Skip to main content

Full text of "Bulletins of American paleontology"

See other formats


y nds ny) 


ois ae 


“a Neca 
0 evan 


Drives 


ry 
yt ts tt 


My 
mA oes 
ve Cocaine. 

aes 
< 


4 emis) ; 
we ate Caetiane seh 


wera: 
Poa toer 


mete. 
Ma Betene a UN alan 
POLE POL tener 
Naan Sean 
stayin alan AA ee 


mi Reet 
ata ae gs 


ahs 
wiry! 
. 


bla En 
Oh Bae 
at TY Fase 
Vas iy, 
ay w 


mR 
es 


= * 
SE ae 
Beth 


c * 


MSPS wee 
waheue 


per) 
SoRiyeat se 


Sey 
Pop ate - 
ion as ee a 
eae oly 
tae “ther 4 tread 
Bia 
ie f : nites su 
; FF ew ay 
Sine es 
Faw curate 
‘ 
r Hoe e 
: | Wa ep 
me Pete 
ve 
ont, 


HARVARD UNIVERSITY 
e 
Library of the 
Museum of 


Comparative Zoology 


a ee ee eee 
VOLUMES 82 & 83, NUMBER 315 DECEMBER 23, 1982 
a a ee ae eae re 


Devonian Brachiopods 
of 


New Mexico 


by 
G. Arthur Cooper 
and 
J. Thomas Dutro, Jr. 


PRI 


5 On 


Paleontological Research Institution RITTVEESETY. 
1259 Trumansburg Road year 
Ithaca, New York, 14850 U.S.A. 


Library of Congress Card Number: 82-62780 


Printed in the United States of America 
Allen Press, Inc. 
Lawrence, KS 66044 U.S.A. 


CONTENTS 


ADSUTRGE “os Golco SSPE RO Be APIS ECS) Onno Hee eta n RM RE RRS Ieee 5 
MALT OAUIG EN OOM ameapevene ist fevcyeneyare rane enstceneter tt sverer axes avovs louse vase cavstovaes 6 
AG TIOVRGERIGHIS saw haas obese AOR one a Soo tae GOmrnoee 7 
BI EVIOUSBW, OL Keener area eres setae rece exse tye ol Les teres, Sc casievseshou cloves 8 
(OWMALS [OMT ONT taco cree GAG coe Core oO OOo SEC OT rte RG 10 
iva GaDMROLMAtONE mite nyer-cactets te ciciete-sitieieiueieseicieis se.e ce eens 15 
Gontaderowkormati ones seer rcs eee ee ore criteria oer 19 
borouUrhPoOOdNROLMAtLOM! Gerace rclsyene-wcreneietesckec tens Te euassiciaaoele 20 
Riodest Canyonvrhonmationi retry let acne rt coe eicerice acer. 21 
Percha Formation (Ready Pay Member) .................. 22 
Rercha Kormation (Box Member) 2..-...2---2-<-05--+-es- 23 
Conodont) Biostratigraphy, 4.0... ee eee ee ees ni cine eee 25 
MGS OLO PIC RELISE OG Vip eis icy ode nyo ap eave ve cab ok sens yas WesohaiSesyavsbsuagonseelele 27 
BAC MOpPOdS MOM LOWas ie < «esol elaleverays aielclorsreves 2 creroejer ove ers 30 


Systematic Paleontology 
MATKOMUCHIONY crepe cersievavs sfeyetere.Votareieieictcisiecsr eel sidietra ies ioyaseeiee te 30 
Class Inarticulata 
Order Lingulida 


Family Lingulidae Menke ....................-..5: 32 
Order Acrotretida 
KamilyDiscinidae)Gray) 2.-- 3.24... 0560+ sseesee ene 33 
Family Craniidae Menke ................000000000 34 
Class Articulata 
Order Orthida 
Family Skenidiidae Kozlowski ..................... 34 
Order Enteletidae 
Family Enteletidae Waagen ........................ 35 
Family Onniellidae Opik ...................0..2-0. 38 
Family Rhipidomellidae Schuchert ................. 4] 
Family Tropidoleptidae Schuchert .................. 43 
Order Strophomenida 
Family Stropheodontidae Caster ................... 44 
Ramilyauncentaims sor ar pvrjmisicleraisteisrascre crocncerenre eles 53 
Family Schuchertellidae Williams .................. 56 
Family Chonetidae Bronn ......................... 57 
Family Strophalosiidae Schuchert .................. 60 


Family Productellidae Schuchert ................... 

Family Leioproductidae Muir-Wood and Cooper ..... 

Family Overtoniidae Muir-Wood and Cooper ........ 
Order Pentamerida 

Family Pentameridae M’Coy 
Order Rynchonellida 

Family Trigonirhynchiidae McLaren ................ 


Family Uncinulidae Rzhonsnitskaya ................ 
Family Pugnacidae Rzhonsnitskaya ................ 
Family Camarotoechiidae Schuchert ................ 
Family Yunnanellidae Rzhonsnitskaya .............. 
Order Spiriferida 
EamilysAtrypidae.Gilll Sassen eee eon erence re 
Family Athyrididae M’Coy ......................0 
Family Ambocoeliidae George ..................... 
Family Mucrospiriferidae Pitrat .................... 
Family Spinocyrtiidae Ivanova ..................... 
Family Thomasariidae new family .................. 
Family Cyrtospiriferidae Termier and Termier ....... 
Family Reticulariidae Waagen 
Family Elythidae Fredericks ....................... 


Family Martiniidae Waagen ....................... 
Family Cyrtinidae Fredericks ....................4. 
Order Terebratulida 
Family Cranaenidae Cloud ........................ 
Family Heterelasminidae Licharew ................. 
Clarification of Taxonomic and Stratigraphic Problems 
raised by Stainbrook’s 1947 Paper on the Percha Brachi- 
(0) 010) KERN ae ie Gonna OO OCALA COD DOT ara abd Soe oO ac 
Appendices 
Ee ocalityalists\(Georraphic) seee ee eaeere eee renee 
2a ocalityelists: (Stratigraphic) meeeraeenieceeieie ee 


3. Locality Descriptions and Faunal Lists 
ReferencesiGited srrojsscreryernotsictoraeener vos Cia eee terete 


Text-figure 


il, 


i) 


LIST OF ILLUSTRATIONS 


Page 

Index map of south-central New Mexico, showing major 
outcrop areas of Devonian rocks ............--.+.5-- 6 

. Map of a part of the Sacramento Mountains, showing 

location of measured sections of Onate Formation, Sly 
Gap Formation, and related strata .................5. 7 

. Map of the San Andres Mountains, showing locations of 
measured sections of Devonian and related strata ..... 9 


. Correlated stratigraphic sections of Devonian rocks, 


Sacramento Mtns., New Mexico .. foldout inside front cover 


. Correlated stratigraphic sections of Devonian rocks, San 


Andres Mtns., New Mexico ...... foldout inside back cover 


. Correlated stratigraphic sections of Devonian rocks at 


selected localities west of the San Andres Mountains, 
New Mexico 


. Correlation diagram of Devonian rocks in New Mex- 


100 ee ear ae Ia aacitinipi ded roicion oie. eon cater nae racnai 1213 


. Correlation of New Mexico Devonian strata with select- 


ed Devonian sequences in North America 


MAlcutoniavangulatamew SPECleSmei)amidernteeiterrreritel= 101 


Table 


LIST OF TABLES 


Page 
. Brachiopods of the Onate Formation ............... 14 
. Brachiopods of the Sly Gap Formation ............. 19 
. Brachiopods of the Contadero Formation ........... 20 
. Brachiopods of the Thoroughgood Formation ........ 21 
. Brachiopods of the Rhodes Canyon Formation ...... 22 
. Brachiopods of the Percha Formation (Ready Pay Mem- 
BEB), ces. fissetesensie-eieuans oxavs sale, «tipo ine ee eee 23 
. Brachiopods of the Percha Formation (Box Member) .. 24 
. Conodont occurrences in the basal Percha Formation, 
southwestern New Mexico, and related strata ....... 28 
Brachiopods of the Independence Shale described in 
ChiSMPAPER rcecsstesreiee ction ee calle eee ae 30 
. Brachiopods from the Cerro Gordo Member and Cedar 
Valley Formation described in this paper ........... 30 


VOLUMES 82 & 83, NUMBER 315 


Devonian Brachiopods 
of 


New Mexico 


by 
G. Arthur Cooper 
and 
J. Thomas Dutro, Jr. 


Paleontological Research Institution yy) aes 
1259 Trumansburg Road iednlonace 
Ithaca, New York, 14850 U.S.A. 5 


PALEONTOLOGICAL RESEARCH INSTITUTION 


Officers 
PRESTIDENTE 250s Rees ee Le oe Oe eR eee Cane BRUCE M. BELL 
WICE-PRESIDENIE” orc cc rate SN a 2 oro WILLIAM A. OLIVER, JR. 
SEGRETAIRW tose cans Guar ris iete hee eRe PS Ea ora eee PHILIP C. WAKELEY 
TPRIEAS WIRIER a tahpa Sate ise ye edge ss See coco eheo eh ashy aetna ROBERT E. TERWILLEGAR 
ASSISTANT SECRETARY AND ASSISTANT TREASURER ............ JOHN L. CISNE 
DIRE GEORG ssh ced corse RO ne ee aE TPES: one ee eae ose Tee PETER R. HOOVER 
LEGAL: COUNSELS sepa pete ee OC: Cin rie ARMAND L. ADAMS 
Trustees 
BrucE M. BELL (to 6/30/84) WILLIAM A. OLIVER, JR. (to 6/30/83) 
RICHARD E. ByRD (to 6/30/83) JOHN POJETA, JR. (to 6/30/85) 
JOHN L. CISNE (to 6/30/85) JAMES E. SORAUF (to 6/30/85) 
LEE B. GIBSON (to 6/30/83) ROBERT E. TERWILLEGAR (to 6/30/84) 
REBECCA S. HARRIS (Life) RAYMOND VAN HOuTtTTE (to 6/30/85) 
PORTER KIER (to 6/30/84) WILLIAM P. S. VENTRESS (to 6/30/84) 
DUANE O. LERoy (to 6/30/84) PHILIP C. WAKELEY (to 6/30/83) 


BULLETINS OF AMERICAN PALEONTOLOGY 
and 


PALEONTOGRAPHICA AMERICANA 
PETERARS NEIOOVERG oss iors eine tk SN tea Han ITA ee EDITOR 


Reviewers for this issue 


ARTHUR J. BOUCOT 
J. G. JOHNSON 
GILBERT KLAPPER 


A list of titles in both series, and available numbers and volumes may be 
had on request. Volumes 1—23 of Bulletins of American Paleontology have been 
reprinted by Kraus Reprint Corporation, Route 100, Millwood, New York 10546 
USA. Volume | of Paleontographica Americana has been reprinted by Johnson 
Reprint Corporation, 111 Fifth Ave., New York, NY 10003 USA. 


Subscriptions to Bulletins of American Paleontology may be started at any 
time, by volume or year. Current price is US $25.00 per volume. Numbers of 
Paleontographica Americana are priced individually, and are invoiced separately 
on request. Purchases for professional use by U.S. citizens are tax-deductible. 


for additional information, write or call: 


Paleontological Research Institution 
1259 Trumansburg Road 
Ithaca, NY 14850 USA 


The Paleontological Research Institution 
acknowledges with special thanks 
the contributions of the following individuals and institutions 


PATRONS 


($1000 or more at the discretion of the contributor) 


ARMAND L. ADAMs (1976) 

James A. ALLEN (1967) 

AMERICAN OIL Company (1976) 
ATLANTIC RICHFIELD COMPANY (1978) 
Miss ETHEL Z. BAILEY (1970) 
CHRISTINA L. BALK (1970) 

Mr. & Mrs. KENNETH E. CASTER (1967) 
CHEVRON O1L ComPAny (1978, 1982) 
Exxon Company (1977 to date) 

Lois S. FOGELSANGER (1966) 

GuLF OIL CORPORATION (1978) 
MERRILL W. Haas (1975) 


REBECCA S. HARRIS (1967) 

RosBert C. HOERLE (1974-77) 

RICHARD I. JOHNSON (1967) 

J. M. MCDONALD FOUNDATION (1972, 1978) 
Mobi O1t CorRPORATION (1977 to date) 
N.Y. State Arts Councit (1970, 1975) 
SAMUEL T. PEEs (1981) 

Rosert A. PoHowsky (1982) 

CASPAR RAPPENECKER (1976) 

Texaco, Inc. (1978, 1982) 

UNION OIL OF CALIFORNIA (1982) 

UNITED STATES STEEL FOUNDATION (1976) 


Mr. & Mrs. PHILIP C. WAKELEY (1976 to date) 


INDUSTRIAL SUBSCRIBERS 


(1982) 
($250 per annum) 


EXXON PRODUCTION RESEARCH COMPANY 
MosiL_ EXPLORATION AND PRODUCING SERVICES 
UNION OIL OF CALIFORNIA 


SUSTAINING MEMBERS 


(1982) 
($75 per annum) 


TOMPKINS COUNTY GEM AND MINERAL CLUB 


(continued overleaf) 


R. TUCKER ABBOTT 
ARMAND L. ADAMS 
JAMES E. ALLEN 
ETHEL Z. BAILEY 
CHRISTINA L. BALK 
Rosert A. BLACK 
HANs BOLLI 

RUTH G. BROWNE 
ANNELIESE S. CASTER 
KENNETH E. CASTER 
JOHN E. DUPONT 


ARTHUR N. DUSENBURY, JR. 


R. H. FLOWER 

Lois S. FOGELSANGER 
A. EUGENE FRITSCHE 
Ernest H. GILMOUR 
MERRIL W. HAAS 
ANITA G. HARRIS 
REBECCA S. HARRIS 
RoBerRT C. HOERLE 
F. D. HOLLAND 
RICHARD I. JOHNSON 
Davip B. JONES 
PETER JUNG 
CAROLINE H. KIERSTEAD 
CeciL H. KINDLE 


LIFE MEMBERS 


($200) 


Mary ETHYL KINDLE 
WILLIAM F. Kose, II 
Jiri Kriz 

THORWALD KRUCKOW 
HAns G. KUGLER 
EGBertT G. LEIGH, JR. 
GERARD A. LENHARD 
DONALD R. MOORE 
SAKAE OHARA 
SAMUEL T. PEES 

JOHN POJeETA, JR. 
DONALD E. RANSOM, JR. 
CASPAR RAPPENECKER 
ANTHONY RESO 
ARTHUR W. ROCKER 
JOHN B. SAUNDERS 
JUDITH SCHIEBOUT 
THomas J. M. ScHoPF 
MirIAM W. SCHRINER 
Davip H. STANSBERY 
HARRELL L. STRIMPLE 
Emity H. VOKES 
HAROLD E. VOKES 
CHRISTINE C. WAKELEY 
Puitie C. WAKELEY 
NORMAN E. WEISBORD 


Membership dues, subscriptions, and contributions are all important sources of funding, and allow 
the Paleontological Research Institution to continue its existing programs and services. The P. R. I. 
publishes two series of respected paleontological monographs, Bulletins of American Paleontology and 
Palaeontographica Americana, that give authors a relatively inexpensive outlet for the publication of 
significant longer manuscripts. In addition, it reprints rare but important older works from the paleon- 
tological literature. The P. R. I. headquarters in Ithaca, New York, houses a collection of invertebrate 
type and figured specimens, among the five largest in North America; an extensive collection of well- 
documented and curated fossil specimens that can form the basis for significant future paleontologic 
research; and a comprehensive paleontological research library. The P. R. I. wants to grow, so that it 
can make additional services available to professional paleontologists, and maintain its position as a 
leader in providing Resources for Paleontologic Research. 

The Paleontological Research Institution is a non-profit, non-private corporation, and all contri- 
butions are U. S. income tax deductible. For more information on P. R. I. programs, memberships, 
or subscriptions to P. R. I. publications, call or write: 


Peter R. Hoover 
Director 


Paleontological Research Institution 
1259 Trumansburg Road 
Ithaca, New York 14850 U.S.A. 
607-273-6623 


DEVONIAN BRACHIOPODS OF NEW ME¥ICO 


By 
G. ARTHUR COOPER J. THOMAS DuTRO, Jr. 
Smithsonian Institution and United States Geological Survey 
Washington, DC Washington, DC 
ABSTRACT 


The Devonian of New Mexico consists of six formations: the late Middle Devonian Onate Formation; the early Late Devonian 
(Frasnian—Chemungian) Sly Gap Formation and Contadero Formation; the early Famennian (Cassadagan) Thoroughgood For- 
mation and Rhodes Canyon Formation; and the middle Famennian (Cassadagan-early Conewangoan) Percha Formation. The 
Onate and Sly Gap formations are found in the Sacramento, San Andres and Caballo Mountains, the Derry Hills and Mud 
Springs Mountains and westward to Hermosa in the Black Range, south and west of which they disappear. The Contadero, 
Thoroughgood and Rhodes Canyon formations are confined to a small part of the San Andres Mountains. 

The Percha Formation consists of two members: the Ready Pay Member of black shale below; and the Box Member above, 
a somewhat calcareous and cobbly shale replete with fossils. The Percha Formation appears near Hillsboro in the Black Range, 
south of Hermosa, and extends westward to Santa Rita and Silver City and south to Cooks Peak, north of Deming. 

The Onate Formation at the eastern and western ends of its exposure is composed mostly of carbonate rocks; but, in the 
Caballo Mountains, Derry Hills and Mud Springs Mountains, it is composed mainly of fossiliferous calcareous shale. The Sly 
Gap Formation is predominantly shaly and usually cobbly; except for color and fauna, it resembles the Box Member of the 
Percha Formation and is often confused with it. The Contadero, Thoroughgood and Rhodes Canyon formations are, respectively, 
silty shale, sandstone and sandy shale. 

The fauna of the late Middle Devonian Onate Formation contains a species of Rhyssochonetes Johnson (1970) and two species 
of Warrenella Crickmay (1953) in an assemblage of 34 genera (two new) and 42 species. The Sly Gap fauna is extensive, 
consisting of 66 species (27 new) in 38 genera (one new). Distinctive in the Sly Gap fauna are: Douvillinaria Stainbrook (1945), 
Calvinaria Stainbrook (1945), Thomasaria Stainbrook (1945), Hypothyridina Buckman (1906) and the ammonoid Manticoceras 
Hyatt (1884). The Contadero fauna with Ripidiorhynchus Sartenaer (1966b) is more modest, consisting of 24 species (nine new) 
in 20 genera (two new). The Thoroughgood fauna, with a new species of Camarotoechia (ss) Hall (1893), consists of 12 species 
(three new) in 10 genera (two new). The fauna of the Rhodes Canyon Formation numbers 16 species (four new) and 12 genera 
(including Crinisarina new genus, present in both the Thoroughgood and Rhodes Canyon formations). 

The Ready Pay Member of the Famennian Percha Formation has few brachiopods and those only at top and bottom, non- 
diagnostic and poorly preserved. The Box Member, on the other hand, is extremely rich in fossils and has corals, bryozoans, 
some bivalves and gastropods, the ammonoid Falciclymenia Schindewolf (1923) as well as numerous brachiopods. Thirty 
brachiopod genera (three new) have been identified in 44 species, of which 13 are new. Important genera are: Paurorhyncha 
Cooper (1942), Syringospira Kindle (1909), and Cyrtiorina new genus. 

Correlation based on brachiopods, buttressed by conodont and ammonoid information where available, is indicated on cor- 
relation charts. It is clearly shown that the entire Percha Formation is of Famennian age and that the Sly Gap Formation is 
older and is not correlative to any part of the Percha. 

Eleven new genera and 85 new species are described in this paper. The new genera are: Dichacaenia, Morphorhynchus, 
Acutoria, Floweria, Stainbrookia, Colophragma, Crinisarina, Petasmaria, Cyrtiorina, Hispidaria and Oiosia. The new species 
and subspecies are: Lingulipora porifera, Aulacella elliptica, Stainbrookia rotunda, Cariniferella alleni, C. costellata, Thiemella 
platys, T. quadrata, Tropidoleptus platys, Strophodonta umbonata, Strophonelloides dorsoconvexus, S. parvulus, Nervostrophia 
extensa, N. geniculata, N. mucrospina, N. plana, N. solida, Douvillina contaderoensis, D. minuta, D. parviuscula, D. rectan- 
gulata, Douvillinaria diversa, Floweria transversalis, F. magnacicatrix, Schuchertella resupinata, Striatochonetes nanus, S. 
subcarinatus, Retichonetes obscurus, R. perchaensis, Rhyssochonetes johnsoni, Eostrophalosia inexpectata, Dichacaenia per- 
plexa, D. umbonata, Leioproductus pauperculus, Devonoproductus intermedius, Gypidula bowsheri, G. mimica, G. stainbrooki, 
G. subcarinata, Cupularostrum? stevensoni, Ripidiorhynchus newmexicoense, Morphorhynchus varicostatum, Hypothyridina 
emmonsi rotunda, Hadrorhynchia alta, H. mclareni, Coeloterorhynchus magnificum, Colophragma ellipticum, Trifidorostellum 
obscurum, Camarotoechia latisinuata, Leiorhynchus sphaericum, Platyterorhynchus obscurum, Petasmaria compacta, P. mul- 
ticostata, P. patens, Atryparia rubra, Desquamatia costata, Costatrypa extensa, lowatrypa rara, Spinatrypa compacta, S. 
obsolescens, S. trulla decorticata, Crinisarina reticulata, Ambocoelia capillata, Echinocoelia pretiosa, Eleutherokomma exten- 
sa, Mucrospirifer parvus, Tylothyris compacta, T. sulcocostata, Acutoria angulata, Acutatheca prolifica, Thomasaria demissa, 
T. warreni, Cyrtospirifer arenarius, C. convexus, C. pyramidalis, C. rotundus, Hispidaria posterogranulosa, H. spinosa, Uch- 
tospirifer? rarus, Indospirifer anomala, I. exfoliata, Warrenella magna, W. floweri, Elita magister, Cyrtina variabilis, Afilasma 
inexpectatum. 


6 BULLETIN 315 


INTRODUCTION 
PURPOSE AND SCOPE 


The geologic history of the Devonian of New Mex- 
ico is developed from analysis of the strata and fossils, 
with special emphasis on the systematic description of 
the brachiopods. Devonian rocks are widely distrib- 
uted in the south-central and southwestern parts of the 
state, especially in the Bap meure, San Andres, and 
Caballo Mountains (Text-figs. 1, 2, 3). Other critical 
exposures are in the Black Range, Derry Hills, Mud 
Springs Mountains, and Cooks Peak area. Detailed 
stratigraphic sections and biostratigraphic data are 
presented, from which correlations, facies distribu- 
tions, and paleogeographic conclusions are drawn. A 
faunal zonation, based primarily on brachiopods, is 
given and its age significance is discussed. This is sup- 
ported by evidence based on occurrences of con- 
odonts, corals and cephalopods. 


GEOGRAPHIC DISTRIBUTION 


Devonian rocks are thin over most of New Mexico, 
ranging up to about 100 m in the thickest sections. 
Most sections, however, are less than 30 m (100 ft) 
thick. This fact led most early workers, who were 
mainly concerned with geologic mapping, to place all 
Devonian rocks in a single formation—the Percha Shale 
(Cooper et al., 1942). However, this thin sequence 
contains a variety of lithic facies and faunas and is 
now known to range in age from late Middle Devonian 
to mid-Famennian. Full appreciation of these com- 
plexities has come slowly as data accumulated over 


SOCORRO 


Text-figure 1.—Index map of south-central New Mexico, showing 
major Outcrop areas of Devonian rocks; location of Figures 2 and 
3: 1, Rincon; 2, southern Caballo Mountains; 3, Mud Springs 


Mountains; 4, Hermosa; 5, Chise; 6, Hillsboro area; 7, Silver City 
area; 8, Cooks Range. 


the past 25 years, but the present state of knowledge 
permits a detailed synthesis that places the Devonian 
of New Mexico in modern perspective. 

Despite statements to the contrary in earlier papers, 
several distinct rock units can be recognized in the 
sequence, and each contains one or more distinct fau- 
nas. The oldest unit, the Onate Formation, crops out 
mainly in the San Andres and Sacramento Mountains 
(Text-figs. 4, 5), although it is also exposed in the Ca- 
ballo Mountains, the Derry Hills, and in the Mud 
Springs Mountains west of Truth-or-Consequences. A 
critical locality for regional interpretation of the Onate 
is near Hermosa, the westernmost outcrop in which 
the Onate has been recognized. A southern and south- 
eastern facies of the Onate is the Canutillo Formation 
of the Franklin Mountains in west Texas. 

The Sly Gap Formation overlies the Onate in most 
places, although it is missing in the Caballo Mountains 
and in the southernmost sections of the Sacramento 
Mountains. This discontinuity may be due to a period 
of late Devonian erosion. In the central part of the San 
Andres Mountains, the Sly Gap is overlain by the Con- 
tadero Formation, which does not occur elsewhere. 
Still more limited in extent are the post-Contadero 
units, the Thoroughgood and Rhodes Canyon forma- 
tions, which are restricted to a small area in the San 
Andres Mountains. 

The Percha Formation, widely exposed in the Black 
Range between Hillsboro and Silver City, extends 
south to the Cooks Peak area and beyond. This unit 
is divided into a lower black shale (the Ready Pay, or 
Silver*, Member) and an upper shaly nodular lime- 
stone (the Box, or Bella*, Member). Relationship of 
the Percha to older Devonian units is displayed near 
Hermosa where the Ready Pay Member unconform- 
ably overlies the Sly Gap Formation which, in turn, 
succeeds the Onate Formation in normal stratigraphic 
sequence (Text-fig. 6). Although occurrences east of 
Hillsboro are recorded in the literature, we found no 
exposures of the Box Member of the Percha in the 
ranges east of that place; the occurrence so designat- 
ed, Derry Hills, proved to be of Pennsylvanian age. 


NATURE OF OUTCROPS 


The Devonian strata are predominantly shaly and 
occur as partly covered slopes between more resistant 
formations above and below. Debris from overlying 
units commonly covers the shaly slopes, and complete 
sections of the Devonian are hard to find. Good out- 


* The Silver and Bella members are terms proposed by C. R. 
Keyes (1908); although prior in publication to Stevenson's (1945) 
names, they have not received acceptance because of poor defini- 
tion and poor type sections. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 7 


(W@W worroro |! 
LOOKOUT /| 
a 


S 
“Onego ! 


Peok Al) 


Bor b/e Con 
— 


B Joplin Ridge 


Dog Conte 
o 9 
Burnt| Ridge 
| 


GobbierA 
1 


A. Lightning 
aa) Loke 


) 
i 


~ 


Principal drainage divides 
">< Bose of mountain escarpment 
— + — Lincoln Notional! Forest boundary 
SSSSS5 Areo of geologic maps (Pray, 196C) 
@ Fire jookout tower 
Paved road 
~™~-— !mproved road 


Gj ACulp Peok 


i 2 3 4 3 
Scale in miles 
RIZE 


Text-figure 2.—Map of a part of the Sacramento Mountains, showing location of measured sections of Onate Formation, Sly Gap Formation, 
and related strata. 1, Pig Canyon anticline; 2, Arcente Canyon; 3, Alamo Peak; 4, Alamo Canyon; 5, Indian Wells Canyon; 6, Deadman 
Canyon; 7, Mule Canyon; 8, San Andres Canyon; 9, Dog Canyon; 10, Escondido Canyon. Base map is from Pray, 1961, figure 2. 


crops occur in the canyons of the San Andres and 
Sacramento Mountains, but care must be taken in as- 
signing a stratigraphic position to fossils collected, be- 
cause these migrate downslope from bedrock expo- 
sures. 


ACKNOWLEDGMENTS 


Although C. C. Branson is now deceased, it is still 
fitting to acknowledge our obligations to him for ac- 
quainting Cooper with the geology and geography of 
the Devonian of New Mexico and for the collections 
he presented to the National Museum of Natural His- 
tory. We recall our work with A. L. Bowsher, later 


with the U.S. Geological Survey, with great pleasure. 
We also acknowledge the help of W. T. Allen who 
assisted Bowsher in the field. 

Our special thanks go to R. Flower, New Mexico 
Bureau of Mines and Mineral Resources, for help in 
the field and for the loan of his useful Devonian col- 
lections. We thank the officers of the White Sands 
Proving Ground for permission to examine sections in 
the San Andres Mountains. The many roads bulldozed 
into the poorly accessible parts of the range proved a 
boon to our efforts. 

We are greatly indebted to F. V. Stevenson, of 
Mountain Home, Arkansas, for the extensive collec- 


8 BULLETIN 315 


tion of New Mexico Devonian fossils he presented to 
the Smithsonian Institution in 1947. 

We appreciate the careful reviews of the manuscript 
by A. J. Boucot, J. G. Johnson and G. Klapper. Help- 
ful suggestions on brachiopod systematics by Boucot 
and Johnson, and on conodont ages and correlations 
by Klapper and A. G. Harris, are gratefully acknowl- 
edged. P. Sartenaer also provided comments on the 
brachiopod taxonomy, which we deeply appreciate. 
However, the final decisions on the taxonomy, age 
designations and correlations are our own and do not 
always agree with the views of the reviewers. 

Finally, we thank T. Lierman for his superb drafting 
of the text-figures, N. P. Dutro for her suggestions on 
illustration design and an indefatigable dedication to 
production of the index, and P. R. Hoover for innu- 
merable editorial suggestions that greatly improved the 
final product. 


PREVIOUS WORK 


Geologic work in the Devonian of New Mexico in 
the early part of this century was concentrated on the 
black shales in the southwestern part of the state. These 
strata were named the Percha Shale by Gordon (1907) 
and were later (Paige, 1916) described in the Silver 
City area. Formal member names were proposed by 
Stevenson (1945) who called the “‘lower Percha shale”’ 
the Ready Pay Member and the “‘upper Percha shale’’ 
the Box Member. Stevenson also established the 
stratigraphic framework for distinguishing other De- 
vonian units in south-central New Mexico and named 
the Sly Gap Formation (Stevenson, 1941b) and the 
Onate and Contadero formations (Stevenson, 1945). A 
good summary of work on the Devonian of New Mex- 
ico up to the mid-1940’s is given in Stevenson (1945). 

Prior to Stevenson’s studies, Darton (1917, p. 44— 
46) described Devonian strata in the various ranges 
from Cooks Range to the Sacramento Mountains. The 
strata, regardless of lithology and fossil content, were 
called Percha Shale. However, he noted that ‘‘The 
beds in the San Andres Mountains appear to carry a 
smaller number of the typical lower Ouray Percha 
forms than are present in Lake Valley, and more of 
the forms characteristic of the Martin, Hackberry and 
Nevada limestone.” 

In a short paper describing the fauna and correlation 
of the Sly Gap Formation, M. A. Stainbrook (1948) 
compared that assemblage with the Independence Shale 
fauna of lowa, which he had recently described (Stain- 
brook, 1945). Several sections of the Upper Devonian 
strata were graphically presented and briefly described 
in two excellent papers on the Mississippian of south- 
western New Mexico (Laudon and Bowsher, 1941; 
1949). 


Three regional monographs of large areas in south- 
ern New Mexico give many details of Devonian stra- 
tigraphy and geologic history. In a paper on the ge- 
ology of the Caballo Mountains by Kelly and Silver 
(1952), the Devonian shales are discussed as Percha 
Formation. However, the sequence at Haystack Can- 
yon is considered of Sly Gap age and Stevenson is 
quoted (1945) as assigning the beds in the Mud Springs 
Mountains and Caballo Mountains to the Sly Gap For- 
mation. The exposures at Nackaye Mines were over- 
looked by Kelly and Silver. 

Several detailed sections are presented by Kott- 
lowski et al. (1956) in their stratigraphic study of the 
San Andres Mountains. With some modification and 
correction, the stratigraphic data on the Devonian of 
the Rhodes Canyon, Hembrillo Canyon and Ash Can- 
yon areas are used as a basis for our biostratigraphic 
analysis. 

In his paper on the geology of the Sacramento 
Mountains escarpment, Pray (1961) gives some addi- 
tional details on the Devonian formations, especially 
the Onate Formation. He also discusses the problem 
of the black shales, in certain local areas. He assigns 
these black sediments to the Percha Shale. Pray’s 
mapping is invaluable for locating critical Devonian 
sections in the Sacramento Mountains. 

Jicha (1954) mapped the Percha Formation in the 
Lake Valley quadrangle; and Zeller (1965) discusses 
Devonian formations in the Big Hatchet Mountains 
area. 

Several papers by Flower (1958, 1959, 1965, 1970) 
elucidate the early Paleozoic and Devonian in New 
Mexico. Flower (1959) redefined the Contadero For- 
mation and named two new post-Contadero forma- 
tions in the central San Andres Mountains. He showed 
that the Thoroughgood Formation overlies the Sly Gap 
in the central San Andres area and proposed the Rhodes 
Canyon Formation to describe about 23 m (75 ft) of 
beds above the Thoroughgood in the Rhodes Canyon 
section. 

Bowsher (1967) summarized all earlier published 
work, and included data from field work accomplished 
in company of L. R. Laudon, and by the present au- 
thors, in a general paper on the Devonian System in 
New Mexico. 


Text-figure 3.—Map of the San Andres Mountains, showing lo- 
cations of measured sections of Devonian and related strata. 1, 
Johnson Park; 2, Capital Peak; 3, Sheep Mountain; 4, Rhodes Can- 
yon; 5, Ash Canyon (north); 6, Hembrillo Canyon; 7, San Andres 
Canyon; 8, Ash Canyon (south); 9, Bear Canyon. Localities west 
of San Andres Mountains: R = Rincon; D = Derry; M = Mud 
Springs Mountains. (from Kottlowski ef al., 1956, fig. 1.) 


Ms aaynatey 
Seneca yaateet 


EONS aan ae pean tte 


° 


MUD ~© 

SPRING 

f emTs 
+ 


ruth Butt 


=xor E 


Consequences © 


aye 


Ren) 


> 


ui 
7 


4\ 
4 


: Elephani- 


he 


ey 


«0g 4 
ce? eS 


ihe, 
eed LL 


ll leva. 


oy 
tery” 


DEVONIAN BRACHIOPODS OF NEW MExICO: COOPER AND DuTRO 


10 Miles OSCURA 


MOUNTAINS 


t) 
o. 


— 


hodes Canyon 


UU 


ae 


x 25 
2 
e Black Top 
SRM 
sulphur Canyon EEG P As 
Bye | e 


5 "% 
peeel'y 
Z 


%, 
% 
4 
% 
%, 
, 
t 
= 


wey 
ri Dead Mon Canyon 


equal 


, 


Three Rivers 


Tularosa 


> 
“s 


Kaylor ‘i Se il aed gees 
LeMin ESD Gwe Oe ita Cae 
Hembrillo Canyén-- -.. 7 - +=: 


Alamogordo 


: a eWhite Sands 


WAN atremee 


San Andres x, 


Peok 

% 8 9o 04, 

. vornada Pec “4 
Ronge 


glove Ranch= 
9 8 e0r Canyon 
<%, > 
> 


8eor Peaks® 


% I 


e 
wy, oY 
~ oe st, 


e* SSon Augustin Poss 
\ 
eWhite Sands 
Headquorters 


Oe, s 


mash ngs tt its 


Los Cruces 
Stote.. 


College 
Mesilla © 
Park 


ae 
“uy, ev 
° 


> 
2 


fe) 
va, 
“ aprnwarnarh, 


= 
4 
“ 
° 


Pre deems ore 


Location 
Figure 


” 


National Monument 


A “s 
£ 


asthe 


PU 


RN 


i 


ae 


ye, 
C4 


Orogrande 


a 
ae 


of 


BULLETIN 315 


ONATE FORMATION 
(Pl. A) 


Type SECTION AND GENERAL 
STRATIGRAPHY 


The type section of the Onate Formation, named by 
Stevenson (1945), is in San Andres Canyon in the San 
Andres Mountains (loc. 523s). The formation consists 
predominantly of ferruginous dolomitic siltstone, shale 
and limestone and is about 27 m (90 ft) thick in the 
type area. Several thickness measurements, made in 
San Andres Canyon by various investigators, range 
from about 25 to 28 m (85 to 95 ft). The thickness of 
the Onate is variable in the San Andres Mountains, 
and it thins both to the north and south of the type 
section. 

In the Sacramento Mountains (Pl. A, figs. 1, 3), the 
Onate is lithologically similar to its outcrops in the San 
Andres Mountains, and varies from about 6 to 18 m 
thick (20 to 60 ft). The thickest section is in Dog Can- 
yon and the unit generally thins both to the north and 
south. 

West of the San Andres Mountains, the Onate is 
present in the southern Caballo Mountains, near Rin- 
con. In the Derry Hills and the Mud Springs Moun- 
tains it is mostly shale. The Onate has been recognized 
as far west as Hermosa where it is 7 m (23 ft) thick 
and is mostly limestone. 

The Onate Formation is regionally unconformable 
above various early Paleozoic formations (Text-fig. 7). 
In the southern San Andres and southern Sacramento 
Mountains, it lies on the Silurian Fusselman Dolomite. 
North of Hembrillo Canyon in the San Andres Moun- 
tains, the underlying units are the upper formations 
and members of the Montoya Group (late Ordovician). 
In the more western exposures, from Mud Springs 
Mountains to Hermosa, the Onate again rests uncon- 
formably on the Fusselman Dolomite. 

The Sly Gap Formation disconformably overlies the 
Onate Formation except where the Sly Gap has been 
removed by late Devonian erosion. In the Sacramento 
Mountains, from Pig Canyon south to Mule Canyon, 
reddish-weathering, ferruginous, silty dolomite forms 
the uppermost beds of the Onate. Basal beds of the 
Sly Gap consist of greenish to gray clay shale with 
scattered limestone nodules. South of Mule Canyon, 
the early Mississippian Caballero Formation rests un- 
conformably on the Onate. In the southern San Andres 
Mountains, the Onate is overlain by a dominantly dark 
grey shale sequence that we assign to the Sly Gap. 


FossiL CONTENT AND DISTRIBUTION 


Fossils are widely, but sparsely, distributed in the 
Onate throughout its area of outcrop. Although bra- 
chiopods are the most abundant faunal element, there 
are also many bryozoans (mainly Cystodictya Ulrich, 
1882), a few tabulate corals (Aulopora Goldfuss, 1829), 
trace fossils (burrows and borings) and, in the Mud 
Springs Mountains (locs. 522f, 5231), the receptaculitid 
Sphaerospongia cf. S. tesselata (Phillips, 1841). 

In the type section in San Andres Canyon in the San 
Andres Mountains, the lower 7 m (25 ft) contains 
abundant specimens of Rhyssochonetes Johnson (1970) 
with a few individuals of Echinocoelia Cooper and 
Williams (1935) near the top. A large species of Platy- 
terorhynchus Sartenaer (1970) is abundant in the over- 
lying 2.4 m (8 ft) thick unit. The next higher bed (4 m, 
or 13 ft thick) contains abundant Cystodictya, along 
with Platyterorhynchus, Nervostrophia Caster (1939) 
and Rhyssochonetes. The overlying unit, 2.7 m (9 ft) 
thick, has abundant bryozoans and the brachiopods 
Schizophoria King (1850), Nervostrophia, Dichacaen- 
ia n. gen. and Atryparia Copper (1966). In the upper 
part of the formation, 4.5 to 8 m (15 to 27 ft) below 
the top, large individuals of Warrenella magna n. sp. 
are present, and Schizophoria and Desquamatia Alek- 
seeva (1960) are abundant. This sequence serves to tie 
together the Onate of the western exposures with that 
of the Sacramento Mountains in that it includes the 
western Warrenella beds, not present in the Sacra- 
mentos, and the Platyterorhynchus-Cystodictya as- 
semblage, common over the entire extent of Onate 
exposures. 

Typical of the Sacramento Mountains Onate is the 
sequence at Alamo Peak where Platyterorhynchus and 
Cystodictya dominate the assemblage and no partic- 
ular vertical zonation is evident. Other genera, com- 
mon to the type section, are: Schizophoria, Nervo- 
strophia, Atryparia, Desquamatia and Echinocoelia. 
Neither Warrenella nor Rhyssochonetes was found in 
the Sacramentos. Striatochonetes Mikryukov (1968) 
apparently occupies a place in the Sacramento assem- 
blage similar to that of Rhyssochonetes in the western 
exposures. 

The Mud Springs Mountains section is characteris- 
tic of the western part of the region. Rhyssochonetes 
occurs commonly in the lower 3 m (10 ft). The over- 
lying 6 m (20 ft) contains a varied assemblage, includ- 
ing Schizophoria, Leiorhynchus Hall (1860a), Mor- 
phorhynchus n. gen., Atryparia, Desquamatia and 
Eleutherokomma Crickmay (1950), in addition to 
Sphaerospongia. The uppermost 6 m (20 ft) contains 
the same brachiopods plus Nervostrophia, Dicha- 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DUTRO 11 


PERCHA 
TYPE AREA 
(composite) 


Lake Valley 
HERMOSA 


Andrecito Mbr 


RIN 
CHISE CON 


\ ZA MUD SPRINGS Miss.? or 
Miss Ya ? 
ph \ MOUNTAINS Contadero 
(50ft missing) // 
ib ch \ / / 
Ready Pay © 
Member 
135 ft \ / 
Ready Pay bi 
Member a == / i) 
bl a 
Aecbe = cono / 
Ready Pay / gd 
Member Vy 526d 
ub \ Penn / } 
y, Sly Gap 
Formation 
20ft Vy, 
i, va ] 526c Ree 
/. ye ; ™~ 
a —_——— a—_ = = == : 
Wm Ha D 
Fusselman NG ] = are m Ha Des Hite Per 
At? . ke 
X Sch a Formation 
a Montoya covered 
= (Valmont) Ss 


Montoya 
y covered 


Fusselman 


Text-figure 6.—Correlated stratigraphic sections of Devonian rocks at selected localities west of the San Andres Mountains, New Mexico 
(See Text-figure | for location of sections and Text-figure 5 for explanations of abbreviations and lithic patterns). 


caenia, Hypothyridina Buckman (1906) and small in- 
dividuals of Warrenella floweri n. sp. 

At the westernmost known exposures of the Onate 
Formation, near Hermosa, Rhyssochonetes occurs in 
the lower 2 m (7 ft) and a characteristic assemblage is 
found in the upper | m (4 ft): Schizophoria, Nervo- 
strophia, Dichacaenia, Hypothyridina, Eleutherokom- 
ma, and Desquamatia, together with Cystodictya. 

An interesting variant on the Onate brachiopod as- 
semblage occurs in the northern part of the San Andres 
Mountains where the formation is only about 3 m (10 


ft) thick. Two species: Tropidoleptus platys n. sp., 
occurring only on Capitol Peak and in Johnson Park 
Canyon, and Acutoria angulata n. sp., found only in 
Johnson Park Canyon, are especially abundant. These 
occur with species found elsewhere in the Onate, in- 
cluding: Nervostrophia plana n. sp., Nervostrophia 
solida n. sp., Schizophoria lata Stainbrook (1940), and 
Platyterorhynchus obscurum n. sp. Also present is 
Mucrospirifer? aff. M? sculptilis (Hall, 1843) and Pen- 
tamerella sp., both links to the Middle Devonian of 
the eastern United States. 


BULLETIN 315 


Cephalopod Conodont 


Sacramento Mtns. 


Zones 


Caballero Fm. (Kinderhookian) 
Wocklumeria Bispatho. ar 
Pe ee costatus ais 
Clymenia Poly. _ 
styriacus = = 
a) 

& Scaph. “_ 

a 5 
3 velifer =e 

= ; 

re) do lila Palin 

marginifera 
Famennian 


do ll B 
Palm. rhomboidea 


Cheiloceras 


Q 
) 
= 
2 
Palm. 
Crepida 


triangularis 


Palm. gigas 


Sly Gap 


Formation 


Manticoceras 


Frasnian ? 


Upper 


Middle 


Polygnathus 
asymmetricus 


Lower 


Lowermost 
P.dengleri 


hermanni-cristatus 


Maenioceras 


Formation 


Ju. Ord N_ Fusselman Dolo. 


Givetian 


Polygnathus varcus 


(upper) 


Underlying units 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 


Southern} Mud : Silver 
__ |Hillsb 

N  SanAndres Mtns.  §| Caballo | Springs Chise Jo el City 
Mtns. Mtns. : 


Lake Valley Fm./Caballero Fm. |Miss.\ Penn] Penn. | Miss. | Miss.? | Caballero | Caballero 


? 
Rhodes 
Canyon 

Fm. 


? 
Thorough - 
good Fm. 

Si 


? 
Contadero 
Formation 
—_e— 
\ 


NX 
/ 
Sly Gap “XN 


Formation 


Onate Formation 


ea ei AGE 
Dolo. Dolo. (Ord.) (Ord.) Dolo. Dolo. 


Text-figure 7.—Correlation diagram of Devonian rocks in New Mexico. 


14 BULLETIN 315 


BRACHIOPOD ASSEMBLAGE OF THE 
ONATE FORMATION 


The total brachiopod assemblage of the Onate con- 
sists of 34 genera and 41 species. Of these, 17 genera 
and 20 species characterize the assemblage biologi- 
cally and biostratigraphically. Three new genera, Di- 
chacaenia (a strophalosiid), Morphorhynchus (a rhyn- 
chonellid), and Acutoria (a spiriferid), and 20 new 
species are described herein. 


AGE OF THE ONATE FORMATION 


The Onate Formation is of late Middle Devonian age 
(Taghanician Stage—late Givetian) and is approximate- 
ly correlative with the lower Tully Limestone of New 
York and the lower Cedar Valley Limestone (Solon 
Member) of Iowa and Illinois (Text-fig. 8). The 
common brachiopod species in the Onate are: Schizo- 
phoria lata Stainbrook (1940), Rhyssochonetes john- 
soni n. sp., Nervostrophia plana n. sp., Morphorhyn- 
chus varicostatum n. sp., Warrenella floweri n. sp., 
W. magna n. sp., and Desquamatia costata n. sp. 

Schizophoria lata was described by Stainbrook from 
the Cedar Valley Limestone of Iowa; the Rhyssocho- 
netes is related both to R. aurora (Hall, 1867) from 
the lower Tully Limestone of New York and the Pine 
Point Formation in Canada and also to the Rhyssocho- 
netes that occurs in the northern Roberts Mountains, 
Nevada (Interval 21 of Johnson, 1978), in the Winne- 
pegosan Limestone of Manitoba and the Solon Mem- 
ber of the Cedar Valley Formation of Iowa. Nervo- 
strophia plana is very like N. tulliensis (H. S. Wil- 
liams, 1890); and Morphorhynchus varicostatum seems 
related to Middle Devonian species of *‘Hadrorhyn- 
chia’ reported from western Canada (McLaren, 1962). 

Of the rarer forms, Leiorhynchus sphaericum 1s close 
to L. castaneum Meek (1867) from the Nevada Middle 
Devonian and from the Pine Point Formation. Had- 
rorhynchia alta and H. mclareni are related to H. san- 
dersoni (Warren, 1944), also from the Pine Point For- 
mation. . 

In addition to the brachiopods, the bryozoan Cys- 
todictya anomalotruncata (Fritz, 1944) is common in 
the upper part of the formation in several places. This 
species is most similar to species described by McNair 
(1937) from the Middle Devonian of Michigan, as Fritz 
indicated in her paper. 

Conodonts are rare in the Onate, but a few speci- 
mens from the lower part of the formation in the Mud 
Springs Mountains, where they occur with Rhysso- 
chonetes johnsoni n. sp., were assigned by Huddle 
(written commun., 1974) to the Polygnathus varcus 
Zone. In New York, this zone is found in the upper 


part of the Middle Devonian, from the Centerfield 
through the Tully formations (Klapper et al., 1971). In 
a paper revising the varcus Zone (Ziegler, Klapper, 
and Johnson, 1976), Rhyssochonetes is reported both 
in Nevada and New York with middle varcus Subzone 
conodonts. 

One of the more exotic non-brachiopod elements of 
the fauna is Sphaerospongia cf. S. tesselata (Phillips, 
1841) which is abundant in the Mud Springs Mountains 
in the *‘Sponge beds’’. This receptaculitid genus oc- 
curs in England and Germany in the late Middle De- 
vonian (Warren, 1944). Wells (1940) described a spec- 
imen from the upper Tully Limestone in New York 
and discussed occurrences in the Winnepegosan 
Limestone in Manitoba and the Torquay Limestone in 


Table 1.—Brachiopods of the Onate Formation (+ = especially 
important elements). 


Lingula species 3 
Lingula species 4 
Trigonoglossa? species 
Orbiculoidea species 
Petrocrania species | 
+  Schizophoria lata Stainbrook 
Schizophoria species 1 
Rhipidomella species 2 
+  Tropidoleptus platys new species 
Strophodonta species | 
+  Nervostrophia plana new species 
“Schuchertella’’ species 
Devonochonetes? species 
+  Striatochonetes nanus new species 
S. subcarinatus new species 
+ Rhyssochonetes johnsoni new species 
Eostrophalosia species | 
+  Dichacaenia perplexa new genus and species 
+ D.umbonata new genus and species 
Pentamerella species 
Cupularostrum? species 
+ Morphorhynchus varicostatum new genus and species 
Hypothyridina species 
Hadrorhynchia alta new species 
H. mclareni new species 
Leiorhynchus sphaericum new species 
Platyterorhynchus obscurum new species 
Atryparia rubra new species 
Desquamatia costata new species 
Emanuella species 
Echinocoelia pretiosa new species 
+  Eleutherokomma extensa new species 
+ Mucrospirifer parvus new species 
cf. M.? sculptilis (Hall) 
+  Acutoria angulata new genus and species 
Orthospirifer species 
Spinocyrtia? species 
+ Warrenella floweri new species 
W. magna new species 
+  Cyrtina variabilis new species 
Cranaena species | 


+++ tte + 


+ 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DuTRO 15 


Devonshire, England. A similar species also occurs in 
the Middle Devonian Hare Indian Shale of northwest- 
ern Canada (Warren and Stelck, 1956). 


SLY GAP FORMATION 
(Pl. B; Pl. C, figs. 1, 2) 


Type SECTION AND GENERAL STRATIGRAPHY 


The type section of the Sly Gap Formation (Steven- 
son, 1945) is on the south slope of Sheep Mountain 
(loc. 506j) in the northern part of the San Andres Range 
(Pl. B, figs. 1, 2). The sequence is dominantly shaly 
and is composed of dark gray laminated shaly units 
alternating with greenish gray nodular units, not unlike 
Percha lithic types but lighter in color. The calcareous 
nodules are particularly dense in the lower, middle and 
uppermost parts of the formation. These fossiliferous 
nodular zones are, essentially, units D, G, and upper- 
most unit H of Stevenson (1945). Although Stevenson 
originally considered the Sly Gap to be about 35 m 
(115 ft) thick, our work has shown that: unit C, at the 
base, is the Onate Formation; units I and J belong to 
the Contadero Formation; and the upper two units (K 
and L) are assigned to the Thoroughgood Formation. 

Consequently, the revised Sly Gap of the type sec- 
tion is about 20 m (65 ft) thick. This includes Steven- 
son’s units D through H (1945, fig. 9). Measurements 
made by us, although differing in details, indicate that 
the Sly Gap varies in thickness from about 20 to 22 m 
(65 to 70 ft) in the type area. To the north, it thins to 
about 12 m (40 ft) at Johnson Park (formerly Mock- 
ingbird Gap) where rocks of early Mississippian age 
unconformably overly this formation. Post-Sly Gap 
strata apparently were removed by late Devonian ero- 
sion in the northernmost part of the San Andres Moun- 
tains. In the Rhodes Pass area, south of the type sec- 
tion, the Sly Gap Formation is about 24 m (80 ft) thick. 
The formation thins to less than 12 m (40 ft) at San 
Andres Canyon and is either missing in the southern 
San Andres Mountains or is replaced by a dark shale 
facies to the south. 

In the Sacramento Mountains, the Sly Gap varies in 
thickness from about 9 to 20 m (30 to 65 ft). The vari- 
ation is again apparently due to late Devonian erosion, 
as the early Mississippian Caballero Formation every- 
where lies disconformably on the Sly Gap. South of 
Mule Canyon, the Caballero lies disconformably on 
the Onate Formation and the Sly Gap is missing. 

In the Mud Springs Mountains, the Sly Gap is about 
16 m (54 ft) thick and is overlain unconformably by 
strata of Pennsylvanian age. This is also the case in 
the Derry Hills, where the Sly Gap is probably no 
thicker than 12 m (40 ft). 


Layers of black shale that occur in the Sly Gap in 
Marble Canyon and in the southern part of the Sac- 
ramento Mountains have been erroneously interpreted 
as tongues of Ready Pay black shale, producing the 
equally erroneous interpretation that the Sly Gap is a 
facies of the Ready Pay. 

The westernmost exposures near Hermosa contain 
about 12 m (40 ft) of Sly Gap beds that overlie the 
Onate and are disconformably overlain by the Ready 
Pay Member of the Percha Shale. 


FosstL CONTENT AND DISTRIBUTION 


In the type area, and elsewhere in the northern part 
of the San Andres Mountains north of Rhodes Pass, 
the characteristic brachiopod assemblages of the Sly 
Gap are superbly developed. Although many of the 
species range through the entire thickness of the for- 
mation, occurrences suggest there is an an approxi- 
mate five-fold zonation, discussed below: 


Colonial Coral Zone 


The upper 1.5 to 2.5 m (5 to 8 ft) contains abundant 
corals, including Alveolites Lamarck (1801), Pachy- 
phyllum Milne-Edwards and Haime (1850), “‘“Hexa- 
gonaria’’, together with Macgeea Webster (1889b) and 
a few brachiopods. This zone is equivalent to the up- 
per part of Stevenson’s (1945) unit H. 


Upper Brachiopod Zone 


About 5 to 6 m (17 to 20 feet) of strata below the 
coral zone consists of characteristically nodular shale 
with many brachiopod species and the coral Macgeea. 
It was also from this level that the ammonite Manti- 
coceras Hyatt (1884) was reported. Collection 521q 
represents this assemblage well. Common species are: 
Aulacella elliptica n. sp., Nervostrophia geniculata n. 
sp., N. mucrospina n. sp., Douvillina parviuscula n. 
sp., Devonoproductus intermedius n. sp., Costatrypa 
varicostata (Stainbrook, 1945), Jowatrypa rara n. sp., 
Pseudoatrypa devoniana (Webster, 1921), Thomas- 
aria warreni n. sp., Elita magister n. sp., Cyrtina in- 
ulta Stainbrook (1945), and Indospirifer anomala n. 
sp. The highest occurrence of Hypothyridina Buck- 
man (1906) is near the base of this unit. This zone is 
equivalent to the middle part of Stevenson’s unit H. 


Lower Brachiopod Zone 


The next lower 6 m (20 ft), or so, also contains a 
variety of brachiopods, and Macgeea—the common 
solitary coral found all through the Sly Gap. Collection 
521k contains many of the species that comprise the 
typical Sly Gap brachiopod assemblage, including: 


BULLETIN 315 


Cephalopod Conodont 


Zones Zones 


Overlying units | _——SsSSC—~*™ 


|_ Miss. N._Penn_ | __Escabrosa_Fm.__| 


Bs 

Wocklumeria Bispatho. | MM. | 
— costatus 
Clymenia 

2 Poly. styriacus | M_| 
(SES) 

Maes 

Platyclymenia velifer 
Palm 

marginifera 


Famennian 
Palm. rhomboidea 


horoug 
Cheiloceras 
Middle 


Lower 


Palm. 
crepida 


Upper 


Middle 


—_?—_ — 


Palm 
triangularis 


Lower 


Contadero 


Portal 


Uppermost Formation 


Cc 
° 
S 
8 oO 
B=) _————— Fm. E 
> 2 
: Upper = 
E Sly Gap t 
3 o 
a = 
Lower Formation 
Manticoceras 
: Ancyrognathus _—_—?—_- 
Frasnian triangularis 
2 § 
<x 2 Mi 
ss iddle 
£0 
aE 
= & 
3S 
a 4 
% 
Lowermost 
(P. dengleri) 
? — ? —— 
hermanni-cristatus 
Onate 
Givetian Formation 


Maenioceras Polygnathus 
(upper) 


varcus 


| Lower| 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 17 


[Wiss] 


[Knapp Col] 
Ellsworth 


Colorado 
| Miss. __| 


Leadville Ls. | Miss. | 


mes R. 


Fitchville 
Fm. (part) 


Pinyon 
Peak 
Limestone 


? 


Dyer | Ouray| Chaffee Amity Sh 
is ? 
Parting 


Mbr. 


? 


Palliser 


Conewangoan 


Maple Mill 


Shale 
? 


Aplington 


Shale 


Formation 


Ellicott Shale 
Dexterville Shale 


Northeast Shale 
|_Shumla Ss. 
Westfield Shale 


Dunkirk 
Shale 


Hanover Shale — 


Forks 


Three 


Sheffield 


Cassadagan 


Wiscoy Fm. 


Formation 
Cohoctonian 


Norwood Rhinestreet 


Shale 


Jefferson 


Limestone 


Formation 


West River 
Shale 


Penn Yan Shale 


Geneseo Shale 


? 
Jordan R. 


Squaw Ba 


Fingerlakesian 


Bluebell 


Dolo. 


Guilmette 


t 
Denay (part) 


| Rapid | Thunder 
ay 
Potter 
Farm 
ap 


? 
reamn 

i Point I 
nae 


Simonson Beartooth 
[Fasselman] M.dev. [Simonson | siurien [Beers er 


Canutillo | Limestone 


Tully 


Formation 


Formation 


ir TE Fm. 
Taghanician 


(upper) 


Ludlowville 


Centerfield 


cavers Hamilton Grp. ae 


Text-figure 8.—Correlation of New Mexico Devonian strata with selected Devonian sequences in North America. Members of Guilmette 
Formation from Drewes (1967). 


8 BULLETIN 315 


Calvinaria bransoni Stainbrook (1948), Cariniferella 
alleni n. sp., Coeloterorhynchus magnificum, Nn. sp., 
Costatrypa extensa n. sp., Cyrtina inulta, Devono- 
productus intermedius n. sp., Gypidula subcarinata n. 
sp., Hypothyridina emmonsi rotunda n. subsp., In- 
dospirifer aff. I. orestes (Hall and Whitfield, 1872), 
Nervostrophia mucrospina n. sp., Productella rugu- 
lata Stainbrook (1945), Pseudoatrypa devoniana, 
Schizophoria iowensis (Hall, 1858), Spinatrypa trulla 
decorticata n. subsp., Thomasaria warreni n. sp., T. 
demissa n. sp., and Tylothyris sulcocostata n. sp. This 
zone is equivalent to the lower part of Stevenson’s 
unit H and his unit G (Stevenson, 1945). 


Atrypa Zone 


This zone is essentially equivalent to Stevenson’s 
(1945) unit E and includes a variety of the atrypids 
that are listed above for the lower brachiopod zone, 
but lacks the wide variety of species of that zone. 


Macgeea Zone 


Although the little horn coral Macgeea is found 
throughout the Sly Gap, its lowest occurrence is near 
the base of the formation where it is found with a 
sparse brachiopod fauna that includes Cyrtina David- 
son (1858) and Coeloterorhynchus Sartenaer (1966a). 
This zone is equivalent to Stevenson’s unit D (1945), 
the lower 2.5 m (8 ft) of the type section. 


These five occurrence zones hold up reasonably well 
in the San Andres Mountains wherever the upper beds 
have not been removed by later erosion. Several col- 
lections, especially those made in the earlier days and 
those collected from slope intervals where there may 
be mixing from the higher beds, contain elements of 
two or more of these intervals. 

A similar zonation is found in the Sacramento 
Mountains, although species abundances differ and the 
uppermost coral beds are nowhere found. The Hy- 
pothyridina beds, near the middle of the formation, 
provide a good marker horizon throughout the Sacra- 
mentos and, where they occur at the top of the for- 
mation—as at Alamo Peak and Indian Wells Canyon— 
they demonstrate the depth of post-Sly Gap erosion in 
this area. 

In the Mud Springs Mountains section, the diversity 
of species is considerably less than in areas to the east. 
Nevertheless, a number of species in the upper 6 m 
(20 ft) can be related to those of the lower brachiopod 
zone of the San Andres Mountains. In the Mud Springs 
Mountains, pre-Pennsylvanian erosion (perhaps sev- 
eral episodes) has removed both the upper brachiopod 
zone and the uppermost colonial coral zone. 


At Hermosa (PI. B, fig. 2), only four brachiopod 
species were collected from near the top of the Sly 
Gap beds, but these strongly suggest a sparse repre- 
sentation of the upper brachiopod zone. 


BRACHIOPOD ASSEMBLAGE OF THE 
Sty GAP FORMATION 


The total brachiopod assemblage of the Sly Gap 
consists of 38 genera and 66 species. Twenty-eight 
genera and 37 species are particularly useful in char- 
acterizing the paleontologic make-up of the assem- 
blage and in correlating it with other Frasnian faunas. 
An indication of the widespread and well-known dis- 
tribution of this assemblage, at the generic level, is 
that only one new genus is described in this paper, this 
even though many man-weeks were spent in the field 
and hundreds of specimens were studied during the 
description of this large number of species. 


AGE OF THE SLY GAP FORMATION 


The Sly Gap Formation is early Late Devonian 
(Frasnian-Cohoctonian Stage) in age on the basis of 
the rich brachiopod assemblage that was first de- 
scribed by Stainbrook (1948). While Stainbrook de- 
scribed 25 species, assigned to 24 genera, our collect- 
ing since 1948 has resulted in the accumulation of the 
large brachiopod assemblage listed in Table 2. Twenty 
of these species were previously described, 31 are new, 
and 16 are either too rare or too poorly preserved to 
warrant formal description. 

Although the Sly Gap brachiopods and the Indepen- 
dence Shale fauna of Iowa are related, as indicated by 
Stainbrook (1948), they are not as similar as it would 
appear from his earlier work (1945). 

The Sly Gap appears to be mid-Frasnian in age, ap- 
proximately equivalent to the Lime Creek Formation 
of Iowa. A number of common or related species in 
the Mount Hawk Formation of Alberta indicates cor- 
relation with a part of that formation (Text-fig. 8). 

Conodonts in the Sly Gap itself are rare and all that 
have been identified so far are non-diagnostic, long- 
ranging forms. An upper limit on the Frasnian age, 
however, is provided by the conodonts and brachio- 
pods of the overlying Contadero Formation. These 
brachiopods are clearly late Frasnian and conodonts 
from the basal Contadero Formation at Sheep Moun- 
tain indicate a maximum age range of Ancyrognathus 
triangularis through upper Palmatolepis gigas zones 
(J. W. Huddle, written comm., 1973; A. G. Harris, 
oral commun., 1977). 

The Frasnian cephalopod, Manticoceras, is report- 
ed from the middle part of the type section. Corals 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 19 


from the Sly Gap include: Macgeea Webster (1889b), 
Alveolites Lamarck (1801), ““Hexagonaria’’ Gtirich 
(1896), and Pachyphyllum Milne-Edwards and Haime 
(1850), all similar to the well-known Frasnian coral 
fauna of the Lime Creek Formation of Iowa. James 
Sorauf (State University of New York, Binghamton, 
NY), who has recently completed a detailed study of 
the Iowa coral fauna, is also studying the Frasnian 
corals of New Mexico. His research, when completed, 
will provide much detail on the nature of the coral 
fauna and its correlatives elsewhere in North America. 


CONTADERO FORMATION 


Type SECTION AND GENERAL 
STRATIGRAPHY 


The type section of the Contadero Formation, pro- 
posed by Stevenson (1945), is in Rhodes Canyon in 
the San Andres Mountains (loc. 526g). Although Ste- 
venson originally included all the post-Sly Gap beds 
in this formation, later refinements by Flower (1958, 
1959, 1965, 1969, 1970) place the higher beds in Flow- 
ers Thoroughgood and Rhodes Canyon formations. 


Thickness measurements in the type section vary 
slightly, but the Contadero is about 14 m (between 45 
and 50 ft) thick. Flower (1970) indicates that the re- 
vised Contadero is 13.5 m (45 ft) thick; we measured 
14.5 m (49 ft) of strata there that we consider to belong 
to the Contadero. The lower I1 m (36 ft) consists of 
alternating layers of dark gray shale and yellowish 
brown siltstone or fine-grained sandstone, and the bas- 
al 0.6 m (2 ft) is predominantly sandstone. There are 
about 20 thin silty or sandy units through this lower 
part of the formation. The next higher unit, about 1.2 
m (4 ft), is calcareous shale with many brachiopods 
and the uppermost 2.5 m (9 ft) consists of dark gray, 
nodular limestone containing large horn corals. 

The Contadero Formation also is present at the type 
section of the Sly Gap (PI. C, fig. 3), although Steven- 
son did not recognize it as such, in his 1945 paper. On 
Sheep Mountain, the Contadero consists of about 6 m 
(20 ft) of shale and siltstone with corals in the upper- 
most foot or so. The formation also is found to the 
south of Rhodes Canyon as far as San Andres Canyon 
and varies between 6 and 20 m (20 to 65 ft) thick 
throughout the area of exposure. 


Table 2.—Brachiopods of the Sly Gap Formation (+ = especially important elements). 


Philhedra species 
Petrocrania species unident. 

+  Schizophoria amanaensis Stainbrook 

+  Schizophoria iowensis (Hall) 
Schizophoria species 2 

+  Aulacella elliptica new species 

+ Cariniferella alleni new species 
Cariniferella iowensis Stainbrook 
C. species 

+ Rhipidomella species | 

+  Strophodonta umbonata new species 
S. species 2 

+ Strophonelloides dorsoconvexus new species 

+ S. parvulus new species 
Nervostrophia extensa new species 

+ N. geniculata new species 

+ N. mucrospina new species 
N. multinervosa Stainbrook 

+ N. solida new species 

+ Pseudodouvillina euglyphea Stainbrook 
Sulcatostrophia calvini (Miller) 
Douvillina arcuata (Hall) 

D. minuta new species 

+ D. parviuscula new species 
D. rectangulata new species 
D. species | 

+ Douvillinaria diversa new species 

+  D. variabilis (Calvin) 

Floweria prava (Hall) [new genus] 

+ F. magnacicatrix new species 
Eostrophalosia inexpectata new species 
Productella rugatula Stainbrook 

+ Devonoproductus intermedius new species 


D. vulgaris Stainbrook 
+  Gypidula bowsheri new species 
+ G. mimica new species 
+  G. subcarinata new species 
Athabaschia? species | 
+  Hypothyridina emmonsi rotunda new subspecies 
+  Coeloterorhynchus magnificum new species 
+  Calvinaria bransoni Stainbrook 
Hyborhynchella bransoni Cooper 
+  Pseudoatrypa devoniana (Webster) 
+  Costatrypa extensa new species 
+  C. varicostata (Stainbrook) 
+ 
+ 


TIowatrypa rara new species 
Spinatrypa trulla decorticata new subspecies 
Ambocoelia species | 
+  Tylothyris sulcocostata new species 
+  Acutatheca prolifica new species 
+  Thomasaria warreni new species 
+  Thomasaria demissa new species 
+  Cyrtospirifer cf. C. thalattodoxa Crickmay 
C. whitneyi (Hall) 
C. species 3 
C. species 4 
C. species 5 
Tenticospirifer cf. T. cyrtinaformis (Hall & Whitfield) 
Indospirifer anomala new species 
+ I. exfoliata new species 
I. aff. J. orestes (Hall & Whitfield) 
+ Elita magister new species 
+  Cyrtina inulta Stainbrook 
Cranaena calvini (Hall & Whitfield) 
C. species 2 


BULLETIN 315 


The Contadero Formation has not been recognized 
outside of the San Andres Mountains. 


FosstL CONTENT AND DISTRIBUTION 


Except for the undescribed coral assemblage at its 
top, the fauna of the Contadero is composed mainly 
of brachiopods. These are concentrated at two levels, 
one in the basal 3 m (10 ft) or so and the other near 
the top, just below and including the coral-bearing beds. 
No particular zonation is seen; the several common 
species appear to be present throughout the formation. 
Only four species range upward from the Sly Gap: 
Aulacella elliptica n. sp., Coeloterorhynchus magni- 
ficum n. sp., Cyrtospirifer whitneyi (Hall, 1858) and 
Cyrtina inulta Stainbrook (1945). Two new genera ap- 
pear in this assemblage: Stainbrookia and Colophrag- 
ma. 

As indicated in the discussion of the age of the Sly 
Gap, conodonts were found in the lowermost bed of 
the Contadero, and are helpful in establishing a datum 
for the ages of both units. 

Both the Sly Gap and Contadero coral assemblages 
are under study by J. Sorauf (State University of New 
York, Binghamton, NY). 


BRACHIOPOD ASSEMBLAGE OF THE 
CONTADERO FORMATION 


There are fewer brachiopod species in the Conta- 
dero than in either the Onate or Sly Gap formations, 
but the assemblage of the Contadero is about twice as 
diverse as those of the Thoroughgood and Rhodes 
Canyon formations. 

The Contadero brachiopod assemblage consists of 
20 genera and 24 species, of which nine genera and 11 
species serve to distinguish it from other Devonian 
assemblages in New Mexico. Three new genera and 
10 new species are recognized. 


AGE OF THE CONTADERO FORMATION 


The brachiopods clearly indicate that the Contadero 
is late Frasnian in age. The presence of true atrypids, 
with Douvillina Oehlert (1887) and a group of distinc- 
tive rhynchonellid genera, indicates that it is pre-Fa- 
mennian. The abundance of Cyrtospirifer whitneyi 
(Hall, 1858) and Uchtospirifer? rarus n. sp. suggests 
that the assemblage is approximately a correlative of 
the lower part of the Cyrtospirifer Zone, which spans 
the late Frasnian-early Famennian interval in the west- 
ern part of the United States (Text-fig. 8). 

As indicated in the discussion of the age of the Sly 
Gap, the late Frasnian conodonts at the base of the 
Contadero support the late Frasnian age shown by the 


brachiopods. Although no precise upper limit can be 
placed on the age of the Contadero, the presence of 
two species of Spinatrypa in the upper part of the 
formation indicates that it probably is entirely Fras- 
nian. No atrypids are known from Famennian strata 
anywhere in the world. Therefore, the Contadero can- 
not be placed in the Famennian as Poole et al. (1967, 
p. 887) indicate in their correlation chart. 


THOROUGHGOOD FORMATION 


Type SECTION AND GENERAL 
STRATIGRAPHY 


The type section of the Thoroughgood Formation is 
on Sheep Mountain (Flower, 1958, 1969, 1970) where 
it consists of 3 to 4 m (10-12 ft) of beds that lie above 
the Contadero (loc. 522). Along with the Contadero 
strata, these beds were also originally a part of Ste- 
venson’s Sly Gap Formation (1945), and his units K 
and L are now included in the Thoroughgood (PI. C, 
fig. 4). A thin remnant of the formation is preserved 
in Rhodes Canyon and Deadman Canyon to the south. 
In each place, only about 15 to 20 cm of sandstone 
remain, but this contains early Famennian conodonts. 

The Thoroughgood is essentially a light gray, fine- 
grained sandstone that weathers yellowish brown. It 
is calcareous in places. The basal few cm contains 
phosphatic pebbles and, although Stevenson (1945) re- 
fers to it as the “‘fish zone’’, no fish remains were 


Table 3.—Brachiopods of the Contadero Formation (+ = espe- 
cially important elements). 


Barroisella species 
Orbiculoidea species 1| 
Schizophoria species 
Aulacella elliptica new species 
+  Stainbrookia rotunda new genus and species 
Thiemella quadrata new species 
+  Douvillina aff. D. cayuta (Hall) 
+ D. species 2 
+ Floweria aff. F. chemungensis (Conrad) [new genus] 
Productella? species 2 
+  Cupularostrum? stevensoni new species 
+  Ripidiorhynchus newmexicoense new species 
Coeloterorhynchus magnificum new species 
+ Colophragma ellipticum new genus and species 
C. species | 
+  Spinatrypa compacta new species 
+ §. obsolescens new species 
Ambocoelia species 2 
+  Tylothyris novamexicana Stainbrook 
T. species | 
Cyrtospirifer whitneyi (Hall) 
+  Uchtospirifer? rarus new species 
Cyrtina inulta Stainbrook 
Cranaena navicella (Hall) 


DEVONIAN BRACHIOPODS OF NEw MEXICO: COOPER AND DUTRO 21 


found by us. The formation has not been identified 
elsewhere in south-central New Mexico. 


Fossitt CONTENT AND DISTRIBUTION 


Only one collection of megafossils has come from 
the Thoroughgood (loc. 522) and this is from the low- 
ermost part. In addition to the small brachiopod as- 
semblage, an indeterminate orthoconic nautiloid ceph- 
alopod and a loxonematid gastropod were also found 
at this level (Flower, 1970). 


BRACHIOPOD ASSEMBLAGE OF THE 
THOROUGHGOOD FORMATION 


This small, though distinctive, group of brachiopods 
includes nine genera and 11 species of which five gen- 
era and six species serve to characterize the assem- 
blage for biostratigraphic and age considerations. Col- 
ophragma and Crinisarina are new, as are three 
species. 


AGE OF THE THOROUGHGOOD FORMATION 


None of the characteristic Frasnian species found 
in the Sly Gap and Contadero formations carries over 
into the Thoroughgood. The presence of abundant 
Cyrtospirifer Nalivkin (in Fredericks, 1924) and Cri- 
nisarina n. gen., together with Evanescirostrum Sar- 
tenaer (1965), and the absence of atrypids indicate an 
early Famennian age (Text-fig. 8). According to Sar- 
tenaer (1969), Evanescirostrum does not occur before 
the Famennian. 

Conodonts in the thinned remnant of the Thorough- 
good in Deadman Canyon in the San Andres Moun- 
tains are of early Famennian age (J. Huddle, written 
commun., 1974). 

Apparently, the Thoroughgood and Rhodes Canyon 
formations of the San Andres Mountains are approx- 
imate equivalents to the Ready Pay Member of the 
Percha Formation, all three units being of early Fa- 
mennian age. 


Table 4.—Brachiopods of the Thoroughgood Formation (+ = es- 
pecially important elements). 


Productella species 3 
Athabaschia? species 2 
Colophragma species 2 [new genus] 
+  Camarotoechia latisinuata new species 
+ Evanescirostrum species | 
Ambocoelia species 3 
+  Tylothyris compacta new species 
+  Cyrtospirifer arenarius new species 
+  C. species | 
C. species 2 
+  Crinisarina reticulata (Stainbrook) [new genus] 
Afilasma species 


RHODES CANYON FORMATION 


TYPE SECTION AND GENERAL 
STRATIGRAPHY 


The type section of the Rhodes Canyon Formation 
is in Rhodes Canyon (locs. 509t, u) where Flower (1958, 
1959, 1969, 1970) applied the name to about 18 m (60 
ft) of dark gray, predominantly non-calcareous shale 
that lies above the thin remnant of the Thoroughgood 
Formation. The lower 3 m (10 ft) is mainly silty, dark 
gray, non-calcareous shale with a few brachiopods in 
the upper 25 cm. The overlying 6 m (20 ft) is fissile, 
non-calcareous shale with interbeds of siltstone. This 
interval is very sparsely fossiliferous. The next highest 
interval, about 3 m (10 ft) thick, is gray, calcareous 
siltstone and silty shale with abundant brachiopods. 
Finally, at the top, there is about 6 m (20 ft) of slightly 
calcareous silty shale and dark gray, fissile clay shale 
with a few brachiopods, as below. 

The Rhodes Canyon Formation has been definitely 
identified elsewhere only at Ash Canyon, to the south 
of the type area. Beds considered to be post-Conta- 
dero, but Devonian, by Kottlowski et al. (1956) in 
Hembrillo Canyon are Mississippian in age. 

Based on stratigraphic position, age relationships and 
general lithic appearance, the Rhodes Canyon For- 
mation is probably correlative to all, or a part of, the 
Ready Pay Member of the Percha Formation. Con- 
sequently, the Rhodes Canyon and Thoroughgood for- 
mations are interpreted as more shoreward facies of 
the dominantly unfossiliferous Ready Pay dark shales 
that are indicative of deeper or stagnant water. 


Fossit CONTENT AND DISTRIBUTION 


Brachiopods are common in the Rhodes Canyon, 
but there is little differentiation in the fauna from bot- 
tom to top of the unit. Collections from the bottom 2 
m (6 ft) are similar to those from the calcareous shales 
higher up and to the highest collection, about 1 m (3 
ft) below the top of the type section. 

No fossils other than brachiopods were collected 
and no conodonts have been identified from this for- 
mation. 


BRACHIOPOD ASSEMBLAGE OF THE RHODES 
CANYON FORMATION 


The brachiopod assemblage consists of 11 genera 
and 16 species. Seven genera and eight species are 
considered significant for characterization of the as- 
semblage. Particularly abundant are the spiriferids: 
Cyrtospirifer pyramidalis n. sp., Tylothyris novamex- 
icana Stainbrook (1947), and Crinisarina reticulata 
(Stainbrook, 1947). The last is assigned to a new genus 


BULLETIN 315 


of athyrids that is common in other Famennian assem- 
blages in western North America. Also important are 
the rhynchonellids, including Evanescirostrum gla- 
braventrum (Stainbrook, 1947), Ripidiorhynchus new- 
mexicoense n. sp., and Cupularostrum? stevensoni n. 
sp. The latter two species are long-ranging forms, but 
the first is restricted to this formation. 


AGE OF THE RHODES CANYON FORMATION 


Brachiopods indicate a definite Famennian age, 
probably mid-Famennian. Crinisarina n. gen. is pro- 
posed to include a group of species that are common 
in middle Famennian strata throughout the western 
part of North America. Species in the new genus pre- 
viously have been referred to Athyris or Cleiothyri- 
dina by many writers. Species closely related to the 
Rhodes Canyon form are found in the upper Palliser 
Formation in Alberta, the Trident Member of the Three 
Forks Formation in Montana, the Port Refugio For- 
mation of Alaska, and the lower Pilot Shale in Nevada. 
The occurrence of Evanescirostrum Sartenaer (1965) 
with Crinisarina also strengthens the Famennian cor- 
relation (see Sartenaer, 1969). Consequently, we feel 
that the Rhodes Canyon assemblage is approximately 
equivalent to the early Platyclymenia Zone (Text-fig. 
8). 

Sartenaer (1969, fig. 3) correlated the Rhodes Can- 
yon with the Box Member of the Percha Shale, making 
both units late Famennian in age. The brachiopods of 
the Rhodes Canyon have little in common with those 
of the Box Member and, while we agree with Sarte- 
naer that the Box Member is probably early late Fa- 
mennian, we think the Rhodes Canyon is older than 
the Trident Member of the Three Forks. 


Table 5.—Brachiopods of the Rhodes Canyon Formation (+ = 
especially important elements). 


+  Aulacella elliptica new species 
+ A. manticula (Stainbrook) 
“Schuchertella’’ species undetermined 
Productella species | 
+ Cupularostrum? stevensoni new species 
Ripidiorhynchus newmexicoense new species 
+ Evanescirostrum glabraventrum (Stainbrook) 
Crinisarina reticulata (Stainbrook) [new genus] 
+ Tylothyris novamexicana Stainbrook 
T. species | 
Cyrtospirifer breviposticus Stainbrook 
+ C. pyramidalis new species 
C. species indeterminate 
+ Afilasma inexpectatum new species 


Cyrtina species indeterminate 


PERCHA FORMATION (READY PAY MEMBER) 
(PI. D, fig. 2) 


Type SECTION AND GENERAL 
STRATIGRAPHY 


The type section of the Percha Formation was cho- 
sen by Stevenson (1945) although the formation had 
been named years earlier by Gordon (1907). As Ste- 
venson explained, Gordon did not select a type section 
but indicated that the type locality was on Percha Creek 
in the vicinity of Hillsboro, New Mexico. As described 
by Stevenson, the Ready Pay Member consists of about 
36 m (120 ft) of black, fissile shale with occasional 
green shale lenses, that grades upward into about 3.6 
m (12 ft) of gray, argillaceous shale. Stevenson re- 
ports, and our examination of relationships in the type 
area confirms, that the Ready Pay grades upward into 
the calcareous nodular shale and limestone of the low- 
er Box Member. 

Throughout the type region, the Ready Pay lies un- 
conformably on the Fusselman Dolomite of Silurian 
age. In several places we collected samples of the 
sandy, phosphatic basal beds that occur just above the 
unconformity. In all instances where definitive con- 
odonts were found, they represent early Famennian 
conodont zones or ranges that are approximately the 
same as those reported from just above the Sly Gap 
Formation in the Hermosa section. There is no ques- 
tion that all of the Percha Shale is younger than Fras- 
nian and, thus, younger than the Sly Gap and Conta- 
dero formations of the San Andres Mountains. 

Shales with conodonts of the same age are reported 
from three other places in south-central New Mexico 
(Huddle, written commun., 1973, 1974). At two local- 
ities in the southern Caballo Mountains, shale that is 
the equivalent of the Ready Pay lies unconformably 
on the Onate Formation. The third locality is in Dead- 
man Canyon, in the Sacramento Mountains. Here the 
conodonts occur in a thin black shale unit (about 1.5 
m thick) that lies above the Sly Gap and below the 
Mississippian Caballero Formation. 

As indicated above, conodonts at the base of the 
Thoroughgood Formation are also of early Famennian 
age. Thus, the Thoroughgood and Rhodes Canyon for- 
mations are approximate correlatives of the Ready Pay 
Member. In none of the eastern areas, however, is an 
equivalent of the Box Member found. 


Fosstt CONTENT AND DISTRIBUTION 


Except for the conodonts mentioned above, fossils 
are extremely rare in the Ready Pay Member. Three 
species of brachiopods (Retichonetes obscurus n. sp., 
Paurorhyncha cooperi Stainbrook (1947) and Syrin- 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DUTRO M33) 


gospira prima Kindle (1909)) were found in the upper 
part of the member at Bear Mountain, north of Silver 
City. All also are present in the brachiopod assem- 
blage of the Box Member. A fourth species, Linguli- 
pora porifera n. sp., was found only in the basal beds 
of the Ready Pay Member about 1.5 mi south of the 
Wilson Ranch in the Hillsboro (15’) quadrangle (loc. 
3037). 

A few scraps of plant material were seen in the low- 
er m or so of silty beds at the Bear Mountain locality, 
but they are indeterminate. 


BRACHIOPOD ASSEMBLAGE OF THE 
READY PAY MEMBER 


Only six species of brachiopods have been collected 
from the Ready Pay Member. The species of Lingu- 
lipora in the basal beds confirms the shallow-water 
origin of those silts, the initial deposits in the Famen- 
nian onlap. The other species in the upper part, being 
common elements of the overlying Box Member as- 
semblage, reflect the gradational nature of the contact 
between the two members. 


AGE OF THE READY PAY MEMBER 


Conodonts from several localities in the basal sandy 
and silty beds of this member indicate that it is no 
older than early Famennian (Text-fig. 8). Four collec- 
tions studied by Huddle (written commun., 1973, 1974) 
yielded abundant conodonts and he suggested that, 
although various elements of the assemblage indicate 
a range from the crepida to the lower marginifera 
zones, a few short-ranging forms indicate that the lat- 
ter age is more likely. 

The conodonts indicate that the Ready Pay is prob- 
ably a black facies of the Thoroughgood—Rhodes Can- 
yon formations, not of the Sly Gap as proposed by 
Poole et al. (1967, p. 887). 

Goniatites from the middle or upper parts of the Box 
Member were discussed by House (1962) who suggests 
that they indicate an age no older than the upper Platy- 
clymenia Zone. His comparison of a large specimen 
of Cyrtoclymenia Hyatt (1884) with one from the Cly- 
menia-Stufe at Dasburg implies that the age may be 


Table 6.—Brachiopods of the Percha Formation (Ready Pay 
Member) (+ = especially important elements). 


+  Lingulipora porifera new species 
+ Retichonetes obscurus new species 
+  Paurorhyncha cooperi Stainbrook 
Petasmaria patens new genus and species 
+ Syringospira prima Kindle 
Cyrtiorina kindlei (Stainbrook) [new genus] 


slightly younger. In any case, the goniatites from the 
Box Member point to a correlation slightly younger 
than the Platyclymenia Zone assemblage of the Three 
Forks Formation. This would be the equivalent of the 
lower or middle styriacus Zone, in terms of the con- 
odont zonation. House (1962, p. 262) states, ““The Per- 
cha ammonoids suggest a slightly younger age than 
those of the Three Forks Shale.’ 


PERCHA FORMATION (BOX MEMBER) 
(Pl. D, figs. 1, 3, 4) 


TyPE SECTION AND GENERAL STRATIGRAPHY 


The type section location was discussed above, rel- 
ative to that of the Ready Pay Member. The Box 
Member, at the type section (loc. 507h) is 14 m (46.5 
ft) thick and is composed essentially of gray calcar- 
eous shale that contains fine-grained limestone nod- 
ules and a few hard gray limestone beds (PI. D, fig. 
1). The member is dominantly shaly, as at the type 
section, at several sections near Kingston, George- 
town, and Hillsboro in the Black Range region, and 
thicknesses vary from about 9 m (30 ft) to 15 m (50 
ft). The entire formation thickens westward to the vi- 
cinity of Silver City where the Box Member is about 
20 to 22 m (70-75 ft) thick and the Ready Pay Member 
is about 70 m (230 ft) thick. The Box Member also 
becomes more calcareous westward with an increase 
in the concentration of limy nodules and thin beds of 
limestone. 

The gradational nature of the contact between the 
two members of the Percha is well displayed at Bear 
Mountain, north of Silver City. The base of the Box 
Member is placed at the bottom of a 3 m (10 ft) unit 
of dark shale with limy nodules that are characteristic 
lithic components of the member. This unit yields a 
number of the brachiopods characteristic of the Box 
assemblage, including Paurorhyncha Cooper (1942), 
Cyrtiorina n. gen., Syringospira Kindle (1909), pro- 
ductids, and a few horn corals. Immediately below, at 
the top of the Ready Pay, is about 6 to 9 m (20-30 ft) 
of dark gray to black, non-calcareous, unfossiliferous 
shale. But below this unit, and above the main mass 
of the black Ready Pay shale, is about 3 to 4 m (10- 
15 ft) of gray, slightly calcareous shale with a few cal- 
careous nodules that contain Syringospira and Pau- 
rorhyncha. 

The Box Member is disconformably overlain by 
Mississippian strata. In some places, as at the type 
section described by Stevenson (1945), the basal Mis- 
sissippian strata are yellowish-weathering, calcareous 
siltstones with phosphate pebbles, conodonts and trace 
fossils (including Zoophycos Massalongo (1855)). In 


BULLETIN 315 


other places, the nodular limestone of the Lower Mis- 
sissippian Caballero Formation lies directly on the Box 
Member. Usually, a lag concentrate or pebbly con- 
glomerate bed can be found at the contact and this 
zone has yielded early Mississippian (late Kinder- 
hookian) conodonts in many places. 


FossiL CONTENT AND DISTRIBUTION 


The Box Member has yielded a diversified assem- 
blage of invertebrate fossils, of which the majority are 
brachiopods. However, there are horn corals, bryo- 
zoans, molluscs (including pelecypods, gastropods, 


Table 7.—Brachiopods of the Percha Formation (Box Member) 
(+ = especially important elements). 


Lingula species | 
Lingula species 2 
Orbiculoides species 2 
Petrocrania ourayensis (Kindle) 
+  Schizophoria australis Kindle 
+  Thiemella platys new species 
+  Schuchertella percha (Stainbrook) 
S. resupinata new species 
Retichonetes obscurus new species 
R. perchaensis new species 
Acanthatia nupera (Stainbrook) 
Leioproductus plicatus (Kindle) 
L. defessa (Stainbrook) 
L. coloradensis (Kindle) 
L. pauperculus new species 
L. varispinosus (Stainbrook) 
Galeatella galeata Muir-Wood and Cooper 
Planoproductus hillsboroensis (Kindle) 
Laminatia laminata (Kindle) 
Sentosia praecedens (Stainbrook) 
S. praecursor (Stainbrook) 
S. spinigera (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Trifidorostellum obscurum new species 
Camarotoechia? species 
Paurorhyncha cooperi Stainbrook 
Eoparaphorhynchus mclareni Sartenaer 
Porostictia perchaensis (Stainbrook) 
Petasmaria compacta new genus and species 
P. multicosta new species 
P. patens new species 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Ambocoelia capillata new species 
+  Strophopleura notabilis (Kindle) 
Cyrtospirifer? animasensis (Girty) 
+  Cyrtospirifer convexus new species 


+++ 


++t+ett¢e4+4+ 


ae 


+++ 4+ +++ 


+ C. rotundus new species 
Hispidaria posterogranulosa new genus and species 
H. spinosa new species 
Cyrtiorina kindlei (Stainbrook) [new genus] 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 


Oiosia putilla (Stainbrook) [new genus] 


cephalopods and scaphopods), Hexacrinus Austin and 
Austin (1844) and echinoderm debris. All the samples 
processed for conodonts proved to be barren thereof. 

Most species of the Box Member range from top to 
bottom. Some apparent exceptions were noted, how- 
ever. Syringospira prima Kindle (1909), which is pres- 
ent in the uppermost part of the Ready Pay, is most 
common in the bottom few m of the Box Member. 
Productids are extremely abundant and varied in the 
lower 6 to 9 m (20-30 ft) in the area around Silver 
City. The lower part of the member also contains 
abundant Cyrtiorina and Paurorhyncha cooperi Stain- 
brook (1947); and Composita bellula Stainbrook (1947) 
seems to be limited to the lower quarter of the Box 
Member of the Percha. 

In many sections, the size of the individuals in- 
creases upward through the member, although the 
number of specimens decreases accordingly. One par- 
ticularly distinctive species, Paurorhyncha cooperi 
Stainbrook (1947), was collected from the upper Ready 
Pay Member but ranges through the Box Member and 
was found within 25 cm of the top at one locality east 
of Kingston, New Mexico. 


BRACHIOPOD ASSEMBLAGE OF THE 
Box MEMBER 


This rich and diversified assemblage consists of 30 
genera and 44 species, of which 25 genera and 32 
species are characteristic. Many of the species were 
described by Kindle (1909) and Stainbrook (1947) and 
most of the productid genera were established by Muir- 
Wood and Cooper (1960). However, four new genera 
are described here: Petasmaria, Hispidaria, Cyrtiori- 
na, and Oiosia. In addition, 12 new species are de- 
scribed. 


AGE OF THE BOX MEMBER 


The lower limit on the age of the Box fauna, as 
discussed above, is suggested by the occurrence of 
goniatites of the upper Platyclymenia Zone. The bra- 
chiopod fauna is definitely younger than that described 
from the West Range Limestone (Johnson ef al., 1969) 
and is older than uppermost Devonian assemblages in 
the Louisiana Limestone and correlative units in west- 
ern North America. Sartenaer (1969) shows ranges for 
Famennian rhynchonellid genera that indicate an age 
no older than the upper Platyclymenia Zone for the 
combination of genera found in the Box Member. This, 
of course, agrees with the meager goniatite data. 

On the mistaken notion that the Box Member was 
Mississippian, Stainbrook (1947) supplied several of 
the productids with Mississippian generic names and 
also included in his faunal list some float specimens, 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 


such as Leptagonia M’Coy (1844) and Buxtonia 
Thomas (1914) from the overlying Mississippian and 
Pennsylvanian. The productids all proved to be pro- 
ductellids and are Devonian in age. 

The mutual occurrence of Paurorhyncha Cooper 
(1942), Porostictia Cooper (1955), Ptychomalotoechia 
Sartenaer (196la), and Trifidorostellum Sartenaer 
(1961la) suggests to us that the age of the Box fauna is 
probably restricted to the Clymenia Zone, in a general 
way (Text-fig. 8). This is equivalent to a range from 
the upper part of the styriacus Zone to the middle part 
of the costatus Zone, in terms of the conodonts. 

Bryozoans from the Box Member were described 
by Fritz (1944). All the species were described as new 
and were assigned to seven genera. Because of a lack 
of comparably described assemblages, however, Fritz 
was unable to make any pertinent statement about the 
age of the Box bryozoans. 


CONODONT BIOSTRATIGRAPHY 


We are greatly indebted to the late J. W. Huddle, 
of the U.S. Geological Survey, who processed, iden- 
tified and analyzed the conodonts in samples taken 
during this study. Because conodont faunas from the 
Devonian of New Mexico have not been discussed in 
any detail, we include Huddle’s determinations and 
remarks here. As indicated in the foregoing biostrati- 
graphic sections of this paper, the conodonts have pro- 
vided valuable clues to the age and correlation of most 
of the Devonian units in New Mexico. In addition, 
many of the enigmatic sequences of black shale in the 
region have been placed in stratigraphic perspective 
as a result of Huddle’s studies. Huddle’s remarks are 
enclosed in quotation marks to distinguish them from 
Our Own interpretations. 

We also acknowledge the assistance of A. G. Harris, 
of the U.S. Geological Survey, who brought some of 
the nomenclature up-to-date and sharpened some of 
the age assignments. 

The collections are discussed below, from oldest to 
youngest, and their significance indicated. All collec- 
tions that contain identifiable conodonts have been 
given U.S.G.S. locality numbers in the Silurian-De- 
vonian catalogue (e.g., USGS 9207-SD) and are stored 
with the USGS conodont reference set at the U.S. 
National Museum of Natural History in Washington, 
D.C. 


CONODONTS FROM THE ONATE FORMATION 


Three of the four collections from the Onate yielded 
conodonts and one of these contains Polygnathus var- 
cus Stauffer (1940), indicating that the formation can 
be assigned to the varcus Zone in the broad sense. 


No 
nn 


Other evidence cited above leads us to place the Onate 
in the middle varcus Subzone. 


USGS locality 9312-SD (loc. 522] herein), Onate Formation, Mud 
Spring Mtns., basal 19 ft of the formation. Collectors: Cooper, Allen 
and Bowsher, 7/22/51. 


No. specimens 


Polygnathus linguiformis Hinde (1879) 5 
Polygnathus varcus s.l. Stauffer (1940) 5 


“The presence of P. varcus suggests that this collection is not 
older than late Givetian. The oldest occurrences of this species in 
eastern U.S. are in the Centerfield Ls. in New York and the Beech- 
wood Ls. in Indiana, and equivalent rocks in the middle West.”’ 


USGS locality 9208-SD (loc. 3023b herein), Onate Formation, John- 
son Park Canyon (Mockingbird Gap). Collectors: Bowsher and Al- 
len. 


No. specimens 


Icriodus nodosus (Huddle, 1934) 5i7/ 
Neoprioniodus sp. 

Polygnathus linguiformis Hinde (1879) 
Polygnathus sp. 


i 


“P. linguiformis and I. nodosus both range from Middle Devonian 
to early Late Devonian. The abundance of /criodus suggests a shal- 
low-water environment.” 


USGS locality 9220-SD; Onate Formation, 200 yd S. of bend in road 
in Johnson Park (Mockingbird Gap), northern San Andres Mtns.; 
1 ft above base of formation. Collector: Bowsher, 1956. 


Icriodus sp. 3 specimens 


“‘Icriodus is common only in the Devonian. Abundant in shallow- 
water deposits.” 

A fourth collection from the Onate Formation in the 
Rhodes Canyon area, San Andres Mountains, yielded 
no conodonts. 


Another collection, from fine-grained sandstone just 
above the Fusselman Dolomite near Lone Mountain, 
south of Hillsboro, yielded conodonts of about this 
same age. Although there was no lithic indication of 
the presence of the Onate Formation when the area 
was examined, the conodonts indicate that a lag con- 
centrate of Onate age was preserved. This is one of 
two places in the region where we know of pre-Fa- 
mennian conodonts from beds at the base of the Ready 
Pay Member that could have been reworked from old- 
er strata. 


USGS locality 9214-SD; field No. M24(1). Lone Mtn. area; fine- 
grained sandstone |-2 in above Fusselman Dolomite. Collectors: 


Cooper and Dutro, 3/24/64. 


No. of 
specimens 


Polygnathus linguiformis Hinde (1879) 3 
Polygnathus tuberculatus? Bryant (1921) 1 


26 BULLETIN 315 


. the surfaces show only a few conodont fragments. P. lin- 
guiformis and P. tuberculatus range from Middle Devonian to early 
Frasnian.”” 


CONODONTS FROM THE SLY GAP FORMATION 


Three collections from the Sly Gap yielded only bro- 
ken, unidentifiable fragments of conodonts to which 
Huddle could assign no age significance. 


CONODONTS FROM THE CONTADERO 
FORMATION 


Two collections from the Contadero yielded con- 
odonts, assigned to the late Frasnian or, perhaps, early 
Famennian by Huddle. A reassessment of these faunas 
by Harris (oral commun., 1977) makes it likely that 
they represent the Ancyrognathus triangularis Zone 
(middle Frasnian). A more detailed knowledge of the 
ranges of conodont species in Frasnian strata of west- 
ern Europe, developed during the past few years, 
makes possible this more precise dating. The clearly 
late Frasnian brachiopod assemblage of the Contadero 
is nicely bracketed by dated conodont collections. 


USGS locality 9215-SD; San Andres Mtns., SE'44NE sec. 26, T. 
11 S., R. 5 E.; type section of Sly Gap Fm. on Sheep Mtn.; basal 
beds of Contadero Fm. Collectors: Cooper and Dutro, 4/7/64. 


No. specimens 


Ancyrodella nodosa Ulrich & Bassler (1926) 14 
Ancyrognathus species 1 
Bryantodus species 1 
Hibbardella species 2 
Ligonodina species 3 
Palmatolepis species 17 
Polygnathus species 16 
Poly. decorosus? Stauffer (1940) 11 


‘The fragments are too small to be certain, but it is probable that 
the following species are present: Palm. subrecta and Polygnathus 
homoirregularis.’’ Harris indicates that the mutual occurrence of 
Ancyrodella nodosa, Ancyrognathus sp. and Palm. subrecta make 
an assignment to the Ancyrognathus triangularis Zone a reasonable 
one. According to Klapper (oral commun., 1981) the assemblage 
could range from the Ancyrognathus triangularis Zone through the 
Palm. gigas Zone, and a slightly younger age would fit better with 
the correlations based on brachiopods (Text-fig. 8). In any case, this 
appears to be a late Frasnian assemblage. 

One collection made by R. H. Flower from 40-45 ft 
above the base of the Contadero in the Rhodes Canyon 
section (collection not numbered) contains a broken 
specimen of Ancyrognathus, probably A. triangularis 
Youngquist, according to Huddle (written commun., 
1968). 


CONODONTS FROM THE THOROUGHGOOD 
FORMATION 


One collection from the thin remnant of Thorough- 
good in Deadman Canyon, San Andres Mountains, 
yielded a few conodonts that are significant in that 
they suggest correlation with the basal Ready Pay 
Member to the west. The Famennian age of the bra- 
chiopods of both the Thoroughgood and Rhodes Can- 
yon formations is not in doubt and this conodont date, 
although qualified because of poor preservation, pro- 
vides a valuable link in the regional stratigraphic 
scheme. 


{no locality number]. San Andres Range, Deadman Canyon; center 
NW'‘4SE% sec. 12, T. 17 S., R. 3 E.; bed N (fish bed) of Thor- 
oughgood Formation. Collectors: Cooper and Dutro, 4/9/64. 

‘There are broken and altered conodonts in the sandstone. The 
5 recognizable fragments seem to be Icriodus, Polygnathus, perhaps 
P. semicostatus, and another species of Polygnathus.” 


Although Huddle did not venture an age assignment, 
P. semicostatus occurs in collections from USGS lo- 
calities 9210-SD and 9213-SD, both of which were as- 
signed an early Famennian age by Huddle (written 
commun., 1973). The species is long-ranging in the 
Famennian, but is unknown below the rhomboidea 
Zone. 


CONODONTS FROM THE READY PAY MEMBER 
OF THE PERCHA SHALE 


Perhaps the most significant contribution of the con- 
odont studies of Huddle is the dating of the base of 
the Ready Pay Member. Three collections from the 
base of the member, two from near Wilson Ranch and 
one from the Cooks Range, indicate an early Famen- 
nian age. 

Although ranges of some of the more distinctive 
species, in terms of conodont studies elsewhere, ex- 
tend from the uppermost Palmatolepis triangularis 
Zone to the upper Palmatolepis marginifera Zone, 
overlapping ranges suggest that an assignment to the 
Palmatolepis rhomboidea Zone is reasonable. Two 
collections from the base of the black shale that 
overlies the Sly Gap Formation near Hermosa give a 
similar age assignment. As mentioned above, the base 
of the Thoroughgood in the San Andres Mountains can 
also be considered of about this age. 

Three collections from black shale above the Onate 
Formation in the southern Caballo Mountains yield 
conodonts of this same approximate age and can be 
interpreted to represent the Ready Pay. This provides 
a stratigraphic correlation between the type region of 
the Ready Pay to the west and the Famennian for- 
mations of the San Andres Range. 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DUTRO DoT 


A final collection that contains conodonts of early 
Famennian age comes from the meter or so of black 
shale that overlies the Sly Gap Formation in Deadman 
Canyon in the Sacramento Mountains. As this collec- 
tion also contains some reworked specimens of Fras- 
nian species, it probably indicates that the shale was 
deposited during a period in the early Famennian when 
Sly Gap and Contadero strata were being eroded. 

The species identified by Huddle (written commun., 
1973; 1974), with their occurrences in the several col- 
lections from New Mexico, and the ranges of the 
species where known elsewhere, are shown in Ta- 
ble 8. 


CONODONTS FROM BEDS OVERLYING 
THE DEVONIAN 


Four collections from beds above the Devonian yield 
early Mississippian conodonts. Three of these are from 
the Caballero Formation in the Kingston-Silver City 
area. The Caballero contains a diversified upper Kin- 
derhookian megafossil assemblage, so this conodont 
evidence is simply confirmatory. The fourth collec- 
tion, however, from Deadman Canyon is from sandy 
beds above the Contadero and just below the Caba- 
llero. Similar sandy beds, with trace fossils including 
Zoophycos, are found elsewhere below the Caballero. 
This Mississippian collection is, therefore, helpful for 
dating this pre-Caballero unit. 

Mississippian collections have been assigned num- 
bers in the U.S. Geological Survey Upper Paleozoic 
catalogue (for example: USGS locality 25242-PC) and 
are stored with the conodont reference set in the United 
States National Museum of Natural History, Wash- 
ington, D.C. 


USGS locality 25242-PC; field No. M27; Kingston quad. (15’); near 
center SW!4SE'4 sec. 14, T. 16 S., R. 27 W.; 1-2 in fish tooth bed 
at base of Mississippian. Collectors: Cooper and Dutro. 


No. 
specimens 

Polygnathus inornatus E. R. Branson (1934) 12 
P. longiposticus Branson and Mehl (1934b) I 
P. scobiniformis E. R. Branson (1934) 5 
P. symmetricus E. R. Branson (1934) 3 
Pseudopolygnathus primus Branson and Mehl (1934b) 3 
Siphonodella cooperi Hass (1959) 12 
5 


S. obsoleta Hass (1959) 
Spathognathodus aculeatus (Branson and Mehl, 1934a) 1 


‘‘All the conodonts in this collection are worn but they all indicate 
an age of middle or late Kinderhookian.” 


USGS locality 25243-PC; San Andres Mtns., Bear Canyon section; 
10-15 ft above base of Caballero Fm. Collectors: Cooper and Dutro. 


No. 
specimens 
Apatognathus sp. 1 
Elictognathus sp. l 
Gnathodus sp. 4 
Hindeodella sp. 4 
Ozarkodina sp. 4 
Polygnathus allocotus C. L. Cooper (1939) 1 
Poly. communis Branson and Mehl (1934b) 10 
Poly. symmetricus E. R. Branson (1934) | 
Siphonodella crenulata (C. L. Cooper, 1939) 5 
S. obsoleta Hass (1959) 5 
Spathognathodus sp. 4 
Synprioniodina sp. 3 


“The presence of §. crenulata indicates a late Kinderhookian age 
for the sample.’ 


USGS locality 25244-PC; Silver City quad., Bear Mtn. section; bed 
E, nodules at base of Caballero Fm. Collectors: Cooper and Dutro. 


No. specimens 


Hindeodella sp. 1 


Neoprioniodus sp. 1 


USGS locality 25245-PC; San Andres Mtn., center NW'4SE"% sec. 
12, T. 17 S., R. 3 E.; Deadman Canyon section, bottom 2 ft of bed 
O. Collectors: Cooper and Dutro. 


No. specimens 


Siphonodella sp. 5 
Pseudopolygnathus sp. 1 
Polygnathus sp. 1 


“This collection is early Mississippian in age.” 


Additional information on conodonts of these inter- 
vals may be found in W. I. Anderson (1966), C. L. 
Cooper (1931), Ellison (1950), and Ethington (1965). 


GEOLOGIC HISTORY 


The foregoing discussion of the stratigraphy and 
biostratigraphy of the Devonian of south-central and 
southwestern New Mexico suggests that a sequence 
of geological events can be described with some con- 
fidence. 

During the late Middle Devonian (late Givetian) the 
seas again covered southern New Mexico after a long 
period during which erosion of early Paleozoic rocks 
had taken place. The Onate Formation represents these 
first, shallow marine deposits, which were unconform- 
ably deposited on an irregular surface that was devel- 
oped on Silurian and Upper Ordovician rocks. 

Fine-grained, dolomitic clastic rocks characterize the 
Onate, which also contains many trace fossils and evi- 
dence of extensive bioturbation. The thin sequence in 


8 BULLETIN 315 


Table 8—Conodont occurrences in the basal Percha Formation, southwestern New Mexico, and related strata. USGS locality 9221-SD, 
base of Ready Pay Member, Cooks Range; USGS locality 9212-SD, base of Ready Pay Member, 0.75 mi SW of Wilson Ranch, Hillsboro 
area; USGS locality 9216-SD, base of Ready Pay Member, just W. of Wilson Ranch, Hillsboro area; USGS locality 9211-SD, base of Ready 
Pay Member, Hermosa A section: USGS locality 9217-SD, base of Ready Pay Member, Hermosa B section; USGS locality 9213-SD, black 
shale near Nakaye Mines, southern Caballo Range; USGS locality 9210-SD, black shale at Red House Mountain, southern Caballo Mountains; 
USGS locality 9207-SD, black shale at top of section, Deadman Canyon, Sacramento Mountains. An asterisk marks each species that may 
have been reworked from older strata. 


U.S. Geological Survey Localities 


Conodont Taxa 9221- 9212- 9216- 9211- 9217- 9213- 9210- 9207- 
SD SD SD SD SD SD SD SD 

FNCOY TIN ab cebene paneodb.sadess oanvneiedsEnoconcobb oboogduoGsessouppooaccrooRoSaEoapdatnnosaaccagrocbarppopabEoceconponacte unseen: l 

Ancyrodellaicurvaral(Bransoniands Mens O54) saree nesses eee eit asec sr eer er eer ae ree errr eee ree et Ge 1 

Ancyrodella Pods Ulrich and Bassler (1926)* ........... sds aR OOUR AERO BORE OR OCC aan cons 1? 

Ancyrodellaysps* x. cccuncmta hrc he une ae ae mea ese netted plete eelesie sie selslorasIe sake [Pe ceist vies toeowereett aetdtee memes Liisa ete steel ete SRE RES l 

Ancyrognathus asymmetricus (Ulrich and Bassler, 1926)*. 4 


Ancyrognathus sp.* yes saya Ree EDEL Sie mya eaieicia, 5 8 ncaTh aaa RV e Merete aie nan OE Eee a ae etree Rie Cee B Bama [eerie ic 1 
Rellodellais pres. serene eee eesti cee ae serene eee eC Ecc car chiter-tatoec asses <e ne ee eee 1 

Bryantoaotusicollingatus! (Buyante 92) peee-eeereeeeeeeee ec ee ee eReee rect erat tac eeee se ee eEeEreeeeeee RAMA rics son denarii sadooloboctaaas 1 ef 
TT AAHOUITIN Cos, gadaosno nee Bacdasnenssssee csubncedastded sanespedooogs seedecesstopspopodc Uae as aAEE AEB OONO Ose sconce eb baaddr Hepaenoos Hoseienone I 
Diplododella sp ras ccit- was essere ee eee LS a TA eT eee EIN ISToS SLL atais ealors eiaetate Pelee errata a steteeaks af eet eter ref aite e eite eee ere ee 1 

Jef NXMCAUAG )aeoccsannadoonssoosassseepo sss" opoG0NIOd osNab boo se Serooonadeenonaass ToD EKOURES sop HsOnS odo agdensaenauDnoscss Adbeb oO“ CGaAt sso cD 1 
Hindeodellaisprrecrn cece ieee tee ser eee eae Chea Rec ee er aera clan oa teins =a | RerPoonernaonnnne Discse putrs saat an Penney Seceaaee 20 
TOnIOAUSISD Waccsat oes ssc cece Oo neem eee aoe ee Meee sees Elke tesisnteisis seis else efels Cli ae ca etna Ot ee conuce Sean  enaree sdrcocd 2 

WIG ONOGINGIS Psectere en ier elosacis se seeateesteiess ese oe oes ee ase teesSeae aseeeoeeeeeCee 6055.) IsopbanesnsabopssbouonabotuctosSs050008 2 

Ia OX Hi) enobasuacne ipsa pesadacein deeaausapdragdaannneuns cco oObhavnocacsspuninaccotounbrossgscrgaun cote tn ippceSe caonaddonsashcmapnts ! 

NCO AROMAGH OG, csndanoounsnndessacbotcenes 340000 00ccaaDtn nde supGONNNASABOBODHONAS 20° Wossseancnoeanabeo6 4 

OZANKOAINGIS Pere Kees e ee gees stl eater ee ai latte aleratose = BEPEDOSC DC llgsexsaeg dager cec encase econ ane yeneecaber 4, eins 2 

IRA CHO FOS EU, 12, GTS SVAN CNIS (OTM) cvconeccccosnonbconbonesannecenanse ssenncaisdebacieauaceaonnatmonscccaere l 

Palmatolepis crepida? Sanneman (1955)................ 

Palmatolepisseigasy Mille rand ax O WMS GUIs ty (19457) mereterenrertetatelatstchetatetols fee als s/o) nl ctaletetetetetehetelcteletetateletelstefst sets 

Palmatolepis glabra Ulrich and Bassler (1926) ...............000000cseeee eee e ener ee 

Palmatolepis glabra acuta Helms (1963).................22.5-.055- sididlajaia a aicie'd er ctarmre ontalua ieee choca. Ae bye tektites sess eee eee CORES eee I 
Palmatolepisieabyargjabrawirichyand) Bassler (1926) eeeseereceesaceace eek ttasc eetseeeehe ce erect sere ctaeneeeet eee eet eee ae 10 
PalmatolepisyelabravleptavZreglerand ud dlei( 1969) irae ctetetsisiseiets-1e eeyeleinieisttelele iets elec aiete eel ciel tereieiensi ae sss ae eee eee eee eee 4 
Palmatolepis glabra prima Ziegler and Huddle (1969) .................. 0002 eee eee ee IS east ta Sipepeone (Bosssiserc 8 
Palmatolepis'glabrajpectinata Ziegler (W962) ec arain ate esate ele «2 = oie vis erence oes alae eee DG. cae hance v anton sale Se ec ee eee ee 14 
Palmatolepis minutajlobaiHelmsi (968) )ssceceeet sae seeest eee see eee ates 2 

Palmatolepis minuta minuta Branson and Mehl (1934)....................22.020055 3 

Palmatolepis perlobata Ulrich and Bassler (1926) ................... 0.0 cece ee scene 

Palmatolepis quadrantinodosa Branson and Mehl (1934) ...............00 000 cece eee eeee eee 

PalmatolepisiquadantnodosannplexapNuuller (956) eeee-eoeeree re ee ce sccacree eee eee eee teen ee eee ere eae eenee eee eee eee nee 

Palmatolepis quadrantinodosalobata Sanneman (1955) Bric 

PalmatalepisysubrectauMillerandpyounzquists((li94i/) ercrrtttercrsisisis siete ieeietetetsteiarstattstereelsteasiaetetetetetets 

Palmatolepis superlobata Branson and Mehl (1934) .................. 0.0 c cece eee 

Palmatolepis:tenuipunciaraysannemany (955) erases cere ere ee ees eileen oot eee esas aa testo eee eee ee seer eee dete eiteceeeetaetennteees 
PalmatolepisitriansularissSsannemany (955) jee ecese sete ceereas- eee eee econ ee CECE EE CEE CEE CE EER Ee eee eee Rec et 1 

Palmatolepis sp..........-- Bee Capea aoe oo coco aa IC iccanencaseadarsendbouuEtoaconacdse macuracurcdsonen ae ude dsonanaan Snr he ammeucomaeedadandaasecdcnnasc scccc 3 
Polygnathus brevilaminus Bransontand! Men) (1934) ) 2 os.-.ot-saynicissieciajs a.-/a's nis1olsiataisieiste sieynicietsrat=rater ote efeisislsleisis a1etcleiereistelat=ia 2 

Polysnathusidecorosuss tatters (1938) acces tees ok cece eee atte aes nee eae Meme ele ee eee ee 

Polysnathus glabersUlrichvand Bassler (i926). -css-neeeseceeiecteeete resellers : epemcce 1 
Polygnathus homoirregularis Ziegler? (1962)................. 

Polygnathus: linguiformiss Hinde. (i879) oem ccceve oe ne ceemciensiaem ame es 

Polygnathus nodocostatus Branson and Mehl (1934) ........... 

Polygnathus normalis Miller and Youngquist (1947)...............00..200scee cues 

Polygnathus perplexus Thomas (1949) ..... 

Polygnathus semicostatus Branson and Mehl (1934). HEdaccan 2 
Polygnathus sp. ae msisiatate Rave ais 2 

Polylophodonta confluens (Ulrich mae Bassler, 1926) Eber aaboncrne SyoAppososcantce asses yo 3 

Polylophodonta pergyrata (Holmes, V928) «as -.o.ssissecscersscsem isis ass.aags oa «hee nialeg sistaietaianteoslels + coer ab Oak Bee eer ee eae ctcteaete eet eee 1 
Polylophodonta sp. Au . a6 Reese o COSHER ARSC or cUcoeocoall onsormpnoncaonoddapovonnsncnAanndoodnodoorcbouuadsotemoaradecoueLn” 1 
Prioniodonta dialata (Bryant, 1921) oan sisi. clot ate eysielafet a sletetata waveratais| leveraveleic eteemGetaeeeictert deere eieIs welt BE ena OnenAOuOSpancececccodne 1 
Spathognathodus regularis\(Branson and: Meh, !1934))-iceicioss.5.c 8285. a eb visas sents Wiclctioto sis Aue + avevelelalate aisis cia tte tote MED Siete rere eletate ec te cee siete ete 8 


Synprioniodina sp. 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DUTRO 29 


the northern part of the San Andres Mountains con- 
tains abundant specimens of /criodus Branson and 
Mehl (1938), a conodont that characterizes shallow- 
water environments (Huddle, written commun., 1974). 
The general thickening of the sequence southward in 
the San Andres Mountains, and the increase in the 
amount of shale, suggests deepening of the original 
basin in that direction. The Canutillo Formation (Nel- 
son, 1940) of the Franklin Mountains is of probable 
Middle Devonian age and could be a deeper-water 
equivalent of the Onate Formation. 

West of the Rio Grande in the Mimbres Range, the 
type area of the Percha Shale, there is little evidence 
of pre-Famennian Devonian deposition in New Mex- 
ico. However, in the Swisshelm and Pedregosa Moun- 
tains of southeastern Arizona, the lower part of the 
Swisshelm Formation is an equivalent of the Onate 
(Epis et al., 1957). 

According to Epis et al. (1957, p. 2255-2256) two 
regional pictures can be developed for the Devonian 
deposits of Arizona and New Mexico. They discuss 
these briefly and prefer the simpler one, in which the 
Percha Shale is interpreted as a deeper-water facies of 
the Sly Gap and its equivalents. Unfortunately, our 
analysis of the faunas and age of the Devonian se- 
quence demonstrates that the Percha is, indeed, youn- 
ger than the Onate-Sly Gap-Contadero sequence. Con- 
sequently, the more complex history, involving onlap 
around a positive area in southwestern New Mexico, 
is the more likely explanation. 

Johnson (1970) presented an intriguing series of ar- 
guments for progressive onlap of the continental 
‘“‘backbone”’ during the Taghanician Stage. Insofar as 
New Mexico and Arizona are concerned, the evidence 
for onlap of this age is overwhelming, with the basal 
Onate strata lying unconformably on formations as old 
as the El Paso (in Arizona) and as young as the Fus- 
selman (in most parts of the San Andres and Sacra- 
mento Mountains). However, this region shows no def- 
inite evidence of progressive overlap unless it be in 
the main outcrop belt of the Martin Group where no 
demonstrably Middle Devonian marine faunas have 
been found. However, detailed stratigraphic and pa- 
leogeographic studies by Teichert (1965) demonstrate 
Frasnian onlap of the Defiance uplift from the south 
and southwest. It is possible that some part of the 
dolomite units at the base of the Jerome Member may 
be equivalents of the Onate Formation, but no fossils 
have been reported that would verify such a relation- 
ship. 

The succeeding strata of the Sly Gap Formation rep- 
resent renewed marine deposition in an open shelf en- 
vironment, after a depositional hiatus. Only fine-grained 


clastic material reached the shelf during the middle 
Frasnian, as the Sly Gap is essentially a calcareous 
clay shale unit. Limestone nodules are characteristic 
of the Sly Gap, which has a diverse marine inverte- 
brate fauna including both solitary and compound ru- 
gose corals. The hiatus is inferred because of the ab- 
sence of early Frasnian fossils, approximately of the 
Shell Rock—Waterways interval of Iowa and western 
Canada. 

In addition, black shale including phosphatic lag de- 
posits at the base of the Sly Gap at Alamo Peak and 
Bear Canyon (PI. A, fig. 2) and clastic strata at the 
base of the Sly Gap at Hermosa and Rincon provide 
sedimentologic evidence of this hiatus. Consequently, 
the contact between the Onate and the Sly Gap must 
be a regionally disconformable one. 

The Contadero Formation indicates an influx of silt 
and fine sand in the later Frasnian and a decrease in 
limestone, although calcareous beds do occur. The 
marine invertebrate fauna is less diverse than that of 
the Sly Gap, but it also includes rugose corals. 

Deposition from the Sly Gap into the Contadero must 
have been essentially continuous, although there are 
many discontinuities between individual beds. Sedi- 
ment accumulation was relatively slow, apparently, 
for much of late Frasnian time is recorded in a few 
tens of meters of strata. By way of contrast, many 
hundreds of meters of beds were deposited in western 
Utah and eastern Nevada during this same time inter- 
val. 

Another major depositional hiatus seems to have 
occurred in latest Frasnian and earliest Famennian 
time. Rather than being an episode of non-deposition, 
this must have been a period of slight uplift and ero- 
sion. The sections in the San Andres Mountains record 
removal of varying amounts of Frasnian strata before 
deposition of the Famennian. 

Marine onlap in the early Famennian (about rhom- 
boidea Zone time) is recorded in many places by sandy 
phosphatic lag deposits containing conodonts repre- 
senting this zone. This was the time of initiation of 
Percha deposition in southwestern New Mexico. It was 
also the time of deposition of the sandy Thoroughgood 
Formation in the San Andres Mountains and black 
shale in the southern part of the Sacramentos. 

The Percha beds record deposition in a deep basin 
resulting in the black, essentially unfossiliferous, shales 
of the Ready Pay Member. Shallow-water, more nor- 
mal, marine shelf conditions are recorded by the cal- 
careous shale and nodular limestone of the overlying 
Box Member with its diverse fauna of brachiopods, 
solitary rugose corals, bryozoans and molluscs. Gra- 
dational stratigraphic relationships between the two 


30 BULLETIN 315 


members indicate continuous deposition through mid- 
dle and early late Famennian time, with a progression 
from deeper water to shallow-shelf conditions. 

In the latest Famennian there was again a period of 
slight uplift during which the Devonian beds were re- 
moved to various depths throughout the region. Sub- 
sequently, middle and late Kinderhookian strata were 
deposited unconformably on Devonian beds as young 
as the Box Member, in the Hillsboro area, and as old 
as the Onate Formation, in the southern part of the 
Sacramento Mountains. 


Table 9.—Brachiopods of the Independence Shale described in 
this paper. 


Schizophoria amanaensis Stainbrook 
Stainbrookia infera (Calvin) [new genus] 
Cariniferella costellata new species (=C. iowensis part) 
Strophonelloides deeringi Stainbrook 
Nervostrophia extensa new species 
Nervostrophia multinervosa Stainbrook 
Pseudodouvillina euglyphea Stainbrook 
Sulcatostrophia calvini (Miller) 

Douvillinaria variabilis (Calvin) 

Floweria transversalis new genus and species 
Productella rugatula Stainbrook 
Devonoproductus vulgaris Stainbrook 
Gypidula cornuta Fenton and Fenton 
Gypidula stainbrooki new species 

Costatrypa varicostata (Stainbrook) 
Iowatrypa americana (Stainbrook) 
Spinatrypa trulla (Stainbrook) 

Acutatheca propria Stainbrook 

Thomasaria altumbona Stainbrook 

Cyrtina inulta Stainbrook 


Table 10.—Brachiopods of the Cerro Gordo Member and Cedar 
Valley Formation, described in this paper. 


Schizophoria lata Stainbrook [Cedar Valley Formation] 

Schizophoria iowensis (Hall) 

Strophonelloides hybridus (Hall & Whitfield) [see S. dorsoconvex- 
us| 

Douvillina arcuata (Hall) 

Floweria prava (Hall) [new genus] 

Eostrophalosia rockfordensis (Hall & Clarke) 

Gypidula cornuta Fenton and Fenton 

Calvinaria bransoni Stainbrook 

Pseudoatrypa devoniana (Webster) 

Tenticospirifer cyrtinaformis (Hall & Whitfield) 

Cranaena calvini (Hall & Whitfield) 

C. navicella (Hall) 


BRACHIOPODS FROM IOWA 


In the course of describing the Frasnian and late 
Middle Devonian brachiopods from New Mexico, it 
was necessary to discuss or describe a number of 
species from correlative strata in Iowa. In his paper 
on the Independence Shale brachiopods, Stainbrook 
(1945) described 35 genera and 40 species. We have 
redescribed about half of these taxa and have added 
two new genera and four new species. 

Eleven species from the Hackberry Formation (Cer- 
ro Gordo Member) and one from the Cedar Valley 
Formation are also redescribed herein. 


SYSTEMATIC PALEONTOLOGY 


INTRODUCTION 


Philosophical Considerations 


The species named and delineated here are mor- 
phospecies in the sense of Hoover (1981). No other 
approach to presenting the wealth of morphologic de- 
tail appears practical. As discussed above, most of the 
New Mexico Devonian brachiopod genera have al- 
ready been distinguished. This is especially true of the 
Frasnian Sly Gap assemblage, for which only one new 
genus is established here. Consequently, in order to 
support detailed biochronologic, biostratigraphic, and 
biogeographic conclusions, every attempt was made 
to separate morphospecies on consistent differences, 
no matter how minute. This may seem to be an ulti- 
mate ‘“‘splitting’’ technique, but we present our re- 
sults, unabashedly, with the full understanding that 
most fossil assemblages fall far short of reflecting the 
biologic diversity of the geologic interval represented 
by the paleontologic sampling. 

The species names are merely conveniences for dis- 
cussing clusters of morphologic variability within what 
are, for the most part, well-established generic con- 
cepts. When similar studies of comparably rich assem- 
blages in various parts of the world are completed, the 
paleontologist will be ready to start formulating sound 
biologic and geologic syntheses. This incompleteness 
has not prevented us from drawing preliminary con- 
clusions concerning the age, correlation and biogeo- 
graphic significance of the New Mexico Devonian bra- 
chiopods. Nor have we shied away from presenting a 
Devonian geologic history of the region that is now 
south-central New Mexico. All these ideas must be 
considered only geologic and biologic hypotheses, to 
be tested by additional work in New Mexico and else- 
where. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuUTRO 31 


Format 


Suprageneric hierarchies are standard for brachio- 
pod systematics and follow, for the most part, those 
of the brachiopod volume of the Treatise on Inverte- 
brate Paleontology (Williams et al., 1965). Minor 
changes were made where we consider earlier workers 
in error. One clarification deserves special mention. 
Generic and suprageneric categories that have classi- 
cally been attributed to Schuchert and LeVene (1929) 
are herein attributed solely to Schuchert who wrote 
(1929, p. 4), ‘The senior author presents a new clas- 
sification (p. 10) built upon the older ones. . .”’ 

Simple generic and specific diagnoses are given for 
most of the taxa, and detailed descriptions are given 
for the new ones. Variability of morphologic features 
is presented both qualitatively and quantitatively, 
where feasible. Some single specimens or small sam- 
ples, however, are so distinct morphologically that they 
are described as new. 

The Percha brachiopod assemblage is well-known 
and extensively described by earlier workers. We dis- 
cuss all the taxa known to us, but illustrate here only 
those species that we describe as new or redescribe 
on the basis of new material or additional observa- 
tions. For illustrations of the better-known Percha 
genera and species, the reader is directed to Girty 
(1900), Kindle (1909), Stainbrook (1947), Cooper (1942, 
1954, 1955) and Muir-Wood and Cooper (1960). Some 
of the brachiopods discussed by Stainbrook (1947) led 
him to erroneous conclusions about the age of the Per- 
cha. We list and discuss these taxa at the end of the 
systematics section, and give our views and interpre- 
tations of each. 


Terminology 


Most of the descriptive terms used here are defined 
in the glossary of the brachiopod volume of the Trea- 
tise (Williams et al., 1965). Special terminology for the 
orthoids and pentameroids is that of Schuchert and 
Cooper (1932), and productoid terminology follows 
Muir-Wood and Cooper (1960). 


The Collection 


The collection on which this monograph is based 
was brought to the U.S. National Museum of Natural 
History by many individuals. The earliest acquisitions 
were made by the U.S. Geological Survey (G. H. Gir- 
ty, L. C. Graton, S. Paige and N. H. Darton, collec- 
tors). A collection from the Percha Shale at Lake Val- 
ley was presented to the U.S. Geological Survey by 
Frank Springer. 

Cooper was introduced to this prolific Devonian se- 


quence in 1939 when he and Dr. Josiah Bridge of the 
U.S. Geological Survey visited the fine exposures of 
Percha Shale at the abandoned settlement at George- 
town, New Mexico. In 1947, E. L. Yochelson and 
Cooper visited Dr. C. C. Branson, who was surveying 
in the Sacramento and San Andres Mountains, and 
collected from several canyon exposures, especially 
at Sly Gap, then difficult of access. Dr. Branson from 
time to time presented the Museum with small lots of 
Sly Gap fossils. 

In 1947, Frank V. Stevenson presented his exten- 
sive collection of New Mexico Devonian fossils to the 
Museum. 

In 1948, Arthur L. Bowsher and W. T. Allen of the 
U.S. National Museum mapped and made collections 
in the Sacramento Mountains and Silver City area. 
This was followed by further collecting in 1952. In 
1956, Bowsher, Dutro, and Cooper, joined by Dr. 
Rousseau Flower from the New Mexico Bureau of 
Mines and Mineral Resources, studied and collected 
in the San Andres and Sacramento Mountains. In 1964, 
Dutro and Cooper measured sections in the Sacra- 
mento and San Andres Mountains and westward to 
Silver City and made additional collections. 

Dr. L. A. Nelson of El Paso presented the Museum 
with choice specimens from the Sly Gap and Percha 
formations. Mr. C. O. Levorson, avid collector of the 
Iowa Devonian, presented the Museum with rare 
specimens of fossils from the Cerro Gordo Member 
that are illustrated herein. 

Merrill A. Stainbrook, then at Texas Tech. at Lub- 
bock, Texas, presented the Museum with fine speci- 
mens of Independence Shale fossils, some of which 
are illustrated in this monograph. After his death, the 
University of Iowa presented Stainbrook’s collections 
of Sly Gap and Percha specimens to the Museum. 

Collections made at the same locality in different 
years generally have received different locality num- 
bers. 

Collections at localities listed in Appendix 3 for 
which no collectors’ names are given were made by 
the authors. 


Type Repository 


All the type specimens cited in the following section 
are identified as USNM specimens, reposited in the 
U.S. National Museum of Natural History, Smithson- 
ian Institution, Washington, D.C. 20560, U.S.A. Some 
of the types originally illustrated by Stainbrook also 
bear an M.A.S. number; this refers to the M. A. Stain- 
brook collection, now also reposited in the U.S. Na- 
tional Museum of Natural History. 


BULLETIN 315 


Measurements 


Measurements given herein are standard for bra- 
chiopod systematics, unless otherwise qualified, and 
are given in millimeters. The use of an asterisk in the 
tables of measurements indicates a number derived 
from the half-measurement of a bilaterally symmetri- 
cal feature. A plus sign after a number indicates that 
the specimen is broken and that the true measure is 
somewhat greater. 


Class INARTICULATA Huxley, 1869 
Order LINGULIDA Waagen, 1885 
Superfamily LINGULACEA Menke, 1828 
Family LINGULIDAE Menke, 1828 
Genus BARROISELLA Hall and Clarke, 1892 


Barroisella species 
Plate 1, figures 1, 2 


Two specimens of a small, narrowly elongate lin- 
guloid are referred to this genus because of the internal 
markings on the ventral valve. One (USNM 2008 18a) 
is a small dorsal valve. The second (USNM 200818b) 
specimen is a larger ventral valve exhibiting only the 
interior. 

The ventral valve is more than twice as long as wide 
and tapers at both ends. The sides are gently rounded. 
Internally the pedicle groove is long and deep. Two 
pallial trunks diverge from the anterior part of the ped- 
icle groove. On the outside of the divergent grooves 
is an elongate muscle scar. Other indefinite muscle 
scars appear at above midvalve. 

The dorsal valve is from a smaller individual but it 
has essentially the same proportions as the ventral 
valve. Some of the exterior is preserved and shows 
fine concentric lines. The posterior part is exfoliated 
but the specimen is too young to show any interior 
details. 

Measurements (in mm).—USNM 200818b (ventral 
valve): length 6.7, width 3.4. USNM 200818a (dorsal 
valve): length 5.0, width 2.5. 

Diagnosis.—Small Barroisella having the length 
more than twice the width. 

Stratigraphic occurrence. 

Locality.—5240. 

Types.—Figured Specimens: USNM 2008 18a, b. 

Discussion.—Specimens of linguloids having essen- 
tially the same size and proportions of this Contadero 
species occur in late Middle and early Late Devonian 
sediments of the eastern United States but none of 
them has been determined to be Barroisella. 


Contadero Formation. 


Genus LINGULA Bruguiere, 1797 
Lingula species | 

Two crushed specimens are all of this genus that 
have so far been found in the Percha Formation (Box 
Member). One specimen (USNM 200834) is 16 mm 
long and about 10 mm wide and is ornamented by very 
closely spaced raised concentric lines. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—S506b. 

Types.—Described Specimen: USNM 200834. 


Lingula species 2 


Another linguloid specimen (USNM 200839) is sim- 
ilarly crushed, and much of the shell is torn away. It 
is 13 mm long and 9 mm wide. Bits of shell retained 
are ornamented with concentric lines, coarser and less 
crowded than those of Lingula sp. 1. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—S07c. 

Types.—Described Specimen: USNM 200839. 


Lingula species 3 


The single dorsal valve (USNM 207697) is small 
(length 12 mm and width about 7 mm), with well- 
rounded sides and rounded anterior margin; posterior 
margin narrowly rounded. Its anterior half is marked 
by four strong, distant concentric lines. The posterior 
half is without strong concentric marks. The species 
has some resemblance to Lingula complanata H. S. 
Williams (1882) of the Ithaca Formation of New York, 
but has stronger concentric markings and is more nar- 
row posteriorly. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—529n, o. 

Types.—Described Specimen: USNM 207697. 


Lingula species 4 


Two specimens measure about 10 mm in length and 
5 mm in width, tapering at both anterior and posterior; 
sides gently rounded. These specimens are unlike Lin- 
gula sp. 3 in their more acuminate ends and gently 
rounded sides. This species is shaped somewhat like 
L. ligea Hall, 1860b from the Middle and lower Upper 
Devonian rocks of New York. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—529r. 

Types.—Described Specimens: USNM 207698a, b. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 33 


Genus TRIGONOGLOSSA Dunbar and 
Condra, 1932 


Trigonoglossa? species 


This large, triangular specimen is badly crushed and 
marked by strong, concentric lines, which are elevated 
near the anterior. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—526h. 

Types.—Described Specimen: USNM 207695. 


Genus LINGULIPORA Girty, 1898 


Lingulipora porifera new species 
Plate 39, figures 52-56 


Small, longitudinally elliptical in outline; length ap- 
proximately twice width; posterior margin forming an- 
gle of about 70 degrees; sides slightly convex, nearly 
parallel; anterior margin broadly rounded; anterolat- 
eral extremities narrowly rounded. Surface marked by 
unequal, concentric lines closely crowded on sides, 
more widely spaced anteriorly on body of shell. Inner 
layer of shell punctate, with coarse punctae scattered 
on umbonal region anterior to beak, but fine and 
crowded punctae on remainder of shell (USNM 
200899k). 

Ventral valve most convex medially with sides slop- 
ing moderately to margins in anterior half but more 
steeply in posterior where valve is deepest; lateral pro- 
file broadly and gently convex, maximum height oc- 
curring about one-third distance from beak; umbonal 
region subcarinate and extending posteriorly to sharp 
and narrow beak. Propareas prominent, pedicle groove 
deep and narrow; interior details indistinct. 

Dorsal valve in lateral profile with maximum height 
in umbonal region, descending abruptly posteriorly but 
gently to anterior margin; anterior profile forms broad 
low triangle roundly subcarinate at apex. Beak region 
roundly angular. Posterior margin thickened on inside 
of valve; interior with low, faint median ridge slightly 
posterior to midvalve. 

Measurements (in mm).— 


ventral valves dorsal valves 


USNM USNM USNM USNM USNM_ USNM 
200899a 200899d 200899) 200899e 200899f 200899h 


Length 5.0 7.8 5.0 S27 4.8 4.4 
Width 2.9 3355) 2.6 3.0 ee) Ds) 


Diagnosis. —Small Lingulipora having nearly par- 
allel sides, length about twice width and subcarinate 
umbonal regions. 

Stratigraphic occurrence.—Percha Formation (bas- 
al Ready Pay Member). 


Locality.—3037. 

Types.—Holotype: USNM 200899b. Figured Para- 
types: USNM 200899d, f, h, and j. Unfigured Para- 
types: USNM 200899a, c, g, i. 

Discussion and comparison.—This species differs 
from Lingulipora williamsana Girty (1898) from the 
Chattanooga Shale in its larger size, longer valves and 
more nearly parallel sides. Girty’s is the only species 
of this genus so far described from this country. In his 
discussion of the genus Lingulipora, Girty (1939, p. 
60) found it confusing that punctae seem to appear 
sporadically in some linguloid species and in the genus 
Trigonoglossa Dunbar and Condra (1932), a fact that 
led him to have doubts about the validity of Linguli- 
pora. It 1s true that little is known of the role and 
occurrence of punctae in the inarticulate brachiopods, 
especially the linguloids. Bistramia Hoek (1912) from 
the Devonian of Bolivia is strongly punctate. There- 
fore, we recognize Lingulipora, because the punctae 
are a convenient means of distinguishing this Ready 
Pay species. 


Order ACROTRETIDA Kuhn, 1949 
Suborder ACROTRETIDINA Kuhn, 1949 
Superfamily DISCINACEA Gray, 1840 
Family DISCINIDAE Gray, 1840 
Subfamily ORBICULOIDEINAE Schuchert, 1929 
Genus ORBICULOIDEA d Orbigny, 1847 


Orbiculoidea species | 
Plate 1, figures 3-6 


A single large dorsal valve (USNM 200817a) refer- 
able to Orbiculoidea, taken from the Contadero For- 
mation, is broadly elliptical, wider than long, and ir- 
regularly conical. In lateral profile the cone is 
asymmetrical, the anterior slope long and convex, the 
posterior slope short and precipitous, probably some- 
what deformed. The apex of the cone is located about 
three-fourths the valve length from the anterior mar- 
gin. In anterior profile the specimen forms a low arch. 
The apex is smooth but the remainder of the shell is 
marked by fine concentric lines and faint radial cos- 
tellae. 

A second specimen (USNM 200817b), smaller than 
the former and probably a young shell of the same 
species, is subcircular, a low cone. The ventral valve 
is concave and has a large foramen. 

Measurements (in mm).—USNM 200817a: length 
13.7, width 17.4 (based on the half measure); height 
of cone 5.6. USNM 200817b: length 6.5, width 6.0; 
height 2.0. 


34 BULLETIN 315 


Stratigraphic occurrence.—Contadero Formation 
(lower 5 ft). 

Locality.—524o0. 

Types.—Figured Specimens: USNM 200817a, b. 


Orbiculoidea species 2 


A single dorsal valve from the Percha Formation 
(Box Member) at locality 506b is so poorly preserved 
that it cannot be identified specifically. It is 15 mm in 
diameter, a low cone with most of the posterior third 
eroded away. The surface is marked by broad con- 
centric undulations. 

Types.—Described Specimen: USNM 200863. 


Suborder CRANIIDINA Waagen, 1885 
Superfamily CRANIACEA Menke, 1828 
Family CRANIIDAE Menke, 1828 
Genus PHILHEDRA Koken, 1889 


Philhedra species 
Plate 1, figure 22 


A single specimen was taken with Rhipidomella at 
Sly Gap. It is small, 8.6 mm long by about 9 mm wide. 
The sides are well rounded, the anterior margin nar- 
rowly rounded. The posterior margin is truncated. The 
smooth apex is located about one-third the length from 
the posterior margin. Costellae measure four to the 
millimeter at the front margin. 

Stratigraphic occurrence.—Sly Gap Formation in 
beds below Macgeea Webster (1889b), at the base of 
the formation. 

Locality.—506}. 

Types.—Figured Specimen: USNM 203534. 


Genus PETROCRANIA Raymond, 1911 


Petrocrania ourayensis (Kindle) 
Plate 1, figures 23-26 


Crania ourayensis Kindle, 1909, p. 391, p. 15, pl. I, figs. 2, 2a. 
Petrocrania ourayensis (Kindle) Stainbrook, 1947, p. 302, pl. 45, 
fig. 10. 


This uncommon species, like all members of the ge- 
nus, is variable with its shape and profile depending 
on the contours of its host. Specimens often reflect, 
on the dorsal valve, the ornamentation of the host. 
The species selects a variety of hosts of which Leio- 
productus coloradoensis (Kindle) and Schizophoria 
australis Kindle are favorites. 

Diagnosis.—Fairly large Petrocrania, subcircular 
cones with apex slightly posterior of the center. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 


Localities.—506, q; 507m, n, 0; 509b, c, e; 527a, d; 
3026, a. 

Types.—Figured Hypotypes: USNM 200826a, 
200827. 


Petrocrania species | 
Plate 1, figure 7 


This is a small conical dorsal valve perched on the 
posterolateral margin of Hadrorhynchia and thus 
somewhat misshapen. The anterior slope is long and 
convex; the posterior slope is short, steep and nearly 
flat. The apex is about three-quarters the length from 
the anterior margin. The surface is marked by fairly 
strong concentric lines. 

Measurements (in mm).—Length 9.2, width 11.5, 
height 3.0. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—523d. 

Types.—Figured Specimen: USNM 201153. 


Petrocrania specifically unidentified 
Plate 1, figures 8, 9; plate 38, figure 24 


Eight additional specimens are referable to this ge- 
nus and no two of them are alike. The variation is 
undoubtedly due, at least in part, to the surface irreg- 
ularity of their hosts. All of the specimens are de- 
tached from their hosts and are conical dorsal valves, 
consequently the true nature of the host is unknown. 
Faint markings on some suggest that the host was 
probably a costellate brachiopod. Five of the speci- 
mens are flatly conical with the apex slightly eccentric, 
and the valves nearly circular. One specimen is con- 
cave across its middle; another is narrowly elliptical 
in outline and forms a high cone. The eighth specimen 
is a small round cone. The largest specimen (USNM 
200819) has a diameter of 20 mm. It and the other three 
flattish forms may be referable to Petrocrania fame- 
lica (Hall and Whitfield). The others may be aberra- 
tions of this species but the specimens are too poorly 
preserved to be sure. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —S06c, 1; 521c, m, q; 522). 

Types.—Figured Specimens: USNM 200819, 200954. 


Class ARTICULATA Huxley, 1869 
Order ORTHIDA Schuchert and Cooper, 1932 
Suborder ORTHIDINA Schuchert and Cooper, 1932 
Superfamily ORTHACEA Woodward, 1852 
Family SKENIDIIDAE Kozlowski, 1929 
Genus SKENIDIUM Hall, 1860a 


DEVONIAN BRACHIOPODS OF NEW MEXxICO: COOPER AND DuUTRO 35 


Skenidium independense Stainbrook 
Skenidium independense Stainbrook 1945, p. 14, pl. 1, figs. 14-18. 


A single dorsal valve of this rare and tiny species 
was taken at locality 506c. This occurrence helps to 
reinforce the relationship between the Independence 
Shale of Iowa and the Sly Gap Formation of New 
Mexico. 

Locality.—S06c. 

Types.—Mentioned Specimen: USNM 252628. 


Order ENTELETIDA Waagen, 1884 
Superfamily ENTELETACEA Waagen, 1884 
Family ENTELETIDAE Waagen, 1884 
Subfamily SCHIZOPHORIINAE Schuchert, 1929 
Genus SCHIZOPHORIA W. King, 1850 


Schizophoria amanaensis Stainbrook 
Plate 4, figures 28-33; plate 39, figures 30-34 


Schizophoria amanaensis Stainbrook, 1945, p. 17, pl. 1, figs. 1-13; 
1948, p. 770, pl. 1, figs. 5, 6. 


Specimens that are nearly as wide as long are re- 
ferred to this species although they do not conform 
precisely to Stainbrook’s description. Sixteen mea- 
sured specimens have a mean L/W ratio of 0.88. Five 
of the specimens have a L/W ratio of over 0.90, indi- 
cating specimens nearly as wide as long. These gross 
measurements are in accordance with Stainbrook’s 
original description and illustrations. All the complete 
specimens he illustrated (1945, pl. 1, figs. 1-4, 9, 10, 
12, 13), and the smaller ventral interior (1945, pl. 1, 
fig. 11) have the length and width nearly equal. They 
are rotund specimens resembling Schizophoria mac- 
farlani but they definitely are not comparable to Meek’s 
type specimen (USNM 5901a). 

Stainbrook mentions a sulcus on the dorsal valve as 
one of the characters of §. amanaensis; this feature is 
most unusual in Schizophoria. Three complete speci- 
mens of this species from the Independence Shale in 
the USNM collections show no trace of a sulcus, nor 
do any of his figures, except figure 9, which appears 
to be an aberrant or obese specimen having its length 
slightly greater than its width. None of the Sly Gap 
specimens referred here have any trace of a dorsal 
sulcus. It is likely that this is a character of old age or 
one of sporadic occurrence. The folding of S. ama- 
naensis is like that of S. iowensis (Hall, 1858), a sulcus 
on the ventral valve deep in the front half and a dorsal 
valve with the fold scarcely elevated except at the very 
anterior and shown mostly as a deep re-entrant in the 
anterior commissure of the dorsal valve. 


Measurements (in mm).— 


dorsal — mid- 
length valve width hinge thick- L/W 


(L) length  (W) width ness ratio 
USNM 200880 
(loc. 521d) 232 23.2 28.3 18.6? 17.6 0.82 
USNM 201196a 
(loc. 523m) 25.3 25.3 29.2 17.5 20.0 0.87 
USNM 201195a 
(loc. 521le) 20.0 20.1 23.3 —_ 13.7 0.86 
USNM 201197a 
(loc. 526a) 21.4 21.4 23-2 13.0 18.5 0.92 
USNM 201198a 
(loc. 509r) 22.0 22.0 Qn 12.7 17.0 0.88 
USNM 201199a 
(loc. 509n) 23.0 23.0 DSS) 15.0 16.6 0.90 
USNM 201201 
(loc. 5091) 24.0 24.0 28.0 13.0 — 0.86 
USNM 201202a 
(loc. 509i) 23.8 23.8 27.3 18.2 17.8 0.87 
USNM 201200a 
(loc. 362a) 19.0 19.0 22.0 12.0 14.9 0.86 


Mean L/W = 0.87. 


Stratigraphic occurrence.—Independence Forma- 
tion; Sly Gap Formation. 

Localities.—362a, b; 506c, i; 509p, x, z; 521d; 523 
m; 524n, s; 526a, k; 528w, y:; 529y; 3045c, i. 

Types.—Figured Hypotypes: USNM 200880, 
201197a. 

Discussion.—Stainbrook compared S. amanaensis 
only to other Iowa species of Schizophoria, overlook- 
ing the close similarity of his species to §. macfarlani 
(Meek, 1868). The type specimen of this species is in 
the U.S. National Museum of Natural History (USNM 
5901a). Its measurements in mm are: length 21.7, dor- 


thickness 18.7, length/midwidth ratio = 0.98, mid- 
width/length ratio = 1.02. The specimen is thus a very 
thick and rotund one, externally like §. amanaensis. 
The Canadian species, however, seems to be more 
finely costellate than the lowa and New Mexico forms, 
has a narrower dorsal umbonal region and more con- 
vex flanks bounding the ventral valve sulcus. Com- 
parison of the few specimens from New Mexico called 
S. amanaensis with the holotype of 8. macfarlani is 
not a satisfactory one. The National Collection has 
only a few specimens identified as §. macfarlani. Some 
of these do not conform to the type and are from sev- 
eral different formations. So few specimens of S. 
amanaensis are now known, and so few are likely to 
be found in the future, because of the sporadic (cave 
deposit) nature of the Independence Shale outcrop, 
that it is unlikely that a satisfactory comparison be- 
tween the two species can be established. 


BULLETIN 315 


It is also possible that S. amanaensis has an unex- 
pected relationship to §. iowensis Hall (1858). Stain- 
brook (1945, p. 18) mentions the occurrence of another 
species of Schizophoria in the Independence Shale that 
is thinner, broader and less quadrate than S$. ama- 
naensis. Indeed his figure 6 is of the ventral valve of 
a specimen answering this description. The National 
Collection includes a few specimens of these wide 
forms that approach §. iowensis (Hall) very closely. 
Unfortunately, enough specimens are not at hand, and 
those that are have been crushed, to show that these 
belong to S$. iowensis. 

Specimens like S. amanaensis of the Independence 
Shale occur in the “‘Hay River Formation on Hay Riv- 


* 


er about mile 26 in Northwest Territories, Canada.’”’ 


Schizophoria australis Kindle 


Schizophoria striatula Girty, 1900, p. 39. 

Schizophoria striatula var. australis Kindle, 1909, p. 21, pl. 1, figs. 
4-8; pl. 2, figs. 1—Sa; pl. 3, fig. 1. 

Schizophoria australis Kindle, Stainbrook, 1947, p. 303, pl. 45, figs. 
lp hy Up te 


This is one of the largest species of Schizophoria 
found in the United States and one of the largest known 
anywhere. It is thus very easily recognized in any Per- 
cha Formation (Box Member) locality. The specimens 
figured by Stainbrook do not indicate the full size of 
the species. Two of the largest specimens measure (in 
mm): USNM 200890 (loc. 506q): length 44.1, dorsal 
valve length 43.2, hinge width 28.6, midwidth 58.2, 
thickness 28.7; USNM 200910 (loc. 507m): length 44.4, 
dorsal valve length 45.7, hinge width 26.3, maximum 
width 51.8, thickness 32.3. 

Localities.—506, b, p, q; 507a, c, e, g, h, m, n, 0, 
t, w, z; 509a, b, c; 527a, b, c, d, e, f, 1, p; 3026; 3026a; 
3029; 3031a; 3034 (=S07h); 3035. USGS (Kirk) 461; 
508; 512. 

Types.—Lectotype: USNM 62005e. Paratypes fig- 
ured by Kindle (1909): USNM 62005a—d. Hypotypes 
figured by Stainbrook (1947): USNM 135229 (=M.A.S. 
1600). Measured Hypotypes: USNM 200890, 200910. 

Discussion.—A type was never selected for this 
species from Kindle’s syntypes. The specimen figured 
by Kindle (1909, pl. 1, fig. 8; pl. 2, figs. 5, 5a—USNM 
62005e) is here chosen as lectotype. 


Schizophoria lata Stainbrook 
Plate 3, figures 6-16; plate 38, figures 25-29 


Schizophoria lata Stainbrook, 1940, p. 488, pl. 2, figs. 1-7, 15, 16. 


Locality information from label that accompanied specimens. 


Medium to large for the genus, wider than long but 
somewhat variable; sides rounded; subrectangular to 
broadly elliptical in outline; cardinal extremities ob- 
tuse; anterolateral extremities rounded; valves un- 
equally deep, ventral valve shallower; anterior com- 
missure narrowly uniplicate. Beak narrowly pointed; 
interarea slightly curved, strongly apsacline. Hinge 
equal in width to about two-thirds maximum width and 
approximately equal to thickness. Surface costellate, 
two to four costellae in one mm depending on size of 
shell. 

Pedicle valve flatly convex in lateral profile, varying 
from nearly flat to slightly concave or slightly convex 
in anterior profile; umbonal region gently inflated; um- 
bolateral slopes moderately steep; sulcus originating 
slightly posterior of midvalve, deepening rapidly an- 
teriorly to occupy about half valve width. Flanks 
bounding sulcus, flat or nearly so. Tongue long, sub- 
acute to acute. 

Dorsal valve strongly convex, nearly hemispherical 
in lateral profile; anterior profile a strongly rounded 
dome with steeply sloping sides. Umbonal and median 
regions strongly inflated. Fold visible only as deep, 
narrowly rounded to angular re-entrant in anterior 
commissure. Flanks swollen. 

Ventral valve with muscle field extending about one- 
third valve length, and about same part of valve width, 
strongly bilobed; diductors separated by thick median 
ridge. Vascula media strong. 

Dorsal valve interior moderately thickened; adduc- 
tor field large; brachiophore supports strong and long. 

Measurements (in mm).— 


dorsal 
length valve width hinge thick- L/W 
(L) length (W) width ness ratio 


USNM 201204a 

(loc. 506d) 25.8 25.8 34.0 18.2 14.2 0.79 
USNM 201204b 

(loc. 506d) 25.6 25.0 31.9 19.0 17.9 0.80 
USNM 201203 

(loc. S09f) 25.8 26.4 S129) 18.3 17.0 0.81 
USNM 201206a 


(loc. 522c) 26.2 26.2 32.8 19.0 17.3 0.80 
USNM 200924 

(loc. 522h) 26.1 26.1 32.0 19.5 18.7 0.78 
USNM 201211 

(loc. 523a) 30.2 30.2 34.1 20.0 19.2? 0.89 


USNM 200853 

(loc. 523d) 24.6 24.4 30.7 16.6 16.0 0.80 
USNM 201209a 

(loc. 523d) 25.2 2552, 30.5 15.4 17.0 0.83 
USNM 201209b 

(loc. 523d) 25.2 25.2 28.6 17.8 18.0 0.88 
USNM 201209c 

(loc. 523d) 26.4 26.4 313)57/ 17.8 15.0? 0.78 


DEVONIAN BRACHIOPODS OF NEW MEXIco: COOPER AND DuTRO 37 


dorsal 
length valve width hinge  thick- L/W 
(L) length (W) ~~ width ness ratio 


USNM 201209d 


(loc. 523d) 25.6 25.0 28.7 15.0 15.7 0.76 
USNM 201209e 

(loc. 523d) 15.7 15.7 20.6 11.0 9.2 0.76 
USNM 200956 


(loc. 523e) 3253 823 41.7 22.6 26.0 0.76 
USNM 201210 

(loc. 523e) PHS) 27.9 38.0 18.8 18.7 0.73 
USNM 201205a 


(loc. 523i) 26.0 26.0 30.0 12.7? 16.9 0.86 
USNM 201208a 

(loc. 526c) 27.3 27.3 32.0 15.0 19.6 0.85 
USNM 201208b 

(loc. 526c) DSi) 25.4 315 13.8 16.9 0.82 
USNM 201207 

(loc. 526)) DIED 25.5 32.3 — 17.8 0.79 


Diagnosis.—Medium to large Schizophoria having 
a strongly sulcate ventral valve, little or no fold, and 
mean L/W = 0.81. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—S06d; 509f; 522c, f, g, h, j; 523a, b, d, 
Copieuepinukey Sos GU XenVe1 Zs 247 Guxs) S26Cyj9 3023s 

Types.—Figured Hypotypes: USNM 200853, 200924, 
200956. 

Discussion.—The large size, fairly thin shell, and 
transverse outline distinguish this species readily from 
Schizophoria amanaensis Stainbrook (1945), in which 
the length and width are nearly equal. It is distin- 
guished from §. magna Fenton and Fenton (1924), 
which has a similar L/W ratio, by its generally larger 
size, less emarginate anterior margin, and somewhat 
broader and less angulated sulcus. Schizophoria lata 
is unlike other species of the genus from the Cedar 
Valley Formation, as it is larger than S. iowensis (Hall, 
1858) and S. meeki Fenton and Fenton (1928) and more 
transverse than the latter and §. /audoni Stainbrook 
(1940), another large species. It is also unlike any of 
the species described from Michigan by Imbrie (1959) 
all of which are from older strata, except S. mesacar- 
ina Imbrie (1959). 


Schizophoria iowensis (Hall) 
Plate 3, figures 1-5 


Orthis iowensis Hall, 1858, p. 488, pl. 2, fig. 4. 

Orthis (Schizophoria) impressa Calvin (not Hall), 1897, p. 167. 

Orthis (Schizophoria) iowensis (Hall) Hall and Clarke, 1892, pl. 6A, 
fig. 29. 


This species is abundant in the Cerro Gordo Mem- 
ber of the Hackberry Formation at Rockford, Iowa. 
The size is variable and it has a mean L/W ratio of 
0.82 (W/L ratio = 1.21) with the range of the former 


from 0.74 to 0.89, indicating a moderately to roundly 
transverse outline. Although Schizophoria is not com- 
mon in the Sly Gap Formation, a number of localities 
yielded specimens, here referred to that genus, that 
are well within the range of variation of Schizophoria 
iowensis. In addition to the agreement in dimensions, 
these specimens also agree in the character of the fold 
and sulcus, many having the deeply indented anterior 
seen in lowa specimens. 

Diagnosis. —Medium sized, transverse Schizopho- 
ria, with L/W ratio ranging from 0.74 to 0.89, and 
marked by a deep sulcus in the anterior half of the 
ventral valve, but without a fold, the tongue of the 
ventral valve fitting into a deep re-entrant in commis- 
sure of the dorsal valve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —S06c, j; 509g, n, s, y; 521c, d, k, m, q, 
s, w; 524m, n. 

Types.—Figured Hypotype: USNM 200955. 


Schizophoria species | 
Plate 4, figures 1-6 


Medium sized for the genus, wider than long, round- 
ly subrectangular, cardinal extremities and sides 
strongly rounded. Maximum width posterior to mid- 
valve. Anterior commissure broadly uniplicate. Inter- 
area long and steeply apsacline, only slightly curved. 
Twelve costellae in five mm. 

Ventral valve resupinate in lateral profile and broad- 
ly concave in anterior profile. Beak small and narrow, 
umbonal region nearly flat; umbolateral slopes convex 
and steeply inclined to the cardinal extremities. Sulcus 
originating slightly posterior of midvalve, broad and 
moderately deep, involving more than half anterior; 
lateral slopes flattened and sloping toward sulcus rath- 
er than toward margin as is usual in Schizophoria. 

Dorsal valve strongly convex in lateral profile and 
strongly domed in anterior profile, sides dipping 
strongly to margins. No fold visible except as a deep 
re-entrant in anterior commissure. Lateral slopes flat- 
tened; umbonal and median regions strongly swollen. 

Measurements (in mm).—Length 29.5, dorsal valve 
length 29.5, maximum width 34.2, hinge width 20.0, 
thickness 19.0, L/W ratio = 0.86. 

Diagnosis. —Medium sized Schizophoria having a 
broadly concave ventral valve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—506d. 

Types.—Figured Specimen: USNM 200908. 

Discussion.—Only a single specimen of this species 
was obtained. It is exceptionally well preserved but it 
occurs with abundant S. /ata Stainbrook (1940) and 
may be an aberration of that species. Without more 


38 BULLETIN 315 


knowledge of the variation of S. /ata, the specimen is 
not named although its characters are unusual and dis- 
tinctive. 


Schizophoria species 2 
Plate 4, figures 22-27 
One unusually well-preserved specimen has the form 
and dimensions of S. amanaensis Stainbrook (1945) 
but with much stronger costellae, especially in the sul- 
cus. The fold is somewhat better defined than that of 
S. amanaensis and the sulcus is shallower. The hinge 
is also narrower than that of S. amanaensis. 
Measurements (in mm).—USNM 200852: length 
22.3, dorsal valve length 22.5, midwidth 25.5, hinge 
width 15.0, thickness 16.4, L/W ratio = 0.86. 
Stratigraphic occurrence.—Sly Gap Formation. 
Locality.—509p. 
Types.—Figured Specimen: USNM 200852. 


Schizophoria species indeterminate 


Several fragmentary specimens of a small Schizo- 
phoria were found in the Contadero Formation at lo- 
calities 523n; 524c; and 526f, g. They are not specifi- 
cally identifiable. 


Superfamily RHIPIDOMELLACEA Alichova, 1960 
Family ONNIELLIDAE Opik, 1933 
Genus AULACELLA Schuchert and Cooper, 1931 


Aulacella elliptica new species 
Plate 2, figures 11-20, 31-59, 73-77; 
plate 39, figure 22 


About medium size, roundly elliptical in outline; 
sides strongly rounded; anterior margin broadly 
rounded; maximum width at midvalve in majority of 
specimens; posterior with beak of ventral valve slight- 
ly protuberant. Hinge narrow, about half valve width. 
Valves of nearly equal convexity. Anterior commis- 
sure slightly curved in ventral direction. Surface mul- 
ticostellate with fine, narrowly-rounded costellae, im- 
planted and intercalated in four or five generations; 
twelve costellae in five mm at anterior margin of large 
specimen. 

Ventral valve having gently and evenly convex lat- 
eral profile; anterior profile gently and broadly convex, 
similar to that of lateral profile. Umbonal region nar- 
rowly convex and extending anteriorly as low, narrow 
fold most prominent at midvalve, less so anteriorly; 
flanks gently convex, separated from the fold by shal- 
low depression on each side of fold. Interarea short, 
curved and apsacline. 


Dorsal valve having about same convexity in lateral 
profile as that of ventral valve; anterior profile broadly 
convex, slightly more so than that of ventral valve and 
marked medially by slight narrow depression repre- 
senting the sulcus. Umbonal region slightly swollen; 
posterolateral extremities depressed. Sulcus originat- 
ing at beak, expanding anteriorly, deepening to about 
midvalve, then becoming shallower. Flanks gently 
swollen. 

Interior not seen. 

Measurements (in mm).— 


maxi- 

dorsal mum 
length valve width hinge thick- L/W 
(L) length (W) width ness_ ratio 


USNM 200885 
(Holotype; loc. 52le) 12.3. 11.7 15.1 7.4 5.6 0.82 
USNM 200884 


(loc. 526e) IS 4 SES eGs0) 5.0 0.83 
USNM 200888 

(loc. 509n) 9.6 SIS NESE DeS) 4.2 0.80 
USNM 200887 

(loc. 521d) 8.8 Ciy/ie mL O22 ee 8) 49 0.86 
USNM 201192a 

(loc. 524m) 10:8) 1016) 122555 6:6 5.3 0.87 
USNM 203562 

(loc. 527r) 8.8 8.7 98 4.6 43 0.90 
USNM 201192b 

(loc. 524m) 9.1 Hoy IU © HGS) 5.0 0.79 
USNM 201193 

(loc. 5240) 10.1 9.9 12.4 6.0? 4.6 0.81 
USNM 201194 

(loc. 506j) 9:5 10.6 97 16:0 47 0.93 


Diagnosis.—Roundly elliptical, finely costellate Au- 
lacella having compressed valves and tendency for fold 
and sulcus to merge anteriorly with overall valve con- 
vexity. 

Stratigraphic occurrence.—Sly Gap Formation; 
Contadero Formation. 

Localities. —Sly Gap Formation: 506c, j, y; 509j, n, 
Pp, q,1r, 8, u, x, y; S21c, d,e, k, m, q, s, t; 5230a; 524m, 
x; 526a, e, k, r; 527r; 528y; 3016 (=506)). 

USGS (Kirk) 543. Contadero Formation: 5240. 

Types.—Holotype: USNM 200885; Figured Para- 
types: USNM 200884, 200887, 200888, 200909, 203562. 

Discussion.—Stainbrook (1948, p. 770, pl. 1, figs. 
15, 16) refers specimens from Alamo Canyon to A. 
infera (Calvin, 1878). The specimens figured are ju- 
veniles, enlarged 5, that might be any juvenile punc- 
tate orthid. The specimens from the Sly Gap in the 
National Collection are unlike Calvin’s species (see PI. 
2, figs. 21-30). Aulacella infera is a squarer form with 
L/W ratio of 0.90, is much thicker and the costellae 
are more distant and less numerous. No specimens 
identifiable with Calvin’s species were taken from New 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuUTRO 39 


Mexico in all of the collecting for the U.S. National 
Museum of Natural History. 


Aulacella manticula (Stainbrook) 


Rhipidomella manticula Stainbrook, 1947, p. 303, pl. 45, figs. 3, 5, 
6 (not fig. 4 = Thiemella platys n. sp.). 


The narrowed hinge region, stout thick valves and 
anteriorly located maximum width of this species in- 
dicate assignment to Aulacella Schuchert and Cooper 
(1931) rather than to Rhipidomella Oehlert (1890). The 
specimens illustrated by Stainbrook in figures 3, 5 and 
6 are similar and exhibit the characters mentioned 
above. The specimen illustrated by figure 4 is differ- 
ently shaped, and has a somewhat shouldered appear- 
ance rather than the sloping posterior margins of the 
other specimens. This individual (fig. 4) is identical to 
specimens coming from the Percha Formation (Box 
Member) farther west in New Mexico and here de- 
scribed under the name of Thiemella platys, new 
species. We suggest that Stainbrook’s specimen orig- 
inated at Hillsboro, New Mexico, at the type locality 
of the Box Member. It has bits of pink shale adhering 
to it, as is common with specimens coming from the 
type locality of the Percha, but this is not seen in any 
specimens from Rhodes Canyon. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation. 

Locality.—East end of Rhodes Pass, San Andres 
Mountains, New Mexico. 

Types.—Holotype: M.A.S. 1589A (=USNM 
135260). Figured Paratype: M.A.S. 1589B (=USNM 
135261la) (not M.A.S. 1589C = Thiemella platys n. sp.). 


STAINBROOKIA new genus 


Small, strongly biconvex, subquadrate in outline with 
rounded sides and truncated anterior margin; ventral 
valve with median fold in young specimens reverting 
to sulcus at the anterior in old ones; dorsal valve ini- 
tially with shallow sulcus but changing to an anterior 
fold in adult or large specimens; anterior commissure 
uniplicate or sulcate depending on size and age. Ven- 
tral valve interarea short and curved; delthyrium open. 
Surface multicostellate, costellae increasing by im- 
plantation in at least three generations. 

Ventral valve interior with long diductor scars form- 
ing bilobed patch with muscle scars separated by low 
ridge; pallial trunks diverging anterolaterally from an- 
terior ends of the diductors. Teeth thick and having 
deep fossettes on their anterior surface. 

Dorsal valve interior with thickened cardinalia in 
adult; brachiophores flattish blades placed obliquely 
and supported by strong thickenings of adventitious 


shell; brachiophores bearing knoblike teeth to articu- 
late with dental fossettes; crura (?) short and blunt. 
Adductor field subcircular, muscle scars deeply im- 
pressed; median ridge variable, usually not strongly 
developed. Cardinal process with short shaft and tri- 
lobed myophore. 

Type species.—Orthis infera Calvin (1878, p. 728). 

Diagnosis.—Small, strongly biconvex Onniellidae 
having a bilobed ventral muscle field; short-shafted 
cardinal process; ventral fold and dorsal sulcus re- 
verting to ventral sulcus and dorsal fold in adult stages. 

Comparison.—The species described herein were 
referred previously to Aulacella Schuchert and Coo- 
per (1931) which has an interior similar to that of 
Stainbrookia but an entirely different exterior. Aula- 
cella is biconvex but its valves are flatly convex and 
the fold of the ventral valve, although often not strong- 
ly developed, is narrow and extends to the anterior 
margin. The exterior of Aulacella is more like that of 
Rhipidomella Oehlert (1890), while the interior of 
Stainbrookia is suggestive of a small species of /sor- 
this Kozlowski (1929). 


Stainbrookia infera (Calvin) 
Plate 2, figures 21-30; plate 39, figures 18-21 


Orthis infera Calvin, 1878, p. 728. 
Aulacella infera (Calvin) Stainbrook, 1945, p. 15, pl. 1, figs. 19-28. 


Figures of this strongly biconvex species are intro- 
duced for comparison with Aulacella from the Sly Gap 
Formation in New Mexico. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—526w. 

Types.—Figured Hypotypes: USNM 200886, 
200886b-e. 


Stainbrookia rotunda new species 
Plate 39, figures 1-7 


Small, strongly biconvex shell with ventral valve 
slightly deeper than dorsal one; subquadrate in out- 
line; wider than long; greatest width at midvalve; sides 
narrowly rounded; anterior commissure narrowly sul- 
cate. Beak low, strongly incurved and protruding only 
slightly posterior to hinge, which is narrower than 
midvalve. Surface multicostellate, costellae strong, in- 
creasing in two generations and somewhat fasciculate 
at the anterior. 

Ventral valve strongly convex in lateral profile; an- 
terior profile broadly convex with suggestion of keeled 
middle. Sides convex and steeply sloping. Fold low, 
formed by two prominent median costellae; interarea 
short and strongly curved. 


40 BULLETIN 315 


Dorsal valve strongly convex but less so than ven- 
tral valve; anterior profile strongly domed and indent- 
ed medially by narrow and deep sulcus; flanks narrow- 
ly convex and steep-sided. 

Interior unknown. 

Measurements (in mm).—Holotype: length 5.6, dor- 
sal valve length 5.5, hinge width 5.8, midwidth 7.3, 
thickness 4.9. 

Diagnosis.—Small, strongly convex Stainbrookia 
having a deep and narrow sulcus. 

Stratigraphic occurrence.—Contadero Formation. 

Locality.—524s; 526a. 

Types.—Holotype: USNM 207700a; Figured 
Paratype: 207700b; Unfigured Paratypes: USNM 
207700c-g. 

Comparison.—This rare species is separated readily 
from S. infera (Calvin) by its small size, strongly con- 
vex valves and the deep dorsal sulcus. 


Genus CARINIFERELLA Schuchert and Cooper, 1931 


Cariniferella alleni new species 
Plate 1, figures 27-47; plate 3, figures 17-25 


Medium size for genus, slightly wider than long, 
roundly subquadrate in outline, cardinal and antero- 
lateral extremities fairly broadly rounded. Hinge 
straight, about three-fourths valve width; greatest width 
at midvalve. Anterior margin with narrow re-entrant 
medially; anterior commissure sulcate. Surface fasci- 
costellate; costellae narrowly rounded, increasing by 
bifurcation and implantation, appearing in three gen- 
erations. At anterior margin of large specimen, one 
costella separating two or three finer ones from next 
larger one. Seven or eight costellae in five mm on dor- 
sal valve and seven to ten in same distance on ventral 
valve, fasciculation of two or three costellae taking 
place at anterior margin. 

Ventral valve evenly and gently convex in lateral 
profile; broadly convex in anterior profile and narrow- 
ly elevated medially. Beak small, narrow, incurved; 
umbonal region carinate; interarea short, concave and 
apsacline. Fold low and narrow, formed by a fascicle 
of costellae somewhat smaller than the largest costel- 
lae and thus conspicuous because of this difference. 
Flanks bounding fold, slightly concave adjacent to fold 
but faintly convex to flat beyond fold. 

Dorsal valve deeper and more convex than ventral 
valve, moderately convex in lateral profile; anterior 
profile strongly convex but with median region nar- 
rowly depressed by sulcus. Flanks slightly convex and 
steep in profile. Sulcus originating at beak, narrow and 
deep but widening anteriorly to occupy about one-third 
valve width. Costellae of sulcus crowded and of small- 
er size than those of flanks. 


Ventral valve interior with elongate, narrow teeth; 
dental plates short, obsolescent because of filling of 
lateral cavities by adventitious shell material; muscle 
scar cordate, not reaching midvalve; diductor scars 
narrow, enclosing small adductor scars. Median part 
of muscle area narrowly elevated and rising to crest 
at anterior of scar and overhanging a pit anterior to 
crest which is origin of trough that extends to anterior 
margin. Genital areas reniform, occupying the pos- 
terolateral areas. 

Dorsal valve interior greatly thickened, brachio- 
phores appearing as flat blades supported by adven- 
titious thickening; notothyrial platform thick; cardinal 
process bilobed, small. Adductor scars on a thickened 
platform; median ridge, formed by inwardly depressed 
shell making the sulcus, strong and widening anterior- 
ly. 

Measurements (in mm).— 


dorsal mid- 
length valve width hinge thick- L/W 
(L) length (W) width ness _ ratio 


USNM 200833 

(loc. 509i) 1910) S1SE7. 9 2378 AO 9.7 0.80 
USNM 200831 

(Holotype; loc. 509p) 17.3. 16.5 21.4 14.3 10.0 0.81 


USNM 200832 
(loc. 521s) 1835 18:0) 21k4> 1220 20 Or a7 
USNM 200837c 
(loc. 506c) 146 146 17.8 10.8 OF O82 
USNM 200837d 
(loc. 506c) 1228) 12-2 S700 8.7 0.82 
USNM 200837e 
(loc. 506c) 8.9  — 10.7 7.7 — 0.83 
USNM 200836 
(loc. 521p) — 18.5 23.5 13.22 — — 


Diagnosis.—Roundly subquadrate Cariniferella 
having narrow fold, deep sulcus and fairly strong bi- 
convexity. 

Stratigraphic occurrence.—Sly Gap Formation (up- 
per part). 

Localities. —S06c, i, v; 5O9f, h, 1, n, p, r; 521c, e, k, 
0, p, u; 524m, u, y; 526k; 529s; 3054. 

Types.—Holotype: USNM 200831. Figured Para- 
types: USNM 200833, 200836, 200837a—d. Unfigured 
Paratype: USNM 200837e. 

Comparison.—Stainbrook identified specimens of 
Cariniferella from the Sacramento Mountains with his 
species C. iowensis from the Independence Shale of 
Iowa. The two specimens are similar in some details 
but are sufficiently unlike to be separate species. Al- 
though the measurements are similar and the L/W ra- 
tio nearly alike for Stainbrook’s (1945) figured speci- 
mens, the Independence species has a deeper dorsal 
valve, a narrower and more erect umbo on the dorsal 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 41 


valve and the flanks of the dorsal valve are steeper. 
Inside the ventral valve the muscle field is larger than 
that of C. alleni. On the exterior, the ornament of C. 
iowensis is Somewhat stronger and the costellae more 
distant than on C. alleni. 

Discussion.—Cariniferella alleni is a rare fossil. It 
proves to be more common in the Sacramento Moun- 
tains, especially in Indian Wells Canyon, than in the 
San Andres Mountains or west of that range. 

C. alleni n. sp. is named for William T. Allen who, 
with Arthur L. Bowsher, assisted in our investigations 
of the Devonian of New Mexico. 


Cariniferella costellata new species 
Plate 38, figures 13-19 


Cariniferella iowensis Stainbrook, 1945, pl. 1, fig. 29 only. 


Small for genus, elliptical in outline with well-round- 
ed cardinal and anterolateral extremities. Valves sub- 
equally deep. Greatest width at midvalve; anterior 
commissure narrowly sulcate. Beak small, incurved. 
Hinge straight, measuring about two-thirds maximum 
width. Surface costellate, ten in five mm at anterior of 
holotype, and appearing in three generations. 

Ventral valve nearly flat in lateral profile except for 
umbonal region which is convex; anterior profile sub- 
carinate medially and gently concave on sides; fold 
beginning at beak, widening anteriorly but narrow 
throughout and essentially a median fascicle. 

Dorsal valve gently convex in lateral profile, broadly 
convex in anterior profile but narrowly depressed in 
middle. Sulcus formed at beak, widening and deep- 
ening anteriorly and taking about one-third valve width. 
Flanks bounding sulcus gently convex. 

Interior not known. 

Measurements (in mm).—Length 11.9, dorsal valve 
length 11.7, midwidth 16.0, hinge width 11.0, thickness 
4.7, L/W = 0.73. 

Diagnosis.—Small, transverse Cariniferella having 
fine costellae. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—526u. 

Types.—Holotype: USNM 201214. 

Comparison.—A specimen of this species was fig- 
ured by Stainbrook as C. iowensis Stainbrook but 
compared to the young of that species it proves to 
have finer costellae, is more transverse, and does not 
have the narrow swollen umbonal region so charac- 
teristic of C. iowensis. The species is suggestive of the 
young of C. tioga (Hall, 1867) but that species attains 
a large size. It is also suggestive of C. virginia (H. S. 
Williams, 1908), but that species is less rounded lat- 


erally and has higher shoulders than the Iowa speci- 
men. 


Cariniferella iowensis Stainbrook 
Plate 2, figures 78-82 


Cariniferella iowensis Stainbrook, 1945, p. 16, pl. 1, figs. 29-37. 


A single complete specimen has the narrow, rotund 
form, deep dorsal sulcus, and deep valves of Stain- 
brook’s species and these are the significant characters 
distinguishing it from C. alleni n. sp. of the Sly Gap. 

Stratigraphic occurrence.—Sly Gap Formation (bed 
H). 

Locality.—52\s. 

Types.—Figured Hypotype: USNM 200832. 


Cariniferella species 
Plate 3, figure 26 


Two large ventral valves and a small, complete 
specimen have many characters, beside size, that sep- 
arate them from Cariniferella alleni n. sp. and C. 
iowensis Stainbrook (1945). These two species have a 
L/W ratio of about 0.80 and are thus well-rounded 
shells with length slightly less than the width. The larg- 
est specimen of the three mentioned as different is 
strongly transverse in its adult form and also in the 
smaller increments that can be seen as growth halts 
on the shell. The specimen (USNM 200835a) measures 
(in mm): length 26.6, midwidth 35.5, hinge width 23.0, 
L/W ratio = 0.75. 

This large species suggests Cariniferella tioga (Hall, 
1867) but differs in having an outline that is more rect- 
angular than broadly elliptical, and in having a smaller 
muscle area. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—521e. 

Types.—Figured Specimen: USNM 200835a. 


Family RHIPIDOMELLIDAE Schuchert, 1913 
Genus RHIPIDOMELLA Ocehlert, 1890 


Rhipidomella species | 
Plate 1, figures 10-21 


About medium size for the genus, nearly circular in 
outline; sides and anterior margins strongly rounded; 
posterior margin somewhat narrowly rounded. Ante- 
rior COmmissure rectimarginate. Beak small, about 
even with dorsal umbo or slightly anterior thereto. 
Hinge width at least one-third of maximum width, 
which is at about mid-valve. Surface finely costellate, 
costellae separated by spaces of width equal to that of 
costellae and numbering three in one mm. 


42 BULLETIN 315 


Ventral valve gently convex in posterior third but 
moderately concave anteriorly; anterior profile very 
broadly concave. Beak small and narrow; interarea 
short, gently concave and strongly apsacline. Cardinal 
extremities depressed. 

Dorsal valve moderately convex in lateral profile 
and broadly domed in anterior profile, the side sloping 
fairly strongly; umbonal and median regions strongly 
swollen; flanks flattened and steep. 

Measurements (in mm).— 


maxi- 

dorsal mum 
length valve width hinge thick- L/W 
(L) length (W) width ness _ ratio 


USNM 204950a 


(loc. 528v) 23:6) 2357 8.6 24.4 8.3 0.97 
USNM 204950b 

(loc. 528v) bes) 24.1 9.2 24.6 7.8 0.91 
USNM 204950c 

(loc. 528v) 22:3 23.0 10.0 25.0 9.8 0.89 
USNM 204950d 

(loc. 528v) 25.0 24.6 10.0 26.2 _- 0.95 
USNM 204950e 

(loc. 528v) 7A Il 22.3 9.0 23:2 7.0 0.91 
USNM 204950f 

(loc. 528v) 23.0 23.0 OF 25.2 7.4 0.91 
USNM 204950g 

(loc. 528v) PANS) 21.0 9.0 22.0 5.9 0.98 
USNM 204950h 

(loc. 528v) 20.0 19.5 9.0 21.2 8.5 0.94 
USNM 204950i 

(loc. 528v) 24.1 232 a 24.4 — 0.98 
USNM 204950) 

(loc. 528v) 19.1 18.7 US 20.4 — 0.94 
USNM 201213 

(loc. 528v) 24.1 23.6 8.0 25.1 8.7 0.96 


Diagnosis.—Nearly circular Rhipidomella having the 
hinge width less than half the maximum shell width. 

Stratigraphic occurrence.—Sly Gap Formation 
(lower part). 

Localities. —506j; 509s; 528v. 

Types.—Figured Specimens: USNM 204950a, b; 
201213. 


Discussion.—Rhipidomella is a rare genus in the 
Upper Devonian rocks of New Mexico. It was found 
at three localities but was not taken in sufficient quan- 
tity to differentiate it satisfactorily from such Middle 
Devonian species as Rhipidomella vanuxemi (Hall, 
1857). It seems to be somewhat more convex dorsally 
than the Middle Devonian species but other details are 
quite similar. Rhipidomella is not only rare in New 
Mexico, it is also unusual in the Upper Devonian of 
New York where it has been called either Rhipido- 
mella leucosia (Hall, 1860a) or R. vanuxemi. The for- 
mer is a Middle Devonian species but is less common 


than R. vanuxemi. Rhipidomella has not yet been found 
in the Independence Shale nor has it been taken from 
the probably related Hackberry Formation (Cerro 
Gordo Member) in Iowa. 


Rhipidomella species 2 


A poorly preserved and fragmentary specimen of 
Rhipidomella was collected from the Onate Formation 
at locality 529w. It is longitudinally and roundly ellip- 
tical as it tapers somewhat anteriorly and posteriorly. 

Types.—Mentioned Specimen USNM 207701. 


Genus THIEMELLA H. S. Williams, 1908 


Thiemella platys new species 
Plate 2, figures 1-10 


Rhipidomella manticula Stainbrook, 1947, p. 303, fig. 4. 


Small for the genus, inequivalved, ventral valve 
having greater depth; transversely elliptical in outline; 
sides rounded; anterior margin gently rounded; hinge 
narrow, measuring slightly more than half maximum 
width, which is at midvalve. Beak small, only slightly 
protuberant; interarea short and curved, apsacline. 
Anterior commissure rectimarginate or nearly so. 

Ventral valve with faint narrow fold in posterior half, 
becoming obsolete anteriorly; sulcus on posterior half 
of dorsal valve becoming obsolete anteriorly. Both 
valves costellate, costellae narrowly rounded, sepa- 
rated by spaces about equal to or slightly wider than 
width of costellae; about ten costellae in space of five 
mm at front of a specimen ten mm long. 

Interior not known. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum_ thick- 
length length width width ness 


USNM 200889 
(Holotype; loc. 527a) 10.0 9.4 5.6? 12.1 BE9) 
USNM 200849 


(Hypotype; loc. 506) 9.6 9.3 6.4 11.4 4.2 
USNM 135261 
(loc. 506) 10.2 10.0 6.7 12.0 — 


Diagnosis.—Small, roundly elliptical, flatly convex 
Thiemella. 

Stratigraphic occurrence. 
Member). 

Localities. —506, q; 507h; 527a, p. 

Types.—Holotype: USNM 200889. Stainbrook’s 
unfigured Paratype: USNM 135261b (Paratype of 
Rhipidomella manticula Stainbrook, 1947). Hypotype: 
USNM 200849. 


Percha Formation (Box 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 43 


Comparison.—This species is much smaller than the 
type species T. villanovia H. S. Williams (1908) and 
the fold and sulcus are obliterated anteriorly at a much 
earlier stage of development. See Thiemella quadrata 
n. sp. for comparison with that species. 


Thiemella quadrata new species 
Plate 2, figures 60-72 


About medium size for the genus, quadrate in out- 
line with rounded cardinal extremities, moderately 
curved sides and slightly curved anterior margin. Hinge 
narrower than shell width, approximately half maxi- 
mum width, which is at about midvalve. Interarea 
short, curved, apsacline. Anterior commissure recti- 
marginate. Surface multicostellate, costellae narrowly 
rounded, increasing by bifurcation and implantation, 
five costellae in two mm at anterior margin. 

Ventral valve gently convex in lateral profile, and 
flatly convex in anterior view. Fold low, extending 
from beak to anterior margin and marked by costellae 
somewhat stronger than those on flanks. Fold outlined 
by shallow lateral impressions that are faintly demar- 
cated from slightly convex flanks. 

Dorsal valve very gently convex in anterior and lat- 
eral profiles. Sulcus shallow, one-third to one-quarter 
valve width, becoming shallower anteriorly. Sulcus 
bounded by two indistinct folds that correspond to 
troughs bounding fold, flattening of sulcus anteriorly 
tending to produce a rectimarginate anterior commis- 
sure in adult. 

Interior unknown. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum thick- 
length length width width ness 


USNM 203533a 


(Holotype) 11.0 10.9 5.0 11.8 3.8 
USNM 203533b 
(Paratype) 12.0 11.7 6.0 13.3 4.8 


Diagnosis.—Medium-sized Thiemella having nearly 
square outline, moderately strong fold and sulcus and 
well-rounded contours. 

Stratigraphic occurrence.—Contadero Formation. 

Locality.—527q. 

Types.—Holotype: USNM 203533a. Figured Para- 
type: USNM 203533b. 

Comparison.—Although the shape of this species is 
similar to that of Thiemella villanovia H. S. Williams 
(1908) it is a smaller species, squarer in outline and 
with sulcus extending farther anteriorly before it be- 
gins to swell near the anterior margin. It differs from 
Thiemella platys n. sp., in its nearly square outline, 


somewhat finer costellation and in the more prominent 
development of the fold and sulcus. 

This is a rare species, only two specimens having 
been found. 


Family TROPIDOLEPTIDAE Schuchert, 1896 
Genus TROPIDOLEPTUS Hall, 1857 


Tropidoleptus platys new species 
Plate 4, figures 13-21 


Of usual size for genus, subquadrate in outline; hinge 
narrower than midwidth; gently concavo-convex. Sides 
rounded; cardinal extremities narrowly rounded. An- 
terior commissure with faint ventrad wave. Surface 
multicostate, costae low and rounded, well-developed 
in posterior and median parts but becoming obsoles- 
cent at the anterior. Median costa of ventral valve 
stronger than the others and opposed by a groove 
deeper than others on the dorsal valve. 

Ventral valve gently convex in lateral profile, most 
convex at midvalve and posterior thereto; anterior 
profile slightly carinate medially but broadly convex. 
Umbolateral slopes moderately steep. 

Dorsal valve fairly evenly and gently concave in both 
profiles, deepest in umbonal region. 

Interior not seen. 

Measurements (in mm).—USNM 20110le (holo- 
type): length 15.0, dorsal valve length 13.9, hinge width 
14.0?, width 17.6, thickness 2.8. USNM 201101b 
(paratype): length 16.7, dorsal valve length ?, hinge 
width 16.8?, width 21.8, thickness ?. 

Diagnosis. —Gently convex, thin-bodied Tropido- 
leptus, with anterior costation becoming obsolescent 
in the adult, especially on the dorsal valve. 

Stratigraphic occurrence. —Onate Formation. 

Localities.—521z; 529g; USGS (Kirk) 540. 3023b. 

Types.—Holotype: USNM 20110le. Figured Para- 
types: USNM 201100, 201101a—c. Unfigured Paratype: 
USNM 201101d. 

Discussion.—Comparison of this New Mexico 
species with Tropidoleptus from elsewhere is difficult 
because the common 7. carinatus (Conrad, 1839) from 
the Appalachian region and New York is quite vari- 
able. This species is abundant in the Kashong Shale 
of the New York Hamilton Group but the Kashong 
specimens are much larger and more convex than 7. 
platys. Some specimens of 7. carinatus show a ten- 
dency to obsolescence of the anterior costae but this 
is not a common feature. Specimens referred to 7. 
carinatus from the Tully Limestone of New York are 
smaller, more convex, and more strongly costate than 
the New Mexico species. Specimens from the Silica 
Shale at Milan, southern Michigan, are about the same 


BULLETIN 315 


size as the New Mexico species but are more strongly 
costate and more concave. 

South American and European species of Tropido- 
leptus are more strongly costate than the New Mexico 
species, and the European 7. rhenanus (Frech, 1897) 
is more transverse. Upper Devonian specimens of 7. 
carinatus are more strongly ribbed than the New Mex- 
ico species. These, like the Hamilton forms, are vari- 
able and in need of systematic revision. 

In New Mexico, Tropidoleptus occurs in swarms, 
mostly of young specimens and with scattered adults, 
in the lower five feet of the Onate Formation on Cap- 
itol Peak and in the Mockingbird Gap (~Johnson Park 
Canyon). 


Order STROPHOMENIDA Opik, 1934 
Suborder STROPHOMENIDINA Opik, 1934 


Superfamily STROPHEODONTACEA 
Sokolskaya, 1960 


Family STROPHEODONTIDAE Caster, 1939 
Subfamily STROPHEODONTINAE Caster, 1939 
Genus STROPHODONTA Hall, 1850 


Strophodonta umbonata new species 
Plate 4, figures 7-12; plate 5, figures 9-18 


This is a medium-sized Strophodonta having a 
strongly convex and umbonate ventral valve and deep- 
ly concave dorsal valve. The hinge is narrower than 
the midwidth; the ventral interarea is anacline and the 
dorsal one is hypercline, so strongly are the valves 
arched. The umbonal region is strongly costellate and 
the costellae extend directly from the beak for ten mm 
where there is a break or growth interruption. At this 
line another generation of costellae is implanted and 
after that another generation appears. On the opposite 
valve the costellae in the first growth segment ante- 
riorly become fasciculate as the new generation of cos- 
tellae appears. 

Measurements (in mm).—USNM 201083 (Holo- 
type): length 22.0, dorsal valve length 16.4, hinge width 
20.0 (based on half measure), midwidth 24.8 (based on 
half measure), thickness 6.5, curvature (surface length 
of ventral valve) 34.0. 

Diagnosis.—Strongly umbonate and deeply con- 
cave Strophodonta having a narrow hinge. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —5S09j, n, p, s; 521d, e, t; 524m, s. 

Types.—Holotype: USNM 201083. Figured Para- 
types: USNM 201084a, b. 

Discussion.—The shape and strongly umbonate form 
of this species are suggestive of Strophodonta boon- 


ensis (Swallow, 1860) from the Snyder Creek Forma- 
tion of Missouri but the Sly Gap species has finer cos- 
tellae, differently arranged, and a narrower shell. This 
and Strophodonta sp. 2 are the only specimens of 
Strophodonta taken from the Sly Gap. Strophodonta 
was on its way toward extinction in Sly Gap time. 


Strophodonta species | 
Plate 5, figures 1-4 


A single specimen of a very thick-shelled Stropho- 
donta has a long (in the direction of valve length), 
nearly orthocline ventral interarea and a moderately 
convex ventral valve in lateral and anterior profiles. 
Unfortunately, the exterior is so badly worn that only 
traces of the costellae appear. These are strong in the 
umbonal region but anteriorly increase suddenly at 
about the middle of the valve. The dorsal valve is mod- 
erately concave, most so just anterior to the umbo. 
The margin on the left side has been damaged but that 
on the right side is gently convex. 

Measurements (in mm).—Length 24.0, dorsal valve 
length 19.6, hinge width 31.0, midwidth 2.0, thickness 
10.2. 

Diagnosis.—Thick-shelled, moderately concave 
Strophodonta having a long ventral interarea. 

Stratigraphic occurrence.—Onate Formation (beds 
C to D). 

Localities.—523, a, e, t. 

Types.—Figured Specimen: USNM 201085. 

Discussion.—We know of no other specimen of 
Strophodonta like this one but the specimen is too 
poor on which to base a species. One other specimen 
of Strophodonta, very poorly preserved, comes from 
the Onate Formation at locality 523. It is small, 15 mm 
long by 17 mm wide, but with all ornament badly worn. 
It is strongly concavo-convex and is suggestive both 
of Strophodonta erratica (Winchell, 1866) of the Trav- 
erse Group of Michigan and of §. costata (Owen, 1852) 
of the Cedar Valley Formation of Iowa. 


Strophodonta species 2 
Plate 5, figures 5-8 


Small for genus, slightly wider than long and with 
hinge narrower than midwidth. Dorsal valve moder- 
ately concave, ventral valve fairly strongly convex in 
both profiles. Surface badly worn, but costellate, cos- 
tellae strong and increasing by implantation. 

Measurements (in mm).—USNM 201181: length 
11.6, dorsal valve length 10.0, hinge width 10.5, mid- 
width 12.6, surface length 16.0, thickness 3.7. 

Diagnosis.—Small coarsely costate Strophodonta 
with narrow hinge. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 45 


Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—526e; 3054b. 

Types.—Figured Specimen: USNM 201181. 

Discussion.—This specimen suggests Strophodonta 
thomasi Fenton and Fenton, 1924 in size, but it does 
not attain the great convexity of that species. It is also 
suggestive of the Middle Devonian S$. erratica (Win- 
chell, 1866) but it is not so wide-hinged nor is it so 
strongly costellate. It seems to be an undescribed 
species, but the single specimen is too poorly pre- 
served to be the basis of a good description. 


Genus STROPHONELLOIDES Caster, 1939 


Strophonelloides dorsoconvexus new species 
Plate 5, figures 40-51 


About average size for the genus, thick-shelled, sub- 
rectangular in outline, wider than long; hinge forming 
widest part; cardinal extremities almost a right angle; 
sides slightly rounded; anterior margin broadly round- 
ed. Interareas long, flat; pseudodeltidium flush with 
interarea; ventral valve interarea strongly apsacline, 
the dorsal one strongly anacline. Surface marked by 
subangular costellae increasing by bifurcation in four 
generations; number of costellae increasing suddenly 
at about midvalve where some costellae produce fas- 
cicles of three finer ones. 

Ventral valve moderately convex in posterior half 
but reverting to strongly concave in anterior half; car- 
dinal extremities flattened. 

Dorsal valve having umbo slightly concave and pos- 
terior half flattened; anterior half strongly convex, an- 
terior profile appearing as a broadly rounded dome. 
Anterior and lateral slopes steep. Cardinal extremities 
flattened to concave. 

Interior not known. 

Measurements (in mm).— 


dorsal 
valve hinge thick- 
length length width midwidth ness 


USNM 200862 

(Holotype; loc. 509n) 24.8 24.6 32.0 32.0 7.0 
USNM 203532 

(loc. 527t) 20.9 20.3 23.3 26.1 6.4 


Diagnosis.—Thick-shelled, wide-hinged Stropho- 
nelloides with strong dorsal convexity and deep ven- 
tral concavity. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—509n, 0, p, r; 521d; 524m; 527t. 

Types.—Holotype: USNM 200862; Figured Para- 
types: USNM 200864, 203532. 

Discussion.—This species strongly suggests Stro- 
Phonelloides hybridus (Hall and Whitfield, 1873) but 


proves to have numerous differences when compared 
with specimens of that species. Its ventral concavity 
and anterior dorsal convexity are much greater than 
those of S. hybridus (Hall and Whitfield, 1873) from 
the Cerro Gordo Member of the Hackberry Forma- 
tion. It also has longer interareas, a less concave dor- 
sal umbo, and lacks the narrow depression that ex- 
tends from the dorsal umbo onto the convex anterior 
of the dorsal valve of S. hybridus. 

This is a very rare species because only six speci- 
mens were found in all the collecting. All the features 
of this species relate it to the §. hybridus tribe of 
Strophonelloides. 


Strophonelloides deeringi Stainbrook 
Plate 5, figures 19-23 


Strophodonta reversa Calvin, 1878, p. 728. 
Strophonelloides deeringi Stainbrook, 1945, p. 19, pl. 2, figs. 25- 
33; 1948, p. 771, pl. 1, fig. 25 (nor fig. 24 = S. parvulus n. sp.). 


Figures of this rare Independence Shale species are 
introduced for comparison with Sly Gap species of 
Strophonelloides. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—S527s. 

Type.—Figured Hypotype: USNM 20086Sa. 


Strophonelloides parvulus new species 
Plate 5, figures 24-39 


Strophonelloides deeringi Stainbrook, 1948, p. 771, pl. 1, fig. 24. 


Small, thick-shelled; elongate rectangular in outline; 
hinge narrower than maximum width which is near 
midvalve; sides gently rounded; anterior margin nar- 
rowly rounded to somewhat nasute. Ventral interarea 
long and strongly apsacline. Profile resupinate. 
Pseudodeltidium flush with interarea. Valves multi- 
costellate, costellae increasing by implantation in two 
or three generations; costellae narrowly angular and 
strong. 

Ventral valve moderately swollen in the posterior 
half, moderately concave at the anterior; cardinal ex- 
tremities flattened. 

Dorsal valve with posterior half gently convex to 
flattened; anterior half strongly convex. Cardinal ex- 
tremities flattened. 

Ventral valve interior having a shallow visceral cav- 
ity and fairly strong anterior ridge; muscle scars oc- 
cupying much of the interior of the visceral cavity; 
palintrope buttressed by a stout pillar. Dorsal valve 
interior having greatly thickened cardinal process, 
muscle platform and median ridge. 


46 BULLETIN 315 


Measurements (in mm).— 


dorsal 
valve _ hinge thick- 
length length width midwidth ness 


USNM 200866 
(Holotype; loc. 


509g) 16.0 (Sei 14.6 115)58) 4.8 
USNM 200867b 

(loc. 506c) 13.6 12.6 12.8 14.2 3.4 
USNM 200867a 

(loc. 506c) 13.8 — 14.0 14.5 — 
USNM 201212 

(loc. 509n) 19.0 18.2 18.9? 21.9 7.3 
USNM 20121 1a 

(loc. 524n) 15.2 14.8 13.6 15.9 3.4 
USNM 201211b 

(loc. 524m) Se) 13.8 15.2 18.2 3.2 


Diagnosis.—Small, thick-shelled Strophonelloides 
with gently rounded, nearly parallel sides and hinge 
narrower than midwidth. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, 1, j; 509g, i, j, 1, n, r, s, z; 521d, 
e, u; 523m; 524n, u; 528x. 

Types.—Holotype: USNM 200866. Figured Para- 
types: USNM 200867a, b, 200872. 

Discussion.—Like all members of this genus, this 
species is quite variable but it is characterized by a 
hinge narrower than the midwidth, obtuse cardinal ex- 
tremities, slightly expanding sides, and narrowly 
rounded anterior margin. It differs from Strophonel- 
loides deeringi Stainbrook (1945) in the narrow hinge, 
rectangular outline, expanded sides and generally 
smaller size of S. parvulus n. sp. and not being strong- 
ly nasute anteriorly, or in having the strong median 
fascicle that forms a prominent fold in Stainbrook’s 
species. Stainbrook’s (1948) specimen from the Sly 
Gap of the Sacramento Mountains was not seen be- 
cause it was not in the collections presented to the 
U.S. National Museum of Natural History. The figure 
of this specimen is strongly suggestive in outline and 
ornament of S. parvulus n. sp. The shape of S. par- 
vulus is similar to that of young S. reversa gravis (Fen- 
ton and Fenton, 1924) but that is a thinner-shelled form 
with finer ornament. 


Subfamily LEPTOSTROPHIINAE Caster, 1939 
Genus NERVOSTROPHIA Caster, 1939 


Nervostrophia extensa new species 
Plate 6, figures 7-14; plate 9, figure 1 


Nervostrophia sp. Stainbrook, 1945, pl. 2, figs. 13-17. 


Of usual size for genus, twice as wide as long, and 
extended laterally into large ears. Valves concavo- 
convex and geniculate about 15 mm anterior to beak. 


Angle of geniculation about 100°, geniculation in pro- 
file rather narrowly rounded with the anterior slope 
steep but shorter than more gentle posterior slope. 
Ventral valve initially costellate with costellae few and 
separated by numerous radial capillae that occupy in- 
terspaces on remainder of the valve. Primary costellae 
very narrowly rounded and discontinuous as normal 
in genus. Ornament of dorsal valve contrasting strong- 
ly to that of ventral valve in having fewer large primary 
costellae and with surface mainly covered by fine ca- 
pillae. 
Measurements (in mm).— 


dorsal 
valve hinge surface thick- 
length length width length ness 


USNM 201088a 
(Holotype; loc. 526y) 19.7 18.8 89:25 23.5 2.9 


USNM 201088c 15.2 14.6 32.4* 17.5 1.8 
USNM 201180a 
(loc. 362a) 17.0 15.9 38.0 20.0 3.6 


Diagnosis.—Nervostrophia having the width about 
twice the length. 

Stratigraphic occurrence.—Independence Forma- 
tion; Sly Gap Formation. 

Localities.—Independence Formation: 362a; 526y; 
Sly Gap: 506c; 521p. 

Types.—Holotype: USNM 201088a. Figured Para- 
types: USNM 201034, 201093. 

Discussion.—In its great width this species is unlike 
any described species of Nervostrophia. Two poorly 
preserved Sly Gap specimens are referred here on the 
basis of their shape, size and ornamentation. 


Nervostrophia geniculata new species 
Plate 6, figures 15-31 


About medium size or moderately large for genus, 
subrectangular in outline and concavo-convex in lon- 
gitudinal section; sides nearly straight to gently con- 
vex; anterior margin broadly rounded; length less than 
greatest width, which is at about midvalve; hinge wide 
but not exceeding the midwidth; beak small, slightly 
protuberant; interarea short, apsacline; pseudodeltid- 
ium small. Ornament of valves different, that of ven- 
tral valve having strong elevated discontinuous cos- 
tellae that often swell locally; primary costellae 
separated by narrow interspaces occupied by one or 
two radial capillae, except on umbonal region where 
radial capillae are more numerous. Dorsal valve with 
primary costellae present but distant and separated by 
numerous radial capillae (as many as five). 

Ventral valve unevenly convex, posterior half gently 
swollen, geniculation taking place near midvalve, an- 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 47 


terior slope convex and steep. Anterior profile broad- 
ly, but not strongly convex. Lateral slopes moderately 
steep; cardinal extremities obtuse. 

Dorsal valve closely fitting the profile of the ventral 
valve, the maximum concavity about two-thirds the 
length from the beak and having steep slopes dipping 
toward the posterior margin of the concavity. 

Interior not seen. 

Measurements (in mm).— 


dorsal maxi- — sur- 
valve hinge mum face _ thick- 
length length width width length ness 


USNM 200934 


(Holotype; 

loc. 521p) 24.8 2355 23.6? 29} 29.5 2.7 
USNM 200933a 

(loc. 506c) 24.7 _— DSi? 32.0 30.0 _ 
USNM 200932 

(loc. 509h) 23.7 — 25.2? 28.8 28.0 _ 


Diagnosis.—Moderately large Nervostrophia genic- 
ulated at midvalve and having the length about 80 per- 
cent of the midwidth. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —5S06c; 509h, 1, q; 521d, p, q, w; 524m, 
n; 526d, r. 

Types.—Holotype: USNM 200934. Figured Para- 
types: USNM 200932, 200933a, b. 

Discussion.—This species differs from Nervostro- 
phia rockfordensis (Fenton and Fenton, 1924) and N. 
solida n. sp., in being geniculate at midvalve, being 
somewhat smaller, and with shorter interareas. It is 
unlike N. calvini Stainbrook (1945) in its longer inter- 
area, more crowded primary costellae, and stronger 
geniculation. 


Nervostrophia mucrospina new species 
Plate 7, figures 1-16 


Small for genus, wider than long, narrowly subrec- 
tangular in outline; strongly concavo-convex in pro- 
file; hinge wider than midvalve and drawn laterally 
into long points of five to seven mm; sides rounded 
and sloping moderately medially; posterolateral ex- 
tremities forming small ears and set off by oblique, 
shallow depression. Anterior margin broadly rounded. 
Interarea short (in direction of valve length), apsacline 
to orthocline; pseudodeltidium flush with surface of 
interarea. Surface of ventral valve costellate, primary 
costellae narrow, elevated, about five or six in space 
of five mm at anterior of average specimen; inter- 
spaces occupied by about four radial capillae; dorsal 
valve surface with primary costellae subdued and ra- 
dial capillae dominating. 


Ventral valve unevenly convex in lateral profile, 
posterior gently convex, median region bent at angle 
of about 110°, front half gently convex; anterior profile 
fairly strongly domed with median region somewhat 
narrowly convex and sides sloping fairly steeply; pos- 
terior and umbonal regions moderately swollen; lateral 
slopes moderately steep. Median anterior half may be 
subnasute. 

Dorsal valve moderately concave, valve fitting 
closely to contours of ventral valve, most concave a 
short distance anterior to midvalve; lateral slopes 
moderately steep; cardinal extremities slightly con- 
cave. 

Interior of neither valve seen. 

Measurements (in mm).— 


dorsal sur- 
valve hinge mid- thick- face 
length length width width ness length 


USNM 201009 


(loc. 521m) 17.8 16.6 20.02 23.3 310) 2225 
USNM 201178a 

(loc. 509n) NB) SS TO Ba IKS AA) 
USNM 201039 

(loc. 521k) 13°45 1228) 16:02) 1657 _— — 
USNM 201024a 

(Holotype; 16.0 — 31.0% 19.0 — 10) 

loc. 509p) 
USNM 201024b 

(loc. 509p) 1637 SUS!) 212871910) — 20:0 
USNM 201024c 

(loc. 509p) 8.7 _ 18.2 12.8 EO 
USNM 201026a 

(loc. 506) 12.4 11.4 14.3? 16.0 — — 
USNM 201026b 

(loc. 506j) 11.5 — 33.8 15.4 — 13.5 
USNM 201209 

(loc. 521e) 12.3 —_ 32.8 16.7 — 13:02 


Diagnosis.—Small, concavo-convex Nervostrophia 
with mucronate cardinal extremities. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —506j; 509f, h, i, n, p, r, s, Z; 521d, e, k, 
m, q, Ss, t, u, w; 524n; 526d, s; 3016. USGS (Kirk) 538. 

Types.—Holotype: USNM 201024a. Figured Para- 
types: USNM 201024b, c; 201026a, b; 201028; 201039. 

Discussion.—This species suggests Nervostrophia 
canace (Hall and Whitfield, 1873) from the Cerro Gor- 
do Member of the Hackberry Formation. It differs, 
however, in being much smaller, more narrowly ge- 
niculated, and with finer ornament. It is less wide than 
N. extensa n. sp., from the Independence Shale. It 
differs from N. multinervosa Stainbrook (1945), also 
from the Independence Shale, in greater curvature, 
finer ornament, and smaller size. 


1g BULLETIN 315 


Nervostrophia multinervosa Stainbrook 
Plate 7, figures 22—25 


Nervostrophia multinervosa Stainbrook, 1945, p. 32, pl. 2, figs. 5— 
8; 1948, pl. 1, fig. 4. 


Two specimens are referred to this species. Another 
fairly large transverse specimen (USNM 201087), is 
free of matrix but is so badly exfoliated and worn that 
little of the ornament is revealed and it can only ten- 
tatively be assigned to N. multinervosa. The other two 
(USNM 201035a, b) are smaller, and reveal only the 
ventral valve, but have fairly well preserved ornament 
that consists of fine radial lines separated by capillae. 
The smaller of these two specimens has the radii more 
crowded than in the specimen herein referred to N. 
cf. N. multinervosa. 

Stratigraphic occurrence.—Sly Gap Formation; In- 
dependence Formation. 

Localities.—S09g; 521d. 

Types.—Figured Specimens: USNM 201035a, b. 

Discussion.—Stainbrook (1945) figured, but did not 
describe, a specimen (USNM 135271) from loc. 506c. 
This is a poorly preserved specimen, lacking the dorsal 
valve but displaying well the ventral valve ornament. 
The species also occurs in the Independence Forma- 
tion at Brandon and Independence, Iowa (Stainbrook, 
1945, p. 33). 


Nervostrophia cf. N. multinervosa Stainbrook 
Plate 7, figures 17-21 


A specimen showing both valves but with exfoliated 
ornament (USNM 201087) measures (in mm): length 
19.6, dorsal valve length 18.7, hinge width 23.2, max- 
imum width 28.2, surface length 26.0, thickness 3.8, 
height 8.7. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—S06c. 

Type.—Figured Specimen: USNM 201087. 


Nervostrophia plana new species 
Plate 6, figures 1-6 


About medium size for genus, wider than long and 
subrectangular in outline; hinge wide, commonly ex- 
tended into acute points, profile slightly concavo- to 
plano-convex. Sides gently rounded; anterior margin 
broadly rounded. Interarea short, apsacline. Pseudo- 
deltidium narrow. Ventral valve marked by low, 
rounded discontinuous primary costellae separated by 
wide interspaces occupied by as many as six radial 
capillae. Dorsal valve similarly ornamented but pri- 
mary costellae more subdued and radial capillae more 
prominent. 


Ventral valve faintly convex in both profiles; um- 
bonal and median regions faintly swollen; postero- 
lateral extremities slightly concave. Mucronate points 
thin and extending laterally about 5 mm. 

Dorsal valve faintly concave in both profiles; pos- 
terior third nearly flat but median third gently concave 
and anterior third flat. Cardinal extremities flattened. 

Ventral valve interior having hinge nearly complete- 
ly dentate; paradental plates stout and solid forming 
an angle of about 57°; muscle area flabellate; adductor 
marks low but thick; pseudodeltidium buttressed by 
an apical callosity. 

Measurements (in mm).— 


dorsal 
valve hinge mid- _ thick- 
length length width width ness 


USNM 20084 1a 


(Holotype; loc. 523u) 22.3 _— 40.0* 27.3 — 
USNM 200841b 

(loc. 523u) 20.0 — 41.2* 26.0* — 
USNM 203531 

(loc. 527u) 22.6 22.0 _ 29.4 <1.0 


Diagnosis.—Very flat Nervostrophia having distant 
low primary costellae in a matrix of fine radial capillae. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —522d, h, n, z; 523a, b, c, d, e, i, s, u, 
x; 524a, 1, k, 1, p, q; 526b, c, h; 527u; 528t; 529g, k, 
m, n; 3023d. 

Types.—Holotype: USNM 20084la. Figured Para- 
types: USNM 200841b; 200929; 203531. 

Discussion.—This species is distinguished immedi- 
ately from Nervostrophia rockfordensis (Fenton and 
Fenton, 1924) and N. solida n. sp., in being flatter than 
either of them and in having much finer ornament. 
Nervostrophia plana suggests N. tulliensis (H. S. Wil- 
liams, 1890), as it is similarly ornamented, but the Tul- 
ly species is smaller and more convex. Nervostrophia 
nervosa (Hall, 1843) is also more convex than the 
Onate species. 


Nervostrophia solida new species 
Plate 7, figures 26-35; plate 8, figures 40, 41 


Nervostrophia thomasi Stainbrook, 1948, p. 772, pl. 1, fig. 1 (not 
fig. 2). 


Large for genus, and thick-shelled; subrectangular 
in outline, sides gently rounded; anterior margin 
broadly rounded; hinge forming widest part of shell in 
well-preserved specimens; valves of unequal depth, 
the ventral valve faintly convex, the dorsal valve from 
slightly convex to gently concave. Ventral interarea 
moderately long; pseudodeltidium flat but bearing a 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 49 


narrow median ridge (monticulus); chilidium narrowly 
rounded, strongly elevated. Surface of ventral valve 
costellate, costellae of two sizes: largest costellae often 
discontinuous; costellae separated by wide interspaces 
marked by fine radial capillae crossed by fine concen- 
tric fila. Dorsal valve surface costellate, the larger cos- 
tellae widely spaced and separated by radial capillae 
that are cancellated by concentric fila more numerous 
than on the opposite valve. The dorsal valve thus gives 
the appearance of a more subdued ornament than that 
of ventral valve. 

Ventral valve faintly convex in both profiles; um- 
bonal region slightly convex; cardinal extremities flat- 
tened. Dorsal valve in old specimens somewhat thick- 
ened submarginally. 

Ventral interior with large flabelliform muscle area 
like that of Protoleptostrophia Caster (1939). Dorsal 
valve interior with adductors deeply inset in adventi- 
tious shell; cardinal process with narrow lobes; bra- 
chiophores short and stout, set close to the cardinal 
process. 

Measurements (in mm).— 


dorsal 
valve hinge mid- thick- 
length length width width ness 


USNM 200842 


(loc. 509k) 26.4 24.6 83:65 33.6 4.9 
USNM 200843a 

(loc. 506c) 29.8 27.8 40.4* 38.2* 4.8 
USNM 200843c 

(Holotype) 29.8 28.0 — 34.0* 3.0 


Diagnosis.—Coarsely costellate, stout-shelled Ner- 
vostrophia having radial capillae on most of shell sur- 
face in both mature and early stages. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, v; 507; 509g, h, i,j, k, 1, r,s; S21c, 
d, e, 1, n, 0, s, u, v, x; 522k; 523c; 524m; 526k; 528); 
3025. USGS (Kirk) 538. 

Types.—Holotype: USNM 200843c. Figured Para- 
types: USNM 200842, 200843a, 200879. Unfigured 
Paratype: USNM 200843b. 

Discussion.—This species is similar to Nervostro- 
phia thomasi Stainbrook (1945) and N. rockfordensis 
(Fenton and Fenton, 1924). From the former it differs 
in having stronger, more distant costellae, and the in- 
terspaces between the costellae in midvalve and the 
anterior are marked by fine radial capillae crossed by 
fine concentric fila. Nervostrophia thomasi is not so 
strongly costellate and the interspaces have fewer ca- 
pillae and stronger concentric lines. Nervostrophia 
rockfordensis has strong costellae that are like those 
of N. solida n. sp. but are more crowded and do not 


have radial intercostellar capillae, but, like N. thom- 
asi, it is provided with strong concentric fila. 

This is a very fragile shell because of the thinness 
of the anterior portions; consequently no complete 
specimens appear in the collection. Some specimens 
are fragments, but are typically ornamented. The 
species is most abundant at locality 506c in the Sac- 
ramento Mountains. 


Genus PS9EUDODOUVILLINA Stainbrook, 1945 


Pseudodouvillina euglyphea Stainbrook 
Plate 8, figures 1-7 


Pseudodouvillina euglyphea Stainbrook, 1945, p. 27, pl. 3, figs. 17— 
20; 1948, p. 771, pl. 1, figs. 28-30. 


This is a small species that is difficult to separate 
from young Douvillina Oehlert (1887) unless the in- 
terior characters are clearly visible. Specimens are 
rectangular, slightly wider than long, with gently 
rounded sides and narrowly rounded front margin. The 
hinge is narrower than the midwidth. The lateral pro- 
file is strongly but unevenly convex with the maximum 
convexity in the posterior third; the anterior profile is 
narrowly convex with precipitous sides and the top 
often somewhat keeled. The valves are ornamented 
by primary costellae, moderately crowded and sepa- 
rated by radial capillae. 

Internally, the ventral valve is like that of Nervo- 
strophia but has prominent divergent paradental plates 
and the long slender diductor scars are separated by 
a prominent median septum. The dorsal valve interior 
is not supplied with the brace plates so characteristic 
of Douvillina. 

A typical specimen (USNM 20105Sa) measures (in 
mm): length 7.8, dorsal valve length 6.8, hinge width 
8.3, maximum width 8.8, surface length 12.0, thick- 
ness 2.2 

Diagnosis.—Small, rectangular, strongly concavo- 
convex shells having an interior like that of Nervo- 
strophia. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S506c, i, v; 509g, i, j, k, m, n, w; 5211, 
0, r; 526f, k; 3045b. 

Types.—Figured Hypotypes: USNM 20105Sa, b. 

Discussion.—This species is commonest in the Sac- 
ramento Mountains. Although Pseudodouvillina was 
placed in synonymy with Nervostrophia Caster (1939) 
by A. Williams (1965, p. H398), the size, shape, or- 
namentation and details of the ventral valve interior 
are sufficiently distinct to warrant the retention of 
Stainbrook’s genus, in our opinion. 


BULLETIN 315 


Genus SULCATOSTROPHIA Caster, 1939 


Sulcatostrophia calvini (Miller) 
Plate 8, figures 20-25 


Strophodonta quadrata Calvin, 1878, p. 728. 

Strophodonta calvini Miller, 1883, p. 298. 

Strophodonta exilis Calvin, 1883, p. 443. 

Sulcatostrophia calvini (Miller) Stainbrook, 1945, p. 29, pl. 3, figs. 
6-11, 23-24; 1948, p. 772, pl. 1, figs. 31, 37. 


This is a small species that occurs with specimens 
of Douvillina Oehlert (1887) and Pseudodouvillina 
Stainbrook (1945); it has both shape and ornament 
similar to those genera. It is readily distinguished, 
however, by having a broad shallow sulcus on the ven- 
tral valve. The interior is like that of Nervostrophia 
Caster (1939). Sulcatostrophia has been put into syn- 
onymy with the latter genus (Williams, A., 1953, p. 
41; 1965, p. H398), but we consider it a valid genus. 

Diagnosis.—Small, rectangular shells ornamented 
as in Nervostrophia but having a broad, shallow sulcus 
on the ventral valve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—5S06c, v; 509m, w, y; 521c, d, 1; 526k. 

Types.—Figured Hypotypes: USNM 201138, 
201059a, b. 

Discussion.—This is a rare species and all speci- 
mens come from exposures in the Sacramento Moun- 
tains. In our opinion, the combination of characters 
listed above serve to set this apart from Douvillina 
and Pseudodouvillina, on the one hand, and Nervo- 
strophia on the other. 


Subfamily DOUVILLININAE Caster, 1939 
Genus DOUVILLINA Oehlert, 1887 


Douvillina arcuata (Hall) 
Plate 8, figures 14-19 


Strophodonta arcuata Hall 1858, p. 492, pl. 3, figs. la—c. 
Douvillina arcuata (Hall) Fenton and Fenton, 1924, p. 87 (for syn- 
onymy); Williams, A., 1953, pl. 11, figs. 7-9. 


In the Hackberry Formation (Cerro Gordo Member) 
of Iowa, this is an abundant but variable form. Most 
specimens, in the early growth stages of the shell, have 
fairly closely crowded costellae with only one or two 
radial capillae intervening. Anteriorly, the larger cos- 
tellae are also crowded. One specimen from the Sly 
Gap conforms to this species. 

Stratigraphic occurrence. 

Locality.—506c. 

Types.—Figured Hypotype: USNM 201044. 


Sly Gap Formation. 


Douvillina contaderoensis new species 
Plate 8, figures 26-39; plate 38, figures 20-23 


About medium size for genus, wider than long; fairly 
deeply concavo-convex. Hinge forming widest part 
when well preserved; cardinal extremities mucronate. 
Sides slightly oblique; anterior margin broadly round- 
ed. Ornament consisting of six or seven larger costel- 
lae in the space of five mm at the front of an average 
specimen; spaces between costellae occupied by fine 
radial capillae. 

Ventral valve moderately to strongly convex with 
maximum convexity at midvalve when viewed in lat- 
eral profile. Anterior profile forming broad moderately 
swollen dome with gently descending sides. Anterior 
slope gently convex and steep; beak small; umbonal 
region gently convex; interarea short, apsacline. 

Dorsal valve closely fitting inside of ventral valve; 
maximum depth slightly anterior to midvalve; sides 
sloping fairly steeply toward midvalve. Interarea short, 
anacline. Ears concave. 

Measurements (in mm).— 


dorsal sur- 
valve hinge mid- face thick- 
length length width width length height ness 


USNM 200970 


(loc. 521) GIGS: 022099 aS Ose Hell 
USNM 200971 

(loc. 521)) 16.0 14.1 22.6 21.4 20.0 5.4? 3.1 
USNM 200991b 

(loc. 509t) 16.0 — 228) 2035 2010 aes _ 
USNM 201009 

(loc. 523n) 17.4 16.0 — 22.8 22.3 6.4 3.0 


Diagnosis.—Fairly large, moderately convex Dou- 
villina. 

Stratigraphic occurrence.—Contadero Formation. 

Localities.—S09t; 521j; 522a; 523n; 524s, t; 526a, g: 
3054k. 

Types.—Figured Hypotypes: USNM 200970; 200971; 
20099 1a, b; 201009. 

Discussion.—This species is fairly common in the 
Contadero Formation and is confined to it. It is distin- 
guished by its relatively large size, for Douvillina, but 
is distinguished by its width and lesser convexity than 
D. arcuata (Hall, 1858) and the other species of the 
genus described here. It is similar to D. cayuta (Hall, 
1867) of the New York Chemung Formation but is 
usually sornewhat smaller and more convex than that 
species. 


Douvillina minuta new species 
Plate 8, figures 8-13; plate 9, figures 2—9 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DuUTRO Syl 


Small for the genus, subcircular in outline with the 
length equal to or exceeding the width; sides and an- 
terior rounded; hinge wide but not equal to midwidth, 
extended into minute ears. Lateral profile strongly but 
unequally convex, the maximum convexity at about 
one-third length from beak; anterior slope long and 
convex; posterior slope short, slightly convex but very 
steep. Ornament consisting of strong angular primary 
costellae forming crest of low plications, median cos- 
tellae most prominent and forming a sort of keel. In- 
terspaces occupied by a few radial capillae. Dorsal 
valve with low angular plications occupied by a cos- 
tella and with interspaces marked by radial capillae. 

Interior not seen. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ surface thick- 
length length width width length ness 


USNM 201056 

(Holotype; 

loc. 506c) Weil 6.9 6.5 7.6 11.0 ?}\1) 
USNM 201076 

(loc. 521n) 7.0 6.4 5.6 6.4 10.0 — 
USNM 201075 

(loc. 524m) 8.5 6.8 6.1 7.4 12.0 — 


Diagnosis.—Very small, rotund, strongly costellate 
Douvillina. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S06c; 521n; 524m. 

Types.—Holotype: USNM 201056. Figured Para- 
types: USNM 201075, 201076. 

Discussion.—The small size and strong ornamen- 
tation distinguish this species from all other described 
species of Douvillina. The species is a rare one. 


Douvillina parviuscula new species 
Plate 9, figures 14-25 


Small for the genus, subquadrate in outline but with 
width slightly greater than length; hinge narrower, equal 
to, or slightly wider than width at midvalve; cardinal 
extremities slightly auriculate; sides nearly straight to 
gently rounded; anterior margin strongly curved. In- 
terarea short, curved, orthocline to anacline. Pseudo- 
deltidium flat but commonly with median elevation 
(monticulus). Surface of ventral valve multicostellate 
and capillate, six primary costellae in five mm at mid- 
valve but two in one mm at the anterior margin, and 
often raised to form crest of a low plication; primary 
costellae increasing abundantly beyond midvalve; in- 
terspaces on posterior half occupied by as many as 
nine radial capillae, but fewer than that anteriorly where 


costellae are less distantly spaced. Dorsal valve with 
primary costellae subdued and radial capillae domi- 
nant. 

Ventral valve strongly convex in lateral profile, the 
posterior slope shorter and steeper than the anterior 
slope; geniculated at three to four mm anterior to beak; 
anterior profile a narrow dome with steeply dipping 
sides. Beak small, umbonal region gently swollen, 
swelling continuing anteriorly to form indistinct fold 
that is lost in strong swelling at midvalve; posterolater- 
al slopes steep and gently concave, producing a small, 
narrowly rounded ear. 

Dorsal valve conforming closely to contour of ven- 
tral valve, thus deeply concave in median region with 
anterior slopes steeply dipping toward posterior; um- 
bonal region flattened; posterolateral region slightly 
concave at small ears. 

Ventral valve interior with thick rim around diduc- 
tor field. Dorsal valve interior not known. 

Measurements (in mm).— 


dorsal sur- 
valve hinge mid- face  thick- 
length length width width length ness 


USNM 201057a 


(loc. 521e) 10.6 9.5 9.4% 10.4 15.5 2.4 
USNM 201054 

(Holotype; 

loc. 509r) 10.8 9.8 10.8 11.4 14.0 2.0 


Diagnosis. —Small, strongly convex, narrow-hinged 
Douvillina having small ears. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S06c, 1, j; 509h, j, n, r, s, y, z; 521d, e, 
m, q, r; 524m, n; 526f, m; 528y; 3025a; 3053; 3054f; 
USGS (Kirk) 543. 

Types.—Holotype: USNM 201054. Figured Para- 
type: USNM 201057a. Unfigured Paratypes: USNM 
201057b-e. 

Discussion.—This species is slightly smaller than 
Douvillina navicula Stainbrook (1945) from the Inde- 
pendence Formation of Iowa, but it also differs in 
lacking the somewhat keeled median region, in having 
less pronounced ears, and in not having strong mu- 
cronate points extending from the ears. 


Douvillina rectangulata new species 
Plate 9, figures 26-37 


Small, length and width nearly equal, deeply con- 
cave, ventral valve moderately geniculated just pos- 
terior to midvalve, dorsal valve fitting interior of ven- 
tral valve closely. Hinge slightly wider than midvalve 


Di BULLETIN 315 


and extended onto small ears, not quite forming right 
angle. Sides subparallel and anterior margin broadly 
rounded. Ventral interarea short, curved, orthocline 
to anacline; dorsal interarea short, hypercline. Surface 
costellate and capillate, costellae numbering about 
three in two mm and separated by up to six capillae. 

Ventral valve strongly convex in lateral profile and 
narrowly domed in anterior profile; lateral slopes steep; 
ears narrowly rounded. Umbonal region very slightly 
protuberant beyond the posterior margin. 

Dorsal valve deepest at midvalve with sides and an- 
terior sloping steeply toward midvalve; ears narrowly 
concave. 

Interior of both valves not known. 

Measurements (in mm).— 


dorsal sur- 
valve hinge mid-_ face thick- 
length length width width length ness height 


USNM 201031 
(Holotype; 
loc. 521e) 919) 8:8) 106% 1025 1425) 2247 5.0) 
USNM 201030 
(loc. 509n) 10.4 9.0 QP ili) 16.0 2.8 5.4 


Diagnosis.—Small Douvillina having length and 
width nearly equal, subparallel sides, and moderate 
geniculation posterior to midvalve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —509ha, n; 521e. 

Types.—Holotype: USNM 201031. Figured Para- 
type: USNM 201030. 

Discussion.—This species is most similar to Dou- 
villina parviuscula n. sp., but differs in having a more 
compact form, more rectangular outline, stronger con- 
vexity and, consequently, deeper dorsal valve; it also 
has a different geniculation and more subdued orna- 
ment. This species is rare in the Sly Gap. 


Douvillina species | 
Plate 9, figures 45-50 


About medium-sized to large for genus, deeply con- 
cave dorsally and strongly convex on ventral side. 
Hinge slightly narrower than midvalve width, where 
shell is widest. Anterior strongly rounded; sides mod- 
erately rounded. Interarea strongly apsacline and 
moderately long. Valves geniculated at about mid- 
valve, the anterior slope convex and steep, the pos- 
terior slope flattened and steeper than anterior one. 
Surface of umbonal part marked by distant narrow 
costellae separated by numerous radial capillae; pri- 
mary costellae increasing and becoming crowded at 
anterior side of geniculated part of shell and with few 


radial capillae intervening. Dorsal valve with costellae 
often elevated on low plications. 

Measurements (in mm).—USNM 201040: length 
13.4, dorsal valve length 11.9, hinge width 15.5 (based 
on half measure), midwidth 17.4, surface measure 18.0, 
thickness 2.7. 

Diagnosis.—Medium to large Douvillina with dis- 
tant primary costellae in posterior half but crowded 
primary costellae on the anterior slope. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c; 509j, n, s, x; 521d, m. 

Types.—Figured Specimen: USNM 201040. 

Discussion.—Not one of the localities listed above 
yielded enough specimens to get a good idea of this 
species. It is larger than Douvillina parviuscula n. sp., 
although similarly marked. The specimens referred here 
vary in degree of convexity. 


Douvillina species 2 
Plate 8, figures 42-54 


About medium size for genus, wider than long, sub- 
rectangular, hinge width variable, usually less than 
midwidth; sides gently rounded; anterior more broadly 
rounded. Geniculation sharp, occurring 3-4 mm an- 
terior to beak. Interarea anacline. Ventral valve 
strongly but unevenly convex in lateral profile; an- 
terior three-fourths moderately convex, posterior 
quarter narrowly curved with dorsal slope directed an- 
teriorly. Surface ornamented as in Douwvillina parvius- 
cula n. sp. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum = surface thick- 
length length width width length ness 


USNM 203561la 


(loc. 527v) es 6.0 7.3 92 12.0 Do 
USNM 203561b 

(loc. 527v) DOS: 51 10.0 10.6? 17.0 — 
USNM 204951 

(loc. 527v) 7.8 6.4 8.0 8.9 12.0 2.9 


Diagnosis. —Small, strongly geniculated Douvillina 
with short posterior slope inclined ventrally. 

Stratigraphic occurrence.—Contadero Formation 
(coral bed). 

Localities. —523r; 524s; 526a, g; 527v. 

Types.—Figured Specimens: USNM 20356la, b; 
204951. 

Discussion.—This species is similar in size and or- 
nament to Douvillina parviuscula n. sp. but differs in 
having the posterior slope so strongly curved that it is 
actually directed ventrally, so strong is the genicula- 
tion. 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DUTRO 


Genus DOUVILLINARIA Stainbrook, 1945 


Douvillinaria diversa new species 
Plate 9, figures 51, 52; plate 10, figures 1-14 


Fairly large for the genus, squarish in outline; hinge 
narrower than greatest width which is at or near mid- 
valve; valves unequal in depth, dorsal valve deeper; 
sides gently rounded; anterior margin strongly round- 
ed; hinge straight; anterior commissure rectimargin- 
ate. Interareas nearly equal in length (direction of valve 
length); ventral interarea strongly apsacline and dorsal 
strongly anacline. Pseudodeltidium narrow, flush with 
interarea but marked medially by narrow elevation or 
monticulus. Surface multicostellate, costellae fine and 
rounded at the posterior but thickening anteriorly and 
capping low plicae; costellae increasing mainly by im- 
plantation. Costellae separated by wide interspaces 
occupied by fine longitudinal threads or capillae. 

Pedicle valve usually unevenly convex, flatly con- 
vex posteriorly but gently convex anteriorly; anterior 
profile broadly and slightly convex. Ventral valve 
slightly concave near midvalve and anterior half gently 
convex. 

Dorsal valve, in lateral profile, gently convex in pos- 
terior third but flattened or slightly concave in anterior 
half to two-thirds. Umbonal region flattened or slightly 
concave but region anterior to umbo convex, to about 
one-third the valve length, becoming slightly concave 
or flat to anterior margin. Anterior profile broadly and 
gently convex with median part narrowly humped and 
sides gently sloping to margins. 

Ventral valve interior having a short, broadly heart- 
shaped muscle area with elevated rim. Dorsal valve 
interior with all structures thickened; brachiophores 
conspicuous; brace plates thick, expanding anteriorly 
and making contact posteriorly with the anterior end 
of the median ridge. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum_ thick- 
length length width width ness 


USNM 200870a 


(loc. 509h) 13.3 13.1 10.7 14.1 3.4 
USNM 200870b 9.1 8.9 9.1 Q7/ YS) 
USNM 200869a 

(Holotype; loc. 521c) 14.1 13.9 12.4 16.0 387) 
USNM 201191a 

(loc. 506c) 12.8 12.8 11.2 14.3 5.0 
USNM 201191b 15.0 14.5 14.7 le 4.0 


Diagnosis.—Fairly large Douvillinaria without a 
strong median fold and sulcus and with the initial stages 
usually biconvex, convexo-plane or convexo-concave. 

Stratigraphic occurrence.—Sly Gap Formation. 


nn 
Ww 


Localities.—S06c, j; 509h, k; 521c, m, s, t, v; 524m, 
u; 526d. 

Types.—Holotype: USNM 200869a. Figured Para- 
types: USNM 200868, 200869b, 200870a. Unfigured 
Paratypes: USNM 200870b, 201191a. 

Discussion.—This species attains the size of D. var- 
iabilis (Calvin, 1878) but has a different lateral profile 
which is usually convexo-plane in the posterior half 
but concavo-convex in the anterior half. The dorsal 
valve in early life is strongly convex but in maturity it 
flattens or becomes gently concave. It is larger than 
D. perversa Stainbrook (1945) and has a different pro- 
file. It is smaller than D. delicata (Fenton and Fenton, 
1924), which has a concave dorsal valve and does not 
have the early part of that valve swollen. 


Douvillinaria variabilis (Calvin) 
Plate 9, figures 38-44 


Strophodonta variabilis Calvin, 1878, p. 727; Hall and Clarke, 1892, 
p. 289, pl. 15B, figs. 4-6. 

Douvillinella variabilis (Calvin) Stainbrook, 1935, pl. 83, fig. 9. 

Douvillinaria variabilis (Calvin) Stainbrook, 1945, p. 24, pl. 3, figs. 
1-4, 25; 1948, p. 771, pl. 1, figs. 26, 27. 


Specimens assigned to this species have a fairly con- 
spicuous fold on the ventral valve and a sulcus on the 
dorsal valve. The specimens are large and compare 
favorably with material from the Independence For- 
mation of Iowa. 

Diagnosis.—Large, moderately concavo-convex 
Douvillinaria with a well-marked fold and sulcus. 

Stratigraphic occurrence.—Sly Gap Formation; In- 
dependence Formation. 

Locality.—506c; 509h, i; 521e, m; 526m. 

Types.—Figured Hypotypes: USNM 20087la, b. 


Family uncertain 
FLOWERIA new genus 


Medium to large size, subrectangular in outline with 
sides gently rounded, anterior margin broadly round- 
ed; valves plano-convex to unequally biconvex, great- 
er convexity on ventral valve. Interareas on both 
valves, that of dorsal valve greatly reduced, that of 
ventral valve moderately long (longitudinally). 
Pseudodeltidium moderately convex and rounded; 
chilidium reduced, marked medially by a depression 
or notch. Anterior commissure variable. Surface cos- 
tellate, costellae appearing in several generations. Shell 
substance without taleolae (not pseudopunctate). 

Ventral valve interior with thick teeth; muscle field 
subcircular. 


SZ BULLETIN 315 


Dorsal valve with bilobed cardinal process, each lobe 
of which is grooved medially; cardinal process welded 
to brachiophores, which are short rods encased in tis- 
sue; forming a cuplike socket. Anterior surface of car- 
dinalia medially marked by small angular node repre- 
senting depression in chilidium that divides cardinal 
process lobes. Adductor field roundly flabellate and 
marked medially by low or indistinct myophore. 

Type species.—Orthis prava Hall, 1858, p. 490 
(=Schuchertella prava (Hall) Fenton and Fenton, 1924, 
p. 117, pl. 20, figs. 21-28). 

Diagnosis. —Medium sized costellate shells having 
ventral valve deeper and more convex than the flat or 
nearly flat dorsal valve. Interareas unequally devel- 
oped. Brachiophores contained in cuplike sockets. 

Discussion.—This genus is very similar in form, 
outline and ornament to Schuchertella (S. lens White, 
1862, type species) but differs in having a much re- 
duced dorsal interarea and having no pseudopunctae. 
It is thus similar to Eoschuchertella Gratsianova (1974) 
but differs in cardinalia, as the Russian genus lacks 
the cuplike cardinalia and has a lesser development of 
the cardinal process. Gratsianova likens her genus to 
Schuchertella but her figures do not conform well to 
the Mississippian species; however, she makes clear 
that Eoschuchertella is not pseudopunctate. Her fig- 
ures do not show the cuplike brachiophore supports 
and the cardinal process figured is more erect than that 
of Schuchertella. Schuchertellopsis Maillieux (1939) 
was erected for brachiopods of this type which are 
cemented to other shells. Floweria n. gen. has been 
found cemented to spiriferids. It is possible that 
Schuchertellopsis is close to, if not the same as, Flow- 
eria but the inner details are not known. 

The American “‘schuchertellas’’, as indeed all the 
related forms, are in great need of revision. The prob- 
lem is complicated by the homeomorphic development 
of impunctate and pseudopunctate forms. Many of the 
species of the Early, Middle and early Late Devonian 
are impunctate while some of those of the latest De- 
vonian (Famennian-Percha) are pseudopunctate. The 
pseudopunctate forms should not be classified with the 
impunctate ones as Havlicek (1967, p. 197) has done 
in making his composition of the Schuchertellidae. 

The genus is named for Dr. Rousseau Flower, So- 
corro, New Mexico, who helped us in our field work 
and provided us with specimens from his own col- 
lecting in the Devonian of New Mexico. 


Floweria prava (Hall) 
Plate 10, figures 15-27 


Orthis prava Hall, 1858, p. 490. 

Orthotetes prava (Hall) Hall and Clarke, 1892, p. 225, pl. 11A, fig. 
13. 

Schuchertella prava (Hall) Fenton and Fenton, 1924, p. 117, pl. 20, 
figs. 21-28; Stainbrook, 1945, p. 34, pl. 2, figs. 19, 20; ? 1948, p. 
772, pl. 1, figs. 7, 8. 


This is a variable species not only in the Hackberry 
Formation (Cerro Gordo Member) where it is best 
known, but also in the Sly Gap of New Mexico. These 
shells are cemented early in life; consequently, their 
ultimate shape depends in part on the nature of the 
host and their mode of growth later after they have 
broken from their attachment. The Sly Gap specimens 
agree well in ornament with the Iowa species, there 
being usually two costellae per mm, but in some parts 
of a shell there may be three costellae in a mm. Flow- 
eria prava commonly has an obtuse cardinal extremity 
but some are acute, nearly at a right angle, occasion- 
ally with a small and insignificant ear. So it is with the 
New Mexico specimens. A wide one (USNM 200855) 
has a small ear on one side and the other is broken. 
Most specimens from the Sly Gap Formation have ob- 
tuse cardinal extremities. The convexity of the dorsal 
valve is variable; most specimens are moderately con- 
vex but a fairly large number are flat or concave, prob- 
ably a reflection of growth on an unusual host. 

The pseudodeltidium of the New Mexico specimens 
is often minutely and fairly evenly wrinkled and a few 
specimens from the Hackberry show a similar wrin- 
kling. No interiors of the New Mexico specimens were 
found. 

Measurements (in mm).— 


dorsal 
valve hinge mid- thick- 
length length width width ness 


USNM 200855 


(loc. 509h) 15.7 14.6 22.6? 23.2 6.0 
USNM 200854 

(loc. 509z) 15.7 14.6 16.7 20.0 6.4 
USNM 201190 

(loc. 509p) 17.2 16.7 21.8? 24.1 6.8 
USNM 200874 

(loc. 521e) 14.3 12.5 14.2 17.3 7.0 


Diagnosis. —Small to medium flattish Floweria of 
subrectangular outline having two or three costellae in 
a mm at the anterior margins of adults, and with a flat 
to slightly convex dorsal valve. 


DEVONIAN BRACHIOPODS OF NEw MExico: COOPER AND DUTRO 


Stratigraphic occurrence.—Sly Gap Formation; 
Hackberry Formation (Cerro Gordo Member). 

Localities. —506c, i, j; 509i, n, p, s, v, x, Z; 52Ic, e, 
m; 524m, n; 3054g. 

Types.—Figured Hypotypes: USNM 200854, 200874. 

Discussion.—The species is very rare in the Sly Gap 
Formation. 


Floweria aff. F. chemungensis (Conrad) 
Plate 11, figures 1-5 


Strophomena chemungensis Conrad, 1842, p. 257, pl. 14, fig. 12. 
Streptorhynchus chemungensis (Conrad) Hall, 1867, p. 67, pl. 10. 


This large, misshapen Floweria is very different from 
F. prava. The specimen measures 25 mm in length and 
30.2 mm in width, but its right side is narrower than 
the left (seen from the ventral side) apparently due to 
impingement against some obstacle to its uniform 
growth. The growth of this shell was very uneven at 
the beak as well as at the sides. The beak is elongated 
and narrow with a narrow interarea for half its length 
but the interarea suddenly widens to measure 14.5 mm 
on the left side (seen from the dorsal side) of the shell; 
but it is only 9.7 mm wide on the right side. In lateral 
view, the ventral valve is resupinate; the dorsal valve 
is fairly strongly convex at midvalve but is flattened 
anteriorly. The primary costellae are narrowly round- 
ed and are separated by one or two costellae of lesser 
strength. Nine or ten costellae occupy a space of five 
mm at the front margin. 

Interior not seen. 

Diagnosis.—Fairly large, irregularly shaped Flow- 
eria, having a resupinate ventral valve, with nine or 
ten costellae in five mm at the anterior. 

Stratigraphic occurrence.—Contadero Formation 
(top). 

Locality.—523n. 

Types.—Figured Hypotype: USNM 200875. 

Discussion.—This specimen suggests relationship to 
Schuchertella [=Floweria| chemungensis (Conrad) 
which also has a resupinate ventral valve. Although 
the eastern species is often larger and flatter, it has 
about the same type of ornament. 

Two other fragmentary specimens of a large schu- 
chertellid may belong here but they are not complete 
enough for identification. One is from the Rhodes Can- 
yon Formation at locality 523p; the other comes from 
the Sly Gap Formation at locality 526n. 


n 
in 


Floweria transversalis new species 
Plate 10, figures 41-46 


About medium size for genus, widely rectangular in 
outline, length about two-thirds the width; concavo- 
convex; sides rounded; hinge slightly less wide than 
midwidth; anterior commissure narrowly flexed dor- 
sally. Cicatrix of attachment occupying about half the 
surface of the ventral valve. Interarea moderately long: 
apsacline. Pseudodeltidium strongly convex. Surface 
costellate, costellae fairly uniform in size, three to a 
mm at the front margin. 

Ventral valve fairly strongly concave and having 
large attachment surface; anterior profile nearly flat; 
cardinal extremities slightly deflected dorsally to form 
small obtuse ears. Anterior with narrow sulcus. Dorsal 
valve moderately convex in lateral profile and broadly 
convex in anterior profile. Median region with low nar- 
row fold not extending posteriorly beyond midvalve. 
Umbo lateral slopes fairly steep and posterolateral ex- 
tremities dorsally deflected. 

Interior not known. 

Measurements (in mm).—USNM 200873 (Holo- 
type): length 19.0, dorsal valve length 16.7, hinge width 
27.6, maximum width 30.5, thickness 7.3. 

Diagnosis.—Concavo-convex Floweria having a 
large attachment surface, length two-thirds the width, 
anterior commissure uniplicate. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—362a. 

Types.—Holotype: USNM 200873. 

Discussion.—Compared to the largest specimen of 
F. prava (Hall, 1858) from Rockford, Iowa, and vicin- 
ity, which has similar length-width dimensions, the In- 
dependence specimen is ventrally concave, rather than 
convex as is the Hackberry specimen. The dorsal valve 
of the Independence specimen is much more convex 
than F. prava from Rockford. The volumes of the 
valves of the Hackberry species are the reverse of 
those of the specimen from the Independence For- 
mation. 


Floweria magnacicatrix new species 
Plate 10, figures 28-33 


About medium-sized, rectangular in outline, length 
about two-thirds width. Cardinal extremities with small 
ears; sides gently rounded; valves convexo-concave. 
Hinge straight, slightly wider (see *‘Discussion’’) than 
midwidth; interarea strongly apsacline, moderately 


14 BULLETIN 315 


long; pseudodeltidium strongly convex. Anterior com- 
missure with a broad dorsal fold. Surface costellate, 
costellae narrowly rounded, strong, about four in five 
mm at the anterior of the dorsal valve, and separated 
by spaces slightly wider than the costellae. 

Ventral valve depressed by a wide cicatrix that oc- 
cupies more than half surface; anterior third forming 
a broad sulcus. Dorsal valve gently convex in lateral 
profile, most convex in umbonal region. Lateral slopes 
gentle; margins somewhat deflected and flattened. 

Measurements (in mm).—USNM 200855 (Holo- 
type): length 15.6, dorsal valve length 14.6, hinge width 
22.8 plus, midwidth 23.2, thickness 5.2, height 6.4. 

Diagnosis. —Floweria having a large cicatrix of at- 
tachment, widely rectangular outline, and strong cos- 
tellation. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—509h. 

Types.—Holotype: USNM 200855. 

Discussion.—This species has characters suggestive 
of Floweria prava (Hall, 1858) and F. transversalis, 
n. sp. It resembles the former in outline but differs in 
having a wider hinge, more prominent fold and sulcus, 
stronger ornament, and much larger cicatrix of attach- 
ment. In the latter respect, F. magnacicatrix is like F. 
transversalis but the ornamentation of the Sly Gap 
species is stronger and the costellae are separated by 
wider interspaces. 


Superfamily DAVIDSONIACEA W. King, 1850 
Family SCHUCHERTELLIDAE Williams, 1953 
Subfamily SCHUCHERTELLINAE Williams, 1953 
Genus SCHUCHERTELLA Girty, 1904 


Schuchertella percha (Stainbrook) 
Plate 10, figure 47 


Schuchertella chemungensis Kindle, 1909, p. 16, pl. 3, figs. 2-4a. 
Schellwienella percha Stainbrook, 1947, p. 304, pl. 45, figs. 9, 12, 
13. 


Sectioning of the ventral beak of a specimen from 
locality 509b failed to show the pseudo-dental plates 
that usually characterize Schellwienella 1. Thomas 
(1910). This Percha Formation (Box Member) species 
appears to be a true Schuchertella having a fairly uniform 
strophomenoid convexo-concave shape. A few spec- 
imens have a greater development of the interarea than 
typical and these have a somewhat distorted umbonal 
region, but the ornament is similar to the typical forms 
and there is no apparent reason to exclude them from 
the species. 

The dorsal valve has a short but well-developed 
chilidium and the cardinal process is bilobed and mod- 


erately elevated. The ridges producing the sockets are 
fairly thick. A poorly developed myophragm appears 
anterior to the cardinal process. This species grows to 
a much larger size than indicated by Stainbrook. 

The species is strongly and finely pseudopunctate 
and thus must be assigned to Schuchertella rather than 
to Floweria. 

Measurements (in mm).—A large and fairly sym- 
metrical specimen (USNM 201133) from locality 3026a 
measures: length 27.3, dorsal valve length 23.2, hinge 
width 27.0, maximum width 32.1, and thickness 8.3. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b; 507a, b, h, 0, v; 509, b; 527a, 
d, p; 3026, 3026a; 3034; 3035; 3042. 

Types.—Figured Hypotype: USNM 201133. 


Schuchertella resupinata new species 
Plate 10, figures 34-36 


About usual size for genus, transversely and broadly 
elliptical in outline, sides rounded; maximum width at 
or posterior to midvalve; hinge wide but not equalling 
maximum shell width. Interarea fairly long, nearly flat 
and strongly apsacline. Pseudodeltidium large and 
convex; chilidium well-developed, in close contact with 
the pseudodeltidium when the valves are closed. Sur- 
face costellate, costellae unequal in size, each suc- 
ceeding generation of three intercalated ones smaller 
than preceding one and separated by spaces as wide 
or wider than costellae. Costellae of all generations 
numbering 13 in five mm at the front margin in inter- 
spaces occupied by closely crowded concentric fila. 

Ventral valve with a sigmoidal lateral profile, the 
posterior umbonal region convex but the anterior be- 
coming concave. Anterior profile having a narrowly 
domed median region with long flattened lateral slopes. 
Dorsal valve with concave umbonal region and convex 
anterior but flattened posterolaterally. 

Interior not seen. 

Measurements (in mm).—USNM 201189a (Holo- 
type): length 23.0, dorsal valve length 16.2, hinge width 
26.4, maximum width 29.3, thickness 9.0?. 

Diagnosis.—Resupinate Schuchertella having 13 
costellae in five mm at the anterior of an adult. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—509e. 

Types.—Holotype: USNM 201189a. Unfigured 
Paratype: USNM 201189b. 

Discussion.—This species differs from Schuchert- 
ella percha (Stainbrook, 1947) in having the posterior 
of the ventral valve strongly convex and the umbonal 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 5)7/ 


region of the dorsal valve concave. Stainbrook’s 
species is much more evenly convexo-concave, al- 
though the ornament is similar. 


‘*Schuchertella’> species 


A small “‘Schuchertella”’ with a crushed dorsal valve 
has alternating fine and coarse costellae, one or two 
of the finer ones separated by the stronger costellae. 
The specimen is subrectangular, wider than long. The 
ventral valve is nearly flat and the dorsal valve fairly 
strongly convex. The specimen is too poorly pre- 
served for satisfactory comparison with other species. 

Measurements (in mm).—Length 13.8, dorsal valve 
length 11.7, hinge width 13.4, maximum width 17.8, 
thickness greater than 5.5. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—523u. 

Types.—Described Specimen: USNM 201188. 


Suborder CHONETIDINA Muir-Wood, 1955 
Superfamily CHONETACEA Bronn, 1862 
Family CHONETIDAE Bronn, 1862 
Subfamily DEVONOCHONETINAE Muir-Wood, 1962 
Genus DEVONOCHONETES Muir-Wood, 1962 


Devonochonetes? species 
Plate 10, figures 37-40 


About medium size for genus, rectangular in outline, 
length about two-thirds midwidth; hinge forming the 
widest part; cardinal extremities acute. Sides nearly 
straight or slightly oblique; anterior margin broadly 
rounded. Posterior margin with at least three short, 
oblique spines. Surface finely costellate, five or six 
costellae in one mm. 

Ventral valve gently convex in both profiles; median 
region gently swollen; slopes to cardinal extremities 
gently concave. Dorsal valve flatly concave. 

Ventral interior with a long median septum; dorsal 
valve with a low, small, bilobed cardinal process with 
low median ridge. 

Measurements (in mm).— 


hinge mid- thick- 
length width width ness 
USNM 203542 
(loc. 527w) 12.5 18.2* 18.6 — 
USNM 203543a 
USNM 201089a 11.1 19.8 16.4 — 
(loc. 526b) 14.2 23.4 21.5 _ 


Diagnosis.—Finely costellate, thin Devonochonetes 

with length 65-70 percent of the midwidth. 
Stratigraphic occurrence.—Onate Formation. 
Localities.—526b; 527w. 


Types.—Figured Specimens: USNM 201089a, b; 
203542; 203543a. 

Discussion.—These specimens are so poorly pre- 
served that they cannot be definitely identified as to 
genus or intelligently compared with any already de- 
scribed species. 


Genus STRIATOCHONETES Mikryukov, 1968 


Striatochonetes nanus new species 
Plate 11, figures 13-18 


Small, subrectangular in outline, wider than long; 
sides rounded; anterior margin broadly rounded. Hinge 
straight, narrower than maximum width, which is at 
midvalve; cardinal extremities obtusely rounded. Pos- 
terior margin with two or three short, oblique spines. 
Surface costellate, except for cardinal extremities; 
about 21 costellae on a side, increasing by implanta- 
tion and bifurcation in three generations. 

Pedicle valve moderately convex in lateral profile 
and broadly domed, but moderately elevated, in an- 
terior profile. Umbonal and median regions swollen. 
Interarea fairly long, curved, orthocline to anacline. 
Pseudodeltidium short and convex. 

Dorsal valve closely fitting the ventral valve and 
thus moderately concave. Interarea strongly hyper- 
cline. Chilidium thin and short; cardinal process tri- 
lobed, filling delthyrial cavity of ventral valve; median 
lobe of cardinal process grooved. 

Ventral valve interior with long, thin median sep- 
tum. Dorsal valve having small, slender prosocket 
ridges, small, thin, divergent anderidia and a thick me- 
dian septum reaching to about midvalve. Lateral septa 
bifurcating near midvalve and dying on the anterior 
slope. 

Measurements (in mm).— 


dorsal 
valve hinge mid-_ thick- 
length length width width ness _ height 


USNM 201077b 


(loc. 522d) 5:3 — 5.6 Hell — 2.0 
USNM 201078 

(loc. 524g) 5.0 — 5.5 7.0 — 1.4 
USNM 201079a 

(loc. 522i) 5.6 — Sa7/ 7.8 — 1.3 


Diagnosis.—Small Striatochonetes having length 
0.7-0.8 the width, and about 40 costellae. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—522d, e, 1; 523b, f, s, w, x; 524a, g, h, 
j, 1; 526h; 529j, n; 3025a; 3053. 

Types.—Holotype: USNM 201079b. Figured Para- 
types: USNM 201077a, b; 201078a, b; 201079a. Un- 
figured Paratype: USNM 201079c. 


BULLETIN 315 


Discussion.—This species resembles several small 
chonetids from the Devonian: C. scitulus Hall (1857), 
C. setigerus Hall (1843), C. schucherti Cleland (1911), 
C. lepidus Hall (1857), and C. scitulites Cooper (1945). 
The New Mexico species is smaller than all of these 
except C. setigerus, but the latter has erect cardinal 
spines rather than oblique ones. 


Striatochonetes subcarinatus new species 
Plate 11, figures 6—12 


Small, roundly elliptical in outline, strongly conca- 
vo-convex in lateral profile; sides strongly rounded and 
anterior broadly rounded. Hinge narrower than great- 
est width, which is at midvalve. Ventral interarea 
moderately long, curved, anacline; pseudodeltidium 
well-developed. Cardinal process with trilobed myo- 
phore bounded by small chilidial plates. Dorsal valve 
interarea short, hypercline. Surface costellate, costel- 
lae low, strongest on umbonal region, numbering about 
four in one mm at the anterior of the dorsal valve. 

Ventral valve strongly convex in lateral profile, 
roundly carinate in anterior profile and with straight, 
steeply-dipping lateral slopes. Umbonal region sub- 
carinate and marked by the strongest costellae. Dorsal 
valve deeply concave and with a deeply concave um- 
bonal region. Deepest in midvalve and with steep slopes 
to midvalve. 

Interior not known. 

Measurements (in mm).—USNM 201080 (Holo- 
type): length 6.8, dorsal valve length 5.5, hinge width 
6.2, midwidth 8.3, thickness 1.5, height 3.8. 

Diagnosis.—Striatochonetes with strongly convex 
and subcarinate anterior profile. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—523f. 

Types.—Holotype: USNM 201080. 

Discussion.—This species is about the same size as 
Striatochonetes nanus n. sp., but differs in being much 
more convex, especially in anterior view, in having 
finer costellae, a narrower hinge and a deeper dorsal 
valve. The species appears to be very rare. 


Genus RETICHONETES Muir-Wood, 1962 


Retichonetes obscurus new species 
Plate 11, figures 19-25 
Small, transversely rectangular in outline; hinge wide 
but narrower than maximum shell width which is at 
about midvalve; sides rounded; anterior margin broad- 
ly rounded; posterior margin bearing two or three slen- 
der, long (2.25 mm) vertical spines. Interarea short, 
apsacline; delthyrium closed by convex pseudodeltid- 
ium; notothyrium plugged by cardinal process; small 


remnantal chilidium present. Surface costellate, cos- 
tellae rounded and separated by interspaces narrower 
than costellae, about five costellae in one mm at an- 
terior of an adult specimen. 

Ventral valve moderately strongly convex in lateral 
profile; broadly but unevenly convex in anterior pro- 
file, the median region swollen and the sides descend- 
ing steeply but lessening to gently concave toward the 
cardinal extremities. Umbonal region not strongly 
swollen and not protruding posteriorly beyond the 
posterior margin. 

Dorsal valve closely fitting into ventral valve and 
having its most concave part slightly posterior of mid- 
valve; cardinal extremities slightly concave; slopes to 
midvalve steepest laterally but less so at the anterior. 

Interior of ventral valve with median septum reach- 
ing to about one-third valve length. Dorsal valve hav- 
ing short prosocket ridges and unusually thick ander- 
idia; entire anterior half strongly pustulose. Cardinal 
process having two central ridges and one on each side 
of the central pair. 

Measurements (in mm).— 


dorsal 
valve hinge mid- surface thick- 
length length width width length ness 


USNM 201177 

(loc. 527f) SS) 4.6 6.0 TES 6.5 0.8 
USNM 201124 

(loc. 506q) 5.0 4.6 6.9 6.2 6.0 0.7 


Diagnosis.—Small, moderately convex Reticho- 
netes having five costellae in one mm at the front mar- 
gin. 

Stratigraphic occurrence.—Percha Formation 
(Ready Pay Member; Box Member). 

Localities.—Percha Formation (Ready Pay Mem- 
ber): 3031; Percha Formation (Box Member): 506, b, 
P, q; 5070; 509b; 527f; 3043. 

Types.—Holotype: USNM 201124a. Figured Para- 
types: USNM 201115, 201116a. Unfigured Paratypes: 
USNM 201124b, 201177, 201116b. 

Discussion.—This species differs from Retichonetes 
perchaensis n. sp. in being more transverse, less con- 
vex, and in having five costellae per mm rather than 
three. This species suggests Chonetes scitulus Hall 
(1857), which is common in eastern United States, but 
that species is a much smaller shell, differently orna- 
mented, and is found low in the Middle Devonian. 
Retichonetes nana (Verneuil, 1845) from the Famen- 
nian (D 2/3) of central Russia has the same appearance 
as R. obscurus but it too is larger, more convex and 
with stronger, more distant costellae. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 59 


Retichonetes perchaensis new species 
Plate 11, figures 26-36 


Small, subquadrate in outline, wider than long with 
hinge forming widest part. Sides slightly oblique; an- 
terior margin broadly and fairly strongly rounded. 
Hinge forming the widest part in perfect specimens; 
cardinal extremities forming small rounded ears. In- 
terarea moderately long, apsacline; delthyrium cov- 
ered by a small convex pseudodeltidium. Surface cos- 
tellate; costellae broad, separated by narrower 
interspaces, three to one mm at the front margin of a 
large specimen. 

Ventral valve strongly convex in lateral profile, 
strongly domed in anterior profile; sides steep. Um- 
bonal region strongly swollen and protruding posterior 
to posterior margin. Median region swollen from pos- 
terior to anterior; anterior slope steep and strongly 
rounded. Dorsal valve deeply concave with greatest 
concavity in the median region. 

Interior not known. 

Measurements (in mm).— 


dorsal 
valve hinge mid- surface thick- 
length length width width length ness 


USNM 201122 

(Holotype; 

loc. 527p) 6.5 5.6 8.0 7.8 8.0 0.6 
USNM 201117 

(loc. 5070) 7.0 — 9.6*? 7.3 OF5 — 
USNM 201119 

(loc. 3043) 6.4 — 11.4*? 6.6 8.0? — 
USNM 201120 

(loc. 506) 5.0 — U3 Sys) 7.0? — 


Diagnosis.—Strongly swollen, subquadrate Reti- 
chonetes having three costae to the mm at the anterior 
of an adult. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b; 507m, 0; 509a, e; 527d, p; 3043. 

Types.—Holotype: USNM 201122. Figured Para- 
types: USNM 201117, 201119, 201120, 201123. 

Discussion.—VThis species differs from Retichonetes 
obscurus n. sp. in its more quadrate form, stronger 
convexity and less numerous costellae. 


Genus RHYSSOCHONETES Johnson, 1970 


Rhyssochonetes johnsoni new species 
Plate 11, figures 37-43 


Large for genus, rectangular in outline; wider than 
long, length about 70 percent of width; maximum width 
at about midvalve; sides rounded, anterior margin 
broadly rounded. Hinge wide, as wide as or slightly 


greater than midwidth; cardinal extremities slightly 
obtuse or slightly acute. Interarea moderately long; 
delthyrium partially closed by convex pseudodeltid- 
ium. Posterior margin bearing a few needle-like spines, 
the longest spine at the cardinal extremity attaining a 
length of 3.0 mm ona specimen 6.5 mm long; another 
fairly long spine appearing about half way between 
terminal spine and beak. Other spines not seen. Sur- 
face of both valves finely costellate, the costellae 
overlaid by a layer of raised wavy concentric lines 
usually stronger than costellae and dominating orna- 
ment. 

Ventral valve gently convex in lateral profile with 
maximum curvature in posterior; anterior flattened; 
anterior profile a broad, moderately convex arch with 
long lateral slopes. Dorsal valve moderately concave 
and fitting snugly into ventral valve. 

Ventral valve interior with flat, stout teeth and a 
long slender median septum. Dorsal valve with thick 
socket ridges, deep pit beneath the cardinal process, 
which is small and quadrilobed. 

Measurements (in mm).— 


dorsal =maxi- 
valve mum hinge thick- 
length length width width ness 


USNM 201061b 

(loc. 526b) 8.2 — 11.7 10.0 _- 
USNM 20106Ic 

(loc. 526b) 7.0 — 9.1 9.6 — 
USNM 20106le 

(loc. 526b) 7.0 — 10.3 9.4? _ 
USNM 201061d 

(loc. 526b) 5)-) — 7.6 8.1 _ 
USNM 204953b 

(loc. 527x) 35) — UL) 6.6 — 


Diagnosis.—Large, strongly rugose Rhyssocho- 
netes with length about three-quarters of the width. 
Stratigraphic occurrence.—Onate Formation. 
Localities. —522), n, z; 524p, q, r; 526b; 527x; 529m. 
Types.—Holotype: USNM 201081c. Figured Para- 
types: USNM 204953a, b; 201081a, b, d, e. Unfigured 
Paratypes: USNM 20106la—k, 201081f-j. 
Discussion.—This is a common species in the Onate 
Formation in the Mud Springs Mountains and at Her- 
mosa. The species is the largest of the genus so far 
found. It is much larger than the Tully Rhyssochonetes 
aurora (Hall, 1867), is differently shaped, and has a 
stronger concentric element in its ornament. Rhysso- 
chonetes johnsoni n. sp. is larger than R. bellarugosus 
(Stainbrook, 1943) from the Cedar Valley Formation 
and is much differently ornamented. Stainbrook’s 
species is costate rather than costellate and its con- 
centric markings are coarse and distant. The New 


60 BULLETIN 315 


Mexico species is also much larger and more strongly 
ornamented than R. aurora solox Johnson (1970) from 
the Denay Limestone of Nevada. 

Specimens in the National collections having about 
the same size and type of ornamentation come from 
the Bituminous Shale Member of the Pine Point For- 
mation of Northwest Territories, Canada. Specimens 
referred to R. aurora (Hall) have been described by 
McCammon (1960, p. 47, pl. 8, figs. 1, 2) from the 
Dawson Bay Formation of Manitoba. 


Suborder PRODUCTIDINA Waagen, 1883 
Superfamily STROPHALOSIACEA Schuchert, 1913 
Family STROPHALOSIIDAE Schuchert, 1913 
Subfamily STROPHALOSIINAE Schuchert, 1913 
Genus EOSTROPHALOSIA Stainbrook, 1943 


Eostrophalosia rockfordensis (Hall and Clarke) 
Plate 12, figures 1, 2 


Strophalosia rockfordensis Hall and Clarke, 1893, p. 316, pl. 17A, 
figs. 1-3. 


Through the kindness of Mr. C. O. Levorson, Rice- 
ville, Iowa, we are able to reproduce here a dorsal 
interior of this rare species, which is the type species 
of Eostrophalosia. This specimen shows the deep 
sockets, the bilobed cardinal process and brevisep- 
tum. The specimen is from the Strophonella Zone of 
the Cerro Gordo Member of the Hackberry Formation 
at locality 5S27y. 

Types.—Figured Hypotype: USNM 200838. 


Eostrophalosia inexpectata new species 
Plate 12, figures 3—12 


About normal size for genus, as now understood, 
transversely elliptical in outline with strongly rounded 
sides and rounded anterior margin; hinge narrower than 
maximum width, which is at about midvalve. Interarea 
anacline, moderately long; delthyrium provided with 
wide, flat pseudodeltidium; dorsal valve interarea 
moderately long but shorter than that of ventral valve, 
hypercline. Notothyrium covered by short, low chi- 
lidium. Cicatrix of attachment small. Ventral valve 
strongly and closely nodose, each node bearing a fairly 
long slender spine; dorsal valve marked by rugae and 
nodes bearing long, slender spines. 

Ventral valve fairly strongly and evenly convex, with 
a rounded, swollen umbonal region moderately pro- 
tuberant beyond posterior margin; anterior profile high 
narrow dome with strongly sloping sides. Lateral slopes 
steep, those to the cardinal extremities steep and 
slightly concave. 


Dorsal valve deeply concave, with all slopes steeply 
dipping to midvalve; cardinal extremities slightly con- 
cave at the indistinct ears. Umbo a flattish smooth 
circle. 

Interior not seen. 

Measurements (in mm).—USNM 200926 (Holotype; 
loc. 509r): length 11.7, dorsal valve length 9.4, hinge 
width 8.8, midwidth 14.0, surface length 19.0, thick- 
ness 2.7, and height 5.9. 

Diagnosis.—Eostrophalosia of moderate size hav- 
ing slender spines on both valves, a deeply concave 
dorsal valve, and the length about 80 percent of the 
width. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—S509r. 

Types.—Holotype: USNM 200926. 

Discussion.—The three known species of this genus 
are all unlike the present one. Eostrophalosia inde- 
pendensis (Stainbrook, 1945) is a much smaller species 
with more delicate spines, a wider hinge and a pro- 
portionately larger cicatrix of attachment. Eostropha- 
losia rockfordensis (Hall and Clarke, 1893) is a smaller 
species with rounder outline than that of E. inexpec- 
tata. Eostrophalosia pedderi Crickmay (1963) is a much 
larger species with stronger spines. The Sly Gap form 
is a very rare species, only one good specimen and 
two fragmentary ones having been collected. 


Eostrophalosia species | 


Incomplete specimens from the Onate Formation 
have an apical cicatrix and closely crowded spines. 
The specimens suggest E. pedderi Crickmay (1963) but 
the spines are more closely crowded and the speci- 
mens are larger. 

Localities.—S522g, 1; 523e; 3025a. 

Types.—Mentioned Specimen: USNM 201176. 


DICHACAENIA new genus 
[Gr. dichas, half; akaeinia, thorn] 


Concavo-convex, subrectangular Strophalosiacea 
having a non-spinose dorsal valve and a ventral valve 
with scattered recumbent spines. Ventral valve inte- 
rior with strong teeth. Dorsal valve interior with sock- 
ets for the teeth, a median septum and widely bilobed 
cardinal process. 

Type species.—Dichacaenia perplexa n. sp. 

Diagnosis.—Strophalosiids of medium size, mod- 
erately concavo-convex with sparsely spinose ventral 
valve and dorsal valve without spines. 

Comparison.—A genus close to this one is Eo- 
strophalosia Stainbrook (1943), which differs from Di- 
chacaenia in having a completely spinose dorsal valve 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 61 


whereas that of the new genus is without spines. The 
dorsal valve of Eostrophalosia is like that of Dicha- 
caenia in having a widely bilobed cardinal process that 
differs from most of the early strophalosiids that have 
a cardinal process with laterally compressed lobes. 

The exterior characters of the new genus suggest 
Heteralosia R. H. King (1938) in having a spiny ven- 
tral valve but non-spinose dorsal valve. However, the 
cardinal process of Heteralosia is of the laterally com- 
pressed type and is entirely different from the open, 
bilobed cardinal process of Dichacaenia. 

Dichacaenia differs from Cyphotalosia Carter (1967) 
that has a non-spinose dorsal valve, in its larger size, 
different shape, flat pseudodeltidium and aberrant car- 
dinal process. 

The same features distinguish Dichacaenia from 
Devonalosia Muir-Wood and Cooper (1960) which has 
a narrowly compressed cardinal process like that of 
young Heteralosia. Another distinction that separates 
Devonalosia from the new genus is the large cicatrix 
of attachment, which in Devonalosia may involve 
nearly the entire surface of the ventral valve, as in D. 
wrightorum Muir-Wood and Cooper (1960). The ci- 
catrix of Dichacaenia is small and no specimens were 
found attached; this suggests that the adult lived loose 
on the bottom, torn from its moorings early in life. 


Dichacaenia perplexa new species 
Plate 12, figures 47—55 


Moderately large, attaining a width of almost an inch 
(24 mm); subrectangular in outline with rounded sides 
and a broadly rounded anterior margin. Hinge wide 
but not equalling the midwidth, which is the widest 
part. Cardinal extremities obtusely rounded. Shell 
concavo-convex. Interareas on both valves well de- 
veloped but that on the ventral valve being the longer 
and orthocline to anacline; dorsal interarea anacline. 
Pseudodeltidium and chilidium not developed. Beak 
having a small scar of attachment. Surface of ventral 
valve smooth and spinose, spine bases short and nar- 
row, spines scattered fairly evenly but distantly over 
surface; spines recumbent, slender, and at least four 
mm long. Dorsal valve without spines, but in places 
dimpled corresponding to the spine bases of the op- 
posite valve. 

Ventral valve evenly and gently convex in lateral 
profile; anterior profile moderately strongly domed. 
Umbonal region small, narrowly swollen, only slightly 
protuberant beyond the posterior margin. Postero- 
lateral slopes slightly concave; anterior slope fairly 
steep. 


Dorsal valve moderately concave, most so slightly 
anterior to midvalve; posterolateral extremities mod- 
erately concave. 

Ventral valve interior with stout teeth; dorsal valve 
with bilobed sessile cardinal process. 

Measurements (in mm).— 


dorsal sur- 
valve hinge mid- face thick- 
length length width width length ness height 


USNM 200957a 


(Holotype; 
loc. 5230) Woy MS) as) 253) PPS Se) 
USNM 200957d 
(loc. 5230) 1G! O14 Gis 22 87) eel 9) On 22247, 


Diagnosis.—Large Dichacaenia having low con- 
vexity and fairly evenly scattered, recumbent spines. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—523e; 523), 0. 

Types.—Holotype: USNM 200957a. Figured Para- 
type: USNM 200957b. Unfigured Paratypes: USNM 
200957c-g. 

Comparison.—This species differs from Devonalo- 
sia wrightorum Muir-Wood and Cooper (1960), which 
is of about the same size, in the smaller cicatrix and 
the different spination. It differs from Eostrophalosia 
rockfordensis (Hall and Clarke, 1893) in its larger size 
and more spreading form. From Eostrophalosia ped- 
deri Crickmay (1963), this Onate form differs in its 
flatter and wider shell and more scattered spines. 

Dichacaenia perplexa n. sp. differs from D. umbo- 
nata n. sp. in its flatter umbonal region and less con- 
vex ventral valve. 


Dichacaenia umbonata new species 
Plate 12, figures 41-46 


Fairly large, elliptical in outline, wider than long; 
sides rounded; anterior margin broadly rounded. Hinge 
narrower than the maximum width, which is at mid- 
valve. Cicatrix of attachment small. Median region 
swollen; umbonal region only slightly protuberant, but 
broad and swollen. Posterolateral slopes short; car- 
dinal extremities somewhat flattened. Surface covered 
by short, distantly scattered, recumbent spines. 

Dorsal valve exterior without spines and flatly la- 
mellose. Dorsal valve interior with long slender brevi- 
septum reaching to midvalve, expanding posteriorly 
but not reaching the cardinal process; socket narrow, 
moderately deep. Cardinal process small, widely bi- 
lobed. 


62 BULLETIN 315 


Measurements (in mm).— 


Measurements (in mm).— 


dorsal maxi- 
valve hinge mum_ thick- surface 
length length width width ness length 


dorsal maxi- sur- 
valve hinge mum _ face _thick- 
length length width width length ness height 


USNM 201012a 
(loc. 523s) 17.8 a 19.5? DDeiih — 23.0 
USNM 201012b 13.3 — 15.0? 73) _— 20.0 


Diagnosis.—Dichacaenia with swollen ventral um- 
bonal region. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —522j; 523b, i, s, t, u, Z; 526c, 529}. 

Types.—Holotype: USNM 201012a. Figured Para- 
types: USNM 201012b, d; 201013; 204954. Unfigured 
Paratypes: USNM 201012c, eH. 

Comparison.—This species differs from D. perplexa 
n. sp. in having a larger size, more inflated ventral 
valve, and wider, more swollen umbonal region. The 
material on which this species is based is scanty and 
preservation is not the best. Only two specimens pre- 
serve the exterior of the dorsal valve. 


Superfamily PRODUCTACEA Gray, 1840 
Family PRODUCTELLIDAE Schuchert, 1929 
Subfamily PRODUCTELLINAE Schuchert, 1929 
Genus PRODUCTELLA Hall, 1867 


Productella rugatula Stainbrook 
Plate 12, figures 13-20 


Productella rugatula Stainbrook, 1945, p. 36, pl. 3, figs. 48, 49; 1948, 
p. 772, pl. 1, figs. 20-22. 


This is a small productellid having a subcircular out- 
line, with rounded sides and anterior; hinge narrower 
than maximum width. The interarea is short and nar- 
row. The cardinal extremities form small, obtuse ears. 
The ventral umbonal region is narrowed and swollen, 
with irregular concentric wrinkles. Concentric wrin- 
kles appear infrequently on the shell body but are 
prominent along the posterior margin. The spines of 
the ventral valve are scattered, variable in number and 
strength. All parts of the ventral valve have spines but 
there seems to be little system to their arrangement. 
They are present on the ears, lateral and anterior 
slopes, median and umbonal regions; one specimen 
may be more abundantly adorned in one part than 
another. The spines have a length of seven mm on a 
shell only eight mm long. The dorsal valve closely 
follows the concavity of the ventral valve and is dim- 
pled in places corresponding to the position of spines 
on the opposite valve. 


USNM 201175a 
(loc. 509n) 1120) 928) 102) 127 Ste 16:05: Oma 
USNM 201175b 
(loc. 509n) 11.4 9.6 SiS) eile S S172 ON 2 See 


Diagnosis. —Small Productella having variable spine 
adornment. 

Stratigraphic occurrence.—Sly Gap Formation; In- 
dependence Formation. 

Localities. —S06c; 509h, j, 1, n, s; 521c, d, e, k, m, 
p, 1, S, V, W; 522k; 524m, w; 526n. 

Types.—Figured Hypotypes: USNM 201118, 201120. 

Discussion.—This is a rare and unsatisfactory 
species because of its invariably poor preservation. 
Few of the specimens preserve spines and most of 
those having both valves in contact are broken around 
the margins. Furthermore, the shells are variable, not 
only in the spine arrangement but also in presence of 
concentric wrinkles. A few specimens have the um- 
bonal region fairly crowded with spine bases. 

The holotype was described by Stainbrook from the 
Independence Shale at Brandon, Iowa. 


Productella species | 
Plate 12, figures 29-40 


Small to medium for genus, subquadrate in outline, 
wider than long; hinge narrower than midwidth; car- 
dinal extremities obtuse; interarea short and narrow. 
Ventral valve strongly convex especially in the um- 
bonal region, which is narrowly curved and strongly 
protuberant beyond the posterior margin; anterior pro- 
file high dome with nearly straight, precipitous sides. 
Dorsal valve deeply concave, most so at midvalve. 
Ventral valve surface marked by concentric undula- 
tions on the umbonal and posterior regions, and by 
long slender spines scattered distantly on surface, and 
without elevated bases. Anterior marked by occasion- 
al short costae, more numerous on lateral slopes. Dor- 
sal valve without spines but with concentric undula- 
tions on posterior margins and around umbonal pit. 

Interior of dorsal valve with long slender brevisep- 
tum. 


DEVONIAN BRACHIOPODS OF NEw Mexico: COOPER AND DuTRO 63 


Measurements (in mm).— 


dorsal maxi- — sur- 
valve hinge mum _ face thick- 
length length width width length ness height 


USNM 201011b 

(loc. 521h) ASO 2 7 62799240) 3) 1825 
USNM 20101 1a 

(loc. 521h) [3X0 ES S23 14723. Ol — 7-0) 
USNM 203563a 

(loc. 527z) 12.4 — EA S208) 210) —=s 720 


Diagnosis.—Small to medium-sized Productella with 
distantly scattered spines and length about 85 percent 
of width. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation. 

Localities.—5S09t, u; 521h; 524e; 527z; 3054g, q. 

Types.—Figured Specimens: USNM 201011a, b; 
203563a-—c. 

Discussion.—These specimens cannot be identified 
with any described species but they are too poor to 
use as the basis for a new species. They are much 
larger than P. rugatula Stainbrook (1945). The illus- 
trated specimens suggest P. thomasi Stainbrook (1945) 
but are larger than that species and have a greater 
development of costae anterior to the spines. 


Productella? species 2 
Plate 11, figures 44—46 


Large for genus, subquadrate in outline and with 
subparallel sides and rounded anterior margin. Strong- 
ly convex, the greatest convexity at midvalve; anterior 
and posterior slopes steep; umbonal region swollen 
and strongly protuberant beyond the posterior margin. 
Interarea short; hinge narrower than midwidth. Sur- 
face marked by numerous slender spines, usually with 
small bases, but anteriorly the spines have interrupted 
elongate ridges on their anterior sides that give the 
impression of anterior costellation. 

Measurements (in mm).—USNM 201143a: length 
26.0, dorsal valve length 22.0, hinge width 14.0, max- 
imum width 26.6*, surface measure 43.0, thickness un- 
known, height 14.0. 

Stratigraphic occurrence.—Contadero Formation. 

Localities. —Contadero: 509u; 523n. Probably Con- 
tadero: 5280. 

Types.—Figured Specimen: USNM 201143a. 

Discussion.—Three lots of large Productella are so 
poorly preserved that they cannot be satisfactorily de- 
scribed. The measured and illustrated specimen is the 
best preserved, but it is damaged on one side. We have 
been unable to make a satisfactory comparison of this 
species with any form described by Hall from the Up- 
per Devonian of New York. 


Productella species 3 
Plate 13, figure 1 


A single specimen of Productella was collected from 
the Thoroughgood Formation (loc. 522). It is so poorly 
preserved that nothing but its outline is definite. 

Types.—Figured Specimen: USNM 201086. 


Subfamily CHONOPECTINAE Muir-Wood and 
Cooper, 1960 


Genus ACANTHATIA Muir-Wood and Cooper, 1960 


Acanthatia nupera (Stainbrook) 


Heteralosia nupera Stainbrook, 1947, p. 309, pl. 46, figs. 16, 17. 
Acanthatia nupera (Stainbrook) Muir-Wood and Cooper, 1960, p. 
158, pl. 38, figs. 1-14. 


This species was inadequately figured by Stainbrook 
but later was described in detail by Muir-Wood and 
Cooper (1960). Its cardinal process is productellid 
rather than strophalosiid and the genus is now placed 
in the Productellidae. It is not a common fossil in the 
Box Member except at locality 506b (=507o). 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —506, a, b, p, q; 507a, h, m, 0, r; 509a, 
b; 527e, f, p; USGS (Kirk) 508; 3026, a; 3029, 3034 = 
507h; 3035; 3042. 

Types.—Holotype: M.A.S. 1598a (=USNM 135240). 
Figured Paratype: M.A.S. 1598b (=USNM 135241). 
Figured Hypotypes: USNM 123437, 123438a-e, g, 
123921a, 123922. All types figured by Stainbrook (1947). 


Family LEIOPRODUCTIDAE Muir-Wood and 
Cooper, 1960 


Subfamily LEIOPRODUCTINAE Muir-Wood and 
Cooper, 1960 


Genus LEIOPRODUCTUS Muir-Wood and 
Cooper, 1960 


Leioproductus plicatus (Kindle) 


Productella coloradensis var. plicatus Kindle, 1909, p. 18, pl. 4, 
figs. 9-11, 712. 

Leioproductus plicatus (Kindle) Stainbrook, 1947, p. 308, pl. 46, 
figs. 21-23; Muir-Wood and Cooper, 1960, p. 168, pl. 40, figs. 1- 


128 


This is a small, rather smooth productid having a 
median plica or median row of spines; it is common 
at many localities. It is difficult to distinguish L. pli- 
catus from L. coloradensis (Kindle, 1909), which is 
very rare in New Mexico. Evidently, Stainbrook se- 
lected this “‘variety”’ as type species for Leioproduc- 
tus because of its abundance as compared to the pau- 
city of numbers of L. coloradensis. 


64 BULLETIN 315 


Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —506, b, p, q; 507a, b, c, d, g, h, k 
hy Oy Gly tts Wo WS SWS fly le SIG Ge ii, (5 (OR Sil) 
3029; 3034; 3035; 3042, s; USGS (Kirk) 508; 512. 


, l,m 
26, a; 


Leioproductus defessa (Stainbrook) 
Avonia defessa Stainbrook, 1947, p. 309, pl. 46, figs. 6, 7. 


The single specimen described and figured by Stain- 
brook is the only one of this species that we have seen. 
Although we have extensive collections from the type 
section of the Percha Formation (Box Member, local- 
ity 507h), from which A. defessa came, no other spec- 
imen like A. defessa has appeared. The unusual fea- 
ture of A. defessa is the development of costae anterior 
to the spine bases which give the appearance of cos- 
tae, the development of which on the anterior slope 
of a productid is a distinguishing character of Avonia 
Thomas (1914). Stainbrook’s specimen has umbonal 
undulations which are not usually present on Avonia. 
The entire aspect of Stainbrook’s specimen is that of 
Leioproductus Muir-Wood and Cooper (1960), but its 
spines are finer than those seen on most specimens of 
Leioproductus. 

Types.—Holotype: M.A.S. 1612 (=USNM 135255). 


Leioproductus coloradensis (Kindle) 


Productella semiglobosa Girty, 1900, p. 42, pl. 3, figs. 7-10. 

Productella coloradensis Kindle, 1909, p. 17, pl. 4, figs. 2-8. 

Leioproductus coloradensis (Kindle) Stainbrook, 1947, p. 308, pl. 
46, figs. 18-20. 


This is a much less common species in New Mexico 
than Leioproductus plicatus (Kindle, 1909) and may 
be identified by its lack of either a median plica or 
vertical row of spines on the ventral valve. Stain- 
brook’s figured specimens were not received by the 
U.S. National Museum of Natural History in the gift 
of his Percha collection. 

Stratigraphic occurrence.—Ouray Formation (in 
Colorado); Percha Formation (Box Member). 

Localities.—Rockwood, Colorado; 507h. 

Types.—Holotype: USNM 62009c. Paratypes: 
USNM 62009a, b, d, e. Figured Hypotypes: USNM 
155811-3 (=M.A.S. 1606). Types figured by Stain- 
brook (1947). 


Leioproductus pauperculus new species 
Plate 12, figures 21-28; plate 13, figures 2-9 

Small for genus, quadrate in outline, slightly wider 
than long; sides slightly oblique to rounded; anterior 
margin broadly rounded; hinge wide, slightly narrower 
than midwidth, in well-preserved specimens. No in- 
terareas. Surface smooth except for spines and con- 
centric wrinkles. Spines fairly long, scattered, leaving 
rounded bases, when broken, that seldom have ante- 
rior ridges. A few spines located on ears and posterior 
margin, a few on the lateral slopes and a few on the 
anterior slope, in some specimens forming an indis- 
tinct row. Concentric undulations generally fairly wide 
and located chiefly in umbonal region. 

Ventral valve with strongly swollen umbo protrud- 
ing well beyond hinge line. Lateral profile strongly 
convex especially in umbonal region where it forms a 
semicircle. Anterior profile a strongly convex dome 
with very steeply sloping sides. Anterior slope long 
and moderately steep. Ears small and rounded. 

Dorsal valve smooth, with few concentric undula- 
tions best developed in the vicinity of the hinge and 
ears. Valve deepest posteromedially, and all slopes 
dipping steeply to the middle; umbonal region a deep 
pit. 

Interior not known. 

Measurements (in mm).— 


dorsal 
valve hinge mid- thick- surface 
length length width width ness length height 


USNM 201111 

(loc. 506p) 10.4 — 9.4* 11.3 — 17 OFS 72032; 
USNM 201110a 

(loc. 506) 10.2 — 10.5 11.8 — 17.5 6.8 
USNM 201112 

(loc. 509a) les; sy 19 1320) 1227 18.5 6.0 


Diagnosis.—Small Leioproductus, with length 0.9 
the width, having umbonal region of ventral valve with 
concentric undulations and scattered distant spines on 
the surface. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —506, b, p, q; 507b, c, h, k, m, n; 509, 
a, b, c; 527d, e, p; 3026; 3042, s. 

Types.—Holotype: USNM 201112a. Figured Para- 
types: USNM 201110a, 201111. Unfigured Paratypes: 
USNM 201110b, 201112b—d. 

Discussion.—This species can be recognized by its 
small size and scattered spines. It differs from both L. 
coloradensis (Kindle, 1909) and L. plicatus (Kindle, 
1909) in its size, nearly equal length and width, more 
numerous spines and less strong concentric undula- 
tions. The species is rare. 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DuTRO 65 


Leioproductus varispinosus (Stainbrook) 


Productella coloradensis var. plicatus Kindle, 1909 (part), pl. 4, fig. 
13 (not figs. 9, 10-1 1a). 

Bispinoproductus varispinosus Stainbrook, 1947, p. 311, pl. 46, figs. 
30-33. 

Leioproductus varispinosus (Stainbrook) Muir-Wood and Cooper, 
1960, p. 168, pl. 40, figs. 14-25. 


Muir-Wood and Cooper (1960, p. 168) put this species 
in Leioproductus Stainbrook (1947) when they sup- 
pressed Bispinoproductus Stainbrook (1947), which is 
regarded as a synonym of Leioproductus. This is a 
large productid, easily distinguished from the other 
species by its more numerous spines, larger size and 
more strongly geniculated profile. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, a, b, p, q; 507a, b, c, f, h, m, n, 
0, p; 509, a, b, e; 527a, d, e, f, g, 1, p; 3026, a; 3029: 
303 1b; 3034; 3042, s. 

Types.—Holotype: USNM 155786. Paratype: USNM 
155787. Primary types figured by Stainbrook (1947). 
Figured Hypotypes: USNM 123947a, b; 123945a, b; 
123948. All types in U.S. National Museum of Natural 
History. 


Genus GALEATELLA Muir-Wood and Cooper, 1960 
Galeatella galeata Muir-Wood and Cooper 


Galeatella galeata Muir-Wood and Cooper, 1960, p. 174, pl. 41, 
figs. 15-24. 


This is a fairly large, smooth leioproductid having a 
broad sulcus at the anterior. It is an uncommon species. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, a, b, p, q; 507a, b, h, k; 509b, c; 
527d; USGS (Kirk) 461; 3026, a; 3034; 3042. 

Types.—Holotype: USNM 123447. Figured Para- 
types: USNM 123448a, b; 123449a, b. Types figured 
in Muir-Wood and Cooper (1960). 


Genus PLANOPRODUCTUS Stainbrook, 1947 


Planoproductus hillsboroensis (Kindle) 


Productella hillsboroensis Kindle, 1909, p. 19, pl. 5, fig. 6. 

Planoproductus hillsboroensis (Kindle) Stainbrook, 1947, p. 310, pl. 
46, figs. 27-29; Muir-Wood and Cooper, 1960, p. 176, pl. 42, figs. 
1-15. 


This is the largest of the Devonian productids so far 
described in North America and can be readily rec- 
ognized even from fragments. It is an uncommon 
species. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 


Localities. —506, a, b, q; 507a, g, h, m, n, 0; 527d, 
f, p; 3026; 3029; 3025; 3042, s. 

Types.—Holotype: USNM 62017. Hypotypes fig- 
ured by Stainbrook (1947) and Muir-Wood and Cooper 
(1960): USNM_ 155793-4 (=M.A.S. 1590); USNM 
123440; 123441; 123443; 123444a; 123942; 123943a. 


Subfamily DEVONOPRODUCTINAE Muir-Wood and 
Cooper, 1960 


Genus DEVONOPRODUCTUS Stainbrook, 1943 


Devonoproductus intermedius new species 
Plate 13, figures 36-56 


Medium size for genus, squarish in outline with 
length and width nearly equal; sides gently rounded to 
subparallel; anterior margin strongly rounded; cardinal 
extremities forming small obtuse ears. Interarea short 
and strongly curved; beak small, incurved and mod- 
erately protuberant beyond the posterior margin. Ven- 
tral valve surface finely costellate and spinose; cos- 
tellae low, crowded, wider than interspaces and 
numbering four to one mm at the anterior margin of 
large adult; spines small, distant, scattered on shell 
body, sparse on ears. Dorsal valve lamellose, lamellae 
closely crowded, about three to one mm and about 26 
in shell 12 mm long; lamellae crowded at anterior. 

Ventral valve moderately convex in lateral profile, 
most so at umbo, which is narrowly rounded; anterior 
profile fairly strongly convex and having steep lateral 
slopes. Umbonal region narrowly rounded and swol- 
len, the swelling extending to midvalve and then de- 
creasing anteriorly. Posterolateral slopes steeply de- 
scending to the slightly rounded ears. 

Dorsal valve moderately concave, most so at about 
midvalve; lateral and anterior slopes descending mod- 
erately steeply toward valve center. Umbonal region 
slightly concave; ears flattened. 

Ventral valve interior not seen. Dorsal valve interior 
having a long, slender breviseptum and small cardinal 
process with myophore lobes separate. Posterolateral, 
lateral and anterior slopes marked by nodes or short 
endospines, the latter concentrated on the anterior 
slope. 

Measurements (in mm).— 


dorsal 
valve hinge maximum surface thick- 
length length width width length ness 


USNM 201027a 


(Holotype; 

loc. 509n) 14.1 12.1 12.3 14.6 PN) GLa 
USNM 201027b 

(loc. 509n) 12.0 10.3 10.4 12.6 16.0 
USNM 201027c ='11.1 9:9 9.4 11.6 14.0 3.2 
USNM 201027d_ —s'11..2 10.0 9.1 11.2 160 3k2 


66 BULLETIN 315 


dorsal 
valve hinge maximum surface thick- 
length length width width length ness 
USNM 20105la 
(loc. 506j) 12.5 11.0 10.0 12.9 KJe5) 3-3 
USNM 201045a 
(loc. 521le) 11.0 _ 9.6 11.3 16.0 — 
USNM 201174a 
(loc. 521k) 10.8 10.0 8.5 11.4 14.0 2.9 


Diagnosis.—Devonoproductus of medium size hav- 
ing numerous overlapping lamellae on the dorsal valve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S06c, j; 509n, q, r, s, z; 521d, e, k, m, 
Pp, q, r, u, w; 522k; 524m, n; 526g, k, n; USGS (Kirk) 
534; 538. 

Types.—Holotype: USNM 201027a. Figured Para- 
types: USNM 201027d; 201045a; 20105la; 201052a; 
201058. Unfigured Paratypes: USNM 201027b, c; 
201045b; 201051b; 201052b. 

Discussion.—This species is intermediate between 
the Independence Formation species, D. vulgaris 
Stainbrook (1945) and D. walcotti (Fenton and Fen- 
ton, 1924) from the Cerro Gordo Member of the Hack- 
berry Formation. Devonoproductus intermedius dif- 
fers from the first in its larger size, less deeply concave 
dorsal valve, less elongated and elevated umbonal re- 
gion, and less pronounced ears. It differs from D. wal- 
cotti in being smaller, less inflated, having a less deep 
dorsal valve, and much finer and more numerous over- 
lapping lamellae on the dorsal valve. Devonoproduc- 
tus intermedius differs from D. hallianus (Walcott, 
1884) from the Eureka District, Nevada, in its smaller 
size, less depth, less protuberant and narrower umbo 
of the ventral valve and more crowded dorsal lamellae. 

Devonoproductus intermedius is a fairly common 
and distinctive fossil in the Sly Gap Formation and the 
genus is one of the more characteristic of Frasnian 
fossils. 


Devonoproductus vulgaris Stainbrook 


Devonoproductus vulgaris Stainbrook, 1945, p. 37, pl. 3, figs. 44, 
51-54. 


Four specimens taken from the Sacramento Moun- 

tains conform to Stainbrook’s species because they 
have high, narrow umbonal regions and a strong, al- 
most subcarinate anterior profile. Unfortunately, no 
dorsal valve is available to check its degree of con- 
cavity and lamellation. 
Small Devonoproductus with strongly 
swollen ventral valve, swollen umbonal region, deeply 
concave dorsal valve and variable lamellation on the 
dorsal valve. 


Diagnosis. 


Stratigraphic occurrence.—Sly Gap Formation. 
Localities. —5S06c; 509v, w. 

Types.—Stainbrook Hypotype: USNM 135282. 
Discussion.—Stainbrook’s specimen came from lo- 


cality S06c. 


Family OVERTONIIDAE Muir-Wood and 
Cooper, 1960 


Genus LAMINATIA Muir-Wood and Cooper, 1960 


Laminatia laminata (Kindle) 


Productella laminatus Kindle, 1909, p. 18, pl. 4, figs. 13, 14. 

Echinoconchus laminatus (Kindle) Stainbrook, 1947, p. 314, pl. 46, 
figs. 3-5. 

Laminatia laminata (Kindle) Muir-Wood and Cooper, 1960, p. 189, 
vl. 41, figs. 1-14. 


This is a small member of the Overtoniidae having 
strong lamellae bearing spines and thus shows a ho- 
meomorphic development toward Echinoconchus S. 
Weller (1914). It is generally a rare species. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b, p, q; 507a, b, h, k, m, q, z; 509, 
a, b, e; 527a, d, e, p; USGS 508; 3026a; 3042, s. 

Types.—Holotype: USGS 62014a. Paratype: USNM 
62014b, figured in Kindle (1909). Hypotypes figured in 
Stainbrook (1947): M.A.S. 1588a, b (=USNM 135247a, 
b). Hypotypes figured in Muir-Wood and Cooper 
(1960): USNM 12395Sa; 123956a, c; 123957; 124127. 


Genus SENTOSIA Muir-Wood and Cooper, 1960 


Sentosia praecedens (Stainbrook) 


Productella spinigera (part) Kindle, 1909, pl. 5, figs. 2, 2a, b. 

Pustula praecedens Stainbrook, 1947, p. 312, pl. 46, figs. 24-26. 

Sentosia praecedens (Stainbrook) Muir-Wood and Cooper, 1960, p. 
196. 


This species is distinguished from S. praecursor 
(Stainbrook, 1947) by its greater convexity, fewer 
spines, and obscure concentric bands. It is a com- 
moner species than §. praecursor but is rather rare 
nevertheless. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b, p, q; 507d, h, m, n, 0; 509a, b, 
e; 527, d, e, p; 3026; 3029; 3042, s. USGS 461, 510. 

Types.—Holotype: M.A.S. 1610a (=USNM 135242). 
Figured Paratype of Stainbrook (1947): M.A.S. 1610b 
(=USNM 123243). Paratype of Productella spinigera 
Kindle (1909): USNM 62015d. 


DEVONIAN BRACHIOPODS OF NEW MExICo: COOPER AND DuTRO 67 


Sentosia praecursor (Stainbrook) 


Productella spinigera Kindle (part), 1909, pl. 5, fig. 4. 

Krotovia praecursor Stainbrook, 1947, p. 313, pl. 46, figs. 11-15. 

Sentosia praecursor (Stainbrook) Muir-Wood and Cooper, 1960, p. 
196, pl. 43, figs. 8-22, text-fig. 7c. 


Muir-Wood and Cooper proposed this species for 


the type of their genus Sentosia, a very distinctive 
form because of its fine mat of spines and broadly 
flattened anterior. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b, p, q; 5070; 509a, b, e; 527c, d, 
f; USGS (Kirk) 461, 510; 3026a, 3042s. 

Types.—Holotype: M.A.S. 1609A = 155790. Fig- 
ured Paratype: M.A.S. 1609 = 155791, primary types 
figured in Stainbrook (1947). Unfigured Paratype: 
USNM 155972. Figured Paratype of Productella spi- 
nigera Kindle (1909) = USNM 62015Sc. Hypotypes fig- 
ured by Muir-Wood and Cooper (1960): USNM 
123959a, b; 123968a, b; 124174. 


Sentosia spinigera (Kindle) 


Productella spinigera Kindle, 1909, p. 19, pl. 5, fig. 1. 
Krotovia spinigera (Kindle) Stainbrook, 1947, p. 313, pl. 46, figs. 
1, la. 


This species is distinguished from the other two 
members of this genus occurring in the Percha For- 
mation (Box Member) by its small size, squarish out- 
line, and strong convexity. It is a very rare species. 

Localities.—506, b, p; 507a, c, h, 0; 509b; 527d, f; 
3026a; USGS 461. 

Types.—Holotype: USNM 62015b. Figured Para- 
types: USNM 62015a, c, d, primary types figured in 
Kindle (1909). Figured Hypotype of Stainbrook (1947): 
M.A.S. 1599 (=USNM 155778). 


Order PENTAMERIDA Schuchert and Cooper, 1931 


Suborder PENTAMERIDINA Schuchert and 
Cooper, 1931 


Superfamily PENTAMERACEA M’Coy, 1844 
Family PENTAMERIDAE M’Coy, 1844 
Subfamily GYPIDULINAE Schuchert, 1929 
Genus GYPIDULA Hall, 1867 


Gypidula bowsheri new species 
Plate 13, figures 26-35; plate 14, figures 1-10 


Small, longer than wide and pentagonal in outline; 
anterior commissure moderately sulcate; surface 


smooth in the posterior half but costate in the anterior 


parts, two or three costae on the fold and one or two 
in the sulcus. Flanks marked by two or three costae. 


Ventral valve moderately convex in lateral profile; 
umbonal region narrowly rounded; anterior profile 
strongly rounded, sides precipitous. Fold originating 
at about midvalve or slightly anterior thereto, low and 
poorly defined, gently rounded transversely. Median 
and umbonal regions strongly swollen; beak moder- 
ately protuberant. Beak strongly incurved, closely ap- 
proaching that of the dorsal valve. 

Dorsal valve gently convex in the posterior half in 
lateral profile but flattened in anterior half; anterior 
profile broadly and slightly convex. Sulcus originating 
abruptly just anterior to midvalve, deepening rapidly 
and forming short, serrate tongue. Flanks flattened. 

Measurements (in mm).— 


dorsal 
valve 
length length width thickness 
USNM 200823 
(loc. 509}) 10.3 8.2 10.0 sl 
USNM 200824 
(Holotype; loc. 526f) 9.3 8.0 9.6 6.5 


Diagnosis.—Small Gypidula, elongate, anteriorly 
costate, one or two costae in the sulcus. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, 1, j; 509j, n, q, s, v; 521e; 524n; 
526f; USGS (Kirk) 541; 3045b. 

Types.—Holotype: USNM 200824. Figured Para- 
type: USNM 200823. 

Comparison.—Gypidula bowsheri n. sp. at once 
suggests another small Gypidula of the Late Devoni- 
an, G. munda (Calvin, 1878), known from a few spec- 
imens only. Comparison of Stainbrook’s figure of the 
type specimen of G. munda indicates that it is a small- 
er and wider species. It is possible that the type spec- 
imen is immature and that the adult would be consid- 
erably larger. Although this is speculation, specimens 
from the Mount Hawk Formation at Winnifred Pass, 
Alberta, received through Dr. D. J. McLaren, are larg- 
er but similar to G. munda in most other characters. 
Gypidula lotis (Walcott, 1884) is also similar if not the 
same as G. munda. It is much wider than G. bowsheri 
and is more strongly costate. 


Gypidula cornuta Fenton and Fenton 
Plate 14, figures 66-73 


Gypidula comis Hall and Clarke (not Owen, 1852), 1894, pl. 72, figs. 
15-17, 21. 

Gypidula cornuta Fenton and Fenton, 1924, p. 121, pl. 25, figs. 26— 
31; Belanski, 1928, p. 14, pl. 1, figs. 12-24, pl. 2, fig. 14. 


This species has hitherto been known only from the 
basal ‘‘zonule’’ of the Cerro Gordo Member of the 
Hackberry Formation where it is rather rare. Although 


68 BULLETIN 315 


the Sly Gap fauna shares elements with the Cerro Gor- 
do Member, Gypidula cornuta was reported by Ste- 
venson (1945, p. 239). The Stevenson specimen is now 
in the U.S. National Museum of Natural History and 
is better referred to Gypidula bowsheri n. sp. 

Gypidula cornuta can now be recorded from the In- 
dependence Formation of Iowa, a formation nearly 
contemporaneous with the Sly Gap Formation. A 
specimen (USNM 200821) presented to the U.S. Na- 
tional Museum of Natural History by Dr. M. A. Stain- 
brook, is well within the range of variation of G. cor- 
nuta. The specimen has an elliptical dorsal valve with 
deep sulcus occupied by a single costa. The fold of 
the ventral valve is low and narrow, defined only in 
the front half of the valve and marked by a slight me- 
dian depression corresponding to the costa in the sul- 
cus. The beaks of both valves are closely appressed 
and the flanks have one obscure costa. 

Measurements (in mm).—USNM 200821: length 
20.8, dorsal valve length 16.5, width 21.0, thickness 
15.8. 

Stratigraphic occurrence. 
tion. 

Locality.—S526t. 

Types.—Figured Hypotype: USNM 200821. 


Independence Forma- 


Gypidula mimica new species 
Plate 13, figures 18—25 


About medium size for genus, length and width 
nearly equal; outline roundly pentagonal; anterior 
commissure strongly sulcate. Surface of posterior half 
smooth, that of anterior half costate; four costae in 
fold and three in sulcus; flanks marked by three costae 
on right side and four on left when viewed from ventral 
side. 

Ventral valve moderately convex in lateral profile, 
most convex in umbonal region; anterior profile 
strongly and roundly convex; umbonal and median re- 
gion strongly swollen; fold originating 23 mm anterior 
to the beak (measured along the surface); fold occu- 
pying slightly less than half valve width, low and flat- 
tened, forming a moderate recess at anterior margin; 
flanks moderately swollen and descending steeply to 
margins. Ventral valve protruding 5.5 mm posterior to 
beak of the dorsal valve. 

Dorsal valve flatly convex in lateral profile, most 
curved anteriorly; anterior profile broadly and gently 
convex; sulcus originating abruptly about 10 mm an- 
terior to beak and producing short truncated tongue; 
flanks bounding sulcus flattened. 

Measurements (in mm).—USNM 200820 (Holo- 
type): length 21.0, dorsal valve length 16.4, width 22.0, 
thickness 14.0. 


Diagnosis.—Gypidula with posterior half smooth, 
anterior part costate with strongly protuberant ventral 
valve. 

Stratigraphic occurrence.—Sly Gap Formation (up- 
per part). 

Locality.—506c. 

Types.—Holotype: USNM 200820. 

Comparison.—This species is more like the Cedar 
Valley Gypidula occidentalis (Hall, 1858) than it is like 
G. cornuta Fenton and Fenton (1924) and forms re- 
lated to that species. Gypidula mimica differs from G. 
occidentalis in having a less convex dorsal valve, less 
strongly costate margins, less incurved ventral beak, 
stronger tongue on the dorsal valve, and more strongly 
rounded lateral profile. Gypidula comis (Owen, 1852) 
is another similar Cedar Valley species but it is larger, 
with narrower, more protuberant beak, stronger cos- 
tation and deeper sulcus than G. mimica. Gypidula 
mimica differs from G. cornuta Fenton and Fenton in 
being more costate anteriorly, less protuberant pos- 
teriorly, in having a shallower sulcus and less approx- 
imate beaks. 


Gypidula stainbrooki new species 
Plate 14, figures 11-18 


About medium size for genus, angularly pentagonal 
in outline; length and width nearly equal. Posterior 
quarter of exterior smooth, anterior three-quarters 
strongly costate. Costae subangular and prominent; 
two on the fold separated by a deep furrow; one in the 
sulcus and two, plus an incipient third costa, on each 
flank. 

Ventral valve strongly convex in lateral profile, nar- 
rowly domed and having precipitous sides in anterior 
profile. Fold originating eight mm from beak (mea- 
sured along surface); fold visible on the umbonal re- 
gion, strengthened and elevated slightly anteriorly but 
never strongly elevated above the flanks; median re- 
gion strongly inflated; flanks swollen and steep. 

Dorsal valve gently convex in posterior third but 
flattened or slightly concave anteriorly; anterior profile 
nearly flat; sulcus originating seven mm anterior to 
beak, widening abruptly, deepening anteriorly to pro- 
duce a fairly long, serrate tongue. Flanks flattened or 
slightly concave. 

Measurements (in mm).—USNM 200822 (Holo- 
type): length 17.1, dorsal valve length 14.3, width 18.3, 
thickness 12.8. 

Diagnosis. —Gypidula resembling G. cornuta Fen- 
ton and Fenton (1924) but costae extending for three- 
fourths the valve length. 

Stratigraphic occurrence. 
tion. 


Independence Forma- 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 69 


Locality.—526t. 

Types.—Holotype: USNM 200822. 

Comparison.—As mentioned above, this specimen 
is suggestive of G. cornuta Fenton and Fenton (1924) 
but differs in the very strong costation, and in less 
protuberance of the ventral beak beyond the dorsal 
valve posterior margin. 

Discussion.—This species was taken by Stainbrook 
from the Brandon, Iowa quarry from which the spec- 
imen of G. cornuta, discussed above, was found. It 
might be argued that G. stainbrooki n. sp. is a variant 
of G. cornuta but none of the specimens of that species 
in the National Collection exhibits such strong costa- 
tion and none with strong costation is mentioned by 
Fenton and Fenton (1924) in their discussion of G. 
cornuta. 


Gypidula subcarinata new species 
Plate 13, figures 10-17 


About medium size for genus, rhomboidal in out- 
line, sides narrowly rounded; anterior margin angular; 
apical angle 91°. Ventral valve posterior moderately 
protuberant. Anterior commissure narrowly sulcate. 
Beak incurved and closely approaching that of dorsal 
valve. Posterior three-quarters smooth, anterior quar- 
ter marked by narrow costae, five or six on the flanks, 
four on the fold, and three in the sulcus. 

Ventral valve moderately convex in lateral profile, 
greatest convexity in umbonal region. Anterior profile 
subcarinate, lateral slopes gently convex and very 
steep. Fold originating six mm anterior to the beak as 
measured along the surface, low initially but fairly 
prominent anteriorly. Fold narrowly convex and form- 
ing a deep re-entrant. 

Dorsal valve moderately convex in lateral profile but 
subgeniculate somewhat posterior to midvalve where 
the sulcus forms; anterior profile gently and broadly 
convex. Sulcus deep and narrow in anterior half of 
valve and producing a long narrow tongue. Flanks 
bounding sulcus gently convex. 

Measurements (in mm).—USNM 200850 (Holo- 
type): length 15.2, dorsal valve length 14.1, width 17.8, 
thickness 12.1, apical angle 91°. 

Diagnosis.—Gypidula of medium size, rhomboidal 
outline, narrow fold and numerous narrow costae. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—52\k. 

Types.—Holotype: USNM 200850. 

Comparison.—This species is smaller and has a less 
extended ventral umbonal region, as well as more nu- 
merous costae, than G. cornuta Fenton and Fenton 
(1924). It is smaller, has more narrowly rounded lateral 


extremities, narrower fold and deeper sulcus, and an 
entirely different shape from G. mimica n. sp. 


Genus PENTAMERELLA Hall, 1867 


Pentamerella species 
Plate 39, figure 29 


A single ventral valve of this genus was found in the 
Onate Formation at the exposures in Johnson Park 
Canyon, locality 3023c. The single ventral valve is of 
about average size for the genus, wider than long with 
acutely pointed beak (85°). A shallow, barely percep- 
tible sulcus originates at about midvalve. The sulcus 
is occupied by five rounded costae and the flanks by 
nine or ten. 

Types.—Figured Specimen: USNM 207699. 


Order RHYNCHONELLIDA Kuhn, 1949 
Superfamily RHYNCHONELLACEA Gray, 1848 
Family TRIGONIRHYNCHIIDAE McLaren, 1965 

Genus CUPULAROSTRUM Sartenaer, 1961b 


Cupularostrum? stevensoni new species 
Plate 14, figures 19-44; plate 41, figures 18-26 


Small for genus, subtriangular in outline, slightly 
wider than long; widest slightly anterior to midvalve; 
anterior commissure strongly uniplicate. Posterolater- 
al extremities forming angle of 100° to 120°. Beak small, 
moderately protuberant, nearly straight to suberect. 
Deltidial plates small, conjunct to disjunct. Foramen 
hypothyridid. Surface completely costate, costae sub- 
angular to narrowly rounded, four on fold, three in 
sulcus, and four or five on flanks. 

Pedicle valve gently but unevenly convex in lateral 
profile and broadly concave in anterior profile. Um- 
bonal and median regions gently convex; sulcus wide 
and deep starting abruptly just posterior to midvalve 
and occupying half or slightly more of valve width; 
tongue long, serrated. Flanks narrow and flattened; 
anterolateral extremities strongly deflected in ventral 
direction. 

Dorsal valve moderately and unevenly convex in 
lateral profile, the maximum convexity in the poste- 
rior; anterior profile rounded dome with median area 
slightly elevated above the convex and steeply de- 
scending flanks. Fold originating posterior to mid- 
valve, elevating above the flanks at about midvalve. 

Ventral valve interior with short stout dental plates. 
Dorsal valve with greatly thickened structures, the 
apical chamber small and narrow; hinge plate divided 
and median septum short and very thick. Inner mar- 
gins of both valves with a row of small interlocking 
ridges. 


70 BULLETIN 315 


Measurements (in mm).— 


dorsal maxi- 
valve mum_ thick- apical 
length length width ness _ angle 


USNM 200896a 


(Holotype; loc. 523v) 11.0 9.8 12.7 7.4 10S° 
USNM 200896b 9.4 8.5 11.5 6.4 107° 
USNM 201029a 

(loc. 509u) 11.2 10.0 13.0 7.4 107° 
USNM 201029b 95 8.4 11.0 6.0 103° 
USNM 200846 

(loc. 524z) 10.4 Sy) 11.9 6.5 103° 


Diagnosis.—Small Cupularostrum with four costae 
on the fold and greatly thickened cardinalia with di- 
vided hinge plate. 

Stratigraphic occurrence.—Contadero and Rhodes 
Canyon formations. 

Localities. —Contadero: 509t; 523q; 526g; Rhodes 
Canyon: 509u; 521g, j; 523p, v; 524z; 526a. 

Types.—Holotype: USNM 200896a. Figured Para- 
types: USNM 200896b, 201029a, b. 

Discussion.—This species is most similar to Pry- 
chomalotoechia sobrina (Stainbrook, 1947) from the 
Percha Formation (Box Member). The Percha species 
is larger than the Rhodes Canyon form when com- 
pared to the largest specimens of C.? stevensoni. Oth- 
er minor differences are in the greater reflection ven- 
trally of the anterolateral margins of P. sobrina, and 
its slightly greater width. A more important difference 
appears on the inside of P. sobrina which has delicate 
cardinalia and an undivided hinge plate. 

Cupularostrum saxitilis (Hall, 1867) is another 
species resembling C.? stevensoni but that species has 
more costae, often with five on the fold, and less 
prominent anterolateral extremities. 

A species of Cupularostrum from the Three Forks 
Formation of Montana, often referred to C. contracta 
(Hall, 1843), is very similar to C.? stevensoni but its 
interior has not been described. Cupularostrum con- 
tracta is a larger and more numerously costate species 
than C.? stevensoni. 


Cupularostrum? species 
Plate 14, figures 45-49 


This species, represented by a single specimen, is 
characterized by having its costation extending onto 
the beaks. It is thus different from Hadrorhynchia 
McLaren (1961), with which it occurs, because that 
genus has smooth umbonal regions. The specimen in 
question is small, with a flattened dorsal valve having 
a broad serrated re-entrant to receive the short tongue 
of the ventral valve. Five strong rounded costae oc- 
cupy the sulcus and six occur on the fold. The sulcus 
begins near midvalve but is shallow and broad. The 
flanks are marked by four broadly rounded costae. 


Measurements (in mm).—USNM 200895: length 
12.8, dorsal valve length 11.4, width 13.4, thickness 
8.8, apical angle 111°. 

Diagnosis.—Completely costate rhynchonellid hav- 
ing six costae on the fold and five in the sulcus. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—523d. 

Types.—Figured Specimen: USNM 200895. 

Discussion.—This specimen is placed questionably 
in Cupularostrum because of its completely costate 
shell, long median septum in the dorsal valve, and 
presence of dental plates. Its nearest relatives seem to 
be undescribed species from the Middle Devonian 
Hamilton Group of New York. 


Genus PTYCHOMALOTOECHIA Sartenaer, 196la 


Ptychomalotoechia sobrina (Stainbrook) 
Plate 41, figures 8-17 


Camarotoechia contracta Kindle (not Hall, 1843), 1909, p. 22, pl. 
6, figs. 1-2a. 
Camarotoechia sobrina Stainbrook, 1947, p. 314, pl. 47, figs. 19-22. 


Serial sections indicate that this species belongs to 
Ptychomalotoechia. It is a very abundant species and 
is found at most exposures of the Percha Formation 
(Box Member). 

Localities. —506, a, b, p, q; 507a, c, d, f, h, k, 1, m, 
Ns O})t5 Uy ys 509as bs C) e2 5276) de. oesn els O26E 
a; 3029; 3030; 3031b; 3042, s. 

Types.—Holotype: M.A.S. 1594 (=USNM 135244). 
Figured Paratypes: M.A.S. 1594a-c (=USNM 
135245a—c). Camarotoechia contracta Kindle (not Hall, 
1843), USNM 62018a-c. 


Genus ATHABASCHIA Crickmay, 1963 


Athabaschia? species | 
Plate 14, figures 50-57; plate 16, figures 7-11 


Small, widely pentagonal in outline; maximum width 
at about midvalve; sides rounded and anterior margin 
truncated. Apical angle broadly obtuse; beak small, 
protuberant. Fold and sulcus wide, originating in an- 
terior third; margins costate, costae subangular, three 
on fold, two in sulcus and four on flanks. Denial plates 
and median septum short. 

Measurements (in mm).— 


dorsal maxi- 
valve mum thick- apical 
length _—_ length width ness angle 
USNM 200914a 
(loc. S09f) 10.0 8.9 12.8 7.0 128° 
USNM 203525 
(loc. 524u) 134-3} 17 16.4 7.6 123° 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 71 


Diagnosis.—Widely pentagonal Athabaschia? with 
two costae in the sulcus. 

Stratigraphic occurrence. 
(Thomasaria Zone). 

Locality.—S09f; 524u. 

Types.—Figured Specimens: USNM 200914a, 
203525. 

Discussion.—This species suggests ‘‘Pugnoides”’ 
solon Thomas and Stainbrook (1922) but is smaller, 
has a proportionally narrower sulcus and fold and more 
strongly geniculated tongue on the ventral valve. It 
differs from Athabaschia? sp. 2 in its smaller size, 
narrower outline and more strongly developed antero- 
lateral extremities bounding the sulcus. 


Sly Gap Formation 


Athabaschia? species 2 
Plate 21, figures 32-37 


Two other specimens are referred questionably to 
this genus. Both have a long median septum and both 
have the characteristic exterior: broadly elliptical, 
smooth posterior half, costate anterior with three cos- 
tae on fold and two in sulcus. These specimens have 
stronger and more rounded costae than the other two 
lots referred to Athabaschia? species 1. 

Stratigraphic occurrence.—Thoroughgood Forma- 
tion. 

Locality.—S22. 

Types.—Figured Specimens: USNM 20104 1a, b. 


Genus RIPIDIORHYNCHUS Sartenaer, 1966b 


Ripidiorhynchus newmexicoense new species 
Plate 15, figures 1-20; plate 41, figures 34-40 


Large, broadly subtriangular in outline; wider than 
long; maximum width slightly anterior to midvalve. 
Anterior commissure strongly uniplicate. Sides nar- 
rowly rounded; anterior margin truncated, postero- 
lateral margins forming an angle of 100° to 120°. Beak 
small, suberect; foramen small, deltidial plates small, 
questionably disjunct. Surface completely costate, 
costae strong, usually four on fold, although a costa 
may be emplaced on one or both lateral slopes of fold, 
usually three in sulcus and five or six on flanks, which 
are not always symmetrical, one flank having six cos- 
tae, the other only five. 

Ventral valve in lateral profile slightly convex in 
posterior half or less, becoming strongly geniculated 
anteriorly; anterior profile fairly deeply concave, an- 
terolateral extremities strongly protuberant and 
bounding curve. Sulcus originating suddenly at mid- 
valve, deepening rapidly and occupying about 60 per- 
cent of maximum width. Flanks narrow, slightly con- 
vex and having a gentle slope to the margins. Tongue 
long, anteriorly serrate, slightly rounded and nearly 
flat. 


Dorsal valve moderately convex in lateral profile, 
most so in posterior part; anterior profile strongly 
semielliptical, slightly elevated medially by fold. Um- 
bonal median region and flanks swollen. Fold origi- 
nating slightly posterior to midvalve, moderately ele- 
vated above flanks at anterior. 

Ventral valve interior having fairly long divergent 
dental plates. Dorsal valve with delicate cardinalia, 
hinge plate undivided, outer hinge plates fairly wide, 
inner hinge plates joined and forming a low arch over 
the apical chamber; socket ridges high and thin. Me- 
dian septum thin and long, extending about to mid- 
valve. 

Measurements (in mm).— 


dorsal maxi- 
valve mum _ thick- apical 
length length width ness _ angle 


USNM 200920a 


(Holotype; loc. 509t) 15.5 14.2 17.8 11.6 110° 
USNM 200920b 15.8 14.7 17.8 11.1 110° 
USNM 200920c 17.4 16.3 21.0 13.2 ite 
USNM 200892 

(loc. 526g) 17.6 16.2 20.3 13.4 Se 


Diagnosis.—Large, strongly costate Ripidiorhyn- 
chus having the length 85 to 90 percent of the width. 

Stratigraphic occurrence.—Contadero Formation; 
Rhodes Canyon Formation. 

Localities. —S09t, u; 521j; 522a; 524t; 526g; 528a; 
3054k. 

Types.—Holotype: USNM 200920a. Figured Para- 
types: USNM 200892, 200920e, 204952. Unfigured 
Paratypes: USNM 200920b-d. 

Discussion.—The nearest species to this one is Ri- 
pidiorhynchus orbicularis (Hall, 1860a) from the Che- 
mung Formation of New York. Although the two 
species are similarly marked, that from New York and 
Pennsylvania is rounder, has a wider fold and sulcus, 
and less extended or reflected anterolateral extremi- 
ties. 

MORPHORHYNCHUS new genus 
[Gr. Morpho = the shapely one] 


About medium-sized for a rhynchonellid, subrectan- 
gular in outline; anterior commissure broadly unipli- 
cate; umbones smooth; ventral beak small, not pro- 
truding and closely pressed to the dorsal umbo; 
foramen small; deltidial plates small, disjunct. Surface 
costate, costae strong and subangular, dying at um- 
bones. 

Ventral valve interior with fairly long, closely spaced 
dental plates and small teeth. Other features not visi- 
ble. Dorsal valve interior with long median septum 
supporting a short, narrow septalium covered by a 
moderately long inner hinge plate that is perforate at 
the rear. Socket ridges slender. 


BULLETIN 315 


Type species.—Morphorhynchus varicostatum n. sp. 

Diagnosis.—Medium-sized rhynchonellid having 
subrectangular outline, smooth umbones, costate shell 
uniplicate with closely-spaced dental plates and small, 
narrow septalium partially covered by a posteriorly 
perforate inner hinge plate. 

Comparison.—The form and construction of the 
hinge plate suggest relationship to Cupularostrum Sar- 
tenaer (1961b) and, in external form, to Hadrorhyn- 
chia McLaren (1961). It differs from the former in its 
rectangular form, very wide fold and sulcus, and the 
different interior. Cupularostrum is described as hav- 
ing a convex inner hinge plate but this is quite unlike 
the inner hinge plate of Morphorhynchus, whichis flat. 
Furthermore, the crura of Morphorhynchus splay hor- 
izontally while those of Cupularostrum are flattened 
laterally. 

The interior of Hadrorhynchia is entirely different 
from that of Morphorhynchus because its septalium is 
filled with limy tissue rather than being covered by an 
inner plate (McLaren, 1962, fig. 16, p. 60). 


Morphorhynchus varicostatum new species 
Plate 15, figures 21-28; plate 41, figures 41-49 


Medium sized, subrectangular in outline; wider than 
long, length equal to slightly more than 80 percent of 
the width; widest at midvalve. Sides narrowly round- 
ed; anterior truncated; posterolateral margins forming 
an angle of 120° to 140°. Beak small, closely pressed 
onto dorsal umbo; foramen small, deltidial plates small, 
disjunct. Anterior commissure broadly uniplicate. 
Surface costate, costae thick and subangular, two or 
usually three in sulcus and three, but more often, four 
on fold. Micro-ornament mostly worn off but occa- 
sional patches of closely spaced concentric lines are 
preserved. Beaks of both valves smooth. Flanks 
marked by four or five costae. 

Ventral valve gently convex in the posterior half but 
strongly geniculated in dorsal direction in anterior half; 
anterior profile broadly concave, anterolateral extrem- 
ities strongly protuberant. Sulcus broad and deep, 
originating at about midvalve and forming a long, gently 
rounded and serrate tongue. Sulcus usually occupied 
by three costae, the median one of which is stronger 
than the lateral ones. Sulcus occupying about 60 per- 
cent of the valve width. Flanks narrow, flattened, or 
slightly concave. 

Dorsal valve unevenly convex in lateral profile, the 
posterior part gently convex, the anterior flattened; 
anterior profile narrow, rounded dome with steep sides. 
Fold originating slightly posterior to midvalve, elevat- 
ing and widening anteriorly where it stands above the 
narrow, convex flanks. 


Ventral valve interior with narrowly divergent, 
closely spaced dental plates. Dorsal valve with undi- 
vided hinge plate, small apical chamber and unusually 
long median septum extending nearly half valve length. 
Crura long, curved, expanding and flattening distally. 

Measurements (in mm).— 


dorsal maxi- 
valve mum _ thick- apical 
length length width ness _ angle 
USNM 200922a 
(Holotype; loc. 522f) 13.5 21.8 15.8 9.8 Sia 
USNM 200922b 10.7 10.1 13.5 7.8 127° 
USNM 201173 
(loc. 523i) 12.4 11.8 14.3 11.7 122° 
USNM 200919a 
(loc. 506d) 14.3 13.0 17.3 11.4 126° 


Diagnosis.—Medium-sized transversely subrectan- 
gular Morphorhynchus with four costae on the fold. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—506d; 522f, g, 1; 523e, 1. 

Types.—Holotype: USNM 200922a. Unfigured 
Paratypes: USNM 200919a, b; 200922b, c; 201173. 

Discussion.—This species is much smaller than 
Hadrorhynchia mclareni n. sp., and is also smaller and 
differently shaped than Hadrorhynchia alta n. sp., both 
from the Onate Formation. It is most like Hadrorhyn- 
chia eurekaensis Johnson (1966), considered a Cassi- 
dirostrum by Johnson (written commun., 1981), from 
the Middle Devonian (Denay Limestone) of Nevada. 
It differs in having stronger costae in the fold and sul- 
cus, although the number of costae is the same, in the 
more lateral sweep to the costae on the flanks and in 
the smaller, less protuberant beak. 


Family UNCINULIDAE Rzhonsnitskaya, 1956 


Subfamily HYPOTHYRIDININAE 
Rzhonsnitskaya, 1956 


Genus HYPOTHYRIDINA Buckman, 1906 


Hypothyridina emmonsi rotunda new subspecies 
Plate 15, figures 39-56; plate 16, figures 1-6; 
plate 39, figures 8-17 


Hypothyridina sp. B. McLaren and Norris, 1962, p. 30, pl. 14, figs. 
7-9. 


About medium-sized for genus, subcircular in out- 
line; length varying from 0.84 to 1.12 of the width; 
sides broadly rounded, anterior truncated and pos- 
terolateral extremities forming broadly obtuse angle. 
Maximum width at midvalve. Anterior commissure 
strongly uniplicate, the tongue of the ventral valve oc- 
cupying about two-thirds valve width. Beak small and 
incurved, often pressed closely to the dorsal valve 
umbo. Surface marked by flatly rounded costae, ten 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 73 


to 14 on the fold, nine to 13 on the sulcus, and up to 
21 on the flanks. 

Ventral valve gently convex in lateral view and 
broadly concave in anterior view, the median part flat- 
tened but the flanks strongly deflected in ventral di- 
rection. Median region swollen; sulcus wide, shallow, 
and defined anteriorly by the strongly deflected an- 


terolateral extremities; tongue long, strongly genicu- 
lated, and broadly rounded anteriorly. 

Dorsal valve very deep, gently convex in lateral pro- 
file, the surface sloping as the valve thickens ante- 
riorly; anterior profile high, rounded dome with nearly 
straight sides; flanks gently convex and slightly folded 
medially at the anterolateral extremities. 

Measurements (in mm).— 


dorsal 
length valve width fold costellae 
(L) length (W) thickness width on tongue L/W ratio 

USNM 200947b 

(Holotype; loc. 506v) 21.0 19.7 21.8 17.6 15.0 10 0.96 
USNM 200950a 

(loc. 509g) 23.4 21.4 25.5 19.1 16.6 11 0.92 
USNM 200950c 

(loc. 509g) 19.8 18.4 21.4 10.6 12.3 10 0.92 
USNM 201160c 

(loc. 509h) 14.0 13.0 12.5 11.2 8.5 7 12, 
USNM 201172 

(loc. 509m) 26.9 25.0 26.0 23.6 18.2 13 1.07 
USNM 201171b 

(loc. 521d) 14.4 13.6 15e5 12.3 10.2 9 0.93 
USNM 200946 

(loc. 521le) 24.1 PSS) 28.2 21.0? 20.0 12 0.85 
USNM 200948b 

(loc. 506c) 20.0 18.0 22.0 17.1 10.7 9 0.91 
USNM 203499a 

(loc. 506c) 16.3 14.7 16.0 13.0 11.9 11 1.02 
USNM 203499b 

(loc. 506c) 15.2 14.6 ISS) 13.1 13.0 14 0.98 
USNM 203496 

(loc. 506i) 22.9 21.5 24.6 19.7 19.1 12 0.93 
USNM 20116la 

(loc. 506j) 14.6 13.7 15.8 12.8 8.7 8 0.92 
USNM 14023a 

Holotype of Hypothyridina emmonsi 

(Hall and Whitfield) 23.1 21.6 26.4 20.6 17.0 13 0.87 


Diagnosis. —Like Hypothyridina emmonsi (Hall and 
Whitfield, 1877), but less transverse than the typical 
subspecies. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S506c, i, j, v; SO9f, g, h, i, m,n, s, w, Z; 
521c, d, e, k, m,n, p, r, u; 524m, w; 526k. 

Types.—Holotype: USNM 200947b. Figured Para- 
types: USNM 200946, 200947c, 200948b, 200949a, 
200950c, 201160a, 201162. Unfigured Paratypes: all 
listed Measured Specimens. 

Discussion.—Hypothyridina 1s fairly common in the 
upper part of the Sly Gap Formation in the Sacramen- 
to Mountains, but it is less abundant in the San Andres 
Range. In many places, two size grades of specimens 
occur, one that ranges up to about 15 mm long and 
another that is larger. The size range suggests a second 
species but the small ones have the same proportions 
as the larger ones and are probably specimens that 
reached adulthood before they attained the more nor- 


mal larger size. All of the specimens are here regarded 
as a single subspecies. 

Compared to the type specimen of H. emmonsi, the 
New Mexico specimens have the same proportions of 
thickness to width and of width of fold to width, but 
when the length/width ratio is compared the New 
Mexico specimens are nearly all less transverse than 
the Nevada specimen. The mean length/width ratio 
suggests a more nearly circular shell—that is, length 
approximately equals width—than the Nevada species, 
which is visibly transverse. Unfortunately, only one 
good specimen is known from Nevada and, conse- 
quently, its variation is unknown. 


Hypothyridina species 
Plate 15, figures 29-38; plate 23, figures 1-6 
About medium-sized for genus, rectangularly 


subquadrate in outline; sides moderately rounded; an- 
terior truncated. Anterior commissure strongly unipli- 


74 BULLETIN 315 


cate. Surface marked by fairly broad costae, about 12 
on the flanks, eight in the sulcus, and nine on the fold. 

Ventral valve with posterior half moderately swol- 
len, swollen area separated from the lateral parts by 
gently concave areas; anterior profile broadly concave 
but with median part slightly convex. Posterolateral 
extremities not prominent. Sulcus forming about 65 
percent of the midwidth. 

Dorsal valve fairly strongly convex in posterior half, 
anterior half flattened; anterior profile narrow, steep- 
sided dome; flanks gently inflated; median region 
strongly swollen. 

Measurements (in mm).—USNM 201090a: length (L) 
18.7?, dorsal valve length 17.3?, maximum width (W) 
21.4, thickness (T) 15.8, width of fold (F) 12.2, L/W 
ratio = 0.88, T/W ratio = 0.73, F/W ratio = 0.57. 

Diagnosis.—Medium-sized, somewhat transverse 
Hypothyridina having strong crowded costae of which 
eight occupy the tongue of the ventral valve. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—506d; 509p; 522n; 523d; 524r; 526b, c: 
528s. 

Types.—Figured Specimens: USNM 201090a, b; 
201141. 

Discussion.—A few poor specimens of Hypothyri- 
dina from the Onate Formation are reminiscent of H. 
venustula (Hall, 1867) or H. procuboides (Kayser, 
1871), both from the Givetian. The costation of Hy- 
pothyridina cameroni Warren (1944) has some resem- 
blance to the Onate form, but it is larger, wider, and 
has more flaring lateral extremities. Specimen USNM 
201141 appears to be an immature specimen of this 
Onate species of Hypothyridina. 


Subfamily HADRORHYNCHIINAE McLaren, 1965 
Genus HADRORHYNCHIA McLaren, 1961 


Hadrorhynchia alta new species 
Plate 16, figures 12-31; plate 18, figures 8-17 


Small for genus, narrowly pentagonal in outline; 
length and width nearly equal; sides broadly rounded, 
anterior margin truncated; posterolateral extremities 
forming an angle of 98° to 118°. Anterior commissure 
strongly uniplicate; beak low, incurved, suberect. Del- 
tidial plates disjunct. Posterior smooth, noncostate but 
marked by fine, flattened radial lines. Anterior marked 
by strong angular costae, three or four on fold with 
one less in sulcus, and usually three, occasionally four, 
on lateral extremities. 

Ventral valve convex in posterior half in lateral pro- 
file but strongly geniculated anteriorly; anterior profile 


deeply concave, flanks swept up into narrow points. 
Umbonal region narrowly swollen, swelling extending 
onto anterior before inception of sulcus; umbolateral 
slopes short and rounded. Sulcus beginning at mid- 
valve, deepening rapidly and occupying three-fifths the 
shell width. Tongue long, geniculated and flattened an- 
teriorly, the costae and interspaces having narrow, 
flattened longitudinal bands. 

Dorsal valve deep, moderately convex in lateral 
profile but narrowly domed in anterior profile with fold 
protruding beyond the rounded, precipitous flanks. 
Fold originating at midvalve, narrow, strongly elevat- 
ed above flanks at anterior. Anterior margin genicu- 
lated to meet that of ventral valve. Troughs between 
costae extended anteriorly and laterally as flattened 
bands to meet troughs and ridges of opposite valve. 

Measurements (in mm).— 


dorsal 

valve thick- apical 
length length width ness angle 

USNM 201139a 
(loc. 523b) 18.8 16.4 20.1 15:5 Se 
USNM 201139b 16.9 15.1 18.2 10.2 108° 

USNM 201140b 
(Holotype; loc. 523d) 19.8 17.0 20.0 13.3 101° 
USNM 201140a 19.6 17.1 20.7 15.0 118° 
USNM 201140d 16.3 14.3 18.0 12.0 WISE 
USNM 201140e 12.6 11.5 13.5 Sh) 108° 


Diagnosis.—Small Hadrorhynchia having a narrow 
fold, and acute anterolateral extremities that are ex- 
tended strongly in a ventral direction. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —522f, 523a, b, d, e, f, h, i; 3025c. 

Types.—Holotype: USNM 201140b. Figured Para- 
types: USNM 200898a, b; 201139a; 201140a; 203526. 
Unfigured Paratypes: USNM 201139b; 201140c—e. 

Discussion. —Compared to Hadrorhynchia sander- 
soni (Warren, 1944), in the length range of 18 to 22 
mm, the specimens from New Mexico prove to have 
nearly equal length and width (mean L/W ratio = 0.94). 
McLaren’s specimens have a mean T/L ratio = 1.14 
whereas the same ratio for H. alta n. sp. is 1.06. The 
mean W/L ratio of H. sandersoni in this size range is 
0.78 but that of H. alta is 0.72. Compared to H. mclar- 
eni n. sp., H. alta does not attain the large size of the 
former, has more strongly deflected anterolateral ex- 
tremities, fewer costae on the flanks, and a narrower 
fold. 

This species, like its relative H. mclareni, is re- 
stricted to the shaly facies of the Onate Formation, 
but it has not been found outside of the Caballos 
Mountains and Derry Hills. 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DUTRO 75 


Hadrorhynchia mclareni new species 
Plate 17, figures 20—43 


Medium-sized for genus, subpentagonal to broadly 
oval in outline; valves unequal in depth and convexity, 
dorsal valve having greater dimensions in both; sides 
rounded, anterior truncated, posterolateral extremities 
forming angle of 118° to 128°. Beak small, incurved; 
beak ridges sharp, defining small, curved interareas. 
Deltidial plates disjunct. Anterior commissure strong- 
ly uniplicate. Fold having four or five costae, the flanks 
five or six. Anterior surface plicate, posterior without 
plication but whole surface covered by fine, flat, ra- 
diating capillae and crossed by fine concentric threads. 
Costae in sulcus bearing a median flat groove. 

Ventral valve gently convex in lateral profile, the 
posterior half flatly convex but the anterior strongly 
curved to geniculate; anterior profile flatly to strongly 
concave; umbonal region slightly swollen; postero- 
lateral areas rounded and sloping steeply to posterior 


margins; sulcus originating abruptly at about mid- 
valve, wide and deep, occupying slightly more than 
half valve width. Flanks bounding sulcus variable from 
flattened and slightly flaring to strongly elevated. 
Tongue long and anteriorly geniculated in the adult. 

Dorsal valve unevenly convex in lateral profile, an- 
terior part gently convex but umbonal region moder- 
ately curved. Anterior profile forming strong dome 
having steeply sloping sides, becoming abruptly ver- 
tical at place of geniculation in adults. Median region 
strongly swollen; flanks fairly strongly swollen. Fold 
forming only at the anterior and extending a variable 
distance posteriorly but usually not extending beyond 
midvalve. Anterior margin geniculated in the adult. 

Dorsal valve interior with broad, concave outer hinge 
plates and elevated socket ridges; crura long, flat- 
tened, distally expanded and concave in an anterior 
direction. 

Measurements (in mm).— 


dorsal maximum 
length valve width thickness 
(L) length (W) (T) apical angle L/W ratio 

USNM 200897b 

(loc. 523b) 14.0 13.0 16.4 ee 120° 0.85 
USNM 200897a 

(loc. 523b) 25.0+ 24.0 31.5 18.5 127° 0.84 
USNM 20089 1a 

(Holotype; loc. 523d) 26.0 23.8 29.0 16.5 W233 0.90 
USNM 200891b 

(loc. 523d) 22.7 20.3 27.0 18.5 ip 0.83 
USNM 20089 1c 

(loc. 523d) 25.0? 23.0 29.8 18.0 252 0.85? 
USNM 200891d 

(loc. 523d) 24.8 21.3 BST 18.0 118° 0.97 
USNM 203500 

(loc. 523k) 25.4? 24.0 31.4 16.4 128° 0.90 
USNM 203501la 

(loc. 523b) 23.5 21.4 27.8 15.0 119° 0.85 
USNM 203501b 

(loc. 523b) 21.5 19.6 25.5 12.5 125° 0.84 
USNM 203502a 

(loc. 523f) 23.0? 21.2 25.0 20.0 118° 0.88 
USNM 203502b 

(loc. 523f) 11.7 10.5 12.3 5.0 106° 0.87 


Mean T/W ratio = 0.65; Mean L/W ratio = 0.87; Mean W/L ratio = 1.07. 


Diagnosis.—Hadrorhynchia of medium size and 
moderate thickness. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—523b, d, f, h, k. 

Types.—Holotype: USNM 200891a. Figured Para- 
types: USNM 200891d, e, g; 200897a, b; 200903; 
200918. Unfigured Paratypes: USNM 200891b, c, f; 
200892a; 203500; 203501a, b; 203502a, b. 

Discussion.—This species is very close to H. san- 


dersoni (Warren, 1944) but that species has a much 
thicker and generally rounder shell than the New Mex- 
ico form. McLaren (1962, p. 62) gives the mean T/L 
ratio for H. sandersoni as 0.94 and the mean W/L ratio 
as 1.07 for specimens 22 mm or more in length. The 
same proportions of H. mclareni of this size group are: 
mean T/L ratio = 0.65 and mean W/L ratio = 0.87. A 
specimen from the Beavertail Limestone on McKenzie 
River attains larger dimensions than any specimen from 


76 BULLETIN 


New Mexico, and is also larger than any listed by 
McLaren, but its proportions are in accordance with 
those cited by McLaren. 

Hadrorhynchia mclareni is an uncommon species 
and, so far as we now know, it is restricted to the 
shaly facies of the Onate Formation in Mud Springs 
Mountains, the Derry Hills and Caballos Mountains. 


Family PUGNACIDAE Rzhonsnitskaya, 1956 
Genus COELOTERORHYNCHUS Sartenaer, 1966a 


Coeloterorhynchus magnificum new species 
Plate 17, figures 4-19; plate 18, figures 31-57; 
plate 40, figures 1-10 


Pugnoides schucherti Stainbrook, 1948, p. 774, pl. 2, figs. 34, 35, 
37. 


Large for genus, rounded pentagonal in outline, 
wider than long; maximum width at or near midvalve; 
sides strongly rounded; anterior margin truncated; 
posterolateral margins forming angle of approximately 
120°. Anterior commissure strongly uniplicate. Valves 
very unequal in depth, the ventral valve shallow, the 
dorsal valve deep. Beak small, incurved, erect; fora- 
men small, often bisecting the beak; deltidial plates 
disjunct. Posterior two-thirds smooth, margins varia- 
bly costate, flanks having four or five costae, fold two 
to five, usually three or four, and sulcus one to four. 

Ventral valve fairly strongly convex medially but 
broadly concave in anterior profile and having the an- 
terolateral extremities swept up in a ventral direction. 
Median region flatly convex, posterolateral slopes short 
and convex; sulcus wide, originating slightly posterior 
to midvalve, deepening anteriorly, and occupying about 
0.7 the valve width; tongue long and strongly bent in 
a dorsal direction, its end strongly serrate and narrow- 
ly rounded. Flanks steeply dipping toward middle of 
valve. 

Dorsal valve unevenly convex in lateral profile, pos- 
terior half moderately convex but anterior half flat- 
tened; anterior profile amost hemispherical, sides 
sloping strongly, and flanks gently convex. Fold form- 
ing at about midvalve usually narrow, occupying about 
half valve width; fold variable, usually with the one or 
two median costae elevated above the others, which 
appear on the slope of fold. Anterolateral extremities 
strongly directed ventrally. 

Measurements (in mm).— 


maxi- 

dorsal mum 
length valve width thick- apical L/W 
(L) length (W) ness angle ratio 


USNM 200915a 
(loc. 506c) 


10.9 i 11.8 5.3 107° 0.93 


Ww 
—_ 
in 


maxi- 

dorsal mum 
length valve width thick- apical L/W 
(L) length (W) ness angle ratio 


USNM 200915b 


(loc. 506c) iW ill WP 60 ie a9) 
USNM 200915c 

(loc. 506c) 13.4 12.0 16.0 6.6 112° 0.84 
USNM 200915d 

(loc. 506c) Vie iy il Ga ISP O95 
USNM 200915e 

(loc. 506c) 16.4 15.0 16.8 9,6 122° 0.98 
USNM 200915f 

(loc. 506c) We) 195 we TO ie Oss 
USNM 200915g 

(loc. 506c) 18:5 17.0) 1955 | 12:4.) 1252 mO195 
USNM 200915h 

(loc. 506c) Is I Al iil ia O.S7 
USNM 200915i 

(loc. 506c) 16.4 15.0 18.7 10.8 120° 0:87 
USNM 200915} 

(loc. 506c) 20.9 19.2 21.4 13.0 125° 0.98 
USNM 200915k 

(loc. 506c) Mi iY) WH WA WEP O95 
USNM 2009151 

(Holotype) 22.6 20.2 25.6 14.8 124° 0.88 
USNM 200913 

(loc. 506c) 20.4 18.3 20.7 14.8 114° 0.99 
USNM 200912 

(loc. 521p) mi iS WIG i682) iP OES 
USNM 200828c 

(loc. 526k) AM i WR 16) iE O97 
USNM 200828a 

(loc. 526k) ey A Ais 1 iP Ose 
USNM 200828b 

(loc. 526k) me We a IG ie OSs 


Diagnosis.—Large, nearly circular to slightly trans- 
verse Coeloterorhynchus with moderate thickness. 

Stratigraphic occurrence.—Sly Gap Formation; 
Contadero Formation. 

Localities. —Sly Gap: 506c, i, j, v, y; 509f, g, h, ha, 
i, j, mM, n, 0, p, q, S, V, W; 52I1c, e, k, 1, m, 0, p, r; 
524m, v; 526k, 1, x; 529s, x; USGS (Kirk) 541; 543. 
Contadero: 521h. 

Types.—Holotype: USNM 2009151. Figured Para- 
types: USNM 200915b, c, g, i, k; 200912; 200913; 
200916; 200917a. Unfigured Paratypes: all Measured 
Specimens. 

Discussion.—The young of this species are nearly 
circular as they have length and width nearly equal 
and the L/W ratio usually equals 0.90 or more. The 
species attains a large size, the largest species of the 
genus so far found in this country. In this regard, it 
differs strongly from Coeloterorhynchus calvini (Fen- 
ton and Fenton, 1924) from the Cerro Gordo Member 
of the Hackberry Formation, which is small and has 
an exaggerated depth dimension. The fold of C. calvini 
is strongly angular and the marginal costae are strong- 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 77 


ly angular. Coeloterorhynchus magnificum n. sp. also 
attains a larger size than C. schucherti (Stainbrook, 
1945) from the Independence Formation of Iowa. The 
Independence species is quite variable and has a L/W 
ratio like that of C. magnificum but its T/W ratio is 
much greater and indicates a shell with exaggerated 
depth like that of C. calvini. Comparison of Indepen- 
dence species is usually difficult because of the rare 
and spasmodic outcrops and general unavailability of 
collections. The ten specimens of C. schucherti in the 
National Collections are mostly small and the largest 
one is smaller and much differently shaped than C. 
magnificum; C. schucherti is much more like C. cal- 
vini and is probably related to that species rather than 
to the one from New Mexico. 

Coeloterorhynchus magnificum n. sp. is fairly com- 
mon in the Sly Gap Formation of the Sacramento 
Mountain Range, but it is less common in the San 
Andres Range and is very rare west of there. The 
species is variable in the degree of costation, the an- 
gularity of the fold and, to some extent, in its dimen- 
sions. 


Coeloterorhynchus species 
Plate 14, figures 58-65 


Small, pentagonal in outline, pugnacid in appear- 
ance with posterior regions smooth and anterior mar- 
gins strongly costate; length and width about equal: 
apical angle nearly 90°; sides bulging; anterior margin 
truncated. Anterior commissure strongly uniplicate. 
Beak small, moderately protuberant. Anterolateral ex- 
tremities bounding the sulcus projecting strongly. Cos- 
tae narrowly subangular, three on the fold, two in the 
sulcus, and four on the flanks. 

Ventral valve gently convex in posterior half, genic- 
ulated in a dorsal direction in anterior half; anterior 
profile gently concave; sulcus originating at anterior 
third, narrow, occupying slightly less than half width. 
Dorsal valve nearly flat in lateral profile but narrowly 
domed in anterior view. Fold short, originating at about 
two-thirds valve length from the beak. Flanks mod- 
erately swollen. 

Ventral valve interior with dental plates and dorsal 
valve having a short median septum. 

Measurements (in mm).— 


dorsal = maxi- 
valve mum thick- apical 
length length width ness angle 


USNM 200894 
(loc. 509f) 12.1 10.5 11.8 8.6 104° 


Stratigraphic occurrence.—Sly Gap Formation. 
Localities.—SO9f; 522k. 
Types.—Figured Specimen: USNM 200894. 


Discussion.—This species is suggestive of some 
forms of *““Pugnoides”’ alta Williams (1890) from the 
State Quarry Formation of Iowa, but it differs in its 
narrower fold, greater proportional width and sharper 
costae. The species is very rare. 


COLOPHRAGMA new genus 
[Gr. kolos, shortened; phragma, fence (septum)] 


Medium-sized rhynchonellid, transversely elliptical 
to transversely pentagonal in outline; biconvex, the 
dorsal valve having the greater convexity and depth; 
anterior commissure strongly uniplicate. Beak small, 
closely pressed onto the dorsal umbo; deltidial plates 
conjunct; foramen small, mesothyridid. Surface semi- 
costate, costae strong and subangular. 

Ventral valve interior with long dental plates defin- 
ing a deep delthyrial cavity. 

Dorsal valve interior having a short median septum 
supporting a small, short septalium; outer hinge plates 
broad and flattened, nearly meeting medially; crural 
bases thick. Other details lacking. 

Type species.—Colophragma ellipticum n. sp. 

Diagnosis.—Rhynchonellacea having the exterior of 
pugnacids, with an aborted median septum and broad 
outer hinge plates. 

Comparison and discussion.—This genus has the 
exterior of Athabaschia Crickmay (1963), Coelote- 
rorhynchus Sartenaer (1966a), Ningbingella Roberts 
(1971), Shumardella Weller (1910), and Parapugnax 
Schmidt (1964). It differs from all of these, except 
Coeloterorhynchus, in the brevity of its median sep- 
tum. From Coeloterorhynchus, it differs in the pres- 
ence of a short median septum, a feature lacking in 
that genus. 

Colophragma is like Chapinella Savage, Eberlein, 
and Churkin (1978) in external appearance but differs 
in internal details. Chapinella has no median septum 
and the hinge plate is divided. In Colophragma the 
hinge plate is undivided and there is a short median 
septum. 


Colophragma ellipticum new species 
Plate 16, figures 49-58; plate 17, figures 1-3; 
plate 18, figures 3-7; plate 40, figures 11-19 


About medium size for genus, broadly triangular in 
outline; wider than long; sides narrowly rounded; an- 
terior margin broadly and gently rounded; postero- 
lateral margins forming angle of 110°-120°. Anterior 
commissure strongly uniplicate. Beak small, suberect 
to erect. Surface paucicostate, the posterior half 
smooth, the anterior half costate; two costae in sulcus, 
three on fold, and two indistinct ones on flanks. 

Ventral valve in lateral profile gently convex at the 
posterior, but bent near midvalve into a broad curve 


78 BULLETIN 315 


to form a long, sharply serrated tongue. Anterior pro- 
file broadly concave and having strongly projecting 
anterolateral extremities. Sulcus beginning abruptly at 
midvalve, deep, and forming about 60 percent of the 
midwidth. Flanks narrow and flat. 

Dorsal valve gently convex in lateral profile, most 
convex in the umbonal region. Anterior a broad and 
high dome having steep sides. Fold originating at about 
midvalve just slightly posterior of the initiation of the 
three costae forming it. Fold moderately elevated above 
narrow but inflated flanks. 

Ventral valve interior with widely divergent dental 
plates. Dorsal valve with short median septum. 

Measurements (in mm).— 


dorsal = maxi- 
valve mum _ thick- apical 
length length width ness angle 
USNM 200921 
(Holotype; loc. S210) 15.2 14.0 18.2 12.5 116° 
USNM 200953 
(loc. 5240) 13.8 12.7 17.6 12.0 118° 
USNM 203537 
(loc. 528b) 13.7 2 17.5+ 9.7 120° 


Diagnosis.—Broadly elliptical Colophragma having 
two costae in the sulcus and two on the flanks. 

Stratigraphic occurrence.—Contadero Formation. 

Localities. —523q; 5240; 528b. 

Types.—Holotype: USNM 200921. Figured Para- 
types: USNM 200953; 203537. 

Discussion.—This species somewhat resembles some 
forms of Coeloterorhynchus magnificum n. sp., but it 
differs in being wider, with shallower sulcus, and nev- 
er with the fold so far extended as in C. magnificum 
n. sp. Resemblance to Petasmaria n. gen. is also shown 
in the wide outline, but that genus usually has a deeper 
sulcus and the median septum is long and prominent. 
The interior details will also serve to distinguish this 
species from specimens of Athabaschia Crickmay 
(1963), which have a strong external resemblance. 


Colophragma species | 


A specimen of Colophragma occurs with Colo- 
phragma ellipticum n. sp., from the Contadero For- 
mation. Compared with specimens of C. ellipticum it 
is much larger, has more convex flanks, a more convex 
ventral valve, lower ventral beak, and a more rounded 
dorsal fold with four costae and a poor development 
of costae on the flanks. 

Measurements (in mm).—USNM 203503: length 
18.6, brachial valve length 16.5, maximum width 20.3, 
thickness 15.0. 

Locality.—524o0. 

Types.—Mentioned Specimen: USNM 203503. 


Colophragma species 2 
Plate 18, figures 1, 2 


About medium size for genus, transversely elliptical 
in outline, sides rounded; maximum width at about 
midvalve. Ventral valve flatly convex and having a 
wide sulcus occupying nearly three-fourths the valve 
width. Sulcus occupied by three costae that originate 
at about midvalve. Dorsal valve moderately convex, 
much deeper than the ventral valve and having the fold 
originating at about midvalve. Fold moderately ele- 
vated at the anterior and marked by four narrowly 
rounded costae. Posterior half of both valves smooth, 
flanks with two costae at the margin. 

Ventral valve interior having short, nearly obsolete 
dental plates. Dorsal valve with a very short median 
septum. 

Measurements (in mm).—USNM 200952: length 
14.7, dorsal valve length 14.1, maximum width 21.0, 
thickness 10.8, apical angle 123°. 

Stratigraphic occurrence.—Thoroughgood Forma- 
tion. 

Locality.—S522. 

Types.—Figured Specimen: USNM 200952. 


Genus TRIFIDOROSTELLUM Sartenaer, 196la 


Trifidorostellum obscurum new species 
Plate 16, figures 32-48; plate 40, figures 28-31 


About medium size, roundly elliptical in outline, 
wider than long; strongly inequivalve, dorsal valve 
having greater depth than ventral one. Sides narrowly 
rounded; anterior margin broadly rounded. Apical an- 
gle 125° to 135°. Anterior commissure strongly unipli- 
cate. Beak small, not prominent, closely pressed onto 
the dorsal umbo. Valves costate, costae narrowly 
rounded, not reaching the beaks, usually three on 
flanks, two in sulcus and three on fold. 

Ventral valve moderately convex in lateral profile, 
medially deeply concave in anterior profile; flanks 
bounding sulcus narrow, gently convex and with steep 
slopes to the posterolateral extremities. Sulcus origi- 
nating about one-fourth the length from the beak (about 
five mm measured on the surface) deepening rapidly 
and forming a long narrow tongue. 

Dorsal valve strongly convex in lateral profile but 
highly domed in anterior profile, the fold standing 
strongly above the narrow, convex flanks. Fold begin- 
ning just anterior to the umbo, narrow, becoming 
strongly elevated anteriorly and forming deep re-en- 
trant anteriorly. 

Ventral interior with short, delicate dental plates. 
Hinge plate divided; no median septum. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 79 


Measurements (in mm).— 


dorsal 
valve thick- apical 
length length width ness angle 


USNM 200980 


(loc. 5070) 12.8 12.4 16.3 14.1 136° 
USNM 200981 

(Holotype; loc. 3029) 11.6 11.6 15.6 11.1 128° 
USNM 201154 

(loc. 3042s) 10.6 9.6 12.3 10.7 127° 
USNM 203504 

(loc. 506b) 11.0? 11.0 15.4 11.2 135° 
USNM 203505 

(Kirk loc. 512) 13.1 11.0 17.3 15.6 127° 


Diagnosis.—Small, rotund Trifidorostellum having 
two costae in the sulcus and three on the flanks. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —S506b, p, q; 507a, n, 0; 527e; 3029; 3042s; 
USGS 512. 

Types.—Holotype: USNM 200981. Figured Para- 
types: USNM 200980; 201154. Unfigured Paratypes: 
USNM 203504; 203505. 

Comparison.—This is the smallest species of the ge- 
nus so far described. The largest specimen is slightly 
narrower than 20 mm, somewhat less than large spec- 
imens of 7. madisonense (Haynes, 1916). Although 
size is significant, the major differences between 7. 
obscurum and the species described by Sartenaer (1969) 
are in the ornament. Sartenaer’s (196la) cited species 
T. uralicum Nalivkin (1947) and T. dunbarense 
(Haynes, 1916) are usually multicostate and have from 
two to five costae in the fold. The flanks of these species 
are variable from smooth to having numerous costae. 


Family CAMAROTOECHIIDAE Schuchert, 1929 
Subfamily CAMAROTOECHIINAE Schuchert, 1929 
Genus CAMAROTOECHIA Hall and Clarke, 1893 


Camarotoechia latisinuata new species 
Plate 19, figures 51-65; plate 40, figures 20-25 


About medium-sized for genus, wider than long; 
subpentagonal to broadly ovate in outline; maximum 
width located anterior to midvalve; posterolateral ex- 
tremities nearly straight or slightly curved; sides con- 
vex and anterolateral extremities narrowly rounded. 
Valves subequally convex. Beak low and usually 
closely appressed onto the dorsal umbo; apical angle 
113° to 127°. Anterior commissure broadly uniplicate. 
Surface multicostate, flanks on each side marked by 
five or more costae, fold by eight and sulcus by one 
less. Costae narrow and rounded, increasing by bifur- 
cation and implantation. 


Ventral valve gently convex in lateral profile and 
most convex in the posterior half; anterior profile 
broadly concave. Umbonal region moderately swol- 
len; posterolateral slopes rounded; sulcus originating 
on umbonal region about five mm anterior to beak, 
widening and deepening rapidly to occupy about 0.8 
the valve width at the front; tongue long and broadly 
rounded. 

Dorsal valve slightly deeper than ventral valve, 
moderately convex in lateral profile, unevenly and 
strongly domed in anterior profile, median region ris- 
ing slightly above flanks. Fold broad and gently con- 
vex in profile, originating on umbo and rising gradually 
anteriorly; flanks convex, steep and well-depressed 
below the fold. 

Ventral valve interior having very short and thin 
dental plates and a small, triangular strongly im- 
pressed muscle field. Dorsal valve with thin, blade- 
like median septum expanded at the proximal end 
where it joins the cardinalia. 

Measurements (in mm).— 


dorsal = maxi- 
valve mum __ thick- apical 
length length width ness angle 


USNM 200945a 


(Holotype; loc. 522) 20.6 19.0 DDE 13.0 125° 
USNM 900945b 18.0 17.6 23.0 10.1 Pa 
USNM 200945c 17.8 16.7 19.8 11.0 116° 


Diagnosis.—Multicostate Camarotoechia having a 
wide sulcus and fold bearing seven and eight costae 
respectively. 

Stratigraphic occurrence. 
tion. 

Locality.—S22. 

Types.—Holotype: USNM 200945a. Figured Para- 
types: USNM 200945b-e. Unfigured Paratype: USNM 
200945f. 

Discussion.—This species strongly resembles spec- 
imens of Camarotoechia multicostata (Hall, 1860a), 
C. mesacostalis (Hall, 1843) and C. sinuata (Hall, 
1867). This is a smaller and less strongly costate species 
than the first; it is larger and has more strongly costate 
flanks than the second; it is also larger than the third, 
has a wider sulcus, stronger costae on the flanks and 
has its maximum width farther to the anterior. 

Camarotoechia latisinuata is common in the Tho- 
roughgood Formation at Sly Gap. 


Thoroughgood Forma- 


Camarotoechia? species 

Plate 18, figures 18-22 
A single specimen, possibly an immature one, was 
found in the Percha Formation (Box Member) at Bear 


80 BULLETIN 315 


Mountain. The fold and sulcus are well-defined re- 
spectively by four and three costae and the flanks have 
two incipient costae. The specimen is too small to re- 
late to other species from Upper Devonian rocks and 
its reference to Camarotoechia is based on exterior 
characters alone. 

Measurements (in mm): USNM 203506: length 9.8, 
dorsal valve length 9.4, width 9.6, thickness 5.2, apical 
angle 112°. 

Stratigraphic occurrence.—Percha Formation (Box 
Member) 

Locality.—509b. 

Types.—Figured Specimen: USNM 203506. 

Discussion.—This specimen does not suggest the 
young of Paurorhyncha Cooper (1942), which are usu- 
ally flatter and widely triangular, rather than nearly 
circular. 


Genus CALVINARIA Stainbrook, 1945 


Calvinaria bransoni Stainbrook 
Plate 18, figures 23-27; plate 19, figures 22-50; plate 
40, figures 32-39 


Calvinaria bransoni Stainbrook, 1948, p. 774, pl. 2, figs. 12-14 (not 
figs. 15, 16); McLaren, 1962, p. 24, 30, text-fig. 6A (p. 28). 


This species was compared by Stainbrook (1948) to 
Calvinaria ambigua (Calvin, 1878) that occurs in the 
Independence Formation, but C. ambigua is a much 
bigger shell and the differences are obvious. Compar- 
ison is better made with Calvinaria albertensis albert- 
ensis (Warren, 1928) redescribed and illustrated by 
McLaren (1962). The two species share many char- 
acters, and McLaren remarks “‘It is considered likely 
that C. bransoni belongs to the same species group as 
C. albertensis but is subspecifically distinct.’’ After 
studying several hundred specimens of C. bransoni, 
we believe that its characters are sufficiently strong 
and distinct from those of the Alberta species to war- 
rant full specific value to C. bransoni. 

C. bransoni is a less transverse species than C. al- 
bertensis. Measurements of 45 specimens between 13 
and 21 mm wide from each of two localities (506c and 
509h) yielded a mean W/L ratio of 1.18 in the first 
locality and 1.19 in the second. According to Mc- 
Laren’s figures (1962, p. 29) the same ratio for speci- 
mens of C. albertensis ranging from 13 mm to the 
maximum width yielded a mean W/L ratio of 1.33- 
1.34, indicating a more transverse species. Other dif- 
ferences appear in the more pronounced costation of 
fold, sulcus and flanks of the New Mexico species. 
McLaren mentions that about a third of the shells of 
the Alberta species develop more than two costae on 
the fold. This feature was seen in only six specimens 
in the entire collection of slightly more than 500 spec- 
imens from New Mexico. 


Calvinaria bransoni is also a variable species, the 
variation being expressed in shape as well as orna- 
ment. Some specimens are rounder than others but 
generally the maximum width remains close to mid- 
valve. A few specimens have subdued costae, espe- 
cially those at locality 521d. The New Mexico species 
is thin-shelled, as a rule, and the thickness of the 
species is variable but tends to be less than that of C. 
albertensis. The latter has a mean T/L ratio of 0.70 for 
specimens in the 13.0 mm to maximum width size class 
for the species. Forty-five specimens of C. bransoni, 
of the same range of measurement, have a mean T/L 
ratio of 0.61, indicating a more compressed shell than 
that of C. albertensis. 

Calvinaria bransoni is most abundant in the Sacra- 
mento Mountains, especially at localities 506c and 
509h. It is fairly common in the San Andres Range, 
but its abundance there is not comparable to that in 
the Sacramento Mountains. West of the San Andres 
Range it is rare. In general, the Sly Gap is less fossil- 
iferous in the more western localities of the formation. 

Measurements of illustrated specimens (in mm).— 


dorsal 
valve thick- apical 
length length width ness angle 


USNM 135269 
(Holotype; loc. 506c) 15.9 15.0 18.4 10.5 144° 
USNM 135270 


(M.A.S. paratype) 17.6 16.8 20.8 10.3 144° 
USNM 200861la 

(loc. 509h) 16.0 15.0 20.4 10.6 148° 
USNM 200861b 

(loc. 509h) 17.2 16.4 20.8 NiS7/ 150° 
USNM 200861c 

(loc. 509h) 17.0 16.3 21.0 11.4 151° 
USNM 200856 

(loc. 509h) 14.3 13.7 19.6 8.5 148° 
USNM 200857 

(loc. 521e) 14.5 13.8 19.6 8.6 149° 
USNM 200858 

(loc. 521k) 16.7 16.0 22.0 10.5 150° 
USNM 200860 

(loc. 526k) 17.8 16.8 22.8 11.0 144° 


Diagnosis.—Strongly costate Calvinaria having two 
prominent costae defining the fold and one in the sul- 
cus. 

Stratigraphic occurrence.—Sly Gap Formation (up- 
per part); Hackberry Formation (Cerre Gordo Mem- 
ber). 

Localities.—S06c, i, j, v, y; 509f, h, j, n, q, S, V, Zz; 
S2ilce ds es ke ms, Ss Vis 523C> 52405 venws 52 Oeemke 
528d, 529x. 

Types.—Figured Hypotypes: USNM 200856b, d; 
200857; 200858; 200859; 200860; 200861a, e. 

Discussion.—lt is interesting to record the presence 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 81 


of Calvinaria in the Cerro Gordo Member of the Hack- 
berry Formation. This specimen seems to accord well 
with C. bransoni (Pl. 19, figs. 46-50) even though it is 
slightly damaged. The specimen (USNM 200859) mea- 
sures (in mm): length 15.7, length of dorsal valve (L) 
15.2, width (W) 18.0, thickness (T) 10.4, apical angle 
115°, W/L ratio = 1.15, T/L ratio = 0.66. The appar- 
ent roundness of the specimen is due to a small amount 
of lateral compression. It is also likely that this small 
distortion accounts for an apical angle that is relatively 
small compared to that usual in C. bransoni. 


Genus LEIORHYNCHUS Hall, 1860a 


Leiorhynchus sphaericum new species 
Plate 19, figures 11-21 


About medium-sized for genus, globular, nearly cir- 
cular in outline; valves unequal in depth, dorsal valve 
having greater depth. Sides broadly rounded, posterior 
somewhat narrowly rounded and anterior margin trun- 
cated. Apical angle 106° to 128°. Anterior commissure 
strongly uniplicate. Ventral beak low, in some speci- 
mens not extending posterior to dorsal umbo but al- 
ways pressed down on dorsal valve. Exterior sparsely 
costate at anterior but smooth posteriorly. Fold marked 
by three or four broad, low costae, sulcus by one less, 
flanks unmarked. 

Ventral valve moderately convex in lateral profile 
but with the anterior tongue fairly strongly geniculat- 
ed; anterior profile gently and broadly convex. Pos- 
terolateral regions steep; umbonal region somewhat 
swollen and marked by initial appearance of costae 
appearing in sulcus, which begins just anterior to mid- 
valve; sulcus abrupt and wide, occupying two-thirds 
valve width; tongue long and anteriorly truncated. 

Dorsal valve almost hemispherical in lateral profile 
and strongly and narrowly domed in anterior profile, 
the sides being almost straight. Umbonal, median and 
flank regions inflated. Costation of fold appearing just 
anterior to umbo, not forming a fold until about mid- 
valve; fold inconspicuous and only slightly raised above 
flanks. 

Measurements (in mm).— 


dorsal 
valve thick- apical 
length length width ness _ angle 


USNM 200851 

(Holotype; loc. 5221) 17.6 17.6 17.8 15.0 120° 
USNM 200893 

(loc. 523e) 17.0 16.4 16.3 12.8 106° 
USNM 201187a 

(loc. S09f) 18.6 18.6 17.3 16.8 152 
USNM 201187b 

(loc. SO9f) 16.5 16.4 15.1 12.2 114° 


Diagnosis.—Small, globular Leiorhynchus having 
coarse anterior costae. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—522f, 1, m; 523d, e; 524x; 302Sa. 

Types.—Holotype: USNM 200851. Figured Para- 
type: USNM 200893. Unfigured Paratypes: USNM 
201187a, b. 

Discussion.—This species is suggestive of Leiorhyn- 
chus castaneum (Meek, 1867) in the strictest sense. 
As that species is now understood, it includes a large 
number of specimens much larger than the holotype, 
which is a small specimen (McLaren, 1962, p. 83). The 
type specimen of L. castaneum (Meek, 1867) (USNM 
13854) measures (in mm): length 21.5, length of dorsal 
valve 21.5, width 20.0, thickness 15.2, apical angle 
127°. Although the dimensions of this specimen are 
close to those of L. sphaericum, as listed above, other 
differences readily separate the two. The anterior 
tongue of the New Mexico species is more strongly 
geniculated, the costae are stronger on fold and sulcus, 
and the dorsal valve is nearly hemispherical. No other 
species of Leiorhynchus known to us or in the Na- 
tional Collections is like this one. 

Leiorhynchus sphaericum is rare. Most of the spec- 
imens in the collection come from Mud Springs Moun- 
tains, but a single specimen was found in Rhodes Can- 
yon and two were taken in the Caballos Mountains, 
all from the shaly facies of the Onate. 


Genus PLATYTERORHYNCHUS Sartenaer, 1970 


Platyterorhynchus obscurum new species 
Plate 20, figures 39-48 


Large, roundly subpentagonal in outline; sides 
rounded; cardinal extremities rounded; anterior mar- 
gin truncated; apical angle 123°. Anterior commissure 
strongly uniplicate. Beak small, strongly incurved and 
pressed to dorsal umbo. Dorsal valve deeper than ven- 
tral one. Posterior half of exterior smooth or nearly 
sO, anterior irregularly costate, fold having four to six 
costae and sulcus five; flanks usually without costae. 

Ventral valve moderately convex in lateral profile, 
broadly and gently convex in anterior profile. Poste- 
rior and posterolateral areas strongly swollen; sulcus 
originating abruptly at about midvalve, wide and shal- 
low, occupying two-thirds the valve width; flanks 
bounding sulcus gently convex and only slightly raised 
above sulcus; tongue long and broadly rounded. 

Dorsal valve gently convex in lateral profile with the 
most curvature in the umbonal region. Anterior profile 
strongly domed, flanks gently convex and very steep. 
Umbonal region swollen; umbolateral slopes steep; fold 
originating at midvalve, elevating moderately ante- 
riorly, where it is fairly strong at front margin. 


BULLETIN 315 


Ventral valve interior having small teeth, obsoles- 
cent dental plates and narrowly triangular muscle field 
set deeply into adventitious tissue. Dorsal valve inte- 
rior with strong, elevated socket ridges, thick outer 
hinge plates, strong, narrow, elevated crural bases and 
long slender distally expanded crura. Median septum 
varying from sharp and blade-like to thick and mas- 
sive. All hinge structures strengthened and softened 
in contour by adventitious shell. 

Measurements (in mm).—USNM 200877 (Holo- 
type): length (L) 28.6, dorsal valve length 26.2, max- 
imum width (W) 29.1, thickness 18.3, apical angle 123°, 
L/W ratio = 0.98. 

Diagnosis.—Large Platyterorhynchus having a cos- 
tate fold and sulcus but smooth flanks. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —5S22d, e, 1, j, m; 523s, u, w, x, y, Z; 524, 
a, b, g, h, j, k; 526h, n, 0, p, v; 528u; 529j, k, n, q, t, 
w; 3013; 3023c; 3025a; 3052a. 

Types.—Holotype: USNM 200877. Figured Para- 
types: USNM 200840; 200878a, d. Unfigured Para- 
types: USNM 200878b, c, e, f. 

Discussion.—This species is akin to a number of 
Camarotoechiidae Schuchert (1929) occurring in Mid- 
dle Devonian strata of the eastern United States. It is 
similar to some forms of *‘Camarotoechia”™’ multicos- 
tatum (Hall, 1860a) but that species, as its name im- 
plies, is strongly costate on fold, sulcus, and flanks. 
This is true also of ‘°C.’ laura (Billings, 1860), from 
Ontario, but that species is more triangular and also 
more costate than Platyterorhynchus obscurum. “*Ca- 
marotoechia’’ kelloggi (Hall, 1867), from Ohio, is a 
smaller and more costate species. Platyterorhynchus 
russelli (McLaren, 1962), type of the genus from the 
Waterways Formation in Alberta, is more strongly 
costate than P. obscurum. 

This species is common in the upper part of the 
Onate Formation in the Sacramento Mountains but is 
rare in the San Andres Range. 


Genus HYBORHYNCHELLA Cooper, 1955 


Hyborhynchella bransoni Cooper 
Plate 20, figures 33-38; plate 21, figures 14-31; 
plate 41, figures 1—7 


Hyborhynchella bransoni Cooper, 1955, p. 62, pl. 12, figs. 1-11, 
text-fig. 1A, 1-9. 


This is such a rare species that very few specimens 
have been collected. It is found most frequently in the 
Sacramento Mountains but it also has been taken in 
the Mud Springs and San Andres Mountains, where it 
is very rare indeed. 


Although serial sections were made of several spec- 
imens, little new information was obtained on the in- 
terior details of the genus. The new sections reveal the 
same adventitiously-thickened dorsal umbonal region 
and cryptic hinge plates. The crural bases are revealed 
as stout but no new information was obtained on the 
crura. 


Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —S06c, 1; 509g, h, j, w; 521c, d; 524m; 
3025a; 3054h. 

Types.—Holotype: USNM 123344b. Figured Para- 
types: USNM 123343a, c; 123344a, c, primary types 
figured in Cooper (1955). Unfigured Paratypes: USNM 
123343b, d-f; 123344d-j. Figured Hypotypes: USNM 
201062; 201063; 201064a; 201069. 


Genus PAURORHYNCHA Cooper, 1942 


Paurorhyncha cooperi Stainbrook 
Plate 20, figure 49; plate 40, figure 40 


Camarotoechia (Plethorhyncha) endlichi (Meek) (part) Kindle, 1909, 
pl. 6, figs. 11, 11a; pl. 7, figs. 1, 2. 

Camarotoechia endlichi (Meek) Girty, 1900, p. 56, pl. 6, figs. 1-3; 
pl. 7, figs. la—e. 

Paurorhyncha sp. Cooper in Shimer and Shrock, 1944, pl. 119, figs. 
23-26. 

Paurorhycha cooperi Stainbrook, 1947, p. 315, pl. 47, figs. 1-5. 


This is one of the commonest and most distinctive 
of all the Percha (Box Member) brachiopods. It has 
been well described by Stainbrook but its interior de- 
tails are not well known. Fortunately, a few specimens 
in the National Collections show some of the details 
of this species. Other information is displayed in serial 
sections. 

The ventral valve has short and delicate dental plates 
that support small, sharp teeth. The ventral muscle 
field is somewhat longitudinally elliptical in outline with 
some thickening on the lateral margins. The diductor 
scars are small. 

The dorsal valve has a short but elevated median 
septum that supports a small, short and wide apical 
chamber, which is partially covered by narrow inner 
hinge plates. The outer hinge plates are narrow or non- 
existent, the crural bases being attached directly to the 
wall of the stout socket ridges. Crural bases thick, 
triangular in cross section; crura stout but their full 
extent not seen. 

Stratigraphic occurrence.—Percha Formation 
(Ready Pay Member; Box Member). 

Localities.—506, a, b, p, q; 507a, d, f, g, h, n, 0, t, 
Vv, W, X, y; 509b, d, e; 527a, d, e, g; USGS (Kirk) 460; 
461; 508; 509; 512; 3026, a; 3029; 3030; 303 1a, b; 3034; 
3042, s. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 83 


Types.—Holotype: M.A.S. 1603a (=USNM 135238). 
Figured Paratypes: M.A.S. 1603b—d (not in National 
Collection) primary types figured by Stainbrook (1947). 
Figured Specimens: USNM 108212a; 10821 1a. Figured 
Hypotypes: USNM 200815; 200816. 


Family YUNNANELLIDAE Rzhonsnitskaya, 1959 
Genus EOPARAPHORHYNCHUS Sartenaer, 196la 


Eoparaphorhynchus mclareni Sartenaer 
Plate 18, figures 28-30 


A single poorly preserved specimen is referred to 
this species. Most of the shell has been altered or de- 
stroyed, consequently fine radial lines are not visible 
on the surface. The specimen has a fold made up of 
three strong angular costae, and the flanks are marked 
by two costae. 

Measurements (in mm).—USNM 201091: length 
18.9, dorsal valve length 16.3, width 21.4, thickness 
13.0?, apical angle 104°. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—3029. 

Types.—Figured Hypotype: USNM 201091. 


Genus POROSTICTIA Cooper, 1955 


Porostictia perchaensis (Stainbrook) 


Pugnax pugnus Kindle (not Martin, 1809), 1909, p. 22, pl. 6, figs. 
3, 4-4b, 5, 6—6b (not figs. 7, 7a, 8, 8a). 

Paraphorhynchus perchaensis Stainbrook, 1947, p. 316, pl. 47, figs. 
6-8 (nor figs. 9-13). 

Porostictia perchaensis (Stainbrook) Cooper, 1955, p. 62, pl. 14, 
figs. 8-18, text-fig. 1C, 11-20. 


This species is commonest in the lower part of the 
Percha Formation (Box Member), especially in the vi- 
cinity of the Wilson Ranch, Hillsboro (15’) Quadrangle 
and east of Santa Rita. 

Localities.—506, a, b, q; 507a, b, h, q; 509a, c; 527c, 
d, i; 3026; 3026a; 3042, s. 

Types.—Holotype: M.A.S. 1596 (=USNM 155807) 
figured by Stainbrook (1947). Figured Specimens of 
Kindle (1909): Pugnax pugnus Kindle (not Martin, 
1809) (=USNM 62019a—-d). Figured Hypotypes of 
Cooper (1955): USNM 123394a; 123395a—c; 123396a. 


Genus EVANESCIROSTRUM Sartenaer, 1965 


Evanescirostrum glabraventrum (Stainbrook) 
Plate 19, figures 7-10 


Shumardella glabraventra Stainbrook, 1947, p. 317, pl. 47, figs. 14— 
18. 

Evanescirostrum glabraventra (Stainbrook) Johnson, Reso, and 
Stevens, 1969, p. 1357, pl. 158, figs. 1-15. 


This is an uncommon species in the Rhodes Canyon 
Formation, rather than the Percha Formation as orig- 
inally described by Stainbrook. The species has a stout 
median septum and short apical chamber that is cov- 
ered by thin inner hinge plates that in turn are contin- 
ued forward from the end of the chamber. The crura 
are long and flattened. 

Localities.—5240; 528c. 

Types.—Holotype: USNM 135258 (not figured 
herein). Figured Paratypes: USNM 135259a, b, spec- 
imens figured by Stainbrook (1947). 


Evanescirostrum species | 
Plate 19, figures 1-6 


About normal size for genus, wider than long and 
with widest part at midvalve. Fold consisting of three 
strong rounded costae developed somewhat posterior 
to midvalve. Fold moderately elevated anteriorly. Sul- 
cus wide, moderately deep, occupying about two-thirds 
of midwidth. Sulcus occupied by two strong costae. 

Interior of ventral valve with short dental plates. 
Interior of dorsal valve having a long slender median 
septum reaching nearly to midvalve and supporting a 
small apical chamber. 

Stratigraphic occurrence.—Thoroughgood Forma- 
tion. 

Locality.—S522. 

Types.—Figured Specimen: USNM 201042. 


PETASMARIA new genus 
[Gr. petasma = anything spread out] 


About medium-sized for a rhynchonellid, wider than 
long, outline widely elliptical; valves unequal in depth, 
dorsal valve having greater depth than ventral; ante- 
rior cOmmissure strongly uniplicate. Beak small, 
closely pressed onto dorsal umbo. Deltidial plates 
small, disjunct; foramen small, mesothyridid. Surface 
strongly costate, costae not reaching beaks, some cos- 
tae increasing by implantation and some by bifurca- 
tion; fine radial lines appearing on costae in anterior 
parts of shell. 

Ventral valve interior having strong, long dental 
plates. Dorsal valve interior with long median septum 
supporting a short, narrow, uncovered septalium. 
Socket ridges strong; outer hinge plates broad. Crura 
somewhat triangular in section. 

Type species.—Petasmaria patens n. sp. 

Diagnosis.—Large Yunnanellidae having a widely 
elliptical outline and multicostate shell. 

Comparison.—Petasmaria is somewhat suggestive 
of Rugaltarostrum Sartenaer (1961a) internally, but it 
differs in exterior and some interior characters. It has 


84 BULLETIN 315 


more strongly costate shells, especially on the flanks, 
whereas Rugaltarostrum generally has smooth flanks 
or only slight costation; Petasmaria is generally a more 
strongly folded shell and has a narrower fold and sul- 
cus. Inside the dorsal valve, the outer hinge plates are 
wider than those of Rugaltarostrum and the septalium 
is short, small and shallow. 

Petasmaria is wider and more strongly uniplicate 
than Eoparaphorhynchus Sartenaer (1961a), which 
usually has a much narrower and more triangular shell 
than species of the Percha genus. Some difficulty may 
be experienced in separating Porostictia Cooper (1955) 
from Petasmaria. The two occur together but, in ad- 
dition to its punctate ornament, Porostictia is a squar- 
ish rhynchonellid and is not widely extended, as is 
Petasmaria. There is external resemblance of the Mis- 
sissippian Pleuropugnoides Ferguson (1966) to Petas- 
maria. The Devonian genus however, possesses a 
strong median septum, a structure not found in Pleu- 
ropugnoides. 


Petasmaria compacta new species 
Plate 20, figures 22-32 


Small, transversely elliptical to subpentagonal in 
outline; sides narrowly rounded; anterior margin trun- 
cated. Anterior commissure strongly uniplicate. Beak 
small, closely pressed onto the dorsal valve umbo. 
Foramen small, mesothyridid?. Beaks smooth. Apical 
angle about 120°. Surface, except for beaks, costate; 
costae few, wide and subangular, two to four on fold, 
one to three in sulcus; flanks marked by two strong 
and one incipient costa. Fine radial capillae on surface 
where exfoliated. 

Ventral valve gently convex in lateral profile, deeply 
concave in anterior profile, with the anterolateral ex- 
tremities rising strongly on each side. Sulcus deep and 
short, originating four mm anterior to the beak and 
extended as long, narrowly-rounded, strongly serrate 
tongue. Flanks narrow, flattened. Fold steep-sided, in 
some specimens bearing one costa on each slope. 

Dorsal valve much deeper than ventral valve, mod- 
erately convex in lateral profile, and strongly trilobed 
in anterior profile; fold narrowly rounded, originating 
just posterior to midvalve; umbonal region having a 
shallow, narrow sulcus from which a bifurcating costa 
arises to form the principal two costae of the fold. 

Ventral valve interior with dental plates; dorsal valve 
interior having a strong median septum. 


Measurements (in mm).— 


dorsal maxi- 
valve mum _ thick- apical 
length length width ness angle height 


USNM 200942a 


(Holotype; 

loc. 5070) 10.7 LOLO 132957820 121° 10.7 
USNM 200942b 10.3 ey ust, Aci 127° 9.5 
USNM 200942c 12.4 1S 1357/8 118° 9.8 
USNM 200942d 1154? 20:0 «1458; 785 118° 8.2 


Diagnosis.—Small Petasmaria having few, strong 
costae. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—S07o. 

Types.—Holotype: USNM 200942a. Figured Para- 
type: USNM 200942b. Unfigured Paratypes: USNM 
200942c-g. 

Comparison.—Although this species is ornamented 
unlike P. patens n. sp. and P. multicostata n. sp., it 
has the general outline and fold and sulcus develop- 
ment of Petasmaria. In addition, it has the capillae on 
exfoliated areas shown in the larger species of the ge- 
nus. The species is rare and was found at only one 
locality. 


Petasmaria multicostata new species 
Plate 21, figures 1-5 


About normal size for genus, wider than long, widely 
elliptical in outline. Sides narrowly rounded, anterior 
truncated. Anterior commissure strongly uniplicate. 
Ventral beak small, not protuberant. Apical angle 141°. 
Beaks smooth. Surface, except for beaks, costate; 
costae narrowly subangular, increasing by bifurcation 
and implantation. Strong concentric wrinkles undulat- 
ing over costae on flanks. 

Ventral valve gently convex in lateral profile but 
deeply concave when viewed from anterior. Sulcus 
deep and wide, originating about five mm anterior to 
beak, and extended into long, rounded and distally 
serrate tongue. Flanks angular and extended strongly 
beyond sulcus. Eight costae occupying the sulcus, the 
median one largest and widest. Flanks marked by five 
costae. 

Dorsal valve triangular in lateral profile and forming 
an angle of about 50°. Anterior profile strongly tri- 
lobed, tongue and fold forming a large median lobe; 
rounded, moderately convex flanks forming the small- 
er lateral lobes. Fold elongated, moderately rounded 
in transverse section, occupied by nine costae, the 


DEVONIAN BRACHIOPODS OF NEw MEXICO: COOPER AND DUTRO 85 


median two bifurcating on umbo and matching median 
one of tongue. Flanks marked by five costae and an 
incipient sixth. Umbonal region with a shallow narrow 
sulcus just posterior to median costae. 

Measurements (in mm).—USNM 200940 (Holo- 
type): length 11.0, dorsal valve length 11.4, maximum 
width 20.2, thickness 14.6, apical angle 141°, height 
15.8. 

Diagnosis.—Petasmaria having exaggerated ventral 
tongue, numerous costae having concentric wrinkles, 
and length about half the width. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—506b; 509e. 

Types.—Holotype: USNM 200940. 

Discussion.—This species differs from P. patens n. 
sp., in its anterioposterior compressed form, narrowly 
elliptical outline, exaggerated fold and sulcus, and nu- 
merous costae. In addition to the concentric markings, 
exfoliated patches show some fine radial capillae. 


Petasmaria patens new species 
Plate 20, figures 1-21; plate 21, figures 6-13; plate 42, 
figures 27-32; plate 44, figures 1-10 


Pugnax pugnus Kindle (not Martin, 1809) 1909, p. 22, pl. 6, figs. 8, 
8a (only). 

Paraphorhynchus perchaensis Stainbrook (part), 1947, p. 316, pl. 
321, figs. 9-13 [not figs. 6-9 = Porostictia perchaensis (Stain- 
brook)]. 


A fairly large rhynchonellid, elliptical in outline, 
wider than long; sides narrowly rounded; anterior 
margin broadly rounded to truncated. Posterior margin 
broadly obtuse and forming angle of 130-145° in adult. 
Anterior commissure strongly uniplicate. Ventral beak 
small, low, closely pressed onto dorsal umbo in large 
adults. Foramen small, mesothyridid?; deltidial plates 
small, disjunct. Umbo narrowly convex to subcari- 
nate. Valves costate; costae narrowly rounded to sub- 
angular, originating just anterior to umbones, which 
are smooth. Costae variable, bifurcating and implant- 
ed; sulcus with three to seven, fold with three to ten 
costae, some formed on slope of fold not meeting the 
sulcus. Flanks usually with three or four strong costae. 
Fine radial lines often present on the flanks, especially 
posterolaterally. 

Ventral valve flatly convex in lateral profile; anterior 
profile deeply concave with the flanks strongly ele- 
vated; flanks narrow, flat to slightly convex. Sulcus 
originating just anterior to umbo (three to four mm 
measured on the surface), widening very rapidly and 
forming a long, narrow strongly serrated tongue. Sides 
of sulcus steep. 


Dorsal valve moderately convex in lateral profile but 
strongly domed in anterior profile, the dome having 
an elevated, narrow median lobe and moderately con- 
vex lobes on its sides. Umbo concave. Fold originating 
about three mm anterior to beak, usually with three 
costae, the outer two often bifurcating or trifurcating; 
development of costae on the fold is quite variable. 
Costae on flanks broad and strongly curved to the mar- 
gins. 

Ventral valve interior with strong and long dental 
plates. Dorsal valve interior with divided hinge plate 
having slender but strong socket ridges, wide outer 
hinge plates and short, shallow septalium supported 
by a long median septum. Crura triangular in section. 

Measurements (in mm).— 


dorsal 
valve thick- apical 
length length width ness angle 


USNM 200939 

(loc. 5070) 16.3 15.9 24.1 18.0 1835 
USNM 200943a 

(loc. 507d) 15.1 14.6 23.3 15.8 133° 
USNM 201186 

(loc. 507n) 16.0 IS\o7/ 28.3 13.6? 148° 
USNM 200938 

(loc. 507c) 15.4 16.0 23.8 19.7 130° 
USNM 200937 

(Holotype; loc. 507h) 14.6 14.6 23.7 23.1 140° 
USNM 200941 

(loc. 509) 11.2 11.0 16.2 8.1 122° 


Diagnosis.—Large, wide Petasmaria, with extend- 
ed fold bearing four to five costae. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, a, b, p, q; 507a, b, c, d, h, k, 1, m, 
Ms Oy Gly It X85 WR OS, SUPE lo, G3 SHA, Gl, Gp ti, In, mn, 
p; 3026, a; 3029; 3030; 303 1a; 3034, 3035; 3042, s; USGS 
(Kirk) 460; 461; 508; 509; 512. 

Types.—Holotype: USNM 200937. Figured Para- 
types: USNM 200938, 200939, 200941, 200943a, b. Un- 
figured Paratypes: USNM 200943c, 201185, 201186. 
Stainbrook (1947) figured specimens: M.A.S. 1596— 
155808, 155809. Kindle (1909) figured specimen: USNM 
62019e. 

Comparison.—Petasmaria patens n. sp. differs from 
Porostictia perchaensis (Stainbrook, 1947), with which 
it occurs and with which it has been confused, in lack- 
ing the rows of pores so characteristic of Porostictia 
Cooper (1955). It also differs in having an entirely dif- 
ferent shape, being wider and with a more swollen and 
elevated median fold. Petasmaria multicostata n. sp. 
is similar to P. patens in shape but differs in the finer 
and more numerous costae. 


BULLETIN 315 


Discussion.—The ornament of this species is highly 
variable as noted above. The flanks of the shell are 
fairly uniform in most specimens but the fold and sul- 
cus are highly variable. The few young specimens 
available have a carinate ventral umbo, the carina ex- 
tending anteriorly as a median costa in the sulcus. This 
carina corresponds to the sulcus in the dorsal umbo. 
Usually one or more costae are implanted in the sulcus 
on either side of the median rib. On the fold there are 
usually two primary costae that soon divide. In some 
specimens these further divide or intercalation takes 
place to produce a multicostate fold. Occasional spec- 
imens deviate so strongly in their costation as to sug- 
gest other species, but the specimens are too few to 
establish this as a fact. 


Order SPIRIFERIDA Waagen, 1883 
Superfamily ATRYPACEA Gill, 1871 
Suborder ATRYPIDINA Moore, 1952 
Family ATRYPIDAE Gill, 1871 
Subfamily ATRYPINAE Gill, 1871 
Genus ATRYPARIA Copper, 1966 


Atryparia rubra new species 
Plate 21, figures 38-53 


Large, subquadrate in outline, length and width 
nearly equal but usually slightly wider than long; sides 
rounded posteriorly but somewhat oblique anteriorly; 
anterior margin narrowly truncated to narrowly na- 
sute. Anterior commissure strongly and commonly 
narrowly uniplicate. Beak small, slightly or not at all 
protuberant beyond the posterior margin. Surface 
strongly costate, four costae in five mm at the front, 
increasing by bifurcation and implantation. Frill about 
eight mm wide. 

Ventral valve lateral profile unequal, posterior part 
gently convex, anterior part geniculated gently to 
strongly toward dorsal valve. Anterior profile flatly 
convex to broadly concave depending on the degree 
of deflection of lateral margins. Posterior moderately 
and somewhat narrowly swollen to midvalve; sulcus 
originating anterior to midvalve, variable from broad 
and shallow to narrow and fairly deep. Frilled margins 
deflected toward ventral side in narrow band. 

Dorsal valve strongly convex in lateral profile and 
most convex posteriorly; anterior profile strongly and 
fairly narrowly domed, sides gently convex and steep. 
Median region greatly swollen, swelling extending to 
the anterior where it forms a short fairly narrow fold. 


Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- frill 
length length width width ness length 


USNM 201005a 


(loc. 5230) Sl SHO a) 
USNM 201005b 26:8) 26:9) 21-6) 30!8) 16:6) a 
USNM 201021 

(Holotype: 

loc. 5230) 34:0) 3339" 2710) 3724. 1G Geeese0 
USNM 201014a 

(loc. 5230) 21:8 2722) 19385" 2/63 
USNM 201014b 273) 27/3. 923 29 LOL OOO 
USNM 201014c 24.1 24.0 20.0 29.6 140 7.0 
USNM 201014d 25:0 24:0! 216 27/7) IBA e0 
USNM 201014e 28:2, 27-7 2610) 3220 Omen 


Diagnosis.—Large frilled Atryparia having four 
costae in five mm at the anterior. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—522f, 1, t, v; 523a, 0; 5241; 3025Sa. 

Types.—Holotype: USNM 201021. Figured Para- 
types: USNM 201005a, b. Unfigured Paratypes: USNM 
201014a-e. 

Discussion.—This species 1s similar in size and out- 
line to Desquamatia costata n. sp., but differs in hav- 
ing stronger costae, greater development of frills, and 
less convex dorsal valve. This species occurs chiefly 
in the Caballos Range, especially at the Nackaye *‘Paint 
Mine”’ locality (5230). 


Genus DESQUAMATIA Alekseeva, 1960 


Desquamatia costata new species 
Plate 22, figures 17-38 


Large, subcircular in outline, length and width sub- 
equal; strongly inequivalve; sides rounded, anterior 
margin truncated to subnasute. Cardinal extremities 
rounded. Anterior commissure narrowly uniplicate. 
Beak and foramen small. Beak slightly protuberant; 
deltidial plates visible. Hinge narrower than maximum 
width, which is at midvalve. Surface costate, costae 
narrowly rounded, generally wider than the inter- 
spaces; imbrications distant on shell body but closely 
crowded at anterior. Five to seven costae in five mm 
at the front margin. 

Ventral valve unevenly convex in lateral profile, 
posterior half gently convex but anterior half flattened. 
Anterior profile variable from gently concave to gently 
convex. Beak narrowly swollen; umbonal region mod- 
erately swollen to midvalve; flanks flattened and be- 
coming slightly deflected in ventral direction near mar- 
gins. Sulcus wide, developing gradually near midvalve 
but not becoming pronounced until near the anterior 
margin; tongue moderately long, narrowly to broadly 
rounded. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 87 


Dorsal valve strongly convex in lateral profile, 
strongly domed in anterior profile, sides gently convex 
or flat and steeply dipping to margins. Umbonal and 
median regions greatly swollen; slopes to cardinal ex- 
tremities concave. Fold formed as a wave in anterior 
commissure, poorly distinguishable on valve surface 
except at very anterior where it may be slightly nasute. 

Measurements (in mm).— 


maxi- 

dorsal mum 
length valve hinge width thick- L/W 
(L) length width (W)~ ness _ ratio 


USNM 200973 


(loc. 522g) Bile AOS) BSS), 18.2 0.90 
USNM 200972 
(Holotype; 
loc. 523d) 2919 283 18.8 33.5 17.8 0.89 
USNM 201007a 
(loc. 5230) 2StOn 20r25 22 ON Sk 2 5 19450191 
USNM 201114 
(loc. 523h) 26 See Ol 7402910), 82901911 
USNM 201006 
(loc. 528e) BDI 25 4 OF S3520 26:45 0L98: 


Diagnosis.—Large finely costate Desquamatia, with 
five costae per five mm at front of valve, and shallow 
anterior sulcus. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —S06d; 522f, g, 1, 0; 523b, d, e, f, g, i, j, 
k, 0, t, u, x, z; 524b, d, 1; 526c, h; 528e; 529j, k, 0; 
3025; 3054f. 

Types.—Holotype: USNM 200972. Figured Para- 
types: USNM 200973, 201003, 201006, 201007a. Un- 
figured Paratypes: USNM 201007b, 200909. 

Comparison.—The distinction of this species from 
Atryparia rubra n. sp. is mentioned under that species. 
Desquamatia costata n. sp. is unlike any of the species 
from the Cedar Valley Formation assigned to “‘Atry- 
pa’ by Stainbrook (1938). Several undescribed atry- 
pas from Michigan and New York seem referable to 
Desquamatia, but are not identifiable with the Onate 
species. Atrypa dignata Fenton and Fenton (1930) re- 
sembles the Onate species but is larger, with a more 
convex dorsal valve, stronger fold and sulcus, and more 
protuberant beak. 


Genus PSEUDOATRYPA Copper, 1973 


Pseudoatrypa devoniana (Webster) 
Plate 22, figures 1-16; plate 23, figures 32-38 


Atrypa devoniana Webster, 1921, p. 19 (not pl. 8, figs. 9-11); Fenton 
and Fenton, 1924, p. 134, pl. 26, figs. 16-24; Stainbrook, 1938, p. 
240; 1945, p. 47, pl. 5, figs. 24-26, 29-32, fig. 1 [1]; 1948, p. 776, 
pl. 2, figs. 23, 24, 48. 


This species can be readily recognized in a collec- 
tion from the Sly Gap Formation by the relatively fine 


ornament. As pointed out by Stainbrook (1945), it is 
variable, not only as to size and shape but in devel- 
opment of fold and sulcus and in ornament. Much of 
the material from the Sly Gap is so badly weathered 
that the ornament is difficult to compare with well- 
preserved specimens. Most of the material consists of 
very costellate young specimens; adults are rare and 
often variable. The sulcus at the anterior is usually 
fairly wide, occupying nearly one-third of the valve 
width. No sharply folded specimens, as occasionally 
occur in large collections from the Hackberry (Cerro 
Gordo Member) Formation, were seen. Occasional 
specimens preserve the frill, which is narrow, about 
seven mm wide on a specimen 21 mm in length. 
Measurements (in mm).— 


dorsal maxi- 
valve hinge mum thick- 
length length width width ness 


USNM 200965 


(loc. 509s) 21.0 20.0 9.1 21.6 12.7 
USNM 200964 

(loc. 509z) 21.3 20.5 9.9 21.8 13.8 
USNM 201018 

(loc. 506) 25.6 25.2 13.0 26.0 17.6 
USNM 200927 

(loc. 521m) 17.3 16.4 11.8 20.0 10.1 
USNM 200944 

(loc. 521d) 26.0 25.0 16.9 26.6 IN7/55) 


Diagnosis.—Medium-sized, fine-lined Pseudoatry- 
pa with moderately strong anterior fold. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, j, v; 509g, h, i, n, r, s, y, z; 521c, 
d, e, k, m, q, s, t, u, w, y; 523m, n, u, w; 526k, s; 
529v; 3054f; USGS (Kirk) 538. 

Types.—Figured Hypotypes: USNM 200927, 200944, 
200964, 200965, 200966, 201094. 


Genus COSTATRYPA Copper, 1973 


Costatrypa extensa new species 
Plate 23, figures 39-56; plate 24, figures 1-12 


About medium in size for genus, subrectangular to 
semielliptical in outline; hinge wide but variable, equal 
to or wider than midvalve but usually slightly narrow- 
er; sides nearly straight to oblique; anterior broadly 
rounded. Beak and foramen small; valves shallow and 
shell compressed. Valves costate, costae fine and sep- 
arated by spaces about equal in width to width of cos- 
tae or wider; four to six costae in five mm at midvalve 
and the anterior margin. Entire surface covered by 
lamellae, distant posteriorly but becoming crowded to- 
ward the anterior. Frills narrow but attaining a width 
of nine mm and having coarser costae than the shell 
body. 


88 BULLETIN 315 


Ventral valve moderately convex in lateral profile, 
most convex posteriorly and with anterior gently ge- 
niculated for form small rounded tongue. Anterior pro- 
file broadly concave when frilled but moderately con- 
vex when frill is lacking, with median region forming 
small mound. Median region from beak to midvalve 
slightly swollen but from there to anterior depressed 
to form broad but shallow sulcus. 

Dorsal valve fairly strongly and evenly convex in 
lateral profile, broadly domed in anterior profile, and 
having fairly steep sides that become concave near 
margins. Median region strongly swollen and ante- 
riorly forming a poorly defined fold, best seen by view- 
ing anterior commissure. 

Ventral valve interior with a narrow, small, oval 
deeply impressed muscle area. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum_ thick- frill 


length length width width ness length 

USNM 200984 

(Holotype; 

loc. 521k) 19.4 18.2 23.5 27.2 1010) 824. 
USNM 200993 

(loc. 521k) 16.6 16.0 20.5 24.0 9:2) 6:6 
USNM 200983 

(loc. 521k) KeyS soy ep 90 — 
USNM 200967 

(loc. 506j) 14.6 13.7 18.0 23102 ele eT, 
USNM 200960a 

(loc. 521d) 14.7 14.2 14.62 18.2 85 — 
USNM 200960b 

(loc. 521d) 14.6 14.2 23.0* 22.0 Weil —"Wotl 
USNM 20095la 

(loc. 3054) 14.0 13.5 18.0 19.6 Sole 5-0 
USNM 200951b 

(loc. 3054) 14.6 13.7 24.0* 24.0 Ul 3H 


Diagnosis.—Shallow, compressed, finely costate 
Costatrypa having a short wide anterior sulcus on ven- 
tral valve and with fairly wide marginal frill. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S506c, j; 509f, p; 521d, e, k, m, u, w; 
524m, v; 526s; 529y; 3054, c; USGS (Kirk) 538. 

Types.—Holotype: USNM 200984. Figured Para- 
types: USNM 20095la, b, 200960a, 200967, 200983, 
200993. Unfigured Paratype: USNM 200960b. 

Discussion.—This species is distinguished from all 
others in the Sly Gap Formation by its transverse out- 
line, shallow valves, broad, short sulcus and ill-defined 
dorsal valve fold. The species is distinctive, but is not 
always easy to separate from young specimens of C. 
varicostata (Stainbrook, 1945). 

The development of the ornament on this species 
and on C. varicostata, as well as on Spinatrypa Stain- 
brook (1951), is similar. The beak of the ventral valve 


starts with a single costa that bifurcates almost im- 
mediately on appearance, and the beak of the dorsal 
valve starts with two costae that form a shallow sulcus 
in which a single costa is implanted not far anterior to 
the beak. All of the costae may bifurcate and some 
costae are implanted. 


Costatrypa varicostata (Stainbrook) 
Plate 23, figures 7-31 
Atrypa varicostata Stainbrook, 1945, p. 47, pl. 5, figs. 13-17. 
Atrypa variacostata [sic] Stainbrook, 1948, p. 776, pl. 2, figs. 44— 

47. 

This and C. extensa n. sp. are the commonest and 
perhaps most distinctive brachiopods in the Sly Gap 
Formation. They occur in countless numbers in the 
exposures on Sheep Mountain. The large quantities of 
specimens available show them to belong to an ex- 
tremely variable species. The distinctly shouldered 
posterior margin of C. varicostata, and ornament 
coarser than that of Pseudoatrypa devoniana, serves 
to separate these two species. 

The valves of C. varicostata are unequal in depth. 
The ventral valve is the shallower and is marked by a 
pronounced fold that originates anterior to midvalve 
and varies from fairly wide to narrow and produces a 
tongue with angular to narrowly rounded extremity. 
The dorsal valve fold originates near midvalve and 
varies from narrowly rounded to moderately rounded. 

The ornament of the pedicle valve in C. varicostata 
begins with a single costa at the beak that immediately 
divides, and between the resulting two another is in- 
serted. All of the costae are divided two or three times 
to the margin. The beak of the dorsal valve has an 
initial sulcus in which a costa is inserted. Costae are 
increased by insertion and division as on the ventral 
valve. The valves are covered by imbricating, closely- 
spaced lamellae that become crowded anteriorly. Frills 
narrow. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- frill 
length length width width ness length 


USNM 200992 

(loc. 506c) 16.2 15.8 7.8 19.4 14.2 — 
USNM 200963 

(loc. 3054b) Psy 1G )5) 10.5 24.2 13.8 — 
USNM 200996 

(loc. 521d) 17.6 16.4 9.0 17.9 14.2 
USNM 200979a 

(loc. 3054) 16.2 15:6) 135359 20:0) leo en 
USNM 200979b 

(loc. 3054) 18.3 W726) 7208 2225 13.0 — 


Diagnosis.—Compact, moderately strongly costate 
Costatrypa having fold and sulcus anterior to mid- 
valve. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 89 


Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, e, j, y; 509f, g, i, p, v, z; 521d, 
e, k, 1, m, q, t, u, v, w, x; 522k, x; 523m; 524m, n, v, 
w; 526d, e, s; 528p; 529p, u, y; 3054b, 1; USGS (Kirk) 
534; 538. 

Types.—Figured Hypotypes: USNM 200963, 
200979a, b, 200992, 200994, 200996, 201004. 

Discussion.—See under Costatrypa extensa n. sp. 


Genus IOWATRYPA Copper, 1973 


Iowatrypa rara new species 
Plate 24, figures 15—29 


Small, generally slightly longer than wide; unequally 
biconvex, the ventral valve having deeper shell in 
adults. Outline subrectangular with subparallel sides, 
obtusely rounded cardinal extremities and rounded an- 
terolateral extremities and truncate anterior margin. 
Anterior commissure gently and narrowly uniplicate. 
Hinge narrower than maximum width, which is just 
anterior to cardinal extremities. Posterolateral margins 
having an angle of about 120°. Interarea short and 
curved; beak small, incurved; foramen small; deltidial 
plates conjunct. Surface as in Atrypa Dalman (1828), 
costellate, costellae increasing by bifurcation; over- 
lapping lamellae distant posteriorly, crowded at ante- 
rior. 

Ventral valve fairly strongly convex in lateral pro- 
file, narrowly domed and with steep sides in anterior 
profile. Umbonal and median regions strongly inflated; 
anterior quarter depressed to form narrow, shallow 
sulcus that creates short rounded tongue. 

Dorsal valve moderately convex in lateral profile, 
less so than the dorsal valve. Anterior profile a mod- 
erately convex dome with flattened, moderately steep 
slopes. Umbonal region only moderately inflated but 
median region strongly swollen, swelling continuing to 
anterior margin to form poorly defined fold. 

Interior not known. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- apical 
length length width width ness _ angle 


USNM 201099 
(loc. 506j) 14.2 12.6 9.5 13.0 9.8 119° 
USNM 201098 
(Holotype; 
loc. 521q) [4a IStOn LO ii 122955 1020) PAP 
USNM 203507 
(loc. 521k) tile72, aK)? dts iio 610) 138i 


Diagnosis.—Fairly strongly costellate /Jowatrypa 
with length greater than, or equal to, the width. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506j; 509r; 521d, k, q. 


Types.—Holotype: USNM 201098. Figured Para- 
types: USNM 201099, 201135a. Unfigured Paratypes: 
USNM 203507, 203508, 201135b. 

Discussion.—This species resembles /. americana 
(Stainbrook, 1945) from the Independence Formation 
and J. owenensis (Webster, 1921) from the Owen 
Member of the Hackberry Formation. The New Mex- 
icO species is ornamented like the former but, accord- 
ing to Stainbrook (1945, p. 52), that species is typically 
slightly wider than long, whereas the reverse is true 
of J. rara. Webster’s species is much longer than wide 
and has a stronger and narrower fold and sulcus than 
I. rara. A species very similar to J. rara occurs in the 
Mt. Hawk Formation of western Canada (Warren and 
Stelck, 1956, pl. 17, figs. 19-24; called Gruenwaldtia 


Sp.). 


Iowatrypa americana (Stainbrook) 
Plate 24, figures 13, 14 


Gruenwaldtia americana Stainbrook, 1945, p. 52, pl. 5, figs. 18-23, 
27, 28; fig. 1 [6]. 


Two views of this species are introduced for com- 
parison with the New Mexico species. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—528f. 

Types.—Figured Hypotype: USNM 201134a. 


Genus SPINATRYPA Stainbrook, 1951 


Spinatrypa compacta new species 
Plate 24, figures 30-45 


Small, subquadrate in outline, length and width 
nearly equal; sides slightly convex to nearly straight 
and subparallel; anterior margin rounded. Anterior 
commissure broadly uniplicate. Beak small and closely 
appressed onto the dorsal umbo. Hinge wide, forming 
small flattened ears. Dorsal valve much deeper than 
ventral valve. Surface distantly costate, costae be- 
coming vague anteriorly; ventral valve umbo with an 
initial costa bifurcating soon after formation, the two 
costae extending nearly to the anterior margin and 
forming a narrow, low fold; umbonal costae flanking 
initial costae occupying the flanks. Dorsal valve umbo 
sulcate; sulcus occupied by a median costa about three 
mm anterior to the beak, it and the flanking costae 
extending to the anterior; eight or nine costae in all. 
Entire surface covered by distant concentric lamellae. 

Ventral valve gently convex in lateral profile, broad- 
ly and flatly convex in anterior profile; sulcus poorly 
developed, defined only at anterior third. Flanks gently 
convex. 


BULLETIN 315 


Dorsal valve strongly convex in lateral profile and 
narrowly domed in anterior profile, sides of dome 
nearly vertical. Fold defined only at commissure. Pos- 
terolateral slopes concave to small cardinal extremi- 
ties. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- 
length length width width ness 


USNM 203538 
(Holotype; loc. 528g) 
USNM 200978a 
(loc. 523n) 17.3 16.8 113357) 17.9 11.4 
USNM 200978b 17.4 16.3 16.2 18.8 13.0 


20.0 19.5 17.0 21.5 13.4 


Diagnosis.—Small, nearly square, paucicostate 
Spinatrypa having small flattened ears. 

Stratigraphic occurrence.—Contadero Formation 
(top); Sly Gap Formation? 

Localities. —S09g?; 522a; 523n; 528g. 

Types.—Holotype: USNM 203538. Figured Para- 
types: USNM 203540a, b. 

Discussion.—This species occurs in abundance at 
the top of the Contadero Formation at Sheep Moun- 
tain. It is smaller than either of the other New Mexico 
species and is more sparsely costate than either; its 
costae cover more of the valve than those of S. ob- 
solescens n. sp. It is a more rotund and thicker species 
than Spinatrypa rockfordensis (Fenton and Fenton, 
1924). It is suggestive of Spinatrypa mascula (Stain- 
brook, 1938) from the Cedar Valley Formation (Solon 
Member) but is smaller, has a straighter hinge, better 
defined ears, and anteriorly obsolescent costae. 


Spinatrypa obsolescens new species 
Plate 24, figures 46-60 


Medium-sized for genus, variable in outline, sub- 
rectangular to longitudinally oval; sides and anterior 
margin rounded; hinge narrower than widest part. Fold 
and sulcus moderately developed and, in some spec- 
imens, poorly defined. Surface partially costate, cos- 
tae well defined only in umbonal region, the ventral 
valve having a median costa that divides soon after 
appearance, and is matched by narrow sulcus on op- 
posite valve. Two costae on each side of median di- 
vided one on ventral umbo, making six on umbo. Dor- 
sal valve having median costa in early fold flanked by 
two or three costae on each side. Costae becoming 
vague or disappearing anterior to the umbonal region. 
Entire surface covered by strong concentric lamellae. 
No spines seen. 

Interior unknown. 


Measurements ( in mm).— 


dorsal maxi- 
valve hinge mum _ thick- 
length length width width ness 
USNM 200962 
(Holotype; loc. 526g) 21.5 21.0 20.0? 26.6 11.9 
USNM 200997d 
(loc. 3054) 28.7 VA) 17202 ees 13.6 
USNM 200997b 
(loc. 3054) 23.7 24.0 19:02 e27e5 13.5 
USNM 200997a 
(loc. 3054) 22.8 23.0 18.0? 24.6 12.1 


Diagnosis.—Spinatrypa having costae vague or ob- 
solescent anterior to the umbonal region. 

Stratigraphic occurrence.—Contadero Formation. 

Localities.—S09t; 521j; 522a; 524t; 526g; 3054k. 

Types.—Holotype: USNM 200962. Figured Para- 
types: USNM 200997a, d. Unfigured Paratypes: USNM 
200997b, c. 

Discussion.—In the umbonal region of both valves, 
this species is marked like that portion of Spinatrypa 
trulla decorticata n. ssp., but the anterior parts of S. 
obsolescens are devoid of costae or show only vague 
traces of them. The species is not so big as S. trulla 
decorticata and can be distinguished from that form 
by the lack of anterior costae. The loss of costae an- 
teriorly takes place in some specimens of S. trulla 
decorticata but usually the obsolescence has not pro- 
ceeded as far as in the Contadero species. It is likely 
that S. obsolescens is descended from the Sly Gap 
subspecies. 


Spinatrypa trulla (Stainbrook) 
Plate 25, figures 1, 2 


Figures of Spinatrypa trulla (Stainbrook, 1945) are 
introduced for comparison with the New Mexico sub- 
species. Spinatrypa trulla, one of the commoner 
species in the Independence Formation of Iowa, is 
usually exceptionally well preserved and some speci- 
mens exhibit the spines to perfection. 

Types.—Figured Hypotypes: USNM 201082a, b. 

Localities. —362a; 528h. 


Spinatrypa trulla decorticata new subspecies 
Plate 25, figures 3-40 


Large, variable in shape and ornament, usually wid- — 


er than long and subquadrate in outline; sides rounded; 
anterior margin rounded; hinge narrower than maxi- 
mum width, which is at about midvalve; beak small, 
low, usually damaged, and always closely pressed onto 
the dorsal valve umbo. Biconvex, dorsal valve swollen 
and much deeper than ventral valve. Surface ornament 
variable from strongly-defined to obsolescence of some 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 9] 


features. Valves costate; ventral umbo usually having 
one costa at beak dividing almost immediately and oc- 
cupying sulcus to the margin; median costa flanked at 
beak by two on each side, the inner two forming sides 
of sulcus but, like their companions on each side, often 
becoming vague and obsolescent. Some specimens with 
costae intercalated anterior of midvalve. Dorsal valve 
umbo marked by a median narrow sulcus correspond- 
ing to initial costa of opposite valve; sulcus occupied 
early in development by median costa that extends 
with two flanking ones to anterior margin to form the 
crest of median fold. Surface strongly lamellose, spines 
being produced at junction of costae and lamellae. 
Spines stout but rarely preserved. 

Ventral valve variable, fairly strongly to gently con- 
vex in lateral profile, broadly and gently concave to 
broadly and moderately convex in anterior profile. 
Sulcus usually broad and shallow, originating at about 
midvalve and producing a broadly rounded moderately 
long tongue. Flanks varying from gently concave to 
gently convex. 

Dorsal valve moderately to strongly convex in lat- 
eral profile and strongly domed with long sloping sides 
in anterior profile. Umbonal and median regions 
strongly swollen. Fold broad and low, originating at 
or near midvalve, seldom very prominent and strong 
only at anterior but pronounced in anterior view. 

Interior not seen. 

Measurements (in mm).— 


maxi- 

dorsal mum 
length valve hinge width thick- L/W 
(L) length width (W) ~ ness_ ratio 


Diagnosis.—Large Spinatrypa with variable orna- 
ment and strongly convex dorsal valve. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —506c, i, j, Vv; 507; 509f, h, i, j, n, p, q, 
Is Sy Va) Wa Xe Ya. Zs D2 cade es ke lems oF mts 523m: 
524m, n, u?; 526f, k, s; 529s; 3054. 

Types.—Holotype: USNM 200998. Figured Para- 
types: USNM 200931, 200982b, d, f, 200999, 201002, 
201159. Unfigured Paratypes: USNM 200961, 200982a, 
c, e, 201182a, 201183a, 201184. 

Discussion.—This is a quite variable subspecies and, 
when compared to a collection of specimens of Spin- 
atrypa trulla (Stainbrook, 1945), many differences may 
be noted in individual specimens. However, they all 
retain their resemblance to Stainbrook’s species. The 
Sly Gap subspecies attains a larger size than that from 
Iowa and is often rounded in outline; occasional spec- 
imens are elongated. The ornament is basically that of 
S. trulla but some specimens have intercalated costae 
more numerous than usual in the Iowa subspecies, in 
which the costae tend to be fairly uniform. Many of 
the New Mexico specimens (e.g., USNM 201159) show 
obsolescence of costae anteriorly and laterally, a phe- 
nomenon rarely seen in S. trulla. 

A few specimens of S. trulla decorticata vary from 
the main line in the development of the fold and sul- 
cus. Specimens from localities 5O9y, 5210 and 524u 
have an exaggerated fold and sulcus, the latter being 
narrow and very deep while the fold is also narrow 
and elevated from umbo to anterior margin. The crest 
of the fold is occupied by three costae, the median 
costa of the initial, juvenile sulcus and two of those 
flanking it on the umbo. Stainbrook (1945, pl. 5, figs. 


USNM 201182a 
(loc. 506i) 30.3 29.1 20.02 35.7 19:4 0.85 
USNM 201183a 30.4 29.8 22.0? 33.7 18.1 0.90 
USNM 200998 
(Holotype; 
loc. 509}) SEAS 30195237425 3887) 9920) 10393) 
USNM 201184 
(loc. S09y) 29:3 24:3) 2253 3055 *16:4) (0583 
USNM 200982b 
(loc. 506c) ZO TE Zee 92 SOLO 1620) 10298 
USNM 200982f 
(loc. 506c) Piles — PX) ag} Rad (ESS) 
USNM 200982d 
(loc. 506c) 14.6 13.9 8.7 15.8 7.1 — 
USNM 200982c 
(loc. 506c) P20) slit il 8.2 12.0 5.9 1.00 
USNM 200982e 
(loc. 506c) 10.0 9.4 6.4 10.0 5.3} — 
USNM 200961 
(loc. 506c) 7S PADS GBS COLESH/ 
USNM 200982a 
(loc. 506c) 3.3) 4.7 — Sel 2.0 = 


5-7) illustrates a specimen (M.A.S. 896; Iowa Univer- 
sity) from the Independence Formation showing the 
same aberrations as those shown by the Sly Gap in- 
dividual (USNM 200999) from locality 524u. This is 
the most exaggerated specimen seen. This latter spec- 
imen measures (in mm): length 23.9, dorsal valve length 
25.3, hinge width 22.0?, maximum width 33.4, and 
thickness 22.4. 


Suborder ATHYRIDIDINA Boucot, Johnson, and 
Staton, 1964 


Superfamily ATHYRIDACEA M’Coy, 1844 
Family ATHYRIDIDAE M Coy, 1844 
Subfamily ATHYRIDINAE M’Coy, 1844 
Genus ATHYRIS M’Coy, 1844 


92 BULLETIN 315 


Athyris transversa (Stainbrook) 


Athyris coloradensis Kindle (not Girty, 1900), 1909, p. 24, pl. 6, fig. 9. 

Cleiothyridina? coloradensis Stainbrook (not Girty, 1900), 1947, p. 
327, pl. 45, figs. 25-30. 

Cleiothyridina? transversa Stainbrook, 1947, p. 327, pl. 45, figs. 14, 
19-22. 


The frills of this species are like those of the more 
familiar Athyris lamellosa (Léveillé, 1835) of the Mis- 
sissippian. Like the Percha species, the frills of this 
large athyrid are made up of closely compacted spine- 
like fibers that often give the ends of the frills a some- 
what spinose appearance, but very minutely so and 
quite unlike Cleiothyridina Buckman (1906). Conse- 
quently, it seems necessary to assign the Percha species 
to Athyris M’Coy (1844) rather than Cleiothyridina. 
Stainbrook evidently had doubts about the assignment 
because he questioned his own use of Cleiothyridina. 

This genus is abundant and varied in the Percha 
Formation (Box Member). Generally, specimens that 
retain much of their frilly ornament seem transverse 
and are assignable to this species. Specimens shorn of 
their frills tend to be rather rounder and suggest C.? 
coloradensis, as interpreted by Stainbrook. It is sug- 
gested that only one species is involved and that re- 
tention or stripping off of the frills is the clue to their 
identification. 

Athyris coloradensis Girty (1900) from the Ouray 
Limestone is a much larger species than the Percha 
forms and cannot be so identified. Inasmuch as A. 
coloradensis is a very different species, and those 
specimens so assigned from the Percha can be iden- 
tified with A. transversa, the latter name is available 
for the Percha species. The name is not entirely an apt 
one because most of the specimens are not transverse 
when stripped of their frills. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—506, b, p; 507a, b, d, e, h, k, 1, m, 0, r, 
t, x; 509a, b, c, e; 527e, 1, p; 3035; 3042, s; USGS 
(Kirk) 461. 

Types.—Holotype: M.A.S. 1595A (=USNM 
155779). Paratype: M.A.S. 1595B (=USNM 155780). 
Figured specimen of Kindle (1909): USNM 62020. Fig- 
ured specimens of C.? coloradensis Stainbrook (not 
Girty) are not in the National Collections. 


CRINISARINA new genus 
{Gr. crinis = hair] 

Shells athyridid, attaining a width of about 25 mm 
or less; length and width subequal; outline subpen- 
tagonal; valves biconvex, subequal in depth. Beak 
small, erect, with moderately large, round foramen. 


Anterior commissure moderately to strongly unipli- 
cate. Surface strongly and closely lamellate, the la- 
mellae bearing a fringe of long, thin, solid spines. 

Ventral valve interior having long stout dental plates 
and thick teeth. Dorsal valve interior with stout nar- 
row hinge plate. 

Type species.—Cleiothyridina reticulata Stain- 
brook, 1947, p. 326, pl. 45, figs. 31-35. Holotype: 
USNM 135248. 

Diagnosis.—Compact, pentagonal athyridids having 
uniplicate anterior commissure and lamellae bearing 
fringes of long thin spines. 

Comparison.—tThe brachiopods referred here have 
been variously placed in Athyris and Cleiothyridina. 
They partake of the interior details of both genera but 
differ in their pentagonal outline, subequal depth, di- 
mensions and strong uniplication. 

Although several species are assigned here, seldom 
are specimens found that have the exterior details well 
preserved. Fortunately, a few specimens of each 
species show the cleiothyridid ornament. Although 
many specimens are weathered smooth, some show 
the concentric crowded lamellae but are completely 
stripped of spines. It is incredible that most specimens 
can be so completely shorn of their hairy spines. A 
few specimens preserve the spines in more or less good 
condition. On one such specimen [C. angelicoides 
(Merriam, 1940), USNM 205263a], they are about three 
mm long. 

The following five species are placed in Crini- 
sarina: Athyris angelicoides Merriam (1940) (USNM 
205263), Cleiothyridina devonica (Raymond, 1909), C. 
humerosa Stainbrook (1950), C. reticulata Stainbrook 
(1950), and C. prouti (Swallow, 1860). The first two 
are, respectively, from the “‘Upper Nevada Lime- 
stone’’ and Three Forks Formation; the third comes 
from the Famennian Aplington Formation of Iowa. 
Crinisarina reticulata is from the Rhodes Canyon For- 
mation, not the Percha as claimed by Stainbrook. 
Crinisarina prouti is from Early Mississippian rocks. 
All except C. devonica have the typical compact form, 
but the Three Forks specimens have a somewhat wid- 
er outline and less strongly developed fold and sulcus 
than usual in the other species. 


Crinisarina reticulata (Stainbrook) 
Plate 26, figures 31-51; plate 45, figures 1-15 


Cleiothyridina reticulata Stainbrook, 1947, p. 326, pl. 45, figs. 31— 
35; Johnson, Reso, and Stephens, 1969, p. 1362, pl. 159, figs. 1- 
26; text-fig. 5. 


Stainbrook described this species as coming from 
the Percha Formation a mile south of the east entrance 
to Rhodes Pass in the San Andres Range. The collec- 


| 
| 
| 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 93 


tions of the U.S. National Museum of Natural History 
include a fair number of specimens of this species but 
their provenance is the Rhodes Canyon Formation 
rather than the Percha, which was not identified in this 
range. The species is related to Crinisarina devonica 
(Raymond) from the Three Forks Formation of Mon- 
tana, but differs in having a more pronounced fold and 
sulcus, deeper valves that give a thicker lateral profile, 
and more strongly reticulate ornamentation; this was 
discussed in some detail by Johnson, Reso, and Ste- 
phens (1969, p. 1356). 

Diagnosis.—Crinisarina having a strong resem- 
blance to Athyris M’Coy (1844) in shape and profile, 
strongly reticulate ornament and strong fold and sul- 
cus. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation; Thoroughgood Formation. 

Localities.—Rhodes Canyon: 509t, u; 521f, g, h, i; 
523n, p, q, Vv; 524c, e, 0, z; 526a; 528c, j; 3054q. Tho- 
roughgood Formation: 522. 

Types.—Holotype: USNM 135248. Figured Para- 
type: M.A.S. 1588 (=USNM 135249), primary types 
figured by Stainbrook (1947). Figured Hypotypes: 
USNM 200925, 200974, 200928, 203539a, b. 

Discussion.—Four specimens from the lower foot 
of the Thoroughgood Formation are included here. 
These specimens are badly exfoliated but they have 
the characteristic size, form, fold and sulcus of C. 
reticulata. 


Genus COMPOSITA Brown, 1849 


Composita bellula Stainbrook 
Plate 26, figures 52-66; plate 45, figures 16-32 


Meristella barrisi? Kindle (not Hall, 1860a), 1909, p. 30, pl. 9, figs. 
7-8a. 

Dielasma coreiforme Stainbrook, 1947, p. 318, pl. 47, figs. 34-37. 

Composita bellula Stainbrook, 1947, p. 325, pl. 45, figs. 15-18, 23, 
24. 


This species is not a Meristella as Kindle thought 
but is the earliest species of Composita, a long line of 
brachiopods that extended into the Permian. The spire 
of C. bellula is similar to that of C. trinuclea (Hall, 
1856) of the late Mississippian but is unlike the jugum 
of C. subtilita (Hall, 1852) which is forked at the an- 
terior. The differences seem insufficient on which to 
base a genus. 

Stratigraphic occurrence. 
Member). 

Localities. —506, b, p; 507a, c, f, h, n, 0, z; 509a, 
c; 527d, f, h; 3026, a; 3029; 3030; 3031b; 3042s. 

Types.—Holotype: M.A.S. 1583A (=USNM 
155799); Figured Paratype: M.A.S. 1583B (=USNM 
155801). Unfigured Paratype: M.A.S. 1538B (=USNM 


Percha Formation (Box 


155800). Figured Specimens of Meristella barrisi? 
Kindle: USNM 62037a-c. Figured Hypotypes herein: 
USNM 201010, 201037, 201038b; 203527a, b. 


Suborder SPIRIFERIDINA Waagen, 1883 
Superfamily CYRTIACEA Fredericks, (1919) 1924 
Family AMBOCOELIIDAE George, 1931 
Genus AMBOCOELIA Hall, 1860a 


Ambocoelia capillata new species 
Plate 26, figures 19-24 


Two specimens of Ambocoelia were taken from the 
Percha Formation (Box Member) at localities S09b and 
3029. The ventral valve is moderately convex in lateral 
profile and narrowly domed in anterior view. A narrow 
sulcus originates at the beak and extends to the an- 
terior margin. An indistinct groove occurs on each flank 
and gives the appearance of the sulcus being bounded 
by two strong costae. The interarea is fairly long and 
strongly curved. The delthyrium is open. 

The dorsal valve is flatly convex in both profiles and 
is marked medially by a broad sulcus corresponding 
to the two costae of the ventral valve. The median 
part of the sulcus is occupied by a broad costa oppos- 
ing the median sulcus of the ventral valve. This ar- 
rangement produces a sulciplicate anterior commis- 
sure. 

The surface of both valves is covered by very fine 
capillae. No features of the interior could be deter- 
mined. 

Measurements (in mm).—USNM 201136 (Holo- 
type): length 5.0, dorsal valve length 4.1, width 4.8, 
thickness 3.8. 

Diagnosis.—Small Ambocoelia marked by fine cap- 
illae. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—S09b; 3029. 

Types.—Holotype: USNM 201136. 

Discussion.—This species is small for the genus and 
differs from described species by the presence on it of 
fine capillae. 


Ambocoelia species | 
Plate 26, figures 7-18 


About normal size for genus, longer than wide, sub- 
pentagonal in outline with hinge wider than valves at 
middle; sides rounded gently and sloping gently me- 
dially; cardinal extremities rounded; anterior margin 
narrowly rounded; posterolateral margins forming an 
angle of 85°. Anterior commissure faintly sulcate. In- 
terarea short, curved and apsacline. Delthyrium 


BULLETIN 315 


bounded by narrow, oblique deltidial plates. Surface 
smooth. 

Ventral valve strongly convex in lateral profile and 
narrowly domed in anterior profile; flanks slightly con- 
vex and steep. Umbonal and median regions strongly 
swollen; median region marked by a narrow sulcus 
extending from umbo to anterior margin. 

Dorsal valve gently but unevenly convex, most con- 
vex in the posterior third; anterior profile forming a 
flat triangle. Anterior margin deflected in a dorsal di- 
rection to form a rim around the anterior and sides, 
the ventral valve fitting over the deflected surface. 

Interior not known. 

Measurements (in mm).—USNM 201073a: length 
8.8, dorsal valve length 7.0, hinge width 7.3, midwidth 
8.0, thickness 5.8. 

Diagnosis.—Medium-sized, elongate Ambocoelia 
having a narrow shallow sulcus on the ventral valve, 
inwardly tapering sides and fairly wide hinge. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —S5061; 526k; 5291. 

Types.—Figured Specimens: USNM 201073a, b. 

Discussion.—These are the only specimens of Am- 
bocoelia taken from the Sly Gap Formation. They dif- 
fer in important respects from Ambocoelia sp. 2 in 
being smaller, with more prominent lateral shoulders, 
less rounded sides, and less anteriorly depressed dor- 
sal valve. These specimens suggest A. umbonata 
(Conrad, 1842) from the Hamilton Group of New York, 
but they are smaller than a well-formed adult of the 
latter species, have less angular sides, a more convex 
dorsal valve and less of a marginal depression in the 
dorsal valve. 


Ambocoelia species 2 
Plate 26, figures 25-30 


About normal size for genus, subpentagonal in out- 
line; length and width nearly equal; hinge slightly nar- 
rower than maximum width, which is just anterior to 
hinge. Sides moderately rounded and sloping gently 
medially; anterior margin narrowly rounded; anterior 
commissure broadly sulcate; posterolateral margins 
forming an angle of 83°; interarea short and strongly 
curved, apsacline. Beak strongly incurved. Surface 
smooth. 

Ventral valve gently convex in lateral profile, broad- 
ly domed in anterior profile, which has short, moder- 
ately steep sides. Umbonal and median regions swol- 
len; median region from beak to anterior margin marked 
by narrow, shallow sulcus. 

Dorsal valve, in lateral profile, with posterior half 
gently convex but anterior half gently concave; ante- 
rior profile broadly but very gently convex; median 


region moderately swollen; anterior margin fairly 
strongly deflected in a dorsad direction, margin form- 
ing rim around concave anterior part; median region 
marked by shallow, narrow sulcus extending from 
midvalve to anterior margin. 

Interior unknown. 

Measurements (in mm).—USNM 201074: length 9.7, 
dorsal valve length 7.3, hinge width 8.6, maximum 
width 9.8, thickness 5.7. 

Diagnosis.—Medium-sized Ambocoelia with mod- 
erately deep ventral valve, short strongly curved in- 
terarea, and anteriorly strongly concave dorsal valve. 

Stratigraphic occurrence.—Contadero Formation. 

Localities.—S09o0, t; 521); 5240; 526g; 3054k. 

Types.—Figured Specimen: USNM 201074. 

Discussion.—This species is abundant in the upper 
part of the Contadero Formation, in places forming a 
coquina consisting mostly of ventral valves. It is 
suggestive of Ambocoelia gregaria Hall (1860a) but 
appears to be wider and lacks the fairly strong ventral 
median sulcus of the New York Chemung species. It 
differs from A. umbonata (Conrad, 1842) of the New 
York Middle Devonian, a widely identified species, in 
having a more rounded outline with rounded cardinal 
extremities, a more convex dorsal valve and smaller 
size. This Contadero Ambocoelia is probably a new 
species but, though it is common in places, our ma- 
terial is inadequate in quality and quantity to provide 
a basis for a formal species description. 


Ambocoelia species 3 


Several poor specimens of a large Ambocoelia were 
taken from the basal foot of the Thoroughgood For- 
mation at locality 522. The species is longer than wide, 
with a long interarea and moderately incurved beak. 
The sulcus of the ventral valve is fairly strongly de- 
veloped. The dorsal valve is swollen posteriorly but 
shallowly concave at the anterior and has a reflected 
margin. 

Measurements of largest specimen (in mm).—USNM 
203509a: length 11.5, dorsal valve length 8.0, width 
10.0, thickness 6.7. 

Types.—Described Specimens: USNM 203509a-g. 


Ambocoelia species 4 


Ambocoelia is abundant in some beds of the Onate 
Formation as at Nackaye Mine above the Derry Hills 
and in San Andres Canyon. Unfortunately, the spec- 
imens are invariably crushed and none suitable for de- 
scription or illustration was found. An average speci- 
men measures about 10 mm in width and 7.5 mm in 
length. Other features are in accordance with the 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 95 


smooth shell and the anteriorly situated growth la- 
mellae usually seen in Ambocoelia. 
Localities.—522m, q; 523e; 5241, p, q, x; 526b; 5291, 
1, m, w. 
Types.—Measured Specimen: USNM 252627. 


Genus EMANUELLA Grabau, 1925 


Emanuella species 
Plate 26, figures 1-6 


About medium-sized for genus, slightly wider than 
long, subcircular in outline. Maximum width just an- 
terior to hinge; sides and anterior margin strongly 
rounded; posterolateral margins forming angle of 106°; 
anterior commissure rectimarginate. Interarea short, 
apsacline; beak strongly incurved, only moderately 
protruding beyond posterior margin. Surface smooth. 
Ventral valve fairly strongly convex in lateral profile 
with maximum convexity at midvalve; anterior profile 
somewhat narrowly domed and having flattened, 
steeply sloping flanks. Umbonal and median regions 
strongly swollen. 

Dorsal valve moderately convex in lateral profile 
and moderately and broadly convex in anterior profile. 
Umbonal and median regions swollen but anterior part 
somewhat flattened. Dorsal valve interior with short 
median septum. 

Measurements (in mm).—USNM 203559: length 
10.0, dorsal valve length 8.8, hinge width 5.5, maxi- 
mum width 10.3, thickness 6.4. 

Diagnosis.—Nearly circular Emanuella having 
strongly incurved beak and interarea, and both valves 
smooth. 

Stratigraphic occurrence. —Onate Formation. 

Localities.—522n; 5241, q, x; 526b; 528i; Lost Can- 
yon, Hermosa, New Mexico (R. Flower, collector). 

Types.—Figured Specimen: USNM 203559. 

Discussion.—Only one good specimen of this species 
was taken and it is assigned to Emanuella rather than 
to Crurithyris George (1931) because of its narrow 
hinge, strongly rounded sides, smoothly convex dorsal 
valve and strongly incurved beak. It suggests E. sub- 
umbona (Hall, 1857) from the top of the Hamilton and 
“E.”” meristoides (Meek, 1868) from the Middle De- 
vonian of the lower MacKenzie Valley, Canada, but 
is smaller and more transversely oval than either of 
these species. 


Genus ECHINOCOELIA Cooper and Williams, 1935 


Echinocoelia pretiosa new species 
Plate 27, figures 20-29 


About medium-sized for genus, length and width 


nearly equal; subpentagonal in outline, sides well- 
rounded; anterior strongly rounded; cardinal extrem- 
ities narrowly rounded; hinge narrower than maximum 
width, which is at about midvalve; posterolateral mar- 
gins forming angle of 87°. Anterior commissure recti- 
marginate. Beak fairly long and moderately protuber- 
ant beyond the posterior margin. Interarea long and 
moderately curved, strongly apsacline. Surface marked 
by fine concentric lines bearing fine spines, three or 
four concentric lines to one mm on the body of ventral 
valve. 

Ventral valve moderately convex in lateral profile 
and narrowly rounded at the umbo where the beak is 
moderately incurved; anterior profile narrowly domed 
with steeply sloping sides. Umbonal and median re- 
gions strongly swollen. Flanks flattened but steep. 

Dorsal valve very gently convex in lateral profile but 
nearly flat in anterior profile. Umbonal region slightly 
swollen; posterolateral areas gently concave. Antero- 
median region flattened. 

Ventral valve with thick dental ridges and long nar- 
row muscle field divided medially by a low myo- 
phragm. Dorsal valve interior having a moderately large 
cardinal process, rounded and without shaft. Socket 
ridges strong and elevated, bounding deep sockets. 
Crural bases triangular and descending close to the 
inner surface of the valve; descending lamellae of the 
spire closely following the inside of the dorsal valve; 
complete spire not seen. 

Measurements (in mm).—USNM 201072 (Holo- 
type): length 8.8, dorsal valve length 7.3, hinge width 
5.5, maximum width 8.7, thickness 5.5. 

Diagnosis.—Echinocoelia of intermediate size hav- 
ing a fairly long interarea, moderately incurved beak, 
and three or four concentric lines to the mm on the 
body of the ventral valve. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—522i, n; 523b, f, s; 5241. 

Types.—Holotype: USNM 201072. Figured Para- 
types: USNM 201065, 201156. 

Discussion.—This species is much smaller than 
Echinocoelia ambocoelioides Cooper and Williams 
(1935) and E. incurva Cooper and Williams (1935). 
These two species also have longer interareas and less 
incurved beaks than the New Mexico species. Echin- 
ocoelia denayensis Johnson (1966) is similar in size, 
but its interarea is much longer and the beak less in- 
curved than that of E. pretiosa. Moreover, the New 
Mexico species has a narrower hinge and more round- 
ed sides than those of the Nevada species. 

Echinocoelia pretiosa is a very rare species, only 
seven specimens having been found in the Onate For- 
mation. 


BULLETIN 315 


Superfamily SPIRIFERACEA W. King, 1846 
Family MUCROSPIRIFERIDAE Boucot 1959 


Genus ELEUTHEROKOMMA Crickmay, 1950 


Eleutherokomma extensa new species 
Plate 27, figures 30-35 


About medium-sized for genus, greatly extended 
laterally with sharply pointed extremities. Width about 
three times the length. Anterior commissure unipli- 
cate. Beak moderately extended; interarea fairly long, 
curved, apsacline. Surface marked by 14 costae on 
each side of the fold and sulcus; costae covered by 
crowded, thin, concentric lamellae, four to one mm. 
Surface of lamellae in sulcus and flanks marked by fine 
radial capillae. 

Ventral valve moderately convex in both profiles; 
sulcus originating at beak, deep and narrow, occupy- 
ing about one-seventh of width. Sulcus narrowly 
V-shaped and forming a short, sharp tongue anterior- 
ly. Flanks gently convex near the sulcus but becoming 
gently concave laterally. 

Dorsal valve moderately convex in lateral profile; 
broadly convex medially in anterior profile with flanks 
becoming gently concave laterally. Fold originating at 
beak, rather narrow throughout its length, forming a 
V-shaped arch anteriorly. Fold barely perceptibly set 
off from flanks. 

Ventral valve interior with strong dental plates. 

Measurements (in mm).— 


dorsal maxi- 
valve mum mid-  thick- 
length length width width ness 


USNM 200876a 
(Holotype; loc. 3052a) 10.6 8.2 29.0 19.3 7.4 


Diagnosis.—Eleutherokomma having width three 
times its length. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—3052a. 

Types.—Holotype: USNM 200876a. Unfigured 
Paratype: USNM 200876b. 

Comparison.—This species has a strongly extended 
hinge like that of E. reidfordi Crickmay (1950), but the 
length is much less in relation to width than in that 
species. Furthermore, E. reidfordi bears a costa in the 
sulcus but the sulcus of E. extensa is without a costa 
in the sulcus and the fold does not have a correspond- 
ing median depression as in E. reidfordi. The paratype 
was partially sectioned and exhibits strong dental 
plates. 


Eleutherokomma species 
Plate 39, figures 45-47 


About usual size for genus, wider than long; widest 
along the hinge; cardinal extremities acutely pointed. 
Sides obliquely rounded; anterior margin broadly 
rounded. Valves unequally convex, the dorsal valve 
having the greater convexity. Interarea long, ortho- 
cline and fairly strongly curved. Fold and sulcus with- 
out costae; flanks marked by 12 to 15 (?) costae, those 
of the extremities fine and indistinct. Entire surface 
marked by unevenly distributed concentric lamellae 
that have faint, fine capillae on their margins. 

Ventral valve moderately convex in lateral profile 
but broadly and fairly strongly convex in anterior pro- 
file. Sulcus beginning at the beak, widening gradually 
anteriorly to occupy about one-fourth the valve width. 
Tongue short. Flanks gently convex; flanks at ears 
concave. 

Dorsal valve more convex than the ventral valve in 
lateral profile; broadly domed in anterior profile; flanks 
flattened and extremities less well demarcated than 
those of the ventral valve. Fold originating at the beak 
low and narrow, forming a fairly strongly uniplicate 
anterior commissure. 

Measurements (in mm).—USNM 203560a: length 
14.0, hinge-width 21.0+, dorsal valve length 11.7, mid- 
width 19.4, thickness 11.5. 

Diagnosis.—Eleutherokomma with strongly convex 
dorsal valve, long orthocline interarea, and irregularly 
lamellose ornamentation. 

Stratigraphic occurrence.—Sly Gap Formation? 
(uppermost sandstone). 

Locality.—S22s. 

Types.—Figured Specimens: USNM 203560a, b, c. 

Discussion.—This species is represented by two © 
ventral valves and a complete but very poorly pre- 
served individual. The ventral valves are impressions, | 
one of the exterior only, the other of both interior and 
exterior. The fact that the interior bears no median | 
septum separates this species from Tylothyris North — 
(1920). The exteriors are not well preserved but very 
fine capillae can be detected with careful study, thus — 
helping to confirm the assignment to Eleutherokom- — 
ma. Most species of Eleutherokomma are strongly © 
mucronate and thus are not to be identified with the | 
form under discussion. Eleutherokomma aechmorpha 
Crickmay (1963), from the Waterways Formation, is — 
perhaps the closest described species but it has more | 
closely crowded concentric lamellae, less convex | 
valves and a long, nearly procline interarea. The Ca- 
ballos Mountains form appears to be a new one, but | 
the material is not adequate to establish a species. 


DEVONIAN BRACHIOPODS OF NEW MEXxICO: COOPER AND DuTRO 97 


Genus MUCROSPIRIFER Grabau, 1931 


Mucrospirifer parvus new species 
Plate 27, figures 1-19 


Small for genus, semielliptical in outline; hinge about 
equal to the maximum width, which is located at about 
midvalve; cardinal extremities nearly a right angle. 
Sides slightly oblique and slightly rounded; anterior 
margin broadly rounded; anterior commissure unipli- 
cate. Interarea long, slightly curved and with beak in- 
curved, apsacline. Delthyrium wide, open. Surface 
costate, nine or ten costae on each flank; entire surface 
covered by closely crowded imbricating lamellae with- 
out any micro-ornament. 

Ventral valve unevenly convex in lateral profile, the 
posterior part having the greater convexity; anterior 
profile forming a broad curve, indented medially, with 
lateral slopes gently convex and moderately steep. 
Sulcus originating at beak, widening anteriorly to oc- 
cupy about one-third valve width at midvalve; sulcus 
shallow and flat-floored and extending onto a short, 
abruptly truncated tongue. 

Dorsal valve fairly evenly and gently convex in lat- 
eral profile but broadly and fairly strongly convex in 
anterior profile. Fold beginning at the beak, elevating 
slightly anteriorly but never strongly elevated above 
the convex flanks; fold flat-topped but with an indis- 
tinct median depression. 

Interior not known except for lack of myophragm 
in ventral valve. 

Measurements (in mm).— 


dorsal mid- 
length valve hinge width thick- L/W 
(L) length width (W)~ ness _ ratio 


USNM 203541 

(Holotype; 

loc. 528k) 1224 LOS 1G: 6) 17-0 10! 0%65 
USNM 201046a 

(loc. 526c) 9.6 85 14.4 13.8 7-35 10569 
USNM 201104 

(loc. 524)) 15 E5428 17205 2010 12-3 OMT 


Diagnosis.—Small, non-extended Mucrospirifer 
having a flat fold and a sulcus with flat floor and trun- 
cated tongue. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —522f, g, i; 523e, i, u, z; 524a, g, j, 1, u; 
526c, h; 528k; 529), o. 

Types.—Holotype: USNM 203541. Figured Para- 
types: USNM 201046a, 201104. Unfigured Paratypes: 
USNM 201046b, c. 

Discussion.—Externally, this species is very similar 
to some specimens identified herein as Tylothyris North 
(1920), but it differs from them in not being laterally 


extended and in not having a myophragm in the muscle 
field of the ventral valve. 

Mucrospirifer parvus n. sp. suggests Mucrospirifer 
consobrinus (dOrbigny, 1850) from the Hamilton 
Group of New York, but that species is larger and has 
more laterally extended ears than the New Mexico 
species. A number of small mucrospiriferids in the 
Middle Devonian of the Hamilton and Traverse groups 
are suggestive of M. parvus, but they have not yet 
been described. 

This is a rare species in the Onate Formation. 


Mucrospirifer? cf. M.? sculptilis (Hall) 


A few specimens of a coarsely plicated mucronate 
spiriferoid are suggestive of Mucrospirifer? sculptilis 
(Hall, 1843), which is common in the Middle Devonian 
of New York. The specimens are so badly worn that 
they do not show well the strong concentric, zigzag 
lamellae characteristic of the species. The specimens 
are found associated with Tropidoleptus platys n. sp. 
in the Onate Formation at localities 528t, 529g, 3023b, 
in Johnson Park Canyon. 

Types.—Mentioned Specimens: USNM 201185a—d. 


Genus TYLOTHYRIS North, 1920 


Tylothyris compacta new species 
Plate 27, figures 57-67; plate 39, figures 23-26 


About medium size for genus, subrectangular in out- 
line, hinge forming the widest part but not extended 
beyond small points; sides rounded and anterior mar- 
gin broadly truncated. Anterior commissure strongly 
uniplicate. Surface of both valves costate, nine costae 
to a side in adult; whole surface covered by closely 
crowded, nonspinose, concentric lamellae, about two 
to amm near midvalve. Lamellae more closely crowd- 
ed at the anterior. 

Ventral valve slightly deeper than dorsal valve, fair- 
ly strongly convex in lateral profile and broadly con- 
vex in anterior profile, dome having a deep excavation 
at middle representing sulcus. Flanks convex and steep. 
Interarea short, curved and apsacline. Sulcus begin- 
ning at beak, wide and deep, forming a broad V and 
occupying about half shell width. Each costa bounding 
sulcus elevated slightly over flanks. Sulcus, in an oc- 
casional specimen, bearing an indistinct costa. Tongue 
forming sharp angle. 

Dorsal valve unevenly convex in lateral profile, pos- 
terior half having fair curvature but anterior half only 
slightly convex. Anterior profile broad triangle. Fold 
originating at beak, moderately elevated above flanks 
and somewhat narrowly rounded, bearing a slight me- 
dian depression. Flanks flat in profile but rounded in 
longitudinal direction. 


98 BULLETIN 315 


Ventral valve interior with greatly thickened delthy- 
rial region and short but deeply entrenched diductor 
scars; myophragm short and slender. 

Measurements (in mm).— 


dorsal 
valve hinge mid- __ thick- 
length length width width ness 


USNM 200986a 


(Holotype; loc. 522) 16.6 14.0 — 23.6 13.2 
USNM 200986b 
(loc. 522) 12.1 10.2 16.4 15.7 97 


Diagnosis.—Medium sized Tylothyris having a wide 
sulcus, nine costae on a side, short interarea, and 
greatly thickened ventral interior. 

Stratigraphic occurrence.—Thoroughgood Forma- 
tion and Contadero Formation. 

Localities. —Thoroughgood: 522. Contadero: 522a, 
5240a. 

Types.—Holotype: USNM 200986a. Figured Para- 
types: USNM 200986b, c; 201032. 

Discussion.—This species ts larger and less laterally 
extended than any other species of Ty/lothyris from the 
New Mexico Upper Devonian. It suggests some forms 
of T. mesacostalis (Hall, 1843) from the Chemung 
Group of New York, but the eastern specimens have 
a stronger costa in the sulcus, with a deeper depres- 
sion on the fold, and the myophragm of the interior is 
much stronger than that of the New Mexico species. 


Tylothyris novamexicana Stainbrook 
Plate 27, figures 36-50; plate 28, figures 25-32 


Tylothyris novamexicana Stainbrook, 1947, p. 323, pl. 47, figs. 31—- 
33. 


Small, wider than long, with hinge drawn out to long 
points. Sides slightly convex and sloping medially. In- 
terarea moderately long, curved, apsacline. Anterior 
commissure strongly uniplicate. Surface of both valves 
marked by costae, eight to 13 on a side depending on 
valve width; costae covered by closely crowded, 
smooth, non-spinose, lamellae—three or four to the 
mm. 

Ventral valve strongly convex in lateral profile but 
forming a broad dome in anterior profile with sides 
flattened and dipping steeply away from a deep median 
indentation formed by the sulcus, which originates at 
beak. Sulcus deep and subangular, forming long acute 
tongue at anterior. 

Dorsal valve about equal in depth to the ventral valve 
and fairly strongly convex in lateral profile; anterior 
view broadly triangular with the fold forming subacute 
apex and sides descending steeply. Fold originating at 
beak, elevating anteriorly above flanks, narrowly 
rounded to subangular. 


Ventral valve interior with strong muscle myo- 
phragm. Cardinal process comb-like. 
Measurements (in mm).— 


dorsal 
valve hinge mid- thick- 
length length width width ness 


USNM 135263 8.8 6.6 12.8 10.0 6.4 
USNM 201107 

(loc. 523q) 11.2 8.8 17.4 14.1 11.0 
USNM 201060 

(loc. 521g) 9.2 7.8 17.0 12.7 7.8 
USNM 203511 

(loc. 509u) 9.6 8.4 16.2 13.8 8.5 


Diagnosis.—Small Tylothyris having subangular fold 
and sulcus. 

Stratigraphic occurrence.—Contadero and Rhodes 
Canyon formations. 

Localities. —Contadero: 509u; 523q. Rhodes Can- 
yon: 521g; 523v; 524z; 528c, j. 

Types.—Figured Hypotypes: USNM 201060, 201017, 
203545a. 

Discussion.—This species is most like T. sulcocos- 
tata n. sp. in size and general appearance but it has a 
deeper sulcus with more sharply angular tongue, and 
a narrowly rounded fold that is not medially grooved. 


Tylothyris sulcocostata new species 
Plate 27, figures 51-56; plate 28, figures 1-24; plate 39, 
figures 48-51; plate 44, figures 21-26 

Small, semielliptical in outline; inequivalved, ven- 
tral valve being deeper. Hinge forming widest part and 
often drawn into sharp angles or blunt points. Sides 
sloping strongly medially; anterior subtruncate. An- 
terior commissure strongly uniplicate. Surface cos- 
tate, costae varying from eight to eleven on each side; 
costae covered by fine, concentric lamellae from three 
to five in one mm near midvalve. 

Ventral valve moderately convex in lateral profile 
but broadly triangular in anterior profile, sides flat and 
moderately steep, apex of triangle truncated or nar- 
rowly and gently concave where the sulcus is located. 
Interarea fairly long, slightly to moderately curved and 
apsacline. Delthyrium fairly wide and open. Sulcus 
originating at beak, widening rapidly anteriorly to oc- 
cupy about one-third to a quarter valve width; sulcus 
shallow and occupied medially by a low costa of vari- 
able strength. Costae on each side of sulcus stronger 
than those of flanks and slightly elevated above flanks. 

Dorsal valve gently convex in lateral profile; ante- 
rior profile a broad dome slightly elevated medially by 
the flatly rounded fold, the sides of the dome slightly 
convex and moderately sloping to the extremities. Fold 
originating at beak, flatly convex and marked medially 
by low narrow depression that extends from beak to 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 99 


anterior margin and matches the costa of the ventral 
sulcus. 

Ventral valve interior with strong muscle myo- 
phragm. 

Measurements (in mm).— 


dorsal 
valve _ hinge mid-  thick- 
length length width width ness 


USNM 201020 

(loc. 521d) 9.4 8.8 16.4+ 14.0 79 
USNM 201067a 

(loc. 506)) 8.4 8.3 17.4+ 13.7 8.3 
USNM 201067b 

(loc. 506j) 8.7 8.2 16.7+ 14.2 8.0 
USNM 201067c 

(loc. 506j) 8.9 8.0 17.1+ 14.1 7.5 
USNM 201067d 

(loc. 506j) 8.6 8.2 14.7? 13.0 6.7 
USNM 201019 


(Holotype; loc. 509z) —-10.0 9.6 17.3+ 14.8 8.8 
USNM 201016 


(loc. 509p) 11.2 10.0 19.2* 14.6 10.4 
USNM 201043 
(loc. 509r) 9.8 8.4 16.0* 12.2 ks) 


Diagnosis.—Small Tylothyris with wide hinge, costa 
in the sulcus and depression on the fold, and with eight 
to eleven costae on a side. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506j, y; 509f, 1, p, r, s, x, z; 521d, e, k, 
m; 522x; 523m; 524m, w; 526e; 3054c. 

Types.—Holotype: USNM 201019. Figured Para- 
types: USNM 201016, 201020, 201043, 201067c. Un- 
figured Paratypes: USNM 201033, 201067a, b, d. 

Discussion.—This species is similar to T. novamex- 
icana Stainbrook (1947) in size but is readily distin- 
guished by its flatter fold, with shallow median groove, 
and the corresponding sulcus on the ventral valve, 
which bears a median costa. 


Tylothyris species | 
Plate 29, figures 29-41 


This species is larger than T. novamexicana Stain- 
brook (1947) and has a strong costa in the sulcus of 
the ventral valve and a corresponding groove along 
the center of the fold of the dorsal valve. It is also 
larger and wider than T. su/cocostata n. sp., the fold 
is narrow and provided with a strong groove, and the 
sulcus bears a much stronger costa than that of the 
Sly Gap species. Although this species approaches 7. 
compacta N. sp. in size it is differently proportioned, 
not being so chunky and in being more laterally ex- 
tended. Tylothyris sp. 1 occurs in the silty limestone 
of the Contadero Formation. It has proved impossible, 
with the material at hand, to produce enough clean 
specimens to give good descriptive material. Its de- 


scription must therefore await discovery of better 
specimens. 

Diagnosis.—Wide-hinged, fairly large Tylothyris 
having a narrow fold and sulcus and a strong median 
costa and groove. 

Stratigraphic occurrence.—Contadero Formation. 

Localities. —S09t; 521f, j; 523n, p; 5240, t; 526g; 528q. 

Types.—Figured Specimens: USNM 201047, 201048, 
201049. 


Genus STROPHOPLEURA Stainbrook, 1947 


Strophopleura notabilis (Kindle) 
Plate 29, figures 1-21 


Spirifer bimesialis Girty (not Hall, 1858), 1900, p. 55. 

Spirifer notabilis Kindle, 1909, p. 26, pl. 7, figs. 3-6c. 

Strophopleura notabilis (Kindle) Stainbrook, 1947, p. 324, pl. 44, 
figs. 17-20. 


This is a fairly rare species and it is difficult to obtain 
good specimens with the mucronate points intact. The 
illustrations are introduced to better the understanding 
of this genus. 

Stratigraphic occurrence. 
Member). 

Localities. —506, a, b, p, q; 507b, d, h, k, m, 0; 509a, 
b, e; 527a, d, e, f, g, j, 1, p; USGS (Kirk) 508, 509; 
3026, a; 3034; 3042, s. 

Types.—Hypotype: USNM 62025. Figured Hypo- 
types: M.A.S. 1586A, B (=USNM 155804). Figured 
Hypotypes, herein: USNM 200847, 200848a, b, 201146, 
201147a, 201148a, 201149a. 


Percha Formation (Box 


Family SPINOCYRTIIDAE Ivanova, 1959 
Genus ORTHOSPIRIFER Pitrat, 1975 


Orthospirifer species 
Plate 29, figures 22-28 


Large, wider than long, hinge forming the widest 
part in young specimens; cardinal extremities acute 
and sharply pointed in young but rounded in adult 
shells; sides rounded and sloping obliquely medially; 
anterior commissure uniplicate. Interarea moderately 
long, curved, apsacline; delthyrium open. Surface cos- 
tate, fold and sulcus without costae; flanks marked by 
21 costae on a large specimen; micro-ornament con- 
sisting of fine radial capillae on fold and sulcus but not 
seen elsewhere on valves. 

Ventral valve gently convex in lateral profile, broad- 
ly and moderately convex in anterior view. Sulcus 
originating at beak, only moderately deep anteriorly 
and occupying about one-fourth the valve width. Flanks 
gently convex. Tongue short and broadly rounded. 

Dorsal valve moderately convex in lateral profile 
with maximum convexity in posterior part; anterior 


100 BULLETIN 315 


profile broadly but gently convex. Fold originating at 
beak, very gently convex and scarcely elevated above 
flanks at anterior. Flanks moderately convex, more so 
than those of the ventral valve. 

Ventral valve interior with strong, divergent dental 
plates and a short apical plate. Details of the interior 
of the dorsal valve not seen. 

Measurements (in mm). 


maxi- 

dorsal mum 
length valve hinge width thick- L/W 
(L) length width (W) ness _ ratio 


USNM 201128a 

(loc. 3023c) 25.5 a 34.8 37.0 7.6? 0.70 
USNM 201128b 

(loc. 3023c) 37.8 37.0 40.0 59.2 11.0 0.66 


Diagnosis.—Orthospirifer apparently having flanks 
devoid of micro-ornament. 

Stratigraphic occurrence.—Onate Formation. 

Locality.—526z; 3023c. 

Types.—Figured Specimens: USNM 201128a-e. 

Discussion.—The specimens on which this species 
is based are poor. Most of them are exfoliated, but 
where the preservation of the costae on the flanks 
seems well preserved, no capillae were observed on 
them. This is the only example among the many species 
of Orthospirifer in the National Collections on which 
radial lines do not appear on the costae of the flanks. 
Because some fairly worn specimens of other species 
of Orthospirifer exhibit traces of the lines, it is prob- 
able that the Onate species had them and better spec- 
imens would reveal their presence. 


Genus SPINOCYRTIA Fredericks, 1916 
Spinocyrtia? species 


Three specimens, preserved in a chert replete with 
Tentaculites Schlotheim (1820), are referred with some 
doubt to this genus. The specimens are partially silic- 
ified and the single dorsal valve is an inner impression. 
The characteristic granules of Spinocyrtia were not 
detected but the ventral valves have the nearly or- 
thocline, curved interarea and subangular sulcus often 
seen in this genus. The costae are closely crowded, 
low and rounded, about 13 on the flanks of the ventral 
valve. The fold of the dorsal valve is narrowly rounded 
and evenly, but moderately, elevated above the round- 
ed flanks. The specimens measure about 20 mm long 
and 30 mm wide. 

Stratigraphic occurrence. 

Locality.—529g. 

Types.—Mentioned Specimens: USNM 200985a-—c. 


Onate Formation. 


ACUTORIA new genus 
[Latin acutus = acute, sharp] 


Spiriferoid, large, transverse; sides rounded; hinge 
narrower than midwidth; concavo-convex, ventral 
valve with concave flanks and deep angular sulcus; 
dorsal valve swollen, with convex flanks and high, an- 
gular fold. Beak strongly obtuse. Costae wide, flat, 
separated by narrow striations. Entire surface covered 
by fine radial capillae, strongest on fold and in sulcus. 

Ventral valve with open delthyrium and horizontal 
apical plate between long divergent dental lamellae. 
Dorsal valve with short median septum buttressing 
cardinal process and cardinalia. 

Type species.—Acutoria angulata n. sp. 

Diagnosis.—Like Orthospirifer internally but with 
concave ventral valve, acute fold and broad flat cos- 
tae. 

Comparison.—This brachiopod was originally mis- 
taken for Paraspirifer Wedekind (1926) and identified 
with P. acuminatus (Conrad, 1839) of the Onondaga in 
the eastern United States and Canada (Stevenson, 
1943). It differs from Paraspirifer in the concavity of 
the ventral valve and in its fine radial ornament. The 
costate surface of Paraspirifer is covered by close rows 
of concentric lines bearing minute spines; the fine or- 
nament of Acutoria consists of fine longitudinal ca- 
pillae. Distinction from Orthospirifer is based on the 
flat costae with the very fine grooves separating them, 
the concave ventral valve, the acute dorsal fold and 
the short median septum. The fold of Orthospirifer is 
invariably broadly rounded and low, and its costae are 
more or less narrowly rounded. 


Acutoria angulata new species 
Plate 28, figures 33-42; Text-figure 9 


Large, subrectangular in outline; hinge narrower than 
maximum width which is at midvalve. Sides rounded, 
postero-lateral extremities forming angle of 126°; an- 
terior margin broadly emarginate. Anterior commis- 
sure angularly uniplicate. Ventral valve interarea long, 
plane, apsacline. Surface with both macro- and micro- 
ornament; macro-ornament on flanks consisting of 
broad, flat costae separated by narrow interspaces, 
eleven or twelve costae on each side; fold and sulcus 
non-costate; micro-ornament consisting of fine, ele- 
vated, radial lines (capillae) covering entire surface of 
both valves. 

Ventral valve unevenly convex along center line, 
umbonal region flatly convex, anterior and middle re- 
gions strongly concave; anterior profile broadly, deep- 
ly concave, forming broad V. Sulcus originating just 
anterior to beak, deepening and widening rapidly to 


DEVONIAN BRACHIOPODS OF NEW MExICco: COOPER AND DUTRO 101 


Text-figure 9.—Acutoria angulata n. sp. 1. Part of the dorsal valve exterior showing radial capillae on the costae and fold, x5, paratype 
USNM 201127f; 2. Apical part of the ventral valve interarea, showing the apical plate, x2, paratype USNM 201127g; 3. Beak of the dorsal 
valve showing silicified median septum, buttress plates and cardinal process, x4, paratype USNM 201127i. Onate Formation, locality 526z. 


occupy nearly half maximum width. Sulcus V-shaped, 
forming angular tongue; flanks flattened to gently con- 
cave. 

Dorsal valve very deep; gently convex in lateral pro- 
file but forming a high, pointed dome in anterior pro- 
file; flanks slightly convex, steep. Fold originating at 
beak, elevating anteriorly, steep-sided and fairly 
strongly elevated above flanks. 

Ventral valve interior with broad and thick apical 
plate; strong dental plates. Dorsal valve interior with 
short median septum just anterior of umbo. 

Measurements (in mm).— 


dorsal 
valve hinge mid-_ thick- 
length length width width ness_ height 


USNM 201127d 
(Holotype: 36.4 — 42.0? 52.0 — 4.0? 
loc. 526z) 
USNM 201127a = 
(loc. 526z) 35.0 _— 39.5 42.4 3227 
USNM 201127b 
(loc. 526z) — 37.2 31.8 480 — _ 26.0 
USNM 201127f 
(loc. 526z) oa 36.4 38.02? 47.0? — 16.7 


Diagnosis.—Large Acutoria having concave ventral 
valve with sharply angular fold and sulcus; entire sur- 
face radially capillate. 

Stratigraphic occurrence.—Onate Formation. 

Localities. —526z; 528t; 3023c. 

Types.—Holotype: USNM 201127d. Figured Para- 
types: USNM 201127a-c, e, f, g, i. 

Discussion.—The most similar species to this one 
are: the American Paraspirifer acuminatus (Conrad, 
1839), P. bownockeri (Stewart, 1927), and the German 
P. cultrijugatus (Roemer, 1844); but all of these differ 
in their micro-ornament from that of Acutoria angu- 


lata n. sp. The micro-ornament of Paraspirifer We- 
dekind (1926) consists of crowded concentric lines 
bearing minute spines, whereas that of the New Mex- 
ico species has fine radial lines only. Acutoria angu- 
lata differs in another respect in that the flanks of the 
ventral valve are concave or flat; they are not convex 
as in the species of Paraspirifer mentioned above. 

Acutoria angulata also suggests Orthospirifer me- 
sastrialis (Hall, 1843), both being liberally adorned with 
fine radial lines; however, the New York species has 
a rounded fold and sulcus, is much wider, and its ven- 
tral valve is not deeply concave. 

Some of the specimens illustrated here were used 
by Stevenson (1943, p. 222) in his announcement of 
an “‘Onondagan equivalent in New Mexico’’, based 
on the identification of *‘Paraspirifer cf. acuminatus’’. 
The specimens now prove not to be Paraspirifer and 
their occurrence is in the Onate Formation, as indi- 
cated by evidence discussed elsewhere in this paper, 
a much younger formation than the Onondaga. 


Genus ACUTATHECA Stainbrook, 1945 


Acutatheca prolifica new species 
Plate 36, figures 1-26; plate 44, figures 11-20 


Not Acutatheca propria = A. cf. propria Stainbrook, 1948; p. 777, 
Plae2aiesse2 e222 


Small, strongly inequivalved; pedicle valve semi- 
pyramidal in profile, dorsal valve subpentagonal in 
outline; sides sloping medially; hinge slightly less wide 
than the greatest width, which is near the posterior of 
the dorsal valve; cardinal extremities narrowly round- 
ed; anterior commissure strongly uniplicate. Ventral 
interarea long (in direction of valve length), gently 
curved, steeply apsacline to catacline. Delthyrium long 
and narrow, partially closed by apical plate (pseudo- 


102 BULLETIN 315 


deltidium?) that occupies the posterior third to half of 
delthyrium; delthyrial edges marked by narrow plates 
oblique to surface of interarea. Surface of both valves 
costate, costae broadly rounded, two to four ona side, 
and separated by spaces about equal to width of cos- 
tae. Entire surface covered by fine radial capillae, seen 
only in unusually well preserved specimens. 

Ventral valve triangular in lateral profile with ven- 
tral valve gently convex; anterior profile narrowly 
domed. Umbonal region narrow; sulcus originating near 
beak and widening anteriorly to occupy about 40 per- 
cent of maximum width of valve. Flanks flattened in 
profile, depressed below two strong costae bounding 
sulcus. 

Dorsal valve fairly strongly convex in lateral profile 
but having the umbonal region narrowly curved; an- 
terior profile a strongly domed curve having steeply 
dipping sides. Flanks gently swollen. Narrow fold be- 
ginning at the beak, widening gradually anteriorly, 
somewhat flat-topped, and with a median depression 
throughout its length. Flanks depressed well below the 
fold and separated by deep bounding grooves. 

Ventral valve interior with two strong, slightly di- 
vergent dental plates extending for one-sixth to one- 
fifth the surface length of the ventral valve. Dorsal 
valve interior with two convergent plates that form a 
small chamber with the median ridge. 

Measurements (in mm).— 


dorsal maxi- 
valve hinge mum_ thick- apical 
length length width width ness angle 
USNM 20110Sa 
(loc. 506c) 1274 1222 15.9 VO 23 G2 
USNM 201105c 
(loc. 506c) 12.5 Mie ILO 16:67) 107 922 
USNM 201105d 
(loc. 506c) 10.7 11.2 11.4 14.2. 10.0 96° 
USNM 201105b 
(loc. 506c) 18205 See i720 20.3 14.5 94° 
USNM 201001a 
(loc. 506i) 1242052517729) S29 Oia 
USNM 203524 
(loc. 509g) 10.7. = 10.3 10.7 14.4 10.6 97° 
USNM 200958 
(loc. 509h) 12.6 12.7 [Seo 7e3 182) 922 
USNM 201008 
(loc. 509w) 1332 12.8 16.0 19:4 13.8 92° 


Diagnosis.—Large, strongly costate Acutatheca 
having a median depression in the fold. 

Stratigraphic occurrence.—Sly Gap Formation (up- 
per part). 

Localities.—506c, 1, v; 509f, g, h, i, j, k, 1, m, n, v, 
w; 5211, 0, p, r; 526k, 3045b. 

Types.—Holotype: USNM 201050a. Figured Para- 
types: USNM 200958, 201001la, 201008, 201132b, 


201142. Unfigured Paratypes: USNM 201001b, 
201050b-d, 201132a. 

Discussion.—Although Stainbrook assigned speci- 
mens from Alamo Canyon to his species A. propria in 
his description of the Independence Shale fauna (1945, 
p. 56), he was not so positive in his paper on the Sly 
Gap fauna (1948, p. 777, pl. 2, figs. 29-32). In the latter 
paper, the species here described was referred to as 
Acutatheca cf. propria in the plate legend but as A. 
propria in the text. The Independence shale specimen 
figured in 1945 (pl. 1, figs. 39-41) is a very small shell 
that differs considerably from the New Mexico species. 
The New Mexico specimens are twice the size of the 
Iowa species or even more than that. In addition, they 
are more strongly costate and have fewer costae on 
the lateral slopes. 

The apical plate in some of the specimens is strongly 
convex, in some it is nearly flat, and in others it is 
slightly concave. Probably, the normal condition is 
convex but, most likely, in many specimens the apical 
plate has been flattened during fossilization. Many 
specimens exhibit lateral plates as well as the apical 
plate. These are located on the sides of the delthyrium, 
highest at the base and descending toward the apex 
where they fade out. They are oblique to the surface 
of the interarea. 


Family THOMASARIIDAE new family 


Small, hemipyramidal shells, with strong, long den- 
tal plates and thick apical plate. Brachial valve interior 
with vertically lamellate cardinal process and socket 
plates free. Surface marked by fine pustules. 

The family assignment of Thomasaria has been in 
question since Stainbrook proposed the genus in 1945 
and suggested ambocoeliid affinities. Pitrat (in Wil- 
liams et al., 1965, p. H711) did not place the genus in 
any known family. Because of its unusual combination 
of characters, we are erecting a new family which, at 
present, includes only the genus Thomasaria. 


Genus THOMASARIA Stainbrook, 1945 


Thomasaria altumbona Stainbrook 
Plate 35, figures 1-3 


Thomasaria altumbona Stainbrook, 1945, p. 58, pl. 4, figs. 22-30, 
text-fig. 2 [14]. 


This is a generally smaller species than the New 
Mexico forms referred to Thomasaria warreni n. sp. 
Specimens are illustrated to show details of the inte- 
rior and interarea of the ventral valve. 

Stratigraphic occurrence.—Independence Forma- 
tion. 

Locality.—5281, m. 

Types.—Figured Hypotypes: USNM 135294a, 
201066. 


DEVONIAN BRACHIOPODS OF NEW MEXIco: COOPER AND DuTRO 103 


Thomasaria demissa new species 
Plate 33, figures 55-65; plate 35, figures 4-24; plate 39, 
figures 35-37; plate 41, figures 27—33 


Small, but about normal size for genus, usually wid- 
er than long with narrowly rounded to sharply pointed 
cardinal extremities. Hinge wide, greater than, equal 
to, or slightly less than maximum width. Sides gently 
curved and sloping medially; anterior narrowly trun- 
cated. Interarea long (in the direction of valve length), 
flat to faintly curved, catacline to procline to steeply 
apsacline. Delthyrium long and narrow, partially 
plugged by narrow plates deposited on inner sides of 
dental plates; also partially closed by narrow, oblique 
plates on margin of the delthyrium; beak sharply 
pointed. Anterior commissure narrowly uniplicate. 
Surface marked by fine concentric, radial lines each 
bearing a row of minute, single-barreled spines. 

Ventral valve semipyramidal in lateral profile with 
the valve gently convex; anterior profile broadly 
domed. Umbonal region narrow; sulcus originating at 
beak, shallow throughout its length and widening an- 
teriorly to occupy about 40 percent of the hinge width. 
Tongue short and flatly rounded. 

Dorsal valve very gently convex in lateral profile; 
broadly and gently convex in anterior profile. Umbo- 
nal region moderately swollen, the swelling continued 
anteriorly to join the fold, which originates at about 
midvalve and is low and conspicuous only at anterior; 
flanks sloping to margins, steepest anterolaterally. 

Measurements (in mm).— 


maxi- 
dorsal mum __ thick- 
length valve hinge width ness apical 
(L) length width (W) (T) angle 


USNM 200936a 


(loc. 521e) LEA OS i7e4 7) 13.8 ile 
USNM 200936b 

(Holotype) OF OMemenl7e4: 17.4 12.2 89° 
USNM 201022b 

(loc. 509r) 9.4 10.2 17.8 17.8 11.6 106° 
USNM 201022c 8.4 9.8 19.2 19.2 10.6 108° 
USNM 201022a 10.0 10.5 17.5 17.5 12.3 88° 
USNM 203523a 

(loc. 509p) 9:9 -10'5 12.4 13.0 9.6 96° 
USNM 200987b 

(loc. 5090) 9.3 10.0 N3e/ 14.3 8.7 Oe 
USNM 200987c 

(loc. 5090) 96 OL6ye 2105 21h 0S 11.0 91° 
USNM 200987a 

(loc. 5090) LOWS) 112? 2316 2316-5 1456 OLY 
USNM 200987d 

(loc. 5090) 13.3 13.2 20.0* 20.0? 14.2 93° 
USNM 201157 

(loc. 5090) 9.4 9.2 14.6 14.6 11.8 84° 
USNM 203522a 

(loc. 521k) 8.7 S)7/ 17.8 17.8 11.3 91° 
USNM 203522b 11.5 12.0 18.8* 18.7 13.7 90° 


Mean L/W ratio = 0.56; Mean T/W ratio = 0.66. 


Diagnosis.—Strongly catacline, usually transverse 
Thomasaria having a low fold. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506e, j, y; 509n, 0, p, q, r; 521le, k, v; 
522b, k; 524v; 526d; 528p; 529p; USGS (Kirk) 534. 

Types.—Holotype: USNM 200936b. Figured Para- 
types: USNM 200936a, 200987d, 201022b, c, 201157, 
205261. Unfigured Paratypes: USNM 200987a-c, 
201022a. 

Discussion.—The main distinction between Thom- 
asaria warreni n. sp. and T. demissa n. sp. is in the 
nature of the ventral palintrope and the length/width 
ratio. Most of the specimens of 7. demissa have the 
palintrope usually catacline but varying to slightly ap- 
sacline or definitely procline. Variants of 7. warreni 
may attain a catacline palintrope but they are a mi- 
nority. Most specimens of 7. demissa have a length/ 
width ratio less than 0.80 because the catacline pal- 
intrope is usually flat and only seldom is curved at the 
apex. Other distinctions appear in the generally less 
pronounced fold on the dorsal valve of T. demissa 
when compared to that of 7. warreni. The apical angle 
of T. demissa tends to be less obtuse, nearer a right 
angle, than that of T. warreni, which is usually fairly 
strongly obtuse. 

Most specimens of 7. demissa come from the San 
Andres Range and to the west as far as Hermosa where 
several specimens were taken from the Sly Gap. The 
species is rare in the Sacramento Mountains, but some 
of the variants of T. warreni with strongly apsacline 
interareas resemble 7. demissa. 


Thomasaria warreni new species 
Plate 35, figures 25-38; plate 36, figures 27-44; 
plate 40, figures 26, 27; plate 43, figures 1-9 


Not Thomasaria altumbona Stainbrook, 1948, p. 777, pl. 2, figs. 38— 

43. 

Small but variable, large adults attaining a width of 
20 mm; shells unequal in depth, the ventral valve hav- 
ing a semipyramidal profile, the dorsal valve moder- 
ately convex in profile. Hinge wide but narrower than 
maximum shell width, which is near midvalve anterior 
to hinge. Sides with distinct shoulders just anterior to 
hinge but rounded and sloping medially anterior to 
midvalve. Cardinal extremities varying from angular 
to rounded. Apical angle greater than 90°. Interarea 
usually long (in the direction of valve length), variable, 
strongly to slightly curved and ranging from nearly 
orthocline to strongly apsacline to procline. Delthyri- 
um long and narrow, partially closed in well-preserved 
specimens by narrow, oblique marginal plates widest 
near the dorsal valve and tapering to disappearance 
apically. Closure of lateral plates not observed. Sur- 
face, except lateral margins, without costae but lateral 
margins provided with three or four obscure costae 


104 BULLETIN 315 


best seen in decorticated specimens. Micro-ornament 
consisting of fine concentric lines bearing minute uni- 
ramous spines, usually seen only in specimens with a 
thin coating of fine shaly matrix. 

Ventral valve variable in profile from semipyrami- 
dal, with an elongated palintrope, to moderately con- 
vex with a nearly orthocline palintrope; anterior pro- 
file strongly domed; sulcus originating on the umbo, 
generally shallow and narrow and producing a narrow 
tongue that is moderately rounded at its extremity. 
Sulcus bounded by poorly formed plication on each 
side; flanks convex but depressed slightly below pli- 
cations bounding the sulcus especially at anterior. Beak 
sharply pointed, slightly to fairly strongly incurved. 

Dorsal valve shallow and gently convex in lateral 
profile; broadly and gently convex in anterior profile; 
cardinal extremities slightly flattened; fold originating 
at umbo but poorly defined to midvalve where it be- 
comes slightly elevated above slightly convex flanks. 
Beak small, poorly defined; umbonal region moder- 
ately swollen. 

Ventral valve interior with strong dental plates and 
small teeth; dental plates extending along valve floor 
for about one-quarter valve length measured on its 
surface. Umbonal and delthyrial chambers often filled 
apically by adventitious shell substance; apical filling 
extending about half, or slightly more, of distance from 
apex to dorsal valve; trace of deltidial structure rarely 
seen. 

Dorsal valve with short flaring socket ridges and 
thick crural bases. 

Measurements (in mm).— 


maxi- 


dorsal mum __ thick- 
length valve hinge width ness _ apical 
(L) length width (W) (T) angle 
USNM 203520a 
(loc. 506c) 13283214 2a 1820 11270 92° 
USNM 203520b 
(loc. 506c) 15.0 14.6 14.7 20.0 11.6 99° 
USNM 203520c 
(loc. 506c) 5, ISO) TIS as} TTS) 108° 
USNM 200988c 
(loc. 506)) HAE lS) AE TIO) 103° 
USNM 200988a 
(Holotype; 
loc. 506j) 15.0 12.8 14.0 17.0 10.8 98° 
USNM 200988b 
(loc. 506j) jes GTI IES 0) 92° 
USNM 201025c 
(loc. S090) 14.5 13.3 15S 18.0 11.9 113° 
USNM 201025a 
(loc. S090) 17.3 14.1 13.0 18.4 ie 101° 
USNM 201025b 
(loc. 5090) 14.2 12.5 13.7 15.8 11.0 110° 
USNM 203521la 
(loc. 524m) 14.3 12.3 13.7 16.7 10.6 972 


USNM 203521b 


(loc. 524m) 1379) 273513245 1828 O oe 
USNM 20352Ic 

(loc. 524m) 135320 14 T Slee ls O ni 
USNM 200959 

(loc. 509n) 1223) EO OP a4 Gy 07) 94° 
USNM 200935b 

(loc. 521c) 12°39) 1S ASS 83 OG 
USNM 20093Sc 

(loc. 521c) 1374 12-45 126) eS Ole 94° 
USNM 200935a 

(loc. 521c) 1B 122 126 S402 Nits 


Mean L/W ratio = 0.86; mean T/W ratio = 0.64. 


Diagnosis.—Moderately transverse Thomasaria 
having a moderately to strongly curved interarea in 
the adult. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506c, 1, j, v; 509g, h, i, k, 1, m, n, 0, q, 
I, S3 V5 Ws Xe S21c. de. ky ms neon dn i522 bP Se: 
524m, n; 526k, m; 3045b; 3054h. 

Types.—Holotype: USNM 200988a; Figured Para- 
types: USNM 200935b, c; 201025a; 201130; 201148; 
203520b; Unfigured Paratypes: USNM 200935a; 
201025b, c; 203520a, c. 

Discussion.—See Thomasaria demissa n. sp. for 
comparison with this species. 

This species agrees with 7. altumbona Stainbrook 
(1948) in the incomplete development of the lateral 
plates bounding the delthyrium. It is, in general, a larg- 
er form than the one from the Independence Forma- 
tion and has a stronger fold and sulcus. Stainbrook’s 
illustrated specimens (1945) are unusually large. 
Thomasaria warreni has strong resemblance to T. 
rockymontana (Warren, 1928) but differs in the lack 
of development of the delthyrial region. In the Alberta 
species, the lateral plates grow together at the distal 
part of the delthyrium to form a convex cover, leaving 
an elongated opening toward the apex. The Alberta 
thomasarias are evidently as variable as those in New 
Mexico, and they may be divided ultimately into sev- 
eral species. 


Family CYRTOSPIRIFERIDAE Termier and 
Termier, 1949 


Genus Cyrtospirifer Nalivkin (in Fredericks, 1924) 
Cyrtospirifer? animasensis (Girty) 


Spirifer disjunctus var. animasensis Girty, 1900, p. 48, pl. 4, figs. 
1-10. 

Spirifer whitneyi var. animasensis Kindle (not Girty, 1900), 1909, 
Dao pleo nese lay nes ar 

Cyrtospirifer animasensis (Girty) Stainbrook, 1947, p. 318, pl. 44, 
figs. 21-24. 
This is an often misidentified species, consequently 

we have queried the citation above. The name has 

been used for cyrtospiriferids having a wide hinge and 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 105 


a long steeply apsacline or catacline interarea. Kin- 
dle’s specimen (1909, pl. 9, figs. 1—-3a) is entirely unlike 
the one figured by Stainbrook. Furthermore, Girty’s 
figures indicate a species larger and less transverse 
than that figured by Stainbrook. Possibly, Girty’s 
specimens include more than one species because the 
larger ones figured by him are quadrate and not widely 
extended at the hinge. This is a rare species in the 
Percha (Box Member) and our specimens and Stain- 
brook’s figured one are inadequate to settle the prob- 
lem. 

Localities.—507h; 527a. 

Types.—?Hypotype: M.A.S. 1594 (=USNM 135250) 
in Stainbrook (1947, p. 318, pl. 44, figs. 21-24). 


Cyrtospirifer arenarius new species 
Plate 30, figures 35—46 


About medium size for genus, subrectangular in out- 
line, wider than long, hinge narrower than maximum 
shell width, which is at about midvalve. Valves sub- 
equal in depth. Sides rounded; anterior margin trun- 
cated to broadly emarginate. Cardinal extremities ob- 
tuse; posterolateral margins varying from 115° to 125°. 
Interarea long, curved, apsacline to nearly orthocline. 
Delthyrium wide, open. Surface costate, costae low 
and rounded, crowded, wider than width of inter- 
spaces and 16 to 18 ona side. 

Ventral valve moderately convex in lateral profile; 
broadly convex in anterior profile with median region 
deeply bisected by deep angular depression repre- 
senting sulcus. Flanks short and steep. Sulcus origi- 
nating at beak, widening anteriorly to occupy about 
half valve width. Sulcus deep and forming broad 
V-shaped trough ending in fairly long subangular 
tongue. Sulcus marked by 13 costae. Slopes of flanks 
to cardinal extremities concave and cardinal extremi- 
ties slightly concave and having small ears. 

Dorsal valve moderately convex in lateral profile but 
broadly and moderately convex in anterior profile, fold 
of adult scarcely affecting apex of curve. Fold starting 
on somewhat swollen umbo, widening to anterior mar- 
gin but standing only slightly above flanks or flush with 
them. Flanks convex and moderately steep. 

Ventral valve interior with strong, widely divergent 
dental plates. 

Measurements (in mm).— 


dorsal 
valve hinge mid- thick- 
length length width width ness 


USNM 201015Sd 

(loc. 522) 21.4 17.1 32,0% 31.0 15.1 
USNM 201015a 

(loc. 522) 225) 17.0 26.0* 27.0 17.0 


USNM 20101Sc 

(loc. 522) 20.6 15.3 22.0 26.4 15.0 
USNM 20101S5e 

(loc. 522) 22.8 18.6 25.8 28.3 16.8 
USNM 20101Sb 

(Holotype) 21.3 15.0 Zallea¥ Dies 15.8 


Diagnosis.—Medium-sized, subrectangular Cyrto- 
spirifer with wide fold and sulcus and broadly emar- 
ginate anterior. 

Stratigraphic occurrence. 
tion. 

Locality.—S22. 

Types.—Holotype: USNM 201015b. Figured Para- 
types: USNM 201015a, c. Unfigured Paratype: USNM 
201015d. 

Discussion.—This species suggests some forms of 
Cyrtospirifer portae Merriam (1940) but, when com- 
pared to the type specimen of that species, which has 
similar measurements to USNM 201015c, the Nevada 
species proves to be more closely costate on the fold 
and sulcus, and is a thicker specimen with steeper 
flanks in both valves. Cyrtospirifer arenarius n. sp. 
seems not to have any close relatives in the Devonian 
of eastern New York, as revealed by the species of 
Cyrtospirifer described and illustrated by Greiner 
(1957). 

Although C. arenarius is similar in size to C. whit- 
neyi (Hall, 1858), it differs from that species in having 
a less conspicuous, flatter fold, and a more erect beak; 
and it is less costate. 


Thoroughgood Forma- 


Cyrtospirifer breviposticus Stainbrook 
Plate 29, figures 42-44 


Cyrtospirifer breviposticus Stainbrook, 1947, p. 320, pl. 44, figs. 3- 
6. 

Not Cyrtospirifer breviposticus Stainbrook. Johnson, Reso, and 
Stephens, 1969, p. 1363, pl. 160, figs. 1-18, text-fig. 6. 


This is a small, subpyramidal Cyrtospirifer (USNM 
135257) with narrow hinge and maximum width at mid- 
valve. This may be a young specimen of a larger species 
but, if so, that species was not found by any of the 
parties collecting for the Smithsonian Institution. 
Stainbrook’s specimen comes from the Rhodes Can- 
yon Formation ‘‘at the east end of Rhodes Pass, San 
Andres Mountains’’, from a rock unit that Stainbrook 
equated with the Percha (Bella = Box Member) that 
is exposed west of the San Andres Range. The beds 
from which this specimen (USNM 135257) came are 
in the type section of the Rhodes Canyon Formation. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation. 

Locality.—528c. 

Types.—Holotype: USNM 135257, figured in Stain- 
brook (1947). 


106 BULLETIN 315 


Discussion.—The identity of Stainbrook’s type with 
the specimens illustrated by Johnson, Reso and Ste- 
phens may be questioned. That Stainbrook’s specimen 
is a youthful one is almost certain, but it is difficult to 
visualize its growing into the strongly convex forms 
figured by Johnson, Reso and Stephens. Furthermore, 
the interarea of the ventral valve would tend to be 
flatter than that shown in the specimens illustrated by 
Johnson, Reso and Stephens. 


Cyrtospirifer convexus new species 
Plate 31, figures 1-10 


Small, subtrapezoidal in outline; hinge forming the 
widest part; cardinal extremities acute, slightly mu- 
cronate in some specimens. Sides oblique; anterior 
margin truncated. Interarea long, slightly curved to- 
ward apex, strongly apsacline to catacline. Delthyrium 
narrow, Open except at apex where it is closed by a 
short apical plate. Surface finely costate, costae low, 
rounded, separated by spaces narrower than costae; 
micro-ornament consisting of fine pustules. Twenty- 
one to 28 costae on each flank of adults and old spec- 
imens. 

Ventral valve fairly strongly convex and much deep- 
er than dorsal valve in lateral profile; anterior profile 
a misshapen dome, excavated medially and having 
sides steeply descending and slightly convex but be- 
coming flat or concave toward the cardinal extremi- 
ties. Sulcus originating at the beak, widening and 
deepening anteriorly to occupy about one-quarter of 
valve width. Sulcus occupied by 11 costae that in- 
crease from two at beak by insertion and bifurcation. 

Dorsal valve strongly convex in lateral profile; an- 
terior profile broadly convex and having a medial knob 
representing fold, flanks flat to slightly convex. Fold 
originating at beak, narrowly rounded and moderately 
elevated above flanks. Sides of fold not strongly de- 
marcated from flanks in most specimens. 

Measurements (in mm).— 


dorsal 
valve _ hinge mid- thick 
length length width width ness 
USNM 201095a 
(loc. 506b) 13.7 13.4 24.8* 18.0 14.6 
USNM 201095b 10.5 10.3 20.3 16.0 9.1 
USNM 201092a 
(Holotype; loc. 3026a) 16.4 13.5 27.4%? 19:8 15.0 
USNM 201092b 11.8 11.4 PPE) AIS) 12.3 
USNM 201103a 
(loc. 506) 16.8 16.0 34.0? 19.8 16.8 
USNM 201103b 
20.6*? 14.6 10.5 


(loc. 506) 11.4 11.0 


Diagnosis.—Wide-hinged Cyrtospirifer with long, 
slightly curved interarea, 21—28 fine costae on each 
flank, and a convex dorsal valve. 


Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —506, a, p, q; 507h, m, 0; 509b, e; 527a, 
d, e, f; 3026, a. 3042, s; 3054g. 

Types.—Holotype: USNM 201092a. Figured Para- 
type: USNM 201103a. Unfigured Paratypes: USNM 
201103b, 201092b, c, 201095a, b. 

Comparison.—This species is akin to Cyrtospirifer? 
animasensis (Girty, 1900) in having a wider hinge that 
may be slightly mucronate, a more convex dorsal valve, 
a narrower fold and sulcus and smaller size. It has 
been usual to identify any small, subconical Cyrto- 
spirifer from the Percha Formation as C.? animasensis 
(Girty), but the Colorado form is quite different from 
those of the Percha (see Cyrtospirifer? animasensis 
above). 


Cyrtospirifer pyramidalis new species 
Plate 31, figures 17-42 


About medium-sized for genus, wider than long, 
subrectangular in outline; sides extending obliquely 
medially; anterior emarginate in dorsal view; valves 
unequal, ventral valve semipyramidal and dorsal valve 
shallow; interarea very long and flat; delthyrium about 
twice as long as wide, with apical plate and pseudo- 
deltidium in rare specimens. Beak flat or slightly 
curved; interarea usually catacline. Cardinal extremi- 
ties acute, hinge forming widest part. Valves finely 
costate, about 20 costae on a side, the ones in cardinal 
extremities fine and often indistinct. Entire surface 
covered by fine radial lines and granules. 

Ventral valve gently convex in lateral profile; sulcus 
originating at beak, widening and deepening anteriorly 
to occupy about one-third the midwidth; sulcus bear- 
ing six or seven costae; tongue varying from suban- 
gular to moderately rounded. Flanks at cardinal ex- 
tremities flattened to slightly concave. 

Dorsal valve fairly evenly and moderately convex 
in lateral profile; anterior profile broadly and fairly 
strongly domed, the median apex bearing a small 
rounded knob representing fold. Fold originating at 
beak, narrow and only gently elevated above flanks, 
which are flattened in anterior view but gently swollen 
in lateral view. 

Ventral valve interior with long divergent dental 
plates extending for about one-third valve length. 

Measurements (in mm).— 


dorsal 
valve hinge mid- _ thick- 
length length width width ness 
USNM 201036a 
(Holotype; loc. 524z) ‘11.0 12.7 23.4* je).3} ES 
USNM 201036b 7.8 9.7 18.2* 14.8 11.8 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DuTRO 107 


dorsal 
valve hinge mid- _ thick- 
length length width width ness 


USNM 201053a 


(loc. 521h) 8.2 9.1 17.4* 13.4 9.1 
USNM 201000a 

(loc. 509u) 10.5 11.6 — 15.7 12.5 
USNM 203544 

(loc. 528)) 11.4 ile? 18.0+ 15.6 11.8 


Diagnosis.—Transverse, hemipyramidal Cyrtospi- 
rifer with long catacline interarea. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation; Contadero Formation. 

Localities.—Contadero Formation: 509t; 521f; 
Rhodes Canyon Formation: 509u; 521h; 523p; 524e, v 
(float), z; 528); 3054q. 

Types.—Holotype: USNM 201036a. Figured Para- 
types: 201000a, b, 201036b, 201053a, 203544. Unfig- 
ured Paratypes: USNM 201000b, 201053b. 

Discussion.—It was at first thought that this species 
is an adult form of Cyrtospirifer breviposticus Stain- 
brook (1947), which comes from the Rhodes Canyon 
Formation a mile south of the entrance to Rhodes Can- 
yon. Comparison of C. pyramidalis with Stainbrook’s 
type specimen (USNM 135257) shows only remote 
similarities. Both species are semipyramidal but that 
is where the similarity ends because Stainbrook’s 
species has rounded cardinal extremities and a hinge 
narrower than the midwidth. The sides of his species 
are well-rounded and the fold and sulcus are relatively 
wide. Although the two species apparently occur at 
the same stratigraphic levels, Stainbrook’s cannot even 
be construed as the young of C. pyramidalis because 
all of the immature specimens of that species are wide- 
hinged. Cyrtospirifer pyramidalis is different from C. 
posticus Johnson, Reso, and Stephens (1969) (not 
Stainbrook, 1947) in being smaller, wider, with acute 
cardinal extremities and a more steeply inclined ven- 
tral palintrope. 


Cyrtospirifer rotundus new species 
Plate 30, figures 1-23 


Spirifer whitneyi var. animasensis (Girty) Kindle, 1909, p. 25, pl. 
9, figs. 21-24. 


Small for genus, subquadrate in outline, width great- 
er than length; hinge usually widest part; cardinal ex- 
tremities acute or extended into small ears. Sides 
rounded, slightly oblique; anterior margin truncated 
narrowly; interarea long, gently curved, steeply ap- 
sacline. Delthyrium apically closed by a short plate. 
Surface finely costate, 18—20 costae on each flank, last 
three or four very small; costae flatly rounded, fairly 
wide and separated by spaces narrower than costae. 


Ventral valve semipyramidal, moderately convex in 
lateral profile but narrowly domed in anterior profile 
and having sides flatly convex, steep and becoming 
concave at the cardinal extremities. Sulcus originating 
at beak, narrow and shallow, occupying about one- 
third maximum width but producing a fairly long, nar- 
rowly rounded to subangular tongue. 

Dorsal valve fairly strongly convex in lateral profile 
and narrowly domed in anterior profile, the center oc- 
cupied by narrow knob representing fold. Flanks gently 
rounded and steep. Fold beginning at beak and wid- 
ening moderately anteriorly, fold fairly strongly ele- 
vated above flanks at anterior, and moderately round- 
ed. 

Measurements (in mm).— 


dorsal 
valve _— hinge mid- __ thick- 
length length width width ness 


USNM 201097b 


(loc. 527a) 13.7 12.2 20.0% 17.0 12.8 
USNM 201097a 16.4 15.1 18.4* Wed 15.0 
USNM 201096 

(Holotype; loc. 506) IS:3 1-7) 20.4* 16.9 14.8 
USNM 200911 

(loc. 3029) 11.8 11.4 24.8* 14.7 10.8 


Diagnosis.—Small, rather narrow, semipyramidal 
Cyrtospirifer having 18-20 costae on each flank. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—S06, b, q; 527a, p; 3029; USGS 509. 

Types.—Holotype: USNM 201096. Figured Para- 
types: USNM 201097a, b. 

Discussion.—This species is readily separated from 
Cyrtospirifer convexus n. sp. by its much narrower form, 
less extended hinge, less oblique lateral margins and 
more elevated fold. A similar appearing, but larger, 
species occurs in the West Range Limestone on Dutch 
John Mountain, north of Pioche, Nevada (USNM 
204956). 

Cyrtospirifer rotundus is similar to C. posticus 
Johnson, Reso, and Stephens (1969) in size and gen- 
eral appearance, but C. rotundus has a stronger, more 
rounded fold, shallower sulcus and more incurved 
beak. 


Cyrtospirifer cf. C. thalattodoxa Crickmay 
Plate 30, figures 24~29 


Cyrtospirifer thalattodoxa Crickmay, 1952, p. 601, pl. 72, figs. 1-7. 


Referred here are specimens of Cyrtospirifer having 
a wide and barely perceptible fold by virtue of lack of 
elevation of the flanks. Most of the specimens are poor, 
but the best one, a fairly young adult (USNM 200968), 
measures (in mm): length 13.7, dorsal valve length 14.0, 


BULLETIN 315 


hinge width 28.4 (points on both sides broken off), 
midwidth 27.0, thickness 14.8. The ventral valve is 
subconical in lateral profile and the interarea is long 
and only slightly curved near the beak. The dorsal 
valve is moderately convex and the fold is only slightly 
elevated and occupies about a third the valve width. 
About 20 pustulose costellae on a side. 

Diagnosis.—Wide-hinged, finely costate Cyrtospi- 
rifer with wide, scarcely elevated fold. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —S21\d, h. 

Types.—Figured Hypotype: USNM 200968. 


Cyrtospirifer whitneyi (Hall) 
Plate 30, figures 30-34 


Spirifer whitneyi Hall, 1858, p. 502, pl. 4, fig. 2; 1867, p. 245, 417; 
Hall and Clarke, 1893, p. 24, 27, pl. 30, figs. 18, 19. 


A few specimens of compact form and moderately 
elevated fold and sulcus are referred to Hall’s species. 
The specimens are all very poorly preserved. 

Diagnosis.—Cyrtospirifer with wide sulcus, wide 
fold moderately elevated above the flanks. 

Stratigraphic occurrence.—Sly Gap Formation; 
Contadero Formation. 

Localities. —Sly Gap: 509s, z; 521d, u. Contadero: 
509t; 521g; 523n, q; 526g. 

Types.—Figured Hypotype: USNM 201102. 


Cyrtospirifer species | 
Plate 31, figures 48-58 


About medium size for genus, roundly elliptical in 
outline, hinge narrower than greatest width at mid- 
valve; sides rounded, anterior margin gently emargin- 
ate; posterolateral margins forming angle of 90° in 
young specimens to 120° in old specimens. Cardinal 
extremities obtuse. Interarea long, fairly to strongly 
curved; beak suberect to strongly incurved. Surface 
finely costate, costae low, rounded and separated by 
spaces narrower than costae. Sixteen costae ona side. 

Ventral valve fairly strongly convex in lateral profile 
but fairly strongly domed in anterior profile, median 
region deeply indented by a wide sulcus, the flanks 
convex and steep. Sulcus originating at beak, wide and 
deep, occupying about half maximum width. Tongue 
fairly long and narrowly rounded. 

Dorsal valve moderately convex in lateral profile; 
broadly domed in anterior profile with flattish steeply 
descending flanks. Fold originating at beak, moderate- 
ly elevated at mid-fold to form a somewhat carinate 
but low elevation. 

Ventral valve with fairly long apical plate. 


Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- apical 
length length width width ness _ angle 


USNM 203535 
(loc. 522) 21.1 15.6 19.8 23.4 15.4 O58 
USNM 203536 2523; 18:9) 26:85) 321) 2033 IAS 


Diagnosis.—Elliptical Cyrtospirifer, anteriorly 
emarginate, with wide fold and sulcus and curved in- 
terarea. 

Stratigraphic occurrence.—Thoroughgood Forma- 
tion. 

Locality.—S522. 

Types.—Figured Specimens: USNM 203535, 203536. 

Discussion.—All of Greiner’s (1957) species from 
the New York Upper Devonian have acute cardinal 
extremities and usually are mucronate. Cyrtospirifer 
portae Merriam (1940) has some resemblance to this 
New Mexico species, but the Nevada form does not 
have the rounded sides and subcarinate fold. 


Cyrtospirifer species 2 
Plate 30, figures 47-50; plate 31, figures 43-47 


Medium-sized for genus, length equal to 60 percent 
of width; outline forming a wide pentagon; hinge wide, 
not equal to maximum width, which is just anterior to 
narrowly rounded cardinal extremities; anterolateral 
margins straight, strongly oblique to midline; anterior 
margin truncated; posterior margins broadly obtuse and — 
forming angle of 145°. Interarea fairly long, curved, 
gently apsacline; delthyrium questionably open. Sur- 
face finely costate, costae broadly rounded and sepa- 
rated by spaces narrower than the costae, which num- — 
ber about 22 on a side. 

Ventral valve gently convex in lateral profile, broad- — 
ly convex in anterior profile, the lateral slopes flat-— 
tened and forming a long, only moderately steep de- 
scent, median part having subangular depression. 
Sulcus originating at beak, widening anteriorly to oc- 
cupy somewhat less than one-third of width and broadly | 
V-shaped. | 

Dorsal valve gently convex in lateral profile, broadly 
and unevenly convex in anterior profile; median region — 
of profile subangular with sides sloping strongly, flat- | 
tened toward midvalve but becoming gently concave | 
toward extremities. Fold beginning at the beak, wid- 
ening rapidly anteriorly but merging laterally into 
slightly convex flanks at anterior, thus poorly defined. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 109 


Measurements (in mm).— 


dorsal maxi- 
valve hinge mum _ thick- apical 
length length width width ness angle 


USNM 200969 
(loc. 522) 20:4 16:8) 3220% 35.522) 1325) 145° 


Diagnosis. —Wide, somewhat compressed Cyrto- 
spirifer having a low fold but moderately deep V-shaped 
sulcus. 

Stratigraphic occurrence. 
tion. 

Locality.—522. 

Types.—Figured Specimens: USNM 200969, 205260. 

Discussion.—In its great width and compressed 
form, this species is unlike any other New Mexico 
form. It is entirely unlike any of the species of Cyr- 
tospirifer figured by Greiner (1957) from the Upper 
Devonian strata of western New York. 


Thoroughgood Forma- 


Cyrtospirifer species 3 
Plate 31, figures 11-16 


Small, wider than long with narrowly and laterally 
extended cardinal extremities. Sides forming a sig- 
moidal curve; anterior margin narrowly emarginate. 
Beak slightly incurved, interarea long and approxi- 
mately catacline. Delthyrium open. Costate, costae 
narrow and separated by spaces narrower than costae; 
21 costae on a side; twelve costae, finer than those on 
the flanks, marking fold; one less in sulcus. 

Ventral valve slightly deeper than dorsal valve, 
moderately convex in lateral profile, strongly and 
broadly domed in anterior profile; sulcus originating 
at beak, marked by costae finer than those of flanks; 
tongue short and rounded. Flanks moderately convex, 
moderately steep becoming flattened at cardinal ex- 
tremities. 

Dorsal valve moderately convex in lateral view, 
strongly domed in anterior view and with fold forming 
a knob on dome. Fold beginning at beak, rounded in 
cross section but not strongly elevated above flanks. 
Flanks having about same convexity as those of ven- 
tral valve, becoming flattened to slightly concave to- 
ward the cardinal extremities. 

Diagnosis. —Wide-hinged, short Cyrtospirifer, fine- 
ly costate with finer costae on the fold and sulcus than 
on the flanks. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality.—521d. 

Types.—Figured Specimen: USNM 201023. 

Discussion.—This species is most like C. convexus 
from the Percha Formation, but it differs in having a 
more acute ventral tongue, narrower cardinal extrem- 
ities, more swollen flanks. We have only a single spec- 
imen of this distinctive species. 


Cyrtospirifer species 4 
Plate 39, figures 38—42 

A single specimen of about average size for the ge- 
nus measures (in mm): length 16.5, dorsal valve length 
17.0, hinge width ca. 33.0 (based on half measure), 
midwidth 24.0, thickness 16.0. The specimen has a 
long, procline, slightly curved interarea. The sides are 
oblique and slightly concave to the prominent ear(s). 
The ventral valve is moderately convex and somewhat 
pyramidal; the dorsal valve is gently convex in lateral 
profile and broadly domed in anterior profile. The sul- 
cus of the ventral valve is broad and fairly deep, 
broadly V-shaped in profile. The fold is low and 
scarcely defined. The costae are narrow and fairly even 
over the whole surface, about 20 on each flank and 15 
on the fold. The anterior commissure is broadly uni- 
plicate and slightly angulated in the middle. Surface 
finely pustulose. 

Diagnosis.—Medium-sized Cyrtospirifer having a 
poorly defined fold but well-marked sulcus and fairly 
uniform costae over the whole surface. 

Stratigraphic occurrence.—Sly Gap Formation. 

Locality. —USGS 543. 

Types.—Figured Specimen: USNM 207696. 

Discussion.—This species is unlike any figured by 
Greiner (1957) from the Upper Devonian rocks of New 
York. The combination of a procline interarea and 
faintly developed fold seem to be unique in the genus. 
The species is reminiscent of C. thalattodoxa Crick- 
may (1952), but that species has a more prominent fold 
and the interarea is not procline. 


Cyrtospirifer species 5 
Plate 39, figures 43, 44 

A small Cyrtospirifer from the Caballos Mountains 
is represented by three valves, two ventral and one 
dorsal, all impressions in a fine-grained sandstone. The 
specimens are wider than long and with slightly mu- 
cronate cardinal extremities. The valves are orna- 
mented by low broadly rounded costae separated by 
narrow interspaces. The sulcus is shallow and is oc- 
cupied by a median costa and two lateral ones. The 
dorsal valve fold is narrow and low, demarcated by 
narrow grooves on each side and marked by four cos- 
tae. The entire surface is strongly granulose. The only 
measurable specimen (USNM 207703a) is 12 mm long, 
and about 25 mm wide (based on the half measure). 

Diagnosis.—Small, wide, alate Cyrtospirifer with 
subdued fold and sulcus. 

Stratigraphic occurrence.—Sly Gap? (uppermost 
sandstone). 

Locality.—522s. 

Types.—Figured Specimens: USNM 207703a, b. 

Discussion.—We have been unable to find a de- 
scribed species like this one. It is possible that these 


110 BULLETIN 315 


specimens represent youthful forms of a normally larg- 
er species. Close check of all the specimens from the 
Sly Gap Formation in the San Andres and Sacramento 
Mountains failed to bring to light any specimens like 
these or any that might be larger adults. 


HISPIDARIA new genus 
[Latin, hispidus = bristly] 


Type species.—Hispidaria posterogranulosa n. sp. 

Diagnosis.—Small, strongly costate, wide-hinged 
cyrtospiriferoids having the bristly to granular orna- 
ment which also occupies most of the interarea. 

Discussion.—This is a small and obscure group, re- 
lated to Cyrtospirifer Nalivkin (in Fredericks, 1924), 
that occurs in the Famennian of New Mexico. The 
shells are fairly strongly costate and the costae are 
densely covered by coarse granules that are often 
lengthened into spines. This micro-ornament is not 
confined to the costate part of the shells but covers 
the parts of the interarea adjacent to the posterior mar- 
gin and extending in a dorsad direction nearly to the 
interarea of the dorsal valve. 


Hispidaria posterogranulosa new species 
Plate 32, figures 22-34 


Small, wider than long, with widely extended hinge 
and acute cardinal extremities. Sides strongly oblique. 
Interarea long, strongly apsacline, divisible into two 
parts, one vertically striated part surrounding the del- 
thyrium and a granulose or bristly part between the 
striated portion and the posterior margin. Delthyrium 
open except for a tiny apical plate. Surface completely 
costate, costae elevated, narrowly rounded and sep- 
arated by spaces nearly equal to the width of costae; 
12 costae on a side in adult. Micro-ornament consist- 
ing of coarse, crowded granules and a few rows of 
spines in the sulcus and on the fold. 

Ventral valve moderately convex in lateral profile 
and broadly triangular in anterior profile, apex being 
indented by sulcus that originates at beak and occupies 
about one-quarter maximum width. Sulcus extended 
into a short angular tongue. Sulcus occupied by one 
costa at apex dividing almost immediately after its ap- 
pearance; other costae intercalated, primary pair bear- 
ing spines. Flanks flattened, moderately steep. 

Dorsal valve very gently convex in lateral profile 
and very broadly and flatly triangular in anterior pro- 
file, the apex of the triangle marked by a rounded knob. 
Flanks barely convex and gently sloping. Fold origi- 
nating at the beak, narrowly rounded, moderately el- 
evated anteriorly and marked initially by two costae. 
Initial two costae forming sides of fold; inserted inside 
them is a large anteriorly bifurcating costa that bears 
minute spines. 


Measurements (in mm).— 


dorsal 
valve _ hinge mid- _ thick- 
length length width width ness 


USNM 201106 
(Holotype; loc. 
3026a) 10.0 8.6 18.0* 13.6 6.7 


Diagnosis.—Small, wide-hinged Hispidaria with in- 
terarea partially granulose. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Locality.—3026a; 3042. 

Types.—Holotype: USNM 201106. Figured Para- 
type: USNM 201109. 

Discussion.—This species is unlike any Cyrtospiri- 
fer known to us. It differs from H. spinosa n. sp., 
which also has a partially granulose interarea, in its 
greater size, greater width, and strongly oblique lateral 
margins. This is a very rare species. 


Hispidaria spinosa new species 
Plate 32, figures 1-21 


Syringospira prima Cooper (not Kindle, 1909), 1954, pl. 37, figs. 
15-18. 


Small, cyrtiniform, with hinge forming widest part; 
cardinal extremities acute; sides oblique, nearly straight 
to gently rounded; anterior margin truncated. Inter- 
area long, apsacline, portion bounding posterolateral 
margin strongly granulose or spinose. Delthyrium with 
short apical foramen. Each flank occupied by 11 or 12 
moderately elevated, rounded costae separated by 
spaces slightly less wide than costae; flanks occasion- 
ally with intercalated costae. Micro-ornament consist- 
ing of mat of coarse granules that are often extended 
into minute spines, on costae and ventral valve inter- 
area. 

Pedicle valve gently convex, most so in anterior half 
in lateral profile; anterior profile forming a fairly high 
triangle with fairly steep sides and apex slightly de- 
pressed by the sulcus. Sulcus originating at beak with 
single costa that divides early. Sulcus moderately deep, 
occupying about a third of width, and usually occupied 
anteriorly by five costae. Tongue short and rounded. 

Dorsal valve fairly evenly and gently convex in lat- 
eral profile, broadly and very gently convex in anterior 
profile. Fold originating at beak, low throughout its 
extent but moderately elevated anteriorly. Fold con- 
sisting of one costa at apex, bifurcating close to the 
apex, the two costae thus formed bounding the sides 
of fold; usually three costae intercalated between the 
primary two. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DuTRO ete 


Measurements (in mm).— 


dorsal 
valve hinge mid- __ thick- 
length length width width ness 


USNM 201126 


(Holotype; loc. 527d) 8.6 6.9 1322% 9.8 5.2 
USNM 201108 

(loc. 527a) 8.4 7.4 10.8* 9.0 S)6) 
USNM 203512 

(loc. 527e) 9.4 8.3 12.8*? 10.8 6.4 
USNM 201125 

(loc. 3026a) 8.8 8.0 16.6* 11.7 7.3 
USNM 201107 

(loc. 506q) 7.0 Si5// 10.6 8.2 4.3 


Diagnosis.—Small, cyrtinoid Hispidaria with very 
coarse granules and spines. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities.—S06, q; 507h, n, u; 509a; 527a, d, e; 
3026, a; 3042. 

Types.—Holotype: USNM 201126. Figured Para- 
types: USNM 201107a, 201108, 201125. Unfigured 
Paratypes: USNM 201107b, 203512. 

Discussion.—Like Hispidaria posterogranulosa, n. 
sp., this one has a granulose outer part of the inter- 
area. It differs from that species in being smaller, nar- 
rower, with less oblique sides, and an outline and pro- 
file like Cyrtina Davidson (1858). The micro-ornament 
of H. spinosa is also stronger than that of H. poster- 
ogranulosa and is very prominent for such a small 
shell. This is also a very rare species. 


CYRTIORINA new genus 


Large, wider than long, hinge less than maximum 
shell width at midvalve. Unequally convex, ventral 
valve more convex. Anterior commissure strongly uni- 
plicate; ventral tongue long. Beak strongly incurved 
over moderately long, concave, apsacline interarea. 
Delthyrium covered by convex, often thick, pseudo- 
deltidium. Surface completely covered by costellae 
rounded posteriorly, flattening anteriorly, and becom- 
ing marginally obsolete in old shells. Costellae covered 
by minute concentric and radial lines that form fine 
granules at their intersection. 

Ventral valve interior with apical plate, often thick- 
ened; dental plates reaching nearly to midvalve. Dor- 
sal valve interior with narrow socket plates supporting 
long, bowed crura. 

Type species.—Cyrtospirifer kindlei Stainbrook, 
1947, p. 318, pl. 44, figs. 1, 2, 7-12. 

Diagnosis.—Large, costellate, uniplicate Cyrtospi- 
riferidae with pseudospondylium and apical plate. 

Discussion.—Stainbrook’s holotype is a somewhat 
distorted specimen having an extended point on one 
side. In all of our collections, and among Stainbrook’s 


unfigured paratypes, not one has lateral points at the 
cardinal extremities. Except for its cardinal extension, 
Stainbrook’s type has all other features conforming to 
those of the majority as outlined above. 

Cyrtiorina is externally similar to Uchtospirifer 
Lyaschenko (1957) but differs in having an apical plate. 
It is like Cyrtiopsis Grabau (1923) in having a pseu- 
dospondylium, often showing traces of a round fora- 
men, a feature of Cyrtiopsis. The latter, however, has 
no apical plate. Dmitria Sidiachenko (1961), a Russian 
genus, resembles Cyrtiorina in its fine costellae and 
large size, but differs in having barely perceptible fold 
and sulcus. 

Spirifer monticola Haynes (1916), of the Three Forks 
Formation in Montana, is another species of Cyrtio- 
rina. 

Cyrtiorina kindlei (Stainbrook) 
Plate 32, figures 35-41; plate 41, figures 50-53 


Spirifer whitneyi Kindle, 1909 (not Hall, 1858), p. 24, pl. 8, figs. 2— 

Sa. 

Cyrtospirifer kindlei Stainbrook, 1947, p. 318, pl. 44, figs. 1, 2, 7- 

122 

This large and distinctive brachiopod is one of the 
commonest species in the Percha. It occurs from up- 
permost nodular beds of the Ready Pay Member 
throughout the succeeding Box Member. 

Stratigraphic occurrence.—Percha Formation 
(Ready Pay Member, Box Member). 

Localities.—506, a, b, p, q; 507a, b, d, e, f, g, h, n, 
0, q, r, t, v, x; 509a, c, d; 527a, c, d, e, f, g, n; 3026, 
a; 3029; 3030, 3031a, b; 3034; 3035; 3042, s; USGS 
(Kirk) 459; 509; 512. 

Types.—Holotype M.A.S. 1591 (=USNM 135286). 
Paratypes: M.A.S. 1519A (=USNM 135237); M.A.S. 
1591B (=USNM 135237); M.A.S. 1591C (=USNM 
135283), figured in Stainbrook (1947). Spirifer whitneyi 
Kindle (not Hall) figured by Kindle, 1909, USNM 
62029a, b; 62031. Figured Hypotypes: USNM 201150a; 
201151a, 1; 203528a, c; 203529b, c. 


Genus UCHTOSPIRIFER Lyaschenko, 1957 


Uchtospirifer? rarus new species 
Plate 33, figures 1-8 


Small for genus, quadrate in outline, about 0.8 as 
long as wide; hinge wide but not quite equal to the 
maximum valve width, which is located at about mid- 
valve; cardinal extremities forming small ears. Sides 
fairly well rounded; anterior margin truncate; postero- 
lateral margins forming an angle of 110°. Interarea 
moderately long, curved, orthocline; flattened pseudo- 
deltidium present. Beak strongly incurved over inter- 
area. Anterior commissure strongly uniplicate. Sur- 
face costate, each flank bearing about 13 narrowly 


BULLETIN 315 


rounded costae separated by spaces nearly as wide as 
costae. Sulcus occupied by four costae and fold by 
five. Strong concentric lamellae appearing at the an- 
terior. Micro-ornament not preserved. 

Ventral valve strongly convex in lateral profile and 
most so in posterior part; anterior profile broad, evenly 
convex dome with moderately sloping sides but ex- 
cavated medially by the sulcus. Sulcus originating at 
beak, deepening and widening anteriorly to occupy 
about 60 percent of width. Sulcus occupied by one 
costa near the beak, others being intercalated ante- 
riorly. Flanks gently convex and steep-sided. Tongue 
long, rounded. 

Dorsal valve less convex and less deep than the ven- 
tral valve in lateral profile; anterior profile forming 
broad triangle, with fold forming low bulge at apex. 
Fold initiating at beak narrowly rounded, moderately 
elevated above flanks anteriorly. Fold initially with 
two costae that bifurcate not far anterior of beak; other 
costae intercalated. 

Measurements (in mm).—USNM 200989 (Holo- 
type): length 15.6, dorsal valve length 12.2, hinge width 
15.2, maximum width 17.7, thickness 11.2, apical an- 
gle 110°. 

Diagnosis. —Small Uchtospirifer? having 13 costae 
on each flank, a flattened pseudodeltidium, and narrow 
fold and sulcus. 

Stratigraphic occurrence.—Contadero Formation. 

Locality.—523q. 

Types.—Holotype: USNM 200989. 

Discussion.—This species is doubtfully placed in 
Uchtospirifer even though it has a pseudodeltidium. 
It is completely unlike Cyrtiorina kindlei (Stainbrook, 
1947) in size and ornament. 


Genus SYRINGOSPIRA Kindle, 1909 
Syringospira prima Kindle 


Syringospira prima Kindle, 1909, p. 29, pl. 7, figs. 8-8d; pl. 8, figs. 
l-la. Cooper in Shimer and Shrock, 1944, p. 327, pl. 125, figs. 
12-14; Cooper, 1954, p. 326, pl. 36, figs. 1-7, 11; pl. 37, figs. 1- 
14 (not figs. 15-18). 


This is an uncommon species in the Percha For- 
mation (Box Member), and is also found in the upper 
part of the black shale (Ready Pay Member) at the 
appearance of the first of the calcareous nodules. It is 
commonly enclosed partially or wholly by the nodules. 
It also occurs in all parts of the calcareous, greenish 
nodular shales of the Box Member but is scattered and 
often rare. There too, it may be enclosed by a nodule. 

Cooper (1954) described this species in detail. 


Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —Percha Fm. (Box Mbr.): 506, a, b, p, 
Gp 20/ab-wenG.e, hi, J, mMs.0; plxerz 5 09amIbaNes 


527a, d, e, f, g, j, k, p; USGS 4804; 3026, a; 3031a, b; 
3034; 3042, s. 

Types.—Holotype: USNM 62027. Figured Para- 
type: USNM 62028; primary types figured by Kindle 
(1909). Hypotypes figured by Stainbrook (1947): 
M.A.S. 1582A, B (=USNM 135239a, b). Hypotypes 
figured by Cooper (1954): USNM 123358; 123359a—c; 
12336la; 123363a; 123364; 12336Sa, b; 123366a, b; 
123367; 123368a; 123369a; 123370a. 


Genus TENTICOSPIRIFER Tien, 1938 


Tenticospirifer cf. T. cyrtinaformis (Hall and Whitfield) 
Plate 33, figures 9-15 
Spirifera cyrtinaformis Hall and Whitfield, 1872, p. 238, pl. 11, figs. 
21-24. 
Platyrachella cyrtinaformis (Hall and Whitfield) Fenton and Fenton, 
1924, p. 162, pl. 29, figs. 1-7. 


This species is represented by eight specimens from 
four localities. The largest, best, and illustrated spec- 
imen is about normal size for the species. It is widest 
at the hinge and the sides slope gently medially. The 
ventral valve is semipyramidal, the interarea long and 
catacline, the delthyrium narrow. The dorsal valve is 
gently convex and the flanks slope to cardinal extrem- 
ities. The fold is poorly defined but is best seen at the 
anterior commissure, which is uniplicate. The surface 
is marked by fine costae separated by spaces of about 
equal width. 

Measurements (in mm).—USNM 201105: length 
10.6, dorsal valve length 11.3, hinge width 16.6 (not 
complete), midwidth 12.5, thickness 10.8. 

Diagnosis.—Finely costate, medium-sized Tenti- 
cospirifer having a poorly developed fold and sulcus. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506j; 509q; 521d; 526x. 

Types.—Figured Hypotype: USNM 201105. 


Genus INDOSPIRIFER Grabau, 1931 


Indospirifer anomala new species 
Plate 33, figures 45-54 


About medium-sized for genus, transversely rect- 
angular in outline; hinge forming widest part and car- 
dinal extremities an acute angle, about 85°. Sides gently 
rounded and sloping slightly toward midvalve; anterior 
narrowly truncated. Interarea wide and long (in the 
direction of the valve length), curved and strongly ap- 
sacline. Beak small and strongly incurved. Valves 
multicostate, each flank having 11 or 12 narrowly 
rounded costae with interspaces slightly wider than 
costae; fold with initial costa that divides anteriorly 
into two that form the crest of the fold; sides of fold 
having two or three costae; costae in sulcus variable. 

Ventral valve moderately convex in lateral profile 
but with umbo strongly curved; anterior profile broad- 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO iS 


ly convex and median region gently depressed by sul- 
cus. Flanks bounding sulcus moderately swollen; sul- 
cus originating at beak, widening anteriorly to occupy 
slightly less than half hinge width; tongue long and 
anteriorly rounded. 

Dorsal valve flatly convex in lateral profile but with 
umbo slightly more curved; anterior profile forming a 
broad triangle with the fold forming crest and flattened 
flanks sloping laterally. Fold originating at beak, nar- 
rowly rounded and occupying one-third or somewhat 
less of hinge width. Fold strongly elevated above flanks 
only in anterior half. 

Measurements (in mm).— 


dorsal 
valve hinge mid-__ thick- 
length length width width ness 


USNM 200905 

(loc. 509z) 21.0 16.9 22.0? 26.4 IS 
USNM 200904 

(Holotype; loc. 506j) 23.8 18.0 33.0 29.2 16.8 


Diagnosis.—Indospirifer of medium size having a 
rounded fold, slightly acute cardinal extremities, and 
11-12 costae on each flank. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—506j; 509s, z; 521d, q; 524m, w. 

Types.—Holotype: USNM 200904. Figured Para- 
type: USNM 20090S. 

Discussion.—This species differs from J. orestes 
(Hall and Whitfield, 1872) and J. exfoliata n. sp. in its 
larger size, rounder fold, less acute cardinal extremi- 
ties and in having more costae on the flanks. This is a 
rare species in the New Mexico Frasnian. 


Indospirifer exfoliata new species 
Plate 33, figures 21-34 


Spirifer strigosus Stainbrook (not Meek, 1876), 1948, p. 777, pl. 2, 
figs. 5-8. 


Small- to medium-sized for genus, wider than long, 
semielliptical in outline; hinge forming widest part; 
cardinal extremities usually acute. Anterior commis- 
sure narrowly uniplicate. Interarea wide, short in di- 
rection of valve length, moderately curved and apsa- 
cline. Beak strongly incurved. Surface multicostate, 
each flank marked by eight costae, the narrow fold by 
two prominent summit costae and one on each slope; 
sulcus marked by strong median costa and variable 
number of lateral subsidiary ones. 

Ventral valve moderately convex in lateral profile 
but broadly convex in anterior profile, crest of curve 
indented medially by shallow sulcus. Sulcus originat- 
ing at beak, widening and deepening anteriorly; sulcus 
width equalling less than half hinge width; flanks 
bounding sulcus swollen and steep. 

Dorsal valve having about same depth as ventral 


valve; broadly convex, middle with a narrow bulge 
representing fold; gently convex in lateral profile; flanks 
flattened, sloping moderately. Fold originating at beak 
and formed of one costa that divides at beak to form 
crest of fold; additional costae added on steep slopes 
of fold. 

Measurements (in mm).— 


dorsal hinge 
length valve width mid- thick- L/W 
(L) length (W) width ness_ ratio 


USNM 200881 
(loc. 506c) 

USNM 200882 S27) 1229) 120102 21k0 6 O75 
(Holotype; loc. 
S06c) 17.1 14.0 22.3 21.0 12.9 0.81 


Diagnosis.—Small to medium Indospirifer with nar- 
row fold and sulcus. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities.—S06c; 509g, i, k, m, w; 521d, 1, n, r; 
524n; 526q. 

Types.—Holotype: USNM 200882. Figured Para- 
type: USNM 200881. 

Discussion.—This species is clearly related to In- 
dospirifer orestes (Hall and Whitfield, 1872) as it has 
the same type of narrow fold crested by two costae 
with subsidiary ones appearing on the slopes. How- 
ever, it is smaller, with a less broad interarea and a 
more strongly incurved ventral beak. 

Stainbrook (1948) referred specimens of this species 
to I. strigosus (Meek, 1860) but the latter is a large 
species with medially sloping sides, rather acute car- 
dinal extremities (63°), and a proportionately narrower 
fold and sulcus. Furthermore, J. exfoliata has no bi- 
furcating lateral costae. 


Indospirifer aff. I. orestes (Hall and Whitfield) 
Plate 33, figures 16-20 


Spirifer orestes Hall and Whitfield, 1872, p. 237, pl. 11, figs. 16~20. 


Two specimens having characters of some examples 
of Indospirifer orestes was taken from Capitol Peak in 
the San Andres Range (locality 521k, u). Measure- 
ments (in mm) of one (USNM 200990) are: length 16.0, 
dorsal valve length 11.4, hinge width 13.2, maximum 
width 18.3, thickness 10.3. The ventral valve is strong- 
ly umbonate and the beak overhangs the long curved 
apsacline interarea. The cardinal extremities and sides 
are well-rounded. The specimen suggests the one fig- 
ured by Hall and Whitfield (1872, pl. 11, fig. 16; re- 
pository unknown), which has well-rounded sides. 
Fenton (1931, p. 138) in his discussion of the Spirifer 
orestes Phratry mentions the similarity of Hall’s spec- 
imen (fig. 16) to S. euginus Fenton (1931). Spirifer 
euginus repositus Fenton (1931), which Fenton illus- 
trated as figure B on p. 201, is closest to the Capitol 


114 BULLETIN 315 


Peak specimen. This specimen has strongly rounded 
sides and a narrow hinge and thus is, in these gross 
characters, like the New Mexico form. Unfortunately, 
the Sly Gap species does not retain any of the fine 
ornament, thus negating a more positive identification. 
Stratigraphic occurrence.—Sly Gap Formation. 
Localities. —S52\k, u. 
Types.—Figured Hypotype: USNM 200990. 


Indospirifer strigosus (Meek) 
Plate 33, figures 35-44 


Spirifer macra Meek (not Hall, 1857), 1860, p. 309. 

Spirifera strigosa Meek, 1860; note appended to extras of the above 
(see Meek, 1877). 

Spirifer strigosus Meek, 1876, p. 347, pl. 1, fig. 5. 

Spirifer (Trigonotreta) strigosus Meek, 1877, p. 43, pl. 3, fig. 5. 

Not Spirifer strigosus Meek. Stainbrook, 1945, p. 52, pl. 5, figs. 14— 
21; 1948, p. 777, pl. 2, figs. 5-8. 


Inasmuch as “‘Spirifer” strigosus Meek has been 
identified in the Independence and Sly Gap faunas, it 
is not amiss to make a few remarks on the type spec- 
imens of that species and to point out the important 
differences between them and the specimens erro- 
neously identified. The type lot of S$. strigosus Meek 
is preserved in the U.S. National Museum of Natural 
History as specimens USNM 629a and 629b. Both 
specimens have been badly decorticated and mutilat- 
ed: 629a, the holotype, lacks much of the posterior 
half, and one side of the dorsal valve. Both of these 
parts have been restored in beeswax; the restoration 
is not very accurate and the shell is now badly scarred 
and worn. Specimen 629b lacks most of the anterior 
half of both valves. The beak and umbonal region of 
the ventral valve have been restored in beeswax. 

The unrestored parts of the holotype indicate a 
specimen with hinge wider than the midwidth and with 
lateral margins sloping strongly medially, thus empha- 
sizing the angularity of the cardinal extremity, which 
is about 62°. The fold at the anterior of the holotype 
is narrow and strongly elevated, occupying about 45 
percent of the width. As shown by the paratype, the 
fold originates on the umbo and is formed of two cos- 
tae that extend on the crest of the fold to the anterior 
margin. Two additional costae are added anteriorly on 
the fold slopes. The sulcus has one large median costa 
and two secondary costae on each side. Each flank of 
the holotype has nine or ten costae, the tenth obscure 
and questionable. 

Stainbrook (1945, p. 53) speaks of bifurcated costae 
in S. strigosus Meek, but the holotype shows no such 
costae on the dorsal valve, perhaps because the pos- 
terior half of the dorsal valve is restored in wax. On 
the ventral valve, a costa is intercalated near the beak 
but none of the other costae show bifurcation. On the 


other hand, the paratype has bifurcation of the costae 
immediately adjacent to the fold on the left side, and 
on the right side of the fold a fascicle of three costae 
appears. Bifurcation of the costae is rare in /. orestes 
(Hall and Whitfield, 1872). Stainbrook figures bifur- 
cation on specimens referred to S. strigosus, but the 
shape of the specimens and the bifurcation are unlike 
either of Meek’s specimens. This suggests that S. stri- 
gosus Meek occurs neither in the Independence Shale 
of Iowa nor in the Sly Gap of New Mexico. 

Stratigraphic distribution. —Upper Devonian. 

Locality.—Neils Valley, east of Eureka, Nevada. 

Types.—Holotype: USNM 629a; Figured Paratype: 
USNM 629b. 


Superfamily RETICULARIACEA Waagen, 1883 
Family RETICULARIIDAE Waagen, 1883 
Genus WARRENELLA Crickmay, 1953 


Warrenella magna new species 
Plate 34, figures 1-19 


Large, broadly elliptical in outline; greatest width at 
midvalve; sides strongly rounded; anterior margin 
truncated; posterior with ventral umbonal region 
strongly protuberant beyond posterior margin of dor- 
sal valve. Hinge wide, narrower than midwidth and 
from 0.6 to 0.75 of midwidth. Beak strongly incurved; 
interarea fairly long (in direction of valve length), 
curved, orthocline to strongly apsacline. Delthyrium 
covered by moderately convex apical plate. Surface 
covered by closely crowded concentric lines of minute 
single-barreled spines, about 12 to the mm. 

Ventral valve moderately convex in lateral profile 
but having the umbonal region narrowly convex; an- 
terior profile strongly and broadly convex and having 
the median region marked by a fairly deep depression 
representing sulcus. Umbonal region and flanks swol- 
len; sulcus bounded by a fairly strong plication that 
merges laterally into swollen flank. Sulcus originating 
five to ten mm anterior to beak, shallow but widening 
anteriorly to equal 40 to 45 percent of midwidth. 
Tongue moderately long, broadly rounded to truncat- 
ed at its free end. 

Dorsal valve gently convex in lateral profile but with 
umbonal region forming most convex part; anterior 
profile broadly triangular, fold forming the apex of the 
triangle and flattened flanks forming the two sides. 
Umbonal region narrowly swollen, swelling extending 
into fold, which is elevated above flanks at about mid- 
valve, continuing to heighten anteriorly; flanks de- 
pressed or slightly deflected in ventrad direction to- 
ward the cardinal and anterolateral extremities; but 
gently swollen between. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 115 


Ventral valve interior with strong dental plates. 
Measurements (in mm).— 


maxi- 
dorsal mum thick- 
length valve hinge width ness L/W T/W 
(L) length width (W) (T)_ ratio ratio 


USNM 200900 


(Holotype; 
loc. 523d) AST S910 3726) 58-20 Sil 10835 10!55 
USNM 200901a 
(loc. 523h) 39.1 31.6 37.0 48.5 26.6 0.80 0.55 


USNM 200901b 43.7 34.7 39.8 53.7 38.8 0.81 0.73 
USNM 200902 


(loc. 523b) 34.4 28.5 29.0 41.8 21.0 0.82 0.48 
USNM 203514 44.4 35.0 38.9? 54.3 32.6 0.89 0.60 
USNM 203515a 

(loc. 523f) 36.3 29.8 32.0 42.4 244 0.86 0.58 
USNM 2035 16a 

(loc. 523)) So 27M 278) ZOO BOY OL O52 
USNM 203513a 

(loc. 526i) 36.7 31.4 33.4 48.0 21.6 0.76 0.46 


Diagnosis.—Large Warrenella with length about 80 
percent of the width, and fine concentric lines with 
about 12 spines to the mm. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—523a, b, d, f, g, h, j, k, 0, s, t; 526i. 

Types.—Holotype: USNM 200900. Figured Para- 
types: USNM 200901a, 200902, 201129a, b, c. Unfig- 
ured Paratypes: USNM 200901b-e, 201129d-h. 

Discussion.—This species resembles Warrenella 
occidentalis (Merriam, 1940) and W. timetea Crick- 
may (1960). It differs from the former in having a less 
protuberant beak of the ventral valve, a deeper sulcus, 
stronger fold, and much wider hinge. Warrenella mag- 
na differs from W. timetea in being more transverse, 
in having finer concentric lines, narrower fold and sul- 
cus, and less strongly protuberant ventral valve beak. 
Warrenella franklini (Meek, 1868) is more elongate, 
deeper, with a less pronounced fold, narrower sulcus, 
and proportionately wider hinge. 

Warrenella magna is sufficiently abundant to give 
some idea as to how the deltidial structure forms. In 
the adult condition, this plate simulates a pseudodel- 
tidium, but it is formed initially of two oblique plates 
on the margins of the delthyrium. The pedicle evi- 
dently was protruded near the apex of the delthyrium 
between these two plates. Anteriorly, these plates grew 
together leaving an opening for the pedicle. In time, 
this pedicle opening was plugged and the pedicle be- 
came no longer functional. Nearly all specimens dis- 
play this plugged foramen. The deltidial structure an- 
terior to the plug shows continuous concentric growth 
lines anterior to the plug, and for some distance an- 
teriorly, similar to the henidium of Cloud (1942, p. 12). 


Warrenella floweri new species 
Plate 35, figures 39-54; plate 42, figure 26; 
plate 43, figures 10-22 


About medium-sized for genus, roundly quadrate in 
outline. Hinge narrower than widest part, which is at 
midwidth; sides broadly rounded; anterior margin 
truncated to slightly emarginate; ventral valve poste- 
rior moderately protuberant beyond the dorsal umbo; 
interarea moderately long, curved, orthocline to ap- 
sacline. Valves unequally deep, ventral valve much 
deeper than dorsal one; anterior commissure unipli- 
cate. Surface marked by closely crowded concentric 
lines bearing minute single-barreled spines, about 14 
to the mm. 

Ventral valve with fairly strongly convex lateral 
profile and a strongly curved umbonal region; anterior 
profile strongly convex and with median region de- 
pressed by sulcus. Flanks bounding sulcus inflated; 
sulcus originating about ten mm anterior to beak, wid- 
ening and deepening anteriorly to occupy somewhat 
less than half midwidth. Tongue moderately long and 
narrowly rounded. 

Dorsal valve gently convex in lateral profile, most 
convex in posterior half; anterior profile broadly con- 
vex, fold not affecting median region; flanks gently 
inflated, sloping moderately. Umbonal region slightly 
swollen; median region moderately inflated; fold 
scarcely visible posterior to midvalve, low and gently 
rounded in anterior third. 

Ventral valve interior having small teeth, thin but 
strong, subparallel dental plates; muscle area narrow, 
partially enclosed by dental plates. 

Measurements (in mm).— 


dorsal 
valve hinge mid- — thick- 
length length width width ness 


USNM 200883b 

(Holotype; loc. 5230) 31.9 25.6 25.6 33.3 21.8 
USNM 200883a 22S 23.4 733.3) 34.3 20.0 
USNM 200883c 30.0 24.0 fee? 33.6 20.6 
USNM 200883d 26.6 22.6 20.6 30.5 16.9 
USNM 200883e 33.0 26.0 33.8 40.4 24.3 
USNM 200883f 8.0 Weil 6.0? 9.4 4.6 
USNM 200923a 

(loc. 524x) 27.8 22.6 Del 32.6 20.4 
USNM 200923b BileS 252 27.0 36.0 PMS) 
USNM 200923c 30.0 23.6 26.4 33.2 22.8 


Diagnosis.—Medium-sized, wide-hinged Warrenel- 
la with rotund appearance and fold conspicuous only 
at anterior. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—506d; 523e, i, k, 0; 524x. 

Types.—Holotype: USNM 200883b. Figured Para- 


116 BULLETIN 315 


types: USNM 200883a, c, d, 200923a. Unfigured Para- 
types: USNM 200883e-f, 200923b-d. 

Discussion.—Warrenella eclectea Crickmay (1953), 
type species of the genus, is of about the same size as 
that of W. floweri n. sp., but it has narrowly rounded 
sides, a more pronounced fold and sulcus, and more 
sharply defined ridges bounding the sulcus on the ven- 
tral valve. Warrenella floweri differs from W. labre- 
quei Crickmay (1960) in having less narrowly rounded 
sides, narrower hinge, more truncated anterior, short- 
er interarea, and a more rounded, less protuberant 
beak. 

Formerly, Warrenella floweri was common at the 
Nackaye ‘‘Paint Mine’’ near Nackaye mine in the Ca- 
ballos Mountains. The specimens came from a tem- 
porary excavation and are not now so abundant. The 
species is rare in the Mud Springs Mountains west of 
Truth-or-Consequences, New Mexico. 

For a discussion of species of Warrenella and their 
stratigraphic meaning, see Ludvigson and Perry (1975). 


Genus TORYNIFERELLA Weyer, 1967 


Toryniferella spinosa (Kindle) 
Plate 42, figures 1-10 
Reticularia spinosa Kindle, 1909, p. 27, pl. 9, figs. 7-9a. 
Torynifer spinosus (Kindle) Stainbrook, 1947, p. 325, pl. 47, figs. 

38-42. 

Stainbrook (1947) referred this species to Torynifer 
but it lacks the median septum and apical chamber 
characteristic of Torynifer. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —S06, a, b, p, q; 507a, c, d, h, k, m, 0, 
u, z; 509a, b, c, e; 527d, e, f, g; 3026, a; 3029; 3034; 
3035; 3042, s; USGS 508, 512. 

Types.—Holotype: USNM 62034. Figured Para- 
type: USNM 62035, primary types figured by Kindle 
(1909). Hypotypes figured by Stainbrook (1947): 


Family ELYTHIDAE Fredericks, 1919 (1924) 
Genus ELITA Fredericks, 1918 


Elita magister new species 
Plate 36, figures 45—55 


Not Elytha diversa Stainbrook, 1948, p. 777, pl. 2, figs. 21, 22. 


Large, subrectangular in outline, subequivalve, ven- 
tral valve slightly deeper. Sides strongly rounded; an- 
terior margin truncated. Anterior commissure strongly 
uniplicate. Beak small, incurved. Interarea long, near- 
ly flat, strongly apsacline. Fold and sulcus without 
costae; flanks costate, costae low and broad, separat- 
ed by narrow interspaces; six or seven costae on each 
flank. 

Ventral valve gently convex in lateral profile but 
with umbonal region more strongly convex than rest 
of profile; anterior profile moderately and broadly con- 
vex, median region narrowly depressed by sulcus. 
Umbonal region narrowly swollen; posterolateral 
slopes rounded and descending steeply to margins. 
Sulcus originating on umbo near beak, deepening an- 
teriorly and occupying nearly half valve width. Flanks 
bounding sulcus moderately inflated. Tongue long and 
narrowly rounded. Delthyrium bounded by narrow, 
obliquely elevated deltidial plates. 

Dorsal valve gently convex in lateral profile except 
for the umbonal region, which is more narrowly 
rounded; anterior profile a broad dome, narrowly el- 
evated medially where fold is located. Umbonal and 
median regions swollen. Fold originating on umbo, 
poorly defined for one-third of its distance anteriorly, 
elevated moderately at midvalve and prominent at an- 
terior, where its width is one-third or somewhat less 
of valve width. Fold narrowly rounded. Flanks gently 
swollen and depressed well below fold anterior to mid- 
valve. 

Ventral valve interior having strong and long dental 


M.A.S. 1589A (=USNM 155774); M.A.S. 1589B plates and a median septum that fails to reach the apex. 
(=USNM 155775); M.A.S. 1589C (=USNM 155776). Measurements (in mm).— 
dorsal maximum 
length valve width hinge thickness 
(L) length (W) width (T) L/W ratio T/W ratio 

USNM 203518 

(loc. 506j) 22.0 19.0 26.0 17.5 14.2 0.84 0.55 
USNM 203517a 

(loc. 509z) 16.1 15.5 18.3 11.2 10.1 0.88 0.56 
USNM 203517b 24.1 22.5 30.0 7.9 18.2 0.80 0.61 
USNM 203519 

(loc. 523m) 24.1 20.8 27.8 14.7 16.5 0.85 0.59 
USNM 200906 

(Holotype; loc. 524m) 26.8 23.7 30.1 16.9 19.6 0.89 0.65 
USNM 200907a 

(loc. 524n) 28.2 25e5 36.6 wh 22.0? 0.77 0.60 

38.0 22.8 25.6 0.79 0.68 


USNM 200907b 30.0? 26.0? 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuUTRO 117 


Diagnosis.—Large moderately transverse Elita with 
the fold scarcely visible on the umbo and broad costae. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —506c, j; 509s; 521d, m, q; 523m; 524m, 
n; 529s; USGS (Kirk) 536; 30541. 

Types.—Holotype: USNM 200906. Figured Para- 
type: USNM 200907. 

Discussion.—Compared to specimens of E. incon- 
sueta (Fenton and Fenton, 1924) the New Mexico 
species attains a much larger size, has a narrower and 
less protuberant ventral umbo, and has stronger cos- 
tae. Elita magister resembles E. diversa Stainbrook 
(1948) but it is larger, less transverse, and has a more 
elevated beak than the Independence species. The 
specimen figured by Stainbrook (1948, pl. 2, fig. 21 
[USNM 135279]) is an immature specimen in which 
the costae have not yet formed. 

Elita magister is a rare species in the Sly Gap For- 
mation and most of the specimens come from the San 
Andres Range. 


Family MARTINIIDAE Waagen, 1883 


OIOSIA new genus 
[Greek oios = peculiar] 


Small, unequally biconvex, ventral valve having 
greater depth than dorsal valve; hinge narrower than 
maximum width; interarea narrow, delthyrium wide, 
partially closed by apical plate. Anterior commissure 
uniplicate. Fold and sulcus non-costate; flanks marked 
by low rounded costae, often only faintly visible. Mi- 
crosculpture consisting of mat of minute spines or pa- 
pillae that leave a pitted surface where stripped from 
the external layer. 

Ventral valve interior with small teeth and long, thin 
dental plates. Myophragm in old shells. Dorsal valve 
interior having narrow sockets bounded by thin and 
delicate socket ridges that lack supports. Spire loosely 
coiled, six volutions on a side. 

Type species.—Brachythyris putilla Stainbrook, 
1947, p. 322, pl. 47, figs. 24-28. 

Diagnosis.—Small martiniids having long dental 
plates and matted papillae on exterior. 

Comparison.—The presence of dental plates in the 
ventral valve and absence of strong costae on the fold 
and sulcus effectively separate Stainbrook’s species 
from Brachythyris M’Coy (1844). The nearest de- 
scribed genus is Eobrachythyris Brice (1970) which 
possesses dental plates and an apical plate, both of 
which are present in the Percha genus. Oiosia n. gen. 
differs from Eobrachythyris in its microsculpture, the 
non-costate fold and sulcus, subdued costation, closely 
disposed dental plates and shell form. Species de- 
scribed and mentioned by Brice are strongly costate 


and one of them is rather mucronate and thus entirely 
unlike the Percha genus. This genus is suggestive of 
Eomartiniopsis Sokolskaya (1941), but there are no 
septal plates in Oiosia as required by Sokolskaya’s 
description, and the exterior is papillose, not pitted as 
in the Russian genus. 


Oiosia putilla (Stainbrook) 
Plate 9, figures 10-13; plate 36, figures 56-60; 
plate 37, figures 42-65; plate 42, figures 11-25; 
plate 44, figures 27-35 


Brachythyris putilla Stainbrook, 1947, p. 322, pl. 47, figs. 24-28. 


A very rare species at all Percha exposures but geo- 
graphically widely represented. 

Stratigraphic occurrence.—Percha Formation (Box 
Member). 

Localities. —506, a, b, p, q; 507b, h, 0; 509a, b, e; 
527d, e; 3026a; 3042s. 

Types.—Holotype: M.A.S. 1585A (=USNM 
155797). Unfigured Paratypes: M.A.S. 1585 (=USNM 
155798). Figured Hypotypes, herein: USNM 200975, 
200976, 200977b, e, 201144, 201145a, 204955a, 204956. 


Superfamily CYRTINACEA Fredericks, 1912 
Family CYRTINIDAE Fredericks, 1912 
Genus CYRTINA Davidson, 1858 


Cyrtina inulta Stainbrook 
Plate 37, figures 1-35 
Cyrtina inulta Stainbrook, 1945, p. 59, pl. 6, figs. 26-29; 1948, p. 

778, pl. 2, figs. 17-20. 

About medium-sized, transversely rectangular in 
dorsal view, hemipyramidal in lateral view; hinge width 
less than or equal to maximum width; cardinal extremi- 
ties obtusely rounded to acute. Anterior commissure 
strongly uniplicate, sides varying from moderately 
rounded to nearly straight and oblique to midvalve; 
interarea long, varying from moderately curved and 
apsacline to nearly flat and catacline. Beak sharp, oc- 
casionally twisted. Deltidium narrow and moderately 
convex; foramen apical, often long. Surface costate, 
costae strongly rounded and separated by spaces about 
equal in width to width of costae; about five to seven 
costae on a side. Entire surface covered by fine gran- 
ules or spines that are seldom preserved and usually 
seen only with difficulty. 

Ventral valve in lateral profile forming an angle of 
50° to 60°, the lower side of the triangle gently convex; 
anterior profile moderately domed but with deep me- 
dian depression representing sulcus. Flanks gently 
swollen and steep. Sulcus originating at beak, deep- 
ening and widening anteriorly to occupy one-third to 
one-half shell width. Tongue moderately long and 
sharply rounded. 


BULLETIN 315 


Dorsal valve evenly and gently convex in lateral 
profile but very broadly and gently convex in anterior 
profile; fold originating at beak, widening anteriorly 
and sharply set off from flanks by deep grooves; flanks 
flatly convex. 

Dorsal valve interior with a long and spreading car- 
dinal process. 

Measurements (in mm).— 


dorsal mid- 
length valve hinge width thick- L/W 
(L) length width (W)_ ness _ ratio 


USNM 201068a 


(loc. 506c) DAO LOSS 16.0 16:0) 13820) 0575 
USNM 201068b 

(loc. 506c) 9.1 9.9 14.7 14.2 12.1 0.64 
USNM 201068c 

(loc. 506c) 10.0 8.4 11.1 12.6 SOROS 
USNM 201068d 

(loc. 506c) Test 7.6 10.5 12.4 10.2 0.57 
USNM 201068e 

(loc. 506c) 7e9) 7.9 IEC Grell Op OS63 


Diagnosis. —Small to medium Cyrtina with five to 
seven costae on each side, a narrowly rounded fold 
often with slight median depression posteriorly, and a 
broad, spreading, four-pronged cardinal process. 

Stratigraphic occurrence.—Sly Gap Formation; 
Contadero Formation. 

Localities.—Sly Gap: 506c, 1, j, v; 509g, h, i, j, 1, n, 
5 195 S56 Wo 0S WR DIES GE Ce Is, 115 Oy Gp ir US D2 <, WR 
524m, n, y; 526k, m, n, q; 529s; 3054b. Contadero: 
526g. 

Types.—Figured Hypotypes: USNM 201068a-f. 

Discussion.—Like most species of the genus, this 
One is quite variable but seems, nevertheless, to con- 
form well with Stainbrook’s illustrated specimens. 
Some of the specimens are larger than those recorded 
by Stainbrook. The species is commonest in the Sac- 
ramento Mountains, especially at the Indian Wells 
Canyon locality (506c). 


Cyrtina variabilis new species 
Plate 37, figures 36-41; plate 38, figures 1-12 


Small for the genus, wider than long, subrectangular 
in dorsal view, subpyramidal in lateral view; anterior 
commissure broadly uniplicate; sides nearly straight 
and subparallel, anterior margin broadly rounded; hinge 
about equal to midwidth and with small cardinal points. 
Interarea fairly long, curved and catacline. Surface 
costate, costae and interspaces of about equal width; 
six Or seven costae on a side. Sulcus broad and shal- 
low, occupying 40 percent of the valve width; tongue 
short and rounded. Fold originating at the beak, low 
and broadly rounded. Deltidium fairly wide and with 


the foramen occupying about 60 percent of its length. 
Interior not seen. 

Measurements (in mm).—USNM 201270: length 8.7, 
dorsal valve length 7.2, hinge width 9.0, midwidth 9.7, 
thickness 7.0. 

Diagnosis.—Small Cyrtina with six or seven costae 
on a side, a broad low fold and shallow sulcus, fairly 
wide deltidium with foramen occupying 60 percent of 
its length. 

Stratigraphic occurrence.—Onate Formation. 

Localities.—5220,; 523d, f, i, 0, t; 524a, d; 526c; 528n; 
3023c; 3052a. 

Types.—Holotype USNM 201270; Figured Para- 
types: USNM 201271, 203558. 

Discussion.—We have assigned specimens from 12 
localities to this species. Each specimen has its own 
peculiarities but the variability seems consistent with 
that seen in other species, Cyrtina inulta Stainbrook 
(1947), for example. Specimens from the “‘Paint Mine”’ 
at Nackaye Mountain (5230) are extremely variable, 
no two seeming to be connected except generically, 
but all having a lower, wider dorsal fold and smaller 
size than Stainbrook’s species. 


Order TEREBRATULIDA Waagen, 1883 | 
Suborder TEREBRATULIDINA Waagen, 1883 
Superfamily DIELASMATACEA Schuchert, 1913 | 
Family CRANAENIDAE Cloud, 1942 | 
Subfamily CRANAENINAE Cloud, 1942 
Genus CRANAENA Hall and Clarke, 1893 


Cranaena calvini (Hall and Whitfield) 
Plate 38, figures 44-53 


Cryptonella eudora Hall and Whitfield, 1873, p. 225. 

Cryptonella calvini Hall and Whitfield, 1873, p. 239. 

(2?) Dielasma calvini (Hall and Whitfield) Hall and Clarke, 1893, p. 
296 (nor pl. 80, figs. 20-22). 

Cranaena calvini (Hall and Whitfield) C. L. Fenton, 1919, p. 372; 
Belanski, 1928, p. 17, pl. 3, figs. 1-7; Cloud, 1942, p. 135; Stain- 
brook, 1945, p. 46, pl. 4, fig. 35. 

Cranaenella calvini (Hall and Whitfield) Fenton and Fenton, 1924, 
p. 132, pl. 25, figs. 23-25. 


A few specimens referred to this species were taken 
from the Sly Gap Formation. One of the specimens 
(USNM 200844) exceeds the dimensions of those from 
the Hackberry Group of Iowa but has the same sub- 
rhomboidal form and flattened profile. 

Stratigraphic occurrence.—Sly Gap Formation. 

Localities. —506j; 5090, w, z; 5210, r; 524m. 

Types.—Figured Hypotypes: USNM 200844a, 
200845. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 119 


Cranaena navicella (Hall) 
Plate 38, figure 30 


Terebratula navicella Hall, 1867, p. 391, pl. 60, figs. 38-44. 

Centronella (?) navicella (Hall) Hall and Clarke, 1895, p. 79, figs. 
40-42. 

Centronella navicella (Hall) Fenton, 1919, p. 372. 

Cranaenella navicella (Hall) Fenton and Fenton, 1924, p. 130, pl. 
25, figs. 18-22. 


A single specimen, somewhat larger than normal but 
with subcarinate dorsal valve, is referred to this Hack- 
berry form. The specimen comes from the Rhodes 
Canyon Formation at locality 528). 

Types.—Figured Hypotype: USNM 203564. 


Cranaena species | 
Plate 39, figures 27, 28 


Three specimens of Cranaena were taken from the 
Onate Formation at localities 522f, 529k, and 3023c, 
the largest one from the former locality. The speci- 
mens are of about average size for the genus. The 
largest one is somewhat rhomboidal in outline, with 
the maximum width at midvalve. The anterior margins 
taper to a narrowly rounded extremity. The posterior 
margins form an angle of 90°. The valves are nearly 
equally, but gently, convex. The largest specimen re- 
sembles Cranaena amana Stainbrook and C. calvini 
(Hall and Whitfield) from the Hackberry Formation, 
but it is more tapering anteriorly than either of those 
species. 

Measurements (in mm).—USNM 207694: length 
19.1, midwidth 18.2*, thickness 8.0. 

Types.—Figured Specimen: USNM 207694. 


Cranaena species 2 


A small, probably immature, Cranaena was found 
at Indian Wells Canyon, locality 506c. The specimen 
is somewhat suggestive of C. navicella (Hall, 1867), 
but its obvious immaturity makes a positive specific 
identification impossible. 

Types.—Mentioned Specimen: USNM 207702. 


Family HETERELASMINIDAE Licharew, 1956 
Genus AFILASMA Stehli, 1961 


Afilasma inexpectatum new species 
Plate 38, figures 32-43; plate 44, figures 36—47 


Near average size for a heterelasminid brachiopod 
(nearly 25 mm in length), subequally biconvex, lon- 
gitudinally elliptical in outline; sides broadly rounded; 
anterior margin slightly rounded to truncate. Anterior 
commissure rectimarginate. Lateral commissure near- 
ly straight. Beak erect, short, labiate and with a large 
submesothyridid foramen. Surface marked only by 
concentric lines of growth. 


Ventral valve of about equal depth to that of dorsal 
valve, moderately and evenly convex in lateral profile; 
moderately domed in anterior profile, with short steep 
sides. Umbonal region narrowly swollen, the swelling 
reaching maximum at about midvalve then declining 
anteriorly to form a long gently curved, moderately 
steep anterior slope. 

Dorsal valve gently convex in lateral profile, some- 
what narrowly domed in anterior profile, more so than 
ventral valve. Median region swollen; umbonal region 
gently convex; anterior slope moderately steep. 

Ventral valve interior with widely divergent dental 
plates. Dorsal valve with long diverging inner hinge 
plates reaching nearly one-third dorsal valve length. 

Measurements (in mm).— 


dorsal maxi- 
valve mum __ thick- apical 
length length width ness _ angle 


USNM 200930 


(loc. 509u) 23.4 20.6 16.5 172 58° 
USNM 203530 
(Holotype; loc. 528a) 20.6 18.2 13.9 lo 60° 


Diagnosis.—Elongate, rectimarginate Afilasma with 
the width about two-thirds of the length. 

Stratigraphic occurrence.—Rhodes Canyon For- 
mation; Contadero Formation. 

Localities. —Contadero Formation: 509u; 528a. 
Rhodes Canyon Formation: 523v. 

Types.—Holotype: USNM 203530. Figured Para- 
type: USNM 200930. 

Discussion.—This species differs strongly from Af- 
ilasma beecheri Stehli (1961) from the Chemung at 
Hornellsville, New York. The New Mexico species is 
smaller, has longer inner hinge plates, is more slender, 
and has gently convex lateral margins. Afilasma inex- 
pectatum is a very rare fossil, only three specimens 
having been found. 


Afilasma species 
Plate 38, figure 31 


Two specimens of another species of Afilasma were 
taken from the Thoroughgood Formation, but they are 
too poor for naming. The best specimen indicates a 
species larger than A. inexpectatum, having a length 
of 28 mm and a width of 20 mm. The specimen is more 
expanded anteriorly and less convex than the Rhodes 
Canyon species, but it has the same elongated and 
divergent inner hinge plates in the dorsal valve and 
short divergent dental plates in the ventral valve. 

Locality.—522. 

Types.—Figured Specimen: USNM 200995b. 

Comparison.—This species differs from A. beecheri 
Stehli (1961) in its outline although it is about the same 
size. Because Afilasma sp. tapers posteriorly and ex- 


120 BULLETIN 315 


pands anteriorly, it is quite different from A. inexpec- 
tatum from the Contadero and Rhodes Canyon for- 
mations. 


CLARIFICATION OF TAXONOMIC AND STRATIGRAPHIC 
PROBLEMS RAISED BY STAINBROOK’S 1947 
PAPER ON THE PERCHA BRACHIOPODS 


Recurrent misinterpretations of the age of the Per- 
cha Formation stem from Stainbrook’s assessment of 
a few brachiopod taxa. During this study, we exam- 
ined all specimens upon which Stainbrook based his 
conclusion that the Percha fauna was of Mississippian 
age. We discuss each suspect taxon below. 

Dielasma coreiforme (Stainbrook 1947, p. 318, pl. 47, 
figs. 34-37) is placed in the synonymy of Composita 
bellula Stainbrook (1947) because it has an impunctate 
shell and the form of a young specimen of C. bellula. 
No unequivocal terebratulaceans have yet been found 
in the Box Member of the Percha Formation. 

Locality.—Hillsboro (Stainbrook 1947, p. 318). 

Types.—M.A.S. 1618 (=USNM 135262). 

Leptaena analoga [Stainbrook (not Phillips, 1836), 
1947, p. 304] was collected by Kindle (1909, p. 16) as 
well as by Stainbrook, who said that his specimens 
were demonstrably from the overlying Mississippian 
Lake Valley Formation. Examination of Kindle’s 
specimen of Leptaena (USNM 62004) suggests that it, 
too, came from the Mississippian. The specimen is 
occupied by a crinoid holdfast, unlikely for a Percha 
specimen because Percha echinoderms are extremely 
rare. Collecting by National Museum of Natural His- 
tory parties over some 20 years failed to locate a Lep- 
taena unequivocally in place in Upper Devonian beds. 
However, float specimens from the overlying Missis- 
sippian strata are frequently encountered. 

Buxtonia singularis (Stainbrook 1947, p. 314, pl. 46, 
figs. 1, 2) is of questionable position for two reasons: 
first, its collecting locale is the outcrop of Rhodes Can- 
yon Formation (not Percha) in Rhodes Pass; second, 
it appears to be a Pennsylvanian Linoproductus Chao 
(1927). The flattish dorsal valve, large spine bases on 
the ears, lack of dorsal spines, and an extremely small 
beak that could not have accommodated the large 
curved cardinal process of Buxtonia Thomas (1914), 
all point to Linoproductus. 

Stratigraphic occurrence.—Float from Pennsylva- 
nian strata at locality near entrance to Rhodes Pass. 

Types.—Holotype: USNM 135253. 

Shumardella glabraventra (Stainbrook 1947, p. 317, 


pl. 47, figs. 14-18). Stainbrook described this species 
as coming from the Percha Formation at Hillsboro, 
New Mexico, and from east of Rhodes Pass. The spec- 
imen is not Shumardella Weller (1910), a Mississip- 
pian genus, but belongs to a rare species of Evanes- 
cirostrum Sartenaer (1965) and comes from the Rhodes 
Canyon Formation. 

Types.—Holotype: M.A.S. 1587 (=USNM 135258; 
not figured herein). 

Cyrtospirifer breviposticus (Stainbrook 1947, p. 320, 
pl. 44, figs. 3-6). Stainbrook’s type specimen is said 
to come from the Percha Formation in Rhodes Pass. 
There is no Percha in Rhodes Pass. The specimen 
comes from the Rhodes Canyon Formation and is de- 
scribed herein. 

Brachythyris bisbeensis (Stainbrook 1947, p. 321, pl. 
47, figs. 23, 29, 30) is definitely not a Brachythyris 
M’Coy (1844) because it has dental plates, barely vis- 
ible on the shell surface. The specimen shows no mi- 
cro-ornament and, thus, is difficult to relate to Indo- 
spirifer Grabau (1931), which has never been found in 
the Percha. The specimen may be a small Anthraco- 
spirifer Lane (1963), possibly related to A. rockymon- 
tana (Marcou, 1858) from the Pennsylvanian. 

Locality.—Mount Martin, near Bisbee, Arizona. 

Types.—Holotype: USNM 135256. 

Tylothyris novamexicana (Stainbrook 1947, p. 323, 
pl. 47, figs. 31-33) is another species said to come from 
the Percha Formation in Rhodes Pass. There is no 
Percha Formation in Rhodes Pass; this species, ac- 
tually from the Rhodes Canyon Formation, is de- 
scribed herein. 

Types.—Holotype: M.A.S. 1620 (=USNM 135263). 

Cleiothyridina reticulata (Stainbrook 1947, p. 326, 
pl. 45, figs. 31-35) is another species said to come from 
the non-existent Percha Formation in Rhodes Canyon. 
It is actually from the Rhodes Canyon Formation and 
is made the type species of Crinisarina, described 
herein. 

Types.—Holotype: M.A.S. 1588a (=USNM 135249); 
Paratype: M.A.S. 1588b (=USNM 135249b). 

Rhipidomella manticula [Stainbrook 1947, p. 303, pl. 
45, figs. 3, 5, 6 (not fig. 4 = Thiemella platys n. sp.)]. 
Three specimens from the Rhodes Canyon Formation 
are more properly referred to Aulacella Schuchert and 
Cooper (1931) and are described herein as Aulacella 
manticula (Stainbrook). The fourth specimen, from the 
Box Member of the Percha Formation, is described 
herein as Thiemella platys n. sp. 


DEVONIAN BRACHIOPODS OF NEw MExico: COOPER AND DUTRO 121 


APPENDIX 1 


LocaALity Lists (GEOGRAPHIC) 


Lava Gap: 509z. 


List of Localities in New Mexico en 
Lost Canyon: 528i. 


Agua Chichita Canyon: 529m Marble Canyon: 506i; 509}; 526q, r; 5290, r. 
Alamo Canyon: 509i, y; 521p; 524; 526k, m, n; 528u, w; 529}, n. Mud Springs Mountains: 506d, e, y; 509f; S21v; 522f, g, h, j, k, 1, 
Alamo Peak: 506s; 509g, k, v; 522d, e, i; 524, g, h, i, j, k, 1; 528x; N, O, U, V, W, X, y, Z; 523c, e, i; 524p, r; 5261; 528e. 
3053. Mule Canyon: 509h, ha; 52Ic. 
Arcente Canyon: 509m; 521n; 529w; 3045. Nackaye Mine: 506r, t; 522p, q; 523d, j, k, 0; 524x; 526e, i; 5291. 
Ash Canyon: 523p, q; 524m; 528r. Nunn Ranch: 527k; 3029. 
Bear Canyon: 529t. Percha Creek: 506p; 507g; 3031. 
Bear Mountain: 506q; 507j, k, 1, m, p, r; 509a, b, e; 527a, b, e, |, Pig Canyon: 509w, x; 5211; 526h, 0; 3052a. 
m,n, 0, p; 3042, s. Red House Mountain: 523g, h. 
Caballero Canyon: 5091. Rhodes Canyon: 506w, x; 509p, r, t, u; 52le, f, g, h, i, j; 522a, b, 
Capitol Peak: 52lu, w, x, y, z; 526s; USGS 538, 540. Gym o23 Fim vis 524e> tenviy Zs 205 Xt 27 te ez) 2Oas Callas Ss 
Chise: S29y. 3045; 3054, b, c, f-i, k, q. 
Chloride Flat: 509; 527g. Rincon: 522r, s; 523a. 
Deadman Canyon (Sacramento Mountains): 5210; 524a; 526f, p, v; San Andres Canyon: 523r, s, t, u; 524q; 529v; USGS 536. 
529k; 3013. Santa Rita: 506b; 507a, c, 0, s, z; 527f; 3043; USGS 459. 
Deadman Canyon (San Andres Mountains): 521s; 523y; 524s; 526a; Sawpit Canyon: S07y. 
S528y; 529q, s, u. Sheep Mountain: 506j; 5090, q, s; 521d, k, m, q, r, t; 522; 523n; 
Derry Hills: 506u; 522p, q; 523b, f; 526). 524c, n; 527r, v; 528g, 0, v; 3016; USGS 534. 
Dog Canyon: 524b, d. Sierra Caballo Mountains fault: 522t; 524w. 
Dry Canyon: 524y. Sly Gap: 521d. 
Escondido Canyon: 523w, x, z; 524f. Sulphur Gap: USGS 534. 
Georgetown: 506, a; 507b. Tierra Blanca Creek: 507q; 3029. 
Hembrillo Canyon: 5240, u; 527q; 528b. Trujillo Creek: 3030. 
Hembrillo Gap: USGS 541. Wilson Ranch: 509c; 527d, h; 3026, a; 3037. 
Hermosa: 526b, c, d; 527w, x; 528i, k, n, p; 529p. 
Hillsboro: 507h; 527); 3025a; 3034; USGS 461. List of Localities in lowa 


Indian Wells Canyon: 506c, v; 507; 509n; 529x. APOE ED) 5 +S) 
Johnson Park Canyon: 526z; 528t; 529g, i; 3023, a-d. ISSO STE SHETES SOMES, [oF SPAS Wh i Vis 2S Sia SAE Ta NG 


Kingston: 507d, e, f, i, n, t, u, v, w, x; 509d; 3035; USGS 460: 4804. yD ; 
Lake Valley: 527c, i; USGS 512. Hackberry Formation: 527y; 528d. 


APPENDIX 2 


Loca.ity Lists (STRATIGRAPHIC) 


Onate Formation: 506d, r, t, u; 521z; 522c-j, I-v, z; 523a, b, d-k, Contadero Formation: 506w, x; 509t; 521f, j; 522a; 5231, n, q, r; 
0, s—u, w-z; 524a, b, d, f-l, p-r, x; 526b, c, h-j, 0, p, v, z; 527u, 524c, 0, oa, s, t; 526a, g; 527q, v; 528a, b, g, j, 0, q; 3054k. 
w, x; 528e, i, k, n, s, t, u; 529g, j, k-o, q, r, t, w; 3013; 3023; Rhodes Canyon Formation: 509u; 521g—i, 523p, v; 524e, z; 527z; 
3025a, c; 3053; USGS 540. 528c, j; 3054q. 
Sly Gap Formation: 506c, e, i, j, s, v, y; 507; S09f, g, h, ha, i-s, v— Thoroughgood Formation: 522. 
z; 521c, d, e, k-y; 522b, k, w—y; 523c, m; 524m, n, u, v, w, y; Percha Formation (Box and Ready Pay members): 506, a, b, p, q; 
$26d-f, k—n, q-s, x; 527r, t; 528p, r, v-y; 5291, p, s, u, V, X, y; 507a-z; 509, a-e; 527a—p; 3026, a; 3039; 3030; 3031la, b; 3034; 
3016; 3025; 3045b, c, g; 3054b, c, fH. 3035; 3037; 3042, s; 3043; USGS 459; 460; 461; 508; 509; 510; 512; 
4804. 
APPENDIX 3 
LocaALity DESCRIPTIONS AND FAUNAL LISTS 
362a. Independence Formation 506. Percha Formation (Box Member) 
On Iowa Highway 220, 0.3 mile west of Middle Amana, Iowa. Ravine just north of hairpin turn of road on west edge of George- 
Coeloterorhynchus schucherti (Stainbrook) (not found in New town, SE'4SE'% sec. 1, T. 17 S., R. 12 W., Santa Rita (71%4’) 
Mexico) quadrangle, New Mexico. Collectors: J. Bridge and G. A. Coo- 
Floweria transversalis new species per. 
Nervostrophia extensa new species Acanthatia nupera (Stainbrook) 
Schizophoria amanaensis Stainbrook Athyris transversa (Stainbrook) 
Spinatrypa trulla (Stainbrook) Composita bellula Stainbrook 
362b. Independence Formation Cyrtiorina kindlei (Stainbrook) 
West Amana, Iowa. Collector: M. A. Stainbrook. Cyrtospirifer convexus new species 


Schizophoria amanaensis Stainbrook C. rotundus new species 


PLZ 


$06. (continued) 


Galeatella galeata Muir-Wood and Cooper 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 

L. varispinosus (Stainbrook) 

Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 

R. perchaensis new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 

S. spinigera (Kindle) 

Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 

Thiemella platys new species 
Toryniferella spinosa (Kindle) 


506a. Percha Formation (Box Member) 


1.4 miles east of hairpin bend in west edge of Georgetown, and 
about 4-4 mile south of Georgetown, Silver City (30’) quadrangle, 
New Mexico. Collectors: J. Bridge and G. A. Cooper. 

Acanthatia nupera (Stainbrook) 

Cyrtiorina kindlei (Stainbrook) 

Cyrtospirifer convexus new species 

Galeatella galeata Muir-Wood and Cooper 

Leioproductus varispinosus (Stainbrook) 

Oiosia putilla (Stainbrook) 

Paurorhyncha cooperi Stainbrook 

Planoproductus hillsboroensis (Kindle) 

Porostictia perchaensis (Stainbrook) 

Ptychomalotoechia sobrina (Stainbrook) 

Strophopleura notabilis (Kindle) 

Syringospira prima Kindle 

Toryniferella spinosa (Kindle) 


506b. Percha Formation (Box Member) 


North side of New Mexico Highway 180, about center W12NE%4 
sec. 20, T. 17 S., R. 11 W., 3.25 miles east of Santa Rita, Santa 
Rita (72') quadrangle, New Mexico. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer rotundus new species 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Lingula species | 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria multicostata new species 
P. patens new species 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 


BULLETIN 315 


Retichonetes obscurus new species 

R. perchaensis new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 

S. spinigera (Kindle) 

Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 


506c. Sly Gap Formation (mostly upper) 
Indian Wells Canyon, NW'44SW!14NW' sec. 14, T. 16 S., R. 10 


E., 3 miles due east of School for Blind at north edge of Ala- 
mogordo, Otero County, Sacramento Mountains, New Mexico. 
Acutatheca prolifica new species 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Cranaena species 2 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
D. vulgaris Stainbrook 
Douvillina arcuata (Hall) 
D. minuta new species 
D. parviuscula new species 
D. species | 
Douvillinaria diversa new species 
D. variabilis (Calvin) 
Elita magister new species 
Floweria prava (Hall) 
Gypidula bowsheri new species 
G. mimica new species 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer exfoliata new species 
Nervostrophia extensa new species 
N. geniculata new species 
N. multinervosa Stainbrook 
N. cf. N. multinervosa Stainbrook 
N. solida new species 
Petrocrania specifically unidentified 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Pseudodouvillina euglyphea Stainbrook 
Schizophoria amanaensis Stainbrook 
S. iowensis (Hall) 
Skenidium independense Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Sulcatostrophia calvini (Miller) 
Thomasaria warreni new species 


506d. Onate Formation 


Sec. 24, T. 13 S., R. 5 W., Mud Springs Mountains, west of Truth- 
or-Consequences, Sierra County, New Mexico. Collector: Carl 
C. Branson (=loc. 522 

Desquamatia costata new species 

Hypothyridina species 

Morphorhynchus varicostatum new species 

Schizophoria lata Stainbrook 

Warrenella floweri new species 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 123 


506e. Sly Gap Formation 
(same as locs. 506d and S06y). 
Costatrypa varicostata (Stainbrook) 
Thomasaria demissa new species 
506i. Sly Gap Formation 
South side of Marble Canyon, one mile east of mouth, NW'%4 sec. 
26, T. 16 S., R. 10 E., east of Alamogordo, New Mexico. Col- 
lectors: C. C. Branson, E. L. Yochelson, and G. A. Cooper. 
Acutatheca prolifica new species 
Ambocoelia species | 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Douvillina parviuscula new species 
Floweria prava (Hall) 
Gypidula bowsheri new species 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Petrocrania specifically unidentified 
Pseudodouvillina euglyphea Stainbrook 
Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Thomasaria warreni new species 
506j. Sly Gap Formation 
Type section of Sly Gap Formation, south end of Sheep Mountain 
on north side of Sly Gap, sec. 25, T. 11 S., R. 5 E., San Andres 
Mountains, Capitol Peak (15’) quadrangle, New Mexico. Collec- 
tors: C. C. Branson, E. L. Yochelson, and G. A. Cooper (=loc. 
§21d). 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Cranaena calvini (Hall and Whitfield) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
Douvillinaria diversa new species 
Elita magister new species 
Floweria prava (Hall) 
Gypidula bowsheri new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer anomala new species 
Iowatrypa rara new species 
Nervostrophia mucrospina new species 
Philhedra species 
Pseudoatrypa devoniana (Webster) 
Rhipidomella species | 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Tenticospirifer cf. T. cyrtinaformis (Hall and Whitfield) 
Thomasaria demissa new species 
T. warreni new species 
Tylothyris sulcocostata new species 
506p. Percha Formation (Box Member) 
Middle Fork of Percha Creek, NE'4 sec. 13, T. 16 S., R. 9 W., 
one mile west of Kingston, Hillsboro (15') quadrangle, New Mex- 
ico. 
Acanthatia nupera (Stainbrook) 


Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus plicatus (Kindle) 
L. pauperculus new species 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
S. spinigera (Kindle) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
506q. Percha Formation (Box Member) 
Bear Mountain, 6—7 miles northwest of Silver City, New Mexico 
(=loc. 507m). 
Acanthatia nupera (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
C. rotundus new species 
Galeatella galeata Muir-Wood and Cooper 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Thiemella platys new species 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
506r. Onate Formation 
4 miles N. 73° E. of Derry, 0.7 mile S. 10° E. of Nackaye Mine, 
SEM4SW!I4NE% sec. 28, T. 17 S., R. 4 W., Sierra County, New 
Mexico (=loc. 522q). 
506s. Sly Gap Formation 
East side of Alamo Peak, NY2NE!'4SE'% sec. 25, T. 16 S., R. 10 
E., east of Alamogordo, Alamogordo (15’) quadrangle, New Mex- 
ico (=loc. 509k). 
506t. Onate Formation 
Nackaye Mine, 4 miles east of Derry, Sierra County, New Mexico 
(=loc. 523d [see for list]). 
506u. Onate Formation 
Derry Hills, 2 mile east of Derry, Sierra County, New Mexico 
(=loc. 523f). 


BULLETIN 315 


506v. Sly Gap Formation 
Second canyon south of Indian Wells Canyon, SE% sec. 14, T. 
16S.,R. 10 E., 4 miles east of the railroad station in Alamogordo, 
New Mexico (=loc. 507). 
Acutatheca prolifica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia solida new species 
Pseudoatrypa devoniana (Webster) 
Pseudodouwvillina euglyphea Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Sulcatostrophia calvini (Miller) 
Thomasaria warreni new species 
506w. Contadero Formation 
One-quarter mile north of Rhodes Canyon, SW'% sec. 3, T. 13 
S., R. 4 E., Socorro County, Black Top Mountain (15’) quadran- 
gle, New Mexico (=loc. 526g [see for list]). 
506x. Contadero Formation 
(same as loc. 526g). 
506y. Sly Gap Formation 
SW'ANEMSE sec. 24, T. 13 S., R. 5 W., %2 mile N. 45° E. of 
Mud Mountain (el. 5749’), Mud Springs Mountains, 3.6 miles N. 
45° W. of Truth-or-Consequences, (formerly Hot Springs), Cu- 
chillo (742') quadrangle, New Mexico. Collectors: C. C. Branson, 
A. L. Bowsher, J. T. Dutro, and G. A. Cooper. 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
Costatrypa varicostata (Stainbrook) 
Thomasaria demissa new species 
507. (=loc. 506v). 
507a. Percha Formation (lower Box Member) 
WI4NE sec. 20, T. 17 S., R. 11 W., 3.2 miles each of the junc- 
tion of New Mexico Hwy. 180 (90) and road to Georgetown, '4— 
14 mile north of New Mexico Rt. 180, Santa Rita (7!2') quadran- 
gle. New Mexico. Collectors: A. L. Bowsher, G. A. Cooper, and 
J. T. Dutro (=locs. 506b, 5070). 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia spinigera (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
507b. Percha Formation (Box Member) 
SE corner, SE% sec. 1, T. 17 S., R. 12 W., at Georgetown, Santa 
Rita (72') quadrangle, New Mexico. Collectors: J. T. Dutro and 
G. A. Cooper (=loc. 506a). 
Athyris transversa (Stainbrook) 


Cyrtiorina kindlei (Stainbrook) 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Petasmaria patens new species 
Porostictia perchaensis (Stainbrook) 
Schuchertella percha (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
507c. Percha Formation (lower Box Member) 
East of bridge on New Mexico highway 180 (90), 3.8 miles west 
of its junction with New Mexico highway 61, 4 miles east of Santa 
Rita, New Mexico. Collectors: J. T. Dutro and G. A. Cooper. 
Composita bellula Stainbrook 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Kindle) 
Lingula species 2 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Sentosia spinigera (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
507d. Percha Formation (Box Member) 
Middle Percha Creek, NE% sec. 13, T. 16 S., R. 9 W., one mile 
west of Kingston, Hillsboro (15’) quadrangle, New Mexico. 
Athyris transversa (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 


$07e. Percha Formation (Box Member) 


Five miles by road (New Mexico 180) southwest of Kingston, 
New Mexico. 

Athyris transversa (Stainbrook) 

Cyrtiorina kindlei (Stainbrook) 

Schizophoria australis Kindle 

Syringospira prima Kindle 


507f. Percha Formation (big nodule zone at top of Ready Pay 
Member) 


On New Mexico highway 180, 2 miles east of Kingston, NW 
corner NE sec. 16, T. 16S., R. 8 W., Hillsboro (15’) quadrangle, 
New Mexico. 

Composita bellula Stainbrook 

Cyrtiorina kindlei (Stainbrook) 

Leioproductus varispinosus (Strainbrook) 

Paurorhyncha cooperi Stainbrook 

Ptychomalotoechia sobrina (Stainbrook) 


507g. Percha Formation (Box Member) 


Canyon of North Percha Creek, SW corner SE4SW% sec. 24, 
T. 16 S., R. 9 W., Hillsboro (15’) quadrangle, New Mexico. 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus plicatus (Kindle) 


DEVONIAN BRACHIOPODS OF NEW MEXxICo: COOPER AND DuTRO 1 


507g. (continued) 
Paurorhyncha cooperi Stainbrook 
Planoproductus hillsboroensis (Kindle) 
Schizophoria australis Kindle 
507h. Percha Formation (Box Member) 
Type section of the Box Member, SW!4SE'%4 sec. 14, T. 16 S., 
R. 7 W., 2 mile south of the east end of the Box Canyon, 24 
miles S. 75° E. of the center of Hillsboro, Hillsboro (15’) quad- 
rangle, New Mexico. Collectors: R. E. Grant, G. A. Cooper, and 
J. T. Dutro. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer? animasensis (Girty) 
C. convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus coloradensis (Kindle) 
Leioproductus defessa Stainbrook 
L. pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
S. spinigera (Kindle) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Thiemella platys new species 
Toryniferella spinosa (Kindle) 
507i. Percha Formation (top of Ready Pay Member or base of Box 
Member, nodules with Syringospira) 
SE cor. NE4SW'4SW'4 sec. 18, T. 17 S., R. 14 W., Silver City 
(7¥2') quadrangle, New Mexico. Collectors: G. A. Cooper and J. 
T. Dutro. 
Syringospira prima Kindle 
507j. Percha Formation (nodular bed at top of Ready Pay or base 
of Box Members) 
West side of Bear Mountain, SW'4SE'4SW' sec. 11, T. 17 S., 
R. 15 W., Silver City (7!2') quadrangle, New Mexico. (=loc. 507p) 
Syringospira prima Kindle 
507k. Percha Formation (Box Member) 
Same as locality 507). 
Athyris transversa (Stainbrook) 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Strophopleura notabilis (Kindle) 
Toryniferella spinosa (Kindle) 
5071. Percha Formation (Box Member) (lower hard bed to top in 
measured section) 
Bear Mountain, SW'44SE'44NE'4SE sec. 13, T. 17S., R. 15 W., 
Silver City (712') quadrangle, New Mexico. Collectors: J. T. Du- 
tro and G. A. Cooper. 


i) 
tn 


Athyris transversa (Stainbrook) 
Leioproductus plicatus (Kindle) 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
507m. Percha Formation (Box Member) 
SE'% sec. 13, T. 17 S., R. 15 W., Bear Mountain, Silver City 
(7¥2') quadrangle, New Mexico. Collectors: J. T. Dutro and G. 
A. Cooper. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Cyrtospirifer convexus new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes perchaensis new species 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
507n. Percha Formation (Box Member) 
Middle Percha Creek, one mile west of Kingston, Hillsboro (15’) 
quadrangle, New Mexico (=loc. 507d). 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Hispidaria spinosa new species 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Trifidorostellum obscurum new species 
5070. Percha Formation (Box Member) 
Same as localities 506b and 507a. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Leioproductus plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria compacta new species 
P. patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 
R. perchaensis new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
S. spinigera (Kindle) 


126 BULLETIN 315 


5070. (continued) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
507p. Percha Formation (nodular bed, top of Ready Pay or base 
of Box Members) 
Bear Mountain, SE% sec. 13, T. 17 S., R. 15 W., Silver City 
(7¥4') quadrangle, New Mexico. Collectors: J. T. Dutro and G. 
A. Cooper. 
Leioproductus varispinosus (Stainbrook) 
Syringospira prima Kindle 
507q. Percha Formation (Box Member) 
Upper Tierra Blanca Creek, Hillsboro (15’) quadrangle, New 
Mexico (=loc. 3039). Collector: A. L. Bowsher. 
Cyrtiorina kindlei (Stainbrook) 
Laminatia laminata (Kindle) 
Leioproductus plicatus (Kindle) 
Petasmaria patens new species 
Porostictia perchaensis (Stainbrook) 
507r. Percha Formation (Box Member) 
SE cor., NE4SW'% sec. 18, T. 17 S., R. 14 W., Silver City 
(7¥4') quadrangle, east of Bear Mountain, New Mexico. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Petasmaria patens new species 
507s. (=loc. 507a) (see for list). 
507t. Percha Formation (Box Member) 
1.5 miles west of Kingston at mine, Hillsboro (15’) quadrangle, 
New Mexico. Collector: F. V. Stevenson. 
Athyris transversa (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Ptychomalotoechia sobrina (Stainbrook) 
507u. Percha Formation (Box Member) 
Two miles west of Kingston, New Mexico. Collector: A. L. 
Bowsher. 
Hispidaria spinosa new species 
Ptychomalotoechia sobrina (Stainbrook) 
Toryniferella spinosa (Kindle) 
507v. Percha Formation (Box Member) 
At Ranger Station, north end of trail on North Percha Creek, SE 
corner, SW'%4 sec. 24, T. 15 S., R. 8 W., Hillsboro (15’) quadran- 
gle, New Mexico. 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Schuchertella percha (Stainbrook) 
507w. Percha Formation (Box Member) 
Four miles up North Percha Creek from branch of West and South 
Percha Creeks, Sierra County, New Mexico. Collector: A. L. 
Bowsher. 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
507x. Percha Formation (Box Member—lower) 
Fork of Percha Creek, center, north boundary of sec. 16, T. 16 
S., R. 8 W., on New Mexico highway 180, 2 miles east of Kings- 
ton, Hillsboro (15') quadrangle, New Mexico. Collector: F. V. 
Stevenson. 
Athyris transversa (Stainbrook) 


Cyrtiorina kindlei (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Syringospira prima Kindle 
507y. Percha Formation (Box Member) 
Sawpit Canyon near Mud Spring, center, NY2SW'4 sec. 5, T. 16 
S., R. 8 W., Hillsboro (15’) quadrangle, New Mexico. Collector: 
A. L. Bowsher. 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
507z. Percha Formation (Box Member) 
Road cut, 4.6 miles east of Santa Rita on New Mexico Hwy. 180, 
New Mexico. Collectors: J. T. Dutro and G. A. Cooper. 
Composita bellula Stainbrook 
Laminatia laminata (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
509. Percha Formation (Box Member) 
1.5 miles north of US highway 260 to Cliff in Chloride Flat, Silver 
City (72') quadrangle, New Mexico. Collector: F. V. Stevenson. 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Petasmaria patens new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
509a. Percha Formation (Box Member) 
Bear Mountain Road, south side of Bear Mountain, 6 miles north- 
west of Silver City, Silver City (712’), quadrangle, New Mexico. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Petasmaria patens new species 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes perchaensis new species 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
509b. Percha Formation (Box Member) 
South side of Bear Mountain, along Bear Mountain road, 6.5 miles 
northwest of Silver City, Silver City (7!2') quadrangle, New Mex- 
ico. 
Acanthatia nupera (Stainbrook) 
Ambocoelia capillata new species 
Athyris transversa (Stainbrook) 
Camarotoechia? species 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 127 


509b. (continued) 


Leioproductus varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 

S. spinigera (Kindle) 

Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 

Toryniferella spinosa (Kindle) 


509c. Percha Formation (Box Member) 


0.4 mile SSE of Wilson Ranch, SE4NE“4SW'4 sec. 10, T. 18 S, 
R. 7 W., Hillsboro (15’) quadrangle, New Mexico. Collectors: F. 
V. Stevenson and A. L. Bowsher. 

Athyris transversa (Stainbrook) 

Composita bellula Stainbrook 

Cyrtiorina kindlei (Stainbrook) 

Galeatella galeata Muir-Wood and Cooper 

Leioproductus pauperculus new species 

Petrocrania ourayensis (Kindle) 

Porostictia perchaensis (Stainbrook) 

Ptychomalotoechia sobrina (Stainbrook) 

Schizophoria australis Kindle 


Costatrypa extensa new species 

C. varicostata (Stainbrook) 

Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia mucrospina new species 
Spinatrypa trulla decorticata new subspecies 
Schizophoria lata Stainbrook 

Tylothyris sulcocostata new species 


509g. Sly Gap Formation 


Alamo Peak, east slope, 5 miles S. 60° E. of Alamogordo, 0.5 
mile west of Holmes Mine, center, E¥2NW!4NE'4SE sec. 25, 
T. 16 S., R. 10 E., Otero County, Lincoln Forest Service Map, 
New Mexico. Collector: A. L. Bowsher. 

Acutatheca prolifica new species 

Coeloterorhynchus magnificum new species 

Costatrypa varicostata (Stainbrook) 

Cyrtina inulta Stainbrook 

Douvillina parviuscula new species 

Hyborhynchella bransoni Cooper 

Hypothyridina emmonsi rotunda new subspecies 

Indospirifer exfoliata new species 

Nervostrophia multinervosa Stainbrook 

N. solida new species 

Pseudoatrypa devoniana (Webster) 

Pseudodouvillina euglyphea Stainbrook 

Schizophoria iowensis (Hall) 

Spinatrypa compacta new species 

Strophonelloides parvulus new species 

Thomasaria warreni new species 


Toryniferella spinosa (Kindle) 509h. Sly Gap Formation 
509d. Percha Formation (lower Box Member or uppermost Ready Hypothyridina beds to base of sill, 0.63 mile N. 75° E of the mouth 
Pay Member—nodular zone) of Mule Canyon, Sacramento Mountains, NE4NW14 sec. 9, T. 


On New Mexico highway 180 (90), 2 miles each of Kingston, 
Hillsboro (15') quadrangle, New Mexico (=locs. 507f, 507x). 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Paurorhyncha cooperi Stainbrook 


509e. Percha Formation (Box Member) 


Bear Mountain road, west side of Bear Mountain, SE14NE' sec. 
10, T. 17S., R. 15 W., Silver City (712) quadrangle, New Mexico. 

Athyris transversa (Stainbrook) 

Cyrtospirifer convexus new species 

Laminatia laminata (Kindle) 

Leioproductus varispinosus (Stainbrook) 

Oiosia putilla (Stainbrook) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

P. multicostata new species 

Petrocrania ourayensis (Kindle) 

Ptychomalotoechia sobrina (Stainbrook) 

Retichonetes perchaensis new species 

Schuchertella resupinata new species 

Sentosia praecedens (Stainbrook) 

Strophopleura notabilis (Kindle) 

Syringospira prima Kindle 

Toryniferella spinosa (Kindle) 
509f. Sly Gap Formation (beds with Thomasaria = Bed 1) 
One half mile northeast of the highest peak, Mud Springs Moun- 
tains, 3.7 miles northwest of Truth-or-Consequences, New Mex- 
ico (=locs. S06y; 3025). 

Acutatheca prolifica new species 

Athabaschia? sp. 1 

Calvinaria bransoni Stainbrook 

Cariniferella alleni new species 

Coeloterorhynchus magnificum new species 


17 S., R. 10 E., Alamogordo (15') quadrangle, New Mexico. Col- 
lectors: J. T. Dutro and G. A. Cooper. 
Acutatheca prolifica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Douvillina parviuscula new species 
Douvillinaria diversa new species 
D. variabilis (Calvin) 
Floweria magnicicatrix new species 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia geniculata new species 
N. mucrospina new species 
N. solida new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 


509ha. Sly Gap Formation 


Just below the black shale, locality same as 509h. 
Coeloterorhynchus magnificum new species 
Douvillina rectangulata new species 


5091. Sly Gap Formation 


2’ miles up Alamo Canyon from mouth, SE'4NE'%4 sec. 2, and 
SW'4NW'4 sec. 1, T. 17 S., R. 10 E., Sacramento Mountains, 
Alamogordo (15’) quadrangle, New Mexico. 

Acutatheca prolifica new species 

Cariniferella alleni new species 

Coeloterorhynchus magnificum new species 

Costatrypa varicostata (Stainbrook) 

Cyrtina inulta Stainbrook 


BULLETIN 315 


509i. (continued) 509n. Sly Gap Formation 
Douvillinaria variabilis (Calvin) NW/4NE'4 sec. 22, T. 16 S., R. 10 E., 3 miles east of Alamo- 
Floweria prava (Hall) gordo, Alamogordo (15’) quadrangle, New Mexico. Collector: C. 


Hypothyridina emmonsi rotunda new subspecies C. Branson. 
Indospirifer exfoliata new species 
Nervostrophia geniculata new species 
N. mucrospina new species 
N. solida new species 
Pseudoatrypa devoniana (Webster) 
Pseudodouvillina euglyphea Stainbrook 
Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Thomasaria warreni new species 
Tylothyris sulcocostata new species 
509j. Sly Gap Formation 
Marble Canyon, east-southeast of Alamogordo, Alamogordo (15’) 
quadrangle, New Mexico. Collector: F. V. Stevenson (=loc. 506i). 
Acutatheca prolifica new species 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Douvillina parviuscula new species 
D. species | 
Gypidula bowsheri new species 
Hyborhynchella bransoni Cooper 
Nervostrophia solida new species 
Productella rugatula Stainbrook 


Acutatheca prolifica new species 

Aulacella elliptica new species 

Calvinaria bransoni Stainbrook 

Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 

Devonoproductus intermedius new species 
Douvillina parviuscula new species 

D. rectangulata new species 

D. species | 

Floweria prava (Hall) 

Gypidula bowsheri new species 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia mucrospina new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Pseudodouvillina euglyphea Stainbrook 
Schizophoria amanaensis Stainbrook 

S. iowensis (Hall) 

Spinatrypa trulla decorticata new subspecies 
Strophodonta umbonata new species 
Strophonelloides dorsoconvexus new species 
S. parvulus new species 

Thomasaria demissa new species 

T. warreni new species 


Pseudodouvillina euglyphea Stainbrook 5090. Sly Gap Formation 

Spinatrypa trulla decorticata new subspecies Sheep Mountain, north side of Sly Canyon, sec. 25, T. 11 S., R. 
Strophodonta umbonata new species 5 E., San Andres Mountains, Capitol Peak (15’) quadrangle, New 
Strophonelloides parvulus new species Mexico. Collector: F. V. Stevenson. 


509k. Sly Gap Formation 
East side of Alamo Peak, NY2NE'4SE% sec. 25, T. 16 S., R. 10 
E., Alamogordo (15’) quadrangle, New Mexico. Collectors: A. 
L. Bowsher and G. A. Cooper. 
Acutatheca prolifica new species 
Douvillinaria diversa new species 
Indospirifer exfoliata new species 


Ambocoelia species 2 

Coeloterorhynchus magnificum new species 
Cranaena calvini (Hall and Whitfield) 
Cyrtina inulta Stainbrook 

Strophonelloides dorsoconvexus new species 
Thomasaria demissa new species 

T. warreni new species 


509p. Sly Gap Formation (beds b-g) 
One quarter mile north of Rhodes Canyon, SW'%4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico 
(=loc. 3054). 


Nervostrophia solida new species 
Pseudodouvillina euglyphea Stainbrook 
Thomasaria warreni new species 


5091. Sly Gap Formation 
North side of Caballero Canyon, a branch of Alamo Canyon, 
southeast of Alamogordo, Sacramento Mountains, Alamogordo 
(15') quadrangle, New Mexico. Collector: F. V. Stevenson. 
Acutatheca prolifica new species 
Cyrtina inulta Stainbrook 
Nervostrophia solida new species 
Productella rugatula Stainbrook 
Strophonelloides parvulus new species 
Thomasaria warreni new species 
509m. Sly Gap Formation (9 feet, 3 inches below the 4-foot black 
shale) 
Arcente Canyon, NW'4 sec. 20, T. 16 S., R. 11 E., Lincoln Forest 
Service Map, about 7 miles east of Alamogordo, New Mexico. 
Collectors: A. L. Bowsher and G. A. Cooper. 


Aulacella elliptica new species 

Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Costatrypa extensa new species 

C. varicostata (Stainbrook) 

Cyrtina inulta Stainbrook 

Nervostrophia mucrospina new species 
Schizophoria amanaensis Stainbrook 

S. species 2 

Spinatrypa trulla decorticata new subspecies 
Strophodonta umbonata new species 
Strophonelloides dorsoconvexus new species 
Thomasaria demissa new species 

Tylothyris sulcocostata new species 


Acutatheca prolifica new species 509q. Sly Gap Formation (lower half) 

Sec. 25, T. 11 S., R. 5 E., Sheep Mountain, north side of Sly 
Canyon, San Andres Mountains, Capitol Peak (15') quadrangle, 
New Mexico (=loc. 3016). Collectors: W. T. Allen and A. L. 
Pseudodouvillina euglyphea Stainbrook Bowsher. 

Sulcatostrophia calvini (Miller) Aulacella elliptica new species 

Calvinaria bransoni Stainbrook 


Coeloterorhynchus magnificum new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer exfoliata new species 


Thomasaria warreni new species 


DEVONIAN BRACHIOPODS OF NEw MExiIco: COOPER AND DUTRO 129 


509q. (continued) 


Coeloterorhynchus magnificum new species 
Devonoproductus intermedius new species 
Gypidula bowsheri new species 
Nervostrophia geniculata new species 
Spinatrypa trulla decorticata new subspecies 
Tenticospirifer cf. T. cyrtinaformis (Hall and Whitfield) 
Thomasaria demissa new species 
T. warreni new species 
509r._ Sly Gap Formation 
Same as locality 509p. Collectors: W. T. Allen and A. L. Bow- 
sher. 
Aulacella elliptica new species 
Cariniferella alleni new species 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
Eostrophalosia inexpectata new species 
Iowatrypa rara new species 
Nervostrophia mucrospina new species 
N. solida new species 
Pseudoatrypa devoniana (Webster) 
Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides dorsoconvexus new species 
S. parvulus new species 
Thomasaria demissa new species 
T. warreni new species 
Tylothyris sulcocostata new species 


509s. Sly Gap Formation 


South slope of Sheep Mountain, opposite windmill in Sly Canyon, 
San Andres Mountains, Capitol Peak (15’) quadrangle, New Mex- 
ico. Collectors: C. C. Branson, E. L. Yochelson, and G. A. Coo- 
per. 

Aulacella elliptica new species 

Calvinaria bransoni Stainbrook 

Coeloterorhynchus magnificum new species 

Cyrtina inulta Stainbrook 

Cyrtospirifer whitneyi (Hall) 

Devonoproductus intermedius new species 

Douvillina parviuscula new species 

D. species 1 

Elita magister new species 

Floweria prava (Hall) 

Gypidula bowsheri new species 

Hypothyridina emmonsi rotunda new subspecies 

Indospirifer anomala new species 

Nervostrophia mucrospina new species 

N. solida new species 

Productella rugatula Stainbrook 

Pseudoatrypa devoniana (Webster) 

Rhipidomella species 1 

Schizophoria iowensis (Hall) 

Spinatrypa trulla decorticata new subspecies 

Strophodonta umbonata new species 

Strophonelloides parvulus new species 

Thomasaria warreni new species 

Tylothyris sulcocostata new species 


509t. Contadero Formation (zones k—-m) 


One quarter mile north of Rhodes Canyon, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15') quadrangle, New Mexico. 
Collectors: A. L. Bowsher and G. A. Cooper (=loc. 3054k). 

Ambocoelia species 2 

Crinisarina reticulata (Stainbrook) 

Cupularostrum? stevensoni new species 


Cyrtospirifer pyramidalis new species 

C. whitneyi (Hall) 

Douvillina contaderoensis new species 
Productella species 1 

Ripidiorhynchus newmexicoense new species 
Spinatrypa obsolescens new species 
Tylothyris species | 


509u. Rhodes Canyon Formation (beds o-r) 


Same locality as 509t. 
Afilasma inexpectatum new species 
Aulacella elliptica new species 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Cyrtospirifer pyramidalis new species 
Productella species | 
P. species 2 
Ripidiorhynchus newmexicoense new species 
Tylothyris novamexicana Stainbrook 


509v. Sly Gap Formation 


Upper 10 feet, Alamo Peak, sec. 25, T. 16 S., R. 10 E. Lincoln 
Forest Service Map, east of Alamogordo, New Mexico. 
Acutatheca prolifica new species 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
Costatrypa varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Devonoproductus vulgaris Stainbrook 
Floweria prava (Hall) 
Gypidula bowsheri new species 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 


509w. Sly Gap Formation (0-10 feet below the 4-foot black shale) 


W'2SW'4 sec. 7, T. 16 S., R. 11 E., first canyon north of Pig 
Canyon, 6 miles east-northeast of Alamogordo, Sacramento 
Mountains, Lincoln Forest Service Map, New Mexico. 

Acutatheca prolifica new species 

Coeloterorhynchus magnificum new species 

Cranaena calvini (Hall and Whitfield) 

Devonoproductus vulgaris Stainbrook 

Hyborhynchella bransoni Cooper 

Hypothyridina emmonsi rotunda new subspecies 

Indospirifer exfoliata new species 

Pseudodouvillina euglyphea Stainbrook 

Spinatrypa trulla decorticata new subspecies 

Sulcatostrophia calvini (Miller) 

Thomasaria warreni new species 


509x. Sly Gap Formation (above the 4-foot black shale) 
Same as locality 509w. 


Aulacella elliptica new species 

Cyrtina inulta Stainbrook 

Douvillina parviuscula new species 

D. species 1 

Floweria prava (Hall) 

Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 

Tylothyris sulcocostata new species 


S09y. Sly Gap Formation (above 3-foot bed of black shale) 


Alamo Canyon, 2.5 miles up from mouth, Sacramento Mountains, 
Alamogordo (15’) quadrangle, New Mexico. 

Aulacella elliptica new species 

Cyrtina inulta Stainbrook 

Douvillina parviuscula new species 

Pseudoatrypa devoniana (Webster) 


130 BULLETIN 315 


509y. (continued) 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Sulcatostrophia calvini (Miller) 
509z. Sly Gap Formation 
Collection made by students of Dr. C. L. Baker in Lava Gap, 
Sheep Mountain, Capitol Peak (15’) quadrangle, New Mexico. 
Calvinaria bransoni Stainbrook 
Costatrypa varicostata (Stainbrook) 
Cranaena calvini (Hall and Whitfield) 
Cyrtospirifer whitneyi (Hall) 
Devonoproductus intermedius new species 
Floweria prava (Hall) 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer anomala new species 
Nervostrophia mucrospina new species 
Pseudoatrypa devoniana (Webster) 
Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Tylothyris sulcocostata new species 
521c. Sly Gap Formation (below the sill) 
SWI4SEY“4NW'4 sec. 9, T. 17 S., R. 10 E., 0.75 mile N. 60° E. 
of the mouth of Mule Canyon, Sacramento Mountains, Alamo- 
gordo (15') quadrangle, New Mexico. Collectors: J. T. Dutro and 
G. A. Cooper. 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Douvillinaria diversa new species 
Floweria prava (Hall) 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia solida new species 
Petrocrania specifically unidentified 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Sulcatostrophia calvini (Miller) 
Thomasaria warreni new species 
521d. Sly Gap Formation 
Type section Sly Gap Formation, sec. 25, T. 11 S., R. 5 E., San 
Andres Mountains, Capitol Peak (15') quadrangle, New Mexico. 
Collectors: F. V. Stevenson and A. L. Bowsher. 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Cyrtospirifer cf. C. thalattodoxa Crickmay 
C. whitneyi (Hall) 
C. species 3 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
D. species | 
Elita magister new species 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer anomala new species 
I. exfoliata new species 


$21f. Contadero Formation (Bed F, near top) 


521g. Rhodes Canyon Formation (Zone E) 


521h. Rhodes Canyon Formation (Zone D) 


lowatrypa rara new species 

Nervostrophia geniculata new species 

N. mucrospina new species 

N. multinervosa Stainbrook 

N. solida new species 

Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria amanaensis Stainbrook 

S. iowensis (Hall) 

Spinatrypa trulla decorticata new subspecies 
Strophodonta umbonata new species 
Strophonelloides dorsoconvexus new species 
S. parvulus new species 

Sulcatostrophia calvini (Miller) 
Tenticospirifer cf. T. cyrtinaformis (Hall and Whitfield) 
Thomasaria warreni new species 

Tylothyris sulcocostata new species 


52le. Sly Gap Formation 


One quarter mile north of Rhodes Canyon, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: F. V. Stevenson. 

Aulacella elliptica new species 

Calvinaria bransoni Stainbrook 

Cariniferella alleni new species 

C. species 

Coeloterorhynchus magnificum new species 

Costatrypa extensa new species 

C. varicostata (Stainbrook) 

Cyrtina inulta Stainbrook 

Devonoproductus intermedius new species 

Douvillina parviuscula new species 

D. rectangulata new species 

Douvillinaria variabilis (Calvin) 

Floweria prava (Hall) 

Gypidula bowsheri new species 

Hypothyridina emmonsi rotunda new subspecies 

Nervostrophia mucrospina new species 

N. solida new species } 

Productella rugatula Stainbrook 

Pseudoatrypa devoniana (Webster) 

Schizophoria amanaensis Stainbrook 

Spinatrypa trulla decorticata new subspecies 

Strophodonta umbonata new species 

Strophonelloides parvulus new species 

Thomasaria demissa new species 

T. warreni new species 

Tylothyris sulcocostata new species 


Rhodes Canyon, San Andres Mountains, Black Top Mountain 
(15') quadrangle, New Mexico. Collector: F. V. Stevenson. 
Crinisarina reticulata (Stainbrook) 
Cyrtospirifer pyramidalis new species 
Schizophoria species indeterminate 
Tylothyris species | 


Same as locality 52le. Collector: F. V. Stevenson. 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Cyrtospirifer whitneyi (Hall) 

Tylothyris novamexicana Stainbrook 


Same as locality 52le. Collector: F. V. Stevenson. 
Crinisarina reticulata (Stainbrook) 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DuTRO 131 


521h. (continued) 


Cyrtospirifer pyramidalis new species 

C. cf. C. thalattodoxa Crickmay 

Productella species | 

521i. Rhodes Canyon Formation (Zone C) 

Same as locality 52le. Collector: F. V. Stevenson. 
Crinisarina reticulata (Stainbrook) 

521j. Contadero Formation (Zone G) 

Same as locality 52le. Collector: F. V. Stevenson. 
Ambocoelia species 2 
Cupularostrum? stevensoni new species 
Douvillina contaderoensis new species 
Ripidiorhynchus newmexicoense new species 
Spinatrypa obsolescens new species 
Tylothyris species | 

521k. Sly Gap Formation (lower) 

Sheep Mountain, NW'4SW'4 sec. 25, T. 11 S., R. 5 E., Capitol 

Peak (15') quadrangle, New Mexico. Collectors: J. T. Dutro and 

G. A. Cooper (=loc. 521m). 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Gypidula subcarinata new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer aff. I. orestes (Hall and Whitfield) 
Towatrypa rara new species 
Nervostrophia mucrospina new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Thomasaria demissa new species 
T. warreni new species 
Tylothyris sulcocostata new species 


$211. Sly Gap Formation 


Pig Canyon anticline, 2 miles N. 7° E. of Ortega Peak, 4.5 miles 
N. 80° E. of Alamogordo, SW!'4NE!4SW'4 sec. 7, T. 16 S., R. 
11 E., Lincoln Forest Service Map, New Mexico. Collectors: W. 
T. Allen and A. L. Bowsher. 

Acutatheca prolifica new species 

Coeloterorhynchus magnificum new species 

Costatrypa varicostata (Stainbrook) 

Cyrtina inulta Stainbrook 

Indospirifer exfoliata new species 

Nervostrophia solida new species 

Pseudodouvillina euglyphea Stainbrook 

Spinatrypa trulla decorticata new subspecies 

Sulcatostrophia calvini (Miller) 

Thomasaria warreni new species 
521m. Sly Gap Formation (lower 20 feet) 
Center NW!4SW'4 sec. 25, T. 11 S., R. 5 E., Sheep Mountain, 
San Andres Mountains, Capitol Peak (15’) quadrangle, New Mex- 
ico. Collectors: J. T. Dutro and G. A. Cooper (=loc. 521k). 

Aulacella elliptica new species 

Calvinaria bransoni Stainbrook 

Coeloterorhynchus magnificum new species 

Costatrypa extensa new species 

C. varicostata (Stainbrook) 

Devonoproductus intermedius new species 


Douvillina parviuscula new species 
D. sp. 1 
Douvillinaria diversa new species 
D. variabilis (Calvin) 
Elita magister new species 
Floweria prava (Hall) 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia mucrospina new species 
Petrocrania specifically unidentified 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 
Tylothyris sulcocostata new species 
521n. Sly Gap Formation 
Arcente Canyon, 7 miles south of High Rolls, SE44NW'4NE!4 
sec. 20, T. 16 S., R. 11 E., Lincoln Forest Service Map, Sacra- 
mento Mountains, New Mexico. Collectors: A. L. Bowsher and 
W. T. Allen. 
Douvillina minuta new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer exfoliata new species 
Nervostrophia solida new species 
Thomasaria warreni new species 
52lo. Sly Gap Formation 
Deadman Canyon, 3 miles southwest of Alamogordo, Sacramento 
Mountains, Alamogordo (15’) quadrangle, New Mexico (=loc. 
526f). 
Acutatheca prolifica new species 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cranaena calvini (Hall and Whitfield) 
Cyrtina inulta Stainbrook 
Nervostrophia solida new species 
Pseudodouvillina euglyphea Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 
521p. Sly Gap Formation 
Alamo Canyon, Sacramento Mountains, Alamogordo (15’) quad- 
rangle, New Mexico. Collector: F. V. Stevenson (=loc. 526n). 
Acutatheca prolifica new species 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Devonoproductus intermedius new species 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia geniculata new species 
N. extensa new species 
Productella rugatula Stainbrook 
521q. Sly Gap Formation (upper 25 feet) 
Center NW!4SW'4 sec. 25, T. 11 S., R. 5 E., Sheep Mountain. 
San Andres Mountains, Capitol Peak (15’) quadrangle, New Mex- 
ico. Collectors: J. T. Dutro and G. A. Cooper. 
Aulacella elliptica new species 
Costatrypa varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
Elita magister new species 
Indospirifer anomala new species 
lowatrypa rara new species 
Nervostrophia geniculata new species 
N. mucrospina new species 
Petrocrania specifically unidentified 


132 BULLETIN 315 


521q. (continued) 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 
Thomasaria warreni new species 
52Ir. Sly Gap Formation 
Sheep Mountain, San Andres Range, sec. 25, T. 11 S., R. 


SES 
Capitol Peak (15’) quadrangle, New Mexico. Collectors: G. A. 


Cooper and J. T. Dutro (=loc. 521d). 
Acutatheca prolifica new species 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
Cranaena calvini (Hall and Whitfield) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer exfoliata new species 
Productella rugatula Stainbrook 
Pseudodouvillina euglyphea Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Thomasaria warreni new species 


521s. Sly Gap Formation (lower) 


Center NW'4SE% sec. 12, T. 17 S., R. 3 E., Deadman Canyon, 
San Andres Range, Kaylor Mountain (15’) quadrangle, New Mex- 


ico. Collectors: G. A. Cooper and J. T. Dutro. 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Cariniferella iowensis Stainbrook 
Douvillinaria diversa new species 
Nervostrophia mucrospina new species 
N. solida new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 


§21t. Sly Gap Formation (bed just above Onate Formation) 


Sheep Mountain, NW'4SW'4 sec. 25, T. 11 S., R. 5 E., Capitol 
Peak Quadrangle, New Mexico. Collectors: G. A. Cooper and J. 


T. Dutro. 
Aulacella elliptica new species 
Costatyrpa varicostata (Stainbrook) 
Cyrtina inulata Stainbrook 
Douvillinaria diversa new species 
Nervostrophia mucrospina new species 
Pseudoatrypa devoniana (Webster) 
Spinatrypa trulla decorticata new subspecies 
Strophodonta umbonata new species 


52lu. Sly Gap Formation (top 23 feet) 


Capitol Peak, San Andres Mountains, Capitol Peak (15') quad- 
rangle, New Mexico. Collectors: A. L. Bowsher and J. T. Dutro. 


Cariniferella alleni new species 

Costatrypa extensa new species 

C. varicostata (Stainbrook) 

Cyrtospirifer whitneyi (Hall) 

Devonoproductus intermedius new species 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer aff. I. orestes (Hall and Whitfield) 
Nervostrophia mucrospina new species 

N. solida new species 

Pseudoatrypa devoniana (Webster) 
Strophonelloides parvulus new species 


52lv. Sly Gap Formation (Bed K—Thomasaria Bed) 


Mud Springs Mountains, NE44SE™% sec. 24, T. 13 S., R. 5 W., 
west of Truth-or-Consequences, Cuchillo (7/2) quadrangle, New 


Mexico. Collectors: G. A. Cooper and J. T. Dutro. 


Calvinaria bransoni Stainbrook 
Costatyrpa varicostata (Stainbrook) 
Douvillinaria diversa new species 
Nervostrophia solida new species 
Productella rugulata Stainbrook 
Thomasaria demissa new species 


521w. Sly Gap Formation (Number 2 of A. L. Bowsher) 
Capitol Peak, San Andres Moutnains, Capitol Peak (15) quad- 
rangle, New Mexico. Collectors: A. L. Bowsher and J. T. Dutro. 


Costatrypa extensa new species 

C. varicostata (Stainbrook) 
Devonoproductus intermedius new species 
Nervostrophia geniculata new species 

N. mucrospina new species 

Productella rugulata Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 


521x. Sly Gap Formation (Number 3 of A. L. Bowsher) 
Same as locality S2lu. 


Costatrypa varicostata (Stainbrook) 
Nervostrophia solida new species 


52ly. Sly Gap Formation (Number 4 of Bowsher) 
Same as locality 521lu. 

522. Thoroughgood Formation (basal bed) 
Sheep Mountain, SE'44NE% sec. 26, T. 11 S., R. 5 E., Capitol 
Peak (15') quadrangle, New Mexico. Collectors: J. T. Dutro, G. 
A. Cooper and F. V. Stevenson. 


Afilasma species 

Ambocoelia species 3 
Athabaschia? species 2 
Camarotoechia latisinuata new species 
Colophragma species 2 

Crinisarina reticulata (Stainbrook) 
Cyrtospirifer arenarius new species 
C. species | 

C. species 2 

Evanescirostrum species | 
Productella species 3 

Tylothyris compacta new species 


$22a. Contadero Formation (Zone 4) 


Amphitheater, north of Rhodes Canyon, center N¥2SW!4NE'%4 
sec. 9, T. 13 S., R. 4 E., San Andres Mountains, Black Top 
Mountain (15’) quadrangle, New Mexico. Collector: F. V. Ste- 
venson. 

Douvillina contaderoensis new species 

Ripidiorhynchus newmexicoense new species 

Spinatrypa compacta new species 

S. obsolescens new species 

Tylothyris compacta new species 


522b. Sly Gap Formation 


Same as locality 522a. Collector: F. V. Stevenson. 
Thomasaria demissa new species 
T. warreni new species 


§22c. Onate Formation (Bed F) 


Rhodes Canyon, San Andres Mountains, New Mexico. Collector: 
A. L. Bowsher 
Schizophoria lata Stainbrook 


§22d. Onate Formation (Zone I) 


East side of Alamo Peak, % mile west of Holmes Mine, center 
EYNW'4NEMSE' sec. 25, T. 16 S., R. 10 E., Lincoln Forest 
Service Map, east of Alamogordo, New Mexico. Collector: A. L. 
Bowsher. 

Nervostrophia plana new species 

Platyterorhynchus obscurum new species 

Striatochonetes nanus new species 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 1 


522e. Onate Formation (Collection C) 
East side of Alamo Peak (same as locs. 522d, 3053). 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
522f. Onate Formation (Sphaerospongia Bed = Bed E of Cooper 
and Dutro) 
NE'M4SE'% sec. 24, T. 13 S., R. 5 W., Mud Springs Mountains, 
Cuchillo (7/2') quadrangle, New Mexico. Collector: C. C. Bran- 
son. 
Atryparia rubra new species 
Cranaena species | 
Desquamatia costata new species 
Leiorhynchus sphaericum new species 
Morphorhynchus varicostatum new species 
Mucrospirifer parvus new species 
Schizophoria lata Stainbrook 
522g. Same as locality 522f. 
522h. Onate Formation (Bed F) 
Same as locality 506y. Collectors: G. A. Cooper and J. T. Dutro. 
Nervostrophia plana new species 
Schizophoria lata Stainbrook 
5221. Onate Formation (Beds B and C) 
Same as locality 522d. Collector: A. L. Bowsher. 
Echinocoelia pretiosa new species 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
522j. Onate Formation (40-50 feet above the base) 
Same as locality 506y. Collector: A. L. Bowsher. 
Dichacaenia umbonata new species 
Petrocrania specifically unidentified 
Platyterorhynchus obscurum new species 
Rhyssochonetes johnsoni new species 
Schizophoria lata Stainbrook 
522k. Sly Gap Formation (43-83 feet above base) 
Same as locality 506y. Collectors: W. T. Allen, A. L. Bowsher 
and G. A. Cooper. 
Coeloterorhynchus species 
Costatrypa varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Nervostrophia solida new species 
Productella rugatula Stainbrook 
Thomasaria demissa new species 
5221. Onate Formation (basal 19 feet of section) 
Same as locality 506y. Collectors: A. L. Bowsher, W. T. Allen 
and G. A. Cooper. 
Atryparia rubra new species 
Desquamatia costata new species 
Leiorhynchus sphaericum new species 
Morphorhynchus varicostatum new species 
(For conodonts see USGS 9312-SD) 
522m. Onate Formation 
Rhodes Pass, SW'4 sec. 3, T. 13 S., R. 4 E., San Andres Moun- 
tains, Capitol Peak (15’) quadrangle, New Mexico. 
Ambocoelia species 4 
Cyrtina variabilis new species 
Leiorhynchus sphaericum new species 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
522n. Onate Formation (Bed B) 
Same as locality 506y. 
Echinocoelia pretiosa new species 
Emanuella species 


1S) 
nr 


Hypothyridina species 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
$220. Onate Formation 
Same as locality 506y. Collector: F. V. Stevenson. 
Cyrtina variabilis new species 
Desquamatia costata new species 
522q. Onate Formation (Ambocoelia beds) 
Base of section, north end of Derry Hills, northeast of Derry, 
Garfield (712') quadrangle, New Mexico. 
Ambocoelia species 4 
522s. Sly Gap Formation? 
Base of blocky sandstone at top of section; sec. 1, T. 18 S., R. 3 
W., northwest of Rincon, Rincon (7!2') quadrangle, New Mexico. 
Collectors: G. A. Cooper and J. T. Dutro. 
Cyrtospirifer species 5 
Eleutherokomma species 
522t. Onate Formation 
NESE, sec. 22, T. 17 S., R. 4 W., Caballo Mountains, Upham 
(15') quadrangle, New Mexico. Collector: F. V. Stevenson. 
Atryparia rubra new species 
522u. Onate Formation 
Same as locality 506y. 
522v. Same as locality 5221. 
5$22w. Same as locality 52lv. 
522x. Sly Gap Formation (Bed J) 
Same as locality 506y. 
Costatrypa varicostata (Stainbrook) 
Tylothyris sulcocostata new species 
§22y. Sly Gap Formation (Bed I) 
Same as locality S06y. 
Cyrtina inulta Stainbrook 
522z. Onate Formation 
Same as locality S06y. 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
523. Onate Formation 
Same as locality S06y. Collector: F. V. Stevenson. 
Strophodonta species | 
Warrenella floweri new species 
523a. Onate Formation 
Sec. 1, T. 18 S., R. 3 W., northwest of Rincon, New Mexico. 
Atryparia rubra new species 
Hadrorhynchia alta new species 
Nervostrophia plana new species 
Schizophoria lata Stainbrook 
Strophodonta species 1 
Warrenella magna new species 
523b. Onate Formation (beds with Warrenella and Hadrorhyn- 
chia) 
North end of Derry Hills, northeast of Derry, Garfield (72') quad- 
rangle, New Mexico (=loc. 523f). Locality now under roadbed of 
Interstate Highway 25. 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Echinocoelia pretiosa new species 
Hadrorhynchia alta new species 
H. mclareni new species 
Nervostrophia plana new species 
Striatochonetes nanus new species 
Warrenella magna new species 
523c. Sly Gap Formation (Bed G) 
Cobbly zone, same as locality 506y. 
Calvinaria bransoni Stainbrook 


134 BULLETIN 315 


§23c. (continued) Dichacaenia umbonata new species 
Nervostrophia solida new species Hadrorhynchia alta new species 
Thomasaria warreni new species Morphorhynchus varicostatum new species 
§23d. Onate Formation Mucrospirifer parvus new species 
Nackaye Mine, center SW'4 sec. 15 and NW'4 sec. 22, T. 17S., Nervostrophia plana new species 
R. 4 W., Caballos Mountains, Upham (15’) quadrangle, New Schizophoria lata Stainbrook 
Mexico. Collectors: G. A. Cooper and J. T. Dutro. Warrenella floweri new species 
Cupularostrum? species §23j. Onate Formation 
Cyrtina variabilis new species Old mine shaft, NW'4NW'4 sec. 22, T. 17 S., R. 4 W., Caballo 
Desquamatia costata new species Mountains, Upham (15’) quadrangle, New Mexico (=loc. 523d). 
Hadrorhynchia alta new species Desquamatia costata new species 
H. mclareni new species Dichacaenia perplexa new species 
Hypothyridina species Warrenella magna new species 
Leiorhynchus sphaericum new species 523k. Onate Formation (lower Warrenella) 
Nervostrophia plana new species SWI4NE sec. 22, T. 17 S., R. 4 W., southeast of Nackaye 
Petrocrania species | Mines, Caballo Mountains, Upham (15’) quadrangle, New Mex- 
Schizophoria lata Stainbrook ico. 
Warrenella magna new species Desquamatia costata new species 
§23e. Onate Formation (lower brown beds = c-d) Hadrorhynchia mclareni new species 
East of measured section at locality 506y. Schizophoria lata Stainbrook 
Ambocoelia species 4 Warrenella magna new species 
Desquamatia costata new species W. floweri new species 
Dichacaenia perplexa new species $231. Contadero Formation 
Hadrorhynchia alta new species Rhodes Pass, same as localities 506w and 526g. 
Leiorhynchus sphaericum new species 523m. Sly Gap Formation (Bed J of Cooper, field designation) 
Morphorhynchus varicostatum new species Rhodes Pass, same as locality 509p. 
Mucrospirifer parvus new species Costatrypa varicostata (Stainbrook) 
Nervostrophia plana new species Elita magister new species 
Schizophoria lata Stainbrook Pseudoatrypa devoniana (Webster) | 
Strophodonta species | Schizophoria amanaensis Stainbrook 
Warrenella floweri new species Spinatrypa trulla decorticata new subspecies | 
523f. Onate Formation Strophonelloides parvulus new species 
Derry Hills, center, east boundary, SW'%4 sec. 29, T. 17 S., R. 4 Tylothyris sulcocostata new species 
W., just northwest of Derry warm spring, Caballo Mountains, 523n. Contadero Formation (topmost) : 
Garfield (7/2') quadrangle, New Mexico. Collectors: J. T. Dutro Sheep Mountain, SE44NE1 sec. 26, T. 11S., R. 5 E., Capitol 
and G. A. Cooper (= loc. 523b). [Now under Interstate Highway Peak (15') quadrangle, New Mexico. Collectors: G. A. Cooper 
25.] and J. T. Dutro. 


Crinisarina reticulata (Stainbrook) \ 

Cyrtospirifer whitneyi (Hall) 

Douvillina contaderoensis new species 

Floweria aff. F. chemungensis (Conrad) 

Productella? species 2 

Schizophoria species indeterminate 

Spinatrypa compacta new species 

Tylothyris species 1 

§230. Onate Formation (upper Warrenella bed) 

Paint Mine, NW!4NE'4 sec. 27, T. 17 S., R. 4 W., one mile 
southeast of Nackaye Mine, Caballo Mountains, Upham (15’) 
quadrangle, New Mexico. Collectors: G. A. Cooper and J. T. 
Dutro. 

Atryparia rubra new species 

Cyrtina variabilis new species 


Cyrtina variabilis new species 
Desquamatia costata new species 
Echinocoelia pretiosa new species 
Hadrorhynchia alta new species 
H. mclareni new species 
Schizophoria lata Stainbrook 
Striatochonetes nanus new species 
S. subcarinatus new species 
Warrenella magna new species 


523g. Onate Formation (Bed B) 
Red House Mountain, NW'4NW'4 sec. 14, T. 18 S., R. 3 W., 
north-northeast of Hatch, Rincon (72’) quadrangle, New Mexico. 
Desquamatia costata new species 
Warrenella magna new species 


523h. Onate Formation (Bed B) Desquamatia costata new species 
Red House Mountain, SW'44SW!4SW'"4 sec. 11, T. 18 S., R. 3 Dichacaenia perplexa new species 
W., just east of barite mine, Caballo Mountains, Upham (15’) Warrenella magna new species 
quadrangle, New Mexico. W. floweri new species 
Hadrorhynchia alta new species 5§230a. (loose at loc. 5230.) 
H. mclareni new species Aulacella elliptica new species 
Schizophoria lata Stainbrook 523p. Rhodes Canyon Formation 
Warrenella magna new species Ash Canyon, center NE!4 sec. 20, T. 14 S., R. 4 E., San Andres 
523i. Onate Formation (beds C—D) Mountains, Black Top Mountain (15’) quadrangle, New Mexico. 
Sphaerospongia beds, same as locality 522f. Cupularostrum? stevensoni new species 
Cyrtina variabilis new species Cyrtospirifer pyramidalis new species 


Desquamatia costata new species Tylothyris species | 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 13 


523q. Contadero Formation 
Same as locality 523p. 
Colophragma ellipticum new species 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Cyrtospirifer whitneyi (Hall) 
Tylothyris novamexicana Stainbrook 
Uchtospirifer? rarus new species 
523r. Contadero Formation 
San Andres Canyon, San Andres Mountains, Bear Peak (15') 
quadrangle, New Mexico. Collector: A. L. Bowsher. 
Douvillina species 2 
523s. Onate Formation 
Type section of the formation, north side San Andres Canyon, 
sec. 18, T. 18 S., R. 4 E., Bear Peak (15’) quadrangle, New Mex- 
ico. Near old lead mine. 
Dichacaenia umbonata new species 
Echinocoelia pretiosa new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
Striatochonetes nanus new species 
Warrenella magna new species 
523t. Onate Formation (bed with Warrenella) 
Same as locality 523s. 
Cyrtina variabilis new species 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Schizophoria lata Stainbrook 
Strophodonta species | 
Warrenella magna new species 
523u. Onate Formation 
San Andres Canyon, San Andres Range, Bear Peak (15’) quad- 
rangle, New Mexico. Collector: F. V. Stevenson. 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Mucrospirifer parvus new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
““Schuchertella’’ species 
523v. Rhodes Canyon Formation 
Same as locality 506w. Collector: A. L. Bowsher. 
Afilasma inexpectatum new species 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Tylothyris novamexicana Stainbrook 
523w. Onate Formation (Bed B) 
SEM4SE'% sec. 26, T. 18 S., R. 10 E., 0.25 mile southeast of 
Escondido Canyon, Sacramento Mountains, Escondido Canyon 
(15’) quadrangle, New Mexico. Collectors: G. A. Cooper and J. 
T. Dutro. 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
523x. Onate Formation (Bed C) 
Same as locality 523w. Collectors: G. A. Cooper and J. T. Dutro. 
Desquamatia costata new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
Striatochonetes nanus new species 
523y. Onate Formation (‘‘Leiorhynchus’’ Bed) 
NW1'4SW'4 sec. 12, T. 17 S., R. 3 E., Deadman Canyon, San 


in 


Andres Mountains, Kaylor (15’) quadrangle, New Mexico. Col- 
lectors: G. A. Cooper and J. T. Dutro. 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
523z. Onate Formation (Bed E) 
Same as locality 523w. 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
524. Onate Formation 
SEM4NE% sec. 2, T. 17 S., R. 10 E., Alamo Canyon, Alamogordo 
(15’) quadrangle, New Mexico. 
Platyterorhynchus obscurum new species 
524a. Onate Formation (6 to 8 feet below the top) 
Deadman Canyon, Sacramento Mountains, Alamogordo (15’) 
quadrangle, New Mexico. 
Cyrtina variabilis new species 
Mucrospirifer parvus new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
524b. Onate Formation 
Dog Canyon, Sacramento Mountains, Alamogordo (15’) quadran- 
gle, New Mexico. 
Desquamatia costata new species 
Platyterorhynchus obscurum new species 
524c. Contadero Formation 
Center NW'4SW"4 sec. 25, T. 11 S., R. 5 E., Sheep Mountain, 
San Andres Mountains, Capitol Peak (15’) quadrangle, New Mex- 
ico. 
Crinisarina reticulata (Stainbrook) 
Schizophoria species indeterminate 
524d. Onate Formation 
First small canyon tributary of Dog Canyon, Sacramento Moun- 
tains, Alamogordo (15’) quadrangle, New Mexico. Collector: F. 
V. Stevenson. 
Cyrtina variabilis new species 
Desquamatia costata new species 
524e. Rhodes Canyon Formation 
Same as localities 506w, 3054q. 
Crinisarina reticulata (Stainbrook) 
Cyrtospirifer pyramidalis new species 
Productella species | 
524f. Onate Formation (25 feet above base) 
South wall of Escondido Canyon, center sec. 26, T. 18 S., R. 10 
E., Escondido Canyon (15’) quadrangle, New Mexico. 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Schizophoria lata new species 
524g. Onate Formation 
Alamo Peak, sec. 25, T. 16 S., R. 10 E., east of Alamogordo, 
Lincoln Forest Service Map, New Mexico. 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
524h. Onate Formation (11.8 to 16.6 feet) 
Same as locality 524g. 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 


524i. Onate Formation (Bed C of Cooper) 
Same as locality 524g. 
Ambocoelia species 4 
Atryparia rubra new species 
Echinocoelia pretiosa new species 
Emanuella species 
Nervostrophia plana new species 
524}. Onate Formation (Bed F of Cooper) 
Same as locality 524g. 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
Striatochonetes nanus new species 
524k. Onate Formation (Bed K of Cooper) 
Same as locality 524g. 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
5241. Onate Formation (Beds G-K of Cooper) 
Same as locality 524g. 
Desquamatia costata new species 
Mucrospirifer parvus new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
524m. Sly Gap Formation 
Center NE% sec. 20, T. 14 S., R. 4 E., Ash Canyon, San Andres 
Mountains, Black Top Mountain (15') quadrangle, New Mexico. 
Aulacella elliptica new species 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Cranaena calvini (Hall and Whitfield) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Douvillina minuta new species 
D. parviuscula new species 
Douvillinaria diversa new species 
Elita magister new species 
Floweria prava (Hall) 
Hyborhynchella bransoni Cooper 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer anomala new species 
Nervostrophia geniculata new species 
N. solida new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Schizophoria iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Strophodonta umbonata new species 
Strophonelloides dorsoconvexus new species 
Thomasaria warreni new species 
Tylothyris sulcocostata new species 
524n. Sly Gap Formation 
Same as locality 521t. Collectors: G. A. Cooper and J. T. Dutro. 
Costatrypa varicostata (Stainbrook) 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Douvillina parviuscula new species 
Elita magister new species 
Floweria prava (Hall) 
Gypidula bowsheri new species 
Indospirifer exfoliata new species 


36 BULLETIN 315 


Nervostrophia geniculata new species 
N. mucrospina new species 
Pseudoatrypa devoniana (Webster) 
Schizophoria amanaensis Stainbrook 
S. iowensis (Hall) 
Spinatrypa trulla decorticata new subspecies 
Strophonelloides parvulus new species 
Thomasaria warreni new species 
5240. Contadero Formation (lower 5 feet) 
Center SW'%4 sec. 2, T. 16 S., R. 4 E., Hembrillo Canyon, San 
Andres Mountains, Kaylor Mountain (15’) quadrangle, New Mex- 
ico. Collectors: G. A. Cooper and J. T. Dutro. 
Ambocoelia species 2 
Aulacella elliptica new species 
Barroisella species 
Colophragma ellipticum new species 
C. species | 
Crinisarina reticulata (Stainbrook) 
Evanescirostrum glabraventrum (Stainbrook) 
Orbiculoidea species | 
5240a. Contadero Formation (middle) 
Same as locality 5240. 
Tylothyris compacta new species 
524p. Onate Formation (upper 2 feet of Bed D) 
Same as locality 506y. Collectors: G. A. Cooper and J. T. Dutro. 
Ambocoelia species 4 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
524q. Onate Formation (Bed A of Cooper) 
San Andres Canyon (=loc. 523s). 
Ambocoelia species 4 
Emanuella species 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
Schizophoria lata Stainbrook 
524r. Onate Formation (basal foot of Bed D) 
Same as locality S06y. 
Hypothyridina species 
Rhyssochonetes johnsoni new species 
524s. Contadero Formation (Bed K) 
Center NW'4SE sec. 12, T. 17 S., R. 3 E., Deadman Canyon, 
San Andres Mountains, Kaylor Mountain (15’) quadrangle, New 
Mexico. Collectors: J. T. Dutro and G. A. Cooper. 
Douvillina contaderoensis new species 
D. species 2 
Schizophoria amanaensis Stainbrook 
Stainbrookia rotunda new species 
Strophodonta umbonata new species 
524t. Contadero Formation (Bed K) 
Same as locality 506w. Collectors: A. L. Bowsher and W. T. 
Allen. 
Douvillina contaderoensis new species 
Ripidiorhynchus newmexicoense new species 
Spinatrypa obsolescens new species 
Tylothyris species 1 
524u. Sly Gap Formation (Beds I-N) 
Same as locality 5240. Collectors: G. A. Cooper and J. T. Dutro. 
Athabaschia? species | 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 
Douvillinaria diversa new species 
Pseudoatrypa devoniana (Webster) 
Spinatrypa trulla decorticata new subspecies 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuUTRO 137 


524u. (continued) 
Strophonelloides parvulus new species 
(Float from Contadero Formation: Cyrtospirifer pyramidalis new 
species) 
524w. Sly Gap Formation 
Sierra Caballo Mountains, north of saddle at fault, New Mexico. 
Collector: F. V. Stevenson. 
Calvinaria bransoni Stainbrook 
Costatrypa varicostata (Stainbrook) 
Hypothyridina emmonsi rotunda new subspecies 
Indospirifer anomala new species 
Productella rugatula Stainbrook 
Pseudoatrypa devoniana (Webster) 
Tylothyris sulcocostata new species 
524x. Onate Formation (Sly Gap, mixed) 
NE'M4SE% sec. 22, T. 17 S., R. 4 W., Upham (15’) quadrangle, 
New Mexico. Collector: F. V. Stevenson. 
Ambocoelia species 4 
Aulacella elliptica new species 
Emanuella species 
Leiorhynchus sphaericum new species 
Schizophoria lata Stainbrook 
Warrenella floweri new species 
524y. Sly Gap Formation 
Dry Canyon, off Gordon Canyon, SE4SW"% sec. 2, T. 17 S., R. 
10E., 0.5 mile south of junction with Caballero Canyon, Alamo 
Canyon, Sacramento Mountains, Alamogordo (15’) quadrangle, 
New Mexico. Collector: F. V. Stevenson. 
Cariniferella alleni new species 
Cyrtina inulta Stainbrook 
524z. Rhodes Canyon Formation (Bed P) 
Same as locality S06w. 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Cyrtospirifer pyramidalis new species 
Tylothyris novamexicana Stainbrook 
526a. Contadero Formation 
Sec. 8, T. 17 S., R. 4 E., Deadman Canyon, San Andres Moun- 
tains, Kaylor Mountain (15') quadrangle, New Mexico. Collector: 
F. V. Stevenson. 
Aulacella elliptica new species 
Crinisarina reticulata (Stainbrook) 
Cupularostrum? stevensoni new species 
Douvillina contaderoensis new species 
D. species 2 
Schizophoria amanaensis Stainbrook 
Stainbrookia rotunda new species 
526b. Onate Formation 
NYNW'4 sec. 19, T. 13 S., R. 8 W., 2 miles N. 60° E. of Her- 
mosa, Sugar Loaf Peak (15’) quadrangle, New Mexico. Collec- 
tors: G. A. Cooper and J. T. Dutro. 
Ambocoelia species 4 
Devonochonetes? species 
Emanuella species 
Hypothyridina species 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
§26c. Onate Formation (Bed D) 
Same as locality 526b. Collectors: G. A. Cooper and J. T. Dutro. 
Cyrtina variabilis new species 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Hypothyridina species 


Nervostrophia plana new species 
Schizophoria lata Stainbrook 
526d. Sly Gap Formation (Bed E-7 feet below top) 
Same as locality 526b. Collectors: G. A. Cooper and J. T. Dutro. 
Costatrypa varicostata (Stainbrook) 
Douvillinaria diversa new species 
Nervostrophia geniculata new species 
N. mucrospina new species 
Thomasaria demissa new species 
526e. Sly Gap Formation 
0.7 mile S. 10° E. of Nackaye Mines, 4 miles N. 73° E. of Derry, 
SE'4SW'4 sec. 22, T. 17 S., R. 4 W., Upham (15’) quadrangle, 
New Mexico. 
Aulacella elliptica new species 
Calvinaria bransoni Stainbrook 
Costatrypa varicostata (Stainbrook) 
Strophodonta species 2 
Tylothyris sulcocostata new species 
526f. Sly Gap Formation 
Deadman Canyon, 3 miles southwest of Alamogordo, Sacramento 
Mountains, Alamogordo (15’) quadrangle, New Mexico (=loc. 
$210). 
Douvillina parviuscula new species 
Gypidula bowsheri new species 
Pseudodouvillina euglyphea Stainbrook 
Spinatrypa trulla decorticata new subspecies 
526g. Contadero Formation 
Rhodes Canyon, same as locality 506w. 
Ambocoelia species 2 
Cupularostrum? stevensoni new species 
Cyrtina inulta Stainbrook 
Cyrtospirifer whitneyi (Hall) 
Devonoproductus intermedius new species 
Douvillina contaderoensis new species 
D. species 2 
Ripidiorhynchus newmexicoense new species 
Schizophoria species indeterminate 
Spinatrypa obsolescens new species 
Tylothyris species 1 
526h. Onate Formation (middle) 
W'2SW'4 sec. 7, T. 16 S., R. 11 E., first canyon north of Pig 
Canyon, Sacramento Mountains, 6 miles east of Alamogordo, 
Alamogordo (15') quadrangle, New Mexico. 
Desquamatia costata new species 
Mucrospirifer parvus new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
Trigonoglossa? species 
526i. Onate Formation 
Nackaye Mines, same as locality 523d. 
Warrenella magna new species 
526j. Onate Formation 
Derry Hills, same as locality 523f. 
Schizophoria lata Stainbrook 
526k. Sly Gap Formation 
SEY4NE% sec. 2, and SWIANW%4% sec. 1, T. 17 S., R. 10 E., 
Alamo Canyon, Alamogordo (15’) quadrangle, New Mexico. 
Acutatheca prolifica new species 
Ambocoelia species | 
Aulacelia elliptica new species 
Calvinaria bransoni Stainbrook 
Cariniferella alleni new species 


138 BULLETIN 315 


526k. (continued) 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Devonoproductus intermedius new species 
Hypothyridina emmonsi rotunda new subspecies 
Nervostrophia solida new species 
Pseudoatrypa devoniana (Webster) 
Pseudodouvillina euglyphea Stainbrook 
Schizophoria amanaensis Stainbrook 
Spinatrypa trulla decorticata new subspecies 
Sulcatostrophia calvini (Miller) 
Thomasaria warreni new species 
5261. Sly Gap Formation 
Same as locality 506y. 
Coeloterorhynchus magnificum new species 
526m. Sly Gap Formation 
First Canyon south of Alamo Canyon, Alamogordo (15') quadran- 
gle. Collector: F. V. Stevenson. 
Cyrtina inulta Stainbrook 
Douvillina parviuscula new species 
Douvillinaria variabilis (Calvin) 
Thomasaria warreni new species 
526n. Sly Gap Formation 
Alamo Canyon, Sacramento Mountains, Alamogordo (15') quad- 
rangle, New Mexico. Collector: F. V. Stevenson (=loc. 521p). 
Cyrtina inulta new species 
Devonoproductus intermedius new species 
Productella rugulata Stainbrook 
5260. Onate Formation 
Pig Canyon, same as locality 5211. 
Platyterorhynchus obscurum new species 
526p. Onate Formation 
SE corner NE4SW'% sec. 3, T. 17 S., R. 10 E., Deadman Can- 
yon, Sacramento Mountains, Alamogordo (15') quadrangle, New 
Mexico. Collector: A. L. Bowsher (=loc. 3013). 
Platyterorhynchus obscurum new species 
§26q. Sly Gap Formation 
North branch of Marble Canyon, Sacramento Mountains, east of 
Alamogordo, Alamogordo (15') quadrangle, New Mexico. Col- 
lector: F. V. Stevenson. 
Cyrtina inulta Stainbrook 
Indospirifer exfoliata new species 
526r. Sly Gap Formation 
First unnamed canyon north of Marble Canyon, Sacramento 
Mountains, east of Alamogordo, Alamogordo (15’) quadrangle, 
New Mexico. 
Aulacella elliptica new species 
Nervostrophia geniculata new species 
526s. Sly Gap Formation 
North side of Capitol Peak, San Andres Mountains, Capitol Peak 
(15') quadrangle, New Mexico. 
Costatrypa varicostata (Stainbrook) 
C. extensa new species 
Nervostrophia mucrospina new species 
Pseudoatrypa devoniana (Webster) 
Spinatrypa trulla decorticata new subspecies 
526t. Independence Shale Formation 
Quarry at Brandon, lowa. Collector: M. A. Stainbrook. 
Gypidula cornuta Fenton and Fenton 
G. stainbrooki new species 
§26u. Independence Shale Formation 
Benton County highway 71, 0.25 mile south of lowa highway 283, 
0.5 mile east of Brandon, lowa. 
Cariniferella costellata new species 


526v. Onate Formation 
NW corner, sec. 3, T. 17 S., R. 10 E., Sacramento Mountains, 
southeast of Alamogordo, Alamogordo (15’) quadrangle, New 
Mexico. Collector: L. C. Pray. 
Platyterorhynchus obscurum new species 
526w. Independence Shale Formation 
Poor Farm, one mile northeast of Independence, Iowa. 
Stainbrookia infera (Calvin) 
526x. Sly Gap Formation (Bed F) 
Same as localities 506w, 3054. 
Coeloterorhynchus magnificum new species 
Tenticospirifer cf. T. cyrtinaformis (Hall and Whitfield) 
526y. Independence Shale Formation 
West Amana, Iowa. Collector: M. A. Stainbrook. 
Nervostrophia extensa new species 
526z. Onate Formation 
Bend of road near Little Tank, Johnson Park Canyon, S¥2NW'%4 
sec. 5, T. 10 S., R. 5 E., north end of San Andres Mountains, 
Capitol Peak (15’) quadrangle, New Mexico. Collector: F. V. Ste- 
venson. Mocking Bird Gap of F. V. Stevenson (1943, p. 222). 
Acutoria angulata new species 
Orthospirifer species 
527a. Percha Formation (Box Member) 
South side of Bear Mountain, along Bear Mountain road, SE%4 
sec. 13 and SW sec. 18, T. 17 S., R. 14 and 15 W., Silver City 
(742') quadrangle, New Mexico. 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer animasensis (Girty) 
C. convexus new species 
C. rotundus new species 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Thiemella platys new species 
527b. Percha Formation (Box Member) 
North of Bear Mountain road, center E%2SE'% sec. 13, T. 17S., 
R. 15 W., southeast side of Bear Mountain, Silver City (7!2’) 
quadrangle, New Mexico. Collectors: R. E. Grant and G. A. Coo- 
per. 
Schizophoria australis Kindle 
§27c. Percha Formation (Box Member) 
Lake Valley, Lake Valley quadrangle, New Mexico. 
Cyrtiorina kindlei (Stainbrook) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
527d. Percha Formation (Box Member) 
Wilson Ranch, NE44NWI4NW'44 sec. 11, T. 18 S., R. 7 W., Hills- 
boro (15’) quadrangle, New Mexico. Collectors: F. V. Stevenson, 
A. L. Bowsher and W. T. Allen. 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 


DEVONIAN BRACHIOPODS OF NEW MExICO: COOPER AND DUTRO 139 


527d. (continued) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes perchaensis new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
S. spinigera (Kindle) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 


527e. Percha Formation (Box Member) 
Bear Mountain road, 5.4 miles north of junction of Alabama and 
Atlantic Streets, Silver City, Silver City (7'2') quadrangle, New 
Mexico. Collector: F. V. Stevenson. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Hispidaria spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 


527f. Percha Formation (Box Member) 
New Mexico highway 180 (90), 5 miles east of Santa Rita, New 
Mexico. 
Acanthatia nupera (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Leioproductus plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Ptychomalotoechia sobrina (Stainbrook) 
Retichonetes obscurus new species 
Schizophoria australis Kindle 
Sentosia spinigera (Kindle) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
527g. Percha Formation (Box Member) 
Chloride Flat, 1.5 miles by road west of Silver City, Silver City 
(7¥2') quadrangle, New Mexico. Collector: F. V. Stevenson. 
Cyrtiorina kindlei (Stainbrook) 


Leioproductus plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
527h. Percha Formation (Box Member) 
0.6 mile N. 60° E. of Wilson Ranch, SW'4SW'4 sec. 2, T. 18 S., 
R. 7 W., Hillsboro (15') quadrangle, New Mexico. 
Composita bellula Stainbrook 
Petasmaria patens new species 
Ptychomalotoechia sobrina (Stainbrook) 
527i. Percha Formation (Box Member) 
One mile north of Lake Valley, New Mexico. 
Porostictia perchaensis (Stainbrook) 
527j.  Percha Formation (Box Member) 
2 miles S. 50° E. of Hillsboro, Hillsboro (15’) quadrangle, New 
Mexico. Collector: L. A. Nelson (=loc. 507h). 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
527k. Percha Formation (Box Member) 
0.3 mile west-southwest of J. P. Nunn Ranch, NW corner sec. 
10, T. 17 S., R. 8 E., Hillsboro (15’) quadrangle, New Mexico. 
Syringospira prima Kindle 
5271. Percha Formation (Box Member—upper 25 feet) 
Bear Mountain, NE'4SE'4SE'% sec. 13, T. 17 S., R. 15 W., Silver 
City (742') quadrangle, New Mexico. 
Athyris transversa (Stainbrook) 
Leioproductus varispinosus (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Strophopleura notabilis (Kindle) 
527m. Same as locality 5271] 
527n. Percha Formation (Ready Pay Member) 
Below the lowest cobbly limestone, 20-30 feet above the base of 
the green shale (top of black shale), west side of Bear Mountain, 
SE corner sec. 3, T. 17 S., R. 15 W., Silver City (712) quadrangle, 
New Mexico. Collector: A. L. Bowsher. 
Cyrtiorina kindlei (Stainbrook) 
Petasmaria patens new species 
5270. Percha Formation (Ready Pay Member) 
15 feet above the base of the green shale or top of the black shale, 
northwest corner of Bear Mountain (=loc. 527n). 
527p. Percha Formation (Box Member) 
South side of Bear Mountain, NE'4SE' sec. 13, T. 17 S., R. 15 
W., Silver City (742') quadrangle, New Mexico. Collectors: G. A. 
Cooper and R. E. Grant (=loc. 5071). 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Cyrtospirifer rotundus new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Retichonetes perchaensis new species 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Thiemella platys new species 


BULLETIN 315 


527q. Contadero Formation 
South side of Hembnillo Canyon, San Andres Mountains, Kaylor 
Mountain (15’) quadrangle, New Mexico. Collector: R. Flower. 
Thiemella quadrata new species 
§27r. Sly Gap Formation 
Sly Gap, San Andres Mountains, Capitol Peak (15’) quadrangle, 
New Mexico (=loc. 521d). 
Aulacella elliptica new species 
527s. Independence Shale Formation 
Lime Creek, 1.25 miles northeast of Brandon, Iowa. 
Strophonelloides deeringi Stainbrook 
527t. Sly Gap Formation (lower 30 feet) 
Rhodes Canyon, San Andres Mountains, Black Top Mountain 
(15’) quadrangle, New Mexico. Collector: R. Flower. 
Strophonelloides dorsoconvexus new species 
527u. Onate Formation 
Rhodes Canyon, San Andres Mountains, Black Top Mountain 
(15’) quadrangle, New Mexico. Collector: R. Flower. 
Nervostrophia plana new species 
527v. Contadero Formation (Coral Bed) 
Sly Gap, San Andres Mountains, Capitol Peak (15’) quadrangle, 
New Mexico. Collector: R. Flower. 
Douvillina species 2 
527w. Onate Formation 
Hermosa, same as locality 526b. Collector: R. Flower. 
Devonochonetes? species 
527x. Onate Formation (basal 5 feet) 
Hermosa, same as locality 526b. Collector: R. Flower. 
Rhyssochonetes johnsoni new species 
527y. Hackberry Formation (Cerro Gordo Member—Strophonella 
Zone) 
0.4 mile south of Bird Hill, in ditch, 5 miles west of Rockford, 
Floyd County, Iowa. Donor: C. O. Levorson, Riceville, Iowa. 
Eostrophalosia rockfordensis (Hall and Clarke) 
§27z. Rhodes Canyon Formation 
Rhodes Canyon, San Andres Mountains, Black Top Mountain 
(15') quadrangle, New Mexico. Collectors: R. Flower and F. V. 
Stevenson. 
Productella species | 
528a. Contadero Formation 
Same as locality 527z. Collector: R. Flower. 
Afilasma inexpectatum new species 
Ripidiorhynchus newmexicoense new species 
528b. Contadero Formation 
Just below the purple shale, north side of Hembrillo Canyon, San 
Andres Mountains, Kaylor Mountain (15’) quadrangle, New Mex- 
ico. Collector: R. Flower. 
Colophragma ellipticum new species 
§28c. Rhodes Canyon Formation 
East end of Rhodes Pass, San Andres Mountains, Black Top 
Mountain (15’) quadrangle, New Mexico. Collector: M. A. Stain- 
brook. 
Crinisarina reticulata (Stainbrook) 
Cyrtospirifer breviposticus Stainbrook 
Evanescirostrum glabraventrum (Stainbrook) 
Tylothyris novamexicana Stainbrook 
528d. Hackberry Formation (Cerro Gordo Member) 
Tile Yard, southwest of Rockford, Floyd County, Iowa. Collector 
and Donor: C. O. Levorson, Riceville, lowa. 
Calvinaria bransoni Stainbrook 
528e. Same as locality 522f. 
Desquamatia costata new species 


528f. Independence Shale Formation 
Cedar River, 2.5 miles south and 1.5 miles due west of Brandon, 
Iowa. Collector: M. A. Stainbrook. 
Towatrypa americana (Stainbrook) 
528g. Contadero Formation 
Same as locality 527v. Collector: R. Flower. 
Spinatrypa compacta new species 
528h. Independence Formation 
West Amana, lowa County, Iowa. Collector: C. H. Belanski 
Spinatrypa trulla (Stainbrook) 
528i. Onate Formation 
Lost Canyon, Hermosa, Sugarloaf Peak (15’) quadrangle, New 
Mexico. Collector: R. Flower. 
Emanuella species 
528). Rhodes Canyon Formation 
Rhodes Canyon, same locality as 527u. Collector: R. Flower. 
Cranaena navicella (Hall) 
Crinisarina reticulata (Stainbrook) 
Cyrtospirifer pyramidalis new species 
Tylothyris novamexicana Stainbrook 
528k. Onate Formation 
Engle Camp, Hermosa, Sugarloaf Peak (15’) quadrangle, New 
Mexico. Collector: R. Flower. 
Mucrospirifer parvus new species 
5281. Independence Shale Formation 
Brandon, Iowa. Collector: M. A. Stainbrook. 
Thomasaria altumbona Stainbrook 
528m. Independence Formation 
Same as locality 526t. 
Thomasaria altumbona Stainbrook 
528n. Onate Formation (above Chonetes Bed) 
Same as locality 526b. Collector: R. Flower 
Cyrtina variabilis new species 
5280. Probably Contadero Formation 
Sly Gap section, same as locality 5090. Collector: R. Flower. 
Productella? species 2 
528p. Sly Gap Formation (2 feet above “‘Chonetes’’) 
Hermosa, same as locality 526b. Collector: R. Flower. 
Costatrypa varicostata (Stainbrook) 
Thomasaria demissa new species 
528q. Contadero Formation (Zone G) 
Same as localities 52le, 521j. Collector: F. V. Stevenson. 
Tylothyris species 1 
528r. Sly Gap Formation 
Same as locality 523p. 
Hypothyridina emmonsi rotunda new subspecies 
528s. Onate Formation 
Same as locality 509p. 
Hypothyridina species 
528t. Onate Formation (‘‘Spirifer’’ beds) 
Johnson Park Canyon, north end of the San Andres Mountains, 
Capitol Peak (15’) quadrangle. New Mexico. 
Mucrospirifer? cf. M.? sculptilis (Hall) 
Nervostrophia plana new species 
Acutoria angulata new species 
528u. Onate Formation 
Alamo Canyon, Sacramento Mountains, Alamogordo (15’) quad- 
rangle, New Mexico. Collector: F. V. Stevenson (=loc. 529j). 
Platyterorhynchus obscurum new species 
§28v. Sly Gap Formation (below lowest occurrence of the coral 
Macgeea) 


DEVONIAN BRACHIOPODS OF NEW MEXIco: COOPER AND DUTRO 141 


528v. (continued) 
Sly Gap, San Andres Mountains, Capitol Peak (15’) quadrangle, 
New Mexico. Collector: R. Flower. 
Rhipidomella species | 
528w. Sly Gap Formation 
Alamo Canyon, Sacramento Mountains, Alamogordo (15’) quad- 
rangle, New Mexico. Collector: F. V. Stevenson. 
Schizophoria amanaensis Stainbrook 
528x. Sly Gap Formation 
East side of Alamo Peak, 2 mile west of Holmes Mine, center 
EVaNWM4NEM“SE sec. 25, T. 16 S., R. 10 E., Lincoln Forest 
Service Map, east of Alamogordo, New Mexico. Collector: A. L. 
Bowsher. 
Strophonelloides parvulus new species 
528y. Sly Gap Formation 
Deadman Canyon, San Andres Mountains, sec. 8, T. 17 S., R. 4 
E., Kaylor Mountain (15’) quadrangle, New Mexico. Collector: 
F. V. Stevenson. 
Aulacella elliptica new species 
Douvillina parviuscula new species 
Nervostrophia solida new species 
Schizophoria amanaensis Stainbrook 
Thomasaria demissa new species 
529g. Onate Formation (lower 5 feet) 
Johnson Park Canyon, north end of San Andres Mountains, Cap- 
itol Peak (15') quadrangle, New Mexico (=loc. 528t). 
Mucrospirifer? cf. M.? sculptilis (Hall) 
Nervostrophia plana new species 
Spinocyrtia? species 
Tropidoleptus platys new species 
5291. Sly Gap Formation 
15 feet below the base of the Mississippian, Johnson Park Can- 
yon, Capitol Peak (15’) quadrangle, New Mexico. 
Ambocoelia species 4 
529}. Onate Formation 
Alamo Canyon, Sacramento Mountains, Alamogordo (15’) quad- 
rangle, New Mexico. Collector: F. V. Stevenson. 
Desquamatia costata new species 
Dichacaenia umbonata new species 
Mucrospirifer parvus new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 


529k. Onate Formation (6-8 feet below the top) 
Deadman Canyon, Sacramento Mountains, Alamogordo (15’) 
quadrangle, New Mexico. 
Desquamatia costata new species 
Nervostrophia plana new species 
Platyterorhynchus obscurum new species 
529]. Onate Formation 
Nackaye Mines, northeast of Derry, Upham (15’) quadrangle, 
New Mexico (=loc. 523d). 
Ambocoelia species 4 
$29m. Onate Formation 
Agua Chichita Canyon, Sacramento Mountains, New Mexico. 
Collector: F. V. Stevenson. 
Ambocoelia species 4 
Nervostrophia plana new species 
Rhyssochonetes johnsoni new species 
529n. Onate Formation 
2.5 miles up Alamo Canyon from its mouth, Alamogordo (15’) 
quadrangle, New Mexico. 
Lingula species 3 
Nervostrophia plana new species 


Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
5290. Onate Formation 
Fault block, Marble Canyon, Alamogordo (15’) quadrangle, New 
Mexico. Collector: F. V. Stevenson. 
Desquamatia costata new species 
Mucrospirifer parvus new species 
529p. Sly Gap Formation 
Same as locality 526b. 
Costatrypa varicostata new species 
Thomasaria demissa new species 
529q. Onate Formation (upper) 
Deadman Canyon, center NW!4SE'% sec. 12, T. 17 S., R. 3 E., 
Kaylor Mountain (15’) quadrangle, New Mexico. 
Platyterorhynchus obscurum new species 
529r. Onate Formation 
Marble Canyon, Alamogordo (15’) quadrangle, New Mexico. Col- 
lector: F. V. Stevenson. 
Lingula species 4 
529s. Sly Gap Formation (Bed I) 
Center NW'4SE'% sec. 12, T. 17 S., R. 3 E., Deadman Canyon, 
San Andres Mountains, Kaylor Mountain (15’) quadrangle, New 
Mexico. 
Cariniferella alleni new species 
Coeloterorhynchus magnificum new species 
Cyrtina inulta Stainbrook 
Elita magister new species 
Spinatrypa trulla decorticata new subspecies 
529t. Onate Formation (Bed C) 
Bear Canyon, Lake Lucero SW (7!4’) quadrangle, New Mexico. 
Platyterorhynchus obscurum new species 
529u. Sly Gap Formation 
South side of Deadman Canyon, San Andres Mountains, center 
W!2 sec. 8, T. 17S., R. 4 W., Kaylor Mountain (15') quadrangle, 
New Mexico. Collector: F. V. Stevenson. 
Costatrypa varicostata (Stainbrook) 
529v. Sly Gap Formation 
San Andres Canyon, San Andres Mountains, New Mexico. 
Pseudoatrypa devoniana (Webster) 
529w. Onate Formation 
Arcente Canyon, Alamogordo (15’) quadrangle, New Mexico. 
Collector: F. V. Stevenson. 
Ambocoelia species 4 
Platyterorhynchus obscurum new species 
Rhipidomella species 2 
529x. Sly Gap Formation (10-14 feet above the Onate) 
Same as locality 506c. 
Calvinaria bransoni Stainbrook 
Coeloterorhynchus magnificum new species 
529y. Sly Gap Formation (Bed A—S-10 feet above base) 
NSW" sec. 9, T. 12 S., R. 7 W., 2 miles N. 60° W. of Chise, 
Sierra County (1960) map, New Mexico. 
Costatrypa extensa new species 
C. varicostata (Stainbrook) 
Schizophoria amanaensis Stainbrook 
USGS 459. Percha Formation (Box Member) 
Northeast of Santa Rita, New Mexico. Collector: H. A. Thorne. 
Cyrtiorina kindlei (Stainbrook) 
USGS 460. Percha Formation (Box Member) 
Mining District, Kingston, New Mexico. Collector: L. C. Graton. 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
USGS 461. Percha Formation (Box Member) 
Same as locality 507h. Collector: L. C. Graton. 


142 BULLETIN 315 


USGS 508. Percha Formation (Box Member) 
Silver City quadrangle, New Mexico. Collectors: G. H. Girty and 
S. Paige. 
Acanthatia nupera (Stainbrook) 
Laminatia laminata (Kindle) 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Schizophoria australis Kindle 
Strophopleura notabilis (Kindle) 
Toryniferella spinosa (Kindle) 
USGS 509. Percha Formation (Box Member) 
Silver City quadrangle, New Mexico. Collectors: G. H. Girty and 
S. Paige. 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer rotundus new species 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Strophopleura notabilis (Kindle) 
USGS 510. Percha Formation (Box Member) 
Silver City quadrangle, New Mexico. Collectors: G. H. Girty and 
S. Paige. 
Sentosia praecedens (Stainbrook) 
USGS 512. Percha Formation (Box Member) 
Lake Valley, New Mexico. Collector and Donor: F. Springer. 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus plicatus (Kindle) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Schizophoria australis Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
USGS 534. Sly Gap Formation 
North slope of Sheep Mountain, San Andres Mountains, New 
Mexico. Collector: N. H. Darton. 
Costatrypa varicostata (Stainbrook) 
Devonoproductus intermedius new species 
Thomasaria demissa new species 
USGS 536. Sly Gap Formation 
San Andres Canyon, San Andres Mountains, New Mexico. Col- 
lector: N. H. Darton. 
Elita magister new species 
USGS 538. Sly Gap Formation 
One mile south of Capitol Peak, San Andres Mountains, New 
Mexico. Collector: N. H. Darton. 
Costatrypa varicostata (Stainbrook) 
C. extensa new species 
Devonoproductus intermedius new species 
Nervostrophia solida new species 
N. mucrospina new species 
Pseudoatrypa devoniana (Webster) 
USGS 540. Onate Formation 
One mile south of Capitol Peak, San Andres Mountains, New 
Mexico. Collector: N. H. Darton. 
Mucrospirifer? cf. M.? sculptilis (Hall) 
Tropidoleptus platys new species 
USGS 541. Sly Gap Formation 
Hembrillo Canyon, San Andres Mountains, New Mexico. Col- 
lector: N. H. Darton. 
Coeloterorhynchus magnificum new species 
Gypidula bowsheri new species 
USGS 543. Sly Gap Formation 
Sulphur Gap, San Andres Mountains, New Mexico. Collector: N. 
H. Darton. 
Aulacella elliptica new species 


Coeloterorhynchus magnificum new species 
Cyrtospirifer species 4 
Douvillina parviuscula new species 
USGS 4804 (Kindle). Percha Formation (Box Member) 
Kingston, New Mexico. Collectors: G. H. Girty and S. Paige. 
Syringospira prima Kindle 
3013. Onate Formation (6 feet below ruddy brown bed at top) 
Deadman Canyon, same as locality 526p. 
Platyterorhynchus obscurum new species 
3016. Sly Gap Formation 
Type section of the Sly Gap Formation, Sheep Mountain, sec. 
25, T. 11 S., R. 15 E., San Andres Mountains, Capitol Peak (15’) 
quadrangle, New Mexico. Collectors: W. T. Allen and A. L. 
Bowsher. 
Aulacella elliptica new species 
Nervostrophia mucrospina new species 
3023. Onate Formation (See 3023b, c, and d below) 
3023b. Onate Formation 
One mile S. 6° W. of Little Tank, Johnson Park Canyon, S¥24NW'4 
sec. 5, T. 10 S., R. 5 E., north end of San Andres Mountains, 
Capitol Peak (15’) quadrangle, New Mexico. Collector: A. L. 
Bowsher (=loc. 526z). For conodonts see USGS loc. 9208-SD. 
Mucrospirifer? cf. M.? sculptilis (Hall) 
Tropidoleptus platys new species 
3023c. Onate Formation 
One mile S. 6° W. of Little Tank, Johnson Park Canyon, S'% 
NW4 sec. 5, T. 10 S., R. 5 E., north end of San Andres Moun- 
tains, Capitol Peak (15’) quadrangle, New Mexico. Collector: A. 
L. Bowsher (=loc. 526z). 
Acutoria angulata new species 
Cyrtina variabilis new species 
Orthospirifer species 
Pentamerella species 
Platyterorhynchus obscurum new species 
Schizophoria lata Stainbrook 
3023d. Onate Formation 
One mile S. 6° W. of Little Tank, Johnson Park Canyon, S2NW'4 
sec. 5, T. 10 S., R. 5 E., north end of San Andres Mountains, 
Capitol Peak (15') quadrangle, New Mexico. Collector: A. L. 
Bowsher (=loc. 5262). 
Nervostrophia plana new species 
3025. Sly Gap Formation 
Same as locality S06y. 
Nervostrophia solida new species 
Hyborhynchella bransoni Cooper 
3025a. Onate Formation 
Same as locality S06y. 
Atryparia rubra new species 
Eleutherokomma extensa new species 
Hadrorhynchia alta new species 
Leiorhynchus sphaericum new species 
Platyterorhynchus obscurum new species 
Striatochonetes nanus new species 
3026. Percha Formation (Box Member) 
0.6 mile N. 65° E. of Earl Wilson Ranch, SE4SE%4SW' sec. 11, 
T. 18 S., R. 7 W., Hillsboro (15’) quadrangle, New Mexico. Col- 
lector: A. L. Bowsher. 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Hispidaria spinosa new species 
Leioproductus pauperculus new species 


DEVONIAN BRACHIOPODS OF NEW MExICO: COOPER AND DuTRO 143 


3026. (continued) 


L. plicatus (Kindle) 

L. varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Petrocrania ourayensis (Kindle) 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 

Toryniferella spinosa (Kindle) 
Falciclymenia bowsheri (Miller and Collinson) 1951* 


3026a. Percha Formation (Box Member) 


West slope of scarp east of tank, 0.7 mile N. 60° E. of Earl Wilson 
Ranch, SW'4SW'4 sec. 2, T. 18 S., R. 7 W., Hillsboro (15’) quad- 
rangle, New Mexico. Collector: A. L. Bowsher. 

Acanthatia nupera (Stainbrook) 

Athyris transversa (Stainbrook) 

Composita bellula Stainbrook 

Cyrtiorina kindlei (Stainbrook) 

Cyrtospirifer convexus new species 

Galeatella galeata Muir-Wood and Cooper 

Hispidaria posterogranulosa new species 

H. spinosa new species 

Laminatia laminata (Kindle) 

Leioproductus plicatus (Kindle) 

L. varispinosus (Stainbrook) 

Oiosia putilla (Stainbrook) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

Petrocrania ourayensis (Kindle) 

Planoproductus hillsboroensis (Kindle) 

Porostictia perchaensis (Stainbrook) 

Ptychomalotoechia sobrina (Stainbrook) 

Schizophoria australis Kindle 

Schuchertella percha (Stainbrook) 

Sentosia spinigera (Kindle) 

Strophopleura notabilis (Kindle) 

Syringospira prima Kindle 

Toryniferella spinosa (Kindle) 


3029. Percha Formation (Box Member) 


East facing scarp on the west side of gap on Tierra Blanca Creek 
road, 2.2 miles N. 75° W. of J. P. Nunn Ranch, 1.2 miles N. 30° 
W. of Tierra Blanca Mountain, SE'4SE"% sec. 5, T. 17 S., R. 8 
W.. Hillsboro (15') quadrangle, New Mexico. Collector: A. L. 
Bowsher. 

Acanthatia nupera (Stainbrook) 

Ambocoelia capillata new species 

Composita bellula Stainbrook 

Cyrtiorina kindlei (Stainbrook) 

Cyrtospirifer rotundus new species 

Eoparaphorhynchus mclareni Sartenaer 

Leioproductus plicatus (Kindle) 

L. varispinosus (Stainbrook) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

Planoproductus hillsboroensis (Kindle) 

Ptychomalotoechia sobrina (Stainbrook) 

Schizophoria australis Kindle 


Sentosia praecedens (Stainbrook) 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 

3030. Percha Formation (Box Member) 

North of Trujillo Creek, on west facing scarp along canyon run- 
ning north from Trujillo Creek, 5 miles S. 50° E. of Hillsboro, 
center, south boundary, sec. 26, T. 16 S., R. 7 W., Hillsboro (15’) 
quadrangle, New Mexico. Collector: A. L. Bowsher. 

Composita bellula Stainbrook 

Cyrtiorina kindlei (Stainbrook) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

Ptychomalotoechia sobrina (Stainbrook) 

3031. Percha Formation (Box Member) 

Fork of Percha Creek, SW'4SE'4 sec. 9, and N12NE'4, sec. 16, 
T. 16 S., R. 8 W., Hillsboro (15') quadrangle, New Mexico. Col- 
lector: A. L. Bowsher. 

Leioproductus varispinosus (Stainbrook) 

Retichonetes obscurus new species 

303la. Percha Formation (Box Member) 

N'’%SW'4SE'4 sec. 9, T. 16 S., R. 8 W., Hillsboro (15’) quadran- 
gle, New Mexico. Collector: A. L. Bowsher. 

Athyris transversa (Stainbrook) 

Cyrtiorina kindlei (Stainbrook) 

Laminatia laminata (Kindle) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

Schizophoria australis Kindle 

Syringospira prima Kindle 

3031b. Percha Formation (lower Box or uppermost Ready Pay 

members) 

Gully on north side of road, SE44NW'4SE'4 sec. 9, T. 16 S., R. 
8 W., Hillsboro (15’) quadrangle, New Mexico. Collector: A. L. 
Bowsher. 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Leioproductus varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Ptychomalotoechia sobrina (Stainbrook) 
Syringospira prima Kindle 
3034. Percha Formation (Box Member) 
Same as locality 507h. 

3035. Percha Formation (Box Member) 

0.44 mile west of Kingston and to the north of Ladrone Gulch, 
SE'4NW'4 sec. 18, T. 16 S., R. 8 W., Hillsboro (15’) quadrangle, 
New Mexico. Collector: A. L. Bowsher. 

Acanthatia nupera (Stainbrook) 

Athyris transversa (Stainbrook) 

Cyrtiorina kindlei (Stainbrook) 

Leioproductus plicatus (Kindle) 

Paurorhyncha cooperi Stainbrook 

Petasmaria patens new species 

Planoproductus hillsboroensis (Kindle) 

Schizophoria australis Kindle 

Schuchertella percha (Stainbrook) 

Toryniferella spinosa (Kindle) 

3037. Percha Formation (Ready Pay Member—basal 10-20 feet) 
In gully, north side of road, 1.2 miles south of Wilson Ranch, 
Hillsboro (15') quadrangle, New Mexico. Collector: A. L. Bow- 
sher. 

Lingulipora porifera new species 


* This specimen, an ammonite, is in the National Museum of 3042. Percha Formation (Box Member) 
Natural History (USNM 116343). It is a rare but important specimen Northwest slope of Bear Mountain, SW!'4NW'4 sec. 11, T. 17 S., 
because it links the Box Member of the Percha to the Clymenia R. 15 W., Silver City (72') quadrangle, New Mexico. Collector: 


Zone of the German Devonian. A. L. Bowsher. 


144 BULLETIN 315 


3042. (continued) 
Acanthatia nupera (Stainbrook) 
Athyris transversa (Stainbrook) 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Galeatella galeata Muir-Wood and Cooper 
Hispidaria posterogranulosa new species 
H. spinosa new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
L. varispinosus (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Schuchertella percha (Stainbrook) 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
3042s. Percha Formation (Box Member) 
East of Bear Mountain road, 0.9 mile southeast of Bear Mountain, 
E'% sec. 13, T. 17 S., R. 15 W., Silver City (7/2') quadrangle, 
New Mexico. Collector: A. L. Bowsher. 
Athyris transversa (Stainbrook) 
Composita bellula Stainbrook 
Cyrtiorina kindlei (Stainbrook) 
Cyrtospirifer convexus new species 
Laminatia laminata (Kindle) 
Leioproductus pauperculus new species 
L. plicatus (Kindle) 
Oiosia putilla (Stainbrook) 
Paurorhyncha cooperi Stainbrook 
Petasmaria patens new species 
Planoproductus hillsboroensis (Kindle) 
Porostictia perchaensis (Stainbrook) 
Ptychomalotoechia sobrina (Stainbrook) 
Schizophoria australis Kindle 
Sentosia praecedens (Stainbrook) 
Strophopleura notabilis (Kindle) 
Syringospira prima Kindle 
Toryniferella spinosa (Kindle) 
Trifidorostellum obscurum new species 
3043. Percha Formation 
Same as locality 5090. 
3045. Sly Gap Formation 
Arcente Canyon, west side road, 6.2 miles south of High Rolls, 
SEY44ANW'I4NE'4 sec. 20, T. 16 S., R. 11 E., Lincoln National 
Forest Service Map, New Mexico. Collector: A. L. Bowsher 
(=loc. 509m). 
3045b. Sly Gap Formation 
Arcente Canyon, west side of road, 6.2 miles south of High Rolls, 
SEY“NW'ANE sec. 20, T. 16 S., R. 11 E., Lincoln National 
Forest Service Map, New Mexico: Collector: A. L. Bowsher (=loc. 
509m). 
Acutatheca prolifica new species 
Gypidula bowsheri new species 
Pseudodouvillina euglyphea Stainbrook 
Thomasaria warreni new species 


3045c. Sly Gap Formation 
Arcente Canyon, west side of road, 6.2 miles south of High Rolls, 
SEY4NWIANE sec. 20, T. 16 S., R. 11 E., Lincoln National 
Forest Service Map, New Mexico. Collector: A. L. Bowsher (=loc. 
509m). 
Schizophoria amanaensis Stainbrook. 
3045g. Sly Gap Formation 
Arcente Canyon, west side of road, 6.2 miles south of High Rolls, 
SEY4ANWI4ANE sec. 20, T. 16 S., R. 11 E., Lincoln National 
Forest Service Map, New Mexico. Collector: A. L. Bowsher (=loc. 
509m). 
Spinatrypa trulla decorticata new subspecies 
3052a. Onate Formation 
Pig Canyon Anticline, 1.8 miles N. 7° E. of Ortega Peak, 4 miles 
N. 80° E. of Alamogordo, SW'4NE!4SW'4 sec. 7, T. 16 S., R. 
11 E., Lincoln Forest Service Map, New Mexico. Collector: A. 
L. Bowsher. 
Cyrtina variabilis new species 
Eleutherokomma extensa new species 
Hadrorhynchia mclareni new species 
Platyterorhynchus obscurum new species 
3053. Onate Formation (Zone I) 
Same as locality 522d. 
3054. Sly Gap Formation 
0.25 mile north of Rhodes Canyon highway, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Cariniferella alleni new species 
Costatrypa extensa new species 
Spinatrypa trulla decorticata new subspecies 
3054b. Sly Gap Formation 
0.25 mile north of Rhodes Canyon highway, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15') quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Costatrypa varicostata Stainbrook 
Cyrtina inulta Stainbrook 
Strophodonta species 2 
3054c. Sly Gap Formation 
0.25 mile north of Rhodes Canyon highway, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Costatrypa extensa new species 
Tylothyris sulcocostata new species 
3054f. Sly Gap Formation 
0.25 miles north of Rhodes Canyon highway, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Calvinaria bransoni Stainbrook 
Douvillina parviuscula new species 
Pseudoatrypa devoniana (Webster) 
3054g. Sly Gap Formation 
0.25 mile north of Rhodes Canyon highway, SW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Floweria prava (Hall) 
3054h. Sly Gap Formation 
0.25 mile north of Rhodes Canyon highway, NW'4 sec. 3, T. 13 
S., R. 4 E., Black Top Mountain (15’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Hyborhynchella bransoni Cooper 
Thomasaria warreni new species 
30541. Sly Gap Formation (beds with Pachyphyllum) 
0.25 mile north of Rhodes Canyon highway, NW'4 sec. 3, T. 13 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 145 


30541. (continued) 
S., R. 4 E., Black Top Mountain (15) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 
Costatrypa varicostata (Stainbrook) 
Elita magister new species 
Schizophoria amanaensis Stainbrook 
3054k. Contadero Formation 
Same as locality 524t. 
Ambocoelia species 2 
Douvillina contaderoensis new species 


Ripidiorhynchus newmexicoense new species 
Spinatrypa obsolescens new species 
3054q. Rhodes Canyon Formation 

0.25 miles north of Rhodes Canyon highway, NW'4 sec. 3, T. 13 
S. R. 4 E., Black Top Mountain (15S’) quadrangle, New Mexico. 
Collector: A. L. Bowsher. 

Crinisarina reticulata (Stainbrook) 

Cyrtospirifer pyramidalis new species 

Productella species | 


REFERENCES CITED 


Alekseeva, R. E. 

1960. Ona new Subgenus of Atrypa (Desquamatia) subgen. nov. 
Family Atrypidae Gill. Akad. Nauk SSSR, Dokl. 131, No. 
2, pp. 421-424. 

Alichova, T. N. 

1960. Otryad Orthida: Mshanki, Brakhiopody [T. G. Sarycheva, 
ed.] [pp. 183-197, pls. 7-19, text-figs. 85-97] in Osnovy 
Paleontologii (Orlov, Y. ed.| Moscow. 

Anderson, W. I. 

1966. Upper Devonian Conodonts and the Devonian-Mississip- 
pian Boundary of North-central lowa. J. Paleontol., vol. 
40, No. 2, pp. 395-415, pls. 48-52, 3 text-figs. 

Austin, T., and Austin, T. Jr. 

1844. A monograph on Recent and fossil Crinoidea. London and 

Bristol. 128 pp., 16 pls. 
Belanski, C. H. 

1928. The Shellrock Stage of the Devonian of Iowa. Am. Midl. 

Nat., vol. 9, No. 5, pp. 165-212, pls. 12-17. 
Billings, E. 

1860. On the Devonian fossils of Canada West. Canadian J., 

n.s., Vol. 5, pp. 249-282, figs. 26-28. 
Boucot, A. J. 

1959. Brachiopods of the Lower Devonian rocks at Highland 
Mills, New York. J. Paleontol., vol. 33, No. 5, pp. 727- 
769, pls. 90-103. 

Boucot, A. J., Johnson, J. G., and Staton, R. D. 

1964. On some Atrypoid, Retzioid, and Athyidoid Brachiopoda. 

J. Paleontol., vol. 38, No. 5, pp. 805-822. 
Bowsher, A. L. 

1967. The Devonian System of New Mexico. Tulsa Geol. Soc. 

Dig., vol. 35, pp. 259-276, 7 figs. 
Branson, C. C., and Mehl, M. G. 

1934a. Conodont studies No. 3, Conodonts from the Grassy Creek 
shale of Missouri: Missouri Univ. Stud., vol. 8, No. 3, 
pp. 171-259, pls. 13-21. 

1934b. Conodont studies No. 4, Conodonts from the Bushberg 
sandstone and equivalent formations of Missouri. Mis- 
souri Univ. Stud., vol. 8, No. 4, pp. 265-300, pls. 22-24. 

1938. The conodont genus Icriodus and its stratigraphic distri- 
bution. J. Paleontol., vol. 12, No. 2, pp. 156-166, pl. 26. 


Branson, E. R. 
1934. Conodonts from the Hannibal formation of Missouri. Mis- 
souri Univ. Stud., vol. 8, pt. 2, No. 4, pp. 301-343, pls. 
25-28. 
Brice, D. 
1970. Etude Paléontologique et Stratigraphique du Devonien de 
U Afghanistan. Mus. Nat. d’Hist. Nat., Notes et Mem. 
Moyen-Orient. Paris, 364 pp., 20 pls., 65 figs. 


Bronn, H. G. 

1862. Die Klassen und Ordnungen der Weichtiere (Malacozoa). 

vol. 3, No. 1, 518 pp., 40 pls. Leipzig and Heidelberg. 
Brown, T. 

1849. Illustrations of the Fossil Conchology of Great Britain and 
Ireland with Descriptions and Localities of all Species. 
London. 273 pp., 98 pls. 

Bruguiere, J. G. 

1797. Encyclopédie Méthodique. Tableau Encyclopédique et 
Meéthodique des Trois Régnes de la Nature. Vers, Co- 
qilles, Mollusques et Polypiers. vol. 2, pls. 96-314. 

Bryant, W. L. 

1921. The Genesee conodont with descriptions of new species. 

Buffalo Soc. Nat. Hist., Bull., vol. 13, pp. 1-59, 16 pls. 
Buckman, S. S. 

1906. Brachiopod Nomenclature. Ann. Mag. Nat. Hist., Ser. 7, 

vol. 18, pp. 323-327. 
Calvin, S. 

1878. On some Dark Shale Recently Discovered below the De- 
vonian Limestones at Independence, lowa; with a notice 
of its Fossils and Descriptions of New Species. U.S. Geol. 
Surv. Terr. (Hayden), Bull., vol. 4, No. 3, pp. 725-730. 

1883. On the Fauna found at Lime Creek, lowa and its Relation 
to other Geological Faunas. Am. J. Sci., 3rd Ser., vol. 
25, pp. 432-436. 

1897. Geology of Cerro Gordo County, lowa. lowa Geol. Surv., 
vol. 7, pp. 117-195, maps. 

Carter, J. L. 

1967. Mississippian brachiopods from the Chappel Limestone 
of Central Texas. Bull. Am. Paleontol., vol. 53, No. 238, 
pp. 253-488, pls. 13-45, 56 figs., 45 tables. 

Caster, K. E. 

1939. A Devonian Fauna from Colombia. Bull. Am. Paleontol., 

vol. 24, No. 83, 218 pp., 14 pls. 
Chao, Y. T. 

1927. Productidae of China, Pt. 1. Producti. Palaeontol. Sinica, 

ser. B, vol. 5, fasc. 2, 192 pp., 16 pls. 
Cloud, P. E. 

1942. Terebratuloid Brachiopoda of the Silurian and Devonian. 
Geol. Soc. Am., Spec. Paper 38, i-xii + 182 pp., 26 pls., 
17 text-figs. 

Conrad, T. A. 

1839. Second annual report of the Paleontological Department 
of the survey. New York Geol. Surv., Ann. Rept. 3, pp. 
57-66. 

1842. Observations on the Silurian and Devonian Systems of the 
United States with Descriptions of New Organic remains. 
Acad. Nat. Sci., Philadelphia, J., vol. 8, pp. 228-280, pls. 
12-17. 


146 BULLETIN 315 


Cooper, C. L. 

1931. New Conodonts from the Woodford Formation of Okla- 
homa. J. Paleontol., vol. 5, No. 3, pp. 230-243, pl. 28. 

1939. Conodonts from a Bushberg-Hannibal horizon in Okla- 
homa. J. Paleontol., vol. 13, pp. 379-422, pls. 39-47. 

Cooper, G. A. 

1942. New Genera of North American Brachiopods. Washing- 
ton Acad. Sci., J., vol. 32, No. 8, pp. 228-235. 

1944. Phylum Brachiopoda [ch. 9] pp. 277-365, pls. 105-143 in 
Shimer, H. W., and Shrock, R. R., Index Fossils of North 
America. Wiley and Sons, New York. ix + 837 pp., 303 
pls. 

1945. New species of brachiopods from the Devonian of Illinois 
and Missouri. J. Paleontol., vol. 19, No. 3, pp. 479-489, 
pls. 63, 64. 

1954. Unusual Devonian Brachiopods. J. Paleontol., vol. 28, 
No. 3, pp. 325-332, pls. 36, 37, 5 text-figs. 

1955. New Genera of Middle Paleozoic Brachiopods. J. Paleon- 
tol., vol. 29, No. 1, pp. 45-63, pls. 11-14, 1 text-fig. 

Cooper, G. A., Butts, C., Caster, K. E., Chadwick, G. H., Goldring, 

W., Kindle, E. M., Kirk, E., Merriam, C. W., Swartz, F. M., 

Warren, P. S., Warthin, A. S., and Willard, B. 

1942. Correlation of the Devonian Sedimentary Formation|s | of 
North America. Geol. Soc. Am., Bull. 53, No. 12, pp. 
1729-1794, correlation chart. 

Cooper, G. A., and Williams, J. S. 

1935. Tully Formation of New York. Geol. Soc. Am., Bull. 46, 

No. 5, pp. 781-868, pls. 54-60. 
Copper, P. 

1966. European Mid-Devonian Correlations. Nature, vol. 209, 
No. 5027, pp. 982-984. 

1973. New Siluro-Devonian Atrypoid Brachiopods. J. Paleon- 
tol., vol. 47, No. 3, pp. 484-500, 3 pls., 3 text-figs. 

Crickmay, C. H. 

1950. Some Devonian Spiriferidae from Alberta. J. Paleontol., 
vol. 24, No. 2, pp. 219-255, pls. 36, 37. 

1952. Discrimination of late Upper Devonian. J. Paleontol., vol. 
26, No. 4, pp. 585-609, pls. 70-78. 

1953. Warrenella, a new genus of Devonian Brachiopods. J. Pa- 
leontol., vol. 27, No. 4, pp. 596-600, 25 text-figs. 

1960. The Older Devonian Faunas of the Northwest Territories. 
Evelyn de Mille Books, Calgary, Canada. 20 pp., 11 pls., 
text-figs. 25, 26. 

1963. Significant New Devonian Brachiopods from Western 
Canada. Evelyn de Mille Books, Calgary, Canada. 63 pp., 

16 pls. 
Dalman, J. W. 

1828. Uppstdllning och Beskrifning af de i Sverige funne Tere- 
bratuliter. K. Svensk. Vetenskapsakad. Handl. for 1827, 
pp. 85-155, pls. 1-6. 

Darton, N. H. 

1917. A Comparison of Paleozoic Sections in Southern New 
Mexico. U.S. Geol. Surv., Prof. Paper 108C, pp. 31-55, 
pls. 13-21. 

Davidson, T. 
1858-1863. The Carboniferous Brachiopoda. [|vol. 2, pt. 5, 280 
pp., 55 pls.] in A Monograph of the British Fossil Brach- 
iopoda. Palaeontogr. Soc. London [6 vols: 1853-1886]. 
Drewes, H. 
1967. Geology of the Connors Pass Quadrangle, Schell Creek 
Range, east-central Nevada. U.S. Geol. Suryv., Prof. Pa- 
per 557, pp. 25-29, table 3. 


Dunbar, C. O., and Condra, G. E. 

1932. Brachiopoda of the Pennsylvanian System of Nebraska. 
Nebraska Geol. Surv., Bull., 2nd Ser., vol. 51, 377 pp., 
44 pls., 25 text-figs. 

Ellison, S. P., Jr. 

1950. Subsurface Woodford Black Shale, West Texas and 
Southeast New Mexico. Texas Bur. Econ. Geol. Rep. In- 
vest., vol. 7, 20 pp. 3 pls., 6 figs. 

Epis, R. C., Gilbert, C. M., and Langenheim, R. L. 

1957. Upper Devonian Swisshelm Formation of Southeastern 
Arizona. Am. Assoc. Petrol. Geol., Bull., vol. 41, No. 10, 
pp. 2243-2256, 3 figs. 

Ethington, R. L. 

1965. Late Devonian and Early Mississippian Conodonts from 
Arizona and New Mexico. J. Paleontol., vol. 39, No. 4, 
pp. 566-589, pl. 67. 

Fenton, C. L. 

1919. The Hackberry Stage of the Upper Devonian of Iowa. 
Am. J. Sci., 4th Ser., vol. 48, pp. 355-376, 5 figs. 

1931. Studies of evolution in the genus Spirifer. Wagner Free 
Inst. Sci., Philadelphia, vol. 2, pp. 8-436, 50 pls., 204 figs. 

Fenton, C. L., and Fenton, M. A. 

1924. The Stratigraphy and Fauna of the Hackberry Stage of 
the Upper Devonian. Univ. Michigan Mus. Geol. Con- 
trib., vol. 1, xi + 260 pp.. 44 pls. 

1928. A new species of Schizophonia from the Devonian of Iowa. 
Am. Midl. Nat., vol. 11, Nos. 3/4, pp. 160-161, 1 pl. 

1930. Studies on the genus Atrypa. Am. Midl. Nat., Notre Dame, 
vol. 12, pp. 1-13, 2 pls. 

Ferguson, J. 

1966. Variation in two species of the Carboniferous brachiopod 
Pleuropugnoides. Yorkshire Geol. Soc., Proc., vol. 35, 
pp. 353-374, 1 pl., 12 figs. 

Flower, R. H. 

1958. Cambrian-Mississippian Beds of Southern New Mexico. 
Roswell Geol. Soc., 11th Field Conf., Guideb., pp. 61-78, 
2 figs. 

1959. Cambrian-Devonian Beds of Southern New Mexico. Ros- 
well Geol. Soc. and Permian Basin Sect., Soc. Econ. Pa- 
leontol. Mineral., Joint Conf., Sacramento Mtns., Otero 
Co., Guideb., pp. 154-171, 2 figs. 

1965. Early Paleozoic of New Mexico. In Guidebook of South- 
western New Mexico, II. New Mexico Geol. Soc., 16th 
Field Conf., New Mexico Bur. Mines and Mineral Res., 
pp. 112-131. 

1969. Early Paleozoic of New Mexico and the El Paso region, 
in The Ordovician symposium. El Paso Geol. Soc. 3rd 
Ann. Field Trip, 1969. El Paso, Texas, pp. 32-101. 

1970. Early Paleozoic of New Mexico and the El Paso Region. 
New Mexico State Bur. Mines and Mineral Res., Spec. 
Reprint, 44 pp., 5 figs. [Reprinted with minor corrections 
from El Paso Geol. Soc., 3rd Ann. Field Trip Guideb. 
(1969), pp. 32-101]. 

Frech, F. 

1897. Lethaea geognostica oder Beschreibung und Abbildung 
fiir die Gebirgs-Formationen bezeichnendsten Versteine- 
rungen. | Theil, Lethaea palaeozoica. Bd. 2, 256 pp., 35 
pls., 6 maps, 55 figs. 

Fredericks, G. N. 

1912. Bemerkungen tiber einige oberpalaeozoische Fossilien von 
Krasnoufimsk. Kazan Univ., Nat. Hist. Soc., Protoc. 
Sess., vol. 269, pp. I-9, | pl. 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DuTRO 147 


Fredericks, G. N. (continued) 

1916. Uber Einige Oberpaldozoic Brachiopoden von Eurasien. 
Comite Geol. Snt. Petersburg, Mem. 156, 87 pp., 5 pls., 
24 text-figs. 

1918. [Concerning the application of the subdivision of the api- 
cal apparatus to brachiopod classification). Russkoe Pa- 
leontol. Obshchestvo, vol. 2, (1917), pp. 85-91 [In Rus- 
sian]. 

1924. O Verkhne-Kammennougol nykh Spiriferidakh Urala. Geol. 
Komit., Izv., vol. 38, No. 2, pp. 295-324, 7 text-figs. [On 
Upper Carboniferous Spiriferids from the Urals. | 

Fritz, M. A. 

1944. Upper Devonian Bryozoa from New Mexico. J. Paleon- 

tol., vol. 18, No. 1, pp. 31-41, pls. 12, 13. 
George, T. N. 

1931. Ambocoelia Hall and Certain Other British Spiriferidae. 
Geol. Soc. London, Q. J., vol. 87, pp. 30-61, pls. 3-5, 3 
text-figs. 

Gill, T. 

1871. Arrangements of the Families of Mollusks. Smithson. Misc. 

Coll., vol. 227, 49 pp. 
Girty, G. H. 

1898. Description of a Fauna found in the Black Shale of East- 
ern Kentucky. Am. J. Sci., 4th Ser., vol. 6, pp. 384-394. 

1900. Devonian Fossils from Southwestern Colorado. The Fau- 
na of the Ouray Limestone. U.S. Geol. Surv., 20th Ann. 
Rep., pt. 2, pp. 31-36, pls. 3-7. 

1904. New Molluscan genera from the Carboniferous. U.S. Nat. 
Mus., Proc., vol. 27, pp. 721-736, pls. 16-18. 

1939. Some Linguloid Shells from the Late Devonian and Early 
Carboniferous Rocks of Pennsylvania and Ohio. U.S. Geol. 
Surv. Prof. Paper 193C, pp. 47-67, pl. 9. 

Goldfuss, A. 

1826-1833. Petrefacta Germaniae. Erste Theil. Diisseldorf. pp. 

1-252, pls. 1-61. 
Gordon, C. H. 

1907. Mississippian (Lower Carboniferous) Formations in the 
Rio Grande Valley, New Mexico. Am. J. Sci., 4th Ser., 
vol. 24, pp. 58-64. 

Grabau, A. W. 

1923-1924. Stratigraphy of China, Part 1. Palaeozoic and Old- 
er. China Geol. Surv., 528 pp., 306 text-figs., 6 pls. 

1931-1933. Devonian Brachiopoda of China, pt. 1. Palaeontol. 
Sinica, Peking, (B ili) vol. 3, 545 pp., 67 text-figs., 55 pls. 

Gratsianova, R. T. 

1974. [“Schuchertellas’’ of the Early and Middle Devonian in 
the south of western Siberia.| In Sreda i Zhizn v Geolog. 
Proshlom (Paleoekologicheskie Problema), pp. 77-87, No- 
vosibirsk. [In Russian]. 

Gray, J. E. 

1840. Synopsis of the Contents of the British Museum, 42nd ed., 
London, 370 pp. 

1848. On the Arrangement of the Brachiopoda. Ann. Mag. Nat. 
Hist., Ser. 2, vol. 2, pp. 435-440. 

Greiner, H. 

1957. Spirifer disjunctus: /ts Evolution and Paleoecology in the 
Catskill Delta. Yale Univ., Peabody Mus. Nat. Hist., Bull., 
vol. 11, 75 pp., 13 pls., 5 figs. 

Gurich, G. 

1896. Das Palaeozoicum des Polnischen Mittelgebirges. Russ.- 
Kais. Mineral. Gesellsch. Verhandl., ser. 2, vol. 32, 539 
pp.. pls. 1-15. 


Hall, J. 

1843. Geology of New York. Part IV. Comprising the geology 
of the Fourth Geological District. pp. i-xxii, 1-683, 74 
figs., 19 pls., map. 

1850. Description of New Species of Fossils and Observations 
upon Some Other Species Previously not well known, from 
the Trenton Limestone. New York St. Cab. Nat. Hist., 
Ann. Rep. vol. 3, pp. 173-183. 

1852. Geology and Paleontology. In Stansbury, H., Exploration 
and survey of the Great Salt Lake of Utah, including a 
reconnaissance of a new route through the Rocky Moun- 
tains. U.S. 32nd Cong. Spec. Sess., Senate Exec. Doc. 
3, pp. 399-414. 

1856. Descriptions and notices of the fossils collected upon the 
route [Whipple's reconnaissance near the thirty-fifth par- 
allel}. U.S. Pacific R. R. Explor. (U.S. 33rd Cong. Sess., 
Senate Exec. Doc. 78, and House Exec. Doc. 91) vol. 3, 
pt. 4, pp. 99-105. 

1857. Descriptions of Paleozoic Fossils. New York St. Cab. Nat. 
Hist., Ann. Rep., vol. 10, pp. 39-180. 

1858. In Hall, J. and Whitney, J. D. Report on the Geological 
Survey of the State of lowa, Embracing the Results of 
Investigations made during . . . 1855, 1856 and 1857. vol. 
1, pt. 1, Geology: xv + 472 pp., pt. 2, Palaeontology: pp. 
473-724, 30 pls. 

1860a. Observations of Brachiopoda. New York St. Cab. Nat. 
Hist., Ann. Rep., vol. 13, pp. 65-75. 

1860b. Descriptions of new species of fossils, from the Hamilton 
group of western New York, with notices of others from 
the same horizon in lowa and Indiana. New York St. Cab. 
Nat. Hist., Ann. Rep., vol. 13, pp. 76-94. 

1867. Descriptions and figures of the fossil Brachiopoda of the 
Upper Helderberg, Hamilton, Portage, and Chemung 
groups. New York Geol. Surv., Palaeontol., vol. 4, pt. 1, 
428 pp., 63 pls. 

Hall, J., and Clarke, J. M. 

1892-1895. An Introduction to the study of the Genera of Pa- 
laeozoic Brachiopoda. Nat. Hist. New York, Palaeontol., 
vol. 8, pt. 1: 367 pp., 20 pls., 37 text-figs., (1892): vol. 8, 
pt. 2: 217 pp., (1893); pp. 319-394, pls. 21-84, (1895). 

1894. An Introduction to the Study of the Brachiopoda. 13th 
Ann. Rep. New York St. Geol. (1893), Palaeontol., vol. 
2, pp. 751-943, pls. 23-54, text-figs. 287-669. 

Hall, J., and Whitfield, R. P. 

1872. Description of New Species of Fossils from the Vicinity 
of Louisville, Ky., and the Falls of the Ohio. New York 
St. Mus., Ann. Rep., vol. 24, pp. 181-200. 

1873. Descriptions of New Species of Fossils from the Devonian 
Rocks of lowa. New York St. Cab. Nat. Hist., Ann. Rep., 
vol. 23, pp. 223-243, pls. 9-12. 

1877. Paleontology. U.S. Geol. Expl. 40th Parallel (King), vol. 
4, pp. 197-302. in Annual report, geological exploration 
of the fortieth parallel from the Sierra Nevada to the east- 
ern Slope of the Rocky Mountains. U.S. [War Dept.] Chief 
Eng. Ann. Rep. 1877 (U.S. Cong. 2d sess. House Exec. 
Doc. 1, pt. 2, vol. 2, pt. 2) App. MM: 1207. 

Hass, W. H. 

1959. Conodonts from the Chappel Limestone of Texas. U.S. 

Geol. Surv., Prof. Paper 294-J, pp. 365-399, pls. 46-50. 
Havlicek, V. 

1967. Brachiopoda of the Suborder Strophomenidina in Czech- 
oslovakia. Ustréd. ustavo geol., Rozpr. Suazek 33, 235 
pp.. 52 pls., 89 text-figs. 


148 BULLETIN 315 


Haynes, W. P. 

1916. The fauna of the Upper Devonian in Montana. Part 2. 
The stratigraphy and Brachiopoda. Carnegie Mus., Ann., 
vol. 10, pp. 13-54, pls. 3-8. 

Helms, J. 

1963. Zur ‘‘Phylogenese’’ und Taxonomie von Palmatolepis 
(Conodontida, Oberdevon). Geologie, vol. 12, pp. 449- 
485, 4 pls. 

Hinde, G. J. 

1879. On conodonts from the Chazy and Cincinnati Groups of 
the Cambro-Silurian, and from the Hamilton and Gene- 
see-Shale divisions of the Devonian, in Canada and the 
United States. Geol. Soc. London, Q. J., vol. 35, pp. 351- 
369, pls. 15-17. 

Hoek, H. von 

1912. Versteinerungen des Cambriums und Silurs in des Hoch- 
landes von Bolivia und ihre Fauna. Neues Jahrb. Geol. 
Palaontol., vol. 6, pp. 305-484, pls. 10-14. 

Holmes, G. B. 

1928. A bibliography of the conodonts with descriptions of early 
Mississippian species. U.S. Nat. Museum, Proc., vol. 72, 
art. 5, No. 2701, pp. 1-38, pls. 1-11. 

Hoover, P. R. 

1981. Paleontology, Taphonomy and Paleoecology of the Pal- 
marito Formation (Permian of Venezuela). Bull. Am. Pa- 
leontol., vol. 80, No. 313, 138 pp., 10 pls., 11 text-figs., 
10 tables. 

House, M. R. 

1962. Observations on the Ammonoid Successions of the North 
American Devonian. J. Paleontol., vol. 36, No. 2, pp. 247— 
284, pls. 43-48, 15 figs. 

Huddle, J. W. 

1934. Conodonts from the New Albany Shale of Indiana. Bull. 

Am. Paleontol., vol. 21, No. 72, pp. 1-136, 12 pls. 
Huxley, T. H. 

1869. An Introduction to the Classification of Animals. London. 

147 pp., 47 text-figs. 
Hyatt, A. 

1884. Genera of fossil cephalopods. Boston Soc. Nat. Hist., 

Proc., vol. 22, pp. 273-338. 
Imbrie, J. 

1959. Brachiopods of the Traverse Group (Devonian) of Mich- 
igan. Am. Mus. Nat. Hist., Bull., vol. 116, No. 4, pp. 
349-409, pls. 49-67. 

Ivanova, E. A. 

1959. K sistematike i evoliutsii spiriferid (Brachiopoda). Akad. 
Nauk SSSR, vol. 4, pp. 47-63, 2 pls., 9 figs. [The System- 
atics and Evolution of Spiriferid Brachiopoda]. 

Jicha, H. L., Jr. 

1954. Geology and Mineral Deposits of Lake Valley Quadran- 
gle, Grant, Luna, and Sierra Counties, New Mexico. New 
Mexico St. Bur. Mines Mineral Res., Bull., vol. 37, 93 
pp., 5 pls., 13 figs. 

Johnson, J. G. 

1966. Middle Devonian brachiopods from the Roberts Moun- 
tains, Central Nevada. Palaeontology, vol. 9, pp. 152- 
181, 6 pls., 4 figs. 

1970. Taghanic Onlap and the end of North American Provin- 
ciality. Geol. Soc. Am., Bull., vol. 81, pp. 2077-2106, 4 
pls., 6 figs. 

1978. Devonian, Givetian age brachiopods and biostratigraphy, 
central Nevada. Geol. et Palaeontol., vol. 12, pp. 117— 


150, 8 pls., 6 figs., 3 tables 


Johnson, J. G. and Norris, A. W. 

1972. Tecnocyrtina, a new genus of Devonian brachiopods. J. 
Paleontol., vol. 46, No. 4, pp. 565-572, 2 pls., 3 text-figs. 

Johnson, J. G., Reso, A., and Stephens, M. 

1969. Late Upper Devonian Brachiopods from the West Range 
Limestone of Nevada. J. Paleontol., vol. 43, No. 6, pp. 
1351-1368, pls. 157-160, 7 text-figs. 

Kayser, E. 

1871. Die Brachiopoden des Mittel- und Oberdevon der Eifel. 
Zeitschr. deutsch, geol. Gesellsch., Bd. 23, pp. 491-647, 
pls. 8-14. 

Kelly, V. C., and Silver, C. 

1952. Geology of the Caballo Mountains. New Mexico Univ. 
Publ., Geol. Ser., vol. 4, 286 pp., 19 pls., 26 figs. 

Keyes, C. R. 

1908. Genesis of the Lake Valley, New Mexico, silver deposits. 
Am. Inst. Mining Eng., Bull., vol. 19, pp. 1-31. 

Kindle, E. M. 

1909. The Devonian Fauna of the Ouray Limestone. U.S. Geol. 
Surv., Bull., vol. 391, 60 pp., 10 pls. 

King, R. H. 

1938. New Chonetidae and Productidae from Pennsylvanian and 
Permian strata of North Central Texas. J. Paleontol., vol. 
12, No. 3, pp. 257-279, pls. 36-39. 

King, W. 

1846. Remarks on certain Genera belonging to the Class Pal- 
liobranchiata. Ann. Mag. Nat. Hist., vol. 18, pp. 26-42, 
83-94. 

1850. A Monograph of the Permian Fossils of England. London. 
Palaeontogr. Soc., xxxvii + 258 pp., 29 pls. 

1859. On Gwynia, Dielasma and Macandrevia, three new Gen- 
era, etc. Dublin Univ. Zool. Botan. Assoc., Proc., vol. 1, 
No. 3, pp. 256-262. 

Klapper, G., Sandberg, C. A., Collinson, C., Huddle, J. W., Orr, 
R. W., Rickard, L. V., Schumacher, D., Seddon, G., and Uyeno, 
Is Us 
1971. North American Devonian Conodont Biostratigraphy (pp. 

285-316] in Symposium on Conodont Biostratigraphy [W. 
C. Sweet and S. M. Bergstrém, eds.]. Geol. Soc. Am., 
Mem. 127, 499 pp. 


Koken, E. F. R. 
1889. Ueber die Entwickelung der Gastropoden vom Cambrium 
bis zur Trias. Neues Jahrb. Mineral., Geol. Palaontol., 
vol. 6, pp. 305—484, pls. 10-14. 


Kottlowski, F. E., Flower, R. H., Thompson, M. L., and Foster, 
R. W. 
1956. Stratigraphic Studies of the San Andres Mountains, New 
Mexico. New Mexico St. Bur. Mines Mineral Res., Mem. 
1, 132 pp., 5 pls., 15 figs. 
Kozlowski, R. 
1929. Les Brachiopodes de la Podolie Polonaise. Palaeontol. 
Pol., vol. 1, xiii + 254 pp., 12 pls. 
Kuhn, O. 
1949. Lehrbuch der Paldozoologie. Stuttgart. 326 pp., 244 figs. 
Lamarck, J. B. P. A. de M. de 
1801. Systeme des animaux sans vertébres. (Paris). viii + 432 
pp. 
Lane, N. G. 
1963. A. silicified Morrowan brachiopod fauna from the Bird 
Spring Formation, southern Nevada. J. Paleontol., vol. 
37, No. 2, pp. 379-392, 3 pls. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DutTRO 149 


Laudon, L. R., and Bowsher, A. L. 

1941. Mississippian Formations of the Sacramento Mountains, 
New Mexico. Am. Assoc. Petrol. Geol., Bull., vol. 25, 
No. 5, pp. 2107-2160, 31 figs. 

1949. Mississippian Formations of Southwestern New Mexico. 
Geol. Soc. Am., Bull., vol. 60, No. 1, pp. 1-87, 44 figs. 

Leveille, C. 

1835. Apercu géologique de quelques localités tres riches en 
coquilles sur les frontiéres de France et de Belgique. Géol. 
Soc. France, Mem., vol. 2, pp. 29-40, pl. 2. 

Licharew, B. K. 

1956. Brachiopoda in Kiparisova, L. D., Markovsky, B. P., and 
Radchenko, G. P. Materialy po paleontologii novye se- 
meystva i rody; Vses. Nauchno-Issled. Geol. Inst. Mater., 
new ser., vol. 12, Paleontol., 267 pp. 

Ludvigson, R., and Perry, D. G. 

1975. The Brachiopod Warrenella in the Lower and Middle De- 
vonian Formations of Northwestern Canada. Geol. Surv. 
Canada, Bull., vol. 235, pp. 59-106, 22 text-figs., pls. 12- 
19. 

Lyashenko, A. I. 

1957. Novyy rod Devonski Brakhiopod Uchtospirifer. [A new 
Genus of Devonian Brachiopods: Uchtospirifer] Akad. 
Nauk SSSR, Dokl., vol. 117, No. 5, pp. 885-888, 1 pl. 

McCammon, H. 

1960. Fauna of the Manitoba Group in Manitoba. Prov. Mani- 

toba, Dep. Mines Nat. Res., Publ. 59-6, 80 pp., 11 pls. 
M’Coy, F. 

1844. A Synopsis of the Characters of the Carboniferous Lime- 

stone Fossils of Ireland. Dublin. viii + 207 pp., 29 pls. 
McLaren, D. J. 

1961. Three new Genera of Givetian and Frasnian (Devonian) 
Rhynchonelloid Brachiopods. Inst. R. Sci. Nat. Belgique, 
Bull., vol. 37, No. 23, 7 pp., 2 pls. 

1962. Middle and Upper Devonian Rhynchonelloid Brachiopods 
from Western Canada. Geol. Surv. Canada, Bull. 86, x + 
122 pp., 18 pls., 1 chart. 

1965. Jn Schmidt and McLaren, Paleozoic Rhynchonellacea, pp. 
H552-H597, figs. 419-477A, in Williams er al., Treatise 
on Invertebrate Paleontology, pt. H. Brachiopoda, Geol. 
Soc. Amer., Univ. Kansas Press, xxxii + 927 pp., 746 
figs. 

McLaren, D. J., and Norris, A. W. 

1962. Illustrations of Fossils. Devonian of Western Canada. In- 
vertebrates. Geol. Surv. Canada, Paper 62-4, 33 pp., 15 
pls. 

MeNair, A. H. 

1937. Cryptostomatous Bryozoa from the Middle Devonian Tra- 
verse Group of Michigan. Michigan Univ. Mus. Paleon- 
tol., Contrib., vol. 5, No. 9, pp. 103-170, 15 pls. 

Maillieux, E. 

1939. La faune des schistes de Barvaux-sur-Ourthe (Frasnien 
Supérieur). Mus. Roy. d’Hist. Nat. de Belgique, Bull. 15, 
No. 53, pp. I-8. 

Marcou, J. 

1858. Geology of North America; with two reports on the prai- 
ries of Arkansas and Texas, the Rocky Mountains of New 
Mexico, and the Sierra Nevada of California. Zurich. 144 
pp., 7 pls. 

Martin, W. 

1809. Petrifacta Derbiensia; or figures and descriptions of petri- 

factions collected in Derbyshire. London. 28 pp., 52 pls. 
Massalongo, A. 

1855. Monografia delle nereide fossili del M{onte| Bolca. Ve- 

rona. 55 pp., 6 pls. 


Meek, F. B. 

1860. Descriptions of new Fossil Remains collected in Nebraska 
and Utah, by the Exploring Expeditions under the com- 
mand of Capt. J. H. Simpson. Acad. Nat. Sci., Philadel- 
phia, Proc. for 1860, pp. 308-315. 

1867 [1868]. Remarks on the Geology of the Valley of the Mac- 
Kenzie River, with figures and descriptions of fossils from 
that region, in the Museum of the Smithsonian Institution, 
chiefly collected by the late Robert Kennicott. Chicago 
Acad. Sci., Trans., vol. 1, pp. 61-114. 

1876. Report on the Paleontological Collections of the Expedi- 
tion. In Simpson, J. H. Report of Explorations across the 
Great Basin of the Territory of Utah in 1859. pp. 337-373, 
illus. 

1877. Paleontology. U.S. Geol. Explor. 40th Par. (King), vol. 
4, 197 pp., illus. 

Menke, C. T. 

1828. Synopsis methodica Molluscorum generum omnium et 
specierum earum quae in Museo Menkeano adservantur. 
Pyrmonti. xii + 91 pp. 

Merriam, C. W. 

1940. Devonian Stratigraphy and Paleontology of the Roberts 
Mountains Region, Nevada. Geol. Soc. Am., Spec. Pa- 
per, No. 25, 114 pp., 16 pls., 8 tables, map. 

Mikryukoy, M. F. 

1968. A new Genus and Species of Chonetidae from the Russian 
Platform. In Markovskii, B. P. [ed.]. Novye vidy drevnikh 
rastenii | bespozvonochnykh. Vipusk, chast II, p. 90-92. 

Miller, A. K. and Collison, C. 

1951. A Clymenoid Ammonoid from New Mexico. Am. J. Sci., 

vol. 249, No. 8, pp. 600-603, 1 fig. 
Miller, A. K., and Youngquist, W. 

1947. Conodonts from the type section of the Sweetland Creek 
shale in Iowa. J. Paleontol., vol. 21, No. 6, pp. 501-517, 
pls. 72-75. 

Miller, S. A. 

1883. Catalogue of American Paleozoic Fossils. 2nd Edition. 
298 pp. 

Milne-Edwards, H., and Haime, J. 

1850. A monograph of the British fossil corals. Palaeontogr. Soc. 
Monogr., pp. i-Ixxxv, 1-71, pls. I-11. 

Moore, R. C. 

1952. In Moore, R. C., Lalicker, C. G., and Fischer, A. G. 

Invertebrate Fossils. McGraw-Hill, New York. 776 pp. 
Muir-Wood, H. M. 

1955. A History of the Classification of the Phylum Brachiop- 
oda. Brit. Mus. [Nat. Hist.]. London. viii + 24 pp., 12 
figs. 

1962. On the Morphology and Classification of the Brachiopod 
Suborder Chonetoidea. Brit. Mus. [Nat. Hist.]. London. 
132 pp., 16 pls., 24 text-figs. 

Muir-Wood, H. M., and Cooper, G. A. 

1960. Morphology, Classification and Life Habits of the Pro- 
ductoidea (Brachiopoda). Geol. Soc. Am., Mem. 81. xii + 
447 pp., 135 pls., 8 text-figs. 

Muller, K. J. 

1956. Die Gattung Palmatolepis (Zur Kenntniss der Conodon- 
ten-Fauna des europdischen Devons, 1.). Senckenb. na- 
turf. Ges., Abh., Bd. 494, pp. 1-70, 11 pls. 

Nalivkin, D. V. 

1918. Devonian Fauna of Turkestan. Mem. Com. Géol., Len- 
ingrad, n. s., vol. 180, xvi + 221 pp., 10 pls. 

1947. Devonien. Atlas of the Guide Forms of the fossil fauna of 
the USSR, vol. 3, 245 pp., 56 pls. Moscow: Vses. Nauch- 
no-Issled. Geol. Inst. (VSEGEI) Minist. Geol. SSSR [In 
Russian]. 


BULLETIN 315 


Nelson, L. A. 
1940. Paleozoic Stratigraphy of the Franklin Mountains of West 
Texas. Am. Assoc. Petrol. Geol., Bull., vol. 24, No. 1, 
pp. 157-172, 5 figs. 


North, F. J. 
1920. On Syringothyris Winchell and Certain Carboniferous 
Brachiopods referred to Spiriferina d’Orbigny. Geol. Soc. 
London, Q. J., vol. 76, No. 2, pp. 162-227, pls. 11-13, 6 
text-figs. 


Oehlert, D. P. 

1887. Jn Fischer, P. H.. Manuel de Paléontologie Conchylio- 
gique ou Histoire Naturelle des Mollusques Vivants et 
Fossiles. F. Savy, Paris. Pt. 2, pp. 1189-1334, pl. 15, text- 
figs. 892-1138. 

1890. Note sur Différents Groupes Etablis dans le Genre Orthis 
et en Particulier sur Rhipidomella Oehlert (Rhipidomys 
Oehlert olim). J. Conchyl., Ser. 3, vol. 30, pp. 366-374. 

Opik, A. 

1933. Uber einige Dalmanellacea aus Estland. Univ. Tartu. 
(Dorpat), Acta et Comment., ser. A, vol. 25, No. 1, pp. 
1-25, 6 pls. 

1934. Uber Klitamboniten. Univ. Tartu. (Dorpat), Acta et Com- 
ment., ser. A, vol. 26, No. 3, 239 pp., 48 pls, 55 text-figs. 

d’Orbigny, A. 

1847. Considérations Zoologiques et Géologiques sur les Bra- 
chiopodes ou Palliobranches. Acad. Sci. Paris, C. R., vol. 
25, pp. 193-195, 266-269. 

1850. Prodréme de paléontologie stratigraphique universelle. vol. 
1, (1849), 394 pp. 

Owen, D. D. 

1852. Report of a geological Survey of Wisconsin, Iowa and 
Minnesota and incidentally of a portion of Nebraska Ter- 
ritory. XXXviii + 638 pp., 27 pls., 16 sections, map. 

Paige, S. 

1916. Silver City Folio, New Mexico. Geol. Atlas U.S., No. 199, 

19 pp., 3 maps. 
Phillips, J. 

1836. Illustrations of the geology of Yorkshire: Pt. 2, the Moun- 
tain Limestone district. John Murray (London). 253 pp.. 
25 pls. 

1841. Figures and descriptions of the Palaeozoic fossils of Corn- 
wall, Devon and West Somerset. Geol. Sury. Gt. Brit., 
Mem., vol. 1, xii + 231 pp., 60 pls. 

Pitrat, C. W. 

1965. Spiriferidina [pp. H667—H728, figs. 542-593] in Williams 
et al., Treatise of Invertebrate Paleontology, pt. H. 
Brachiopoda, Geol. Soc. Am., Univ. Kansas Press, xxxii_ + 
927 pp., 746 figs. 

1975. Orthospirifer, New Genus of Spinocyrtiid Brachiopods. J. 
Paleontol., vol. 49, No. 2, pp. 387-394, 2 pls., 1 text-fig. 

Poole, F. G., Baars, D. L., Drewes, H., Hayes, P. T., Ketner, kK. B., 

Mckee, E. E., Teichert, C., and Williams, J. S. 

1967. Devonian of the Southwestern United States [vol. 1, pp. 
879-912] in International Symposium on the Devonian 
System. |D. H. Oswald, ed.] Calgary. [2 vols.] vol. I: xii + 
1055 pp., vol. I, xi + 1377 pp. 

Pray, L. C. 

1961. Geology of the Sacramento Mountains escarpment, Otero 
County, New Mexico. New Mexico St. Bur. Mines Min- 
eral Res., Bull., vol. 35, 144 pp., 3 pls., 34 figs. 

Raymond, P. E. 

1909. The fauna of the Upper Devonian in Montana. Part 1. 

The fossils of the red shales. Carnegie Mus., Ann., vol. 
pp. 141-158, pls. 3-8 


Raymond, P. E. (continued) 

1911. The Brachiopoda and Ostracoda of the Chazy. Carnegie 

Mus., Ann., vol. 7, pp. 215-259, pls. 33-36. 
Roberts, J. 

1971. Devonian and Carboniferous brachiopods from the Bo- 
naparte Gulf Basin, Northwestern Australia. vol. 1 (Text): 
ix + 284 pp., 87 figs: Appendix, pp. 285-319, 14 figs.; vol. 
2: 59 pls., index pp. i-xiil. 

Roemer, C. F. 

1844. Das Rheinische Uebergangsgebirge. Eine Paldontolo- 

gisch-geognostische Darstellung. Hanover, 96 pp., 6 pls. 
Rzhonsnitskaya, M. A. 

1956. New Families and Genera. In Kiparisova, Markovski and 
Radchenko. Materials on Paleontology. Vses. Nauchno- 
Issled. Inst. Mater., n. ser., Trudy, Vipusk 12, 267 pp., 
43 pls., 53 figs. 

1959. K Sistematike Rinchonellid. Paleontol. Zh., vol. 1, pp. 25— 
35. [On the Systematics of the Rhynchonellids.} 

Sanneman, D. 

1955. Beitrag zur Untergliederung des Oberdevons nach Con- 
odonten. N. Jahrb. Geol. Palaontol., Abh. 100, pp. 324— 
331, 1 pl. 

Sartenaer, P. 

196la. Late Upper Devonian (Famennian) Rhynchonelloid Bra- 
chiopods. Inst. R. Sci. Nat. Belgique, Bull., vol. 37, No. 
24, pp. 1-10, 2 pls. 

1961b. Etude Nouvelle en Deux Parties du Genre: Camarotoechia 
Hall and Clarke, 1893. Premiére Partie: Atrypa congregata 
Conrad, Espéce-Type (1). Inst. R. Sci. Nat. Belgique, Bull., 
vol. 37, No. 22, pp. I-11, | pl. 

1965. Trois Nouveaux Genres de Brachiopodes Rhynchonellides 
du Famennien. Inst. R. Sci. Belgique, Bull., vol. 41, No. 
3, pp. 1-12, 2 pls. 

1966a. Frasnian Rhynchonellida from East Iran. Partie 2. Fras- 
nian Rhynchonellida from the Ozbak-Kuh and Tabas Re- 
gions (East Iran). Geol. Surv. Iran, Rep., No. 6, pp. 25— 
53, 2 pls., 2 figs. 

1966b. Ripidiorhynchus, Nouveau Genre de Brachiopode Rhyn- 
chonellide du Frasnien. Inst. R. Sci. Nat. Belgique, Bull., 
vol. 42, No. 30, pp. 1-13, 2 pls. 

1969. Late Upper Devonian (Famennian) Rhynchonellid Bra- 
chiopods from Western Canada. Geol. Surv. Canada, Bull. 
169, 269 pp., 19 pls., 41 text-figs. 

1970. Nouveaux Genres Rhynchonellides (Brachiopodes) du Pa- 
léozoique. Inst. R. Sci. Nat. Belgique, Bull., vol. 46, 32 
pp- 

Savage, N. M., Eberlein, G. D., and Churkin, M., Jr. 

1978. Upper Devonian Brachiopods from the Port Refugio For- 
mation, Suemez Island, Southeastern Alaska. J. Paleon- 
tol., vol. 52, No. 2, pp. 370-393, pls. 1, 2, 24 text-figs. 


Schindewolf, O. E. 
1923. Entwurf einer natiirlichen Systematik der Clymenoidea. 
Zentralbl. Mineral. Geol. Paléont., Jahrg. 1923, pp. 23- 
30, 59-64. 
Schlotheim, E. F. von 
1820. Die petrefactenkunde auf ihrem jetzigen Standpunkte durch 
die Beschreibung einer Sammlung versteinerte und fos- 
siler Uberreste der Tier- und Pflanzenreichs der Vorwelt 
erldutert. Gotha. vol. 1, lxii + 378 pp. 
Schmidt, H. 
1964. Neue Gattungen palaeozoischer Rhynchonellacea 
(Brachiopoda). Senckenberg. Leth., vol. 45, No. 6, pp. 
505-506. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 151 


Schuchert, C. 

1896. Brachiopoda. in Zittel, K. A. von [translated and edited 
by C. R. Eastman]. Textbook of Paleontology, First Edi- 
tion 1. Macmillan and Company, Limited, London. pp. 
291-343, text-figs. 489-587. 

1913. Brachiopoda. in Zittel, K. A. von [translated and edited 
by C. R. Eastman] Textbook of Paleontology, Second Edi- 
tion 1. Macmillan and Company, Limited, London. pp. 
355-420, text-figs. 526-636. 

1929. In Schuchert and Levene, C. M.* Brachiopoda. Fossilium 
Catalogus 1: Animalia, Pars 42. W. Junk, Berlin. 140 pp. 
Schuchert, C. and Cooper, G. A. 

1931. Synopsis of the Brachiopod Genera of the Suborders Or- 
thoidea and Pentameroidea, with notes on the Telotre- 
mata. Am. J. Sci., ser. 5, vol. 22, pp. 241-251. 

1932. Brachiopod Genera of the Suborders Orthoidea and Pen- 
tameroidea. Yale Univ., Peabody Mus. Nat. Hist., Mem. 
4, No. 1, xii + 270 pp., 29 pls., 36 text-figs. 


Sidiachenko, A. I. 
1961. [The Upper Devonian Cyrtospiriferid subgenus Dmitria.] 
Paleontolog. Zhurnal, 1961, No. 2, pp. 80-85, | pl., | fig. 
{In Russian]. 


Sokolskaya, A. N. 

1941. [Lower Carboniferous and Devonian-Carboniferous bra- 
chiopods of the Moscow Basin (Tschernysshino, Upa and 
Malevka-Muraevnya Beds).| Akad. Nauk, Paleontol. Inst., 
Trudy, vol. 12, Pt. 2, pp. 1-138, 12 pls., 39 figs. [In Rus- 
sian]. 

1960. Otryad Strophomenida in Mshanki Brakhiopody, Sary- 
cheva, T. G. [ed.], in Osnovy Paleontologii, Orlov, Yu A. 
{ed.], pp. 206-220, pls. 28-32, text-figs. 138-193. Moskva. 
{in Russian] 


Stainbrook, M. A. 

1935. A Devonian Fauna from the Sacramento Mountains, New 
Mexico. J. Paleontol., vol. 9, No. 8, pp. 709-714, plate 
83. 

1938. Pentameridae of the Cedar Valley Beds of lowa. Am. Midl. 
Nat., vol. 19, No. 3, pp. 723-738, 2 pls., 8 text-figs. 

1940. Orthoid Brachiopods of the Cedar Valley Limestone of 
Iowa. Am. Midl. Nat., vol. 23, pp. 482-492, 2 pls. 

1943. Strophomenacea of the Cedar Valley Limestone of Iowa. 
J. Paleontol., vol. 17, No. 1, pp. 39-59, pls. 6, 7. 

1945. Brachiopoda of the Independence Shale of lowa. Geol. 
Soc. Am., Mem. 14, vi + 74 pp., 6 pls. 

1947. Brachiopoda of the Percha Shale of New Mexico and Ar- 
izona. J. Paleontol., vol. 21, No. 4, pp. 297-328, pls. 44— 
47. 

1948. Age and Correlation of the Devonian Sly Gap Beds near 
Alamogordo, New Mexico. Am. J. Sci., vol. 246, pp. 765— 
790, 2 pls. 

1950. Brachiopoda and Stratigraphy of the Aplington Forma- 
tion of Northern Iowa. J. Paleontol., vol. 24, No. 3, pp. 
365-385, pls. 53, 54. 

1951. Substitution for the Preoccupied Brachiopod Name Hys- 
tricina. Washington Acad. Sci., J., vol. 41, No. 6, p. 196. 


* Schuchert alone is responsible for the classification proposed in 
this paper. He says (page 4). “‘The senior author presents a new 
classification (p. 10) built upon the older ones, with the purpose of 
facilitating the work of other authors in finding closely related gen- 
era and thus avoiding the making of synonyms.”’ [Italics are ours.] 


Stauffer, C. R. 

1938. Conodonts of the Olentangy shale. J. Paleontol., vol. 12, 
No. 5, pp. 411-443, pls. 48-53. 

1940. Conodonts from the Devonian and associated clays of 
Minnesota. J. Paleontol., vol. 14, pp. 417-435, pls. 58-60. 

Stehli, F. G. 

1961. New Genera of Upper Paleozoic Terebratuloids. J. Pa- 

leontol., vol. 35, No. 3, pp. 457-466, pl. 62, 8 text-figs. 
Stevenson, F. V. 

1941a. Denovian Formations in New Mexico. Illinois Acad. 
Sci., Trans., vol. 34, No. 2, p. 163. 

1941b. The Devonian Sly Gap Formation of New Mexico 
{Abstr.]. Oil and Gas J., vol. 39, No. 47, p. 65. 

1943. Onondagan Equivalent in New Mexico. Am. Assoc. Pe- 
trol. Geol., Bull., vol. 27, No. 2, pp. 222-223. 

1945. The Devonian of New Mexico. J. Geol., vol. 53, pp. 217- 
245, 13 figs. 

Stewart, G. A. 

1927. Fauna of the Silica Shale of Lucas County [Ohie]. Geol. 

Surv. Ohio, 4th ser., Bull. 32, 76 pp., 5 pls. 
Swallow, G. C. 

1860. Descriptions of new fossils from the Carboniferous and 
Devonian rocks of Missouri. Acad. Sci. St. Louis, Trans., 
vol. 1, pp. 635-639. 

Szulczewski, M. 

1971. Upper Devonian conodonts, stratigraphy and facial de- 
velopment in the Holy Cross Mts. Acta Geol. Polonica, 
vol. 21, 129 pp., 34 pls., 10 figs. 

Teichert, C. 

1965. Devonian Rocks and Paleogeography of Central Arizona. 

U.S. Geol. Surv., Prof. Paper 464, 181 pp. 
Termier, H., and Termier, G. 

1949. Essai sur l'Evolution des Spiriféridés. Notes et Mem. Serv. 
Mines et Carte Geol. Maroc, vol. 74, pp. 85-112, 12 text- 
figs. 

Thomas, A. O., and Stainbrook, M. A. 

1922. Status of certain rhynchonellid brachiopods from the De- 
vonian of Iowa. lowa Acad. Sci., Proc., vol. 29, pp. 93— 
98, | pl. 

Thomas, I. 

1910. British Carboniferous Orthotetinae. Gt. Brit. Geol. Surv., 
Mem., vol. 1, pt. 2, pp. 83-134, pl. 13. 

1914. The British Carboniferous Producti. 1. Gt. Brit. Geol. Surv. 
(Palaeontol.), Mem., vol. 1, pt. 4, pp. 197-366, pls. 17— 
20. 

Thomas, L. A. 

1949. Devonian-Mississippian formations of southeast Iowa. 
Geol. Soc. Am., Bull., vol. 60, No. 3, pp. 403-438, pls. 
1-4. 

Tien, C. C. 

1938. Devonian Brachiopoda of Hunan. Palaeontol. Sinica, n. 

ser. B, vol. 4 [whole series 113], 192 pp., 22 pls. 
Ulrich, E. O. 

1882. American Paleozoic Bryozoa. Cincinnati Soc. Nat. Hist., 

vol. 5, pp. 121-175, 232-257. 
Ulrich, E. O., and Bassler, R. S. 

1926. A classification of toothlike fossils, conodonts, with de- 
scriptions of American Devonian and Mississippian 
species. U.S. Nat. Mus., Proc., vol. 68, art. 12, pp. 1-63, 
11 pls. 

Verneuil, E. de 

1845. Paléontologie, mollusques, brachiopodes. In Murchison, 
R. I., Verneuil, E. de, and Keyserling, A. de, Géologie 
de la Russie d'Europe, vol. 2, pt. 3, pp. 37-294, pls. 11, 
15-18. 


Waagen, W. H. 
1882-1885. Salt Range Fossils, Part 4. (2) Brachiopoda. Pa- 
laeontol. Indica, Mem., Ser. 13, vol. 1, pp. 329-770, pls. 
25-86 (fasc. 1, pp. 329-390, pls. 25-28, Dec. 1882; fasc. 
2, pp. 391-546, pls. 29-49, Aug. 1883; fasc. 3, pp. 547— 
610, pls. 50-57, May 1884; fasc. 4, pp. 611-728, pls. 58- 
81, Dec. 1884; fasc. 5, pp. 729-770, pls. 82-86, July 1885). 
Walcott, C. D. 
1884. Paleontology of the Eureka District, Nevada. U.S. Geol. 
Surv. Monogr. 8, xili + 298 pp., 24 pls. 
Warren, P. S. 
1928. The Paleozoics of the Crowsnest Pass, Alberta. R. Soc. 
Canada, Trans., sect. 4, vol. 22, pp. 109-119, 1 pl. 
1944. The Role of Sphaerospongia tesselata in the MacKenzie 
River Devonian. Canadian Field Nat., vol. 58, pp. 28-29, 
1 pl. 
Warren, P. S., and Stelck, C. R. 
1956. Reference Fossils of Canada. Part 1. Devonian Faunas 
of Western Canada. Geol. Assoc. Canada, Spec. Paper |, 
15 pp., 29 pls. 
Webster, C. L. 
1889a. A description of the Rockford Shales of Iowa. Davenport 
Acad. Sci., Proc., vol. 5, pp. 100-109. 
1889b. Description of a new genus of corals (MacGeea) from 
the Devonian Rocks of Iowa. Am. Nat., vol. 23, pp. 710- 
712. 
1921. Notes on the Genus Atrypa with Descriptions of New 
Species. Am. Midl. Nat., vol. 7, pp. 13-20. 
Wedekind, R. 
1926. Die devonische Formation. In W. Salomon, Grundziige 
der Geologie. vol. 2, pp. 194-226, 6 pls. Stuttgart. 
Weller, S. 
1910. Internal Characters of Some Mississippian Rhynchonel- 
liform Shells. Geol. Soc. Am., Bull., vol. 21, pp. 497-516, 
18 figs. 
1914. The Mississippian Brachiopoda of the Mississippi Valley 
Basin. Illinois Geol. Surv., Monogr. 1, 508 pp., 83 pls. 
Wells, J. W. 
1940. Sphaerospongia in the Tully Formation. J. Paleontol., vol. 
14, No. 5, pp. 502-504, 2 figs. 
Weyer, D. 
1967. Kitakamithyris Minato 195] (Brachiopoda, Spiriferida) aus 
dem Etroeungt (Oberdevon) und Tournai (Unterkarbon) 
des Rheinischen Schiefergebirges. Geol. [Berlin], vol. 16, 
No. 4, pp. 433-451, 2 pls., 3 text-figs. 
White, C. A. 
1862. Description of new species of fossils from the Devonian 
and Carboniferous rocks of the Mississippi Valley. Boston 
Soc. Nat. Hist., Proc., vol. 9, pp. 8-33. 


BULLETIN 315 


Williams, A. 

1953. North American and European Stropheodontids: their 
Morphology and Systematics. Geol. Soc. Am., Mem. 56, 
vi + 67 pp., 13 pls., 5 text-figs. 

Williams, A., Rowell, A. J., Muir-Wood, H. M., Pitrat, C. W., 
Schmidt, H., Stehli, F. G., Ager, D. V., Wright, A. D., Elliott, G. 
F., Amsden, T. W., Rudwick, M. J. S., Hatai, K., Biernat, G., 
McLaren, D. J., Boucot, A. J., Johnson, J. G., Staton, R. D., 
Grant, R. E., and Jope, H. M. 

1965. Treatise on Invertebrate Paleontology, Part H, Brachi- 
opoda [R. C. Moore, ed.], Geol. Soc. Am., Univ. Kansas 
Press [2 vols.], xxxii + 927 pp., 746 text-figs. 

Williams, H. S. 

1882. The recurrence of faunas in the Devonian rocks of New 
York. Am. Assoc., Proc., vol. 30, pp. 186-191. 

1890. The Cuboides Zone and its fauna; a discussion of methods 
of correlation. Geol. Soc. Am., Bull., vol. 1, pp. 481-500, 
pls: sl25M1B° 

1908. The Dalmanellas of the Chemung Formation and a closely 
Related new Brachiopod Genus Thiemella. U.S. Nat. Mus., 
Proc., vol. 34, No. 1596, pp. 35-64, 4 pls. 

Winchell, A. 

1866. The Grand Traverse region. A report on the geology and 
industrial resources of the counties of Antrim, Grand 
Traverse, Benzie, and Leelanaw in the Lower Peninsula 
of Michigan (including appendix). Ann Arbor. 97 pp., map. 

Woodward, S. P. 

1851-1856. A Manual of the Mollusca, or Rudimentary Treatise 
of Recent and Fossil Shells. Parts I-III. xvi + 486 pp. 

Zeller, R. A., Jr. 

1965. Stratigraphy of the Big Hatchet Mountains Area, New 
Mexico. New Mexico St. Bur. Mines Mineral Res., Mem. 
16, 128 pp., 6 pls., 18 figs. 

Ziegler, W. 

1962. Die Conodonten aus den Geréllen des Zechsteinkonglom- 
erates von Rossenray (stidwestlich Rheinberg/Nieder- 
rhein). Geol. Rheinl.-Westf., Fortschr., Bd. 6, pp. I-15, 
pls. 1-4. 

Ziegler, W., and Huddle, J. W. 

1969. Die Palmatolepis glabra-Gruppe (Conodonta) nach der 
Revision der Typen von Ulrich & Bassler durch J. W. 
Huddle. Geol. Rheinl.-Westf., Fortschr., Bd. 16, pp. 377— 
386. 

Ziegler, W., Klapper, G., and Johnson, J. G. 

1976. Redefinition and Subdivision of the varcus Conodont Zone 
(Middle—? Upper Devonian) in Europe and North Ameri- 
ca. Geol. et Palaeontol., vol. 10, pp. 109-132, pls. 1-4. 


PLATES 


PLATE A 


VOLUMES 82, 83 


AMERICAN PALEONTOLOGY, 


OF} 


loa) 
yy 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO 


‘tadooD “y ‘5 Aq o10Ug 


‘OOIXAT, MAN “ALIA ‘Soul, AARYORN ‘(1awWWeRY MOlaq IsNf) aUO]SAWT] aAISSeUL ‘UOTRWUIOY aIVUC * 


‘laysmog “JT “vy Aq o10yg 


“SUIRJUNOW OJUIWBIIVG ‘UOAURD Big ‘UONPWAO] aeUG * 


‘radood “y ‘D5 Aq o10ug 


‘suIR]UNOW Saipuy uRg ‘UoAURD Ikag ‘(1aWWRY Mojaq IsNf) do} UO pag Ysy YUM UONRWIOY aevUG - 


“Taysmog “J “Vy Aq o10yug 


‘SUIRJUNOP] OJUWRIOLG ‘youRIg UPUIpPRaq ‘UOAURD OWWPY ‘UOnRWIO4 a1eUC * 


UONeULIO a1vUQ 


V ALV Id AO NOILVNV 1dX4 


a 


BULLETIN 315 


Vaal 


‘tadoog ‘Vv ‘5 Aq 0104d 


“suIR]UNOY|, Saipuy ues ‘UoAURD URWIpREg ‘UPIUOAAG JO UONDSAS [NJ ~ 


‘radooa “y “9 Aq 0104d 


“suIRJUNO|, SdIpuy URS ‘UOAURD URWIprag ‘UONeWIO, deDH Ajg 1addy ~ 


‘tadoog ‘Vv ‘D5 Aq 0104d 


“‘OOIXa MON ‘vSOWIAaH ‘uONeULIO dey A[g Ie[NPON * 


-yadooa “y ‘5 Aq 01044 


“suIeJUNO, OJUSWIRIORS ‘UOAURD OWR]Y ‘UONRUIO, ded AIS Jo [IY [eoIUOD * 


uonewmioy dey AIS 


q divig 10 NOILVNV1dXq 


a 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE B 


PRATEae 


AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


rINS OF 


van 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 


‘radoog ‘Vy ‘Dd Aq o10ug 


“SUIRJUNO| Saipuy uPs ‘ureUNOW daayg ‘uoNeUIOY poosysno10y], * 


‘radoog “Vy ‘D5 Aq o10ug 
‘suleJUNOW Saipuy ues ‘uleyUNoW dsayg ‘uOnRWIOY O1ape}UOD 
‘radoo9 “y ‘5 Aq o10Ug 


“suIeJUNOW Soipuy uRg ‘ure}UNOW| daayg ‘uoNeWio4 dey A[g ay) Jo uoNsod Ayeys ° 


‘tadooD “y 'D Aq o10Ug 


‘suIeJUNO| SaiIpuy US ‘YyINOS ay} Woy UIeJUNO| dasys ° 


urejunoyw daayg ‘uetuoaag 


OJ ALVTd JO NOILVNV1dXq 


nm 


BULLETIN 315 


156 


‘radooa “y ‘9 Aq 104d 


“OOIXAP MAN ‘UMOJATIOADH ‘1aquiawi Jo yed 1aMO] Ul Sa[npou ‘(1aquay XOg) UONPUIO eYdIAg ~ 


‘tadooD “y ‘9 Aq o104d 


‘ODIXIP MAN ‘UMOIITIOADH 3e UONDdas | [NJ ‘(laquayy XOg) UONRWIO BYdIdg * 


‘tadoo9 “Vy *5 Aq o10ug 


OOIXAW MAN ‘AapIN ‘ureyunop auo7T ‘(auojsy[is Avg Apeoy) UoNneWIO eYydiog * 


iadooa “y ‘5 Aq 004g 


OOIXIW MAN ‘OLOQSTIIH JO IseayINos ‘(YLOS Ai|eoo]) UONDas adAq ‘(1aqWay XOG) UONeWAIOY eYyosag ° 


UONeULIO BYd.1ag 


d ALV1g 40 NOILVNV1dXq 


a 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE D 


ig A 


LLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


PLATE 


Figure 


10-21. 


23-26. 


27-47. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 157 


EXPLANATION OF PLATE | 


Page 


> LEAT En) Ges Se pan macemotts ob onio Cr moO any as Geel ear Scere cri chtartas oierctio 6 Aree nena et hoy che emo achts Cromer nian crs ree cloth 32 


1, Interior of a small, narrow individual, «6, figured specimen USNM 200818b; 2, Exterior of a smaller dorsal valve, <4, figured 
specimen USNM 20008 18a. 
Contadero Formation, locality 5240. 


PE ORDICUIOIMCAESPECIE SE co teretare came pesana tous ears VT RMR eae Gre cotta Se seats a teeee eee tna te SO ch ty cnsaney SVN oS fa ene ao acne rare OE anne! sa ARRON E Leet tetoed 33 


3, 4, Side and dorsal views of a dorsal valve, <1, figured specimen USNM 200817a; 5, Exterior of the same dorsal valve, <2; 6, 
View of the ventral side of a small specimen, showing pedicle opening, <3, figured specimen USNM 200817b. 
Contadero Formation (lower 5 ft), locality 5240. 


POP CUrOCKAaAMiasSPECLES ie as. < sin cycvevereie eyereloiccs ele as oe Spatevagagat erect choi ass, Sz) ces casayous tenes cues (onevavayevorey apace oicuavararagegersys) sr) zie er atsuene ever sweney eneteeenene er ciara 34 


Dorsal view of a specimen attached to a large Hadrorhynchia, x2, figured specimen USNM 201153. 
Onate Formation, locality 523d. 


MRetrocraniaispecitically unidentified yy. e:< leo save -vanctover stags sreehel 6 5 oi ay 31s fos 2s adeustoy vena yetctevers wis lo, v allepofalinis tolanedssevene sis) sic esi le el elegetevevere sreusce 34 


Side and dorsal views of a large depressed specimen, 1, figured specimen USNM 200819. 

Sly Gap Formation, locality 521Ic. 
Ii pid omel APS PECLESilew-reeyaes saps te ces eaa Stars “ov gave coz sc, sichs¥oh= cam atet atonal au cores senate ynie uate stele tera some trictske nie ceeretel kote net ter corer rata erene tert Torr 41 
10-14, Anterior, dorsal, side, posterior and ventral views of a complete specimen, x1, USNM 204950a; 15-19, Dorsal, ventral, 
posterior, side, and anterior views of another complete specimen, x1, USNM 204950b; 20, 21, Dorsal and ventral views of a 
third specimen, x1, USNM 201213. 

Sly Gap Formation (below Macgeea Webster, 1889), locality 528v. 


me EHIME CARS PE CLES eeecerekeretedere oe feet eiseeisee neyeyevernits ci So ucirarerenensieteian ane 151 ene PRPSYEOENES « tocvan‘x ae feceawia Pores elec cavers ates Stccdt che ane Posey ee ate RonereTenene eer eneye renee ears 34 


Dorsal view of a small, incomplete dorsal valve, <3, figured specimen USNM 203534. 

Sly Gap Formation (below Macgeea Webster, 1889; at the base of the formation), locality 506). 

Retrocraniajourayensis) (Kane) e555 sc. <z2 yave'tey ataceveieneisusie16) xeresc1 si elev enone she oheveterdiesslerievoren’ © ay avavenevevabater cleo: s/evele|« fovea </eveusbereleieueleier veversteis 34 
23, 24, Side and dorsal views of a specimen attached to Leioproductus, <1, hypotype USNM 200826a; 25, Same view and 
specimen, x2. 

Percha Formation (Box Member), locality 3026a. 

26, Dorsal view of another specimen, attached to Schizophoria australis Kindle, x2, hypotype USNM 200827. 

Percha Formation (Box Member), locality 506. 
CariniferellaralleniincwaSpeCieseme-rererereretcsterrtertrretrettoretere ete elobenedeterateer el eteteforek tated cteratarte tet tereciy-aetetistoielenere iarciereetr tte toet iterate 40 
27-31, Ventral, side, posterior, dorsal and anterior views, x1, holotype USNM 200831; 32, 33, Dorsal and ventral views, <2, of 
the holotype. 

Sly Gap Formation, locality 509p. 

34, 35, Interior of the dorsal valve, showing cardinal process and brachiophores, <1 and x2, paratype USNM 200837a; 36, 37, 
Interior of the ventral valve, showing bilobed muscle marks, <1 and x2, paratype USNM 200837b; 38-41, 43, Posterior, ventral, 
side, anterior and dorsal views of a young adult, x1, paratype USNM 200837c; 42, 44-47, Side, posterior, anterior, ventral and 
dorsal views of a specimen smaller than the preceding, x1, paratype USNM 200837d. 

Sly Gap Formation, locality 506c. 


Figure 
1-10. 


11-20. 


31-59. 


60-72. 


73-77. 


78-82. 


BULLETIN 315 


EXPLANATION OF PLATE 2 


Thiemella platyssnew SpECieSs ee feiec sccre/=yer- stevet- ievelousis 2 eheleaehe = ein nin) sieve teiever=belel sleted tevelcleastrekeietat 1c oie teiel tise eee ee 42 
1-5, Anterior, ventral, posterior, side and dorsal views of a complete specimen, <1, holotype USNM 200889; 6-10, Anterior, 
side, ventral, dorsal and posterior views, x2, of the same specimen. 

Percha Formation (Box Member), locality 527a. 
Aulacellatellipticainew SPECIES) a..y5:. 5. sieve o) sles ousteteyeverevsiol =! =10.0\> + insere.0 ol aneteheter skatetetspetenoteyatets eteierstelsls)ietetersr=tetel tte -teeete tet tee eee 38 
11-15, Ventral, dorsal, side, anterior and posterior views of a small specimen, <1, paratype USNM 200887; 16-20, Dorsal, 
anterior, posterior, side and ventral views, <2, of the same paratype. 

Sly Gap Formation, locality 3016 = 506). 


:. Stainbrookia infera (Calvin) ses tesinscss 5 gece sere weonevovere vo epyeynniens wo iar eS) che ove ses yeMByavereIO eats fate fone arora Gorse unrevelel coe AOE Cena 39 


21-25, Side, dorsal, ventral, anterior and posterior views of a strongly convex individual, x1, hypotype USNM 200886; 26-30, 
Anterior, posterior, ventral, side and dorsal views, <2, of the same specimen. 

Independence Formation, locality 526w. 
Aulacellaellipticasne Ww, SPECleS gate. -icy-y keg) -sevetetegetatene el sheter shot ake se") San0 e oust shee [Suetehsyeic\ ote 04 os ine ite ee ole) “isch eee CLS eee 38 
31-35, Ventral, posterior, side, dorsal and anterior views of a small specimen, x1, paratype USNM 200888; 36-39, Dorsal, 
anterior, side and ventral views, <2, of the same paratype. 

Sly Gap Formation, locality 509n. 
40-44, Side, posterior, dorsal, ventral and anterior views of a complete specimen, <1, paratype USNM 200884; 45-49, Anterior, 
posterior, dorsal, side and ventral views, 2, of the same paratype. 

Sly Gap Formation, locality 526e. 
50-54, Dorsal, ventral, side, posterior and anterior views, <1, holotype, USNM 200885; 55-59, Anterior, dorsal, side, posterior 
and ventral views, x2, of the holotype. 

Sly Gap Formation, locality 52le. 
Thiemella: quadrata new SPeCleS. jo 6. ac reyaisrs,o: cess scapace: ana tesee eoeve opoicpay aide ees brersuene/ stay siete opened seeien: eel ails See ee Eee eee 43 
60-63, Dorsal, side, ventral, and anterior views, x1, holotype USNM 203533a; 64-66, Dorsal, anterior, and side views, x2, of 
the holotype: 67-70, Posterior, dorsal, ventral and side views, <1, paratype USNM 203533b; 71, 72, Anterior and side views, 
x2, of the same paratype. 

Contadero Formation, locality 527q. 
Aulacellaielliptica: new 'SpECies: oss ius ie:sicca’escnans aisiaraca a eraveuesevelayerd ore ae 6 a a eneiorahehepebere eee atenctes ets, aioe: Siete 16) 2! keene PAT Siete One eRReR a 38 
Ventral, anterior, side, dorsal and posterior views of a complete specimen, x2, paratype USNM 203562. 

Sly Gap Formation, locality 527r. 
Cariniferella io wenisis Stainbrook ossacciaieyosd cusps agave siaoi sche olopsue vente aueue siete vey auels Gesele rsh ckiakes ene ayeyeyele aneler eae « ere OC eee eee 41 
Anterior, posterior, dorsal, side and ventral views of an adult, x1, hypotype USNM 200832. 

Sly Gap Formation (Bed H), locality 521s. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


Wy 
% 


ih 


y] 
" 


| 
it’s 


62 


PLATE 


> 


PINS OF 


AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


PLATE 3 


Figure 
1=5 


6-16. 


17-25. 


DEVONIAN BRACHIOPODS OF NEw MEXICO: COOPER AND DuTRO 159 


EXPLANATION OF PLATE 3 


Schizophoriawiowensisi(ELAlL) sc yeveyezevesetens)=isloreceste av cienede tenereve tse stesetonch ote etexstatay ereteus cteveieievstsaseevsnsu-iel=a-Uorsrsnet-1steterenaKaeete iaiererexearireeererereL Tete 37 
Posterior, anterior, side, ventral and dorsal views of a large specimen, x1, hypotype USNM 200955. 

Sly Gap Formation, locality 509n. 
SchizophoriailatayS tain brooke 3 5 = sree fare yesei eee ete i ste peet a veo os sh es salen te sy els Pavey evra fet ore eyorstane dev ays = /aylsss¥s eteya sade et ovevopeene teen aiasebodet 36 
6-10, Posterior, ventral, side, dorsal and anterior views of a small specimen, x1, hypotype USNM 200853; 11, Ornament of the 
same specimen, ventral view, x2. 

Onate Formation, locality 523d. 
12-16, Anterior, dorsal, ventral, posterior and side views of a large adult, x1, hypotype USNM 200956. 

Onate Formation, locality 523e. 
Canriniferellatalleniine w/SpeCles sy iiec cir isiche teeta re teue erties donne seaehcscia ore codec yedavevice faxeucucis) Shaper overs vamp steers a ais teers tocieeaetel es 40 
17, 18, Interior of the dorsal valve showing cardinalia, x1 and x2, paratype USNM 200836. 

Sly Gap Formation (18 inches above base of bed 8), locality 521p. 
19-23, Posterior, side, ventral, dorsal and anterior views of a large adult, x1, paratype USNM 200833; 24, 25, Dorsal and ventral 
views of the same specimen, 1.5. 

Sly Gap Formation, locality 5091. 


Me CariniferellaySPEClES weet crete trees ake peers oe eisai civoral's esl ofcfen spa vayeehe Poy otenoneeeeeIcWswaps eke ce navekot uc vs davelievaregsfoye, seastedeys dopo Saye PRC EY stensT= irene Botte 41 


Dorsal (interior) view of exceptionally large ventral valve, <1, figured specimen USNM 200835a. 
Sly Gap Formation, locality 52le. 


160 


Figure 
1-6. 


22-27 


28-33. 


BULLETIN 315 


EXPLANATION OF PLATE 4 


Schizophoria’ Species Wey yoyo ete) excce: ayes ave oto se seh s\s (w elepehatevate tate loleke to iataralenae= olace a) aol eis aiae/or os olatetsienet Reeperayo) satel tafele tte tate itee ett 37 
1-5, Dorsal, ventral, side, posterior and anterior views of a complete specimen, x1, figured specimen USNM 200908; 6, Ventral 
view of the same specimen showing hollow costellae, <2. 

Sly Gap Formation (lower), locality 506d. 


2 Strophodontaliumbonata new SpeCies. < .<.orere\ers ekcre rs el == tee neda da fore ed aay eyet cde eye olassn esos ojanata aferseatebepeRelei salads ts ohelskelel ste kek tetet te tet a eens 44 


7-11, Posterior, side, dorsal, anterior and ventral views of a young specimen, <1, paratype USNM 201084b; 12, Dorsal view of 
same specimen, x2. 
Sly Gap Formation, locality 524m. 


2 Propidoleptus) platys mew SPECIES) cret-iaterersietss tered -foverslat)<ayops clay ara) eps eketoteyateleranea(elelelc clemeateltetelepevatatetaret- Pets ttet iets tetet tie tee 43 


13, Rubber impression of a large specimen seen from the dorsal side, x1, paratype USNM 201100. 

Onate Formation, USGS (Kirk) locality 540. 
14, 15, Dorsal and ventral views of a young specimen, <1, paratype USNM 201101a; 16, Exterior of a large ventral valve, x1, 
paratype USNM 201101b; 17-19, Ventral, dorsal and side views, x1, holotype USNM 201101le; 20, Dorsal view of the holotype, 
x2; 21, Dorsal (exterior) view of a dorsal valve, x1, paratype USNM 20110Ic. 

Onate Formation, locality 521z. 


» SchizophoriarspECies) 2 save jcieyeyscerersh nes evsse vores lave toate ete uss ceueedes chee) ete iecelotererels avai e eile state spear Rokenscale iets cite skeen 38 


22-26, Posterior, anterior, side, dorsal and ventral views of a well-formed specimen, x1, hypotype USNM 200852; 27, Exterior 
of the ventral side of the same specimen, showing hollow costellae, x2. 
Sly Gap Formation, locality 509p. 
Schizophoria:amanaensis:StainbroOke: ccs csi sae 5 ccce0s sesse ts wie = eats aleve a Rua lotager era einleveuets\tese ¥ eve ohoteiotae iene Sere eres Patete area en ner eR ene eo 35 
28-32, Posterior, anterior, side, dorsal and ventral views of a complete specimen, <1, hypotype USNM 200880; 33, Exterior of 
the ventral valve of the same specimen, showing ornament, x2. 
Sly Gap Formation, locality 521d. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 


Figure 
1+4. 


9-18. 


19-23. 


24-39. 


40-51. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 161 


EXPLANATION OF PLATE 5 


Strophodontayspeciesw Lerrterenyerererois eaten vee reverie rere renee PeT ech sls yeverene aie onauereva kone rene ievere oucvareievercteae(orsycverexsuslieychereravccieysrenetarencrstaiara cee meeenetalete 44 
Ventral, posterior, side, and dorsal views of a large thick-shelled specimen with large interareas, <1, figured specimen USNM 
201085. 

Onate Formation (beds C to D), locality 523e. 


> SERIA Goss As syecenoeessnnacedepposaccoopEsucsonesonooDonoacens GeeGoonenSo ones Guo psecdodaroocpoDUOUOODO OD 44 


5-7, Side, ventral and dorsal views of a poorly preserved complete specimen, x1, figured specimen USNM 201181; 8, Dorsal 
view of the same specimen, x2. 

Sly Gap Formation, locality 526e. 
Sirophodontarampona lane WaSPEClES stem et ee teyeys iterate terres never rete etesta et orchete nea ote ete dete tedet = eliotete atl ofetetoceyatetl tet er<t tele E-yevett ier 44 
9-13, Dorsal, posterior, anterior, side and ventral views of a large specimen, <1, holotype USNM 201083. 

Sly Gap Formation, locality 509n. 
14-18, Side, ventral, anterior, posterior and dorsal views of a young adult, x1, paratype USNM 201084a. 

Sly Gap Formation, locality 524m. 
Strophonelloidesideeringt |S tain DROOKs yey: fepai tay) tar cote ore anata ehedct eters heyeltevenstsisy yore ore otter sep are insite ee ronclo eee eee rerkeiiete tee e Erol 45 
Ventral, posterior, side, anterior and dorsal views of a well-preserved specimen, x1, hypotype USNM 200865a. 

Independence Formation, locality 527s. 
CERT OTE GES AAS WEN GUSEESS sraeccaccacndoncnonn bse sdoccooonn on ddonobOoon aEdoDCoUsUDCEoSSEs Oooo oEEBdooobesuE 45 
24-28, Ventral, posterior, anterior, side and dorsal views of a complete adult, x1, holotype USNM 200866; 29, 30, Ventral and 
dorsal views of the same specimen, x2. 

Sly Gap Formation, locality 509g. 
31-35, Anterior, side, posterior, ventral and dorsal views of another complete specimen, x1, paratype USNM 200867b; 36, 37, 
Dorsal and ventral views of the same specimen, x2; 38, Interior of the ventral valve, x2, paratype USNM 200867a. 

Sly Gap Formation, locality 506c. 
39, Interior of an obese dorsal valve, x2, paratype USNM 200872. 

Sly Gap Formation, locality 528x. 
Strophonelloides dorsoconvexus NEW SPECIES. .. 2.2. ee eee eee ee ee nee ee eeeeeeseeeees 45 
40-44, Anterior, dorsal, posterior, ventral and side views, <1, holotype USNM 200862; 45, Exterior of the ventral valve of the 
same specimen showing the ornament, 1.5. 

Sly Gap Formation, locality 509n. 
46-50, Dorsal, side, ventral, anterior and posterior of a smaller specimen, <1, paratype USNM 203532. 

Sly Gap Formation (lower 30 feet), locality 527t. 
51, Interior of a dorsal valve, x1, paratype USNM 200864. 

Sly Gap Formation, locality 509r. 


Figure 
|-6. 


BULLETIN 315 


EXPLANATION OF PLATE 6 


Nervostrophiaiplama mews SpeCiessy cir carere fee celetene fale ele) fe cle ot leveroteyed ctafokersis te cteroyoteite arate kelstoketetsie seers eters letersi chet sts eee 48 
1, 2, Interior of the ventral valve showing muscle marks, x1 and x1.5, paratype USNM 200929. 

Onate Formation, locality 522h. 
3, 4, Two ventral valves showing hinge extensions and ornament, respectively, both x1, holotype USNM 200841la, and paratype 
USNM 200841b. 

Onate Formation, locality 523u. 
5, 6, Exterior of a dorsal valve showing ornament, <1 and x2, paratype USNM 203531. 

Onate Formation, locality 527u. 


» Nervostrophiaextensa new SPECIES: ae. se.c.< i505 sssie:snveseve (w/e: ars isis sie v-scorsi sie eve retesele elete/ovele. oie tors -cishehetetesve (ele) svsieds ehh / tens ete arate eet eee 46 


7, Dorsal view of a nearly complete specimen, <1, holotype USNM 201088a. 

Independence Formation, locality 526y. 
8-12, Ventral, anterior, posterior, dorsal and side views of an imperfect specimen, <1, paratype USNM 201034; 13, 14, Ventral 
and dorsal views of the same specimen, showing ornament, x2. 

Sly Gap Formation, locality 506c. 


: Nervostrophiaygeniculataimew SPECIES. -yarrey-tclers ateh= crore. cies oles + «ve cin olcceratetaledeleleterehevebeieistetetstate sielslatatelsratele leit teyetetetststeteta eet ites 46 


15-18, Ventral, side, anterior and posterior of a ventral valve, x1, paratype USNM 200932. 
Sly Gap Formation, locality 509h. 
19-23, Anterior, posterior, ventral, side and dorsal views, x1, holotype USNM 200934. 
Sly Gap Formation, locality 521p. 
24, 25, Dorsal and side views of another specimen, x1, paratype USNM 200933b; 26, 27, Dorsal views of the same specimen, 
x1.5 and x2; 28-31, Ventral, anterior, posterior and side views of a ventral valve, x1, paratype USNM 200933a. 
Sly Gap Formation, locality 506c. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 6 


INS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 7 


Figure 
1-16. 


17-21. 


26-35. 


DEVONIAN BRACHIOPODS OF NEW MEXxICco: COOPER AND DuTRO 


EXPLANATION OF PLATE 7 


INelvostrophiay mucrospiname Ww) SPECIESt. erie vets crencrit recat se car yereus ose erences cuershartinieae ei oueletegegaial susiaytictorcurusrerorsteterme ric tencietasesteretene 
1-4, Posterior anterior, side, and ventral views of a ventral valve, x1, paratype USNM 201024b; 5, Ventral exterior of the 
preceding specimen, x2, showing ornament; 6, A small ventral valve, x2, paratype USNM 201024c; 7, 8, Ventral views of a 
well-formed specimen, x1, x2, holotype USNM 201024a. 

Sly Gap Formation, locality 509p. 
9, 10, A ventral valve showing the long hinge extensions, x1, x2, paratype USNM 201026b; 11, 12, Exterior of a small dorsal 
valve, x1, x2, paratype USNM 201026a. 

Sly Gap Formation, locality 506). 
13, 14, Another ventral valve showing lateral points, x1, x2, paratype USNM 201028. 

Sly Gap Formation, locality 52le. 
15, 16, Dorsal valve showing some details of the ornament, x1, x2, paratype USNM 201039. 

Sly Gap Formation (lower), locality 521k. 
NegvostrophiaiciaN: multinervosa StainbroOk: oo... ws jje cue wesnrs rae eles Sires om ee ere elale sie nie syesnieie uereynie,« wisuminloceeieie. cre eesexetetelstelofes0/ei0 
Posterior, side, anterior, dorsal and ventral valves of an imperfect specimen, <1, hypotype USNM 201087. 

Sly Gap Formation, locality 506c. 


7 Nervostrophiaymultineryosai StaimbrOOk: 1) s)sie/ ccc 2.5/0) 9 o0ss wrevein sisinv eve) sig. ckerore niciaie aisieieleve oe: #1 veil steamiihs info) staie iotei share reveyarerevoiehelesehaler= elsies< 


22, 23, Exterior of a small ventral valve, x1, <2, figured specimen USNM 201035b; 24, 25, Another ventral valve with mucronate 
cardinal extremity preserved, x1, <2, figured specimen USNM 201035Sa. 

Sly Gap Formation, locality 521d. 
Nerv Ostropliasolidanew?SPOCleS 2 eiz tae :ny-csresvs2y ane ous: svoveyereney ep eae teensy oh ele ovncn ste ar erates eves note teueves ests <tayetepe ele weqenete etekensy se ameteretenetersereketeas fev 
26-29, Posterior, dorsal, side and ventral views of an imperfect specimen, x1, paratype USNM 200842. 

Sly Gap Formation, locality 509k. 
30-33, Side, ventral, dorsal and posterior views, <1, holotype USNM 200843c; 34, Dorsal view of another imperfect specimen, 
x1, paratype USNM 200843a; 35, The posterior region of the same specimen enlarged to show the chilidium and narrow elevation 
of the pseudodeltidium, x2. 

Sly Gap Formation, locality 506c. 


48 


48 


48 


Figure 


BULLETIN 315 


EXPLANATION OF PLATE 8 


1-7... Pseudodouvillina:euglypheaStainbrook., ~ <.<.615.5..12:-roois sre ole of ie1sie otete ove) wie o¥e 010 ,c0/eisis els olelot helolele iterate oietel setter ic Eerie eteieiee eee tenemee 49 


8-13. 


14-19. 


20-25. 


26-39. 


40, 41. 


1, Dorsal view of a complete specimen x1, hypotype USNM 20105Sa; 2—6, dorsal, anterior, posterior, side and ventral views of 
the same specimen, x2; 7, Interior of the ventral valve, x3, hypotype USNM 201055b. 

Sly Gap Formation, locality 506c. 
Douvillina minutainew species: 2.05. s.0eceizys.¢ aie s scvtssateysiaiece oxa.e cca s oe soe, erelers/alevo-0 01ar5;q\ averse ois nbove) eras a sve ahels |p eREIETeT= oe ee kee ee tee eee 50 
8, Dorsal view, <1, holotype USNM 201056; 9-13, Ventral, dorsal, posterior, side and anterior views of the holotype, <2. 

Sly Gap Formation, locality 506c. 
Douvillina: arcuata; (Fall): :<.<.o::<.000e:sa.5; sy 0135.05 aceus 3 ese aids ewes oni tu royevers.np8 8/0) ate falas ecsy <yoee eyo) xsl ny es voieusyekese sede stole [ates eee anak eee eee 50 
14, Dorsal view, x1, hypotype USNM 201044; 15-19, Anterior, side, posterior, dorsal and ventral views of the same hypotype, 
xD: 

Sly Gap Formation, locality 506c. 
Sulcatostrophiatcalyvimis (Miller) 5 cyec vv petete ater = ein) ov yovere mo ceteveve oie toretet =) levelereeroteie) a lel 1olniolotebeeeteheetetalal=i-iefeceteeciate stellata tetanic eee 50 
20, Ventral view, x1, hypotype USNM 201138; 21-23, Posterior, ventral and side views of the same hypotype, x2. 

Sly Gap Formation, locality 509m. 
24, 25, Dorsal and ventral views of two specimens, <3, hypotypes USNM 201059a and b, respectively. 

Sly Gap Formation, locality 521d. 
Douvillina contaderoensisinew SPECIES! <q) /)e.eccre 5 orereisiate © orace o\si 6 © 6 se elehe er eteeloxorere\® uel o4s epefeley sl sls\a.e “Leelee ere ee eee 50 
26-30, Dorsal, ventral, side, anterior and posterior views of a complete specimen, x1, hypotype USNM 200970; 31-35, Ventral, 
dorsal, posterior, anterior and side views of another complete specimen, x1, hypotype USNM 200971; 36, Dorsal view of the 
same specimen, x2. 

Contadero Formation, locality 521). 
37, 38, Exterior views of a ventral valve showing ornament, x1, x2, hypotype USNM 200991b; 39, Interior of a poorly preserved 
dorsal valve, x1, hypotype USNM 200991a. 

Contadero Formation, locality 509t. 
Nervostrophia solidai Mew. SPeCleS: «roi ,csescitejexece 1 oe o:0/anys\'s;etescre ovescsiayalays tiers Aetectarormetene ove eusuchaienereldiekausr evs: casiotie le eielere tetra ton toro eee 48 
40, Exterior view showing ornament in detail, x2, holotype USNM 200843c. 

Sly Gap Formation, locality 506c. 
41, Posterior of the interior of a dorsal valve showing the cardinalia, x2, paratype USNM 200879. 

Sly Gap Formation, locality 521n. 


42, Dorsal view of a complete specimen, x1, figured specimen USNM 203561a; 43-47, Ventral, dorsal, side, anterior and posterior 
views of the same specimen, 2; 48, Ventral valve of another specimen, x1, figured specimen USNM 203561b. 49, Dorsal view 
of a complete specimen, x1, figured specimen USNM 204951; 50-54, Dorsal, ventral, side, anterior and posterior views of the 
same specimen, 2. 

Contadero Formation (coral bed), locality 527v. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 8 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 9 


Figure 
ite 


2-9. 


10-13. 


14-25. 


26-37. 


38-44. 


45-50. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 165 


EXPLANATION OF PLATE 9 


INenv ostrophiagextensaine wi SPCCles segs sre rece cs, shes averenetet enon eter euoteuotcnere  pelciatcrer omnes oncerenoanet el lescaiavs dar atic Mon aaa VaInTE eiois Cat ccee ee Tee 46 
Ventral view of an adult specimen, «1, paratype USNM 201093. 

Sly Gap Formation, locality 521p. 
DouvillinapminutameweSpeCles ack vweterene orc istenese eres eee ty co wie ehereretere Nee eve EN OE Nels akc Gs aoahe sean TCL EEE ene 50 
2-5, Side, anterior, posterior and ventral views of a ventral valve, <3, paratype USNM 201076. 

Sly Gap Formation, locality 521n. 
6-9, Side, anterior, posterior and ventral views of another ventral valve, x3, paratype USNM 201075. 

Sly Gap Formation, locality 524m. 
Oiosia;putillay(StainblOOk) hy tae steers ele ihete lov Ame rove Meypee bores arse eel hore rare lade Sic Tee eae ee Sec ne ER oR epee 117 
10-12, Anterior, dorsal and side views of a well-formed individual, x1, hypotype USNM 204955a. 

Percha Formation (Box Member), locality 506b. 
13, Fragment showing pits left by stripping of small exterior spines, x4, hypotype USNM 204956. 

Percha Formation (Box Member), locality 527d. 
DouvillimayparvilsculamMewASPeCiess (<6 2 evoys\esciszasis eigicis siayensy a Gebel asec oiercteys.< Salas s) <1 4 steroid RG eI Oe ER ERE OE re eine 51 
14, Dorsal view of a paratype, x1, USNM 201057a; 15-19, Side, ventral, dorsal, posterior and anterior views of the same specimen, 
x2. 

Sly Gap Formation, locality 52le. 
20, Dorsal view, x1, holotype USNM 201054; 21-25, Posterior, ventral, side, anterior and dorsal views of the same specimen, 
SOD, 

Sly Gap Formation, locality 509r. 
Douvillinayrectangulataimew SPeCles =: ve ctereyo stele ici ere eee vo eee Leasu cide ene pe eave Sra Shaves ae Cente oie eens elie 51 
26, Dorsal view of a complete specimen, <1, paratype USNM 201030; 27-31, Side, dorsal, ventral, posterior and anterior views 
of the same specimen, x2. 

Sly Gap Formation, locality 509n. 
32, Dorsal view of another complete specimen, x1, holotype USNM 201031; 33-37, Dorsal, anterior, side, ventral and posterior 
views of the same specimen, x2. 

Sly Gap Formation, locality 521e. 
Douvillinaria‘variabilis\(Calvan)'s, cz. ers evecerore eis) oesavere eas avere artic era etree ooo eaves accle eel ue Hee ORI SiGe aE Cee 53 
38-42, Ventral, anterior, dorsal, posterior and side views, x1, hypotye USNM 20087 1a; 43, Dorsal view of the same hypotype, 
<2; 44, Interior of a dorsal valve, x2, hypotype USNM 20087 |b. 

Sly Gap Formation, locality 506c. 
IBMT GCC Ih. i.e doa ciaets Sno ad ob AGI pS camcdia co crc eR noeencrant ar aceon Mork aes on comoom.cadoomeccododud ohabhe oe 52 
45, Dorsal view of a complete specimen, 1, figured specimen USNM 201040; 46-50, Posterior, ventral, side, anterior and dorsal 
views of the same specimen, x2. 

Sly Gap Formation, locality 509j. 


51, Dorsal view of a complete specimen, <2, paratype USNM 200869b. 
Sly Gap Formation, locality 52Ic. 

52, Interior of a small dorsal valve, x2, paratype USNM 200868. 
Sly Gap Formation, locality 509k. 


Figure 
1-14. 


34-36. 


37-40. 


41-46. 


47. 


BULLETIN 315 


EXPLANATION OF PLATE 10 


Douvillinaria:diversa NeW sSPEClES®, 0. Ay.5sscjarete eicienestor sires socs sre ete (v= Sotelo) 1asavevoNe ohereher Sieve ce averevel a fela araye ee ee aeRO Reece eee een 53 
1-5, Dorsal, posterior, side, anterior and ventral views of a complete specimen, <1, holotype USNM 200869a; 6, 7, Dorsal and 
ventral views of the same specimen, x2. 

Sly Gap Formation, locality 521c. 
8-12, Posterior, anterior, ventral, side and dorsal views of a specimen smaller than the holotype, x1, paratype USNM 200870a; 
13, 14, Ventral and dorsal views of the same paratype, <2. 

Sly Gap Formation, locality 509h. 


. Floweriaiprava (El alll)... .q... c-cxsueteyeinuce a erty receketelete pee (ie ele ee 0ay seven ete. Sei le eusieho inlet eded ovehe delete ones ai cCRC PS lc es ote ate 54 


15-19, Dorsal, posterior, anterior, ventral and side views of a small individual, x1, hypotype USNM 200874; 20, 21, Posterior 
and dorsal views of the same specimen, x2. 

Sly Gap Formation, locality 521e. 
22-26, Posterior, side, ventral, dorsal, and anterior views of a flatter specimen, x1, hypotype USNM 200854; 27, Dorsal view of 
same specimen, x2. 

Sly Gap Formation, locality 509z. 


33. Floweria magnacicatrix! NEW'SPEGCIES.. «sis .:i:ere ie peer 5 51s ye:crsiepe oe o-o.506 o arcs oie epsilesac4.6/ai «/eyciele) s+ © 6 shes) alisivic o1els"s/s/eial oleate s GleReteteteeeletetee 55 


28-32, Anterior, dorsal, side, ventral and posterior views of a complete specimen, x1, holotype USNM 200855; 33, Dorsal view 
of the same specimen, 2. 
Sly Gap Formation (below sill), locality 509h. 
Schuchertella'resupinata new SPeCieS. «2/0 cie cscs ccigs osinersies ove sacle etetn ws a 45 o/s) sreiaire'eys are cle avetele alee keeper eens 56 
Dorsal, side and ventral views, <1, holotype USNM 201189a. 
Percha Formation (Box Member), locality 509e. 
Devomochonetes? Sp. e.scccstevesaieiese sce Guarentee w cncreveseev's © wey avin aie a separ's erelioya’ ave ene) devs-inve rar Cnchee ava Va cel ek cuahe yoy ep STaN chal re Seo) 6 SLT eee ae eae 57 
37, 38, Large ventral valves, x1, USNM 203543a, 203542. 
Onate Formation (base), locality 527w. 
39, 40, Small and large ventral valves, <1, figured specimens USNM 201089b, a. 
Onate Formation (Bed B), locality 526b. 
Hloweriaitramsversalis: MEW SPECIES. xe ae.e.:, 015 erie ose eres sere: a.e paar eer sre (sh cye vesserevoy ovoid 0 ne OR eer eT ona o SICT ORTOP eI eI TCE earn ae 55 
41-45, Anterior, ventral, posterior, side and dorsal views, x1, holotype USNM 200873; 46, Dorsal exterior of the holotype, <2. 
Independence Formation, locality 362a. 
Schuchertella\perchia:(Stain brook). 3.27c/saie rare cascicns stats conse ego tes ceuia ca ese, cy cyeen ape tr cheer apenas Sr ezever eke re)S  eealors Pee RPT RES peer Re eae eee 56 
Dorsal view of a complete specimen, x1, hypotype USNM 201133. 
Percha Formation (Box Member), locality 3026a. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 10 


LETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


PLATE 11 


Figure 


13-18. 


19-25. 


26-36. 


37-43. 


44-46. 


DEVONIAN BRACHIOPODS OF NEW MExiIco: COOPER AND DuTRO 167 


EXPLANATION OF PLATE I1 


Page 
Plo weriavaiieehchemunpensisy(Gonrad) scree cct tec. csisraetr ctr atars scerstereett ic <i sferevarein eqchenenetstentte eietekatetchenc)s, eit sieeue cee eteevensearere etal eteayaehetel 55 
Ventral, dorsal, posterior, anterior and side views of a complete specimen, x1, hypotype USNM 200875. 
Contadero Formation (top), locality 523n. 
seStriatochonetes/subCcarimatus MEW SPECLES bey eys.c = areas exe okevar si wictaxsuntolele) ehevecer ay wisioysie oln)=ie)s)avaveretelatayelieyo/e nisi anc ere/ofofel el eieve ere eicletstatoval™ olele/« 58 


6, Dorsal view of a complete specimen, 1, holotype USNM 201080; 7-11, Side, dorsal, posterior, anterior and ventral views of 
the same specimen, 2; 12, The same specimen, showing the ornament, 3. 
Onate Formation, locality 523f. 
Striatochonetes mans me wsSpeClesommess cia v-verieeretertereeeserareiereieicictcteyeietctereieieeteteie esorerststete retro teteten-c= icici deol eetsistet-Taiieteteteh tele tetera crest: 57 
13, Exterior of a ventral valve, x3, paratype USNM 201078a; 14, Interior of a dorsal valve, x3, paratype USNM 201078b. 
Onate Formation, locality 524g. 
15, Exterior of another ventral valve, x3, holotype USNM 201079b; 16, Interior of a dorsal valve, x3, paratype USNM 201079a. 
Onate Formation, locality 522i. 
17, Exterior of a ventral valve, x3, paratype USNM 201077b; 18, Small slab, showing interior of two dorsal valves, <3, paratype 
USNM 201077a. 
Onate Formation, locality 522d. 
RetichonetesyobScurus NEW SPECIESs ec iso.<. 5:5; o:c.scseeueyercvene si spate ssstevaia ieiie sone = areucfave:‘e.e a, se evaysts cbeieteyo oyars sie egeves ale [elu iecovesegeteleneteveretorepe ecto. = 58 
19-23, Posterior, anterior, side, ventral and dorsal views, x4, holotype USNM 201124a; 24, Interior of a dorsal valve, <4, 
paratype USNM 2011 16a. 
Percha Formation (Box Member), locality 506q. 
25, Exterior of a small ventral valve, x4, paratype USNM 201115. 
Percha Formation (Box Member), locality 5070. 
Retichonetes)perchacusisille wiSPeCles smeteteye tere etetet ole tevetedotel ates teredetetatet ster ofetesietetckeketetetatetetalereteetetereieiebstsistsieteferereietetetetateretetateeerteenee ret ea 59 
26-30, Anterior, ventral, dorsal, posterior and side views, x4, holotype USNM 201122. 
Percha Formation (Box Member), locality 527p. 
31, Exterior of a ventral valve, x4, paratype USNM 201123. 
Percha Formation (Box Member), locality 506b. 
32, Decorticated exterior of a ventral valve, x3, paratype USNM 201119. 
Percha Formation (Box Member), locality 3043. 
33, Exterior of a small ventral valve, x3, paratype USNM 201120. 
Percha Formation (Box Member), locality 506. 
34-36, Anterior, side and ventral views of a ventral valve, x3, paratype USNM 201117. 
Percha Formation (Box Member), locality 507o. 
Rhyssochonetespy OMNSOMILNE WAS PE CLES emp yetetetepegedelel ete reterereleratevabete/ olla toterrtestalisteroyerelolatetev=fatets ares re erste hete tet shereiataolaieteesitetetetetee rats eters 59 
37, 38, Two ventral valves, the smaller one showing a long spine on the cardinal extremity, x3, paratypes USNM 204953a, b. 
Onate Formation, locality 527x. 
39, Ventral valve exterior, showing concentric lines, x3, paratype USNM 201081a; 40, Exterior of another ventral valve, showing 
concentric lines, x3, holotype USNM 201081c; 41, 42, Interior of a dorsal valve, x4, paratypes USNM 20108Id, e; 43, Interior 
of a ventral valve, x4, paratype USNM 201081b. 
Onate Formation, locality 522n. 
roductellarispeChes: 2 ieavssese sca ia cqetenee strate ss eeovovo Gn 16 choise a wreceve tc ouaraneiere tale nears is MoRoke sieves cue se ete ye job tush geiayanas oueelotehe leseieas atecayevofers ohere 63 
Posterior, side and ventral views of a large specimen, x1, figured specimen USNM 201143a. 
Contadero Formation (?), locality 5280. 


21-28. 


29-40. 


41-46. 


BULLETIN 315 


EXPLANATION OF PLATE 12 


Page 
;. Eostrophalosiairockfordensis| (Hall and!'@larke). 7 oo... se. ecg ckele esi ee Hiss ie ele ore 2 il eeleleieiafeie cralenioieie oie 2 ele) elsiet sae eet eres 60 
Exterior and interior of a dorsal valve, x2, hypotype USNM 200838. 
Hackberry Formation (Cerro Gordo Member—Strophonella Zone), locality 527y. 
; Eostrophalosia}inexpectata New SPECleSs. <icr-yarers sve miele: leo ee lahore velnoin ole /e(ejonel ele iste euekor= ohoeevaac foteserlererelerete sl Ton ter keiek tte eaent 60 


3-7, Dorsal, anterior, side, ventral and posterior views, <1, holotype USNM 200926; 8-12, Anterior, posterior, ventral, dorsal 
and side views, showing cicatrix of attachment, of the holotype, 2. 
Sly Gap Formation (zones b-1), locality 509r. 


sProductellayrupatulayStainbroOk., fe rrscre tee cuss syelevessdels) steves-vele)beheyeusl=) o)ateroratetole ete teteroetettetever selene cfele Le else hayes 62 


13-17, Anterior, ventral, side, posterior and dorsal views of a complete specimen, x2, hypotype USNM 201120. 

Sly Gap Formation, locality 526n. 
18-20, Posterior, anterior, and ventral views of a ventral valve, x2, hypotype USNM 201118. 

Sly Gap Formation (43-83 feet above base), locality 522k. 
Leioproductus) paupercultis new SPECIES. 2. 0)<) 5.6: siesevoie as ciov5s 1s 10) sce nls: opsyarasefate) fe) cierererenaia) okeiaeveveretetetet-torejcyehelelrais elie Ratettehe tet tte tae iat 64 
21-23, Ventral, side and anterior views of a ventral valve, x1, paratype USNM 201110a. 

Percha Formation (Box Member), locality 506. 
24-27, Side, anterior, posterior and ventral views, <1, paratype USNM 201111; 28, Ventral view of the same paratype, x2. 

Percha Formation (Box Member), locality 506p. 
Productella:species: Ws, isis. c:e:cccssus, ste wiery susie es aisness tecerenscererenees fede) Suh $x sca dbs aera cyojiera be veney ayes hepahe ce Vie Meee See Ie etere eee eRe ee eee 62 
29-32, Dorsal, side, ventral and posterior views of a complete specimen, 1, figured specimen USNM 201011b; 33, Anterior of 
another ventral valve, <1, figured specimen USNM 20101 1a. 

Rhodes Canyon Formation, locality 521h. 
34-37, Side, posterior, anterior and ventral views of a ventral valve, x1, figured specimen USNM 203563a; 38, 39, Side and 
anterior views of another ventral valve, <1, figured specimen USNM 203563b; 40, Interior of a dorsal valve, 2, figured specimen 
USNM 203563c. 

Rhodes Canyon Formation, locality 527z. 
Dichacaenia umbonata NEw SPECLES . 6. e.ciz isis csissrciece avevele dose orm wisvejotesaiele wre teyeve sya 54 asaue ol sca este ve mueione oyereers fetelterene lel «cetera er eee 61 
41, 42, Exterior of two ventral valves, <1, paratype (USNM 201012b) and holotype (USNM 201012a); 43, 44, Interior of a dorsal 
valve, x1 and x2, paratype USNM 201012d. 

Onate Formation, locality 523s. 
45, Interior of another dorsal valve showing strong breviseptum, x2, paratype USNM 201013. 

Onate Formation, locality 523i. 
46, Interior of an imperfect dorsal valve, x2, paratype USNM 204954. 

Onate Formation, locality 526c. 


55. Dichacaenia perplexa NEW SPECIES... :5.5-s.2;e:510)50512 75,5: 9 oysioys1 0 5'ev0¥0) obese) tus rate o 416) ef ap spapous svar acdhei eus'epeiee ele) ahi o <rese/eroeece ele ee ene 61 


47-49, Dorsal, ventral and side views, x1, holotype USNM 200957a; 50-54, Ventral, side, anterior, posterior, and dorsal views, 
showing short recumbent spines, x2, holotype USNM 200957a; 55, Posterior of a paratype, showing interareas, x2, USNM 
200957b. 

Onate Formation, locality 5230. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 12 


INS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 13 


Figure 


10-17. 


36-56. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DuUTRO 169 


EXPLANATION OF PLATE 13 


2-6, Side, posterior, ventral, anterior and dorsal views of complete specimen, x1, holotype USNM 201112a; 7-9, Anterior, dorsal 
and ventral views, <2, of the holotype. 

Percha Formation (Box Member), locality 509a. 
Gypidularsubcarinatalnew/ SPECless toy-rctereyereretets stereo) shel eyed ve PoToTe ee chet scr eden Vest ver ele spor esol a elaiaieteratetalancisvokerotalailietelssefetatstenet=yabetet=later= 
10-14, Posterior, anterior, ventral, side and dorsal views, x1, holotype USNM 200850; 15-17, Side, anterior and dorsal views, 
«1.5, of the holotype. 

Sly Gap Formation, locality 521k. 


Pa GY PIGUlAayIMiMICARNE WASPECLES he aiereevteeeystel srs etetet ote Relea tote emetenet ater = ets het steered cacieveletsbey oicionts ter ere tel vercveveteteneie rect rete feketenelens Estee Re Reeeete mer Ki 


18-22, Posterior, ventral, side, dorsal and anterior views, <1, holotype USNM 200820; 23-25, Anterior, side and dorsal views, 
1.5, of the holotype. 
Sly Gap Formation, locality 506c. 


> Gxt oye WOW SESE, posocescopnasooun cos ou DORE GGHDoCdEesO0NS dnbobbenBoobUL SoouuEHOuUoKBDdoDEDONS OOCOBeODEC 


26-30, Anterior, ventral, dorsal, posterior, and side views, x1, holotype USNM 200824; 31-35, Posterior, ventral, side, anterior 
and dorsal views, <2, of the holotype. 
Sly Gap Formation, locality 526f. 
Devyonoproductus intermedius new SpECi€S. .. 2... 2655. cee ce ee ee ee eet eee net tee teens er esenssensecarcces 
36, Dorsal view, x1, paratype USNM 201027d; 37-41, Ventral, dorsal, side, posterior and anterior views of the same paratype, 
x2: 42, Dorsal view, x1, holotype USNM 201027a; 43-47, Anterior, dorsal, side, posterior and ventral views, <2, of the holotype. 
Sly Gap Formation, locality 509n. 
48, Dorsal view of a specimen smaller than the holotype, x1, paratype USNM 201051a; 49-53, Dorsal, posterior, side, anterior 
and ventral views of the same paratype, 2. 
Sly Gap Formation, locality 506). 
54, Exterior of a ventral valve, showing costellae, x2, paratype USNM 201045a. 
Sly Gap Formation, locality 52le. 
55, Interior of a dorsal valve, x3, paratype USNM 201052a. 
Sly Gap Formation, locality 521k. 
56, Interior of another dorsal valve, x3, paratype USNM 201058. 
Sly Gap Formation, locality 506c. 


Page 
PREPROCUCTEN AS PE CLES HS 2: wersyaretevecenscoterst averevcre stone neRe ia evevs seers uss ors cereieivey Saar afchete ere ree etcrsye reer ete teu eaalveye 76 ieiecevere¥eve fanevayole speket apsaeierevoRee RMR Re eremeeee 63 
Exterior of a ventral valve, <1, figured specimen USNM 201086. 
Thoroughgood Formation (basal bed), locality 522. 
> Letarad hating PeNTraanS MENT GoeTIES: cossoogsonpedocoosanmunnnscacopon dn boDBUEUUonooDOUUCAnEsEDOOCDeHOOODGSeU ODOC ALAND 64 


69 


68 


67 


Figure 
1-10. 


11-18. 


19-44, 


45-49, 


66-73. 


5. Coeloterorhynchus species. 


BULLETIN 315 


EXPLANATION OF PLATE 14 


Gypidulaibowsheriinew/ Species ys .c/r-!-t cite ole eee sired) ticle sheer feloletetacreratoheleleieteleotekeie eeteleteeteleteleie aol ieee 67 
1-5, Dorsal, ventral, posterior, anterior and side views, <1, paratype USNM 200823; 6-10, Dorsal, posterior, anterior, side and 
ventral views of the same paratype, x2. 
Sly Gap Formation, locality 509j. 
Gypidulaistainbroolkime wi SpeCleSe ers wreroyetel=\oletote) otete|-fevetere ele) oforeletesetel ore tetaleeteletsRel el etet=!= let atetetetatehet= teers el =tetetelet feta tate test tet eet 68 
11-15, Dorsal, posterior, ventral, side and anterior views, <1, holotype USNM 200822; 16-18, Side, anterior, and dorsal views, 
1.5, of the holotype. [Compare with Gypidula cornuta Fenton and Fenton below, figs. 66—73.] 
Independence Formation, locality 526t. 
Cupularostrum? istevensomiMew SPECIES: ...y6.6.c.m 5 ccsierepe ev ee © ede ie (ole, oNo\e wieie/ ei esiescichel 8 elerenetelalsicie «eles eleteheispeetexsic tte ate cette eae 69 
19-23, Dorsal, anterior, side, posterior and ventral views, x1, holotype USNM 200896a; 24-28, Dorsal, ventral, posterior, side 
and anterior views of a complete specimen, x1, paratype USNM 200896b; 29-31, Ventral, anterior and side views, x2, paratype. 
Rhodes Canyon Formation, locality 523v. 
32-36, Anterior, side, posterior, dorsal and ventral views of a small specimen, x1, paratype USNM 201029a; 37-39, Anterior, 
side and dorsal views of the same specimen, <2; 40-44, Side, dorsal, ventral, anterior and posterior views of a large specimen, 
x1, paratype USNM 201029b. 
Rhodes Canyon Formation, locality 509u. 
Cupularostrum SPECIES. We sioyecraielacne oes cteisl ere) evevede Leleie el ial shel ess) ais sieveneteltstheyeteket tere teic eo artittaeteie keboldeha ie eee ae eet eee ene 70 
Posterior, anterior, side, dorsal and ventral views, <1, figured specimen USNM 200895. 
Onate Formation, locality 523d. 


S7iAthabaschia? Species ile ese aeccsty cus fee Bretears ess a eaeee aie vs ANE Sa wge vee whee wwe nse ded etme caste etal e ere cope ohe Sere oe oee Se Ce Pate oe 70 


50-54, Ventral, posterior, anterior, side and dorsal views of a complete specimen, 1, figured specimen USNM 200914a; 55-57, 
Dorsal, side and anterior views, <2, of the same specimen. 

Sly Gap Formation (Thomasaria Zone), locality 509f. 
bleeds av seanyerelis tenses oubie 6, one Snayn @ alesatbpawevatane (6 lle eleyavaiianss See fees, al CM SEEN STR SECS, SSeS hae 77 
58-62, Posterior, anterior, side, dorsal and ventral views, <1, figured specimen USNM 200894; 63-65, Dorsal, side, and anterior 
views of the same specimen, 2. 

Sly Gap Formation, locality SO9f. 
Gypidula‘cornuta Fentonand! Renton’, «.2/..:2/<y2svepecsenctever acess nie a saeteychenstereteme eseueren seus choles i tefeyetereenenore oetenate: olclefeeeeeekeee aaeeae 67 
66-70, Posterior, anterior, side, dorsal and ventral views of a well-preserved specimen, x1, hypotype USNM 200821; 71-73, 
Dorsal, anterior and side views of the same specimen, x1.5. 

Independence Formation, locality 526t. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 14 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 15 


Figure 
1-20. 


21-28. 


29-38. 


39-56. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 171 


EXPLANATION OF PLATE 15 


Ripidiorhynchus newmexicoense new species. 
1-S, Dorsal, anterior, side, posterior and ventral views, <1, paratype USNM 204952. 

Contadero Formation, locality 528a. 
6-10, Side, dorsal, ventral, posterior and anterior views of a large specimen, <1, paratype USNM 200892. 

Contadero Formation, locality 526g. 
11-15, Dorsal, anterior, posterior, ventral and side views, x1, holotype USNM 200920a; 16-20, Ventral, posterior, side, anterior 
and dorsal views of a large specimen, 1, paratype USNM 200920e. 

Contadero Formation, locality 509t. 
Morphorh ynchusivaricostatumapmeweSPeCles sassy svercy-y-0syoevsispecevereherstshelecsyodsy yale letdencuaberaisvedecoveisteysiais sSeyerensiete tate toleperenel eteteeerete re toler eer dente: 72 
21-25, Dorsal, posterior, side, anterior and ventral views, <1, holotype USNM 200922a; 26-28, Anterior, dorsal and side views 
of the holotype, x2. 

Onate Formation, locality 522f. 
Ey pOthyriGinAaySPECLES 2 ce yayscver-verete cto set-toteteVe) ayes ovene ic ccssVereneie cee ve t= vero assis lene eae (orev enaye ie mushe cisieseialehsicrsqereke olelenteaehere corner MrT Teena: 73 
29-33, Anterior, posterior, side, dorsal and ventral views of a small imperfect specimen, 1, figured specimen USNM 201090b; 
34-38, Anterior, posterior, side, dorsal and ventral views of a larger specimen, x1, figured specimen USNM 201090a. 

Onate Formation, locality 523d. 
Hypothyridina emmonsi rotunda new subspecies. ............. 00. cece eee ee e eee t ee ee eee e teeta ee eee 72 
39-41, Dorsal, side and anterior views of a small specimen, x1, paratype USNM 200947c; 42-46, Ventral, anterior, posterior, 
side and dorsal views of another specimen, x1, holotype USNM 200947b. 

Sly Gap Formation, locality 506v. 
47-51, Side, anterior, dorsal, ventral and posterior views of a slender, young specimen, <1, paratype USNM 200950c. 

Sly Gap Formation, locality 509g. 
52-56, Dorsal, anterior, side, posterior and ventral views of a very large specimen, 1, paratype USNM 200946. 

Sly Gap Formation, locality 52le. 


172 


Figure 
1-6. 


49-58. 


BULLETIN 315 


EXPLANATION OF PLATE 16 


Page 


Hypothyridina!emmonsi( rotunda new SUbSPeClesen eryertery-rereirateleretetareratelener el etete tat ate\elatet etatete lela cteyetere Ts ae etete) ates fete ete tate eet tele ete 
1-3, Ventral, side and anterior views, <1, paratype USNM 200948b; 4-6, Side, anterior and dorsal views of another specimen, 
x1, paratype USNM 200949a. 

Sly Gap Formation (upper), locality 506c. 


. Athabaschia? species: tls. ac. syesiecigvere sat eus.svere-oosset pevaveneve rete: efess (6100 ny oytssa arayeroyayieie johabeieiens « ccrafetevcheneteere tag eaei rial peverat cet kveiete kee eee eee 


7-10, Side, anterior, dorsal and ventral views of a complete specimen, x1, figured specimen USNM 203525; 11, Dorsal view of 
the same specimen, x2. 
Sly Gap Formation, locality 524u. 


- Hadrorhynchia‘alta new Species’ <.:2).< <s.cfure ssocte eee ie «eee ee eae ee isis susie eietsre stele’ oda [else esere els ciecltis ine sine Sit alee ie Rael 


12-16, Anterior, side, posterior, ventral and dorsal views of a large individual, x1, paratype USNM 201139a. 

Onate Formation, locality 523b. 
17-21, Posterior, dorsal, anterior, ventral and side views, <1, holotype USNM 201140b; 22, Left side of ventral valve of the 
holotype, showing the fine linear capillae, x3; 23-27, Side, dorsal, posterior, anterior and ventral views, x1, paratype USNM 
201140a; 28, Anterior of the same paratype, showing peculiar arrangement of costae, <2. 

Onate Formation, locality 523d. p 
29-31, Dorsal, side and anterior views of a young individual, x1, paratype USNM 203526. 

Onate Formation, locality 522f. 


. Drifidorostellum obscurum New SPCCIES=; <)-.-).:2721 eree,-elapene: a:eve +c ekovev aves eve rete potene reyes sec] oketaveleoeleerenet eee let svete father alee eee eke ee 


32-36, Dorsal, posterior, ventral, anterior and side views, x1, paratype USNM 200980; 37, 38, Side and anterior views of the 
same paratype, x2. 

Percha Formation (Box Member), locality 5070. 
39_43, Dorsal, posterior, anterior, ventral and side views, <1, holotype USNM 200981; 44, 45, Side and anterior views of the 
holotype, 2. 

Percha Formation (Box Member), locality 3029. 
46-48, Anterior, side and dorsal views of a young specimen, x2, paratype USNM 201154. 

Percha Formation (Box Member), locality 3042s. 
Colophragma ellipticum Mew Species. oi. 5 o- ec.5  sisst oie cca nol cele oie eis oe ¥ eres ereleieie ©) iis oie ol =tcie leis ove eked etee tReet Recetas 
49-53, Side, anterior, posterior, ventral and dorsal views, x1, holotype USNM 200921. 

Contadero Formation, locality 5240. 
54-58, Posterior, side, dorsal, ventral and anterior views, <1, paratype USNM 203537. 

Contadero Formation, locality 528b. 


72 


70 


74 


78 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 16 


ULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 17 


Figure 
1=3" 


4-19. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 173 


EXPLANATION OF PLATE 17 


Colophragmayellipticum! Ne wiSPECIES >, ices. s.ere os co weve aisyey sso covered ars Savers epeces cree wieieiey sel “ie/orysrenetelese) refers eis isre as cisrelereeweisloleekeyel steiner 77 
Dorsal, anterior and side views, «1.5, holotype USNM 200921. 

Contadero Formation, locality 5240. 
‘Coeloterorhynchusmagnificum NEW SPECIES. © 2 ses, asic cre wie o whore 4 6) sis me ops s:apetere vee) 414 rere Grell al ieee ete. des sie0e Gps) sl oncneue el siehenehene icine « 76 
4-6, Ventral, side and anterior views of a very young specimen, <1, paratype USNM 200915b: 7-9, Side, ventral and anterior 
views of another young specimen larger than the preceding, x1, paratype USNM 200915c; 10-14, Side, ventral, anterior, dorsal 
and posterior views of a large adult with one costa in sulcus, x1, paratype USNM 200913. 

Sly Gap Formation, locality 506c. 
15-19, Anterior, side, posterior, dorsal and ventral views of a large spreading specimen, x1, paratype USNM 200912. 

Sly Gap Formation, locality 52Ip. 


PH Aadrorhynchiapmclareni sme Ww SPCCLESe, oojc.s «ace: vzspvysuecesevss cee epsuezs (ove sees 2 eusyorsue  susyo,opeisisl a,avepa, a eusyacete  exentuopeleyeheyascicks pega tetente eacaeesvete elses 75 


20-24, Ventral, anterior, posterior, dorsal and side views of a well-preserved individual, x1, paratype USNM 200903. 

Onate Formation, locality 523h. 
25-29, Dorsal, posterior, anterior, side and ventral views of a youthful individual in which the costae have not yet formed, x1, 
paratype USNM 200897b; 30, Anterior view of a large specimen showing peculiar costae, x1, paratype USNM 200897a. 

Onate Formation, locality 523b. 
31, Posterior of a large specimen showing long flattened crura, x2, paratype USNM 200918. 

Onate Formation, locality 523k. 
32-36, Posterior, dorsal, side, ventral and anterior views of a large specimen, 1, paratype USNM 200891d; 37-41, Dorsal, 
posterior, anterior, side, and ventral views of a large individual, x1, holotype USNM 200891a; 42, Anterior view of a large 
specimen showing linear depression in costae, x2, paratype USNM 200891e; 43, Exterior of a crushed specimen showing capillae 
on the ventral valve, x4, paratype USNM 200891g. 

Onate Formation, locality 523d. 


3-7. 


8-17. 


18-22. 


23-27. 


BULLETIN 315 


EXPLANATION OF PLATE 18 


Dorsal and side views of an imperfect specimen, x1, figured specimen USNM 200952. 

Thoroughgood Formation, locality 522. 
Colophragmatellipticum'Me Ww SPECIES sap rayerey pore r sero tete ate) irl e evoyeel= Yayo rela clones ltecetore eletei-lsvainietataleelspotelstelstetet-telct-\ foe reteset 77 
Anterior, ventral, side, posterior and dorsal views, x1, paratype USNM 200953. 

Contadero Formation, locality 5240. 
Hadrorhynchiajaltainew: Species vrata: ayar = jeteeg 10: < aes re was celeyevevase epeietenevasele «1 overs a ateverova telat etepeystayelsletere?=foreiel ai et=tete cso et terete 74 
8-11, Dorsal, ventral, side and anterior views of a complete young specimen, <1, paratype USNM 200898a; 12, Dorsal view of 
the same specimen, 2; 13-16, Ventral, side, dorsal, and anterior views of another youthful specimen, x1, paratype USNM 
200898b; 17, Dorsal view of the same specimen, x2. 

Onate Formation, locality 3025c. 
Camarotoechia? Species: pis Sayters co ciecase tices usta e010 ee acs ap See wee tat eee crake eye reise evecare eo cre euetGhevers Tee NeTC, reTCR See oC TR OEA OE ae 79 
18-20, Dorsal, side and anterior views, <1, figured specimen USNM 203506; 21, 22, Ventral and dorsal views of the same 
specimen, x2. 

Percha Formation (Box Membér), locality 509b. 
Calvinariabransoni:Stain brooks, <2 .5.cc cei. s.2ssiecsrzys. is eis eilare ons overs sqenessueol aie aoe Susi ¥.0 cm ebelenebelecereeveieretea.+ «veils ©. SI oRe ern ener R CES eee 80 
Posterior, anterior, ventral, side and dorsal views of a large specimen, x1, hypotype USNM 200858. 

Sly Gap Formation (lower), locality 521k. 


 oparaphorhynchusimelareniSantenacr erties rise ile siete iets icteric terisieieeiersisiel=t sinter ede stalelele) tet sretatee aerial eteieletiaht ec heeeeeeete 83 


Anterior, side and dorsal views of an imperfect specimen, x1, hypotype USNM 201091. 
Percha Formation (Box Member), locality 3029. 


- (Coeloterorhynchus; magnificum) NEW SPECIESS x. os o:< wie onic ieusvers ciciarelc's pint uavors old Seer tel veto eeiete ararere eieke aera eee ieee ries 76 


31-35, Dorsal, ventral, anterior, side and posterior views of a young specimen, x1, paratype USNM 200915i; 36-40, Dorsal, 
anterior, posterior, side and ventral views of a large specimen, x1, paratype USNM 200915k; 41-45, Dorsal, side, anterior, 
posterior and ventral views, x1, holotype USNM 2009151; 46, 47, Ventral interior views, showing muscle field and pallial marks, 
xl and x2, paratype USNM 200915g. 

Sly Gap Formation (upper), locality 506c. 
48-52, Posterior, dorsal, ventral, anterior and side views, <1, paratype USNM 200916. 

Sly Gap Formation, locality 506). 
53-57, Posterior, anterior, dorsal, ventral, and side views of another specimen, x1, paratype USNM 200917a. 

Sly Gap Formation, locality 506i. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 18 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 19 


Figure 
1-6. 


7-10. 


51-65. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DuTRO LES 


EXPLANATION OF PLATE 19 


FE VANESCIFOSET UIT SP CCLES cy sysp ts coke opel s ozs cis everensy Scher eae atta Re To ee oT on UREN eS IIIS Ee TeTE IS orien bars st Sus tose a ele rote ye ees epeeES bes 83 
Dorsal, anterior, side, posterior, anterior tilted and ventral views, <1, figured specimen USNM 201042. 

Thoroughgood Formation, locality 522. 
Eyanescirostrum|glabrayentrumi (Stain brooks) .)727-9.¢5 5. u</-0=1<'s) eps svek eins cca eee eters ss ore. «tole lel ai sauna eee ate tese cciereia ate = eae CE ee nereere 83 
7-9, Anterior, dorsal and side views of a somewhat flattened individual, x1, paratype USNM 135259a; 10, Posterior view of an 
interior filling showing median septum, 2, paratype USNM 135259b. 

**Percha”’ Formation of Stainbrook = Rhodes Canyon Formation, locality 528c. 


Pea LelOrhynCchus:sSpRACLicuM NEW SPECIES., = <)<yare-=, -ne.9 sveve\osei0 SyeTaTGy ES Si o/ovors¥ersresese Si sieisieve ove re lelavcvoheve ere cicveh oierare evs caval tote overe ete rabe eee eveleh sis 81 


11-15, Anterior, dorsal, posterior, side and ventral views, x1, paratype USNM 200893. 
Onate Formation, locality 523e. 

16-20, Posterior, anterior, dorsal, side and ventral views, x1, holotype USNM 200851; 21, Anterior view of the holotype, x2. 
Onate Formation, locality 5221. 


me Cal vinariayDransOmils tain DLOOK sy cc -veraye eveveicisic role yore olor Tore em ieVele a eitraneee nie ie aie sleeve Glele PRINS cas OTS TEE ohio tee eect non care 80 


22-26, Anterior, side, ventral, dorsal and posterior views of a large individual with a single costa in the sulcus, x1, hypotype 
USNM 200856d; 27-30, Ventral, dorsal, side, and anterior views of a specimen with two costae in the sulcus, x1, hypotype 
USNM 200856b. 

Sly Gap Formation, locality 509h. 
31-35, Ventral, posterior, side, anterior and dorsal views of a wide individual, x1, hypotype USNM 200860; 36, Dorsal view of 
same specimen, 1.5. 

Sly Gap Formation, locality 526k. 
37-39, Side, anterior, and dorsal views of a specimen with two costae in the sulcus, x1, hypotype USNM 20086le; 40, Anterior 
of another hypotype with one costa in the sulcus, x1, USNM 20086la. 

Sly Gap Formation, locality 509h. 
41-45, Posterior, dorsal, anterior, ventral and side views of a young specimen, x1, hypotype USNM 200857. 

Sly Gap Formation, locality 52le. 
46-50, Posterior, anterior, side, ventral and dorsal views of an imperfect specimen, x1, hypotype USNM 200859. 

Hackberry Formation (Cerro Gordo Member), locality 528d. 
Camarotoechia\latisinuatalbmew :SPeECIeSs, mreyeceveve-srstere: cielo Vernie cynicra te eke kd eve ieee Laer cTo\e e lei) saree ule RoR erate areeeleye eee ei a ares eet abere 79 
51-55, Dorsal, posterior, anterior, side and ventral views, x1, holotype USNM 200945a; 56-60, Anterior, posterior, ventral, 
side and dorsal views, 1, paratype USNM 200945b; 61-63, Dorsal, side and anterior views of a young individual, x1, paratype 
USNM 200945c; 64, 65, Posterior views of two specimens showing imprints of the cardinalia, both x2, paratypes USNM 200945d, e. 

Thoroughgood Formation, locality 522. 


Figure 


1-21. 


39-48. 


49. 


BULLETIN 315 


EXPLANATION OF PLATE 20 


Petasmaria patens NeW SPECIES... occcosus-eiseocs:eraite/esaitesn cotnfouenctceevayetteys cision, a+ io /sleyenla tol efoto aleueaisboveietope keke Naieisetoiaeta fase ea ket ea ee 85 
1-5, Anterior, ventral, side, posterior and dorsal views, <1, holotype USNM 200937. 

Percha Formation (Box Member), locality 507h. 
6-10, Anterior, dorsal, posterior, side and ventral views, x1, paratype USNM 200939. 

Percha Formation (Box Member), locality 5070. 
11-15, Dorsal, posterior, ventral, side and anterior views of an unusually folded specimen, x1, paratype USNM 200938. 

Percha Formation (Box Member), locality 507c. 
16-20, Anterior, dorsal, side, posterior and ventral views, <1, paratype USNM 200943a; 21, Enlargement of part of the dorsal 
valve, showing linear ornament, <3, paratype USNM 200943b. 

Percha Formation (Box Member), locality 507d. 


-s Petasmaria compacta ne Wi SPECIES |= scs-1.vercte e.ccac<fousnsesv tee s0so) avaiiayove esi2hoioyevs7 sas <nencvohecvereweketed Totes fatayetelel tetas oratoie Teneo tia EE ee aes 84 


22-26, Anterior, ventral, posterior, dorsal and side views, <1, holotype USNM 200942a; 27-29, Ventral, anterior and side views 
of the holotype, x2; 30-32, Side, anterior and dorsal views, x2, paratype USNM 200942b. 
Percha Formation (Box Member), locality 5070. 


, Hyborhynchella\bransoni (Coopers csr.) <scpes oe ct shen ocean oie ete, oe) o(edcois ede etean) oj our al oleyeke etekebseey exe ele (ebsiel skate ie sterateeleteisl ole eee eae hen 82 


33, Dorsal view, x1, hypotype USNM 201063; 34-38, Anterior, posterior, side, dorsal and ventral views of the same hypotype, 
x2. 

Sly Gap Formation, locality 302Sa. 
Platyterorhynchus’obscurum’ NeW. SPECIES wo: i:5)5.<j215 55 seus eiagets eucnoses > wiainy slay ebe lenge: svar yeresavevaustabejccerer eters) ebaps)'s eneteratets terete toler tee tetera ements 81 
39_43, Anterior, posterior, ventral, side and dorsal views, x1, holotype USNM 200877; 44, 45, Cardinalia of the dorsal valve, 
xl and x2, paratype USNM 200878d; 46, 47, Interior views of the ventral valve, x1 and x2, paratype USNM 200878a; 48, 
Interior of a complete specimen showing the long, distally flattened crura, x2, paratype USNM 200840. 

Onate Formation, locality 526p = 3013. 
Paurorhyncha’ cooperi (Stainbrook)).. <. 5.:<<:2)5)s)sisre::2 2 ececavostco. c=: austatiote apy slevnze jeeie easy eyniel eke aus syste fereys ‘abei=fesaisss)ofeishetetetenelele teeaebetele eters 82 
Interior view of both valves showing cardinalia and short dental plates, x2, hypotype USNM 200815. 

Percha Formation (Box Member), locality 506b. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 20 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 21 


14-31. 


32-37. 


38-53. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO a7, 


EXPLANATION OF PLATE 21 


Page 
se betasmaniaymulticostatammewispeClese, fer miese renee ricer ries cieveicic oi cielcketasieustchersiehvelt ieee eiclele lai tevele slat itentet ree tenetenst= rier 84 
Anterior, side, dorsal, ventral and posterior views, <1, holotype USNM 200940. 
Percha Formation (Box Member), locality 506b. 
> LATEST AIA PPETONS WOW GeeSIEs gooousscaapeudsaccoun > s anesoespoasnadoDCOGddoc so0Gemo DOOD O DUR GOddOGoconeESoOoDODEDICODE 85 


6-10, Ventral, anterior, side, dorsal and posterior views of a young specimen, 1, paratype USNM 200941; 11-13, Anterior, side 
and dorsal views of same specimen, 1.5. 

Percha Formation (Box Member), locality 509. 
Hy borhynchellajbransoni| COOpelemerc etic eyekiee eet ie tere eer rte teicher det sie tote ole) eee ere tet feted Lele ie rae Peete eet tee et Rear 82 
14, Dorsal view of a large specimen, x1, hypotype USNM 201064a; 15-19, Anterior, posterior, ventral, side and dorsal views of 
the same hypotype, x2. 

Sly Gap Formation, locality 524m. 
20, Dorsal view, x1, hypotype USNM 201062; 21-25, Anterior, posterior, side, ventral and dorsal views of same hypotype, x2. 

Sly Gap Formation, locality 3045h. 
26, Dorsal view, x1, hypotype USNM 201069; 27-31, Anterior, side, posterior, dorsal and ventral views of the same hypotype, 
x2. 

Sly Gap Formation, locality 509h. 
ACE SET? Gaa6IGS 2, Soascovenccvcvnnadguseandaosoounon ooUODaRdomundds00sDUUSDDoUN OU UOEUUEUGODUdedCooSOUOsDDCONOGD 71 
32-34, Anterior, side and dorsal views of a small specimen, x1, figured specimen USNM 201041a; 35-37, Anterior, dorsal and 
side views of a larger individual, x1, figured specimen USNM 201041b. 

Thoroughgood Formation, locality 522. 
AGATA EDD WOW GISTs. aavonceancoososcncoencsncoooDencosceg DUD soTIN DON IDODGOOSHUODSOOuESSIGOD I HOCODSONEAe DOE 86 
38-42, Ventral, dorsal, side, posterior and anterior views of a specimen showing part of its frills, x1, paratype USNM 201005a; 
43-47, Posterior, side, dorsal, ventral and anterior views, <1, paratype USNM 201005b; 48, Ventral view of the same specimen 
showing details of the ornament, <1.5; 49-53, Dorsal, ventral, side, anterior and posterior views of a specimen preserving some 
of its frills, x1, holotype USNM 201021. 

Onate Formation, locality 5230. 


178 


Figure 
1-16. 


17-38. 


BULLETIN 315 


EXPLANATION OF PLATE 22 


Pseudoatrypa\devomiamal(WeEDStEL) ir reye rm tcicte ose ieee ene hele retad olde er natedete rebel ate ekefapa elena sheeted tel ley =) = ake els steteete) ots iets eee 87 
1-5, Ventral, dorsal, anterior, posterior and side views, x1, hypotype USNM 200965. 

Sly Gap Formation, locality 509s. 
6-10, Ventral, anterior, posterior, side and dorsal views of a small specimen, x1, hypotype USNM 200927. 

Sly Gap Formation, locality 521m. 
11-15, Anterior, posterior, dorsal, side and ventral views of a large individual, x1, hypotype USNM 200944; 16, Rubber replica 
of the exterior of a large specimen showing costation, x1, hypotype USNM 201094. 

Sly Gap Formation, locality 521d. 
Desquamatialcostata Dew SpeCleSs eyiertey-l-fteienre ol ioeis “sever icralerelvevehetone airs e otereloletetatefateloteketatat= fetal sleet =tel «lett =h= lets ets tate eee teen 86 
17-21, Dorsal, posterior, anterior, ventral and side views of a large specimen, x1, paratype USNM 200973. 

Onate Formation (Bed E = Bed with Sphaerospongia cf. S. tesselata Phillips (1841)), locality 522g (=522f). 
22-26, Posterior, anterior, ventral, side and dorsal views of another adult, x1, paratype USNM 201007a. 

Onate Formation, locality 5230. 
27-29, Dorsal, side and anterior views of a large obese specimen, 1, paratype USNM 201006; 30-32, Ventral, anterior and dorsal 
views of a specimen with injured anterior margin, x1, paratype USNM 201003. 

Onate Formation, locality 528e. 
33-37, Anterior, posterior, ventral, side and dorsal views of a well-formed individual, x1, holotype USNM 200972; 38, Ventral 
view of the holotype, enlarged to show the ornament, x1.5. 

Onate Formation, locality 523d. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 22 


st 
) uy 
, anh 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 23 


Figure 
1-6. 


7-31. 


39-56. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 179 


EXPLANATION OF PLATE 23 


18 Rf ELC d CMTE XX ess ieee eee eto pete Oiaiaie ORIeeec 6 ora oC G. cioieateiess a cicresctro 5 GARG WA GIO EE ao Oe an EE Gm cnc coca cee 73 
1, Dorsal view, 1, figured specimen USNM 201141; 2-6, Dorsal, posterior, side, anterior and ventral views of the same specimen, 
ape 

Onate Formation, locality 526c. 
Costatrypabvaricostatay (Stain DLOOK) eget cyocs.ccco) 2 sic nats areas tetetsi eer eilexo) ovary Siete eetorehe ey sniorel s Deke REIN OICIE Gere l Ena 88 
7-11, Side, anterior, dorsal, ventral and posterior views, <1, hypotype USNM 200992. 

Sly Gap Formation, locality 506c. 
12-16, Dorsal, side, posterior, ventral and anterior views of a sharply-folded specimen, <1, hypotype USNM 200996. 

Sly Gap Formation, locality 521d. 
17-21, Ventral, anterior, posterior, dorsal and side views of a large individual, x1, hypotype USNM 200963; 22, Ventral view of 
a small specimen, <1, hypotype USNM 200979a; 23-27, Ventral, anterior, posterior, side and dorsal views of an imperfect 
specimen, <1, hypotype USNM 200979b; 28, Ventral view of the same specimen, x1.5; 29, 30, Ventral views of a large specimen 
preserving the frills, x1 and x1.5, hypotype USNM 201004. 

Sly Gap Formation, locality 3054b. 
31, Ventral view of a small specimen preserving its frills, x 1.5, hypotype USNM 200994. 

Sly Gap Formation, locality 509p. 


mLseudoatrypardeyomianial (W.EDSUEL) =. says ears 1-12 fete Merete onctncee te evr Meret T ee covey sped PRCT re teres AER one Tene cot ATEN OTE ROR Nas 87 


32-36, Posterior, anterior, dorsal, side and ventral views of a large specimen, <1, hypotype USNM 200964. 

Sly Gap Formation, locality 509z. 
37, 38, Ventral views showing frills, <1 and «1.5, hypotype USNM 200966. 

Sly Gap Formation, locality 506c. 
Costa thy pakext EnSaen EW sSDECIES eters -nenevay= cere seek revere ach eateyer eae enacts crates ce eeP ae Ree cos ener eoeioee Ree tsratebe 1s RRR cro 87 
39, Ventral views with frill partially preserved, <1, paratype USNM 200967. 

Sly Gap Formation, locality 506). 
40-44, Dorsal, side, posterior, anterior and ventral views of a large specimen, x1, holotype USNM 200984. 

Sly Gap Formation, locality 521k. 
45-49, Posterior, side, anterior, ventral and dorsal views of a small individual, x1, paratype USNM 200951la; 50, Ventral view 
of the same specimen, 1.5; 51-55, Posterior, anterior, dorsal, ventral and side views of a specimen preserving some of the frills, 
x1, paratype USNM 200951b; 56, Ventral view of the same specimen, x1.5. 

Sly Gap Formation, locality 3054. 


Figure 


BULLETIN 315 


EXPLANATION OF PLATE 24 


1-12... Costatrypa‘extensa New SPECIES. 655 .<.<51<evejele cpoveisielestrs oie eje lore: d)s neve ie e-vininyme johns ele 1 apstebelatelosesckeae =i a/eies0l hots eRe ete ete ae nen 


Ww 
Ig 


46-60. 


1-5, Ventral, anterior, side, posterior and dorsal views of a large specimen, <1, paratype USNM 200993; 6-10, Dorsal, anterior, 
side, ventral and posterior views of another specimen, x1, paratype USNM 200983. 

Sly Gap Formation, locality 521k. 
11, 12, Ventral and dorsal views, x1, paratype USNM 200960a. 

Sly Gap Formation, locality 521d. 


= Jowatrypa americana’ (Stainbrook)% ((oeicryernte sce cereals ore sim open el st ia) sie eile) lel) ele etal ee) ataloke tote teroie talento eke eae eee 


Dorsal and side views, for comparision with /. rara, new species, <2, hypotype USNM 201134a. 
Independence Formation, locality 528f. 


: Nowatrypa Fara New SPECIES oe oq 2: cie,<s sys, ocetstarsfatesete ie] oscjo'e) ais 6/0 aay slniosecefereieteneye otorol/ensierteuetes ened fel ateke =sieta/1 tolerates eee eee ea 


15-19, Dorsal, ventral, anterior, posterior and side views, x1, paratype USNM 201099; 20, 21, Dorsal and side views of the same 
paratype, x2; 22, 23, Dorsal and side views of another individual, x2, paratype USNM 201135a. 
Sly Gap Formation, locality 506). 
24-28, Posterior, anterior, side, dorsal and ventral views, <1, holotype USNM 201098; 29, Dorsal view of the holotype, x2. 
Sly Gap Formation, locality 521q. 


. Spinatrypa compactaime ws SPCCles sierra tela lereteeaeatey ss aiay esas = keke edeloNetene teeta) ai-Weatateoleter ates Neneteiefeietenstersiotereiat leis eitel (elect ett ee eee 


30-34, Side, posterior, anterior, dorsal and ventral views, <1, holotype USNM 203538; 35, Ventral view of the holotype, x2. 
Contadero Formation, locality 528g. 
36-40, Anterior, side, ventral, dorsal and posterior views of a well-preserved adult, «1, paratype USNM 203540a; 41-45, Posterior, 
dorsal, anterior, side and ventral views, <1, paratype USNM 203540b. 
Contadero Formation, locality 523n. 
Spinatrypa ‘obsolescens new Species. = cecicteieiciie mie cisco 3 5is10 = lets c elerere aie wie eieiey « neve sye¥e el oleFeRe iets > oie 2 aie te ae cee ee eee 
46-50, Ventral, side, dorsal, anterior and posterior views, <1, holotype USNM 200962; 51, Ventral view of the holotype, x1.5. 
Contadero Formation, locality 526g. 
52-55, Ventral, dorsal, anterior and side views, x1, paratype USNM 200997a; 56-60, Anterior, dorsal, side, posterior and ventral 
views of a large specimen, x1, paratype USNM 200997d. 
Contadero Formation, locality 3054k. 


89 


89 


89 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 24 


PLATE 25 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


Figure 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 181 


EXPLANATION OF PLATE 25 


IPE mo pinatry parerullay(StaimbirOO ky) json. jeuenstsse, ave eusyssske sues dosed chee ueore va eronsrexaveteeuwiere eleife ay a ear easie ose seve seksi ee eee ee 90 


3-40. 


Dorsal views of two large specimens showing spines, 1, hypotypes USNM 201082a, b. 
Independence Formation, locality 528h. 
Spinatrypaltrullardecorticataimew: SUDSPECIES ty + .<.<. 5. «eis \ec. espe eral teres aio et Mey crake Toler ale aia Seal SPT Vn aE ees cbs Sree Pe oT eel te 90 
3-7, Dorsal, ventral, side, anterior and posterior views of a large, strongly folded specimen, <1, paratype USNM 200999. 
Sly Gap Formation, locality 524u. 
8-12, Ventral, posterior, dorsal, anterior and side views of a specimen with obsolescent costae, 1, paratype USNM 201159. 
Sly Gap Formation, locality 506i. 
13-17, Dorsal, anterior, posterior, ventral and side views of a large specimen, x1, paratype USNM 200982b; 18-22, Side, ventral, 
anterior, dorsal and posterior views of an immature specimen, x1, paratype USNM 200982d; 23-27, Anterior, ventral, side, 
posterior and dorsal views of a young adult, x1, paratype USNM 200982f. 
Sly Gap Formation, locality 506c. 
28-32, Anterior, posterior, side, ventral and dorsal views of a thick adult, x1, paratype USNM 200931; 33, Ventral view of the 
same paratype, x2. 
Sly Gap Formation, locality 509y. 
34-38, Anterior, posterior, ventral, side and dorsal views of a large adult, x1, holotype USNM 200998. 
Sly Gap Formation, locality 509j. 
39, 40, Dorsal and side views of an injured adult, 1, paratype USNM 201002. 
Sly Gap Formation, locality 506). 


Figure 


1-6. 


19-24. 


52-66. 


BULLETIN 315 


EXPLANATION OF PLATE 26 


Emanuela: SpeGieSss (Sive:.c 202 s/sccve; tne sre choca petetelave overetoyeus ie) oes ich ei susvase lace (eyeighete taleleesisy eteimbous eis Sutetsle Slee oF NMI a eRe eae ee 


1, Dorsal view, <1, figured specimen USNM 203559; 2-6, Posterior, anterior, dorsal, side and ventral views of the same individual, 
9 


XL. 


Onate Formation, locality 528i. 


~ AmmbocoeliavSpeGies) We. scsi. .)c)=.2, sete sensisvofe,ap20deaiase1sistadeneysyater aie elisdey exe 218s ig) oye: <5.) oS arreystopetoye toyeneie che ve te teyahe = lepcloto-c> 1 1 een ae ee 


7, Dorsal view, <1, figured specimen USNM 201073b; 8-12, Posterior, anterior, side, ventral and dorsal views of the same 
specimen, 2: 13, Dorsal view, x1, figured specimen USNM 201073a; 14-18, Anterior, posterior, side, dorsal, and ventral views 
of the same specimen, x2. 
Sly Gap Formation, locality 5061. 
Ambocoelia capillatai mew SpeCleSs 2... ejare cere. voyage co ce-s cis) fe eucie ecko sxe eicyeresc¥olais’ » wis) srageicicheiata sbeleperoveuarsieisiere fohots steseyeretets aero tae eae 
19-23, Anterior, posterior, side, ventral and dorsal views, x4, holotype USNM 201136; 24, Ventral view of the holotype, x6. 
Percha Formation (Box Member), locality 509b. 


25, Dorsal view, x1, figured specimen USNM 201074; 26-30, Anterior, posterior, dorsal, side and ventral views of the same 
specimen, X2. 
Contadero Formation, locality 526g. 


: Crinisarina reticulata:(Stainbrook) =< j<. se). 5 case. ccere tere, oco spare: oy ouar e-cter cys arabe eeereeMapa wieneseyo: ace. cra: <pelereye’srona vere cane rates ee ee 


31-35, Anterior, posterior, side, ventral and dorsal views of a small individual, x1, hypotype USNM 203539a; 36, Enlarged area 
in sulcus showing spines, x4, hypotype USNM 203539b. 
Rhodes Canyon Formation, locality 528). 
37-41, Side, anterior, posterior, dorsal and ventral views of a large individual, x1, hypotype USNM 200925. 
Rhodes Canyon Formation, locality 521g. 
42-46, Side, dorsal, ventral, anterior and posterior views of a large adult, x1, hypotype USNM 200928. 
Rhodes Canyon Formation, locality 523v. 
47-51, Ventral, side, posterior, anterior and dorsal views, x1, hypotype USNM 200974. 
Thoroughgood Formation, locality 522. 
Composita bellula ‘Stainbrook. ojo coc ce sie careers se rere cies sper 6 oe wpa SIGs IS BE So S(ore Sele wFeLEE Is) © ore sysraie see reyelel ets eI Tee ERR 
52-56, Anterior, ventral, posterior, dorsal and side views of a large individual, x1, hypotype USNM 201037. 
Percha Formation (Box Member), locality 527d. 
57-59, Side, anterior and dorsal views of a narrow specimen, x1, hypotype USNM 201038b. 
Percha Formation (Box Member), locality 507c. 
60-64, Anterior, dorsal, side, ventral and posterior views of a large symmetrical specimen, x1, hypotype USNM 201010. 
Percha Formation (Box Member), locality 3026a. 
65, 66, Ventral views of two specimens showing the spires, x1, hypotypes USNM 203527a, b. 
Percha Formation (Box Member), locality 509c. 


93 


93 


94 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 26 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 27 


Figure 
1-19. 


20-29. 


57-67. 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DuTRO 183 


EXPLANATION OF PLATE 27 


Mucrospinifersparvus ne WwaspeClessmertorre skerete ors ais) ereueserssoiore erect erate te ne cre ie rel tenance elie eee one eee ee Ee eer 97 
1-4, Side, posterior, dorsal and anterior views, <1, holotype USNM 203541; 5-8, Anterior, ventral, side and posterior views of 
the holotype, <2. 
Onate Formation, locality 528k. 
9, Dorsal view of a small individual, x1, paratype USNM 201046a; 10-14, Dorsal, anterior, ventral, posterior and side views of 
the same paratype, x2. 
Onate Formation, locality 526c. 
15-19, Anterior, dorsal, side, posterior and ventral views of a decorticated specimen, <1, paratype USNM 201104. 
Onate Formation (Bed F), locality 524). 
Hehinococlajpretiosa mew SPeCless siete ie toss oe verti crs Terao ee eile ers aca ea eee severe ace teenaeEs avers susie feisieie ie istoe eteeerieineeters 95 
20-24, Dorsal, anterior, posterior, ventral and side views, x2, holotype USNM 201072; 25, Dorsal valve of the holotype, x3; 26, 
Ventral valve of the holotype to show concentric bands, <4. 
Onate Formation, locality 522i. 
27, Interior of the dorsal valve showing cardinalia, x3, paratype USNM 201065. 
Onate Formation, locality 523b. 
28, 29, Dorsal and ventral views of an interior filling showing traces of muscle marks and cardinalia, x2, paratype USNM 201156. 
Onate Formation, locality 522n. 


a HleutherokommarextensagmewsSPECleSs, ictsce,2 isin sewaberereseue sucverstey ete eel les oncsse ane aor meet tes lek Siar SHAT Tene ee Ee eee aes 96 


30-34, Anterior, posterior, side, dorsal and ventral views, «1, holotype USNM 200876a; 35, Dorsal view of the holotype, x2. 
Onate Formation, locality 3052a. 


we hylothyrisinovamexicanay StalnbLOOKs, -/cvre folate eyo tatese stele oles settee chatere QS ieee Oe ee oso se Oe overs Pore Cet ee nane 98 


36-40, Posterior, side, ventral, anterior and dorsal views, x1, hypotype USNM 201060; 41, 42, Dorsal and anterior views of the 
same hypotype showing sharp fold, x2. 

Rhodes Canyon Formation, locality 521g. 
43-47, Dorsal, anterior, side, ventral and posterior views of a large specimen, x1, hypotype USNM 20354Sa; 48, Dorsal view of 
the same specimen, x1.5; 49, 50, Anterior and ventral views of the same specimen, showing angular sulcus and concentric 
lamellae, x2. 

Rhodes Canyon Formation, locality 528}. 


MlylothyrisisulcocostatanDe Ww SPECLES. fers ceicreje1 cies sie ca1s) fe) shes elo) sh) seede/= SinIACi ue ceheenia (es 2iayis aussie nls ocala RISE ere CE eee nee ictore 98 


51-55, Ventral, anterior, side, posterior and dorsal views, x1, paratype USNM 201016; 56, Dorsal view of the same paratype, 
aa2s 

Sly Gap Formation, locality 3054c = S09p. 
hylothyrisicompacta New. SPECLESs veyers cuatarec exeytevoves sy sts eh ounsovo-= olay oney eee Cau eneteneneveraceteoret onesie revetey on cxieysrevars. «eishiayavahaper-rass ena iarnee eecroreneteraie ee 97 
57-61, Dorsal, side, anterior, posterior and ventral views, <1, paratype USNM 200986b; 62-66, Posterior, side, anterior, dorsal 
and ventral views, x1, holotype USNM 200986a; 67, Interior filling of a ventral valve showing dental plates and median septum, 
x1, paratype USNM 200986c. 

Thoroughgood Formation, locality 522. 


184 


Figure 
1-24. 


33-42. 


BULLETIN 315 


EXPLANATION OF PLATE 28 


Tylothyris'sulcocostataimew.SpeECles:, Fryer: -\epeieicsn=. 2122 + cco4es os .=sctere nis oles saekn avn mel evatene en Yeve teva keNeketeteyasal =P fale) si-fets enateroae ene ed 98 
1-5, Anterior, dorsal, posterior, ventral and side views, x1, holotype USNM 201019; 6-9, Posterior, anterior, dorsal and ventral 
views of the holotype, x2. 

Sly Gap Formation, locality 509z. 
10-14, Posterior, side, ventral, dorsal and anterior views, x1, paratype USNM 201020; 15-18, Dorsal, posterior, anterior and 
ventral views of the same paratype, x2. 

Sly Gap Formation, locality 521d. 
19-23, Side, posterior, ventral, anterior and dorsal views of another specimen, <1, paratype USNM 201067c; 24, Dorsal view of 
the same paratype, x2. 

Sly Gap Formation, locality 506j. 


. Lylothyris|novamexicana Stain brooks, 2/35 9.c.sss:s0s 6.2c06 (35% = jayssenocs lousy is idveyesateneuvegn: alntale) ened sianays,wityovasztesebelaievorebere ashe ete el ee eee nesote 98 


25-29, Ventral, dorsal, anterior, posterior and side views of an obese individual, «1, hypotype USNM 201017; 30-32, Ventral, 
dorsal and anterior views of the same hypotype, x2. 

Contadero Formation, locality 523q. 
Acutoria’angulata New sSPECIES |. 0.005 /c. oyster sie 3 fe Yors orceve vin eia = Sicyerey ene sucyecay avers eyeie) ocyevar wets Stele rs isvereieie rele Sees elke efa Ree eee 100 
33-36, Anterior, oblique side, posterior and dorsal views of a dorsal valve showing angular fold, «1, paratype USNM 201127b; 
37, Rubber impression of an incomplete ventral valve showing flattened costae, x1, prepared from paratype USNM 200127e; 38, 
Exterior of the same fragmentary ventral valve, <1; 39, Rubber replica of the exterior of a decorticated ventral valve showing 
the fine capillae on the surface, x1, made from paratype USNM 201127c; 40, 41, Anterior and ventral views of a nearly complete 
ventral valve showing angular sulcus, <1, holotype USNM 201127d; 42, Exterior of a much decorticated ventral valve, x1, 
paratype USNM 201127a. 

Onate Formation, locality 526z. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 28 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 29 


i 4 


, YQ 


Figure 
1-21. 


29-41. 


42-44. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DUTRO 185 
EXPLANATION OF PLATE 29 
Page 
Strophoplearamotabilis) (Kindle) tamevactyee,- <r eri etter ocean felietel sy aysne tte erearatereietetercrsret siereiceeie te helerorelstaratolascresvetarerorenevianstetsis 99 


1, 2, Ventral views of an exceptionally mucronate specimen, <1 and 1.5, hypotype USNM 200847; 3-7, Anterior, dorsal, 
posterior, ventral and side views of a nearly complete individual, x1, hypotype USNM 200848a; 8, Dorsal view of the same 
specimen, <2; 9, Ventral view of an immature, mucronate individual, x1, hypotype USNM 200848b. 

Percha Formation (Box Member), locality 527p. 
10-14, Anterior, dorsal, ventral, posterior and side views of a large specimen, x1, hypotype USNM 201149a; 15, 16, Ventral and 
dorsal views of the same hypotype, x2. 

Percha Formation (Box Member), locality 507h. 
17, 18, Dorsal views of a narrow, stunted specimen, x1 and x2, hypotype USNM 201147a. 

Percha Formation (Box Member), locality 506q. 
19, 20, Ventral views of another stunted and narrow individual, x1 and x2, hypotype USNM 201148a. 

Percha Formation (Box Member), locality 506. 
21, Dorsal view of a mucronate but narrowed dorsal valve, x1, hypotype USNM 201146. 

Percha Formation (Box Member), locality 527a. 


MOVE CH OSD ATTLCLAS PC CLES a race ose tele v chet elo s bse Seo sicaes cr le or Sense ter as Pata sge coef ow Fo epeusanerteieso trea) ay shafebs felelajn ees = obs eueieielobetotac be korsEePen tate ackatasate seeeonete 


22, Exterior of a ventral valve showing flattened costae and capillae in sulcus, x1, USNM 201128a; 23, Rubber replica of the 
exterior of another ventral valve, x1, made from USNM 201128e; 24, Dorsal view of dorsal valve, x1, USNM 201128c; 25, 
Exterior of another dorsal valve showing capillae on the fold, «1, USNM 201128d; 26-28, Posterior, dorsal and side views of a 
large dorsal valve, x1, USNM 201128b. 

Onate Formation, locality 3023c. 
I yloth yriseSDECLESM lh. coyoyscc.o: 3.4 re lovome rene PVs seexelin 8 sv cuc. xt to 6 Gustcneneae potey Meskevars "a elevanss os avsvlats eteeto¥eUe 21 ey tesatede a0 2s coir, cifes cheer eee ode ta eye setae eeen= a 
29-33, Anterior, posterior, ventral, side and dorsal views of an exfoliated specimen, <1, figured specimen USNM 201047. 

Contadero Formation, locality 526g. 
34-38, Posterior, anterior, ventral, side and dorsal views of another damaged specimen, x1, figured specimen USNM 201048; 39, 
Dorsal view of the same specimen, <2; 40, 41, Dorsal views of a specimen preserving the exterior ornament, <1 and x2, figured 
specimen USNM 201049. 

Contadero Formation, locality 509t. 
CyrtospiriferDrevipOsticus! Salo LOO Kegee persis mene rn ee teted eee areata alee eueesyeke teeter te keereay tos ote fake taken ener ater geben tts otenene cee revera eateries 
Side, anterior and dorsal views, «2, holotype USNM 135257. 

Rhodes Canyon Formation, (locality) Rhodes Pass. 


186 


Figure 
1-23. 


24-29. 


35-46. 


47-S0. 


BULLETIN 315 


EXPLANATION OF PLATE 30 


Page 


Cyrtospirifer: rotundus mew SPECleS =. cts ret tere ue) oot st re s-¥ es aero olod ohio etalosoous ints) sloval ss «1s at osetene eie(-israta) a) eee ee eer ae ee eke eee 
1-5, Posterior, ventral, anterior, side and dorsal views of a complete specimen, 1, paratype USNM 201097b; 6-8, Side, anterior 
and dorsal views of the same paratype, x2; 9-13, Ventral, dorsal, posterior, side and exterior views of another specimen, x1, 
paratype USNM 201097a; 14-16, Dorsal, side and anterior views of the same specimen, x2. 

Percha Formation (Box Member), locality 527a. 
17-21, Anterior, side, posterior, ventral and dorsal views, <1, holotype USNM 201096; 22, 23, Side and anterior views of the 
holotype, <2. 

Percha Formation (Box Member), locality 506. 
Cyrtospirifer cf: C: thalattodoxa Crick may... 6:.:5:40sci secsissepescsrs. 5 25015 05 eons 5 ole eyeneyencdeeveue s:0ycile «ep cls ys do) slishstuslioys\ faves jet oY take IER te Ree eet ree 
24-28, Posterior, side, dorsal, ventral and anterior views, x1, hypotype USNM 200968; 29, Enlargement of the right side of the 
dorsal valve of the same specimen showing granules, <4. 

Sly Gap Formation, locality 52th. 


; \Cyrtospirifer:whitmeyii(EPall) soy eeiny cucu = cienore te scree feel se says = oveks epeueiersin wiedelcioha cies Peksteeret ole leysistake eleyarcistcaae petals kent eRe ee ene 


Dorsal, posterior, side, anterior and ventral views of a poorly preserved individual, x1, hypotype USNM 201102. 

Sly Gap Formation, locality 521d. 
Cyrtospirifer: arenarius NEW: SPECLES: «:<:is/siaie ove eros: oie acs 616 wore ave): o's 61 sien) le everson evaleve leekeex eve oll tehisle rel over olois se lelcl ioe tenant ttt tet ae 
35-37, Side, dorsal and anterior views, x1, paratype USNM 201015c; 38-41, Anterior, dorsal, side and ventral views, <1, paratype 
USNM 201015a; 42-46, Posterior, side, dorsal, ventral and anterior views, x1, holotype USNM 201015b. 

Thoroughgood Formation, locality 522. 
CyrtospiriferspeCies! 25. hc ox cis cecescte't hue Wiesunitis Gasineie ohsiaiatermece ait ays scasiecicuamsgc hie peekowitiel esas aCe ee eee eee eens 
Dorsal, anterior, side and posterior views of a wide individual, x1, figured specimen USNM 205260. 

Thoroughgood Formation, locality 522. 


107 


107 


108 


105 


108 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 30 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 31 


Heit 


Uf 


Figure 


1-10. 


11-16. 


17-42. 


43-47. 


48-58. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 


EXPLANATION OF PLATE 31 


Cyr-tospirifersCONVEXUS MEW SPECLES= 1. wciskisi ee ite ete recieve iy a eit eaeTe Tee tee ore Le ele To oes AL MRC Ts Sot ara eT SI CI SEE TET TEE eerie 
1-5, Ventral, side, posterior, anterior and dorsal views, x1, paratype USNM 201103a. 

Percha Formation (Box Member), locality 506. 
6-10, Side, dorsal, ventral, anterior and posterior views, x1, holotype USNM 201092a. 

Percha Formation (Box Member), locality 3026a. 
Gyr tospirifer: ss Pe CLES) Be) te rarscsren< teeave tevetorsYereyateyehane oye ess PMNS Sey coe eH EVA S TS Ee Yaron) 203 OSHIP Sr SEST NSS DELS SED a VORTICES eects 
11-15, Posterior, anterior, side, ventral and dorsal views, <1, figured specimen USNM 201023; 16, Dorsal view of same specimen, 
XD 

Sly Gap Formation, locality 521d. 
Cyrtospinifergpyramidalis ME Ws SPECLES ss, sco. oje ease ere aleve ever ereTe rere e NATIT ar or = VS e pd seev=y esos sys cosh sass eae OUR ee ReTE erSIR ase) Seah RTO Tos 
17-21, Anterior, side, dorsal, posterior and ventral views, x1, paratype USNM 203544; 22-24, Anterior, dorsal and side views 
of the same paratype, x2. 

Rhodes Canyon Formation, locality 528). 
25, Anterior view, x1, paratype USNM 201000a; 26, Side view of same paratype, <2; 27, Posterior view, showing a pseudodel- 
tidium, <1, paratype USNM 201000b. 

Rhodes Canyon Formation, locality 509u. 
28-32, Anterior, dorsal, side, ventral and posterior views of a small specimen, <1, paratype USNM 201053a; 33, 34, Anterior 
and dorsal views of the same specimen, 2. 

Rhodes Canyon Formation, locality 521h. 
35-39, Dorsal, posterior, ventral, anterior and side views, <1, holotype USNM 201036a; 40, 41, Side and anterior views of the 
holotype, 1.5; 42, Side view of a small specimen, <1, paratype USNM 201036b. 

Rhodes Canyon Formation, locality 524z. 


Ventral, posterior, anterior, side and dorsal views, x1, figured specimen USNM 200969. 

Thoroughgood Formation, locality 522. 
Gyn lospiniferssPe cies sete cacy sey cece sie reetensyapeteners oder Re isos S STAC eats Sa asus as ce 3) SASTRY H Ne Ge Pedate eae EO OTR ORATOR CTR Peters issuers 
48-52, Anterior, posterior, ventral, side and dorsal views of a young specimen, x1, figured specimen USNM 203535; 53-55, 
Dorsal, side and anterior views of the same specimen, 1.5; 56-58, Side, anterior and dorsal views of an adult, <1, figured 
specimen USNM 203536. 

Thoroughgood Formation, locality 522. 


187 


108 


108 


188 


Figure 
1-21. 


22-34. 


BULLETIN 315 


EXPLANATION OF PLATE 32 


Hispidariayspinosainew, SPECleSeyareter te yedelolepenereyaiapaseneto) <r aheferete eeteleteicien oleyaferoters | ol=/=veteterelelsaeteae fotelshettetetet eR st= eee ter alate ae teeta 110 
1, Dorsal view, <1, paratype USNM 201108; 2-6, Posterior, anterior, side, ventral and dorsal views of the same paratype, <2; 
7, 8, Posterior and dorsal views of the same specimen, enlarged to show spinules, x3. 

Percha Formation (Box Member), locality 527a. 
9-11, Anterior, dorsal and side views, x2, paratype USNM 201107a; 12, 13, Posterior and ventral views of the same specimen, 
showing spinules on interarea and exterior surface, <4. 

Percha Formation (Box Member), locality 506q. 
14, Dorsal view of an adult, <1, paratype USNM 201125; 15, 16, Ventral and posterior views of the same specimen, x2. 

Percha Formation (Box Member), locality 3026a. 
17, Dorsal view of a mucronate specimen, x1, holotype USNM 201126; 18-21, Anterior, side, dorsal and ventral views of the 
holotype, x2. 

Percha Formation (Box Member), locality 527d. 
Hispidaria} posterogranulosa MEW SPECIES. 5.<.0:6.55.651< aye seis svspsisy eis iejere o s/ers: stave Siele) syeyo-s bueieie lave o/sieis ereye ele #10] elev erste el ele afore eee ee 110 
22-26, Posterior, ventral, anterior, dorsal and side views, x2, paratype USNM 201109; 27, Dorsal view of the same specimen, 
Sal 

Percha Formation (Box Member), locality 3042. 
28, Dorsal view of the holotype, x1, USNM 201106; 29-33, Anterior, ventral, side, dorsal and posterior views of the holotype, 
x2; 34, Posterior view of the holotype, showing granules on interarea, <4. 

Percha Formation (Box Member), locality 3026a. 


- Cyrtiorinaikindleil (Stainbrook)s, -<../.:<-<5j0:5 05000 aie are-oie wvevarete eres crera tates revereee le eats yay alte) =) Sota (okenebenevenete ars Spree oleie ke eeron tener eee eee 111 


35-39, Posterior, side, anterior, ventral and dorsal views of a large adult, x1, hypotype USNM 20115la; 41, Exterior of shell, 
enlarged to show granular ornament, x3, hypotype USNM 20115li. 

Percha Formation (Box Member), locality 507o. 
40, Posterior view showing pseudodeltidium with plugged foramen, x1, hypotype USNM 201150a. 

Percha Formation (Box Member), locality 506q. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 32 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 33 


Figure 
1-8. 


9-15. 


16-20. 


21-34. 


35-44. 


DEVONIAN BRACHIOPODS OF NEW MExICo: COOPER AND DUTRO 189 


EXPLANATION OF PLATE 33 


Page 

Wehtospinifer/;ranus ne wsspeclesumemnnerretrercieiiac eerie eaicysl-creiister cr iter C aired ra areranntcryralert orient eke 111 
1-5, Dorsal, posterior, side, anterior and ventral views, x1, holotype USNM 200989; 6-8, Anterior, side and dorsal views of the 
holotype, 1.5. 

Contadero Formation, locality 523q. 
Menticospiriternich sl cyrtinaformis) (Hallland sWhittield)s ser crc icccre- eve) ereterteteter ie detetete eel =veteseteted -teleleterereyevetsiete ete etetdeiey eter ehelelaiaiore 112 
9-13, Anterior, posterior, ventral, dorsal and side views, x1, hypotype USNM 201105; 14, 15, Side and dorsal views of the same 
hypotype, x2. 

Sly Gap Formation, locality 521d. 
Indospiriferzattsls;orestes) (allkand! Wihithield) Seecrerciececn eye once rereuc cote ts te teyenee sayeuseyeve aleve) = o-ouanete lelthaeseje eletensiel chalayeters rsrenenstoletsyertenene cic ot 113 
Posterior, anterior, side, ventral and dorsal views, 1, hypotype USNM 200990. 

Sly Gap Formation, locality 521u. 
IndospititersexfoliatamMewiSPeCCleS sc ccratevevc niapdeyevereravetarer habe reve dete tevat cher leks ekler one asatck =! wisyar'ole 0 a taso)ny are¥etereteveretatetevadevalens eer heteetersl s eiereter anal outs 113 
21-25, Side, anterior, posterior, ventral and dorsal views, <1, paratype USNM 200881; 26, Dorsal view of the same specimen, 
1.5; 27-31, Dorsal, posterior, ventral, anterior and side views, x1, holotype USNM 200882; 32-34, Side, anterior and dorsal 
views of the holotype, =1.5. 

Sly Gap Formation, locality 506c. 
Indospicitersstrivosusi (MCEK) iran tep-cctentsltetencreter set ctereccraveye etcleve rater cv sta leneycretevesai cee ccraver a cyet-v-forclat ayansveteretnts fenate toeacKal aterecer NR Nea earner tere e esa 114 
35-39, Ventral, side, dorsal, anterior and posterior views, <1, holotype USNM 629a; 40-44, Posterior, side, anterior, dorsal and 
ventral views, <1, paratype USNM 629b. 

Upper Devonian, Neils Valley, east of Eureka, Nevada. 


me lndospiriteranomalame ws SpeCleSamrer rir rtecnteie einer ri raiieke asters) veiaverercieiaieicl adeteestoleelciavarstokeeaie vel kekotetelcieratatretieietet eters tseetere tate 112 


45_49, Ventral, posterior, side, anterior and dorsal views, <1, holotype USNM 200904. 
Sly Gap Formation, locality 506). 

50-54, Dorsal, ventral, side, anterior and posterior views, <1, paratype USNM 200905. 
Sly Gap Formation, locality 509z. 


RENN OMASALI ALC EINISSA TIE WASPECLOS occ. cer ec s<\=sai=.isy spare oxetot orarcl edd yeeteneve totes ravers teleneKenetorsl ctevsierel sneleltereteremenste) =< ccopsrspnuciege terete tannery etehsy eect 103 


55-59, Dorsal, ventral, posterior, anterior and side views, <1, paratype USNM 201022c; 60-64, Anterior, posterior, side, dorsal 
and ventral views of an unusually wide specimen, x1, paratype USNM 201022b; 65, Dorsal view of the same paratype, <2. 
Sly Gap Formation, locality 509r. 


190 BULLETIN 315 


EXPLANATION OF PLATE 34 
Figure 


Page 
1-19. Warrenella magna new species. 


1-5, Ventral, dorsal, anterior, posterior and side views of a young specimen, x1, paratype USNM 200902. 
Onate Formation, locality 523b. 


showing pseudodeltidium, x1.5. 


6-10, Posterior, ventral, anterior, dorsal and side views, <1, paratype USNM 200901a; 11, Posterodorsal view of the same paratype, 
Onate Formation, locality 523h. 


12-16, Posterior, side, anterior, ventral and dorsal views of a large adult, x1, holotype USNM 200900; 17, Dorsal view of an 
immature individual, x1, paratype USNM 201129b; 18, Posterodorsal view of an immature specimen showing the pseudodeltidium, 


x2, paratype USNM 201129c; 19, Enlargement of the exterior of a large adult, showing concentric rows of fine spines, x4, paratype 
USNM 201129a. 


Onate Formation, locality 523d. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 


PLATE 


34 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 35 


Figure 
1-3. 


4-24. 


39-54. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 191 


EXPLANATION OF PLATE 35 


WhomasariayaltumbonayS tai brooks) esc yereycysiezs aye cece crete seers ov is as ics rev ane ore eue yaya tor nate yoy SUG fo loon Milas ae TTR eae efoneee 102 
1, 2, Interior and posterior views of a ventral valve, showing details of the dental plates and interarea, x3, hypotype USNM 
201066. 

Independence Formation, locality 5281. 
3, Posterior of a complete specimen showing traces of a deltidial cover, x2, hypotype USNM 135294a. 

Independence Formation, locality 528m. 
shomasariaidemissa new SPECIES! ears 5.5. cece. <farene esis orate) eo ote oraeche Cian are Ta Soto OA tie, re ee eee Ee eee 103 
4-8, Posterior, side, ventral, anterior and dorsal views, x1, paratype USNM 200987d. 

Sly Gap Formation, locality 5090. 
9-13, Ventral, posterior, dorsal, side and anterior views, 1, holotype USNM 200936b; 14, Side view of the holotype, x2; 15— 
19, Ventral, posterior, dorsal, side and anterior views of a strongly pyramidal specimen, x1, paratype USNM 200936a. 

Sly Gap Formation, locality 52le. 
20-24, Dorsal, side, posterior, ventral and anterior views, 2, paratype USNM 201157. 

Sly Gap Formation, locality 5090. 


ceehomasarialwarrenmiine Ww SPECIES = 2y.:'sy</ousfais)= eresarsie eis eosvevmaue rai e s¥elssesei  aeaerajo051 61506 eyar acids ebei,  shsvavs yee eevee avenegesclevosa eres erenorey sete ie teeters 103 


25-29, Anterior, dorsal, posterior, ventral and side views, x1, paratype USNM 200959. 

Sly Gap Formation, locality 509n. 
30-34, Anterior, side, posterior, dorsal and ventral views, x1, holotype USNM 200988a. 

Sly Gap Formation, locality 506). 
35-38, Side, anterior, posterior and dorsal views of a large individual, x1, paratype USNM 201025a. 

Sly Gap Formation, locality 5090. 
IW armenellaimOwerisne WSPECLS mo sejcccsclcvees stevet ers cveys hoi eya soya re regeTe Rete eneyS rete ne eeey ey ova Sus ce ony 5 Pera ee Pee ever Lakes ea eRe 115 
39-43, Dorsal, posterior, side, ventral and anterior views, 1, paratype USNM 200883a; 44-48, Side, dorsal, ventral, anterior 
and posterior views, <1, holotype USNM 200883b; 49, Interior of an imperfect ventral valve showing dental plates and pseudo- 
deltidium, <1.5, paratype USNM 200883c. 

Onate Formation, locality 5230. 
50-54, Ventral, anterior, posterior, side and dorsal views, x1, paratype USNM 200923a. 

Onate Formation, locality 524x. 


192 


Figure 


27-44. 


56-60. 


BULLETIN 315 


EXPLANATION OF PLATE 36 


1-S, Posterior, ventral, side, anterior and dorsal views, <1, paratype USNM 201001a. 
Sly Gap Formation, locality 5061. 
6-10, Anterior, side, dorsal, ventral and posterior views, <1, holotype USNM 201050a. 
Sly Gap Formation, locality 506c. 
11-15, Ventral, side, anterior, posterior and dorsal views, <1, paratype USNM 200958; 16-18, Dorsal, side and anterior views 
of the same paratype, 2. 
Sly Gap Formation, locality 509h. 
19-21, Anterior, side and dorsal views of a large specimen, <1, paratype USNM 201008. 
Sly Gap Formation, locality 509w. 
22-24, Dorsal, side and anterior views, <1.5, paratype USNM 201132b; 25, Posterior of the same paratype showing deltidial 
plate, x2. 
Sly Gap Formation, locality 3045b. 
26, Posterior of another specimen showing deltidial plate, x2, paratype USNM 201142. 
Sly Gap Formation, locality 509h. 
Thomasaria warreniinew Species. = 25,<terecs -lesine we eneeiy ae a8 Date. cuarernte sy oe SAIS Ae oIe Be RE SOs eclare este alae ein) Tae Eee eee nee 103 
27-31, Ventral, dorsal, anterior, posterior and side views, x1, hypotype USNM 200935b; 32-36, Anterior, dorsal, side, ventral 
and posterior views of a small specimen, x1, hypotype USNM 200935Sc; 37, Side view of the same specimen, 1.5. 
Sly Gap Formation, locality 52Ic. 
38-42, Posterior, anterior, side, ventral and dorsal views of a large specimen, x1, hypotype USNM 203520b. 
Sly Gap Formation, locality 506c. 
43, Posterior view showing apical plates, x2, hypotype USNM 201130. 
Sly Gap Formation, locality 509h. 
44, Exterior of the left side of a dorsal valve, showing concentric arrangement of very fine spines, x5, hypotype USNM 201148. 
Sly Gap Formation, locality 3045b. 


; Hlitabmiagister: new SPECleS: <5 %s.ecc.ecs-vve ya. sjo-sceee tis in, siecle Sie tyes sia eine crys SheieceveLs © OGCs onlay ale OPV ISIGINTS oll STSTe TOTS TO STOO een ee 116 


4549, Ventral, dorsal, posterior, side and anterior views, x1, holotype USNM 200906; 50, Posterior of the holotype, showing 
oblique, lateral plates on the sides of the delthyrium, 2. 

Sly Gap Formation, locality 524m. 
51-55, Anterior, posterior, ventral, side and dorsal views of a large specimen, <1, paratype USNM 200907. 

Sly Gap Formation, locality 524n. 
Oiosia\putillai(Stain brook). syave ceveie/cccyaschs ferepasaveve: «ete araceratvetese nereacotepey exeMe Ra soe) av:ta eterene etmceeearete ere Testor ere yatober = er aay te Ter toTaT ee 117 
Anterior, posterior, side, ventral and dorsal views of a complete specimen, <1, hypotype USNM 200976. 

Percha Formation (Box Member), locality 506b. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 36 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 37 


Figure 
1-35. 


36-41. 


42-65. 


DEVONIAN BRACHIOPODS OF NEW MEXxIco: COOPER AND DUTRO 193 


EXPLANATION OF PLATE 37 


CyrtinatinultayS tain brooke eer rrr eects lever eve eevee leseke tele sedes tate es evoneve eel fete elal she toletotetefetetenetaneteetetetey tele pete teke tenet e-Iter Tot 117 
1-5, Dorsal, ventral, posterior, anterior and side views of a small specimen, x1, hypotype USNM 201068f; 6-10, Side, posterior, 
dorsal, ventral and anterior views of a large specimen, <1, hypotype USNM 201068a; 11-15, Posterior, side, anterior, ventral 
and dorsal views, x1, hypotype USNM 201068b; 16-20, Posterior, dorsal, anterior, ventral and side views of a small specimen, 
x1, hypotype USNM 201068c; 21-25, Ventral, dorsal, anterior, posterior and side views of the same hypotype, x2; 26-30, 
Ventral, posterior, anterior, dorsal and side views of a wide specimen, <2, hypotype USNM 201068e; 31-35, Anterior, dorsal, 
side, ventral and posterior views of an elongate specimen, x2, hypotype USNM 201068d. 

Sly Gap Formation, locality 506c. 
(CORINA TEES OW GOES papaccocanbovonnoogson abc onn|bdoccbHacsD Doo Ub Hoo DO DOB DoEcdOOCODLQUOODSODDODODOGCUGOSENE 118 
36, Dorsal view, x1, holotype USNM 201270; 37-41, Posterior, dorsal, side, ventral and anterior views of the same specimen, 
xDe 

Onate Formation, locality 5230. 
Cis MITA (GAMO), csdocacecsvoocagncsoocaoosHvedosedOdHDMnGoGUDDsUOUePDSo GBI OKUEEDUOgGHUGOOocnOgoadouoRsadaS 117 
42-46, Dorsal, posterior, anterior, ventral and side views of a complete specimen, <1, hypotype USNM 200977b; 47-49, Dorsal, 
anterior and side views of the same hypotype, 2; 50, Posterior of a dorsal valve, showing small dorsal plates, x2, hypotype 
USNM 200977e. 

Percha Formation (Box Member), locality 506q. 
51-55, Side, dorsal, anterior, ventral and posterior views, x1, hypotype USNM 200975; 56-58, Side, anterior and dorsal views 
of the same hypotype, x2. 

Percha Formation (Box Member), locality 3026a. 
59, Dorsal view of a complete specimen, x1, hypotype USNM 201145a; 60-64, Side, anterior, posterior, dorsal and ventral views 
of the same specimen, x2. 

Percha Formation (Box Member), locality 509e. 
65, Enlargement of the right side of a dorsal valve showing fine, spinose ornament, x5, hypotype USNM 201144. 

Percha Formation (Box Member), locality 506p. 


194 


Figure 
1-12. 


13-19. 


20-23. 


30. 


Si; 


44-53. 


BULLETIN 315 


EXPLANATION OF PLATE 38 


GCyrtinatyariabilis Mew SPEClES = veye ere ere) = clove sfetes shes! o)ete te tater ey vol oy ealobel ct otor-tatoratetetato(oyotelereetelete stereo retateteeinint ie) ister et tenet tet ee 118 
1, Dorsal view, x1, paratype USNM 203558; 2-6, Ventral, side, anterior, dorsal and posterior views of the same specimen, x2. 
Onate Formation, locality 528n. 
7, Dorsal view, 1, paratype USNM 201271; 8-12, Dorsal, anterior, side, ventral and posterior views of the same specimen, x2. 
Onate Formation, locality 523i. 
Cariniferella\ costellata MEW SPECIES ~ 1.4.5: <.ésc. css 3 <:c; cvsneue te: wtetouzre) #:fceae ein iovsi = overs wis] eye ¥ssepedtersucipy ayers a) ausitevene yeysyeteireseleto e126) siete o leds teachin 41 
13-17, Side, posterior, anterior, ventral and dorsal views, x1, holotype USNM 201214; 18, 19, Dorsal and ventral views of the 
holotype, x2. 
Independence Formation, locality 526u. 
Douvillina ‘contaderoensis: DEW SPECIES 5 -<:<c6)<)<.e-ersisyevsisteyey erative <i avs,o)ekoyeisis. eloisistojeverhe oetelao seieeia suelo oes nis Slee elo ee eter 50 
Ventral, posterior, side and dorsal views of an imperfect specimen, x1, hypotype USNM 201009. 
Contadero Formation, locality 523n. 


» Petrocraniaispecifically sunidenmtise dy yr yajer- areal tevevaleteraiekeesreyehenetct arene = eloloi= aie cs) eedeneeeetetetene ek ceeeehe etetchiehelel ee asta eee 34 


Exterior of the dorsal valve, x2, hypotype USNM 200954. 
Sly Gap Formation, locality 506c. 


i (Schizophoriailata:S tain brOOks Wa rereeceeyec t= et onetetetesescro Sy sree evere: sve en cre seteteeenenc Ped Mees ots voles REN aera er ec etee steve el aL NS enact eee 36 


Anterior, posterior, ventral, side and dorsal views, x1, hypotype USNM 200924. 

Onate Formation, locality 522h. 
Cranaena’ navicellaa (Pall) e° i. csicacateetaearate fe sas sata eaters pate es eee al au te ails aa fovea 08 UGTS aris fo a eras la (BETS lene < peteyebaife fe eee Ud (ele ce ata eee eae 119 
Dorsal view of an imperfect specimen, <1, hypotype USNM 203564. 

Rhodes Canyon Formation, locality 528). 
Afilasmia'SPeCiGS: cisieiesisie.e sue. ai 3: epayeeyetaveus ele ai 8 63: aie where 0 a: avarice, auerececa se) Sve tn evn lb ylee coy apa deceteylohs ay alaneho kate fol euaistcte- joy etasieken etcetera teeta 119 
Dorsal view of a large specimen, 1, figured specimen USNM 200995b. 

Thoroughgood Formation, locality 522. 


> Afilasma inexpectatuum new SPECIES: se x:-5-.00:-ceierey ater aorare esevedaetetereseuslnte 21s tein sla dete bate ate etetetevet sore tevenet siete ciciat ete Sicko fete eens 119 


32-36, Anterior, ventral, posterior, dorsal and side views of a distorted individual, «1, paratype USNM 200930. 

Contadero Formation, locality 509u. 
37-41, Anterior, posterior, side, dorsal and ventral views, <1, holotype USNM 203530; 42, 43, Side and dorsal views of the 
holotype, <2. 

Contadero Formation, locality 528a. 
Cranaena’ calvini) (Fall ‘and: Whitfield) =, s(ove)s.6:<ieveisvess ieee: 6.03: 2se Svein re eves 51 Su erered os © oi cee coVeVeUeroueye RSG uels eles 6) acere oleae Loneh CPS eRe eRe 118 
44-48, Posterior, anterior, side, dorsal and ventral views of a restored specimen, x1, hypotype USNM 200844a (left half restored 
when viewed from the dorsal side). 

Sly Gap Formation, locality 52Ir. 
49-53, Anterior, posterior, ventral, side and dorsal views, x1, hypotype USNM 200845. 

Sly Gap Formation, locality 509z. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 38 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 39 


Figure 


DEVONIAN BRACHIOPODS OF NEW MEXIco: COOPER AND DuTRO 195 


EXPLANATION OF PLATE 39 


1S tain brookiayr otun dayne wsSPeCles: 15/5 assteisis cielo terosiasioteteteevorstedetes eRe era eet oMea cvs eee ie Ocho shed eee eee 39 


18-21. 


23-26. 


30-34. 


35-37. 


38-42. 


43, 44. 


45-47. 


48-S1. 


1, Dorsal view, x1, holotype USNM 207700a; 2-6, Posterior, side, ventral, anterior and dorsal views of the holotype, 2; 7, 
Ventral view of a larger specimen, «2, paratype USNM 207700b. 
Contadero Formation, locality 526a. 


‘wHypothyridinajemmonsijrotundanew, subspecies, «meee eeciee kien eee nea ee eee senna Eee eee eeee 72 


8-12, Posterior, ventral, anterior, side and dorsal views of a complete specimen, <1, paratype USNM 201160a. 
Sly Gap Formation, locality 509h. 
13-17, Ventral, posterior, anterior, side and dorsal views of larger specimen than the preceding, <1, paratype USNM 201162. 
Sly Gap Formation, locality 506c. 
Stainbrookiaymtenad (Calvi) es ejscaxc cy etegss crete te eyetesereae ree eackeeee sas Ne ees Cova yee eh cate Ta ava eM Ces ER TR al ct sete ML te 39 
18, 19, Interior of two ventral valves, showing muscle marks and median ridge, x2, hypotypes USNM 200886b, c; 20, 21, Interior 
of two dorsal valves, showing brachiophores and median ridge, x2, hypotypes USNM 200886d, e. 
Independence Formation, locality 526w. 


A ulacellavellipGeaiNe Ww. SPECIESS ices) vers eves sere clorcisrar-ye a csepeseiateteptere oes eHLD soo eave NT Sie ose vei omayas rer Sool csc Eo ote GIES 38 


Interior of the dorsal valve, showing cardinalia, x2, paratype USNM 200909. 

Sly Gap Formation, locality 528y. 
HylothyrisicompactaMeweSpeClesi. rec. .yeretey so sis) s/o, ecier MME eve ev eeyeyor ol shersr aves oj 0) cle ro ae Ieners lege eT ert eateraiele eer eee eee ea 97 
Dorsal, side, anterior and ventral views of an exfoliated specimen, showing median costa in the sulcus, x1, paratype USNM 
201032. 

Contadero Formation, locality 522a. 


PCC ATIACIALSDECLES tle e Me varrese, cree yy ass eps fac ssyes 6, <hstayeseheba eicases ovo vss ahs hel eo ees atees SII cate roy Bakley alin 8v SRE Gy OCT SE Ee 119 


Side and dorsal views of an imperfect specimen, 1, figured specimen USNM 207694. 
Onate Formation, locality 522f. 


meen tamerellay spe Cie Sey tps sy sscisiey staat echoes tote) seca, Sees ape Tete av ics ta ave RELS SPRL S USE ST 6-0 co SOTA ROI EV Oa OIG) re SETA EER eT ee ee 69 


Ventral view of an imperfect valve, <1, figured specimen USNM 207699. 

Onate Formation, locality 3023c. 
Schizophoriayamanaensis:s tan Oro Okie 5 fe(cszyssasavapsiess oie sy cto Oho) ere MTOR aoe PU Gee Sta eS ORS PaO Ce TE Se Tey Ne ah eee IA cet Re 35 
Dorsal, posterior, side, anterior and ventral views of a rotund specimen, x1, hypotype USNM 201197a. 

Contadero Formation, locality 526a. 
MhOmMasarialGemissawMlewW/ SPECIES 0%. ass sce ceeho¥e cpr ay esc eet cys pe gee oReT STS NSU SE Ge oy oI aT SER Tr: Ter ee 103 
Anterior, side and dorsal views of a complete specimen showing incipient costation, x1, paratype USNM 205261. 

Sly Gap Formation, locality 529p. 
CyrtOspinifEr;SPE CESS. Mrs eoraters st ousr oat othe ose eV CAPs (o-aeyeu cteray te dovahen poe cue eves otSretxe late oleh oR PORCRS AL ISI SIGE GE ea Paice 109 
Dorsal, posterior, side, anterior and ventral views of a complete but exfoliated specimen, x1, figured specimen USNM 207696. 

Sly Gap Formation, USGS locality 543. 
(CORTON TM GR Ges CSF tonto app en nace do DonE eo Aaprondovcce se coaeccoan Serena caemEn Sra cot coco suneam conn etos sonnbasddees 109 
Dorsal and ventral views of rubber replicas of exterior molds showing granulose ornament x2, figured specimens USNM 
207703b, a. 

Sly Gap Formation?, locality 522s. 
Bileuth err ok ommmmais pe CLES ceyacsysssy ogo ou ch ov sievey sev nsver ost ots fever Sey oper oye ysstre rete al one exer cy ctsteaversyiapere eiicievaselione ete POR Paes et ae OES Oe eT 96 
Views of rubber replicas of the ventral exterior, ventral interior and dorsal exterior, <2, figured specimens USNM 203560b, c, 
a. Note absence of a median septum in the ventral interior. 

Sly Gap Formation?, locality 522s. 
hylothyrisisulcocostata MEWeSPECLES = o.y.censis,o,s:o myn cies ns snctetese el cietose's eieie noua «eget stercuekereaetetetcyonoteceeye Or aGie rer eaevers cick otis erent ee areas 98 
48-50, Dorsal, side and anterior views, <1, paratype USNM 201043; 51, Ventral view of the same specimen, showing costa in 
the sulcus, x2. 

Sly Gap Formation, locality 509r. 


Se LingulipOrayporiterayneweSPECIES.< cm /sioscie: Melek eek ey fo seuei > “valet skereteyen clas snovs sexerere seo orerers Suerolstevehe fe cierto eSNG stoutetadey fare eee eee Tere 33 


52, Exterior of the ventral valve, x4, replica of holotype USNM 200899b; 53, 54, Interior and exterior of ventral valves, x4, 
paratypes USNM 200899d, j; 55, 56, Interior and exterior of dorsal valves, x4, paratypes USNM 200899f, h. 
Percha Formation (Ready Pay Member), locality 3037. 


196 BULLETIN 315 
EXPLANATION OF PLATE 40 
Serial Sections 
Figure Page 
1=10: Coeloterorhynchus: magnificum NEW SPECIES = sa ci-s 5 ace 2 oe ess sie vere reseces seis aes) aust) ovs|osasetsv eve aueyora/oSieapa/acheve tor svars ale eee eer ae eee 76 
Serial sections of a large adult showing dental plates and lack of a median septum, in mm from the beak: | = 0.1, 2 = 1.1, 3 = 
1.8, 4 = 2.1, 5 = 2.4, 6 = 2.9, 7 =3.1, 8 = 3.6, 9 = 4.2, 10 = 4.6. All x3. Paratype USNM 200913a. 
Locality 506c. 
I1=195'Colophragmayellipticuminew/SPEClES = ayeya yatelote= tel eletatetey ele) avotcteletaleteteteketelatolstatetetototaer =) elaleteyevetstst tel elicestelel leks steele teeta ee eee 77 
Serial sections showing dental plates and short median septum in mm from a point near the beak = 11: 12 = 0.3, 13 = 0.5, 14 
0.6, 15 = 0.8, 16 = 1.0, 17 = 1.1, 18 = 1.4, 19 = 1.7. All x3. Paratype USNM 200953a. 
Locality 5240. 
20=25.. Camarotoechialatisinuata NEW. SPECIES. fcc se)ysiec cache ons: st cetavayecs oo) 2) sispeyererciesa1= erchurel ayers ovayavonevaltesyehets eheleo=sclt steer hotel evel ae eee et eee 79 
Serial sections showing thickened dorsal structures in mm from the beak: 20 = 1.1, 21 1.6, 22 = 1.9, 23 = 2.4, 24 = 2.4 at 
<4, 25 = 2.9. All except 24 are x3. Paratype USNM 200945g. 
Locality 522. 
26327... Lhomasaria\warrenisnewSpCCleSs 2. cl-tcicio evensierseeici=) Vee ko nels ee eer Sereleileieis sheild a eeeiia aoe eel eee eee eee 103 
Serial sections 2.0 mm and 2.6 mm from beak showing cardinalia and dental plates, x2. Paratype USNM 341931. 
Locality 5090. 
28-3); “Trifidorostellum obscurum MEW SPECIES. ciaicc ccs seis gece sicyaieisy ev ovepe te ayeieis:s /ayoie Sua veuetswlechye uate eisrete axelererate terete ee oleae eek ene een 78 
Serial sections showing short dental plates and absence of median septum in mm from the beak: 28 = 0.8, 29 = 1.1, 30 = 1.5, 
31 = 1.7. All x4. Paratype USNM 341932. 
Locality 507n. 
32=39;. Calyinariaybransomi/ Stain brook... -...5.<sstezsvave eo os an pce eue) s 31a shecapstel'n aie (ov eyata Guat eile Ue housn yay oy abaletel skcteess ogous eke toyota erate eed nce aoe eee ee 80 
Serial sections showing thick cardinalia in mm from beak: 32 = 0.1, 33 = 0.3, 34 = 0.45, 35 = 0.6, 36 = 0.9, 37 = 1.0, 38 = 1.2, 
39 = 1.4. All x2. Hypotype USNM 341933. 
Locality S506c. 
40: Paurorhyncha:cooperi(Stainbrook. xo och Siccccsteca rege ecara este crave rectie sy wheres wn wea cpticea aera wfetevah Grate aepferev eben ct enees ayener ates rato natal eae tee Tne eee eee 82 


Section cut on plane where teeth and sockets join, x2. Hypotype USNM 200816. 
Locality 507h. 


Figure 
1-7. 


8-17. 


18-26. 


34-40. 


4149. 


50-53. 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 197 


EXPLANATION OF PLATE 41 
Serial Sections 


Hy borhynchellaibransomi!COOper) y.ie:s,esecsue ks cei eh esos cise seperate oie oe ants se Coyey na iS ave vache eval os ante vabee ayes Seca al eeay cue ote Lavell aye yeleev ane evenaiteks 82 
Serial sections showing internal structures. Sections in mm from the beak: 1 = 0.1, 2 = 0.2, 3 = 0.7, 4 = 0.9, 5 = 1.0, 6 = 1.7, 
= 2.4. All x4. Compare these sections with those of Camarotoechia and Calvinaria (P|. 40). Hypotype USNM 341934. 
Locality S506c. 
Etychomalotoechia;sobrinal(Stainbrook):. joys. cr. /Syercc ce ecerose les euccarctever<yereee mie sy 20ers tabi a ch Speve et aonn yap el wte levey eves casters) ayohevetet ote aresstere sveretere 70 
Serial sections showing dental plates and cardinalia, in mm from the beak: 8 = 0.5, 9 = 0.7, 10 = 0.8, 11 = 0.9, 12 = 1.1, 13 
1.2, 14 = 1.5, 1S = 1.8, 16 = 1.9, 17 = 2.5. All x3. Hypotype USNM 341943. 
Locality 506b. 
Cuapularostrum?:stevensomiine Ww, SPeCleS >. y:01<eter-ser-vassactte eee oe oteholo re oy ees rose oils ence lose lole ein eto sen ok Oe eee 69 
Serial sections showing dental plates and cardinalia in mm from the beak: 18 = 0.2, 19 = 0.8, 20 = 1.1, 21 1.2, 23 = 1.5, 24 
1.6, 25 = 1.8, 26 = 2.2. All x4. Compare with Prychomalotoechia above. Stippling = adventitious shell. Paratype USNM 201029c. 
Locality 509u. 


Pe LHOMASAFIAy CEMMUSSAUNEW)SPECIES:: 5 teva eitte oy cue avsun esis neve aves aes ay singe. alls vs epaveraieiene a syevav ever ele. eye overavore & cteyerroree rain sreventie encievare eee eons 103 


Serial sections through the beak of the ventral valve in mm from the beak. 27 = 0.2, 28 = 0.7, 29 = 1.6, 30 = 2.2, 31 = 2.3, 32 
2.4, 33 = 2.6. All x3. Paratype USNM 200936c. 

Locality 52le. 
Ripidiorhynchus newmexicoense NEW SPECIES: <<< /-geicresa2ee) caravans ore erase seb aveues gai w/e dso: ai ayoressveueile eevee aqaye e ayaysie dine srelesreietels alereiaienal sts 71 
Serial sections showing dental plates and hinge plate in mm from the beak: 34 = 0.3?, 35 = 0.5?, 36 = 0.6?, 37 = 0.92, 38 = 
1.1?, 39 = 1.3?, 40 = 1.5?. All x3. Exact distances from tip of beak to sections are estimates because the actual tip of the beak 
was missing. Paratype 200920c. 

Locality 509t. 
Morphorhynchus:varicostatum' Dew. 'SpeCieS., ia ccscc nie areeieielelsiainis oie cree, oscvene tereiese oyeie cl cteleisi ain eiatolotescleierevacciansieloreielal tereretern ciation 72 
Serial sections showing long dental plates and hinge plate in mm from the beak: 41 = 0.2, 42 = 0.6, 43 = 0.9, 44 = 1.0, 45 
1.1, 46 = 1.2, 47 = 1.3, 48, 49 = 1.6. All x3, except 49 = x6. Paratype USNM 200922d. 

Locality 522f. 
Cyrtiorinalkindleti(StainbrOoke)!, ciz.s{s:evesereis, speysesve ie sage evens wie etosaye calor saye ovevaas ees) nseienopn cos sole ayer vere copie eotecnesore ee eee ytteiets Gaels eletaleistereiome 111 
50, Section of a ventral valve showing dental plates and deltidial cover, x2. Stippling indicates adventitious shell. Hypotype 
USNM 203528a. Section taken 8 mm anterior to the beak. 51, Another specimen with section cut about 5 mm anterior to the 
beak and showing a deltidial cover with irregular external surface, x2, hypotype USNM 203528c. Stippling showing adventitious 
shell. 

Locality 507h. 
52, 53, Two specimens sectioned about 5 mm anterior to the beak to show the relationship of the deltidial cover to the apical 
plate, x2, hypotypes USNM 203529b, c. 

Locality 507a. 


198 


Figure 
1-10: 


11-25. 


27-32. 


BULLETIN 315 


EXPLANATION OF PLATE 42 
Serial Sections 


Moryuiferellayspinosal (Kindle) Sarr leery ler creleiptelteleiteteterelo steietecietee tke cicisicn ieee ase oie rete steteieiele ete teenie ee 116 
Serial sections showing dental plates, ventral median ridge and cardinalia in mm from the beak. 1 = 0.4, 2 = 1.4, 3 = 1.6, 4 
1.8, 5 = 2.1, 6 = 2.5, 7 = 2.8, 8 = 3.0, 9 = 3.3, 10 = 3.6. All x3. Hypotype USNM 341936. 
Locality 506b. 
COHUSE PUM (Sei GI pancanomonconnndocosnnoosHbenoesim ac ceo bondeoUbodunonoe Doc orEescoonpocron loeb cdbERdornon’ 117 
Serial sections showing septa and cardinalia in mm from the beak: 11 = 0.3, 12 = 0.6, 13 = 1.1, 14 = 1.3, 15S = 1.7, 16 = 1.9, 
17 = 2215 18 = 2:2, 19) = 2335 20 = 2:5, 21 = 2-7, 22 = 2.9) 23° = 321,24 = 3°35 25 — 3:9) All x3 Hypotype USNM3419352 
Locality 509b. 


= Warrenella' HOwerlinGW.SPEClESs rer roayoreteisie lsieiarcfeyeheieyereselo wiexs-ere lovoxcle Vane ehereiaete elche/eisi at ule luchekey treteke siocsele lo] scien 115 


Section at about midvalve showing position of spire, <1.5, paratype USNM 200883d. 
Locality 5230. 
Petasmariaypatens)new) SPECIES ayers t-¥- Varroa ce erer ene pene ep ev ee eel favre sey PaYoder ests eis ote lest srey ave otketstareysieliektelefe lets rslce te cick ree eee ieee 85 
Serial sections showing septa and cardinalia in mm from the beak: 27 = 0.4, 28 = 1.1, 29 = 1.4, 30 = 1.7, 31 1.9, 32 = 2:0: 
All x3. Paratype USNM 200937a. 
Locality 507h. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 43 


Figure 
1-9. 


10-22. 


DEVONIAN BRACHIOPODS OF NEW MExIco: COOPER AND DUTRO 199 


EXPLANATION OF PLATE 43 
Serial Sections 


Mhomasanria: WarrenisMew SPECIES.) <5 2s. 2 axiche mie = uchsncteyasccce oo ete Sy ar sora toray ents) 5 a76o) aucys es o,a ce e/ah oS ayaa deV ev ayavevaveyercusvevavelovecereuete rere eevee eres 103 
Serial sections showing dental plates and cardinalia. Stippling indicates adventitious shell. Distance from beak in mm: 1 = 0.7, 
2 = 1.2,3 = 1.5, 4 = 1.6, 5 = 1.8, 6 = 2.6, 7 = 2.9, 8 = 3.0, 9 = 3.2. 1-5 = x4; 6-9 = x3. Paratype USNM 341937. 
Locality 506c. 
Wiarrenellaihowerl NEW SPECIES =. fei. oes. orcs ovevs: siste esse av aioeetaycvereraysreseledire  <cbus eye gates euegeser seas wives ovate vereneieeanes esare iste cetarstereiciers leisieneiate ersten 115 
Serial sections showing thick shell, dental plates and cardinalia. Distances of sections from beak in mm: 10 = 1.5, 11 = 3.0, 12 = 
3.8, 13 = 4.4, 14 = 5.0, 15 = 5.6, 16 = 6.1, 17 = 7.1, 18 = 7.5, 19 = 7.7, 20 = 8.0, 21 = 8.2. All x2. Paratype USNM 200883a. 
22, Section of a dorsal valve about one mm anterior to the beak, x2. Paratype USNM 200883e. 
Locality 5230. 


200 


Figure 
1-10. 


11-20. 


21-26. 


36-47. 


BULLETIN 315 


EXPLANATION OF PLATE 44 
Serial Sections 


Page 

Petasmariavpatensimew/SpeCes yrs veteyers)-\-es- footer retake ere ete erase aed oles arel eae t=) al lelistate el ley eRe tee totel ole crete ee tee eee 85 
Serial sections showing septa and cardinalia in mm from the beak: 1 = 0.7, 2 = 1.5, 3 = 1.8, 4 = 2.5, 5 = 2.8, 6 = 2.9, 7 = 3.0, 
8 = 3.2, 9 = 3.3, 10 = ? [>3.3]. All x3. Paratype USNM 341940. 

Locality 506b. 
AcutathecayprolificaynewsSPeClesey serie rvstoieteretersreteteherersete ieee terarstatercnenove le reveke ten oreheyetsge enetetere fetched evere tet telseaiet irate ee 101 
Serial sections showing septa and cardinalia in mm from the beak: 11 1:0; 12;= 0-5; 13°=" 168, 14 = 2:05 15 = 2335 16 ale 
17 = 3.4, 18 = 3.8, 19 = 4.0, 20 = 4.4. Figures 11-15 = x3; figures 16-20 = x1.5. Paratype 341938. 

Locality 509g. 
Mylothyrisssulcocostatanne wi SpeClesepeeeye terete deter de detde et efotied -Tetetet folie shetetete ta eel tateiarelet ie tarene te etete rete te elise ale oe oie e eee 98 


25 = 2.1, 26 = 2.5. All x1.5. Paratype USNM 201067d. 
Locality 506). 


352 Oiosia)putillay(Stainbrook) © mance eer ieeetle tele rrat iirc iether tertete cee oers aii ciiensintenett tier etcetera 117 


Serial sections showing dental plates and cardinalia in mm from the beak: 27 = 1.0, 28 = 1.5, 29 = 1.8, 30 = 2.6, 31 = 4.0, 32 
4.3, 33 = 4.5, 34 = 4.7, 35 = 5.1. All x3 except 35 = 6 of the cardinalia. Hypotype USNM 341944. 

Locality 509b. 
Afilasmavinexpectatumime waSpeCles eer verettelete stacy petal onetayarat-tatehevenctenstetetetere oleate cesnevole cere et atare ett eileen ee teke te ee ete 119 
Serial sections showing plates and cardinalia in mm from the beak: 36 = ca. 0.3, 37 = 1.8, 38 = 2.7, 39 = 3.0, 40 = 3.8, 41 = 
4.1, 42 = 4.2, 43 = 4.4, 44 = 6.1, 45 = 7.6, 46 = 7.9, 47 = 10.4. All x2. Paratype USNM 341939. 

Locality 523v. 


BULLETINS OF AMERICAN PALEONTOLOGY, VOLUMES 82, 83 PLATE 45 


uP aS no aN 


/ 
SU 
ny 
‘ 13 


\ 


f 
/ 
\t\ Dy, 
; 
! / 
DIN G7, 


SS 


Figure 
1-15. 


16-32. 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DuTRO 201 


EXPLANATION OF PLATE 45 
Serial Sections 


Crinisarinabreticulata (Stain brook) ss sists corey soc vaetes ccc varer sue ro ass crea a ape Voreieyehepor co ee rogersne es enevene eecie fave) oo Poereteyonere = lat aE SEO Ee 92 
Serial sections showing the interior, hinge plate, dental plates and development of the spire. The jugum is only partially shown. 
Distances in mm from beak are cumulative: 1 = 0.5, 2 = 1.5, 3 = 2.3, 4 = 2.4, 5 = 2.8, 6 = 3.1, 7 = 3.2, 8 = 3.6, 9 = 4.3, 10 
4.6, 11 = 5.5, 12 = 6.1, 13 = 6.4, 14 = 6.5, 15 = 8.1. All x3. Hypotype USNM 341941. 

Locality 509u. 
Compositabellulagstain brooks. zee. cys: srs; sssyseraye:oyetecsuet a: apsasoersueWaans el eve setayn owereuety eeyevs tee ant iepenayele here elev eseve ieieteta slsueie secre aroteaueter ante meets 93 
Serial sections showing hinge plate of dorsal valve, dental plates part of spire. Jugum not completely shown. Cumulative distances 
fomubeaksinumm: 16.— 2-2) X3- 1 — 22918. — 227 19) — Sl 20) 3222 340 2D — 307/23) — 4. ON 240— 4030 25) — le 26 
5.3, 27 = 6.6, 28 = 7.6, 29 = 8.8, 30 = 10.0, 31 = 10.4. All except 16 are ca. x2. Hypotype USNM 341942. 

Locality 527d. 
32 = about 16 mm from beak, 1.5. Hypotype USNM 341945. 

Locality 509c. 


202 BULLETIN 315 


INDEX 


Note: Page numbers are in light face, plate numbers are in bold face type; numbers in italics indicate principal discussions. F = foldout, 


inside front cover. 


Acanthatia nupera (Stainbrook, 1947).............2:00esseeeseeeeee 
AIGOGINGUSD: teee acne stock ses eee ses oene eee see eases 
ACUIECALUS ES PALNOLNAINOAUSecee nena etcetera te eee 27 
CETL ATTOS, JEAIRERTDIAHAR cocaccooasencseacocceacéeaéanoocnoenqsccece 100,101 
HOTTA (Cie JAAN RUAYAr  codccoccoscosopecansomunococnagnosesee 101 
Acutatheca Stainbrook: (1945) eeseeascsce cece ece sone eneneneeteaeceee 102 
Clee prOprigks taln brooks (1.9455) serseseneee eee eee ee eeneceeee 101,102 
JARO POR EATS, sogespeonocosscisceces 36,44 ...... 19,101,102 
propriasstainbrook: (1945) -c2.c-esesseesessseses eee se eens 30,101,102 
AIGHTON (GUI BENS ueeicces ss cece eer es sence enc annce 11,14,/00, 101 
ANG UlGLA WASP aise cco saeco ncces condenses sce. 28) si cace 11,14,/00,/01 
aechmorphnawEleutheroKOnundercectccsceenccsce et ecotcenteesae eee 96 
AjflasmarSteblii (96) eeeecceccseseceesseosecscerees 119 
beechent Stehlis(lOGi) excep gsceceeseacaceon eeaaees soe tecsememeecee 119 
inmexPeEClalumM SPecece season te eee ese scares 38,44 ...... 22,119,120 
SDit ces voc svansen doo- stme Seca tatonesesaeeake ame aaeeehs 38 xc. 21,119,120 
Alas karen. views seas eeceo neces sates oat gone ocdend cove ceunassnccneree ne che 22 
albertensis \Calpinarigimpen tena: conte we erence eee eee eee eee 80 
albertensis albertensis, Calvinaria .......c..c1-cecceceeceeceecensenees 80 
Allekseeval(l O60) rer ares 5 eterac ccc sosn sone e trons coe meee een neat 10,86 
ALiGhOVas (1960) Seeeresss ce ranocmenee sec omare cone eee Eee 38 
Allen, W. T. ...... -  7,31,41,128,129,131,133,136,138,142 
allent-«G@ariniferellaimeccss cere paces ienecenes See 1 Ie lesa 18,19,40,41 
GlloGotus, “ROL ENGTRUS|. san. <0 sites ee sence eos cee eee ee 27 
alta, 
EI AAE ONLY NChIAgsrer eo sterescenaenee ere we, LOSS) eee ee 14,72,74 
pele OVIOIGES erect meee note cet Necae sane ese sent oe cee eeOR ones oneee 77 
altumbona, Thomasaria ....... 
Alveolites Lamarck (1801) .... 
amana, Cranaena _ ............- 
amanaensis, Schizophoria ..............+. 
ambi gia Galvinanigiemesacc.cx <5 cc csecesee=cc vets dort nene cones eee donee 
MANOEL) 15 ENN (HEXSUE) scosaqnoocponopooo daar aosHesnoncesopooanonoe 
Capillataune Sp, cs.esasess cevesesssesese suse vee esdanes tee 26 ...... 24,93 
gregaria Hall (1860a) 
Spal Gyacsce tence: aces costae eenccesecsssenese se seeeoes 
SP cone re nassnawe ence sone Sawcanaoneces ce scacuuesaeeeen aes 26) 2545.5 20,94 
SPediawwase sca eeet ae sate re sac ctaaah coer eon e «Sus aene sees eane eee oreo eas 21,94 
GING. Tan sneespearcrOcenn ier aSecOetE ar bar CC aE Rene Cnn a aE ROPe asaer aoAE 94,95 
LenDonatal (Conrad 1842) eetete.seccccrece en tatteesteeee cee 94 
AmbOcocloides WPEChinOCcOeligharec tacos --eterceene eee 95 
americana, 
(CRUCIAL HLT Tee eacnceeOScOrpccpOonAEe CUS HRRDD ECO AoE EET EEE EE RAGEEOSECE ERG 89 
TOWQUY POO.) coces ssacinnste toons see ewan scosscertraeeees DA oes 30,89 
ANGLO GA MeDta ONah «arcane ene ese eee OO 120 
Ancyrodella 
curvata (Bransoniand Mehl; 1934)) <o0o. ooo... .sesases0scc-cceeesne 28 
nodosa Ulrich and Bassler(1926)\.........:..000s00--seseseereeees 26,28 
SD i eawnnaaacecennsaserenren-cusree ites soecscasdeen socrac antec set nu enaeer es 28 
Ancyrognathus 
asymmetricus (Ulrich and Bassler, 1926) ............-.0.00e0002+ 28 


SPirem caste tac sce cre emer aacinn tase meee eae ae eon: EERE eae 26,28 
IMOARCUIGTIS. YOUNR GUISE) .cesrsmesecteec cee eee ee ee 
Ancyrognathus triangularis Zone... 
Andersoni(1 966)! i2scuccssc secs esate e.cee-cae 
angelicoides, 
PASLY I 1Sipgoe tas xpratcte nase Aux a eeraene sealewuveeee tenet e eae 92 
SIIPIS OVIN Gi vioncnnawerme tare aoe sans eos conse Se seee EOE 29 


Crs tehal, INQOMOTHE. recancacannoansanocee7n 7700000 28) sone 11,14,/00,101 

animasensis, 

GY HtOSPIFifer Pee Te enc Ms dnd Anas nn ads Oe OE 104 
GCyrtospiniferd seecchess estos eee eon soe Soe 24,104,106 
anomala, Indospirifer ..........0..0..00005 ves OS) steaee 15,19,112,113 

anomalotruncata, Cystodictya .... 14 

Anthracospirifer Lane (1963)......... area 174%) 
rockymontara) (Marcos 1858)) e.cese-ses-ceeseeeee eee eee 120 

ApatoQnathus: SP. .c-.2004 cases soxeoeseneessuesseass vote see toes eee 27 

Aplington Formation! ~222-.--<-<e-<otoa-ceetecrecee state tee ene nee 92 

Appalachian C6 gi OM 22 ...s xcececeras oc casusveacenceess soereereeeee eee 43 

arcuata, 

Douvillina ee.ssss-eeere 
Strophodonta 

arenarius, \GyrtOspmnifer, cvscse-csesse esceese eee 30P =e 21,105 

AMIZONAl 02 ses ceneccs a cous ens sco sesvecenedeavenssedettee cane: Serene eee 29 
Defiance uplift: 22ee..c ses: scgec. cadences nsec ete e ee 29 
Mount) Martin; near Bisbee) fasces--s1----eeeee tee eee eee 120 
Redregosa Mountainsmee-ss-see-eeee cease reat seep ed) 
Swisshelm Mountains ............. san 29) 

asymmetricus; Ancyrognathus) .:.-2.<2s<- -caees--eeeetee eam 28 

AithabaschiaGnickmay (1963) iecesseeseseeeeenete eee eee eee eee 77,78 

Athabaschia? 

SPoulll cececweascup sees Mtoe eceeteecec sre seenaee wore 14,16 ...... 19,70,71 
LO ee ret cctccossocre eee nsceecciccses PA caeeer 21,71 

Athyris M’Coy (1844) ............... 
angelicoides Merriam (1940) ... 
coloradensis Girty (1900) ...... 

Tamellosa\(éveilléy 1835), c.cc2..e<nee ss oeee soeceeeeeeeenesec eee 
transversa (Stainbrook, 1947) 

Atrypa’ Dalmamn'(1828))... <2-<-2..---e<cassts-co8-e2e- pecs ome eee eee 
devoniana Webster(19211)\ cic .ceencsos ec swasese sees ee eee eee 
dignata Fenton and Fenton (1930) ............. eae, 87 
variacostata Stainbrook (1948) .................05 aos 88 
varicostata Stainbrook (1945) EES 

“SAtry pa” .cceshis cs isvesoncacscsccdeceeszete season soscesees Ge tee ee RE eee 87 

Ary partial Copper (1966) Pr-ccesc-c5-ceeossee eet teen ce eee 10,86 
TUDIG WeiSPe. ss osdbeeceeavsscesdsiaessaceasedsuesesceee Pa eases 14,86,87 

Aulacella Schuchert and Cooper (1931) .................000- 38,39,120 
ellipticaimaspe eanses-aseeeeseeeteee ee 2539) weenie 15,19,20,22,38,39 
infera) (Calvin; 1878)! Ssecceeccevseceascs-eteceeaoce eee eeee eee 38,39 
manticula (Stainbrook, 1947).... ss 2239120 

Anloporas Goldfussi(1829)) pec -cesesen-eeeee cetera nee eee heceeeeeeeee 10 

QUYOTASIRRYSSOCRONELES sence sence soacsere see sete oee eee 14,59,60 

GUTOTASOLOX, (RIYSSOCHONELES) eee ee-a-2eeececdne-eh ete e eee eee eee 60 

AustiniandlAustini (1844)! ere weseccceteececeee se sake eee 

australis, Schizophoria ....... 

Avonia Thomas (1914)............. 


defessa Stainbrook (1947) 


Baker, (C. Lo. .g.. scscsacacesdensnessnsneustsnclaeeccines successes ceteee see eeeetee 130 

barrisi?, Meristella@ 0. s.0.s.cccc jescavescanscouc eee sen ss eseuceeeee ee eeee 93 

Barroisella Halltand!' Clarke (1892) esssceccenseseeee seeeeeeeeeeeeneee 32 
SD eo aeiecsechiserenctesheaeeeecous. seen enie ee aoe eee 

Beavertail Limestone ........ 

beecheri, Afilasma ......... : 


Beechwood! (eimestone) “icecesceces-esecesseressuets eresseiecinestit se se eeee 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 203 


Belanskiey GM eceee cnc naecontansacossencesr coors mor erec srne tenes: aceon eoe 140 
Belanskau (928) poe. ncerc seeks sree eerceneeeenmees 

bellarugosus, Rhyssochonetes 
Bellodell ass pe woreecccoe- eens eae aeee ee 
bella Gompositamrerencmceneercs eters 26:45) eee 
Billings (1860) .......... 
bimesialis, Spirifer  .. 
bisbeensis, Brachythyris 


IBISPaTHOSNANOAUSCOSTATUS LONG) sesecesceeeeteckeee eee eeeeeee eee 25 
Bispinoproductus Stainbrook (1947) .............00..0.00-ceeceeeeeeeees 65 
VAFISPILOSUS StalnbrOOk: (1947) wexscessseeeseecescens.csceceseeseeeecs 65 
Bistramia Hoek (1912) 33 
BOliViae Se) sd ecc tec saxececeeceeses 33 
boonensis, Strophodonta 44 
BS OUCOUs RAMU Ratmecnss fled eetne. Geet Sods bessca fa teivsceuenesoeteatioust esate 8 
BOUCOM (LISS) eens sinessccsanausi tee wansuenen snes cece nsorectmesenesemert sans 96 
Boncotnenta len (lOG4 ee o.oo. cece -casecste sem aceeaciact IRR RRes eee eS 91 
DOWMOCKEREWROIASPINUfeR) saoscecceacaos 2a aeese nonce se ese oee econ eee 101 
BowsherpAn Wen fess escsavancsiasecess 7,31,41,124,126-133,135, 136, 
138,139,141-145,153 
Bowsers (U9GT) Weccs soe. sscci cncssciss so yoaeack Cecuadeseae a oe eae eee 8 
Bows tertenGy pidilamarsncecuce teasessesceaceeceseree 135145 sh 19,67, 68 
BrachythyrisnNis Coys (844) eccseccc2.c2-s500-c ce eee os ootiene sone 117,120 
bishzensisy stain brooks (1947) ee---cessecs=cecce<seeseecosee eeeses eee 120 
putillags tain brook: (1947) 8 cs sseseee<ennsenae see eceonoree eeseeccserte 117 
Bransousi Gn Gane ee-e aes ..-. 7,31,122—124,128, 129,133 
Bransonme wy Rei (L934) loot soccesesvassesmesssaesuaasensauacwbere wets seen 27 
BransonmandMebl(1934a) tos. cccnceces<scsceasenscoeeesdat eueaeerere 27,28 
BransoniandsMenli(1934b) |<. -2-.-sccesqc-s--0ee=scuseeeceocstsce once: 27,28 
BransougangeMehlli(938)iessceece-tecsecetecssccssese ce resensecsteassers 29 
bransoni, 
(CHAN GIOTHE® cotoncaooseaaobe been BsEbOnG see 18,19,40 ...... 18,19,30,80,8/ 
IERIDCAAIAOHANE — coccoocnasnesonasno;0ss050560b00C 20214 tie 19,82 
Avi AiIUl Se OLY SNAURU Stece-cacaeaceee o-ceteec te reenee rece eee eee 28 
breviposticus, Cyrtospirifer ............ QDS ros 22,105,106, 107,120 
1synteS (ICID) osascoseocnnceocesaR eer CeBBerC Reece Acer acenaarceRcoseerenebaantoce 117 
BI POU zee fais eee Netiese aeauss sae aedsagcesuaseresaedesseaes 3151215122 
BRONTE CEOO2) emth ec oee aces sneer dssnsiese oesiss os cllon ah ameaeiecemeeedacs amen 57 


Layee (R721) a cpodopdonceece bl coonccanccr aance a anOpacnnesenrpcrcceceo tear Oi, 2 
Bryantodus 
callitagaing (ssmeilts ISP) Sosancsscoedesssooanes9s2oncccccaopedcndseses 28 
SDs. .cdobqadadoo nea SCOH eRe aH ORp DAR ECE CER Erneta Rriccoceserechaaerepecnera 26,28 
hy OZOANS MRM ss seen cece nes Sess ses c doacenn sSoeWae es Sok canes Sas 10,14,25 
Buckman (1906) 11,15,72,92 
BUXTON IGMENOMASs (914) meecceee teeererseeces soe cense see eseeeaeee 25,120 
SI CUIATISE SAIN DLOOKS (1945) ences pec secs ae seatceseeseeec cess oneeeaee 120 
GaballeropRonmationee secs tee ss-ce-ceecece sore ceecescss 10,15,22,23,27 
(Caliitim (E3743) cccnepech eee cenep pee acemEaadarneeaceaasar 38,39,45,50,53,67,80 
Calvin (1883) 50 
Calvin (1897) 37 
GalvinarianStainbrooks (1945) on cse see -eeene sae eeat ase ese 80,81 
IDET LenSiSt testes cedae scene seseex eased sauce na sae ste saeaaisgeoes gence tases 80 
albertensis albertensis (Warren, 1928).............02-0e:c0eeeeeeeee 80 
arabic iang (Calvineel 878) wc. scenc< cert =< see see- eee n-ne seeeeee tances 80 
bransoni Stainbrook (1948) ...... 18,19,40 ...... 18,19,30,80,8/ 
calvini, 
Goelotenorhync Huser nercniessccep de ecceneae eden eee eens 76,77 
(CRA ATT! “sbeeccscdoancSOen eRe CCSeCEP OST OCC 38) sc 19,30,/78, 119 
Granaenell an rr re woes co ske cw ecds tech <i sacs «ae eae ee eS 118 
(GEORDIE . coecadaqsttodocse seco see DEO ERCCbO ACen ecoscod eeaseaapeccs 118 


(C)Dielas ria ceascunatewaescz-2 30 ss os odavae tives Sade caee coee as Roe 118 


calvini, 
Nervostrophia 
Strophodonta 
Sulcatostrophia 


Camarotoechia Hall and Clarke (1893) ..............0.0.0e0ee eee 79,80 
Contraciaibandlel (ops tially1 843) seseesese geese eeanee ae eeeeeeaene 70 
endlichix(Meek) iserces-orces seeces eerste re oe ener 82 
LEHISTDIGHA NE S05, ceccoscsesccunpesboauenosceasposased 19,40 ...... 21,79 


mesacostalis (Hall, 1843) 79 
multicostata (Hall, 1860a) .... 79 
SINUATAW(ELALLE PS O7) asec ctoonteccsecce ee reenter Err oe ae 79 
Rae SHAN OY) = ((WIEYT)) sposencooncanconasoasone tooseasnqososcosenocs 70 
(CON MAT OUO AG HARE Soy socomnconessocseceocenononeocecenen 18 ...... 24,79,80 
““Camarotoechia”™ 
Leal rari (IG EVIE LRSS7) hacenesencascedscnoadasasesoctcossaenndeanccos0x0cbace 82 
emia: (USNS. IRSD)  onoseceecgococcnoconcaStonsedbooncocoooebecceose: 82 
ACOSTA (US EN, NESIA)) cocosssnsocosonsoosocdanscacnesoconconncuc 82 
Camarotoechia (Plethorhyncha) endlichi (Meek) ................-. 82 
CAMERONiE Ly D OLY RIG INn Gaara eeereee ce aeet ete ne tence eee 74 
CONHOR ING MOGHOD ATE -cenansecanceccoconeqsaoocooantboneconeponbasonceoon 47 
@anadagee es 14,35, 100 
7. | oy Cane ne area pene Ban aspnoraeocce torpecreaconeoredec 18,22,80,82,104 
Wanniifred (Pass). ccseccc sv seveacapes sqeeek ern ese Oe 67 
MackenzieiRiver (Valley, ieesees sees: snceetecees eeenee ae eee eee DOS 
Manitobal (eat siteets soc srn cece ores eres eee ee 
MOKCHWEStEMMG: Jaron cacecetcs see eee eee ee eee SE eee 


Hay River ... 


Ontantojeee 

WESCELD |)... cnencccesseccs sscate cussnencadeeccudecs ede easos see meee 
Canutillo Formation 
capillataAmbocoeliderscerseetete eee eee ZO aos: 24,93 
Carinatus, elropidolepiusmaccsameceacessdeee center eee 43,44 
Cariniferella Schuchert and Cooper (1931)................2..02005 40,41 

UGH TDS SO nopeaccooseseeosa¢ 18,19,40,41 

costellata n. sp. 30,4] 

iowensis Stainbrook (1945)................2.00c0e0008 19,40,4] 

G)Dk: sciaudadnqcncuapaceochoodsabcddonsdaneossoocesueccsoscondcosta 19,4] 

tio ga) (Halle L867) cc.sccacqcuccescccaeceusce crac at osene a eee 4] 

Wigyiare Cel So \Wallbernigs JEU) cceacocccccsoccoonpedenorocoonccccscos 4] 
Carter (1967) Ol 
CASSIGIFOSIFUIN Zo. osdenasces acces ere oere et Re 72 
CON CTIA D, ILGIOA IN UIORS cesoscosscosncocecaeasonoce soaasnssasases090: 14,81 
Gaster (1939) \jccccuectees <svacsscseeetascrscw este mates 10,44-46,49,50 
Cayuta, Douvillina: ec sccccucescesaees conses sion OTe 50 
GDI (EX), IDOE. coca cescmoscconscooocéecdaonacosessscencosececonse 
CedarWalley himestoneesse-ceeeesseseeeeere 

Solon’ Member ---.-.-----------e- 
Centerfield Limestone 
Centronella navicella (Hall, 1867)  ................00.2005- 


Centronella(?) navicella (Hall, 1867) 
Phao (MOLT Re Ssocearotectek coces coe cecdc ee eee COR eee 


Chapinella Savage, Eberlein, and Churkin (1978).................. 77 
ChattanoogatShaley 22-2 carne <coceavacecccseescees se eee eee eee 33 
chemungensis, 

FLOW CNIG a. sot wedeas tues uocenen sick coat ou stae neat ec ee 55 

Schuchentella oct sccctcctss rece geee cee ewe eee oe PE 56 

SIKEplOriyNGhus” he: =cee-- aoc tates ctvede conaeea esc on oee ero 55 

SLOPNOMENG stescncdc wee acacia hese scenes vce tecshecesesoecee nee arte 55 
chemungensis (aff.), Floweria .... LD eee 20,55 
Chemuneskonmationieees eae eee 50,71,94,98,119 
Chonetes 

lépidusekiallk(1 851) veccecc- acs. coeceesss een seavaeostsesessomee eee 58 

schuchertiq@leland| (U9) ieeneesesee-ce- ea seese ee eee eee ee seeneeee 58 


204 BULLETIN 315 


Chonetes 


Scifulites: Coope4;. (1945)! ess.c-cscssscees ssc seeczsenseaeceseceeeceenaes 58 
Seituluspialli(U857)) sscceecescscseecs-c-cs cae ecch ace sec vesececese seer ce 58 
Seliperusubd alli (843) is es.nccecteccrcrodeseecccceossucotcercecesest eee 
Gircularisy (att.) wha lmatolepismmeescceeaeen-ceeee eee eee eeee 
Cleiothyridina Buckman (1906)............. 
devonica (Raymond, 1909)..... 
humerosa Stainbrook (1950) .. ‘. 
proutt (Swallow; 1860). ceesccc-s-2sce-scececsene eee seeps e sees 
TEtGULALARS CAIN OTOOKS (L947) Nereree eae ete eee eee cee eee 
Cleiothyridina? 
coloradensis Stainbrook (not Girty, 1900) ...................02005 92 
transversaystainbrooks (1947) reese.s-tesserae ose eece aeaceneseeeee ese 92 
Cloud) (1942). oatacs cqestcsse dousesecteenc meen ero ence ete 115,118 
@lymenia=S tuberin set secs ee ac ee cease ee eset se ee Eo eS eee 23 
Gly menial Zone essere new ctscicy eeeneestonsweses as tee senececaeeees 25,143 
Coeloterorhynchus Sartenaer (1966a) .............0.2.0.0000 18,76,77 
calvini (Fenton and Fenton, 1924) .............s.ceceeeeeeeneees 76,77 
(ETON Ws SOs sencceceoeencoecce 17,18,40 ...... 18-20, 76,77,78 
SGhucherti (StalnblOOksw1 945) leenceeseceeteet reer tesee eee eeee reese 77 
Spies Betas. Sees ceiaauseeatebats cosewe enous 
Cohoctonian Stage 
Collins ats Bry Antodusmeccres-ceeeceeree cate cece eee eR eee 28 
(COMMAOHGTE Do tESNG soasogssocgosadocodeosadoougeaecodedsoobe0 20,21,77,78 
ellipticurinSDom rte esse eesee see ceec ee 16-18,40 ...... 20,77,78 
S Ppl ots sacereatees sci-snawtvescecsuebens avue deve venir steamers seam noteee 20,78 
SP ipl) vac sdacee vite ww acount stasinesisces acgeeseadesnsenaevenmetece Ey pescoe ZAt oi 
coloradensis, 
PAILILY TIS amet e Cea cis douse eae e eee aOR SE orien Sa oe ae Ee 92 
Gletothyidinaneeracccn nen tree eee e eee Cee 92 
WSCIOPKOACUCLUS. Gennes caen cose nose sos sete oes fone ae oe 24,34,63,64 
IPF OUUCTEIIG. Girone svssiecsucteen cacseecsioe se eecacc intron ee aeneee ene ee eee 64 
coloradensis var. plicatus, Productella .............00-00c00e0000+ 63,65 
C@oloradoy Fetes ccoseaccne shane ate erate ee 
ROCKWOOd i225: Wade stececk ease seceeesnc: 


comis, Gypidula 
communis, Polygnathus 


compacta, 
IP CLASIMAT IGE oe sa RE Re Bee etaiec aces oo ese ooo at one 20 ...... 24,84 
IS PIMMAUIY DQ ods cccescinasadscenesogeceiendene soot eee DAN se 20,89, 90 
iy lothiyrismeere tad Roe Se eececo 21,97,98,99 
Complanatawlingul acres sosctssetrs deserter Meo ee 32 
Composita Brown (1849) 93 
bellula Stainbrook (1947) ..............2005 26,45 ...... 24,93,120 
GAA TGE (5 ENUSSIK SPI ecoanoapeeceosneelaeecoadcebee cau aoadesescabeoscne: 93 
trinucleas (tall i856) veescevecssee!nsencce- oscar oeraee eee ere 93 
CONMMENS WeOly OD LOdON! dracresetescereea eee eee eee enero 28 
@onodontibiostratioraphy, meee sscecoseee settee eee eee 25-27 
Conrad (1839) aces. seer esee sete se races n ne see 43,100,101 
Conrad) (1842) Oe ae. cecsiece sense ase Oe acc eee AAC eee ae LOA 
COMSODTINUS PVIUGOSDINI fan sessereee eee nee ence e ee 97 
Contaderoensis, Douvilling ....<.0.--.-.-.--c-e.s+2s0eces0s S338ieee- 50 
Contadero formation ........ ( Cane 6,8,18, 19,20, 21,22,26,27,29,32, 
33,38,50,78 ,90,94,99,119,121 
TNCA CAC») th Sane BEDE DEC RAED OEOECORE CORED EE GOERS URED ROLE abner concedoo 20 
Brachiopod assemblage of 20 
Conodonts from ve 26 
Onitlhae BAPE DAC CECE RC nCer ce DOLE ELEGCODC von lS 
LUTE onc a ERR ence eiacee CED Dochos acer Ree parE niRem are rmcas arassoncend 15 

contracta, 
GAMOALOlOECHIG vec. caess2s0s02esc0ee0: sgoededtebevusceesseestarncecaeatrs 70 
CUPUIGROSUUIN: secccuncene cus tavens ace ve cuisnaws dev naGe eeu ee 70 
CONVEXUS GYTIOSPINIED (ensucessticenecesezes OL. ..--.28 24,100 MO7a109) 
Coopers: Case (193i) seni oeavecess dence se ; 27 
Coopers (I9S9) Mee cere cctie. es esere 27 


Cooper; (Gy Avec nebo ccasie savsivacee getaeaee teases tenneia te 7,31,153-156 
Cooper, G. A wae 23525,51580582 
Coopers'G A944): ac cscies derweesenstdeteaccuceaeneee cues oeees 82,112 
Cooper,:G: Ax (1945) cccccsessxotacedateveienedoreteesleeessnatereeorenee 58 
Coopers GirAs (1954) essen ectesccnace sence eoeeten 31,110,112 
Gooper:sG AN G1955) hesesaweeseceienejosace ison peeeeeteeeremee 25,31,82-85 
Cooper;(G.sAmemal:(1942)) 1. crccascesseenecoceac tee backer 6 
‘CooperandsWilliamis ((193'5)) mesccsseeese-eee ase eee eee eee eee 10,95 
cooperi, 

Raurorhynchamessenccesae ere 20:40) 2225 22-24 82,83 

Siphonod ellavecies nia saeaetedscucnteas sons esteee toe ee eee ee Pi] 
Copper: (1966)! see. ss. cess aeeace evewcceescvsijacees se beneeeh deo eee eee 10,86 
Copper (LOTS) eessbs ose eS cen acne odee acon eee ne ee 87,89 
Coralifaunas Frasnian)s202-..scsee-cs-neens-eceseteeeee eee eee 15,19,29 

TOWay so-cccrte eee ak aes 


New Mexico 
coreiforme, Dielasma 


GONMULQ | Gry DIGI were acene eect ane teesete renee 
costata, 
IDEAS CTO AOTHE sonenptea 25068. 6en68e00 “i8209005020068850- 
Strophodontas 202. cseeeses se dacescansssecices ce sodaeessene ss eee 
Costatrypa ‘Copper (1973)) <c.casce.+ccevsee-ocosseeeoe one eceaeeeate aeeeee 88 
CXTENSA MISPe teeceee eae -e ee - soe 23,24) coh. 18,19,87,88, 89 
varicostata (Stainbrook, 1945) ............ 23, es 15,19,30,88,89 
costellata|\Gariniferellavmrsacaasseareaeee-ceetere ee eee 3B eo 30,41 
GrandenarHalltand\\@larkei(1893) meeessosie- ieee eee eee 119 
amana Stainbrook: c2vecscc2aseeacceseeden-ee lanes on -eoot eee 119 
calvini (Hall and Whitfield, 1873) ......... 38) 4562 19,30,7178, 119 
navicella (Hall, 1867) 38iec8 20,30,119 
SPs Di aces seecenseescrcehenes op ieeeeucse -ausecaeccaseetees eee 39 ES 14,119 
SP eh Qe ceieectaues sews de vephacweaceces suck cv ncce cae se ue noe eee eee 19,119 
Cranaenella 
calyint (Halland) Whitfield) 1873) )\<------.0----22-e-eeeseese sees 118 
navicella) (Halls, 1867) xeecevescvececvceceescencet coe ce seer ee ee eeeeeee 119 
Graniavourayensismkundlei(909) eere.se encase eae eeee ee eeteee eee 34 


crenulata, Siphonodella 
crepida?, Palmatolepis 


Crickmay \(U950)» sccegesewseseassass cistiicescseoseeaees acre te eeeeeteoees 
Crickimay) (1952) 22 cecaerccee ois at Sere acdsee eee eee 
Crickmiay.((1953))\ e.2: Heveeeese A hedesecest eatoniendom oreo 114,116 
Grickmay\(1960)> svscacseneeis deere dacesatdectestaaseeceeen eee 115,116 
Grickmay) (1963) recesses acme eeecersre see ee ee ere 60,61,70,77,78,96 
(CONAN, 1s (428s paoecmbangascasaseasnoassoso9e002e50C 21,22,92,93,120 
angelicoides (Merriam, 1940)... shuseaanackeecse tease ee eRe 92 
devonica (Raymond, 1909)..................2..- ee 92293) 
humerosa\(Stainbrooksy 1950) peeeessee-snesee ees sees ee eee 92 
prouti (Swallow, 1860)) spsecccsenseeceeenecc-ceeeee eee eer ee eeteceeeee 92 
reticulata (Stainbrook, 1947) ......... 26,45 es 21,22,92,93, 120 
Grurithyris! George: (931) cennes-ceesaecesceese cease acta aeeeeeeeee eee 95 
Cryptonella 
calvini Halltand) Whitfieldi(1873)) scree serac ene eee eee 118 
eudora Hall and Whitfield (1873) ..............csceeeeeeeeeeeeeeeeees 118 
GUILrijUGAtUS PRANGS DINU Clareecceer de cenceac eee cre caeetee ease sees 101 
Gupularostrum Sartenaer (1961b))<2-----..2--2-2-e 10+ tosses eee 70,72 
contracta (Hall, 1843) 70 
Saxitilis (Hall, 1867) scsonesceewestore docteees cote tee cece eeeeee 70 
Cupularostrum? 
Spi Wececsscecancstsen cc cneses ease natin en tenae- cee eee 14) cs 14,70 
SLEVENSONIOM A SDaresecceeeeseaeencercadesscae 144i ee ee 20,22,69,70 
curvata; Ancyrodella ;:.2cscccevescncwesesnevesscssauaeaeecedesetoeenate 28 
Gyphotalosiay Carter (1967) Werecrneie-tece en seriece rece esser teeta see 61 
Cyrtina Davidson (1858) pecsscssaderctecseteaseere sce eeaee 18,111,118 
inulta Stainbrook (1945) .... STi 15,18-20,30,//7,118 
SPi Indet. © ...ccssSeescsvisesased seman oe enadaces cecasnecrcsnaaseen ese sees 22 


VArlaDiliS Ne 'SPracse cases snes ectemencecenstenearseeese SVIESH) coocce 14,118 


DEVONIAN BRACHIOPODS OF NEW MEXICO: COOPER AND DuTRO 205 


cyrtinaformis, 


el ahy racial ee tecn se seagate ean cela a ssc oso cts s TUR Gow RO EE SRS 
YZ ARL! Sonace space nRCOOSE pO ce ere acce pa CEBSConTE GHC SGCOSEE RH CCERnEeaacrac ee 
TAORCOQ DIAG? cosnappsbosesoodeeeacbacsce: 

cyrtinaformis (cf.), Tenticospirifer 

Gyrtiopsis)Grabaul (1923) eese..-3-2e ace -2oe 

(GY PHOLt GANS BEN en cemeeiracecesena- censsccchs eek srenere ce eeea sone ZOE ; 
kindlei (Stainbrook, 1947) ............... SVM cooceg SII NN 
monticola (Haynes, 1916) ... 111 

Cyrtoclymenia Hyatt (1884) 23 

Cyrtospirifer Nalivkin (in Fredericks, 1924) ... + 21,105—110 
CA ATOA OSES (GTB, VECO) saacosesscacacceocstoncoosbaanessneeBansaccae 104 
UTES UGS) 0 ponb none r OAc OACOnOCOOUEEEGETDAEEOEES SOs LOS 
breviposticus Stainbrook (1947) ...... 29 ...... 22,105,106, 107,120 
ef. C. thalattodoxa Crickmay (1952) ...... 30) .i28. 19,107,108 
CONVEXUSIN IS Pvt rcins so dec seaseee--22-5.000 31 ...... 24,106, 107,109 
kindlewpstainbrOoks (1947) es ccc.ecnasees sees eee eee eee 111 
portae Merriam (1940) 105,108 
Posticusastainbrooks (1947) i sses-ncne. ose ss meen se see eee eee eke 107 
JESIICATOTALIIS Tals, Ss padceosocaosecusssodesudone See E22 OG 1OZ. 
ROL CG TAS! TNs 6) 2) seep ecsberbenicopoasocbececBeece tone cbcercs 30 ...... 24,107 
S ean Mee ee eo sia cise scsecleosceane ies seca na Seana Oe aS SH eaccen ZAR 
sp. . 21,108,109 
SPO Meso Senco secu satee soos sees seaesi on sicheeee seen uO cokes 19,109 
SDRAM reer coce eeces oro ch oecsececduns SovesiasekeseesseGess, . OL ccnes 19,709 
sp. 5 19,/09,110 
sp. 1 Rte selre acco ose seve soos se ninctriomches tee ccwiies Sues boaations oaeneceene 22 
thalattodoxa’ Crickmay; (1952)) ecs.s-cs-wee test ecco eee 107,109 
WRLC IA ELAS SSS8)) ccessesesecerseccsen sense see 30) 035 19,20, 1/08 

Cyrtospirifer? animasensis (Girty, 1900).................. 24,104,106 


GyrlospirifereZONe) se---.4.---.-4- 
Cystodictya Ulrich (1882) .............. 
anomalotruncata (FritZ, 1944) .........cccccceeeecseceeeeeees 
Al mMABN LSZS) Meese meee ce ners con cclcw were seeas sens ooacstove eet een eeecee 89 
ID ATCON IN MELE oP ae ch aclcc a rsh <tyacsetecsktnecstenst teseeecare 31,142 
ID Fiona (ICI) 6 Scand SCO pee eo CEERE REDE CEO GADRCECEEEE See aE pEanErEr eRe cas 8 
David Sonn 858) er satess ooede sca versecosces ices doesewsaestesese se 18,111,117 
DAWSONED AYRE ORMAION. wen eee- see cease ne ecacdeee saeco te ceercer neces 60 
C2CORIHIS Ro JOM VTLS cosccescuasceoscoosassoeseocoosecsuacouncecs Aap ds) 
deeringi, Strophonelloides <...--2--2---ce2c0ccaceazeeess 5) =encee 30,45,46 
defessa, 
PAU OMIM ee are cn Nena Seth et eee ae stiooe cece wane ent eces ee ESE se naeeeeee 64 
HPC LOD. OGU CLUS Riera Tete ne dete cen eee nase SST ee AE ERC RERC ER EASE ee 24,64 
ENGAGED OUVIILINATIG: sesnee secre na cace cee aseamer eset cence eee 53 
demissa, Thomasaria ............... 33,35,39,41 ...... 18,19, /03, 104 
ETA CNSISHEP GHINOGOEIIA ie eee eae ee eee 95 
Me navalli mes tone sank a keene ce o-oo es eac co sacaccbeasctes aeeeeteee 60,72 
Desquamatia Alekseeva (1960) .............2...c0eeeseeceeneees 10,11,87 
GOSHEG No S05 tacdecececosenoseneEeeeceGrUnocene ence D2 sre: 14,86 ,87 
Devonalosia Muir-Wood and Cooper (1960) ......................4- 61 
wrightorum Muit-Wood and Cooper (1960) ..................6.- 61 
Devoniant rene sees tse oe 6-8 ,11,13,17,25,45,54,67,98, 120,143 
TYG TRE) Greondciseaenscocded eOnceSGce CUE E EES EeE SESSoeOSEREe costes eeBERrrAapode 33 
Deadman Canyon, San Andres Mountains ............. Bitte 
Middle, 
eastempUnitedsStatesi-.-cs-------se--- 11,14,32,42,58,82,94,97 
MackenzieaValleyrrs:saccecst-ncca ces coer eee ee 95 
IMIG Gane pe ens ea es Soo ese oe se See TST ESTE To 14 
IN@Vadlaeeseesse nanan fi tae rads oie fone setasccde ioseneern ee cece eee 14,72 
SacramMentomMountainspres nese s ea tetesesaaneemenecreseoacmasenese F 
Sheep Mountain, San Andres Mountains ............... Cas 


Upper, New York .................222....... 32,42,44,63, 105,108,109 


devoniana, 
ANTI DG svaaels ss ca clec ess Jods css eaeoee es saule valsiouate uae do dede sees nee abeeeaseenee 
Pseudoatrypa 

devonica, 
Cleiothyridina 


GTINIS ONIN GES... scenes s coscseecsaeeseceneaeees : 
Devonochonetes Muir-Wood (1962) ..............2.2.0c0sesssncosessees 57 
Devonochonetesnaspun-ycuncececcnasseeceteere eer eet LO} econ: 14,57 
Devonoproductus Stainbrook (1943) .......ccccecceceeeeeeeeeeeeeeees 66 

hallianusn(Wealcottw884)mrrecssscenceaeeca sneer eeee cee eeeee 66 

MAM BAGH ING SOs ceccscacosoasecensadsaaacac 13) 2253: 15,18,19,65,66 

PURE ARS LENE LD WYO) < os scoganoscocerbeace Saeneaaszacaocuatecesnoe 19,30,66 

walcotti (Fenton and Fenton, 1924) .................s0sceeceeereeeee 66 
dialataSRrionodonidwerecncencerceeeten cee ne en eee eee eee 28 
Dichacaeniagngce lees seereescesteeescectencena teen 10,11,14,60,6/,62 

DEMPIEXG! NSP vaccsenees cesses sass ses eeceeeem- sae 12)... 25: 14,60,6/,62 

LEI DONALARIGES Deen ere ree nee nee eee 12) anes 14,6/,62 
(2)Dielasma calvini (Hall and Whitfield, 1873) ..................... 118 
Dielasma coreiforme Stainbrook (1947) ..................0200 93,120 
GIBNALA WAIPY DAs sasciectisten cee secae tae tee hex See ITe Sed AAR Re oO 87 
Diplodellaisp jr veces treason tacter ase eos eoe eae ona I aOR 28 
disjunctus Var. AnimasensiS, Spirifer ........10cc.cccceeceeeeeccesens 104 
diversa, 

Douvillinariaipessseneeee eee 19,53 

ELLY tha eiomoce nes soceeen ce ee se 116,117 
Dmitriagsidiachenko) (196) meeeeeeeeeeace ener ere eee ence eee eee 111 
dorsoconvexus, Strophonelloides ...........00.00000e00ceeee $,19,30,45 
Douvillina.@Oehlerti (1887) eereeceecseeceeee eee eee ee 20,49-52 

aff. D: cayutai(Hall)) ic. .cosecssaeanauesees see sceese aoe eee 20 

arcuata (Hall, 1858)...... sae) (Sleweoss 19,30,50 

Cayuta (Tal 1867) sc ecsece coun cessectecavesecess en senseeceeer tree eee 50 

COPHACGROTAGNW WW, QDs. sosesasacncocsosouesnsopesss0o700000 8538) 0-c 50 

MINION. SPs. 0cccesees cose cececssecscese saseec ee ha 8:9) 5.2 19,50,51 

MaviculayS tainbrooki (945) eeseeeesee ete ee eee eee eee ee eee 51 

DORIS OTNA We G2) sseusccescscnescooanseacacoeac6ec Oe IOneD 15,19,51,52 

RACHA GTA WS SDs, cneisconasnegcoasonede90se9000000000 Decca ses 19,51, 52 

sp. 1 D ccccaes 19,52 

SPii 2) Re tewciee cemesesascumcmnee de: Coc eeet eee naeeeetee aac tasersee 8) sseccs 20,52 
Douvillinaria;Stainbrooks (1945) We seees see cases ceaeeeseeee eect eeeeeee 53 

delicata) (Rentonyands Benton sl 924) pees nee eee eres 53 

(Co ASTA ACY  Vegto) 2) a ecbecaecuasereEoceatescnsececoanacscacane D910 ice 19,53 

penversa: Stainbrooks (1945) seesseeenenease seen erences see eee eee eeee eee 53 

variabilis (Calvin, 1878) ................+ peat asec 19,30,53 
Douvillinella variabilis (Calvin, 1878) ............0.c0ceeceeeeeeeeeenee 53 
Dre WesiUL96 1) oss. scasuanaceas sahh a suececeeretesuee soe een eoe See eRe 17 
Dunbarand) Condray (1932) pescescesseccoscesecese nce eee eee eee 33 
dunbarense wlrifidorostelliummcanescestee eneeeene eee eee 79 
Dutro; Ue TICs cascct sagen slope cts.csdeesees sees eeae- ee 31 
Dutro Nie Re, oi ccs sacedes ex testens ec cecaaee egos ees ee eee 8 
Echinocoelia Cooper and Williams (1935) ......................-. 10,95 

ambocoelioides Cooper and Williams (1935) ..................-.. 95 

denayensis JOHNSON (LOGO) peameseaenecatee eee eee eters 95 

incurva Cooper and Williams (1935)  ................se2eeeeeeeeeee 95 

PL CLIOSG’D. SP n eeecseneens ese oa see ss ae essen meee tector Ot fence 14,95 
Echinoconchus S. Weller (1914) 66 

laminatus (Kindle, 1909)............. 66 
eclectea, Warrenella — .........02.0.00-06+ 116 
Eleutherokomma Crickmay (1950) ........-...2-+s0eeeeeeee ee 10,11,96 

aechmorpha Grckmay, (1963) seneessss-se ace oeere eee eee 96 

CXLENSA ESP eee eee eee eee 

reidfordi Crickmay (1950) 

Jos. cap satenbseanoasdocnocasqcanodasadaoganposboccoscoHSaeCaCueb: 

Elicto gnathus’ Spitec.cccecs ec sess econ tee se sae sewee Sea can coeds cote ence 27 


206 BULLETIN 315 


Elita \Predericksi(1918)) ccsscctseses- ce oteeacecoseeosesesres ence -oseen cree 117 
inconsueta (Fenton and Fenton, 1924)  ................0..0.e00es 117 
MEAPISTEF MASP co-c cose st ee see sete eee escce ess 3672-8 15,19,/16,117 

elliptica, Aulacella ............+.. Bes, sa Ose ees 15,19,20,22,38,39 

ellipticum, Colophragma 16-18,40 ...... 20,77,78 

Bllisomi(N95SO)) aeec2sscdsccc carbene o= odes ondecs Samcuees te seuassessedesesenece 27 

BlPasoGimeStOne: 2.52. cosex dies cece cesses sccee-eaeeceseseser se seeseeces 29 

Elytha diversa Stainbrook (1948) ..............-...2sssse-seeeees 116,117 

Emanuella Grabau (1925) 

Gt iy aeeecberpocenncrReEneenp acacia 
subumbona (Hall, 1857) 

“Emanuella”’ meristoides (Meek, 1868) .............0.02200ec0eeeeeee 95 

COTURD IS, EPROM AM TEHAGT — sscoconscsodacocanacocanxcecoononenepoascceccens 763 

emmonsi rotunda, Hypothyridina ....... 15,16,39 ...... 18,19, 72,7. 

endlichi, 

(CAMANOLOCCHIA re wssace onde oa see soek eee ee nee deen tendo one ee ice seee eee 82 
Camarotoechia (Plethorhyncha) 82 

England 22cs5:teic.cevevcnosecerce esses cece tice 14 
DEV ONSHING: Sircsteecsecenescicnagicces este sets ene dene es awed omeeeceeee eee 15 

EQS SIS ITE (STAD) sececescacnop9gnccenasconncqencozooese0c000 117 

Eomartiniopsis Sokolskaya (1941) ....-....02--2c00--e-0scee-ceseoses 117 

Eoparaphorhynchus Sartenaer(1961a)  ......-....-.-.-0eeeeseeeen ee 84 
mclaren Savtenaen (1 9G)'a))ne.cseseeeeee soe se ete see 

Eoschuchertella Gratsianova (1974) 

Eostrophalosia Stainbrook (1943)........ 
independensis (Stainbrook, 1945) ............-.-:.ceceececeeceeeeeeee 60 
IMEXDECLALANN. *SPostarsescoseececeseecetaece nose oaee eee | Cece 19,60 
peddermG@rickmay, (963) saesseeacc eet ete cee eeae nese eee 60,61 
rockfordensis (Hall and Clarke, 1893) ......... 1 Pease 30,60,61 
SP sill woeseaetheacaee soon sesso de Rieah oe atestsaiaeeee cm aerersosaneeoaestes 14,60 

Episier ale (L957) actecns suc voseseees-custeccescencdeassceteate seaceesesenece 29 

erosionsilate Devontani ccecescssosecesccecteneeetees 6,10,15,18,27,29,30 

Cine, SHAQ COGLTTG! GnasoondsccosodonosescunbabaosoceDeqeooedeAceGor 44,45 

Bithineton (L965) eres sete ree eros noe core caeen eee een een 27 

Eudora Gry DiOnell A jccescsccc- dene a encsh ox sasscactoesdeet Tneseeenocereeee 118 

CUTS TIOS IS) 0111 RA RRC DEOPO REE BCEIOR COCO ORE CCDC OPER COSE OO CORCIOHOSOAIaE 113 

CULINUSHTECDOSILUS OS DINU C)> evecare seceesseseaccece satectereseeeereee se 113 

euglyphea, Psuedodouvillina ... fis (Ol Saccee 19,30,49 

CULEKAENSIS= LAGFOMAYNCHIG sees eoneasescoreeeeesenasee eeeeee eee 72 

Evanescirostrum Sartenaer (1965)! <.2..-.-+-c2-c-ns-ceeeeees 21,22,120 
LA DTAVIEMILY Decca eect teen secret cen Sete nee INNS. eC ee 83 
glabraventrum (Stainbrook, 1947) .................. 1) cased 22,83 
Sty TO acecanccaacecedea-cosaboSertaee ee aaconeancruaerecocbacck 

exfoliata, Indospirifer 

EXLIS OSTNOPHOGONI Ares sccene ase ener ate eee 

extensa, 

(CONSTI AY QO merce ere ER EUBOG ION IOOCD OES ECLOCER 
PL CUIN GY OKOMNITIG! Rat ens scoehce erect eee eee eee Dh Ane 14,96 
INCIAHORU TO DLHIG?. tasecseroanadosnecseciscncdbeseooscegan 639 res. 19,30,46,47 

WAIELI COME CLF OCTONIG eatnossers tn chen ote oe ne ee 34 

Pamennian Stage: :....-00c2.+..00s05- 20-24 ,26,27,29,30,54,58,92,110 

Benton (1919) ieee a. etek eadneencacene a eta ee eee 118,119 

Rentoni(lS28) i" eaine-cccesestescuesseeseensece ete ee ee 3i7/ 

Rentomi(l93i)), o2.--<teseecsacct of.cbs onc etsos saee ar scene ae eeee e 113 

Fenton and Fenton (1924) .................. 37,45-50,53,54,66-69,76, 

87,90,112,117-119 

Rentonraudiventom)(1930)i.csrsssecessescensee eeeeee eee eC e eee 87 

Herpusomi(l 966) 0) .vscecccevsaseeccs. vee eiss eee ee ee 84 

POWER VEL ieee sat rec sesconeceseectetee Sees ee 7,26,31,54,95, 140,141 

BIOWERLGSS) ire sixes ssursencwwecuacatesecescasccece nee eee 8,19-21 

BOW PIA) lated sores tacos sanded neces iw as oh ert ee ee 8,19,21 

Flower (1965) ......... sie SeRO esas vegies acucscs eRe nae eee 8,19 

Flower (1969) ......... Bea sdersenatee ki Rea eB 19-2] 


Plower' (970)... \a.ccsssesc<te03 cosoneseeecpep epceden ee cee eee 8,19-21 
flowers Warnell ssees sn. s ees ee 42 AS eer 11,14,//5,116 
PlOWeriaitnQemss wsctecc.cecces ocean costes toes 53,54,55,56 
aff. F. chemungensis (Conrad, 1842) ........ Pag Ui arenes 20,55 
Ghemungensisa(Conrads842) i eseereeetes ete eee eee eee 55 
IA ONACICAINUGMeSP wa cessseeeaseesntesesere eters 1Oses 19,55,56 
prava’ (eal i858) easces se ssene sen eens secre 10) 225258 19,30,54,55,56 
PANSVENSAS HM SDa see acnce ee see sceteaee ee nsee eee 10uA 30,55, 56 
jfranklini= \Warrenella” cce-cccecnesctcoe le chee eee 115 
Frasnian Stage .... 18—22 25-27 ,29,30,66,113 
Brechi (897) cise sccazads cc acsnecscccsecscnsstsctaet ty aero ee 44 
Fredericks \((1912)). cccccetcessce.cicccsecesaceve-ete soot ost ee soe eres 117 
Fredéricks\((1916)!.. sseesz:cee.c enn ce eenee tec spanseseesceateere ee enero 100 
Fredericks! (1918) iti xcenses-sesessseeecocee scot tet tate etree EEE 116 
Bred ericks' (1924) ie. 2: ccconsetaee te see teeeeeee eee 21,93,104,110,116 
BP ritZ (944) 9s ic cwees 2oke aot secon oh tees tas ese cet oe ees ee 14,25 
Fusselman Dolomite 10,22,25,29 
galeata, Galéatella) ccoc.c. ocprteccsescteeeedex toes ok oe 24,65 
Galeatella Muir-Wood and Cooper (1960) 
galeata Muir-Wood and Cooper (1960) .............2......0065 24,65 
geniculata, Nervostrophia 15,19,46,47 
George (193'l) ee eseecteeee cesses 93,95 
Germany 2.4 oesccsssscaanensee ie ace es saeoeterees 4 14,101,143 
DaSbut8 wae cessacea. Saeed scdsear ss sea desucesvecoueeasseoneus Se eeeeee eee 23 
gigas, Palmatolepiseticccosesss Goede 2 sees iene aes esse ee 28 
Gill (UST) ica cseasemansinenenes does Pens aesbace ees ass eke ea ae ete eee eee eee 86 
Girtys' Gu A, 2..scssccarcueatcseencce teed. che ee ee 31,142 
Gift yi(1898)) fencsSoessetdeessincg eased aaacises sxe ceeneetteeee eee ee eee 33 
Girty (1900) .... 
Girty (1904) 
Girty (1939) 
Givetian: Stage: sc tesaeeeeb owes See 14,25,27,74 
glaber. Poly gnathttsveks ccc xc: 2c o0soss2sn5ee-taseese er eee 28 
glabra JPalmatolepismmasererstaeter caterer eee <a e28 
glapravacuid weal piatolepismecee serene ce eee eee sees 
glabra glabra, Palmatolepis .. oes 
glabra lepta, Palmatolepis .......... ages 
glabra pectinata, Palmatolepis .... sen 28 
glabraiprima, Ralmatolepis: secsssssccesee aces reece eee ee 28 
glabraventra, 
IEVQNESCINOSITUIMN) Secskcice cos soce secs odeocoe se eee ee 83 
Shiurmard ell) ercicccsccseawck sc ccvecev nase ooenecee eee ee 83,120 
glabraventrum, EVaneSCirOStrum ..........00000000-0000+ 19. 22,83 
Gnathodiusispseaeeereseseee ree Bee 24 
Goldfuss(1826=1833)) 355.2580 oecde  soceccc nee ooeee eee 10 
Gordoni(d S07) i Sc. acne seecec ae ec cette aoe ne ee eee 8,22 
Grabaut\ (1923); ssssaseaccadesseedeedewaeceaceses sagwae tess eevee ee beo reece 11 
Graba\(1925)). «aan. <ceweeeses occas seceee coescca doe aca 95 
Grabaui(l93 1) \..cccascencnenscasseesorc so cece eee 97,112,120 
GiantaWlRen Bye. 2 fce tesa cere saeco eae ee ee 125,138,139 
Graton, [bsC.. sssc.esceceth eases oe oe coco 31,141 
Gratsianova:(1974)) _<-sesskcmwescsqace socan- ce eee 54 
(TENT OY (Oi prossessecocecoschetebacsocBcsernetennetaeatoascnd onssaGaGacKa 33,62 
Gray (1848) oct shes cesctsessenc sanscorse see ee oe eso eee ee 69 
PreZAariay AMDOCOELIA acsneccssaessdetseenece ne stae eee eae 94 
Greiner((l957)) Feces sacesoeaedoee te cetce ee rere eee ee 105,108,109 
Gruenwaldtia 
americana) Stalnbrook: (1945) see-seeeeees cee eee eee eee 89 
SPs Gav ensen sneer asines cecfecn secure wetee tees eaten aeteece acer eases eee 89 
Guilmette Rormation ).cose.cs. aces ccaswoseos se cern eenere sae ee eee eee 17 
Gunich (1896) eevens.cctheateeeesecevens ccadenansss seeedewer meebo eee 19 
GypidulawAalli(8Gi) eae acces tee eaetion eee eeoee eccrine 67-69 


bowsheri: NSP: stsseces once tact eseav ieee tate 135140 19,67,68 


DEVONIAN BRACHIOPODS OF NEw Mexico: CooPpER AND DuUTRO 207 


Gypidula 

GOMISE( Owens ml 852)i-etesecstede ac sh oreo secede serene eee 67,68 
cornuta Fenton and Fenton (1924) ......... 14) scte5: 30,67,68,69 
LOtisa (WealCOtt lS 84) ve rewecose~ ssc uiseseue sapecon ese orotic ben econ: 67 
TPLEIILE GOMMSD eUaecinajaccsen foaine soso escseceaheneesc te epee 19,68,69 
Fatraater ((CANGING 1h7k3) ccacocradepcecpaaeonee a cneceaasneeteetcncnce-ee- ee 67 
mit Da czania {ie cteieavewss zacenawongs Se SRO 68 
14 ...... 30,68,69 
1 Se oaceer 18,19,69 
Hackberry Formation .................. 8,30,55,81,87,89, 118,119,121 
@erroiGordosMembene.ces-s-----e eee 30,31,37,42,45,47,50,54, 
66-68 ,76,81,87 
SO DHOMEI ANZOUES tae derctacaccasascemseeetencoseose sess enba cocateorsees 60 
Owen Membetirecs case racemes see ccs tc soe aenenewnsoaeek sanuscecnaaeeweds 89 
Hadrorhynchia McLaren (1961) 34,70,72,74,75 
CLILARIERSD eleee eee scc a0 <cajfovacsectssassi cases act HOBIE codecs 14,72,74 
CULEKAEISYSH ORNS OM) (1 9G6) In sseserene-- sence see saecceceter te ne eeeee 72 
RMLGIALETIBNSP tice =o eoaeescescesaceets se sceuet 7, oe 14,72,74,75,76 
Sandersonii (Warren, 1944) .c< 2 2- .c.2s-deecseesesasenesscesee 14,74,75 
pT ACE OUMY TIC Att cere sacs sscasewsc eset casas Sea wasnoot es sana anaee oe 14 
Ffalli(843)ecce..---ee ..  11,48,58,70,79,97,98,101 
FLAN USS) Betscces gates anciesesecceceeinene staitinnssaiehsiecnnseeeeeneces tes 44 
Tal QUSS2) owe meee cecisciececosstcncactent ne dete Sevens veseteecte eka tale dese see 93 
VAIN GSS G) Premecree nae corcareens sxc oe ste eskns co teee ak Rasen ee 93 
IS GUI! ((TEISS7/) -ceiesdeoceecaasecesoe eaacence: Ean ae Ree eceenen 42 ,43,58,95,114 
ISA (GIES) Sooceeeosecasescenee 18,20,35—37,50,54—-56,68 ,99, 105,108,111 
ISAT (IGISUED)” Gonscesacsaseeesdoneenceessaaeone 10,34,42,71,79,81,82,93,94 
EL AUIS GOD) Meese eee oan secant oseescnscen cc uscdsseonees concen eet ons eae 32 
ET alli((867) Wacssecs--=-25- 14,41,50,55,59,62,67,69,70,74,79,82, 108,119 
Fallfandi@larke(892) i ire-e.ne-2-cc-.ncseceseseaeae oeee eee 32,37,53,54 
FfallFandi@larkes (1893) 5 cnc s-c0-2e-eeeceesceeeeecsosees 60,61,79, 108,118 
Hall and Clarke (1894) 67 
Hall and Clarke (1895) 119 
Erallvandmwinitheldi(872)) ete sec seca ce nwonss doen scsnace ne naee 18,112-114 
lal fandaw hithield (878) ie. -e-s5-<- soe cecenesetee es -eneee 45,47, 118 
lal Wang awh thield (Si) esese: -caeceote cseee ene serseneeee steer see es 73 
RalianusMMeY ONOPLOUUClUs) cone seresteasees cen secncesdeteseeeeeneet eee 66 
fami toni Gronpieeecestcsseces seco cee eee eee 43,44 ,70,94,95,97 
Elarep Giannis Dal@w cates. ccc 22k occ eee ac svncsuaseacvewaes<cnsctasceeeese 15 
amiss At sicce ssa: sess eeee: 8,18,25,26 
EU cES NLS DR REE toons cc ticidcia ocean ve coh nose woatece daviecd Sean eae 27 
In ERalltey < ((IRLSAY coco ase ce peor enGuS Sco Se ne COBH BORO IRR R Geers sonassencor: 54 
PAVESI LOI G) Pace tecscctc cana tseacceesc see anu cceascassvowencscorcaseeenees 79,111 
HaveRiv ere ROrmationm! 23; .c-.c5<2.05 evaseac0see Soccss sncceeeseeeseeenee ces 36 
ELE LN Sy (1963) Beeetees se ce nnnoctewes eiiasaise a scnn whe se ad eeeeee tee sae SnOSS 28 
Heteralosia R. H. King (1938) 61 
nupera Stainbrook (1947) ................ 63 
Hexacrinus Austin and Austin (1844) ..................ecceceeeee neon 24 
midexaponarianGUtichy (896) esec-5-=--ssnceee en cesansaes eee eeoee 15,19 
FFUID DOLGELIGESP tates aren Se coe ds savie olson desfpoo.ciasdels sels alcs sabe eee ROS 26,28 

hillsboroensis, 
RIAN OPY OAUCLUSWeroxse cen esa ee oneee ies sakes ose eenae eterna LALO) 
IBV OGUGIEL AiO eescea seen aso de soce tae 5 sce ae oe Does LEE SEES Sos ROS EO 65 
TING Og (US /9) Mreecmsceciocecs nev cosecsccsecsecs scwenee 25,28 
HAI NAECAECILARSP wate ete aecets asst ccceeerec hivies costs eee eee 27,28 
ELES PIC AY ICMU Ss BED he aosan sone see cccese ace Sabet eeesscc eee 24,110,111 
PosterOgranulOSUs DN. SP. ......ceceeeeeeeeneeeeeees .. 24,710,111 
SPIMOSGMMS SP eee cecccsu swe saswasescesedesowssetesees S2iencce 24,110,111 
GOERS (912) orecnee ee occ cicacecen see ce elceccsunlasecssesupccuctonesersescae 33 
VOLES) (928) Wrenca veces secs cee <ccatcdcaceecscacwct ss scessachenscossecessaee 28 
NOMOMNTESUIATIS, ROL SNAINUS) --.-<2-ee- ccs ens: - acess <r oeanca <2 26,28 
ET OOV EDGR AIRG i ceecert cosnecet suc sos woek lob anaceessastivecesscd ses coed acecee 8 
ELOOV.era O81) Preemcceasschee cece ctne eat ac ecsena pack oanseaecumocsaenedcoseree 30 


FHOUSEs (962) Meatetecea ces ceccte swat cass conn wesc eotan cet eas eae ee 23 
FLU les W a cesscnerccwocesns sreecoves see cee aeeeees 14,18,21—23 ,25—27,29 
Pd dle) (1934) Whe occmseseasewcsvcordtaess concsee Marans noses oni ak Sooo 25 
humerosa, 
Cleiothipridlinite dc Wrasccss cc ss nis Sacesie nat coeser see oot aetna cac ene eae 92 
Grins anim asec ris coccorcs sseee ee Sessa 5 onc Seen 92 
16 Ub EAA IESE) S aeacecocdenccsocnocdeCe sce GBDCRDSECMRACEREOGUBESEGcCCAEeLEEE Riaee 
Elly ate) (884) peepee saeco eee cc oe ran ae sees sece seen 15,23 
Hyborhynchella bransoni Cooper (1955) ....... 20,21,41 ...... 19,82 
hy Dridus sp SIrophorne loidespscensesseeeseeeeeeee eee eee ee ES 
Hypothyridina Buckman (1906)........... 11,15,73,74 
CameronimWarreni (1944) ocr: pease sca satan seeecoe te eee eater nee 74 
emmorst (Hallbandawnithie] dial Si) meseeeeneeneteeeteeeeenaee tees 73 
EMMONS FOtUNdA DN. SP. ......0e0e0eee ee 15516339 ease 18,19,72,73 
(ARAN NO CLAS (WEARS USD). cccocsceosenscobaceccnoneecncocscosocne see 74 
SD Biase ecctesiccscwovcesh ote scercseececeseneeeseessmeees 1S-23 Sees 14,73,74 
Spa BiMclbarentandsNomisi(1962) esse seceeceeeece eee eee 72 
venustulay (E1all P1867) ec-ceorcsceecenastese-seocee nce caer eee 74 


Icriodus Branson and Mehl (1938) 


nodosus,(Huddlew1934) eee ese eee ee 
sp. 
Mlin@is Sess ces. cwessaeeseetncdnc sesake teases eee neoseee 
Imbrie (L959) servsck. ocseae ect sats cw seccsees eee seaeen ones 
ipipressaaOrinisiGS ChizophOria) meena eee 
UNCONSUCTGS ELitG oer seen sess enna sence ee ee 
INGUIVO PE CRINOGOEIIQME caanereeter ene e ee ee ee 
Independence Formation (Shale) ...... 8, 18,30, 31,35,36,40,42,45, 
47,48,51,53,55,62,66,68,77, 80, 
89-91 ,102,104,114,117,121 
independensemSKEnidiummecraseese eaecee eee eee Ee 35 
independensis, Bostrophalosia: .-..:<2-..++00ss0+se+sasseseeseavers son: 60 
Undianial ose sete nae eee ce seats eee. caeseees sacseasen eee 25 
indospirifers Grabatl (93)l) meeesecesese saeco eee eae 113,120 
aff. J. orestes (Hall and Whitfield, 1872)...... 334.25. 18,19, //3 
HUONG Bids. cosoeooasocosadancdscseecs00000 33). 2.2333 ISSO SIZES: 
exfoliatawnees Dues oee ee eee 66) poses 19,713 
orestes (Hall and Whitfield, 1872) ..................02.0.0.00+ 113,114 
SHI ZOSUSa (Meek S60) ieesceesraeeeeeee seen eeeeeceaeeee S3i8A% 113,//4 
IMexPEClatasHOStrOPNGlosiaweraaesesseeere ee eee 12) oes 19,60 
inexpectatum, Afilasma <2...1-c.<n0---2eceee 38,44 ...... 22,//9,120 
infera, 
Aulacellas.: iit a ae ee ee I 38,39 
OMNIS 552.532. Jacket ct seaecte ce saeenias Aas 39 
ISZQINDIOOKIGHcwe ears eca sense aaaee te eens DY) somes 30,39, 40 
INOLNALULS OLY SATUS meeeeent ea eee eae eee eee 27 
intermedius, Devonproductus .............-. | ecace 15,18,19,65,66 
inulta, Cyrtina it e8 37 wees 15,18—20,30,/17 
Lowa Aesth teens mere 14,18,19,31,35,40— 
42,44 ,50,51,53,54,60,68,77,90,91 ,92,102,114,118,121 
Brachiopods) swesseceteeoshs cnecroreonch qeaseetadon. ae. disceaeee aaseeaseneeene 30 
Brand On jsvier: 3. -.esecscseesshassussnel bee ee eee 48 62,69 
Independence; <<:.cs2-0 2 <2 aersacecossnotetse neta 
ROCKION tem a8 esas ace sone ine ce nance Tae tas rove coe ee 
Towatrypa Copper (1973) ............ 
americana (Stainbrook, 1945) 
owenensis (Webster, 1921).............. 
171.2710 VCR) » Saenepecnener es pec ace oaree eoeEeerre noo: ccbe DAY cbse: 15,19,89 
iowensis, 
Cariniferellam wevccr serene es ee eee 19,40,4/ 
Orthis 37 
Orthis (Schizophoria) .... 37 
SCRIZOphoriaw esteem 18,19,30,35,36,37 
Psorthistiozlowskill (1929) eseces-csenensce eee ease ene eee 39 


IthacavPormationt. .co.ctesss sconce se soeater cue icek wee ee 32 


208 BULLETIN 315 


IRVETa CK 1 US) pensemeoceeeocossecoundascocanasococosoondeqearocctaqcnascnéon 99 
Vicha= (1954) i. ccecoc see vs cccesccuea seecens sees oacde see du onaesck easier are eee 8 
WONNSON Iss Gene -c at ete sors eowon See ne soso cere eclace tensenetee 8,72 
Vohnsoni (L966); ccseceek tes caswe eae shee ea sotcho teow Pe ee cms ceemeeatety 72,95 
Fohnsonti (1970). ..ve..scsscse-cscecsseessenscsecsdecces see seesee 10,29,59,60 
TOHNSONIMCU978)) oc ceeseoahes ee. Seseacs once weeceeecte seutiesmrecceeese miteaees 14 
dcorieon Gr eth, (ILS) conpoccocsopousansessocoaseae0 24,83 ,92,93,105—107 
Johnsoni, Rhyssochonetes ...........00.0cces0eceeees THe konaee 14,59,60 
Iashongi shales icc. seco-eccvasccscchientneecnececenenesmecoseesn ss esses 43 
Kayser (1871) 74 
kelloggi, ““Camarotoechia™ 82 
Kellygand@Silven(1952)) (ice.ccss-cessesasa-tee cescece cessteosceecceses sess 8 
reyes: (1908) iiiieocsccetecevacs coca uctaeedanereanausmecsaiactieesdemeseeemasceses 6 
and ler(@1909) peseeeseeee ee 23,24 ,31,34 ,36,56,63—67 ,70,82,83,85,92, 
93,99, 104,105,107,110-112,116,120 
kindlei, 
GY HONING -sencnanececevees<aeee serene ene S24 ea, 2 eet 
(ENT AROXY 1/11 (a secon obobnbacGeepastbaraboonanonsoonooasGaacopacocnAebonecdsac 111 
Kings (Re(1938)" cchcescethed.becdseeceacoag 2 reasdovtaecyetvervecnen case 61 
anges W((1IS46) i scores oe acninesen ed te nsesovouahaneracadouneswevadseasse 96 
IT Tea AWVER( (1135) 0) dae peoneanncc ne noorcorcperesoseneeneanececerapcccs 10,35,56 
Klapper Gen Aatawe sosccccenatart cies sondsaseeae des orouseaeascetee eyesore 8,26 
Klapper er al. (1971) 14 
IReOKen (1889) sae ccessevcncaw acs c sesh sean oe seeeues Sau oe ode seceye ness teeeeeeee 34 
Mottlowskivenal ai (956)esesssenercer econo nce eee eee ee renee 8,21 
KOZIOWSKIN(1929) SP oie cewoonccadecera rene eee ot on oats eave eter eee ae 34,39 
Krotovia 
DFA CUursorss tain bLOOks (1947) aeeeesee-meeeeeeeneceeeetetee see ee eee 67 
ispinigeran (Kindle 1909) oacsesems cone secsecsee eter cree tee ee sseseneeee 67 
UH TN(W949) ae eres ek chee ec ctaas socetors cone coe See eeer on are ee ees OSH OO, 
labrequet, Warrenella sec o0.s2cesecwee secoecnescneeseceraeedes sceeeeteee 116 
Make: Valley Hormation: s.3sesctescecse. eoene soe reece enna eneee seco 120 
Mamarcka (SOD) secccseca snes onc eacens hones seaee. se ssates concer eee ee 15,19 
LamellosaetAlthy rise secese geese ssexseveses cod coe ec ones doc ow enone ee 92 
laminatasLaminatiaawencccccscescee sce ee ee ee 24,66 
Laminatia laminata (Kindle, 1909) ..................0c0ceeeeeeeeees 24,66 
laminatus, 
E-CRINOCONGHUS Se vescxscdecets sxceaesecsint eens HEC Ee 66 
J eLRaye TGS (lsc pee CoReFRCEE TCE POCOC POC OR ECO EEE EE ORE COCHORT.CE ACB 66 
Wane) (1963) im sec zcscaew en aeccnese toss asses eecane anes sodhiseee ae eee ee 120 
lafamSchizopnoria® Weosssccssetesesecoseeeeceee 3338) 25s: 11,14,30,36,37 
latisinuata, Camarotoechia  .......0.0.0c0cceceeeees 19,40 ...... 21,79 
Wealid ony los Rens scere.ostessoneeseon: 8 
audontandiBowsher (1941) ersss.ces-2 seeeeeeeee cee eeeeeeeee r28 
EaudomandiBowsher (1949) ascsoscccssseceosecene scar eeece eee 8 
[AUC OVI AS CHIZOPNOMG we. etesneee see aeaen eeeene e e 37 
laUy ae CAMaArOloeChiawae-texe nee cere ECE 82 
Leioproductus Muir-Wood and Cooper (1960) ............... 63-65 
coloradensis (sindles 1909) esses coon cneses eee 24 34,63 ,64 
ACTESSal(StainbrOOks 1947) pesessseesee eee ee eee 24,64 
PATI AKOTITS Ts 5) )s,. cecootorordbocoosacbocaanchoonons 12513 24,64 
DEO (OST CES ICO) eo conmcphoostossenacobeacnecoscodsocde 24,63 ,64 
varispinosus (Stainbrook, 1947) ........2..s0.0cssececsessecereee 24,65 
LI CLOVAYNGHUSMELANN (1 S60a) le ceesses eoetere eee eee eee 10,81 
castaneum Meek (1867) 
SDHACTICUYN NASP --ser. 95 ek dese eeco ee ea 


lens, Schuchertella 
lepidus, Chonetes 


Leptaena ......... Sees ie reas: ‘ 
analoga Stainbrook (not Phillips, 1836) 
EDL AR OnInAVis COV) (1844), Oiies:- ese oscc 20s ese ee 25 


leucosiaRhipidomellasy eccs.:-2: sear aeeeasesecese teaeeae serene eee 42 
Wévetlle (183 S)ii.co.evcceasszpense aa daca cececvuseaneche tence coe eee reer eeree 92 
Levorson; (CAO: -l.eccccasawscsadstwersecccsenestesceesoeste scree 31,60,140 
Licharew (1956) 119 
PELOLIMAM Herat cans oeceuracciete eDetoeetestet ve cisco aenmaptecitl cease tema Neoee ee meee 8 
ligea, Lingula 32 
Ei ZONOGING SP. co<ssscerimwsides seit avenstansetacees treet een rere noe eon 
[ime @reeksBormationhcc.cc2-ce cee ecees-eee eee eee reset eee 18,19 
linguiformis Poly gnathus secccscsceseeeseeseeeceeeseee ete ae heeee 25,26,28 
Lingula Bruguiere (1797) 

complanata We S= Williams) (1882))) ieee. 22st sesso eee 32 

ligea Hall (1860) 

SDipl) ecstete-ccseceee 

SPso2 J. vomcarawccscstese ected anil sch eceieced tee seceaeenan ce ane eee nere 

SPs 3 wvcccdh hesee cues cusavierssteaws vac veleeeeSeenaeeeree cae st cece eeeeee 

SP34o esi decveiee Scenes ccc casaecanas vaceesesesoueceee doseenene ater eeeeee 
Lingulipora: Gintyi (1898) se -n.eseeceseeenesech see seeet ee eee eee eeeree 

JXOMEIARE! 18, JO seecocasnaccts 


williamsana Girty (1898)... 
Linoproductus Chao (1927) 


Eonchodina: Spe. .cssetescestas tokees« ones ee ee <0 Cael Coe ees ee 
longiposticus, Polygnathus 
Llotis) GyPiduld) soiwececcscwmosiecsieonccececedvascivetse delodurs seca ee eeeReeene 
Louisiana-Limestone:, 2f... 4. s.22 2. aes- sos nae een ese eee 
Ludvigson and Perry (1975) 
Isyashenkol (195i) ecaescscese sees reece 
macfarlant; ‘SGRIZOPROVIG Jeencecscces-e-2-- 5+ ee caceeosseeeracedeeet eee 35 
Macgeea Webster (1889) eens neeces: secs cece eee 15,18,19,34 
MNACIGs SPIVifeT css success teomewe ceases as ones once tes dae semen eee 114 
madisonense> Mrifidorostellurn) esc ..2222<- seas sees eae 79 
MaAQISCer ELLA! a ccwsteceaae eiomeediu sete aaacseeee 36 ...... 15,19,/16,117 
magna, 
S CHIZOPROPIG voc. ewe seuwscowaeseussccseueeectsacaictseectaee eee eeee eee 37 
Warrenellaceytiscccnteordcuaeniveedes coerce 34 00... 10,14, 714,115 
IMALAGICALNIXA OWEN Ge arenses oe sen eee een aoe 10) 52.2: 19,55,56 
magnificum, Coeloterorhynchus ...... 17,18,40 ...... 18-20, 76,77,78 
Maillieux (1939) cvec. focavewnwacsecncvasecoesessavorseessenneeeeeeetateeeeee 54 
Manticoceras: Hyatt (1884) iiss .cseeece- see eee eee eee 15,18 
manticula, 
Aldlacellan s2csccsestcxteeuerenacc versace dete eaeee eee eee 22,39, 120 
Rhipidomella® sos scaptoazctecutsesstencsesegese sae eae ee 39,42,120 
Marco) (1858) cs2ccc< seseces oases vex cneeses cowesssvesscecse coat ee cee eee 120 
Martin (1809) <nsof.fssieeatesaceete ge cncp eine severe ance se siose ee eee eee 
Martin Formations sencsqesenccsnacoq-ceeresceseacace ene teteenee eee eee 


Jerome Member 
mascula, Spinatrypa 
Massalongo (1855) 
McCammon (1960) 


McLaren Di Ui cisseesvnatewesastranneaaeeeeecnc cca ee se tein ce eee eee 
Me Laren\(1961)) <ccececcesscctes.cusstiowcccetecasscaccueesesesmeeee 70,72,74 
Melearent(1962)i) susseestecsecnceeeae se ceqaentenceteetens 14,72,75,80-82 
McWareni (1965) eeseeseeee eee 69,74 
Mearenvand sNoris) (962) aensnesc-scceeereese see apes eee eee nee 72 
mclareni, 

POD Ar APNOLMYNGHUSicscoreceadeaeseene es eaneee near 18%. 24,83 

ETQA OTMYAGHIG arses cecertesesetse en aaees 1 reer 14,72,74,75,76 
IMGCNalri(937)) stevia sacssegesseecceecasleseeestet acne deineemece cee areca 14 
M'Goyi(844)) siccseiiscceccenewonteadesescnsssnoee 25,67,91—93, 117,120 
IIS SORTED). SantoosseoncusbscasoocasococnosescocpacoaceédsGecanoanncesh 113,114 
Meeki(1867) [1868]) <2. ccccsc. sacececsensceaveceeavessseee 14,35,81,95,115 
Meek \(1876)i ccccscivac saawavvcacanavamacsennantenece poe sonem testa 113,114 
Meeki(1 872) inate ce st.cencdecdacunneccecescnsars ccekcd ee seer eee eee ae eee 114 


MeeKi, SCHiIZOPHOria’ «225. acexcdustaessuwcessaeccescatesecucnecessceeteees 37 


DEVONIAN BRACHIOPODS OF NEw MEXICO: COOPER AND DuTRO 209 


Menken (1828) 8 teniesecesescsccswccien see ceaceete coste ct ante Mastuaoaseeasan S2ES4 
Meristella 93 
barrisi ? Kindle (not Hall, 1860a) 93 
ROMIMIQIEIDSs UTA O AIA ars sono opsosbanonnbesoasigosnodann0d00s900600000000 95 
IMermiam (L940) sc cceucneresacuiuica cate teacesaeioee sane nesects 92,105,108, 115 
ROSACEA, NEURO O MORO. ccosccocassosococo-cosccequooondocesannosccooase 37 
mesacostalis, 
(COR OUCOG Te caostes cudeddocrce onbeceontetacuscher oat doudssonncusdonsd 79 
TESIOUAN IDS Sacooacbepeoedot bc een Sere ericcoEsConeaosescrcRecaecaEn ond 98 
mesastrialus, Orthospirifer 101 
INT EIT da ceeee po cd cc cd os rice aR DSaAnCRASEE DaLCRCERRCRSBE Reamer 14,37,44,87 
SOUGH ERM Mill AN a yeemewes soceaentes decoeaesecsckeecoueete esate ease 43 
Mikryukov (1968)........ coon WOLSi7/ 
IN DTT re. (AGES) econersteroccteet epcersnasctdtecracercnosbeaarare.aenterebocoesced 50 
Millerandayvounsquisty(194i7)ereccescsecsceseusnecesnseaceenssatesce scones 28 
Milne-Edwards and Haime (1850)...............0.....0.00c00cceeee ee 15,19 
UIGOn GYDIGUIGeesncaseacse cert teecee eee ceeceenee ne 118} seacae 19,68,69 
VAULT PIOUVINUIG: Seccsceaccacsecawscshuscwcecesncias $9) see 19,50,51 
POLO VOLNGI, JACUTACEO GAS. cacooacoocoooncpnpcopeaceboooDpaaxneecece0Es0n00 28 
TEA) TELAT al IAL HO FB} a cecadeaasocopjacacencondenocasecosooesaccos 28 
Mississippian...................0.0+ 8,10,15,21-25,27,54,84,92,93,120 
Conodontss iE arly: Jc. vevesue cee sens odewesstincaec cust ovbwcase aeeewases 27 
ING STM OOKIAN faer sires eiantceie cies oeutesemmasiatte cnace se newesees 
IMSS OW Reet en tack ocna sec acassi nd a<cetah Aatecee suanedsanaanee cs ecthecsdaie 
IMOM CAN amc cracte rover anes cence ss cnot scseauseenen ekoe eta eens 
monticola, 
(CAOAUTOTULION saa concen GanRensOon cL SeT EEE Cnc ROTC REC EE RCr EE ricooserereoarone 
SPITIer eee. ee 
Montoya Group 
IMO OTeN (952) bate sa-hs sen eciecen cs deisc s doussa des sect eetsh use saesseoneeee oes 
WOR OOH DAG HES 306 (42 soaseeaessqncods0sac0sG008—000000000000 10,14,7/,72 
WATIGOSTALUMIAD RISD -tascece se ceas tence eee scee see ceee 15,41 osne2 14,72 
Mount Hawk Formation...................:ecceeeeeeeeceeeeeeeees 18,67,89 
mucrospina, Nervostrophia 5 U ease 15,18,19,47 
IMMGTOSDIniferl Grabatl(193)l) escesceseeatseerecnerec te reee ee tee eee rerees 97 
Gonsobrinuss(GiOrbiony. L850) esneseee eee sees eee see eeeeseeeereee ee 97 
IDGUVIES MTS Dispeced soeceaeoacincsosesewecde ae saneseeeuek eineswcwe DM] seco 14,97 
Mucrospirifer? 
Gi AML? Se@nTGHS (BEING, TE48))  jcoopssosonesooosusenonaqecses 11,14,97 
sculptilis (Hall, 1843) .... 97 
Muir-Wood (1955) .......... Sy/ 
IMIMITEWOOGNGLOG2)) ees .s scans cae av scien serasesiiee sitions de aetneeeeotss 57,58 
Muir-Wood and Cooper (1960) .................0206068 24,31,61,63-67 
10 (Cal ere (IO GS) ee ee nes Sea a Ree ee Ree ee 28 
multicostata, 
(GAMA OLOE CRIA ete e oe aoe a8: sede is orate Uo ekenne seeeeaeseescetasees 79 
Petasmaria Dy aieseenn 24,84 ,85 
aAlteOSe a, “(COVA ROOCA TE” — cocceonoandecoocosacuoeeenesesdcoace 82 
multinervosa, Nervostrophia ............0.0000065 Viroeer 19,30,47,48 
multinervosa (cf.), Nervostrophia —.........0c0cc0ceecneneee es TT 68606 48 
RFLILTUCA Cem Gy PLLA Gewese are ae oe ota ee Selassie dis dares pace ome eeae dase etme ele 67 
INalavikims (19477) peesscesseae a cscesnssseneseose ae sce stacy aelaeae Sarees otocule 79 
Nalivkin (in Fredericks, 1924) ... oe 21,104,110 
WIAD NOMA VOGIAH AS: poquenanedeehece paSreecrpaEaeoe seqodnoncobpeaseccsaosnoedo 58 
MANUS trlatochonetesy seraceecesscasecesansneeseeee MM, ete 14,57,58 
National Museum of Natural History ......... 7,25,27,31,35,39,46, 
64,65,68,114,143 
navicella, 
Centr onellammer. ce ccetc sass tas ooeeeeisiens Sanco iene Mesa pase ee Ne eoet Reds 119 
Centronella (?) ... eae 119 
GrANACNA recan saws cee aac cee swee Sous een, (OO eee 20,30,/19 
Granaenell aimee ease ses eo acc catenin canes San ec a ons w neste mae ee ate nese 119 
GRAD RENTON, cescaenocone decane tenDEDROOORE eT ocercetncsde ide cerenoere cee nce 119 


NavicdlamDouvillindecscnn ccc tes eee eRe ae eee 

INES) fo 1a Bear = \a cinco a coo Ka ReRAR CDRS cE spe. ccooondscHaaaasorcendeaccees 

Nelson (1940) ............ 

Neoprioniodus sp. ...... 

nervosa, Nervostrophia 

Nenvostropniae@astera (1939) peese- a= seee ee neesteseeee tees 10,11,46—-S0 
Calving Stainbrooki G45) ire seeereree tees eee ee 47 
canace’(Halltandhwihitfield 5118/3) ssssenseateceseseeecn coeeceneeeaee 47 
cf. N. multinervosa Stainbrook (1945)............0....000- Tite 48 
CNLENSG MASP y nsec ce een es hoses eee enero 6:9).2k- 5: 19,30,46,47 
PENIGUIALAEN SSP ce sneteses seseereerneeee eens 6: ssee5: 15,19,46,47 
IMUCTOSPING NASP ae orecesesenesrasseaeaeeaeceeees Hace 15,18,19,47 
multinervosa Stainbrook (1945)............... Titec 19,30,47,48 
nervosa (ELAM IS43) lence vette ators chose senonn aoececeneseeeeene eoeeeee 48 
TALTIICT AV GRC) 9 smsocccadsceatdchca eter prroeenecreeperctoccaas 6. st scte 11,14,48 
rockfordensis (Fenton and Fenton, 1924) .................. 47-49 
SOLIGANINES Dit eo otese ee eeaee cortege seen Up Stoetiona 11,19,47,48,49 
SPy Stain brooks(1945)Meces esters eacen cee eee ee cca eee 46 
thomasia stain brooks (1948) maessesceeecesteaseee ee eee teeters 48,49 
tullven'sish (Hen SeaWilliamsanl890) mecsssenee ee see ee seeeeeeeee esac eee 14,48 

Nevada isis jac sicctean ance caacce coon nneceees 
Dutch John Mountain ... 
GERI socecscudosnseosdoane 
Eureka District ............ i 

INGils: Walley» fiiev esti tteeasceetnessace cs ieteenee teat ees eee eee 

Biocheftesth: Shee ede ee ones aac eee ee 
Roberts Mountains ... 

INGWEGEYILTTNESIOS: cenapscopancascocccosccopcoscourqoecousesoopasonsoEReC 

New Mexico 
Alamo Cany One. sq-cescees sesautssceeacsnsecaeceeneen tes 7,102,153, 154 
AMlamoiPeak inn fo aes saseeaeaaeeee ne ane sea 7,10,18,29,38 
ATcentel any On ihe. .secewenene seeistaneodeeee eee 7/ 
AshtGany oni (north) kessccoscesecescoe see sce cetera ee nee eee 8 
Ash Canyon (South) stscsccsecssssasee sce seca nee aeeeeeee ee 8,21 
Bear Canyon ere sees cece cecese tees eee son Telecare 829,153 
Beary Mountain ie. s-cececcsaoseaeresee eee eee eee eee Ree 23,27,29 
Big gHatchetsMountainsyersnsseeeecesce eter eee teecene see a tere eee eee eee 8 
Black#Range ss) caters scace oseasee ses secncnes oe eee 6,23 
Caballo Mountains ...... 6,8,10,22,26,28,74,76,81,86,96, 109,116 
Capital Peaks siccsttediveccaec che soneshnceccaneaeetaeseeeee nies 8,10,44,113 
GEIS Cero aa eaubisc prcintacios naa sree One e a cls Na ANS EROS RE 6 
Cooks Peak area .... 6,8,26,28 
Deadman Canyon (Sacramento Mountains) ......... 22,27,28,153 
Deadman Canyon (San Andres Mountains) ..... 20,21,26,27,154 
IDY anf) 6 11 | Wiaeaoeecoasacceebopscascesceseusc ios uosodreeas 6,8,10,15,74,76,94 
Dog! Cany Ones ..vskinaeisevescheceenv un hesthaseeeoeee eee eee 7,10 
Escondido Canyon :5¢ sicncsee oes veeeecew ss eae tense eee 7 
Rranklins Mountains \encescsce--ecncoesecreccines cece eee eee eee 6,29 
Georgetown ............ 23,31,156 
Haystack: Gany one’ ccactteaiwatceie cect naan ee haere nee ae 8 
Hembrillo}Canyonleercesasi ieee teen eee eee eee 8,10,21 
| ole) na\o het Werananccoaoeseseanaancees 6,10,11,15,18,22,28,29,59, 103,154 
} SUT Khoy0) Colm maenaaansenccescenicearcnacaaecercs 6,22,23,25,28,30,39, 120,156 
IndiansWells) Canyonieenenseeesce eee eee eee 7,18,41,118,119 
Johnson Park Canyon ................c00c00eeee eee 8,11,15,25,44,69,97 
IRIN SSCON. dW see ode taete eae ces cciwnnacoacenstereaees-eteee ee nae 23,24,27 
Make: Valley? 32502 so Site Secttereccnctecncoeeane yeudees se ocean ee 831 
onesMountaini (Hunley) ias.eaeseeteeeneeenece teen r eee serene 25,156 
Marble*Ganyon sesitecasacassceescnccccce: oecscen seen sev arecenesecerencare 15 
Mimbres*Ranp evevesrs ear. cea teen eee ee ete eerteeeeee cereee 29 
Mockingbird Gap 15,25,44 
Mud Springs Mountains ....... 6,8,10,14,15,18,25,59,76,81,82,116 


NY ut Fe OF1 11.400) | GonencasasnossécacneoRocsecdadcarsosuasucasaetocsaccodccdcgae 10,15 


210 BULLETIN 315 


New Mexico 


Nackaye Mines (=Paint Mine) ............ 8 ,28,86,94 116,118,153 
Nackayes Mountain rsssasstiseccsetrecsect eran lee casiiect steer tare 118 
[Rese] PeN ORES <rapanconoodocesuanssucboocecesbangocqoSUcTADGoooosEeoCDonoDon. 22 
lfte (CEN) _ sponcccsdosseecuonscnoncHcocncmcnoneacauaccadecdaecsnn 7,10,153 
RediHonsesMountainisseerace-ateeesce aces snese cect ce tecenseaseetenes 28 
Rhodes Canyon = 8,19,20,21,25,26,39,81,107 
ROdESHPASS 12, cecsesecens saweeeds dent see caceeeeeeeee emotes 15,92,105,120 
TTI (S{0) 1 Wil Baap eee Reece Reesor to parecrnarnocanacee casa espaedacect 6,8,10,29 
Sacramento Mountains ... 6-8,10,15,18,22,28-31,40,41,44,49,5S0, 
66,73,77,80,82,103,110,118,153,154 
Shan ZAC IRES (CEO) Sosecnonocescconcacéoncdoncacassaaaqa” 7,8,10,15,19,94 
San Andres Mountains ...... 6-8,10,11,15,18-22,25—27,29,31,41, 
73,77 ,80,82,103,105,110,113,117,153-155 
SantavRitapeer.sc-seeeess 83 
Sheep Mountain 8,15,18-20,155 
Silver! Gitysanease-cctacs.sscaosucencesesnceeesen cesar 6,8,23,24,27,31 
SR CO ee eeer enceticontneoc crore acnaeenBEtarnoceasnh cosaceccnaicosbaGonon 79 
south-central .. peeOs 2 L275 0 
South western seen. ..cncs-5: bosons saeeeec eset mae etecececeee sete 8,27-29 
slinith=or=COnSequenCesay crs. saccetccsmesteeastesaeesteescee cane 6,116 
Wihite Sands) Proving) Grounds) ce. -pcsecs<eceesseesaeseenesaeacenenese 7 
WilsSoneRanchiws.-eccsccesees sc oncanacscecseoseneecenocaescens 23,26,28,83 
New Mexico Bureau of Mines and Mineral Resources ...... 7,31 
newmexicoense, Ripidiorhynchus ............-+. 15,41 ...... 20,22,71 
ING WY Onkgiien ce deseccsrenererstdeovetseed 14,25 ,32,42,43,50,63,70,71,87, 
94,97 ,98, 101,105,108, 109 
Hornell Swill ex jeccetccede cache cava ws Oecctoues te neednnscoeeysorootesen sn ae 119 
Ningoingellavnoberts: (197i) eacesese-csseeec ee seetecnce en eeatee teeta 77 
TOMO, To) MAH TIS! — coccenenossenononecnconseanoonjannecdsoe60000 28 
NOACOSGHAnGyrod ellaw y..sossuescienecen sean seas saenesseteensuecerececes 26,28 
NOGOSUS> ACTIORUS soso acs va casa ves sowendcsosswene seeesssauiacue seve sncesss 
normalis, Polygnathus 
IN foudeh C710) Iiperane neon noeree teen en pacepncrnarae erronccroorcrsaiccondc: 
notabilis, 
SPPUPT ON mc secon ache ovcele sab gacive oss sista coals weaise dz toinniaee eeeeseoeanaeeareee 99 
STOPNODICUT OM amas se seme et ice eter eee ee 29) nes 24,99 
novamexicana, Tylothyris. ............++5 P9233. seson0 20-22,98, 99,120 
nupera, 
FA COMING) soo cescnivas dc de ise s dese sicuans tages ioe sesenesstienece eemetentee 24,63 
Heteralosia 63 
obscurum, 
TALS OROTIMTA IIT ccaconccoaseanqecseosecenesece ZAN) cossec 11,14,8/,82 
Trifidorostellum weseee 24578879, 
ODSCULUSHIRELICNONELES) saseeetresenseehonteretece 1 eeeeeen 22-24, 58,59 
ODIO AXEAOS), SAMA YO cooceccooncencoxcoeccencnopaesG000 DA se ee. 20,90 
obsoleta, Siphonodella 27 
occidentalis, 
(COTA hate ce nciicoc cue eE Ec oReCOBECCEO CECT CEE BERR EHS SROE Rai recencee 68 
Wiarrenell dines wastsicnoccncs calcnoe ender see 115 
Mehler (US87) mereses ceive seeks one acts aN soe ee oY 20,49,50 
WehlertiGlS90) Mec ccerctessrescret seat inet ocaues ese eeneeeeaenes 39,41,120 
(OU UN) cea deaer oo Reon Cop SEE ORO TORE TET CART MEE RP EOARER CREA ha gers 82 
QlOSIAMUN RODS Were scsines oesss sat ct erence 24,117 
putilla (Stainbrook, 1947) ............ 9,36,37,42,44 ...... 24,117 
Onate Formation ......... J. Neaoens F,6—-8,/0,/1,14,15,20,22,25-27, 
29,30,44,48,59-61,69,72,74,76, 
81,82,87,94,95,97, 101,119 
PAB OLOLtecr ae ccotnes naterd seen treee cies eee ecto ee 


Brachiopod assemblage of 
Conodonts in. 

Onivie2s. ; 

Warrenella beds 


Onondaga Limestone 


(Opiks((1938) Ieeeeeteeresestes 

OpikiG O34) bow See as cccve.cosnhaccet eas ee ee 

OnDiGularism RIptGlor iyi G HU Sierese ee saeese- eae ane oe eee eee eee 71 

Orbiculoideakdi@rbignyi (847) ere ceeser eee ee eee eee eee 33 
SPs ofadooseyuatawaiceas eosaenes marenataceaetawas eldinae seea ete eee eee 14 
Spi seectecdidvesdh.. qeszgevia fh ecieseeemescsaceeassseeeveess Wi Ass 20,33 


sp. 2 


MOrbigny (L845) czas. ce tanes cecessatdsce ws qenestncestenaes-¢eosseeeeeee 33 
Gi Orbigny (1850) tse esk- osssccesccascuesancseccdecnseeWeaeeeaeee eee 97 
orestes, 
TNAOS DIN ifier set ohe scree se oseogee wc ookae nee tegen tees eee ee 113,114 
SDUNILER: cacccevese.sbaaecanoana ceseus cies ictlvasswessteenenneteee tee ae 113 
ONneStesn (att) snd OS DIT Clauses eee seeeee ne tener 330 es 18,19, 7/3 
Orthis 
inferasCalvini((1878) xc: ...cseseccctsb ee seleesadlcsen ep esac aa eee eres 39 
Towensis Hallli(1858) ees: cxcve seccsesteneceasyenec nce aeeeee eee 37 
praval Hallli(1858)\..sedecees de sen cise coss eens cece sce eRe eee 54 
Orthis (Schizophoria) 
tmpressa Calvin (1897) esecc-a-enee 2 erence net eee ee eee eee 37 
iowensis (Hall, 1858) 37 
Orthospirifersbitrati (1975) eee--ce-eeeceeeeee aces e erate eee cee Rees 100 
mesastrialis) (Hall 1843) eecsc.scaseescccnes-ceeees nee 101 
So sacs coven hoa tee estat eeae sass deuceseoe.petecssees 29) eee 14,99, 100 
Orthotetesiprava (Ela 858) is cescssccecensseceeeeenceeetaaee sees 54 
ourayensis, 
CRANIG 5. cs ssdiss nasi axeascsieescacncessede xe aecweu nen macedes te vere ooo eeeEES 34 
Petrocrania ...... Ll eee 24,34 
Ouray Formation ... 64,92 
Owen (852) i-eeeeercses sees 44,67,68 
OWENENSIS) LOW QAtrY POs <esectee estes soos. sole does os Puech s aaah 89 
OZArKOGINAES Pee cerns econ ae sees tase son eee saeco eee eee 27,28 
Pachyphyllum Milne-Edwards and Haime (1850) ............... 15,19 
[ECT its Rem Re OR RE SERCO Ron bod Ecc ch Scaoaeeceeenb eda cqaemncuncssuodacic 31,142 
Paige (1916): 1.0.5 cise ci tessecstatscsscscacteccescetanss teccer eee seme meeeeee 8 
Palliser Formation? (. i veco.cs.csesencectasonterecmeree tone tec een eames 22 
Palmatolepis 
ait. circularis) Szulezewska) (1971) nese: secese= see see sesseneseease eee 28 
crepida ? Sannemann (1955) ............-. 28 
gigas Miller and Youngquist (1947) 28 
glabra Ulrichand’ Bassler/(1926)) c2..ecese ees ones eee sees eee 28 
actita THelmsi(1963) | nccscesacan-seccceseateeee eee teeter eeeeeeeee 28 
glabra Ulrich and Bassler (1926) 28 
lepta Ziegler and Huddle (1969) 28 
prima Ziegler and Huddle (1969) 28 
pectinata’ Ziegler (962) eecsste ene -c sees cece eee ee 28 
minuta 
loba Helms\(1963)) ss.cesccscecccccadves seccseerecesueen tesco eee 28 
minuta Branson and Mehl (1934) ..............c0.ceeeeeueeen eres 28 
perlobata Ulrich and Bassler (1926)...............:c.eeeeeeeeeee eens 28 
quadrantinodosa Branson and Mehl (1934) .................0665 28 
inflexay Muller (1956) easse-mscescees csc cececce eee sceree eee 76 
quadrantinodosalobata Sannemann (1955) ................0200006+ 28 
sp. wees, 26328) 
subrecta Miller and Youngquist (1947) .............cecceeseeee eee 26 
superlobata Branson and Mehl (1934) .................c0eeceeeeeees 28 
tenuipunctata Sannemann (1955) ...........0.sccceceeeeeeneeenenenes 28 
irianeularisySannemann! (1955) ie eeeseeeee sete ssettaete eer eee ee ets 28 
Palmatolepis crepida Zone ........... 
Palmatolepis gigas Zone .............6+ 
Palmatolepis marginifera Zone 
Palmatolepis rhomboidea ZOMG eseecsese-eeeaee seer ease saat 26,29 


Palmatolepisitriangulanisw Zone serecess-ceeeeeseecee see se eee eee 26 


DEVONIAN BRACHIOPODS OF NEw Mexico: CoopER AND DuTRO 211 


Paraphorhynchus perchaensis Stainbrook (1947) ............... 83,85 
arapuonaxcSchimdts (1964) iec.meccteeceeeereeereereeecsmereeee dieses 77 
Paraspirifer, Wedekind) (1926) <..2-.-cscccsesssesscceceseooes sores 100,101 
acuminatus (Conrad, 1839) 100,101 
bownockeri (Stewart, 1927) 101 
cultrijugatus (Roemer, 1844) ... 101 
“‘Paraspirifer cf. ACUMINAIUS”? .............00000000 Seco KOI! 
parviuscula, Douvillind ........2...c0ee0eeeeeesees Oe. 1S S19 S552 
panvulus, Strophonelloides\.........<-«-----0c--+-0=00 Si nese 19,45,46 
DAGUUS sR MILGHOSDINIf CIs sone ea.nes= scene tecsecceaceeeaesoeeees QT ips 14,97 
patens, Petasmaria ............ 20,21,42,44 ...... 23,24,83,84,85,86 
pauperculus, Leioproductus Sth nL Ses. 50, 2404 
Raurornynchai Cooper (1942) hec..cesncs c-senceeresceseshineceris 23 ,25,80 
cooperi Stainbrook (1947) ............... 20,40 ...... 22-24,82,83 
G2)> .. cadaaaboasnssOOC DS SoU BSCE ARCASrISCRUER ACER Raa BOpOCOS COC OSB ERC rACeCrocrscn 82 
DECUEYIMEOSILOPNGLOSIAM mee ac meee e eee ene ee eee 60,61 
Penns Vallare attend. cs. csscsccressecnescscccocussseerseue eerencesutiret ewe 71 
Pennsylvanian 6,15,25,120 
PEM tGIERELLGNSD nee sec sescotne tose ceeetieurencsseeneneees 39) casdee 11,14,69 
percha, 
SGUGINTARAIIGL tadaacsotencposcebsco CS aEeBaacnoChScseeEOrcetee chocRaoceecos 56 
S@ANGICGHIGQUG Scans basceseouedsccucaaeercee Cee Medcenocnee LO}: 24,56 
perchaensis, 
Paraphorhynchus ....... 
IRQ ROSCA coceoecaconosoec 
FRELLGNOMELES ere eerie vc saseceses ocoaeasacee soe ee 
Percha Formation (Shale) ............ Divecvecs 6,8,15,21,22,23,24,25, 
26,29,31,36,39,54,64,70, 83,84,93, 
105,106, 109,111,117,120,121 
Bella Vie Delupecares ease atecs usc Gavscus vavensstenstechu se watens aroneeeee 6 
BoxeMembeteess-ss-seesees DD) eee 6,8,22,23,24,25,29,30,32,36,39, 
56,63 ,64,67,70, 79,82,92,93,105,112,121,143 
PND CHO lige ae as nt cetera nie soe disses csisunalasiys olseoameniedewadee tne 24,25 
Brachiopodiassemblagerofies.c.cessen: seesteer reese eeeeseee eee 24 
Ready Pay Member............... Dros 6,8,15,21,22,23, 24-26, 
29,33,112,121 
NO CMO lem ne trecthiate oc ailde 5 csis soe vs ovaleaua ap sacacsn se aaeeausiesse veeseus 23 
BrachiopodiassemblagelOfcasaeser scene eaceceeacceseteeeccene 23 
Conodonts from ................ Pe ZOs2i/n28) 
SilVermMem De thar. sosetee anc. s ces tose nels seeaqeasess leveet ee oteccieceaeest 6 
Dereyrata ee olylOPROGONIA sieseaccerereeces er se ses ceureh seas eee seen 28 
ERLODALAME RA IMOLOlepisarneseseac-taccane ser enoeetecenseee esate eneteeee 28 
DENDICXGMDIGNAGGENIG. eeccecceese-ceee- eo pee eens 1202 14,60,6/,62 
DEN DICKUS ROLY SNGINUS: Me nacnsce onion oeeceatseeee cconseesbes sieaeete eee 28 
perversa, Douvillinaria ..... wee, 
Petasmaria n. gen. ..............++ .. 24,78,83,84,85 
GOIMPAGCLAWNA SP werene cone secee se soemnmen es oes ose se casas 7A) scone 24,84 
MU ILLGOSTAlaaMy SD iiss can cesesseeee sees --aeasisecere 21 24,84,85 
JACHOGS 1b Gos cpaagoodasessaesoe 20,21,42,44 ...... 23 ,24,83,84,85,86 


Petrocrania Raymond (1911) 
famelica (Hall and Whitfield)... 
ourayensis (Kindle, 1909) 


CPD “Ih . GosaecanscenacannansoosonOcnge deoarscooeceaHbodaooooRaaTS 

SP-sUMIAEN Geen os. cs -seee ie waice ee eanosdscee se deuisecoce 
PRIME Er GUS Drake seracetswplosiewiels clas seseacie sees stjeselaee woisele oe 
Phillips (1836) 120 
Phillips (1841) ee lOr 
Bilotishalepeeceesceece tence 22 
EineyRointyRonmatione nc: ose .cctecerccnsscsee cfeed~ case sesede caterers 14,60 

BituminouseShalesMembersncescesce ec se sess etee seen e eee eocecees 60 
ita ty (1 9GS) area eeceacst. ccc. naccssk conics sae seceees te Saahe eeeeees seen 102 
Pi trata (OTS) weeesesce oe. cecctias soaccoecncss che Taste bec cae sbeed eee secede tees 99 


plana, Nervostrophia 3 
Planoproductus hillsboroensis (Kindle, 1909) .................+ 24,65 


ElatyclymrenianZoneneneserseee ae eeeccereesmatten caesar 22-24 
Platyrachella cyrtinaformis (Hall and Whitfield, 1872) ......... 112 
platys, 
Phiemellai es toss cenccces didoaiasseae po cnscs Dcatene 24 ,39,42,43, 120 
TQ OEHIO DIS cock asccnenectooutosbocsoonsocc t eeoricc 11,14,43,44,97 
ElatyterorhynchussSartenach ((970))e.ce-ceseseeeesseeceeeeeeeenates 10,82 


OQUSCUUMENGS DP Senate eee ‘ aca) 20) acne 11,14,8/,82 
russelli (McLaren, 1962)............. 

Pleuropugnoides Ferguson (1966))............2.....00sesecssecosceseevee 

POR CCTTIS, LEAHOOROGIACTT). ceccobecpospoceso0ns0000s000000G000000000 

POY BNAthUs BIS. acececesks ae Sei tec soseitanwsedwancheacse seeesaanee cect 
allocotus| Gale Coope4n(939) iesecseccsassssece erect rese see eeeee ee 
brevilaminus Branson and Mehl (1934) 
communis Branson and Mehl (1934b) ...... 
decorosus ? Stauffer (1940) ............... 
glaber, Ulrich and Bassler (1926) ....0.-.......cccc-sseccseseeseeeee 
homoirregularisyZiegler7i (1962) pescesecesceneeceeececacsesenesee eee 26,28 
PLOCRALUS IE. Reo BTANSOM (L954) tee cach acececnceeneeneaceeceaeeees 27 
linguiformisy Elinde} (187.9) teacc-cssceseoteceseeseccescteeeecere 25,26,28 
longiposticus Branson and Mehl (1934b)... 7H 
nodocostatus Branson and Mehl (1934) ...............0ce0eeeeeeee 28 
normalis Miller and Youngquist (1947) 
perplexus@bhomasi(1949) mi. .c.cs.encsecoe osc deere eee ee eee 
scobiniformis E. R. Branson (1934) ............0c0ccccecceseeeseees 
semicostatus Branson and Mehl (1934) ... 


tuberculatusy 7. Bryant) (L921) eesresee ence atee eee ebeeeeeae eee 25,26 

varcus Stautter, (1940) icchesee recone ee ceee eres eet ee eee 25 
Roly gnathusistynlacuseZ One wanneerence ce eect eae cee eee 23,25 
RolysnathusivarcuseLOne wecsccecercssecesevcseenstertereecececeseatee 14,25 

middlevarcuspSUbZONG seer eseeee tere eeeee eee eee eee eeee eee 14,25 
Polylophodonta 

confluens (Ulrich and Bassler, 1926) ...............sec0eceseeeeees 28 

DOR SUGHE (LEONE, WEPAS)) cocoocmosacesascccacconedoncscconaccascaecs 28 

SD iy sbsceshcatdescccedcons senstadoswcsunas aces sumees seseeesecen deseeeeseeeee 28 
Poolevetial:i(1967)), (acc ee ss ewes ossac sce detec sence teeees See 20,23 
ROMA ICR ONO RE. saconscsqecongseonmnonpbocaeaceCenooaon 397 23,33 
Porostictia. Cooper (1955) eccsscn-scestececeee 25,84,85 

perchaensis (Stainbrook, 1947) 24,83,85 
DOME, GHAOQOTVOP —scoseconcecnoouecesvouaoee 105,108 
PortyRefugioshormati onleceeeseeaeeetee tc eaeee eee eee eee eaeee 22 
posterogranulosa, Hispidaria  .............6.06+ BV, eacto 24,110,111 
DOSTIGUS GYILOS Din lf Cloeractccesescrscest crear ce eater eee 107 
praecedens, 

US TUIGS sozka cetyl oaaes sr ae een eo ceas cee onesea Nee Sore Serato ae 66 

S CHLOS 1G i202 ocean ds tees hes seen oho deed boda 24,66 
praecursor, 

KF OLOVIG swat aren sees cane neae eee seb te sconces se cuEe cep ecules eee re oa 67 

S CNLOSIO! vassucceescoreweaccegas cuts sess yeu cue oreneusesemousete settee 24 66,67 
prava, 

PLOW Ch iG cee. 2jn ce scenpacnemes cee eeteseseeaeeeen 10) osece 19,30,54,55,56 

ORS. ess sc sock neste daewee soe Seas ooew a ees RR Oo Eee 54 

Onthotetesy vesessews chic cdasveucieossktecostes Wome mesos eae roe 54 

SGHUCKEFIENG caemaseiseunsdesienhoostent Shatuesstseeetoe nese ae eee oes 54 
Pray Tes @s csuiciay sed cevavetsiseends cenputeveseunsGeh tence does ee eee 138 
Pray, (L9G) o5ca5 SB voice fe See aetv ce Soe ee eet ators sh aise oe ade eee omen 7,8 
pretiosa, Echinocoelia... Pe kaener 14,95 
prima, Syringospira ...... see 23,241 ON) 
DUUTUSH eS CLG ODOLY SNA NU Swmeestenecansccee teen ee ee eee ee eee 27 
Prioniodontardialatal (Bry antes 9211) seeseaeee scons ceneeene cease eeeee eee 28 
JD ROCH OA LERPROW MENTE coo cocspencoscconcedouococsecsuncoatosacesn- 74 
Productellamballi(SG7) meesccesceecencesteccr cee cea eee eee eee 62,63 

coloradensispbandlel (909) mes ssasetee cece eee eaten nee eee 64 


2D, BULLETIN 315 


Productella 

coloradensis var. plicatus Kindle (1909).... 

hillsboroensis Kindle (1909) .............02.06- 

laminatusemnd|el (1909) eeencsnce te ececeeeteeeteet en eetecescrereccecs 

rugatula Stainbrook (1945)...............665 Vee enane 18,19,30,62,63 

Saag tolexaxa (GIN (CUED) ccacoseeccossanancecondsecoAcnseanoannGecbocs 64 

Gob: II|) cocoaseonceeae.oebsoccaeced sococcoscconesonarones MS econo (Los) 

SPS see seceeeres cisiesete/artersnnts 2 

spinigera Kindle (1909) ...... 

thomasi Stainbrook (1945) ... 
JAROGICHID BS: 22 negcecacceosooosscccorecacnscaccadonps9bict Me recee ZAC) 
ProlpficawAcutathecdenanten.eseetere eee 36.44 ...... 19,101,102 
DLODTIG vA GULAINe Gaerne: eee eeseenec ae eeeae aren eset 30,101,102 
[DrOprian (Cis) A cutatheca esse ce eet eee ee eeaceeter ee eee 101,102 
ProtoleptostrophianG@asterm (1939) eeecsssestes esterase ese teaee ee 49 
prouti, 

Gleiothynidinas sas sie csusovesstuacustecees tedceneee seen eee eae eae eae 92 

Grim SQNin GanicaSacwendecsgcesdeseecena tens See eee ene oe we eee eee 92 
lsanilociinyaa (Goysyee (S/S) eocogoasecoopsooddoseonossonadteosassenscobo° 87 

devoniana (Webster, 1921)......... 2223 Ree 15,18,19,30,87,88 
Pseudodouvillina Stainbrook (1945) .............ce cece eee cree eee es 49,50 

euglyphea Stainbrook (1945) ...............:.00eee Skike 19,30,49 
Pseudopolygnathus 

primus Branson and Mehl (1934b)  ................cceeeeene seen e ees 27 

SD cc acaeseaed ewes te aeseaeee aes not cea a earnce eae Manet eaeae ene 27 
Ptychomalotoechia Sartenaer (19614) ................0+-se0eeeeee ee 25,70 

Sobrina’ (Stainbrook;, 1947). -sec..-..2---e se seconde es alow: 24,70 
Pugnax pugnus Kindle (not Martin, 1809) ..................0000e 83,85 
Pugnoides schucherti Stainbrook (1948) .............0.00.c0eeseeeeee 76 
“Pugnoides”” 

alta Williamsi(1890)\scc ccc. <ccncsesscscscocscasteeso-esecesaoresedeseeee 77 

solon Thomas and Stainbrook (1922) ...............:seceeeeeee eee 71 
IDUBNUS TRUP NAN aM estes cetec ecco ne cen sc cer ar cece aa scesee eee ae 83,85 
Pustula praecedens Stainbrook (1947) ..............00sccecnereenee ees 66 
putilla, 

IBrachiy thy ri Site 2a. aecjnctmeaeeee corssese estos ae deat ease tet eases 117 

(OUCKL7 esnarececerornecoererccenpracecceecenacs 9,36,37,42,44 ...... 24,117 
pyramidalis, Cyrtospirifer ...........0.0000000 SI sche 21,22, 106,107 
quadrantinodosa,-Palmatolepisic.cssesesoe-sseeetene ee acae eee 28 
quadrantinodosa inflexa, Palmatolepis  ..........00.0c0ceceee ee ee eens 28 
quadrantinodosalobata, Palmatolepis ...........0.c00c0ec0ecveeeeee ees 28 
quadrata, 

ISUFOPHOGONIGE taewescasuececwewaieea sanchinca selene acne mentees eae sea 50 

Dhiermel lal eh eacnc ath cs vnsseeanae on seee cust antes oeneseese Dears 20,43 
TATA; TOWALY PA “wiscecsecwssceckecessceceessecceaosans D4 acts 15,19,89 
HANG, MUA OQ DIANA 1 cooemoncnnoocoscoseosb0Nds000 33) ccnsee 20,711,112 
Raymond (1909) ) ssee: csenessecunecece cesta ss deers doatvevedgssterteeses 92 
Raymon (Lil) Meeeeseeseeeeeenes 34 
rectangulata, Douvillina ......... 19,51,52 
regularis, Spathognathodus ... 28 
reidfordi, Eleutherokomma 96 
resupinata, Schuchertella ........0cccseseseseseeeees ... 24,56,57 
Retichonetes, Muir-Woodi(962)) iiesamunscssertesasenccstenecceees 58,59 

nana (Wermeuil ysl 845) oyseccasesscceesseacdeaeeaeieene-eearer oe ceceee 58 

ODSCULUS TUS cscceesescecen ator eaenenenence es 11 ...... 22-24,58,59 

DER CHAERSIS Nai SDemite- nae ar aver eereeeceree ere ete i eee 24,58,59 
REeticulariagspinosa anGlev((909) pe-cecsaceseacseeseeceeteeneeeen cette 116 
reticulata, 

(GOAT AIH, tl teeer corm oree entered Pechecer bono O-teprcaccntcEassos 92,120 

CHIMISAFING pe oncesonse cee coreeae doce ence 26,45 ...... 21,22,92,93,120 
FEVENSA, SIFOPNOGOM Gs .csscene: cancels isas once ses steedeateen he eae eeee 45 
reversa gravis, Strophonelloides 46 
TAENANUS MULOPIGOLEDIUS teasees sense a enenceen seiner : 44 


Rhipidomellai@ehlert (1890)! ses-c eee se-reee cee eee eceeeeeeeee 34,39,42,120 
leucosia: (Hallo 860a):ctckccccscssesede ones nstseetioees eee aor eeeeee 42 
Manticulasstainbrooks (1947) eee eee eee eee eee 39,42,120 
SP a sce ea tees sttatd cance sie seen desea aeeeeer, 

SPQ ave ecastatastaaaseataaes 
vanuxemi (Hall, 1857) 
Rhodes Canyon Formation ............ 6,8,19,20,2/,22,23,26,55,70, 
83 ,92,93,105,107,119-121 
AG TOfs See e sie s be tecactiseeuttese acu seeie aets oe eee eee eee 22 
Brachiopodtassemblageyofices-s-ac-<.4-ssse eae teeee eee neem 21 
Rhyssochonetes Johnson (1970) 10,11,14,59 
(entiraye! (Vatelll, IOI) cenonnccnecneconen 14,59,60 
Solox JOHNSON (1970)! eeecccssesoseesee reece ene eee eee 60 
bellarugosus (Stainbrook, 1943) ............ccccceceeseeeseseecesere 59 
VOHhASONT NASP: a2<-.see<otexedshe noses seseaseee steers 1b aces 14,59,60 

Ripidiorhynchus Sartenaer (1966b) ............2.0.00cecesceceeserenees 71 
NEWMEXICOENSE N. SP. ........0c0eeeeeeee 15:41) ee 20 22271 
orbicularisi (ELalls 1860a) e-ce-csesesscrosancesecenaee ee eee e neater 71 

Roberts:(197)) voeecceswucecapeaticvssconevacses ovewstoo dec sae eee eReeee Wil 

rockfordensis, 

JOUR OME ON TE socoonscentocenencpoomonaposannpepccces 12) 25s 30,60, 61 

Nervostrophia) cits sisssactocssseevedaasttcasesatemedseaeeeeee meee 47-49 

SDINGEFY PA snaise ese tsencte deeiaseeltastnescaacceedseesecesiecaesceee ceeteeette 90 

SUPOPhalOSiG: :cwsseicn.s cossevesnaes canes veseseews «asgess see ee ee CEE 60 
rockymontana, 

Anthracospirifer: ..vseccsses sceawsusnsasstenessoiecced osseous 120 

PROMGSGNIG ceiovens yucetSeaen ceccs ote se gh + sseses sees RE 104 

Roemer \(1844)) \sicecicea adds cacanabvcecanoviusdencteosaeos hosee eee eee 101 

FOTUNG GENS TAI DROO KIC ee eee eeeeee eee eters 39) 20,39,40 

ROtundus Gy ilOSpInifCrameese ene ceseeeee eee eee 302 24,107 

TUDrayY-Alry Paria: <M essatsceeecescselen sone ee ee 14,86,87 

Rugaltarostrum Sartenaer (1961a)  .............0.ecececeeeceeeee ees 83,84 

rugatula, Productella 18,19,30,62,63 

russelli, PlatyterorhynGhusmcss.-dec+sc.ceeseee ene scee eee eee eee 82 

Rzhonsnitskayai(l956)! cac.cccscecncss ecco secre see ceeecee neem e ee eRe eeee 72,76 

Rzhonsnitskaya’(1959)\e--ceeecees sass cesese es seace eee eee ee eee eee 83 

SANGErSONI,. LAGKOLHYNGHIGN ser eeeeceee eee ce cee ee en eee eee 14,74,75 

Sannemann (1955)! .ss2ccs<sene.-.6-e2eqeseseoesesteaee saeco eee eee eee 28 

Sartenaer,, Piss ccwcocesaesiwenas-scabercuueiene onewetvene sere ese aee Re eee eee eRe Ee 8 

Sarfenaeni(196ia))iccpccconceceasectens cee oceenene 25,70,76,78,79,83 84 

Sartenaer\(19G1B)), aciiccceete niece cocuseaiseonack aoe eeee enone eee ee meee nee 69,72 

Sartenaer (1965) ... . 21,22,83,120 

Sartenaer (1966a) ... 18,77 

Sarténaer/(1966D)) -.sccsccseaucesnamacewcs event soveeame sete ee ee eeeeeeeeee 71 

Sartenaer (1969) ... 21,22,24,79 

Sartenger(1970)h wccceseacesensessutesus-enomancesceaceereacte ete 10,81 

Savage, Eberlein, and Churkin (1978) ..................ceccecoeceeeees fli 

SQXitilis, CUpUlarOSIrlln-ceestseceeeeese ene 70 

Schellwienella 1. Thomas (1910)... 56 
percha Stainbrook (1947) ......... 56 

Schizophoria W. King (1850)...........c0ceceecneceeenecn eens 10,11,35-38 
amanaensis Stainbrook (1945) ....... 4539) o5005. 19,30,35,36, 37,38 
‘australis Kindlei(1909) ee scssscesseeeeeeeeeteseeereees Dice 24 34,36 
iowensis (Hall, 1858) ................0++0 Sissceene 18,19,30,35,36,37 
lata Stainbrook (1940)............ oe es Herta: 11,14,30,36,37 
laudoni¢Stainbrook (1940) esseeeeceeeceee tee ae seee eee seas 37 
macharlanta(Meek. 1867/1 868])eanrccenccercecineseaneetetss cee 35 
magna Fenton and Fenton (1924)...............:sssssesssencsessens 37 
meeki Fenton and Fenton (1928) .............0ccceeeeeeeeeenenenenee 37 
mesacarina Umbrie\(1959)) Wonc.ccsesse-eeeetes este se eceeaeeeteeeeee 37 
SPiz dee sh ce gsdecasiawanaaiansiaeacldnerasdectstgemadee tec mse cee one ee eee ane 20 
SPi ll sscsceuasoneuce cheswienecenees sce aaseres coneeemeecss Auer tae 14,37,38 
sp. 2 tN 19,38 


DEVONIAN BRACHIOPODS OF NEw MExIco: COOPER AND DUTRO 213 


Schizophoria 


SDL legge setts es sens cerns viv rsa cpa naiasjns cnteanna rs sae eh ac aa tee Settee 38 
wineries (GSiAS7 (WEDD) socecoscespsosdsseocdesqusssvagcedesusnadoobdosd200 36 
striatulanvan. australis: Kindle((1909)) Seescceseesecesse-eceeeeees 36 
Sielnloidevetrin ((IEVAD) ocosocdocessoencbabadoosscsenbencspdenandoseseasurcpsesaaoe 100 
Schmnidty (1964) toece see se oancs cee Phil. 
Schuchert (1896) 43 
Schuchert (1913) 41,60, 118 
Schuchent (929) ive ses. acevesscesseacscsosp sentence 31,33,35,62,67,79,82 
Schuchert and! Coopen ((193)1).-er----cs-ro-rca--seeeeesoess- 38539.075120 
Schuchertand| Cooper (1932) ersrescesseccceseeeseecee ese eeteretee EOS: 
ScnuchertellaxGinty (1904) <2. .25sc<c0scececseseesscceiesncsescseesceses D4550 
Ghemunvcerisismbnandles (1909) snceeseeateeeaeeeeresseasenece estes 56 
HEVISMWIDILOL (B62) Mins ass cnecec ccs cscaceseecreeet aecstecsaaeeueanee teens 54 
percha (Stainbrook, 1947) ... 96> 10) seene2 24556 
joazing: (0s EV IS) ee depenaecer oaepacbecdocoboccesacprecartacrrrocacees 54 
VESUPINAIAMNASD:. ac -0s-ceasensesscasceossesoesees LOM e407, 


““Schuchertella”’ 


Schuchertella [=Floweria] chemungensis (Conrad) .............-. 55 
NignuchertellopsissMatllieuxd(1939)) seensseds.eceeeeee ses eessecaee cee 54 
schucherti, 
GHONGTE SP rece acc en ek on ote seed duc taee e aaee ewan Rose eee 58 
Coeloterorhynchus .... 77 
TEACHES acoceasdeceaee 76 
scitulites, Chonetes ..... ay 8 
IS CLLMILES PR GHOMELE SS. asec acieeilse sels noe seacae seeiecnasi elton cae oneen tes 58 
SGODIMI OLS OLVONGINUST mescccher canes sdccsiacess cossec nator eeeneeenes 27 
SCHIDHENS,, IMMOROQAIATIAP I ceaeacdoqueecsecodneaLboacuccoCdoaosceqeEdoDeeCeS 97 
GUI DETIISA (Cle) IMI CROSPINIFEN 2i-.cesesrerccesete meeeee eee 11,14,97 
semicostatus, Polygnathus ......... SHORGEeB ECORI, ZAMRP AS) 
SOT AIQIOINONH, JEROGINENALGE “SasceAsocosconuscuccodenecodsoccuacscodeocasceds 64 
Sentosia Muir-Wood and Cooper (1960) 
praecedens (Stainbrook, 1947) ..............cecsceeeeeeeeseseeeees 24,66 
praecursor (Stainbrook, 1947) .......2....0..-:-.seese-eess 24 66,67 
SPIMIEPENC (ANGLES O09) lenses -zcncsecenste-aceesen tees seceoce se eee alas, 
ISCLI PENIS GHOMELES) tevin a cweadutee stasnic soe osseaseseaeseanstatenewdeasenees 58 
Shell Rock Formation......... 29 
Shumardellamweller (90) i ccacesececscs-scstes oceceressentecece ces 77,120 
glabraventra Stainbrook (1947) ..............cc0ccee cence eenenes 83,120 
Sidiachenk oy (LOG) me: soe ates se Soecescks aveveesacnececupecseanes aoeneeeeeese 111 
Silica Nal emmeette tes cest ona sf anelctnsone che oe seisnece came sa sees eases 43 
STLOULLGTUSHMB UM! ON Giaaanencais ne sacsesccenacenecsa sent casmeesieates aaeescase 120 
SIVILG La OC IMAL OLOCGNIG:: o2~.2 eas ceedeecdssces sete sas rene ne eteereane ee 79 
Siphonodella 
GOOPErIMEVASS) (1959) \ieeass wc. eteeteercoe ss oss cesteeaneee dees sceseinester es 27 
Grenularany (Cale COOPEN, 1939) 2 ccsstetenesecesee ne dase eeeere 27 
Ob Soletantlassy (L959) bec sesces cece secaseae esses cnceaseeecesoetsaeeaee 27 
SD eM ae ne eee saceeccis seca out Saeanesteacednnew st aocusadteataecseesetedud 27 
Skenidium independense Stainbrook (1945) ...........2...02.020 es 3b) 
Sly Gap Formation .................. BiG veces F,6-8,10,/5,/8, 19,20, 
21-23 ,26,27,29-31,35,37-39,41,44-46,54-56, 
60,6668 ,73,80,87,88,90,91,94, 
99,102,103,110,114,117,118,121 
IAB EXON Rone eeenc sc ava cee eo cones ccbiansvscueseseacus seanetereuem ooteusesens 
AIL ry DON LONG eats scene 32Sce send scue sauce seeowewacaeansensaeresseenee stenee 


Brachiopod assemblage of ... 
Colonial coral Zone.... 
Conodonts in............. 
Ely pothyridinapnoecdsy,.cucsssrsaeascecee ese stecer ee ote eeeeencen sees 
MowemBrachiopod Zone seesccrees- cesses een aatns ee eeenesesase 
NAGYS AN AEN TAOS (Ch occas ep sccenOcce RCE nod oee ee Bac aee Oe prng Paseo csc ercee 
MR OMASANIANLIONG wacesee se nase ooes aos cans sence = eee Soe eee eee eese 
WniteD) WofeS tevensoMbecsesecre eae renee scceate sees seeaaceecey aires 


Sly Gap Formation 
WnitsyD=HofiStevensoutescesrcccosntteccet eset oncecee esse 15 
Unit E, of Stevenson 
Unit G, of Stevenson 


WinitsHstofiStevensSOn!s....:.c-<cstcs:oscceceseseosceseresaeseus ascends 15,18 

Wppen Brachiopodi Zone ersescessseceeeoseee sect ce sere eee eee ease 15 
Snyder Greekshormationipa-ssscs-csss--cees coseeeneccenerce -caaeeesce 44 
sobrina, 

CAMANOL OE CHIA masesszeews = seacensceciose se ates otcccee secu neaet ee cetntee 70 

JE MMARO IMM OOAG ATG scoseacacencogosepocoaceqs0q000s00¢00000 Al See 70) 
SokolskayaiGU94ll)| 2.canceeswcteowssoawseccesesuucueossmeecsdorsentaades 117 
Sokolskayai(1960))" c.sccssscsasaceteeeeseonses sovevesoscsessescr once meee 44 
Solida, Nervostrophid ..........c0.0.ceceenee V8 vesses 11,19,47,48,49 


solon, ‘‘Pugnoides” 
SOTA LS UL. Greate acessire ct ee seo clat wele ls petaiscisctes side eicie cia stew oisitecio eee tema 
Spathognathodus 


aculeatus (Branson and Mehl, 1934a) .............2..2.00200ec0eee 27 
regularis (Branson and Mehl, 1934) .................c:ceeceeeeeee eee 28 
SDu cde cactgoueetees ree adadavas utssae dived eaeviseeseucanetnuontne nee ace eases 27 
sphaericum, Leiorhynchus .. Baa! be eee e 14,8] 


SPHACrOSPON GIA’ sree shoo oe doje cnc sone ane ale sacae eee eae 10 
Chensetesselatan (ebillipsss184il)essseseee sees eeseeee eee eens 10,14 
Spinatrypa Stainbrooks (195i) sesesscesea-e-eeeee eee see 20,88,90,91 
COMPAGCIA MSP) wesceseeateeee ee eee eeeee eee 2420589590 
Mas culaa(Stain brooks. 93.8) peecseerecesee sateen ate ce eee 90 
rockfordensis (Fenton and Fenton, 1924) .................0.0005 90 
Obsoles CenstN Sper careascenee aceon eee ee 24 ...... 20,90 
trulla (Stainbrook, 1945)... ieee Sees 05902911 
ORCORNGHG Wo SIN. cascasacosacasncoessocs2c60 DS scs2e3 18,19,90,91 
spinigera, 
IR OLOVIG scenes fact oes coarse naes ea dsae te eee eee sedans sacescsecse ee eeeeeaees 
Produ ctella’ wrscicacaniccnases ches oxcaeeecee nse een 
SYA LLOOL leoennnsnontneceoaoconeisorcannoosooanonneeacOnco> 
Spinocyrtia Fredericks (1916) 
SPINOGYTiQNESP A Seeesese were socom eased sea se eeeieereccaeceeeee 
spinosa, 
ELIS DIG AMI Grice ec owen sete see teat nee eee 
RELI CUIAFIG Foocc soctsces sae se nc dne8 S50 Hae eee TSIEN CRORE 
Toryniferella ......... 
spinosus, Torynifer 
Spirifer 
bismestalissGintyi (nor tlalls858) saecesee sees neretere eee eee 99 
disjunctus var. animasensis Girty (1900) .........2....eeseeee eee 104 
euginus: Hentoni(193i1) secssecccsasvsccuaccassc-ceenee eee ee neae ae 113 
repositus Penton (l93ill)) i eceesescese sess esses ce ecsesaereeseesesees 113 
macra Meek (not Hall, 1857)... 114 
monticola Haynes (1916) 111 
notabilis) Kindle (@1909)) eee seseee- see -e ons adage 99: 
onestes) Halland \Wihitfiel di(11872)masseseeaeeeeee se eater ee ee eee 113 
WH AFOOG (UNDIES TIES) Sonacecsoceocscnecaocencebononongsooosccde 113,114 
whitneyt Hrallii(1858) ocsceecewe secs cecentecceessen can ee seesnnereseceree 
whitneyi var. animasensis (Girty, 1900) .............00..2eee scenes 
SQA Ar > SUTSHOSIS WHOS Space canenccobcoesesacssogeneee 
Spirifer (Trigonotreta) strigosus (Meek, 1860) ...... 
2Spirifer whitneyi var. animasensis (Girty, 1900) 
Spirifera 
cyrtinaformis Hall and Whitfield (1872) ....................2c000 112 
Strigosa Meeks (1860) ga..tscsace sacs cess cece oo ence ee ee acne ee seenee 114 
Springers es: a... sosecpecaeceer sade asus steam aese encase So. ceneentenecene 31,142 
Stainbrook, M. A. .... 31,68,69,121,138,140 
Brachiopod!icollectionie-cas-s.ssccseacerosere re eeec eee cae aca setacetenes 31 
Stainbrook«(l935)\sescctcssteewe5.- 20s cncbeesseteces cet scteseee eters eens 53 
Stainbrooki (1938) it sov.c seen scacceecsccece saceesecass conc erenaseemee ees 87,90 
Stainbrook:((1940)): <<<. sciss.sssscesceescensecscscecesscccsece srs 11,14,36,37 


Stain brooke\((1943) i eicencesceseseceion acca cc sesseee eennase saree 59,60,65 


214 BULLETIN 315 


Stainbrook (1945) ............ 8,15,18,20,30,35—-41 ,45-51,53,54,60,62, 
63 ,66,77,80,87—-91,101,102,104,114,117,118 

Stainbrook (1947) ........... 21,22,24,31,34,36,39,42,56,63-67,70,82, 
83,85,92,93 98,99, 104,105, 

107,111,112,116-118,120 


Stainbrook (1948) ............. 8,18,38,45,46,48-50,53 ,54,62,76,80,87, 
88, 101-104, 113,114,116,117 
Stainbrooks (1950) ) cccvscecscceccvscevecscacscscasusectadedsseeectenc see. ences 92 
Stambrooky (1951) i onciewe conc csecssacsanedesedessccusaeacsseas wee ssesetees 88,89 
Sfainbrookin Gypidulaiecnaece-eaeesnescee seeace asec 145 525 30,68 ,69 
Stainbrookia| MAGens ~es...5:25scosscauueeoveessenwcecccseotne ne 20,39,40 
infer. @.( Galvin yl 87.8) eesececcecceseatessccesestee 239" oi.ks 30,39,40 
REUDIR ING Ss soscogcaasenssccde aivd) See 20,39,40 
State Quarry shonmation sasceseceoeeae es seeeee cee eeeee seo er erce eee eee 77 
Stauitier((1938)) ssc cse se Soeck ave snc eeee necean es Se aus oee dase aoe sex eo 28 
Stauferi(S40)b ace ot cca Sento eee ee ee aces nie ae ee 25,26 
Stehlit 9G) Wass. decwssaciees eeese ts No ee ae ee oe 119 
StevenSOns Eee Viescseenceaseeeees 7,31,126—128,130—-133,135,137-141 
Stevenson (O41 D) ir veces sseencece si sedececnos cenesee cee one eaneoaeconee noes 8 
Stevenson (1943)! mesa case ce ceae faces cane seeese ee encweceeoeee ones 100,101 
SLEVEDSONI (L945) eee neceeetee ere see ect eseeseeace 8,10,15,18-20,22,23,68 
stevensoni, Cupularostrum ? ............0000+ 14,41 ...... 20,22,69,70 
S fe wart (GODT) aeseceteceneecesasssuee ie ese eee Mee ene ee anes meena 101 
Streptorhynchus chemungensis (Conrad, 1842) .............2......- Bp) 
Striatochonetes Mikryukov (1968) 10,57,58 
MLQNLUSETICISD cpatsoncecrectcoecs sees eta asec 14,57,58 
subcarinatus n. sp. .....- Soll ees 14,58 
SERTALUI@ AS CHIZODNONIG) a esesee een eee eee ec Cree eee 36 
Striatula vax. australis, SChizophoria 2:2 -2---2c--<--2escsseeceseose=e 36 
SET EP OSQMUS DINU CNG xc ces awae seu enue cece ssece eee reach onc Rete eee 114 
Strigosus, 
UNA OSPUIfer? pareca ces ad eva yece 02 cae oeuwetss seat devs SSiecee 113,114 
AY 110 4 aa paccerr ance aecenronsor eon scbeenoren ierrcernccstecourneeerecere 113,114 
SY 2 0 4 pat echo perenne odor CSE ace CEE CCR PEGE Ea oa he CANCE PES SER ERE 114 
Spider (ri gonotr eta) scewaceecsestcas eaeee ae eee ee 


Srarapel or AO VS ENGR) ececececooccececocacnossdencocodcocnacscdooases 
arcuata Hall (1858) 
boonensis (Swallow, 1860) 


CalvinisMiller\(1883)-cc-ccse-coo odes oes oe a eee 

COSTATANO Wen. L852) he ceaee re ee aeeee reece cece ene eee 

erratican (Winchell 866) e.-caeet ee eece ee eer eneeee 

exilis Calvin (1883) ............ 

quadrata Calvin (1878) 

reversa Calvin (1878) ....... 

Spire fesse ste scc shdevadews cobs dacs tensdecs scence ceeceeee tes 

SPie2 es eee ee ee a ete ee 

thomasi Fenton and Fenton (1924) 

Winbonatamesps csssssee sees eee oer 

IAAT OTEKY (CRANIIN (CIE AD) _ osnoocenanasosoosnecannoscnooncsccancebacnoabed 
Stropholosia rockfordensis Hall and Clarke (1893) ............... 60 
Strophomena chemungensis Conrad (1842) ........2.2.c0e00ec0000e 55 
SLLODNOVElordess Caster (1939) iecese--seseceeseseeeee eee eneteees 45,46 

deeringi Stainbrook (1945) ...............ccececeeeeee Cigeaeeee 30,45, 46 

OFS OCONVEXIS IES Pumeesas cece censtenerosaeeec eter ere By cases 19,30,45 

hybridus (Hall and Whitfield, 1873) .... 30,45 

DAY VUES VSD es 82 codecc. code eee a 19,45 ,46 

reversa gravis (Fenton and Fenton, 1924) ...............-.c00e00+ 46 
Strophopleura notabilis (Kindle, 1909) i eee 24,99 
SUDCOIINGIA, GY DIAUO\....)---r eeeett en ee ta ee ee Lee 18,19,69 
subcarinatus, StriatOChOnetes .......cc0cccececeeeecenecene Miscsves 14,58 
SUDFECION EL AIMAIOLEDIS:, secc-s sees neste 26 
SUDA (COMPOSIIA! Po. c<teesscaseceseeene ee 93 
subumbona, Bmanuelld c.cccsstesent at eae 95 


Suleatostrophiak Caster (1939) ee-seneeceesseecereeee ee ee epee seers 50 

Calvinil(Miller 883) aecesesee area eee eee eee 8. eae 19,30,50 
sulcocostata, Tylothyris ............... 27,28,39,44 ...... 18,19,98,99 
Superlobata,sPalmatolepis) acces cnass aero ener eases eee 28 
Swallow (1860) iis. cciccticescevesccscecssssacueveveessedesssteresetdemeeree 44,92 


Taghanician Stage 


Meicherti (1965) esses eee 
tenuipunctata» Palmatolepisy ..-2-2se-<0--ease- ee se eeete eee ene 28 
Hentaculites: Schlothetm(1820) tse .cse--ee see eee teee eee eee 100 
Tenticospirifer Wien (1938) ce cccsensces sesso eee ee ee eee eee ee 112 
cf. T. cyrtinaformis (HalJ and Whitfield, 1872) ..... 3354 19,712 
cyrtinaformis (Hall and Whitfield, 1872) ............-.......0200e 30 
Terebratula navicella Hall (1867) 
Termier and Termier (1949) ......... 
tesselata'(ch.) )Sphacrospongia--secc-o-0-2-eatee eee eee ee 
Texas Mech University: <ccccscc.se-cocsseseccsee eee eee 
thalattodoxas Gyrtospirifensnesscccace--eoeerce eae ee eee ee 107,109 
thalattodoxa (cf.), Cyrtospirifer  ............... 30.2 19,107,108 
Thiemella Hes S; Williams (1908) 22-c--2s2-seeeeeceee ee ee 43 
platys n. sp. ...... eatap Dios 24 ,39,42,43,120 
GUAAN ALA I.) SP :dsace= Seca ove Jasdveccsnseawseeseeseeenee 2a 20,43 
villanovia H. S. Williams (1908) ...........2..2-00.-2-cessseeees 42,43 
Thomas,, T.)\(1910) |. c0cc8 tecdewewcusoscewcuens sduncste seoe oe ete ee 56 
Thomas; T1914) ctecssrsces seas saseesseecascoec hero eeeee te ae 25,64,120 
Phomas; L.A (1949) eo. caeeelecdsaccecse.chetsunseeeee aoe eect 28 
Thomasyand!Stainbrook (1922) ce sccscee eee ee eee 71 
Thomasaria Stainbrook (1945) ... 102,104 
altumbona Stainbrook (1945)............ 30,1702, 103,104 
GeMISSa NSISP ieee acest eteees oes 18,19, /03, 104 
rockymontana (Warren; 1928), i acccce-eseeseee eee ee eee eee eee 104 
TAIT ING 95 ecesossesone 35,36,40,43 ...... 15,18,19,102,/03,104 
Thomasariidae n.ifam! 24s). .csccevtevceesessan eee otee eee 102 
thomasi, 
Nervostrophia ... 
Productella ...... 
Strophodonta 
Thortie; Ei-Awy: sescvecuaverneavossaadedceosewes ace occ toe cena eee 141 
Thoroughgood Formation ...... Cie 6,8,19,20,21,22,23,26,29, 
79,93 94,119,121 
ABO OF wicceieets ovesetecetertesecs eoeeceapoentvaneacces stecesee eee eters 
Brachiopod assemblage of .... 
Conodonts from ................ 
Unit K,-of Stevenson 2.2 .c5.50c.ccce 465. doccawseean acseereeeeeeeeeee 
Wnit L, of Stevenson) (isi. 5.c1.d5s0ee ves cceaeswcedan aus soeeee eee 
Three Forks Formation’ $2cs-ns0e-esseessseeeeeeoeeees 
Prident'Member 22... :..25. 20. <ecseseesene cesiewavecnecceeens saereeeeeee 
diven} (938) ees-ecseseeesee 


timetea, Warrenella ... 
tioga, Cariniferella ... - 
Torquay Limestone’... <.<2cc2scevccsseescssousesces eroevoeesecsteeeerontemes 
TOrYNEfer: cei Riv cos aecawe done eeeaetataues sostedetcnecactece hace a nee 

spinosus) (Kindle, 1909)) so oeo ct ei cdecene=csecececccesceceeee ee seneeees 
Toryniferella spinosa (Kindle, 1909) 
transversa, 

Althiynis! deeccsts cs eee se 

Cleiothyridina ? 
transversalis, Blowerial wasseecteseesseteetereaieee 


DEVONIAN BRACHIOPODS OF NEW MEXxICcOo: COOPER AND DUTRO 215 


MTAVETSELGTOUP) fF ecwdohececansvccesese on cees veeaeee sates evosceceuesese ses 44,97 
triangularis, 
VAIN CNN OBNQLNUS 2.2 va scnecxensssessenssconeeesneens sesonecceocss nash aeons 26 
THAT TOKO IDES Goqecassoceesodennenaecabseecone 
Trifidorostellum Sartenaer (1961a) ..........ceseneeneeeeeneeneeenes Dil 
CIVAETATSE (nN, WOO) sgoscoeccooscosanuddoconsenocondsoaaseden 
IMMCISONEMSEA LAYNES 1916) eacenecscen assess nace ssce eastern serses 
MISGNATGD Ts Ss ceapsoapednoshsodseeadedecuecooss 
MralicurmmeNalvkinn (947) wessseesestaceeceare: sareeeseesseeaeeaserer 
Trigonoglossa Dunbar and Condra (1932) .... 
MrieonOglossaiy SPs cacesencccscceacerascecoseececedss 


MT UMULGLE Gl COMP OSL Carer enccntanca cesta he cereteme asec ovecteetoecseaete see 
Mrapidolepiiusperalli(85i/) mecenctenaeacsceeeteoretcecncneecertsteaeseses 
CMTS (COTE, NEIE)) ~ césoccocesoocenecpaacbocapaadeneesuecds000 43,44 
IDLGLY SINUS Pvq ace vac «con oscscesocsdsdescceesesese (I aten 11,14,43,44,97 
FHENGMUSH (LECH ISOM) ccs eoceaccoatecesesceeeere ve sacceoesocensnes 44 
EMU LOPES DIN ALY PGi eescncces osc asec ee . 30,90,91 
trulla decorticata, Spinatrypa 18,19,90,9] 
EOIN 25 JZOBAAGTHAIIS. cacooascesooococescomeopnoopvesocuuonadece Z2SPAS) 
(UP OQSIS, INGANOMITOUDTE GeBespoccseccteonb00s ascenoodcaooBOceHoeuaECD 14,48 
Mull vabimestOne we sees sec ect cc soes sede ce eavewscenatere sept eeenes 14,43 48,59 
Tyla omnis: IN@TTATIEPAD)  pooeaocnpsaeaqnossancnqpooepenanecosoocsodssn 96-99 
(AOUMOGGT Ws GOs soccsepocsonaceods <6 CUE) soccca LULU LE Ee) 
mesacostalis (Hall, 1843) 98 
novamexicana Stainbrook (1947) ..... 27,28 ...... 20—22,98, 99,120 
SPsdl vec sce sercaes ccsesccseessessdosevesdeccoancezexese D9) Fences 20522599 
SUIGOCOStAta NSP. -cescceeececen sess 27,28,39,44 ...... 18,19,98,99 
Wehtospirifer Wyaschenko) (1957) se---e-reeseee- eee se seeee cesses ee 111,112 
Wehtospinifereigeccnes ete tera ies seen ss cetee te everiivee cose eecemetaseneeen 112 
rarus N. Sp. 5) sconce AOL 
WTI HY 882) eee was nos ses ctenwss covccenccamcoeseuseetnecees seveseusee tenes 10 
WinichyandeBassleri (1926) ee--paccseseceer emcee ere setae cess cect cce= 26,28 
umbonata, 
Ambocoelia 
Dichacaenia 
Strophodonta 
OTNV ENSIG Ye OMMLO Waly secs cores ewnewerte ves cv astievescee dese memevenert aces 31,91 
VAG R, TEAC OSUOITTT. cosccdcosenceaqeecbsosacaecG55000Gecdo000G000006 79 
WeSsGeolopicaliSunVey ce-c----<cocccee set eeceeseesecesucse 7,25,27,28,31 
Silurian-Devonian (=Kirk) catalogue ............... 25,28,141,142 
Wippembaleozoic catalogue ser-n-e-s-sedesesee eee se seeceetes sees ee 27 
WESESeRe (RUSSIA) fac scce-ecee ence eee .. 54,111,117 
Cem tial ln mmmeera areca secs anae scone ole ancetisanccesdsiemscaseorece er 58 
(UIE shoootnaent docpbate cee nrc oSe cree peace conser tone Ceericcerere ee mear erent 29 
W ESLEMM weremuat acts scent secu once son ranevcu ce ance cedeelstesislesles's'ecs tele 29 
VAMUXEMIUM RIT DIC OLE A assceexcetee ses aasee sete beeen eae e ee eacnee esos 42 
EIEIO, (PONE YETITIS) eagabaacadaobeecancecoosnccocoAnene spe scqNcoD0BEaIe00 25 
variabilis, 
Gyr linear cerrecse cons sc <te ccs ce catitt cuaeemenee oss 37-38) 3.20. 14,//8 
ID orauillinaytaterecs sacee sane saec sees ween eee sees Quests 19,30,53 
Douvillinell Gimme sescck coos ccs esse ae ten oe ene ee eee eees 53 
SHAQRINORIORTIGT “ccoraaseddoosdoacnocnasoeOnaEencoech 53 
variacosta, Atrypa 88 
varicostata, 
ALVA DD Giccte da ddcec race cccastee sisecercacsccrsscsdecveceerscisesiwse'scsene siete 88 
Costatrypa PS erie 15,19,30,88,89 


varicostatum, MorphorhynchuS — ..........0.-+-0+ 1554 eee 14,72 


varispinosus, 


IBISDINODROGU CLUSu secre seth ane coteee aecr ee eteee doce cetiese ee seer ene eeee 65 
PFCLOPKOGUCLUS! <5.20 ees. a vole eccieec eae ne hod ea eee eae 24,6: 
venustula, Hypothyridina 


Wemeuilli((84'5) masacssaseencades 
villanovia, Thiemella 
virginia, Cariniferella 
VU SATIS; sDeVONOPLOAUCIUS) wesesesenercsccce tessdecseceseeaeee 


Waagen (1883) 
Waagen (1884) 
Waagen (1885) 
Walcott (1884) 
walcotti, Devonoproductus 


Warreni(1928) i soosc.cocscsescstenas ssoseseeeenese seo sona eeu eateoreeeee 
Warren (1944) ) sacra ihe siee tease caceanstee tustnc cate aaeeee nae 
Warrensandistelcks (1956) imeese-cessecesescceesncesteeeseeeneeceeeerces 15,89 
Warrenella Crickmay (1953) 10,115,116 
eclectea Crickmay (1953) 116 
PlOWEFL MG (SPhizesecescceene see eeeese secs 35,42,43 ...... 11,14,/15,116 
ifranklini (Meeks 1868) ie saccececeacscwacte sce untentiossacae sae sneiecesere 115 
labrequerm CrickmayalGl960) aeeeesereece tesa eeee eae eee ee eee eee 116 
MaAGNa NaSPerereteteee ter eee estes e 10,14,//4,/15 
occidentalis (Merriam, 1940) ..... poo NS) 
timetea’ Crickmay) (1960) mec-seseesecsne ses ceeeceerereceeceeeseaenen eaeee 115 
warreni, Thomasaria ......... 35,36,40,43 ...... 15,18,19,102,/03,104 
Watenway sik onmation=eresee sce esseeeeeeee ce tece ener eee eeee 29,82,96 
Webster(1889b) i) seccc it eos.nc-. cscs seascoredeseccnceeceeeesere ee 15,19,34 
‘Webster (921) Misia sees cate omy sore aes accent meee 15,87,89 
Wedekind (1926) .... 100,101 
Weller (1910))..-....... ZO) 
Weller (1914) tirue.eisiessveacisoveisae scene sesnewas seeeetecnen teen anaeoee ees eess 66 
A,1(2) KK @ 1-1) eeeese coca cen eenceccececresccscrcondencodricocnnecreaceacscaconccc 14 
West Range vboimestones -ccnceeessesecse see cenenteeseeee taser eeere eee 24,107 
Weyer (1967) ici Sevace caer sess opcissemebave sec cecsee mee ete taccuetietnemenes 116 
Whites(1862)) 22215 esc corse seascvesstecptecessctrene meena ee ee eaee eee 54 
whitneyi, 
Cyr tOspirifer® cisccsascsdeeescnsssazpeaseectsacbeecsdact 30) 5.220 19,20, 108 
SPIER was awes ss anee sa cecene sale ve cute ceasebonore seem esc ebse meee meee 111 
whitneyi var. animasensis, Spirifer ...........:c0.ceeceeeeeeeeee 104,107 
Williams: A‘((1953) eee.sececae sees ence aeeueceseoeneeeeeenere reseeeesees 50,56 
WilliamsssAv" etali (L965) eeesssscesceeseccce see nenteaetence sees 31,49,50 
Williams; H./S/ (1882) iccb us: cateaecesceessectsee teen estecaee seaeeeceeenecr 32 
Williams, H. S. (1890)... 14,48,77 
Williams; H./S (1908) i... so.cseccesace.cseavces <cueseusescoedeceroes 41-43 
willlamsana EINnGUlipOnd \penedesescsanese=-seeeeeeee en eseceeeaee deena 33 
Winchelll((1866) iii vccz cecees sae .vcences- cus -sonseeereee eee roe eeesmccesetes 44,45 
Winneperosanbleimestone yestee sec esas nee sae eee eee eae eee 14 
Woodward (ESS2) saci roc coerce cecccies sense ee ee eee 34 
Ane MOA, IDYAO CEO TE! sotocosscncondnccnontonocasboscoocbaccassccouce 61 
Yiochelsons, Es ccsesancecuavennntescoueseccernaeneecescen eee 31,123,129 
Leller (L965) |\.ce3-ccceeens ds hc dennserosesseereereseeeece tee ceeceeete ee tenes 8 
Ziegler (U962)i wsseseccatssescsusesurecssxestace eee eee eatec ene teen 28 
Hie glewandyrHuddleys (1969) taesnccccee ee -eesee eee wiz 28 
Ziegler etal. (US76)) scceelocseceedeseee secede ease canoe sceeeeer oa nee 14 
Zoophy cos Massalongo\ (855) en-cssecsoeseseensteeneeeee eee eece eee 23°25], 


6197 O04 


Text-figure 5.—Correlated stratigraphic sections of Devonian ik 


rocks, San Andres Mountains, New Mexico. Datum is the top of the 6 
EXPLANATIONS OF Ojate Formation; colonial coral beds in the Sly Gap Formation RHODES CANYON 
HEMBRILLO 
ABBREVIATIONS FOR extend as far south as Rhodes Canyon; T, Thoroughgood (type Contadero) CANYON 


Formation, present only from Sheep Mountain to Rhodes Canyon; 

TEXT-FIGURES 4, 5, AND 6 Rhodes Canyon Formation present only from Rhodes Canyon to 
FAUNAL pea ae Ash Canyon (north). (see Text-figure 3 for location of sections). 

algae 

colonial corals 

horn corals 

Macgeea 

Pachyphyllum 

bryozoans 

Sphaerospongia 

inarticulate brachiopods 

chonetids 

spiriferids 20 Ft. 

Aulacella 

Acutoria 

Alryparia 3 

large Camarotoechia / 

small Camarotoechia SHEEP MTN. = 

ae type sivcsp) —_//7/ - 
mani Ld {from Flower} 


Desquamatia Lake Valley ¢/ 


Elita ay, 
Hyborhynchella Sparaclte 


Hadrorhynchia 
Hypothyridina 
Nervostrophia T 
Paurorhyncha (_— 
Platyrerorhynchus 


Pseudoatrypa 2 
Rhyssochonetes 


(modified from 
Kottlowski, et al, 1956) 
Andrecito Mbr. 


Miss. 


BX : 
ae ASH CANYON 


(north) 


pc 


pe 


Caballero 
(Mi 


8 
\ 7 ASH CANYON 9 
(south) 


SAN ANDRES (from Kottlowski, BEAR CANYON 
CANYON et al, 1956) 


(type Onate) 


523n 524c Caballero 


527v 528g 


521q 524m 
j Ma 521d 


1 
Rhipidomella CAPITAL PK. 


Syringospira JOHNSON PARK 
Schizophoria Lake 


Striatochonetes | Valley uh 
Thomasaria (Andrecito Ae 


Caballero 
(Miss.) 


Tropidoleptus Lake Valley Mbr. 
Warrenella floweri 
Warrenella magna 
Sly Gap fauna 
Three Forks fauna 
Manticoceras 
vertical worm burrows 
horizontal worm burrows 
fucoids 

cono conodonts 

ft fish teeth 

OTHER ABBREVIATIONS 

MS mafic sill 

cov covered 

pe partly covered 

ph phosphatic nodules 

T Thoroughgood Formation 

ROCK MODIFIERS 

black 
brown 
cobbly 
gray 
gray-black 
dark gray 
gypsiferous 
green 
green with nodules 
olive 
red-weathering 
red and green 


524u 
bi 


ATr ruseze SPEEA 


528t 35212-——- 
Montoya 5262 Montoya 
(Alana (Aleman 
Mbr.) Mbr.) — 
Montoya 


(Cutter Mbr.) 


On 
nate Fm. Fusselman 


Fusselman 
Montoya 


EXPLANATIONS OF LITHIC PATTERNS FOR TEXT-FIGURES 4, 5, AND 6 \ 


= =] Conglomerate; ie 
phosphatic pebbles Sandstone, dolomitic Limestone 


Sandstone, mostly 


Siltstone Limestone nodules 
fine-grained S22 


Siltstone, dolomitic eas Dolomite 


Sandstone, calcareous 


Fusselman 


PREPARATION OF MANUSCRIPTS 


Bulletins of American Paleontology usually comprises two or more sep- 
arate monographs in two volumes each year. This series is a publication outlet 
for significant longer paleontological monographs for which high quality photo- 
graphic illustrations and the large quarto format are a requisite. 


Manuscripts submitted for publication in this monograph series must be 
typewritten, and double-spaced throughout (including direct quotations and ref- 
erences). All manuscripts should contain a table of contents, lists of text-figures 
and (or) tables, and a short, informative abstract that includes names of all new 
taxa. Format should follow that of recent numbers in the series. All measurements 
must be stated in the metric system, alone or in addition to the English system 
equivalent. The maximum dimensions for photographic plates are 178 mm X 
229 mm (7” X 9”; outlined on this page). Single-page text-figures should be drafted 
for reproduction as single column (82 mm; 34”) or full page (178 mm; 7”) width, 
but arrangements can be made to publish text-figures that must be larger. Any 
lettering in illustrations should follow the recommendations of Collinson (1962). 


Authors must provide three (3) copies of the text and accompanying illus- 
trative material. The text and line-drawings may be reproduced xerographically, 
but glossy prints at publication scale must be supplied for all half-tone illustrations 
and photographic plates. These prints should be identified clearly on the back. 


All dated text-citations must be referenced, except those that appear only 
within long-form synonymies. Additional references may be listed separately if 
their importance can be demonstrated by a short general comment, or individual 
annotations. Referenced publication titles must be spelled out in their entirety. 
Citations of illustrations within the monograph bear initial capitals (e.g., Plate, 
Text-figure), but citations of illustrations in other articles appear in lower-case 
letters (e.g., plate, text-figure). 


Original plate photomounts should have oversize cardboard backing and 
strong tracing paper overlays. These photomounts should be retained by the author 
until the manuscript has been formally accepted for publication. Explanations of 
text-figures should be interleaved on separate numbered pages within the text, 
and the approximate position of the text-figure in the text should be indicated. 
Explanations of plates follow the Bibliography. 


Authors are requested to enclose $10 with each manuscript submitted, to 
cover costs of postage during the review process. 


Collinson, J. 
1962. Size of lettering for text-figures. Journal of Paleontology, vol. 36, 
p. 1402. 


N 
1 
PIG CANYON 
ANTICLINE 2 
ARCENTE 
CANYON 


Caballero 
(Miss.) 


Caballero 


M 
bl “ 
D TANI ae” 


- 
Hy Th] 509m 


Montoya 
(Valmont) 


gn 
cov 
— a SS =] ee eon 
wbh f 
PI 
wby 
g 
PI St Des ] 526h — 
a Fusselman — 
> 
PI H, ] 3052a 
cov 
Z 


Text-figure 4.— Correlated stratigraphic sections of Devonian rocks, Sacramento Mountains, New Mexico. Datum 
is the top of the Ofate Formation; internal datum in the Sly Gap Formation is the Hyporhyridina bed (see Text-figure 
wo location of sections, Text-figure 5 for explanations of abbreviations and lithic patterns). 


4 5 
3 ALAMO INDIAN WELLS 
6 
YON ANYON 
ALAMO SAN c 
DEADMAN 
CANYON 


Caballero 


ee 


Da 


Fusselman 


Fusselman 


Fusselman 


ph 
Caballero (type section) 
SS iS Caballero 
AD 50 _ 
] WS pass) 5 
D Hy 
bl 
506c 
Hy Th M SG 
509; 
Hy N 
abies Hb 
Ma eS 
c > 529x 
gn 
f 
Des 
D PI St 529] 
ee ] : ——s bn-bI 
PI ee wae SS 
Fusselman — 


Ss 
7 
MULE CANYON 20 ft 
(composite) 
9 
Caballero DOG 
fl CANYON 
Lake Valley 
(Andrecito Mbr.) 8 10 
SAN ANDRES ESCONDIDO 
gnn 
CANYON CANYON 
Caballero Caballero Caballero 
PI 523z 
Des | s2at 
br 
eee 523: 
Fusselman Des x 
cov vi \ se 
NS 5 
7 \ EEZd erst] 5230 
cov 
Fusselman 


Fusselman 


Gilbert Dennison Harris 
(1864 - 1952) 


Founder of the Bulletins of American Paleontology (1895) 


TORN 0.27710-390-9 


uit 


et od SF 48 OR Ha 0 
bE ‘ ERA HN otbiel ot bh : On 
; Ra Teas prin si one Ph ee 
Prius ay SHAW wee ve ‘ 
ana Fe + 
pee ee ree cen 
. Basia Fate, 


ol Cai . 
MAGN EME IT @ 1- F cre Dey eres , F * 
Wriaa raven gimatel Feira okigeh. aie e ie “Ves , ; 
pi neteatased pensineess nes f 4 t ‘ , 
ee wae tt 3 cs s s1 v Pas 
aie nome Crd Were) , . * 
peo ? ; : i teh A 
err 2 oo AA i v4 ‘ 
Stat ni i ee ieee ays 2 fi ; ' ‘ 
? th ae wa Pinks : 
ome ey ae ee ait Coe u f 
ees aie aero 
iol y Sabu n on UR Le . 
pepe fia Nae Oa st ; 
ene : ‘ 
Ct Bh ti : r Protye iia 7 : 
EAA okey onie sat ra re ‘ i 
PAO EE AA es MEE AAR A tie oh fa i oe tise : 
ede Pedtne : orn ‘ { . 
FAA EaEie Aes oanek 9 * a hee “ i J f 
Fe 3 * oe 
Fe atin BA od ph psi a : 
aie f 
fr niet wy op o th iF re: i 4 ; 
RPA pave oe backed Or ti yy tg Ms 
ht. MeO Leet hdr y 7 
ase boa ted 4 : , 
, J Ly 1 . . 
eras, ' ¢ F 
foe ae - 
, 
Ware ere : 
HS sony ete 
tine 
SOE ari 
ion tg a Rady 
a - a hs @ 3; 3 , : cing c : 
MS gS SME A one taee ga e 4.0 Mae - ” 2 , 
LPI ER ERT ANT aD aE ee f 
te ae 
3 s 
Ft Wd 2 q 
a ¥: 7 
et Des : 4 : T bbe. 4 ? 
er ees F 
ihe porate wife bale 
tat sie Oa WS ee ny 
Bie ewe ; 
het ote AE 
Nea eae ene 
AER SRD OR 
BRR 
oa 4 ‘ , G 
A 
Bites 
eo é ¢ . 
Sot x 3 
WAST TSE : 
cht ne | em, ¥ 
* y . 4 
Yea ed n . 
on oe Se eat ¥ 4 ‘ 
rea iy bs 
oe es 
1 f : i 
SPAR a % va c Shae a a 
brads aT if Ae 
ve ‘ 
rf On oii Saran eta ad 
; ea RES kh ‘ . ‘ 
el DRO art Set : 
Sioa inv ene yh d 
ATE: = ‘ 
va wy erences v f i b 
um CRY a aed ‘ . 
iA he OU eth 


Mt * oe * i , s ; 
x we “ a f 

ve 
see