Brasileiras Reunidos, Ibicuf,
Mangaratiba (see Mangaratiba); Rio de Ja-
neiro, Brazil
Cidade Caete (= Caete, Cidade); Minas Gerais,
Brazil (1952'S, 4340'W, Varejao and Valle,
1982)
Didelphis aurita, 940 m, ca. 1 100 m
Cincinati (= Valparaiso); Magdalena, Colombia
(1109'N, 7406'W, Paynter, 1997)
Didelphis marsupialis, 4500-5500 ft (= 1400-
1700 m)
Marmosa robinsoni
Cinco Rancho, 28 km NE of Icabarii; Bolivar, Ve-
nezuela (Icabani = 0420'N, 6145'W, Geo-
net)
Metachirus nudicaudatus, 775 m
Cinincay (see Cuenca); Azuay, Ecuador
Cituro; Darien, Panama (0800'N, 7736'W, Wen-
zel and Tipton, 1966)
Didelphis marsupialis
Ciudad Bolivar; Bolivar, Venezuela (0808'N,
6333'W, Paynter, 1982)
Caluromys philander, ca. 100 m
Didelphis marsupialis, ca. 100 m
Marmosa murina, ca. 100 m
Marmosa robinsoni, ca. 100 m
Monodelphis brevicaudata, ca. 100 m
Ciudad Cortes (see Puerto Cortez); Puntarenas,
Costa Rica
Ciudade de Sao Goncalo do Rio Abaixo (= Fa-
zenda de Cascata = Esta?ao de Pesquisas de
Peti = Sao Goncalo do Rio Abaixo); Minas
Gerais, Brazil (1951'S, 4315'W, Varejao
and Valle, 1982)
Didelphis albiventris
Didelphis aurita (1950'S, 4324'W)
Marmosa incanus
Ciudad Guayana (= San Felix), right bank Rib
Orinoco; Bolivar, Venezuela (0823'N,
6240'W, Paynter, 1982)
Caluromys philander, 20 m
Ciudad Victoria, 70 km S (= El Carrizo); Tamau-
lipas, Mexico (2344'N, 9908'W, Geonet)
Philander opossum, 758 m, 2500 ft (= 780 m)
Clarines, 14 km W; Anzoategui, Venezuela
(0957'N, 6518'W, Handley, 1976)
Didelphis marsupialis, 100 m
Clarksdale, Demerara River (see New Saint Eu-
statius); Essequibo Islands-West Demerara,
Guyana
Clemente Onelli, E of; Rio Negro, Argentina
(4114'S, 7001'W, Paynter, 1985)
Lestodelphys halli, 95 1 m
Clevia, W bank Suriname River (= Plantation
Clevia), NE of "De Morgenstond"; Com-
mewijne, Suriname (0552'N, 5506'W, Geo-
net)
Caluromys philander
Didelphis marsupialis
Marmosa murina (0550'N, 5510'W)
Monodelphis brevicaudata
Philander opossum
Club Campestre, 1 mi S of El Poblado; Antioquia,
Colombia (El Poblado - 0613'N, 7534'W,
Paynter, 1997)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
201
Didelphis marsupialis
Coast Hills; Valparaiso, Chile (not located)
Thylamys elegans
Coban: Alta Verapaz. Guatemala (1529'N.
9022'W, Geonet)
Marmosa mexicana
Philander opossum
Cobija. 10 km S: Pando. Bolivia (1102'S,
6844'W, Paynter. 1992)
Didelphis marsupialis
Cocal. upper Rio San Juan; Cauca. Colombia
(0231'N, 77WW, Paynter. 1997 and Hersh-
kovitz, 1997)
Philander opossum, 125 m. 187 m
Cochabamba. cerca de; Cochabamba. Bolivia
(1724'S, 6609'W, Paynter. 1992)
Didelphis albiventris
Thylamys venustus, 2557 m
Cochabamba. km 5 from Camino; Santa Cruz.
Bolivia (1747'S. 6312'W. Paynter, 1992)
Didelphis marsupialis
Cocha Cashu; Madre de Dios, Peru (ca. H5rS,
7119'W. Stephens and Traylor. 1983)
Glironia venusta, 380 m
Metachirus nudieaudatus, 380 m (N bank Rio
Manu)
Micoureus regina, 380 m
Philander opossum, 380 m
Cocha Cashu Biological Station, ca. 80 mi up-
stream from mouth of Rio Manu. Parque Na-
cional del Manu; Madre de Dios. Peru (ca.
H51'S, 7119'W, Stephens and Traylor.
1983)
Caluromys lanatus, ca. 400 m
Caluromysiops irrupta, 400 m
Chironectes minimus
Didelphis marsupialis
Mannosops noctivagus
Micoureus demerarae
Cocoli; Panama. Panama (0859'N, 7935'W,
Geonet)
Philander opossum
Cocollar; Monagas. Venezuela (1010'N,
6347'W, Paynter, 1982)
Didelphis marsupialis, 800 m. 1 100 m
Cocos; Maranhao, Brazil (0435'S. 4340'W.
Paynter and Traylor, 1991)
Gracilinanus agilis, 72 m
Marmosa murina, 72 m
Coco Solo: Colon, Panama (0922'N. 7954'W.
Geonet)
Philander opossum
Codajas; Amazonas, Brazil (0350'S, 6205'W.
Paynter and Traylor, 1991)
Gracilinanus emiliae
Marmosa murina, 38 m (Rio Solimoes)
Micoureus demerarae, 38 m (Rio Solimoes)
Codo; Maranhao, Brazil (0429'S. 4353'W. Payn-
ter and Traylor, 1991)
Marmosa murina, 58 m
Coiba Island (see Isla Coiba): Veraguas. Panama
Colima; Cuscatlan. El Salvador (1403'N.
8908'W. Geonet)
Didelphis marsupialis
Collon Cura: Neuquen, Argentina (4007'S.
7044'W, Paynter. 1985)
Thylamys pallidior
Colocacao Vira-Volta. left bank Rio Jurua; Ama-
zonas. Brazil (0317'S. 6614'W. Patton et al.,
2000)
Caluromys lanatus (on Igarape Arabidi)
Mannosops impavidus
Mannosops panidens
Metachirus nudieaudatus
Micoureus demerarae
Colomi, 5 km N: Cochabamba. Bolivia (1721'S,
6552'W, Paynter. 1994)
Didelphis albiventris. 3270 m
Colon; Colon. Panama (0921'N. 7954'W, Geo-
net)
Didelphis marsupialis
Colon; Montevideo. Uruguay (3448'S, 5614'W,
Paynter. 1994)
Lutreolina crassicaudata
Monodelphis dimidiata
Colon (see San Juan de Colon): Tachira. Vene-
zuela
Colon and Code. W of; Veraguas, Panama (not
located)
Mannosa mexicana
Colonia Agn'cola de Caracolicito (de Santa Mar-
ta). Rio Ariguani: Cesar. Colombia (Caracol-
icito = 1012'N. 7358'W. Paynter, 1997)
Caluromys lanatus, 335 m
Didelphis marsupialis, 335-400 m
Colonia Agricola Turen (= Colonia Turen): Por-
tuguesa. Venezuela (0915'N. 6905'W. Geo-
net)
Chironectes minimus, 130 m
Didelphis marsupialis, 130 m
Colonia Alocar (see Colonia Alvear): Mendoza,
Argentina
Colonia Alpina. 16 km N of Teresopolis; Rio de
Janeiro. Brazil (Teresopolis = 2226'S,
4259'W, Paynter and Traylor. 1991)
Didelphis aurita
Chironectes minimus
Colonia Alvear (= Colonia Alocar = General Al-
202
FIELDIANA: ZOOLOGY
vear); Mendoza, Argentina (3458'S, 6742'W,
Geonet)
Thylamys pallidior
Colonia de Agrfcola de Caracolicito; (see Colonia
de Agrfcola de Santa Marta); Magdalena, Co-
lombia
Colonia de Agrfcola de Santa Marta (= Colonia
de Agrfcola de Caracolicito); Magdalena, Co-
lombia (Santa Marta = 1115'N, 7413'W,
Paynter, 1997)
Chironectes minimus
Colonia do Prata (see Igarape Acu); Para, Brazil
Colonia Hansa (see Corupa); Santa Catarina, Bra-
zil
Colonia Mennonita (= Mennoniten Kolinien);
Boqueron, Paraguay (2330'S, 6000'W,
Paynter, 1989)
Didelphis albiventris
Colonia Nueva Italia, Vileta; Central, Paraguay
(2537'S, 5730'W, Paynter, 1989)
Gracilinanus agilis
Lutreolina crassicaudata, 100 m
Colonia Somerfield #11; Caaguazu, Paraguay
(2526'S, 5543'W, Geonet)
Didelphis albiventris, 375 m
Didelphis marsupialis, 375 m
Metachirus nudicaudatus
Colonia Suiza; Colonia, Uruguay (3419'S,
5713'W, USBGN)
Lutreolina crassicaudata
Colonia Turen (see Colonia Agrfcola Turen); Por-
tuguesa, Venezuela
Colonnello; Zulia, Venezuela (not located)
Metachirus nudicaudatus
Comaccka (see Takama); Upper Demerara-Ber-
bice, Guyana
Comackka, 80 mi up Demerara River (see Taka-
ma); Upper Demerara-Berbice, Guyana
Comallo; Rio Negro, Argentina (4102'S,
7016'W, Paynter, 1985)
Lestodelphys halli
Comarapa, 25 km W; Cochabamba, Bolivia
(1751'S, 6440'W, Anderson, 1997)
Graciliananus agilis
Comarapa, 5 km SE; Santa Cruz, Bolivia
(1758'S, 6429'W, Anderson, 1997)
Thylamya pallidior
Comarapa, 6 km S, 10 km E; Santa Cruz, Bolivia
(1757'S, 6424'W, Anderson, 1997)
Didelphis albiventris, 1527 m
Thylamya pallidior
Comarapa, 15 km W; Santa Cruz, Bolivia
(1753'S, 6435'W, Anderson, 1997)
Didelphis albiventris, 2700 m
Combe; Suburb of Paramaribo, Suriname (Para-
maribo = 0550'N, 5511'W, Stephens and
Tray lor, 1985)
Caluromys philander
"Comechatiba" (see Ponta Cumuruxatiba); Ba-
hia, Brazil
Comisarfa del Vichada; Vichada, Colombia
(0515'N, 6752'W, Boher Bentti, 1988)
Lutreolina crassicaudata, 100 m
Comitan, 20 mi W; Chiapas, Mexico (1615'N,
9208'W, Geonet)
Didelphis marsupialis
Commewijne River; Commewijne, Suriname
(0554'N, 5505'W, Geonet)
Philander opossum
Comodoro Rivadavia (see Estancia Machijada);
Chubut, Argentina
Comoxatiba (see Ponta Cumuruxatiba); Bahia,
Brazil
Comunidade Colina, right bank Rio Tiquie, Mun-
icfpio Sao Gabriel da Cachoeira; Amazonas,
Brazil (0072'N, 6004'W, Patton et al., 2000)
Micoureus demerarae
Philander andersoni
Complejo Astronomico El Leoncito; San Juan,
Argentina (not located)
Thylamys pallidior
Conceicao; Roraima, Brazil (0211'N, 6057'W,
Paynter and Traylor, 1991)
Didelphis marsupialis
Conceicao do Araguaia, left bank Rio Araguaia;
Para, Brazil (0815'S, 4917'W, Paynter and
Traylor, 1991)
Caluromys philander
Conceicao do Mato Dentro; Minas Gerais, Brazil
(1901'S, 4325'W, Paynter and Traylor,
1991)
Marmosops incanus, 800 m
Ph dander frenata
Concepcion; Concepcidn, Paraguay (2325'S,
5717'W, Paynter, 1989)
Gracilinanus agilis, 175 m (2320'S, 5725'W)
Lutreolina crassicaudata
Concepcion (see Concepcion del Mar); Esquintla,
Guatamala
Concepcion; Tucuman, Argentina (2720'S,
6535'W, Paynter, 1985)
Didelphis albiventris, 400 m (2720'S,
6530'W)
Monodelphis dimidiata
Thylamys venustus, ca. 400 m
Concepcion, 2 mi E; Chiriquf, Panama (0831'N,
8237'W, Geonet)
Didelphis marsupialis
BROWN: ATLAS OF NEW WORLD MARSUPIALS
203
Concepcion del Mar (= Concepcion); Esquintla,
Guatemala (1412'N. 9118'W. Geonet)
Philander opossum
Concepcion de San Rafael: Heredia. Costa Rica
(1002'N. 8404'W, USBGN)
Micoureus alstoni. 1500 m
Concordia: Antioquia. Colombia (0603'N.
7555'W. Paynter. 1997)
Didelphis marsupialis, 1740-2030 m
Condoto. Rio Condoto. tributary of Rio San Juan:
Choco. Colombia (0506'N, 7637'W. Payn-
ter. 1997)
Marmosa robinsoni, 70 m
Philander opossum, 300 ft (= 90 m)
Contamana; Loreto. Peru (0720'S, 7501 'W. Geo-
net)
Marmosa murina, 134 m (right bank Rio Uca-
yali)
Marmosops impavidus, 134 m (Rio Ucayali)
Metachirus nudicaudatus. 500 ft (= 150 m)
(NE bank Rio Ucayali)
Micoureus regina, 134 m (right bank Rio Uca-
yali)
Contao. Palena. Rio Negro. 19.7 km N. 26.7 km
S: Los Lagos. Chile (4158'S. 7238'W, Mes-
erve et al.. 1991)
Dromiciops gliroides, 200 m
Rhyncholestes raphanurus, 200 m (19.7 km N.
11.1 km WNW of Rio Negro, and 26.7 km
S of Contao)
Contulmo: Biobio. Chile (3800'S. 7314'W. Geo-
net)
Dromiciops gliroides. All m
Coomacka (see Takama); Upper Demerara-West
Coast Berbice. Guyana
Copan; Copan. Honduras (1451'N. 8908'W.
Geonet)
Didelphis marsupialis
Copataza: Pastaza, Ecuador (0207'S. 7727'W.
Paynter. 1993)
Didelphis marsupialis, 419 m
Coray: Valle. Honduras (1339'N, 8731'W. Geo-
net)
Philander opossum
Corazon. forest NW of: Pichincha. Ecuador
(0032'S, 7839'W. Paynter. 1993)
Caenolestes fuliginosus
Cordilera Yuto. 10 km S of Camiri (see Cordillera
Yuto): Santa Cruz. Bolivia
Cordillera Balzar (see Montanas de Balzar): Man-
abi. Ecuador
Cordillera Basilio. 3.4 km S of Basilio (see Bas-
ilio); Santa Cruz. Bolivia
Cordillera Carpish. E slope; Huanuco. Peru (ca.
0940'S. 7609'W. Stephens and Traylor,
1983)
Didelphis albiventris, ca. 2500-3500 m
Cordillera de Chilla. Taraguacocha: El Oro, Ec-
uador (0332'S. 7937'W. Paynter, 1993)
Caenolestes caniventer, 2950 m
Cordillera de Colconche, near Manglaralto: Guay-
as. Ecuador (0020'S, 8020'W, Paynter,
1993)
Didelphis marsupialis
Cordillera de La Rusia (see Cordillera de Rusia):
Bocaya/Santander. Colombia
Cordillera de los Andes: Biobio. Chile (3656'S,
7122'W. USBGN)
Dromiciops gliriodes
Cordillera de Merida. S of Lago de Maracaibo:
Merida, Venezuela (ca. 0830'N. 7140'W,
Geonet)
Monodelphis adusta, 1000 m
Cordillera de Rusia (= Cordillera de La Rusia =
La Rusia = Paramo de La Rusia); BocayaV
Santander, Colombia (0559'N, 7305'W.
Paynter. 1997)
Caluromys lanatus, 4320 m
Cordilleras del Transito. near Vallenar; Atacama,
Chile (Vallenar = 2834'S, 7045'W, Geonet)
Thylamys elegans
Cordillera Yuti (= Cordilera Yuto), 10 km S of
Camiri: Santa Cruz, Bolivia (2015'S,
6330'W, Anderson. 1997)
Thylamys venustus
Cordoba, 3 mi SE of Yongas; Veracruz-Llave,
Mexico (1854'N. 9656'W. Geonet)
Philander opossum
Cornes. 30 mi above Rio Viejo; Veraguas, Pana-
ma (Rio Viejo = 0744'N. 8101'W, Geonet)
Chironectes minimus
Corn Islands (see Islas del Maiz): Zelaya. Nica-
ragua
Coro, 49 km N. 32 km W; Falcon, Venezuela
(1149'N. 6957'W. Lopez-Fuster et al..
2000)
Marmosa robinsoni
Coronel Oviedo; Caaguazii. Paraguay (2525'S,
5627'W. USBGN)
Didelphis albiventris
Coropina Kreek. near Republiek: Commewijne,
Suriname (0532'N. 5510'W. Geonet)
Metachirus nudicaudatus
Corosha: Amazonas, Peru (0550'S. 7745'W, Ste-
phens and Traylor, 1983)
Didelphis albiventris, 6900 ft (= 2150 m)
Corozal Army Reservation (see Corozal); Pana-
ma. Panama
204
FIELDIANA: ZOOLOGY
Corozal (= Corozal Army Reservation); Panama,
Panama (0858'N, 7935'W, Geonet)
Didelphis marsupialis
Corozal, Parque Nacional "El Avila"; Distrito
Federal, Venezuela (1033'N, 6657'W, Geo-
net)
Didelphis marsupialis
Marmosops fuscatus
Monodelphis palliolata
Corral Quemado Formation; Catamarca, Argenti-
na (Corral Quemado = 2708'S, 6657'W,
Paynter, 1985)
Lutreolina crassicaudata
Corrientes; Corrientes, Argentina (2758'S,
5850'W, Paynter, 1985)
Didelphis albiventris, 55 m (20 km SE of Cor-
rientes)
Lutreolina crassicaudata (left bank Rio Parana)
Cortada del Guayabo; Miranda, Venezuela
(1021'N, 6654'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni
Monodelphis palliolata
Cortes (see Puerto Cortez); Puntarenas, Costa
Rica
Corumba; Mato Grosso do Sul, Brazil (1901'S,
5739'W, Paynter and Traylor, 1991)
Didelphis albiventris, 115 m
Philander opossum (W bank Rio Paraguay)
Corupa (= Colonia Hansa = Hansa = Humboldt);
Santa Catarina, Brazil, (2626'S, 4914'W,
Paynter and Traylor, 1991)
Chironectes minimus, 62 m
Didelphis aurita
Gracilinanus microtarsus (ca. 2630'S,
5030'W)
Micoureus demerarae, 62 m
Monodelphis iheringi
Philander frenata
Cosanga, Rio Aliso; Napo, Ecuador (0034'N,
7752'W, Paynter, 1993)
Caenolestes fuliginosus
Cosiguina; Chinandega, Nicaragua (1255'N,
8730'W, Geonet)
Philander opossum
Costa da Serra; Rio Grande do Sul, Brazil (not
located)
Chironectes minimus
Costao dos Engenhos; Sao Paulo, Brazil (2441'S,
4725'W, Hershkovitz, 1997)
Philander frenata
Cotia, 12 km W of Sao Paulo; Sao Paulo, Brazil
(Sao Paulo = 2330'S, 4630'W, Paynter and
Traylor, 1991)
Philander frenata
Cotoca, 2 km SE; Santa Cruz, Bolivia (1746'S,
6258'W, Anderson, 1997)
Philander opossum
Coxim; Mato Grosso do Sul, Brazil (1830'S,
5445'W, Paynter and Traylor, 1991)
Didelphis albiventris
Crasto; Sergipe, Brazil (1125'S, 3725'W,
USBGN)
Metachirus nudicaudatus, 10 m
Crato; Ceara, Brazil (0714'S, 3923'W, Paynter
and Traylor, 1991)
Didelphis albiventris, ATI m
Gracilinanus emiliae, ATI m, 427 m
Monodelphis domestica, ATI m (Sitio Constan-
tino, Sitio Olaria)
Crique Tigre; French Guiana (0538'-0458'N,
5358'-5302'W, Geonet)
Didelphis albiventris (Petit Saut)
Didelphis marsupialis
Cristalina; Goias, Brazil (1645'S, 4736'W, Payn-
ter and Traylor, 1991)
Gracilinanus agilis, 1 1 89 m (12 km NE of
Cristalina)
Monodelphis domestica
Cristobal; Colon, Panama (0921'N, 7954'W,
Geonet)
Metachirus nudicaudatus
Philander opossum
"Crovetto Field", ca. 6 km N of Balcarce; Buenos
Aires, Argentina (3751'S, 5816'W, Mares
and Braun, 2000)
Monodelphis diminiata, 100 m
Cruz del Eje; Cordoba, Argentina (3044'S,
6448'W, Paynter, 1985)
Didelphis albiventris, 457 m
Cruzeiro (near Brasilia); Distrito Federal, Brazil
(Brazilia = 1547'S, 4755'W, Paynter and
Traylor, 1991)
Didelphis marsupialis, ca. 1 1 00 m
Cruzeiro; Sao Paulo, Brazil (2235'S, 4445'W,
Hershkovitz files)
Didelphis albiventris, 530 m
Didelphis aurita, 530 m
Cuabre (= Cubre); Limon, Costa Rica (0936'N,
8247'W, Geonet)
Micoureus alstoni (30 km S of Limon)
Cuan Creek; Bocas del Toro, Panama (0912'N,
8217'W, Geonet)
Caluromys derbianus
Cuartal Zaraza-San Juan de los Morros; Guarico,
Venezuela (0955'N, 6722'W, Perez-Hernan-
dez, 1989)
Monodelphis brevicaudata, 530 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
205
Cuarto Rio Victoria, Rt. 14, km 265; Misiones,
Argentina (not located)
Chironectes minimus
Cuba; Limon, Costa Rica (Rio Cuba = 1001'N,
8311'W, Geonet)
Micoureus alstoni
Cubre (see Cuabre); Limon, Costa Rica
Cucao, Isla Chiloe; Los Lagos, Chile (4238'S,
7407'W, Geonet)
Dromiciops gliroides
Rhyncholestes raphanurus
Cuchivano; Sucre, Venezuela (ca. 1015'N,
6355'W, Paynter, 1982)
Didelphis marsupialis, 200 m
Marmosa robinsoni, 213 m
Cuenca (= Cinincay); Azuay, Ecuador (0253'S,
7859'W, Paynter, 1993)
Didelphis albiventris, 8400 ft (= 2600 m)
Cuesta del Huron, 29 km W of Cineguillas, on
road 64; Jujuy, Argentina (2216'S, 6603'W,
Flores et al., 2000)
Thylamys pallidior, 3835 m
Cuiaba (= Angaba = Cuyaba); Mato Grosso, Bra-
zil (1535'S, 5605'W, Paynter and Traylor,
1991)
Didelphis albiventris, 219 m (4-10 km NW, 10
km N)
Monodelphis domestica, 450 m
Philander opossum (10 km N)
Culata (see La Culata); Merida, Venezuela
Culiacan; Sinaloa, Mexico (2447'N, 10723'W,
Geonet)
Marmosa canescens
Culturutuin (see Cultuurtuin, Agricultural Exper-
imental Station); Paramaribo, Suriname
Cultuurtuin (= Culturutuin), Agricultural Experi-
mental Station; Paramaribo, Suriname
(0550'N, 5511'W, Hershkovitz, 1997)
Caluromys philander
Didelphis marsupialis
Marmosa murina
Monodelphis brevicaudata
Philander opossum
Culumani (see Chulumani); La Paz, Bolivia
Cumaca (= Cummacca); St. Andrew, Trinidad
and Tobago (1042'N, 6109'W, Handley and
Gordon, 1979)
Didelphis marsupialis
Marmosops fuscatus
Cumana; Sucre, Venezuela (1028'N, 6410'W,
Geonet)
Caluromys philander, 25-30 m (21 km E of
Cumana)
Didelphis marsupialis, 0-30 m (24 km E of
Cumana)
Marmosa robinsoni, 1-50 m (16-21 km E of
Cumana)
Cumanacoa; Sucre, Venezuela (1015'N,
6355'W, Paynter, 1982)
Didelphis albiventris, 200 m
Cumana (= Cumeria); Ucayali, Peru (0951'S,
7401'W, Geonet)
Philander andersoni, 303 m
Philander opossum, 1000 ft (= 300 m)
Cumbre #1, Cerro Duida; Amazonas, Venezuela
(Cerro Duida = 0325'N, 6540'W, Paynter,
1982)
Didelphis marsupialis, 1400 m
Cumbre #13, Despilodero, Cerro Duida (see Des-
filadero, Cumbre #13); Amazonas, Venezuela
Cumbre #15, Cerro Duida; Amazonas, Venezuela
(Cerro Duida = 0325'N, 6540'W, Paynter,
1982)
Didelphis albiventris, 6600 ft (= 2060 m)
Didelphis marsupialis, 2000 m (Soma #15)
Cumbre del Cerro, Marahuaca; Amazonas, Ve-
nezuela (0334'N, 6527'W, Geonet)
Didelphis albiventris, 2550 m
Cumeria (see Cumana); Ucayali, Peru
Cummacca (see Cumaca); St. Andrew, Trinidad
and Tobago
Cumuruxatiba (see Ponta Cumuruxatiba); Bahia,
Brazil
Cupa (= Montanas de Cupa); Esmeraldas, Ecua-
dor (0026'N, 7925'W, Baker, 1974)
Didelphis marsupialis
Curacautin (near Rio Colorado); Araucania, Chile
(3826'S, 7153'W, Geonet)
Dromiciops gliroides, 900 m, 1035 m
Curitiba; Parana, Brazil (2525'S, 4918'W, Ca-
ceres and Montero-Filho, 1998)
Didelphis aurita
Curques, 24 km N of Susques on road 74; Jujuy,
Argentina (2314'S, 6623'W, Flores et al.,
2000)
Thylamys pallidior, 4100 m
Curuguaty; Canendiyu, Paraguay (2431'S,
5542'W, Paynter, 1989)
Didelphis albiventris
Caluromys lanatus (13.3 km N of Curuguaty)
Chironectes minimus (6.3 km N of Curuguaty)
Lutreolina crassicaudata
Metachirus nudicaudatus
Curundu; Panama, Panama (0859'N, 7933'W,
Geonet)
Caluromys derbianus
Didelphis marsupialis
206
FIELDIANA: ZOOLOGY
Philander opossum
Curupao; Miranda, Venezuela (1030'N, 6638'W,
Paynter, 1982)
Caluromys philander, 1 1 60 m
Marmosa robinsoni, 1130-1190 m
Marmosops fuscatus, 1160m
Micoureus demerarae, 1150-1160 m
Monodelphis palliolata, 1160m
Cusco; Cusco, Peru (1331'S, 7159'W, Stephens
and Traylor, 1983)
Didelphis albiventris, 3496 m
Cuyaba (see Cuiaba); Mato Grosso, Brazil
Cuyagua; Aragua, Venezuela (1029'N, 6742'W,
Geonet)
Chironectes minimus
Cuyambuyo; Tarija, Bolivia (2216'S, 6433'W,
Anderson, 1997)
Lutreolina crassicaudata
Thylamys venustus
Cylindro, NE of Boquete; Bocas del Toro, Pana-
ma (Boquete = 0910'N, 8216'W, Geonet)
Marmosops invictus, 1220+ m
D
Dabeiba; Antioquia, Colombia (0701'N,
7616'W, Paynter, 1997)
Didelphis marsupialis, 600 m
Dadanawa, 15 km E; Upper Takutu-Upper Esse-
quibo, Guyana (0250'N, 5931'W, Geonet)
Didelphis marsupialis, 150 m
Gracilinanus emiliae
Dagua (= Caldas); Valle del Cauca, Colombia
(0340'N, 7641'W, Paynter, 1997)
Didelphis marsupialis, 816 m
Daule; Guayas, Ecuador (0150'S, 7956'W,
Paynter, 1993)
Didelphis marsupialis, sea level
Delta del Parana; Buenos Aires, Argentina
(3343'S, 5915'W, Paynter, 1985)
Lutreolina crassicaudata
Delta del Parana (see Brazo Largo); Entre Rfos,
Argentina
Demerara River, 29 mi above Georgetown; De-
merara-Mahaica, Guyana (Georgetown =
0648'N, 5810'W, Geonet)
Didelphis marsupialis
Marmosa murina
"De Morgenstond" (= Morgenstond = Plantage
de Morgenstond), NE Paramaribo, left bank
Suriname River; Paramaribo, Suriname
(0551'N, 5508'W Geonet)
Marmosa murina
Monodelphis brevicaudata
Departamento Capital; Salta, Argentina (not lo-
cated)
Thylamys pusilla
Department of General Belgrano; Misiones, Ar-
gentina (not precisely located)
Chironectes minimus
Desaguadero; Mendoza, Argentina (3325'S,
6511'W, Paynter, 1985)
Thylamys pallidior
Deseado, General Balgano; Misiones, Argentina
(2547'S, 5403'W, Geonet)
Didelphis aurita
Desfiladero, Cumbre #13 (= Cumbre #13, Des-
pilodero = Despilodero); Amazonas, Vene-
zuela (ca. 0319'N, 6535'W, Paynter, 1982)
Didelphis albiventris, 6200 ft (= 1930 m)
Didelphis marsupialis, 1825 m
Despilodero (see Desfiladero, Cumbre #13); Ama-
zonas, Venezuela
Dique de Cadillal (see El Cadillal Dike); Tucu-
man, Argentina
Dique Escaba; Tucuman, Argentina (2740'S,
6546'W, Mares et al., 1966)
Didelphis albiventris
Diriamba; Choluteca, Nicaragua (1 151 'N,
8614'W, Geonet)
Caluromys derbianus
Didelphis marsupialis, 600 m
Philander opossum, 600 m
Divala; Chiriquf, Panama (0825'N, 8243'W,
Geonet)
Caluromys derbianus
Divisadero; Morazan, El Salvador (1336'N,
8803'W, Geonet)
Didelphis marsupialis, 750 ft (= 230 m), 1000
ft (= 300 m)
Philander opossum (2 mi N of Divisadero)
Doctor Facundo Zuvina (see Zuviria); Salta, Ar-
gentina
Dois Hermanos (see Dois Irmaos); Pernambuco,
Brazil
Dois Irmaos (= Dois Hermanos = Hermanos);
Pernambuco, Brazil (ca. 0803'S, 3554'W,
Paynter and Traylor, 1991)
Didelphis marsupialis
Marmosa murina
Metachirus nudicaudatus
Micoureus demerarae
Monodelphis americana
Monodelphis domestica
Dolavon, ca. 200 km W; Chubut, Argentina
(3439'S, 6807'W, Mares and Braun, 2000)
Thylamys pallidior
BROWN: ATLAS OF NEW WORLD MARSUPIALS
207
Dolavon, 280 km W; Chubut, Argentina (4345'S,
6857'W, Mares and Braun, 2000)
Thylamys pallidior
Dominical, 0.5 mi N; Puntarenas, Costa Rica
(0915'N, 8352'W, Geonet)
Caluromys derbianus
Don Amo [Donama], Sierra Nevada de Santa
Marta; Magdalena, Colombia (ca. 1115'N,
7358'W, Paynter, 1997)
Metachirus nudicaudatus, 1000 ft (= 300 m)
Dos Caminos (see Los Dos Caminos); Guarico,
Venezuela
Dos Ramon, Rancho Grande (see Estacion Biol-
ogica de Rancho Grande); Aragua, Venezuela
Douradoquara; Minas Gerais, Brazil (1828'S,
4735'W, Varejao and Valle, 1982)
Didelphis albiventris
Dourados; Mato Grosso do Sul, Brazil (2213'S,
5448'W, Paynter and Traylor, 1991)
Didelphis albiventris (25 km S of Dourados)
Metachirus nudicaudatus (12 km N of Doura-
dos)
Philander opossum
Dragones; Salta, Argentina (2315'S, 6321'W,
Geonet)
Didelphis albiventris
Thylamys pusilla
Dubulay Ranch; Upper Demerara-Berbice, Guy-
ana (not precisely located, but Dubulay =
0537'N, 5732'W, USBGN)
Monodelphis brevicaudata
Duefias (see San Miguel Duefias); Suchitepequez,
Guatemala
Dunoon; Demerara-Mahaica; Guyana (0626'N,
5818'W, Geonet)
Marmosa murina
Duraznillar (= El Duraznillar), Parque Nacional
Calilegua; Jujuy, Argentina (2335'-2400'S,
6430'-6500'W, Paynter, 1985)
Thylamys venustus, 2500 m
E
Ebano; San Luis Potosf, Mexico (2213'N,
9822'W, Geonet)
Didelphis marsupialis
Eirunepe (see Joao Pessoa, Amazonas, Brazil)
Ejido Santa Isabel; Tamaulipas, Mexico (23 C 14'N,
9900'W, Gardner, 1973)
Didelphis marsupialis
El Alto, Valle de Suaza; Huila, Colombia (ca.
0307'N, 7523'W, Geonet)
Didelphis marsupialis
El Arenal (see Arenal); Jujuy, Argentina
El Bracho; Tucuman, Argentina (2659'S,
6511'W, Paynter, 1985)
Thylamys pallidior, 400 m
Thylamys pusilla, 400 m
El Breal, 6 km SW of Santa Victoria (= Riva-
davia); Salta, Argentina (Santa Victoria =
2215'S, 6458'W, Paynter, 1985)
Didelphis albiventris, 2000 m
El Cadillal, Aguas Chiquitas, Sierras de Medina;
Tucuman, Argentina (2637'S, 6212'W, Ma-
res et al., 1996)
Lutreolina crassicaudata
Thylamys pallidior
Thylamys pusilla
Thylamys venustus
El Cadillal Dike (= Dique de Cadillal), 25 km
NW of San Miguel de Tucuman; Tucuman,
Argentina (2637'S, 6512'W, Mares et al.,
1966)
Didelphis albiventris
Thylamys pallidior, 700 m
Thylamys pusilla, 1 00 m
El Cajon; Tucuman, Argentina (2632'S, 6452'W,
Mares et al., 1966)
Didelphis albiventris
El Callao (see El Peru); Bolivar, Venezuela
El Callao (see Challao); Mendoza, Argentina
El Candelo, Tucuragua; Cojedas, Venezuela
(0953'N, 6839'W, Perez-Hernandez, 1989)
Monodelphis palliolata, 640 m
El Carmen, near, a campground on Highway 9;
Jujuy, Argentina (2428'S, 6521'W, Flores et
al., 2000)
Thylamys venustus, 4600 ft (= 440 m)
El Carmen, Rfo Blanco; Beni, Bolivia (1357'S,
6343'W, Hershkovitz, 1997)
Philander opossum
El Carrizo, 2 km W and 70 km S of Ciudad Vic-
toria, on Pan American Highway (see Ciudad
Victoria); Tamaulipas, Mexico
El Castillo, near Mindo; Pichincha, Ecuador
(Mindo = 0002'S, 7848'W, Paynter, 1993)
Caenolestes convelatus
El Cerro del Oropuche (= Heights of Ourepouche
= Oropuche); St. David, Trinidad and Tobago
(1046'N, 6109'W, USBGN)
Didelphis marsupialis
Marmosa robinsoni (Victoria, ca. 1015'N,
6133'W)
El Chaupi, NW of; Pichincha, Ecuador (0036'S,
7831'W, Geonet)
Caenolestes fuliginosus, 3410 m
El Chiral; El Oro, Ecuador (0338'S, 7941'W,
Paynter, 1993)
208
FIELDIANA: ZOOLOGY
Caenolestes caniventer, 5350 ft (= 1670 m)
Didelphis albiventris, 5350 ft (= 1670 m)
Philander opossum, 5350 ft (= 1670 m)
(0339'S, 7943'W)
El Colorado; Formosa, Argentina (2618'S,
5922'W, Geonet)
Didelphis albiventris
Lutreolina crassicaudata
El Consejo, 10 km NE; Aragua, Venezuela
(1014'N, 6716'W, Paynter, 1982)
Chironectes minimus, ca. 600 m
El Contento; Cordoba, Colombia (0908'N,
7532'W, Adler et al., 1997)
Didelphis marsupialis
Marmosa robinsoni
El Corazon; Pichincha, Ecuador (0032'S,
7839'W, Paynter, 1993)
Didelphis albiventris, 4191 m
"El Corte"; Maldonado, Uruguay (ca. 3440'S,
5455'W, Paynter, 1994)
Didelphis albiventris, 100 m
El Cruce; Misiones, Argentina (not located)
Monodelphis dimidiata
El Destino, Rio Cuaque (= Rio Coaque); Manabi,
Ecuador (Rio Coaque = 0001'N, 8006'W,
Paynter, 1993)
Marmosa robinsoni, 900 m
El Dividive (see El Dividivi); Trujillo, Venezuela
El Dividivi (= El Dividive); Trujillo, Venezuela
(0929'N, 7044'W, Paynter, 1982)
Philander opossum, 90 m
El Dorado; Misiones, Argentina (2624'S,
5438'W, Paynter, 1985)
Didelphis albiventris
El Dorado, 65 km SSE (also see km 121 and 125);
Bolivar, Venezuela (0644'N, 6138'W, Payn-
ter, 1982)
Marmosops pinheiroi
Monodelphis brevicaudata
El Duraznillar (see Duraznillar); Jujuy, Argentina
El Eden, Parque Nacional "El Avila"; Distrito
Federal, Venezuela (1031'N, 6652'W, Perez-
Hernandez, 1989)
Didelphis marsupialis, 1500 m
Marmosops fuscatus
Monodelphis palliolata, 1500 m
El Encantado, Hacienda El Limon; Distrito Fed-
eral, Venezuela (1029'N, 6705'W, Geonet)
Didelphis marsupialis, 1500 m
Gracilinanus marica, 1500 m
El Encino, 10 km S of Rio Sabinas; Tamaulipas,
Mexico (ca. 2240'N, 9916'W, Alvarez,
1963)
Philander opossum, 400 ft (= 125 m)
El Estor-Los Dantas; Izabal, Guatemala (El Estor
= 1532'N, 8921'W, Geonet)
Philander opossum, 100 ft (= 30 m)
El Guayabal (see Guayabal); Norte de Santander,
Colombia
El Guri (see El Manteco); Bolivar, Venezuela
El Hatillo, La Union; Miranda, Venezuela
(1022'N, 6648'W, Geonet)
Didelphis marsupialis, ca. 200 m
Marmosa robinsoni
Monodelphis palliolata
El Hato, Antisana; Pichincha, Ecuador (0000'S,
7807'W, Geonet)
Caenolestes convelatus, 4100 m
El Higo, 6.5 km NNW; Veracruz-Llave, Mexico
(2146'N, 9828'W, Geonet)
Didelphis marsupialis
El Indio, Ruta 307; Tucuman, Argentina (2702'S,
6540'W, Mares et al., 1996)
Lutreolina crassicaudata
El Junquito (= Carretera El Junquito); Distrito
Federal, Venezuela (1028'N, 6705'W, Geo-
net)
Gracilinanus marica, 1000 m
Marmosops fuscatus, 1000 m
El Manaco; Bolivar, Venezuela (0619'-0609'N,
6122'-6119'W, Handley, 1976)
Marmosa murina, 150 m
Monodelphis brevicaudata, 150 m
El Mangrullo; Chaco, Argentina (not located)
Thylamys pusilla
El Mante; Tamaulipas, Mexico (2244'N,
9858'W)
Philander opossum
El Manteco (= El Guri); Bolivar, Venezuela
(0739'N, 6250'W, Geonet)
Micoureus demerarae
Monodelphis brevicaudata
El Monolito (see Abra de Cafias); Jujuy, Argen-
tina
El Muneco; Cartago, Costa Rica (0948'N,
8355'W, Geonet)
Didelphis marsupialis, 3800 ft (= 1180 m),
4200 ft (= 1300 m)
El Naranjo (= Naranjos); San Luis Potosi, Mexico
(2232'N, 9920'W, Geonet)
Didelphis marsupialis
El Orinoco, Rio Cesar; Cesar, Colombia (10 10'N,
7324'W, Geonet)
Didelphis marsupialis
El Palmar, Rio Grande; Boh'var, Venezuela
(0801'N, 6155'W, Hershkovitz, 1997)
Philander opossum
BROWN: ATLAS OF NEW WORLD MARSUPIALS
209
El Palmar; Santa Cruz, Bolivia (1751'S,
6320'W. Anderson, 1997)
Philander opossum
El Palmar, Sierra de Santa Barbara, 4 Km E; Ju-
juy. Argentina (ca. 2004'S, 6434'W, Mares
and Braun, 2000)
Thylamys vemistus
El Palmar Road, Ibanez: Santa Cruz, Bolivia
(1748'S, 6308'W, Anderson. 1997)
Marmosops dorothea
Philander opossum, 500 m
El Panorama. Rio Aurare; Zulia. Venezuela (ca.
1040'N, 7125'W, Paynter, 1982)
Didelphis marsupialis, sea level
El Papayo, 7 km SE of Puebloviejo; Los Ribs.
Ecuador (Puebloviejo = 0134'S, 7930'W.
Paynter, 1993)
Philander opossum
El Papayo, Rio Saija; Cauca, Colombia (Rio Saija
= 0252'N, 7741'W, Paynter. 1997)
Chironectes minimus, 500 m
El Pauji; Bolivar, Venezuela (0423'N, 6129'W.
Perez-Hernandez. 1989)
Didelphis albiventris, 900 m
El Peru (= El Callao = Peru Mines): Bolivar, Ve-
nezuela (0719'N, 6149'W, Paynter. 1982)
Didelphis marsupialis, ca. 200 m
El Poblado: Antioquia. Colombia (0613'N,
7534'W, Paynter, 1997)
Didelphis marsupialis
El Pool. Bluefields: Zelaya. Nicaragua (1202'N.
8344'W. USBGN)
Metachirus nudicaudatus
El Porvenir. 5 km N: Beni, Bolivia (1451'S.
6621'W, Anderson, 1997)
Mieoureus constantiae
El Pueblito (= Pueblo Viejo). N slope of Sierra
Nevada, Santa Marta; La Guajira. Colombia
(1059'N, 7327'W, Paynter, 1997)
Caluromys lanatus, 4000 ft (= 1250 m)
Didelphis marsupialis, 610 m
Marmosa robinsoni
El Quebrachal; Salta. Argentina (2517'S.
6404'W, Paynter. 1985)
Didelphis albiventris, 339 m
El Real: Chiapas. Mexico (1706'N, 9155'W,
Geonet)
Didelphis marsupialis, 1800 ft (= 560 m)
El Real (= El Real de Santa Maria, Cerro Mali):
Darien. Panama (0808'N, 7743'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Marmosa robinsoni
El Real. Cerro Mali (see El Real); Darien. Panama
El Real de Santa Maria, Cerro Mali (see Real);
Darien, Panama
El Recreo; Zelaya, Nicaragua (1210'N. 8419'W,
Geonet)
Caluromys derbianus
Chironectes minimus, 25 m (S side Rio Mico)
Didelphis marsupialis, 25 m
Philander opossum, 20 m
El Rincon (= Rincon); San Luis, Argentina
(3221'S, 6459'W. Paynter, 1985)
Didelphis albiventris
El Risa de Oro, near Cocahuatel (see Risa de
Oro): Chiapas. Mexico
El Roble: Cundinamarca, Colombia (ca. 0423'N,
7419'W. Paynter, 1997)
Didelphis albiventris, 2475 m
Marmosops impavidus, 8000 ft (= 2500 m)
El Roble; Quindio, Colombia (0441'N, 7536'W,
Paynter. 1997)
Marmosops fuscatus, 2175 m
El Rosario: Zulia, Venezuela (0909'N. 7236'W,
Handley. 1976)
Caluromys lanatus
Didelphis marsupialis, 24-125 m
Marmosa murina, 24-125 m
Philander opossum, 24-125 m
El Salado; Magdalena, Colombia (1028'N,
7441'W, Geonet)
Didelphis marsupialis, 430 m
El Salto: San Luis Potosi, Mexico (2236'N,
9924'W. Gardner, 1973)
Didelphis marsupialis (Rio Naranja)
Philander opossum, 2000 ft (= 625 m) (also El
Salto Falls)
El Sance Peralta; Costa Rica (not located)
Philander opossum
El Simbolar: Jujuy, Argentina (2411'S, 6428'W,
Flores et al., 2000)
Thylamys venustus
El Soche; Cundinamarca. Colombia (0432'N,
7420'W, Geonet)
Caenolestes fuliginosus, 2700 m
El Socorro, cerca de; Guarico. Venezuela
(0859'N, 6544'W, Paynter, 1982)
Didelphis marsupialis
El Sosneado. 8 km NW; Mendoza, Argentina
(3500'S. 6938'W. Mares and Braun, 2000)
Thylamys pallidior
El Talisman (= Talisman); Chiapas, Mexico
(1542'N. 9242'W. Geonet)
Philander opossum. 1 27 m
El Tirol. NNE of Encarnacion: Itapua. Paraguay
(Encarnacion = 2720'S. 5554'W, Paynter,
1989)
210
FIELDIANA: ZOOLOGY
Didelphis marsupialis, 230 m
El Tocuyo; Lara, Venezuela (0947'N, 6948'W,
Paynter, 1982)
Didelphis marsupialis, 617 m
El Trapiche; Beni, Bolivia (1448'S, 6618'W, An-
derson, 1997)
Didelphis marsupialis
Metachirus nudicaudatus
El Triumfo (see El Triunfo); Cundinamarca, Co-
lombia
El Triunfo (= El Triumfo); Cundinamarca, Co-
lombia (ca. 0432'N, 7430'W, Paynter, 1997)
Marmosa robinsoni, 600 ft (= 180 m)
El Trompillo; Carabobo, Venezuela (1004'N,
6746'W, Paynter, 1982)
Marmosa robinsoni, ca. 500 m
El Tukuko (see Tukoko); Zulia, Venezuela
El Valle; Code, Panama (0836'N, 8008'W, Geo-
net)
Didelphis marsupialis
Metachirus nudicaudatus (6 mi E of El Valle)
Philander opossum
El Valle, Caracas; Distrito Federal, Venezuela
(1027'N, 6655'W, Paynter, 1982)
Caluromys lanatus
El Vertigo, Zongo Valley; La Paz, Bolivia
(1605'S, 6802'W, Anderson, 1997)
Didelphis marsupialis
El Volcan; Chiriqui, Panama (0846'N, 8240'W,
Geonet)
Philander opossum (14.5 km NW of Finca San-
ta Clara, 0846'N, 8238'W)
El Zapote, 7 mi S of Sabana Grande; Francisco
Morazan, Honduras (1345'N, 8725'W, Geo-
net)
Didelphis marsupialis
Embalse Huechun (= Huechun, Embalse); Santi-
ago, Chile (3304'S, 7048'W, Paynter, 1988)
Thylamys elegans, ca. 560 m
Embalase Rio Moron; Carabobo, Venezuela
(1027'N, 6800'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Embocada (= Estancia La Cabana); Beni, Bolivia
(1503'S, 6658'W, Anderson, 1997)
Didelphis marsupialis, less than 600 m
Marmosops impavidus, less than 600 m
Marmosops noctivagus, less than 600 m
Empire; Panama, Panama (0904'N, 7940'W,
Geonet)
Didelphis marsupialis
Philander opossum
Encarnacion; Itapiia, Paraguay (2720'S, 5554'W,
Paynter, 1989)
Didelphis albiventris
Encontrados; Zulia, Venezuela (0903'N,
7214'W, Paynter, 1982)
Philander opossum (60 km WNW of Encontra-
dos)
Engenheiro Reeve (= Rive); Espirito Santo, Bra-
zil (2046'S, 4128'W, Paynter and Traylor,
1991)
Philander frenata, 600 m
Ensenada Cuaranta; Sucre, Venezuela (1038'N,
6215'W, Handley, 1976)
Didelphis marsupialis, 1-4 m
Entre Rios; Argentina (not located)
Chironectes minimus
Entre Rios; La Paz, Bolivia (1539'S, 6726'W,
Anderson, 1997)
Didelphis marsupialis
Entre Rios, 5 km NNW; Tarija, Bolivia (2129'S,
6412'W, Anderson, 1997)
Micoureus constantiae
Thylamys pusilla (2132'S, 6412'W)
Thylamys venustus
Erquis; Tarija, Bolivia (2128'S, 6448'W, Ander-
son, 1997)
Thylamys venustus
Erval (= Serra do Herval); Rio Grande do Sul,
Brazil (3025'S, 5155'W, Geonet)
Chironectes minimus
Escarcega; Campeche, Mexico (1837'N,
9043'W, Geonet)
Caluromys derbianus (near Escarcega)
Didelphis marsupialis (7.5 km W of Escarcega)
Philander opossum (65 km S, 128 km of Es-
carcega)
Escazii; San Jose, Costa Rica (0955'N, 8408'W,
Geonet)
Micoureus alstoni
Escola Agricola de Exu; Pernambuco, Brazil (Exu
= 0731'S, 3943'W, Paynter and Traylor,
1991)
Monodelphis domestica
Thylamys pusilla (.7 km S of Exu)
Escola Agropecuaria (= Escuela Agropecuaria);
Concepcion, Paraguay (2321'S, 5723'W,
Palma, 1995)
Thylamys macrura
Escopeta Camp, 23 km NNE of San Felix; Chi-
riqui, Panama (San Felix = 0817'N,
8152'W, Geonet)
Chironectes minimus
Escuela Agropecuaria (see Escola Agropecuaria);
Concepcion, Paraguay
Escuela Normal, Monteros; Tucuman, Argentina
(2647'S, 6533'W, Mares et al., 1966)
Didelphis albiventris
BROWN: ATLAS OF NEW WORLD MARSUPIALS
211
Esmeralda; Amazonas, Venezuela (0310'N,
6533'W, Paynter, 1982)
Caluromys lanatus, 145 m (48 km NW of Es-
meralda, 0311'N, 6533'W)
Didelphis marsupialis, 143 m
Metachirus nudicaudatus (Cerro Duida)
Micoureus demerarae, 325 ft (= 95 m) (foot of
Cerro Duida, upper Rio Orinoco)
Monodelphis brevicaudata, 135 m
Philander andersoni, 135 m
Esmeraldas; Esmeraldas, Ecuador (0059'N,
7942'W, Paynter, 1993)
Didelphis marsupialis, sea level
Philander opossum, near sea level
Espiritu; Beni, Bolivia (1408'S, 6621'W, Ander-
son, 1997)
Philander opossum
Esquinapa; Sinaloa, Mexico (2250'N, 10547'W,
Gardner, 1973)
Marmosa canescens
Esquipulas; Matagalpa, Nicaragua (1240'N,
8547'W, Geonet)
Caluromys derbianus
Philander opossum, 420 m (ca. 1200'N,
8600'W)
Essequibo (see Oko Mts.); Cuyuni-Mazaruni,
Guyana
Estacao Biologica de Boraceia (see also Bora-
ceia), Biological Station in Salesopolis; Sao
Paulo, Brazil (2339'S, 4554'W, Heyer et al.,
1990)
Didelphis aurita, ca. 737 m, 850 m
Marmosops incanus, 820 m
Marmosops paulensis, 850 m
Philander frenata, 850 m
Estacao Biologica de Caratinga, Fazenda Montes
Claros, 45 km E of Caratinga; Minas Gerais,
Brazil (Caratinga = 1950'S, 4150'W, Mus-
trangi and Patton, 1997)
Marmosops incanus, 500 m
Estacao Biologica de Santa Lucia, Santa Tereza;
Espirito Santo, Brazil (1950'S, 4022'W,
Mustrangi and Patton, 1997)
Marmosops incanus
Metachirus nudicaudatus
Estacao de Pesquisas de Peti, Sao Goncalo do Rio
Abaixo (see Ciudade de Sao Goncalo do Rio
Abaixo); Minas Gerais, Brazil
Estacao Ecologica do Tapacura (see Sao Lourenco
da Mata); Pernambuco, Brazil
Estacion Biologica de Calabozo; Guarico, Vene-
zuela (0852'N, 6723'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
Estacion Biologica de La Selva (= Finca La Selva
= La Selva Biological Reserve), 3 km S of
Puerto Viejo; Heredia, Costa Rica (Puerto
Viejo = 1028'N, 8401'W, Geonet)
Caluromys derbianus, 70 m
Estacion Biologica de los Llanos, 9 km SE of Cal-
abozo; Guarico, Venezuela (0856'N,
6725'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 100 m
Marmosa robinsoni, 100-115 m (0852'N,
6723'W)
Monodelphis brevicaudata, 100 m (0852'N,
6723'W)
Monodelphis palliolata
Estacion Biologica de Rancho Grande (= Rancho
Grande = Rancho Grande Biological Station);
Aragua, Venezuela (1021'N, 6740'W, Perez-
Hernandez, 1989)
Caluromys philander, 910-1170 m
Chironectes minimus, 910-1170 m (also
Parque Nacional de Rancho Grande at
1022'N, 6741'W)
Didelphis marsupialis, 910-1170 m
Gracilinanus marica, 1 100 m (also Dos Ra-
mon, Guacamaya, La Quinta, Pica Periquito,
El Portachuela at 1021'N, 6741'W, Guami-
tas, Palo de Vaca)
Marmosops fuscatus, 910-1170 m, 1050 m
(1022'N, 6741'W) (also Dos Ramon, Gua-
camaya, La Quinta, Pica Periquito, El Porta-
chuela at 1021'N, 6741'W, Guamitas)
Micoureus demerarae, 910-1170 m, 1100 m
(also Dos Ramon, Guacamaya, La Quinta,
Pica Periquito, El Portachuela at 1021'N,
6741'W, Guamitas)
Monodelphis brevicaudata
Monodelphis palliolata, 910-1170 m
Philander opossum
Estacion Biologica R. Schreiter, San Javier; Tu-
cuman, Argentina (not located)
Thylamys venustus
Estacion de Kasmera, Rio Yasa; Zulia, Venezuela
(1005'N, 7245'W, Perez-Hernandez, 1989)
Monodelphis palliolata
Estacion de Piscicultura, El Limon; Aragua, Ve-
nezuela (1019'N, 6738'W, Perez-Hernandez,
1989)
Didelphis marsupialis
Estacion Experimental del Instituto Nacional de
Tecnologia Agropecuaria (= INTA areas), ca.
15 km NW of Balcarce; Buenos Aires, Ar-
gentina (3751'S, 5816'W, Mares and Braun,
2000)
Monodelphis dimidiata, ca. 100-150 m
212
FIELDIANA: ZOOLOGY
Estacion Experimental de Rio Negro; Miranda,
Venezuela (1020'N, 6015'W, Perez-Hernan-
dez, 1989)
Micoureus demerarae, 50-70 m
Estacion Pailon; Santa Cruz, Bolivia (1739'S,
6245'W, Anderson, 1997)
Marmosops dorothea
Estacion Tambo, Nuevo y Ruta 188, km 60;
Buenos Aires, Argentina (3358'S, 6035'W,
USBGN)
Monodelphis dimidiata
Estacion Vipos (see Vfpos); Tucuman, Argentina
Estacion Yuruma (see Yuruma); Potosi, Bolivia
Estanaque de Curupao; Miranda, Venezuela
(1030'N, 6638'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Estancia Barranquita (see San Joaquin); Beni, Bo-
livia
Estancia Bolivar; Tarija, Bolivia (2138'S,
6234'W, Anderson, 1997)
Monodelphis domestica
Thylamys pusilla (also at 2138'N, 6237'W)
Estancia Cachuela Esperanza; Santa Cruz, Bolivia
(1647'S, 6314'W, Anderson, 1997)
Gracilinanua agdis
Philander opossum
"Estancia Concordia," 15 mi SW of Dolores; So-
riano, Uruguay (Dolores = 3333'S, 5813'W,
Paynter, 1994)
Didelphis albiventris, ca. 100 m
Estancia Coropa, Goya; Corrientes, Argentina
(Goya = 2908'S, 5916'W, Paynter, 1985)
Thylamys pusilla
Estancia Cuevas; Santa Cruz, Bolivia (1811'S,
6344'W, Anderson, 1997)
Micoureus constantiae
Estancia Guampacha; Santiago del Estero, Argen-
tina (2759'S, 6445'W, Flores et al., 2000)
Thylamys pusilla
Estancia Isibobo; Santa Cruz, Bolivia (1931'S,
6336'W, Anderson, 1997)
Monodelphis domestica
Estancia Ita Caabo, Mercedes; Corrientes, Argen-
tina (Mercedes = 2912'S, 5805'W, Geonet)
Lutreolina crassicaudata
Estancia Jahue (see Quine, Estancia Jahue); Santa
Cruz, Bolivia
Estancia Juan de Zalazar, left bank, 2 km N of
Rio Verde, near Fortin Juan Salazar in Presi-
dente Hayes; Boqueron, Paraguay (Fortin
Juan Salazar = 2338'S, 5918'W, Paynter,
1989)
Didelphis albiventris, 200 m
BR(
Estancia La Cabana, 1 km E of La Embocada (see
Embocada); Beni, Bolivia
Estancia La Geraldina; Santa Fe, Santiago del Es-
tero, Argentina (ca. 3005'S, 6200'W Payn-
ter, 1985)
Didelphis albiventris
Estancia La Germania; Santa Fe, Argentina
(3233'S, 6124'W, Paynter, 1985)
Didelphis albiventris
Estancia La Germania (see Nueva Germania); San
Pedro, Paraguay
Estancia Laja (see Quine, Estancia Laja); Santa
Cruz, Bolivia
Estancia La Selva, Bogota (see La Selva); Cun-
dinamarca, Colombia
Estancia Leoncito; San Juan, Argentina (3148'S,
6920'W, USBGN)
Thylamys pallidior
Estancia Linda Vista, NNE Formosa (see Riacho
Pilaga); Formosa, Argentina
Estancia Los Alpes (see La Lata); Colonia, Uru-
guay
Estancia Los Ingleses (= Los Ingleses = Los Yn-
gleses); Buenos Aires, Argentina (3631'S,
5635'W, Paynter, 1985)
Didelphis albiventris, sea level
Lutreolina crassicaudata, sea level
Estancia Los Manantiales, Languifieo (= Los
Manantiales); Chubut, Argentina (4316'S,
6951'W, Geonet)
Lestodelphys halli
Estancia La Madrugada, near Puerto Deseado;
Santa Cruz, Argentina (4706'S, 6629'W,
Mares and Braun, 2000)
Lestodelphys halli
Estancia Machijada, near Puetro Deseado ( =
Comodoro Rivadavia); Chubut, Argentina
(4552'S, 6730'W, Thomas, 1929)
Lestodelphys halli
Estancia Marfa Sofia; Rio Negro, Argentina
(4037'S, 7009'W, Mares and Braun, 2000)
Thylamys pallidior
Estancia Melo, 6 km SE; Cerro Largo, Uruguay
(Melo = 3222'S, 541 l'W, Paynter, 1994)
Didelphis albiventris
Estancia San Ignacio, 24 km NNW of Carayao;
Caaguazii, Paraguay (Carayao = 2510'S,
5626'W, Paynter, 1989)
Didelphis albiventris
Lutreolina crassicaudata
Estancia Tapikiole (= Tapikiole = La Urbana =
La Urbana Vieja), 30 km S of Rio Pilcomayo;
Formosa, Argentina (2501'S, 5919'W, Payn-
ter, 1985)
IOWN: ATLAS OF NEW WORLD MARSUPIALS
213
Didelphis albiventris, 100 m
Metachirus nudicaudatus
Thylamys pusilla, ca. 100 m
Estancia Tehuel Malal; Neuquen, Argentina (ca.
4100'S, 7115'W, Paynter, 1985)
Lestodelphys halli
Estancia Yutiole, 20 km S of San Joaquin; Beni,
Bolivia (1305'S, 6448'W, Anderson, 1997)
Caluromys lanatus
Didelphis marsupialis
Estero de Chirigiiare; Portuguesa, Venezuela
(0833'N, 6842'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Esteros, Rio Pilcomayo; Boqueron, Paraguay
(2347'S, 61 o 01'W, Paynter, 1989)
Thylamys pusilla
Esteros; Santiago del Estero, Argentina (2637'S,
6339'W, Geonet)
Thylamys pallidior
Thylamys pusilla
Estrada Ferro de Braganca; Para, Brazil (Bragana
= 0103'S, 4646'W, Paynter and Traylor,
1991)
Didelphis marsupialis
Estrada Picarreira, rigth bank Rio Canaburi,
Parque Nacional do Pica da Neblina; Ama-
zonas, Brazil (0036'N, 6605'W, Patton, pers.
comm.)
Didelphis marsupialis
Eten; Lambayeque, Peru (0654'S, 7952'W, Ste-
phens and Traylor, 1983)
Didelphis marsupialis, 5 m
Exaltacion, W bank Rio Mamore; Beni, Bolivia
(1316'S, 6515'W, Paynter, 1992)
Didelphis marsupialis (8 km N of Exaltacion)
Philander opossum (8 km N of Exaltacion)
Explorer's Inn Reserve; Madre de Dios, Peru
(1250'S, 6917'W, Emmons et al., 1994)
Caluromys lanatus, 190-270 m (E bank Rio
Tambopata)
Marmosops noctivagus, 270-290 m (E bank
Rio Tambopata)
Metachirus nudicaudatus, 270-290 m (E bank
Rio Tambopata)
Micoureus regina, 270-290 m
Philander opossum, 270-290 m (E bank Rio
Tambopata)
Exu; Pernambuco, Brazil (0731'S, 3943'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 500 m
Monodelphis domestica, ca. 500 m
Thylamys pusilla, ca. 500 m
Fabiao, Itacarambi; Minas Gerais, Brazil
(1508'S, 4414'W, Geonet)
Marmosops incanus
Fabrica de Papel, N of Cuyambuyo; Tarija, Boliv-
ia (not located)
Glironia venusta, 980 m
Faldas Iliniiza; Pichincha, Ecuador (0040'S,
7842'W, Paynter, 1993)
Caenolestes fuliginosus
Faldas Occidentales del Pichincha; Pichincha, Ec-
uador (0010'S, 7833'W, Geonet)
Caenolestes fuliginosus
Faro (= Sao Jose); Para, Brazil (0211'S,
5644'W, Paynter and Traylor, 1991)
Didelphis marsupialis, ca. 50 m
Monodelphis brevicaudata
Fazenda Agua Limpa; Distrito Federal, Brazil
(1557'S, 4756'W, Paynter and Traylor, 1991)
Gracilinanus agilis, 1000 m
Philander frenata (gallery forest)
Fazenda Almada, Ilheus (= Sao Jose da Lagoa);
Bahia, Brazil (1438'S, 3912'W, Hershkovitz
files)
Didelphis aurita, sea level
Metachirus nudicaudatus
Fazenda Almeida, Rio do Braco, Ilheus; Bahia,
Brazil (1449'S, 3902'W, Mares et al., 1981)
Micoureus demerarae
Fazenda Batente, 5.5 km SE of Exu; Pernambuco,
Brazil (Exu = 0731'S, 3943'W, Paynter and
Traylor, 1991)
Monodelphis domestica
Fazenda Beijo Grande (see Itabuna); Bahia, Brazil
Fazenda Boa Fe, 16 km NE of Teresopolis; Rio
de Janeiro, Brazil (Boa Fe = 2222'S,
4153'W, Geonet)
Gracilinanus microtarsus
Philander frenata
Fazenda Boa Vista, Teresopolis (see Teresopolis);
Rio de Janeiro, Brazil
Fazenda Cacua, Serrinha (see Serrinha); Bahia,
Brazil
Fazenda Caianinha, Pesqueira; Pernambuco, Bra-
zil (Pesqueira = 0822'S, 3642'W, USBGN)
Monodelphis domestica
Fazenda Capoeira do Rosario, Feira; Bahia, Brazil
(Feira = 1215'S, 3857'W, USBGN)
Monodelphis domestica
Fazenda Capricornio, Ubatuba; Sao Paulo, Brazil
(2325'S, 4505'W, Paynter and Traylor, 1991)
Metachirus nudicaudatus, 150 m
214
FIELDIANA: ZOOLOGY
Fazenda Cardosa, NE of Caparao; Minas Gerais,
Brazil (2222'S, 4148'W, Hershkovitz, 1997)
Philander frenata, 1018 m (Serra de Caparao)
Fazenda de Cascata (see Ciudade Sao Goncalo
Rio Abaixo); Minas Gerais, Brazil
Fazenda Cayumba, Feira; Bahia, Brazil (Feira =
1215'S, 3857'W, USBGN)
Monodelphis domestica
Fazenda da Floresta, Rio Matipo (= Alvarenga);
Minas Gerais, Brazil (1953'S, 4233'W,
Paynter and Traylor, 1991)
Philander frenata
Fazenda da Toca, Ilhabela (see Ilha de Sao Se-
bastiao); Sao Paulo, Brazil
Fazenda Esmeralda; Minas Gerais, Brazil
(1920'S, 4250'W, Hershkovitz, 1997)
Gracilinanus agilis (Rio Casca)
Marmosops incanus
Metachirus nudicaudatus
Philander frenata (Rio Casca)
Fazenda Guarani, 1.9 km N of Exu; Pernambuco,
Brazil (Exu = 0731'S, 3943'W, Paynter and
Traylor, 1991)
Thylamys pusilla
Fazenda Guaricana, Guaratuba, Morretes; Parana,
Brazil (2545'S, 4855'W, Paynter and Tray-
lor, 1991)
Didelphis aurita
Fazenda Intervales, Base Saibadela, Sete Barros
in the Serra de Parnapiacaba (= Base de Car-
mo = Capao Bonito = Intervales); Sao Paulo,
Brazil (2420'S, 4825'W, Mustrangi and Pat-
ton, 1997)
Didelphis aurita, 100 m
Marmosops incanus, 700 m (Base de Carmo,
2423'S, 4755'W)
Marmosops paulensis, 700 m (2423'S,
4755'W)
Metachirus nudicaudatus, 100 m
Micoureus demerarae
Philander frenata, 700 m (2423'S, 4855'W)
Fazenda Jacu, Feira; Bahia, Brazil (Jacu =
1219'S, 3836'W, Geonet)
Monodelphis domestica
Fazenda Montes Claros; Minas Gerais, Brazil (ca.
1900'S, 4100'W, da Fonseca et al., 1988)
Gracilinanus agilis
Marmosops incanus, 500 m
Metachirus nudicaudatus (ca. 1925'S,
4235'W)
Monodelphis americana (ca. 1920'S,
4100'W)
Philander frenata (ca. 1925'S, 4235'W)
Fazenda Nazareth (see Ilha Mexiana); Para, Brazil
Fazenda Nhumirim, 150 km SE of Corumba;
Mato Grosso do Sul, Brazil (Corumba =
1901'S, 5739'W, Paynter and Traylor, 1991)
Monodelphis domestica
Fazenda Oiteiro, Serrinha (see Serrinha); Bahia,
Brazil
Fazenda Olho da Agua, 2 km N of Valenca do
Piauf; Piauf, Brazil (0624'S, 4145'W,
USBGN)
Monodelphis domestica
Fazenda Pinheira, 1.5 km SW of Exu; Pernam-
buco, Brazil (Exu = 0731'S, 3743'W, Payn-
ter and Traylor, 1991)
Monodelphis domestica
Fazenda Pirataquisse, Ilheus; Bahia, Brazil (Ilheus
= 1449'S, 3902'W, Paynter and Traylor,
1991)
Didelphis aurita
Fazenda Saco, 6.6 km N of Serra Talhada; Per-
nambuco, Brazil (Serra Talhada = 0759'S,
3813'W, USBGN)
Monodelphis domestica
Fazenda Salgado, Feira; Bahia, Brazil (Feira =
1215'S, 3857'W, USBGN)
Monodelphis domestica
Fazenda Santa Fe (see Flora); Acre, Brazil
Fazenda Santa Terezinha, 33 km NE of Linhares;
Espirito Santo, Brazil (Linhares = 1925'S,
4004'W, Paynter and Traylor, 1991)
Micoureus demerarae
Fazenda Sororoca, Pesqueira; Pernambuco, Brazil
(Pesqueira = 0822'S, 3642'W, USBGN)
Monodelphis domestica
Fazenda Tesa, Soure, Ilha de Marajo; Para, Brazil
(Soure = 0044'S, 4831'W, Paynter and
Traylor, 1991)
Monodelphis maraxina
Fazenda Umburana, Serrinha (see Serrinha); Ba-
hia, Brazil
Feira (see Feira de Santana); Bahia, Brazil
Feira de Santana (= Feira); Bahia, Brazil
(1215'S, 3857'W, USBGN)
Didelphis albiventris
Monodelphis domestica
Ferrovia Amapa, 192 km from Serra do Navio;
Amapa, Brazil (not located)
Philander opossum
Fervedouro; Minas Gerais, Brazil (2043'S,
4229'W, Mustrangi and Patton, 1997)
Marmosops incanus, 1300 m
Philander frenata
Finca Amayo; Rivas, Nicaragua (Rivas =
1126'N, 8550'W, Geonet)
Caluromys derbianus (13 km E of Rivas)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
215
Philander opossum, 40 m (SE of Rivas)
Finca America, Managua; Managua, Nicaragua
(Managua = 1209'N, 8617'W, Geonet)
Caluromys derbianus
Philander opossum
Finca "Bengala," Carretera Salento-Cocora;
Quindio, Colombia (Salento = 0438'N,
7534'W, Paynter and Traylor, 1981)
Caenolestes fuliginosus, 3250 m
Finca Carolina, 10 mi S of Tumbador; San Mar-
cos, Guatemala (1450'N, 9154'W, Geonet)
Philander opossum, 3500 ft (= 1090 m)
Finca Cipres, 9 mi from Mazatenango; Suchite-
pequez, Guatemala (Mazatenango =
1432'N, 9130'W, Geonet)
Philander opossum, 2000 ft (= 625 m)
Finca El Buque, Villavicencio; Meta. Colombia
(Villavicencio = 0409'N, 7337'W, Paynter,
1997)
Didelphis marsupialis, 500 m
Finca El Capricho, Villavicencio; Meta. Colombia
(Villavicencio = 0409'N, 7337'W, Paynter,
1997)
Didelphis marsupialis, 300 m
Philander opossum, 300 m (38 km E of Villa-
vicencio, 0409'N, 7316'W)
Finca El Jagiiel, 45 km W on road 304, on way
to Requelme; Tucuman, Argentina (2628'S,
6448'W, Flores et al, 2000)
Thy I amy s venustus
Finca El Progresso; Santa Rosa, Guatemala
(1405'N, 9021'W, Geonet)
Didelphis marsupialis
Philander opossum
Finca El Salto, 1 mi E of Esquintla; Esquintla,
Guatemala (1408'N, 9045'W, Geonet)
Philander opossum
Finca El Soche, 15 km W of Soacha; Cundina-
marca, Colombia (0435'N, 7413'W, Patter-
son and Albuja, 1996)
Caenolestes fuliginosus, 2630 m, 2800 m
Finca Germania, 24 km NE of Huixtla; Chiapas,
Mexico (Huixtla = 1509'N, 9228'W, Geo-
net)
Didelphis marsupialis
Finca Hollanda, near Paramo de Chinche; Valle
del Cauca, Colombia (Paramo de Chinche =
0337'N, 7614'W, Geonet)
Didelphis albiventris, 2700 m
Finca Jimenez, 0.5 mi E; Guanacaste, Costa Rica
(not located)
Caluromys derbianus
Philander opossum, 30 m
Finca La Esperanza, 45 km N of Huixtla; Chiapas,
Mexico (Huixtla = 1509'N, 9228'W, Geo-
net)
Didelphis marsupialis
Finca La Lola; Limon, Costa Rica (not located)
Caluromys derbianus, 50 m
Finca La Selva (see Estacion Biologica de La Sel-
va), 3 mi S of Puerto Viejo; Heredia, Costa
Rica
Finca Ligia, Parrita; Puntarenas, Costa Rica (Par-
rita = 0931'N, 8419'W, Geonet)
Didelphis marsupialis
Finca Lornessa, 2 km NW of Santa Ana; San
Jose, Costa Rica (Santa Ana = 0956'N,
8411'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Finca Ocuilapa, Tonala; Chiapas, Mexico (Tonala
= 1605'N, 9345'W, Geonet)
Didelphis marsupialis (10 km SE of Tonala)
Philander opossum (10 km S of Tonala)
Finca Pacifica, 3 mi N of Cafias; Guanacaste, Cos-
ta Rica (Canas = 1026'N, 8506'W,
USBGN)
Chironectes minimus
Finca Prusia, cerca; Chiapas, Mexico (1544'N,
9244'W, Gardner, 1973)
Didelphis marsupialis
Finca Rincon Santo, 6 km from Carretera Salento-
Cocora, Municipio Salento; Quindio, Colom-
bia (0438'N, 7534'W, Patterson and Albuja,
1996)
Caenolestes fuliginosus, 2750 m, 3250 m
Finca Rio Frio, Morales; Izabal, Guatemala (Mo-
rales = 1541'N, 8855'W, Geonet)
Chironectes minimus
Finca Santa Cecilia, 6.5 km S of Guanacoste; Gra-
nada, Nicaragua (Guanacoste = 1152'N,
8615'W, USBGN)
Caluromys derbianus
Finca Santa Clara, El Volcan; Chiriqui, Panama
(El Volcan = 0846'N, 8238'W, Geonet)
Chironectes minimus
Finca Selache, ca. 6 mi S of Cipres; Suchitepe-
quez, Guatemala (Cipres = ca. 1438'N,
9129'W, Gardner, 1973)
Didelphis marsupialis
Finca Sepacuite, 40 mi E of Coban (1529'N,
9032'W) and S of Catabon (1534'N,
8949'W); Alta Verapaz, Guatemala (ca.
1540'N, 8950'W, Geonet)
Philander opossum, 3200 ft (= 970 m)
Finca Tepeyac, near Matagalpa; Matagalpa, Nic-
aragua (Matagalpa = 1255'N, 8555'W,
Geonet)
216
FIELDIANA: ZOOLOGY
Didelphis marsupialis
Finca Yakulfca, Angosto del Rio Pescado; Salta,
Argentina (2233'S, 6432'W, Flores et al.,
2000)
Thylamys venustus
Finisanti, right bank Saramacca River, opposite
Loksie Hattie; Brokopondo, Suriname
(0508'N, 5529'W, Pine, 1981)
Marmosa murina
Marmosops parvidens
Monodelphis brevicaudata
First Falls on Cuyuni River; Cuyuni-Mazaruni,
Guyana (Cuyuni River = 0623'N, 5841'W,
Stephens and Tray lor, 1985)
Monodelphis brevicaudata
Fish Creek, Cerro Duida; Amazonas, Venezuela
(Cerro Duida = 0325'N, 6540'W, Hershko-
vitz files)
Metachirus nudicaudatus, 109-350 ft (= 30-
110 m)
Flexal (= Frechal), Rio Surumu; Roraima, Brazil
(0350'N, 6032'W, Paynter and Traylor,
1991)
Didelphis marsupialis
Flexal, 1 2 km N, W bank Rio Tapajos; Para, Bra-
zil (0534'S, 5713'W, Pine and Handley,
1984)
Monodelphis emiliae
Flor de Oro, left bank Rio Itenez, Parque Noel
Kempff Mercado; Santa Cruz, Bolivia
(1535'S, 6034'W, Anderson, 1997)
Micoureus demerarae
Flor do Prado, near Quatipuru; Para, Brazil (Qua-
tipuru = 0052'S, 4659'W, Paynter and Tray-
lor, 1991)
Philander opossum
Flora (= Fazenda Santa Fe); Acre, Brazil
(0836'S, 7251'W, Patton et al., 2000)
Caluromys lanatus
Metachirus nudicaudatus
Philander opossum
Floresta da Tijuca and Silvestre; Rio de Janeiro,
Brazil (ca. 2256'S, 4317'W, Paynter and
Traylor, 1991)
Metachirus nudicaudatus
Marmosops incanus
Philander frenata
Floresta Nacional Tapirape-Aquiri, Municipio de
Maraba; Para, Brazil (0548'S, 5030' W, Pat-
ton et al., 2000)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
Florestal; Minas Gerais, Brazil (1956'S, 4427'W,
Varejao and Valle, 1982)
Didelphis albiventris
Fordlandia; Para, Brazil (0340'S, 5530' W, Payn-
ter and Traylor, 1991)
Caluromys philander (right bank Rio Tapajos)
Didelphis marsupialis
Metachirus nudicaudatus (E bank Rio Tapajos)
Monodelphis emiliae (right bank Rio Tapajos)
Philander opossum (E bank Rio Tapajos)
Forest Preserve Nova Lombardia, 8 km S of
Caixa D'Agua; Espfrito Santo, Brazil (Caixa
D'Agua = 1238'S, 4055'W, USBGN)
Monodelphis scalops
Forestry Camp, 2.1 mi N of Salamanca; Toledo,
Belize (1651'N, 8901'W, Geonet)
Marmosa robinsoni
Formosa; Goias, Brazil (1532'S, 4720'W, Hersh-
kovitz, 1997)
Philander frenata (gallery forest)
Fort Clayton; Panama, Panama (0900'N,
7945'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Fort Davis; Colon, Panama (0917'N, 7954'W,
Geonet)
Caluromys derbianus
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
Forte do Rio Branco (= Forteleza de Sao Joa-
quim); Roraima, Brazil (0301'N, 6028'W,
Paynter and Traylor, 1991)
Caluromys philander
Forteleza de Sao Joaquim (see Forte do Rio Bran-
co); Roraima, Brazil
Fort Gulick; Colon, Panama (0919'N, 7952'W,
Geonet)
Caluromys derbianus
Fortin; Veracruz-Llave, Mexico (1854'N, 9700'W,
Geonet)
Didelphis marsupialis, 3200 ft (= 1000 m)
Philander opossum, 3200 ft (= 1000 m)
Fortin de Zalazar (see Fortin Juan de Zalazar);
Presidente Hayes, Paraguay
Fortfn General Genes (= General Genes), 1.6 km
NE of Asuncion; Presidente Hayes, Paraguay
(Asuncion = 2401'S, 5933'W, Paynter,
1989)
Didelphis albiventris
Fortfn Guachalla; Boqueron, Paraguay (2227'S,
6220'W, Paynter, 1989)
Didelphis albiventris, 250 m
Fortin Juan de Zalazar (= Fortin de Zalazar =
BROWN: ATLAS OF NEW WORLD MARSUPIALS
217
Juan de Zalazar); Presidente Hayes, Paraguay
(2306'S, 5918'W, Paynter, 1989)
Didelphis albiventris (left bank Rio Verde)
Monodelphis brevicaudis, ca. 150 m (left bank
Rio Verde)
Thylamys pusilla, 1 50 m
Fort Kobbe; Panama, Panama (0854'N, 7935'W,
Geonet)
Caluromys derbianus
Didelphis marsupialis
Metachirus nudicaudatus
Fort Sherman; Colon, Panama (0921 'N, 7957'W,
Geonet)
Caluromys derbianus (Camp Pifia)
Didelphis marsupialis
Metachirus nudicaudatus
Fort Sherman Military Reservation; Colon, Pan-
ama (0919'N, 7958'W, Geonet)
Marmosa robinsoni
Fortuna, Pandora; Limon, Costa Rica (Pandora =
0945'N, 8257'W, Geonet)
Caluromys derbianus
Foz do Curua (= Rio Curua do Sul); Para, Brazil
(0239'S, 5410'W, Paynter and Traylor,
1991)
Caluromys philander
Foz do Iguacu [Iguazu], 14 km NE, Itaipu Bina-
cional, Refugio Bela Vista (see Puerto Igua-
zu); Misiones, Argentina
Fracran, San Pedro; Misiones, Argentina
(2646'S, 5416'W, Paynter, 1985)
Philander opossum
Fragmentos Florestais da Cai, Aracruz Calulore,
Aracruz; Espfrito Santo, Brazil (Aracruz =
1949'S, 4016'W, Paynter and Traylor, 1991)
Metachirus nudicaudatus, 50 m
Franca; Sao Paulo, Brazil (2032'S, 4724'W,
Paynter and Traylor, 1991)
Didelphis albiventris
Gracilinanus microtarsus, 997 m
Lutreolina crassicaudata, 997 m
France Field; Colon, Panama (0921'N, 7953'W,
Geonet)
Caluromys derbianus
Francisco de C; Minas Gerais, Brazil (not located)
Philander opossum, 1580 m
Fray Bentos-Puerto Ungue, Rio Negro; Rio Ne-
gro, Uruguay (3307'S, 5519'W, Hershko-
vitz, 1992)
Gracilinanus agilis, 6 m
Frechal (see Flexal); Roraima, Brazil
Frederick Willem IV Vallen (falls); Nickerie, Su-
riname (0328'N, 5737'W, Stephens and
Traylor, 1985)
Monodelphis brevicaudata, 112 m
Friburgo (see Nova Friburgo); Rio de Janeiro,
Brazil
Frijoles, Bahia Peninsula; Colon, Panama
(0910'N, 7947'W, Geonet)
Caluromys derbianus
Frontera (see Alvaro Obregon); Tabasco, Mexico
Fuenteydueno; Nueva Esparta, Venezuela
(1100'N, 6400'W, Perez-Hernandez, 1989)
Marmosa robinsoni, 200 m
Fuentes; San Jose, Costa Rica (0956'N, 8402'W,
Geonet)
Caluromys derbianus
Fundao; Pernambuco, Brazil (0806'S, 3636'W,
Geonet)
Didelphis marsupialis, sea level
Fundo Caricacho, Urama; Carabobo, Venezuela
(1028'N, 6815'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
Fundo El Venado, Isla Chiloe, 5 km W of Palo-
mar; Los Lagos, Chile (not located)
Rhyncholestes raphanurus
Fundo Pecuario Masaguaral, 45 km S, Calabozo;
Guarico, Venezuela (0831'N, 6735'W,
Paynter, 1982)
Didelphis marsupialis, 90 m, 100 m
Marmosa robinsoni, ca. 63-100 m
Fusagasuga; Cundinamarca, Colombia (0421'N,
7422'W, Paynter, 1997)
Caluromys lanatus, 1746 m
Didelphis albiventris, 1750 m
Marmosa robinsoni, 1746 m
Marmosops fuscatus, 6000 ft (= 1875 m)
Marmosops impavidus, 6000 ft (= 1875 m)
Galaya (= Saloya Occidente), W of, Rio Saloya;
Pichincha, Ecuador (Rio Saloya = 0001'N,
7857'W, Paynter, 1993)
Caenolestes convelatus, 1 1 00 m
Caenolestes fuliginosus
Gallon Jug; Orange Walk District, Belize
(1733'N, 8902'W, Geonet)
Philander opossum
Galvez; Santa Fe, Argentina (3202'S, 6113'W,
Paynter, 1985)
Lutreolina crassicaudata, 53 m
Gamboa; Colon, Panama (0906'N, 7942'W,
Geonet)
Caluromys derbianus
Philander opossum
Gamboa, 5 mi NE (see Juan Mina Station); Pan-
ama, Panama
218
FIELDIANA: ZOOLOGY
Gansee, W of Paramaribo, now covered by Bro-
kopondo Lake; Brokopondo, Suriname
(0449'N, 5504'W, Geonet)
Marmosa lepida
Ganso Azul; Huanuco, Peru (0849'S, 7441'W,
Geonet)
Caluromys lanatus
Garanhuns; Pernambuco, Brazil (0854'S,
3628'W, USBGN)
Didelphis albiventris, 835 m
Monodelphis domestica (Sftio Riacho Fundo)
Garrafao, Estacion Rio Terezopolis [= Teresopol-
is], Maje; Rio de Janeiro, Brazil (2114'S,
4124'W, Geonet)
Philander frenata
Gatun, Rio Indio; Colon, Panama (0916'N,
7955'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni
Metachirus nudicaudatus
Philander opossum
Gavea; Rio de Janeiro, Brazil (ca. 2257'S,
4315'W, Paynter and Tray lor, 1991)
Didelphis aurita, sea level
Metachirus nudicaudatus
General Alvear (see Colonia Alvear); Mendoza,
Argentina
General Genes (see Fortin General Genes); Pres-
idente Hayes, Paraguay
General Roca; Rio Negro, Argentina (3902'S,
6735'W, Geonet)
Thylamys pallidior, 23 1 m
General Vedia, Rio de Oro; Chaco, Argentina
(2656'S, 5840'W, Paynter, 1985)
Lutreolina crassicaudata
Georgetown; Demerara-Mahaica, Guyana
(0648'N, 5810'W, Geonet)
Monodelphis brevicaudata
Philander opossum, sea level (E bank Demer-
ara River)
Glaura (see Cachoeira do Campo); Minas Gerais,
Brazil
Glycon Ridge (= Glycon Swamp); Bolivar, Ve-
nezuela (0512'N, 6044'W, Paynter, 1982)
Didelphis albiventris, 5900 ft (= 1840 m)
Glycon Swamp (see Glycon Ridge); Bolivar, Ve-
nezuela
Goias, Brazil (not precisely located)
Chironectes minimus
Golfo de Campeche; Tabasco, Mexico (Campeche
= 1950'N, 9042'W, Geonet)
Chironectes minimus
Gonzalez Catan; Buenos Aires, Argentina
(3446'S, 5839'W, Paynter, 1985)
Lutreolina crassicaudata, ca. 15 m
Gorotire (= Novo Horizonte = Gradaus); Para,
Brazil (0743'S, 51H'W, Paynter and Tray-
lor, 1991)
Micoureus demerarae, 252 m
Monodelphis emiliae, ca. 100 m
Philander opossum
Goya; Corrientes, Argentina (2908'S, 5916'W,
Paynter, 1985)
Didelphis albiventris, 600 ft (= 180 m)
Lutreolina crassicaudata, 37 m
Thylamys macrura, 37 m
Thylamys pusilla, 600 ft (= 180 m)
Gradaus (see Gorotire); Para, Brazil
Gran Chaco (see Chaco); Presidente Hayes, Par-
aguay
Grand Etang; Grenada (1205'N, 6142'YV, Geo-
net)
Marmosa robinsoni
Grande Riviere, near Tree Station; St. David, Trin-
idad and Tobago (ca. 1050'N, 6103'W,
USBGN)
Didelphis marsupialis
Great Corn Island (see Isla de Maiz); Zelaya, Nic-
aragua
Greytown {- San Juan del Norte); Rib San Juan,
Nicaragua (1055'N, 8342'W, Geonet)
Didelphis marsupialis
Philander opossum
Groningen, Saramacca River; Saramacca, Suri-
name (0548'N, 5528'W, Geonet)
Didelphis marsupialis
Groto Grande, Turmalina; Minas Gerais, Brazil
(Turmalina = 1717'S, 4245'W, Geonet)
Marmosops incanus
Guabala; Chiriqui, Panama (0813'N, 8144'W,
Geonet)
Caluromys derbianus
Didelphis marsupialis
Guaiabamba (see Santa Rosa de Huayabamba);
Amazonas, Peru
Guacamaya, Rancho Grande (see Estacion Biol-
ogica de Rancho Grande); Aragua, Venezuela
Guaicaramo (= Guaycaramo); Meta, Colombia
(0443'N, 7302'W, Paynter, 1997)
Didelphis marsupialis, 600-700 m
Marmosops impavidus, 600-700 m
Metachirus nudicaudatus
Guaillabamba, Riobamba; Pichincha, Ecuador
(0004'S, 7821'W, Paynter, 1993)
Didelphis albiventris, 3500 m
Guainche, between Alamor (0402'S, 8002'W)
and Celica; Loja, Ecuador (0407'S, 7959'W,
Paynter, 1993)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
219
Didelphis marsupialis. 975 m
Marmosops impavidus, 975 m
Micoureus regina. 975 m (near Alamor)
Gualaquiza; Morona-Santiago. Ecuador (03 ; 24'S.
7833'W. Paynter. 1993)
Marmosa lepida, 971 m
Marmosa murina, 971 m (Rio Zamora)
Marmosops noctivagus. 971 m
Gualea. Ilambo Valley: Pichincha. Ecuador
(0007'N, 7850'W. Paynter. 1993)
Caenolestes convelatus. 1100 m
Caenolestes fuliginosus
Chironectes minimus. 3000 ft (= 930 m)
Didelphis albiventris, 1 2 1 8- 1 875 m
Marmosa robinsoni, ca. 1500 m
Philander opossum, ca. 1250 m
Gualffn (see Hualfih): Catamarca. Argentina
Guama (= Iriteria = Sao Miguel do Guama):
Para. Brazil (0137'S. 4727'W. Paynter and
Traylor, 1991)
Metachirus nudicaudatus
Philander opossum
Guamalito: Norte de Santander. Colombia
(0834'N. 7327'W. Paynter. 1997)
Didelphis marsupialis. 600-700 m
Guamitas. Rancho Grande (see Estacion Biologi-
ca de Rancho Grande): Aragua. Venezuela
Guanabara. Baia de: Rio de Janeiro. Brazil
(2250'S. 4310'W. Paynter and Traylor. 1991)
Didelphis aurita
Guanagazapa: Esquintla, Guatemala (14 = 14'N.
90=39'W. Geonet)
Philander opossum
Guanay: La Paz. Bolivia (1528'S, 67=52 "W.
Paynter. 1992)
Didelphis marsupialis
Micoureus regina, 1700 ft (= 530 m) (also 5
km SE of Guanay)
Metachirus nudicaudatus
Monodelphis brevicaudata (also at 15=30'S.
6750'W)
Guanico (= Las Palmitas): Los Santos. Panama
(0718'N. 8026'W. Geonet)
Caluromxs derbianus
Chironectes minimus
Didelphis marsupialis
Guapaton. Valle de Suaza: Huila. Colombia (Rio
Suaza = 02=10'N. 7541'W. USBGN)
Didelphis marsupialis
Guaquitas: Barinas. Venezuela (0728'N. 7139'W.
Geonet)
Philander opossum
Guara. gallery forest: Distrito Federal. Brazil (not
located)
Philander opossum
Guaraciaba: Minas Gerais. Brazil (2032'S.
42=58 'W. Varejao and Valle. 1982)
Didelphis aurita
Guaraunos. Finca Vuelta Larga: Sucre. Venezuela
( 1033'N. 6307'W. Lopez-Fuster et al.. 2000)
Marmosa robinsoni
Guaratuba: Sao Paulo. Brazil (2345'S. 4555'W,
Paynter and Traylor. 1991)
Didelphis aurita. sea level
Metachirus nudicaudatus (5 km S of Casa
Grande. Varejao and Valle. 1982)
Guariba: Sao Paulo. Brazil (2121'S. 4814'W,
USBGN)
Lutreolina crassicaudata
Guasca. forest near Rio Balcones [Quebrada Bal-
cones]: Cundinamarca. Colombia (0440'N,
73=33 'W. Paynter. 1997)
Caenolestes fuliginosus. 2700-2750 m
Didelphis albiventris. 2720 m
Guatemala, lowlands, east-central Guatemala (not
located)
Caluromxs derbianus
Guatopo National Park ( see Parque Nacional Gua-
topo): Miranda-Guarico. Venezuela
Guayabal (= El Guayabal): Norte de Santander.
Colombia (0801'N. 72 C 30'W, Paynter. 1997)
Marmosa robinsoni. ca. 200 m
Guayabamba (see Santa Rosa de Huayabamba):
Amazonas, Peru
Guayabo. near Jaque: Darien. Panama (0723'N,
7802'W, Wenzel and Tipton. 1966)
Chironectes minimus
Guayabo. near Gualaquiza: Morona-Santiago. Ec-
uador (Gualaquiza = 0324'S, 7833'W, Geo-
net)
Marmosops noctivagus
Guayaquil: Guayas. Ecuador (0210'S. 7950'W,
Paynter. 1993)
Didelphis marsupialis. sea level
Marmosa robinsoni. sea level
Philander opossum, near sea level
Guayaramerin: Beni. Bolivia (1048'S. 6523'W,
Paynter. 1992)
Didelphis marsupialis
Philander opossum (1051'S. 6523'W)
Guaycaramo (see Guaicaramo): Meta. Colombia
Guayquitas: Barinas. Venezuela (0830'N,
70=23 'W. Perez-Hernandez. 1989)
Philander opossum
Giiemes. Highway 34. km 1383: Salta. Argentina
(2440'S, 6403'W, Flores et al., 2000)
Thxlamxs venustus
220
FIELDIANA: ZOOLOGY
Guengue; Valle del Cauca, Colombia (ca.
0314'N, 7621'W, Paynter, 1997)
Didelphis marsupialis, 3650 ft (= 1 140 m)
Giiiniquina (= Uinikina); Delta Amacuro Terri-
torio Federal, Venezuela (0910'N, 6103'W,
Hershkovitz, 1997)
Didelphis marsupialis, sea level
Philander opossum, sea level
Gutierrez; Santa Cruz, Bolivia (1925'S, 6334'W,
Anderson, 1997)
Thy I amy s venustus
H
Hacienda Amazonia, Alto Rio Madre de Dios;
Madre de Dios, Peru (1256'S, 7110'W, Pa-
checo et al., 1993)
Micoureus regina
Monodelphis adusta, 825 m (Hacienda Rio Ma-
dre de Dios, Alto Rio Madre de Dios)
Hacienda Azacualpa, near Villa El Carmen; Ma-
nagua, Nicaragua (Villa El Carmen =
1159'N, 8631'W, Geonet)
Didelphis marsupialis
Philander opossum (5 km NW of Villa El Car-
men)
Hacienda Bellavista, Volcan Casita; Chinandega,
Nicaragua (Volcan Casita = 1242'N,
8658'W, Geonet)
Caluromys derbianus
Hacienda Buena Vista, few km above Vista Ale-
gre on Rio Chinchao; Huanuco, Peru (Vista
Alegre = ca. 0931'S, 7552'W, Stephens and
Traylor, 1983)
Caluromys lanatus
Didelphis albiventris, 3500 ft (= 1090 m),
1250 m
Didelphis marsupialis, 3500-4000 ft (= 1090-
1250 m)
Hacienda Cadena, Quisipanchi; Cusco, Peru
(1324'S, 7043'W, Stephens and Traylor,
1983)
Caluromys lanatus, ca. 1000 m
Chironectes minimus, 1000 m
Hyladelphys kalinowskii, 1000 m
Marmosa murina, ca. 1000 m
Marmosops noctivagus, 1000 m
Micoureus regina, ca. 1000 m
Philander opossum, 890 m (left bank Rfo Mar-
capata, 1325'S, 7146'W)
Hacienda Capulin; San Luis Potosf, Mexico
(2130'N, 9937'W, Gardner, 1973)
Didelphis marsupialis
Hacienda Carapiche; Distrito Federal, Venezuela
(1029'N, 6719'W, Handley, 1976)
Didelphis marsupialis, 398 m, 600 m (near El
Limon)
Monodelphis palliolata, 398 m (near El Limon)
Hacienda Chilata; Sonsonate, El Salvador
(1339'N, 8934'W, Gardner, 1973)
Didelphis marsupialis, 2000 ft (= 625 m)
Philander opossum, 2000 ft (= 625 m)
Hacienda Cincinnati, Santa Marta; Magdalena,
Colombia (1106'N, 7406'W, Paynter, 1997)
Caluromys lanatus
Hacienda Conzulman [Conzulmann], N Cuyam-
buyo; Tarija, Bolivia (not located)
Glironia venusta, 1000 m
Hacienda de Carmen; Yucatan, Mexico (not lo-
cated)
Philander opossum
Hacienda El Limon; Distrito Federal, Venezuela
(1028'N, 6717'W, Paynter, 1982)
Didelphis marsupialis, 100 m (Suapo), 600 m
(La Cochita), 1400 m (Zona Guacatal)
Gracilinanus marica, 600-1400 m, 1400 m
(Zona Guacatal)
Monodelphis palliolata, 100 m (Suapo), 600 m,
850 m (La Cochita), 1400 m (Zona Guacatal)
Hacienda El Merey (see Montalban); Carabobo,
Venezuela
Hacienda Erika, Rio Salvacion; Cusco, Peru (Rio
Salvacion = 1355'S, 7112'W, Geonet)
Chironectes minimus, 300-500 m
Didelphis marsupialis, 350-500 m
Metachirus nudicaudatus, 350-500 m (Rio
Alto Madre de Dios, opposite Salvacion)
Philander opossum, 300-500 m (Rfo Alto Ma-
dre de Dios)
Hacienda Exito; Huanuco, Peru (0926'S,
7600'W, USBGN)
Didelphis marsupialis, 3000 ft (= 930 m)
Metachirus nudicaudatus, 3000 m (Rfo Cayum-
ba)
Hacienda Garzon, Cerro Pichincha; Pichincha,
Ecuador (Cerro Pichincha = 0010'S,
7833'W, Paynter, 1993)
Caenolestes fuliginosus
Hacienda La Carbonera (= La Carbonera), Para-
mo Tambor; Merida, Venezuela (0838'N,
7121'W, Paynter, 1982)
Didelphis albiventris, 6000-7000 ft (= 1875-
2180 m), 2916 m
Hacienda La Esperanza, 8 km de Magdaleno,
Lago de Valencia; Aragua, Venezuela
(1006'N, 6737'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 51 A m
Monodelphis brevicaudata
Hacienda La Laguna, near Andahuaylas; Apurf-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
221
mac, Peru (Andahuaylas = 1339'S, 7323'W,
Stephens and Traylor, 1983)
Didelphis albiventris, 3040 m
Hacienda La Palmira, near Andahuaylas; Apurf-
mac, Peru (Andahuaylas = 1339'S, 7323'W,
Stephens and Traylor, 1983)
Didelphis albiventris, 2200 m
Hacienda La Palmita (see Hato La Palmita); Gua-
rico, Venezuela
Hacienda La Primavera, east Andes, E side; Boy-
aca, Colombia (ca. 0700'N, 7220'W, Payn-
ter, 1997)
Caenolestes fuliginosus, 7000 ft (= 2180 m)
Hacienda La Trampa; Jinotega, Nicaragua (ca.
1308'N, 8554'W, Gardner, 1973)
Didelphis marsupialis (near Jinotega)
Philander opossum (16 km NE of Jinotega)
Hacienda La Vega, 5 km ESE of San Pedro del
Lago; Imbabura, Ecuador (0013'N, 7812'W,
Patterson and Albuja, 1996)
Caenolestes convelatus, 2573 m
Hacienda Limon, near Balsas; Cajamarca, Peru
(ca. 0650'S, 7805'W, Stephens and Traylor,
1983)
Didelphis albiventris, 3000 ft (= 930 m)
Hacienda Llagueda; La Libertad, Peru (ca.
0746'S, 7835'W, Stephens and Traylor,
1983)
Didelphis marsupialis, 2000 ft (= 625 m)
Hacienda Luisiana (see Luisiana); Ayacucho,
Peru
Hacienda Magdalena (= Pueblo Juarez); Colima,
Mexico (1910'N, 10355'W, Geonet)
Marmosa canescens
Hacienda Mecatepe, near Nandaime; Granada,
Nicaragua (Nandaime = 1145'N, 8603'W,
Geonet)
Didelphis marsupialis
Hacienda Misisi; Trujillo, Venezuela (0921'N,
7018'W, Perez-Hernandez, 1989)
Gracilinanus dryas, 2210-2360 m
Marmosops fuscatus, 2210-2350 m
Hacienda Ochenta; Nueva Esparta, Venezuela
(1100'N, 6400'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Hacienda Paramba (= Paramba), Rio Mira; Im-
babura, Ecuador (0049'N, 7821'W, Paynter,
1993)
Caluromys derbianus
Marmosa robinsoni, 111 m
Philander opossum, 1 1 00 m
Hacienda Pelicano; Santa Cruz, Bolivia (1825'S,
6210'W, Anderson, 1997)
Marmosops dorothea
Hacienda Porvenir, 4 mi above Hacienda Exito;
Huanuco, Peru (Hacienda Exito = 0926'S,
7600'W, Stephens and Traylor, 1983)
Didelphis marsupialis, 3000 ft (= 930 m)
Hacienda Quetepe, 16 km E of Cumana; Sucre,
Venezuela (1026'N, 6401'W, Lopez-Fuster
et al., 2000)
Marmosa robinsoni
Hacienda San Isidro, El Realejo, 10 km S of Chin-
andega; Chinandega, Nicaragua (Chinandega
= 1237'N, 8709'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Philander opossum
Hacienda San Pedro, Santa Cruz de Mora (see
Santa Cruz); Merida, Venezuela
Hacienda San Ramon, Puna; Guayas, Ecuador
(0219'S, 7928'W, USBGN)
Marmosa robinsoni
Hacienda Santa Elena (= Santa Elena), 35 km NE
of Tingo Maria; Huanuco, Peru (Tingo Maria
= 0918'S, 7559'W, Stephens and Traylor,
1983)
Metachirus nudicaudatus, 1000 m
Hacienda Santa Maria, 15 mi NE of Liberia;
Guanacaste, Costa Rica (1047'N, 8519'W,
USBGN)
Philander opossum
Hacienda Santa Teresi [possibly Hacienda Santa
Teresfta at 0212'S, 8010'W, USBGN], 4 km
SW of Puebloviejo; Los Rfos, Ecuador (Pueb-
loviejo = 0134'S, 7930'W, Paynter, 1993)
Didelphis marsupialis
Philander opossum (ca. 12 km N of Vinces,
Vinces = 0132'S, 7945'W)
Hacienda Tamiahua, Cabo Rojo; Veracruz-Llave,
Mexico (2144'N, 9733'W, Gardner, 1973)
Didelphis marsupialis
Hacienda Taulis; Cajamarca, Peru (0654'S,
7903'W, Stephens and Traylor, 1983)
Didelphis albiventris, 1700 m
Hacienda Tepeyac, near Matagalpa; Matagalpa,
Nicaragua (Matagalpa = 1255'N, 8555'W,
Geonet)
Didelphis marsupialis
Philander opossum
Hacienda Tepello; Matagalpa, Nicaragua (not lo-
cated)
Philander opossum
Hacienda Villa Carmen; Cusco, Peru (ca. 1250'S,
7115'W, Stephens and Traylor, 1983)
Caluromys lanatus, 600 m
Chironectes minimus, 600 m
Marmosa andersoni, 600 m
222
FIELDIANA: ZOOLOGY
Marmosa rubra, 600 m
Marmosops noctivagus, 600 m
Marmosops parvidens, 600 m
Hacienda Zapotitan, Pacific Coast, Rio Sucio; La
Libertad, El Salvador (1348'N, 8927'W,
USBGN)
Chironectes minimus, 1500 ft (= 460 m)
Hama (see Cafetal de Chama); Merida, Venezuela
Hamacas [Hamecas]; Santa Cruz, Bolivia
(1744'S, 6311'W, Hershkovitz, 1997)
Philander opossum
Hamonia (see Ibirama); Santa Catarina, Brazil
Hansa (see Corupa); Santa Catarina, Brazil
Hato Acapulco, 25 km S of La Trinidad de Arau-
ca; Apure, Venezuela (0701'N, 6907'W,
Geonet)
Marmosa robinsoni
Hatillo; San Jose, Costa Rica (0955'N, 8406'W,
Geonet)
Didelphis marsupialis
Hato Cariben, Rio Cinaruco, 46 km ENE of
Puerto Paez; Apure, Venezuela (0633'N,
6713'W, Handley, 1976)
Didelphis marsupialis, 76 m
Hato El Frio, W of El Saman de Apure; Apure,
Venezuela (0749'N, 6859'W, Lopez-Fuster
et al., 2000)
Marmosa robinsoni
Hato El Tirado; Cojedes, Venezuela (0904'N,
6822'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Marmosa robinsoni
Hato Flores Moradas; Guarico, Venezuela
(0816'N, 6736'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Hato La Fe, Corozpando; Guarico, Venezuela
(0903'N, 6750'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Hato La Florida; Bolivar, Venezuela (0730-
0725'N, 6557'-6539'W, Handley, 1976)
Didelphis marsupialis, 43-45 m
Micoureus demerarae, 43-45 m
Hato Las Palmitas (= Hacienda La Palmita), San
Francisco de Tiznados; Guarico, Venezuela
(0936'N, 6727'W, Handley, 1976)
Marmosa robinsoni, 181m
Monodelphis brevicaudata
Hato Mata de Bejuco; Monagas, Venezuela
(0919'N, 6256'W, Handley, 1976)
Didelphis marsupialis, 18 m
Gracilinanus marica, 18 m
Marmosa robinsoni, 18 m
Hato Nuevo, Cojedes, Venezuela (0913'N,
6805'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Monodelphis brevicaudata
Hato Pinedo; Cojedes, Venezuela (0853'N,
6804'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Hato San Jose, 20 km W of La Paragua; Bolivar,
Venezuela (La Paragua = 0650'N, 6320'W,
Handley, 1976)
Didelphis marsupialis, 324 m (0649'N,
6329'W)
Lutreolina crassicaudata, 298 m
Marmosa murina, 306 m (0649'N, 6329'W)
Micoureus demerarae, 306 m (0649'N,
6329'W)
Philander opossum, 302 m, 524 m (146 km S,
7 km NE of Hato San Jose, 0644'N,
6327'W)
Heights of Ourepouche [Oropuche] (see El Cerro
del Oropuche); St. David, Trinidad and To-
bago
Hermanos (see Dois Irmaos); Pernambuco, Ar-
gentina
Hickmann; Salta, Argentina (2312'S, 6334'W,
Paynter, 1985)
Thylamys pusilla, 280 m
Highway 3 at junction of Highways 3 and 34;
Jujuy, Argentina (2348'S, 6448'W, Flores et
al., 2000)
Thylamys pallidior
Thylamys venustus
Highway 29, 10 km W of Tfrafi; Jujuy, Argentina
(2359'S, 6523'W, Paynter, 1985)
Thylamys venustus, 5800 ft. (= 1810 m)
Highway 62 (see Andalgala); Catamarca, Argen-
tina
Highway 65, between Andalgala and San Miguel
de Tucuman; Catamarca, Argentina (not lo-
cated)
Didelphis albiventris, 2000 f (= 625 m)
Highway 338 on road to Horco Molle, 4 km W
of junction (see also Horco Molle); Tucuman,
Argentina (2649'S, 6519'W, Caplonch et al.,
1997)
Didelphis albiventris, 2750 ft (= 860 m)
Thylamys venustus, 2750 ft (= 860 m)
Higuerilla, 20 km E of Tilcara; Jujuy, Argentina
(Tilcara = 2334'S, 6522'W, Paynter, 1985)
Thylamys venustus, 2000 m
"Hill Plot" (see Balcarce); Buenos Aires, Argen-
tina
Honda; Tolima, Colombia (0512'N, 7445'W,
Paynter, 1997)
Didelphis marsupialis, 230 m
Marmosa robinsoni, 230 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
223
Honda Bay (see Bahia Honda); La Guajira, Co-
lombia
Hoolie Island, Essequibo River; Rupununi, Guy-
ana (not located)
Marmosa murina
Horco Molle, 25 km NW of San Miguel de Tu-
cuman (also see Highway 338 on road to Hor-
co Molle); Tucuman, Argentina (2645'S.
6521'W, Mares et al., 1996)
Didelphis albiventris (Reserva Biologica [ =
Biological Reserve] de Horco Molle)
Lutreolina crassicaudata, 650 m
Thylamys venustus, 2400 ft (= 750 m) (Reserva
Biologica [= Biological Reserve] de Horco
Molle)
Horcus; Chusquisaca, Bolivia (1929'S, 6433'W.
USBGN)
Thylamys venustus, 6000 ft (= 1875 m)
Horquilla, Rio Machupo, 15 km above; Beni, Bo-
livia (1241'S, 6432'W. Anderson, 1997)
Didelphis marsupialis
Hotel Las Pozas, Villa Tunari; Cochabamba. Bo-
livia (1657'S, 6523'W, Anderson, 1997)
Didelphis marsupialis
Hotel Tunquelen, 25 km E of San Carlos de Bar-
iloche; Rio Negro, Argentina (San Carlos de
Bariloche = 4109'S. 7118'W. Paynter, 1985)
Dromiciops gliroides
Huacapistana; Junfn, Peru (1 114'S, 7529'W, Ste-
phens and Traylor, 1983)
Didelphis albiventris, ca. 2500 m
Huachipa; Huanuco, Peru (ca. 0930'S. 7552'W,
Stephens and Traylor. 1983)
Micoureus regina, 3000 ft ( = 930 m). 4600 ft
(= 1430 m)
Huajchilla; La Paz, Bolivia (1637'S, 6803'W,
Anderson, 1997)
Thylamys pallidior
Thylamys venustus
Huaijumbe (see Huajyumbe); Cusco, Peru
Huajyumbe (= Huaijumbe = Huayjumbe); Cusco.
Peru (ca. 1315'S, 7035'W, Stephens and
Traylor, 1983)
Chironectes minimus, 630 m
Metachirus nudicaudatus
Hualfin (= Gualfin); Catamarca, Argentina
(2714'S, 6650'W, Paynter, 1985)
Thylamys pallidior, 2220 m
Hualtaran, Parque Nacional Sierra de Quijadas;
San Luis, Argentina (ca. 3229'S, 6700'W.
Mares and Braun, 2000)
Thylamys pallidior, ca. 2800 ft (= 875 m)
Huampami (= Kayamas), near Aguaruna Village.
Rio Cenepa (also see Rio Cenepa); Amazon-
as. Peru (0428'S. 7810'W, Mustrangi and
Patton. 1997)
Didelphis marsupialis
Marmosa rubra, 700 ft (= 220 m)
Metachirus nudicaudatus
Micoureus regina, 212 m
Philander andersoni, 700 ft (= 220 m)
Huancabamba. Canchaque: Piura. Peru (0515'S,
7925'W. Stephens and Traylor, 1983)
Didelphis albiventris, 1198 m
Huancabamba, km 30 on road to San Ignacio; Pi-
ura. Peru (0515'S, 7929'W, Stephens and
Traylor. 1983)
Caenolestes caniventer, 3000 m
Huanhuachayo; Ayacucho. Peru (ca. 1244'S.
7347'W. Stephens and Traylor, 1983)
Metachirus nudicaudatus, 1660 m
Philander andersoni, 1660 m
Philander opossum, 1 660 m
Huanuco. on banks of Rio Huallaga; Huanuco.
Peru (0956'S. 7615'W. Stephens and Tray-
lor. 1983)
Didelphis albiventris, 6600 ft (= 2060 m)
Huaraco; La Paz, Bolivia (1710'S, 6755'W. An-
derson. 1997)
Thylamys pallidior
Huarandosa, Rio Chinchipe; Cajamarca. Peru (Rio
Chinchipe = 0528'S. 7832'W, Stephens and
Traylor, 1983)
Didelphis marsupialis, 915 m
Huaras [Huarez]; Ancash, Peru (0930'S,
7736'W, Stephens and Traylor. 1983)
Thylamys pallidior, 3052 m
Huayjumbe (see Huajyumbe); Cusco. Peru
Huechun. Embalse (see Embalse Huechiin); San-
tiago. Chile
Huehuetan; Chiapas, Mexico (1501'N. 9224'W,
Geonet)
Marmosa mexicana
Philander opossum
Huemul. Nahuel Huapi; Neuquen. Argentina (ca.
4055'S. 7130'W, Paynter. 1985)
Dromiciops gliroides, 167 m
Huepil; Biobio, Chile (3714'S. 7156'W, Geonet)
Dromiciops gliroides
Huichihuyan; San Luis Postosi, Mexico (2130'N.
9857'W. USBGN)
Philander opossum
Huitanaa (= Iutanaa = Hyutanaha = Hyutanaham
= "Hyunaaham"), upper Rio Purus; Ama-
zonas. Brazil (0740'S, 6546'W, Paynter and
Traylor, 1991)
Marmosops noctivagus, ca. 50 m
224
FIELDIANA: ZOOLOGY
Huite (= Puerto Huite), Isla Chiloe; Los Lagos,
Chile (4207'S, 7327'W, Geonet)
Dromiciops gliroides
Humahuaca, 1 1 km E, 2 km E of Pucara on road
to Cianzo; Chubut, Argentina (2312'S,
6516'W, Flores et al., 2000)
Thylamys pallidior, 1 1500 ft (= 3590 m)
Humberto de Campos (see Miritiba); Maranhao,
Brazil
Humboldt (see Corupa); Santa Catarina, Brazil
Hurlingham; Buenos Aires, Argentina (3436'S,
5838'W, USBGN)
Lutreolina crassicaudata, ca. 20 m
Hyama Creek, Demerara River; Demerara-Mahai-
ca, Guyana (0627'N, 5818'W, Geonet)
Marmosa murina
Hyde Park; Demerara-Mahaica, Guyana (0630'N,
5816'W, Geonet and Times Atlas, 1992)
Caluromys philander
Marmosa murina
Marmosops parvidens, 20 ft (= 6 m), 30 m
Micoureus demerarae
Philander opossum (Demerara River)
"Hyunaaham" (see Huitanaa); Amazonas, Brazil
Hyutanaba (see Huitanaa); Amazonas, Brazil
Hyutanaham (see Huitanaa); Amazonas, Brazil
I
Ibarra; Imbabura, Ecuador (0021'N, 7807'W,
Paynter, 1993)
Didelphis albiventris, 6600-7500 ft (= 2060-
2340 m)
Ibiapaba (see Serra Ibiapaba); Ceara, Brazil
Ibiato; Beni, Bolivia (1448'S, 6418'W, Ander-
son, 1997)
Didelphis marsupialis
Ibicui [Ibicuhy] (see Mangaratiba); Rio de Janei-
ro, Brazil
Ibirama (= Hamonia); Santa Catarina, Brazil
(2704'S, 4931'W, Paynter and Traylor, 1991)
Monodelphis iheringi
Ibitf (see Monte Alegre do Sul); Sao Paulo, Brazil
Icabarii, 23 km NE; Bolivar, Venezuela (0420'N,
6245'W, Geonet)
Didelphis albiventris, 824 m
Didelphis marsupialis (23-45 km NE of Ica-
barii)
Marmosa murina, 473-982 m (45 km NE of
Icabarii)
Monodelphis brevicaudata, 741-851 m (18-45
km NE of Icabarii)
Igarape Acu (= Colonia do Prata = Igarape Assu
= Lazaopolis do Prata); Para, Brazil (0107'S,
4737'W, Paynter and Traylor, 1991)
Marmosa murina, 50 m
Metachirus nudicaudatus
Philander opossum
Igarape Amorin (= Amarim Igarape = Limontu-
ba), Rio Tapajos; Para, Brazil (0226'S,
5500'W, Hershkovitz, 1977)
Didelphis marsupialis
Metachirus nudicaudatus (E bank Rio Tapajos)
Igarape Assu (see Igarape-Acu); Para, Brazil
Igarape Bravo, Rio Tapajos; Para, Brazil (0224'S,
5441'W, Paynter and Traylor, 1991)
Didelphis marsupialis
Igarape Cacao Pereira (see Cacau Pirera); Ama-
zonas, Brazil
Igarape Grande, Rio Jurua; Amazonas, Brazil (ca.
0635'S, 6950'W, Paynter and Traylor, 1991)
Caluromys lanatus
Gracilinanus emiliae
Marmosa murina (W bank Rio Jurua, ca.
0630'S, 6950'W)
Igarape Maroi (see Marai); Para, Brazil
Igarape Nova Empressa (= Nova Empresa), left
bank Rio Jurua; Amazonas, Brazil (0648'S,
7044'W, Patton et al., 2000)
Marmosops neb Una
Micoureus regina
Philander opossum
Igarape Porongaba, Rio Jurua; Acre, Brazil
(0840'S, 7247'W, Patton et al., 2000)
Caluromys lanatus (right and left banks)
Didelphis marsupialis (right and left banks)
Marmosops impavidus (right bank)
Marmosops neblina (right bank)
Marmosops noctivagus (right and left banks)
Metachirus nudicaudatus (right and left banks)
Micoureus demerarae (right and left banks)
Micoureus regina (right and left banks)
Monodelphis emiliae (right bank)
Philander mcilhennyi (right and left banks)
Philander opossum (right bank)
Igarape Rio Branco, upper Rio Maraca, Mazagao;
Amapa, Brazil (see Rio Maraca = 0032'S,
5112'W, Paynter and Traylor, 1991)
Monodelphis brevicaudata
Igarape Tapereba, Chaves; Para, Brazil (0010'S,
4955'W, Hershkovitz, 1997)
Philander opossum
Iguape; Sao Paulo, Brazil (2443'S, 4733'W,
Paynter and Traylor, 1991)
Monodelphis iheringi
Philander frenata
Iguasii (see Puerto Iguazii); Misiones, Argentina
Iguazu (see Puerto Iguazu); Misiones, Argentina
BROWN: ATLAS OF NEW WORLD MARSUPIALS
225
llama; Yoro, Honduras (ca. 1456'N, 8813'W,
Geonet)
Caluromys derbianus
Ilha das Oncas, left bank Rio Negro, near Miran-
da, left bank Rio Jurua; Amazonas, Brazil
(0555'S, 6138'W, Geonet)
Philander opossum
Ilha das On^as, opposite Belem; Para, Brazil
(0127'S, 4833'W, Paynter and Traylor, 1991)
Chironectes minimus, ca. 10 m
Ilha da Vitoria (= Ilha Victoria); Sao Paulo, Bra-
zil (2350'S, 4500'W, Hershkovitz, 1992)
Gracilinanus microtarsus, near sea level
Ilha de Guahyba (see Ilha Guaiba); Rio Grande
do Sul, Brazil
Ilha de Marajo (= Caldeirao = Marajo); Para,
Brazil (OTOO'S, 4930'W, Paynter and Tray-
lor, 1991)
Didelphis marsupialis, 10 m
Marmosa murina, ca. 10 m
Micoureus demerarae, ca. 10 m
Monodelphis maraxina, ca. 10 m
Ilha de Sao Sebastiao (= Fazenda da Toca); Sao
Paulo, Brazil (2350'S, 4518'W. USBGN)
Didelphis aurita
Gracilinanus microtarsus
Marmosops incanus, 150 m
Philander frenata, 150 m (2348'S, 4525'W,
Paynter and Traylor, 1991)
Ilha do Guahyba (see Ilha Guaiba); Rio Grande
do Sul, Brazil
Ilha do Mosqueiro, near Belem; Para, Brazil
(0107'S, 4824'W, Paynter and Traylor, 1991)
Chironectes minimus, ca. 10 m
Ilha do Taiuna, Rio Tocantins, opposite Cameta;
Para, Brazil (0215'S, 4929'W, Hershkovitz,
1997)
Didelphis marsupialis
Marmosops parvidens
Metachirus nudicaudatus
Philander opossum
Ilha Grande; Rio de Janeiro. Brazil (2309'S,
4430'W, Hershkovitz, 1992)
Didelphis aurita
Gracilinanus microtarsus, sea level (near Ilha
Grande)
Marmosops incanus, sea level
Monodelphis americana
Ilha Guahyba (see Ilha Guaiba); Rio Grande do
Sul, Brazil
Ilha Guaiba ( = Ilha Guahyba = Ilha de Guahyba
= Ilha do Guahyba = Isla Guahyba), Porto
Alegre; Rio Grande do Sul, Brazil (3004'S,
51H'W, Hershkovitz, 1992)
Gracilinanus microtarsus, sea level
Ilha Mexiana, Fazenda Nazareth; Para, Brazil
(0002'N, 4935'W, Paynter and Traylor,
1991)
Didelphis marsupialis, sea level
Ilha Paxiuba. right bank Rio Jurua; Amazonas,
Brazil (0319'S, 6600'W, Patton et al., 2000)
Caluromys lanatus
Didelphis marsupialis
Marmosa murina
Marmosops parvidens
Metachirus nudicaudatus
Micoureus demerarae
Ilha Victoria (see Ilha da Vitoria); Sao Paulo, Bra-
zil
Ilhazinha, left bank Rio Jurua; Amazonas, Brazil
(0317'S, 6614'W, Mustrangi and Patton,
1997)
Didelphis marsupialis
Marmosops parvidens
Ilheus; Bahia, Brazil (1439'S, 3902'W, Mustran-
gi and Patton, 1997)
Didelphis aurita, sea level
Marmosops incanus, sea level
Iliapaba (= Cerro Iliapaba = Serra Iliapaba);
Ceara, Brazil (0503'S, 4055'W, Hershkovitz,
1992)
Gracilinanus agilis
Monodelphis domestica
Illapel. between Illapel and Salamanca; Coquim-
bo, Chile (Illapel = 3137'S, 7109'W, Payn-
ter, 1988)
Thyla??iys elegans
Imperatriz (see Parintins); Amazonas, Brazil
Inajatuba, lower Rio Tapajos; Para, Brazil (not lo-
cated)
Didelphis marsupialis
Inanez; Pasco, Peru (not located)
Philander andersoni, 5000 ft (= 1560 m)
Incachaca; Cochabamba, Bolivia (1714'S,
6549'W, Paynter, 1992)
Didelphis albiventris, 7700 ft (= 2400 m)
Marmosops impavidus
Marmosops noctivagus, 2350 m
Monodelphis osgoodi, 2600 m
Inca Mines (not "Juliaca"), Rio Inambari (= San-
to Domingo); Puno, Peru (1351'S, 6941'W,
Stephens and Traylor, 1983)
Chironectes minimus (coordinates incorrectly
given as 1530'S, 7008'W)
Didelphis albiventris, 7000 ft (= 2180 m)
Marmosops noctivagus, 1689 m
Micoureus regina, 6000 ft (= 1875 m)
226
FIELDIANA: ZOOLOGY
Independencia; Pando, Bolivia (1 126'S, 6734'W,
Anderson, 1997)
Gracilinanus agilis
Marmosops impavidus
Monodelphis brevicaudata
Philander opossum
Indillama (= Intillama); Pastaza, Ecuador
(0135'S, 7754'W, Geonet)
Philander andersoni (Rio Napo)
Inga Pirca; Pichincha, Ecuador (0010'S,
7823'W, USBGN)
Caenolestes fuliginosus
Ingeniero Guillermo N. Juarez (= Ingeniero Jua-
rez); Formosa, Argentina (2354'S, 6151'W,
Paynter, 1985)
Gracilinanus agilis, 100 m
Thylamys pusilla, 518 ft (= 160 m) (35 km S
of Ingeniero Guillermo N. Juarez, 2411'S,
6153'W)
Ingeniero Juarez (see Ingeniero Guillermo N. Jua-
rez); Formosa, Argentina
Ingeniero Mora; Santa Cruz, Bolivia (1808'S,
6312'W, Anderson, 1997)
Gracilinanus agilis
Marmosops dorothea
Monodelphis domestica
Ingenio La Esperanza; Jujuy, Argentina (2414'S,
6452'W, Flores et al., 2000)
Gracilinanus agilis
Ingenio San Martin del Tabacal, Lote "Milagros";
Salta, Argentina (2316'S, 6415'W, Flores et
al, 2000)
Gracilinanus agilis
Instituto Agronomico do Norte (see Belem); Para,
Brazil
Instituto Venezolano de Investigaciones Cientffi-
cas (IVIC); Miranda-Guarico, Venezuela
(1024'N, 6658'W, Handley, 1976)
Caluromys philander, 1 600 m
Didelphis marsupialis, 1460-1665 m
Marmosops fuscatus, 1460 m
INTA areas (see Estacion Experimental del Insti-
tuto Nacional de Tecnologia Agropecuaria);
Buenos Aires, Argentina
Intag, Penaherra; Imbabura, Ecuador (ca.
0024'N, 7836'W, Paynter, 1993)
Philander opossum
Intervales (see Fazenda Intervales); Sao Paulo,
Brazil
Intervales Biological Reserve; Sao Paulo, Brazil
(2420'S, 4825'W, Mustrangi, 1994)
Marmosops incanus
Intillama (see Indillama); Pastaza, Ecuador
Ipanema ( = Bacaetava = Rio Ipanema = Rio
Ypanema = Varnhagen = Ypanema); Sao
Paulo, Brazil (2326'S, 4736'W, Paynter and
Traylor, 1991)
Caluromys philander, 950 m
Chironectes minimus, 600 m
Didelphis aurita, 600 m (Rio Ipanema)
Gracilinanus microtarsus, 600 m
Metachirus nudicaudatus
Monodelphis americana, 950 m
Philander frenata, 600 m
Thylamys velutinus, 950 m
Ipeau-Apez, suburb of Belem (see Belem); Para,
Brazil
Ipiranga; Sao Paulo, Brazil (ca. 2336'S, 4637'W,
Paynter and Traylor, 1991)
Gracilinanus microtarsus
Ipitinga, right bank Rio Acara; Para, Brazil
(0157'S, 4811'W, Hershkovitz, 1997)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
Iporanga, Lajeado; Sao Paulo, Brazil (2436'S,
4834'W, Hershkovitz, 1997)
Philander frenata (Lajeado)
Philander opossum (Parque Estadual Petar)
Ipu; Ceara, Brazil (0420'S, 4042'W, Paynter and
Traylor, 1991)
Gracilinanus agilis, 250 m, 300 m
Monodelphis domestica, 250 m
Ipure (= Ypure), near Cumana; Sucre, Venezuela
(Cumana = 1028'N, 6410'W, Paynter, 1982)
Caluromys philander, 2350 ft (= 700 m)
Marmosa robinsoni
Iquiri (see Ituxi); Acre, Brazil
Iquitos (= Itaya); Loreto, Peru (0346'S, 7315'W,
Stephens and Traylor, 1983)
Didelphis marsupialis, 106 m
Metachirus nudicaudatus (Rio Maranon)
Philander andersoni (0345'S, 7312'W)
Philander opossum (left bank Rio Maranon-
Itaya, 0345'S, 7312'W)
Irazu; Cartago, Costa Rica (0959'N, 8351'W,
Geonet)
Caluromys derbianus
Ireng Valley; Upper Takutu-Upper Essequibo,
Guyana (ca. 0405'N, 5930'W, Times Atlas,
1992, and Hershkovitz files)
Micoureus demerarae
Iriteria (see Guama); Para, Brazil
Irocanga; Para, Brazil (ca. 0230'S, 5510'W, Pat-
terson, 1992)
Marmosa murina
Irupana, 2 km S; La Paz, Bolivia (1630'S,
6728'W, Anderson, 1997)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
227
Metachirus nudicaudatus
Iscayachi, Rio Tomayapo; Tarija, Bolivia
(2129'S, 6457'W, Anderson, 1997)
Thylamys pallidior
Isla Barro Colorado (see Barro Colorado): Colon/
Panama, Panama
Isla Bastimentos; Bocas del Toro. Panama
(0918'N, 8208'W, Geonet)
Caluromys derbianus
Philander opossum
Isla Brava; Chiriqui, Panama (0813'N, 8216'W,
Geonet)
Caluromys derbianus
Isla Buenaventura; (Colon?), Panama (not locat-
ed)
Philander opossum
Isla Cebaco; Veraguas, Panama (0732'N,
8109'W, USBGN)
Caluromys derbianus
Chironectes minimus
Philander opossum
Isla Cigarron, Rio Negro; Amazonas. Venezuela
(0144'N, 6702'W. Perez-Hernandez. 1989)
Micoureus demerarae
Isla Coiba (= Coiba Island); Veraguas, Panama
(0730'N, 8142'W, Geonet)
Didelphis marsupialis
Isla Colon; Bocas del Toro, Panama (0924'N,
8217'W, Geonet)
Philander opossum
Isla de Canas, 43.7 km NW of junction of High-
ways 50 and 18 on road east (see junction of
Highways 50 and 18): Salta, Argentina
Isla de La Tuna (= La Tuna): Rocha. Uruguay
(3440'S, 5409'W, Paynter, 1994)
Didelphis albiventris, sea level
Lutreolina crassieaudata, sea level
Isla del Carmen; Campeche, Mexico (1842'N,
9139'W, Geonet)
Didelphis marsupialis
Isla de los Pajaros; Chubut, Argentina (ca.
4225'S, 64 31'VV, Paynter, 1985)
Thylamys pallidior, sea level
Isla del Rey, near Isla San Miguel: Panama, Pan-
ama (Isla San Miguel = 0823'N, 7856'W.
Gardner, 1973)
Marmosa robinsoni
Isla de Maiz (= Great Corn Island): Zelaya, Nic-
aragua (1215'N, 8300'W, Geonet)
Marmosa mexicana
Isla de Margarita (see Isla Margarita); Nueva Es-
parta, Venezuela
Isla Ella, Parana delta: Buenos Aires. Argentina
(Parana delta boca = ca. 3343'S, 5915'W,
Geonet)
Lutreolina crassieaudata
Isla Escudo de Veraguas: Bocas del Toro, Panama
(0906'N. 8133'W, Geonet)
Caluromys derbianus
Isla Gargantiia; Pando, Bolivia (1224'S, 6835'W,
USBGN)
Caluromys lanatus
Didelphis marsupialis
Philander opossum
Isla Gobernadora: Veraguas, Panama (0733'N,
8112'W. Geonet)
Caluromys derbianus
Philander opossum
Isla Guahyba (see Uha Guafba); Rio Grande do
Sul. Brazil
Isla Margarita ( = Margarita Island = Isla de Mar-
garita): Nueva Esparta, Venezuela (lTOO'N,
6400'W. Paynter, 1982)
Caluromys philander
Marmosa robinsoni, 2-425 m
Isla Maria Madre (= Maria Madre Island), Islas
Tres Man'as; Sinaloa. Mexico (2135'N,
10633'W. Geonet)
Marmosa canescens
Isla Nicolas Suarez (see Nicolas Suarez Island);
Beni. Bolivia
Isla Parida: Chiriqui. Panama (0807'N, 8220'W,
Geonet)
Caluromys derbianus
Isla Popa: Bocas del Toro. Panama (0911'N,
8207'W. Geonet)
Didelphis marsupialis
Philander opossum
Isla Puma: Colon. Panama (0914'N, 7954'W,
Geonet)
Philander opossum
Isla Puna (= Puna = Puna Island); Guayas, Ec-
uador (0250'S, 8008'W, Paynter, 1993)
Didelphis marsupialis
Marmosa robinsoni
Isla Roble. Delta de Parana: Buenos Aires, Ar-
gentina (Rio Parana ca. 3343'S, 5915'W,
Paynter. 1985)
Gracilinanus agilis, near sea level
Isla Saboga (= Saboga Island); Panama, Panama
(0837'N, 7904'W, Geonet)
Marmosa robinsoni
Isla San Cristobal; Bocas del Toro, Panama
(0915'N. 8216'W, Geonet)
Philander opossum
Isla San Miguel (= San Miguel Island); Panama,
Panama (0823'N. 7856'W. Gardner, 1973)
228
FIELDIANA: ZOOLOGY
Didelphis marsupialis
Marmosa robinsoni
Islas del Mafz (= Corn Islands); Zelaya, Nicara-
gua (1210'N, 8302'W, Geonet)
Marmosa mexicana
Islas de Ometepe (see Ometepe); Rivas; Nicara-
gua
Islas Tres Marias; Sinaloa, Mexico (2125'N,
10628'W, Geonet)
Marmosa canescens
Isla Victoria, Lago Nahuel Huapi; Neuquen, Ar-
gentina (4056'S, 7133'W, USBGN)
Dromiciops gliroides
Istmo Ameghino; Chubut, Argentina (4225'S,
6435'W, Mares and Braun, 2000)
Thylamys pallidior
Itabana al Sur de las Vegas; Cojedes, Venezuela
(0930'N, 6836'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Itabira; Minas Gerais, Brazil (1938'S, 4312'W,
Varejao and Valle, 1982)
Didelphis albiventris, 763 m
Didelphis aurita, 763 m
Itabirito; Minas Gerais, Brazil (2015'S, 4348'W,
USBGN)
Didelphis aurita
Itabu Creek Head (see Boundary Camp); Upper
Takutu-Upper Essequibo, Guyana
Itabuna, 25 km W of Ilheus (= Fazenda Beijo
Grande); Bahia, Brazil (1448'S, 3916'W,
Mustrangi and Patton, 1997)
Didelphis aurita
Marmosops incanus, 50 m
Metachirus nudicaudatus
Itacoatiara; Amazonas, Brazil (0308'S, 5825'W,
Paynter and Traylor, 1991)
Caluromys philander, 28 m (N bank Rio Ama-
zonas)
Didelphis marsupialis
Marmosa murina, 28 m (N bank Rio Amazon-
as)
Monodelphis brevicaudata, 28 m (N bank Rio
Amazonas)
Itacuruca (see Itacurussa); Rio de Janeiro, Brazil
Itacurussa (= Itacuruca); Rio de Janeiro, Brazil
(2255'S, 4355'W, Paynter and Traylor, 1991)
Didelphis aurita
Itahuania (= Itanhuania); Madre de Dios, Peru
(1247'S, 7113'W, Stephens and Traylor,
1983)
Caluromysiops irrupta, 450 m
Micoureus regina
Itaituba-Jacare-Acanga (= km 212); Para, Brazil
(0417'-0616'S, 5559'-5739'W, Paynter
and Traylor, 1991)
Didelphis marsupialis, 45 m
Metachirus nudicaudatus
Monodelphis emiliae
Itanhaem; Sao Paulo, Brazil (241 l'S, 4647'W,
Paynter and Traylor, 1991)
Metachirus nudicaudatus
Itanhandu; Minas Gerais, Brazil (2223'S,
4451'W, Mustrangi and Patton, 1997)
Marmosops paulensis, 1400 m
Itanhuania (see Itahuania); Madre de Dios, Peru
Itape Camp; Guaira, Paraguay (2551'S, 5638'W,
Hershkovitz, 1992)
Gracilinanus agilis, 200 m
Itapetininga; Sao Paulo, Brazil (2336'S, 4803'W,
Paynter and Traylor, 1 99 1 )
Micoureus demerarae, 641 m
Itapoama, Rio Tapajos; Para, Brazil (ca. 0315'S,
5510'W, Paynter and Traylor, 1991)
Marmosa murina
Itapocu (see Rio Itapocu); Santa Catarina, Brazil
Harare; Sao Paulo, Brazil (2407'S, 4920'W,
Paynter and Traylor, 1 99 1 )
Caluromys philander
Monodelphis unistriata, 131 m
Itati; Corrientes, Argentina (2716'S, 5815'W,
Paynter, 1985)
Didelphis albiventris
Itatiaya [Itatiaia] (see Pico das Agulhas Negras);
Rio de Janeiro, Brazil
Itaya (see Iquitos); Loreto, Peru
Itonama; Beni, Bolivia (1228'S, 6424'W, Payn-
ter, 1992)
Philander opossum
Ituanaa (see Huitanaa); Amazonas, Brazil
Ituverava; Sao Paulo, Brazil (2020'S, 4746'W,
Paynter and Traylor, 1 99 1 )
Didelphis albiventris, 635 m
Gracilinanus microtarsus, 605 m (2020'S,
47047 ' W)
Lutreolina crassicaudata, 635 m (2020'S,
4727'W)
Ituxi (= Iquiri); Acre, Brazil (0718'S, 6451'W,
Paynter and Traylor, 1991)
Didelphis marsupialis
Iucai (= Yucabf = Yucahy = Yucalf); Amazonas,
Brazil (0013'S, 6649'W, Paynter and Tray-
lor, 1991)
Didelphis marsupialis, ca. 75 m
Iutanaa (see Huitanaa); Amazonas, Brazil
Ivinheima (see Rio Ivinheima); Mato Grosso do
Sul, Brazil
BROWN: ATLAS OF NEW WORLD MARSUPIALS
229
Iwokrama Reserve; Potaro-Siparuni. Suriname
(ca. 0500'N. 59WW. Voss et al.. 2001)
Marmosa pinheiroi
Micoureus demerarae
Ixiamas: La Paz, Bolivia (1345'S, 6809'W.
Paynter. 1992)
Didelphis albiventris
Mannosops noctivagus, 222 m (13 km SW. 13
km W of Ixiamas)
Micoureus constantiae
Micoureus demerarae (13 km W of Ixiamas)
Micoureus regina
Izamal; Yucatan, Mexico (2056'N. 8901'W.
Geonet)
Marmosa mexicana
Jabaquara; Bahia. Brazil (1331'S. 3958'W. Cer-
queira. 1984)
Didelphis albiventris
Jabuticabal; Sao Paulo. Brazil (2116'S. 4819'W.
Paynter and Traylor. 1 99 1 )
Gracilinanus microtarsus, 605 m
Jacare-Acanga. W bank Rio Tapajos: Para. Brazil
(0616'S. 5739'W. Paynter and Traylor. 1991 )
Metachirus nudicaudatus
Jacarehy (see Jacarei): Sao Paulo. Brazil
Jacarei (= Jacarehy); Sao Paulo. Brazil (2319'S.
4558'W. Paynter and Traylor. 1991)
Caluromys philander
Jacarepagua; Rio de Janeiro. Brazil (2255'S.
4321'W. Paynter and Traylor. 1991)
Chironectes minimus
Jaco. 10 km N; Puntarenas. Costa Rica (0937'N.
8438'W. Geonet)
Didelphis marsupialis
Jaguanum; Rio de Janeiro. Brazil (2300'S.
" 4356'W. USBGN)
Didelphis aurita
Jaguaquara: Bahia. Brazil (1332'S. 3958'W.
Mustrangi and Patton. 1997)
Mannosops incanus. 600 m
Jainu. right bank Rio Jurua: Amazonas. Brazil
(0628'S. 6846'W. Patton et al.. 2000)
Didelphis marsupialis
Mannosops neblina
Micoureus regina
Jalapa: Nueva Segovia. Nicaragua (1355'N.
8608'W. Geonet)
Chironectes minimus, 600 m. 660 m
Didelphis marsupialis, 680 m
Philander opossum. 660 m
Jalapa: Veracruz-Llave. Mexico (1932'N.
9655'W. Geonet)
Didelphis marsupialis (5 km N of Jalapa)
Mannosa mexicana. 4200 ft (= 1300 m). 4500
ft (= 1400 m)
Jamachuma: Cochabamba. Bolivia (1732'S.
6607'W. Anderson. 1997)
Didelphis albiventris
Thylamys pallidior
Jaque: Darien. Panama (0731'N. 7810'W. Geo-
net)
Caluromys derbianus (Rio Imamado)
Chironectes minimus (Rio Imamado. Rio
Jaque)
Didelphis marsupialis (Rio Imamado)
Metachirus nudicaudatus (Rio Imamado. Rio
Jaque)
Philander opossum
Jardim Botanico: Rio de Janeiro. Brazil (ca.
22 55'S. 4312'W. Paynter and Traylor, 1991)
Didelphis aurita. ca. 1 100 m
'"Jarikaba" (= Jarikaba Kreek); Saramacca. Su-
riname (05 C 47'N. 55 D 22'W. Geonet)
Mannosa murina
Monodelphis brevicaudata
Jarikaba Kreek (see ""Jarikaba"); Saramacca. Su-
riname
Jauja: Araucania. Chile (3804'S. 7155'W. Geo-
net)
Dromiciops gliroides, 610 m
Jaunache (see Jauneche): Los Rios. Ecuador
Jauneche (= Jaunache); Los Ribs. Ecuador
(0120'S. 7935'W. Paynter. 1993)
Caluromys derbianus. 50-70 m
Chironectes minimus
Didelphis marsupialis. 50-70 m
Jenaro Herrera Biological Station; Loreto. Peru
(0455'S. 73 C 40'W. Fleck and Harder. 1995)
Philander mcilhennyi. 1 25 m
Jequie; Bahia. Brazil (1351'S, 4004'W. Paynter
and Traylor. 1991)
Didelphis albiventris. 200 m
Mannosops incanus. 200 m
Jequitinhonha: Minas Gerais. Brazil (1626'S.
41WW. Paynter and Traylor. 1991)
Didelphis aurita. 245 m
Jerico: Antioquia. Colombia (0547'N. 7547'W.
Paynter. 1997)
Micoureus alstoni. ca. 1500 m
Jesus Carranza (= Santa Lucrecia); Veracruz-Lla-
ve. Mexico (1726'N. 9502'W. Geonet)
Caluromys derbianus. 300 ft (= 90 m)
Didelphis marsupialis. 300 ft (= 90 m), 450 ft
(= 140 m)
Philander opossum. 200-300 ft (= 60-90 m)
(1726'N. 9501'W)
230
FIELDIANA: ZOOLOGY
Jico (= Xico); Veracruz-Llave, Mexico (1925'N,
9700'W, Geonet)
Didelphis marsupialis, 4200 ft (= 1300 m)
Jimba (see La Jimba); Veracruz-Llave, Mexico
Jimenez, Rio Jimenez; Limdn, Costa Rica
(1013'N, 8343'W, USBGN)
Marmosa mexicana, 700 ft (= 218 m)
Joao Pessoa; Parafba, Brazil (0707'S, 3452'W,
Paynter and Traylor, 1991)
Monodelphis americana, 45 m
Joao Pessoa (= Eirunepe = Sao Felipe), W bank
Rio Jurua; Amazonas, Brazil (0640'S,
6952'W, Paynter and Traylor, 1991)
Marmosa murina
Joinvile (= Joinville); Santa Catarina, Brazil
(2618'S, 4850'W, Paynter and Traylor, 1991)
Didelphis aurita
Gracilinanus microtarsus, sea level (near Join-
vile)
Micoureus demerarae, 4 m
Monodelphis iheringi, 4 m
Philander frenata
Joinville (see Joinvile); Santa Catarina, Brazil
Jordao Riberao (= Rio Jordao); Minas Gerais,
Brazil (1826'S, 4806'W, Paynter and Tray-
lor, 1991)
Chironectes minimus
Didelphis albiventris, 800 m
Micoureus constantiae
Jose de Melo; Minas Gerais, Brazil (1940'S,
4335'W, Varejao and Valle, 1982)
Didelphis albiventris
Jose Marti, 19 km ESE; San Juan, Argentina
(3153'S, 6802'W, Mares and Braun, 2000)
Thylamys pallidior
Jua, near Iguatu; Ceara, Brazil (Iguatu = 0622'S,
3918'W, Paynter and Traylor, 1991)
Gracilinanus agilis
Juan (probably Jua, see above), near Iguatu;
Ceara, Brazil (Iguatu = 0622'S, 3918'W,
Paynter and Traylor, 1 99 1 )
Didelphis albiventris
Juanchito; Valle del Cauca, Colombia (0328'N,
7629'W, Paynter, 1997)
Chironectes minimus, 985 m
Juan de Zalazar (see Fortin Juan de Zalazar);
Presidente Hayes, Paraguay
Juan Mina Station; Panama, Panama (Juan Mina
= 0910'N, 7939'W, Geonet)
Didelphis marsupialis (Rio Chagres)
Philander opossum (also 5 mi NE of Gamboa)
Juazeiro; Bahia, Brazil (0925'S, 4030'W, Payn-
ter and Traylor, 1 99 1 )
Didelphis albiventris, 371 m
Juiz de Fora; Minas Gerais, Brazil (2145'S,
4320'W, Paynter and Traylor, 1991)
Didelphis aurita, 667 m
Marmosops incanus, 700 m
Philander frenata
Jujuy (see San Salvador de Jujuy); Jujuy, Argen-
tina
"Juliaca" (see Inca Mines); Puno, Peru
Junction of Highway 14 and 2 de Mayo Road
(Dos de Mayo Road); Misiones, Argentina
(not located)
Didelphis aurita
Junction of Highway 21 and Arroyo Oveja Negra,
2 km W of Parque Provincial Mocona; Mi-
siones, Argentina (not located)
Didelphis aurita
Junction of Highways 50 and 18, 43.7 km N on
road to Isla de Cafias (= 43.7 km N on road
to Isla de Cafias); Salta, Argentina (2257'S,
6433'W, Flores et al., 2000)
Micoureus constantiae
Thylamys venustus
Junction of Highway 338 and road to Horco Mol-
le, 4 km W (see Highway 338 on road to Hor-
co Molle); Tucuman, Argentina
Junction of road 34 and Puesto Viejo, 4 km E;
Jujuy, Argentina (2429'S, 6459'W, Flores et
al., 2000)
Thylamys venustus
Juntas de San Antonio; Salta, Argentina (San An-
tonio = ca. 2308'S, 6420'W, Paynter, 1985)
Thylamys venustus
Juquia (= Rio Juquia), SW of Serra do Mar, N of
Iguape; Sao Paulo, Brazil (2419'S, 4736'W,
Paynter and Traylor, 1991)
Didelphis aurita
Philander frenata
Juquila; Oaxaca, Mexico (not precisely located,
Geonet lists 3 localities of same name all
around 1600'N, 9700'W)
Marmosa mexicana, 1500 m
K
Kaiserberg Airstrip (see Kayser Gebirgte Air-
strip); Nickerie, Suriname
Kalacoon (= Kalakun); Cayuni-Mazaruni, Guya-
na (0624'N, 5839'W, Stephens and Traylor,
1985)
Didelphis marsupialis, ca. 100 m
Micoureus demerarae
Kalakun (see Kalacoon); Mazaruni-Potaro, Guy-
ana
Kamakusa; Cuyuni-Mazaruni, Guyana (0557'N,
5954'W, Stephens and Traylor, 1985)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
231
Monodelphis brevicaudata
Kanavayen; Bolivar, Venezuela (0538'N,
6148'W, Geonet)
Didelphis marsupialis, 3100 ft (= 970 m)
Kanuku Mountains; Upper Takutu-Upper Esse-
quibo, Guyana (0312'N, 5935'W, Geonet)
Didelphis marsupialis, 2000 ft (= 625 m)
Marmosa murina (N of Kanuku Mts.)
Metachirus nudicaudatus, 182 m (0300'N,
5945'W)
Philander opossum, 240 ft (= 75 m)
Karanambo, Rupunumi River; Upper Takutu-Up-
per Essequibo, Guyana (0345'N, 5918'W,
Mustrangi and Patton, 1997)
Marmosops parvidens, 1 00 m
Kartabo; Mazaruni-Potaro, Guyana (0623'N,
5841'W, Stephens and Traylor, 1985, and
Times Atlas, 1992)
Didelphis marsupialis, 100 m
Marmosa murina
Metachirus nudicaudatus
Micoureus demerarae
Monodelphis brevicaudata
Kashmera (see Kasmera); Zulia, Venezuela
Kasmera, base of Serranfa de Perija; Zulia, Ve-
nezuela (1005'N, 7245'W, Perez-Hernandez,
1989)
Monodelphis palliolata, 20 m
Kasmera (= Kashmera), Rio Yasa; Zulia, Vene-
zuela (0957'N, 7245'W, Geonet)
Didelphis marsupialis, 270 m
Metachirus nudicaudatus, 270 m (1005'N,
7245'W)
Monodelphis palliolata, 272-273 m (0952'N,
7243'W)
Kates Lagoon; Cayo, Belize (1759'N, 8828'W,
Geonet)
Caluromys derbianus
Kayamas, Aguaruna Village, Rio Cenepa (see
Aguaruna Village); Amazonas, Peru
Kayser Gebirgte Airstrip (= Kaiserberg Airstrip);
Nickerie, Suriname (0310'N, 5627'W, Ste-
phens and Traylor, 1985)
Didelphis marsupialis, ca. 900 ft (= 280 m)
Marmosops parvidens, 900 ft (= 280 m)
Monodelphis brevicaudata
Philander opossum, 900 ft (= 280 m)
Kiteni; Cusco, Peru (ca. 1220'S, 7250'W, Ste-
phens and Traylor, 1983)
Didelphis marsupialis
Metachirus nudicaudatus, 1500 ft (= 460 m),
650 m ("60 km from Rosalina on road from
Quellabamba")
km 19, Itaituba-Jacareacanga; Para, Brazil
(0417'S, 5559'W, Hershkovitz, 1997)
Metachirus nudicaudatus (0420'S, 5605'W)
Philander opossum (W bank Rio Tapajos)
km 35 NW of Yucumo; Beni, Bolivia (1452'S,
6707'W, Anderson, 1997)
Gracilinanus agilis
km 42, Highway 64, S of San Pedro de Colalao;
Tucuman, Argentina (2620'S, 6533'W, Ma-
res, 1996)
Didelphis albiventris, 4700 ft (= 1465 m)
Thylamys venustus, 4700 ft (= 1465 m)
km 43, Altamira-Itaituba; Para, Brazil (0320'S,
5215'W, Hershkovitz files)
Metachirus nudicaudatus
km 43.7 NW of junction of Highways 50 and 18
on road to Isla de Canas (see Junction of
Highways 50 and 18); Salta, Argentina
km 46, Manaus-Itacoatiara; Amazonas, Brazil (ca.
0308'S, 6045'W, Hershkovitz files)
Caluromys philander
Metachirus nudicaudatus
km 50, Manaus-Itacoatiara RR; Amazonas, Brazil
(0309'S, 6043'W, Herskovitz, 1992)
Caluromys philander
Didelphis marsupialis (ca. 0308'S, 5825'W)
Gracilinanus emiliae, 100-200 m
km 51 (see Paramo de Guanacas); Cauca, Colom-
bia
km 70-80, Carretera El Dorado-Santa Elena de
Uairen; Bolfvar, Venezuela (0600'-0629'N,
6127'W, Perez-Hernandez, 1989)
Didelphis albiventris, 100 m
km 84, Santarem-Cuiaba; Para, Brazil (0232'S,
5443'W, Hershkovitz files)
Metachirus nudicaudatus
km 87-94, Belem-Brasflia (see Capim); Para,
Brazil
km 88, Carretera El Dorado-Santa Elena; Bolivar,
Venezuela (0600'N, 6127'W, Perez Hernan-
dez, 1989)
Monodelphis brevicaudata, 100 m
km 90, Rodovia Belem-Brasflia; Para, Brazil (Be-
lem = 0127'S, 4829'W, Paynter and Traylor,
1991)
Philander opossum
km 94, Belem-Brasflia; Para, Brazil (0127'S,
4829'W, Paynter and Traylor, 1991)
Philander opossum
km 121, 85 km SSE of El Dorado; Bolivar, Ve-
nezuela (0609'N, 6122'W, Pine, 1981)
Marmosops pinheiroi, 1032 m
km 124-134, Carretera El Dorado-Santa Elena de
232
FIELD1ANA: ZOOLOGY
Uairen; Bolivar, Venezuela (0556'N,
6127'W, Perez-Hernandez, 1989)
Didelphis albiventris, 1100-1350 m
Monodelphis brevicaudata, 1100-1350 m
km 125, 85 km SSE of El Dorado; Bolivar, Ve-
nezuela (0609'-0559'N, 6126'-61 22'W,
Handley, 1976)
Didelphis albiventris, 1032-1165 m
Didelphis marsupialis, 193-1165 m, 1032-
1165 m
Marmosa murina, 1032 m
Marmosops parvidens
Marmosops pinheiroi, 1032 m
Micoureus demerarae, 324-1032 m
km 212, Itaituba-Jacare-Acanga Highway (see
Itaituba-Jacare-Acanga); Para, Brazil
km 216, Estrada de Santarem-Cuiaba (BR165);
Para, Brazil (0417'S, 5559'W, Hershkovitz,
1997)
Philander opossum
km 217, Santarem-Cuiaba Highway, E bank Rio
Tapajos; Para, Brazil (0405'S, 5506'W, Pine
and Handley, 1984)
Monodelphis emiliae
km 514, Highway 20, 6 km N; San Juan, Argen-
tina (3154'S, 6804'W, Geonet)
Thylamys pallidior
Kuitaro River, 40 mi E of Dadanawa; Upper Tak-
utu-Upper Essequibo, Guyana (Dadanawa =
0250'N, 5931'W, Geonet)
Micoureus demerarae
Kunana, Sierra de Perija; Zulia, Venezuela
(0925'N, 7225'W, Perez-Hernandez, 1989)
Caluromys lanatus, 1140 m
Kurinwas River (see Rio Kurinwas); Zelaya, Nic-
aragua
Kwatamang Landing (see Quartermile Landing);
Rupununi, Guyana
La Argentina, Rio Cubugon; Boyaca, Colombia
(0703'N, 7215'W, Paynter, 1997)
Metachirus nudicaudatus, 2400 ft (= 750 m)
La Azotea (see Parque Nacional Diamante); Entre
Rios, Argentina
La Azulita; Merida, Venezuela (0843'N,
7127'W, Paynter, 1982)
Marmosa robinsoni, 3500 ft (= 1090 m)
Marmosops fuscatus, 1135m
Metachirus nudicaudatus, 3500 ft (= 1090 m)
La Batea, 5 km SW of Maron; Carabobo, Vene-
zuela (1028'N, 6813'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
La Belgica, 2 km S; Santa Cruz, Bolivia (1731'S,
6312'W, Anderson, 1997)
Philander opossum
La Blanquita; Apure, Venezuela (0717'N,
7150'W, Geonet)
Philander opossum, 24 m
La Blanquita, Parque Nacional Yacambu; Lara,
Venezuela (0942'N, 6940'W, Perez-Hernan-
dez, 1989)
Micoureus demerarae
La Boca Rio Saija (see Rio Saija); Cauca, Colom-
bia
La Cabana; Antioquia, Colombia (0718'N,
7536'W, Geonet)
Didelphis marsupialis (0710'N, 7526'W)
Marmosops parvidens, 1200-2000 m
La Calera; Managua, Nicaragua (not located, may
be La Calera in Granada, Nicaragua)
Caluromys derbianus
La Calera (= Calera); Valparaiso, Chile (3247'S,
7112'W, Paynter, 1988)
Thylamys elegans, 216 m
La Calera, Nandaime; Granada, Nicaragua (Nan-
daime = 1144'N, 8606'W, Geonet)
Caluromys derbianus
Didelphis marsupialis (near Nandaime)
Philander opossum (W of Nandaime)
La Candela; Huila, Colombia (ca. 0150'N,
7620'W, Paynter, 1997)
Micoureus regina, 6000 ft (= 1875 m)
La Carbonera (see Hacienda La Carbonera); Me-
rida, Venezuela
La Carolina, 3 mi N of Quito; Pichincha, Ecuador
(Quito = 0013'S, 7830'W, Paynter, 1993)
Caenolestes fuliginosus, 9400 ft (= 2930 m)
La Ceiba, 7 mi W; Atlantida, Honduras (1547'N,
8648'W, Geonet)
Didelphis marsupialis
La Ceiba; Trujillo, Venezuela (0928'N, 7104'W,
Paynter, 1982)
Didelphis marsupialis, 29 m
Marmosa robinsoni, 16-29 m
Monodelphis palliolata, 29 m
La Chicharras (see Las Chicharras); Chiapas,
Mexico
La Chorera; Panama, Panama (0852'N, 7948'W,
Wenzel and Tipton, 1966)
Caluromys derbianus
La Cienega, near Baruta; Miranda, Venezuela
(1027'N, 6652'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni
Micoureus demerarae, ca. 900-1000 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
233
Monodelphis palliolata
La Concepcion; Carchi, Ecuador (0036'N.
7807'W, Paynter, 1993)
Philander opossum
La Concepcion, Sierra Nevada de Santa Marta;
Magdalena, Colombia (944'N, 7425'W,
Geonet)
Marmosa robinsoni, 800 m, 3000 ft (= 930 m)
La Conchita (see Hacienda El Limon); Distrito
Federal, Venezuela
La Concordia (see Caserfo Boro); Lara, Venezuela
La Copa, 4 km NW of Montalban (see Montal-
ban); Carabobo, Venezuela
La Cruz; Valparaiso, Bolivia (1124'S, 6713'W,
Anderson, 1997)
Micoureus constantiae
La Cruz; Santiago, Chile (3250'S, 7114 f W, Geo-
net)
Thylamys elegans, ca. 100 m
La Culata (= Culata = Selva Culata). 17 km NE
of Merida; Merida, Venezuela (ca. 0844'N,
7105'W, Geonet)
Gracilinanus dryas, 4000 m
La Cumbre de Valencia; Carabobo, Venezuela (ca.
1020'N, 6800'W, Paynter, 1982)
Marmosops fuscatus, 5000 ft (= 1560 m),
1700 m
Laderas Iliniza; Pichincha, Ecuador (0040'S.
7842'W, Paynter, 1993)
Caenolestes fuliginosus
La Embocada; Beni, Bolivia (1503'S, 6658'W,
Anderson, 1997)
Didelphis marsupialis
Marmosa murina
La Erica; Barinas, Venezuela (0828'N, 7021'W,
Geonet)
Marmosa robinsoni
Metachirus nudicaudatus
La Esmeralda, Monte [Cerro] Duida; Amazonas.
Venezuela (0310'N, 6533'W, Paynter, 1982)
Caluromys lanatus, 100 m, 145 m (48 km NW
of La Esmeralda)
Micoureus demerareae
La Esperanza; Cusco, Peru (ca. 1313'S, 7125'W.
Pacheco et al., 1993)
Lestoros inca, 2850 m
La Esperanza, near San Carlos; Rio San Juan.
Nicaragua (San Carlos = 1107'N, 8447'W,
Geonet)
Didelphis marsupialis
Philander opossum (ca. 1206'N, 8600'W)
La Florida; Distrito Federal, Venezuela (1030'N,
6652'W, Geonet)
Monodelphis brevicaudata
Monodelphis palliolata, 1000 m
La Florida (= Piliguaya = Pitiguaya); La Paz, Bo-
livia (1621'S, 6746'W, Paynter, 1992)
Didelphis albiventris, 5600 ft (= 1750 m)
Didelphis marsupialis, 1750 m
Gracilinanus agilis, 5800 ft (= 1800 m)
Marmosops dorothea, 1700 m, 1750 m
Micoureus regina
La Fortuna, Cerro San Miguel; Amazonas, Ve-
nezuela (not located)
Didelphis marsupialis, 970 m
La Fria, Colonia Agropecuaria del IAN; Tachira,
Venezuela (0813'N, 7214'W, Geonet)
Caluromys lanatus
La Frijolera; Antioquia, Colombia (0710'N,
7525'W, Paynter and Traylor, 1981)
Didelphis marsupialis, 1500 m
Metachirus nudicaudatus, 5000 ft (= 1560 m)
(western slope Cordillera Central)
Lagarto; Ucayali, Peru (1040'S, 7354'W, Ste-
phens and Traylor, 1983)
Didelphis marsupialis (ca. 1041'S, 7349'W)
Marmosa lepida (Rio Ucayali)
Marmosops impavidus (Rio Ucayali)
Lagarto Cocha, right bank, upper Rio Ucayali;
Ucayali, Peru (1041'S, 7348'W, Hershko-
vitz, 1993)
Philander opossum, 250 m
Lagartococha. Rio Napo; Loreto, Peru (0039'S,
7516'W, Stephens and Traylor, 1983)
Micoureus regina
Lagoa de Juparana; Espi'rito Santo, Brazil
(1920'S, 4005'W, Hershkovitz files)
Metachirus nudicaudatus
Lagoa do Monteiro; Parafba, Brazil (Monteiro =
0753'S. 3707'W, Paynter and Traylor, 1991)
Monodelphis domestica
Lago Agrio; Napo, Ecuador (0006'S, 7654'W,
Geonet)
Caluromys lanatus (left bank Rio Aguarico)
Lagoa Santa; Minas Gerais, Brazil (1938'S,
4353'W, Paynter and Traylor, 1991)
Caluromys lanatus, 760 m
Chironectes minimus
Didelphis albiventris, 760 m, 850 m
Didelphis aurita, 850 m
Gracilinanus agilis, 760 m
Lutreolina crassicaudata
Marmosops incanus, 760 m (also Mandi)
Micoureus constantiae
Micoureus demerarae, 700 m, 760 m
Monodelphis americana, 760 m
Monodelphis domestica, 760 m
Monodelphis sorex (Rio das Velhas)
234
FIELDIANA: ZOOLOGY
Philander frenata (1940'S, 4353'W)
Thylamys velutinus, 760 m
Lago Atescatempa (see Laguna de Atescatempa);
Jutiapa, Guatemala
Lago Balzon; Bolivia (not located)
Gracilinanus agilis
Lago Calima; Valle del Cauca, Colombia
(0408'N, 7704'W, Paynter, 1997)
Didelphis marsupialis, 1450 m
Lago Catemaco; Veracruz-Llave, Mexico
(1825'N, 9505'W, USBGN)
Didelphis marsupialis
Lago Chapo, south side; Los Lagos, Chile
(4127'S, 7230'W, Geonet)
Rhyncholestes raphanurus
Lago de Maracaibo (see Cordillera de Merida);
Tachira-Merida, Venezuela
Lago de Tefe (see Santo Isidoro); Amazonas, Bra-
zil
Lago de Valencia (see Punta Cabito); Carabobo,
Venezuela
Lago Diamante (see Laguna Diamante); Mendo-
za, Argentina
Lago do Arara; Amazonas, Brazil (0326'S,
6122'W, USBGN)
Micoureus demerarae
Lago do Baptista; Amazonas, Brazil (0318'S,
5815'W, Hershkovitz, 1997)
Caluromys philander
Philander opossum
Lago do Capiranga; Amazonas, Brazil (0314'S,
6749'W, USBGN)
Micoureus demerarae
Lago do Ipixuna; Amazonas, Brazil (0352'S,
5352'W, Hershkovitz, 1997)
Philander opossum (S bank Rio Solim5es)
Lago Gatun (= Lake Gatun), Peninsula Buena
Vista; Colon, Panama (0912'N, 7955'W,
Geonet)
Marmosa robinsoni
Philander opossum
Lago Jiloa (= Laguna Jiloa); Managua, Nicaragua
(1213'N, 8619'W, USBGN)
Didelphis marsupialis
Lago Lacar; Neuquen, Argentina (4011'S,
7130'W, Paynter, 1985)
Dromiciops gliroides
Lago Meduinim, left bank Rio Negro; Amazonas,
Brazil (0147'S, 6123'W, Patton, pers.
comm.)
Didelphis marsupialis
Marmosops parvidens
Metachirus nudicaudatus
Micoureus demerarae
Philander opossum
Lago Miranes, Rio Napo (see Rio Mazan); Loreto,
Peru
Lago Muyo; Santiago del Estero, Argentina
(2842'S, 6250'W, Geonet)
Thylamys pallidior
Thylamys pusilla
Lago Nicaragua (also see Rivas); Rivas, Nicara-
gua (1126'N, 8550'W, Geonet)
Marmosa mexicana
Lago Olomega; San Miguel, El Salvador (ca.
1323'N, 8803'W, Gardner, 1973)
Didelphis marsupialis, 200 ft (= 60 m)
Philander opossum, 200 ft (= 60 m)
Lago Paraiso, Parque Nacional Puyehue; Los La-
gos, Chile (4048'S, 7217'W, Geonet)
Dromiciops gliroides, 950 m
Lago Quillen; Neuquen, Argentina (3925'S,
7118'W, Paynter, 1985)
Dromiciops gliroides, 1000 m
Lago Sandoval, Rio Madre de Dios; Madre de
Dios, Peru (Sandoval = 1237'S, 6905'W,
Geonet)
Metachirus nudicaudatus, 133 m
Philander opossum, 133 m
Lago Tapaiuna (see Lago Tapayuna); Para, Brazil
Lago Tapayuna (= Lago Tapaiuna); Para, Brazil
(0323'S, 5816'W, Paynter and Tray lor,
1991)
Monodelphis emiliae, 25 m
Lago Todos Los Santos; Los Lagos, Chile
(4106'S, 7215'W, Geonet)
Dromiciops gliroides, ca. 200 m
Lago Vai-Quem-Quer (= Vai-Quem-Quer), right
bank Rio Jurua; Amazonas, Brazil (0319'S,
6601'W, Patton et al., 2000)
Didelphis marsupialis
Marmosops noctivagus
Marmosops parvidens
Metachirus nudicaudatus
Micoureus demerarae
La Granja, Itenez; Beni, Bolivia (1239'S,
6410'W, Geonet)
Monodelphis domestica
Monodelphis kunsi, below 200 m (W bank of
Rio Itonamas, 4 km N of Magdalena)
La Grita; Tachira, Venezuela (0808'N, 7158'W,
Geonet)
Didelphis marsupialis
La Gruta, Isla Colon; Bocas del Toro, Panama
(0923'N, 8216'W, Geonet)
Caluromys derbianus
La Guaira; Distrito Federal, Venezuela (1036'N,
6656'W, Paynter, 1982)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
235
Caluromys philander, sea level
Didelphis marsiipialis, sea level
Laguna (see La Laguna): Darien. Panama
Laguna (see Laguna Verde): Valparaiso. Chile
Laguna Blanca: Formosa. Argentina (2508'S.
5815'W, Geonet)
Didelphis albiventris
Lutreolina crassicaudata
Philander opossum
Laguna Chasico: Buenos Aires. Argentina
(3838'S. 6306'W. Paynter. 1985)
Didelphis albiventris, 25 m
Thylamys pallidior, 25 m
Laguna de Atescatempa (= Lago Atescatempa):
" Jutiapa. Guatemala (1412'N, 8942'W. Geo-
net)
Philander opossum
Laguna de Chacopata: Sucre. Venezuela
(1041'N. 6348'W. Lopez-Fuster et al..
2000)
Micoureus robinsoni
Laguna de Las Catas. Tartagal: Salta. Argentina
(2216'S, 6352'W. Flores et al.. 2000)
Thylamys venustus
Laguna de Los Padres. 17 km WNW of Mar del
Plata: Buenos Aires, Argentina (3750'S.
5750'W. Reig. 1964)
Didelphis albiventris
Laguna de Pozuelos: Jujuy. Argentina (2222'S.
" 6601'W. Paynter. 1985)
Thylamys pallidior
Laguna de Purace. 0.8 E of Laguna San Rafael
(0225'N. 7625'W. Paynter and Traylor.
1981); Cauca. Colombia (0224'N. 7623'W.
Patterson and Albuja. 1996)
Caenolestes fuliginosus. 3100 m. 3300 m.
3320 m
Laguna Diamante (= Lago Diamante): Mendoza.
" Argentina (3410'S. 6941'W. Paynter. 1985)
Thylamys pallidior
Laguna Gonzalez. Santa Ana: Corrientes. Argen-
" tina (Santa Ana = 2901'S. 5629'W. Paynter.
1985)
Gracilinanus agilis
Laguna Jiloa (see Lago Jiloa): Managua. Nicara-
gua
Laguna La Brea: Jujuy. Argentina (2356'S.
6428'W. Flores et al.. 2000)
Thylamys venustus
Laguna Malleco: Araucania. Chile (3813'S.
7149'W. Geonet)
Dromiciops gliroides, 1050 m
Laguna Mucubajf (see Laguna Negra): Merida.
Venezuela
Laguna Negra. Paramo de Mucubaji (= Laguna
Mucubajf): Merida. Venezuela (ca. 0846'N.
70 C 48'W. Perez-Hernandez. 1989)
Didelphis albiventris, 3500 m
Laguna San Rafael. Paramo de Purace: Cauca.
" Colombia (0225'N. 7625'W, Paynter and
Traylor. 1981)
Caenolestes fuliginosus, 3300-3320. 3375 m
Laguna Verde (= Laguna): Valparaiso. Chile
" (3306'S. 71 C 41'W. Vaynter. 1988)
Thylamys elegans, sea level
La Guneta [Laguneta]: Quindio. Colombia
(0435'N. 7530'W. Paynter and Traylor.
1981)
Caenolestes fuliginosus, 10300 ft (= 3200 m)
La Higuera: Tucuman. Argentina (2613'S.
6526'W. USBGN)
Thylamys venustus
La Horqueta. Tiara; Aragua. Venezuela (Tiara =
1008'N. 6709'W. Geonet)
Caluromys philander
La Invernada: La Rioja. Argentina (3007'S.
66 C 14'W. USBGN)
Thylamys pallidior
La Irma. Rio Abangares: Guanacaste. Costa Rica
(Rio Abangares = 1010'N. 8505'W. Geo-
net)
Didelphis marsiipialis
La Isla. near Cojoro. 37 km NNE of Paraguaipoa
in Zulia. Venezuela: La Guajira. Colombia
(Cojoro = 1138'N. 7150'W. Handley and
Gardner. 1979)
Marmosa xerophila, 5-15 m
La Jimba (= Jimba): Veracruz-Llave. Mexico
(17 C 55'N. 9525'W. Geonet)
Philander opossum, 350 ft (= 110 m)
La Joya. near Xilitla: San Luis Potosi. Mexico
(Xilitla = 2226'N. 10048'W. Geonet)
Didelphis marsiipialis
Lake Gatun. Peninsula Buena Vista (see Lago Ga-
tiin): Colon. Panama
La Laguna (= Laguna): Darien. Panama
(08%4'N. 8101'W. Geonet)
Didelphis marsiipialis
Marmosops invictus. 975 m
La Laja: Bolivar. Venezuela (0603'N. 6129'W.
Geonet)
Didelphis albiventris, 500 m
La Laja. upper Rio Orinoco: Amazonas. Venezue-
la (03 C 10'N. 6533'W. Paynter. 1982)
Marmosa murina
La Laja. Campamento MOP. 109 km S of El Do-
rado: Bolivar. Venezuela (600'N. 6127'W,
Perez-Hernandez. 1989)
236
FIELDIANA: ZOOLOGY
Monodelphis brevicaudata, 500 m
La Lata (= Estancia Los Alpes), 10 km S (La Lata
is 40 km N of Rosario); Colonia, Uruguay
(ca. 3355'S, 5723'W, Paynter, 1994)
Didelphis albiventris, ca. 150 m
La Lengueta; Barinas, Venezuela (0830'N,
7023'W, Hershkovitz, 1997)
Caluromys lanatus
Metachirus nudicaudatus
Philander opossum
La Libertad (= Libertad); Peten, Guatemala
(1647'N, 9007'W, Geonet)
Didelphis marsupialis, 540 ft (= 160 m)
Philander opossum
La Libertad, 10 mi W; Libertad, El Salvador
(1329'N, 8919'W, Geonet)
Didelphis marsupialis
La Macarena (= Serrania de Macarena); Meta,
Colombia (0245'N, 7355'W, Paynter, 1997)
Didelphis marsupialis, 300 m
Lamarao; Bahia, Brazil (1145'S, 3855'W, Payn-
ter and Traylor, 1991)
Marmosops incanus, 300 m
La Maria; Valle del Cauca, Colombia (0338'N,
7643'W, Paynter, 1997)
Didelphis marsupialis, 1400 m
Lambari; Minas Gerais, Brazil (2159'S, 4525'W,
Varejao and Valle, 1982)
Didelphis aurita
Lambayeque; Lambayeque, Peru (0642'S,
7955'W, Stephens and Traylor, 1983)
Didelphis marsupialis, 17 m
La Mendoxa; Panama, Panama (not located)
Caluromys derbianus
La Merced; Catamarca, Argentina (2810'S,
6541'W, Paynter, 1985)
Didelphis albiventris
La Merced; Junin, Peru (1103'S, 7519'W, Ste-
phens and Traylor, 1983)
Caluromys lanatus, 1000 m
Didelphis albiventris, 1200 m
La Montana, 3-4 km SE of Merida; Merida, Ve-
nezuela (0835'N, 7108'W, Geonet)
Gracilinanus dryas, 2250-2600 m
La Mucuy; Merida, Venezuela (0837'N,
7103'W, Geonet)
Chironectes minimus, 2050 m
Didelphis albiventris, 2050 m
Gracilinanus dryas
La Neblina, Cerro La Neblina (see Cerro de La
Neblina); Amazonas, Venezuela
Langamankondre; Marowijne, Suriname (0543'N,
5401'W, Geonet)
Gracilinanus emiliae
Micoureus demerarae
Monodelphis brevicaudata (left bank Marowi-
jne River)
La Palmilla (= Palmilla = Pamilla), Quillota; Val-
paraiso, Chile (3248'S, 7114'W, Geonet)
Thylamys elegans, 150 m
Lapango, below mouth of Rio Pilcomayo, S of
Asumcion; Central, Paraguay (Asuncion =
2526'S, 5740'W, Paynter, 1989)
Philander opossum
La Papaya; Manabi, Ecuador (not located)
Philander opossum, 50 m
La Paraguay, 20 km W; Bolivar, Venezuela
(0650'N, 6320'W, Paynter, 1982)
Lutreolina crassicaudata
La Pastora, 14 km E of Mirimire; Falcon, Vene-
zuela (lril'N, 6837'W, Paynter, 1982)
Caluromys philander, 122 m (near La Pastora)
Didelphis marsupialis, ca. 250 m
Marmosa robinsoni, 150 m
Marmosops cracens, 150 m (near La Pastora)
La Paz; Cordoba, Argentina (3213'S, 6503'W,
Paynter, 1985)
Didelphis albiventris
La Paz; Entre Rios, Argentina (3045'S, 5939'W,
Paynter, 1985)
Thylamys pusilla, ca. 50 m
La Paz; La Paz, Bolivia (1630'S, 6809'W, An-
derson, 1997)
Thylamys pallidior
La Paz; Mendoza, Argentina (3328'S, 6733'W,
Paynter, 1985)
Didelphis albiventris, 501 m
Thylamys pallidior
La Pedregosa (see Quebrada La Pedregosa); Me-
rida, Venezuela
La Pena; Atlantico, Colombia (1035'N, 7502'W,
Paynter, 1997)
Didelphis marsupialis, 15 m
La Picada (see Volcan Osorno); Los Lagos, Chile
La Picada Bajo (see Valle de La Picada); Los La-
gos, Chile
"La Picada Forest," Parque Nacional Vicente
Perez Rosales (= Parque Nacional Vicente
Perez Rosales = Refugio); Los Lagos, Chile
(Refugio = 4104'S, 7228'W, Geonet)
Dromiciops gliroides, 600 m
Rhyncholestes raphanurus, 450 m
La Plata; Buenos Aires, Argentina (3455'S,
5757'W, Paynter, 1985)
Didelphis albiventris
Lutreolina crassicaudata, 19 m
La Ponchera; Tachira, Venezuela (0726'N,
7152'W, Hershkovitz, 1997)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
237
Philander opossum. 1 000 m
La Poule: Saramacca. Suriname (0547'N.
5525'W, Geonet)
Caluromys philander
Didelphis marsupialis (left bank Saramacca
River)
Monodelphis brevicaudata
Philander opossum (left bank Saramacca River)
La Poza. Rio Santiago: Amazonas. Peru (04 C 03'S.
7746'W. Stephens and Tray lor. 1983)
Caluromys lanatus. 180 m
Didelphis marsupialis. 180 m (04 C 24'S.
7738'W)
Metachirus nudicaudatus. 180 m
Philander andersoni. 1 80 m
La Providencia. Turgua (see Turgua): Miranda.
Venezuela
La Purfsima: Tamaulipas. Mexico (24 C 17'N.
9928'W. Geonet)
Philander opossum (also 7 km S\V of La Pur-
fsima)
La Qiiiaca. 17 km W. 3 km S: Jujuy. Argentina
(22WS. 6544'W. Flores et al..'2000)
Thylamys pallidior
La Quinta. Rancho Grande (see Estacion Biolo-
gica de Rancho Grande): Aragua. Venezuela
La Quinta: Barinas. Venezuela (0821'N, 7033'W,
Geonet)
Metachirus nudicaudatus. 600-1070 m
Larangeiras. 20 km W; Beni. Bolivia (1313'S.
62 D 21'W. Anderson. 1997)
Didelphis marsupialis
La Reserva: La Paz. Bolivia (15'44'S. 6731'W,
Anderson. 1997)
Chironectes minimus. 840 m
Didelphis marsupialis. 840 m
Marmosops dorothea. 840 m
Marmosops panidens. 840 m
Metachirus nudicaudatus. 840 m
Micoureus constantiae. 840 m. 950 m
Monodelphis adust a. 840 m
Largarto Cocho (see Boca de Lagarto Cocha):
Napo. Ecuador
La Rusia (see Cordillera de Rusia): Bocaya/San-
tander. Colombia
Las Agiiitas. El Taficillo: Tucuman. Argentina
(26 C 42'S. 65 C 22'W. Flores et al.. 2000)
Thylamys venustus
Las Bonitas. Cano Anaru: Barinas. Venezuela
(0723'N. 70 C 44'W. Perez-Hernandez. 1989)
Philander opossum
Las Cascadas. near Gamboa; Colon. Panama
(0905'N. 7942'W. Wenzel and Tipton.
1966)
Caluromys derbianus
Las Chicharras (= Chicharras = La Chicharras):
Chiapas. Mexico (1506'N, 9215'W. Geonet)
Marmosa mexicana
Philander opossum
Las Condes: Santiago. Chile (3322'S. 7031'W.
Geonet)
Thylamys elegans
Las Cruces: Santa Cruz. Bolivia (1747'S.
6322'W. Anderson. 1997)
Marmosa murina. 480 m
Las Cuevas: Entre Rfos. Argentina (3221'S.
60 C 29'W. Geonet)
Lutreolina crassicaudata
Las Dos Bocas. La Sierra: Carabobo. Venezuela
(09 = 57'N. 67 C 59'W. Geonet)
Didelphis marsupialis
La Selva Biological Reserve (see Estacion Biol-
ogica de La Selva): Heredia. Costa Rica
La Selva. Bogota (= Estancia La Selva): Cundi-
namarca. Colombia (Bogota = 04 C 36'N.
74"05 W. Paynter. 1997)
Caenolestes fuliginosus
Gracilinanus marica (near Bogota)
Las Flores. Gracias (see Las Flores Pucca): Lem-
pfra. Honduras
Las Flores Pucca (= Las Flores. Gracias): Lem-
pfra. Honduras (1442'N. 88 C 37'W, Geonet)
Chironectes minimus
Didelphis marsupialis
Las Heras: Mendoza. Argentina (3251'S.
68"49'W. Paynter, 1985)
Thylamys pallidior
Las Higuerillas. 5 km S on Highway 9: Catamar-
ca. Argentina (27 r 47'S. 65 C 51'W. Mares et
al.. 1997)
Thylamys venustus. 2900 ft (= 900 m). 3580 ft
(= 1120 m)
La Sierra. Margarita: Nueva Esparta. Venezuela
(1100'N. 64 C 00'W. Perez-Hemandez. 1989)
Marmosa robinsoni. 100 m
Las Lajas: Neuquen. Argentina (3931'S.
70 C 22'W. Paynter. 1985)
Thylamys pallidior. 640 m
Las Lomitas: Formosa. Argentina (2442'S.
60 C 36'W. Paynter. 1985)
Didelphis albiventris. 135 m
Las Lomitas: Santa Cruz. Bolivia (1753'S.
63 C 07'W. Anderson. 1997)
Gracilinanus agilis
Marmosops dorothea (1755'S. 6308'W)
Las Lomitas (see Lomitas): Valle del Cauca. Co-
lombia
238
FIELDIANA: ZOOLOGY
Las Maquinas, Santo Domingo Trail; Pichincha,
Ecuador (0026'S, 7844'W, Bublitz, 1987)
Caenolestes convelatus, 7000 ft (= 2180 m)
Caenolestes fuliginosus
Didelphis albiventris, 7000 ft (= 2180 m)
Las Marias, 10 km NW del Paso Las Marias, 6
km SE of Melo; Cerro Largo, Uruguay
(3222'S, 5411'W, Paynter, 1994)
Didelphis albiventris
Las Marimondas (see Marimonda); La Guajira,
Colombia
Las Mesadas, Trancas; Tucuman, Argentina
(2627'S, 6530'W, Flores et al., 2000)
Thylamys pallidior, 2000 m
Thylamys pusilla
Las Mesas, 17 km NE of San Juan de Colon;
Tachira, Venezuela (0810'N, 7210'W, Han-
dley, 1976)
Didelphis marsupialis, 460 m
Las Minas; Veracruz-Llave, Mexico (1942'N,
9707'W, Geonet)
Didelphis marsupialis
Las Palmas; Chaco, Argentina (2704'S, 5842'W,
Paynter, 1985)
Gracilinanus agilis
Las Palmitas (see Guanico); Los Santos, Panama
Las Piedras; Canelones, Uruguay (3444'S,
5613'W, Paynter, 1994)
Didelphis albiventris, ca. 100 m
Lutreolina eras sic audata, ca. 100 m
Las Porqueras; Merida, Venezuela (0903'N,
7037'W, Geonet)
Didelphis albiventris, 2450 m
Micoureus demerarae, 2450 m (circa de Tim-
otes)
Las Quemadas, Valle de Suaza; Huila, Colombia
(Rio Suaza = 0210'N, 7541'W, Geonet)
Didelphis marsupialis
Las Quiguas, San Esteban; Carabobo, Venezuela
(1026'N, 6800'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 250 m
Monodelphis palliolata
Las Rosas; Santa Fe, Argentina (3228'S,
6143'W, Paynter, 1985)
Lutreolina crassicaudata
Lassance; Minas Gerais, Brazil (1758'S,
4405'W, Varejao and Valle, 1982)
Didelphis albiventris, ca. 500 m
Las Ventanas, near Belen Gualcho; Ocotepeque,
Honduras (Belen Gualcho = 1429'N,
8848'W, Geonet)
Didelphis marsupialis
Las Ventanas, Lago Yojoa; Santa Barbara, Hon-
duras (Lago Yojoa = 1452'N, 8759'W,
Geonet)
Didelphis marsupialis
Latal; Sucre, Venezuela (ca. 1010'N, 6355'W,
Paynter, 1982)
Didelphis marsupialis, 900 m
Laterite Valley, Cerro Duida; Amazonas, Vene-
zuela (Cerro Duida = 0325'N, 6540'W,
Paynter, 1982)
Didelphis albiventris, 6200 ft (= 1930 m)
Didelphis marsupialis
La Tigra; Bolivar, Venezuela (0730'N, 6135'W,
Geonet)
Didelphis albiventris, 250 m
La Tinta (see Sierra de La Tinta); Buenos Aires,
Argentina
La Tirana (see Tamara), Rio Nechi; Antioquia,
Colombia
La Toma Capaya, Rio Marismita; Miranda, Ve-
nezuela (1026'N, 6616'W, Perez-Hernan-
dez, 1989)
Chironectes minimus, 300 m
Didelphis marsupialis, 300 m
Monodelphis palliolata, 300 m
La Torrecita (see Urdampilleta); Buenos Aires,
Argentina
La Trinidad, Hato La Trinidad de Arauca; Apure,
Venezuela (071 l'N, 6905'W, Perez-Hernan-
dez, 1989)
Marmosa robinsoni
La Trinite Montains; French Guiana (0437'N,
5322'W, Francois Catzeflis, pers. com.)
Philander opossum
La Trinitaria (see Zapaluta); Chiapas, Mexico
La Tuna (see Isla de la Tuna); Rocha, Uruguay
La Tuxpena (= Chompoton); Campeche, Mexico
(1827'N, 9005'W, Geonet)
Philander opossum
La Union; Miranda, Venezuela (1024'N,
6647'W, Geonet)
Didelphis marsupialis
Marsmosa robinsoni (El Hatillo)
Monodelphis palliolata (El Hatillo)
La Union, left bank Rio Caura; Bolfvar, Venezuela
(0655'N, 6455'W, Paynter, 1982)
Marmosa murina, ca. 200 m
La Union (= Union); Los Lagos, Chile (4017'S,
7305'W, Geonet)
Dromiciops gliroides, 29 m
La Urbana (see Estancia Tapikiole); Formosa, Ar-
gentina
La Urbana Vieja (see Estancia Tapikiole); For-
mosa, Argentina
Lavala (see Matagalpa); Matagalpa, Nicaragua
BROWN: ATLAS OF NEW WORLD MARSUPIALS
239
Lavalle [La Valle]; Santiago del Estero, Argentina
(2812'S, 6508'W, Paynter, 1985)
Thylamys pallidior, 463 m
Thylamys pusilla
Lava Pes, Facao Road, Sao Luis de Caceres; Mato
Grosso, Brazil (Sao Luis de Caceres =
1604'S, 5741'W, Paynter and Traylor, 1991)
Monodelphis domestica
La Vega, E of Bogota; Cundinamarca, Colombia
(0500'N, 7421'W, Paynter and Traylor,
1981)
Chironectes minimus
La Vega del Rio Chama (Rio Chama = 0903'N,
7137'W, Paynter, 1982); Merida, Venezuela
(3 La Vega localities given, one nearest to
Rio at 0830'N, 7118'W, Geonet; ca.
0833'N, 7113'W, Hershkovitz files)
Didelphis marsupialis, 1400 m
La Venta; Tabasco, Mexico (1806'N, 9401'W,
Gardner, 1973)
Caluromys derbianus
Didelphis marsupialis
Philander opossum
Lazaropolis (see also Belem); Para, Brazil (not lo-
cated)
Metachirus nudicaudatus
Lazaropolis do Prata (see Igarape Acu); Para, Bra-
zil
La Zumbadora (see Cerro Azul); Panama, Panama
Leguia, Rio Pachitea; Loreto, Peru (not located)
Marmosops impavidus
Lelydorp (see Lelydorpplan); Para, Suriname
Lelydorp; Wanica, Suriname (0542'N, 5514'W,
Geonet)
Marmosa murina
Philander opossum
Lelydorplan (see Lelydorpplan); Para, Suriname
Lelydorpplan (= Lelydorp = Lelydorpplan); Para,
Suriname (0537'N, 5512'W, Geonet)
Didelphis marsupialis (0545'N, 5516'W)
Marmosa murina
Philander opossum
Leon; Jujuy, Argentina (2403'S, 6526'W, Payn-
ter, 1985)
Thylamys venustus, 1500 m, 1622 m, 5800 ft
(= 1800 m)
Leopoldina; Minas Gerais, Brazil (2128'S,
4240'W, Varejao and Valle, 1982)
Didelphis aurita
Les Nouragues; French Guiana (ca. 0415'N,
5245'W, Voss et al., 2001)
Marmosa murina
Leticia; Amazonas, Colombia (0409'N, 6957'W,
Paynter, 1997)
Caluromysiops irrupta, 100 m
Libertad (see La Libertad); Peten, Guatemala
Libertad; Loreto, Peru (0351'S, 7404'W, Geo-
net)
Metachirus nudicaudatus, 120 m
Lilhue-Calel; La Pampa, Argentina (3800'S,
6534'W, Paynter, 1985)
Lestodelphys halli, ca. 200 m
Lima; Lima, Peru (1203'S, 7703'W, Stephens
and Traylor, 1983)
Didelphis albiventris
Limache; Valparaiso, Chile (3301'S, 7116'W,
Geonet)
Thylamys elegans, ca. 85 m
Limacpunco, near Hacienda Cadena; Cusco, Peru
(1328'S, 7055'W, Bublitz, 1987)
Didelphis albiventris, 2400 m
Lestoros inca, 2400 m
Limao, right bank Rio Cotinga; Roraima, Brazil
(0356'N, 6030'W, Paynter and Traylor,
1991)
Monodelphis brevicaudata
Limao, Rio Jurua; Mato Grosso, Brazil (0848'S,
5732'W, USBGN)
Philander opossum (E and W banks)
Lima Pareja, 4 km SW of Pueblo Viejo; Los Ribs,
Ecuador (Pueblo Viejo = 0134'S, 7930'W,
Paynter, 1993)
Philander opossum
Limbani; Puno, Peru (1408'S, 6942'W, Stephens
and Traylor, 1983)
Didelphis albiventris, 3350 m
Lestoros inca, 8800-9500 ft (= 2750-2960 m)
(3 mi N of Puno)
Limoal, near Igarape Amorin; Para, Brazil (not
located)
Didelphis marsupialis
Limoeira da Anadia; Alagoas, Brazil (0944'S,
3631'W, USBGN)
Monodelphis domestica (Sitio Barracao)
Limon; Ecuador (not located)
Philander opossum, 1200 ft (= 375 m)
Limon; Los Rios, Ecuador (0109'S, 7929'W),
Geonet
Didelphis marsupialis
Limona; Loja, Ecuador (not precisely located)
Didelphis marsupialis
Limoncocha; Napo, Ecuador (0024'S, 7637'W,
Geonet)
Caluromys lanatus
Didelphis marsupialis, 300 m
Marmosops noctivagus
Metachirus nudicaudatus, 300 m
Micoureus regina
240
FIELDIANA: ZOOLOGY
Philander andersoni
Limontuba (see Igarape Amorin); Para, Brazil
Linhares; Espirito Santo, Brazil (1925'S,
4004'W, Paynter and Tray lor, 1991)
Metachirus nudicaudatus
Lins; Sao Paulo, Brazil (2140'S, 4945'W, Payn-
ter and Traylor, 1991)
Metachirus nudicaudatus
Lion Hill; Colon/Panama, Panama (0913'N,
7954'W, Wenzel and Tipton, 1966)
Didelphis marsupialis
Llamachaque, near Pelechuco; La Paz, Bolivia
(Pelechuco = 1448'S, 6905'W, Geonet)
Lestoros inca, 3150 m
Llano de Merida; Merida, Venezuela (0800'N,
7110'W, Hershkovitz, 1992)
Gracilinanus marica
Llera, Santa Isabel; Tamaulipas, Mexico
(2319'N, 9901'W, Geonet)
Didelphis marsupialis, 2000 ft (= 625 m)
Lloa, 3.5 km NE, Rio Condor Huachana; Pichin-
cha, Ecuador (0015'S, 7835'W, Paynter,
1993)
Caenolestes fuliginosus
Llunchi, Rio Napo; Napo, Ecuador (0037'S,
7646'W, Hershkovitz files)
Philander andersoni
Loja; Loja, Ecuador (0400'S, 7913'W, Paynter,
1993)
Didelphis albiventris, 2200 m
Loksie Hattie; Brokopondo, Suriname (0509'N,
5528'W, Hershkovitz, 1997)
Didelphis marsupialis (Saramacca River)
Philander opossum
Loma Cana, near Cerro Pirre; Darien, Panama
(Cerro Pirre = 0751'N, 7744'W, Geonet)
Marmosops impavidus, 4900 ft (= 1530 m)
Loma de Famatima (= Cerro de Famatina); La
Rioja, Argentina (2850'S, 6727'W, Paynter,
1985)
Thylamys pallidior, 3200 m
Loma Jincho Asjata; Oruro, Bolivia (1803'S,
6856'W, Anderson, 1997)
Thylamys pallidior
Lomitas (= Las Lomitas); Valle del Cauca, Co-
lombia (0338'N, 7638'W, Paynter, 1997)
Didelphis marsupialis, 4526 ft (= 1415 m)
Marmosa robinsoni, 1400 m
Loncovaca; Mendoza, Argentina (3639'S,
6839'W, Geonet)
Thylamys pallidior
Longatoma; Valparaiso, Chile (3222'S, 7118'W,
Paynter, 1988)
Thylamys elegans, 23 m
Long Bay, Isla Bastimentos; Bocas del Toro, Pan-
ama (0923'-0906'N, 8135'-8214'W,
USBGN)
Caluromys derbianus
Lonquimay; Araucania, Chile (3826'S, 7114'W,
Geonet)
Dromiciops gliroides, 915 m, 1340 m, 1520 m
Los Amates; Izabal, Guatemala (1516'N,
8906'W, Geonet)
Philander opossum
Los Campamentos, Patenemo; Carabobo, Vene-
zuela (1026'N, 6800'W, Perez-Hernandez,
1989)
Didelphis marsupialis
Monodelphis palliolata
Los Canales le Naiguata, Parque Nacional El Avi-
la; Distrito Federal, Venezuela (1036'N,
6644'W, Perez-Hernandez, 1989)
Marmosa robinsoni, 800 m
Monodelphis palliolata, 800 m
Los Chorros, Caracas; Distrito Federal, Venezuela
(1030'N, 6655'W, Perez-Hernandez, 1989)
Didelphis marsupialis, ca. 980 m
Los Chupones, Rio San Carlos; Cojedes, Vene-
zuela (0945'N, 6838'W, Geonet)
Didelphis marsupialis, 380 m
Los Cocos, 14 km S of Boaco; Boaco, Nicaragua
(1206'N, 8554'W, USBGN)
Caluromys derbianus
Chironectes minimus, 220 m
Los Colorados, 17 km E of Santo Domingo; Salta,
Argentina (2437'S, 6320'W, Flores et al.,
2000)
Thylamys pusilla
Los Dos Caminos (= Dos Caminos); Guarico, Ve-
nezuela (0952'N, 6720'W, Perez-Hernan-
dez, 1989)
Caluromys philander
Didelphis marsupialis
Marmosa robinsoni
Los Encuentros, 4 km ENE; Zamora-Chinchipe,
Ecuador (not located)
Metachirus nudicaudatus
Los Guayabitos; Miranda, Venezuela (1022'N,
6645'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 1 360 m
Marmosa robinsoni
Monodelphis palliolata, 1 360 m
Los Giiires; Delta Amacuro Territorio Federal,
Venezuela (0914'N, 6153'W, Geonet)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
BROWN: ATLAS OF NEW WORLD MARSUPIALS
241
Los Ingleses (see Estancia Los Ingleses); Buenos
Aires, Argentina
Los Manantiales (see Estancia Los Manantiales);
Chubut, Argentina
Los Menucos, 15 km SE; Rio Negro, Argentina
(4050'S, 6808'W, Paynter, 1985)
Lestodelphys halli
Los Micos, San Juan de Arama; Meta, Colombia
(0317'N, 7353'W, Paynter, 1997)
Chironectes minimus, 400 m
Gracilinanus emiliae, 396 m
Metachirus nudicoudatus, 1300 ft (= 400 m)
(Rio Giiejar)
Los Palmares; Santa Cruz, Bolivia (1525'S,
6100'W, Anderson, 1997)
Didelphis marsupialis
Philander opossum
Los Parques Nacionales Conguillio; Araucania,
Chile (3842'S, 7139'W, Jimenez and Ra-
geot, 1979)
Dromiciops gliroides
Los Patos, 25 km SE of El Manteco; Bolivar, Ve-
nezuela (OTll'N, 6222'W, Handley, 1976)
Caluromys philander, 150 m
Marmosa murina, 350 m
Los Pozos, near Arenillas; El Oro, Ecuador (Ar-
enillas = 0333'S, 8004'W, Paynter, 1993)
Marmosa robins on i
Los Reyes; Michoacan de Ocampo, Mexico
(1935'N, 10230'W, Geonet)
Marmosa canescens, 4000 ft (= 1250 m)
Los Romanos; Tucuman, Argentina (2724'S,
6508'W, Mares et al., 1996)
Didelphis albiventris
Los Sarmientos (see Sarmiento); Tucuman, Ar-
gentina
Los Tajibos (= Tojibos), 40 km E of San Borja;
Beni, Bolivia (1450'S, 6622'W, Anderson,
1997)
Caluromys lanatus
Los Teques; Miranda, Venezuela (1021'N,
6702'W, Paynter, 1982)
Didelphis marsupialis
Marmosops fu scat us
Los Totumu; Beni, Bolivia (1400'S, 6350'W,
Anderson, 1997)
Didelphis marsupialis
Los Venados, Parque Nacional El Avila; Distrito
Federal, Venezuela (1033'N, 6654'W, Geo-
net)
Caluromys philander, 1500 m, 1600 m (Monte
Avila)
Didelphis marsupialis, 1400 m, 1475 m
Gracilinanus marica, 1400 m, 1500 m, 1739 m
(1032'N, 6654'W)
Marmosops fuscatus, 1443-1500 m
Los Vilos, 4 km S; Coquimbo, Chile (3154'S,
7130'W, Geonet)
Thylamys elegans, sea level
Los Yngleses (see Estancia Los Ingleses); Buenos
Aires, Argentina
Lota; Biobio, Chile (3705'S, 7310'W, Geonet)
Dromiciops gliroides, sea level
Luca; Oruro, Bolivia (1936'S, 6753'W, Ander-
son, 1997)
Thylamys pallidior
Lucie River; Sipaliwini, Suriname (0335'N,
5740'W, Geonet)
Didelphis marsupialis
Luisiana (= Hacienda Luisiana); Ayacucho, Peru
(ca. 1239'S, 7344'W, Stephens and Traylor,
1983)
Didelphis marsupialis, 500 m
Metachirus nudicaudatus, 300 m, 1900 ft ( =
590 m) (left bank Rio Apurimac)
Lujan de Cuyo; Mendoza, Argentina (3426'S,
5832'W, Paynter, 1985)
Thylamys pallidior
Lules; Tucuman, Argentina (2656'S, 6523'W,
Mares et al., 1996)
Thylamys venustus
Lumbrera; Salta, Argentina (2512'S, 6455'W,
Geonet)
Didelphis albiventris
Luque; Central, Paraguay (2516'S, 5734'W,
Paynter, 1989)
Lutreolina crassicaudata
Philander opossum (17 km E of Luque)
Luya; Amazonas, Peru (0611'S, 7754'W, Geo-
net)
Didelphis marsupialis, 3600 ft (= 1 125 m)
Luz; Minas Gerais, Brazil (1957'S, 4510'W, Va-
rejao and Valle, 1982)
Didelphis albiventris
M
Macaco, left bank Rio Jau; Amazonas, Brazil
(0205'S, 6207'W, Patton, pers. comm.)
Marmosops noctivagus
Metachirus nudicaudatus
Micoureus demerarae
Philander andersoni
Macaco Secco, near Serra Andaraf (= Andarahy);
Bahia, Brazil, (Serra Andaraf = 1248'S,
4120'W, Paynter and Traylor, 1991)
Didelphis albiventris, ca. 366 m
242
FIELDIANA: ZOOLOGY
Macapa (= Belem Virus Laboratory); Amapa,
Brazil (0002'N, 5103'W, Paynter and Tray-
lor, 1991)
Didelphis marsupialis, 15 m
Marmosa murina, 15 m
Philander opossum (Canal del Norte)
Macchu Pichu, Ri'o Urubamba; Cusco, Peru
(1307'S, 7234'W, Stephens and Traylor,
1983)
Didelphis albiventris, 10000-14000 ft (=
3125-4375 m)
Lestoros inca, 12000-14000 ft (= 3750-
4375 m)
Marmosops noctivagus, 2060 m
Macchu [Machu] Pichu [Picchu], Urubamba,
Cedrobamba Ruins; Cusco, Peru (1315'S,
7233'W, Albuja and Patterson, 1996)
Lestoros inca, 12000 ft (= 3750 m)
Macchu [Machu] Pichu [Picchu], Urubamba,
Runcaraccay Ruins; Cusco, Peru (ca.
1314'S, 7229'W, Stephans and Traylor,
1983)
Lestoros inca, 13000 ft (= 4000 m)
Maceira [Macieiras], Itatiaya [Itatiaia] (see Pico
das Agulhas Negras); Rio de Janeiro, Brazil
Machado Portela; Bahia, Brazil (1309'S,
4045'W, USBGN)
Monodelphis domestica
Machado; Minas Gerais, Brazil (1957'S,
4510'W, Varejao and Valle, 1982)
Didelphis albiventris
Machiques, Kasmera; Zulia, Venezuela (1004'N,
7236'W, Geonet)
Didelphis marsupialis, 320 m
Metachirus nudicaudatus, 1150 m
Macieiras, Pico das Agulhas Negras (see Pico das
Aguulhas Negras); Rio de Janeiro, Brazil
Mackerie (= Makerie), West River, Wilhelmina
Mts.; Nickerie, Suriname 0326'N, 5645'W,
Hershkovitz, 1997)
Philander opossum, 3000 ft (= 930 m)
Macuspana; Tabasco, Mexico (1746'N, 9236'W,
Geonet)
Didelphis marsupialis
Philander opossum (19 mi N, 5 mi SW of Ma-
cuspana)
Madden Road; Panama, Panama (ca. 0905'N,
7938'W, Gardner, 1973)
Didelphis marsupialis
Philander opossum
Mafil; Los Lagos, Chile (3939'S, 7257'W, Geo-
net)
Dromiciops gliroides
Magdalena; Beni, Bolivia (1320'S, 6408'W,
Paynter, 1992)
Philander opossum
Maicolpue, 2 km S of Bahia Mansa; Los Lagos,
Chile (Bahia Mansa = 4033'S, 7346'W,
Geonet)
Dromiciops gliroides, 50-110 m
Rhyncholestes raphanurus, 100 m, 110 m (2
km S of Bahia Mansa)
Maimara; Jujuy, Argentina (2337'S, 6524'W,
Paynter, 1985)
Thylamys pallidior, 2230 m, 2300 m, 2500 m
Maingot Estates, 5 mi up Sangre Grande River;
St. Andrew, Trinidad and Tobago (Sangre
Grande River = 1035'N, 6105'W, USBGN)
Didelphis marsupialis
Makerie, West River, Wilhelmina Mts. (see Mack-
erie); Nickerie, Suriname
Malabrigo; Santa Fe, Argentina (2921'S,
5959'W, Geonet)
Didelphis albiventris
Lutreolina crassicaudata
Mala Mala; Tucuman, Argentina (2647'S,
6533'W, Mares et at, 1966)
Didelphis albiventris
Malargue, 12.8 km E; Mendoza, Argentina
(3529'S, 6926'W, Mares and Braun, 2000)
Thylamys pallidior
Maldonado; Maldonado, Uruguay (3454'S,
5457'W, Paynter, 1994)
Didelphis albiventris
Lutreolina crassicaudata
Monodelphis dimidiata
Mamatoco; Magdalena, Colombia (1114'N,
7410'W Paynter, 1997)
Marmosa robinsoni, ca. 25 m
Mambita; Cundinamarca, Colombia (0446'N,
7319'W, Paynter, 1997)
Marmosops impavidus, ca. 1000 m
Metachirus nudicaudatus (S of Rio Guavio)
Mamore (see Rio Mamore); Beni, Bolivia
Manacal; Sucre, Venezuela (1036'N, 6228'W,
Geonet)
Caluromys philander, 180-575 m
Didelphis marsupialis, 185-470 m
Monodelphis palliolata, 415-417 m
Manacapuru, near mouth Rio Negro; Amazonas,
Brazil (0318'S, 6037'W, Paynter and Tray-
lor, 1991)
Marmosa murina, 34 m
Managua; Managua, Nicaragua (1209'N,
8617'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
BROWN: ATLAS OF NEW WORLD MARSUPIALS
243
Mannosa mexicana
Philander opossum
Manaos (see Manaus); Amazonas, Brazil
Manaus (= Barra do Rio Negro = Manaos):
Amazonas, Brazil (0308'S, 6001'W, Paynter
and Traylor. 1991)
Caluromys lanatus (left bank Rio Negro)
Caluromys philander (left bank Rio Negro)
Didelphis marsupialis
Metachirus nudicaudatus
Micoureus demerarae 21 m (also 80 km S of
Manaus)
Monodelphis brevicaudata, 21 m (left bank Rio
Negro)
Mandi. Lagoa Santa (see Lagoa Santa): Minas
Gerais. Brazil
Mandinga; San Bias. Panama (0928'N. 79 C 05'W.
Geonet)
Chironectes minimus
Didelphis marsupialis
Metachirus nudicaudatus
Mangabeira (= Mangabeiras): Alagoas. Brazil,
(ca. 0955'S. 3608'W. Paynter and Traylor.
1991)
Caluromys philander
Didelphis aurita
Mannosa murina, sea level
Mangabeiras (see Mangabeira): Alagoas. Brazil
Mangaratiba. (= Ibicuf = Ibicuhy): Rio de Janei-
ro. Brazil (2257'S. 4402'W, Paynter and
Traylor, 1991)
Mannosops in can us, 50 m
Metachirus nudicaudatus, 50 m (Ciadade Mi-
neradoras Brasileiras Reunidos)
Philander frenata
Manhua^u; Minas Gerais. Brazil (2015'S.
4210'W, Varejao and Valle. 1982)
Didelphis aurita
Manicore; French Guaina (not located)
Didelphis marsupialis
Manu: Madre de Dios. Peru (1215'S. 7050'W.
Stephens and Traylor. 1983)
Chironectes minimus, 365 m
Manu National Park (see Parque Nacional del
Manu); Madre de Dios-Cusco. Peru
Manuinhos: Brazil (not located)
Didelphis marsupialis
Manuripi Health National Wild Life Reserve:
Pando. Bolivia (1 1 = 50'S. 6845'W. Anderson.
1997)
Didelphis marsupialis
Mapastepec: Chiapas, Mexico (1526'N. 9254'W.
Geonet)
Philander opossum, 8 m
Mapenna Creek, ca. 6 km from Corentyne River:
East Berbice-Corentyne, Guyana (0523'N,
5722'W, Patton. pers. comm.)
Didelphis marsupialis
Mapiri: La Paz. Bolivia (1515'S, 6810'W, Payn-
ter. 1992)
Didelphis marsupialis, 491 m
Mannosops dorothea, 491 m, 2000 ft (= 625 m)
Mannosops noctivagus
Metachirus nudicaudatus, 2000 ft (= 625 m)
Micoureus regina, 2000 ft (= 625 m)
Maraba: Para. Brazil (0521'S, 4907'W, Paynter
and Traylor. 1991)
Caluromys philander
Philander opossum (left bank Rio Tocantins.
0520'S. 4910'W)
Maraca. Serra do Navio (see Serra do Navio);
Amapa. Brazil
Maracaibo: Zulia. Venezuela (1040'N. 7137'W,
Geonet)
Metachirus nudicaudatus
Maracaju; Mato Grosso do Sul. Brazil (2138'S,
55WW. USBGN)
Didelphis albiventris, 500 m
Lutreolina crassicaudata
Maracas Valley. St. George: Port-of-Spain. Trini-
dad and Tobago (ca. 10 o 41'N. 6124'W, Geo-
net)
Mannosops fuscatus
Maracay: Aragua. Venezuela (1015'N. 6736'W,
Paynter. 1982)
Chironectes minimus, 445 m
Didelphis marsupialis, 445 m
Micoureus demerarae, 445 m
Marai (= Igarape Maroi). right bank Rio Tapajos;
Para. Brazil (0251'S, 5503'W, Patterson,
1992)
Monodelphis brevicaudata
Marajo (see Ilha de Marajo); Para. Brazil
Maranhao (see Sao Luis): Maranhao, Brazil
Marcapata: Cusco. Peru (1330'S. 7055'W, Ste-
phens and Traylor. 1983)
Chironectes minimus
Didelphis albiventris, 9000 ft ( = 2800 m), 3150
m (13 C 30'S. 7054'W)
Lestoros inca, 9000 ft (= 2800 m), 2812 m
Mannosa murina
Mannosops noctivagus, 3150 m
Marco (suburb of Belem) (see Belem): Para. Bra-
zil
Mar del Plata (Cuidad): Buenos Aires. Argentina
(3800'S. 5733'W. Paynter, 1985)
Didelphis albiventris, sea level (Baliza Caviu)
Monodelphis dimidiata (Baliza Cavivi)
244
FIELDIANA: ZOOLOGY
Mar del Sur, 12.5 km SW of Miramar (see Mir-
amar); Buenos Aires, Argentina
Mar del Sur; Buenos Aires, Argentina (3820'S,
5759'W, USBGN)
Didelphis albiventris
Lutreolina crassicaudata
Ma Retraite, W bank Suriname River; Paramari-
bo, Suriname (0552'N, 5508'W, Geonet)
Caluromys philander
Margarita Island (see Isla Margarita); Nueva Es-
parta, Venezuela
Maria Madre Island (see Isla Maria Madre); Sin-
aloa, Mexico
Marian; Napo, Ecuador (0002'S, 7620'W, Geo-
net)
Caluromys lanatus
Didelphis albiventris
Mariana; Minas Gerais, Brazil (2023'S, 4325'W,
Paynter and Tray lor, 1991)
Monodelphis americana
Mariano Moreno (= Moreno); Buenos Aires, Ar-
gentina (3439'S, 5848'W, Paynter, 1985)
Lutreolina crassicaudata
Mariato; Veragua, Panama (0743'N, 8058'W,
Geonet)
Marmosa mexicana
Marica; Rio de Janeiro, Brazil (2255'S, 4249'W,
Geonet)
Philander frenata
Marienburg, left bank Commewijne River; Com-
mewijne, Suriname (0553'N, 5503'W, Geo-
net)
Caluromys philander
Marimonda (= Las Marimondas), Serrania de
Perija; La Guajira, Colombia (ca. 1035'N,
7245'W, Paynter, 1997)
Caluromys lanatus
Didelphis marsupialis, 240 m, 312 m, 1000 m
(1037'N, 7258'W)
Gracilinanus marica, 1000 m
Gracilinanus perijae, 1450 m (1052'N,
7243'W)
Marmosa robinsoni, ca. 1000 m
Maripa; Bolivar, Venezuela (0726'N, 6509'W,
Paynter, 1982)
Caluromys philander, ca. 100 m (right bank
Rio Caura)
Metachirus nudicaudatus (E bank Rio Caura)
Philander opossum (right bank Rio Caura)
Mariquita; Tolima, Colombia (0511'N, 7454'W,
Geonet)
Marmosa robinsoni, 535 m
Marmato, Valle de Suaza; Huila, Colombia (Rio
Suaza = 0210'N, 7541'W, USBGN)
Didelphis marsupialis
Maroba (see Nova Vicosa); Bahia, Brazil
Maromba (see Pico das Agulhas Negras); Rio de
Janeiro, Brazil
Maroni (see Maroni River); Marowijne, Suriname
Maroni River (= Maroni); Marowijne, Suriname
(0545'N, 5358'W, Geonet)
Marmosa murina
Masagua, Finca Santillana del Esquintla; Esquin-
tla, Guatemala (1412'N, 9051'W, Geonet)
Philander opossum
Masaya; Masaya, Nicaragua (1158'N, 8606'W,
Geonet)
Caluromys derbianus
Macuspana; Tabasco, Mexico (1745'N, 9236'W,
Geonet)
Didelphis marsupialis
Masisea; Ucayali, Peru (0836'S, 7419'W, Ste-
phens and Traylor, 1983)
Caluromys lanatus, 225 m (right bank Rio Uca-
yali)
Micoureus regina, 225 m (near Masisea, left
bank Rio Ucayali)
Mata de Caixa D'Agua; Espirito Santo, Brazil
(2038'S, 4055'W, Geonet)
Didelphis aurita
Matagalpa; Matagalpa, Nicaragua (1255'N,
8555'W, Geonet)
Caluromys derbianus
Didelphis marsupialis (Savala [Lavala])
Marmosa mexicana
Mata Salada, Hato Acapulco; Apure, Venezuela
(ca. 0707'N, 6945'W, Perez-Hernandez,
1989)
Marmosa robinsoni
Mateus Leme; Minas Gerais, Brazil (2000'S,
4428'W, Varejao and Valle, 1982)
Didelphis albiventris
Matfas Romero, N of Rio Sarabia; Oaxaca, Mex-
ico (1653'N, 9502'W, Geonet)
Chironectes minimus, 1 800 m
Philander opossum (S of Matfas Romero)
Matichaca, near Puente Malalma; Junin, Peru
(1114'S, 7532'W, Geonet)
Didelphis albiventris, ca. 6400 ft (= 2000 m)
Matipoo (see Rio Matipo); Minas Gerais, Brazil
Mato do Larva d'Agua, 1.7 km from Santa Tereza
(see Santa Teresa); Espirito Santo, Brazil
Mato Grosso [Matogrosso]; Mato Grosso, Brazil
(1500'S, 5957'W, Paynter and Traylor, 1991)
Caluromys philander (Rio Jurare)
Didelphis marsupialis
Philander opossum
BROWN: ATLAS OF NEW WORLD MARSUPIALS
245
Mato Verde; Minas Gerais, Brazil (1523'S,
4253'W, Varejao and Valle, 1982)
Didelphis albiventris
Matozinhos; Minas Gerais, Brazil (19 C 30'S,
4408'W, Paynter and Traylor, 1991)
Didelphis albiventris, 812 m
Mattas do Piroculina (see Piroculina); Rondonia.
Brazil
Matucana, Rio Rimac, Huarochiri; Lima, Peru
(1151'S, 7624'W, Stephens and Traylor,
1983)
Thylamys pallidior, 8200 ft (= 2560 m)
Matura; St. Andrew, Trinidad and Tobago
(1040'N, 6104'W, Geonet)
Didelphis marsupialis
Maturfn; Monagas, Venezuela (0945'N, 631 l'W.
Paynter, 1982)
Didelphis marsupialis, 74 m (32 km SE of Ma-
turin)
Gracilinanus emiliae
Lutreolina crassicaudata, 1A m
Micoureus demerarae, 74 m
Mavaca, Alto Rio Orinoco; Amazonas, Venezuela
(0231'N, 6510'W, Perez-Hernandez, 1989)
Marmosa murina, 200 m
Philander andersoni, 200 m
Maxon Ranch; Panama, Panama (0857'N,
8000'W, Wenzel and Tipton, 1966)
Marmosa robinsoni
Metachirus nudicaudatus
Maynas; Loreto, Peru (not located)
Philander andersoni
Mazagao, Rio Maraca; Amapa, Brazil (0007'S,
5117'W, Paynter and Traylor, 1991)
Caluromys philander
Philander opossum (right bank)
Mazagao; Para. Brazil (0220'S, 4925'W, Paynter
and Traylor, 1991)
Philander opossum (right bank Rio Tocantins)
Mazan; Azuay, Ecuador (0252'S, 7908'W. Bub-
litz. 1987)
Caenolestes caniventer, 2700 m
Mazatlan; Sinaloa, Mexico (2313'N, 10625'W,
Geonet)
Marmosa canescens
Mecapaca; La Paz, Bolivia (1640'S, 6801'W,
Anderson. 1997)
Thylamys pallidior
Thylamys venustus
Mecatepe, Nandaime; Granada, Nicaragua
(1146'N, 8557'W, Geonet)
Caluromys derbianus
Philander opossum (2 km N, 1 1 km E of Nan-
daime)
Medelh'n; Antioquia, Colombia (0615'N,
7535'W, Paynter, 1997)
Caluromys derbianus, 1538 m
Chironectes minimus
Marmosa murina
Marmosa robinsoni
Medina; Cundinamarca. Colombia (0430'N,
7321'W, Paynter, 1997)
Marmosops impavidus, 576 m
Metachirus nudicaudatus (eastern slope Andes)
Mendes; Rio de Janeiro, Brazil (2232'S,
4344'W, Mustrangi and Patton, 1997)
Marmosops incanus, 400 m
Mendoza (= San Nicolas); Amazonas, Peru
(0620'S. 7724'W. Stephens and Traylor,
1983)
Didelphis marsupialis, 4500 ft (= 1400 m)
Marmosops impavidus, 4500 ft (= 1400 m)
Metachirus nudicaudatus, 4500 ft (= 1400 m)
(Rio Huambe)
Mendoza, 37 km N; Mendoza, Argentina
(3253'S, 6849'W, Paynter, 1985)
Didelphis albiventris
Thylamys pallidior
Menocucho, 16 km ENE; La Libertad, Peru
(080LS, 7850'W, Stephens and Traylor,
1983)
Didelphis marsupialis, 1000 ft (= 300 m)
Mennoniten Kolinien (see Colonia Mennonita);
Boqueron, Paraguay
Mera; Pastaza. Ecuador (0127'S, 7807'W, Payn-
ter. 1993)
Chironectes minimus
Didelphis albiventris, 1100 m
Marmosa murina
Marmosops noctivagus, 1140 m (0128'S,
7808'W)
Metachirus nudicaudatus
Monodelphis adust a
Philander opossum
Mercedes. 6 km S and opposite Buena Hora, Bra-
zil, Rio Guapore; Santa Cruz, Bolivia
(1536'S, 6022'W, Hershkovitz, 1997)
Philander opossum
Merey, opposite Brazo Casiquiare; Amazonas,
Venezuela (0217'N, 6711'W, Hershkovitz,
1997)
Philander andersoni
Merida; Merida, Venezuela (0836'N, 7108'W,
Paynter. 1982)
Didelphis albiventris, 1630 m, 1641 m, 7000 ft
(= 2180 m), 9000 ft (= 2800 m)
Didelphis marsupialis, 7000 ft (= 2180 m)
Gracilinanus dryas, 1630 m
246
FIELDIANA: ZOOLOGY
Gracilinanus marica, 1641 m
Marmosa robinsoni, 1641 m
Marmosops fuscatus, 1600 m, 1630 m, 1641 m
Philander opossum
Merredor; Beni, Bolivia (ca. 1 141 'S, 6505'W,
Hershkovitz, 1997)
Philander opossum
Mesa Bolivar; Merida, Venezuela (0829'N,
7136'W, Paynter, 1982)
Metachirus nudicaudatus
Mesada de las Colmenas, Parque Nacional Cali-
legua; Jujuy, Argentina (2342'S, 6452'W,
Flores et al., 2000)
Micoureus constantiae, 1150 m, 550 m
Thylamys venustus, 1150 m
Meseta de Huanchaca, Parque Nacional Noel
Kempff Mercado; Santa Cruz, Bolivia
(1425'S, 6050'W, Tarifa and Anderson,
1997)
Glironia venusta
Meseta El Pedrero; Santa Cruz, Argentina
(4617'S, 6905'W, Geonet)
Lestodelphys halli
Meta and Vichada (Sabanas of); Colombia (not
precisely located)
Lutreolina crassicaudata
Metates, 5.3 km SSW of, Sierra Juarez; Oaxaca,
Mexico (1546'N, 9618'W, USBGN)
Philander opossum, 1280 m
Metlaltoyuca; Puebla, Mexico (2045'N, 9751'W,
Geonet)
Didelphis marsupialis
Philander opossum
Middle Camp (= Central Camp), Cerro Duida;
Amazonas, Venezuela (Cerro Duida =
0325'N, 6540'W, Paynter, 1982)
Didelphis marsupialis, 100 m
Marmosa murina, 350 ft (= 110 m), 4800 ft
(= 1500 m) (foot of Cerro Duida)
Milagres; Ceara, Brazil (0717'S, 3857'W,
USBGN)
Monodelphis domestica (Sitio Camara)
Milla, Sierra de Merida; Merida, Venezuela
(0830'N, 7122'W, Paynter, 1982)
Marmosa robinsoni, ca. 1110m
Mina Altamira; Atacama, Chile (2627'S,
7019'W, Geonet)
Thylamys elegans
Mina Carbon, Zea; Merida, Venezuela (0823'N,
7147'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Monodelphis palliolata
Philander opossum
Mina Pirquitas, 31 km SE on road 74b, Sierra de
Quichagua; Jujuy, Argentina (2254'S,
6619'W, Flores et al., 2000)
Thylamys pallidior, 4200 m
Minas Capillitas; Catamarca, Argentina (2720'S,
6625'W, Mares et al., 1997)
Thylamys pallidior, 3200 m
Minas de Aroa, 19-30 km NW of San Felipe;
Yaracuy, Venezuela (1025'N, 6854'W, Han-
dley, 1976)
Chironectes minimus, 395 m
Didelphis marsupialis, 395-1400 m
Monodelphis palliolata, 400-430 m
Minas de Oro; Comayagua, Honduras (1425'N,
8738'W, Geonet)
Chironectes minimus
Minca; Magdalena, Colombia (1109'N, 7407'W,
Paynter, 1997)
Caluromys lanatus, 2200 m
Marmosa robinsoni, 2000 ft (= 625 m), 675 m
Mindo; Pichincha, Ecuador (0002'S, 7848'W,
Geonet)
Caluromys derbianus, 4200 ft (= 1270 m)
Chironectes minimus
Didelphis albiventris, 1100 m
Marmosa robinsoni, 1264 m
Marmosops impavidus, 4200 ft (= 1300 m)
Metachirus nudicaudatus, 4200 ft (= 1300 m),
4213 ft (= 1300 m)
Micoureus regina, 5500 ft (= 1700 m)
Philander opossum, 3500 ft (= 1090 m) (below
Mindo)
Minnehaha Creek; Potaro-Suparuni, Guyana
(0508'N, 5907'W, Geonet)
Monodelphis brevicaudata
Mirador, Rio Napo; Tungurahua, Ecuador (ca.
0126'S, 7815'W, Paynter, 1993)
Marmosa murina
Marmosops noctivagus
Mirador (= Montanas de Mirador); Guayas, Ec-
uador (0135'S, 7953'W, Geonet)
Didelphis marsupialis
Mirador; Veracruz-Llave, Mexico (1913'N,
9651'W, Geonet)
Didelphis marsupialis
Marmosa mexicana
Miraflores; Boyaca, Colombia (0512'N, 7312'W,
Paynter, 1997)
Micoureus alstoni
Micoureus demerarae
Miramar; Buenos Aires, Argentina (3816'S,
5751'W, Paynter, 1985)
Didelphis albiventris, sea level (Mar del Sur,
12.5 km SW of Miramar)
Lutreolina crassicaudata, sea level
BROWN: ATLAS OF NEW WORLD MARSUPIALS
247
Monodelphis dimidiata. sea level
Miramar: Tabasco. Mexico (1831'N. 9244'W.
Geonet)
Philander opossum
Miranda; Mato Grosso do Sul. Brazil (20 C 14'S.
5622'W, Paynter and Tray lor. 1991)
Monodelphis domestica
Philander opossum
Mirapinima (see Miripinima): Amazonas. Brazil
Mirimire: Falcon. Venezuela (1110'N. 6843'W.
Geonet)
Caluromys philander ( 14 km ENE of Mirimire)
Didelphis marsupialis. 130-250 m (14 km
ENE of Mirimire)
Marmosa robinsoni, 90-250 m (near Mirimire)
Marmosops cracens, 170 m (5 km and 13 km
E of Mirimire. 1 115'N. 6813'W)
Monodelphis palliolata
Miripinima (= Mirapinima): Amazonas. Brazil
(0211'S. 6108'W. Paynter and Traylor.
1991)
Didelphis marsupialis, ca. 25 m
Miritiba (= Humberto de Campos): Maranhao.
Brazil (0237'S. 4327'W. Paynter and Tray-
lor. 1991)
Caluromys philander
Mision. W of Concepcion (= Misiones): Chaco.
Paraguay (not located)
Thylamys pusilla. 100 m
Mision del Tocuco (= Mision Tukuko): Zulia. Ve-
nezuela (0945'N. 7245'W. Perez-Hernan-
dez. 1989)
Caluromys lanatus. 250 m
Metachirus nudicaudatus
Misiones. W of Concepcion (see Mision): Chaco.
Paraguay
Mision Tukuko (see Mision del Tucuco): Zulia.
Venezuela
Missao Velha: Ceara. Brazil (0715'S. 3908'W.
USBGN)
Monodelphis domestica (Si'tio Araruna. Si'tio
Cinta da Solidade. Si'tio Emboscada. Si'tio
Lapinha)
Mitla. 25 mi S of Oaxaca: Oaxaca. Mexico
(1655'N, 9624'W. Geonet)
Didelphis marsupialis
Miyuyoc: Jujuy. Argentina (2251'S. 6518'W.
Flores et al.. 2000)
Thylamys pallidior
Mocajuba: Para. Brazil (0235'S. 4930'W. Payn-
ter and Traylor. 1991)
Didelphis marsupialis. 30 m (Rio Tocantins)
Philander opossum (E bank Rio Tocantins)
Mocombinho. Manga. Rio Sao Francisco: Minas
Gerais. Brazil (1610'S. 4321'W. Geonet)
Marmosops incanus. 500 m
Moengo: Marowijne. Suriname (0537'N.
5424'W. Geonet)
Marmosa murina
Monodelphis brevicaudata
Philander opossum
Mogi das Cruzes: Sao Paulo. Brazil (2331'S,
4611'W, Paynter and Traylor. 1991)
Monodelphis sore.x. 737 m
Mohinga Valley (= Rio Mojinga). 3 mi S of Fort
Sherman: Colon. Panama (0917'N. 7958'W.
Geonet)
Chironectes minimus
Moira Camp (see Moire): La Paz. Bolivia
Moire (= Moire Camp = Moira Camp); Alto Rio
Madidi: La Paz. Bolivia (1335'S. 6846'W.
Anderson. 1997)
Marmosa murina. 170 m
Marmosops noctivagus
Moire Camp (see Moire): La Paz. Bolivia
Mojinga Swamp; Colon. Panama (0918'N,
7957'W. Geonet)
Didelphis marsupialis
Moju (see Rio Mojui): Para. Brazil
Mojui dos Campos (see Rio Mojui); Para. Brazil
Molleturo: Azuay. Ecuador (02 D 48'S. 7926'W.
Paynter. 1993)
Caenolestes caniventer. 7600 ft (= 2375 m)
Caenolestes fidiginosus. 7600 ft (= 2375 m)
Didelphis albiventris. 7600 ft (= 2375 m)
Montalban (= Hacienda El Merey): Carabobo.
Venezuela <1009'-1017'N, 6821-
68 C 12'W. Handley. 1976)
Caluromys philander. 598 m (near Montalban.
1011'N. 6820'W)
Didelphis marsupialis. 562-1000 m (1013'N.
6820'W)
Marmosa robinsoni. 562-1000 m
Marmosops fuscatus. 1513-1537 m (La Copa.
4 km NW of Montalban. 1013'N. 6820'W)
Monodelphis palliolata. 562-598 m
Montalvo: Pastaza. Ecuador (0204'S. 7658'W.
Paynter. 1993)
Caluromys lanatus. 250 m
Didelphis marsupialis. 250 m
Glironia venusta. 250 m
Metachirus nudicaudatus. 314 m (Rio Bobon-
aza)
Philander andersoni (right bank Rio Bobonaza)
Montana de Edrama. Sierra de Merida: Merida.
Venezuela (Sierra de Merida = 0840'N.
7100'W. Geonet)
248
FIELDIANA: ZOOLOGY
Caluromys lanatus, 2000 m
Montana de Vitoc (see Pueblo Nuevo); Junin,
Peru
Montafias de Balzar (= Balzar Mountains = Cor-
dillera Balzar); Manabf, Ecuador (0055'S,
7955'W, Paynter, 1993)
Caluromys derbianus, 300 m
Marmosa robinsoni, ca. 300 m
Montafias de Cupa (see Cupa); Esmeraldas, Ec-
uador
Montafias de la Sierra (= Montes de la Serra =
Sierra Montafias), near Merida; Merida, Ve-
nezuela (ca. 0840'N, 7100'W, Paynter,
1993)
Caluromys lanatus
Gracilinanus dryas, 3000 m
Montafias de Mirador (see Mirador); Guayas, Ec-
uador
Montafias Uchisera (see Uchisera); Merida, Ve-
nezuela
Montafiita, Florencia, left bank Rio Orteguaza;
Caqueta, Colombia (0130'N, 7528'W, Payn-
ter, 1997)
Metachirus nudicaudatus, 400 m
Monteagudo, 2 km SW; Chuquisaca, Bolivia
(1949'S, 6359'W, Anderson, 1997)
Thylamys venustus
Montealegre; Huanuco, Peru (ca. 0930'S,
7457'W, Stephens and Traylor, 1983)
Marmosops bishopi (provisionally referred to
M. bishopi by Voss et al., 2001)
Monte Alegre (see Monte Alegre do Sul); Sao
Paulo, Brazil
Monte Alegre, Palmeiras; Mato Grosso, Brazil
(1603'S, 5530'W, Hershkovitz, 1997)
Philander opossum
Monte Alegre do Sul (= Ibiti = Monte Alegre);
Sao Paulo, Brazil (2240'S, 4641'W, Paynter
and Traylor, 1991)
Caluromys philander
Monodelphis sorex
Philander frenata (Monte Alegre, Amparo)
Monte Belo; Minas Gerais, Brazil (2120'S,
4620'W, Varejao and Valle, 1982)
Didelphis albiventris
Didelphis aurita
Monte Cano, Pueblo Nuevo; Falcon, Venezuela
(1157'N, 6955'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
Monte Chimborazo; Chimborazo, Ecuador
(0128'S, 7848'W, Geonet)
Caenolestes fuliginosus
Monte Chocolatal (see San Ramon); Beni, Bolivia
Monte Cristo; Tabasco, Mexico (1745'N,
9146'W, Geonet)
Philander opossum
Monte Cristo Mine; Morazan, El Salvador (not
located)
Didelphis marsupialis, 218 m, 700 ft (= 220
m)
Monte Hermoso Formation; Buenos Aires, Argen-
tina (not located)
Lutreolina crassicaudata
Montenegro (see Sao Joao do Monte Negro); Rio
Grande do Sul, Brazil
Montero, 3 km SE; Santa Cruz, Bolivia (1723'S,
6314'W, Anderson, 1997)
Gracilinanus agilis
Philander opossum
Monterrey [Monte Rey], 22 km S of San Jose;
San Jose, Costa Rica (0945'N, 8440'W,
Geonet)
Caluromys derbianus
Philander opossum, 1100 m
Monte Roraima (see Cerro Roraima); Bolivar, Ve-
nezuela
Montes de las Serra (see Montafias de la Sierra);
Merida, Venezuela
Montes de Pedregoso, Sierra de Merida (see Que-
brada La Pedregosa); Merida, Venezuela
Monte Serrat, Itatiaya, Parque Nacional de Itatiaia
(see Pico das Agulhas Negras); Rio de Ja-
neiro, Brazil
Monteverde; Puntarenas, Costa Rica (1018'N,
8449'W, Geonet)
Didelphis marsupialis, 1400-1450 m
Philander opossum, 1380 m (Rio Guacimal, ca.
1018'N, 8450'W)
Monteverde Forest Reserve; Puntarenas, Costa
Rica (Monteverde = 1018'N, 8449'W, Geo-
net)
Didelphis marsupialis, 1500-1800 m
Montevideo; Montevideo, Uruguay (3453'S,
5611'W, Paynter, 1994)
Didelphis albiventris, sea level
Lutreolina crassicaudata, sea level
Monodelphis dimidiata
Montufar, San Gabriel; Carchi, Ecuador (San Ga-
briel = 0036'N, 7749'W, Geonet)
Didelphis albiventris
Monumento al Soldado Pionero; Bolivar, Vene-
zuela (0556'N, 6127'W, Perez-Hernandez,
1989)
Didelphis albiventris, 1500 m
Moon Mountains; Upper Takutu-Upper Essequi-
bo, Guyana (OTOO'N, 5900'W, Times Atlas,
1992, and Hershkovitz files)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
249
Marmosa murina
Moreno (see Mariano Moreno); Buenos Aires,
Argentina
Morgenstond (see "De Morgenstond"); Paramar-
ibo, Suriname
Morichal Largo; Anzoategui, Venezuela (0829'N,
6351'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Morichal Villasmil, Calabozo; Guarico, Venezue-
la (0856'N, 6725'W, Perez-Hernandez,
1989)
Didelphis marsupialis
Marmosa robinsoni
Monodelphis brevicaudata
Moromoro; El Oro, Ecuador (0343'S, 7944'W,
Geonet)
Philander opossum
Morretenho, 30 km NNW of Juquia; Sao Paulo,
Brazil (Juquia = 2419'S, 4748'W, Paynter
and Tray lor, 1991)
Didelphis aurita, 40 m
Morro de Arara [Morro d'Arara], 30 km W of
mouth of Rio Mucuri; Bahia, Brazil (1718'S,
3940'W, Avila Pires, 1965)
Micoureus demerarae
Motatan; Trujillo, Venezuela (0924'N, 7036'W,
Paynter, 1982)
Didelphis marsupialis, 290 m
Metachirus nudicaudatus, 250 m
Philander opossum, 229 m (5 km NNE of Mo-
tatan), 330 m (1 km E of Motatan)
Motupe; Lambayeque, Peru (0609'S, 7944'W,
Stephens and Traylor, 1983)
Didelphis marsupialis, 150 m
Motzorongo; Veracruz-Llave, Mexico (1838'N,
9644'W, Geonet)
Caluromys derbianus
Philander opossum, 1500 ft (= 460 m)
(1834'N, 9644'W)
Mount Sajama (see Sajama Nevado); Oruro, Bo-
livia
Moyobamba; San Martin, Peru (0603'S,
7658'W, Stephens and Traylor, 1983)
Chironectes minimus, 860 m
Didelphis marsupialis, 860 m
Marmosa murina, 860 m
Marmosops impavidus, 855 m, 860 m
Metachirus nudicaudatus, 2820 ft (= 880 m)
Micoureus regina, 860 m
Monodelphis adust a, 860 m
Philander opossum, 2700 ft (= 840 m) (left
bank Rio Mayo)
Mt. Harris, Torure Ward; St. Andrew, Trinidad
and Tobago (1030'N, 6107'W, Geonet)
Caluromys philander, 844 ft (= 260 m)
Didelphis marsupialis, 884 ft (= 270 m)
Mt. Illininiza (see Cerro Illiniza); Pichincha, Ec-
uador
Mt. Pine Ridge; Cayo, Belize (not precisely lo-
cated, Cayo = 1659'N, 8848'W, Geonet)
Didelphis marsupialis
Philander opossum
Mt. Roraima (see Cerro Roraima); Bolivar, Ve-
nezuela
Mt. Roraima; Cuyuni-Mazaruni, Guyana (0512'N,
6044'W, Stephens and Traylor, 1985, Times
Atlas, 1992)
Didelphis marsupialis, 8000 ft (= 2500 m)
Munchique; Cauca, Colombia (0232'N, 7657'W,
Paynter, 1997)
Caenolestes fuliginosus, 2400 m
Didelphis albiventris, 2100-2400 m, 8225 ft ( =
2750 m), 9000 ft (= 2800 m)
Municipio Parapara, Districto Rosco (see Para-
para); Guarico, Venezuela
Murucutu (see Belem); Para, Brazil
Muy Muy; Matagalpa, Nicaragua (1246'N,
8538'W, Geonet)
Caluromys derbianus
Muzo; Boyaca, Colombia (0532'N, 7406'W,
Paynter, 1997)
Caluromys lanatus, 850 m
Marmosops parvidens, 1300 m (0530'N,
7410'W)
Metachirus nudicaudatus, 850 m, 1100 m
N
Nacaome; Valle, Honduras (1331'N, 8730'W,
Geonet)
Philander opossum (Rio de Coray)
Nacional Park of Brazil (see Parque Nacional de
Brasilia); Distrito Frederal, Brazil
Nacunan, Reserva Ecologica de Nacunan; Men-
doza, Argentina (3403'S, 6758'W, Geonet)
Didelphis albiventris
Thylamys pallidior, 4700 ft (= 1465 m)
Nanegal; Pichincha, Ecuador (0007'N, 7846'W,
Paynter. 1993)
Caenolestes fuliginosus, 6000 ft (= 1875 m)
(road to Nanegal)
Chironectes minimus, ca. 1400 m
Didelphis albiventris, 4000 ft (= 1250 m)
Marmosa robinsoni, 1400 m, 5000 ft (= 1560 m)
Micoureus regina, ca. 1400 m
Naranjal, Valle de Suaza; Huila, Colombia
(0217'N, 7559'W, Geonet)
Caluromys lanatus
Didelphis marsupialis
Naranjos (see El Naranjo); San Luis Potosi, Mex-
ico
250
FIELDIANA: ZOOLOGY
Narinja; Mato Grosso, Brazil (not located)
Didelphis marsupialis
Nassau Mountains; Sipaliwini, Suriname
(0448'N, 5436'W, Geonet)
Marmosa lepida
Natagaima, left bank Rio Magdalena; Tolima, Co-
lombia (0351'N, 7506'W, Paynter, 1997)
Caluromys lanatus
National Park of Brazil (see Parque Nacional de
Brasilia); Distrito Federal, Brazil
Nauta, left bank Rio Samiria; Loreto, Peru
(0432'S, 7333'W, Stephens and Tray lor,
1983)
Didelphis marsupialis, 150 m
Philander andersoni, ca. 130 m
Nautla, 9 km NW; Veracruz-Llave, Mexico
(2013'N, 9647'W, Geonet)
Didelphis marsupialis, 10 ft ( 3 m)
Navarro, near Orosi; Cartago, Costa Rica
(0949'N, 8353'W, Geonet)
Marmosa mexicana, 2500-3000 ft (= 780-
930 m)
Nequejahuira; La Paz, Bolivia (ca. 1620'S,
6750'W, Paynter, 1992)
Didelphis albiventris, 8000 ft (= 2500 m)
Micoureus regina, 8000 ft (= 2500 m) (Rio
Unduavi)
Nevati Mission; Pasco, Peru (ca. 1021'S,
7451'W, Stephens and Traylor, 1983)
Didelphis marsupialis, 900 ft (= 280 m)
Neverf; Sucre, Venezuela (1006'N, 6438'W,
Paynter, 1982)
Didelphis marsupialis, 2400 ft (= 750 m)
New River Falls; East Berbice-Corentyne, Guy-
ana (0321'N, 5735'W, USBGN)
Hyladelphys kalinowskii
New Saint Eustatius (= Clarksdale), Demerara
River; Essequibo Islands-West Demerara,
Guyana (0628'N, 5819'W, Geonet)
Marmosa murina
Nicaragua Boquete; Chiriqui, Panama (not locat-
ed)
Didelphis marsupialis
Nickerie River; Sipaliwini, Suriname (0447'N,
5646'W, Geonet)
Philander opossum (upper Nickerie River)
Nicolas Suarez Island (= Isla Nicolas Suarez),
Rio Mamore, near Guayaramerfn; Beni, Bo-
livia (1048'S, 6525'W, Anderson, 1997)
Philander opossum
Nieuw-Amsterdam, right bank Suriname River;
Commewijne, Suriname (0553'N, 5505'W,
Stephens and Traylor, 1985)
Caluromys philander
Didelphis marsupialis
Nievecita Farm; Bocas del Toro, Panama (ca.
0934'N, 8244'W, Geonet)
Didelphis marsupialis
Noetinger; Cordoba, Argentina (3222'S,
6219'W, Paynter, 1985)
Lutreolina crassicaudata
No Leon (= Alto No Leon); Distrito Federal, Ve-
nezuela (1025'N, 6710'W, Paynter, 1982)
Didelphis marsupialis, 1665-2050 m
Marmosops fuscatus, 1750-2000 m
Norco, 20 km W of Vipos; Tucuman, Argentina
(2629'S, 6522'W, Paynter, 1985)
Thylamys venustus, 2500 m
Norte de Yaritagua, 30 km N (see Yaritagua); Yar-
acuy, Venezuela
Northern Range; Simla, Trinidad and Tobago (not
precisely located)
Caluromys philander
Nova Empresa (see Igarape Nova Empressa);
Amazonas, Brazil
Nova Friburgo (= Friburgo = Novo Friburgo);
Rio de Janeiro, Brazil (2216'S, 4232'W,
Paynter and Traylor, 1991)
Chironectes minimus, 846 m
Didelphis aurita, 846 m (2217'S, 4218'W)
Monodelphis americana, 846 m
Philander frenata
Nova Lima; Minas Gerais, Brazil (1959'S,
4351'W, Paynter and Traylor, 1991)
Didelphis albiventris, 745 m
Nova Odessa; Sao Paulo, Brazil (2247'S,
4718'W, USBGN)
Didelphis albiventris
Nova Venecia; Espirito Santo, Brazil (1843'S,
4024'W, Paynter and Traylor, 1991)
Didelphis aurita, 50 m
Nova Vicosa (= Villa Vicosa = Maroba), Rio
Peruipe; Bahia, Brazil (1753'S, 3922'W,
Paynter and Traylor, 1 99 1 )
Didelphis aurita, sea level
Nova Vida, right bank Rio Jurua; Acre, Brazil
(0822'S, 7249'W, Patton et al., 2000)
Caluromys lanatus
Didelphis marsupialis
Marmosops neblina
Marmosops noctivagus
Metachirus nudicaudatus
Micoreus demerarae
Micoureus regina
Monodelphis emiliae
Philander mcilhennyi
Philander opossum
Novita, Rio Tamana; Choco, Colombia (0457'N,
7634'W, Paynter, 1997)
Philander opossum, 400 ft (= 125 m)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
251
Novito; Zulia, Venezuela (1002'N, 7243'W,
Hershkovitz files)
Marmosa robinsoni, 1 132-1 150 m
Metachirus nudicaudatus, 1 135-1 150 m
Novo Friburgo (see Nova Friburgo); Rio de Ja-
neiro, Brazil
Novo Horizonte (see Gorotire); Para, Brazil
Nucleo El Laurel; Miranda, Venezuela (not locat-
ed)
Didelphis marsupialis, 1300 m
Nucleo Pionefro Humboldt (see Aripuana); Mato
Grosso, Brazil
Nueva Germania, 88 km SE of Concepcion (=
Estancia La Germania); San Pedro, Paraguay
(2354'S, 5634'W, Paynter, 1989)
Didelphis albiventris, ca. 150 m
Didelphis aurita, ca. 150 m
Lutreolina crassicaudata
Micoureus demerarae, ca. 150 m
Nueva Ocotepeque; Ocotepeque. Honduras
(1426'N, 8911'W, Geonet)
Didelphis marsupialis
Nuevo San Juan; Colon/Panama, Panama (not lo-
cated)
Chironectes minimus
Nuevo San Juan, Rio Galvez; Loreto, Peru (Rio
Galvez = 0512'S, 7253'W, Geonet)
Hyladelphys kalinowskii
Nula, 3 km N of Nulita; Apure, Venezuela (Nulita
= 0718'N, 7154'W, Geonet)
Caluromxs lanatus
Nulita; Apure, Venezuela (0718'N, 7154'W,
Geonet)
Caluromxs lanatus, 24 m (Selvas de San Cam-
ilo, 29 km SSW of Santo Domingo)
Didelphis marsupialis, 24 m
Philander opossum, 24 m (0719'N, 7155'W)
Nun; Bocas del Toro, Panama (0855'N, 8149'W.
Geonet)
Metachirus nudicaudatus
Philander opossum
O
Oaxaca; Oaxaca, Mexico (1702'N, 9644'W,
Geonet)
Marmosa canescens, 4600 ft (= 1430 m)
Ocabamba (see Ocobamba); Cusco, Peru
Ocampo; Tamaulipas. Mexico (2250'N, 9920'W.
Geonet)
Philander opossum, 2400 ft (= 750 m)
Ocaneque (see Oconeque); Puno. Peru
Occobamba Valley (see Rio Ocobamba); Cusco,
Peru
Ocidente, opposite left bank Rio Jurua; Acre. Bra-
zil (0834'S, 7248'W, Patton et al., 2000)
Marmosops neblina
Metachirus nudicaudatus
Philander opossum
Ocoa River, near, on the Villavicencio-Acacias
Road (see Rio Ocoa); Meta, Colombia
Ocobamba (= Ocabamba); Cusco. Peru (1252'S,
7222'W. Geonet)
Didelphis albiventris, 3900 m (below La Raya
Pass)
Marmosa murina, ca. 9000 ft (= 2800 m)
Ocobamba Pass; Cusco. Peru (1328'S, 7333'W,
Geonet)
Didelphis albiventris, 11000-13000 ft ( =
3400-4000 m)
Ocobamba Valley (see Rio Ocobamba): Cusco,
Peru
Oconeque (= Ocaneque); Puno, Peru (1403'S,
6942'W. Stephens and Traylor, 1983)
Didelphis albiventris, 1956 m, 7000 ft ( =
2180 m)
Ocotal: Nueva Segovia, Nicaragua (1338'N,
8629'W. Geonet)
Marmosa me.xicana, 4500 ft (= 1400 m) (near
Honduras border)
Ocumare (see Ocumare de la Costa); Aragua, Ve-
nezuela
Ocumare de la Costa (= Ocumare); Aragua, Ve-
nezuela (1028'N, 6746'W, Paynter, 1982)
Chironectes minimus
Didelphis marsupialis, ca. 25 m
Marmosa robinsoni, 180 m
Oelemarie Airstrip: Sipaliwini. Suriname
(0306'N. 5432'W, Geonet)
Didelphis marsupialis
Marmosa murina
Monodelphis brevicaudata
Ojo Negro (road from Nanegal to Anca); Pichin-
cha. Ecuador (Nanegal = 0007'S. 7846'W,
Paynter. 1993)
Micoureus regina
Oko Mts. (= Essequibo): Cuyuni-Mazaruni, Guy-
ana (0628'N. 5857'W, Times Atlas, 1992,
and Stephens and Traylor. 1985)
Caluromxs philander
Didelphis marsupialis, 1000 ft (= 300 m)
Old Point, Isla Bastimentos: Bocas del Toro. Pan-
ama (0917'N, 8205'W. Geonet)
Caluromxs derbianus
Philander opossum
Old Santo Domingo Trail; Pichincha. Ecuador
(not located)
Caenolestes convelatus, 6500 ft (= 2030 m)
Caenolestes fuliginosus, 8750 ft (= 2730 m)
Ollantaytambo; Cusco, Peru (1316'S. 7216'W,
Stephens and Traylor, 1991)
252
FIELDIANA: ZOOLOGY
Didelphis albiventris, 2846 m
Lestoros inca, 3700 m (25-26 km NW of Ol-
lantaytambo), 3530 m (32 km NW of Ollan-
taytambo)
Olmue; Valparaiso, Chile (3300'S, 7112'W,
Geonet)
Thylamys elegans, 152 m
Ometepe (= Islas de Ometepe); Rivas, Nicaragua
(1130'N, 8535'W, Geonet)
Philander opossum
Onkone Yare (see Parque Nacional Yasunf);
Napo, Ecuador
Onverwacht, 25 km S of Paramaribo; Para, Suri-
name (0535'N, 5511'W, Geonet)
Micoureus demerarae
Oran (see San Ramon de la Nueva Oran); Salta,
Argentina
Oriente (see Seringal Oriente); Acre, Brazil
Oriximina; Para, Brazil (0145'S, 5552'W, Payn-
ter and Tray lor, 1991)
PhUander opossum
Orizaba; Veracruz-Llave, Mexico (1851'N,
9706'W, Geonet)
Didelphis marsupialis, 5500 ft (= 1700 m)
Philander opossum, 5500 ft (= 170 m)
Oropuche (see El Cerro del Oropuche); Trinidad
and Tobago
Orosa; Loreto, Peru (0326'S, 7208'W, Hershko-
vitz, 1997)
Marmosa murina, 100 m (right bank opposite
mouth Rio Napo)
Marmosops noctivagus, ca. 100 m
Micoureus regina, ca. 100 m (mouth Rio Napo)
Philander andersoni
Oruro; Oruro, Bolivia (1759'S, 6709'W, Ander-
son, 1997)
Thylamys venustus
Oruro, 64 km S (= 40 mi S); Oruro, Bolivia
(1834'S, 6709'W, Anderson, 1997)
Thylamys venustus, 12000 ft (= 3750 m)
Osorno; Los Lagos, Chile (4034'S, 7309'W,
Geonet)
Dromiciops gliroides, 145 m, 175 m, (32 km
SSE, 54 km SSE of Osorno)
Rhyncholestes raphanurus, 100 m, 140 m (32
km SSE of Osorno, 4102'S, 7230'W)
Otro Cerro, 1 8 km W of Chumbicha; Catamarca,
Argentina (Chumbicha = 2852'S, 6614'W,
Geonet)
Thylamys pallidior, 3000 ft (= 930 m)
Ouanary Riviere; French Guiana (0414'N,
5139'W, Stephens and Traylor, 1985)
Didelphis marsupialis, sea level
Philander opossum
Ouro Preto; Minas Gerais, Brazil (2023'S,
4330'W, Paynter and Traylor, 1991)
Didelphis aurita, 1061 m
Oxapampa (= San Juan de Oxapampa); Pasco,
Peru (1034'S, 7524'W, Geonet)
Caluromys lanatus
Metachirus nudicaudatus, 900 ft (= 280 m)
Philander andersoni, 900 ft (= 280 m)
Oyapock Fluve (see Oyapock River); French Gui-
ana
Oyapock River (= Oyapock Fluve = Yapock Riv-
er); French Guiana (0408'N, 5140'W, Ste-
phens and Traylor, 1985)
Chironectes minimus
Oyeros, 3 km N; Jujuy, Argentina (2430'S,
6500'W, Flores et al., 2000)
Thylamys venustus
Pacasmayo; La Libertad, Peru (0724'S, 7934' W,
Stephens and Traylor, 1983)
Didelphis marsupialis, 8 m
Pachahal [Pachatal], below Mindo; Pichincha, Ec-
uador (Mindo = 0002'S, 7848'W, Paynter,
1993)
Metachirus nudicaudatus, 3000 ft (= 930 m)
Pacific coast; Chiapas, Mexico (not precisely lo-
cated)
Chironectes minimus
Pacuare, Rio Pacuare; Cartago, Costa Rica
(0944'N, 8333'W, Geonet)
Caluromys derbianus
Philander opossum (ca. 1014'N, 8317'W)
Padcoya, 1 1.5 km N and 5.5 km E; Tarija, Bolivia
(2147'S, 6440'W, Anderson, 1997)
Thylamys venustus
Padilla, 9 km N; Chiquisaca, Bolivia (1912'S,
6927'W, Anderson, 1997)
Thylamys pusilla
Pad van Wanica, just S of Paramaribo; Paramar-
ibo, Suriname (Paramaribo = 0550'N,
5510'W, Geonet)
Caluromys philander
Paeto, below Gualea; Pichincha, Ecuador (Gualea
= 0007'N, 7850'W, Paynter, 1993)
Caluromys derbianus, 3000 ft (= 930 m)
Pagancillo; La Rioja, Argentina (2934'S,
6803'W, Geonet)
Thylamys pallidior
Pagash; La Libertad, Peru (not located)
Didelphis albiventris, 1400 m
Paiguano; Coquimbo, Chile (3001'S, 7032'W,
Geonet)
Thylamys elegans, 3300 ft (= 1030 m)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
253
Pailon; Santa Cruz. Bolivia (1733'S. 6244'W.
Anderson, 1997)
Monodelphis domestiea (also at 7 km N of Pai-
lon = 1734'S, 6234'W)
Paime: Cundinamarca. Colombia (0522'N.
7410'W. Paynter. 1997)
Marrnosa robinsoni, 1038 m
Pajaral; Santa Cruz, Bolivia (1458'S. 6232'W.
Anderson, 1997)
Micoureus constantiae
Philander opossum
Pakani: Upper Takutu-Upper Essequibo. Guyana
(0415'N. 5828'W. Geonet)
Didelphis marsupialis
Pakitza; Madre de Dios. Peru (ca. 1150'S.
7117'W. Pacheco et al.. 1993)
Gracilinonus agilis. 350 m (N bank Rio Manu)
Marrnosa rubra. 350 m
Mannosops noctivagus. 350 m (N bank Rio
Manu)
Mannosops panidens. 350 m (N bank Rio
Manu)
Metachirus nudicaudatus. 350-500 m (N bank
Rio Manu)
Micoureus regina, 350 m
Monodelphis brevicaudata. 350 m
Philander opossum, 350 m (N bank Rio Manu)
Palacios; Beni, Bolivia (1334'S. 6519'W. Hersh-
kovitz. 1997)
Philander opossum
Palambla: Piura. Peru (0523'S. 7937'W. Ste-
phens and Traylor. 1983)
Didelphis marsupialis. 1190-1985 m
Palenque: Chiapas. Mexico (1731'N. 9158'W.
Geonet)
Caluromys derbianus
Didelphis marsupialis (16 km NW of Palenque)
Philander opossum
Palenque. Ruinas de: Chiapas. Mexico (Palenque
= 1731'N. 9158'W. Geonet)
Didelphis marsupialis. 90 m. 300 m
Pallatanga: Chimborazo. Ecuador (01 C 59'S.
7857'W. Paynter. 1993)
Didelphis marsupialis. 1500 m
Mannosops impavidus, 1500 m
Palma: Goias. Brazil (ca. 1233'S. 47 C 46'W. Payn-
ter and Traylor, 1991)
Micoureus demerarae, ca. 250 m
Palmar (= Palmar Sur): Puntarenas. Costa Rica
(08 C 58'N. 8327'W. Geonet)
Didelphis marsupialis
Palmar Sur (see Palmar); Puntarenas. Costa Rica
Palmarito. Rio San Julian; Santa Cruz, Bolivia
(1649'S. 6237'W. Anderson, 1997)
Mannosops dorothea
Philander opossum
254
Palma Sola; Jujuy. Argentina (240'S. 6449'W.
Flores et al.. 2000)
Thxlamxs venustus
Palmeira:' Parana. Brazil (25 C 25'S, 5000'W.
Hershkovitz. 1992)
Chironectes minimus
Didelphis albiventris
Gracilinanus microtarsus. 865 m
Monodelphis dimidiata
Palmeira dos Indios: Alagoas. Brazil (0925'S.
3637'W. Paynter and Traylor. 1991)
Didelphis albiventris. 342 m
Monodelphis domestiea (Sitio Capuma. Sftio
Dormiao. Sitio Riveira. Sitio Sabia)
Palmeiras: Bahia. Brazil (1227'S. 4134'W. Cer-
queira. 1984)
Didelphis albiventris
Palmera. Rio Pastaza: Napo. Ecuador (not locat-
ed)
Mannosops impavidus. 4000 ft (= 1250 m)
Palmera. E of Banos (see Banos Oriente); Tun-
guarahua. Ecuador
Palmilla (see La Palmilla); Valparaiso. Chile
Palmira: Valle del Cauca. Colombia (0332'N.
7616'W. Paynter. 1997)
Chironectes minimus
Palmira de Zarcero: Alajuela. Costa Rica
(101 l'N. 8424'W. Geonet)
Didelphis marsupialis
Palmira: Pando. Bolivia (H42'S. 6756'W. An-
derson. 1997)
Mannosops noctivagus
Palm Swamp. 12 km ESE of Resistencia; Cha-
co. Argentina (2727'S. 5859'W. Paynter.
1985)
Lutreolina crassicaudata. 170 m
Palo de Vaca. Rancho Grande (see Estacion Biol-
ogica de Rancho Grande); Aragua, Venezuela
Paloemeu Airstrip. Tapanahoni River: Sipaliwini.
Suriname (0321'N, 5527'W. Geonet)
Monodelphis brevicaudata, 210 m
Palomino. Sierra Nevada de Santa Marta; Mag-
dalena. Colombia (1102'N. 7339'W. Payn-
ter. 1997)
Caluromys lanatus, ca. 600 m
Mannosa robinsoni. ca. 600 m
Metachirus nudicaudatus (on Caribbean Sea. N
of Sierra Nevada de Santa Marta)
Palo Santo: Chiriquf. Panama (0849'N. 8241'W.
Geonet)
Didelphis marsupialis
Palos Blancos: La Paz. Bolivia (1534'S.
6716'W. Anderson. 1997)
Didelphis marsupialis
Micoureis constantiae
Thxlamxs venustus
FIELDIANA: ZOOLOGY
Palos Quemados; O'Higgins, Chile (3402'S,
7034'W, Geonet)
Thylamys elegans
Pambilar, 10 km N of Concepcion; Esmeraldas,
Ecuador (Concepcion = 0103'N, 7850'W,
USBGN)
Didelphis marsupialis, 20 m
Pamilla (see La Palmilla); Valparaiso, Chile
Pampa de Agnia, 30 km N; Chubut, Argentina
(4328'S, 6949'W, Mares and Braun, 2000)
Lestodelphys halli
Pampa de Guanacos, 6 km S, 2 km E; Santiago
del Estero, Argentina (2617'S, 6201'W, Flo-
res et al., 2000)
Thylamys pusilla, ca. 1 20 m
Pampa de Meio; Beni, Bolivia (1230'S, 6415'W,
Anderson, 1997)
Gracilinanus agilis
Pampa Linda, 5 km N, 44 km W, Bariloche; Rio
Negro, Argentina (Bariloche = 4109'S,
7118'W, Paynter, 1985)
Dromiciops gliroides, 950 m
Pamplona; Norte de Santander, Colombia
(0723'N, 7239'W, Paynter, 1997)
Didelphis albiventris, 7300 ft (= 2280 m)
Panama City; Panama, Panama (0858'N,
7932'W, Geonet)
Caluromys derbianus
Panama Viejo; Panama, Panama (0900'N,
7929'W, Geonet)
Caluromys derbianus
Pandora; Limon, Costa Rica (0944'N, 8258'W,
Geonet)
Didelphis marsupialis
Panguana; Huanuco, Peru (ca. 0937'S, 7457'W,
Stephens and Traylor, 1983)
Philander mcilhennyi
Pantanal; Mato Grosso do Sul, Brazil (1800'S,
5600'W, Geonet)
Caluromys lanatus, ca. 50 m
Pantoja, Carretera Maracay-Ocumare; Aragua,
Venezuela (1021'N, 6741'W, Perez Hernan-
dez, 1989)
Gracilinanus marica
Micoureus demerarae
Marmosops fuscatus
Papallacta; Napo, Ecuador (0022'S, 7808'W,
Paynter, 1983)
Caenolestes fuliginosus
Didelphis albiventris, 9980 ft (= 3120 m)
Papantla; Veracruz-Llave, Mexico (2027'N,
9719'W, Geonet)
Didelphis marsupialis, 300 ft (= 90 m)
Philander opossum
Papudo; Valparaiso, Chile (3231'S, 7027'W,
Geonet)
Thylamys elegans, sea level
Para (see Belem); Para, Brazil
Para de Minas; Minas Gerais, Brazil (1951'S,
4437'W, Paynter and Traylor, 1991)
Didelphis albiventris, 796 m
Paracou; French Guiana (0523'N, 5254'W, Geo-
net)
Didelphis marsupialis
Chironectes minimus
Gracilinanus emiliae
Hyladelphys kalinowskii
Marmosa murina
Marmosops parvidens
Marmosops pinheiroi
Metachirus nudicaudatus
Micoureus demerarae
Monodelphis brevicaudata
Philander opossum
Parahyba; Brazil (see Parnaiba); Piauf, Brazil
Paraiba do Sul; Rio de Janeiro, Brazil (2206'S,
4317'W, Paynter and Traylor, 1991)
Didelphis aurita, 280 m
Paramaribo (= Rijweg); Paramaribo, Suriname
(0550'N, 5510'W, Geonet)
Caluromys philander
Didelphis marsupialis
Marmosa lepida
Marmosa murina
Monodelphis brevicaudata (35 km S of Para-
maribo)
Philander opossum (ca. 9 km W of Paramaribo)
Paramba (see Hacienda Paramba); Imbabura, Ec-
uador
Paramillo; Antioquia, Colombia (0659'N,
7551'W, Paynter, 1997)
Didelphis albiventris, 12500 ft (= 3900 m)
Paramito; Merida, Venezuela (0859'N, 7046'W,
Handley, 1976)
Didelphis albiventris, 2016 m, 3020-3275 m
Paramo de Barragan; Valle del Cauca, Colombia
(0400'N, 7540'W, Patterson and Albuja,
1996, and Geonet)
Caenolestes fuliginosus
Paramo de Guanacas, km 5 1 , between Totoro and
Inza (= km 51); Cauca, Colombia (0230'N,
7615'W, Patterson and Albuja, 1996, and
Paynter and Traylor, 1 98 1 )
Caenolestes fuliginosus, 3100 m
Paramo de La Rusia (see Cordillera de Rusia);
Bocaya/Santander, Colombia
Paramo de Mucubaji (see Laguna Negra); Merida,
Venezuela
Paramo de Purace, near Popayan; Cauca, Colom-
bia (0224'N, 7623'W, Paynter, 1997)
Caenolestes fuliginosus, 3500 m
Paramo de Tama; Tachira, Venezuela (0725'N,
7226'W, Paynter, 1982)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
255
Caenolestes fuliginosus, 2450-3329 m
Didelphis albiventris, 6000-7000 ft (= 1875-
2180 m)
Paramo Frontino; Antioquia, Colombia (0628'N,
7604'W, Paynter, 1997)
Caenolestes convelatus, 3100-3600 m
Caenolestes fuliginosus, 3100 m, 3300 m
Paramos El Soche; Cundinamarca, Colombia
(0432'N, 7420'W, USBGN)
Caenolestes fuliginosus
Paramos of Chimborazo or Pichincha; Pichincha,
Ecuador (not located)
Caenolestes fuliginosus
Paramo Sonson; Antioquia, Colombia (0543'N,
7515'W, Paynter, 1997)
Caenolestes fuliginosus, 8800 ft (= 2750 m),
3050 m, 3100 m (7 km E of Paramo Sonson)
Paramo Tambor (see Hacienda La Carbonera);
Merida, Venezuela
Paramo Zumbader; Tachira, Venezuela (0758'N,
7204'W, Geonet)
Didelphis albiventris, 2250 m
Parana do Pocao (see Caracaraf); Roraima, Brazil
Paraopeba; Minas Gerais, Brazil (1915'S,
4425'W, Varejao and Valle, 1982)
Didelphis albiventris, 509 m
Parapara (= Municipio Parapara); Guarico, Ve-
nezuela (0944'N, 6718'W, Paynter, 1982)
Caluromys philander
Para River, ca. 10 km SE of Paramaribo; Para-
maribo, Suriname (0545'N, 5508'W, Geo-
net)
Philander opossum
Paratani; Cochabamba, Bolivia (1734'S,
6621'W, Paynter, 1992)
Thylamys venustus, 2450 m, 2800 m
Paratubal [Paratudal]; Mato Grosso, Brazil (not
precisely located)
Marmosa murina
Pareditas, 35 km S, 3 km E on Highway 40; Men-
doza, Argentina (3356'S, 6904'W, Geonet)
Thylamys pallidior, 4700 ft (= 1468 m)
Pariacoto, Rio Pira Valley; Ancash, Peru
(0931'S, 7753'W, Stephens and Traylor,
1983)
Thylamys pallidior, 8500 ft (= 2650 m)
Parintins (= Villa Bella Imperatriz = Bella Im-
peratriz = Imperatriz); Amazonas, Brazil
(0236'S, 5644'W, Paynter and Traylor,
1991)
Didelphis marsupialis, 24 m (boca Rio Andira)
Marmosa murina, sea level
Micoureus demerarae, 24 m
Philander opossum
Parita; Herrera, Panama (0800'N, 8031'W, Geo-
net)
Caluromys derbianus
Parla Chonto; Amazonas, Venezuela (not located)
Chironectes minimus
Parnahyba (see Parnaiba); Piaui, Brazil
Parniba (see Parnaiba); Piaui, Brazil
Parnaiba (= Parahyba = Parnahyba = Parniba);
Piaui, Brazil (0254'S, 4146'W, Geonet)
Marmosa murina
Parque Biologico Sierra de San Javier; Tucuman,
Argentina (2647'S, 6522'W, Capllonch et
al., 1997)
Didelphis albiventris
Lutreolina crassicaudata
Thylamys venustus
Parque de La Macarena (= Parque La Macarena),
Rio Guapaya; Meta, Colombia (0245'N,
7355'W, Paynter, 1997)
Caluromys lanatus
Parque de Purace; Cauca, Colombia (0224'N,
7623'W, Patterson and Albuja, 1996)
Caenolestes fuliginosus, 3500 m
Parque Estadual de Ilhabela, Ilha de Sao Sebas-
tiao; Sao Paulo, Brazil (Ilha de Sao Sebastiao
= 2340'S, 4518'W, Paynter and Traylor,
1991)
Philander frenata, 650 m
Parque Estadual de Florestal do Rio Doce (see
Parque Estadual do Rio Doce); Minas Gerais,
Brazil
Parque Estadual do Ibitipoca, 30 km N of Lima
Duarte; Minas Gerais, Brazil (ca. 2133'S,
4355'W, Paynter and Traylor, 1991)
Philander frenata
Parque Estadual do Rio Doce (= Parque Estadual
de Florestal do Rio Doce = Parque Florestal
Estadual do Rio Doce = Rio Doce State For-
estry Park = Rio Doce State Park); Minas
Gerais, Brazil (1955'-1930'S, 4356'-
4231'W, Paynter and Traylor, 1991)
Caluromys philander, 250-515 m
Didelphis aurita (1940'S, 4233'W, Varejao
and Valle, 1982)
Gracilinanus agilis
Gracilinanus microtarsus, 230-515 m
Marmosops incanus, 200-500 m (1930'S,
4230'W)
Metachirus nudicaudatus
Micoureus demerarae
Parque Estadual da Pedra Blanca; Rio de Janeiro,
Brazil (2256'S, 4317'W, Mustrangi and Pat-
ton, 1997)
Marmosops incanus, sea level
256
FIELDIANA: ZOOLOGY
Parque Estadual do Rfo Preto, 15 km S of Sao
Goncalo do Rio Preto (= Sao Goncalo do
Supacaf); Minas Gerais, Brazil (2154'S,
4536'W, Paynter and Traylor, 1991)
Micoureus demerarae
Parque Florestal Estadual da Jaiba; Minas Gerais,
Brazil (Jaiba = 1212'S, 3853'W, Geonet)
Marmosops incanus
Parque Florestal Estadual do Rio Doce (see Flo-
restal Estadual do Rio Doce); Minas Gerais,
Brazil
Parque La Macarena (see Parque de La Macar-
ena); Meta, Colombia
Parque Lecoq; Montevideo, Uruguay (ca.
3453'S, 5611'W, Paynter, 1994)
Lutreolina crassicaudata
Parque Nacional Baritu, desembocadura Arroyo
Santelmita; Salta, Argentina (2231'S,
6437'W, Flores et al., 2000)
Didelphis albiventris
Micoureus constantiae, 700 m
Thylamys venustus
Parque Nacional Braulio Carrillo, near San Mi-
guel; Heredia, Costa Rica (ca. 1028'N,
8405'W, Timm et al., 1987)
Didelphis marsupialis, 200-750 m
Micoureus alstoni
Parque Nacional Calilegua, Caimancito (= Aguas
Negras); Jujuy, Argentina (2345'S, 6456' W,
Flores et al., 2000)
Didelphis albiventris, 1700 m, 2500 m, 2800 m
Lutreolina crassicaudata
Micoureus constantiae, 550 m
Thylamys venustus, 600 m
Parque Nacional Chaco; Chaco, Argentina
(2640'S, 5948'W, Heinonen and Chebez,
1997)
Didelphis albiventris
Gracilinanus agilis
Lutreolina crassicaudata
Philander opossum
Parque Nacional da Amazonia (Tapajos); Para,
Brazil (0342'-0450'S, 57 21'-5622'W,
Paynter and Traylor, 1991)
Didelphis marsupialis
Parque Nacional da Serra do Cfpo, Serra do Es-
pinha^o; Minas Gerais, Brazil (1914'S,
4333'W, Mustrangi and Patton, 1997)
Marmosops incanus, 900-1200 m
Parque Nacional de Brasilia (= National Park of
Brazil); Distrito Federal, Brazil (1535'S,
4854'W, Hershkovitz, 1997)
Chironectes minimus
Didelphis albiventris
Gracilinanus agilis, 700-1300 m
Philander frenata
Parque Nacional del Manu (= Manu National
Park = Parque Nacional Manu); Madre de
Dios-Cusco, Peru (1200'S, 7130'W, Ste-
phens and Traylor, 1983)
Chironectes minimus
Glironia venustus
Parque Nacional de Puyehue; Los Lagos, Chile
(4039'-4042'S, 7230'-7218'W, Patterson
and Albuja, 1996, Geonet)
Dromiciops gliroides
Rhyncholestes raphanurus, 700 m (4042'S,
7218'W)
Parque Nacional de Rancho Grande (see Estacion
Biologica de Rancho Grande); Aragua, Ve-
nezuela
Parque Nacional de Sierra Nevada; Merida, Ve-
nezuela (0833'N, 7045'W, Geonet)
Gracilinanus dryas, 2200 m
Parque Nacional Diamante (= La Azotea); Entre
Rios, Argentina (3207'S, 6038'W, Heinonen
and Chebez, 1997)
Didelphis albiventris
Parque Nacional do Caparao; Minas Gerais, Bra-
zil (2030'S, 4140'W, Mustrangi and Patton,
1997)
Didelphis aurita
Marmosops incanus, 1000 m
Marmosops paulensis, 1000 m
Philander frenata (Pedra Roxa = 2149'S,
4409'W)
Parque Nacional do Itatiaia (see Pico das Agulhas
Negras); Rio de Janeiro, Brazil
Parque Nacional "El Avila"; Distrito Federal, Ve-
nezuela (1035'N, 6635'W, Geonet)
Didelphis marsupialis, 1000 m
Marmosa robinsoni
Marmosops fuscatus, 1000 m
Monodelphis palliolata, 1000 m
Parque Nacional El Palmar; Entre Rfos, Argentina
(3150'S, 5817'W, Heinonen and Chebez,
1997)
Didelphis albiventris
Lutreolina crassicaudata, ca. 50 m
Thylamys pusilla, ca. 50 m
Parque Nacional El Rey; Salta, Argentina
(2442'S, 6438'W, Paynter, 1985)
Didelphis albiventris
Lutreolina crassicaudata
Thylamys venustus
Parque Nacional Guatopo (= Guatopo National
Park); Miranda-Guarico, Venezuela (ca.
BROWN: ATLAS OF NEW WORLD MARSUPIALS
257
10H'-0955'N, 66 31'-66 19'W, Handley,
1976)
Caluromys philander, 250-1500 m (ca.
1005'N, 6630'W)
Chironectes minimus, 250-1500 m
Didelphis marsupialis, 250-1500 m
Marmosa murina, 250-1500 m (ca. 1005'N,
6630'W)
Marmosops fuscatus, 250-1500 m (ca.
1005'N, 6630'W)
Micoureus demerarae, 250-1500 m (ca.
1005'N, 6630'W)
Monodelphis palliolata, 250-1500 m
Parque Nacional Iguazu [Parque Nacional Igua-
q\i]; Misiones, Argentina-Parana, Brazil
(2515'-25 41'S, 5436'-5426'W, Paynter
and Traylor, 1991)
Chironectes minimus
Didelphis albiventris, 5 m
Didelphis aurita
Lutreolina crassicaudata
Metachirus nudicaudatus
Micoureus demerarae
Monodelphis iheringi
Monodelphis scalops
Monodelphis sorex
Philander opossum
Parque Nacional Lago Puelo; Chubut, Argentina
(4211'S, 7142'W, Heinonen and Chebez,
1997)
Dromiciops gliroides
Parque Nacional Laguna Blanca; Neuquen, Ar-
gentina (3903'S, 7023'W, Paynter, 1985)
Thy I amy s pallidior
Parque Nacional Lanfn; Neuquen, Argentina
(3955'S, 7125'W, Paynter, 1985)
Dromiciops gliroides
Parque Nacional Lihue Calel; La Pampa, Argen-
tina (3757'S, 6533'W, Heinonen and Che-
bez, 1977)
Lestodelphys halli
Thylamys pallidior
Parque Nacional Los Alcerces; Chubut, Argentina
(4255'S, 7152'W, Heinonen and Chebez,
1997)
Dromiciops gliroides
Parque Nacional Los Arrayanes; Neuquen, Argen-
tina (4050'S, 7138'W, Heinonen and Che-
bez, 1997)
Dromiciops gliroides
Parque Nacional Machalilla; Guayas-Manabi, Ec-
uador (ca. 0142'S, 8046'W, Paynter, 1993)
Caluromys derhianus, 0-800 m
Didelphis marsupialis, 0-800 m
Parque Nacional Manu (see Parque Nacional del
Manu); Madre de Dios-Cusco, Peru
Parque Nacional Mburucuya; Corrientes, Argen-
tina (2801'S, 5801'W, Heinonen and Che-
bez, 1997)
Didelphis albiventris
Lutreolina crassicaudata
Parque Nacional Nahuelbuta; Araucania-Biobio,
Chile (3751'S, 7305'W, Jimenez and Ra-
geot, 1979)
Dromiciops gliroides, 950 m
Parque Nacional Nahuel Huapi (also see Beatriz);
Rio Negro/Neuquen, Argentina (4103'S,
7135'W, Heinonoen Fortabut and Chebez,
1997)
Dromiciops gliroides, 800 m (Villa Angustura)
Rhyncholestes raphanurus
Parque Nacional Rio Pilcomayo; Formosa, Argen-
tina (2502'S, 5812'W, Heinonen and Che-
bez, 1997)
Didelphis albiventris
Lutreolina crassicaudata
Philander opossum
Thylamys pusilla
Parque Nacional Sierra de las Quijadas; San Luis,
Argentina (ca. 3229'S, 6702'W, Heinnonen
and Chebez, 1997)
Thylamys pallidior
Parque Nacional Tikal (= Tikal); Peten, Guate-
mala (1715'N, 8939'W, Geonet)
Didelphis marsupialis
Philander opossum
Parque Nacional Tolhuaca; Los Lagos, Chile
(3910'S, 7143'W, Jimenez and Rageot,
1979)
Dromiciops gliroides
Parque Nacional Vicente Perez Rosales, "La Pi-
cada Forest" (see "La Picada Forest"); Los
Lagos, Chile
Parque Nacional Villarrica; Araucania, Chile
(3927'S, 7143'W, Geonet)
Dromiciops gliroides
Parque Nacional Yapacana, Cano; Amazonas, Ve-
nezuela (0341'N, 6652'W, Perez-Hernan-
dez, 1989)
Caluromys lanatus
Parque Nacional Yasunf, 20 km S of Pompeya Sur
(= Onkone Yare); Napo, Ecuador (Pompeya
= 0026'S, 7637'W, Geonet)
Didelphis marsupialis
Parque Nacional Yasunf, Onkone Sur, 38 km S of
Pompeya Sur; Napo, Ecuador (Pompeya =
0026'S, 7637'W, Geonet)
Philander andersoni
258
FIELD1ANA: ZOOLOGY
Parque Nacional Ybycui; Paraguarf, Paraguay
(2601'S, 5703'W, Paynter, 1989)
Chironectes minimus
Didelphis albiventris
Lutreolina crassicaudata
Metachirus nudicaudatus
Parque Pedro del Rio, near Biobio; Concepcion,
Chile (Concepcion = 3650'S, 7303'W, Geo-
net)
Dromiciops gliroides
Parque Provincial El Cochuna; Tucuman, Argen-
tina (2718'S, 6554'W, Flores et al., 2000)
Thylamys venustus
Pasa Nueva; Veracruz-Llave, Mexico (1759'N,
9511'W, Gardner, 1973)
Marmosa mexicana
Paso Bajito, Rio Moquite; Anzoategui, Venezuela
(0836'N, 6413'W, Geonet)
Lutreolina crassicaudata, 160-180 m
Paso del Rey, along Arroyo de la Canada Honda;
San Luis, Argentina (3257'S, 6600'W, Geo-
net)
Thylamys pallidior, 4700 ft (= 1465 m), 4400
ft (= 1375 m)
Paso Duraznero, Ayo da Mina (= Arroyo de la
Mina); Rio Grande do Sul, Brazil (Arroyo de
la Mina = 3153'S, 5405'W, USBGN)
Didelphis albiventris
Paso Pino Hachado; Araucania, Chile (3840'S,
7054'W, Greer, 1965)
Dromiciops gliroides, 1460 m
Passo Fundo; Rio Grande do Sul, Brazil (2815'S,
5220'W, Paynter and Traylor, 1991)
Didelphis albiventris, 673 m
Philander opossum
Passo Quatro; Minas Gerais, Brazil (2228'S,
4505'W, Mustrangi and Patton, 1997)
Marmosops paulensis, 1500 m
Passos; Minas Gerais, Brazil (2043'S, 4637'W,
Varejao and Valle, 1982)
Caluromys philander
Didelphis albiventris (2043'S, 4635'W)
Monodelphis rubida, 728 m
Monodelphis sorex, 428 m
Pasto, 33 km E; Narino, Colombia (0113'N,
7716'W, Geonet)
Caenolestes fuliginosus
Patos de Minas; Minas Gerais, Brazil (1835'S,
4630'W, Varejao and Valle, 1982)
Didelphis albiventris
Patquia; La Rioja, Argentina (3003'S, 6653'W,
Paynter, 1985)
Thylamys pallidior, 43 1 m
Patuca; Gracias a Dios, Honduras (1548'N,
8417'W, Geonet)
Didelphis marsupialis
Philander opossum
Patuca River; Gracias a Dios, Honduras (1547'N,
8415'W, Geonet)
Didelphis marsupialis
Paucartambo; Cusco, Peru (1318'S, 7140'W,
Stephens and Traylor, 1983)
Lestoros inca, 2140 m, 2190 m, 2850 m, 3240
m, 10400 ft (= 3250 m) (10 km N and 32,
39, 48, 54 km NE of Paucartambo)
Marmosops impavidus, 1460 m (72 km NE of
Paucartambo)
Marmosops noctivagus (72 km NE of Paucar-
tambo)
Paujchi (see Pauji); Azuay, Ecuador
Paujf (= Paujchi); Azuay, Ecuador (0225'S,
7858'W, Bublitz, 1987)
Caenolestes caniventer
Paulo, Rio Cuquenan, Cerro Roraima; Bolivar,
Venezuela (ca. 0508'N, 6049'W, Paynter,
1982)
Marmosa murina, 1200 m
Pavol, 20 km NE of Mapastepec; Chiapas, Mex-
ico (Mapastepec = 1527'N, 9254'W, Geo-
net)
Didelphis marsupialis
Paya Camp; Darien, Panama (0758'N, 7724'W,
Geonet)
Metachirus nudicaudatus
Pazna, 10 km SW; Oruro, Bolivia (1840'S,
6659'W, Anderson, 1997)
Thylamys pallidior
Pebas; Loreto, Peru (0320'S, 7149'W, Stephens
and Traylor, 1983)
Marmosa murina, 101 m (Alto Maranon)
Marmosops noctivagus, 101 m
Metachirus nudicaudatus
Micoureus regina, 101 m (left bank Rio Ma-
ranon)
Pecanha; Minas Gerais, Brazil (1833'S, 4233'W,
Varejao and Valle, 1982)
Didelphis aurita, 780 m
Pedra; Pernambuco, Brazil (0832'S, 3658'W,
Cerqueira, 1984)
Didelphis albiventris
Pedra Branco, Parati; Rio de Janeiro, Brazil (ca.
2313'S, 4443'W, Varejao and Valle, 1982)
Philander frenata
Pedra Roxa (see Parque Nacional do Caparao);
Minas Gerais, Brazil
Pedregosa (see Quebrada La Pedregosa); Merida,
Venezuela
BROWN: ATLAS OF NEW WORLD MARSUPIALS
259
Pedro Leopoldo; Minas Gerais, Brazil (1937'S,
4405'W, Varejao and Valle, 1982)
Didelphis albiventris, 710 m
Pedro Miguel; Panama, Panama (0901'N,
7937'W, Geonet)
Chironectes minimus
Peixe-boi, Belem; Para, Brazil (0112'S, 4718'W,
Paynter and Traylor, 1991)
Chironectes minimus, 45 m
Didelphis marsupialis, 45 m
Philander opossum, 45 m (R. R. Braganca)
Pekin Abajo; Anzoategui, Venezuela (1010'N,
6430'W, Perez-Hernandez, 1989)
Didelphis marsupialis, 100 m
Penedo, right bank Rio Jurua; Amazonas, Brazil
(0650'S, 7045'W, Patton et al., 2000)
Didelphis marsupialis
Marmosops noctivagus
Metachirus nudicaudatus
Micoureus demerarae
Micoureus regina
Monodelphis emiliae
Philander mcilhennyi
Penha; Paraiba, Brazil (0710'S, 3448'W,
USBGN)
Didelphis marsupialis
Marmosa murina
Monodelphis domestica
Penha; Pernambuco, Brazil (0817'S, 3846'W,
USBGN)
Marmosa murina
Pen Hope, 20 mi E of Georgetown; Demerara-
Mahaica, Guyana (Georgetown = 0648'N,
5810'W, Geonet)
Marmosa murina
Peninsula de Paraguana (see Pueblo Nuevo); Fal-
con, Venezuela
Peninsula Valdes; Chubut, Argentina (4230'S,
6400'W, Paynter, 1985)
Thylamys pallidior
Peninsula Valiente; Bocas del Toro, Panama
(0905'N, 8151'W, Geonet)
Metachirus nudicaudatus (Punta Alegre)
Philander opossum (Punta Alegre, Quebrada
Hido)
Penuelas Forest Preserve, 7 km E of Las Tablas;
Valparaiso, Chile (3310'S, 7032'W, Geonet)
Thylamys elegans, 300 m
Peperpot; Commewijne, Suriname (0547'N,
5508'W, Geonet)
Marmosa murina
Perene; Junin, Peru (1058'S, 7518'W, Geonet)
Didelphis albiventris, 800-1000 m
Pergamino, Ruta 188, Curva de Pena; Buenos Ai-
res, Argentina (Pergamino = 3353'S,
6035'W, Paynter, 1985)
Monodelphis dimidiata
Perica; Marowijne, Suriname (Perica Kreek =
0547'N, 5508'W, Stephens and Traylor,
1985)
Phlander opossum
Perico, near, Maranon Valley; Amazonas, Peru
(not precisely located, but there is a Perico in
Cajamarca, near border of Amazonas =
0520'S, 7841'W, Geonet)
Marmosops noctivagus, ca. 1000 ft (= 300 m)
Perseverancia; Santa Cruz, Boliva (1438'S,
6237'W, Anderson, 1997)
Marmosops noctivagus
Peru Mines, El Callao (see El Peru); Bolivar, Ven-
ezuela
Perus, near; Sao Paulo, Brazil (2325'S, 4645'W,
Hershkovitz, 1992)
Gracilinanus microtarsus, sea level
Pesqueira; Pernambuco, Brazil (0822'S,
3642'W, USBGN)
Didelphis albiventris
Peulla, Lago Todos Santos; Los Lagos, Chile
(4106'S, 7202'W, Geonet)
Dromiciops gliroides
Rhyncholestes raphanurus
Petit Saut (see Crique Tigre); French Guiana
Picada de Olmos, 15 km SE, on road 16; Santiago
del Estero, Argentina (2606'S, 6210'W, Flo-
res et al., 2000)
Thylamys pusilla
Pica Periquito, El Portachuela, Rancho Grande
(see Estacion Biologica de Rancho Grande);
Aragua, Venezuela
Pichan; (see Volcan Pichincha); Pichincha, Ecua-
dor
Pichao, Sierra de Quilmas; Tucuman, Argentina
(2621'S, 6603'W, Flores et al., 2000)
Thylamys pallidior
Pichincha; Pichincha, Ecuador (0001'S, 7949'W,
Geonet)
Caenolestes fuliginosus
Pichinde; Valle del Cauca, Colombia (0326'N,
7637'W, Paynter, 1997)
Didelphis albiventris, 1600 m
Didelphis marsupialis, 1600 m
Pichucalco; Chiapas, Mexico (1731'N, 9304'W,
Geonet)
Chironectes minimus, 100 m
Didelphis marsupialis, 100 m
Pico Avila; Distrito Federal-Miranda, Venezuela
(ca. 1032'N, 6652'W, Paynter, 1989)
260
FIELDIANA: ZOOLOGY
Didelphis marsupialis, 1616-2232 m,
(1033'N, 6652'W)
Gracilinanus marica, 2124-2135 m (near Hotel
Humboldt)
Marmosops fuscatus, 1281-2232 m
Pico das Agulhas Negras, Itatiaya [Itatiaia] (=
Aguas Negras = Campo Bello = Monte Ser-
rat = Parque Nacional do Itatiaia); Rio de
Janeiro, Brazil (2223'S, 4438'W, Paynter
and Tray lor, 1991)
Didelphis aurita, 830-850 m
Lutreolina crassicaudata, 1150 m (Maromba)
Marmosa murina, 830 m
Marmosops incanus, 500 m, 2787 m
Micoureus demerarae, 1830 m (Macieiras)
Monodelphis americana, 2787 m
Monodelphis dimidiata, 815 m (Maromba),
1200 m (Maromba-Maua)
Monodelphis so rex
Philander frenata (Macieiras, Itatiaia, 2230'S,
4434'W)
Pico Salamanca; Chubut, Argentina (4535'S,
6720'W, Paynter, 1985)
Lestodelphys halli, 618 m
Picuma, Valle de Suaza; Huila, Colombia (Rio
Suaza = 0210'N, 7541'W, USBGN)
Didelphis marsupialis
Piedras Blancas; Beni, Bolivia (1315'S, 6420'W,
Anderson, 1997)
Caluromys lanatus
Didelphis marsupialis
Philander opossum
Piedras Coloradas; Tucuman, Argentina (2704'S,
6540'W, Mares et al., 1996)
Lutreolina crassicaudata
Piedras Negras; San Jose, Costa Rica (0955'N,
8419'W, USBGN)
Micoureus alstoni
Piedra Tendida; Tucuman, Argentina (2630'S,
6452'W, Mares et al., 1996)
Didelphis albiventris, 2500 ft (= 780 m)
Thylamys venustus, 2500 ft (= 780 m)
Pilar; Neembucii, Paraguay (2625'S, 5823'W,
Paynter, 1989)
Lutreolina crassicaudata
Pilcopata, 10 m W; Cusco, Peru (ca. 1308'S,
7110'W, Stephens and Traylor, 1983)
Marmosops noctivagus, 550 m
Piliguaya (see La Florida); La Paz, Bolivia
Pilimbala; Valle del Cauca, Colombia (not locat-
ed)
Didelphis albiventris, 2900 m
Pillahuata; Cusco, Peru (1308'S, 7125'W, Ste-
phens and Traylor, 1983)
Lestoros inca, 2450-2880 m
Pina; Panama, Panama (0916'N, 8000'W, Geo-
net)
Philander opossum
Pinalitos; Misiones, Argentina (2559'S, 5354'W,
Paynter, 1985)
Chironectes minimus, ca. 500 m
Pinamar; Uruguay (not located)
Didelphis albiventris
Lutreolina crassicaudata
Pinangu; Azuay, Ecuador (0226'S, 7858'W,
Bublitz, 1987)
Caenolestes caniventer
Pinas; El Oro, Ecuador (0342'S, 7942'W, Payn-
ter, 1993)
Micoureus regina, 905 m
Philander opossum, 3600 ft (= 1 125 m)
Pino (see Pinos); Tarija, Bolivia
Pinos (= Pino); Tarija, Bolivia (2136'S, 6454'W,
Paynter, 1992)
Thylamys venustus, 2000 m
Piquete (= Piquette); Sao Paulo, Brazil (2236'S,
4511'W, Paynter and Traylor, 1991)
Caluromys philander
Didelphis albiventris, 646 m
Didelphis aurita, 646 m
Micoureus demerarae, 646 m
Monodelphis scalops, 646 m
Monodelphis sorex, 646 m
Philander frenata
Piquette (see Piquete); Sao Paulo, Brazil
Piquiatuba, Rio Tapajos; Sao Paulo, Brazil
(0240'S, 5458'W, Hershkovitz, 1977)
Didelphis marsupialis
Piquirenda Viejo, 6 km W; Salta, Argentina
(2221'S, 6350'W, Flores et al., 2000)
Thylamys pusilla
Piracicaba; Sao Paulo, Brazil (2243'S, 4738'W,
Paynter and Traylor, 1991)
Chironectes minimus
Gracilinanus microtarsus, 548 m
Monodelphis americana
Monodelphis scalops, 527 m
Pira Pira; Carabobo, Venezuela (0957'N,
6804'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Monodelphis palliolata
Pirapolis (see Piriapolis); Maldonado, Uruguay
Pirapora; Minas Gerais, Brazil (1721'S, 4456'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 412 m
Piraqui; Alto Parana, Paraguay (not located)
Monodelphis brevicaudis
BROWN: ATLAS OF NEW WORLD MARSUPIALS
261
Pirassununga; Sao Paulo. Brazil (2159'S.
4725'W, Paynter and Traylor. 1991)
Lutreolina crassicaudata, 634 m
Pirayii-hi (see Arroyo Pirayu-f); Itapua. Paraguay
Pirebebuy (see Piribebuy); Paraguari. Paraguay
Piriapolis (= Pirapolis); Maldonado. Uruguay
(3454'S, 5517'W, Paynter, 1994)
Didelphis albiventris, sea level
Lutreolina crassicaudata. sea level
Piribebuy (= Pirebebuy); Paraguari. Paraguay
(2529'S, 5703'W, Paynter, 1989)
Chironectes minimus (27 km SW of Piribebuy)
Pirituba, near Sao Paulo: Sao Paulo. Brazil (ca.
2330'S, 4645'W, Paynter and Traylor. 1991 )
Monodelphis sorex
Piroculina (= Mattas do Piroculina), Chapada dos
Parecis; Rondonia. Brazil (1238'S. 6012'W.
Paynter and Traylor, 1991)
Metachirus nudicaudatus
Pirrita (see San Geronimo): Puntarenas. Costa
Rica
Pista Nova [Nueva], 8 km N of Porto Velho (see
Porto Velho); Rondonia. Brazil
Piste de Saint-Elie Ecerex Station (= Piste St.
Elie); French Guiana (0518'N, 5304'W. Ju-
lien-Laferriere. 1997)
Caluromys philander
Didelphis albiventris
Didelphis marsupialis
Mannosa marina
Philander opossum
Piste St. Elie (see Piste de Saint-Elie Ecerex Sta-
tion); French Guiana
Pitalito; Huila, Colombia (0151'N. 7602'W.
Paynter, 1997)
Caluromys lanatus
Pitiguaya (see La Florida): La Paz. Bolivia
Piura; Piura. Peru (0512'S, 8038'W. Stephens
and Traylor, 1983)
Didelphis marsupialis, 29 m
Plantage de Morgenstond (see "De Morgen-
stond"); Paramarino. Suriname
Plantation Clevia (see Clevia): Commewijne. Su-
riname
Plantation San Huberto. Tingo Maria: Huanuco.
Peru (Tingo Maria = 0918'S. 7559'W. Ste-
phens and Traylor. 1983)
Caluromys lanatus
Plantation Hope: Demerara-Mahaica. Guyana
(0642'N, 5758'W. Geonet)
Mannosa murina
Plantation "Mary's Hope." E of Totness: Coronie.
Suriname (0553'N, 5618'W. Geonet)
Micoureus demerarae
Platanales de los Campos; Merida. Venezuela (not
located)
Micoureus demerarae
Playa Ancha; Salta. Argentina (2207'S, 6344'W,
Geonet)
Thylamys venustus
Playa Colorado: Sucre. Venezuela (1016'N.
'6426'W. Perez-Hernandez. 1989)
Chironectes minimus, sea level
Playa del Rio Base. Cerro Duida; Amazonas. Ve-
nezuela (Cerro Duida = 0325'N. 6540'W.
Paynter. 1982)
Philander andersoni
Pocao: Pernambuco. Brazil (0811'S. 36 C 42'W.
USBGN)
Didelphis albiventris. 980 m
Monodelphis domestica, 980 m
Pocao. Boa Vista (see Boa Vista); Roraima. Brazil
Pocao. Caracaraf (see Caracaraf): Roraima. Brazil
Po^o das Antas. Reserva Biologica: Rio de Ja-
neiro. Brazil (2230'-2233'S. 4215'-
4219'W. Pires et al.. 1999)
Caluromys philander
Didelphis aurita
Micoureus demerarae
Poco Preto (see Estacao Biologica de Boraceia):
Sao Paulo. Brazil
Pocone; Mato Grosso. Brazil (1615'S. 5637'W.
Hershkovitz. 1997)
Gracilinanus agilis
Philander opossum (gallery forest)
Pocos de Caldas: Minas Gerais. Brazil (2148'S.
4634'W. Paynter and Traylor. 1991)
Didelphis aurita. 1 1 88 m
Polanco. Minas: Lavalleja. Uruguay (3353'S.
5503'W. Paynter. 1994)
Lutreolina crassicaudata
Pomara: Amazonas. Peru (0516'S, 7826'W, Ste-
phens and Traylor. 1983)
Metachirus nudicaudatus
Pomeroon; Pomeroon-Supenaam. Guyana (Pom-
eroon River = 0737'N. 5845'W, Stephens
and Traylor. 1985)
Philander opossum
Pomona: Stann Creek District. Belize (1659'N,
8822'W. Geonet)
Philander opossum (4 mi E of Pomona)
Pompeu: Minas Gerais. Brazil (1913'S, 4459'W.
Varejao and Valle. 1982)
Didelphis albiventris
Ponta Cumuruxatiba ( = "Comechatiba" = Com-
oxatiba = Cumuruxatiba): Bahia. Brazil
(1706'S. 39H'W. Paynter and Traylor,
1991)
262
FIELDIANA: ZOOLOGY
Metachirus nudicaudatus
Ponta de Boraceia; Sao Paulo, Brazil (2348'S,
4549'W, Hershkovitz, 1997)
Philander opossum
Ponta da Fruta; Espirito Santo, Brazil (2031'S,
4026'W, Paynter and Traylor, 1991)
Didelphis aurita, sea level
Ponte Nova; Minas Gerais, Brazil (2024'S,
4254'W, Paynter and Traylor, 1991)
Didelphis aurita, 402 m
Popoyan; Cauca, Colombia (0227'N, 7636'W,
Paynter, 1997)
Didelphis marsupialis, 1750 m
Porto Bello; Colon, Panama (0933'N, 7939'W,
Geonet)
Didelphis marsupialis
Porto de Moz; Para, Brazil (0145'S, 5214'W,
Paynter and Traylor, 1991)
Didelphis marsupialis, 30 m (Rio Xingu)
Monodelphis brevicaudata, 30 m
Porto Espiridao; Mato Grosso, Brazil (1551'S,
5828'W, Paynter and Traylor, 1991)
Caluromys lanatus, 372 m
Porto Faia, Rio Parana; Mato Grosso, Brazil (not
located)
Micoureus constantiae
Porto Feliz; Santa Catarina, Brazil (not located)
Philander frenata
Port-of-Spain; St. George, Trinidad and Tobago
(1039'N, 6131'W, Geonet)
Didelphis marsupialis
Porto Novo; Minas Gerais, Brazil (2153'S,
4242'W, Mustrangi and Patton, 1997)
Marmosops incanus, 160 m
Porto Real, near; Rio de Janeiro, Brazil (2225'S,
4420'W, Paynter and Traylor, 1991)
Gracilinanus microtarsus, sea level
Porto Seguro; Bahia, Brazil (1626'S, 3905'W,
Mustrangi and Patton, 1997)
Marmosops incanus, sea level
Porto Velho; Rondonia, Brazil (0846'S, 6354'W,
Paynter and Traylor, 1991)
Metachirus nudicaudatus (Pista Nova [Nueva],
8 km N of Porto Velho)
Philander opossum
Portovelo; El Oro, Ecuador (0343'S, 7939'W,
Paynter, 1993)
Didelphis marsupialis, 640 m
Marmosops impavidus, 2000 ft (= 625 m)
Porvenir, 1.3 km SW; Chuquisaca, Bolivia
(2045'S, 6313'W, coordinates given on
specimen tag)
Didelphis albiventris, 675 m
Monodelphis domestica
Thylamys venustus
Potaro; Potaro-Siparuni, Guyana (0523'N,
5908'W, Stephens and Traylor, 1985)
Micoureus demerarae
Monodelphis brevicaudata
Potrerillo; La Rioja, Argentina (2825'S, 6741'W,
Geonet)
Thylamys pallidior, 2600 m
Potrerillos, 3.6 km N; Salta, Argentina (2604'S,
6530'W, Flores et al., 2000)
Thylamys venustus
Potrero; Veracruz-Llave, Mexico (1852'N,
9650'W, Geonet)
Didelphis marsupialis, 1700 ft (= 530 m)
Philander opossum, 1700 ft (= 530 m) (8 km
NW of Potrero)
Potrero Viejo, 5 km W of Potrero; Veracruz-Lla-
ve, Mexico (1852'N, 9650'W, Geonet)
Didelphis marsupialis
Philander opossum, 1700 ft (= 530 m) (5 km
W of Potrero)
Powakka; Suriname, Suriname (0527'N,
5505'W, Stephens and Traylor, 1985)
Philander opossum
Pozo Ancho (town between Cartago and Pacaya);
Cartago, Costa Rica (Cartago = 0952'N,
8355'W, Pacaya = 0935'N, 8404'W Geo-
net)
Micoureus alstoni
Pozo Azul; Puntarenas, Costa Rica (0930'N,
8420'W, Gardner, 1973)
Didelphis marsupialis
Pozo del Tigre, Patino; Formosa, Argentina
(2454'S, 6019'W, Geonet)
Gracilinanus agilis
Pozo Hondo; Tucuman, Argentina (2749'S,
6520'W, Mares et al., 1996)
Didelphis albiventris
Pozuzo; Pasco [formerly Huanuco], Peru
(1004'S, 7532'W, Stephens and Traylor,
1983)
Chironectes minimus, 342 m, 1000 m
Didelphis marsupialis, 1000 ft (= 300 m)
Glironia venusta, 800 m
Marmosa murina, 1000 m
Marmosops noctivagus, 1000 m
Metachirus nudicaudatus, 752-800 m
Micoureus regina, 1000 m (Rio Pachitea)
Prados; Minas Gerais, Brazil (2103'S, 4405'W,
Varejao and Valle, 1982)
Didelphis albiventris
Didelphis aurita
Praia do Felix, Ubatuba; Sao Paulo, Brazil (Uba-
BROWN: ATLAS OF NEW WORLD MARSUPIALS
263
tuba = 2326'S, 4504'W. Paynter and Tray-
lor, 1991)
Philander frenata, 150 m
Pregonero; Tachira, Venezuela, (0801'N,
7145'W Geonet)
Monodelphis adusta, 1 100 m
Presidente Juscelino: Minas Gerais, Brazil
(1840'S, 4411'W, Hershkovitz. 1992)
Didelphis albiventris
Primeiro Morro: Sao Paulo, Brazil (2422'S.
4749'W)
Didelphis aurita
Gracilinanus microtarsus, ca. 25 m
Princesa Isabel; Parafba, Brazil (0744'S.
3800'W. USBGN)
Didelphis albiventris
Princes Town [Princestown]; Victoria, Trinidad
and Tobago (1016'N, 6123'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni, 125 ft (= 40 m)
Progreso: Chiriquf. Panama (0827'N. 8250'W.
Geonet)
Caluromys derbianus
Didelphis marsupialis ( 1 mi SW of Progreso)
Philander opossum
Prosperidade (= Cachoeira de Pancada); Amapa.
Brazil (ca. 0010'S, 5150'W, Paynter and
Traylor. 1991)
Didelphis marsupialis
Prudente de Morais: Minas Gerais. Brazil
(1929'S, 4411'W, USBGN)
Didelphis albiventris
Pucallpa; Ucayali, Peru (0823'S, 7432'W. Ste-
phens and Traylor. 1983)
Chironectes minimus, ca. 150 m (Rio Ucayali)
Didelphis marsupialis, ca. 150 m (56 km and
59 km SW of Pucallpa)
Metachirus nudicaudatus (57-59 km W of Pu-
callpa, W bank Rio Ucayali. 0828'S.
7436'W)
Monodelphis adusta, ca. 150 m
Philander mcilhennyi, 1180 m (59 km NE of
Rio Ucayali. 59 km SW and 59 km W of
Pucallpa)
Puca Tambo: San Martin. Peru (0610'S, 7716'W
Stephens and Traylor, 1983)
Marmosops impavidus, 1480 m
Pueblo Bello. Valledupar: Magdalena. Colombia
(1025'N, 7335'W Geonet)
Caluromys lanatus
Didelphis marsupialis, 1067 m
Pueblo Juarez (see Hacienda Magdalena): Colima.
Mexico
Pueblo Las Virtudes, 17 km from Caja Seca. Cas-
erio El Amparo; Merida, Venezuela
(0907'N. 7057'W, Geonet)
Metachirus nudicaudatus
Pueblo Nuevo (= Vitoc = Montana de Vitoc)
(also see Chanchamayo); Juni'n, Peru
(lTlO'S. 7516'W, Stephens and Traylor,
1983)
Marmosops impavidus (cerca de Chanchama-
yo)
Marmosops noctivagus, ca. 500 m
Pueblo Nuevo. 15-25 km SSW of Peninsula de
Paraguana: Falcon. Venezuela (1150'N,
6959'W. Handley, 1976)
Marmosa xerophila, 13-90 m
Pueblo Viejo [Puebloviejo] (see El Pueblito); La
Guajira. Colombia
Puebloviejo: Los Rios, Ecuador (0134'S,
7930'W, Paynter, 1997)
Philander opossum
Puente Alto. Rio Maipo; Santiago, Chile (3337'S,
7035'W Geonet)
Thylamys elegans, ca. 500 m
Puente de Ayuriquin (= Alluriquin): Pichincha,
Ecuador (0019'S, 7859'W, Paynter, 1997)
Caluromys derbianus, 700 m
Puente de Chimbo (= Rio Chimbo); Chimborazo,
Ecuador (ca. 0210'S, 7906'W Paynter,
1993)
Didelphis marsupialis, 350 m
Marmosa robinsoni, 345 m
Marmosops impavidus
Philander opossum, 1200 ft (= 375 m)
(0210'S, 7907'W)
Puente de Moromoro: El Oro. Ecuador (0350'S,
7950'W USBGN)
Chironectes minimus
Puerto Angel: Oaxaca, Mexico (1540'N,
9629'W, Geonet)
Marmosa canescens
Puerto Armuelles: Chiriquf, Panama (0817'N,
8252'W. Geonet)
Philander opossum
Puerto Asis: Putumayo, Colombia (0030'N,
7631'W, Paynter. 1997)
Didelphis marsupialis, 330 m
Philander andersoni, 330 m (17 km N of
Puerto Asis)
Puerto Ayacucho: Amazonas. Venezuela (0540'-
0506'N. 6745'-6735'W, Handley, 1976)
Caluromys lanatus, 126 m
Caluromys philander, 126 m (30 km S of
Puerto Ayacucho)
Chironectes minimus (0540'N, 6735'W)
264
FIELDIANA: ZOOLOGY
Didelphis albiventris, ca. 100 m (30 km S of
Puerto Ayacucho)
Didelphis marsupialis, 126-135 m (30-32 km
S of Puerto Ayacucho)
Micoureus demerarae, 126-135 m (30-32 km
S of Puerto Ayacucho)
Monodelphis brevicaudata, 114-135 m (30-32
km S of Puerto Ayacucho)
Puerto Bemberg; Misiones, Argentina (2558'S,
5430'W, Paynter, 1989)
Metachirus nudicaudatus
Puerto Bermejo; Chaco, Argentina (2656'S,
5830'W, USBGN)
Didelphis albiventris (10 km S of Puerto Ber-
mejo)
Philander opossum (10 km S of Puerto Ber-
mejo)
Puerto Bertoni, Alto Rio Parana; Alto Parana, Par-
aguay (2538'S, 5440'W, Paynter, 1989)
Caluromys lanatus
Chironectes minimus, 91 m
Metachirus nudicaudatus
Monodelphis dimidiata
Monodelphis sorex, 91 m
Thylamys pusilla, 9 1 m
Puerto Blest, Parque Nacional Nahuel Huapi; Rio
Negro, Argentina (4102'S, 7149'W, Paynter,
1985)
Dromiciops gliroides, 764 m, 770 m
Rhyncholestes raphanurus, 780 m
Puerto Caballo; Beni, Bolivia (1343'S, 6521'W,
Hershkovitz, 1997)
Gracilinanus agilis
Philander opossum
Puerto Cabello; Carabobo, Venezuela (1028'N,
6801'W, Paynter, 1982)
Marmosa robinsoni, sea level
Puerto Camacho, 15 km NW; Pando, Bolivia
(1128'S, 6750'W, Anderson, 1997)
Marmosops noctivagus
Puerto Carmen, Los Lagos; Chiloe, Chile
(4308'S, 7346'W, Geonet)
Dromiciops gliroides
Rhyncholestes raphanurus
Puerto Catatumbo; Zulia, Venezuela (0906'N,
7241'W, Geonet)
Metachirus nudicaudatus, 50 m
Philander opossum, 50 m
Puerto Centinela (see Centinela); Beni, Bolivia
Puerto Cortez (= Ciudad Cortes = Cortes); Pun-
tarenas, Costa Rica (0858'N, 8332'W, Geo-
net)
Caluromys derbianus
Puerto del Triunfo; Usulutan, El Salvador
(1317'N, 8833'W, Geonet)
Didelphis marsupialis, sea level
Philander opossum, sea level
Puerto Gaitan; Arauca, Colombia (0615'N,
7127'W, Paynter, 1997)
Lutreolina crassicaudata
Metachirus nudicaudatus
Puerto Golfito, 9 km ENE; Puntarenas, Costa Rica
(0839'N, 8309'W, Geonet)
Didelphis marsupialis
Puerto Huite (see Huite); Los Lagos, Chile
Puerto Iguazu (= Foz do Iguacu = Iguasu = Igua-
zu); Misiones, Argentina (2534'S, 5434'W,
Paynter, 1985)
Chironectes minimus, ca. 100 m
Didelphis albiventris (2533'S, 5435'W)
Didelphis aurita
Puerto Indiana, N bank Rio Maranon; Loreto,
Peru (ca. 0308'S, 7303'W, Stephens and
Traylor, 1983)
Metachirus nudicaudatus
Puerto Libertad (= Bemberg), Iguazu, Rfo Uru-
gua-i; Misiones, Argentina (2555'S,
5436'W, Geonet)
Gracilinanus agilis
Puerto Linares; La Paz, Bolivia (1529'S,
6731'W, Anderson, 1997)
Didelphis marsupialis
Marmosops noctivagus
Micoureus constantiae
Puerto Lobos; Chubut, Argentina (4203'S,
6508'W, Mares and Braun, 2000)
Thylamys pallidior
Puerto Maldonado; Madre de Dios, Peru (1236'S,
6911'W, Stephens and Traylor, 1983)
Caluromys lanatus (Reserva Cusco Amazoni-
co, 4 km E of Puerto Maldonado)
Didelphis marsupialis, 200 m (Reserva Cusco
Amazonico, 4 km E of Puerto Maldonado)
Metachirus nudicaudatus, 200 m (also at Re-
serva Cusco Amazonico, 4 km E of Puerto
Maldonado)
Philander opossum, 1 30-200 m (Reserva Cus-
co Amazonico, 4 km E and 12 km E of
Puerto Maldonado; left bank Rio Tambo,
1235'S, 6918'W)
Puerto Marquez (see San Marquez); Huanuco,
Peru
Puerto Montt; Los Lagos, Chile (4128'S,
7257'W, Geonet)
Dromiciops gliroides
Puerto Napo, Rio Napo (= Rio Napo); Napo, Ec-
uador (0103'N, 7747'W, Hershkovitz, 1997)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
265
Didelphis marsupialis
Philander andersoni
Puerto Octay; Los Lagos, Chile (4058'S,
7254'W, Geonet)
Rhyncholestes raphanurus, 140 m (15 km
NNW of Puerto Octay)
Puerto Paez; Apure, Venezuela (0613'N,
6728'W, Paynter, 1982)
Didelphis marsupialis
Puerto Piramides; Chubut, Argentina (4234'S,
6417'W, Mares and Braun, 2000)
Thylamys pallidior
Puerto Piray, km 18; Misiones, Argentina
(2628'S, 5442'W, Paynter, 1985)
Chironectes minimus, ca. 50 m
Puerto Quellon, Isla Chiloe (= Quelk5n); Los La-
gos, Chile (4307'S, 7337'W, Geonet)
Dromiciops gliroides, sea level, 20 m (15 km
S of Puerto Quellon, 4308'S, 7344'W, near
mouth of Rio Yaldad)
Rhyncholestes raphanurus (near mouth of Rio
Yaldad, 15 km SW of Quellon)
Puerto Salinas; Beni, Bolivia (1420'S, 6733'W,
Paynter, 1992)
Marmosa murina, 226 m
Puerto San Rafael (= San Rafael); Itapua, Para-
guay (ca. 2640'S. 5453'W, Paynter, 1989)
Chironectes minimus (2 km NNW of Puerto
San Rafael)
Didelphis albiventris
Puerto Schwelm; Misiones, Argentina (not locat-
ed)
Didelphis albiventris
Didelphis aurita
Puerto Siles; Beni, Bolivia (1248'S, 6505'W,
Hershkovitz, 1997)
Didelphis marsupialis
Philander opossum
Puerto Suarez, Sara Province; Santa Cruz, Bolivia
(1857'S, 5751'W, Paynter, 1992)
Gracilinanus agilis, 145 m
Puerto Valdivia; Antioquia, Colombia (0718'N,
7523'W, Paynter, 1997)
Didelphis marsupialis, 1 80 m
Puerto Viejo, Rio Sarapiqui; Heredia, Costa Rica
(1028'N, 8401'W, Geonet)
Chironectes minimus, 300 ft (= 90 m)
Didelphis marsupialis
Philander opossum, 90 m (1043'N, 8356'W)
Puesto Campo Grande, 17 km E of Santo Domin-
go; Salta, Argentina (2437'S, 6320'W; Flo-
res et al., 2000)
Thylamys pusilla
Puesto Perferacion; Santa Cruz, Bolivia (1955'S,
6234'W)
Thylamys pusilla
Puesto Punta del Agua, 2 km S; Mendoza, Ar-
gentina (3534'S, 6803'W, Mares and Braun,
2000)
Thylamys pallidior, 2700 ft (= 840 m)
Puesto San Antonio; Madre de Dios, Peru
(1239'S, 6644'W, Emmons et al., 1994)
Caluromys lanatus, ca. 200 m
Didelphis marsupialis, ca. 200 m
Philander opossum, ca. 200 m (W bank of Rio
Heath)
Puna (see Isla Puna); Guayas, Ecuador
Puna Island (see Isla Puna); Guayas, Ecuador
Punta Alegre, Peninsula Valiente; Bocas del Toro,
Panama (Peninsula Valiente = 0905'N,
8151'W, Geonet)
Caluromys derbianus
Didelphis marsupialis
Punta Arena, Peninsula Macanao; Nuevo Esparta,
Venezuela (1058'N, 6423'W, Lopez-Fuster
et al., 2000)
Marmosa robinsoni
Punta Cabito, Lago de Valencia; Carabobo, Ve-
nezuela (1011'N, 6751'W, Perez-Hernan-
dez, 1989)
Didelphis marsupialis, 415 m, 420 m
Marmosa robinsoni (1012'N, 6749'W)
Punta del Agua; Mendoza, Argentina (3531'S,
6805'W, Geonet)
Thylamys pallidior
Punta Lara; Buenos Aires, Argentina (3449'S,
5759'W, Paynter, 1985)
Didelphis albiventris, sea level
Lutreolina crassicaudata, sea level
Punta Norte (see Cayo de Agua); Bocas del Toro,
Panama
Punta Norte, Peninsula Valdes, 3 km S; Chubut,
Argentina (4204'S, 6345'W, Geonet)
Thylamys pallidior
Punta Rieles; Santa Cruz, Bolivia (1635'S,
6412'W, Anderson, 1997)
Micoureus regina
Punta Santa Ana; Loja, Ecuador (0350'S,
7925'W, Hershkovitz, 1997)
Philander opossum, 3650 ft (= 1140 m) (on
road from Zaruma to Loja)
Purace; Cauca, Colombia (0224'N, 7627'W,
Paynter, 1997)
Didelphis marsupialis, 2646-3650 m
Pun; Antioquia, Colombia (0725'N, 7520'W,
Paynter, 1997)
266
FIELDIANA: ZOOLOGY
Metachirus nudicaudatus, 140 m (Rio Pun, Rio
Cauca, 0724'N, 7520'W)
Quadro Penteado, 30 km from Sete Barras; Sao
Paulo, Brazil (Sete Barras = 2423'S,
4755'W, Paynter and Tray lor, 1991)
Didelphis aurita, 50 m
Quarry Heights; Panama, Panama (0857'N,
7933'W, Geonet)
Didelphis marsupialis
Quartel do Sacramento; Minas Gerais, Brazil
(1944'S, 4231'W, Hershkovitz, 1997)
Philander frenata
Quartermile Landing (= Kwatamang Landing),
Rupununi River, 5 km S of Anna; Rupununi
Guyana (0355'N, 5906'W, Patton, pers.
comm.)
Micoureus demerarae
Quebrada Aucayo; Loreto, Peru (0350'S,
7305'W, Geonet)
Monodelphis emiliae
Quebrada de Las Flores; San Juan, Argentina
(3140'S, 6808'W, Mares and Braun, 2000)
Thylamys pallidior
Quebrada de Lopez, San Francisco del Monte de
Oro (see San Francisco del Monte de Oro);
San Luis, Argentina
Quebrada de los Cuervos, 43 km N of Trienta y
Tres; Trienta y Tres, Uruguay (Trienta y Tres
= 3214'S, 5423'W, Paynter, 1994)
Didelphis albiventris, ca. 100 m
Quebrada de Lules; Tucuman, Argentina
(2652'S, 6525'W, Mares et al., 1966)
Lutreolina crassicaudata
Quebrada de Pichan; (see Volcan Pichincha); Pi-
chincha, Ecuador
Quebrada de San Lorenzo, 12 km NW of Salta;
Salta, Argentina (San Lorenzo [suburb of
Salta] = 2443'S, 6529'W, Paynter, 1985)
Thylamys pallidior
Quebrada Huarandosa, Rio Chinchipe; Cajamar-
ca, Peru (0512'S, 7848'W, Hershkovitz
files)
Metachirus nudicaudatus
Quebrada Kohuiri; Oruro, Bolivia (1808'S,
6857'W, Anderson, 1997)
Thylamys pallidior
Quebrada La Pedregosa (= La Pedregosa = Mon-
tes de Pedregoso = Pedregosa); Merida, Ve-
nezuela (0836'N, 7112'W, Paynter, 1982)
Gracilinanus marica, 1600 m (0837'N,
7142'W)
Marmosa robinsoni, 1600 m
Quebrada Matara; Apunmac, Peru (1345'S,
7254'W, Stephens and Traylor, 1983)
Didelphis albiventris, 6200 ft (= 1930 m)
Quebrada Morro Leguia; Cusco, Peru (ca.
1310'S, 7104'W, Pacheco et al., 1993)
Lestoros inca, 2100-2250 m
Quebrada Oran; Loreto, Peru (Oran = 0321'S,
7231'W, Geonet)
Didelphis marsupialis, 110 m
Philander andersoni, 110m
Quebrada Repressa, Puerto Obaldfa; San Bias,
Panama (Puerto Obaldia = 0840'N,
7725'W, Geonet)
Chironectes minimus
Quebrada Tasinchana; Pichincha, Ecuador
(0039'S, 7836'W, Bublitz, 1987)
Caenolestes fuliginosus, 3520-3640 m
Quebrada Vainilla (= Quebrada Vanilla), 10 km
SSW of mouth of Rio Napo; Loreto, Peru
(mouth Rio Napo = 0330'S, 7243'W, Geo-
net)
Glironia venusta (on east bank)
Quebrada Valdivia; Antioquia, Colombia
(0711'N, 7527'W, Paynter, 1997)
Marmosops parvidens, 900 m
Quebrada Vanilla (see Quebrada Vainilla); Loreto,
Peru
Quebrada Venado, Armila; San Bias, Panama (Ar-
mila = 0840'N, 7728'W, Geonet)
Chironectes minimus
Quebrangulo; Alagoas, Brazil (0920'S, 3629'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 411m
Quellon (see Puerto Quellon); Los Lagos, Chile
Quemapure, Brazo Casiquiare; Amazonas, Vene-
zuela (0229'N, 6627'W, Paynter, 1982)
Micoureus demerarae, ca. 100 in
Quillabamba, 90 km SE, below Abra Malaga;
Cusco, Peru (1310'S, 7232'W, Patterson
and Albuja, 1996)
Lestoros inca, 3540 m
Quillota; Valparaiso, Chile (3253'S, 7116'W,
Geonet)
Thylamys elegans, 128 m
Quilpue; Valparaiso, Chile (3302'S, 7126'W,
Geonet)
Thylamys elegans, ca. 100 m
Quincemil; Cusco, Peru (1316'S, 7038'W, Ste-
phens and Traylor, 1983)
Caluromys lanatus, 650 m
Caluromysiops irrupta, 650 m
Chironectes minimus, 650 m
Metachirus nudicaudatus, 680 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
267
Philander opossum, 680 m (1315'S, 7043'W)
Quindfo, western; Quindfo, Colombia (not locat-
ed)
Marmosops impavidus
Quine, 6 km NNE of Estancia Jahue; Santa Cruz,
Bolivia (1802'S, 6419'W, Anderson, 1997)
Thylamys ven ustus
Quine, 3.2 km NE, 17 km S of Estancia Laja:
Santa Cruz, Bolivia (1824'S, 6419'W, An-
derson, 1997)
Thylamys \ 'en ustus
Quininde (see Rosa Zarate); Esmeraldas, Ecuador
Quinisa, Chanchamayo; Junin, Peru (Chancha-
mayo = 1106'S, 7519'W, USBGN)
Didelphis marsupialis
Quiroga; Buenos Aires, Argentina (3516'S,
6125'W, Geonet)
Lutreolina crassicaudata
Quiros; Catamarca, Argentina (2847'S, 6507'W,
Mares et al., 1997)
Thylamys pallidior
Quispicanchi (Province); Cusco, Peru (1325'S,
7100'W, Stephens and Traylor, 1983)
Chironectes minimus
Quistococha [Quista Cocha], Maynes; Loreto,
Peru (0350'S, 7316'W, USBGN)
Didelphis marsupialis, 110 m
Philander andersoni
Quito; Pichincha, Ecuador (0013'S, 7830'W,
Paynter, 1993)
Didelphis albiventris, 2817 m
Quixada; Ceara, Brazil (0458'S, 3901'W, Payn-
ter and Traylor, 1991)
Monodelphis domestica, 190 m
Raco; Tucuman, Argentina (2638'S, 6256'W,
Paynter, 1985)
Lutreolina crassicaudata, 1000 m
Thylamys venustus, 1000 m
Raleigh Vallen (Falls); Sipaliwini, Suriname
(0443'N. 5612'W, Geonet)
Marmosa murina (E bank Coppename River)
Micoureus demerarae (E bank Coppename
River)
Monodelphis hrevicaudata
Rancho Grande (see Estacion Biologico de Ran-
cho Grande); Aragua, Venezuela
Rancho Grande, Auyan-tepuf; Bolfvar, Venezuela
(Auyan-tepuf = 0555'N, 6232'W, Paynter,
1985)
Monodelphis hrevicaudata
Rancho Grande Biological Station (see Estacion
Biologico de Rancho Grande); Aragua, Ve-
nezuela
Rancho Palo Amarillo, Amatlan; Sinaloa, Mexico
(not located)
Marmosa canescens
Rancho Pano Ayucutle, 6 mi N of Gomez Farias,
25 mi N of El Mante; Tamaulipas, Mexico
(2307'N, 9913'W, Alvarez, 1963)
Philander opossum, 300 ft (= 90 m)
Rancho Sabinal; San Luis Potosf, Mexico (Sabinal
= 2145'N, 9907'W, Geonet)
Didelphis marsupialis
Rancho Tambo, 61 km E; Tarija, Bolivia (2127'S,
6419'W, Anderson, 1997)
Didelphis albiventris, 2100 m
Thylamys venustus
Raudal Cerguera, Rio Autana; Amazonas, Vene-
zuela (0445'N, 6725'W, Perez-Hernandez,
1989)
Caluromys lanatus, 100 m
Micoureus demerarae
Raudal Chalimana; Bolivar. Venezuela (0410'N,
6259'W, Perez-Hernandez, 1989)
Philander opossum (Rio Paramichf, Rio Para-
gua)
Raudalito (see San Felipe); Amazonas, Venezuela
Realango, Serra da Piraquara; Rio de Janeiro, Bra-
zil (2252'S, 4326'W, Paynter and Traylor,
1991)
Metachirus nudicaudatus
Recife; Pernambuco, Brazil (0803'S, 3454'W,
Paynter and Traylor, 1991)
Marmosa murina, sea level
Recoleta, Asuncion; Central, Paraguay (Asuncion
= 2516'S, 5740'W, Paynter, 1989)
Didelphis albiventris
Reconcavo (= Bahia de Todos Santos); Bahia,
Brazil (ca. 1245'S, 3845'W, Paynter and
Traylor, 1991)
Didelphis aurita
Red Tank; Panama, Panama (0900'N, 7936'W,
Gardner, 1973)
Didelphis marsupialis
Refugio (see Volcan Osorno); Los Lagos, Chile
Refugio Juliaca; Madre de Dios, Peru (1257'S,
6852'W, Emmons et al., 1994)
Didelphis marsupialis, ca. 200 m
Refugio Military General Alvarado, 3 km W;
Mendoza, Argentina (not located)
Thylamys pallidior
Refugio, near Paso Coihue; Neuquen, Argentina
(Paso Coihue = 4058'S, 7121'W, Paynter,
1985)
Dromiciops gliroides, 1370 m
268
FIELDIANA: ZOOLOGY
Refugio Neumeyer, SSW San Carlos de Barilo-
che; Rio Negro, Argentina (San Carlos de
Bariloche = 4109'S, 7118'W, Geonet)
Dromiciops gliroides
Refugio Picoplancha; Madre de Dios, Peru
(1248'S, 6849'W, Emmons et al., 1994)
Micoureus regina (W bank Rio Heath)
Refugio, Valle de La Picada; Los Lagos, Chile
(4102'S, 7248'W, Geonet)
Dromiciops gliroides, 425-1135 m
Rhyncholestes raphanurus, 425 m, 450 m, 715
m, 820 m, 1000 m
Registo do Sai; Rio de Janeiro, Brazil (ca.
2256'S, 4400'W, Paynter and Traylor, 1991)
Didelphis aurita
Metachirus nudicaudatus
Represa de Guanapito, Altagracia de Orituco;
Guarico, Venezuela (0955'N, 6625'W,
Perez-Hernandez, 1989)
Chironectes minimus
Marmosa robinsoni
Republiek; Para, Suriname (0530'N, 5512'W,
Geonet)
Metachirus nudicaudatus
Reserva Biologica de Horco Molle (see Horco
Molle); Tucuman, Argentina
Reserva Biologica del Beni; Beni, Bolivia
(1451'S, 6621'W, Anderson, 1997)
Didelphis marsupialis
Reserva Biologica de Tingua, Nova Iguacii; Rio
de Janeiro, Brazil (2236'S, 4326'W, Mus-
trangi and Patton, 1997)
Marmosops incanus
Reserva Biologica Nova Lombardia; Santa Tere-
za, Brazil (ca. 1950'S, 4232'W, Mustrangi
and Patton, 1997)
Marmosops incanus, 500-1200 m
Reserva Cusco Amazonico, 14 km E of Puerto
Maldonado (see Puerto Maldonado); Madre
de Dios, Peru
Reserva Ecologica Biologica do Instituto de Geo-
grafia e Estatfstica (IBGE), 20 km S of Bra-
silia; Distrito Federal, Brazil (Brasilia =
1547'S, 4756'W, Paynter and Traylor, 1991)
Didelphis albiventris, 1 1 00 m
Gracilinanus agilis
Marmosa murina (35 km SE of Brasilia)
Monodelphis americana, ca. 1100 m
Reserva Ecologica de Nacunan (see Nacunan);
Mendoza, Argentina
Reserva Experimental de Flora y Fauna de Horco
Molle; Tucumam, Argentia (2645'S,
6221'W, Capllonch et al., 1997)
Didelphis albiventris
Reserva Forestal Campanha, Vale do Rio Doce;
Linhares, Brazil (1930'S, 4000'W, Mustran-
gi and Patton, 1997)
Marmosops incanus, 50 m
Reserva Forestal Imataca; Bolfvar, Venezuela (not
precisely located)
Monodelphis brevicaudata
Reserva Forestal Ticoporo; Barinas, Venezuela
(0800'N, 7003'W, Ldpez-Fuster et al.,
2000)
Marmosa robinsoni
Metachirus nudicaudatus
Philander opossum
Reserva Natural Estricta El Leoncito; San Juan,
Argentina (3146'S, 6910'W, Heinonen and
Chebez, 1997)
Thylamys pallidior
Reserva Natural Estricta Otamendi; Buenos Aires,
Argentina (3414'S, 5853'W, Heinonen and
Chebez, 1997)
Didelphis albiventris
Gracilinanus agilis
Lutreolina crasssicaudata
Reserva Natural Estricta San Antonio; Misiones,
Argentina (2605'S, 5346'W, Heinonen and
Chebez, 1997)
Didelphis aurita
Gracilinanus agilis
Monodelphis sorex
Reserva Natural Formosa; Formosa, Argentina
(2419'S, 6143'W, Heinonen and Chebez,
1997)
Didelphis albiventris
Monodelphis domestica
Reserva Provincial La Florida; Tucuman, Argen-
tina (2713'S, 6537'W, Flores et al., 2000)
Thylamys venustus
Restrepo (= Caney = Canay); Meta, Colombia
(0415'N, 7333'W, Paynter, 1997)
Didelphis marsupialis (Caney, Rio Guatiqufa)
Metachirus nudicaudatus
Philander opossum
Reyes; Beni, Bolivia (1419'S, 6723'W, Paynter,
1992)
Gracilinanus agilis, 196 m
Riacho Pilaga (= Estancia Linda Vista); Formosa,
Argentina (ca. 2513'S, 5747'W, Paynter,
1985)
Gracilinanus agilis, 100 m
Philander opossum
Ribeirao Bonito; Sao Paulo, Brazil (2204'S,
4810'W, Paynter and Traylor, 1991)
Lutreolina crassicaudata, 585 m
Ribeirao da Fortuna, Ilheus near Buerama; Bahia,
BROWN: ATLAS OF NEW WORLD MARSUPIALS
269
Brazil (Buerama 1457'S, 3919'W.
USBGN)
Metachirus nudicaudatus
Ribeirao da Lagoa, 20 km S of Salesopolis; Sao
Paulo, Brazil (Salesopolis 2332'S.
4551'W, Geonet)
Monodelphis sorex
Ribeirao das Pedras (= Ribeirao Pedra), near
Raneho Pedra, Rio Pardo; Minas Gerais, Bra-
zil (1805'S, 4340'W, USBGN)
Chironectes minimus
Ribeirao do Varjao (see Varjao); Sao Paulo, Brazil
Ribeirao Fundo; Sao Paulo, Brazil (ca. 2415'S,
4745'W, Paynter and Tray lor, 1991)
Didelphis aurita, 7(K) m
Ribeirao Jordao (see Rio Jordao); Minas Gerais,
Brazil
Ribeirao Pedra (see Ribeirao das Pedras); Minas
Gerais, Brazil
Ribeirao Preto; Sao Paulo, Brazil (2110'S,
4748'W, Paynter and Traylor, 1991)
Lutreolina crassicaudata, 518m
Riberalta; Beni, Bolivia (1059'S, 6606'W, An-
derson, 1997)
Didelphis marsupialis
Philander opossum
Ricaurte; Narifio, Colombia (0113'N, 7759'W,
Paynter, 1997)
Marmosops impavidus, 5000 ft (= 1560 m)
Micoureus regina, 5000 ft (= 1560 m)
Richmond; Trinidad and Tobago (1113'N,
6037'W, Geonet)
Marmosa murina
Rijweg (see Paramaribo); Paramaribo, Suriname
Rincao; Sao Paulo, Brazil (2135'S, 4805'W,
Paynter and Traylor, 1991)
Didelphis albiventris
Rincon, Corcovado National Park; Puntarenas,
Costa Rica (0842'N, 8329'W, Geonet)
Chironectes minimus
Didelphis marsupialis (0842'N, 8328'W)
Rincon (see El Rincon); San Luis, Argentina
Rio Acara; Para, Brazil (0140'S, 4825'W, Payn-
ter and Traylor, 1991)
Metachirus nudicaudatus
Rio Aceramarca (see Rio Aceromarca), Rio Un-
duavi, Yungas; La Paz, Bolivia
Rio Aceromarca (= Rio Aceramarca), Rio Un-
duavi, Yungas; La Paz, Bolivia (1618'S,
6753'W, Paynter, 1992)
Didelphis albiventris
Gracilinanus aceramarcae, 2600 m, 10800 ft
( = 3370 m)
Rio Acima; Minas Gerais. Brazil (2007'S,
4315'W, Varejao and Valle, 1982)
Didelphis albiventris
Rfo Aguaray-Guazu, right bank tributary Rio Par-
aguay; Presidente Hayes. Paraguay (24 47 S,
5719'W, Paynter, 1989)
Lutreolina crassicaudata (1.2 km N of Rio
Aguaray-Guazu)
Rio Aguas Claras, near San Adolfo, Acevedo;
Huila, Colombia (0145'N, 7555'W, Pine,
1981)
Marmosops parvidens, 1400 m, 1700 m^
Rio Albarregas; Merida, Venezuela (0835'N,
7109'W, Paynter, 1982)
Gracilinanus marica, 1630 m
Marmosops fuscatus, 1630 m
Rio Alto Madre de Dios, Parque Nacional del
Manu; Madre de Dios, Peru (1217'S,
7052'W, USBGN)
Caluromxsiops irrupta
Rfo Alto Puriis; Ucayali, Peru (0934'S, 7036'W,
USBGN)
Chironectes minimus
Rio Amaca-Yacu [Rio Amacayacu]; Amazonas,
Colombia (0330'N, 7015'W, Lemke, 1982)
Marmosa lepida, 100 m
Rio Amaparf, Macapa; Amapa, Brazil (0043'N,
5132'W, Paynter and Traylor, 1991)
Marmosa murina
Marmosops pinheiroi
Monodelphis brevicaudata
Philander opossum (Serra do Navio)
Rio Amaya, boca, W bank Rio Morona; Loreto,
Peru (0412'S, 7710'W, Hershkovitz files)
Metachirus nudicaudatus, 200 m
Rio Anaure (= Aurare); Zulia, Venezuela
(1036'N, 7130'W, Paynter, 1982)
Didelphis marsupialis, sea level
Rfo Anchicaya, alto; Valle del Cauca, Colombia
(0336'N, 7654'W, Timm, pers. comm.)
Philander opossum
Rfo Andalgala; Catamarca, Argentina (2748'S,
6617'W, Mares et al., 1997)
Thylamys pallidior
Rfo Andira, boca; Amazonas, Brazil (0245'S,
5649'W, Paynter and Traylor, 1991)
Micoureus demerarae
Rio Aquidaban at Paso Horqueta; Concepcion,
Paraguay (Paso Aquidaban = 2307'S,
5720'W, USBGN)
Philander opossum
Rio Araguari, Macapa; Amapa, Brazil (0115'N,
4955'W, Paynter and Traylor, 1991)
Caluromys philander (lower Rfo Araguari)
270
FIELDIANA: ZOOLOGY
Micoureus demerarae
Monodelphis brevicaudata (lower Rio Aragu-
ari)
Rio Arauca; Arauca, Colombia (0703'-1001'N,
7209'-7158'W, Paynter, 1997)
Didelphis marsupialis, 1000 m, 1100 m
Rio Aripuana [Aripuanan]; Amazonas, Brazil
(0507'S, 6024'W, USBGN)
Marmosops noctivagus
Rio Atoyac; Veracruz-Llave, Mexico (1902'N,
9608'W, Geonet)
Philander opossum
Rio Axtla; San Luis Potosi, Mexico (Axtla =
2128'N, 9851'W, Geonet)
Didelphis marsupialis
Rio Balcones (see Guasca, Rio Balcones); Cun-
dinamarca, Colombia
Riobamba; Chimborazo, Ecuador (0140'S,
7838'W, Paynter, 1997)
Didelphis albiventris, 2754 m
Rio Bariatou [Barratow], near Cali (see Cali); Val-
le del Cauca, Colombia
Rio Barragan, cerca de la desembocadura Barin-
tina; Barinas, Venezuela (0840'N, 7025'W,
Perez-Hernandez, 1989)
Marmosa robinsoni, 440 m
Monodelphis palliolata, 440 m
Rio Baudo, Rio Sando; Chocd, Colombia
(0457'N, 7722'W, Paynter, 1997)
Chironectes minimus
Rio Baures, mouth; Beni, Bolivia (1230'S,
6418'W, Anderson, 1997)
Gracilinanus agilis
Rio Beni; Beni, Bolivia (1023'S, 6524'W, Payn-
ter, 1992)
Didelphis marsupialis, 240 m
Rio Beni; La Paz, Bolivia (1315'S, 6718'W, An-
derson, 1997)
Didelphis marsupialis
Philander opossum
Rio Bermejo; Napo, Ecuador (0017'N, 7706'W,
Paynter, 1993)
Caenolestes fuliginosus
Rio Birongo, Birongo; Miranda, Venezuela
(1029'N, 6614'W, Perez-Hernandez, 1989)
Chironectes minimus
Monodelphis palliolata, 60 m
Rio Blanco; Jujuy, Argentina (2425'S, 6523'W,
Flores et al., 2000)
Didelphis albiventris
Thylamys venustus, 1443 m
Rio Blanco, Piedras Negras; Veracruz-Llave,
Mexico (1845'N, 9600'W, Geonet)
Didelphis marsupialis, 400 ft (= 125 m) (20
km W of Piedras Negras)
Philander opossum, 400 ft (= 125 m) (20 km
WNW of Piedras Negras)
Rio Branco, Mazagao; Amapa, Brazil (0007'S,
5117'W, Hershkovitz, 1997)
Philander opossum (tributary Rio Maraca)
Rio Branco; Roraima, Brazil (not precisely locat-
ed)
Caluromys philander
Rio Briceno; Manabf, Ecuador (0031'S, 8027'W,
USBGN)
Marmosa robinsoni
Rio Bubi, near Sona; Veraguas, Panama (Sona =
0801'N, 8119'W, Geonet)
Chironectes minimus
Rio Bubi, Rio Viejo; Veraguas, Panama (0758'N,
8137'W, Geonet)
Chironectes minimus
Rio Burgua; Tachira, Venezuela (0726'N,
7152'W, Paynter, 1982)
Caluromys lanatus
Rio Cachimayo; Potosi, Bolivia (1918'S,
6612'W, Paynter, 1982)
Didelphis albiventris, 3700 ft (= 1150 m)
Rio Cahy (see Rio Cai); Rio Grande do Sul, Bra-
zil
Rio Cai (= Rio Cahy); Rio Grande do Sul, Brazil
(2956'S, 5116'W, Paynter and Traylor,
1991)
Caluromys lanatus
Rio Caoni; Pichincha, Ecuador (0012'N,
7923'W, Paynter, 1993)
Philander opossum
Rio Capihuara; Pastaza, Ecuador (0231'S,
7651'W, Paynter, 1993)
Metachirus nudicaudatus, 300 m
Rio Caris, S of El Tigre; Anzoategui, Venezuela
(0836'N, 6413'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
Rio Casiquiare; Amazonas, Venezuela (0234'N,
6718'W, Perez-Hernandez, 1989)
Didelphis marsupialis
Micoureus demerarae, 130 m
Monodelphis brevicaudata
Philander andersoni, 1 30 m
Rio Castano Overo, W of San Carlos de Barilo-
che; Rio Negro, Argentina (San Carlos de
Bariloche = 4109'S, 7118'W, Paynter,
1985)
Dromiciops gliroides
Rio Cauca (= Cauca Seca) (also see Cali); Valle
del Cauca, Colombia
BROWN: ATLAS OF NEW WORLD MARSUPIALS
271
Rio Cauca Valley (see Cali); Valle del Cauca, Co-
lombia
Rio Cauquita, few km S of Cali; Valle del Cauca.
Colombia (Cali = 0327'N, 7631'W, Payn-
ter, 1997)
Caluromys derbianus
Marmosops impavidus, KKX) m
Rio Cayumba; Junin, Peru (not located)
Marmosops noctivagus
Rio Cayumba, Rio Huallaga: Huanuco, Peru
(0927'S, 7558'W, Stephens and Traylor,
1983)
M icon reus regina, 3(KK) ft (= 930 m)
Rio Ceballos; Cordoba, Argentina (3107'S,
6418'W. Pineet al 1985)
Monodelphis dimidiata, 500 m
Rio Cenepa, Huampami (also see Huampami):
Amazonas, Peru (0439'S, 7808'W, Geonet)
Chironectes minimus (N of Huampanf)
Philander andersoni (NW of Huampanf)
Rio Chagres; Colon/Panama, Panama (0907'N,
7942'W, Geonet)
Didelphis marsupialis
Rio Challana; La Paz, Bolivia (1528'S, 67 51'W,
USBGN)
Metachirus nudicaudatus (SE of Guanay)
Rio Chama (see Cafetal de Chama); Merida, Ve-
nezuela
Rfo Chamicuros (= Chimicuros), a tributary of
the Maranon between Rios Huallaga and
Ucayali: Loreto, Peru (Chamicuros = ca.
0530'S, 7530'W, Stephens and Traylor,
1983)
Caluromys lanatus
Rio Chanchan; Chimborazo, Ecuador (0217'S,
7924'W, Paynter. 1993)
Marmosops impavidus
Rfo Changena Camp; Bocas del Toro, Panama
(Rfo Changena = 0906'N, 8234'W, Wenzel
and Tipton. 1966)
Philander opossum
Rfo Changuinola; Bocas del Toro, Panama
(0928'N, 8227'W, Geonet)
Caluromys derbianus
Chironectes minimus
Rfo Chapare (see also Rfo Mamore); Cochabam-
ba-Santa Cruz, Bolivia (1558'S, 6442'W,
Paynter. 1982)
Didelphis marsupialis (54 km S of mouth of
Rfo Chapare)
Rfo Chico; Chaco, Argentina (not located)
Philander opossum
Rfo Chico; Miranda, Venezuela (1020'N,
6556'W, Geonet)
Didelphis marsupialis, sea level, 1 m (1 km S
and 1-6 km SSE of Rfo Chico)
Monodelphis palliolata, 1 m (6 km SSE of Rfo
Chico)
Rfo Chimate, Rfo Mapiri; La Paz, Bolivia (ca.
1525'S, 6800'W, Paynter, 1992)
Metachirus nudicaudatus, 1900 ft ( = 590 m)
Rfo Chimbo, Coto; Chimborazo, Ecuador
(0217'S, 7924'W, Paynter, 1993)
Didelphis marsupialis, 350 m
Rfo Chimbo. junction of Rfo Coco (see Puente de
Chimbo); Chimborazo, Ecuador
Rfo Chinchao; Huanuco, Peru (0930'S, 7556'W,
Stephens and Traylor, 1983)
Marmosa murina, ca. 1000 m
Marmosops noctivagus
Micoureus regina, ca. 1000 m (Rfo Huallaga)
Rfo Choronf; Aragua, Venezuela (Choronf =
1029'N, 6737 7 W, Paynter, 1982)
Chironectes minimus
Rfo Chucunaque; Darien, Panama (0809'N,
7744'W, Geonet)
Didelphis marsupialis
Rfo Cinaruco, 38 km NNW of Puerto Paez; Apu-
re, Venezuela (0631'N, 6746'W, Handley,
1976)
Didelphis marsupialis, 76 m
Rfo Coaque (see El Destino, Rfo Coaque); Man-
abf, Ecuador
Rfo Cobarfa (= Rfo Covarfa); Boyaca, Colombia
(0703'N, 7204'W, Paynter, 1997)
Chironectes minimus, 1000 ft (= 300 m)
Metachirus nudicaudatus, 1000 ft (= 300 m)
Philander opossum, 1100 ft (= 340 m) (near
mouth, above town of Covarfa [Cobarfa])
Rfo Coca; Napo, Ecuador (0029'S, 7658'W,
Paynter, 1997)
Marmosops impavidus
Marmosops noctivagus
Rfo Coco (= Rfo Segovfa = Segovia River); bor-
der between Honduras and Nicaragua (boca
Rfo Coco = 1500'N, 8310'W, Geonet)
Didelphis marsupialis (in Zelaya, Nicaragua)
Marmosa mexicana (in Zelaya, Nicaragua)
Micoureus alstoni (in Hondorus)
Philander opossum
Rfo Colorado; Chiriquf, Panama (0849'N,
8243'W, Geonet)
Chironectes minimus, 4000 ft (= 1250 m)
Rfo Colorado, near (see Curacautfn); Araucania,
Chile
Rfo Colorado, 30 km E of Guayacan; Santiago,
Chile (Guayacan = 3336'S, 7022'W, Geo-
net)
272
FIELDIANA: ZOOLOGY
Thylamys elegans
Rio Comberciato; Cusco, Peru (1228'S, 7307'W,
Stephens and Traylor, 1983)
Caluromys lanatus, 2000 ft (= 625 m)
Rio Copataza; Pastaza, Ecuador (0207'S,
7727'W, Paynter, 1993)
Marmosops noctivagus
Philander andersoni, 650 m
Rio Covaria (see Rio Cobaria); Boyaca, Colombia
Rio Cumboto; Aragua, Venezuela (1029'N,
6746'W, Paynter, 1982)
Chironectes minimus
Rio Cunucunuma, Belen; Amazonas, Venezuela
(0313'N, 6558'W, Paynter, 1989)
Caluromys lanatus, 150 m
Chironectes minimus, 150 m
Philander andersoni, 1 50 m
Rio Curanja; Ucayali, Peru (0958'S, 7058'W,
Stephens and Traylor, 1985)
Chironectes minimus (near Brazilian border)
Didelphis marsupialis
Micoureus demerarae (near Balta)
Rio Curaray; Pastaza, Ecuador (not precisely lo-
cated)
Metachirus nudicaudatus
Rio Curaray; Loreto, Peru (0222'S, 7405'W, Ste-
phens and Traylor, 1983)
Didelphis marsupialis
Marmosa murina (Oriente, boca)
Marmosa rubra
Marmosops impavidus (at Rio Napo)
Marmosops noctivagus
Metachirus nudicaudatus
Micoureus regina
Philander andersoni
Rio Curaray and Napo junction; Loreto, Peru
(0222'S, 7405'W, Stephens and Traylor,
1983)
Glironia venusta (Anthony gives location as
Ecuador, but it is in Peru)
Rio Curiche; Choco, Colombia (0702'N,
7740'W, Geonet)
Didelphis marsupialis
Rio Curinhuas (see Rio Kurinwas); Zelaya, Nic-
aragua
Rio Curua do Sul (see Foz do Curua); Para, Brazil
Rio das Mortes; Mato Grosso, Brazil (1145'S,
5044'W, Paynter and Traylor, 1991)
Didelphis albiventris
Rio das Velhas, near Lagoa Santa; Minas Gerais,
Brazil (Lagoa Santa = 1938'S, 4353'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 760 m
Gracilinanus agilis
Rio das Velhas, tributary Rio Sao Francisco; Mi-
nas Gerais, Brazil (1713'S, 4449'W, Paynter
and Traylor, 1991)
Didelphis aurita
Monodelphis americana
Rio de Janeiro; Rio de Janeiro, Brazil (2254'S,
4314'W, Paynter and Traylor, 1991)
Chironectes minimus, near sea level (near Rio
de Janeiro)
Didelphis aurita
Gracilinanus microtarsus, near sea level
Marmosops paulensis
Metachirus nudicaudatus
Monodelphis scalops
Philander frenata
Rio de La Plata; NE Argentina, parts of Uruguay
and Paraguay (not precisely located)
Chironectes minimus
Didelphis albiventris
Rio de Las Conchas; Salta, Argentina (2518'S,
1601'W, Flores et al., 2000)
Thylamys venustus
Rio de Oro; Chaco, Argentina (2704'S, 5834'W,
Hershkovitz, 1997)
Philander opossum (boca)
Rio de Oro; Manabi, Ecuador (0210'N, 7922'W,
Hershkovitz, 1997)
Philander opossum
Rio de Oro, boca, 60 km WNW of Encontrados;
Zulia, Venezuela (0906'N, 7245'W, Hersh-
kovitz, 1997)
Philander opossum, 73 m
Rio do Braco, Ilheus; Bahia, Brazil (1438'S,
3914'W, USBGN)
Didelphis aurita
Metachirus nudicaudatus
Rio Docampado; Choco, Colombia (0445'N,
7718'W, Paynter, 1997)
Chironectes minimus, 350 m
Philander opossum, 75-160 m
Rio Doce; Espirito Santo, Brazil (1937'S,
3949'W, Hershkovitz files)
Metachirus nudicaudatus
Rio Doce State Forestry Park (see Parque Estad-
ual do Rio Doce); Minas Gerais, Brazil
Rio Doce State Park (see Parque Estadual do Rio
Doce); Minas Gerais, Brazil
Rio Don Diego, 45 km E of Santa Marta; Mag-
dalena, Colombia (1115'N, 7342'W, Payn-
ter, 1997)
Marmosa robinsoni
Rio Eirii, Rio Jurua; Amazonas, Brazil (0642'S,
6952'W, Paynter and Traylor, 1991)
Caluromys lanatus
BROWN: ATLAS OF NEW WORLD MARSUPIALS
273
Rio Escondido, 50 mi above Bluefields; Zelaya,
Nicaragua (1204'N. 8545'W, Geonet)
Didelphis marsupialis
Philander opossum
Rfo Frio [Rfofrio]; Valle del Cauca, Colombia
(0409'N, 7618'W. Paynter, 1997)
Chironectes minimus
Didelphis marsupialis, 960 m
Philander opossum, 3500 ft (= 1090 m)
Rio Gomes, 7 km E of downtown San Francisco
del Monte del Oro; San Luis, Argentina (San
Francisco del Monte del Oro = 3236'S,
6608'W, Geonet)
Thylamys pallidior, 2800 ft (= 876 m)
Rio Grande, 5 km NE of mouth; Santa Cruz, Bo-
livia (ca. 1551'S, 6437'W, Paynter, 1992)
Didelphis marsupialis
Rio Guaillabamba, trib., N of Quito; Pichincha,
Ecuador (0028'S, 7925'W, Paynter, 1993)
Chironectes minimus, 2106 m
Rio Guaimaral, Valledupar, 5 km E of El Orinoco;
Cesar, Colombia (0949'N, 7337'W, Geonet)
Didelphis marsupialis, 140 m
Rio Guapaya (see Parque de La Macarena); Meta,
Colombia
Rio Guarico, near San Juan de los Morros; Gua-
rico, Venezuela (1000'N, 6742'W, Geonet)
Chironectes minimus
Rio Guayabero; Meta, Colombia (0236'N,
7247'W, Geonet)
Didelphis marsupialis
Metachirus nudicaudatus (above mouth of Rio
Ariari)
Rio Hondo, Samana, Cordillera Central; Caldas,
Colombia (0542'N, 7501'W, Paynter, 1997)
Philander opossum
Rio Huixtla, ca. 13 km N of Huixtla; Chiapas,
Mexico (1501'N, 9232'W, Geonet)
Didelphis marsupialis
Rio Ibare, boca; Beni, Bolivia (1437'S, 6457'W,
Paynter, 1992)
Didelphis marsupialis
Philander opossum (also at 1446'S, 6451'W,
and 1448'S, 6414'W)
Ri'o Ichilo; Cochabamba, Bolivia (1650'S,
6445'W, Hershkovitz, 1997)
Philander opossum (52 km S of mouth, also at
1628'S, 6445'W)
Rio Ichilo, 150 km from mouth of Rio Chimore,
ca. 9 km S of mouth Rio Chapare; Cocha-
bamba-Santa Cruz. Bolivia (1629'S.
6444'W, Anderson, 1997)
Didelphis marsupialis
Metachirus nudicaudatus
M icon reus const an tiae
Rio Iguazii, 60 km E of Puerto Iguazii; Misiones,
Argentina (2536'S, 5436'W, USBGN)
Didelphis aurita
Rfo Imamado (see Jaque); Darien, Panama
Ri'o Inambari; Madre de Dios, Peru (1241'S,
6944'W. Stephens and Traylor, 1983)
Metachirus nudicaudatus
Rfo Indio, near Gatiin; Colon, Panama (0916'N,
7957'W, Geonet)
Didelphis marsupialis
Marmosa robinsoni (0917'N, 7958'W)
Rfo Inio, boca, Isla Chiloe; Los Lagos, Chile
(4321'S, 7407'W, Geonet)
Dromiciops gliroides
Rhyncholestes raphanurus
Rio Ipanema (see Ipanema); Sao Paulo, Brazil
Rio Itapocu (= Rfo Novo do Itapocu), near; Santa
Catarina, Brazil (2634'S, 4840'W, USBGN)
Chironectes minimus
Gracilinanus microtarsus
Rfo Itata; Los Lagos, Chile (3623'S, 7252'W,
Geonet)
Dromiciops gliroides
Rio Itauna (= Rio Itaunas), mouth at; Espfrito
Santo, Brazil (1825'S, 3942'W, Paynter and
Traylor, 1991)
Didelphis aurita
Rio Itaunas (= Rio Itauna); Espfrito Santo, Brazil
Rfo Itenez; Beni, Bolivia (1229'S, 6415'W, An-
derson, 1997)
Micoureus constantiae
Rfo Itenez, opposite Costa Marques, Brazil; Beni,
Bolivia (1228'S, 6416'W, Paynter, 1992)
Didelphis marsupialis
Micoureus constantiae
Rfo Itenez, SW Costa Margurez, Brazil; Beni, Bo-
livia (1230'S, 6418'W, Anderson, 1997)
Didelphis marsupialis
Rio Ivinheima (= Ivinheima); Mato Grosso do
Sul. Brazil (2314'S, 5342'W, Paynter and
Traylor, 1991)
Didelphis alhiventris
Didelphis aurita
Lutreolina crassicaudata
Rioja; San Martfn, Peru (0610'S, 7710'W,
Hershkovitz, 1997)
Philander opossum
Rfo Jaque, Jaque (see Jaque); Darien, Panama
Rio Jaru, upper; Rondonia, Brazil (1056'S,
6304'W, de Vivo and Gomes, 1989)
Caluromysiops irrupta
Rfo Jau, above mouth, right bank; Amazonas,
274
FIELDIANA: ZOOLOGY
Brazil (0157'S, 6129'W, Patton, pers.
comm.)
Didelphis marsupialis
Metachirus nudicaudatus
Micoureus demerarae
Philander andersoni
Rio Jauru, upper Rio Paraguay; Mato Grosso,
Brazil (1622'S, 5746'W, Paynter and Tray-
lor, 1991)
Caluromys lanatus
Metachirus nudicaudatus (1530'S, 5833'W,
Hershkovitz files)
Rio Javari; Amazonas, Brazil (0421'S, 7002'W,
Paynter and Traylor, 1991)
Marmosa murina
Rio Jojui (see Rio Mojui); Para, Brazil
Rio Jordao (= Ribeirao Jordao); Minas Gerais,
Brazil (1826'S, 4806'W, Paynter and Tray-
lor, 1991)
Chironectes minimus
Didelphis albiventris
Micoureus constantiae
Rio Jordao; Parana, Brazil (2546'S, 5207'W,
Paynter and Traylor, 1 99 1 )
Micoureus demerarae
Rio Juquia, barra (see Juquia); Sao Paulo, Brazil
Rio Jurare (see Mato Grosso); Mato Grosso, Bra-
zil
Rio Jurua; Amazonas, Brazil (0237'S, 6544'W,
Paynter and Traylor, 1991)
Didelphis marsupialis
Rio Jurua, upper; Acre, Brazil (ca. 0008'S,
7300'W, Hershkovitz files)
Chironectes minimus
Rio Kagka, headwaters; Amazonas, Peru (ca.
0433'S, 7816'W, Stephens and Traylor,
1983)
Didelphis marsupialis, 2600 ft (= 800 m)
Rio Kurinwas (= Rio Curinhuas = Kurinwas Riv-
er); Zelaya, Nicaragua (1249'N, 8341'W,
Geonet)
Didelphis marsupialis
Philander opossum (1245'N, 8343'W)
Rio Lagoa Juparana; Espirito Santo, Brazil (Lagoa
Juparana = 1935'S, 4018'W, Paynter and
Traylor, 1991)
Micoureus demerarae
Rio Las Canas, Horco Molle; Tucuman, Argentina
(2645'S, 6521'W, Capllonch et al., 1997)
Didelphis albiventris
Rio Las Escobas, 3 mi WSW of Puerto de Santo
Tomas; Izabal, Guatemala (Puerto de Santo
Tomas = 1542'N, 8837'W, Geonet)
Chironectes minimus, 340-600 ft (= 100-180 m)
Rio La Torre, boca; Madre de Dios, Peru
(1251'S, 6917'W, Stephens and Traylor,
1983)
Philander opossum, 260 m (S bank Rio Tam-
bopata)
Riolima, near San Antonio, W of Cali; Valle del
Cauca, Colombia (San Antonio = 0330'N,
7638'W, Paynter, 1997)
Didelphis marsupialis, 1500-1850 m
Rio Limdn; Chuquisaca, Bolivia (1933'S,
6408'W, Anderson, 1997)
Gracilinanus agilis
Marmosops noctivagus
Micoureus constantiae
Monodelphis domestica, 1300 m
Thylamys venustus
Rio Lipeo; Tarija, Bolivia (2241'S, 6426'W,
Paynter, 1992)
Monodelphis kunsi
Rio Lujan; Buenos Aires, Argentina (3426'S,
5832'W, Paynter, 1985)
Lutreolina crassicaudata
Rio Machariapo; La Paz, Bolivia (1418'S,
6835'W, USBGN)
Didelphis marsupialis, 1000 m
Micoureus constantiae (1529'S, 6731'W)
Glironia venusta, 1000 m (Alto Rio Madidi re-
gion, ca. 1430'S, 6835'W)
Rio Madeira; Amazonas, Brazil (not precisely lo-
cated)
Marmosa murina
Rio Madeira, boca; Amazonas, Brazil (0322'S,
5845' W, Paynter and Traylor, 1991)
Philander opossum
Rio Magdalena; Magdalena, Colombia (boca =
1106'N, 7451'W, Paynter, 1997)
Marmosa robinsoni
Rio Magdalena, San Agustin (see San Agustin);
Huila, Colombia
Rio Mamore; Beni, Bolivia (1023'S, 6523'W,
Paynter, 1992)
Didelphis marsupialis (W side of Rio Mamore)
Philander opossum (also at 1226'S, 6500'W)
Rio Mamore, 2 km N, mouth Rio Chapare; Santa
Cruz, Bolivia (1557'S, 6441'W, Anderson,
1997)
Micoureus constantiae
Philander opossum
Rio Mamore, 5 km NW, mouth Rio Grande; Bo-
livia (1550'S, 64 41'W, Anderson, 1997)
Didelphis marsupialis
Rio Mamore; Rondonia, Brazil (not precisely lo-
cated)
Monodelphis brevicaudata
BROWN: ATLAS OF NEW WORLD MARSUPIALS
275
Rio Manapiarc (see San Juan de Manapiare):
Amazonas. Venezuela
Rio Mandinga. Gamboa; Panama. Panama
(0906'N, 7942'W, Geonet)
Didelphis marsupialis (2 mi W of Gamboa)
Philander opossum
Rio Manu. lower: Madre de Dios. Peru (ca.
I206'S. 71 C ()6'W. Pacheco et al.. 1993)
Metachirus nudicaudatus
Rio Man/anares: Magdalena. Colombia (1114'N.
7413'W, Paynter, 1997)
Marmosa robinsoni
Rio Mapuera: Para. Brazil (0056'S. 5702'W, Da
Silva and Langguth. 1989)
Glironia venusta
Rio Maraca: Amapa. Brazil (0026'S. 5126'W,
Paynter and Tray lor. 1991)
Philander opossum
Rio Marajano. Meseta de Jaua: Bolivar. Venezuela
(0445'N. 64 C 32'W. Perez-Hernandez. 1989)
Didelphis albiventris, 1800 m
Marmosa tyleriana. 1800 m
Rio Maranon. upper; Amazonas. Peru (near Rios
Chiriaco, Cenepa. Nieva and Santiago. Santa
Maria de Nieva = 0435'S, 77'54'W. Ste-
phens and Traylor. 1983)
Chironectes minimus
Didelphis marsupialis
Rio Maruanum. 38 km NW of Macapa: Amapa.
Brazil (001 l'N. 5116'W. Paynter and Tray-
lor. 1991))
Didelphis marsupialis
Marmosa murina
Rio Matipo (= Rio Matipoo = Matipoo): Minas
Gerais. Brazil (1953'S. 4233'W, Paynter
and Traylor. 1991)
Philander frenata
Rio Matipoo (= Rio Matipo): Minas Gerais. Bra-
zil
Rio Maule. junction of Rio Claro: Los Lagos.
Chile (3519'S. 72 = 25'W. Geonet)
Thxlamys elegans. ca. 1000 m
Rio Mawarinuma (see Cerro de La Neblina):
Amazonas. Venezuela
Rio Mazan (= Lago Miranes). boca: Loreto. Peru
(0328'S. 73 = 02'W. Hershkovitz. 1997)
Philander andersoni
Rio Mecaya. right bank Rio Caqueta: Putumavo.
Colombia (0028'N. 7520'W. Paynter. 1997)
Didelphis marsupialis, 185 m (00 C 29'N.
75 C 11'W)
Rio Memo, near Altagracia: Guarico. Venezuela
(0955'N. 6625'W, Perez-Hernandez. 1989)
Marmosa robinsoni
Rio Metlac. i km W of Fortin: Veracruz-Llave.
Mexico (Fortin = 18 = 54'N. 97 C 00'W. Geonet)
Didelphis marsupialis
Rio Mice, south side El Recreo: Zelaya. Nicara-
gua ( 121 IN. 84 C 16'W. Geonet)
Didelphis marsupialis
Rio Mindo: Pichincha. Ecuador (0003'N,
7909'W, Paynter. 1993)
Caluromys derbianus
Rio Mojinga (see Mohinga Valley): Colon. Pan-
ama
Rio Mojotoro. bridge of Ferrocarril Belgrano: Sal-
ta. Argentina (24 C 40'S. 65 C 03'W. Flores et al.,
2000 )
Thxlamys venustus
Rio Mojui (= Moju = Mojui dos Campos = Rio
Jojui): Para. Brazil (01 C 40'S. 4825'W, Payn-
ter and Traylor. 1991)
Metachirus nudicaudatus
Rio Monduapo (see Rio Munduapo): Amazonas.
Venezuela
Rio Motatan: Trujillo. Venezuela (0924'N.
70 C 36'W. Paynter. 1982)
Philander opossum. 290 m
Rib Mucujun: Merida. Venezuela (0836'N,
71 C 09'W. Paynter. 1982)
Didelphis albiventris, 9000 ft ( = 2800 m)
Rio Mucuri: Espirito Santo-Bahia-Minas Gerais.
Brazil (18 C 05'S. 3934'W, Paynter and Tray-
lor. 1991)
Marmosa murina
Micoureus demerarae (northern boundary of
Espirito Santo, near Morro de Arara. Bahia)
Rio Munduapo ( = Rio Monduapo). near San Fer-
nando de Atabapo: Amazonas. Venezuela
(San Fernanado de Atabapo = 0403'N,
67 C 28'\V. Paynter. 1982)
Marmosa murina
Rio Nam; Cordoba. Colombia (07 C 54'N. 7618'W.
Paynter. 1997)
Caluromys derbianus
Chironectes minimus, 100 m (upper Rio Sinii)
Rio Napo: Loreto. Peru (0751'S, 7617'W,
Hershkovitz. 1977)
Marmosa murina
Rio Napo. lower: Loreto. Peru (0320'S. 72 C 40'W.
Stephens and Traylor. 1983)
Didelphis marsupialis (north side)
Rib Napo. near (see Puerto Napo): Napo. Ecuador
Rio Nareuda: Pando. Bolivia (1118'S. 68 C 46'W.
USBGN)
Didelphis marsupialis
Metachirus nudicaudatus
Philander opossum
276
FIELDIANA: ZOOLOGY
"Rio Negro"; Amazonas, Brazil (not precisely lo-
cated)
Philander opossum
Rio Negro, 12.4 km WNW; Los Lagos, Chile
(4158'S, 7229'W, Geonet)
Dromiciops gliroides, 150 m
Rhyncholestes raphanurus, 150 m, 200 m
(4158'S, 7233'W)
Rio Negro, Carlo Ardabo; Amazonas, Venezuela
(0151'N, 6703'W, Perez-Hernandez, 1989)
Monodelphis brevicaudata
Rio Negro; Presidente Hayes, Paraguay (2504'S,
5755'W, Paynter, 1989)
Didelphis albiventris
Rio Negro; Rio Negro, Uruguay (3324'S,
5822'W, Paynter, 1994)
Didelphis albiventris (left bank)
Lutreolina crassicaudata
Rio Negro, 10 km above Barra del Rio Tacuarem-
bo; Tacuarembo, Uruguay (Rio Tacuarembo
= ca. 3225'S, 5529'W, Paynter, 1994)
Didelphis albiventris
Rio Novo do Itapocii (see Rio Itapocii); Santa Ca-
tarina, Brazil
Rfo Ocamo (= Rio Ocorno); Amazonas, Vene-
zuela (0248'N, 6514'W, Paynter, 1982)
Didelphis marsupialis (boca Rio Orinoco)
Marmosa murina (upper Rio Orinoco)
Micoureus demerarae
Monodelphis brevicaudata
Rio Ocoa (= Ocoa River), Villavicencio- Acacias
Road; Meta, Colombia (0408'N, 7315'W,
Paynter, 1997)
Caluromys lanatus
Chironectes minimus
Rio Ocobamba (= Occobamba Valley = Ocobam-
ba Valley); Cusco, Peru (1226'S, 7223'W,
Stephens and Traylor, 1983)
Didelphis albiventris
Lestoros inca, 9100 ft (= 2840 m) (1253'S,
7221'W)
Monodelphis adusta (ca. 1250'S, 7226'W)
Rio Ocorno (see Rio Ocamo); Amazonas, Vene-
zuela
Rfo Ocumare; Aragua, Venezuela (1029'N,
6745'W, Geonet)
Chironectes minimus
Rfo Orinoco, upper; Amazonas, Venezuela (not
precisely located)
Micoureus demerarae
Rfo Oritupano; Anzoategui, Venezuela (0913'N,
6350'W, Lopez-Fuster et al., 2000)
Marmosa robinsoni
Rfo Oscuro, near Cali; Valle del Cauca, Colombia
(Cali = 0327'N, 7631'W, Paynter, 1997)
Caluromys derbianus, 1000 m
Rio Otlati; Puebla, Mexico (1922'N, 9831'W,
Geonet)
Didelphis marsupialis, 8700 ft (= 2700 m)
Rfo Ovejera, San Agustin; Huila, Colombia
(0157'N, 7629'W, Paynter, 1997)
Caenolestes fuliginosus, 2350 m
Didelphis albiventris, 2350 m (0155'N,
7630'W)
Rfo Pachitea, tributary of Rfo Huanuco; Huanuco,
Peru (0846'S, 7432'W, Stephens and Tray-
lor, 1983)
Chironectes minimus
Rfo Pacuare; Limdn, Costa Rica (1014'N,
8317'W, Geonet)
Marmosa mexicana
Rfo Palometillas; Santa Cruz, Bolivia (1736'S,
6337'W, Hershkovitz, 1992)
Gracilinanus agilis
Rio Papagaio (= Rio Saueruina), right bank Rio
Juruena; Mato Grosso, Brazil (1200'S,
5843'W, Paynter and Traylor, 1991)
Caluromys philander
Rio Paracaf; Parana, Brazil (2341'S, 5357'W,
Paynter and Traylor, 1991)
Philander frenata
Rfo Parana; Alto Parana, Paraguay (not precisely
located)
Thylamys pusilla
Rfo Parana-Guazu, 1 5 km from banks of Rfo Pa-
rana; Misiones, Argentina (2641'S, 5448'W,
Paynter, 1985)
Chironectes minimus
Rfo Paranapura; Loreto, Peru (0554'S, 7506'W,
Stephens and Traylor, 1983)
Philander andersoni
Rio Pardo, left side upper Rio Parana; Mato Gros-
so do Sul, Brazil (2146'S, 5209'W, Paynter
and Traylor, 1991)
Chironectes minimus (left side upper Rio Pan-
rana)
Rio Parnahyba, lower; Piauf, Brazil (not precisely
located)
Didelphis albiventris
Rfo Pasaje Talavera; Buenos Aires, Argentina
(3353'S, 5855'W, USBGN)
Lutreolina crassicaudata
Rfo Perene; Junfn, Peru (1109'S, 7418'W,
USBGN)
Micoureus regina
Rfo Periquito; Aragua, Venezuela (ca. 1022'N,
6742'W, Geonet)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
277
Chironectes minimus
Rfo Pescado: Manabf, Ecuador (0125'S,
8015'W, Hershkovitz, 1997)
Metachirus nudicaudatus, 16(H) ft (= 500 m)
Philander opossum, 500 m
Rio Pilcomayo. boca; Central, Paraguay (2521'S.
5742'W, Paynter, 1989)
Lutreolina crassicaudata
Rio Pindo; El Oro. Ecuador (0350'S, 7945'W,
Pay men 1993)
Didelphis marsupialis
Rfo Pindo Yacu; Pastaza, Ecuador (0208'S.
7603'W. Hershkovitz, 1997)
Caluromys lanatus
Metachirus nudicaudatus, 250 m
Philander andersoni, 250 m
Rfo Piratucu, boca. 80 km N of Manaus; Ama-
zonas, Brazil (Manaus = 0308'S, 60 o 01'W)
Ma rmosops pa n idens
Rfo Pirn's (see San Geronimo); Puntarenas, Costa
Rica
Rfo Pirrita (see San Geronimo): Puntarenas. Costa
Rica
Rfo Pita: Pichincha. Ecuador (0018'S. 7828'W.
Paynter. 1993)
Caenolestes fuliginosus
Rfo Pita, upper; Pichincha. Ecuador (0020'S.
7825'W. Bublitz. 1963)
Caenolestes fuliginosus
Rfo Pitisama [Rfo Pitasama] (also see Cerro Am-
boro), Parque Nacional de Amboro: Santa
Cruz. Bolivia (1742'S. 6337'W, Anderson
et al., 1997)
Chironectes minimus, 500 m
Rfo Pito: San Bias. Panama (0842'N. 7732'W.
Geonet)
Chironectes minimus
Rfo Platano Biosphere Reserve: Gracias a Dios.
Honduras (1553'N, 8641'W. Geonet)
Philander opossum
Rfo Pucro, head, Cerro Mali: Darien. Panama
(0759'N, 7733'W, Wenzel and Tipton.
1966)
Caluromys derbianus
Rfo Puyacatengo, 15 km E of Teapa; Tabasco.
Mexico (1744'N. 9255'W, Geonet)
Didelphis marsupialis
Rfo Quininde (see Rosa Zarate); Esmeraldas, Ec-
uador
Rfo Quiser (see Rfo Quizar); Santa Cruz. Bolivia
Rfo Quizar (= Rfo Quiser), Nuflo de Chavez;
Santa Cruz. Bolivia (1637'S. 6246'W. Payn-
ter, 1992)
Monodelph is domestica
Rfo Ramos. 20 km N of Montemorelos; Nuevo
Leon. Mexico (2531'N, 9952'W, Geonet)
Didelphis marsupialis, 1000 ft (= 300 m)
Rfo Ranquil; Araucania, Chile (3816'S, 7118'W,
Geonet)
Dromiciops gliroides, 1370 m
Rfo Raposo: Valle del Cauca, Colombia (0343'N,
7708'W. Hershkovitz, 1997)
Didelphis marsupialis
Philander opossum, near sea level (Pacific Vi-
rology Field Camp)
Rfo Reventazon, 5 km SE of Turrialba; Cartago,
Costa Rica (Turrialba = 0954'N, 8341'W,
Geonet)
Chironectes minimus, 1800 ft (= 560 m)
Rio Rodeio, near Rio de Janeiro, Serra do Mar;
Rio de Janeiro. Brazil (Rodeio = 2233'S,
4341'W. Hershkovitz, 1997)
Didelphis aurita
Rfo Sabinas. 10 km N of El Encino (see El En-
cino); Tamaulipas, Mexico
Rfo Saija, boca (= La Boca Rfo Saija); Cauca,
Colombia (0252'N, 7741'W, Paynter, 1997)
Chironectes minimus (0252'N, 7743'W)
Didelphis marsupialis
Philander opossum, near sea level
Rio Saloya; Pichincha, Ecuador (0001'N,
7857'W, Paynter, 1993)
Caenolestes convelatus
Rfo Samiria (= Samiria) (also see Yanayaquillo);
Loreto. Peru (0442'S, 7413'W, Stephens
and Traylor. 1983)
Caluromys lanatus
Didelphis albiventris
Didelphis marsupialis
Rfo Sandd. Rio Baudd; Chocd, Colombia
(0503'N. 7657'W, Paynter, 1997)
Metachirus nudicaudatus, 160 m
Philander opossum, 1 60 m
Rfo San Francisco. Altura de Yuto; Jujuy, Argen-
tina (2339'S. 6427'W, Flores et al., 2000)
Micoureus constantiae, 500 m
Rfo San Miguel; San Miguel, El Salvador
(1314'N, 88 C 22'W, Geonet)
Philander opossum, 225 ft (= 70 m)
Rfo San Rafael, near Santa Rosa; Pichincha, Ec-
uador (0020'S, 7827'W. Bublitz, 1987)
Caenolestes fuliginosus, 9000 ft (= 2800 m)
Rfo Santa Marfa; Salta, Argentina (2317'S,
6414'W. Paynter. 1985)
Thyalmys venustus
Rio Sao Domingos; Goias, Brazil (1324'S,
4712'W, Paynter and Traylor, 1991)
Monodelphis domestica
278
FIELDIANA: ZOOLOGY
Rio Sao Jose; Espfrito Santo, Brazil (2000'S,
4040'W, Mustrangi and Patton, 1997)
Marmosops incanus
Rib Sara, near Buena Vista; Santa Cruz, Bolivia
(Buena Vista = 1727'S, 6340'W, Paynter,
1996)
Micoureus constantiae
Rio Saueruina (see Rio Papagaio); Mato Grosso,
Brazil
Rios Chimbo-Coco; Guayas, Ecuador (0210'S,
7950'W, Hershkovitz, 1997)
Philander opossum, 2400 ft (= 750 m)
Rio Seco, near Cordoba; Veracruz-Llave, Mexico
(Cordoba = 1853'N, 9656'W, Geonet)
Philander opossum
Rio Segovia (see Rio Coco); border between Nic-
aragua and Honduras
Riosinho, left bank Rio Fresco; Para, Brazil (ca.
0707'S, 5139'W, Geonet and Wenzel and
Tipton, 1966)
Monodelphis emiliae
Rio Siquia; Zelaya, Nicaragua (1245'N, 8501 "W,
Geonet)
Philander opossum
Rio Socopito (= Rio Socopo); Falcon, Venezuela
(1030'N, 7044'W, Handley, 1976)
Didelphis marsupialis, 480 m
Marmosa robinsoni, 470 m
Rio Socopo (see Rio Socopito); Falcon, Venezuela
Rio Solimoes, lower; Amazonas, Brazil (not pre-
cisely located)
Didelphis marsupialis
Rib Solocame; La Paz, Bolivia (1616'S,
6724'W, Paynter, 1992)
Marmosops dorothea
Rio Suapure (Caura Valley); Bolivar, Venezuela
(0648'N, 6701'W, Paynter, 1992)
Marmosa murina
Rio Suno, Rio Napo; Napo, Ecuador (0042'S,
7708'W, USBGN)
Micoureus regina
Rio Supamo; Bolivar, Venezuela (0648'N,
6150'W, Geonet)
Didelphis marsupialis, 150 m
Marmosa murina, 350 m
Monodelphis brevicaudata, 150 m
Rib Surutu; Santa Cruz, Bolivia (1724'S,
6351'W, Paynter, 1992)
Metachirus nudicaudatus
Micoureus constantiae (W tributary and 25 km
E of Buenavista)
Rio Taquari, valley of (see Taquari River Valley);
Rio Grande do Sul, Brazil
Rio Tarra, tributary of Rib Catatumbo; Norte de
Santander, Colombia (0839'N, 7301'W,
Paynter, 1992)
Didelphis marsupialis, 200 m
Metachirus nudicaudatus, 200 m (Rio Catatum-
bo)
Rio Tavara, Rib Inambari; Puno, Peru (ca.
1322'S, 6936'W, Stephens and Traylor,
1983)
Micoureus regina
Rio Teapa; Tabasco, Mexico (1758'N, 9254'W,
Geonet)
Chironectes minimus
Rfo Termales, Manizales; Caldas, Colombia
(0458'N, 7523'W, Bublitz, 1987)
Caenolestes fuliginosus, 2600-3300 m
Rio Tesorero; Jujuy, Argentina (2359'S, 6518'W,
Flores et al., 2000)
Thylamys venustus
Rio Tijamuchi, 5 km from mouth; Beni, Bolivia
(1410'S, 6458'W, Anderson, 1997)
Didelphis marsupialis
Philander opossum (1456'S, 6509'W)
Rio Tiraxi; Jujuy, Argentina (2359'S, 6519'W,
Flores et al., 2000)
Thylamys venustus
Rio Tocantins, lower; Para, Brazil (not precisely
located)
Caluromys philander
Rio Tocuyo; Lara, Venezuela (head of Rio =
1103'N, 6820'W, El Tocuyo at head of Rio
Tocuyo = 0947'N, 6820'W, Paynter, 1982)
Marmosa robinsoni
Rio Tuma, E of Matagalpa; Zelaya, Nicaragua
(1303'N, 8444'W, Geonet)
Marmosa mexicana, below 1000 ft (= 300 m)
Rio Tuy, 10 km N of El Consejo; Aragua, Vene-
zuela (El Consejo = 1014'N, 6716'W,
Paynter, 1982)
Chironectes minimus
Rfo Umulla; Honduras (not located)
Marmosa mexicana
Rio Urubamba, boca; Ucayali, Peru (1042'S,
7342'W, Hershkovitz, 1997)
Metachirus nudicaudatus, 250 m
Philander opossum
Rio Urucu (see Alto Rio Urucu); Amazonas, Bra-
zil
Rfo Urugua-I, 30 km of Puerto Bemberg, tributary
of Rfo Parana; Misiones, Argentina (2554'S,
5436'W, Geonet)
Caluromys lanatus
Chironectes minimus
Didelphis albiventris
Didelphis aurita
BROWN: ATLAS OF NEW WORLD MARSUPIALS
279
Gracilinanus agilis
Metachirus nudicaudatus
Philander opossum
Rio Utcubamba, 15 mi above Chachapoyas; Ama-
zonas, Peru (0530'S, 7832'W, Stephens and
Tray lor, 1983)
Didelphis albiventris, 2290 m
Rio Verde; Esmeraldas, Ecuador (0104'N,
7925'W, Paynter, 1993)
Didelphis marsupialis, sea level-300 m
Rio Victoria; Misiones, Argentina (not located)
Monodelphis iheringi
Rio Viejo (see Cornes); Veraguas, Panama
Rio Vilcanota; Cusco, Peru (1319'S, 7159'W,
Geonet)
Micoureus regina
Rio Xingu, E bank, 52 km SSW of Alatamira;
Para, Brazil (0339'S, 5222'W, Patton and
Da Silva, 1997)
Philander opossum
Rio Xiriviny; Amazonas, Peru (0059'S, 6153'W,
Hershkovitz, 1997)
Philander opossum, 260 ft (= 80 m)
Rfo Yana Rumi; Pastaza, Ecuador (0138'S,
7659'W, Hershkovitz, 1997)
Metachirus nudicaudatus
Philander andersoni
Rio Yapacani; Santa Cruz, Bolivia (1724'S,
6346'W, Anderson et al., 1993)
Didelphis marsupialis
Rio Yarapa; Loreto, Peru (0430'S, 7325'W,
Fleck and Harder, 1995)
Philander mcilhennyi
Rio Yavari; Loreto, Peru (0421'S, 7002'W, Ste-
phens and Traylor, 1983)
Marmosa murina
Rio Ypanema (see Ipanema); Sao Paulo, Brazil
Rfo Yuruan; Bolivar, Venezuela (0648'N,
6150'W, Paynter, 1982)
Philander opossum
Rib Zulia; Norte de Santander, Colombia
(0904'N, 7218'W, Paynter, 1997)
Caluromys lanatus
Risa de Oro, 3 km E (= El Risa de Oro); Chiapas,
Mexico (1620'N, 9403'W, Gardner, 1973)
Didelphis marsupialis
Rivadavia; Mendoza, Argentina (3313'S,
6821'W, Paynter, 1985)
Didelphis albiventris, sea level
Rivadavia (see El Breal); Salta, Argentina
Rivas (also see Lago Nicaragua); Rivas, Nicara-
gua (1 126'N, 8550'W, Geonet)
Caluromys derhianus
Rive (see Engenheiro Reeve); Espirito Santo, Bra-
zil
Riviere Arataye (see Arataye Crique); Inini,
French Guiana
Roatan (= Ruatan) Island, one of islands of the
Islas de la Bahi'a; Honduras (1618'N,
8633'W, Geonet)
Marmosa robinsoni
Robles; Santiago del Estero, Argentina (2755'S,
6408'W. Flores et al., 2000)
Thylamys pusilla
Roca Nova, Serra do Mar; Parana, Brazil
(2530'S, 4850'W, Hershkovitz, 1997)
Didelphis aurita, 1000 m
Marmosops paulensis, 1000 m
Philander frenata, 930-1150 m
Rocas Novas (see Caete); Minas Gerais, Brazil
Rocha, ca. 40 km NNW of Juquia; Sao Paulo,
Brazil (2453'S, 4705'W, Hershkovitz files)
Chironectes minimus, ca. 50 m
Didelphis aurita
Metachirus nudicaudatus (also possibly at La-
goa do Rocha = 2153'S, 4705'W)
Rodeio, Serra do Mar; Rio de Janeiro, Brazil
(2233'S, 4341'W, Hershkovitz, 1997)
Philander frenata, 1400 ft (= 430 m)
Rodeo, 7.5 km SE; Cochabamba, Bolivia
(1741'S, 6536'W, Anderson, 1997)
Didelphis albiventris
Thylamys pallidior
Thylamys venustus, 3800 m (1740'S, 6535'W)
Rodman Naval Ammnition Depot; Panama, Pan-
ama (0857'N, 7935'W, Geonet)
Marmosa robinsoni
Metachirus nudicaudatus (0857'N, 7937'W)
Philander opossum
Rosarinho (= Rozarinho); Amazonas, Brazil
(0342'S, 5908'W, Paynter and Traylor,
1991)
Didelphis marsupialis
Marmosa murina (Lago Miguel, Rio Madeira)
Marmosops noctivagus (Lago Miguel, W bank
Rio Madiera)
Philander opossum
Rosario; Sinaloa, Mexico (2300'N, 10552'W,
Geonet)
Marmosa canescens
Rosario de Lerma; Salta, Argentina (2459'S,
6535'W, Paynter, 1985)
Didelphis albiventris, ca. 1000 m
Rosa Zarate (= Quininde = Rfo Quininde); Es-
meraldas, Ecuador (0020'N, 7928'W, Payn-
ter, 1993)
Chironectes minimus
280
FIELDIANA: ZOOLOGY
Didelphis marsupialis, ca. 100 m
Metachirus nudicaudatus
Philander opossum
Rozarinho (see Rosarinho); Amazonas, Brazil
Ruatan (see Roatan Island); Honduras
Runcaraccay Ruins, Machu Pichu (see Machu Pi-
chu, Urubamba, Runcaraccay Ruins); Cusco,
Peru
Rupununi River; Upper Takutu-Upper Essequibo,
Guyana (0403'N, 5835'W, Geonet)
Philander opossum, 200 ft (= 60 m)
Rurrenabaque; Beni, Bolivia (1428'S, 6734'W,
Anderson, 1997)
Didelphis marsupialis
Philander opossum
Ruta 188, Curva de Pena (see Pergamino); Bue-
nos Aires, Argentina
Sabana Grande; Managua, Nicaragua (1207'N,
8610'W, Geonet)
Caluromys derbianus
Philander opossum, 40-50 m
Sabana Grande, Cerro Duida; Amazonas, Vene-
zuela (ca. 0325'N, 6540'W, Paynter, 1982)
Didelphis marsupialis, 100 m
Sabanas de Vichada y Meta; Meta, Colombia (not
located)
Lutreolina crassicaudata
Sabaneta; Distrito Federal, Venezuela (1028'N,
6705'W, Perez-Hernandez, 1989:381)
Gracilinanus marica
Marmosops fuscatus
Sabanetas; Cauca, Colombia (0232'N, 7653'W,
Paynter, 1997)
Chironectes minimus, 2000 m
Didelphis marsupialis, 1800 m, 1900 m, 2000 m
Sabanilla, Rio Zamora, near mouth Rio Sabanilla;
Zamora-Chinchipe, Ecuador (0402'S,
7901'W, Geonet)
Didelphis albiventris, 5700 ft (= 1780 m)
Sahara; Minas Gerais, Brazil (1954'S, 4351'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 705 m
Sabinopolis; Minas Gerais, Brazil (1840'S,
4306'W, USBGN)
Didelphis aurita
Saboga Island (see Isla Saboga); Panama, Panama
Sacado, right bank Rio Jurua; Amazonas, Brazil
(0645'N, 7051'W, Patton et al., 2000)
Caluromys lanatus
Micoureus regina
Philander opossum
Saimiria (see Rio Samiria); Loreto, Peru
Saint Georges; French Guiana (0345'N, 5205'W,
Stephens and Traylor, 1985)
Caluromys philander
Saint-Ignace-Gouazou (see San Ignacio); Mi-
siones, Paraguay
Sajama, 9 km N, 4 km E; Oruro, Bolivia (1803'S,
6856'W, Anderson, 1997)
Thylamys pallidior
Sajama Nevado (= Cerro Sajama = Mount Sa-
jama); Oruro, Bolivia (1807'S, 6900'W,
Paynter, 1992)
Gracilinanus agilis, 4350 m
Salache, ca. 6 mi S of Cipres; Suchetepequez,
Guatemala (Cipres = 1433'N, 9028'W,
Geonet)
Didelphis marsupialis
Salar Cuchari, on road 70; Jujuy, Argentina
(2350'S, 6647'W, Flores et al., 2000)
Thylamys pallidior
Salento; Quindio, Colombia (0438'N, 7534'W,
Paynter, 1997)
Didelphis marsupialis, 1895 m
Marmosops impavidus (near Quindio)
Salesopolis; Sao Paulo, Brazil (2332'S, 4551'W,
Hershkovitz, 1997)
Philander frenata
Salinas; El Oro, Ecuador (0323'S, 8019'W, Geo-
net)
Didelphis marsupialis
Salinas; Minas Gerais, Brazil (1612'S, 4215'W,
Varejao and Valle, 1982)
Didelphis albiventris
Salinas Diamante R R Station (see Salinas del Di-
amante); Mendoza, Argentina
Salinas del Diamante; Mendoza, Argentina
(3456'S, 6852'W, Paynter, 1985)
Thylamys pallidior
Salinas del Bebedero; San Luis, Argentina
(3332'S, 6639'W, Geonet)
Thylamys pallidior, 1000 ft (= 300 m), 1350 ft
(= 420 m)
Salobra; Mato Grosso do Sul, Brazil (2114'S,
5708'W, Hershkovitz, 1997)
Philander frenata (gallery forest)
Saloya Occidente (see Galaya); Pichincha, Ecua-
dor
Salta, 12 km NW; Salta, Argentina (2447'S,
6525'W, Paynter, 1985)
Thylamys venustus
Salt Creek, Isla Bastimentos; Bocas del Toro, Pan-
ama (Isla Bastimentos = 0918'N, 8208'W,
Geonet)
Caluromys derbianus
BROWN: ATLAS OF NEW WORLD MARSUPIALS
281
Salto da Felicidade, Rio Sepotuba; Mato Grosso.
Brazil (1441'S, 574.VW, USBGN)
Caluromys lanatus
Salto de Pirareta, 10 km S of Piribebuy; La Cor-
dillera, Paraguay (Piribebuy = 2529'S,
5703'W. Paynter, 1989)
Chironectes minimus
Salto do Sepotuba, Rio Sepotuba; Mato Grosso.
Brazil (1555'S. 5737'W, Hershkovitz, 1992)
Gracilinanus agilis
Salto Ichum, Rio Paragua; Bolivar, Venezuela
(0450'N, 6320'W, Perez-Hernandez, 1989)
Didelphis albiventris
Saltos de Pirareta; Paraguari, Paraguay (not lo-
cated)
Didelphis albiventris
Salto Utiarity; Mato Grosso, Brazil (1302'S,
5817'W, Paynter and Traylor, 1991)
Caluromys philander
Salud; Colon, Panama (0912'N. 8008'W. Geo-
net)
Philander opossum
Salvador (see Bahia); Bahia, Brazil
Salvias; El Oro, Ecuador (0347'S, 7921'W,
Paynter, 1993)
Marmosops impavidus, 1050 m
Philander opossum, 3500 ft (= 1090 m)
Samana; Caldas, Colombia (ca. 0515'N,
7515'W, Paynter, 1987)
Marmosops parvidens, 1 200 m
Samiria (see Rio Samiria); Loreto, Peru
San Adolfo; Huila, Colombia (0137'N, 7559'W.
Paynter, 1987)
Didelphis marsupialis, 1400 m
San Agustfn; Monagas, Venezuela (1012'N,
6332'W, Geonet)
Caluromys philander, 1 170-1 180 m
Chironectes minimus, 1 150 m, 1 160-1335 m
Didelphis marsupialis, 1160-1335 m
Marmosa murina, 1260-1365 m
Marmosa robinsoni, 854-1690 m
Marmosops fuscatus, 1 1 50- 1339 m
San Agustfn, Aldea Puerto Golpe; El Progresso,
Guatemala (1457'N, 8958'W, Geonet)
Philander opossum
San Agustfn. Rfo Magdalena; Huila, Colombia
(0153'N, 7616'W. Paynter. 1997)
Caenolestes fuliginosus, 2300 m
Didelphis marsupialis, 1 690 m
San Alouatta [sic]; Suriname (not located)
Marmosa lepida
San Andres; Miranda. Venezuela (1022'N,
6650'W, Handley, 1976)
Didelphis marsupialis, 1 140-1 144 m
Marmosa robinsoni, 950-1144 m
San Andres Tuxtla; Veracruz-Llave, Mexico
(1827'N, 9513'W, Geonet)
Caluromys derbianus
Didelphis marsupialis, 1000 ft (= 300 m)
Marmosa mexicana
Philander opossum, 1000 ft (= 300 m)
San Antonio; Chimborazo, Ecuador (0217'S,
7859'W, Gardner. 1993)
Didelphis albiventris
San Antonio; Chinandega, Nicaragua (1232'N,
8703'W, Geonet)
Didelphis marsupialis, 35 m
San Antonio; Misiones, Argentina-Parana, Brazil
(2607'S, 5345'W, Paynter, 1985)
Didelphis albiventris
San Antonio; Nueva Esparta. Venezuela (1057'N,
6355'W, Paynter, 1982)
Marmosa robinsoni, ca. 50 m
San Antonio; Pichincha, Ecuador (0001'N,
7827'W, Paynter, 1997)
Caenolestes caniventer
San Antonio; San Martin, Peru (0715'S,
7644'W, Stephens and Traylor, 1983)
Micoureus regina (SE of Moyobamba)
San Antonio; Santiago del Estero, Argentina
(2646'S, 6437'W, Flores et al., 2000)
Thylamys pusilla
San Antonio; Tungurahua, Ecuador (0109'S,
7828'W, Paynter, 1993)
Caenolestes caniventer
San Antonio; Valle del Cauca, Colombia
(0330'N, 7638'W, Paynter, 1997)
Didelphis albiventris, 6500 ft (= 2030 m)
Mannosops impavidus, 7000 ft (= 2180 m)
San Antonio, 1 hr N of Tambo; Canar, Ecuador
(0229'S, 7857'W, Geonet)
Didelphis albiventris, ca. 1225 m, 11000 ft ( =
3400 m)
San Antonio, Rfo Ulva [Ulba]; Napo, Ecuador
(Hacienda at San Antonio = OTIO'S,
7828'W, Hershkovitz files)
Marmosops impavidus
San Antonio, San Agustfn; Huila, Colombia
(0157'N, 7629'W, Paynter, 1997)
Caenolestes fuliginosus, 2200-2300 m
Didelphis albiventris, 2200 m, 2350 m
San Antonio. 10 km E, near Camiaco-Limoqueje;
Beni, Bolivia (1507'S, 6450'W, Geonet)
Didelphis marsupialis
San Antonio de Maturfn; Monagas, Venezuela
(1007'N. 6343'W, Paynter, 1982)
Marmosa robinsoni, ca. 500 m
282
FIELDIANA: ZOOLOGY
San Bartolo; Loja, Ecuador (ca. 0402'S, 7955'W,
Paynter, 1993)
Didelphis marsupialis, 2300 m
San Bias, 3.5 mi E; Sinoloa, Mexico (2605'N,
10846'W, Geonet)
Marmosa canescens
San Calixto, Rio Tarra; Norte de Santander, Co-
lombia (0824'N, 7512'W, Geonet)
Chironectes minimus
Metachirus nudicaudatus
San Calixto, Valle de Suaza; Huila, Colombia
(Rio Suaza = 0210'N, 7512'W, Geonet)
Chironectes minimus
Didelphis marsupialis
San Carlos; Alajuela, Costa Rica (1037'N,
8410'W, Geonet)
Chironectes minimus
Philander opossum (1035'N, 8410'W)
San Carlos; Mendoza, Argentina (3346'S,
6902'W, Paynter, 1985)
Didelphis albiventris, 943 m
San Carlos de Rio Negro; Amazonas, Venezuela
(0155'N, 6704'W, Paynter, 1982)
Caluromys lanatus, 120 m
Monodelphis brevicaudata, 120 m
San Cristobal (suburb of Bogota); Cundinamarca,
Colombia (0434'N, 7405'W, Paynter, 1997)
Caenolestes fuliginosus, 2800-2900 m
Didelphis albiventris, 2600 m (0436'N,
7405'W)
San Cristobal, Verapaz; Alto Verapaz, Guatemala
(1523'N, 9024'W, Geonet)
Marmosa mexicana
San Cristobal, 41 km SW, near Paramo de Tama;
Tachira, Venezuela (0746'N, 7214'W, Payn-
ter, 1982)
Caenolestes fuliginosus
Sanderije (see Zanderij); Para, Suriname
San Emilio, Lago Nicaragua; Rivas, Nicaragua
(Lago Nicaragua = 1 131'N, 8552'W, Geo-
net)
Marmosa mexicana
San Esteban; Carabobo, Venezuela (1026'N,
6801'W, Paynter, 1982)
Marmosa robinsoni, ca. 200 m
Micoureus demerarae, ca. 200 m
Micoureus regina, ca. 200 m
Monodelphis palliolata, ca. 200 m
San Eusebio, Carbonera; Merida, Venezuela
(0839'N, 7123'W, Perez-Hernandez, 1989)
Marmosops fuscatus, 2200 m
San Felipe; Yaracuy, Venezuela (1020'N,
6844'W, Paynter, 1982)
Marmosa robinsoni
San Felipe ( = Raudalito), Rio Sipapo; Amazonas,
Venezuela (not precisely located; Rio Sipapu
= 0503'N, 6748'W, Paynter, 1982)
Metachirus nudicaudatus
San Felipe Orizatlan, 4 km E; Hidalgo, Mexico
(2111'N, 9837'W, Gardner, 1973)
Didelphis marsupialis
San Felix (see Ciudad Guayana); Bolivar, Vene-
zuela
San Fernando, left bank Rio Yavari; Loreto, Peru
(0412'S, 7014'W, Stephens and Traylor,
1983)
Metachirus nudicaudatus, 100 m
San Fernando; Tucuman, Argentina (2622'S,
6527'W, Mares et al., 1966)
Didelphis albiventris
San Francisco de C; Minas Gerais, Brazil (not lo-
cated)
Philander opossum
San Francisco del Monte de Oro; San Luis, Ar-
gentina (3236'S, 6608'W, Paynter, 1985)
Didelphis albiventris, 1480 ft (= 460 m)
Thylamys pallidior, 2800 ft ( 875 m) (Que-
brada de Lopez)
San Francisco; Tungurahua, Ecuador (0118'S,
7830'W, Paynter, 1993)
Caenolestes caniventer
San Geronimo, Cerro San Miguel; Amazonas, Ve-
nezuela (not located)
Didelphis marsupialis, 960 m
San Geronimo (= San Jeronimo), Pirns (= Rio
Pirris = Rio Parrita = Parrita); Puntarenas,
Costa Rica (1002'N, 8438'W, Geonet)
Didelphis marsupialis
San Gil; Santander, Colombia (0633'N, 7308'W,
Paynter, 1997)
Caluromys lanatus, 1150 m
San Gregorio; Artigas, Uruguay (3033'S,
5752'W, Paynter, 1994)
Lutreolina crassicaudata, ca. 100 m
Sangre Grande, St. Andrew, Rio Grande Forest;
St. Andrew, Trinidad and Tobago (1035'N,
6107'W, Geonet)
Marmosa robinsoni
Marmosops fuscatus
Sangre Grande River; St. Andrew, Trinidad and
Tobago (1035'N, 6105'W, Geonet)
Didelphis marsupialis
San Ignacio; San Jose, Costa Rica (0948'N,
8410'W, Geonet)
Caluromys derbianus
Philander opossum
San Ignacio (see San Ignacio de Velasco); Santa
Cruz, Bolivia
BROWN: ATLAS OF NEW WORLD MARSUPIALS
283
San Ignacio, Cerro [Mt.j Pichincha: Pichincha.
Ecuador (0012'S, 7833'W, Geonet)
Caenolestes fuliginosus. 1 1 150 ft (= 3500 m)
San Ignacio, 20 km W (= Saint-Ignace-Gouazou);
Misiones, Paraguay (2652'S, 5703'W, Payn-
ter, 1989)
Didelphis albiventris
Thylamys pusilla
San Ignacio de Moxos; Beni, Bolivia (1453'S.
6536'W, Paynter. 1992)
Didelphis marsupialis
Philander opossum
San Ignacio de Velasco (= San Ignacio); Santa
Cruz, Bolivia (1622'S, 6057'W, Geonet)
Didelphis albiventris, ca. 300 m (also 1737'S.
6053'W, Paynter, 1992)
Monodelphis domestica
San Ignacio de Yuran (= San Ignacio de Yuruani).
Rio Yuruan; Bolivar, Venezuela (Rio Yuruan
= 0648'N, 6150'W, Paynter, 1982)
Didelphis albiventris
Lutreolina crassicaudata, 900-940 m
San Ignacio de Yuruani (see San Ignacio de Yu-
ran); Bolivar, Venezuela
San Isidro, Valle de Suaza; Huila. Colombia
(0310'N, 7455'W, Geonet)
Caluromys lanatus
Didelphis marsupialis
San Isidro del General (= Urena); San Jose. Costa
Rica (0923'N, 8342'W, Geonet)
Didelphis marsupialis
San Jacinto, Rio Tigre; Loreto. Peru (0218'S,
7551'W, Patton and da Silva, 1997)
Philander andersoni
San Jacinto, Rib Pastaza; Loreto, Peru (ca.
0300'S, 7600'W, Patton et al.. 2000)
Philander andersoni
San Jacinto; Merida, Venezuela (ca. 0836'N.
7105'W, Paynter, 1982)
Didelphis marsupialis, 1450 m
San Javier, 23 km W; Beni. Bolivia (1434'S.
6455'W, Anderson, 1997)
Lutreolina crassicaudata
San Javier; Esmeraldas. Ecuador (0104'N.
7847'W, Paynter. 1993)
Caluromys derbianus (Rio Cachabi [Cachavf])
Didelphis marsupialis, ca. 100 m (Rio Cachabi
[Cachavf])
Marmosa robinsoni, 100 m (Rio Cachabi [Ca-
chavf))
Metachirus nudicaudatus, 60 ft (= 18 m) (Rio
Cachabi [Cachavf])
Philander opossum, 20 m
San Javier to El Puente. 6 mi E on road; Santa
Cruz. Bolivia (1620'S, 6242'W, Tarifa and
Anderson. 1997)
Glironia vemista, 300 m
San Javier. 12 km S; Santa Cruz. Bolivia
(1627'S, 6239'W, Anderson, 1997)
Glironia venusta
San Javier: Tucuman, Argentina (2647'S,
6523'W. Paynter. 1985)
Thylamys venustus, 1000 m
San Javier, right bank Rio Paraguay; Misiones,
Argentina (2753'S, 5508'W, Paynter, 1985)
Chironectes minimus, ca. 50 m
San Jeronimo; Baja Verapaz. Guatemala
(1503'N. 9212'W, Geonet)
Marmosa mexicana, 2950 ft (= 920 m)
San Jeronimo; Cusco. Peru (1334'S. 7145'W,
Stephens and Traylor, 1983)
Philander opossum, 500 ft (= 150 m)
San Jeronimo (see San Geronimo); Puntarenas.
Costa Rica
San Jeronimo. Rfo Ucayali; Loreto, Peru
(0750'S, 7445'W, Geonet)
Marmosops impavidus, 300 m
Micoureus regina, 1000 ft (= 300 m)
Philander andersoni, 300 m
Philander opossum, 500 ft (= 150 m)
San Joaquin (= Camino Vilches = Estancia Bar-
ranquita); Beni. Bolivia (1304'S, 6449'W,
Paynter. 1992)
Caluromys lanatus, 200 m
Didelphis marsupialis, 200 m, 400 m (1 mi E
of San Joaquin)
Gracilinanus agilis, 200 m
Lutreolina crassicaudata, ca. 200 m
Monodelphis domestica, 200 m
Philander opossum, ca. 200 m (W bank Rfo
Machupo. 1 mi E of San Joaqufn)
San Jorge. Rfo Negro, 15 km NNW; Durazno,
Uruguay (3252'S, 5555'W, USBGN)
Didelphis albiventris
San Jose: Caldas. Colombia (0505'N, 7548'W,
Geonet)
Didelphis marsupialis
San Jose, coast belt; Cauca. Colombia (0351'N,
7652'W, Paynter, 1997)
Marmosa robinsoni
San Jose: Manabf. Ecuador (0009'S, 8004'W,
Baker. 1974)
Chironectes minimus, 50 m
San Jose (see San Jose Nuevo); Napo, Ecuador
San Jose. Rfo Pilcomayo; Formosa, Argentina (ca.
2515'S. 5815'W. Paynter, 1997)
Lutreolina crassicaudata, ca. 100 m
San Jose. Rfo Santa Rosa; Ayacucho, Peru (ca.
284
FIELDIANA: ZOOLOGY
1244'S, 7346'W, Stephens and Traylor,
1983)
Metachirus nudicaudatus, 3300 ft (= 1030 m)
Philander andersoni, 1000 m
San Jose (= San Jose de Chiquitos), Rio San Mi-
guel; Santa Cruz, Bolivia (1751'S, 6047'W,
Paynter, 1992)
Didelphis albiventris, 297 m
Lutreolina crassicaudata, ca. 300 m
Metachirus nudicaudatus
Micoureus constantiae
San Jose; San Jose, Costa Rica (0956'N,
8405'W, Geonet)
Chironectes minimus
Didelphis marsupialis
Micoureus alstoni
San Jose; Valle del Cauca, Colombia (0351'N,
7652'W, Paynter, 1997)
Didelphis marsupialis, 115m
Marmosa robinsoni, 115 m
San Jose Abajo (see San Jose Nuevo); Napo, Ec-
uador
San Jose de Chiquitos (see San Jose, Rio San Mi-
guel); Santa Cruz, Bolivia
San Jose del Sacare; Chalatenango, El Salvador
(1415'N, 8910'W, Gardner, 1973)
Didelphis marsupialis, 3600 ft (= 1125 m)
San Jose do Sumaco (see San Jose Nuevo); Napo,
Ecuador
San Jose Nuevo (= San Jose = San Jose de Su-
maco = San Jose Abajo); Napo, Ecuador
(0026'S, 7720'W, Paynter, 1993)
Didelphis marsupialis, 500-1000 m
Marmosa rubra, 500-1000 m (San Jose Nuevo,
near Volcan Sumaco, below)
Marmosops impavidus, ca. 500-1000 m
Marmosops noctivagus, ca. 3500 ft (= 1090 m)
Metachirus nudicaudatus (0031'S, 7720'W)
Micoureus regina, ca. 500-1000 m (Rio Napo)
Philander andersoni
San Juan; Cartago, Costa Rica (0954'N, 8400'W,
USBGN)
Caluromys derbianus
Micoureus alstoni
San Juan (= San Juan de Telpaneca = Telpaneca);
Madriz, Nicaragua (1333'N, 8610'W, Geo-
net)
Philander opossum, 3500 ft (= 1090 m)
San Juan; Pasco, Peru (0952'S, 7513'W, Geonet)
Didelphis albiventris, ca. 275 m
San Juan, Sandia; Puno, Peru (not precisely lo-
cated)
Didelphis marsupialis
San Juan de Arama; Meta, Colombia (0326'N,
7350'W, Paynter, 1997)
Caluromys lanatus, 400 m
Gracilinanus emiliae, 396 m, 400 m (0320'N,
7353'W)
San Juan de Colon (= Colon); Tachira, Venezuela
(0802'N, 7216'W, Paynter, 1982)
Monodelphis palliolata, 2500 ft (= 780 m)
San Juan de la Punta; Veracruz-Llave, Mexico
(1849'N, 9643'W, Geonet)
Caluromys derbianus
Didelphis marsupialis, 400 ft (= 125 m)
San Juan del Norte (see Greytown); Rio San Juan,
Nicaragua
San Juan de los Morros; Guarico, Venezuela
(0955'N, 6721'W, Geonet)
Caluromys philander
Monodelphis brevicaudata (50 km from San
Juan de los Morros)
San Juan de Manapiare, Rio Manapiare; Amazon-
as, Venezuela (0519'N, 6603'W, Perez-Her-
nandez, 1989)
Caluromys lanatus, 155 m
Caluromys philander, 1 55 m
Didelphis marsupialis, 155 m
Marmosa murina, 155 m
Micoureus demerarae, 155 m
Philander andersoni, 150 m, 155 m
San Juan de Nuevo Mundo (see San Juan Nuevo);
Pando, Bolivia
San Juan de Oxapampa (see Oxapampa); Pasco,
Peru
San Juan de Rioseco; Cundinamarca, Colombia
(0451'N, 7438'W, Paynter, 1997)
Didelphis marsupialis, 1303 m
San Juan de Telpaneca (see San Juan); Madriz,
Nicaragua
San Juan Nepomuceno; Bolivar, Colombia
(0957'N, 7505'W, Paynter, 1997)
Didelphis marsupialis, 167 m
San Julian; Distrito Federal, Venezuela (ca.
1037'N, 6650'W, Paynter, 1982)
Caluromys philander, sea level
Monodelphis palliolata, sea level
San Lorenzo; Loreto, Peru (0449'S, 8012'W,
Geonet)
Didelphis marsupialis, 180 m
Marmosa murina, 500 ft (= 150 m)
San Lorenzo (see Sao Lourenco do Sul); Rio
Grande do Sul, Brazil
San Lourenco do Sul (see Sao Lourenco do Sul);
Rio Grande do Sul, Brazil
San Luis; Cusco, Peru (1306'S, 7225'W, Ste-
phens and Traylor, 1983)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
285
Lestoros inca, 9(XX) ft (= 2800 m) (km 185 on
OUantaytambo-Quellabamba Rd)
San Luis de la Sierra; Chaco, Paraguay (not pre-
cisely located)
Didelphis albiventris
San Marcos; Veracruz-Llave. Mexico (2012'N.
9657'W, Geonet)
Didelphis marsupialis, 200 ft (= 60 m)
Philander opossum. 2(X) ft (= 60 m)
San Marque/. (= Puerto Marquez), Rio Pachitea;
Huanuco, Peru (0932'S. 7456'W. Stephens
and Tray lor. 1983)
Marmosa lepida, ca. HX) m
San Martin de Turumban. Rio Cuyuni; Bolivar.
Venezuela (0659'N. 6102'W. Hershkovitz.
1997)
Monodelphis brevicaudata, 100 m
Philander opossum, 100 m
San Mateo; Lima. Peru (1147'S. 7618'W. Ste-
phens and Tray lor. 1983)
Didelphis albiventris, 3149 m
San Mateo Yetla (see Yetla); Oaxaca, Mexico
San Miguel; La Guajira. Colombia (1058'N,
7329'W, Paynter. 1997)
Marmosa robinsoni
San Miguel. Palenque; Amazonas, Venezuela (not
located)
Didelphis marsupialis
San Miguel, Sierra Nevada de Santa Marta; La
Guajira. Colombia (1058'N, 7329'W, Payn-
ter, 1997)
Marmosa robinsoni
San Miguel de Tucuman (see Tucuman); Tucu-
man, Argentina
San Miguel Duenas (= Duenas); Suchitepequez,
Guatemala (1431'N, 9048'W. Geonet)
Philander opossum
San Miguel Island (see Isla San Miguel); Panama.
Panama
San Miguel Rincon; Santa Cruz. Bolivia (1723'S,
6332'W. Anderson et al., 1993)
Didelphis marsupialis, 300 m
Marmosops dorothea
Philander opossum
San Nicolas (see Mendoza); Amazonas, Peru
San Pablo; Narino. Colombia (0106'N. 7801'W,
Paynter. 1997)
Micoureus regina, 1400 m
San Pablo; Pasco. Peru (ca. 1027'S. 7452'W.
Stephens and Tray Kir. 1983)
Didelphis marsupialis. 900 ft (= 280 m)
San Pablo: Tucuman. Argentina (2652'S.
6519'W. Paynter. 1985)
Lutreolina crassicaudata (11 km N of San Pab-
lo)
Thylamys venustus, ca. 500 m
San Pablo. 1.5 km E of El Mojon; Beni, Bolivia
(1352'S. 6536'W. Hershkovitz, 1997)
Philander opossum (also at 1305'S, 6459'W)
San Pablo Village; Beni. Bolivia (1305'S,
6459'W. Anderson. 1997)
Didelph is ma rsupialis
San Pedro. 10 km W; Beni. Bolivia (1420'S.
6455'W, Anderson. 1997)
Lutreolina crassicaudata
San Pedro; Misiones, Argentina (2638'S,
5408'W, Paynter. 1985)
Philander opossum
San Pedro, above, at km 152, 72 km NE of Pau-
cartambo; Cusco. Peru (ca. 1309'S, 7115'W,
Pacheco et al., 1993)
Marmosops impavidus, 1500 m
Monodelphis adusta, 825-1500 m
San Pedro (see San Pedro Sula); Cortes. Honduras
San Pedro de Colalao; Tucuman. Argentina
(2614'S. 6529'W, Flores et al., 2000)
Didelphis albiventris
Thylamys pusilla
Thylamys pallidior
Thylamys venustus
San Pedro Montes de Oca: San Jose, Costa Rica
(0956'N, 8403'W, Geonet and Gardner,
1973)
Didelphis marsupialis
San Pedro Sula. 2 mi W. near Laguna Ticamaya;
Cortes. Honduras (1530'N, 8802'W. Geo-
net)
Caluromys derbianus
Didelphis marsupialis
Philander opossum
San Pedro Tapanatepec (see Tapanatepec); Oaxa-
ca, Mexico
San Rafael (see Puerto San Rafael); Itapua, Par-
aguay
San Rafael; Mendoza, Argentina (3437'S,
6817'W, Paynter, 1985)
Didelphis albiventris, 686 m
San Rafael, 19 km SW of Villa Tunari; Cocha-
bamba, Bolivia (Villa Tunari, ca. 1657'S,
6524'W, Paynter. 1992)
Metachirus nudicaudatus
San Rafael; St. George. Trinidad and Tobago
(1034'N, 6116'W, Geonet)
Didelphis marsupialis, 100 ft (= 30 m)
San Rafael de Amboro: Santa Cruz, Bolivia
(1736'S, 6336'W, Anderson et al., 1993)
Didelphis marsupialis, 400 m
286
FIELDIANA: ZOOLOGY
Marmosops noctivagus
Monodelphis domestica (hunting camp,
1739'S, 6348'W)
San Rafael del Norte; Jinotega, Nicaragua
(1313'N, 8607'W, Geonet)
Marmosa mexicana, 5000 ft (= 1560 m)
San Ramon (= Monte Chocolatal); Beni, Bolivia
(1318'S, 6437'W, Hershkovitz, 1997)
Philander opossum (W bank Rio Machupo)
San Ramon; Junfn, Peru (1108'S, 7520'W, Ste-
phens and Tray lor, 1983)
Caluromys lanatus, 820 m
Didelphis marsupialis, 820 m
Marmosa murina, 820 m
Marmosops impavidus, 820 m, 1275 m
Marmosops noctivagus, 820 m
Metachirus nudicaudatus, 2900 ft (= 900 m)
San Ramon; Matagalpa, Nicaragua (not located)
Chironectes minimus
San Ramon; Santa Cruz, Bolivia (1733'S,
6103'W, Herskovitz, 1997)
Marmosops dorothea
Philander opossum
San Ramon de la Nueva Oran (= Oran), 24 km
NW of Aguas Blancas; Salta, Argentina
(2308'S, 6420'W, Paynter, 1985)
Lutreolina crassicaudata (2244'S, 6422'W)
Micoureus constantiae
Thylamys venustus, 362 m
San Salvador de Jujuy (= Jujuy); Jujuy, Argentina
(2411'S, 651 8' W, Paynter, 1985)
Thylamys venustus, 1259 m
San Sebastian (see San Sebastian de Rabago);
Cesar, Colombia
San Sebastian de Rabago (= San Sebastian); Ces-
ar, Colombia (1034'N, 7336'W, Paynter,
1997)
Didelphis marsupialis, 2000 m, 2200 m
San Sebastian de Yah (see Yalf); Jinotega, Nica-
ragua
Santa Ana; Cusco, Peru (ca. 1252'S, 7243'W,
Stephens and Traylor, 1983)
Didelphis marsupialis, 1060 m
Santa Ana; Loja, Ecuador (0414'S, 7917'W,
USBGN)
Micoureus regina
Santa Ana; San Jose, Costa Rica (0956'N,
8411'W, Geonet)
Caluromys derbianus (2 km N of Santa Ana)
Didelphis marsupialis
Micoureus alstoni (2 km NW of Santa Ana)
Philander opossum
Santa Ana [Anna] de Chapada (see Chapada dos
Guimaraes); Mato Grosso, Brazil
Santa Ana Trail, between Zaruma and Loja; Loja,
Ecuador (0371'S, 7925'W, Zaruma =
0341'S, 7937'W, Loja = 0400'S, 7913'W,
Geonet)
Micoureus regina, 3650 ft (= 1140 m)
Santa Barbara; Antioquia, Colombia (0625'N,
7607'W, Geonet)
Caenolestes convelatus, 2700 m, 2800 m, 3000
m, 3100 m
Santa Barbara; Jujuy, Argentina (2336'S,
6504'W, Paynter, 1985)
Gracilinanus agilis
Thylamys venustus (2417'S, 6424'W, Flores et
al., 2000)
Santa Barbara (see Serra do Caraga); Minas Ger-
ais, Brazil
Santa Barbara; Santa Barbara, Honduras
(1455'N, 8814'W, Geonet)
Didelphis marsupialis
Santa Cecilia; Napo, Ecuador (0003'N, 7658'W,
Paynter, 1993)
Caluromys lanatus
Chironectes minimus
Didelphis marsupialis, 340 m
Metachirus nudicaudatus, 340 m
Micoureus regina
Philander andersoni
Santa Clara, Villa Bella Imperatriz; Amazonas,
Brazil (0250'S, 5655'W, Hershkovitz, 1997)
Philander opossum
Santa Clara del Mar; Buenos Aires, Argentina
(3750'S, 5730'W, Geonet)
Monodelphis dimidiata
Santa Cruz; Barima-Waini, Guyana (0740'N,
5945'W, Geonet)
Philander opossum
Santa Cruz; Loreto, Peru (0533'S, 7548'W, Ste-
phens and Traylor, 1983)
Marmosa lepida, 149 m (right bank Rio Hual-
laga)
Santa Cruz (= Hacienda San Pedro = Santa Cruz
de Mora); Merida, Venezuela (0825'N,
7139'W, Paynter, 1982)
Didelphis marsupialis
Micoureus demerarae, 950 m
Monodelphis palliolata, 950 m
Santa Cruz (see Santa Cruz de la Sierra); Santa
Cruz, Bolivia
Santa Cruz, right bank, mouth of Rio Eini, a trib-
utary of Rio Jurua; Amazonas, Brazil (Rio
Eiru = 0642'S, 6952'W, Paynter and Tray-
lor, 1991)
Caluromys lanatus
Santa Cruz de Cafia (see Cana); Darien, Panama
BROWN: ATLAS OF NEW WORLD MARSUPIALS
287
Santa Cruz, 27 km SE; Santa Cruz, Bolivia
(1758'S, 63()3'W, Anderson, 1997)
Marmosops dorothea
Micoureus constantiae
Monodelphis domestic a
Philander opossum (15 km SW of Santa Cruz
= 1754'S, 6326'W, 15 km S of Santa Cruz
= 1753'S, 63()7'W)
Santa Cruz de la Sierra (= Santa Cruz); Santa
Cruz, Bolivia (1748'S, 6310'W, Paynter,
1992)
Caluromys lanatus
Didelphis marsupialis, sea level (5 km from
Santa Cruz on Rio Yapacani = 1750'S,
6320'W), 600 m
Marmosops dorothea, ca. 500 m (15 km S of
Santa Cruz = 1753'S, 6307'W)
Metachirus nudicaudatus
Micoureus constantiae, 480 m
Monodelphis domestica
Philander opossum (18 km SW of Santa Cruz)
Thylamys pusilla
Santa Cruz de Mora (see Santa Cruz); Merida,
Venezuela
Santa Domingo (see Inca Mines); Puno, Peru
Santa Elena (see Hacienda Santa Elena); Huanu-
co, Peru
Santa Elena, Rio Samiria; Loreto, Peru (ca.
0450'S, 7413'W, Stephens and Traylor,
1983)
Caluromys lanatus, 130 m
Didelphis marsupialis, 130 m (0442'S,
7412'W)
Metachirus nudicaudatus, 100 m, 130 m
Philander andersoni, ca. 130 m (0442'S,
7412'W)
Santa Elena de Guairen (see Santa Elena de Uair-
en); Bolivar, Venezuela
Santa Elena de Uairen (= Santa Elena de Guair-
en); Bolivar, Venezuela (0437'N, 6108'W,
Paynter, 1982)
Marmosa murina
Santa Fe; Santa Fe, Argentina (3138'S, 6042'W,
Paynter, 1985)
Lutreolina crassicaudata
Santa Fe, Rio Santa Maria; Veraguas, Panama
(0831'N, 8105'W, Geonet)
Caluromys derhianus
Didelphis marsupialis
Santafe de Bogota (see Bogota); Cundinamarca,
Colombia
Santa Gertrudes, Araras; Para, Brazil (ca. 0322'S,
5514'W, Hershkovitz files)
Caluromys philander
Santa Ines, 5 km ESE of Caripe; Monagas, Ve-
nezuela (Caripe = 1012'N, 6329'W, Payn-
ter. 1982)
Marmosops fuscatus, 800 m
Santa Isabel (see Tapurucuara); Amazonas, Brazil
Santa Juana, 10 km NE of Quepos; San Jose, Cos-
ta Rica (0931'N, 8407'W, Geonet)
Philander opossum, 500 m
Santa Lucrecia (see Jesus Carranza); Veracruz-
Llave, Mexico
Santa Luisa, Rio Nanay; Loreto, Peru (0335'S,
7430'W, Hershkovitz, 1997)
Metachirus nudicaudatus, 160 m
Philander andersoni
Santa Luzia; Minas Gerais, Brazil (1947'S,
4352'W, Paynter and Traylor, 1991)
Didelphis albiventris, 681 m
Santa Maria, Braganca; Para, Brazil (0103'S,
4646'W, Hershkovitz, 1997)
Philander opossum
Santa Maria, Mariscal Ramos Castilla; Loreto,
Peru (not located)
Caluromys lanatus
Santa Maria da Serra; Sao Paulo, Brazil (2233'S,
4810'W, USBGN)
Didelphis albiventris
Santa Maria de Itabira; Minas Gerais, Brazil
(1927'S, 4308'W, USBGN)
Didelphis aurita
Santa Maria del Erebato; Bolivar, Venezuela
(0458'N, 6450'W, Perez-Hernandez, 1989)
Chironectes minimus
Santa Maria de Ostuma; Matagalpa, Nicaragua
(1257'N, 8558'W, USBGN)
Didelphis marsupialis, 1250 m
Philander opossum, 1250 m
Santa Marta; Magdalena, Colombia (1115'N,
7413'W, Paynter, 1997)
Caluromys lanatus
Chironectes minimus, sea level
Didelphis marsupialis, sea level (Santa Marta
region)
Marmosa robinsoni, sea level
Santa Marta, near Sierra Nevada de Santa Marta;
Magdalena, Colombia (1114'N, 7405'W,
Paynter, 1997)
Caluromys lanatus, 4000 ft (= 1250 m) (near
Bonda)
Santa Marta, San Agustfn; Huila, Colombia
(0156'N, 7632'W, Bublitz, 1987)
Caenolestes fuliginosus, 2700 m
Santana do Cariri; Ceara, Brazil (0711'S,
3944'W, USBGN)
Monodelphis domestica
288
FIELDIANA: ZOOLOGY
Santana do Ipanema; Alagoas, Brazil (0922'S,
3714'W, USBGN)
Monodelphis domestica (Sitio Lagoinha)
Santarem, right bank Rio Tapajos; Para, Brazil
(0226'S, 5442'W, Paynter and Traylor,
1991)
Caluromys philander (Mojui dos Campos)
Didelphis marsupialis, 15 m (Mojui dos Cam-
pos)
Gracilinanus emiliae, 51 m (mouth Rio Tapa-
jos)
Marmosa murina, 15 m (Mojui dos Campos)
Metachirus nudicaudatus
Santa Rita, near Rio Manapire; Guarico, Vene-
zuela (0608'N, 6615'W, Perez-Hernandez,
1989)
Marmosa robinsoni
Santa Rita (= Santa Rita do Weil), Tabatinga, left
bank Rio Solimoes; Amazonas, Brazil
(0329'S, 6919'W, Paynter and Traylor,
1991)
Marmosops noctivagus
Santa Rita, right bank Rio Nanay, near Iquitos;
Loreto, Peru (Iquitos = 0346'S, 7315'W,
Stephens and Traylor, 1983)
Caluromys lanatus, 20 m
Didelphis marsupialis, 120 m
Santa Rita, 2 km and 4 km NW; Jujuy, Argentina
(2427'S, 6450'W, Flores et al., 2000)
Thylamys venustus
Santa Rita do Supucai; Minas Gerais, Brazil
(2215'S, 4540'W, USBGN)
Didelphis aurita
Santa Rita do Weil (see Santa Rita); Amazonas,
Brazil
Santa Rosa; Beni, Bolivia (1301'S, 6511'W,
Hershkovitz, 1997)
Philander opossum
Santa Rosa; Boaco, Nicaragua (1237'N, 8531'W,
Geonet)
Chironectes minimus, 300 m (7 km N, 15 km
E of Boaca)
Didelphis marsupialis (also at 7 km N, 15 km
E of Boaca)
Santa Rosa; Cartago, Costa Rica (ca. 0955'N,
8351'W, Geonet)
Didelphis marsupialis, 2300 m
Santa Rosa; El Oro, Ecuador (0327'S, 7958'W
Paynter, 1993)
Didelphis albiventris, ca. 100 m
Didelphis marsupialis, ca. 100 m
Marmosa robinsoni, ca. 100 m
Santa Rosa; Mendoza, Argentina (3315'S,
6809'W, Paynter, 1985)
Thylamys pallidior
Santa Rosa; Pando, Bolivia (1213'S, 6824'W,
Anderson, 1997)
Marmosops noctivagus (also at 8 km SW of
Santa Rosa = 1216'S, 6827'W)
Metachirus nudicaudatus
Monodelphis brevicaudata
Santa Rosa; Pichincha, Ecuador (not located)
Didelphis albiventris
Santa Rosa, above Rio Pita; Pichincha, Ecuador
(0020'S, 7826'W, Geonet)
Caenolestes caniventer
Santa Rosa, Alto Ucayali; Ucayali, Peru (1043'S,
7353'W, Hershkovitz, 1997)
Didelphis marsupialis
Philander opossum
Santa Rosa, Mindo; Pichincha, Ecuador (Mindo
= 0002'S, 7848'W, Paynter, 1983)
Caenolestes convelatus
Caenolestes fuliginosus, 9600 ft (= 3000 m)
Santa Rosa, 1 km N of Merida; Merida, Venezue-
la (Merida = 0836'N, 7108'W, Paynter,
1982)
Chironectes minimus, 1860 m
Santa Rosa, 7 km N; Santa Cruz, Bolivia
(1703'S, 6335'W, Anderson, 1997)
Didelphis marsupialis
Micoureus constantiae
Philander opossum
Santa Rosa de Huayabamba (= Guaiabamba =
Guayabamba); Amazonas, Peru (0622'S,
7725'W Stephens and Traylor, 1983)
Metachirus nudicaudatus, 5500 ft (= 1700 m)
Santa Rosa de Leales; Tucuman, Argentina
(2709'S, 6515'W, Mares et al., 1966)
Didelphis albiventris
Santa Rosita, 3 mi S of Warnes; Santa Cruz, Bo-
livia (1730'S, 6310'W, Hershkovitz, 1997)
Didelphis marsupialis
Marmosops dorothea
Philander opossum
Santa Teresa (= Mato do Larva d'Agua); Espfrito
Santo, Brazil (1955'S, 4036'W, Paynter and
Traylor, 1991)
Chironectes minimus, 659 m
Didelphis aurita, 659 m
Philander frenata
Santa Teresa (see Urucum); Mato Grosso do Sul,
Brazil
Santa Teresa, near San Ignacio; Misiones, Argen-
tina (San Ignacio = 2716'S, 5532'W, Payn-
ter, 1985)
Lutreolina crassicaudata (30 km W of San Ig-
nacio)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
289
Santa Teresa, Peralta; Cartago, Costa Rica
(0956'N. 8347'W, Geonet)
Didelphis marsupialis
Santa Victoria (see El Breal); Salta, Argentina
Santiago; Veraguas. Panama (08()6'N. 8059'W.
Geonet)
Caluromys derbianus
Santiago; Santa Cruz, Bolivia (18 19'S. 5935'W,
Anderson. 1997)
Gracilinanus agilis
Philander opossum
Santiago de Chiquitos; Santa Cruz. Bolivia
(1818'S, 5936'W, Anderson. 1997)
Gracilinanus agilis, 700 m. 730 m
Monodelphis domestica
Santisma Trinidad (see Trinidad); Central. Para-
guay
Santo Antonio da Amatary (= Santo Antonio do
Amaterei = Santo Antonio do Amateri);
Amazonas. Brazil (ca. 300'S. 58WW. Voss
et al.. 2001)
Monodelphis brevicaudata
Santo Antonio de Uayara, Rio Eiru; Amazonas,
Brazil (0643'S, 6952'W, Hershkovitz, 1997)
Philander opossum
Santo Antonio do Amaterei (see Santo Antonio
da Amatary); Amazonas. Brazil
Santo Antonio do Amateri (see Santo Antonio da
Amatary); Amazonas, Brazil
Santo Dios, 2 km NE of San Joaquin; Beni, Bo-
livia (San Joaquin = 1304'S, 6449'W.
Hershkovitz. 1997)
Philander opossum
Santo Domingo (= Santo Domingo Petapa).
mountains near; Oaxaca. Mexico (1650'N.
9509'W, Gardner, 1973, and Geonet)
Didelphis marsupialis
Santo Domingo, Rfo Inambari (see Inca mines);
Puno, Peru
Santo Domingo (see Santo Domingo de los Co-
lorados); Pichincha. Ecuador
Santo Domingo; Santiago del Estero. Argentina
(not located)
Didelphis albiventris
Santo Domingo de Bogota; Cundinamarca. Co-
lombia (Bogota = 0424'N, 7433'W.
USBGN)
Micoureus alstoni
Micoureus demerarae
Santo Domingo de Guzman, Isthmus de Tehuan-
tepec; Oaxaca. Mexico (1620'N. 9514'W.
Geonet)
Marmosa canescens
Santo Domingo de los Colorados (= Santo Do-
mingo); Pichincha, Ecuador (0015'S,
7909'W, Paynter. 1993)
Marmosa robinsoni, 1600 ft (= 500 m)
Marmosops impavidus, 500 m
Santo Domingo Petape (see Santo Domingo); Oa-
xaca, Mexico
Santo Isidoro. Tefe (= Lago de Tefe = Tefe);
Amazonas. Brazil (0327'S. 6447'W. Hersh-
kovitz. 1997)
Metachiris nudicaudaius
Philander opossum
San Vicente de Chucurf; Santander. Colombia
(0654'N. 7325'W, Paynter, 1997)
Didelphis marsupialis, 692 m
Metachirus nudicaudatus (0653'N, 7325'W)
San Vito de Java; Puntarenas. Costa Rica
(0850'N, 8258'W, Geonet)
Didelphis marsupialis, 900-1800 m
Sao Bento; Goias, Brazil (1827'S, 5147'W, Avi-
la Pires. 1968)
Gracilinanus agilis
Sao Domingos; Mato Grosso. Brazil (ca. 1330'S.
5123'W. Paynter and Traylor. 1991)
Didelphis albiventris, 457 m
Sao Domingos do Prata: Minas Gerais. Brazil
(1958'S. 4255'W, Varejao and Valle, 1982)
Didelphis aurita
Sao Felipe (see Joao Pessoa); Amazonas. Brazil
Sao Francisco; Espfrito Santo. Brazil (1845'S.
4053'W. Varejao and Valle. 1982)
Didelphis aurita
Sao Felipe (see Joao Pessoa); Amazonas, Brazil
Sao Gabriel (see Uaupes); Amazonas. Brazil
Sao Goncalo do Rio Abaixo (see Ciudade de Sao
Goncalo do Rio Abaixo); Minas Gerais. Bra-
zil
Sao Goncalo dos Campos. 30 km SW of Feira de
Santana; Bahia, Brazil (1225'S, 3858'W.
Geonet)
Marmosops incanus
Sao Goncalo do Supacaf. Parque Estadual do Rio
Doce (see Parque Estadual do Rfo Preto); Mi-
nas Gerais. Brazil
Sao Joao da Serra do Norte; Mato Grosso. Brazil
(1645'S. 5303'W, Hershkovitz, 1992)
Gracilinanus agilis
Sao Joao de Jacutinga (see Caratinga); Minas
Gerais. Brazil
Sao Joao de Petropolis: Espfrito Santo. Brazil
(1949'S. 4040'W. Paynter and Traylor.
1991)
Philander frenata
Sao Joao do Monte Negro (= Montenegro); Rio
290
FIELDIANA: ZOOLOGY
Grande do Sul, Brazil (2942'S, 5128'W,
Paynter and Tray lor, 1991)
Didelphis albiventris
Sao Joao dos Patos; Maranhao, Brazil (0630'S,
4342'W, USBGN)
Didelphis albiventris, 220 m
Sao Joao Marcos; Rio de Janeiro, Brazil (2254'S,
4358'W, Hershkovitz, 1997)
Marmosops incanus
Philander frenata
Sao Jose, near Faro (see Faro); Para, Brazil
Sao Jose da Lagoa, Rio doce, upper Rio Piricaba
(see Fazenda Almada); Bahia, Brazil
Sao Lorenzo (see Sao Lourenco do Sul); Rio
Grande do Sul, Brazil
Sao Lourenco (see Sao Lourenco do Sul); Rio
Grande do Sul, Brazil
Sao Lourenco do Sul (= San Lorenzo = San Lou-
renco do Sul = Sao Lourenco = Sao Lor-
enzo); Rio Grande do Sul, Brazil (3122'S,
5158'W, Paynter and Traylor, 1991)
Chironectes minimus, sea level
Gracilinanus microtarsus, near sea level
Lutreolina crassicaudata, sea level
Monodelphis dimidiata, sea level
Monodelphis sorex, sea level
Sao Luis (= Maranhao); Maranhao, Brazil
(0236'S, 4414'W, Paynter and Traylor,
1991)
Didelphis marsupialis, sea level
Sao Luis de Caceres (see Caceres); Mato Grosso,
Brazil
Sao Marcus Road, Urucum; Mato Grosso do Sul,
Brazil (Urucum = 1909'S, 5738'W, Paynter
and Traylor, 1991)
Monodelphis domestica
Sao Miguel do Guama (see Guama); Para, Brazil
Sao Miguel dos Campos; Alagoas, Brazil
(0947'S, 3605'W, Paynter and Traylor,
1991)
Didelphis aurita, 40 m
Sao Paulo; Sao Paulo, Brazil (2332'S, 4637'W,
Paynter and Traylor, 1991)
Didelphis aurita, 580 m
Gracilinanus microtarsus, 580 m
Marmosops incanus, 600 m
Philander frenata (2330'S, 4630'W)
Sao Roque de Minas; Minas Gerais, Brazil
(2020'S, 4623'W, Paynter and Traylor,
1991)
Didelphis albiventris, ca. 500 m
Sao Sebastiao; Sao Paulo, Brazil (2348'S,
4525'W, Paynter and Traylor, 1991)
Caluromys philander
Gracilinanus microtarsus, near sea level
Marmosops incanus, sea level
Monodelphis scalops, sea level
Philander frenata, sea level
Sapitiba (see Sepetiba); Rio de Janeiro, Brazil
Sapitibi (see Sepetiba); Rio de Janeiro, Brazil
Sapoa; Rivas, Nicaragua (1115'N, 8538'W, Geo-
net)
Caluromys derbianus
Philander opossum, 40 m
Sapucai (= Sapucay); Paraguarf, Paraguay
(2540'S, 5655'W, Paynter, 1989)
Didelphis albiventris, ca. 220 m
Gracilinanus agilis, ca. 220 m
Metachirus nudicaudatus
Micoureus demerarae, ca. 220 m
Philander opossum
Thylamys pusilla, ca. 220 m
Sapucajuba (suburb of Belem) (see Belem); Para,
Brazil
Sapucay (see Sapucai); Paraguarf, Paraguay
Sarayacu, Rio Bobonaza; Pastaza, Ecuador
(0144'S, 7729'W, Paynter, 1993)
Caluromys lanatus, 700 m
Didelphis marsupialis, 700 m
Marmosa lepida, 700 m
Metachirus nudicaudatus
Micoureus regina, 2500 ft (= 780 m)
Philander andersoni, 450-500 m
Sarayacu, left bank Rio Ucayali; Loreto, Peru
(0644'S, 7506'W, Stephens and Traylor,
1991)
Didelphis marsupialis, 125 m
Micoureus regina
Philander opossum (0644'S, 7507'W)
Sarcuta; Norte de Santander, Colombia (not lo-
cated)
Caluromys lanatus
Sarmiento, vicinity of; Chubut, Argentina
(4536'S, 6905'W, USBGN)
Lestodelphys halli
Sarmiento (= Los Sarmientos); Tucuman, Argen-
tina (2724'S, 6541'W, Paynter, 1985)
Didelphis albiventris, 4000 ft (= 1250 m)
Sasaima; Cundinamarca, Colombia (0458'N,
7426'W, Paynter, 1997)
Caluromys lanatus
Saul; French Guiana (0337'N, 5312'W, Stephens
and Traylor, 1985)
Marmosa murina
Saut, Tamanoir, Rio Mana; French Guiana
(0509'N, 5345'W, Stephens and Traylor,
1991)
Monodelphis brevicaudata
BROWN: ATLAS OF NEW WORLD MARSUPIALS
291
Savala (see Matagalpa); Matagalpa, Nicaragua
Savana Grande (see Sabana Grande); Amazonas,
Venezuela
Savanillas de Pirns; San Jose, Costa Rica (there
is a Pirn's in Puntarenas on the Pacific Coast,
SW of San Jose = 0930'N, 8420'W. but Sa-
vanillas de Pirn's not precisely located)
Philander opossum
Savanna Grande; St. George, Trinidad and Tobago
(1018'N, 6122'W. USBGN)
Marmosa robinsoni
Saynani; La Paz, Bolivia (1607'S, 6805'W, An-
derson, 1997)
Didelphis albiventris
Seabra; Bahia, Brazil (1225'S, 4146'W,
USBGN)
Didelphis albiventris
Marmosops incanus
Seboyal (see Cebollal); Loja, Ecuador
Segovia River (see Rio Coco); Honduras
Selva Culata (= see La Culata); Merida, Vene-
zuela
Selva Oscura; Araucania, Chile (3822'S,
7211'W, Geonet)
Dromiciops gliroides, 305 m
Selvas de San Camilo, 29 km SSW of Santo Do-
mingo; Apure, Venezuela (0721'N, 7101'W,
Paynter and Traylor, 1 99 1 )
Caluromys lanatus, 24 m
Seminario Cora9ao de Jesus, Corupa; Santa Ca-
tarina, Brazil, (Corupa = 2626'S, 4914'W,
Paynter and Traylor, 1 99 1 )
Gracilinanus microtarsus
Monodelphis sorex
Sena Madureira; Acre, Brazil (0904'S, 6840'W,
Paynter and Traylor, 1991)
Caluromys lanatus
Didelphis marsupialis, 135 m
Senor do Bomfim (see Villa Nova); Bahia, Brazil
Seno Reloncavi, 40 km S; Los Lagos, Chile
(4136'S, 7250'W, Geonet)
Dromiciops gliroides
Sepetiba (= Sapitiba = Sapitibi); Rio de Janeiro,
Brazil (2258'S, 4342'W, Paynter and Tray-
lor, 1991)
Didelphis aurita (2259'S, 4342'W)
Philander frenata
Seringal Altamira (see Altamira). right bank Rio
Jurua; Amazonas. Brazil
Seringal Condor, left bank Rfo Jurua; Amazonas,
Brazil (0645'S. 7()51'W. Patton et al.. 2000)
Didelphis marsupialis
Marmosops impavidus
Marmosops nocti vagus
Metachirus nudicaudatus
Micoureus demerarae
Micoureus regina
Monodelphis emiliae
Philander mcilhennyi
Serinsal Ocidente. right and left banks of Rio Ju-
rua; Acre. Brazil (0834'S. 7248'W, Patton
et al., 2000)
Philander opossum
Seringal Oriente (= Oriente); Acre, Brazil
(0848'S, 7246'W, Paynter and Traylor,
1991)
Philander opossum (E bank Rio Jurua)
Seringal Penedo, right bank Rio Jurua; Amazonas,
Brazil (0818'S, 7109'W, Geonet)
Philander mcilhennyi
Seringal Sobral, left bank Rio Jurua; Acre, Brazil
(0642'S, 7003'W, Geonet)
Philander mcilhennyi
Serinha (see Serrinha); Sao Paulo, Brazil
Serra; Espirito Santo, Brazil (2007'S, 4018'W,
Hershkovitz, 1997)
Philander frenata
Serra Boa Vista, Rio Ipiranga (see Boa Vista); Sao
Paulo, Brazil
Serra da Cantareira (= Cantareira = Cantareira
Florestal Preserve); Sao Paulo, Brazil
(2325'S, 4639'W, Paynter and Traylor,
1991)
Monodelphis iheringi
Philander frenata (upper Rio Tiete)
Serra da Lua; Roraima, Brazil (0215'N, 6045'W,
Paynter and Traylor, 1991)
Didelphis marsupialis (near Boa Vista)
Micoureus demerarae (near Boa Vista, Rio
Branco)
Serra de Ibiapaba (see Serra Ibiapaba); Ceara,
Brazil
Sena de Itatiaia; Sao Paulo, Brazil (2255'S,
4528'W, Hershkovitz. 1997)
Philander frenata
Sena de Macae; Rio de Janeiro, Brazil (2219'S,
4220'W, Paynter and Traylor, 1991)
Monodelphis sorex
Serra de Parintins; Amazonas, Brazil (0235'S,
5625'W, Hershkovitz, 1997)
Philander opossum
Serra de Tucuman; Tucuman, Argentina (not lo-
cated)
Thylamys pallidior
Serra de Tumucamaque (= Tumucamaque); Ama-
pa, Brazil (0220'N, 5500'W, Paynter and
Traylor, 1991)
Monodelphis brevicaudata
292
FIELDIANA: ZOOLOGY
Serra do Bananal; Sao Paulo, Brazil (ca. 2245'S,
4420'W, Paynter and Traylor, 1991)
Didelphis aurita
Serra do Caraca (= Santa Barbara); Minas Gerais,
Brazil (1908'S, 4330'W, Varejao and Valle,
1982)
Didelphis albiventris, 1950 m
Serra do Herval (see Erval); Rio Grande do Sul,
Brazil
Serra do Japf, 7 km W of Jundiai; Sao Paulo, Bra-
zil (2314'S, 4657'W, Patton and Da Silva,
1997)
Philander frenata
Serra do Mar; Sao Paulo, Brazil (2325'S,
4505'W, Hershkovitz, 1977)
Didelphis aurita
Serra do Navio; Amapa, Brazil (0059'N,
5203'W, Paynter and Traylor, 1991)
Caluromys philander
Didelphis marsupialis, ca. 100 m
Marmosops pinheiroi, ca. 100 m (Rio Amapari)
Metachirus nudicaudatus (Maraca)
Monodelphis brevicaudata
Philander opossum
Serra dos Orgaos; Rio de Janeiro, Brazil (2222'S,
4245'W, Paynter and Traylor, 1991)
Chironectes minimus
Didelphis aurita
Gracilinanus microtarsus
Marmosops incanus
Monodelphis americana, 2318 m
Monodelphis theresa, 2318 m
Philander frenata (2256'S, 4332'W)
Serra dos Parecis (= Chapadao dos Parecis); Mato
Grosso, Brazil (1300'S, 6000'W, Paynter
and Traylor, 1991)
Metachirus nudicaudatus
Serra Ibiapaba (= Cerra de Ibiapaba = Cerra Ibia-
paba = Ibiapaba = Serra de Ibiapaba); Ceara,
Brazil (0503'S, 4055'W, Hershkovitz, 1992)
Gracilinanus agilis, 300 m
Monodelphis domestica, 900 m
Serrania Bella Vista; La Paz, Bolivia (1541'S,
6730'W, Anderson, 1997)
Marmosops dorothea, 1525 m
Metachirus nudicaudatus
Serrania de Calilegua (= Calilegua Mountain =
Cerro Calilegua); Jujuy, Argentina (2335'S,
6455'W, Paynter, 1985)
Didelphis albiventris, 2600 m
Lutreolina crassicaudata
Thylamys pallidior, 2000 m
Thylamys venustus, ca. 2000 m
Serrania de Copey, Margarita; Nueva Esparta, Ven-
ezuela (lTOO'N, 6400'W, Perez-Hernandez,
1989)
Marmosa robinsoni, 800 m
Serrania del Darien; Chocd, Colombia (ca.
0825'-0830'N, 7730'W, Paynter, 1997)
Monodelphis adusta, 0-1700 m
Serrania del Turimiquire; Sucre, Venezuela
(1003'N, 6354'W, Lopez-Fuster et al.,
2000)
Marmosa robinsoni
Serrania de Macarena (see La Macarena); Meta,
Colombia
Serrania Sama; Tarija, Bolivia (2129'S, 6502'W,
Paynter, 1992)
Thylamys pallidior, 3200 m
Serrinha; Bahia, Brazil (1139'S, 3900'W, Cer-
queira, 1984)
Didelphis albiventris
Marmosops incanus
Monodelphis domestica (Fazenda Cacua, Fa-
zenda Oiteiro, and Fazenda Umburana =
1332'S, 3936'W, USBGN)
Serrinha (= Serinha); Sao Paulo, Brazil (2109'-
2129'S, 5047'-4731'W, USBGN)
Didelphis aurita
"S. Franciso de C"; Minas Gerais, Brazil (not lo-
cated)
Philander opossum, 1580 m
Shahuia; Ucayali, Peru (0952'S, 7401'W, Hersh-
kovitz, 1997)
Philander opossum, 312 m (Rio Ucayali, op-
posite Cumana)
Shintuya, right bank Rio Alto Madre de Dios;
Madre de Dios, Peru (ca. 1240'S, 7113'W,
Stephens and Traylor, 1983)
Chironectes minimus, 420 m
Sibube; Bocas del Toro, Panama (0936'N,
8249'W, Geonet)
Caluromys derbianus
Chironectes minimus
Sierra de Aroa; Yaracuy, Venezuela (1026'N,
6854'W, Paynter, 1982)
Didelphis marsupialis, 500 m
Sierra de La Peregrina; Buenos Aires, Argentina
(ca. 3755'S, 5742'W, Reig, 1964)
Monodelphis dimidiata
Sierra de La Tinta (= La Tinta); Buenos Aires,
Argentina (3737'S, 5933'W, Paynter, 1985)
Monodelphis dimidiata
Sierra de Famatima (see Loma de Famatima); La
Rioja, Argentina
Sierra de la Ventana (= Abra de La Ventana);
Buenos Aires, Argentina (3809'S, 6159'W,
Paynter, 1985)
BROWN: ATLAS OF NEW WORLD MARSUPIALS
293
M on odelph is dim idia ta
Thylamys pallidior, 1 243 m
Sierra de Merida; Merida, Venezuela (0840'N.
7100'W, Paynter, 1982)
Gracilinanus marica, 1600 m
Sierra de Tilcara; Jujuy, Argentina (2335'S,
6512'W, Floreset al., 2000)
Thylamys pallidior, 4500 m
Sierra Madre Oriental, E slope; Tamaulipas, Mex-
ico (not preeisely located)
Philander opossum
Sierra Montanas (see Montanas de la Sierra); Me-
rida, Venezuela
Sierra Nahuelbuta (see Cerro Nahuelbuta); Los
Lagos, Chile
Sierra Negra, E of Villanueva; La Guajira, Co-
lombia (1037'N, 7255'W, Paynter, 1997)
Caluromys lanatus
Marmosa robinsoni, 1265 m
Marmosops impavidus, 1265 m
Sierra Nevada de Santa Marta (see Palomino and
Santa Marta); Magdalena, Colombia
Sierra Nevada de Merida; Merida, Venezuela
(0833'N, 7103'W, Paynter, 1982)
Didelphis marsupialis, 7500 ft (= 2340 m)
Sierra Parima; Amazonas, Venezuela (0240'N,
6430'W, Hershkovitz, 1997)
Philander andersoni, 200 m
Sierra Santa Barbara (see Cerro Santa Barbara);
Jujuy, Argentina
Sierras de Medina; Tucuman, Argentina (2624'S,
6507'W, USBGN)
Lutreolina crassicaudata
Siguatepeque, Lempira; Honduras (1442'N,
8838'W, Geonet)
Chironectes minimus
Simao Pereira; Minas Gerais, Brazil (2157'S,
4320'W, Mustrangi and Patton, 1997)
Marmosops incanus
Sinche; Bolfvar, Ecuador (0132'S, 7859'W,
Paynter, 1993)
Caenolestes fuliginosus, 11000 ft (= 3400 m)
Sinnamarey; Cayenne, French Guiana (0523'N,
5257'W, Stephens and Traylor, 1985)
Philander opossum
Sinnamarey River; French Guiana (0527'N,
5300'W, Stephens and Traylor, 1985)
Didelphis alhiventris
Didelphis marsupialis
Metachirus nudicaudatus
Sipaliwini Airstrip; Nickerie, Suriname (ca.
0205'N, 5610'W, Stephens and Travlor
1985)
Caluromys philander
Didelphis marsupialis
Philander opossum
Sipurio (= "Talamanca"); Limon, Costa Rica
(0932'N, 8255'W, Geonet)
Caluromys derhianus
Didelphis marsupialis
Siquirres, Rio Pacuare; Limdn, Costa Rica
(1006'N, 8331'W, Geonet)
Micoureus alstoni
Sitio Araruna, Missao Velha (see Missao Velha);
Ceara, Brazil
Sitio Barracao, Limoeira da Anadia (see Limoeira
da Anadia); Alagoas, Brazil
Sitio Belem, Bodoco (see Bodoco); Pernambuco,
Brazil
Sitio Boa Esperan?a de Jerico, Triunfo (see Triun-
fo); Pernambuco, Brazil
Sitio Borgens, Triunfo (see Triunfo); Pernambuco,
Brazil
Sitio Cachoeira Grande, Vicosa (see Vicosa); Al-
agoas, Brazil
Sitio Camara, Milagres (see Milagres); Ceara,
Brazil
Sitio Cana Brava de Jerico, Triunfo (see Triunfo);
Pernambuco, Brazil
Sitio Capuma, Palmeira dos Indios (see Palmeira
dos Indios); Alagoas, Brazil
Sitio Cereado, Jardim, Solonopole (see Solono-
pole); Ceara, Brazil
Sitio Cinta da Solidade, Missao Velho (see Missao
Velho); Ceara, Brazil
Sitio Constantino, Crato (see Crato); Ceara, Brazil
Sitio Corredor do Vento, Triunfo (see Triunfo);
Pernambuco, Brazil
Sitio Dormiao, Palmeira dos Indios (see Palmeira
dos Indios); Alagoas, Brazil
Sitio Emboscada, Missao Velho (see Missao Vel-
ho); Ceara, Brazil
Sitio Lagoinha, Santana do Ipanema (see Santana
do Ipanema); Alagoas, Brazil
Sitio Lapinha, Missao Velho (see Missao Velho);
Ceara, Brazil
Sitio Macaco de Baixa Verde, Triunfo (see Triun-
fo); Pernambuco, Brazil
Sitio Novo, Triunfo (see Triunfo); Pernambuco,
Brazil
Sitio Oiti, Triunfo (also see Triunfo); Pernambuco,
Brazil (0833'S, 3926'W, USBGN)
Monodelphis domestica
Sitio Olaria, Crato (see Crato); Ceara, Brazil
Sitio Paus Preto, Bodoco (see Bodoco); Pernam-
buco, Brazil
Sitio Riacho Fundo, Garanhuns (see Garanhuns);
Pernambuco, Brazil
294
FIELDIANA: ZOOLOGY
Sitio Riveira, Palmeira dos indios (see Palmeira
dos Indios); Alagoas, Brazil
Sitio Sabia, Palmeira dos Indios (see Palmeira dos
Indios); Alagoas, Brazil
Sitio Sao Goncalo, Bodoco (see Bodoco); Per-
nambuco, Brazil
Sitio Sao Mateus, Triunfo (see Triunfo); Pernam-
buco, Brazil
Sitio Volta, Campo Sales (see Campo Sales);
Ceara, Brazil
Sitio Xique-Xique, Bodoco (also see Bodoco);
Pernambuco, Brazil (0821'S, 3800'W,
USBGN)
Monodelphis domestica
Soacha, San Miguel-Sibate; Cundinamarca, Co-
lombia (0420'N, 7416'W, Paynter, 1997)
Caenolestes fuliginosus, 2600 m, 2630 m
Sochi; Guerrero, Mexico (not located)
Marmosa canescens
Sobral, left bank Rio Jurua; Acre, Brazil (0822'S,
7249'W, Patton et al., 2000)
Caluromys lanatus
Didelphis marsupialis
Metachirus nudicaudatus
Micoureus demerarae
Micoureus regina
Philander mcilhennyi
Philander opossum
Socorre; Cordoba, Colombia (0751'N, 7617'W,
Paynter, 1997)
Chironectes minimus, 100 m (upper Rio Sinu)
Didelphis marsupialis, 100 m
Metachirus nudicaudatus, 100 m (Rio Sinu)
Solonopole; Ceara, Brazil (0544'S, 3901'W,
Paynter and Traylor, 1991)
Monodelphis domestica (Sitio Cereado)
Solusuchiapa; Chiapas, Mexico (1726'N,
9303'W, Geonet)
Didelphis marsupialis
Sonson; Antioquia, Colombia (0542'N, 7518'W,
Paynter, 1997)
Didelphis albiventris, 2400 m
Didelphis marsupialis, 2545 m
Sophie; French Guiana (0358'N, 5329'W,
USBGN)
Monodelphis brevicaudata
Soure, Ilha de Marajo; Para, Brazil (0044'S,
4831'W, Paynter and Traylor, 1991)
Monodelphis maraxina, ca. 10 m
Soyalo; Chiapas, Mexico (1654'N, 9255'W,
Geonet)
Didelphis marsupialis
Speyside; Trinidad and Tobago (1118'N,
6032'W, Geonet)
Marmosa robinsoni
St. Agustine; St. David, Trinidad and Tobago
(1039'N, 6124'W, Paynter, 1982)
Marmosops fuscatus
St. Eugene; French Guiana (0451'N, 5304'W,
USBGN)
Monodelphis brevicaudata
St. George's; Grenada (1203'N, 6145'W, Geo-
net)
Marmosa robinsoni
Suapo (see Hacienda El Li'mon); Distrito Federal,
Venezuela
Suapure, left bank Rio Caura; Bolivar, Venezuela
(0714'N, 6510'W, Paynter, 1982)
Caluromys philander, ca. 200 m
Marmosa murina, ca. 100 m
Monodelphis brevicaudata, ca. 200 m
Sucre; Chusquisaca, Bolivia (1902'S, 6517'W,
Paynter, 1982)
Thylamys venustus, 3000 m
Sumidouro, near Rio de Janeiro; Minas Gerais,
Brazil (2024'S, 4321'W, Paynter and Tray-
lor, 1991)
Marmosops paulensis, ca. 700 m
Summit; Panama, Panama (0904'N, 7939'W,
Geonet)
Caluromys derbianus
Sumwood Channel, Isla Popa; Bocas del Toro,
Panama (0909'N, 8209'W, Geonet)
Caluromys derbianus
Sunchal, Sierra de Santa Barbara; Jujuy, Argen-
tina (2416'S, 6427'W, Geonet)
Thylamys venustus, 1200 m
Supacay (see Sapucaf)
Supenaam River; Pomeroon-Supenaam, Guyana
(0659'N, 5831'W, Geonet)
Caluromys philander
Didelphis marsupialis
Marmosa murina
Metachirus nudicaudatus
Monodelphis brevicaudata
Philander opossum (0658'N, 5831'W)
Surama, 30 km NE; Potaro-Sipaliwini, Guyana
(0410'N, 5905'W, Paynter and Traylor,
1985)
Philander opossum
Micoureus demerarae
Surco; Lima, Peru (1152'S, 7628'W, Stephens
and Traylor, 1991)
Didelphis albiventris, 2018 m, 2050 m
Thylamys pallidior, 6700 ft (= 2090 m)
Suriname (no specific locality)
Chironectes minimus
BROWN: ATLAS OF NEW WORLD MARSUPIALS
295
Tabacal; Salta, Argentina (2316'S, 6415'W,
Geonet)
Didelphis albiventris
Tabay; Merida, Venezuela (0838'N, 7104'W,
Paynter, 1982)
Didelphis albiventris, 3 1 55 m
Didelphis marsupialis, 1710 m, 2100 m
Gracilinanus dr\as
Tabernilla; Panama, Panama (0907'N, 7949'W,
Wenzel and Tipton, 1966)
Caluromys derbianus
Philander opossum
Tablada (= Tallada); Tarija, Bolivia (2133'S,
6447'W, Hershkovitz, 1997)
Gracilinanus agilis, 2000 m
Thylamys venustus, 2000 m
Tablas Monte, 9.5 km NE; Cochabamba, Bolivia
(1702'S, 6559'W, Anderson, 1997)
Marmosops noctivagus
Micoureus constantiae
Tablas Monte, 4.4 km N; Cochabamba, Bolivia
(1704'S, 6601'W, Anderson, 1997)
Marmosops noctivagus
Micoureus constantiae
Monodelphis osgoodi
Tablahuasi occidente, 8 km W of Quito; Pichin-
cha, Ecuador (Quito = 0013'S, 7830'W,
Paynter, 1993)
Caenolestes fuliginosus
Tabori; Bocas del Toro, Panama (not located)
Philander opossum
Tacarcuna (= Tocarcuna); Darien, Panama (near
Panama-Colombia border) (0805'N, 7717'W,
Gardner, 1973)
Caluromys derbianus
Chironectes minimus, 1950 ft (= 610 m)
Didelph is ma rsup ia lis
Marmosops invictus, 2700 ft (= 840 m) (Ta-
carcuna Casita Camp), 4000 ft (= 1500 m)
(Tacarcuna Laguna Camp)
Metachirus nudicaudatus, 2650 ft (= 830 m),
879 m (Tacarcuna Village Camp)
Monodelphis adusta, 1950 ft (= 610 m), 2000
ft (= 625 m) (also at Tacarcuna Village
Camp)
Tacarigua; Nueva Esparta, Venezuela (lTOO'N,
6400'W, Perez-Hernandez, 1989)
Marmosa robinsoni
Tacuati; San Pedro, Paraguay (2327'S, 5635'W,
Paynter, 1989)
Metachirus nudicaudatus (Aca Poi, S bank Rio
Ypanf)
Philander opossum
Tafi del Valle; Tucuman, Argentina (2652'S,
6541'W, Paynter, 1985)
Didelphis albiventris
Thylamys pallidior, ca. 2000 m
Tafi Viejo (= Cerro de Tafi Viejo); Tucuman, Ar-
gentina (2644'S, 6516'W, Mares et al., 1996)
Lutreolina crassicaudata
Thylamys venustus
Taganga; Magdalena, Colombia (1116'N,
7412'W, Paynter, 1997)
Marmosa robinsoni, sea level
Takama (= Comaccka = Comackka = Coom-
acka); East Demerara-West Coast Berbice,
Guyana (0534'N, 5755'W, Geonet)
Marmosa murina, 50 ft (= 15 m) (180 mi up
Demerara River)
Micoureus demerarae, 160 ft (= 50 m)
"Talamanca" (= see Sipurio), from Limon to
Panama border; Limon, Costa Rica
Tale, 2 mi NE; Chiriqui, Panama (0815'N,
8141'W, Gardner, 1973)
Didelphis marsupialis
Talisman (see El Talisman); Chiapas, Mexico
Tallada (see Tablada); Tarija, Bolivia
Tamanoir; French Guiana (0509'N, 5345'W, Ste-
phens and Traylor, 1985)
Monodelphis brevicaudata
Tamara (= La Tirana), Rio Nechi; Antioquia, Co-
lombia (0722'N, 750O'W, Hershkovitz files)
Metachirus nudicaudatus, 530 m
Tamatama, N bank Rio Orinoco; Amazonas, Ve-
nezuela (0309'N, 6550'W, Paynter, 1982)
Caluromys lanatus
Caluromys philander, 1 35 m
Didelphis marsupialis, 100-135 m
Marmosa murina, 135 m
Metachirus nudicaudatus, 135 m
Monodelphis brevicaudata, 130-135 m
Philander andersoni, 135 m (0308'N,
6552'W)
Tamazunchale; San Luis Potosi, Mexico
(2116'N, 9847'W, Geonet)
Didelphis marsupialis
Tambapu (= Tambaii); Sao Paulo, Brazil
(2142'S, 4716'W, Geonet)
Micoureus demerarae
Tambaii (see Tambapu); Sao Paulo, Brazil
Tambi (see Tanabi); Sao Paulo, Brazil
Tambillo; Cajamarca, Peru (ca. 0610'S, 7845'W,
Stephens and Traylor, 1983)
Marmosa robinsoni, 5800 ft (= 1800 m)
Tambillo, 3 km S; Pichincha, Ecuador (0025'S,
7832'W. Paynter, 1993)
296
FIELDIANA: ZOOLOGY
Didelphis albiventris, 2785 m
Tambo, 2.5 km N; Santa Cruz, Bolivia (1800'S,
6428'W, Anderson, 1997)
Thylamys venustus
Tambo Almirante, near Uchco [Uscho]; San Mar-
tin, Peru (ca. 0608'S, 7720'W, Stephens and
Traylor, 1983)
Marmosops impavidus, 2030 m
Tambo Carrizal, 40 km S of Chachapoyas; Araa-
zonas, Peru (ca. 0647'S, 7752'W, Stephens
and Traylor, 1983)
Marmosops noctivagus, 9000 ft (= 2000 m)
Tambopata; Madre de Dios, Peru (1236'S,
69H"W, Stephens and Traylor, 1983)
Didelphis marsupialis
Metachirus nudicaudatus
Tambo Ventija (see Ventilla); Amazonas, Peru
Tambor, left bank Rfo Jau; Amazonas, Brazil (ca.
0230'S, 6400'W, Patton et al., 2000)
Didelphis marsupialis
Marmosops noctivagus
Metachirus nudicaudatus
Micoureus demerarae
Tanabi (= Tambi); Sao Paulo, Brazil (2037'S,
4937'W, Geonet)
Caluromys philander
Tangoshari, 2 km SW of Rio Pangoreni; Cusco,
Peru (1146'S, 7326'W, Patton et al., 2000)
Micoureus demerarae, 1000 m
Tapacari; Cochabamba, Bolivia (1731'S,
6636'W, Anderson, 1997)
Didelphis albiventris
Tapaiuna, right bank Rio Tapajos; Para, Brazil
(0254'S, 5505'W, Paynter and Traylor,
1991)
Caluromys philander
Tapajos; Roraima, Brazil (not located)
Monodelphis brevicaudata
Tapaliza, near Cerro Tacarcuna; Darien, Panama
(0759'N, 7726'W, Wenzel and Tipton,
1966)
Marmosa robinsoni
Tapanahoni, Paloemeu; Sipaliwini, Suriname
(0422'N, 5427'W, Geonet)
Didelphis marsupialis, 210 m
Tapanatepec (= San Pedro Tapanatepec); Oaxaca,
Mexico (1621'N, 9412'W, Geonet)
Didelphis marsupialis
Philander opossum
Tapecua; Tarija, Bolivia (2126'S, 6355'W, An-
derson, 1997)
Didelphis albiventris
Monodelphis kunsi
Thylamys venustus
Taperinha; Para, Brazil (0232'S, 5417'W, Payn-
ter and Traylor, 1991)
Monodelphis emiliae, ca. 15 m
Tapia; Panama, Panama (0904'N, 7925'W, Geo-
net)
Didelphis marsupialis
Tapia; Tucuman, Argentina (2336'S, 6518'W,
Paynter, 1985)
Thylamys pallidior, 689 m
Thylamys pusilla
Tapikiole (see Estancia Tapikiole); Formosa, Ar-
gentina
Tapirapoan (see Tapirapua); Mato Grosso, Brazil
Tapirapua (= Tapirapoan = Tapirapuan), Rio Se-
potuba; Mato Grosso, Brazil (1451'S,
5745'W, Paynter and Traylor, 1991)
Caluromys philander
Metachirus nudicaudatus
Tapirapuan (see Tapirapua); Mato Grosso, Brazil
Tapua, a few mi E of Asuncion; Central, Paraguay
(Asuncion = 2516'S, 5740'W, Paynter,
1989)
Thylamys macrura
Tapurucuara (= Santa Isabel); Amazonas, Brazil
(0024'S, 6502'W, Paynter and Traylor,
1991)
Didelphis marsupialis, 70 m
Taquara (= Taquara do Mundo Novo); Rio
Grande do Sul, Brazil (2939'S, 5047'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 29 m
Didelphis aurita, 29 m
Gracilinanus microtarsus, 29 m
Monodelphis dimidiata, 29 m
Monodelphis iheringi, 29 m
Monodelphis sorex, 29 m
Philander opossum (2931'S, 5047'W)
Taquara do Mundo Novo (see Taquara); Rio
Grande do Sul, Brazil
Taquari River Valley (= Rio Taquari); Rio Grande
do Sul, Brazil (Taquari = 2948'S, 5151'W,
Paynter and Traylor, 1991)
Didelphis albiventris, 76 m
Tarabuco, 12 km N and 1 km E; Chuquisaca, Bo-
livia (1904'S, 6449'W, Anderson, 1997)
Didelphis albiventris, 2450 m
Thylamys pusilla (also at 1988'S, 6449'W and
1910'S, 6456'W)
Taraguacocha, Cordillera de Chilla; El Oro, Ec-
uador (0340'S, 7940'W, Paynter, 1993)
Didelphis albiventris, 9750 ft (= 3040 m)
Tarapoto, 28 km NE; San Martin, Peru (0630'S,
7625'W, Stephens and Traylor, 1983)
Caluromys lanatus, 750 m
BROWN: ATLAS OF NEW WORLD MARSUPIALS
297
Tarija: Tarija. Bolivia (2131'S, 6445'W, Paynter.
1992)
Thylamys venustus, 6700 ft (= 2090 m)
Tarma; Junin, Peru (1125'S, 7542'W. Stephens
and Tray lor, 1983)
Chironectes minimus
Didelphis albiventris, 8000 ft (= 2500 m) (17
km below Tarma)
Marmosops juninensis
Tartagal; Salta. Argentina (2232'S, 6349'W.
Paynter, 1985)
Monodelphis dimidiata
Thylamys venustus, 502 m (27 km E of Tarta-
gal)
Tatamalas (= Tatemeles): Sinaloa. Mexico (not
located)
Marmosa canescens
Tatemeles (see Tatamalas); Sinaloa. Mexico
Tauari (= Tauary), Rio Tapajos; Para. Brazil
(0305'S, 5506'W, Paynter and Traylor.
1983)
Didelphis marsupialis, ca. 25 m
Tauary (see Tauari); Para. Brazil
Taulis; Cajamarca. Peru (ca. 0654'S. 7903'W.
Stephens and Traylor. 1983)
Marmosops impavidus, 9000 ft (= 2800 m)
Teapa; Tabasco. Mexico (1733'N, 9257'W. Geo-
net)
Caluromys derbianus
Chironectes minimus (2 mi NW of Teapa)
Didelphis marsupialis
Philander opossum ( 1 9 mi N of Teapa)
Teapa; Veracruz-Llave. Mexico (1724'N.
9252'W, Geonet)
Caluromys derbianus
Tefe (see Santo Isidoro); Amazonas. Brazil
Tegucigalpa; Francisco Morazan. Honduras
(1406'N. 8713'W. Geonet)
Chironectes minimus
Didelphis marsupialis
Tehuacan: Puebla. Mexico (1827'N. 9723'W,
Geonet)
Marmosa canescens, 1700 m
Tehuantepec: Oaxaca. Mexico (1620'N. 9514'W.
Geonet)
Marmosa canescens
Tekom; Yucatan. Mexico (2039'N. 8758'W. 4
localities listed within a few km of each oth-
er. Geonet)
Philander opossum
Telpaneca (see San Juan): Madriz. Nicaragua
Temuco: Araucania. Chile (38 C 44'S 7~ >C WW
USBGN)
Dromiciops gliroides, !
Teniente Ochoa; Nueva Asuncion, Paraguay
(21 S 42'S. 6102'W. USBGN)
Thylamys pusilla
Tenochtitlan; Veracruz-Llave, Mexico (1745'N,
9445'W. coordinates given on specimen tag)
Didelphis marsupialis
Philander opossum, 60 m
Teocelo: Veracruz-Llave, Mexico (1923'N,
9658'W. Geonet)
Philander opossum, 4000 ft (= 1250 m)
Teofilo Otoni; Minas Gerais, Brazil (1751'S,
41 C 30'W. Paynter and Traylor. 1991)
Didelphis aurita, 318 m
Tepic; Nayarit. Mexico (2130'N, 10454'W, Geo-
net)
Marmosa canescens, 3000 ft (= 930 m)
Terebinto; Santa Cruz, Bolivia (1743'S, 6323'W,
Geonet)
Micoureus constantiae
Teresinha (= Terezinha); Amapa, Brazil (0058'N,
5202'W, Paynter and Traylor, 1991)
Philander opossum (Rio Amapari)
Teresopolis (= Theresopolis = Therezopolis =
Fazenda Boa Vista); Rio de Janeiro. Brazil
(2226'S, 4259'W, Paynter and Traylor,
1991)
Caluromys philander, 902 m (2227'S,
4257'W)
Chironectes minimus
Didelphis aurita, 902 m. 951 m
Gracilinanus microtarsus, 902 m. 950 m
Marmosops incanus, 902 m
Marmosops paulensis
Metachirus nudicaudatus (2227'S, 4257'W)
Monodelphis americana, 902 m
Monodelphis scalops, 902 m
Monodelphis theresa, 902 m, 3800 ft (= 1180
m), 4000 ft (= 1250 m)
Philander frenata, 3000 ft (= 930 m) (2227'S,
4257'W)
Termales de Ruiz. Manizales; Caldas, Colombia
(0454'N. 7518'W, Patterson and Albuja,
1996)
Caenolestes fuliginosus, 3100-3300 m
Terezinha (see Teresinha); Amapa. Brazil
Teustepe (= Tuestepe); Boaco, Nicaragua
(1225'N, 8548'W, Geonet)
Caluromys derbianus
Chironectes minimus, 140 m
Philander opossum, 140 m
Texolo. first pueblo of Teocleo: Veracruz-Llave,
Mexico (Teocelo = 1923'N, 9858'W, Geo-
net)
Marmosa mexicana
298
FIELDIANA: ZOOLOGY
Theresopolis (see Teresopolis); Rio de Janeiro,
Brazil
Therezopolis (see Teresopolis); Rio de Janeiro,
Brazil
Thermas de San Luis, 30 km E of Pucon; Arau-
cania, Chile (Pucon = 3916'S, 7158'W,
Geonet)
Dromiciops gliroides, 600 m
Ticrapo; Huancavelica, Peru (1321'S, 7525'W,
Stephens and Traylor, 1991)
Thylamys pallidior, 2184 m, 8000 ft (= 2500 m)
Ticunhuaya; La Paz, Bolivia (ca. 1528'S,
6818'W, Paynter, 1992)
Marmosops noctivagus, 4800 ft (= 1500 m)
Micoureus regina, 4800 ft (= 1500 m) (road
from Sorata-Guanay)
Tierra Oscura; Bocas del Toro, Panama (0911'N,
8217'W, Geonet)
Philander opossum
Tignamar; Tarapaca, Chile (1835'S, 6930'W,
Geonet)
Thylamys elegans, 11500 ft (= 3485 m)
Tiguino, 1 30 km S of Coca; Pastaza, Ecuador (Rio
Tigiiino = 0118'S, 7629'W, Geonet)
Metachirus nudicaudatus
Tihuatlan; Veracruz-Llave, Mexico (2043'N,
9732'W, Geonet)
Didelphis marsupialis, 700 ft (= 220 m)
Philander opossum, 300 ft (= 90 m)
Tihumayu; Chuquisaca, Bolivia (1934'S,
6408'W, Anderson, 1997)
Monodelphis domestica
Tijuca; Rio de Janeiro, Brazil (ca. 2256'S,
4317'W, Paynter and Traylor, 1991)
Didelphis aurita
Tikal (see Parque Nacional Tikal); Peten, Guate-
mala
Timotes; Merida, Venezuela (0859'N, 7044'W,
Paynter, 1982)
Didelphis albiventris, 2016 m
Tingo Maria; Huanuco, Peru (0918'S, 7559'W,
Stephens and Traylor, 1 99 1 )
Chironectes minimus, 649 m
Didelphis marsupialis, 649 m
Gracilinanus agilis, 600 m, 649 m
Marmosa murina, 2000 ft (= 625 m), 649 m
Marmosops noctivagus, 2000 ft (= 625 m)
Metachirus nudicaudatus, 3000 ft (= 930 m)
Philander opossum, 2000 ft (= 625 m)
Tita; Santa Cruz, Bolivia (1825'S, 6210'W, Mus-
trangi and Patton, 1997)
Marmosops dorothea, 300 m
Marmosops impavidus
Monodelphis domestica
Philander opossum
Thylamys venustus
Tlapa, Rio Xochihuehuellen; Guerrero, Mexico
(1733'N, 9833'W, Geonet)
Marmosa canescens
Tlapacingo (= Tlapancingo); Oaxaca, Mexico
(1728'N, 9814'W, Geonet)
Marmosa canescens
Tlapacoyan; Veracruz-Llave, Mexico (1958'N,
9713'W, Geonet)
Didelphis marsupialis, 1700 ft (= 530 m)
Tlapancingo (see Tlapacingo); Oaxaca, Mexico
Tobago, Trinidad and Tobago (ca. 1120'N,
61 o 00'-6020'W, Geonet)
Caluromys philander
Didelphis marsupialis
Tobati'; La Cordillera, Paraguay (2515'S,
5704'W, Paynter, 1989)
Didelphis albiventris
Lutreolina crassicaudata
Philander opossum (12 km N of Tobati)
Tobesobe, Guayo; Delta Amacuro Territorio Fed-
eral, Venezuela (0900'N, 6125'W, Hershko-
vitz, 1997)
Metachirus nudicaudatus
Philander opossum
Tocantins; Goias, Brazil (not located)
Didelphis albiventris
Tocarcuna (see Tacarcuna); Darien, Panama
Tocomechi, 15 km NW of Warnes; Santa Cruz,
Bolivia (1735'S, 6255'W, Hershkovitz,
1997, Paynter, 1992)
Didelphis marsupialis
Philander opossum
Tocopoqueu, Ocobamba Valley; Cusco, Peru (ca.
1253'S, 7221'W, Stephens and Traylor,
1983)
Lestoros inca, 9100 ft (= 2840 m)
Monodelphis osgoodi, 2780 m
Tocuco (see Tukuko); Zulia, Venezuela
Todos Santos; Cochabamba, Bolivia (1648'S,
6508'W, Paynter, 1992)
Didelphis marsupialis, 1300 ft (= 400 m)
Tojibos (see Los Tojibos); Beni, Bolivia
Toledo; Norte de Santander, Colombia (0719'N,
7228'W, Paynter, 1997)
Caluromys lanatus
Tolima, near Bogota; Cundinamarca, Colombia
(Bogota = 0436'N, 7405'W, Paynter, 1997)
Caluromys lanatus
Tolloche; Salta, Argentina (2530'S, 6332'W,
Geonet)
Thylamys pusilla
BROWN: ATLAS OF NEW WORLD MARSUPIALS
299
Tomima: Chuquisaca. Bolivia ( 191 IS. 6430'W,
Paynter, 1992)
Monodelphis domestica, 207 1 m
Toocoq; Peten. Guatemala (1641'N. 9002'W.
Gardner. 1973)
Didelphis marsupialis
Torixoreu; Mato Grosso. Brazil (1615'S.
5226'W. Geonet)
Philander opossum
Tornquist; Buenos Aires, Argentina (3806'S.
6214'W. Paynter, 1985)
Monodelphis dimidiata, 550 m
Torono-tepui, Chimanta; Bolivar. Venezuela (ca.
0524'N. 6200'W. Geonet)
Didelphis albiventris, 2 1 00 m
Torontoy; Cuseo. Peru (1310'S, 7230'W. Ste-
phens and Traylor. 1983)
Didelphis albiventris, ca. 2400 m
Lestoros inca. 14000 ft (= 4375 m)
Marniosops impavidus, 8000 ft (= 2500 m)
Toro Rapids; Rio San Juan. Nicaragua (1102'N.
8428'W. Gardner. 1973)
Didelphis marsupialis
Totaitu; Santa Cruz. Bolivia (16WS. 6230'W.
Anderson. 1997)
Didelphis marsupialis
Philander opossum
Totness; Coronie. Suriname (0553'N, 5619'W.
Geonet)
Didelphis marsupialis
Philander opossum
Totora. Tinkusiri, 17 km E; Cochabamba. Bolivia
(1745'S, 6502'W. Anderson. 1997)
Thylamys venustus
Totoro; Cauca. Colombia (0230'N, 7624'W.
Paynter. 1997)
Didelphis marsupialis, 2500 m
Towashing [to washing'?] Stream. Glycon Swamp:
Bolivar. Venezuela (Glycon Swamp = ca.
0512'N, 6044'W. Paynter. 1982)
Didelphis albiventris, 5400 ft (= 1680 m)
Trapicheiro; Rio 
