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BLM    LIBRARY 


0.  97-12       IDAHO  BUREAU  OF  LAND  MANAGEMENT 


July  1997 


88055390 


QL 

84.2 

.L352 

no. 

97-12 


Breeding  Bird  Survey  of  Old  Growth/Serai, 
Prescribed  Burn,  and  Clearcut  Stands  of 

Western  Juniper 


by 

Golden  Eagle  Audubon  Society 


ru 

BREEDING  BIRD  SURVEY  OF 
OLD-GROWTH/SERAL,  PRESCRIBED  BURN,  AND  CLEARCUT  STANDS  OF 

WESTERN  JUNIPER 

Golden  Eagle  Audubon  Society 
6  January  1 997 


Cooperative  Challenge  Cost  Share  Project 


A 


Bureau  of  Land  Management 

Boise  District  Office  ,\^ 

3948  Development  Ave 

Boise,  ID  83705  *n>\ 


-\-N 


and 


Golden  Eagle  Audubon  Society 

P.O.  Box  8261 

Boise,  ID  83707 


Abstract:  A  survey  of  avian  use  of  western  juniper  stands  subjected  to  three  management 
strategies  was  conducted  during  May  and  June  1 993  in  the  Owyhee  and  Bruneau  Resource  Areas, 
Owyhee  County,  Idaho.  The  three  managment  strategies  included:    untreated  (old-growth/seral), 
prescribed  burns,  and  clearcuts.  Two-hundred  and  ten  eight-minute  samples  were  conducted  at 
70  points.  Fourty-one  species  and  1755  birds  were  observed.  Old-growth/seral  plots  averaged 
4.15  species  and  6.44  birds  per  plot,  prescribed  burn  plots  averaged  1.89  species  and  2.87  birds 
per  plot,  and  clearcut  plots  averaged  1.74  species  and  2.49  birds  per  plot.  Gray  flycatcher, 
mountain  bluebird,  American  robin,  chipping  sparrow,  dark-eyed  junco,  and  Cassin's  finch  were 
most  abundant  in  old-growth/seral  stands.  Vesper  sparrow  were  most  abundant  in  prescribed 
burn  stands.  Tree  swallow,  violet  green  swallow,  house  wren,  green-tailed  towhee,  Brewer's 
sparrow,  and  Brewer's  blackbird  were  most  abundant  in  clearcut  areas. 


Introduction 

Western  juniper  (Juniperus  occidentalism  stands  were  historically  found  in  rock  outcrops,  along 
streams,  and  on  scablands  (Eddleman  1984).  Distribution  was  limited  by  seed  dispersal  and 
periodic  fires.  During  the  past  150  years,  western  juniper  has  expanded  its  range  into  adjacent 
grass  and  shrublands  as  a  result  of  fire  suppression,  overgrazing,  and  climatic  changes  (Burkhardt 
and  Tisdale  1976,  Meeuwig  and  Murray  1978).  Understory  grass  and  forb  productivity  declines 
as  juniper  stands  mature,  thereby  reducing  forage  available  to  grazing  ungulates.  For  this  reason 
juniper  has  been  considered  an  invader  in  some  areas  and  is  targeted  for  removal.  A  variety  of 
removal  strategies  have  been  proposed  including  clearcutting  and  prescribed  burning. 

Several  studies  have  shown  the  importance  of  pinyon-juniper  stands  to  avian  species  (Balda  and 
Masters  1980,  Sedgewick  1987).  Few  studies  have  been  conducted  to  determine  avian  use  of 
western  juniper  stands.  Western  juniper  provided  perching  and  nesting  sites  for  at  least  27  species 
of  birds  (Maser  and  Gashwiler  1978).  Juniper  berries  were  an  important  food  source  for 
migratory  and  winter  resident  birds  such  as  robins  and  Townsend  solitaires  (Eddleman  1984). 

Overstory  removal  in  pinyon-juniper  stands  may  result  in  significant  declines  in  avian  use 
(CMeara  et  al.  1981,  Sedgewick  and  Ryder  1987).  The  objective  of  this  study  was  to  inventory 
avian  use  during  May  and  June  in  prescribed  burn,  clearcut,  and  old-growth/seral  stands  of 
western  juniper  in  southwest  Idaho. 


Methods 

Survey  points  were  located  within  a  three-mile  buffer  along  the  Mud  Flat  road  between  Juniper 
Mountain  and  the  Mud  Flat  Guard  Station,  Owyhee  County,  Idaho  (Fig.  1).  Three  treatment 
types  were  surveyed:  old-growth/seral  (OG/S)  (25  points,  5  transects),  prescribed  burn  (PB)  (26 
points,  3  transects),  and  clearcut  (CC)  (19  points,  3  transects)  stands.  Survey  points  were  located 
approximately  250  m  apart  and  125  m  from  ecotones. 

Habitat  Parameters 

Live  tree  stems  were  counted  in  a  fixed  radius  (1 1 .3  m  for  moderate  and  high  density  stands  or  25 
m  for  low  density  stands)  circle  centered  on  the  survey  point.  Stems  were  recorded  by  diameter 
breast  height  (dbh)  classes  (seedling,  <5  cm,  5-10  cm,  10.1-20  cm,  20.1-40  cm,  >40  cm). 

Snags  and  downed  logs  were  counted  in  a  fixed  radius  (1 1.3  m  or  25  m)  circle  centered  on  the 
survey  point.  Snags  and  logs  were  classified  by  decay  class  (undecayed,  no  bark,  moderate  decay, 
very  decayed)  and  size  (1-10  cm,  10.1-20  cm,  >20  cm). 

Avian  Population  Survey 

The  point  count  method  was  used  to  determine  avian  species  use.  An  attempt  was  made  to 
sample  each  point  three  times  between  1  May  1992  and  30  June  1993.  The  same  observer 


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conducted  all  avian  surveys.  Surveys  were  conducted  only  when  climatic  conditions  Thigh  wind, 
rain)  did  not  affect  detectabilhy.  Surveys  began  at  sunrise  and  were  terminated  at  1 100.  Each 
point  was  surveyed  for  eight  minutes  per  sample.  Observations  were  recorded  for  0-50  m,  51-125 
m,  and  >125  m  distances  from  plot  center.  Data  recorded  included:  species,  activity,  number  of 
birds,  distance  buffer  from  point  (0-50  m,  51-125  m,  >125  m),  and  quadrant  (NE,  SE,  SW,  NW). 
Incidental  observations  were  recorded  separately. 


Avian  Population  Data  Analyses 

Analyses  were  based  on  birds  observed  within  50  m  of  observation  points  unless  otherwise  noted. 
The  mean  number  of  species  and  birds  observed  was  determined  for  sample  periods  and  for  all 
periods  combined  by  transect  and  treatment  type.  Observations  of  unidentified  birds  were 
included  in  determining  abundance.  Observations  of  identified  species  were  used  to  calculate 
diversity.  Differences  in  abundance  between  treatments  and  sampling  periods  were  determined 
using  ANOVA  (McGarigal  and  McComb  1992).  Similarity  in  bird  species  composition  between 
treatments  was  determined  using  Sorenson's  Index  (Mueller-Dombois  and  Ellenberg  1974): 

Sorenson's  Index  =  2  W  x  1 00/(a+b) 

where  W  was  the  number  of  species  shared  between  the  two  treatments  being 
compared 

a  was  the  number  of  species  in  treatment  A 
b  was  the  number  of  species  in  treatment  B 

Rarefaction  was  used  to  determine  expected  species  diversity  in  different  treatments  based  on 
number  of  birds  observed  (James  and  Rathbun  1981). 


Results 

Climate  and  Habitat  Parameters 

Precipitation  for  the  period  October- July  measured  at  Silver  City  was  44.5  cm  in  1992  and  60.3 
cm  in  1993.  Live  juniper  densities  were  3  times  greater  in  OG/S  stands  than  PB  or  CC  treatments 
for  >  20  cm  dbh  classes  (Fig.  2).  Juniper  snag  and  log  densities  were  greatest  in  PB  and  CC  plots 
(Fig.  2).  Fire  removed  live  mountain  mahogany  in  PB  treatments;  however,  mahogany  densities 
were  also  reduced  in  mature  juniper  stands  (Fig.  3). 

Avian  Populations 

Two-hundred  and  ten  separate  eight-minute  bird  surveys  were  conducted  between  2  May  and  28 
June  1993.  Fourty-one  species  and  1755  birds  (including  154  birds  of  unidentified  species)  were 
observed  at  all  distances  for  all  plots  combined  (Appendix  A).  Thirty-five  species  and  859  birds 
(including  84  birds  of  unidentified  species)  were  observed  within  50  m  of  all  points  (Table  1). 


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Table  I.  Number  of  birds  by  species  seen  within  50  m  of  sample  points  in  old-growth/seral  (OGS), 
prescribed  burn  (PB),  and  clearcut  (CC)  stands  sampled  2  May  through  28  June  1993,  Owyhee 
Count,  Idaho.  Season  of  use,  habitat,  and  nest  location  are  from  Saab  and  Groves  (1992). 


Species 

w/in  50  m 

Season 

Habitat 

Nest  Loc. 

OG/S 

PB 

CC 

Turkey  vulture 

TUVU 

1 

Red-tailed  hawk 

RTHA 

3 

3 

R* 

R,SS,C,W 

D,C 

American  kestrel 

AMKE 

3 

1 

R* 

SS,R,A 

Sn,Cl 

Common  snipe 

COSN 

3 

1 

Mourning  dove 

MODO 

3 

1 

Calliope  hummingbird 

CAHU 

1 

1 

B* 

R,C,W 

D/C,S 

Unknown  hummingbird 

UNHU 

2 

2 

1 

Red-naped  sapsucker 

RNSA 

1 

Northern  flicker 

NOFL 

9 

4 

1 

R 

Downy  woodpecker 

DO  WO 

1 

R 

Gray  flycatcher 

GRTL 

68 

37 

15 

B* 

Dusky  flycatcher 

DUFL 

2 

Unknown  flycatcher 

UNFL 

1 

2 

Tree  swallow 

TRES 

10 

Violet-green  swallow 

VGSW 

2 

8 

B* 

W,C,R 

Sn,Cl,T 

Common  raven 

CORA 

16 

1 

2 

R 

Mountain  chickadee 

MOCH 

7 

4 

R 

Red-breasted  nuthatch 

RBNU 

1 

House  wren 

HOWR 

9 

10 

13 

B* 

R,W,C 

D,Sn 

Rock  WTen 

ROWR 

3 

3 

1 

B* 

SS 

G.Cr 

Ruby-crowned  kinglet 


RCKI 


Species 


w/in  50  m 


Season 


Habitat 


Nest  Loc. 


OG/S         PB 


CC 


Mountain  bluebird  MOBL 

Hermit  thrush  HETH 

American  robin  AMRO 

Yellow-rumped  warbler  AUWA 
Black- throated  gray  warbler        BTYW 

Unknown  warbler  UNWA 

Green-tailed  towhee  GTTO 

Vesper  sparrow  VESP 

Chipping  sparrow  CHSP 

Brewer's  sparrow  BRSP 

Dark-eyed  junco  DEJU 


44  32 

10  1 

30  6 

4  3 

3 

1 


7 
5 

70 
1 

35 


26 

32 

5 

8 


12 


11 
13 
10 
12 


B* 
B* 
R* 

B* 
B* 


B* 


ss,w,c 

Sn,T 

c,w 

C,W,R 

D/C.S 

C,R 

D/C 

w 

D/C 

SS,W,R 

S,G 

B* 

B* 

B* 

R* 

Red-winged  blackbird  RWBL 

Western  meadowlark  WEME 

Brewer's  blackbird  BRBL 

Brown-headed  cowbird  BHCO 

Cassin's  finch  CAFI 

Pine  siskin  PISI 


R* 


3 
21 

53 


2 
2 
1 

15 

2 


WeJIA 

C,W,SS 
C,W 


R,S 

D,S 
T 


Unknown 


UNKN 


58 


20 


Totals 


483       234         142 


Identified  Species 
Unknown 


30  23 

4  2 


22 
3 


Season  -  R=resident,  B=breeding,  *=neotropical  migrant 

Habitat  Association  -  C=coniferous  and  mixed  coniferous/deciduous,  W=woodland,  aspen,  mountain 

mahogany,  juniper,  R=riparian,  SS=sagebrush,  grassland,  We=wetland 
Nest  Location  -  C=coniferous  tree,  D=deciduous  tree,  C/D=coniferous  or  deciduous  tree,  Cl=cliff, 

Cr=crevice,  G=ground,  S=shrub,  Sn=snag,  T=tree 


Thirty-two  (18  obligate,  14  facultative)  species  of  neotropical  migrants  were  observed  overall, 
with  25  (17  obligate,  8  facultative)  species  occurring  within  50  m  of  points. 

Gray  flycatcher,  mountain  bluebird,  American  robin,  chipping  sparrow,  dark-eyed  junco,  brown- 
headed  cowbird,  and  Cassin's  finch  made  up  66%  of  observations  in  OG/S  plots  (Table  1).  Gray 
flycatcher,  mountain  bluebird,  vesper  sparrow,  and  chipping  sparrow  made  up  54%  of 
observations  in  PB  plots.  Gray  flycatcher,  house  wren,  mountain  bluebird,  green-tailed  towhee, 
vesper  sparrow,  chipping  sparrow,  and  Brewer's  sparrow  made  up  73%  of  observations  in  CC 
plots. 

Species  diversity  and  abundance  were  greatest  in  OG/S  plots  and  lowest  in  CC  plots  (Table  2). 
Abundance  increased  between  the  first  sample  and  the  last  sample  in  OGS  plots,  whereas  diversity 
and  abundance  were  greatest  in  sample  2  for  PB  and  CC  plots  (Appendix  B).  Species  diversity 
and  abundance  were  not  different  between  OG/S  and  PB  plots;  however,  diversity  and  abundance 
were  lower  (r2  =  0.239-0.445,  P  <  0.001)  in  CC  plots  than  OG/S  and  PB  plots  for  each  sampling 
period. 

The  Sam  Noble  PB  plot  had  consistently  lower  species  diversity  and  abundance  than  all  other 
plots  (r2  =  0.513  -  0.638,  P  <  0.001).  CC  plots  generally  had  lower  species  diversity  than  OG/S 
(P  <  0.001);  however,  there  was  no  consistent  pattern  for  diversity  and  abundance  between 
transects  and  samples. 

Similarity  indices  for  species  seen  within  50  m  of  the  point  were  greatest  for  the  OG/S-PB 
comparison  and  lowest  for  the  PB-CC  comparison  (Table  3).  Similarity  indices  between  sample 
periods  were  greatest  between  sample  periods  1  (2-16  May)  and  2  (17-29  May)  and  lowest 
between  sample  periods  1  and  3  (1 1-28  June)  (Table  4).  Rarefaction  predicted  similar  species 
diversity  in  OG/S,  PB,  and  CC  areas  (Figure  4).  Twenty-two  species  were  predicted  in  OG/S,  20 
species  were  predicted  in  PB  and  21  species  were  predicted  in  CC  areas  when  abundance  was 
held  constant  between  the  three  areas. 


Yearly  Variations 

Seven  species  were  observed  in  1993  that  were  not  observed  in  1992  (Canada  goose,  prairie 
falcon,  dusky  flycatcher,  red-breasted  nuthatch,  yellow  warbler,  black-headed  grosbeak,  and 
Brewer's  blackbird)  (Appendix  A).  Brewer's  blackbird  and  Canada  goose  were  the  only  species 
with  more  than  2  observations.  Eighteen  species  were  observed  in  1992  that  were  not  observed 
in  1993  (mallard,  sharp-shinned  hawk,  Cooper's  hawk,  goshawk,  great-horned  owl,  cordillerian 
flycatcher,  olive-sided  flycatcher,  western  wood  pewee,  American  crow,  bushtit,  Townsend's 
solitaire,  sage  thrasher,  red-eyed  vireo,  warbling  vireo,  lazuli  bunting,  lark  sparrow,  yellow- 
headed  blackbird,  and  western  tanager).  Great-horned  owl,  downy  woodpecker,  common  bushtit, 
and  Townsend's  solitaire  were  the  only  species  with  more  than  2  observations. 

Changes  in  relative  abundance  (percentage  of  total  known  observations  for  a  given  species) 
between  1992  and  1993  were  variable.  Northern  flicker  numbers  decreased  in  PB  plots  (5.1%- 


Table  2.    Average  number  of  bird  species  and  individuals  observed  within  50  m  of  points  by 
treatment  for  sampling  2  May  through  28  June  1993,  Owyhee  Co.,  Idaho.  Average 
number  of  species  includes  unique  unknowns  (ie.  unidentified  flycatcher  species  in  a 
plot  with  no  identified  flycatchers). 


Treatment 

Transect 
Location 

N 

#  species 

#  birds 

mean 

SD 

mean 

SD 

Old-growth/ 

Stoneman  Cr. 

18 

3.39 

0.52 

4.00 

0.63 

Serai 

Rays  Res.  East 

24 

4.29 

1.92 

6.58 

3.32 

Pleasant  Valley 

9 

4.11 

1.05 

8.22 

2.44 

Juniper  Mtn 

9 

4.78 

2.05 

6.89 

3.89 

Rays  Res.  West 

12 

4.50 

1.45 

6.67 

2.67 

Combined 

75 

4.15 

1.57 

6.44 

2.91 

Prescribed 

Sam  Noble 

36 

0.89 

0.67 

1.22 

1.20 

Burn 

Pleasant  Valley 

24 

3.17 

1.31 

5.08 

2.32 

Cottonwood  Cr. 

18 

2.17 

1.54 

3.22 

2.32 

Combined 

78 

1.89 

1.54 

2.87 

2.50 

Clear  Cut 

Pleasant  Valley 

21 

1.62 

1.32 

2.10 

1.73 

Juniper  Mtn 

21 

1.71 

1.06 

2.05 

1.24 

Hanley  Cabin 

15 

1.93 

0.88 

3.67 

3.85 

Combined 

67 

1.74 

1.11 

2.49 

2.41 

Table  3,    Similarity  indices  (based  on  Sorenson's  index)  for  avian  species  observed  in  old- 

growth/seral  (OGS),  prescribed  burn  (PB),  and  clearcut  (CC)  stands  during  May-June, 
1993  in  Owyhee  County,  Idaho. 


OGS 

PB 

#  spp.  in 
common 

OGS 

CC 

#  spp.  in 
common 

PB 

CC 

#  spp.  in 
common 

50  m 

% 

30 

23 
71.7 

19 

30 

22 
69.2 

18 

23 

22 
66.7 

15 

similarity 


125  m 

32 

30 

% 

77.4 

similarity 

overall 

33 

31 

% 

78.1 

24 


32 


25 


33 


26 
79.3 


31 
78.1 


23 


30 


25 


31 


26 
78.6 


31 
80.6 


22 


25 


similarity 


Table  4.    Similarity  indices  (based  on  Sorenson's  index)  for  avian  species  observed  during  three 

sample  periods  [A  =  2-16  May  1993  (Pleasant  Valley  PB  sample  A  was  conducted  on  30 
May),  B  =  17-29  May  1993  (Pleasant  Valley  PB  sample  B  was  conducted  on  13  June),  C  = 
1 1-28  June  1993]  in  Owyhee  County,  Idaho. 


A 

B 

#  spp.  in 
common 

A 

C 

#  spp.  in 
common 

B 

C 

#  spp.  in 
common 

50  m 

22 

23 

16 

22 

26 

16 

23 

26 

15 

% 

67.9 

52.4 

63.2 

similarity 

10 


1.9%).  Gray  flycatcher  numbers  increased  in  all  treatments  (OG/S  -  10.7%- 16%,  PB  -  10.6%- 
1 7.2,  CC  -  4.9%-l  1%).  Mountain  chickadee  numbers  decreased  in  OG/S  plots  (10.2%-1 .6%). 
House  wren  numbers  decreased  in  CC  plots  (13.4%-9.5%).  Mountain  bluebird  numbers 
increased  in  OG/S  plots  (6%-10.4%)  and  decreased  in  PB  (18.2%- 15%)  and  CC  (19.5%-8.8%) 
plots.  Green-tailed  towhee  numbers  increased  in  CC  plots  (6.1%-8.1%).  Vesper  sparrow 
numbers  decreased  in  CC  plots  (1 5.8%-9.6%).  Chipping  sparrow  numbers  decreased  in  OG/S 
(2 1.4%- 16.5%)  and  CC  (19.5%-7.4%)  plots  and  increased  in  PB  plots  (1 1.4%- 15%).  Dark-eyed 
junco  numbers  increased  in  OG/S  plots  (5.2%-8.2%).  Brown-headed  cowbird  numbers  decreased 
in  PB  plots  (6.4%-0.5%).  Cassin's  finch  numbers  increased  in  OG/S  (6.6%- 12.5)  and  PB  (4.2%- 
7%)  plots. 

Number  of  species  (r2  =  0.31,  F  =  91.1,  P  <  0.001)  and  number  of  birds  seen  (r2  =  0.25,  F  =  67.1, 
P  <  0.001)  were  different  between  treatments  for  1992  and  1993  combined.  However,  they  were 
not  different  between  years  (species:  F  =  2.6,  P  =  0.107;  total:  F  =  2.7,  P  =  0.101).  Number  of 
species  (r2  =  0.44,  F  =  28.8,  P  <  0.001)  and  number  of  birds  (r2  =  0.37,  F  =  20.7,  P  <  0.001) 
were  different  between  transects  for  1 992  and  1 993  combined.  OG/S  and  the  Pleasant  Valley  PB 
transect  had  greater  numbers  of  species  and  birds  than  the  other  PB  and  CC  transects. 

Similarity  indices  were  3.5%  (PB-CC),  7.4%  (OG/S-PB),  and  16.8%  (OG/S-CC)  greater  in  1993 
than  in  1992  for  the  50  m  plots.  Indices  were  even  greater  when  overall  numbers  were  compared. 

Discussion 

Greater  avian  abundance  and  species  diversity  in  OG/S  was  probably  best  explained  by  greater 
structural  diversity.  Understory  components  were  similar  between  treatments,  however,  cover 
was  greater  in  OG/S  than  PB  or  CC  treatments  (McCoy  1993).  While  OG/S  stands  had  no 
juniper  snags  and  few  logs,  they  had  greater  diversity  and  abundance  of  live  junipers.  Vegetation 
composition  and  structure  in  the  Pleasant  Valley  PB  closely  resembled  OG/S  stands  resulting  in 
similar  avian  abundance  and  diversity  (Appendix  C).  PB  treatments  with  a  >  20%  kill  rate  had 
relatively  open  understories  that  provided  less  cover  for  ground  foraging  species  compared  to  CC 
sites  with  slash. 

Weather  may  have  influenced  changes  in  species  composition  and  abundance  between  years. 
Greater  precipitation  in  1993  than  1992  may  have  had  variable  effects  on  avian  populations. 
Increased  precipitation  could  result  in  greater  insect  and  seed  production  and  consequently 
greater  avian  productivity,  however,  extended  cool,  wet  weather  may  have  caused  some  nest 
failures.  Generalist  species  may  have  been  favored  resulting  in  lower  diversity  and  greater 
similarity  of  avian  species  between  treatments. 


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Figure  4.  Rarefa 
and  clearcut  stan 

12 


Literature  Cited 

Balda,  R.P.  and  N.  Masters.   1980.  Avian  communities  in  the  pinyon-juniper  woodland:  a 
descriptive  analysis.  Pages  146-169  in  U.S.  For.  Serv.  Gen.  Tech.  Rep.  INT-86. 

Burkhardt,  J.W.  and  E.  W.  Tisdale.   1976.  Causes  of  juniper  invasion  in  southwestern  Idaho. 
Ecology  57(3):264-270. 

Eddleman,  L.E.   1984.  Ecological  studies  on  wester  juniper  in  central  Oregon.  Pages  29-35  in 
Proceedings  -  western  juniper  management  short  course  (T.E.  Bedell,  editor).  Oregon 
State  University,  Corvallis,  Oregon. 

James,  F.C.  and  S.  Rathbun.  1981.  Rarefaction,  relative  abundance,  and  diversity  of  avian 
communities.  Auk  98:785-800. 

Maser  C.  and  J.S.  Gashwiler.  1978.  Interrelationships  of  wildlife  and  western  juniper.  Pages  37- 
82  in  Proceedings  of  the  western  juniper  ecology  and  management  workshop  (R.E.  Martin, 
J.E.  Dealy,  D.L.  Caraher,  editors).  U.S.  For.  Serv.  Gen.  Tech.  Rep.  PNW-74.  Pac. 
Northwest  For.  and  Ran.  Exp.  Stn.,  Portland,  Oregon. 

McCoy,  M.   1993.  Breeding  bird  survey  of  clearcut,  prescribed  burn,  and  old-growth/seral  stands 
of  western  juniper.  CCS  report  for  Boise  District,  BLM.   12  pp. 

McGarigal,  K.  and  W.C.  McComb.  1992.  Streamside  versus  upslope  breeding  bird  communities 
in  the  central  Oregon  coast  range.  J.  Wildl.  Manage.  56:10-23. 

Meeuwig,  R.O.  and  R.B  Murray.   1978.  Current  research  on  pinyon-juniper  in  the  Great  Basin. 
Pages  97-103  in  Proceedings  of  the  western  juniper  ecology  and  management  workshop 
(R.E.  Martin,  J.E.  Dealy,  D.L.  Caraher,  editors).  U.S.  For.  Serv.  Gen.  Tech.  Rep.  PNW- 
74.  Pac.  Northwest  For.  and  Ran.  Exp.  Stn.,  Portland,  Oregon. 

Mueller-Dombois,  D.  and  H.  Ellenberg.   1974.  Aims  and  methods  of  vegetation  ecology.  John 
Wiley  and  Sons,  New  York. 

O'Meara,  T.E.,  J.B.  Haufler,  L.H.  Stelter,  J.G.  Nagy.   1981.  Nongame  wildlife  responses  to 
chaining  of  pinyon-juniper  woodland.  J.  Wildl.  Manage.  45:381-389. 

Saab,  V.A.  and  C.R.  Groves.   1992.  Idaho's  migratory  landbirds.  Idaho  Dept.  of  Fish  and  Game. 
Nongame  Leaflet  No.  10. 

Sedgewick,  J.A.   1987.  Avian  habitat  relationships  in  a  pinyon-juniper  woodland.  Wilson  Bull. 
99:413-431. 


13 


and  R.A.  Ryder.  1987.  Effects  of  chaining  pinyon-juniper  on  nongame  wildlife.  Pages 


541-551  in  Proceedings  of  the  pinyon-juniper  conference,  Reno,  Nevada  (R.L.  Everett, 
editor).  Int.  Mtn  Res.  Sta.,  Ogden,  Utah. 


14 


Appendix  A.  Numbers  of  birds  observed  during  2  May  through  28  June  1993  at  70  points  in  old- 
growth/seral  (OGS),  prescribed  burn  (PB),  and  clearcut  (CC)  stands  in  Owyhee  County,  Idaho. 


All  distances 

w/in  125 

m. 

w/in  50  m 

Species 

Species 
Code 

OGS 

PB 

CC 

OGS 

PB 

CC 

OGS 

PB       CC 

Canada  goose 

CAGO 

4 

Common  snipe 

COSN 

1 

5 

1 

1 

5 

1 

3         1 

Turkey  vulture 

TUVU 

2 

8 

2 

1 

1 

Red-tailed  hawk 

RTHA 

3 

5 

3 

5 

3 

3 

American  kestrel 

AMKE 

5 

3 

5 

3 

3 

1 

Prairie  falcon 

PRFA 

2 

2 

Mourning  dove 

MODO 

9 

4 

5 

9 

4 

4 

3 

1 

Great-horned  owl 

GHOW 

2 

Common  poorwill 

COPW 

1 

1 

Common  nighthawk 

CONI 

1 

1 

Calliope  hummingbird 

CAHU 

1 

1 

1 

1 

1         1 

Unknown  hummingbird 

UNHU 

3 

2 

1 

3 

2 

1 

2 

2         1 

Northern  flicker 

NOFL 

20 

25 

16 

19 

25 

10 

9 

4         1 

Red-naped  sapsucker 

RNSA 

2 

2 

2 

1 

1 

Downy  woodpecker 

DOWO 

1 

1 

1 

Gray  flycatcher 

GRFL 

84 

64 

32 

84 

64 

32 

68 

37       15 

Dusky  flycatcher 

DUFL 

2 

2 

2 

Unknown  flycatcher 

UNFL 

1 

2 

1 

2 

1 

2 

Tree  swallow 

TRES 

11 

11 

10 

Violet-green  swallow 

VGSW 

2 

10 

2 

10 

2 

8 

Black-billed  magpie 

BBMA 

1 

1 

Common  raven 

CORA 

111 

14 

20 

103 

4 

6 

16 

1         2 

Mountain  chickadee 

MOCH 

10 

2 

7 

10 

2 

7 

7 

4 

Red-breasted  nuthatch 

RBNU 

1 

1 

1 

House  wren 

HOWR 

13 

15 

25 

13 

14 

22 

9 

10       13 

Species 

Species 
Code 

All  distances 
OGS        PB        CC 

w/in  125 
OGS       PB 

m 
CC 

w/in  50  m 

OGS        PB 

CC 

Rock  wren 

ROWR 

7 

4 

5 

7 

4 

5 

3 

3 

1 

Ruby-crowned  kinglet 

RCKI 

2 

2 

1 

2 

2 

1 

2 

2 

Mountain  bluebird 

MOBL 

68 

65 

30 

68 

64 

30 

44 

32 

12 

Hermit  thrush 

HETH 

38 

1 

8 

36 

1 

4 

10 

1 

American  robin 

AMRO 

59 

24 

25 

59 

23 

22 

.0 

6 

5 

*■ 

Yellow  warbler 

YEWA 

1 

• 

Yellow-rumped  warbler 
Black-throated  gray  warbler 
Unknown  warbler 

AUWA 
BTYW 
UNWA 

5 
4 

1 

3 

2 

5 
4 

1 

3 

4 
3 

1 

3 

Green-tailed  towhee 

GTTO 

8 

2 

21 

8 

2 

21 

7 

11 

Vesper  sparrow 

VESP 

7 

62 

32 

7 

59 

32 

5 

26 

13 

Chipping  sparrow 

CHSP 

103 

56 

16 

103 

56 

16 

70 

32 

10 

Brewer's  sparrow 

BRSP 

5 

47 

49 

5 

46 

49 

1 

5 

12 

Dark-eyed  junco 

DEJU 

49 

11 

1 

49 

11 

35 

8 

Western  meadowlark 

WEME 

5 

21 

1 

10 

2 

Red-winged  blackbird 

RWBL 

3 

3 

3 

3 

3 

3 

3 

2 

Brewer's  blackird 

BRBL 

3 

4 

19 

3 

4 

19 

3 

2 

9 

Brown-headed  cowbird 

BHCO 

25 

2 

3 

25 

2 

2 

21 

1 

1 

Pine  siskin 

PISI 

2 

4 

2 

4 

2 

4 

> 

Cassin's  finch 

CAFI 

85 

23 

8 

85 

23 

8 

53 

15 

2 

* 

Unknown 

UNKN 

85 

46 

23 

85 

46 

23 

58 

20 

6 

Totals 

Identified  species 
Unknown 


835      523      397        818      49      344        483      234       14 

33        31         31  32       30        26  30        23       22 

423  423  423 


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Appendix  C.  Transect  descriptions  for  avian  population  sampling  in  old-growth/seraL  prescribed 
burn,  and  clear  cut  stands  of  western  juniper  in  Owyhee  County,  Idaho. 

Old-growth/Seral 

Stoneman  Creek  (6  points)  -  This  transect  was  located  on  both  sides  of  the  Mud  Flat  road  on  a 
saddle  between  two  drainages.  One  point  on  the  west  side  of  the  road  was  in  a  moderately  older- 
aged  stand  with  a  few  trees  removed  during  prior  logging  activity.  The  other  point  on  the  west 
side  of  the  road  was  located  in  a  younger  (estimated  <45  years  old)  serai  stand.  Points  on  the  east 
side  of  the  road  were  located  in  primarily  old-growth  stands  ranging  from  fairly  shallow  soils  with 
rocky  outcrops  to  a  somewhat  deeper  soil  with  moderate-aged  (about  100-125  years)  trees. 
Understory  cover  was  greatest  in  areas  with  deeper  soils  and  younger  or  more  open  stands  of 
juniper.  The  nearest  water  was  approximately  one-mile  away. 

Rays  Reservoir  East  (8  points)  -  This  transect  was  divided  by  a  series  of  low  rocky  ridges  and  dry 
to  semi- wet  drainages.  Old-growth  juniper  dominated  the  ridges  and  upper  slopes  with  younger 
(estimated  <  75  years  old)  trees  on  lower  slopes  and  into  meadows.  Sample  points  were  generally 
located  in  moderate-aged  (50-150  years  old)  stands.  There  was  a  spring  and  reservoir  on  the 
transect.  A  clear  cut  was  initiated  on  the  southeast  border  of  the  transect  in  1994.  One  point  was 
clear  cut  after  sampling  was  completed. 

Rays  Reservoir  West  (4  points)  -  This  transect  was  located  on  two  ridges  with  old-growth  stands 
of  juniper,  bisected  by  a  small  semi-wet  meadow.  A  spring  also  existed  on  the  northwest  side  of 
the  transect. 

Pleasant  Valley  (3  points)  -  This  transect  was  located  adjacent  to  the  Pleasant  Valley  clear  cut 
transect.  The  transect  was  dominated  by  trees  <  100  years  old  with  scattered  older  (>  150  years 
old)  trees.  Pleasant  Valley  Creek  or  the  spring  on  the  Rays  Reservoir  West  transect  were  the 
closest  water  sources. 

Juniper  Mountain  (3  points)  -  This  transect  was  dominated  by  moderate-aged  (50-80  years  old) 
trees  with  old-growth  trees  on  rocky  outcrops.  This  transect  had  the  greatest  canopy  cover  and 
sparsest  understory.  It  was  located  adjacent  to  a  perennial  stream. 

Prescribed  Burn 

Sam  Noble  (12  points)  -  This  site  was  burned  in  1985  with  approximately  40-50%  kill  of  junipers. 
The  stand  was  fairly  open  and  consisted  of  younger  (<60  years  old)  trees  prior  to  burning. 
Pockets  of  live  junipers  were  widely  scattered  throughout  the  transect.  Most  of  the  dead  junipers 
were  still  standing  during  the  sample  period.  Most  points  were  located  in  fairly  open  areas  with  < 
half  containing  some  live  junipers  within  50  m.  Water  was  available  within  1/4  to  1  mile  to  the 
west  of  points  in  the  transect. 

Pleasant  Valley  (8  points)  -  This  site  was  burned  in  1985  with  <  20%  kill  of  junipers.  The  site 
contained  a  mixture  of  old  and  moderate  aged  trees.  Burned  areas  were  generally  <  50  m  in 


18 


diameter.  This  transect  was  very  similar  in  structure  to  old-growth/seral  stands.  A  reservoir  was 
located  within  Vz  mile  of  any  point. 

Cottonwood  Creek  (6  points)  -  The  site  was  burned  in  1985.  The  western  portion  (2  points) 
experienced  a  <25%  kill  and  was  adjacent  to  a  perennial  stream.  It  had  a  mixture  of  old  and 
moderate  aged  live  trees.  The  eastern  portion  (4  points)  experienced  a  45%  kill  and  was  located 
within  Vi  mile  of  water.  This  portion  had  been  dominated  by  trees  <  65  years  old  prior  to 
treatment.  Pockets  of  live  juniper  remained  within  125  m  of  each  point.  This  site  also  included 
on  old-growth/seral  point. 

Clear  Cut 

Pleasant  Valley  (7  points)  -  This  site  was  cut  in  1986-91  The  site  had  fairly  shallow  soils  and  was 
dominated  by  old-growth  trees  prior  to  cutting.  Slash  up  to  1  m  deep  was  common  on  the  site. 
Old-growth  stands  ringed  the  site  and  extended  into  the  cut  area.  Pleasant  Valley  Creek  and  the 
spring  in  Rays  Reservoir  West  were  the  nearest  sources  of  water. 

Hanley  Cabin  (5  points)  -  This  site  was  cut  in  1990-92.  Prior  to  cutting  there  was  a  mixture  of 
young  to  old  trees  in  moderately  deep  soils  interspersed  with  semi-wet  to  wet  stringer  meadows. 
There  was  a  small  (<  5  acres)  stand  of  aspen  around  a  spring  on  the  perimeter  of  the  cut.  Slash 
up  to  1  m  deep  was  common  on  the  site. 

Juniper  Mountain  (7  points)  -  This  site  was  burned  in  1981  and  cut  in  1982-83.  Prior  to 
treatment,  there  was  a  mixture  of  moderate  to  old  (>  50  years  old)  junipers.  There  was  a  complex 
interspersion  of  live  trees,  snags,  and  open  spaces  after  treatment.  Some  pockets  of  partially  or 
un-burned  slash  were  present. 


19 


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QL  84.2  .L352  no.  97-12 

88055390 
Breeding  bird  survey  of  old 
growth/seral,  prescribed 


i_-'_ivi  u;u>i 


BLDG50.ST-150A 
DENVER  FEDERAL  CENTER 

P.O.  BOX  25047 
DENVER,  COLORADO  80225 


Bureau  of  Land  Management 

Idaho  State  Office 
1387  S.  Vinnell  Way 
Boise,  Idaho  83709 

BLM/ID/PT-97/015+1150